Etiology of bronze leaf disease of Populus
Jason A. Smith; R. A. Blanchette; M. E. Ostry; N. A. Anderson
2002-01-01
Bronze leaf disease is a potentially destructive disorder of the Populus section of the genus Populus. The causal agent has been reported to be Apioplagiostoma populi (anarnorph: Discula sp.). Based on etiological and symptomological studies, field observations of symptom development suggest that the pathogen...
How Can We Best Achieve Contracting Unity of Effort in the CENTCOM Area of Responsibility?
2013-11-20
Major Torres’ military decorations include the Bronze Star Medal, the Army Meritorious Medal with oak leaf (OLC), the Army Commendation Medal...1990–May 1991). Major Ross’ military decorations include the Bronze Star Medal, the Army Meritorious Medal with two oak leaf (OLC), the Army...Berteau, D. J., Maddox, D. M., Oliver , D. R., Jr., Salomon, L. E., & Singley, G. T., III. (2007). Urgent reform required: Army expeditionary contracting
USDA-ARS?s Scientific Manuscript database
Landscape irrigation is the second largest user of reclaimed water in industrialized countries; however its high concentration of soluble salts, especially Na+ and Cl-, may induce growth reduction and leaf necrosis or bronzing in ornamental species. The present study was conducted to determine the g...
Portable X-ray diffractometer equipped with XRF for archaeometry
NASA Astrophysics Data System (ADS)
Uda, M.; Ishizaki, A.; Satoh, R.; Okada, K.; Nakajima, Y.; Yamashita, D.; Ohashi, K.; Sakuraba, Y.; Shimono, A.; Kojima, D.
2005-09-01
A portable X-ray diffractometer equipped with an X-ray fluorescence spectrometer was improved so as to get a diffraction pattern and a fluorescence spectrum simultaneously in air from one and the same small area on a specimen. Here, diffraction experiments were performed in two modes, i.e. an angle rotation mode and an energy dispersive mode. In the latter a diffraction pattern and a fluorescence spectrum were simultaneously recorded in a short time, 100 s or less, on one display. The diffractometer was tested in the field to confirm its performance. Targets chosen for this purpose were a bronze mirror from the Eastern Han Dynasty (25-220), and a stupa and its pedestal which are part of the painted statue of "Tamonten holding a stupa" from the Heian Period (794-1192), enshrined in the Engyouji temple founded in 996. The bronze mirror was identified as a product of the Han Dynasty from its chemical composition and the existence of the δ phase in the Cu-Sn alloy. The stupa and its pedestal were decorated with gold powder and gold leaf, respectively. From the XRF data of the pedestal, the underlying layer of gold leaf seems to have been painted with emerald green.
Sampson, Clare; Kirk, William D. J.
2013-01-01
The western flower thrips Frankliniella occidentalis (Pergande) (Thysanoptera: Thripidae) is a cosmopolitan, polyphagous insect pest that causes bronzing to fruit of strawberry (Fragaria x ananassa). The main aim of this study was to test whether mass trapping could reduce damage and to predict whether this approach would be economically viable. In semi-protected strawberry crops, mass trapping of F. occidentalis using blue sticky roller traps reduced adult thrips numbers per flower by 61% and fruit bronzing by 55%. The addition of the F. occidentalis aggregation pheromone, neryl (S)-2-methylbutanoate, to the traps doubled the trap catch, reduced adult thrips numbers per flower by 73% and fruit bronzing by 68%. The factors affecting trapping efficiency through the season are discussed. Damage that would result in downgrading of fruit to a cheaper price occurred when bronzing affected about 10% of the red fruit surface. Cost-benefit analysis using this threshold showed that mass trapping of thrips using blue sticky roller traps can be cost-effective in high-value crops. The addition of blue sticky roller traps to an integrated pest management programme maintained thrips numbers below the damage threshold and increased grower returns by a conservative estimate of £2.2k per hectare. Further work is required to develop the F. occidentalis aggregation pheromone for mass trapping and to determine the best timing for trap deployment. Mass trapping of thrips is likely to be cost-effective in other countries and other high-value crops affected by F. occidentalis damage, such as cucumber and cut flowers. PMID:24282554
Electronic band structure and charge density wave transition in quasi-2D KMo6O17 purple bronze
NASA Astrophysics Data System (ADS)
Valbuena, M. A.; Avila, J.; Vyalikh, D. V.; Guyot, H.; Laubschat, C.; Molodtsov, S. L.; Asensio, M. C.
2008-03-01
High resolution angle-resolved photoemission of quasi-2D KMo6O17 purple bronze has been performed in the range from room temperature to 130 K, slightly above the charge density wave (CDW) transition (Tc = 110 K), and down to 35 K (well below Tc). In this paper we report a detailed study of how electronic band structure is affected by this transition driven by the hidden nesting scenario. The expected spectroscopic fingerprints of the CDW phase transition have been found and discussed according to the hidden one dimension and the development of a quasi-commensurate CDW. The excellent agreement between theory and our experimental results makes of potassium purple bronze a reference system for studying this type of instabilities.
NASA Astrophysics Data System (ADS)
Masi, G.; Chiavari, C.; Avila, J.; Esvan, J.; Raffo, S.; Bignozzi, M. C.; Asensio, M. C.; Robbiola, L.; Martini, C.
2016-03-01
Gilded bronzes are often affected by severe corrosion, due to defects in the Au layer and Au/Cu alloy galvanic coupling, stimulated by large cathodic area of the gilded layer. Galvanic corrosion, triggered by gilding defects, leads to products growth at the Au/bronze interface, inducing blistering or break-up of the Au layer. In this context, fire-gilded bronze replicas prepared by ancient methods (use of spreadable Au-Hg paste) was specifically characterised by compiling complementary spectroscopic and imaging information before/after accelerated ageing with synthetic rain. Fire-gilded bronze samples were chemically imaged in cross-section at nano-metric scale (<200 nm) using high energy and lateral resolution synchrotron radiation photoemission (HR-SRPES) of core levels and valence band after conventional characterisation of the samples by Glow Discharge optical Emission Spectroscopy (GD-OES) and conventional X-ray photoelectron spectroscopy (XPS). We have found a net surface enrichment in Zn and Sn after fire-gilding and presence of metallic Hg, Pb and Cu within the Au layer. Moreover, the composition distribution of the elements together with their oxidation has been determined. It was also revealed that metallic phases including Hg and Pb remain in the gilding after corrosion. Moreover, selective dissolution of Zn and Cu occurs in the crater due to galvanic coupling, which locally induces relative Sn species enrichment (decuprification). The feasibility advantages and disadvantages of chemical imaging using HR-SRPES to study artworks have been investigated on representative replicas.
NASA Astrophysics Data System (ADS)
de Luna, Martina Salzano; Buonocore, Giovanna; Di Carlo, Gabriella; Giuliani, Chiara; Ingo, Gabriel M.; Lavorgna, Marino
2016-05-01
Protective coatings based on polymers synthesized from renewable sources (chitosan or an amorphous vinyl alcohol based polymer) have been prepared for the protection of bronze artifacts from corrosion. Besides acting as an effective barrier against corrosive species present in the environment, the efficiency of the coatings has been improved by adding corrosion inhibitor compounds (benzotriazole or mercaptobenzothiazole) to the formulations. The liquid medium of the formulations has been carefully selected looking at maximizing the wettability on the bronze substrate and optimizing the solvent evaporation rate. The minimum amount of inhibitor compounds has been optimized by performing accelerated corrosion tests on coated bronze substrates. The inhibitors have been directly dissolved in the coating-forming solutions and/or introduced by means of nanocarriers, which allow to control the release kinetics. The free dissolved inhibitor molecules immediately provide a sufficient protection against corrosion. On the other hand, the inhibitor molecules contained in the nanocarriers serve as long-term reservoir, which can be activated by external corrosion-related stimuli in case of particularly severe conditions. Particular attention has been paid to other features which affect the coating performances. Specifically, the adhesion of the protective polymer layer to the bronze substrate has been assessed, as well as its permeability properties and transparency, the latter being a fundamental feature of protective coating for cultural heritages. Finally, the protective efficiency of the produced smart coatings has been assessed through accelerated corrosion tests.
Origin of the mysterious Yin-Shang bronzes in China indicated by lead isotopes.
Sun, Wei-dong; Zhang, Li-peng; Guo, Jia; Li, Cong-ying; Jiang, Yu-hang; Zartman, Robert E; Zhang, Zhao-feng
2016-03-18
Fine Yin-Shang bronzes containing lead with puzzlingly highly radiogenic isotopic compositions appeared suddenly in the alluvial plain of the Yellow River around 1400 BC. The Tongkuangyu copper deposit in central China is known to have lead isotopic compositions even more radiogenic and scattered than those of the Yin-Shang bronzes. Most of the Yin-Shang bronzes are tin-copper alloys with high lead contents. The low lead and tin concentrations, together with the less radiogenic lead isotopes of bronzes in an ancient smelting site nearby, however, exclude Tongkuangyu as the sole supplier of the Yin-Shang bronzes. Interestingly, tin ingots/prills and bronzes found in Africa also have highly radiogenic lead isotopes, but it remains mysterious as to how such African bronzes may have been transported to China. Nevertheless, these African bronzes are the only bronzes outside China so far reported that have lead isotopes similar to those of the Yin-Shang bronzes. All these radiogenic lead isotopes plot along ~2.0-2.5 Ga isochron lines, implying that deposits around Archean cratons are the most likely candidates for the sources. African cratons along the Nile and even micro-cratons in the Sahara desert may have similar lead signatures. These places were probably accessible by ancient civilizations, and thus are the most favorable suppliers of the bronzes.
75 FR 14257 - Pricing for Bronze Medals
Federal Register 2010, 2011, 2012, 2013, 2014
2010-03-24
... DEPARTMENT OF THE TREASURY United States Mint Pricing for Bronze Medals AGENCY: United States Mint, Department of the Treasury. ACTION: Notice. SUMMARY: The United States Mint is announcing the price of the 1\\5/16\\- inch bronze medals, 1\\1/2\\-inch bronze medals and three-inch bronze medals. Beginning March...
Origin of the mysterious Yin-Shang bronzes in China indicated by lead isotopes
Sun, Wei-dong; Zhang, Li-peng; Guo, Jia; Li, Cong-ying; Jiang, Yu-hang; Zartman, Robert E.; Zhang, Zhao-feng
2016-01-01
Fine Yin-Shang bronzes containing lead with puzzlingly highly radiogenic isotopic compositions appeared suddenly in the alluvial plain of the Yellow River around 1400 BC. The Tongkuangyu copper deposit in central China is known to have lead isotopic compositions even more radiogenic and scattered than those of the Yin-Shang bronzes. Most of the Yin-Shang bronzes are tin-copper alloys with high lead contents. The low lead and tin concentrations, together with the less radiogenic lead isotopes of bronzes in an ancient smelting site nearby, however, exclude Tongkuangyu as the sole supplier of the Yin-Shang bronzes. Interestingly, tin ingots/prills and bronzes found in Africa also have highly radiogenic lead isotopes, but it remains mysterious as to how such African bronzes may have been transported to China. Nevertheless, these African bronzes are the only bronzes outside China so far reported that have lead isotopes similar to those of the Yin-Shang bronzes. All these radiogenic lead isotopes plot along ~2.0–2.5 Ga isochron lines, implying that deposits around Archean cratons are the most likely candidates for the sources. African cratons along the Nile and even micro-cratons in the Sahara desert may have similar lead signatures. These places were probably accessible by ancient civilizations, and thus are the most favorable suppliers of the bronzes. PMID:26988425
46 CFR 160.044-3 - General requirements.
Code of Federal Regulations, 2014 CFR
2014-10-01
... free from any defects affecting serviceability. Where a choice of materials is permitted, the materials... pump size No. 3. Discharge outlets shall be provided with a tee of cast bronze or other corrosion...
46 CFR 160.044-3 - General requirements.
Code of Federal Regulations, 2013 CFR
2013-10-01
... free from any defects affecting serviceability. Where a choice of materials is permitted, the materials... pump size No. 3. Discharge outlets shall be provided with a tee of cast bronze or other corrosion...
46 CFR 160.044-3 - General requirements.
Code of Federal Regulations, 2012 CFR
2012-10-01
... free from any defects affecting serviceability. Where a choice of materials is permitted, the materials... pump size No. 3. Discharge outlets shall be provided with a tee of cast bronze or other corrosion...
NASA Astrophysics Data System (ADS)
Alakkam, Abdulla Ahmed
The enamels of one hundred and ninety two teeth were analyzed for their oxygen isotopic composition from five archaeological sites in northern Jordan (Ya'amoun, Pella, Yasieleh, Sa'ad and Waqqas) to reconstruct the climate from the Middle Bronze Age II (1800--1500 BCE) to the Late Byzantine Period (491--640 CE). Results showed that the Middle Bronze Age II delta 18O values were the highest indicating warmer and drier climate conditions. The Late Bronze Age II and the Iron Age I/II experienced more favorable climate conditions. The Early Byzantine period delta18O values indicate wetter conditions compared to the Late Byzantine period but the latter was wetter than the Middle Bronze Age II. The geographic distribution of the archaeological sites in Jordan was affected by climate that changed over the prehistory of the country (from the Paleolithic to the Ottoman Period). Archaeological sites decreased in number during dry periods like the Middle Bronze Age II but increased during wet periods like the Roman one. Proximity to water sources was favored during the various cultural periods, where the northwestern and the mid-western areas witnessed dense occupation. On the other hand, areas that presently receive less than 200mm annual rainfall were not favored even during the wet periods. There were major climate events that overran the whole Mediterranean region during the Holocene, where Jordan was subject to them as well. These climatic events (prolonged droughts and/or insufficient precipitation) were identified using Bryson's archaeoclimate model that simulates the amount of precipitation, evapotranspiration and temperature during the last 12,000 years. These climate events coincided with the collapse of many societies in the Levant, such as the Middle Bronze Age societies of Mesopotamian cities.
Corrosion on prehistoric Cu-Sn-alloys: the influence of artificial environment and storage
NASA Astrophysics Data System (ADS)
Mödlinger, Marianne; Piccardo, Paolo
2013-12-01
The paper contributes to the identification of different corrosion products detected on the cross-section specimens sampled from Bronze Age swords and one helmet found between 60-160 years ago. The objects are kept in 1889 built oak showcases at the Natural History Museum Vienna, having suffered unknown restoration treatments. The identified corrosion products not only affect further eventual treatment in conservation science of copper base objects but also contribute to identify the often unknown find context, which is meant to facilitate archaeological interpretation of the Bronze Age weapons. The analyses of the samples were carried out using SEM-EDXS-EBSD and optical microscopy.
NASA Astrophysics Data System (ADS)
Masi, G.; Balbo, A.; Esvan, J.; Monticelli, C.; Avila, J.; Robbiola, L.; Bernardi, E.; Bignozzi, M. C.; Asensio, M. C.; Martini, C.; Chiavari, C.
2018-03-01
Application of a protective coating is the most widely used conservation treatment for outdoor bronzes (cast Cu-Sn-Zn-Pb-Sb alloys). However, improving coating protectiveness requires detailed knowledge of the coating/substrate chemical bonding. This is particularly the case for 3-mercapto-propyl-trimethoxy-silane (PropS-SH) applied on bronze, exhibiting a good protective behaviour in outdoor simulated conditions. The present work deals with X-Ray Photoelectron Spectroscopy (XPS) and Electron Microscopy (FEG-SEM + FIB (Focused Ion Beam)) characterization of a thin PropS-SH film on bronze. In particular, in order to better understand the influence of alloying elements on coating performance, PropS-SH was studied first on pure Cu and Sn substrates then on bronzes with increasing alloy additions: Cu8Sn as well as a quinary Cu-Sn-Zn-Pb-Sb bronze. Moreover, considering the real application of this coating on historical bronze substrates, previously artificially aged ("patinated") bronze samples were prepared and a comparison between bare and "patinated" quinary bronzes was performed. In the case of coated quinary bronze, the free surface of samples was analysed by High Resolution Photoelectron Spectroscopy using Synchrotron Radiation (HR-SRPES) at ANTARES (Synchrotron SOLEIL), which offers a higher energy and lateral resolution. By compiling complementary spectroscopic and imaging information, a deeper insight into the interactions between the protective coating and the bronze substrate was achieved.
Knipper, Corina; Friedrich, Ronny; Kromer, Bernd; Lindauer, Susanne; Radosavljević, Jelena; Wittenborn, Fabian; Krause, Johannes
2015-01-01
The transition from the Neolithic to the Early Bronze Age in Central Europe has often been considered as a supra-regional uniform process, which led to the growing mastery of the new bronze technology. Since the 1920s, archaeologists have divided the Early Bronze Age into two chronological phases (Bronze A1 and A2), which were also seen as stages of technical progress. On the basis of the early radiocarbon dates from the cemetery of Singen, southern Germany, the beginning of the Early Bronze Age in Central Europe was originally dated around 2300/2200 BC and the transition to more complex casting techniques (i.e., Bronze A2) around 2000 BC. On the basis of 140 newly radiocarbon dated human remains from Final Neolithic, Early and Middle Bronze Age cemeteries south of Augsburg (Bavaria) and a re-dating of ten graves from the cemetery of Singen, we propose a significantly different dating range, which forces us to re-think the traditional relative and absolute chronologies as well as the narrative of technical development. We are now able to date the beginning of the Early Bronze Age to around 2150 BC and its end to around 1700 BC. Moreover, there is no transition between Bronze (Bz) A1 and Bronze (Bz) A2, but a complete overlap between the type objects of the two phases from 1900–1700 BC. We thus present a revised chronology of the assumed diagnostic type objects of the Early Bronze Age and recommend a radiocarbon-based view on the development of the material culture. Finally, we propose that the traditional phases Bz A1 and Bz A2 do not represent a chronological sequence, but regionally different social phenomena connected to the willingness of local actors to appropriate the new bronze technology. PMID:26488413
Code of Federal Regulations, 2012 CFR
2012-10-01
...-basic steel, aluminum, brass, bronze, or copper mill products-representation. 252.216-7007 Section 252....216-7007 Economic price adjustment—basic steel, aluminum, brass, bronze, or copper mill products... Steel, Aluminum, Brass, Bronze, or Copper Mill Products—Representation (MAR 2012) (a) Definitions. The...
Code of Federal Regulations, 2013 CFR
2013-10-01
...-basic steel, aluminum, brass, bronze, or copper mill products-representation. 252.216-7007 Section 252....216-7007 Economic price adjustment—basic steel, aluminum, brass, bronze, or copper mill products... Steel, Aluminum, Brass, Bronze, or Copper Mill Products—Representation (MAR 2012) (a) Definitions. The...
Code of Federal Regulations, 2014 CFR
2014-10-01
...-basic steel, aluminum, brass, bronze, or copper mill products-representation. 252.216-7007 Section 252....216-7007 Economic price adjustment—basic steel, aluminum, brass, bronze, or copper mill products... Steel, Aluminum, Brass, Bronze, or Copper Mill Products—Representation (MAR 2012) (a) Definitions. The...
NASA Astrophysics Data System (ADS)
Nerantzis, Nerantzis
The compositions of copper-base tools, weapons, ornaments and ceremonial metalwork from numerous Late Bronze Age Aegean sites reveal a pattern of specific alloy combinations for the fabrication of certain classes of objects. Thus the majority of weapons and tools were made of high tin bronze whereas bronze statuettes, tripods and cauldrons contain small amounts of lead and in some cases tin is present in low amounts. Such diversity reflects the direct relationship between the compositions of prehistoric bronze objects and the art of their fabrication, because both the alloy additions and the impurities exert a pronounced effect on the forming capacity of alloys. In order to understand the correlation between composition and formability of Minoan bronzes, replica compositions with varying tin and lead contents were experimentally reproduced and their forming capacities were tested. Deformation and heat treatment of five tin and two leaded tin bronze alloys was attempted in order to replicate the forming stages for the shaping of cutting tools and bronze sheet for vessels and cauldrons. The amount of cold-working and annealing intervals, required to test the effects of workability on alloy properties, has been reflected as hardness values and transformations of the structural characteristics for each sample. It has been shown through the course of the experiment that high tin bronzes could be formed by frequent, short annealing stages at 600˚C and that it is possible to work-harden leaded bronze as long as time and temperatures are closely monitored.
75 FR 13345 - Pricing for Certain United States Mint Products
Federal Register 2010, 2011, 2012, 2013, 2014
2010-03-19
... is announcing the price of First Spouse Bronze Medals and 2010 First Spouse Bronze Medal Series: Four... currently on sale, effective March 18, 2010. The 2010 First Spouse Bronze Medal Series: Four-Medal Set will be priced at $15.95 and will contain all four 2010 First Spouse Bronze Medals. The release date for...
Federal Register 2010, 2011, 2012, 2013, 2014
2012-03-12
... DEPARTMENT OF THE TREASURY United States Mint Pricing for 2012 Kennedy Half-Dollar Bags and Rolls, Bronze Medals, the First Spouse Bronze Medal Set and the Birth Set AGENCY: United States Mint, Department of the Treasury. ACTION: Notice. SUMMARY: The United States Mint is announcing 2012 pricing for...
Code of Federal Regulations, 2014 CFR
2014-10-01
...-basic steel, aluminum, brass, bronze, or copper mill products. 252.216-7000 Section 252.216-7000 Federal... adjustment—basic steel, aluminum, brass, bronze, or copper mill products. As prescribed in 216.203-4-70(a)(1), use the following clause: Economic Price Adjustment—Basic Steel, Aluminum, Brass, Bronze, or Copper...
Code of Federal Regulations, 2013 CFR
2013-10-01
...-basic steel, aluminum, brass, bronze, or copper mill products. 252.216-7000 Section 252.216-7000 Federal... adjustment—basic steel, aluminum, brass, bronze, or copper mill products. As prescribed in 216.203-4-70(a)(1), use the following clause: Economic Price Adjustment—Basic Steel, Aluminum, Brass, Bronze, or Copper...
Code of Federal Regulations, 2012 CFR
2012-10-01
...-basic steel, aluminum, brass, bronze, or copper mill products. 252.216-7000 Section 252.216-7000 Federal... adjustment—basic steel, aluminum, brass, bronze, or copper mill products. As prescribed in 216.203-4-70(a)(1), use the following clause: Economic Price Adjustment—Basic Steel, Aluminum, Brass, Bronze, or Copper...
Code of Federal Regulations, 2011 CFR
2011-10-01
...-basic steel, aluminum, brass, bronze, or copper mill products. 252.216-7000 Section 252.216-7000 Federal... adjustment—basic steel, aluminum, brass, bronze, or copper mill products. As prescribed in 216.203-4-70(a), use the following clause: Economic Price Adjustment—Basic Steel, Aluminum, Brass, Bronze, or Copper...
Effect of alloying elements on the physicomechanical properties of copper and tin bronze
NASA Astrophysics Data System (ADS)
Ri, Kh.; Komkov, V. G.; Ri, E. Kh.
2014-09-01
The effect of alloying elements (Al, Si, Mn, Zn, Ni, As) on the physicomechanical properties of copper and tin bronze (6 wt % Sn) is studied. These alloying elements are found to increase the hardness and the microhardness of the structural constituents of Cu- X alloys due to hardening the α solid solution and eutectoid, and this effect of alloying elements is most effective in tin bronze. Alloyed copper and tin bronze have a lower thermal conductivity and corrosion resistance as compared to plain copper and tin bronze.
Thermal and Electrical Conductivity Measurements of Cda 510 Phosphor Bronze
NASA Astrophysics Data System (ADS)
Tuttle, J.; Canavan, E.; DiPirro, M.
2010-04-01
Many cryogenic systems use electrical cables containing phosphor bronze wire. While phosphor bronze's electrical and thermal conductivity values have been published, results vary among different phosphor bronze formulations. The James Webb Space Telescope (JWST) will use several phosphor bronze wire harnesses containing a specific formulation (CDA 510, annealed temper). These harnesses dominate the heat conducted into the JWST instrument stage, and approximately half of the harness conductance is due to the phosphor bronze wires. Since the JWST radiators are expected to keep the instruments at their operating temperature with limited cooling margin, it is important to know the thermal conductivity of the actual alloy being used. We describe an experiment that measured its electrical and thermal conductivity between 4 and 295 Kelvin.
Thermal and Electrical Conductivity Measurements of CDA 510 Phosphor Bronze
NASA Technical Reports Server (NTRS)
Tuttle, James E.; Canavan, Edgar; DiPirro, Michael
2009-01-01
Many cryogenic systems use electrical cables containing phosphor bronze wire. While phosphor bronze's electrical and thermal conductivity values have been published, there is significant variation among different phosphor bronze formulations. The James Webb Space Telescope (JWST) will use several phosphor bronze wire harnesses containing a specific formulation (CDA 510, annealed temper). The heat conducted into the JWST instrument stage is dominated by these harnesses, and approximately half of the harness conductance is due to the phosphor bronze wires. Since the JWST radiators are expected to just keep the instruments at their operating temperature with limited cooling margin, it is important to know the thermal conductivity of the actual alloy being used. We describe an experiment which measured the electrical and thermal conductivity of this material between 4 and 295 Kelvin.
Corrosion and Preservation of Bronze Artifacts.
ERIC Educational Resources Information Center
Walker, Robert
1980-01-01
Reviews chemical information relating to the corrosion of bronze artifacts. Properties of copper alloys are reviewed, with a thorough discussion of the specialized properties of bronze. Techniques to reduce or eliminate corrosion are listed. (CS)
Metallography and microstructure interpretation of some archaeological tin bronze vessels from Iran
DOE Office of Scientific and Technical Information (OSTI.GOV)
Oudbashi, Omid, E-mail: o.oudbashi@aui.ac.ir; Davami, Parviz, E-mail: pdavami@razi-foundation.com
2014-11-15
Archaeological excavations in western Iran have recently revealed a significant Luristan Bronzes collection from Sangtarashan archaeological site. The site and its bronze collection are dated to Iron Age II/III of western Iran (10th–7th century BC) according to archaeological research. Alloy composition, microstructure and manufacturing technique of some sheet metal vessels are determined to reveal metallurgical processes in western Iran in the first millennium BC. Experimental analyses were carried out using Scanning Electron Microscopy–Energy Dispersive X-ray Spectroscopy and Optical Microscopy/Metallography methods. The results allowed reconstructing the manufacturing process of bronze vessels in Luristan. It proved that the samples have been manufacturedmore » with a binary copper–tin alloy with a variable tin content that may relates to the application of an uncontrolled procedure to make bronze alloy (e.g. co-smelting or cementation). The presence of elongated copper sulphide inclusions showed probable use of copper sulphide ores for metal production and smelting. Based on metallographic studies, a cycle of cold working and annealing was used to shape the bronze vessels. - Highlights: • Sangtarashan vessels are made by variable Cu-Sn alloys with some impurities. • Various compositions occurred due to applying uncontrolled smelting methods. • The microstructure represents thermo-mechanical process to shape bronze vessels. • In one case, the annealing didn’t remove the eutectoid remaining from casting. • The characteristics of the bronzes are similar to other Iron Age Luristan Bronzes.« less
Preparation and evaluation of thin-film sodium tungsten bronzes
NASA Technical Reports Server (NTRS)
Kautz, H. E.; Fielder, W. L.; Singer, J.; Fordyce, J. S.
1974-01-01
Thin films of sodium tungsten bronze (NaxWO3) were investigated as reversible sodium ion electrodes for solid electrolytes. The films were made by electron beam evaporation of the three phases, W metal, Na2WO4, and WO3, followed by sintering. The substrates were sodium beta alumina disks and glass slides. X-ray diffraction analyses of the films showed that sintering in dry nitrogen with prior exposure to air lead to mixed phases. Sintering in vacuum with no air exposure produced tetragonal I bronze with a nominal composition of Na0.31WO3, single phase within the limits of X-ray diffraction detectability. The films were uniform and adherent on sodium beta alumina substrates. The ac and dc conductivities of the beta alumina were measured with the sodium tungsten bronze films as electrodes. These experiments indicated that the tetragonal I bronze electrodes were not completely reversible. This may have resulted from sodium ion blocking within the bronze film or at the bronze beta alumina interface. Methods for attempting to make more completely reversible electrodes are suggested.
Nacre biomimetic design--a possible approach to prepare low infrared emissivity composite coatings.
Zhang, Weigang; Xu, Guoyue; Ding, Ruya; Duan, Kaige; Qiao, Jialiang
2013-01-01
Mimicking the highly organized brick-and-mortar structure of nacre, a kind of nacre-like organic-inorganic composite material of polyurethane (PU)/flaky bronze composite coatings with low infrared emissivity was successfully designed and prepared by using PU and flaky bronze powders as adhesives and pigments, respectively. The infrared emissivity and microstructure of the coatings were systematically investigated by infrared emissometer and scanning electron microscopy, respectively, and the cause of low infrared emissivity of the coatings was discussed by using the theories of one-dimensional photonic structure. The results show that the infrared emissivity of the nacre-like PU/flaky bronze composite coatings can be as low as 0.206 at the bronze content of 60 wt. %, and it is significantly lower than the value of PU/sphere bronze composite coatings. Microstructure observation illustrated that the nacre-like PU/flaky bronze composite coatings have similar one-dimensional photonic structural characteristics. The low infrared emissivity of PU/flaky bronze composite coatings is derived from the similar one-dimensional photonic structure in the coatings. Copyright © 2012 Elsevier B.V. All rights reserved.
Positron lifetime in vanadium oxide bronzes
NASA Astrophysics Data System (ADS)
Dryzek, J.; Dryzek, E.
2003-09-01
The positron lifetime (PL) and Doppler broadening (DB) of annihilation line measurements have been performed in vanadium oxide bronzes MxV2O5. The dependence of these annihilation characteristics on the kind and concentration of the metal M donor has been observed. In the PL spectrum only one lifetime component has been detected in all studied bronzes. The results indicate the positron localization in the structural tunnels present in the crystalline lattice of the vanadium oxide bronzes. (
Kattner, W.T.
1959-08-11
A process is described for recovering tin from bronze comprising melting the bronze; slowly cooling the melted metal to from 280 to 240 deg C whereby eta- phase bronze crystallizes; separating the eta-bronze crystals from the liquid metal by mechanical means; melting the separated crystals; slowly cooling the melted eta-crystals to a temperature from 520 to 420 deg C whereby crystals of epsilonbronze precipitate; removing said epsilon-crystals from the remaining molten metal; and reintroducing the remaining molten metal into the process for eta-crystallization.
Jia, La-jiang; Jin, Pu-jun
2015-01-01
The present paper analyzes the interior rust that occurred in bronze alloy sample from 24 pieces of Early Qin bronze wares. Firstly, samples were processed by grinding, polishing and ultrasonic cleaning-to make a mirror surface. Then, a confocal micro-Raman spectrometer was employed to carry out spectroscopic study on the inclusions in samples. The conclusion indicated that corrosive phases are PbCO3 , PbO and Cu2O, which are common rusting production on bronze alloy. The light-colored circular or massive irregular areas in metallographic structure of samples are proved as Cu2O, showing that bronze wares are not only easy to be covered with red Cu2O rusting layer, but also their alloy is easy to be eroded by atomic oxygen. In other words, the rust Cu2O takes place in both the interior and exterior parts of the bronze alloy. In addition, Raman spectrum analysis shows that the dark grey materials are lead corrosive products--PbCO3 and PbO, showing the corroding process of lead element as Pb -->PbO-->PbCO3. In the texture of cast state of bronze alloy, lead is usually distributed as independent particles between the different alloy phases. The lead particles in bronze alloy would have oxidation reaction and generate PbO when buried in the soil, and then have chemical reaction with CO3(2-) dissolved in the underground water to generate PbCO3, which is a rather stable lead corrosive production. A conclusion can be drawn that the external corrosive factors (water, dissolved oxygen and carbonate, etc) can enter the bronze ware interior through the passageway between different phases and make the alloy to corrode gradually.
Federal Register 2010, 2011, 2012, 2013, 2014
2010-09-22
... DEPARTMENT OF STATE [Public Notice 7181] Culturally Significant Objects Imported for Exhibition Determinations: ``Ancient Chinese Bronzes From the Shouyang Studio: The Katherine and George Fan Collection... ``Ancient Chinese Bronzes from the Shouyang Studio: The Katherine and George Fan Collection,'' imported from...
21 CFR 73.2646 - Bronze powder.
Code of Federal Regulations, 2012 CFR
2012-04-01
... ADDITIVES EXEMPT FROM CERTIFICATION Cosmetics § 73.2646 Bronze powder. (a) Identity and specifications. The....1646 (a)(1) and (b). (b) Uses and restrictions. Bronze powder may be safely used in coloring cosmetics generally, including cosmetics intended for use in the area of the eye, in amounts consistent with good...
21 CFR 73.2646 - Bronze powder.
Code of Federal Regulations, 2014 CFR
2014-04-01
... ADDITIVES EXEMPT FROM CERTIFICATION Cosmetics § 73.2646 Bronze powder. (a) Identity and specifications. The....1646 (a)(1) and (b). (b) Uses and restrictions. Bronze powder may be safely used in coloring cosmetics generally, including cosmetics intended for use in the area of the eye, in amounts consistent with good...
21 CFR 73.2646 - Bronze powder.
Code of Federal Regulations, 2013 CFR
2013-04-01
... ADDITIVES EXEMPT FROM CERTIFICATION Cosmetics § 73.2646 Bronze powder. (a) Identity and specifications. The....1646 (a)(1) and (b). (b) Uses and restrictions. Bronze powder may be safely used in coloring cosmetics generally, including cosmetics intended for use in the area of the eye, in amounts consistent with good...
21 CFR 73.2646 - Bronze powder.
Code of Federal Regulations, 2011 CFR
2011-04-01
... 21 Food and Drugs 1 2011-04-01 2011-04-01 false Bronze powder. 73.2646 Section 73.2646 Food and Drugs FOOD AND DRUG ADMINISTRATION, DEPARTMENT OF HEALTH AND HUMAN SERVICES GENERAL LISTING OF COLOR....1646 (a)(1) and (b). (b) Uses and restrictions. Bronze powder may be safely used in coloring cosmetics...
21 CFR 73.2646 - Bronze powder.
Code of Federal Regulations, 2010 CFR
2010-04-01
... 21 Food and Drugs 1 2010-04-01 2010-04-01 false Bronze powder. 73.2646 Section 73.2646 Food and Drugs FOOD AND DRUG ADMINISTRATION, DEPARTMENT OF HEALTH AND HUMAN SERVICES GENERAL LISTING OF COLOR....1646 (a)(1) and (b). (b) Uses and restrictions. Bronze powder may be safely used in coloring cosmetics...
Federal Register 2010, 2011, 2012, 2013, 2014
2010-03-30
... DEPARTMENT OF STATE [Public Notice 6936] Culturally Significant Objects Imported for Exhibition Determinations: ``Gods of Angkor: Bronzes From the National Museum of Cambodia'' SUMMARY: Notice is hereby given... objects to be included in the exhibition ``Gods of Angkor: Bronzes from the National Museum of Cambodia...
Federal Register 2010, 2011, 2012, 2013, 2014
2011-07-08
... the Federal Register on February 10, 2011 (76 FR 7584). Workers were engaged in the production of cast... their purchases of cast bronze memorial products (and like or directly competitive articles) in 2007... Department determines that imports of articles like or directly competitive with the cast bronze memorial...
NASA Astrophysics Data System (ADS)
Zhao, Jian-hua; Li, Pu; Tang, Qi; Zhang, Yan-qing; He, Jian-sheng; He, Ke
2017-02-01
In this study, the bronze-matrix/x-graphite (x = 0, 1, 3 and 5%) composites were fabricated by powder metallurgy route by using Cu-coated graphite, Ni-coated graphite and pure graphite, respectively. The microstructure, mechanical properties and corrosive behaviors of bronze/Cu-coated-graphite (BCG), bronze/Ni-coated-graphite (BNG) and bronze/pure-graphite (BPG) were characterized and investigated. Results show that the Cu-coated and Ni-coated graphite could definitely increase the bonding quality between the bronze matrix and graphite. In general, with the increase in graphite content in bronze-matrix/graphite composites, the friction coefficients, ultimate density and wear rates of BPG, BCG and BNG composites all went down. However, the Vickers microhardness of the BNG composite would increase as the graphite content increased, which was contrary to the BPG and BCG composites. When the graphite content was 3%, the friction coefficient of BNG composite was more stable than that of BCG and BPG composites, indicating that BNG composite had a better tribological performance than the others. Under all the values of applied loads (10, 20, 40 and 60N), the BCG and BNG composites exhibited a lower wear rate than BPG composite. What is more, the existence of nickel in graphite powders could effectively improve the corrosion resistance of the BNG composite.
Population genomics of Bronze Age Eurasia.
Allentoft, Morten E; Sikora, Martin; Sjögren, Karl-Göran; Rasmussen, Simon; Rasmussen, Morten; Stenderup, Jesper; Damgaard, Peter B; Schroeder, Hannes; Ahlström, Torbjörn; Vinner, Lasse; Malaspinas, Anna-Sapfo; Margaryan, Ashot; Higham, Tom; Chivall, David; Lynnerup, Niels; Harvig, Lise; Baron, Justyna; Della Casa, Philippe; Dąbrowski, Paweł; Duffy, Paul R; Ebel, Alexander V; Epimakhov, Andrey; Frei, Karin; Furmanek, Mirosław; Gralak, Tomasz; Gromov, Andrey; Gronkiewicz, Stanisław; Grupe, Gisela; Hajdu, Tamás; Jarysz, Radosław; Khartanovich, Valeri; Khokhlov, Alexandr; Kiss, Viktória; Kolář, Jan; Kriiska, Aivar; Lasak, Irena; Longhi, Cristina; McGlynn, George; Merkevicius, Algimantas; Merkyte, Inga; Metspalu, Mait; Mkrtchyan, Ruzan; Moiseyev, Vyacheslav; Paja, László; Pálfi, György; Pokutta, Dalia; Pospieszny, Łukasz; Price, T Douglas; Saag, Lehti; Sablin, Mikhail; Shishlina, Natalia; Smrčka, Václav; Soenov, Vasilii I; Szeverényi, Vajk; Tóth, Gusztáv; Trifanova, Synaru V; Varul, Liivi; Vicze, Magdolna; Yepiskoposyan, Levon; Zhitenev, Vladislav; Orlando, Ludovic; Sicheritz-Pontén, Thomas; Brunak, Søren; Nielsen, Rasmus; Kristiansen, Kristian; Willerslev, Eske
2015-06-11
The Bronze Age of Eurasia (around 3000-1000 BC) was a period of major cultural changes. However, there is debate about whether these changes resulted from the circulation of ideas or from human migrations, potentially also facilitating the spread of languages and certain phenotypic traits. We investigated this by using new, improved methods to sequence low-coverage genomes from 101 ancient humans from across Eurasia. We show that the Bronze Age was a highly dynamic period involving large-scale population migrations and replacements, responsible for shaping major parts of present-day demographic structure in both Europe and Asia. Our findings are consistent with the hypothesized spread of Indo-European languages during the Early Bronze Age. We also demonstrate that light skin pigmentation in Europeans was already present at high frequency in the Bronze Age, but not lactose tolerance, indicating a more recent onset of positive selection on lactose tolerance than previously thought.
DOE Office of Scientific and Technical Information (OSTI.GOV)
Friedman, E. S.; Brody, A. J.; Young, M. L.
Seven bronze bangles from Tell en-Nasbeh, northern Judah, were investigated to understand the phase composition and manufacturing process of the artifacts, and possibly suggest a provenance for their origin. Synchrotron x-ray radiation diffraction (XRD) and fluorescence (XRF) were used in the analysis to avoid any destructive sampling and at the same time penetrate through the surface into the core metal. These techniques enabled us to determine that the bangles were not just tin bronze, but leaded tin bronze. Based on excavation reports, it is unlikely that the metal objects were manufactured locally at Tell en-Nasbeh; rather, preliminary XRD and XRFmore » data point towards the neighboring region of Edom as their origin. Despite their political enmity during the Iron Age II, the data suggest that Judahite social demands for bronze may have fostered a strong economic relationship between these two polities.« less
Code of Federal Regulations, 2010 CFR
2010-10-01
...-basic steel, aluminum, brass, bronze, or copper mill products. 252.216-7000 Section 252.216-7000 Federal... adjustment—basic steel, aluminum, brass, bronze, or copper mill products. As prescribed in 216.203-4-70(a... Mill Products (JUL 1997) (a) Definitions. As used in this clause— Established price means a price which...
Propolis as a green corrosion inhibitor for bronze in weakly acidic solution
NASA Astrophysics Data System (ADS)
Varvara, Simona; Bostan, Roxana; Bobis, Otilia; Găină, Luiza; Popa, Florin; Mena, Vicente; Souto, Ricardo M.
2017-12-01
In the present work, the inhibitive action of natural propolis on bronze corrosion in a weakly acidic solution containing Na2SO4 and NaHCO3 at pH 5 was evaluated using multiscale electrochemical techniques, namely potentiodynamic polarization, electrochemical impedance spectroscopy and scanning vibrating electrode technique measurements. The major constituents of propolis were identified by HPLC. Surface characterization was performed by SEM-EDX and AFM analysis. Experiments were performed as a function of the propolis concentration and immersion time in the corrosive electrolyte. The obtained results showed that propolis presents good anticorrosive properties on bronze, acting as a mixed-type inhibitor, but its protective effectiveness is time-dependent. The highest inhibiting efficiency of 98.9% was obtained in the presence of 100 ppm propolis, after about 12 h of exposure to inhibitor-containing electrolyte through the stabilization of Cu2O on the bronze surface. The inhibitive properties of propolis on bronze corrosion are likely due to the adsorption of its main constituents (flavonoids and phenolic compounds), through the oxygen atoms in their functional groups and aromatic rings, which have been evidenced by FT-IR spectra. The adsorption of propolis on bronze was found to follow Langmuir adsorption isotherm.
Fabrication of a Bronze Age Sword using Ancient Techniques
NASA Astrophysics Data System (ADS)
Sapiro, David; Webler, Bryan
2016-12-01
A khopesh was cast and forged for the TMS 2016 Bladesmithing Symposium. The khopesh was the first sword style, originating during the Bronze Age in the Near East. The manufacturing process used in this study closely followed Bronze Age techniques to determine the plausibility of open mold casting coupled with cold work and annealing cycles. Forging and annealing cycles substantially increased blade strength and diminished intergranular δ-phase inclusions. While a functional blade was not completed due to casting defects, the process gives valuable insight into the effort required to fabricate a khopesh during the Bronze Age. Forging and annealing cycles following casting were necessary to produce the mechanical properties desired in a sword.
Study of wear mechanism of chopped fiber reinforced epoxy composite filled with graphite and bronze
NASA Astrophysics Data System (ADS)
Patil, Nitinchand; Prasad, Krishna
2018-04-01
The combined effect of graphite and sintered bronze with a short glass fiber reinforced epoxy composites was investigated in this work. A pin on disc wear test was carried out to study the wear behaviour and mechanism of the composites. The objective of this work is to develop an alternate friction resistance material for the application of sliding bearing. It was observed that the addition of sintered bronze improved mechanical and thermal stability of the composites as bronze has low contact resistance with graphite and has high thermal conductivity. It was observed from the test results that increased volume percentage of graphite and presence of bronze are play significant role in wear mechanism of the composites. It was observed from the scanning electronic microscopes (SEM) that the abrasive and adhesive wear mechanism was prominent in this study. It was also evident from the result that the frictional force remains stable irrespective of the applied normal load.
Anderung, Cecilia; Bouwman, Abigail; Persson, Per; Carretero, José Miguel; Ortega, Ana Isabel; Elburg, Rengert; Smith, Colin; Arsuaga, Juan Luis; Ellegren, Hans; Götherström, Anders
2005-01-01
The geographic situation of the Iberian Peninsula makes it a natural link between Europe and North Africa. However, it is a matter of debate to what extent African influences via the Straits Gibraltar have affected Iberia's prehistoric development. Because early African pastoralist communities were dedicated to cattle breeding, a possible means to detect prehistoric African–Iberian contacts might be to analyze the origin of cattle breeds on the Iberian Peninsula. Some contemporary Iberian cattle breeds show a mtDNA haplotype, T1, that is characteristic to African breeds, generally explained as being the result of the Muslim expansion of the 8th century A.D., and of modern imports. To test a possible earlier African influence, we analyzed mtDNA of Bronze Age cattle from the Portalón cave at the Atapuerca site in northern Spain. Although the majority of samples showed the haplotype T3 that dominates among European breeds of today, the T1 haplotype was found in one specimen radiocarbon dated 1800 calibrated years B.C. Accepting T1 as being of African origin, this result indicates prehistoric African–Iberian contacts and lends support to archaeological finds linking early African and Iberian cultures. We also found a wild ox haplotype in the Iberian Bronze Age sample, reflecting local hybridization or backcrossing or that aurochs were hunted by these farming cultures. PMID:15941827
Barbosa, Eduardo R M; Tomlinson, Kyle W; Carvalheiro, Luísa G; Kirkman, Kevin; de Bie, Steven; Prins, Herbert H T; van Langevelde, Frank
2014-01-01
Changes in land use may lead to increased soil nutrient levels in many ecosystems (e.g. due to intensification of agricultural fertilizer use). Plant species differ widely in their response to differences in soil nutrients, and for savannas it is uncertain how this nutrient enrichment will affect plant community dynamics. We set up a large controlled short-term experiment in a semi-arid savanna to test how water supply (even water supply vs. natural rainfall) and nutrient availability (no fertilisation vs. fertilisation) affects seedlings' above-ground biomass production and leaf-nutrient concentrations (N, P and K) of broad-leafed and fine-leafed tree species. Contrary to expectations, neither changes in water supply nor changes in soil nutrient level affected biomass production of the studied species. By contrast, leaf-nutrient concentration did change significantly. Under regular water supply, soil nutrient addition increased the leaf phosphorus concentration of both fine-leafed and broad-leafed species. However, under uneven water supply, leaf nitrogen and phosphorus concentration declined with soil nutrient supply, this effect being more accentuated in broad-leafed species. Leaf potassium concentration of broad-leafed species was lower when growing under constant water supply, especially when no NPK fertilizer was applied. We found that changes in environmental factors can affect leaf quality, indicating a potential interactive effect between land-use changes and environmental changes on savanna vegetation: under more uneven rainfall patterns within the growing season, leaf quality of tree seedlings for a number of species can change as a response to changes in nutrient levels, even if overall plant biomass does not change. Such changes might affect herbivore pressure on trees and thus savanna plant community dynamics. Although longer term experiments would be essential to test such potential effects of eutrophication via changes in leaf nutrient concentration, our findings provide important insights that can help guide management plans that aim to preserve savanna biodiversity.
Warm Spraying of High-Strength Ni-Al-Bronze: Cavitation Characteristics and Property Prediction
NASA Astrophysics Data System (ADS)
Krebs, Sebastian; Kuroda, Seiji; Katanoda, Hiroshi; Gaertner, Frank; Klassen, Thomas; Araki, Hiroshi; Frede, Simon
2017-01-01
Bronze materials such as Ni-Al-bronze show exceptional performances against cavitation erosion, due to their high fatigue strength and high strength. These materials are used for ship propellers, pump systems or for applications with alternating stresses. Usually, the respective parts are cast. With the aim to use resources more efficiently and to reduce costs, this study aimed to evaluate opportunities to apply bronze as a coating to critical areas of respective parts. The coatings should have least amounts of pores and non-bonded areas and any contaminations that might act as crack nuclei and contribute to material damages. Processes with low oxidation and high kinetic impacts fulfill these criteria. Especially warm spraying, a nitrogen-cooled HVOF process, with similar impact velocities as cold gas spraying but enhanced process temperature, allows for depositing high-strength Ni-Al-bronze. This study systematically simulates and evaluates the formation and performance of warm-sprayed Ni-Al-bronze coatings for different combustion pressures and nitrogen flow rates. Substrate preheating was used to improve coating adhesion for lower spray parameter sets. Furthermore, this study introduces an energy-based concept to compare spray parameter sets and to predict coating properties. Coatings with low porosities and high mechanical strengths are obtained, allowing for a cavitation resistance similar to bulk material.
Effects of combination of leaf resources on competition in container mosquito larvae.
Reiskind, M H; Zarrabi, A A; Lounibos, L P
2012-08-01
Resource diversity is critical to fitness in many insect species, and may determine the coexistence of competitive species and the function of ecosystems. Plant material provides the nutritional base for numerous aquatic systems, yet the consequences of diversity of plant material have not been studied in aquatic container systems important for the production of mosquitoes. To address how diversity in leaf detritus affects container-inhabiting mosquitoes, we examined how leaf species affect competition between two container inhabiting mosquito larvae, Aedes aegypti and Aedes albopictus, that co-occur in many parts of the world. We tested the hypotheses that leaf species changes the outcome of intra- and interspecific competition between these mosquito species, and that combinations of leaf species affect competition in a manner not predictable based upon the response to each leaf species alone (i.e. the response to leaf combinations is non-additive). We find support for our first hypothesis that leaf species can affect competition, evidence that, in general, leaf combination alters competitive interactions, and no support that leaf combination impacts interspecific competition differently than intraspecific competition. We conclude that combinations of leaves increase mosquito production non-additively such that combinations of leaves act synergistically, in general, and result in higher total yield of adult mosquitoes in most cases, although certain leaf combinations for A. albopictus are antagonistic. We also conclude that leaf diversity does not have a different effect on interspecific competition between A. aegypti and A. albopictus, relative to intraspecific competition for each mosquito.
NASA Astrophysics Data System (ADS)
Nishida, Takamasa; Eda, Kazuo
2018-02-01
Hydrothermal syntheses of alkali-metal blue molybdenum bronze nanoribbons, which are expected to exhibit unique properties induced by a combined effect of extrinsic and intrinsic low-dimensionalities, from hydrated-alkali-metal molybdenum bronzes were investigated. Nanoribbons grown along the quasi-one-dimensional (1D) conductive direction of Cs0.3MoO3, which is difficult to prepare by the conventional methods, were first synthesized. The nanomorphology formation is achieved by a solid-state conversion (or crystallite splitting) and subsequent crystallite growth, and the structural changes of the starting material related to the conversion were first observed by powder X-ray diffraction and scanning transmission electron microscopy as a result of finely tuned reaction system and preparation conditions. The structural changes were analyzed by model simulations and were attributed to the structural modulations that were concerned with the intralayer packing disorder and with two-dimensional long-range ordered structure, formed in MoO3 sheets of the hydrated molybdenum bronze. Moreover, the modulations were related to displacement defects of the Mo-O framework units generated along the [100] direction in the hydrated molybdenum bronze. Then, it was suggested that the solid-state conversion into blue molybdenum bronze and the crystallite splitting to nanomorphology were initiated by the breaking of the Mo-O-Mo bonds at the defects. [Figure not available: see fulltext.
NASA Astrophysics Data System (ADS)
Hauer, Michél; Henkel, Knuth Michael; Krebs, Sebastian; Kroemmer, Werner
2017-01-01
Within a research project regarding cavitation erosion-resistant coatings, arc spraying was used with different traverse speeds to influence heat transfer and the resulting residual stress state. The major reason for this study is the lack of knowledge concerning the influence of residual stress distribution on mechanical properties and coating adhesion, especially with respect to heterogeneous aluminum bronze alloys. The materials used for spray experiments were the highly cavitation erosion-resistant propeller alloys CuAl9Ni5Fe4Mn (Ni-Al-Bronze) and CuMn13Al8Fe3Ni2 (Mn-Al-Bronze). Analyses of cavitation erosion behavior were carried out to evaluate the suitability for use in marine environments. Further microstructural, chemical and mechanical analyses were realized to examine adhesive and cohesive coating properties. Residual stress distribution was measured by modified hole drilling method using electronic speckle pattern interferometry (ESPI). It was found that the highest traverse speed led to higher tensile residual stresses near the surface and less cavitation erosion resistance of the coatings. Moreover, high oxygen affinity of main alloying element aluminum was identified to severely influence the microstructures by the formation of large oxides and hence the coating properties. Overall, Mn-Al-Bronze coatings showed lower residual stresses, a more homogeneous pore and oxide distribution and less material loss by cavitation than Ni-Al-Bronze coatings.
Copper smelting and sediment pollution in Bronze Age China.
NASA Astrophysics Data System (ADS)
Zhang, S.; Dong, G.
2017-12-01
The emergence and diffusion of metallurgical technology had tremendous environmental consequence, however, the spatial-temporal consequences of the metallurgy during Bronze Age are not clear in China. Here, Xray fluorescence (XRF) measurement and principal component analysis (PCA) were conducted on heavy metal element (Cu, Ni, Pb, Zn, Cr and As) concentrations (HMEC) of natural and anthropogenic sediment samples systematically collected from 22 late Neolithic-Bronze Age sites in Hexi corridor to explore the potential for subcontinental-wide changes in soil geochemistry. We place this data within the context of the Cu concentrations in lacustrine sediments located near smelting and mining centers in Bronze Age China. Our results show that variation of HMEC in anthropogenic sediment in Hexi corridor is contemporaneous with the increases of the Cu concentrations in lacustrine sediment around 4000 BP. Comparative data suggests the metallurgical production diffused from the Hexi corridor to central and southwestern China around 3600 BP. We argue that sediment pollution is not an isolated phenomenon during the Bronze Age China, but rather occurred on regional scales and is closely related to the intensity of smelting activities.
Graetz, Ilana; McKillop, Caitlin N; Kaplan, Cameron M; Waters, Teresa M
2017-05-01
Since 2014, average premiums for health plans available in the Affordable Care Act marketplaces have increased. We examine how premium price changes affected the amount consumers pay after subsidies for the lowest-cost bronze and silver plans available by age in the federally facilitated exchanges. Between 2015 and 2016, benchmark plan premiums increased in 83.3% of counties. Overall, rising benchmark premiums were associated with lower average after-subsidy premiums for the lowest-cost bronze and silver plans for older subsidy-eligible adults, but with higher after-subsidy premiums for younger adults purchasing the same plans, regardless of income. With recent discussions to replace or overhaul the Affordable Care Act, it is critical that we learn from the successes and failures of the current policy. Our findings suggest that the subsidy design, which makes rising premiums costlier for younger adults looking to purchase an entry-level plan, may be contributing to adverse selection and instability in the marketplace.
The effect of uric acid on outdoor copper and bronze.
Bernardi, E; Bowden, D J; Brimblecombe, P; Kenneally, H; Morselli, L
2009-03-15
Bird droppings are often quoted as a decay agent for outdoor goods, in particular buildings and statues. Undoubtedly, they represent one of the major causes of aesthetic damage on outdoor materials, but the real chemical damage they are able to induce, in particular on metals, is not so well studied. This work focused on the short term role of uric acid, the main constituent of bird urine, with respect to copper, which make such an important contribution to architectural elements of buildings and outdoor sculpture. Preliminary results of laboratory tests and analyses on real exposed samples showed that uric acid chemically affects copper and bronzes: the surface of the metal is modified and copper urates formed. Also natural patina, formed on statues and roof, react with uric acid, even if it seems to afford some protection toward bird droppings. In general, experimental results confirm that the potential chemical damage by bird droppings is significant when considering external cultural heritage such as statues, metal monuments and buildings with historic copper roofs.
USDA-ARS?s Scientific Manuscript database
Leaf architectural traits, such as length, width and angle, directly influence canopy structure and light penetration, photosynthate production and overall yield. We discovered and characterized a maize (Zea mays) mutant with aberrant leaf architecture we named drooping leaf1 (drl1), as leaf blades ...
Bronze rainbow hologram mirrors
NASA Astrophysics Data System (ADS)
Dawson, P.
2006-02-01
This project draws on holographic embossing techniques, ancient artistic conventions of bronze mirror design and modelling and casting processes to accomplish portraiture of reflection. Laser scanning, 3D computer graphics and holographic imaging are employed to enable a permanent 3D static holographic image to appear integrated with the real-time moving reflection of a viewer's face in a polished bronze disc. The disc and the figure which holds it (caryatid) are cast in bronze from a lost wax model, a technique which has been used for millennia to make personal mirrors. The Caryatid form of bronze mirror which went through many permutations in ancient Egyptian, Greece and Rome shows a plethora of expressive figure poses ranging from sleek nudes to highly embellished multifigure arrangements. The prototype of this series was made for Australian choreographer Graeme Murphy, Artistic Director of the Sydney Dance Company. Each subsequent mirror will be unique in figure and holographic imagery as arranged between artist and subject. Conceptually this project references both the modern experience of viewing mirrors retrieved from ancient tombs, which due to deterioration of the surface no longer reflect, and the functioning of Chinese Magic mirrors, which have the ability to project a predetermined image. Inspired by the metaphorical potential of these mirrors, which do not reflect the immediate reality of the viewer, this bronze hologram mirror series enables each viewer to reflect upon himself or herself observing simultaneously the holographic image and their own partially obliterated reflection.
NASA Technical Reports Server (NTRS)
Huemmrich, Karl F.
2013-01-01
The leaf inclination angle distribution (LAD) is an important characteristic of vegetation canopy structure affecting light interception within the canopy. However, LADs are difficult and time consuming to measure. To examine possible global patterns of LAD and their implications in remote sensing, a model was developed to predict leaf angles within canopies. Canopies were simulated using the SAIL radiative transfer model combined with a simple photosynthesis model. This model calculated leaf inclination angles for horizontal layers of leaves within the canopy by choosing the leaf inclination angle that maximized production over a day in each layer. LADs were calculated for five latitude bands for spring and summer solar declinations. Three distinct LAD types emerged: tropical, boreal, and an intermediate temperate distribution. In tropical LAD, the upper layers have a leaf angle around 35 with the lower layers having horizontal inclination angles. While the boreal LAD has vertical leaf inclination angles throughout the canopy. The latitude bands where each LAD type occurred changed with the seasons. The different LADs affected the fraction of absorbed photosynthetically active radiation (fAPAR) and Normalized Difference Vegetation Index (NDVI) with similar relationships between fAPAR and leaf area index (LAI), but different relationships between NDVI and LAI for the different LAD types. These differences resulted in significantly different relationships between NDVI and fAPAR for each LAD type. Since leaf inclination angles affect light interception, variations in LAD also affect the estimation of leaf area based on transmittance of light or lidar returns.
Calcium oxalate druses affect leaf optical properties in selenium-treated Fagopyrum tataricum.
Golob, Aleksandra; Stibilj, Vekoslava; Nečemer, Marijan; Kump, Peter; Kreft, Ivan; Hočevar, Anja; Gaberščik, Alenka; Germ, Mateja
2018-03-01
Plants of the genus Fagopyrum contain high levels of crystalline calcium oxalate (CaOx) deposits, or druses, that can affect the leaf optical properties. As selenium has been shown to modify the uptake and accumulation of metabolically important elements such as calcium, we hypothesised that the numbers of druses can be altered by selenium treatment, and this would affect the leaf optical properties. Tartary buckwheat (Fagopyrum tataricum Gaertn.) was grown outdoors in an experimental field. At the beginning of flowering, plants were foliarly sprayed with sodium selenate solution at 10 mg selenium L -1 or only with water. Plant morphological, biochemical, physiological and optical properties were examined, along with leaf elemental composition and content. Se spraying did not affect leaf biochemical and functional properties. However, it increased leaf thickness and the contents of Se in the leaves, and decreased the density of calcium oxalate druses in the leaves. Except Se content, Se spraying did not affect contents of other elements in leaves, including total calcium per dry mass of leaf tissue. Redundancy analysis showed that of all parameters tested, only the calcium oxalate druses parameters were significant in explaining the variability of the leaf reflectance and transmittance spectra. The density of CaOx druses positively correlated with the reflectance in the blue, green, yellow and UV-B regions of the spectrum, while the area of CaOx druses per mm 2 of leaf transection area positively correlated with the transmittance in the green and yellow regions of the spectrum. Copyright © 2018 Elsevier B.V. All rights reserved.
Mytych, Jennifer; Satora, Leszek; Kozioł, Katarzyna
2018-02-01
Bronze corydoras (Corydoras aeneus) uses the distal part of the intestine as accessory respiratory organ. Our previous study showed the presence of epidermal growth factor receptor (EGFR) cytoplasmic domain in the digestive tract of the bronze corydoras. In this study, using Western Blot method, we validated the results presented in the previous research. In detail, results of Western Blot analysis on digestive and respiratory part of bronze corydoras intestine homogenates confirmed the immunoreactivity of anti-cytoplasmic domain (C-terminal) human EGFR antibodies with protein band of approximately 180kDa (EGFR molecular weight). This indicates a high homology of EGFR domain between these species and the possibility of such antibody use in bronze corydoras. Statistically significantly higher EGFR expression was observed in the respiratory part of intestine when compared to the digestive part. This implies higher proliferation activity and angiogenesis of epithelium in this part of intestine, creating conditions for air respiration. Therefore, Corydoras aeneus may be considered as a model organism for the molecular studies of the mechanisms of epithelial proliferation initiation and inhibition depending on hypoxia and normoxia. Copyright © 2017. Published by Elsevier GmbH.
Detail view of the bronze gates, lit from behind by ...
Detail view of the bronze gates, lit from behind by the sun - United States Department of Commerce, Bounded by Fourteenth, Fifteenth, and E streets and Constitution Avenue, Washington, District of Columbia, DC
Bronze Alloy Development for Zinc Vapor Capture
Korinko, Paul S.
2017-04-24
After gamma-emitting 65Zinc was detected in a vacuum pumping system contained in a tritium glovebox, a series of experiments were undertaken to develop a method and material to trap zinc vapors in an area that is more suitable for preventing dose to workers. In this study, bronze alloys with 0–30% tin were prepared using a powder metallurgical process and exposed to three levels of zinc vapors. Furthermore, all of the alloys demonstrated acceptable zinc gettering capacity; however, low tin content bronzes are considered for further testing.
Electrochromic device containing metal oxide nanoparticles and ultraviolet blocking material
DOE Office of Scientific and Technical Information (OSTI.GOV)
Garcia, Guillermo; Koo, Bonil; Gregoratto, Ivano
An electrochromic device includes a nanostructured transition metal oxide bronze layer that includes one or more transition metal oxide and one or more dopant. The electrochromic device also includes nanoparticles containing one or more transparent conducting oxide (TCO), a solid state electrolyte, a counter electrode, and at least one protective layer to prevent degradation of the one or more nanostructured transition metal oxide bronze. The nanostructured transition metal oxide bronze selectively modulates transmittance of near-infrared (NIR) and visible radiation as a function of an applied voltage to the device.
Isotopic Evidence for Early Trade in Animals between Old Kingdom Egypt and Canaan.
Arnold, Elizabeth R; Hartman, Gideon; Greenfield, Haskel J; Shai, Itzhaq; Babcock, Lindsay E; Maeir, Aren M
2016-01-01
Isotope data from a sacrificial ass and several ovicaprines (sheep/goat) from Early Bronze Age household deposits at Tell es-Safi/Gath, Israel provide direct evidence for the movement of domestic draught/draft and husbandry animals between Old Kingdom Egypt (during the time of the Pyramids) and Early Bronze Age III Canaan (ca. 2900-2500 BCE). Vacillating, bi-directional connections between Egypt and Canaan are known throughout the Early Bronze Age, but here we provide the first concrete evidence of early trade in animals from Egypt to Canaan.
Hemphill, Brian E; Mallory, J P
2004-07-01
Numerous Bronze Age cemeteries in the oases surrounding the Täklamakan Desert of the Tarim Basin in the Xinjiang Uyghur Autonomous Region, western China, have yielded both mummified and skeletal human remains. A dearth of local antecedents, coupled with woolen textiles and the apparent Western physical appearance of the population, raised questions as to where these people came from. Two hypotheses have been offered by archaeologists to account for the origins of Bronze Age populations of the Tarim Basin. These are the "steppe hypothesis" and the "Bactrian oasis hypothesis." Eight craniometric variables from 25 Aeneolithic and Bronze Age samples, comprising 1,353 adults from the Tarim Basin, the Russo-Kazakh steppe, southern China, Central Asia, Iran, and the Indus Valley, are compared to test which, if either, of these hypotheses are supported by the pattern of phenetic affinities possessed by Bronze Age inhabitants of the Tarim Basin. Craniometric differences between samples are compared with Mahalanobis generalized distance (d2), and patterns of phenetic affinity are assessed with two types of cluster analysis (the weighted pair average linkage method and the neighbor-joining method), multidimensional scaling, and principal coordinates analysis. Results obtained by this analysis provide little support for either the steppe hypothesis or the Bactrian oasis hypothesis. Rather, the pattern of phenetic affinities manifested by Bronze Age inhabitants of the Tarim Basin suggests the presence of a population of unknown origin within the Tarim Basin during the early Bronze Age. After 1200 B.C., this population experienced significant gene flow from highland populations of the Pamirs and Ferghana Valley. These highland populations may include those who later became known as the Saka and who may have served as "middlemen" facilitating contacts between East (Tarim Basin, China) and West (Bactria, Uzbekistan) along what later became known as the Great Silk Road. Copyright 2004 Wiley-Liss, Inc.
NASA Astrophysics Data System (ADS)
Ilogebe, Amamchukwu Bernard
Binder-jet 3D printing has been one of the additive manufacturing techniques employed in fabrication of intricate parts, by utilizing metal powders. Liquid metal infiltration of bronze into binder-jet printed structural amorphous metal resulted in a net shape, fully-dense parts were made. The final part was characterized by means of scanning electron microscopy, electron dispersive x-ray spectroscopy and computed tomography. The densification in the binder-jet samples was also compared to die-pressed ones, and was found to be 3.96g/cm 3 and 3.89g/cm3. Thus, binder-jet can be used to model a die-pressed part. Scanning electron micrograph displayed the presence of considerable porosity in the sintered binder-jet samples, as well as some limited porosity in the infiltrated samples. Evident also from SEM analysis was the presence of internal powder micro pores. Electron dispersive spectroscopy results show that the bronze filled out the pores as was expected. According to the computed tomography results, the un-infiltrated sample has an average porosity of 34%, while the bronze-infiltrated samples have an average porosity of 1%. Micro-indentation was also performed on the infiltrated and uninfiltrated samples to evaluate the mechanical properties. The un-infiltrated sample had 2.98GPa hardness, while bronze infiltrated sample had 4.00GPa hardness using Vickers hardness method. Generally, it was found that infiltration of bronze into structural amorphous metal improved homogeneity of the material, as well as the mechanical properties. Further research needs to be done on the mechanical properties of binder-jet printed parts of SAM alloy, infiltrated with bronze. Variation of thickness needs to be included for further research to ascertain the critical achievable depth of infiltration using bronze as the reinforcement material.
Detail view of bronze light fixture on on north facade, ...
Detail view of bronze light fixture on on north facade, lower level - Blue Ridge Sanatorium, Building No. 1902, East side of State Route 20, .25 mile south of I-64, Charlottesville, Independent City, VA
Detail view of stylized railing and bronze light fixture on ...
Detail view of stylized railing and bronze light fixture on southwest facade - Blue Ridge Sanatorium, Building No. 1901, East side of State Route 20, .25 mile south of I-64, Charlottesville, Independent City, VA
18. Photocopy of drawing of bronze dedication plaque, circa 1903 ...
18. Photocopy of drawing of bronze dedication plaque, circa 1903 (original drawing in possession of City Engineer's Office Grand Rapids, Michigan) DEDICATION PLAQUE. - Bridge Street Bridge, Spanning Grand River, Michigan & Bridge Streets, Grand Rapids, MI
Interior view, detail to show the ornate bronze doors to ...
Interior view, detail to show the ornate bronze doors to the entrance lobby elevators - United States Department of Commerce, Bounded by Fourteenth, Fifteenth, and E streets and Constitution Avenue, Washington, District of Columbia, DC
Corrosion of 85-5-5-5 bronze in natural and synthetic acid rain
NASA Astrophysics Data System (ADS)
Morselli, L.; Bernardi, E.; Chiavari, C.; Brunoro, G.
In order to investigate the decay of bronzes exposed to acid wet depositions, a comparative study has been performed by following the corrosion behaviour of different sets of bronze specimens exposed either to natural rain or to a similar solution, without organic compounds, artificially reproduced in laboratory. The as cast G85 bronze specimens were exposed to aggressive solutions for different periods through a wet-dry technique. The pH trend of the solutions and the amount of metals transferred into the solutions were periodically monitored. OM, SEM, XRD, RAMAN analyses and ac electrochemical measurements were performed on the artificially weathered specimens. Preliminary results, showing the difference between the ageing in natural and synthetic rain, suggest the influence of the organic components on the corrosion process. In particular, the growth of a more uniform protective layer of corrosion products on the metal surface exposed to the natural rain could be attributed to these components.
The first horse herders and the impact of early Bronze Age steppe expansions into Asia.
de Barros Damgaard, Peter; Martiniano, Rui; Kamm, Jack; Moreno-Mayar, J Víctor; Kroonen, Guus; Peyrot, Michaël; Barjamovic, Gojko; Rasmussen, Simon; Zacho, Claus; Baimukhanov, Nurbol; Zaibert, Victor; Merz, Victor; Biddanda, Arjun; Merz, Ilja; Loman, Valeriy; Evdokimov, Valeriy; Usmanova, Emma; Hemphill, Brian; Seguin-Orlando, Andaine; Yediay, Fulya Eylem; Ullah, Inam; Sjögren, Karl-Göran; Iversen, Katrine Højholt; Choin, Jeremy; de la Fuente, Constanza; Ilardo, Melissa; Schroeder, Hannes; Moiseyev, Vyacheslav; Gromov, Andrey; Polyakov, Andrei; Omura, Sachihiro; Senyurt, Süleyman Yücel; Ahmad, Habib; McKenzie, Catriona; Margaryan, Ashot; Hameed, Abdul; Samad, Abdul; Gul, Nazish; Khokhar, Muhammad Hassan; Goriunova, O I; Bazaliiskii, Vladimir I; Novembre, John; Weber, Andrzej W; Orlando, Ludovic; Allentoft, Morten E; Nielsen, Rasmus; Kristiansen, Kristian; Sikora, Martin; Outram, Alan K; Durbin, Richard; Willerslev, Eske
2018-05-09
The Yamnaya expansions from the western steppe into Europe and Asia during the Early Bronze Age (~3000 BCE) are believed to have brought with them Indo-European languages and possibly horse husbandry. We analyze 74 ancient whole-genome sequences from across Inner Asia and Anatolia and show that the Botai people associated with the earliest horse husbandry derived from a hunter-gatherer population deeply diverged from the Yamnaya. Our results also suggest distinct migrations bringing West Eurasian ancestry into South Asia before and after but not at the time of Yamnaya culture. We find no evidence of steppe ancestry in Bronze Age Anatolia from when Indo-European languages are attested there. Thus, in contrast to Europe, Early Bronze Age Yamnaya-related migrations had limited direct genetic impact in Asia. Copyright © 2018, American Association for the Advancement of Science.
Balliana, Eleonora; Aramendía, Maite; Resano, Martin; Barbante, Carlo; Vanhaecke, Frank
2013-03-01
Although in many cases Pb isotopic analysis can be relied on for provenance determination of ancient bronzes, sometimes the use of "non-traditional" isotopic systems, such as those of Cu and Sn, is required. The work reported on in this paper aimed at revising the methodology for Cu and Sn isotope ratio measurements in archaeological bronzes via optimization of the analytical procedures in terms of sample pre-treatment, measurement protocol, precision, and analytical uncertainty. For Cu isotopic analysis, both Zn and Ni were investigated for their merit as internal standard (IS) relied on for mass bias correction. The use of Ni as IS seems to be the most robust approach as Ni is less prone to contamination, has a lower abundance in bronzes and an ionization potential similar to that of Cu, and provides slightly better reproducibility values when applied to NIST SRM 976 Cu isotopic reference material. The possibility of carrying out direct isotopic analysis without prior Cu isolation (with AG-MP-1 anion exchange resin) was investigated by analysis of CRM IARM 91D bronze reference material, synthetic solutions, and archaeological bronzes. Both procedures (Cu isolation/no Cu isolation) provide similar δ (65)Cu results with similar uncertainty budgets in all cases (±0.02-0.04 per mil in delta units, k = 2, n = 4). Direct isotopic analysis of Cu therefore seems feasible, without evidence of spectral interference or matrix-induced effect on the extent of mass bias. For Sn, a separation protocol relying on TRU-Spec anion exchange resin was optimized, providing a recovery close to 100 % without on-column fractionation. Cu was recovered quantitatively together with the bronze matrix with this isolation protocol. Isotopic analysis of this Cu fraction provides δ (65)Cu results similar to those obtained upon isolation using AG-MP-1 resin. This means that Cu and Sn isotopic analysis of bronze alloys can therefore be carried out after a single chromatographic separation using TRU-Spec resin. Tin isotopic analysis was performed relying on Sb as an internal standard used for mass bias correction. The reproducibility over a period of 1 month (n = 42) for the mass bias-corrected Sn isotope ratios is in the range of 0.06-0.16 per mil (2 s), for all the ratios monitored.
Isotopic Evidence for Early Trade in Animals between Old Kingdom Egypt and Canaan
Greenfield, Haskel J.; Shai, Itzhaq; Babcock, Lindsay E.; Maeir, Aren M.
2016-01-01
Isotope data from a sacrificial ass and several ovicaprines (sheep/goat) from Early Bronze Age household deposits at Tell es-Safi/Gath, Israel provide direct evidence for the movement of domestic draught/draft and husbandry animals between Old Kingdom Egypt (during the time of the Pyramids) and Early Bronze Age III Canaan (ca. 2900–2500 BCE). Vacillating, bi-directional connections between Egypt and Canaan are known throughout the Early Bronze Age, but here we provide the first concrete evidence of early trade in animals from Egypt to Canaan. PMID:27322197
NASA Astrophysics Data System (ADS)
Li, Jingxiang; Zhao, Shengdun; Ishihara, Kunihiko
2013-05-01
A novel approach is presented to study the acoustical properties of sintered bronze material, especially used to suppress the transient noise generated by the pneumatic exhaust of pneumatic friction clutch and brake (PFC/B) systems. The transient exhaust noise is impulsive and harmful due to the large sound pressure level (SPL) that has high-frequency. In this paper, the exhaust noise is related to the transient impulsive exhaust, which is described by a one-dimensional aerodynamic model combining with a pressure drop expression of the Ergun equation. A relation of flow parameters and sound source is set up. Additionally, the piston acoustic source approximation of sintered bronze silencer with cylindrical geometry is presented to predict SPL spectrum at a far-field observation point. A semi-phenomenological model is introduced to analyze the sound propagation and reduction in the sintered bronze materials assumed as an equivalent fluid with rigid frame. Experiment results under different initial cylinder pressures are shown to corroborate the validity of the proposed aerodynamic model. In addition, the calculated sound pressures according to the equivalent sound source are compared with the measured noise signals both in time-domain and frequency-domain. Influences of porosity of the sintered bronze material are also discussed.
Cold Spraying of Cu-Al-Bronze for Cavitation Protection in Marine Environments
NASA Astrophysics Data System (ADS)
Krebs, S.; Gärtner, F.; Klassen, T.
2015-01-01
Traveling at high speeds, ships have to face the problem of rudder cavitation-erosion. At present, the problem is countered by fluid dynamically optimized rudders, synthetic, and weld-cladded coatings on steel basis. Nevertheless, docking and repair is required after certain intervals. Bulk Cu-Al-bronzes are in use at ships propellers to withstand corrosion and cavitation. Deposited as coatings with bulk-like properties, such bronzes could also enhance rudder life times. The present study investigates the coating formation by cold spraying CuAl10Fe5Ni5 bronze powders. By calculations of the impact conditions, the range of optimum spray parameters was preselected in terms of the coating quality parameter η on steel substrates with different temperatures. As-atomized and annealed powders were compared to optimize cavitation resistance of the coatings. Results provide insights about the interplay between the mechanical properties of powder and substrate for coating formation. Single particle impact morphologies visualize the deformation behavior. Coating performance was assessed by analyzing microstructures, bond strength, and cavitation resistance. These first results demonstrate that cold-sprayed bronze coatings have a high potential for ensuring a good performances in rudder protection. With further optimization, such coatings could evolve towards a competitive alternative to existing anti-cavitation procedures.
Gortari, Fermín; Guiamet, Juan José; Graciano, Corina
2018-06-01
Rust produced by Melampsora sp. is considered one of the most relevant diseases in poplar plantations. Growth reduction in poplar plantations takes place because rust, like other pathogens, alters leaf physiology. There is not a complete evaluation of several of the physiological traits that can be affected by rust at leaf level. Therefore, the aim of this work was to evaluate, in an integrative way and in the same pathosystem, which physiological processes are affected when Populus deltoides Bartr. ex Marsh. leaves are infected by rust (Melampsora medusae Thümen). Leaves of two clones with different susceptibility to rust were analyzed. Field and pot experiments were performed, and several physiological traits were measured in healthy and infected leaves. We conclude that rust affects leaf mesophyll integrity, and so water movement in the leaf in liquid phase is affected. As a consequence, gas exchange is reduced, affecting both carbon fixation and transpiration. However, there is an increase in respiration rate, probably due to plant and fungal respiration. The increase in respiration rate is important in the reduction of net photosynthetic rate, but also some damage in the photosynthetic apparatus limits leaf capacity to fix carbon. The decrease in chlorophyll content would start later and seems not to explain the reduction in net photosynthetic rate. Both clones, although they have different susceptibility to rust, are affected in the same physiological mechanisms.
Retta, Moges; Yin, Xinyou; van der Putten, Peter E L; Cantre, Denis; Berghuijs, Herman N C; Ho, Quang Tri; Verboven, Pieter; Struik, Paul C; Nicolaï, Bart M
2016-11-01
The mechanism of photosynthesis in C 4 crops depends on the archetypal Kranz-anatomy. To examine how the leaf anatomy, as altered by nitrogen supply and leaf age, affects the bundle sheath conductance (g bs ), maize (Zea mays L.) plants were grown under three contrasting nitrogen levels. Combined gas exchange and chlorophyll fluorescence measurements were done on fully grown leaves at two leaf ages. The measured data were analysed using a biochemical model of C 4 photosynthesis to estimate g bs . The leaf microstructure and ultrastructure were quantified using images obtained from micro-computed tomography and microscopy. There was a strong positive correlation between g bs and leaf nitrogen content (LNC) while old leaves had lower g bs than young leaves. Leaf thickness, bundle sheath cell wall thickness and surface area of bundle sheath cells per unit leaf area (S b ) correlated well with g bs although they were not significantly affected by LNC. As a result, the increase of g bs with LNC was little explained by the alteration of leaf anatomy. In contrast, the combined effect of LNC and leaf age on S b was responsible for differences in g bs between young leaves and old leaves. Future investigations should consider changes at the level of plasmodesmata and membranes along the CO 2 leakage pathway to unravel LNC and age effects further. Copyright © 2016 Elsevier Ireland Ltd. All rights reserved.
Kothari, Adit R; Burnett, Nicholas P
2017-09-01
In nature, plants regularly interact with herbivores and with wind. Herbivores can wound and alter the structure of plants, whereas wind can exert aerodynamic forces that cause the plants to flutter or sway. While herbivory has many negative consequences for plants, fluttering in wind can be beneficial for plants by facilitating gas exchange and loss of excess heat. Little is known about how herbivores affect plant motion in wind. We tested how the mass of an herbivore resting on a broad leaf of the tulip tree Liriodendron tulipifera , and the damage caused by herbivores, affected the motion of the leaf in wind. For this, we placed mimics of herbivores on the leaves, varying each herbivore's mass or position, and used high-speed video to measure how the herbivore mimics affected leaf movement and reconfiguration at two wind speeds inside a laboratory wind tunnel. In a similar setup, we tested how naturally occurring herbivore damage on the leaves affected leaf movement and reconfiguration. We found that the mass of an herbivore resting on a leaf can change that leaf's orientation relative to the wind and interfere with the ability of the leaf to reconfigure into a smaller, more streamlined shape. A large herbivore load slowed the leaf's fluttering frequency, while naturally occurring damage from herbivores increased the leaf's fluttering frequency. We conclude that herbivores can alter the physical interactions between wind and plants by two methods: (1) acting as a point mass on the plant while it is feeding and (2) removing tissue from the plant. Altering a plant's interaction with wind can have physical and physiological consequences for the plant. Thus, future studies of plants in nature should consider the effect of herbivory on plant-wind interactions, and vice versa.
Phytohormones signaling and crosstalk regulating leaf angle in rice.
Luo, Xiangyu; Zheng, Jingsheng; Huang, Rongyu; Huang, Yumin; Wang, Houcong; Jiang, Liangrong; Fang, Xuanjun
2016-12-01
Leaf angle is an important agronomic trait in rice (Oryza sativa L.). It affects both the efficiency of sunlight capture and nitrogen reservoirs. The erect leaf phenotype is suited for high-density planting and thus increasing crop yields. Many genes regulate leaf angle by affecting leaf structure, such as the lamina joint, mechanical tissues, and the midrib. Signaling of brassinosteroids (BR), auxin (IAA), and gibberellins (GA) plays important roles in the regulation of lamina joint bending in rice. In addition, the biosynthesis and signaling of BR are known to have dominant effects on leaf angle development. In this review, we summarize the factors and genes associated with the development of leaf angle in rice, outline the regulatory mechanisms based on the signaling of BR, IAA, and GA, and discuss the contribution of crosstalk between BR and IAA or GA in the formation of leaf angle. Promising lines of research in the transgenic engineering of rice leaf angle to increase grain yield are proposed.
DOE Office of Scientific and Technical Information (OSTI.GOV)
Li Yang; Bao Zhirong; Wu Taotao
In this paper, a kind of specific stalactitic product was found on the interior surface of a covered bronze wine vessel with loop-handle (Chinese name is you), which was fabricated in Shang Dynasty (1700 B.C.-1100 B.C.) and now is collected in Xiaogan Museum, Hubei province of China. The microstructures of the product were characterized systematically by using optical microscopy, scanning electron microscope, transmission electron microscope, X-ray diffraction, and Raman microscopy. The experimental results revealed that the product belonged to a kind of malachite with high purity and high crystallinity. The growth of the product was considered to be a possiblemore » reason that the vessel was overly airtight within a museum display cabinet besides a lid of the vessel, which made the excess of H{sub 2}O and CO{sub 2} gas concentrations inside the vessel during long-term storage. This corrosion product is very harmful to bronze cultural relics, because of a large amount of copper consumption from the matrix which will reduce its life. The growth mechanism of the specific stalactitic product and the suggestions for preservation of the similar bronze relics in museum were proposed. - Highlights: Black-Right-Pointing-Pointer The stalactitic product was the high purity and good crystallinity malachite. Black-Right-Pointing-Pointer Its growth was related to the excess of H{sub 2}O and CO{sub 2} gas concentrations in museum. Black-Right-Pointing-Pointer It is harmful to the bronzes, because copper will be consumed from the matrix. Black-Right-Pointing-Pointer The suggestions for preservation of the similar bronzes in museum were proposed.« less
Detail view to show the bronze gates hanging in the ...
Detail view to show the bronze gates hanging in the driveway portals; the open grille is foliated and crowned with patriotic eagle emblems - United States Department of Commerce, Bounded by Fourteenth, Fifteenth, and E streets and Constitution Avenue, Washington, District of Columbia, DC
Analysis of 3800-year-old Yersinia pestis genomes suggests Bronze Age origin for bubonic plague.
Spyrou, Maria A; Tukhbatova, Rezeda I; Wang, Chuan-Chao; Valtueña, Aida Andrades; Lankapalli, Aditya K; Kondrashin, Vitaly V; Tsybin, Victor A; Khokhlov, Aleksandr; Kühnert, Denise; Herbig, Alexander; Bos, Kirsten I; Krause, Johannes
2018-06-08
The origin of Yersinia pestis and the early stages of its evolution are fundamental subjects of investigation given its high virulence and mortality that resulted from past pandemics. Although the earliest evidence of Y. pestis infections in humans has been identified in Late Neolithic/Bronze Age Eurasia (LNBA 5000-3500y BP), these strains lack key genetic components required for flea adaptation, thus making their mode of transmission and disease presentation in humans unclear. Here, we reconstruct ancient Y. pestis genomes from individuals associated with the Late Bronze Age period (~3800 BP) in the Samara region of modern-day Russia. We show clear distinctions between our new strains and the LNBA lineage, and suggest that the full ability for flea-mediated transmission causing bubonic plague evolved more than 1000 years earlier than previously suggested. Finally, we propose that several Y. pestis lineages were established during the Bronze Age, some of which persist to the present day.
USDA-ARS?s Scientific Manuscript database
Leaf shape is an important plant architecture trait that is affected by plant hormones, especially auxin. In Arabidopsis, PINOID (PID), a regulator for the auxin polar transporter PIN (PIN-FORMED) affects leaf shape formation, but this function of PID in crop plants has not been well studied. From a...
Zheng, Liang; Van Labeke, Marie-Christine
2017-01-01
Light quality critically affects plant development and growth. Development of light-emitting diodes (LEDs) enables the use of narrow band red and/or blue wavelengths as supplementary lighting in ornamental production. Yet, long periods under these wavelengths will affect leaf morphology and physiology. Leaf anatomy, stomatal traits, and stomatal conductance, leaf hydraulic conductance (K leaf ), and photosynthetic efficiency were investigated in three ornamental pot plants, namely Cordyline australis (monocot), Ficus benjamina (dicot, evergreen leaves), and Sinningia speciosa (dicot, deciduous leaves) after 8 weeks under LED light. Four light treatments were applied at 100 μmol m -2 s -1 and a photoperiod of 16 h using 100% red (R), 100% blue (B), 75% red with 25% blue (RB), and full spectrum white light (W), respectively. B and RB resulted in a greater maximum quantum yield (F v /F m ) and quantum efficiency (Φ PSII ) in all species compared to R and W and this correlated with a lower biomass under R. B increased the stomatal conductance compared with R. This increase was linked to an increasing stomatal index and/or stomatal density but the stomatal aperture area was unaffected by the applied light quality. Leaf hydraulic conductance (K leaf ) was not significantly affected by the applied light qualities. Blue light increased the leaf thickness of F. benjamina , and a relative higher increase in palisade parenchyma was observed. Also in S. speciosa , increase in palisade parenchyma was found under B and RB, though total leaf thickness was not affected. Palisade parenchyma tissue thickness was correlated to the leaf photosynthetic quantum efficiency (Φ PSII ). In conclusion, the role of blue light addition in the spectrum is essential for the normal anatomical leaf development which also impacts the photosynthetic efficiency in the three studied species.
Marcelo Ardon; Catherine M. Pringle; Susan L. Eggert
2009-01-01
Comparisons of the effects of leaf litter chemistry on leaf breakdown rates in tropical vs temperate streams are hindered by incompatibility among studies and across sites of analytical methods used to...
Federal Register 2010, 2011, 2012, 2013, 2014
2011-02-10
... DEPARTMENT OF LABOR Employment and Training Administration [TA-W-72,953] Matthews International Corporation, Bronze Division, Kingwood, WV; Notice of Affirmative Determination Regarding Application for.... Conclusion After careful review of the application, I conclude that the claim is of sufficient weight to...
Gold, Silver and Bronze Citations.
ERIC Educational Resources Information Center
American School & University, 2003
2003-01-01
Presents the gold, silver, and bronze winners of a competition, which judged the most outstanding learning environments at educational institutions nationwide. Jurors spent two days reviewing projects, focusing on concepts and ideas that made them exceptional. For each citation, the article offers information on the firm, client, total area, total…
Leaf drop affects herbivory in oaks.
Pearse, Ian S; Karban, Richard
2013-11-01
Leaf phenology is important to herbivores, but the timing and extent of leaf drop has not played an important role in our understanding of herbivore interactions with deciduous plants. Using phylogenetic general least squares regression, we compared the phenology of leaves of 55 oak species in a common garden with the abundance of leaf miners on those trees. Mine abundance was highest on trees with an intermediate leaf retention index, i.e. trees that lost most, but not all, of their leaves for 2-3 months. The leaves of more evergreen species were more heavily sclerotized, and sclerotized leaves accumulated fewer mines in the summer. Leaves of more deciduous species also accumulated fewer mines in the summer, and this was consistent with the idea that trees reduce overwintering herbivores by shedding leaves. Trees with a later leaf set and slower leaf maturation accumulated fewer herbivores. We propose that both leaf drop and early leaf phenology strongly affect herbivore abundance and select for differences in plant defense. Leaf drop may allow trees to dispose of their herbivores so that the herbivores must recolonize in spring, but trees with the longest leaf retention also have the greatest direct defenses against herbivores.
Genetic variation of the bronze locus (MC1R) in turkeys from Southern Brazil
Corso, Josmael; Hepp, Diego; Ledur, Mônica C.; Peixoto, Jane O.; Fagundes, Nelson J. R.; Freitas, Thales R. O.
2017-01-01
Abstract Domestic turkeys present several color phenotypes controlled by at least five genetic loci, but only one of these has been identified precisely: the bronze locus, which turned out to be the melanocortin-1 receptor (MC1R) gene. MC1R variation is important for breeders interested in maintaining or developing different color varieties. In this study, we sequenced most of the MC1R gene from 16 White Holland (the main commercial turkey variety) and 19 pigmented turkeys from southern Brazil with two purposes. The first was to describe the MC1R diversity in White Holland turkeys, which may serve as reservoirs of genetic diversity at this locus. The second was to test whether the traditional color classification used by Brazilian breeders is related to previously known MC1R alleles. White Holland turkeys had four different haplotypes corresponding to the bronze (b +) and black-winged bronze (b 1) alleles. Pigmented turkeys also had four haplotypes corresponding to the b + and b 1 alleles, but different haplotypes represent the most common b + allele in these two groups. The black (B) allele was absent from our samples. Overall, our results suggest that white and pigmented individuals form two different populations, and that the traditional color classification used by Brazilian breeders cannot accurately predict the genotypes at the bronze locus. PMID:28323301
Genetic variation of the bronze locus (MC1R) in turkeys from Southern Brazil.
Corso, Josmael; Hepp, Diego; Ledur, Mônica C; Peixoto, Jane O; Fagundes, Nelson J R; Freitas, Thales R O
2017-01-01
Domestic turkeys present several color phenotypes controlled by at least five genetic loci, but only one of these has been identified precisely: the bronze locus, which turned out to be the melanocortin-1 receptor (MC1R) gene. MC1R variation is important for breeders interested in maintaining or developing different color varieties. In this study, we sequenced most of the MC1R gene from 16 White Holland (the main commercial turkey variety) and 19 pigmented turkeys from southern Brazil with two purposes. The first was to describe the MC1R diversity in White Holland turkeys, which may serve as reservoirs of genetic diversity at this locus. The second was to test whether the traditional color classification used by Brazilian breeders is related to previously known MC1R alleles. White Holland turkeys had four different haplotypes corresponding to the bronze (b+) and black-winged bronze (b1) alleles. Pigmented turkeys also had four haplotypes corresponding to the b+ and b1 alleles, but different haplotypes represent the most common b+ allele in these two groups. The black (B) allele was absent from our samples. Overall, our results suggest that white and pigmented individuals form two different populations, and that the traditional color classification used by Brazilian breeders cannot accurately predict the genotypes at the bronze locus.
Punwong, Paramita; Juprasong, Yotin; Traiperm, Paweena
2017-09-01
This study investigated the short-term impacts of an oil spill on the leaf anatomical structures of Terminalia catappa L. from crude oil leakage in Rayong province, Thailand, in 2013. Approximately 3 weeks after the oil spill, leaves of T. catappa were collected along the coastline of Rayong from one affected site, five adjacent sites, and a control site. Slides of the leaf epidermis were prepared by the peeling method, while leaf and petiole transverse sections were prepared by paraffin embedding. Cell walls of adaxial epidermal cell on leaves in the affected site were straight instead of the jigsaw shape found in leaves from the adjacent and control sites. In addition, the stomatal index of the abaxial leaf surface was significantly lower in the affected site. Leaf and petiole transverse sections collected from the affected site showed increased cuticle thickness, epidermal cell diameter on both sides, and palisade mesophyll thickness; in contrast, vessel diameter and spongy mesophyll thickness were reduced. These significant changes in the leaf anatomy of T. catappa correspond with previous research and demonstrate the negative effects of oil spill pollution on plants. The anatomical changes of T. catappa in response to crude oil pollution are discussed as a possible indicator of pollution and may be used in monitoring crude oil pollution.
Zheng, Liang; Van Labeke, Marie-Christine
2017-01-01
Light quality critically affects plant development and growth. Development of light-emitting diodes (LEDs) enables the use of narrow band red and/or blue wavelengths as supplementary lighting in ornamental production. Yet, long periods under these wavelengths will affect leaf morphology and physiology. Leaf anatomy, stomatal traits, and stomatal conductance, leaf hydraulic conductance (Kleaf), and photosynthetic efficiency were investigated in three ornamental pot plants, namely Cordyline australis (monocot), Ficus benjamina (dicot, evergreen leaves), and Sinningia speciosa (dicot, deciduous leaves) after 8 weeks under LED light. Four light treatments were applied at 100 μmol m−2 s−1 and a photoperiod of 16 h using 100% red (R), 100% blue (B), 75% red with 25% blue (RB), and full spectrum white light (W), respectively. B and RB resulted in a greater maximum quantum yield (Fv/Fm) and quantum efficiency (ΦPSII) in all species compared to R and W and this correlated with a lower biomass under R. B increased the stomatal conductance compared with R. This increase was linked to an increasing stomatal index and/or stomatal density but the stomatal aperture area was unaffected by the applied light quality. Leaf hydraulic conductance (Kleaf) was not significantly affected by the applied light qualities. Blue light increased the leaf thickness of F. benjamina, and a relative higher increase in palisade parenchyma was observed. Also in S. speciosa, increase in palisade parenchyma was found under B and RB, though total leaf thickness was not affected. Palisade parenchyma tissue thickness was correlated to the leaf photosynthetic quantum efficiency (ΦPSII). In conclusion, the role of blue light addition in the spectrum is essential for the normal anatomical leaf development which also impacts the photosynthetic efficiency in the three studied species. PMID:28611818
Fischer, Ravit; Nitzan, Nadav; Chaimovitsh, David; Rubin, Baruch; Dudai, Nativ
2011-05-11
The aroma in sweet basil is a factor affecting the commercial value of the crop. In previous studies leaf age was considered to be a factor that influences the composition of essential oil (EO). In this study it was hypothesized that a single observation of the EO content in leaves from different positions on the main stem (young vs old) could predict the developmental changes in the plant during its life cycle. Plants harvested at week 16 demonstrated an exponential increase (R(2) = 0.92) in EO concentration in leaves on the main stem and lateral shoots, indicating higher EO concentrations in younger than in older leaves. Eugenol and methyleugenol predominated (28-77%) in the extract. Eugenol levels were higher in younger leaves (∼53%), and methyl-eugenol levels predominated in older leaves (∼68%). Linalool was lower in mature leaves than in younger leaves. This suggested that eugenol converted into methyleugenol and linalool decreased as leaf mature. However, in weekly monitored plants, the levels of these compounds in the EO had limited variation in the maturing leaf regardless of its position on the stem. This proposed that the EO composition in an individual leaf is mostly affected by the leaf position on the stem and not by its maturation process. Because leaf position is related to plant development, it is probable that the plant's physiological age at the time of leaf formation from the primordial tissue is the factor affecting the EO composition. It was concluded that interpretation of scientific observations should be carried out with caution and that hypotheses should be tested utilizing multifaceted approaches.
James Lashomb; Alan Iskra; Ann Brooks Gould; George Hamilton
2003-01-01
Bacterial leaf scorch (BLS) of amenity trees is caused by the bacterium Xylella fastidiosa, a xylem-limited pathogen that causes water stress resulting in leaf scorch, decline, and eventual death of affected trees. Recent surveys indicate that BLS is widespread throughout the eastern half of the United States. In New Jersey, BLS primarily affects red and pin oaks...
NASA Astrophysics Data System (ADS)
Janeček, Štěpán; Lepš, Jan
2005-09-01
The effects of litter removal, leaf cover of established plants and cover of basal internodes of a dominant species Molinia caerulea on seedling germination and the dynamics of established plants were studied in a field experiment in an oligotrophic wet meadow. Although the negative influence of litter on total seedling number and seedling species composition was non-significant, litter significantly affected the dynamics of the established vegetation and caused inhibition of total leaf cover development. The effects of total leaf cover of established plants on seedling establishment changed during the vegetation season. Whereas the effect of total leaf cover was positive at the start and in the middle of the vegetation season, at the end the total leaf cover negatively affected seedling establishment. Both total leaf cover and cover of basal internodes affected seedling composition. Effects of these two variables were statistically separable suggesting that they are based on different mechanisms. The response of seedling establishment to these factors was species specific and, consequently, our data support the hypothesis that that biotically generated spatial heterogeneity can promote species co-existence through the differentiation of species regeneration niches.
Clay Corner: Recreating Chinese Bronze Vessels.
ERIC Educational Resources Information Center
Gamble, Harriet
1998-01-01
Presents a lesson where students make faux Chinese bronze vessels through slab or coil clay construction after they learn about the history, function, and design of these vessels. Utilizes a variety of glaze finishes in order to give the vessels an aged look. Gives detailed guidelines for creating the vessels. (CMK)
Citizen Warrior: Major General Joshua Lawrence Chamberlain a Study in Command Leadership
2012-04-29
1999) p. 303. 44 Wallace, p. 68. 45 James E. Kelly, Generals in Bronze, ed. William B. Styple, read Patrick Cullen (Ashland, Blackstone Audio, Inc...Bronze, ed. William B. Styple, read Patrick Cullen. Ashland, Blackstone Audio, Inc., 2008. Knott, Steven W. “Knowledge Must Become Capability
15. A CLOSE UP VIEW OF ONE OF THE EXPOSED ...
15. A CLOSE UP VIEW OF ONE OF THE EXPOSED AGGREGATE LAMP STANDARDS, ITS BRONZE DECORATED FERRULE AND BRONZE LENS RECEIVER. - County Line Bridge, Spanning St. Joseph River at State Route 219, 0.6 mile south of U.S. Route 20, Osceola, St. Joseph County, IN
Federal Register 2010, 2011, 2012, 2013, 2014
2011-09-13
... DEPARTMENT OF STATE [Public Notice: 7579] Culturally Significant Objects Imported for Exhibition Determinations: ``Antico: The Golden Age of Renaissance Bronzes'' Summary: Notice is hereby given of the... exhibition ``Antico: The Golden Age of Renaissance Bronzes,'' imported from abroad for temporary exhibition...
46 CFR 162.017-3 - Materials, construction, and workmanship.
Code of Federal Regulations, 2010 CFR
2010-10-01
... serviceability. (b) Bodies of pressure-vacuum relief valves must be made of bronze or such corrosion-resistant..., and seats shall be made of bronze or such corrosion-resistant material as may be approved by the... springs shall be made of corrosion-resistant material. Springs plated with corrosion-resistant material...
USDA-ARS?s Scientific Manuscript database
Leafing out phenology affects a wide variety of ecosystem processes and ecological interactions, and it affects how natural and artificial ecosystems respond to different weather conditions in the spring. There is, however, relatively little information available on the factors affecting species dif...
DOE Office of Scientific and Technical Information (OSTI.GOV)
Lyons, P.C.; Evans, J.J.; Bacon, C.W.
Infection by the fungal endophyte Acremonium coenophialum affected the accumulation of inorganic and organic N in leaf blades and leaf sheaths of KY31 tall fescue (Festuca arundinacea Schreb.) grown under greenhouse conditions. Total soluble amino acid concentrations were increased in either the blade or sheath of the leaf from infected plants. A number of amino acids were significantly increased in the sheath, but only asparagine increased in the blade. Infection resulted in higher sheath NH{sub 4}{sup +} concentrations, whereas NO{sub 3}{sup {minus}} concentrations decreased in both leaf parts. The effects on amino acid, NO{sub 3}{sup {minus}}, and NH{sub 4}{sup +}more » concentrations were dependent upon the level of N fertilization and were usually apparent only at the high rate (10 millimolar) of application. Administration of {sup 14}CO{sub 2} to the leaf blades increased the accumulation of {sup 14}C in their amino acid fraction but not in the sheaths of infected plants. This may indicate that infection increased amino acid synthesis in the blade but that translocation to the sheath, which is the site of fungal colonization, was not affected. Glutamine synthetase activity was greater in leaf blades of infected plants at high and low N rates of fertilization, but nitrate reductase activity was not affected in either part of the leaf. Increased activities of glutamine synthetase together with the other observed changes in N accumulation and metabolism in endophyte-infected tall fescue suggest that NH{sub 4}{sup +} reassimilation could also be affected in the leaf blade.« less
Mineral resource of the month: tin
Carlin, James F.
2011-01-01
Tin was one of the earliest-known metals. Because of its hardening effect on copper, tin was used in bronze implements as early as 3500 B.C. Bronze, a copper-tin alloy that can be sharpened and is hard enough to retain a cutting edge, was used during the Bronze Age in construction tools as well as weapons for hunting and war. The geographical separation between tin-producing and tin-consuming nations greatly influenced the patterns of early trade routes. Historians think that as early as 1500 B.C., Phoenicians traveled by sea to the Cornwall district of England to obtain tin. The pure metal was not used unalloyed until about 600 B.C.
Symbolic Metal Bit and Saddlebag Fastenings in a Middle Bronze Age Donkey Burial
Bar-Oz, Guy; Nahshoni, Pirhiya; Motro, Hadas; Oren, Eliezer D.
2013-01-01
Here we report the unprecedented discovery of the skeleton of a ritually interred donkey with a metal horse bit in association with its teeth and saddlebag fastenings on its back. This discovery in the Middle Bronze Age III sacred precinct (1700/1650-1550 BCE) at Tel Haror, Israel, presents a unique combination of evidence for the early employment of equid harnessing equipment, both for chariot bridling (horse bit) and pack animals (saddlebags). The ritually deposited donkey with its unique accoutrements advances our understanding of the broad social and religious significance of equids in the Levantine Bronze Age, previously known mainly from textual and iconographical sources. PMID:23484046
Pérez-Pérez, José Manuel; Rubio-Díaz, Silvia; Dhondt, Stijn; Hernández-Romero, Diana; Sánchez-Soriano, Joaquín; Beemster, Gerrit T S; Ponce, María Rosa; Micol, José Luis
2011-12-01
Despite the large number of genes known to affect leaf shape or size, we still have a relatively poor understanding of how leaf morphology is established. For example, little is known about how cell division and cell expansion are controlled and coordinated within a growing leaf to eventually develop into a laminar organ of a definite size. To obtain a global perspective of the cellular basis of variations in leaf morphology at the organ, tissue and cell levels, we studied a collection of 111 non-allelic mutants with abnormally shaped and/or sized leaves, which broadly represent the mutational variations in Arabidopsis thaliana leaf morphology not associated with lethality. We used image-processing techniques on these mutants to quantify morphological parameters running the gamut from the palisade mesophyll and epidermal cells to the venation, whole leaf and rosette levels. We found positive correlations between epidermal cell size and leaf area, which is consistent with long-standing Avery's hypothesis that the epidermis drives leaf growth. In addition, venation parameters were positively correlated with leaf area, suggesting that leaf growth and vein patterning share some genetic controls. Positional cloning of the genes affected by the studied mutations will eventually establish functional links between genotypes, molecular functions, cellular parameters and leaf phenotypes. © 2011 Blackwell Publishing Ltd.
7 CFR 51.1145 - U.S. No. 1 Bronze.
Code of Federal Regulations, 2010 CFR
2010-01-01
...) Internal quality: Lots meeting the internal requirements for “U.S. Grade AA Juice (Double A)” or “U.S... 7 Agriculture 2 2010-01-01 2010-01-01 false U.S. No. 1 Bronze. 51.1145 Section 51.1145 Agriculture Regulations of the Department of Agriculture AGRICULTURAL MARKETING SERVICE (Standards, Inspections, Marketing...
46 CFR 193.10-10 - Fire hydrants and hose.
Code of Federal Regulations, 2010 CFR
2010-10-01
... decks where no protection is afforded to the hose in heavy weather, the hose may be temporarily removed... a garden hose nozzle that is bronze or metal with strength and corrosion resistance equivalent to... and couplings shall be as follows: (1) Couplings shall be of brass, bronze, or other equivalent metal...
DOE Office of Scientific and Technical Information (OSTI.GOV)
Mattox, Tracy M.; Koo, Bonil; Garcia, Guillermo
An electrochromic device includes a nanostructured transition metal oxide bronze layer that includes one or more transition metal oxide and one or more dopant, a solid state electrolyte, and a counter electrode. The nanostructured transition metal oxide bronze selectively modulates transmittance of near-infrared (NIR) spectrum and visible spectrum radiation as a function of an applied voltage to the device.
NASA Astrophysics Data System (ADS)
Mercuri, F.; Caruso, G.; Orazi, N.; Zammit, U.; Cicero, C.; Colacicchi Alessandri, O.; Ferretti, M.; Paoloni, S.
2018-05-01
In this paper, a new method based on the use of infrared thermography is proposed for the characterization of repairs and inserted parts on ancient bronzes. In particular, the quality of the contact between different kind of insertions and the main body of bronze statues is investigated by analysing the heat conduction process occurring across the interface between them. The thermographic results have been used to establish the nature of these inserted elements and the way they have been coupled to the main body of the statue during and after the manufacturing process. A model for the heat conduction based on the numerical finite elements method has been applied to compare the obtained results to the theoretical predictions. Measurements have been first carried out on test samples and then in the field on the Boxer at Rest (Museo Nazionale Romano in Rome), a masterpiece of the Greek Statuary, which contains a large variety of inserted items and repairs which are typical of the manufacturing process of bronze artefacts in general.
NASA Astrophysics Data System (ADS)
Zhang, N.; Dong, G.; Yang, X.; Zuo, X.; Kang, L.; Ren, L.; Liu, H.; Li, H.; Min, R.; Liu, X.; Zhang, D.; Chen, F.
2017-12-01
The extracted microfossils from the dental calculus of ancient teeth are a new form of archaeological evidence which can provide direct information on the plant diet of a population. Here, we present the results of analyses of starch grains and phytoliths trapped in the dental calculus of humans who occupied the Bronze Age site of Shilinggang ( 2500 cal yr BP) in Yunnan Province, southwestern China. The results demonstrate that the inhabitants consumed a wide range of plants, including rice, millet, and palms, together with other food plants which have not previously been detected in Yunnan. The discovery of various underground storage organs (USOs; tubers, roots, bulbs, and rhizomes) and acorns complements the application of conventional macrofossil and isotope studies to understand the diet of the Bronze Age human population of Yunnan. The wide variety of plant foods consumed suggests that the inhabitants adopted a broad-spectrum strategy of gathering food and cultivating crops in northwest Yunnan Province in the late Bronze Age at a time when agricultural societies were developed in the central plains of China.
Band structure of the quasi two-dimensional purple molybdenum bronze
NASA Astrophysics Data System (ADS)
Guyot, H.; Balaska, H.; Perrier, P.; Marcus, J.
2006-09-01
The molybdenum purple bronze KMo 6O 17 is quasi two-dimensional (2D) metallic oxide that shows a Peierls transition towards a metallic charge density wave state. Since this specific transition is directly related to the electron properties of the normal state, we have investigated the electronic structure of this bronze at room temperature. The shape of the Mo K1s absorption edge reveals the presence of distorted MoO 6 octahedra in the crystallographic structure. Photoemission experiments evidence a large conduction band, with a bandwidth of 800 meV and confirm the metallic character of this bronze. A wide depleted zone separates the conduction band from the valence band that exhibits a fourfold structure, directly connected to the octahedral symmetry of the Mo sites. The band structure is determined by ARUPS in two main directions of the (0 0 1) Brillouin zone. It exhibits some unpredicted features but corroborates the earlier theoretical band structure and Fermi surface. It confirms the hidden one-dimensionality of KMo 6O 17 that has been proposed to explain the origin of the Peierls transition in this 2D compound.
Escalation of a coevolutionary arms race through host rejection of brood parasitic young.
Langmore, Naomi E; Hunt, Sarah; Kilner, Rebecca M
2003-03-13
Cuckoo nestlings that evict all other young from the nest soon after hatching impose a high reproductive cost on their hosts. In defence, hosts have coevolved strategies to prevent brood parasitism. Puzzlingly, they do not extend beyond the egg stage. Thus, hosts adept at recognizing foreign eggs remain vulnerable to exploitation by cuckoo nestlings. Here we show that the breach of host egg defences by cuckoos creates a new stage in the coevolutionary cycle. We found that defences used during the egg-laying period by host superb fairy-wrens (Malurus cyaneus) are easily evaded by the Horsfield's bronze-cuckoo (Chrysococcyx basalis), a specialist fairy-wren brood parasite. However, although hosts never deserted their own broods, they later abandoned 40% of nests containing a lone Horsfield's bronze-cuckoo nestling, and 100% of nests with a lone shining bronze-cuckoo nestling (Chrysococcyx lucidus), an occasional fairy-wren brood parasite. Our experiments demonstrate that host discrimination against evictor-cuckoo nestlings is possible, and suggest that it has selected for the evolution of nestling mimicry in bronze-cuckoos.
NASA Astrophysics Data System (ADS)
Deng, H.; Chen, G. Y.; Zhou, C.; Zhou, X. C.; He, J.; Zhang, Y.
2014-09-01
A series of theoretical analyses and experimental investigations were performed to examine a pulsed fiber-laser tangential profiling and radial sharpening technique for bronze-bonded diamond grinding wheels. The mechanisms for the pulsed laser tangential profiling and radial sharpening of grinding wheels were theoretically analyzed, and the four key processing parameters that determine the quality, accuracy, and efficiency of pulsed laser dressing, namely, the laser power density, laser spot overlap ratio, laser scanning track line overlap ratio, and number of laser scanning cycles, were proposed. Further, by utilizing cylindrical bronze wheels (without diamond grains) and bronze-bonded diamond grinding wheels as the experimental subjects, the effects of these four processing parameters on the removal efficiency and the surface smoothness of the bond material after pulsed laser ablation, as well as the effects on the contour accuracy of the grinding wheels, the protrusion height of the diamond grains, the sharpness of the grain cutting edges, and the graphitization degree of the diamond grains after pulsed laser dressing, were explored. The optimal values of the four key processing parameters were identified.
Maize YABBY Genes drooping leaf1 and drooping leaf2 Regulate Plant Architecture[OPEN
Briggs, Sarah; Bradbury, Peter J.
2017-01-01
Leaf architecture directly influences canopy structure, consequentially affecting yield. We discovered a maize (Zea mays) mutant with aberrant leaf architecture, which we named drooping leaf1 (drl1). Pleiotropic mutations in drl1 affect leaf length and width, leaf angle, and internode length and diameter. These phenotypes are enhanced by natural variation at the drl2 enhancer locus, including reduced expression of the drl2-Mo17 allele in the Mo17 inbred. A second drl2 allele, produced by transposon mutagenesis, interacted synergistically with drl1 mutants and reduced drl2 transcript levels. The drl genes are required for proper leaf patterning, development and cell proliferation of leaf support tissues, and for restricting auricle expansion at the midrib. The paralogous loci encode maize CRABS CLAW co-orthologs in the YABBY family of transcriptional regulators. The drl genes are coexpressed in incipient and emergent leaf primordia at the shoot apex, but not in the vegetative meristem or stem. Genome-wide association studies using maize NAM-RIL (nested association mapping-recombinant inbred line) populations indicated that the drl loci reside within quantitative trait locus regions for leaf angle, leaf width, and internode length and identified rare single nucleotide polymorphisms with large phenotypic effects for the latter two traits. This study demonstrates that drl genes control the development of key agronomic traits in maize. PMID:28698237
Saini, Kumud; Markakis, Marios N.; Zdanio, Malgorzata; Balcerowicz, Daria M.; Beeckman, Tom; De Veylder, Lieven; Prinsen, Els; Beemster, Gerrit T. S.; Vissenberg, Kris
2017-01-01
In plants many developmental processes are regulated by auxin and its directional transport. PINOID (PID) kinase helps to regulate this transport by influencing polar recruitment of PIN efflux proteins on the cellular membranes. We investigated how altered auxin levels affect leaf growth in Arabidopsis thaliana. Arabidopsis mutants and transgenic plants with altered PID expression levels were used to study the effect on auxin distribution and leaf development. Single knockouts showed small pleiotropic growth defects. Contrastingly, several leaf phenotypes related to changes in auxin concentrations and transcriptional activity were observed in PID overexpression (PIDOE) lines. Unlike in the knockout lines, the leaves of PIDOE lines showed an elevation in total indole-3-acetic acid (IAA). Accordingly, enhanced DR5-visualized auxin responses were detected, especially along the leaf margins. Kinematic analysis revealed that ectopic expression of PID negatively affects cell proliferation and expansion rates, yielding reduced cell numbers and small-sized cells in the PIDOE leaves. We used PIDOE lines as a tool to study auxin dose effects on leaf development and demonstrate that auxin, above a certain threshold, has a negative affect on leaf growth. RNA sequencing further showed how subtle PIDOE-related changes in auxin levels lead to transcriptional reprogramming of cellular processes. PMID:28659952
Matsuda, Ikki; Clauss, Marcus; Tuuga, Augustine; Sugau, John; Hanya, Goro; Yumoto, Takakazu; Bernard, Henry; Hummel, Jürgen
2017-01-01
Free-living animals must make dietary choices in terms of chemical and physical properties, depending on their digestive physiology and availability of food resources. Here we comprehensively evaluated the dietary choices of proboscis monkeys (Nasalis larvatus) consuming young leaves. We analysed the data for leaf toughness and digestibility measured by an in vitro gas production method, in addition to previously reported data on nutrient composition. Leaf toughness, in general, negatively correlated with the crude protein content, one of the most important nutritional factors affecting food selection by leaf-eating primates. This result suggests that leaf toughness assessed by oral sensation might be a proximate cue for its protein content. We confirmed the importance of the leaf chemical properties in terms of preference shown by N. larvatus; leaves with high protein content and low neutral detergent fibre levels were preferred to those of the common plant species. We also found that these preferred leaves were less tough and more digestible than the alternatives. Our in vitro results also suggested that N. larvatus were little affected by secondary plant compounds. However, the spatial distribution pattern of plant species was the strongest factor explaining the selection of the preferred leaf species. PMID:28211530
Leaf traits and herbivory levels in a tropical gymnosperm, Zamia stevensonii (Zamiaceae).
Prado, Alberto; Sierra, Adriel; Windsor, Donald; Bede, Jacqueline C
2014-03-01
Slow-growing understory cycads invest heavily in defenses to protect the few leaves they produce annually. The Neotropical cycad Zamia stevensonii has chemical and mechanical barriers against insect herbivores. Mechanical barriers, such as leaf toughness, can be established only after the leaf has expanded. Therefore, chemical defenses may be important during leaf expansion. How changes in leaf traits affect the feeding activity of cycad specialist insects is unknown. We investigated leaf defenses and incidence of specialist herbivores on Z. stevensonii during the first year after leaf flush. Herbivore incidence, leaf production, and leaf traits that might affect herbivory-including leaf age, lamina thickness, resistance-to-fracture, work-to-fracture, trichome density, and chlorophyll, water, and toxic azoxyglycoside (AZG) content-were measured throughout leaf development. Principal component analysis and generalized linear models identified characteristics that may explain herbivore incidence. Synchronized leaf development in Z. stevensonii is characterized by quick leaf expansion and delayed greening. Specialist herbivores feed on leaves between 10 and 100 d after flush and damage ∼37% of all leaflets produced. Young leaves are protected by AZGs, but these defenses rapidly decrease as leaves expand. Leaves older than 100 d are protected by toughness. Because AZG concentrations drop before leaves become sufficiently tough, there is a vulnerable period during which leaves are susceptible to herbivory by specialist insects. This slow-growing gymnosperm invests heavily in constitutive defenses against highly specialized herbivores, underlining the convergence in defensive syndromes by major plant lineages.
Neuwirthová, Eva; Lhotáková, Zuzana; Albrechtová, Jana
2017-01-01
The aims of the study were: (i) to compare leaf reflectance in visible (VIS) (400–700 nm), near-infrared (NIR) (740–1140 nm) and short-wave infrared (SWIR) (2000–2400 nm) spectral ranges measured monthly by a contact probe on a single leaf and a stack of five leaves (measurement setup (MS)) of two broadleaved tree species during the vegetative season; and (ii) to test if and how selected vegetation indices differ under these two MS. In VIS, the pigment-related spectral region, the effect of MS on reflectance was negligible. The major influence of MS on reflectance was detected in NIR (up to 25%), the structure-related spectral range; and weaker effect in SWIR, the water-related spectral range. Vegetation indices involving VIS wavelengths were independent of MS while indices combining wavelengths from both VIS and NIR were MS-affected throughout the season. The effect of leaf stacking contributed to weakening the correlation between the leaf chlorophyll content and selected vegetation indices due to a higher leaf mass per area of the leaf sample. The majority of MS-affected indices were better correlated with chlorophyll content in both species in comparison with MS-unaffected indices. Therefore, in terms of monitoring leaf chlorophyll content using the contact probe reflectance measurement, these MS-affected indices should be used with caution, as discussed in the paper. If the vegetation indices are used for assessment of plant physiological status in various times of the vegetative season, then it is essential to take into consideration their possible changes induced by the particular contact probe measurement setup regarding the leaf stacking. PMID:28538685
Alternate alloys for UF/sub 6/ cylinder valve packing nuts
DOE Office of Scientific and Technical Information (OSTI.GOV)
Blue, S.C.
One-inch cylinder valve packing nuts made from aluminum bronze alloy C63600 are susceptible to stress corrosion cracking. The use of alloys which are more resistant to cracking should eliminate the occasional nut failures. It is proposed that packing nuts be produced from either aluminum bronze C61300 or Monel (NO4400). 2 refs., 4 tabs.
Noncyanide Stripper Placement Program. Phase 1
1989-05-01
bronze (brazing material ) from low-alloy steels , heat and corrosion resistant...STRIPPERS AND BASIS MATERIALS FROM WHICH THE COATINGS ARE REMOVED (FROM T.O.42C2-1-7) Surface Coating Basis Material Brass Low-Alloy Steels Bronze Low...braze materials , low alloy steels , and heat and corrosion resistant steels . Additional tests were performed on three masking materials routinely
Federal Register 2010, 2011, 2012, 2013, 2014
2013-09-16
... amended by CBP Dec. 08-40, are due to expire on September 19, 2013, unless extended. The Assistant...: Effective Date: September 19, 2013. FOR FURTHER INFORMATION CONTACT: For legal aspects, Lisa Burley, Cargo... until September 19, 2013, and amended them to include archaeological material from the Bronze Age...
De Maeseneer, M; Buls, N; Cleeren, N; Lenchik, L; De Mey, J
2006-01-01
We present an unusual application of multidetector CT and shaded surface rendering in the investigation of a soil sample, containing an ancient Roman bronze bowl. The CT findings were of fundamental importance in helping the archaeologists study the bronze bowl from the soil sample.
Ancient Chinese Bronzes: Teacher's Packet.
ERIC Educational Resources Information Center
Smithsonian Institution, Washington, DC. Arthur M. Sackler Gallery.
The focus of this teacher's packet is the bronze vessels made for the kings and great families of the early Chinese dynasties between 1700 B.C. and 200 A.D. The materials in the guide are intended for use by teachers and students visiting the exhibition, "The Arts of China," at the Arthur M. Sackler Gallery of the Smithsonian Institution…
Investigation of archaeological metal artefacts by laser-induced breakdown spectroscopy (LIBS)
NASA Astrophysics Data System (ADS)
Tankova, V.; Malcheva, G.; Blagoev, K.; Leshtakov, L.
2018-03-01
In this work, laser-induced breakdown spectroscopy was applied to determining the elemental composition of a set of ancient bronze artefacts dated from the Late Bronze Age and Early Iron Age (14th – 10th century BC). We used a Nd:YAG laser at 1064 nm with pulse duration of 10 ns and energy of 10 mJ and determined the elemental composition of the bronze alloy that was used in manufacturing the samples under study. The concentrations of tin and lead in the bulk of the examined materials was estimated after generating calibration curves for a set of four standard samples. The preliminary results of the analysis will provide information on the artefacts provenance and on the production process.
NASA Astrophysics Data System (ADS)
Ryashin, N. S.; Malikov, A. G.; Shikalov, V. S.; Gulyaev, I. P.; Kuchumov, B. M.; Klinkov, S. V.; Kosarev, V. F.; Orishich, A. M.
2017-10-01
The paper presents results of the cold spraying of aluminum bronze coatings on substrates profiled with WC/Ni tracks obtained by laser cladding. Reinforcing cermet frames shaped as grids with varied mesh sizes were clad on stainless steel substrates using a CO2 laser machine "Siberia" (ITAM SB RAS, Russia). As a result, surfaces/substrates with heterogeneous shape, composition, and mechanical properties were obtained. Aluminum bronze coatings were deposited from 5lF-NS powder (Oerlikon Metco, Switzerland) on those substrates using cold spraying equipment (ITAM SB RAS). Data of profiling, microstructure diagnostics, EDS analysis, and mechanical tests of obtained composites is reported. Surface relief of the sprayed coatings dependence on substrate structure has been demonstrated.
Marcelo Ard& #243; n; Catherine M. Pringle; Susan L. Eggert
2009-01-01
Comparisons of the effects of leaf litter chemistry on leaf breakdown rates in tropical vs temperate streams are hindered by incompatibility among studies and across sites of analytical methods used to measure leaf chemistry. We used standardized analytical techniques to measure chemistry and breakdown rate of leaves from common riparian tree species at 2 sites, 1...
Guo, Wei Hong; Wang, Hua; Yu, Mu Kui; Wu, Tong Gui; Han, You Zhi
2017-03-18
We analyzed the rules of Metasequoia glyptostroboides along with latitude, including leaf length, leaf width, leaf perimeter, leaf area, ratio of leaf length to width, specific leaf area (SLA), and leaf dry mass based on eight stands growing at different latitudes in the coastal area of eastern China, as well as their relationships with climatic and soil factors. The results showed that the leaf length, leaf width and leaf perimeter increased with increasing latitude, while the leaf area and SLA firstly increased and then decreased. The mean annual temperature and annual precipitation were the major environmental factors affecting the leaf traits along latitude gradient. With the increase of soil N content, the SLA decreased firstly and then increased, while the leaf mass decreased significantly. With the increase of soil P content, the SLA increased, and the leaf mass decreased significantly.
UV radiation is the primary factor driving the variation in leaf phenolics across Chinese grasslands
Chen, Litong; Niu, Kechang; Wu, Yi; Geng, Yan; Mi, Zhaorong; Flynn, Dan FB; He, Jin-Sheng
2013-01-01
Due to the role leaf phenolics in defending against ultraviolet B (UVB) under previously controlled conditions, we hypothesize that ultraviolet radiation (UVR) could be a primary factor driving the variation in leaf phenolics in plants over a large geographic scale. We measured leaf total phenolics, ultraviolet-absorbing compounds (UVAC), and corresponding leaf N, P, and specific leaf area (SLA) in 151 common species. These species were from 84 sites across the Tibetan Plateau and Inner Mongolian grasslands of China with contrasting UVR (354 vs. 161 mW/cm2 on average). Overall, leaf phenolics and UVAC were all significantly higher on the Tibetan Plateau than in the Inner Mongolian grasslands, independent of phylogenetic relationships between species. Regression analyses showed that the variation in leaf phenolics was strongly affected by climatic factors, particularly UVR, and soil attributes across all sites. Structural equation modeling (SEM) identified the primary role of UVR in determining leaf phenolic concentrations, after accounting for colinearities with altitude, climatic, and edaphic factors. In addition, phenolics correlated positively with UVAC and SLA, and negatively with leaf N and N: P. These relationships were steeper in the lower-elevation Inner Mongolian than on the Tibetan Plateau grasslands. Our data support that the variation in leaf phenolics is controlled mainly by UV radiation, implying high leaf phenolics facilitates the adaptation of plants to strong irradiation via its UV-screening and/or antioxidation functions, particularly on the Tibetan Plateau. Importantly, our results also suggest that leaf phenolics may influence on vegetation attributes and indirectly affect ecosystem processes by covarying with leaf functional traits. PMID:24363898
Earthquake and the Catastrophic End of the Late Bronze Age in the Eastern Mediterranean
NASA Astrophysics Data System (ADS)
Nur, A.
2009-04-01
The reasons for the catastrophic and wide spread political as well as physical collapse in the Aegean and Eastern Mediterranean areas that define the end of the Bronze age ca. 1225 BC to 1175 BC remain a major enigma. It has been attributed by historian to attacks by outsiders with the most favored group being the (enigmatic) so-called sea people. Unfortunately there is no real evidence for this. However combined geological, geophysical and archaeological evidence suggests that earthquakes may have played a key role in this extraordinary collapse during the late 13th and early 12th centuries . Based on the instrumentally recorded earthquakes occurring in the Aegean and Eastern Mediterranean region during the 20th century, several events that have clear historical information, and the geography of seismically active faults it is obvious that numerous earthquakes of magnitude 6·5 or greater (enough to destroy modern buildings, let alone those of antiquity) occurred here frequently in the past. Furthermore major earthquakes often occur in this region in groups, known as ‘‘sequences'' or ‘‘storms'', in which one large quake is followed days, months, or a few years later by others elsewhere on the plate boundary fault lines. When a map of the areas in the Aegean and Eastern Mediterranean region shaken by 20th century earthquakes of magnitude 6·5 and greater and with an intensity of VII or greater is overlaid on Robert Drews' map of sites destroyed in these same regions during the so-called ‘‘Catastrophe'' near the end of the Late Bronze Age, it is readily apparent that virtually all of these LBA sites lie within the affected (‘‘high-shaking'') areas. This would suggest that a major ‘‘earthquake storm'' may have occurred in the Late Bronze Age Aegean and Eastern Mediterranean during the years 1225-1175 . This ‘‘storm'' may have interacted with societal, political and economic forces at work in these areas c. 1200 and merits consideration by archaeologists and prehistorians. Similarly several other unexplained civilization collapses may also be linked to catastrophic earthquakes such as the collapse of the Cassas Grandes civilization in Sonora, Mexico, and the Indus valley civilization,
Altinoz, M A; Ince, B; Sav, A; Dincer, A; Cengiz, S; Mercan, S; Yazici, Z; Bilgen, M N
2014-02-01
Undecomposed human bodies and organs always attracted interest in terms of understanding biological tissue stability and immortality. Amongst these, cases of natural mummification found in glaciers, bog sediments and deserts caused even more attention. In 2010, an archeological excavation of a Bronze Age layer in a tumulus near the Western Anatolia city Kütahya revealed fire affected regions with burnt human skeletons and charred wooden objects. Inside of the cracked skulls, undecomposed brains were discernible. To analyze the burial taphonomy of the rare phenomenon of brain preservation, we analyzed brains, bone, teeth and surrounding soils elements using Inductively Coupled Plasma-Mass Spectrometer (ICP-MS). Adipocere formation or saponification of postmortem tissue fat requires high levels of alkalinity and especially potassium. Indeed, ICP-MS analysis of the brain, teeth and bone and also of the surrounding soil revealed high levels of potassium, magnesium, aluminum and boron, which are compatible with the famous role of Kütahya in tile production with its soil containing high level of alkalines and tile-glazing boron. Fatty acid chromatography revealed simultaneous saturation of fats and protection of fragile unsaturated fatty acids consistent with soil-presence of both pro-oxidant and anti-oxidant trace metals. Computerized tomography revealed protection of diencephalic, metencephalic and occipital tissue in one of the best-preserved specimens. Boron was previously found as an intentional preservative of Tutankhamen and Deir el Bahari mummies. Here, in natural soil with its insect-repellant, anti-bacterial and fire-resistance qualities it may be a factor to preserve heat-affected brains as almost bioporcellain specimens. Copyright © 2013 Elsevier GmbH. All rights reserved.
Nielsen, David G; Muilenburg, Vanessa L; Herms, Daniel A
2011-06-01
Bronze birch borer (Agrilus anxius Gory) is the key pest of birches (Betula spp.) in North America, several of which have been recommended for ornamental landscapes based on anecdotal reports of borer resistance that had not been confirmed experimentally. In a 20-yr common garden experiment initiated in 1979 in Ohio, North American birch species, including paper birch (Betula papyrifera Marshall), 'Whitespire' gray birch (Betula populifolia Marshall), and river birch (Betula nigra L.), were much more resistant to bronze birch borer than species indigenous to Europe and Asia, including European white birch (Betula pendula Roth), downy birch (Betula pubescens Ehrh.), monarch birch (Betula maximowicziana Regel), and Szechuan white birch (Betula szechuanica Jansson). Within 8 yr of planting, every European white, downy, and Szechuan birch had been colonized and killed, although 100% of monarch birch had been colonized and 88% of these plants were killed after nine years. Conversely, 97% of river birch, 76% of paper birch, and 73% Whitespire gray birch were alive 20 yr after planting, and river birch showed no evidence of colonization. This pattern is consistent with biogeographic theory of plant defense: North American birch species that share a coevolutionary history with bronze birch borer were much more resistant than naïve hosts endemic to Europe and Asia, possibly by virtue of evolution of targeted defenses. This information suggests that if bronze birch borer were introduced to Europe or Asia, it could threaten its hosts there on a continental scale. This study also exposed limitations of anecdotal observation as evidence of host plant resistance.
Ancient X chromosomes reveal contrasting sex bias in Neolithic and Bronze Age Eurasian migrations.
Goldberg, Amy; Günther, Torsten; Rosenberg, Noah A; Jakobsson, Mattias
2017-03-07
Dramatic events in human prehistory, such as the spread of agriculture to Europe from Anatolia and the late Neolithic/Bronze Age migration from the Pontic-Caspian Steppe, can be investigated using patterns of genetic variation among the people who lived in those times. In particular, studies of differing female and male demographic histories on the basis of ancient genomes can provide information about complexities of social structures and cultural interactions in prehistoric populations. We use a mechanistic admixture model to compare the sex-specifically-inherited X chromosome with the autosomes in 20 early Neolithic and 16 late Neolithic/Bronze Age human remains. Contrary to previous hypotheses suggested by the patrilocality of many agricultural populations, we find no evidence of sex-biased admixture during the migration that spread farming across Europe during the early Neolithic. For later migrations from the Pontic Steppe during the late Neolithic/Bronze Age, however, we estimate a dramatic male bias, with approximately five to 14 migrating males for every migrating female. We find evidence of ongoing, primarily male, migration from the steppe to central Europe over a period of multiple generations, with a level of sex bias that excludes a pulse migration during a single generation. The contrasting patterns of sex-specific migration during these two migrations suggest a view of differing cultural histories in which the Neolithic transition was driven by mass migration of both males and females in roughly equal numbers, perhaps whole families, whereas the later Bronze Age migration and cultural shift were instead driven by male migration, potentially connected to new technology and conquest.
Ren, Lele; Li, Xin; Kang, Lihong; Brunson, Katherine; Liu, Honggao; Dong, Weimiao; Li, Haiming; Min, Rui; Liu, Xu; Dong, Guanghui
2017-01-01
Reconstructing ancient diets and the use of animals and plants augment our understanding of how humans adapted to different environments. Yunnan Province in southwest China is ecologically and environmentally diverse. During the Neolithic and Bronze Age periods, this region was occupied by a variety of local culture groups with diverse subsistence systems and material culture. In this paper, we obtained carbon (δ13C) and nitrogen (δ15N) isotopic ratios from human and faunal remains in order to reconstruct human paleodiets and strategies for animal exploitation at the Bronze Age site of Shilinggang (ca. 2500 Cal BP) in northwest Yunnan Province. The δ13C results for human samples from Shilinggang demonstrate that people's diets were mainly dominated by C3-based foodstuffs, probably due to both direct consumption of C3 food and as a result of C3 foddering of consumed animals. Auxiliary C4 food signals can also be detected. High δ15N values indicate that meat was an important component of the diet. Analysis of faunal samples indicates that people primarily fed pigs and dogs with human food waste, while sheep/goats and cattle were foddered with other food sources. We compare stable isotope and archaeobotanical data from Shilinggang with data from other Bronze Age sites in Yunnan to explore potential regional variation in subsistence strategies. Our work suggests that people adopted different animal utilization and subsistence strategies in different parts of Yunnan during the Bronze Age period, probably as local adaptations to the highly diversified and isolated environments in the region.
Ren, Lele; Li, Xin; Kang, Lihong; Brunson, Katherine; Liu, Honggao; Dong, Weimiao; Li, Haiming; Min, Rui; Liu, Xu
2017-01-01
Reconstructing ancient diets and the use of animals and plants augment our understanding of how humans adapted to different environments. Yunnan Province in southwest China is ecologically and environmentally diverse. During the Neolithic and Bronze Age periods, this region was occupied by a variety of local culture groups with diverse subsistence systems and material culture. In this paper, we obtained carbon (δ13C) and nitrogen (δ15N) isotopic ratios from human and faunal remains in order to reconstruct human paleodiets and strategies for animal exploitation at the Bronze Age site of Shilinggang (ca. 2500 Cal BP) in northwest Yunnan Province. The δ13C results for human samples from Shilinggang demonstrate that people’s diets were mainly dominated by C3-based foodstuffs, probably due to both direct consumption of C3 food and as a result of C3 foddering of consumed animals. Auxiliary C4 food signals can also be detected. High δ15N values indicate that meat was an important component of the diet. Analysis of faunal samples indicates that people primarily fed pigs and dogs with human food waste, while sheep/goats and cattle were foddered with other food sources. We compare stable isotope and archaeobotanical data from Shilinggang with data from other Bronze Age sites in Yunnan to explore potential regional variation in subsistence strategies. Our work suggests that people adopted different animal utilization and subsistence strategies in different parts of Yunnan during the Bronze Age period, probably as local adaptations to the highly diversified and isolated environments in the region. PMID:28531221
Bioarchaeology of adaptation to a marginal environment in bronze age Western China.
Berger, Elizabeth; Wang, Hui
2017-07-08
This study examines human adaptation to the 4000 BP climate change event, which is said to have increased the marginality of Inner Asian environments. We propose to define "marginal" environments not in relation to a specific economic activity (e.g., agriculture), but in relation to whether humans living there are physiologically stressed. Three sites in the Hexi Corridor of Gansu were studied, one from the early and two from the late Bronze Age (N = 125). The study includes three indicators of physiological stress: linear enamel hypoplasias (LEH); tibial periosteal lesions; and fertility. The early and late Bronze Age groups were compared to examine whether human physiological stress increased. The percent of individuals with LEH declined dramatically, indicating fewer growth disruptions. Tibial periosteal reactions also changed, from mostly active to mostly healing at the time of death, indicating that frailty declined. Fertility, which is sensitive to changes in population health and resource availability, did not change significantly. Counter to the dominant narrative of environmental deterioration and subsistence system collapse, the Bronze Age residents of the Hexi Corridor show no skeletal evidence that they suffered from resource shortages or struggled to adapt in the fluctuating climate that pertained after the 4000 BP climate event. In fact, this study found that people suffered from less frailty and fewer growth disruptions after the unstable climate had persisted for some time. Therefore, in human biological terms, the Hexi Corridor did not become more marginal for human habitation during the Bronze Age. © 2017 Wiley Periodicals, Inc.
Stoepler, Teresa M; Lill, John T
2013-10-01
A variety of ecological factors influence host use by parasitoids, including both abiotic and biotic factors. Light environment is one important abiotic parameter that differs among habitats and influences a suite of plant nutritional and resistance traits that in turn affect herbivore performance. However, the extent to which these bottom-up effects "cascade up" to affect higher trophic levels and the relative importance of direct and indirect effects of sunlight on tritrophic interactions are unclear. The objective of this study was to test how light environment (light gap vs. shaded forest understory) and leaf type (sun vs. shade leaves) affect the performance and incidence of parasitism of two species of moth larvae, Euclea delphinii and Acharia stimulea (Limacodidae). We manipulated the leaf phenotype of potted white oak saplings by growing them in either full sun or full shade throughout leaf expansion to produce sun and shade leaves, respectively. These saplings were then placed in light gap and adjacent shaded understory habitats in the forest in a full-factorial design, and stocked with sentinel larvae that were exposed to parasitism ("exposed" experiments). We reared additional larvae in sleeve cages (protected from parasitism) to isolate light environment and leaf phenotype treatment effects on larval performance in the absence of enemies ("bagged" experiments). In the exposed experiments, light environment strongly affected the likelihood of parasitism, while leaf phenotype did not. Euclea delphinii larvae were up to 6.6 times more likely to be parasitized in light gaps than in shaded understory habitats. This pattern was consistent for both tachinid fly and wasp parasitoids across two separate experiments. However, the larval performance of both species in the bagged experiments was maximized in the shade-habitat/sun-leaf treatment, a habitat/leaf-type combination that occurs infrequently in nature. Taken together, our results suggest that the direct effects of light environment on the incidence of parasitism supersede any indirect effects resulting from altered leaf quality and reveal inherent ecological trade-offs for herbivores confronted with choosing between sunny (high leaf quality, harsh environment, high parasitism) and shaded (reduced leaf quality less harsh environment, reduced parasitism) habitats.
ERIC Educational Resources Information Center
Donais, Mary Kate; Whissel, Greg; Dumas, Ashley; Golden, Kathleen
2009-01-01
A unique, interdisciplinary collaboration between chemistry and classics has led to the development of an experiment for nonscience majors. This instrumental analysis experiment was designed for use in an archaeology course to quantify the amount of lead in ancient bronze coins. The coins were corroded beyond visual identification, so provenance…
Exposition and Synthesis of Benin Bronze Casting: Emphasis on the Olotan Casters of Benin
ERIC Educational Resources Information Center
Ifeta, Chris Funke
2016-01-01
The introduction of Western education to Nigeria has brought in its wake great strides toward development. Changes in Benin dates far back to the dawn of the 20th century. This paper investigates the critical role of education in development. The paper integrates interview data collected from bronze casters in Benin. The first section of the paper…
Enhancement of bronze alloy surface properties by FSP second-phase particle incorporation
Ajayi, O. O.; Lorenzo-Martin, Cinta
2017-06-15
This study presents results of an experimental study to evaluate friction stir processing (FSP) with and without hard second-phase particle incorporation as a means to enhance surface properties and wear performance of C86300 manganese bronze alloy. FSP of flat bronze alloy specimens was conducted with hardened H-13 tool steel to create a 3-mm-thick processed surface layer. The process was also used to incorporate B 4C particles, thereby creating a metal-matrix composite layer on the alloy surface. FSP alone was observed to produce substantial reduction in grain size (from an initial value of 350 mu m to 1-5 μm). FSP withoutmore » particle incorporation resulted in modest surface hardening due to grain refinement and dispersion hardening. Under lubricated contact in block-on-ring testing with a hardened steel counter face, FSP produced substantial reduction (about 3X) in bronze wear after polishing of processing surface roughening. FSP with hard B 4C second-phase particle incorporation further reduced wear by up to 20X. The improvement in wear behavior is attributed to grain refinement and load shielding by second-phase particles, as determined by wear mechanism analysis.« less
Higham, Charles F W; Douka, Katerina; Higham, Thomas F G
2015-01-01
There are two models for the origins and timing of the Bronze Age in Southeast Asia. The first centres on the sites of Ban Chiang and Non Nok Tha in Northeast Thailand. It places the first evidence for bronze technology in about 2000 B.C., and identifies the origin by means of direct contact with specialists of the Seima Turbino metallurgical tradition of Central Eurasia. The second is based on the site of Ban Non Wat, 280 km southwest of Ban Chiang, where extensive radiocarbon dating places the transition into the Bronze Age in the 11th century B.C. with likely origins in a southward expansion of technological expertise rooted in the early states of the Yellow and Yangtze valleys, China. We have redated Ban Chiang and Non Nok Tha, as well as the sites of Ban Na Di and Ban Lum Khao, and here present 105 radiocarbon determinations that strongly support the latter model. The statistical analysis of the results using a Bayesian approach allows us to examine the data at a regional level, elucidate the timing of arrival of copper base technology in Southeast Asia and consider its social impact.
NASA Astrophysics Data System (ADS)
Brunetti, Antonio; Depalmas, Anna; di Gennaro, Francesco; Serges, Alessandra; Schiavon, Nicola
2016-07-01
The chemical composition of a unique bronze artifact known as the "Cesta" ("Basket") belonging to the ancient Nuragic civilization of the Island of Sardinia, Italy has been analyzed by combining X-ray Fluorescence Spectroscopy (XRF) with Monte Carlo simulations using the XRMC code. The "Cesta" had been discovered probably in the XVIII century with the first graphic representation reported around 1761. In a later draft (dated 1764), the basket has been depicted as being carried upside-down on the shoulder of a large bronze warrior Barthélemy (1761), Pinza (1901), Winckelmann (1776) . The two pictorial representations differed only by the presence of handles in the most recent one. XRF measurements revealed that the handles of the object are composed by brass while the other parts are composed by bronze suggesting the handles as being a later addition to the original object. The artifact is covered at its surface by a fairly thick corrosion patina. In order to determine the bronze bulk composition without the need for removing the outer patina, the artifact has been modeled as a two layer object in Monte Carlo simulations.
Higham, Charles F. W.
2015-01-01
There are two models for the origins and timing of the Bronze Age in Southeast Asia. The first centres on the sites of Ban Chiang and Non Nok Tha in Northeast Thailand. It places the first evidence for bronze technology in about 2000 B.C., and identifies the origin by means of direct contact with specialists of the Seima Turbino metallurgical tradition of Central Eurasia. The second is based on the site of Ban Non Wat, 280 km southwest of Ban Chiang, where extensive radiocarbon dating places the transition into the Bronze Age in the 11th century B.C. with likely origins in a southward expansion of technological expertise rooted in the early states of the Yellow and Yangtze valleys, China. We have redated Ban Chiang and Non Nok Tha, as well as the sites of Ban Na Di and Ban Lum Khao, and here present 105 radiocarbon determinations that strongly support the latter model. The statistical analysis of the results using a Bayesian approach allows us to examine the data at a regional level, elucidate the timing of arrival of copper base technology in Southeast Asia and consider its social impact. PMID:26384011
Arnone, J A; Zaller, J G; Körner, Ch; Ziegler, C; Zandt, H
1995-09-01
Results from laboratory feeding experiments have shown that elevated atmospheric carbon dioxide can affect interactions between plants and insect herbivores, primarily through changes in leaf nutritional quality occurring at elevated CO 2 . Very few data are available on insect herbivory in plant communities where insects can choose among species and positions in the canopy in which to feed. Our objectives were to determine the extent to which CO 2 -induced changes in plant communities and leaf nutritional quality may affect herbivory at the level of the entire canopy. We introduced equivalent populations of fourth instar Spodoptera eridania, a lepidopteran generalist, to complex model ecosystems containing seven species of moist tropical plants maintained under low mineral nutrient supply. Larvae were allowed to feed freely for 14 days, by which time they had reached the seventh instar. Prior to larval introductions, plant communities had been continuously exposed to either 340 μl CO 2 l -1 or to 610 μl CO 2 l -1 for 1.5 years. No major shifts in leaf nutritional quality [concentrations of N, total non-structural carbohydrates (TNC), sugar, and starch; ratios of: C/N, TNC/N, sugar/N, starch/N; leaf toughness] were observed between CO 2 treatments for any of the species. Furthermore, no correlations were observed between these measures of leaf quality and leaf biomass consumption. Total leaf area and biomass of all plant communities were similar when caterpillars were introduced. However, leaf biomass of some species was slightly greater-and for other species slightly less (e.g. Cecropia peltata)-in communities exposed to elevated CO 2 . Larvae showed the strongest preference for C. peltata leaves, the plant species that was least abundant in all communites, and fed relatively little on plants species which were more abundant. Thus, our results indicate that leaf tissue quality, as described by these parameters, is not necessarily affected by elevated CO 2 under relatively low nutrient conditions. Hence, the potential importance of CO 2 -induced shifts in leaf nutritional quality, as determinants of herbivory, may be overestimated for many plant communities growing on nutrient-poor sites if estimates are based on traditional laboratory feeding studies. Finally, slight shifts in the abundance of leaf tissue of various species occurring under elevated CO 2 will probably not significantly affect herbivory by generalist insects. However, generalist insect herbivores appear to become more dependent on less-preferred plant species in cases where elevated CO 2 results in reduced availability of leaves of a favoured plant species, and this greater dependency may eventually affect insect populations adversely.
Leaf litter breakdown of native and exotic tree species in two Hawaiian streams that differ in flow
Megan Roberts; Ayron M. Strauch; Tracy Wiegner; Richard A. Mackenzie
2016-01-01
Riparian leaf litter is a major source of allochthonous organic material to temperate and tropical streams, promoting primary and secondary productivity in lotic and nearshore habitats. In tropical island streams, where native leaf-shredding macroinvertebrates are absent, physical fragmentation from stream flow is an important factor affecting leaf litter breakdown and...
NASA Technical Reports Server (NTRS)
Spalvins, T.
1973-01-01
Solid film lubricants of radio frequency sputtered molybdenum disulfide (MoS2) were applied to silver, gold, copper, and bronze surfaces that had various pretreatments (mechanical polishing, sputter etching, oxidation, and sulfurization). Optical and electron transmission micrographs and electron diffraction patterns were used to interpret the film formation characteristics and to evaluate the sputtering conditions in regard to the film and substrate compatibility. Sputtered MoS2 films flaked and peeled on silver, copper, and bronze surfaces except when the surfaces had been specially oxidized. The flaking and peeling was a result of sulfide compound formation and the corresponding grain growth of the sulfide film. Sputtered MoS2 films showed no peeling and flaking on gold surfaces regardless of surface pretreatment.
Schroeder, Oskar; Benecke, Norbert; Frölich, Kai; Peng, Zuogang; Kaniuth, Kai; Sverchkov, Leonid; Reinhold, Sabine; Belinskiy, Andrey; Ludwig, Arne
2017-01-01
Sheep were one of the first livestock species domesticated by humans. After initial domestication in the Middle East they were spread across Eurasia. The modern distribution of endogenous Jaagsiekte sheep retrovirus insertions in domestic sheep breeds suggests that over the course of millennia, successive introductions of improved lineages and selection for wool quality occurred in the Mediterranean region and most of Asia. Here we present a novel ancient DNA approach using data of endogenous retroviral insertions in Bronze and Iron Age domestic sheep from the Caucasus and Pamir mountain areas. Our findings support a secondary introduction of wool sheep from the Middle East between the Late Bronze Age and Iron Age into most areas of Eurasia. PMID:28632161
Schroeder, Oskar; Benecke, Norbert; Frölich, Kai; Peng, Zuogang; Kaniuth, Kai; Sverchkov, Leonid; Reinhold, Sabine; Belinskiy, Andrey; Ludwig, Arne
2017-06-20
Sheep were one of the first livestock species domesticated by humans. After initial domestication in the Middle East they were spread across Eurasia. The modern distribution of endogenous Jaagsiekte sheep retrovirus insertions in domestic sheep breeds suggests that over the course of millennia, successive introductions of improved lineages and selection for wool quality occurred in the Mediterranean region and most of Asia. Here we present a novel ancient DNA approach using data of endogenous retroviral insertions in Bronze and Iron Age domestic sheep from the Caucasus and Pamir mountain areas. Our findings support a secondary introduction of wool sheep from the Middle East between the Late Bronze Age and Iron Age into most areas of Eurasia.
The role of oxide structure on copper wire to the rubber adhesion
NASA Astrophysics Data System (ADS)
Su, Yea-Yang; Shemenski, Robert M.
2000-07-01
Most metals have an oxide layer on the surface. However, the structure of the oxide varies with the matrix composition, and depends upon the environmental conditions. A bronze coating, nominal composition of 98.5% Cu and balance of Sn, is applied to steel wire for reinforcing pneumatic tire beads and to provide adhesion to rubber. This work studied the influence of copper oxides on the bronze coating on adhesion during vulcanization. To emphasize the oxide structures, electrolytic tough pitch (ETP) copper wire was used instead of the traditional bronze-coated tire bead wire. Experimental results confirmed the hypothesis that cuprous oxide (Cu 2O) could significantly improve bonding between copper wire and rubber, and demonstrated that the interaction between rubber and oxide layer on wire is an electrochemical reaction.
Poiré, Richard; Wiese-Klinkenberg, Anika; Parent, Boris; Mielewczik, Michael; Schurr, Ulrich; Tardieu, François; Walter, Achim
2010-06-01
Diel (24 h) leaf growth patterns were differently affected by temperature variations and the circadian clock in several plant species. In the monocotyledon Zea mays, leaf elongation rate closely followed changes in temperature. In the dicotyledons Nicotiana tabacum, Ricinus communis, and Flaveria bidentis, the effect of temperature regimes was less obvious and leaf growth exhibited a clear circadian oscillation. These differences were related neither to primary metabolism nor to altered carbohydrate availability for growth. The effect of endogenous rhythms on leaf growth was analysed under continuous light in Arabidopsis thaliana, Ricinus communis, Zea mays, and Oryza sativa. No rhythmic growth was observed under continuous light in the two monocotyledons, while growth rhythmicity persisted in the two dicotyledons. Based on model simulations it is concluded that diel leaf growth patterns in mono- and dicotyledons result from the additive effects of both circadian-clock-controlled processes and responses to environmental changes such as temperature and evaporative demand. Apparently very distinct diel leaf growth behaviour of monocotyledons and dicotyledons can thus be explained by the different degrees to which diel temperature variations affect leaf growth in the two groups of species which, in turn, depends on the extent of the leaf growth control by internal clocks.
Microbial decomposition is highly sensitive to leaf litter emersion in a permanent temperate stream.
Mora-Gómez, Juanita; Duarte, Sofia; Cássio, Fernanda; Pascoal, Cláudia; Romaní, Anna M
2018-04-15
Drought frequency and intensity in some temperate regions are forecasted to increase under the ongoing global change, which might expose permanent streams to intermittence and have severe repercussions on stream communities and ecosystem processes. In this study, we investigated the effect of drought duration on microbial decomposition of Populus nigra leaf litter in a temperate permanent stream (Oliveira, NW Portugal). Specifically, we measured the response of the structural (assemblage composition, bacterial and fungal biomass) and functional (leaf litter decomposition, extracellular enzyme activities (EEA), and fungal sporulation) parameters of fungal and bacterial communities on leaf litter exposed to emersion during different time periods (7, 14 and 21d). Emersion time affected microbial assemblages and litter decomposition, but the response differed among variables. Leaf decomposition rates and the activity of β-glucosidase, cellobiohydrolase and phosphatase were gradually reduced with increasing emersion time, while β-xylosidase reduction was similar when emersion last for 7 or more days, and the phenol oxidase reduction was similar at 14 and 21days of leaf emersion. Microbial biomass and fungal sporulation were reduced after 21days of emersion. The structure of microbial assemblages was affected by the duration of the emersion period. The shifts in fungal assemblages were correlated with a decreased microbial capacity to degrade lignin and hemicellulose in leaf litter exposed to emersion. Additionally, some resilience was observed in leaf litter mass loss, bacterial biomass, some enzyme activities and structure of fungal assemblages. Our study shows that drought can strongly alter structural and functional aspects of microbial decomposers. Therefore, the exposure of leaf litter to increasing emersion periods in temperate streams is expected to affect decomposer communities and overall decomposition of plant material by decelerating carbon cycling in streams. Copyright © 2017 Elsevier B.V. All rights reserved.
Panchen, Zoe A.; Primack, Richard B.; Gallinat, Amanda S.; Nordt, Birgit; Stevens, Albert-Dieter; Du, Yanjun; Fahey, Robert
2015-01-01
Background and Aims Autumn leaf senescence marks the end of the growing season in temperate ecosystems. Its timing influences a number of ecosystem processes, including carbon, water and nutrient cycling. Climate change is altering leaf senescence phenology and, as those changes continue, it will affect individual woody plants, species and ecosystems. In contrast to spring leaf out times, however, leaf senescence times remain relatively understudied. Variation in the phenology of leaf senescence among species and locations is still poorly understood. Methods Leaf senescence phenology of 1360 deciduous plant species at six temperate botanical gardens in Asia, North America and Europe was recorded in 2012 and 2013. This large data set was used to explore ecological and phylogenetic factors associated with variation in leaf senescence. Key Results Leaf senescence dates among species varied by 3 months on average across the six locations. Plant species tended to undergo leaf senescence in the same order in the autumns of both years at each location, but the order of senescence was only weakly correlated across sites. Leaf senescence times were not related to spring leaf out times, were not evolutionarily conserved and were only minimally influenced by growth habit, wood anatomy and percentage colour change or leaf drop. These weak patterns of leaf senescence timing contrast with much stronger leaf out patterns from a previous study. Conclusions The results suggest that, in contrast to the broader temperature effects that determine leaf out times, leaf senescence times are probably determined by a larger or different suite of local environmental effects, including temperature, soil moisture, frost and wind. Determining the importance of these factors for a wide range of species represents the next challenge for understanding how climate change is affecting the end of the growing season and associated ecosystem processes. PMID:25808654
Panchen, Zoe A; Primack, Richard B; Gallinat, Amanda S; Nordt, Birgit; Stevens, Albert-Dieter; Du, Yanjun; Fahey, Robert
2015-11-01
Autumn leaf senescence marks the end of the growing season in temperate ecosystems. Its timing influences a number of ecosystem processes, including carbon, water and nutrient cycling. Climate change is altering leaf senescence phenology and, as those changes continue, it will affect individual woody plants, species and ecosystems. In contrast to spring leaf out times, however, leaf senescence times remain relatively understudied. Variation in the phenology of leaf senescence among species and locations is still poorly understood. Leaf senescence phenology of 1360 deciduous plant species at six temperate botanical gardens in Asia, North America and Europe was recorded in 2012 and 2013. This large data set was used to explore ecological and phylogenetic factors associated with variation in leaf senescence. Leaf senescence dates among species varied by 3 months on average across the six locations. Plant species tended to undergo leaf senescence in the same order in the autumns of both years at each location, but the order of senescence was only weakly correlated across sites. Leaf senescence times were not related to spring leaf out times, were not evolutionarily conserved and were only minimally influenced by growth habit, wood anatomy and percentage colour change or leaf drop. These weak patterns of leaf senescence timing contrast with much stronger leaf out patterns from a previous study. The results suggest that, in contrast to the broader temperature effects that determine leaf out times, leaf senescence times are probably determined by a larger or different suite of local environmental effects, including temperature, soil moisture, frost and wind. Determining the importance of these factors for a wide range of species represents the next challenge for understanding how climate change is affecting the end of the growing season and associated ecosystem processes. © The Author 2015. Published by Oxford University Press on behalf of the Annals of Botany Company. All rights reserved. For Permissions, please email: journals.permissions@oup.com.
Jones, Ian M; Koptur, Suzanne
2015-01-01
• Extrafloral nectar (EFN) mediates food for protection mutualisms between plants and defensive insects. Understanding sources of variation in EFN production is important because such variations may affect the number and identity of visitors and the effectiveness of plant defense. We investigated the influence of plant developmental stage, time of day, leaf age, and leaf damage on EFN production in Senna mexicana var. chapmanii. The observed patterns of variation in EFN production were compared with those predicted by optimal defense theory.• Greenhouse experiments with potted plants were conducted to determine how plant age, time of day, and leaf damage affected EFN production. A subsequent field study was conducted to determine how leaf damage, and the resulting increase in EFN production, affected ant visitation in S. chapmanii.• More nectar was produced at night and by older plants. Leaf damage resulted in increased EFN production, and the magnitude of the response was greater in plants damaged in the morning than those damaged at night. Damage to young leaves elicited a stronger defensive response than damage to older leaves, in line with optimal defense theory. Damage to the leaves of S. chapmanii also resulted in significantly higher ant visitation in the field.• Extrafloral nectar is an inducible defense in S. chapmanii. Developmental variations in its production support the growth differentiation balance hypothesis, while within-plant variations and damage responses support optimal defense theory. © 2015 Botanical Society of America, Inc.
Leaf gas exchange of mature bottomland oak trees
Rico M. Gazal; Mark E. Kubiske; Kristina F. Connor
2009-01-01
We determined how changes in environmental moisture affected leaf gas exchange in Nuttall (Quercus texana Buckley), overcup (Q. lyrata Walt.), and dominant and codominant swamp chestnut (Q. michauxii Nutt.) oak trees in Mississippi and Louisiana. We used canopy access towers to measure leaf level gas...
Husbands, Aman Y.; Benkovics, Anna H.; Nogueira, Fabio T.S.; Lodha, Mukesh; Timmermans, Marja C.P.
2015-01-01
Flattened leaf architecture is not a default state but depends on positional information to precisely coordinate patterns of cell division in the growing primordium. This information is provided, in part, by the boundary between the adaxial (top) and abaxial (bottom) domains of the leaf, which are specified via an intricate gene regulatory network whose precise circuitry remains poorly defined. Here, we examined the contribution of the ASYMMETRIC LEAVES (AS) pathway to adaxial-abaxial patterning in Arabidopsis thaliana and demonstrate that AS1-AS2 affects this process via multiple, distinct regulatory mechanisms. AS1-AS2 uses Polycomb-dependent and -independent mechanisms to directly repress the abaxial determinants MIR166A, YABBY5, and AUXIN RESPONSE FACTOR3 (ARF3), as well as a nonrepressive mechanism in the regulation of the adaxial determinant TAS3A. These regulatory interactions, together with data from prior studies, lead to a model in which the sequential polarization of determinants, including AS1-AS2, explains the establishment and maintenance of adaxial-abaxial leaf polarity. Moreover, our analyses show that the shared repression of ARF3 by the AS and trans-acting small interfering RNA (ta-siRNA) pathways intersects with additional AS1-AS2 targets to affect multiple nodes in leaf development, impacting polarity as well as leaf complexity. These data illustrate the surprisingly multifaceted contribution of AS1-AS2 to leaf development showing that, in conjunction with the ta-siRNA pathway, AS1-AS2 keeps the Arabidopsis leaf both flat and simple. PMID:26589551
7 CFR 906.365 - Texas Orange and Grapefruit Regulation 34.
Code of Federal Regulations, 2010 CFR
2010-01-01
... Bright, U.S. No. 1 Bronze, U.S. Combination (with not less than 60 percent, by count, of the oranges in....S. No. 1 Bright, or U.S. No. 1 Bronze, or meet the quality requirements of “Texas Fancy” or “Texas... within 48 hours prior to the time of shipment. No handler may transport by motor vehicle or cause the...
Shanks, H.R.; Bevolo, A.J.; Danielson, G.C.; Weber, M.F.
An oxygen electrode for a fuel cell utilizing an acid electrolyte has a substrate of an alkali metal tungsten bronze of the formula: A/sub x/WO/sub 3/ where A is an alkali metal and x is at least 0.2, which is covered with a thin layer of platinum tungsten bronze of the formula: Pt/sub y/WO/sub 3/ where y is at least 0.8.
Shanks, Howard R.; Bevolo, Albert J.; Danielson, Gordon C.; Weber, Michael F.
1980-11-04
An oxygen electrode for a fuel cell utilizing an acid electrolyte has a substrate of an alkali metal tungsten bronze of the formula: A.sub.x WO.sub.3 where A is an alkali metal and x is at least 0.2, which is covered with a thin layer of platinum tungsten bronze of the formula: Pt.sub.y WO.sub.3 where y is at least 0.8.
Shifts in leaf litter breakdown along a forest-pasture-urban gradient in Andean streams.
Iñiguez-Armijos, Carlos; Rausche, Sirkka; Cueva, Augusta; Sánchez-Rodríguez, Aminael; Espinosa, Carlos; Breuer, Lutz
2016-07-01
Tropical montane ecosystems of the Andes are critically threatened by a rapid land-use change which can potentially affect stream variables, aquatic communities, and ecosystem processes such as leaf litter breakdown. However, these effects have not been sufficiently investigated in the Andean region and at high altitude locations in general. Here, we studied the influence of land use (forest-pasture-urban) on stream physico-chemical variables (e.g., water temperature, nutrient concentration, and pH), aquatic communities (macroinvertebrates and aquatic fungi) and leaf litter breakdown rates in Andean streams (southern Ecuador), and how variation in those stream physico-chemical variables affect macroinvertebrates and fungi related to leaf litter breakdown. We found that pH, water temperature, and nutrient concentration increased along the land-use gradient. Macroinvertebrate communities were significantly different between land uses. Shredder richness and abundance were lower in pasture than forest sites and totally absent in urban sites, and fungal richness and biomass were higher in forest sites than in pasture and urban sites. Leaf litter breakdown rates became slower as riparian land use changed from natural to anthropogenically disturbed conditions and were largely determined by pH, water temperature, phosphate concentration, fungal activity, and single species of leaf-shredding invertebrates. Our findings provide evidence that leaf litter breakdown in Andean streams is sensitive to riparian land-use change, with urban streams being the most affected. In addition, this study highlights the role of fungal biomass and shredder species (Phylloicus; Trichoptera and Anchytarsus; Coleoptera) on leaf litter breakdown in Andean streams and the contribution of aquatic fungi in supporting this ecosystem process when shredders are absent or present low abundance in streams affected by urbanization. Finally, we summarize important implications in terms of managing of native vegetation and riparian buffers to promote ecological integrity and functioning of tropical Andean stream ecosystems.
Determining past leaf-out times of New England's deciduous forests from herbarium specimens.
Everill, Peter H; Primack, Richard B; Ellwood, Elizabeth R; Melaas, Eli K
2014-08-01
• There is great interest in studying leaf-out times of temperate forests because of the importance of leaf-out in controlling ecosystem processes, especially in the face of a changing climate. Remote sensing and modeling, combined with weather records and field observations, are increasing our knowledge of factors affecting variation in leaf-out times. Herbarium specimens represent a potential new source of information to determine whether the variation in leaf-out times observed in recent decades is comparable to longer time frames over past centuries.• Here we introduce the use of herbarium specimens as a method for studying long-term changes in leaf-out times of deciduous trees. We collected historical leaf-out data for the years 1834-2008 from common deciduous trees in New England using 1599 dated herbarium specimens with young leaves.• We found that leaf-out dates are strongly affected by spring temperature, with trees leafing out 2.70 d earlier for each degree C increase in mean April temperature. For each degree C increase in local temperature, trees leafed out 2.06 d earlier. Additionally, the mean response of leaf-out dates across all species and sites over time was 0.4 d earlier per decade. Our results are of comparable magnitude to results from studies using remote sensing and direct field observations.• Across New England, mean leaf-out dates varied geographically in close correspondence with those observed in studies using satellite data. This study demonstrates that herbarium specimens can be a valuable source of data on past leaf-out times of deciduous trees. © 2014 Botanical Society of America, Inc.
Freitas, A F; Pereira, F F; Formagio, A S N; Lucchetta, J T; Vieira, M C; Mussury, R M
2014-10-01
Spodoptera frugiperda (J.E. Smith) (Lepidoptera: Noctuidae) causes significant losses in corn crops and necessitates the use of alternative control strategies, such as the application of bioinsecticides. We report the effect of methanolic leaf extracts of Annona dioica, Annona cacans, and Annona coriacea on the development and reproduction of S. frugiperda. A quantitative analysis was carried out to determine the total concentration of phenolics, flavonoids, and condensed tannin (CT) in leaf extracts. Corn leaves were immersed in a 1% methanolic leaf extract solution and fed to second instars of S. frugiperda. Leaf disks dipped in the synthetic insecticide Connect® (Bayer CropScience Ltda) composed of a neonicotinoid (imidacloprid) and a pyrethroid (β-cyfluthrin), which are harmful to S. frugiperda, was used as positive control. Distilled water was used as a negative control treatment. The leaf extract of A. coriacea decreased larval survivorship, arrested pupal development, and affected the weight gain of S. frugiperda. A. dioica also affected larval survivorship, but its effects were more pronounced for the adult stage, as fecundity, fertility, egg hatchability, and embryonic development were severely affected. Leaf extracts from A. cacans had no effect on S. frugiperda. The leaf extracts of A. dioica and A. coriacea showed a higher content of flavonoids and phenols, respectively. Our results indicated that both A. dioica and A. coriacea have the potential for development as botanical insecticides.
Antibacterial drugs as corrosion inhibitors for bronze surfaces in acidic solutions
NASA Astrophysics Data System (ADS)
Rotaru, Ileana; Varvara, Simona; Gaina, Luiza; Muresan, Liana Maria
2014-12-01
The present study is aiming to investigate the effect of four antibiotics (amoxicillin, ciprofloxacin, doxycycline and streptomycin,) belonging to different classes of antibacterial drugs on bronze corrosion in a solution simulating an acid rain (pH 4). Due to their ability to form protective films on the metal surface, the tested antibiotics act as corrosion inhibitors for bronze. The antibiotics were tested at various concentrations in order to determine the optimal concentration range for the best corrosion inhibiting effect. In evaluating the inhibition efficiency, polarization curves, electrochemical impedance spectroscopy, SEM and XPS measurements were used. Moreover, a correlation between the inhibition efficiency of some antibacterial drugs and certain molecular parameters was determined by quantum chemical computations. Parameters like energies EHOMO and ELUMO and HOMO-LUMO energy gap were used for correlation with the corrosion data.
A-15 Superconducting composite wires and a method for making
Suenaga, Masaki; Klamut, Carl J.; Luhman, Thomas S.
1984-01-01
A method for fabricating superconducting wires wherein a billet of copper containing filaments of niobium or vanadium is rolled to form a strip which is wrapped about a tin-alloy core to form a composite. The alloy is a tin-copper alloy for niobium filaments and a gallium-copper alloy for vanadium filaments. The composite is then drawn down to a desired wire size and heat treated. During the heat treatment process, the tin in the bronze reacts with the niobium to form the superconductor niobium tin. In the case where vanadium is used, the gallium in the gallium bronze reacts with the vanadium to form the superconductor vanadium gallium. This new process eliminates the costly annealing steps, external tin plating and drilling of bronze ingots required in a number of prior art processes.
Wrapping process for fabrication of A-15 superconducting composite wires
Suenaga, M.; Klamut, C.J.; Luhman, T.S.
1980-08-15
A method for fabricating superconducting wires wherein a billet of copper containing filaments of niobium or vanadium is rolled to form a strip which is wrapped about a tin-alloy core to form a composite. The alloy is a tin-copper alloy for niobium filaments and a gallium-copper alloy for vanadium filaments. The composite is then drawn down to a desired wire size and heat treated. During the heat treatment process, the tin in the bronze reacts with the niobium to form the superconductor niobium tin. In the case where vanadium is used, the gallium in the gallium bronze reacts with the vanadium to form the superconductor vanadium gallium. This new process eliminates the costly annealing steps, external tin plating and drilling of bronze ingots required in a number of prior art processes.
NASA Astrophysics Data System (ADS)
Martyushev, Nikita V.; Risto, Nikolay A.
2014-10-01
This paper investigates the use of nanopowders in the composition of foundry coatings when casting leaded tin bronzes. Influence of the composition of the applied protective coating on surface finish is studied. The effects of the coatings of the following compositions are compared: non-stick coating (a mixture of low-dispersed chromium oxide powder and heat-treated vegetable oil); non-stick lubricant ASPF-2/RgU on the basis of low- dispersed graphite powder and heat-treated vegetable oil; patent #2297300 (a mixture of superdispersed zirconium dioxide powder with industrial oil). It is demonstrated that application of foundry coatings containing superdispersed metal oxide powders with low thermal conductivity makes it possible to significantly reduce irregularities and eliminate gas porosity on the surface of tin-leaded bronze castings.
NASA Technical Reports Server (NTRS)
Brewe, D. E.; Wisander, D. W.; Scribbe, H. W.
1972-01-01
Forty-millimeter-bore ball bearings with lead- and lead-alloy-coated retainers were operated in liquid hydrogen at 30,000 rpm under a thrust load of 1780 N (400 lb.) Four different substrate materials were used for the retainer. Longer bearing run times were achieved with a lead-tin-copper alloy coating plated onto a leaded-bronze material (22.5 hr) and an aluminum-bronze alloy (19.3 hr). One bearing with a pure lead coating achieved the desired objective of 10 hr. This bearing had an aluminum - bronze substrate retainer and ran successfully for 12.4 hr. Additions of antimony to the lead provided an alloy coating with better wear resistance than pure lead; however, this coating was abrasive to the outer-race lands.
NASA Technical Reports Server (NTRS)
Gausman, H. W.; Cardenas, R.; Berumen, A.
1974-01-01
Pepper and sorghum plants (characterized by porous and compact leaf mesophylls, respectively) were used to study the influence of leaf age on light reflectance. Measurements were limited to the upper five nodal positions within each growth stage, since upper leaves make up most of the reflectance surfaces remotely sensed. The increase in leaf thickness and water content with increasing leaf age was taken into consideration, since each of these factors affects the reflectance as well as the selection of spectral wavelength intervals for optimum discrimination of vegetation.
Jianwei Zhang; Marcus Schaub; Jonathan A. Ferdinand; John M. Skelly; Kim C. Steiner; James E. Savage
2010-01-01
We investigated the effect of leaf age on the response of net photosynthesis (A), stomatal conductance (gwv), foliar injury, and leaf nitrogen concentration (NL) to tropospheric ozone (O3) on Prunus serotina seedlings grown in open-plots (AA) and open-top...
Association of microRNAs with Types of Leaf Curvature in Brassica rapa.
Ren, Wenqing; Wang, Han; Bai, Jinjuan; Wu, Feijie; He, Yuke
2018-01-01
Many vegetable crops of Brassica rapa are characterized by their typical types of leaf curvature. Leaf curvature in the right direction and to the proper degree is important for the yield and quality of green vegetable products, when cultivated under stress conditions. Recent research has unveiled some of the roles of miRNAs in Brassica crops such as how they regulate the timing of leafy head initiation and shape of the leafy head. However, the molecular mechanism underlying the variability in leaf curvature in B. rapa remains unclear. We tested the hypothesis that the leaf curvature of B. rapa is affected by miRNA levels. On the basis of leaf phenotyping, 56 B. rapa accessions were classified into five leaf curvature types, some of which were comparable to miRNA mutants of Arabidopsis thaliana in phenotype. Higher levels of miR166 and miR319a expression were associated with downward curvature and wavy margins, respectively. Overexpression of the Brp - MIR166g-1 gene caused rosette leaves to change from flat to downward curving and folding leaves to change from upward curving to flat, leading to the decrease in the number of incurved leaves and size of the leafy head. Our results reveal that miRNAs affect the types of leaf curvature in B. rapa . These findings provide insight into the relationship between miRNAs and variation in leaf curvature.
Association of microRNAs with Types of Leaf Curvature in Brassica rapa
Ren, Wenqing; Wang, Han; Bai, Jinjuan; Wu, Feijie; He, Yuke
2018-01-01
Many vegetable crops of Brassica rapa are characterized by their typical types of leaf curvature. Leaf curvature in the right direction and to the proper degree is important for the yield and quality of green vegetable products, when cultivated under stress conditions. Recent research has unveiled some of the roles of miRNAs in Brassica crops such as how they regulate the timing of leafy head initiation and shape of the leafy head. However, the molecular mechanism underlying the variability in leaf curvature in B. rapa remains unclear. We tested the hypothesis that the leaf curvature of B. rapa is affected by miRNA levels. On the basis of leaf phenotyping, 56 B. rapa accessions were classified into five leaf curvature types, some of which were comparable to miRNA mutants of Arabidopsis thaliana in phenotype. Higher levels of miR166 and miR319a expression were associated with downward curvature and wavy margins, respectively. Overexpression of the Brp-MIR166g-1 gene caused rosette leaves to change from flat to downward curving and folding leaves to change from upward curving to flat, leading to the decrease in the number of incurved leaves and size of the leafy head. Our results reveal that miRNAs affect the types of leaf curvature in B. rapa. These findings provide insight into the relationship between miRNAs and variation in leaf curvature. PMID:29467771
Wuyts, Nathalie; Massonnet, Catherine; Dauzat, Myriam; Granier, Christine
2012-09-01
Light and soil water content affect leaf surface area expansion through modifications in epidermal cell numbers and area, while effects on leaf thickness and mesophyll cell volumes are far less documented. Here, three-dimensional imaging was applied in a study of Arabidopsis thaliana leaf growth to determine leaf thickness and the cellular organization of mesophyll tissues under moderate soil water deficit and two cumulative light conditions. In contrast to surface area, thickness was highly conserved in response to water deficit under both low and high cumulative light regimes. Unlike epidermal and palisade mesophyll tissues, no reductions in cell number were observed in the spongy mesophyll; cells had rather changed in volume and shape. Furthermore, leaf features of a selection of genotypes affected in leaf functioning were analysed. The low-starch mutant pgm had very thick leaves because of unusually large palisade mesophyll cells, together with high levels of photosynthesis and stomatal conductance. By means of an open stomata mutant and a 9-cis-epoxycarotenoid dioxygenase overexpressor, it was shown that stomatal conductance does not necessarily have a major impact on leaf dimensions and cellular organization, pointing to additional mechanisms for the control of CO(2) diffusion under high and low stomatal conductance, respectively. © 2012 Blackwell Publishing Ltd.
Medeiros, Juliana S.; Ward, Joy K.
2013-01-01
Summary Changes in atmospheric carbon dioxide concentration ([CO2]) affect plant carbon/water trade-offs, with implications for drought tolerance. Leaf-level studies often indicate that drought tolerance may increase with rising [CO2], but integrated leaf and xylem responses are not well understood in this respect. In addition, the influence of low [CO2] of the last glacial period on drought tolerance and xylem properties is not well understood.We investigated the interactive effects of a broad range of [CO2] and plant water potentials on leaf function, xylem structure and function and the integration of leaf and xylem function in Phaseolus vulgaris.Elevated [CO2] decreased vessel implosion strength, reduced conduit specific hydraulic conductance, and compromised leaf specific xylem hydraulic conductance under moderate drought. By contrast, at glacial [CO2], transpiration was maintained under moderate drought via greater conduit specific and leaf specific hydraulic conductance in association with increased vessel implosion strength.Our study involving the integration of leaf and xylem responses suggests that increasing [CO2] does not improve drought tolerance. We show that under glacial conditions changes in leaf and xylem properties could increase drought tolerance, while under future conditions greater productivity may only occur when higher water use can be accommodated. PMID:23668237
Survey of Nickel-Aluminium-Bronze Casting Alloys on Marine Applications,
1981-04-01
and corrosion performance of nickel-aluminium bronze (NAB)/covered by naval specification DGS-8520 and DGS-348 have been investigated. No evidence was...found to suggest that there would be any significant difference in corrosion performance between alloys meeting the two specifications. Early... corrosion problems associated with the weld repair areas of castings have been overcome largely by using improved foundry and welding techniques followed by a
Towards reconstruction of the lost Late Bronze Age intra-caldera island of Santorini, Greece.
Karátson, Dávid; Gertisser, Ralf; Telbisz, Tamás; Vereb, Viktor; Quidelleur, Xavier; Druitt, Timothy; Nomikou, Paraskevi; Kósik, Szabolcs
2018-05-04
During the Late Bronze Age, the island of Santorini had a semi-closed caldera harbour inherited from the 22 ka Cape Riva Plinian eruption, and a central island referred to as 'Pre-Kameni' after the present-day Kameni Islands. Here, the size and age of the intracaldera island prior to the Late Bronze Age (Minoan) eruption are constrained using a photo-statistical method, complemented by granulometry and high-precision K-Ar dating. Furthermore, the topography of Late Bronze Age Santorini is reconstructed by creating a new digital elevation model (DEM). Pre-Kameni and other parts of Santorini were destroyed during the 3.6 ka Minoan eruption, and their fragments were incorporated as lithic clasts in the Minoan pyroclastic deposits. Photo-statistical analysis and granulometry of these lithics, differentiated by lithology, constrain the volume of Pre-Kameni to 2.2-2.5 km 3 . Applying the Cassignol-Gillot K-Ar dating technique to the most characteristic black glassy andesite lithics, we propose that the island started to grow at 20.2 ± 1.0 ka soon after the Cape Riva eruption. This implies a minimum long-term lava extrusion rate of ~0.13-0.14 km 3 /ky during the growth of Pre-Kameni.
Corrosion of dental aluminium bronze in neutral saline and saline lactic acid.
Tibballs, J E; Erimescu, Raluca
2006-09-01
To compare the corrosion behaviours of two aluminium bronze, dental casting alloys during a standard immersion test and for immersion in neutral saline. Cast specimens of aluminium bronzes with 1.4 wt% Fe (G) and 4 wt% Fe (N) were subject to progressively longer periods (up to in total 7 days) immersed in 0.1 M saline, 0.1 M lactic acid solutions and examined by scanning electron microscopy with EDX analysis. Immersion in 0.1M neutral saline was for 7 days. In the acidic solution, exposed interdendritic volumes in alloy N corroded completely away in 7 days with dissolution of Ni-enriched precipitate species as well as the copper-rich matrix. Alloy G begins to corrode more slowly but by a similar mechanism. The number density of an Fe-enriched species is insufficient to maintain a continuous galvanic potential to the copper matrix, and dissolution becomes imperceptible. In neutral saline solution, galvanic action alone caused pit-etching, without the dissolution of either precipitate species. The upper limit for the total dissolution of metallic ions in the standard immersion test can be set at 200 microg cm(-2). Aluminium bronze dental alloys can be expected to release both copper and nickel ions into an acidic oral environment.
Satora, Leszek; Mytych, Jennifer; Bilska-Kos, Anna
2018-05-23
Bronze corydoras (Corydoras aeneus) is a small diurnal activity fish from South America. Under hypoxia conditions, it uses the posterior part of the intestine as an accessory respiratory organ. The present PCR studies demonstrated higher expression of HIF-1α (hypoxia-inducible factor) gene in the respiratory than that in digestive part of bronze corydoras intestine. Further, immunolocalization studies using antibodies specific to HIF-1α and transmission electron microscopy (TEM) revealed the presence of HIF-1α epitopes in the intestine of Corydoras aeneus. In the respiratory intestine, the numerous clusters of gold particles visualizing HIF-1α antibody were observed within fibroblasts, whereas in the digestive tract of this species, single gold grains in the epithelial cells were noted. On the other hand, the presence of HIF-1α and the cytoplasmic domain of the epidermal growth factor receptor (EGFR) in the respiratory intestine of bronze corydoras assumes their interactions in the system where these factors appeared for the first time. The non-obligatory air-breathing fishes using their digestive tract as an accessory respiratory organ during hypoxia conditions are very interesting for the studies of the processes that control HIF-1α expression and squamous cell proliferation.
NASA Astrophysics Data System (ADS)
Gulyaev, P.; Kotvanova, M.; Omelchenko, A.
2017-05-01
The mechanism of abnormal photo-thermal effect of laser radiation on nanoparticles of oxide bronzes has been proposed in this paper. The basic features of the observed effect are: a) sub-threshold absorption of laser radiation by the excitation of donor-like levels formed in the energy gap due to superficial defects of the oxide bronze nano-crystals; b) an interband radiationless transition of energy of excitation on deep triplet levels and c) consequent recombination occurring at the plasmon absorption. K or Na atoms thermally intercalated to the octahedral crystal structure of TiO2 in the wave SHS combustion generate acceptor levels in the gap. The prepared oxide bronzes of the non-stoichiometric composition NaxTiO2 and KxTiO2 were examined by high resolution TEM, and then grinded in a planetary mill with powerful dispersion energy density up to 4000 J/g. This made it possible to obtain nanoparticles about 50 nm with high surface defect density (1017-1019 cm-2 at a depth of 10 nm). High photo-thermal effect of laser radiation on the defect nanocrystals observed after its impregnation into cartilaginous tissue exceeds 7 times in comparison with the intact ones.
NASA Astrophysics Data System (ADS)
Kaniewski, D.; Paulissen, E.; Van Campo, E.; Weiss, H.; Otto, T.; Bretschneider, J.; Van Lerberghe, K.
2010-09-01
The alluvial deposits near Gibala-Tell Tweini provide a unique record of environmental history and food availability estimates covering the Late Bronze Age and the Early Iron Age. The refined pollen-derived climatic proxy suggests that drier climatic conditions occurred in the Mediterranean belt of Syria from the late 13th/early 12th centuries BC to the 9th century BC. This period corresponds with the time frame of the Late Bronze Age collapse and the subsequent Dark Age. The abrupt climate change at the end of the Late Bronze Age caused region-wide crop failures, leading towards socio-economic crises and unsustainability, forcing regional habitat-tracking. Archaeological data show that the first conflagration of Gibala occurred simultaneously with the destruction of the capital city Ugarit currently dated between 1194 and 1175 BC. Gibala redeveloped shortly after this destruction, with large-scale urbanization visible in two main architectural phases during the Early Iron Age I. The later Iron Age I city was destroyed during a second conflagration, which is radiocarbon-dated at circa 2950 cal yr BP. The data from Gibala-Tell Tweini provide evidence in support of the drought hypothesis as a triggering factor behind the Late Bronze Age collapse in the Eastern Mediterranean.
NASA Astrophysics Data System (ADS)
Afshari, Elham; Ghambari, Mohammad; Farhangi, Hasan
2016-11-01
In this study, jet milling was used to recycle tin bronze machining chips into powder. The main purpose of this study was to assess the effect of the microstructure of tin bronze machining chips on their breakage behavior. An experimental target jet mill was used to pulverize machining chips of three different tin bronze alloys containing 7wt%, 10wt%, and 12wt% of tin. Optical and electron microscopy, as well as sieve analysis, were used to follow the trend of pulverization. Each alloy exhibited a distinct rate of size reduction, particle size distribution, and fracture surface appearance. The results showed that the degree of pulverization substantially increased with increasing tin content. This behavior was attributed to the higher number of machining cracks as well as the increased volume fraction of brittle δ phase in the alloys with higher tin contents. The δ phase was observed to strongly influence the creation of machining cracks as well as the nucleation and propagation of cracks during jet milling. In addition, a direct relationship was observed between the mean δ-phase spacing and the mean size of the jet-milled product; i.e., a decrease in the δ-phase spacing resulted in smaller particles.
Long term leaf phenology and leaf exchange strategies of a cerrado savanna community
NASA Astrophysics Data System (ADS)
de Camargo, Maria Gabriela G.; Costa Alberton, Bruna; de Carvalho, Gustavo H.; Magalhães, Paula A. N. R.; Morellato, Leonor Patrícia C.
2017-04-01
Leaf development and senescence cycles are linked to a range of ecosystem processes, affecting seasonal patterns of atmosphere-ecosystem carbon and energy exchanges, resource availability and nutrient cycling. The degree of deciduousness of tropical trees and communities depend on ecosystems characteristics such as amount of biomass, species diversity and the strength and length of the dry season. Besides defining the growing season, deciduousness can also be an indicator of species response to climate changes in the tropics, mainly because severity of dry season can intensify leaf loss. Based on seven-years of phenological observations (2005 to 2011) we describe the long-term patterns of leafing phenology of a Brazilian cerrado savanna, aiming to (i) identify leaf exchange strategies of species, quantifying the degree of deciduousness, and verify whether these strategies vary among years depending on the length and strength of the dry seasons; (ii) define the growing seasons along the years and the main drivers of leaf flushing in the cerrado. We analyzed leafing patterns of 107 species and classified 69 species as deciduous (11 species), semi-deciduous (29) and evergreen (29). Leaf exchange was markedly seasonal, as expected for seasonal tropical savannas. Leaf fall predominated in the dry season, peaking in July, and leaf flushing in the transition between dry to wet seasons, peaking in September. Leafing patterns were similar among years with the growing season starting at the end of dry season, in September, for most species. However, leaf exchange strategies varied among years for most species (65%), except for evergreen strategy, mainly constant over years. Leafing patterns of cerrado species were strongly constrained by rainfall. The length of the dry season and rainfall intensity were likely affecting the individuals' leaf exchange strategies and suggesting a differential resilience of species to changes of rainfall regime, predicted on future global change scenarios.
Leaf primordium size specifies leaf width and vein number among row-type classes in barley.
Thirulogachandar, Venkatasubbu; Alqudah, Ahmad M; Koppolu, Ravi; Rutten, Twan; Graner, Andreas; Hensel, Goetz; Kumlehn, Jochen; Bräutigam, Andrea; Sreenivasulu, Nese; Schnurbusch, Thorsten; Kuhlmann, Markus
2017-08-01
Exploring genes with impact on yield-related phenotypes is the preceding step to accomplishing crop improvements while facing a growing world population. A genome-wide association scan on leaf blade area (LA) in a worldwide spring barley collection (Hordeum vulgare L.), including 125 two- and 93 six-rowed accessions, identified a gene encoding the homeobox transcription factor, Six-rowed spike 1 (VRS1). VRS1 was previously described as a key domestication gene affecting spike development. Its mutation converts two-rowed (wild-type VRS1, only central fertile spikelets) into six-rowed spikes (mutant vrs1, fully developed fertile central and lateral spikelets). Phenotypic analyses of mutant and wild-type leaves revealed that mutants had an increased leaf width with more longitudinal veins. The observed significant increase of LA and leaf nitrogen (%) during pre-anthesis development in vrs1 mutants also implies a link between wider leaf and grain number, which was validated from the association of vrs1 locus with wider leaf and grain number. Histological and gene expression analyses indicated that VRS1 might influence the size of leaf primordia by affecting cell proliferation of leaf primordial cells. This finding was supported by the transcriptome analysis of mutant and wild-type leaf primordia where in the mutant transcriptional activation of genes related to cell proliferation was detectable. Here we show that VRS1 has an independent role on barley leaf development which might influence the grain number. © 2017 The Authors. The Plant Journal published by John Wiley & Sons Ltd and Society for Experimental Biology.
Husbands, Aman Y; Benkovics, Anna H; Nogueira, Fabio T S; Lodha, Mukesh; Timmermans, Marja C P
2015-12-01
Flattened leaf architecture is not a default state but depends on positional information to precisely coordinate patterns of cell division in the growing primordium. This information is provided, in part, by the boundary between the adaxial (top) and abaxial (bottom) domains of the leaf, which are specified via an intricate gene regulatory network whose precise circuitry remains poorly defined. Here, we examined the contribution of the ASYMMETRIC LEAVES (AS) pathway to adaxial-abaxial patterning in Arabidopsis thaliana and demonstrate that AS1-AS2 affects this process via multiple, distinct regulatory mechanisms. AS1-AS2 uses Polycomb-dependent and -independent mechanisms to directly repress the abaxial determinants MIR166A, YABBY5, and AUXIN RESPONSE FACTOR3 (ARF3), as well as a nonrepressive mechanism in the regulation of the adaxial determinant TAS3A. These regulatory interactions, together with data from prior studies, lead to a model in which the sequential polarization of determinants, including AS1-AS2, explains the establishment and maintenance of adaxial-abaxial leaf polarity. Moreover, our analyses show that the shared repression of ARF3 by the AS and trans-acting small interfering RNA (ta-siRNA) pathways intersects with additional AS1-AS2 targets to affect multiple nodes in leaf development, impacting polarity as well as leaf complexity. These data illustrate the surprisingly multifaceted contribution of AS1-AS2 to leaf development showing that, in conjunction with the ta-siRNA pathway, AS1-AS2 keeps the Arabidopsis leaf both flat and simple. © 2015 American Society of Plant Biologists. All rights reserved.
Gautier, Hélène; Massot, Capucine; Stevens, Rebecca; Sérino, Sylvie; Génard, Michel
2009-02-01
The mechanisms involving light control of vitamin C content in fruits are not yet fully understood. The present study aimed to evaluate the impact of fruit and leaf shading on ascorbate (AsA) accumulation in tomato fruit and to determine how fruit sugar content (as an AsA precursor) affected AsA content. Cherry tomato plants were grown in a glasshouse. The control treatment (normally irradiated fruits and irradiated leaves) was compared with the whole-plant shading treatment and with leaf or fruit shading treatments in fruits harvested at breaker stage. In a second experiment, the correlation between sugars and AsA was studied during ripening. Fruit shading was the most effective treatment in reducing fruit AsA content. Under normal conditions, AsA and sugar content were correlated and increased with the ripening stage. Reducing fruit irradiance strongly decreased the reduced AsA content (-74 %), without affecting sugars, so that sugar and reduced AsA were no longer correlated. Leaf shading delayed fruit ripening: it increased the accumulation of oxidized AsA in green fruits (+98 %), whereas it decreased the reduced AsA content in orange fruits (-19 %), suggesting that fruit AsA metabolism also depends on leaf irradiance. Under fruit shading only, the absence of a correlation between sugars and reduced AsA content indicated that fruit AsA content was not limited by leaf photosynthesis or sugar substrate, but strongly depended on fruit irradiance. Leaf shading most probably affected fruit AsA content by delaying fruit ripening, and suggested a complex regulation of AsA metabolism which depends on both fruit and leaf irradiance and fruit ripening stage.
Gautier, Hélène; Massot, Capucine; Stevens, Rebecca; Sérino, Sylvie; Génard, Michel
2009-01-01
Background and Aims The mechanisms involving light control of vitamin C content in fruits are not yet fully understood. The present study aimed to evaluate the impact of fruit and leaf shading on ascorbate (AsA) accumulation in tomato fruit and to determine how fruit sugar content (as an AsA precursor) affected AsA content. Methods Cherry tomato plants were grown in a glasshouse. The control treatment (normally irradiated fruits and irradiated leaves) was compared with the whole-plant shading treatment and with leaf or fruit shading treatments in fruits harvested at breaker stage. In a second experiment, the correlation between sugars and AsA was studied during ripening. Key Results Fruit shading was the most effective treatment in reducing fruit AsA content. Under normal conditions, AsA and sugar content were correlated and increased with the ripening stage. Reducing fruit irradiance strongly decreased the reduced AsA content (−74 %), without affecting sugars, so that sugar and reduced AsA were no longer correlated. Leaf shading delayed fruit ripening: it increased the accumulation of oxidized AsA in green fruits (+98 %), whereas it decreased the reduced AsA content in orange fruits (−19 %), suggesting that fruit AsA metabolism also depends on leaf irradiance. Conclusions Under fruit shading only, the absence of a correlation between sugars and reduced AsA content indicated that fruit AsA content was not limited by leaf photosynthesis or sugar substrate, but strongly depended on fruit irradiance. Leaf shading most probably affected fruit AsA content by delaying fruit ripening, and suggested a complex regulation of AsA metabolism which depends on both fruit and leaf irradiance and fruit ripening stage. PMID:19033285
NASA Astrophysics Data System (ADS)
Mayes, M. T.; Estes, L. D.; Gago, X.; Debats, S. R.; Caylor, K. K.; Manfreda, S.; Oudemans, P.; Ciraolo, G.; Maltese, A.; Nadal, M.; Estrany, J.
2016-12-01
Leaf area is an important ecosystem variable that relates to vegetation biomass, productivity, water and nutrient use in natural and agricultural systems globally. Since the 1980s, optical satellite image-based estimates of leaf area based on indices such as Normalized Difference Vegetation Index (NDVI) have greatly improved understanding of vegetation structure, function, and responses to disturbance at landscape (10^3 km2) to continental (10^6 km2) spatial scales. However, at landscape scales, satellites have failed to capture many leaf area patterns indicative of vegetation succession, crop types, stress and other conditions important for ecological processes. Small drones (UAS - unmanned aerial systems) offer new means for assessing leaf area and vegetation structure at higher spatial resolutions (<1 m) and land cover features such as substrate exposure that may affect estimates of vegetation structure in satellite data. Yet it is unclear how differences in spatial and spectral resolution between UAS and satellite data affect their relationships to each other, and to common field measurements of leaf area (e.g. LiCOR photosensors) and land cover. Constraining these relationships is important for leveraging UAS data to improve scaling of field data on leaf area and biomass to satellite data from Landsat, Sentinel-2, and increasing numbers of commercial sensors. Here, we quantify relationships among field, UAS and satellite estimates of vegetation leaf area and biomass in three case study landscapes spanning semi-arid Mediterranean (Matera, Southern Italy and Mallorca, Spain) and North American temperate ecosystems (New Jersey, USA). We assess how land cover and sensor spectral characteristics affect UAS and satellite-derived NDVI, leaf-area and biomass estimates. Then, we assess the fidelity of UAS, WorldView-2, and Landsat leaf-area and biomass estimates to field-measured landscape changes and variability, including vegetation recovery from fire (Mallorca), and leaf-area and biomass variability due to orchard type and agro-ecosystem management (Matera, New Jersey). Finally, we highlight promising ways forward for improving field data collection and the use of UAS observations to monitor vegetation leaf-area and biomass change at landscape scales in natural and agricultural systems.
Selection in Europeans on Fatty Acid Desaturases Associated with Dietary Changes
Buckley, Matthew T.; Racimo, Fernando; Allentoft, Morten E.; Jensen, Majken K.; Jonsson, Anna; Huang, Hongyan; Hormozdiari, Farhad; Sikora, Martin; Marnetto, Davide; Eskin, Eleazar; Jørgensen, Marit E.; Grarup, Niels; Pedersen, Oluf; Hansen, Torben; Kraft, Peter; Willerslev, Eske
2017-01-01
Abstract FADS genes encode fatty acid desaturases that are important for the conversion of short chain polyunsaturated fatty acids (PUFAs) to long chain fatty acids. Prior studies indicate that the FADS genes have been subjected to strong positive selection in Africa, South Asia, Greenland, and Europe. By comparing FADS sequencing data from present-day and Bronze Age (5–3k years ago) Europeans, we identify possible targets of selection in the European population, which suggest that selection has targeted different alleles in the FADS genes in Europe than it has in South Asia or Greenland. The alleles showing the strongest changes in allele frequency since the Bronze Age show associations with expression changes and multiple lipid-related phenotypes. Furthermore, the selected alleles are associated with a decrease in linoleic acid and an increase in arachidonic and eicosapentaenoic acids among Europeans; this is an opposite effect of that observed for selected alleles in Inuit from Greenland. We show that multiple SNPs in the region affect expression levels and PUFA synthesis. Additionally, we find evidence for a gene–environment interaction influencing low-density lipoprotein (LDL) levels between alleles affecting PUFA synthesis and PUFA dietary intake: carriers of the derived allele display lower LDL cholesterol levels with a higher intake of PUFAs. We hypothesize that the selective patterns observed in Europeans were driven by a change in dietary composition of fatty acids following the transition to agriculture, resulting in a lower intake of arachidonic acid and eicosapentaenoic acid, but a higher intake of linoleic acid and α-linolenic acid. PMID:28333262
USDA-ARS?s Scientific Manuscript database
Cotton leaf curl disease (CLCuD) in the Indian subcontinent is associated with several distinct monopartite begomoviruses and DNA satellites. However, only a single begomovirus was associated with breakdown of resistance against CLCuD in previously resistant cotton varieties. The monopartite begomov...
Steven L. Voelker; J. Renee Brooks; Frederick C. Meinzer; Rebecca Anderson; Martin K.-F. Bader; Giovanna Battipaglia; Katie M. Becklin; David Beerling; Didier Bert; Julio L. Betancourt; Todd E. Dawson; Jean-Christophe Domec; Richard P. Guyette; Christian K??rner; Steven W. Leavitt; Sune Linder; John D. Marshall; Manuel Mildner; Jerome Ogee; Irina Panyushkina; Heather J. Plumpton; Kurt S. Pregitzer; Matthias Saurer; Andrew R. Smith; Rolf T. W. Siegwolf; Michael C. Stambaugh; Alan F. Talhelm; Jacques C. Tardif; Peter K. Van de Water; Joy K. Ward; Lisa Wingate
2016-01-01
Rising atmospheric [CO2], ca, is expected to affect stomatal regulation of leaf gas-exchange of woody plants, thus influencing energy fluxes as well as carbon (C), water, and nutrient cycling of forests. Researchers have proposed various strategies for stomatal regulation of leaf gas-exchange that include maintaining a constant leaf internal [CO...
NASA Technical Reports Server (NTRS)
Latimer, J. G.; Mitchell, C. A.
1988-01-01
Container-grown eggplant (Solanum melongena L. var esculentum Nees. 'Burpee's Black Beauty') seedlings were conditioned with brief, periodic mechanical stress or abscisic acid (ABA) in a greenhouse prior to outdoor exposure. Mechanical stress consisted of seismic (shaking) or thigmic (stem flexing) treatment. Exogenous ABA (10(-3) or 10(-4)M) was applied as a soil drench 3 days prior to outdoor transfer. During conditioning, only thigmic stress reduced stem elongation and only 10(-3) M ABA reduced relative growth rate (RGR). Both conditioning treatments increased leaf specific chlorophyll content, but mechanical stress did not affect leaf ABA content. Outdoor exposure of unconditioned eggplant seedlings decreased RGR and leaf-specific chlorophyll content, but tended to increase leaf ABA content relative to that of plants maintained in the greenhouse. Conditioning did not affect RGR of plants subsequently transferred outdoors, but did reduce stem growth. Seismic stress applied in the greenhouse reduced dry weight gain by plants subsequently transferred outdoors. Mechanical stress treatments increased leaf water potential by 18-25% relative to that of untreated plants.
Comparison of Upward and Downward Translocation of 14C From a Single Leaf of Sunflower
Shiroya, Michi
1968-01-01
When single leaves attached at a given node were allowed to carry on photosynthesis in 14CO2 for 30 min, younger plants showed a higher proportion of upward translocation than did older plants. Downward translocation of 14C-photosynthate was stimulated by ATP pre-treatment of the translocating leaf, while upward translocation was not affected by ATP. A similar phenomenon was observed in the translocation of 14C-sucrose infiltrated into a leaf with or without ATP. Downward translocation of photosynthate was inhibited by DNP pre-treatment of a fed leaf. Upward translocation, however, was not affected by DNP. Thirty min after infiltration of 14C-glucose into a leaf, almost all the 14C translocated upwards was found to be in the form of glucose, while a great part of the 14C translocated downwards was in the form of sucrose. In the case of translocation of infiltrated 14C-sucrose, 14C found both above and below the fed leaf was mainly in the form of sucrose. PMID:16656944
Peach leaf responses to soil and cement dust pollution.
Maletsika, Persefoni A; Nanos, George D; Stavroulakis, George G
2015-10-01
Dust pollution can negatively affect plant productivity in hot, dry and with high irradiance areas during summer. Soil or cement dust were applied on peach trees growing in a Mediterranean area with the above climatic characteristics. Soil and cement dust accumulation onto the leaves decreased the photosynthetically active radiation (PAR) available to the leaves without causing any shade effect. Soil and mainly cement dust deposition onto the leaves decreased stomatal conductance, photosynthetic and transpiration rates, and water use efficiency due possibly to stomatal blockage and other leaf cellular effects. In early autumn, rain events removed soil dust and leaf functions partly recovered, while cement dust created a crust partially remaining onto the leaves and causing more permanent stress. Leaf characteristics were differentially affected by the two dusts studied due to their different hydraulic properties. Leaf total chlorophyll decreased and total phenol content increased with dust accumulation late in the summer compared to control leaves due to intense oxidative stress. The two dusts did not cause serious metal imbalances to the leaves, except of lower leaf K content.
The Neolithic and Bronze Age Monument Complex of Thornborough, North Yorkshire, UK
NASA Astrophysics Data System (ADS)
Harding, Jan
The Neolithic and Bronze Age monument complex of Thornborough in North Yorkshire highlights the significance of heavenly bodies to religious belief and practice. Its investigation by a novel methodology employing virtual reality technology demonstrates alignments with Orion's Belt and the midwinter sunrise. It is argued that these were deliberate, replicating the seasonal movement of those using the complex and thereby suggesting a close relationship between people's skyscape and life cycles.
Low Energy Consumption Hydraulic Techniques
1988-08-30
usually at welds . 1-15 SECTION II PHASE I - ADVANCED AIRCRAFT HYDRAULIC SYSTEM SELECTION Phase I included Task 1 selection of the aircraft and definition...face was bronze plated. The bearings were 52100 tool steel and the pistons were M50 tool steel. The shoe faces were 4140 with bronze plate and the back...o Magnet assembly o Coil assembly DDV Force Motor - -- ,..._(First Stage) oeMain Control Valve __(Second Sae Main Control Valve LVDT Figure 282 Direct
Bronzed cowbird taken in Florida
Matteson, R.E.
1970-01-01
On 8 November 1968 in Gainesville, Florida, I removed a male Bronzed Cowbird (Tangavius a. aeneus) from a blackbird decoy trap containing a large number of Brown-headed Cowbirds (Malothrus ater). Oliver L. Austin, Jr., at the Florida State Museum, verified the species identification by noting the notched inner webs of the outer three primaries, a characteristic of the genus. The subspecific identification was made at the U. S. National Museum where the bird is now specimen number 531666. The subspecies normally ranges from southcentral Texas and the Yucatan Peninsula south through Central America to Panama (Check-list of North American birds, fifth ed., Baltimore, Amer. Ornithol. Union, 1957, p. 542). This Gainesville specimen apparently is the first Bronzed Cow- bird taken in Florida. Alexander Sprunt, Jr., (Florida bird life. In Addendum to Florida bird life, New York, Coward-McCann, 1963, p. 18) lists three photographed sightings at Sarasota, Florida, in April 196
Studies on bronze pre-monetary signs found in Dobroudja using XRF and micro-PIXE
NASA Astrophysics Data System (ADS)
Constantinescu, B.; Cristea-Stan, D.; Talmatchi, G.; Ceccato, D.
2016-03-01
We performed compositional analyses on 180 Scythian-type arrowheads and pre-monetary signs using XRF method and on 60 small fragments of such items (approx. 100 microns diameter), sampling being performed on previously corrosion-cleaned areas on their surface, using micro-PIXE. The items are found in Dobroudja, Istros-Histria region. The most relevant for numismatists result is that for each finding place the same type of alloy was used both for fighting arrowheads and for pre-monetary signs. Our analyses revealed three types of alloys: Cu-Sn-Pb ("normal" bronze), Cu-Sn-Mn-Pb and Cu-Sn-Sb-Pb. The presence of antimony suggests the use of fahlore-type poly-metals deposits, most probably from Caucasus Mountains. The problem of ancient bronze containing manganese is more complicated; an explanation could be the use of manganese oxides as flux necessary to smelt oxidized ores.
NASA Astrophysics Data System (ADS)
Esfandiari, M.; Shirmardi, S. P.; Medhat, M. E.
2014-06-01
In this study, element analysis and the mass attenuation coefficient for matrixes of gold, bronze and water with various impurities and the concentrations of heavy metals (Cu, Mn, Pb and Zn) are evaluated and calculated by the MCNP simulation code for photons emitted from Barium-133, Americium-241 and sources with energies between 1 and 100 keV. The MCNP data are compared with the experimental data and WinXCom code simulated results by Medhat. The results showed that the obtained results of bronze and gold matrix are in good agreement with the other methods for energies above 40 and 60 keV, respectively. However for water matrixes with various impurities, there is a good agreement between the three methods MCNP, WinXCom and the experimental one in low and high energies.
Visual mimicry of host nestlings by cuckoos
Langmore, Naomi E.; Stevens, Martin; Maurer, Golo; Heinsohn, Robert; Hall, Michelle L.; Peters, Anne; Kilner, Rebecca M.
2011-01-01
Coevolution between antagonistic species has produced instances of exquisite mimicry. Among brood-parasitic cuckoos, host defences have driven the evolution of mimetic eggs, but the evolutionary arms race was believed to be constrained from progressing to the chick stage, with cuckoo nestlings generally looking unlike host young. However, recent studies on bronze-cuckoos have confounded theoretical expectations by demonstrating cuckoo nestling rejection by hosts. Coevolutionary theory predicts reciprocal selection for visual mimicry of host young by cuckoos, although this has not been demonstrated previously. Here we show that, in the eyes of hosts, nestlings of three bronze-cuckoo species are striking visual mimics of the young of their morphologically diverse hosts, providing the first evidence that coevolution can select for visual mimicry of hosts in cuckoo chicks. Bronze-cuckoos resemble their own hosts more closely than other host species, but the accuracy of mimicry varies according to the diversity of hosts they exploit. PMID:21227972
Magnetization and magnetoresistance of common alloy wires used in cryogenic instrumentation.
Abrecht, M; Adare, A; Ekin, J W
2007-04-01
We present magnetization and magnetoresistance data at liquid-helium and liquid-nitrogen temperatures for wire materials commonly used for instrumentation wiring of specimens, sensors, and heaters in cryogenic probes. The magnetic susceptibilities in Systeme International units at 4.2 K were found to be: Manganin 1.25x10(-2), Nichrome 5.6x10(-3), and phosphor bronze -3.3x10(-5), indicating that phosphor bronze is the most suitable for high-field applications. We also show the ferromagnetic hysteresis loop of Constantan wire at liquid-helium temperature. The magnetoresistance of these four wires was relatively small: the changes in resistance at 4 K due to a 10 T transverse magnetic field are -2.56% for Constantan, -2.83% for Manganin, +0.69% for Nichrome, and +4.5% for phosphor bronze, compared to about +188% for a typical copper wire under the same conditions.
XRF and micro-PIXE studies of inhomogeneity of ancient bronze and silver alloys
NASA Astrophysics Data System (ADS)
Vasilescu, A.; Constantinescu, B.; Stan, D.; Talmatchi, G.; Ceccato, D.
2017-09-01
New results regarding alloy composition and microstructure for a series of ancient bronze and silver items by X-ray Fluorescence and micro-Particle Induced X-ray Emission spectrometry were obtained in the framework of an extensive numismatic project (Scythian-type arrowheads, arrowhead-shaped monetary signs and wheel coins produced by Histria, 7th-4th century of BCE, and Dacian Radulesti-Hunedoara-type silver tetradrachms, 2nd-1st century of BCE). In Histria, warfare arrowheads were used for trade with Barbarian neighbors at first, then mechanically modified, next melted and cast as dedicated monetary signs, being, in the end, replaced by wheel coins. Three different types of alloys have been identified, and Cu-Mn and Cu-Pb segregation shown. In a blank for Radulesti-Hunedoara-type coins, Ag-(Cu+Pb) segregation has been demonstrated, suggesting an imperfectly alloyed silver-leaded bronze.
Nettle as a distinct Bronze Age textile plant.
Bergfjord, C; Mannering, U; Frei, K M; Gleba, M; Scharff, A B; Skals, I; Heinemeier, J; Nosch, M-L; Holst, B
2012-01-01
It is generally assumed that the production of plant fibre textiles in ancient Europe, especially woven textiles for clothing, was closely linked to the development of agriculture through the use of cultivated textile plants (flax, hemp). Here we present a new investigation of the 2800 year old Lusehøj Bronze Age Textile from Voldtofte, Denmark, which challenges this assumption. We show that the textile is made of imported nettle, most probably from the Kärnten-Steiermark region, an area which at the time had an otherwise established flax production. Our results thus suggest that the production of woven plant fibre textiles in Bronze Age Europe was based not only on cultivated textile plants but also on the targeted exploitation of wild plants. The Lusehøj find points to a hitherto unrecognized role of nettle as an important textile plant and suggests the need for a re-evaluation of textile production resource management in prehistoric Europe.
Nettle as a distinct Bronze Age textile plant
Bergfjord, C.; Mannering, U.; Frei, K. M.; Gleba, M.; Scharff, A. B.; Skals, I.; Heinemeier, J.; Nosch, M. -L; Holst, B.
2012-01-01
It is generally assumed that the production of plant fibre textiles in ancient Europe, especially woven textiles for clothing, was closely linked to the development of agriculture through the use of cultivated textile plants (flax, hemp). Here we present a new investigation of the 2800 year old Lusehøj Bronze Age Textile from Voldtofte, Denmark, which challenges this assumption. We show that the textile is made of imported nettle, most probably from the Kärnten-Steiermark region, an area which at the time had an otherwise established flax production. Our results thus suggest that the production of woven plant fibre textiles in Bronze Age Europe was based not only on cultivated textile plants but also on the targeted exploitation of wild plants. The Lusehøj find points to a hitherto unrecognized role of nettle as an important textile plant and suggests the need for a re-evaluation of textile production resource management in prehistoric Europe. PMID:23024858
Magnetization and magnetoresistance of common alloy wires used in cryogenic instrumentation
NASA Astrophysics Data System (ADS)
Abrecht, M.; Adare, A.; Ekin, J. W.
2007-04-01
We present magnetization and magnetoresistance data at liquid-helium and liquid-nitrogen temperatures for wire materials commonly used for instrumentation wiring of specimens, sensors, and heaters in cryogenic probes. The magnetic susceptibilities in Systeme International units at 4.2 K were found to be: Manganin 1.25×10-2, Nichrome 5.6×10-3, and phosphor bronze -3.3×10-5, indicating that phosphor bronze is the most suitable for high-field applications. We also show the ferromagnetic hysteresis loop of Constantan wire at liquid-helium temperature. The magnetoresistance of these four wires was relatively small: the changes in resistance at 4 K due to a 10 T transverse magnetic field are -2.56% for Constantan, -2.83% for Manganin, +0.69% for Nichrome, and +4.5% for phosphor bronze, compared to about +188% for a typical copper wire under the same conditions.
NASA Astrophysics Data System (ADS)
Bilgen, Nejat; Olgun, Asim
This paper focuses on the spectroscopic and thermal analysis of the archaeological samples of mortar and plaster from middle Bronze Age and Achaemenid period in Seyitömer Höyük. The composition of the samples was investigated using X-ray diffraction (XRD), scanning electron microscopy (SEM), Fourier transform infrared (FTIR) spectroscopy and thermogravimetric thermal analysis (TG-DTA). The results showed that human used different types of raw materials in the preperation of the mortar and plaster in the Middle Bronze Age and Achaemenid period. The material used in middle Bronze Age contains muscovite whereas the material in Achaemenid period contains albite. Although, the chemical composition of the mortar and plaster used in the period were similar, the calcium content of the plaster is relatively higher than the one of the mortar indicating people's awareness of the binding properties of calcite.
Yang, Yongil; Karlson, Dale
2012-08-01
The cold shock domain is among the most evolutionarily conserved nucleic acid binding domains from prokaryotes to higher eukaryotes, including plants. Although eukaryotic cold shock domain proteins have been extensively studied as transcriptional and post-transcriptional regulators during various developmental processes, their functional roles in plants remains poorly understood. In this study, AtCSP3 (At2g17870), which is one of four Arabidopsis thaliana c old s hock domain proteins (AtCSPs), was functionally characterized. Quantitative RT-PCR analysis confirmed high expression of AtCSP3 in reproductive and meristematic tissues. A homozygous atcsp3 loss-of-function mutant exhibits an overall reduced seedling size, stunted and orbicular rosette leaves, reduced petiole length, and curled leaf blades. Palisade mesophyll cells are smaller and more circular in atcsp3 leaves. Cell size analysis indicated that the reduced size of the circular mesophyll cells appears to be generated by a reduction of cell length along the leaf-length axis, resulting in an orbicular leaf shape. It was also determined that leaf cell expansion is impaired for lateral leaf development in the atcsp3 loss-of-function mutant, but leaf cell proliferation is not affected. AtCSP3 loss-of-function resulted in a dramatic reduction of LNG1 transcript, a gene that is involved in two-dimensional leaf polarity regulation. Transient subcellular localization of AtCSP3 in onion epidermal cells confirmed a nucleocytoplasmic localization pattern. Collectively, these data suggest that AtCSP3 is functionally linked to the regulation of leaf length by affecting LNG1 transcript accumulation during leaf development. A putative function of AtCSP3 as an RNA binding protein is also discussed in relation to leaf development.
Christa P.H. Mulder; Bitty A. Roy; Sabine Gusewell
2008-01-01
Parasite damage strongly affects dynamics of boreal forests. Damage levels may be affected by climate change, either directly or indirectly through changes in properties of host trees. We examined how herbivore and pathogen damage in Alnus viridis subsp. fruticosa (Rupr.) Nym. depend on leaf morphology and chemistry, tree size...
USDA-ARS?s Scientific Manuscript database
Cotton is an economically important crop affected by a number of abiotic and biotic stresses. Cotton leaf curl disease (CLCuD) is caused by virus in the genus Begomovirus (family Geminiviridae), collectively called cotton leaf curl viruses (CLCuVs). It is one of the most devastating virual diseases ...
Army Communicator. Volume 27, Number 3, Fall 2002
2002-01-01
The BOM is presented to individuals who stand above their peers in their contributions to the Signal Regiment and SCRA. A minimum one-year membership...www.gordon.army.mil/ocos/rdiv/. Army Communicator 39 AR – Army regulation BOM – Bronze Order of Mercury BW – Bronze Wahatchee CONUS – continental United States CS...service. Medical technicians checked Dudley’s vital signs but couldn’t determine his status, so they recommended he go to the hospital for a more
Tamil Chola Bronzes and Swamimalai Legacy: Metal Sources and Archaeotechnology
NASA Astrophysics Data System (ADS)
Srinivasan, Sharada
2016-08-01
This review explores the great copper alloy image casting traditions of southern India from archaeometallurgical and ethnometallurgical perspectives. The usefulness of lead isotope ratio and compositional analysis in the finger-printing and art historical study of more than 130 early historic, Pallava, Chola, later Chola, and Vijayanagara sculptures (fifth-eighteenth centuries) is highlighted, including Nataraja, Buddha, Parvati, and Rama images made of copper, leaded bronze, brass, and gilt copper. Image casting traditions at Swamimalai in Tamil Nadu are compared with artistic treatises and with the technical examination of medieval bronzes, throwing light on continuities and changes in foundry practices. Western Indian sources could be pinpointed for a couple of medieval images from lead isotope analysis. Slag and archaeometallurgical investigations suggest the exploitation of some copper and lead-silver sources in the Andhra and Karnataka regions in the early historic Satavahana period and point to probable copper sources for the medieval images in Karnataka, Tamil Nadu, and Andhra Pradesh. The general lower iron content in southern Indian bronzes perhaps renders the proximal copper-magnetite reserves of Seruvila in Sri Lanka as a less likely source. Given the lack of lead deposits in Sri Lanka, however, the match of the lead isotope signatures of a well-known Ceylonese Buddhist Tara in British Museum with a Buddha image from Nagapattinam in Tamil Nadu may underscore ties between the island nation and the southern Indian Tamil regions.
NASA Astrophysics Data System (ADS)
Schiavon, Nick; de Palmas, Anna; Bulla, Claudio; Piga, Giampaolo; Brunetti, Antonio
2016-09-01
A spectrometric protocol combining Energy Dispersive X-Ray Fluorescence Spectrometry with Monte Carlo simulations of experimental spectra using the XRMC code package has been applied for the first time to characterize the elemental composition of a series of famous Iron Age small scale archaeological bronze replicas of ships (known as the ;Navicelle;) from the Nuragic civilization in Sardinia, Italy. The proposed protocol is a useful, nondestructive and fast analytical tool for Cultural Heritage sample. In Monte Carlo simulations, each sample was modeled as a multilayered object composed by two or three layers depending on the sample: when all present, the three layers are the original bronze substrate, the surface corrosion patina and an outermost protective layer (Paraloid) applied during past restorations. Monte Carlo simulations were able to account for the presence of the patina/corrosion layer as well as the presence of the Paraloid protective layer. It also accounted for the roughness effect commonly found at the surface of corroded metal archaeological artifacts. In this respect, the Monte Carlo simulation approach adopted here was, to the best of our knowledge, unique and enabled to determine the bronze alloy composition together with the thickness of the surface layers without the need for previously removing the surface patinas, a process potentially threatening preservation of precious archaeological/artistic artifacts for future generations.
Density-functional studies of tungsten trioxide, tungsten bronzes, and related systems
NASA Astrophysics Data System (ADS)
Ingham, B.; Hendy, S. C.; Chong, S. V.; Tallon, J. L.
2005-08-01
Tungsten trioxide adopts a variety of structures which can be intercalated with charged species to alter the electronic properties, thus forming “tungsten bronzes.” Similar effects are observed upon removing oxygen from WO3 . We present a computational study of cubic and hexagonal alkali bronzes and examine the effects on cell size and band structure as the size of the intercalated ion is increased. With the exception of hydrogen (which is predicted to be unstable as an intercalate), the behavior of the bronzes are relatively consistent. NaWO3 is the most stable of the cubic systems, although in the hexagonal system the larger ions are more stable. The band structures are identical, with the intercalated atom donating its single electron to the tungsten 5d valence band. A study of fractional doping in the NaxWO3 system (0⩽x⩽1) showed a linear variation in cell parameter and a systematic shift in the Fermi level into the conduction band. In the oxygen-deficient WO3-x system the Fermi level undergoes a sudden jump into the conduction band at around x=0.2 . Lastly, three compounds of a layered WO4•α,ω -diaminoalkane hybrid series were studied and found to be insulating, with features in the band structure similar to those of the parent WO3 compound that relate well to experimental UV-visible spectroscopy results.
NASA Astrophysics Data System (ADS)
Hauer, Michél; Henkel, Knuth Michael; Krebs, Sebastian; Kroemmer, Werner
2018-01-01
The highly cavitation erosion-resistant propeller alloys CuAl9Ni5Fe4Mn (Ni-Al-Bronze) and CuMn13Al8Fe3Ni2 (Mn-Al-Bronze) were arc-sprayed using a mixture of nitrogen and 2% of hydrogen as atomizing gas and different traverse speeds. The objective was to identify the influences of the different spraying conditions, such as temperature regime and melting loss, on the resulting residual stress states and coating properties. Residual stresses were measured by the incremental hole-drilling method using ESPI. Temperature measurements were carried out by thermographic imaging. Microstructural, chemical and mechanical analyses were realized to examine adhesive and cohesive properties. Additionally, the cavitation erosion behavior was investigated to analyze cohesive coating properties. The spraying process itself was improved, which was apparent by mainly enhanced deposition efficiency and reduced surface temperatures. The amount of oxides and pores as well as the melting loss of alloying elements were reduced. Moreover, an increased cavitation erosion resistance and thus coating cohesion as well as less residual stresses were identified. The change in atomizing gas diminished the impact of the quenching stresses on the coating properties. In contrast, the adhesive strength, Young's moduli and partially the hardness were slightly reduced. With regard to materials, Ni-Al-Bronze revealed superior coating properties in comparison with Mn-Al-Bronze.
Research and application of surface heat treatment for multipulse laser ablation of materials
NASA Astrophysics Data System (ADS)
Cai, Song; Chen, Genyu; Zhou, Cong
2015-11-01
This study analysed a laser ablation platform and built heat transfer equations for multipulse laser ablation of materials. The equations include three parts: laser emission after the material melt and gasification; end of laser emission after the material melts and there is the presence of a super-hot layer and solid-phase heat transfer changes during material ablation. For each of the three parts, the effects of evaporation, plasma shielding and energy accumulation under the pulse interval were considered. The equations are reasonable, and all the required parameters are only related to the laser parameters and material properties, allowing the model to have a certain versatility and practicability. The model was applied for numerical simulation of the heat transfer characteristics in the multipulse laser ablation of bronze and diamond. Next, experiments were conducted to analyse the topography of a bronze-bonded diamond grinding wheel after multipulse laser ablation. The theoretical analysis and experimental results showed that multipulse laser can merge the truing and dressing on a bronze-bonded diamond grinding wheel. This study provides theoretical guidance for optimising the process parameters in the laser ablation of a bronze-bonded diamond grinding wheel. A comparative analysis showed that the numerical solution to the model is in good agreement with the experimental data, thus verifying the correctness and feasibility of the heat transfer model.
NASA Technical Reports Server (NTRS)
Brombacher, W G; Melton, E R
1931-01-01
Experimental data are presented on the variation of the modulus of rigidity in the temperature range -20 to +50 degrees C. of a number of metals which are of possible use for elastic elements for aircraft and other instruments. The methods of the torsional pendulum was used to determine the modulus of rigidity and its temperature coefficient for aluminum, duralumin, monel metal, brass, phosphor bronze, coin silver, nickel silver, three high carbon steels, and three alloy steels. It was observed that tensile stress affected the values of the modulus by amounts of 1 per cent or less.
Martorell, Sebastian; Medrano, Hipolito; Tomàs, Magdalena; Escalona, José M; Flexas, Jaume; Diaz-Espejo, Antonio
2015-03-01
Previous studies have reported correlation of leaf hydraulic vulnerability with pressure-volume parameters related to cell turgor. This link has been explained on the basis of the effects of turgor on connectivity among cells and tissue structural integrity, which affect leaf water transport. In this study, we tested the hypothesis that osmotic adjustment to water stress would shift the leaf vulnerability curve toward more negative water potential (Ψ leaf ) by increasing turgor at low Ψ leaf . We measured leaf hydraulic conductance (K leaf ), K leaf vulnerability [50 and 80% loss of K leaf (P50 and P80 ); |Ψ leaf | at 50 and 80% loss of K leaf , respectively), bulk leaf water relations, leaf gas exchange and sap flow in two Vitis vinifera cultivars (Tempranillo and Grenache), under two water treatments. We found that P50 , P80 and maximum K leaf decreased seasonally by more than 20% in both cultivars and watering treatments. However, K leaf at 2 MPa increased threefold, while osmotic potential at full turgor and turgor loss point decreased. Our results indicate that leaf resistance to hydraulic dysfunction is seasonally plastic, and this plasticity may be mediated by osmotic adjustment. © 2014 Scandinavian Plant Physiology Society.
Flores, L; Banjac, Z; Farré, M; Larrañaga, A; Mas-Martí, E; Muñoz, I; Barceló, D; Elosegi, A
2014-04-01
The intensification of agriculture has promoted the use of pesticides such as fungicides and insecticides. Many pesticides readily leach into natural water bodies and affect both organisms and ecosystem processes such as leaf breakdown, a crucial process in headwater streams. As leaf breakdown in streams involves sequential steps by different groups of organisms (first microbial conditioning, then invertebrate shredding), pesticides targeting different organisms are likely to affect one or the other step, and a mixture of contaminants might have interactive effects. Our objective was to evaluate the effect of a fungicide (imazalil) and an insecticide (diazinon) on stream fungal and invertebrate activities, and their effects on leaf consumption. After an initial assay to define 'effective concentration' of both pesticides in a laboratory experiment, we manipulated pesticide presence/absence during the conditioning and shredding phases. Both pesticides affected fungal community and reduced the performance of the shredding amphipod Echinogammarus berilloni, and leaf consumption. The impact of pesticides on fungal sporulation depended on the length of the exposure period. In addition, pesticides seemed to cause an energetic imbalance in the amphipod, affecting body condition and mortality. The combined effect of both pesticides was similar to those of the fungicide. Overall, our results show that the effects of pesticide mixtures on leaf breakdown are hard to predict from those observed in either fungi or macroinvertebrate performance. Copyright © 2014 Elsevier B.V. All rights reserved.
NASA Astrophysics Data System (ADS)
Fouache, Eric; Desruelles, Stéphane; Eddargach, Wassel; Cammas, Cecilia; Wattez, Julia; Martin, Chloé; Tengberg, Margareta; Beuzen-Waller, Tara; Cable, Charlotte; Thornton, Christopher
2014-05-01
Registered as a World Heritage Site by UNESCO in 1989, the extensive archaeological site of Bat is situated within the Wadi Sharsah and around the modern village and palm grove of Bat, 24 km from the modern city of Ibri in northwestern Oman. The archaeological remains from the Bronze Age excavated by the Bat Archaeological Project are located in two mains areas. The northern area consists of a chain of low limestone hills cut by wadi tributaries leading to the main Wadi Sharsah. It is characterised by an exceptionnally high density of graves from two successive Bronze Age periods : Hafit (ca. 3100-2700 BCE) and Umm an-Nar (ca. 2700-2000 BCE). South of the Bat cemetery, in the flat part of the valley, there are several large circular structures (known historically as « towers ») and remains from both Hafit and Umm an-Nar periods, as well as later periods. Geomorphological mapping of the floodplain, associated with archaeological survey, have identified walls suggesting that during the Umm an-Nar period there was a system of irrigation which controlled flood water. Sedimentological, malacological, C14 dating and micromorphological studies of a 10 m long and 2.5 m high section located 143 m northeast of the Tower 1146 on the left bank of a small tributary of the Wadi Sharsah provide strong argument for the presence of an irrigation system that began before the Hafit period. New C14 datings confirm this hypothesis. Botanical macro-remains collected during the excavation of early Bronze Age structures at Bat further indicate the presence of date palm gardens since the 3rd millenium BCE allowing the cultivation of several crop species, in particular cereals. Most generally the global palaeoenvironmental reconstruction from our data supports a model of a general trend of aridification from Bronze Age to iron Age. Key words : Bronze Age, Holocene, Geomoephology, Micromorphology, Irrigation, Oman.
When was irrigation first used in Bat (Wadi Sharsah, northwestern Oman)?
NASA Astrophysics Data System (ADS)
Fouache, E.; Desruelles, S.; Eddargach, W.; Cammas, C.; Wattez, J.; Martin, C.; Tengberg, M.; Cable, C.; Thornton, C.
2012-04-01
The extensive archaeological site of Bat, registered as a World Heritage Site by UNESCO in 1989, is situated within the Wadi Sharsah and around the modern village and palm grove of Bat, approximately 24 km from the modern city of Ibri in northwestern Oman. The archaeological remains from the Bronze Age excavated by the Bat Archaeological Project are located in two main areas. The northern area consists of a chain of low limestone hills cut by wadi tributaries leading to the main Wadi Sharsah. It is characterised by an exceptionally high density of graves from two successive Bronze Age periods: Hafit (ca. 3100-2700 BCE) and Umm an-Nar (ca. 2700-2000 BCE). South of the Bat cemetery, in the flat part of the valley, there are several large circular structures (known historically as "towers") and remains from both Hafit and Umm an-Nar periods, as well as later periods. Geomorphological mapping of the floodplain, associated with archaeological survey, have identified walls suggesting that during the Umm an-Nar period there was a system of irrigation which controlled flood water. Sedimentological, malacological, C14 dating and micromorphological studies of a 10 m long and 2.5 m high section located 143 m northeast of the Tower 1146 on the left bank of a small tributary of the Wadi Sharsah provide strong arguments for the presence of an irrigation system that began before the Hafit period. Botanical macro-remains collected during the excavation of early Bronze Age structures at Bat further indicate the presence of date palm gardens since the early 3rd millennium BCE allowing the cultivation of several crop species, in particular cereals. Most generally, the global palaeoenvironmental reconstruction from our data supports a model of a general trend of aridification from Bronze to Iron Ages. Key words : Bronze Age, Holocene, Geomorphology, Micromorphology, Irrigation, Oman
Pulito, Claudio; Mori, Federica; Sacconi, Andrea; Casadei, Luca; Ferraiuolo, Maria; Valerio, Maria Cristina; Santoro, Raffaela; Goeman, Frauke; Maidecchi, Anna; Mattoli, Luisa; Manetti, Cesare; Di Agostino, Silvia; Muti, Paola; Blandino, Giovanni; Strano, Sabrina
2015-07-20
Malignant pleural mesothelioma is a poorly treated neoplasia arising from the pleural mesothelial lining. Here we document that the leaf extract of Cynara scolymus exerts broad antitumoral effects both in vitro and in vivo on mesothelioma cell lines. We found that Cynara scolymus treatment affects strongly cell growth, migration and tumor engraftment of mesothelioma cell lines. Strikingly, dietary feeding with Cynara scolymus leaf extract reduces the growth of mesothelioma xenografted tumors similarly to pemetrexed, a commonly employed drug in the treatment of mesothelioma. In aggregate our findings suggest that leaf extract of Cynara scolymus holds therapeutic potential for the treatment of mesothelioma.
Pulito, Claudio; Mori, Federica; Sacconi, Andrea; Casadei, Luca; Ferraiuolo, Maria; Valerio, Maria Cristina; Santoro, Raffaela; Goeman, Frauke; Maidecchi, Anna; Mattoli, Luisa; Manetti, Cesare; Di Agostino, Silvia; Muti, Paola; Blandino, Giovanni; Strano, Sabrina
2015-01-01
Malignant pleural mesothelioma is a poorly treated neoplasia arising from the pleural mesothelial lining. Here we document that the leaf extract of Cynara scolymus exerts broad antitumoral effects both in vitro and in vivo on mesothelioma cell lines. We found that Cynara scolymus treatment affects strongly cell growth, migration and tumor engraftment of mesothelioma cell lines. Strikingly, dietary feeding with Cynara scolymus leaf extract reduces the growth of mesothelioma xenografted tumors similarly to pemetrexed, a commonly employed drug in the treatment of mesothelioma. In aggregate our findings suggest that leaf extract of Cynara scolymus holds therapeutic potential for the treatment of mesothelioma. PMID:26136339
J. S. King; K. S. Pregitzer; D. R. Zak; M. E. Kubiske; W. E. Holmes
2001-01-01
Rising atmospheric carbon dioxide has the potential to alter leaf litter chemistry, potentially affecting decomposition and rates of carbon and nitrogen cycling in forest ecosystems. This study was conducted to determine whether growth under elevated atmospheric CO2 altered the quality and microbial decomposition of leaf litter of a widely...
Terfa, Meseret Tesema; Solhaug, Knut Asbjørn; Gislerød, Hans Ragnar; Olsen, Jorunn Elisabeth; Torre, Sissel
2013-05-01
Alterations in light quality affect plant morphogenesis and photosynthetic responses but the effects vary significantly between species. Roses exhibit an irradiance-dependent flowering control but knowledge on light quality responses is scarce. In this study we analyzed, the responses in morphology, photosynthesis and flowering of Rosa × hybrida to different blue (B) light proportions provided by light-emitting diodes (LED, high B 20%) and high pressure sodium (HPS, low B 5%) lamps. There was a strong morphological and growth effect of the light sources but no significant difference in total dry matter production and flowering. HPS-grown plants had significantly higher leaf area and plant height, yet a higher dry weight proportion was allocated to leaves than stems under LED. LED plants showed 20% higher photosynthetic capacity (Amax ) and higher levels of soluble carbohydrates. The increase in Amax correlated with an increase in leaf mass per unit leaf area, higher stomata conductance and CO2 exchange, total chlorophyll (Chl) content per area and Chl a/b ratio. LED-grown leaves also displayed a more sun-type leaf anatomy with more and longer palisade cells and a higher stomata frequency. Although floral initiation occurred at a higher leaf number in LED, the time to open flowers was the same under both light conditions. Thereby the study shows that a higher portion of B light is efficient in increasing photosynthesis performance per unit leaf area, enhancing growth and morphological changes in roses but does not affect the total Dry Matter (DM) production or time to open flower. Copyright © Physiologia Plantarum 2012.
Leaf litter decomposition and elemental change in three Appalachian mountain streams of different pH
Steven W. Solada; Sue A. Perry; William B. Perry
1996-01-01
The decomposition of leaf litter provides the primary nutrient source for many of the headwater mountain streams in forested catchments. An investigation of factors affected by global change that influence organic matter decomposition, such as temperature and pH, is important in understanding the dynamics of these systems. We conducted a study of leaf litter elemental...
Huang, Yong-Ju; Evans, Neal; Li, Zi-Qin; Eckert, Maria; Chèvre, Anne-Marie; Renard, Michel; Fitt, Bruce D L
2006-01-01
Near-isogenic Brassica napus lines carrying/lacking resistance gene Rlm6 were used to investigate the effects of temperature and leaf wetness duration on phenotypic expression of Rlm6-mediated resistance. Leaves were inoculated with ascospores or conidia of Leptosphaeria maculans carrying the effector gene AvrLm6. Incubation period to the onset of lesion development, number of lesions and lesion diameter were assessed. Symptomless growth of L. maculans from leaf lesions to stems was investigated using a green fluorescent protein (GFP) expressing isolate carrying AvrLm6. L. maculans produced large grey lesions on Darmor (lacking Rlm6) at 5-25 degrees C and DarmorMX (carrying Rlm6) at 25 degrees C, but small dark spots and 'green islands' on DarmorMX at 5-20 degrees C. With increasing temperature/wetness duration, numbers of lesions/spots generally increased. GFP-expressing L. maculans grew from leaf lesions down leaf petioles to stems on DarmorMX at 25 degrees C but not at 15 degrees C. We conclude that temperature and leaf wetness duration affect the phenotypic expression of Rlm6-mediated resistance in leaves and subsequent L. maculans spread down petioles to produce stem cankers.
NASA Astrophysics Data System (ADS)
Hertanto, B. S.; Kartikasari, L. R.; Swastike, Winny; Cahyadi, M.; Yuliani, A.; Nuhriawangsa, A. M. P.
2017-04-01
The objective of this study was to determine the effect of cincau leaf (Cyclea barbata L.Miers) on the physical properties of milk cincau curd. The materials of this research were milk cow of Local Friesian Holstein and leaves of cincau. This research used one way randomized design. The treatment of this research was concentration ratio between cincau leaf and cow milk (w/v): A1 = 10%:90%; A2 = 20%:80%; A3 = 30%:70%. The data was analyzed using ANOVA, and differences between treatment means were further analysed using Duncan’s New Multiple Range Test. Our study revealed that different concentrations of cincau leaf significantly affected cohesiveness, chewiness, hardness, gumminess, springiness, pH and syneresis (p<0.01). However, it did not affect adhesiveness. In addition, the level of 30% of cincau leaf increased cohesiveness, and the level of 20% increased chewiness, hardness, gumminess, springiness, pH. On the other hand, syneresis decreased at the level of 20%. It can be concluded that the addition cincau leaf up to a level of 20% improved the physical properties of milk cincau curd.
McGrath, Justin M; Karnosky, David F; Ainsworth, Elizabeth A
2010-04-01
Early spring leaf out is important to the success of deciduous trees competing for light and space in dense forest plantation canopies. In this study, we investigated spring leaf flush and how long-term growth at elevated carbon dioxide concentration ([CO(2)]) and elevated ozone concentration ([O(3)]) altered leaf area index development in a closed Populus tremuloides (aspen) canopy. This work was done at the Aspen FACE experiment where aspen clones have been grown since 1997 in conditions simulating the [CO(2)] and [O(3)] predicted for approximately 2050. The responses of two clones were compared during the first month of spring leaf out when CO(2) fumigation had begun, but O(3) fumigation had not. Trees in elevated [CO(2)] plots showed a stimulation of leaf area index (36%), while trees in elevated [O(3)] plots had lower leaf area index (-20%). While individual leaf area was not significantly affected by elevated [CO(2)], the photosynthetic operating efficiency of aspen leaves was significantly improved (51%). There were no significant differences in the way that the two aspen clones responded to elevated [CO(2)]; however, the two clones responded differently to long-term growth at elevated [O(3)]. The O(3)-sensitive clone, 42E, had reduced individual leaf area when grown at elevated [O(3)] (-32%), while the tolerant clone, 216, had larger mature leaf area at elevated [O(3)] (46%). These results indicate a clear difference between the two clones in their long-term response to elevated [O(3)], which could affect competition between the clones, and result in altered genotypic composition in future atmospheric conditions. Published by Elsevier Ltd.
Pectin Methylesterification Impacts the Relationship between Photosynthesis and Plant Growth1[OPEN
Kim, Sang-Jin; Renna, Luciana; Brandizzi, Federica
2016-01-01
Photosynthesis occurs in mesophyll cells of specialized organs such as leaves. The rigid cell wall encapsulating photosynthetic cells controls the expansion and distribution of cells within photosynthetic tissues. The relationship between photosynthesis and plant growth is affected by leaf area. However, the underlying genetic mechanisms affecting carbon partitioning to different aspects of leaf growth are not known. To fill this gap, we analyzed Arabidopsis plants with altered levels of pectin methylesterification, which is known to modulate cell wall plasticity and plant growth. Pectin methylesterification levels were varied through manipulation of cotton Golgi-related (CGR) 2 or 3 genes encoding two functionally redundant pectin methyltransferases. Increased levels of methylesterification in a line over-expressing CGR2 (CGR2OX) resulted in highly expanded leaves with enhanced intercellular air spaces; reduced methylesterification in a mutant lacking both CGR-genes 2 and 3 (cgr2/3) resulted in thin but dense leaf mesophyll that limited CO2 diffusion to chloroplasts. Leaf, root, and plant dry weight were enhanced in CGR2OX but decreased in cgr2/3. Differences in growth between wild type and the CGR-mutants can be explained by carbon partitioning but not by variations in area-based photosynthesis. Therefore, photosynthesis drives growth through alterations in carbon partitioning to new leaf area growth and leaf mass per unit leaf area; however, CGR-mediated pectin methylesterification acts as a primary factor in this relationship through modulation of the expansion and positioning of the cells in leaves, which in turn drive carbon partitioning by generating dynamic carbon demands in leaf area growth and leaf mass per unit leaf area. PMID:27208234
Pectin Methylesterification Impacts the Relationship between Photosynthesis and Plant Growth.
M Weraduwage, Sarathi; Kim, Sang-Jin; Renna, Luciana; C Anozie, Fransisca; D Sharkey, Thomas; Brandizzi, Federica
2016-06-01
Photosynthesis occurs in mesophyll cells of specialized organs such as leaves. The rigid cell wall encapsulating photosynthetic cells controls the expansion and distribution of cells within photosynthetic tissues. The relationship between photosynthesis and plant growth is affected by leaf area. However, the underlying genetic mechanisms affecting carbon partitioning to different aspects of leaf growth are not known. To fill this gap, we analyzed Arabidopsis plants with altered levels of pectin methylesterification, which is known to modulate cell wall plasticity and plant growth. Pectin methylesterification levels were varied through manipulation of cotton Golgi-related (CGR) 2 or 3 genes encoding two functionally redundant pectin methyltransferases. Increased levels of methylesterification in a line over-expressing CGR2 (CGR2OX) resulted in highly expanded leaves with enhanced intercellular air spaces; reduced methylesterification in a mutant lacking both CGR-genes 2 and 3 (cgr2/3) resulted in thin but dense leaf mesophyll that limited CO2 diffusion to chloroplasts. Leaf, root, and plant dry weight were enhanced in CGR2OX but decreased in cgr2/3. Differences in growth between wild type and the CGR-mutants can be explained by carbon partitioning but not by variations in area-based photosynthesis. Therefore, photosynthesis drives growth through alterations in carbon partitioning to new leaf area growth and leaf mass per unit leaf area; however, CGR-mediated pectin methylesterification acts as a primary factor in this relationship through modulation of the expansion and positioning of the cells in leaves, which in turn drive carbon partitioning by generating dynamic carbon demands in leaf area growth and leaf mass per unit leaf area. © 2016 American Society of Plant Biologists. All Rights Reserved.
Hydrogen-isotope permeation barrier
Maroni, Victor A.; Van Deventer, Erven H.
1977-01-01
A composite including a plurality of metal layers has a Cu-Al-Fe bronze layer and at least one outer layer of a heat and corrosion resistant metal alloy. The bronze layer is ordinarily intermediate two outer layers of metal such as austenitic stainless steel, nickel alloys or alloys of the refractory metals. The composite provides a barrier to hydrogen isotopes, particularly tritium that can reduce permeation by at least about 30 fold and possibly more below permeation through equal thicknesses of the outer layer material.
2010-11-23
Disaster Relief? Consider Creation of Campaign Medal National Security Threat? Advocate for Activation of National Defense Service Medal Consider...2) Arrowhead Device: The arrowhead device is a bronze replica of an Indian arrowhead 1/4 inch high. It is a Department of the Army device that...device is a bronze replica of an Indian arrowhead 1/4 inch high. It is a Department of the Army device that is authorized for wear on the AFEM. (3
Structural Engineering. Loads. Design Manual 2.2.
1981-11-01
cast, rolled 534 Locust 46 Bronze, 7.9 to 14% Sn 509 Maple, hard 43 Bronze, aluminum 481 Maple, white 33 Copper , cast, rolled 556 Oak, chestnut 54... Copper ore, pyrites 262 Oak, live 59 Gold, cast, hammered 1205 Oak, red, black 41 Gold, bars, stacked 1133 Oak, white 46 Gold, coin in bags 1084 Pine...Phosphate rock, apatite 200 Glass, crystal 184 Porphyry 172 Hay and straw - bales 20 Pumice, natural 40 Leather 59 Quartz, flint 165 Paper 58 Sandstone
Evolution of the Structure of Cu-1% Sn Bronze under High Pressure Torsion and Subsequent Annealing
NASA Astrophysics Data System (ADS)
Popov, V. V.; Popova, E. N.; Stolbovsky, A. V.; Falahutdinov, R. M.
2018-04-01
The evolution of the structure of tin bronze under the room-temperature high-pressure torsion with different degrees of deformation and the subsequent annealing has been investigated. The thermal stability of the structure formed, namely, its behavior upon annealing in the temperature range of 150-400°C has been studied. The possibility of alloying copper with tin has been analyzed with the purpose of obtaining a thermally stable nanostructure with high strength characteristics.
Hydrostatic Response of Submarine Nickel Aluminum Bronze Valves with Corrosion Damage
2008-07-01
Engineering PO Box 1000 Halifax, Nova Scotia B3J 2X4 Project Manager: Dr. T.S. Koko , 902-425-5101 Contract Number: W7707-078022/001/HAL Contract...Manager: Dr. T.S. Koko , 902-425-5101 ext 243 Contract Number: W7707-078022/001/HAL Contract Scientific Authority: Dr. Y. Wang, 902-427-3035...Hydrostatic Response of Submarine Nickel Aluminum Bronze Valves with Corrosion Damage B.K.C. Yuen; T.S. Koko ; R. Warner; DRDC Atlantic CR 2008
Zubrod, Jochen P; Englert, Dominic; Wolfram, Jakob; Rosenfeldt, Ricki R; Feckler, Alexander; Bundschuh, Rebecca; Seitz, Frank; Konschak, Marco; Baudy, Patrick; Lüderwald, Simon; Fink, Patrick; Lorke, Andreas; Schulz, Ralf; Bundschuh, Mirco
2017-08-01
Leaf litter is a major source of carbon and energy for stream food webs, while both leaf-decomposing microorganisms and macroinvertebrate leaf shredders can be affected by fungicides. Despite the potential for season-long fungicide exposure for these organisms, however, such chronic exposures have not yet been considered. Using an artificial stream facility, effects of a chronic (lasting up to 8 wk) exposure to a mixture of 5 fungicides (sum concentration 20 μg/L) on leaf-associated microorganisms and the key leaf shredder Gammarus fossarum were therefore assessed. While bacterial density and microorganism-mediated leaf decomposition remained unaltered, fungicide exposure reduced fungal biomass (≤71%) on leaves from day 28 onward. Gammarids responded to the combined stress from consumption of fungicide-affected leaves and waterborne exposure with a reduced abundance (≤18%), which triggered reductions in final population biomass (18%) and in the number of precopula pairs (≤22%) but could not fully explain the decreased leaf consumption (19%), lipid content (≤43%; going along with an altered composition of fatty acids), and juvenile production (35%). In contrast, fine particulate organic matter production and stream respiration were unaffected. Our results imply that long-term exposure of leaf-associated fungi and shredders toward fungicides may result in detrimental implications in stream food webs and impairments of detrital material fluxes. These findings render it important to understand decomposer communities' long-term adaptational capabilities to ensure that functional integrity is safeguarded. Environ Toxicol Chem 2017;36:2178-2189. © 2017 SETAC. © 2017 SETAC.
Chondrogiannis, Christos; Grammatikopoulos, George
2016-12-01
Leaf development is influenced by almost all the prevailing environmental conditions as well as from the conditions at the time of bud formation. Furthermore, the growth form of a plant determines the leaf longevity and subsequently the investment in biomass and the internal structure of the mesophyll. Therefore, photosynthetic traits of a growing leaf, though, partly predetermined, should also acclimate to temporal changes during developmental period. In addition, the age of the plant can affect photosynthesis of the growing leaf, yet, in the majority of studies, the age is associated to the size of the plant. To test if the reproductive status of the plant affects the time kinetics of the photosynthetic capacity of a growing leaf and the relative contribution of the plants' growth form to the whole procedure, field measurements were conducted in juveniles (prereproductive individuals) and adults (fully reproductive individuals) of an evergreen sclerophyllous shrub (Nerium oleander), a semi-deciduous dimorphic shrub (Phlomis fruticosa), and a winter deciduous tree with pre-leafing flowering (Cercis siliquastrum). PSII structural and functional integrity was progressively developed in all species, but already completed, only some days after leaf expansion in P. fruticosa. Developing leaf as well as fully developed leaf in adults of C. siliquastrum showed enhanced relative size of the pool of final PSI electron acceptors. Photosynthetic traits between juveniles and adults of P. fruticosa were similar, though the matured leaf of adults exhibited lower transpiration rates and improved water-use efficiency than that of juveniles. Adults of the evergreen shrub attained higher CO 2 assimilation rate than juveniles in matured leaf which can be attributed to higher electron flow devoted to carboxylation, and lower photorespiration rate. The reproductive phase of the plant seemed to be involved in modifications of the PSII and PSI functions of the deciduous tree, in carboxylation and photorespiration traits of the evergreen shrub, and in water conductance efficiency of the semi-deciduous shrub. However, it is interesting, that regardless of the growth form of the plant and the prospective leaf longevity of the developing leaf, adults need to support flowering outmatch juveniles, in terms of photosynthesis.
Determining Prehistoric Mining Practices in Southeastern Europe Using Copper Isotopes
NASA Astrophysics Data System (ADS)
Powell, Wayne; Mathur, Ryan; Bankoff, H. Arthur; Bulatović, Aleksandar; Filipović, Vojislav
2017-04-01
Copper was first smelted from malachite at 5000 BCE in Serbia. There the Eneolithic (Copper Age) began with the production of small jewelry pieces and progressed to the casting of massive copper tools near its end, approximately 2000 years later. However, copper metallurgy in southeastern Europe ceased or significantly decreased in the later third millennium, several centuries before the Bronze Age began. Whether this metallurgical hiatus was the result a cultural shift or depletion of natural resources remains an ongoing subject of debate. It has been speculated that the marked reduction in metal production at the Eneolithic-Bronze Age transition was due to the exhaustion of surficial weathered oxide ores and the technical inability to smelt the underlying sulfide minerals. The behavior of copper isotopes in near-surface environments allows us to differentiate highly weathered oxide ores that occur at Earth's surface from non-weathered sulfide ores that occur at greater depth. The oxidation of copper generates fluids and associated minerals that are enriched in the 65Cu isotope. Thus, oxidative weathering of sulfide ores leads to the development of three stratified isotopic reservoirs for copper: 1) oxides above the water table that are enriched in 65Cu; 2) residual weathered sulfides minerals at the water table that are depleted in 65Cu; and 3) non-fractionated, non-weathered sulfide ore below the water table. And so, the transformative shift to sulfide-based metallurgy will be delineated by a significant decrease in δ65Cu in copper artifacts corresponding to the first use of 65Cu-depleted residual ore. The degree of variability of primary ore composition from numerable ore deposits would likely result in the overlap of copper isotope composition between populations of artifacts. Therefore, shifts in the mean copper isotope values and associated standard deviations would best reflect changes in ores use. A baseline value of -0.2‰ ±0.5 (1) was determined from an average of 164 published measurements from chalcopyrite and bornite from 8 epithermal and massive sulfide deposits. Twenty-two (88%) of Eneolithic artifacts (n=25) have values greater than this, whereas eight (73%) of the Early Bronze age artifacts (n=11) yield compositions less than -0.2‰. The mean of Middle Bronze Age, Late Bronze Age and Early Iron Age (n=86) cluster near -0.2‰. This pattern is consistent with a progression to the mining of ore assemblages from increasing depths through prehistory. The shift from 65Cu-enriched to 65Cu-depleted copper in artifacts across the Eneolithic-Bronze Age boundary at 2500 BCE indicates that accessible near-surface oxide ore reserves were depleted after approximately two millennia of mining, and that the beginning of the Bronze Age in the Balkans corresponded to the acquisition of pyrotechnology which allowed for the extraction of metals from sulfide minerals and the resumption of copper mining activity in the region.
NASA Astrophysics Data System (ADS)
Walaszek, Damian; Senn, Marianne; Wichser, Adrian; Faller, Markus; Wagner, Barbara; Bulska, Ewa; Ulrich, Andrea
2014-09-01
This work describes an evaluation of a strategy for multi-elemental analysis of typical ancient bronzes (copper, lead bronze and tin bronze) by means of laser ablation inductively coupled plasma mass spectrometry (LA-ICPMS).The samples originating from archeological experiments on ancient metal smelting processes using direct reduction in a ‘bloomery’ furnace as well as historical casting techniques were investigated with the use of the previously proposed analytical procedure, including metallurgical observation and preliminary visual estimation of the homogeneity of the samples. The results of LA-ICPMS analysis were compared to the results of bulk composition obtained by X-ray fluorescence spectrometry (XRF) and by inductively coupled plasma mass spectrometry (ICPMS) after acid digestion. These results were coherent for most of the elements confirming the usefulness of the proposed analytical procedure, however the reliability of the quantitative information about the content of the most heterogeneously distributed elements was also discussed in more detail.
NASA Astrophysics Data System (ADS)
Feldshtein, E.; Kiełek, P.; Kiełek, T.; Dyachkova, L.; Letsko, A.
2017-05-01
In the paper, the changes in some mechanical properties and wear behavior of CuSn10 sintered bronze and MMCs based on this bronze reinforced with composite ultrafine aluminide powders FeAl/15 % Al2O3, NiAl/15 % Al2O3 and Ti-46Al-8Cr are described. It was observed that the presence of aluminides in the MMCs leads to an increase in the hardness, but the flexural strength may increase or decrease depending on the type of aluminide. The presence of aluminides in the MMC reduces the wear rate considerably. It is decreased in the direction of FeAl/15 % Al2O3 → NiAl/15 % Al2O3 → Ti-46Al-8Cr aluminides and for the best MMC composition the advantage is about 20 times. In the MMCs wear process, micro-craters are formed on the contact surface and it is the principal reason of a decrease in the wear rate.
A Roman bronze statuette with gilded silver mask from Sardinia: an EDXRF study
NASA Astrophysics Data System (ADS)
Cesareo, Roberto; Brunetti, Antonio; D'Oriano, Rubens; Canu, Alba; Demontis, Gonaria Mattia; Celauro, Angela
2013-12-01
A Roman bronze statuette from the 2nd Century BC was recovered from a nuragic sanctuary close to Florinas, in the north of Sardinia. The facial portion of the statuette is covered by a silver mask, partially gilded and attached to the bronze by tin-lead welding. The silver mask was carefully analyzed by portable energy-dispersive X-ray fluorescence (EDXRF), a non-destructive and non-invasive method. The aim of the analysis was to reconstruct the layered structure of the silver gilt mask, and to determine homogeneity and thickness of the gold, silver and lead-tin sheets. This is possible by using the internal ratio of the X-ray lines, i.e. starting from the surface, Au (L α/L β), Ag (K α/K β), Au-L α/Ag-K α and Pb (L α/L β).The results were compared with those obtained with simulated X-ray spectra, obtained both experimentally and by using the Monte Carlo simulation technique.
Chu, Chenxiao; Yang, Jing; Zhang, Qianqian; Wang, Nana; Niu, Feier; Xu, Xuena; Yang, Jian; Fan, Weiliu; Qian, Yitai
2017-12-20
Flower-like assembly of ultrathin nanosheets composed of anatase and bronze TiO 2 embedded in carbon is successfully synthesized by a simple solvothermal reaction, followed with a high-temperature annealing. As an anode material in sodium-ion batteries, this composite exhibits outstanding electrochemical performances. It delivers a reversible capacity of 120 mA h g -1 over 6000 cycles at 10 C. Even at 100 C, there is still a capacity of 104 mA h g -1 . Besides carbon matrix and hierarchical structure, abundant interfaces between anatase and bronze greatly enhance the performance by offering additional sites for reversible Na + storage and improving the charge-transfer kinetics. The interface enhancements are confirmed by discharge/charge profiles, rate performances, electrochemical impedance spectra, and first-principle calculations. These results offer a new pathway to upgrade the performances of anode materials in sodium-ion batteries.
Casting Simulation of an Austrian Bronze Age Sword Hilt
NASA Astrophysics Data System (ADS)
Pola, Annalisa; Mödlinger, Marianne; Piccardo, Paolo; Montesano, Lorenzo
2015-07-01
Bronze Age swords with a metal hilt can be considered the peak of Bronze Age casting technologies. To reconstruct the casting techniques used more than 3000 years ago, a metal hilted sword of the Schalenknauf type from Lower Austria was studied with the aid of macroscopic analyses and simulation of mold filling and casting solidification. A three-dimensional model of the hilt was created based on optical scanner measurements performed on a hilt recently discovered during archaeological excavations. Three different configurations of the gating system were considered, two on the pommel disk and one on the knob, and the effect of its location on the formation of casting defects was investigated. Three-dimensional computed tomography was used to detect internal defects, such as gas and shrinkage porosity, which were then compared with those calculated by simulation. The best match between actual and predicted hilt quality demonstrated the location of the gating system, which turned out to be on the pommel disk.
Comparative modeling of Bronze Age land use in the Malatya Plain (Turkey)
NASA Astrophysics Data System (ADS)
Arıkan, Bülent; Restelli, Francesca Balossi; Masi, Alessia
2016-03-01
Computational modeling in archeology has proven to be a useful tool in quantifying changes in the paleoenvironment. This especially useful method combines data from diverse disciplines to answer questions focusing on the complex and non-linear aspects of human-environment interactions. The research presented here uses various proxy records to compare the changes in climate during the Bronze Age in the Malatya Plain in eastern Anatolia, which is situated at the northern extremity of northern Mesopotamia. Extensive agropastoral land use modeling was applied to three sites of different size and function in the Malatya Plain during the Early Bronze Age I period to simulate the varying scale and intensity of human impacts in relation to changes in the level of social organization, demography, and temporal length. The results suggest that even in land use types subjected to a light footprint, the scale and intensity of anthropogenic impacts change significantly in relation to the level of social organization.
Animal exploitation in northeast Tibetan Plateau during Neolithic-Bronze Age
NASA Astrophysics Data System (ADS)
Dong, Guanghui; Ren, Lele; Li, Zhipeng; Zhang, Dongju; Chen, Fahu
2017-04-01
The history and driving force for prehistoric human occupation of the Tibetan Plateau has been intensively concerned in recent years. Multidisciplinary studies reveal that human extensively and year-roundly settled below 2500 m asl in northeast Tibetan Plateau (NETP) since 5200BP mainly with the utilization of millet crops, and above 3000 m asl post 3600BP primarily with cold-tolerant barley. However, the strategy for human animal utilization during Neolithic and Bronze periods in NETP still remains enigmatic. We collected and identified hundreds of animal bones from 14 Neolithic and Bronze sites in NETP, with the application of radiocarbon dating, carbon and nitrogen isotopes analysis, and ancient DNA analysis, we explore the changing patterns of animal utilization, and potential human behaviours for taming wild animals in this area throughout late prehistoric times. These works shed lights for understanding when and how husbandry and pastoral economies emerged, and how prehistoric human adapted to the high-cold environment of the Tibetan Plateau..
USDA-ARS?s Scientific Manuscript database
Leaf rust (Puccinia triticina Eriks. & Henn.) is a major disease affecting durum wheat production. The Lr14a leaf rust resistant gene present in the durum wheat cv. Creso and its derivative Colosseo is one of the best characterized leaf rust resistance sources presently deployed in durum wheat breed...
USDA-ARS?s Scientific Manuscript database
Parastagonospora nodorum is a necrotrophic pathogen of wheat, causing Septoria nodorum blotch (SNB) affecting both the leaf and glume. P. nodorum is the major leaf blotch pathogen on spring wheat in Norway. Resistance to the disease is quantitative, but several host-specific interactions between nec...
Rising atmospheric [CO2], ca, is expected to affect stomatal regulation of leaf gas-exchange of woody plants, thus influencing energy fluxes as well as carbon (C), water and nutrient cycling of forests. Researchers have reported that stomata regulate leaf gas-exchange around “set...
USDA-ARS?s Scientific Manuscript database
Rust diseases caused by Puccinia spp. pose a major threat to global wheat production. Puccinia triticina (Pt), an obligate basidiomycete biotroph, causes leaf rust disease which incurs yield losses of up to 50% in wheat. Historically, resistant wheat cultivars have been used to control leaf rust, bu...
[Spectrum Variance Analysis of Tree Leaves Under the Condition of Different Leaf water Content].
Wu, Jian; Chen, Tai-sheng; Pan, Li-xin
2015-07-01
Leaf water content is an important factor affecting tree spectral characteristics. So Exploring the leaf spectral characteristics change rule of the same tree under the condition of different leaf water content and the spectral differences of different tree leaves under the condition of the same leaf water content are not only the keys of hyperspectral vegetation remote sensing information identification but also the theoretical support of research on vegetation spectrum change as the differences in leaf water content. The spectrometer was used to observe six species of tree leaves, and the reflectivity and first order differential spectrum of different leaf water content were obtained. Then, the spectral characteristics of each tree species leaves under the condition of different leaf water content were analyzed, and the spectral differences of different tree species leaves under the condition of the same leaf water content were compared to explore possible bands of the leaf water content identification by hyperspectral remote sensing. Results show that the spectra of each tree leaf have changed a lot with the change of the leaf water content, but the change laws are different. Leaf spectral of different tree species has lager differences in some wavelength range under the condition of same leaf water content, and it provides some possibility for high precision identification of tree species.
Ledger, Paul M; Miras, Yannick; Poux, Matthieu; Milcent, Pierre Yves
2015-01-01
Early human societies and their interactions with the natural world have been extensively explored in palaeoenvironmental studies across Central and Western Europe. Yet, despite an extensive body of scholarship, there is little consideration of the environmental impacts of proto-historic urbanisation. Typically palaeoenvironmental studies of Bronze and Iron Age societies discuss human impact in terms of woodland clearance, landscape openness and evidence for agriculture. Although these features are clearly key indicators of human settlement, and characterise Neolithic and early to Middle Bronze Age impacts at Corent, they do not appear to represent defining features of a protohistoric urban environment. The Late Iron Age Gallic Oppidum of Corent is remarkable for the paucity of evidence for agriculture and strong representation of apophytes associated with disturbance. Increased floristic diversity - a phenomenon also observed in more recent urban environments - was also noted. The same, although somewhat more pronounced, patterns are noted for the Late Bronze Age and hint at the possibility of a nascent urban area. High percentages of pollen from non-native trees such as Platanus, Castanea and Juglans in the late Bronze Age and Gallic period also suggest trade and cultural exchange, notably with the Mediterranean world. Indeed, these findings question the validity of applying Castanea and Juglans as absolute chronological markers of Romanisation. These results clearly indicate the value of local-scale palaeoecological studies and their potential for tracing the phases in the emergence of a proto-historic urban environment.
NASA Astrophysics Data System (ADS)
Duman, M. S.; Kaplan, E.; Cuvalcı, O.
2018-01-01
The present paper is based on experimental studies and numerical simulations on the surface fatigue failure of the PTFE-bronze layered journal bearings under real-time loading. ‘Permaglide Plain Bearings P10’ type journal bearings were experimentally tested under different real time dynamic loadings by using real time journal bearing test system in our laboratory. The journal bearing consists of a PTFE-bronze layer approximately 0.32 mm thick on the steel support layer with 2.18 mm thick. Two different approaches have been considered with in experiments: (i) under real- time constant loading with varying bearing widths, (ii) under different real-time loadings at constant bearing widths. Fatigue regions, micro-crack dispersion and stress distributions occurred at the journal bearing were experimentally and theoretically investigated. The relation between fatigue region and pressure distributions were investigated by determining the circumferential pressure distribution under real-time dynamic loadings for the position of every 10° crank angles. In the theoretical part; stress and deformation distributions at the surface of the journal bearing analysed by using finite element methods to determine the relationship between stress and fatigue behaviour. As a result of this study, the maximum oil pressure and fatigue cracks were observed in the most heavily loaded regions of the bearing surface. Experimental results show that PTFE-Bronze layered journal bearings fatigue behaviour is better than the bearings include white metal alloy.
Structural and compositional analysis of a casting mold sherd from ancient China.
Zong, Yunbing; Yao, Shengkun; Lang, Jianfeng; Chen, Xuexiang; Fan, Jiadong; Sun, Zhibin; Duan, Xiulan; Li, Nannan; Fang, Hui; Zhou, Guangzhao; Xiao, Tiqiao; Li, Aiguo; Jiang, Huaidong
2017-01-01
Casting had symbolic significance and was strictly controlled in the Shang dynasty of ancient China. Vessel casting was mainly distributed around the Shang capital, Yin Ruins, which indicates a rigorous centralization of authority. Thus, for a casting mold to be excavated far from the capital region is rare. In addition to some bronze vessel molds excavated at the Buyao Village site, another key discovery of a bronze vessel mold occurred at Daxinzhuang. The Daxinzhuang site was a core area in the east of Shang state and is an important site to study the eastward expansion of the Shang. Here, combining synchrotron X-rays and other physicochemical analysis methods, nondestructive three-dimensional structure imaging and different elemental analyses were conducted on this mold sherd. Through high penetration X-ray tomography, we obtained insights on the internal structure and discovered some pores. We infer that the generation of pores inside the casting mold sherd was used to enhance air permeability during casting. Furthermore, we suppose that the decorative patterns on the surface were carved and not pasted onto it. Considering the previous compositional studies of bronze vessels, the copper and iron elements were analyzed by different methods. Unexpectedly, a larger amount of iron than of copper was detected on the surface. According to the data analysis and archaeological context, the source of iron on the casting mold sherd could be attributed to local soil contamination. A refined compositional analysis confirms that this casting mold was fabricated locally and used for bronze casting.
NASA Astrophysics Data System (ADS)
Brunelli, D.; Levi, S. T.; Fragnoli, P.; Renzulli, A.; Santi, P.; Paganelli, E.; Martinelli, M. C.
2013-12-01
An integrated microchemical-petrographic approach is here proposed to discriminate the provenance of archaeological pottery artefacts from distinct production centres. Our study focuses on a statistically significant sampling ( n=186) of volcanic temper-bearing potteries representative of the manufacturing and dispersion among the islands of the Aeolian Archipelago during the Bronze Age. The widespread establishment of new settlements and the abundant recovery of Aeolian-made ceramic in southern Italy attest for the increased vitality of the Archipelago during the Capo Graziano culture (Early Bronze Age-Middle Bronze Age 2; 2300-1430 BC). Potteries from three of the main known ancient communities (Lipari, Filicudi and Stromboli) have been studied integrating old collections and newly excavated material. Volcanic tempers have been first investigated through multivariate analyses of relative abundances of mineral and rock clasts along with petrographic characters. In addition, we performed in-situ mineral chemistry microanalyses by Electron Microprobe and Laser Ablation—Inductively Coupled Plasma Mass Spectrometry to assess major and trace element composition of the most common mineral phases. Four Temper Compositional Reference Units have been recognised based on compositional trends. Two units (AI and AX) are unequivocally distinct by their peculiar trace element enrichment and petrographic composition; they mostly contain samples from the sites of Lipari and Stromboli, respectively. Units AIV and AVIII, restricted to the sites of Filicudi and Stromboli, show distinct petrographic characters but overlapped geochemical fingerprints.
DOE Office of Scientific and Technical Information (OSTI.GOV)
Alekseev, Evgeny V., E-mail: e.alekseev@fz-juelich.de; Institut für Kristallographie, RWTH Aachen, Jägerstraße 17–19 D-52066 Aachen; Felbinger, Olivier
K[AsW{sub 2}O{sub 9}], prepared by high-temperature solid-state reaction, is the first member of the arsenate–tungsten bronze family. The structure of K[AsW{sub 2}O{sub 9}] is based on a 3-dimensional (3D) oxotungstate–arsenate framework with the non-centrosymmetric P2{sub 1}2{sub 1}2{sub 1} space group, a=4.9747(3) Å, b=9.1780(8) Å, c=16.681(2) Å. The material was characterized using X-ray diffraction, scanning electron microscopy (SEM), differential scanning calorimetry (DSC), Raman and infrared (IR) spectroscopic techniques. The results of DSC demonstrate that this phase is stable up to 1076 K. Second harmonic generation (SHG) measurements performed on a powder sample demonstrate noticeable (0.1 of LiIO{sub 3}) non-linear optical (NLO)more » activity. - Graphical abstract: K[AsW{sub 2}O{sub 9}], the first member of arsenate–tungsten bronze family exhibit new three dimensional structure type, significant thermal stability and NLO properties. Highlights: • K[AsW{sub 2}O{sub 9}], the first member of the arsenate–tungsten bronze family was synthesized with solid state reaction technique. • Structure of this phase was investigated with X-ray diffraction, IR and Raman spectroscopy and electron microscopy. • Thermal stability of the phase was determinate with DSC techniques. • NLO properties were investigated.« less
Processing parameter optimization for the laser dressing of bronze-bonded diamond wheels
NASA Astrophysics Data System (ADS)
Deng, H.; Chen, G. Y.; Zhou, C.; Li, S. C.; Zhang, M. J.
2014-01-01
In this paper, a pulsed fiber-laser dressing method for bronze-bonded diamond wheels was studied systematically and comprehensively. The mechanisms for the laser dressing of bronze-bonded diamond wheels were theoretically analyzed, and the key processing parameters that determine the results of laser dressing, including the laser power density, pulse overlap ratio, ablation track line overlap ratio, and number of scanning cycles, were proposed for the first time. Further, the effects of these four key parameters on the oxidation-damaged layer of the material surface, the material removal efficiency, the material surface roughness, and the average protrusion height of the diamond grains were explored and summarized through pulsed laser ablation experiments. Under the current experimental conditions, the ideal values of the laser power density, pulse overlap ratio, ablation track line overlap ratio, and number of scanning cycles were determined to be 4.2 × 107 W/cm2, 30%, 30%, and 16, respectively. Pulsed laser dressing experiments were conducted on bronze-bonded diamond wheels using the optimized processing parameters; next, both the normal and tangential grinding forces produced by the dressed grinding wheel were measured while grinding alumina ceramic materials. The results revealed that the normal and tangential grinding forces produced by the laser-dressed grinding wheel during grinding were smaller than those of grinding wheels dressed using the conventional mechanical method, indicating that the pulsed laser dressing technology provides irreplaceable advantages relative to the conventional mechanical dressing method.
Li, Wen-Qiang; Zhang, Min-Juan; Gan, Peng-Fei; Qiao, Lei; Yang, Shuai-Qi; Miao, Hai; Wang, Gang-Feng; Zhang, Mao-Mao; Liu, Wen-Ting; Li, Hai-Feng; Shi, Chun-Hai; Chen, Kun-Ming
2017-12-01
Leaf rolling is considered as one of the most important agronomic traits in rice breeding. It has been previously reported that SEMI-ROLLED LEAF 1 (SRL1) modulates leaf rolling by regulating the formation of bulliform cells in rice (Oryza sativa); however, the regulatory mechanism underlying SRL1 has yet to be further elucidated. Here, we report the functional characterization of a novel leaf-rolling mutant, curled leaf and dwarf 1 (cld1), with multiple morphological defects. Map-based cloning revealed that CLD1 is allelic with SRL1, and loses function in cld1 through DNA methylation. CLD1/SRL1 encodes a glycophosphatidylinositol (GPI)-anchored membrane protein that modulates leaf rolling and other aspects of rice growth and development. The cld1 mutant exhibits significant decreases in cellulose and lignin contents in secondary cell walls of leaves, indicating that the loss of function of CLD1/SRL1 affects cell wall formation. Furthermore, the loss of CLD1/SRL1 function leads to defective leaf epidermis such as bulliform-like epidermal cells. The defects in leaf epidermis decrease the water-retaining capacity and lead to water deficits in cld1 leaves, which contribute to the main cause of leaf rolling. As a result of the more rapid water loss and lower water content in leaves, cld1 exhibits reduced drought tolerance. Accordingly, the loss of CLD1/SRL1 function causes abnormal expression of genes and proteins associated with cell wall formation, cuticle development and water stress. Taken together, these findings suggest that the functional roles of CLD1/SRL1 in leaf-rolling regulation are closely related to the maintenance of cell wall formation, epidermal integrity and water homeostasis. © 2017 The Authors The Plant Journal © 2017 John Wiley & Sons Ltd.
Fini, Alessio; Frangi, Piero; Amoroso, Gabriele; Piatti, Riccardo; Faoro, Marco; Bellasio, Chandra; Ferrini, Francesco
2011-11-01
The aim of this work was to evaluate the effects of selected mycorrhiza obtained in the urban environment on growth, leaf gas exchange, and drought tolerance of containerized plants growing in the nursery. Two-year-old uniform Acer campestre L., Tilia cordata Mill., and Quercus robur L. were inoculated with a mixture of infected roots and mycelium of selected arbuscular (maple, linden) and/or ectomycorrhiza (linden, oak) fungi and grown in well-watered or water shortage conditions. Plant biomass and leaf area were measured 1 and 2 years after inoculation. Leaf gas exchange, chlorophyll fluorescence, and water relations were measured during the first and second growing seasons after inoculation. Our data suggest that the mycelium-based inoculum used in this experiment was able to colonize the roots of the tree species growing in the nursery. Plant biomass was affected by water shortage, but not by inoculation. Leaf area was affected by water regime and, in oak and linden, by inoculation. Leaf gas exchange was affected by inoculation and water stress. V(cmax) and J(max) were increased by inoculation and decreased by water shortage in all species. F(v)/F(m) was also generally higher in inoculated plants than in control. Changes in PSII photochemistry and photosynthesis may be related to the capacity of inoculated plants to maintain less negative leaf water potential under drought conditions. The overall data suggest that inoculated plants were better able to maintain physiological activity during water stress in comparison to non-inoculated plants.
Ferreira, Verónica; Koricheva, Julia; Duarte, Sofia; Niyogi, Dev K; Guérold, François
2016-03-01
Many streams worldwide are affected by heavy metal contamination, mostly due to past and present mining activities. Here we present a meta-analysis of 38 studies (reporting 133 cases) published between 1978 and 2014 that reported the effects of heavy metal contamination on the decomposition of terrestrial litter in running waters. Overall, heavy metal contamination significantly inhibited litter decomposition. The effect was stronger for laboratory than for field studies, likely due to better control of confounding variables in the former, antagonistic interactions between metals and other environmental variables in the latter or differences in metal identity and concentration between studies. For laboratory studies, only copper + zinc mixtures significantly inhibited litter decomposition, while no significant effects were found for silver, aluminum, cadmium or zinc considered individually. For field studies, coal and metal mine drainage strongly inhibited litter decomposition, while drainage from motorways had no significant effects. The effect of coal mine drainage did not depend on drainage pH. Coal mine drainage negatively affected leaf litter decomposition independently of leaf litter identity; no significant effect was found for wood decomposition, but sample size was low. Considering metal mine drainage, arsenic mines had a stronger negative effect on leaf litter decomposition than gold or pyrite mines. Metal mine drainage significantly inhibited leaf litter decomposition driven by both microbes and invertebrates, independently of leaf litter identity; no significant effect was found for microbially driven decomposition, but sample size was low. Overall, mine drainage negatively affects leaf litter decomposition, likely through negative effects on invertebrates. Copyright © 2015 Elsevier Ltd. All rights reserved.
Acid rain attack on outdoor sculpture in perspective
NASA Astrophysics Data System (ADS)
Livingston, Richard A.
2016-12-01
A major concern motivating research in acid rain materials effects has been the potential for damage to cultural heritage, particularly outdoor marble and bronze sculpture. However, a combination of field and laboratory studies has failed to show a correlation between rain pH and loss of materials. In order to understand this counterintuitive lack of acid rain effect, an aqueous geochemical modeling approach was used to analyze rain runoff chemistry for the relative importance of acid rain neutralization, dry deposition, and in the case of marble, natural carbonate dissolution. This approach involved the development of pH - SO42- phase diagrams for marble (calcium carbonate) and bronze (copper) under ambient environmental conditions. This then enabled reaction path modeling of the acid neutralization process using the pH range typically found in wet deposition (3.5-6). The results were for marble that the theoretical maximum amount of Ca2+ ion that could be lost due acid rain neutralization would be 0.158 mmol/l compared to 10.5 mmol/l by dry deposition, and for bronze, the Cu2+ ion losses would be 0.21 mmol/l and 47.3 mmol/l respectively. Consequently dry deposition effects on these materials have the potential to dominate over wet deposition effects. To test these predictions the geochemical models were applied to examples of data sets from mass balance (runoff vs rainfall) studies on a marble statue in New York City and some bronze memorial plaques at Gettysburg PA. Although these data sets were collected in the early 1980s they remain valid for demonstrating the mass balance method. For the marble statue, the mean Ca2+ losses by dry deposition was about 69% of the total compared 0.3% for acid rain neutralization, which was less than the natural carbonate dissolution losses of 0.8%. For the bronze, the mean Cu2+ losses were 70.6% by SO42- dry deposition and 23% by NO3- dry deposition compared to 6.4% by acid rain neutralization. Thus for both cases the wet deposition component was less than the variability of the dry deposition components, which explains the observed lack of correlation between the rain pH and the material losses. In addition, for the marble case, there was evidence for HCl acid vapor attack resulting from nitric acid/sea salt interactions and for bronze, ammonium ion may be important. In both cases, significant imbalances suggested that unmeasured organic acids may have a significant effect.
Impact of Meloidogyne incognita on Physiological Efficiency of Vitis vinifera.
Melakeberhan, H; Ferris, H
1989-01-01
Four-week-old French Colombard plants rooted from green cuttings were inoculated with 0, 1,000, 2,000, 4,000, or 8,000 Meloidogyne incognita second-stage juveniles and maintained at 25 C night and 30 C day. Leaf area and dry weight and the rates of photosynthesis, stomatal conductance, and internal leaf CO concentration were measured at intervals up to 59 days after inoculation. Nematode stress dosage, measured as the product of cumulative number of juveniles and females and their total energy (calories) demand, was up to 3.4 kcal and accounted for up to 15% of the energy assimilated by the plants. There was a decline in the rate of leaf area expansion and leaf, stem, shoot, root (excluding nematode weight), and total plant dry weight with increasing nematode stress. Root weight including nematodes was not affected. Total respiration, plant photosynthesis, energy assimilated into plant tissue and respiration, and gross production efficiency decreased significantly with nematode stress. Photosynthetic rate, transpiration rate, stomatal conductance, and internal CO concentration were not affected. This study demonstrates that the energy demand for growth and reproduction of M. incognita accounts for a significant portion of the total energy entering the plant system. As a result, less energy is partitioned into leaf area expansion which, in turn, affects the energy entering the system and results in decreased productivity of nematode-infected grape vines.
NASA Astrophysics Data System (ADS)
Seitz, Steffen; Goebes, Philipp; Assmann, Thorsten; Schuldt, Andreas; Scholten, Thomas
2017-04-01
In subtropical parts of China, high rainfall intensities cause continuous soil losses and thereby provoke severe harms to ecosystems. In woodlands, it is not the tree canopy, but mostly an intact forest floor that provides protection from soil erosion. Although the protective role of leaf litter covers against soil losses is known for a long time, little research has been conducted on the processes involved. For instance, the role of different leaf species and leaf species diversity has been widely disregarded. Furthermore, the impact of soil meso- and macrofauna within the litter layer on soil losses remains unclear. To investigate how leaf litter species and diversity as well as soil meso- and macrofauna affect sediment discharge in a subtropical forest ecosystem, a field experiment was carried out in Xingangshan, Jiangxi Province, PR China (BEF China). A full-factorial random design with 96 micro-scale runoff plots and seven domestic leaf species in three diversity levels and a bare ground feature were established. Erosion was initiated with a rainfall simulator. This study confirms that leaf litter cover generally protects forest soils from water erosion (-82 % sediment discharge on leaf covered plots compared to bare plots) and this protection is gradually removed as the litter layer decomposes. Different leaf species showed variable impacts on sediment discharge and thus erosion control. This effect can be related to different leaf habitus, leaf decomposition rates and food preferences of litter decomposing meso- and macrofauna. In our experiment, runoff plots with leaf litter from Machilus thunbergii in monoculture showed the highest sediment discharge (68.0 g m-2), whereas plots with Cyclobalanopsis glauca in monoculture showed the smallest rates (7.9 g m-2). At the same time, neither leaf species diversity, nor functional diversity showed any significant influence, only a negative trend could be observed. Nevertheless, the protective effect of the leaf litter layer was influenced by the presence (or absence) of soil meso- and macrofauna. Fauna presence increased soil erosion rates significantly by 58 %. It was assumed that this faunal effect arose from arthropods loosening and processing the soil surface as well as fragmenting and decomposing the protecting leaf litter covers. Thus, effects of this fauna group on sediment discharge have to be considered in soil erosion experiments.
Tsutsumi, Koichi; Konno, Masae; Miyazawa, Shin-Ichi; Miyao, Mitsue
2014-02-01
Elevated CO2 concentrations (eCO2) trigger various plant responses. Despite intensive studies of these responses, the underlying mechanisms remain obscure. In this work, we investigated when and how leaf physiology and anatomy are affected by eCO2 in rice plants. We analyzed the most recently fully expanded leaves that developed successively after transfer of the plant to eCO2. To discriminate between the effects of eCO2 and those of nitrogen deficiency, we used three different levels of N application. We found that a decline in the leaf soluble protein content (on a leaf area basis) at eCO2 was only observed under N deficiency. The length and width of the leaf blade were reduced by both eCO2 and N deficiency, whereas the blade thickness was increased by eCO2 but was not affected by N deficiency. The change in length by eCO2 became detectable in the secondly fully expanded leaf, and those in width and thickness in the thirdly fully expanded leaf, which were at the leaf developmental stages P4 and P3, respectively, at the onset of the eCO2 treatment. The decreased blade length at eCO2 was associated with a decrease in the epidermal cell number on the adaxial side and a reduction in cell length on the abaxial side. The decreased width resulted from decreased numbers of small vascular bundles and epidermal cell files. The increased thickness was ascribed mainly to enhanced development of bundle sheath extensions at the ridges of vascular bundles. These observations enable us to identify the sites of action of eCO2 on rice leaf development.
NASA Astrophysics Data System (ADS)
Santiago, L. S.; Sickman, J. O.; Goulden, M.; DeVan, C.; Pasquini, S. C.; Pivovaroff, A. L.
2011-12-01
Leaf carbon isotopic composition and leaf water isotopic enrichment reflect physiological processes and are important for linking local and regional scale processes to global patterns. We investigated how seasonality affects the isotopic composition of bulk leaf carbon, leaf sugar carbon, and leaf water hydrogen under a Mediterranean climate. Leaf and stem samples were collected monthly from four tree species (Calocedrus decurrens, Pinus lambertiana, Pinus ponderosa, and Quercus chrysolepis) at the James San Jacinto Mountain Reserve in southern California. Mean monthly bulk leaf carbon isotopic composition varied from -34.5 % in P. ponderosa to -24.7 % in P. lambertiana and became more depleted in 13C from the spring to the summer. Mean monthly leaf sugar varied from -29.3 % in P. ponderosa to -21.8 % in P. lambertiana and was enriched in 13C during the winter, spring and autumn, but depleted during the mid-summer. Leaf water hydrogen isotopic composition was 28.4 to 68.8 % more enriched in deuterium than source water and this enrichment was greater as seasonal drought progressed. These data indicate that leaf carbon and leaf water hydrogen isotopic composition provide sensitive measures that connect plant physiological processes to short-term climatic variability.
Elevated CO{sub 2} and leaf shape: Are dandelions getting toothier?
DOE Office of Scientific and Technical Information (OSTI.GOV)
Thomas, S.C.; Bazzaz, F.A.
1996-01-01
Heteroblastic leaf development in Taraxacum officinale is compared between plants grown under ambient (350 ppm) vs. elevated (700 ppm) CO{sub 2} levels. Leaves of elevated CO{sub 2} plants exhibited more deeply incised leaf margins and relatively more slender leaf laminae than leaves of ambient CO{sub 2} plants. These differences were found to be significant in allometric analyses that controlled for differences in leaf size, as well as analyses that controlled for leaf development order. The effects of elevated CO{sub 2} on leaf shape were most pronounced when plants were grown individually, but detectable differences were also found in plants grownmore » at high density. Although less dramatic than in Taraxacum, significant effects of elevated CO{sub 2} on leaf shape were also found in two other weedy rosette species, Plantago major and Rumex crispus. These observations support the long-standing hypothesis that leaf carbohydrate level plays an important role in regulating heteroblastic leaf development, though elevated CO{sub 2} may also affect leaf development through direct hormonal interactions or increased leaf water potential. In Taraxacum, pronounced modifications of leaf shape were found at CO{sub 2} levels predicted to occur within the next century. 33 refs., 5 figs.« less
Effects of ammonium sulfate aerosols on vegetation—II. Mode of entry and responses of vegetation
NASA Astrophysics Data System (ADS)
Gmur, Nicholas F.; Evans, Lance S.; Cunningham, Elizabeth A.
These experiments were designed to provide information on the rates of aerosol deposition, mode of entry, and effects of deposition of submicrometer ammonium sulfate aerosols on foliage of Phaseolus vulgaris L. A deposition velocity of 3.2 × 10 3cms-1 was constant during 3-week exposures of plants to aerosol concentrations of 26mg m -3 (i.e. about two orders of magnitude above ambient episode concentrations). Mean deposition rate on foliage was 4.1 × 10 -11 μg cm -2s -1. Visible injury symptoms included leaf chlorosis, necrosis and loss of turgor. Chlorosis was most frequent near leaf margins causing epinasty and near major veins. Internal injury occurred initially in spongy mesophyll cells. Eventually abaxial epidermal and palisade parenchyma cells were injured. These results suggest that submicrometer aerosols enter abaxial stomata and affect more internal cells before affecting leaf surface cells. Exposure to aerosols decreased both abaxial and adaxial leaf resistances markedly. Although visible injury to foliage occurred, no changes in dry mass of roots and shoots or leaf area occurred. These results suggest that for the plant developmental stage studied, while leaf resistances decreased and cellular injury occurred in foliage, these factors were not significantly related to plant growth and development.
NASA Astrophysics Data System (ADS)
Leonard, N. E.
2005-05-01
As wetlands are invaded by Chinese tallow trees (Triadica sebifera), native trees are displaced and detrital inputs to amphibian breeding ponds are altered. I used a mesocosm experiment to examine the effect of Chinese tallow leaf litter on the survival to, size at, and time to metamorphosis of amphibian larvae. Fifty 1000-L cattle watering tanks were treated with 1500 g dry weight of one of five leaf litter treatments: Chinese tallow, laurel oak (Quercus laurifolia), water tupelo (Nyssa aquatica), slash pine (Pinus elliottii), or a 3:1:1:1 mixture. Each tank received 45 tadpoles of Pseudacris feriarum, Bufo terrestris, and Hyla cinerea in sequence according to their natural breeding phonologies. Every Pseudacris feriarum and Bufo terrestris tadpole exposed to Chinese tallow died prior to metamorphosis. Hyla cinerea survival in tanks with tallow-only was significantly lower than that observed for all other leaf treatments. Hyla cinerea tadpoles from tallow-only and mixed-leaf treatments were larger at metamorphosis and transformed faster than those in tanks with native leaves only. These results suggest that Chinese tallow leaf litter may negatively affect tadpoles of early breeding frogs and that Chinese tallow invasion may change the structure of amphibian communities in temporary ponds.
Yang, Bin; Knyazikhin, Yuri; Lin, Yi; Yan, Kai; Chen, Chi; Park, Taejin; Choi, Sungho; Mõttus, Matti; Rautiainen, Miina; Myneni, Ranga B.; Yan, Lei
2017-01-01
Leaf scattering spectrum is the key optical variable that conveys information about leaf absorbing constituents from remote sensing. It cannot be directly measured from space because the radiation scattered from leaves is affected by the 3D canopy structure. In addition, some radiation is specularly reflected at the surface of leaves. This portion of reflected radiation is partly polarized, does not interact with pigments inside the leaf and therefore contains no information about its interior. Very little empirical data are available on the spectral and angular scattering properties of leaf surfaces. Whereas canopy-structure effects are well understood, the impact of the leaf surface reflectance on estimation of leaf absorption spectra remains uncertain. This paper presents empirical and theoretical analyses of angular, spectral, and polarimetric measurements of light reflected by needles and shoots of Pinus koraiensis and Picea koraiensis species. Our results suggest that ignoring the leaf surface reflected radiation can result in an inaccurate estimation of the leaf absorption spectrum. Polarization measurements may be useful to account for leaf surface effects because radiation reflected from the leaf surface is partly polarized, whereas that from the leaf interior is not. PMID:28868160
Fei, Hailong; Wu, Xiaomin; Li, Huan; Wei, Mingdeng
2014-02-01
A simple and versatile method for preparation of novel sodium intercalated (NH4)2V6O16 is developed via a simple hydrothermal route. It is found that ammonium sodium vanadium bronze displays higher discharge capacity and better rate cyclic stability than ammonium vanadium bronze as lithium-ion battery cathode material because of smaller charge transfer resistance, which would favor superior discharge capacity and rate performance. Crown Copyright © 2013. Published by Elsevier Inc. All rights reserved.
Formation of nanocrystalline tetragonal oxide tungsten bronzes on platinum
NASA Astrophysics Data System (ADS)
Kosov, A. V.; Semerikova, O. L.; Vakarin, S. V.; Pankratov, A. A.; Plaksin, S. V.; Zaykov, Yu. P.
2017-02-01
Cyclic voltammetry is used to study the formation of tetragonal oxide tungsten bronze of the composition K x Na y WO3 on a Pt(110) substrate during electrodeposition from a K2WO4-Na2WO4-WO3 melt. The potential ranges in which cathode products of various compositions and morphologies form are found. K x Na(0.66- x)WO3 crystals are shown to form according to the nucleation/growth mechanism. A general scheme is proposed and used to write equations for cathode reactions.
Minoan and Mycenaean medicine and its Near Eastern contacts.
Arnott, R
2004-01-01
Few scholars of ancient medicine have considered that Hippocratic practice may be based in part upon the experience and tradition, handed-down from generation to generation, starting before the end of the Bronze Age in 1100 BC. This paper examines the evidence for medical practitioners in the Aegean in the second millennium BC, and of medical contacts between the Aegean and contemporary bronze age societies of Egypt and the Near East at this time, and suggests that some of these contacts may have been the start of Near Eastern influence on Greek medicine.
Tests of Lead-bronze Bearings in the DVL Bearing-testing Machine
NASA Technical Reports Server (NTRS)
Fischer, G
1940-01-01
The lead-bronze bearings tested in the DVL machine have proven themselves very sensitive to load changes as in comparison with bearings of light metal. In order to prevent surface injuries and consequently running interruptions, the increase of the load has to be made in small steps with sufficient run-in time between steps. The absence of lead in the running surface, impurities in the alloy (especially iron) and surface irregularities (pores) decreases the load-carrying capacity of the bearing to two or three times that of the static load.
Inverse Calibration Free fs-LIBS of Copper-Based Alloys
NASA Astrophysics Data System (ADS)
Smaldone, Antonella; De Bonis, Angela; Galasso, Agostino; Guarnaccio, Ambra; Santagata, Antonio; Teghil, Roberto
2016-09-01
In this work the analysis by Laser Induced Breakdown Spectroscopy (LIBS) technique of copper-based alloys having different composition and performed with fs laser pulses is presented. A Nd:Glass laser (Twinkle Light Conversion, λ = 527 nm at 250 fs) and a set of bronze and brass certified standards were used. The inverse Calibration-Free method (inverse CF-LIBS) was applied for estimating the temperature of the fs laser induced plasma in order to achieve quantitative elemental analysis of such materials. This approach strengthens the hypothesis that, through the assessment of the plasma temperature occurring in fs-LIBS, straightforward and reliable analytical data can be provided. With this aim the capability of the here adopted inverse CF-LIBS method, which is based on the fulfilment of the Local Thermodynamic Equilibrium (LTE) condition, for an indirect determination of the species excitation temperature, is shown. It is reported that the estimated temperatures occurring during the process provide a good figure of merit between the certified and the experimentally determined composition of the bronze and brass materials, here employed, although further correction procedure, like the use of calibration curves, can be demanded. The reported results demonstrate that the inverse CF-LIBS method can be applied when fs laser pulses are used even though the plasma properties could be affected by the matrix effects restricting its full employment to unknown samples provided that a certified standard having similar composition is available.
Holes in teeth - Dental caries in Neolithic and Early Bronze Age populations in Central Germany.
Nicklisch, Nicole; Ganslmeier, Robert; Siebert, Angelina; Friederich, Susanne; Meller, Harald; Alt, Kurt W
2016-01-01
This study provides diachronic insight into the epidemiology of carious defects in teeth of Neolithic and Early Bronze Age populations in Central Germany over a period of 4000 years. The data were retrieved from skeletal remains uncovered at 21 sites throughout the Middle Elbe-Saale region (MES), comprising a total of 494 individuals with preserved teeth. The data generated were examined for age- and sex-related differences in order to gain information about the dietary habits and socio-economic structures of the period with the goal of identifying potential diachronic changes. The results indicated that dietary habits changed over the course of the Neolithic period: the prevalence of caries significantly decreased between the Early and Late Neolithic. The adults from the Early Neolithic sample, particularly those from the LBK bore the highest rate of caries. This highlights the essential importance of cereals in the diet of the early farmers in the Middle Elbe-Saale region. As time went on, meat and dairy products became more and more important, which had a positive impact on dental health. The data also show sex-specific differences: women were more often affected by caries than men and female jaws also generally exhibited greater numbers of carious teeth than their male counterparts. Dental health is a reflection of both biological factors and of economic and sociocultural structures. Copyright © 2015 Elsevier GmbH. All rights reserved.
DOE Office of Scientific and Technical Information (OSTI.GOV)
Crutsinger, Greg; Habenicht, Melissa N; Classen, Aimee T
2008-01-01
Plant-insect interactions can alter ecosystem processes, especially if the insects modify plant architecture, quality, or the quantity of leaf litter inputs. In this study, we investigated the interactions between the gall midge Rhopalomyia solidaginis and tall goldenrod, Solidago altissima, to quantify the degree to which the midge alters plant architecture and how the galls affect rates of litter decomposition and nutrient release in an old-field ecosystem. R. solidaginis commonly leads to the formation of a distinct apical rosette gall on S. altissima and approximately 15% of the ramets in a S. altissima patch were galled (range: 3-34%). Aboveground biomass ofmore » galled ramets was 60% higher and the leaf area density was four times greater on galled leaf tissue relative to the portions of the plant that were not affected by the gall. Overall decomposition rate constants did not differ between galled and ungalled leaf litter. However, leaf-litter mass loss was lower in galled litter relative to ungalled litter, which was likely driven by modest differences in initial litter chemistry; this effect diminished after 12 weeks of decomposition in the field. The proportion of N remaining was always higher in galled litter than in ungalled litter at each collection date indicating differential release of nitrogen in galled leaf litter. Several studies have shown that plant-insect interactions on woody species can alter ecosystem processes by affecting the quality or quantity of litter inputs. Our results illustrate how plant-insect interactions in an herbaceous species can affect ecosystem processes by altering the quality and quantity of litter inputs. Given that S. altissima dominates fields and roadsides and that R. solidaginis galls are highly abundant throughout eastern North America, these interactions are likely to be important for both the structure and function of old-field ecosystems.« less
Ochagavía, Helga; Prieto, Paula; Savin, Roxana; Griffiths, Simon; Slafer, GustavoA
2018-04-27
Wheat adaptation is affected by Ppd genes, but the role of these alleles in the rates of leaf and spikelet initiation has not been properly analysed. Twelve near isogenic lines (NILs) combining Ppd-1a alleles from different donors introgressed in A, B, and/or D genomes were tested under field conditions during two growing seasons together with the wild type, Paragon. Leaf initiation rate was unaffected by Ppd-1a alleles so the final leaf number (FLN) was reduced in parallel with reductions in the duration of the vegetative phase. Spikelet primordia initiation was accelerated and consequently the effect on spikelets per spike was less than proportional to the effect on the duration of spikelet initiation. The magnitude of these effects on spikelet plastochron depended on the doses of Ppd-1 homoeoalleles and the specific insensitivity alleles carried. Double ridge was consistently later than floral initiation, but the difference between them was not affected by Ppd-1a alleles. These findings have potential for selecting the best combinations from the Ppd-1 homoeoallelic series for manipulating adaptation taking into consideration particular effects on spikelet number.
Ochagavía, Helga; Prieto, Paula; Griffiths, Simon
2018-01-01
Abstract Wheat adaptation is affected by Ppd genes, but the role of these alleles in the rates of leaf and spikelet initiation has not been properly analysed. Twelve near isogenic lines (NILs) combining Ppd-1a alleles from different donors introgressed in A, B, and/or D genomes were tested under field conditions during two growing seasons together with the wild type, Paragon. Leaf initiation rate was unaffected by Ppd-1a alleles so the final leaf number (FLN) was reduced in parallel with reductions in the duration of the vegetative phase. Spikelet primordia initiation was accelerated and consequently the effect on spikelets per spike was less than proportional to the effect on the duration of spikelet initiation. The magnitude of these effects on spikelet plastochron depended on the doses of Ppd-1 homoeoalleles and the specific insensitivity alleles carried. Double ridge was consistently later than floral initiation, but the difference between them was not affected by Ppd-1a alleles. These findings have potential for selecting the best combinations from the Ppd-1 homoeoallelic series for manipulating adaptation taking into consideration particular effects on spikelet number. PMID:29562296
Leaf-trait plasticity and species vulnerability to climate change in a Mongolian steppe.
Liancourt, Pierre; Boldgiv, Bazartseren; Song, Daniel S; Spence, Laura A; Helliker, Brent R; Petraitis, Peter S; Casper, Brenda B
2015-09-01
Climate change is expected to modify plant assemblages in ways that will have major consequences for ecosystem functions. How climate change will affect community composition will depend on how individual species respond, which is likely related to interspecific differences in functional traits. The extraordinary plasticity of some plant traits is typically neglected in assessing how climate change will affect different species. In the Mongolian steppe, we examined whether leaf functional traits under ambient conditions and whether plasticity in these traits under altered climate could explain climate-induced biomass responses in 12 co-occurring plant species. We experimentally created three probable climate change scenarios and used a model selection procedure to determine the set of baseline traits or plasticity values that best explained biomass response. Under all climate change scenarios, plasticity for at least one leaf trait correlated with change in species performance, while functional leaf-trait values in ambient conditions did not. We demonstrate that trait plasticity could play a critical role in vulnerability of species to a rapidly changing environment. Plasticity should be considered when examining how climate change will affect plant performance, species' niche spaces, and ecological processes that depend on plant community composition. © 2015 John Wiley & Sons Ltd.
Lotus leaf extract and L-carnitine influence different processes during the adipocyte life cycle.
Siegner, Ralf; Heuser, Stefan; Holtzmann, Ursula; Söhle, Jörn; Schepky, Andreas; Raschke, Thomas; Stäb, Franz; Wenck, Horst; Winnefeld, Marc
2010-08-05
The cellular and molecular mechanisms of adipose tissue biology have been studied extensively over the last two decades. Adipose tissue growth involves both an increase in fat cell size and the formation of mature adipocytes from precursor cells. To investigate how natural substances influence these two processes, we examined the effects of lotus leaf extract (Nelumbo nucifera-extract solution obtained from Silab, France) and L-carnitine on human preadipocytes and adipocytes. For our in vitro studies, we used a lotus leaf extract solution alone or in combination with L-carnitine. Utilizing cultured human preadipocytes, we investigated lotus leaf extract solution-induced inhibition of triglyceride incorporation during adipogenesis and possible effects on cell viability. Studies on human adipocytes were performed aiming to elucidate the efficacy of lotus leaf extract solution to stimulate lipolytic activity. To further characterize lotus leaf extract solution-mediated effects, we determined the expression of the transcription factor adipocyte determination and differentiation factor 1 (ADD1/SREBP-1c) on the RNA- and protein level utilizing qRT-PCR and immunofluorescence analysis. Additionally, the effect of L-carnitine on beta-oxidation was analyzed using human preadipocytes and mature adipocytes. Finally, we investigated additive effects of a combination of lotus leaf extract solution and L-carnitine on triglyceride accumulation during preadipocyte/adipocyte differentiation. Our data showed that incubation of preadipocytes with lotus leaf extract solution significantly decreased triglyceride accumulation during adipogenesis without affecting cell viability. Compared to controls, adipocytes incubated with lotus leaf extract solution exhibited a significant increase in lipolysis-activity. Moreover, cell populations cultivated in the presence of lotus leaf extract solution showed a decrease in adipocyte differentiation capacity as indicated by a decrease in the ADD1/SREBP-1c signal. Importantly, our results demonstrated that a combination of lotus leaf extract solution and L-carnitine reduced triglyceride accumulation to a greater extent compared to incubation with either substance alone. Overall, our data demonstrate that a combination of lotus leaf extract and L-carnitine reduced triglyceride accumulation in human (pre)adipocytes by affecting different processes during the adipocyte life cycle. For this reason, this combination might represent a treatment option for obesity-related diseases.
Rozema, Jelte; Cornelisse, Danny; Zhang, Yuancheng; Li, Hongxiu; Bruning, Bas; Katschnig, Diana; Broekman, Rob; Ji, Bin; van Bodegom, Peter
2015-01-01
Salt tolerance of higher plants is determined by a complex set of traits, the timing and rate of evolution of which are largely unknown. We compared the salt tolerance of cultivars of sugar beet and their ancestor, sea beet, in hydroponic studies and evaluated whether traditional domestication and more recent breeding have changed salt tolerance of the cultivars relative to their ancestor. Our comparison of salt tolerance of crop cultivars is based on values of the relative growth rate (RGR) of the entire plant at various salinity levels. We found considerable salt tolerance of the sea beet and slightly, but significantly, reduced salt tolerance of the sugar beet cultivars. This indicates that traditional domestication by selection for morphological traits such as leaf size, beet shape and size, enhanced productivity, sugar content and palatability slightly affected salt tolerance of sugar beet cultivars. Salt tolerance among four sugar beet cultivars, three of which have been claimed to be salt tolerant, did not differ. We analysed the components of RGR to understand the mechanism of salt tolerance at the whole-plant level. The growth rate reduction at higher salinity was linked with reduced leaf area at the whole-plant level (leaf area ratio) and at the individual leaf level (specific leaf area). The leaf weight fraction was not affected by increased salinity. On the other hand, succulence and leaf thickness and the net assimilation per unit of leaf area (unit leaf rate) increased in response to salt treatment, thus partially counteracting reduced capture of light by lower leaf area. This compensatory mechanism may form part of the salt tolerance mechanism of sea beet and the four studied sugar beet cultivars. Together, our results indicate that domestication of the halophytic ancestor sea beet slightly reduced salt tolerance and that breeding for improved salt tolerance of sugar beet cultivars has not been effective. PMID:25492122
Zhang, Chaowen; Chen, Feifan; Zhao, Ziyao; Hu, Liangliang; Liu, Hanqiang; Cheng, Zhihui; Weng, Yiqun; Chen, Peng; Li, Yuhong
2018-06-01
Two round-leaf mutants, rl-1 and rl-2, were identified from EMS-induced mutagenesis. High throughput sequencing and map-based cloning suggested CsPID encoding a Ser/Thr protein kinase as the most possible candidate for rl-1. Rl-2 was allelic to Rl-1. Leaf shape is an important plant architecture trait that is affected by plant hormones, especially auxin. In Arabidopsis, PINOID (PID), a regulator for the auxin polar transporter PIN (PIN-FORMED) affects leaf shape formation, but this function of PID in crop plants has not been well studied. From an EMS mutagenesis population, we identified two round-leaf (rl) mutants, C356 and C949. Segregation analysis suggested that both mutations were controlled by single recessive genes, rl-1 and rl-2, respectively. With map-based cloning, we show that CsPID as the candidate gene of rl-1; a non-synonymous SNP in the second exon of CsPID resulted in an amino acid substitution and the round leaf phenotype. As compared in the wild type plant, CsPID had significantly lower expression in the root, leaf and female flowers in C356, which may result in the less developed roots, round leaves and abnormal female flowers, respectively in the rl-1 mutant. Among the three copies of PID genes, CsPID, CsPID2 and CSPID2L (CsPID2-like) in the cucumber genome, CsPID was the only one with significantly differential expression in adult leaves between WT and C356 suggesting CsPID plays a main role in leaf shape formation. The rl-2 mutation in C949 was also cloned, which was due to another SNP in a nearby location of rl-1 in the same CsPID gene. The two round leaf mutants and the work presented herein provide a good foundation for understanding the molecular mechanisms of CsPID in cucumber leaf development.
Xie, Hongtao; Yu, Mukui; Cheng, Xiangrong
2018-03-01
Light availability greatly affects plant growth and development. In shaded environments, plants must respond to reduced light intensity to ensure a regular rate of photosynthesis to maintain the dynamic balance of nutrients, such as leaf non-structural carbohydrates (NSCs), carbon (C), nitrogen (N) and phosphorus (P). To improve our understanding of the nutrient utilization strategies of understory shade-tolerant plants, we compared the variations in leaf NSCs, C, N and P in response to heterogeneous controlled light conditions between two subtropical evergreen broadleaf shade-tolerant species, Elaeocarpus sylvestris (E. sylvestris) and Illicium henryi (I. henryi). Light intensity treatments were applied at five levels (100%, 52%, 33%, 15% and 6% full sunlight) for 30 weeks to identify the effects of reduced light intensity on leaf NSC allocation patterns and leaf C:N:P stoichiometry characteristics. We found that leaf soluble sugar, starch and NSC concentrations in E. sylvestris showed decreasing trends with reduced light intensity, whereas I. henryi presented slightly increasing trends from 100% to 15% full sunlight and then significant decreases at extremely low light intensity (6% full sunlight). The soluble sugar/starch ratio of E. sylvestris decreased with decreasing light intensity, whereas that of I. henryi remained stable. Moreover, both species exhibited increasing trends in leaf N and P concentrations but limited leaf N:P and C:P ratio fluctuations with decreasing light intensity, revealing their adaptive strategies for poor light environments and their growth strategies under ideal light environments. There were highly significant correlations between leaf NSC variables and C:N:P stoichiometric variables in both species, revealing a trade-off in photosynthesis production between leaf NSC and carbon allocation. Thus, shade-tolerant plants readjusted their allocation of leaf NSCs, C, N and P in response to light acclimation. Redundancy analysis showed that leaf morphological features of both E. sylvestris and I. henryi affected their corresponding leaf nutrient traits. These results improve our understanding of the dynamic balance between leaf NSCs and leaf C, N and P components in the nutritional metabolism of shade-tolerant plants. Two species of understory shade-tolerant plants responded differently to varying light intensities in terms of leaf non-structural carbohydrate allocation and the utilization of carbon, nitrogen and phosphorus to balance nutritional metabolism and adapt to environmental stress. Copyright © 2018 Elsevier Masson SAS. All rights reserved.
Coca Leaf and Cocaine Addiction: Some Historical Notes
Blejer-Prieto, H.
1965-01-01
Coca-leaf habituation has affected millions of Andean natives for over 400 years. In the last half-century it has also involved millions more Malayans. Coca leaf, from which cocaine and extracts for some commercial carbonated soft drinks are obtained, remains relatively unknown by the medical and allied professions elsewhere. A review of the original medical, historical and other pertinent literature of the last 350 years illustrates the origins of the use of coca leaf, its spread, the isolation of cocaine and its first uses, as well as some of the euphoric and other effects of both substances. PMID:5318484
NASA Astrophysics Data System (ADS)
Riccucci, Cristina; Ingo, Gabriel Maria; Faustoferri, Amalia; Pierigè, Maria Isabella; Parisi, Erica Isabella; Di Carlo, Gabriella; De Caro, Tilde; Faraldi, Federica
2013-12-01
The Samnite bronze belts and the chest disk cuirasses (VIII-IV BC) are the distinctive defensive weapons of the Samnite warriors having likely also a symbolic relevance. These artefacts were mainly found during the archaeological excavations of warriors' graves from ancient Abruzzo (central Italy). Their chemical composition, metallurgical features and corrosion products formed during the long-term burial have been studied by means of the combined use of analytical techniques such as optical microscopy (OM), scanning electron microscopy coupled with energy-dispersive X-ray micro-analysis (SEM-EDS), X-ray photoelectron spectroscopy (XPS) and X-ray diffraction (XRD). The micro-chemical and structural results show that the bronze belts have often been produced by using unusual high-tin bronze alloys achieving a silver-like appearance and by performing tailored cycles of thermal treatments under reducing conditions and hot mechanical working aimed to shape the high-tin alloys in the form of a thin bronze sheet. Furthermore, the investigation has shown that the main alloying elements have been transformed during the burial into mineral species giving rise to the formation of stratified structures constituted by different mineral phases such as tin oxides, cuprous oxide (Cu2O) and copper carbonates (azurite (Cu3(CO3)2(OH)2 and malachite (CuCO3Cu(OH)2)) as well as dangerous chlorine-based compounds such as nantokite (CuCl) and atacamite (Cu2(OH)3Cl) polymorphs. This information evidences the strict interaction of the alloying elements with the soil components as well as the occurrence of the copper cyclic corrosion as a post-burial degradation phenomenon. The present study confirms that the combined micro-chemical and micro-structural investigation techniques such as SEM-EDS, XPS, XRD and OM can be successfully used to investigate the technological production processes of the ancient artefacts and to achieve the detailed micro-chemical and structural description of the corrosion products useful for the identification of degradation agents and mechanisms and, thereafter, to propose a reliable tailored strategy for the conservation.
Shi, Zuomin; Haworth, Matthew; Feng, Qiuhong; Cheng, Ruimei; Centritto, Mauro
2015-01-01
Plant growth at high elevations necessitates physiological and morphological plasticity to enable photosynthesis (A) under conditions of reduced temperature, increased radiation and the lower partial pressure of atmospheric gases, in particular carbon dioxide (pCO2). Previous studies have observed a wide range of responses to elevation in plant species depending on their adaptation to temperature, elevational range and growth habit. Here, we investigated the effect of an increase in elevation from 2500 to 3500 m above sea level (a.s.l.) on three montane species with contrasting growth habits and leaf economic strategies. While all of the species showed identical increases in foliar δ13C, dark respiration and nitrogen concentration with elevation, contrasting leaf gas exchange and photosynthetic responses were observed between species with different leaf economic strategies. The deciduous shrub Salix atopantha and annual herb Rumex dentatus exhibited increased stomatal (Gs) and mesophyll (Gm) conductance and enhanced photosynthetic capacity at the higher elevation. However, evergreen Quercus spinosa displayed reduced conductance to CO2 that coincided with lower levels of photosynthetic carbon fixation at 3500 m a.s.l. The lower Gs and Gm values of evergreen species at higher elevations currently constrains their rates of A. Future rises in the atmospheric concentration of CO2 ([CO2]) will likely predominantly affect evergreen species with lower specific leaf areas (SLAs) and levels of Gm rather than deciduous species with higher SLA and Gm values. We argue that climate change may affect plant species that compose high-elevation ecosystems differently depending on phenotypic plasticity and adaptive traits affecting leaf economics, as rising [CO2] is likely to benefit evergreen species with thick sclerophyllous leaves. PMID:26433706
Campos, Diana; Alves, Artur; Lemos, Marco F L; Correia, António; Soares, Amadeu M V M; Pestana, João L T
2014-07-01
Detritus processing is vital for freshwater ecosystems that depend on the leaf litter from riparian vegetation and is mediated by microorganisms and aquatic invertebrates. Shredder invertebrates transform coarse particulate organic matter into fine particulate organic matter used as food by collector species. Direct and indirect effects of contaminants can impair detritus processing and thus affect the functioning of these ecosystems. Here, we assessed the combined effects of a toxic metal (cadmium) and resource quality (leaf species) on detritus processing and shredder-collector interactions. We considered two types of leaves, alder and eucalyptus that were microbially conditioned under different Cd concentrations in the laboratory. The microbial communities present on leaves were analyzed by Denaturing Gradient Gel Electrophoresis (DGGE), and we also measured microbial respiration rates. Sericostoma vittatum (a caddisfly shredder) and Chironomus riparius (a midge collector) were also exposed to Cd and allowed to consume the corresponding alder or eucalyptus leaves. We evaluated C. riparius growth and leaf mass loss in multispecies microcosms. Cadmium exposure affected leaf conditioning and fungal diversity on both leaf species, as assessed by DGGE. Cadmium exposure also affected the mass loss of alder leaves by reductions in detritivore feeding, and impaired C. riparius growth. Chironomus riparius consumed alder leaf discs in the absence of shredders, but S. vittatum appear to promote C. riparius growth in treatments containing eucalyptus. These results show that indirect effects of contaminants along detritus-processing chains can occur through effects on shredder-collector interactions such as facilitation but they also depend on the nutritional quality of detritus and on sensitivity and feeding plasticity of detritivore species.
Stomatal and pavement cell density linked to leaf internal CO2 concentration
Šantrůček, Jiří; Vráblová, Martina; Šimková, Marie; Hronková, Marie; Drtinová, Martina; Květoň, Jiří; Vrábl, Daniel; Kubásek, Jiří; Macková, Jana; Wiesnerová, Dana; Neuwithová, Jitka; Schreiber, Lukas
2014-01-01
Background and Aims Stomatal density (SD) generally decreases with rising atmospheric CO2 concentration, Ca. However, SD is also affected by light, air humidity and drought, all under systemic signalling from older leaves. This makes our understanding of how Ca controls SD incomplete. This study tested the hypotheses that SD is affected by the internal CO2 concentration of the leaf, Ci, rather than Ca, and that cotyledons, as the first plant assimilation organs, lack the systemic signal. Methods Sunflower (Helianthus annuus), beech (Fagus sylvatica), arabidopsis (Arabidopsis thaliana) and garden cress (Lepidium sativum) were grown under contrasting environmental conditions that affected Ci while Ca was kept constant. The SD, pavement cell density (PCD) and stomatal index (SI) responses to Ci in cotyledons and the first leaves of garden cress were compared. 13C abundance (δ13C) in leaf dry matter was used to estimate the effective Ci during leaf development. The SD was estimated from leaf imprints. Key Results SD correlated negatively with Ci in leaves of all four species and under three different treatments (irradiance, abscisic acid and osmotic stress). PCD in arabidopsis and garden cress responded similarly, so that SI was largely unaffected. However, SD and PCD of cotyledons were insensitive to Ci, indicating an essential role for systemic signalling. Conclusions It is proposed that Ci or a Ci-linked factor plays an important role in modulating SD and PCD during epidermis development and leaf expansion. The absence of a Ci–SD relationship in the cotyledons of garden cress indicates the key role of lower-insertion CO2 assimilation organs in signal perception and its long-distance transport. PMID:24825295
Pollastrini, Martina; Nogales, Ana Garcia; Benavides, Raquel; Bonal, Damien; Finer, Leena; Fotelli, Mariangela; Gessler, Arthur; Grossiord, Charlotte; Radoglou, Kalliopi; Strasser, Reto J; Bussotti, Filippo
2017-02-01
An assemblage of tree species with different crown properties creates heterogeneous environments at the canopy level. Changes of functional leaf traits are expected, especially those related to light interception and photosynthesis. Chlorophyll a fluorescence (ChlF) properties in dark-adapted leaves, specific leaf area, leaf nitrogen content (N) and carbon isotope composition (δ13C) were measured on Picea abies (L.) H.Karst., Pinus sylvestris L. and Betula pendula Roth. in monospecific and mixed boreal forests in Europe, in order to test whether they were affected by stand species richness and composition. Photosynthetic efficiency, assessed by induced emission of leaf ChlF, was positively influenced in B. pendula by species richness, whereas P. abies showed higher photosynthetic efficiency in monospecific stands. Pinus sylvestris had different responses when it coexisted with P. abies or B. pendula. The presence of B. pendula, but not of P. abies, in the forest had a positive effect on the efficiency of photosynthetic electron transport and N in P. sylvestris needles, and the photosynthetic responses were positively correlated with an increase of leaf δ13C. These effects on P. sylvestris may be related to high light availability at the canopy level due to the less dense canopy of B. pendula. The different light requirements of coexisting species was the most important factor affecting the distribution of foliage in the canopy, driving the physiological responses of the mixed species. Future research directions claim to enhance the informative potential of the methods to analyse the responses of pure and mixed forests to environmental factors, including a broader set of plant species' functional traits and physiological responses. © The Author 2017. Published by Oxford University Press. All rights reserved. For Permissions, please email: journals.permissions@oup.com.
Stomatal and pavement cell density linked to leaf internal CO2 concentration.
Santrůček, Jiří; Vráblová, Martina; Simková, Marie; Hronková, Marie; Drtinová, Martina; Květoň, Jiří; Vrábl, Daniel; Kubásek, Jiří; Macková, Jana; Wiesnerová, Dana; Neuwithová, Jitka; Schreiber, Lukas
2014-08-01
Stomatal density (SD) generally decreases with rising atmospheric CO2 concentration, Ca. However, SD is also affected by light, air humidity and drought, all under systemic signalling from older leaves. This makes our understanding of how Ca controls SD incomplete. This study tested the hypotheses that SD is affected by the internal CO2 concentration of the leaf, Ci, rather than Ca, and that cotyledons, as the first plant assimilation organs, lack the systemic signal. Sunflower (Helianthus annuus), beech (Fagus sylvatica), arabidopsis (Arabidopsis thaliana) and garden cress (Lepidium sativum) were grown under contrasting environmental conditions that affected Ci while Ca was kept constant. The SD, pavement cell density (PCD) and stomatal index (SI) responses to Ci in cotyledons and the first leaves of garden cress were compared. (13)C abundance (δ(13)C) in leaf dry matter was used to estimate the effective Ci during leaf development. The SD was estimated from leaf imprints. SD correlated negatively with Ci in leaves of all four species and under three different treatments (irradiance, abscisic acid and osmotic stress). PCD in arabidopsis and garden cress responded similarly, so that SI was largely unaffected. However, SD and PCD of cotyledons were insensitive to Ci, indicating an essential role for systemic signalling. It is proposed that Ci or a Ci-linked factor plays an important role in modulating SD and PCD during epidermis development and leaf expansion. The absence of a Ci-SD relationship in the cotyledons of garden cress indicates the key role of lower-insertion CO2 assimilation organs in signal perception and its long-distance transport. © The Author 2014. Published by Oxford University Press on behalf of the Annals of Botany Company. All rights reserved. For Permissions, please email: journals.permissions@oup.com.
Satora, Leszek; Kozioł, Katarzyna; Zebrowski, Jacek
2017-06-01
Accessory respiratory organs in fish exhibit great diversity but share the presence of numerous capillaries covered by a simple squamous epithelium. The adoption of the intestine for respiratory function needs certain special modifications. In this study, we explored immunohistochemical and metabolic fingerprint features that could underlay this adaptation in bronze corydoras Corydoras aeneus. Immunohistochemical localization of the cytoplasmic domain of epidermal growth factor receptor (EGFR) in the respiratory part of intestine demonstrated a strong positive immunoreaction in epithelial cells and connective tissue. Fourier Transfer Infrared (FTIR) spectroscopy coupled with chemometrics discriminated between anterior and posterior region of intestine in terms of secondary structure of proteins and the abundance of p-cresol and other phenolics. The latter were reduced in the posterior part of intestine, indicating the cessation of digestive function in this region. It has been suggested that aquatic hypoxia via endocrine cells (hypoxia-sensitive) activate EGFR, which induce proliferation of squamous epithelial cells, thereby enabling gas diffusion in the posterior part of intestine. It seems that hypoxia and normoxia are opposed conditions adjusting the production of squamous epithelial cells in this intestine. The physiological role of EGFR in the respiratory intestine of bronze corydoras is of interest not only from an evolutionary aspect but also in terms of a potential model for observations process proliferation squamous epithelial cells. Future investigations on the molecular responses to different water oxygen levels in air-breathing bronze corydoras fish are required to clarify the mechanism responsible for squamous cell proliferation. Copyright © 2017 Elsevier GmbH. All rights reserved.
Structural and compositional analysis of a casting mold sherd from ancient China
Zong, Yunbing; Yao, Shengkun; Lang, Jianfeng; Chen, Xuexiang; Fan, Jiadong; Sun, Zhibin; Duan, Xiulan; Li, Nannan; Fang, Hui; Zhou, Guangzhao; Xiao, Tiqiao; Li, Aiguo; Jiang, Huaidong
2017-01-01
Casting had symbolic significance and was strictly controlled in the Shang dynasty of ancient China. Vessel casting was mainly distributed around the Shang capital, Yin Ruins, which indicates a rigorous centralization of authority. Thus, for a casting mold to be excavated far from the capital region is rare. In addition to some bronze vessel molds excavated at the Buyao Village site, another key discovery of a bronze vessel mold occurred at Daxinzhuang. The Daxinzhuang site was a core area in the east of Shang state and is an important site to study the eastward expansion of the Shang. Here, combining synchrotron X-rays and other physicochemical analysis methods, nondestructive three-dimensional structure imaging and different elemental analyses were conducted on this mold sherd. Through high penetration X-ray tomography, we obtained insights on the internal structure and discovered some pores. We infer that the generation of pores inside the casting mold sherd was used to enhance air permeability during casting. Furthermore, we suppose that the decorative patterns on the surface were carved and not pasted onto it. Considering the previous compositional studies of bronze vessels, the copper and iron elements were analyzed by different methods. Unexpectedly, a larger amount of iron than of copper was detected on the surface. According to the data analysis and archaeological context, the source of iron on the casting mold sherd could be attributed to local soil contamination. A refined compositional analysis confirms that this casting mold was fabricated locally and used for bronze casting. PMID:28296963
Ledger, Paul M.; Miras, Yannick; Poux, Matthieu; Milcent, Pierre Yves
2015-01-01
Early human societies and their interactions with the natural world have been extensively explored in palaeoenvironmental studies across Central and Western Europe. Yet, despite an extensive body of scholarship, there is little consideration of the environmental impacts of proto-historic urbanisation. Typically palaeoenvironmental studies of Bronze and Iron Age societies discuss human impact in terms of woodland clearance, landscape openness and evidence for agriculture. Although these features are clearly key indicators of human settlement, and characterise Neolithic and early to Middle Bronze Age impacts at Corent, they do not appear to represent defining features of a protohistoric urban environment. The Late Iron Age Gallic Oppidum of Corent is remarkable for the paucity of evidence for agriculture and strong representation of apophytes associated with disturbance. Increased floristic diversity – a phenomenon also observed in more recent urban environments – was also noted. The same, although somewhat more pronounced, patterns are noted for the Late Bronze Age and hint at the possibility of a nascent urban area. High percentages of pollen from non-native trees such as Platanus, Castanea and Juglans in the late Bronze Age and Gallic period also suggest trade and cultural exchange, notably with the Mediterranean world. Indeed, these findings question the validity of applying Castanea and Juglans as absolute chronological markers of Romanisation. These results clearly indicate the value of local-scale palaeoecological studies and their potential for tracing the phases in the emergence of a proto-historic urban environment. PMID:25853251
Makki, Mahsasadat; Dupouy-Camet, Jean; Sajjadi, Seyed Mansour Seyed; Naddaf, Saied Reza; Mobedi, Iraj; Rezaeian, Mostafa; Mohebali, Mehdi; Mowlavi, Gholamreza
2017-04-01
Shahr-e Sukhteh (meaning burnt city in Persian) in Iran is an archeological site dated back to around 3,200-1,800 BC. It is located in Sistan and Baluchistan Province of Iran and known as the junction of Bronze Age trade routes crossing the Iranian plateau. It was appointed as current study area for paleoparasitological investigations. Excavations at this site have revealed various archeological materials since 1967. In the present study, sheep and carnivore coprolites excavated from this site were analyzed by means of rehydration technique using TSP solution for finding helminth eggs. Dicrocoelium dendriticum, Capillaria sp., and Taenia sp. eggs were identified, while some other objects similar to Anoplocephalidae and Toxocara spp. eggs were also retrieved from the samples but their measured parameters did not match those of these species. The present paper illustrates the first paleoparasitological findings of Bronze Age in eastern Iran supporting the economic activities, peopling, and communication as well as the appropriate condition for zoonotic helminthiasis life cycle in Shahr-e Sukhteh archeological site.
Makki, Mahsasadat; Dupouy-Camet, Jean; Sajjadi, Seyed Mansour Seyed; Naddaf, Saied Reza; Mobedi, Iraj; Rezaeian, Mostafa; Mohebali, Mehdi; Mowlavi, Gholamreza
2017-01-01
Shahr-e Sukhteh (meaning burnt city in Persian) in Iran is an archeological site dated back to around 3,200-1,800 BC. It is located in Sistan and Baluchistan Province of Iran and known as the junction of Bronze Age trade routes crossing the Iranian plateau. It was appointed as current study area for paleoparasitological investigations. Excavations at this site have revealed various archeological materials since 1967. In the present study, sheep and carnivore coprolites excavated from this site were analyzed by means of rehydration technique using TSP solution for finding helminth eggs. Dicrocoelium dendriticum, Capillaria sp., and Taenia sp. eggs were identified, while some other objects similar to Anoplocephalidae and Toxocara spp. eggs were also retrieved from the samples but their measured parameters did not match those of these species. The present paper illustrates the first paleoparasitological findings of Bronze Age in eastern Iran supporting the economic activities, peopling, and communication as well as the appropriate condition for zoonotic helminthiasis life cycle in Shahr-e Sukhteh archeological site. PMID:28506043
DOE Office of Scientific and Technical Information (OSTI.GOV)
Ajayi, O. O.; Lorenzo-Martin, Cinta
This study presents results of an experimental study to evaluate friction stir processing (FSP) with and without hard second-phase particle incorporation as a means to enhance surface properties and wear performance of C86300 manganese bronze alloy. FSP of flat bronze alloy specimens was conducted with hardened H-13 tool steel to create a 3-mm-thick processed surface layer. The process was also used to incorporate B 4C particles, thereby creating a metal-matrix composite layer on the alloy surface. FSP alone was observed to produce substantial reduction in grain size (from an initial value of 350 mu m to 1-5 μm). FSP withoutmore » particle incorporation resulted in modest surface hardening due to grain refinement and dispersion hardening. Under lubricated contact in block-on-ring testing with a hardened steel counter face, FSP produced substantial reduction (about 3X) in bronze wear after polishing of processing surface roughening. FSP with hard B 4C second-phase particle incorporation further reduced wear by up to 20X. The improvement in wear behavior is attributed to grain refinement and load shielding by second-phase particles, as determined by wear mechanism analysis.« less
Huang, L X
2017-11-28
This compilation process of Mian xue tang zhen jiu ji cheng ( Mian Xuetang ' s Synthetic Work of Acu - moxibustion )was quite mysterious, the book's advent coincidentally met with several major historical events which made its value of decoding its historical functions far beyond its academic value. The editor of this book is Le Xianyang, the founder of Tong Ren Tang. While compiling this book, he also made a replica of Ming-Dynasty's Zhengtong bronze statue. More than 200 years later, Le Jingyu, the 12th generation of the Le family revised Ming Tang Tu inherited from his ancestors and engraved Ming Tang Tu on the stone for its standardization. This great achievement of Le family was no less than the recasting of the Tiansheng bronze statue and the reengraving Tong ren shu xue zhen jiu tu jing ( Illustrated Manual of Acupuncture Points of the Bronze Figure ) by support of state power for building the Beijing Imperial Hospital in the eighth year of the Zhengtong reign (1443).
Nanoscale building blocks in a novel lithium arsenotungsten bronze: Synthesis and characterization
Zhao, Pei; Mangir Murshed, M.; Huq, Ashfia; ...
2015-02-19
Here we report on a novel compound Li 3AsW 7O 25 obtained by solid-state reaction and characterized by diffraction and spectroscopic methods. The bronze-type compound crystallizes in the orthorhombic space group Pbca with a=724.38(3) pm, b=1008.15(4) pm, c=4906.16(17) pm and Z=8. The structure is built up by chains of WO 6 octahedra interconnected by AsO 4 tetrahedra and WO 6 octahedra forming a polyhedral arrangement as seen in intergrowth tungsten bronzes. The X-ray single crystal structure refinement allows solving the complex arsenotungstate framework. The powder neutron diffraction data analysis locates the lithium atoms. Thermal analysis showed that Li 3AsW 7Omore » 25 is stable up to its melting at 1135(3) K followed by a decomposition at 1182(5) K. Finally, the Kubelka-Munk treatment of the UV-vis spectrum revealed a wide band gap in the range of 2.84-3.40 eV depending on the presumed electron transition type.« less
Environmental Roots of the Late Bronze Age Crisis
Kaniewski, David; Van Campo, Elise; Guiot, Joël; Le Burel, Sabine; Otto, Thierry; Baeteman, Cecile
2013-01-01
The Late Bronze Age world of the Eastern Mediterranean, a rich linkage of Aegean, Egyptian, Syro-Palestinian, and Hittite civilizations, collapsed famously 3200 years ago and has remained one of the mysteries of the ancient world since the event’s retrieval began in the late 19th century AD/CE. Iconic Egyptian bas-reliefs and graphic hieroglyphic and cuneiform texts portray the proximate cause of the collapse as the invasions of the “Peoples-of-the-Sea” at the Nile Delta, the Turkish coast, and down into the heartlands of Syria and Palestine where armies clashed, famine-ravaged cities abandoned, and countrysides depopulated. Here we report palaeoclimate data from Cyprus for the Late Bronze Age crisis, alongside a radiocarbon-based chronology integrating both archaeological and palaeoclimate proxies, which reveal the effects of abrupt climate change-driven famine and causal linkage with the Sea People invasions in Cyprus and Syria. The statistical analysis of proximate and ultimate features of the sequential collapse reveals the relationships of climate-driven famine, sea-borne-invasion, region-wide warfare, and politico-economic collapse, in whose wake new societies and new ideologies were created. PMID:23967146
NASA Astrophysics Data System (ADS)
Deckers, Katleen; Pessin, Hugues
2010-09-01
Vegetation changes are reconstructed based on more than 51,000 charcoal fragments of more than 380 samples from nine Bronze Age sites in northern Syria and southern Turkey. In addition to fragment proportions, special attention was paid to the frequency of Pistacia relative to Quercus and Populus/ Salix relative to Tamarix, fruit-tree ubiquity, and riverine diversity in order to gain an improved understanding of the human versus climatic impact on the vegetation. The results indicate that human impacts first took place within the riverine forest. This phase was followed by land clearing within the woodland steppe, especially in the northern portion of the study area. In the south near Emar, the woodland steppe probably disappeared by the Late Bronze Age. It is uncertain whether this was caused by aridification and/or human clearing. The northward shift of the Pistacia-woodland steppe is very likely a result of climatic drying that occurred throughout the entire period under investigation. Although increased deforestation is evident through time, the small proportions of imported wood indicate that local resources were still available.
Zhang, Jianwei; Schaub, Marcus; Ferdinand, Jonathan A; Skelly, John M; Steiner, Kim C; Savage, James E
2010-08-01
We investigated the effect of leaf age on the response of net photosynthesis (A), stomatal conductance (g(wv)), foliar injury, and leaf nitrogen concentration (N(L)) to tropospheric ozone (O(3)) on Prunus serotina seedlings grown in open-plots (AA) and open-top chambers, supplied with either carbon-filtered or non-filtered air. We found significant variation in A, g(wv), foliar injury, and N(L) (P < 0.05) among O(3) treatments. Seedlings in AA showed the highest A and g(wv) due to relatively low vapor pressure deficit (VPD). Older leaves showed significantly lower A, g(wv), N(L), and higher foliar injury (P < 0.001) than younger leaves. Leaf age affected the response of A, g(wv), and foliar injury to O(3). Both VPD and N(L) had a strong influence on leaf gas exchange. Foliar O(3)-induced injury appeared when cumulative O(3) uptake reached 8-12 mmol m(-2), depending on soil water availability. The mechanistic assessment of O(3)-induced injury is a valuable approach for a biologically relevant O(3) risk assessment for forest trees. Published by Elsevier Ltd.
Bacterial Leaf Spot of Lettuce: Request for Samples
USDA-ARS?s Scientific Manuscript database
Bacterial leaf spot of lettuce caused by by Xanthomonas campestris pv. vitians has been affecting coastal California crops for many years and has become a chronic problem. Differences in pathogen genotypes have been demonstrated and correlated to disease responses on resistant and susceptible cultiv...
Ken W. Krauss; Robert R. Twilley; Thomas w. Doyle; Emile S. Gardiner
2006-01-01
We determined how different hydroperiods affected leaf gas exchange characteristics of greenhouse-grown seedlings (2002) and saplings (2003) of the mangrove species Avicennia germinans (L.) Stearn., Laguncularia racemosa (L.) Gaertn. f., and Rhizophora mangle L. Hydroperiod treatments included...
Multi-modal sensor system for plant water stress assessment
USDA-ARS?s Scientific Manuscript database
Plant stress critically affects plant growth and causes significant loss of productivity and quality. When the plant is under water stress, it impedes photosynthesis and transpiration, resulting in changes in leaf color and temperature. Leaf discoloration in photosynthesis can be assessed by measu...
A molecular insight into papaya leaf curl-a severe viral disease.
Varun, Priyanka; Ranade, S A; Saxena, Sangeeta
2017-11-01
Papaya leaf curl disease (PaLCuD) caused by papaya leaf curl virus (PaLCuV) not only affects yield but also plant growth and fruit size and quality of papaya and is one of the most damaging and economically important disease. Management of PaLCuV is a challenging task due to diversity of viral strains, the alternate hosts, and the genomic complexities of the viruses. Several management strategies currently used by plant virologists to broadly control or eliminate the viruses have been discussed. In the absence of such strategies in the case of PaLCuV at present, the few available options to control the disease include methods like removal of affected plants from the field, insecticide treatments against the insect vector (Bemisia tabaci), and gene-specific control through transgenic constructs. This review presents the current understanding of papaya leaf curl disease, genomic components including satellite DNA associated with the virus, wide host and vector range, and management of the disease and suggests possible generic resistance strategies.
Moura, Bárbara B; Alves, Edenise S
2014-11-01
Phenotypic plasticity of the leaves can interfere with the plant sensitivity to ozone (O3) toxic effect. This study aimed to assess whether the leaf structure of Ipomoea nil changes due to climatic variations and whether these changes affect the species' sensitivity. Field exposures, in different seasons (winter and spring) were made. The leaves that developed during the winter were thinner, with a lower proportion of photosynthetic tissues, higher proportion of intercellular spaces and lower density and stomatal index compared to those developed during the spring. The temperature and relative humidity positively influenced the leaf thickness and stomatal index. The visible injuries during winter were positively correlated with the palisade parenchyma thickness and negatively correlated with the percentage of spongy parenchyma; during the spring, the symptoms were positively correlated with the stomatal density. In conclusion, the leaf structure of I. nil varied among the seasons, interfering in its sensitivity to O3. Copyright © 2014 Elsevier Ltd. All rights reserved.
Diversity of Riparian Plants among and within Species Shapes River Communities
Jackrel, Sara L.; Wootton, J. Timothy
2015-01-01
Organismal diversity among and within species may affect ecosystem function with effects transmitting across ecosystem boundaries. Whether recipient communities adjust their composition, in turn, to maximize their function in response to changes in donor composition at these two scales of diversity is unknown. We use small stream communities that rely on riparian subsidies as a model system. We used leaf pack experiments to ask how variation in plants growing beside streams in the Olympic Peninsula of Washington State, USA affects stream communities via leaf subsidies. Leaves from red alder (Alnus rubra), vine maple (Acer cinereus), bigleaf maple (Acer macrophyllum) and western hemlock (Tsuga heterophylla) were assembled in leaf packs to contrast low versus high diversity, and deployed in streams to compare local versus non-local leaf sources at the among and within species scales. Leaves from individuals within species decomposed at varying rates; most notably thin leaves decomposed rapidly. Among deciduous species, vine maple decomposed most rapidly, harbored the least algal abundance, and supported the greatest diversity of aquatic invertebrates, while bigleaf maple was at the opposite extreme for these three metrics. Recipient communities decomposed leaves from local species rapidly: leaves from early successional plants decomposed rapidly in stream reaches surrounded by early successional forest and leaves from later successional plants decomposed rapidly adjacent to later successional forest. The species diversity of leaves inconsistently affected decomposition, algal abundance and invertebrate metrics. Intraspecific diversity of leaf packs also did not affect decomposition or invertebrate diversity. However, locally sourced alder leaves decomposed more rapidly and harbored greater levels of algae than leaves sourced from conspecifics growing in other areas on the Olympic Peninsula, but did not harbor greater aquatic invertebrate diversity. In contrast to alder, local intraspecific differences via decomposition, algal or invertebrate metrics were not observed consistently among maples. These results emphasize that biodiversity of riparian subsidies at the within and across species scale have the potential to affect aquatic ecosystems, although there are complex species-specific effects. PMID:26539714
Diversity of Riparian Plants among and within Species Shapes River Communities.
Jackrel, Sara L; Wootton, J Timothy
2015-01-01
Organismal diversity among and within species may affect ecosystem function with effects transmitting across ecosystem boundaries. Whether recipient communities adjust their composition, in turn, to maximize their function in response to changes in donor composition at these two scales of diversity is unknown. We use small stream communities that rely on riparian subsidies as a model system. We used leaf pack experiments to ask how variation in plants growing beside streams in the Olympic Peninsula of Washington State, USA affects stream communities via leaf subsidies. Leaves from red alder (Alnus rubra), vine maple (Acer cinereus), bigleaf maple (Acer macrophyllum) and western hemlock (Tsuga heterophylla) were assembled in leaf packs to contrast low versus high diversity, and deployed in streams to compare local versus non-local leaf sources at the among and within species scales. Leaves from individuals within species decomposed at varying rates; most notably thin leaves decomposed rapidly. Among deciduous species, vine maple decomposed most rapidly, harbored the least algal abundance, and supported the greatest diversity of aquatic invertebrates, while bigleaf maple was at the opposite extreme for these three metrics. Recipient communities decomposed leaves from local species rapidly: leaves from early successional plants decomposed rapidly in stream reaches surrounded by early successional forest and leaves from later successional plants decomposed rapidly adjacent to later successional forest. The species diversity of leaves inconsistently affected decomposition, algal abundance and invertebrate metrics. Intraspecific diversity of leaf packs also did not affect decomposition or invertebrate diversity. However, locally sourced alder leaves decomposed more rapidly and harbored greater levels of algae than leaves sourced from conspecifics growing in other areas on the Olympic Peninsula, but did not harbor greater aquatic invertebrate diversity. In contrast to alder, local intraspecific differences via decomposition, algal or invertebrate metrics were not observed consistently among maples. These results emphasize that biodiversity of riparian subsidies at the within and across species scale have the potential to affect aquatic ecosystems, although there are complex species-specific effects.
Zhang, Wenting; Mirlohi, Shirin; Li, Xiaorong; He, Yuke
2018-06-01
Leaf traits affect plant agronomic performance; for example, leaf hair number provides a morphological indicator of drought and insect resistance. Brassica rapa crops have diverse phenotypes, and many B. rapa single-nucleotide polymorphisms (SNPs) have been identified and used as molecular markers for plant breeding. However, which SNPs are functional for leaf hair traits and, therefore, effective for breeding purposes remains unknown. Here, we identify a set of SNPs in the B. rapa ssp. pekinenesis candidate gene BrpHAIRY LEAVES1 ( BrpHL1 ) and a number of SNPs of BrpHL1 in a natural population of 210 B. rapa accessions that have hairy, margin-only hairy, and hairless leaves. BrpHL1 genes and their orthologs and paralogs have many SNPs. By intensive mutagenesis and genetic transformation, we selected the functional SNPs for leaf hairs by the exclusion of nonfunctional SNPs and the orthologous and paralogous genes. The residue tryptophan-92 of BrpHL1a was essential for direct interaction with GLABROUS3 and, thus, necessary for the formation of leaf hairs. The accessions with the functional SNP leading to substitution of the tryptophan-92 residue had hairless leaves. The orthologous BrcHL1b from B. rapa ssp. chinensis regulates hair formation on leaf margins rather than leaf surfaces. The selected SNP for the hairy phenotype could be adopted as a molecular marker for insect resistance in Brassica spp. crops. Moreover, the procedures optimized here can be used to explain the molecular mechanisms of natural variation and to facilitate the molecular breeding of many crops. © 2018 American Society of Plant Biologists. All rights reserved.
Jiang, Dan; Fang, Jingjing; Lou, Lamei; Zhao, Jinfeng; Yuan, Shoujiang; Yin, Liang; Sun, Wei; Peng, Lixiang; Guo, Baotai; Li, Xueyong
2015-01-01
Leaf morphology is closely associated with cell division. In rice, mutations in Narrow leaf 1 (NAL1) show narrow leaf phenotypes. Previous studies have shown that NAL1 plays a role in regulating vein patterning and increasing grain yield in indica cultivars, but its role in leaf growth and development remains unknown. In this report, we characterized two allelic mutants of NARROW LEAF1 (NAL1), nal1-2 and nal1-3, both of which showed a 50% reduction in leaf width and length, as well as a dwarf culm. Longitudinal and transverse histological analyses of leaves and internodes revealed that cell division was suppressed in the anticlinal orientation but enhanced in the periclinal orientation in the mutants, while cell size remained unaltered. In addition to defects in cell proliferation, the mutants showed abnormal midrib in leaves. Map-based cloning revealed that nal1-2 is a null allelic mutant of NAL1 since both the whole promoter and a 404-bp fragment in the first exon of NAL1 were deleted, and that a 6-bp fragment was deleted in the mutant nal1-3. We demonstrated that NAL1 functions in the regulation of cell division as early as during leaf primordia initiation. The altered transcript level of G1- and S-phase-specific genes suggested that NAL1 affects cell cycle regulation. Heterogenous expression of NAL1 in fission yeast (Schizosaccharomyces pombe) further supported that NAL1 affects cell division. These results suggest that NAL1 controls leaf width and plant height through its effects on cell division. PMID:25658704
Greenwood, J.L.; Rosemond, A.D.; Wallace, J.B.; Cross, W.F.; Weyers, H.S.
2007-01-01
Most nutrient enrichment studies in aquatic systems have focused on autotrophic food webs in systems where primary producers dominate the resource base. We tested the heterotrophic response to long-term nutrient enrichment in a forested, headwater stream. Our study design consisted of 2 years of pretreatment data in a reference and treatment stream and 2 years of continuous nitrogen (N) + phosphorus addition to the treatment stream. Studies were conducted with two leaf species that differed in initial C:N, Rhododendron maximum (rhododendron) and Acer rubrum (red maple). We determined the effects of nutrient addition on detrital resources (leaf breakdown rates, litter C:N and microbial activity) and tested whether nutrient enrichment affected macroinvertebrate consumers via increased biomass. Leaf breakdown rates were ca. 1.5 and 3?? faster during the first and second years of enrichment, respectively, in the treatment stream for both leaf types. Microbial respiration rates of both leaf types were 3?? higher with enrichment, and macroinvertebrate biomass associated with leaves increased ca. 2-3?? with enrichment. The mass of N in macroinvertebrate biomass relative to leaves tended to increase with enrichment up to 6?? for red maple and up to 44?? for rhododendron leaves. Lower quality (higher C:N) rhododendron leaves exhibited greater changes in leaf nutrient content and macroinvertebrate response to nutrient enrichment than red maple leaves, suggesting a unique response by different leaf species to nutrient enrichment. Nutrient concentrations used in this study were moderate and equivalent to those in streams draining watersheds with altered land use. Thus, our results suggest that similarly moderate levels of enrichment may affect detrital resource quality and subsequently lead to altered energy and nutrient flow in detrital food webs. ?? 2006 Springer-Verlag.
Krauss, Ken W.; Twilley, Robert R.; Doyle, Thomas W.; Gardiner, Emile S.
2006-01-01
We determined how different hydroperiods affected leaf gas exchange characteristics of greenhouse-grown seedlings (2002) and saplings (2003) of the mangrove species Avicennia germinans (L.) Stearn., Laguncularia racemosa (L.) Gaertn. f., and Rhizophora mangle L. Hydroperiod treatments included no flooding (unflooded), intermittent flooding (intermittent), and permanent flooding (flooded). Plants in the intermittent treatment were measured under both flooded and drained states and compared separately. In the greenhouse study, plants of all species maintained different leaf areas in the contrasting hydroperiods during both years. Assimilation–light response curves indicated that the different hydroperiods had little effect on leaf gas exchange characteristics in either seedlings or saplings. However, short-term intermittent flooding for between 6 and 22 days caused a 20% reduction in maximum leaf-level carbon assimilation rate, a 51% lower light requirement to attain 50% of maximum assimilation, and a 38% higher demand from dark respiration. Although interspecific differences were evident for nearly all measured parameters in both years, there was little consistency in ranking of the interspecific responses. Species by hydroperiod interactions were significant only for sapling leaf area. In a field study, R. mangle saplings along the Shark River in the Everglades National Park either demonstrated no significant effect or slight enhancement of carbon assimilation and water-use efficiency while flooded. We obtained little evidence that contrasting hydroperiods affect leaf gas exchange characteristics of mangrove seedlings or saplings over long time intervals; however, intermittent flooding may cause short-term depressions in leaf gas exchange. The resilience of mangrove systems to flooding, as demonstrated in the permanently flooded treatments, will likely promote photosynthetic and morphological adjustment to slight hydroperiod shifts in many settings..
Leaf-out phenology of temperate woody plants: from trees to ecosystems.
Polgar, Caroline A; Primack, Richard B
2011-09-01
Leafing-out of woody plants begins the growing season in temperate forests and is one of the most important drivers of ecosystem processes. There is substantial variation in the timing of leaf-out, both within and among species, but the leaf development of almost all temperate tree and shrub species is highly sensitive to temperature. As a result, leaf-out times of temperate forests are valuable for observing the effects of climate change. Analysis of phenology data from around the world indicates that leaf-out is generally earlier in warmer years than in cooler years and that the onset of leaf-out has advanced in many locations. Changes in the timing of leaf-out will affect carbon sequestration, plant-animal interactions, and other essential ecosystem processes. The development of remote sensing methods has expanded the scope of leaf-out monitoring from the level of an individual plant or forest to an entire region. Meanwhile, historical data have informed modeling and experimental studies addressing questions about leaf-out timing. For most species, onset of leaf-out will continue to advance, although advancement may be slowed for some species because of unmet chilling requirements. More information is needed to reduce the uncertainty in predicting the timing of future spring onset. © 2011 The Authors. New Phytologist © 2011 New Phytologist Trust.
F-Box Protein FBX92 Affects Leaf Size in Arabidopsis thaliana
Baute, Joke; Polyn, Stefanie; De Block, Jolien; Blomme, Jonas; Van Lijsebettens, Mieke
2017-01-01
F-box proteins are part of one of the largest families of regulatory proteins that play important roles in protein degradation. In plants, F-box proteins are functionally very diverse, and only a small subset has been characterized in detail. Here, we identified a novel F-box protein FBX92 as a repressor of leaf growth in Arabidopsis. Overexpression of AtFBX92 resulted in plants with smaller leaves than the wild type, whereas plants with reduced levels of AtFBX92 showed, in contrast, increased leaf growth by stimulating cell proliferation. Detailed cellular analysis suggested that AtFBX92 specifically affects the rate of cell division during early leaf development. This is supported by the increased expression levels of several cell cycle genes in plants with reduced AtFBX92 levels. Surprisingly, overexpression of the maize homologous gene ZmFBX92 in maize had no effect on plant growth, whereas ectopic expression in Arabidopsis increased leaf growth. Expression of a truncated form of AtFBX92 showed that the contrasting effects of ZmFBX92 and AtFBX92 gain of function in Arabidopsis are due to the absence of the F-box-associated domain in the ZmFBX92 gene. Our work reveals an additional player in the complex network that determines leaf size and lays the foundation for identifying putative substrates. PMID:28340173
Phloem unloading in developing leaves of sugar beet
DOE Office of Scientific and Technical Information (OSTI.GOV)
Schmalstig, J.G.
1985-01-01
Physiological and transport data support a symplastic pathway for phloem unloading in developing leaves of sugar beet (Beta vulgaris L. Klein E, multigerm). The sulfhydryl inhibitor parachloromercuribenzene sulfonic acid (PCMBS) inhibited uptake of (/sup 14/C)-sucrose added to the free space of developing leaves, but did not affect import of (/sup 14/C)-sucrose during steady-state /sup 14/CO/sub 2/ labeling of a source leaf. The passively-transported xenobiotic sugar, (/sup 14/C)-L-glucose did not readily enter mesophyll cells when supplied through the cut end of the petiole of a sink leaf as determined by whole leaf autoradiography. In contrast, (/sup 14/C)-L-glucose translocated through the phloemmore » from a mature leaf, rapidly entered mesophyll cells, and was evenly distributed between mesophyll and veins. Autoradiographs of developing leaves following a pulse of /sup 14/CO/sub 2/ to a source leaf revealed rapid passage of phloem translocated into progressively higher order veins as the leaf developed. Entry into V order veins occurred during the last stage of import through the phloem. Import into developing leaves was inhibited by glyphosate (N-phosphomethylglycine), a herbicide which inhibits the aromatic amino acid pathway and hence protein synthesis. Glyphosate also stopped net starch accumulation in sprayed mature leaves, but did not affect export of carbon from treated leaves during the time period that import into developed leaves was inhibited.« less
Niinemets, Ülo; Portsmuth, Angelika; Tena, David; Tobias, Mari; Matesanz, Silvia; Valladares, Fernando
2007-01-01
Background Broad scaling relationships between leaf size and function do not take into account that leaves of different size may contain different fractions of support in petiole and mid-rib. Methods The fractions of leaf biomass in petiole, mid-rib and lamina, and the differences in chemistry and structure among mid-ribs, petioles and laminas were investigated in 122 species of contrasting leaf size, life form and climatic distribution to determine the extent to which differences in support modify whole-lamina and whole-leaf structural and chemical characteristics, and the extent to which size-dependent support investments are affected by plant life form and site climate. Key Results For the entire data set, leaf fresh mass varied over five orders of magnitude. The percentage of dry mass in mid-rib increased strongly with lamina size, reaching more than 40 % in the largest laminas. The whole-leaf percentage of mid-rib and petiole increased with leaf size, and the overall support investment was more than 60 % in the largest leaves. Fractional support investments were generally larger in herbaceous than in woody species and tended to be lower in Mediterranean than in cool temperate and tropical plants. Mid-ribs and petioles had lower N and C percentages, and lower dry to fresh mass ratio, but greater density (mass per unit volume) than laminas. N percentage of lamina without mid-rib was up to 40 % higher in the largest leaves than the total-lamina (lamina and mid-rib) N percentage, and up to 60 % higher than whole-leaf N percentage, while lamina density calculated without mid-rib was up to 80 % less than that with the mid-rib. For all leaf compartments, N percentage was negatively associated with density and dry to fresh mass ratio, while C percentage was positively linked to these characteristics, reflecting the overall inverse scaling between structural and physiological characteristics. However, the correlations between N and C percentages and structural characteristics differed among mid-ribs, petioles and laminas, implying that the mass-weighted average leaf N and C percentage, density, and dry to fresh mass ratio can have different functional values depending on the importance of within-leaf support investments. Conclusions These data demonstrate that variation in leaf size is associated with major changes in within-leaf support investments and in large modifications in integrated leaf chemical and structural characteristics. These size-dependent alterations can importantly affect general leaf structure vs. function scaling relationships. These data further demonstrate important life-form effects on and climatic differentiation in foliage support costs. PMID:17586597
Multifilamentary niobium tin magnet conductors
DOE Office of Scientific and Technical Information (OSTI.GOV)
Larbalestier, D.C.; Madsen, P.E.; Lee, J.A.
1975-03-01
Practical magnet conductors of multifilamentary Nb$sub 3$Sn have been produced. Evaluation of these bronze route conductors is described. Conductors studied range from a 1369 filament all-bronze matrix conductor to 5143 and approximately 42,000 filament conductors, containing internal high purity copper protected by diffusion barriers. Filament sizes vary from approximately 3 to 8 $mu$m diameter. The effect of heat treatment conditions on critical current and transition temperature is presented and it is shown that overall critical current densities greater than those available in niobium titanium can now be produced in multifilamentary Nb$sub 3$Sn magnet conductor.
Sesbania bispinosa, a new host of a begomovirus-betasatellite complex in Pakistan
USDA-ARS?s Scientific Manuscript database
Severe leaf curling, yellowing and vein thickening symptoms, typical of begomoviruses infection, were observed on Sesbania bispinosa grown in cotton leaf curl disease affected cotton field in Pakistan. A begomovirus and its associated betasatellite were amplified and sequenced from these plants. Com...
Potential impact of soil microbiomes on the leaf metabolome and on herbivore feeding behavior
USDA-ARS?s Scientific Manuscript database
: It is known that environmental factors can affect the biosynthesis of leaf metabolites. Similarly, specific pairwise plant-microbe interactions modulate specifically the plant’s metabolome by stimulating production of phytoalexins and other defense-related compounds. However, there is no informati...
The Liguleless narrow mutation affects proximal distal signaling and leaf growth
USDA-ARS?s Scientific Manuscript database
How cells acquire competence to differentiate according to position is an essential question in developmental biology. Maize leaves provide a unique opportunity to study positional information. In the developing leaf primordium, a line is drawn across a field of seemingly identical cells. Above the ...
Feng, Hui; Skinkis, Patricia A; Qian, Michael C
2017-01-01
The impacts of fruit zone leaf removal on volatile and anthocyanin compositions of Pinot noir wine were investigated over two growing seasons. Wine volatiles were analyzed by multiple techniques, including headspace solid phase microextraction-GC-MS (HS-SPME-GC-MS), headspace-GC-FID (HS-GC-FID) and stir bar sorptive extraction-GC-MS (SBSE-GC-MS). Fruit zone leaf removal affected the concentration of many grape-derived volatile compounds such as terpene alcohols and C13-norisoprenoids in wine, although the degree of impact depended on the vintage year and severity of leaf removal. Fruit zone leaf removal resulted in greater concentrations of linalool, α-terpineol and β-damascenone but had no impact on other terpene alcohols or β-ionone. Fruit zone leaf removal had no consistent impact on C6 alcohols, volatile phenols, lactones, fermentation-derived alcohols, acids, or most esters. Fruit zone leaf removal increased anthocyanins in final wine. Copyright © 2016 Elsevier Ltd. All rights reserved.
Hamaoka, Norimitsu; Yasui, Hideshi; Yamagata, Yoshiyuki; Inoue, Yoko; Furuya, Naruto; Araki, Takuya; Ueno, Osamu; Yoshimura, Atsushi
2017-12-01
High water use efficiency is essential to water-saving cropping. Morphological traits that affect photosynthetic water use efficiency are not well known. We examined whether leaf hairiness improves photosynthetic water use efficiency in rice. A chromosome segment introgression line (IL-hairy) of wild Oryza nivara (Acc. IRGC105715) with the genetic background of Oryza sativa cultivar 'IR24' had high leaf pubescence (hair). The leaf hairs developed along small vascular bundles. Linkage analysis in BC 5 F 2 and F 3 populations showed that the trait was governed by a single gene, designated BLANKET LEAF (BKL), on chromosome 6. IL-hairy plants had a warmer leaf surface in sunlight, probably due to increased boundary layer resistance. They had a lower transpiration rate under moderate and high light intensities, resulting in higher photosynthetic water use efficiency. Introgression of BKL on chromosome 6 from O. nivara improved photosynthetic water use efficiency in the genetic background of IR24.
Leaf Shape Responds to Temperature but Not CO2 in Acer rubrum
Royer, Dana L.
2012-01-01
The degree of leaf dissection and the presence of leaf teeth, along with tooth size and abundance, inversely correlate with mean annual temperature (MAT) across many plant communities. These relationships form the core of several methods for reconstructing MAT from fossils, yet the direct selection of temperature on tooth morphology has not been demonstrated experimentally. It is also not known if atmospheric CO2 concentration affects leaf shape, limiting confidence in ancient climate reconstructions because CO2 has varied widely on geologic timescales. Here I report the results of growing Acer rubrum (red maple) in growth cabinets at contrasting temperature and CO2 conditions. The CO2 treatment imparted no significant differences in leaf size and shape, while plants grown at cooler temperatures tended to have more teeth and more highly dissected leaves. These results provide direct evidence for the selection of temperature on leaf shape in one species, and support a key link in many leaf-climate methods. More broadly, these results increase confidence for using leaf shape in fossils to reconstruct paleoclimate. PMID:23152921
Plant structure predicts leaf litter capture in the tropical montane bromeliad Tillandsia turneri.
Ospina-Bautista, F; Estévez Varón, J V
2016-05-03
Leaves intercepted by bromeliads become an important energy and matter resource for invertebrate communities, bacteria, fungi, and the plant itself. The relationship between bromeliad structure, defined as its size and complexity, and accumulated leaf litter was studied in 55 bromeliads of Tillandsia turneri through multiple regression and the Akaike information criterion. Leaf litter accumulation in bromeliads was best explained by size and complexity variables such as plant cover, sheath length, and leaf number. In conclusion, plant structure determines the amount of litter that enters bromeliads, and changes in its structure could affect important processes within ecosystem functioning or species richness.
Buitrago, Sindy; Vanegast, Leidy; Ramos, Carolina
2015-09-01
Espeletia paipana is an endangered giant caulescent rosette endemic to (Asteraceae), Boyacdi-Colombia. Espelelia paipana is an endangered giant caulescent rosette endemic to Boyacá department. In order to establish whether a plant disease, characterized by the loss of leaf pubescence (PPF) and attributed to the pathogenic action of endophytic microorganisms, is the cause of the increasing mortality of population, the physiological performance of the species was evaluated with and without PPF. The incidence (% leaves affected in each of the 27 individuals in the current population) and severity (% leaf area affected on 135 leaves) of the PPF were monitored over a period of nine months, in three topographic zones of different heights. During four consecutive days in both dry and wet season, physiological parameters as chlorophyll content index (ICC), stomatal conductance (Gs) and leaf temperature (Tfol) were measured in healthy and affected leaves. The study was complemented with isolations and pathogenicity tests to identify the causal agent of the PPF. Overall, although the disease incidence in E. paipana was constant over time, the severity progressed surpassing 60 % of the leaf area. The increasing of severity in the upper side of leaves was attributed to the photo-oxidative effect of high radiation between 11:00 h and 14:00 h of the day. The reduction of functional leaf area because of the PPF, led to low Gs with serious implications for carbon fixation and thus limiting growth and biomass renewal. The effect of season in Tfol varied according to the topographic zone, while the ICC did not present a defined pattern with respect to the PPF; its low values could be associated with the production of other pigments. Finally, although it is not possible to ensure that Botrytis sp. is the causative of the loss of leaf pubescence, it is postulated as the most probably causal agent due to its high representativeness in the isolates and its infectious potential during the pathogenicity tests. In general, the reduction of healthy leaf biomass and decrease of physiological performance suggest that PPF affect negatively the survival of E. paipana, which means that the use of biological controllers could be a strategy to mitigate its effect on the population.
Effect of water availability on tolerance of leaf damage in tall morning glory, Ipomoea purpurea
NASA Astrophysics Data System (ADS)
Atala, Cristian; Gianoli, Ernesto
2009-03-01
Resource availability may limit plant tolerance of herbivory. To predict the effect of differential resource availability on plant tolerance, the limiting resource model (LRM) considers which resource limits plant fitness and which resource is mostly affected by herbivore damage. We tested the effect of experimental drought on tolerance of leaf damage in Ipomoea purpurea, which is naturally exposed to both leaf damage and summer drought. To seek mechanistic explanations, we also measured several morphological, allocation and gas exchange traits. In this case, LRM predicts that tolerance would be the same in both water treatments. Plants were assigned to a combination of two water treatments (control and low water) and two damage treatments (50% defoliation and undamaged). Plants showed tolerance of leaf damage, i.e., a similar number of fruits were produced by damaged and undamaged plants, only in control water. Whereas experimental drought affected all plant traits, leaf damage caused plants to show a greater leaf trichome density and reduced shoot biomass, but only in low water. It is suggested that the reduced fitness (number of fruits) of damaged plants in low water was mediated by the differential reduction of shoot biomass, because the number of fruits per shoot biomass was similar in damaged and undamaged plants. Alternative but less likely explanations include the opposing direction of functional responses to drought and defoliation, and resource costs of the damage-induced leaf trichome density. Our results somewhat challenge the LRM predictions, but further research including field experiments is needed to validate some of the preliminary conclusions drawn.
Bours, Ralph; van Zanten, Martijn; Pierik, Ronald; Bouwmeester, Harro; van der Krol, Alexander
2013-10-01
In the natural environment, days are generally warmer than the night, resulting in a positive day/night temperature difference (+DIF). Plants have adapted to these conditions, and when exposed to antiphase light and temperature cycles (cold photoperiod/warm night [-DIF]), most species exhibit reduced elongation growth. To study the physiological mechanism of how light and temperature cycles affect plant growth, we used infrared imaging to dissect growth dynamics under +DIF and -DIF in the model plant Arabidopsis (Arabidopsis thaliana). We found that -DIF altered leaf growth patterns, decreasing the amplitude and delaying the phase of leaf movement. Ethylene application restored leaf growth in -DIF conditions, and constitutive ethylene signaling mutants maintain robust leaf movement amplitudes under -DIF, indicating that ethylene signaling becomes limiting under these conditions. In response to -DIF, the phase of ethylene emission advanced 2 h, but total ethylene emission was not reduced. However, expression analysis on members of the 1-aminocyclopropane-1-carboxylic acid (ACC) synthase ethylene biosynthesis gene family showed that ACS2 activity is specifically suppressed in the petiole region under -DIF conditions. Indeed, petioles of plants under -DIF had reduced ACC content, and application of ACC to the petiole restored leaf growth patterns. Moreover, acs2 mutants displayed reduced leaf movement under +DIF, similar to wild-type plants under -DIF. In addition, we demonstrate that the photoreceptor PHYTOCHROME B restricts ethylene biosynthesis and constrains the -DIF-induced phase shift in rhythmic growth. Our findings provide a mechanistic insight into how fluctuating temperature cycles regulate plant growth.
Neufeld, Howard S; Chappelka, Arthur H; Somers, Greg L; Burkey, Kent O; Davison, Alan W; Finkelstein, Peter L
2006-03-01
The ability of the SPAD-502 chlorophyll meter to quantify chlorophyll amounts in ozone-affected leaves of cutleaf coneflower (Rudbeckia laciniata var. digitata) was assessed in this study. When relatively uninjured leaves were measured (percent leaf area affected by stipple less than 6%), SPAD meter readings were linearly related to total chlorophyll with an adjusted R (2) of 0.84. However, when leaves with foliar injury (characterized as a purple to brownish stipple on the upper leaf surface affecting more than 6% of the leaf area) were added, likelihood ratio tests showed that it was no longer possible to use the same equation to obtain chlorophyll estimations for both classes of leaves. Either an equation with a common slope or a common intercept was necessary. We suspect several factors are involved in altering the calibration of the SPAD meter for measuring chlorophyll amounts in visibly ozone-injured leaves, with the most likely being changes in either light absorption or scattering resulting from tissue necrosis.
Effect of solution and leaf surface polarity on droplet spread area and contact angle.
Nairn, Justin J; Forster, W Alison; van Leeuwen, Rebecca M
2016-03-01
How much an agrochemical spray droplet spreads on a leaf surface can significantly influence efficacy. This study investigates the effect solution polarity has on droplet spreading on leaf surfaces and whether the relative leaf surface polarity, as quantified using the wetting tension dielectric (WTD) technique, influences the final spread area. Contact angles and spread areas were measured using four probe solutions on 17 species. Probe solution polarity was found to affect the measured spread area and the contact angle of the droplets on non-hairy leaves. Leaf hairs skewed the spread area measurement, preventing investigation of the influence of surface polarity on hairy leaves. WTD-measured leaf surface polarity of non-hairy leaves was found to correlate strongly with the effect of solution polarity on spread area. For non-polar leaf surfaces the spread area decreases with increasing solution polarity, for neutral surfaces polarity has no effect on spread area and for polar leaf surfaces the spread area increases with increasing solution polarity. These results attest to the use of the WTD technique as a means to quantify leaf surface polarity. © 2015 Society of Chemical Industry. © 2015 Society of Chemical Industry.
Pérez, D; Galindo, L
2000-12-01
The year 1996 had a high pluviosity in Morrocoy National Park (western coastal zone, Venezuela) and low salinity in December 1996 affected the seagrass beds, dominated by Thalassia testudinum. Patches without T. testudinum were observed in localities of the park that used to have very dense populations of this plant. Sampling was done at Las Luisas to determine leaf productivity, turnover rate, short shoot density and relative biomass of plant sections, in order to compare with data obtained in September 1996, previous to the event. Green leaves, roots and rhizomes were the most affected parts. Mean green leaf biomass decreased in January and February 1997 to 5% of the plant's total biomass; mean root biomass decreased in March to 40% and mean rhi zome biomass decreased in February to 30%. The density of the active short shoots decreased to a minimum in February, but in April it reached a value similar to that of September 1996. The density of the inactive short shoots decreased to a minimum in March, and they dissappeared in April, matching the increasing density of the active short shoots between these two months. In February 1997, 56% of the inactive short shoots showed evidence of leaf initiation. In January 1997 the leaf productivity and turnover rate values (2.72 +/- 0.35 g/m2/d and 2.15% leaf DW/d) were similar to the annual mean previously determined from Las Luisas (2.35 +/- 0.72 g/m2/d and 1.96% leaf DW/d). Nevertheless, the values of productivity and turnover rate detected at Las Luisas in April 1997 (4.88 +/- 2.14 g/m2/d and 4.66% leaf DW/d) were higher than those values previously reported for this location. In response to the mortality episode, the leaf productivity and turnover rate of T. testudinum increased and the leaf initiation was activated in the inactive short shoots.
Interaction of Water Supply and N in Wheat 1
Morgan, Jack A.
1984-01-01
The purpose of this study was to investigate effects of N nutrition and water stress on stomatal behavior and CO2 exchange rate in wheat (Triticum aestivum L. cv Olaf). Wheat plants were grown hydroponically with high (100 milligrams per liter) and low (10 milligrams per liter) N. When plants were 38 days old, a 24-day water stress cycle was begun. A gradual increase in nutrient solution osmotic pressure from 0.03 to 1.95 mega Pascals was achieved by incremental additions of PEG-6,000. Plants in both N treatments adjusted osmotically, although leaf water potential was consistently lower and relative water content greater for low N plants in the first half of the stress cycle. Leaf conductance of high N plants appeared greater than that of low N plants at high water potentials, but showed greater sensitivity to reductions in water potential as indicated by earlier stomatal closure during the stress cycle. The apparent greater stomatal sensitivity of high N plants was associated with a curvilinear relationship between leaf conductance and leaf water potential; low N plants exhibited more of a threshold response. Trends in [CO2]INT throughout the stress cycle indicated nonstomatal effects of water stress on CO2 exchange rate were greater in high N plants. Although estimates of [CO2]INT were generally lower in high N plants, they were relatively insensitive to leaf water potential-induced changes in leaf conductance. In contrast, [CO2]INT of low N plants dropped concomitantly with leaf conductance at low leaf water potentials. Oxygen response of CO2 exchange rate for both treatments was affected less by reductions in water potential than was CO2 exchange rate at 2.5% O2, suggesting that CO2 assimilation capacity of the leaves was affected more by reductions in leaf water potential than were processes related to photorespiration. PMID:16663780
Because the rate of isoprene (2-methyl-1,3-butadiene) emission from plants is highly temperature-dependent, we investigated the natural fluctuations on leaf temperature and the effects of rapid temperature change on isoprene emission of red oak (Quercus rubra L.) leaves at the to...
Chloroplast Response to Low Leaf Water Potentials
Boyer, J. S.; Potter, J. R.
1973-01-01
The effect of decreases in turgor on chloroplast activity was studied by measuring the photochemical activity of intact sunflower (Helianthus annuus L. cv. Russian Mammoth) leaves having low water potentials. Leaf turgor, calculated from leaf water potential and osmotic potential, was found to be affected by the dilution of cell contents by water in the cell walls, when osmotic potentials were measured with a thermocouple psychrometer. After the correction of measurements of leaf osmotic potential, both the thermocouple psychrometer and a pressure chamber indicated that turgor became zero in sunflower leaves at leaf water potentials of −10 bars. Since most of the loss in photochemical activity occurred at water potentials below −10 bars, it was concluded that turgor had little effect on the photochemical activity of the leaves. PMID:16658486
Chloroplast response to low leaf water potentials: I. Role of turgor.
Boyer, J S; Potter, J R
1973-06-01
The effect of decreases in turgor on chloroplast activity was studied by measuring the photochemical activity of intact sunflower (Helianthus annuus L. cv. Russian Mammoth) leaves having low water potentials. Leaf turgor, calculated from leaf water potential and osmotic potential, was found to be affected by the dilution of cell contents by water in the cell walls, when osmotic potentials were measured with a thermocouple psychrometer. After the correction of measurements of leaf osmotic potential, both the thermocouple psychrometer and a pressure chamber indicated that turgor became zero in sunflower leaves at leaf water potentials of -10 bars. Since most of the loss in photochemical activity occurred at water potentials below -10 bars, it was concluded that turgor had little effect on the photochemical activity of the leaves.
Coming of leaf age: control of growth by hydraulics and metabolics during leaf ontogeny.
Pantin, Florent; Simonneau, Thierry; Muller, Bertrand
2012-10-01
Leaf growth is the central process facilitating energy capture and plant performance. This is also one of the most sensitive processes to a wide range of abiotic stresses. Because hydraulics and metabolics are two major determinants of expansive growth (volumetric increase) and structural growth (dry matter increase), we review the interaction nodes between water and carbon. We detail the crosstalks between water and carbon transports, including the dual role of stomata and aquaporins in regulating water and carbon fluxes, the coupling between phloem and xylem, the interactions between leaf water relations and photosynthetic capacity, the links between Lockhart's hydromechanical model and carbon metabolism, and the central regulatory role of abscisic acid. Then, we argue that during leaf ontogeny, these interactions change dramatically because of uncoupled modifications between several anatomical and physiological features of the leaf. We conclude that the control of leaf growth switches from a metabolic to a hydromechanical limitation during the course of leaf ontogeny. Finally, we illustrate how taking leaf ontogeny into account provides insights into the mechanisms underlying leaf growth responses to abiotic stresses that affect water and carbon relations, such as elevated CO2, low light, high temperature and drought. © 2012 INRA. New Phytologist © 2012 New Phytologist Trust.
Lotus leaf extract and L-carnitine influence different processes during the adipocyte life cycle
2010-01-01
Background The cellular and molecular mechanisms of adipose tissue biology have been studied extensively over the last two decades. Adipose tissue growth involves both an increase in fat cell size and the formation of mature adipocytes from precursor cells. To investigate how natural substances influence these two processes, we examined the effects of lotus leaf extract (Nelumbo nucifera-extract solution obtained from Silab, France) and L-carnitine on human preadipocytes and adipocytes. Methods For our in vitro studies, we used a lotus leaf extract solution alone or in combination with L-carnitine. Utilizing cultured human preadipocytes, we investigated lotus leaf extract solution-induced inhibition of triglyceride incorporation during adipogenesis and possible effects on cell viability. Studies on human adipocytes were performed aiming to elucidate the efficacy of lotus leaf extract solution to stimulate lipolytic activity. To further characterize lotus leaf extract solution-mediated effects, we determined the expression of the transcription factor adipocyte determination and differentiation factor 1 (ADD1/SREBP-1c) on the RNA- and protein level utilizing qRT-PCR and immunofluorescence analysis. Additionally, the effect of L-carnitine on beta-oxidation was analyzed using human preadipocytes and mature adipocytes. Finally, we investigated additive effects of a combination of lotus leaf extract solution and L-carnitine on triglyceride accumulation during preadipocyte/adipocyte differentiation. Results Our data showed that incubation of preadipocytes with lotus leaf extract solution significantly decreased triglyceride accumulation during adipogenesis without affecting cell viability. Compared to controls, adipocytes incubated with lotus leaf extract solution exhibited a significant increase in lipolysis-activity. Moreover, cell populations cultivated in the presence of lotus leaf extract solution showed a decrease in adipocyte differentiation capacity as indicated by a decrease in the ADD1/SREBP-1c signal. Importantly, our results demonstrated that a combination of lotus leaf extract solution and L-carnitine reduced triglyceride accumulation to a greater extent compared to incubation with either substance alone. Conclusions Overall, our data demonstrate that a combination of lotus leaf extract and L-carnitine reduced triglyceride accumulation in human (pre)adipocytes by affecting different processes during the adipocyte life cycle. For this reason, this combination might represent a treatment option for obesity-related diseases. PMID:20687953
Macintosh, Alison A; Pinhasi, Ron; Stock, Jay T
2014-01-01
Humeral morphology has been shown to reflect, in part, habitual manipulative behaviors in humans. Among Central European agricultural populations, long-term social change, increasing task specialization, and technological innovation all had the potential to impact patterns of habitual activity and upper limb asymmetry. However, systematic temporal change in the skeletal morphology of agricultural populations in this region has not been well-characterized. This study investigates diachronic patterns in humeral biomechanical properties and lengths among 174 adult Central European agriculturalists through the first ∼ 5400 years of farming in the region. Greater asymmetry in biomechanical properties was expected to accompany the introduction of metallurgy, particularly in males, while upper limb loading patterns were expected to be more similar between the Bronze and Iron Ages. Results revealed a divergence in the lateralization of upper limb biomechanical properties by sex between the Early/Middle Neolithic and Early/Middle Bronze Age. Neolithic females had significantly more variable properties than males in both humeri, while Bronze Age female properties became homogeneous and very symmetrical relative to the right-biased lateralization of contemporaneous males. The Bronze Age to Iron Age transition was associated with morphological change among females, with a significant increase in right-biased asymmetry and a concomitant reduction in sexual dimorphism. Relative to biomechanical properties, humeral length variation and asymmetry were low though some significant sexual dimorphism and temporal change was found. It was among females that the lateralization of humeral biomechanical properties, and variation within them, changed most profoundly through time. This suggests that the introduction of the ard and plow, metallurgical innovation, task specialization, and socioeconomic change through ∼ 5400 years of agriculture impacted upper limb loading in Central European women to a greater extent than men.
Barash, Alon; Eisenberg-Degen, Davida; Grosman, Leore; Oron, Maya; Berger, Uri
2017-01-01
The Intermediate Bronze Age (IB) in the Southern Levant (ca. 2350–2000 BCE) is known as the “Dark Ages,” following the collapse of Early Bronze urban society and predating the establishment of the Middle Bronze cities. The absence of significant settlements and monumental building has led to the reconstruction of IB social organization as that of nomadic, tribal society inhabiting rural villages with no central governmental system. Excavation in the Shamir Dolmen Field (comprising over 400 dolmens) on the western foothills of the Golan Heights was carried out following the discovery of rock art engravings on the ceiling of the central chamber inside one of the largest dolmens ever recorded in the Levant. Excavation of this multi-chambered dolmen, covered by a basalt capstone weighing some 50 tons, revealed a secondary multi-burial (of both adults and children) rarely described in a dolmen context in the Golan. Engraved into the rock ceiling above the multi-burial is a panel of 14 forms composed of a vertical line and downturned arc motif. 3D-scanning by structured-light technology was used to sharpen the forms and revealed the technique employed to create them. Building of the Shamir dolmens required a tremendous amount of labor, architectural mastery, and complex socio-economic organization well beyond the capacity of small, rural nomadic groups. The monumental megalithic burial of the Shamir dolmens indicates a hierarchical, complex, non-urban governmental system. This new evidence supports a growing body of recent criticism stemming from new discoveries and approaches that calls for rethinking our views of the Levantine IB “Dark Ages.” PMID:28253312
Haber, Marc; Doumet-Serhal, Claude; Scheib, Christiana; Xue, Yali; Danecek, Petr; Mezzavilla, Massimo; Youhanna, Sonia; Martiniano, Rui; Prado-Martinez, Javier; Szpak, Michał; Matisoo-Smith, Elizabeth; Schutkowski, Holger; Mikulski, Richard; Zalloua, Pierre; Kivisild, Toomas; Tyler-Smith, Chris
2017-08-03
The Canaanites inhabited the Levant region during the Bronze Age and established a culture that became influential in the Near East and beyond. However, the Canaanites, unlike most other ancient Near Easterners of this period, left few surviving textual records and thus their origin and relationship to ancient and present-day populations remain unclear. In this study, we sequenced five whole genomes from ∼3,700-year-old individuals from the city of Sidon, a major Canaanite city-state on the Eastern Mediterranean coast. We also sequenced the genomes of 99 individuals from present-day Lebanon to catalog modern Levantine genetic diversity. We find that a Bronze Age Canaanite-related ancestry was widespread in the region, shared among urban populations inhabiting the coast (Sidon) and inland populations (Jordan) who likely lived in farming societies or were pastoral nomads. This Canaanite-related ancestry derived from mixture between local Neolithic populations and eastern migrants genetically related to Chalcolithic Iranians. We estimate, using linkage-disequilibrium decay patterns, that admixture occurred 6,600-3,550 years ago, coinciding with recorded massive population movements in Mesopotamia during the mid-Holocene. We show that present-day Lebanese derive most of their ancestry from a Canaanite-related population, which therefore implies substantial genetic continuity in the Levant since at least the Bronze Age. In addition, we find Eurasian ancestry in the Lebanese not present in Bronze Age or earlier Levantines. We estimate that this Eurasian ancestry arrived in the Levant around 3,750-2,170 years ago during a period of successive conquests by distant populations. Copyright © 2017 The Author(s). Published by Elsevier Inc. All rights reserved.
Sharon, Gonen; Barash, Alon; Eisenberg-Degen, Davida; Grosman, Leore; Oron, Maya; Berger, Uri
2017-01-01
The Intermediate Bronze Age (IB) in the Southern Levant (ca. 2350-2000 BCE) is known as the "Dark Ages," following the collapse of Early Bronze urban society and predating the establishment of the Middle Bronze cities. The absence of significant settlements and monumental building has led to the reconstruction of IB social organization as that of nomadic, tribal society inhabiting rural villages with no central governmental system. Excavation in the Shamir Dolmen Field (comprising over 400 dolmens) on the western foothills of the Golan Heights was carried out following the discovery of rock art engravings on the ceiling of the central chamber inside one of the largest dolmens ever recorded in the Levant. Excavation of this multi-chambered dolmen, covered by a basalt capstone weighing some 50 tons, revealed a secondary multi-burial (of both adults and children) rarely described in a dolmen context in the Golan. Engraved into the rock ceiling above the multi-burial is a panel of 14 forms composed of a vertical line and downturned arc motif. 3D-scanning by structured-light technology was used to sharpen the forms and revealed the technique employed to create them. Building of the Shamir dolmens required a tremendous amount of labor, architectural mastery, and complex socio-economic organization well beyond the capacity of small, rural nomadic groups. The monumental megalithic burial of the Shamir dolmens indicates a hierarchical, complex, non-urban governmental system. This new evidence supports a growing body of recent criticism stemming from new discoveries and approaches that calls for rethinking our views of the Levantine IB "Dark Ages."
A Novel Cathode Material for Cathodic Dehalogenation of 1,1-Dibromo Cyclopropane Derivatives.
Gütz, Christoph; Selt, Maximilian; Bänziger, Markus; Bucher, Christoph; Römelt, Christina; Hecken, Nadine; Gallou, Fabrice; Galvão, Tomás R; Waldvogel, Siegfried R
2015-09-28
Leaded bronze turned out to be an excellent cathode material for the dehalogenation reaction of cyclopropanes without affecting the strained molecular entity. With this particular alloy, beneficial properties of lead cathodes are conserved, whereas the corrosion of cathode is efficiently suppressed. The solvent in the electrolyte determines whether a complete debromination reaction is achieved or if the process can be selectively stopped at the monobromo cyclopropane intermediate. The electroorganic conversion tolerates a variety of functional groups and can be conducted at rather complex substrates like cyclosporine A. This approach allows the sustainable preparation of cyclopropane derivatives. © 2015 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim.
NASA Astrophysics Data System (ADS)
Ding, J.; Johnson, E. A.; Martin, Y. E.
2017-12-01
Leaf is the basic production unit of plants. Water is the most critical resource of plants. Its availability controls primary productivity of plants by affecting leaf carbon budget. To avoid the damage of cavitation from lowering vein water potential t caused by evapotranspiration, the leaf must increase the stomatal resistance to reduce evapotranspiration rate. This comes at the cost of reduced carbon fixing rate as increasing stoma resistance meanwhile slows carbon intake rate. Studies suggest that stoma will operate at an optimal resistance to maximize the carbon gain with respect to water. Different plant species have different leaf shapes, a genetically determined trait. Further, on the same plant leaf size can vary many times in size that is related to soil moisture, an indicator of water availability. According to metabolic scaling theory, increasing leaf size will increase total xylem resistance of vein, which may also constrain leaf carbon budget. We present a Constrained Maximization Model of leaf (leaf CMM) that incorporates metabolic theory into the coupling of evapotranspiration and carbon fixation to examine how leaf size, stoma resistance and maximum net leaf primary productivity change with petiole xylem water potential. The model connects vein network structure to leaf shape and use the difference between petiole xylem water potential and the critical minor vein cavitation forming water potential as the budget. The CMM shows that both maximum net leaf primary production and optimal leaf size increase with petiole xylem water potential while optimal stoma resistance decreases. Narrow leaf has overall lower optimal leaf size and maximum net leaf carbon gain and higher optimal stoma resistance than those of broad leaf. This is because with small width to length ratio, total xylem resistance increases faster with leaf size. Total xylem resistance of narrow leaf increases faster with leaf size causing higher average and marginal cost of xylem water potential with respect to net leaf carbon gain. With same leaf area, total xylem resistance of narrow leaf is higher than broad leaf. Given same stoma resistance and petiole water potential, narrow leaf will lose more xylem water potential than broad leaf. Consequently, narrow leaf has smaller size and higher stoma resistance at optimum.
Genetic dissection and validation of candidate genes for flag leaf size in rice (Oryza sativa L.).
Tang, Xinxin; Gong, Rong; Sun, Wenqiang; Zhang, Chaopu; Yu, Sibin
2018-04-01
Two major loci with functional candidate genes were identified and validated affecting flag leaf size, which offer desirable genes to improve leaf architecture and photosynthetic capacity in rice. Leaf size is a major determinant of plant architecture and yield potential in crops. However, the genetic and molecular mechanisms regulating leaf size remain largely elusive. In this study, quantitative trait loci (QTLs) for flag leaf length and flag leaf width in rice were detected with high-density single nucleotide polymorphism genotyping of a chromosomal segment substitution line (CSSL) population, in which each line carries one or a few chromosomal segments from the japonica cultivar Nipponbare in a common background of the indica variety Zhenshan 97. In total, 14 QTLs for flag leaf length and nine QTLs for flag leaf width were identified in the CSSL population. Among them, qFW4-2 for flag leaf width was mapped to a 37-kb interval, with the most likely candidate gene being the previously characterized NAL1. Another major QTL for both flag leaf width and length was delimited by substitution mapping to a small region of 13.5 kb that contains a single gene, Ghd7.1. Mutants of Ghd7.1 generated using CRISPR/CAS9 approach showed reduced leaf size. Allelic variation analyses also validated Ghd7.1 as a functional candidate gene for leaf size, photosynthetic capacity and other yield-related traits. These results provide useful genetic information for the improvement of leaf size and yield in rice breeding programs.
Liu, Yang; Qian, Chenyun; Ding, Sihui; Shang, Xulan; Yang, Wanxia; Fang, Shengzuo
2016-12-01
As a highly valued and multiple function tree species, Cyclocarya paliurus is planted and managed for timber production and medical use. However, limited information is available on its genotype selection and cultivation for growth and phytochemicals. Responses of growth and secondary metabolites to light regimes and genotypes are useful information to determine suitable habitat conditions for the cultivation of medicinal plants. Both light regime and provenance significantly affected the leaf characteristics, leaf flavonoid contents, biomass production and flavonoid accumulation per plant. Leaf thickness, length of palisade cells and chlorophyll a/b decreased significantly under shading conditions, while leaf areas and total chlorophyll content increased obviously. In the full light condition, leaf flavonoid contents showed a bimodal temporal variation pattern with the maximum observed in August and the second peak in October, while shading treatment not only reduced the leaf content of flavonoids but also delayed the peak appearing of the flavonoid contents in the leaves of C. paliurus. Strong correlations were found between leaf thickness, palisade length, monthly light intensity and measured flavonoid contents in the leaves of C. paliurus. Muchuan provenance with full light achieved the highest leaf biomass and flavonoid accumulation per plant. Cyclocarya paliurus genotypes show diverse responses to different light regimes in leaf characteristics, biomass production and flavonoid accumulation, highlighting the opportunity for extensive selection in the leaf flavonoid production.
Sun, Jinwei; Wu, Jiabing; Guan, Dexin; Yao, Fuqi; Yuan, Fenghui; Wang, Anzhi; Jin, Changjie
2014-01-01
Leaf respiration is an important component of carbon exchange in terrestrial ecosystems, and estimates of leaf respiration directly affect the accuracy of ecosystem carbon budgets. Leaf respiration is inhibited by light; therefore, gross primary production (GPP) will be overestimated if the reduction in leaf respiration by light is ignored. However, few studies have quantified GPP overestimation with respect to the degree of light inhibition in forest ecosystems. To determine the effect of light inhibition of leaf respiration on GPP estimation, we assessed the variation in leaf respiration of seedlings of the dominant tree species in an old mixed temperate forest with different photosynthetically active radiation levels using the Laisk method. Canopy respiration was estimated by combining the effect of light inhibition on leaf respiration of these species with within-canopy radiation. Leaf respiration decreased exponentially with an increase in light intensity. Canopy respiration and GPP were overestimated by approximately 20.4% and 4.6%, respectively, when leaf respiration reduction in light was ignored compared with the values obtained when light inhibition of leaf respiration was considered. This study indicates that accurate estimates of daytime ecosystem respiration are needed for the accurate evaluation of carbon budgets in temperate forests. In addition, this study provides a valuable approach to accurately estimate GPP by considering leaf respiration reduction in light in other ecosystems. PMID:25419844
High resolution estimates of the corrosion risk for cultural heritage in Italy.
De Marco, Alessandra; Screpanti, Augusto; Mircea, Mihaela; Piersanti, Antonio; Proietti, Chiara; Fornasier, M Francesca
2017-07-01
Air pollution plays a pivotal role in the deterioration of many materials used in buildings and cultural monuments causing an inestimable damage. This study aims to estimate the impacts of air pollution (SO 2 , HNO 3 , O 3 , PM 10 ) and meteorological conditions (temperature, precipitation, relative humidity) on limestone, copper and bronze based on high resolution air quality data-base produced with AMS-MINNI modelling system over the Italian territory over the time period 2003-2010. A comparison between high resolution data (AMS-MINNI grid, 4 × 4 km) and low resolution data (EMEP grid, 50 × 50 km) has been performed. Our results pointed out that the corrosion levels for limestone, copper and bronze are decreased in Italy from 2003 to 2010 in relation to decrease of pollutant concentrations. However, some problem related to air pollution persists especially in Northern and Southern Italy. In particular, PM 10 and HNO 3 are considered the main responsible for limestone corrosion. Moreover, the high resolution data (AMS-MINNI) allowed the identification of risk areas that are not visible with the low resolution data (EMEP modelling system) in all considered years and, especially, in the limestone case. Consequently, high resolution air quality simulations are suitable to provide concrete benefits in providing information for national effective policy against corrosion risk for cultural heritage, also in the context of climate changes that are affecting strongly Mediterranean basin. Copyright © 2017 Elsevier Ltd. All rights reserved.
Primack, Richard B; Laube, Julia; Gallinat, Amanda S; Menzel, Annette
2015-11-01
Climate change is advancing the leaf-out times of many plant species and mostly extending the growing season in temperate ecosystems. Laboratory experiments using twig cuttings from woody plant species present an affordable, easily replicated approach to investigate the relative importance of factors such as winter chilling, photoperiod, spring warming and frost tolerance on the leafing-out times of plant communities. This Viewpoint article demonstrates how the results of these experiments deepen our understanding beyond what is possible via analyses of remote sensing and field observation data, and can be used to improve climate change forecasts of shifts in phenology, ecosystem processes and ecological interactions. The twig method involves cutting dormant twigs from trees, shrubs and vines on a single date or at intervals over the course of the winter and early spring, placing them in containers of water in controlled environments, and regularly recording leaf-out, flowering or other phenomena. Prior to or following leaf-out or flowering, twigs may be assigned to treatment groups for experiments involving temperature, photoperiod, frost, humidity and more. Recent studies using these methods have shown that winter chilling requirements and spring warming strongly affect leaf-out and flowering times of temperate trees and shrubs, whereas photoperiod requirements are less important than previously thought for most species. Invasive plant species have weaker winter chilling requirements than native species in temperate ecosystems, and species that leaf-out early in the season have greater frost tolerance than later leafing species. This methodology could be extended to investigate additional drivers of leaf-out phenology, leaf senescence in the autumn, and other phenomena, and could be a useful tool for education and outreach. Additional ecosystems, such as boreal, southern hemisphere and sub-tropical forests, could also be investigated using dormant twigs to determine the drivers of leaf-out times and how these ecosystems will be affected by climate change. © The Author 2015. Published by Oxford University Press on behalf of the Annals of Botany Company. All rights reserved. For Permissions, please email: journals.permissions@oup.com.
Organ-specific proteomics of soybean seedlings under flooding and drought stresses.
Wang, Xin; Khodadadi, Ehsaneh; Fakheri, Baratali; Komatsu, Setsuko
2017-06-06
Organ-specific analyses enrich the understanding of plant growth and development under abiotic stresses. To elucidate the cellular responses in soybean seedlings exposed to flooding and drought stresses, organ-specific analysis was performed using a gel-free/label-free proteomic technique. Physiological analysis indicated that enzyme activities of alcohol dehydrogenase and delta-1-pyrroline-5-carboxylate synthase were markedly increased in leaf and root of plants treated with 6days of flooding and drought stresses, respectively. Proteins related to photosynthesis, RNA, DNA, signaling, and the tricarboxylic acid cycle were predominately affected in leaf, hypocotyl, and root in response to flooding and drought. Notably, the tricarboxylic acid cycle was suppressed in leaf and root under both stresses. Moreover, 17 proteins, including beta-glucosidase 31 and beta-amylase 5, were identified in soybean seedlings under both stresses. The protein abundances of beta-glucosidase 31 and beta-amylase 5 were increased in leaf and root under both stresses. Additionally, the gene expression of beta-amylase 5 was upregulated in leaf exposed to the flooding and drought, and the expression level was highly correlated with the protein abundance. These results suggest that beta-amylase 5 may be involved in carbohydrate mobilization to provide energy to the leaf of soybean seedlings exposed to flooding and drought. This study examined the effects of flooding and drought on soybean seedlings in different organs using a gel-free/label-free proteomic approach. Physiological responses indicated that enzyme activities of alcohol dehydrogenase and delta-1-pyrroline-5-carboxylate synthase were increased in leaf and root of soybean seedlings exposed to flooding and drought for 6days. Functional analysis of acquired protein profiles exhibited that proteins related to photosynthesis, RNA, DNA, signaling, and the tricarboxylic acid cycle were predominated affected in leaf, hypocotyl, and root under both stresses. Moreover, the tricarboxylic acid cycle was suppressed in leaf and root of stressed soybean seedlings. Additionally, increased protein abundance of beta-amylase 5 was consistent with upregulated gene expression in the leaf under both stresses, suggesting that carbohydrate metabolism might be governed in response to flooding and drought of soybean seedlings. Copyright © 2017 Elsevier B.V. All rights reserved.
Comparing intra- and inter-specific effects on litter decomposition in an old-field ecosystem
DOE Office of Scientific and Technical Information (OSTI.GOV)
Crutsinger, Greg; Sanders, Dr. Nathan James; Classen, Aimee T
2009-09-01
Plant species can differ in the quantity and quality of leaf litter they produce, and many studies have examined whether plant species diversity affects leaf-litter decomposition and nutrient release. A growing number of studies have indicated that intra-specific variation within plant species can also affect key ecosystem processes. However, the relative importance of intra- versus inter-specific variation for the functioning of ecosystems remains poorly known. Here, we investigate the effects of intra-specific variation in a dominant old-field plant species, tall goldenrod (Solidago altissima), and inter-specific variation among goldenrod species on litter quality, decomposition, and nitrogen (N) release. We found thatmore » the nutrient concentration of leaf litter varied among genotypes, which translated into 50% difference in decomposition rates. Variation among other goldenrod species in decomposition rate was more than twice that of genetic variation within S. altissima. Furthermore, by manipulating litterbags to contain 1, 3, 6, or 9 genotypes, we found that S. altissima genotype identity had much stronger effects than did genotypic diversity on leaf-litter quality, decomposition, and N release. Taken together, these results suggest that the order of ecological importance for controlling leaf-litter decomposition and N release dynamics is plant species identitygenotype identity>genotypic diversity.« less
BIG LEAF is a regulator of organ size and adventitious root formation in poplar
DOE Office of Scientific and Technical Information (OSTI.GOV)
Yordanov, Yordan S.; Ma, Cathleen; Yordanova, Elena
Here we report the discovery through activation tagging and subsequent characterization of the BIG LEAF (BL) gene from poplar. In poplar, BL regulates leaf size via positively affecting cell proliferation. Up and downregulation of the gene led to increased and decreased leaf size, respectively, and these phenotypes corresponded to increased and decreased cell numbers. BL function encompasses the early stages of leaf development as native BL expression was specific to the shoot apical meristem and leaf primordia and was absent from the later stages of leaf development and other organs. Consistently, BL downregulation reduced leaf size at the earliest stagesmore » of leaf development. Ectopic expression in mature leaves resulted in continued growth most probably via sustained cell proliferation and thus the increased leaf size. In contrast to the positive effect on leaf growth, ectopic BL expression in stems interfered with and significantly reduced stem thickening, suggesting that BL is a highly specific activator of growth. Additionally, stem cuttings from BL overexpressing plants developed roots, whereas the wild type was difficult to root, demonstrating that BL is a positive regulator of adventitious rooting. Large transcriptomic changes in plants that overexpressed BL indicated that BL may have a broad integrative role, encompassing many genes linked to organ growth. Here, we conclude that BL plays a fundamental role in control of leaf size and thus may be a useful tool for modifying plant biomass productivity and adventitious rooting.« less
BIG LEAF is a regulator of organ size and adventitious root formation in poplar
Yordanov, Yordan S.; Ma, Cathleen; Yordanova, Elena; Meilan, Richard; Strauss, Steven H.; Busov, Victor B.
2017-01-01
Here we report the discovery through activation tagging and subsequent characterization of the BIG LEAF (BL) gene from poplar. In poplar, BL regulates leaf size via positively affecting cell proliferation. Up and downregulation of the gene led to increased and decreased leaf size, respectively, and these phenotypes corresponded to increased and decreased cell numbers. BL function encompasses the early stages of leaf development as native BL expression was specific to the shoot apical meristem and leaf primordia and was absent from the later stages of leaf development and other organs. Consistently, BL downregulation reduced leaf size at the earliest stages of leaf development. Ectopic expression in mature leaves resulted in continued growth most probably via sustained cell proliferation and thus the increased leaf size. In contrast to the positive effect on leaf growth, ectopic BL expression in stems interfered with and significantly reduced stem thickening, suggesting that BL is a highly specific activator of growth. In addition, stem cuttings from BL overexpressing plants developed roots, whereas the wild type was difficult to root, demonstrating that BL is a positive regulator of adventitious rooting. Large transcriptomic changes in plants that overexpressed BL indicated that BL may have a broad integrative role, encompassing many genes linked to organ growth. We conclude that BL plays a fundamental role in control of leaf size and thus may be a useful tool for modifying plant biomass productivity and adventitious rooting. PMID:28686626
BIG LEAF is a regulator of organ size and adventitious root formation in poplar
Yordanov, Yordan S.; Ma, Cathleen; Yordanova, Elena; ...
2017-07-07
Here we report the discovery through activation tagging and subsequent characterization of the BIG LEAF (BL) gene from poplar. In poplar, BL regulates leaf size via positively affecting cell proliferation. Up and downregulation of the gene led to increased and decreased leaf size, respectively, and these phenotypes corresponded to increased and decreased cell numbers. BL function encompasses the early stages of leaf development as native BL expression was specific to the shoot apical meristem and leaf primordia and was absent from the later stages of leaf development and other organs. Consistently, BL downregulation reduced leaf size at the earliest stagesmore » of leaf development. Ectopic expression in mature leaves resulted in continued growth most probably via sustained cell proliferation and thus the increased leaf size. In contrast to the positive effect on leaf growth, ectopic BL expression in stems interfered with and significantly reduced stem thickening, suggesting that BL is a highly specific activator of growth. Additionally, stem cuttings from BL overexpressing plants developed roots, whereas the wild type was difficult to root, demonstrating that BL is a positive regulator of adventitious rooting. Large transcriptomic changes in plants that overexpressed BL indicated that BL may have a broad integrative role, encompassing many genes linked to organ growth. Here, we conclude that BL plays a fundamental role in control of leaf size and thus may be a useful tool for modifying plant biomass productivity and adventitious rooting.« less
BIG LEAF is a regulator of organ size and adventitious root formation in poplar.
Yordanov, Yordan S; Ma, Cathleen; Yordanova, Elena; Meilan, Richard; Strauss, Steven H; Busov, Victor B
2017-01-01
Here we report the discovery through activation tagging and subsequent characterization of the BIG LEAF (BL) gene from poplar. In poplar, BL regulates leaf size via positively affecting cell proliferation. Up and downregulation of the gene led to increased and decreased leaf size, respectively, and these phenotypes corresponded to increased and decreased cell numbers. BL function encompasses the early stages of leaf development as native BL expression was specific to the shoot apical meristem and leaf primordia and was absent from the later stages of leaf development and other organs. Consistently, BL downregulation reduced leaf size at the earliest stages of leaf development. Ectopic expression in mature leaves resulted in continued growth most probably via sustained cell proliferation and thus the increased leaf size. In contrast to the positive effect on leaf growth, ectopic BL expression in stems interfered with and significantly reduced stem thickening, suggesting that BL is a highly specific activator of growth. In addition, stem cuttings from BL overexpressing plants developed roots, whereas the wild type was difficult to root, demonstrating that BL is a positive regulator of adventitious rooting. Large transcriptomic changes in plants that overexpressed BL indicated that BL may have a broad integrative role, encompassing many genes linked to organ growth. We conclude that BL plays a fundamental role in control of leaf size and thus may be a useful tool for modifying plant biomass productivity and adventitious rooting.
Leaf habit and woodiness regulate different leaf economy traits at a given nutrient supply.
Ordoñez, Jenny C; van Bodegom, Peter M; Witte, Jan-Philip M; Bartholomeus, Ruud P; van Dobben, Han F; Aerts, Rien
2010-11-01
The large variation in the relationships between environmental factors and plant traits observed in natural communities exemplifies the alternative solutions that plants have developed in response to the same environmental limitations. Qualitative attributes, such as growth form, woodiness, and leaf habit can be used to approximate these alternative solutions. Here, we quantified the extent to which these attributes affect leaf trait values at a given resource supply level, using measured plant traits from 105 different species (254 observations) distributed across 50 sites in mesic to wet plant communities in The Netherlands. For each site, soil total N, soil total P, and water supply estimates were obtained by field measurements and modeling. Effects of growth forms, woodiness, and leaf habit on relations between leaf traits (SLA, specific leaf area; LNC, leaf nitrogen concentration; and LPC, leaf phosphorus concentration) vs. nutrient and water supply were quantified using maximum-likelihood methods and Bonferroni post hoc tests. The qualitative attributes explained 8-23% of the variance within sites in leaf traits vs. soil fertility relationships, and therefore they can potentially be used to make better predictions of global patterns of leaf traits in relation to nutrient supply. However, at a given soil fertility, the strength of the effect of each qualitative attribute was not the same for all leaf traits. These differences may imply a differential regulation of the leaf economy traits at a given nutrient supply, in which SLA and LPC seem to be regulated in accordance to changes in plant size and architecture while LNC seems to be primarily regulated at the leaf level by factors related to leaf longevity.
F-Box Protein FBX92 Affects Leaf Size in Arabidopsis thaliana.
Baute, Joke; Polyn, Stefanie; De Block, Jolien; Blomme, Jonas; Van Lijsebettens, Mieke; Inzé, Dirk
2017-05-01
F-box proteins are part of one of the largest families of regulatory proteins that play important roles in protein degradation. In plants, F-box proteins are functionally very diverse, and only a small subset has been characterized in detail. Here, we identified a novel F-box protein FBX92 as a repressor of leaf growth in Arabidopsis. Overexpression of AtFBX92 resulted in plants with smaller leaves than the wild type, whereas plants with reduced levels of AtFBX92 showed, in contrast, increased leaf growth by stimulating cell proliferation. Detailed cellular analysis suggested that AtFBX92 specifically affects the rate of cell division during early leaf development. This is supported by the increased expression levels of several cell cycle genes in plants with reduced AtFBX92 levels. Surprisingly, overexpression of the maize homologous gene ZmFBX92 in maize had no effect on plant growth, whereas ectopic expression in Arabidopsis increased leaf growth. Expression of a truncated form of AtFBX92 showed that the contrasting effects of ZmFBX92 and AtFBX92 gain of function in Arabidopsis are due to the absence of the F-box-associated domain in the ZmFBX92 gene. Our work reveals an additional player in the complex network that determines leaf size and lays the foundation for identifying putative substrates. © The Author 2017. Published by Oxford University Press on behalf of Japanese Society of Plant Physiologists.
Li, Xiaoqiong; Guo, Wenfeng; Siemann, Evan; Wen, Yuanguang; Huang, Wei; Ding, Jianqing
2016-12-01
Spatially separated aboveground (AG) and belowground (BG) herbivores are closely linked through shared host plants, and both patterns of AG-BG interactions and plant responses may vary among plant genotypes. We subjected invasive (USA) and native (China) genotypes of tallow tree (Triadica sebifera) to herbivory by the AG specialist leaf-rolling weevil Heterapoderopsis bicallosicollis and/or the root-feeding larvae of flea beetle Bikasha collaris. We measured leaf damage and leaves rolled by weevils, quantified beetle survival, and analyzed flavonoid and tannin concentrations in leaves and roots. AG and BG herbivores formed negative feedbacks on both native and invasive genotypes. Leaf damage by weevils and the number of beetle larvae emerging as adults were higher on invasive genotypes. Beetles reduced weevil damage and weevils reduced beetle larval emergence more strongly for invasive genotypes. Invasive genotypes had lower leaf and root tannins than native genotypes. BG beetles decreased leaf tannins of native genotypes but increased root tannins of invasive genotypes. AG herbivory increased root flavonoids of invasive genotypes while BG herbivory decreased leaf flavonoids. Invasive genotypes had lower AG and BG herbivore resistance, and negative AG-BG herbivore feedbacks were much stronger for invasive genotypes. Lower tannin concentrations explained overall better AG and BG herbivore performances on invasive genotypes. However, changes in tannins and flavonoids affected AG and BG herbivores differently. These results suggest that divergent selection on chemical production in invasive plants may be critical in regulating herbivore performances and novel AG and BG herbivore communities in new environments.
Peach leaf curl disease shifts sugar metabolism in severely infected leaves from source to sink.
Moscatello, Stefano; Proietti, Simona; Buonaurio, Roberto; Famiani, Franco; Raggi, Vittorio; Walker, Robert P; Battistelli, Alberto
2017-03-01
Peach leaf curl is a disease that affects the leaves of peach trees, and in severe cases all of the leaf can be similarly affected. This study investigated some effects of this disease on the metabolism of peach leaves in which all parts of the leaf were infected. These diseased leaves contained very little chlorophyll and performed little or no photosynthesis. Compared to uninfected leaves, diseased leaves possessed higher contents of fructose and especially glucose, but lowered contents of sucrose, sorbitol and especially starch. The activities of soluble acid invertase, neutral invertase, sorbitol dehydrogenase and sucrose synthase were all higher in diseased leaves, whereas, those of aldose-6-phosphate reductase and sucrose phosphate synthase were lower. The activities of hexokinase and fructokinase were little changed. In addition, immunblots showed that the contents of Rubisco and ADP-glucose phosphorylase were reduced in diseased leaves, whereas, the content of phosphoenolpyruvate carboxylase was increased. The results show that certain aspects of the metabolism of diseased leaves are similar to immature sink leaves. That is photosynthetic function is reduced, the leaf imports rather than exports sugars, and the contents of non-structural carbohydrates and enzymes involved in their metabolism are similar to sink leaves. Further, the effects of peach leaf curl on the metabolism of peach leaves are comparable to the effects of some other diseases on the metabolism of photosynthetic organs of other plant species. Copyright © 2016 Elsevier Masson SAS. All rights reserved.
DOE Office of Scientific and Technical Information (OSTI.GOV)
Dillenburg, L.R.; Sullivan, J.H.; Teramura, A.H.
1995-07-01
In order to perform their functions as photosynthetic organs, leaves must cope with excess heat and potentially damaging ultraviolet radiation. Possible increases in the UV-B portion of the solar spectrum may place an additional burden on leaves, and this could be particularly important for young expanding leaves with poorly developed UV-B defense mechanisms. We evaluated the effects of supplemental UV-B radiation on leaf expansion and the development of photosynthetic capacity and pigments in sweetgum (Liquidambar styraciflua L.) seedlings. Seedlings were grown in the field under either ambient or ambient plus 3 or 5.0 kJ of biologically effective supplemental UV-B radiation.more » Although final leaf size was unaffected, the rate of leaf elongation and accumulation of leaf area was slower in leaves exposed to the lower supplemental UV-B irradiance. In contrast, chlorophyll accumulation and the development of photosynthetic capacity was more rapid in plants exposed to the higher, compared to the lower supplemental UV-B irradiance. The accumulation of anthocyanins and other putative flavonoids or UV-absorbing compounds was scarcely affected by exposure to supplemental UV-B radiation. These results suggest that the UV-B portion of the solar spectrum may, in the absence of gross affects on biomass, exert subtle influences on leaf ontogeny and the development of photosynthetic pigments and capacity in sweetgum. 44 refs., 6 figs.« less
USDA-ARS?s Scientific Manuscript database
Almond leaf scorch (ALS) disease has been a chronic problem for California almond growers. This disease is caused by the bacterial pathogen Xylella fastidiosa and is transmitted by xylem-feeding insects. Previous research suggested that retaining, rather than roguing, ALS-affected trees may be more ...
QTLs for resistance to the leaf rust Puccinia brachypodii in the model grass Brachypodium distachyon
USDA-ARS?s Scientific Manuscript database
The wild grass Brachypodium distachyon is a useful new model for temperate cereals, but its potential to study the interactions with pathogens remains underexploited. Leaf rust is one of the major fungal diseases affecting cereals, and recently the host status of Brachypodium to Puccinia rusts was i...
Predicting the presence of whiteflies and tomato yellow leaf curl virus in Florida tomato fields
USDA-ARS?s Scientific Manuscript database
Florida is one of the leading states for production of fresh market tomatoes. Production is severely affected by Tomato yellow leaf curl virus (TYLCV). The objective of this study was to identify landscape and climatic factors that drive whitefly populations and TYLCV incidence in commercial tomato ...
Timothy D. Meehan; Michael S. Crossley; Richard L. Lindroth
2010-01-01
Human alteration of atmospheric composition affects foliar chemistry and has possible implications for the structure and functioning of detrital communities. In this study, we explored the impacts of elevated carbon dioxide and ozone on aspen (Populus tremuloides) leaf litter chemistry, earthworm (Lumbricus terrestris) individual...
Leaf fall, humus depth, and soil frost in a northern hardwood forest
George Hart; Raymond E. Leonard; Robert S. Pierce
1962-01-01
In the mound-and-depression microtopography of the northern hardwood forest, leaves are blown off the mounds and collect in the depressions. This influence of microtopography on leaf accumulation is responsible for much of the variation in humus depth; and this, in turn, affects the formation and depth of soil frost.
USDA-ARS?s Scientific Manuscript database
Organic rice production has significantly increased in the U. S. over the last decade. Growers lack effective tools to manage sheath blight, caused by Rhizoctonia solani, and narrow brown leaf spot (NBLS), caused by Cercospora janseana, two major diseases affecting organic rice production. An experi...
Southern corn leaf blight a story worth retelling
USDA-ARS?s Scientific Manuscript database
The Southern Corn Leaf Blight Epidemic of 1970-1971 was one of the most costly disease outbreaks to affect North American agriculture, destroying 15% of the crop at a cost of $1.0 billion (US). It resulted from an over reliance on cytoplasmic Texas male sterile (cms-T) lines in hybrid seed producti...
NASA Astrophysics Data System (ADS)
Festa, G.; Senesi, R.; Alessandroni, M.; Andreani, C.; Vitali, G.; Porcinai, S.; Giusti, A. M.; Materna, T.; Paradowska, A. M.
2011-03-01
Quantitative neutron studies of cultural heritage objects provide access to microscopic, mesoscopic, and macroscopic structures in a nondestructive manner. In this paper we present a neutron diffraction investigation of a Ghiberti Renaissance gilded bronze relief devoted to the measurement of cavities and inhomogeneities in the bulk of the sample, along with the bulk phase composition and residual strain distribution. The quantitative measurements allowed the determination of the re-melting parts extension, as well as improving current knowledge about the manufacturing process. The study provides significant and unique information to conservators and restorators about the history of the relief.
Strengthening of ferrous binder jet 3D printed components through bronze infiltration
DOE Office of Scientific and Technical Information (OSTI.GOV)
Cordero, Zachary C.; Siddel, Derek H.; Peter, William H.
Fully-dense, net shape objects have been fabricated from a rapidly-solidified ferrous powder using binder-jet 3D printing and molten bronze infiltration. X-ray diffraction, scanning electron microscopy, and differential thermal analysis were used to characterize the structural evolution of the powder feedstock during an infiltration heating cycle. Microindentation and bend tests were performed on the infiltrated material to evaluate its mechanical properties. The infiltrated material had an 11 GPa hardness and moderate damage tolerance. It was found that infiltration improved both the ductility and strength of the sintered preforms by eliminating the stress concentration at the interparticle necks.
Strengthening of ferrous binder jet 3D printed components through bronze infiltration
Cordero, Zachary C.; Siddel, Derek H.; Peter, William H.; ...
2017-04-08
Fully-dense, net shape objects have been fabricated from a rapidly-solidified ferrous powder using binder-jet 3D printing and molten bronze infiltration. X-ray diffraction, scanning electron microscopy, and differential thermal analysis were used to characterize the structural evolution of the powder feedstock during an infiltration heating cycle. Microindentation and bend tests were performed on the infiltrated material to evaluate its mechanical properties. The infiltrated material had an 11 GPa hardness and moderate damage tolerance. It was found that infiltration improved both the ductility and strength of the sintered preforms by eliminating the stress concentration at the interparticle necks.
Erosion of phosphor bronze under cavitation attack in a mineral oil
NASA Technical Reports Server (NTRS)
Rao, B. C. S.; Buckley, D. H.
1986-01-01
Experimental investigations on erosion of a copper alloy, phosphor bronze, under cavitation attack in a viscous mineral oil are presented. The details of pit formation and erosion were studied using scanning electron microscopy. The mean depth of penetration, the variations in surface roughness, and the changes in erosion pit size were studied. Cavitation pits formed initially over the grain boundaries while the surface grains were plastically deformed. Erosion of surface grains occurred largely by ductile fracture involving microcracking and removal in layers. The ratio h/a of the depth h to half width a of cavitation pits increased with test duration from 0.047 to 0.55.
Bronx Zoo cogeneration project
DOE Office of Scientific and Technical Information (OSTI.GOV)
Rivet, P.H.
The New York Zoological Society commenced feasibility studies for a proposed cogeneration and district heating system for the Bronz Zoo in spring 1982. Early studies focused on evaluating the Zoo's energy loads, infrastructure, and energy delivery and financing systems. The Zoological Society and New York City joined in the decision to support the construction of a system which would serve not only the Bronx Zoo but also five nearby City-funded installations, including the adjacent New York Botanical Garden. Since the submission of that study, the project has been modified in scope, scaling back to a generating capacity designed to servemore » only the Bronz Zoo.« less
Processing and problems in manufacturing a Ti-modified Nb/sub 3/Sn MJR billet. Volume 2
DOE Office of Scientific and Technical Information (OSTI.GOV)
McDonald, W.K.; Smathers, D.; Geno, J.D.
1985-06-18
This report is submitted to complete Task II of University of California Order Number 4321405. Task I had Teledyne Wah Chang Albany (TWCA) assemble and process by the Modified Jelly Roll (MJR) method a Ti-modified Nb/sub 3/Sn superconductor billet. This billet was identified as M103 by TWCA. The billet matrix is nominally composed of copper 13.5 wt % tin bronze sheet and niobium 1.2 wt % titanium expanded metal with a volume ratio of three parts bronze to one part niobium alloy. All processing steps and problems encountered in manufacturing billet M103 are described in this report.
Dáder, Beatriz; Gwynn-Jones, Dylan; Moreno, Aránzazu; Winters, Ana; Fereres, Alberto
2014-09-05
Ultraviolet (UV) radiation directly regulates a multitude of herbivore life processes, in addition to indirectly affecting insect success via changes in plant chemistry and morphogenesis. Here we looked at plant and insect (aphid and whitefly) exposure to supplemental UV-A radiation in the glasshouse environment and investigated effects on insect population growth. Glasshouse grown peppers and eggplants were grown from seed inside cages covered by novel plastic filters, one transparent and the other opaque to UV-A radiation. At a 10-true leaf stage for peppers (53 days) and 4-true leaf stage for eggplants (34 days), plants were harvested for chemical analysis and infested by aphids and whiteflies, respectively. Clip-cages were used to introduce and monitor the insect fitness and populations of the pests studied. Insect pre-reproductive period, fecundity, fertility and intrinsic rate of natural increase were assessed. Crop growth was monitored weekly for 7 and 12 weeks throughout the crop cycle of peppers and eggplants, respectively. At the end of the insect fitness experiment, plants were harvested (68 days and 18-true leaf stage for peppers, and 104 days and 12-true leaf stage for eggplants) and leaves analysed for secondary metabolites, soluble carbohydrates, amino acids, total proteins and photosynthetic pigments. Our results demonstrate for the first time, that UV-A modulates plant chemistry with implications for insect pests. Both plant species responded directly to UV-A by producing shorter stems but this effect was only significant in pepper whilst UV-A did not affect the leaf area of either species. Importantly, in pepper, the UV-A treated plants contained higher contents of secondary metabolites, leaf soluble carbohydrates, free amino acids and total content of protein. Such changes in tissue chemistry may have indirectly promoted aphid performance. For eggplants, chlorophylls a and b, and carotenoid levels decreased with supplemental UV-A over the entire crop cycle but UV-A exposure did not affect leaf secondary metabolites. However, exposure to supplemental UV-A had a detrimental effect on whitefly development, fecundity and fertility presumably not mediated by plant cues as compounds implied in pest nutrition - proteins and sugars - were unaltered. Copyright © 2014 Elsevier B.V. All rights reserved.
Szécsényi-Nagy, Anna; Roth, Christina; Brandt, Guido; Rihuete-Herrada, Cristina; Tejedor-Rodríguez, Cristina; Held, Petra; García-Martínez-de-Lagrán, Íñigo; Arcusa Magallón, Héctor; Zesch, Stephanie; Knipper, Corina; Bánffy, Eszter; Friederich, Susanne; Meller, Harald; Bueno Ramírez, Primitiva; Barroso Bermejo, Rosa; de Balbín Behrmann, Rodrigo; Herrero-Corral, Ana M; Flores Fernández, Raúl; Alonso Fernández, Carmen; Jiménez Echevarria, Javier; Rindlisbacher, Laura; Oliart, Camila; Fregeiro, María-Inés; Soriano, Ignacio; Vicente, Oriol; Micó, Rafael; Lull, Vicente; Soler Díaz, Jorge; López Padilla, Juan Antonio; Roca de Togores Muñoz, Consuelo; Hernández Pérez, Mauro S; Jover Maestre, Francisco Javier; Lomba Maurandi, Joaquín; Avilés Fernández, Azucena; Lillios, Katina T; Silva, Ana Maria; Magalhães Ramalho, Miguel; Oosterbeek, Luiz Miguel; Cunha, Claudia; Waterman, Anna J; Roig Buxó, Jordi; Martínez, Andrés; Ponce Martínez, Juana; Hunt Ortiz, Mark; Mejías-García, Juan Carlos; Pecero Espín, Juan Carlos; Cruz-Auñón Briones, Rosario; Tomé, Tiago; Carmona Ballestero, Eduardo; Cardoso, João Luís; Araújo, Ana Cristina; Liesau von Lettow-Vorbeck, Corina; Blasco Bosqued, Concepción; Ríos Mendoza, Patricia; Pujante, Ana; Royo-Guillén, José I; Esquembre Beviá, Marco Aurelio; Dos Santos Goncalves, Victor Manuel; Parreira, Rui; Morán Hernández, Elena; Méndez Izquierdo, Elena; Vega Y Miguel, Jorge; Menduiña García, Roberto; Martínez Calvo, Victoria; López Jiménez, Oscar; Krause, Johannes; Pichler, Sandra L; Garrido-Pena, Rafael; Kunst, Michael; Risch, Roberto; Rojo-Guerra, Manuel A; Haak, Wolfgang; Alt, Kurt W
2017-11-15
Agriculture first reached the Iberian Peninsula around 5700 BCE. However, little is known about the genetic structure and changes of prehistoric populations in different geographic areas of Iberia. In our study, we focus on the maternal genetic makeup of the Neolithic (~ 5500-3000 BCE), Chalcolithic (~ 3000-2200 BCE) and Early Bronze Age (~ 2200-1500 BCE). We report ancient mitochondrial DNA results of 213 individuals (151 HVS-I sequences) from the northeast, central, southeast and southwest regions and thus on the largest archaeogenetic dataset from the Peninsula to date. Similar to other parts of Europe, we observe a discontinuity between hunter-gatherers and the first farmers of the Neolithic. During the subsequent periods, we detect regional continuity of Early Neolithic lineages across Iberia, however the genetic contribution of hunter-gatherers is generally higher than in other parts of Europe and varies regionally. In contrast to ancient DNA findings from Central Europe, we do not observe a major turnover in the mtDNA record of the Iberian Late Chalcolithic and Early Bronze Age, suggesting that the population history of the Iberian Peninsula is distinct in character.
Invasion of cemeteries in Florida by Aedes albopictus.
O'Meara, G F; Gettman, A D; Evans, L F; Scheel, F D
1992-03-01
Aedes albopictus has been found in 53 of the 67 Florida counties. The initial discoveries in 11 of these counties were made in cemeteries. At several locations, Ae. albopictus became well-established in cemeteries before appearing in nearby accumulations of waste tires. The recycling of plastic floral baskets may be aiding the spread of Ae. albopictus. Mosquitoes were commonly found in all types of flower-holding containers in cemeteries, except bronze vases. In the laboratory, most Aedes aegypti eggs laid in bronze vases hatched, but larvae subsequently died. The spread of Ae. albopictus in cemeteries seems to occur at the expense of Ae. aegypti populations. At one cemetery immature Ae. albopictus and Ae. aegypti were found in about 70% of the Aedes-positive containers at the start of a monitoring program. In subsequent collections from this site, Ae. albopictus was found in nearly all Aedes-positive containers, whereas there was a progressive decrease in containers with Ae. aegypti. This trend did not appear to be the result of any seasonal pattern because in a nearby cemetery where Ae. albopictus was absent, Ae. aegypti did not show a similar decline. Limiting flower-holding containers to those with drain holes or to bronze vases would greatly limit mosquito production.
Di Turo, Francesca; Proietti, Chiara; Screpanti, Augusto; Fornasier, M Francesca; Cionni, Irene; Favero, Gabriele; De Marco, Alessandra
2016-11-01
The interaction of pollutants with Cultural Heritage materials leads to artworks and materials degradation and loss, causing an unpriceless damage. This works aims to estimate the impacts of air pollution and meteorological conditions on limestone, copper and bronze and represents the European risk assessment for corrosion of Cultural Heritage materials. The measures and policies for atmospheric pollution reduction have cut off the SO 2 concentration and consequently its impact on materials is drastically reduced. Indeed, in 1980 the number of UNESCO sites in danger was extremely high (94% for limestone, 54% for copper and 1% for bronze) while in 2010 these sites did not exceed the tolerable value of surface recession and corrosion. However, some problem related to air pollution persists. In particular, Random Forest Analysis (RFA), highlights PM 10 as the main responsible for materials corrosion, in 2010. Two scenarios in 2030 have been tested, highlighting that the corrosion levels of limestone, copper and bronze exceed the tolerable limits only in the Balkan area and Turkey. Our results show the importance in the air quality modelling as a powerful tool for the UNESCO sites conservation. Copyright © 2016 Elsevier Ltd. All rights reserved.
Economic and social activities on ancient Cypriot terraced landscapes.
Ridder, Elizabeth; Galletti, Christopher S; Fall, Patricia L; Falconer, Steven E
2017-11-01
We investigate ancient agricultural terraces and their associated social and economic activities across the site complex consisting of the village at Politiko-Troullia and its more extensive associated taskscape. Surface artifact distributions mapped over 12 ha are integrated with evidence excavated from this Bronze Age settlement in central Cyprus. Contrary to expectations, artifact densities do not diminish with distance from the village architecture. In particular, concentrations of Prehistoric Bronze Age ceramics and ground stone artifacts are most pronounced on nearby terraced hillsides. These terraces were not utilized for domestic structures, but for extensive processing of agricultural crops and copper ore. Bronze Age excavated plant remains indicate cultivation of olives, grapes and figs, with wood resources dominated by olive and pine. Larger, non-portable ground stones and gaming stones are associated with communal social and economic activities in open courtyard settings in Politiko-Troullia. This category of ground stone also is particularly common on the terraced hillsides around Troullia, suggesting that similar behaviors occurred beyond village structures. The terraced landscape of Politiko-Troullia exemplifies a multi-faceted taskscape with a range of agricultural, metallurgical and social activities. Copyright © 2016 Elsevier Ltd. All rights reserved.
Cheng, Ruimei; Shi, Zuomin; Xu, Gexi; Liu, Shirong; Centritto, Mauro
2018-01-01
Variation in photosynthetic-nitrogen use efficiency (PNUE) is generally affected by several factors such as leaf nitrogen allocation and leaf diffusional conductances to CO2, although it is still unclear which factors significantly affect PNUE in tropical montane rain forest trees. In this study, comparison of PNUE, photosynthetic capacity, leaf nitrogen allocation, and diffusional conductances to CO2 between five Fagaceae tree species and five Leguminosae tree species were analyzed in Jianfengling tropical montane rain forest, Hainan Island, China. The result showed that PNUE of Fagaceae was significantly higher than that of Leguminosae (+35.5%), attributed to lower leaf nitrogen content per area (Narea, –29.4%). The difference in nitrogen allocation was the main biochemical factor that influenced interspecific variation in PNUE of these tree species. Fagaceae species allocated a higher fraction of leaf nitrogen to the photosynthetic apparatus (PP, +43.8%), especially to Rubisco (PR, +50.0%) and bioenergetics (PB +33.3%) in comparison with Leguminosae species. Leaf mass per area (LMA) of Leguminosae species was lower than that of Fagaceae species (-15.4%). While there was no significant difference shown for mesophyll conductance (gm), Fagaceae tree species may have greater chloroplast to total leaf surface area ratios and that offset the action of thicker cell walls on gm. Furthermore, weak negative relationship between nitrogen allocation in cell walls and in Rubisco was found for Castanopsis hystrix, Cyclobalanopsis phanera and Cy. patelliformis, which might imply that nitrogen in the leaves was insufficient for both Rubisco and cell walls. In summary, our study concluded that higher PNUE might contribute to the dominance of most Fagaceae tree species in Jianfengling tropical montane rain forest. PMID:29390007
Tang, Jingchao; Cheng, Ruimei; Shi, Zuomin; Xu, Gexi; Liu, Shirong; Centritto, Mauro
2018-01-01
Variation in photosynthetic-nitrogen use efficiency (PNUE) is generally affected by several factors such as leaf nitrogen allocation and leaf diffusional conductances to CO2, although it is still unclear which factors significantly affect PNUE in tropical montane rain forest trees. In this study, comparison of PNUE, photosynthetic capacity, leaf nitrogen allocation, and diffusional conductances to CO2 between five Fagaceae tree species and five Leguminosae tree species were analyzed in Jianfengling tropical montane rain forest, Hainan Island, China. The result showed that PNUE of Fagaceae was significantly higher than that of Leguminosae (+35.5%), attributed to lower leaf nitrogen content per area (Narea, -29.4%). The difference in nitrogen allocation was the main biochemical factor that influenced interspecific variation in PNUE of these tree species. Fagaceae species allocated a higher fraction of leaf nitrogen to the photosynthetic apparatus (PP, +43.8%), especially to Rubisco (PR, +50.0%) and bioenergetics (PB +33.3%) in comparison with Leguminosae species. Leaf mass per area (LMA) of Leguminosae species was lower than that of Fagaceae species (-15.4%). While there was no significant difference shown for mesophyll conductance (gm), Fagaceae tree species may have greater chloroplast to total leaf surface area ratios and that offset the action of thicker cell walls on gm. Furthermore, weak negative relationship between nitrogen allocation in cell walls and in Rubisco was found for Castanopsis hystrix, Cyclobalanopsis phanera and Cy. patelliformis, which might imply that nitrogen in the leaves was insufficient for both Rubisco and cell walls. In summary, our study concluded that higher PNUE might contribute to the dominance of most Fagaceae tree species in Jianfengling tropical montane rain forest.
Rodrigues, Cleiton G; Krüger, Alexandra P; Barbosa, Wagner F; Guedes, Raul Narciso C
2016-04-11
The ongoing concern about bee decline has largely focused on honey bees and neonicotinoid insecticides, while native pollinators such as Neotropical stingless bees and agrochemicals such as other insecticide groups, pesticides in general, and fertilizers-especially leaf fertilizers-remain neglected as potential contributors to pollination decline. In an effort to explore this knowledge gap, we assessed the lethal and sublethal behavioral impact of heavy metal-containing leaf fertilizers in a native pollinator of ecological importance in the Neotropics: the stingless beeFriesella schrottkyi(Friese). Two leaf fertilizers-copper sulfate (24% Cu) and a micronutrient mix (Arrank L: 5% S, 5% Zn, 3% Mn, 0.6% Cu, 0.5% B, and 0.06% Mo)-were used in oral and contact exposure bioassays. The biopesticide spinosad and water were used as positive and negative controls, respectively. Copper sulfate compromised the survival of stingless bee workers, particularly with oral exposure, although less than spinosad under contact exposure. Sublethal exposure to both leaf fertilizers at their field rates also caused significant effects in exposed workers. Copper sulfate enhanced flight take-off on stingless bee workers, unlike workers exposed to the micronutrient mix. There was no significant effect of leaf fertilizers on the overall activity and walking behavior of worker bees. No significant effect was observed for the respiration rate of worker bees under contact exposure, but workers orally exposed to the micronutrient mix exhibited a reduced respiration rate. Therefore, leaf fertilizers do affectF. schrottkyi, what may also occur with other stingless bees, potentially compromising their pollination activity deserving attention. © The Authors 2016. Published by Oxford University Press on behalf of Entomological Society of America. All rights reserved. For Permissions, please email: journals.permissions@oup.com.
Inhibitory activities of Moringa oleifera leaf extract against α-glucosidase enzyme in vitro
NASA Astrophysics Data System (ADS)
Natsir, H.; Wahab, A. W.; Laga, A.; Arif, A. R.
2018-03-01
Alpha-glucosidase is a key enzyme in the final process of breaking carbohydrates into glucose. Inhibition of α-glucosidase affected more absorption of glucose, so it can reduce hyperglycemia condition. The aims of this study is to determine the effectiveness of inhibition wet and dried Moringa oleifera leaf extract through α-glucosidase activity in vitro. The effectiveness study of inhibition on the activity of α-glucosidase enzyme obtained from white glutinous rice (Oryza sativa glutinosa) was carried out using wet and dried kelor leaf extract of 13% (w/v) with 10 mM α-D-glucopyranoside (PNPG) substrate. A positive control used 1% acarbose and substrate without addition of extract was a negative control. Inhibitory activity was measured using spectrophotometers at a wavelength of 400 nm. The result showed that the inhibition activity against α-glucosidase enzyme of dried leaf extract, wet leaf extract and acarbose was 81,39%, 83,94%, and 95,4%, respectively on pH 7,0. The effectiveness inhibition of the wet Moringa leaf extract was greater than the dried leaf extract. The findings suggest that M. oleifera leaf has the potential to be developed as an alternative food therapy for diabetics.
Salinity stress inhibits bean leaf expansion by reducing turgor, not wall extensibility
NASA Technical Reports Server (NTRS)
Neumann, P. M.; Van Volkenburgh, E.; Cleland, R. E.
1988-01-01
Treatment of bean (Phaseolus vulgaris L.) seedlings with low levels of salinity (50 or 100 millimolar NaCl) decreased the rate of light-induced leaf cell expansion in the primary leaves over a 3 day period. This decrease could be due to a reduction in one or both of the primary cellular growth parameters: wall extensibility and cell turgor. Wall extensibility was assessed by the Instron technique. Salinity did not decrease extensibility and caused small increases relative to the controls after 72 hours. On the other hand, 50 millimolar NaCl caused a significant reduction in leaf bulk turgor at 24 hours; adaptive decreases in leaf osmotic potential (osmotic adjustment) were more than compensated by parallel decreases in xylem tension potential and the leaf apoplastic solute potential, resulting in a decreased leaf water potential. It is concluded that in bean seedlings, mild salinity initially affects leaf growth rate by a decrease in turgor rather than by a reduction in wall extensibility. Moreover, long-term salinization (10 days) resulted in an apparent mechanical adjustment, i.e. an increase in wall extensibility, which may help counteract reductions in turgor and maintain leaf growth rates.
NASA Astrophysics Data System (ADS)
Goebes, Philipp; Seitz, Steffen; Kühn, Peter; Scholten, Thomas
2016-04-01
Soil erosion is crucial for degradation of carbon (C) from their pools in the soil. If C of the eroded sediment and runoff are not only related to soil pools but also resulting additively from decomposition of litter cover, the system gets more complex. The role of these amounts for C cycling in a forest environment is not yet known properly and thus, the aim of this study was to investigate the role of leaf litter diversity, litter cover and soil fauna on C redistribution during interrill erosion. We established 96 runoff plots that were deployed with seven domestic leaf litter species resulting in none species (bare ground), 1-species, 2-species and 4-species mixtures. Every second runoff plot was equipped with a fauna extinction feature to investigate the role of soil meso- and macrofauna. Erosion processes were initiated using a rainfall simulator at two time steps (summer 2012 and autumn 2012) to investigate the role of leaf litter decomposition on C redistribution. C fluxes during 20 min rainfall simulation were 99.13 ± 94.98 g/m². C fluxes and C contents both were affected by soil fauna. C fluxes were higher with presence of soil fauna due to loosening and slackening of the soil surface rather than due to faster decomposition of leaves. In contrast, C contents were higher in the absence of soil fauna possibly resulting from a missing dilution effect in the top soil layer. Leaf litter diversity did not affect C fluxes, but indirectly affected C contents as it increased the soil fauna effect with higher leaf litter diversity due to superior food supply. Initial C contents in the soil mainly determined those of the eroded sediment. For future research, it will be essential to introduce a long-term decomposition experiment to get further insights into the processes of C redistribution.
Leaf-on canopy closure in broadleaf deciduous forests predicted during winter
Twedt, Daniel J.; Ayala, Andrea J.; Shickel, Madeline R.
2015-01-01
Forest canopy influences light transmittance, which in turn affects tree regeneration and survival, thereby having an impact on forest composition and habitat conditions for wildlife. Because leaf area is the primary impediment to light penetration, quantitative estimates of canopy closure are normally made during summer. Studies of forest structure and wildlife habitat that occur during winter, when deciduous trees have shed their leaves, may inaccurately estimate canopy closure. We estimated percent canopy closure during both summer (leaf-on) and winter (leaf-off) in broadleaf deciduous forests in Mississippi and Louisiana using gap light analysis of hemispherical photographs that were obtained during repeat visits to the same locations within bottomland and mesic upland hardwood forests and hardwood plantation forests. We used mixed-model linear regression to predict leaf-on canopy closure from measurements of leaf-off canopy closure, basal area, stem density, and tree height. Competing predictive models all included leaf-off canopy closure (relative importance = 0.93), whereas basal area and stem density, more traditional predictors of canopy closure, had relative model importance of ≤ 0.51.
Palisade cell shape affects the light-induced chloroplast movements and leaf photosynthesis.
Gotoh, Eiji; Suetsugu, Noriyuki; Higa, Takeshi; Matsushita, Tomonao; Tsukaya, Hirokazu; Wada, Masamitsu
2018-01-24
Leaf photosynthesis is regulated by multiple factors that help the plant to adapt to fluctuating light conditions. Leaves of sun-light-grown plants are thicker and contain more columnar palisade cells than those of shade-grown plants. Light-induced chloroplast movements are also essential for efficient leaf photosynthesis and facilitate efficient light utilization in leaf cells. Previous studies have demonstrated that leaves of most of the sun-grown plants exhibited no or very weak chloroplast movements and could accomplish efficient photosynthesis under strong light. To examine the relationship between palisade cell shape, chloroplast movement and distribution, and leaf photosynthesis, we used an Arabidopsis thaliana mutant, angustifolia (an), which has thick leaves that contain columnar palisade cells similar to those in the sun-grown plants. In the highly columnar cells of an mutant leaves, chloroplast movements were restricted. Nevertheless, under white light condition (at 120 µmol m -2 s -1 ), the an mutant plants showed higher chlorophyll content per unit leaf area and, thus, higher light absorption by the leaves than the wild type, which resulted in enhanced photosynthesis per unit leaf area. Our findings indicate that coordinated regulation of leaf cell shape and chloroplast movement according to the light conditions is pivotal for efficient leaf photosynthesis.
NASA Astrophysics Data System (ADS)
Lestari, R. P.; Nissa, C.; Afifah, D. N.; Anjani, G.; Rustanti, N.
2018-02-01
Alternative treatment for metabolic syndrome can be done by providing a diet consist of functional foods or beverages. Synbiotic yoghurt containing binahong leaf extract which high in antioxidant, total LAB and fiber can be selected to reduce the risk of metabolic syndrome. The effect of binahong leaf extract in synbiotic yoghurt against total LAB, antioxidant activity, and acceptance were analyzed. The experiment was done with complete randomized design with addition of binahong leaf extract 0% (control); 0.12%; 0.25%; 0.5% in synbiotic yoghurt. Analysis of total LAB using Total Plate Count test, antioxidant activity using DPPH, and acceptance were analyzed by hedonic test. The addition of binahong leaf extract in various doses in synbiotic yoghurt decreased total LAB without significant effect (p=0,145). There was no effect of addition binahong leaf extract on antioxidant activity (p=0,297). The addition of binahong leaf extract had an effect on color, but not on aroma, texture and taste. The best result was yoghurt synbiotic with addition of 0,12% binahong leaf extract. Conclusion of the research was the addition of binahong leaf extract to synbiotic yogurt did not significantly affect total LAB, antioxidant activity, aroma, texture and taste; but had a significant effect on color.
Baker, Robert L; Leong, Wen Fung; Brock, Marcus T; Markelz, R J Cody; Covington, Michael F; Devisetty, Upendra K; Edwards, Christine E; Maloof, Julin; Welch, Stephen; Weinig, Cynthia
2015-10-01
Improved predictions of fitness and yield may be obtained by characterizing the genetic controls and environmental dependencies of organismal ontogeny. Elucidating the shape of growth curves may reveal novel genetic controls that single-time-point (STP) analyses do not because, in theory, infinite numbers of growth curves can result in the same final measurement. We measured leaf lengths and widths in Brassica rapa recombinant inbred lines (RILs) throughout ontogeny. We modeled leaf growth and allometry as function valued traits (FVT), and examined genetic correlations between these traits and aspects of phenology, physiology, circadian rhythms and fitness. We used RNA-seq to construct a SNP linkage map and mapped trait quantitative trait loci (QTL). We found genetic trade-offs between leaf size and growth rate FVT and uncovered differences in genotypic and QTL correlations involving FVT vs STPs. We identified leaf shape (allometry) as a genetic module independent of length and width and identified selection on FVT parameters of development. Leaf shape is associated with venation features that affect desiccation resistance. The genetic independence of leaf shape from other leaf traits may therefore enable crop optimization in leaf shape without negative effects on traits such as size, growth rate, duration or gas exchange. © 2015 The Authors. New Phytologist © 2015 New Phytologist Trust.
Impacts of drought and crayfish invasion on stream ecosystem structure and function
Magoulick, Daniel D.
2014-01-01
Drought and seasonal drying can be important disturbance events in many small streams, leading to intermittent or isolated habitats. Many small streams contain crayfish populations that are often keystone or dominant species in these systems. I conducted an experiment in stream mesocosms to examine the effects of drought and potential ecological redundancy of a native and invasive crayfish species. I examined the effects of drought (drought or control) and crayfish presence (none, native crayfish Orconectes eupunctus or invasive crayfish Orconectes neglectus) on stream mesocosm structure and function (leaf breakdown, community metabolism, periphyton, sediment and chironomid densities) in a fully factorial design. Each mesocosm contained a deep and shallow section, and drought treatments had surface water present (5-cm depth) in deep sections where tiles and leaf packs were placed. Drought and crayfish presence did not interact for any response variable. Drought significantly reduced leaf breakdown, and crayfish presence significantly increased leaf breakdown. However, the native and invasive crayfish species did not differ significantly in their effects on leaf breakdown. Drought significantly reduced primary production and community respiration overall, whereas crayfish presence did not significantly affect primary production and community respiration. Neither drought nor crayfish presence significantly affected periphyton overall. However, drought significantly reduced autotrophic index (AI), and crayfish presence increased AI. Inorganic sediment and chironomid density were not affected by drought, but both were significantly reduced by crayfish presence. O. eupunctus reduced AI and sediment more than O. neglectus did. Neither drought nor crayfish species significantly affected crayfish growth or survival. Drought can have strong effects on ecosystem function, but weaker effects on benthic structure. Crayfish can have strong effects on ecosystem structure and function regardless of drought. In stream mesocosms, native and invasive crayfish species appeared largely ecologically redundant, although subtle differences in crayfish effects could cascade throughout the food web, and further research is needed to address this question.
NASA Astrophysics Data System (ADS)
McLaren, J.; van de Weg, M. J.; Shaver, G. R.; Gough, L.
2013-12-01
Changes in global climate have resulted in a ';greening' of the Arctic as the abundance of deciduous shrub species increases. Consequently, not only the living plant community, but also the litter composition changes, which in turn can affect carbon turnover patterns in the Arctic. We examined effects of changing litter composition (both root and leaf litter) on decomposition rates with a litter bag study, and specifically focused on the impact of deciduous shrub Betula nana litter on litter decomposition from two evergreen shrubs (Ledum palustre, and Vaccinium vitis-idaea) and one graminoid (Eriophorum vaginatum) species. Additionally, we investigated how decomposition was affected by nutrient availability by placing the litterbags in an ambient and a fertilized moist acidic tundra environment. Measurements were carried out seasonally over 2 years (after snow melt, mid-growing season, end growing season). We measured litter mass loss over time, as well as the respiration rates (standardized for temperature and moisture) and temperature sensitivity of litter respiration at the time of harvesting the litter bags. For leaves, Betula litter decomposed faster than the other three species, with Eriophorum leaves decomposing the slowest. This pattern was observed for both mass loss and litter respiration rates, although the differences in respiration became smaller over time. Surprisingly, combining Betula with any other species resulted in slower overall weight loss rates than would be predicted based on monoculture weight loss rates. This contrasted with litter respiration at the time of sampling, which showed a positive mixing effect of adding Betula leaf liter to the other species. Apparently, during the first winter months (September - May) Betula litter decomposition is negatively affected by mixing the species and this legacy can still be observed in the total mass loss results later in the year. For root litter there were fewer effects of species identity on root decomposition rates; only Ledum roots decomposed slower than the other three species and the overall root litter respiration rates increased with the duration of the experiment (in contrast to leaf liter respiration). A fertilized environment had no effect on overall weight loss of the leaf or root litter within the time of our study, but leaf and root litter respiration rates were significantly larger at the end of the study in the fertilized tundra.The temperature sensitivity of leaf respiration was significantly lower for leaf litter respiration than root liter respiration after fist snow melt, but this difference disappeared throughout the first growing season and neither was influenced by species composition or fertilization with N+P. Overall, our results suggest that as arctic vegetation shifts towards shrub-dominated tundra, both species composition and seasonally dependent processes will determine effects of changing vegetation types on carbon turnover in arctic ecosystems.
Kumar, Jitendra; Gunapati, Samatha; Singh, Sudhir P; Kumar, Abhinav; Lalit, Adarsh; Sharma, Naresh C; Puranik, Rekha; Tuli, Rakesh
2013-06-01
A begomovirus and its associated alpha- and betasatellite were detected in tomato plants affected with leaf curl disease. Based on a nucleotide sequence identity of 99 %, this begomovirus was designated an isolate of cotton leaf curl Burewala virus (CLCuBuV). The alphasatellite exhibited 93 % sequence identity to cotton leaf curl Burewala alphasatellite (CLCuBuA) and is hence referred to here as a variant of CLCuBuA. The detected betasatellite was recombinant in nature and showed 70 % sequence identity to the known betasatellites. Inoculation of healthy tomato with CLCuBuV plus betasatellite, either in the presence or the absence of alphasatellite, led to typical leaf curling, while inoculation with CLCuBuV in the absence of betasatellite resulted in mild symptoms. This confirmed the role of the betasatellite in expression of disease symptoms. We propose to name the newly detected betasatellite tomato leaf curl Hajipur betasatellite (ToLCHJB).
Kaiser, Elias; Kromdijk, Johannes; Harbinson, Jeremy; Heuvelink, Ep; Marcelis, Leo F M
2017-01-01
Plants depend on photosynthesis for growth. In nature, factors such as temperature, humidity, CO 2 partial pressure, and spectrum and intensity of irradiance often fluctuate. Whereas irradiance intensity is most influential and has been studied in detail, understanding of interactions with other factors is lacking. We tested how photosynthetic induction after dark-light transitions was affected by CO 2 partial pressure (20, 40, 80 Pa), leaf temperatures (15·5, 22·8, 30·5 °C), leaf-to-air vapour pressure deficits (VPD leaf-air ; 0·5, 0·8, 1·6, 2·3 kPa) and blue irradiance (0-20 %) in tomato leaves (Solanum lycopersicum). Rates of photosynthetic induction strongly increased with CO 2 partial pressure, due to increased apparent Rubisco activation rates and reduced diffusional limitations. High leaf temperature produced slightly higher induction rates, and increased intrinsic water use efficiency and diffusional limitation. High VPD leaf-air slowed down induction rates and apparent Rubisco activation and (at 2·3 kPa) induced damped stomatal oscillations. Blue irradiance had no effect. Slower apparent Rubisco activation in elevated VPD leaf-air may be explained by low leaf internal CO 2 partial pressure at the beginning of induction. The environmental factors CO 2 partial pressure, temperature and VPD leaf-air had significant impacts on rates of photosynthetic induction, as well as on underlying diffusional, carboxylation and electron transport processes. Furthermore, maximizing Rubisco activation rates would increase photosynthesis by at most 6-8 % in ambient CO 2 partial pressure (across temperatures and humidities), while maximizing rates of stomatal opening would increase photosynthesis by at most 1-3 %. © The Author 2016. Published by Oxford University Press on behalf of the Annals of Botany Company. All rights reserved. For Permissions, please email: journals.permissions@oup.com.
NASA Astrophysics Data System (ADS)
Tucić, Branka; Tomić, Vladimir; Avramov, Stevan; Pemac, Danijela
1998-12-01
A multivariate selection analysis has been used to test the adaptiveness of several Iris pumila leaf traits that display plasticity to natural light conditions. Siblings of a synthetic population comprising 31 families of two populations from contrasting light habitats were grown at an open dune site and in the understory of a Pinus nigra stand in order to score variation in phenotypic expression of six leaf traits: number of senescent leaves, number of live leaves, leaf length, leaf width, leaf angle, and specific leaf area. The ambient light conditions affected the values of all traits studied except for specific leaf area. In accordance to ecophysiological expectations for an adaptive response to light, both leaf length and width were significantly greater while the angle between sequential leaves was significantly smaller in the woodland understory than at the exposed dune site. The relationship between leaf traits and vegetative fitness (total leaf area) differed across light habitats as predicted by functional hypotheses. The standardized linear selection gradient ( β') for leaf length and width were positive in sign in both environments, but their magnitude for leaf length was higher in the shade than under full sunlight. Since plasticity of leaf length in the woodland shade has been recognized as adaptive, fitness cost of producing plastic change in leaf length was assessed. In both of the available methods used, the two-step and the multivariate regression procedures, a rather high negative association between the fitness value and the plasticity of leaf length was obtained, indicating a cost of plasticity. The selection gradient for leaf angle was weak and significant only in the woodland understory. Genetic correlations between trait expressions in contrasting light environments were negative in sign and low in magnitude, implying a significant genetic variation for plasticity in these leaf traits. Furthermore, leaf length and leaf width were found to be genetically positively coupled, which indicates that there is a potential for these two traits to evolve toward their optimal phenotypic values even faster than would be expected if they were genetically independent.
HOW to Identify and Control Black Walnut Mycosphaerella Leaf Spots
Kenneth J. Jr. Kessler; Linda B.H. Swanson
1985-01-01
This leaf-spot disease, caused by the fungus Mycosphaerella juglandis, attacks black walnut, Juglans nigra, and Persian walnut, J. regia. Thus far, the disease has been found in North Carolina, Georgia, Illinois, and Iowa. It is important in young walnut plantations, where it causes premature defoliation, thus reducing growth and nut production. Affected walnut trees...
Yigen Chen; Therese M. Poland
2009-01-01
Biotic and abiotic environmental factors affect plant nutritional quality and defensive compounds that confer plant resistance to herbivory. Influence of leaf age, light availability, and girdling on foliar nutrition and defense of green ash (Fraxinus pennsylvanica Marsh) was examined in this study. Longevity of the emerald ash borer, ...
Richard A. MacKenzie; Tracy N. Wiegner; Frances Kinslow; Nicole Cormier; Ayron M. Strauch
2013-01-01
Abstract. We examined how invasion of tropical riparian forests by an exotic N-fixing tree (Falcataria moluccana) affects organic-matter dynamics in a Hawaiian river by comparing early stages of leaf-litter breakdown between the exotic F. moluccana and native Metrosideros polymorpha trees. We examined early...
Effects of elevated CO2 leaf diet on gypsy moth (Lepidoptera: Lymantriidae) respiration rates
Anita R. Foss; William J. Mattson; Terry M. Trier
2013-01-01
Elevated levels of CO2 affect plant growth and leaf chemistry, which in turn can alter host plant suitability for insect herbivores. We examined the suitability of foliage from trees grown from seedlings since 1997 at Aspen FACE as diet for the gypsy moth (Lymantria dispar L.) Lepidoptera: Lymantriidae: paper birch (...
USDA-ARS?s Scientific Manuscript database
Leaf and neck anthracnose caused by Colletotrichum coccodes is a new disease of onion in Michigan. To test the effect of inoculation method, ‘Prince’ onion seedlings were grown in the greenhouse and inoculated with either a conidial suspension of C. coccodes (alone or with an abrasive agent) or infe...
The importance of tree size and fecundity for wind dispersal of big-leaf mahogany
Julian M. Norghauer; Charles A. Nock; James Grogan
2011-01-01
Seed dispersal by wind is a critical yet poorly understood process in tropical forest trees. How tree size and fecundity affect this process at the population level remains largely unknown because of insufficient replication across adults. We measured seed dispersal by the endangered neotropical timber species big-leaf mahogany (Swietenia macrophylla King, Meliaceae)...
Effects of Foliar Insecticides on Leaf-Level Spectral Reflectance of Soybean.
Alves, Tavvs M; Marston, Zachary P; MacRae, Ian V; Koch, Robert L
2017-12-05
Pest-induced changes in plant reflectance are crucial for the development of pest management programs using remote sensing. However, it is unknown if plant reflectance data is also affected by foliar insecticides applied for pest management. Our study assessed the effects of foliar insecticides on leaf reflectance of soybean. A 2-yr field trial and a greenhouse trial were conducted using randomized complete block and completely randomized designs, respectively. Treatments consisted of an untreated check, a new systemic insecticide (sulfoxaflor), and two representatives of the most common insecticide classes used for soybean pest management in the north-central United States (i.e., λ-cyhalothrin and chlorpyrifos). Insecticides were applied at labeled rates recommended for controlling soybean aphid; the primary insect pest in the north-central United States. Leaf-level reflectance was measured using ground-based spectroradiometers. Sulfoxaflor affected leaf reflectance at some red and blue wavelengths but had no effect at near-infrared or green wavelengths. Chlorpyrifos affected leaf reflectance at some green, red, and near-infrared wavelengths but had no effect at blue wavelengths. λ-cyhalothrin had the least effect on spectral reflectance among the insecticides, with changes to only a few near-infrared wavelengths. Our results showing immediate and delayed effects of foliar insecticides on soybean reflectance indicate that application of some insecticides may confound the use of remote sensing for detection of not only insects but also plant diseases, nutritional and water deficiencies, and other crop stressors. © The Author(s) 2017. Published by Oxford University Press on behalf of Entomological Society of America. All rights reserved. For permissions, please e-mail: journals.permissions@oup.com.
Monroy, Silvia; Menéndez, Margarita; Basaguren, Ana; Pérez, Javier; Elosegi, Arturo; Pozo, Jesús
2016-12-15
Drought, an important environmental factor affecting the functioning of stream ecosystems, is likely to become more prevalent in the Mediterranean region as a consequence of climate change and enhanced water demand. Drought can have profound impacts on leaf litter decomposition, a key ecosystem process in headwater streams, but there is still limited information on its effects at the regional scale. We measured leaf litter decomposition across a gradient of aridity in the Ebro River basin. We deployed coarse- and fine-mesh bags with alder and oak leaves in 11 Mediterranean calcareous streams spanning a range of over 400km, and determined changes in discharge, water quality, leaf-associated macroinvertebrates, leaf quality and decomposition rates. The study streams were subject to different degrees of drought, specific discharge (Ls -1 km -2 ) ranging from 0.62 to 9.99. One of the streams dried out during the experiment, another one reached residual flow, whereas the rest registered uninterrupted flow but with different degrees of flow variability. Decomposition rates differed among sites, being lowest in the 2 most water-stressed sites, but showed no general correlation with specific discharge. Microbial decomposition rates were not correlated with final nutrient content of litter nor to fungal biomass. Total decomposition rate of alder was positively correlated to the density and biomass of shredders; that of oak was not. Shredder density in alder bags showed a positive relationship with specific discharge during the decomposition experiment. Overall, the results point to a complex pattern of litter decomposition at the regional scale, as drought affects decomposition directly by emersion of bags and indirectly by affecting the functional composition and density of detritivores. Copyright © 2016 The Authors. Published by Elsevier B.V. All rights reserved.
NASA Astrophysics Data System (ADS)
Kahmen, A.; Merchant, A.; Callister, A.; Dawson, T. E.; Arndt, S. K.
2006-12-01
Stable isotopes have been a valuable tool to study water or carbon fluxes of plants and ecosystems. In particular oxygen isotopes (δ18O) in leaf water or plant organic material are now beginning to be established as a simple and integrative measure for plant - water relations. Current δ18O models, however, are still limited in their application to a broad range of different species and ecosystems. It remains for example unclear, if species-specific effects such as different leaf morphologies need to be included in the models for a precise understanding and prediction of δ18O signals. In a common garden experiment (Currency Creek Arboretum, South Australia), where over 900 different Eucalyptus species are cultivated in four replicates, we tested effects of leaf morphology and anatomy on δ18O signals in leaf water of 25 different species. In particular, we determined for all species enrichment in 18O of mean lamina leaf water above source water (Δ18O) as related to leaf physiology as well as leaf thickness, leaf area, specific leaf area and weight and selected anatomical properties. Our data revealed that diurnal Δ18O in leaf water at steady state was significantly different among the investigated species and with differences up to 10% at midday. Fitting factors (effective path length) of leaf water Δ18O models were also significantly different among the investigated species and were highly affected by species-specific morphological parameters. For example, leaf area explained a high percentage of the differences in effective path length observed among the investigated species. Our data suggest that leaf water δ18O can act as powerful tool to estimate plant - water relations in comparative studies but that additional leaf morphological parameters need to be considered in existing δ18O models for a better interpretation of the observed δ18O signals.
Remote detection of air pollution stress to vegetation - Laboratory-level studies
NASA Technical Reports Server (NTRS)
Westman, Walter E.; Price, Curtis V.
1987-01-01
An experimental investigation of the role of leaf chemistry, anatomy, moisture content, and canopy density on spectral reflectance in healthy and pollution stressed western conifer needles and broad-leafed species of California coastal sage scrub is presented. Acid mist at a level of pH 2.0 is found to more severely effect chlorophyll loss and leaf death than ozone at a level of 0.2 ppm for a four-week period. Both pollutants cause water loss, affecting Bands 4 and 5 in nonlinear ways. The infrared bands initially rise as free water is lost, and subsequently, scattering and reflectance decline. The net effect is shown to be a reduction in TM 4/3 and a rise in TM 5/4 with pollution stress. Under more severe pollution stresses, the decline of leaf area indices due to accelerated leaf drop accentuates the expected TM 4/3 and TM 5/4 changes.
Fundamental trade-offs generating the worldwide leaf economics spectrum.
Shipley, Bill; Lechowicz, Martin J; Wright, Ian; Reich, Peter B
2006-03-01
Recent work has identified a worldwide "economic" spectrum of correlated leaf traits that affects global patterns of nutrient cycling and primary productivity and that is used to calibrate vegetation-climate models. The correlation patterns are displayed by species from the arctic to the tropics and are largely independent of growth form or phylogeny. This generality suggests that unidentified fundamental constraints control the return of photosynthates on investments of nutrients and dry mass in leaves. Using novel graph theoretic methods and structural equation modeling, we show that the relationships among these variables can best be explained by assuming (1) a necessary trade-off between allocation to structural tissues versus liquid phase processes and (2) an evolutionary tradeoff between leaf photosynthetic rates, construction costs, and leaf longevity.
Are leaf chemistry signatures preserved at the canopy level?
DOE Office of Scientific and Technical Information (OSTI.GOV)
Borel, C.C.; Gerstl, S.A.W.
1994-05-01
Imaging spectrometers have the potential to be very useful in remote sensing of canopy chemistry constituents such as nitrogen and lignin. In this study under the HIRIS project the question of how leaf chemical composition which is reflected in leaf spectral features in the reflectance and transmittance is affected by canopy architecture was investigated. Several plants were modeled with high fidelity and a radiosity model was used to compute the canopy spectral signature over the visible and near infrared. We found that chemical constituent specific signatures such as absorptions are preserved and in the case of low absorption are actuallymore » enhanced. For moderately dense canopies the amount of a constituent depends also on the total leaf area.« less
Duarte, Sofia; Cássio, Fernanda; Ferreira, Verónica; Canhoto, Cristina; Pascoal, Cláudia
2016-08-01
Ongoing climate change is expected to affect the diversity and activity of aquatic microbes, which play a key role in plant litter decomposition in forest streams. We used a before-after control-impact (BACI) design to study the effects of warming on a forest stream reach. The stream reach was divided by a longitudinal barrier, and during 1 year (ambient year) both stream halves were at ambient temperature, while in the second year (warmed year) the temperature in one stream half was increased by ca. 3 °C above ambient temperature (experimental half). Fine-mesh bags containing oak (Quercus robur L.) leaves were immersed in both stream halves for up to 60 days in spring and autumn of the ambient and warmed years. We assessed leaf-associated microbial diversity by denaturing gradient gel electrophoresis and identification of fungal conidial morphotypes and microbial activity by quantifying leaf mass loss and productivity of fungi and bacteria. In the ambient year, no differences were found in leaf decomposition rates and microbial productivities either between seasons or stream halves. In the warmed year, phosphorus concentration in the stream water, leaf decomposition rates, and productivity of bacteria were higher in spring than in autumn. They did not differ between stream halves, except for leaf decomposition, which was higher in the experimental half in spring. Fungal and bacterial communities differed between seasons in both years. Seasonal changes in stream water variables had a greater impact on the activity and diversity of microbial decomposers than a warming regime simulating a predicted global warming scenario.
NASA Astrophysics Data System (ADS)
González-Zurdo, P.; Escudero, A.; Nuñez, R.; Mediavilla, S.
2016-11-01
In temperate climates, evergreen leaves have to survive throughout low temperature winter periods. Freezing and chilling injuries can lead to accelerated senescence of part of the leaf surface, which contributes to a reduction of the lifespan of the photosynthetic machinery and of leaf lifetime carbon gain. Low temperatures are also associated with changes in foliar chemistry and morphology that affect consumption by herbivores. Therefore, the severity of foliar area losses caused by accelerated senescence and herbivory can change along winter temperature gradients. The aim of this study is to analyse such responses in the leaves of three evergreen species ( Quercus ilex, Q. suber and Pinus pinaster) along a climatic gradient. The leaves of all three species presented increased leaf mass per area (LMA) and higher concentrations of structural carbohydrates in cooler areas. Only the two oak species showed visible symptoms of damage caused by herbivory, this being less intense at the coldest sites. The leaves of all three species presented chlorotic and necrotic spots that increased in size with leaf age. The foliar surface affected by chlorosis and necrosis was larger at the sites with the coldest winters. Therefore, the effects of the winter cold on the lifespan of the photosynthetic machinery were contradictory: losses of leaf area due to accelerated senescence increased, but there was a decrease in losses caused by herbivory. The final consequences for carbon assimilation strongly depend on the exact timing of the appearance of the damage resulting from low temperature and grazing by herbivores.
Girardoz, S; Tomov, R; Eschen, R; Quicke, D L J; Kenis, M
2007-10-01
The horse-chestnut leaf miner, Cameraria ohridella, is an invasive alien species defoliating horse-chestnut, a popular ornamental tree in Europe. This paper presents quantitative data on mortality factors affecting larvae and pupae of the leaf miner in Switzerland and Bulgaria, both in urban and forest environments. Two sampling methods were used and compared: a cohort method, consisting of the surveying of pre-selected mines throughout their development, and a grab sampling method, consisting of single sets of leaves collected and dissected at regular intervals. The total mortality per generation varied between 14 and 99%. Mortality was caused by a variety of factors, including parasitism, host feeding, predation by birds and arthropods, plant defence reaction, leaf senescence, intra-specific competition and inter-specific competition with a fungal disease. Significant interactions were found between mortality factors and sampling methods, countries, environments and generation. No mortality factor was dominant throughout the sites, generations and methods tested. Plant defence reactions constituted the main mortality factor for the first two larval stages, whereas predation by birds and arthropods and parasitism were more important in older larvae and pupae. Mortality caused by leaf senescence was often the dominant mortality factor in the last annual generation. The cohort method detected higher mortality rates than the grab sampling method. In particular, mortality by plant defence reaction and leaf senescence were better assessed using the cohort method, which is, therefore, recommended for life table studies on leaf miners.
Hao, Xingyu; Li, Ping; Feng, Yongxiang; Han, Xue; Gao, Ji; Lin, Erda; Han, Yuanhuai
2013-01-01
Traditional Chinese medicine relies heavily on herbs, yet there is no information on how these herb plants would respond to climate change. In order to gain insight into such response, we studied the effect of elevated [CO2] on Isatis indigotica Fort, one of the most popular Chinese herb plants. The changes in leaf photosynthesis, chlorophyll fluorescence, leaf ultrastructure and biomass yield in response to elevated [CO2] (550±19 µmol mol(-1)) were determined at the Free-Air Carbon dioxide Enrichment (FACE) experimental facility in North China. Photosynthetic ability of I. indigotica was improved under elevated [CO2]. Elevated [CO2] increased net photosynthetic rate (P N), water use efficiency (WUE) and maximum rate of electron transport (J max) of upper most fully-expended leaves, but not stomatal conductance (gs), transpiration ratio (Tr) and maximum velocity of carboxylation (V c,max). Elevated [CO2] significantly increased leaf intrinsic efficiency of PSII (Fv'/Fm') and quantum yield of PSII(ΦPS II ), but decreased leaf non-photochemical quenching (NPQ), and did not affect leaf proportion of open PSII reaction centers (qP) and maximum quantum efficiency of PSII (Fv/Fm). The structural chloroplast membrane, grana layer and stroma thylakoid membranes were intact under elevated [CO2], though more starch grains were accumulated within the chloroplasts than that of under ambient [CO2]. While the yield of I. indigotica was higher due to the improved photosynthesis under elevated [CO2], the content of adenosine, one of the functional ingredients in indigowoad root was not affected.
Kaur, Jagdeep; Fellers, John; Adholeya, Alok; Velivelli, Siva L S; El-Mounadi, Kaoutar; Nersesian, Natalya; Clemente, Thomas; Shah, Dilip
2017-02-01
Rust fungi of the order Pucciniales are destructive pathogens of wheat worldwide. Leaf rust caused by the obligate, biotrophic basidiomycete fungus Puccinia triticina (Pt) is an economically important disease capable of causing up to 50 % yield losses. Historically, resistant wheat cultivars have been used to control leaf rust, but genetic resistance is ephemeral and breaks down with the emergence of new virulent Pt races. There is a need to develop alternative measures for control of leaf rust in wheat. Development of transgenic wheat expressing an antifungal defensin offers a promising approach to complement the endogenous resistance genes within the wheat germplasm for durable resistance to Pt. To that end, two different wheat genotypes, Bobwhite and Xin Chun 9 were transformed with a chimeric gene encoding an apoplast-targeted antifungal plant defensin MtDEF4.2 from Medicago truncatula. Transgenic lines from four independent events were further characterized. Homozygous transgenic wheat lines expressing MtDEF4.2 displayed resistance to Pt race MCPSS relative to the non-transgenic controls in growth chamber bioassays. Histopathological analysis suggested the presence of both pre- and posthaustorial resistance to leaf rust in these transgenic lines. MtDEF4.2 did not, however, affect the root colonization of a beneficial arbuscular mycorrhizal fungus Rhizophagus irregularis. This study demonstrates that the expression of apoplast-targeted plant defensin MtDEF4.2 can provide substantial resistance to an economically important leaf rust disease in transgenic wheat without negatively impacting its symbiotic relationship with the beneficial mycorrhizal fungus.
Perdomo, Juan Alejandro; Conesa, Miquel À; Medrano, Hipólito; Ribas-Carbó, Miquel; Galmés, Jeroni
2015-10-01
This study evaluates the long-term individual and combined effects of high temperature (HT) and water deficit (WD) stress on plant growth, leaf gas-exchange and water use efficiency in cultivars of the three most important crops worldwide, rice, wheat and maize. Total plant biomass (B t ) accumulation decreased under all treatments, being the combined HT-WD treatment the most detrimental in all three species. Although decreases in B t correlated with adjustments in biomass allocation patterns (i.e. the leaf area ratio), most of the variation observed in B t was explained by changes in leaf gas exchange parameters. Thus, integrated values of leaf carbon balance obtained from daily course measurements of photosynthesis and respiration were better predictors of plant growth than the instantaneous measurements of leaf gas exchange. Leaf water use efficiency, assessed both by gas exchange and carbon isotope measurements, was negatively correlated with B t under WD, but not under the combined WD and HT treatment. A comparative analysis of the negative effects of single and combined stresses on the main parameters showed an additive component for WD and HT in rice and maize, in contrast to wheat. Overall, the results of the specific cultivars included in the study suggest that the species native climate plays a role shaping the species acclimation potential to the applied stresses. In this regard, wheat, originated in a cold climate, was the most affected species, which foretells a higher affectation of this crop due to climate change. © 2014 Scandinavian Plant Physiology Society.
Moore, Bd.; Palmquist, D. E.; Seemann, J. R.
1997-01-01
We have examined the possible role of leaf cytosolic hexoses and the expression of mannitol metabolism as mechanisms that may affect the repression of photosynthetic capacity when plants are grown at 1000 versus 380 [mu]L L-1 CO2. In plants grown at high CO2, leaf ribulose-1,5-bisphosphate carboxylase/oxygenase content declined by [greater than or equal to]20% in tobacco (Nicotiana sylvestris) but was not affected in the mannitol-producing species snapdragon (Antirrhinum majus) and parsley (Petroselinum hortense). In the three species mesophyll glucose and fructose at midday occurred almost entirely in the vacuole (>99%), irrespective of growth CO2 levels. The estimated cytosolic concentrations of glucose and fructose were [less than or equal to]100 [mu]M. In the three species grown at high CO2, total leaf carbohydrates increased 60 to 100%, but mannitol metabolism did not function as an overflow mechanism for the increased accumulation of carbohydrate. In both snapdragon and parsley grown at ambient or high CO2, mannitol occurred in the chloroplast and cytosol at estimated midday concentrations of 0.1 M or more each. The compartmentation of leaf hexoses and the metabolism of alternate carbohydrates are further considered in relation to photosynthetic acclimation to high levels of CO2. PMID:12223804
Haworth, Matthew; Belcher, Claire M; Killi, Dilek; Dewhirst, Rebecca A; Materassi, Alessandro; Raschi, Antonio; Centritto, Mauro
2018-04-18
Global warming events have coincided with turnover of plant species at intervals in Earth history. As mean global temperatures rise, the number, frequency and duration of heat-waves will increase. Ginkgo biloba was grown under controlled climatic conditions at two different day/night temperature regimes (25/20 °C and 35/30 °C) to investigate the impact of heat stress. Photosynthetic CO 2 -uptake and electron transport were reduced at the higher temperature, while rates of respiration were greater; suggesting that the carbon balance of the leaves was adversely affected. Stomatal conductance and the potential for evaporative cooling of the leaves was reduced at the higher temperature. Furthermore, the capacity of the leaves to dissipate excess energy was also reduced at 35/30 °C, indicating that photo-protective mechanisms were no longer functioning effectively. Leaf economics were adversely affected by heat stress, exhibiting an increase in leaf mass per area and leaf construction costs. This may be consistent with the selective pressures experienced by fossil Ginkgoales during intervals of global warming such as the Triassic - Jurassic boundary or Early Eocene Climatic Optimum. The physiological and morphological responses of the G. biloba leaves were closely interrelated; these relationships may be used to infer the leaf economics and photosynthetic/stress physiology of fossil plants.
Gontijo, Lessando M; Nechols, James R; Margolies, David C; Cloyd, Raymond A
2012-01-01
The arrangement, number, and size of plant parts may influence predator foraging behavior, either directly, by altering the rate or pattern of predator movement, or, indirectly, by affecting the distribution and abundance of prey. We report on the effects of both plant architecture and prey distribution on foraging by the predatory mite, Phytoseiulus persimilis Athias-Henriot (Acari: Phytoseiidae), on cucumber (Cucumis sativus L.). Plants differed in leaf number (2- or 6-leafed), and there were associated differences in leaf size, plant height, and relative proportions of plant parts; but all had the same total surface area. The prey, the twospotted spider mite Tetranychus urticae Koch (Acari: Tetranychidae), were distributed either on the basal leaf or on all leaves. The effect of plant architecture on predator foraging behavior varied depending on prey distribution. The dimensions of individual plant parts affected time allocated to moving and feeding, but they did not appear to influence the frequency with which predators moved among different plant parts. Overall, P. persimilis moved less, and fed upon prey longer, on 6-leafed plants with prey on all leaves than on plants representing other treatment combinations. Our findings suggest that both plant architecture and pattern of prey distribution should be considered, along with other factors such as herbivore-induced plant volatiles, in augmentative biological control programs.
NASA Astrophysics Data System (ADS)
Ghiara, G.; Grande, C.; Ferrando, S.; Piccardo, P.
2018-01-01
In this study, tin-bronze analogues of archaeological objects were investigated in the presence of an aerobic Pseudomonas fluorescens strain in a solution, containing chlorides, sulfates, carbonates and nitrates according to a previous archaeological characterization. Classical fixation protocols were employed in order to verify the attachment capacity of such bacteria. In addition, classical metallurgical analytical techniques were used to detect the effect of bacteria on the formation of uncommon corrosion products in such an environment. Results indicate quite a good attachment capacity of the bacteria to the metallic surface and the formation of the uncommon corrosion products sulfates and sulfides is probably connected to the bacterial metabolism.
DOE Office of Scientific and Technical Information (OSTI.GOV)
Krause, T. W.; Harlley, D.; Babbar, V. K.
Nickel Aluminum Bronze (NAB) is a material with marine environment applications that under certain conditions can undergo selective phase corrosion (SPC). SPC involves the removal of minority elements while leaving behind a copper matrix. Pulsed eddy current (PEC) was evaluated for determination of SPC thickness on a NAB valve section with access from the surface corroded side. A primarily linear response of PEC amplitude, up to the maximum available SPC thickness of 4 mm was observed. The combination of reduced conductivity and permeability in the SPC phase relative to the base NAB was used to explain the observed sensitivity ofmore » PEC to SPC thickness variations.« less
Classification of Corrosion Defects in NiAl Bronze Through Image Analysis
2010-01-01
the sub-band coefficient matrix as: EntropypðkÞ ¼ X i X j jCpk;ði;jÞj 2 log jCpk;ði;jÞj 2 ð2Þ EnergypðkÞ ¼ X i X j jCpk;ði;jÞj 2 ð3Þ where...Wharton, R.C. Barik , G. Kear, R.J.K. Wood, K.R. Stokes, F.C. Walsh, The corrosion of nickel–aluminum bronze in seawater, Corros. Sci. 47 (2005) 3336...corrosion under fatigue conditions, J . Aircraft (7–8) (2009) 1253–1259. [10] H.S. Isaacs, Initiation of stress corrosion cracking of sensitized type
NASA Technical Reports Server (NTRS)
1997-01-01
HyComp(R), Inc. development a line of high temperature carbon fiber composite products to solve wear problems in the harsh environment of steel and aluminum mills. WearComp(R), self-lubricating composite wear liners and bushings, combines carbon graphite fibers with a polyimide binder. The binder, in conjunction with the fibers, provides the slippery surface, one that demands no lubrication, yet wears at a very slow rate. WearComp(R) typically lasts six to ten times longer than aluminum bronze. Unlike bronze, WearComp polishes the same surface and imparts a self-lube film for years of service. It is designed for continuous operation at temperatures of 550 degrees Fahrenheit and can operate under high compressive loads.
Solar Orientations of Bronze Age Shrines in Gournia, Crete
NASA Astrophysics Data System (ADS)
Henriksson, G.; Blomberg, M.
2009-08-01
The measurements of orientations at Gournia are part of the Uppsala University project to investigate the nature of Minoan astronomy. The town is typical of Minoan settlements and therefore was chosen for the selection of representative buildings from different kinds of Minoan sites. We use classical archaeoastronomical methods and our own computer programs for calculation of the positions of the sun, moon and stars in the Aegean Bronze Age. We discovered that Gournia is the fourth Minoan site with shrines that are oriented to sunrise at lunar month intervals with respect to the equinoxes. There is also a Mycenaean shrine in the town and we compared its orientation to Mycenaean buildings of the same type in Crete.
Geoscience rediscovers Phoenicia's buried harbors
NASA Astrophysics Data System (ADS)
Marriner, Nick; Morhange, Christophe; Doumet-Serhal, Claude; Carbonel, Pierre
2006-01-01
After centuries of archaeological debate, the harbors of Phoenicia's two most important city states, Tyre and Sidon, have been rediscovered, and including new geoarcheological results reveal how, where, and when they evolved after their Bronze Age foundations. The early ports lie beneath their present urban centers, and we have indentified four harbor phases. (1) During the Bronze Age, Tyre and Sidon were characterized by semi-open marine coves that served as protoharbors. (2) Biostratigraphic and lithostratigraphic data indicate the presence of early artificial basins after the first millennium B.C. (3) The harbors reached their apogees during the Greco-Roman and Byzantine periods. (4) Silting up and coastal progradation led to burial of the medieval basins, lost until now.
Synthesis and characterization of thin-transparent nanostructured films for surface protection
NASA Astrophysics Data System (ADS)
Veltri, S.; Sokullu, E.; Barberio, M.; Gauthier, M. A.; Antici, P.
2017-01-01
This work demonstrates that very thin and optically transparent nanocomposite films can be conveniently applied on surface materials, displaying potent antibacterial properties without affecting the aesthetics of the underlying material. In our approach we propose new composite materials, which ensure the surface protection by inactivating the bacteria before a biofilm can be formed. The films contain very small loadings of TiO2, graphene, or fullerene, and can easily be applied on large surfaces using conventional brushes or air-brushes. These nanocomposite films are very promising candidates for the preservation of statues, mosaics, floors, buildings, and other objects that are exposed to challenging environmental conditions such as Architectonical Heritage or building materials (materials featuring stone, pigments, bronze, granite, marble, and glass).
Non-linear direct effects of acid rain on leaf photosynthetic rate of terrestrial plants.
Dong, Dan; Du, Enzai; Sun, Zhengzhong; Zeng, Xuetong; de Vries, Wim
2017-12-01
Anthropogenic emissions of acid precursors have enhanced global occurrence of acid rain, especially in East Asia. Acid rain directly suppresses leaf function by eroding surface waxes and cuticle and leaching base cations from mesophyll cells, while the simultaneous foliar uptake of nitrates in rainwater may directly benefit leaf photosynthesis and plant growth, suggesting a non-linear direct effect of acid rain. By synthesizing data from literature on acid rain exposure experiments, we assessed the direct effects of acid rain on leaf photosynthesis across 49 terrestrial plants in China. Our results show a non-linear direct effect of acid rain on leaf photosynthetic rate, including a neutral to positive effect above pH 5.0 and a negative effect below that pH level. The acid rain sensitivity of leaf photosynthesis showed no significant difference between herbs and woody species below pH 5.0, but the impacts above that pH level were strongly different, resulting in a significant increase in leaf photosynthetic rate of woody species and an insignificant effect on herbs. Our analysis also indicates a positive effect of the molar ratio of nitric versus sulfuric acid in the acid solution on leaf photosynthetic rate. These findings imply that rainwater acidity and the composition of acids both affect the response of leaf photosynthesis and therefore result in a non-linear direct effect. Copyright © 2017 Elsevier Ltd. All rights reserved.
Rasheed, Fahad; Dreyer, Erwin; Richard, Béatrice; Brignolas, Franck; Brendel, Oliver; Le Thiec, Didier
2015-04-01
Poplar genotypes differ in transpiration efficiency (TE) at leaf and whole-plant level under similar conditions. We tested whether atmospheric vapour pressure deficit (VPD) affected TE to the same extent across genotypes. Six Populus nigra genotypes were grown under two VPD. We recorded (1) (13)C content in soluble sugars; (2) (18)O enrichment in leaf water; (3) leaf-level gas exchange; and (4) whole-plant biomass accumulation and water use. Whole-plant and intrinsic leaf TE and (13)C content in soluble sugars differed significantly among genotypes. Stomatal conductance contributed more to these differences than net CO2 assimilation rate. VPD increased water use and reduced whole-plant TE. It increased intrinsic leaf-level TE due to a decline in stomatal conductance. It also promoted higher (18)O enrichment in leaf water. VPD had no genotype-specific effect. We detected a deviation in the relationship between (13)C in leaf sugars and (13)C predicted from gas exchange and the standard discrimination model. This may be partly due to genotypic differences in mesophyll conductance, and to its lack of sensitivity to VPD. Leaf-level (13)C discrimination was a powerful predictor of the genetic variability of whole-plant TE irrespective of VPD during growth. © 2014 John Wiley & Sons Ltd.
Limousin, Jean-Marc; Bickford, Christopher P; Dickman, Lee T; Pangle, Robert E; Hudson, Patrick J; Boutz, Amanda L; Gehres, Nathan; Osuna, Jessica L; Pockman, William T; McDowell, Nate G
2013-10-01
Leaf gas-exchange regulation plays a central role in the ability of trees to survive drought, but forecasting the future response of gas exchange to prolonged drought is hampered by our lack of knowledge regarding potential acclimation. To investigate whether leaf gas-exchange rates and sensitivity to drought acclimate to precipitation regimes, we measured the seasonal variations of leaf gas exchange in a mature piñon-juniper Pinus edulis-Juniperus monosperma woodland after 3 years of precipitation manipulation. We compared trees receiving ambient precipitation with those in an irrigated treatment (+30% of ambient precipitation) and a partial rainfall exclusion (-45%). Treatments significantly affected leaf water potential, stomatal conductance and photosynthesis for both isohydric piñon and anisohydric juniper. Leaf gas exchange acclimated to the precipitation regimes in both species. Maximum gas-exchange rates under well-watered conditions, leaf-specific hydraulic conductance and leaf water potential at zero photosynthetic assimilation all decreased with decreasing precipitation. Despite their distinct drought resistance and stomatal regulation strategies, both species experienced hydraulic limitation on leaf gas exchange when precipitation decreased, leading to an intraspecific trade-off between maximum photosynthetic assimilation and resistance of photosynthesis to drought. This response will be most detrimental to the carbon balance of piñon under predicted increases in aridity in the southwestern USA. © 2013 John Wiley & Sons Ltd.
Halilou, Oumarou; Hissene, Halime Mahamat; Clavijo Michelangeli, José A; Hamidou, Falalou; Sinclair, Thomas R; Soltani, Afshin; Mahamane, Saadou; Vadez, Vincent
2016-12-01
Rapid leaf area development may be attractive under a number of cropping conditions to enhance the vigor of crop establishment and allow rapid canopy closure for maximizing light interception and shading of weed competitors. This study was undertaken to determine (1) if parameters describing leaf area development varied among ten peanut ( Arachis hypogeae L.) genotypes grown in field and pot experiments, (2) if these parameters were affected by the planting density, and (3) if these parameters varied between Spanish and Virginia genotypes. Leaf area development was described by two steps: prediction of main stem number of nodes based on phyllochron development and plant leaf area dependent based on main stem node number. There was no genetic variation in the phyllochron measured in the field. However, the phyllochron was much longer for plants grown in pots as compared to the field-grown plants. These results indicated a negative aspect of growing peanut plants in the pots used in this experiment. In contrast to phyllochron, there was no difference in the relationship between plant leaf area and main stem node number between the pot and field experiments. However, there was genetic variation in both the pot and field experiments in the exponential coefficient (PLAPOW) of the power function used to describe leaf area development from node number. This genetic variation was confirmed in another experiment with a larger number of genotypes, although possible G × E interaction for the PLAPOW was found. Sowing density did not affect the power function relating leaf area to main stem node number. There was also no difference in the power function coefficient between Spanish and Virginia genotypes. SSM (Simple Simulation model) reliably predicted leaf canopy development in groundnut. Indeed the leaf area showed a close agreement between predicted and observed values up to 60000 cm 2 m -2 . The slightly higher prediction in India and slightly lower prediction in Niger reflected GxE interactions. Until more understanding is obtained on the possible GxE interaction effects on the canopy development, a generic PLAPOW value of 2.71, no correction for sowing density, and a phyllochron on 53 °C could be used to model canopy development in peanut.
NASA Astrophysics Data System (ADS)
Yuan, Chuan; Gao, Guangyao; Fu, Bojie
2017-03-01
Stemflow transports nutrient-enriched precipitation to the rhizosphere and functions as an efficient terrestrial flux in water-stressed ecosystems. However, its ecological significance has generally been underestimated because it is relatively limited in amount, and the biotic mechanisms that affect it have not been thoroughly studied at the leaf scale. This study was conducted during the 2014 and 2015 rainy seasons at the northern Loess Plateau of China. We measured the branch stemflow volume (SFb), shrub stemflow equivalent water depth (SFd), stemflow percentage of incident precipitation (SF %), stemflow productivity (SFP), funnelling ratio (FR), the meteorological characteristics and the plant traits of branches and leaves of C. korshinskii and S. psammophila. This study evaluated stemflow efficiency for the first time with the combined results of SFP and FR, and sought to determine the inter- and intra-specific differences of stemflow yield and efficiency between the two species, as well as the specific bio-/abiotic mechanisms that affected stemflow. The results indicated that C. korshinskii had a greater stemflow yield and efficiency at all precipitation levels than that of S. psammophila. The largest inter-specific difference generally occurred at the 5-10 mm branches during rains of ≤ 2 mm. Precipitation amount was the most influential meteorological characteristic that affected stemflow yield and efficiency in these two endemic shrub species. Branch angle was the most influential plant trait on FR. For SFb, stem biomass and leaf biomass were the most influential plant traits for C. korshinskii and S. psammophila, respectively. For SFP of these two shrub species, leaf traits (the individual leaf area) and branch traits (branch size and biomass allocation pattern) had a great influence during lighter rains ≤ 10 mm and heavier rains > 15 mm, respectively. The lower precipitation threshold to start stemflow allowed C. korshinskii (0.9 mm vs. 2.1 mm for S. psammophila) to employ more rains to harvest water via stemflow. The beneficial leaf traits (e.g., leaf shape, arrangement, area, amount) might partly explain the greater stemflow production of C. korshinskii. Comparison of SFb between the foliated and manually defoliated shrubs during the 2015 rainy season indicated that the newly exposed branch surface at the defoliated period and the resulting rainfall intercepting effects might be an important mechanism affecting stemflow in the dormant season.
Yu, Runze; Cook, Michael G; Yacco, Ralph S; Watrelot, Aude A; Gambetta, Gregory; Kennedy, James A; Kurtural, S Kaan
2016-11-02
The relationships between variations in grapevine (Vitis vinifera L. cv. Merlot) fruit zone light exposure and water deficits and the resulting berry flavonoid composition were investigated in a hot climate. The experimental design involved application of mechanical leaf removal (control, pre-bloom, post-fruit set) and differing water deficits (sustained deficit irrigation and regulated deficit irrigation). Flavonol and anthocyanin concentrations were measured by C18 reversed-phased HPLC and increased with pre-bloom leaf removal in 2013, but with post-fruit set leaf removal in 2014. Proanthocyanidin isolates were characterized by acid catalysis in the presence of excess phloroglucinol followed by reversed-phase HPLC. Post-fruit set leaf removal increased total proanthocyanidin concentration in both years, whereas no effect was observed with applied water amounts. Mean degree of polymerization of skin proanthocyanidins increased with post-fruit set leaf removal compared to pre-bloom, whereas water deficit had no effect. Conversion yield was greater with post-fruit set leaf removal. Seed proanthocyanidin concentration was rarely affected by applied treatments. The application of post-fruit set leaf removal, regardless of water deficit. increased the proportion of proanthocyanidins derived from the skin, whereas no leaf removal or pre-bloom leaf removal regardless of water deficit increased the proportion of seed-derived proanthocyanidins. The study provides fundamental information to viticulturists and winemakers on how to manage red wine grape low molecular weight phenolics and polymeric proanthocyanidin composition in a hot climate.
Harnessing Genetic Variation in Leaf Angle to Increase Productivity of Sorghum bicolor
Truong, Sandra K.; McCormick, Ryan F.; Rooney, William L.; Mullet, John E.
2015-01-01
The efficiency with which a plant intercepts solar radiation is determined primarily by its architecture. Understanding the genetic regulation of plant architecture and how changes in architecture affect performance can be used to improve plant productivity. Leaf inclination angle, the angle at which a leaf emerges with respect to the stem, is a feature of plant architecture that influences how a plant canopy intercepts solar radiation. Here we identify extensive genetic variation for leaf inclination angle in the crop plant Sorghum bicolor, a C4 grass species used for the production of grain, forage, and bioenergy. Multiple genetic loci that regulate leaf inclination angle were identified in recombinant inbred line populations of grain and bioenergy sorghum. Alleles of sorghum dwarf-3, a gene encoding a P-glycoprotein involved in polar auxin transport, are shown to change leaf inclination angle by up to 34° (0.59 rad). The impact of heritable variation in leaf inclination angle on light interception in sorghum canopies was assessed using functional-structural plant models and field experiments. Smaller leaf inclination angles caused solar radiation to penetrate deeper into the canopy, and the resulting redistribution of light is predicted to increase the biomass yield potential of bioenergy sorghum by at least 3%. These results show that sorghum leaf angle is a heritable trait regulated by multiple loci and that genetic variation in leaf angle can be used to modify plant architecture to improve sorghum crop performance. PMID:26323882
Kebrom, Tesfamichael H.; Mullet, John E.
2014-12-12
Shoot branches or tillers develop from axillary buds. The dormancy versus outgrowth fates of buds depends on genetic, environmental and hormonal signals. Defoliation inhibits bud outgrowth indicating the role of leaf-derived metabolic factors such as sucrose in bud outgrowth. In this study, the sensitivity of bud outgrowth to selective defoliation was investigated. At 6 d after planting (6 DAP), the first two leaves of sorghum were fully expanded and the third was partially emerged. Therefore, the leaves were selectively defoliated at 6 DAP and the length of the bud in the first leaf axil was measured at 8 DAP. Budmore » outgrowth was inhibited by defoliation of only 2 cm from the tip of the second leaf blade. The expression of dormancy and sucrose-starvation marker genes was up-regulated and cell cycle and sucrose-inducible genes was down-regulated during the first 24 h postdefoliation of the second leaf.At 48 h, the expression of these genes was similar to controls as the defoliated plant recovers. Our results demonstrate that small changes in photosynthetic leaf area affect the propensity of tiller buds for outgrowth. Therefore, variation in leaf area and photosynthetic activity should be included when integrating sucrose into models of shoot branching.« less
NASA Astrophysics Data System (ADS)
Xu, Jun; Papanikolaou, Nikos; Shi, Chengyu; Jiang, Steve B.
2009-08-01
Synchronized moving aperture radiation therapy (SMART) has been proposed to account for tumor motions during radiotherapy in prior work. The basic idea of SMART is to synchronize the moving radiation beam aperture formed by a dynamic multileaf collimator (DMLC) with the tumor motion induced by respiration. In this paper, a two-dimensional (2D) superimposing leaf sequencing method is presented for SMART. A leaf sequence optimization strategy was generated to assure the SMART delivery under realistic delivery conditions. The study of delivery performance using the Varian LINAC and the Millennium DMLC showed that clinical factors such as collimator angle, dose rate, initial phase and machine tolerance affect the delivery accuracy and efficiency. An in-house leaf sequencing software was developed to implement the 2D superimposing leaf sequencing method and optimize the motion-corrected leaf sequence under realistic clinical conditions. The analysis of dynamic log (Dynalog) files showed that optimization of the leaf sequence for various clinical factors can avoid beam hold-offs which break the synchronization of SMART and fail the SMART dose delivery. Through comparison between the simulated delivered fluence map and the planed fluence map, it was shown that the motion-corrected leaf sequence can greatly reduce the dose error.
Xu, Jun; Papanikolaou, Nikos; Shi, Chengyu; Jiang, Steve B
2009-08-21
Synchronized moving aperture radiation therapy (SMART) has been proposed to account for tumor motions during radiotherapy in prior work. The basic idea of SMART is to synchronize the moving radiation beam aperture formed by a dynamic multileaf collimator (DMLC) with the tumor motion induced by respiration. In this paper, a two-dimensional (2D) superimposing leaf sequencing method is presented for SMART. A leaf sequence optimization strategy was generated to assure the SMART delivery under realistic delivery conditions. The study of delivery performance using the Varian LINAC and the Millennium DMLC showed that clinical factors such as collimator angle, dose rate, initial phase and machine tolerance affect the delivery accuracy and efficiency. An in-house leaf sequencing software was developed to implement the 2D superimposing leaf sequencing method and optimize the motion-corrected leaf sequence under realistic clinical conditions. The analysis of dynamic log (Dynalog) files showed that optimization of the leaf sequence for various clinical factors can avoid beam hold-offs which break the synchronization of SMART and fail the SMART dose delivery. Through comparison between the simulated delivered fluence map and the planed fluence map, it was shown that the motion-corrected leaf sequence can greatly reduce the dose error.
Leaf traits within communities: context may affect the mapping of traits to function.
Funk, Jennifer L; Cornwell, William K
2013-09-01
The leaf economics spectrum (LES) has revolutionized the way many ecologists think about quantifying plant ecological trade-offs. In particular, the LES has connected a clear functional trade-off (long-lived leaves with slow carbon capture vs. short-lived leaves with fast carbon capture) to a handful of easily measured leaf traits. Building on this work, community ecologists are now able to quickly assess species carbon-capture strategies, which may have implications for community-level patterns such as competition or succession. However, there are a number of steps in this logic that require careful examination, and a potential danger arises when interpreting leaf-trait variation among species within communities where trait relationships are weak. Using data from 22 diverse communities, we show that relationships among three common functional traits (photosynthetic rate, leaf nitrogen concentration per mass, leaf mass per area) are weak in communities with low variation in leaf life span (LLS), especially communities dominated by herbaceous or deciduous woody species. However, globally there are few LLS data sets for communities dominated by herbaceous or deciduous species, and more data are needed to confirm this pattern. The context-dependent nature of trait relationships at the community level suggests that leaf-trait variation within communities, especially those dominated by herbaceous and deciduous woody species, should be interpreted with caution.
Predation-related odours reduce oviposition in a herbivorous mite.
Choh, Yasuyuki; Uefune, Masayoshi; Takabayashi, Junji
2010-01-01
When adult females of the herbivorous mite, Tetranychus urticae, were exposed to the predatory mite, Phytoseiulus persimilis, they laid fewer eggs than females that had not been exposed to P. persimilis when transferred onto a new leaf patch. However, when T. urticae females were exposed to either products of P. persimilis or artificially damaged conspecific eggs on a leaf patch, the number of T. urticae eggs on a new leaf patch did not differ significantly from the control. The reduced oviposition was neither due to the feeding activity on the leaf patch with P. persimilis nor to that on the new leaf patch. There was also no significant difference between the number of T. urticae eggs produced on a new leaf patch following exposure to the odours of a neighbouring leaf patch where there had previously been either P. persimilis or T. urticae adults. However, female T. urticae that had been exposed to odours from neighbouring leaf patches on which both T. urticae and P. persimilis had been placed produced significantly fewer eggs on a new leaf patch than those that had not been exposed to such odours. Neither odours from neighbouring intact leaf patches on which T. urticae eggs were preyed on by P. persimilis, nor odours from a neighbouring Parafilm patch on which T. urticae was preyed on by P. persimilis affected the oviposition of T. urticae. These data suggest that the presence of T. urticae, P. persimilis and a leaf patch are needed for the emission of odours to reduce oviposition in T. urticae.
USDA-ARS?s Scientific Manuscript database
Comparison of spinach (Spinacia oleracea L.) cultivars Lazio and Samish grown during the summer solstice in the sub-arctic versus the winter solstice in the sub-tropics provided insight into interactions between plant environment (day length, light intensity, ambient temperatures), cultivar and leaf...
Ebba K. Peterson; Niklaus J. Grünwald; Jennifer L. ParkeSoil
2017-01-01
Soilborne inoculum (infested leaf debris which has become incorporated into the soil) may be an important contributor to the persistence of the sudden oak death pathogen Phytophthora ramorum in recurrently positive nurseries. To initiate new epidemics, soilborne inoculum must not only be able to survive over time, but also be capable of...
Xiangming Xiao; Stephen Hagen; Qingyuan Zhang; Michael Keller; Berrien Moore III
2006-01-01
Leaf phenology of tropical evergreen forests affects carbon and water fluxes. In an earlier study of a seasonally moist evergreen tropical forest site in the Amazon basin, time series data of Enhanced Vegetation Index (EVI) from the VEGETATION and Moderate Resolution Imaging Spectroradiometer (MODIS) sensors showed an unexpected seasonal pattern, with higher EVI in the...
Wang, Y S; Liu, Z Y; Li, Y F; Zhang, Y; Yang, X F; Feng, H
2013-04-02
Artistic diversiform leaf color is an important agronomic trait that affects the market value of ornamental kale. In the present study, genetic analysis showed that a single-dominant gene, Re (red leaf), determines the red leaf trait in ornamental kale. An F2 population consisting of 500 individuals from the cross of a red leaf double-haploid line 'D05' with a white leaf double-haploid line 'D10' was analyzed for the red leaf trait. By combining bulked segregant analysis and sequence-related amplified polymorphism technology, we identified 3 markers linked to the Re/re locus. A genetic map of the Re locus was constructed using these sequence-related amplified polymorphism markers. Two of the markers, Me8Em4 and Me8Em17, were located on one side of Re/re at distances of 2.2 and 6.4 cM, whereas the other marker, Me9Em11, was located on the other side of Re/re at a distance of 3.7 cM. These markers could be helpful for the subsequent cloning of the red trait gene and marker-assisted selection in ornamental kale breeding programs.
Ni, Qinxue; Wang, Zhiqiang; Xu, Guangzhi; Gao, Qianxin; Yang, Dongdong; Morimatsu, Fumiki; Zhang, Youzuo
2013-01-01
Indocalamus latifolius (Keng) McClure leaf is a popular food material in East Asia due to its antioxidant and anticorrosive activities. To utilize it more effectively, we investigated the discrepancy of antioxidant activities and active compound content in Indocalamus latifolius leaf along with the altitude change. Total flavonoids, phenolics, titerpenoids and eight characteristic active constituents, i.e, orientin, isoorientin, vitexin, homovitexin, p-coumaric acid, chlorogenic acid, caffeic acid, and ferulic acid, were determined by UV-spectrophotometer and synchronous RP-HPLC, respectively. Antioxidant activity was measured using DPPH and FRAP methods. Our data showed that the content of TP and TF, DPPH radical scavenging ability and ferric reduction power of Indocalamus latifolius leaf changed as altitude altered, with the trends of decreasing gradually when lower than 700 m and then increasing to 1,000 m. Chlorogenic acid and orientin were the main characteristic compounds in Indocalamus latifolius leaf and were also affected by altitude. Our result indicated that higher altitude with an adverse environment is conducive to secondary metabolite accumulation for Indocalamus latifolius. It would provide a theoretical basis to regulate the leaf collection conditions in the industrial use of Indocalamus latifolius leaf.
Neurosurgery during the Bronze Age: a skull trepanation in 1900 BC Greece.
Papagrigorakis, Manolis J; Toulas, Panagiotis; Tsilivakos, Manolis G; Kousoulis, Antonis A; Skorda, Despoina; Orfanidis, George; Synodinos, Philippos N
2014-02-01
Paleoneurosurgery represents a comparatively new developing direction of neurosurgery dealing with archaeological skull and spine finds and studying their neurosurgical aspects. Trepanation of the cranial vault was a widespread surgical procedure in antiquity and the most convincing evidence of the ancient origin of neurosurgery. The present study considers a case of trepanation from the Middle Bronze Age Greece (1900-1600 B.C.). The skull under study belongs to skeletal material unearthed from Kirra, Delphi (Central Greece). Macroscopic examination and palpation, as well as three-dimensional computed tomography, were used in this study. There is osteological evidence that the skull belongs to a man who died at 30-35 years of age. The procedure of trepanation was performed on the right parietal bone. Both macroscopic and computed tomography evaluation demonstrate an intravital bone reaction at the edges of the aperture. Projected on the right surface of the brain, the trepanation is located on the level of the central groove. The small dimensions and the symmetrical shape of this hole give us an indication that it was made by a metal tool. We conclude that this paleopathological case provides valuable information about the condition of life and the pre-Hippocratic neurosurgical practice in Bronze Age Greece. Copyright © 2014 Elsevier Inc. All rights reserved.
NASA Astrophysics Data System (ADS)
Yi, Zhiran; Yang, Bin; Li, Guimiao; Liu, Jingquan; Chen, Xiang; Wang, Xiaolin; Yang, Chunsheng
2017-07-01
This letter presents a high performance bimorph piezoelectric MEMS harvester with bulk PZT thick films on both sides of a flexible thin beryllium-bronze substrate via bonding and thinning technologies. The upper and lower PZT layers are thinned down to about 53 μm and 76 μm, respectively, and a commercial beryllium bronze with the thickness of about 50 μm is used as the substrate. The effective volume of this device is 30.6 mm3. The harvester with a tungsten proof mass generated the close-circuit peak-to-peak voltage of 53.1 V, the output power of 0.979 mW, and the power density of 31.99 mW/cm3 with the matching load resistance of 360 kΩ at the applied acceleration amplitude of 3.5 g and the applied frequency of 77.2 Hz. Meanwhile, in order to evaluate the stability, the device was measured continuously under applied acceleration amplitudes of 1.0 g and 3.5 g for one hour and demonstrated a good stability. Then, the harvester was utilized to light up LEDs and about twenty-one serial LEDs were lighted up at resonance under an applied acceleration amplitude of 3.0 g.
NASA Astrophysics Data System (ADS)
Shizuma, Kiyoshi; Kajimoto, Tsuyoshi; Endo, Satoru; Matsugi, Kazuhiro; Arimatsu, Yui; Nojima, Hisashi
2017-09-01
Eight ancient bimetal swords held by Hiroshima University, Japan were analyzed non-destructively through γ-ray radiography and X-ray fluorescence (XRF). 137Cs and 60Co γ-ray irradiation sources were used to obtain transmission images of swords. A scanning radiography method using a 60Co γ-ray source was developed. XRF was used for qualitative elemental analysis of the swords. The presence of iron cores in the hilts of some swords had been observed and it was assumed that the cores were a ritual symbol or had a functional purpose. However, our work reveals that these swords were originally bronze-hilted iron swords and that the rusty blades were replaced with bronze blades to maintain the swords' commercial value as an antique. Consequently, the rest of the iron blade was left in the hilt as an iron tang. The junction of the blade and the guard was soldered and painted to match the patina color. XRF analysis clearly showed that the elemental Sn/Cu ratios of the blades and the hilts were different. These findings are useful for clarifying the later modifications of the swords and are important for interpreting Bronze Age and Iron Age history correctly.
Composition and microstructure of Roman metallic artefacts of Southwestern Iberian Peninsula
NASA Astrophysics Data System (ADS)
Valério, P.; Voráčová, E.; Silva, R. J. C.; Araújo, M. F.; Soares, A. M. M.; Arruda, A. M.; Pereira, C.
2015-10-01
The Roman invasion introduces new alloys and metallurgical practices in Iberian Peninsula. The southwestern end of this region has many evidences of connections with the Roman World, but there are no studies about the manufacture and use of copper-based artefacts during this period. Therefore, a set of about 20 ornaments, tools and small attachments recovered at the Roman sites of Monte Molião and Cidade das Rosas was studied by an analytical approach combining micro-EDXRF, optical microscopy, SEM-EDS and Vickers microhardness testing. The artefact composition shows a good correlation with function, namely pure copper for nails and rivets, low-tin bronze (2-6 wt% Sn) for basic tools, high-tin bronze (14 wt% Sn) for fibulae and high-lead bronze (19 wt% Pb) for a decorated jug handle. The manufacture also depends on function because most artefacts were subjected to thermomechanical processing, except the ornaments that would not benefit from post-casting work. Brass and gunmetal were only present in the site with a later chronology. A metallurgy visibly ruled by economical, aesthetical and technological concerns reinforces the evidences about the total integration of Southwestern Iberian Peninsula in the Roman World, but further studies will be essential to determine the evolution of copper-based alloys in Lusitania under Roman influence.
Chmura, D J; Modrzyński, J; Chmielarz, P; Tjoelker, M G
2017-03-01
Mechanisms of shade tolerance in tree seedlings, and thus growth in shade, may differ by leaf habit and vary with ontogeny following seed germination. To examine early responses of seedlings to shade in relation to morphological, physiological and biomass allocation traits, we compared seedlings of 10 temperate species, varying in their leaf habit (broadleaved versus needle-leaved) and observed tolerance to shade, when growing in two contrasting light treatments - open (about 20% of full sunlight) and shade (about 5% of full sunlight). We analyzed biomass allocation and its response to shade using allometric relationships. We also measured leaf gas exchange rates and leaf N in the two light treatments. Compared to the open treatment, shading significantly increased traits typically associated with high relative growth rate (RGR) - leaf area ratio (LAR), specific leaf area (SLA), and allocation of biomass into leaves, and reduced seedling mass and allocation to roots, and net assimilation rate (NAR). Interestingly, RGR was not affected by light treatment, likely because of morphological and physiological adjustments in shaded plants that offset reductions of in situ net assimilation of carbon in shade. Leaf area-based rates of light-saturated leaf gas exchange differed among species groups, but not between light treatments, as leaf N concentration increased in concert with increased SLA in shade. We found little evidence to support the hypothesis of a increased plasticity of broadleaved species compared to needle-leaved conifers in response to shade. However, an expectation of higher plasticity in shade-intolerant species than in shade-tolerant ones, and in leaf and plant morphology than in biomass allocation was supported across species of contrasting leaf habit. © 2016 German Botanical Society and The Royal Botanical Society of the Netherlands.
Rezende, Renan de Souza; Gonçalves Júnior, José Francisco; Lopes, Aline; Piedade, Maria Teresa Fernandez; Cavalcante, Heloide de Lima; Hamada, Neusa
2017-01-01
Climate change may affect the chemical composition of riparian leaf litter and, aquatic organisms and, consequently, leaf breakdown. We evaluated the effects of different scenarios combining increased temperature and carbon dioxide (CO2) on leaf detritus of Hevea spruceana (Benth) Müll. and decomposers (insect shredders and microorganisms). We hypothesized that simulated climate change (warming and elevated CO2) would: i) decrease leaf-litter quality, ii) decrease survival and leaf breakdown by shredders, and iii) increase microbial leaf breakdown and fungal biomass. We performed the experiment in four microcosm chambers that simulated air temperature and CO2 changes in relation to a real-time control tracking current conditions in Manaus, Amazonas, Brazil. The experiment lasted seven days. During the experiment mean air temperature and CO2 concentration ranged from 26.96 ± 0.98ºC and 537.86 ± 18.36 ppmv in the control to 31.75 ± 0.50ºC and 1636.96 ± 17.99 ppmv in the extreme chamber, respectively. However, phosphorus concentration in the leaf litter decreased with warming and elevated CO2. Leaf quality (percentage of carbon, nitrogen, phosphorus, cellulose and lignin) was not influenced by soil flooding. Fungal biomass and microbial leaf breakdown were positively influenced by temperature and CO2 increase and reached their highest values in the intermediate condition. Both total and shredder leaf breakdown, and shredder survival rate were similar among all climatic conditions. Thus, low leaf-litter quality due to climate change and higher leaf breakdown under intermediate conditions may indicate an increase of riparian metabolism due to temperature and CO2 increase, highlighting the risk (e.g., decreased productivity) of global warming for tropical streams. PMID:29190723
How do leaf veins influence the worldwide leaf economic spectrum? Review and synthesis.
Sack, Lawren; Scoffoni, Christine; John, Grace P; Poorter, Hendrik; Mason, Chase M; Mendez-Alonzo, Rodrigo; Donovan, Lisa A
2013-10-01
Leaf vein traits are implicated in the determination of gas exchange rates and plant performance. These traits are increasingly considered as causal factors affecting the 'leaf economic spectrum' (LES), which includes the light-saturated rate of photosynthesis, dark respiration, foliar nitrogen concentration, leaf dry mass per area (LMA) and leaf longevity. This article reviews the support for two contrasting hypotheses regarding a key vein trait, vein length per unit leaf area (VLA). Recently, Blonder et al. (2011, 2013) proposed that vein traits, including VLA, can be described as the 'origin' of the LES by structurally determining LMA and leaf thickness, and thereby vein traits would predict LES traits according to specific equations. Careful re-examination of leaf anatomy, published datasets, and a newly compiled global database for diverse species did not support the 'vein origin' hypothesis, and moreover showed that the apparent power of those equations to predict LES traits arose from circularity. This review provides a 'flux trait network' hypothesis for the effects of vein traits on the LES and on plant performance, based on a synthesis of the previous literature. According to this hypothesis, VLA, while virtually independent of LMA, strongly influences hydraulic conductance, and thus stomatal conductance and photosynthetic rate. We also review (i) the specific physiological roles of VLA; (ii) the role of leaf major veins in influencing LES traits; and (iii) the role of VLA in determining photosynthetic rate per leaf dry mass and plant relative growth rate. A clear understanding of leaf vein traits provides a new perspective on plant function independently of the LES and can enhance the ability to explain and predict whole plant performance under dynamic conditions, with applications towards breeding improved crop varieties.
Source of lead-210 and polonium-210 in tobacco.
Tso, T C; Harley, N; Alexander, L T
1966-08-19
Test plants were grown within a chamber enriched with radon-222 in the atmosphere, in tobacco fields with different sources of phosphate-containing fertilizer, and in culture containing lead-210 in the nutrient solution. Harvested leaves were subjected to three curing conditions. The major portion of the lead-210 in the plant was probably absorbed through the roots. Airborne radon 222 and its daughters contributed much less to the plant's content of lead-210 and of polonium-210. The stage of leaf development and the methods used to cure the leaf affected the final amount of polonium-210 in tobacco leaf.
USDA-ARS?s Scientific Manuscript database
Huanglongbing (HLB)-affected citrus often display zinc deficiency symptoms. In this study, supplemental zinc was applied to citrus to determine its effect on Candidatus Liberibacter asiaticus (Las) titer, HLB symptoms, and leaf microbiome. HLB-affected citrus were treated with various amounts of zi...
NASA Astrophysics Data System (ADS)
Kalayci, Tuna
This study investigates the relationship between rainfall variation and rain-fed agricultural production in Upper Mesopotamia with a specific focus on Early Bronze Age urban settlements. In return, the variation in production is used to explore stability of urban settlement systems. The organization of the flow of agricultural goods is the key to sustaining the total settlement system. The vulnerability of a settlement system increases due to the increased demand for more output from agricultural lands. This demand is the key for the success of urbanization project. However, without estimating how many foodstuffs were available at the end of a production cycle, further discussions on the forces that shaped and sustained urban settlement systems will be lacking. While large scale fluctuations in the flow of agricultural products between settlements are not the only determinants of hierarchical structures, the total available agricultural yield for each urban settlement in a hierarchy must have influenced settlement relations. As for the methodology, first, Early Bronze Age precipitation levels are estimated by using modern day associations between the eastern Mediterranean coastal areas and the inner regions of Upper Mesopotamia. Next, these levels are integrated into a remote-sensing based biological growth model. Also, a CORONA satellite imagery based archaeological survey is conducted in order to map the Early Bronze Age settlement system in its entirety as well as the ancient markers of agricultural intensification. Finally, ancient agricultural production landscapes are modeled in a GIS. The study takes a critical position towards the traditionally held assumption that large urban settlements (cities) in Upper Mesopotamia were in a state of constant demand for food. The results from this study also suggest that when variations in ancient precipitation levels are translated into the variations in production levels, the impact of climatic aridification on ancient settlement systems becomes less visible in the archaeological record.
DOE Office of Scientific and Technical Information (OSTI.GOV)
Huang, Weijiu; Chongqing Municipal Key Laboratory of Institutions of Higher Education for Mould Technology, Chongqing University of Technology, Chongqing 400054; Chai, Linjiang, E-mail: chailinjiang@cqut.edu.cn
2016-04-15
Specimens cut from a rolled tin bronze sheet were annealed at 400–800 °C for 1 h and evolution of their microstructures was then characterized in details by electron channeling contrast imaging and electron backscatter diffraction techniques. Particularly, statistics on special boundaries (SBs) with Σ ≤ 29 and network connectivity of random high angle boundaries (HABs) in the annealed specimens were examined to probe optimization potentials of grain boundary character distribution (GBCD) for this material. Results show that the deformed microstructure in the as-received material begins to be recrystallized when the annealing temperature increase to 500 °C and average grain sizesmore » surge with further increasing temperatures. As a result of the recrystallization, a large number of annealing twins (with Σ3 misorientation) are produced, leading to remarkably increased fractions of SBs (f{sub SBs}). Thanks to preexisting dense low angle boundaries, the majority of SBs in the 500 °C specimen with only partial recrystallization are Σ3{sub ic} (incoherent) boundaries, which effectively disrupt connectivity of random HABs network. Although the f{sub SBs} can be further increased (up to 72.5%) in specimens with full recrystallization (at higher temperatures), the Σ3{sub ic} boundaries would be replaced to some extent by Σ3{sub c} (coherent) boundaries which do not contribute directly to optimizing the GBCD. This work should be able to provide clear suggestions on applying the concept of grain boundary engineering to tin bronze alloys. - Highlights: • The rolled tin bronze begins to be recrystallized as temperature increases to 500 °C. • A lot of SBs are produced after recrystallization and the highest f{sub SBs} is 72.5%. • Partially recrystallized specimen has the optimum GBCD due to more Σ3{sub ic} boundaries. • The Σ3{sub ic} boundaries are replaced by Σ3{sub c} boundaries after full recrystallization.« less
Matus, José Tomás; Loyola, Rodrigo; Vega, Andrea; Peña-Neira, Alvaro; Bordeu, Edmundo; Arce-Johnson, Patricio; Alcalde, José Antonio
2009-01-01
Anthocyanins, flavan-3-ols, and flavonols are the three major classes of flavonoid compounds found in grape berry tissues. Several viticultural practices increase flavonoid content in the fruit, but the underlying genetic mechanisms responsible for these changes have not been completely deciphered. The impact of post-veraison sunlight exposure on anthocyanin and flavonol accumulation in grape berry skin and its relation to the expression of different transcriptional regulators known to be involved in flavonoid synthesis was studied. Treatments consisting of removing or moving aside the basal leaves which shade berry clusters were applied. Shading did not affect sugar accumulation or gene expression of HEXOSE TRANSPORTER 1, although in the leaf removal treatment, these events were retarded during the first weeks of ripening. Flavonols were the most drastically reduced flavonoids following shading and leaf removal treatments, related to the reduced expression of FLAVONOL SYNTHASE 4 and its putative transcriptional regulator MYB12. Anthocyanin accumulation and the expression of CHS2, LDOX, OMT, UFGT, MYBA1, and MYB5a genes were also affected. Other regulatory genes were less affected or not affected at all by these treatments. Non-transcriptional control mechanisms for flavonoid synthesis are also suggested, especially during the initial stages of ripening. Although berries from the leaf removal treatment received more light than shaded fruits, malvidin-3-glucoside and total flavonol content was reduced compared with the treatment without leaf removal. This work reveals that flavonol-related gene expression responds rapidly to field changes in light levels, as shown by the treatment in which shaded fruits were exposed to light in the late stages of ripening. Taken together, this study establishes MYB-specific responsiveness for the effect of sun exposure and sugar transport on flavonoid synthesis. PMID:19129169
Matus, José Tomás; Loyola, Rodrigo; Vega, Andrea; Peña-Neira, Alvaro; Bordeu, Edmundo; Arce-Johnson, Patricio; Alcalde, José Antonio
2009-01-01
Anthocyanins, flavan-3-ols, and flavonols are the three major classes of flavonoid compounds found in grape berry tissues. Several viticultural practices increase flavonoid content in the fruit, but the underlying genetic mechanisms responsible for these changes have not been completely deciphered. The impact of post-veraison sunlight exposure on anthocyanin and flavonol accumulation in grape berry skin and its relation to the expression of different transcriptional regulators known to be involved in flavonoid synthesis was studied. Treatments consisting of removing or moving aside the basal leaves which shade berry clusters were applied. Shading did not affect sugar accumulation or gene expression of HEXOSE TRANSPORTER 1, although in the leaf removal treatment, these events were retarded during the first weeks of ripening. Flavonols were the most drastically reduced flavonoids following shading and leaf removal treatments, related to the reduced expression of FLAVONOL SYNTHASE 4 and its putative transcriptional regulator MYB12. Anthocyanin accumulation and the expression of CHS2, LDOX, OMT, UFGT, MYBA1, and MYB5a genes were also affected. Other regulatory genes were less affected or not affected at all by these treatments. Non-transcriptional control mechanisms for flavonoid synthesis are also suggested, especially during the initial stages of ripening. Although berries from the leaf removal treatment received more light than shaded fruits, malvidin-3-glucoside and total flavonol content was reduced compared with the treatment without leaf removal. This work reveals that flavonol-related gene expression responds rapidly to field changes in light levels, as shown by the treatment in which shaded fruits were exposed to light in the late stages of ripening. Taken together, this study establishes MYB-specific responsiveness for the effect of sun exposure and sugar transport on flavonoid synthesis.
NASA Astrophysics Data System (ADS)
Zhao, Wenqiang; Reich, Peter B.; Yu, Qiannan; Zhao, Ning; Yin, Chunying; Zhao, Chunzhang; Li, Dandan; Hu, Jun; Li, Ting; Yin, Huajun; Liu, Qing
2018-04-01
Understanding leaf stoichiometric patterns is crucial for improving predictions of plant responses to environmental changes. Leaf stoichiometry of terrestrial ecosystems has been widely investigated along latitudinal and longitudinal gradients. However, very little is known about the vertical distribution of leaf C : N : P and the relative effects of environmental parameters, especially for shrubs. Here, we analyzed the shrub leaf C, N and P patterns in 125 mountainous sites over an extensive altitudinal gradient (523-4685 m) on the Tibetan Plateau. Results showed that the shrub leaf C and C : N were 7.3-47.5 % higher than those of other regional and global flora, whereas the leaf N and N : P were 10.2-75.8 % lower. Leaf C increased with rising altitude and decreasing temperature, supporting the physiological acclimation mechanism that high leaf C (e.g., alpine or evergreen shrub) could balance the cell osmotic pressure and resist freezing. The largest leaf N and high leaf P occurred in valley region (altitude 1500 m), likely due to the large nutrient leaching from higher elevations, faster litter decomposition and nutrient resorption ability of deciduous broadleaf shrub. Leaf N : P ratio further indicated increasing N limitation at higher altitudes. Interestingly, drought severity was the only climatic factor positively correlated with leaf N and P, which was more appropriate for evaluating the impact of water status than precipitation. Among the shrub ecosystem and functional types (alpine, subalpine, montane, valley, evergreen, deciduous, broadleaf, and conifer), their leaf element contents and responses to environments were remarkably different. Shrub type was the largest contributor to the total variations in leaf stoichiometry, while climate indirectly affected the leaf C : N : P via its interactive effects on shrub type or soil. Collectively, the large heterogeneity in shrub type was the most important factor explaining the overall leaf C : N : P variations, despite the broad climate gradient on the plateau. Temperature and drought induced shifts in shrub type distribution will influence the nutrient accumulation in mountainous shrubs.
Native arbuscular mycorrhizal symbiosis alters foliar bacterial community composition.
Poosakkannu, Anbu; Nissinen, Riitta; Kytöviita, Minna-Maarit
2017-11-01
The effects of arbuscular mycorrhizal (AM) fungi on plant-associated microbes are poorly known. We tested the hypothesis that colonization by an AM fungus affects microbial species richness and microbial community composition of host plant tissues. We grew the grass, Deschampsia flexuosa in a greenhouse with or without the native AM fungus, Claroideoglomus etunicatum. We divided clonally produced tillers into two parts: one inoculated with AM fungus spores and one without AM fungus inoculation (non-mycorrhizal, NM). We characterized bacterial (16S rRNA gene) and fungal communities (internal transcribed spacer region) in surface-sterilized leaf and root plant compartments. AM fungus inoculation did not affect microbial species richness or diversity indices in leaves or roots, but the AM fungus inoculation significantly affected bacterial community composition in leaves. A total of three OTUs in leaves belonging to the phylum Firmicutes positively responded to the presence of the AM fungus in roots. Another six OTUs belonging to the Proteobacteria (Alpha, Beta, and Gamma) and Bacteroidetes were significantly more abundant in NM plants when compared to AM fungus-inoculated plants. Further, there was a significant correlation between plant dry weight and leaf microbial community compositional shift. Also, there was a significant correlation between leaf bacterial community compositional shift and foliar nitrogen content changes due to AM fungus inoculation. The results suggest that AM fungus colonization in roots has a profound effect on plant physiology that is reflected in leaf bacterial community composition.
Crous, Kristine Y; O'Sullivan, Odhran S; Zaragoza-Castells, Joana; Bloomfield, Keith J; Negrini, A Clarissa A; Meir, Patrick; Turnbull, Matthew H; Griffin, Kevin L; Atkin, Owen K
2017-08-01
Nitrogen (N) and phosphorus (P) have key roles in leaf metabolism, resulting in a strong coupling of chemical composition traits to metabolic rates in field-based studies. However, in such studies, it is difficult to disentangle the effects of nutrient supply per se on trait-trait relationships. Our study assessed how high and low N (5 mM and 0.4 mM, respectively) and P (1 mM and 2 μM, respectively) supply in 37 species from six plant functional types (PTFs) affected photosynthesis (A) and respiration (R) (in darkness and light) in a controlled environment. Low P supply increased scaling exponents (slopes) of area-based log-log A-N or R-N relationships when N supply was not limiting, whereas there was no P effect under low N supply. By contrast, scaling exponents of A-P and R-P relationships were altered by P and N supply. Neither R : A nor light inhibition of leaf R was affected by nutrient supply. Light inhibition was 26% across nutrient treatments; herbaceous species exhibited a lower degree of light inhibition than woody species. Because N and P supply modulates leaf trait-trait relationships, the next generation of terrestrial biosphere models may need to consider how limitations in N and P availability affect trait-trait relationships when predicting carbon exchange. © 2017 The Authors. New Phytologist © 2017 New Phytologist Trust.
Evaluation of transfer films of Salox M on 440C for HPOTP bearing cage applications, task 119
NASA Technical Reports Server (NTRS)
Barber, S. A.; Kannel, J. W.; Dufrane, K. F.
1986-01-01
The objective of the task was to evaluate the suitability of a bronze-filled polytetra fluoroethylene (Salox M) as the cage material in Space Shuttle Main Engine (SSME) high pressure oxygen turbo pump (HPOTP) bearings. The role of the cage pocket material will be to provide a transferred lubricating interface at the ball-race contact region. A series of experiments was conducted which involved block-on-ring tests (the block was the polytetrafluoroethylene (PTFE) -filled material and the ring was through-hardened 440C steel) and high speed traction tests of two 440C disks with one disk rubbed with a PTFE block to generate a transfer film. Measurements included post test visual observations of the condition of the 440C, wear rate measurements of the blocks, and traction measurements between the disks. It was observed that both Salox M and glass-filled PTFE (Armalon) transferred PTFE to 440C at cryogenic temperatures. Bronze is also transferred to uncoated 440C from the Salox M. At room temperature no PTFE transfer was observed in the high speed disk tests due to severe frictional heating, although bronze transfer still occurred with the bronze-PTFE Salox M material. Since the bearing will operate at cryogenic temperature, transfer films are very probable. Salox M wore slightly, although probably tolerably, more than glass-filled PTFE against 440C. However, Salox M is clearly less abrasive to 440C than is the glass-filled material. When the surface layer of PTFE is depleted from the glass-filled material, the glass fibers tend to seriously abrade the steel. This problem does not occur with Salox M. The surface roughness studies indicate that smooth balls are quite reasonable for transfer films. No significant difference in wear rates of the candidate cage materials was observed when the 440C surface finish was increased from 0.025 micron to 0.1 micron cla. At higher levels of roughness, the wear rate increased. Two surface coatings (MoS2 and TiN) were tested. The surface coatings tended to reduce the Salox M wear rate and reduce bronze transfer. PTFE transfer at cryogenic temperatures still occurred with the surface coatings. Two cage design concepts are presented utilizing Salox M material. One design involves a metal-reinforced Salox M cage, while the second design uses Salox M inserts in a metal structure.
Leaf water potentials measured with a pressure chamber.
Boyer, J S
1967-01-01
Leaf water potentials were estimated from the sum of the balancing pressure measured with a pressure chamber and the osmotic potential of the xylem sap in leafy shoots or leaves. When leaf water potentials in yew, rhododendron, and sunflower were compared with those measured with a thermocouple psychrometer known to indicate accurate values of leaf water potential, determinations were within +/- 2 bars of the psychrometer measurements with sunflower and yew. In rhododendron. water potentials measured with the pressure chamber plus xylem sap were 2.5 bars less negative to 4 bars more negative than psychrometer measurements.The discrepancies in the rhododendron measurements could be attributed, at least in part, to the filling of tissues other than xylem with xylem sap during measurements with the pressure chamber. It was concluded that, although stem characteristics may affect the measurements, pressure chamber determinations were sufficiently close to psychrometer measurements that the pressure chamber may be used for relative measurements of leaf water potentials, especially in sunflower and yew. For accurate determinations of leaf water potential, however, pressure chamber measurements must be calibrated with a thermocouple psychrometer.
Leaf Water Potentials Measured with a Pressure Chamber
Boyer, J. S.
1967-01-01
Leaf water potentials were estimated from the sum of the balancing pressure measured with a pressure chamber and the osmotic potential of the xylem sap in leafy shoots or leaves. When leaf water potentials in yew, rhododendron, and sunflower were compared with those measured with a thermocouple psychrometer known to indicate accurate values of leaf water potential, determinations were within ± 2 bars of the psychrometer measurements with sunflower and yew. In rhododendron. water potentials measured with the pressure chamber plus xylem sap were 2.5 bars less negative to 4 bars more negative than psychrometer measurements. The discrepancies in the rhododendron measurements could be attributed, at least in part, to the filling of tissues other than xylem with xylem sap during measurements with the pressure chamber. It was concluded that, although stem characteristics may affect the measurements, pressure chamber determinations were sufficiently close to psychrometer measurements that the pressure chamber may be used for relative measurements of leaf water potentials, especially in sunflower and yew. For accurate determinations of leaf water potential, however, pressure chamber measurements must be calibrated with a thermocouple psychrometer. PMID:16656476
NASA Astrophysics Data System (ADS)
Natyanun, S.; Unai, S.; Yu, L. D.; Tippawan, U.; Pussadee, N.
2017-09-01
This study was aimed at understanding elemental concentration distribution in local longan leaf for how the plant was affected by the environment or agricultural operation. The analysis applied the MeV-microbeam particle induced X-ray emission (PIXE) mapping technique using a home-developed tapered glass capillary microbeam system at Chiang Mai University. The microbeam was 2-MeV proton beam in 130 µm in diameter. The studying interest was in the difference in the elemental concentrations distributed between the leaf midrib and lamina areas. The micro proton beam analyzed the leaf sample across the leaf midrib edge to the leaf lamina area for total 9 data requisition spots. The resulting data were colored to form a 1D-map of the elemental concentration distribution. Seven dominant elements, Al, S, Cl, K, Ca, Sc and Fe, were identified, the first six of which were found having higher concentrations in the midrib area than in the lamina area, while the Fe concentration was in an opposite trend to that of the others.
DOE Office of Scientific and Technical Information (OSTI.GOV)
Blatt, C.R.
1982-01-01
Preplant applications of Borate -65 at 0.56, 1.12 and 2.24 kg B/ha were reflected in significant increases in soil and leaf B levels up to one year following boron application. After 2 cropping seasons soil B level did not reflect rate of applied B and Solubor was applied broadcast at 1.12, 2.24 and 4.48 kg B/ha in the spring of the 3rd cropping season. Soil and leaf B levels and leaf marginal necrosis increased compared with control plots at all rates of applied B at full bloom in the 3rd cropping season. Rate of applied B was reflected in significantmore » soil and leaf B increases one year following application. Fruit yields through four cropping seasons were not affected by any source of rate of applied B. A soil B range of 0.15-0.25 ppM and a leaf B range of 20-30 ppm will give optimum crop response from the Midway strawberry.« less
Lunar gravity affects leaf movement of Arabidopsis thaliana in the International Space Station.
Fisahn, Joachim; Klingelé, Emile; Barlow, Peter
2015-06-01
Cyclic leaf ascent and descent occur in synchrony and phase congruence with the lunisolar tidal force under a broad range of conditions. Digitized records of the vertical leaf movements of Arabidopsis thaliana were collected under space flight conditions in the International Space Station (ISS). Oscillations of leaf movements with periods of 45 and 90 min were found under light-adapted conditions, whereas in darkness, the periods were 45, 90, and 135 min. To demonstrate the close relationship between these oscillations and cyclical variations of the lunisolar gravitational force, we estimated the oscillations of the in-orbit lunisolar tide as they apply to the ISS, with the aid of the Etide software application. In general, in-orbit lunisolar gravitational profiles exhibited a periodicity of 45 min. Alignment of these in-orbit oscillations with the oscillations of Arabidopsis leaf movement revealed high degrees of synchrony and a congruence of phase. These data corroborate previous results which suggested a correlative relationship and a possible causal link between leaf movement rhythms obtained on ground and the rhythmic variation of the lunisolar tidal force.
Quantitative trait loci controlling leaf venation in Arabidopsis.
Rishmawi, Louai; Bühler, Jonas; Jaegle, Benjamin; Hülskamp, Martin; Koornneef, Maarten
2017-08-01
Leaf veins provide the mechanical support and are responsible for the transport of nutrients and water to the plant. High vein density is a prerequisite for plants to have C4 photosynthesis. We investigated the genetic variation and genetic architecture of leaf venation traits within the species Arabidopsis thaliana using natural variation. Leaf venation traits, including leaf vein density (LVD) were analysed in 66 worldwide accessions and 399 lines of the multi-parent advanced generation intercross population. It was shown that there is no correlation between LVD and photosynthesis parameters within A. thaliana. Association mapping was performed for LVD and identified 16 and 17 putative quantitative trait loci (QTLs) in the multi-parent advanced generation intercross and worldwide sets, respectively. There was no overlap between the identified QTLs suggesting that many genes can affect the traits. In addition, linkage mapping was performed using two biparental recombinant inbred line populations. Combining linkage and association mapping revealed seven candidate genes. For one of the candidate genes, RCI2c, we demonstrated its function in leaf venation patterning. © 2017 John Wiley & Sons Ltd.
Growth and photosynthesis of Japanese flowering cherry under simulated microgravity conditions
NASA Technical Reports Server (NTRS)
Sugano, Mami; Ino, Yoshio; Nakamura, Teruko
2002-01-01
The photosynthetic rate, the leaf characteristics related to photosynthesis, such as the chlorophyll content, chlorophyll a/b ratio and density of the stomata, the leaf area and the dry weight in seedlings of Japanese flowering cherry grown under normal gravity and simulated microgravity conditions were examined. No significant differences were found in the photosynthetic rates between the two conditions. Moreover, leaf characteristics such as the chlorophyll content, chlorophyll a/b ratio and density of the stomata in the seedlings grown under the simulated microgravity condition were not affected. However, the photosynthetic product of the whole seedling under the simulated microgravity condition increased compared with the control due to its leaf area increase. The results suggest that dynamic gravitational stimulus controls the partitioning of the products of photosynthesis.
ERIC Educational Resources Information Center
Hunt, Susan
1979-01-01
An exhibit designed for visually handicapped persons presented by the Kalamazoo (Michigan) Institute of Art included bronze sculptures and oil paintings from the institute's permanent collection. (CL)
Martínez-Ballesté, Andrea; Martorell, Carlos
2015-01-01
Traditional management practices are usually thought to be sustainable. The Maya manage Sabal (Arecaceae) palms in homegardens, using their leaves for thatching. The sustainability of such production systems depends on the long-term persistence of palm populations, whereas resource availability also depends on the number of leaves on individual palms. We examined how leaf harvest affects Sabal yapa and S. mexicana population growth rates (λ) and leaf production, comparing traditional and alternative harvest regimes in terms of sustainability and productivity. Demographic, harvest and leaf production data were recorded for three years in two homegardens. We used general integral projection models linked to leaf-production models to describe population dynamics and productivity. Harvest had no effect on S. yapa's vital rates or on λ, but it changed the growth rate of individuals of S. mexicana, with a negligible impact on λ. Homegardens affected λ values, reflecting the species' ecological affinities. S. mexicana, introduced from mesic forests, required watering and shade; therefore, its population declined rapidly in the homegarden that lacked both water and shade. The λ of the xerophilic S. yapa was slightly larger without watering than with watering. Palms usually compensated for leaf extraction, causing the number of leaves harvested per individual to increase with harvest intensity. Nevertheless, traditional management is relatively mild, allowing standing leaves to accumulate but reducing the homegarden's yield. Apparently, the Maya do not seek to maximize annual production but to ensure the availability of large numbers of leaves in homegardens. These leaves may then be used when the entire roof of a hut needs to be replaced every few years.
Martínez-Ballesté, Andrea; Martorell, Carlos
2015-01-01
Traditional management practices are usually thought to be sustainable. The Maya manage Sabal (Arecaceae) palms in homegardens, using their leaves for thatching. The sustainability of such production systems depends on the long-term persistence of palm populations, whereas resource availability also depends on the number of leaves on individual palms. We examined how leaf harvest affects Sabal yapa and S. mexicana population growth rates (λ) and leaf production, comparing traditional and alternative harvest regimes in terms of sustainability and productivity. Demographic, harvest and leaf production data were recorded for three years in two homegardens. We used general integral projection models linked to leaf-production models to describe population dynamics and productivity. Harvest had no effect on S. yapa’s vital rates or on λ, but it changed the growth rate of individuals of S. mexicana, with a negligible impact on λ. Homegardens affected λ values, reflecting the species’ ecological affinities. S. mexicana, introduced from mesic forests, required watering and shade; therefore, its population declined rapidly in the homegarden that lacked both water and shade. The λ of the xerophilic S. yapa was slightly larger without watering than with watering. Palms usually compensated for leaf extraction, causing the number of leaves harvested per individual to increase with harvest intensity. Nevertheless, traditional management is relatively mild, allowing standing leaves to accumulate but reducing the homegarden’s yield. Apparently, the Maya do not seek to maximize annual production but to ensure the availability of large numbers of leaves in homegardens. These leaves may then be used when the entire roof of a hut needs to be replaced every few years. PMID:25803029
Macarisin, Dumitru; Patel, Jitendra; Bauchan, Gary; Giron, Jorge A; Ravishankar, Sadhana
2013-11-01
Similar to phytopathogens, human bacterial pathogens have been shown to colonize the plant phylloplane. In addition to environmental factors, such as temperature, UV, relative humidity, etc., the plant cultivar and, specifically, the leaf blade morphological characteristics may affect the persistence of enteropathogens on leafy greens. This study was conducted to evaluate the effect of cultivar-dependent leaf topography and the role of strain phenotypic characteristics on Escherichia coli O157:H7 persistence on organic spinach. Spinach cultivars Emilia, Lazio, Space, and Waitiki were experimentally inoculated with the foodborne E. coli O157:H7 isolate EDL933 and its isogenic mutants deficient in cellulose, curli, or both curli and cellulose production. Leaves of 6-week-old plants were inoculated with 6.5 log CFU per leaf in a biosafety level 2 growth chamber. At 0, 1, 7, and 14 days, E. coli O157:H7 populations were determined by plating on selective medium and verified by laser scanning confocal microscopy. Leaf morphology (blade roughness and stoma density) was evaluated by low-temperature and variable-pressure scanning electron microscopy. E. coli O157:H7 persistence on spinach was significantly affected by cultivar and strain phenotypic characteristics, specifically, the expression of curli. Leaf blade roughness and stoma density influenced the persistence of E. coli O157:H7 on spinach. Cultivar Waitiki, which had the greatest leaf roughness, supported significantly higher E. coli O157:H7 populations than the other cultivars. These two morphological characteristics of spinach cultivars should be taken into consideration in developing intervention strategies to enhance the microbial safety of leafy greens.
Timing and duration of autumn leaf development in Sweden, a 4-year citizen science study
NASA Astrophysics Data System (ADS)
Bolmgren, Kjell; Langvall, Ola
2017-04-01
Phenology monitoring has traditionally focused on the start of phenological phases and the start of the growing season, especially when it comes to species-specific observations on the ground. The patterns of and the mechanisms behind the end of particular phases and the growing season itself are less studied and poorly understood. With a changing climate, the need to understand and predict effects on the length as well as on the end of phenological phases increase in importance, e.g. in relation to estimations of carbon budgets and validation of remote sensing data. Furthermore, different species may be affected in different ways by changing conditions. In this 4-year-study, tens of thousands of pupils in ages from 6 to 19 years old were involved in observing autumn leaf development of common deciduous tree species. Their observations were made near schools all over Sweden (55-68°N). Observations were made weekly between late August and early November and followed an image-based observation protocol, classifying autumn leaf development into five levels, from a summer-green (level 0) to a 100% autumn-colored (level 4) canopy. As expected, there was a general (negative) correlation between latitude and the start of leaf senescence (level 2; 1/3 autumn-colored canopy), but the correlation differed largely among years and between species. There was a week correlation between latitude and duration of the leaf senescence period, defined as the period between 1/3 (level 2) and 100% (level 4) of autumn-colored canopy. A delayed onset of the leaf senescence affected the duration of the leaf senescence period more strongly; One (1) day later start was correlated with a 5-day shorter period. Different species had different length of their senescence period, with oak (mainly Quercus robur) and birches (Betula pendula and B. pubescence) having on average a 50% longer period than trembling aspen (Populus tremula) and Norway maple (Acer platanoides).
Schollert, Michelle; Kivimäenpää, Minna; Michelsen, Anders; Blok, Daan; Rinnan, Riikka
2017-02-01
Climate change in the Arctic is projected to increase temperature, precipitation and snowfall. This may alter leaf anatomy and gas exchange either directly or indirectly. Our aim was to assess whether increased snow depth and warming modify leaf anatomy and affect biogenic volatile organic compound (BVOC) emissions and CO 2 exchange of the widespread arctic shrubs Betula nana and Empetrum nigrum ssp. hermaphroditum METHODS: Measurements were conducted in a full-factorial field experiment in Central West Greenland, with passive summer warming by open-top chambers and snow addition using snow fences. Leaf anatomy was assessed using light microscopy and scanning electron microscopy. BVOC emissions were measured using a dynamic enclosure system and collection of BVOCs into adsorbent cartridges analysed by gas chromatography-mass spectrometry. Carbon dioxide exchange was measured using an infrared gas analyser. Despite a later snowmelt and reduced photosynthesis for B. nana especially, no apparent delays in the BVOC emissions were observed in response to snow addition. Only a few effects of the treatments were seen for the BVOC emissions, with sesquiterpenes being the most responsive compound group. Snow addition affected leaf anatomy by increasing the glandular trichome density in B. nana and modifying the mesophyll of E. hermaphroditum The open-top chambers thickened the epidermis of B. nana, while increasing the glandular trichome density and reducing the palisade:spongy mesophyll ratio in E. hermaphroditum CONCLUSIONS: Leaf anatomy was modified by both treatments already after the first winter and we suggest links between leaf anatomy, CO 2 exchange and BVOC emissions. While warming is likely to reduce soil moisture, melt water from a deeper snow pack alleviates water stress in the early growing season. The study emphasizes the ecological importance of changes in winter precipitation in the Arctic, which can interact with climate-warming effects. © The Author 2017. Published by Oxford University Press on behalf of the Annals of Botany Company. All rights reserved. For Permissions, please email: journals.permissions@oup.com.
Schollert, Michelle; Kivimäenpää, Minna; Michelsen, Anders; Blok, Daan; Rinnan, Riikka
2017-01-01
Background and Aims Climate change in the Arctic is projected to increase temperature, precipitation and snowfall. This may alter leaf anatomy and gas exchange either directly or indirectly. Our aim was to assess whether increased snow depth and warming modify leaf anatomy and affect biogenic volatile organic compound (BVOC) emissions and CO2 exchange of the widespread arctic shrubs Betula nana and Empetrum nigrum ssp. hermaphroditum. Methods Measurements were conducted in a full-factorial field experiment in Central West Greenland, with passive summer warming by open-top chambers and snow addition using snow fences. Leaf anatomy was assessed using light microscopy and scanning electron microscopy. BVOC emissions were measured using a dynamic enclosure system and collection of BVOCs into adsorbent cartridges analysed by gas chromatography–mass spectrometry. Carbon dioxide exchange was measured using an infrared gas analyser. Key Results Despite a later snowmelt and reduced photosynthesis for B. nana especially, no apparent delays in the BVOC emissions were observed in response to snow addition. Only a few effects of the treatments were seen for the BVOC emissions, with sesquiterpenes being the most responsive compound group. Snow addition affected leaf anatomy by increasing the glandular trichome density in B. nana and modifying the mesophyll of E. hermaphroditum. The open-top chambers thickened the epidermis of B. nana, while increasing the glandular trichome density and reducing the palisade:spongy mesophyll ratio in E. hermaphroditum. Conclusions Leaf anatomy was modified by both treatments already after the first winter and we suggest links between leaf anatomy, CO2 exchange and BVOC emissions. While warming is likely to reduce soil moisture, melt water from a deeper snow pack alleviates water stress in the early growing season. The study emphasizes the ecological importance of changes in winter precipitation in the Arctic, which can interact with climate-warming effects. PMID:28064192
Christopher B. Allen; Rodney E. Will; Robert C. McGravey; David R. Coyle; Mark D. Coleman
2005-01-01
We investigated how water and nutrient availability affect radiation-use effeciency (e) and assessed leaf gas exchange as a possible mechanism for shifts in e. We measured aboveground net primary production (ANPP) and annual photosynthetically active radiation (PAR) capture to calculate e as well as leaf-level physiological variables (light-saturated net photosynthesis...
Ebba K. Peterson; Niklaus J. Grünwald; Jennifer L. Parke
2017-01-01
Soilborne inoculum (infested leaf debris which has become incorporated into the soil) may be an important contributor to the persistence of the sudden oak death pathogen Phytophthora ramorum in recurrently positive nurseries. To initiate new epidemics, soilborne inoculum must not only be able to survive over time, but also be capable of producing...
Long-term leaf fall mass from three watersheds on the Fernow Experimental Forest, West Virginia
Mary Beth Adams
2008-01-01
Foliar biomass may serve as an indicator of site productivity, and spatial and temporal changes can help us understand effects of important variables affecting productivity. Leaf litterfall mass is one way to estimate foliar biomass, and has been measured on three watersheds on the Fernow Experimental Forest in West Virginia for 19 years. These watersheds all contain...
Bacterial Leaf Scorch Affects New Jersey State Tree (Pest Alert)
USDA Forest Service; Northeastern Area, State and Private Forestry
2000-01-01
Recent surveys indicate that Bacterial Leaf Scorch (BLS) of northern red oak is widespread within New Jersey (see map) with many communities experiencing a high disease incidence. BLS is considered a threat not only to the state tree, northern red oak, but also to pin and scarlet oaks and other urban trees such as sycamore and elm. The disease in oaks has been observed...
Program to develop sprayed, plastically deformable compressor shroud seal materials
NASA Technical Reports Server (NTRS)
Schell, J. D.; Schell, J. D.
1980-01-01
A study of fundamental rub behavior for 10 dense, sprayed materials and eight current compressor clearance materials was conducted. A literature survey of a wide variety of metallurgical and thermophysical properties was conducted and correlated to rub behavior. Based on the results, the most promising dense rub material was Cu-9A1. Additional studies on the effects of porosity, incursion rate, blade solidity, and ambient temperature were carried out on aluminum bronze (Cu-9Al-1Fe) with and without a 515B Feltmetal underlayer. A further development effort was conducted to assess the property requirements of a porous, aluminum bronze, seal material. Strength, thermal cycle capabilities, erosion and oxidation resistance, machinability, and abradability at several porosity levels were examined.
NASA Technical Reports Server (NTRS)
Green, G.; Mattauch, R. J. (Inventor)
1983-01-01
A method is described for forming sharp tips on thin wires, in particular phosphor bronze wires of diameters such as one-thousandth inch used to contact micron size Schottky barrier diodes, which enables close control of tip shape and which avoids the use of highly toxic solutions. The method includes dipping an end of a phosphor bronze wire into a dilute solution of sulfamic acid and applying a current through the wire to electrochemically etch it. The humidity in the room is controlled to a level of less than 50%, and the voltage applied between the wire and another electrode in the solutions is a half wave rectified voltage. The current through the wire is monitored, and the process is stopped when the current falls to a predetermined low level.
Doebrich, Jeff
2009-01-01
Copper was one of the first metals ever extracted and used by humans, and it has made vital contributions to sustaining and improving society since the dawn of civilization. Copper was first used in coins and ornaments starting about 8000 B.C., and at about 5500 B.C., copper tools helped civilization emerge from the Stone Age. The discovery that copper alloyed with tin produces bronze marked the beginning of the Bronze Age at about 3000 B.C. Copper is easily stretched, molded, and shaped; is resistant to corrosion; and conducts heat and electricity efficiently. As a result, copper was important to early humans and continues to be a material of choice for a variety of domestic, industrial, and high-technology applications today.
Pressure dependence of the Peierls transition in the quasi two-dimensional purple bronze KMo 6O 17
NASA Astrophysics Data System (ADS)
Rötger, A.; Beille, J.; Laurant, J. M.; Schlenker, C.
1993-09-01
The electrical resistivity and the lattice parameters have been studied as a function of pressure on the quasi-twodimensional purple bronze KMo 6O 17 which shows a Peierls transition towards a commensurate charge density wave state. The Peierls temperature is found to be first slightly decreased for pressures smaller than 6 kbar, then strongly increased above. This increase is associated to an anomalous contraction of the lattice parameters in the plane of the layers. The corresponding large increase of the compressibility above 16 kbar at 300 K is associated to the pretransitional regime of the Peierls transition as a function of pressure. These results are attributed mainly to an improved nesting of the Fermi surface under pressure.
NASA Technical Reports Server (NTRS)
Richards, Hadley T.
1954-01-01
A turbine blade with a porous stainless-steel shell sintered to a supporting steel strut has been fabricated for tests at the NACA by Federal-Mogul Corporation under contract from the Bureau of Aeronautics, Department of the Navy. The apparent permeability of this blade, on the average, more nearly approaches the values specified by the NAGA than did two strut-supported bronze blades in a previous investigation. Random variations of permeability in the present blade are substantialy greater than those of the bronze blades, but projected improvements in certain phases of the fabrication process are expected to reduce these variations.
Hydrology, phenology and the USA National Phenology Network
Kish, George R.
2010-01-01
Phenology is the study of seasonally-recurring biological events (such as leaf-out, fruit production, and animal reproduction and migration) and how these events are influenced by environmental change. Phenological changes are some of the most sensitive biological indicators of climate change, and also affect nearly all aspects of ecosystem function. Spatially extensive patterns of phenological observations have been closely linked with climate variability. Phenology and hydrology are closely linked and affect one another across a variety of scales, from leaf intercellular spaces to the troposphere, and over periods of seconds to centuries. Ecosystem life cycles and diversity are also influenced by hydrologic processes such as floods and droughts. Therefore, understanding the relationships between hydrology and phenology is increasingly important in understanding how climate change affects biological and physical systems.
NASA Astrophysics Data System (ADS)
Kruijt, B.; Barton, C.; Rey, A.; Jarvis, P. G.
The 3-dimensional forest model MAESTRO was used to simulate daily and annual photosynthesis and transpiration fluxes of forest stands and the sensitivity of these fluxes to potential changes in atmospheric CO2 concentration ([CO2]), temperature, water stress and phenology. The effects of possible feed-backs from increased leaf area and limitations to leaf nutrition were simulated by imposing changes in leaf area and nitrogen content. Two different tree species were considered: Picea sitchensis (Bong.) Carr., a conifer with long needle longevity and large leaf area, and Betula pendula Roth., a broad-leaved deciduous species with an open canopy and small leaf area. Canopy photosynthetic production in trees was predicted to increase with atmospheric [CO2] and length of the growing season and to decrease with increased water stress. Associated increases in leaf area increased production further only in the B. pendula canopy, where the original leaf area was relatively small. Assumed limitations in N uptake affected B. pendula more than P. sitchensis. The effect of increased temperature was shown to depend on leaf area and nitrogen content. The different sensitivities of the two species were related to their very different canopy structure. Increased [CO2] reduced transpiration, but larger leaf area, early leaf growth, and higher temperature all led to increased water use. These effects were limited by feedbacks from soil water stress. The simulations suggest that, with the projected climate change, there is some increase in stand annual `water use efficiency', but the actual water losses to the atmosphere may not always decrease.
Santiago, Louis S; Kitajima, Kaoru; Wright, S Joseph; Mulkey, Stephen S
2004-05-01
We investigated leaf physiological traits of dominant canopy trees in four lowland Panamanian forests with contrasting mean annual precipitation (1,800, 2,300, 3,100 and 3,500 mm). There was near complete turn-over of dominant canopy tree species among sites, resulting in greater dominance of evergreen species with long-lived leaves as precipitation increased. Mean structural and physiological traits changed along this gradient as predicted by cost-benefit theories of leaf life span. Nitrogen content per unit mass (Nmass) and light- and CO2-saturated photosynthetic rates per unit mass (Pmass) of upper canopy leaves decreased with annual precipitation, and these changes were partially explained by increasing leaf thickness and decreasing specific leaf area (SLA). Comparison of 1,800 mm and 3,100 mm sites, where canopy access was available through the use of construction cranes, revealed an association among extended leaf longevity, greater structural defense, higher midday leaf water potential, and lower Pmass, Nmass, and SLA at wetter sites. Shorter leaf life spans and more enriched foliar delta15N values in drier sites suggest greater resorption and re-metabolism of leaf N in drier forest. Greater dominance of short-lived leaves with relatively high Pmass in drier sites reflects a strategy to maximize photosynthesis when water is available and to minimize water loss and respiration costs during rainless periods. Overall, our study links coordinated change in leaf functional traits that affect productivity and nutrient cycling to seasonality in lowland tropical forests. Copyright 2004 Springer-Verlag
Responses of rubber leaf phenology to climatic variations in Southwest China
NASA Astrophysics Data System (ADS)
Zhai, De-Li; Yu, Haiying; Chen, Si-Chong; Ranjitkar, Sailesh; Xu, Jianchu
2017-11-01
The phenology of rubber trees (Hevea brasiliensis) could be influenced by meteorological factors and exhibits significant changes under different geoclimates. In the sub-optimal environment in Xishuangbanna, rubber trees undergo lengthy periods of defoliation and refoliation. The timing of refoliation from budburst to leaf aging could be affected by powdery mildew disease (Oidium heveae), which negatively impacts seed and latex production. Rubber trees are most susceptible to powdery mildew disease at the copper and leaf changing stages. Understanding and predicting leaf phenology of rubber trees are helpful to develop effective means of controlling the disease. This research investigated the effect of several meteorological factors on different leaf phenological stages in a sub-optimal environment for rubber cultivation in Jinghong, Yunnan in Southwest China. Partial least square regression was used to quantify the relationship between meteorological factors and recorded rubber phenologies from 2003 to 2011. Minimum temperature in December was found to be the critical factor for the leaf phenology development of rubber trees. Comparing the delayed effects of minimum temperature, the maximum temperature, diurnal temperature range, and sunshine hours were found to advancing leaf phenologies. A comparatively lower minimum temperature in December would facilitate the advancing of leaf phenologies of rubber trees. Higher levels of precipitation in February delayed the light green and the entire process of leaf aging. Delayed leaf phenology was found to be related to severe rubber powdery mildew disease. These results were used to build predictive models that could be applied to early warning systems of rubber powdery mildew disease.
Roa-Fuentes, Lilia L; Templer, Pamela H; Campo, Julio
2015-10-01
Leaf traits are closely associated with nutrient use by plants and can be utilized as a proxy for nutrient cycling processes. However, open questions remain, in particular regarding the variability of leaf traits within and across seasonally dry tropical forests. To address this, we considered six leaf traits (specific area, thickness, dry matter content, N content, P content and natural abundance (15)N) of four co-occurring tree species (two that are not associated with N2-fixing bacteria and two that are associated with N2-fixing bacteria) and net N mineralization rates and inorganic N concentrations along a precipitation gradient (537-1036 mm per year) in the Yucatan Peninsula, Mexico. Specifically we sought to test the hypothesis that leaf traits of dominant plant species shift along a precipitation gradient, but are affected by soil N cycling. Although variation among different species within each site explains some leaf trait variation, there is also a high level of variability across sites, suggesting that factors other than precipitation regime more strongly influence leaf traits. Principal component analyses indicated that across sites and tree species, covariation in leaf traits is an indicator of soil N availability. Patterns of natural abundance (15)N in foliage and foliage minus soil suggest that variation in precipitation regime drives a shift in plant N acquisition and the openness of the N cycle. Overall, our study shows that both plant species and site are important determinants of leaf traits, and that the leaf trait spectrum is correlated with soil N cycling.
Roy, Somnath; Muraleedharan, Narayanannair; Mukhopadhyay, Ananda
2014-08-01
Oligonychus coffeae Nietner (Acari: Tetranychidae), the red spider mite (RSM), is a major pest of tea (Camellia sinensis) in most tea-producing countries. Nymphs and adults of RSM lacerate cells, producing minute characteristic reddish brown marks on the upper surface of mature leaves, which turn red in severe cases of infestation, resulting in crop loss. The pest is present on tea all the year round, although numbers vary depending on season. Their number increases as the weather warms up and decreases markedly once rains set in. Under optimal conditions there may be 22 overlapping generations in a year. Parthenogenesis is known to occur; consequently, all mite stages can be found at a given time. Their infestation is mainly confined to the upper surface of the mature leaves and could readily be identified by the bronzing of the leaf. There are several naturally occurring insect predators, such as coccinellid and staphylinid larvae, lacewing larvae, and mite predators, most importantly species of the families Phytoseiidae and Stigmaeidae. Integrated management has been adopted to control this mite pest, involving cultural, mechanical, physical, biological and chemical methods. This review collates the most important works carried out on biology, ecology and management of O. coffeae. Also the scope of future studies for better management of this regular mite pest of tea is discussed.
Influence of vesicular arbuscular mycorrhizae and leaf age on net gas exchange of citrus leaves.
Syvertsen, J P; Graham, J H
1990-11-01
The purpose of this study was to test the hypothesis that vesicular arbuscular mycorrhizal (VAM) fungi affect net assimilation of CO(2) (A) of different-aged citrus leaves independent of mineral nutrition effects of mycorrhizae. Citrus aurantium L., sour orange plants were grown for 6 months in a sandy soil low in phosphorus that was either infested with the VAM fungus, Glomus intraradices Schenck & Smith, or fertilized with additional phosphorus and left nonmycorrhizal (NM). Net CO(2) assimilation, stomatal conductance, water use efficiency, and mineral nutrient status for expanding, recently expanded, and mature leaves were evaluated as well as plant size and relative growth rate of leaves. Nutrient status and net gas exchange varied with leaf age. G. intraradices-inoculated plants had well-established colonization (79% of root length) and were comparable in relative growth rate and size at final harvest with NM plants. Leaf mineral concentrations were generally the same for VAM and NM plants except for nitrogen. Although leaf nitrogen was apparently sufficient for high rates of A, VAM plants did have higher nitrogen concentrations than NM at the time of gas exchange measurements. G. intraradices had no effect on A, stomatal conductance, or water use efficiency, irrespective of leaf age. These results show that well-established VAM colonization does not affect net gas exchange of citrus plants that are comparable in size, growth rate, and nutritional status with NM plants.
Autumn leaf subsidies influence spring dynamics of freshwater plankton communities.
Fey, Samuel B; Mertens, Andrew N; Cottingham, Kathryn L
2015-07-01
While ecologists primarily focus on the immediate impact of ecological subsidies, understanding the importance of ecological subsidies requires quantifying the long-term temporal dynamics of subsidies on recipient ecosystems. Deciduous leaf litter transferred from terrestrial to aquatic ecosystems exerts both immediate and lasting effects on stream food webs. Recently, deciduous leaf additions have also been shown to be important subsidies for planktonic food webs in ponds during autumn; however, the inter-seasonal effects of autumn leaf subsidies on planktonic food webs have not been studied. We hypothesized that autumn leaf drop will affect the spring dynamics of freshwater pond food webs by altering the availability of resources, water transparency, and the metabolic state of ponds. We created leaf-added and no-leaf-added field mesocosms in autumn 2012, allowed mesocosms to ice-over for the winter, and began sampling the physical, chemical, and biological properties of mesocosms immediately following ice-off in spring 2013. At ice-off, leaf additions reduced dissolved oxygen, elevated total phosphorus concentrations and dissolved materials, and did not alter temperature or total nitrogen. These initial abiotic effects contributed to higher bacterial densities and lower chlorophyll concentrations, but by the end of spring, the abiotic environment, chlorophyll and bacterial densities converged. By contrast, zooplankton densities diverged between treatments during the spring, with leaf additions stimulating copepods but inhibiting cladocerans. We hypothesized that these differences between zooplankton orders resulted from resource shifts following leaf additions. These results suggest that leaf subsidies can alter both the short- and long-term dynamics of planktonic food webs, and highlight the importance of fully understanding how ecological subsidies are integrated into recipient food webs.
Effects of inert dust on olive (Olea europaea L.) leaf physiological para.
Nanos, George D; Ilias, Ilias F
2007-05-01
Cement factories are major pollutants for the surrounding areas. Inert dust deposition has been found to affect photosynthesis, stomatal functioning and productivity. Very few studies have been conducted on the effects of cement kiln dust on the physiology of perennial fruit crops. Our goal was to study some cement dust effects on olive leaf physiology.effects on olive leaf physiology. On Cement kiln dust has been applied periodically since April 2003 onto olive leaves. Cement dust accumulation and various leaf physiological parameters were evaluated early in July 2003. Measurements were also taken on olive trees close to the cement factory. Leaf dry matter content and specific leaf weight increased with leaf age and dust content. Cement dust decreased leaf total chlorophyll content and chlorophyll a/chlorophyll b ratio. As a result, photosynthetic rate and quantum yield decreased. In addition, transpiration rate slightly decreased, stomatal conductance to H2O and CO2 movement decreased, internal CO2 concentration remained constant and leaf temperature increased. The changes in chlorophyll are possibly due to shading and/or photosystem damage. The changes in stomatal functioning were possibly due to dust accumulation between the peltates or othe effects on stomata. Dust (in this case from a cement kiln) seems to cause substantial changes to leaf physiology, possibly leading to reduced olive productivity. Avoidance of air contamination from cement factories by using available technology should be examined together with any possible methodologies to reduce plant tissue contamination from cement dust. Longterm effects of dust (from cement kiln or other sources) on olive leaf, plant productivity and nutritional quality of edible parts could be studied for conclusive results on dust contamination effects to perennial crops.
Yoon, Leena; Liu, Ya-Nan; Park, Hyunjin; Kim, Hyun-Sook
2015-07-01
We hypothesized that olive leaf extract might alleviate dyslipidemia resulting from estrogen deficiency. Serum lipid profile and mRNA expression of the related genes in the liver and adipose tissue were analyzed after providing olive leaf extract (200 or 400 mg/kg body weight; n=7 for each group) to ovariectomized rats for 10 weeks. After 10 weeks' administration, the rats in the olive leaf extract-administered groups showed significantly lower levels of serum triglyceride and very-low-density lipoprotein (VLDL)-cholesterol compared with the rats in the control group, whereas the administration of olive leaf extract did not significantly change the elevated low-density lipoprotein cholesterol levels. In addition, administration of high dose of olive leaf extract significantly decreased the liver triglyceride and increased serum estradiol levels. mRNA expressions of peroxisome proliferator-activated receptor alpha (PPAR α) and acyl-CoA oxidase (ACO) were not affected by ovariectomy, however, administration of olive leaf extract significantly increased both PPAR α and ACO mRNA expression. Expression of adiponectin mRNA in adipose tissue was significantly decreased in the ovariectomized control group. Rats administered low-dose olive leaf extract showed significantly elevated adiponectin mRNA expression compared with rats in the ovariectomized control group. Even though dose-dependent effects were not observed in most of the measurements, these results suggest that genes involved in lipid metabolism may be regulated by olive leaf extract administration in ovariectomized rats.
Effects of dew deposition on transpiration and carbon uptake in leaves
NASA Astrophysics Data System (ADS)
Gerlein-Safdi, C.; Koohafkan, M.; Chung, M.; Rockwell, F. E.; Thompson, S. E.; Caylor, K. K.
2017-12-01
Dew deposition occurs in ecosystems worldwide, even in the driest deserts and in times of drought. Although some species absorb dew water directly via foliar uptake, a ubiquitous effect of dew on plant water balance is the interference of dew droplets with the leaf energy balance, which increases leaf albedo and emissivity and decreases leaf temperature through dew evaporation. Dew deposition frequency and amount are expected to be affected by changing environmental conditions, with unknown consequences for plant water stress and ecosystem carbon, water and energy fluxes. Here we present a simple leaf energy balance that characterizes the effect of deposition and the evaporation of dew on leaf energy balance, transpiration, and carbon uptake. The model is driven by five common meteorological variables and shows very good agreement with leaf wetness sensor data from the Blue Oak Ranch Reserve in California. We explore the tradeoffs between energy, water, and carbon balances for leaves of different sizes across a range of relative humidity, wind speed, and air temperature conditions. Our results show significant water savings from transpiration suppression up to 30% for leaf characteristic lengths of 50 cm due to the decrease in leaf temperature. C. 25% of water savings from transpiration suppression in smaller leaves arise from the effect of dew droplets on leaf albedo. CO2 assimilation is decreased by up to 15% by the presence of dew, except for bigger leaves in windspeed conditions below 1 m/s when an increase in assimilation is expected.
A dynamic leaf gas-exchange strategy is conserved in woody ...
Rising atmospheric [CO2], ca, is expected to affect stomatal regulation of leaf gas-exchange of woody plants, thus influencing energy fluxes as well as carbon (C), water and nutrient cycling of forests. Researchers have reported that stomata regulate leaf gas-exchange around “set points” that include a constant leaf internal [CO2], ci, a constant drawdown in CO2 (ca - ci), and a constant ci/ca. Because these set points can result in drastically different consequences for leaf gas-exchange, it will be essential for the accuracy of Earth systems models that generalizable patterns in leaf gas-exchange responses to ca be identified if any do exist. We hypothesized that the concept of optimal stomatal behavior, exemplified by woody plants shifting along a continuum of these set point strategies, would provide a unifying framework for understanding leaf gas-exchange responses to ca. We analyzed studies reporting C stable isotope ratio (δ13C) or photosynthetic discrimination (∆13C) from woody plant taxa that grew across ca spanning at least 100 ppm for each species investigated. From these data we calculated ci, and in combination with known or estimated ca, leaf gas-exchange regulation strategies were assessed. Overall, our analyses does not support the hypothesis that trees are canalized towards any of the proposed set points, particularly so for a constant ci. Rather, the results are consistent with the hypothesis that stomatal optimization regulates leaf gas
NASA Astrophysics Data System (ADS)
Pedentchouk, N.; Lawson, T.; Eley, Y.; McAusland, L.
2012-04-01
Stable isotopic compositions of oxygen and hydrogen are used widely to investigate modern and ancient water cycles. The D/H composition of organic compounds derived from terrestrial plants has recently attracted significant attention as a proxy for palaeohydrology. However, the role of various plant physiological and biochemical factors in controlling the D/H signature of leaf wax lipids in extant plants remains unclear. The focus of this study is to investigate the effect of plant transpiration on the D/H composition of n-alkanes in terrestrial plants. This experiment includes 4 varieties of Arabidopsis thaliana that differ with respect to stomatal density and stomatal geometry. All 4 varieties were grown indoors under identical temperature, relative humidity, light and watering regimes and then sampled for leaf wax and leaf water stable isotopic measurements. During growth, stomatal conductance to carbon dioxide and water vapour were also determined. We found that the plants varied significantly in terms of their transpiration rates. Transpiration rates were significantly higher in Arabidopsis ost1 and ost1-1 varieties (2.4 and 3.2 mmol m-2 s-1, respectively) than in Arabidopsis RbohD and Col-0 (1.5 and 1.4). However, hydrogen isotope measurements of n-alkanes extracted from leaf waxes revealed a very different pattern. Varieties ost1, ost1-1, and RbohD have very similar deltaD values of n-C29 alkane (-125, -128, and -127 per mil), whereas the deltaD value of Col-0 is more negative (-137 per mil). The initial results of this work suggest that plant transpiration is decoupled from the D/H composition of n-alkanes. In other words, physical processes that affect water vapour movement between the plant and its environment apparently cannot account for the stable hydrogen isotope composition of organic compounds that comprise leaf waxes. Additional, perhaps biochemical, processes that affect hydrogen isotope fractionation during photosynthesis might need to be invoked to explain the reason for this decoupling. Our current work that also includes leaf water isotopic measurements will provide further details regarding the role of transpiration in controlling the deltaD values of leaf lipids.
TALE and Shape: How to Make a Leaf Different.
Di Giacomo, Elisabetta; Iannelli, Maria Adelaide; Frugis, Giovanna
2013-05-06
The Three Amino acid Loop Extension (TALE) proteins constitute an ancestral superclass of homeodomain transcription factors conserved in animals, plants and fungi. In plants they comprise two classes, KNOTTED1-LIKE homeobox (KNOX) and BEL1-like homeobox (BLH or BELL, hereafter referred to as BLH), which are involved in shoot apical meristem (SAM) function, as well as in the determination and morphological development of leaves, stems and inflorescences. Selective protein-protein interactions between KNOXs and BLHs affect heterodimer subcellular localization and target affinity. KNOXs exert their roles by maintaining a proper balance between undifferentiated and differentiated cell state through the modulation of multiple hormonal pathways. A pivotal function of KNOX in evolutionary diversification of leaf morphology has been assessed. In the SAM of both simple- and compound-leafed seed species, downregulation of most class 1 KNOX (KNOX1) genes marks the sites of leaf primordia initiation. However, KNOX1 expression is re-established during leaf primordia development of compound-leafed species to maintain transient indeterminacy and morphogenetic activity at the leaf margins. Despite the increasing knowledge available about KNOX1 protein function in plant development, a comprehensive view on their downstream effectors remains elusive. This review highlights the role of TALE proteins in leaf initiation and morphological plasticity with a focus on recent advances in the identification of downstream target genes and pathways.
Coupled atmosphere/canopy model for remote sensing of plant reflectance features
NASA Technical Reports Server (NTRS)
Gerstl, S. A.; Zardecki, A.
1985-01-01
Solar radiative transfer through a coupled system of atmosphere and plant canopy is modeled as a multiple-scattering problem through a layered medium of random scatterers. The radiative transfer equation is solved by the discrete-ordinates finite-element method. Analytic expressions are derived that allow the calculation of scattering and absorption cross sections for any plant canopy layer form measurable biophysical parameters such as the leaf area index, leaf angle distribution, and individual leaf reflectance and transmittance data. An expression for a canopy scattering phase function is also given. Computational results are in good agreement with spectral reflectance measurements directly above a soybean canopy, and the concept of greenness- and brightness-transforms of Landsat MSS data is reconfirmed with the computed results. A sensitivity analysis with the coupled atmosphere/canopy model quantifies how satellite-sensed spectral radiances are affected by increased atmospheric aerosols, by varying leaf area index, by anisotropic leaf scattering, and by non-Lambertian soil boundary conditions. Possible extensions to a 2-D model are also discussed.
Local auxin production underlies a spatially restricted neighbor-detection response in Arabidopsis
Michaud, Olivier
2017-01-01
Competition for light triggers numerous developmental adaptations known as the “shade-avoidance syndrome” (SAS). Important molecular events underlying specific SAS responses have been identified. However, in natural environments light is often heterogeneous, and it is currently unknown how shading affecting part of a plant leads to local responses. To study this question, we analyzed upwards leaf movement (hyponasty), a rapid adaptation to neighbor proximity, in Arabidopsis. We show that manipulation of the light environment at the leaf tip triggers a hyponastic response that is restricted to the treated leaf. This response is mediated by auxin synthesized in the blade and transported to the petiole. Our results suggest that a strong auxin response in the vasculature of the treated leaf and auxin signaling in the epidermis mediate leaf elevation. Moreover, the analysis of an auxin-signaling mutant reveals signaling bifurcation in the control of petiole elongation versus hyponasty. Our work identifies a mechanism for a local shade response that may pertain to other plant adaptations to heterogeneous environments. PMID:28652343
Black leaf streak disease affects starch metabolism in banana fruit.
Saraiva, Lorenzo de Amorim; Castelan, Florence Polegato; Shitakubo, Renata; Hassimotto, Neuza Mariko Aymoto; Purgatto, Eduardo; Chillet, Marc; Cordenunsi, Beatriz Rosana
2013-06-12
Black leaf streak disease (BLSD), also known as black sigatoka, represents the main foliar disease in Brazilian banana plantations. In addition to photosynthetic leaf area losses and yield losses, this disease causes an alteration in the pre- and postharvest behavior of the fruit. The aim of this work was to investigate the starch metabolism of fruits during fruit ripening from plants infected with BLSD by evaluating carbohydrate content (i.e., starch, soluble sugars, oligosaccharides, amylose), phenolic compound content, phytohormones, enzymatic activities (i.e., starch phosphorylases, α- and β-amylase), and starch granules. The results indicated that the starch metabolism in banana fruit ripening is affected by BLSD infection. Fruit from infested plots contained unusual amounts of soluble sugars in the green stage and smaller starch granules and showed a different pattern of superficial degradation. Enzymatic activities linked to starch degradation were also altered by the disease. Moreover, the levels of indole-acetic acid and phenolic compounds indicated an advanced fruit physiological age for fruits from infested plots.
Fanwoua, Julienne; Bairam, Emna; Delaire, Mickael; Buck-Sorlin, Gerhard
2014-01-01
Understanding the role of branch architecture in carbon production and allocation is essential to gain more insight into the complex process of assimilate partitioning in fruit trees. This mini review reports on the current knowledge of the role of branch architecture in carbohydrate production and partitioning in apple. The first-order carrier branch of apple illustrates the complexity of branch structure emerging from bud activity events and encountered in many fruit trees. Branch architecture influences carbon production by determining leaf exposure to light and by affecting leaf internal characteristics related to leaf photosynthetic capacity. The dynamics of assimilate partitioning between branch organs depends on the stage of development of sources and sinks. The sink strength of various branch organs and their relative positioning on the branch also affect partitioning. Vascular connections between branch organs determine major pathways for branch assimilate transport. We propose directions for employing a modeling approach to further elucidate the role of branch architecture on assimilate partitioning. PMID:25071813
Spring Constants for Stacks of Curved Leaves of Pyrolytic Boron Nitride
NASA Technical Reports Server (NTRS)
Kaforey, M. L.; Deeb, C. W.; Matthiesen, D. H.
1999-01-01
Stacks of curved leaves of pyrolytic boron nitride (PBN) were deflected and the force versus deflection data was recorded. From this data, the spring constant for a given spring geometry (radius of curvature of a leaf, width of a leaf, thickness of a leaf, and number of leaves in the stack) was determined. These experiments were performed at room temperature, 500 C and 1000 C. However, temperature was not found to affect the spring constant. The measured values were generally within one order of magnitude of predictions made using a previously derived equation for a simply supported cylindrical section with a line force at the center.
Simonin, Kevin A; Santiago, Louis S; Dawson, Todd E
2009-07-01
Although crown wetting events can increase plant water status, leaf wetting is thought to negatively affect plant carbon balance by depressing photosynthesis and growth. We investigated the influence of crown fog interception on the water and carbon relations of juvenile and mature Sequoia sempervirens trees. Field observations of mature trees indicated that fog interception increased leaf water potential above that of leaves sheltered from fog. Furthermore, observed increases in leaf water potential exceeded the maximum water potential predicted if soil water was the only available water source. Because field observations were limited to two mature trees, we conducted a greenhouse experiment to investigate how fog interception influences plant water status and photosynthesis. Pre-dawn and midday branchlet water potential, leaf gas exchange and chlorophyll fluorescence were measured on S. sempervirens saplings exposed to increasing soil water deficit, with and without overnight canopy fog interception. Sapling fog interception increased leaf water potential and photosynthesis above the control and soil water deficit treatments despite similar dark-acclimated leaf chlorophyll fluorescence. The field observations and greenhouse experiment show that fog interception represents an overlooked flux into the soil-plant-atmosphere continuum that temporarily, but significantly, decouples leaf-level water and carbon relations from soil water availability.