Sample records for acadian flycatcher empidonax

  1. Sex determination of the Acadian Flycatcher using discriminant analysis

    USGS Publications Warehouse

    Wilson, R.R.

    1999-01-01

    I used five morphometric variables from 114 individuals captured in Arkansas to develop a discriminant model to predict the sex of Acadian Flycatchers (Empidonax virescens). Stepwise discriminant function analyses selected wing chord and tail length as the most parsimonious subset of variables for discriminating sex. This two-variable model correctly classified 80% of females and 97% of males used to develop the model. Validation of the model using 19 individuals from Louisiana and Virginia resulted in 100% correct classification of males and females. This model provides criteria for sexing monomorphic Acadian Flycatchers during the breeding season and possibly during the winter.

  2. Pathways and consequences of contaminant flux to Acadian flycatchers (Empidonax virescens) in urbanizing landscapes of Ohio, USA.

    PubMed

    Rowse, Linnea M; Rodewald, Amanda D; Sullivan, S Mažeika P

    2014-07-01

    A prevalent environmental contaminant, mercury (Hg) is mobile and persistent in aquatic systems, where it often occurs in its bioavailable form methylmercury. Because methylmercury can bioaccumulate in aquatic insects and then transfer to terrestrial food webs, riparian consumers reliant upon aquatic emergent insects, should be disproportionately affected. Using the aerial insectivore Acadian flycatcher (Empidonax virescens) as a focal species, we examined (1) the extent to which total Hg loads in breeding flycatchers affected body condition and reproductive output and (2) potential pathways of contaminant flux in 19 riparian forest fragments distributed across an urban-to-rural landscape gradient in Ohio, USA. From April-August 2011-2012, we collected blood samples from adult (n=76) and nestling (n=17 from 7 nests) flycatchers, monitored their annual reproductive success (i.e., total number of fledglings), and sampled water, sediment, and aquatic emergent insects at each site. Hg concentrations in adult flycatcher blood (47 to 584 μg/kg, x¯=211.8, SD=95.5) were low relative to published advisory levels and not related to body condition. However, even at low concentrations, blood Hg was negatively related to reproductive success, with a 0.83 decline in the number of fledglings per μg/kg (loge) increase of blood Hg. Adult flycatchers had 11× greater concentrations of blood Hg than their offspring. Hg levels in flycatcher blood were not predicted by Hg concentrations in sediment, water, or aquatic emergent insects, with the exception of rural landscapes alone, in which flycatcher Hg was negatively related to sediment Hg. In addition to illustrating the difficulty of predicting exposure pathways that may vary among landscape contexts, our study provides evidence that even trace levels of contaminants may impair reproductive success of free-living songbirds. Copyright © 2014 Elsevier B.V. All rights reserved.

  3. Southwestern willow flycatchers (Empidonax traillii extimus) in a grazed landscape: factors influencing brood parasitism

    Treesearch

    Katherine M. Brodhead; Scott H. Stoleson; Deborah M. Finch

    2007-01-01

    Brood parasitism by Brown-headed Cowbirds (Molothrus ater; hereafter "cowbirds") is an important factor contributing to the endangered status of the Southwestern Willow Flycatcher (Empidonax traillii extimus, hereafter "flycatcher"). We report on factors that influence brood parasitism on the flycatcher using...

  4. Risky business: Site selection by Acadian Flycatchers under threat of nest predation and brood parasitism

    USGS Publications Warehouse

    HazIer, K.R.; Cooper, R.J.; Twedt, D.J.

    2006-01-01

    Habitat quality is determined not only by habitat structure and the availability of resources, but also by competitors, cooperators, predators, and parasites. We hypothesized that, for passerines, minimizing risk from avian nest predators and brood parasites is an important factor in selecting a breeding site. Through the early part of two breeding seasons, we spot-mapped locations of Acadian Flycatchers (Empidonax virescens, territory selectors), Red-bellied Woodpeckers (Melanerpes carolinus, nest predators) and Brown-headed Cowbirds (Molothrus ater, brood parasites) in a 56-ha study area within an extensive bottomland hardwood forest. We were thereby able to determine the order of flycatcher territory settlement and nest initiation in relation to risk of predation and parasitism, while accounting for habitat structure. Male settlement was influenced by both habitat structure and risk avoidance. However, risk trom woodpeckers was relatively more important in the first season and risk from cowbirds in the second, evidently due to differences in the relative abundance of predator and brood-parasite in each year. For male flycatchers, settlement choices appear to be flexible in the face of changing 'risk landscapes.' For females, habitat structure was the most important predictor of nest site selection. Even so, there was evidence that females avoided cowbirds. Surprisingly, nest site selection was positively associated with woodpecker abundance in the first season when woodpeckers were present in greater numbers. Possible explanations for this contradictory result are discussed.

  5. Alder flycatcher

    Treesearch

    Scott H. Stoleson

    2010-01-01

    The alder flycatcher is a small flycatcher in the confusing genus Empidonax. It so closely resembles its look-alike congener, the willow flycatcher (E. traillii), that until 1973 they were considered a single species, Traill's flycatcher (American Ornithologists' Union 1973). As most published studies of Traill's...

  6. A survey protocol for the Southwestern Willow Flycatcher (Empidonax traillii extimus)

    USGS Publications Warehouse

    Tibbitts, Timothy J.; Sogge, Mark K.; Sferra, Susan J.

    1994-01-01

    The southwestern willow flycatcher (Empidonax traillii extimus) is a riparian obligate neotropical migrant, nesting in cottonwood-willow associations and structurally similar riparian vegetation associations. The southwestern willow flycatcher has declined through the twentieth century, primarily due to a number of factors, including loss and fragmentation of riparian habitat, brood parasitism by brown-headed cowbirds (Molothrus ater), invasion of riparian habitat by the exotic tamarisk (Tamarix sp.), and predation (Hunter et al. 1987), Unitt 1987, Hunter et al. 1988, Whitfield 1990, Harris 1991, Rosenberg et al. 1991). In 1991 the U.S. Fish and Wildlife Service (USFWS) designated the southwestern willow flycatcher as a candidate category 1 species (USFWS 1991), indicating that the USFWS had sufficient information to support listing under the Endangered Species Act of 1973, as amended (Act), but that a proposal to list was precluded by other listing actions of higher priority. In July 1993, the USFWS proposed to list E. t. extimus as an endangered species and to designate critical habitat under the Act (USFWS 1993). The states of Arizona, New Mexico, and California comprise most of the southwestern willow flycatcher's historic and current range. Each of these states lists the species as endangered [Arizona Game and Fish Department (AGFD) 1988, New Mexico Department of Game and Fish (NMDGF) 1988, California Department of Game and Fish 1991]. Because of the precarious status of the southwestern willow flycatcher (Unitt 1987, USFWS 1993), there is a need to identify as many remaining breeding locations as possible. This survey protocol was developed to facilitate and standardize breeding surveys, and is based primarily on extensive 1992 and 1993 field surveys. It was developed at the request of the Arizona Partners in flight, and organization of Federal and State agencies, nongovernmental organizations, and individuals. This protocol is intended to be useful

  7. Habitat and sex differences in physiological condition of breeding Southwestern Willow Flycatchers (Empidonax traillii extimus)

    USGS Publications Warehouse

    Owen, J.C.; Sogge, M.K.; Kern, M.D.

    2005-01-01

    The Southwestern Willow Flycatcher (Empidonax traillii extimus; here- after “flycatcher”) is a federally listed endangered species that breeds in densely vegetated riparian habitats dominated by native and exotic plants, including introduced monotypic saltcedar (Tamarix ramosissima). Some workers have theorized that saltcedar is unsuitable habitat for the flycatcher, primarily because it generally supports a smaller and less diverse invertebrate community (the flycatcher's food base) than native habitats (e.g. Salix spp.). However, differences in insect communities between native and saltcedar habitats are not proof that saltcedar habitats are inferior. The only way to evaluate whether the habitats differ in dietary or energetic quality is to document actual food limitation or its manifestations. Measurements of an individual's body condition and metabolic state can serve as indicators of environmental stressors, such as food limitation and environmental extremes. We captured 130 flycatchers breeding in native and saltcedar habitats in Arizona and New Mexico and measured 12 variables of physiological condition. These variables included body mass, fat level, body condition index, hematocrit, plasma triglycerides, plasma free fatty acids and glycerol, plasma glucose and beta-hydroxybutyrate, plasma uric acid, total leukocyte count, and heterophil-to-lymphocyte ratio. We found substantial sex-based differences in the condition of male and female flycatchers. Ten of the 12 measures of physiological condition differed significantly between the sexes. In all cases where male and female condition differed (except mass), the differences suggest that males were in poorer condition than females. We found few habitat-based differences in flycatcher condition. Only 3 of the 12 physiological condition indices differed significantly between habitats. Our data show that, at least in some parts of the flycatcher's range, there is no evidence that flycatchers breeding in

  8. Ecology and behaviour of the Alder Flycatcher Empidonax alnorum, Tyrannidae in winter

    USGS Publications Warehouse

    Foster, M.; Adams, N.J.; Slotow, R.H.

    1999-01-01

    The Alder Flycatcher Empidonax alnorum (Tyrannidae) winters from Colombia and Venezuela, south to northern Argentina. We studied the species in Manu National Park, Madre de Dios, Peru. In this area the species occupies riparian zones along lowland meander rivers as well as adjacent transition zone forest in the floodplain. These habitats have a number of age-related longitudinal zones (parallel to the river margin) that are short term (I-2 year) transitory stages in the succession to floodplain forest. The fast growing plant species that make up these habitats are suitable for commercial harvest for paper pulp. We examined habitat use of the species in an attempt to identify the effects of such harvest and resulting habitat loss on these birds. The Alder Flycatcher favours two longitudinal zones where it establishes territories. The territories may be long-term or transitory, and are often occupied and defended by two individuals, generally of different age. One bird tends to be more active in territory defence than the other. Defence involves patrolling, vocal duels, and chases. Birds sally for insects but also eat large quantities of fruit.

  9. Ecology and behaviour of the Alder Flycatcher Empidonax alnorum, Tyrannidae in winter

    USGS Publications Warehouse

    Foster, M.; Adams, N.J.; Slotow, R.H.

    1998-01-01

    The Alder Flycatcher Empidonax alnorum (Tyrannidae) winters from Colombia and Venezuela, south to northern Argentina. We studied the species in Manu National Park, Madre de Dios, Peru. In this area the species occupies riparian zones along lowland meander rivers as well as adjacent transition zone forest in the floodplain. These habitats have a number of age-related longitudinal zones (parallel to the river margin) that are short term (I-2 year) transitory stages in the succession to floodplain forest. The fast growing plant species that make up these habitats are suitable for commercial harvest for paper pulp. We examined habitat use of the species in an attempt to identify the effects of such harvest and resulting habitat loss on these birds. The Alder Flycatcher favours two longitudinal zones where it establishes territories. The territories may be long-term or transitory, and are often occupied and defended by two individuals, generally of different age. One bird tends to be more active in territory defence than the other. Defence involves patrolling, vocal duels, and chases. Birds sally for insects but also eat large quantities of fruit.

  10. Winter distribution of willow flycatcher subspecies (Distribución Invernal de las Subespecies de Empidonax traillii)

    USGS Publications Warehouse

    Paxton, Eben H.; Unitt, Philip; Sogge, Mark K.; Whitfield, Mary; Keim, Paul

    2011-01-01

    Documenting how different regions across a species' breeding and nonbreeding range are linked via migratory movements is the first step in understanding how events in one region can influence events in others and is critical to identifying conservation threats throughout a migratory animal's annual cycle. We combined two studies that evaluated migratory connectivity in the Willow Flycatcher (Empidonax traillii), one using mitochondrial DNA sequences from 172 flycatchers sampled throughout their winter range, and another which examined morphological characteristics of 68 museum specimens collected in the winter range. Our results indicate that the four subspecies occupy distinct but overlapping regions of the winter range. Connectivity between specific breeding and winter grounds appears to be moderate to strong, with distributions that suggest migration patterns of both the chain and leap-frog types connecting the breeding and nonbreeding grounds. The Pacific lowlands of Costa Rica appear to be a key winter location for the endangered Southwestern Willow Flycatcher (E. t. extimus), although other countries in Central America may also be important for the subspecies.

  11. Primer registro de Empidonax fulvifrons en Nicaragua

    Treesearch

    Francisco J. Muñoz; Wayne J Arendt; Liliana Chavarria; Pablo Somarriba; Aura L. Cruz

    2009-01-01

    English title: First documented report of buff-breasted fl ycatcher Empidonax fulvifrons in Nicargua. A buff-breasted flycatcher (Empidonax fulvifrons) was observed on 4 January 2008 in Loma Fría, Dipilto Municipality, Department of Nueva Segovia. The bird was sited in a pasture with scattered pines (Pinus oocarpa) adjacent to a pine-oak forest. It formed part of a...

  12. Geographic variation in the plumage coloration of willow flycatchers Empidonax traillii

    USGS Publications Warehouse

    Paxton, Eben H.; Sogge, Mark K.; Koronkiewicz, Thomas J.; McLeod, Mary Anne; Theimer, Tad C.

    2010-01-01

    The ability to identify distinct taxonomic groups of birds (species, subspecies, geographic races) can advance ecological research efforts by determining connectivity between the non-breeding and breeding grounds for migrant species, identifying the origin of migrants, and helping to refine boundaries between subspecies or geographic races. Multiple methods are available to identify taxonomic groups (e.g., morphology, genetics), and one that has played an important role for avian taxonomists over the years is plumage coloration. With the advent of electronic devices that can quickly and accurately quantify plumage coloration, the potential of using coloration as an identifier for distinct taxonomic groups, even when differences are subtle, becomes possible. In this study, we evaluated the degree to which plumage coloration differs among the four subspecies of the willow flycatcher Empidonax traillii, evaluated sources of variation, and considered the utility of plumage coloration to assign subspecies membership for individuals of unknown origin. We used a colorimeter to measure plumage coloration of 374 adult willow flycatchers from 29 locations across their breeding range in 2004 and 2005. We found strong statistical differences among the mean plumage coloration values of the four subspecies; however, while individuals tended to group around their respective subspecies' mean color value, the dispersion of individuals around such means overlapped. Mean color values for each breeding site of the three western subspecies clustered together, but the eastern subspecies' color values were dispersed among the other subspecies, rather than distinctly clustered. Additionally, sites along boundaries showed evidence of intergradation and intermediate coloration patterns. We evaluated the predictive power of colorimeter measurements on flycatchers by constructing a canonical discriminant model to predict subspecies origin of migrants passing through the southwestern U

  13. Status, ecology, and conservation of the southwestern willow flycatcher

    Treesearch

    Deborah M. Finch; Scott H. Stoleson

    2000-01-01

    This publication was prepared in response to a need expressed by southwestern agencies and organizations for a comprehensive assessment of the population status, history, biology, ecology, habitats, threats, and conservation of the southwestern willow flycatcher (Empidonax traillii extimus). The southwestern willow flycatcher was federally listed as...

  14. Winter behavior and ecology of the Alder Flycatcher (Empidonax alnorum) in Peru

    USGS Publications Warehouse

    Foster, M.S.

    2007-01-01

    The winter ecology and behavior of Alder Flycatchers (Empidonax alnorum) were studied along the Manu, a white-water meander river in Manu National Park, Madre de Dios, Peru during October and November, 1993 to 1997. The birds occupied territories in primary-succession habitats on growing point bars. They were most common in mixed stands of Tessaria integrifolia (Asteraceae) and Gynerium sagittatum (Gramineaceae) interspersed with bare sand areas. The uneven height of the Tessaria canopy, which resulted in openings in the vegetation large enough for the birds to flycatch, was an important habitat feature. Birds obtained insects, which formed about 96% of the diet, by aerial hawking (91%), perch gleaning (6%), and hover gleaning (3%). They also fed on fruit. Territory sizes ranged from 0.04 to 0.25 ha. Of nine territories that we observed closely, six were occupied by two birds each, two by one bird each, and one by three birds. Every territory had one dominant individual who was primarily responsible for territory defense; the other birds were associates. Vocalizations given included the fee-bee-o song, a two-syllable song, and the pit note, which are also given on the breeding grounds. A series of pits given increasingly rapidly signaled a territorial interaction. In aggressive encounters, the birds (1) interacted vocally, remaining on their territories and counter calling or exchanging agitated calls; (2) moved toward a common territorial boundary and engaged in a vocal duel; or (3) the dominant chased intruders out of the territory. Chases were most common when a wave of new birds entered the area. Dominant birds, which sang the full song, were probably adult males. Immature males do not sing a full song, and females are not known to sing in nature. Associate individuals were likely females or young males.

  15. Winter behavior and ecology of the Alder Flycatcher (Empidonax alnorum) in Peru

    USGS Publications Warehouse

    Foster, M.S.

    2007-01-01

    The winter ecology and behavior of Alder Flycatchers (Empidonax alnorum) were studied along the Manu, a white-water meander river in Manu National Park, Madre de Dios, Peru?? during October and November, 1993 to 1997. The birds occupied territories in primary-succession habitats on growing point bars. They were most common in mixed stands of Tessaria integrifolia (Asteraceae) and Gynerium sagittatum (Gramineaceae) interspersed with bare sand areas. The uneven height of the Tessaria canopy, which resulted in openings in the vegetation large enough for the birds to flycatch, was an important habitat feature. Birds obtained insects, which formed about 96% of the diet, by aerial hawking (91%), perch gleaning (6%), and hover gleaning (3%). They also fed on fruit. Territory sizes ranged from 0.04 to 0.25 ha. Of nine territories that we observed closely, six were occupied by two birds each, two by one bird each, and one by three birds. Every territory had one dominant individual who was primarily responsible for territory defense; the other birds were associates. Vocalizations given included the fee-bee-o song, a two-syllable song, nd the pit note, which are also given on the breeding grounds. A series of pits given increasingly rapidly signaled a territorial interaction. In aggressive encounters, the birds (1) interacted vocally, remaining on their territories and counter calling or exchanging agitated calls; (2) moved toward a common territorial boundary and engaged in a vocal duel; or (3) the dominant chased intruders out of the territory. Chases were most common when a wave of new birds entered the area. Dominant birds, which sang the full song, were probably adult males. Immature males do not sing a full song, and females are not known to sing in nature. Associate individuals were likely females or young males. ?? The Neotropical Ornithological Society.

  16. Primary song by a juvenile willow flycatcher

    USGS Publications Warehouse

    Sogge, M.K.

    1997-01-01

    The timing of song development in suboscines, in which song appears not to be learned from other adults is poorly known. The Willow Flycatcher (Empidonax traillii) is a suboscine with a primary song typically referred to as fitz-bew. I report here an instance of very early singing by a 6-8-wk-old Willow Flycatcher, which sang in an aggressive context in response to a recording of adult flycatcher song. This is exceptionally early development of primary song, even among suboscines. Early song development may assist in the defense of winter territories.

  17. Parents or predators: Examining intraseasonal variation in nest survival for migratory passerine

    Treesearch

    Robin Hirsch-Jacobson; W. Andrew Cox; Emily E. Tewes; Frank R., III Thompson; John Faaborg

    2012-01-01

    For birds, risk of nest predation can vary within a breeding season, but few data exist that explain why such variation occurs. We investigated intraseasonal variation of nest survival of the Acadian Flycatcher (Empidonax virescens) in Midwestern forests and tested whether four of the adults' reproductive strategies (clutch size, nest...

  18. Landscape forest cover and edge effects on songbird nest predation vary by nest predator

    Treesearch

    W. Andrew Cox; Frank R. III Thompson; John Faaborg

    2012-01-01

    Rates of nest predation for birds vary between and within species across multiple spatial scales, but we have a poor understanding of which predators drive such patterns. We video-monitored nests and identified predators at 120 nests of the Acadian Flycatcher (Empidonax virescens) and the Indigo Bunting (Passerina cyanea) at eight...

  19. Species and temporal factors affect predator-specific rates of nest predation for forest songbirds in the midwest

    Treesearch

    W. Andrew Cox; Frank R. III Thompson; John Faaborg

    2012-01-01

    Knowledge of the relative contributions of predator species to overall rates of nest predation can improve our understanding of why predation risk varies, but the identity of predators is seldom known. We used video technology to identify nest predators of the tree-nesting Acadian Flycatcher (Empidonax virescens) and the shrub-nesting Indigo Bunting...

  20. The role of ecologic diversification in sibling speciation of Empidonax flycatchers (Tyrannidae): multigene evidence from mtDNA.

    PubMed

    Johnson, N K; Cicero, C

    2002-10-01

    Avian genera characterized by sibling species with distinctive habitat preferences present an evolutionary enigma in view of the more commonplace occurrence of syntopic congeners that differ strikingly in colour and pattern. No existing theory has explained the evolutionary background that led to these differences. Here we propose that great phenotypic similarity among some groups of sibling species limits their coexistence and that clues to their radiation can be seen in patterns of geographical occurrence. To illustrate our thesis we focused on the New World flycatcher genus Empidonax, a group of 15 species notorious for their great phenotypic similarity. Using 3069 base pairs of mitochondrial DNA from four genes, we produced a complete molecular phylogeny that identified four clades, three of which represent close relatives. The fourth clade includes only E. virescens, which apparently has no close living relatives. The majority of species, including many distant relatives, are completely (58.1%) or essentially (6.7%) allopatric in breeding distribution and exhibit striking ecological segregation into distinctive climate-vegetation zones. Even where ranges overlap, occupancy of the same habitat by different species is rare. Phylogenetic and distributional patterns in Empidonax suggest a peripatric model of stepwise colonization and then range expansion of small groups of pioneers during glacial periods into initially enlarging, distinctive habitats destined to be widespread during interglacials. Vicariance is not indicated in the absence of barriers of appropriate age and geographical position. Rapoport's rule that northern species have larger ranges than southern species is strongly supported.

  1. Saltcedar and Southwestern Willow Flycatchers: Lessons From Long-term Studies in Central Arizona

    Treesearch

    M. K. Sogge; E. H. Paxton; April A. Tudor

    2006-01-01

    The endangered Southwestern Willow Flycatcher (Empidonax traillii extimus: SWWF) is a riparian-obligate bird that breeds only in dense, typically wet riparian vegetation. Since the mid-1990s, biologists have discovered a substantial number of flycatchers breeding in habitat dominated by exotic saltcedar (Tamarix ramossisima) in...

  2. Female-biased sex ratio, polygyny, and persistence in the endangered Southwestern Willow Flycatcher (Empidonax traillii extimus)

    USGS Publications Warehouse

    Kus, Barbara E.; Howell, Scarlett; Wood, Dustin A.

    2017-01-01

    Demographic changes in populations, such as skewed sex ratios, are of concern to conservationists, especially in small populations in which stochastic and other events can produce declines leading to extirpation. We documented a decline in one of the few remaining populations of Southwestern Willow Flycatcher (Empidonax traillii extimus) in southern California, USA, which dropped from 40 to 5 adults between 2000 and 2015. Declines were unequal between sexes (94% for males, 82% for females). Adult sex ratios were female-biased in 10 of 16 yr. The proportion of paired males that were polygynous ranged from 0% to 100%, depending on the ratio of females to males in the adult population. Some males paired with up to 5 females simultaneously. We investigated the role of nestling sex ratio in the female-biased adult sex ratio by using genetic techniques to determine sex from blood samples collected from 162 nestlings in 72 nests from 2002 to 2009. Both population-level and within-brood nestling sex ratios were female-biased, and were not influenced by nest order (first or subsequent), parental mating type (monogamous or polygynous), or year. Disproportionately more females than males were recruited into the breeding population, mirroring nestling and fledgling sex ratios. It thus appears that a skewed nestling sex ratio has contributed to a female-biased adult population, which in turn has influenced mating behavior. We propose that the capacity for polygyny, which generally occurs at low levels in Southwestern Willow Flycatchers, has allowed this population to persist through a decline that might otherwise have resulted in extinction.

  3. Microhabitat use by breeding Southwestern Willow Flycatchers on the Gila River, New Mexico

    Treesearch

    Scott H. Stoleson; Deborah M. Finch

    2003-01-01

    The endangered Southwestern Willow Flycatcher (Empidonax traillii extimus) breeds at numerous sites throughout its range that vary greatly in floristics, vegetation structure, and the extent of human alteration of the habitat. Here we present information on nesting habitat characteristics of Willow Flycatchers in the largest extant population of the subspecies along...

  4. Site fidelity, territory fidelity, and natal philopatry in Willow Flycatchers (Empidonax traillii)

    USGS Publications Warehouse

    Sedgwick, James A.

    2004-01-01

    I investigated the causes and consequences of adult breeding-site fidelity, territory fidelity, and natal philopatry in Willow Flycatchers (Empidonax traillii) in southeastern Oregon over a 10-year period, testing the general hypothesis that fidelity and dispersal distances are influenced by previous breeding performance. Willow Flycatchers adhered to the generally observed tendencies of passerine birds for low natal philopatry and high breedingsite fidelity. Site fidelity (return to the study area) of adult males (52.0%) and females (51.3%), and median dispersal distances between seasons (16 m vs. 19 m) were similar. Previous breeding performance and residency (age-experience), but not study-site quality, explained site fidelity in females. Site fidelity of females rearing 4–5 young (64.4%) exceeded that of unsuccessful females (40.0%), breeding dispersal was less (successful: 15 m; unsuccessful: 33 m), and novice residents were more site-faithful than former residents. Probability of site fidelity was higher for previously successful females (odds ratio = 4.76), those with greater seasonal fecundity (odds ratio = 1.58), novice residents (odds ratio = 1.41), and unparasitized females (odds ratio = 2.76). Male site fidelity was not related to residency, site quality, or previous breeding performance. Territory fidelity (return to the previous territory) in females was best explained by previous breeding performance, but not by site quality or residency. Previously successful females were more likely to return to their territory of the previous season than either unsuccessful (odds ratio = 14.35) or parasitized birds (odds ratio = 6.38). Male territory fidelity was not related to residency, site quality, or previous breeding performance. Natal philopatry was low (7.8%) and similar for males and females. Site quality appeared to influence philopatry, given that no birds reared at a low-quality study site returned there to breed, and birds reared there dispersed

  5. Final recovery plan of the southwestern willow flycatcher (Empidonax traillii extimus)

    Treesearch

    Deborah M. Finch; Stephen I. Rothstein; Jon C. Boren; William L. Graf; Jerry L. Holechek; Barbara E. Kus; Robert M. Marshall; Molly M. Pohl; Susan J. Sferra; Mark K. Sogge; Julie C. Stromberg; Bradley A. Valentine; Mary J. Whitfield; Sartor O. Williams

    2002-01-01

    The Southwestern Willow Flycatcher Recovery Team is composed of a Technical Subgroup (pg. ii), six Implementation Subgroups (Appendix A), and a Tribal Working Group. The Technical Subgroup consists of 14 academic scientists, researchers, and resource managers with a wide range of expertise in avian biology and ecology, southwestern willow flycatcher ecology, cowbird...

  6. Parasitism, productivity, and population growth: response of Least Bell's Vireos (Vireo bellii pusillus) and Southwestern Willow Flycatchers (Empidonax traillii extimus) to cowbird (Molothrus spp.) control

    USGS Publications Warehouse

    Kus, Barbara E.; Whitfield, Mary J.

    2005-01-01

    Cowbird (Molothrus spp.) control is a major focus of recovery-oriented management of two endangered riparian bird species,the Least Bell's Vireo (Vireo bellii pusillus) and Southwestern Willow Flycatcher (Empidonax traillii extimus). During the past 20 years, annual trapping of cowbirds at Least Bell's Vireo and Southwestern Willow Flycatcher breeding sites has eliminated or reduced parasitism in comparison with pretrapping rates and, thereby, significantly increased seasonal productivity of nesting pairs. Enhanced productivity, in turn, has resulted in an 8-fold increase in numbers of Least Bell's Vireos; Southwestern Willow Flycatcher abundance, however, has changed little, and at some sites has declined despite cowbird control. Although generally successful by these short-term measures of host population response, cowbird control poses potential negative consequences for long-term recovery of endangered species. As currently employed, cowbird control lacks predetermined biological criteria to trigger an end to the control, making these species' dependence on human intervention open-ended. Prolonged reliance on cowbird control to manage endangered species can shift attention from identifying and managing other factors that limit populations--in particular, habitat availability. On the basis of our analysis of these long-term programs, we suggest that cowbird control be reserved for short-term crisis management and be replaced, when appropriate, by practices emphasizing restoration and maintenance of natural processes on which species depend. /// El manejo orientado hacia la recuperación de dos especies de aves ribereñas Vireo belli pusillus y Empidonax trailli extimus se ha focalizado principalmente en el control de los Molothrus spp parásitos. Durante los pasados 20 años, la captura anual de los Molothrus en las áreas de nidificación de Vireo belli pusillus y Empidonax trailli extimus ha eliminado o reducido el parasitismo en comparación con las tasas

  7. Nest site characteristics of Hammond's and Pacific-slope flycatchers in northwest California

    Treesearch

    H.F. Sakai; B.R. Noon

    1991-01-01

    Thirty nests of Hammond’s (Empidonax hammondii) and 88 nests of Pacificslope (E. difficilis) Flycatchers were located in different-aged Douglas-fir (Pseudotsuga menziesii)/ tanoak (Lithocarpus densiflorus) dominated forests at 12 study sites in northwestern California during the...

  8. Recovering southwestern willow flycatcher populations will benefit riparian health

    Treesearch

    Deborah M. Finch

    1999-01-01

    When the U.S. Fish and Wildlife Service (USFWS) listed the southwestern willow flycatcher (Empidonax traillii exigua) as federally endangered in 1995, new incentives, controversies and energy were generated to conserve and restore southwestern riparian ecosystems. Close attention has been focused on river and stream conservation in the Southwest since at least 1977,...

  9. Willow Flycatcher nonbreeding territory defense behavior in Costa Rica

    USGS Publications Warehouse

    Sogge, M.K.; Koronkiewicz, T.J.; van Riper, Charles; Durst, S.L.

    2007-01-01

    We studied the intraspecific territorial defense behavior of wintering Willow Flycatchers (Empidonax traillii) in Costa Rica using a randomized playback experiment that exposed male and female birds to recordings of Willow Flycatcher songs and calls, Lesser Ground Cuckoo (Morococcyx erythropygius) vocalizations, and random noise. Flycatchers of both sexes responded most strongly to simulated conspecific territory intrusion, and the agonistic behaviors that we observed were similar to those seen during natural intraspecific encounters in winter. Both males and females engaged in song and aggressive behaviors in defense of territories, and there was no significant difference between the sexes in scored agonistic responses. The similarity between the sexes in intraspecific territorial defense behaviors and aggressiveness may account for both sexes of flycatchers using the same habitats at our study sites in Costa Rica, and wintering females defending territories against males. The Willow Flycatcher, a sexually monomorphic species, differs in this way from a number of sexually dimorphic passerines, in which behaviorally dominant males occur in more optimal winter habitats. ?? The Cooper Ornithological Society 2007.

  10. Breeding biology and behavior of Hammond's and western flycatchers in northwestern California

    Treesearch

    H.F. Sakai

    1988-01-01

    The Hammond's (Empidonax hammondii) and Western (E. difficilis) flycatchers are sympatric within portions of their breeding ranges in north- western California. Westerns are common in a variety of habitats (Bent 1942:247, Johnson 1980:11-23); Hammond's are more abundant at higher elevations (Bent 1942:226, Johnson...

  11. Migration of the Willow Flycatcher along the Middle Rio Grande

    Treesearch

    Wang Yong; Deborah M. Finch

    1997-01-01

    We studied timing, abundance, subspecies composition, fat stores, stopover length, and habitat use of Willow Flycatchers (Empidonax traillii) during spring and fall stopover along the Middle Rio Grande, New Mexico. Spring migration started in mid-May and lasted about a month. Fall migration started in early-August and also lasted about a month. The most abundant...

  12. Geographic variation in the song of Willow Flycatchers: Differentiation between Empidonax traillii adastus and E. t. extimus

    USGS Publications Warehouse

    Sedgwick, J.A.

    2001-01-01

    The vocal signatures of the primary song form (“fitz-bew”) of the endangered Southwestern Willow Flycatcher (Empidonax traillii extimus) and its northern counterpart, E. t. adastus, are distinctive. Songs of the extimus subspecies are longer (total song, note, internote) and frequencies at maximum amplitude are lower than those of adastus. I used vocal evidence to clarify the distributional limits of the Southwestern Willow Flycatcher and that of the geographically adjacent subspecies, E. t. adastus. Unweighted pair-group method using averaging (UPGMA) cluster analysis and canonical discriminant analysis revealed that (1) low elevation, southerly desert populations (Arizona, New Mexico, and southern Utah) have a unique vocal identity corresponding to populations in the range of E. t. extimus; (2) northerly song groups (Oregon, Colorado, and northern Utah) share a different song type corresponding to populations in the range of E. t. adastus; and (3) a departure from vocal and morphological congruence occurs for a population of high-elevation Arizona birds that, although in the currently accepted range of E. t. extimus, sings songs acoustically similar to more northern populations (E. t. adastus). Multiple regression of song distance on latitude and elevation, and a comparison of a matrix of song distances with a matrix of latitude and elevation dissimilarities, demonstrated that song populations sort out by both latitude and elevation: birds with the vocal identity of extimus occur as far north as 37°N if at low elevation, and those acoustically similar to adastus occur as far south as 33.7°N if at high elevation. The vocal background of northern New Mexico birds appears to be intermediate between that of extimus and adastus, suggesting that northern New Mexico is a zone of intermixing and intergradation between the subspecies. Pure forms of E. t. extimus apparently do not occur in Colorado because even the southernmost populations are acoustically similar to

  13. Winter distribution of willow flycatcher subspecies

    USGS Publications Warehouse

    Paxton, E.H.; Unitt, P.; Sogge, M.K.; Whitfield, M.; Keim, P.

    2011-01-01

    Documenting how different regions across a species' breeding and nonbreeding range are linked via migratory movements is the first step in understanding how events in one region can influence events in others and is critical to identifying conservation threats throughout a migratory animal's annual cycle. We combined two studies that evaluated migratory connectivity in the Willow Flycatcher (Empidonax traillii), one using mitochondrial DNA sequences from 172 flycatchers sampled throughout their winter range, and another which examined morphological characteristics of 68 museum specimens collected in the winter range. Our results indicate that the four subspecies occupy distinct but overlapping regions of the winter range. Connectivity between specific breeding and winter grounds appears to be moderate to strong, with distributions that suggest migration patterns of both the chain and leap-frog types connecting the breeding and nonbreeding grounds. The Pacific lowlands of Costa Rica appear to be a key winter location for the endangered Southwestern Willow Flycatcher (E. t. extimus), although other countries in Central America may also be important for the subspecies. ?? The Cooper Ornithological Society 2011.

  14. Reproductive ecology of Dusky Flycatchers in western Montana

    USGS Publications Warehouse

    Sedgwick, James A.

    1993-01-01

    Breeding ecology of Dusky Flycatchers (Empidonax oberholseri) was studied in western Montana from May-August 1974. Dusky Flycatchers were monogamous and single-brooded although some pairs made renesting attempts after first nests failed. Length of the reproductive cycle for first nesting attempts, from arrival on the breeding grounds to fledging, was about 70 days. All nests were placed in shrubs, primarily ninebark (Physocarpus malvaceus) and Rocky Mountain maple (Acer glabrum), and were an average of 150 cm above the ground. Females incubated a clutch with an average of 4.0 eggs for 15-16 days, and the average nestling period was 17.5 days. Both sexes fed nestlings and fledglings, but only females brooded nestlings. Egg survival was 63.8%, hatching success was 95.4%, and nestling survival was 61.9% for an overall probability of 0.376 percent that an egg produced a fledgling. Predation was the major cause of nest failure. Dusky Flycatchers reared an average of 1.9 fledglings/pair.

  15. The status of the Willow and Pacific-slope flycatchers in northwestern California and southern Oregon

    Treesearch

    C. John Ralph; K. Hollinger

    2003-01-01

    The Willow (Empidonax traillii) and Pacific-slope (E. dificilis) flycatchers are generally similar in their morphology and foraging, but differ in their habitat and population dynamics. Through a concentration of constant-effort mist-netting stations, we documented the movements and composition of populations over a relatively...

  16. A satellite model of Southwestern Willow Flycatcher (Empidonax traillii extimus) breeding habitat and a simulation of potential effects of tamarisk leaf beetles (Diorhabda spp.), southwestern United States

    USGS Publications Warehouse

    Hatten, James R.

    2016-08-08

    The study described in this report represents the first time that a satellite model has been used to identify potential Southwestern Willow Flycatcher (Empidonax traillii extimus) (hereinafter referred to as “flycatcher”) breeding habitat rangewide for 2013–15. Fifty-seven Landsat scenes were required to map the entire range of the flycatcher, encompassing parts of six States and more than 1 billion 30-meter pixels. Predicted flycatcher habitat was summarized in a hierarchical fashion from largest to smallest: regionwide, State, U.S. Fish and Wildlife Service (FWS) management unit, 7.5-minute quadrangle, and critical-habitat reach. The term “predicted habitat” is used throughout this report to distinguish areas the satellite model predicts as suitable flycatcher habitat from what may actually exist on the ground. A rangewide accuracy assessment was done with 758 territories collected in 2014, and change detection was done with yearly habitat maps to identify how and where habitat changed over time. Additionally, effects of tamarisk leaf beetles (Diorhabda spp.) on flycatcher habitat were summarized for the lower Virgin River from 2010 to 2015, and simulations of how tamarisk leaf beetles may affect flycatcher habitat in the lower Colorado and upper Gila Rivers were done for 2015. Model results indicated that the largest areas of predicted flycatcher habitat at elevations below 1,524 meters were in New Mexico and Arizona, areas followed in descending order by California, Texas, Nevada, Utah, and Colorado. By FWS management unit, the largest area of flycatcher habitat during all 3 years were the Middle Rio Grande (New Mexico), followed by the Upper Gila (Arizona and New Mexico) and Middle Gila/San Pedro (Arizona) management units. The area of predicted flycatcher habitat varied considerably in 7.5-minute quadrangles, ranging from 0 to1,398 hectares (ha). Averaged across 3 years, the top three producing quadrangles were Paraje Well (New Mexico), San Marcial

  17. A habitat overlap analysis derived from Maxent for Tamarisk and the South-western Willow Flycatcher

    Treesearch

    Patricia York; Paul Evangelista; Sunil Kumar; James Graham; Curtis Flather; Thomas Stohlgren

    2011-01-01

    Biologic control of the introduced and invasive, woody plant tamarisk (Tamarix spp, saltcedar) in south-western states is controversial because it affects habitat of the federally endangered South-western Willow Flycatcher (Empidonax traillii extimus). These songbirds sometimes nest in tamarisk where floodplain-level invasion replaces native habitats. Biologic control...

  18. The ecology of the Southwestern Willow Flycatcher in central Arizona - A 10-year synthesis report

    USGS Publications Warehouse

    Paxton, Eben H.; Sogge, Mark K.; Durst, Scott L.; Theimer, Tad C.; Hatten, James R.

    2007-01-01

    BACKGROUND From 1996 to 2005, the U.S. Geological Survey (USGS) conducted a demographic study of the Southwestern Willow Flycatcher (Empidonax traillii extimus) in Arizona in collaboration with the Arizona Game and Fish Department (AGFD). The study was begun the year following the listing of the Southwestern Willow Flycatcher as an endangered species. At the time of the listing, very little was known about the biology and threats to the flycatcher, and one of the main objectives of the study was to gather detailed long-term information on the biology of the flycatcher. This report is organized into eight chapters. Following the introductory chapter, we deal with specific aspects of flycatcher ecology and habitat use in each of six separate chapters. We end with a concluding chapter that synthesizes information into broad topical themes that address key management issues. Each of the core chapters (chapters 2 through 7) conclude with a list of management considerations derived from the findings of the respective chapter.

  19. Injury due to leg bands in willow flycatchers

    USGS Publications Warehouse

    Sedgwick, J.A.; Klus, R.J.

    1997-01-01

    We report an apparently unusually high incidence of leg injury in Willow Flycatchers (Empidonax traillii) as a result of banding and color banding. Color bands and U.S. Fish and Wildlife Service (USFWS) bands applied to Willow Flycatchers from 1988-1995 resulted in an overall leg injury rate of 9.6% to birds returning to our study areas in subsequent years. Most injuries occurred on legs with only color band(s) (58.3%) or on legs with both a USFWS band and a color band (35%); only 6.7% of injuries (4/60) were due to USFWS bands alone, yielding an overall USFWS band injury rate of only 0.6%. Injuries ranged from severe (swollen, bleeding legs; a missing foot) to relatively minor (irritations on the tarsus). Amputation of the foot occurred in 33.9% of the cases. Return rates of adult injured birds in the year(s) following injury were significantly lower than for the population at large.

  20. Nestling sex ratio in the Southwestern Willow Flycatcher

    USGS Publications Warehouse

    Paxton, E.H.; Sogge, M.K.; McCarthey, T.D.; Keim, P.

    2002-01-01

    Using molecular-genetic techniques, we determined the gender of 202 Southwestern Willow Flycatcher (Empidonax traillii extimus) nestlings from 95 nests sampled over a five-year period. Overall nestling sex ratio did not vary significantly from 50:50 among years, by clutch order, or by mating strategy (monogamous vs. polygamous pairings). However, we did observe significant differences among the four sites sampled, with sex ratios biased either toward males or females at the different sites. Given the small population sizes and geographic isolation of many of the endangered subspecies' breeding populations, sex-ratio differences may have localized negative impacts. ?? The Cooper Ornithological Society 2002.

  1. Nestling sex ratios in the southwestern willow flycatcher

    USGS Publications Warehouse

    Paxton, E.H.; Sogge, M.K.; McCarthey, Tracy; Keim, Paul

    2002-01-01

    Using molecular-genetic techniques, we determined the gender of 202 Southwestern Willow Flycatcher (Empidonax traillii extimus) nestlings from 95 nests sampled over a five-year period. Overall nestling sex ratio did not vary significantly from 50:50 among years, by clutch order, or by mating strategy (monogamous vs. polygamous pairings). However, we did observe significant differences among the four sites sampled, with sex ratios biased either toward males or females at the different sites. Given the small population sizes and geographic isolation of many of the endangered subspecies' breeding populations, sex-ratio differences may have localized negative impacts.

  2. Available data support protection of the Southwestern Willow Flycatcher under the Endangered Species Act

    USGS Publications Warehouse

    Theimer, Tad C.; Smith, Aaron D.; Mahoney, Sean M.; Ironside, Kirsten E.

    2016-01-01

    Zink (2015) argued there was no evidence for genetic, morphological, or ecological differentiation between the federally endangered Southwestern Willow Flycatcher (Empidonax traillii extimus) and other Willow Flycatcher subspecies. Using the same data, we show there is a step-cline in both the frequency of a mtDNA haplotype and in plumage variation roughly concordant with the currently recognized boundary between E. t. extimus and E. t adastus, the subspecies with which it shares the longest common boundary. The geographical pattern of plumage variation is also concordant with previous song analyses differentiating those 2 subspecies and identified birds in one low-latitude, high-elevation site in Arizona as the northern subspecies. We also demonstrate that the ecological niche modeling approach used by Zink yields the same result whether applied to the 2 flycatcher subspecies or to 2 unrelated species, E. t. extimus and Yellow Warbler (Setophaga petechia). As a result, any interpretation of those results as evidence for lack of ecological niche differentiation among Willow Flycatcher subspecies would also indicate no differentiation among recognized species and would therefore be an inappropriate standard for delineating subspecies. We agree that many analytical techniques now available to examine genetic, morphological, and ecological differentiation would improve our understanding of the distinctness (or lack thereof) of Willow Flycatcher subspecies, but we argue that currently available evidence supports protection of the Southwestern Willow Flycatcher under the Endangered Species Act.

  3. A Natural History Summary and Survey Protocol for the Southwestern Willow Flycatcher

    USGS Publications Warehouse

    Sogge, Mark K.; ,; Ahlers, Darrell; ,; Sferra, Susan J.; ,

    2010-01-01

    The Southwestern Willow Flycatcher (Empidonax traillii extimus) has been the subject of substantial research, monitoring, and management activity since it was listed as an endangered species in 1995. When proposed for listing in 1993, relatively little was known about the flycatcher's natural history, and there were only 30 known breeding sites supporting an estimated 111 territories rangewide (Sogge and others, 2003a). Since that time, thousands of presence/absences surveys have been conducted throughout the historical range of the flycatcher, and many studies of its natural history and ecology have been completed. As a result, the ecology of the flycatcher is much better understood than it was just over a decade ago. In addition, we have learned that the current status of the flycatcher is better than originally thought: as of 2007, the population was estimated at approximately 1,300 territories distributed among approximately 280 breeding sites (Durst and others, 2008a). Concern about the Southwestern Willow Flycatcher on a rangewide scale was brought to focus by Unitt (1987), who described declines in flycatcher abundance and distribution throughout the Southwest. E. t. extimus populations declined during the 20th century, primarily because of habitat loss and modification from activities, such as dam construction and operation, groundwater pumping, water diversions, and flood control. In 1991, the U.S. Fish and Wildlife Service (USFWS) designated the Southwestern Willow Flycatcher as a candidate category 1 species (U.S. Fish and Wildlife Service, 1991). In July 1993, the USFWS proposed to list E. t. extimus as an endangered species and to designate critical habitat under the Act (U.S. Fish and Wildlife Service, 1993). A final rule listing E. t. extimus as endangered was published in February 1995 (U.S. Fish and Wildlife Service, 1995); critical habitat was designated in 1997 (U.S. Fish and Wildlife Service, 1997). The USFWS Service released a Recovery Plan for

  4. Vocal frequency change reflects different responses to anthropogenic noise in two suboscine tyrant flycatchers

    PubMed Central

    Francis, Clinton D.; Ortega, Catherine P.; Cruz, Alexander

    2011-01-01

    Anthropogenic noise is prevalent across the globe and can exclude birds from otherwise suitable habitat and negatively influence fitness; however, the mechanisms responsible for species' responses to noise are not always clear. One effect of noise is a reduction in effective acoustic communication through acoustic masking, yet some urban songbirds may compensate for masking by noise through altering their songs. Whether this vocal flexibility accounts for species persistence in noisy areas is unknown. Here, we investigated the influence of noise on habitat use and vocal frequency in two suboscine flycatchers using a natural experiment that isolated effects of noise from confounding stimuli common to urban habitats. With increased noise exposure, grey flycatcher (Empidonax wrightii) occupancy declined, but vocal frequency did not change. By contrast, ash-throated flycatcher (Myiarchus cinerascens) occupancy was uninfluenced by noise, but individuals in areas with greater noise amplitudes vocalized at a higher frequency, although the increase (≈200 kHz) may only marginally improve communication and may represent a secondary effect from increased vocal amplitude. Even so, the different flycatcher behavioural responses suggest that signal change may help some species persist in noisy areas and prompt important questions regarding which species will cope with an increasingly noisy world. PMID:21123268

  5. Survivorship across the annual cycle of a migratory passerine, the willow flycatcher

    USGS Publications Warehouse

    Paxton, Eben H.; Durst, Scott L.; Sogge, Mark K.; Koronkiewicz, Thomas J.; Paxton, Kristina L.

    2017-01-01

    Annual survivorship in migratory birds is a product of survival across the different periods of the annual cycle (i.e. breeding, wintering, and migration), and may vary substantially among these periods. Determining which periods have the highest mortality, and thus are potentially limiting a population, is important especially for species of conservation concern. To estimate survival probabilities of the willow flycatcher Empidonax traillii in each of the different periods, we combined demographic data from a 10-year breeding season study with that from a 5-year wintering grounds study. Estimates of annual apparent survival for breeding and wintering periods were nearly identical (65–66%), as were estimates of monthly apparent survival for both breeding and wintering stationary periods (98–99%). Because flycatchers spend at least half the year on the wintering grounds, overall apparent survivorship was lower (88%) on the wintering grounds than on the breeding grounds (97%). The migratory period had the highest mortality rate, accounting for 62% of the estimated annual mortality even though it comprises only one quarter or less of the annual cycle. The migratory period in the willow flycatcher and many other neotropical migrants is poorly understood, and further research is needed to identify sources of mortality during this crucial period.

  6. 77 FR 43107 - Endangered and Threatened Species Permit Applications

    Federal Register 2010, 2011, 2012, 2013, 2014

    2012-07-23

    ... presence/absence surveys for southwestern willow flycatcher (Empidonax traillii extimus) within Arizona...), southwestern willow flycatcher (Empidonax trailii extimus), and Yuma clapper rail (Rallus longirostris... conduct presence/absence surveys and collect plant material (leaves, flowers) of white bladderpod...

  7. Describing Willow Flycatcher habitats: scale perspectives and gender differences

    USGS Publications Warehouse

    Sedgwick, James A.; Knopf, Fritz L.

    1992-01-01

    We compared habitat characteristics of nest sites (female-selected sites) and song perch sites (male-selected sites) with those of sites unused by Willow Flycatchers (Empidonax traillii) at three different scales of vegetation measurement: (1) microplot (central willow [Salix spp.] bush and four adjacent bushes); (2) mesoplot (0.07 ha); and, (3) macroplot (flycatcher territory size). Willow Flycatchers exhibited vegetation preferences at all three scales. Nest sites were distinguished by high willow density and low variability in willow patch size and bush height. Song perch sites were characterized by large central shrubs, low central shrub vigor, and high variability in shrub size. Unused sites were characterized by greater distances between willows and willow patches, less willow coverage, and a smaller riparian zone width than either nest or song perch sites. At all scales, nest sites were situated farther from unused sites in multivariate habitat space than were song perch sites, suggesting (1) a correspondence among scales in their ability to describe Willow Flycatcher habitat, and (2) females are more discriminating in habitat selection than males. Microhabitat differences between male-selected (song perch) and female-selected (nest) sites were evident at the two smaller scales; at the finest scale, the segregation in habitat space between male-selected and female-selected sites was greater than that between male-selected and unused sites. Differences between song perch and nest sites were not apparent at the scale of flycatcher territory size, possibly due to inclusion of (1) both nest and song perch sites, (2) defended, but unused habitat, and/or (3) habitat outside of the territory, in larger scale analyses. The differences between nest and song perch sites at the finer scales reflect their different functions (e.g., nest concealment and microclimatic requirements vs. advertising and territorial defense, respectively), and suggest that the exclusive use

  8. Elucidating spatially explicit behavioral landscapes in the Willow Flycatcher

    USGS Publications Warehouse

    Bakian, Amanda V.; Sullivan, Kimberly A.; Paxton, Eben H.

    2012-01-01

    Animal resource selection is a complex, hierarchical decision-making process, yet resource selection studies often focus on the presence and absence of an animal rather than the animal's behavior at resource use locations. In this study, we investigate foraging and vocalization resource selection in a population of Willow Flycatchers, Empidonax traillii adastus, using Bayesian spatial generalized linear models. These models produce “behavioral landscapes” in which space use and resource selection is linked through behavior. Radio telemetry locations were collected from 35 adult Willow Flycatchers (n = 14 males, n = 13 females, and n = 8 unknown sex) over the 2003 and 2004 breeding seasons at Fish Creek, Utah. Results from the 2-stage modeling approach showed that habitat type, perch position, and distance from the arithmetic mean of the home range (in males) or nest site (in females) were important factors influencing foraging and vocalization resource selection. Parameter estimates from the individual-level models indicated high intraspecific variation in the use of the various habitat types and perch heights for foraging and vocalization. On the population level, Willow Flycatchers selected riparian habitat over other habitat types for vocalizing but used multiple habitat types for foraging including mountain shrub, young riparian, and upland forest. Mapping of observed and predicted foraging and vocalization resource selection indicated that the behavior often occurred in disparate areas of the home range. This suggests that multiple core areas may exist in the home ranges of individual flycatchers, and demonstrates that the behavioral landscape modeling approach can be applied to identify spatially and behaviorally distinct core areas. The behavioral landscape approach is applicable to a wide range of animal taxa and can be used to improve our understanding of the spatial context of behavior and resource selection.

  9. Territoriality, site fidelity, and survivorship of willow flycatchers wintering in Costa Rica

    USGS Publications Warehouse

    Koronkiewicz, T.J.; Sogge, M.K.; van Riper, Charles; Paxton, E.H.

    2006-01-01

    We studied wintering Willow Flycatchers (Empidonax traillii) in two seasonal freshwater wetland habitats in northwestern Costa Rica during five boreal winters, to determine habitat occupancy, overwinter and between-year site and territory fidelity, and the degree to which the sexes maintain and defend winter territories. Both males and females used agonistic displays, song, and other vocalizations to maintain and defend mutually exclusive winter territories. Males were generally more abundant than females, but this varied by site and year. There was no significant difference in male and female territory size, nor any indication of sexual habitat segregation. Similarity in morphology and aggressiveness between the sexes may account for the lack of habitat segregation and the ability of females to maintain territories at wintering sites. Each year, 80%-92% of banded flycatchers that were present in midwinter remained at the site until late winter; of these, 86%-100% of individuals maintained the same territories throughout the entire period. We also observed nonterritorial floaters that subsequently established and held winter territories. Between-year site fidelity averaged 68%, and almost all returning birds established territories with boundaries similar to the previous year. Between-year apparent survivorship estimates ranged annually from 54%-72%, with no difference between sites but weak support for higher survivorship of males compared to females. Values for winter site and territory fidelity were generally higher than those reported for other species and for Willow Flycatchers on the breeding grounds; between-year survivorship estimates were similar to those reported for breeding flycatchers. ?? The Cooper Ornithological Society 2006.

  10. Food habits of the southwestern willow flycatcher during the nesting season

    USGS Publications Warehouse

    Drost, Charles A.; Paxton, Eben H.; Sogge, Mark K.; Whitfield, Mary J.

    2003-01-01

    The food habits and prey base of the endangered Southwestern Willow Flycatcher (Empidonax traillii extimus) are not well known. We analyzed prey remains in 59 fecal samples from an intensively-studied population of this flycatcher at the Kern River Preserve in southern California. These samples were collected during the nesting season in 1996 and 1997 from adults caught in mist nets, and from nestlings temporarily removed from the nest for banding. A total of 379 prey individuals were identified in the samples. Dominant prey taxa, both in total numbers and in percent occurrence, were true bugs (Hemiptera), flies (Diptera), and beetles (Coleoptera). Leafhoppers (Homoptera: Cicadellidae), spiders (Araneae), bees and wasps (Hymenoptera), and dragonflies and damselflies (Odonata) were also common items. Diet composition was significantly different between years, due to a large difference in the numbers of spiders between 1996 and 1997. There was also a significant difference between the diet of young and adults, with the diet of young birds having significantly higher numbers of odonates and beetles. There was a trend toward diet differences between males and females, but this was not significant at the P = 0.05 level.

  11. Southwestern Willow Flycatcher Breeding Site and Territory Summary - 2007

    USGS Publications Warehouse

    Durst, Scott L.; Sogge, Mark K.; Stump, Shay D.; Walker, Hira A.; Kus, Barbara E.; Sferra, Susan J.

    2008-01-01

    The Southwestern willow flycatcher (Empidonax traillii extimus; hereafter references to willow flycatcher and flycatcher refer to E.t. extimus, except where specifically noted) is an endangered bird that breeds only in dense riparian habitats in parts of six Southwestern states (Arizona, New Mexico, southern California, extreme southern Nevada, southern Utah, and southwestern Colorado). Since 1993, hundreds of Southwestern willow flycatcher surveys have been conducted each year, and many new flycatcher breeding sites located. This document synthesizes the most current information available on all known Southwestern willow flycatcher breeding sites. This rangewide data synthesis was designed to meet two objectives: (1) identify all known Southwestern willow flycatcher breeding sites and (2) assemble data to estimate population size, location, habitat, and other information for all breeding sites, for as many years as possible, from 1993 through 2007. This report provides data summaries in terms of the number of flycatcher sites and the number of territories. When interpreting and using this information, it must be kept in mind that a 'site' is a geographic location where one or more willow flycatchers establishes a territory. Sites with unpaired territorial males are considered breeding sites, even if no nesting attempts were documented. A site is often a discrete patch of riparian habitat but may also be a cluster of riparian patches; there is no standardized definition for site, and its use varies within and among states. For example, five occupied habitat patches along a 10-km stretch of river might be considered five different sites in one state but only a single site in another state. This lack of standardization makes comparisons based on site numbers problematic. Researchers for this report generally deferred to statewide summary documents or to local managers and researchers when delineating a site for inclusion in the database. However, to avoid inflating

  12. Genetic variation in the endangered Southwestern Willow Flycatcher

    USGS Publications Warehouse

    Busch, Joseph; Miller, Mark P.; Paxton, E.H.; Sogge, M.K.; Keim, Paul

    2000-01-01

    The Southwestern Willow Flycatcher (Empidonax traillii extimus) is an endangered Neotropical migrant that breeds in isolated remnants of dense riparian habitat in the southwestern United States. We estimated genetic variation at 20 breeding sites of the Southwestern Willow Flycatcher (290 individuals) using 38 amplified fragment length polymorphisms (AFLPs). Our results suggest that considerable genetic diversity exists within the subspecies and within local breeding sites. Statistical analyses of genetic variation revealed only slight, although significant, differentiation among breeding sites (Mantel's r = 0.0705, P < 0.0005; θ = 0.0816, 95% CI = 0.0608 to 0.1034; ΦST = 0.0458, P < 0.001). UPGMA cluster analysis of the AFLP markers indicates that extensive gene flow has occurred among breeding sites. No one site stood out as being genetically unique or isolated. Therefore, the small level of genetic structure that we detected may not be biologically significant. Ongoing field studies are consistent with this conclusion. Of the banded birds that were resighted or recaptured in Arizona during the 1996 to 1998 breeding seasons, one-third moved between breeding sites and two-thirds were philopatric. Low differentiation may be the result of historically high rangewide diversity followed by recent geographic isolation of breeding sites, although observational data indicate that gene flow is a current phenomenon. Our data suggest that breeding groups of E. t. extimus act as a metapopulation.

  13. A habitat overlap analysis derived from maxent for tamarisk and the south-western willow flycatcher

    NASA Astrophysics Data System (ADS)

    York, Patricia; Evangelista, Paul; Kumar, Sunil; Graham, James; Flather, Curtis; Stohlgren, Thomas

    2011-06-01

    Biologic control of the introduced and invasive, woody plant tamarisk ( Tamarix spp, saltcedar) in south-western states is controversial because it affects habitat of the federally endangered South-western Willow Flycatcher ( Empidonax traillii extimus). These songbirds sometimes nest in tamarisk where floodplain-level invasion replaces native habitats. Biologic control, with the saltcedar leaf beetle ( Diorhabda elongate), began along the Virgin River, Utah, in 2006, enhancing the need for comprehensive understanding of the tamarisk-flycatcher relationship. We used maximum entropy (Maxent) modeling to separately quantify the current extent of dense tamarisk habitat (>50% cover) and the potential extent of habitat available for E. traillii extimus within the studied watersheds. We used transformations of 2008 Landsat Thematic Mapper images and a digital elevation model as environmental input variables. Maxent models performed well for the flycatcher and tamarisk with Area Under the ROC Curve (AUC) values of 0.960 and 0.982, respectively. Classification of thresholds and comparison of the two Maxent outputs indicated moderate spatial overlap between predicted suitable habitat for E. traillii extimus and predicted locations with dense tamarisk stands, where flycatcher habitat will potentially change flycatcher habitats. Dense tamarisk habitat comprised 500 km2 within the study area, of which 11.4% was also modeled as potential habitat for E. traillii extimus. Potential habitat modeled for the flycatcher constituted 190 km2, of which 30.7% also contained dense tamarisk habitat. Results showed that both native vegetation and dense tamarisk habitats exist in the study area and that most tamarisk infestations do not contain characteristics that satisfy the habitat requirements of E. traillii extimus. Based on this study, effective biologic control of Tamarix spp. may, in the short term, reduce suitable habitat available to E. traillii extimus, but also has the potential

  14. Species-specific variation in nesting and postfledging resource selection for two forest breeding migrant songbirds.

    PubMed

    Jenkins, Julianna M A; Thompson, Frank R; Faaborg, John

    2017-01-01

    Habitat selection is a fundamental component of community ecology, population ecology, and evolutionary biology and can be especially important to species with complex annual habitat requirements, such as migratory birds. Resource preferences on the breeding grounds may change during the postfledging period for migrant songbirds, however, the degree to which selection changes, timing of change, and whether all or only a few species alter their resource use is unclear. We compared resource selection for nest sites and resource selection by postfledging juvenile ovenbirds (Seiurus aurocapilla) and Acadian flycatchers (Empidonax virescens) followed with radio telemetry in Missouri mature forest fragments from 2012-2015. We used Bayesian discrete choice modeling to evaluate support for local vegetation characteristics on the probability of selection for nest sites and locations utilized by different ages of postfledging juveniles. Patterns of resource selection variation were species-specific. Resource selection models indicated that Acadian flycatcher habitat selection criteria were similar for nesting and dependent postfledging juveniles and selection criteria diverged when juveniles became independent from adults. After independence, flycatcher resource selection was more associated with understory foliage density. Ovenbirds differed in selection criteria between the nesting and postfledging periods. Fledgling ovenbirds selected areas with higher densities of understory structure compared to nest sites, and the effect of foliage density on selection increased as juveniles aged and gained independence. The differences observed between two sympatric forest nesting species, in both the timing and degree of change in resource selection criteria over the course of the breeding season, illustrates the importance of considering species-specific traits and postfledging requirements when developing conservation efforts, especially when foraging guilds or prey bases differ

  15. Resources for Teaching About Acadian Regions of Canada.

    ERIC Educational Resources Information Center

    Deveau, J. Alphonse

    1978-01-01

    Provides an annotated list of references for use in teaching about Acadian regions of Canada, along with a brief statement about the movements of eighteenth century Acadian settlers. References include general histories of the Maritime provinces which contain information on Acadian history, and publications on Acadian history in particular. (BC)

  16. Distribution and abundance of Least Bell’s Vireos (Vireo bellii pusillus) and Southwestern Willow Flycatchers (Empidonax traillii extimus) on the Middle San Luis Rey River, San Diego, southern California—2016 data summary

    USGS Publications Warehouse

    Allen, Lisa D.; Howell, Scarlett L.; Kus, Barbara E.

    2017-09-29

    Executive SummaryWe surveyed for Least Bell’s Vireos (LBVI) (Vireo bellii pusillus) and Southwestern Willow Flycatchers (SWFL) (Empidonax traillii extimus) along the San Luis Rey River, between College Boulevard in Oceanside and Interstate 15 in Fallbrook, California (middle San Luis Rey River), in 2016. Surveys were done from March 30 to July 11 (LBVI) and from May 18 to July 30 (SWFL). We found 142 LBVI territories, at least 106 of which were occupied by pairs. Six additional transient LBVIs were detected. Of 20 banded LBVIs detected in the survey area, 9 had been given full color-band combinations prior to 2016, although we were unable to determine the exact color combination of 1 female LBVI. Seven other LBVIs with single (natal) federal bands were recaptured and banded in 2016. Four vireos with single dark blue federal bands indicating that they were banded as nestlings on the lower San Luis Rey River could not be recaptured for identification.Three SFWL territories were observed in the survey area in 2016. Two territories were occupied by pairs and one by a male of unknown breeding status. Both pairs attempted to nest at least once, and both pairs were successful, fledging three young each. Nesting began in early June and continued into July. Brown-Headed Cowbird (Molothrus ater) eggs were not observed in either nest. An additional 12 transient Willow Flycatchers of unknown subspecies were detected in 2016.Two of the five resident SWFLs were originally banded as nestlings on Marine Corps Base Camp Pendleton. One male and one female were banded as nestlings on Camp Pendleton in 2009 and 2011, respectively. One natal male of unknown breeding status, originally banded as a nestling on the middle San Luis Rey River in 2015, was recaptured and given a unique color combination in 2016. This male was later detected on Marine Corps Base Camp Pendleton.

  17. Timing of the Acadian Orogeny in Northern New Hampshire.

    PubMed

    Eusden Jr; Guzofski; Robinson; Tucker

    2000-03-01

    New U-Pb geochronology constrains the timing of the Acadian orogeny in the Central Maine Terrane of northern New Hampshire. Sixteen fractions of one to six grains each of zircon or monazite have been analyzed from six samples: (1) an early syntectonic diorite that records the onset of the Acadian; (2) a schist, a migmatite, and two granites that together record the peak of the Acadian; and (3) a postkinematic pluton that records the end of the Acadian. Zircon from the syntectonic Wamsutta Diorite gives a 207Pb/206Pb age of circa 408 Ma, the time at which the boundary between the deforming orogenic wedge and the foreland basin was in the vicinity of the Presidential Range. This age agrees well with the Emsian position of the northwest migrating Acadian orogenic front and records the beginning of the Acadian in this part of the Central Maine Terrane. We propose a possible Acadian tectonic model that incorporates the geochronologic, structural, and stratigraphic data. Monazite from the schist, migmatite, Bigelow Lawn Granite, and Slide Peak Granite gives 207Pb/206U ages, suggesting the peak of Acadian metamorphism and intrusion of two-mica granites occurred at circa 402-405 Ma, the main pulse of Acadian orogenesis. Previously reported monazite ages from schists that likely record the peak metamorphism in the Central Maine Terrane of New Hampshire and western Maine range from circa 406-384 Ma, with younger ages in southeastern New Hampshire and progressively older ages to the west, north, and northeast. Acadian orogenesis in the Presidential Range had ended by circa 355 Ma, the 207Pb/235U age of monazite from the Peabody River Granite. From 408 to perhaps at least 394 Ma, Acadian orogenesis in the Presidential Range was typical of the tectonic style, dominated by synkinematic metamorphism, seen in central and southern New Hampshire, Massachusetts, and Connecticut. From no earlier than 394 Ma to as late as 355 Ma, the orogenesis was typical of the style in parts of

  18. Timing of the Acadian Orogeny in northern New Hampshire

    USGS Publications Warehouse

    Eusden, J.D.; Guzofski, C.A.; Robinson, A.C.; Tucker, R.D.

    2000-01-01

    New U-Pb geochronology constrains the timing of the Acadian orogeny in the Central Maine Terrane of northern New Hampshire. Sixteen fractions of one to six grains each of zircon or monazite have been analyzed from six samples: (1) an early syntectonic diorite that records the onset of the Acadian, (2) a schist, a migmatite, and two granites that together record the peak of the Acadian; and (3) a postkinematic pluton that records the end of the Acadian. Zircon from the syntectonic Wamsutta Diorite gives a 207Pb/206Pb age of circa 408 Ma, the time at which the boundary between the deforming orogenic wedge and the foreland basin was in the vicinity of the Presidential Range. This age agrees well with the Emsian position of the northwest migrating Acadian orogenic front and records the beginning of the Acadian in this part of the Central Maine Terrane. We propose a possible Acadian tectonic model that incorporates the geochronologic, structural, and stratigraphic data. Monazite from the schist, migmatite, Bigelow Lawn Granite, and Slide Peak Granite gives 207Pb/206U ages, suggesting the peak of Acadian metamorphism and intrusion of two-mica granites occurred at circa 402-405 Ma, the main pulse of Acadian orogenesis. Previously reported monazite ages from schists that likely record the peak metamorphism in the Central Maine Terrane of New Hampshire and western Maine range from circa 406-384 Ma, with younger ages in southeastern New Hampshire and progressively older ages to the west, north, and northeast. Acadian orogenesis in the Presidential Range had ended by circa 355 Ma, the 207Pb/235U age of monazite from the Peabody River Granite. From 408 to perhaps at least 394 Ma, Acadian orogenesis in the Presidential Range was typical of the tectonic style, dominated by synkinematic metamorphism, seen in central and southern New Hampshire, Massachusetts, and Connecticut. From no earlier than 394 Ma to as late as 355 Ma, the orogenesis was typical of the style in parts of

  19. 78 FR 28837 - Acadian Gas Pipeline System; Notice of Petition

    Federal Register 2010, 2011, 2012, 2013, 2014

    2013-05-16

    ... DEPARTMENT OF ENERGY Federal Energy Regulatory Commission [Docket No. PR11-129-001] Acadian Gas Pipeline System; Notice of Petition Take notice that on May 6, 2013, Acadian Gas Pipeline System (Acadian... concerns filed in the September 26, 2011 filing, as more fully detailed in the petition. Any person...

  20. Distribution and abundance of Least Bell’s Vireos (Vireo bellii pusillus) and Southwestern Willow Flycatchers (Empidonax traillii extimus) on the Middle San Luis Rey River, San Diego County, southern California—2017 data summary

    USGS Publications Warehouse

    Allen, Lisa D.; Howell, Scarlett L.; Kus, Barbara E.

    2018-04-20

    We surveyed for Least Bell’s Vireos (LBVI) (Vireo bellii pusillus) and Southwestern Willow Flycatchers (SWFL) (Empidonax traillii extimus) along the San Luis Rey River, between College Boulevard in Oceanside and Interstate 15 in Fallbrook, California (middle San Luis Rey River), in 2017. Surveys were conducted from April 13 to July 11 (LBVI) and from May 16 to July 28 (SWFL). We found 146 LBVI territories, at least 107 of which were occupied by pairs. Five additional transient LBVIs were detected. LBVIs used five different habitat types in the survey area: mixed willow, willow-cottonwood, willow-sycamore, riparian scrub, and upland scrub. Forty-four percent of the LBVIs occurred in habitat characterized as mixed willow and 89 percent of the LBVI territories occurred in areas with greater than 50 percent native plant cover. Of 16 banded LBVIs detected in the survey area, 8 had been given full color-band combinations prior to 2017. Four other LBVIs with single (natal) federal bands were recaptured and banded in 2017. Three LBVIs with single dark blue federal bands indicating that they were banded as nestlings on the lower San Luis Rey River and one LBVI with a single gold federal band indicating that it was banded as a nestling on Marine Corps Base Camp Pendleton (MCBCP) could not be recaptured for identification. One banded LBVI emigrated from the middle San Luis Rey River to the lower San Luis Rey River in 2017.One resident SWFL territory and one transient Willow Flycatcher of unknown subspecies (WIFL) were observed in the survey area in 2017. The resident SWFL territory, which was comprised of mixed willow habitat (5–50 percent native plant cover), was occupied by a single male from May 22 to June 21, 2017. No evidence of pairing or nesting activity was observed. The SWFL male was banded with a full color-combination indicating that he was originally banded as a nestling on the middle San Luis Rey River in 2014 and successfully bred in the survey area in 2016

  1. Chapter 7: Migration and winter ecology

    Treesearch

    Deborah M. Finch; Jeffrey F. Kelly; Jean-Luc E. Cartron

    2000-01-01

    The willow flycatcher (Empidonax traillii) is a Neotropical migrant that breeds in North America, but winters in Central and northern South America. Little specific information is known about migration and wintering ecology of the southwestern willow flycatcher (E. t. extimus) (Yong and Finch 1997). Our report applies principally...

  2. Chapter 3: Threats

    Treesearch

    Robert M. Marshall; Scott H. Stoleson

    2000-01-01

    The continued survival of the southwestern willow flycatcher (Empidonax traillii extimus) is threatened indirectly by the loss, modification, and fragmentation of riparian habitat, and directly by factors that impact the survival and reproductive success of flycatchers. Because the impact of habitat loss on small populations can be particularly...

  3. Chapter 6: Breeding season ecology

    Treesearch

    Mark K. Sogge

    2000-01-01

    The willow flycatcher (Empidonax traillii) breeds across much of the conterminous United States and in portions of extreme southern Canada. As might be expected in such a wide-ranging species, willow flycatchers in different portions of the range exhibit differences in appearance, song, and ecological characteristics. The intent of this chapter is to...

  4. Response

    Treesearch

    Wang Yong; Deborah M. Finch

    1999-01-01

    Hubbard (1999) criticizes our paper "Migration of the Willow Flycatcher along the middle Rio Grande" (Yong and Finch 1997), where we reported aspects of stopover ecology of the species including timing, abundance, fat stores, stopover length, and habitat use. Hubbard questions our identification of subspecies of the Willow Flycatcher (Empidonax traillii) and...

  5. Archaeological and Historical Reconnaissance and Literature Search of Cultural Resources within the Pembina River Project, Pembina and Cavalier Counties, North Dakota. Volume 1.

    DTIC Science & Technology

    1982-12-01

    SAythya americana Redhead Aythya valisineria Canvasback Aix sponsa Wood Duck Aythya collaris Ring-necked Ducks Aythya affinis Lesser Scaup Lophodytes...Chickadee Vireo olivaceus Red-eyed Vireo Vireo gilvus Warbling Vireo Empidonax Tinimus Least Flycatcher Contopus virens Eastern Wood Pewee Contopus...sordidulus Western Wood Pewee Myiarchus crinitus Great Crested Flycatcher Catharus fuscescens Veery Sejurus aurocapillus Ovenbird Mniotilta varia Black-and

  6. 75 FR 19987 - Endangered and Threatened Species Permit Applications

    Federal Register 2010, 2011, 2012, 2013, 2014

    2010-04-16

    ..., during normal business hours at the U.S. Fish and Wildlife Service, 500 Gold Ave. SW., Room 6034... seminuda), woundfin (Plagopterus argentissimus), Colorado pikeminnow (Ptychocheilus lucius), razorback... southwestern willow flycatcher (Empidonax taillii extimus) within Arizona, Colorado, and Utah. Authority: 16 U...

  7. Memoires d'une famille acadienne de Van Buren, Maine (Memoirs of an Acadian Family of Van Buren, Maine).

    ERIC Educational Resources Information Center

    Cyr, Marguerite

    These memoirs of an Acadian family present aspects of the cultural history of the Acadians in the St. John River Valley in Maine. The six chapters deal with the following topics: (1) a brief history of the land and the people; (2) the chronicles of a large Acadian family from the time of the arrival of their ancestors from France until the…

  8. Inferring the demographic history of European Ficedula flycatcher populations

    PubMed Central

    2013-01-01

    Background Inference of population and species histories and population stratification using genetic data is important for discriminating between different speciation scenarios and for correct interpretation of genome scans for signs of adaptive evolution and trait association. Here we use data from 24 intronic loci re-sequenced in population samples of two closely related species, the pied flycatcher and the collared flycatcher. Results We applied Isolation-Migration models, assignment analyses and estimated the genetic differentiation and diversity between species and between populations within species. The data indicate a divergence time between the species of <1 million years, significantly shorter than previous estimates using mtDNA, point to a scenario with unidirectional gene-flow from the pied flycatcher into the collared flycatcher and imply that barriers to hybridisation are still permeable in a recently established hybrid zone. Furthermore, we detect significant population stratification, predominantly between the Spanish population and other pied flycatcher populations. Conclusions Our results provide further evidence for a divergence process where different genomic regions may be at different stages of speciation. We also conclude that forthcoming analyses of genotype-phenotype relations in these ecological model species should be designed to take population stratification into account. PMID:23282063

  9. Niche separation in flycatcher-like species in the lowland rainforests of Malaysia.

    PubMed

    Mansor, Mohammad Saiful; Ramli, Rosli

    2017-07-01

    Niche theory suggests that sympatric species reduce interspecific competition through segregation of shared resources by adopting different attack manoeuvres. However, the fact that flycatcher-like bird species exclusively use the sally manoeuvre may thus challenge this view. We studied the foraging ecology of three flycatcher-like species (i.e. Paradise-flycatcher Terpsiphone sp., Black-naped Monarch Hypothymis azurea, and Rufous-winged Philentoma Philentoma pyrhoptera) in the Krau Wildlife Reserve in central Peninsular Malaysia. We investigated foraging preferences of each bird species and the potential niche partitioning via spatial or behavioural segregation. Foraging substrate was important parameter that effectively divided paradise-flycatcher from Black-naped Monarch and Rufous-winged Philentoma, where monarch and philentoma foraged mainly on live green leaves, while paradise-flycatcher foraged on the air. They also exhibited different foraging height preferences. Paradise-flycatcher, for instance, preferred the highest studied strata, while Black-naped Monarch foraged mostly in lower strata, and Rufous-winged Philentoma made use of the lowest strata. This study indicates that niche segregation occurs among sympatric species through foraging substrate and attack manoeuvres selection. Copyright © 2017 Elsevier B.V. All rights reserved.

  10. A multiscaled model of southwestern willow flycatcher breeding habitat

    USGS Publications Warehouse

    Hatten, J.R.; Paradzick, C.E.

    2003-01-01

    The southwestern willow flycatcher (SWFL; Empidonax traillii extimus) is an endangered songbird whose habitat has declined dramatically over the last century. Understanding habitat selection patterns and the ability to identify potential breeding areas for the SWFL is crucial to the management and conservation of this species. We developed a multiscaled model of SWTL breeding habitat with a Geographic Information System (GIS), survey data, GIS variables, and multiple logistic regressions. We obtained presence and absence survey data from a riverine ecosystem and a reservoir delta in south-central Arizona, USA, in 1999. We extracted the GIS variables from satellite imagery and digital elevation models to characterize vegetation and floodplain within the project area. We used multiple logistic regressions within a cell-based (30 X 30 m) modeling environment to (1) determine associations between GIS variables and breeding-site occurrence at different spatial scales (0.09-72 ha), and (2) construct a predictive model. Our best model explained 54% of the variability in breeding-site occurrence with the following variables: vegetation density at the site (0.09 ha), proportion of dense vegetation and variability in vegetation density within a 4.5-ha neighborhood, and amount of floodplain or flat terrain within a 41-ha neighborhood. The density of breeding sites was highest in areas that the model predicted to be most suitable within the project area and at an external test site 200 km away. Conservation efforts must focus on protecting not only occupied patches, but also surrounding riparian forests and floodplain to ensure long-term viability of SWTL. We will use the multiscaled model to map SWTL breeding habitat in Arizona, prioritize future survey effort, and examine changes in habitat abundance and quality over time.

  11. Extreme drought alters frequency and reproductive success of floaters in Willow Flycatchers

    USGS Publications Warehouse

    Theimer, Tad; Sogge, Mark K.; Cardinal, Suzanne N.; Durst, Scott L.; Paxton, Eben H.

    2018-01-01

    Changes in habitat quality, including those caused by extreme events like droughts and floods, could alter costs and benefits of territoriality and thereby the prevalence and reproductive consequences for individuals capable of breeding that do not do so (floaters). We studied floating behavior in a population of Southwestern Willow Flycatchers (Empidonax traillii extimus) in central Arizona during one year of extreme drought, one year of lake inundation, and three years of near average precipitation. In all years, most floaters were second year (SY) males, and most subsequently settled outside of the patch where they were detected in the floating year, suggesting that floaters did not “queue” at high-quality territories in order to achieve higher reproductive success in subsequent years. Instead, cohorts that floated in non-drought years had lower apparent survival and lower reproductive success compared to territorial birds. In the extreme drought year, however, the number of floaters was 1.5 times greater than in all other years combined, more females floated, and apparent survival and mean annual productivity in subsequent years was higher for males that floated in that year than for those that were territorial. Inundation of habitat due to rising reservoir levels did not result in an increase in floaters because many birds nested in inundated areas where trees projected above the water so that the relative amount of available habitat was not reduced to the extent habitat models predicted. Overall, our results indicate that the prevalence and reproductive and demographic consequences of floating can change under extreme climatic events like severe drought.

  12. Chapter 1: Introduction of the conservation assessment concept

    Treesearch

    Deborah M. Finch

    2000-01-01

    The goal of this document is to describe the current status, ecology, habitat, and threats of the southwestern willow flycatcher (Empidonax traillii extimus); to offer guidance for managing and protecting this Neotropical migrant and its habitats; and to identify gaps in our knowledge of the bird and its requirements. Goals, processes, and target...

  13. Implications of climate change for bird conservation in the southwestern U.S

    Treesearch

    Megan M. Friggens; Deborah M. Finch

    2015-01-01

    Future expected changes in climate and human activity threaten many riparian habitats, particularly in the southwestern U.S. Using Maximum Entropy (MaxEnt3.3.3) modeling, we characterized habitat relationships and generated spatial predictions of habitat suitability for the Lucy’s warbler (Oreothlypis luciae), the Southwestern willow flycatcher (Empidonax...

  14. Demographic Divergence History of Pied Flycatcher and Collared Flycatcher Inferred from Whole-Genome Re-sequencing Data

    PubMed Central

    Nadachowska-Brzyska, Krystyna; Burri, Reto; Olason, Pall I.; Kawakami, Takeshi; Smeds, Linnéa; Ellegren, Hans

    2013-01-01

    Profound knowledge of demographic history is a prerequisite for the understanding and inference of processes involved in the evolution of population differentiation and speciation. Together with new coalescent-based methods, the recent availability of genome-wide data enables investigation of differentiation and divergence processes at unprecedented depth. We combined two powerful approaches, full Approximate Bayesian Computation analysis (ABC) and pairwise sequentially Markovian coalescent modeling (PSMC), to reconstruct the demographic history of the split between two avian speciation model species, the pied flycatcher and collared flycatcher. Using whole-genome re-sequencing data from 20 individuals, we investigated 15 demographic models including different levels and patterns of gene flow, and changes in effective population size over time. ABC provided high support for recent (mode 0.3 my, range <0.7 my) species divergence, declines in effective population size of both species since their initial divergence, and unidirectional recent gene flow from pied flycatcher into collared flycatcher. The estimated divergence time and population size changes, supported by PSMC results, suggest that the ancestral species persisted through one of the glacial periods of middle Pleistocene and then split into two large populations that first increased in size before going through severe bottlenecks and expanding into their current ranges. Secondary contact appears to have been established after the last glacial maximum. The severity of the bottlenecks at the last glacial maximum is indicated by the discrepancy between current effective population sizes (20,000–80,000) and census sizes (5–50 million birds) of the two species. The recent divergence time challenges the supposition that avian speciation is a relatively slow process with extended times for intrinsic postzygotic reproductive barriers to evolve. Our study emphasizes the importance of using genome-wide data to

  15. Demographic divergence history of pied flycatcher and collared flycatcher inferred from whole-genome re-sequencing data.

    PubMed

    Nadachowska-Brzyska, Krystyna; Burri, Reto; Olason, Pall I; Kawakami, Takeshi; Smeds, Linnéa; Ellegren, Hans

    2013-11-01

    Profound knowledge of demographic history is a prerequisite for the understanding and inference of processes involved in the evolution of population differentiation and speciation. Together with new coalescent-based methods, the recent availability of genome-wide data enables investigation of differentiation and divergence processes at unprecedented depth. We combined two powerful approaches, full Approximate Bayesian Computation analysis (ABC) and pairwise sequentially Markovian coalescent modeling (PSMC), to reconstruct the demographic history of the split between two avian speciation model species, the pied flycatcher and collared flycatcher. Using whole-genome re-sequencing data from 20 individuals, we investigated 15 demographic models including different levels and patterns of gene flow, and changes in effective population size over time. ABC provided high support for recent (mode 0.3 my, range <0.7 my) species divergence, declines in effective population size of both species since their initial divergence, and unidirectional recent gene flow from pied flycatcher into collared flycatcher. The estimated divergence time and population size changes, supported by PSMC results, suggest that the ancestral species persisted through one of the glacial periods of middle Pleistocene and then split into two large populations that first increased in size before going through severe bottlenecks and expanding into their current ranges. Secondary contact appears to have been established after the last glacial maximum. The severity of the bottlenecks at the last glacial maximum is indicated by the discrepancy between current effective population sizes (20,000-80,000) and census sizes (5-50 million birds) of the two species. The recent divergence time challenges the supposition that avian speciation is a relatively slow process with extended times for intrinsic postzygotic reproductive barriers to evolve. Our study emphasizes the importance of using genome-wide data to

  16. Environmental Assessment for Leasing Nellis Air Force Base Land for Construction and Operation of a Solar Photovoltaic System, Clark County, Nevada

    DTIC Science & Technology

    2006-08-01

    oxides OSHA Occupational Safety and Health Administration PL Public Law PM10 Particulate matter ≤ 10 microns in diameter POL Petroleum, oils , and...several years, costs and demand for energy produced through non- renewable resources, such as crude oil , have increased dramatically. In response to...flycatcher Empidonax difficilus Rough-winged swallow Stelgidopteryx serripennis Ruby-crowned kinglet Regulus calendula Northern mockingbird Mimus

  17. Russian Olive Biology, Invasion, and Ecological Impacts in Western North America

    DTIC Science & Technology

    2016-07-01

    habitat for bird species , but is not functionally equivalent to native vegetation. Leatherman (2011) lists a variety of birds that consume fruits, sap...abundance, but intermediate avian species richness and diversity, in Russian olive stands. Russian olive bird communities were somewhat similar to...provided distinctive habitat for some bird species (e.g., least flycatchers [Empidonax minimus], red-eyed vireos [Vireo olivaceous]), while providing

  18. [sup 40]Ar/[sup 39]Ar evidence for delayed post-Acadian cooling in the southernmost Connecticut Valley Synclinorium

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Moecher, D.P.; Cosca, M.A.

    1992-01-01

    Available Ar-40/Ar-39 data for the Connecticut Valley Synclinorium (CVS) of the New England segment of the Appalachian Orogen indicate rapid post-Acadian cooling. However, new data indicate this pattern does not extend the entire length of the CVS. Ar-40/Ar-39 ages obtained from hornblende and muscovite in The Straits Schist indicate delayed cooling and a more complex post-Acadian thermal history. Data for the Seymour area are consistent with the studies above for the vicinity of the Waterbury Dome. The data farther south indicate one or more of the following: (1) slow (2--3C/Ma) post-Acadian cooling and uplift through the Permian; (2) post-Acadian coolingmore » through Hbl closure in the Mississippian with a subsequent Alleghanian metamorphism that did not exceed 500 C; or (3) post-Acadian cooling with subsequent metamorphism that approached 500 C or involved ductile recrystallization, partly resetting hornblende and totally resetting muscovite south of Derby. Petrologic evidence supporting (2) or (3) consists of widespread but not pervasive greenschist facies retrogression of Hbl + Pl + Sph assemblages in amphibolites to Act + Ep, and Grt + Ky + St assemblages in metapelites to Chl + Bt + Qz. The present data cannot resolve between (2) or (3). However, both are consistent with results of a study in the Bridgeport Synform that yield (1) a U-Pb monazite age of 296 [+-] 2 Ma from the Ansonia Leucogranite, implying the occurrence of an Alleghanian thermal event that promoted monazite growth; and, (2) a U-Pb cooling age of 360 Ma from sphene in the Pumpkin Ground Granodiorite, indicating that Alleghanian events did not exceed ca. 550 C.« less

  19. Southwestern willow flycatchers recaptured at wintering sites in Costa Rica

    USGS Publications Warehouse

    Koronkiewicz, Thomas J.; Sogge, Mark K.

    2001-01-01

    An adult Southwestern Willow Flycatcher banded in summer 1998 at Ash Meadows National Wildlife Refuge, Nevada, was recaptured the following winter in Santa Cruz, Costa Rica, then relocated at Ash Meadows during the 1999 breeding season. Another Southwestern Willow Flycatcher banded in 1999 as a nestling at Roosevelt Lake, Arizona, was recaptured in January 2000 on its wintering grounds in Bolsen, Costa Rica. The bird was recaptured at Roosevelt Lake in summer 2000, returned to the same wintering site in Bolsen the following winter, and was relocated at Roosevelt Lake in summer 2001.

  20. Using a remote sensing/GIS model to predict southwestern Willow Flycatcher breeding habitat along the Rio Grande, New Mexico

    USGS Publications Warehouse

    Hatten, James R.; Sogge, Mark K.

    2007-01-01

    Introduction The Southwestern Willow Flycatcher (Empidonax traillii extimus; hereafter SWFL) is a federally endangered bird (USFWS 1995) that breeds in riparian areas in portions of New Mexico, Arizona, southwestern Colorado, extreme southern Utah and Nevada, and southern California (USFWS 2002). Across this range, it uses a variety of plant species as nesting/breeding habitat, but in all cases prefers sites with dense vegetation, high canopy, and proximity to surface water or saturated soils (Sogge and Marshall 2000). As of 2005, the known rangewide breeding population of SWFLs was roughly 1,214 territories, with approximately 393 territories distributed among 36 sites in New Mexico (Durst et al. 2006), primarily along the Rio Grande. One of the key challenges facing the management and conservation of the Southwestern Willow Flycatcher is that riparian areas are dynamic, with individual habitat patches subject to cycles of creation, growth, and loss due to drought, flooding, fire, and other disturbances. Former breeding patches can lose suitability, and new habitat can develop within a matter of only a few years, especially in reservoir drawdown zones. Therefore, measuring and predicting flycatcher habitat - either to discover areas that might support SWFLs, or to identify areas that may develop into appropriate habitat - requires knowledge of recent/current habitat conditions and an understanding of the factors that determine flycatcher use of riparian breeding sites. In the past, much of the determination of whether a riparian site is likely to support breeding flycatchers has been based on qualitative criteria (for example, 'dense vegetation' or 'large patches'). These determinations often require on-the-ground field evaluations by local or regional SWFL experts. While this has proven valuable in locating many of the currently known breeding sites, it is difficult or impossible to apply this approach effectively over large geographic areas (for example, the

  1. Mesohabitat use of threatened hemlock forests by breeding birds of the Delaware River basin in northeastern United States

    USGS Publications Warehouse

    Ross, R.M.; Redell, L.A.; Bennett, R.M.; Young, J.A.

    2004-01-01

    Avian biodiversity may be at risk in eastern parks and forests due to continued expansion of the hemlock woolly adelgid (Adelges tsugae), an exotic homopteran insect native to East Asia. To assess avian biodiversity, mesohabitat relations, and the risk of species loss with declining hemlock forests in Appalachian park lands, 80 randomly distributed fixed-radius plots were established in which territories of breeding birds were estimated on four forest-terrain types (hemlock and hardwood benches and ravines) in the Delaware Water Gap National Recreation Area. Both species richness and number of territories were higher in hardwood than hemlock forest types and in bench than ravine terrain types. Four insectivorous species, Acadian flycatcher (Empidonax virescens), blue-headed vireo (Vireo solitarius), black-throated green warbler (Dendroica virens), and Blackburnian warbler (Dendroica fusca), showed high affinity for hemlock forest type and exhibited significantly greater numbers of territories in hemlock than hardwood sites. These species are hemlock-associated species at risk from continued hemlock decline in the Delaware River valley and similar forests of the mid-Atlantic east slope. Two of these species, the blue-headed vireo and Blackburnian warbler, appeared to specialize on ravine mesohabitats of hemlock stands, the vireo a low-to-mid canopy species, the warbler a mid-to-upper canopy forager. Unchecked expansion of the exotic adelgid and subsequent hemlock decline could negatively impact 3,600 pairs from the park and several million pairs from northeastern United States hemlock forests due to elimination of preferred habitat.

  2. Building hierarchical models of avian distributions for the State of Georgia

    USGS Publications Warehouse

    Howell, J.E.; Peterson, J.T.; Conroy, M.J.

    2008-01-01

    To predict the distributions of breeding birds in the state of Georgia, USA, we built hierarchical models consisting of 4 levels of nested mapping units of decreasing area: 90,000 ha, 3,600 ha, 144 ha, and 5.76 ha. We used the Partners in Flight database of point counts to generate presence and absence data at locations across the state of Georgia for 9 avian species: Acadian flycatcher (Empidonax virescens), brownheaded nuthatch (Sitta pusilla), Carolina wren (Thryothorus ludovicianus), indigo bunting (Passerina cyanea), northern cardinal (Cardinalis cardinalis), prairie warbler (Dendroica discolor), yellow-billed cuckoo (Coccyxus americanus), white-eyed vireo (Vireo griseus), and wood thrush (Hylocichla mustelina). At each location, we estimated hierarchical-level-specific habitat measurements using the Georgia GAP Analysis18 class land cover and other Geographic Information System sources. We created candidate, species-specific occupancy models based on previously reported relationships, and fit these using Markov chain Monte Carlo procedures implemented in OpenBugs. We then created a confidence model set for each species based on Akaike's Information Criterion. We found hierarchical habitat relationships for all species. Three-fold cross-validation estimates of model accuracy indicated an average overall correct classification rate of 60.5%. Comparisons with existing Georgia GAP Analysis models indicated that our models were more accurate overall. Our results provide guidance to wildlife scientists and managers seeking predict avian occurrence as a function of local and landscape-level habitat attributes.

  3. Two countries, one forest: Working beyond political boundaries in the Northern Appalachian/Acadian Forest

    Treesearch

    James Sullivan

    2007-01-01

    Two Countries, One Forest (2C1Forest) is a collaboration of conservation organizations and researchers committed to the long-term ecological health of the Northern Appalachian/ Acadian ecoregion of the United States and Canada.

  4. Prenuptial dental extractions in Acadian women: first report of a cultural tradition.

    PubMed

    Gordon, Sara C; Kaste, Linda M; Barasch, Andrei; Safford, Monika M; Foong, W Choong; ElGeneidy, Adry

    2011-12-01

    Prenuptial tooth extractions, extractions of all teeth in at least one dental arch before marriage, are not identified in the dental literature. Driven by a professional encounter, the purpose of this study was to confirm the existence of this practice among Acadian women. An 8-item survey instrument with space for comments was mailed to 182 dentists from traditionally Acadian regions of Canada. The survey was provided in English and French. Ninety dentists responded (50.3%); 8 of them (9%) had been asked to perform prenuptial extractions, and an additional 9 volunteered awareness of this practice. Awareness and requests were associated with dental practice in a county with a ≥20% French-speaking population. Prenuptial extractions in this population have been confirmed by the current cohort of dentists. The potential public health, clinical, and systemic health research implications for women who are edentulous for most of their adult life merit further study. Additionally, it is important to determine if interventions are needed to curtail cultural expectations of such practices.

  5. Modeling the Emergent Impacts of Harvesting Acadian Forests over 100+ Years

    NASA Astrophysics Data System (ADS)

    Luus, K. A.; Plug, L. J.

    2007-12-01

    Harvesting strategies and policies for Acadian forest in Nova Scotia, Canada, presently are set using Decision Support Models (DSMs) that aim to maximize the long-term (>100y) value of forests through decisions implemented over short time horizons (5-80 years). However, DSMs typically are aspatial, lack ecological processes and do not treat erosion, so the long-term (>100y) emergent impacts of the prescribed forestry decisions on erosion and vegetation in Acadian forests remain poorly known. To better understand these impacts, we created an equation-based model that simulates the evolution of a ≥4 km2 forest in time steps of 1 y and at a spatial resolution of 3 m2, the footprint of a single mature tree. The model combines 1) ecological processes of recruitment, competition, and mortality; 2) geomorphic processes of hillslope erosion; 3) anthropic processes of tree harvesting, replanting, and road construction under constraints imposed by regulations and cost/benefit ratio. The model uses digital elevation models, parameters (where available), and calibration (where measurements are not available) for conditions presently found in central Cape Breton, Nova Scotia. The model is unique because it 1) deals with the impacts of harvesting on an Acadian forest; and 2) vegetation and erosion are coupled. The model was tested by comparing the species-specific biomass of long-term (40 y) forest plot data to simulated results. At the spatial scale of individual 1 ha plots, model predictions presently account for approximately 50% of observed biomass changes through time, but predictions are hampered by the effects of serendipitous "random" events such as single tree windfall. Harvesting increases the cumulative erosion over 3000 years by 240% when compared to an old growth forest and significantly suppresses the growth of Balsam Fir and Sugar Maple. We discuss further tests of the model, and how it might be used to investigate the long-term sustainability of the

  6. Vortex wake, downwash distribution, aerodynamic performance and wingbeat kinematics in slow-flying pied flycatchers.

    PubMed

    Muijres, Florian T; Bowlin, Melissa S; Johansson, L Christoffer; Hedenström, Anders

    2012-02-07

    Many small passerines regularly fly slowly when catching prey, flying in cluttered environments or landing on a perch or nest. While flying slowly, passerines generate most of the flight forces during the downstroke, and have a 'feathered upstroke' during which they make their wing inactive by retracting it close to the body and by spreading the primary wing feathers. How this flight mode relates aerodynamically to the cruising flight and so-called 'normal hovering' as used in hummingbirds is not yet known. Here, we present time-resolved fluid dynamics data in combination with wingbeat kinematics data for three pied flycatchers flying across a range of speeds from near hovering to their calculated minimum power speed. Flycatchers are adapted to low speed flight, which they habitually use when catching insects on the wing. From the wake dynamics data, we constructed average wingbeat wakes and determined the time-resolved flight forces, the time-resolved downwash distributions and the resulting lift-to-drag ratios, span efficiencies and flap efficiencies. During the downstroke, slow-flying flycatchers generate a single-vortex loop wake, which is much more similar to that generated by birds at cruising flight speeds than it is to the double loop vortex wake in hovering hummingbirds. This wake structure results in a relatively high downwash behind the body, which can be explained by the relatively active tail in flycatchers. As a result of this, slow-flying flycatchers have a span efficiency which is similar to that of the birds in cruising flight and which can be assumed to be higher than in hovering hummingbirds. During the upstroke, the wings of slowly flying flycatchers generated no significant forces, but the body-tail configuration added 23 per cent to weight support. This is strikingly similar to the 25 per cent weight support generated by the wing upstroke in hovering hummingbirds. Thus, for slow-flying passerines, the upstroke cannot be regarded as inactive

  7. Decaying wood and tree regeneration in the Acadian Forest of Maine, USA

    Treesearch

    Jamie K. Weaver; Laura S. Kenefic; Robert S. Seymour; John C. Brissette

    2009-01-01

    We examined the effect of management history on the availability of decayed downed wood and the use of downed wood as a regeneration substrate in mixed-species stands in the Acadian Forest of Maine. Regeneration of red spruce (Picea rubens Sarg.), eastern hemlock (Tsuga canadensis (L.) Carr.), balsam fir (Abies balsamea L. Mill), and red maple (Acer rubrum L.) was...

  8. A technique to produce aluminum color bands for avian research

    USGS Publications Warehouse

    Koronkiewicz, T.J.; Paxton, E.H.; Sogge, M.K.

    2005-01-01

    We developed a technique to produce metal (aluminum) color bands, in response to concerns about leg injuries caused by celluloid-plastic color bands applied to Willow Flycatchers (Empidonax traillii). The technique involves color-anodized aluminum bands (unnumbered blanks and federal numbered bands), with auto pin-striping tape and flexible epoxy sealant, to create a variety of solid, half- and triple-split colors. This allows for hundreds of unique, high-contrast color combinations. During six consecutive years of application, these colored metal bands have resisted color fade compared to conventional celluloid-plastic bands, and have reduced leg injuries in the flycatcher. Although not necessarily warranted for all color-banding studies, these metal bands may provide a lower-impact option for studies of species known to be impacted by plastic color bands.

  9. Cystic fibrosis mutations in North American populations of French ancestry: Analysis of Quebec French-Canadian and Louisiana Acadian families

    PubMed Central

    Rozen, Rima; Schwartz, Robert H.; Hilman, Bettina C.; Stanislovitis, Pat; Horn, Glenn T.; Klinger, Katherine; Daigneault, Jocelyne; De Braekeleer, Marc; Kerem, Bat-sheva; Tsui, Lap-Chee; Fujiwara, T. Mary; Morgan, Kenneth

    1990-01-01

    A 3-bp deletion (ΔF508) in the cystic fibrosis (CF) gene is the mutation on the majority of CF chromosomes. We studied 112 CF families from North American populations of French ancestry: French-Canadian families referred from hospitals in three cities in Quebec and from the Saguenay-Lac St. Jean region of northeastern Quebec and Acadian families living in Louisiana. ΔF508 was present on 71%, 55%, and 70% of the CF chromosomes from the major-urban Quebec, Saguenay-Lac St. Jean, and Louisiana Acadian families, respectively. A weighted estimate of the proportion of ΔF508 in the French-Canadian patient population of Quebec was 70%. We found that 95% of the CF chromosomes with ΔF508 had D7S23 haplotype B, the most frequent haplotype on CF chromosomes. In the Saguenay-Lac St. Jean families, 86% of the CF chromosomes without ΔF508 had the B haplotype, compared with 31% for the major-urban Quebec and Louisiana Acadian families. The incidence of CF in the Saguenay-Lac St. Jean population was 1/895 live-born infants. PMID:2220803

  10. Estimation of linkage disequilibrium and interspecific gene flow in Ficedula flycatchers by a newly developed 50k single-nucleotide polymorphism array

    PubMed Central

    Kawakami, Takeshi; Backström, Niclas; Burri, Reto; Husby, Arild; Olason, Pall; Rice, Amber M; Ålund, Murielle; Qvarnström, Anna; Ellegren, Hans

    2014-01-01

    With the access to draft genome sequence assemblies and whole-genome resequencing data from population samples, molecular ecology studies will be able to take truly genome-wide approaches. This now applies to an avian model system in ecological and evolutionary research: Old World flycatchers of the genus Ficedula, for which we recently obtained a 1.1 Gb collared flycatcher genome assembly and identified 13 million single-nucleotide polymorphism (SNP)s in population resequencing of this species and its sister species, pied flycatcher. Here, we developed a custom 50K Illumina iSelect flycatcher SNP array with markers covering 30 autosomes and the Z chromosome. Using a number of selection criteria for inclusion in the array, both genotyping success rate and polymorphism information content (mean marker heterozygosity = 0.41) were high. We used the array to assess linkage disequilibrium (LD) and hybridization in flycatchers. Linkage disequilibrium declined quickly to the background level at an average distance of 17 kb, but the extent of LD varied markedly within the genome and was more than 10-fold higher in ‘genomic islands’ of differentiation than in the rest of the genome. Genetic ancestry analysis identified 33 F1 hybrids but no later-generation hybrids from sympatric populations of collared flycatchers and pied flycatchers, contradicting earlier reports of backcrosses identified from much fewer number of markers. With an estimated divergence time as recently as <1 Ma, this suggests strong selection against F1 hybrids and unusually rapid evolution of reproductive incompatibility in an avian system. PMID:24784959

  11. Status and breeding ecology of the southwestern willow flycatcher in the Grand Canyon

    USGS Publications Warehouse

    Sogge, M.K.; Tibbitts, T.J.; Petterson, J.R.

    1997-01-01

    Ernpidonax trailIll extirnus is one of several recognized subspecies of the Willow Flycatcher (Unitt 1987, Browning 1993), a neotropical migrant that breeds across much of North America. This southwestern race is a riparian obligate, nesting in dense patches of willow (Salix sp.), willow-cottonwood (Populus sp.), or other similarly structured habitats. In some areas of the Southwest, it nests in dense stands of tamarisk (Tamarix sp.). Willow Flycatchers were once widespread and locally common in the Southwest (Unitt 1987) but have declined to the point that E. t. extirnus was listed as an endangered subspecies in 1995 (USFWS 1995).

  12. Early reproductive success of western bluebirds and ash-throated flycatchers: a landscape-contaminant perspective.

    PubMed

    Fair, Jeanne M; Myers, Orrin B

    2002-01-01

    Eggshell quality, clutch size, sex ratio, and hatching success of western bluebirds (Sialia mexicana) and ash-throated flycatchers (Myiarchus cinerascens) were studied on a landscape-soil contaminant gradient at Los Alamos National Laboratory (LANL) in New Mexico from 1997 to 1999. A variety of contaminants (heavy metals, chemicals, insecticides, polychlorinated biphenyls (PCBs), organochlorines, and radioactive isotopes) range across different spatial scales and concentrations on LANL land. This study is an example of a monitoring program over a large area with varying degree of contamination that is used to highlight locations of concern for future research. There were two locations where the flycatcher had a lower hatching success. The bluebirds at Sandia wetland, a location of concern for PCBs. had a thinner eggshell thickness index (RATCLIFFE) and the eggs were smaller than at other locations. The flycatcher had thinner eggshells than bluebirds, which could add to sensitivity to exposure to contaminants. There was no variation in clutch size or sex ratio between locations or areas closer to contaminant release sites for both species. Percent females in the clutch ranged from 0 to 100% in the WEBL and from 33 to 67% for ATFL.

  13. 78 FR 343 - Endangered and Threatened Wildlife and Plants; Designation of Critical Habitat for Southwestern...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2013-01-03

    ...We, the U.S. Fish and Wildlife Service (Service), designate revised critical habitat for the southwestern willow flycatcher (Empidonax traillii extimus) (flycatcher) under the Endangered Species Act. In total, approximately 1,975 stream kilometers (1,227 stream miles) are being designated as critical habitat. These areas are designated as stream segments, with the lateral extent including the riparian areas and streams that occur within the 100-year floodplain or flood-prone areas encompassing a total area of approximately 84,569 hectares (208,973 acres). The critical habitat is located on a combination of Federal, State, tribal, and private lands in Inyo, Kern, Los Angeles, Riverside, Santa Barbara, San Bernardino, San Diego, and Ventura Counties in California; Clark, Lincoln, and Nye Counties in southern Nevada; Kane, San Juan, and Washington Counties in southern Utah; Alamosa, Conejos, Costilla, and La Plata Counties in southern Colorado; Apache, Cochise, Gila, Graham, Greenlee, La Paz, Maricopa, Mohave, Pima, Pinal, Santa Cruz, and Yavapai Counties in Arizona; and Catron, Grant, Hidalgo, Mora, Rio Arriba, Socorro, Taos, and Valencia Counties in New Mexico. The effect of this regulation is to conserve the flycatcher's habitat under the Endangered Species Act.

  14. The Acadian Forest Ecosystem Research Program: an example of natural disturbance-based silviculture in the northeast

    Treesearch

    Mike R. Saunders; Robert S. Seymour; Robert G. Wagner

    2014-01-01

    The Acadian Forest Ecosystem Research Program (AFERP) began in 1994 as one of the nation's first trials of natural disturbance-based silviculture. The study tests the ecological impacts of two versions of expanding-gap silvicultural systems that are designed to emulate the spatial extent and frequency of natural disturbances in northeastern North America. The...

  15. A new species of Muscicapa flycatcher from Sulawesi, Indonesia.

    PubMed

    Harris, J Berton C; Rasmussen, Pamela C; Yong, Ding Li; Prawiradilaga, Dewi M; Putra, Dadang Dwi; Round, Philip D; Rheindt, Frank E

    2014-01-01

    The Indonesian island of Sulawesi, a globally important hotspot of avian endemism, has been relatively poorly studied ornithologically, to the extent that several new bird species from the region have been described to science only recently, and others have been observed and photographed, but never before collected or named to science. One of these is a new species of Muscicapa flycatcher that has been observed on several occasions since 1997. We collected two specimens in Central Sulawesi in 2012, and based on a combination of morphological, vocal and genetic characters, we describe the new species herein, more than 15 years after the first observations. The new species is superficially similar to the highly migratory, boreal-breeding Gray-streaked Flycatcher Muscicapa griseisticta, which winters in Sulawesi; however, the new species differs strongly from M. griseisticta in several morphological characters, song, and mtDNA. Based on mtDNA, the new species is only distantly related to M. griseisticta, instead being a member of the M. dauurica clade. The new species is evidently widely distributed in lowland and submontane forest throughout Sulawesi. This wide distribution coupled with the species' apparent tolerance of disturbed habitats suggests it is not currently threatened with extinction.

  16. Long-term research on classical silvicultural approaches in the Acadian Forest: Penobscot Experimental Forest Part I

    Treesearch

    John C. Brissette; Michael R. Saunders; Laura S. Kenefic; Paul E. Sendak

    2006-01-01

    The most comprehensive study of stand dynamics in the Acadian Forest Region is an experiment by the USDA Forest Service at the Penobscot Experimental Forest (PEF) in Maine. It was established from 1952-1957 to study changes in structure, composition, and productivity from an array of silvicultural treatments. Ingrowth, accretion, and mortality of individual trees (!Y0....

  17. Status and migration of the Southwestern willow flycatcher in New Mexico

    Treesearch

    Deborah M. Finch; Jeffrey F. Kelly

    1999-01-01

    In the Southwestern United States, recent degradation of riparian habitats has been linked to decline of the Southwestern subspecies of the Willow Flycatcher. During a 2-year banding effort, migration patterns and bird fat content were analyzed. Recommendations for managers, and outlines for conservation plans, are included.

  18. Ecological genomics predicts climate vulnerability in an endangered southwestern songbird.

    PubMed

    Ruegg, Kristen; Bay, Rachael A; Anderson, Eric C; Saracco, James F; Harrigan, Ryan J; Whitfield, Mary; Paxton, Eben H; Smith, Thomas B

    2018-05-09

    Few regions have been more severely impacted by climate change in the USA than the Desert Southwest. Here, we use ecological genomics to assess the potential for adaptation to rising global temperatures in a widespread songbird, the willow flycatcher (Empidonax traillii), and find the endangered desert southwestern subspecies (E. t. extimus) most vulnerable to future climate change. Highly significant correlations between present abundance and estimates of genomic vulnerability - the mismatch between current and predicted future genotype-environment relationships - indicate small, fragmented populations of the southwestern willow flycatcher will have to adapt most to keep pace with climate change. Links between climate-associated genotypes and genes important to thermal tolerance in birds provide a potential mechanism for adaptation to temperature extremes. Our results demonstrate that the incorporation of genotype-environment relationships into landscape-scale models of climate vulnerability can facilitate more precise predictions of climate impacts and help guide conservation in threatened and endangered groups. © 2018 John Wiley & Sons Ltd/CNRS.

  19. A New Species of Muscicapa Flycatcher from Sulawesi, Indonesia

    PubMed Central

    Yong, Ding Li; Prawiradilaga, Dewi M.; Putra, Dadang Dwi; Round, Philip D.; Rheindt, Frank E.

    2014-01-01

    The Indonesian island of Sulawesi, a globally important hotspot of avian endemism, has been relatively poorly studied ornithologically, to the extent that several new bird species from the region have been described to science only recently, and others have been observed and photographed, but never before collected or named to science. One of these is a new species of Muscicapa flycatcher that has been observed on several occasions since 1997. We collected two specimens in Central Sulawesi in 2012, and based on a combination of morphological, vocal and genetic characters, we describe the new species herein, more than 15 years after the first observations. The new species is superficially similar to the highly migratory, boreal-breeding Gray-streaked Flycatcher Muscicapa griseisticta, which winters in Sulawesi; however, the new species differs strongly from M. griseisticta in several morphological characters, song, and mtDNA. Based on mtDNA, the new species is only distantly related to M. griseisticta, instead being a member of the M. dauurica clade. The new species is evidently widely distributed in lowland and submontane forest throughout Sulawesi. This wide distribution coupled with the species' apparent tolerance of disturbed habitats suggests it is not currently threatened with extinction. PMID:25419968

  20. Variation in habitat use by juvenile Acadian redfish, Sebastes fasciatus

    USGS Publications Warehouse

    Auster, P.J.; Lindholm, J.; Valentine, P.C.

    2003-01-01

    A basic paradigm in behavioral ecology is that organisms expand their distribution as preferred sites become saturated with individuals that reduce the availability of resources (e.g., shelter, prey) on a per capita basis. Previous fish community studies at Stellwagen Bank National Marine Sanctuary have shown that juvenile Acadian redfish Sebastes fasciatus (20 cm TL) also occurred in dense cerianthid habitats. Two explanations for these distributions can be advanced. The simplest is that redfish use both boulder and cerianthid habitats on an encounter basis, regardless of habitat saturation or predation pressure. Alternatively, boulder reefs serve as recruitment habitats and cerianthid habitats serve as a conduit for redfish moving away from saturated boulder reef sites, essentially serving as elements of a 'redfish pump'.

  1. Detrital zircon geochronology of pre- and syncollisional strata, Acadian orogen, Maine Appalachians

    USGS Publications Warehouse

    Bradley, Dwight C.; O'Sullivan, Paul B.

    2017-01-01

    The Central Maine Basin is the largest expanse of deep-marine, Upper Ordovician to Devonian metasedimentary rocks in the New England Appalachians, and is a key to the tectonics of the Acadian Orogeny. Detrital zircon ages are reported from two groups of strata: (1) the Quimby, Rangeley, Perry Mountain and Smalls Falls Formations, which were derived from inboard, northwesterly sources and are supposedly older; and (2) the Madrid, Carrabassett and Littleton Formations, which were derived from outboard, easterly sources and are supposedly younger. Deep-water deposition prevailed throughout, with the provenance shift inferred to mark the onset of foredeep deposition and orogeny. The detrital zircon age distribution of a composite of the inboard-derived units shows maxima at 988 and 429 Ma; a composite from the outboard-derived units shows maxima at 1324, 1141, 957, 628, and 437 Ma. The inboard-derived units have a greater proportion of zircons between 450 and 400 Ma. Three samples from the inboard-derived group have youngest age maxima that are significantly younger than the nominal depositional ages. The outboard-derived group does not share this problem. These results are consistent with the hypothesised provenance shift, but they signal potential problems with the established stratigraphy, structure, and (or) regional mapping. Shallow-marine deposits of the Silurian to Devonian Ripogenus Formation, from northwest of the Central Maine Basin, yielded detrital zircons featuring a single age maximum at 441 Ma. These zircons were likely derived from a nearby magmatic arc now concealed by younger strata. Detrital zircons from the Tarratine Formation, part of the Acadian foreland-basin succession in this strike belt, shows age maxima at 1615, 980 and 429 Ma. These results are consistent with three episodes of zircon recycling beginning with the deposition of inboard-derived strata of the Central Maine Basin, which were shed from post-Taconic highlands located to the

  2. Breeding birds in managed forests on public conservation lands in the Mississippi Alluvial Valley

    USGS Publications Warehouse

    Twedt, Daniel J.; Wilson, R. Randy

    2017-01-01

    Managers of public conservation lands in the Mississippi Alluvial Valley have implemented forest management strategies to improve bottomland hardwood habitat for target wildlife species. Through implementation of various silvicultural practices, forest managers have sought to attain forest structural conditions (e.g., canopy cover, basal area, etc.) within values postulated to benefit wildlife. We evaluated data from point count surveys of breeding birds on 180 silviculturally treated stands (1049 counts) that ranged from 1 to 20 years post-treatment and 134 control stands (676 counts) that had not been harvested for >20 years. Birds detected during 10-min counts were recorded within four distance classes and three time intervals. Avian diversity was greater on treated stands than on unharvested stands. Of 42 commonly detected species, six species including Prothonotary Warbler (Prothonotaria citrea) and Acadian Flycatcher (Empidonax virescens) were indicative of control stands. Similarly, six species including Indigo Bunting (Passerina cyanea) and Yellow-breasted Chat (Icteria virens) were indicative of treated stands. Using a removal model to assess probability of detection, we evaluated occupancy of bottomland forests at two spatial scales (stands and points within occupied stands). Wildlife-forestry treatment improved predictive models of species occupancy for 18 species. We found years post treatment (range = 1–20), total basal area, and overstory canopy were important species-specific predictors of occupancy, whereas variability in basal area was not. In addition, we used a removal model to estimate species-specific probability of availability for detection, and a distance model to estimate effective detection radius. We used these two estimated parameters to derive species densities and 95% confidence intervals for treated and unharvested stands. Avian densities differed between treated and control stands for 16 species, but only Common Yellowthroat

  3. A removal model for estimating detection probabilities from point-count surveys

    USGS Publications Warehouse

    Farnsworth, G.L.; Pollock, K.H.; Nichols, J.D.; Simons, T.R.; Hines, J.E.; Sauer, J.R.

    2002-01-01

    Use of point-count surveys is a popular method for collecting data on abundance and distribution of birds. However, analyses of such data often ignore potential differences in detection probability. We adapted a removal model to directly estimate detection probability during point-count surveys. The model assumes that singing frequency is a major factor influencing probability of detection when birds are surveyed using point counts. This may be appropriate for surveys in which most detections are by sound. The model requires counts to be divided into several time intervals. Point counts are often conducted for 10 min, where the number of birds recorded is divided into those first observed in the first 3 min, the subsequent 2 min, and the last 5 min. We developed a maximum-likelihood estimator for the detectability of birds recorded during counts divided into those intervals. This technique can easily be adapted to point counts divided into intervals of any length. We applied this method to unlimited-radius counts conducted in Great Smoky Mountains National Park. We used model selection criteria to identify whether detection probabilities varied among species, throughout the morning, throughout the season, and among different observers. We found differences in detection probability among species. Species that sing frequently such as Winter Wren (Troglodytes troglodytes) and Acadian Flycatcher (Empidonax virescens) had high detection probabilities (∼90%) and species that call infrequently such as Pileated Woodpecker (Dryocopus pileatus) had low detection probability (36%). We also found detection probabilities varied with the time of day for some species (e.g. thrushes) and between observers for other species. We used the same approach to estimate detection probability and density for a subset of the observations with limited-radius point counts.

  4. DNA metabarcoding of nestling feces reveals provisioning of aquatic prey and resource partitioning among Neotropical migratory songbirds in a riparian habitat.

    PubMed

    Trevelline, Brian K; Nuttle, Tim; Hoenig, Brandon D; Brouwer, Nathan L; Porter, Brady A; Latta, Steven C

    2018-05-01

    Riparian habitats are characterized by substantial flows of emergent aquatic insects that cross the stream-forest interface and provide an important source of prey for insectivorous birds. The increased availability of prey arising from aquatic subsidies attracts high densities of Neotropical migratory songbirds that are thought to exploit emergent aquatic insects as a nestling food resource; however, the prey preferences and diets of birds in these communities are only broadly understood. In this study, we utilized DNA metabarcoding to investigate the extent to which three syntopic species of migratory songbirds-Acadian Flycatcher, Louisiana Waterthrush, and Wood Thrush-breeding in Appalachian riparian habitats (Pennsylvania, USA) exploit and partition aquatic prey subsidies as a nestling food resource. Despite substantial differences in adult foraging strategies, nearly every nestling in this study consumed aquatic taxa, suggesting that aquatic subsidies are an important prey resource for Neotropical migrants nesting in riparian habitats. While our results revealed significant interspecific dietary niche divergence, the diets of Acadian Flycatcher and Wood Thrush nestlings were strikingly similar and exhibited significantly more overlap than expected. These results suggest that the dietary niches of Neotropical migrants with divergent foraging strategies may converge due to the opportunistic provisioning of non-limiting prey resources in riparian habitats. In addition to providing the first application of DNA metabarcoding to investigate diet in a community of Neotropical migrants, this study emphasizes the importance of aquatic subsidies in supporting breeding songbirds and improves our understanding of how anthropogenic disturbances to riparian habitats may negatively impact long-term avian conservation.

  5. African departure rather than migration speed determines variation in spring arrival in pied flycatchers.

    PubMed

    Ouwehand, Janne; Both, Christiaan

    2017-01-01

    Properly timed spring migration enhances reproduction and survival. Climate change requires organisms to respond to changes such as advanced spring phenology. Pied flycatchers Ficedula hypoleuca have become a model species to study such phenological adaptations of long-distance migratory songbirds to climate change, but data on individuals' time schedules outside the breeding season are still lacking. Using light-level geolocators, we studied variation in migration schedules across the year in a pied flycatcher population in the Netherlands, which sheds light on the ability for individual adjustments in spring arrival timing to track environmental changes at their breeding grounds. We show that variation in arrival dates to breeding sites in 2014 was caused by variation in departure date from sub-Saharan Africa and not by environmental conditions encountered en route. Spring migration duration was short for all individuals, on average 2 weeks. Males migrated ahead of females in spring, while migration schedules in autumn were flexibly adjusted according to breeding duties. Individuals were therefore not consistently early or late throughout the year. In fast migrants like our Dutch pied flycatchers, advancement of arrival to climate change likely requires changes in spring departure dates. Adaptation for earlier arrival may be slowed down by harsh circumstances in winter, or years with high costs associated with early migration. © 2016 The Authors. Journal of Animal Ecology published by John Wiley & Sons Ltd on behalf of British Ecological Society.

  6. Niltavinae, a new taxon of Old World flycatchers (Aves: Muscicapidae).

    PubMed

    Sangster, George; Alström, Per; Forsmark, Emma; Olsson, Urban

    2016-11-23

    The relationships among Old World chats and flycatchers (Muscicapidae) have recently been clarified in two independent molecular phylogenetic studies (Sangster et al. 2010, Zuccon & Ericson 2010). Both studies recovered a well-supported clade of predominantly blue flycatchers of the genera Niltava, Cyornis, Eumyias and Cyanoptila, some species traditionally included in the genus Rhinomyias and one species traditionally included in Ficedula (F. monileger). The family-group name Niltavinae Sangster, Alström, Forsmark & Olsson, 2010, was introduced for this clade (Sangster et al. 2010). Unfortunately, our description of the new taxon Niltavinae did not include a diagnosis (sensu ICZN article 13.1.1) and thus inadvertently made this name unavailable for nomenclatural purposes (Zuccon 2011). The supplementary data associated with our original paper included evidence by which the taxon Niltavinae can be diagnosed: a 1 base pair (bp) deletion in the ornithine decarboxylase gene, as shown in Figure S1 in the online version (doi:10.1016/j.ympev.2010.07.008), but this was not mentioned in the printed version of our paper. Given that Niltavinae Sangster, Alström, Forsmark & Olsson, 2010 represents a nomen nudum, and the clade for which this name was intended thus remains unnamed, we here provide the following description.

  7. Water Quality in the Acadian-Pontchartrain Drainages; Louisiana and Mississippi, 1999-2001

    USGS Publications Warehouse

    Demcheck, Dennis K.; Tollett, Roland W.; Mize, Scott V.; Skrobialowski, Stanley C.; Fendick, Robert B.; Swarzenski, Christopher M.; Porter, Stephen

    2004-01-01

    This report contains the major findings of a 1999?2001 assessment of water quality in the Acadian-Pontchartrain Drainages Study Unit. It is one of a series of reports by the National Water-Quality Assessment (NAWQA) Program that present major findings in 51 major river basins and aquifer systems across the Nation. In these reports, water quality is discussed in terms of local, State, and regional issues. Conditions in a particular basin or aquifer system are compared to conditions found elsewhere and to selected national benchmarks, such as those for drinking-water quality and the protection of aquatic organisms. This report is intended for individuals working with water-resource issues in Federal, State, or local agencies, universities, public interest groups, or in the private sector. The information will be useful in addressing a number of current issues, such as the effects of agricultural and urban land use on water quality, human health, drinking water, source-water protection, hypoxia and excessive growth of algae and plants, pesticide registration, and monitoring and sampling strategies. This report also is for individuals who wish to know more about the quality of streams and ground water in areas near where they live, and how that water quality compares to other areas across the Nation. The water-quality conditions in the Acadian-Pontchartrain Drainages Study Unit summarized in this report are discussed in detail in other reports that can be accessed from (http://la.water.usgs.gov/nawqa/default.htm). Detailed technical information, data and analyses, collection and analytical methodology, models, graphs, and maps that support the findings presented in this report in addition to other reports in this series from other basins can be accessed from the national NAWQA Web site (http://water.usgs.gov/nawqa).

  8. Myiarchus flycatchers are the primary seed dispersers of Bursera longipes in a Mexican dry forest

    PubMed Central

    Almazán-Núñez, R. Carlos; Eguiarte, Luis E.; Arizmendi, María del Coro

    2016-01-01

    We evaluated the seed dispersal of Bursera longipes by birds along a successional gradient of tropical dry forest (TDF) in southwestern Mexico. B. longipes is an endemic tree to the TDF in the Balsas basin. The relative abundance of frugivorous birds, their frequency of visits to B. longipes and the number of removed fruits were recorded at three study sites with different stages of forest succession (early, intermediate and mature) characterized by distinct floristic and structural elements. Flycatchers of the Myiarchus and Tyrannus genera removed the majority of fruits at each site. Overall, visits to B. longipes were less frequent at the early successional site. Birds that function as legitimate dispersers by consuming whole seeds and regurgitating or defecating intact seeds in the process also remove the pseudoaril from seeds, thereby facilitating the germination process. The highest germination percentages were recorded for seeds that passed through the digestive system of two migratory flycatchers: M. cinerascens and M. nutingii. Perch plants, mainly composed of legumes (e.g., Eysenhardtia polystachya, Acacia cochliacantha, Calliandra eryophylla, Mimosa polyantha), serve also as nurse plants since the number of young individuals recruited from B. longipes was higher under these than expected by chance. This study shows that Myiarchus flycatchers are the most efficient seed dispersers of B. longipes across all successional stages. This suggests a close mutualistic relationship derived from adaptive processes and local specializations throughout the distribution of both taxa, as supported by the geographic mosaic theory of coevolution. PMID:27326382

  9. An USH2A founder mutation is the major cause of Usher syndrome type 2 in Canadians of French origin and confirms common roots of Quebecois and Acadians.

    PubMed

    Ebermann, Inga; Koenekoop, Robert K; Lopez, Irma; Bou-Khzam, Lara; Pigeon, Renée; Bolz, Hanno J

    2009-01-01

    Congenital hearing loss affects approximately one child in 1000. About 10% of the deaf population have Usher syndrome (USH). In USH, hearing loss is complicated by retinal degeneration with onset in the first (USH1) or second (USH2) decade. In most populations, diagnostic testing is hampered by a multitude of mutations in nine genes. We have recently shown that in French Canadians from Quebec, USH1 largely results from a single USH1C founder mutation, c.216G>A ('Acadian allele'). The genetic basis of USH2 in Canadians of French descent, however, has remained elusive. Here, we have investigated nine USH2 families from Quebec and New Brunswick (the former Acadia) by haplotype analyses of the USH2A locus and sequencing of the three known USH2 genes. Seven USH2A mutations were identified in eight patients. One of them, c.4338_4339delCT, accounts for 10 out of 18 disease alleles (55.6%). This mutation has previously been reported in an Acadian USH2 family, and it was found in homozygous state in the three Acadians of our sample. As in the case of c.216G>A (USH1C), a common haplotype is associated with c.4338_4339delCT. With a limited number of molecular tests, it will now be possible in these populations to estimate whether children with congenital hearing impairment of different degrees will develop retinal disease - with important clinical and therapeutic implications. USH2 is the second example that reveals a significant genetic overlap between Quebecois and Acadians: in contrast to current understanding, other genetic disorders present in both populations are likely based on common founder mutations as well.

  10. An USH2A founder mutation is the major cause of Usher syndrome type 2 in Canadians of French origin and confirms common roots of Quebecois and Acadians

    PubMed Central

    Ebermann, Inga; Koenekoop, Robert K; Lopez, Irma; Bou-Khzam, Lara; Pigeon, Renée; Bolz, Hanno J

    2009-01-01

    Congenital hearing loss affects approximately one child in 1000. About 10% of the deaf population have Usher syndrome (USH). In USH, hearing loss is complicated by retinal degeneration with onset in the first (USH1) or second (USH2) decade. In most populations, diagnostic testing is hampered by a multitude of mutations in nine genes. We have recently shown that in French Canadians from Quebec, USH1 largely results from a single USH1C founder mutation, c.216G>A (‘Acadian allele'). The genetic basis of USH2 in Canadians of French descent, however, has remained elusive. Here, we have investigated nine USH2 families from Quebec and New Brunswick (the former Acadia) by haplotype analyses of the USH2A locus and sequencing of the three known USH2 genes. Seven USH2A mutations were identified in eight patients. One of them, c.4338_4339delCT, accounts for 10 out of 18 disease alleles (55.6%). This mutation has previously been reported in an Acadian USH2 family, and it was found in homozygous state in the three Acadians of our sample. As in the case of c.216G>A (USH1C), a common haplotype is associated with c.4338_4339delCT. With a limited number of molecular tests, it will now be possible in these populations to estimate whether children with congenital hearing impairment of different degrees will develop retinal disease – with important clinical and therapeutic implications. USH2 is the second example that reveals a significant genetic overlap between Quebecois and Acadians: in contrast to current understanding, other genetic disorders present in both populations are likely based on common founder mutations as well. PMID:18665195

  11. Bayesian Inference of Allele-Specific Gene Expression Indicates Abundant Cis-Regulatory Variation in Natural Flycatcher Populations

    PubMed Central

    Wang, Mi

    2017-01-01

    Abstract Polymorphism in cis-regulatory sequences can lead to different levels of expression for the two alleles of a gene, providing a starting point for the evolution of gene expression. Little is known about the genome-wide abundance of genetic variation in gene regulation in natural populations but analysis of allele-specific expression (ASE) provides a means for investigating such variation. We performed RNA-seq of multiple tissues from population samples of two closely related flycatcher species and developed a Bayesian algorithm that maximizes data usage by borrowing information from the whole data set and combines several SNPs per transcript to detect ASE. Of 2,576 transcripts analyzed in collared flycatcher, ASE was detected in 185 (7.2%) and a similar frequency was seen in the pied flycatcher. Transcripts with statistically significant ASE commonly showed the major allele in >90% of the reads, reflecting that power was highest when expression was heavily biased toward one of the alleles. This would suggest that the observed frequencies of ASE likely are underestimates. The proportion of ASE transcripts varied among tissues, being lowest in testis and highest in muscle. Individuals often showed ASE of particular transcripts in more than one tissue (73.4%), consistent with a genetic basis for regulation of gene expression. The results suggest that genetic variation in regulatory sequences commonly affects gene expression in natural populations and that it provides a seedbed for phenotypic evolution via divergence in gene expression. PMID:28453623

  12. What Do Owls, Salamanders, Flycatchers and Cuckoos Have In Common?

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Musgrave, Maria A.

    This is an article from the Los Alamos Living magazine. Los Alamos National Laboratory sits on a beautiful and unique landscape that provides important protected habitat to many species, including a few that are federally-listed as threatened or endangered. These species are the Jemez Mountains Salamander, the Mexican Spotted Owl, the Southwestern Willow Flycatcher, the Yellow-billed Cuckoo, and the New Mexico Meadow Jumping Mouse. Part of the job of the Laboratory's wildlife biologists is to survey for these species each year and determine what actions need to be taken if they are found.

  13. Climate adaptation and speciation: particular focus on reproductive barriers in Ficedula flycatchers.

    PubMed

    Qvarnström, Anna; Ålund, Murielle; McFarlane, S Eryn; Sirkiä, Päivi M

    2016-01-01

    Climate adaptation is surprisingly rarely reported as a cause for the build-up of reproductive isolation between diverging populations. In this review, we summarize evidence for effects of climate adaptation on pre- and postzygotic isolation between emerging species with a particular focus on pied (Ficedula hypoleuca) and collared (Ficedula albicollis) flycatchers as a model for research on speciation. Effects of climate adaptation on prezygotic isolation or extrinsic selection against hybrids have been documented in several taxa, but the combined action of climate adaptation and sexual selection is particularly well explored in Ficedula flycatchers. There is a general lack of evidence for divergent climate adaptation causing intrinsic postzygotic isolation. However, we argue that the profound effects of divergence in climate adaptation on the whole biochemical machinery of organisms and hence many underlying genes should increase the likelihood of genetic incompatibilities arising as side effects. Fast temperature-dependent co-evolution between mitochondrial and nuclear genomes may be particularly likely to lead to hybrid sterility. Thus, how climate adaptation relates to reproductive isolation is best explored in relation to fast-evolving barriers to gene flow, while more research on later stages of divergence is needed to achieve a complete understanding of climate-driven speciation.

  14. Arthropods of native and exotic vegetation and their association with willow flycatchers and Wilson's warblers

    Treesearch

    Linda S. DeLay; Deborah M. Finch; Sandra Brantley; Richard Fagerlund; Michael D. Means; Jeffrey F. Kelly

    1999-01-01

    We compared abundance of migrating Willow Flycatchers and Wilson's Warblers to the abundance of arthropods in exotic and native vegetation at Bosque del Apache National Wildlife Refuge. We trapped arthropods using glue-boards in 1996 and 1997 in the same cottonwood, saltcedar, and willow habitats where we mist-netted birds during spring and fall migration. There...

  15. Multi-locus phylogenetic analysis of Old World chats and flycatchers reveals extensive paraphyly at family, subfamily and genus level (Aves: Muscicapidae).

    PubMed

    Sangster, George; Alström, Per; Forsmark, Emma; Olsson, Urban

    2010-10-01

    The chats and flycatchers (Muscicapidae) represent an assemblage of 275 species in 48 genera. Defining natural groups within this assemblage has been challenging because of its high diversity and a paucity of phylogenetically informative morphological characters. We assessed the phylogenetic relationships of 124 species and 34 genera of Muscicapidae, and 20 species of Turdidae, using molecular sequence data from one mitochondrial gene and three nuclear loci, in total 3240bp. Bayesian and maximum likelihood analyses yielded a well-resolved tree in which nearly all basal nodes were strongly supported. The traditionally defined Muscicapidae, Muscicapinae and Saxicolinae were paraphyletic. Four major clades are recognized in Muscicapidae: Muscicapinae, Niltavinae (new family-group name), Erithacinae and Saxicolinae. Interesting relationships recovered by this analysis include: (i) a clade comprising the 'blue' flycatcher genera Niltava, Cyornis, Cyanoptila and Eumyias and some species of Rhinomyias; (ii) the position of Erithacus rubecula in a clade of otherwise exclusively African species; (iii) a close relationship between the shortwing Heinrichia calligyna and the flycatcher Rhinomyias insignis; (iv) a sister-relationship between forktails Enicurus and whistling thrushes Myophonus; and (v) a sister relationship of Ficedula and the 'chats'Monticola, Phoenicurus, Saxicola and Oenanthe. A high number of traditionally defined genera was found to be paraphyletic or polyphyletic. Copyright 2010 Elsevier Inc. All rights reserved.

  16. Variation in the foraging behaviors of two flycatchers: associations with stage of the breeding cycle

    Treesearch

    H.F. Sakai; B.R. Noon

    1990-01-01

    The foraging characteristics of Hammond’s and Western flycatchers in northwestern California varied with different stages of the breeding cycle during the breeding seasons (early April-mid August) in 1984 and 1985. The species’ behaviors did not always vary in parallel nor were all foraging behaviors distributed equally during the breeding cycle. For example, the...

  17. PSMC analysis of effective population sizes in molecular ecology and its application to black-and-white Ficedula flycatchers.

    PubMed

    Nadachowska-Brzyska, Krystyna; Burri, Reto; Smeds, Linnéa; Ellegren, Hans

    2016-03-01

    Climatic fluctuations during the Quaternary period governed the demography of species and contributed to population differentiation and ultimately speciation. Studies of these past processes have previously been hindered by a lack of means and genetic data to model changes in effective population size (Ne ) through time. However, based on diploid genome sequences of high quality, the recently developed pairwise sequentially Markovian coalescent (PSMC) can estimate trajectories of changes in Ne over considerable time periods. We applied this approach to resequencing data from nearly 200 genomes of four species and several populations of the Ficedula species complex of black-and-white flycatchers. Ne curves of Atlas, collared, pied and semicollared flycatcher converged 1-2 million years ago (Ma) at an Ne of ≈ 200 000, likely reflecting the time when all four species last shared a common ancestor. Subsequent separate Ne trajectories are consistent with lineage splitting and speciation. All species showed evidence of population growth up until 100-200 thousand years ago (kya), followed by decline and then start of a new phase of population expansion. However, timing and amplitude of changes in Ne differed among species, and for pied flycatcher, the temporal dynamics of Ne differed between Spanish birds and central/northern European populations. This cautions against extrapolation of demographic inference between lineages and calls for adequate sampling to provide representative pictures of the coalescence process in different species or populations. We also empirically evaluate criteria for proper inference of demographic histories using PSMC and arrive at recommendations of using sequencing data with a mean genome coverage of ≥18X, a per-site filter of ≥10 reads and no more than 25% of missing data. © 2016 The Authors. Molecular Ecology Published by John Wiley & Sons Ltd.

  18. The geology of a part of Acadia and the nature of the Acadian orogeny across Central and Eastern Maine

    USGS Publications Warehouse

    Tucker, R.D.; Osberg, P.H.; Berry, H.N.

    2001-01-01

    The zone of Acadian collision between the Medial New England and Composite Avalon terranes is well preserved in Maine. A transect from northwest (Rome) to southeast (Camden) crosses the eastern part of Medial New England comprising the Central Maine basin, Liberty-Orrington thrust sheet, and Fredericton trough, and the western part of Composite Avalon, including the Graham Lake, Clarry Hill, and Clam Cove thrust sheets. U-Pb geochronology of events before, during, and after the Acadian orogeny helps elucidate the nature and distribution of tectonostrati& graphic belts in this zone and the timing of some Acadian events in the Northern Appalachians. The Central Maine basin consists of sedimentary and volcanic rocks of Middle Ordovician (???470 to ???460 Ma) age overlain with probable conformity by latest Ordovician(?) through earliest Devonian marine rift and flysch sedimentary rocks; these are intruded by weakly to undeformed plutonic rocks of Early and Middle Devonian age (???399??378 Ma). The Fredericton trough consists of Early Silurian gray pelite and sandstone to earliest Late Silurian calcareous turbidite, deformed and variably metamorphosed prior to the emplacement of Late Silurian (???422 Ma) and Early to Late Devonian (???418 to ???368 Ma) plutons. The Liberty-Orrington thrust sheet consists of Cambrian(?)-Ordovician (>???474 to ???469 Ma and younger) clastic sedimentary and volcanic rocks intruded by highly deformed Late Silurian (???424 to ???422 Ma) and Devonian (???418 to ???389 Ma) plutons, possibly metamorphosed in Late Silurian time (prior to ???417 Ma), and metamorphosed to amphibolite facies in Early to Middle Devonian time (???400 to ???381 Ma). The Graham Lake thrust sheet contains possible Precambrian rocks, Cambrian sedimentary rocks with a volcanic unit dated at ???503 Ma, and Ordovician rocks with possible Caradocian Old World fossils, metamor& phosed and deformed in Silurian time and intruded by mildly to undeformed Late Silurian (???421 Ma

  19. Female ornamentation and territorial conflicts in collared flycatchers ( Ficedula albicollis)

    NASA Astrophysics Data System (ADS)

    Hegyi, Gergely; Garamszegi, László Zsolt; Eens, Marcel; Török, János

    2008-10-01

    Female ornaments in species with conventional sex roles often indicate individual quality, but the evolutionary forces maintaining them are less clear. Sexual competition for breeding opportunities may represent an important role for female signals, especially in polygynous species, but there is little experimental evidence for this. The wing patch size (WPS) of female collared flycatchers indicates age and body condition and predicts social mating patterns. We challenged nest-building females with decoy females of varying WPS and found that the aggressive response of residents increased with decoy WPS, suggesting a role for this female ornament in territorial competition. Our results explain why female WPS predicts territorial distances when mated to a polygynous male and indicate that the role of WPS in female competitive interactions is similar to that in males of the same population.

  20. Using Molecular Genetic Markers to Resolve a Subspecies Boundary: The Northern Boundary of the Southwestern Willow Flycatcher in the Four-Corner States

    USGS Publications Warehouse

    Paxton, Eben H.; Sogge, Mark K.; Theimer, Tad C.; Girard, Jessica; Keim, Paul

    2008-01-01

    *Executive Summary* The northern boundary of the endangered Southwestern Willow Flycatcher (Empidonax traillii extimus) is currently approximated as running through southern Colorado and Utah, but the exact placement is uncertain because this subspecies shares a border with the more northern and non-endangered E. t. adastus. To help resolve this issue, we evaluated the geographic distribution of mitochondrial and nuclear DNA by sampling breeding sites across the four-corner states (Arizona, Colorado, New Mexico, and Utah). We found that breeding sites clustered into two major groups generally consistent with the currently designated boundary, with the exception of three sites situated along the current boundary. However, delineating a precise boundary that would separate the two subspecies is made difficult because (1) we found evidence for a region of intergradation along the boundary area, suggesting the boundary is not discreet, and (2) the boundary region is sparsely populated, with too few extant breeding populations to precisely locate a boundary. The boundary region encompasses an area where elevation changes markedly over relatively short distances, with low elevation deserts to the south and more mesic, higher elevation habitats to the north. We hypothesized that latitudinal and elevational differences and their concomitant ecological effects could form an ecological barrier that inhibited gene flow between the subspecies, forming the basis for the subspecies boundary. We modeled changes in geographic patterns of genetic markers as a function of latitude and elevation finding significant support for this relationship. The model was brought into a GIS environment to create multiple subspecies boundaries, with the strength of each predicted boundary evaluated on the basis of how much genetic variation it explained. The candidate boundary that accounted for the most genetic variation was situated generally near the currently recognized subspecies boundary

  1. Gene flow during glacial habitat shifts facilitates character displacement in a Neotropical flycatcher radiation.

    PubMed

    Chattopadhyay, Balaji; Garg, Kritika M; Gwee, Chyi Yin; Edwards, Scott V; Rheindt, Frank E

    2017-09-01

    Pleistocene climatic fluctuations are known to be an engine of biotic diversification at higher latitudes, but their impact on highly diverse tropical areas such as the Andes remains less well-documented. Specifically, while periods of global cooling may have led to fragmentation and differentiation at colder latitudes, they may - at the same time - have led to connectivity among insular patches of montane tropical habitat with unknown consequences on diversification. In the present study we utilized ~5.5 kb of DNA sequence data from eight nuclear loci and one mitochondrial gene alongside diagnostic morphological and bioacoustic markers to test the effects of Pleistocene climatic fluctuations on diversification in a complex of Andean tyrant-flycatchers of the genus Elaenia. Population genetic and phylogenetic approaches coupled with coalescent simulations demonstrated disparate levels of gene flow between the taxon chilensis and two parapatric Elaenia taxa predominantly during the last glacial period but not thereafter, possibly on account of downward shifts of montane forest habitat linking the populations of adjacent ridges. Additionally, morphological and bioacoustic analyses revealed a distinct pattern of character displacement in coloration and vocal traits between the two sympatric taxa albiceps and pallatangae, which were characterized by a lack of gene flow. Our study demonstrates that global periods of cooling are likely to have facilitated gene flow among Andean montane Elaenia flycatchers that are more isolated from one another during warm interglacial periods such as the present era. We also identify a hitherto overlooked case of plumage and vocal character displacement, underpinning the complexities of gene flow patterns caused by Pleistocene climate change across the Andes.

  2. Changing climate and the phenological response of great tit and collared flycatcher populations in floodplain forest ecosystems in Central Europe.

    PubMed

    Bauer, Zdenek; Trnka, Miroslav; Bauerová, Jana; Mozný, Martin; Stepánek, Petr; Bartosová, Lenka; Zalud, Zdenek

    2010-01-01

    This study is based on 47 years of observations (1961-2007) on two common bird species, the Great Tit (Parus major) and the Collared Flycatcher (Ficedula albicollis), and a dominant tree species in their habitat, the English Oak (Quercus robur). The study took place at four research sites in the Czech Republic located in full-grown, multi-aged floodplain forests with no forestry management. An increase in air temperature over the evaluated period clearly influenced the length of phenological phases. The full foliage date of English Oak has advanced by 8.7 days during the past 47 years. Great Tit and Collared Flycatcher populations have reacted to the changing climate in the same way, with first laying date and mean laying date advancing by between 6.0 and 9.0 days. In all cases, the trends are highly significant and consistent over all sites. Despite the ongoing shift in phenological stages toward the beginning of the year, the change does not appear to have led to mistiming in the trophic food chain. Overall, this study shows almost identical rates of change in egg laying dates for both bird species in all the floodplain forests studied, and these trends are coherent with those of English Oak and peak herbivorous caterpillar activity.

  3. Problems in separating species with similar habits and vocalizations

    USGS Publications Warehouse

    Robbins, C.S.; Stallcup, R.W.; Ralph, C. John; Scott, J. Michael

    1981-01-01

    The possibilities for species misidentification based on vocalization or habitat association are high. However, the magnitude of the errors actually perpetrated is generally within an acceptable range in most types of bird survey work. Examples of problems discussed are: congeners that are similar in appearance or in song (such as Chimney and Vaux's Swifts, Chaetura pelagica, C. vauxi; Hammond's, Dusky and Gray Flycatchers, Empidonax hammondii, E. oberholseri, E. wrightii; Willow and Alder Flycatchers, E. traillii, E. alnorum; Common and Fish Crows, Corvus brachyrhynchos, C. ossifragus); birds that are misidentified because they are not expected by the observer (House Finches, Carpodacus mexicanus, invading new areas of eastern U.S.); birds that imitate other species (especially Starling, Sturnus vulgaris, and Mockingbird, Mimus polyglottos); birds in mixed flocks; birds with geographic differences in vocalizations (Solitary Vireo, Vireo solitarius); woodpeckers that are only heard drumming; and nests or eggs that are misidentified. Equally serious problems are the errors resulting from undetected species and from careless recording or failure to check manuscripts against original data. The quality of published count work can be improved considerably by (1) recognizing the problems that exist, (2) standardizing techniques for dealing with situations where not all birds can be identified, and (3) routinely applying all appropriate safeguards such as verification by mist netting and measuring, photography, tape recording or playback, additional observations, and careful verification of all entries in the final manuscript.

  4. The Mouse-colored Tyrannulet (Phaeomyias murina) is a species complex that includes the Cocos Flycatcher (Nesotriccus ridgwayi), an island form that underwent a population bottleneck.

    PubMed

    Zucker, Marc R; Harvey, Michael G; Oswald, Jessica A; Cuervo, Andrés; Derryberry, Elizabeth; Brumfield, Robb T

    2016-08-01

    Simultaneous examination of evolutionary history in island forms and closely related mainland relatives can provide reciprocal insight into the evolution of island and mainland faunas. The Cocos Flycatcher (Nesotriccus ridgwayi) is a small tyrant flycatcher (Tyrannidae) endemic to Cocos Island, an oceanic island in the eastern Pacific Ocean. We first established its close relationship to the mainland species Mouse-colored Tyrannulet (Phaeomyias murina) using a phylogeny from genome-wide ultraconserved elements and exons. We then used mitochondrial DNA to explore the relationships between Nesotriccus and Phaeomyias populations from across its distribution in Central and South America. We found that Nesotriccus is nested within the Phaeomyias evolutionary tree, and that Phaeomyias represents a complex of at least four evolutionarily distinct species that differ in plumage, voice, and habitat association. Nesotriccus underwent a population bottleneck subsequent to its divergence from Central American and northern South American Phaeomyias populations in the middle Pleistocene. The 46 UCE loci containing alleles that are fixed between the two species are widely distributed across the genome, which suggests that selective or neutral processes responsible for divergence have occurred genome-wide. Overall, our simultaneous examination of Phaeomyias and Nesotriccus revealed divergent levels of genetic diversity and evolutionary histories between island and mainland forms. Copyright © 2016 Elsevier Inc. All rights reserved.

  5. Contrasting patterns of nest survival and postfledging survival in ovenbirds and Acadian flycatchers in Missouri forest fragments

    Treesearch

    Julianna M. A. Jenkins; Frank R. Thompson; John Faaborg

    2016-01-01

    We can improve our ability to assess population viability and forecast population growth under different scenarios by understanding factors that limit population parameters in each stage of the annual cycle. Postfledging mortality rates may be as variable as nest survival across regions and fragmentation gradients, although factors that negatively impact nest survival...

  6. Effects of reintroduced beaver (Castor canadensis) on riparian bird community structure along the upper San Pedro River, southeastern Arizona and northern Sonora, Mexico

    USGS Publications Warehouse

    Johnson, Glenn E.; van Riper, Charles

    2014-01-01

    Chapter 1.—We measured bird abundance and richness along the upper San Pedro River in 2005 and 2006, in order to document how beavers (Castor canadensis) may act as ecosystem engineers after their reintroduction to a desert riparian area in the Southwestern United States. In areas where beavers colonized, we found higher bird abundance and richness of bird groups, such as all breeding birds, insectivorous birds, and riparian specialists, and higher relative abundance of many individual species—including several avian species of conservation concern. Chapter 2.—We conducted bird surveys in riparian areas along the upper San Pedro River in southeastern Arizona (United States) and northern Sonora (Mexico) in order to describe factors influencing bird community dynamics and the distribution and abundance of species, particularly those of conservation concern. These surveys were also used to document the effects of the ecosystem-altering activities of a recently reintroduced beavers (Castor canadensis). Chapter 3.—We reviewed Southwestern Willow Flycatcher (Empidonax traillii extimus) nest records and investigated the potential for future breeding along the upper San Pedro River in southeastern Arizona, where in July 2005 we encountered the southernmost verifiable nest attempt for the species. Continued conservation and management of the area’s riparian vegetation and surface water has potential to contribute additional breeding sites for this endangered Willow Flycatcher subspecies. Given the nest record along the upper San Pedro River and the presence of high-density breeding sites to the north, the native cottonwood-willow forests of the upper San Pedro River could become increasingly important to E. t. extimus recovery, especially considering the anticipated effect of the tamarisk leaf beetle (Diorhabda carinulata) on riparian habitat north of the region.

  7. Heavy metals and metalloids in egg contents and eggshells of passerine birds from Arizona

    USGS Publications Warehouse

    Mora, Miguel A.

    2003-01-01

    Concentrations of inorganic elements were determined in eggs of passerine birds including the endangered southwestern willow flycatcher (Empidonax traillii extimus) from four regions in Arizona. The main aim of the study was to determine the distribution of metals in egg contents and eggshells, with emphasis on the deposition of Sr in eggshells. Seventy eggs of 11 passerine species were collected at four nesting locations during 2000. Aluminum, Ba, Cr, Cu, Mn, Se, Sr, and Zn, were detected primarily in egg contents of all bird species. Arsenic, Ni, Pb, and V were detected primarily in eggshells. A proportion of most inorganic elements accumulated in the eggshell. Concentrations of Ba, Cu, Mn, Se, Sr, and Zn in egg contents and As, Ba, Cu, and V in eggshells of yellow-breasted chats (Icteria virens) were similar among locations. However, concentrations of Mn, Ni, Sr, and Zn in eggshells were significant different among locations. Except for Cu, Mn, Se, and Zn, concentrations of inorganic elements were 2–35 times greater in eggshells than in eggs. Most concentrations of metals and metalloids in eggs and eggshells of all the bird species were below levels known to affect reproduction or that have other deleterious effects. However, I found somewhat elevated concentrations of Sr in eggshells (highest mean=1505 μg/g dw, n=3) of yellow-breasted chats and willow flycatchers, and in egg contents of yellow warblers (Dendroica petechia) and song sparrows (Melospiza melodia). Whether current observed concentrations of Sr in eggshells are affecting nesting birds in Arizona remains to be determined. Strontium and other metals could be associated with lower hatching success in some areas. This study shows that a proportion of many inorganic elements accumulates in the eggshell and that the potential effects on the proper structure and functioning of the eggshell should not be ignored.

  8. Polygyny and its fitness consequences for primary and secondary female pied flycatchers

    PubMed Central

    Huk, Thomas; Winkel, Wolfgang

    2006-01-01

    In polygynous species with biparental care, the amount of paternal support often varies considerably. In the pied flycatcher (Ficedula hypoleuca), females mated with monogamous males receive more male assistance during the nestling phase than females mated with bigynous males, as the latter have to share their mates with another female. Bigynous males, however, give more support to their primary broods than to their secondary broods. Using a long-term dataset (31 years), the present study revealed that direct reproductive success, i.e. number of fledglings, was lower in females that mated with bigynous males, especially in secondary broods without male assistance, than in females that mated with monogamous males. Secondary broods with male assistance were more affected than primary broods. Female survival was independent of mating status. In primary broods, a delayed compensation for inferior direct reproductive success was found in terms of the number of grandoffspring, a phenomenon that did not occur in secondary broods. Delayed compensation in primary broods refers to indirect effects, i.e. good genes. According to the sexy son hypothesis, genetically superior (i.e. sexy) males may have sons with a higher number of broods belonging to a polygynous breeding status than do sons from broods with a monogamous father. This was indeed the case for sons descending from primary broods, but not for sons descending from secondary broods. PMID:16769641

  9. Trade-offs between sexual advertisement and immune function in the pied flycatcher (Ficedula hypoleuca).

    PubMed

    Kilpimaa, Janne; Alatalo, Rauno V; Siitari, Heli

    2004-02-07

    Good genes models of sexual selection assume that sexual advertisement is costly and thus the level of advertisement honestly reveals heritable viability. Recently it has been suggested that an important cost of sexual advertisement might be impairment of the functioning of the immune system. In this field experiment we investigated the possible trade-offs between immune function and sexual advertisement by manipulating both mating effort and activity of immune defence in male pied flycatchers. Mating effort was increased in a non-arbitrary manner by removing females from mated males during nest building. Widowed males sustained higher haematocrit levels than control males and showed higher expression of forehead patch height, suggesting that manipulation succeeded in increasing mating effort. Males that were experimentally forced to increase mating effort had reduced humoral immune responsiveness compared with control males. In addition, experimental activation of immune defence by vaccination with novel antigens reduced the expression of male ornament dimensions. To conclude, our results indicate that causality behind the trade-off between immune function and sexual advertisement may work in both directions: sexual activity suppresses immune function but immune challenge also reduces sexual advertisement.

  10. Trade-offs between sexual advertisement and immune function in the pied flycatcher (Ficedula hypoleuca).

    PubMed Central

    Kilpimaa, Janne; Alatalo, Rauno V.; Siitari, Heli

    2004-01-01

    Good genes models of sexual selection assume that sexual advertisement is costly and thus the level of advertisement honestly reveals heritable viability. Recently it has been suggested that an important cost of sexual advertisement might be impairment of the functioning of the immune system. In this field experiment we investigated the possible trade-offs between immune function and sexual advertisement by manipulating both mating effort and activity of immune defence in male pied flycatchers. Mating effort was increased in a non-arbitrary manner by removing females from mated males during nest building. Widowed males sustained higher haematocrit levels than control males and showed higher expression of forehead patch height, suggesting that manipulation succeeded in increasing mating effort. Males that were experimentally forced to increase mating effort had reduced humoral immune responsiveness compared with control males. In addition, experimental activation of immune defence by vaccination with novel antigens reduced the expression of male ornament dimensions. To conclude, our results indicate that causality behind the trade-off between immune function and sexual advertisement may work in both directions: sexual activity suppresses immune function but immune challenge also reduces sexual advertisement. PMID:15058434

  11. Low genetic variance in the duration of the incubation period in a collared flycatcher (Ficedula albicollis) population.

    PubMed

    Husby, Arild; Gustafsson, Lars; Qvarnström, Anna

    2012-01-01

    The avian incubation period is associated with high energetic costs and mortality risks suggesting that there should be strong selection to reduce the duration to the minimum required for normal offspring development. Although there is much variation in the duration of the incubation period across species, there is also variation within species. It is necessary to estimate to what extent this variation is genetically determined if we want to predict the evolutionary potential of this trait. Here we use a long-term study of collared flycatchers to examine the genetic basis of variation in incubation duration. We demonstrate limited genetic variance as reflected in the low and nonsignificant additive genetic variance, with a corresponding heritability of 0.04 and coefficient of additive genetic variance of 2.16. Any selection acting on incubation duration will therefore be inefficient. To our knowledge, this is the first time heritability of incubation duration has been estimated in a natural bird population. © 2011 by The University of Chicago.

  12. A test of the "sexy son" hypothesis: sons of polygynous collared flycatchers do not inherit their fathers' mating status.

    PubMed

    Gustafsson, Lars; Qvarnström, Anna

    2006-02-01

    According to the original "sexy son" hypothesis, a female may benefit from pairing with an already-mated male despite a reduction in fecundity because her sons inherit their father's attractiveness. We used data from a long-term study of collared flycatchers (Ficedula albicollis) collected during 24 years to test this prediction. Our results show that the sons of polygynously mated females fledged in poor condition and therefore did not inherit their father's large forehead patch (a condition-dependent display trait) or mating status. From the female's perspective, polygynous pairing resulted in fewer recruited grandchildren than did a monogamous pairing. The reproductive value of sons did not outweigh the fecundity costs of polygyny because the low paternal care reduced the attractiveness of sons. When there are long-lasting parental effects on offspring attractiveness, costs of polygyny may include the production of nonsexy sons.

  13. Analyzing spatial and temporal trends in Aboveground Biomass within the Acadian New England Forests using the complete Landsat Archive

    NASA Astrophysics Data System (ADS)

    Kilbride, J. B.; Fraver, S.; Ayrey, E.; Weiskittel, A.; Braaten, J.; Hughes, J. M.; Hayes, D. J.

    2017-12-01

    Forests within the New England states and Canadian Maritime provinces, here described as the Acadian New England (ANE) forests, have undergone substantial disturbances due to insect, fire, and anthropogenic factors. Through repeated satellite observations captures by USGS's Landsat program, 45 years of disturbance information can be incorporated into modeling efforts to better understand the spatial and temporal trends in forest above ground biomass (AGB). Using Google's Earth Engine, annual mosaics were developed for the ANE study area and then disturbance and recovery metrics were developed using the temporal segmentation algorithm VeRDET. Normalization procedures were developed to incorporate the Landsat Multispectral Scanner (MSS, 1972 - 1985) data alongside the modern era of Landsat Thematic Mapper (TM, 1984-2013), Enhanced Thematic Mapper plus (ETM+, 1999 - present), and Operational Land Imager (OLI, 2013- present) data products. This has enabled the creation of a dataset with an unprecedented spatial and temporal view of forest landscape change. Model training was performed using was the Forest Inventory Analysis (FIA) and New Brunswick Permanent Sample Plot data datasets. Modeling was performed using parametric techniques such as mixed effects models and non-parametric techniques such as k-NN imputation and generalized boosted regression. We compare the biomass estimate and model accuracy to other inventory and modeling studies produced within this study area. The spatial and temporal patterns of stock changes are analyzed against resource policy, land ownership changes, and forest management.

  14. Avian habitat relationships in pinyon-juniper woodland

    USGS Publications Warehouse

    Sedgwick, James A.

    1987-01-01

    Habitat relationships of breeding birds were examined in northwestern Colorado in pinyon-juniper (Pinus edulis-Juniperus osteosperma) woodland and in openings where most overstory trees had been knocked down by anchor chaining. Vegetation characteristics and physical habitat features were measured in 233 0.04-ha circular plots around singing males of 13 species of birds from 15 May to 15 July 1980. Thirteen-group discriminant function analysis ordinated bird species along three habitat dimensions described by (1) canopy height; (2) slope, shrub size, and shrub species diversity; and (3) percentage canopy cover, large tree density, distance from a habitat edge, litter cover, and green cover. Woodland, open-area, and intermediate edge species were clearly segregated along the first discriminant axis, and species' associations with shrubs, inclination, ground cover, and edges were revealed by the ordinations along the second and third discriminant axes. Two-group discriminant analyses comparing occupied and available plots identified additional and more specific habitat associations. For example, Hermit Thrushes (Catharus guttatus) were associated with mature forested habitats and forest interiors, Virginia's Warblers (Vermivora virginiae) favored steep, oak-covered draws, Rock Wrens (Salpinctes obsoletus) selected areas where percentage log cover and small tree density were high, and Dusky Flycatchers (Empidonax oberholseri) preferred shrubby slopes with scattered large trees near woodland edges.

  15. Evaluation de la validite d'un programme francais televise en Nouvelle-Ecosse (Evaluation of the Validity of a French Program Televised in Nova Scotia).

    ERIC Educational Resources Information Center

    Starets, Moshe

    1983-01-01

    A television series for Acadians was evaluated by the Ministry of Education according to its stated objectives of vocabulary development and strengthening of ethnic appreciation in the Acadian community. The series was found to be an effective method of both teaching Acadian culture and reinforcing students' competence in French. (MSE)

  16. 78 FR 65843 - General Provisions; Revised List of Migratory Birds

    Federal Register 2010, 2011, 2012, 2013, 2014

    2013-11-01

    ..., 2008, 2009, and 2010), on matters of taxonomy, nomenclature, and the sequence of species and other... Flycatcher, Ficedula albicilla (AOU 1982, 1983, 1998, 2006); Dark-sided Flycatcher, Muscicapa sibirica (AOU...). Mugimaki Flycatcher, Ficedula mugimaki (1). Taiga Flycatcher, Ficedula albicilla (1). Dark-sided Flycatcher...

  17. Foveolar Müller Cells of the Pied Flycatcher: Morphology and Distribution of Intermediate Filaments Regarding Cell Transparency.

    PubMed

    Zueva, Lidia; Golubeva, Tatiana; Korneeva, Elena; Makarov, Vladimir; Khmelinskii, Igor; Inyushin, Mikhail

    2016-04-01

    Specialized intermediate filaments (IFs) have critical importance for the clearness and uncommon transparency of vertebrate lens fiber cells, although the physical mechanisms involved are poorly understood. Recently, an unusual low-scattering light transport was also described in retinal Müller cells. Exploring the function of IFs in Müller cells, we have studied the morphology and distribution pattern of IFs and other cytoskeletal filaments inside the Müller cell main processes in the foveolar part of the avian (pied flycatcher) retina. We found that some IFs surrounded by globular nanoparticles (that we suggest are crystallines) are present in almost every part of the Müller cells that span the retina, including the microvilli. Unlike IFs implicated in the mechanical architecture of the cell, these IFs are not connected to any specific cellular membranes. Instead, they are organized into bundles, passing inside the cell from the endfeet to the photoreceptor, following the geometry of the processes, and repeatedly circumventing numerous obstacles. We believe that the presently reported data effectively confirm that the model of nanooptical channels built of the IFs may provide a viable explanation of Müller cell transparency.

  18. Foreland-forearc collisional granitoid and mafic magmatism caused by lower-plate lithospheric slab breakoff: The Acadian of Maine, and other orogens

    USGS Publications Warehouse

    Schoonmaker, A.; Kidd, W.S.F.; Bradley, D.C.

    2005-01-01

    During collisional convergence, failure in extension of the lithosphere of the lower plate due to slab pull will reduce the thickness or completely remove lower-plate lithosphere and cause decompression melting of the asthenospheric mantle; magmas from this source may subsequently provide enough heat for substantial partial melting of crustal rocks under or beyond the toe of the collisional accretionary system. In central Maine, United States, this type of magmatism is first apparent in the Early Devonian West Branch Volcanics and equivalent mafic volcanics, in the slightly younger voluminous mafic/silicic magmatic event of the Moxie Gabbro-Katahdin batholith and related ignimbrite volcanism, and in other Early Devonian granitic plutons. Similar lower-plate collisional sequences with mafic and related silicic magmatism probably caused by slab breakoff are seen in the Miocene-Holocene Papuan orogen, and the Hercynian-Alleghenian belt. Magmatism of this type is significant because it gives evidence in those examples of whole-lithosphere extension. We infer that normal fault systems in outer trench slopes of collisional orogens in general, and possibly those of oceanic subduction zones, may not be primarily due to flexural bending, but are also driven by whole-lithosphere extension due to slab pull. The Maine Acadian example suggests that slab failure and this type of magmatism may be promoted by pre-existing large margin-parallel faults in the lower plate. ?? 2005 Geological Society of America.

  19. Nest survival of forest birds in the Mississippi Alluvial Valley

    USGS Publications Warehouse

    Twedt, D.J.; Wilson, R.R.; Henne-Kerr, J.L.; Hamilton, R.B.

    2001-01-01

    vimns, 7%), eastern towhee (14%), indigo bunting (14%), and northern cardinal (17%) did not differ from nest success in cottonwood plantations that were coppiced from root sprouts following pulpwood harvest. Within bottomland hardwood forests, uneven-aged group-selection timber harvest reduced the combined daily nest survival of all species from 0.958 to 0.938, which reduced nest success by about 14%. Specifically, timber harvest reduced nest success of species that nest in the forest midstory and canopy, such as Acadian flycatcher (Empidonax virescens)--from 32% before harvest to 14% after harvest. Conversely, those species that nest primarily in the shrubby understory--such as northern cardinal--were not affected by timber harvest and maintained an overall nest success of about 33%. Thus, birds nesting in the understory of bottomland hardwood forests are not adversely impacted by selective timber harvest, but there is a short-term reduction in nest success for birds that nest in the canopy and midstory.

  20. Divergent patterns of age-dependence in ornamental and reproductive traits in the collared flycatcher.

    PubMed

    Evans, Simon R; Gustafsson, Lars; Sheldon, Ben C

    2011-06-01

    Sexual ornaments are predicted to honestly signal individual condition. We might therefore expect ornament expression to show a senescent decline, in parallel with late-life deterioration of other characters. Conversely, life-history theory predicts the reduced residual reproductive value of older individuals will favor increased investment in sexually attractive traits. Using a 25-year dataset of more than 5000 records of breeding collared flycatchers (Ficedula albicollis) of known age, we quantify cross-sectional patterns of age-dependence in ornamental plumage traits and report long-term declines in expression that mask highly significant positive age-dependency. We partition this population-level age-dependency into its between- and within-individual components and show expression of ornamental white plumage patches exhibits within-individual increases with age in both sexes, consistent with life-history theory. For males, ornament expression also covaries with life span, such that, within a cohort, ornamentation indicates survival. Finally, we compared longitudinal age-dependency of reproductive traits and ornamental traits in both sexes, to assess whether these two trait types exhibit similar age-dependency. These analyses revealed contrasting patterns: reproductive traits showed within-individual declines in late-life females consistent with senescence; ornamental traits showed the opposite pattern in both males and females. Hence, our results for both sexes suggest that age-dependent ornament expression is consistent with life-history models of optimal signaling and, unlike reproductive traits, proof against senescence. © 2011 The Author(s). Evolution© 2011 The Society for the Study of Evolution.

  1. Nutritional correlates and mate acquisition role of multiple sexual traits in male collared flycatchers

    NASA Astrophysics Data System (ADS)

    Hegyi, Gergely; Szöllősi, Eszter; Jenni-Eiermann, Susanne; Török, János; Eens, Marcel; Garamszegi, László Zsolt

    2010-06-01

    The information content of a sexual signal may predict its importance in a multiple signal system. Many studies have correlated sexual signal expression with the absolute levels of nutrient reserves. In contrast, the changes of nutrient reserves associated with signal expression are largely unknown in the wild due to technical limitations although they are important determinants of signal information content. We compared two visual and eight acoustic sexual traits in male collared flycatchers to see whether the nutritional correlates of expression predict the role of the signal in sexual selection. We used single point assays of plasma lipid metabolites to estimate short-term changes in nutritional state in relation to sexual trait expression during courtship. As a measure of sexual selection, we estimated the relationship with pairing latency after arrival in a 4-year dataset. Males which found a mate rapidly were characterized by large wing and forehead patches, but small song strophe complexity and small figure repertoire size. Traits more strongly related to pairing latency were also more closely related to changes in nutrient reserves. This indicates a link between signal role and information content. Small wing patches and, surprisingly, complex songs seemed to indicate poor phenotypic quality and were apparently disfavoured at mate acquisition in our population. Future studies of the information content of sexual traits, especially dynamic traits such as song, may benefit from the use of plasma metabolite profiles as non-invasive indicators of short-term changes in body condition.

  2. Geographical trends in the yolk carotenoid composition of the pied flycatcher (Ficedula hypoleuca).

    PubMed

    Eeva, Tapio; Ruuskanen, Suvi; Salminen, Juha-Pekka; Belskii, Eugen; Järvinen, Antero; Kerimov, Anvar; Korpimäki, Erkki; Krams, Indrikis; Moreno, Juan; Morosinotto, Chiara; Mänd, Raivo; Orell, Markku; Qvarnström, Anna; Siitari, Heli; Slater, Fred M; Tilgar, Vallo; Visser, Marcel E; Winkel, Wolfgang; Zang, Herwig; Laaksonen, Toni

    2011-02-01

    Carotenoids in the egg yolks of birds are considered to be important antioxidants and immune stimulants during the rapid growth of embryos. Yolk carotenoid composition is strongly affected by the carotenoid composition of the female's diet at the time of egg formation. Spatial and temporal differences in carotenoid availability may thus be reflected in yolk concentrations. To assess whether yolk carotenoid concentrations or carotenoid profiles show any large-scale geographical trends or differences among habitats, we collected yolk samples from 16 European populations of the pied flycatcher, Ficedula hypoleuca. We found that the concentrations and proportions of lutein and some other xanthophylls in the egg yolks decreased from Central Europe northwards. The most southern population (which is also the one found at the highest altitude) also showed relatively low carotenoid levels. Concentrations of β-carotene and zeaxanthin did not show any obvious geographical gradients. Egg yolks also contained proportionally more lutein and other xanthophylls in deciduous than in mixed or coniferous habitats. We suggest that latitudinal gradients in lutein and xanthophylls reflect the lower availability of lutein-rich food items in the northern F. hypoleuca populations and in montane southern populations, which start egg-laying earlier relative to tree phenology than the Central European populations. Similarly, among-habitat variation is likely to reflect the better availability of lutein-rich food in deciduous forests. Our study is the first to indicate that the concentration and profile of yolk carotenoids may show large-scale spatial variation among populations in different parts of the species' geographical range. Further studies are needed to test the fitness effects of this geographical variation.

  3. Tamarix and Diorhabda leaf beetle interactions: implications for Tamarix water use and riparian habitat

    USGS Publications Warehouse

    Nagler, Pamela; Glenn, Edward P.

    2013-01-01

    Tamarix leaf beetles (Diorhabda carinulata) have been widely released on western United States rivers to control introduced shrubs in the genus Tamarix, with the goals of saving water through removal of an assumed high water-use plant, and of improving habitat value by removing a competitor of native riparian trees. We review recent studies addressing three questions: (1) to what extent are Tamarix weakened or killed by recurrent cycles of defoliation; (2) can significant water salvage be expected from defoliation; and (3) what are the effects of defoliation on riparian ecology, particularly on avian habit? Defoliation has been patchy at many sites, and shrubs at some sites recover each year even after multiple years of defoliation. Tamarix evapotranspiration (ET) is much lower than originally assumed in estimates of potential water savings, and are the same or lower than possible replacement plants. There is concern that the endangered southwestern willow flycatcher (Empidonax trailli extimus) will be negatively affected by defoliation because the birds build nests early in the season when Tamarix is still green, but are still on their nests during the period of summer defoliation. Affected river systems will require continued monitoring and development of adaptive management practices to maintain or enhance riparian habitat values. Multiplatform remote sensing methods are playing an essential role in monitoring defoliation and rates of ET on affected river systems.

  4. Mechanical power curve measured in the wake of pied flycatchers indicates modulation of parasite power across flight speeds.

    PubMed

    Johansson, L Christoffer; Maeda, Masateru; Henningsson, Per; Hedenström, Anders

    2018-01-01

    How aerodynamic power required for animal flight varies with flight speed determines optimal speeds during foraging and migratory flight. Despite its relevance, aerodynamic power provides an elusive quantity to measure directly in animal flight. Here, we determine the aerodynamic power from wake velocity fields, measured using tomographical particle image velocimetry, of pied flycatchers flying freely in a wind tunnel. We find a shallow U-shaped power curve, which is flatter than expected by theory. Based on how the birds vary body angle with speed, we speculate that the shallow curve results from increased body drag coefficient and body frontal area at lower flight speeds. Including modulation of body drag in the model results in a more reasonable fit with data than the traditional model. From the wake structure, we also find a single starting vortex generated from the two wings during the downstroke across flight speeds (1-9 m s -1 ). This is accomplished by the arm wings interacting at the beginning of the downstroke, generating a unified starting vortex above the body of the bird. We interpret this as a mechanism resulting in a rather uniform downwash and low induced power, which can help explain the higher aerodynamic performance in birds compared with bats. © 2018 The Author(s).

  5. Out of Amazonia again and again: episodic crossing of the Andes promotes diversification in a lowland forest flycatcher

    PubMed Central

    Miller, Matthew J; Bermingham, Eldredge; Klicka, John; Escalante, Patricia; do Amaral, Fabio S. Raposo; Weir, Jason T; Winker, Kevin

    2008-01-01

    Most Neotropical lowland forest taxa occur exclusively on one side of the Andes despite the availability of appropriate habitat on both sides. Almost all molecular phylogenies and phylogenetic analyses of species assemblages (i.e. area cladograms) have supported the hypothesis that Andean uplift during the Late Pliocene created a vicariant barrier affecting lowland lineages in the region. However, a few widespread plant and animal species occurring in lowland forests on both sides of the Andes challenge the generality of this hypothesis. To understand the role of the Andes in the history of such organisms, we reconstructed the phylogeographic history of a widespread Neotropical flycatcher (Mionectes oleagineus) in the context of the other four species in the genus. A molecular phylogeny based on nuclear and mitochondrial sequences unambiguously showed an early basal split between montane and lowland Mionectes. The phylogeographic reconstruction of lowland taxa revealed a complex history, with multiple cases in which geographically proximate populations do not represent sister lineages. Specifically, three populations of M. oleagineus west of the Andes do not comprise a monophyletic clade; instead, each represents an independent lineage with origins east of the Andes. Divergence time estimates suggest that at least two cross-Andean dispersal events post-date Andean uplift. PMID:18285279

  6. A removal model for estimating detection probabilities from point-count surveys

    USGS Publications Warehouse

    Farnsworth, G.L.; Pollock, K.H.; Nichols, J.D.; Simons, T.R.; Hines, J.E.; Sauer, J.R.

    2000-01-01

    We adapted a removal model to estimate detection probability during point count surveys. The model assumes one factor influencing detection during point counts is the singing frequency of birds. This may be true for surveys recording forest songbirds when most detections are by sound. The model requires counts to be divided into several time intervals. We used time intervals of 2, 5, and 10 min to develop a maximum-likelihood estimator for the detectability of birds during such surveys. We applied this technique to data from bird surveys conducted in Great Smoky Mountains National Park. We used model selection criteria to identify whether detection probabilities varied among species, throughout the morning, throughout the season, and among different observers. The overall detection probability for all birds was 75%. We found differences in detection probability among species. Species that sing frequently such as Winter Wren and Acadian Flycatcher had high detection probabilities (about 90%) and species that call infrequently such as Pileated Woodpecker had low detection probability (36%). We also found detection probabilities varied with the time of day for some species (e.g. thrushes) and between observers for other species. This method of estimating detectability during point count surveys offers a promising new approach to using count data to address questions of the bird abundance, density, and population trends.

  7. Possible roles for corticosterone and critical size in the fledging of nestling pied flycatchers.

    PubMed

    Kern, M; Bacon, W; Long, D; Cowie, R J

    2001-01-01

    Our study was designed to see whether corticosterone (B) rises abruptly in the blood of nestling pied flycatchers (Ficedula hypoleuca) at the time they fledge, as reported recently for kestrels, and if so, why. We measured the growth and blood levels of B and selected nutrients of nestlings in broods of five, seven, and nine chicks during 1998 and 1999. In half of the broods, we clipped selected wing and tail feathers of both parents with the intention of making it more difficult for them to provide their chicks with food. We collected blood samples when the chicks were six to 10 d old (period of rapid growth) and 15 d of age or older (0-5 d before fledging). B increased substantially several days before the chicks left the nest and then declined somewhat. We found no differences in rates of growth or blood levels of B, nutrients, and hematocrit as a function of either brood size or parental handicapping. Nestlings within a day of fledging appear to have been food deprived in 1998; their glucose was significantly reduced, and B, free fatty acids, and glycerol were significantly elevated compared to levels in chicks 1-4 d younger. Such changes did not occur in 1999. Blood levels of B were significantly correlated with brood size near the day of fledging, but not earlier, in both years of the study. It was possible to predict the day on which chicks would leave the nest, using their wing length when 12 d old. These results suggest that high blood levels of B associated with food restriction and sibling competition induce chicks to fledge, provided they have reached a critical size, and that the importance of fasting, sibling competition, and B may vary from year to year.

  8. Isotopic evidence for closed-system anatexis at midcrustal levels: An example from the Acadian Appalachians of New England

    NASA Astrophysics Data System (ADS)

    Lathrop, Alison S.; Blum, Joel D.; Chamberlain, C. Page

    1994-05-01

    We have investigated the Sr and O isotope systematics of granitoid and metasedimentary samples from the Central Main Terrane (CMT) of New England. Granitoid samples were taken from interior and contact zones within the Acadian-aged (approximately 410 m.y.), synmetamorphic and syntectonic Kinsman Quartz Monzonite (KQM), which is a member of the New Hampshire Plutonic Series. Metasedimentary samples were taken from Silurian and Devonian formations hosting the KQM. Initial Sr isotope ratios (Sr(sub i) and delta O-18 values for the KQM range from 0.70799 to 0.71246 and 7.6% to 12.9%, respectively, and Sr(sub i) and delta O-18 values of the metasedimentary rocks range from 0.70770 to 0.75008 and 6.2% to 14.1%, respectively. We observe a linear and slightly positive correlation between Sr(sub i) and delta O-18 for interior KQM samples that can be duplicated by a mixing curve calculated for metasedimentary endmembers, whereas the Sr(sub i) and delta O-18 values of contact KQM samples cluster near the Sr(sub i) and delta O-18 values of the metasedimentary rocks with which they are in contact. Mixing calculations provide no evidence for a measurable primitive mantle component in either interior or contact KQM samples, and we conclude that the Sr-O isotopic composition of the KQM is most likely a reflection of isotopic heterogeneities inherited from a complex package of midcrustal metasedimentary source rocks. We propose that the KQM is the product of midcrustal partial melting that was initiated due to excess thermal energy from the decay of anomalously high concentrations of heat-producing elements in Silurian source rocks within the CMT. Because we see no isotopic evidence for a lower-crustal or mantle component in the KQM, we suggest that midcrustal anatexis may have occurred as a closed-system process, requiring no accompanying mantle-derived magma or above normal mantle heat flow.

  9. Integrated plumage colour variation in relation to body condition, reproductive investment and laying date in the collared flycatcher

    NASA Astrophysics Data System (ADS)

    Laczi, Miklós; Hegyi, Gergely; Herényi, Márton; Kiss, Dorottya; Markó, Gábor; Nagy, Gergely; Rosivall, Balázs; Szöllősi, Eszter; Török, János

    2013-10-01

    The possible integration of different sexual ornaments into a composite system, and especially the information content of such ornament complexes, is poorly investigated. Many bird species display complex plumage coloration, but whether this represents one integrated or several independent sexual traits can be unclear. Collared flycatchers ( Ficedula albicollis) display melanised and depigmented plumage areas, and the spectral features (brightness and UV chroma) of these are correlated with each other across the plumage. In a 5-year dataset of male and female plumage reflectance, we examined some of the potential information content of integrated, plumage-level colour attributes by estimating their relationships to previous and current year body condition, laying date and clutch size. Females were in better condition the year before they became darker pigmented, and males in better current year condition were also darker pigmented. Female pigment-based brightness was positively, while male structurally based brightness was negatively related to current laying date. Finally, the overall UV chroma of white plumage areas in males was positively associated with current clutch size. Our results show that higher degree of pigmentation is related to better condition, while the structural colour component is associated with some aspects of reproductive investment. These results highlight the possibility that correlated aspects of a multiple plumage ornamentation system may reflect together some aspects of individual quality, thereby functioning as a composite signal.

  10. [Visual input affects the expression of the early genes c-Fos and ZENK in auditory telencephalic centers of pied flycatcher nestlings during the acoustically-guided freezing].

    PubMed

    Korneeva, E V; Tiunova, A A; Aleksandrov, L I; Golubeva, T B; Anokhin, K V

    2014-01-01

    The present study analyzed expression of transcriptional factors c-Fos and ZENK in 9-day-old pied flycatcher nestlings' (Ficedula hypoleuca) telencephalic auditory centers (field L, caudomedial nidopallium and caudomedial mesopallium) involved in the acoustically-guided defense behavior. Species-typical alarm call was presented to the young in three groups: 1--intact group (sighted control), 2--nestlings visually deprived just before the experiment for a short time (unsighted control) 3--nestlings visually deprived right after hatching (experimental deprivation). Induction of c-Fos as well as ZENK in nestlings from the experimental deprivation group was decreased in both hemispheres as compared with intact group. In the group of unsighted control, only the decrease of c-Fos induction was observed exclusively in the right hemisphere. These findings suggest that limitation of visual input changes the population of neurons involved into the acoustically-guided behavior, the effect being dependant from the duration of deprivation.

  11. Effects of pollution on land snail abundance, size and diversity as resources for pied flycatcher, Ficedula hypoleuca.

    PubMed

    Eeva, Tapio; Rainio, Kalle; Suominen, Otso

    2010-09-01

    Passerine birds need extra calcium during their breeding for developing egg shells and proper growth of nestling skeleton. Land snails are an important calcium source for many passerines and human-induced changes in snail populations may pose a severe problem for breeding birds. We studied from the bird's viewpoint how air pollution affects the shell mass, abundance and diversity of land snail communities along a pollution gradient of a copper smelter. We sampled remnant snail shells from the nests of an insectivorous passerine, the pied flycatcher, Ficedula hypoleuca, to find out how the availability of land snails varies along the pollution gradient. The total snail shell mass increased towards the pollution source but declined abruptly in the vicinity of the smelter. This spatial variation in shell mass was evident also within a single snail species and could not be wholly explained by spatially varying snail numbers or species composition. Instead, the total shell mass was related to their shell size, individuals being largest at the moderately polluted areas. Smaller shell size suggests inferior growth of snails in the most heavily polluted area. Our study shows that pollution affects the diversity, abundance (available shell mass) and individual quality of land snails, posing reproductive problems for birds that rely on snails as calcium sources during breeding. There are probably both direct pollution-related (heavy metal and calcium levels) and indirect (habitat change) effects behind the observed changes in snail populations. Copyright 2010 Elsevier B.V. All rights reserved.

  12. Linguistic Minorities and the Multilingual Turn: Constructing Language Ownership through Affect in Cultural Production

    ERIC Educational Resources Information Center

    McLaughlin, Mireille

    2016-01-01

    The "multilingual turn" brings questions of language ownership to the forefront of debates about linguistic minority governance. Acadian minority cultural producers construct language ownership using multiple languages and targeting multilingual publics, but use ideologies of monolingualism to situate Acadian authenticity in place and…

  13. Nest site preference depends on the relative density of conspecifics and heterospecifics in wild birds.

    PubMed

    Samplonius, Jelmer M; Kromhout Van Der Meer, Iris M; Both, Christiaan

    2017-01-01

    Social learning allows animals to eavesdrop on ecologically relevant knowledge of competitors in their environment. This is especially important when selecting a habitat if individuals have relatively little personal information on habitat quality. It is known that birds can use both conspecific and heterospecific information for social learning, but little is known about the relative importance of each information type. If provided with the choice between them, we expected that animals should copy the behaviour of conspecifics, as these confer the best information for that species. We tested this hypothesis in the field for Pied Flycatchers Ficedula hypoleuca arriving at their breeding grounds to select a nest box for breeding. We assigned arbitrary symbols to nest boxes of breeding pied flycatchers (conspecifics) and blue and great tits, Cyanistes caeruleus and Parus major (heterospecifics), in 2014 and 2016 in two areas with different densities of tits and flycatchers. After ca 50% of flycatchers had returned and a flycatcher symbol was assigned to their nest box, we gave the later arriving flycatchers the choice between empty nest boxes with either a conspecific (flycatcher) or a heterospecific (tit) symbol. As expected, Pied Flycatchers copied the perceived nest box choice of conspecifics, but only in areas that were dominated by flycatchers. Against our initial expectation, flycatchers copied the perceived choice of heterospecifics in the area heavily dominated by tits, even though conspecific minority information was present. Our results confirm that the relative density of conspecifics and heterospecifics modulates the propensity to copy or reject novel behavioural traits. By contrasting conspecific and heterospecific ecology in the same study design we were able to draw more general conclusions about the role of fluctuating densities on social information use.

  14. Nesting success of grassland and savanna birds on reclaimed surface coal mines of the midwestern United States

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Galligan, E.W.; DeVault, T.L.; Lima, S.L.

    2006-12-15

    Reclaimed surface coal mines in southwestern Indiana support many grassland and shrub/savanna bird species of conservation concern. We examined the nesting success of birds on these reclaimed mines to assess whether such 'unnatural' places represent productive breeding habitats for such species. We established eight study sites on two large, grassland-dominated mines in southwestern Indiana and classified them into three categories (open grassland, shrub/savanna, and a mixture of grassland and shrub/savanna) based on broad vegetation and landscape characteristics. During the 1999 and 2000 breeding seasons, we found and monitored 911 nests of 31 species. Daily nest survival for the most commonlymore » monitored grassland species ranged from 0.903 (Dickcissel, Spiza americana) to 0.961 (Grasshopper Sparrow, Ammodramus savannarum). Daily survival estimates for the dominant shrub/savanna nesting species ranged from 0.932 (Brown Thrasher, Toxostoma rufum) to 0.982 (Willow Flycatcher, Empidonax traillii). Vegetation and landscape effects on nesting success were minimal, and only Eastern Meadowlarks (Sturnella magna) showed a clear time-of-season effect, with greater nesting success in the first half of the breeding season. Rates of Brown-headed Cowbird (Molothrus ater) parasitism were only 2.1% for grassland species and 12.0% for shrub/savanna species. The nesting success of birds on reclaimed mine sites was comparable to that in other habitats, indicating that reclaimed habitats on surface mines do not necessarily represent reproductive traps for birds.« less

  15. Birds of a high-altitude cloud forest in Alta Verapaz, Guatemala.

    PubMed

    Eisermann, Knut; Schulz, Ulrich

    2005-01-01

    The Northern Central American Highlands have been recognized as endemic bird area, but little is known about bird communities in Guatemalan cloud forests. From 1997 to 2001 a total of 142 bird species were recorded between 2000 and 2400 masl in cloud forest and agricultural clearings on Montaña Caquipec (Alta Verapaz, Guatemala). The bird community is described based on line transect counts within the forest. Pooling census data from undisturbed and disturbed forest, the Gray-breasted Wood-Wren (Henicorhina leucophrys) was found to be the most abundant species, followed in descending order by the Common Bush-Tanager (Chlorospingus ophthalmicus), the Paltry Tyrannulet (Zimmerius vilissimus), the Yellowish Flycatcher (Empidonax flavescens), the Ruddy-capped Nightingale-Thrush (Catharus frantzi), and the Amethyst-throated Hummingbird (Lampornis amethystinus). Bird communities in undisturbed and disturbed forest were found to be similar (Serensen similarity index 0.85), indicating low human impact. Of all recorded species, approximately 27% were Nearctic-Neotropical migratory birds. The most abundant one was the Wilson's Warbler (Wilsonia pusilla). The Montaña Caquipec is an important area for bird conservation, which is indicated by the presence of four species listed in the IUCN Red List (Highland Guan Penelopina nigra, Resplendent Quetzal Pharomachrus mocinno, Pink-headed Warbler Ergaticus versicolor, Golden-cheeked Warbler Dendroica chrysoparia), and 42 Mesoamerican endemics, of which 14 species are endemic to the Central American Highlands. The results presented here will be useful as baseline data for a long-term monitoring.

  16. Tamarisk biocontrol using tamarisk beetles: Potential consequences for riparian birds in the southwestern United States

    USGS Publications Warehouse

    Paxton, Eben H.; Theimer, Tad C.; Sogge, Mark K.

    2011-01-01

    The tamarisk beetle (Diorhabda spp.), a non-native biocontrol agent, has been introduced to eradicate tamarisk (Tamarix spp.), a genus of non-native tree that has become a dominant component of riparian woodlands in the southwestern United States. Tamarisk beetles have the potential to spread widely and defoliate large expanses of tamarisk habitat, but the effects of such a widespread loss of riparian vegetation on birds remains unknown. We reviewed literature on the effects of other defoliating insects on birds to investigate the potential for tamarisk beetles to affect birds positively or negatively by changing food abundance and vegetation structure. We then combined data on the temporal patterns of tamarisk defoliation by beetles with nest productivity of a well-studied riparian obligate, the Southwestern Willow Flycatcher (Empidonax traillii extimus), to simulate the potential demographic consequences of beetle defoliation on breeding riparian birds in both the short and long term. Our results highlight that the effects of tamarisk biocontrol on birds will likely vary by species and population, depending upon its sensitivity to seasonal defoliation by beetles and net loss of riparian habitat due to tamarisk mortality. Species with restricted distributions that include areas dominated by tamarisk may be negatively affected both in the short and long term. The rate of regeneration and/or restoration of native cottonwoods (Populus spp.) and willows (Salix spp.) relative to the rate of tamarisk loss will be critical in determining the long-term effect of this large-scale ecological experiment.

  17. Breeding phenology of birds: mechanisms underlying seasonal declines in the risk of nest predation.

    PubMed

    Borgmann, Kathi L; Conway, Courtney J; Morrison, Michael L

    2013-01-01

    Seasonal declines in avian clutch size are well documented, but seasonal variation in other reproductive parameters has received less attention. For example, the probability of complete brood mortality typically explains much of the variation in reproductive success and often varies seasonally, but we know little about the underlying cause of that variation. This oversight is surprising given that nest predation influences many other life-history traits and varies throughout the breeding season in many songbirds. To determine the underlying causes of observed seasonal decreases in risk of nest predation, we modeled nest predation of Dusky Flycatchers (Empidonax oberholseri) in northern California as a function of foliage phenology, energetic demand, developmental stage, conspecific nest density, food availability for nest predators, and nest predator abundance. Seasonal variation in the risk of nest predation was not associated with seasonal changes in energetic demand, conspecific nest density, or predator abundance. Instead, seasonal variation in the risk of nest predation was associated with foliage density (early, but not late, in the breeding season) and seasonal changes in food available to nest predators. Supplemental food provided to nest predators resulted in a numerical response by nest predators, increasing the risk of nest predation at nests that were near supplemental feeders. Our results suggest that seasonal changes in foliage density and factors associated with changes in food availability for nest predators are important drivers of temporal patterns in risk of avian nest predation.

  18. Breeding Phenology of Birds: Mechanisms Underlying Seasonal Declines in the Risk of Nest Predation

    PubMed Central

    Borgmann, Kathi L.; Conway, Courtney J.; Morrison, Michael L.

    2013-01-01

    Seasonal declines in avian clutch size are well documented, but seasonal variation in other reproductive parameters has received less attention. For example, the probability of complete brood mortality typically explains much of the variation in reproductive success and often varies seasonally, but we know little about the underlying cause of that variation. This oversight is surprising given that nest predation influences many other life-history traits and varies throughout the breeding season in many songbirds. To determine the underlying causes of observed seasonal decreases in risk of nest predation, we modeled nest predation of Dusky Flycatchers (Empidonax oberholseri) in northern California as a function of foliage phenology, energetic demand, developmental stage, conspecific nest density, food availability for nest predators, and nest predator abundance. Seasonal variation in the risk of nest predation was not associated with seasonal changes in energetic demand, conspecific nest density, or predator abundance. Instead, seasonal variation in the risk of nest predation was associated with foliage density (early, but not late, in the breeding season) and seasonal changes in food available to nest predators. Supplemental food provided to nest predators resulted in a numerical response by nest predators, increasing the risk of nest predation at nests that were near supplemental feeders. Our results suggest that seasonal changes in foliage density and factors associated with changes in food availability for nest predators are important drivers of temporal patterns in risk of avian nest predation. PMID:23776566

  19. 76 FR 50541 - Endangered and Threatened Wildlife and Plants; Designation of Revised Critical Habitat for...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-08-15

    ... centimeters (5.8 inches) in length. The flycatcher is one of four subspecies of the willow flycatcher... clustering into two groups separated approximately along the currently recognized boundary; however, they... central Arizona. Flycatchers are believed to exist and interact as groups of metapopulations (Service 2002...

  20. Two families from New England with usher syndrome type IC with distinct haplotypes.

    PubMed

    DeAngelis, M M; McGee, T L; Keats, B J; Slim, R; Berson, E L; Dryja, T P

    2001-03-01

    To search for patients with Usher syndrome type IC among those with Usher syndrome type I who reside in New England. Genotype analysis of microsatellite markers closely linked to the USH1C locus was done using the polymerase chain reaction. We compared the haplotype of our patients who were homozygous in the USH1C region with the haplotypes found in previously reported USH1C Acadian families who reside in southwestern Louisiana and from a single family residing in Lebanon. Of 46 unrelated cases of Usher syndrome type I residing in New England, two were homozygous at genetic markers in the USH1C region. Of these, one carried the Acadian USH1C haplotype and had Acadian ancestors (that is, from Nova Scotia) who did not participate in the 1755 migration of Acadians to Louisiana. The second family had a haplotype that proved to be the same as that of a family with USH1C residing in Lebanon. Each of the two families had haplotypes distinct from the other. This is the first report that some patients residing in New England have Usher syndrome type IC. Patients with Usher syndrome type IC can have the Acadian haplotype or the Lebanese haplotype compatible with the idea that at least two independently arising pathogenic mutations have occurred in the yet-to-be identified USH1C gene.

  1. Waiting for trees to grow: nest survival, brood parasitism, and the impact of reforestation efforts

    USGS Publications Warehouse

    Hazler, K.R.; Twedt, D.J.; Cooper, R.J.

    2005-01-01

    Of the forested wetlands that once covered the Mississippi Alluvial Valley, only -25% remain due to large-scale conversion to agriculture. Reforestation efforts are currently underway, but tracts planted with slow-growing oaks maintain the structure of a grassland for 5 yr or longer, and will require at least 40 yr to resemble a mature forest. Nonetheless, it is hoped that reforestation, even in early stages, can effectively increase core area in extant tracts of mature forest by reducing higher rates of nest failure and brood parasitism often associated with forest-agriculture interfaces. To test this, we monitored nests of a mature-forest specialist, the Acadian Flycatcher, in extensive bottomland forests adjacent to agricultural fields and reforested tracts (<20 yr-old). We used an information-theoretic approach to evaluate alternative hypotheses regarding the relative impacts of agriculture and reforestation in the landscape. Controlling for year, season, and stand basal area, there was little evidence that landscape context significantly affected nest survival, although survival tended to increase with decreasing amounts of agriculture. The probability of brood parasitism increased with greater proportions of open habitats in the landscape. There was much stronger support for the hypothesis that parasitism rates depended on the sum of agricultural and reforested tracts, rather than on the amount of agriculture alone. Thus, reforested tracts are not expected to have the desired effect of reducing parasitism rates in the adjacent mature forest until several decades have passed.

  2. "Bonjour History": Materials and Ideas for Teaching Louisiana's Cajun History = "Bonjour l'Histoire": Quelques Idees, Deux ou Trois Activites, et Plusieurs Materiaux pour l'Enseignement de l'Histoire des Cadiens en Louisiane.

    ERIC Educational Resources Information Center

    Mounier, Brenda

    The goals of this teacher's guidebook and videotape are designed to incorporate Acadian (Cajun) history into the 4th grade social studies curriculum and the 4th and 5th grade Louisiana 30-minute daily French programs and French immersion programs. Another goal is to create an awareness, appreciation, and understanding of Acadian history in…

  3. Emsian synorogenic paleogeography of the Maine Applachians

    USGS Publications Warehouse

    Bradley, D.; Tucker, R.

    2002-01-01

    The Acadian deformation front in the northern Appalachians of Maine and New Hampshire can now be closely located during the early Emsian (Early Devonian; 408-406 Ma). Tight correlations between paleontologically and isotopically dated rocks are possible only because of a new 408-Ma time scale tie point for the early Emsian. The deformation front lay between a belt of Lower Devonian flysch and molasse that were deposited in an Acadian foreland basin and had not yet been folded and a belt of early Emsian plutons that intruded folded Lower Devonian rocks. This plutonic belt includes the newly dated Ore Mountain gabbro (U/Pb; 406 Ma), which hosts magmatic-sulfide mineralization. Along the deformation front, a 407-Ma pluton that locally truncates Acadian folds (Katahdin) was the feeder to volcanic rocks (Traveler Rhyolite; 406-407 Ma) that are part of the foreland-basin succession involved in these same folds. The Emsian igneous rocks thus define a syncollisional magmatic province that straddled the deformation front. These findings bear on three alternative subduction geometries for the Acadian collision.

  4. Monitoring breeding and migration of neotropical migratory birds at Point Loma, San Diego County, California, 5-year summary, 2011–15

    USGS Publications Warehouse

    Lynn, Suellen; Madden, Melanie C.; Kus, Barbara E.

    2017-04-27

    Executive SummaryWe operated a bird banding station on the Point Loma peninsula in western San Diego County, California, during spring and summer from 2011 to 2015. The station was established in 2010 as part of a long-term monitoring program for neotropical migratory birds during spring migration and for breeding birds as part of the Monitoring Avian Productivity and Survivorship (MAPS) program.During spring migration (April and May), 2011–15, we captured 1,760 individual birds of 54 species, 91 percent (1,595) of which were newly banded, fewer than 1 percent (3) of which were recaptures that were banded in previous years, and 9 percent (143 hummingbirds, 2 hawks, and 17 other birds) of which we released unbanded. We observed an additional 22 species that were not captured. Thirty-four individuals were captured more than once. Bird capture rate averaged 0.49 ± 0.07 captures per net-hour (range 0.41–0.56). Species richness per day averaged 6.87 ± 0.33. Cardellina pusilla (Wilson’s warbler) was the most abundant spring migrant captured, followed by Empidonax difficilis (Pacific-slope flycatcher), Vireo gilvus (warbling vireo), Zonotrichia leucophrys (white-crowned sparrow), and Selasphorus rufus (rufous hummingbird). Captures of white-crowned sparrow decreased, and captures of Pacific-slope flycatcher increased, over the 5 years of our study. Fifty-six percent of known-sex individuals were male and 44 percent were female. The peak number of new species arriving per day ranged from April 1 (2013-six species) to April 16 (2012-five species). A significant correlation was determined between the number of migrants captured each day per net-hour and the density of echoes on the Next-Generation Radar (NEXRAD) images across all 5 years, and in each year except 2014. NEXRAD radar imagery appears to be a useful tool for detecting pulses in migration.Our results indicate that Point Loma provides stopover habitat during migration for 76 migratory species, including 20

  5. Natural selection and inheritance of breeding time and clutch size in the collared flycatcher.

    PubMed

    Sheldon, B C; Kruuk, L E B; Merilä, J

    2003-02-01

    Many characteristics of organisms in free-living populations appear to be under directional selection, possess additive genetic variance, and yet show no evolutionary response to selection. Avian breeding time and clutch size are often-cited examples of such characters. We report analyses of inheritance of, and selection on, these traits in a long-term study of a wild population of the collared flycatcher Ficedula albicollis. We used mixed model analysis with REML estimation ("animal models") to make full use of the information in complex multigenerational pedigrees. Heritability of laying date, but not clutch size, was lower than that estimated previously using parent-offspring regressions, although for both traits there was evidence of substantial additive genetic variance (h2 = 0.19 and 0.29, respectively). Laying date and clutch size were negatively genetically correlated (rA = -0.41 +/- 0.09), implying that selection on one of the traits would cause a correlated response in the other, but there was little evidence to suggest that evolution of either trait would be constrained by correlations with other phenotypic characters. Analysis of selection on these traits in females revealed consistent strong directional fecundity selection for earlier breeding at the level of the phenotype (beta = -0.28 +/- 0.03), but little evidence for stabilising selection on breeding time. We found no evidence that clutch size was independently under selection. Analysis of fecundity selection on breeding values for laying date, estimated from an animal model, indicated that selection acts directly on additive genetic variance underlying breeding time (beta = -0.20 +/- 0.04), but not on clutch size (beta = 0.03 +/- 0.05). In contrast, selection on laying date via adult female survival fluctuated in sign between years, and was opposite in sign for selection on phenotypes (negative) and breeding values (positive). Our data thus suggest that any evolutionary response to selection on

  6. Male-male competition and parental care in collared flycatchers (Ficedula albicollis): an experiment controlling for differences in territory quality.

    PubMed

    Qvarnström, A; Grifffith, S C; Gustafsson, L

    2000-12-22

    Females are known to benefit from mate choice in several different ways but the relationship between these benefits has received little attention. The quality of resources provided by males, such as nest sites, and paternal care are often assumed to covary positively However, because the location of the nest affects the cost of parental care, these two benefits from mate choice can easily be confounded. To investigate the provisioning ability of successful competitors while controlling for differences in territory quality we removed early-settled pairs of collared flycatchers (Ficedula albicollis) and allowed replacement by later-arriving males or floaters (i.e.'poor competitors'). A control group of early-settled males (i.e. 'good competitors') had their females removed. Females paired to good competitors enjoyed a significantly higher reproductive success and tended to receive more parental assistance from their mates compared with females mated to poor competitors. Thus, some males seem able not only to compete successfully over resources but also to feed their offspring at a relatively higher rate. An alternative explanation, that poor competitors invested less in offspring quality in response to a lower share of paternity, could be rejected. The rate of extra-pair paternity did not differ between the two treatment groups. Our results suggest that male- male competition can sometimes facilitate female choice of superior care-givers. Thus, a female's benefit from choosing a competitive male may not be restricted to the quality of the resource he defends but can also include superior paternal care.

  7. Avian use of forest habitats in the Pembina Hills of northeastern North Dakota

    USGS Publications Warehouse

    Faanes, Craig A.; Andrew, Jonathan M.

    1983-01-01

    North Dakota has the least extensive total area of forested habitats of any of the 50 United States. Although occurring in limited area, forest communities add considerably to the total ecological diversity of the State. The forests of the Pembina Hills region in northeastern North Dakota are one of only three areas large enough to be considered of commercial value. During 1981 we studied the avifauna of the upper valley of the Pembina River in the Pembina Hills. Field work extended from 20 April to 23 July; breeding bird censuses were conducted 7 June to 2 July. Of the 120 bird species recorded during the study period, 79 species were recorded during the breeding season. The total breeding population was estimated at nearly 76,000 breeding pairs. The wood warblers (Parulidae) were the most numerous family, accounting for about 28,000 breeding pairs. The yellow warbler (Dendroica petechia) was the most abundant breeding species, making up 19.4% of the population. American redstart (Setophaga ruticilla) was second in abundance, accounting for 10.5% of the breeding population. Largest breeding densities occurred in the willow (Salix sp.) shrub community. Although supporting the lowest mean breeding density, quaking aspen (Populus tremuloides) forests supported the highest species diversity. First State breeding records were recorded for alder flycatcher (Empidonax alnorum) and golden-winged warbler (Vermivora chrysoptera). Records were obtained for 12 species considered rare or unusual in North Dakota during the breeding season. The status of all species known to have occurred in the study area is described in an annotated species list.

  8. Tamarisk biocontrol using tamarisk beetles: Potential consequences for riparian birds in the southwestern United States

    USGS Publications Warehouse

    Paxton, E.H.; Theimer, T.C.; Sogge, M.K.

    2011-01-01

    The tamarisk beetle (Diorhabda spp.), a non-native biocontrol agent, has been introduced to eradicate tamarisk (Tamarix spp.), a genus of non-native tree that has become a dominant component of riparian woodlands in the southwestern United States. Tamarisk beetles have the potential to spread widely and defoliate large expanses of tamarisk habitat, but the effects of such a widespread loss of riparian vegetation on birds remains unknown. We reviewed literature on the effects of other defoliating insects on birds to investigate the potential for tamarisk beetles to affect birds positively or negatively by changing food abundance and vegetation structure. We then combined data on the temporal patterns of tamarisk defoliation by beetles with nest productivity of a wellstudied riparian obligate, the Southwestern Willow Flycatcher (Empidonax traillii extimus), to simulate the potential demographic consequences of beetle defoliation on breeding riparian birds in both the short and long term. Our results highlight that the effects of tamarisk biocontrol on birds will likely vary by species and population, depending upon its sensitivity to seasonal defoliation by beetles and net loss of riparian habitat due to tamarisk mortality. Species with restricted distributions that include areas dominated by tamarisk may be negatively affected both in the short and long term. The rate of regeneration and/or restoration of native cottonwoods (Populus spp.) and willows (Salix spp.) relative to the rate of tamarisk loss will be critical in determining the long-term effect of this large-scale ecological experiment. ?? The Cooper Ornithological Society 2011.

  9. Planned flooding and Colorado River riparian trade-offs downstream from Glen Canyon Dam, Arizona

    USGS Publications Warehouse

    Stevens, Lawrence E.; Ayers, T.J.; Bennett, J.B.; Christensen, K.; Kearsley, M.J.C.; Meretsky, V.J.; Phillips, A. M.; Parnell, R.A.; Spence, J.; Sogge, M.K.; Springer, A.E.; Wegner, D.L.

    2001-01-01

    Regulated river restoration through planned flooding involves trade-offs between aquatic and terrestrial components, between relict pre-dam and novel post-dam resources and processes, and between management of individual resources and ecosystem characteristics. We review the terrestrial (wetland and riparian) impacts of a 1274 m3/s test flood conducted by the U.S. Bureau of Reclamation in March/April 1996, which was designed to improve understanding of sediment transport and management downstream from Glen Canyon Dam in the Colorado River ecosystem. The test flood successfully restored sandbars throughout the river corridor and was timed to prevent direct impacts to species of concern. A total of 1275 endangered Kanab ambersnail (Oxyloma haydeni kanabensis) were translocated above the flood zone at Vaseys Paradise spring, and an estimated 10.7% of the total snail habitat and 7.7% of the total snail population were lost to the flood. The test flood scoured channel margin wetlands, including potential foraging habitats of endangered Southwestern Willow Flycatcher (Empidonax traillii extimus). It also buried ground-covering riparian vegetation under >1 m of fine sand but only slightly altered woody sandbar vegetation and some return-current channel marshes. Pre-flood control efforts and appropriate flood timing limited recruitment of four common nonnative perennial plant species. Slight impacts on ethnobotanical resources were detected >430 km downstream, but those plant assemblages recovered rapidly. Careful design of planned flood hydrograph shape and seasonal timing is required to mitigate terrestrial impacts during efforts to restore essential fluvial geomorphic and aquatic habitats in regulated river ecosystems.

  10. Subtle but ubiquitous selection on body size in a natural population of collared flycatchers over 33 years.

    PubMed

    Björklund, M; Gustafsson, L

    2017-07-01

    Understanding the magnitude and long-term patterns of selection in natural populations is of importance, for example, when analysing the evolutionary impact of climate change. We estimated univariate and multivariate directional, quadratic and correlational selection on four morphological traits (adult wing, tarsus and tail length, body mass) over a time period of 33 years (≈ 19 000 observations) in a nest-box breeding population of collared flycatchers (Ficedula albicollis). In general, selection was weak in both males and females over the years regardless of fitness measure (fledged young, recruits and survival) with only few cases with statistically significant selection. When data were analysed in a multivariate context and as time series, a number of patterns emerged; there was a consistent, but weak, selection for longer wings in both sexes, selection was stronger on females when the number of fledged young was used as a fitness measure, there were no indications of sexually antagonistic selection, and we found a negative correlation between selection on tarsus and wing length in both sexes but using different fitness measures. Uni- and multivariate selection gradients were correlated only for wing length and mass. Multivariate selection gradient vectors were longer than corresponding vector of univariate gradients and had more constrained direction. Correlational selection had little importance. Overall, the fitness surface was more or less flat with few cases of significant curvature, indicating that the adaptive peak with regard to body size in this species is broader than the phenotypic distribution, which has resulted in weak estimates of selection. © 2017 European Society For Evolutionary Biology. Journal of Evolutionary Biology © 2017 European Society For Evolutionary Biology.

  11. Genomic distribution and estimation of nucleotide diversity in natural populations: perspectives from the collared flycatcher (Ficedula albicollis) genome.

    PubMed

    Dutoit, Ludovic; Burri, Reto; Nater, Alexander; Mugal, Carina F; Ellegren, Hans

    2017-07-01

    Properly estimating genetic diversity in populations of nonmodel species requires a basic understanding of how diversity is distributed across the genome and among individuals. To this end, we analysed whole-genome resequencing data from 20 collared flycatchers (genome size ≈1.1 Gb; 10.13 million single nucleotide polymorphisms detected). Genomewide nucleotide diversity was almost identical among individuals (mean = 0.00394, range = 0.00384-0.00401), but diversity levels varied extensively across the genome (95% confidence interval for 200-kb windows = 0.0013-0.0053). Diversity was related to selective constraint such that in comparison with intergenic DNA, diversity at fourfold degenerate sites was reduced to 85%, 3' UTRs to 82%, 5' UTRs to 70% and nondegenerate sites to 12%. There was a strong positive correlation between diversity and chromosome size, probably driven by a higher density of targets for selection on smaller chromosomes increasing the diversity-reducing effect of linked selection. Simulations exploring the ability of sequence data from a small number of genetic markers to capture the observed diversity clearly demonstrated that diversity estimation from finite sampling of such data is bound to be associated with large confidence intervals. Nevertheless, we show that precision in diversity estimation in large outbred population benefits from increasing the number of loci rather than the number of individuals. Simulations mimicking RAD sequencing showed that this approach gives accurate estimates of genomewide diversity. Based on the patterns of observed diversity and the performed simulations, we provide broad recommendations for how genetic diversity should be estimated in natural populations. © 2016 The Authors. Molecular Ecology Resources Published by John Wiley & Sons Ltd.

  12. Habitat shifts in the evolutionary history of a Neotropical flycatcher lineage from forest and open landscapes

    PubMed Central

    2008-01-01

    Background Little is known about the role ecological shifts play in the evolution of Neotropical radiations that have colonized a variety of environments. We here examine habitat shifts in the evolutionary history of Elaenia flycatchers, a Neotropical bird lineage that lives in a range of forest and open habitats. We evaluate phylogenetic relationships within the genus based on mitochondrial and nuclear DNA sequence data, and then employ parsimony-based and Bayesian methods to reconstruct preferences for a number of habitat types and migratory behaviour throughout the evolutionary history of the genus. Using a molecular clock approach, we date the most important habitat shifts. Results Our analyses resolve phylogenetic relationships among Elaenia species and confirm several species associations predicted by morphology while furnishing support for other taxon placements that are in conflict with traditional classification, such as the elevation of various Elaenia taxa to species level. While savannah specialism is restricted to one basal clade within the genus, montane forest was invaded from open habitat only on a limited number of occasions. Riparian growth may have been favoured early on in the evolution of the main Elaenia clade and subsequently been deserted on several occasions. Austral long-distance migratory behaviour evolved on several occasions. Conclusion Ancestral reconstructions of habitat preferences reveal pronounced differences not only in the timing of the emergence of certain habitat preferences, but also in the frequency of habitat shifts. The early origin of savannah specialism in Elaenia highlights the importance of this habitat in Neotropical Pliocene and late Miocene biogeography. While forest in old mountain ranges such as the Tepuis and the Brazilian Shield was colonized early on, the most important colonization event of montane forest was in conjunction with Pliocene Andean uplift. Riparian habitats may have played an important role in

  13. Monitoring avian productivity and survivorship (MAPS) 5-year summary, Naval Outlying Landing Field, Imperial Beach, southwestern San Diego County, California, 2009-13

    USGS Publications Warehouse

    Lynn, Suellen; Madden, Melanie C.; Houston, Alexandra; Kus, Barbara E.

    2015-01-01

    During 2009–13, a Monitoring Avian Productivity and Survivorship (MAPS) banding station was operated at the Naval Outlying Landing Field (NOLF), Imperial Beach, in southwestern San Diego County, California. The station was established as part of a long-term monitoring program of Neotropical migratory bird populations on NOLF and helps Naval Base Coronado (NOLF is a component) meet the goals and objectives of Department of Defense Partners in Flight program and the Birds and Migratory Birds Management Strategies of the Naval Base Coronado Integrated Natural Resources Management Plan. During 2009–13, captures averaged 644 ±155 per year. Fifty-seven species were captured, of which 44 are Neotropical migratory species and 33 breed at the MAPS station. Twenty-two sensitive species were detected, including Least Bell’s Vireo (Vireo bellii pusillus), Willow Flycatcher (Empidonax traillii), Yellow-breasted Chat (Icteria virens) and Yellow Warbler (Setophaga petechia). Local population trends varied among species and years, as did annual productivity (number of young per adult). We found no significant relationship between productivity and the observed population size in the subsequent year for any species, nor did we find an association between productivity and precipitation for the current bio-year. Similarly, survivorship varied across species and years, and there was no obvious relationship between adult survivorship and observed population size for any species except Wrentit (Chamaea fasciata), for which the relationship was positive. Adult survivorship was unrelated to precipitation at the MAPS station. Additional years of data will be required to generate sample sizes adequate for more rigorous analyses of survivorship and productivity as predictors of population growth.

  14. 77 FR 41147 - Endangered and Threatened Wildlife and Plants; Designation of Revised Critical Habitat for the...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2012-07-12

    ... segments and associated flycatcher habitat are anticipated to support the strategy, rationale, and science... strategy, rationale, and science of flycatcher conservation in order to meet territory and habitat-related... costs associated with monitoring and education activities, fencing, habitat restoration and creation...

  15. Whole-genome patterns of linkage disequilibrium across flycatcher populations clarify the causes and consequences of fine-scale recombination rate variation in birds.

    PubMed

    Kawakami, Takeshi; Mugal, Carina F; Suh, Alexander; Nater, Alexander; Burri, Reto; Smeds, Linnéa; Ellegren, Hans

    2017-08-01

    Recombination rate is heterogeneous across the genome of various species and so are genetic diversity and differentiation as a consequence of linked selection. However, we still lack a clear picture of the underlying mechanisms for regulating recombination. Here we estimated fine-scale population recombination rate based on the patterns of linkage disequilibrium across the genomes of multiple populations of two closely related flycatcher species (Ficedula albicollis and F. hypoleuca). This revealed an overall conservation of the recombination landscape between these species at the scale of 200 kb, but we also identified differences in the local rate of recombination despite their recent divergence (<1 million years). Genetic diversity and differentiation were associated with recombination rate in a lineage-specific manner, indicating differences in the extent of linked selection between species. We detected 400-3,085 recombination hotspots per population. Location of hotspots was conserved between species, but the intensity of hotspot activity varied between species. Recombination hotspots were primarily associated with CpG islands (CGIs), regardless of whether CGIs were at promoter regions or away from genes. Recombination hotspots were also associated with specific transposable elements (TEs), but this association appears indirect due to shared preferences of the transposition machinery and the recombination machinery for accessible open chromatin regions. Our results suggest that CGIs are a major determinant of the localization of recombination hotspots, and we propose that both the distribution of TEs and fine-scale variation in recombination rate may be associated with the evolution of the epigenetic landscape. © 2017 The Authors. Molecular Ecology Published by John Wiley & Sons Ltd.

  16. Effects of sea-level rise and anthropogenic development on priority bird species habitats in coastal Georgia, USA.

    PubMed

    Brittain, Ross A; Craft, Christopher B

    2012-02-01

    We modeled changes in area of five habitats, tidal-freshwater forest, salt marsh, maritime shrub-scrub (shrub), maritime broadleaf forest (oak) and maritime narrowleaf (pine) forest, in coastal Georgia, USA, to evaluate how simultaneous habitat loss due to predicted changes in sea level rise (SLR) and urban development will affect priority bird species of the south Atlantic coastal plain by 2100. Development rates, based on regional growth plans, were modeled at 1% and 2.5% annual urban growth, while SLR rates, based on the Intergovernmental Panel on Climate Change's A1B mean and maximum scenarios, were modeled at 52 cm and 82 cm, respectively. SLR most greatly affected the shrub habitat with predicted losses of 35-43%. Salt marsh and tidal forest also were predicted to lose considerable area to SLR (20-45 and 23-35%, respectively), whereas oak and pine forests had lesser impact from SLR, 18-22% and 11-15%, respectively. Urban development resulted in losses of considerable pine (48-49%) and oak (53-55%) habitat with lesser loss of shrub habitat (21-24%). Under maximum SLR and urban growth, shrub habitat may lose up to 59-64% compared to as much as 62-65% pine forest and 74-75% oak forest. Conservation efforts should focus on protection of shrub habitat because of its small area relative to other terrestrial habitats and use by Painted Buntings (Passerina ciris), a Partners In Flight (PIF) extremely high priority species. Tidal forests also deserve protection because they are a likely refuge for forest species, such as Northern Parula and Acadian Flycatcher, with the decline of oak and pine forests due to urban development.

  17. Implications of Climate Change for Bird Conservation in the Southwestern U.S. under Three Alternative Futures

    PubMed Central

    Friggens, Megan M.; Finch, Deborah M.

    2015-01-01

    Future expected changes in climate and human activity threaten many riparian habitats, particularly in the southwestern U.S. Using Maximum Entropy (MaxEnt3.3.3) modeling, we characterized habitat relationships and generated spatial predictions of habitat suitability for the Lucy’s warbler (Oreothlypis luciae), the Southwestern willow flycatcher (Empidonax traillii extimus) and the Western yellow-billed cuckoo (Coccyzus americanus). Our goal was to provide site- and species-specific information that can be used by managers to identify areas for habitat conservation and/or restoration along the Rio Grande in New Mexico. We created models of suitable habitat for each species based on collection and survey samples and climate, biophysical, and vegetation data. We projected habitat suitability under future climates by applying these models to conditions generated from three climate models for 2030, 2060 and 2090. By comparing current and future distributions, we identified how habitats are likely to change as a result of changing climate and the consequences of those changes for these bird species. We also examined whether land ownership of high value sites shifts under changing climate conditions. Habitat suitability models performed well. Biophysical characteristics were more important that climate conditions for predicting habitat suitability with distance to water being the single most important predictor. Climate, though less important, was still influential and led to declines of suitable habitat of more than 60% by 2090. For all species, suitable habitat tended to shrink over time within the study area leaving a few core areas of high importance. Overall, climate changes will increase habitat fragmentation and reduce breeding habitat patch size. The best strategy for conserving bird species within the Rio Grande will include measures to maintain and restore critical habitat refugia. This study provides an example of a presence-only habitat model that can be

  18. Implications of Climate Change for Bird Conservation in the Southwestern U.S. under Three Alternative Futures.

    PubMed

    Friggens, Megan M; Finch, Deborah M

    2015-01-01

    Future expected changes in climate and human activity threaten many riparian habitats, particularly in the southwestern U.S. Using Maximum Entropy (MaxEnt3.3.3) modeling, we characterized habitat relationships and generated spatial predictions of habitat suitability for the Lucy's warbler (Oreothlypis luciae), the Southwestern willow flycatcher (Empidonax traillii extimus) and the Western yellow-billed cuckoo (Coccyzus americanus). Our goal was to provide site- and species-specific information that can be used by managers to identify areas for habitat conservation and/or restoration along the Rio Grande in New Mexico. We created models of suitable habitat for each species based on collection and survey samples and climate, biophysical, and vegetation data. We projected habitat suitability under future climates by applying these models to conditions generated from three climate models for 2030, 2060 and 2090. By comparing current and future distributions, we identified how habitats are likely to change as a result of changing climate and the consequences of those changes for these bird species. We also examined whether land ownership of high value sites shifts under changing climate conditions. Habitat suitability models performed well. Biophysical characteristics were more important that climate conditions for predicting habitat suitability with distance to water being the single most important predictor. Climate, though less important, was still influential and led to declines of suitable habitat of more than 60% by 2090. For all species, suitable habitat tended to shrink over time within the study area leaving a few core areas of high importance. Overall, climate changes will increase habitat fragmentation and reduce breeding habitat patch size. The best strategy for conserving bird species within the Rio Grande will include measures to maintain and restore critical habitat refugia. This study provides an example of a presence-only habitat model that can be used

  19. Songbird response to increased willow (Salix spp.) growth in Yellowstone's northern range.

    PubMed

    Baril, Lisa M; Hansen, Andrew J; Renkin, Roy; Lawrence, Rick

    2011-09-01

    After nearly a century of height suppression, willows (Salix spp.) in the northern range of Yellowstone National Park, U.S.A., are increasing in height growth as a possible consequence of wolf (Canis lupus) restoration, climate change, or other factors. Regardless of the drivers, the recent release of this rare but important habitat type could have significant implications for associated songbirds that are exhibiting declines in the region. Our objective was to evaluate bird response to releasing willows by comparing willow structure and bird community composition across three willow growth conditions: height suppressed, recently released, and previously tall (i.e., tall prior to the height increase of released willows). Released and previously tall willows exhibited high and similar vertical structure, but released willows were significantly lower in horizontal structure. Suppressed willows were significantly shorter and lower in horizontal cover than released or previously tall willows. Bird richness increased along a gradient from lowest in suppressed to highest in previously tall willows, but abundance and diversity were similar between released and previously tall willows, despite lower horizontal cover in the released condition. Common Yellowthroat (Geothlypis trichas) and Lincoln's Sparrow (Melospiza lincolnii) were found in all three growth conditions; however, Yellow Warbler (Dendroica petechia), Warbling Vireo (Vireo gilvus), Willow Flycatcher (Empidonax traillii), and Song Sparrow (Melospiza melodii) were present in released and previously tall willows only. Wilson's Warbler (Wilsonia pusilla) was found in previously tall willows only, appearing to specialize on tall, dense willows. The results of our a priori habitat models indicated that foliage height diversity was the primary driver of bird richness, abundance, and diversity. These results indicate that vertical structure was a more important driver of bird community variables than horizontal

  20. Retrospective multicenter evaluation of the "fly-catching syndrome" in 24 dogs: EEG, BAER, MRI, CSF findings and response to antiepileptic and antidepressant treatment.

    PubMed

    Wrzosek, Marcin; Płonek, Marta; Nicpoń, Józef; Cizinauskas, Sigitas; Pakozdy, Akos

    2015-12-01

    The fly-catching syndrome (FCS) is a rare canine condition of sudden, occasional, or constant episodes of biting the air. It may be accompanied by jumping, licking, and swallowing. The etiology of FCS is unknown and controversial. Various explanations for its occurrence have included epileptoid disorders such as visual cortex epileptiform disturbances and simple and complex partial seizures as well as compulsive disorders, hallucinatory behavior, and stereotypy. A retrospective multicenter analysis of 24 dogs with clinical symptoms of FCS is presented. Clinical signs at the time of presentation, the mean age at onset of the disease, the response to treatment, and the clinical outcome were recorded and analyzed in all patients. All dogs underwent clinical, neurological, and otoscopic examinations. Complete blood cell counts (CBCs) and serum chemistry panels were obtained from each dog. Diagnostic testing included MRI and EEG examinations in 21 cases, BAER in 19 cases, and CSF analysis in 20 cases. The EEG revealed spike activity in 8 (38%) of the 21 cases, 7 of which had activity in the occipital lobes. The brainstem auditory evoked response (BAER) revealed three cases of bilateral deafness. The MRI revealed six cases of Chiari malformation (CM), one case of syringohydromyelia (SM), and one case of a falx cerebri meningioma. The dogs were divided into groups according to their treatment protocol. Group A included dogs treated with phenobarbital (PB), and group B consisted of dogs treated with fluoxetine (FLX). Thirty-six percent of the dogs in group A responded to PB, while 100% of the dogs in group B responded to FLX. The results suggest that FCS is more responsive to FLX than PB. However, the etiology of this behavior remains unclear in most cases. Copyright © 2015 The Authors. Published by Elsevier Inc. All rights reserved.

  1. The tectono-thermal evolution of the Waterbury dome, western Connecticut, based on U-Pb and 40Ar/39Ar ages

    USGS Publications Warehouse

    Dietsch, Craig; Kunk, Michael J.; Aleinikoff, John; Sutter, John F.

    2010-01-01

    Level 3 nappes were emplaced over the Waterbury dome along an Acadian décollement synchronous with the formation of a D3 thrust duplex in the dome. The décollement truncates the Ky + Kfs-in (migmatite) isograd in the dome core and a St-in isograd in level 3 nappes, indicating that peak metamorphic conditions in the dome core and nappe cover rocks formed in different places at different times. Metamorphic overgrowths on zircon from the felsic orthogneiss in the Waterbury dome have an age of 387 ± 5 Ma. Rocks of all levels and the décollement are folded by D4 folds that have a strongly developed, regional crenulation cleavage and D5 folds. The Waterbury dome was formed by thrust duplexing followed by fold interference during the Acadian orogeny. The 40Ar/39Ar ages of amphibole, muscovite, biotite, and K-feldspar from above and below the décollement are ca. 378 Ma, 355 Ma, 360 Ma (above) and 340 (below), and 288 Ma, respectively. Any kilometer-scale vertical movements between dome and nappe rocks were over by ca. 378 Ma. Core and cover rocks of the Waterbury dome record synchronous, post-Acadian cooling.

  2. Offshore medicine.

    PubMed

    Baker, D

    2001-03-01

    Offshore life can be refreshing for medics who are looking for a little change of pace; however, it is not for everyone. Working offshore can be the easiest or most boring job you'll ever have. It takes a specific type of medic to fit this mold. So, if you are considering a career in the offshore field, take all of the above into consideration. You are not just making a change in jobs, but a change in lifestyle. Once you become accustomed to this lifestyle, it will be hard to go back to the everyday hustle and bustle of the streets. For more information about working offshore, contact Acadian Contract Services at 800/259-333, or visit www.acadian.com.

  3. Nasal mites from birds of a Guatemalan cloud forest (Acarina: rhinonyssidae)

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Spicer, G.S.

    1984-01-01

    A survey of the nasal mites from Guatemalan cloud forest birds is reported. Seventy-eight birds, representing 10 families and 18 species, were examined. Prevalence of infection was 24%. Two new species are described: Sternostoma darlingi from Mitrephanes phaeocercus (Tyrannidae) and S. pencei from Empidonax flavescens (Tyrannidae). New host records are reported for S. priangae from Chlorospingus opthalmicus (Thraupidae), S. hutsoni from Catharus dryas (Turdidae), Ptilonyssus sairae from Chlorospingus opthalmicus (Thraupidae), and Myioborus miniatus (Parulidae), P. euroturdi from Catharus dryas (Turdidae), P. tyrannus from Empidonax flavescens and Mitrephanes phaeocercus (both Tyannidae), and Tinaminyssus ixoreus from Catharus dryas (Turdidae). The subspecies Ptilonyssusmore » euroturdi mimicola Fain and Hyland is synonymized with the nominate subspecies. Data are presented to suggest that the Rhinonyssidae may be a polyphyletic assemblage. 35 references, 12 figures, 1 table.« less

  4. Females discriminate against heterospecific sperm in a natural hybrid zone.

    PubMed

    Cramer, Emily R A; Ålund, Murielle; McFarlane, S Eryn; Johnsen, Arild; Qvarnström, Anna

    2016-08-01

    When hybridization is maladaptive, species-specific mate preferences are selectively favored, but low mate availability may constrain species-assortative pairing. Females paired to heterospecifics may then benefit by copulating with multiple males and subsequently favoring sperm of conspecifics. Whether such mechanisms for biasing paternity toward conspecifics act as important reproductive barriers in socially monogamous vertebrate species remains to be determined. We use a combination of long-term breeding records from a natural hybrid zone between collared and pied flycatchers (Ficedula albicollis and F. hypoleuca), and an in vitro experiment comparing conspecific and heterospecific sperm performance in female reproductive tract fluid, to evaluate the potential significance of female cryptic choice. We show that the females most at risk of hybridizing (pied flycatchers) frequently copulate with multiple males and are able to inhibit heterospecific sperm performance. The negative effect on heterospecific sperm performance was strongest in pied flycatcher females that were most likely to have been previously exposed to collared flycatcher sperm. We thus demonstrate that a reproductive barrier acts after copulation but before fertilization in a socially monogamous vertebrate. While the evolutionary history of this barrier is unknown, our results imply that there is opportunity for it to be accentuated via a reinforcement-like process. © 2016 The Author(s). Evolution © 2016 The Society for the Study of Evolution.

  5. Spatial Patterns of Subtidal Benthic Invertebrates and Environmental Factors in the Nearshore Gulf of Maine

    EPA Science Inventory

    The spatial patterns of subtidal benthic invertebrates and physical-chemical variables in the nearshore Gulf of Maine (Acadian Biogeographic Province) were studied to provide information needed to calibrate benthic indices of environmental condition, determine physical-chemical f...

  6. 76 FR 26313 - Endangered and Threatened Species Permit Applications

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-05-06

    ..., during normal business hours at the U.S. Fish and Wildlife Service, 500 Gold Ave., SW., Room 6034...: Rhea Environmental Consulting, Mancos, Colorado. Applicant requests a renewal to a current permit for... (Empidonax traillii extimus) within Arizona, New Mexico, Colorado, and Utah. Permit TE-150490 Applicant: John...

  7. Vitamin profiles in two free-living passerine birds under a metal pollution gradient - A calcium supplementation experiment.

    PubMed

    Ruiz, Sandra R; Espín, Silvia; Sánchez-Virosta, Pablo; Salminen, Juha-Pekka; Lilley, Thomas M; Eeva, Tapio

    2017-04-01

    Vitamin and carotenoid deficiency may impair development in free-living vertebrates, because of the importance of these micronutrients to growth, antioxidant defense and calcium regulation. Micronutrient and calcium insufficiency can be intensified by metal pollution which can interfere with nutrient homeostasis or indirectly reduce food availability. Furthermore, absorption of dietary heavy metals is dependent on food calcium and vitamin levels. We investigated the effect of calcium on plasma vitamin and carotenoid profiles and how these affected growth and survival in two passerine birds with different calcium turnover living along a metal pollution gradient. Vitamins (A, D 3 and E) and carotenoids were quantified from blood plasma of great tit (Parus major) and pied flycatcher (Ficedula hypoleuca) nestlings. Metal concentrations in soil and in feces from the same nestlings were used to assess the exposure to air pollution. Additionally, we examined the vitamin level variation between developmental stages (eggs and nestlings within the same brood). Our results showed that generally higher concentrations of vitamins and carotenoids circulate in blood of great tits than in pied flycatchers. In general, birds inhabiting the polluted zone presented lower concentrations of the studied micronutrients. Calcium supplementation and metal pollution decreased vitamin A concentration in pied flycatcher, but not in great tit, while vitamin A affected growth and survival in great tit and pied flycatcher respectively. Our results suggest that populations under exposure to metal pollution may experience increased vitamin A deficiency, and that the two passerine species, while obtaining similar micronutrients in food, respond differently to environmental disturbance of nutrients. Copyright © 2017 Elsevier Inc. All rights reserved.

  8. Insights into the genetic architecture of morphological traits in two passerine bird species.

    PubMed

    Silva, C N S; McFarlane, S E; Hagen, I J; Rönnegård, L; Billing, A M; Kvalnes, T; Kemppainen, P; Rønning, B; Ringsby, T H; Sæther, B-E; Qvarnström, A; Ellegren, H; Jensen, H; Husby, A

    2017-09-01

    Knowledge about the underlying genetic architecture of phenotypic traits is needed to understand and predict evolutionary dynamics. The number of causal loci, magnitude of the effects and location in the genome are, however, still largely unknown. Here, we use genome-wide single-nucleotide polymorphism (SNP) data from two large-scale data sets on house sparrows and collared flycatchers to examine the genetic architecture of different morphological traits (tarsus length, wing length, body mass, bill depth, bill length, total and visible badge size and white wing patches). Genomic heritabilities were estimated using relatedness calculated from SNPs. The proportion of variance captured by the SNPs (SNP-based heritability) was lower in house sparrows compared with collared flycatchers, as expected given marker density (6348 SNPs in house sparrows versus 38 689 SNPs in collared flycatchers). Indeed, after downsampling to similar SNP density and sample size, this estimate was no longer markedly different between species. Chromosome-partitioning analyses demonstrated that the proportion of variance explained by each chromosome was significantly positively related to the chromosome size for some traits and, generally, that larger chromosomes tended to explain proportionally more variation than smaller chromosomes. Finally, we found two genome-wide significant associations with very small-effect sizes. One SNP on chromosome 20 was associated with bill length in house sparrows and explained 1.2% of phenotypic variation (V P ), and one SNP on chromosome 4 was associated with tarsus length in collared flycatchers (3% of V P ). Although we cannot exclude the possibility of undetected large-effect loci, our results indicate a polygenic basis for morphological traits.

  9. Evaluation of a Biostimulant (Pepton) Based in Enzymatic Hydrolyzed Animal Protein in Comparison to Seaweed Extracts on Root Development, Vegetative Growth, Flowering, and Yield of Gold Cherry Tomatoes Grown under Low Stress Ambient Field Conditions

    PubMed Central

    Polo, Javier; Mata, Pedro

    2018-01-01

    The objectives of this experiment were to determine the effects of different application rates of an enzyme hydrolyzed animal protein biostimulant (Pepton) compared to a standard application rate of a biostimulant derived from seaweed extract (Acadian) on plant growth parameters and yield of gold cherry tomatoes (Solanum lycopersicum L.). Biostimulant treatments were applied starting at 15 days after transplant and every 2 weeks thereafter for a total of 5 applications. One treatment group received no biostimulant (Control). Three treatment groups (Pepton-2, Pepton-3, Pepton-4) received Pepton at different application rates equivalent to 2, 3, or 4 kg/ha applied by foliar (first 2 applications) and by irrigation (last 3 applications). Another treatment group (Acadian) received Acadian at 1.5 L/ha by irrigation for all five applications. All groups received the regular fertilizer application for this crop at transplantation, flowering, and fruiting periods. There were four plots per treatment group. Each plot had a surface area of 21 m2 that consisted of two rows that were 7 m long and 1.5 m wide. Plant height, stem diameter, distance from head to bouquet flowering, fruit set distance between the entire cluster and cluster flowering fruit set, leaf length, and number of leaves per plant was recorded for 20 plants (5 plants per plot) at 56 and 61 days after the first application. Root length and diameter of cherry tomatoes were determined at harvest from 20 randomly selected plants. Harvesting yield per plot was registered and production per hectare was calculated. Both biostimulants improved (P < 0.05) all vegetative parameters compared with the control group. There was a positive linear (P < 0.001) effect of Pepton application rate for all parameters. The calculated yield was 7.8 and 1 Ton/ha greater that represent 27 and 2.9% higher production for Pepton applied at 4 kg/ha compared to the control and to Acadian, respectively. In conclusion, Pepton was effective

  10. Actes des Journees de linguistique (Proceedings of the Linguistics Conference) (7th, March 1993).

    ERIC Educational Resources Information Center

    Drouin, Patrick, Ed.; And Others

    Papers from a 1993 conference on linguistics, all in French, include essays on the following: Yoruba morphophonology; literary Arabic morphophonology; grammatical cohesion in Burushaski; phonological and lexical variation in French Canadian dialects, including Acadian; insults in Madrid Spanish; discourse analysis; maintenance of meaning in…

  11. Concept Categories as Measures of Culture Distance.

    ERIC Educational Resources Information Center

    Mackey, William F.

    The research reported in this paper tests the possibility of using differences in conceptual categories in the measurement of differences between two cultures. In the initial experiment, which contrasts French and French-Acadian linguistic habits, the authors limit themselves to conceptual categories reflected in concrete nouns representing the…

  12. (Re)-Introduction to French: Four Education Models to Revitalise an Endangered Group in Eastern Canada

    ERIC Educational Resources Information Center

    Cormier, Marianne; Bourque, Jimmy; Jolicoeur, Manon

    2014-01-01

    This study explores early francization models for a linguistic minority currently struggling to preserve its language. The French Acadians of New Brunswick, Canada, represent 30% of the province's total population, yet their numbers and their linguistic vitality are decreasing. New Brunswick has two public school systems: the English language…

  13. Culturally and economically important nontimber forest products of northern Maine

    Treesearch

    Michelle J. Baumflek; Marla R. Emery; Clare Ginger

    2010-01-01

    Nontimber forest products (NTFPs) gathered for food, medicine, craft, spiritual, aesthetic, and utilitarian purposes make substantial contributions to the economic viability and cultural vitality of communities. In the St. John River watershed of northern Maine, people identifying with cultural groups including Acadian, Maliseet, Mi'kmaq, Scotch-Irish, and Swedish...

  14. A Study of the Linguistic Features of Cajun English.

    ERIC Educational Resources Information Center

    Cox, Juanita

    The study contrasts Acadian English (Cajun) spoken in Louisiana with the local standard English, describing the linguistic features (pronunciation, grammar, vocabulary) of the dialect in non-technical language. The objective is to inform elementary and secondary school teachers and others involved in education and curriculum development for a…

  15. 15 CFR Appendix I to Part 921 - Biogeographic Classification Scheme

    Code of Federal Regulations, 2014 CFR

    2014-01-01

    ... 15 Commerce and Foreign Trade 3 2014-01-01 2014-01-01 false Biogeographic Classification Scheme I Appendix I to Part 921 Commerce and Foreign Trade Regulations Relating to Commerce and Foreign Trade... Part 921—Biogeographic Classification Scheme Acadian 1. Northern of Maine (Eastport to the Sheepscot...

  16. 15 CFR Appendix I to Part 921 - Biogeographic Classification Scheme

    Code of Federal Regulations, 2013 CFR

    2013-01-01

    ... 15 Commerce and Foreign Trade 3 2013-01-01 2013-01-01 false Biogeographic Classification Scheme I Appendix I to Part 921 Commerce and Foreign Trade Regulations Relating to Commerce and Foreign Trade... Part 921—Biogeographic Classification Scheme Acadian 1. Northern of Maine (Eastport to the Sheepscot...

  17. 15 CFR Appendix I to Part 921 - Biogeographic Classification Scheme

    Code of Federal Regulations, 2012 CFR

    2012-01-01

    ... 15 Commerce and Foreign Trade 3 2012-01-01 2012-01-01 false Biogeographic Classification Scheme I Appendix I to Part 921 Commerce and Foreign Trade Regulations Relating to Commerce and Foreign Trade... Part 921—Biogeographic Classification Scheme Acadian 1. Northern of Maine (Eastport to the Sheepscot...

  18. 15 CFR Appendix I to Part 921 - Biogeographic Classification Scheme

    Code of Federal Regulations, 2010 CFR

    2010-01-01

    ... 15 Commerce and Foreign Trade 3 2010-01-01 2010-01-01 false Biogeographic Classification Scheme I Appendix I to Part 921 Commerce and Foreign Trade Regulations Relating to Commerce and Foreign Trade... Part 921—Biogeographic Classification Scheme Acadian 1. Northern of Maine (Eastport to the Sheepscot...

  19. 15 CFR Appendix I to Part 921 - Biogeographic Classification Scheme

    Code of Federal Regulations, 2011 CFR

    2011-01-01

    ... 15 Commerce and Foreign Trade 3 2011-01-01 2011-01-01 false Biogeographic Classification Scheme I Appendix I to Part 921 Commerce and Foreign Trade Regulations Relating to Commerce and Foreign Trade... Part 921—Biogeographic Classification Scheme Acadian 1. Northern of Maine (Eastport to the Sheepscot...

  20. A new model for the provenance of the Upper Devonian Old Red Sandstone (UORS) of southern Ireland

    NASA Astrophysics Data System (ADS)

    Ennis, Meg; Meere, Pat; Timmerman, Martin

    2010-05-01

    The geology of Southern Ireland is dominated by the influence of both the Caledonian and Variscan orogenies which have shaped the landscape of today. The Old Red Sandstone (ORS) sequences of the Middle - Upper Devonian Munster Basin have traditionally been viewed as a post-orogenic molasse deposit sourced from the Caledonides (Friend et al. 2000 & references therein), which were subsequently deformed by the Late Carboniferous Variscan Orogeny. This model does not take into account the potential impact of the Acadian Orogeny, an Early to Mid Devonian transpressional tectonic event which culminated in Mid Emsian times and resulted in the deformation and inversion of Lower ORS (LORS) basins across Britain and Ireland (Soper & Woodcock 2003; Meere & Mulchrone 2006). Evidence of Acadian deformation in Southern Ireland is recorded in the LORS sequence of the Lower-Middle Devonian basin, the Dingle Basin. Meere & Mulchrone (2006) show that penetrative deformation visible in the LORS of the Dingle Basin has an Acadian signature and is not associated with Late Carboniferous Variscan compression (Parkin 1976; Todd 2000). The role of the Acadian Orogeny in the tectono-sedimentary evolution of Southern Ireland has been analyzed in this study using a multidisciplinary approach. Petrographic analysis of both the LORS and Upper ORS (UORS) of southern Ireland suggests an alternative provenance model in which there is a direct genetic link between the two Devonian deposits. There is a fining-up relationship between the two basins and the volcanic lithic fragments - while extremely limited in occurrence in the Munster Basin - are strikingly similar in both units. The absence of conglomeratic units at the base of the Munster Basin provide further evidence that the UORS does not represent a classic molasse deposit. This is supported by EMPA data from both basins which indicates identical mica chemistries in both the LORS and UORS. A comparison with the white mica chemistries from a

  1. Final Supplemental Environmental Impact Statement for Airborne Laser Program at Kirtland Air Force Base (AFB) and White Sands Missile Range/Holloman AFB, NM, and Edwards AFB and Vandenberg AFB, CA

    DTIC Science & Technology

    2003-06-01

    californicus- _" -~ T ~ - 1" T - E I T Coho salmon Unarmoured three- spined stickleback I Tidewater goby 1 Eucyclogobius n e w b e p i ,.I...ENDANGERED Black-footed ferret (Mustela nigripes)"" Northern aplomado falcon (Falcofemoralis septentrionalis) Kuenzler hedgehog cachls (Echi...Empidonax traillii ertimus) Kuenzler hedgehog cactus (Echinocer~.~rendleri var. kuenzleri) Todsen’s pennyroyal (Hedeoma fodsenii) Sacramento

  2. Active Research in the Age of Reconciliation: A Relationship-Based Approach for Non-Indigenous Researchers

    ERIC Educational Resources Information Center

    Carreau, Robert; Robinson, Daniel B.

    2017-01-01

    This article recounts a five-year journey connecting Acadian and Mi'kmaw schools in Nova Scotia, Canada. We trace the unfolding of relationships between minority school students and administrators through both natural and semi-structured dialogues. We believe this work contributes to the emerging field of research called for by the Truth and…

  3. BASIC EDUCATION FOR SPANISH-SPEAKING DISADVANTAGED PUPILS.

    ERIC Educational Resources Information Center

    OTT, ELIZABETH H.

    A BILINGUAL EDUCATION PROGRAM HAS BEEN FIELD TESTED IN SAN ANTONIO, TEXAS, IN CLASSES OF DISADVANTAGED MEXICAN-AMERICAN CHILDREN. THIS SOUTHWEST EDUCATIONAL DEVELOPMENT LABORATORY (SWEDL) PROGRAM IS ALSO APPLICABLE TO TEACHING FRENCH ACADIANS AND NEGRO AMERICANS. THE STRENGTH OF THE PROGRAM, ACCORDING TO THE AUTHOR, LIES IN WHAT IT DOES TO CHANGE…

  4. Leaf area prediction models for Tsuga canadensis in Maine

    Treesearch

    Laura S. Kenefic; R.S. Seymour

    1999-01-01

    Tsuga canadensis (L.) Carr. (eastern hemlock) is a common species throughout the Acadian forest. Studies of leaf area and growth efficiency in this forest type have been limited by the lack of equations to predict leaf area of this species. We found that sapwood area was an effective leaf area surrogate in T. canadensis, though...

  5. 78 FR 2249 - Magnuson-Stevens Act Provisions; Fisheries of the Northeastern United States; Northeast...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2013-01-10

    ...This action reopens the comment period for an Acadian redfish- related proposed rule that published on November 8, 2012. The original comment period closed on November 23, 2012. This action clarifies a bycatch threshold incorrectly explained in the proposed rule. The public comment period is being reopened to solicit additional public comment on this correction.

  6. Timing of terrane accretion in eastern and east-central Maine

    NASA Astrophysics Data System (ADS)

    Ludman, Allan

    1986-05-01

    The Norumbega fault zone is often cited as a post-Acadian suture between exotic blocks, even though stratigraphic, structural, and metamorphic data indicate that there is little offset of the Silurian-Devonian strata that the zone cuts in eastern Maine. Similarly, the Kingman fault zone has been shown by gravity and geochemical studies to separate distinct crustal blocks, whereas mapping shows that it lies entirely within a Silurian turbidite package. These conflicts are resolved if the two fault zones represent boundaries between Ordovician or older crustal blocks that had accreted to form a composite terrane prior to deposition of the cover sequences. The faults now mapped within these younger rocks formed by reactivation of the pre-Silurian boundaries during late Acadian time; movement continued until the late Carboniferous. Most of the accretionary history of Maine had thus ended before the Silurian. A complex composite terrane may have formed during Cambrian-Ordovician time that (1) interacted with cratonic North America during the Taconian orogeny and (2) became the “basement” upon which the Silurian and Lower Devonian strata of eastern Maine were deposited.

  7. Comparison of statistical and theoretical habitat models for conservation planning: the benefit of ensemble prediction

    USGS Publications Warehouse

    Jones-Farrand, D. Todd; Fearer, Todd M.; Thogmartin, Wayne E.; Thompson, Frank R.; Nelson, Mark D.; Tirpak, John M.

    2011-01-01

    Selection of a modeling approach is an important step in the conservation planning process, but little guidance is available. We compared two statistical and three theoretical habitat modeling approaches representing those currently being used for avian conservation planning at landscape and regional scales: hierarchical spatial count (HSC), classification and regression tree (CRT), habitat suitability index (HSI), forest structure database (FS), and habitat association database (HA). We focused our comparison on models for five priority forest-breeding species in the Central Hardwoods Bird Conservation Region: Acadian Flycatcher, Cerulean Warbler, Prairie Warbler, Red-headed Woodpecker, and Worm-eating Warbler. Lacking complete knowledge on the distribution and abundance of each species with which we could illuminate differences between approaches and provide strong grounds for recommending one approach over another, we used two approaches to compare models: rank correlations among model outputs and comparison of spatial correspondence. In general, rank correlations were significantly positive among models for each species, indicating general agreement among the models. Worm-eating Warblers had the highest pairwise correlations, all of which were significant (P , 0.05). Red-headed Woodpeckers had the lowest agreement among models, suggesting greater uncertainty in the relative conservation value of areas within the region. We assessed model uncertainty by mapping the spatial congruence in priorities (i.e., top ranks) resulting from each model for each species and calculating the coefficient of variation across model ranks for each location. This allowed identification of areas more likely to be good targets of conservation effort for a species, those areas that were least likely, and those in between where uncertainty is higher and thus conservation action incorporates more risk. Based on our results, models developed independently for the same purpose

  8. Sixty years of research, 60 years of data: long-term US Forest Service data management on the Penobscot Experimental Forest

    Treesearch

    Matthew B. Russell; Spencer R. Meyer; John C. Brissette; Laura Kenefic

    2014-01-01

    The U.S. Department of Agriculture, Forest Service silvicultural experiment on the Penobscot Experimental Forest (PEF) in Maine represents 60 years of research in the northern conifer and mixedwood forests of the Acadian Forest Region. The objective of this data management effort, which began in 2008, was to compile, organize, and archive research data collected in the...

  9. Growth comparison of northern white-cedar to balsam fir and red spruce by site class

    Treesearch

    Philip V. Hofmeyer; Laura S. Kenefic; Robert S. Seymour; John C. Brissette

    2006-01-01

    Though northern white-cedar is a common and economically important component of the Acadian Forest of Maine and adjacent Canada, there is little regional data about the growth and development of this species. Sixty sites in northern Maine were used to compare growth of cedar to that of red spruce and balsam fir along a range of site classes and light exposures. On...

  10. Bedrock geologic and joint trend map of the Pinardville quadrangle, Hillsborough County, New Hampshire

    USGS Publications Warehouse

    Burton, William C.; Armstrong, Thomas R.

    2013-01-01

    The bedrock geology of the Pinardville quadrangle includes the Massabesic Gneiss Complex, exposed in the core of a regional northeast-trending anticlinorium, and highly deformed metasedimentary rocks of the Rangeley Formation, exposed along the northwest limb of the anticlinorium. Both formations were subjected to high-grade metamorphism and partial melting: the Rangeley during the middle Paleozoic Acadian orogeny, and the Massabesic Gneiss Complex during both the Acadian and the late Paleozoic Alleghanian orogeny. Granitoids produced during these orogenies range in age from Devonian (Spaulding Tonalite) to Permian (granite at Damon Pond), each with associated pegmatite. In the latest Paleozoic the Massabesic Gneiss Complex was uplifted with respect to the Rangeley Formation along the ductile Powder Hill fault, which also had a left-lateral component. Uplift continued into the early Mesozoic, producing the 2-kilometer-wide Campbell Hill fault zone, which is marked by northwest-dipping normal faults and dilational map-scale quartz bodies. Rare, undeformed Jurassic diabase dikes cut all older lithologies and structures. A second map is a compilation of joint orientations measured at all outcrops in the quadrangle. There is a great diversity of strike trends, with northeast perhaps being the most predominant.

  11. 75 FR 9281 - General Provisions; Revised List of Migratory Birds

    Federal Register 2010, 2011, 2012, 2013, 2014

    2010-03-01

    ... York (AOU 1982, 1983, 1998); Flycatcher, La Sagra's, Myiarchus sagrae--Alabama, Florida (AOU 1982, 1983..., Belcher's, Larus belcheri--Florida (AOU 1982, 1983, 1998, 2003); Hawk, Roadside, Buteo magnirostis--Texas..., Southern, Progne elegans--Florida (AOU 1982, 1983, 1998); Mockingbird, Bahama, Mimus gundlachii--Florida...

  12. Organic Chemical Concentrations in Eggs and Nestlings of Cavity Nesting Birds at and around Los Alamos National Laboratory

    DOE PAGES

    Gaukler, Shannon M.; Hathcock, Charles D.; Fair, Jeanne M.

    2018-02-13

    In 1943, Los Alamos National Laboratory (LANL) was established as part of the Manhattan project to design atomic weapons. LANL now operates as a multidisciplinary research institution. As part of an ongoing assessment of siterelated ecological risk, organochlorine pesticides, their metabolites, polycyclic aromatic hydrocarbons, polychlorinated biphenyls (PCBs), and 2,3,7,8-tetrachlorodibenzo-p-dioxin toxic equivalents (TEQs) were evaluated in western bluebird (Sialia mexicana) and ash-throated flycatcher (Myiarchus cinerascens) eggs relative to a developed but non-industrial reference area; PCBs and TEQs were also evaluated in nestlings. Chemicals were below detection limits in the majority of samples. Western bluebird eggs collected from the study area hadmore » significantly lower concentrations of dieldrin, oxychlordane, and trans-nonachlor when compared with eggs from the reference area. No differences were observed in concentrations of dichlorodiphenyldichloroethylene (DDE), dichlorodiphenyltrichloroethane (DDT), and heptachlor epoxide. Ash-throated flycatcher eggs contained higher total TEQ concentrations when compared with western bluebird eggs; however, no differences in concentrations of DDE, DDT, dieldrin, or total PCBs were observed. No differences were observed in total PCBs or TEQs in nestlings between the two species. Western bluebird eggs contained higher levels of total PCBs and TEQs when compared with nestlings; no differences were observed in total PCBs or TEQs between ash-throated flycatcher eggs and nestlings. Chemical concentrations detected in eggs of both species were below levels that are associated with adverse effects reported in the scientific literature, suggesting that concentrations of organic chemicals observed here appear to be at levels causing negligible risks to local bird populations.« less

  13. Organic Chemical Concentrations in Eggs and Nestlings of Cavity Nesting Birds at and around Los Alamos National Laboratory

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Gaukler, Shannon M.; Hathcock, Charles D.; Fair, Jeanne M.

    In 1943, Los Alamos National Laboratory (LANL) was established as part of the Manhattan project to design atomic weapons. LANL now operates as a multidisciplinary research institution. As part of an ongoing assessment of siterelated ecological risk, organochlorine pesticides, their metabolites, polycyclic aromatic hydrocarbons, polychlorinated biphenyls (PCBs), and 2,3,7,8-tetrachlorodibenzo-p-dioxin toxic equivalents (TEQs) were evaluated in western bluebird (Sialia mexicana) and ash-throated flycatcher (Myiarchus cinerascens) eggs relative to a developed but non-industrial reference area; PCBs and TEQs were also evaluated in nestlings. Chemicals were below detection limits in the majority of samples. Western bluebird eggs collected from the study area hadmore » significantly lower concentrations of dieldrin, oxychlordane, and trans-nonachlor when compared with eggs from the reference area. No differences were observed in concentrations of dichlorodiphenyldichloroethylene (DDE), dichlorodiphenyltrichloroethane (DDT), and heptachlor epoxide. Ash-throated flycatcher eggs contained higher total TEQ concentrations when compared with western bluebird eggs; however, no differences in concentrations of DDE, DDT, dieldrin, or total PCBs were observed. No differences were observed in total PCBs or TEQs in nestlings between the two species. Western bluebird eggs contained higher levels of total PCBs and TEQs when compared with nestlings; no differences were observed in total PCBs or TEQs between ash-throated flycatcher eggs and nestlings. Chemical concentrations detected in eggs of both species were below levels that are associated with adverse effects reported in the scientific literature, suggesting that concentrations of organic chemicals observed here appear to be at levels causing negligible risks to local bird populations.« less

  14. Cultural Resources Survey of Three Iberville Parish Levee Enlargement and Revetment Construction Items

    DTIC Science & Technology

    1993-09-22

    SURVEY OF THREE IBERVILLE PARISH LEVEE ENLARGEMENT AND REVETMENT CONSTRUCTION ITEMS September 1993 Sam .4 D2 FINAL REPORT E R. Christopher Goodwin...LEVEE ENLARGEMENT ANj REVETMENT CONSTRUCTION ITEMS 12. PERSONAL AUTHOR(S) R. Christopher Goodwin, Ph.d., Rebecca E. Bruce, Lawrence L Hewitt, and E... block number) FIELD GROUP SUB-GROUP Acadian Coast Historic Arche6cogy Rice Antebellum Iberville Parish Saw Mill Plantation Carville Leprosarium Ophelia

  15. Biogeography of dinoflagellate cysts in northwest Atlantic ...

    EPA Pesticide Factsheets

    Few biogeographic studies of dinoflagellate cysts include the near-shore estuarine environment. We determine the effect of estuary type, biogeography, and water quality on the spatial distribution of organic-walled dinoflagellate cysts from the Northeast USA (Maine to Delaware) and Canada (Prince Edward Island). A total of 69 surface sediment samples were collected from 27 estuaries, from sites with surface salinities >20. Dinoflagellate cysts were examined microscopically and compared to environmental parameters using multivariate ordination techniques. The spatial distribution of cyst taxa reflects biogeographic provinces established by other marine organisms, with Cape Cod separating the northern Acadian Province from the southern Virginian Province. Species such as Lingulodinium machaerophorum and Polysphaeridinium zoharyi were found almost exclusively in the Virginian Province, while others such as Dubridinium spp. and Islandinium? cezare were more abundant in the Acadian Province. Tidal range, sea surface temperature (SST), and sea surface salinity (SSS) are statistically significant parameters influencing cyst assemblages. Samples from the same type of estuary cluster together in canonical correspondence analysis when the estuaries are within the same biogeographic province. The large geographic extent of this study, encompassing four main estuary types (riverine, lagoon, coastal embayment, and fjord), allowed us to determine that the type of estuary has

  16. Telescoping metamorphic isograds: Evidence from 40Ar/39A dating in the Orange-Milford belt, southern Connecticut

    USGS Publications Warehouse

    Kunk, Michael J.; Walsh, Gregory J.; Growdon, Martha L.; Wintsch, Robert P.

    2013-01-01

    New 40Ar/39Ar ages for hornblende and muscovite from the Orange-Milford belt in southern Connecticut reflect cooling from Acadian amphibolite facies metamorphism between ∼380 to 360 Ma followed by retrograde recrystallization of fabric-forming muscovite and chlorite during lower greenschist facies Alleghanian transpression at ∼280 Ma. Reported field temperature and pressure gradients are improbably high for these rocks and a NW metamorphic field gradient climbing from chlorite-grade to staurolite-grade occurs over less than 5 km. Simple tilting cannot account for this compressed isograd spacing given the geothermal gradient of ∼20 °C/km present at the time of regional metamorphism. However, post-metamorphic transpression could effectively telescope the isograds by stretching the belt at an oblique angle to the isograd traces. Textures in the field and in thin section reveal several older prograde schistosities overprinted by lower greenschist facies fabrics. The late cleavages commonly occur at the scale of ∼100 μm and these samples contain multiple age populations of white mica. 40Ar/39Ar analysis of these poly-metamorphic samples with mixed muscovite populations yield climbing or U-shaped age spectra. The ages of the low temperature steps are late Paleozoic, while the ages of the older steps are late Devonian. These results support our petrologic interpretation that the younger cleavage developed under metamorphic conditions below the closure temperature for Ar diffusion in muscovite, that is, in the lower greenschist facies. The correlation of a younger regionally reproducible age population with a pervasive retrograde muscovite ± chlorite cleavage reveals an Alleghanian (∼280 Ma) overprint on the Acadian metamorphic gradient (∼380 Ma). Outcrop-scale structures including drag folds and imbricate boudins suggest that Alleghanian deformation and cleavage development occurred in response to dextral transpression along a northeast striking boundary

  17. 77 FR 75646 - Kenai National Wildlife Refuge, Soldotna, AK; Draft Environmental Impact Statement for the...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2012-12-21

    ..., and Cook Inlet Region, Inc. (CIRI), owns the subsurface estate of coal, oil, and gas in the project..., snowshoe hares, and numerous species of Neotropical birds, such as olive-sided flycatchers, myrtle warblers... received the subsurface oil, gas, and coal estate to nearly 200,000 acres within the Refuge as part of its...

  18. A Field Guide to Common Educator Birds.

    ERIC Educational Resources Information Center

    Smock, Judith N.

    1990-01-01

    Teaching may be for the birds, but many educator birds have been abandoning the field of late for downier nests. Several species still remain, including the crown-tufted superintendent bird, the red-throated assistant superintendent bird, the crested fly-catcher principal bird, the exotic scenery bird, the roadrunner psychologist bird, and the…

  19. Silence is not golden: the hissing calls of tits affect the behaviour of a nest predator.

    PubMed

    Zub, Karol; Czeszczewik, Dorota; Ruczyński, Ireneusz; Kapusta, Anna; Walankiewicz, Wiesław

    2017-01-01

    Nest predation is one of the most important mortality factors of birds. Field observations showed that tits (Paridae) produce hissing calls and, usually, have lower breeding losses than nesting Ficedula flycatchers, which do not make such calls. We hypothesise that differences in fledgling success can be directly attributed to the vocal reaction of tits. We tested experimentally whether the hissing calls can affect the behaviour of a potential predator, analysing the response of the Yellow-necked Mouse Apodemus flavicollis to playback of calls of three Parid species. The number of visits by mice to two types of cavities (with playback and control) was not significantly different, but the average time spent by mice in cavities with playback (3.9 s) was significantly shorter than in cavities without playback (26.3 s). This suggests that hissing behaviour of tits significantly changes the exploration activity of predators, which may ultimately increase the breeding success of this group of birds relative to the flycatchers. Nest predation is one of the most important mortality factors of small land birds, but some anti-predatory mechanisms are still poorly recognised. Numerous studies demonstrate that incubating tits make hissing sounds, when a predator is near, but despite almost a century of research, there is little evidence these calls indeed affect behaviour of predators. By using a simple laboratory experiment, we demonstrated that the hissing acoustic signals used by tits may change the behaviour of yellow-necked mice, which are an important predator of cavity-nesting birds in temperate forests. Intruding mice withdrew from cavities where hissing sounds were played back. Our results suggest that the hissing behaviour of tits can change the exploration activity of potential predators and may increase breeding success of this group of birds relative to the flycatchers, which stay silent when their nest is threatened.

  20. Bird populations in logged and unlogged western larch/Douglas-fir forest in northwestern Montana

    Treesearch

    Bret W. Tobalske; Raymond C. Shearer; Richard L. Hutto

    1991-01-01

    Of 32 species of abundant breeding birds, populations of 10 species differed significantly between small cutting units and adjacent uncut forest. Foliage foragers and tree gleaners were less abundant in cutting units, while flycatching species and ground foragers were more common there. Of nesting guilds, conifer tree nesters were least abundant in cutting units, and...

  1. Chapter 10: Management recommendations

    Treesearch

    Deborah M. Finch; Janie Agyagos; Tracy McCarthey; Robert M. Marshall; Scott H. Stoleson; Mary J. Whitfield

    2000-01-01

    This chapter was developed over a series of meetings using a group-consensus process. Our recommendations are based on published results, on information compiled in the previous chapters, on expert opinion, and on unpublished data of conservation team members. This chapter is available as temporary guidance until the Recovery Plan for the southwestern willow flycatcher...

  2. Bacteria-killing ability of fresh blood plasma compared to frozen blood plasma

    DOE PAGES

    Jacobs, Anne C.; Fair, Jeanne Marie

    2015-10-09

    In recent years, the bacteria-killing assay (BKA) has become a popular technique among ecoimmunologists. New variations of that assay allow researchers to use smaller volumes of blood, an important consideration for those working on small-bodied animals. However, this version of the assay requires access to a lab with a nanodrop spectrophotometer, something that may not be available in the field. One possible solution is to freeze plasma for transport; however, this assumes that frozen plasma samples will give comparable results to fresh ones. Here, we tested this assumption using plasma samples from three species of birds: chickens (Gallus gallus), ash-throatedmore » flycatchers (Myiarchus cinerascens), and western bluebirds (Sialia mexicana). Chicken plasma samples lost most or all of their bacterial killing ability after freezing. This did not happen in flycatchers and bluebirds; however, frozen plasma did not produce results comparable to those obtained using fresh plasma. Finally, we caution researchers using the BKA to use fresh samples whenever possible, and to validate the use of frozen samples on a species-by-species basis.« less

  3. Bacteria-killing ability of fresh blood plasma compared to frozen blood plasma.

    PubMed

    Jacobs, Anne C; Fair, Jeanne M

    2016-01-01

    In recent years, the bacteria-killing assay (BKA) has become a popular technique among ecoimmunologists. New variations of that assay allow researchers to use smaller volumes of blood, an important consideration for those working on small-bodied animals. However, this version of the assay requires access to a lab with a nanodrop spectrophotometer, something that may not be available in the field. One possible solution is to freeze plasma for transport; however, this assumes that frozen plasma samples will give comparable results to fresh ones. We tested this assumption using plasma samples from three species of birds: chickens (Gallus gallus), ash-throated flycatchers (Myiarchus cinerascens), and western bluebirds (Sialia mexicana). Chicken plasma samples lost most or all of their bacterial killing ability after freezing. This did not happen in flycatchers and bluebirds; however, frozen plasma did not produce results comparable to those obtained using fresh plasma. We caution researchers using the BKA to use fresh samples whenever possible, and to validate the use of frozen samples on a species-by-species basis. Copyright © 2015 Elsevier Inc. All rights reserved.

  4. Bacteria-killing ability of fresh blood plasma compared to frozen blood plasma

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Jacobs, Anne C.; Fair, Jeanne Marie

    In recent years, the bacteria-killing assay (BKA) has become a popular technique among ecoimmunologists. New variations of that assay allow researchers to use smaller volumes of blood, an important consideration for those working on small-bodied animals. However, this version of the assay requires access to a lab with a nanodrop spectrophotometer, something that may not be available in the field. One possible solution is to freeze plasma for transport; however, this assumes that frozen plasma samples will give comparable results to fresh ones. Here, we tested this assumption using plasma samples from three species of birds: chickens (Gallus gallus), ash-throatedmore » flycatchers (Myiarchus cinerascens), and western bluebirds (Sialia mexicana). Chicken plasma samples lost most or all of their bacterial killing ability after freezing. This did not happen in flycatchers and bluebirds; however, frozen plasma did not produce results comparable to those obtained using fresh plasma. Finally, we caution researchers using the BKA to use fresh samples whenever possible, and to validate the use of frozen samples on a species-by-species basis.« less

  5. Chapter 2: Population status on breeding grounds

    Treesearch

    Robert M. Marshall

    2000-01-01

    In his review of the overall population status of the southwestern willow flycatcher, Unitt (1987) concluded that "Even if a few hundred pairs persist in New Mexico, the total population of the subspecies is well under 1000 pairs; I suspect 500 is more likely." Since Unitt's review, a substantial amount of information has been collected rangewide on the...

  6. The role of indicator species: Neotropical migratory song birds

    Treesearch

    Theodore R. Simons; Kerry N. Rabenold; David A. Buehler; Jaime A. Collazo; Kathleen E. Franzreb

    1999-01-01

    Southern Appalachian forests support some of the richest avian diversity in North America, including some 75 species of Neotropical migrants, birds that perform the remarkable feat of making much of the Western Hemisphere their home. This diverse group includes the swallows, kingbirds, and other flycatchers that feed in the air on flying insects. The Eastern kingbird...

  7. The Role of MC1R in Speciation & Phylogeny

    ERIC Educational Resources Information Center

    Offner, Susan

    2013-01-01

    A point mutation in the MC1R gene, a G-protein-coupled receptor, has been found that could have led to the formation of two subspecies of Solomon Island flycatcher from a single ancestral population. I discuss the many roles that G-protein-coupled receptors play in vertebrate physiology and how one particular point mutation can have enormous…

  8. Chapter 11: Research needs

    Treesearch

    Scott H. Stoleson; Janie Agyagos; Deborah M. Finch; Tracy McCarthey; Jamie Uyehara; Mary J. Whitfield

    2000-01-01

    Until the southwestern willow flycatcher was proposed for listing as an endangered species in 1993, it was subject to relatively little scientific scrutiny, in part because it is drab, prone to inhabiting dense, wet thickets, and difficult to identify and observe. Consequently, it remained one of the lesser-known of North American birds. Since that time, it has become...

  9. Environmental Assessment for Demolition of Building 2148 and Construction of a New Facility

    DTIC Science & Technology

    2004-02-01

    Athene cunicularia SP Caspian Tern Sterna caspia SP Ferruginous Hawk Buteo regalis T Grasshopper Sparrow Ammodramus savannarum SP/SD Lewis’ Woodpecker...northern latitudes winter along streams and lakes throughout Utah and have been observed at the Weber River just north of the Base. Willow Flycatchers...Pelecanus erythrorhynchos SD Black Tern Chlidonias niger SP Blue Grosbeak Guiraca caerulea SP/SD Bobolink Dolichonyx oryzivorus SP/SD Burrowing Owl

  10. Legacy Bird Species at Risk Monitoring in and Around Camp Navajo and the Naval Observatory Flagstaff Station, AZ

    DTIC Science & Technology

    2010-04-16

    19th century due to a convergence of complex stressors such as overgrazing, timber harvest, drought , and fire suppression (Allen et al. 2002, Moore... pinus 16 Olive Warbler Peucedramus taeniatus 5 Plumbeous Vireo Vireo solitarius 60 Pygmy Nuthatch Sitta pygmaea 177 Red Crossbill Loxia...Dove Zenaida macroura 14 Northern Flicker Colaptes auratus 50 Pine Siskin Carduelis pinus 58 Olive-sided Flycatcher Contopus cooperi 1

  11. Natal habitat imprinting counteracts the diversifying effects of phenotype-dependent dispersal in a spatially structured population.

    PubMed

    Camacho, Carlos; Canal, David; Potti, Jaime

    2016-08-08

    Habitat selection may have profound evolutionary consequences, but they strongly depend on the underlying preference mechanism, including genetically-determined, natal habitat and phenotype-dependent preferences. It is known that different mechanisms may operate at the same time, yet their relative contribution to population differentiation remains largely unexplored empirically mainly because of the difficulty of finding suitable study systems. Here, we investigate the role of early experience and genetic background in determining the outcome of settlement by pied flycatchers (Ficedula hypoleuca) breeding in two habitat patches between which dispersal and subsequent reproductive performance is influenced by phenotype (body size). For this, we conducted a cross-fostering experiment in a two-patch system: an oakwood and a conifer plantation separated by only 1 km. Experimental birds mostly returned to breed in the forest patch where they were raised, whether it was that of their genetic or their foster parents, indicating that decisions on where to settle are determined by individuals' experience in their natal site, rather than by their genetic background. Nevertheless, nearly a third (27.6 %) moved away from the rearing habitat and, as previously observed in unmanipulated individuals, dispersal between habitats was phenotype-dependent. Pied flycatchers breeding in the oak and the pine forests are differentiated by body size, and analyses of genetic variation at microsatellite loci now provide evidence of subtle genetic differentiation between the two populations. This suggests that phenotype-dependent dispersal may contribute to population structure despite the short distance and widespread exchange of birds between the study plots. Taken together, the current and previous findings that pied flycatchers do not always settle in the habitat to which they are best suited suggest that their strong tendency to return to the natal patch regardless of their body size

  12. Interspecific variation in the relationship between clutch size, laying date and intensity of urbanization in four species of hole-nesting birds.

    PubMed

    Vaugoyeau, Marie; Adriaensen, Frank; Artemyev, Alexandr; Bańbura, Jerzy; Barba, Emilio; Biard, Clotilde; Blondel, Jacques; Bouslama, Zihad; Bouvier, Jean-Charles; Camprodon, Jordi; Cecere, Francesco; Charmantier, Anne; Charter, Motti; Cichoń, Mariusz; Cusimano, Camillo; Czeszczewik, Dorota; Demeyrier, Virginie; Doligez, Blandine; Doutrelant, Claire; Dubiec, Anna; Eens, Marcel; Eeva, Tapio; Faivre, Bruno; Ferns, Peter N; Forsman, Jukka T; García-Del-Rey, Eduardo; Goldshtein, Aya; Goodenough, Anne E; Gosler, Andrew G; Grégoire, Arnaud; Gustafsson, Lars; Harnist, Iga; Hartley, Ian R; Heeb, Philipp; Hinsley, Shelley A; Isenmann, Paul; Jacob, Staffan; Juškaitis, Rimvydas; Korpimäki, Erkki; Krams, Indrikis; Laaksonen, Toni; Lambrechts, Marcel M; Leclercq, Bernard; Lehikoinen, Esa; Loukola, Olli; Lundberg, Arne; Mainwaring, Mark C; Mänd, Raivo; Massa, Bruno; Mazgajski, Tomasz D; Merino, Santiago; Mitrus, Cezary; Mönkkönen, Mikko; Morin, Xavier; Nager, Ruedi G; Nilsson, Jan-Åke; Nilsson, Sven G; Norte, Ana C; Orell, Markku; Perret, Philippe; Perrins, Christopher M; Pimentel, Carla S; Pinxten, Rianne; Richner, Heinz; Robles, Hugo; Rytkönen, Seppo; Senar, Juan Carlos; Seppänen, Janne T; Pascoal da Silva, Luis; Slagsvold, Tore; Solonen, Tapio; Sorace, Alberto; Stenning, Martyn J; Tryjanowski, Piotr; von Numers, Mikael; Walankiewicz, Wieslaw; Møller, Anders Pape

    2016-08-01

    The increase in size of human populations in urban and agricultural areas has resulted in considerable habitat conversion globally. Such anthropogenic areas have specific environmental characteristics, which influence the physiology, life history, and population dynamics of plants and animals. For example, the date of bud burst is advanced in urban compared to nearby natural areas. In some birds, breeding success is determined by synchrony between timing of breeding and peak food abundance. Pertinently, caterpillars are an important food source for the nestlings of many bird species, and their abundance is influenced by environmental factors such as temperature and date of bud burst. Higher temperatures and advanced date of bud burst in urban areas could advance peak caterpillar abundance and thus affect breeding phenology of birds. In order to test whether laying date advance and clutch sizes decrease with the intensity of urbanization, we analyzed the timing of breeding and clutch size in relation to intensity of urbanization as a measure of human impact in 199 nest box plots across Europe, North Africa, and the Middle East (i.e., the Western Palearctic) for four species of hole-nesters: blue tits (Cyanistes caeruleus), great tits (Parus major), collared flycatchers (Ficedula albicollis), and pied flycatchers (Ficedula hypoleuca). Meanwhile, we estimated the intensity of urbanization as the density of buildings surrounding study plots measured on orthophotographs. For the four study species, the intensity of urbanization was not correlated with laying date. Clutch size in blue and great tits does not seem affected by the intensity of urbanization, while in collared and pied flycatchers it decreased with increasing intensity of urbanization. This is the first large-scale study showing a species-specific major correlation between intensity of urbanization and the ecology of breeding. The underlying mechanisms for the relationships between life history and

  13. An autochthonous Avalonian basement source for the latest Ordovician Brenton Pluton in the Meguma terrane of Nova Scotia: U-Pb-Hf isotopic constraints and paleogeographic implications

    NASA Astrophysics Data System (ADS)

    Duncan Keppie, J.; Gregory Shellnutt, J.; Dostal, Jaroslav; Fraser Keppie, D.

    2018-04-01

    The Ediacaran-Ordovician Meguma Supergroup was thrust over Avalonia basement prior to the intrusion of post-Acadian, ca. 370 Ma, S-type granitic batholiths. This has led to two main hypotheses regarding the original location of the Meguma terrane, a continental rise prism bordering either NW Africa or Avalonia. On the other hand, the pre-Acadian, ca. 440 Ma Brenton pluton has yielded the following U/Pb LA-ICP-MS zircon data: (1) 448 ± 3 Ma population peak inferred to be the intrusive age and (2) ca. 550 and 700 Ma inherited ages common to both Avalonia and NW Africa. In contrast, Hf isotopic analyses of zircon yielded model ages ranging from 814 to 1127 Ma with most between 940 and 1040 Ma: such ages are typical of Avalonia and not NW Africa. The ages of the inherited zircons found within the Brenton pluton suggest that it was probably derived by partial melting of sub-Meguma, mid-crustal Avalonian rocks, upon which the Meguma Supergroup was deposited. Although Avalonia is commonly included in the peri-Gondwanan terranes off NW Africa or Amazonia, paleomagnetic data, faunal provinciality, and Hf data suggest that, during the Ediacaran-Early Cambrian, it was an island chain lying near the tropics (ca. 20-30 °S) and was possibly a continuation of the Bolshezemel volcanic arc accreted to northern Baltica during the Ediacaran Timanide orogenesis. This is consistent with the similar derital zircon population in the Ediacaran-Cambrian Meguma Supergroup and the Dividal Group in northeastern Baltica.

  14. Geometry of an outcrop-scale duplex in Devonian flysch, Maine

    USGS Publications Warehouse

    Bradley, D.C.; Bradley, L.M.

    1994-01-01

    We describe an outcrop-scale duplex consisting of 211 exposed repetitions of a single bed. The duplex marks an early Acadian (Middle Devonian) oblique thrust zone in the Lower Devonian flysch of northern Maine. Detailed mapping at a scale of 1:8 has enabled us to measure accurately parameters such as horse length and thickness, ramp angles and displacements; we compare these and derivative values with those of published descriptions of duplexes, and with theoretical models. Shortening estimates based on line balancing are consistently smaller than two methods of area balancing, suggesting that layer-parallel shortening preceded thrusting. ?? 1994.

  15. Late cretaceous extensional tectonics and associated igneous activity on the northern margin of the Gulf of Mexico Basin

    NASA Technical Reports Server (NTRS)

    Bowen, R. L.; Sundeen, D. A.

    1985-01-01

    Major, dominantly compressional, orogenic episodes (Taconic, Acadian, Alleghenian) affected eastern North America during the Paleozoic. During the Mesozoic, in contrast, this same region was principally affected by epeirogenic and extensional tectonism; one episode of comparatively more intense tectonic activity involving extensive faulting, uplift, sedimentation, intrusion and effusion produced the Newark Series of eposits and fault block phenomena. This event, termed the Palisades Disturbance, took place during the Late Triassic - Earliest Jurassic. The authors document a comparable extensional tectonic-igneous event occurring during the Late Cretaceous (Early Gulfian; Cenomanian-Santonian) along the southern margin of the cratonic platform from Arkansas to Georgia.

  16. Sex chromosome linked genetic variance and the evolution of sexual dimorphism of quantitative traits.

    PubMed

    Husby, Arild; Schielzeth, Holger; Forstmeier, Wolfgang; Gustafsson, Lars; Qvarnström, Anna

    2013-03-01

    Theory predicts that sex chromsome linkage should reduce intersexual genetic correlations thereby allowing the evolution of sexual dimorphism. Empirical evidence for sex linkage has come largely from crosses and few studies have examined how sexual dimorphism and sex linkage are related within outbred populations. Here, we use data on an array of different traits measured on over 10,000 individuals from two pedigreed populations of birds (collared flycatcher and zebra finch) to estimate the amount of sex-linked genetic variance (h(2)z ). Of 17 traits examined, eight showed a nonzero h(2)Z estimate but only four were significantly different from zero (wing patch size and tarsus length in collared flycatchers, wing length and beak color in zebra finches). We further tested how sexual dimorphism and the mode of selection operating on the trait relate to the proportion of sex-linked genetic variance. Sexually selected traits did not show higher h(2)Z than morphological traits and there was only a weak positive relationship between h(2)Z and sexual dimorphism. However, given the relative scarcity of empirical studies, it is premature to make conclusions about the role of sex chromosome linkage in the evolution of sexual dimorphism. © 2012 The Author(s). Evolution© 2012 The Society for the Study of Evolution.

  17. DNA-DNA hybridization-based phylogeny for "higher" nonpasserines: reevaluating a key portion of the avian family tree.

    PubMed

    Bleiweiss, R; Kirsch, J A; Lapointe, F J

    1994-09-01

    A matrix of delta T mode values for 10 birds, including 9 nonpasserines and a suboscine passerine flycatcher, was generated by DNA-DNA hybridization. Within the most derived lineages, all bootstrapped and jackknifed FITCH trees lend strong support to sister-groupings of the two swift families, of hummingbirds to swifts, and of these to a clade containing both owls and night-hawks. The outgroup duck roots the tree between the woodpecker (Piciformes) and the remaining taxa, indicating that Piciformes are among the earliest branches within nonpasserines. However, the succeeding branches to kingfisher, mousebird, and suboscine passerine flycatcher are based on short internodes that are poorly supported by bootstrapping and that give inconsistent results in jackknifing. Although these 3 orders may have arisen through rapid or near-simultaneous divergence, placement of the "advanced" Passeriformes deep within a more "primitive" radiation indicates that nonpasserines are paraphyletic, echoing the same distinction for reptiles with respect to their advanced descendants. Despite significant rate variation among different taxa, these results largely concur with those obtained with the same technique by Sibley and Ahlquist, who used the delta T50H measure and UPGMA analysis. This agreement lends credence to some of their more controversial claims.

  18. Effects of nocturnal illumination on life-history decisions and fitness in two wild songbird species

    PubMed Central

    de Jong, Maaike; Ouyang, Jenny Q.; Da Silva, Arnaud; van Grunsven, Roy H. A.; Kempenaers, Bart; Visser, Marcel E.; Spoelstra, Kamiel

    2015-01-01

    The effects of artificial night lighting on animal behaviour and fitness are largely unknown. Most studies report short-term consequences in locations that are also exposed to other anthropogenic disturbance. We know little about how the effects of nocturnal illumination vary with different light colour compositions. This is increasingly relevant as the use of LED lights becomes more common, and LED light colour composition can be easily adjusted. We experimentally illuminated previously dark natural habitat with white, green and red light, and measured the effects on life-history decisions and fitness in two free-living songbird species, the great tit (Parus major) and pied flycatcher (Ficedula hypoleuca) in two consecutive years. In 2013, but not in 2014, we found an effect of light treatment on lay date, and of the interaction of treatment and distance to the nearest lamp post on chick mass in great tits but not in pied flycatchers. We did not find an effect in either species of light treatment on breeding densities, clutch size, probability of brood failure, number of fledglings and adult survival. The finding that light colour may have differential effects opens up the possibility to mitigate negative ecological effects of nocturnal illumination by using different light spectra. PMID:25780240

  19. Nd, Sr and O isotopic study of the petrogenesis of two syntectonic members of the New Hampshire Plutonic Series

    NASA Astrophysics Data System (ADS)

    Lathrop, A. S.; Blum, Joel D.; Chamberlain, C. Page

    1996-07-01

    Nd, Sr and O isotope systematics were used to investigate the petrogenesis of two adjacent plutons of the Bethlehem Gneiss (BG) and the Kinsman Quartz Monzonite (KQM), exposed within the Central Maine Terrane (CMT) of New England. Both are Acadian-aged (≈413 Ma) synmetamorphic and syntectonic members of the New Hampshire Plutonic Series (NHPS). Potential source rocks analyzed for this study include Silurian and Devonian metasedimentary rocks of the CMT, and Ordovician metasedimentary rocks and granitic gneisses of the Bronson Hill Anticlinorium (BHA), which border the CMT to the west. The ɛSr(413), ɛNd(413) and δ18O values for the KQM range from 56.3 to 120.0, 2.8 to -6.4, and 7.6‰ to 12.9‰, respectively; values for the BG range from 7.4 to 144.7, 0.6 to -9.3, and 8.3‰ to 11.3‰, respectively; and values for possible source rocks range from 38.1 to 654.2, -10.7 to 5.4, and 6.2‰ to 14.1‰, respectively. Both the BG and KQM have extremely heterogeneous initial isotopic compositions consistent with mixing of multiple crustal source rocks, and neither contains a volumetrically significant (i.e., ≥10%) mantlederived component. Overlapping values of ɛNd(413), ɛSr(413) and δ18O values for both the BG and KQM samples resemble values for metasedimentary host rocks of the CMT and BHA. We observe no systematic correlations between ɛNd and ɛSr values for either the BG or the KQM. The ɛSr and δ18O values for the BG do not form any simple mixing trends, nor is there any direct correlation between the isotopic compositions of contact BG samples and their adjacent host rocks, in contrast to our observations for the KQM (Lathrop et al. 1994). We propose that the KQM and BG magmas were generated through anatexis of metasedimentary rocks from both the BHA and CMT in response to crystal thickening during the Acadian orogeny. Melting may have been initiated within CMT metasediments in response to high heat production in these mid-crustal rocks combined with

  20. A combined geochronological approach to investigating long lived granite magmatism, the Shap granite, UK

    NASA Astrophysics Data System (ADS)

    Miles, A. J.; Woodcock, N. H.

    2018-04-01

    With the advent of more precise dating methods, it has become apparent that zircon dates from granite plutons frequently indicate older emplacement ages than other dating methods. Here we attempt to reconcile a number of dating methods from the c. 5 km2 Caledonian Shap granite, Northern England. The results reveal a more complex and protracted evolution than indicated by application of any single dating method. Zircon U-Pb dates give a weighted mean age of 415.6 ± 1.4 (2σ) Ma. A mafic enclave, dated at 412 ± 2 (2σ) Ma (revised Rb-Sr feldspar age from Davidson et al., 2005), contains resorbed K-feldspar and zircon crystals scavenged from the host crystal mush. These ages are at odds with field relations in the thermal aureole that suggest final emplacement at approximately 404 Ma or later during Acadian deformation. Previously reported Re-Os ages on molybdenites associated with magmatic fluids, have given ages of 405.2 ± 1.8 (2σ) Ma (Selby et al., 2008) and confirm the overlap of at least some magmatic activity with Acadian deformation. A similar emplacement age is supported by Rb-Sr whole-rock-mineral and biotite K-Ar dates when adjusted for revised decay constants (402 ± 3 Ma and 401 ± 7 Ma, respectively, Wadge et al., 1978). The lower closure temperatures of these systems relative to the U-Pb system in zircon means that they are more likely to record the timing of final granite emplacement. These data suggest that most zircons grew before final granite emplacement, by about 10 Ma on average. We suggest that the majority of zircon crystals record pre-emplacement magmatic activity within a deeper part of the system. Mafic enclaves and their scavenged cargo of crystals record the assembly of a mid-crustal batholith where crystals remained at least locally mobile at 412 Ma. Gravity data support the existence of an extensive, 1500 km2 intrusive body, originally at about 15 km depth beneath Shap. This batholith is likely to have remained below the granite

  1. Estimating tree crown widths for the primary Acadian species in Maine

    Treesearch

    Matthew B. Russell; Aaron R. Weiskittel

    2012-01-01

    In this analysis, data for seven conifer and eight hardwood species were gathered from across the state of Maine for estimating tree crown widths. Maximum and largest crown width equations were developed using tree diameter at breast height as the primary predicting variable. Quantile regression techniques were used to estimate the maximum crown width and a constrained...

  2. Closure of George Air Force Base, San Bernardino County, California

    DTIC Science & Technology

    1990-05-01

    and welfare , crops, livestock, and property. " Resource Conservation and Recovery Act (RCRA). Regulates the disposal of haza,’dous wastes. 1-4...Willow flycatcher X (ErnroidQon tail) Yellow breasted chat X (Icteria virens) Summer tanager X (EPcrag rubra) Bendire’s thrasher X (Toxostoma bendirei...Veteran’s Administration. An Ld. of 55 dB is recognized by HUD, DOT, and EPA as an outdoor goal for protecting public welfare in residential areas. This

  3. Igneous petrogenesis and tectonic setting of granitic rocks from the eastern Blue Ridge, Alabama Appalachians

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Drummond, M.S.; Allison, D.T.; Tull, J.F.

    1994-03-01

    A span of 150 my of orogenic activity is recorded within the granitic rocks of the eastern Blue Ridge of Alabama (EBR). Four discrete episodes of plutonism can be differentiated, each event exhibiting distinct field relations and geochemical signatures. (1) Penobscotian stage: this initial stage of plutonic activity is represented by the Elkahatchee Quartz Diorite (EQD), a premetamorphic (495 Ma) batholith and the largest intrusive complex (880 km[sup 2]) exposed in the Blue Ridge. Calc-alkaline I-type tonalite-granodiorite are the principal lithologies, with subordinate cumulate hbl-bt diorite, metadacite, granite and trondhjemite. The parental tonalitic magmas are interpreted to have been derivedmore » from a subducted MORB source under eclogite to get amphibolite conditions. (2) Taconic stage: the Kowaliga augen gneiss (KAG) and the Zana granite gneiss (ZG) are 460 Ma granitic bodies that reside in the SE extremity and structurally highest portion of the EBR. Both of these bodies are pre-metamorphic with strongly elongate sill- and pod-like shapes concordant with S[sub 1] foliation. Granite and granodiorite comprise the bulk of the KAG. (3) Acadian stage: Rockford Granite (RG), Bluff springs Granite (BSG, 366 Ma), and Almond Trondhjemite represent a suite of pre- to syn-metamorphic granitic intrusions. (4) late-Acadian stage: The Blakes Ferry pluton (BFP) is a post-kinematic pluton displaying spectacular by schlieren igneous flow structures, but no metamorphic fabric. The pluton's age can be bracketed between a 366 Ma age on the BSG and a 324 Ma K-Ar muscovite age on the BFP. BFP's petrogenesis has involved partial melting a MORB source followed by assimilation of metasedimentary host rock.« less

  4. A Late Silurian U-Pb zircon age for Linville metadiabase, Grandfather Mountain window, North Carolina

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Fetter, A.H.; Goldberg, S.A.

    1993-03-01

    Linville metadiabase intrudes both Precambrian basement within the Grandfather Mountain window and nonconformably overlying Grandfather Mountain Formation. It occurs as sill-like and dike-like bodies, and is apparently not recognized outside of the window. Major element data classify the composition of the rock as tholeiitic basalt. Zircon was separated from a metadiabase body intruding metasiltstone of the Grandfather Mountain Formation west of Cranberry Knob. Zircon are clear, euhedral, with a l/w ratio of 3:1. Two abraded fractions (75-150 and < 75 [mu]m, both NM-2[degree]) yield concordant ages, which the authors report as 415 [plus minus] 3 Ma (2[sigma]), a weighted meanmore » of the two [sup 207]Pb/[sup 206]Pb ages. This Late Silurian (Ludlow) age is interpreted as the time of crystallization. Previously, similarities in mode of occurrence and major element composition have been used to correlate Linville metadiabase with late Precambrian mafic rocks within and outside of the window, as field relations do not constrain its age. Linville metadiabase thus is an unlikely candidate for feeders to the Montezuma metabasalt, which occurs as a flow immediately above metarhyolite dated as 742 [plus minus] 2 Ma (2[sigma]). Linville metadiabase may be one component of a magmatic pulse spanning 10-20 m.y. associated with the Acadian orogeny. The new zircon age places constraints on the timing of metamorphism and deformation, as Linville metadiabase is foliated, containing metamorphic assemblages from the biotite zone of the greenschist facies. The age and fabric relations are permissive evidence of post-Taconic, Acadian or Alleghanian orogeny.« less

  5. Effects of nocturnal illumination on life-history decisions and fitness in two wild songbird species.

    PubMed

    de Jong, Maaike; Ouyang, Jenny Q; Da Silva, Arnaud; van Grunsven, Roy H A; Kempenaers, Bart; Visser, Marcel E; Spoelstra, Kamiel

    2015-05-05

    The effects of artificial night lighting on animal behaviour and fitness are largely unknown. Most studies report short-term consequences in locations that are also exposed to other anthropogenic disturbance. We know little about how the effects of nocturnal illumination vary with different light colour compositions. This is increasingly relevant as the use of LED lights becomes more common, and LED light colour composition can be easily adjusted. We experimentally illuminated previously dark natural habitat with white, green and red light, and measured the effects on life-history decisions and fitness in two free-living songbird species, the great tit (Parus major) and pied flycatcher (Ficedula hypoleuca) in two consecutive years. In 2013, but not in 2014, we found an effect of light treatment on lay date, and of the interaction of treatment and distance to the nearest lamp post on chick mass in great tits but not in pied flycatchers. We did not find an effect in either species of light treatment on breeding densities, clutch size, probability of brood failure, number of fledglings and adult survival. The finding that light colour may have differential effects opens up the possibility to mitigate negative ecological effects of nocturnal illumination by using different light spectra. © 2015 The Author(s) Published by the Royal Society. All rights reserved.

  6. Bimodal Silurian and Lower Devonian volcanic rock assemblages in the Machias-Eastport area, Maine

    USGS Publications Warehouse

    Gates, Olcott; Moench, R.H.

    1981-01-01

    Exposed in the Machias-Eastport area of southeastern Maine is the thickest (at least 8,000 m), best exposed, best dated, and most nearly complete succession of Silurian and Lower Devonian volcanic strata in the coastal volcanic belt, remnants of which crop out along the coasts of southern New Brunswick, Canada, and southeastern New England in the United States. The volcanics were erupted through the 600-700-million-year-old Avalonian sialic basement. To test the possibility that this volcanic belt was a magmatic arc above a subduction zone prior to presumed Acadian continental collision, samples representing the entire section in the Machias-Eastport area of Maine were chemically analyzed. Three strongly bimodal assemblages of volcanic rocks and associated intrusives are recognized, herein called the Silurian, older Devonian, and younger Devonian assemblages. The Silurian assemblage contains typically nonporphyritic high-alumina tholeiitic basalts, basaltic andesites, and diabase of continental characterand calc-alkalic rhyolites, silicic dacites, and one known dike of andesite. These rocks are associated with fossiliferous, predominantly marine strata of the Quoddy, Dennys, and Edmunds Formations, and the Leighton Formation of the Pembroke Group (the stratigraphic rank of both is revised herein for the Machias-Eastport area), all of Silurian age. The shallow marine Hersey Formation (stratigraphic rank also revised herein) of the Pembroke Group, of latest Silurian age (and possibly earliest Devonian, as suggested by an ostracode fauna), contains no known volcanics; and it evidently was deposited during a volcanic hiatus that immediately preceded emergence of the coastal volcanic belt and the eruption of the older Devonian assemblage. The older Devonian assemblage, in the lagoonal to subaerial Lower Devonian Eastport Formation, contains tholeiitic basalts and basaltic andesites, typically with abundant plagioclase phenocrysts and typically richer in iron and

  7. Height development of shade-tolerant conifer saplings in multiaged Acadian forest stands

    Treesearch

    Andrew R. Moores; Robert S. Seymour; Laura S. Kenefic

    2007-01-01

    Understory growth dynamics of northern conifer species were studied in four stands managed under multiaged silvicultural systems in eastern Maine. Height growth of Picea rubens Sarg., Abies balsamea (L.) Mill., and Tsuga canadensis (L.) Carr. saplings between 0.5 and 6.0 m in height was related to the proportion...

  8. Mitigation of Shore Damage Attributed to the Federal Navigation Structures at Hammond Bay Harbor.

    DTIC Science & Technology

    1976-11-01

    34.’,u:e bz ma :’s Winter wren :,. UsatN2II Golden crown kinglet , f, cI’OPuM Cedar waxwing ’ ’’w .": x?,r’ ,7jq Northern shrike * , ~ PIPar Starling...flycatcher Pct-,hlidon Bank swallows Progue sabis Purple martins Troo Zodytes aedon House wren Dv,,’tolZa oaroliensis Catbird T ,xosterna rufern Brown...and Related Land Resources b the Wic ’ , sir ,S ,ounci 1. S. ai 9.t Mr. and Mi_rs. Joseph__ Mser I. Comment: We have read the environin.ntaIl impact

  9. Remote-Sensing Survey of the Atchafalaya Basin Main Channel, Atchafalaya Channel Training Project, Sts. Martin and Mary Parishes, Louisiana

    DTIC Science & Technology

    1991-11-01

    Just above Cornay’s Bridge they sunk the steamer Flycatcher and a schooner loaded with bricks, plus live oak trees were cut down and thrown into the...contour level) (Feet) Single Objects Engine camshaft 20 fi x 2 m 45 45 x 50 feet 15 Cas’ Iron soil pipe 10 ft long. 100 lbs 1407 45 x 65 feet 4 Iron...hitting any of the numerous fallen trees , snags, submerged logs, shallow sand bars, etc., 52 Chapter 3. Remote-Sensing Survey which occur along much of the

  10. Paleomagnetic Progress in Peri-Gondwanan Terranes of Cape Breton Island, Nova Scotia

    NASA Astrophysics Data System (ADS)

    Grunow, A. M.; Thompson, M. D.; Barr, S. M.; White, C. E.

    2009-05-01

    Paleopoles from primary Ediacaran magnetization directions established the Gondwanan origin of northern Appalachian Avalonian terranes, but magnetic overprints in the same rocks also provide useful tectonic information. Thus, in the Southeastern New England Avalon Zone, virtual geomagnetic poles (VGPs) calculated from magnetic B and C components in both 595 Ma Lynn-Mattapan volcanic rocks and 490-488 Ma Nahant Gabbro track mid- and late Paleozoic segments of the North American apparent polar wander path (APWP), suggesting the influence of Acadian and Neo-Acadian accretionary events. We report here on multi- vectorial magnetizations in pilot samples from Cape Breton Island, Nova Scotia where the Bras d'Or and Mira terranes represent both Ganderian and Avalonian elements transferred from Gondwana. Overprint relationships in these terranes may constrain their amalgamation with each other as well their docking with Laurentia. As in southeastern New England, secondary remanences can be identified in Cape Breton Island as consistent magnetization directions in rocks of differing ages. The S- to SSE-trending and gently downward pointing direction reported in 1985 by Johnson and Van der Voo in Middle Cambrian sedimentary rocks of the Bourinot Group (Bras d'Or terrane), for example, is also present in the 563 Ma Main à Dieu Formation and in 620 Ma Chisholm Brook Granite and East Bay Hill rhyolite (Mira terrane). This magnetization represents the C component already found around Boston, MA. The resulting VGPs in both areas occupy positions on the North American APWP consistent with a Neo-Acadian overprint, possibly related to the docking of the Meguma terrane against previously accreted Avalonia. Other overprint directions encountered in this investigation give rise to VGPs that do not coincide with the North American APWP, hence appear to reflect tectonic events independent of Laurentia. One such cluster comprising both Mira and Bras d'Or VGPs includes the paleopole also

  11. French-Canadians, Acadians and the French in New England. A Learning Activity Packet.

    ERIC Educational Resources Information Center

    Maine Univ., Orono. New England - Atlantic Provinces - Quebec Center.

    The history of the French-Canadians from their beginnings to the present time is the topic of this Learning Activity Packet (LAP). Designed to acquaint students with the French, Canada's earliest permanent settlers following the Indians, the unit is divided into six objectives which include learning activities for each. Students are expected to…

  12. Science-Driven Management of Protected Areas: A Philippine Case Study

    NASA Astrophysics Data System (ADS)

    Mallari, Neil Aldrin D.; Collar, Nigel J.; McGowan, Philip J. K.; Marsden, Stuart J.

    2013-06-01

    The lack of scientific baseline information hinders appropriate design and management of protected areas. To illustrate the value of science to management, we consider five scenarios for the 202.0 km² Puerto Princesa Subterranean River National Park, Philippines: (1) closure to human activities, (2) and (3) two levels of increase in unplanned human activities, (4) creation of a forest corridor and (5) additional allocation of land for permanent or shifting agriculture. We then use habitat-specific bird density estimates to simulate the net effect of each scenario on 18 focal bird populations. Closure has significant benefits—populations of five species are predicted to increase by >50 % and nine by >25 %, but two secondary forest flycatchers, including the endemic and `Vulnerable' Palawan flycatcher, decline dramatically, while the creation of a 4.0 km² forest corridor yields average increases across species of 2 ± 4 % (SD). In contrast, heavier unplanned park usage produces declines in all but a few species, while the negative effects of an extra 2.0 km² of shifting cultivation are 3-5 times higher than for a similar area of permanent agriculture and affect species whose densities are highest in primary habitats. Relatively small changes within the park, especially those associated with agricultural expansion, has serious predicted implications for local bird populations. Our models do not take into account the full complexities of bird ecology at a site, but they do provide park managers with an evidence base from which to make better decisions relating to biodiversity conservation obligations which their parks are intended to meet.

  13. Science-driven management of protected areas: a Philippine case study.

    PubMed

    Mallari, Neil Aldrin D; Collar, Nigel J; McGowan, Philip J K; Marsden, Stuart J

    2013-06-01

    The lack of scientific baseline information hinders appropriate design and management of protected areas. To illustrate the value of science to management, we consider five scenarios for the 202.0 km² Puerto Princesa Subterranean River National Park, Philippines: (1) closure to human activities, (2) and (3) two levels of increase in unplanned human activities, (4) creation of a forest corridor and (5) additional allocation of land for permanent or shifting agriculture. We then use habitat-specific bird density estimates to simulate the net effect of each scenario on 18 focal bird populations. Closure has significant benefits-populations of five species are predicted to increase by >50 % and nine by >25 %, but two secondary forest flycatchers, including the endemic and 'Vulnerable' Palawan flycatcher, decline dramatically, while the creation of a 4.0 km² forest corridor yields average increases across species of 2 ± 4 % (SD). In contrast, heavier unplanned park usage produces declines in all but a few species, while the negative effects of an extra 2.0 km² of shifting cultivation are 3-5 times higher than for a similar area of permanent agriculture and affect species whose densities are highest in primary habitats. Relatively small changes within the park, especially those associated with agricultural expansion, has serious predicted implications for local bird populations. Our models do not take into account the full complexities of bird ecology at a site, but they do provide park managers with an evidence base from which to make better decisions relating to biodiversity conservation obligations which their parks are intended to meet.

  14. Fitness consequences of northward dispersal as possible adaptation to climate change, using experimental translocation of a migratory passerine.

    PubMed

    Burger, Claudia; Nord, Andreas; Nilsson, Jan-Åke; Gilot-Fromont, Emmanuelle; Both, Christiaan

    2013-01-01

    Climate change leads to rapid, differential changes in phenology across trophic levels, often resulting in temporal mismatches between predators and their prey. If a species cannot easily adjust its timing, it can adapt by choosing a new breeding location with a later phenology of its prey. In this study, we experimentally investigated whether long-distance dispersal to northern breeding grounds with a later phenology could be a feasible process to restore the match between timing of breeding and peak food abundance and thus improve reproductive success. Here, we report the successful translocation of pied flycatchers (Ficedula hypoleuca) to natural breeding sites 560 km to the Northeast. We expected translocated birds to have a fitness advantage with respect to environmental phenology, but to potentially pay costs through the lack of other locally adapted traits. Translocated individuals started egg laying 11 days earlier than northern control birds, which were translocated only within the northern site. The number of fledglings produced was somewhat lower in translocated birds, compared to northern controls, and fledglings were in lower body condition. Translocated individuals were performing not significantly different to control birds that remained at the original southern site. The lack of advantage of the translocated individuals most likely resulted from the exceptionally cold spring in which the experiment was carried out. Our results, however, suggest that pied flycatchers can successfully introduce their early breeding phenotype after dispersing to more northern areas, and thus that adaptation through dispersal is a viable option for populations that get locally maladapted through climate change.

  15. Guild structure of a riparian avifauna relative to seasonal cattle grazing

    USGS Publications Warehouse

    Knopf, F.L.; Sedgwick, J.A.; Cannon, R. W.

    1988-01-01

    Knopf et al. found that summer cattle grazing has an adverse effect on the presence of certain willow-dependent songbirds. Pastures that have historical summer grazing no longer have the Willow flycatcher, Lincoln's sparrow and the White-crowned sparrow present. Yet in these same areas, birds like the American Robin, Brown-headed cowbird and the Red-winged blackbird have increased in density. One possible answer for the decrease in some songbirds is the fact that the main focus of the Arapaho National Wildlife Refuge is on waterfowl habitat, which requires large amounts of open space (opposite of desirable songbird habitat).

  16. Silurian extension in the Upper Connecticut Valley, United States and the origin of middle Paleozoic basins in the Québec embayment

    USGS Publications Warehouse

    Rankin, D.W.; Coish, R.A.; Tucker, R.D.; Peng, Z.X.; Wilson, S.A.; Rouff, A.A.

    2007-01-01

    Pre-Silurian strata of the Bronson Hill arch (BHA) in the Upper Connecticut Valley, NH-VT are host to the latest Ludlow Comerford Intrusive Suite consisting, east to west, of a mafic dike swarm with sheeted dikes, and an intrusive complex. The rocks are mostly mafic but with compositions ranging from gabbro to leucocratic tonalite. The suite is truncated on the west by the Monroe fault, a late Acadian thrust that carries rocks of the BHA westward over Silurian-Devonian strata of the Connecticut Valley-Gaspe?? trough (CVGT). Dikes intrude folded strata with a pre-intrusion metamorphic fabric (Taconian?) but they experienced Acadian deformation. Twenty fractions of zircon and baddeleyite from three sample sites of gabbrodiorite spanning nearly 40 km yield a weighted 207Pb/206Pb age of 419 ?? 1 Ma. Greenschist-facies dikes, sampled over a strike distance of 35 km, were tholeiitic basalts formed by partial melting of asthenospheric mantle, with little or no influence from mantle or crustal lithosphere. The dike chemistry is similar to mid-ocean ridge, within-plate, and back-arc basin basalts. Parent magmas originated in the asthenosphere and were erupted through severely thinned lithosphere adjacent to the CVGT. Extensive middle Paleozoic basins in the internides of the Appalachian orogen are restricted to the Que??bec embayment of the Laurentian rifted margin, and include the CVGT and the Central Maine trough (CMT), separated from the BHA by a Silurian tectonic hinge. The NE-trending Comerford intrusions parallel the CVGT, CMT, and the tectonic hinge, and indicate NW-SE extension. During post-Taconian convergence, the irregular margins of composite Laurentia and Avalon permitted continued collision in Newfoundland (St. Lawrence promontory) and coeval extension in the Que??bec embayment. Extension may be related to hinge retreat of the northwest directed Brunswick subduction complex and rise of the asthenosphere following slab break-off. An alternative hypothesis is

  17. Contrasting tectonothermal domains and faulting in the Potomac terrane, Virginia-Maryland - Discrimination by 40Ar/39Ar and fission-track thermochronology

    USGS Publications Warehouse

    Kunk, Michael J.; Wintsch, R.P.; Naeser, C.W.; Naeser, N.D.; Southworth, C.S.; Drake, Avery A.; Becker, J.L.

    2005-01-01

    New 40Ar/39Ar data reveal ages and thermal discontinuities that identify mapped and unmapped fault boundaries in the Potomac terrane in northern Virginia, thus confirming previous interpretations that it is a composite terrane. The rocks of the Potomac terrane were examined along the Potomac River, where it has been previously subdivided into three units: the Mather Gorge, Sykesville, and Laurel Formations. In the Mather Gorge Formation, at least two metamorphic thermal domains were identified, the Blockhouse Point and Bear Island domains, separated by a fault active in the late Devonian. Early Ordovician (ca. 475 Ma) cooling ages of amphibole in the Bear Island domain reflect cooling from Taconic metamorphism, whereas the Blockhouse Point domain was first metamorphosed in the Devonian. The 40Ar/39Ar data from muscovites in a third (eastern) domain within the Mather Gorge Formation, the Stubblefield Falls domain, record thrusting of the Sykesville Formation over the Mather Gorge Formation on the Plummers Island fault in the Devonian. The existence of two distinctly different thermal domains separated by a tectonic boundary within the Mather Gorge argues against its status as a formation. Hornblende cooling ages in the Sykesville Formation are Early Devonian (ca. 400 Ma), reflecting cooling from Taconic and Acadian metamorphism. The ages of retrograde and overprinting muscovite in phyllonites from domain-bounding faults are late Devonian (Acadian) and late Pennsylvanian (Alleghanian), marking the time of assembly of these domains and subsequent movement on the Plummers Island fault. Our data indicate that net vertical motion between the Bear Island domain of the Mather Gorge complex and the Sykesville Formation across the Plummers Island fault is east-side-up. Zircon fission-track cooling ages demonstrate thermal equillbrium across the Potomac terrane in the early Permian, and apatite fission-track cooling ages record tilting of the Potomac terrane in the Cretaceous

  18. Paleocurrent analysis of a deformed Devonian foreland basin in the northern Appalachians, Maine, USA

    USGS Publications Warehouse

    Bradley, D.C.; Hanson, L.S.

    2002-01-01

    New paleocurrent data indicate that the widespread Late Silurian and Devonian flysch and molasse succession in Maine was deposited in an ancestral, migrating foreland basin adjacent to an advancing Acadian orogenic belt. The foreland-basin sequence spread across a varied Silurian paleogeography of deep basins and small islands-the vestiges of an intraoceanic arc complex that not long before had collided with the Laurentian passive margin during the Ordovician Taconic Orogeny. We report paleocurrents from 43 sites representing 12 stratigraphic units, the most robust and consistent results coming from three units: Madrid Formation (southwesterly paleoflow), Carrabassett Formation (northerly paleoflow), and Seboomook Group (westerly paleoflow). Deformation and regional metamorphism are sufficiently intense to test the limits of paleocurrent analysis requiring particular care in retrodeformation. ?? 2002 Elsevier Science B.V. All rights reserved.

  19. Comparison of historical bottleneck effects and genetic consequences of re-introduction in a critically endangered island passerine.

    PubMed

    Bristol, Rachel M; Tucker, Rachel; Dawson, Deborah A; Horsburgh, Gavin; Prys-Jones, Robert P; Frantz, Alain C; Krupa, Andy; Shah, Nirmal J; Burke, Terry; Groombridge, Jim J

    2013-09-01

    Re-introduction is an important tool for recovering endangered species; however, the magnitude of genetic consequences for re-introduced populations remains largely unknown, in particular the relative impacts of historical population bottlenecks compared to those induced by conservation management. We characterize 14 microsatellite loci developed for the Seychelles paradise flycatcher and use them to quantify temporal and spatial measures of genetic variation across a 134-year time frame encompassing a historical bottleneck that reduced the species to ~28 individuals in the 1960s, through the initial stages of recovery and across a second contemporary conservation-introduction-induced bottleneck. We then evaluate the relative impacts of the two bottlenecks, and finally apply our findings to inform broader re-introduction strategy. We find a temporal trend of significant decrease in standard measures of genetic diversity across the historical bottleneck, but only a nonsignificant downward trend in number of alleles across the contemporary bottleneck. However, accounting for the different timescales of the two bottlenecks (~40 historical generations versus <1 contemporary generation), the loss of genetic diversity per generation is greater across the contemporary bottleneck. Historically, the flycatcher population was genetically structured; however, extinction on four of five islands has resulted in a homogeneous contemporary population. We conclude that severe historical bottlenecks can leave a large footprint in terms of sheer quantity of genetic diversity lost. However, severely depleted genetic diversity does not render a species immune to further genetic erosion upon re-introduction. In some cases, the loss of genetic diversity per generation can, initially at least, be greater across re-introduction-induced bottlenecks. © 2013 John Wiley & Sons Ltd.

  20. First year results from an experimental study of the ecohydrologic benefits of beaver and beaver dam analogue restoration techniques in Childs Meadow, CA

    NASA Astrophysics Data System (ADS)

    Yarnell, S. M.; Pope, K.; Podolak, K.; Wolf, E.; Burnett, R.

    2016-12-01

    Due to extensive livestock grazing and widespread removal of beaver and willows, headwater meadows have transformed from multi-thread channels with seasonally active floodplains into single thread, incised channels that store less carbon, retain less water, and are lower in habitat quality for a diverse suite of meadow-dependent wildlife. Meadow restoration techniques often include willow planting and cattle exclosures; however, few studies have rigorously tested the long-term efficacy of these methods or evaluated alternative restoration techniques such as reintroduction of beaver or installation of beaver dam analogues (BDAs). This project seeks to evaluate the installation of BDAs as a restoration technique in Childs Meadow, a heavily grazed meadow in the Cascade Range representative of low-gradient meadows across northern California. Using a before-after-control-impact study design, the study tests the impacts of two restoration techniques (willow planting with cattle exclusion and willow planting with cattle exclusion and BDAs) on hydrology, carbon sequestration, and sensitive species. Results will be compared with measurements in an unrestored section of the meadow that currently supports an active beaver population and two imperiled species (Cascades Frog and Willow Flycatcher). One specific project objective is to measure the response of hydrogeomorphic conditions (e.g. groundwater, surface water, temperature, habitat) and Cascades Frog and Willow Flycatcher to restorative actions. Pre-treatment data was collected in summer 2015, a cattle exclosure was established and willows were planted in fall 2015, and installation of the BDAs is planned for fall 2016. Three years of post-implementation monitoring will be completed to assess impacts of the treatments. Here, we will present our sampling design and first year results following initiation of the treatments.

  1. A neotropical migrant bird's dilemma: where to stop for a good meal

    USGS Publications Warehouse

    Fontaine, Joseph J.; van Riper, Charles

    2009-01-01

    To learn how migrating birds determine where to stop and find food, scientists from the U.S. Geological Survey, Northern Arizona University, and The University of Arizona studied the behavior of 28 species of neotropical migrant songbirds - warblers, flycatchers, tanagers, and vireos - along the lower Colorado River from 2001 to 2004. They found that, like interstate travelers greeted by restaurant billboards, songbirds flying over Cibola National Wildlife Refuge, Arizona, relied on the flowering of honey mesquite (Prosopis glandulosa) to detect the availability of insects that they prey on. Understanding where and why migrant birds stop will help land managers better protect key habitats used by these tiny travelers.

  2. Streamflow and Endangered Species Habitat in the Lower Isleta Reach of the Middle Rio Grande

    USGS Publications Warehouse

    Bovee, Ken D.; Waddle, Terry J.; Spears, J. Mark

    2008-01-01

    San Acacia Dam is located in a reach of the Rio Grande that has been designated as critical habitat for two endangered species, the Rio Grande silvery minnow (Hybognathus amarus) and the southwestern willow flycatcher (Empidonax traillii extimus). Under present operations, the Rio Grande upstream from the dam is used to convey irrigation water to the Socorro main canal at San Acacia Dam. In order to increase operational flexibility and improve irrigation delivery efficiency, the 'Bernardo Siphon' has been proposed to intercept up to 150 cubic feet per second from the Lower San Juan Riverside Drain on the east side of the Rio Grande and transport it under the river into a drainage canal on the west side. Irrigation deliveries to the Socorro main canal would be conveyed by way of the drainage canal rather than the Rio Grande. The objective of this study was to provide the Bureau of Reclamation (BOR) and other stakeholders with a tool to evaluate the effects of different operational modes of the Bernardo siphon on habitat for H. amarus and E. t. extimus in this section of river. We used a two-dimensional hydraulic simulation model to simulate hydraulic conditions for a range of discharges at three study sites in the Rio Grande between the proposed siphon location and San Acacia Dam. Suitable habitat characteristics were defined for H. amarus by consensus of a panel of experts and for E. t. extimus on the basis of a study conducted in 2003 by BOR. Habitat suitability maps for each targeted life stage and simulated discharge were constructed using a Geographic Information System (ArcGIS) and the results compiled into tables relating discharge to areas of suitable habitat. A separate analysis was conducted to calculate an index of connectivity among habitat patches at low flows. A hydrologic model was constructed to synthesize flows, by reach, without the siphon, which was used as a baseline for comparison with similarly-synthesized discharges with the siphon under

  3. Political Partisanship, Bureaucratic Pragmatism and Acadian Nationalism: New Brunswick, Canada's 1920 History Textbook Controversy

    ERIC Educational Resources Information Center

    Helyar, Frances

    2014-01-01

    During a time of post-war sensitivity to Canadian nationalism and patriotism, public feeling was aroused in 1920 New Brunswick regarding a world history textbook with a new chapter about the First World War. The American author made no reference to Canada's war efforts. The subsequent public discussion focused on issues of patriotism, citizenship,…

  4. Quality of Water from Shallow Wells in Urban Residential and Light Commercial Areas in Lafayette Parish, Louisiana, 2001 through 2002

    USGS Publications Warehouse

    Fendick, Robert B.; Tollett, Roland W.

    2004-01-01

    In 2001-02, the U.S. Geological Survey installed and sampled 28 shallow wells in urban residential and light commercial areas in Lafayette Parish, Louisiana, for a land-use study in the Acadian-Pontchartrain Study Unit of the National Water-Quality Assessment (NAWQA) Program. The wells were installed in the Chicot aquifer system, the primary source of water for irrigation and public-water supplies in southwestern Louisiana. The purpose of this report is to describe the quality of water from the 28 shallow wells and to relate that water quality to natural factors and to human activities. Ground-water samples were analyzed for general ground-water properties and about 240 water-quality contituents, including dissolved solids, major inorganic ions, trace elements, nutrients, dissolved organic carbon (DOC), radon, chlorofluorocarbons, selected stable isotopes, pesticides, pesticide degradation products, and volatile organic compounds (VOC's).

  5. Resolving Evolutionary Relationships in Closely Related Species with Whole-Genome Sequencing Data

    PubMed Central

    Nater, Alexander; Burri, Reto; Kawakami, Takeshi; Smeds, Linnéa; Ellegren, Hans

    2015-01-01

    Using genetic data to resolve the evolutionary relationships of species is of major interest in evolutionary and systematic biology. However, reconstructing the sequence of speciation events, the so-called species tree, in closely related and potentially hybridizing species is very challenging. Processes such as incomplete lineage sorting and interspecific gene flow result in local gene genealogies that differ in their topology from the species tree, and analyses of few loci with a single sequence per species are likely to produce conflicting or even misleading results. To study these phenomena on a full phylogenomic scale, we use whole-genome sequence data from 200 individuals of four black-and-white flycatcher species with so far unresolved phylogenetic relationships to infer gene tree topologies and visualize genome-wide patterns of gene tree incongruence. Using phylogenetic analysis in nonoverlapping 10-kb windows, we show that gene tree topologies are extremely diverse and change on a very small physical scale. Moreover, we find strong evidence for gene flow among flycatcher species, with distinct patterns of reduced introgression on the Z chromosome. To resolve species relationships on the background of widespread gene tree incongruence, we used four complementary coalescent-based methods for species tree reconstruction, including complex modeling approaches that incorporate post-divergence gene flow among species. This allowed us to infer the most likely species tree with high confidence. Based on this finding, we show that regions of reduced effective population size, which have been suggested as particularly useful for species tree inference, can produce positively misleading species tree topologies. Our findings disclose the pitfalls of using loci potentially under selection as phylogenetic markers and highlight the potential of modeling approaches to disentangle species relationships in systems with large effective population sizes and post

  6. Pleistocene land bridges act as semipermeable agents of avian gene flow in Wallacea.

    PubMed

    Garg, Kritika M; Chattopadhyay, Balaji; Wilton, Peter R; Malia Prawiradilaga, Dewi; Rheindt, Frank E

    2018-08-01

    Cyclical periods of global cooling have been important drivers of biotic differentiation throughout the Quaternary. Ice age-induced sea level fluctuations can lead to changing patterns of land connections, both facilitating and disrupting gene flow. In this study, we test if species with differing life histories are differentially affected by Quaternary land connections. We used genome-wide SNPs in combination with mitochondrial gene sequences to analyse levels of divergence and gene flow between two songbird complexes across two Wallacean islands that have been repeatedly connected during glaciations. Although the two bird complexes are similar in ecological attributes, the forest and edge-inhabiting golden whistler Pachycephala pectoralis is comparatively flexible in its diet and niche requirements as compared to the henna-tailed jungle-flycatcher Cyornis colonus, which is largely restricted to the forest interior. Using population-genomic and coalescent approaches, we estimated levels of gene flow, population differentiation and divergence time between the two island populations. We observed higher levels of differentiation, an approximately two to four times deeper divergence time and near-zero levels of gene flow between the two island populations of the more forest-dependent henna-tailed jungle-flycatcher as compared to the more generalist golden whistler. Our results suggest that Quaternary land bridges act as semipermeable agents of gene flow in Wallacea, allowing only certain taxa to connect between islands while others remain isolated. Quaternary land bridges do not accommodate all terrestrial species equally, differing in suitability according to life history and species biology. More generalist species are likely to use Quaternary land connections as a conduit for gene flow between islands whereas island populations of more specialist species may continue to be reproductively isolated even during periods of Quaternary land bridges. Copyright © 2018

  7. An Ecohydrological Approach to Riparian Restoration Planning in the American Southwest

    NASA Astrophysics Data System (ADS)

    Leverich, G. T.; Orr, B.; Diggory, Z.; Dudley, T.; Hatten, J.; Hultine, K. R.; Johnson, M. P.; Orr, D.

    2014-12-01

    Riparian systems across the American southwest region are under threat from a growing and intertwined cast of natural and anthropogenic stressors, including flooding, drought, invasion by non-native plants, wildfire, urban encroachment, and land- and water-use practices. In relatively remote and unregulated systems like the upper Gila River in Arizona, riparian habitat value has persisted reasonably well despite much of it being densely infested with non-native tamarisk (salt cedar). A new concern in the watershed, however, is the eventual arrival of the tamarisk leaf beetle that is expected to soon colonize the tamarisk-infested riparian corridor as the beetle continues to spread across the southwest region. While there are numerous potential benefits to tamarisk suppression (e.g., groundwater conservation, riparian habitat recovery, fire-risk reduction), short-term negative consequences are also possible, such as altered channel hydraulics and canopy defoliation during bird nesting season (e.g., the endangered southwestern willow flycatcher). In preparation for anticipated impacts following beetle colonization, we developed a holistic restoration framework to promote recovery of native riparian habitat and subsequent local increases in avian population. Pivotal to this process was an ecohydrological assessment that identified sustainable restoration sites based on consideration of natural and anthropogenic factors that, together, influence restoration opportunities—flood-scour dynamics, vegetation community structure and resilience, surface- and groundwater availability, soil texture and salinity, wildfire potential, and land-use activities. Data collected included high-resolution remote-sensing products, GIS-based delineation of geomorphic activity, and vegetation field mapping. These data along with other information generated, including pre-biocontrol vegetation monitoring and flycatcher-habitat modeling, were synthesized to produce a comprehensive

  8. Recommendations for cowbird management in recovery efforts for the southwestern willow flycatcher

    USGS Publications Warehouse

    Rothstein, S.I.; Kus, B.E.; Whitfield, M.J.; Sferra, S.J.

    2003-01-01

    Three generations of mallard ducks (Anas platyrhynchos) were fed either a control diet or a diet containing 0.5 ppm mercury in the form of methylmercury. The levels of mercury in adult tissues and eggs remained about the same over 3 generations. The methylmercury diet had no effect on adult weights or weight changes during the reproductive season. Females fed a diet containing 0.5 ppm mercury laid a greater percentage of their eggs outside their nestboxes than did controls, and also laid fewer eggs and produced fewer ducklings. Methylmercury in the diet appeared to result in a small amount of eggshell thinning. Ducklings from parents fed methylmercury were less responsive than, controls to tape-recorded maternal calls, but were hyper-responsive to a frightening stimulus in avoidance tests; there were no significant differences in locomotor activity in an open-field test.

  9. Mycobacteriosis due to Mycobacterium genavense in six pet birds.

    PubMed Central

    Hoop, R K; Böttger, E C; Ossent, P; Salfinger, M

    1993-01-01

    Six cases of mycobacteriosis due to Mycobacterium genavense in three budgerigars (Melopsittacus undulatus), one orange-winged amazon (Amazona amazonica), one flycatcher (Cyanoptila cyanomelana), and one zebra finch (Taeniopygia guttata) are discussed. Gross lesions associated with the infection included a high degree of muscular wasting (five cases), hepatomegaly (four cases), and thickening of the wall of the small intestine (four cases). Granulomas were found in the lung (one case) and the subcutis (one case). Acid-fast bacilli were detected in the liver of all six birds. Only the use of acidic BACTEC mediums consistently led to growth, whereas the egg-based medium failed. These findings point to a possible role of the environment as a reservoir for M. genavense. Images PMID:8463407

  10. A field trip guidebook to the type localities of Marland Billings' 1935 Paleozoic bedrock stratigraphy near Littleton, New Hampshire

    USGS Publications Warehouse

    Rankin, Douglas W.; Rankin, Mary B.

    2014-01-01

    Marland Billings' classic paper published in 1937 in the Geological Society of America Bulletin established a succession of six stratigraphic units in rocks of low metamorphic grade near Littleton, New Hampshire. The two youngest units are fossiliferous in the area, with ages established at the time as “middle” Silurian and Early Devonian. Billings and students mapped the same stratigraphic section in adjacent areas of progressively higher regional metamorphic grade. This work laid the foundation upon which a major part of subsequent work in New England has been directly or indirectly built. This guidebook was written for a field trip held in March 2013 to visit roadcuts that are as close as possible in March to the type localities or areas of Billings’ six-fold stratigraphic succession. Ten stops are in rocks of chlorite grade of Acadian(?) metamorphism; the final stop visits amphibolite of the Ammonoosuc Volcanics. Fieldwork by the authors over the past 20 years confirms Billings’ broad conclusions.

  11. The Usher syndrome in the Lebanese population and further refinement of the USH2A candidate region.

    PubMed

    Saouda, M; Mansour, A; Bou Moglabey, Y; El Zir, E; Mustapha, M; Chaib, H; Nehmé, A; Mégarbané, A; Loiselet, J; Petit, C; Slim, R

    1998-08-01

    Usher syndrome (USH) is an autosomal-recessive disease characterized by neurosensory deafness and progressive retinitis pigmentosa. So far, three clinical types of Usher syndrome have been defined, and are caused by defects at more than eight loci. We report the linkage analysis of seven Lebanese families with Usher syndrome, two with type I (USH1) and five with type II (USH2). We demonstrate that one family is linked to the USH1C locus, a rare form of USH1 only reported in the French Acadian population. Linkage analysis of the five USH2 families with recently mapped loci allowed us to reduce the USH2A candidate region to a very small interval flanked by D1S2646/D1S2629 and D1S2827. Furthermore, haplotype comparison between the different families suggests a founder effect for the USH2A mutation among the different Lebanese ethnic groups, while a genetic heterogeneity is noted for Usher syndrome type I.

  12. Alternative Speech Communication System for Persons with Severe Speech Disorders

    NASA Astrophysics Data System (ADS)

    Selouani, Sid-Ahmed; Sidi Yakoub, Mohammed; O'Shaughnessy, Douglas

    2009-12-01

    Assistive speech-enabled systems are proposed to help both French and English speaking persons with various speech disorders. The proposed assistive systems use automatic speech recognition (ASR) and speech synthesis in order to enhance the quality of communication. These systems aim at improving the intelligibility of pathologic speech making it as natural as possible and close to the original voice of the speaker. The resynthesized utterances use new basic units, a new concatenating algorithm and a grafting technique to correct the poorly pronounced phonemes. The ASR responses are uttered by the new speech synthesis system in order to convey an intelligible message to listeners. Experiments involving four American speakers with severe dysarthria and two Acadian French speakers with sound substitution disorders (SSDs) are carried out to demonstrate the efficiency of the proposed methods. An improvement of the Perceptual Evaluation of the Speech Quality (PESQ) value of 5% and more than 20% is achieved by the speech synthesis systems that deal with SSD and dysarthria, respectively.

  13. Change Points in the Population Trends of Aerial-Insectivorous Birds in North America: Synchronized in Time across Species and Regions.

    PubMed

    Smith, Adam C; Hudson, Marie-Anne R; Downes, Constance M; Francis, Charles M

    2015-01-01

    North American populations of aerial insectivorous birds are in steep decline. Aerial insectivores (AI) are a group of bird species that feed almost exclusively on insects in flight, and include swallows, swifts, nightjars, and flycatchers. The causes of the declines are not well understood. Indeed, it is not clear when the declines began, or whether the declines are shared across all species in the group (e.g., caused by changes in flying insect populations) or specific to each species (e.g., caused by changes in species' breeding habitat). A recent study suggested that population trends of aerial insectivores changed for the worse in the 1980s. If there was such a change point in trends of the group, understanding its timing and geographic pattern could help identify potential causes of the decline. We used a hierarchical Bayesian, penalized regression spline, change point model to estimate group-level change points in the trends of 22 species of AI, across 153 geographic strata of North America. We found evidence for group-level change points in 85% of the strata. Change points for flycatchers (FC) were distinct from those for swallows, swifts and nightjars (SSN) across North America, except in the Northeast, where all AI shared the same group-level change points. During the 1980s, there was a negative change point across most of North America, in the trends of SSN. For FC, the group-level change points were more geographically variable, and in many regions there were two: a positive change point followed by a negative change point. This group-level synchrony in AI population trends is likely evidence of a response to a common environmental factor(s) with similar effects on many species across broad spatial extents. The timing and geographic patterns of the change points that we identify here should provide a spring-board for research into the causes behind aerial insectivore declines.

  14. Change Points in the Population Trends of Aerial-Insectivorous Birds in North America: Synchronized in Time across Species and Regions

    PubMed Central

    Smith, Adam C.; Hudson, Marie-Anne R.; Downes, Constance M.; Francis, Charles M.

    2015-01-01

    North American populations of aerial insectivorous birds are in steep decline. Aerial insectivores (AI) are a group of bird species that feed almost exclusively on insects in flight, and include swallows, swifts, nightjars, and flycatchers. The causes of the declines are not well understood. Indeed, it is not clear when the declines began, or whether the declines are shared across all species in the group (e.g., caused by changes in flying insect populations) or specific to each species (e.g., caused by changes in species’ breeding habitat). A recent study suggested that population trends of aerial insectivores changed for the worse in the 1980s. If there was such a change point in trends of the group, understanding its timing and geographic pattern could help identify potential causes of the decline. We used a hierarchical Bayesian, penalized regression spline, change point model to estimate group-level change points in the trends of 22 species of AI, across 153 geographic strata of North America. We found evidence for group-level change points in 85% of the strata. Change points for flycatchers (FC) were distinct from those for swallows, swifts and nightjars (SSN) across North America, except in the Northeast, where all AI shared the same group-level change points. During the 1980s, there was a negative change point across most of North America, in the trends of SSN. For FC, the group-level change points were more geographically variable, and in many regions there were two: a positive change point followed by a negative change point. This group-level synchrony in AI population trends is likely evidence of a response to a common environmental factor(s) with similar effects on many species across broad spatial extents. The timing and geographic patterns of the change points that we identify here should provide a spring-board for research into the causes behind aerial insectivore declines. PMID:26147572

  15. Temporal and spatial distribution of Paleozoic metamorphism in the southern Appalachian Blue Ridge and Inner Piedmont delimited by ion microprobe U-Pb ages of metamorphic zircon

    USGS Publications Warehouse

    Merschat, Arthur J.; Bream, Brendan R.; Huebner, Matthew T.; Hatcher, Robert D.; Miller, Calvin F.

    2017-01-01

    Ion microprobe U-Pb zircon rim ages from 39 samples from across the accreted terranes of the central Blue Ridge, eastward across the Inner Piedmont, delimit the timing and spatial extent of superposed metamorphism in the southern Appalachian orogen. Metamorphic zircon rims are 10–40 µm wide, mostly unzoned, and dark gray to black or bright white in cathodoluminescence, and truncate and/or embay interior oscillatory zoning. Black unzoned and rounded or ovoid-shaped metamorphic zircon morphologies also occur. Th/U values range from 0.01 to 1.4, with the majority of ratios less than 0.1. Results of 206Pb/238U ages, ±2% discordant, range from 481 to 305 Ma. Clustering within these data reveals that the Blue Ridge and Inner Piedmont terranes were affected by three tectonothermal events: (1) 462–448 Ma (Taconic); (2) 395–340 Ma (Acadian and Neoacadian); and (3) 335–322 Ma, related to the early phase of the Alleghanian orogeny. By combining zircon rim ages with metamorphic isograds and other published isotopic ages, we identify the thermal architecture of the southern Appalachian orogen: juxtaposed and superposed metamorphic domains have younger ages to the east related to the marginward addition of terranes, and these domains can serve as a proxy to delimit terrane accretion. Most 462–448 Ma ages occur in the western and central Blue Ridge and define a continuous progression from greenschist to granulite facies that identifies the intact Taconic core. The extent of 462–448 Ma metamorphism indicates that the central Blue Ridge and Tugaloo terranes were accreted to the western Blue Ridge during the Taconic orogeny. Zircon rim ages in the Inner Piedmont span almost 100 m.y., with peaks at 395–385, 376–340, and 335–322 Ma, and delimit the Acadian-Neoacadian and Alleghanian metamorphic core. The timing and distribution of metamorphism in the Inner Piedmont are consistent with the Devonian to Mississippian oblique collision of the Carolina superterrane

  16. Bedrock geology of the Mount Carmel and Southington quadrangles, Connecticut

    USGS Publications Warehouse

    Fritts, Crawford Ellswroth

    1962-01-01

    thinly layered sediments that contained rather high percentages of fine-grained volcanic debris. Metadiabase and metabasalt extrusives above Derby Hill Schist south of the map area represent more intense volcanic activity before or during the early stages of the Taconic disturbance in Late Ordovician time. Impure argillaceous, siliceous, and minor calcareous sediments of the Wepawaug Schist, which is Silurian and Devonian in age, were deposited unconformably on older rocks during renewed subsidence of a geosyncline. The Wepawaug now occupies the trough of a tight syncline, which formed before and during progressive regional metamorphism at the time of the Acadian orogeny in middle to Late Devonian time. Felsic igneous rocks were intruded into the metasedimentary formations of Paleozoic age before the climax of the latest progressive regional metamorphism. Intrusives that gave rise to the Prospect and Ansonia Gneisses were emplaced mainly in the Southington Mountain Schist, and the igneous rocks as well as the host rocks were metamorphosed in the staurolite zone. Although it is possible that these two intrusives were emplaced during the Taconic disturbance, the writer believes it more likely that the igneous rocks from which the Prospect and Ansonia Gneisses formed were emplaced during the Acadian orogeny. Woodbridge Granite, which intruded the Wepawaug Schist, is Devonian in age and undoubtedly was emplaced during the Acadian orogeny. In this area the granite is essentially unmetamorphosed, because it is in the chlorite, garnet, and biotite zones. Southwest of the map area, however, metamorphic equivalents of the Woodbridge are found in Wepawaug Schist in the staurolite zone. The Ansonia Gneiss, therefore, may be a metamorphic equivalent of the Woodbridge Granite. Rocks of Late Triassic age formerly covered the entire map area, but were eroded from the western part after tilting and faulting in Late Triassic time. The New Haven Arkose of the Newark

  17. The Northeast Kingdom batholith, Vermont: magmatic evolution and geochemical constraints on the origin of Acadian granitic rocks

    USGS Publications Warehouse

    Ayuso, R.A.; Arth, Joseph G.

    1992-01-01

    Five Devonian plutons (West Charleston, Echo Pond, Nulhegan, Derby, and Willoughby) that constitute the Northeast Kingdom batholith in Vermont show wide ranges in elemental abundances and ratios consistent with major crustal contributions during their evolution. The batholith consists of metaluminous quartz gabbro, diorite and quartz monzodiorite, peraluminous granodiorite and granite, and strongly peraluminous leucogranite. Contents of major elements vary systematically with increasingSiO40) and have small negative Eu anomalies. The strongly peraluminous Willoughby leucogranite has unique trace-element abundances and ratios relative to the rest of the batholith, including low contents of Hf, Zr, Sr, and Ba, low values of K/Rb (80-164), Th/Ta (<9), Rb/Cs (7-40), K/Cs (0.1-0.5), Ce/Pb (0.5-4), high values of Rb/Sr (1-18) low to moderate REE contents and light-REE enriched patterns (with small negative Eu anomalies). Flat REE patterns (with large negative Eu anomalies) are found in a small, hydrothermally-altered area characterized by high abundances of Sn (up to 26 ppm), Rb (up to 670 ppm), Li (up to 310 ppm), Ta (up to 13.1 ppm), and U (up to 10 ppm). There is no single mixing trend, fractional crystallization assemblage, or assimilationscheme that accounts for all trace elementvariations from quartz gabbro to granite in the Northeast Kingdom batholith. The plutons originated by mixing mantle-derived components and crustal melts generated at different levels in the heterogeneous lithosphere in a continental collisional environment. Hybrid rocks in the batholith evolved by fractional crystallization and assimilation of country rocks (<50% by mass), and some of the leucogranitic rocks were subsequently disturbed by a mild hydrothermal event that resulted in the deposition of small amounts of sulfide minerals. ?? 1992 Springer-Verlag.

  18. Pre-lithification tectonic foliation development in a clastic sedimentary sequence

    NASA Astrophysics Data System (ADS)

    Meere, Patrick; Mulchrone, Kieran; McCarthy, David; Timmermann, Martin; Dewey, John

    2016-04-01

    The current view regarding the timing of regionally developed penetrative tectonic fabrics in sedimentary rocks is that their development postdates lithification of those rocks. In this case fabric development is achieved by a number of deformation mechanisms including grain rigid body rotation, crystal-plastic deformation and pressure solution (wet diffusion). The latter is believed to be the primary mechanism responsible for shortening and the domainal structure of cleavage development commonly observed in low grade metamorphic rocks. In this study we combine field observations with strain analysis and modelling to fully characterise considerable (>50%) mid-Devonian Acadian crustal shortening in a Devonian clastic sedimentary sequence from south west Ireland. Despite these high levels of shortening and associated penetrative tectonic fabric there is a marked absence of the expected domainal cleavage structure and intra-clast deformation, which are expected with this level of deformation. In contrast to the expected deformation processes associated with conventional cleavage development, fabrics in these rocks are a product of translation, rigid body rotation and repacking of extra-formational clasts during deformation of an un-lithified clastic sedimentary sequence.

  19. The Valley and Ridge Province of eastern Pennsylvania - stratigraphic and sedimentologic contributions and problems ( USA).

    USGS Publications Warehouse

    Epstein, J.B.

    1986-01-01

    The rocks in the area, which range from Middle Ordovician to Late Devonian in age, are more than 7620 m thick. This diversified group of sedimentary rocks was deposited in many different environments, ranging from deep sea, through neritic and tidal, to alluvial. In general, the Middle Ordovician through Lower Devonian strata are a sedimentary cycle related to the waxing and waning of Taconic tectonism. The sequence began with a greywacke-argillite suite (Martinsburg Formation) representing synorogenic basin deepening. This was followed by basin filling and progradation of a sandstone-shale clastic wedge (Shawangunk Formation and Bloomsburg Red Beds) derived from the erosion of the mountains that were uplifted during the Taconic orogeny. The sequence ended with deposition of many thin units of carbonate, sandstone, and shale on a shelf marginal to a land area of low relief. Another tectonic-sedimentary cycle, related to the Acadian orogeny, began with deposition of Middle Devonian rocks. Deep-water shales (Marcellus Shale) preceded shoaling (Mahantango Formation) and turbidite sedimentation (Trimmers Rock Formation) followed by another molasse (Catskill Formation). -from Author

  20. Nova Scotia: ``Feu Follet" At Cheticamp, and Also the Phenomena At L'Sitkuk of the Mi'Kmaw, May Be Electromagnetic In Nature.

    NASA Astrophysics Data System (ADS)

    Ochs, Michael Ann; McLeod, Roger D.

    2001-11-01

    There is a strong tradition that ``feu follet" exists at the cemetery associated with the Acadian French at Cheticamp. It is described as a blue light, and may actually be the equivalent of an ascending, positively charged stream of ions in the atmosphere, just like the blue-light column that is often a precursor of a storm's lightning-strike. Similar phenomena are at America's Stonehenge, at a stone serpent effigy site in Ohio, and just north of the Lakes Memphremagog and Magog of Vermont and Canada. At the Bear River L'sitkuk Reservation area, which seems to us to have been a most unsuitable site, was deliberately chosen by the Mi'kmaw for their living area. Was this because certain properties of the electromagnetic field (EMF) are evident to them there, which also seem to be reflected in their legends? We hope to establish that these disparate cultures and their separate worldviews can be confirmed by the presence of particular EMF signatures. *This paper does not represent the views of the United States Environmental Protection Agency.

  1. Predation as a landscape effect: the trading off by prey species between predation risks and protection benefits.

    PubMed

    Mönkkönen, M; Husby, M; Tornberg, R; Helle, P; Thomson, R L

    2007-05-01

    1. Predators impose costs on their prey but may also provide benefits such as protection against other (e.g. nest) predators. The optimal breeding location in relation to the distance from a nesting raptor varies so as to minimize the sum of costs of adult and nest predation. We provide a conceptual model to account for variation in the relative predation risks and derive qualitative predictions for how different prey species should respond to the distance from goshawk Accipiter gentilis nests. 2. We test the model predictions using a comprehensive collection of data from northern Finland and central Norway. First, we carried out a series of experiments with artificial bird nests to test if goshawks may provide protection against nest predation. Second, we conducted standard bird censuses and nest-box experiments to detect how the density or territory occupancy of several prey species varies with distance from the nearest goshawk nest. 3. Nest predation rate increased with distance from goshawk nest indicating that goshawks may provide protection for birds' nests against nest predation. Abundance (or probability of presence) of the main prey species of goshawks peaked at intermediate distances from goshawk nests, reflecting the trade-off. The abundance of small songbird species decreased with distance from goshawk nests. The goshawk poses little risk to small songbirds and they may benefit from goshawk proximity in protection against nest predation. Finally, no pattern with distance in pied flycatcher territory (nest box) occupation rate or the onset of egg-laying was detected. This is expected, as flycatchers neither suffer from marked nest predation risk nor are favoured goshawk prey. 4. Our results suggest that territory location in relation to the nest of a predator is a trade-off situation where adult birds weigh the risk of themselves being predated against the benefits accrued from increased nest survival. Prey species appear able to detect and measure

  2. Phanerozoic geological evolution of Northern and Central Africa: An overview

    NASA Astrophysics Data System (ADS)

    Guiraud, R.; Bosworth, W.; Thierry, J.; Delplanque, A.

    2005-10-01

    The principal paleogeographic characteristics of North and Central Africa during the Paleozoic were the permanency of large exposed lands over central Africa, surrounded by northerly and northwesterly dipping pediplanes episodically flooded by epicontinental seas related to the Paleotethys Ocean. The intra-continental Congo-Zaire Basin was also a long-lived feature, as well as the Somali Basin from Late Carboniferous times, in conjunction with the development of the Karoo basins of southern Africa. This configuration, in combination with eustatic sea-level fluctuations, had a strong influence on facies distributions. Significant transgressions occurred during the Early Cambrian, Tremadocian, Llandovery, Middle to Late Devonian, Early Carboniferous, and Moscovian. The Paleozoic tectonic history shows an alternation of long periods of predominantly gentle basin subsidence and short periods of gentle folding and occasionally basin inversion. Some local rift basins developed episodically, located mainly along the northern African-Arabian plate margin and near the West African Craton/Pan-African Belt suture. Several arches or spurs, mainly N-S to NE-SW trending and inherited from late Pan-African fault swarms, played an important role. The Nubia Province was the site of numerous alkaline anorogenic intrusions, starting in Ordovician times, and subsequently formed a large swell. Paleozoic compressional events occurred in the latest Early Cambrian ("Iskelian"), Medial Ordovician to earliest Silurian ("pre-Caradoc" and "Taconian"), the end Silurian ("Early Acadian" or "Ardennian"), mid-Devonian ("Mid-Acadian"), the end Devonian ("Late Acadian" or "Bretonnian"), the earliest Serpukhovian ("Sudetic"), and the latest Carboniferous-earliest Permian ("Alleghanian" or "Asturian"). The strongest deformations, including folding, thrusting, and active strike-slip faulting, were registered in Northwestern Africa during the last stage of the Pan-African Belt development around the

  3. 40Ar/39Ar and K-Ar data bearing on the metamorphic and tectonic history of western New England.

    USGS Publications Warehouse

    Sutter, J.F.; Ratcliffe, N.M.; Mukasa, S.B.

    1985-01-01

    40Ar/39Ar ages of coexisting biotite and hornblende from Proterozoic Y gneisses of the Berkshire and Green Mt massifs, as well as 40Ar/39Ar and K/Ar mineral and whole-rock ages from Palaeozoic metamorphic rocks, suggest that the thermal peaks for the dominant metamorphic recrystallization in western New England occurred 465 + or - 5 m.y. (Taconian). 40Ar/39Ar age data from a poorly-defined terrain along the eastern strip of the area suggests that the area has been retrograded during a metamorphism that peaked at least 376 + or - 5 m.y. (Acadian). Available age and petrological data from western New England indicate the presence of at least three separate metamorphic-structure domains of Taconic age: 1) a small area of relict high-P and low-T metamorphism, 2) a broad area of normal Barrovian metamorphism from chlorite to garnet grade characterized by a gentle metamorphic gradient and, 3) a rather narrow belt of steep-gradient, Barrovian series metamorphic rocks. Areas of maximum metamorphic intensity within the last domain coincide with areas of maximum crustal thickening in the later stage of Taconic orogeny. -L.di H

  4. Usher syndrome: definition and estimate of prevalence from two high-risk populations.

    PubMed

    Boughman, J A; Vernon, M; Shaver, K A

    1983-01-01

    The Usher Syndrome (US) refers to the combined neurosensory deficits of profound hearing impairment and retinitis pigmentosa. We have obtained information on 600 cases of deaf-blindness from the registry of the Helen Keller National Center for Deaf-Blind Youths and Adults (HKNC). Of these, 54% met the diagnostic criteria of US, although only 23.8% were so diagnosed. More extensive analysis of 189 Usher clients from HKNC showed an excess of males, some variability in audiograms, and wide ophthalmologic variation. Genetic analysis of 113 sibships showed a segregation ratio consistent with recessive inheritance. The Acadian population of Louisiana has a high frequency of US which contributes significantly to the deaf population of the state. Among 48 cases from the Louisiana School for the Deaf, there was an excess of males, more variability in audiograms than expected, and an increased segregation ratio in the 26 informative sibships. Estimates of prevalence obtained using registry data and statistics from Louisiana clearly suggest that the previous estimate of 2.4 per 100,000 is too low for the United States. Recognizing problems with ascertainment, our prevalence estimate of 4.4 per 100,000 is still considered quite conservative.

  5. Side-specific effect of yolk testosterone elevation on second-to-fourth digit ratio in a wild passerine

    NASA Astrophysics Data System (ADS)

    Nagy, Gergely; Blázi, György; Hegyi, Gergely; Török, János

    2016-02-01

    Second-to-fourth digit ratio is a widely investigated sexually dimorphic morphological trait in human studies and could reliably indicate the prenatal steroid environment. Conducting manipulative experiments to test this hypothesis comes up against ethical limits in humans. However, oviparous tetrapods may be excellent models to experimentally investigate the effects of prenatal steroids on offspring second-to-fourth digit ratio. In this field study, we injected collared flycatcher ( Ficedula albicollis) eggs with physiological doses of testosterone. Fledglings from eggs with elevated yolk testosterone, regardless of their sex, had longer second digits on their left feet than controls, while the fourth digit did not differ between groups. Therefore, second-to-fourth digit ratio was higher in the testosterone-injected group, but only on the left foot. This is the first study which shows experimentally that early testosterone exposure can affect second-to-fourth digit ratio in a wild population of a passerine bird.

  6. You mob my owl, I'll mob yours: birds play tit-for-tat game.

    PubMed

    Krama, Tatjana; Vrublevska, Jolanta; Freeberg, Todd M; Kullberg, Cecilia; Rantala, Markus J; Krams, Indrikis

    2012-01-01

    Reciprocity is fundamental to cooperative behaviour and has been verified in theoretical models. However, there is still limited experimental evidence for reciprocity in non-primate species. Our results more decisively clarify that reciprocity with a tit-for-tat enforcement strategy can occur among breeding pied flycatchers Ficedula hypoleuca separate from considerations of byproduct mutualism. Breeding pairs living in close proximity (20-24 m) did exhibit byproduct mutualism and always assisted in mobbing regardless of their neighbours' prior actions. However, breeding pairs with distant neighbours (69-84 m) either assisted or refused to assist in mobbing a predatory owl based on whether or not the distant pair had previously helped them in their own nest defense against the predator. Clearly, these birds are aware of their specific spatial security context, remember their neighbours' prior behaviour, and choose a situation-specific strategic course of action, which could promote their longer-term security, a capacity previously thought unique to primates.

  7. The risk of predation favors cooperation among breeding prey

    PubMed Central

    Krama, Tatjana; Berzins, Arnis; Rantala, Markus J

    2010-01-01

    Empirical studies have shown that animals often focus on short-term benefits under conditions of predation risk, which reduces the likelihood that they will cooperate with others. However, some theoretical studies predict that animals in adverse conditions should not avoid cooperation with their neighbors since it may decrease individual risks and increase long-term benefits of reciprocal help. We experimentally tested these two alternatives to find out whether increased predation risk enhances or diminishes the occurrence of cooperation in mobbing, a common anti-predator behavior, among breeding pied flycatchers, Ficedula hypoleuca. Our results show that birds attended mobs initiated by their neighbors more often, approached the stuffed predator significantly more closely, and mobbed it at a higher intensity in areas where the perceived risk of predation was experimentally increased. This study demonstrates a positive impact of predation risk on cooperation in breeding songbirds, which might help to explain the emergence and evolution of cooperation. PMID:20714404

  8. You mob my owl, I'll mob yours: birds play tit-for-tat game

    PubMed Central

    Krama, Tatjana; Vrublevska, Jolanta; Freeberg, Todd M.; Kullberg, Cecilia; Rantala, Markus J.; Krams, Indrikis

    2012-01-01

    Reciprocity is fundamental to cooperative behaviour and has been verified in theoretical models. However, there is still limited experimental evidence for reciprocity in non-primate species. Our results more decisively clarify that reciprocity with a tit-for-tat enforcement strategy can occur among breeding pied flycatchers Ficedula hypoleuca separate from considerations of byproduct mutualism. Breeding pairs living in close proximity (20–24 m) did exhibit byproduct mutualism and always assisted in mobbing regardless of their neighbours' prior actions. However, breeding pairs with distant neighbours (69–84 m) either assisted or refused to assist in mobbing a predatory owl based on whether or not the distant pair had previously helped them in their own nest defense against the predator. Clearly, these birds are aware of their specific spatial security context, remember their neighbours' prior behaviour, and choose a situation-specific strategic course of action, which could promote their longer-term security, a capacity previously thought unique to primates. PMID:23150772

  9. Structure of the cold- and menthol-sensing ion channel TRPM8.

    PubMed

    Yin, Ying; Wu, Mengyu; Zubcevic, Lejla; Borschel, William F; Lander, Gabriel C; Lee, Seok-Yong

    2018-01-12

    Transient receptor potential melastatin (TRPM) cation channels are polymodal sensors that are involved in a variety of physiological processes. Within the TRPM family, member 8 (TRPM8) is the primary cold and menthol sensor in humans. We determined the cryo-electron microscopy structure of the full-length TRPM8 from the collared flycatcher at an overall resolution of ~4.1 ångstroms. Our TRPM8 structure reveals a three-layered architecture. The amino-terminal domain with a fold distinct among known TRP structures, together with the carboxyl-terminal region, forms a large two-layered cytosolic ring that extensively interacts with the transmembrane channel layer. The structure suggests that the menthol-binding site is located within the voltage-sensor-like domain and thus provides a structural glimpse of the design principle of the molecular transducer for cold and menthol sensation. Copyright © 2018 The Authors, some rights reserved; exclusive licensee American Association for the Advancement of Science. No claim to original U.S. Government Works.

  10. Migrating songbirds tested in computer-controlled Emlen funnels use stellar cues for a time-independent compass.

    PubMed

    Mouritsen, H; Larsen, O N

    2001-11-01

    This paper investigates how young pied flycatchers, Ficedula hypoleuca, and blackcaps, Sylvia atricapilla, interpret and use celestial cues. In order to record these data, we developed a computer-controlled version of the Emlen funnel, which enabled us to make detailed temporal analyses. First, we showed that the birds use a star compass. Then, we tested the birds under a stationary planetarium sky, which simulated the star pattern of the local sky at 02:35 h for 11 consecutive hours of the night, and compared the birds' directional choices as a function of time with the predictions from five alternative stellar orientation hypotheses. The results supported the hypothesis suggesting that birds use a time-independent star compass based on learned geometrical star configurations to pinpoint the rotational point of the starry sky (north). In contrast, neither hypotheses suggesting that birds use the stars for establishing their global position and then perform true star navigation nor those suggesting the use of a time-compensated star compass were supported.

  11. Food supplementation mitigates dispersal-dependent differences in nest defence in a passerine bird.

    PubMed

    Récapet, Charlotte; Daniel, Grégory; Taroni, Joëlle; Bize, Pierre; Doligez, Blandine

    2016-05-01

    Dispersing and non-dispersing individuals often differ in phenotypic traits (e.g. physiology, behaviour), but to what extent these differences are fixed or driven by external conditions remains elusive. We experimentally tested whether differences in nest-defence behaviour between dispersing and non-dispersing individuals changed with local habitat quality in collared flycatchers, by providing additional food during the nestling rearing period. In control (non-food-supplemented) nests, dispersers were less prone to defend their brood compared with non-dispersers, whereas in food-supplemented nests, dispersing and non-dispersing individuals showed equally strong nest defence. We discuss the importance of dispersal costs versus adaptive flexibility in reproductive investment in shaping these differences in nest-defence behaviour between dispersing and non-dispersing individuals. Irrespective of the underlying mechanisms, our study emphasizes the importance of accounting for environmental effects when comparing traits between dispersing and non-dispersing individuals, and in turn assessing the costs and benefits of dispersal. © 2016 The Author(s).

  12. Chapter 9: The ecology of brown-headed cowbirds and their effects on southwestern willow flycatchers

    Treesearch

    J. C. Uyehara; Mary J. Whitfield; Lloyd Goldwasser

    2000-01-01

    Brown-headed cowbirds (Molothrus ater) are obligate brood-parasites, that is, female cowbirds lay their eggs in nests of other species. If the cowbird eggs are accepted, the host pair may raise the young cowbird, often at a reduction of the hosts' reproductive success. Cowbird females are also known to remove host eggs and nestlings from nests,...

  13. A field test of point relascope sampling of down coarse woody material in managed stands in the Acadian Forest

    Treesearch

    John C. Brissette; Mark J. Ducey; Jeffrey H. Gove

    2003-01-01

    We field tested a new method for sampling down coarse woody material (CWM) using an angle gauge and compared it with the more traditional line intersect sampling (LIS) method. Permanent sample locations in stands managed with different silvicultural treatments within the Penobscot Experimental Forest (Maine, USA) were used as the sampling locations. Point relascope...

  14. Vitality and Ethnolinguistic Attitudes of Acadians, Franco-Ontarians and Francophone Quebecers: Two or Three Solitudes in Canada's Bilingual Belt?

    ERIC Educational Resources Information Center

    Sioufi, Rana; Bourhis, Richard Y.; Allard, Réal

    2016-01-01

    Do French-Canadian (FC) minorities in New Brunswick and Ontario remain as committed as majority Francophone Quebecers in developing their vitality within Canada's bilingual belt? FCs constitute host communities for interprovincial migrants of FC and English-Canadian (EC) background who can bolster or weaken the vitality of FCs. How FCs and ECs…

  15. Indicators and standards of quality across space and time, as reflected in a study in Acadian National Park, Maine

    Treesearch

    Laura E. Anderson; Robert E. Manning; William A. Valliere

    2009-01-01

    Recreation indicators and standards of quality may be influenced by: 1) the popularity (or overall use level) of a recreation area and 2) the time of day that an individual chooses to visit a popular site. To examine these connections, data were collected via on-site self-administered surveys at several "high"- and "low"-use sites in Acadia National...

  16. Modelling metamorphism in the Hoosac Schist, Western Massachusetts: new approaches to a New England problem

    NASA Astrophysics Data System (ADS)

    Bidgood, Anna; Waters, Dave; Gardiner, Nick

    2015-04-01

    Along the western margin of the metamorphic Appalachians in New England, Taconic (Ordovician) tectonism and metamorphism are overprinted towards the east by Acadian (Devonian) structures and metamorphism. The Hoosac Schist, a probable correlate of the well-known Gassetts Schist of Vermont, lies in the region of overprinting. It forms a narrow N-S-trending tectonically-bound zone crossing several Barrovian mineral-assemblage zones from garnet to kyanite grade. Highly aluminous units containing cm-sized garnets (Cheney & Brady, 1992) are noted for the occurrence of textural unconformities within the garnets, separating inclusion-rich cores from inclusion-poor rims. Matrix domains contain both paragonite and muscovite. Muscovite is present in at least two compositionally distinct generations, with broad later laths cutting across a microfolded earlier fabric. Rutile is restricted to inclusions in garnet, whereas the matrix Ti- phase is ilmenite. These features suggest a polymetamorphic history, potentially recording the superimposition of Acadian metamorphism on Taconic, but it has not yet proved possible to demonstrate the presence of two metamorphic cycles. This study aims to test and employ the new and revised activity models recently developed for metapelites in the full system MnNCKFMASHTO (White et al, 2014), for use with the Holland & Powell data-set 6. Features that can now be more explicitly modelled include garnet zonation in relation to its inclusion suites and microstructural features, the occurrence, texture and distribution of Ti-bearing accessory minerals, and the assemblages and compositional trends in white micas. Preliminary modelling, correlated with microstructural observation, indicates (1) some confirmation of the concern expressed by White et al (2014) that the stability of margarite-bearing assemblages may be somewhat overestimated, (2) that apart from this, the early growth history of garnet is consistent with its suite of trapped inclusions

  17. Production of high-resolution forest-ecosite maps based on model predictions of soil moisture and nutrient regimes over a large forested area.

    PubMed

    Yang, Qi; Meng, Fan-Rui; Bourque, Charles P-A; Zhao, Zhengyong

    2017-09-08

    Forest ecosite reflects the local site conditions that are meaningful to forest productivity as well as basic ecological functions. Field assessments of vegetation and soil types are often used to identify forest ecosites. However, the production of high-resolution ecosite maps for large areas from interpolating field data is difficult because of high spatial variation and associated costs and time requirements. Indices of soil moisture and nutrient regimes (i.e., SMR and SNR) introduced in this study reflect the combined effects of biogeochemical and topographic factors on forest growth. The objective of this research is to present a method for creating high-resolution forest ecosite maps based on computer-generated predictions of SMR and SNR for an area in Atlantic Canada covering about 4.3 × 10 6 hectares (ha) of forestland. Field data from 1,507 forest ecosystem classification plots were used to assess the accuracy of the ecosite maps produced. Using model predictions of SMR and SNR alone, ecosite maps were 61 and 59% correct in identifying 10 Acadian- and Maritime-Boreal-region ecosite types, respectively. This method provides an operational framework for the production of high-resolution maps of forest ecosites over large areas without the need for data from expensive, supplementary field surveys.

  18. The emplacement, geochemistry and petrogenesis of two central Morocco Hercynian granites. Geotectonic implications

    NASA Astrophysics Data System (ADS)

    Giuliani, G.; Cheilletz, A.; Zimmermann, J. L.

    New field, petrographic and geochemical data including REE, Rb-Sr, and K-Ar, are presented concerning the emplacement and petrogenesis of two calc-alkaline Moroccan Hercynian granites: the Zaër pluton and the Djebel Aouam stocks. Zonation in the Zaër pluton does not appear to result from simple fractional crystallization nor from hydrothermal alteration but is rather the diapiric intrusion of two interlocked bodies. REE geochemistry supports the interpretation that the biotite-granodiorite magma (301 ± 8.2 M.a.) and the Djebel Aouam stocks appear to have been derived by fusion of crustal materials with possible mantle contamination ( Initial87Sr/ 86Srratio = 0.70514 ). The second two-mica granitic magma (283.4 ± 6.2 M.a.) corresponds to a peraluminous granite (1.22 < A/ CNK < 1.33) generated by fusion of continental crust including mature sedimentarr materials (Initial 87Sr/ 86Sr ratio = 0.70836). These granites are compared to the Acadian granites of north Appalachian and classified in the «post-collision, uplift environmentå type. An illustrative four step model presenting the generation and emplacement of these two granites together with the associated W-Sn-Pb-Zn-Ag mineralizations is proposed: (1) first magma generation, (2) first melt emplacement, (3) second melt emplacement, (4) late convective flow.

  19. Mate choice and imprinting in birds studied by cross-fostering in the wild.

    PubMed Central

    Slagsvold, Tore; Hansen, Bo T; Johannessen, Lars E; Lifjeld, Jan T

    2002-01-01

    Sexual-selection theories generally assume that mating preferences are heritable traits. However, there is substantial evidence that the rearing environment may be important for the development of mating preferences, indicating that they may be learnt, or modified by experience. The relative importance of such sexual imprinting across species remains largely unexplored. Here, we report results of a large-scale cross-fostering experiment in the wild in which nestling birds were raised by parents of a different species. We show that resulting sexual imprinting may have a negative effect on pairing success in one species (the great tit, Parus major), but not in two other species (the blue tit, P. caeruleus and the pied flycatcher, Ficedula hypoleuca). A remarkable variation thus seems to exist, even between species that are congeneric and have similar breeding ecologies. The cross-fostering resulted in heterospecific pairings between the two tit species (female blue tit breeding with male great tit), which has never, to our knowledge, been previously documented. However, the chicks fledging from these nests were all blue tit. PMID:12137574

  20. The excuse principle can maintain cooperation through forgivable defection in the Prisoner's Dilemma game

    PubMed Central

    Krams, Indrikis; Kokko, Hanna; Vrublevska, Jolanta; Āboliņš-Ābols, Mikus; Krama, Tatjana; Rantala, Markus J.

    2013-01-01

    Reciprocal altruism describes a situation in which an organism acts in a manner that temporarily reduces its fitness while increasing another organism's fitness, but there is an ultimate fitness benefit based on an expectation that the other organism will act in a similar manner at a later time. It creates the obvious dilemma in which there is always a short-term benefit to cheating, therefore cooperating individuals must avoid being exploited by non-cooperating cheaters. This is achieved by following various decision rules, usually variants of the tit-for-tat (TFT) strategy. The strength of TFT, however, is also its weakness—mistakes in implementation or interpretation of moves, or the inability to cooperate, lead to a permanent breakdown in cooperation. We show that pied flycatchers (Ficedula hypoleuca) use a TFT with an embedded ‘excuse principle’ to forgive the neighbours that were perceived as unable to cooperate during mobbing of predators. The excuse principle dramatically increases the stability of TFT-like behavioural strategies within the Prisoner's Dilemma game. PMID:23864603

  1. Shiny cowbird parasitism in two avian communities in Puerto Rico

    USGS Publications Warehouse

    Wiley, J.W.

    1985-01-01

    The shiny cowbird (M. bonariensis), a brood parasite, has recently expanded its range from South America to Puerto Rico via the Lesser Antilles. This species is a host generalist and, on reaching Puerto Rico, encounteed avian species with no history of social parasitism. In mangrove habitat study areas, 42% of the resident non-raptorial land bird species were parasitized. Some species were heavily parasitized; e.g., yellow warbler (Dendroica petechia), 76% of nests parasitized black-whiskered vireo (Vireo altiloquus), 82%, Puerto Rican flycatcher (Myiarchus antillarum), 85%, yellow-shouldered blackbird (Agelaius xanthomus), 95%, troupial (Icterus icterus), 100%, black-cowled oriole (I. dominicensis), 100%. Others suffered low rates of parasitism (2-17% of nests examined); e.g., gray kingbird (Tyrannus dominicensis), red-legged thrush (Turdus plumbeus), bronze mannikin (Lonchura cucullata), northern mockingbird (Mimus polyglottos), greater antillean grackle (Quiscalus niger). Cowbird parasitism affected hosts by depressing nest success an average of 41% below non-parasitized nests and reducing host productivity. Parasitized host nests hatched 12% fewer eggs an fledged 67% fewer of their own chicks than non-parasitized pairs.

  2. Ultra-Rapid Vision in Birds.

    PubMed

    Boström, Jannika E; Dimitrova, Marina; Canton, Cindy; Håstad, Olle; Qvarnström, Anna; Ödeen, Anders

    2016-01-01

    Flying animals need to accurately detect, identify and track fast-moving objects and these behavioral requirements are likely to strongly select for abilities to resolve visual detail in time. However, evidence of highly elevated temporal acuity relative to non-flying animals has so far been confined to insects while it has been missing in birds. With behavioral experiments on three wild passerine species, blue tits, collared and pied flycatchers, we demonstrate temporal acuities of vision far exceeding predictions based on the sizes and metabolic rates of these birds. This implies a history of strong natural selection on temporal resolution. These birds can resolve alternating light-dark cycles at up to 145 Hz (average: 129, 127 and 137, respectively), which is ca. 50 Hz over the highest frequency shown in any other vertebrate. We argue that rapid vision should confer a selective advantage in many bird species that are ecologically similar to the three species examined in our study. Thus, rapid vision may be a more typical avian trait than the famously sharp vision found in birds of prey.

  3. Controls of late Palaeozoic and Mesozoic extension in the British Isles: evidence from seismic reflection data in the Central North Sea

    NASA Astrophysics Data System (ADS)

    Smith, K.; Cameron, T. D. J.

    2009-04-01

    Controls of late Palaeozoic and Mesozoic extension in the British Isles: evidence from seismic reflection data in the Central North Sea. Kevin Smith (1) and Don Cameron (2) (1) British Geological Survey, Murchison House, West Mains Road, Edinburgh, EH9 3LA. (ksm@bgs.ac.uk). (2) British Geological Survey, 376 Gilmerton Road, Edinburgh, EH17 7QS. In the area of the British Isles during the late Devonian and early Carboniferous, the oblique convergence of Laurentia and Gondwana imposed a torque on the adjoining terranes of Baltica and Avalonia. Their resulting clockwise rotation was accommodated by widely distributed N-S extension in the intervening zones previously formed by Caledonian and Acadian convergence. South of Laurentia and Baltica, late Palaeozoic extension was focused (1) at terrane margins, (2) in areas of limited Caledonian-Acadian plutonism, and (3) in places where the western (Iapetus) and eastern (Tornquist) convergence zones intersect at a high angle. One of these latter areas lies in Central England immediately north of the Midland Microcraton (part of Eastern Avalonia), where thermal subsidence associated with early Carboniferous extension gave rise to the late Carboniferous Pennine Basin. Interpretation of an extensive set of 3D and 2D long-offset seismic reflection data suggests that a similar area of enhanced extension at a fold belt intersection lies to north of the Mid North Sea High in the middle of the Central North Sea. Variscan uplift and inversion of the late Palaeozoic basins began to predominate in mid-Carboniferous times as final amalgamation of all the different terranes to form Pangaea curtailed the initial episode of extension and thermal subsidence. This change in the tectonic regime was associated with the onset of tholeiitic volcanism within the convergence zones, and was followed by localised extension during the earliest Permian. Evidence obtained from seismic interpretation of the deep structure of the UK sector of the Central

  4. Glastonbury Gneiss and mantling rocks (a modified Oliverian dome) in south-central Massachusetts and north-central Connecticut; geochemistry, petrogenesis, and isotopic age

    USGS Publications Warehouse

    Leo, G.W.; Zartman, R.E.; Brookins, D.G.

    1984-01-01

    The Glastonbury dome is a long, narrow structure trending approximately 70 km north-northeast through Connecticut and Massachusetts along the west side of the Bronson Hill anticlinorium. Structurally and stratigraphically the dome is analogous to the Oliverian domes of New Hampshire. It is cored by Glastonbury Gneiss and is mantled by Ammonoosuc Volcanics and Partridge Formation (or their equivalents) of Ordovician age. The Glastonbury Gneiss intrudes the Ammonoosuc and, thereby, establishes the relative age of the two units. Monson Gneiss, which unconformably underlies the Ammonoosuc Volcanics in the Monson anticline to the east, is not in contact with Glastonbury Gneiss except near Stafford Springs, Conn., where the contact may be gradational. In some places, Monson Gneiss shows evidence of plastic flow and potential anatexis. The northern part of the Glastonbury Gneiss typically is leucocratic, granoblastic, relatively potassium-poor gneiss that appears homogeneous in outcrop, but proves to be chemically and modally inhomogeneous over short distances, as shown by variation diagrams and REE plots. The gneiss straddles the compositional fields of trondhjemite, tonalite, and granodiorite, and partly overlaps that of Monson Gneiss. The southern part of the Glastonbury Gneiss is consistently more potassic than the northern, having compositions ranging from granite to granodiorite. All of the Glastonbury Gneiss show pervasive, strong foliation, deformation, and local shearing related to the Acadian orogeny. Field relations, textures, and chemistry of the northern part of the Glastonbury suggest an origin by anatexis of the premetamorphic Monson sequence at temperatures of about 690 DC to 750 DC and pressures of <3kbars. The southern part of the Glastonbury appears to have been generated contemporaneously but not comagmatically from calcalkaline crust. U-Pb zircon ages for both the northern and southern bodies are slightly discordant with 207PbfosPb ages of 445 to 467

  5. Textural and Rb-Sr isotopic evidence for late Paleozoic mylonitization within the Honey Hill fault zone southeastern Connecticut

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    O'Hara, K.D.; Gromet, L.P.

    A petrographic and Rb-Sr isotopic study of rocks within and near the Honey Hill fault zone places important constraints on its history of movement. Rb-Sr apparent ages for micas and plagioclase from these rocks have been reset and range from Permian to Triassic, considerably younger than the minimum stratigraphic age (Ordovician) of the rocks studied or of Acadian (Devonian) regional metamorphism. Permian Rb-Sr ages of dynamically recrystallized muscovite date the development of mylonite fabric. An older age is precluded by the excellent preservation of unrecovered quartz, which indicates that these rocks did not experience temperatures high enough to anneal quartzmore » or thermally reset Rb-Sr isotopic systems in muscovite since the time of mylonitization. Metamorphic mineral assemblages and mineral apparent ages in rocks north of the fault zone indicate recrystallization under similar upper greenschist-lower amphibolite grade conditions during Permian to Triassic time. Collectively these results indicate that the Honey Hill fault zone was active during the Late Paleozoic and that ductile deformation and metamorphism associated with the Alleghanian orogeny extend well into southern Connecticut. An Alleghanian age for mylonitization within the Honey Hill fault zone suggests it should be considered as a possible site for the major Late Paleozoic strike-slip displacements inferred from paleomagnetic studies for parts of coastal New England and maritime Canada.« less

  6. Upper Devonian outcrop stratigraphy, southwestern Virginia and southeastern West Virginia

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Dennison, J.M.; Filer, J.K.; Rossbach, T.J.

    Ongoing outcrop studies are resulting in the extension of existing formal lithostratigraphic units and revision of previously less refined subdivisions of Upper Devonian strata in southwestern Virginia and southeastern West Virginia. A 425 km (263 mi) long stratigraphic cross-section has been constructed primarily from the outcrop belt along the Allegheny Structural Front, supplemented by sections from nearby outcrop belts. This NE-SW striking cross-section is oblique to the nearly due N-S depositional strike of the Upper Devonian Acadian orogenic wedge. To the southwest, the Upper Devonian section thins from 2,100 meters (6,900 feet) to 230 meters (756 feet) as progressively moremore » distal deposits are encountered. An integrated approach has been taken to establish chronostratigraphic control within the cross-section. The best time markers include particularly regressive parasequences which can be identified across facies boundaries (especially the Pound and Briery Gap Sandstones and their equivalents), volcanic ashes, and an organic-rich shale zone marking the base of a major transgression (equivalent to the base of the Huron Shale in Ohio and the Dunkirk Shale of New York). These tools provide chronostratigraphic correlation through the undivided Brallier Formation. Supplemental control includes biostratigraphic markers as well as marine dull redbeds within the Foreknobs which parallel other time lines and may represent partially reduced influxes of oxidized coastal plain sediments during minor parasequence scale regressions.« less

  7. Pre-lithification tectonic foliation development in a clastic sedimentary rock sequence from SW Ireland

    NASA Astrophysics Data System (ADS)

    Meere, Patrick; Mulchrone, Kieran; McCarthy, David

    2017-04-01

    The current orthodoxy regarding the development of regionally developed penetrative tectonic cleavage fabrics in sedimentary rocks is that it postdates lithification of those rocks. It is well established that fabric development under these circumstances is achieved by a combination of grain rigid body rotation, crystal-plastic deformation and pressure solution. The latter is believed to be the primary mechanism responsible for the domainal nature of cleavage development commonly observed in low grade metamorphic rocks. While there have been advocates for the development of tectonic cleavages before host rock lithification these are currently viewed as essentially local aberrations without regional significance. In this study we combine new field observations with strain analysis, element mapping and modelling to characterise Acadian (>50%) crustal shortening in a Devonian clastic sedimentary sequence from the Dingle Peninsula of south west Ireland. Fabrics in these rocks reflect significant levels of tectonic shortening are a product of grain translation, rigid body rotation and repacking of intra- and extra-formational clasts during deformation of an unconsolidated clastic sedimentary sequence. There is an absence of the expected domainal cleavage structure and intra-clast deformation expected with conventional cleavage formation. This study requires geologists to consider the possibility such a mechanism contributing to tectonic strain in a wide range of geological settings and to look again at field evidence that indicates early sediment mobility during deformation.

  8. Geographical Variation in Egg Mass and Egg Content in a Passerine Bird

    PubMed Central

    Ruuskanen, Suvi; Siitari, Heli; Eeva, Tapio; Belskii, Eugen; Järvinen, Antero; Kerimov, Anvar; Krams, Indrikis; Moreno, Juan; Morosinotto, Chiara; Mänd, Raivo; Möstl, Erich; Orell, Markku; Qvarnström, Anna; Salminen, Juha-Pekka; Slater, Fred; Tilgar, Vallo; Visser, Marcel E.; Winkel, Wolfgang; Zang, Herwig; Laaksonen, Toni

    2011-01-01

    Reproductive, phenotypic and life-history traits in many animal and plant taxa show geographic variation, indicating spatial variation in selection regimes. Maternal deposition to avian eggs, such as hormones, antibodies and antioxidants, critically affect development of the offspring, with long-lasting effects on the phenotype and fitness. Little is however known about large-scale geographical patterns of variation in maternal deposition to eggs. We studied geographical variation in egg components of a passerine bird, the pied flycatcher (Ficedula hypoleuca), by collecting samples from 16 populations and measuring egg and yolk mass, albumen lysozyme activity, yolk immunoglobulins, yolk androgens and yolk total carotenoids. We found significant variation among populations in most egg components, but ca. 90% of the variation was among individuals within populations. Population however explained 40% of the variation in carotenoid levels. In contrast to our hypothesis, we found geographical trends only in carotenoids, but not in any of the other egg components. Our results thus suggest high within-population variation and leave little scope for local adaptation and genetic differentiation in deposition of different egg components. The role of these maternally-derived resources in evolutionary change should be further investigated. PMID:22110579

  9. Search behavior of arboreal insectivorous migrants at gulf coast stopover sites in spring

    USGS Publications Warehouse

    Chen, Chao-Chieh; Barrow, W.C.; Ouchley, K.; Hamilton, R.B.

    2011-01-01

    Search behavior of arboreal insectivorous migrants was studied at three stopover sites along the northern coast of the Gulf of Mexico during spring migrations, 1993–1995. We examined if search behavior was affected by phylogeny, or by environmental factors. A sequence of search movements (hop, flutter, or flight) in a foraging bout was recorded for each migrant encountered. Search rate, frequency, and distance of movements were calculated for each species. Search rate was positively correlated with proportion of hop, but negatively correlated to flight distance. Hop distance was positively correlated to tarsus length, as was flight distance to wing length for the 31 species of migrants. Cluster analysis indicated closely related species generally have similar foraging modes, which range from “sit-and-wait” of flycatchers to “widely foraging” of warblers. Migrants tended to use more hops in dense vegetation, but more flights in areas with sparse vegetation. Migrants also used more flights when foraging in mixed-species flocks and during periods of high migrant density. Logistic models indicated warblers were more influenced by environmental factors than vireos, possibly because warblers are near-perch searchers and more affected by these factors.

  10. The increased risk of predation enhances cooperation

    PubMed Central

    Krams, Indrikis; Bērziņš, Arnis; Krama, Tatjana; Wheatcroft, David; Igaune, Kristīne; Rantala, Markus J.

    2010-01-01

    Theory predicts that animals in adverse conditions can decrease individual risks and increase long-term benefits by cooperating with neighbours. However, some empirical studies suggest that animals often focus on short-term benefits, which can reduce the likelihood that they will cooperate with others. In this experimental study, we tested between these two alternatives by evaluating whether increased predation risk (as a correlate of environmental adversity) enhances or diminishes the occurrence of cooperation in mobbing, a common anti-predator behaviour, among breeding pied flycatchers Ficedula hypoleuca. We tested whether birds would join their mobbing neighbours more often and harass a stuffed predator placed near their neighbours' nests more intensely in areas with a higher perceived risk of predation. Our results show that birds attended mobs initiated by their neighbours more often, approached the stuffed predator significantly more closely, and mobbed it at a higher intensity in areas where the perceived risk of predation was experimentally increased. In such high-risk areas, birds also were more often involved in between-pair cooperation. This study demonstrates the positive impact of predation risk on cooperation in breeding songbirds, which might help in explaining the emergence and evolution of cooperation. PMID:19846454

  11. Artificial cavities enhance breeding bird densities in managed cottonwood forests

    USGS Publications Warehouse

    Twedt, D.J.; Henne-Kerr, J.L.

    2001-01-01

    The paucity of natural cavities within short-rotation hardwood agroforests restricts occupancy by cavity-nesting birds. However, providing 1.6 artificial nesting cavities (nest boxes)/ha within 3- to 10-year-old managed cottonwood forests in the Mississippi Alluvial Valley increased territory density of cavity-nesting birds. Differences in territory densities between forests with and without nest boxes increased as stands aged. Seven bird species initiated 38 nests in 173 boxes during 1997 and 39 nests in 172 boxes during 1998. Prothonotary warblers (Protonotaria citrea) and eastern bluebirds (Sialia sialis) accounted for 67% of nests; nearly all warbler nests were in 1.8-L, plastic-coated cardboard (paper) boxes, whereas bluebird nests were divided between paper boxes and 3.5-L wooden boxes. Larger-volume (16.5-L) wooden nest boxes were used by eastern screech owls (Otus asio) and great crested flycatchers (Myiarchus crinitus), but this box type often was usurped by honey bees (Apis mellifera). To enhance territory densities of cavity-nesting birds in cottonwood agroforests, we recommend placement of plastic-coated paper nest boxes, at a density of 0.5/ha, after trees are >4 years old but at least 2 years before anticipated timber harvest.

  12. Manipulation of parental effort affects plumage bacterial load in a wild passerine.

    PubMed

    Alt, Grete; Saag, Pauli; Mägi, Marko; Kisand, Veljo; Mänd, Raivo

    2015-06-01

    It has been suggested that plumage microorganisms play an important role in shaping the life histories of wild birds. Some bacteria may act as pathogens or cause damage to feathers, and thereby reduce individual fitness. Intense parental care in birds can result in a reduction of self-maintenance and preening behavior in parents and therefore might affect the dynamics of microbiota living on their feathers. However, experimental evidence of this relationship is virtually absent. We manipulated the parental effort of wild breeding pied flycatcher (Ficedula hypoleuca) females by modifying their brood size or temporarily removing male partners. We expected that experimentally decreasing or increasing parental effort would affect feather sanitation in females and therefore also bacterial density on their plumage. In accordance with this hypothesis, manipulation affected the density of free-living bacteria: females with reduced broods had the lowest number of free-living bacteria on their feathers, while females left without male partners had the highest. However, manipulation did not have a significant effect on the densities of attached bacteria. Our results provide experimental evidence that a trade-off between self-maintenance and parental effort affects plumage bacterial densities in birds.

  13. Environmental pollution has sex-dependent effects on local survival

    PubMed Central

    Eeva, Tapio; Hakkarainen, Harri; Laaksonen, Toni; Lehikoinen, Esa

    2006-01-01

    Environmental pollutants cause a potential hazard for survival in free-living animal populations. We modelled local survival (including emigration) by using individual mark–recapture histories of males and females in a population of a small insectivorous passerine bird, the pied flycatcher (Ficedula hypoleuca) living around a point source of heavy metals (copper smelter). Local survival of F. hypoleuca females did not differ between polluted and unpolluted environments. Males, however, showed a one-third higher local-survival probability in the polluted area. Low fledgling production was generally associated with decreased local survival, but males in the polluted area showed relatively high local survival, irrespective of their fledgling number. A possible explanation of higher local survival of males in the polluted area could be a pollution-induced change in hormone (e.g. corticosterone or testosterone) levels of males. It could make them to invest more on their own survival or affect the hormonal control of breeding dispersal. The local survival of males decreased in the polluted area over the study period along with the simultaneous decrease in heavy metal emissions. This temporal trend is in agreement with the stress hormone hypothesis. PMID:17148387

  14. Integration of Spectral Reflectance across the Plumage: Implications for Mating Patterns

    PubMed Central

    Laczi, Miklós; Török, János; Rosivall, Balázs; Hegyi, Gergely

    2011-01-01

    Background In complex sexual signaling systems such as plumage color, developmental or genetic links may occur among seemingly distinct traits. However, the interrelations of such traits and the functional significance of their integration rarely have been examined. Methodology/Principal Findings We investigated the parallel variation of two reflectance descriptors (brightness and UV chroma) across depigmented and melanized plumage areas of collared flycatchers (Ficedula albicollis), and the possible role of integrated color signals in mate acquisition. We found moderate integration in brightness and UV chroma across the plumage, with similar correlation structures in the two sexes despite the strong sexual dichromatism. Patterns of parallel color change across the plumage were largely unrelated to ornamental white patch sizes, but they all showed strong assortative mating between the sexes. Comparing different types of assortative mating patterns for individual spectral variables suggested a distinct role for plumage-level color axes in mate acquisition. Conclusions/Significance Our results indicate that the plumage-level, parallel variation of coloration might play a role in mate acquisition. This study underlines the importance of considering potential developmental and functional integration among apparently different ornaments in studies of sexual selection. PMID:21853088

  15. Comparative study of understorey birds diversity inhabiting lowland rainforest virgin jungle reserve and regenerated forest.

    PubMed

    Nor Hashim, Ezyan; Ramli, Rosli

    2013-01-01

    A comparative study of understorey birds inhabiting different habitats, that is, virgin jungle reserve (VJR) and regenerated forest (RF), was conducted in Ulu Gombak Forest Reserve and Selangor and Triang Forest Reserve, Negeri Sembilan, Peninsular Malaysia. The objective of this study was to assess the diversity of understorey birds in both habitats and the effects of forest regeneration on the understorey bird community. The mist-netting method was used to capture understorey birds inhabiting both habitats in both locations. Species composition and feeding guild indicated that understorey bird populations were similar in the two habitats. However, the number of secondary forest species such as Little spiderhunter (Arachnothera longirostra) in VJR is increasing due to its proximity to RF. This study discovered that RFs in both study areas are not yet fully recovered. However, based on the range of species discovered, the RFs have conservation value and should be maintained because they harbour important forest species such as babblers and flycatchers. The assessment of the community structure of understorey birds in VJR and RF is important for forest management and conservation, especially where both habitats are intact.

  16. Comparative Study of Understorey Birds Diversity Inhabiting Lowland Rainforest Virgin Jungle Reserve and Regenerated Forest

    PubMed Central

    Nor Hashim, Ezyan; Ramli, Rosli

    2013-01-01

    A comparative study of understorey birds inhabiting different habitats, that is, virgin jungle reserve (VJR) and regenerated forest (RF), was conducted in Ulu Gombak Forest Reserve and Selangor and Triang Forest Reserve, Negeri Sembilan, Peninsular Malaysia. The objective of this study was to assess the diversity of understorey birds in both habitats and the effects of forest regeneration on the understorey bird community. The mist-netting method was used to capture understorey birds inhabiting both habitats in both locations. Species composition and feeding guild indicated that understorey bird populations were similar in the two habitats. However, the number of secondary forest species such as Little spiderhunter (Arachnothera longirostra) in VJR is increasing due to its proximity to RF. This study discovered that RFs in both study areas are not yet fully recovered. However, based on the range of species discovered, the RFs have conservation value and should be maintained because they harbour important forest species such as babblers and flycatchers. The assessment of the community structure of understorey birds in VJR and RF is important for forest management and conservation, especially where both habitats are intact. PMID:24453888

  17. Influence of partial harvesting and site factors on the abundance and composition of natural regeneration in the Acadian Forest of Maine, USA

    Treesearch

    Mohammad Bataineh; Laura Kenefic; Aaron Weiskittel; Robert Wagner; John Brissette

    2013-01-01

    Understanding the factors regulating the composition and abundance of natural regeneration in forest ecosystems is critical to sustainable management worldwide. Using a long-term silvicultural experiment in Maine, we partitioned the variation in natural regeneration and examined the contribution of overstory and understory vegetation (biotic factors), substrate and...

  18. Louisiana coastal ecosystem

    USGS Publications Warehouse

    ,

    2000-01-01

    Louisiana's coast and its degradation and restoration are major environmental issues being studied at the National Wetlands Research Center. Coastal ecosystems are vulnerable because of the tremendous amount of human activity that takes place along the coast. Information on ecological processes is essential to guide the development along the coast as well as to protect and restore wildlife habitat.Louisiana has about 40% of coastal wetlands in the lower 48 states; they support fish, waterfowl, and fur-bearing animals as well as unique cultures like that of the Acadians. The fish and wildlife resources of Louisiana's coast produce over $1 billion each year in revenues.But Louisiana has the highest coastal loss rate because of natural and human causes. Over the past three decades, Louisiana has lost as much as 35-40 mi2 (90-104 km2) of coastal wetlands a year.The National Wetlands Research Center is qualified to assess and monitor this ecosystem because of its proximity to the study area, a staff chosen for their expertise in the system, and a number of established partnerships with others who study the areas. The Center is often the lead group in partnerships with universities, other federal agencies, and private entities who study this ecosystem.Most of the Center's research and technology development performed for coastal wetlands are done at the Lafayette headquarters; some work is performed at the National Wetlands Research Center's project office in Baton Rouge, LA.

  19. Chapter 4: The dynamic environmental history of southwest willow flycatcher habitat: A survey of changing riparian conditions through time

    Treesearch

    Richard D. Periman; Jeffrey F. Kelly

    2000-01-01

    The extent of riparian habitat in the southwestern United States during the past 100 years appears to have been reduced by modern land development, urban expansion, and a general increase in human populations (Finch 1996; Shaw and Finch 1996). Thousands of acres of river flood plain along the Rio Grande and Colorado rivers have been cleared for agriculture,...

  20. Thirty-seventh supplement to the American Ornithologists' Union Check-list of North American birds

    USGS Publications Warehouse

    Monroe, Burt L.; Banks, Richard C.; Fitzpatrick, John W.; Howell, Thomas R.; Johnson, Ned K.; Ouellet, Henri; Remsen, J.V.; Storer, Robert W.

    1989-01-01

    This third supplement subsequent to the 6th edition (1983) of the A.O.U. "Check-list of North American Birds" consists of changes adopted by the Committee on Classification and Nomenclature as of 1 March 1989. The changes fall into nine categories: (1) six species are added to the main list (Pterodroma longirostris, Larus crassirostris, Streptopelia decaocto, Cocccyzus julieni, Chrysolampis mosquitus, Emberiza aureola) because of new distributional information; (2) five species (Ara cubensis, Chlorostilbon bracei, Empidonax occidentalis, Polioptila californica, Pipilo crissalis) are added to the main list because of the splitting of species already on the list; (3) one name (Anthus rubescens) is changed because of the splitting of a species from outside the Checklist area; (4) two names (Morus bassanus, Nyctanassa violacea) is removed from the main list to Appendix B because of re-evaluation of Northern Hemisphere records; (6) three species (Pterodrama rostrata, P. alba, P. solandri) are moved from Appendix A to Appendix B, and one (P. defilippiana) is added to Appendix B because of questionable sight records; (7)A.O.U. numbers are added to three species (Ciccaba virgata, Myiopagis viridicata, Molothrus bonariensis) on the basis on new distributional records or supporting data; (8) several corrections in spelling or citations are made; and (9) English names are changed for twelve species to accommodate worldwide usage of these names. No new distributional information is included except as indicated above (i.e. minor changes of distribution are not noted). These actions bring the number of species recognized as occurring in North America (main list) to 1,945.

  1. "Le Cochon", a Washed up Mermaid, and Soldier's Bread: Three Generations of Acadian Women at the Intersection of Museums and Fiction

    ERIC Educational Resources Information Center

    Taber, Nancy

    2018-01-01

    In this autoethnographic narrative article, I apply a feminist antimilitarist lens to discuss how the critical learning that museums can facilitate can provide a powerful pedagogic lens into the past and present, particularly when connected to self. These sites can illuminate the strength of women as they encounter/ed individual and societal…

  2. Effects of intensity and frequency of harvesting on abundance, stocking and composition of natural regeneration in the Acadian Forest of eastern North America

    Treesearch

    John C. Brissette

    1996-01-01

    In a silviculture experiment in east-central Maine, USA, natural regeneration was sampled to measure the effects of: (1) a range of partial harvest intensities, and (2) repeated partial harvest at one intensity. Under the first objective, five treatments were compared with residual basal areas ranging from 15 to 24 m2 ha-1...

  3. Eastern hemlock response to even- and uneven-age management in the Acadian forest: results from the Penobscot Experimental Forest long-term silviculture study

    Treesearch

    John C. Brissette; Laura S. Kenefic

    2000-01-01

    Eastern hemlock (Tsuga canadensis (L.) Carr.) is an important tree species in the mixed-species conifer forests of northern New England and adjacent Canada. Hemlock is very tolerant of understory conditions; consequently, it responds differently to various silvicultural treatments. In a long-term study at the Penobscot Experimental Forest in east-...

  4. Aggressive behavior of the male parent predicts brood sex ratio in a songbird

    NASA Astrophysics Data System (ADS)

    Szász, Eszter; Garamszegi, László Zsolt; Hegyi, Gergely; Szöllősi, Eszter; Markó, Gábor; Török, János; Rosivall, Balázs

    2014-08-01

    Brood sex ratio is often affected by parental or environmental quality, presumably in an adaptive manner that is the sex that confers higher fitness benefits to the mother is overproduced. So far, studies on the role of parental quality have focused on parental morphology and attractiveness. However, another aspect, the partner's behavioral characteristics, may also be expected to play a role in brood sex ratio adjustment. To test this hypothesis, we investigated whether the proportion of sons in the brood is predicted by the level of territorial aggression displayed by the father, in the collared flycatcher ( Ficedula albicollis). The proportion of sons in the brood was higher in early broods and increased with paternal tarsus length. When controlling for breeding date and body size, we found a higher proportion of sons in the brood of less aggressive fathers. Male nestlings are more sensitive to the rearing environment, and the behavior of courting males may often be used by females to assess their future parental activity. Therefore, adjusting brood sex ratio to the level of male aggression could be adaptive. Our results indicate that the behavior of the partner could indeed be a significant determinant in brood sex ratio adjustment, which should not be overlooked in future studies.

  5. Predator encounters have spatially extensive impacts on parental behaviour in a breeding bird community

    PubMed Central

    Moks, Kadri; Thomson, Robert L.; Calhim, Sara; Järvistö, Pauliina E.; Schuett, Wiebke; Velmala, William; Laaksonen, Toni

    2016-01-01

    Predation risk has negative indirect effects on prey fitness, partly mediated through changes in behaviour. Evidence that individuals gather social information from other members of the population suggests that events in a community may impact the behaviour of distant individuals. However, spatially wide-ranging impacts on individual behaviour caused by a predator encounter elsewhere in a community have not been documented before. We investigated the effect of a predator encounter (hawk model presented at a focal nest) on the parental behaviour of pied flycatchers (Ficedula hypoleuca), both at the focal nest and at nearby nests different distances from the predator encounter. We show that nest visitation of both focal pairs and nearby pairs were affected, up to 3 h and 1 h, respectively. Parents also appeared to compensate initial disrupted feeding by later increasing nest visitation rates. This is the first evidence showing that the behaviour of nearby pairs was affected away from an immediate source of risk. Our results indicate that the impacts of short-term predator encounters may immediately extend spatially to the broader community, affecting the behaviour of distant individuals. Information about predators is probably quickly spread by cues such as intra- and heterospecific alarm calls, in communities of different taxa. PMID:27030411

  6. Convergent evolution of Hawaiian and Australo-Pacific honeyeaters from distant songbird ancestors.

    PubMed

    Fleischer, Robert C; James, Helen F; Olson, Storrs L

    2008-12-23

    The Hawaiian "honeyeaters," five endemic species of recently extinct, nectar-feeding songbirds in the genera Moho and Chaetoptila, looked and acted like Australasian honeyeaters (Meliphagidae), and no taxonomist since their discovery on James Cook's third voyage has classified them as anything else. We obtained DNA sequences from museum specimens of Moho and Chaetoptila collected in Hawaii 115-158 years ago. Phylogenetic analysis of these sequences supports monophyly of the two Hawaiian genera but, surprisingly, reveals that neither taxon is a meliphagid honeyeater, nor even in the same part of the songbird radiation as meliphagids. Instead, the Hawaiian species are divergent members of a passeridan group that includes deceptively dissimilar families of songbirds (Holarctic waxwings, neotropical silky flycatchers, and palm chats). Here we designate them as a new family, the Mohoidae. A nuclear-DNA rate calibration suggests that mohoids diverged from their closest living ancestor 14-17 mya, coincident with the estimated earliest arrival in Hawaii of a bird-pollinated plant lineage. Convergent evolution, the evolution of similar traits in distantly related taxa because of common selective pressures, is illustrated well by nectar-feeding birds, but the morphological, behavioral, and ecological similarity of the mohoids to the Australasian honeyeaters makes them a particularly striking example of the phenomenon.

  7. Bird diversity and distribution in relation to urban landscape types in northern Rwanda.

    PubMed

    Gatesire, T; Nsabimana, D; Nyiramana, A; Seburanga, J L; Mirville, M O

    2014-01-01

    Using the point count method, linear mixed models, Shannon's diversity index, and Bray-Curtis cluster analysis, we conducted a study of the effect of urban fabric layout on bird diversity and distribution in northern Rwanda. The results showed a significant effect of city landscapes on bird richness and relative abundance; residential neighborhoods, institutional grounds, and informal settlements had the highest species diversity in comparison to other microlandscape types. Riversides were characterized by specialized bird species, commonly known to be restricted to wetland environments. Built-up areas and open field landscapes had comparable results. One Albertine Rift endemic bird species, the Ruwenzori Double-collared Sunbird (Cinnyris stuhlmanni), was recorded. Three migratory birds were found in Musanze city for the first time: the Common Sandpiper (Actitis hypoleucos), the Spotted Flycatcher (Muscicapa striata), and the Willow Warbler (Phylloscopus trochilus). Two bird species have not been previously reported in Rwanda: the Garden Warbler (Sylvia borin) and the Lesser Spotted Eagle (Aquila pomarina). The implications of this study are particularly relevant to urban decision makers who should consider the existence of a great diversity of avian fauna when developing and implementing master plans, especially when villages and cities are in proximity of protected areas or natural reserves.

  8. Large-scale geographical variation in eggshell metal and calcium content in a passerine bird (Ficedula hypoleuca).

    PubMed

    Ruuskanen, Suvi; Laaksonen, Toni; Morales, Judith; Moreno, Juan; Mateo, Rafael; Belskii, Eugen; Bushuev, Andrey; Järvinen, Antero; Kerimov, Anvar; Krams, Indrikis; Morosinotto, Chiara; Mänd, Raivo; Orell, Markku; Qvarnström, Anna; Slate, Fred; Tilgar, Vallo; Visser, Marcel E; Winkel, Wolfgang; Zang, Herwig; Eeva, Tapio

    2014-03-01

    Birds have been used as bioindicators of pollution, such as toxic metals. Levels of pollutants in eggs are especially interesting, as developing birds are more sensitive to detrimental effects of pollutants than adults. Only very few studies have monitored intraspecific, large-scale variation in metal pollution across a species' breeding range. We studied large-scale geographic variation in metal levels in the eggs of a small passerine, the pied flycatcher (Ficedula hypoleuca), sampled from 15 populations across Europe. We measured 10 eggshell elements (As, Cd, Cr, Cu, Ni, Pb, Zn, Se, Sr, and Ca) and several shell characteristics (mass, thickness, porosity, and color). We found significant variation among populations in eggshell metal levels for all metals except copper. Eggshell lead, zinc, and chromium levels decreased from central Europe to the north, in line with the gradient in pollution levels over Europe, thus suggesting that eggshell can be used as an indicator of pollution levels. Eggshell lead levels were also correlated with soil lead levels and pH. Most of the metals were not correlated with eggshell characteristics, with the exception of shell mass, or with breeding success, which may suggest that birds can cope well with the current background exposure levels across Europe.

  9. Bird Diversity and Distribution in relation to Urban Landscape Types in Northern Rwanda

    PubMed Central

    Gatesire, T.; Nsabimana, D.; Nyiramana, A.; Seburanga, J. L.; Mirville, M. O.

    2014-01-01

    Using the point count method, linear mixed models, Shannon's diversity index, and Bray-Curtis cluster analysis, we conducted a study of the effect of urban fabric layout on bird diversity and distribution in northern Rwanda. The results showed a significant effect of city landscapes on bird richness and relative abundance; residential neighborhoods, institutional grounds, and informal settlements had the highest species diversity in comparison to other microlandscape types. Riversides were characterized by specialized bird species, commonly known to be restricted to wetland environments. Built-up areas and open field landscapes had comparable results. One Albertine Rift endemic bird species, the Ruwenzori Double-collared Sunbird (Cinnyris stuhlmanni), was recorded. Three migratory birds were found in Musanze city for the first time: the Common Sandpiper (Actitis hypoleucos), the Spotted Flycatcher (Muscicapa striata), and the Willow Warbler (Phylloscopus trochilus). Two bird species have not been previously reported in Rwanda: the Garden Warbler (Sylvia borin) and the Lesser Spotted Eagle (Aquila pomarina). The implications of this study are particularly relevant to urban decision makers who should consider the existence of a great diversity of avian fauna when developing and implementing master plans, especially when villages and cities are in proximity of protected areas or natural reserves. PMID:25133203

  10. The eye of the field geologist and the mind of the tectonician: one view of dynamic crustal rheology in convergent orogens

    NASA Astrophysics Data System (ADS)

    Brown, M.

    2004-05-01

    -depleted rocks, implying that they were quasi-continuously drained. Studies of migmatitic granulites demonstrate that melt migrates from grain boundaries to mesoscale networks of structures (mm to m) to steeply-inclined conduits recorded by rod or tabular granite intrusions (m to dm). Melt loss from lower crust yields residual rocks composed of strong minerals (feldspar, pyroxene and garnet) with only minor melt on grain boundaries. Thus, weakening of lower crust due to melting is followed by its strengthening. Around the brittle-to-viscous transition zone granite accumulates in subhorizontal tabular plutons, which implies transient presence of significantly weaker layers in shallow orogenic crust; these are potential detachment horizons. Field studies of exhumed orogens suggest deformation commonly is laterally, transversely and vertically diachronous, reflecting the spatial and temporal variation in the weakening-to-strengthening cycle. There may be important sub-horizontal movement horizons, which allow (partial) decoupling of crustal layers. At upper-to-middle crustal levels rocks are metamorphosed in greenschist-amphibolite facies, with local enhancement by pluton-advected heat to amphibolite-granulite facies and thrust-style brittle-ductile deformation (e.g., Acadian, NH). Rocks from middle crustal levels are in amphibolite facies and have penetrative steep fabrics (e.g., Acadian, western ME) or exhibit a complex network of shallowly- and steeply-dipping fabrics (e.g., St. Malo, France). Rocks from lower crustal levels are in amphibolite-granulite facies and have shallow fabrics due to crustal flow, although these may be steepened by later deformation including core-complex formation (e.g., S. Brittany, France).

  11. Nitrous Oxide Emissions From Northern Forested and Harvested Ecosystems

    NASA Astrophysics Data System (ADS)

    Kavanaugh, K. M.; Kellman, L. M.

    2005-12-01

    Very little is known about how deforestation alters the soil subsurface production and surface emissions of N2O from northern forest soils. Soil N2O surface fluxes and subsurface concentrations from two 3 year old harvested and intact forest pairs of contrasting soil texture were monitored during the 2004 and 2005 growing seasons in the Acadian forest of Atlantic Canada in order to: 1) quantify N2O emissions associated with each land-use type, 2) examine spatial and temporal variations in subsurface concentrations and surface fluxes at each site, and 3) determine the suitability of a photoacoustic gas monitor (PGM) for in- situ field measurements vs. field sample collection and laboratory analysis on a gas chromatograph. Each site was instrumented with 11 permanent collars for surface flux measurements designed to capture the microsite variability at the sites. Subsurface soil gas samplers, designed to identify the important zones of N2O production in the vertical profile were installed at depths of 0, 10, 20 and 35 cm below the organic-mineral soil interface. Surface fluxes were measured with non-steady-state vented surface flux chambers with measurements of all surface flux and subsurface data made on a bi-weekly basis. Results suggest that spatial and temporal variability in surface emissions are very high and routinely close to zero. Subsurface profile concentration data shows vertical concentration profiles at intact forest sites with concentrations close to atmospheric, while harvested sites show a pattern of increasing N2O concentration with depth, reaching a maximum of approximately 27000ppb at 35cm.

  12. Strike-slip faulting at Thebes Gap, Missouri and Illinois; implications for New Madrid tectonism

    USGS Publications Warehouse

    Harrison, Richard W.; Schultz, Art

    1994-01-01

    Numerous NNE and NE striking strike-slip faults and associated normal faults, folds, and transtensional grabens occur in the Thebes Gap area of Missouri and Illinois. These structures developed along the northwestern margin of the buried Reelfoot rift of Precambrian-Cambrian age at the northern edge of the Mississippi embayment. They have had a long-lived and complex structural history. This is an area of recent moderate seismicity, approximately 45 km north of the New Madrid seismic zone. Stratigraphic evidence suggests that these faults were active during the Middle Ordovician. They were subsequently reactivated between the Early Devonian and Late Cretaceous, probably in response to both the Acadian and Ouachita orogenies. Deformation during this period was characterized by strongly faulted and folded Ordovician through Devonian rocks. In places, these deformed rocks are overlain with angular unconformity by undeformed Cretaceous strata. Fault motion is interpreted as dominantly strike slip. A still younger period of reactivation involved Late Cretaceous and Cenozoic formations as young as the Miocene or Pliocene Mounds Gravel. These formations have experienced both minor high-angle normal faulting and subsequent major, right-lateral strike-slip faulting. En echelon north-south folds, ENE striking normal faults, regional fracture patterns, and drag folds indicate the right-lateral motion for this major episode of faulting which predates deposition of Quaternary loess. Several nondefinitive lines of evidence suggest Quaternary faulting. Similar fault orientations and kinematics, as well as recent seismicity and proximity, clearly suggest a structural relationship between deformation at Thebes Gap and tectonism associated with the New Madrid area.

  13. Supra-subduction zone extensional magmatism in Vermont and adjacent Quebec: Implications for early Paleozoic Appalachian tectonics

    USGS Publications Warehouse

    Kim, J.; Coish, R.; Evans, M.; Dick, G.

    2003-01-01

    Metadiabasic intrusions of the Mount Norris Intrusive Suite occur in fault-bounded lithotectonic packages containing Stowe, Moretown, and Cram Hill Formation lithologies in the northern Vermont Rowe-Hawley belt, a proposed Ordovician arc-trench gap above an east-dipping subduction zone. Rocks of the Mount Norris Intrusive Suite are characteristically massive and weakly foliated, have chilled margins, contain xenoliths, and have sharp contacts that both crosscut and are parallel to early structural fabrics in the host metasedimentary rocks. Although the mineral assemblage of the Mount Norris Intrusive Suite is albite + actinolite + epidote + chlorite + calcite + quartz, intergrowths of albite + actinolite are probably pseudomorphs after plagioclase + clinopyroxene. The metadiabases are subalkaline, tholeiitic, hypabyssal basalts with preserved ophitic texture. A backarc-basin tectonic setting for the intrusive suite is suggested by its LREE (light rare earth element) enrichment, negative Nb-Ta anomalies, and Ta/Yb vs. Th/Yb trends. Although no direct isotopic age data are available, the intrusions are broadly Ordovician because their contacts are clearly folded by the earliest Acadian (Silurian-Devonian) folds. Field evidence and geochemical data suggest compelling along-strike correlations with the Coburn Hill Volcanics of northern Vermont and the Bolton Igneous Group of southern Quebec. Isotopic and stratigraphic age constraints for the Bolton Igneous Group bracket these backarc magmas to the 477-458 Ma interval. A tectonic model that begins with east-dipping subduction and progresses to outboard west-dipping subduction after a syncollisional polarity reversal best explains the intrusion of deformed metamorphosed metasedimentary rocks by backarc magmas.

  14. Brown seaweed- (Tasco) treated conserved forage enhances antioxidant status and immune function in heat-stressed wether lambs.

    PubMed

    Saker, K E; Fike, J H; Veit, H; Ward, D L

    2004-04-01

    Twenty-seven wether lambs were utilized to evaluate select innate immunity and oxidative stress in response to diet and heat stress. Dietary treatments were: (i) control (tall fescue) hay = no Tasco (tradename for the extract of the brown seaweed, Ascophyllum nodosum, Acadian Sealants Ltd, Nova Scotia, Canada); (ii) pre-harvest Tasco-Forage-treated hay and (iii) control hay + post-harvest Tasco-EX. Tasco-Forage and Tasco-EX are two forms of the Tasco extract that are either applied to foliage or used for direct feeding, respectively. All lambs were supplemented with soyabean meal and trace mineralized salt. Heat stress was applied for 10 days with measurements obtained at days 0, 4 and 10. A heat x treatment interaction indicated hay with Tasco enhanced monocyte oxidative burst through short duration (p < 0.05) and long duration (p < 0.10) heat stress. Phagocytic activity was influenced by days of heat stress (p < 0.001) and treatment (p = 0.02) with post-harvest Tasco lambs exhibiting the greatest immune enhancement (p < 0.05). Red and white blood cell glutathione peroxidase increased by heat stress day 10 in Tasco lambs. Superoxide dismutase activity was increased and lipid hydroperoxide metabolites minimized (p < 0.01) through long duration heat stress in the pre-harvest Tasco group. Tasco treatment of tall fescue hay prior to harvest appears to provide residual effects on animal antioxidant availability in short-duration heat stress. Tasco supplementation to post-harvest fescue hay enhances immune function and protects against prolonged heat-induced oxidative stress.

  15. Lessons learned from implementing a national infrastructure in Sweden for storage and analysis of next-generation sequencing data

    PubMed Central

    2013-01-01

    Analyzing and storing data and results from next-generation sequencing (NGS) experiments is a challenging task, hampered by ever-increasing data volumes and frequent updates of analysis methods and tools. Storage and computation have grown beyond the capacity of personal computers and there is a need for suitable e-infrastructures for processing. Here we describe UPPNEX, an implementation of such an infrastructure, tailored to the needs of data storage and analysis of NGS data in Sweden serving various labs and multiple instruments from the major sequencing technology platforms. UPPNEX comprises resources for high-performance computing, large-scale and high-availability storage, an extensive bioinformatics software suite, up-to-date reference genomes and annotations, a support function with system and application experts as well as a web portal and support ticket system. UPPNEX applications are numerous and diverse, and include whole genome-, de novo- and exome sequencing, targeted resequencing, SNP discovery, RNASeq, and methylation analysis. There are over 300 projects that utilize UPPNEX and include large undertakings such as the sequencing of the flycatcher and Norwegian spruce. We describe the strategic decisions made when investing in hardware, setting up maintenance and support, allocating resources, and illustrate major challenges such as managing data growth. We conclude with summarizing our experiences and observations with UPPNEX to date, providing insights into the successful and less successful decisions made. PMID:23800020

  16. Understory avifauna exhibits altered mobbing behavior in tropical forest degraded by selective logging.

    PubMed

    Hua, Fangyuan; Sieving, Kathryn E

    2016-11-01

    In understanding the impacts of selective logging on biodiversity, relatively little is known about the critical behavioral link between altered forest conditions and population persistence. Predator-mobbing is a widespread anti-predator behavior in birds that expresses a well-known trade-off influencing prey survival under predation risk. Here, we ask whether the predator-mobbing behavior of understory forest birds is altered by selective logging and associated forest structural changes in the highly endangered lowland rainforest of Sumatra. At four study sites spanning a gradient of logging-induced forest degradation, we used standardized mobbing and owl call playbacks with predator model presentation to elicit the predator-mobbing behavior of understory prey birds, compared birds' mobbing intensity across sites, and related variation in this intensity to forest vegetation structure. We found that selective logging altered birds' predator-mobbing intensity (measured by behavioral conspicuousness and propensity to approach the predator) as well as forest structure, and that vegetative changes to canopy and understory were correlated with contrasting responses by the two major bird foraging guilds, gleaning versus flycatching birds. We additionally discuss the implications of our findings for further hypothesis testing pertaining to the impacts of selective logging on the ecological processes underlying prey mobbing behavior, particularly with regards to predator-prey interactions and prey accruement of energy reserves.

  17. Explaining stasis: microevolutionary studies in natural populations.

    PubMed

    Merilä, J; Sheldon, B C; Kruuk, L E

    2001-01-01

    Microevolution, defined as a change in the genetic constitution of a population over time, is considered to be of commonplace occurrence in nature. Its ubiquity can be inferred from the observation that quantitative genetic divergence among populations usually exceeds that to be expected due to genetic drift alone, and from numerous observations and experiments consistent with local adaptation. Experimental manipulations in natural populations have provided evidence that rapid evolutionary responses may occur in the wild. However, there are remarkably few cases where direct observations of natural populations have revealed microevolutionary changes occurring, despite the frequent demonstration of additive genetic variation and strong directional selection for particular traits. Those few cases where responses congruent with expectation have been demonstrated are restricted to changes over one generation. In this article we focus on possible explanations as to why heritable traits under apparently strong directional selection often fail to show the expected evolutionary response. To date, few of these explanations for apparent stasis have been amenable to empirical testing. We describe new methods, derived from procedures developed by animal breeding scientists, which can be used to address these explanations, and illustrate the approach with examples from long-term studies of collared flycatchers (Ficedula albicollis) and red deer (Cervus elaphus). Understanding why most intensively studied natural populations do not appear to be evolving is an important challenge for evolutionary biology.

  18. Two new species of Quasithelazia Maplestone, 1932 (Nematoda: Acuariidae) from Malaysia, with an amended diagnosis and review of the genus.

    PubMed

    Mutafchiev, Yasen; Mariaux, Jean; Georgiev, Boyko B

    2014-06-01

    Quazithelazia rostrata n. sp. from Ceyx erithaca (L.) (type-host) and Alcedo euryzona Temminck (Coraciiformes, Alcedinidae) and Q. alata n. sp. from Enicurus ruficapillus Temminck (Passeriformes, Muscicapidae) are described from vicinities of Gombak Biological Station, Selangor, Malaysia; both species are parasitic under the koilin lining of the gizzard. Paratypes of Schistogendra pelargopsis Nandi, De & Majumdar, 1985, a parasite of Pelargopsis capensis (L.) (Alcedinidae) from India, are redescribed and the species is recognised as a junior synonym of the type-species of Quasithelazia, Q. tenuis Maplestone, 1932 (new synonymy), a species originally described from Halcyon smyrnensis (L.) (Alcedinidae) in India. An amended diagnosis of the genus Quasithelazia Maplestone, 1932 is proposed. Currently, this genus includes eight species occurring in the Old World, six of them parasitic in kingfishers (Alcedinidae) and two species parasitic in flycatchers (Muscicapidae). These include, inter alia, Q. halcyoni n. comb. for Viktorocara halcyoni Ryzhikov & Khokhlova, 1964 from Halcyon smyrnensis and H. pileata (Boddaert) in Vietnam and the Russian Far East, Q. microcordonis n. comb. for Rusguniella microcordonis Schmidt & Kuntz, 1971 from Halcyon coromanda major (Temminck & Schlegel) in Taiwan and Q. multipapillata n. comb. for Schistogendra multipapillata Zhang, 1993 from Tarsiger cyanurus (Pallas) (Muscicapidae) in China. Comparative morphological data for Quasithelazia spp. are presented. Schistogendra oligopapillata Zhang & An, 2002 from domestic ducks in China is considered a species incertae sedis.

  19. What prevents phenological adjustment to climate change in migrant bird species? Evidence against the ``arrival constraint'' hypothesis

    NASA Astrophysics Data System (ADS)

    Goodenough, Anne E.; Hart, Adam G.; Elliot, Simon L.

    2011-01-01

    Phenological studies have demonstrated changes in the timing of seasonal events across multiple taxonomic groups as the climate warms. Some northern European migrant bird populations, however, show little or no significant change in breeding phenology, resulting in synchrony with key food sources becoming mismatched. This phenological inertia has often been ascribed to migration constraints (i.e. arrival date at breeding grounds preventing earlier laying). This has been based primarily on research in The Netherlands and Germany where time between arrival and breeding is short (often as few as 9 days). Here, we test the arrival constraint hypothesis over a 15-year period for a U.K. pied flycatcher ( Ficedula hypoleuca) population where laying date is not constrained by arrival as the period between arrival and breeding is substantial and consistent (average 27 ± 4.57 days SD). Despite increasing spring temperatures and quantifiably stronger selection for early laying on the basis of number of offspring to fledge, we found no significant change in breeding phenology, in contrast with co-occurring resident blue tits ( Cyanistes caeruleus). We discuss possible non-migratory constraints on phenological adjustment, including limitations on plasticity, genetic constraints and competition, as well as the possibility of counter-selection pressures relating to adult survival, longevity or future reproductive success. We propose that such factors need to be considered in conjunction with the arrival constraint hypothesis.

  20. Host selection by the shiny cowbird

    USGS Publications Warehouse

    Wiley, J.W.

    1988-01-01

    Factors important in Shiny Cowbird (Molothrus bonariensis) host selection were examined within the mangrove community in Puerto Rico. Cowbirds did not parasitize birds in proportion to their abundance. The cowbird breeding season coincided with those of its major hosts, which were 'high-quality' foster species (i.e., species that fledge .gtoreq. 55% of cowbirds hatched: Yellow Warbler, Dendroica petechia; Yellow-shouldered Blackbird, Agelaius xanthomus; Black-whiskered Vireo, Vireo altiloquus; Black-cowled Oriole, Icterus dominicensis; Peurto Rican Flycatcher, Myiarchus antillarum; Troupial, Icterus icterus), and did not extend into other periods even though nests of 'low-quality: species (i.e., species that fledge < 55% of cowbird chicks that hatched: Bronze Mannikin, Lonchura cucullata; Greater Antillean Grackle, Quiscalus niger; Gray Kingbird, Tyrannus dominicensis; Northern Mockingbird, Mimus polyglottos; Red-legged Thrush, Turdus plumbeus) were available. Shiny Cowbird food habits and egg size were similar to those of their hosts, suggesting that cowbirds choose hosts partly on the basis of this combination. Cowbirds located host nests primarily by cryptically watching activities of birds in likely habitats. Other nest locating strategies were active searching of suitable habitat and 'flushing' of hosts by the cowbird's noisy approach. Cowbirds closely monitored nest status with frequent visits that peaked on the host's first day of egg laying. Hosts using covered nests (e.g., cavities, domed nests) were as vulnerable to cowbird parasitism as those building open nests.

  1. What prevents phenological adjustment to climate change in migrant bird species? Evidence against the "arrival constraint" hypothesis.

    PubMed

    Goodenough, Anne E; Hart, Adam G; Elliot, Simon L

    2011-01-01

    Phenological studies have demonstrated changes in the timing of seasonal events across multiple taxonomic groups as the climate warms. Some northern European migrant bird populations, however, show little or no significant change in breeding phenology, resulting in synchrony with key food sources becoming mismatched. This phenological inertia has often been ascribed to migration constraints (i.e. arrival date at breeding grounds preventing earlier laying). This has been based primarily on research in The Netherlands and Germany where time between arrival and breeding is short (often as few as 9 days). Here, we test the arrival constraint hypothesis over a 15-year period for a U.K. pied flycatcher (Ficedula hypoleuca) population where laying date is not constrained by arrival as the period between arrival and breeding is substantial and consistent (average 27 ± 4.57 days SD). Despite increasing spring temperatures and quantifiably stronger selection for early laying on the basis of number of offspring to fledge, we found no significant change in breeding phenology, in contrast with co-occurring resident blue tits (Cyanistes caeruleus). We discuss possible non-migratory constraints on phenological adjustment, including limitations on plasticity, genetic constraints and competition, as well as the possibility of counter-selection pressures relating to adult survival, longevity or future reproductive success. We propose that such factors need to be considered in conjunction with the arrival constraint hypothesis.

  2. A survey of current breeding habitats: Chapter 5 in Status, ecology and conservation of the Southwestern Willow Flycatcher (RMRS-GTR-60)

    USGS Publications Warehouse

    Sogge, Mark K.; Marshall, Robert M.; Finch, Deborah M.; Stoleson, Scott H.

    2000-01-01

    The distribution and abundance of a species across a landscape depends, in part, on the distribution and abundance of appropriate habitat. If basic resource needs such as food, water, and cover are not present, then that species is excluded from the area. Scarcity of appropriate habitat is generally the key reason for the status of most rare and endangered species. An understanding of an endangered species’ habitat characteristics is crucial to effective management, conservation and recovery.

  3. Reported middle Paleozoic fossils and new geochronological data from the southern and central Appalachians: Disposable outrageous hypothesis or justification for major revision of tectonic history

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Hatcher, R.D. Jr.

    Recently published interpretations of fossil fragments from the Walden Creek Group (Ocoee Supergroup) suggesting that these rocks are middle Paleozoic (Devonian to Early Carboniferous), and new geochronological data that yield late Paleozoic age dates on rocks and major faults in the Blue Ridge and piedmont, if taken alone, would permit speculation that most of the deformation and metamorphism affecting this part of the orogen is Alleghanian. The two Ordovician clastic wedges (Sevier, Llanvirn, and Martinsburg, Caradoc-Ashgill) and the Carboniferous-Permian wedge(s), along with many radiometric ages on plutons, indicate uplift and sediment dispersal from the interior of the southern and centralmore » Appalachians (SCA) that may have resulted from Taconian and Alleghanian deformation. Combining the reproducible fossil evidence, including that from Alabama and a recently discovered crinoid fragment from the upper part of the Murphy belt sequence, with the most current geochronological data requires that peak metamorphism and penetrative deformation be at least Devonian or younger at the southwestern end of the orogen, and Late Ordovician or younger in the Carolinas and northern Georgia. Zircon ages reported from large thrust and dextral strike-slip faults bounding the Pine Mountain window indicate all of the faults there may be Alleghanian, except the younger sinistral Mesozoic faults, and requires that both metamorphism and penetrative deformation there also be Alleghanian. As in New England, the southern Appalachian Alleghanian metamorphic core is now known to be much more extensive. The older data require that the Taconian and perhaps the Acadian orogenies were significant events in the SCA, but these new data reconfirm the dominance of Alleghanian continent-continent collision processes here.« less

  4. Infiltration-driven metamorphism, New England, USA: Regional CO2 fluxes and implications for Devonian climate and extinctions

    NASA Astrophysics Data System (ADS)

    Stewart, E. M.; Ague, Jay J.

    2018-05-01

    We undertake thermodynamic pseudosection modeling of metacarbonate rocks in the Wepawaug Schist, Connecticut, USA, and examine the implications for CO2 outgassing from collisional orogenic belts. Two broad types of pseudosections are calculated: (1) a fully closed-system model with no fluid infiltration and (2) a fluid-buffered model including an H2O-CO2 fluid of a fixed composition. This fluid-buffered model is used to approximate a system open to infiltration by a water-bearing fluid. In all cases the fully closed-system model fails to reproduce the observed major mineral zones, mineral compositions, reaction temperatures, and fluid compositions. The fluid-infiltrated models, on the other hand, successfully reproduce these observations when the XCO2 of the fluid is in the range ∼0.05 to ∼0.15. Fluid-infiltrated models predict significant progressive CO2 loss, peaking at ∼50% decarbonation at amphibolite facies. The closed-system models dramatically underestimate the degree of decarbonation, predicting only ∼15% CO2 loss at peak conditions, and, remarkably, <1% CO2 loss below ∼600 °C. We propagate the results of fluid-infiltrated pseudosections to determine an areal CO2 flux for the Wepawaug Schist. This yields ∼1012 mol CO2 km-2 Myr-1, consistent with multiple independent estimates of the metamorphic CO2 flux, and comparable in magnitude to fluxes from mid-ocean ridges and volcanic arcs. Extrapolating to the area of the Acadian orogenic belt, we suggest that metamorphic CO2 degassing is a plausible driver of global warming, sea level rise, and, perhaps, extinction in the mid- to late-Devonian.

  5. The carpenter fork bed, a new - and older - Black-shale unit at the base of the New Albany shale in central Kentucky: Characterization and significance

    USGS Publications Warehouse

    Barnett, S.F.; Ettensohn, F.R.; Norby, R.D.

    1996-01-01

    Black shales previously interpreted to be Late Devonian cave-fill or slide deposits are shown to be much older Middle Devonian black shales only preserved locally in Middle Devonian grabens and structural lows in central Kentucky. This newly recognized - and older -black-shale unit occurs at the base of the New Albany Shale and is named the Carpenter Fork Bed of the Portwood Member of the New Albany Shale after its only known exposure on Carpenter Fork in Boyle County, central Kentucky; two other occurrences are known from core holes in east-central Kentucky. Based on stratigraphic position and conodont biostratigraphy, the unit is Middle Devonian (Givetian: probably Middle to Upper P. varcus Zone) in age and occurs at a position represented by an unconformity atop the Middle Devonian Boyle Dolostone and its equivalents elsewhere on the outcrop belt. Based on its presence as isolated clasts in the overlying Duffin Bed of the Portwood Member, the former distribution of the unit was probably much more widespread - perhaps occurring throughout western parts of the Rome trough. Carpenter Fork black shales apparently represent an episode of subsidence or sea-level rise coincident with inception of the third tectophase of the Acadian orogeny. Deposition, however, was soon interrupted by reactivation of several fault zones in central Kentucky, perhaps in response to bulge migration accompanying start of the tectophase. As a result, much of central Kentucky was uplifted and tilted, and the Carpenter Fork Bed was largely eroded from the top of the Boyle, except in a few structural lows like the Carpenter Fork graben where a nearly complete record of Middle to early Late Devonian deposition is preserved.

  6. The importance of floodplain forests in the conservation and management of neotropical migratory birds in the Midwest

    USGS Publications Warehouse

    Knutson, M.G.; Hoover, J.P.; Klaas, E.E.; Thompson, Frank R.

    1996-01-01

    Bottomland forests of the Central Forest Region of the Upper Midwest are found primarily on the floodplains of large rivers and include at least six types of forest communities. Birds breeding in bottomland forests are affected by extensive variation in latitude, climate, hydrology, forest succession, and change caused by anthropogenic disturbances. The floodplain forest bird community differs in species composition and in relative abundance from adjacent upland habitats. High abundances of some species are found in the floodplain and some species, such as the prothonotary warbler, brown creeper, yellow-billed cuckoo, yellow-bellied sapsucker, and great crested flycatcher, show a clear preference for floodplain forests. Studies of nesting success indicate that, for some species, nest success may be higher in the floodplain than in the uplands. Floodplain birds face threats due to large-scale loss of floodplain forest habitat. Conservation efforts should focus on restoring degraded floodplains by maintaining high tree species diversity and wide corridors. To accomplish this, the underlying hydrodynamics which support a diverse floodplain forest habitat may need to be restored. Large, contiguous tracts of floodplain and upland forests should be maintained where they exist and restored in other locations. This will provide some high quality habitat for area-sensitive neotropical migratory birds (NTMBs) in agricultural landscapes where small, scattered forest fragments are the rule. Future research efforts should examine the importance of floodplain forests in maintaining populations of neotropical migrants, especially birds experiencing population declines in adjacent uplands.

  7. Healing with animals in the Levant from the 10th to the 18th century

    PubMed Central

    Lev, Efraim

    2006-01-01

    Animals and products derived from different organs of their bodies have constituted part of the inventory of medicinal substances used in various cultures since ancient times. The article reviews the history of healing with animals in the Levant (The Land of Israel and parts of present-day Syria, Lebanon, and Jordan, defined by the Muslims in the Middle Ages as Bilad al-Sham) in the medieval and early Ottoman periods. Intensive research into the phenomenon of zootherapy in the medieval and early Ottoman Levant has yielded forty-eight substances of animal origin that were used medicinally. The vast majority of these substances were local and relatively easy to obtain. Most of the substances were domestic (honey, wax, silkworm, etc.), others were part of the local wildlife (adder, cuttle fish, flycatcher, firefly, frog, triton, scorpion, etc.), part of the usual medieval household (milk, egg, cheese, lamb, etc.), or parasites (louse, mouse, stinkbug, etc.). Fewer substances were not local but exotic, and therefore rare and expensive (beaver testicles, musk oil, coral, ambergris, etc.). The range of symptoms that the substances of animal origin were used to treat was extensive and included most of the known diseases and maladies of that era: mainly hemorrhoids, burns, impotence, wounds, and skin, eye, and stomach diseases. Changes in the moral outlook of modern societies caused the use of several substances of animal origin to cease in the course of history. These include mummy, silkworm, stinkbug, scarabees, snail, scorpion, and triton. PMID:16504024

  8. Healing with animals in the Levant from the 10th to the 18th century.

    PubMed

    Lev, Efraim

    2006-02-21

    Animals and products derived from different organs of their bodies have constituted part of the inventory of medicinal substances used in various cultures since ancient times. The article reviews the history of healing with animals in the Levant (The Land of Israel and parts of present-day Syria, Lebanon, and Jordan, defined by the Muslims in the Middle Ages as Bilad al-Sham) in the medieval and early Ottoman periods. Intensive research into the phenomenon of zootherapy in the medieval and early Ottoman Levant has yielded forty-eight substances of animal origin that were used medicinally. The vast majority of these substances were local and relatively easy to obtain. Most of the substances were domestic (honey, wax, silkworm, etc.), others were part of the local wildlife (adder, cuttle fish, flycatcher, firefly, frog, triton, scorpion, etc.), part of the usual medieval household (milk, egg, cheese, lamb, etc.), or parasites (louse, mouse, stinkbug, etc.). Fewer substances were not local but exotic, and therefore rare and expensive (beaver testicles, musk oil, coral, ambergris, etc.). The range of symptoms that the substances of animal origin were used to treat was extensive and included most of the known diseases and maladies of that era: mainly hemorrhoids, burns, impotence, wounds, and skin, eye, and stomach diseases. Changes in the moral outlook of modern societies caused the use of several substances of animal origin to cease in the course of history. These include mummy, silkworm, stinkbug, scarabees, snail, scorpion, and triton.

  9. Summer inventory of landbirds in Kenai Fjords National Park

    USGS Publications Warehouse

    2006-01-01

    As part of the National Park Service Inventory and Monitoring Program, we conducted a summer inventory of landbirds within Kenai Fjords National Park. Using a stratified random sampling design of areas accessible by boat or on foot, we selected sites that encompassed the breadth of habitat types within the Park. We detected 101 species across 52 transects, including 62 species of landbirds, which confirmed presence of 87% of landbird species expected to occur in the Park during the summer breeding season. We found evidence of breeding for three Partners in Flight Watch List species, Rufous Hummingbird (Selasphorus rufus), Olive-sided Flycatcher (Contopus cooperi), and Rusty Blackbird (Euphagus carolinus), which are of particular conservation concern due to recent population declines. Kenai Fjords National Park supports extremely high densities of Hermit Thrush, Orange-crowned Warbler, and Wilson’s Warbler (Wilsonia pusilla) compared with other regions of Alaska. Other commonly observed species included Fox Sparrow (Passerella iliaca), Varied Thrush (Ixoreus naevius), Rubycrowned Kinglet (Regulus calendula), and Yellow Warbler (Dendroica petechia). More than half of the landbird species we observed occurred in needleleaf forests, and several of these species were strongly associated with the coastforest interface. Tall shrub habitats, which occurred across all elevations and in recently deglaciated areas, supported high densities and a diverse array of passerines. Two major riparian corridors, with their broadleaf forests, wetlands, and connectivity to interior Alaska, provided unique and important landbird habitats within the region.

  10. Effects of protective fencing on birds, lizards, and black-tailed hares in the Western Mojave Desert

    USGS Publications Warehouse

    Brooks, M.

    1999-01-01

    Effects of protective fencing on birds, lizards, black-tailed hares (Lepus californicus), perennial plant cover, and structural diversity of perennial plants were evaluated from spring 1994 through winter 1995 at the Desert Tortoise Research Natural Area (DTNA), in the Mojave Desert, California. Abundance and species richness of birds were higher inside than outside the DTNA, and effects were larger during breeding than wintering seasons and during a high than a low rainfall year. Ash-throated flycatchers (Myiarchus cinerascens), cactus wrens (Campylorhynchus brunneicapillus), LeConte's thrashers (Toxostoma lecontei), loggerhead shrikes (Lanius ludovicianus), sage sparrows (Amphispiza belli), and verdins (Auriparus flaviceps) were more abundant inside than outside the DTNA. Nesting activity was also more frequent inside. Total abundance and species richness of lizards and individual abundances of western whiptail lizards (Cnemidophorous tigris) and desert spiny lizards (Sceloporus magister) were higher inside than outside. In contrast, abundance of black-tailed hares was lower inside. Structural diversity of the perennial plant community did not differ due to protection, but cover was 50% higher in protected areas. Black-tailed hares generally prefer areas of low perennial plant cover, which may explain why they were more abundant outside than inside the DTNA. Habitat structure may not affect bird and lizard communities as much as availability of food at this desert site, and the greater abundance and species richness of vertebrates inside than outside the DTNA may correlate with abundances of seeds and invertebrate prey.

  11. Sound settlement: noise surpasses land cover in explaining breeding habitat selection of secondary cavity-nesting birds.

    PubMed

    Kleist, Nathan J; Guralnick, Robert P; Cruz, Alexander; Francis, Clinton D

    2017-01-01

    Birds breeding in heterogeneous landscapes select nest sites by cueing in on a variety of factors from landscape features and social information to the presence of natural enemies. We focus on determining the relative impact of anthropogenic noise on nest site occupancy, compared to amount of forest cover, which is known to strongly influence the selection process. We examine chronic, industrial noise from natural gas wells directly measured at the nest box as well as site-averaged noise, using a well-established field experimental system in northwestern New Mexico. We hypothesized that high levels of noise, both at the nest site and in the environment, would decrease nest box occupancy. We set up nest boxes using a geospatially paired control and experimental site design and analyzed four years of occupancy data from four secondary cavity-nesting birds common to the Colorado Plateau. We found different effects of noise and landscape features depending on species, with strong effects of noise observed in breeding habitat selection of Myiarchus cinerascens, the Ash-throated Flycatcher, and Sialia currucoides, the Mountain Bluebird. In contrast, the amount of forest cover less frequently explained habitat selection for those species or had a smaller standardized effect than the acoustic environment. Although forest cover characterization and management is commonly employed by natural resource managers, our results show that characterizing and managing the acoustic environment should be an important tool in protected area management. © 2016 by the Ecological Society of America.

  12. Birds reveal their personality when singing.

    PubMed

    Garamszegi, László Zsolt; Eens, Marcel; Török, János

    2008-07-09

    Individual differences in social behaviour may have consequences for mate choice and sexual signalling, because partners should develop preferences for personalities that maximize reproductive output. Here we propose that behavioural traits involved in sexual advertisement may serve as good indicators of personality, which is fundamental for sexual selection to operate on temperament. Bird song has a prominent and well-established role in sexual selection, and it displays considerable variation among individuals with a potentially strong personality component. Therefore, we predicted that features of song would correlate with estimates of personality. In a field study of free-living male collared flycatchers, Ficedula albicollis, we characterised personality based on the exploration of an altered breeding environment, and based on the risk taken when a potential predator was approaching during a simulated territorial interaction. We found that explorative and risk-taker individuals consistently sang at lower song posts than shy individuals in the presence of a human observer. Moreover, males from lower posts established pair-bonds relatively faster than males from higher posts. Our results may demonstrate that risk taking during singing correlates with risk taking during aggression and with exploration, thus personality may be manifested in different contexts involving sexual advertisement. These findings are in accordance with the hypothesis that the male's balance between investment in reproduction and risk taking is reflected in sexual displays, and it may be important information for choosy females that seek partners with personality traits enhancing breeding success.

  13. Tectonic overprint on magnetic fabric of the Ordovician Thetford Mines Ophiolite (Canada)

    NASA Astrophysics Data System (ADS)

    Di Chiara, Anita; Morris, Antony; Anderson, Mark W.; Menegon, Luca

    2017-04-01

    Studies in modern oceanic settings suggest locally along low-spreading ridges both lower crust and upper mantle peridotites may be exhumed to the seafloor in features known as oceanic core complexes (OCC). Examples of OCC on geological record can be preserved in ophiolites, relict of oceanic crust obducted onto continental margins, as for example the Jurassic Mirdita Ophiolite (Albania), suggesting that this spreading mode was active in the past. In order to understand such dynamics further, we investigated the OCC preserved in the Thetford Mines Ophiolite (TMO). TMO is part of the Southern Quebec ophiolites in the Canadian Appalachians (Quebec region), divided into three lithotectonic assemblages: The Humber Zone, a remnant of the Laurentian continental margin; The Cambrian-Ordovician Dunnage Zone, a remnant of the Iapetus Ocean and including the TMO and other ophiolites; and Silurian-Devonian Gaspé Belt, the sedimentary cover sequence. These were subjected to polyphase deformation, experiencing two Paleozoic orogenies: The Ordovician Taconian Orogeny (the Humber and Dunnage zones were amalgamated) and the Devonian Acadian orogeny which deformed and metamorphosed both the Dunnage Zone and the overlying Gaspe Belt. Here we present results from 12 paleomagnetic sites sampled on Humber zone on pillow lavas, dykes, layered gabbros and serpentinized dunites. Our results from AMS experiments show that these rocks, formed by fundamentally different magmatic processes, share a common magnetic fabric, with a kmin axis NW-SE orientated and the kmax steeply plunging to the NE. Additional processing of acquired BSE images and chemical mapping analyses at the SEM show that the kmax of the magnetic fabric is parallel to the elongation of magnetic particles (Iron rich minerals). This remarkably consistent fabric has a tectonic origin and is consistent with shortening perpendicular to the regional trend of fold axes.

  14. Structural and Hydrologic Implications of Joint Orientations in the Warner Creek and Stony Clove Drainage Basins, Catskill Mountains, Eastern New York

    NASA Astrophysics Data System (ADS)

    Haskins, M. N.; Vollmer, F. W.; Rayburn, J. A.; Gurdak, J. J.

    2010-12-01

    To investigate joint control on hydrology as well as tectonic implications, we conducted a study of joint orientations near the Stony Clove and Warner Creek drainages of the Catskill Mountains, Eastern New York. Specific goals of this research were to determine joint control on stream orientations and groundwater flow, to compare results with previous studies in the area, and to investigate their tectonic significance. Trails, streams, and road cuts were traversed to locate bedrock outcrops whose positions were determined using topographic maps and a handheld GPS unit. Additional outcrops were located using aerial photographs and GIS data. Joint orientations were measured using a standard Brunton pocket transit. The data was analyzed using Orient (Vollmer, 2010), an orientation analysis program, to plot joint and stream orientations on rose diagrams. ArcGIS was used to produce topographic, hill-shade, and stream drainage maps. Over 500 joint orientations at over 100 outcrop stations were collected. The data were plotted on a rose diagrams, and two major joint sets were found, one with a mean strike of 021° and one with a mean strike of 096°. Stream orientations were also plotted on a rose diagram showing an axial mean of 022°, and indicate that the joint set with mean strike of 021 may have a significant control on stream orientations. The hill-shade maps also demonstrate clearly the strong control of jointing on the topography. The data collected in this research expands on previous joint orientation studies of Engelder and Geiser (1980) in the southwestern and central Catskills, and is similar to joint orientations found by Isachsen et al. (1977) in their study of the Panther Mountain circular structure, a possible impact-related feature. The origin of this jointing is thought to be related to Alleghanian (Permian) and possibly Acadian (Devonian) orogenic events.

  15. A newly discovered K-bentonite zone in the Lower Devonian of the Appalachian Basin; Basal Esopus and Needmore Formations (Late Pragian-Emsian)

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Ver Straeten, C.A.

    1992-01-01

    The K-bentonite-rich interval of the Esopus Formation (eastern New York and northeastern Pennsylvania) overlies the coeval Oriskany/Glenerie/Ridgely Formations and ranges from 1 to 6.3 m in thickness. Six to seventeen soapy-feeling, yellow, tan, green, or gray clay to claystone beds (0.001 to 0.5 m-thick) interbedded with thin siltstone and chert beds (0.02--1 m-thick) characterize outcrops in eastern New York. Heavy mineral separates from these layers yield abundant uncorraded euhedral zircons and apatites, indicating that these are K-bentonites. In eastern Pennsylvania, the westernmost outcrop of the Esopus Formation displays a 2.3 m-thick massive, soapy-feeling clay to claystone-dominated interval. The presence ofmore » both coarse, highly abraded and small, fragile, pristine-appearing zircons and apatites from a 20 cm sampled interval may indicate a complex amalgamation/reworking history to the relatively thick, clay-dominated strata. Similar clay/claystone-rich strata have been found in the lower 0.15 to 1 m of the Beaverdam Member (Needmore Formation) in central Pennsylvania. Interbedded clays and claystones with or without minor siltstone beds characterize some outcrops. Other localities are clay-dominated, with minor amounts of quartz sand present in strata immediately overlying the Ridgely Sandstone. These newly discovered K-bentonite-rich strata mark a transition from shelfal orthoquartzites and carbonates to basinal black/dark gray shales similar to the overlying Middle Devonian Tioga ash interval. Deposition of ash-rich strata, associated with increased volcanic activity, coincided with subsidence of the foreland basin/relative sea level rise. These events were concurrent with a flush of siliciclastic sediments into the basin and are indicative of the onset of an early tectophase of the Devonian Acadian Orogeny.« less

  16. Socially responsible medical education: innovations and challenges in a minority setting.

    PubMed

    Schofield, Aurel; Bourgeois, Daniel

    2010-03-01

    Distributed medical education sites help train, recruit and retain doctors, notably in rural and isolated areas, by providing education and training in these areas and adapting their curriculum to meet the host community's health needs. The Centre de Formation Médicale du Nouveau Brunswick (CFMNB; New Brunswick Medical Education Centre) was established by a partnership between two academic institutions, the Université de Sherbrooke (University of Sherbrooke), situated in the province of Quebec, and the Université de Moncton (University of Moncton), situated in the province of New Brunswick, in Canada. The CFMNB is specifically targeting a minority community (Acadians). Working to establish a high-quality medical education programme, the CFMNB has also set community objectives to meet not only the health needs of this population, but also its social and economic needs. This paper describes the overall objectives of this project, which are: to reduce the gap between community needs and academic institutional needs; to address ethno-cultural and language differences in a defined minority population, and to develop collaboration between the partners involved, including government and community entities which are often perceived as operating in isolation from one another. We also describe why and how the CFMNB developed community-focused objectives and the challenges that came with these innovations, and present lessons from the experience that may be relevant to other sites interested in the social responsibility of medical schools. The CFMNB has produced interesting work and innovations in the field of social responsibility and has encountered many challenges. Continuing interaction between medical education, health research and health services to better address the needs of the population has been established. The information obtained by this process has been used to build a strategic plan for the CFMNB in order to ensure that it is socially responsive and has

  17. K-Ar dating and delta O-18-delta D characterization of nanometric illite from Ordovician K-bentonites of the Appalachians: illitization and the Acadian-Alleghenian tectonic activity

    USGS Publications Warehouse

    Clauer, Norbert; Fallick, Anthony E.; Eberl, Dennis D.; Honty, Miroslav; Huff, Warren D.; Auberti, Amelie

    2013-01-01

    Nanometric (2 diagram that illitization occurred in all fractions by simultaneous nucleation and crystal growth, except for one sample. In that sample, a period of growth without nucleation was detected on top of the nucleation and growth episode. The K-Ar ages organize into two isochrons, the first at 319.9 ± 2.0 Ma with an initial 40Ar/36Ar ratio of 271 ± 66 Ma, and the second at 284.9 ± 1.2 Ma with an initial 40Ar/36Ar ratio of 310 ± 44. One data point above the older isochron and three between the two isochrons suggest a detrital contamination for the former separate and a possible further generation of nanoparticles for the three others. The samples with the older crystallization age consist of illite and illite-rich mixed-layers, and those with the younger age contain smectite-rich mixed-layers without illite, or illite-enriched illite-smectite mixed-layers. The K-Ar ages fit the age trends published previously for similar K-bentonites with regional age patterns between 240 and 270 Ma in the southwestern region, between 270 and 300 Ma in the central zone and the southern Appalachians, and between 315 and 370 Ma in the northernmost. Each of the two generations of illite crystals yields very consistent δ18O (V-SMOW) values at 17 ± 1‰ for the older and at 21 ± 1‰ for the younger. If crystallization temperatures of the nanometric illite were between 100 and 200 °C, as suggested by microthermometric determinations, the hydrothermal fluids had δ18O values of 4 ± 1‰ in the Dalton district and of 8 ± 1‰ in the Lafayette, Trenton, and Dirtseller districts at 100 °C, and of 11 ± 1 and 15 ± 1‰ in the same locations at 200 °C, probably because the water-rock isotope exchanges at elevated temperature occurred in rock-dominated systems. The δ18O of the fluids remained unchanged during local crystal growth, but varied depending on the geographic location of the samples and timing of illitization. The δD (V-SMOW) values of the different size fractions do not provide consistent information; they range from −70 to −45‰ for most nanometric and micrometric fractions (V-SMOW), but with no apparent coherent pattern. Nanometric illite-rich crystals from K-bentonite that underwent tectono-thermal alteration yield constant ages, constant clay mineralogy, constant crystallite size distributions for all of the nucleating and growing illite-type crystals of each sample, as well as constant δ18O values implying constant fluid chemistry, all pointing to geologically sudden crystallization.

  18. MHC-mediated sexual selection on birdsong: Generic polymorphism, particular alleles and acoustic signals.

    PubMed

    Garamszegi, László Zsolt; Zagalska-Neubauer, Magdalena; Canal, David; Blázi, György; Laczi, Miklós; Nagy, Gergely; Szöllősi, Eszter; Vaskuti, Éva; Török, János; Zsebők, Sándor

    2018-06-01

    Several hypotheses predict that the major histocompatibility complex (MHC) drives mating preference in females. Olfactory, colour or morphological traits are often found as reliable signals of the MHC profile, but the role of avian song mediating MHC-based female choice remains largely unexplored. We investigated the relationship between several MHC and acoustic features in the collared flycatcher (Ficedula albicollis), a European passerine with complex songs. We screened a fragment of the class IIB second exon of the MHC molecule, of which individuals harbour 4-15 alleles, while considerable sequence diversity is maintained at the population level. To make statistical inferences from a large number of comparisons, we adopted both null-hypothesis testing and effect size framework in combination with randomization procedures. After controlling for potential confounding factors, neither MHC allelic diversity nor the presence of particular alleles was associated remarkably with the investigated qualitative and quantitative song traits. Furthermore, genetic similarity among males based on MHC sequences was not reflected by the similarity in their song based on syllable content. Overall, these results suggest that the relationship between features of song and the allelic composition and diversity of MHC is not strong in the studied species. However, a biologically motivated analysis revealed that individuals that harbour an MHC allele that impairs survival perform songs with broader frequency range. This finding suggests that certain aspects of the song may bear reliable information concerning the MHC profile of the individuals, which can be used by females to optimize mate choice. © 2018 John Wiley & Sons Ltd.

  19. Tamarix as habitat for birds: Implications for riparian restoration in the Southwestern United States

    USGS Publications Warehouse

    Sogge, M.K.; Sferra, S.J.; Paxton, E.H.

    2008-01-01

    Exotic vegetation has become a major habitat component in many ecosystems around the world, sometimes dramatically changing the vegetation community structure and composition. In the southwestern United States, riparian ecosystems are undergoing major changes in part due to the establishment and spread of the exotic Tamarix (saltcedar, tamarisk). There are concerns about the suitability of Tamarix as habitat for birds. Although Tamarix habitats tend to support fewer species and individuals than native habitats, Arizona Breeding Bird Atlas data and Birds of North America accounts show that 49 species use Tamarix as breeding habitat. Importantly, the relative use of Tamarix and its quality as habitat vary substantially by geographic location and bird species. Few studies have examined how breeding in Tamarix actually affects bird survivorship and productivity; recent research on Southwestern Willow Flycatchers has found no negative effects from breeding in Tamarix habitats. Therefore, the ecological benefits and costs of Tamarix control are difficult to predict and are likely to be species specific and site specific. Given the likelihood that high-quality native riparian vegetation will not develop at all Tamarix control sites, restoration projects that remove Tamarix but do not assure replacement by high-quality native habitat have the potential to reduce the net riparian habitat value for some local or regional bird populations. Therefore, an assessment of potential negative impacts is important in deciding if exotic control should be conducted. In addition, measurable project objectives, appropriate control and restoration techniques, and robust monitoring are all critical to effective restoration planning and execution. ?? 2008 Society for Ecological Restoration International.

  20. Longevity records and survival estimate of birds in a Guatemala rain forest

    USGS Publications Warehouse

    Robbins, C.S.; Dowell, B.; Hines, J.

    2002-01-01

    Birds were mist-netted for ten consecutive 'winter' seasons at two sites on Cerro San Gil and for three to nine seasons at eight other sites on the mountain. Sixteen nets were used at each site for three days; net locations were the same each year. From 1,255 subsequent-year recaptures we computed annual survival using the program MARK. A low annual survival of 0.26+0.03 for Long-tailed Hermit probably reflects extensive wandering in search of food. The only other residents with low survival rates were Ochre-bellied Flycatcher (0.32) and Olive-backed Euphonia (0.38). Other residents tested ranged from 0.49 for Red-capped Manakin to 0.67 for Stub-tailed Spadebill and are within ranges reported from other tropical sites. Rates for migrants were lower, ranging from 0.33 (Worm-eating Warbler) to 0.45 (Kentucky Warbler). Limiting the analysis to known territorial adults (birds that had already returned from a previous year), raised survival rates for residents an average of 0.05, whereas rates for wintering migratory species remained unchanged. The oldest birds recaptured were all residents: Scalythroated Leaftosser (9 years 9 months), Tawny-winged and Wedge-billed Woodcreepers, Northern Bentbill, Tawny-crowned Greenlet, and White-breasted Woodwren (8 years 9 months each). Ages over three years nine months were recorded for 46 species; for the majority of these, new maximum age records were established. A positive relationship was found between survival rate and maximum age and between sample size and maximum age

  1. Ecomorphology, differentiated habitat use, and nocturnal activities of Rhinolophus and Hipposideros species in East Asian tropical forests.

    PubMed

    Lee, Ya-Fu; Kuo, Yen-Min; Chu, Wen-Chen; Lin, Yu-Hsiu; Chang, Hsing-Yi; Chen, Wei-Ming

    2012-02-01

    We investigated the wing morphology and foraging distributions of sympatric Rhinolophus and Hipposideros species by acoustic sampling, measuring wing parameters, and observing bats in different settings of tropical East Asian forests, to evaluate their flexibility in habitat use and edge sensitivity. R. formosae and H. terasensis were more abundant at edges/in open habitats and shared the highest overlap, with R. formosae displaying the greatest breadth in habitat use, whereas R. monoceros had a higher abundance and feeding efficiency in forest interiors with a continuous canopy. H. terasensis was significantly larger and had higher wing loading and aspect ratio than R. formosae and R. monoceros, while R. formosae had higher wing loading but a lower aspect ratio than the smaller-sized R. monoceros. Shrubs and herbs were higher at sites where bats were captured than at those without bat captures, and R. monoceros and R. formosae were associated with greater canopy and ground coverage, respectively. R. monoceros always foraged while flying at lower heights close to the herb/shrub layers, while H. terasensis and R. formosae used perching to different extents, with R. formosae preferably using fly-catching techniques and appearing farther from the path in open forests rather than in forest interiors. Our results indicate that differences in wing parameters account for the different degrees of flexibility in habitat use, yet the deviations of call frequency from the expected values in R. formosae and H. terasensis suggest additional adaptations accounting for their flexibility in exploring habitats. Copyright © 2012 Elsevier GmbH. All rights reserved.

  2. Evaluating blood and excrement as bioindicators for metal accumulation in birds.

    PubMed

    Berglund, Åsa M M

    2018-02-01

    Birds are widely used to assess metal contamination in the environment and there are different approaches to determine the exposure level in individuals, some being destructive (collection of soft tissues) and some non-destructive (blood, feathers and excrement). The use of blood to detect internal concentrations of metals is an acknowledged method, but to what extent blood can predict the concentrations in soft tissues has been less well evaluated in wild terrestrial birds. The same is true for excrements. This study compares the non-destructive methods using blood and excrement with liver sampling, with respect to exposure and accumulation of the elements arsenic, cadmium, copper, lead and zinc in nestling pied flycatchers (Ficedula hypoleuca). Blood, liver and excrement reflected the environmental exposure of non-essential elements and were independent of nestling sex. There were asymptotic relationships between the concentration of arsenic, cadmium and lead in liver and blood, excrement and liver, and excrement and blood, but none for copper or zinc. Those relationships were generally stronger between liver and blood than between excrements and internal concentrations. Lead had the strongest associations for all matrixes. The conclusion is that blood is an appropriate tool to assess accumulation of arsenic, cadmium and especially lead, but that blood can underestimate the accumulation at highly contaminated sites. Excrement can also give an indication of metal accumulation, but may overestimate internal concentrations at high exposure, and individual variability makes direct comparisons between these matrices less appropriate. Copyright © 2017 Elsevier Ltd. All rights reserved.

  3. Rediscovery of the type series of the Acadian Masked Shrew, Sorex acadicus Gilpin, 1865 (Mammalia: Soricidae), with the designation of a neotype and a reevaluation of its taxonomic status

    USGS Publications Warehouse

    Woodman, Neal

    2018-01-01

    The name Sorex acadicus Gilpin, 1865 is currently recognized as the valid name for the Nova Scotian subspecies of the masked shrew, S. cinereus Kerr, 1792 (Mammalia: Soricidae), but a holotype for the taxon was never designated, and the location of the type series has been a mystery. The authority for this species, John Bernard Gilpin, was associated with the Nova Scotia Museum, Halifax, NS, but that institution has no Gilpin specimens in its possession, and I could find no record of Gilpin shrews in any other Canadian Museum. I recently discovered a series of Gilpin specimens in the Mammal Collection of the National Museum of Natural History, Washington, DC (USNM), some of which may have been part of the original type series of S. acadicus, and I show that these specimens best represent Gilpin's concept of the taxon. From this series, I designate a neotype for S. acadicus. I also evaluate the distinctiveness of Nova Scotian S. c. acadicus compared with S. c. cinereus from Maine, New Brunswick, and New Hampshire and determine that S. acadicus should be considered a junior synonym of S. c. cinereus.

  4. Differences in breeding bird assemblages related to reed canary grass cover cover and forest structure on the Upper Mississippi River

    USGS Publications Warehouse

    Kirsch, Eileen M.; Gray, Brian R.

    2017-01-01

    Floodplain forest of the Upper Mississippi River provides habitat for an abundant and diverse breeding bird community. However, reed canary grass Phalaris arundinacea invasion is a serious threat to the future condition of this forest. Reed canary grass is a well-known aggressive invader of wetland systems in the northern tier states of the conterminous United States. Aided by altered flow regimes and nutrient inputs from agriculture, reed canary grass has formed dense stands in canopy gaps and forest edges, retarding tree regeneration. We sampled vegetation and breeding birds in Upper Mississippi River floodplain forest edge and interior areas to 1) measure reed canary grass cover and 2) evaluate whether the breeding bird assemblage responded to differences in reed canary grass cover. Reed canary grass was found far into forest interiors, and its cover was similar between interior and edge sites. Bird assemblages differed between areas with more or less reed canary grass cover (.53% cover breakpoint). Common yellowthroat Geothlypis trichas, black-capped chickadee Parus atricapillus, and rose-breasted grosbeak Pheucticus ludovicianus were more common and American redstart Setophaga ruticilla, great crested flycatcher Myiarchus crinitus, and Baltimore oriole Icterus galbula were less common in sites with more reed canary grass cover. Bird diversity and abundance were similar between sites with different reed canary grass cover. A stronger divergence in bird assemblages was associated with ground cover ,15%, resulting from prolonged spring flooding. These sites hosted more prothonotary warbler Protonotaria citrea, but they had reduced bird abundance and diversity compared to other sites. Our results indicate that frequently flooded sites may be important for prothonotary warblers and that bird assemblages shift in response to reed canary grass invasion.

  5. A study of onchocerciasis with severe skin and eye lesions in a hyperendemic zone in the forest of southwestern Cameroon: clinical, parasitologic, and entomologic findings.

    PubMed

    Somo, R M; Enyong, P A; Fobi, G; Dinga, J S; Lafleur, C; Agnamey, P; Ngosso, A; Ngolle, E M

    1993-01-01

    Prior to the initiation of an onchocerciasis control program based on the mass administration of ivermectin in the rain forest of southwestern Cameroon, a preliminary baseline study of the area was conducted. The results of this study showed that onchocerciasis was hyperendemic in the area. Skin symptoms and signs were observed including pruritus (67.4% of the population examined), onchocerca nodules (51.6%), skin depigmentation (18.5%), and hanging groins (5.7%). Except for pruritus, the prevalence of these symptoms increased with age. Of the eyes examined, 44.9% had microfilariae in the anterior chamber, 33.5% had choroidoretinitis, 28.0% had punctate keratitis, 8.3% had papillary abnormalities, and 3.6% had sclerosing keratitis. Vision in 10.5% of the eyes examined was classified as blind or very poor (visual acuity = 0-0.10), in 15.7% as poor (visual acuity = 0.11-0.39), and in 73.8% as good (visual acuity = 0.4-1.00). Unlike previous reports that have linked serious ocular damage mainly to savanna onchocerciasis, the present study showed that forest onchocerciasis also caused significant ocular pathology, including blindness. Parasitologically, positive skin snips were recorded for 92.7% of the persons examined, with both sexes being equally infected. The parasite load, expressed as the geometric mean number of microfilariae per skin snip, was 53.6, and was much higher in males than in females. The flv vector, Simulium squamosum, had a high infection rate of 7.5% infective females in Bakumba and 6.8% infective females in Ngbandi, the two fly-catching points. The transmission potential was 266 infective larvae per person per month in Bakumba and 189 in Ngbandi.

  6. Natural selection and genetic variation for reproductive reaction norms in a wild bird population.

    PubMed

    Brommer, Jon E; Merilä, Juha; Sheldon, Ben C; Gustafsson, Lars

    2005-06-01

    Many morphological and life-history traits show phenotypic plasticity that can be described by reaction norms, but few studies have attempted individual-level analyses of reaction norms in the wild. We analyzed variation in individual reaction norms between laying date and three climatic variables (local temperature, local rainfall, and North Atlantic Oscillation) of 1126 female collared flycatchers (Ficedula albicollis) with a restricted maximum likehood linear mixed model approach using random-effect best linear unbiased predictor estimates for the elevation (i.e., expected laying date in the average environment) and slope (i.e., adjustment in laying date as a function of environment) of females' reaction norms. Variation in laying date was best explained by local temperature, and individual females differed in both the elevation and the slope of their laying date-temperature reaction norms. As revealed by animal model analyses, there was weak evidence for additive genetic variance of elevation (h2 +/- SE = 0.09 +/- 0.09), whereas there was no evidence for heritability of slope (h2 +/- SE = 0.00 +/- 0.01). Selection analysis, using a female's lifetime production of fledglings or recruits as an estimate of her fitness, revealed significant selection for a lower phenotypic value and breeding value for elevation (i.e., earlier laying date at the average temperature). There was selection for steeper phenotypic values of slope (i.e., greater plasticity in the adjustment of laying date to temperature), but no significant selection on the breeding values of slope. Although these results suggest that phenotypic laying date is influenced by additive genetic factors, as well as by an interaction with the environment, selection on plasticity would not produce an evolutionary response.

  7. Lithospheric buckling and far-foreland deformation during the Laramide and Appalachian orogenies

    NASA Astrophysics Data System (ADS)

    Tikoff, B.; Siddoway, C. S.

    2017-12-01

    Major intraplate tectonics within North America (Laurentia) occurs during times of major orogenesis along the plate margins. During mountain building, typical structures of the hinterland are an orogenic plateau and fold-and-thrust belts, while in the far foreland (intraplate) areas long-wavelength ( 200 km or longer) folds and fault-reactivation features form. Long-wavelength folds are evident in both the Appalachian and Laramide orogenic forelands, with the stratigraphy recording the timing of the uplift. This contribution examines the model of lithospheric buckling - periodic folding associated with a horizontal endload on the edge of the plate - based on scaled, physical experiments and corroborated by numerical models. The Laramide (75-55 Ma) intraplate orogen in the classical location in Wyoming contains basement-cored arches spaced 200 km apart, for which the mechanism of uplift is questioned. Seismic evidence obtained for the Bighorn uplift, Wyoming, obtained by the EarthScope Bighorn project, shows an upwarp of the Moho beneath, but oblique to the trend of the surface exposure of the basement arch. Both the surface and Moho exhibit approximately the same structural relief. The seismic data exhibit no evidence for a regionally continuous decollement, nor is there evidence of rotation of structural markers within these features, of the type that is observed in the detached fold-and-thrust belt. The intraplate region affected by long-wavelength folding includes western Wyoming, with continuation of some features across the E-W-oriented Cheyenne belt (e.g., Rock Springs-Douglas Creek arch), Colorado Plateau, and High Plains east of the Rocky Mountains, where surface and subsurface structures display a series of anticlinal arches ("plains-type" folds). Appalachian mountain building also caused long-wavelength folding, with a spacing consistent with lithospheric buckling, mostly associated with the Devonian Acadian orogeny. The Laramide arches in the High Plains

  8. Predicting the recruitment of established regeneration into the sapling size class following partial cutting in the Acadian Forest Region: Using long-term observations to assess the performance of FVS-NE

    Treesearch

    David Ray; Chad Keyser; Robert Seymour; John Brissette

    2008-01-01

    Forest managers are increasingly called upon to provide long-term predictions of forest development. The dynamics of regeneration establishment, survival and subsequent recruitment of established seedlings to larger size classes is a critical component of these forecasts, yet remains a weak link in available models. To test the reliability of FVS-NE for simulating...

  9. Identification of a feather β-keratin gene exclusively expressed in pennaceous barbule cells of contour feathers in chicken.

    PubMed

    Kowata, Kinue; Nakaoka, Minori; Nishio, Kaori; Fukao, Ayaka; Satoh, Akira; Ogoshi, Maho; Takahashi, Sumio; Tsudzuki, Masaoki; Takeuchi, Sakae

    2014-05-25

    Feathers are elaborate skin appendages shared by birds and theropod dinosaurs that have hierarchical branching of the rachis, barbs, and barbules. Feather filaments consist of β-keratins encoded by multiple genes, most of which are located in tandem arrays on chromosomes 2, 25, and 27 in chicken. The expansion of the genes is thought to have contributed to feather evolution; however, it is unclear how the individual genes are involved in feather formation. The aim of the present study was to identify feather keratin genes involved in the formation of barbules. Using a combination of microarray analysis, reverse-transcription polymerase chain reaction, and in situ hybridization, we found an uncharacterized keratin gene on chromosome 7 that was expressed specifically in barbule cells in regenerating chicken feathers. We have named the gene barbule specific keratin 1 (BlSK1). The BlSK1 gene structure was similar to the gene structure of previously characterized feather keratin genes, and consisted of a non-coding leader exon, an intron, and an exon with an open reading frame (ORF). The ORF was predicted to encode a 98 aa long protein, which shared 59% identity with feather keratin B. Orthologs of BlSK1 were found in the genomes of other avian species, including turkey, duck, zebra finch, and flycatcher, in regions that shared synteny with chromosome 7 of chicken. Interestingly, BlSK1 was expressed in feather follicles that generated pennaceous barbules but not in follicles that generated plumulaceous barbules. These results suggested that the composition of feather keratins probably varies depending on the structure of the feather filaments and, that individual feather keratin genes may be involved in building different portions and/or types of feathers in chicken. Copyright © 2014 Elsevier B.V. All rights reserved.

  10. A sampling plan for riparian birds of the Lower Colorado River-Final Report

    USGS Publications Warehouse

    Bart, Jonathan; Dunn, Leah; Leist, Amy

    2010-01-01

    A sampling plan was designed for the Bureau of Reclamation for selected riparian birds occurring along the Colorado River from Lake Mead to the southerly International Boundary with Mexico. The goals of the sampling plan were to estimate long-term trends in abundance and investigate habitat relationships especially in new habitat being created by the Bureau of Reclamation. The initial objective was to design a plan for the Gila Woodpecker (Melanerpes uropygialis), Arizona Bell's Vireo (Vireo bellii arizonae), Sonoran Yellow Warbler (Dendroica petechia sonorana), Summer Tanager (Piranga rubra), Gilded Flicker (Colaptes chrysoides), and Vermilion Flycatcher (Pyrocephalus rubinus); however, too little data were obtained for the last two species. Recommendations were therefore based on results for the first four species. The study area was partitioned into plots of 7 to 23 hectares. Plot borders were drawn to place the best habitat for the focal species in the smallest number of plots so that survey efforts could be concentrated on these habitats. Double sampling was used in the survey. In this design, a large sample of plots is surveyed a single time, yielding estimates of unknown accuracy, and a subsample is surveyed intensively to obtain accurate estimates. The subsample is used to estimate detection ratios, which are then applied to the results from the extensive survey to obtain unbiased estimates of density and population size. These estimates are then used to estimate long-term trends in abundance. Four sampling plans for selecting plots were evaluated based on a simulation using data from the Breeding Bird Survey. The design with the highest power involved selecting new plots every year. Power with 80 plots surveyed per year was more than 80 percent for three of the four species. Results from the surveys were used to provide recommendations to the Bureau of Reclamation for their surveys of new habitat being created in the study area.

  11. A Water Budget for Riparian Vegetation on the Lower Colorado River: the Myth of Water Salvage

    NASA Astrophysics Data System (ADS)

    Nagler, P. L.; Glenn, E. P.; Webb, R. H.; Howard, K. A.

    2007-05-01

    For many years, river managers have envisaged large saving of water by clearing the exotic plant, saltcedar (Tamarix ramosissima) from western U.S. rivers. Early estimates of evapotranspiration (ET) by saltcedar ranged as high as 3-4 m/yr, and it was estimated that saltcedar on the Lower Colorado River used more water than Los Angeles. Furthermore, saltcedar was considered to have low habitat value, so clearing projects might enhance habitat value by allowing the return of more valuable native species. We have examined these assumptions based on recent evidence. Moisture flux towers set in dense saltcedar stands show that ET is moderate, ranging from 0.8-1.4 m/yr with a mean value of 1 m/yr over five studies on three rivers, similar to wide-area estimates from remote sensing studies. Projected over the 18,200 ha of dense saltcedar monocultures estimated for the Lower Colorado River riparian corridor in the U.S., the potential water saving would only be about 1 percent of the annual flow (assuming no replacement vegetation). A similar acreage of saltcedar monoculture exists in the Colorado River delta in Mexico, but these stands are supported by outflow of brackish water from the irrigation district rather than river water. The assumption of low habitat value is not supported by recent studies. For example, Hinojosa- Huerta (2006) found that saltcedar monocultures away from the river channel supported 65 percent as many bird numbers and 74 percent as many bird species as the best habitat type, mixed saltcedar and native trees in proximity to water, in the delta of the Colorado River in Mexico, and saltcedar provided equal habitat value as native trees for endangered willow flycatchers on Arizona and New Mexico rivers (Owen et al., 2005). Hence, the prospects for saving water without destroying habitat by clearing saltcedar are doubtful for this river system.

  12. Delayed diagnosis of a patient with Usher syndrome 1C in a Louisiana Acadian family highlights the necessity of timely genetic testing for the diagnosis and management of congenital hearing loss.

    PubMed

    Umrigar, Ayesha; Musso, Amanda; Mercer, Danielle; Hurley, Annette; Glausier, Cassondra; Bakeer, Mona; Marble, Michael; Hicks, Chindo; Tsien, Fern

    2017-01-01

    Advances in sequencing technologies and increased understanding of the contribution of genetics to congenital sensorineural hearing loss have led to vastly improved outcomes for patients and their families. Next-generation sequencing and diagnostic panels have become increasingly reliable and less expensive for clinical use. Despite these developments, the diagnosis of genetic sensorineural hearing loss still presents challenges for healthcare providers. Inherited sensorineural hearing loss has high levels of genetic heterogeneity and variable expressivity. Additionally, syndromic hearing loss (hearing loss and additional clinical abnormalities) should be distinguished from non-syndromic (hearing loss is the only clinical symptom). Although the diagnosis of genetic sensorineural hearing loss can be challenging, the patient's family history and ethnicity may provide critical information, as certain genetic mutations are more common in specific ethnic populations. The early identification of the cause of deafness can benefit patients and their families by estimating recurrence risks for future family planning and offering the proper interventions to improve their quality of life. Collaboration between pediatricians, audiologists, otolaryngologists, geneticists, and other specialists are essential in the diagnosis and management of patients with hearing disorders. An early diagnosis is vital for proper management and care, as some clinical manifestations of syndromic sensorineural hearing loss are not apparent at birth and have a delayed age of onset. We present a case of Usher syndrome (congenital deafness and childhood-onset blindness) illustrating the challenges encountered in the diagnosis and management of children presenting with congenital genetic sensorineural hearing loss, along with helpful resources for clinicians and families.

  13. Gondwanan/peri-Gondwanan origin for the Uchee terrane, Alabama and georgia: Carolina zone or Suwannee terrane(?) and its suture with Grenvillian basement of the Pine Mountain window

    USGS Publications Warehouse

    Steltenpohl, M.G.; Mueller, P.M.; Heatherington, A.L.; Hanley, T.B.; Wooden, J.L.

    2008-01-01

    The poorly known, suspect, Uchee terrane occupies a critical tectonic position with regard to how and when peri-Gondwanan (Carolina) and Gondwanan (Suwannee) terranes were sutured to Laurentia. It lies sandwiched between Laurentian(?) continental basement exposed in the Pine Mountain window and adjacent buried Gondwanan crust of the Suwannee terrane. The Uchee terrane has been proposed as both a septum of Piedmont rocks that once was continuous across the erosionally breached Pine Mountain window or part of the Carolina zone. To help resolve this issue, we conducted U-Pb (SHRIMP-RG) (sensitive high-resolution ion microprobe-reverse geometry) zircon studies and whole-rock isotopic analyses of principal metasedimentary and metaplutonic units. U-Pb ages for zircons from the Phenix City Gneiss suggest igneous crystallization at ca. 620 Ma, inheritance ca. 1000 to ca. 1700 Ma, and a ca. 300 Ma (Alleghanian) overprint recorded by zircon rims. Zircons from the metasedimentary/metavolcaniclastic Moffits Mill Schist yield bimodal dates at ca. 620 and 640 Ma. The 620 to 640 Ma dates make these rocks age-equivalent to the oldest parts of the Carolina slate belt (Virgilina and Savannah River) and strongly suggest a Gondwanan (Pan-African and/or Trans-Brasiliano) origin for the Uchee terrane. Alternatively, the Uchee terrane may be correlative with metamorphic basement of the Suwannee terrane. The ca. 300 Ma overgrowths on zircons are compatible with previously reported 295 to 288 Ma 40Ar/39Ar hornblende dates on Uchee terrane rocks, which were interpreted to indicate deep tectonic burial of the Uchee terrane contemporaneous with the Alleghanian orogeny recorded in the foreland. Temperature-time paths for the Uchee terrane are similar to that of the Pine Mountain terrane, indicating a minimum age of ca. 295 Ma for docking. In terms of tectono-metamorphic history of the Uchee terrane, it is important to note that no evidence for intermediate "Appalachian" dates (e.g, Acadian or

  14. Rifting along the northern Gondwana margin and the evolution of the Rheic Ocean: A Devonian age for the El Castillo volcanic rocks (Salamanca, Central Iberian Zone)

    NASA Astrophysics Data System (ADS)

    Gutiérrez-Alonso, G.; Murphy, J. B.; Fernández-Suárez, J.; Hamilton, M. A.

    2008-12-01

    Exposures of volcanic rocks (El Castillo) in the Central Iberian Zone near Salamanca, Spain, are representative of Paleozoic volcanic activity along the northern Gondwanan passive margin. Alkaline basalts and mafic volcaniclastic rocks of this sequence are structurally preserved in the core of the Variscan-Tamames Syncline. On the basis of the occurrence of graptolite fossils in immediately underlying strata, the El Castillo volcanics traditionally have been regarded as Lower Silurian in age. In contrast, most Paleozoic volcanic units in western Iberia are rift-related mafic to felsic rocks emplaced during the Late Cambrian-Early Ordovician, and are attributed to the opening of the Rheic Ocean. We present new zircon U-Pb TIMS data from a mafic volcaniclastic rock within the El Castillo unit. These data yield a near-concordant, upper intercept age of 394.7 ± 1.4 Ma that is interpreted to reflect a Middle Devonian (Emsian-Eifelian) age for the magmatism, demonstrating that the El Castillo volcanic rocks are separated from underlying lower Silurian strata by an unconformity. The U-Pb age is coeval with a widespread extensional event in Iberia preserved in the form of a generalized paraconformity surface described in most of the Iberian Variscan realm. However, in the inner part of the Gondwanan platform, the Cantabrian Zone underwent a major, coeval increase in subsidence and the generation of sedimentary troughs. From this perspective, the eruption age reported here probably represents a discrete phase of incipient rifting along the southern flank of the Rheic Ocean. Paleogeographic reconstructions indicate that this rifting event was coeval with widespread orogeny and ridge subduction along the conjugate northern flank of the Rheic Ocean, the so called Acadian "orogeny". We speculate that ridge subduction resulted in geodynamic coupling of the northern and southern flanks of the Rheic Ocean, and that the extension along the southern flank of the Rheic Ocean is a

  15. Two new genera of songbirds represent endemic radiations from the Shola Sky Islands of the Western Ghats, India.

    PubMed

    Robin, V V; Vishnudas, C K; Gupta, Pooja; Rheindt, Frank E; Hooper, Daniel M; Ramakrishnan, Uma; Reddy, Sushma

    2017-01-23

    A long-standing view of Indian biodiversity is that while rich in species, there are few endemics or in-situ radiations within the subcontinent. One exception is the Western Ghats biodiversity hotspot, an isolated mountain range with many endemic species. Understanding the origins of the montane-restricted species is crucial to illuminate both taxonomic and environmental history. With evidence from genetic, morphometric, song, and plumage data, we show that two songbird lineages endemic to the Western Ghats montane forest each have diversified into multiple distinct species. Historically labeled as single species of widespread Asian genera, these two lineages are highly divergent and do not group with the taxa in which they were previously classified but rather are distinct early divergences in larger Asian clades of flycatchers and babblers. Here we designated two new genera, the Western Ghats shortwings as Sholicola and the laughingthrushes as Montecincla, and evaluated species-limits to reflect distinct units by revising six previously named taxa and describing one novel species. Divergence dating showed that both these montane groups split from their Himalayan relatives during the Miocene, which is coincident with a shift towards arid conditions that fragmented the previously contiguous humid forest across peninsular India and isolated these lineages in the Western Ghats. Furthermore, these two genera showed congruent patterns of diversification across the Western Ghats Sky Islands, coincident with other climatic changes. Our study reveals the existence of two independent endemic radiations in the high montane Western Ghats or Shola Sky Islands with coincident divergence times, highlighting the role of climate in the diversification of these ancient lineages. The endemic and highly divergent nature of these previously unrecognized species underscores the dearth of knowledge about the biogeography of the Asian tropics, even for comparatively well-known groups

  16. Breeding biology of the golden-faced Tyrannulet (Zimmerius chrysops) in venezuela

    USGS Publications Warehouse

    Goulding, W.; Martin, T.E.

    2010-01-01

    We present the first detailed information on the breeding biology of the Golden-faced Tyrannulet (Zimmerius chrysops). Information was gathered from 96 nests in Yacamb National Park, Venezuela during the 2002 to 2008 breeding seasons. The enclosed nest was similar to descriptions of nests of other species in the genus. Eggs were laid on alternate days with mean (?? SE) clutch size of 1.98 ?? 0.02 (n 45) and fresh weight of 1.616 ?? 0.020 g (n 48). Only the female incubated and the incubation period averaged 16.9 ?? 0.3 days (n 10). Nest attentiveness ( time on the nest) averaged 66.0 ?? 1.6 (n 40) and increased from early to mid- and late-incubation. Incubation behavior yielded an average 24-hr egg temperature of 34.88 ?? 0.45?? C (n 7 nests, 43 days). The nestling growth rate constant for body mass (k 0.285 ?? 0.011) was slow even for tropical tyrannids. The nestling period for nests where exact hatch and fledging days were observed ranged from 17 to 19 days with an average of 18.0 ?? 0. 2 days (n 9). Both females and males fed nestlings at a rate that increased over the nestling period with a mean of 4.41 ?? 0.65 trips/hr (n 10) during days 1 and 2 after hatching, and 14.93 ?? 2.36 trips/hr (n 6) at pin-break (days 1011). Daily predation rates were similar in egg-laying (0.052 ?? 0.025; n 76.5 exposure days) and incubation periods (0.068 ?? 0.010; n 575.5 exposure days), but were lower during the nestling period (0.039 ?? 0.010; n 377.0 exposure days). The total daily predation rate (0.057 ?? 0.007; n 989.0 exposure days) indicated only 12 of nests were successful. These breeding biology parameters for Z. chrysops differ substantially from other tyrant-flycatchers and temperate species, further highlighting the diversity within the Tyrannidae. ?? 2010 by the Wilson Ornithological Society.

  17. Documentation of the Ecological Risk Assessment Computer Model ECORSK.5

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Anthony F. Gallegos; Gilbert J. Gonzales

    1999-06-01

    The FORTRAN77 ecological risk computer model--ECORSK.5--has been used to estimate the potential toxicity of surficial deposits of radioactive and non-radioactive contaminants to several threatened and endangered (T and E) species at the Los Alamos National Laboratory (LANL). These analyses to date include preliminary toxicity estimates for the Mexican spotted owl, the American peregrine falcon, the bald eagle, and the southwestern willow flycatcher. This work has been performed as required for the Record of Decision for the construction of the Dual Axis Radiographic Hydrodynamic Test (DARHT) Facility at LANL as part of the Environmental Impact Statement. The model is dependent onmore » the use of the geographic information system and associated software--ARC/INFO--and has been used in conjunction with LANL's Facility for Information Management and Display (FIMAD) contaminant database. The integration of FIMAD data and ARC/INFO using ECORSK.5 allows the generation of spatial information from a gridded area of potential exposure called an Ecological Exposure Unit. ECORSK.5 was used to simulate exposures using a modified Environmental Protection Agency Quotient Method. The model can handle a large number of contaminants within the home range of T and E species. This integration results in the production of hazard indices which, when compared to risk evaluation criteria, estimate the potential for impact from consumption of contaminants in food and ingestion of soil. The assessment is considered a Tier-2 type of analysis. This report summarizes and documents the ECORSK.5 code, the mathematical models used in the development of ECORSK.5, and the input and other requirements for its operation. Other auxiliary FORTRAN 77 codes used for processing and graphing output from ECORSK.5 are also discussed. The reader may refer to reports cited in the introduction to obtain greater detail on past applications of ECORSK.5 and assumptions used in deriving model parameters.« less

  18. Inventory of montane-nesting birds in Katmai and Lake Clark national parks and preserves

    USGS Publications Warehouse

    Ruthrauff, Daniel R.; Tibbitts, Lee; Gill, Robert E.; Handel, Colleen M.

    2007-01-01

    As part of the National Park Service’s Inventory and Monitoring Program, biologists from the U. S. Geological Survey’s Alaska Science Center conducted an inventory of birds in montane regions of Katmai and Lake Clark National Parks and Preserves during 2004–2006. We used a stratified random survey design to allocate samples by ecological subsection. To survey for birds, we conducted counts at 468 points across 29, 10-km x 10-km (6.2-mi x 6.2-mi) sample plots in Katmai and 417 points across 25, 10-km x 10-km sample plots in Lake Clark. We detected 92 and 104 species in Katmai and Lake Clark, respectively, including 40 species of conservation concern. We detected three species not previously recorded in Katmai (Ring-necked Duck [Aythya collaris], Lesser Scaup [Aythya affinis], and White-tailed Ptarmigan [Lagopus leucurus]) and two species not previously recorded in Lake Clark (Northern Flicker [Colaptes auratus ] and Olive-sided Flycatcher [Contopus cooperi]). The most commonly detected species in both parks was Golden-crowned Sparrow (Zonotrichia atricapilla); Fox Sparrow (Passerella iliaca) and American Pipit (Anthus rubescens) were abundant and widely-distributed as well. We defined sites as low (100–350 m), middle (351–600 m), or high (601–1,620 m) elevation based on the distribution of vegetation cover, and similarly categorized the 34 most-commonly detected species based on the mean elevation of sample points at which they were detected. High elevation (i.e., alpine) sites were characterized by high percent cover of dwarf shrub and bare ground habitat and supported species like Rock Ptarmigan (L. mutus), American Golden-Plover (Pluvialis dominica), Wandering Tattler (Tringa incana), Surfbird (Aphriza virgata), and Snow Bunting (Plectrophenax nivalis), all species of conservation concern. This inventory represents the first systematic survey of birds nesting in montane regions of both parks. Results from this inventory can form the foundation of

  19. Potential for Water Savings by Defoliation of Saltcedar (Tamarix spp.) by Saltcedar Beetles (Diorhabda carinulata) in the Upper Colorado River Basin

    NASA Astrophysics Data System (ADS)

    Nagler, P. L.; Nguyen, U.; Bateman, H. L.; Jarchow, C.; van Riper, C., III; Waugh, W.; Glenn, E.

    2016-12-01

    Northern saltcedar beetles (Diorhabda carinata) have spread widely in riparian zones on the Colorado Plateau since their initial release in 2002. One goal of the releases was to reduce water consumption by saltcedar in order to conserve water through reduction of evapotranspiration (ET). The beetle moved south on the Virgin River and reached Big Bend State Park in Nevada in 2014, an expansion rate of 60 km/year. This is important because the beetle's photoperiod requirement for diapause was expected to prevent them from moving south of 37°N latitude, where endangered southwest willow flycatcher habitat occurs. In addition to focusing on the rate of dispersal of the beetles, we used remote sensing estimates of ET at 13 sites on the Colorado, San Juan, Virgin and Dolores rivers and their tributaries to estimate riparian zone ET before and after beetle releases. We estimate that water savings from 2007-2015 was 31.5 million m3/yr (25,547 acre-ft/yr), amounting to 0.258 % of annual river flow from the Upper Colorado River Basin to the Lower Basin. Reasons for the relatively low potential water savings are: 1) baseline ET before beetle release was modest (0.472 m/yr); 2) reduction in ET was low (0.061 m/yr) because saltcedar stands tended to recover after defoliation; 3) riparian ET even in the absence of beetles was only 1.8 % of river flows, calculated as the before beetle average annual ET (472 mm/yr) times the total area of saltcedar (51,588 ha) divided by the combined total average annual flows (1964-2015) from the upper to lower catchment areas of the Colorado River Basin at the USGS gages (12,215 million m3/yr or 9.90 million acre-ft). Further research is suggested to concentrate on the ecological impacts (both positive and negative) of beetles on riparian zones and on identifying management options to maximize riparian health.

  20. Examining the effect of altered redox conditions on deep soil organic matter stability

    NASA Astrophysics Data System (ADS)

    Gabriel, C.; Kellman, L. M.; Ziegler, S. E.

    2013-12-01

    Since subsoil horizons contribute significantly to terrestrial carbon (C) budgets, understanding the influence of disturbances such as forest harvesting on subsoil C stability is critical. Clearcut harvesting leads to changes in the soil physico-chemical environment, including altering redox conditions arising from changes in soil hydrology that increase soil saturation, soil temperature, and pH. These physico-chemical changes have the potential to alter the adsorption of soil organic matter (SOM) to minerals, particularly at depth where SOM is primarily associated with mineral phases. The objective of this study was to determine the effect of differing redox states (aerobic vs. anaerobic) and temperature upon SOM stability of forested soils representative of the Acadian Forest Region of Eastern North America. Composite soil samples through depth (0-10, 10-20, 20-35, and 35-50 cm) from a mature red spruce forest (110 years) were incubated under optimum (aerobic) or saturated (anaerobic) conditions for 1 or 4 months at two temperatures (5 and 15 C). Following incubation, soil leachate was analyzed for dissolved organic carbon (DOC), and UV-vis absorbance in order to determine soil C losses and its optical character. Specific UV-vis absorbance SUVA (254 nm) and spectral slope ratios were calculated in order to assess the composition of chromophoric dissolved organic matter (CDOM). Preliminary results from the 1 month incubation indicate that under anaerobic conditions, all depths released DOC with a higher SUVA than under aerobic conditions, with the largest change observed in the 0-10 cm depth increment. Soil incubated at 5 C produced leachate with significantly less DOC and with a lower absorbance compared to 15 C under both redox conditions. These results suggest that both temperature and redox state are important in determining the aromaticity of DOC released from soils. Spectral slope ratios revealed that a greater proportion of CDOM of lower molecular weight

  1. Provenance of Modern Soils and Limestone and Chert Bedrock of Middle Tennessee Assessed Using Detrital Zircon U-Pb Geochronology

    NASA Astrophysics Data System (ADS)

    Ayers, J. C.; Katsiaficas, N. J.; Wang, X.

    2014-12-01

    overlap with the Grenville, Taconic and Acadian orogenies of eastern North America. This study demonstrates that dating of detrital zircon is a powerful tool for determining the provenance of soil and limestone.

  2. Precipitation of Oriented Rutile and Ilmenite Needles in Garnet, Northeastern Connecticut, USA: Evidence for Extreme Metamorphic Conditions?

    NASA Astrophysics Data System (ADS)

    Ague, J. J.; Eckert, J. O.

    2011-12-01

    We report the discovery of oriented needles of rutile and, less commonly, ilmenite in the cores of garnets from northeastern CT, USA. The rocks preserve granulite facies mineral assemblages, form part of the Merrimack Synclinorium, and underwent metamorphism and deformation during the Acadian orogeny. The needles appear identical to those reported from a number of extreme P-T environments worldwide, including UHP metamorphic rocks, high-P granulites, and garnet peridotites. The needles are predominantly oriented along <111> directions in garnet. The long axes of the rutile needles commonly do not go extinct parallel to the cross hairs under cross-polarized light (e.g., Griffin et al., 1971). This anomalous extinction indicates that the needles do not preserve a specific crystallographic relationship with their garnet hosts (e.g., Hwang et al., 2007). The needles range from a few hundred nm to a few um in diameter, and can be mm-scale in length. Micrometer-scale plates of rutile, srilankite and crichtonite have also been observed in some garnets together with the Fe-Ti oxide needles. Several origins for the needles have been proposed in the literature; we investigate the hypothesis that they precipitated in situ from originally Ti-rich garnet. Chemical profiles across garnets indicate that some retain Ti zoning, with elevated-Ti concentrations in the cores dropping to low values in the rims. For these zoned garnets, high-resolution, 2-D chemical mapping using the JEOL JXA-8530F field emission gun electron microprobe at Yale University reveals that the needles are surrounded by well-defined Ti-depletion halos. Chemical profiles also document strong depletions of Cr (which is present in both rutile and ilmenite) directly adjacent to needles. The observed Ti-depletions demonstrate that the needles precipitated from Ti-bearing garnet, probably during cooling and/or decompression associated with exhumation. The rutile precipitates must be largely incoherent with respect

  3. Marine species distribution shifts on the U.S. Northeast Continental Shelf under continued ocean warming

    NASA Astrophysics Data System (ADS)

    Kleisner, Kristin M.; Fogarty, Michael J.; McGee, Sally; Hare, Jonathan A.; Moret, Skye; Perretti, Charles T.; Saba, Vincent S.

    2017-04-01

    The U.S. Northeast Continental Shelf marine ecosystem has warmed much faster than the global ocean and it is expected that this enhanced warming will continue through this century. Complex bathymetry and ocean circulation in this region have contributed to biases in global climate model simulations of the Shelf waters. Increasing the resolution of these models results in reductions in the bias of future climate change projections and indicates greater warming than suggested by coarse resolution climate projections. Here, we used a high-resolution global climate model and historical observations of species distributions from a trawl survey to examine changes in the future distribution of suitable thermal habitat for various demersal and pelagic species on the Shelf. Along the southern portion of the shelf (Mid-Atlantic Bight and Georges Bank), a projected 4.1 °C (surface) to 5.0 °C (bottom) warming of ocean temperature from current conditions results in a northward shift of the thermal habitat for the majority of species. While some southern species like butterfish and black sea bass are projected to have moderate losses in suitable thermal habitat, there are potentially significant increases for many species including summer flounder, striped bass, and Atlantic croaker. In the north, in the Gulf of Maine, a projected 3.7 °C (surface) to 3.9 °C (bottom) warming from current conditions results in substantial reductions in suitable thermal habitat such that species currently inhabiting this region may not remain in these waters under continued warming. We project a loss in suitable thermal habitat for key northern species including Acadian redfish, American plaice, Atlantic cod, haddock, and thorney skate, but potential gains for some species including spiny dogfish and American lobster. We illustrate how changes in suitable thermal habitat of important commercially fished species may impact local fishing communities and potentially impact major fishing ports

  4. Le vocabulaire disponible du francais, Tome 1. Le vocabulaire concret usuel des enfants francais et acadiens: Etude temoin (The Working French Vocabulary, Volume 1. Common Generic Terms Used by French and Acadian Children: A Field Study).

    ERIC Educational Resources Information Center

    Mackey, William F.; And Others

    The first of a two-volume study of the relative accessibility of French vocabulary in French-speaking Canada presents statistical data concerning the frequency, distribution, valence, and accessibility of vocabulary related to 16 fundamental centers of interest found in normal conversation. The scope, procedures, and results of the study are…

  5. Unpredictable food supply modifies costs of reproduction and hampers individual optimization.

    PubMed

    Török, János; Hegyi, Gergely; Tóth, László; Könczey, Réka

    2004-11-01

    Investment into the current reproductive attempt is thought to be at the expense of survival and/or future reproduction. Individuals are therefore expected to adjust their decisions to their physiological state and predictable aspects of environmental quality. The main predictions of the individual optimization hypothesis for bird clutch sizes are: (1) an increase in the number of recruits with an increasing number of eggs in natural broods, with no corresponding impairment of parental survival or future reproduction, and (2) a decrease in the fitness of parents in response to both negative and positive brood size manipulation, as a result of a low number of recruits, poor future reproduction of parents, or both. We analysed environmental influences on costs and optimization of reproduction on 6 years of natural and experimentally manipulated broods in a Central European population of the collared flycatcher. Based on dramatic differences in caterpillar availability, we classified breeding seasons as average and rich food years. The categorization was substantiated by the majority of present and future fitness components of adults and offspring. Neither observational nor experimental data supported the individual optimization hypothesis, in contrast to a Scandinavian population of the species. The quality of fledglings deteriorated, and the number of recruits did not increase with natural clutch size. Manipulation revealed significant costs of reproduction to female parents in terms of future reproductive potential. However, the influence of manipulation on recruitment was linear, with no significant polynomial effect. The number of recruits increased with manipulation in rich food years and tended to decrease in average years, so control broods did not recruit more young than manipulated broods in any of the year types. This indicates that females did not optimize their clutch size, and that they generally laid fewer eggs than optimal in rich food years. Mean

  6. Evaluating carbon storage, timber harvest, and habitat possibilities for a Western Cascades (USA) forest landscape.

    PubMed

    Kline, Jeffrey D; Harmon, Mark E; Spies, Thomas A; Morzillo, Anita T; Pabst, Robert J; McComb, Brenda C; Schnekenburger, Frank; Olsen, Keith A; Csuti, Blair; Vogeler, Jody C

    2016-10-01

    Forest policymakers and managers have long sought ways to evaluate the capability of forest landscapes to jointly produce timber, habitat, and other ecosystem services in response to forest management. Currently, carbon is of particular interest as policies for increasing carbon storage on federal lands are being proposed. However, a challenge in joint production analysis of forest management is adequately representing ecological conditions and processes that influence joint production relationships. We used simulation models of vegetation structure, forest sector carbon, and potential wildlife habitat to characterize landscape-level joint production possibilities for carbon storage, timber harvest, and habitat for seven wildlife species across a range of forest management regimes. We sought to (1) characterize the general relationships of production possibilities for combinations of carbon storage, timber, and habitat, and (2) identify management variables that most influence joint production relationships. Our 160 000-ha study landscape featured environmental conditions typical of forests in the Western Cascade Mountains of Oregon (USA). Our results indicate that managing forests for carbon storage involves trade-offs among timber harvest and habitat for focal wildlife species, depending on the disturbance interval and utilization intensity followed. Joint production possibilities for wildlife species varied in shape, ranging from competitive to complementary to compound, reflecting niche breadth and habitat component needs of species examined. Managing Pacific Northwest forests to store forest sector carbon can be roughly complementary with habitat for Northern Spotted Owl, Olive-sided Flycatcher, and red tree vole. However, managing forests to increase carbon storage potentially can be competitive with timber production and habitat for Pacific marten, Pileated Woodpecker, and Western Bluebird, depending on the disturbance interval and harvest intensity chosen

  7. Migratory timing, rate, routes and wintering areas of White-crested Elaenia (Elaenia albiceps chilensis), a key seed disperser for Patagonian forest regeneration

    PubMed Central

    Bravo, Susana Patricia; Cueto, Victor Rodolfo; Gorosito, Cristian Andrés

    2017-01-01

    Migratory animals often play key ecological roles within the communities they visit throughout their annual journeys. As a consequence of the links between biomes mediated by migrants, changes in one biome could affect remote areas in unpredictable ways. Migratory routes and timing of most Neotropical austral migrants, which breed at south temperate latitudes of South America and overwinter closer to or within tropical latitudes of South America, have yet to be described in detail. As a result, our understanding about how these birds provide links between South American biomes is almost non-existent. White-crested Elaenia (Elaenia albiceps chilensis) is a long-distance austral migrant that breeds in the Patagonian Forest biome and overwinters in tropical South America. Because this small flycatcher plays a key role in the regeneration of this ecosystem, our objective was to describe the annual cycle of White-crested elaenias to evaluate the degree of migratory connectivity between breeding and wintering areas and therefore to determine if there are specific biomes of northern South America linked by elaenias to Patagonian forests. Fifteen individuals were successfully tracked throughout a complete migration cycle using miniature light-level geolocators. All individuals resided and moved through the same general regions. During fall (March-April-May), elaenias were located in the Caatinga and the Atlantic Forest biomes, from Rio de Janeiro to the region near Salvador da Bahia, Brazil. During winter (June-July-Aug.), birds were located further inland, within the Cerrado biome. Birds used three different routes during fall migration. Our results indicate that some individuals use a direct route, flying between 500–600 km/day, crossing desert and grasslands, while others took a detour, flying 100–200 km/day through forested areas with refueling opportunities. All birds used the Yunga forest during spring migration, with ten out of 15 individuals showing a clear

  8. Seismotectonics of northeastern United States and adjacent Canada

    NASA Astrophysics Data System (ADS)

    Yang, Jih-Ping; Aggarwal, Yash Pal

    1981-06-01

    appears to predominate. The western margin of this province, however, appears to be relatively aseismic. We attribute this relative lack of activity to one or more of the following: the presence of igneous activity postdating rifting of Africa from North America, the occurrence of intense metamorphism during the Acadian orogeny which may have annealed preexisting faults, and the predominance of ductile as opposed to brittle deformation in the geologic past. The inferred axis of maximum horizontal compression along the eastern margin of the Appalachians is rather uniform and trends W-WNW, almost perpendicular to the magnetic lineations offshore. We suggest that this W-WNW compression reflects the gravitational force arising from horizontal density variations in the oceanic lithosphere as it cools and moves away from spreading centers.

  9. Bedrock geologic map of the Lisbon quadrangle, and parts of the Sugar Hill and East Haverhill quadrangles, Grafton County, New Hampshire

    USGS Publications Warehouse

    Rankin, Douglas W.

    2018-04-20

    , siltstone, and phyllite.During the Late Ordovician, a series of arc-related plutons intruded the Ammonoosuc Volcanics including the Moody Ledge pluton and the Scrag granite of Billings (1937). Subsequent plutonism related to the Acadian orogeny occurred after volcanism and deposition resulted in the Littleton Formation during the Late Devonian, including the intrusion of the Haverhill pluton and French Pond Granite found in the southern part of the map.This report consists of a geologic map and an online geographic information systems database that includes contacts of bedrock geologic units, faults, outcrops, and structural geologic information. The geologic map is intended to serve as a foundation for applying geologic information to problems involving land use decisions, groundwater availability and quality, earth resources such as natural aggregate for construction, assessment of natural hazards, and engineering and environmental studies for waste disposal sites and construction projects.

  10. Origin of the Lyme Dome and implications for the timing of multiple Alleghanian deformational and intrusive events in southern Connecticut

    USGS Publications Warehouse

    Walsh, G.J.; Aleinikoff, J.N.; Wintsch, R.P.

    2007-01-01

    unloading and exhumation. Monazite and metamorphic zircon rim ages record this event at ca. 259 Ma. The evolution of the Lyme dome involved D1 mylonitization, intrusion, and migmatization during north-directed contraction, limited late D1 diapirism, D2 migmatization during WNW contraction with associated flexural flow and fold interference, D3 NNW horizontal extension and decompression melting, and final D4 vertical extension and rapid exhumation. Late regional uplift, extension, and normal faulting at higher crustal levels may have been caused by diapiric rise of the lower crust, below the structural level of the Lyme dome. The rocks record no evidence of Acadian metamorphism or deformation, suggesting that the Gander zone here was not tectonically juxtaposed with Avalon until the Alleghanian orogeny.

  11. A transect through the base of the Bronson Hill Terrane in western New Hampshire

    USGS Publications Warehouse

    Walsh, Gregory J.; Valley, Peter M.; Sicard, Karri R.; Thompson, Thelma Barton; Thompson, Peter J.

    2012-01-01

    regards to the stratigraphy of the BHA and CVT, and the distribution, or even existence, of faults ranging in age from Devonian to Mesozoic (e.g., Spear and others, 2008; McWilliams and others, 2010; Walsh and others, 2010). Questions to ponder on this trip include, but are not limited to: 1) Is the Bronson Hill anticlinorium allochthonous? 2) What is the crust beneath the Bronson Hill anticlinorium? 3) Is there a “Big Staurolite nappe” as proposed by Spear and others (2002, 2003, 2008)? 4) What is the role of Taconic, Acadian, and Alleghanian orogenesis in the tectonic development of the region? Modern 1:24,000-scale mapping is the first step towards answering these questions. Mapping will be supplemented by modern geochronology and geochemistry as this project develops. We plan to share some of our provisional results during this field trip.

  12. Sub-pluvial Saqqara and its possible impact on ancient Egyptian civilization in the Old Kingdom Period (4600 - 4100 yrs BP)

    NASA Astrophysics Data System (ADS)

    Welc, Fabian; Marks, Leszek

    2014-05-01

    ). A role of these wet intervals is incontestable and they can represent a sub-pluvial episode with two peaks at about 4600 and 4200 yrs BP, presumably of regional significance for north-eastern Africa. The authors suggest to name this relatively wet period the Saqqara Sub-pluvial. Based on the present knowledge, the extremely dynamic terminal phase of the second peak of the Saqqara Sub-pluvial and the following sudden desertification noted in deposits at archaeological sites in Saqqara and in the adjoining area (Abusir, Giza) should be roughly correlated with global climatic perturbation of the 3rd Bond Event or 4.2 ka Rapid Climate Change. The climate change in that time resulted in a collapse of the Egyptian civilization but also in rapid depopulation of Palestine, disintegration of the Acadian Empire in Mesopotamia and the Harappa Civilization in northwest India. Mechanism of this climate change is investigated within the project funded by National Science Centre in Poland (decision no. DEC-2012/05/B/ST10/00558).

  13. Bedrock geologic map of the Miles Pond and Concord quadrangles, Essex and Caledonia Counties, Vermont, and Grafton County, New Hampshire

    USGS Publications Warehouse

    Rankin, Douglas W.

    2018-04-20

    The bedrock geologic map of the Miles Pond and Concord quadrangles covers an area of approximately 107 square miles (276 square kilometers) in east-central Vermont and adjacent New Hampshire, north of and along the Connecticut River. This map was created as part of a larger effort to produce a new bedrock geologic map of Vermont through the collection of field data at a scale of 1:24,000. The majority of the map area consists of the Bronson Hill anticlinorium, a post-Early Devonian structure that is cored by metamorphosed Cambrian to Silurian sedimentary, volcanic, and plutonic rocks. A major feature on the map is the Monroe fault, interpreted to be a west-directed, steeply dipping Late Devonian (Acadian) thrust fault. To the west of the Monroe fault, rocks of the Connecticut Valley-Gaspé trough dominate and consist primarily of metamorphosed Silurian and Devonian sedimentary rocks. To the north, the Victory pluton intrudes the Bronson Hill anticlinorium. The Bronson Hill anticlinorium consists of the metamorphosed Albee Formation, the Ammonoosuc Volcanics, the Comerford Intrusive Complex, the Highlandcroft Granodiorite, and the Joselin Turn tonalite. The Albee Formation is an interlayered, feldspathic metasandstone and pelite that is locally sulfidic. Much of the deformed metasandstone is tectonically pinstriped. In places, one can see compositional layering that was transposed by a steeply southeast-dipping foliation. The Ammonoosuc Volcanics are lithologically complex and predominantly include interlayered and interfingered rhyolitic to basaltic volcanic and volcaniclastic rocks, as well as lesser amounts of siltstone, phyllite, graywacke, and grit. The Comerford Intrusive Complex crops out east of the Monroe fault and consists of metamorphosed gabbro, diorite, tonalite, aplitic tonalite, and crosscutting diabase dikes. Abundant mafic dikes from the Comerford Intrusive Complex intruded the Albee Formation and Ammonoosuc Volcanics east of the Monroe fault. The

  14. Geology of the Andover Granite and surrounding rocks, Massachusetts

    USGS Publications Warehouse

    Castle, Robert O.

    1964-01-01

    Field and petrographic studies of the Andover Granite and surrounding rocks have afforded an opportunity for an explanation of its emplacement and crystallization. The investigation has contributed secondarily to an understanding of the geologic history of southeastern New England, particularly as it is revealed in the Lawrence, Wilmington, South Groveland, and Reading quadrangles of Massachusetts. The Andover Granite and Sharpners Pond Tonalite together comprise up to 90 percent of the Acadian(?) subalkaline intrusive series cropping out within the area of study. The subalkaline series locally invades a sequence of early to middle Paleozoic and possibly Precambrian metasedimentary and metavolcanic rocks. Much of the subalkaline series and most of the Andover Granite is confined between two prominent east-northeast trending faults or fault systems. The northern fault separates the mildly metamorphosed Middle Silurian(?) Merrimack Group on the north from a highly metamorphosed and thoroughly intruded Ordovician(?) sequence on the south. The southern 'boundary '' fault is a major structural discontinuity characterized by penetrative, diffuse shearing over a zone one-half mile or more in width. The magmatic nature of the Andover Granite is demonstrated by: (1) sharply crosscutting relationships with surrounding rocks; (2) the occurrence of tabular-shaped xenoliths whose long directions parallel the foliation within the granite and whose internal foliation trends at a high angle to that of the granite; (3) continuity with the clearly intrusive Sharpners Pond Tonalite; (4) the compositional uniformity of the granite as contrasted with the compositional diversity of the rocks it invades; (5) its modal and normative correspondence with (a) calculated norms of salic extrusives and (b) that of the ternary (granite) minimum for the system NaAlSi3O8-KAlSi3O8-SiO2. Orogenic granites, as represented by the Andover, contrast with post-orogenic granites, represented locally by

  15. Dissolution Mediated Boron and Carbon Storage during Exhumation of HP Metapelites: Examples from New Hampshire Tourmaline-Graphite Intergrowths

    NASA Astrophysics Data System (ADS)

    Galvez, M.; Rumble, D.; Cody, G. D.; Sverjensky, D. A.

    2013-12-01

    The dynamic of light elements (e.g. C,B) in subduction zones is a complex process ultimately governed by variables such as P, T, fH2 and pH. Interface phenomena at scales from the outcrop to intergranular surfaces play key chemical and mechanical roles on this dynamic (e.g. Galvez et al. 2013). We report here a petrological study of hydrated borosilicate tourmaline intergrown with graphite formed at the contact between igneous intrusives and high grade micaschists in New Hampshire graphite deposits (Rumble and Hoering, 1986). Our study includes Raman scattering, SEM, microprobe analysis and thermodynamic modeling, focusing on the Franklin Pierce and Walpole outcrops. Both localities experienced HP-HT metamorphism during the Acadian orogeny as well as complex metasomatic process during exhumation. The tourmaline-graphite intergrowths are structurally localized at and around contacts between an aplite sill and micaschists - biotite-muscovite-garnet-sillimanite-plagioclase-quartz-ilmenite - (Franklin Pierce), or along shear zones (Walpole) in veins. Tourmalines are dravitic in composition (i.e. Na, Mg rich with minor vacancy and Li content 0.2/0.1 a.p.f.u) and contain multiple primary tubular mixed fluid-solid inclusions containing graphite, quartz and gaseous CO2 and CH4. Sharp optical and compositional radial zonations are observed from core to rim in sections along and perpendicular to the c-axis. Blue-green cores are enriched in Mg and Ca (1.5/0.1 a.p.f.u respectively) whereas rims are enriched in Fe, Na and Ti (0.9/0.6/0.1 a.p.f.u respectively). Alternative interpretations in terms of sector zoning or compositional variability of the mineralizing fluid will be discussed. The carbonaceous material (CM) occurs primarily as flakes directly replacing biotite present in wall rocks. The structural ordering of CM, of unambiguous abiotic origin, reveals a material possessing the 3 dimensional structure of hexagonal graphite. Our results are critically compared to

  16. Quality of water in domestic wells in the Chicot and Chicot equivalent aquifer systems, southern Louisiana and southwestern Mississippi, 2000-2001

    USGS Publications Warehouse

    Tollett, Roland W.; Fendick, Robert B.; Simmons, Lane B.

    2003-01-01

    In 2000-2001, water-quality data were collected from 60 randomly selected domestic wells in the Acadian-Pontchartrain Study Unit, as part of the National Water-Quality Assessment Program. The data were collected from wells screened in shallow sands (less than 350 feet below land surface) in two major aquifer systems--the Chicot aquifer system in southwestern Louisiana and the Chicot equivalent aquifer system in southeastern Louisiana and southwestern Mississippi. The Chicot equivalent aquifer system is part of the Southern Hills regional aquifer system, and both the Chicot aquifer system and the Southern Hills regional aquifer systems are designated as sole-source aquifers by the U.S. Environmental Protection Agency (USEPA). The well depths ranged from 40 to 340 feet below land surface with a median depth of 120 feet. The ground-water-quality data included 5 physiochemical properties, dissolved solids, 9 major inorganic ions, 24 trace elements, 6 nutrients, dissolved organic carbon, 109 pesticides and degradation products, and 85 volatile organic compounds (VOC's); and a subset of the wells were sampled for radon, chlorofluorocarbons, and stable isotopes. Water from 35 of the 60 domestic wells sampled had pH values less than the USEPA Seconday Maximum Contaminant Level (SMCL) range of 6.5 to 8.5 standard units. Specific conductance ranged from 17 to 1,420 microsiemens per centimeter at 25 degrees Celsius. Dissolved-solids concentrations in water from two wells exceeded the SMCL of 500 mg/L (milligrams per liter); the maximum concentration was 858 mg/L. Sodium and calcium were the dominant cations, and bicarbonate and chloride were the dominant anions. One chloride concentration (264 mg/L) exceeded the SMCL of 250 mg/L. One arsenic concentration (55.3 micrograms per liter) exceeded the USEPA Maximum Contaminant Level (MCL) of 10 micrograms per liter. Iron concentrations in water from 22 wells exceeded the SMCL of 300 micrograms per liter; the maximum concentration

  17. Correlation between the Palaeozoic structures from West Iberian and Grand Banks margins using inversion of magnetic anomalies

    NASA Astrophysics Data System (ADS)

    Silva, Elsa A.; Miranda, J. M.; Luis, J. F.; Galdeano, A.

    2000-05-01

    The Ibero-Armorican Arc (IAA) is a huge geological structure of Pre-Cambrian origin, tightened during hercynian times and deeply affected by the opening of the Atlantic Ocean and the Bay of Biscay. Its remnants now lie in Iberia, north-western France and the Canadian Grand Banks margins. The qualitative correlation between these three blocks has been attempted by several authors (e.g. Lefort, J.P., 1980. Un 'Fit' structural de l'Atlantique Nord: arguments geologiques pour correler les marqueurs geophysiques reconnus sur les deux marges. Mar. Geol. 37, 355-369; Lefort, J.P., 1983. A new geophysical criterion to correlate the Acadian and Hercynian orogenies of Western Europe and Eastern America. Mem. Geol. Soc. Am. 158, 3-18; Galdeano, A., Miranda, J.M., Matte, P., Mouge, P., Rossignol, C., 1990. Aeromagnetic data: A tool for studying the Variscan arc of Western Europe and its correlation with transatlantic structures. Tectonophysics 177, 293-305) using magnetic anomalies, mainly because they seem to preserve the hercynian zonation, in spite of the strong thermal and mechanical processes that took place during rifting and ocean spreading. In this paper, we present a new contribution to the study of the IAA structure based on the processing of a compilation of magnetic data from Iberia and Grand Banks margins. To interpret the magnetic signature, a Fourier-domain-based inversion technique was applied, considering a layer with a constant thickness of 10 km, and taking into account only the induced field. The digital terrain model was derived from ETOPO5 (ETOPO5, 1986. Relief map of the earth's surface. EOS 67, 121) and TerrainBase (TerrainBase, 1995. In: Row III, L.W., Hastings, D.A., Dunbar, P.K. (Eds.), Worldwide Digital Terrain Data, Documentation Manual, CD-ROM Release 1.0. GEODAS-NGDC Key to Geophysical Records. Documentation N. 30, April) databases. The pseudo-susceptibility distribution obtained was repositioned for the 156.5 Ma epoch, using the Srivastava and

  18. Quality of water from shallow wells in the rice-growing area in southwestern Louisiana, 1999 through 2001

    USGS Publications Warehouse

    Tollett, Roland W.; Fendick, Robert B.

    2004-01-01

    In 1999-2001, the U.S. Geological Survey installed and sampled 27 shallow wells in the rice-growing area in southwestern Louisiana as part of the Acadian-Pontchartrain Study Unit of the National Water-Quality Assessment Program. The purpose of this report is to describe the waulity of water from shallow wells in the rice-growing area and to relate that water quality to natural and anthropogenic activities, particularly rice agriculture. Ground-water samples were analyzed for general ground-water properties and about 150 water-quality constituents, including major inorganic ions, trace elements, nutrients, dissolved organic carbon (DOC), pesticides, radon, chloroflourocarbons, and selected stable isotopes. Dissolved solids concentrations for 17 wells exceeded the U.S. Environmental Protection Agency secondary minimum containment level of 500 milligrams per liter (mg/L) for drinking water. Concentrations for major pesticides generally were less than the maximum contaminant levels for drinking water. Two major inorganic ions, sulfate and chloride, and two trace elements, iron and manganese, had concentrations that were greater than the secondary maximum containment levels. Three nutrient concentrations were greater than 2 mg/L, a level that might indicate contamination from human activities, and one nutrient concentration (that for nitrite plus nitrite as nitrogen) was greater than the maximum contaminant level of 10 mg/L for drinking water. The median concentration for DOC was 0.5 mg/L, indicating naturally-occurring DOC conditions in the study area. Thirteen pesticides and 7 pesticide degradation products were detected in 14 of the 27 wells sampled. Bentazon, 2, 4-D, and molinate (three rice herbicides) were detected in water from four, one, and one wells, respectively, and malathion (a rice insecticide) was deteced in water fromone well. Low-level concentrations and few detections of nutrients and pesticides indicated that ground-water quality was affected slightly

  19. Stratigraphy, geochronology, and accretionary terrane settings of two Bronson Hill arc sequences, northern New England

    USGS Publications Warehouse

    Moench, R.H.; Aleinikoff, J.N.

    2002-01-01

    off the southern Laurentian margin, but northwest of the principal Iapetan suture, or Red Indian line (RIL). The Boil Mountain-Jim Pond-Hurricane Mountain sequence was ramped northwestward over the Chain Lakes massif at ???475 Ma, on the basal Boil Mountain surface. This obduction probably occurred slightly before obduction on the Baie Verte-Brompton surface (BBL), farther NW, over the Laurentian margin, and was followed by Dead River flysch sedimentation, which ended with the abrupt onset of Ammonoosuc-sequence arc magmatism at ???470 Ma. Ammonoosuc eruptions probably ended at ???460 Ma, when Iapetus closed along the Red Indian line. During a following magmatic hiatus of ???3-5 m.y., now represented by portions of the Partridge Formation that overlie the Ammonoosuc Volcanics, subduction polarity reversed, and subduction resumed below the northwest-dipping Brunswick subduction complex (BSC) of New Brunswick, Canada. Quimby-sequence magmatism (???456-435 Ma) on the the newly accreted Laurentian margin occurred above the BSC, whose footwall is now buried to the southeast by mainly Silurian clastic sediments of the Merrimack-Fredericton trough, deposited in the "Fredericton Sea". In Silurian to Early Devonian time, the NW-dipping BSC footwall was paired with a SE-dipping subduction zone that produced arc magmas of the Coastal Volcanic belt, built on the composite Avalon and adjacent peri-Avalonian terranes. Orogen-normal extension produced by rapid rollback of both subduction zones narrowed the Fredericton Sea, produced the Central Maine and Connecticut Valley-Gaspe?? basins, and culminated in the Acadian orogeny when the sea completely closed in Early Devonian time. Published by Elsevier Science Ltd.

  20. Stratigraphy, geochronology, and accretionary terrane settings of two Bronson Hill arc sequences, northern New England

    USGS Publications Warehouse

    Moench, R.H.; Aleinikoff, J.N.

    2003-01-01

    the southern Laurentian margin, but northwest of the principal Iapetan suture, or Red Indian line. The Boil Mountain-Jim Pond-Hurricane Mountain sequence was ramped northwestward over the Chain Lakes massif at ???475 Ma, on the basal Boil Mountain surface. This obduction probably occurred slightly before obduction on the Baie Vert-Brompton surface (BBL), farther NW, over the Laurentian margin, and was followed by Dead River flysch sedimentation, which ended with the abrupt onset of Ammonoosuc-sequence arc magmatism at ???470 Ma. Ammonoosuc eruptions probably ended at ???460 Ma, when Iapetus closed along the Red Indian line. During a following magmatic hiatus of ???3-5 m.y., now represented by portions of the Partridge Formation that overlie the Ammonoosuc Volcanics, subduction polarity reversed, and subduction resumed below the northwest-dipping Brunswick subduction complex (BSC) of New Brunswick, Canada. Quimby-sequence magmatism (???456-435 Ma) on the newly accreted Laurentian margin occurred above the BSC, whose footwall is now buried to the southeast by mainly Silurian clastic sediments of the Merrimack-Fredericton trough, deposited in the "Fredericton Sea". In Silurian to Early Devonian time, the NW-dipping BSC footwall was paired with a SE-dipping subduction zone that produced arc magmas of the Coastal Volcanic belt, built on the composite Avalon and adjacent peri-Avalonian terranes. Orogen-normal extension produced by rapid rollback of both subduction zones narrowed the Fredericton Sea, produced the Central Maine and Connecticut Valley-Gaspe?? basins, and culminated in the Acadian orogeny when the sea completely closed in Early Devonian time. Published by Elsevier Science Ltd.

  1. Geology of the Stroudsburg quadrangle and Adjacent areas, Pennsylvania--New Jersey

    USGS Publications Warehouse

    Epstein, Jack Burton

    1971-01-01

    The Stroudsburg area is within the Valley and Ridge and Great Valley physiographic provinces, Northampton and Monroe Counties, Pennsylvania, and Warren County, New Jersey. The northeast-trending subparallel valleys and ridges resulted from erosion of folded heterogeneous sedimentary rocks. These are Middle Ordovician to Middle Devonian in age and are more than 17,000 feet thick. Deposition of a thick flysch sequence (Martinsburg Formation of Ordovician age) accompanied onset of Taconic orogenesis. It was followed by deposition of a thick molasse sequence of Silurian and Early Devonian age (continental and marginal-marine clastics--Shawangunk Formation and Bloomsburg Red Beds--overlain by predominantly marginal-marine and subtidal limestone, dolomite, shale, and sandstone--Poxono Island Formation through Oriskany Group). Basin deepening and gradual shallowing occurred during Esopus through Mahantango deposition, heralding the Acadian clastic wedge exposed north of the Stroudsburg area. Interpretation of sedimentary structures and regional stratigraphic relations suggest that the Silurian and Devonian rocks were deposited in the following environments: A1luviated coastal plain (meandering and braided streams), tidal flats (supratidal and intertidal), barrier zone, and neritic zone (upper and lower). The rock stratigraphic units have been grouped into four lithotectonic units, each having a different style of deformation. Folds produced in these rocks are disharmonic, and it is believed that each rock sequence is set off from units above and below by decollements, or zones of detachment. Movement was northwest into the Appalachian basin, primarily by gravitational sliding. The contact between the Shawangunk Formation of Silurian age and Martinsburg Formation of Ordovician age, is one zone of detachment as well as an angular unconformity. Deformational effects of the Middle to Late Ordovician Taconic orogeny are elusive, but it appears that the folds and most minor

  2. Stratigraphy, geochronology, and accretionary terrane settings of two Bronson Hill arc sequences, northern New England

    NASA Astrophysics Data System (ADS)

    Moench, Robert H.; Aleinikoff, John N.

    the southern Laurentian margin, but northwest of the principal Iapetan suture, or Red Indian line. The Boil Mountain-Jim Pond-Hurricane Mountain sequence was ramped northwestward over the Chain Lakes massif at ∼475 Ma, on the basal Boil Mountain surface. This obduction probably occurred slightly before obduction on the Baie Vert-Brompton surface (BBL), farther NW, over the Laurentian margin, and was followed by Dead River flysch sedimentation, which ended with the abrupt onset of Ammonoosuc-sequence arc magmatism at ∼470 Ma. Ammonoosuc eruptions probably ended at ∼460 Ma, when Iapetus closed along the Red Indian line. During a following magmatic hiatus of ∼3-5 m.y., now represented by portions of the Partridge Formation that overlie the Ammonoosuc Volcanics, subduction polarity reversed, and subduction resumed below the northwest-dipping Brunswick subduction complex (BSC) of New Brunswick, Canada. Quimby-sequence magmatism (∼456-435 Ma) on the newly accreted Laurentian margin occurred above the BSC, whose footwall is now buried to the southeast by mainly Silurian clastic sediments of the Merrimack-Fredericton trough, deposited in the “Fredericton Sea”. In Silurian to Early Devonian time, the NW-dipping BSC footwall was paired with a SE-dipping subduction zone that produced arc magmas of the Coastal Volcanic belt, built on the composite Avalon and adjacent peri-Avalonian terranes. Orogen-normal extension produced by rapid rollback of both subduction zones narrowed the Fredericton Sea, produced the Central Maine and Connecticut Valley-Gaspé basins, and culminated in the Acadian orogeny when the sea completely closed in Early Devonian time.

  3. Comparative geomorphic analysis of surficial deposits at three central Appalachian watersheds: Implications for controls on sediment-transport efficiency

    NASA Astrophysics Data System (ADS)

    Taylor, Stephen B.; Steven Kite, J.

    2006-08-01

    Factors that control the routing and storage of sediments in the Appalachian region are poorly understood. This study involves a comparative geomorphic analysis of three watersheds underlain by sandstones and shales of the Acadian clastic wedge. These areas include the Fernow Experimental Forest, Tucker County, West Virginia; the North Fork basin, Pocahontas County, West Virginia; and the Little River basin, Augusta County, Virginia. GIS-based analyses of surficial map units allow first-order approximation of sediment-storage volumes in valley bottoms. Estimates of volumes are examined in tandem with morphometric analyses and the distribution of bedrock channels to make inferences regarding controls on sediment-transport efficiency in the central Appalachians. The Fernow and North Fork areas are characterized by V-shaped valleys with mixed reaches of alluvial-bedrock channels distributed throughout the drainage network. In contrast, the Little River valley is notably wider and gravelly alluvial fill is abundant. Comparator watershed parameters for the Fernow, North Fork and Little River areas include, respectively: (1) basin area = 15.2 km 2, 49.3 km 2, 41.5 km 2; (2) basin relief = 0.586 km, 0.533 km, 0.828 km; (3) drainage density = 4.2 km - 1 , 3.3 km - 1 , 4.7 km - 1 ; (4) ruggedness = 2.5, 1.7, 3.9; (5) Shreve magnitude = 139, 287, 380; (6) total valley-bottom area (km 2) = 0.76 km 2, 1.86 km 2, 3.09 km 2; (7) average hillslope gradients = 17.2°, 18.4°, 22.1°; (8) total debris-fan surface area = 0.113 km 2, 0.165 km 2, 0.486 km 2; and (9) debris-fan frequency = 2.0 km - 2 , 1.0 km - 2 , 2.8 km - 2 . The storage volumes in valley bottoms were estimated using map polygon areas and surface heights above channel grade. The Little River contains significantly higher sediment volumes in floodplain, terrace and fan storage compartments; total volumes of the valley bottoms are approximately twice that of the Fernow and North Fork areas combined. Unit storage

  4. Stratigraphy, geochronology, and accretionary terrane settings of two Bronson Hill arc sequences, northern New England

    NASA Astrophysics Data System (ADS)

    Moench, Robert H.; Aleinikoff, John N.

    2002-01-01

    southern Laurentian margin, but northwest of the principal Iapetan suture, or Red Indian line (RIL). The Boil Mountain-Jim Pond-Hurricane Mountain sequence was ramped northwestward over the Chain Lakes massif at ∼475 Ma, on the basal Boil Mountain surface. This obduction probably occurred slightly before obduction on the Baie Verte-Brompton surface (BBL), farther NW, over the Laurentian margin, and was followed by Dead River flysch sedimentation, which ended with the abrupt onset of Ammonoosuc-sequence arc magmatism at ∼470 Ma. Ammonoosuc eruptions probably ended at ∼460 Ma, when Iapetus closed along the Red Indian line. During a following magmatic hiatus of ∼3-5 m.y., now represented by portions of the Partridge Formation that overlie the Ammonoosuc Volcanics, subduction polarity reversed, and subduction resumed below the northwest-dipping Brunswick subduction complex (BSC) of New Brunswick, Canada. Quimby-sequence magmatism (∼456-435 Ma) on the the newly accreted Laurentian margin occurred above the BSC, whose footwall is now buried to the southeast by mainly Silurian clastic sediments of the Merrimack-Fredericton trough, deposited in the “Fredericton Sea”. In Silurian to Early Devonian time, the NW-dipping BSC footwall was paired with a SE-dipping subduction zone that produced arc magmas of the Coastal Volcanic belt, built on the composite Avalon and adjacent peri-Avalonian terranes. Orogen-normal extension produced by rapid rollback of both subduction zones narrowed the Fredericton Sea, produced the Central Maine and Connecticut Valley-Gaspé basins, and culminated in the Acadian orogeny when the sea completely closed in Early Devonian time.

  5. Vascular Plant and Vertebrate Inventory of Tumacacori National Historical Park

    USGS Publications Warehouse

    Powell, Brian F.; Albrecht, Eric W.; Halvorson, William L.; Schmidt, Cecilia A.; Anning, Pamela; Docherty, Kathleen

    2005-01-01

    Executive Summary This report summarizes the results of the first comprehensive biological inventory of Tumacacori National Historical Park (NHP) in southern Arizona. These surveys were part of a larger effort to inventory vascular plants and vertebrates in eight National Park Service units in Arizona and New Mexico. From 2000 to 2003 we surveyed for vascular plants and vertebrates (fish, amphibians, reptiles, birds, and mammals) at Tumacacori NHP to document presence of species within the administrative boundaries of the park's three units. Because we used repeatable study designs and standardized field techniques, these inventories can serve as the first step in a long-term monitoring program. We recorded 591 species at Tumacacori NHP, significantly increasing the number of known species for the park (Table 1). Species of note in each taxonomic group include: * Plants: second record in Arizona of muster John Henry, a non-native species that is ranked a 'Class A noxious weed' in California; * Amphibian: Great Plains narrow-mouthed toad; * Reptiles: eastern fence lizard and Sonoran mud turtle; * Birds: yellow-billed cuckoo, green kingfisher, and one observation of the endangered southwestern willow flycatcher; * Fishes: four native species including an important population of the endangered Gila topminnow in the Tumacacori Channel; * Mammals: black bear and all four species of skunk known to occur in Arizona. We recorded 79 non-native species (Table E.S.1), many of which are of management concern, including: Bermudagrass, tamarisk, western mosquitofish, largemouth bass, bluegill, sunfish, American bullfrog, feral cats and dogs, and cattle. We also noted an abundance of crayfish (a non-native invertebrate). We review some of the important non-native species and make recommendations to remove them or to minimize their impacts on the native biota of the park. Based on the observed species richness, Tumacacori NHP possesses high biological diversity of plants, fish

  6. Final integrated trip report: site visits to Area 50, Andersen Air Force Base, Guam National Wildlife Refuge, War in the Pacific National Historical Park, Guam, Rota and Saipan, CNMI, 2004-2005

    USGS Publications Warehouse

    Hess, Steven C.; Pratt, Linda W.

    2006-01-01

    Limestone forests are the most diverse natural plant communities of Guam. Like other natural vegetation types, these forests have a long history of anthropogenic disturbances, being altered and shaped by humans for more than 4,000 years. Although this occupation represents a relatively long human influence in comparison to other Pacific islands, animals associated with humans, such as commensal rodents, arrived in these islands beginning only 1,000 years ago, and larger mammals, such as pigs (Sus scrofa), may not have arrived until European contact. Limestone forests, which also occur on several other Mariana Islands, developed in the presence of frequent tropical storms and are therefore well adapted to this type of natural disturbance regime. However, recent human activities including large scale clearing and conversion combined with the presence of high levels of alien herbivores and seed predators, and the loss of ecological services provided by the former native avifauna may be causing the decline of Guam's forests. Limestone forests on northern Guam, much like those of other Mariana Islands, were heavily cleared for the construction of military installations during World War II. The accidental introduction of the Brown Tree Snake (Boiga irregularis; BTS) around this same period subsequently accelerated the disappearance of Guam's native avifauna and other endemic terrestrial vertebrates, and with them, seed dispersal, pollination, and the predatory regulation of herbivorous insects. Guam and the Mariana Islands contained a high proportion (32 pecent) of endemic bird species, with 4 forms endemic to Guam alone: the now extinct Guam Flycatcher (Myiagra freycineti), and Guam Bridled White-eye (Zosterops conspicillatpicillata), one of three island endemic subspecies from the Marianas; Guam rail (Rallus owstonii); and Guam Kingfisher (Todiramphus cinnamominus cinnamominus), an island endemic subspecies of the regionally endemic Micronesian Kingfisher. Guam once

  7. Assessment and management of dead-wood habitat

    USGS Publications Warehouse

    Hagar, Joan

    2007-01-01

    Oregon and Washington, approximately 150 species of wildlife are reported to use dead wood in forests (O’Neil et al., 2001). Forty-seven sensitive and special-status species are associated with dead wood (Appendix A). These are key species for management consideration because concern over small or declining populations is often related to loss of suitable dead-wood habitat (Marshall et al., 1996). Primary excavators (woodpeckers) also are often the focus of dead-wood management, because they perform keystone functions in forest ecosystems by creating cavities for secondary cavity-nesters (Martin and Eadie, 1999; Aubry and Raley, 2002). A diverse guild of secondary cavity-users (including swallows, bluebirds, several species of ducks and owls, ash-throated flycatcher, flying squirrel, bats, and many other species) is unable to excavate dead wood, and therefore relies on cavities created by woodpeckers for nesting sites. Suitable nest cavities are essential for reproduction, and their availability limits population size (Newton, 1994). Thus, populations of secondary cavity-nesters are tightly linked to the habitat requirements of primary excavators. Although managers often focus on decaying wood as habitat for wildlife, the integral role dead wood plays in ecological processes is an equally important consideration for management. Rose et al. (2001) provide a thorough review of the ecological functions of dead wood in Pacific Northwest forests, briefly summarized here. Decaying wood functions in: soil development and productivity, nutrient cycling, nitrogen fixation, and carbon storage. From ridge tops, to headwater streams, to estuaries and coastal marine ecosystems, decaying wood is fundamental to diverse terrestrial and aquatic food webs. Wildlife species that use dead wood for cover or feeding are linked to these ecosystem processes through a broad array of functional roles, including facilitation of decay and trophic interactions with other organisms (Marcot, 2002