Fire regimes of quaking aspen in the Mountain West
Shinneman, Douglas J.; Baker, William L.; Rogers, Paul C.; Kulakowski, Dominik
2013-01-01
Quaking aspen (Populus tremuloides Michx.) is the most widespread tree species in North America, and it is found throughout much of the Mountain West (MW) across a broad range of bioclimatic regions. Aspen typically regenerates asexually and prolifically after fire, and due to its seral status in many western conifer forests, aspen is often considered dependent upon disturbance for persistence. In many landscapes, historical evidence for post-fire aspen establishment is clear, and following extended fire-free periods senescing or declining aspen overstories sometimes lack adequate regeneration and are succeeding to conifers. However, aspen also forms relatively stable stands that contain little or no evidence of historical fire. In fact, aspen woodlands range from highly fire-dependent, seral communities to relatively stable, self-replacing, non-seral communities that do not require fire for persistence. Given the broad geographic distribution of aspen, fire regimes in these forests likely co-vary spatially with changing community composition, landscape setting, and climate, and temporally with land use and climate – but relatively few studies have explicitly focused on these important spatiotemporal variations. Here we reviewed the literature to summarize aspen fire regimes in the western US and highlight knowledge gaps. We found that only about one-fourth of the 46 research papers assessed for this review could be considered fire history studies (in which mean fire intervals were calculated), and all but one of these were based primarily on data from fire-scarred conifers. Nearly half of the studies reported at least some evidence of persistent aspen in the absence of fire. We also found that large portions of the MW have had little or no aspen fire history research. As a result of this review, we put forth a classification framework for aspen that is defined by key fire regime parameters (fire severity and probability), and that reflects underlying biophysical settings and correlated aspen functional types. We propose the following aspen fire regime types: (1) fire-independent, stable aspen; (2) fire-influenced, stable aspen; (3) fire-dependent, seral, conifer-aspen mix; (4) fire-dependent, seral, montane aspen-conifer; and (5) fire-dependent, seral, subalpine aspen-conifer. Closing research gaps and validating our proposed aspen fire regime classification will likely require additional site-specific research, enhanced dendrochronology techniques, charcoal and pollen record analysis, spatially-explicit modeling, and other techniques. We hope to encourage development of site-appropriate disturbance ecology characterizations, in order to aid efforts to manage and restore aspen communities and to diagnose key factors contributing to changes in aspen.
First-year postfire and postharvest soil temperatures in aspen and conifer stands
Michael C. Amacher; Amber D. Johnson; Debra E. Kutterer; Dale L. Bartos
2001-01-01
Aspen (Populus tremuloides Michx.) stands are in decline throughout the Interior Western United States because of fire suppression, overbrowsing by domestic livestock and native ungulates, and forest succession. We measured summertime soil temperatures in stable aspen, decadent aspen, and mixed aspen/conifer stands; a mixed aspen/conifer clearcut;...
Complex interactions shaping aspen dynamics in the Greater Yellowstone Ecosystem
Kathryn Brown; Andrew J. Hansen; Robert E. Keane; Lisa J. Graumlich
2006-01-01
Loss of aspen (Populus tremuloides) has generated concern for aspen persistence across much of the western United States. However, most studies of aspen change have been at local scales and our understanding of aspen dynamics at broader scales is limited. At local scales, aspen loss has been attributed to fire exclusion, ungulate herbivory, and...
Aspen symposium '89 proceedings; 1989 July 25-27; Duluth, MN.
USDA FS
1990-01-01
This proceedings of Aspen Symposium `89 contains 31 papers balanced between the subjects of aspen ecology and silvics, aspen management and silviculture, and aspen products and utilization. It also includes eight brief papers based on poster presentations.
Predicting the Spatial Distribution of Aspen Growth Potential in the Upper Great Lakes Region
Eric J. Gustafson; Sue M. Lietz; John L. Wright
2003-01-01
One way to increase aspen yields is to produce aspen on sites where aspen growth potential is highest. Aspen growth rates are typically predicted using site index, but this is impractical for landscape-level assessments. We tested the hypothesis that aspen growth can be predicted from site and climate variables and generated a model to map the spatial variability of...
Villalba, Juan J; Burritt, Elizabeth A; St Clair, Samuel B
2014-10-01
Aspen (Populus tremuloides Michx.) has evolved a chemical defense system comprised of phenolic glycosides (PG), which effectively deter insect herbivory. However, much less is known about the role of PG and the nutritional quality of the associated plant community on aspen browse susceptibility to mammalian herbivores. In three successive periods during the growing season, we conducted experiments with sheep by offering leaves from two aspen stands with different concentrations of PG (LOW, HIGH) or aspen leaves vs. leaves from a forb (Utah pea, Lathyrus pauciflorus) or a grass (smooth brome, Bromus inermis Leyss.) growing in an aspen understory. Intake of aspen (19 to 35 % PG) was low in all periods (1 to 6 g/Kg(0.75) in 2 hr) supporting the notion that aspen's defense system may contribute to its ecological success. However, lambs ate larger amounts of LOW than of HIGH suggesting that sheep could discriminate between aspen stands with different concentrations of PG, even when both stands were relatively well defended. Concentration of nutrients and chemical defenses in aspen leaves remained fairly stable across the growing season, and preference for aspen increased over the growing season. In contrast, preference for the forb and the grass decreased across the growing season in concert with a decline in the nutritional quality of these plants. The data suggest that nutritional context of aspen and associated forage species drove preference more than contrasts in defense chemistry of aspen. There may be periods of "susceptibility" of aspen use by mammalian herbivores, despite high concentrations of chemical defenses, which can potentially be targeted by management to reduce aspen herbivory.
Sustaining aspen in western landscapes: Symposium proceedings; 13-15 June 2000; Grand Junction, CO
Wayne D. Shepperd; Dan Binkley; Dale L. Bartos; Thomas J. Stohlgren; Lane G. Eskew
2001-01-01
The current status and trend of aspen is a topic of debate; some studies have claimed dramatic reductions in aspen stands while others have found no major changes. The actual picture of aspen forests across the West is variable, and the presence of conifers and ungulates in aspen may or may not indicate a progressive loss of aspen. These proceedings summarize the state...
Assessing aspen using remote sensing
Randy Hamilton; Kevin Megown; Jeff DiBenedetto; Dale Bartos; Anne Mileck
2009-01-01
Large areas of aspen (Populus tremuloides) have disappeared and continue to disappear from western forests due to successional decline and sudden aspen decline (SAD). This loss of aspen ecosystems negatively impacts watersheds, wildlife, plants, and recreation. Much can still be done to restore aspen if timely and appropriate action is taken. However, land managers...
Long-term aspen exclosures in the Yellowstone ecosystem
Charles E. Kay
2001-01-01
Aspen has been declining in the Yellowstone Ecosystem for more than 80 years. Some authors have suggested that aspen is a marginal plant community in Yellowstone and that recent climatic variation has adversely affected aspen, while others contend that excessive browsing by native ungulates is primarily responsible for aspen's widespread decline. To test these...
Aspen regeneration in south-central Colorado, San Isabel National Forest
Tim Benedict
2001-01-01
The potential for aspen regeneration in conifer stands has been underestimated on the Salida Ranger District. Harvest of mature aspen stands on the Salida and San Carlos Districts encouraged regeneration. Following harvest, the Douglas-fir and some Engelmann spruce stands in the Arkansas Hills area regenerated primarily to aspen. Disturbance through aspen harvest,...
Landscape composition in aspen woodlands under various modeled fire regimes
Eva K. Strand; Stephen C. Bunting; Lee A. Vierling
2012-01-01
Quaking aspen (Populus tremuloides) is declining across the western United States. Aspen habitats are diverse plant communities in this region and loss of these habitats can cause shifts in biodiversity, productivity, and hydrology across spatial scales. Western aspen occurs on the majority of sites seral to conifer species, and long-term maintenance of these aspen...
Aspen: Ecology and management in the western United States
Norbert V. DeByle; Robert P. Winokur
1985-01-01
Information about the biology, ecology, and management of quaking aspen on the mountains and plateaus of the interior western United States, and to a lesser extent, Canada, is summarized and discussed. The biology of aspen as a tree species, community relationships in the aspen ecosystem, environments, and factors affecting aspen forests are reviewed. The resources...
Landscape dynamics of aspen and conifer forests
Dale L. Bartos
2001-01-01
Quaking aspen (Populus tremuloides Michx.) is widely dispersed across the landscape of North America. Seventy-five percent of the aspen in the western United States occurs in the states of Colorado (50%) and Utah (25%). Reproduction in aspen is primarily by asexual means, e.g., root sprouts that are generally referred to as suckers. An aspen clone consists of numerous...
Detection of aspen-conifer forest mixes from LANDSAT digital data. [Utah-Idaho Bear River Range
NASA Technical Reports Server (NTRS)
Jaynes, R. A.; Merola, J. A.
1982-01-01
Aspen, conifer and mixed aspen/conifer forests were mapped for a 15-quadrangle study area in the Utah-Idaho Bear River Range using LANDSAT multispectral scanner data. Digital classification and statistical analysis of LANDSAT data allowed the identification of six groups of signatures which reflect different types of aspen/conifer forest mixing. Photo interpretations of the print symbols suggest that such classes are indicative of mid to late seral aspen forests. Digital print map overlays and acreage calculations were prepared for the study area quadrangles. Further field verification is needed to acquire additional information about the nature of the forests. Single date LANDSAT analysis should be a cost effective means to index aspen forests which are at least in the mid seral phase of conifer invasion. Since aspen canopies tend to obscure understory conifers for early seral forests, a second date analysis, using data taken when aspens are leafless, could provide information about early seral aspen forests.
Smith, David Solance; Fettig, Stephen M.; Bowker, Matthew A.
2016-01-01
Quaking aspen (Populus tremuloides) is the most widespread tree species in North America and has supported a unique ecosystem for tens of thousands of years, yet is currently threatened by dramatic loss and possible local extinctions. While multiple factors such as climate change and fire suppression are thought to contribute to aspen’s decline, increased browsing by elk (Cervus elaphus), which have experienced dramatic population increases in the last ∼80 years, may severely inhibit aspen growth and regeneration. Fires are known to favor aspen recovery, but in the last several decades the spatial scale and intensity of wildfires has greatly increased, with poorly understood ramifications for aspen growth. Here, focusing on the 2000 Cerro Grande fire in central New Mexico – one of the earliest fires described as a “mega-fire” - we use three methods to examine the impact of elk browsing on aspen regeneration after a mega-fire. First, we use an exclosure experiment to show that aspen growing in the absence of elk were 3× taller than trees growing in the presence of elk. Further, aspen that were both protected from elk and experienced burning were 8.5× taller than unburned trees growing in the presence of elk, suggesting that the combination of release from herbivores and stimulation from fire creates the largest aspen growth rates. Second, using surveys at the landscape level, we found a correlation between elk browsing intensity and aspen height, such that where elk browsing was highest, aspen were shortest. This relationship between elk browsing intensity and aspen height was stronger in burned (r = −0.53) compared to unburned (r = −0.24) areas. Third, in conjunction with the landscape-level surveys, we identified possible natural refugia, microsites containing downed logs, shrubs etc. that may inhibit elk browsing by physically blocking aspen from elk or by impeding elk’s ability to move through the forest patch. We did not find any consistent patterns between refuge elements and aspen size or canopy cover suggesting that natural refugia are not aiding in aspen recruitment and that all young aspen were susceptible to browsing. In much of their normal range, aspen are not growing to large size classes, which threatens the future of this iconic species and calls into question the ability of ecosystems to recover from mega-fires. Our results highlight the importance of considering multiple interacting factors (i.e. fire and increased elk browsing) when considering aspen management and regeneration.
Wayne D. Shepperd; Frederick W. Smith; Kristen A. Pelz
2015-01-01
An experimental assessment of the use of clearfell harvesting to initiate a regeneration response in commercially managed aspen forests affected by sudden aspen decline (SAD) was conducted in western Colorado in cooperation with the USDA Forest Service. Nine pure commercial quality aspen stands, with three levels of mortality attributed to SAD, were selected (...
Prescribed fire, elk, and aspen in Grand Teton National Park
Ron Steffens; Diane Abendroth
2001-01-01
In Grand Teton National Park, a landscape-scale assessment of regeneration in aspen has assisted park managers in identifying aspen stands that may be at risk due to a number of interrelated factors, including ungulate browsing and suppression of wildland fire. The initial aspen survey sampled an estimated 20 percent of the park's aspen stands. Assessment of these...
Pamela J. Jakes
1981-01-01
The fourth Minnesota Forest Inventory shows that aspen continues to dominate the State's forests. Thirty-nine percent of Minnesota's commercial forest area is in the aspen forest type. Aspen species accounted for the largest portion of growing-stock inventory, net annual growth, and removals.
Tool for Analysis and Reduction of Scientific Data
NASA Technical Reports Server (NTRS)
James, Mark
2006-01-01
The Automated Scheduling and Planning Environment (ASPEN) computer program has been updated to version 3.0. ASPEN as a whole (up to version 2.0) has been summarized, and selected aspects of ASPEN have been discussed in several previous NASA Tech Briefs articles. Restated briefly, ASPEN is a modular, reconfigurable, application software framework for solving batch problems that involve reasoning about time, activities, states, and resources. Applications of ASPEN can include planning spacecraft missions, scheduling of personnel, and managing supply chains, inventories, and production lines. ASPEN 3.0 can be customized for a wide range of applications and for a variety of computing environments that include various central processing units and randomaccess memories. Domain-specific reasoning modules (e.g., modules for determining orbits for spacecraft) can easily be plugged into ASPEN 3.0. Improvements over other, similar software that have been incorporated into ASPEN 3.0 include a provision for more expressive time-line values, new parsing capabilities afforded by an ASPEN language based on Extensible Markup Language, improved search capabilities, and improved interfaces to other, utility-type software (notably including MATLAB).
Fire effects in northeastern forests: aspen.
Cary Rouse
1986-01-01
Fire has been a natural component of the aspen ecosystem. Any fire in an established aspen stand will cause injury. Aspen is easily top-killed, but the roots remain viable. A fire's heat can stimulate sprout growth from these roots, aiding natural regeneration.
Aspen wood characteristics, properties and uses: a review of recent literature.
Fred M. Lamb
1967-01-01
Summarizes information on wood properties and uses of quaking aspen from recent literature. Includes current data on the growth and production of aspen in the Lake States. Outlines additional research needs concerning aspen wood properties and uses.
Toni Lyn Morelli; Susan C. Carr
2011-01-01
We conducted a literature review of the effects of climate on the distribution and growth of quaking aspen (Populus tremuloides Michx.) in the Western United States. Based on our review, we summarize models of historical climate determinants of contemporary aspen distribution. Most quantitative climate-based models linked aspen presence and growth...
Management of aspen in a changing environment
Shinneman, Douglas; Halford, Anne S.; Howell, Cheri; Krasnow, Kevin D.; Strand, Eva K.; Chambers, Jeanne
2015-01-01
Aspen communities are biologically rich and ecologically valuable, yet they face myriad threats, including changing climate, altered fire regimes, and excessive browsing by domestic and wild ungulates. Recognizing the different types of aspen communities that occur in the Great Basin, and being able to distinguish between seral and stable aspen stands, can help managers better identify restoration needs and objectives. Identifying key threats to aspen regeneration and persistence in a given stand or landscape is important to designing restoration plans, and to selecting appropriate treatment types. Although some aspen stands will need intensive treatment (e.g., use of fire) to persist or remain healthy, other stands may only require the modification of current management practices (e.g., reducing livestock browsing) or may not require any action at all (e.g., self-replacing stable aspen communities).
ERIC Educational Resources Information Center
Entman, Robert M.; Katz, Michael L.
The Aspen Institute's Communications and Society Program convened leaders and experts in the telecommunications and related fields to address telecommunications regulation in an IP (Internet Protocols) environment at the 15th annual Aspen Institute Telecommunications Policy Conference (Aspen, Colorado, August 12-16, 2000). The report from this…
NASA Technical Reports Server (NTRS)
Merola, J. A.; Jaynes, R. A.; Harniss, R. O.
1983-01-01
Aspen, conifer and mixed aspen/conifer forests were mapped for a 15-quadrangle study area in the Utah-Idaho Bear River Range using LANDSAT multispectral scanner (MSS) data. The digital MSS data were utilized to devise quantitative indices which correlate with apparently stable and seral aspen forests. The extent to which a two-date LANDSAT MSS analysis may permit the delineation of different categories of aspen/conifer forest mix was explored. Multitemporal analyses of MSS data led to the identification of early, early to mid, mid to late, and late seral stages of aspen/conifer forest mixing.
Norbert V. DeByle; Robert P. Winokur
1985-01-01
Quaking or trembling aspen (Populus tremuloides Michx.) is the only aspen in western North America. Therefore, in this part of the continent, it is commonly and correctly referred to simply as "aspen". Throughout much of the interior West, it is the only upland hardwood. Aspen occupies millions of acres, and, in some states, it is the most...
NASA Technical Reports Server (NTRS)
Rabideau, Gregg; Chien, Steve; Knight, Russell; Schaffer, Steven; Tran, Daniel; Cichy, Benjamin; Sherwood, Robert
2006-01-01
The Automated Scheduling and Planning Environment (ASPEN) computer program has been updated to version 3.0. ASPEN is a modular, reconfigurable, application software framework for solving batch problems that involve reasoning about time, activities, states, and resources. Applications of ASPEN can include planning spacecraft missions, scheduling of personnel, and managing supply chains, inventories, and production lines. ASPEN 3.0 can be customized for a wide range of applications and for a variety of computing environments that include various central processing units and random access memories.
Winnie, John A
2012-12-01
Aspen in the Greater Yellowstone Ecosystem are hypothesized to be recovering from decades of heavy browsing by elk due to a behaviorally mediated trophic cascade (BMTC). Several authors have suggested that wolves interact with certain terrain features, creating places of high predation risk at fine spatial scales, and that elk avoid these places, which creates refugia for plants. This hypothesized BMTC could release aspen from elk browsing pressure, leading to a patchy recovery in places of high risk. I tested whether four specific, hypothesized fine-scale risk factors are correlated with changes in current elk browsing pressure on aspen, or with aspen recruitment since wolf reintroduction, in the Daly Creek drainage in Yellowstone National Park, and near two aspen enclosures outside of the park boundary. Aspen were not responding to hypothesized fine-scale risk factors in ways consistent with the current BMTC hypothesis.
Summary: Aspen decline in the West?
Dennis H. Knight
2001-01-01
No other tree in the Rocky Mountain region is more highly valued for its amenities than aspen (Populus tremuloides). In Colorado, New Mexico, Arizona, and southern Utah, aspen covers entire mountain slopes and plateaus, sometimes forming the landscape matrix in which other cover types occur as patches. Northward aspen occurs in patches (figure 1), forming small groves...
Aspen's ecological role in the West
William H. Romme; Lisa Floyd-Hanna; David D. Hanna; Elisabeth Bartlett
2001-01-01
Aspen exhibits a variety of ecological roles. In southern Colorado, the 1880 landscape mosaic contained a range of stand ages, of which half were >70 years old and half were younger. Pure aspen stands in southern Colorado are widespread and may result from previous short fire intervals that eliminated local conifer seed sources. Aspen regeneration in northern...
Landscape assessment of a stable aspen community in southern Utah, USA
Paul C. Rogers; A. Joshua Leffler; Ronald J. Ryel
2010-01-01
Recent reports of rapid die-off of aspen (Populus tremuloides), coupled with vigorous debate over longterm reduction of aspen cover in western North America, has prompted considerable research given the importance of this forest type for economic and non-economic interests. Despite this interest, indicators of aspen conditions are...
Dale L. Bartos
2007-01-01
Quaking aspen (Populus tremuloides Michx.) is the most widely distributed broadleaf tree in North American (Little 1971; Sargent 1890). Aspen forests occur from Labrador on the east coast to Alaska in the north to Mexico in the south. In its eastern range, aspen is relatively continuously distributed. In the western United States, however, it occurs on the more...
Drought characteristics' role in widespread aspen forest mortality across Colorado, USA.
Anderegg, Leander D L; Anderegg, William R L; Abatzoglou, John; Hausladen, Alexandra M; Berry, Joseph A
2013-05-01
Globally documented widespread drought-induced forest mortality has important ramifications for plant community structure, ecosystem function, and the ecosystem services provided by forests. Yet the characteristics of drought seasonality, severity, and duration that trigger mortality events have received little attention despite evidence of changing precipitation regimes, shifting snow melt timing, and increasing temperature stress. This study draws upon stand level ecohydrology and statewide climate and spatial analysis to examine the drought characteristics implicated in the recent widespread mortality of trembling aspen (Populus tremuloides Michx.). We used isotopic observations of aspen xylem sap to determine water source use during natural and experimental drought in a region that experienced high tree mortality. We then drew upon multiple sources of climate data to characterize the drought that triggered aspen mortality. Finally, regression analysis was used to examine the drought characteristics most associated with the spatial patterns of aspen mortality across Colorado. Isotopic analysis indicated that aspens generally utilize shallow soil moisture with little plasticity during drought stress. Climate analysis showed that the mortality-inciting drought was unprecedented in the observational record, especially in 2002 growing season temperature and evaporative deficit, resulting in record low shallow soil moisture reserves. High 2002 summer temperature and low shallow soil moisture were most associated with the spatial patterns of aspen mortality. These results suggest that the 2002 drought subjected Colorado aspens to the most extreme growing season water stress of the past century by creating high atmospheric moisture demand and depleting the shallow soil moisture upon which aspens rely. Our findings highlight the important role of drought characteristics in mediating widespread aspen forest mortality, link this aspen die-off to regional climate change trends, and provide insight into future climate vulnerability of these forests. © 2013 Blackwell Publishing Ltd.
The extent and characteristics of low productivity aspen areas in Minnesota.
Gerhard K. Raile; Jerold T. Hahn
1982-01-01
Plot data from 1977 Minnesota forest inventory were used to evaluate the productivity of Minnesota's aspen forest. Computer simulation was used to develop equations for evaluating the current and potential productivity of aspen forest stands. The analysis showed that 49% of the state's aspen forest type was producing less than half of potential volume yields...
Lichen community change in response to succession in aspen forests of the southern Rocky Mountains
Paul C. Rogers; Ronald J. Ryel
2008-01-01
In western North America, quaking aspen (Populus tremuloides) is the most common hardwood in montane landscapes. Fire suppression, grazing and wildlife management practices, and climate patterns of the past century are all potential threats to aspen coverage in this region. If aspen-dependent species are losing habitat, this raises concerns about...
Long-term impact of a leaf miner outbreak on the performance of quaking aspen
D. Wagner; P. Doak
2013-01-01
The aspen leaf miner, Phyllocnistis populiella Cham., has caused widespread and severe damage to aspen in the boreal forests of western North America for over a decade. We suppressed P. populiella on individual small aspen ramets using insecticide at two sites near Fairbanks, Alaska, annually for 7 years and compared plant...
Aspen in the Sierra Nevada: Regional conservation of a continental species
Paul C. Rogers; Wayne D. Shepperd; Dale L. Bartos
2007-01-01
Quaking aspen (Populus tremuloides Michx.) a common species in North America, is a minor species in the Sierra Nevada of California. However, the limited coverage of aspen in this area appears to carry a disproportionate biodiversity load: numerous species are dependent on the unique components of aspen forests habitat. Land managers in the region...
Shearing Restores Full Productivity to Sparse Aspen Stands
Donald A. Perala
1983-01-01
Four mature but grossly under-stocked (15 to 23 percent of normal) aspen stands were regenerated by suckering following shearing. Eight years later, aspen standing crop varied with site quality from 3.4 to 8.0 tons per acre -- nearly the potential for these sites at this age. Shearing is as effective as complete clearcutting for regenerating aspen.
Aspen biology, community classification, and management in the Blue Mountains
David K. Swanson; Craig L. Schmitt; Diane M. Shirley; Vicky Erickson; Kenneth J. Schuetz; Michael L. Tatum; David C. Powell
2010-01-01
Quaking aspen (Populus tremuloides Michx.) is a valuable species that is declining in the Blue Mountains of northeastern Oregon. This publication is a compilation of over 20 years of aspen management experience by USDA Forest Service workers in the Blue Mountains. It includes a summary of aspen biology and occurrence in the Blue Mountains, and a...
Planting aspen to rehabilitate riparian areas: a pilot study
Wayne D. Shepperd; Stephen A. Mata
2005-01-01
We planted 742 greenhouse-grown containerized aspen seedlings in the riparian area of Hurd Creek on the Arapaho National Forest east of Tabernash, Colorado. Objectives were to (1) determine whether aspen seedlings can be planted in an operational setting and survive in sufficient numbers to successfully establish a mature aspen stand and (2) determine the effectiveness...
Basic tree-ring sample preparation techniques for aging aspen
Lance A. Asherin; Stephen A. Mata
2001-01-01
Aspen is notoriously difficult to age because of its light-colored wood and faint annual growth rings. Careful preparation and processing of aspen ring samples can overcome these problems, yield accurate age and growth estimates, and concisely date disturbance events present in the tree-ring record. Proper collection of aspen wood is essential in obtaining usable ring...
Above- and below-ground effects of aspen clonal regeneration and succession to conifers
Wayne D. Shepperd; Dale L. Bartos; Stephen A. Mata
2001-01-01
Above- and below-ground characteristics were measured and compared for six sets of paired trembling aspen (Populus tremuloides Michx.) clones on the Fishlake National Forest in central Utah. Three self-regenerating clones were compared with three non-regenerating clones and three pure aspen stands were compared with three mixed aspen-conifer stands. Regenerating clones...
Trembling aspen response to a mixed-severity wildfire in the Black Hills, South Dakota, USA
Tara L. Keyser; Frederick W. Smith; Wayne D. Shepperd
2005-01-01
Trembling aspen (Populus tremuloides Michx.) regeneration dynamics including sprout production, growth, and clone size were measured to determine the effects of fire on small aspen clone persistence following a mixedseverity wildfire in the Black Hills, South Dakota. Four years postfire, 10 small, isolated aspen clones per low and high fire severity...
Modeling aspen cover type diameter distributions in Minnesota
Curtis L. VanderSchaaf
2015-01-01
An attempt was made to model diameter distributions of aspen (Populus spp) stands. The aspen (1,020,150 acres) cover type has the greatest acreage on DNR lands and it contains a variety of hardwood and softwood species. Aspen is a valuable pulpwood species, annually comprising around 50 percent of total timber harvest on DNR lands....
Yang, Jian; Weisberg, Peter J.; Shinneman, Douglas; Dilts, Thomas E.; Earnst, Susan L.; Scheller, Robert M
2015-01-01
Content Changing aspen distribution in response to climate change and fire is a major focus of biodiversity conservation, yet little is known about the potential response of aspen to these two driving forces along topoclimatic gradients. Objective This study is set to evaluate how aspen distribution might shift in response to different climate-fire scenarios in a semi-arid montane landscape, and quantify the influence of fire regime along topoclimatic gradients. Methods We used a novel integration of a forest landscape succession and disturbance model (LANDIS-II) with a fine-scale climatic water deficit approach to simulate dynamics of aspen and associated conifer and shrub species over the next 150 years under various climate-fire scenarios. Results Simulations suggest that many aspen stands could persist without fire for centuries under current climate conditions. However, a simulated 2–5 °C increase in temperature caused a substantial reduction of aspen coverage at lower elevations and a modest increase at upper elevations, leading to an overall reduction of aspen range at the landscape level. Increasing fire activity may favor aspen increase at its upper elevation limits adjacent to coniferous forest, but may also favor reduction of aspen at lower elevation limits adjacent to xeric shrubland. Conclusions Our study highlights the importance of incorporating fine-scale terrain effects on climatic water deficit and ecohydrology when modeling species distribution response to climate change. This modeling study suggests that climate mitigation and adaptation strategies that use fire would benefit from consideration of spatial context at landscape scales.
Buck, Joshua R.; St. Clair, Samuel B.
2012-01-01
Development and change in forest communities are strongly influenced by plant-soil interactions. The primary objective of this paper was to identify how forest soil characteristics vary along gradients of forest community composition in aspen-conifer forests to better understand the relationship between forest vegetation characteristics and soil processes. The study was conducted on the Fishlake National Forest, Utah, USA. Soil measurements were collected in adjacent forest stands that were characterized as aspen dominated, mixed, conifer dominated or open meadow, which includes the range of vegetation conditions that exist in seral aspen forests. Soil chemistry, moisture content, respiration, and temperature were measured. There was a consistent trend in which aspen stands demonstrated higher mean soil nutrient concentrations than mixed and conifer dominated stands and meadows. Specifically, total N, NO3 and NH4 were nearly two-fold higher in soil underneath aspen dominated stands. Soil moisture was significantly higher in aspen stands and meadows in early summer but converged to similar levels as those found in mixed and conifer dominated stands in late summer. Soil respiration was significantly higher in aspen stands than conifer stands or meadows throughout the summer. These results suggest that changes in disturbance regimes or climate scenarios that favor conifer expansion or loss of aspen will decrease soil resource availability, which is likely to have important feedbacks on plant community development. PMID:23285012
Federal Register 2010, 2011, 2012, 2013, 2014
2011-11-08
... to 100 1.5 megawatt (MW) to 3 MW wind turbine generators with a nameplate capacity of 250 MW of power... Intent to Prepare an Environmental Impact Statement for the Quaking Aspen Wind Energy Project, Wyoming... Statement (EIS) for the Quaking Aspen Wind Energy Project (Quaking Aspen). By this notice, BLM is: (1...
Evaluation of burned aspen communities in Jackson Hole, Wyoming
Charles E. Kay
2001-01-01
Aspen has been declining in Jackson Hole for many years, a condition generally attributed to the fact that lightning fires have been aggressively suppressed since the early 1900s. It is also believed that burning will successfully regenerate aspen stands despite high elk numbers. To test this hypothesis, I evaluated 467 burned and 495 adjacent, unburned aspen stands at...
Current trends in the management of aspen and mixed aspen forests for sustainable production
A. J. David; John C. Zasada; D. W. Gilmore; S. M. Landhausser
2001-01-01
Quaking aspen (Populus tremuloides Michx.) is a remarkable species that performs several significant ecological roles throughout its range while at the same time is facing ever-increasing harvesting pressure. Although its full product potential remains untapped, aspen utilization has increased noticeably in the past 15 years as it has become a desired species for...
Paul C. Rogers
2007-01-01
In western North America, quaking aspen (Populus tremuloides Michx.) is the most common hardwood in montane landscapes. Fire suppression, grazing, wildlife management practices, and climate patterns of the past century are some of the threats to aspen coverage in this region. Researchers are concerned that aspen-dependent species may be losing...
Climate variability and fire effects on quaking aspen in the central Rocky Mountains, USA
Vachel A. Carter; Andrea Brunelle; Thomas A. Minckley; John D. Shaw; R. Justin DeRose; Simon Brewer
2017-01-01
Our understanding of how climate and fire have impacted quaking aspen (Populus tremuloides Michx.) communities prior to the 20th century is fairly limited. This study analysed the period between 4500 and 2000 cal. yr BP to assess the pre-historic role of climate and fire on an aspen community during an aspen-dominated period.
HOW to Identify and Minimize White Trunk Rot of Aspen
Michael E. Ostry; James W. Walters
1983-01-01
Phellinus tremulae (=Fomes ignarius var populinus) causes a heart rot of aspen that causes more volume loss than any other disease of aspen. Severity of the disease increases with stand age. In fact, incidence of white trunk rot is a major consideration in determining aspen rotations. Although no consistent relation exists between site and decay, generally less volume...
Ungulate herbivory on Utah aspen: Assessment of longterm exclosures
Charles E. Kay; Dale L. Bartos
2000-01-01
The role of livestock grazing and big-game browsing in the decline of aspen (Populus tremuloides Michx.) in the Intermountain West has long been questioned.All known aspen exclosures (n=8) on the Dixie and Fishlake National Forests in south-central Utah were measured during late summer of 1995 and 1996 to determine aspen stem dynamics, successional status, and...
Kristen A. Pelz; Frederick W. Smith
2013-01-01
There has been speculation that quaking aspen (Populus tremuloides) dominance of forests will increase due to mortality caused by mountain pine beetle (Dendroctonus ponderosae) (MPB). High aspen sucker densities have been observed inthe years following MPB-caused pine mortality, but it remains unclear if this disturbance will result in a pulse of aspen...
Brian Palik; Kory Cease; Leanne Egeland; Charles Blinn
2003-01-01
We examined aspen regeneration under diferent riparian management zone (RMZ) treatments in aspen forests in northern Minnesota. We also compared aspen regeneration in partially harvested RMZs to adjacent upland clearcuts. The four RMZ treatments included: (1) full control (no cutting in RMZ or upland); (2) riparian control (RMZ uncut; upland clearcut); and partially...
Characteristics of aspen infected with heartrot: Implications for cavity-nesting birds
Chris Witt
2010-01-01
Phellinus tremulae is an important fungal decay agent common to aspen and a critical component to the cavity-nesting bird complex found in western aspen stands. Little information exists on the conditions that facilitate infection and spread of P. tremulae in aspen forests. I used Forest Inventory and Analysis (FIA) data to explore the relationships of several tree and...
Simulation of quaking aspen potential fire behavior in Northern Utah, USA
R. Justin DeRose; A. Joshua Leffler
2014-01-01
Current understanding of aspen fire ecology in western North America includes the paradoxical characterization that aspen-dominated stands, although often regenerated following fire, are âfire-proofâ. We tested this idea by predicting potential fire behavior across a gradient of aspen dominance in northern Utah using the Forest Vegetation Simulator and the Fire and...
NASA Astrophysics Data System (ADS)
Meier, G.; Brown, J. F.; Vogelmann, J. E.; Evelsizer, R.
2012-12-01
Trembling aspen (Populus tremuloides, referred hereafter as Aspen) has an especially wide geographical distribution in North America, extending from Alaska across the Canadian provinces, the U.S., and south into Mexico. This deciduous species is successional, shade intolerant, and often exists as a dominant among other species at mid-elevations. Aspen occupies wide latitudinal, elevational, and environmental gradients making it a favorable candidate for a study of phenology and climate relationships. The phenological characterization in our Aspen study is derived from a database of conterminous U.S. phenological indicators hosted by the U.S. Geological Survey (http://phenology.cr.usgs.gov/index.php). Nine satellite-derived phenological indicators are calculated from 250m resolution Moderate Resolution Imaging Spectroradiometer (MODIS) Normalized Difference Vegetation Index (NDVI). From this database, we selected start of season (SOST), end of season (EOST), maximum NDVI (MaxN) and time integrated NDVI (TIN) to characterize and analyze the seasonal patterns of Aspen over a 10-year time period (2001-2010). Areas of continuous Aspen cover (≥ 80% Aspen cover type) derived from the LANDFIRE project were then used to extract elevation, precipitation, temperature, and snow water equivalent data. In the Rocky Mountains, Aspen recently suffered from multi-year drought stress accompanied by insect and disease infestations. Numerous studies have documented the existence of Sudden Aspen Decline (SAD) in Montana, Utah, Arizona, and Colorado, indicating that Aspen may be on the edge of its environmental tolerances in some areas. The satellite-derived phenology metrics, and climate and biogeographical indicators were the basis for characterizing Aspen seasonality and assessing the environmental context of SAD. Between several Aspen study areas, there was reasonably consistent progression in the SOST timing from low elevations to higher elevations. A less obvious progression was observed in the EOST, and year-to-year EOST variability appeared to be related to the timing of episodic widespread snow events. Also, higher and lower elevation Aspen stands showed lower TIN (a proxy for annual net primary productivity) and lower MaxN (a proxy for maximum vegetation vigor) compared with the mid-elevations. In the 10-year MODIS record, there was a slight trend toward a later SOST (delay from 8 to 16 days depending on the locale), but a concurrent increase of approximately 4% in the TIN over the same 10-year period. A severe drought (affecting much of Colorado) in 2002 resulted in a low TIN and very low MaxN for that year. This drought event resulted in a compression of the Aspen SOST over its full elevational range, such that Aspen showed anomalous synchronicity in phenological response apparently due to moisture stress. The occurrence of the severe drought in 2002, followed by additional dry years in 2003 and 2004 are thought to be causal factors in the occurrence of SAD. The preliminary analysis of the phenology and climate of Aspen showed the complexity of these relationships and their variability through time. Further research is proceeding to compare and contrast SAD-affected and non-SAD affected areas.
Zeigenfuss, Linda C.; Binkley, Dan; Tuskan, Gerald A.; Romme, William H.; Yin, Tongming; DiFazio, Stephen; Singer, Francis J.
2008-01-01
Lack of recruitment and canopy replacement of aspen (Populus tremuloides) stands that grow on the edges of grasslands on the low-elevation elk (Cervus elaphus) winter range of Rocky Mountain National Park (RMNP) in Colorado have been a cause of concern for more than 70 years. We used a combination of traditional dendrochronology and genetic techniques as well as measuring the characteristics of regenerating and nonregenerating stands on the elk winter range to determine when and under what conditions and estimated elk densities these stands established and through what mechanisms they may regenerate. The period from 1975 to 1995 at low elevation on the east side had 80-95 percent fewer aspen stems than would be expected based on the trend from 1855 through 1965. The age structure of aspen in the park indicates that the interacting effects of fires, elk population changes, and livestock grazing had more-or-less consistent effects on aspen from 1855 to 1965. The lack of a significant change in aspen numbers in recent decades in the higher elevation and west side parts of the park supports the idea that the extensive effects of elk browsing have been more important in reducing aspen numbers than other factors. The genetic variation of aspen populations in RMNP is high at the molecular level. We expected to find that most patches of aspen in the park were composed of a single clone of genetically identical trees, but in fact just 7 percent of measured aspen patches consisted of a single clone. A large frequency of polyploid (triploid and tetraploid) genotypes were found on the low elevation, east-side elk winter range. Nonregenerating aspen stands on the winter range had greater annual offtake, shorter saplings, and lower density of mid-height (1.5-2.5 m) saplings than regenerating stands. Overwinter elk browsing, however, did not appear to inhibit the leader length of aspen saplings. The winter range aspen stands of RMNP appear to be highly resilient in the face of very intense herbivory by elk and harsh environmental conditions. Conservation efforts through fencing protection and decreased elk browsing pressure are already being planned as part of the park's new elk management plan. If these efforts are undertaken, conditions that encourage stem recruitment to the tree canopy will likely result and the continued survival of these aspen stands will be enhanced.
Biodiversity: Aspen stands have the lead, but will nonnative species take over?
Geneva W. Chong; Sara E. Simonson; Thomas J. Stohlgren; Mohammed A. Kalkhan
2001-01-01
We investigated vascular plant and butterfly diversity in Rocky Mountain National Park. We identified 188 vascular plant species unique to the aspen vegetation type. The slope of the mean species-area curve for the aspen vegetation type was the steepest of the 10 types sampled, thus, an increase in aspen area could have much greater positive impacts on plant species...
Grindstone Flat and Big Flat enclosures - 41-year record of changes in clearcut aspen communities
Walter F. Mueggler; Dale L. Bartos
1977-01-01
The role of deer and cattle in the failure of aspen stands to regenerate on Beaver Mountain in southern Utah was investigated by a series of exclosures constructed in 1934. Three-fourths of each exclosure was clearcut of aspen in 1934. Aspen reproduction, shrubs, and herbaceous understory were measured in 1937, 1942, 1949, and 1975. Implications of wildlife and...
Jennifer DeWoody; Thomas H. Rickman; Bobette E. Jones; Valerie D. Hipkins
2009-01-01
The most widely distributed tree in North America, quaking aspen (Populus tremuloides, Michx.), reproduces sexually via seed and clonally via suckers. The size of aspen clones varies geographically, generally smaller in the east and large in the arid Intermountain West. In order to describe clone size and genetic structure of aspen in the southern Cascade...
Diane L. Wagner; Linda DeFoliart; Patricia Doak; Jenny Schneiderheinze
2008-01-01
The aspen leaf miner, Phyllocnistis populiella, feeds on the contents of epidermal cells on both top (adaxial) and bottom (abaxial) surfaces of quaking aspen leaves, leaving the photosynthetic tissue of the mesophyll intact. This type of feeding is taxonomically restricted to a small subset of leaf mining insects but can cause widespread plant...
Aspen overstory recruitment in northern Yellowstone National Park during the last 200 years
Eric J. Larsen; William J. Ripple
2001-01-01
Using a monograph provided by Warren (1926) and two sets of aspen increment cores collected in 1997 and 1998, we analyzed aspen overstory recruitment in Yellowstone National Park (YNP) over the past 200 years. We found that successful aspen overstory recruitment occurred on the northern range of YNP from the middle to late 1700s until the 1920s, after which it...
Joseph O. Sexton; R. Douglas Ramsey; Dale L. Bartos
2006-01-01
Quaking aspen (Populus tremuloides Michx.) is the most widely distributed tree species in North America, but its presence is declining across much of the Western United States. Aspen decline is complex, but results largely from two factors widely divergent in temporal scale: (1) Holocene climatic drying of the region has led to water limitation of aspen seedling...
Recovering aspen follow changing elk dynamics in Yellowstone: evidence of a trophic cascade?
Painter, Luke E; Beschta, Robert L; Larsen, Eric J; Ripple, William J
2015-01-01
To investigate the extent and causes of recent quaking aspen (Populus tremuloides) recruitment in northern Yellowstone National Park, we measured browsing intensity and height of young aspen in 87 randomly selected aspen stands in 2012, and compared our results to similar data collected in 1997-1998. We also examined the relationship between aspen recovery and the distribution of Rocky Mountain elk (Cervus elaphus) and bison (Bison bison) on the Yellowstone northern ungulate winter range, using ungulate fecal pile densities and annual elk count data. In 1998, 90% of young aspen were browsed and none were taller-than 200 cm, the height at which aspen begin to escape from elk browsing. In 2012, only 37% in the east and 63% in the west portions of the winter range were browsed, and 65% of stands in the east had young aspen taller than 200 cm. Heights of young aspen were inversely related to browsing intensity, with the least browsing and greatest heights in the eastern portion of the range, corresponding with recent changes in elk density and distribution. In contrast with historical elk distribution (1930s-1990s), the greatest densities of elk recently (2006-2012) have been north of the park boundary (approximately 5 elk/km2), and in the western part of the range (2-4 elk/km2), with relatively few elk in the eastern portion of the range (<2 elk/km2), even in mild winters. This redistribution of elk and decrease in density inside the park, and overall reduction in elk numbers, explain why many aspen stands have begun to recover. Increased predation pressure following the reintroduction of gray wolves (Canis lupius) in 1995-1996 played a role in these changing elk population dynamics, interacting with other influences including increased predation by bears (Ursus spp.), competition with an expanding bison population, and shifting patterns of human land use and hunting outside the park. The resulting new aspen recruitment is evidence of a landscape-scale trophic cascade in which a resurgent large carnivore community, combined with other ecological changes, has benefited aspen through effects on ungulate prey.
Quaking aspen productivity recovers after repeated prescribed fire.
D. A. Perala
1995-01-01
Describes how quaking aspen (Populus tremuloides Michx.) stand recovered after logging, and logging and burning. Aspen suckering was profuse after each destructive episode but differences in stockability caused different yield trajectories.
Multiscale habitat selection by Ruffed Grouse at low population densities
Zimmerman, G.S.; Gutierrez, R.J.; Thogmartin, W.E.; Banerjee, S.
2009-01-01
Theory suggests habitats should be chosen according to their relative evolutionary benefits and costs. It has been hypothesized that aspen (Populus spp.) forests provide optimal habitat for Ruffed Grouse (Bonasa umbellus). We used the low phase of a grouse population's cycle to assess the prediction that grouse should occupy aspen and avoid other forest types at low population density because of the presumptive fitness benefits of aspen. On the basis of our observations, we predict how the Ruffed Grouse population will increase in different forest types during the next cycle. In conifer (Pinus spp., Abies balsamea, Picea spp.)-dominated and mixed aspen-conifer landscapes, grouse densities were highest where forest types were evenly distributed. Within these landscapes, male Ruffed Grouse selected young aspen stands that were large and round or square. Although Ruffed Grouse selected young aspen stands strongly, contrary to prediction, they also used other forest types even when young aspen stands remained unoccupied. The relative densities of Ruffed Grouse in aspen and conifer forests indicated that the aspen forest's carrying capacities for grouse was higher than the conifer forest's at least during the low and declining phases of the grouse's cycle. On the basis of our observations, we predict that Ruffed Grouse populations in aspen-dominated landscapes will have higher population densities and fluctuate more than will populations in conifer-dominated landscapes. We suggest that studies of avian habitat selection would benefit from knowledge about the relative densities among habitats at differing population sizes because this information could provide insight into the role of habitat in regulating populations and clarify inferences from studies about habitat quality for birds. ?? 2009 by The Cooper Ornithological Society. All rights reserved.
Ungulate herbivory alters leaf functional traits and recruitment of regenerating aspen.
Rhodes, Aaron C; Anderson, Val; St Clair, Samuel B
2017-03-01
Herbivory by ungulates can affect forest regeneration success, but its long-term impacts on tree function and recruitment are less studied. We evaluated strategies of resistance, tolerance and vertical escape against ungulate herbivory by evaluating leaf traits (photosynthesis, morphology and chemistry) and growth rates of aspen in the presence and absence of ungulate herbivores 1, 2, 3 and 26 years after fires initiated aspen suckering. Over the initial 3-year period, ~60% of aspen stems in unfenced plots showed evidence of being browsed by ungulates. After 3 years, aspen in unfenced plots had smaller leaves, were 50% shorter, and had 33% lower nonstructural carbohydrate concentrations and 33% greater concentrations of condensed tannins, when compared with fenced aspen. Aspen exposed to ungulate herbivory over a 26-year period maintained smaller leaves, had lower annual radial growth rates and were still below the critical height threshold of 2 m required to escape ungulate herbivory for successful recruitment. In contrast, the average height of aspen protected from ungulates was approaching 6 m. Over the 26-year period leaves in unfenced plots had 41% lower nonstructural carbohydrate concentrations and greater expression of defense compounds-condensed tannins (63%) and phenolic glycosides (102%)-than leaves in fenced plots. Photosynthetic rates were slightly higher in aspen that experienced ungulate browsing, suggesting that changes in leaf anatomy and chemistry due to ungulate herbivory did not interfere with photosynthesis. Our data suggest that ungulate browsing increases investment in chemical defense, lowers nonstructural carbohydrate concentrations and reduces leaf area, which decreases the recruitment potential of regenerating aspen. © The Author 2017. Published by Oxford University Press. All rights reserved. For Permissions, please email: journals.permissions@oup.com.
Drought causes reduced growth of trembling aspen in western Canada.
Chen, Lei; Huang, Jian-Guo; Alam, Syed Ashraful; Zhai, Lihong; Dawson, Andria; Stadt, Kenneth J; Comeau, Philip G
2017-07-01
Adequate and advance knowledge of the response of forest ecosystems to temperature-induced drought is critical for a comprehensive understanding of the impacts of global climate change on forest ecosystem structure and function. Recent massive decline in aspen-dominated forests and an increased aspen mortality in boreal forests have been associated with global warming, but it is still uncertain whether the decline and mortality are driven by drought. We used a series of ring-width chronologies from 40 trembling aspen (Populus tremuloides Michx.) sites along a latitudinal gradient (from 52° to 58°N) in western Canada, in an attempt to clarify the impacts of drought on aspen growth by using Standardized Precipitation Index (SPI) and Standardized Precipitation Evapotranspiration Index (SPEI). Results indicated that prolonged and large-scale droughts had a strong negative impact on trembling aspen growth. Furthermore, the spatiotemporal variability of drought indices is useful for explaining the spatial heterogeneity in the radial growth of trembling aspen. Due to ongoing global warming and rising temperatures, it is likely that severer droughts with a higher frequency will occur in western Canada. As trembling aspen is sensitive to drought, we suggest that drought indices could be applied to monitor the potential effects of increased drought stress on aspen trees growth, achieve classification of eco-regions and develop effective mitigation strategies to maintain western Canadian boreal forests. © 2017 John Wiley & Sons Ltd.
Evaluation of pelleted aspen foliage as a ruminant feedstuff
DOE Office of Scientific and Technical Information (OSTI.GOV)
Bas, F.J.; Ehle, F.R.; Goodrich, R.D.
1985-01-01
Growth and digestion trials were made to determine the nutritive value of pelleted aspen (Populus tremuloides) foliage as a dietary ingredient for sheep. Lambs offered diets without or with 25, 50 and 75% aspen leaves, with lucerne as the other dietary ingredient, ate less and gained less weight as the proportion of aspen leaves in the diet increased (P less than 0.05). Digestibility coefficients for DM, organic matter, crude protein, gross energy, neutral detergent fibre, acid detergent fibre, hemicellulose and cellulose decreased linearly (P less than 0.01) as the percentage of aspen foliage eaten increased. Calculated digestibility of individual aspenmore » leaf components gave values as low as 16.6 and 13% for crude protein and cellulose, respectively. Coefficients of determination for the linear regressions indicated no associative effects between lucerne and aspen leaves. Due to the depressed value for crude protein digestibility, the amount of acid detergent fibre-insoluble nitrogen was estimated. Over 50% of the total N in aspen foliage was bound to the acid detergent fibre fraction, reflecting the presence of heat-damaged protein, tannin-protein complexes that are unavailable for digestion or both. After adjustment for unavailable N, the crude protein digestibility of aspen foliage was 61.5%. Balances of 10 minerals were estimated during the digestion trial. Negative mineral balances for the 75% aspen leaf diet suggest that the lambs were in a nutrient deficient condition when fed on this diet. 29 references.« less
Quaking aspen reproduce from seed after wildfire in the mountains of southeastern Arizona
Ronald D. Quinn; Lin Wu
2001-01-01
Quaking aspen regenerated from seed after a stand replacement wildfire in the Chiricahua Mountains of southeastern Arizona. The wildfire had created gaps in the canopy so that aspen were able to establish from seed. Seedlings were found at a mean density of 0.17 m-2, 30 m or more from the nearest potential seed trees. Six clumps of aspen seedlings contained 18-186...
Analysis of aspen stand structure and composition in the Western U.S.: implications for management
J. D. Shaw
2004-01-01
Aspen communities in the western U.S. are considered at risk because of low levels of disturbance and high levels of herbivory by wild and domestic ungulates. There appears to be a trend toward the loss of aspen-dominated stands West-wide. In some cases the loss is caused by succession, with shade-tolerant conifers becoming dominant. On dry sites where aspen is...
Modeling aspen responses to climatic warming and insect defoliation in western Canada
E. H. Ted Hogg
2001-01-01
Effects of climate change at three aspen sites in Saskatchewan were explored using a climate-driven model that includes insect defoliation. A simulated warming of 4-5 °C caused complete mortality due to drought at all three sites. A simulated warming of 2-2.5 °C caused complete mortality of aspen at the parkland site, while aspen growth at two boreal sites showed...
Can aspen persist in conifer dominated forests?
Douglas H. Page; John D. Shaw
2016-01-01
In 1998 we measured a large, old aspen in a mixed spruce-fir-aspen forest on the Utah State University T.W. Daniel Experimental Forest in northern Utah. The tree was 297 years old - about the same age as the oldest spruce in the stand. A search of the forestry literature revealed that the oldest published age for an aspen came from a tree in the Sierra Nevada Range in...
Clone expansion and competition between quaking and bigtooth aspen suckers after clearcutting.
Donald A. Perala
1981-01-01
Quaking aspen clones expanded over more area, and regenerated greater sucker stem densities and biomass than did bigtooth aspen clones. However, sucker height growth was similar between the two species.
Detection of variations in aspen forest habitat from LANDSAT digital data: Bear River Range, Utah
NASA Technical Reports Server (NTRS)
Merola, J. A.; Jaynes, R. A. (Principal Investigator)
1982-01-01
The aspen forests of the Bear River Range were analyzed and mapped using data recorded on July 2, 1979 by the LANDSAT III satellite; study efforts yielded sixty-seven light signatures for the study area, of which three groups were identified as aspen and mapped at a scale of 1:24,000. Analysis and verification of the three groups were accomplished by random location of twenty-six field study plots within the LANDSAT-defined aspen areas. All study plots are included within the Cache portion of the Wasatch-Cache National Forest. The following selected site characteristics were recorded for each study plot: a list of understory species present; average percent cover density for understory species; aspen canopy cover estimates and stem measurements; and general site topographic characteristics. The study plot data were then analyzed with respect to corresponding Landsat spectral signatures. Field studies show that all twenty-six study plots are associated with one of the three aspen groups. Further study efforts concentration on characterizing the differences between the site characteristics of plots falling into each of the three aspen groups.
R. Minocha; S. Long; S. Minocha; P Marquardt; M. Kubiske
2010-01-01
The objective of the present study was to evaluate the long-term (10 years) effects of elevated CO2 and O3 on the carbon and nitrogen metabolism of aspen trees. The study was conducted at the Aspen Free-Air Carbon Dioxide Enrichment (FACE) experimental site, Rhinelander, WI, (USA).
Iterative Repair Planning for Spacecraft Operations Using the Aspen System
NASA Technical Reports Server (NTRS)
Rabideau, G.; Knight, R.; Chien, S.; Fukunaga, A.; Govindjee, A.
2000-01-01
This paper describes the Automated Scheduling and Planning Environment (ASPEN). ASPEN encodes complex spacecraft knowledge of operability constraints, flight rules, spacecraft hardware, science experiments and operations procedures to allow for automated generation of low level spacecraft sequences. Using a technique called iterative repair, ASPEN classifies constraint violations (i.e., conflicts) and attempts to repair each by performing a planning or scheduling operation. It must reason about which conflict to resolve first and what repair method to try for the given conflict. ASPEN is currently being utilized in the development of automated planner/scheduler systems for several spacecraft, including the UFO-1 naval communications satellite and the Citizen Explorer (CX1) satellite, as well as for planetary rover operations and antenna ground systems automation. This paper focuses on the algorithm and search strategies employed by ASPEN to resolve spacecraft operations constraints, as well as the data structures for representing these constraints.
Dale L. Bartos
2008-01-01
The health of quaking aspen (Populus tremuloides) in the Great Basin is of growing concern. The following provides an overview of aspen decline and die-off in areas within and adjacent to the Great Basin and suggests possible directions for research and management.
Pärnik, Tiit; Ivanova, Hiie; Keerberg, Olav; Vardja, Rael; Niinemets, Ulo
2014-06-01
The growth rate of triploid European aspen (Populus tremula L.) and hybrid aspen (P. tremula × Populus tremuloides Michx.) significantly exceeds that of diploid aspen, but the underlying physiological controls of the superior growth rates of these genotypes are not known. We tested the hypothesis that the superior growth rate of triploid and hybrid aspen reflects their greater net photosynthesis rate. Micropropagated clonal plants varying in age from 2.5 to 19 months were used to investigate the ploidy and plant age interaction. The quantum yield of net CO2 fixation (Φ) in leaves of young 2.5-month-old hybrid aspen was lower than that of diploid and triploid trees. However, Φ in 19-month-old hybrid aspen was equal to that in triploid aspen and higher than that in diploid aspen. Φ and the rate of light-saturated net photosynthesis (ANS) increased with plant age, largely due to higher leaf dry mass per unit area in older plants. ANS in leaves of 19-month-old trees was highest in hybrid, medium in triploid and lowest in diploid aspen. Light-saturated photosynthesis had a broad temperature optimum between 20 and 35 °C. Rate of respiration in the dark (RDS) did not vary among the genotypes in 2.5-month-old plants, and the shape of the temperature response was also similar. RDS increased with plant age, but RDS was still not significantly different among the leaves of 19-month-old diploid and triploid aspen, but it was significantly lower in leaves of 19-month-old hybrid plants. The initial differences in the growth of plants with different ploidy were minor up to the age of 19 months, but during the next 2 years, the growth rate of hybrid aspen exceeded that of triploid plants by 2.7 times and of diploid plants by five times, in line with differences in ANS of 19-month-old plants of these species. It is suggested that differences in photosynthesis and growth became more pronounced with tree aging, indicating that ontogeny plays a key role in the expression of superior traits determining the productivity of given genotypes. © The Author 2014. Published by Oxford University Press. All rights reserved. For Permissions, please email: journals.permissions@oup.com.
Quaking aspen woodland after conifer control: herbaceous dynamics
USDA-ARS?s Scientific Manuscript database
Western juniper (Juniperus occidentalis Hook.) woodlands are replacing lower elevation (< 2100 m) quaking aspen (Populus tremuloides Michx.) stands in the northern Great Basin. Restoring aspen woodlands is important because they provide wildlife habitat for many species and contain a high diversity...
Kimball T. Harper; John D. Shane; John R. Jones
1985-01-01
Quaking aspen, or trembling aspen (Populus tremuloides), was named and described by Michaux in 1803. It exhibits marked phenotypic variability throughout its transcontinental range. Numerous authors, especially the early ones, tried to give order to the variability by subdividing it taxonomically. Quahng aspen has been subdivided by various...
Aspen Sucker Production and Growth from Outplanted Root Cuttings
Donald A. Perala
1978-01-01
Aspen suckers from 1-m-long root cuttings survived and grew better than those from 12.5-cm-long cuttings. Sucker survival and growth were also inversely related to parent root diameter. Discusses the practical implications for aspen management.
Barnett, David T.; Stohlgren, Thomas J.
2001-01-01
We investigated aspen (Populus tremuloides)regeneration in the Gros Ventre River Valley, the National Elk Refuge and a small part of Grand Teton National Park, Wyoming, USA to see if elk (Cervus elaphus) browsing was as damaging as previously thought. We conducted a landscape-scale survey to assess aspen regeneration across gradients of wintering elk concentrations using 68 randomly selected aspen stands in the 1090 km2 study area. Forty-four percent of the stands sampled supported some newer regeneration that had reached the canopy. There were no significant differences of regeneration across elk winter range classification (p=0.25) or distance from feedgrounds (p=0.96). However, a multiple linear regression found that the concentration of elk was one of several important predictors of successful aspen regeneration (p=0.005, R2=0.36). Our results suggest that stand-replacing regeneration occurs across the landscape at a variety of elk densities despite some trends of reduced regeneration under greater elk concentrations. We propose that high spatial and temporal variation and scattered patches of successful aspen regeneration characterize aspen persistence between periods of episodic regeneration and recruitment.
Quaking aspen woodland after conifer control: tree and shrub dynamics
USDA-ARS?s Scientific Manuscript database
Western juniper (Juniperus occidentalis spp. occidentalis) woodlands are replacing lower elevation (< 2100 m) quaking aspen (Populus tremuloides) stands of the northern Great Basin. Recovery of aspen is important because these communities provide habitat for many wildlife species. We evaluated two...
ERIC Educational Resources Information Center
Schwartz, Evan I.
This is a report of the 10th annual Aspen Institute Roundtable on Information Technology (Aspen, Colorado, August 2-5, 2001). Participants were also polled after the events of September 11, and these comments have been integrated into the report. The mission of this report is to take a wide-ranging look at the trends that are defining the next new…
Sánchez-Rodríguez, Dolores; Marco, Ester; Ronquillo-Moreno, Natalia; Maciel-Bravo, Liev; Gonzales-Carhuancho, Abel; Duran, Xavier; Guillén-Solà, Anna; Vázquez-Ibar, Olga; Escalada, Ferran; Muniesa, Josep M
2018-01-25
The aim of this study was to assess the prevalence of malnutrition by applying the ASPEN/AND definition and the ESPEN consensus definition in a postacute-care population, and secondly, to determine the metrological properties of the set of six clinical characteristics that constitute the ASPEN/AND basic diagnosis, compared to the ESPEN consensus, based mostly on objective anthropometric measurements. Prospective study of 84 consecutive deconditioned older inpatients (85.4 ± 6.2; 59.5% women) admitted for rehabilitation in postacute care. ASPEN/AND diagnosis of malnutrition was considered in presence of at least two of the following: low energy intake, fluid accumulation, diminished handgrip strength, and loss of weight, muscle mass, or subcutaneous fat. Sensitivity, specificity, positive and negative predictive values, accuracy, likelihood ratios, and kappa statistics were calculated for ASPEN/AND criteria and compared with ESPEN consensus. The prevalence of malnutrition by ASPEN/AND criteria was 63.1% and by ESPEN consensus, 20.2%; both diagnoses were associated with significantly longer length of stay, but the ESPEN definition was significantly associated with poorer functional outcomes after the rehabilitation program. Compared to ESPEN consensus, ASPEN/AND diagnosis showed fair validity (sensitivity = 94.1%; specificity = 44.8%); kappa statistic was 2.217. Applying the ASPEN/AND definition obtained a higher prevalence of malnutrition in a postacute-care population than was identified by the ESPEN definition. ASPEN/AND criteria had fair validity and agreement compared with the ESPEN definition. A simple, evidence-based, unified malnutrition definition might improve geriatric care. Copyright © 2018 Elsevier Ltd and European Society for Clinical Nutrition and Metabolism. All rights reserved.
Orbital Express Mission Operations Planning and Resource Management using ASPEN
NASA Technical Reports Server (NTRS)
Chouinard, Caroline; Knight, Russell; Jones, Grailing; Tran, Daniel
2008-01-01
As satellite equipment and mission operations become more costly, the drive to keep working equipment running with less man-power rises.Demonstrating the feasibility of autonomous satellite servicing was the main goal behind the Orbital Express (OE) mission. Planning the satellite mission operations for OE required the ability to create a plan which could be executed autonomously over variable conditions. The Automated-Scheduling and Planning Environment (ASPEN)tool, developed at the Jet Propulsion Laboratory, was used to create the schedule of events in each daily plan for the two satellites of the OE mission. This paper presents an introduction to the ASPEN tool, the constraints of the OE domain, the variable conditions that were presented within the mission, and the solution to operations that ASPEN provided. ASPEN has been used in several other domains, including research rovers, Deep Space Network scheduling research, and in flight operations for the ASE project's EO1 satellite. Related work is discussed, as are the future of ASPEN and the future of autonomous satellite servicing.
Genetic diversity in aspen and its relation to arthropod abundance
Zhang, Chunxia; Vornam, Barbara; Volmer, Katharina; Prinz, Kathleen; Kleemann, Frauke; Köhler, Lars; Polle, Andrea; Finkeldey, Reiner
2015-01-01
The ecological consequences of biodiversity have become a prominent public issue. Little is known on the effect of genetic diversity on ecosystem services. Here, a diversity experiment was established with European and North American aspen (Populus tremula, P. tremuloides) planted in plots representing either a single deme only or combinations of two, four and eight demes. The goals of this study were to explore the complex inter- and intraspecific genetic diversity of aspen and to then relate three measures for diversity (deme diversity, genetic diversity determined as Shannon index or as expected heterozygosity) to arthropod abundance. Microsatellite and AFLP markers were used to analyze the genetic variation patterns within and between the aspen demes and deme mixtures. Large differences were observed regarding the genetic diversity within demes. An analysis of molecular variance revealed that most of the total genetic diversity was found within demes, but the genetic differentiation among demes was also high. The complex patterns of genetic diversity and differentiation resulted in large differences of the genetic variation within plots. The average diversity increased from plots with only one deme to plots with two, four, and eight demes, respectively and separated plots with and without American aspen. To test whether intra- and interspecific diversity impacts on ecosystem services, arthropod abundance was determined. Increasing genetic diversity of aspen was related to increasing abundance of arthropods. However, the relationship was mainly driven by the presence of American aspen suggesting that species identity overrode the effect of intraspecific variation of European aspen. PMID:25674097
Predicting Ground Fire Ignition Potential in Aspen Communities
S. G. Otway; E. W. Bork; K. R. Anderson; M. E. Alexander
2006-01-01
Fire is one of the key disturbances affecting aspen (Populus tremuloides Michx.) forest ecosystems within western Canadian wildlands, including Elk Island National Park. Prescribed fire use is a tool available to modify aspen forests, yet clearly understanding its potential impact is necessary to successfully manage this disturbance.
Repeated Prescribed Burning in Aspen
Donald A. Perala
1974-01-01
Infrequent burning weather, low flammability of the aspen-hardwood association, and prolific sprouting and seeding of shrubs and hardwoods made repeated dormant season burning a poor tool to convert good site aspen to conifers. Repeat fall burns for wildlife habitat maintenance is workable if species composition changes are not important.
John R. Jones; Norbert V. DeByle
1985-01-01
The broad genotypic variability in quaking aspen (Populus tremuloides Michx.), that results in equally broad phenotypic variability among clones is important to the ecology and management of this species. This chapter considers principles of aspen genetics and variation, variation in aspen over its range, and local variation among clones. For a more...
76 FR 77591 - Surety Companies Acceptable on Federal Bonds: Aspen American Insurance Company
Federal Register 2010, 2011, 2012, 2013, 2014
2011-12-13
... DEPARTMENT OF THE TREASURY Fiscal Service Surety Companies Acceptable on Federal Bonds: Aspen American Insurance Company AGENCY: Financial Management Service, Fiscal Service, Department of the Treasury... hereby issued under 31 U.S.C. 9305 to the following company: Aspen American Insurance Company (NAIC 43460...
Sustaining aspen productivity in the Lake States
Douglas M. Stone
2001-01-01
Sustaining forest productivity requires maintaining soil productivity. Management activities that decrease soil porosity and remove organic matter can reduce productivity. We determined effects of three levels of organic matter removal (OMR) and soil compaction on aspen regeneration and growth following winter harvest of aspen-dominated stands in northern Minnesota,...
Aspen response to prescribed fire and wild ungulate herbivory
Steve Kilpatrick; Diane Abendroth
2001-01-01
Land management agencies in northwest Wyoming have implemented an active prescribed fire program to address historically altered fire regimes, regenerate aspen, and improve overall watershed functions. Treated clones are susceptible to extensive browsing from elk concentrated on supplemental feedgrounds and from wintering moose. Previous attempts at fire-induced aspen...
Minnesota's aspen and its projected supply
Richard W. Groff
1966-01-01
Aspen, Minnesota's largest forest type, covers nearly one-third of the commercial forest land in the State. Its major components are bigtooth aspen, P. tremuloides, both fast-growing, generally short-lived, aggressive pioneer species that readily become established in burns and clear cuttings. They cannot be "stored on the stump," as they become...
Manipulations to regenerate aspen ecosystems
Wayne D. Shepperd
2001-01-01
Vegetative regeneration of aspen can be initiated through manipulations that provide hormonal stimulation, proper growth environment, and sucker protection - the three elements of the aspen regeneration triangle. The correct course of action depends upon a careful evaluation of the size, vigor, age, and successional status of the existing clone. Soils and site...
Effect of conifer encroachment into aspen stands on understory biomass
B. R. Stam; J. C. Malechek; D. L. Bartos; J. E. Browns; E. B. Godfrey
2008-01-01
Conifers (Picea and Abies spp.) have replaced aspen (Populus tremuloides Michx.) over much of aspen's historic range in the western United States. We measured the impact of this change upon the production of understory vegetation potentially useful as forage for livestock and wildlife on two southern Utah...
The effect of aspen wood characteristics and properties on utilization
Kurt H. Mackes; Dennis L. Lynch
2001-01-01
This paper reviews characteristics and properties of aspen wood, including anatomical structure and characteristics, moisture and shrinkage properties, weight and specific gravity, mechanical properties, and processing characteristics. Uses of aspen are evaluated: sawn and veneer products, composite panels, pulp, excelsior, post and poles, animal bedding, animal food...
Effects of water and temperature
John R. Jones; Merrill R. Kaufmann; E. Arlo Richardson
1985-01-01
Aspen's geographic and elevational ranges indicate a species that tolerates severe cold but does not tolerate sustained high temperatures, or semiarid or even dry, subhumid conditions. Much can be inferred from observation of the sites on which quaking aspen grows in the West. Aspen's distribution is related to its regeneration characteristics, its pathology...
Molecular tools and aspen management: A primer and prospectus
Karen E. Mock; Bryce A. Richardson; Paul G. Wolf
2013-01-01
Aspen (Populus tremuloides) isaniconic species in North American landscapes, highly valued for recreation, fiber, wildlife and livestock forage, carbon sequestration, biodiversity, and as a fuelbreak. However, there are rising concerns about the ability of aspen to persist in portions of its range, based on bioclimatic modeling, physiological thresholds and mortality...
Mapping aspen in the Interior West
Charles E. Werstak
2012-01-01
Quaking aspen (Populus tremuloides Michx.) is a critical species that supports wildlife and livestock, watershed function, the forest products industry, landscape diversity, and recreation opportunities in the Interior West (Bartos and Campbell 1998). Studies have indicated that changes in fire regimes, an increase in herbivore presence in young aspen stands, and...
How to Create, Modify, and Interface Aspen In-House and User Databanks for System Configuration 1:
DOE Office of Scientific and Technical Information (OSTI.GOV)
Camp, D W
2000-10-27
The goal of this document is to provide detailed instructions to create, modify, interface, and test Aspen User and In-House databanks with minimal frustration. The level of instructions are aimed at a novice Aspen Plus simulation user who is neither a programming nor computer-system expert. The instructions are tailored to Version 10.1 of Aspen Plus and the specific computing configuration summarized in the Title of this document and detailed in Section 2. Many details of setting up databanks depend on the computing environment specifics, such as the machines, operating systems, command languages, directory structures, inter-computer communications software, the version ofmore » the Aspen Engine and Graphical User Interface (GUI), and the directory structure of how these were installed.« less
Calder, W John; Horn, Kevin J; St Clair, Samuel B
2011-06-01
Disturbance patterns strongly influence plant community structure. What remains less clear, particularly at a mechanistic level, is how changes in disturbance cycles alter successional outcomes in plant communities. There is evidence that fire suppression is resulting in longer fire return intervals in subalpine forests and that these lengthened intervals increase competitive interactions between aspen and conifer species. We conducted a field and greenhouse study to compare photosynthesis, growth and defense responses of quaking aspen and subalpine fir regeneration under light reductions and shifts in soil chemistry that occur as conifers increase in dominance. The studies demonstrated that aspen regeneration was substantially more sensitive to light and soil resource limitations than that of subalpine fir. For aspen, light reductions and/or shifts in soil chemistry limited height growth, biomass gain, photosynthesis and the production of defense compounds (phenolic glycosides and condensed tannins). Biomass gain and phenolic glycoside concentrations were co-limited by light reduction and changes in soil chemistry. In contrast, subalpine fir seedlings tended to be more tolerant of low light conditions and showed no sensitivity to changes in soil chemistry. Unlike aspen, subalpine fir increased its root to shoot ratio on conifer soils, which may partially explain its maintenance of growth and defense. The results suggest that increasing dominance of conifers in subalpine forests alters light conditions and soil chemistry in a way that places greater physiological and growth constraints on aspen than subalpine fir, with a likely outcome being more successful recruitment of conifers and losses in aspen cover.
Richard L. Lindroth
2001-01-01
Quaking aspen (Populus tremuloides) employs two major systems of defense against damage by environmental agents: chemical defense and tolerance. Aspen accumulates appreciable quantities of phenolic glycosides (salicylates) and condensed tannins in most tissues and accumulates coniferyl benzoate in flower buds. Phenolic glycosides are toxic and/or deterrent to pathogens...
D. A. Perala; D.H. Alban
1982-01-01
Compares rates of forest floor decomposition and nutrient turnover in aspen and conifers. These rates were generally most rapid under aspen, slowest under spruce, and more rapid on a loamy fine sand than on a very fine sandy loam. Compares results with literature values.
Ecology, biodiversity, management, and restoration of aspen in the Sierra Nevada
Wayne D. Shepperd; Paul C. Rogers; David Burton; Dale L. Bartos
2006-01-01
This report was commissioned by the USDA Forest Service Lake Tahoe Basin Management Unit to synthesize existing information on the ecology and management of aspen (Populus tremuloides) in the Sierra Nevada of California and surrounding environs. It summarizes available information on aspen throughout North America from published literature, internal...
Differential response of Aspen and Birch trees to heat stress under elevated carbon dioxide
Joseph N.T. Darbah; Thomas D. Sharkey; Carlo Calfapietra; David F. Karnosky
2010-01-01
The effect of high temperature on photosynthesis of isoprene-emitting (aspen) and non-isoprene-emitting (birch) trees were measured under elevated CO2 and ambient conditions. Aspen trees tolerated heat better than birch trees and elevated CO2 protected photosynthesis of both species against moderate heat stress. Elevated CO...
Nitrogen mineralization in the aspen ecosystem
M. C. Amacher; A. D. Johnson; T. Christopherson; D. E. Kutterer; D. L. Bartos; J. Kotuby-Amacher
1999-01-01
Aspen (Populus tremuloides) are in decline throughout the interior western U.S. because of conifer invasion, fire suppression, and overbrowsing by domestic livestock and native ungulates. Aspen restoration requires disturbances such as fire or cutting and exclosures to protect against overbrowsing. We measured the effects of the 1996 Pole Creek fire, Fishlake NF, UT on...
Effects of moisture on aspen-fiber/polypropylene composites
Roger M. Rowell
2002-01-01
Aspen fiber/polypropylene composites were made using several different levels of aspen fiber (0 to 60% by weight) with and without the addition of a compatibilizer (maleic anhydride grafted polypropylene, MAPP). These composites were tested under various relative humidity conditions and in water soaking, boiling water, cyclic liquid water and oven drying tests. In all...
Decline of quaking aspen in the Interior West - examples from Utah
Dale L. Bartos; Robert B. Campbell
1998-01-01
Quaking aspen (Populus tremuloides) are unique because, in contrast to most western forest trees, they reproduce primarily by suckering from the parent root system. Generally disturbance or dieback is necessary to stimulate regeneration of aspen stands. These self-regenerating stands have existed for thousands of years. If they are lost from the...
A synthetic sex pheromone for the large aspen tortrix in Alaska.
Richard A. Werner; J. Weatherston
1980-01-01
Cis-11-tetradecenal was found to be the specific attractant for adult male large aspen tortrix, Choristoneura conflictana (Walker), populations in quaking aspen, Populus tremuloides Michx., forests of interior Alaska. The attractant was dispersed from polyethylene caps in Pherocon® -2 traps placed 1.5 m above ground.
John R. Jones; Norbert V. DeByle
1985-01-01
Aspen has been recognized for many years as being very intolerant of shade (Baker 1918a, Clements 1910, Weigle and Frothingham 1911, Zon and Graves 1911). In dense stands, vigorous aspen trees are confined to the dominant and codominant crown classes. Regardless of size, when they are overtopped by larger trees, aspen trees deteriorate and eventually die. Many well-...
Aspen, climate, and sudden decline in western USA
Gerald E. Rehfeldt; Dennis E. Ferguson; Nicholas L. Crookston
2009-01-01
A bioclimate model predicting the presence or absence of aspen, Populus tremuloides, in western USA from climate variables was developed by using the Random Forests classification tree on Forest Inventory data from about 118,000 permanent sample plots. A reasonably parsimonious model used eight predictors to describe aspen's climate profile. Classification errors...
Effects of moisture on aspen-fiber/polypropylene composites
Roger M. Rowell; Sandra E. Lange; Rodney E. Jacobson
2004-01-01
Moisture sorption in fiber-thermoplastic composites leads to dimensional instability and biological attack. To determine the pick up of moisture this type of composite, aspen fiber/polypropylene composites were made using several different levels of aspen fiber (30 to 60% by weight) with and without the addition of a compatibilizer (maleic anhydride grafted...
Slow lifelong growth predisposes Populus tremuloides to tree mortality
Kathryn B. Ireland; Margaret M. Moore; Peter Z. Fule; Thomas J. Zegler; Robert E. Keane
2014-01-01
Widespread dieback of aspen forests, sometimes called sudden aspen decline, has been observed throughout much of western North America, with the highest mortality rates in the southwestern United States. Recent aspen mortality has been linked to drought stress and elevated temperatures characteristic of conditions expected under climate change, but the role of...
Aspen Competition Drives Innovative Ideas for Community Colleges
ERIC Educational Resources Information Center
Gonzalez, Jennifer
2012-01-01
When Valencia College became the first recipient of the Aspen Prize for Community College Excellence last month, an unsung sector earned uncommon recognition. Now that the speeches are over and the prize money has been awarded, the Aspen Institute is sharing early lessons from its yearlong effort to determine the top community college in the…
Cankers on Western Quaking Aspen
David W. Johnson; Jerome S. Beatty; Thomas E. Hinds
1995-01-01
Long appreciated for its esthetic and shade tree value and its importance for wildlife, aspen is also capable of excellent growth and high yields and thus is an important commercial timber species. However, aspen has one major drawback-its soft bark is easily wounded by abiotic factors, animals, and insects. Subsequently, these wounds can be invaded by disease...
Effects of ugulate browsing on aspen regeneration in northwestern Wyoming
Bruce L. Smith; J. Scott Dieni; Roxane L. Rogers; Stanley H. Anderson
2001-01-01
Although clearcutting has been demonstrated to be an effective means to regenerate aspen, stand replacement may be retarded under conditions of intense browsing of regeneration, such as that experienced near elk feedgrounds in northwestern Wyoming. We studied the effects of ungulate browsing on regenerating aspen following clearcutting on the National Elk Refuge. Nine...
Do pine trees in aspen stands increase bird diversity?
Mark A. Rumble; Lester D. Flake; Todd R. Mills; Brian L. Dykstra
2001-01-01
In the Black Hills of South Dakota, quaking aspen (Populus tremuloides) is being replaced by conifers through fire suppression and successional processes. Although the Black Hills National forest is removing conifers (primarily ponderosa pine [Pinus ponderosa]) to increase the aspen communities in some mixed stands, Forest Plan guidelines allow four conifers per...
What to do with northern hardwood-aspen mixtures
Richard M. Godman
1992-01-01
The aspen type covers the most area in the Lake States, followed by northern hardwoods. Both prefer good soil and are easy to regenerate. Because of past cutting practices, though, the types are now mixed over large areas. Usually there is an overstory of fast-growing aspen interspersed with an understory of dense hardwoods.
Nest-site selection and nest survival of Lewis's woodpecker in aspen riparian woodlands
Karen R. Newlon; Victoria A. Saab
2011-01-01
Riparian woodlands of aspen (Populus tremuloides) provide valuable breeding habitat for several cavity-nesting birds. Although anecdotal information for this habitat is available for Lewis's Woodpecker (Melanerpes lewis), no study has previously examined the importance of aspen woodlands to this species' breeding biology. From 2002 to 2004, we monitored 76...
Using Forest Health Monitoring to assess aspen forest cover change in the southern Rockies ecoregion
Paul Rogers
2002-01-01
Long-term qualitative observations suggest a marked decline in quaking aspen (Populus tremuloides Michx.) primarily due to advancing succession and fire suppression. This study presents an ecoregional coarse-grid analysis of the current aspen situation using Forest Health Monitoring (FHM) data from Idaho, Wyoming, and Colorado. A...
Craig W. Johnson; Thomas C. Brown; Michael L. Timmons
1985-01-01
Aspen is valued for its scenic beauty. One indication of this is the trips to the "high country" that many forest visitors make to view the autumn color changes (fig. 1). Another is the frequency with which aspen is planted in urban and suburban areas. Subjective generalizations about the esthetic uses of aspen, although reasonable, provide only rough...
Miranda T. Curzon; Anthony W. D' Amato; Brian J. Palik; Kris Verheyen
2016-01-01
Questions: Does the increase in disturbance associated with removing harvest residues negatively impact biodiversity and resilience in aspen-dominated forest ecosystems? How do responses of functional diversity measures relate to community recovery and standing biomass? Location: Aspen (Populus tremuloides, Michx.) mixedwood forests in Minnesota...
Growth and decay losses in Colorado aspen
Thomas E. Wengert Hinds
1977-01-01
Decay in Colorado aspen, Populus tremuloides Michx., was extensively surveyed in 1954-56, but volume estimates were presented on a cubic foot basis. This paper reanalyzes the data on a board foot (Scribner) basis. Tree growth and gross and net volumes per acre expected on commercial aspen sites are given. Decay volumes are correlatzd with site class...
Ralph L. Anderson; Gerald W. Anderson; Arthur L. Jr. Schipper
1979-01-01
Hypoxylon canker, caused by the fungus Hypoxylon mammatum (Wahl.) Mill. (formerly H. pruinatum (Klot.) Cke.), is one of the most important killing diseases of aspen in eastern North America. In Michigan, Minnesota, and Wisconsin, the total impact of Hypoxylon canker has been estimated to be 30 percent of the annual net growth of aspen; in 1972, trees worth more than $4...
Growth and crown architecture of two aspen genotypes exposed to interacting ozone and carbon dioxide
Richard E. Dickson; M.D. Coleman; Priit Pechter; David Karnosky
2001-01-01
To study the impact of ozone (O3) and O3 plus CO2 on aspen growth, we planted two trembling aspen clones, differing in sensitivity to O3, in the ground in open-top chambers and exposed them to different concentrations of O3 and O3 plus CO2...
W. F. Mueggler
1985-01-01
Aspen trees grow along moist stream bottoms as well as on dry ridges and southerly exposures, on talus slopes, and on shallow to deep soils of varied origins. Quaking aspen is one of the few plant species that can grow in all mountain vegetational zones from the alpine to the basal plain (Daubenmire 1943). As a consequence, aspen dominated communities are found...
Karen Rachel Newlon
2005-01-01
Aspen (Populus tremuloides) riparian woodlands are extremely limited in distribution throughout the western U.S., yet these habitats have a disproportionate value to breeding birds. Aspen habitats are also considered prime sheep and cattle summer range, particularly in the semiarid Intermountain West. Such concentrated use has raised concern about...
Effect of pruning the parent root on growth of aspen suckers
Ashbel F. Hough
1965-01-01
Various portions of the root systems of bigtooth aspen (Populus grandidentata) suckers were severed, and the subsequent height and radial growth of stems were measured. Aspen vegetative regeneration is heavily dependent on the parent roots for at least 25 years following initial suckering. The distal portion of the parent root contributes more to...
Harvey H. Smith
1947-01-01
Aspen is now the major forest type in the Lake States, and extensive stands are reaching maturity. Increasing quantities are being cut, and much is now being put to new and more exacting uses that require better air-seasoning and kiln-drying practices. The recent favorable market for green aspen lumber appears to be falling off, and the time may soon come when...
Management for esthetics and recreation, forage, water, and wildlife
Norbert V. DeByle
1985-01-01
In the West, aspen forests have not been actively managed for wood products largely because of the lack of markets for quaking aspen timber from the Rocky Mountains (see the WOOD UTILIZATION chapter). Despite this, the aspen ecosystem has been used to provide a variety of resources and opportunities (see PART 111. RESOURCES AND USES).
D.F. Karnosky; Kevin E. Percy; Bixia Xiang; Brenda Callan; Asko Noormets; Blanka Mankovska; Anthony Hopkin; Jaak Sober; Wendy Jones; R.E. Dickson; J.G. Isebrands
2002-01-01
We investigated the interaction of elevated CO2 and/or (Ozone) O3 on the occurrence and severity of aspen leaf rust (Malampsora medusae Thuem. f. sp. tremuloidae) on trembling aspen (Populus tremuloides MIchx.) Furthermore, we examined the role of changes in...
Mycorrhizal fungi of aspen forests: Natural occurrence and potential applications
Cathy L. Cripps
2001-01-01
Native mycorrhizal fungi associated with aspen were surveyed on three soil types in the north-central Rocky Mountains. Selected isolates were tested for the ability to enhance aspen seedling growth in vitro. Over 50 species of ectomycorrhizal fungi occur with Populus tremuloides in this region, primarily basidiomycete fungi in the Agaricales. Almost one-third (30%)...
Appraising fuels and flammability in western aspen: a prescribed fire guide
James K. Brown; Dennis G. Simmerman
1986-01-01
Describes a method for appraising fuels and fire behavior potential in aspen forests to guide the use of prescribed fire and the preparation of fire prescriptions. Includes an illustrated classification of aspen fuels; appraisals of fireline intensity, rate of spread, adjective ratings for fire behavior and probability of burn success; and evaluations of seasonal...
Justin M. Britton; R. Justin DeRose; Karen E. Mock; James N. Long
2016-01-01
Recent concern regarding the potential decline of quaking aspen (Populus tremuloides Michx.) forests in the western United States has sparked concern over whether the species can be effectively regenerated. Using a retrospective approach, we quantified the response of regenerating aspen stems to an ordinary set of silvicultural treatments conducted over...
Decline of aspen (Populus tremuloides) in the Interior West [Abstract 2
Dale L. Bartos
1997-01-01
It is commonly recognized that aspen (Populus tremuloides) ecosystems in the Interior West provide numerous benefits: (1) forage for livestock, (2) habitat for wildlife, (3) water for downstream users, (4) esthetics, (5) sites for recreational opportunities, (6) wood fiber, and (7) landscape diversity. Loss or potential loss of aspen on these lands can be attributed...
Cankers on Western Quaking Aspen (FIDL)
David W. Johnson; Jerome S. Beatty; Thomas E. Hinds
1995-01-01
Quaking aspen (Populus tremuloides Michx.) is one of the most well-known tree species in the western United States (figure 1). It is found from the northern limit of trees in northwestern Alaska through the western United States and into northern Mexico. Quaking aspen is an aggressive pioneer species that frequently colonizes burned sites, making it an important...
Extent of decay associated with Fomes igniarius sporophores in Colorado aspen
Thomas E. Hinds
1963-01-01
The most destructive decay of aspen (Populus tremuloides Michx.) is caused by Fomes igniarius var. populinus (Neu.) Camb. This fungus accounted for 59 percent of the decay found in a recent study of aspen in Colorado. It is almost impossible to find stands of any age that are not damaged to some degree by
Hardeep S. Rai; Karen E. Mock; Bryce A. Richardson; Richard C. Cronn; Katherine J. Hayden; Jessica W. Wright; Brian J. Knaus; Paul G. Wolf
2013-01-01
Aspen (Populus tremuloides) is a temperate North American tree species with a geographical distribution more extensive than any other tree species on the continent. Because it is economically important for pulp and paper industries and ecologically important for its role as a foundation species in forest ecosystems, the decline of aspen in large...
Dynamics of aspen root biomass and sucker production following fire
Roy A. Renkin; Don G. Despain
2001-01-01
Changes in preburn aspen root biomass 8 years following prescribed fire were analyzed for five experimental sites distributed across a moisture gradient. Total root biomass decreased across all sites but was proportionately greater in xeric than mesic sites. Response of post-burn aspen suckers to ungulate browsing varied according to site and treatment. Browsing...
Ross W. Davidson; Thomas E. Hinds; Frank G. Hawksworth
1959-01-01
Quaking aspen (Populus tremuloides Michx.) stands are extensive in the central Rocky Mountains. The species reaches its maximum development in the mountains and high mesas west of the Continental Divide in Colorado (Baker, 1925). On the better sites aspen yields a greater volume of wood in a shorter period than most of the conifers growing at comparable elevations. The...
Leanne M. Vigue; Richard L. Lindroth
2010-01-01
Trembling aspen Populus tremuloides Michaux is an important forest species in the Great Lakes region and displays tremendous genetic variation in foliar chemistry. Elevated carbon dioxide (CO2) and ozone (O3) may also influence phytochemistry and thereby alter the performance of insect herbivores such as...
Competitive status influences tree-growth responses to elevated CO2 and 03 in aggrading aspen stands
E. P. McDonald; E. L. Kruger; D. E. Riemenschneider; J. G. Isebrands
2002-01-01
1. Competition effects on growth of individual trees were examined for 4 years in aggrading, mixed-clone stands of trembling aspen (Populus tremuloides Michx.) at the Aspen-FACE free-air Co2 and O3 enrichment facility in northern Wisconsin, USA. During each growing season stands received one of four...
The aspen mortality summit; December 18 and 19, 2006; Salt Lake City, UT
Dale L. Bartos; Wayne D. Shepperd
2010-01-01
The USDA Forest Service Rocky Mountain Research Station sponsored an aspen summit meeting in Salt Lake City, Utah, on December 18 and19, 2006, to discuss the rapidly increasing mortality of aspen (Populus tremuloides) throughout the western United States. Selected scientists, university faculty, and managers from Federal, State, and non-profit agencies with experience...
Norbert V. DeByle
1985-01-01
Aspen forests provide important habitat for many species of wildlife (Gullion 1977b), especially in the West (see the appendix to this chapter). In the coniferous forests of the interior West, aspen groves may be the only source of abundant forage; in the grasslands they may be the sole source of cover. A primary value of the aspen ecosystem in the West during the past...
Ecology and management of aspen: A Lake States perspective
David T. Cleland; Larry A. Leefers; Donald I. Dickmann
2001-01-01
Aspen has been an ecologically important, though relatively minor, component of the Lake States (Michigan, Wisconsin, and Minnesota) forests for millennia. General Land Office records from the 1800s indicate that aspen comprised a small fraction of the region's eastern forests, but was more extensive on the western edge. Then Euro-American settlement in the 1800s...
Sudden aspen decline in southwest Colorado
J. J. Worrall; R. A. Mask; T. Eager; L. Egeland; W. D. Shepperd
2008-01-01
Sudden aspen decline (SAD) has increased rapidly in recent years, approaching 350,000 acres in Colorado in 2007, or 13% of the aspen cover type. We investigated the severity, site/stand factors and causes associated with SAD in southwest Colorado. First, we documented landscape (GIS-DEM analyses) and stand factors (stand exams). There was a strong inverse relationship...
R. Justin DeRose; Karen E. Mock; James N. Long
2015-01-01
High rates of triploidy have recently been described in quaking aspen (Populus tremuloides Michx.) of the Intermountain West, raising questions about the contributions of triploidy to stand persistence and dynamics. In this study, we investigated cytotype differences between diploid and triploid aspen clones using dendrochronological techniques. We used tree-ring data...
Landscape-scale dynamics of aspen in Rocky Mountain National Park, Colorado
Margot W. Kaye; Kuni Suzuki; Dan Binkley; Thomas J. Stohlgren
2001-01-01
Past studies of quaking aspen in Rocky Mountain National Park suggested that the aspen population is declining due to intensive browsing by elk (Cervus elaphus). These studies were conducted in the elk winter range, an area of intensive elk impact. The elk summer range experiences less intense grazing pressure. We tested the hypothesis that impacts of elk would be...
Establishment of white spruce growth trial in an aspen understory
Chris Maundrell; C. Hawkins
2001-01-01
A conifer release trial was established in a 45-year-old aspen overstory stand in Northeast British Columbia, Canada. Thinning occurred from 0 to 100% in increments of approximately 10% to yield a total of ten units. Treatment consisted of physically cutting aspen stems at the root collar or girdling at breast height (d.b.h.) for trees 10 cm d...
Joseph N.T. Darbah; Mark E. Kubiske; Neil Nelson; Katre Kets; Johanna Riikonen; Anu Sober; Lisa Rouse; David F. Karnosky
2010-01-01
Photosynthetic acclimation under elevated carbon dioxide (CO2) and/or ozone (O3) has been the topic of discussion in many papers recently. We examined whether or not aspen plants grown under elevated CO2 and/or O3 will acclimate after 11 years of exposure at the Aspen Face site...
E. Oksanen; E. Häikiö; J. Sober; D.F. Karnosky
2003-01-01
Saplings of three aspen (Populus tremuloides) genotypes and seedlings of paper birch (Betula papyrifera) were exposed to elevated ozone (1.5x ambient) and 560 p.p.m. CO2, singly and in combination, from 1998 at the Aspen-FACE (free-air CO2 enrichment) site (Rhinelander, USA).
Model-based assessment of aspen responses to elk herbivory in Rocky Mountain National Park
Peter J. Weisberg; Michael B. Coughenour
2001-01-01
In Rocky Mountain National Park, aspen has been observed to decline on elk winter range for many decades. The SAVANNA ecosystem model was adapted to explore interactions between elk herbivory and aspen dynamics on the elk winter range. Several scenarios were explored that considered different levels of overall elk population; different levels of elk utilization of...
Harvesting Impacts on Soil Properties and Tree Regeneration in Pure and Mixed Aspen Stands
Melissa J. Arikian; Kiaus J. Peuttmann; Alaina L. Davis; George E. Host; John Zasada
1999-01-01
Impacts of clearcutting and selective harvesting on pure aspen/mixed aspen hardwood stands were examined in northern Minnesota. We studied these impacts on 18 stands, which were harvested 4 to 11 years ago and received no further treatment. In each stand, residual composition, soil compaction, and tree regeneration were determined along a gradient of disturbance in the...
Chad M. Oukrop; David M. Evans; Dale L. Bartos; R. Douglas Ramsey; Ronald J. Ryel
2011-01-01
Quaking aspen (Populus tremuloides Michx.) are the most widely distributed tree species across North America, but its dominance is declining in many areas of the western United States, with certain areas experiencing rapid mortality events over the past decade. The loss of aspen from western landscapes will continue to profoundly impact biological, commercial, and...
Scaling Aspen-FACE experimental results to century and landscape scales
Eric J. Gustafson; Mark E. Kubiske; Brian R. Sturtevant; Brian R. Miranda
2013-01-01
The Aspen-FACE experiment generated 11 years of empirical data on the effect of CO2 enrichment and elevated ozone on the growth of field-grown trees (maple, birch and six aspen clones) in northern Wisconsin, but it is not known how these short-term plot-level responses might play out at the landscape scale over multiple decades where competition...
Restoration of Aspen-Dominated Ecosystem in the Lake States
Douglas M. Stone; John D. Elioff; Donald V. Potter; Donald B. Peterson; Robert Wagner
2001-01-01
A reserve tree method(RTM)of harvesting was installed in six 70-to75-year-old aspen-dominated stands to determine if retaining 10 to 15 dominant aspen per acre would decrease sucker density to facilitate restoration of a conifer component. A reserve shelterwood cut was applied to three additional stands to evaluate performance of white pine planted under 50% crown...
Restoration of aspen-dominated ecosystems in the Lake States
Douglas M. Stone; John D. Elioff; Donald V. Potter; Donald B. Peterson; Robert Wagner
2000-01-01
A reserve tree method (RTM) of harvesting was installed in six 70-to75-year old aspen-dominated stands to determine if retaining 10 to 15 dominant aspen per acre would decrease sucker density to facilitate restoration of a conifer component. A reserve shelterwood cut was applied to three additional stands to evaluate performance of white pine planted under 50% crown...
Aspen restoration in the Blue Mountains of northeast Oregon
Diane M. Shirley; Vicky Erickson
2001-01-01
In the Blue Mountains of northeast Oregon, quaking aspen is on the western fringe of its range. It exists as small, scattered, remnant stands of rapidly declining trees. Although little is known about the historic distribution of aspen in Oregon, it is believed that stands were once larger and more widely distributed. Decline of the species is attributed to fire...
Aspen encroachment on meadows of the North Rim, Grand Canyon National Park
Margaret M. Moore; David W. Huffman
2001-01-01
Composition and structure data were analyzed to determine the characteristics of trees encroaching on the montane meadows and subalpine grasslands of the North Rim, Grand Canyon National Park. Tree invasion in the 1900s showed a pattern of increasing establishment, with quaking aspen comprising the majority (52%) of encroaching trees. Most aspen established in the last...
Aspen indicator species in lichen communities in the Bear River range of Idaho and Utah
Paul C. Rogers; Roger Rosentreter; Ronald J. Ryel
2007-01-01
Aspen are thought to be declining in this region due to a combination of fire suppression, grazing and wildlife management practices, and potentially cool/wet climates of the past century which favor advancing conifer succession. Many scientists are concerned that aspen's related species may also be losing habitat, thereby threatening the long-term local and...
Restoration of aspen-dominated ecosystems in the Lake States
Douglas M. Stone; John D. Elioff; Donald V. Potter; Donald B. Peterson; Robert Wagner
2001-01-01
A reserve tree method (RTM) of harvesting was installed in six 70- to 75-yearold aspen-dominated stands to determine if retaining 10 to 15 dominant aspen per acre would decrease sucker density to facilitate restoration of a conifer component. A reserve shelterwood cut was applied to three additional stands to evaluate performance of white pine planted under 50% crown...
Mortality of aspen on the Gros Ventre elk winter range
Richard G. Krebill
1972-01-01
Stands of aspen on the Gros Ventre elk winter range of northwestern Wyoming are suffering high mortality and are not regenerating satisfactorily. If the 1970 mortality rate (3.6 percent) continues, about a two-thirds reduction in the numbers of tree-sized aspen can be expected by year 2000. Collected evidence suggests that the mortality rate is unusually high because...
Hybrid Aspen Response to Shearing in Minnesota: Implications for Biomass Production
Grant M. Domke; Andrew J. David; Anthony W. D' Amato; Alan R. Ek; Gary W. Wycoff
2011-01-01
There is great potential for the production of woody biomass feedstocks from hybrid aspen stands; however, little is known about the response of these systems to silvicultural treatments, such as shearing. We sought to address this need by integrating results from more than 20 years of individual tree and yield measurements in hybrid aspen (Populus tremuloides Mich. Ã...
ERIC Educational Resources Information Center
McGuire, Georgianna; Nicolau, Siobhan
This report from the 1997 Aspen Institute seminar concerns how demographic changes in American will affect Hispanic Americans' role in the business community. Section 1, "Lashes: Back, Front, and Sideways" (Harold Hodgkinson), describes pervasive national pessimism over demographic change and documents universal backlash to that change…
Aspen decline on the Coconino National Forest
Mary Lou Fairweather; Brian W. Geils; Mike Manthei
2008-01-01
An accelerated decline of aspen occurred across the Coconino National Forest, in northern Arizona, following a frost event in June 1999, and a long-term drought that included an extremely dry and warm period from 2001 through 2002, and bouts of defoliation by the western tent caterpillar in 2004, 2005, and 2007. From 2003 to 2007, we monitored aspen mortality and...
Insects and other invertebrates
John R. Jones; Norbert V. DeByle; Diane M. Bowers
1985-01-01
Quaking aspen throughout its range appears to be host to several insect and other invertebrate pests (fig. 1). It is a short-lived species that is palatable to a large variety of animals. Furniss and Carolin (1977) listed 33 insect species that use aspen as a food source. Some are quite damaging and may kill otherwise healthy stands of aspen; others feed on weakened or...
Predation risk and elk-aspen foraging patterns
Clifford A. White; Michael C. Feller
2001-01-01
Elk-aspen foraging patterns may be influenced by cover type, distance from roads or trails, the type of user on road or trail (park visitor, human hunter, or predator), and two general states of aspen condition (open-grown or thicket). Pellet group and browse utilization transects in the Canadian Rockies showed that elk were attracted to roads used by park visitors and...
Soil properties associated with various stages of succession in the aspen ecosystem
Michael C. Amacher; Dale L. Bartos
1997-01-01
In the Interior west, if current conditions continue (e.g., lack of fire, wildlife use, grazing by livestock, natural succession) that have prevailed for the past 100 to 140 years, most aspen stands will eventually be replaced by conifers, sagebrush, or possibly tall shrub communities. Current estimates are there has beeh a 60% decrease in aspen dominated lands since...
Decline of aspen (Populus tremuloides) in the Interior West [Abstract 1
Dale L. Bartos; Robert B Campbell
1997-01-01
Western aspen forests are unique because they reproduce primarily by suckering from the parent root system. Generally a disturbance or die back is necessary to stimulate regeneration of the stands. Unlike other tree species, if aspen stands are lost from the landscape, generally they will not return through natural processes. If current conditions continue (e.g., lack...
Evaluation of techniques to protect aspen suckers from ungulate browsing in the Black Hills
Andrew M. Kota; Dale L. Bartos
2010-01-01
Excessive browsing by cattle (Bos taurus L.) and wild ungulates, particularly elk (Cervus elaphus L.), sometimes inhibits growth and maturity of quaking aspen (Populus tremuloides Michx.) suckers in Western North America. In areas where aspen stands are in poor condition or declining, protecting suckers from ungulates may be necessary. This study compared the utility...
Response of transplanted aspen to irrigation and weeding on a Colorado reclaimed surface coal mine
Robert C. Musselman; Wayne D. Shepperd; Frederick W. Smith; Lance A. Asherin; Brian W. Gee
2012-01-01
Successful re-establishment of aspen (Populus tremuloides Michx.) on surface-mined lands in the western United States is problematic because the species generally regenerates vegetatively by sprouting from parent roots in the soil; however, topsoil is removed in the mining process. Previous attempts to plant aspen on reclaimed mine sites have failed because...
The effect of aspen harvest and growth on water yield in Minnesota
Elon S. Verry
1987-01-01
Annual water yield increased following the clearcutting of a mature aspen forest in years 1-9 and year 14 of subsequent aspen regrowth. Maximum increases of 85, 117, and 88 mm year-l occurred during the first 3 years of regrowth. Increases in streamflow volumes from snowmelt and early spring rains were minimal and more variable after harvest and...
Structural lumber properties of Wisconsin-5 hybrid poplar : [abstract
David E. Kretschmann; J. G. Isebrands; Glen Stanosz
2000-01-01
Aspen (Populus tremuloides) is an important component of the resource of the Lake States of Michigan, Minnesota, and Wisconsin. In recent years, a concern has surfaced that the aspen cut will exceed its growth. This concern arises from the increased demand placed on the aspen resource by paper, composite, and solid wood industries. To satisfy this demand for forest...
Aspen lumber for building purposes
Louis W. Rees
1947-01-01
Recent shortages of lumber for building purposes make it desirable to seek out all possible supplies of additional lumber. Aspen, according to a study made in northern Minnesota during 1944, may be a source of lumber largely untapped. The results of that study are the main basis of this paper. In the past, aspen has received little consideration as a source of lumber,...
Grizelle Gonzalez; Timothy R. Seastedt; Zugeily Donato
2003-01-01
We compared the abundance and community composition of earthworms, soil macroarthropods, and litter microarthropods to test faunal effects on plant litter decomposition rates in two forests in the subalpine in Colorado, USA. Litterbags containing recently senesced litter of Populus tremuloides (aspen) and Pinus contorta (lodgepole pine) were placed in aspen and pine...
Forest stand structure, productivity, and age mediate climatic effects on aspen decline
Bell, David M.; Bradford, John B.; Lauenroth, William K.
2014-01-01
Because forest stand structure, age, and productivity can mediate the impacts of climate on quaking aspen (Populus tremuloides) mortality, ignoring stand-scale factors limits inference on the drivers of recent sudden aspen decline. Using the proportion of aspen trees that were dead as an index of recent mortality at 841 forest inventory plots, we examined the relationship of this mortality index to forest structure and climate in the Rocky Mountains and Intermountain Western United States. We found that forest structure explained most of the patterns in mortality indices, but that variation in growing-season vapor pressure deficit and winter precipitation over the last 20 years was important. Mortality index sensitivity to precipitation was highest in forests where aspen exhibited high densities, relative basal areas, quadratic mean diameters, and productivities, whereas sensitivity to vapor pressure deficit was highest in young forest stands. These results indicate that the effects of drought on mortality may be mediated by forest stand development, competition with encroaching conifers, and physiological vulnerabilities of large trees to drought. By examining mortality index responses to both forest structure and climate, we show that forest succession cannot be ignored in studies attempting to understand the causes and consequences of sudden aspen decline.
A simple PCR-based marker to determine sex in aspen.
Pakull, B; Kersten, B; Lüneburg, J; Fladung, M
2015-01-01
The genus Populus features a genetically controlled sex determination system, located on chromosome 19. However, different Populus species vary in the position of the sex-linked region on the respective chromosome and the apparent heterogametic sex, and the precise mechanism of sex determination in Populus is still unknown. Using next generation sequencing of pooled samples of male and female aspens, we identified the aspen homologue of the P. trichocarpa gene Potri.019G047300 ('TOZ19') to be male-specific. While in P. tremuloides, the complete gene is missing in the genome of female plants, a short fragment of the 3'-part of the gene is still present in P. tremula females. The male-specific presence and transcription of TOZ19 was further verified using PCR in various different aspen individuals and RT-PCR expression analysis. TOZ19 is potentially involved in early steps of flower development, and represents an interesting candidate gene for involvement in sex determination in aspen. Regardless of its role as candidate gene, TOZ19 represents an ideal marker for determination of the sex of non-flowering aspen individuals or seedlings. © 2014 German Botanical Society and The Royal Botanical Society of the Netherlands.
Predicting nest success from habitat features in aspen forests of the central Rocky Mountains
Heather M. Struempf; Deborah M. Finch; Gregory Hayward; Stanley Anderson
2001-01-01
We collected nesting data on bird use of aspen stands in the Routt and Medicine Bow National Forests between 1987 and 1989. We found active nest sites of 28 species of small nongame birds on nine study plots in undisturbed aspen forests. We compared logistic regression models predicting nest success (at least one nestling) from nest-site or stand-level habitat...
David T. Barnett; Thomas J. Stohlgren
2001-01-01
We investigated aspen (Populus tremuloides) regeneration in the Gros Ventre River Valley, the National Elk Refuge, and a small part of Grand Teton National Park, Wyoming, to see if elk (Cervus elaphus) browsing was as damaging as previously thought. We conducted a landscape-scale survey to assess aspen regeneration across gradients of wintering elk concentrations using...
Growth and mortality of bigtooth aspen trees stressed by defoliation
Donald D. Davis; Timothy M. Frontz
2003-01-01
A survey conducted by the authors in 1993 in six stands within an area of declining aspen revealed that aspen mortality ranged from 25 to 67 percent per stand. Tree-ring analyses revealed that trees dead at time of sampling in 1993 had been growing slower in four of the six stands during the previous decade than were the surviving trees. However growth declines had...
Jing Cai; Melvin T. Tyree
2010-01-01
The objective of this study was to quantify the relationship between vulnerability to cavitation and vessel diameter within a species. We measured vulnerability curves (VCs: percentage loss hydraulic conductivity versus tension) in aspen stems and measured vessel-size distributions. Measurements were done on seed-grown, 4-month-old aspen (Populus tremuloides...
Diane Wagner; Linda DeFoliart; Patricia Doak; Jenny Schneiderheinze
2008-01-01
We studied the effect of epidermal mining on aspen growth and physiology during an outbreak of Phyllocnistis populiella in the boreal forest of interior Alaska. Experimental reduction of leaf miner density across two sites and 3 years significantly increased annual apsen growth rates relative to naturally mined controls. Leaf mining damage was...
Lingli Liu; John S. King; Christian P. Giardina; Fitzgerald L. Booker
2009-01-01
We examined the effects of elevated CO2 and O3 and their interaction on leaf litter chemistry and decomposition in pure stands of aspen (Populus tremuloides) and mixed stands of birch (Betula papyrifera) and aspen at the Aspen Free Air CO2 Enrichment (FACE) experiment. A 935-day in situ incubation...
Heartrot fungi's role in creating picid nesting sites in living aspen
John H. Hart; D. L. Hart
2001-01-01
To determine the number of cavity-containing aspens in old-growth (>80 years), we counted the number of stems containing cavities in 132 0.02-ha plots in Wyoming. There were 8.7 cavities/ha of aspen type. At least 84% of the cavity stems were alive when the initial cavity was constructed; 60% were alive when examined. Fruiting bodies and Phellinus tremulae (a...
Interaction among cervids, fungi, and aspen in northwest Wyoming
John H. Hart; D. L. Hart
2001-01-01
Eighty-five 0.02-ha plots in the Gros Ventre River drainage of northwestern Wyoming with high elk usage had 39% fewer aspen stems in 1985 than in 1970. Sixtyfive of these plots were remeasured in 1989 and 53 additional plots established in 1986 on the Hoback River drainage (lower winter elk usage) were remeasured in 1990. Overall mortality (average/year) of aspen stems...
Stand development of trembling aspen in Canaan Valley, West Virginia
James S. Rentch; James T. Anderson
2008-01-01
In wetlands of Canaan Valley, West Virginia, trembling aspen occurs as a disjunct population well south of its primary natural range. Based on sample data from 15 stands, we found that aspen occurs as nearly monospecific stands or clones. Eight stands had median ages between 30 and 40 yrs, and we suggest that stand initiation was related to changes in land use after...
Effects of partial cutting on diseases, mortality, and regeneration of Rocky Mountain aspen stands
James W. Walters; Thomas E. Hinds; David W. Johnson; Jerome Beatty
1982-01-01
Logging wounds on residual aspen, in partially cut stands, predisposed wounded trees to attack by insects and diseases. Five to 7 years after cutting, aspen mortality amounted to 20%; 41% of the live trees were infected with canker diseases; and 30% were infested with wood borers. Adequate sprouting occurred even though only 60-80% of the basal area was removed.
Paul C. Rogers
2008-01-01
In recent years the aspen research and management community has witnessed increasing accounts of unexplained aspen die-offs across the Rocky Mountain region. In response, two meetings were held to address the issue; this paper summarizes the most recent gathering, a symposium held in Fort Collins at the USDA Forest Service, Rocky Mountain Research Station, on February...
NASA Astrophysics Data System (ADS)
Boyd, M. A.; Walker, X. J.; Rogers, B. M.; Goetz, S. J.; Wagner, D.; Mack, M. C.
2017-12-01
Climate change has increased tree mortality and growth decline in forested ecosystems worldwide. In response to warming and drying of the boreal forest, trembling aspen (Populus tremuloides) has experienced recent large-scale productivity declines. Although declines in productivity are thought to be primarily a result of moistures stress, infestation is another major driver of aspen decline and may interact strongly with climate. Throughout interior Alaska widespread and consistent foliar damage by the aspen epidermal leaf miner Phyllocnistis populiella has been observed concurrent with some of the warmest and driest growing seasons on record. Here we use tree ring measurements and remote sensing indices of vegetation productivity (NDVI) to study the influence of leaf miner and climate on aspen productivity and physiology in the Alaskan boreal forest, and assess if NDVI reflects variations in these ground-based measurements. We assessed ring width and tree ring stable carbon isotope (d13C) response of aspen to infestation and a climate moisture index (CMI) from 2004 - 2014. We found that when growth was negatively correlated to infestation, then it was no longer positively influenced by moisture availability during the growing season. Regardless of the radial growth response to leaf mining, tree ring d13C decreased with increasing infestation. We also found that NDVI was influenced by leaf mining and showed a positive correlation with tree ring d13C, which suggests that NDVI is reflective of changes in tree characteristics under leaf mining that influence tree ring d13C. This finding also reveals the prospect of using satellite data to monitor fluctuations in tree physiology during leaf miner infestation. Our results indicate that aspen productivity will be severely hindered during leaf miner infestation, and that infestation will inhibit the ability of aspen to respond to favorable climate conditions by increasing growth and potentially photosynthesis. This suggests that the productivity, reproduction, and health of aspen in boreal forests, and in turn any related biophysical or carbon sequestration benefits, may become limited under future warming if infestation by leaf miner continues or accelerates.
NASA Technical Reports Server (NTRS)
Badhwar, G. D.; Macdonald, R. B.; Hall, F. G.; Carnes, J. G.
1986-01-01
Results from analysis of a data set of simultaneous measurements of Thematic Mapper band reflectance and leaf area index are presented. The measurements were made over pure stands of Aspen in the Superior National Forest of northern Minnesota. The analysis indicates that the reflectance may be sensitive to the leaf area index of the Aspen early in the season. The sensitivity disappears as the season progresses. Based on the results of model calculations, an explanation for the observed relationship is developed. The model calculations indicate that the sensitivity of the reflectance to the Aspen overstory depends on the amount of understory present.
NASA Technical Reports Server (NTRS)
1981-01-01
The development of a coal gasification system design and mass and energy balance simulation program for the TVA and other similar facilities is described. The materials-process-product model (MPPM) and the advanced system for process engineering (ASPEN) computer program were selected from available steady state and dynamic models. The MPPM was selected to serve as the basis for development of system level design model structure because it provided the capability for process block material and energy balance and high-level systems sizing and costing. The ASPEN simulation serves as the basis for assessing detailed component models for the system design modeling program. The ASPEN components were analyzed to identify particular process blocks and data packages (physical properties) which could be extracted and used in the system design modeling program. While ASPEN physical properties calculation routines are capable of generating physical properties required for process simulation, not all required physical property data are available, and must be user-entered.
Soil variation and sampling intensity under red pine and aspen in Minnesota.
David H. Alban
1974-01-01
In red pine or aspen stands only two soil samples were needed to estimate (+/- 10%, 95% confidence) pH, bulk density, or sand, but 25 to 60 samples were required to estimate N, P, K, Ca, Mg, available water, or silt + clay. To estimate most forest floor properties required 30 to 50 samples in red pine stands, but only about half as many in aspen stands.
Commercial thinning in small-diameter aspen stands in northern Minnesota: study establishment report
Daniel W. Gilmore; Jennifer D. Glenn; Michael E. Ostry; John C. Zasada; Michael A. Benedict
2006-01-01
In the spring of 1999, a long-term study was established to examine the physical and biological aspects of thinning young aspen stands in Minnesota. Three aspen stands ranging in age from 25 to 35 years were selected on lands owned by the State of Minnesota and UPM Kymmene. Two thinning treatments (low and high density) and an unthinned control were installed at each...
Weight-Volume relationships of Aspen and Winter-Cut Black Spruce Pulpwood in Northern Minnesota
David C. Lothner; Richard M. Marden; Edwin Kallio
1974-01-01
Seasonal weight-volume relationships were determined for rough (bark on) aspen and black spruce 100-inch pulpwood that was delivered withing 1 week after cutting in northern Minnesota during 1971-72. For aspen, the weight of wood and bark per cubic foot of wood averaged 56 pounds in the winter and 61 pounds in the summer. This relationshipfor winter-cut black spruce...
Meier, Gretchen A.; Brown, Jesslyn F.; Evelsizer, Ross J.; Vogelmann, James E.
2014-01-01
Trembling aspen (Populus tremuloides Michx.) occurs over wide geographical, latitudinal, elevational, and environmental gradients, making it a favorable candidate for a study of phenology and climate relationships. Aspen forests and woodlands provide numerous ecosystem services, such as high primary productivity and biodiversity, retention and storage of environmental variables (precipitation, temperature, snow–water equivalent) that affect the spring and fall phenology of the aspen woodland communities of southwestern Colorado. We assessed the land surface phenology of aspen woodlands using two phenology indices, start of season time (SOST) and end of season time (EOST), from the U.S. Geological Survey (USGS) database of conterminous U.S. phenological indicators over an 11-year time period (2001–2011). These indicators were developed with 250 m resolution remotely sensed data from the Moderate Resolution Imaging Spectroradiometer processed to highlight vegetation response. We compiled data on SOST, EOST, elevation, precipitation, air temperature, and snow water equivalent (SWE) for selected sites having more than 80% cover by aspen woodland communities. In the 11-year time frame of our study, EOST had significant positive correlation with minimum fall temperature and significant negative correlation with fall precipitation. SOST had a significant positive correlation with spring SWE and spring maximum temperature.
Defensive effects of extrafloral nectaries in quaking aspen differ with scale.
Mortensen, Brent; Wagner, Diane; Doak, Patricia
2011-04-01
The effects of plant defenses on herbivory can differ among spatial scales. This may be particularly common with indirect defenses, such as extrafloral nectaries (EFNs), that attract predatory arthropods and are dependent on predator distribution, abundance, and behavior. We tested the defensive effects of EFNs in quaking aspen (Populus tremuloides Michx.) against damage by a specialist herbivore, the aspen leaf miner (Phyllocnistis populiella Cham.), at the scale of individual leaves and entire ramets (i.e., stems). Experiments excluding crawling arthropods revealed that the effects of aspen EFNs differed at the leaf and ramet scales. Crawling predators caused similar reductions in the percent leaf area mined on individual leaves with and without EFNs. However, the extent to which crawling predators increased leaf miner mortality and, consequently, reduced mining damage increased with EFN expression at the ramet scale. Thus, aspen EFNs provided a diffuse defense, reducing damage to leaves across a ramet regardless of leaf-scale EFN expression. We detected lower leaf miner damage and survival unassociated with crawling predators on EFN-bearing leaves, suggesting that direct defenses (e.g., chemical defenses) were stronger on leaves with than without EFNs. Greater direct defenses on EFN-bearing leaves may reduce the probability of losing these leaves and thus weakening ramet-scale EFN defense. Aspen growth was not related to EFN expression or the presence of crawling predators over the course of a single season. Different effects of aspen EFNs at the leaf and ramet scales suggest that future studies may benefit from examining indirect defenses simultaneously at multiple scales.
Wooley, Stuart C.; Donaldson, Jack R.; Gusse, Adam C.; Lindroth, Richard L.; Stevens, Michael T.
2007-01-01
Background and Aims A wide variety of plants produce extrafloral nectaries (EFNs) that are visited by predatory arthropods. But very few studies have investigated the relationship between plant genetic variation and EFNs. The presence of foliar EFNs is highly variable among different aspen (Populus tremuloides) genotypes and the EFNs are visited by parasitic wasps and predatory flies. The aim here was to determine the heritability of EFNs among aspen genotypes and age classes, possible trade-offs between direct and indirect defences, EFN induction following herbivory, and the relationship between EFNs and predatory insects. Methods EFN density was quantified among aspen genotypes in Wisconsin on trees of different ages and broad-sense heritability from common garden trees was calculated. EFNs were also quantified in natural aspen stands in Utah. From the common garden trees foliar defensive chemical levels were quantified to evaluate their relationship with EFN density. A defoliation experiment was performed to determine if EFNs can be induced in response to herbivory. Finally, predatory arthropod abundance among aspen trees was quantified to determine the relationship between arthropod abundance and EFNs. Key Results Broad-sense heritability for expression (0·74–0·82) and induction (0·85) of EFNs was high. One-year-old trees had 20% greater EFN density than 4-year-old trees and more than 50% greater EFN density than ≥10-year-old trees. No trade-offs were found between foliar chemical concentrations and EFN density. Predatory fly abundance varied among aspen genotypes, but predatory arthropod abundance and average EFN density were not related. Conclusions Aspen extrafloral nectaries are strongly genetically determined and have the potential to respond rapidly to evolutionary forces. The pattern of EFN expression among different age classes of trees appears to follow predictions of optimal defence theory. The relationship between EFNs and predators likely varies in relation to multiple temporal and environmental factors. PMID:17951361
Kauffman, Matthew J.; Brodie, Jedediah F.; Jules, Erik S.
2010-01-01
Behaviorally mediated trophic cascades (BMTCs) occur when the fear of predation among herbivores enhances plant productivity. Based primarily on systems involving small-bodied predators, BMTCs have been proposed as both strong and ubiquitous in natural ecosystems. Recently, however, synthetic work has suggested that the existence of BMTCs may be mediated by predator hunting mode, whereby passive (sit-and-wait) predators have much stronger effects than active (coursing) predators. One BMTC that has been proposed for a wide-ranging active predator system involves the reintroduction of wolves (Canis lupus) to Yellowstone National Park, USA, which is thought to be leading to a recovery of trembling aspen (Populus tremuloides) by causing elk (Cervus elaphus) to avoid foraging in risky areas. Although this BMTC has been generally accepted and highly popularized, it has never been adequately tested. We assessed whether wolves influence aspen by obtaining detailed demographic data on aspen stands using tree rings and by monitoring browsing levels in experimental elk exclosures arrayed across a gradient of predation risk for three years. Our study demonstrates that the historical failure of aspen to regenerate varied widely among stands (last recruitment year ranged from 1892 to 1956), and our data do not indicate an abrupt cessation of recruitment. This pattern of recruitment failure appears more consistent with a gradual increase in elk numbers rather than a rapid behavioral shift in elk foraging following wolf extirpation. In addition, our estimates of relative survivorship of young browsable aspen indicate that aspen are not currently recovering in Yellowstone, even in the presence of a large wolf population. Finally, in an experimental test of the BMTC hypothesis we found that the impacts of elk browsing on aspen demography are not diminished in sites where elk are at higher risk of predation by wolves. These findings suggest the need to further evaluate how trophic cascades are mediated by predator–prey life history and ecological context.
Kauffman, Matthew J; Brodie, Jedediah F; Jules, Erik S
2010-09-01
Behaviorally mediated trophic cascades (BMTCs) occur when the fear of predation among herbivores enhances plant productivity. Based primarily on systems involving small-bodied predators, BMTCs have been proposed as both strong and ubiquitous in natural ecosystems. Recently, however, synthetic work has suggested that the existence of BMTCs may be mediated by predator hunting mode, whereby passive (sit-and-wait) predators have much stronger effects than active (coursing) predators. One BMTC that has been proposed for a wide-ranging active predator system involves the reintroduction of wolves (Canis lupus) to Yellowstone National Park, USA, which is thought to be leading to a recovery of trembling aspen (Populus tremuloides) by causing elk (Cervus elaphus) to avoid foraging in risky areas. Although this BMTC has been generally accepted and highly popularized, it has never been adequately tested. We assessed whether wolves influence aspen by obtaining detailed demographic data on aspen Stands using tree rings and by monitoring browsing levels in experimental elk exclosures arrayed across a gradient of predation risk for three years. Our study demonstrates that the historical failure of aspen to regenerate varied widely among stands (last recruitment year ranged from 1892 to 1956), and our data do not indicate an abrupt cessation of recruitment. This pattern of recruitment failure appears more consistent with a gradual increase in elk numbers rather than a rapid behavioral shift in elk foraging following wolf extirpation. In addition, our estimates of relative survivorship of young browsable aspen indicate that aspen are not currently recovering in Yellowstone, even in the presence of a large wolf population. Finally, in an experimental test of the BMTC hypothesis we found that the impacts of elk browsing on aspen demography are not diminished in sites where elk are at higher risk of predation by wolves. These findings suggest the need to further evaluate how trophic cascades are mediated by predator-prey life history and ecological context.
1986-07-01
to the fall diet (Beer 1948, Hungerford 1957, Phillips 1967, McGowan 1973). Hazelnut, chokecherry , and juneberry supplement aspen and other foods...bigtooth aspen (Svoboda and Gullion 1972). Chokecherry is a locally preferred winter food (Phillips 1967). Nutrition. Korschgen (1966) determined...grandidentata Quaking aspen P. tremuloide8 Beech Fags app. Birch Betula spp. Blackberry Rubus app. Cherry Prunus spp. Chokecherry P. virginiana
ERIC Educational Resources Information Center
Bollier, David
The 1994 Aspen Institute Roundtable on Information Technology began as a look at the changing nature of the home. In building scenarios of the "new home," the participants expressed many significant insights into issues of personal identity, community-building, and setting boundaries in our lives and environments. This report captures…
Rakesh Minocha; Stephanie Long; Subhash Minocha; Paula Marquardt; Neil Nelson; Mark. Kubiske
2010-01-01
This study was conducted at the Aspen Free-Air Carbon Dioxide Enrichment (FACE) experimental site, Rhinelander, WI, (USA). Since 1998, 12 experimental rings planted in 1997 underwent four different treatments: control; elevated CO2 (560 ppm); elevated O3 (1.5X ambient) and elevated CO2 (560 ppm) + O...
Clinical and Molecular Consequences of NF1 Microdeletion
2005-05-01
and Cell Biology of NFl, NF2 and Schwannomatosis , Aspen, June, 2005 "* Abstract: Shinohara MM, Kuechle MK, Graves J, Stephens K. Neurofibromin is a...caspase target. The CTF International Consortium for the Molecular and Cell Biology of NF1, NF2 and Schwannomatosis , Aspen, June, 2005. "* Abstract...Molecular and Cell Biology of NF1, NF2 and Schwannomatosis , Aspen, June, 2005 Abstract: Shinohara MM, Kuechle MK, Graves J, Stephens K. Neurofibromin is a
Role of Volatility in the Development of JP-8 Surrogates for Diesel Engine Application
2014-01-01
distillation curves of the surrogate fuels were calculated using the Aspen HYSYS [41] software package, and the Peng- Robinson model was chosen to...distillation curves for the surrogate fuels developed in this investigation, the accuracy of Aspen HYSYS software predictions were compared with...and SF3. The distillation curves calculated using Aspen HYSYS software for the five surrogate fuels of Table 1 are shown in Figure 7, along with the
ERIC Educational Resources Information Center
Bollier, David
This document is an interpretive synthesis of the discussion at a conference sponsored by the Aspen Institute that sought to develop new ways to understand how the Internet is changing the powers of the nation-state, the conduct of international relations, and the definitions of nation security. This report examines how the Internet and other…
Updraft Fixed Bed Gasification Aspen Plus Model
DOE Office of Scientific and Technical Information (OSTI.GOV)
2007-09-27
The updraft fixed bed gasification model provides predictive modeling capabilities for updraft fixed bed gasifiers, when devolatilization data is available. The fixed bed model is constructed using Aspen Plus, process modeling software, coupled with a FORTRAN user kinetic subroutine. Current updraft gasification models created in Aspen Plus have limited predictive capabilities and must be "tuned" to reflect a generalized gas composition as specified in literature or by the gasifier manufacturer. This limits the applicability of the process model.
America Promises to Come Back: A New National Strategy
1991-08-05
unveiled in Aspen, Colorado on August 2, 1990, involving a mix of active, reserve, and reconstitutable forces, and General Colin Powell’s "base" force...are vital for the reintroduction of formidable American military power overseas. There is a risk that the end of the Cold War may bring an increased...Winter 1991/1992. (2) "Remarks by the President to the Aspen Institute Symposium" (as delivered), Office of the Press Secretary (Aspen, Colorado ), The
2009-01-01
Schwannomatosis Aspen, Colorado, 2006. (appended Item 1) 2. Crotti TN, Walsh NC, Barnes GL, Gerstenfeld LC, McHugh KP. Neurofibromin expression...Tumor Foundation, International Consortium for the Molecular and Cellular Biology of NF1, NF2, and Schwannomatosis , Aspen, Colorado, 2006...and Cellular Biology of NF1, NF2, and Schwannomatosis , Aspen, Colorado, 2006. A large proportion of patients with Neurofibromatosis Type 1
Influence of elk grazing on soil properties in Rocky Mountain National Park
Binkley, Dan; Singer, F.; Kaye, M.; Rochelle, R.
2003-01-01
We used three 35-year exclosures to examine the effects of high elk populations on a variety of soil properties in three vegetation types: upland sagebrush, aspen, and meadow. Grazing and hoof action by elk significantly increased bulk density (from 0.87 kg/l ungrazed to 0.94 kg/l grazed), with greater effects on soils with fewer rocks. Grazing substantially reduced extractable calcium, magnesium, potassium and phosphorus in the sagebrush type, but not in the aspen or meadow types. The only grazing effect on pH came in aspen types, where grazing prevented aspen establishment, and kept soil pH about 0.7 units higher than under aspen inside the exclosures. Grazing had no overall effect on total soil C and N across all exclosures and vegetation types. The availability of soil nitrogen, indexed by in-field resin bags and net mineralization in soil cores, showed little overall effect of grazing. Limited data on soil leaching indicated a possibility of strong increases in nitrate leaching with grazing for an aspen vegetation type at one exclosure. Although we found little effect of grazing on soil N supply, we note that N fertilization doubled the production of grasses and shrubs; if grazing eventually led to changes in soil N supply, species composition and growth would likely change. ?? 2003 Elsevier B.V. All rights reserved.
Larsen, Peter E; Cseke, Leland J; Miller, R Michael; Collart, Frank R
2014-10-21
Rising atmospheric levels of carbon dioxide and ozone will impact productivity and carbon sequestration in forest ecosystems. The scale of this process and the potential economic consequences provide an incentive for the development of models to predict the types and rates of ecosystem responses and feedbacks that result from and influence of climate change. In this paper, we use phenotypic and molecular data derived from the Aspen Free Air CO2 Enrichment site (Aspen-FACE) to evaluate modeling approaches for ecosystem responses to changing conditions. At FACE, it was observed that different aspen clones exhibit clone-specific responses to elevated atmospheric levels of carbon dioxide and ozone. To identify the molecular basis for these observations, we used artificial neural networks (ANN) to examine above and below-ground community phenotype responses to elevated carbon dioxide, elevated ozone and gene expression profiles. The aspen community models generated using this approach identified specific genes and subnetworks of genes associated with variable sensitivities for aspen clones. The ANN model also predicts specific co-regulated gene clusters associated with differential sensitivity to elevated carbon dioxide and ozone in aspen species. The results suggest ANN is an effective approach to predict relevant gene expression changes resulting from environmental perturbation and provides useful information for the rational design of future biological experiments. Copyright © 2014 Elsevier Ltd. All rights reserved.
Kawaoka, Akiyoshi; Matsunaga, Etsuko; Endo, Saori; Kondo, Shinkichi; Yoshida, Kazuya; Shinmyo, Atsuhiko; Ebinuma, Hiroyasu
2003-01-01
We previously demonstrated that overexpression of the horseradish (Armoracia rusticana) peroxidase prxC1a gene stimulated the growth rate of tobacco (Nicotiana tabacum) plants. Here, the cauliflower mosaic virus 35S::prxC1a construct was introduced into hybrid aspen (Populus sieboldii × Populus grandidentata). The growth rate of these transformed hybrid aspen plants was substantially increased under greenhouse conditions. The average stem length of transformed plants was 25% greater than that of control plants. There was no other obvious phenotypic difference between the transformed and control plants. Fast-growing transformed hybrid aspen showed high levels of expression of prxC1a and had elevated peroxidase activities toward guaiacol and ascorbate. However, there was no increase of the endogenous class I ascorbate peroxidase activities in the transformed plants by separate assay and activity staining of native polyacrylamide gel electrophoresis. Furthermore, calli derived from the transformed hybrid aspen grew faster than those from control plants and were resistant to the oxidative stress imposed by hydrogen peroxide. Therefore, enhanced peroxidase activity affects plant growth rate and oxidative stress resistance. PMID:12857800
Kawaoka, Akiyoshi; Matsunaga, Etsuko; Endo, Saori; Kondo, Shinkichi; Yoshida, Kazuya; Shinmyo, Atsuhiko; Ebinuma, Hiroyasu
2003-07-01
We previously demonstrated that overexpression of the horseradish (Armoracia rusticana) peroxidase prxC1a gene stimulated the growth rate of tobacco (Nicotiana tabacum) plants. Here, the cauliflower mosaic virus 35S::prxC1a construct was introduced into hybrid aspen (Populus sieboldii x Populus grandidentata). The growth rate of these transformed hybrid aspen plants was substantially increased under greenhouse conditions. The average stem length of transformed plants was 25% greater than that of control plants. There was no other obvious phenotypic difference between the transformed and control plants. Fast-growing transformed hybrid aspen showed high levels of expression of prxC1a and had elevated peroxidase activities toward guaiacol and ascorbate. However, there was no increase of the endogenous class I ascorbate peroxidase activities in the transformed plants by separate assay and activity staining of native polyacrylamide gel electrophoresis. Furthermore, calli derived from the transformed hybrid aspen grew faster than those from control plants and were resistant to the oxidative stress imposed by hydrogen peroxide. Therefore, enhanced peroxidase activity affects plant growth rate and oxidative stress resistance.
New dimension analyses with error analysis for quaking aspen and black spruce
NASA Technical Reports Server (NTRS)
Woods, K. D.; Botkin, D. B.; Feiveson, A. H.
1987-01-01
Dimension analysis for black spruce in wetland stands and trembling aspen are reported, including new approaches in error analysis. Biomass estimates for sacrificed trees have standard errors of 1 to 3%; standard errors for leaf areas are 10 to 20%. Bole biomass estimation accounts for most of the error for biomass, while estimation of branch characteristics and area/weight ratios accounts for the leaf area error. Error analysis provides insight for cost effective design of future analyses. Predictive equations for biomass and leaf area, with empirically derived estimators of prediction error, are given. Systematic prediction errors for small aspen trees and for leaf area of spruce from different site-types suggest a need for different predictive models within species. Predictive equations are compared with published equations; significant differences may be due to species responses to regional or site differences. Proportional contributions of component biomass in aspen change in ways related to tree size and stand development. Spruce maintains comparatively constant proportions with size, but shows changes corresponding to site. This suggests greater morphological plasticity of aspen and significance for spruce of nutrient conditions.
BIOASPEN: System for technology development
NASA Technical Reports Server (NTRS)
1986-01-01
The public version of ASPEN was installed in the VAX 11/750 computer. To examine the idea of BIOASPEN, a test example (the manufacture of acetone, butanol, and ethanol through a biological route) was chosen for simulation. Previous reports on the BIOASPEN project revealed the limitations of ASPEN in modeling this process. To overcome some of the difficulties, modules were written for the acid and enzyme hydrolyzers, the fermentor, and a sterilizer. Information required for these modules was obtained from the literature whenever possible. Additional support modules necessary for interfacing with ASPEN were also written. Some of ASPEN subroutines were themselves altered in order to ensure the correct running of the simulation program. After testing of these additions and charges was completed, the Acetone-Butanol-Ethanol (ABE) process was simulated. A release of ASPEN (which contained the Economic Subsystem) was obtained and installed. This subsection was tested and numerous charges were made in the FORTRAN code. Capital investment and operating cost studies were performed on the ABE process. Some alternatives in certain steps of the ABE simulation were investigated in order to elucidate their effects on the overall economics of the process.
Thinning strategies for aspen: a prediction model.
Donald A. Perala
1978-01-01
Derives thinning strategies to maximize volume yields of aspen fiber, sawtimber, and veneer. Demonstrates how yields are affected by growing season climatic variation and periodic defoliation by forest tent caterpillar.
Cross-Grain Knife Planing Improves Surface Quality and Utilization of Aspen
Harold A. Stewart
1971-01-01
Aspen at 6 percent moisture content was planed parallel to the grain and across the grain on a cabinet planer with a 25? rake angle, 1/16- and 1/32-inch depth of cut, and 20 knife marks per inch. Aspen was also cross-grain knife planed with a 45? rake angle, 1/32-, 1/16-, and 1/8-inch depths of cut, and 20, 10, 5, and 2.5 knife marks per inch. Cross-grain knife...
American Society for Parenteral & Enteral Nutrition
... Center Advertising and Sponsorship Learn More ASPEN Enteral Nutrition by the Numbers: EN Data Across the Healthcare Continuum Learn More The ASPEN Adult Nutrition Support Core Curriculum, 3rd Edition Has Arrived! The ...
USDA FS
1972-01-01
Contains 25 technical papers on the aspen resource and its potential, inventory projections, economics, wood and fiber products, silvics and ecology, diseases and insects, growth and yield, and various aspects of management in the Lake States and Canada.
Simulating economic effects of disruptions in the telecommunications infrastructure.
DOE Office of Scientific and Technical Information (OSTI.GOV)
Cox, Roger Gary; Barton, Dianne Catherine; Reinert, Rhonda K.
2004-01-01
CommAspen is a new agent-based model for simulating the interdependent effects of market decisions and disruptions in the telecommunications infrastructure on other critical infrastructures in the U.S. economy such as banking and finance, and electric power. CommAspen extends and modifies the capabilities of Aspen-EE, an agent-based model previously developed by Sandia National Laboratories to analyze the interdependencies between the electric power system and other critical infrastructures. CommAspen has been tested on a series of scenarios in which the communications network has been disrupted, due to congestion and outages. Analysis of the scenario results indicates that communications networks simulated by themore » model behave as their counterparts do in the real world. Results also show that the model could be used to analyze the economic impact of communications congestion and outages.« less
NASA Astrophysics Data System (ADS)
Janaun, J.; Kamin, N. H.; Wong, K. H.; Tham, H. J.; Kong, V. V.; Farajpourlar, M.
2016-06-01
Air heating unit is one of the most important parts in paddy drying to ensure the efficiency of a drying process. In addition, an optimized air heating unit does not only promise a good paddy quality, but also save more for the operating cost. This study determined the suitable and best specifications heating unit to heat air for paddy drying in the LAMB dryer. In this study, Aspen HYSYS v7.3 was used to obtain the minimum flow rate of hot water needed. The resulting data obtained from Aspen HYSYS v7.3 were used in Aspen Exchanger Design and Rating (EDR) to generate heat exchanger design and costs. The designs include shell and tubes and plate heat exchanger. The heat exchanger was designed in order to produce various drying temperatures of 40, 50, 60 and 70°C of air with different flow rate, 300, 2500 and 5000 LPM. The optimum condition for the heat exchanger were found to be plate heat exchanger with 0.6 mm plate thickness, 198.75 mm plate width, 554.8 mm plate length and 11 numbers of plates operating at 5000 LPM air flow rate.
DOE Office of Scientific and Technical Information (OSTI.GOV)
Tuskan, Gerald A; Yin, Tongming
Lack of aspen (Populus tremuloides) recruitment and canopy replacement of aspen stands that grow on the edges of grasslands on the low-elevation elk (Cervus elaphus) winter range of Rocky Mountain National Park (RMNP) in Colorado has been a cause of concern for more than 70 years (Packard, 1942; Olmsted, 1979; Stevens, 1980; Hess, 1993; R.J. Monello, T.L. Johnson, and R.G. Wright, Rocky Mountain National Park, 2006, written commun.). These aspen stands are a significant resource since they are located close to the park's road system and thus are highly visible to park visitors. Aspen communities are integral to the ecologicalmore » structure of montane and subalpine landscapes because they contain high native species richness of plants, birds, and butterflies (Chong and others, 2001; Simonson and others, 2001; Chong and Stohlgren, 2007). These low-elevation, winter range stands also represent a unique component of the park's plant community diversity since most (more than 95 percent) of the park's aspen stands grow in coniferous forest, often on sheltered slopes and at higher elevations, while these winter range stands are situated on the low-elevation ecotone between the winter range grasslands and some of the park's drier coniferous forests.« less
Best Practices Case Study: Shaw Construction Burlingame Ranch Ph.1, Aspen, CO
DOE Office of Scientific and Technical Information (OSTI.GOV)
Pacific Northwest National Laboratory & Oak Ridge National Laboratory
Shaw Construction built 84 energy efficient, affordable condominiums forthe City of Aspen that achieved HERS scores of less than 62 with help from Building America’s research team lead Building Science Corporation.
BOREAS TE-10 Leaf Optical Properties
NASA Technical Reports Server (NTRS)
Hall, Forrest G. (Editor); Papagno, Andrea (Editor); Chan, Stephen S.; Middleton, Elizabeth
2000-01-01
The Boreal Ecosystem-Atmospheric Study (BOREAS) TE-10 (Terrestrial Ecology) team collected several data sets in support of its efforts to characterize and interpret information on the reflectance, transmittance, gas exchange, oxygen evolution, and biochemical properties of boreal vegetation. This data set describes the spectral optical properties (reflectance and transmittance) of boreal forest conifers and broadleaf tree leaves as measured with a Spectron Engineering SE590 spectroradiometer at the Southern Study Area Old Black Spruce (SSA OBS), Old Jack Pine (OJP), Young Jack Pine (YJP), Old Aspen (OA), Old Aspen Auxiliary (OA-AUX), Young Aspen Auxiliary (YA-AUX), and Young Aspen (YA) sites. The data were collected during the growing seasons of 1994 and 1996 and are stored in tabular ASCII files. The data files are available on a CD-ROM (see document number 20010000884), or from the Oak Ridge National Laboratory (ORNL) Distributed Active Archive Center (DAAC).
DOE Office of Scientific and Technical Information (OSTI.GOV)
Basu, N.; Pryor, R.J.
1997-09-01
This report presents a microsimulation model of a transition economy. Transition is defined as the process of moving from a state-enterprise economy to a market economy. The emphasis is on growing a market economy starting from basic microprinciples. The model described in this report extends and modifies the capabilities of Aspen, a new agent-based model that is being developed at Sandia National Laboratories on a massively parallel Paragon computer. Aspen is significantly different from traditional models of the economy. Aspen`s emphasis on disequilibrium growth paths, its analysis based on evolution and emergent behavior rather than on a mechanistic view ofmore » society, and its use of learning algorithms to simulate the behavior of some agents rather than an assumption of perfect rationality make this model well-suited for analyzing economic variables of interest from transition economies. Preliminary results from several runs of the model are included.« less
Phenolic Glycosides in Populus tremuloides and their Effects on Long-Term Ungulate Browsing.
Lastra, R A; Kenkel, N C; Daayf, F
2017-10-01
In the aspen-grassland ecotone of Riding Mountain, Manitoba, lightly browsed vigorous clones of trembling aspen (Populus tremuloides Michx.) occur in close proximity to heavily browsed dieback clones. This study examines whether intraspecific variation in the production of phenolic glycosides is correlated with this strong dichotomy in clonal vigor. Individual clones were sampled over four years at three sites located along a gradient of increasing soil moisture stress. At each site, eight aspen clones of similar size and age were sampled: four vigorous and four dieback clones (total of 24 individual clones). The severity of wapiti (elk) browsing was assessed as the ratio of browse-damaged to total branches per aspen ramet. Statistically significant differences in foliar concentrations of the phenolic glycosides salicortin and tremulacin were observed between vigorous and dieback clones: a mean of 14.8% dry mass for lightly browsed (vigorous) clones, versus just 7.0% for heavily browsed (dieback) clones. Mean concentrations of foliar phenolics were also significantly greater in more moisture-stressed sites. These results demonstrate that the strong dichotomy in clonal vigor (vigorous versus dieback clones) is associated with large differences in phenolic glycoside production. Vigorous clone ramets produce high amounts of phenolic glycosides and have low levels of herbivore browsing and low mortality rates, whereas dieback clone ramets have low amounts of phenolic glycosides and much higher herbivore browsing and mortality rates. This suggests that intraspecific variation in phenolic glycosides in trembling aspen is an important predisposing factor leading to ramet mortality, and by extension to the decline of aspen clones.
Elevated CO2 response of photosynthesis depends on ozone concentration in aspen
Asko Noormets; Olevi Kull; Anu Sober; Mark E. Kubiske; David F. Karnosky
2010-01-01
The effect of elevated CO2 and O3 on apparent quantum yield (ø), maximum photosynthesis (Pmax), carboxylation efficiency (Vcmax) and electron transport capacity (Jmax) at different canopy locations was studied in two aspen (Populus...
Using glyphosate herbicide in converting aspen to conifers.
Donald A. Perala
1985-01-01
Glyphosate at 1.5 to 2.0 lbs. Per acre active ingredient will control aspen suckers, shrubs, and herbs when applied during August to early September. There appears to be some intersite variability in the efficacy of the herbicide.
Manager's handbook for aspen in the north-central states.
Donald A. Perala
1977-01-01
Summarizes information on silvicultural practices to improve yields of timber, water, and wildlife, while minimizing unsightly manipulation of the landscape, for the aspen forest type. A management key outlines recommendations for given stand conditions and management objectives.
Aspen Modeling of the Bayer Process
NASA Astrophysics Data System (ADS)
Langa, J. M.; Russell, T. G.; O'Neill, G. A.; Gacka, P.; Shah, V. B.; Stephenson, J. L.; Snyder, J. G.
The ASPEN simulator was used to model Alcoa's Pt. Comfort Bayer refinery. All areas of the refinery including the lakes and powerhouse were modeled. Each area model was designed to be run stand alone or integrated with others for a full plant model.
Estimating aspen volume and weight for individual trees, diameter classes, or entire stands.
Bryce E. Schlaegel
1975-01-01
Presents allometric volume and weight equations for Minnesota quaking aspen. Volume, green weight, and dry weight estimates can be made for wood, bark, and limbs on the basis of individual trees, diameter classes, or entire stands.
Effects of elevated CO2 and ozone on phenolic glycosides of trembling aspen
James K. Nitao; Muraleedharan G. Nair; William J. Mattson; Daniel A. Herms; Bruce A. Birr; Mark D. Coleman; Terry M. Trier; J. G. Isebrands
1996-01-01
We tested the effects of elevated CO2 and ozone on concentrations of the phenolic glycosides salicortin and tremulacin in immature and mature foliage of the trembling aspen (Populus tremuloides) clones 216, 259, and 271.
Imaging spectroscopy links aspen genotype with below-ground processes at landscape scales
Madritch, Michael D.; Kingdon, Clayton C.; Singh, Aditya; Mock, Karen E.; Lindroth, Richard L.; Townsend, Philip A.
2014-01-01
Fine-scale biodiversity is increasingly recognized as important to ecosystem-level processes. Remote sensing technologies have great potential to estimate both biodiversity and ecosystem function over large spatial scales. Here, we demonstrate the capacity of imaging spectroscopy to discriminate among genotypes of Populus tremuloides (trembling aspen), one of the most genetically diverse and widespread forest species in North America. We combine imaging spectroscopy (AVIRIS) data with genetic, phytochemical, microbial and biogeochemical data to determine how intraspecific plant genetic variation influences below-ground processes at landscape scales. We demonstrate that both canopy chemistry and below-ground processes vary over large spatial scales (continental) according to aspen genotype. Imaging spectrometer data distinguish aspen genotypes through variation in canopy spectral signature. In addition, foliar spectral variation correlates well with variation in canopy chemistry, especially condensed tannins. Variation in aspen canopy chemistry, in turn, is correlated with variation in below-ground processes. Variation in spectra also correlates well with variation in soil traits. These findings indicate that forest tree species can create spatial mosaics of ecosystem functioning across large spatial scales and that these patterns can be quantified via remote sensing techniques. Moreover, they demonstrate the utility of using optical properties as proxies for fine-scale measurements of biodiversity over large spatial scales. PMID:24733949
Final Harvest of Above-Ground Biomass and Allometric Analysis of the Aspen FACE Experiment
DOE Office of Scientific and Technical Information (OSTI.GOV)
Mark E. Kubiske
The Aspen FACE experiment, located at the US Forest Service Harshaw Research Facility in Oneida County, Wisconsin, exposes the intact canopies of model trembling aspen forests to increased concentrations of atmospheric CO2 and O3. The first full year of treatments was 1998 and final year of elevated CO2 and O3 treatments is scheduled for 2009. This proposal is to conduct an intensive, analytical harvest of the above-ground parts of 24 trees from each of the 12, 30 m diameter treatment plots (total of 288 trees) during June, July & August 2009. This above-ground harvest will be carefully coordinated with themore » below-ground harvest proposed by D.F. Karnosky et al. (2008 proposal to DOE). We propose to dissect harvested trees according to annual height growth increment and organ (main stem, branch orders, and leaves) for calculation of above-ground biomass production and allometric comparisons among aspen clones, species, and treatments. Additionally, we will collect fine root samples for DNA fingerprinting to quantify biomass production of individual aspen clones. This work will produce a thorough characterization of above-ground tree and stand growth and allocation above ground, and, in conjunction with the below ground harvest, total tree and stand biomass production, allocation, and allometry.« less
NASA Astrophysics Data System (ADS)
Hogg, E.
2009-05-01
In western Canada, the boundary between boreal forest and prairie grasslands marks a dramatic change in nearly all aspects of ecosystem functioning. These include a steep spatial gradient in hydrological characteristics of the landscape (lake level variability, water runoff and stream flow patterns) that coincides with the southern range limit of peatlands and several species of boreal conifers. Previous studies indicate that the forest-grassland boundary in this region represents a critical "tipping point" (Lenton et al. 2008) where long-term water input by precipitation is barely sufficient to satisfy the water use demands of productive, closed-canopy forests. This concept is consistent with the observed, regional gradient in the character of forests dominated by aspen (Populus tremuloides), the most abundant and widespread deciduous tree in North America. Aspen-dominated forests are productive and continuous in the boreal zone, but are stunted and patchy in the boreal-grassland transition zone, often referred to as the aspen parkland. Based on the "tipping point" concept, there are concerns that aspen forests in this region are especially sensitive to the projected trend toward warmer and drier conditions under human-induced climate change. In response to these concerns, a large-scale study was established across west-central Canada in 2000, entitled "Climate Impacts on Productivity and Health of Aspen" (CIPHA). The study has hierarchical sampling design that is aimed at "scaling up" forest-climate responses from individual trees to the region. During 2001-2002, the region was affected by an exceptionally severe drought that subsequently led to massive dieback and mortality of aspen forests within the boreal-grassland transition zone. Drought severity and extent was quantified using a simple climate moisture index (CMI), and drought impacts were quantified using tree-ring analysis, in combination with plot-based and remotely-sensed measures. Results showed that stand-level productivity, dieback and mortality were governed primarily by moisture variation. Furthermore, during and following this drought there was increasing damage by wood-boring insects and elevated, regional-scale mortality of aspen over at least 6 years (2002-2008). Although it is premature to attribute these impacts to anthropogenic climate change, they provide an excellent analog for what may be expected in future, even under a modest trend toward drying over the next few decades. Furthermore, the recent aspen mortality in western Canada shares many features common to other recent episodes of drought-induced forest mortality that have been documented on all of the earth's forested continents. This suggests the need for an integrated, global research and monitoring system that would enable early detection and attribution of large-scale ecosystem changes, especially in climatically-sensitive regions along forest-grassland boundaries around the world.
Translating MAPGEN to ASPEN for MER
NASA Technical Reports Server (NTRS)
Rabideau, Gregg R.; Knight, Russell L.; Lenda, Matthew; Maldague, Pierre F.
2013-01-01
This software translates MAPGEN (Europa and APGEN) domains to ASPEN, and the resulting domain can be used to perform planning for the Mars Exploration Rover (MER). In other words, this is a conversion of two distinct planning languages (both declarative and procedural) to a third (declarative) planning language in order to solve the problem of faithful translation from mixed-domain representations into the ASPEN Modeling Language. The MAPGEN planning system is an example of a hybrid procedural/declarative system where the advantages of each are leveraged to produce an effective planner/scheduler for MER tactical planning. The adaptation of the planning system (ASPEN) was investigated, and, with some translation, much of the procedural knowledge encoding is amenable to declarative knowledge encoding. The approach was to compose translators from the core languages used for adapting MAGPEN, which consists of Europa and APGEN. Europa is a constraint- based planner/scheduler where domains are encoded using a declarative model. APGEN is also constraint-based, in that it tracks constraints on resources and states and other variables. Domains are encoded in both constraints and code snippets that execute according to a forward sweep through the plan. Europa and APGEN communicate to each other using proxy activities in APGEN that represent constraints and/or tokens in Europa. The composition of a translator from Europa to ASPEN was fairly straightforward, as ASPEN is also a declarative planning system, and the specific uses of Europa for the MER domain matched ASPEN s native encoding fairly closely. On the other hand, translating from APGEN to ASPEN was considerably more involved. On the surface, the types of activities and resources one encodes in APGEN appear to match oneto- one to the activities, state variables, and resources in ASPEN. But, when looking into the definitions of how resources are profiled and activities are expanded, one sees code snippets that access various information available during planning for the moment in time being planned to decide at the time what the appropriate profile or expansion is. APGEN is actually a forward (in time) sweeping discrete event simulator, where the model is composed of code snippets that are artfully interleaved by the engine to produce a plan/schedule. To solve this problem, representative code is simulated as a declarative series of task expansions. Predominantly, three types of procedural models were translated: loops, if statements, and code blocks. Loops and if statements were handled using controlled task expansion, and code blocks were handled using constraint networks that maintained the generation of results based on what the order of execution would be for a procedural representation. One advantage with respect to performance for MAPGEN is the use of APGEN s GUI. This GUI is written in C++ and Motif, and performs very well for large plans.
Eugene M. Wengert; Dennis M. Donnelly; Donald C. Markstrom; Harold E. Worth
1985-01-01
In the past, markets for quaking aspen timber from the Rocky Mountains have been insufficient to support significant harvesting. This shortage of markets severely restrained the potential for aspen management. As a result, many stands protected from wildfire gradually reverted to conifers (see the VEGETATIVE REGENERATION and FIRE chapters).
Thinning Pole-Sized Aspen Has no Effect on Number of Veneer Trees or Total Yield
Bryce E. Schlaegel; Stanley B. Ringlod
1971-01-01
Thinning 37-year-old aspen in north central Minnesota did not increase either total volume production or the number of veneer-sized trees after 10 years. Thinning is not recommended for stands nearing rotation age.
Future timber harvests on the Chippewa and Superior National Forests in Minnesota.
Blair Orr; Joseph Buongiorno; Timothy Young; David C. Lothner; Edwin Kallio
1985-01-01
Timber harvests on the Superior and Chippewa National Forests in Minnesota are projected to increase substantially by 2030. The increasing demand for aspen may affect the total harvests. Both forests need to give high priority to aspen management.
Effects of widespread drought-induced aspen mortality on understory plants.
Anderegg, William R L; Anderegg, Leander D L; Sherman, Clare; Karp, Daniel S
2012-12-01
Forest die-off around the world is expected to increase in coming decades as temperature increases due to climate change. Forest die-off will likely affect understory plant communities, which have substantial influence on regional biological diversity, ecosystem function, and land-atmosphere interactions, but how die-off alters these plant communities is largely unknown. We examined changes in understory plant communities following a widespread, drought-induced die-off of trembling aspen (Populus tremuloides) in the western United States. We assessed shrub and herbaceous cover and volume in quadrats in 55 plots located across a wide range of levels of aspen mortality. We measured species richness and composition of herbaceous plant communities by recording species presence and absence in 12 sets of paired (1 healthy, 1 dying) aspen plots. Although understory composition in healthy and dying stands was heterogeneous across the landscape, shrub abundance, cover, and volume were higher and abundance of herbaceous species, cover, and volume were lower in dying aspen stands. Shrub cover and volume increased from 2009 to 2011 in dying stands, which suggests that shrub growth and expansion is ongoing. Species richness of herbs declined by 23% in dying stands. Composition of herbs differed significantly between dying and healthy stands. Richness of non-native species did not differ between stand types. The understory community in dying aspen stands was not similar to other shrub-dominated plant communities in the region and may constitute a novel community. Our results suggest that changes in understory plant communities as forests die off could be a significant indirect effect of climate change on biological diversity and forest communities. ©2012 Society for Conservation Biology.
Orbital Express mission operations planning and resource management using ASPEN
NASA Astrophysics Data System (ADS)
Chouinard, Caroline; Knight, Russell; Jones, Grailing; Tran, Daniel
2008-04-01
As satellite equipment and mission operations become more costly, the drive to keep working equipment running with less labor-power rises. Demonstrating the feasibility of autonomous satellite servicing was the main goal behind the Orbital Express (OE) mission. Like a tow-truck delivering gas to a car on the road, the "servicing" satellite of OE had to find the "client" from several kilometers away, connect directly to the client, and transfer fluid (or a battery) autonomously, while on earth-orbit. The mission met 100% of its success criteria, and proved that autonomous satellite servicing is now a reality for space operations. Planning the satellite mission operations for OE required the ability to create a plan which could be executed autonomously over variable conditions. As the constraints for execution could change weekly, daily, and even hourly, the tools used create the mission execution plans needed to be flexible and adaptable to many different kinds of changes. At the same time, the hard constraints of the plans needed to be maintained and satisfied. The Automated Scheduling and Planning Environment (ASPEN) tool, developed at the Jet Propulsion Laboratory, was used to create the schedule of events in each daily plan for the two satellites of the OE mission. This paper presents an introduction to the ASPEN tool, an overview of the constraints of the OE domain, the variable conditions that were presented within the mission, and the solution to operations that ASPEN provided. ASPEN has been used in several other domains, including research rovers, Deep Space Network scheduling research, and in flight operations for the NASA's Earth Observing One mission's EO1 satellite. Related work is discussed, as are the future of ASPEN and the future of autonomous satellite servicing.
Mauriat, Mélanie; Petterle, Anna; Bellini, Catherine; Moritz, Thomas
2014-05-01
Knowledge of processes involved in adventitious rooting is important to improve both fundamental understanding of plant physiology and the propagation of numerous plants. Hybrid aspen (Populus tremula × tremuloïdes) plants overexpressing a key gibberellin (GA) biosynthesis gene (AtGA20ox1) grow rapidly but have poor rooting efficiency, which restricts their clonal propagation. Therefore, we investigated the molecular basis of adventitious rooting in Populus and the model plant Arabidopsis. The production of adventitious roots (ARs) in tree cuttings is initiated from the basal stem region, and involves the interplay of several endogenous and exogenous factors. The roles of several hormones in this process have been characterized, but the effects of GAs have not been fully investigated. Here, we show that a GA treatment negatively affects the numbers of ARs produced by wild-type hybrid aspen cuttings. Furthermore, both hybrid aspen plants and intact Arabidopsis seedlings overexpressing AtGA20ox1, PttGID1.1 or PttGID1.3 genes (with a 35S promoter) produce few ARs, although ARs develop from the basal stem region of hybrid aspen and the hypocotyl of Arabidopsis. In Arabidopsis, auxin and strigolactones are known to affect AR formation. Our data show that the inhibitory effect of GA treatment on adventitious rooting is not mediated by perturbation of the auxin signalling pathway, or of the strigolactone biosynthetic and signalling pathways. Instead, GAs appear to act by perturbing polar auxin transport, in particular auxin efflux in hybrid aspen, and both efflux and influx in Arabidopsis. © 2014 The Authors The Plant Journal © 2014 John Wiley & Sons Ltd.
McGrath, Justin M; Karnosky, David F; Ainsworth, Elizabeth A
2010-04-01
Early spring leaf out is important to the success of deciduous trees competing for light and space in dense forest plantation canopies. In this study, we investigated spring leaf flush and how long-term growth at elevated carbon dioxide concentration ([CO(2)]) and elevated ozone concentration ([O(3)]) altered leaf area index development in a closed Populus tremuloides (aspen) canopy. This work was done at the Aspen FACE experiment where aspen clones have been grown since 1997 in conditions simulating the [CO(2)] and [O(3)] predicted for approximately 2050. The responses of two clones were compared during the first month of spring leaf out when CO(2) fumigation had begun, but O(3) fumigation had not. Trees in elevated [CO(2)] plots showed a stimulation of leaf area index (36%), while trees in elevated [O(3)] plots had lower leaf area index (-20%). While individual leaf area was not significantly affected by elevated [CO(2)], the photosynthetic operating efficiency of aspen leaves was significantly improved (51%). There were no significant differences in the way that the two aspen clones responded to elevated [CO(2)]; however, the two clones responded differently to long-term growth at elevated [O(3)]. The O(3)-sensitive clone, 42E, had reduced individual leaf area when grown at elevated [O(3)] (-32%), while the tolerant clone, 216, had larger mature leaf area at elevated [O(3)] (46%). These results indicate a clear difference between the two clones in their long-term response to elevated [O(3)], which could affect competition between the clones, and result in altered genotypic composition in future atmospheric conditions. Published by Elsevier Ltd.
The Fate of Aspen in a World with Diminishing Snowpacks
NASA Astrophysics Data System (ADS)
Kavanagh, K.; Link, T. E.; Seyfried, M. S.; Kemp, K. B.
2010-12-01
Aspen (Populus tremuloides) productivity is tightly coupled with soil moisture. In the mountainous regions of the western USA, annual replenishment of soil moisture commonly occurs during snowmelt. Therefore, snow pack depth and duration can play an important role in sustaining aspen productivity. The presence of almost 50 years of detailed climate data across an elevational transect in the Reynolds Creek Experimental Watershed (RCEW) in southwestern Idaho offers a novel opportunity to better understand the role of shifting precipitation patterns on aspen productivity. Over the past 50 years, the proportion of the precipitation falling in the form of snow decreased by almost a factor of 2 at mid to low elevations in the RCEW, coupled with a roughly four week advance of snow ablation, and decline of large snow drifts that release moisture into the early summer. Results from growth ring increment, stable isotope analysis, sapflux and a process model (Biome BGC), will be used to determine the impact of shifting precipitation patterns on tree productivity along this transect over the past 50 years. Aspen trees located on moist microsites continue to transpire water and maintain high stomatal conductance 21 days later in the growing season relative to individuals on drier microsites. Predictions of net primary productivity (NPP) in aspen are very sensitive to precipitation patterns. NPP becomes negative as early as day 183 (90 days post budbreak) for years with little winter and spring precipitation whereas, in years with ample winter and spring precipitation, NPP remains positive until day 260 when leaf fall occurs. These results give unique insight into the conditions that deciduous tree species will encounter in a warming climate where snow water equivalent continues to diminish and soil moisture declines soon after budbreak occurs.
Robinson, Kathryn M; Hauzy, Céline; Loeuille, Nicolas; Albrectsen, Benedicte R
2015-01-01
Nestedness and modularity are measures of ecological networks whose causative effects are little understood. We analyzed antagonistic plant–herbivore bipartite networks using common gardens in two contrasting environments comprised of aspen trees with differing evolutionary histories of defence against herbivores. These networks were tightly connected owing to a high level of specialization of arthropod herbivores that spend a large proportion of the life cycle on aspen. The gardens were separated by ten degrees of latitude with resultant differences in abiotic conditions. We evaluated network metrics and reported similar connectance between gardens but greater numbers of links per species in the northern common garden. Interaction matrices revealed clear nestedness, indicating subsetting of the bipartite interactions into specialist divisions, in both the environmental and evolutionary aspen groups, although nestedness values were only significant in the northern garden. Variation in plant vulnerability, measured as the frequency of herbivore specialization in the aspen population, was significantly partitioned by environment (common garden) but not by evolutionary origin of the aspens. Significant values of modularity were observed in all network matrices. Trait-matching indicated that growth traits, leaf morphology, and phenolic metabolites affected modular structure in both the garden and evolutionary groups, whereas extra-floral nectaries had little influence. Further examination of module configuration revealed that plant vulnerability explained considerable variance in web structure. The contrasting conditions between the two gardens resulted in bottom-up effects of the environment, which most strongly influenced the overall network architecture, however, the aspen groups with dissimilar evolutionary history also showed contrasting degrees of nestedness and modularity. Our research therefore shows that, while evolution does affect the structure of aspen–herbivore bipartite networks, the role of environmental variations is a dominant constraint. PMID:26306175
Forest statistics for Minnesota's Aspen-Birch Unit.
Neal P. Kingsley
1991-01-01
The fifth inventory of Minnesota's Aspen-Birch Unit reports 8.7 million acres of land, of which 7.4 million acres are forested. This bulletin present statistical highlights and contains detailed tables of forest area, as well as timber volume, growth, removals, mortality, and ownership.
Papermaking properties of aspen ultrahigh-yield mechanical pulps
J. N. McGovern; T. H. Wegner
1991-01-01
Eleven types of aspen ultra-high-yield (90% and above) mechanical pubs were evaluated for their chemical compositions (including sulfur), handsheet strength, and optical properties, fiber length indices, and fiberizing energies. The pulping processes were stone groundwood, pressurized stone groundwood, refiner mechanical, thermomechanical, chemimechanical (alkaline...
Wood property variation in Populus
Dean W. Einspahr; Miles K. Benson; John R. Peckham
1968-01-01
The use of bigtooth aspen (Populus grandidentata Michx.), quaking aspen (P. tremuloides Michx.), and cottonwood (P. deltoides Bartr.) by the pulp and paper industry has increased greatly during the past decade. This expanded use has stimulated research on the genetic improvement of Populus. For the past 12 years...
John R. Jones
1985-01-01
Quaking aspen is the most widely distributed native North American tree species (Little 1971, Sargent 1890). It grows in a great diversity of regions, environments, and communities (Harshberger 1911). Only one deciduous tree species in the world, the closely related Eurasian aspen (Populus tremula), has a wider range (Weigle and Frothingham 1911)....
Thomas E. Hinds
1985-01-01
Although many diseases attack aspen, relatively few kill or seriously injure living trees. The common leaf diseases, in general, are widely distributed throughout the range of aspen, whereas there are subtle differences in distribution between the important decay fungi, and apparently entirely different areas of distribution of major cankercausing organisms. However,...
Immobilized anaerobic fermentation for bio-fuel production by Clostridium co-culture.
Xu, Lei; Tschirner, Ulrike
2014-08-01
Clostridium thermocellum/Clostridium thermolacticum co-culture fermentation has been shown to be a promising way of producing ethanol from several carbohydrates. In this research, immobilization techniques using sodium alginate and alkali pretreatment were successfully applied on this co-culture to improve the bio-ethanol fermentation performance during consolidated bio-processing (CBP). The ethanol yield obtained increased by over 60 % (as a percentage of the theoretical maximum) as compared to free cell fermentation. For cellobiose under optimized conditions, the ethanol yields were approaching about 85 % of the theoretical efficiency. To examine the feasibility of this immobilization co-culture on lignocellulosic biomass conversion, untreated and pretreated aspen biomasses were also used for fermentation experiments. The immobilized co-culture shows clear benefits in bio-ethanol production in the CBP process using pretreated aspen. With a 3-h, 9 % NaOH pretreatment, the aspen powder fermentation yields approached 78 % of the maximum theoretical efficiency, which is almost twice the yield of the untreated aspen fermentation.
Automated Planning and Scheduling for Orbital Express (151)
NASA Technical Reports Server (NTRS)
Knight, Russell
2008-01-01
The challenging timeline for DARPA's Orbital Express mission demanded a flexible, responsive, and (above all) safe approach to mission planning. Because the mission was a technology demonstration, pertinent planning information was learned during actual mission execution. This information led to amendments to procedures, which led to changes in the mission plan. In general, we used the ASPEN planner scheduler to generate and validate the mission plans. We enhanced ASPEN to enable it to reason about uncertainty. We also developed a model generator that would read the text of a procedure and translate it into an ASPEN model. These technologies had a significant impact on the success of the Orbital Express mission.
BOREAS TE-12 SSA Leaf Water Potential Data
NASA Technical Reports Server (NTRS)
Hall, Forrest G. (Editor); Curd, Shelaine (Editor); Walter-Shea, Elizabeth A.; Mesarch, Mark A.; Chen, L.; Yang, Litao
2000-01-01
The Boreal Ecosystem-Atmospheric Study (BOREAS) TE-12 (Terrestrial Ecology) team collected water potential data in 1993 and 1994 from aspen, jack pine, and black spruce leaves/needles. Collections were made at the Southern Study Area Nipawin Fen Site (SSA FEN), Young Jack Pine (YJP), Young Aspen (YA), Old Aspen (OA), and Old Black Spruce (OBS) sites. Measurements were made using a pressure chamber on a platform in the field. The data are provided in tabular ASCII files. The data files are available on a CD-ROM (see document number 20010000884), or from the Oak Ridge National Laboratory (ORNL) Distributed Active Archive Center (DAAC).
Aspen: A microsimulation model of the economy
DOE Office of Scientific and Technical Information (OSTI.GOV)
Basu, N.; Pryor, R.J.; Quint, T.
1996-10-01
This report presents, Aspen. Sandia National Laboratories is developing this new agent-based microeconomic simulation model of the U.S. economy. The model is notable because it allows a large number of individual economic agents to be modeled at a high level of detail and with a great degree of freedom. Some features of Aspen are (a) a sophisticated message-passing system that allows individual pairs of agents to communicate, (b) the use of genetic algorithms to simulate the learning of certain agents, and (c) a detailed financial sector that includes a banking system and a bond market. Results from runs of themore » model are also presented.« less
Interfingering of the Frontier Formation and Aspen Shale, Cumberland Gap, Wyoming.
M'gonigle, J.
1982-01-01
The basal part, or the Chalk Creek Member, of the non-marine lower Frontier Formation (Upper Cretaceous) includes a thin coal bed that grades S into a carbonaceous shale. The latter plus associated sandstones and shales pinch out S of Cumberland Gap and lie stratigraphically below the top of the Aspen Shale. The beds in the upper part of the Aspen, in turn, pinch out within the Frontier Formation. The coal bed and equivalent carbonaceous shale represent in-place accumulation of peat. The interfingering suggests that in SW Wyoming the Lower/Upper Cretaceous boundary is within the Chalk Creek Member. -from Author
Aerogel: From Aerospace to Apparel
NASA Technical Reports Server (NTRS)
2001-01-01
Aspen Systems Inc. developed an aerogel-manufacturing process solved the handling problems associated with aerogel-based insulation products. Their aerogels can now be manufactured into blankets, thin sheets, beads, and molded parts; and may be transparent, translucent, or opaque. Aspen made the material effective for window and skylight insulation, non-flammable building insulation, and inexpensive firewall insulation that will withstand fires in homes and buildings, and also assist in the prevention of forest fires. Another Aspen product is Spaceloft(TM); an inexpensive, flexible blanket that incorporates a thin layer of aerogel embedded directly into the fabric. Spaceloft, is incorporated into jackets intended for wear in extremely harsh conditions and activities, such as Antarctic expeditions.
Aerogels Insulate Against Extreme Temperatures
NASA Technical Reports Server (NTRS)
2010-01-01
In 1992, NASA started to pursue the development of aerogel for cryogenic insulation. Kennedy Space Center awarded Small Business Innovation Research (SBIR) contracts to Aspen Systems Inc., of Marlborough, Massachusetts, that resulted in a new manufacturing process and a new flexible, durable, easy-to-use form of aerogel. Aspen Systems formed Aspen Aerogels Inc., in Northborough, Massachusetts, to market the product, and by 2009, the company had become the leading provider of aerogel in the United States, producing nearly 20 million square feet per year. With an array of commercial applications, the NASA-derived aerogel has most recently been applied to protect and insulate people s hands and feet.
Lake States Aspen Productivity Following Soil Compaction and Organic Matter Removal
Douglas M. Stone
2002-01-01
Aspen (Populus tremuloides Michx. and P. grandidentata Michx.) provides wood products, watershed protection, and wildlife habitat for numerous game and non-game species across the northern Great Lakes region. Sustaining the productivity of these ecosystems requires maintaining soil productivity. Management activities that decrease...
Field studies of pine, spruce and aspen periodically subjected to sulfur gas emissions
A. H. Legge; R. G. Amundson; D. R. Jaques; R. B. Walker
1976-01-01
Field studies of photosynthesis in Pinus contorta/Pinus banksiana (lodgepole pine/jack pine) hybrids, Picea glauca (white spruce) and Populus tremuloides (aspen) subjected to SO2 and H2S from a nearby natural gas processing plant were initiated near Whitecourt,...
Evaluation of a full-time ride service program : Aspen, Colorado's Tipsy Taxi service
DOT National Transportation Integrated Search
2000-10-01
This report summarizes a study of Aspen, Colorado's Tipsy Taxi ride service program. This service, which provides a free ride home for persons who are too intoxicated to drive, has been in place since December 1983. Funded through donations from the ...
The extent and characteristics of low-productivity aspen areas in Wisconsin.
Allen L. Lundgren; Jerold T. Hahn
1978-01-01
An analysis of inventory plots from Wisconsin's forest survey showed that 18% of the state's 3.7 million acres of aspen type was producing less than a quarter of potential volume yields and 47% was producing less than half of potential volume yields.
Projecting the aspen resource in the Lake States.
William A. Leuschner
1972-01-01
Aspen growing stock inventories for nine Lake States forest survey units were updated to the common base year of 1968. Cut and inventory were projected to the year 2000 under three sets of assumptions. Potential shortages were found in northeastern Wisconsin and Michigan if historical trends continue.
From Plant Hydraulics to Ecohydrology: a Case Study of Water Limitation in Aspen Forests
NASA Astrophysics Data System (ADS)
Sperry, J.; Venturas, M.; Love, D.; Anderegg, W.; Mackay, D. S.
2017-12-01
How dry must it get to threaten a standing forest? We answered this question for aspen stands in Utah with a model that predicts tree gas exchange and water status by optimizing photosynthetic gain vs. hydraulic risk from xylem cavitation. The model was parameterized for 10 aspen stands from various elevations and mountain ranges in the state of Utah, USA. The 2016 growing season was simulated from site-specific micrometeorological data under shallow (0.5 m) vs. deep (2m) root depth scenarios starting at field capacity. The model predicted a water-limiting threshold for each stand, defined as the minimum water input required to maximize stand gas exchange. All but one stand was estimated to be near or above its threshold in 2016. In the majority of stands, spring soil moisture and summer rain fell far short of supplying the threshold requirement. Without additional water, these stands would suffer over 70% loss of tree hydraulic conductance and high mortality risk. These more water-demanding stands were predicted to rely on groundwater for 60-95% of their threshold supply. Groundwater dependence suggests a greater sensitivity to winter precipitation than to growing season conditions. All but the sparsest aspen stands would experience significant mortality risk from a 50% reduction in groundwater input. The aspen test case suggests a wider utility for linking plant hydraulics and hydrology.
Duncan, Shona; Jing, Qing; Katona, Adrian; Kazlauskas, Romas J; Schilling, Jonathan; Tschirner, Ulrike; Aldajani, Waleed Wafa
2010-03-01
The recalcitrance of lignocellulosic biomass to enzymatic release of sugars (saccharification) currently limits its use as feedstock for biofuels. Enzymatic hydrolysis of untreated aspen wood releases only 21.8% of the available sugars due primarily to the lignin barrier. Nature uses oxidative enzymes to selectively degrade lignin in lignocellulosic biomass, but thus far, natural enzymes have been too slow for industrial use. In this study, oxidative pretreatment with commercial peracetic acid (470 mM) removed 40% of the lignin (from 19.9 to 12.0 wt.% lignin) from aspen and enhanced the sugar yields in subsequent enzymatic hydrolysis to about 90%. Increasing the amount of lignin removed correlated with increasing yields of sugar release. Unfortunately, peracetic acid is expensive, and concentrated forms can be hazardous. To reduce costs and hazards associated with using commercial peracetic acid, we used a hydrolase to catalyze the perhydrolysis of ethyl acetate generating 60-70 mM peracetic acid in situ as a pretreatment to remove lignin from aspen wood. A single pretreatment was insufficient, but multiple cycles (up to eight) removed up to 61.7% of the lignin enabling release of >90% of the sugars during saccharification. This value corresponds to a predicted 581 g of fermentable sugars from 1 kg of aspen wood. Improvements in the enzyme stability are needed before the enzymatically generated peracetic acid is a commercially viable alternative.
Acute O3 damage on first year coppice sprouts of aspen and maple sprouts in an open-air experiment
DOE Office of Scientific and Technical Information (OSTI.GOV)
Darbah, J.N.; Nagy, J.; Jones, W. S.
2011-10-01
We studied the effect of high ozone (O{sub 3}) concentration (110-490 nmol mol{sup -1}) on regenerating aspen (Populus tremuloides) and maple (Acer saccharum) trees at an open-air O{sub 3} pollution experiment near Rhinelander WI USA. This study is the first of its kind to examine the effects of acute O{sub 3} exposure on aspen and maple sprouts after the parent trees, which were grown under elevated O{sub 3} and/or CO{sub 2} for 12 years, were harvested. Acute O{sub 3} damage was not uniform within the crowns of aspen suckers; it was most severe in the mature, fully expanded photosynthesizing leaves.more » Young expanding leaves showed no visible signs of acute O{sub 3} damage contrary to expectations. Stomatal conductance played a primary role in the severity of acute O{sub 3} damage as it directly controlled O{sub 3} uptake. Maple sprouts, which had lower stomatal conductance, smaller stomatal aperture, higher stomatal density and larger leaf surface area, were tolerant of acute O{sub 3} exposure. Moreover, elevated CO{sub 2} did not ameliorate the adverse effects of acute O{sub 3} dose on aspen and maple sprouts, in contrast to its ability to counteract the effects of long-term chronic exposure to lower O{sub 3} levels.« less
Shrub Nesting of the Red-Eyed Vireo in Relation to Structure of Aspen Forests
William J. Jr Mattson
1979-01-01
Nests were built in five species of high shrubs in four trembling aspen forests of northern Minnesoa. Nest densities ranged between 0.5 and 3.0 per acre and were positively related to the abundance3 of large (>1.2-cm diameter) shrub stems.
Federal Register 2010, 2011, 2012, 2013, 2014
2010-03-23
... SECURITIES AND EXCHANGE COMMISSION [File No. 500-1] Aspen Group Resources Corp., Commercial Concepts, Inc., Desert Health Products, Inc., Equalnet Communications Corp., Geneva Steel Holdings Corp... securities of Commercial Concepts, Inc. because it has not filed any periodic reports since the period ended...
Regeneration and productivity of aspen grown on repeated short rotations.
Donald A. Perala
1979-01-01
Regeneration and productivity of aspen cannot be sustained on rotations shorter than about 8 years. Productivity losses on short rotations are physiological and morphological rather than nutritional in nature. Stump and root collar sprouts, which are rare in mature stands, were more numerous than root suckers.
Timber resource of Minnesota's Aspen-Birch Unit, 1977.
John S. Jr. Spencer; Arnold J. Ostrom
1979-01-01
The fourth inventory of Minnesota's Aspen-Birch Unit shows solid gains in growing-stock and sawtimber volumes between 1962 and 1977, but a 13% decline in commercial forest area. This report gives statistical highlights and contains detailed tables of forest area a well as timber volume, growth, mortality, ownership, and use.
Aspen community types of the Intermountain Region
Walter F. Mueggler
1988-01-01
This vegetation classification is based upon existing community structure and composition in the aspen-dominated forests of the Intermountain Region of the Forest Service. The 56 community types occur within eight tree-cover types. A diagnostic key using indicator species facilitates field identification of the community types. Vegetational composition, productivity,...
Risk Communication, Metacommunication, and Rhetorical Stases in the Aspen-EPA Superfund Controversy.
ERIC Educational Resources Information Center
Stratman, James F.; And Others
1995-01-01
Explores the relationship between current theoretical definitions of risk communication, the unique national role that EPA plays in defining health and environmental risks, and possible explanations for EPA's inability to persuade Aspen, Colorado, to accept a cleanup plan. Explores ownership messages conveyed through metacommunication conflict…
The 2013 Aspen Prize for Community College Excellence
ERIC Educational Resources Information Center
Perlstein, Linda
2013-01-01
For millions of Americans, community colleges provide an essential pathway to well-paying jobs and continuing higher education. The Aspen Prize for Community College Excellence honors those institutions that strive for and achieve exceptional levels of success for all students, while they are in college and after they graduate. Community colleges…
Photosynthetic productivity of aspen clones varying in sensitivity to tropospheric ozone
M.D. Coleman; J.G. Isebrands; R.E. Dickson; D.F. Karnosky
1995-01-01
Rooted cuttings from three aspen (Populus tremuloides Michx.) clones (216, 271 and 259, classified as high, intermediate and low in O3 tolerance, respectively) were exposed to either diumal O3 profiles simulating those of Michigan's Lower Peninsula (episodic treatments), or diurnal square-wave O
Aspen Root Sucker Formation and Apical Dominance
Robert E. Farmer
1962-01-01
Root suckering is the primary mode of regeneration in the aspens, Populus tremuloides Michx. and P. grandidentata Michx. When stems of these species are cut, numerous suckers originating in the root pericycle are formed on their extensive lateral root systems. During their first season of growth, suckers ordinarily reach a height...
Defense.gov Special Report: Aspen Security Forum 2014
Adventurism-Is Europe Ready? Cooperation in Counterterrorism Bringing into Balance the Military Instrument The Future of Warfare Striking Balance between Security and Liberty Assessing Department of Homeland Security Aspen Security Forum. Story Army Needs Balance, Odierno Says In the debate about how large the Army
Federal Register 2010, 2011, 2012, 2013, 2014
2012-10-03
... support the highest level of biodiversity for interior western forests. Productivity of aspen ecosystem... intolerant and sprouts heavily following disturbance; such as fire, which removes shading effects in areas..., mechanical treatments in Inventoried Roadless Areas, effects on wildlife species, effects to watersheds and...
Dennis P. Bradley; Frank E. Biltonen
1973-01-01
Describes the productivity of selected aspen pulpwood loggers in northern Minnesota. The most important factors affecting productivity were the ratio of harvested trees per acre to total trees per acre, harvested volume per acre, and the spacing of nonharvested trees
Modeling aspen and red pine shoot growth to daily weather variations.
Donald A. Perala
1983-01-01
Quantifies daily shoot growth of quaking aspen and red pine in response to daily variation in air temperature, soil moisture, solar radiation, evapotranspiration, and inherent seasonal plant growth rhythm. Discusses potential application of shoot growth equations to silvicultural problems related to microclimatic variation. Identifies limitations and areas for...
ASPEN Plus in the Chemical Engineering Curriculum: Suitable Course Content and Teaching Methodology
ERIC Educational Resources Information Center
Rockstraw, David A.
2005-01-01
An established methodology involving the sequential presentation of five skills on ASPEN Plus to undergraduate seniors majoring in ChE is presented in this document: (1) specifying unit operations; (2) manipulating physical properties; (3) accessing variables; (4) specifying nonstandard components; and (5) applying advanced features. This…
AmeriFlux CA-Oas Saskatchewan - Western Boreal, Mature Aspen
Black, T. Andrew [The University of British Columbia
2017-01-01
This is the AmeriFlux version of the carbon flux data for the site CA-Oas Saskatchewan - Western Boreal, Mature Aspen. Site Description - 53.62889° N, 106.19779° W, elabation of 600.63 m,BOREAS 1994, 1996, BERMS climate and flux measurements began Dec. 1996
Steroid Hormones in NF1 Tumorigenesis
2005-08-01
Schwannomatosis , in Aspen in June this year. Table 3. Summary of current xenograft proliferation/TUNEL (apoptosis) data. Culture I sex I Outlier? I PI/Al I...Biology of NF1, NF2, and schwannomatosis . Aspen, CO, June 2005, poster. CONCLUSIONS: In vitro data indicate that steroid hormone receptor levels in normal
BOREAS TE-12 SSA Shoot Geometry Data
NASA Technical Reports Server (NTRS)
Hall, Forrest G. (Editor); Curd, Shelaine (Editor); Walter-Shea, Elizabeth A.; Mesarch, Mark A.; Cheng, L.; Yang, Litao
2000-01-01
The Boreal Ecosystem-Atmospheric Study (BOREAS) TE-12 (Terrestrial Ecology) team collected shoot geometry data in 1993 and 1994 from aspen, jack pine, and black spruce trees. Collections were made at the Southern Study Area Nipawin Fen Site (SSA FEN), Young Jack Pine (YJP), Old Jack Pine (OJP), Old Aspen (OA), Young Aspen (YA), Mixed Site (MIX), and Old Black Spruce (OBS) sites. A caliper was used to measure shoot and needle lengths and widths. A volume displacement procedure was used to measure the weight of the shoot or twig submerged in water. The data files are available on a CD-ROM (see document number 20010000884), or from the Oak Ridge National Laboratory (ORNL) Distributed Active Archive Center (DAAC).
Identifying environmental features for land management decisions. [Utah
NASA Technical Reports Server (NTRS)
1981-01-01
Advances in digital processing of LANDSAT imagery and in the interpretation of aerial photography are reported as well as the development of a geographic information system and the acquisition of cartographic equipment. Services to technical specialists and dignitaries are also described. The status of the following studies is discussed: Farmington Bay waterfowl; Sevier River wetland and agriculture; Davis County foothill development; Bear River Range aspen habitat; Wasatch-Cache riparian habitats; irrigated acreage in the Bear River Basin; the Parker Mountain rangeland inventory; and the development of techniques for inventorying aspen and aspen conifer stands. The cooperative project with NASA-Ames to verify LANDSAT digital maps of the forest and range resources on the LaSal Mountains is also considered.
Forage quality in burned and unburned aspen communities
Norbert V. DeByle; Philip J. Urness; Deborah L. Blank
1989-01-01
Selected forage species were sampled during the first and second summers after autumn prescribed burning of three sites in southeastern Idaho. They were analyzed for in vitro dry matter digestibility, protein, calcium, and phosphorus. This aspen type has a highly nutritious understory. Burning further improved the quality of the selected species only during the first...
Aspen flakeboard treated with disodium octaborate tetrahydrate
Robert H. White; John Forsman; John R. Erickson
2008-01-01
In this project, we investigated mechanical properties and fire performance of aspen flakeboards manufactured with the fire-retardant chemical disodium octaborate tetrahydrate (DOT). Flakeboards were prepared using two levels of adhesive loading (5% and 7% methylene diphenyl diisocyanate (MDI)) and three levels of fire-retardant treatments (6%, 9%, and 12%). DOT is a...
Norbert V. DeByle
1985-01-01
Quaking aspen dominates several million acres on mountainous watersheds in the West. The sites occupied receive enough precipitation to yield water to lower elevations. Most aspen areas receive 16 inches (40 cm) or more precipitation annually; many receive more than 39 inches (100 cm) (see the CLIMATES chapter), well in excess of on-site loss from evapotranspiration....
Federal Register 2010, 2011, 2012, 2013, 2014
2013-07-31
... providing for human safety. Treatments would be carried out on National Forest System (NFS) Lands within the scope of direction provided in the GMUG Revised Land and Resource Management Plan. DATES: To be most... DEPARTMENT OF AGRICULTURE Forest Service Spruce Beetle Epidemic and Aspen Decline Management...
Federal Register 2010, 2011, 2012, 2013, 2014
2012-05-25
...[bdi1][bdi0] and Fort Collins Carbon Monoxide Maintenance Plans' Motor Vehicle Emissions Budgets for... Aspen Attainment/Maintenance Area'' \\1\\ and its motor vehicle emissions budget, and the ``Revised Carbon... which we determine whether a SIP revision's motor vehicle emissions budget (MVEB) is adequate for...
John R. Jones; George A. Schier
1985-01-01
This chapter considers aspen growth as a process, and discusses some characteristics of the growth and development of trees and stands. For the most part, factors affecting growth are discussed elsewhere, particularly in the GENETICS AND VARIATION chapter and in chapters in PART 11. ECOLOGY. Aspen growth as it relates to wood production is examined in the WOOD RESOURCE...
USDA-ARS?s Scientific Manuscript database
ASPEN Plus based simulation models have been developed to design a pyrolysis process for the on-site production and utilization of pyrolysis oil from equine waste at the Equine Rehabilitation Center at Morrisville State College (MSC). The results indicate that utilization of all available Equine Reh...
W. F. Mueggler
1985-01-01
The extensive forests and isolated clones of quaking aspen in the western United States have been valued for many years as wildlife habitat and livestock summer range (Sampson 1919). The actual amount of forage produced beneath the aspen trees differs appreciably among sites. Houston (1954) indicated that although many sites produce 1,000 to 2,000 pounds per acre (1,...
Colin M. Callahan; Carol A. Rowe; Ronald J. Ryel; John D. Shaw; Michael D. Madritch; Karen E. Mock
2013-01-01
Aspen populations in the south-western portion of the range are consistent with expectations for a historically stable edge, with low within-population diversity, significant geographical population structuring, and little evidence of northward expansion. Structuring within the southwestern cluster may result from distinct gene pools separated during the Pleistocene...
Hereford Garland
1949-01-01
The use of aspen for the manufacture of excelsior is well established in the Lake States. The properties of the wood make it suitable for this general use, and there are adequate supplies available for increased productive capacity. The process of manufacturing exceisior is relatively uncomplicated, and the production equipment is simple and relatively low in cost....
Nutrient transport in surface runoff and interflow from an aspen-birch forest
D.R. Timmons; E.S. Verry; R.E. Burwell; R.F. Holt
1977-01-01
Nutrients transported in surface runoff and interflow from an undisturbed aspen-birch (Populus tremuloides Michx., and Betula papyrifera Marsh.) forest (6.48 ha) in northern Minnesota were measured for 3 years. Surface runoff from snowmelt accounted for 97% of the average annual surface runoff and for 57% of the average annual...
Estimating white trunk rot in aspen stands
Alan C. Jones; Michael E. Ostry
1998-01-01
Advanced decay caused by Phellinus tremulae was estimated in 295 trembling aspen on 30 plots in 2 Minnesota counties using existing inventory guides, and then measured by felling and sectioning the trees. In standing trees, decay volume was underestimated by 38% compared to measured decay volume in felled trees. The most reliable external indicator...
John R. Jones; Norbert V. DeByle
1985-01-01
The broad range of aspen in North America is evidence of its equally broad tolerance of wide variations in climate (Fowells 1965). Given open space for establishment and not too severe competition from other plants, aspen can survive from timberline on the tundra's edge to very warm temperate climates, and from the wet maritime climates of the coasts to very...
Timothy D. Meehan; Michael S. Crossley; Richard L. Lindroth
2010-01-01
Human alteration of atmospheric composition affects foliar chemistry and has possible implications for the structure and functioning of detrital communities. In this study, we explored the impacts of elevated carbon dioxide and ozone on aspen (Populus tremuloides) leaf litter chemistry, earthworm (Lumbricus terrestris) individual...
Aspen ecosystem properties in the Upper Great Lakes.
David H. Alban; Donald A. Perala; Martin F. Jurgensen; Michael E. Ostry; John R. Probst
1991-01-01
Describes four ecosystems in Minnesota and Michigan comprised of mature aspen stands on a range of soils from sandy to clay. The ecosystems are part of a long-term study of the effects of harvesting and species conversion on ecosystem properties. Presents data on geology, weather, soils, vegetation, litterfall, nitrogen dynamics, insects, disease, and wildlife.
Historical patterns in lichen communities of montane quaking aspen forests
Paul C. Rogers; Dale L. Bartos; Ronald J. Ryel
2011-01-01
Climate shifts and resource exploitation in Rocky Mountain forests have caused profound changes in quaking aspen (Populus tremuloides Michx.) structure and function since Euro-American settlement. It therefore seems likely that commensurate shifts in dependent epiphytes would follow major ecological transitions. In the current study, we merge several lines of inquiry...
Defensive effects of extrafloral nectaries in quaking aspen differ with scale
Brent Mortensen; Diane Wagner; Patricia Doak
2010-01-01
The effects of plant defenses on herbivory can differ among spatial scales. This may be particularly common with indirect defenses, such as extrafloral nectarines (EFNs), that attract predatory arthropods and are dependent on predator distribution, abundance, and behavior. We tested the defensive effects of EFNs in quaking aspen (Populus tremuloides...
Quaking aspen and the human experience: Dimensions, issues, and challenges
Stephen F. McCool
2001-01-01
Humans assign four types of meanings to aspen landscapes: (1) instrumental meanings dealing with the attainment of a goal - such as production of pulp or provision of recreation opportunities; (2) aesthetic meanings; (3) cultural/symbolic meanings dealing with spiritual and social attachments to landscapes; and (4) individual/expressive meanings derived out of...
Moisture sorption properties of composite boards from esterified aspen fiber
C. Clemons; R. A. Young; R. M. Rowell
1992-01-01
One barrier to producing wood-plastic composites with wood fiber is the poor thermoplasticity of wood fiber. The objective of our study was to determine the plasticization of chemically modified wood fiber through tests on unmodified and esterified fiberboards. Attrition-milled aspen fiber was esterified with neat acetic, maleic, or succinic anhydride. Fourier...
Development of a protocol for the ecological assessment of a special species
David Burton
2004-01-01
Developing consistent inventory and assessment protocols is important to people working on aspen issues in California and Nevada. Efforts have focused on identifying key indicators of ecological condition within aspen stands. The protocols have incorporated a range of factors that create or affect those indicators. Resulting ecological assessments conducted through the...
ERIC Educational Resources Information Center
Evans, Steven T.; Huang, Xinqun; Cramer, Steven M.
2010-01-01
The commercial simulator Aspen Chromatography was employed to study and optimize an important new industrial separation process, weak partitioning chromatography. This case study on antibody purification was implemented in a chromatographic separations course. Parametric simulations were performed to investigate the effect of operating parameters…
Norbert V. DeByle
1985-01-01
The aspen ecosystem is rich in number and species of animals, especially in comparison to associated coniferous forest types. This natural species diversity and richness has been both increased and influenced by the introduction of domestic livestock. The high value of the aspen type as a forage resource for livestock and as forage and cover for wildlife makes the...
Decay and Discoloration of Aspen
Gerald W. Anderson; Thomas E. Hinds; Donald M. Knutson
1977-01-01
Quaking aspen (Populus tremuloides Michx.) is the most widely distributed tree species in North America. It has a transcontinental range and grows under a wide variety of climatic and edaphic conditions. The commercial range includes the Lake States as well as parts of New England; it also includes Manitoba, Saskatchewan, and Alberta, Canada. Although many stands in...
Stomatal uptake of O3 in aspen and aspen-birch forests under free-air CO2 and O3 enrichment
Johan Uddling; Alan J. Hogg; Ronald M. Teclaw; Mary Anne Carroll; David S. Ellsworth
2010-01-01
Rising atmospheric carbon dioxide (CO2) may alleviate the toxicological impacts of concurrently rising tropospheric ozone (O3) during the present century if higher CO2 is accompanied by lower stomatal conductance (gs), as assumed by many models. We investigated how elevated...
Whole stand volume tables for quaking aspen in the Rocky Mountains
Wayne D. Shepperd; H. Todd Mowrer
1984-01-01
Linear regression equations were developed to predict stand volumes for aspen given average stand basal area and average stand height. Tables constructed from these equations allow easy field estimation of gross merchantable cubic and board foot Scribner Rules per acre, and cubic meters per hectare using simple prism cruise data.
Restoring High Priority Habitats for Birds: Aspen and Pine in the Interior West
Rex Sallabanks; Nils D. Christoffersen; Whitney W. Weatherford; Ralph Anderson
2005-01-01
This paper describes a long-term habitat restoration project in the Blue Mountains ecoregion, northeast Oregon, that we initiated in May 2000. We focused our restoration activities on two habitats previously identified as being high priority for birds: quaking aspen (Populus tremuloides) and ponderosa pine (Pinus ponderosa). In...
Carbon allocation and partitioning in aspen clones varying in sensitivity to tropospheric ozone
M.D. Coleman; R.E. Dickson; J.G. Isebrands; D.F. Karnosky
1995-01-01
Clones of aspen (Populus tremuloides Michx.) were identified that differ in biomass production in response to O3exposure. 14Carbon tracer studies were used to determine if the differences in biomass response were linked to shifts in carbon allocation and carbon partitioning patterns. Rooted cuttings from...
Estimating aspen crown fuels in northeastern Minnesota.
Robert M. Loomis; Peter J. Roussopoulos
1978-01-01
An application section presents tables for estimating foliage and branchwood (wood plus bark) of aspen tree crowns; the branchwood is subdivided by (1) living and dead material and by (2) living material alone into diameter size groups. A documentation section describes the derivation of equations and compares the equations derived with those derived by other...
Aspen sprout production and water use
Robert S. Johnston
1969-01-01
Sprouting response and soil moisture depletion on aspen plots were compared under four experimental conditions: (a) clearcut, (b) clearcut, stumps sprayed with sodium arsenite, (c) basal injection of sodium arsenite, and (d) control. Nunbers of sprouts varied with treatment for 2 years, but after 4 years the nunbers of sprouts on all plots were about equal....
John R. Jones; Norbert V. DeByle
1985-01-01
Edaphic and climatic characteristics of a site quite well define the quality of that site for plant growth. The importance of soil characteristics to the growth and well-being of aspen in the West is apparent from observations by many authors, from inferences resulting from work with other trees and agricultural crops, and from detailed study of aspen soils and site...
Possibilities of managing aspen
Paul Zehngraff
1947-01-01
The management of aspen or popple (Populus tremuloides) has received little attention in the past, largely because of the former abundant supply of other and more valuable tree species in the Lake States. Because of a rapidly expanding market for all forest products during the past decade, particularly during the war, the attention and ingenuity of...
Soil properties and aspen development five years after compaction and forest floor removal
Douglas M. Stone; John D. Elioff
1998-01-01
Forest management activities that decrease soil porosity and remove organic matter have been associated with declines in site productivity. In the northern Lake States region, research is in progress in the aspen (Populus tremuloides Michx. and P. grandidentata Michx.) forest type to determine effects of soil compaction and organic...
Modeling the Removal of Xenon from Lithium Hydrate with Aspen HYSYS
NASA Astrophysics Data System (ADS)
Efthimion, Phillip; Gentile, Charles
2011-10-01
The Laser Inertial Fusion Engine (LIFE) project mission is to provide a long-term, carbon-free source of sustainable energy, in the form of electricity. A conceptual xenon removal system has been modeled with the aid of Aspen HYSYS, a chemical process simulator. Aspen HYSYS provides excellent capability to model chemical flow processes, which generates outputs which includes specific variables such as temperature, pressure, and molar flow. The system is designed to strip out hydrogen isotopes deuterium and tritium. The base design bubbles plasma exhaust laden with x filled with liquid helium. The system separates the xenon from the hydrogen, deuterium, and tritium with a lithium hydrate and a lithium bubbler. After the removal of the hydrogen and its isotopes, the xenon is then purified by way of the process of cryogenic distillation. The pure hydrogen, deuterium, and tritium are then sent to the isotope separation system (ISS). The removal of xenon is an integral part of the laser inertial fusion engine and Aspen HYSYS is an excellent tool to calculate how to create pure xenon.
Growth-climate relationships across topographic gradients in the northern Great Lakes
Dymond, S.F.; D'Amato, A.W.; Kolka, R.K.; Bolstad, P.V.; Sebestyen, S.D.; Bradford, John B.
2016-01-01
Climatic conditions exert important control over the growth, productivity, and distribution of forests, and characterizing these relationships is essential for understanding how forest ecosystems will respond to climate change. We used dendrochronological methods to develop climate–growth relationships for two dominant species, Populus tremuloides (quaking aspen) and Pinus resinosa (red pine), in the upper Great Lakes region to understand how climate and water availability influence annual forest productivity. Trees were sampled along a topographic gradient at the Marcell Experimental Forest (Minnesota, USA) to assess growth response to variations in temperature and different water availability metrics (precipitation, potential evapotranspiration (PET), cumulative moisture index (CMI), and soil water storage). Climatic variables were able to explain 33–58% of the variation in annual growth (as measured by ring-width increment) for quaking aspen and 37–74% of the variation for red pine. Climate–growth relationships were influenced by topography for quaking aspen but not for red pine. Annual ring growth for quaking aspen decreased with June CMI on ridges, decreased with temperature in the November prior to the growing season on sideslopes, and decreased with June PET on toeslopes. Red pine growth increased with increasing July PET across all topographic positions. These results indicate the sensitivity of both quaking aspen and red pine to local climate and show several vulnerabilities of these species to shifts in water supply and temperature because of climate change.
Simulating the Dependence of Aspen on Redistributed Snow
NASA Astrophysics Data System (ADS)
Soderquist, B.; Kavanagh, K.; Link, T. E.; Seyfried, M. S.; Winstral, A. H.
2013-12-01
In mountainous regions across the western USA, the distribution of aspen (Populus tremuloides) is often directly related to heterogeneous soil moisture subsidies resulting from redistributed snow. With decades of climate and precipitation data across elevational and precipitation gradients, the Reynolds Creek Experimental Watershed (RCEW) in southwest Idaho provides a unique opportunity to study the relationship between aspen and redistributed snow. Within the RCEW, the total amount of precipitation has not changed in the past 50 years, but there are sharp declines in the percentage of the precipitation falling as snow. As shifts in the distribution of available moisture continue, future trends in aspen net primary productivity (NPP) remain uncertain. In order to assess the importance of snowdrift subsidies, NPP of three aspen stands was simulated at sites spanning elevational and precipitation gradients using the biogeochemical process model BIOME-BGC. At the aspen site experiencing the driest climate and lowest amount of precipitation from snow, approximately 400 mm of total precipitation was measured from November to March of 2008. However, peak measured snow water equivalent (SWE) held in drifts directly upslope of this stand was approximately 2100 mm, 5 times more moisture than the uniform winter precipitation layer initially assumed by BIOME-BGC. BIOME-BGC simulations in dry years forced by adjusted precipitation data resulted in NPP values approximately 30% higher than simulations assuming a uniform precipitation layer. Using BIOME-BGC and climate data from 1985-2011, the relationship between simulated NPP and measured basal area increments (BAI) improved after accounting for redistributed snow, indicating increased simulation representation. In addition to improved simulation capabilities, soil moisture data, diurnal branch water potential, and stomatal conductance observations at each site detail the use of soil moisture in the rooting zone and the onset of drought stress occurring in stands located along a precipitation phase gradient. These results further emphasize the importance of redistributed snow in heterogeneous landscapes along with the need to account for temporal shifts in water resource availability when assessing ecosystem vulnerability to climate change.
NASA Astrophysics Data System (ADS)
Landhäusser, S. M.; Snedden, J.; Silins, U.; Devito, K. J.
2012-04-01
Spatial root distribution, root morphology, and intra- and inter-clonal connections of mature boreal trembling aspen clones (Populus tremuloides Michx.) were explored to shed light on the functional relationships between vertical and horizontal distribution of roots and the variation in soil water availability along hill slopes. Root systems of mature aspen were hydraulically excavated in large plots (6 m wide and 12 m long) and to a depth of 30 cm. Most aspen roots were located in the upper 20 cm of the soil and fine and coarse root occupancy was highest in the lower slope positions and lowest towards the upper hill slope position likely because of soil moisture availability. Observation of the root system distribution along the hill slope correlated well with the observation of greater leaf area carried by trees growing at the lower portion of the hill slope. Interestingly, trees growing at the bottom of the slope required also less sapwood area to support the same amount of leaf area of trees growing at the top of a slope. These observations appear to be closely related to soil moisture availability and with that greater productivity at the bottom of the slope. However, trees growing on the upper slope tended to have long lateral roots extending downslope, which suggests long distance water transport through these lateral feeder roots. Genetic analysis indicated that both intra- and inter-clonal root connections occur in aspen, which can play a role in the sharing of resources along moisture gradients. Root systems of boreal aspen growing on upper slope positions exhibited a combination of three attributes (1) asymmetric lateral root systems, that are skewed downslope, (2) deeper taproots, and (3) intra and inter-clonal root connections, which can all be considered adaptive strategies to avoid drought stress in upper slope positions.
Ayres, Edward; Steltzer, Heidi; Berg, Sarah; Wallenstein, Matthew D; Simmons, Breana L; Wall, Diana H
2009-06-18
Previous studies have shown that plants often have species-specific effects on soil properties. In high elevation forests in the Southern Rocky Mountains, North America, areas that are dominated by a single tree species are often adjacent to areas dominated by another tree species. Here, we assessed soil properties beneath adjacent stands of trembling aspen, lodgepole pine, and Engelmann spruce, which are dominant tree species in this region and are distributed widely in North America. We hypothesized that soil properties would differ among stands dominated by different tree species and expected that aspen stands would have higher soil temperatures due to their open structure, which, combined with higher quality litter, would result in increased soil respiration rates, nitrogen availability, and microbial biomass, and differences in soil faunal community composition. We assessed soil physical, chemical, and biological properties at four sites where stands of aspen, pine, and spruce occurred in close proximity to one-another in the San Juan Mountains, Colorado. Leaf litter quality differed among the tree species, with the highest nitrogen (N) concentration and lowest lignin:N in aspen litter. Nitrogen concentration was similar in pine and spruce litter, but lignin:N was highest in pine litter. Soil temperature and moisture were highest in aspen stands, which, in combination with higher litter quality, probably contributed to faster soil respiration rates from stands of aspen. Soil carbon and N content, ammonium concentration, and microbial biomass did not differ among tree species, but nitrate concentration was highest in aspen soil and lowest in spruce soil. In addition, soil fungal, bacterial, and nematode community composition and rotifer, collembolan, and mesostigmatid mite abundance differed among the tree species, while the total abundance of nematodes, tardigrades, oribatid mites, and prostigmatid mites did not. Although some soil characteristics were unaffected by tree species identity, our results clearly demonstrate that these dominant tree species are associated with soils that differ in several physical, chemical, and biotic properties. Ongoing environmental changes in this region, e.g. changes in fire regime, frequency of insect outbreaks, changes in precipitation patterns and snowpack, and land-use change, may alter the relative abundance of these tree species over coming decades, which in turn will likely alter the soils.
Variable performance of outbreak defoliators on aspen clones exposed to elevated CO2 and O3
Daniel A. Herms; William J. Mattson; David N. Karowe; Mark D. Coleman; Terry M. Trier; Bruce A. Birr; J. G. Isebrands
1996-01-01
Increasing atmospheric concentrations of ozone and CO2 affect many aspects of tree physiology. However, their effects on tree resistance to insects have received relatively little attention. The objectives of this study were to test the effects of elevated CO2 and ozone on the resistance of three quaking aspen (...
Ruffed grouse selection of drumming sites in the Black Hills National Forest
Christopher P. Hansen; Mark A. Rumble; Joshua J. Millspaugh
2011-01-01
Ruffed grouse (Bonasa umbellus) are important game birds that depend on multiple forest age-classes of aspen (Populus spp.) for food and cover, which makes them an appropriate management indicator species for the condition of quaking aspen (Populus tremuloides) communities in the Black Hills National Forest of western South Dakota and northeastern Wyoming (BHNF)....
Thomas L. Eberhardt; Soo-Hong Min; James S. Han
2006-01-01
Biomass-based filtration media are of interest as an economical means to remove pollutants and nutrients found in stormwater runoff. Refined aspen wood fiber samples treated with iron salt solutions demonstrated limited capacities to remove (ortho)phosphate from test solutions. To provide additional sites for iron complex formation, and thereby impart a greater...
Populus tremuloides mortality near the southwestern edge of its range
Thomas J. Zegler; Margaret M. Moore; Mary L. Fairweather; Kathryn B. Ireland; Peter Z. Fule
2012-01-01
Mortality and crown dieback of quaking aspen (Populus tremuloides) were extensive on the Williams Ranger District, Kaibab National Forest in northern Arizona. We collected data from a random sample of 48 aspen sites to determine the relationship of predisposing site and stand factors and contributing agents to ramet mortality. Mortality of overstory (P10.1 cm DBH)...
Elevated CO2 response of photosynthesis depends on ozone concentration in aspen
A. Noormets; O. Kull; A. Sôber; M.E. Kubiske; D.F. Karnosky
2010-01-01
The effect of elevated CO2 and O3 on apparent quantum yield (f), maximum photosynthesis (Pmax), carboxylation efficiency (Vcmax) and electron transport capacity (Jmax) at different canopy locations was studied in two aspen (Populus tremuloides) clones of contrasting O3 tolerance. Local light climate at every leaf was characterized as fraction of above-canopy...
Is the wide distribution of aspen a result of its stress tolerance?
V. J. Lieffers; S. M. Landhausser; E. H. Hogg
2001-01-01
Populus tremuloides is distributed from drought-prone fringes of the Great Plains to extremely cold sites at arctic treeline. To occupy these conditions aspen appears to be more tolerant of stress than the other North American species of the genus Populus. Cold winters, cold soil conditions during the growing season, periodic drought, insect defoliation, and...
Survivor aspen: Can we predict who will get voted off the island?
F. A. Baker; J. D. Shaw
2008-01-01
During the past few years, aspen have been dying at rates that appear to exceed normal rates. We believe that this mortality should not be unexpected, given the severe drought of the past 10 years. We examine the literature and FIA data and identify several factors that indicate such mortality should be expected.
Jeffrey T. Olson; Allen L. Lundgren
1978-01-01
The 1968 Wisconsin Forest Survey showed large areas of aspen type that are not considered fully stocked. The economic feasibility of converting partially-stocked stands to full stocking is examined, and a rule presented for determining when a partially-stocked stand should be harvested to maximize its present value.
Widespread triploidy in western North American aspen (Populus tremuloides)
Karen E. Mock; Colin M. Callahan; M. Nurul Islam-Faridi; John D. Shaw; Hardeep S. Rai; Stewart C. Sanderson; Carol A. Rowe; Ronald J. Ryel; Michael D. Madritch; Richard S. Gardner; Paul G. Wolf
2012-01-01
We document high rates of triploidy in aspen (Populus tremuloides) across the western USA (up to 69% of genets), and ask whether the incidence of triploidy across the species range corresponds with latitude, glacial history (as has been documented in other species), climate, or regional variance in clone size. Using a combination of microsatellite genotyping, flow...
Advancing Entrepreneurship Education. A Report of the Youth Entrepreneurship Strategy Group
ERIC Educational Resources Information Center
Aspen Institute, 2008
2008-01-01
The Youth Entrepreneurship Strategy Group convened its inaugural meeting from September 26-28, 2007 at the Aspen Institute in Aspen, Colorado. A group of dynamic national leaders from the fields of education, entrepreneurship and business, public policy, media, and philanthropy met over three days to explore the promise of, and obstacles to,…
John R. Jones; Norbert V. DeByle
1985-01-01
In some areas, many aspen stands are all the same age, dating from a single great fire or a year of widespread fires (fig. 1). The 1879 fire in the Jackson Hole region of Wyoming (Loope and Gruell 1973) and the 1904 fires in Arizona's White Mountains (Kallander 1969) are examples. Choate (1966) found that almost all aspen stands in New Mexico were even-aged, many...
Jessica E. Hancock; Wendy M. Loya; Christian P. Giardina; Laigeng Li; Vincent L. Chiang; Kurt S. Pregitzer
2007-01-01
We conducted a glasshouse mesocosm study that combined 13C isotope techniques with wild-type and transgenic aspen (Populus tremuloides) in order to examine how altered lignin biosynthesis affects plant production and soil carbon formation. Our transgenic aspen lines expressed low stem lignin concentration but normal cellulose...
Aspen succession in the Intermountain West: A deterministic model
Dale L. Bartos; Frederick R. Ward; George S. Innis
1983-01-01
A deterministic model of succession in aspen forests was developed using existing data and intuition. The degree of uncertainty, which was determined by allowing the parameter values to vary at random within limits, was larger than desired. This report presents results of an analysis of model sensitivity to changes in parameter values. These results have indicated...
Relationship between foliar chemistry and insect performance: the forest tent caterpillar
Francois Lorenzetti; Yves Mauffette; Eric Bauce
1999-01-01
Forest tent caterpillar (FTC) feeds on several species of deciduous trees (Stehr and Cook 1968), in northeastern North America, quaking aspen is the preferred host of this spring-feeding insect. FTC commonly defoliates several thousands of hectares of aspen stands each year in Quebec (Bordeleau 1990), although its secondary hosts seldom are attacked.
L.F Ohmann; H.O. Batzer; R.R. Buech; D.C. Lothner; D. A. Perala; A.L. Schipper; E.S. Verry
1978-01-01
Describes some harvest options and their consequences in terms of timber investment return, water yield and quality, wildlife, visual quality, and disease and insect impact for the aspen, white birch, red pine, white pine, jack pine, black spruce, spruce-fir, and white-cedar forest types of the Lake States.
Acute O3 damage on first year coppice sprouts of aspen and maple sprouts in an open-air experiment
Joseph N.T. Darbah; Wendy S. Jones; Andrew J. Burton; John Nagy; Mark E. Kubiske
2011-01-01
We studied the effect of high ozone (O3) concentration (110-490 nmol mol-1) on regenerating aspen (Populus tremuloides) and maple (Acer saccharum) trees at an open-air O3 pollution experiment near Rhinelander WI USA. This study is the first of its kind to examine...
Foreword: "We already know all about aspen"
Dan Binkley
2001-01-01
As we developed plans for the symposium on sustaining aspen in western landscapes (held in Grand Junction, Colorado, on June 13-15, 2000), we solicited support from state and federal agencies, universities, and the Rocky Mountain Elk Foundation. The widespread support was very encouraging, and only one agency turned down the opportunity to join in supporting this...
Defining Excellence: Lessons from the 2013 Aspen Prize Finalists
ERIC Educational Resources Information Center
Aspen Institute, 2013
2013-01-01
In many respects, one couldn't find a group of 10 schools more diverse than the finalists for the 2013 Aspen Prize for Community College Excellence. One community college serves 1,500 students, another 56,000. There are institutions devoted primarily--even solely--to technical degrees, and ones devoted mainly to preparing students for further…
Katri Kostiainen; Seija Kaakinen; Elina Warsta; Mark E. Kubiske; Neil D. Nelson; Jaak Sober; David F. Karnosky; Pekka Saranpaa; Elina Vapaavuori
2008-01-01
We investigated the interactive effects of elevated concentrations of carbon dioxide ([CO2]) and ozone ([O3]) on radial growth, wood chemistry and structure of five 5-year-old trembling aspen (Populus tremuloides Michx.) clones and the wood chemistry of paper birch (Betula papyrifera...
Johan Uddling; Ronald M. Teclaw; Mark E. Kubiske; Kurt S. Pregitzer; David S. Ellsworth
2008-01-01
Elevated concentrations of atmospheric carbon dioxide ([CO2]) and tropospheric ozone ([O3]) have the potential to affect tree physiology and structure and hence forest water use, which has implications for climate feedbacks. We investigated how a 40% increase above ambient values in [CO2] and [O
M. E. Kubiske; V. S. Quinn; P. E. Marquardt; D. F. Karnosky
2007-01-01
Three model communities of trembling aspen (monoculture, and mixed with either paper birch or sugar maple) were grown for seven years in elevated atmospheric CO2 and O3 using Free Air CO2 Enrichment (FACE) technology. We utilized trends in species' importance, calculated as an index of volume...
John R. Jones; Norbert V. DeByle; Robert P. Winokur
1985-01-01
Aspen has not been cut extensively in the West; in fact, it has been grossly underutilized. For example, as recently as 1975, the aspen harvest from National Forests in four Forest Service regions in the Rocky Mountain area was 7.64 million board feet. Additional minor volumes were cut on special-use permits for products such as fuel and corral poles. The total amount...
George A. Schier; John R. Jones; Robert P. Winokur
1985-01-01
Aspen is noted for its ability to regenerate vegetatively by adventitious shoots or suckers that arise on its long lateral roots. It also produces sprouts from stumps and root collars; but they are not common. In a survey of regeneration after clearcutting mature aspen in Utah. Baker (1918b) found that 92% of the shoots originated from roots, 7% from root collars, and...
Rapid mortality of Populus tremuloides in southwestern Colorado, USA
James J. Worrall; Leanne Egeland; Thomas Eager; Roy A. Mask; Erik W. Johnson; Philip A. Kemp; Wayne D. Shepperd
2008-01-01
Concentrated patches of recent trembling aspen (Populus tremuloides) mortality covered 56,091 ha of Colorado forests in 2006. Mortality has progressed rapidly. Area affected increased 58% between 2005 and 2006 on the Mancos-Dolores Ranger District, San Juan National Forest, where it equaled nearly 10% of the aspen cover type. In four stands that were...
Scaling ozone responses of forest trees to the ecosystem level in a changing climate
D.F. Karnosky; K.S. Pregitzer; D.R. Zak; M.E. Kubiske; G.R. Hendrey; D. Weinstein; M. Nosal; K.E. Percy
2005-01-01
Many uncertainties remain regarding how climate change will alter the structure and function of forest ecosystems. At the Aspen FACE experiment in northern Wisconsin, we are attempting to understand how an aspen/birch/maple forest ecosystem responds to long-term exposure to elevated carbon dioxide (CO2) and ozone (O3),...
Sudden aspen decline in southwest Colorado: Site and stand factors and a hypothesis on etiology
Jim Worrall; Leanne Egeland; Tom Eager; Roy Mask; Erik Johnson; Phil Kemp; Wayne Shepperd
2008-01-01
An initial assessment of rapid dieback and mortality of aspen in southwest Colorado suggests that it represents a decline disease incited by acute, warm drought. Predisposing factors include low elevation, south and southwest aspects, droughty soils, open stands, and physiological maturity. Contributing factors include Cytospora canker, two bark beetles, poplar borer,...
Comparisons of SPORL and dilute acid pretreatments for sugar and ethanol productions from aspen
S. Tian; W. Zhu; Roland Gleisner; X.J. Pan; Junyong Zhu
2011-01-01
This study reports comparative evaluations of sugar and ethanol production from a native aspen (Populus tremuloides) between sulfite pretreatment to overcome recalcitrance of lignocelluloses (SPORL) and dilute acid (DA) pretreatments. All aqueous pretreatments were carried out in a laboratory wood pulping digester using wood chips at 170°C with a liquid to...
D.F. Karnosky; B. Mankovska; K. Percy; R.E. Dickson; G.K. Podila; J. Sober; A. Noormets; G. Hendrey; Mark D. Coleman; M. Kubiske; K.S. Pregitzer; J.G. Isebrands
1999-01-01
Abstract. Over the years, a series of trembling aspen (Populus tremuloides Michx.) clones differing in O3 sensitivity have been identified from OTC studies. Three clones (216 and 271[(O3 tolerant] and 259 [O3 sensitive]) have been characterized for O3...
Automated Planning for a Deep Space Communications Station
NASA Technical Reports Server (NTRS)
Estlin, Tara; Fisher, Forest; Mutz, Darren; Chien, Steve
1999-01-01
This paper describes the application of Artificial Intelligence planning techniques to the problem of antenna track plan generation for a NASA Deep Space Communications Station. Me described system enables an antenna communications station to automatically respond to a set of tracking goals by correctly configuring the appropriate hardware and software to provide the requested communication services. To perform this task, the Automated Scheduling and Planning Environment (ASPEN) has been applied to automatically produce antenna trucking plans that are tailored to support a set of input goals. In this paper, we describe the antenna automation problem, the ASPEN planning and scheduling system, how ASPEN is used to generate antenna track plans, the results of several technology demonstrations, and future work utilizing dynamic planning technology.
Bedding material affects mechanical thresholds, heat thresholds and texture preference
Moehring, Francie; O’Hara, Crystal L.; Stucky, Cheryl L.
2015-01-01
It has long been known that the bedding type animals are housed on can affect breeding behavior and cage environment. Yet little is known about its effects on evoked behavior responses or non-reflexive behaviors. C57BL/6 mice were housed for two weeks on one of five bedding types: Aspen Sani Chips® (standard bedding for our institute), ALPHA-Dri®, Cellu-Dri™, Pure-o’Cel™ or TEK-Fresh. Mice housed on Aspen exhibited the lowest (most sensitive) mechanical thresholds while those on TEK-Fresh exhibited 3-fold higher thresholds. While bedding type had no effect on responses to punctate or dynamic light touch stimuli, TEK-Fresh housed animals exhibited greater responsiveness in a noxious needle assay, than those housed on the other bedding types. Heat sensitivity was also affected by bedding as animals housed on Aspen exhibited the shortest (most sensitive) latencies to withdrawal whereas those housed on TEK-Fresh had the longest (least sensitive) latencies to response. Slight differences between bedding types were also seen in a moderate cold temperature preference assay. A modified tactile conditioned place preference chamber assay revealed that animals preferred TEK-Fresh to Aspen bedding. Bedding type had no effect in a non-reflexive wheel running assay. In both acute (two day) and chronic (5 week) inflammation induced by injection of Complete Freund’s Adjuvant in the hindpaw, mechanical thresholds were reduced in all groups regardless of bedding type, but TEK-Fresh and Pure-o’Cel™ groups exhibited a greater dynamic range between controls and inflamed cohorts than Aspen housed mice. PMID:26456764
Deacon, Nicholas John; Grossman, Jake Joseph; Schweiger, Anna Katharina; Armour, Isabella; Cavender-Bares, Jeannine
2017-12-01
Aspen groves along the Niobrara River in Nebraska have long been a biogeographic curiosity due to morphological differences from nearby remnant Populus tremuloides populations. Pleistocene hybridization between P. tremuloides and P. grandidentata has been proposed, but the nearest P. grandidentata populations are currently several hundred kilometers east. We tested the hybrid-origin hypothesis using genetic data and characterized putative hybrids phenotypically. We compared nuclear microsatellite loci and chloroplast sequences of Niobrara River aspens to their putative parental species. Parental species and putative hybrids were also grown in a common garden for phenotypic comparison. On the common garden plants, we measured leaf morphological traits and leaf-level spectral reflectance profiles, from which chemical traits were derived. The genetic composition of the three unique Niobrara aspen genotypes is consistent with the hybridization hypothesis and with maternal chloroplast inheritance from P. grandidentata . Leaf margin dentition and abaxial pubescence differentiated taxa, with the hybrids showing intermediate values. Spectral profiles allowed statistical separation of taxa in short-wave infrared wavelengths, with hybrids showing intermediate values, indicating that traits associated with internal structure of leaves and water absorption may vary among taxa. However, reflectance values in the visible region did not differentiate taxa, indicating that traits related to pigments are not differentiated. Both genetic and phenotypic results support the hypothesis of a hybrid origin for these genetically unique aspens. However, low genetic diversity and ongoing ecological and climatic threats to the hybrid taxon present a challenge for conservation of these relictual boreal communities. © 2017 Botanical Society of America.
Couture, John J; Meehan, Timothy D; Rubert-Nason, Kennedy F; Lindroth, Richard L
2017-01-01
Anthropogenic activities are altering levels of atmospheric carbon dioxide (CO 2 ) and tropospheric ozone (O 3 ). These changes can alter phytochemistry, and in turn, influence ecosystem processes. We assessed the individual and combined effects of elevated CO 2 and O 3 on the phytochemical composition of two tree species common to early successional, northern temperate forests. Trembling aspen (Populus tremuloides) and paper birch (Betula papyrifera) were grown at the Aspen FACE (Free-Air Carbon dioxide and ozone Enrichment) facility under four combinations of ambient and elevated CO 2 and O 3 . We measured, over three years (2006-08), the effects of CO 2 and O 3 on a suite of foliar traits known to influence forest functioning. Elevated CO 2 had minimal effect on foliar nitrogen and carbohydrate levels in either tree species, and increased synthesis of condensed tannins and fiber in aspen, but not birch. Elevated O 3 decreased nitrogen levels in both tree species and increased production of sugar, condensed tannins, fiber, and lignin in aspen, but not birch. The magnitude of responses to elevated CO 2 and O 3 varied seasonally for both tree species. When co-occurring, CO 2 offset most of the changes in foliar chemistry expressed under elevated O 3 alone. Our results suggest that levels of CO 2 and O 3 predicted for the mid-twenty-first century will alter the foliar chemistry of northern temperate forests with likely consequences for forest community and ecosystem dynamics.
Randy Kolka; Aaron Steber; Ken Brooks; Charles H. Perry; Matt Powers
2012-01-01
Although a number of harvesting studies have assessed compaction, no study has considered the interacting relationships of harvest season, soil texture, and landscape position on soil bulk density and surface soil strength for harvests in the western Lake States. In 2005, we measured bulk density and surface soil strength in recent clearcuts of predominantly aspen...
D. F. Karnosky; B. Mankovska; K. Percy; R. E. Dickson; G. K. Podila; J. Sober; A. Noormets; G. Hendrey; M. D. Coleman; M. Kubiske; K. S. Pregitzer; J. G. Isebrands
1999-01-01
Over the years, a series of trembling aspen (Populus tremuloides Michx.) clones differing in O3 sensitivity have been identified from OTC studies. Three clones (216 and 271[O3 tolerant] and 259 [O3 sensitive]) have been characterized for O3 sensitivity by growth and biomass...
Aspen fencing in northern Arizona: A 15-year perspective
James M. Rolf
2001-01-01
Aspen clearcuts in the 1960s and 1970s on the Peaks Ranger District of the Coconino National Forest in northern Arizona failed to regenerate successfully because of browsing primarily by elk. Since 1985, over 400 acres have been successfully regenerated using fencing of various designs to exclude elk. The expense and visual impact of establishing and maintaining over...
Mechanical properties of aspen
R.P.A. Johnson
1947-01-01
The clear wood of aspen has inherent properties of merit. Several of these properties are familiar to a large proportion of the general public, for some of the common types of matches are made of this wood. The requirements of wood for matches are exacting in that the wood must combine straightness of grain, ease of splitting, ease of working, and toughness. To meet...
Muzaffer Ahmet Karaaslan; Mandla A. Tshabalala; Gisela Buschle-Diller
2012-01-01
Semi-interpenetrating network hydrogel films were prepared using hemicellulose and chemically crosslinked chitosan. Hemicellulose was extracted from aspen by using a novel alkaline treatment and characterized by HPSEC, and consisted of a mixture of high and low molecular weight polymeric fractions. HPLC analysis of the acid hydrolysate of the hemicellulose showed that...
Leading from the Front of the Classroom: A Roadmap to Teacher Leadership That Works
ERIC Educational Resources Information Center
Aspen Institute, 2014
2014-01-01
In this paper, Leading Educators and the Aspen Institute propose a roadmap to empower teachers to lead from the front of the classroom. This paper outlines key phases that system administrators will need to consider as they build teacher leadership systems that address their highest priorities. For each phase, the Aspen Institute offers a…
Nicholas J. Brazee; Daniel L. Lindner; Shawn Fraver; Anthony W. D' Amato; Amy M. Milo
2012-01-01
To better understand the potential long-term effects of biomass harvesting on biodiversity, the polyporoid fungi community was characterized from 120 plots in four aspen-dominated forests in Minnesota. Four deadwood variables (substratum species, substratum type, decay class and diameter class) were recorded for each polyporoid species occurrence. A total of 2358...
Time-domain NMR study of the drying of hemicellulose extracted aspen (Populus tremuloides Michx.)
Thomas Elder; Carl Houtman
2013-01-01
The effect of hot water on aspen chips has been evaluated using time-domain low-field nuclear magnetic resonance (NMR) spectroscopy. At moisture contents above fiber saturation point, treated chips exhibit relaxation times of free water longer than for the control. This is consistent with the removal of hemicelluloses given the hydrophilicity of these polysaccharides....
Federal Register 2010, 2011, 2012, 2013, 2014
2013-01-25
... Pass, RNAV (GPS)-A, Amdt 1 Jonesboro, AR, Jonesboro Muni, ILS OR LOC RWY 23, Amdt 2 Jonesboro, AR, Jonesboro Muni, VOR RWY 23, Amdt 11 Melbourne, AR, Melbourne Muni--John E Miller Field, RNAV (GPS) RWY 3, Amdt 1 Melbourne, AR, Melbourne Muni--John E Miller Field, RNAV (GPS) RWY 21, Amdt 1 Aspen, CO, Aspen...
Paul C. Rogers; Kori D. Moore; Ronald J. Ryel
2009-01-01
Forty-seven randomly selected mid-elevation aspen stands were sampled for lichens and stand conditions. Plots were characterized according to tree species cover, basal area, stand age, bole scarring, tree damage, and presence of lichen species. We also recorded ammonia emissions with passive sensors at 25 urban and agricultural sites throughout an adjacent populated...
Preliminary study on flakeboard panels made from aspen slash wood
Yan Yu; Alan Rudie; Zhiyong Cai
2010-01-01
The disposal of forest-thinning residue is one of the major problems for sustainable forest management. The purpose of this study was to investigate the technical possibility of utilizing aspen logging slash wood with a diameter ranging from 50 to 76 mm for flakeboard production. Influences of weight ratio between slash wood and commercial flakes on the selected...
Y. Xue; D.R. Veazie; C. Glinsey; M.F. Horstemeyer; R.M. Rowell
2007-01-01
The mechanical properties of newly developed aspen fiberâpolypropylene composites (APC) were experimentally explored and numerically predicted at the temperatures and humidity that are typical for domestic housing applications. The mechanical properties of APCs with five different fiber-loadings were evaluated at the room temperature, 4 [degrees] C, and 40 [degrees] C...
W. A. Laycock; Dale Bartos
1999-01-01
Vegetation was sampled inside and outside eight exclosures in salt desert shrub and sagebrush vegetation types in Southwestern Wyoming and eight exclosures in aspen vegetation in southern Utah. Only species richness has been examined thus far. Five of the eight Wyoming exclosures had an average of 11% more plant species present outside the exclosure than inside.
Dana L. Richter; Jessie A. Glaeser
2015-01-01
Two isolates of Chlorociboria aeruginascens (Nyl.) Kanouse incubated axenically on aspen wood blocks resulted in 18% and 32% mass loss after 134 wks (2 yrs 8 mo). Aspen wood decayed by C. aeruginascens contained cavities in the S2 layer of the secondary cell wall, similar to Type I soft rot attack, as well as erosion troughs and...
D. F. Karnosky; D. R. Zak; K. S. Pregitzer; C. S. Awmack; J. G. Bockheim; R. E. Dickson; G. R. Hendrey; G. E. Host; J. S. King; B. J. Kopper; E. L. Kruger; M. E. Kubiske; R. L. Lindroth; W. J. Mattson; E. P. McDonald; A. Noormets; E. Oksanen; W. F. J. Parsons; K. E. Percy; G. K. Podila; D. E. Riemenschneider; P. Sharma; R. Thakur; A. S& #244ber; J. S& #244ber; W. S. Jones; S. Anttonen; E. Vapaavuori; B. Mankovska; W. Heilman; J. G. Isebrands
2003-01-01
1. The impacts of elevated atmospheric CO2 and/or O3 have been examined over 4 years using an open-air exposure system in an aggrading northern temperate forest containing two different functional groups (the indeterminate, pioneer, 03-sensitive species Trembling Aspen, Populus tremuloides...
W. Zhu; Carl J. Houtman; J.Y. Zhu; Roland Gleisner; K.F. Chen
2012-01-01
A combined hydrolysis factor (CHF) was developed to predict xylan hydrolysis during pretreatments of native aspen (Populus tremuloides) wood chips. A natural extension of previously developed kinetic models allowed us to account for the effect of catalysts by dilute acid and two sulfite pretreatments at different pH values....
Grizelle Gonzalez; William Gould; Andrew T. Hudak; Teresa Nettleton Hollingsworth
2008-01-01
In this study, we set up a wood decomposition experiment to i) quantify the percent of mass remaining, decay constant and performance strength of aspen stakes (Populus tremuloides) in dry and moist boreal (Alaska and Minnesota, USA), temperate (Washington and Idaho, USA), and tropical (Puerto Rico) forest types, and ii) determine the effects of...
Nitrogen mineralization in aspen/conifer soils after a natural fire
Michael C. Amacher; Dale L. Bartos; Tracy Christopherson; Amber D. Johnson; Debra E. Kutterer
2001-01-01
We measured the effects of the 1996 Pole Creek fire, Fishlake National Forest, Utah, on available soil N and net N mineralization for three summers after the fire using an ion exchange membrane (IEM) soil core incubation method. Fire in mixed aspen/conifer increased the amount of available NH4, and a subsequent net increase in soil nitrification was observed. Release...
Prehydrolysis of aspen wood with water and with dilute aqueous sulfuric acid
Edward L. Springer; John F. Harris
1982-01-01
Water prehydrolysis of aspen wood was compared with 0.40% sulfuric acid prehydrolysis at a reaction temperature of 170°C. Acid prehydrolysis gave much higher yields of total anhydroxylose units in the prehydrolyzate and removed significantly less anhydroglucose from the wood than did the water treatment. At maximum yields of total anhydroxylose units in the...
Root growth and physiology of potted and field-grown trembling aspen exposed to tropospheric ozone
M.D. Coleman; R.E. Dickson; J.G. Isebrands; D.F. Karnosky
1996-01-01
We studied root growth and respiration of potted plants and field-grown aspen trees (Populus tremuloides Michx.) exposed to ambient or twice-ambient ozone. Root dry weight of potted plants decreased up to 45% after 12 weeks of ozone treatment, and root system respiration decreased by 27%. The ozone-induced decrease in root system respiration of...
Summary of nutrient and biomass data from two aspen sites in western United States
Robert S. Johnston; Dale L. Bartos
1977-01-01
Summary tables are presented for aboveground biomass and nutrient concentrations for 20 aspen trees (Populus tremuloides Michx.) that were sampled at two study sites in Utah and Wyoming. Trees were divided into seven components - leaves, current twigs, old twigs, deadwood (branches), branches, bark, and bole wood. Samples from each component were analyzed for nitrogen...
Surface compaction estimates and soil sensitivity in Aspen stands of the Great Lakes States
Aaron Steber; Ken Brooks; Charles H. Perry; Randy Kolka
2007-01-01
Aspen forests in the Great Lakes States support much of the regional timber industry. Management-induced soil compaction is a concern because it affects forest health and productivity and soil erosion. Soil compaction increases bulk density and soil strength and can also decrease air and water movement into and through the soil profile. Currently, most inventories, and...
Aspen clearcutting increases snowmelt and storm flow peaks in north central Minnesota
Elon S. Verry; Jeffrey R. Lewis; Kenneth N. Brooks
1983-01-01
Clearcutting aspen from the upland portion of an upland peatland watershed in north central Minnesota caused snowmelt peak discharge to increase 11 to 143 percent. Rainfall peak discharge size increased as much as 250 percent during the first two years after clearcutting, then decreased toward precutting levels in subsequent years. Storm flow volumes from rain during...
Douglas M. Stone; Richard Kabzems
2002-01-01
Forest management practices that decrease soil porosity and remove organic matter can reduce site productivity. We evaluated effects of four treatments-merchantable bole harvest (MBH) with three levels of soil compaction (none, light, or heavy), and total woody vegetation harvest plus forest floor removal (FFR)-on fifth-year regeneration and growth of aspen (...
Structural lumber properties of hybrid poplar
David E. Kretschmann; J. G.. Isebrands; Glen. Stanosz; John R. Dramm; Adele. Olstad; David. Cole; Jay. Samsel
The Lake States of Michigan, Minnesota, and Wisconsin consist of 49.4 million hectares (122 million acres) of land, 19.8 million hectares (49 million acres) of which are forested, with 6% of those having been removed by law from timber production. Aspen (Populus tremuloides) is an important component of this resource. In recent years, a concern that the aspen cut will...
W. A. Laycock; Dale Bartos; Keith Klement
2001-01-01
Recent conservation biology and environmental literature contains claims that livestock grazing has caused and continues to cause reduction in species diversity on Western rangelands, especially public rangelands. This paper present quantitative data on species richness (number of species) inside and outside 24 long-term exclosures; 8 exclosures in aspen vegetation in...
April M. Brough; R. Justin DeRose; Mary M. Conner; James N. Long
2017-01-01
Understanding the degree of spatial fidelity exhibited by individuals within a species increases our ability to manage for desired future outcomes. Elk (Cervus elaphus) is a closely managed species in the Western US, but there is little research evaluating their summer home-range fidelity. Elk summer-fall homeranges overlap considerably with aspen (Populus tremuloides...
Jack Pine and Aspen Forest Floors in Northeastern Minnesota
Robert M. Loomis
1977-01-01
Characteristics of upland forest floors under mature jack pine and aspen in northeastern Minnesota were investigated. These fuel measurements were needed as inputs for fire behavior prediction models -- useful for fire management decisions. The forest floor weight averaged 33,955 kg/ha and depth averaged 7.1 cm. Bulk density averaged 17 kg/m3 for the L (litter)...
Multiple factors affect aspen regeneration on the Uncompahgre Plateau, west-central Colorado
Barry C. Johnston
2001-01-01
In 1996, I inventoried over 90 aspen stands in 12 timber sales that had been clearcut >3 years previously. Units that regenerated adequately were larger, had higher slope angles, and had soils with a thick Mollic surface layer. Units that regenerated inadequately often had plant species that indicated high water tables. The factors associated with inadequate...
Pooja Sharma; Anu Sober; Jaak Sober; Gopi P. Podila; Mark E. Kubiske; William J. Mattson; Judson G. Isebrands; David F. Karnosky
2003-01-01
The greenhouse gases CO2 and 03 are increasing in the earth's atmosphere. Little is known about long-term impacts of these two co-occurring gases on forest trees. We have been examining the impacts of these two gases on the physiology and growth of trembling aspen (Populus tremuloides) and sugar...
Automated Design Space Exploration with Aspen
DOE Office of Scientific and Technical Information (OSTI.GOV)
Spafford, Kyle L.; Vetter, Jeffrey S.
Architects and applications scientists often use performance models to explore a multidimensional design space of architectural characteristics, algorithm designs, and application parameters. With traditional performance modeling tools, these explorations forced users to first develop a performance model and then repeatedly evaluate and analyze the model manually. These manual investigations proved laborious and error prone. More importantly, the complexity of this traditional process often forced users to simplify their investigations. To address this challenge of design space exploration, we extend our Aspen (Abstract Scalable Performance Engineering Notation) language with three new language constructs: user-defined resources, parameter ranges, and a collection ofmore » costs in the abstract machine model. Then, we use these constructs to enable automated design space exploration via a nonlinear optimization solver. We show how four interesting classes of design space exploration scenarios can be derived from Aspen models and formulated as pure nonlinear programs. The analysis tools are demonstrated using examples based on Aspen models for a three-dimensional Fast Fourier Transform, the CoMD molecular dynamics proxy application, and the DARPA Streaming Sensor Challenge Problem. Our results show that this approach can compose and solve arbitrary performance modeling questions quickly and rigorously when compared to the traditional manual approach.« less
Automated Design Space Exploration with Aspen
Spafford, Kyle L.; Vetter, Jeffrey S.
2015-01-01
Architects and applications scientists often use performance models to explore a multidimensional design space of architectural characteristics, algorithm designs, and application parameters. With traditional performance modeling tools, these explorations forced users to first develop a performance model and then repeatedly evaluate and analyze the model manually. These manual investigations proved laborious and error prone. More importantly, the complexity of this traditional process often forced users to simplify their investigations. To address this challenge of design space exploration, we extend our Aspen (Abstract Scalable Performance Engineering Notation) language with three new language constructs: user-defined resources, parameter ranges, and a collection ofmore » costs in the abstract machine model. Then, we use these constructs to enable automated design space exploration via a nonlinear optimization solver. We show how four interesting classes of design space exploration scenarios can be derived from Aspen models and formulated as pure nonlinear programs. The analysis tools are demonstrated using examples based on Aspen models for a three-dimensional Fast Fourier Transform, the CoMD molecular dynamics proxy application, and the DARPA Streaming Sensor Challenge Problem. Our results show that this approach can compose and solve arbitrary performance modeling questions quickly and rigorously when compared to the traditional manual approach.« less
NASA Astrophysics Data System (ADS)
Soderquist, B.; Kavanagh, K.; Link, T. E.; Seyfried, M. S.; Strand, E. K.
2016-12-01
Precipitation regimes in many semiarid ecosystems are becoming increasingly dominated by winter rainfall as a result of climate change. Across these regions, snowpack plays a vital role in the distribution and timing of soil moisture availability. Rising temperatures will result in a more uniform distribution of soil moisture, advanced spring phenology, and prolonged growing seasons. Productive and wide ranging tree species like aspen, Populus tremuloides, may experience increased vulnerability to drought and mortality resulting from both reduced snowpack and increased evaporative demand during the growing season. We simulated the net primary production (NPP) of aspen stands spanning the rain:snow transition zone in the Reynolds Creek Critical Zone Observatory (RCCZO) in southwest Idaho, USA. Within the RCCZO, the total amount of precipitation has remained unchanged over the past 50 years, however the percentage of the precipitation falling as snow has declined by approximately 4% per decade at mid-elevation sites. The biogeochemical process model Biome-BGC was used to simulate aspen NPP at three stands located directly below snowdrifts that provide melt water late into the spring. After adjusting precipitation inputs to account for the redistribution of snow, we assessed climate change impacts on future aspen productivity. Mid-century (2046-2065) aspen NPP was simulated using temperature projections from a multi-model average under high emission conditions using the Multivariate Adaptive Constructed Analogs (MACA) data set. While climate change simulations indicated over a 20% decrease in annual NPP for some years, NPP rates for other mid-century years remained relatively unchanged due to variations in growing season conditions. Mid-century years with the largest decreases in NPP typically showed increased spring transpiration rates resulting from earlier leaf flush combined with warmer spring conditions. During these years, the onset of drought stress occurred earlier due to increased early season soil moisture use and higher summer vapor pressure deficits. These results indicate that vegetation response to decreased snowpack can result in significant drought stress although phenological shifts that better align leaf production and precipitation ameliorate this response in some years. Precipitation regimes in many semiarid ecosystems are becoming increasingly dominated by winter rainfall as a result of climate change. Across these regions, snowpack plays a vital role in the distribution and timing of soil moisture availability. Rising temperatures will result in a more uniform distribution of soil moisture, advanced spring phenology, and prolonged growing seasons. Productive and wide ranging tree species like aspen, Populus tremuloides, may experience increased vulnerability to drought and mortality resulting from both reduced snowpack and increased evaporative demand during the growing season. We simulated the net primary production (NPP) of aspen stands spanning the rain:snow transition zone in the Reynolds Creek Critical Zone Observatory (RCCZO) in southwest Idaho, USA. Within the RCCZO, the total amount of precipitation has remained unchanged over the past 50 years, however the percentage of the precipitation falling as snow has declined by approximately 4% per decade at mid-elevation sites. The biogeochemical process model Biome-BGC was used to simulate aspen NPP at three stands located directly below snowdrifts that provide melt water late into the spring. After adjusting precipitation inputs to account for the redistribution of snow, we assessed climate change impacts on future aspen productivity. Mid-century (2046-2065) aspen NPP was simulated using temperature projections from a multi-model average under high emission conditions using the Multivariate Adaptive Constructed Analogs (MACA) data set. While climate change simulations indicated over a 20% decrease in annual NPP for some years, NPP rates for other mid-century years remained relatively unchanged due to variations in growing season conditions. Mid-century years with the largest decreases in NPP typically showed increased spring transpiration rates resulting from earlier leaf flush combined with warmer spring conditions. During these years, the onset of drought stress occurred earlier due to increased early season soil moisture use and higher summer vapor pressure deficits. These results indicate that vegetation response to decreased snowpack can result in significant drought stress although phenological shifts that better align leaf production and precipitation ameliorate this response in some years.
Lumber Grade Yields for Graded Aspen Logs and Trees
Leland F. Hanks; Robert L. Brisbin
1978-01-01
Green lumber grade yields for aspen were determined for use with the U.S. Forest Service hardwood log and tree grades. The yields for logs are expressed in percent of total lumber tally volume, and those for trees are expressed in board feet. Overruns for the International 1/4-inch and Scribner log rules along with lumber recovery factors are shown by log grade.
Jessica E. Hancock; Kate L. Bradley; Christian P. Giardina; Kurt S. Pregitzer
2008-01-01
Plants influence soil carbon (C) formation through the quality and quantity of C released to soil. Soil type, in turn can modify a plant's influence on soil through effects on plant production, tissue quality and regulation of soil C decomposition and stabilization. Wild-type aspen and three transgenic aspen lines expressing reduced stem lignin concentrations and/...
Hydrologic recovery of aspen clearcuts in northwestern Alberta
R. H. Swanson; R. L. Rothwell
2001-01-01
A 3-year study of evapotranspiration from aspen clearcuts 1 to 14 years of age indicated the following: (1) The annual evapotranspiration from 1- to 5-year-old clearcuts ranges from 0 to 143 mm less than a mature forest on the same site. Evapotranspiration is highly dependent upon the amount of precipitation. (2) These effects can vanish in as few as 2 years with low...
Controlling Hazel, Aspen Suckers, and Mountain Maple with Picloram
Donald A. Perala
1971-01-01
Tests showed that picloram/2,4-D mixture was equal to or superior to 2,4-D alone or a 2,4,5-D/2,4,5-T mixture in controlling hazel, aspen suckers, and mountain maple for reforestation purposes. Survival of red pine planted 9 months after treatment was not influenced by residual soil effects of picloram. However, foliar application contributed to mortality of...
Chris Polson
2001-01-01
Aspen wood, when carefully selected and kiln dried, makes excellent stock for artist stretcher frames. Direct marketing techniques including the Internet and word of mouth give access to national markets, providing a more diverse and stable customer base for operations from a rural area. High-quality service, as shown by product performance and rapid order fulfillment...
Effect of citric acid modification of aspen wood on sorption of copper ion
James D. McSweeny; Roger M. Rowell; Soo Hong Min
2006-01-01
Milled aspen wood was thermochemically modified with citric acid for the purpose of improving the copper (Cu2+) ion sorption capacity of the wood when tested in 24-hour equilibrium batch tests. The wood-citric acid adducts provided additional carboxyl groups to those in the native wood and substantially increased Cu2+ ion uptake of the modified wood compared with that...
William L. Headlee; Richard B. Hall; Ronald S., Jr. Zalesny
2013-01-01
Hybrid poplars have demonstrated high productivity as short rotation woody crops (SRWC) in the Midwest USA, and the hybrid aspen "Crandon" (Populus alba L. × P. grandidenta Michx.) has exhibited particularly promising yields on marginal lands. However, a key obstacle for wider deployment is the lack of economic...
Growth response to fertilizer in a young aspen-birch stand
Miroslaw M. Czapowskyj; Lawrence O. Safford
1978-01-01
A thinned aspen-birch-red maple stand was fertilized with N, P, and N plus P, both with and without lime (L). Overall, treatments with N increased height growth by an average of 79 percent, and volume growth by 69 percent, over treatments without N. Lime tended to increase both average height and volume growth over each corresponding treatment without lime. The amount...
Rebecca E. Ibach; Roger M. Rowell; Sandra E. Lange; Rebecca L. Schumann
2002-01-01
Aspen fiber-polypropylene composites were prepared with various levels of fiber (0,30%, 40%, 50%, and 60%), polypropylene (PP) (100%, 98%, 70%, 68%, 60%, 58%, 50%, 48%, 40%, and 38%), and the compatibilizer maleated polypropylene (MAPP) (0 and 2%). Specimens were either subjected to 10 cycles of 1 week room temperature water soaking-oven drying or 2-hr. boiling...
Janine Rice; Tim Bardsley; Pete Gomben; Dustin Bambrough; Stacey Weems; Allen Huber; Linda A. Joyce
2017-01-01
Aspen ecosystems are valued because they add biodiversity and ecological value to the landscape. They provide rich and productive habitats and increase aesthetic value. Climate change poses the risk of altering and disrupting these ecosystems, and it may worsen the effects of non-climate stressors. To provide scientific information for land managers facing the...
Feeding ecology of sharp-shinned hawks in deciduous and coniferous forests in Colorado
Suzanne M. Joy; Richard T. Reynolds; Richard L. Knight; Richard W. Hoffman
1994-01-01
Feeding ecology of 11 Sharp-skinned Hawk (Accipiter striates) pairs nesting in aspen (Populus tremuloides), conifer (Abies, Picea spp.), and mixed aspen-conifer habitats in southwest Colorado was investigated during 1988-1989. Small birds (x-bar = 20.9 g, SE = 0.8 g) and mammals (x-bar = 41.1 g, SE = 3.3 g) comprised 91 and 9% of...
A severe epidemic of Marssonina leaf blight on quaking aspen in Northern Utah
Roy O. Harniss; David L. Nelson
1984-01-01
The extent of Marssonina leaf blight (Marssonina populi) on quaking aspen (Populus tremuloides) was observed in northern Utah and adjacent States in 1981 aand 1982. Area of the epidemic and symptoms of the disease are described. On 1,000 acres (405 hal) in northern Utah, infection levels were 6 percent slight, 12 percent light, 32 percent moderate, 16 percent...
Emmanuel Ebanyenle; Andrew J. Burton; Andrew J. Storer; Dana L. Richter; Jessie A. Glaeser
2016-01-01
We examined the effects of elevated CO2 and/or O3 on the wood-decaying basidiomycete fungal community and wood decomposition rates at the Aspen Free-Air CO2 and O3 Enrichment (Aspen FACE) project. Mass loss rates were determined after one year of log decomposition on the soil...
J.Y. Zhu; R. Gleisner; C.T. Scott; X.L. Luo; S. Tian
2011-01-01
Native aspen (Populus tremuloides) was pretreated using sulfuric acid and sodium bisulfite (SPORL) and dilute sulfuric acid alone (DA). Simultaneous enzymatic saccharification and fermentation (SSF) was conducted at 18% solids using commercial enzymes with cellulase loadings ranging from 6 to 15 FPU/g glucan and Saccharomyces cerevisiae Y5. Compared with DA...
Richard Voldseth; Brian J. Palik; John Elioff
2011-01-01
Impacts of organic matter removal and compaction on soil properties and productivity are reported from the first 10 years of the Long-Term Soil Productivity Study in Great Lakes aspen ecosystems. Organic matter removal treatments included main bole, total tree harvest, and total tree harvest with forest floor removal. Compaction treatments included minimal compaction,...
Multi-scale habitat use of male ruffed grouse in the Black Hills National Forest
Cassandra L. Mehls; Kent C. Jensen; Mark A. Rumble; Michael C. Wimberly
2014-01-01
Ruffed grouse (Bonasa umbellus) are native upland game birds and a management indicator species (MIS) for aspen (Populus tremuloides) in the Black Hills National Forest (Black Hills). Our objective was to assess resource selection of male ruffed grouse to identify the most appropriate scale to manage for aspen and ruffed grouse in the Black Hills. During spring 2007...
ERIC Educational Resources Information Center
Adler, Richard P.; Breivik, Patricia Senn
This report is an informed observer's interpretation of the discussions that took place at the 1998 annual meeting of the Aspen Institute's Forum on Communications and Society (FOCAS). It summarizes the inquiry made by FOCAS members into the many issues surrounding information literacy, including what information literacy is, why we need an…
Regeneration of aspen by suckering on burned sites in western Wyoming
Dale L. Bartos; Walter F. Mueggler; Robert B. Campbell
1991-01-01
Numbers of suckers produced following burning of aspen stands varied greatly between areas but followed similar trends over a 6-year period: a relative abundance the first year, followed by an abrupt decline by the third year and a gradual decline thereafter. First-year sucker numbers that ranged from 34,000 to 147,000/ha suffered mortality between 54 and 93 percent by...
Mogensen, Kris M; Andrew, Benjamin Y; Corona, Jasmine C; Robinson, Malcolm K
2016-07-01
The Society of Critical Care Medicine (SCCM) and American Society for Parenteral and Enteral Nutrition (ASPEN) recommend that obese, critically ill patients receive 11-14 kcal/kg/d using actual body weight (ABW) or 22-25 kcal/kg/d using ideal body weight (IBW), because feeding these patients 50%-70% maintenance needs while administering high protein may improve outcomes. It is unknown whether these equations achieve this target when validated against indirect calorimetry, perform equally across all degrees of obesity, or compare well with other equations. Measured resting energy expenditure (MREE) was determined in obese (body mass index [BMI] ≥30 kg/m(2)), critically ill patients. Resting energy expenditure was predicted (PREE) using several equations: 12.5 kcal/kg ABW (ASPEN-Actual BW), 23.5 kcal/kg IBW (ASPEN-Ideal BW), Harris-Benedict (adjusted-weight and 1.5 stress-factor), and Ireton-Jones for obesity. Correlation of PREE to 65% MREE, predictive accuracy, precision, bias, and large error incidence were calculated. All equations were significantly correlated with 65% MREE but had poor predictive accuracy, had excessive large error incidence, were imprecise, and were biased in the entire cohort (N = 31). In the obesity cohort (n = 20, BMI 30-50 kg/m(2)), ASPEN-Actual BW had acceptable predictive accuracy and large error incidence, was unbiased, and was nearly precise. In super obesity (n = 11, BMI >50 kg/m(2)), ASPEN-Ideal BW had acceptable predictive accuracy and large error incidence and was precise and unbiased. SCCM/ASPEN-recommended body weight equations are reasonable predictors of 65% MREE depending on the equation and degree of obesity. Assuming that feeding 65% MREE is appropriate, this study suggests that patients with a BMI 30-50 kg/m(2) should receive 11-14 kcal/kg/d using ABW and those with a BMI >50 kg/m(2) should receive 22-25 kcal/kg/d using IBW. © 2015 American Society for Parenteral and Enteral Nutrition.
Earnst, Susan L.; Dobkin, David S.; Ballard, Jennifer A.
2012-01-01
Riparian and quaking aspen (Populus tremuloides) woodlands are centers of avian abundance and diversity in the western United States, but they have been affected adversely by land use practices, particularly livestock grazing. In 1990, cattle were removed from a 112,500-ha national wildlife refuge in southeastern Oregon. Thereafter, we monitored changes in vegetation and bird abundance in years 1–3 (phase 1) and 10–12 (phase 2) in 17 riparian and 9 snow-pocket aspen plots. On each 1.5-ha plot, we sampled vegetation in 6 transects. Three times during each breeding season, observers recorded all birds 50 m to each side of the plot's 150-m centerline for 25 minutes. We analyzed data with multivariate analysis of variance and paired t tests with p values adjusted for multiple comparisons. In both periods, riparian and snow-pocket aspen produced extensive regeneration of new shoots (x̄ = 2646 stems/ha and 7079 stems/ha, respectively). By phase 2, a 64% increase in medium-diameter trees in riparian stands indicated successful recruitment into the overstory, but this pattern was not seen in snow-pocket stands, where the density of trees was over 2 times greater. By phase 2 in riparian and snow-pocket stands, native forb cover had increased by 68% and 57%, respectively, mesic shrub cover had increased by 29% and 58%, and sagebrush cover had decreased by 24% and 31%. Total avian abundance increased by 33% and 39% in riparian and snow-pocket aspen, respectively, ground or understory nesters increased by 133% and 67% and overstory nesters increased by 34% and 33%. Similarly, ground or understory foragers increased by 25% and 32%, aerial foragers by 55% and 57%, and overstory foragers by 66% and 43%. We interpreted the substantial regeneration of aspen shoots, increased densities of riparian forbs and shrubs, and increased avian abundances as a multitrophic-level response to the total removal of livestock and as substantial movement toward recovery of biological integrity.
Riikonen, Johanna; Kets, Katre; Darbah, Joseph; Oksanen, Elina; Sober, Anu; Vapaavuori, Elina; Kubiske, Mark E; Nelson, Neil; Karnosky, David F
2008-02-01
Paper birch (Betula papyrifera Marsh.) and three trembling aspen clones (Populus tremuloides Michx.) were studied to determine if alterations in carbon gain in response to an elevated concentration of CO(2) ([CO(2)]) or O(3) ([O(3)]) or a combination of both affected bud size and carbohydrate composition in autumn, and early leaf development in the following spring. The trees were measured for gas exchange, leaf size, date of leaf abscission, size and biochemical characteristics of the overwintering buds and early leaf development during the 8th-9th year of free-air CO(2) and O(3) exposure at the Aspen FACE site located near Rhinelander, WI. Net photosynthesis was enhanced 49-73% by elevated [CO(2)], and decreased 13-30% by elevated [O(3)]. Elevated [CO(2)] delayed, and elevated [O(3)] tended to accelerate, leaf abscission in autumn. Elevated [CO(2)] increased the ratio of monosaccharides to di- and oligosaccharides in aspen buds, which may indicate a lag in cold acclimation. The total carbon concentration in overwintering buds was unaffected by the treatments, although elevated [O(3)] decreased the amount of starch by 16% in birch buds, and reduced the size of aspen buds, which may be related to the delayed leaf development in aspen during the spring. Elevated [CO(2)] generally ameliorated the effects of elevated [O(3)]. Our results show that both elevated [CO(2)] and elevated [O(3)] have the potential to alter carbon metabolism of overwintering buds. These changes may cause carry-over effects during the next growing season.
Cellulose synthase stoichiometry in aspen differs from Arabidopsis and Norway spruce.
Zhang, Xueyang; Dominguez, Pia Guadalupe; Kumar, Manoj; Bygdell, Joakim; Miroshnichenko, Sergey; Sundberg, Bjorn; Wingsle, Gunnar; Niittyla, Totte
2018-05-14
Cellulose is synthesised at the plasma membrane by cellulose synthase complexes (CSCs) containing cellulose synthases (CESAs). Genetic analysis and CESA isoform quantification indicate that cellulose in the secondary cell walls of Arabidopsis (Arabidopsis thaliana) is synthesised by isoforms CESA4, CESA7 and CESA8 in equimolar amounts. Here, we used quantitative proteomics to investigate whether the CSC model based on Arabidopsis secondary cell wall CESA stoichiometry can be applied to the angiosperm tree aspen (Populus tremula) and the gymnosperm tree Norway spruce (Picea abies). In the developing xylem of aspen the secondary cell wall CESA stoichiometry was 3:2:1 for PtCESA8a/b : PtCESA4 : PtCESA7a/b, while in Norway spruce the stoichiometry was 1:1:1 as previously observed in Arabidopsis. Furthermore, in aspen tension wood the secondary cell wall CESA stoichiometry changed to 8:3:1 for PtCESA8a/b : PtCESA4 : PtCESA7a/b. PtCESA8b represented 73% of the total secondary cell wall CESA pool, and quantitative PCR analysis of CESA transcripts in cryo-sectioned tension wood revealed increased PtCESA8b expression during formation of the cellulose-enriched gelatinous layer while the transcripts of PtCESA4, PtCESA7a/b and PtCESA8a decreased. A wide-angle X-ray scattering analysis showed that the shift in CESA stoichiometry in tension wood coincided with an increase in crystalline cellulose microfibril diameter suggesting that the CSC CESA composition influences microfibril properties. The aspen CESA stoichiometry results raise the possibility of alternative CSC models, and suggest that homomeric PtCESA8b complexes are responsible for cellulose biosynthesis in the gelatinous layer in tension wood. {copyright, serif} 2018 American Society of Plant Biologists. All rights reserved.
Forest floor leachate fluxes under six different tree species on a metal contaminated site.
Van Nevel, Lotte; Mertens, Jan; De Schrijver, An; Baeten, Lander; De Neve, Stefaan; Tack, Filip M G; Meers, Erik; Verheyen, Kris
2013-03-01
Trees play an important role in the biogeochemical cycling of metals, although the influence of different tree species on the mobilization of metals is not yet clear. This study examined effects of six tree species on fluxes of Cd, Zn, DOC, H(+) and base cations in forest floor leachates on a metal polluted site in Belgium. Forest floor leachates were sampled with zero-tension lysimeters in a 12-year-old post-agricultural forest on a sandy soil. The tree species included were silver birch (Betula pendula), oak (Quercus robur and Q. petraea), black locust (Robinia pseudoacacia), aspen (Populus tremula), Scots pine (Pinus sylvestris) and Douglas fir (Pseudotsuga menziesii). We show that total Cd fluxes in forest floor leachate under aspen were slightly higher than those in the other species' leachates, yet the relative differences between the species were considerably smaller when looking at dissolved Cd fluxes. The latter was probably caused by extremely low H(+) amounts leaching from aspen's forest floor. No tree species effect was found for Zn leachate fluxes. We expected higher metal leachate fluxes under aspen as its leaf litter was significantly contaminated with Cd and Zn. We propose that the low amounts of Cd and Zn leaching under aspen's forest floor were possibly caused by high activity of soil biota, for example burrowing earthworms. Furthermore, our results reveal that Scots pine and oak were characterized by high H(+) and DOC fluxes as well as low base cation fluxes in their forest floor leachates, implying that those species might enhance metal mobilization in the soil profile and thus bear a potential risk for belowground metal dispersion. Copyright © 2013 Elsevier B.V. All rights reserved.
Figured grain in aspen is heritable and not affected by graft-transmissible signals
Youran Fan; Kendal Rupert; Alex C. Wiedenhoeft; Keith Woeste; Christian Lexer; Richard Meilan
2013-01-01
Figure can add value to wood products, but its occurrence is unpredictable. A first step in reliably producing figured wood is determining whether it is faithfully transmitted to progeny via sexual and asexual reproduction. We describe a 26-year-old male aspen genotype, designated âCurly Poplarâ, which was shown to be a Populus à canescens hybrid using microsatellite...
Pine hollow exclosures: Effect of browsing on an aspen community sprayed with 2,4-D
Dale L. Bartos; Roy O. Harniss
1990-01-01
The Pine Hollow aspen (Populus tremuloides) exclosures on the Ashley National Forest in eastern Utah were sampled in 1984, 19 years after they were established. The effects of 2,4-D, wildlife, and cattle on plant succession were evaluated. Two exclosures were used to protect the sprayed area from (1) all animal use and (2) only livestock use. A third sprayed area was...
Michael T. Thompson
2009-01-01
There are two events occurring in Colorado that are concerning forest managers in Colorado. There is severe and widespread mortality of lodgepole pine due to the mountain pine beetle and aspen forests in some areas of the state have experienced widespread, severe, and rapid crown deterioration leading to mortality. Implementation of the Forest Inventory and Analysis...
M. M. Dudley; Jose Negron; N. A. Tisserat; W. D. Shepperd; W. R. Jacobi
2015-01-01
We analyzed a series of increment cores collected from 260 adult dominant or co-dominant quaking aspen (Populus tremuloides Michx.) trees from national forests across Colorado and southern Wyoming in 2009 and 2010. Half of the cores were collected from trees in stands with a high amount of crown dieback, and half were from lightly damaged stands. We define the level of...
ERIC Educational Resources Information Center
Rodriguez, America
This publication reports on a 2000 forum that included media business executives, financiers, academics, and other experts who examined: how minority media can leverage their historically strong community base to succeed in the new century; how they can overcome financial challenges; what measures they might adopt to keep abreast of opportunities…
Ruffed grouse (Bonasa umbellus) drumming log and habitat use in Grand Teton National Park, Wyoming
Buhler, M.L.; Anderson, S.H.
2001-01-01
We described 15 Ruffed Grouse (Bonasa umbellus) drumming logs and adjacent habitat within Grand Teton National Park, Wyoming. Drumming logs and adjacent habitat differed from 30 random non-drumming sites. Drumming logs had fewer limbs (8; P = 0.003) and a smaller percentage of bark remaining (12%; P = 0.0001). These logs were in advanced stages of decay but were still firm to the touch. Additionally, drumming logs were found close to clearings but in areas with increased amounts of undergrowth and mature trees. Adjacent habitat analysis (0.04-ha circular plot centered on logs) indicated drumming locations had significantly greater average canopy height, more vegetative cover consisting of conifer and total canopy cover, and more vertical foliage between 0.3 m and 3.0 m in height. Adjacent habitat was in advanced stages of maturity as indicated by significant numbers of both large-diameter logs and large-diameter lodgepole pine (Pinus contorta) and quaking aspen (Populus tremuloides) snags. Tree species dominating the canopy and subcanopy were large-diameter Engelmann spruce (Picea engelmannii), lodgepole pine, and quaking aspen. Subalpine fir (Abies lasiocarpa) and quaking aspen saplings were more numerous at used sites. Ruffed Grouse drummed in coniferous areas within close proximity of quaking aspen.
Chen, Lei; Huang, Jian-Guo; Dawson, Andria; Zhai, Lihong; Stadt, Kenneth J; Comeau, Philip G; Whitehouse, Caroline
2018-02-01
Insects, diseases, fire and drought and other disturbances associated with global climate change contribute to forest decline and mortality in many parts of the world. Forest decline and mortality related to drought or insect outbreaks have been observed in North American aspen forests. However, little research has been done to partition and estimate their relative contributions to growth declines. In this study, we combined tree-ring width and basal area increment series from 40 trembling aspen (Populus tremuloides Michx.) sites along a latitudinal gradient (from 52° to 58°N) in western Canada and attempted to investigate the effect of drought and insect outbreaks on growth decline, and simultaneously partition and quantify their relative contributions. Results indicated that the influence of drought on forest decline was stronger than insect outbreaks, although both had significant effects. Furthermore, the influence of drought and insect outbreaks showed spatiotemporal variability. In addition, our data suggest that insect outbreaks could be triggered by warmer early spring temperature instead of drought, implicating that potentially increased insect outbreaks are expected with continued warming springs, which may further exacerbate growth decline and death in North America aspen mixed forests. © 2017 John Wiley & Sons Ltd.
Analysis of Cryogenic Cycle with Process Modeling Tool: Aspen HYSYS
NASA Astrophysics Data System (ADS)
Joshi, D. M.; Patel, H. K.
2015-10-01
Cryogenic engineering deals with the development and improvement of low temperature techniques, processes and equipment. A process simulator such as Aspen HYSYS, for the design, analysis, and optimization of process plants, has features that accommodate the special requirements and therefore can be used to simulate most cryogenic liquefaction and refrigeration processes. Liquefaction is the process of cooling or refrigerating a gas to a temperature below its critical temperature so that liquid can be formed at some suitable pressure which is below the critical pressure. Cryogenic processes require special attention in terms of the integration of various components like heat exchangers, Joule-Thompson Valve, Turbo expander and Compressor. Here, Aspen HYSYS, a process modeling tool, is used to understand the behavior of the complete plant. This paper presents the analysis of an air liquefaction plant based on the Linde cryogenic cycle, performed using the Aspen HYSYS process modeling tool. It covers the technique used to find the optimum values for getting the maximum liquefaction of the plant considering different constraints of other parameters. The analysis result so obtained gives clear idea in deciding various parameter values before implementation of the actual plant in the field. It also gives an idea about the productivity and profitability of the given configuration plant which leads to the design of an efficient productive plant.
A system for predicting the amount of Phellinus (Fomes) igniarius rot in trembling aspen stands
Robert L. Anderson; Arthur L. Jr. Schipper
1978-01-01
The occurrence of Phellinus (Fomes) igniarius white trunk rot in 45- to 50-year-old trembling aspen stands can be predicted by applying a constant to the stand basal area with P. igniarius conks to estimate the total basal area with P. igniarius rot. Future decay projections can be made by reapplying the basal area of hidden decay for each 6 years projected. This paper...
ERIC Educational Resources Information Center
Kleeman, Michael J.
The goals of the Fourth Annual Aspen Institute Roundtable on International Telecommunications were: to address procedural and substantive solutions for international coordination of Internet policy issues; to suggest different models for dispute resolution and policy coordination that might be employed in the various contexts of Internet usage;…
Humbird, David; Trendewicz, Anna; Braun, Robert; ...
2017-01-12
A biomass fast pyrolysis reactor model with detailed reaction kinetics and one-dimensional fluid dynamics was implemented in an equation-oriented modeling environment (Aspen Custom Modeler). Portions of this work were detailed in previous publications; further modifications have been made here to improve stability and reduce execution time of the model to make it compatible for use in large process flowsheets. The detailed reactor model was integrated into a larger process simulation in Aspen Plus and was stable for different feedstocks over a range of reactor temperatures. Sample results are presented that indicate general agreement with experimental results, but with higher gasmore » losses caused by stripping of the bio-oil by the fluidizing gas in the simulated absorber/condenser. Lastly, this integrated modeling approach can be extended to other well-defined, predictive reactor models for fast pyrolysis, catalytic fast pyrolysis, as well as other processes.« less
Boullata, Joseph I; Holcombe, Beverly; Sacks, Gordon; Gervasio, Jane; Adams, Stephen C; Christensen, Michael; Durfee, Sharon; Ayers, Phil; Marshall, Neil; Guenter, Peggi
2016-08-01
Parenteral nutrition (PN) is a high-alert medication with a complex drug use process. Key steps in the process include the review of each PN prescription followed by the preparation of the formulation. The preparation step includes compounding the PN or activating a standardized commercially available PN product. The verification and review, as well as preparation of this complex therapy, require competency that may be determined by using a standardized process for pharmacists and for pharmacy technicians involved with PN. An American Society for Parenteral and Enteral Nutrition (ASPEN) standardized model for PN order review and PN preparation competencies is proposed based on a competency framework, the ASPEN-published interdisciplinary core competencies, safe practice recommendations, and clinical guidelines, and is intended for institutions and agencies to use with their staff. © 2016 American Society for Parenteral and Enteral Nutrition.
DOE Office of Scientific and Technical Information (OSTI.GOV)
Humbird, David; Trendewicz, Anna; Braun, Robert
A biomass fast pyrolysis reactor model with detailed reaction kinetics and one-dimensional fluid dynamics was implemented in an equation-oriented modeling environment (Aspen Custom Modeler). Portions of this work were detailed in previous publications; further modifications have been made here to improve stability and reduce execution time of the model to make it compatible for use in large process flowsheets. The detailed reactor model was integrated into a larger process simulation in Aspen Plus and was stable for different feedstocks over a range of reactor temperatures. Sample results are presented that indicate general agreement with experimental results, but with higher gasmore » losses caused by stripping of the bio-oil by the fluidizing gas in the simulated absorber/condenser. Lastly, this integrated modeling approach can be extended to other well-defined, predictive reactor models for fast pyrolysis, catalytic fast pyrolysis, as well as other processes.« less
NASA Astrophysics Data System (ADS)
Miller, R. M.; Podila, G. K.
2008-12-01
An understanding of the genetic determinants of keystone symbiotic relationships is essential to elucidating adaptive mechanisms influencing higher-order processes, including shifts in community composition following environmental perturbations. The Aspen FACE project offers a unique opportunity to address adaptive processes with an imposed three way interaction experiment composed of the atmospheric pollutant ozone (eO3), elevated CO2 (eCO2) fumigations, five Populus tremuloides (aspen) genotypes, and both arbuscular mycorrhizal and ectomycorrhizal fungal interactions. The 10 year time span of this experiment has allowed for a realistic and mechanistic understanding of above ground responses of the aspen genotypes to eCO2, eO3 and the interaction effects of eCO2 and eO3. Even so, treatment influences to the below ground, including carbon allocation to roots and associated mycorrhizal symbionts, and rhizosphere dynamics are just beginning to be understood. We hypothesized that mycorrhizal fungal responses to eCO2, eO3, and the interaction effects of eCO2+eO3 are conditioned by the degree of response of their aspen hosts. We intend to describe the molecular mechanisms of an important critical interaction between host and fungus using microarray analysis of expression profiles, as well as metabolic profiling of aspen roots and their associated mycorrhizal partner, the arbuscular mycorrhizal fungus (AMF) Glomus intraradices, under eCO2, eO3 and eCO2+eO3. We present evidence that host-derived factors, expressed in response to eCO2+eO3, trigger responses in Glomus leading to the partitioning or metabolic shift in lipid biosynthesis that is associated with reduced extraradical hyphae growth and altered lipid metabolism. We then scale these lower-level responses to give better insight to fungal intraradical and extraradical allocation of biomass and fungal and root lipid and carbohydrate content in association with aspen genotype responses to the imposed treatments. By evaluating microarray data of more than 2300 genes that are regulated (out of 25,000) in aspen mycorrhizal roots, the eCO2 responsive and eO3 tolerant aspen ecotype 271 demonstrated upregulation for antioxidant genes under eCO2+eO3 conditions. We found decreased expression of both neutral and acid invertase genes indicating that the availability of carbohydrate to the fungus is reduced. We also found an increase in plant amino acid transporters under eO3 and eCO2+eO3 that partitions more nitrogen to the plant from mycorrhizal roots and triggers the fungus into an N-starvation and lipid storage mode. This observation is supported by down-regulation of genes involved in nitrogen utilization in Glomus and the enrichment of hyphal 15N content, as well as an increase in the AMF marker storage lipid (neutral fatty acid 16:1w5c)in the root. The up-regulation of pathways involved in the formation of triglycerides that can be taken up by the fungus may be a critical step for changes in Glomus lipid metabolism. Also, in support of the above findings, is the rather high expression of genes involved in iron sequestration by aspen clone 271 when exposed to both eO3 and eCO2+eO3 fumigation. Iron is needed for both fatty acid (FA) desaturases and fatty acid synthase. Under eCO2+eO3, we found down-regulation of FA desaturases in Glomus, suggesting reduced levels of iron could be a potential signal for the fungus to go into storage mode and reduced growth of extraradical hyphae into the soil.
1976-08-01
American elm Lonicera tatarica - tartarian honeysuckle Ulmus rubra - slippery elm Siiibucus canadenis - common elder Ulmus thoiii~sii - cork elm ...community borders the marshes and swamps. 2.060 The predominant species are the trembling aspen (Populus tremuloides), red maple (Acer rubrum), the elms ...succession. The most numerous trees (in descending order) are: white birch, red maple, aspen, sugar maple, black ash, basswood, elm (Ulmus sp.), hemlock
The Impact of Nitrogen Limitation and Mycorrhizal Symbiosis on Aspen Tree Growth and Development
DOE Office of Scientific and Technical Information (OSTI.GOV)
Tran, Bich Thi Ngoc
Nitrogen deficiency is the most common and widespread nutritional deficiency affecting plants worldwide. Ectromycorrhizal symbiosis involves the beneficial interaction of plants with soil fungi and plays a critical role in nutrient cycling, including the uptake of nitrogen from the environment. The main goal of this study is to understand how limiting nitrogen in the presence or absence of an ectomycorrhizal fungi, Laccaria bicolor, affects the health of aspen trees, Populus temuloides.
Derba-Maceluch, Marta; Awano, Tatsuya; Takahashi, Junko; Lucenius, Jessica; Ratke, Christine; Kontro, Inkeri; Busse-Wicher, Marta; Kosik, Ondrej; Tanaka, Ryo; Winzéll, Anders; Kallas, Åsa; Leśniewska, Joanna; Berthold, Fredrik; Immerzeel, Peter; Teeri, Tuula T; Ezcurra, Ines; Dupree, Paul; Serimaa, Ritva; Mellerowicz, Ewa J
2015-01-01
Certain xylanases from family GH10 are highly expressed during secondary wall deposition, but their function is unknown. We carried out functional analyses of the secondary-wall specific PtxtXyn10A in hybrid aspen (Populus tremula × tremuloides). PtxtXyn10A function was analysed by expression studies, overexpression in Arabidopsis protoplasts and by downregulation in aspen. PtxtXyn10A overexpression in Arabidopsis protoplasts resulted in increased xylan endotransglycosylation rather than hydrolysis. In aspen, the enzyme was found to be proteolytically processed to a 68 kDa peptide and residing in cell walls. Its downregulation resulted in a corresponding decrease in xylan endotransglycosylase activity and no change in xylanase activity. This did not alter xylan molecular weight or its branching pattern but affected the cellulose-microfibril angle in wood fibres, increased primary growth (stem elongation, leaf formation and enlargement) and reduced the tendency to form tension wood. Transcriptomes of transgenic plants showed downregulation of tension wood related genes and changes in stress-responsive genes. The data indicate that PtxtXyn10A acts as a xylan endotransglycosylase and its main function is to release tensional stresses arising during secondary wall deposition. Furthermore, they suggest that regulation of stresses in secondary walls plays a vital role in plant development. © 2014 The Authors. New Phytologist © 2014 New Phytologist Trust.
DOE Office of Scientific and Technical Information (OSTI.GOV)
Liu, Jian; Bearden, Mark D.; Fernandez, Carlos A.
Magnesium (Mg) has many useful applications especially in various Mg alloys which can decrease weight while increasing strength. To increase the affordability and minimize environment consequence, a novel catalyzed organo-metathetical (COMET) process was proposed to extract Mg from seawater aiming to achieve significant reduction in total energy and production cost comparing with the melting salt electrolysis method currently adopted by US Mg LLC. A process flowsheet for a reference COMET process was set-up using Aspen Plus which included five key steps, anhydrous MgCl2 production, transmetallation, dibutyl Mg decomposition, n-BuLi regeneration, and LiCL electrolysis. The energy and production cost and CO2more » emission were estimated based on the Aspen modeling using Aspen economic analyzer. Our results showed that it is possible to produce Mg from seawater with a production cost of $2.0/kg-Mg while consuming about 35.3 kWh/kg-Mg and releasing 7.0 kg CO2/kg-Mg. A simplified US Mg manufacturing process was also generated using Aspen and the cost and emission results were estimated for comparison purpose. Under our simulation conditions, the reference COMET process maintain a comparable CO2 emission rate and can save about 40% in production cost and save about 15% energy compared to the simplified US Mg process.« less
NASA Technical Reports Server (NTRS)
Middleton, Elizabeth M.; Sullivan, Joseph H.; Papagno, Andrea J.
2000-01-01
The role of foliar nitrogen (N) in the seasonal dynamics and vertical canopy distribution of photosynthetic pigments, photosynthetic capacity, and carbon (C) storage was investigated in boreal broadleaved species. The study was conducted at two different aged stands (60 y and 15 y) in 1994 and 1996 in Saskatchewan, Canada as part of the Boreal Ecosystem-Atmosphere Study (BOREAS). Foliage in upper and lower strata was examined for aspen (Populus tremuloides Michx.) and its associated hazelnut shrub (Corylus americana Walt.). We determined that C accumulation, expressed as dry mass per unit leaf area (mg C cm (exp -2)), was linearly dependent on N content (approximately 0.3- 3.5 mg N cm (exp -2))(r (exp 2) = 0.93, n=383, P less than 0.001) when eleven foliage groups were defined according to species, site, and developmental stage. C assembly was greatest in the upper aspen strata of both sites (seasonal average, 40.1 plus or minus 0.6 mg C cm (exp -2)), intermediate in the lower aspen strata (32.7 plus or minus 0.6), and considerably lower, and similar, in the hazelnut shrub layers (23.7 plus or minus 0.6) and in expanding aspen leaves (23.8 plus or minus 0.5); the lowest C assembly per unit N occurred in the two youngest, emerging leaf groups (17.1 plus or minus 0.6). Other relationships among physiological and biochemical variables were typically non-linear and were confounded by inclusion of the three groups of young (i.e., emerging or expanding) leaves, unless these were separately identified. Net C uptake, measured as photosynthetic capacity (A (sub max), micromole CO2 m (exp -2) s (exp -1)), was greater in aspen throughout the season, and optimal in mid-summer at a C:N ratio of approximately 18 (approximately 2.3 %N). When young leaves were excluded and logarithms of both variables were used, A (sub max) was approximately linearly dependent on N (mg N cm (exp-2) (r (exp 2) = 0.85, n= 193, P less than 0.001), attributed to incorporation of N into photosynthetic complexes and enzymes. In mature leaves, differences in pigment content vs. N among canopy strata were accentuated when N was expressed per unit leaf area (Mg cm (exp -2)) . However, the simplest log-linear relationship between a pigment variable and N was obtained for a ratio describing the relative allocation of photosynthetic pigment to Chl a (Chl a/[Chl b + carotenoids], microgram cm (exp -2)/ microgram cm-2) vs. %N (r (exp 2) = 0.90, n=343, P less than 0.001). Attainment of comparable A (sub max) Chl a content and relative Chl a allocation per unit N (mg cm (exp -2)) was achieved at different foliar N levels per canopy group: the lowest N requirement was for hazelnut leaves in the lowest, shaded stratum at the older, closed canopy site; the highest N requirement was in aspen leaves of the upper-most stratum at the younger, more open canopy site. These results highlight the differences in physiological responses between young and fully expanded leaves and show that sustaining those foliar constituents and processes important to C balance may require higher foliar N levels in leaves of establishing vs. mature aspen stands. There may be implications for remote-sensing assessments made for carbon balance in springtime, or over a landscape mosaic comprised of different aged stands.
Reinikainen, Michael; D’Amato, Anthony W.; Bradford, John B.; Fraver, Shawn
2014-01-01
Low-severity canopy disturbance presumably influences forest carbon dynamics during the course of stand development, yet the topic has received relatively little attention. This is surprising because of the frequent occurrence of such events and the potential for both the severity and frequency of disturbances to increase as a result of climate change. We investigated the impacts of low-severity canopy disturbance and average insect defoliation on forest carbon stocks and rates of carbon sequestration in mature aspen mixedwood forests of varying stand age (ranging from 61 to 85 years), overstory composition, stocking level, and site quality. Stocking level and site quality positively affected the average annual aboveground tree carbon increment (CAAI), while stocking level, site quality, and stand age positively affected tree carbon stocks (CTREE) and total ecosystem carbon stocks (CTOTAL). Cumulative canopy disturbance (DIST) was reconstructed using dendroecological methods over a 29-year period. DIST was negatively and significantly related to soil carbon (CSOIL), and it was negatively, albeit marginally, related to CTOTAL. Minima in the annual aboveground carbon increment of trees (CAI) occurred at sites during defoliation of aspen (Populus tremuloides Michx.) by forest tent caterpillar (Malacosoma disstria Hubner), and minima were more extreme at sites dominated by trembling aspen than sites mixed with conifers. At sites defoliated by forest tent caterpillar in the early 2000s, increased sequestration by the softwood component (Abies balsamea (L.) Mill. and Picea glauca (Moench) Voss) compensated for overall decreases in CAI by 17% on average. These results underscore the importance of accounting for low-severity canopy disturbance events when developing regional forest carbon models and argue for the restoration and maintenance of historically important conifer species within aspen mixedwoods to enhance stand-level resilience to disturbance agents and maintain site-level carbon stocks.
Persistent Effects of Fire Severity on Early Successional Forests in Interior Alaska
NASA Technical Reports Server (NTRS)
Shenoy, Aditi; Johnstone, Jill F.; Kasischke, Eric S.; Kielland, Knut
2011-01-01
There has been a recent increase in the frequency and extent of wildfires in interior Alaska, and this trend is predicted to continue under a warming climate. Although less well documented, corresponding increases in fire severity are expected. Previous research from boreal forests in Alaska and western Canada indicate that severe fire promotes the recruitment of deciduous tree species and decreases the relative abundance of black spruce (Picea mariana) immediately after fire. Here we extend these observations by (1) examining changes in patterns of aspen and spruce density and biomass that occurred during the first two decades of post-fire succession, and (2) comparing patterns of tree composition in relation to variations in post-fire organic layer depth in four burned black spruce forests in interior Alaska after 10-20 years of succession.Wefound that initial effects of fire severity on recruitment and establishment of aspen and black spruce were maintained by subsequent effects of organic layer depth and initial plant biomass on plant growth during post-fire succession. The proportional contribution of aspen (Populus tremuloides) to total stand biomass remained above 90% during the first and second decades of succession in severely burned sites, while in lightly burned sites the proportional contribution of aspen was reduced due to a 40- fold increase in spruce biomass in these sites. Relationships between organic layer depth and stem density and biomass were consistently negative for aspen, and positive or neutral for black spruce in all four burns. Our results suggest that initial effects of post-fire organic layer depths on deciduous recruitment are likely to translate into a prolonged phase of deciduous dominance during post-fire succession in severely burned stands. This shift in vegetation distribution has important implications for climate-albedo feedbacks, future fire regime, wildlife habitat quality and natural resources for indigenous subsistence activities in interior Alaska.
Bryophyte species richness on retention aspens recovers in time but community structure does not.
Oldén, Anna; Ovaskainen, Otso; Kotiaho, Janne S; Laaka-Lindberg, Sanna; Halme, Panu
2014-01-01
Green-tree retention is a forest management method in which some living trees are left on a logged area. The aim is to offer 'lifeboats' to support species immediately after logging and to provide microhabitats during and after forest re-establishment. Several studies have shown immediate decline in bryophyte diversity after retention logging and thus questioned the effectiveness of this method, but longer term studies are lacking. Here we studied the epiphytic bryophytes on European aspen (Populus tremula L.) retention trees along a 30-year chronosequence. We compared the bryophyte flora of 102 'retention aspens' on 14 differently aged retention sites with 102 'conservation aspens' on 14 differently aged conservation sites. We used a Bayesian community-level modelling approach to estimate the changes in bryophyte species richness, abundance (area covered) and community structure during 30 years after logging. Using the fitted model, we estimated that two years after logging both species richness and abundance of bryophytes declined, but during the following 20-30 years both recovered to the level of conservation aspens. However, logging-induced changes in bryophyte community structure did not fully recover over the same time period. Liverwort species showed some or low potential to benefit from lifeboating and high potential to re-colonise as time since logging increases. Most moss species responded similarly, but two cushion-forming mosses benefited from the logging disturbance while several weft- or mat-forming mosses declined and did not re-colonise in 20-30 years. We conclude that retention trees do not function as equally effective lifeboats for all bryophyte species but are successful in providing suitable habitats for many species in the long-term. To be most effective, retention cuts should be located adjacent to conservation sites, which may function as sources of re-colonisation and support the populations of species that require old-growth forests.
Non-invasive methods to maintain cervical spine position after pediatric tracheal resections.
Aydinyan, Kahren K; Day, Jonathan D; Troiano, Gina M; Digoy, G Paul
2017-07-01
To present our experience with two methods of neck stabilization after pediatric tracheal resection with primary anastomosis as possible alternatives to the traditional chest-chin suture. Children undergoing tracheal resection and/or cricotracheal resection with anastomosis under tension were placed in cervical spine flexion postoperatively with either a chest-chin (Grillo) suture, an Aspen cervical collar or Trulife Johnson cervical-thoracic orthosis (CTO). A retrospective chart review of tracheal resections performed between 2005 and 2016 was completed to evaluate the positive and negative factors associated with each neck flexion technique. Of the 20 patients, there were 13 patients with the Grillo suture, 4 with the Aspen collar and 3 patients with the Johnson CTO. There were 13 tracheal resection procedures and 7 cricotracheal resections, all of which had anastomosis under tension. One major anastomosis dehiscence was noted with the Grillo suture technique which required reoperation. Two patients with the Grillo suture experienced skin breakdown at the suture site. The Aspen cervical collar, which fixed the cervical spine and prevented lateral and rotational motion, was limited in several cases in that it placed the spine in slight hyperextension. The Johnson CTO provided the most support in a flexed position and prevented cervical spine motion in all directions. No anastomosis complications were noted with the Aspen collar or the Johnson CTO, however, several patients sustained minor cutaneous wounds. In this series the Aspen cervical collar and Johnson CTO were used successfully as non-Grillo alternatives to postoperative neck stabilization in pediatric tracheal resections. Modifications to both devices are proposed to minimize cutaneous injuries and increase immobilization of the cervical spine in the desired flexed position. Although these devices appear to be safe and may be better tolerated, further innovation is needed to improve the design and fit of these devices. Copyright © 2017 Elsevier B.V. All rights reserved.
Genetic Augmentation of Syringyl Lignin in Low-lignin Aspen Trees, Final Report
DOE Office of Scientific and Technical Information (OSTI.GOV)
Chung-Jui Tsai; Mark F. Davis; Vincent L. Chiang
2004-11-10
As a polysaccharide-encrusting component, lignin is critical to cell wall integrity and plant growth but also hinders recovery of cellulose fibers during the wood pulping process. To improve pulping efficiency, it is highly desirable to genetically modify lignin content and/or structure in pulpwood species to maximize pulp yields with minimal energy consumption and environmental impact. This project aimed to genetically augment the syringyl-to-guaiacyl lignin ratio in low-lignin transgenic aspen in order to produce trees with reduced lignin content, more reactive lignin structures and increased cellulose content. Transgenic aspen trees with reduced lignin content have already been achieved, prior to themore » start of this project, by antisense downregulation of a 4-coumarate:coenzyme A ligase gene (Hu et al., 1999 Nature Biotechnol 17: 808- 812). The primary objective of this study was to genetically augment syringyl lignin biosynthesis in these low-lignin trees in order to enhance lignin reactivity during chemical pulping. To accomplish this, both aspen and sweetgum genes encoding coniferaldehyde 5-hydroxylase (Osakabe et al., 1999 PNAS 96: 8955-8960) were targeted for over-expression in wildtype or low-lignin aspen under control of either a constitutive or a xylem-specific promoter. A second objective for this project was to develop reliable and cost-effective methods, such as pyrolysis Molecular Beam Mass Spectrometry and NMR, for rapid evaluation of cell wall chemical components of transgenic wood samples. With these high-throughput techniques, we observed increased syringyl-to-guaiacyl lignin ratios in the transgenic wood samples, regardless of the promoter used or gene origin. Our results confirmed that the coniferaldehyde 5-hydroxylase gene is key to syringyl lignin biosynthesis. The outcomes of this research should be readily applicable to other pulpwood species, and promise to bring direct economic and environmental benefits to the pulp and paper industry.« less
Aquatic Ecosystem Response to Timber Harvesting for the Purpose of Restoring Aspen
Jones, Bobette E.; Krupa, Monika; Tate, Kenneth W.
2013-01-01
The removal of conifers through commercial timber harvesting has been successful in restoring aspen, however many aspen stands are located near streams, and there are concerns about potential aquatic ecosystem impairment. We examined the effects of management-scale conifer removal from aspen stands located adjacent to streams on water quality, solar radiation, canopy cover, temperature, aquatic macroinvertebrates, and soil moisture. This 8-year study (2003–2010) involved two projects located in Lassen National Forest. The Pine-Bogard Project consisted of three treatments adjacent to Pine and Bogard Creeks: (i) Phase 1 in January 2004, (ii) Phase 2 in August 2005, and (iii) Phase 3 in January 2008. The Bailey Project consisted of one treatment adjacent to Bailey Creek in September 2006. Treatments involved whole tree removal using track-laying harvesters and rubber tire skidders. More than 80% of all samples analyzed for NO3-N, NH4-N, and PO4-P at Pine, Bogard, and Bailey Creeks were below the detection limit, with the exception of naturally elevated PO4-P in Bogard Creek. All nutrient concentrations (NO3-N, NH4-N, PO4-P, K, and SO4-S) showed little variation within streams and across years. Turbidity and TSS exhibited annual variation, but there was no significant increase in the difference between upstream and downstream turbidity and TSS levels. There was a significant decrease in stream canopy cover and increase in the potential fraction of solar radiation reaching the streams in response to the Pine-Bogard Phase 3 and Bailey treatments; however, there was no corresponding increase in stream temperatures. Macroinvertebrate metrics indicated healthy aquatic ecosystem conditions throughout the course of the study. Lastly, the removal of vegetation significantly increased soil moisture in treated stands relative to untreated stands. These results indicate that, with careful planning and implementation of site-specific best management practices, conifer removal to restore aspen stands can be conducted without degrading aquatic ecosystems. PMID:24376826
Ted Hogg, Edward H; Michaelian, Michael
2015-05-01
Increases in mortality of trembling aspen (Populus tremuloides Michx.) have been recorded across large areas of western North America following recent periods of exceptionally severe drought. The resultant increase in standing, dead tree biomass represents a significant potential source of carbon emissions to the atmosphere, but the timing of emissions is partially driven by dead-wood dynamics which include the fall down and breakage of dead aspen stems. The rate at which dead trees fall to the ground also strongly influences the period over which forest dieback episodes can be detected by aerial surveys or satellite remote sensing observations. Over a 12-year period (2000-2012), we monitored the annual status of 1010 aspen trees that died during and following a severe regional drought within 25 study areas across west-central Canada. Observations of stem fall down and breakage (snapping) were used to estimate woody biomass transfer from standing to downed dead wood as a function of years since tree death. For the region as a whole, we estimated that >80% of standing dead aspen biomass had fallen after 10 years. Overall, the rate of fall down was minimal during the year following stem death, but thereafter fall rates followed a negative exponential equation with k = 0.20 per year. However, there was high between-site variation in the rate of fall down (k = 0.08-0.37 per year). The analysis showed that fall down rates were positively correlated with stand age, site windiness, and the incidence of decay fungi (Phellinus tremulae (Bond.) Bond. and Boris.) and wood-boring insects. These factors are thus likely to influence the rate of carbon emissions from dead trees following periods of climate-related forest die-off episodes. © 2014 Her Majesty the Queen in Right of Canada Global Change Biology © 2014 John Wiley & Sons Ltd Reproduced with the permission of the Minister of Natural Resources Canada.
Dynamic Simulation of AN Helium Refrigerator
NASA Astrophysics Data System (ADS)
Deschildre, C.; Barraud, A.; Bonnay, P.; Briend, P.; Girard, A.; Poncet, J. M.; Roussel, P.; Sequeira, S. E.
2008-03-01
A dynamic simulation of a large scale existing refrigerator has been performed using the software Aspen Hysys®. The model comprises the typical equipments of a cryogenic system: heat exchangers, expanders, helium phase separators and cold compressors. It represents the 400 W @ 1.8 K Test Facility located at CEA—Grenoble. This paper describes the model development and shows the possibilities and limitations of the dynamic module of Aspen Hysys®. Then, comparison between simulation results and experimental data are presented; the simulation of cooldown process was also performed.
2014-01-01
benefits of enteral nutrition (EN), the American Society for Parenteral and Enteral Nutrition (ASPEN) and the Society of Critical Care Medicine (SCCM...instances where parenteral nutrition was used rather than EN or the subject consumed food orally). Because the change in clinical practice included only...therapy in the adult critically ill patient: Society of Critical Care Medicine (SCCM) and American Society for Parenteral and Enteral Nutrition (A.S.P.E.N
Darkwah, Kwabena; Nokes, Sue E; Seay, Jeffrey R; Knutson, Barbara L
2018-05-22
Process simulations of batch fermentations with in situ product separation traditionally decouple these interdependent steps by simulating a separate "steady state" continuous fermentation and separation units. In this study, an integrated batch fermentation and separation process was simulated for a model system of acetone-butanol-ethanol (ABE) fermentation with in situ gas stripping, such that the fermentation kinetics are linked in real-time to the gas stripping process. A time-dependent cell growth, substrate utilization, and product production is translated to an Aspen Plus batch reactor. This approach capitalizes on the phase equilibria calculations of Aspen Plus to predict the effect of stripping on the ABE fermentation kinetics. The product profiles of the integrated fermentation and separation are shown to be sensitive to gas flow rate, unlike separate steady state fermentation and separation simulations. This study demonstrates the importance of coupled fermentation and separation simulation approaches for the systematic analyses of unsteady state processes.
DOE Office of Scientific and Technical Information (OSTI.GOV)
Burton, Andrew J.; Zak, Donald R.; Kubiske, Mark E.
Two of the most important and pervasive greenhouse gases driving global change and impacting forests in the U.S. and around the world are atmospheric CO2 and tropospheric O3. As the only free air, large-scale manipulative experiment studying the interaction of elevated CO2 and O3 on forests, the Aspen FACE experiment was uniquely designed to address the long-term ecosystem level impacts of these two greenhouse gases on aspen-birch-maple forests, which dominate the richly forested Lake States region. The project was established in 1997 to address the overarching scientific question: “What are the effects of elevated [CO2] and [O3], alone and inmore » combination, on the structure and functioning of northern hardwood forest ecosystems?” From 1998 through the middle of the 2009 growing season, we examined the interacting effects of elevated CO2 and O3 on ecosystem processes in an aggrading northern forest ecosystem to compare the responses of early-successional, rapid-growing shade intolerant trembling aspen and paper birch to those of a late successional, slower growing shade tolerant sugar maple. Fumigations with elevated CO2 (560 ppm during daylight hours) and O3 (approximately 1.5 x ambient) were conducted during the growing season from 1998 to 2008, and in 2009 through harvest date. Response variables quantified during the experiment included growth, competitive interactions and stand dynamics, physiological processes, plant nutrient status and uptake, tissue biochemistry, litter quality and decomposition rates, hydrology, soil respiration, microbial community composition and respiration, VOC production, treatment-pest interactions, and treatment-phenology interactions. In 2009, we conducted a detailed harvest of the site. The harvest included detailed sampling of a subset of trees by component (leaves and buds, fine branches, coarse branches and stem, coarse roots, fine roots) and excavation of soil to a depth of 1 m. Throughout the experiment, aspen and birch photosynthesis increased with elevated CO2 and tended to decrease with elevated O3, compared to the control. In contrast to aspen and birch, maple photosynthesis was not enhanced by elevated CO2. Elevated O3 did not cause significant reductions in maximum photosynthesis in birch or maple. In addition, photosynthesis in ozone sensitive clones was affected to a much greater degree than that in ozone tolerant aspen clones. Treatment effects on photosynthesis contributed to CO2 stimulation of aboveground and belowground growth that was species and genotype dependent, with birch and aspen being most responsive and maple being least responsive. The positive effects of elevated CO2 on net primary productivity NPP were sustained through the end of the experiment, but negative effects of elevated O3 on NPP had dissipated during the final three years of treatments. The declining response to O3 over time resulted from the compensatory growth of O3-tolerant genotypes and species as the growth of O3-sensitive individuals declined over time. Cumulative NPP over the entire experiment was 39% greater under elevated CO2 and 10% lower under elevated O3. Enhanced NPP under elevated CO2 was sustained by greater root exploration of soil for growth-limiting N, as well as more rapid rates of litter decomposition and microbial N release during decay. Results from Aspen FACE clearly indicate that plants growing under elevated carbon dioxide, regardless of community type or ozone level, obtained significantly greater amounts of soil N. These results indicate that greater plant growth under elevated carbon dioxide has not led to “progressive N limitation”. If similar forests growing throughout northeastern North America respond in the same manner, then enhanced forest NPP under elevated CO2 may be sustained for a longer duration than previously thought, and the negative effect of elevated O3 may be diminished by compensatory growth of O3-tolerant plants as they begin to dominate forest communities. By the end of the experiment, elevated CO2 increased ecosystem C content by 11%, whereas elevated O3 decreased ecosystem C content by 9%. Total ecosystem C content in the interaction treatment (elevated CO2 and O3) did not significantly differ from that of the control. Total ecosystem C content responded similarly to the treatments across the three forest communities. The treatment effects on ecosystem C content resulted from differences in tree biomass, particularly woody tissues (branches, stem, and coarse roots), and lower C content in the near-surface mineral soil. During its duration, the Aspen FACE project involved collaboration between scientists from 9 countries, and over the course of the experiment there were over 120 Aspen FACE scientific users. These scientists helped produce 75 publications during the most recent funding period (2008-2014) and 207 peer-reviewed publications (169 in refereed journals) since the beginning of the project.« less
DOE Office of Scientific and Technical Information (OSTI.GOV)
Lindroth, R.L.; Kinney, K.K.
1998-10-01
Elevated concentrations of atmospheric CO{sub 2} are likely to interact with other factors affecting plant physiology to alter plant chemical profiles and plant-herbivore interactions. The authors evaluated the independent and interactive effects of enriched CO{sub 2} and artificial defoliation on foliar chemistry of quaking aspen (Populus tremuloides) and sugar maple (Acer saccharum), and the consequences of such changes for short-term performance of the gypsy moth (Lymantria dispar). They grew aspen and maple seedlings in ambient and enriched CO{sub 2} environments at the University of wisconsin Biotron. Seven weeks after budbreak, trees in half of the rooms were subjected to 50%more » defoliation. Afterwards, foliage was collected for chemical analyses, and feeding trials were conducted with fourth-stadium gypsy moths. Enriched CO{sub 2} altered foliar levels of water, nitrogen, carbohydrates, and phenolics, and responses generally differed between the two tree species. Defoliation induced chemical changes only in aspen. They found no significant interactions between CO{sub 2} and defoliation for levels of carbon-based defenses (phenolic glycosides and tannins). CO{sub 2} treatment altered the performance of larvae fed aspen, but not maple, whereas defoliation had little effect on performance on insects. In general, results from this experimental system do not support the hypothesis that induction of carbon-based chemical defenses, and attendant effects on insects, will be stronger in a CO{sub 2}-enriched world.« less
2012 Aspen Winter Conferences on High Energy and Astrophysics
DOE Office of Scientific and Technical Information (OSTI.GOV)
Campbell, John; Olivier, Dore; Fox, Patrick
Aspen Center for Physics Project Summary DE-SC0007313 Budget Period: 1/1/2012 to 12/31/2012 The Hunt for New Particles, from the Alps to the Plains to the Rockies The 2012 Aspen Winter Conference on Particle Physics was held at the Aspen Center for Physics from February 11 to February 17, 2012. Sixty-seven participants from nine countries, and several universities and national labs attended the workshop titled, The Hunt for New Particles, from the Alps to the Plains to the Rockies. There were 53 formal talks, and a considerable number of informal discussions held during the week. The weeks events included a publicmore » lecture-Hunting the Dark Universe given by Neal Weiner from New York University) and attended by 237 members of the public, and a physics cafe geared for high schoolers that is a discussion with physicists conducted by Spencer Chang (University of Oregon), Matthew Reece (Harvard University) and Julia Shelton (Yale University) and attended by 67 locals and visitors. While there were no published proceedings, some of the talks are posted online and can be Googled. The workshop was organized by John Campbell (Fermilab), Patrick Fox (Fermilab), Ivan Furic (University of Florida), Eva Halkiadakis (Rutgers University) and Daniel Whiteson (University of California Irvine). Additional information is available at http://indico.cern.ch/conferenceDisplay.py?confId=143360. Inflationary Theory and its Confrontation with Data in the Planck Era The 2012 Aspen Winter Conference on Astroparticle physics held at the Aspen Center for Physics was Inflationary Theory and its Confrontation with Data in the Planck Era. It was held from January 30 to February 4, 2012. The 62 participants came from 7 countries and attended 43 talks over five days. Late mornings through the afternoon are reserved for informal discussions. In feedback received from participants, it is often these unplanned chats that produce the most excitement due to working through problems with fellow physicists from other institutions and countries or due to incipient collaborations. In addition, Shamit Kachru of Stanford University gave a public lecture titled The Small (and Large) Scale Structure of Space-Time.There were 237 members of the general public in attendance. Before the lecture, 65 people attended the physics cafe to discuss the current topic with Matthew Kleban (New York University) and Chao-Lin Kuo (Stanford University). This workshop was organized by Olivier Dore (Jet Propulsion Lab), Fabian Schmidt (Caltech), Leonardo Senatore (Stanford University), and Kendrick Smith (Princeton University).« less
NASA Astrophysics Data System (ADS)
Devito, K. J.; Mendoza, C. A.; Petrone, R. M.; Landhäusser, S.; Silins, U.; Qualizza, C.; Gignac, D.
2011-12-01
The Western Boreal Plain (WBP) eco-region of western Canada is experiencing unprecedented industrial development for forest, oil and gas resources, stressing the need to assess and understand the sink and source areas of regional water flow. This requires the development of models that can be use to predict and mitigate the impacts of land use changes on water quantity and quality. The WBP is characterized by low relief, hummocky terrain with complexes of forestland and wetlands, especially peatlands. A paired aspen forest harvest experiment (HEAD2) was conducted on adjacent pond-peatland-aspen forestland complexes of a moraine landform, ubiquitous throughout the WBP, to determine the influence of aspen vegetated forestlands and harvesting relative to wetlands on local and regional water cycling. Reductions in transpiration and interception following aspen harvest resulted in some increase in soil moisture. However, excess water was largely absorbed into deep moraine substrates and resulted in groundwater recharge with little or no lateral flow to adjacent wetlands and aquatic systems. As a result of high soil and groundwater storage there was no observable difference in runoff from the harvested catchment compared to the adjacent reference (uncut) catchment confirming that in most years runoff originates from the wetland (peatland) rather than forestland hydrologic units. Furthermore, soil moisture increases were short lived due to rapid redevelopment of leaf area as result of high-density regeneration of aspen through root suckering. Recovery of transpiration and interception to near pre-harvest conditions occurred within 3 years. Climate cycles, primarily inter-annual variation in snow pack, can overwhelm the influence of aspen harvest in the WBP, as during the harvest experiment large water table rises were observed in both the uncut and the harvested forestlands. This study illustrates that forestlands on deep moraine hummocky substrates of the WBP act essentially as water sinks, with large storage and exchange to the atmosphere and infrequent runoff at a regional scale. This contrasts wetland hydrologic units that contribute most frequently to regional scale runoff due to low soil storage and persistent lateral surface or near surface runoff. Mapping the organization or configuration of these two hydrological (or cryptic) units on distinct geologic landforms rather than the topographic drainage networks appears to best represent water cycling and interactions in the WBP. Using hydrologic units will facilitate better understanding and modeling of regional runoff and aid in determining the influence of geology, climate and land use interactions in heterogeneous glacial landscapes.
NASA Astrophysics Data System (ADS)
Shenoy, A.; Kielland, K.; Johnstone, J. F.
2011-12-01
Increases in the frequency, extent, and severity of fire in the North American boreal region are projected to continue under a warming climate and are likely to be associated with changes in future vegetation composition. In interior Alaska, fire severity is linked to the relative dominance of deciduous versus coniferous canopy species. Severely burned areas have high levels of deciduous recruitment and subsequent stand dominance, while lightly burned areas exhibit black spruce self-replacement. To elucidate potential mechanisms by which differential fire severity results in differential post-fire vegetation development, we examined changes in soil nitrogen (N) supply (NO3- and NH4+) and in situ 15N uptake by young aspen (Populus tremuloides) and black spruce (Picea mariana) trees growing in lightly and severely burned areas. We hypothesized that (a) soil nitrate supply would be higher in severely burned sites and (b) since conifers have been shown to have a reduced physiological capacity for NO3- uptake, aspen would display greater rates of NO3- uptake than spruce in severely burned sites. Our results suggested that the composition and magnitude of inorganic N supply 14 years after the fire was nearly identical in high-severity and low-severity sites, and nitrate represented nearly 50% of the supply. However, both aspen and spruce took up substantially more NH4+-N than NO3- -N regardless of fire severity. Surprisingly, spruce exhibited only a moderately lower rate of NO3- uptake (μg N/g root-1h-1) than aspen. At the stand level, aspen took up nearly an order-of-magnitude more N per hectare in severely burned sites compared to lightly burned sites, while spruce exhibited the opposite pattern of N uptake with respect to fire severity. Whereas ammonium appeared to be preferred by both species, nitrate represented a larger component of N uptake (based on the NO3-:NH4+ uptake ratio) in aspen (0.7) than in spruce (0.4). We suggest that these species-specific differences in N preference coupled with their respective physiological response to fire severity represent a positive feedback loop that reinforce the opposing stand dominance patterns that have developed at the two ends of the fire severity spectrum. Shifts in forest composition from the current dominance by conifers to a future landscape dominated by deciduous forest are of concern due to impacts on climate-albedo feedbacks, forest productivity, ecosystem carbon storage, and wildlife habitat use.
Comparison of pretreatment methods on the enzymatic Saccharification of aspen wood
DOE Office of Scientific and Technical Information (OSTI.GOV)
Pinto, J.H.; Kamden, D.P.
Five different chemical pretreatments, using dilute sulfuric acid, sodium hydroxide, hydrogen peroxide and sodium hydroxide, peroxy-monosulfate, and acetic acid, were applied to aspen thermomechanical fibers. The pretreated fibers were submitted to enzymatic hydrolysis and the liberated glucose was monitored. High glucose concentrations were observed for the peroxymonosulfate and the acetic acid pretreated samples. Glucose concentrations greater than 25 g/L were obtained in these cases. This corresponds to conversions on the order of 90% of the retreated substrate glucose content. 18 refs., 1 fig., 4 tabs.
NASA Astrophysics Data System (ADS)
Soderquist, B.; Kavanagh, K.; Link, T. E.; Strand, E. K.; Seyfried, M. S.
2014-12-01
In mountainous regions across the western USA, the composition of aspen (Populus tremuloides) and sagebrush steppe plant communities is often closely related to heterogeneous soil moisture subsidies resulting from redistributed snow. With decades of climate and precipitation data across elevational and precipitation gradients, the Reynolds Creek Experimental Watershed (RCEW) and critical zone observatory (CZO) in southwest Idaho provides a unique opportunity to study the relationship between vegetation types and redistributed snow. Within the RCEW, the total amount of precipitation has remained unchanged over the past 50 years, however the percentage of the precipitation falling as snow has declined by approximately 4% per decade at mid-elevation sites. As shifts in precipitation phase continue, future trends in vegetation composition and net primary productivity (NPP) of different plant functional types remains a critical question. We hypothesize that redistribution of snow may supplement drought sensitive species like aspen more so than drought tolerant species like mountain big sagebrush (Artemisia tridentata spp. vaseyana). To assess the importance of snowdrift subsidies on sagebrush steppe vegetation, NPP of aspen, shrub, and grass species was simulated at three sites using the biogeochemical process model BIOME-BGC. Each site is located directly downslope from snowdrifts providing soil moisture inputs to aspen stands and neighboring vegetation. Drifts vary in size with the largest containing up to four times the snow water equivalent (SWE) of a uniform precipitation layer. Precipitation inputs used by BIOME-BGC were modified to represent the redistribution of snow and simulations were run using daily climate data from 1985-2013. Simulated NPP of annual grasses at each site was not responsive to subsidies from drifting snow. However, at the driest site, aspen and shrub annual NPP was increased by as much as 44 and 30%, respectively, with the redistribution of snow. These results indicate that as snow water subsidies decrease, ecosystems may shift from tree and shrub dominated to grassland dominated. As climate change progresses, shifts in the precipitation regimes in semi-arid environments may lead to changes in species composition and carbon stores throughout the intermountain west.
Contrasting drivers and trends of coniferous and deciduous tree growth in interior Alaska.
Cahoon, Sean M P; Sullivan, Patrick F; Brownlee, Annalis H; Pattison, Robert R; Andersen, Hans-Erik; Legner, Kate; Hollingsworth, Teresa N
2018-03-22
The boreal biome represents approximately one third of the world's forested area and plays an important role in global biogeochemical and energy cycles. Numerous studies in boreal Alaska have concluded that growth of black and white spruce is declining as a result of temperature-induced drought stress. The combined evidence of declining spruce growth and changes in the fire regime that favor establishment of deciduous tree species has led some investigators to suggest the region may be transitioning from dominance by spruce to dominance by deciduous forests and/or grasslands. Although spruce growth trends have been extensively investigated, few studies have evaluated long-term radial growth trends of the dominant deciduous species (Alaska paper birch and trembling aspen) and their sensitivity to moisture availability. We used a large and spatially extensive sample of tree cores from interior Alaska to compare long-term growth trends among contrasting tree species (white and black spruce vs. birch and aspen). All species showed a growth peak in the mid-1940s, although growth following the peak varied strongly across species. Following an initial decline from the peak, growth of white spruce showed little evidence of a trend, while black spruce and birch growth showed slight growth declines from ~1970 to present. Aspen growth was much more variable than the other species and showed a steep decline from ~1970 to present. Growth of birch, black and white spruce was sensitive to moisture availability throughout most of the tree-ring chronologies, as evidenced by negative correlations with air temperature and positive correlations with precipitation. However, a positive correlation between previous July precipitation and aspen growth disappeared in recent decades, corresponding with a rise in the population of the aspen leaf miner (Phyllocnistis populiella), an herbivorous moth, which may have driven growth to a level not seen since the early 20th century. Our results provide important historical context for recent growth and raise questions regarding competitive interactions among the dominant tree species and exchanges of carbon and energy in the warming climate of interior Alaska. © 2018 by the Ecological Society of America.
NASA Astrophysics Data System (ADS)
Ewers, B. E.; Mackay, D. S.; Samanta, S.; Ahl, D. E.; Burrows, S. S.; Gower, S. T.
2001-12-01
Land use changes over the last century in northern Wisconsin have resulted in a heterogeneous landscape composed of the following four main forest types: northern hardwoods, northern conifer, aspen/fir, and forested wetland. Based on sap flux measurements, aspen/fir has twice the canopy transpiration of northern hardwoods. In addition, daily transpiration was only explained by daily average vapor pressure deficit across the cover types. The objective of this study was to determine if canopy average stomatal conductance could be used to explain the species effects on tree transpiration. Our first hypothesis is that across all of the species, stomatal conductance will respond to vapor pressure deficit so as to maintain a minimum leaf water potential to prevent catostrophic cavitiation. The consequence of this hypothesis is that among species and individuals there is a proportionality between high stomatal conductance and the sensitivity of stomatal conductance to vapor pressure deficit. Our second hypothesis is that species that do not follow the proportionality deviate because the canopies are decoupled from the atmosphere. To test our two hypotheses we calculated canopy average stomatal conductance from sap flux measurements using an inversion of the Penman-Monteith equation. We estimated the canopy coupling using a leaf energy budget model that requires leaf transpiration and canopy aerodynamic conductance. We optimized the parameters of the aerodynamic conductance model using a Monte Carlo technique across six parameters. We determined the optimal model for each species by selecting parameter sets that resulted in the proportionality of our first hypothesis. We then tested the optimal energy budget models of each species by comparing leaf temperature and leaf width predicted by the models to measurements of each tree species. In red pine, sugar maple, and trembling aspen trees under high canopy coupling conditions, we found the hypothesized proportionality between high stomatal conductance and the sensitivity of stomatal conductance to vapor pressure deficit. In addition, the canopy conductance of trembling aspen was twice as high as sugar maple and the aspen trees showed much more variability.
Aspen height, stem-girth and survivorship in an area of high ungulate use
Keigley, R.B.; Frisina, M.R.
2008-01-01
An increase in ungulate population size potentially exposes aspen suckers, saplings, and trees to increased use. This study examined how stem height and girth influenced the selection of stems by ungulates for browsing, rubbing, and gnawing, and reconstructed the history of ungulate use for the study area. Transects were run through each of three aspen clones growing in southwestern Montana to determine height, circumference, and the surface area from which bark was totally and partially removed by rubbing and gnawing. Stems 20-250 cm tall were browsed. Stems 2-13 cm diameter were preferentially selected for rubbing and gnawing. The area of totally removed bark on dead saplings was twice the area of removed bark on live stems of similar diameter, suggesting that bark removal played a major role in the death of some stems. Based on an analysis of stem height and age, ungulate browsing was inferred to have increased from a light-to-moderate level to an intense level in 1991. The depth of scars was used to date scarring events. An increase in rubbing and gnawing was determined to have occurred about 1985. We concluded that elk were primarily responsible for the observed impacts. The combined effect of rubbing, gnawing, and browsing affects a broader span of ages compared to the effect of browsing alone. If prescribed fire is used to rejuvenate aspen stands, the resulting young stems should be protected from heavy browsing, rubbing and gnawing until they reach about 13 cm diameter and have grown out of the browse zone.
Wang, Wei-Qing; Cheng, Hong-Yan; Song, Song-Quan
2013-01-01
Effects of temperature, storage time and their combination on germination of aspen (Populus tomentosa) seeds were investigated. Aspen seeds were germinated at 5 to 30°C at 5°C intervals after storage for a period of time under 28°C and 75% relative humidity. The effect of temperature on aspen seed germination could not be effectively described by the thermal time (TT) model, which underestimated the germination rate at 5°C and poorly predicted the time courses of germination at 10, 20, 25 and 30°C. A modified TT model (MTT) which assumed a two-phased linear relationship between germination rate and temperature was more accurate in predicting the germination rate and percentage and had a higher likelihood of being correct than the TT model. The maximum lifetime threshold (MLT) model accurately described the effect of storage time on seed germination across all the germination temperatures. An aging thermal time (ATT) model combining both the TT and MLT models was developed to describe the effect of both temperature and storage time on seed germination. When the ATT model was applied to germination data across all the temperatures and storage times, it produced a relatively poor fit. Adjusting the ATT model to separately fit germination data at low and high temperatures in the suboptimal range increased the models accuracy for predicting seed germination. Both the MLT and ATT models indicate that germination of aspen seeds have distinct physiological responses to temperature within a suboptimal range. PMID:23658654
DOE Office of Scientific and Technical Information (OSTI.GOV)
Moore, Joel; Rabe, Karin; Nayak, Chetan
2012-05-01
Aspen Center for Physics Project Summary DOE Budget Period: 10/1/2011 to 9/30/2012 Contract # DE-SC0007479 New Paradigms for Low-Dimensional Electronic Materials The 2012 Aspen Winter Conference on Condensed Matter Physics was held at the Aspen Center for Physics from February 5 to 10, 2012. Seventy-four participants from seven countries, and several universities and national labs attended the workshop titled, New Paradigms for Low-Dimensional Electronic Materials. There were 34 formal talks, and a number of informal discussions held during the week. Talks covered a variety of topics related to DOE BES priorities, including, for example, advanced photon techniques (Hasan, Abbamonte, Orenstein,more » Shen, Ghosh) and predictive theoretical modeling of materials properties (Rappe, Pickett, Balents, Zhang, Vanderbilt); the full conference schedule is provided with this report. The week's events included a public lecture (Quantum Matters given by Chetan Nayak from Microsoft Research) and attended by 234 members of the public, and a physics caf© geared for high schoolers that is a discussion with physicists conducted by Kathryn Moler (Stanford University) and Andrew M. Rappe (University of Pennsylvania) and attended by 67 locals and visitors. While there were no published proceedings, some of the talks are posted online and can be Googled. The workshop was organized by Joel Moore (University of California Berkeley), Chetan Nayak (Microsoft Research), Karin Rabe (Rutgers University), and Matthias Troyer (ETH Zurich). Two organizers who did not attend the conference were Gabriel Aeppli (University College London & London Centre for Nanotechnology) and Andrea Cavalleri (Oxford University & Max Planck Hamburg).« less
Yamashita, Takahiro; Yamamoto-Ikemoto, Ryoko
2014-01-01
We investigated the removal of nitrogen and phosphate from the effluent of a sewage treatment plant over a long-term operation in bioreactors packed with different combinations of wood and iron, with a trickling filter packed with foam ceramics for nitrification. The average nitrification rate in the trickling filter was 0.17 kg N/m3∙day and remained at 0.11 kg N/m3∙day even when the water temperature was below 15 °C. The denitrification and phosphate removal rates in the bioreactor packed with aspen wood and iron were higher than those in the bioreactor packed with cedar chips and iron. The bioreactor packed with aspen wood and iron continued to remove nitrate and phosphate for >1200 days of operation. The nitrate removal activity of a biofilm attached to the aspen wood from the bioreactor after 784 days of operation was 0.42 g NO3-N/kg dry weight wood∙ day. There was no increase in the amount of dissolved organic matter in the outflow from the bioreactors. PMID:25247426
Hancock, Jessica E; Loya, Wendy M; Giardina, Christian P; Li, Laigeng; Chiang, Vincent L; Pregitzer, Kurt S
2007-01-01
We conducted a glasshouse mesocosm study that combined (13)C isotope techniques with wild-type and transgenic aspen (Populus tremuloides) in order to examine how altered lignin biosynthesis affects plant production and soil carbon formation. Our transgenic aspen lines expressed low stem lignin concentration but normal cellulose concentration, low lignin stem concentration with high cellulose concentration or an increased stem syringyl to guaiacyl lignin ratio. Large differences in stem lignin concentration observed across lines were not observed in leaves or fine roots. Nonetheless, low lignin lines accumulated 15-17% less root C and 33-43% less new soil C than the control line. Compared with the control line, transformed aspen expressing high syringyl lignin accumulated 30% less total plant C - a result of greatly reduced total leaf area - and 70% less new soil C. These findings suggest that altered stem lignin biosynthesis in Populus may have little effect on the chemistry of fine roots or leaves, but can still have large effects on plant growth, biomass partitioning and soil C formation.
Comparative simulation of a fluidised bed reformer using industrial process simulators
NASA Astrophysics Data System (ADS)
Bashiri, Hamed; Sotudeh-Gharebagh, Rahmat; Sarvar-Amini, Amin; Haghtalab, Ali; Mostoufi, Navid
2016-08-01
A simulation model is developed by commercial simulators in order to predict the performance of a fluidised bed reformer. As many physical and chemical phenomena take place in the reformer, two sub-models (hydrodynamic and reaction sub-models) are needed. The hydrodynamic sub-model is based on the dynamic two-phase model and the reaction sub-model is derived from the literature. In the overall model, the bed is divided into several sections. In each section, the flow of the gas is considered as plug flow through the bubble phase and perfectly mixed through the emulsion phase. Experimental data from the literature were used to validate the model. Close agreement was found between the model of both ASPEN Plus (ASPEN PLUS 2004 ©) and HYSYS (ASPEN HYSYS 2004 ©) and the experimental data using various sectioning of the reactor ranged from one to four. The experimental conversion lies between one and four sections as expected. The model proposed in this work can be used as a framework in developing the complicated models for non-ideal reactors inside of the process simulators.
Modeling Xenon Purification Systems in a Laser Inertial Fusion Engine
NASA Astrophysics Data System (ADS)
Hopkins, Ann; Gentile, Charles
2011-10-01
A Laser Inertial Fusion Engine (LIFE) is a proposed method to employ fusion energy to produce electricity for consumers. However, before it can be built and used as such, each aspect of a LIFE power plant must first be meticulously planned. We are in the process of developing and perfecting models for an exhaust processing and fuel recovery system. Such a system is especially essential because it must be able to recapture and purify expensive materials involved in the reaction so they may be reused. One such material is xenon, which is to be used as an intervention gas in the target chamber. Using Aspen HYSYS, we have modeled several subsystems for exhaust processing, including a subsystem for xenon recovery and purification. After removing hydrogen isotopes using lithium bubblers, we propose to use cryogenic distillation to purify the xenon from remaining contaminants. Aspen HYSYS allows us to analyze predicted flow rates, temperatures, pressures, and compositions within almost all areas of the xenon purification system. Through use of Aspen models, we hope to establish that we can use xenon in LIFE efficiently and in a practical manner.
2014-01-01
Background Wood cell walls are rich in cellulose, hemicellulose and lignin. Hence, they are important sources of renewable biomass for producing energy and green chemicals. However, extracting desired constituents from wood efficiently poses significant challenges because these polymers are highly cross-linked in cell walls and are not easily accessible to enzymes and chemicals. Results We show that aspen pectate lyase PL1-27, which degrades homogalacturonan and is expressed at the onset of secondary wall formation, can increase the solubility of wood matrix polysaccharides. Overexpression of this enzyme in aspen increased solubility of not only pectins but also xylans and other hemicelluloses, indicating that homogalacturonan limits the solubility of major wood cell wall components. Enzymatic saccharification of wood obtained from PL1-27-overexpressing trees gave higher yields of pentoses and hexoses than similar treatment of wood from wild-type trees, even after acid pretreatment. Conclusions Thus, the modification of pectins may constitute an important biotechnological target for improved wood processing despite their low abundance in woody biomass. PMID:24450583
Okińczyc, Piotr; Szumny, Antoni; Szperlik, Jakub; Kulma, Anna; Franiczek, Roman; Żbikowska, Beata; Krzyżanowska, Barbara; Sroka, Zbigniew
2018-05-25
In this work, we studied similarities and differences between 70% ethanol in water extract (70EE) and essential oils (EOs) obtained from propolis, black poplars ( Populus nigra L.) and aspens ( P. tremula L.) to ascertain which of these is a better indicator of the plant species used by bees to collect propolis precursors. Composition of 70EE was analyzed by UPLC-PDA-MS, while GC-MS was used to research the EOs. Principal component analyses (PCA) and calculations of Spearman's coefficient rank were used for statistical analysis. Statistical analysis exhibited correlation between chemical compositions of propolis and Populus buds' 70EE. In the case of EOs, results were less clear. Compositions of black poplars, aspens EOs and propolises have shown more variability than 70EE. Different factors such as higher instability of EOs compared to 70EE, different degradation pattern of benzyl esters to benzoic acid, differences in plant metabolism and bees' preferences may be responsible for these phenomena. Our research has therefore shown that 70EE of propolis reflected the composition of P. nigra or complex aspen⁻black poplar origin.
Experimental Design for CMIP6: Aerosol, Land Use, and Future Scenarios Final Report
DOE Office of Scientific and Technical Information (OSTI.GOV)
Arnott, James
2015-10-30
The Aspen Global Change Institute hosted a technical science workshop entitled, “Experimental design for CMIP6: Aerosol, Land Use, and Future Scenarios,” on August 3-8, 2014 in Aspen, CO. Claudia Tebaldi (NCAR) and Brian O’Neill (NCAR) served as co-chairs for the workshop. The Organizing committee also included Dave Lawrence (NCAR), Jean-Francois Lamarque (NCAR), George Hurtt (University of Maryland), & Detlef van Vuuren (PBL Netherlands Environmental Change). The meeting included the participation of 22 scientists representing many of the major climate modeling centers for a total of 110 participant days.
Double keystone bird in a keystone species complex.
Daily, G C; Ehrlich, P R; Haddad, N M
1993-01-15
Species in a Colorado subalpine ecosystem show subtle interdependences. Red-naped sapsuckers play two distinct keystone roles. They excavate nest cavities in fungus-infected aspens that are required as nest sites by two species of swallows, and they drill sap wells into willows that provide abundant nourishment for themselves, hummingbirds, orange-crowned warblers, chipmunks, and an array of other sap robbers. The swallows thus depend on, and the sap robbers benefit from, a keystone species complex comprised of sapsuckers, willows, aspens, and a heartwood fungus. Disappearance of any element of the complex could cause an unanticipated unraveling of the community.
Evolution, Three-Dimensional Model and Localization of Truncated Hemoglobin PttTrHb of Hybrid Aspen
Dumont, Estelle; Jokipii-Lukkari, Soile; Parkash, Vimal; Vuosku, Jaana; Sundström, Robin; Nymalm, Yvonne; Sutela, Suvi; Taskinen, Katariina; Kallio, Pauli T.; Salminen, Tiina A.; Häggman, Hely
2014-01-01
Thus far, research on plant hemoglobins (Hbs) has mainly concentrated on symbiotic and non-symbiotic Hbs, and information on truncated Hbs (TrHbs) is scarce. The aim of this study was to examine the origin, structure and localization of the truncated Hb (PttTrHb) of hybrid aspen (Populus tremula L. × tremuloides Michx.), the model system of tree biology. Additionally, we studied the PttTrHb expression in relation to non-symbiotic class1 Hb gene (PttHb1) using RNAi-silenced hybrid aspen lines. Both the phylogenetic analysis and the three-dimensional (3D) model of PttTrHb supported the view that plant TrHbs evolved vertically from a bacterial TrHb. The 3D model suggested that PttTrHb adopts a 2-on-2 sandwich of α-helices and has a Bacillus subtilis -like ligand-binding pocket in which E11Gln and B10Tyr form hydrogen bonds to a ligand. However, due to differences in tunnel cavity and gate residue (E7Ala), it might not show similar ligand-binding kinetics as in Bs-HbO (E7Thr). The immunolocalization showed that PttTrHb protein was present in roots, stems as well as leaves of in vitro -grown hybrid aspens. In mature organs, PttTrHb was predominantly found in the vascular bundles and specifically at the site of lateral root formation, overlapping consistently with areas of nitric oxide (NO) production in plants. Furthermore, the NO donor sodium nitroprusside treatment increased the amount of PttTrHb in stems. The observed PttTrHb localization suggests that PttTrHb plays a role in the NO metabolism. PMID:24520401
Cederholm, Tommy; Jensen, Gordon L
2017-02-01
During the ESPEN Congress in Copenhagen, Denmark (September 2016) representatives of the 4 largest global PEN-societies from Europe (ESPEN), USA (ASPEN), Asia (PENSA) and Latin America (FELANPE), and from national PEN-societies around the world met to continue the conversation on how to diagnose malnutrition that started during the Clinical Nutrition Week, Austin, USA (February 2016). Current thinking on diagnostic approaches was shared; ESPEN suggested a grading approach that could encompass various types of signs, symptoms and etiologies to support diagnosis. ASPEN emphasized where the parties agree; i.e. that the three major published approaches (ESPEN, ASPEN/AND and Subjective Global Assessment (SGA)) all propose weight loss as a key indicator for malnutrition. FELANPE suggested that the anticipated consensus approach needs to prioritize a diagnostic methodology that is available for everybody since resources differ globally. PENSA highlighted that BMI varies by ethnicity/race, and that sarcopenia/muscle mass evaluation is important for the diagnosis of malnutrition. A Core Working Committee of the Global Leadership Initiative on Malnutrition (GLIM) has been established (comprised of two representatives each from the 4 largest PEN-societies) that will lead consensus development in collaboration with a larger Working Group with broad global representation, using e-mail, telephone conferences, and face-to-face meetings during the up-coming ASPEN and ESPEN Congresses. Transparency and external input will be sought. Objectives include: 1. Consensus development around evidence-based criteria for broad application. 2. Promotion of global dissemination of the consensus criteria. 3. Seeking adoption by the World Health Organization (WHO) and the International Classification of Diseases (ICD). Copyright © 2017 American Society for Parenteral and Enteral Nutrition, Elsevier Ltd, European Society for Clinical Nutrition and Metabolism. Published by Elsevier Ltd.. All rights reserved.
Steele, Sarah J.; Gower, Stith T.; Vogel, Jason G.; Norman, John M.
1997-01-01
Root biomass, net primary production and turnover were studied in aspen, jack pine and black spruce forests in two contrasting climates. The climate of the Southern Study Area (SSA) near Prince Albert, Saskatchewan is warmer and drier in the summer and milder in the winter than the Northern Study Area (NSA) near Thompson, Manitoba, Canada. Ingrowth soil cores and minirhizotrons were used to quantify fine root net primary production (NPPFR). Average daily fine root growth (m m(-2) day(-1)) was positively correlated with soil temperature at 10-cm depth (r(2) = 0.83-0.93) for all three species, with black spruce showing the strongest temperature effect. At both study areas, fine root biomass (measured from soil cores) and fine root length (measured from minirhizotrons) were less for jack pine than for the other two species. Except for the aspen stands, estimates of NPPFR from minirhizotrons were significantly greater than estimates from ingrowth cores. The core method underestimated NPPFR because it does not account for simultaneous fine root growth and mortality. Minirhizotron NPPFR estimates ranged from 59 g m(-2) year(-1) for aspen stands at SSA to 235 g m(-2) year(-1) for black spruce at NSA. The ratio of NPPFR to total detritus production (aboveground litterfall + NPPFR) was greater for evergreen forests than for deciduous forests, suggesting that carbon allocation patterns differ between boreal evergreen and deciduous forests. In all stands, NPPFR consistently exceeded annual fine root turnover and the differences were larger for stands in the NSA than for stands in the SSA, whereas the difference between study areas was only significant for black spruce. The imbalance between NPPFR and fine root turnover is sufficient to explain the net accumulation of carbon in boreal forest soils.
Following isotopes in pulse-chase enriched aspen seedlings
NASA Astrophysics Data System (ADS)
Norris, C. E.; Wasylishen, R. E.; Landhäusser, S.; Quideau, S. A.
2011-12-01
One method to quantitatively trace biogeochemical fluxes through ecosystems, such as organic matter decomposition, is to use plant material enriched with stable isotopes. However, as plant macromolecules are known to vary in their rate of formation and decomposition, both the enrichment levels and the location of enrichment within the plant material should be characterized prior to decomposition and tracing studies. Aspen (Populus tremuloides Michx.) is a common tree species with a diverse organic matter chemical structure found in the western Canadian boreal forest. This study used a multi pulse and multi chase enrichment of stable isotopes (15N and 13C) on aspen seedlings to determine the seedling enrichment, isotope movement among plant tissues and translocation of isotopes within plant macromolecules e.g., carbohydrates and lignin. As expected, all tissues experienced increased enrichment with multiple pulses. An initial enrichment with 13C was observed in the leaves followed by translocation to the stems and roots while the 15N moved upward from the roots to leaves. The macromolecular chemistry of the organic carbon was further characterized using 13C solid state nuclear magnetic resonance spectroscopy. After the initial two hour chase period enrichment of the O-alkyl type (carbohydrate) carbon within the leaves was identified, followed by redistribution to more complex carbon compounds after the one week chase period. Root and stem tissues did not show the same pattern. Rather, changes in 13C enrichment were observed in shifting ethyl and methyl alkyl (lipid) carbon peak intensities for the stem samples while roots did not preferentially allocate 13C to a specific macromolecule. These results confirm that stable isotope enrichment of plants was non-uniform across macromolecules and tissue types. Enrichment of aspen seedlings was therefore dependant on the pulse-chase sequence used.
Detection of Aspens Using High Resolution Aerial Laser Scanning Data and Digital Aerial Images
Säynäjoki, Raita; Packalén, Petteri; Maltamo, Matti; Vehmas, Mikko; Eerikäinen, Kalle
2008-01-01
The aim was to use high resolution Aerial Laser Scanning (ALS) data and aerial images to detect European aspen (Populus tremula L.) from among other deciduous trees. The field data consisted of 14 sample plots of 30 m × 30 m size located in the Koli National Park in the North Karelia, Eastern Finland. A Canopy Height Model (CHM) was interpolated from the ALS data with a pulse density of 3.86/m2, low-pass filtered using Height-Based Filtering (HBF) and binarized to create the mask needed to separate the ground pixels from the canopy pixels within individual areas. Watershed segmentation was applied to the low-pass filtered CHM in order to create preliminary canopy segments, from which the non-canopy elements were extracted to obtain the final canopy segmentation, i.e. the ground mask was analysed against the canopy mask. A manual classification of aerial images was employed to separate the canopy segments of deciduous trees from those of coniferous trees. Finally, linear discriminant analysis was applied to the correctly classified canopy segments of deciduous trees to classify them into segments belonging to aspen and those belonging to other deciduous trees. The independent variables used in the classification were obtained from the first pulse ALS point data. The accuracy of discrimination between aspen and other deciduous trees was 78.6%. The independent variables in the classification function were the proportion of vegetation hits, the standard deviation of in pulse heights, accumulated intensity at the 90th percentile and the proportion of laser points reflected at the 60th height percentile. The accuracy of classification corresponded to the validation results of earlier ALS-based studies on the classification of individual deciduous trees to tree species. PMID:27873799
To Create a Consensus on Malnutrition Diagnostic Criteria.
Cederholm, Tommy; Jensen, Gordon L
2017-03-01
During the European Society for Clinical Nutrition and Metabolism (ESPEN) Congress in Copenhagen, Denmark (September 2016), representatives of the 4 largest global parenteral and enteral nutrition (PEN) societies from Europe (ESPEN), the United States (American Society for Parenteral and Enteral Nutrition [ASPEN]), Asia (Parenteral and Enteral Nutrition Society of Asia [PENSA]), and Latin America (Latin American Federation of Parenteral and Enteral Nutrition [FELANPE]) and from national PEN societies around the world met to continue the conversation on how to diagnose malnutrition that started during the Clinical Nutrition Week, Austin, Texas (February 2016). Current thinking on diagnostic approaches was shared; ESPEN suggested a grading approach that could encompass various types of signs, symptoms, and etiologies to support diagnosis. ASPEN emphasized where the parties agree; that is, that the 3 major published approaches (ESPEN, ASPEN-Academy of Nutrition and Dietetics, and Subjective Global Assessment [SGA]) all propose weight loss as a key indicator for malnutrition. FELANPE suggested that the anticipated consensus approach needs to prioritize a diagnostic method that is available for everybody since resources differ globally. PENSA highlighted that body mass index varies by ethnicity/race and that sarcopenia/muscle mass evaluation is important for the diagnosis of malnutrition. A Core Working Committee of the Global Leadership Initiative on Malnutrition has been established (comprising 2 representatives each from the 4 largest PEN societies) that will lead consensus development in collaboration with a larger working group with broad global representation, using e-mail, telephone conferences, and face-to-face meetings during the upcoming ASPEN and ESPEN congresses. Transparency and external input will be sought. Objectives include (1) consensus development around evidence-based criteria for broad application, (2) promotion of global dissemination of the consensus criteria, and (3) seeking adoption by the World Health Organization and the International Classification of Diseases.
DOE Office of Scientific and Technical Information (OSTI.GOV)
Anastasia M. Gribik; Ronald E. Mizia; Harry Gatley
This project addresses both the technical and economic feasibility of replacing industrial gas in lime kilns with synthesis gas from the gasification of hog fuel. The technical assessment includes a materials evaluation, processing equipment needs, and suitability of the heat content of the synthesis gas as a replacement for industrial gas. The economic assessment includes estimations for capital, construction, operating, maintenance, and management costs for the reference plant. To perform these assessments, detailed models of the gasification and lime kiln processes were developed using Aspen Plus. The material and energy balance outputs from the Aspen Plus model were used asmore » inputs to both the material and economic evaluations.« less
DOE Office of Scientific and Technical Information (OSTI.GOV)
This is an Aspen Plus process model for in situ and ex situ upgrading of fast pyrolysis vapors for the conversion of biomass to hydrocarbon fuels. It is based on conceptual designs that allow projections of future commercial implementations of the technologies based on a combination of research and existing commercial technologies. The process model was developed from the ground up at NREL. Results from the model are documented in a detailed design report NREL/TP-5100-62455 (available at http://www.nrel.gov/docs/fy15osti/62455.pdf).
Bell, David M.; Bradford, John B.; Lauenroth, William K.
2015-01-01
By examining variation in disease prevalence, mortality of healthy trees, and mortality of diseased trees, we showed that the role of disease in aspen tree mortality depended on the scale of inference. For variation among individuals in diameter, disease tended to expose intermediate-size trees experiencing moderate risk to greater risk. For spatial variation in summer temperature, disease exposed lower risk populations to greater mortality probabilities, but the magnitude of this exposure depended on summer precipitation. Furthermore, the importance of diameter and slenderness in mediating responses to climate supports the increasing emphasis on trait variation in studies of ecological responses to global change.
DOE Office of Scientific and Technical Information (OSTI.GOV)
Heil, T.P.; Lindsay, R.C.
1990-08-01
Extracts from Large Toothed Aspen (Populous grandidenta Michx.) and Jack Pine (Pinus banksiana Lamb.) contained many odor compounds found in flavor-tainted walleye from the Wisconsin River, including alkylphenols. Aspen wood contained 8 ppb of 2-isopropylphenol, and river sediments also contained low ppb levels of many alkylphenols, including 2-isopropylphenol. Thiophenol and thiocresol which sporadically cause offensive sulfury taints in Wisconsin River fish were also found in river sediment. Quantitative analysis of fish for alkylphenols supported a hypothesis involving a food chain-mediated seasonal fluctuation of alkylphenol levels in fish. Thiophenols are postulated to be derived from pulp mill activities on the river.
Using a Portfolio of Algorithms for Planning and Scheduling
NASA Technical Reports Server (NTRS)
Sherwood, Robert; Knight, Russell; Rabideau, Gregg; Chien, Steve; Tran, Daniel; Engelhardt, Barbara
2003-01-01
The Automated Scheduling and Planning Environment (ASPEN) software system, aspects of which have been reported in several previous NASA Tech Briefs articles, includes a subsystem that utilizes a portfolio of heuristic algorithms that work synergistically to solve problems. The nature of the synergy of the specific algorithms is that their likelihoods of success are negatively correlated: that is, when a combination of them is used to solve a problem, the probability that at least one of them will succeed is greater than the sum of probabilities of success of the individual algorithms operating independently of each other. In ASPEN, the portfolio of algorithms is used in a planning process of the iterative repair type, in which conflicts are detected and addressed one at a time until either no conflicts exist or a user-defined time limit has been exceeded. At each choice point (e.g., selection of conflict; selection of method of resolution of conflict; or choice of move, addition, or deletion) ASPEN makes a stochastic choice of a combination of algorithms from the portfolio. This approach makes it possible for the search to escape from looping and from solutions that are locally but not globally optimum.
Cai, Jing; Tyree, Melvin T
2010-07-01
The objective of this study was to quantify the relationship between vulnerability to cavitation and vessel diameter within a species. We measured vulnerability curves (VCs: percentage loss hydraulic conductivity versus tension) in aspen stems and measured vessel-size distributions. Measurements were done on seed-grown, 4-month-old aspen (Populus tremuloides Michx) grown in a greenhouse. VCs of stem segments were measured using a centrifuge technique and by a staining technique that allowed a VC to be constructed based on vessel diameter size-classes (D). Vessel-based VCs were also fitted to Weibull cumulative distribution functions (CDF), which provided best-fit values of Weibull CDF constants (c and b) and P(50) = the tension causing 50% loss of hydraulic conductivity. We show that P(50) = 6.166D(-0.3134) (R(2) = 0.995) and that b and 1/c are both linear functions of D with R(2) > 0.95. The results are discussed in terms of models of VCs based on vessel D size-classes and in terms of concepts such as the 'pit area hypothesis' and vessel pathway redundancy.
Cryopreservation of dormant in vivo-buds of hybrid aspen: timing as a critical factor.
Aronen, T; Ryynanen, L
2014-01-01
For the conservation of hybrid aspen germplasm, cryostorage of dormant in vivo buds is a convenient back-up method for field collections. In practice in Finland, bud collection is performed from February to March. The aim of this study was to assess how this time schedule can be extended without compromising regeneration. In addition, an easily measurable marker for successful cryopreservation was examined. Timing of cryopreservation was tested from August to February, using dormant buds from both outdoor and indoor plants. To find a marker, water content and gene expression of hydrid aspens, as well as environmental factors such as temperature, temperature sum, and light period were followed. Cryopreservation was successful from October to February, when, on an average, at least 75 % of the buds regenerated through micropropagation, and there was no difference to non-frozen controls. Significant genotypic variation was observed in October and February, with regeneration rates of 61-100 % and 37-98 %, respectively. No marker for successful cryopreservation was found among the studied factors. The results provide flexibility for the undertaking of practical work, with a recommendation that cryopreservation can be carried out from November to January - earlier than the current practice.
Mechanical Characterization of Composites and Foams for Aerospace Applications
NASA Technical Reports Server (NTRS)
Veazie, D. R.; Glinsey, C.; Webb, M. M.; Norman, M.; Meador, Michael A. (Technical Monitor)
2000-01-01
Experimental studies to investigate the mechanical properties of ultra-lightweight polyimide foams for space applications, compression after impact (CAI) properties for low velocity impact of sandwich composites, and aspen fiber/polypropylene composites containing an interface adhesive additive, Maleic Anhydride Grafted Polypropylene (MAPP), were performed at Clark Atlanta University. Tensile, compression, flexural, and shear modulus tests were performed on TEEK foams categorized by their densities and relative cost according to ASTM specifications. Results showed that the mechanical properties of the foams increased as a function of higher price and increasing density. The CAI properties of Nomex/phenolic honeycomb core, fiberglass/epoxy facesheet sandwich composites for two damage arrangements were compared using different levels of impact energy ranging from 0 - 452 Joules. Impact on the thin side showed slightly more retention of CAI strength at low impact levels, whereas higher residual compressive strength was observed from impact on the thick side at higher impact levels. The aspen fiber/polypropylene composites studied are composed of various percentages (by weight) of aspen fiber and polypropylene ranging from 30%-60% and 40%-100%, respectively. Results showed that the MAPP increases tensile and flexural strength, while having no significant influence on tensile and flexural modulus.
NASA Astrophysics Data System (ADS)
Welp, L. R.; Liu, H.; Randerson, J. T.
2005-12-01
Recent increases in growing season length at high northern latitudes may allow for greater rates of photosynthesis and carbon accumulation during spring and summer. However, warmer air and soil temperatures may also stimulate higher rates of respiration in boreal and arctic ecosystems. The net effect of these different processes on biome-level carbon fluxes remains challenging to predict. We measured carbon fluxes in three interior Alaskan stands for three years (2002-2004) to test the hypothesis that NEE in early and mid successional ecosystems is more sensitive to climate variability than NEE of older, mature ecosystems. The stands represented a chronosequence of recovery after fire with burn events in 1920 (black spruce and moss), 1987 (aspen and willow) and 1999 (grasses and deciduous shrubs). In 2002, the region experienced a cool, moderately wet spring and wet summer. In contrast, 2003 had a warm, dry spring and dry summer. In 2004, the spring was the warmest and wettest of all three years and a severe summer drought followed. Spring air temperature increased during each year of the study with April-May means of 6.4°C in 2002, 7.7°C in 2003 and 9.9°C in 2004. In each stand, warmer spring temperatures increased spring GPP, as has been reported for other northern forests, and also led to increased carbon uptake at the aspen stand with May-June NEE values of -72, -106 and -138 g C m-2. In contrast, May-June NEE at the black spruce stand increased from -94 g C m-2 in 2002 to -110 g C m-2 in 2003, but returned to -96 g C m-2 in 2004 as Re increased in response to warmer soil temperatures during that year. Sensitivity of spring NEE to climate variability was greatest at the intermediate aged aspen stand. Using a simple atmospheric model, we also found that the amplitude of the seasonal cycle of atmospheric CO2 was more sensitive to climate variability when forced with fluxes from the aspen stand than from the black spruce stand. During each year, we observed mid-summer depressions of carbon uptake rates at the black spruce stand. Despite the 2004 summer drought, Jul-Aug GPP and Re both increased in 2004, resulting in Jul-Aug NEE values comparable to 2003 (-57 and -59 g C m-2 for the black spruce in 2003 and 2004 respectively and -124 and -120 g C m-2 for the aspen), both of which were less than the wet cool summer of 2002 (-76 g C m-2 for the black spruce and -156 g C m-2 for the aspen). Warm springs and cool summers tended to promote carbon uptake in these ecosystems.
Martin, Thomas E.
2015-01-01
Synthesis and applications. Management should target species that specialize in resource selection on a declining resource. Species with greater resource selection generalization can reduce population impacts of environmental change. Resource generalization can allow a species like the wren to take advantage of habitat refuges, such as those provided by the elk exclosures. Yet, resource generalization cannot offset the negative impacts of broad-scale declines in habitat quality on the landscape, as demonstrated by the general decline of wrens. Ultimately, aspen is an important habitat for biodiversity, and land management programmes that protect and aid recovery of aspen habitats may be critical.
Huang, Jian-Guo; Stadt, Kenneth J; Dawson, Andria; Comeau, Philip G
2013-01-01
We examined the effect of competition on stem growth of Picea glauca and Populus tremuloides in boreal mixedwood stands during the stem exclusion stage. We combined traditional approaches of collecting competition data with dendrochronology to provide retrospective measurements of stem diameter growth. Several competition indices including stand basal area (BA), the sum of stem diameter at breast height (SDBH), and density (N) for the broadleaf and coniferous species, as well as similar indices considering only trees with diameters greater than each subject (BAGR, SDBHGR, and NGR), were evaluated. We used a nonlinear mixed model to characterize the basal area increment over the past 5, 10, 15, 20, 25, 30, and 35 years as a function of growth of nearby dominant trees, the size of the subject trees, deciduous and coniferous competition indices, and ecoregions. SDBHGR and BAGR were better predictors for spruce, and SDBHGR and NGR were better for aspen, respectively, than other indices. Results showed strongest correlations with long-term stem growth, as the best models integrated growth for 10-25 years for aspen and ≥ 25 for spruce. Our model demonstrated a remarkable capability (adjusted R(2)>0.67) to represent this complex variation in growth as a function of site, size and competition.
Multi-omics approach identifies molecular mechanisms of plant-fungus mycorrhizal interaction
Larsen, Peter E.; Sreedasyam, Avinash; Trivedi, Geetika; ...
2016-01-19
In mycorrhizal symbiosis, plant roots form close, mutually beneficial interactions with soil fungi. Before this mycorrhizal interaction can be established however, plant roots must be capable of detecting potential beneficial fungal partners and initiating the gene expression patterns necessary to begin symbiosis. To predict a plant root – mycorrhizal fungi sensor systems, we analyzed in vitro experiments of Populus tremuloides (aspen tree) and Laccaria bicolor (mycorrhizal fungi) interaction and leveraged over 200 previously published transcriptomic experimental data sets, 159 experimentally validated plant transcription factor binding motifs, and more than 120-thousand experimentally validated protein-protein interactions to generate models of pre-mycorrhizal sensormore » systems in aspen root. These sensor mechanisms link extracellular signaling molecules with gene regulation through a network comprised of membrane receptors, signal cascade proteins, transcription factors, and transcription factor biding DNA motifs. Modeling predicted four pre-mycorrhizal sensor complexes in aspen that interact with fifteen transcription factors to regulate the expression of 1184 genes in response to extracellular signals synthesized by Laccaria. Predicted extracellular signaling molecules include common signaling molecules such as phenylpropanoids, salicylate, and, jasmonic acid. Lastly, this multi-omic computational modeling approach for predicting the complex sensory networks yielded specific, testable biological hypotheses for mycorrhizal interaction signaling compounds, sensor complexes, and mechanisms of gene regulation.« less
Multi-omics approach identifies molecular mechanisms of plant-fungus mycorrhizal interaction
DOE Office of Scientific and Technical Information (OSTI.GOV)
Larsen, Peter E.; Sreedasyam, Avinash; Trivedi, Geetika
In mycorrhizal symbiosis, plant roots form close, mutually beneficial interactions with soil fungi. Before this mycorrhizal interaction can be established however, plant roots must be capable of detecting potential beneficial fungal partners and initiating the gene expression patterns necessary to begin symbiosis. To predict a plant root – mycorrhizal fungi sensor systems, we analyzed in vitro experiments of Populus tremuloides (aspen tree) and Laccaria bicolor (mycorrhizal fungi) interaction and leveraged over 200 previously published transcriptomic experimental data sets, 159 experimentally validated plant transcription factor binding motifs, and more than 120-thousand experimentally validated protein-protein interactions to generate models of pre-mycorrhizal sensormore » systems in aspen root. These sensor mechanisms link extracellular signaling molecules with gene regulation through a network comprised of membrane receptors, signal cascade proteins, transcription factors, and transcription factor biding DNA motifs. Modeling predicted four pre-mycorrhizal sensor complexes in aspen that interact with fifteen transcription factors to regulate the expression of 1184 genes in response to extracellular signals synthesized by Laccaria. Predicted extracellular signaling molecules include common signaling molecules such as phenylpropanoids, salicylate, and, jasmonic acid. Lastly, this multi-omic computational modeling approach for predicting the complex sensory networks yielded specific, testable biological hypotheses for mycorrhizal interaction signaling compounds, sensor complexes, and mechanisms of gene regulation.« less
Huang, Jian-Guo; Stadt, Kenneth J.; Dawson, Andria; Comeau, Philip G.
2013-01-01
We examined the effect of competition on stem growth of Picea glauca and Populus tremuloides in boreal mixedwood stands during the stem exclusion stage. We combined traditional approaches of collecting competition data with dendrochronology to provide retrospective measurements of stem diameter growth. Several competition indices including stand basal area (BA), the sum of stem diameter at breast height (SDBH), and density (N) for the broadleaf and coniferous species, as well as similar indices considering only trees with diameters greater than each subject (BAGR, SDBHGR, and NGR), were evaluated. We used a nonlinear mixed model to characterize the basal area increment over the past 5, 10, 15, 20, 25, 30, and 35 years as a function of growth of nearby dominant trees, the size of the subject trees, deciduous and coniferous competition indices, and ecoregions. SDBHGR and BAGR were better predictors for spruce, and SDBHGR and NGR were better for aspen, respectively, than other indices. Results showed strongest correlations with long-term stem growth, as the best models integrated growth for 10–25 years for aspen and ≥25 for spruce. Our model demonstrated a remarkable capability (adjusted R2>0.67) to represent this complex variation in growth as a function of site, size and competition. PMID:24204891
... Solution Center NOVEL Project Parenteral Nutrition Resources Intravenous Lipid Emulsions (ILE) Video Series SmartPN Practice and Research Toolkits Publications & ASPEN Journals Journal of Parenteral and ...
Michaelian, Michael; Hogg, Edward H; Hall, Ronald J; Arsenault, Eric
2011-01-01
Drought-induced, regional-scale dieback of forests has emerged as a global concern that is expected to escalate under model projections of climate change. Since 2000, drought of unusual severity, extent, and duration has affected large areas of western North America, leading to regional-scale dieback of forests in the southwestern US. We report on drought impacts on forests in a region farther north, encompassing the transition between boreal forest and prairie in western Canada. A central question is the significance of drought as an agent of large-scale tree mortality and its potential future impact on carbon cycling in this cold region. We used a combination of plot-based, meteorological, and remote sensing measures to map and quantify aboveground, dead biomass of trembling aspen (Populus tremuloides Michx.) across an 11.5 Mha survey area where drought was exceptionally severe during 2001–2002. Within this area, a satellite-based land cover map showed that aspen-dominated broadleaf forests occupied 2.3 Mha. Aerial surveys revealed extensive patches of severe mortality (>55%) resembling the impacts of fire. Dead aboveground biomass was estimated at 45 Mt, representing 20% of the total aboveground biomass, based on a spatial interpolation of plot-based measurements. Spatial variation in percentage dead biomass showed a moderately strong correlation with drought severity. In the prairie-like, southern half of the study area where the drought was most severe, 35% of aspen biomass was dead, compared with an estimated 7% dead biomass in the absence of drought. Drought led to an estimated 29 Mt increase in dead biomass across the survey area, corresponding to 14 Mt of potential future carbon emissions following decomposition. Many recent, comparable episodes of drought-induced forest dieback have been reported from around the world, which points to an emerging need for multiscale monitoring approaches to quantify drought effects on woody biomass and carbon cycling across large areas.
DOE Office of Scientific and Technical Information (OSTI.GOV)
Kinney, K.K.; Lindroth, R.L.; Jung, S.M.
1997-01-01
Increasing concentrations of atmospheric CO{sub 2} will interact with other environmental factors to influence the physiology and ecology of trees. This research evaluated how plant phytochemical responses to enriched atmospheric CO{sub 2} are affected by the availability of soil nitrate (NO{sub 3}{sup -}) and how these chemical changes alter performance of a tree-feeding folivore. Seedlings of three deciduous tree species were grown in ambient or elevated CO{sub 2} in combination with low or high soil NO{sub 3}{sup -} availability. Lymantria dispar larvae were reared on foliage (aspen and maple). Concentrations of nitrogen and soluble protein decreased, whereas concentrations of starch,more » condensed tannins, and ellagitannins increased, in response to elevated CO{sub 2} and/or low NO{sub 3}{sup -}. Responses of simple carbohydrates and phenolic glycosides were variable absolute (net) changes in foliar C:N ratios were greatest for aspen and least for oak, whereas relative changes were greatest for maple and least for aspen. Elevated CO{sub 2} treatments had little effect on gypsy moth development time, growth rate, or larval mass. Larvae reared on aspen foliage grown under elevated CO{sub 2} exhibited increased consumption but decreased conversion efficiencies. Gypsy moth responses to NO{sub 3}{sup -} were strongly host specific. The magnitude of insect response elicited by resource-mediated shifts in host chemistry will depend on how levels of compounds with specific importance to insect fitness are affected. Relatively few true interactions occured between carbon and nitrogen availability and insect performance. Tree species frequently interacted with CO{sub 2} and/or NO{sub 3}{sup -} availability to affect both parameters. The effects of elevated atmospheric CO{sub 2} on terrestrial plant communities will depend on species composition and soil nutrient availability. 54 refs., 9 figs., 4 tabs.« less
Community assembly in epiphytic lichens in early stages of colonization.
Gjerde, Ivar; Blom, Hans H; Lindblom, Louise; Saetersdal, Magne; Schei, Fride Høstad
2012-04-01
Colonization studies may function as natural experiments and have the potential of addressing important questions about community assembly. We studied colonization for a guild of epiphytic lichens in a former treeless heathland area of 170 km2 in southwest Norway. We investigated if epiphytic lichen species richness and composition on aspen (Populus tremula) trees corresponded to a random draw of lichen individuals from the regional species pool. We compared lichen communities of isolated young (55-120 yr) and old (140-200 yr) forest patches in the heathland area to those of aspen forest in an adjacent reference area that has been forested for a long time. All thalli (lichen bodies) of 32 selected lichen species on trunks of aspen were recorded in 35 aspen sites. When data for each site category (young, old, and reference) were pooled, we found the species richness by rarefaction to be similar for reference sites and old sites, but significantly lower for young sites. The depauperated species richness of young sites was accompanied by a skew in species composition and absence of several species that were common in the reference sites. In contrast, genetic variation screened with neutral microsatellite markers in the lichen species Lobaria pulmonaria showed no significant differences between site categories. Our null hypothesis of a neutral species assembly in young sites corresponding to a random draw from the regional species pool was rejected, whereas an alternative hypothesis based on differences in colonization capacity among species was supported. The results indicate that for the habitat configuration in the heathland area (isolated patches constituting < 0.4% of the area) lichen communities may need a colonization time of 100-150 yr for species richness to level off, but given enough time, isolation will not affect species richness. We suggest that this contradiction to expectations from classical island equilibrium theory results from low extinction rates.
Fréchette, Emmanuelle; Ensminger, Ingo; Bergeron, Yves; Gessler, Arthur; Berninger, Frank
2011-11-01
Future climate will alter the soil cover of mosses and snow depths in the boreal forests of eastern Canada. In field manipulation experiments, we assessed the effects of varying moss and snow depths on the physiology of black spruce (Picea -mariana (Mill.) B.S.P.) and trembling aspen (Populus tremuloides Michx.) in the boreal black spruce forest of western Québec. For 1 year, naturally regenerated 10-year-old spruce and aspen were grown with one of the following treatments: additional N fertilization, addition of sphagnum moss cover, removal of mosses, delayed soil thawing through snow and hay addition, or accelerated soil thawing through springtime snow removal. Treatments that involved the addition of insulating moss or snow in the spring caused lower soil temperature, while removing moss and snow in the spring caused elevated soil temperature and thus had a warming effect. Soil warming treatments were associated with greater temperature variability. Additional soil cover, whether moss or snow, increased the rate of photosynthetic recovery in the spring. Moss and snow removal, on the other hand, had the opposite effect and lowered photosynthetic activity, especially in spruce. Maximal electron transport rate (ETR(max)) was, for spruce, 39.5% lower after moss removal than with moss addition, and 16.3% lower with accelerated thawing than with delayed thawing. Impaired photosynthetic recovery in the absence of insulating moss or snow covers was associated with lower foliar N concentrations. Both species were affected in that way, but trembling aspen generally reacted less strongly to all treatments. Our results indicate that a clear negative response of black spruce to changes in root-zone temperature should be anticipated in a future climate. Reduced moss cover and snow depth could adversely affect the photosynthetic capacities of black spruce, while having only minor effects on trembling aspen.
Decker, Vicki H G; Bandau, Franziska; Gundale, Michael J; Cole, Christopher T; Albrectsen, Benedicte R
2017-02-01
Condensed tannin (CT) contents of European aspen (Populus tremula L.) vary among genotypes, and increases in nitrogen (N) availability generally reduce plants' tannin production in favor of growth, through poorly understood mechanisms. We hypothesized that intrinsic tannin production rates may co-vary with gene expression responses to soil N and resource allocation within the phenylpropanoid pathway (PPP). Thus, we examined correlations between soil N levels and both expression patterns of eight PPP genes (measured by quantitative-reverse transcription PCR) and foliar phenolic compounds (measured by liquid chromatography-mass spectrometry) in young aspen genets with intrinsically extreme CT levels. Monitored phenolics included salicinoids, lignins, flavones, flavonols, CT precursors and CTs. The PPP genes were consistently expressed more strongly in high-CT trees. Low N supplements reduced expression of genes throughout the PPP in all genets, while high N doses restored expression of genes at the beginning and end of the pathway. These PPP changes were not reflected in pools of tannin precursors, but varying correlations between gene expression and foliar phenolic pools were detected in young and mature leaves, suggesting that processes linking gene expression and the resulting phenolics vary spatially and temporally. Precursor fluxes suggested that CT-related metabolic rate or sink controls are linked to intrinsic carbon allocation strategies associated with N responses. Overall, we found more negative correlations (indicative of allocation trade-offs) between PPP gene expression and phenolic products following N additions in low-CT plants than in high-CT plants. The tannin-related expression dynamics suggest that, in addition to defense, relative tannin levels may also be indicative of intraspecific variations in the way aspen genets respond to soil fertility. © The Author 2016. Published by Oxford University Press. All rights reserved. For Permissions, please e-mail: journals.permissions@oup.com.
Sirdaarta, Joseph; Maen, Anton; Rayan, Paran; Matthews, Ben; Cock, Ian Edwin
2016-05-01
High antioxidant capacities have been linked to the treatment and prevention of several cancers. Recent reports have identified several native Australian fruits with high antioxidant capacities. Despite this, several of these species are yet to be tested for anticancer activity. Solvent extracts prepared from high antioxidant native Australian fruits were analyzed for antioxidant capacity by the di (phenyl)-(2,4,6-trinitrophenyl) iminoazanium free radical scavenging assay. Antiproliferative activities against CaCo2 and HeLa cancer cells were determined by a multicellular tumor spheroid-based cell proliferation assay. Toxicity was determined by Artemia franciscana bioassay. Methanolic extracts of all plant species displayed high antioxidant contents (equivalent to approximately 7-16 mg of vitamin C per gram of fruit extracted). Most aqueous extracts also contained relatively high antioxidant capacities. In contrast, the ethyl acetate, chloroform, and hexane extracts of most species (except lemon aspen and bush tomato) had lower antioxidant contents (below 1.5 mg of vitamin C equivalents per gram of plant material extracted). The antioxidant contents correlated with the ability of the extracts to inhibit proliferation of CaCo2 and HeLa cancer cell lines. The high antioxidant methanolic extracts of all species were potent inhibitors of cell proliferation. The methanolic lemon aspen extract was particularly effective, with IC50 values of 480 and 769 μg/mL against HeLa and CaCo2 cells, respectively. In contrast, the lower antioxidant ethyl acetate and hexane extracts (except the lemon aspen ethyl acetate extract) generally did not inhibit cancer cell proliferation or inhibited to only a minor degree. Indeed, most of the ethyl acetate and hexane extracts induced potent cell proliferation. The native tamarind ethyl acetate extract displayed low-moderate toxicity in the A. franciscana bioassay (LC50 values below 1000 μg/mL). All other extracts were nontoxic. A total of 145 unique mass signals were detected in the lemon aspen methanolic and aqueous extracts by nonbiased high-performance liquid chromatography-mass spectrometry analysis. Of these, 20 compounds were identified as being of particular interest due to their reported antioxidant and/or anticancer activities. The lack of toxicity and antiproliferative activity of the high antioxidant plant extracts against HeLa and CaCo2 cancer cell lines indicates their potential in the treatment and prevention of some cancers. Australian fruit extracts with high antioxidant contents were potent inhibitors of CaCo2 and HeLa carcinoma cell proliferationMethanolic lemon aspen extract was particularly potent, with IC50 values of 480 μg/mL (HeLa) and 769 μg/mL (CaCo2)High-performance liquid chromatography-mass spectrometry-quadrupole time-of-flight analysis highlighted and putatively identified 20 compounds in the antiproliferative lemon aspen extractsIn contrast, lower antioxidant content extracts stimulated carcinoma cell proliferationAll extracts with antiproliferative activity were nontoxic in the Artemia nauplii assay. Abbreviations used: DPPH: di (phenyl)- (2,4,6-trinitrophenyl) iminoazanium, HPLC: High-performance liquid chromatography, IC50: The concentration required to inhibit by 50%, LC50: The concentration required to achieve 50% mortality, MS: Mass spectrometry.
Rats' preferences for corn versus wood-based bedding and nesting materials.
Ras, T; van de Ven, M; Patterson-Kane, E G; Nelson, K
2002-10-01
Corn by-products can be used as bedding and nesting products. Corn-cob bedding resists ammonia build-up and corn-husk nesting material resists dampness. It is not clear whether these advantages are at the expense of animal comfort. Corn cob was compared to aspen chip bedding, and corn husk to paper strip nesting material. Data from 20 rats with differential early bedding experience suggested that they prefer aspen chip, but are also biased towards the bedding they were raised on. Data from 10 rats with no prior nesting material experience suggested that paper strip was preferred over cornhusk. Thus, corn-cob products are not recommended except in situations where air quality and/or flooding are significant problems.
Orbital Express Mission Operations Planning and Resource Management using ASPEN
NASA Technical Reports Server (NTRS)
Chouinard, Caroline; Knight, Russell; Jones, Grailing; Tran, Danny
2008-01-01
The Orbital Express satellite servicing demonstrator program is a DARPA program aimed at developing "a safe and cost-effective approach to autonomously service satellites in orbit". The system consists of: a) the Autonomous Space Transport Robotic Operations (ASTRO) vehicle, under development by Boeing Integrated Defense Systems, and b) a prototype modular next-generation serviceable satellite, NEXTSat, being developed by Ball Aerospace. Flexibility of ASPEN: a) Accommodate changes to procedures; b) Accommodate changes to daily losses and gains; c) Responsive re-planning; and d) Critical to success of mission planning Auto-Generation of activity models: a) Created plans quickly; b) Repetition/Re-use of models each day; and c) Guarantees the AML syntax. One SRP per day vs. Tactical team
Downregulation of RWA genes in hybrid aspen affects xylan acetylation and wood saccharification.
Pawar, Prashant Mohan-Anupama; Ratke, Christine; Balasubramanian, Vimal K; Chong, Sun-Li; Gandla, Madhavi Latha; Adriasola, Mathilda; Sparrman, Tobias; Hedenström, Mattias; Szwaj, Klaudia; Derba-Maceluch, Marta; Gaertner, Cyril; Mouille, Gregory; Ezcurra, Ines; Tenkanen, Maija; Jönsson, Leif J; Mellerowicz, Ewa J
2017-06-01
High acetylation of angiosperm wood hinders its conversion to sugars by glycoside hydrolases, subsequent ethanol fermentation and (hence) its use for biofuel production. We studied the REDUCED WALL ACETYLATION (RWA) gene family of the hardwood model Populus to evaluate its potential for improving saccharification. The family has two clades, AB and CD, containing two genes each. All four genes are expressed in developing wood but only RWA-A and -B are activated by master switches of the secondary cell wall PtNST1 and PtMYB21. Histochemical analysis of promoter::GUS lines in hybrid aspen (Populus tremula × tremuloides) showed activation of RWA-A and -B promoters in the secondary wall formation zone, while RWA-C and -D promoter activity was diffuse. Ectopic downregulation of either clade reduced wood xylan and xyloglucan acetylation. Suppressing both clades simultaneously using the wood-specific promoter reduced wood acetylation by 25% and decreased acetylation at position 2 of Xylp in the dimethyl sulfoxide-extracted xylan. This did not affect plant growth but decreased xylose and increased glucose contents in the noncellulosic monosaccharide fraction, and increased glucose and xylose yields of wood enzymatic hydrolysis without pretreatment. Both RWA clades regulate wood xylan acetylation in aspen and are promising targets to improve wood saccharification. © 2017 The Authors. New Phytologist © 2017 New Phytologist Trust.
Lihavainen, Jenna; Keinänen, Markku; Keski-Saari, Sarita; Kontunen-Soppela, Sari; Sõber, Anu; Oksanen, Elina
2016-01-01
Relative air humidity (RH) is expected to increase in northern Europe due to climate change. Increasing RH reduces the difference of water vapour pressure deficit (VPD) between the leaf and the atmosphere, and affects the gas exchange of plants. Little is known about the effects of decreased VPD on plant metabolism, especially under field conditions. This study was conducted to determine the effects of artificially decreased VPD on silver birch (Betula pendula Roth.) and hybrid aspen (Populus tremula L.×P. tremuloides Michx.) foliar metabolite and nutrient profiles in a unique free air humidity manipulation (FAHM) field experiment during the fourth season of humidity manipulation, in 2011. Long-term exposure to decreased VPD modified nutrient homeostasis in tree leaves, as demonstrated by a lower N concentration and N:P ratio in aspen leaves, and higher Na concentration and lower K:Na ratio in the leaves of both species in decreased VPD than in ambient VPD. Decreased VPD caused a shift in foliar metabolite profiles of both species, affecting primary and secondary metabolites. Metabolic adjustment to decreased VPD included elevated levels of starch and heptulose sugars, sorbitol, hemiterpenoid and phenolic glycosides, and α-tocopherol. High levels of carbon reserves, phenolic compounds, and antioxidants under decreased VPD may modify plant resistance to environmental stresses emerging under changing climate. PMID:27255929
NASA Astrophysics Data System (ADS)
Arnott, J. C.; Katzenberger, J.
2015-12-01
The impacts of global climate change to regional scales are complex and cut across sectorial and jurisdictional boundaries, and therefore, a unique enterprise of collaboration between scientists, resource managers, and other stakeholders for development of adequate response strategies is required. Such collaboration has been exhibited between stakeholders, researchers, and a boundary organization—the Aspen Global Change Institute—since 2005 in assessing impacts and crafting policies in response with regard to climate change impacts in the mountain watershed surrounding Aspen, CO. A series of structured stakeholder interviews and town hall sessions, impact assessment reports, and intensive collaboration between various information providers and user groups has set the stage for development of both mitigation of and adaptation to climate change impacts. The most recent example of this has included the use of global scale climate model output to inform the development of resiliency strategies in response to extreme precipitation projections. The use of this kind of resource has been considered in a variety of decision-making contexts and has included the development of region- and decision-relevant qualitative scenarios that make use of quantitative model-based information. Results from this line of work that include feedback from actual users', a boundary organization, and researchers' perspectives will be reported along with examples of policy and implementation results.
Improving root-zone soil properties for Trembling Aspen in a reconstructed mine-site soil
NASA Astrophysics Data System (ADS)
Dyck, M. F.; Sabbagh, P.; Bockstette, S.; Landhäusser, S.; Pinno, B.
2014-12-01
Surface mining activities have significantly depleted natural tree cover, especially trembling aspen (Populus tremuloides), in the Boreal Forest and Aspen Parkland Natural Regions of Alberta. The natural soil profile is usually destroyed during these mining activities and soil and landscape reconstruction is typically the first step in the reclamation process. However, the mine tailings and overburden materials used for these new soils often become compacted during the reconstruction process because they are subjected to high amounts of traffic with heavy equipment. Compacted soils generally have low porosity and low penetrability through increased soil strength, making it difficult for roots to elongate and explore the soil. Compaction also reduces infiltration capacity and drainage, which can cause excessive runoff and soil erosion. To improve the pore size distribution and water transmission, subsoil ripping was carried out in a test plot at Genesee Prairie Mine, Alberta. Within the site, six replicates with two treatments each, unripped (compacted) and ripped (decompacted), were established with 20-m buffers between them. The main objective of this research was to characterize the effects of subsoil ripping on soil physical properties and the longevity of those effects.as well as soil water dynamics during spring snowmelt. Results showed improved bulk density, pore size distribution and water infiltration in the soil as a result of the deep ripping, but these improvements appear to be temporary.
Siedlecka, Anna; Wiklund, Susanne; Péronne, Marie-Amélie; Micheli, Fabienne; Lesniewska, Joanna; Sethson, Ingmar; Edlund, Ulf; Richard, Luc; Sundberg, Björn; Mellerowicz, Ewa J
2008-02-01
Wood cells, unlike most other cells in plants, grow by a unique combination of intrusive and symplastic growth. Fibers grow in diameter by diffuse symplastic growth, but they elongate solely by intrusive apical growth penetrating the pectin-rich middle lamella that cements neighboring cells together. In contrast, vessel elements grow in diameter by a combination of intrusive and symplastic growth. We demonstrate that an abundant pectin methyl esterase (PME; EC 3.1.1.11) from wood-forming tissues of hybrid aspen (Populus tremula x tremuloides) acts as a negative regulator of both symplastic and intrusive growth of developing wood cells. When PttPME1 expression was up- and down-regulated in transgenic aspen trees, the PME activity in wood-forming tissues was correspondingly altered. PME removes methyl ester groups from homogalacturonan (HG) and transgenic trees had modified HG methylesterification patterns, as demonstrated by two-dimensional nuclear magnetic resonance and immunostaining using PAM1 and LM7 antibodies. In situ distributions of PAM1 and LM7 epitopes revealed changes in pectin methylesterification in transgenic trees that were specifically localized in expanding wood cells. The results show that en block deesterification of HG by PttPME1 inhibits both symplastic growth and intrusive growth. PttPME1 is therefore involved in mechanisms determining fiber width and length in the wood of aspen trees.
Veksha, Andrei; Bhuiyan, Tazul I.; Hill, Josephine M.
2016-01-01
Several samples of activated carbon were prepared by physical (CO2) and chemical (H3PO4) activation of aspen wood and tested for the adsorption of organic compounds from water generated during the recovery of bitumen using steam assisted gravity drainage. Total organic carbon removal by the carbon samples increased proportionally with total pore volume as determined from N2 adsorption isotherms at −196 °C. The activated carbon produced by CO2 activation had similar removal levels for total organic carbon from the water (up to 70%) to those samples activated with H3PO4, but lower yields, due to losses during pyrolysis and activation. A method to increase the yield when using CO2 activation was proposed and consisted of recycling bio-oil produced from previous runs to the aspen wood feed, followed by either KOH addition (0.48%) or air pretreatment (220 °C for 3 h) before pyrolysis and activation. By recycling the bio-oil, the yield of CO2 activated carbon (after air pretreatment of the mixture) was increased by a factor of 1.3. Due to the higher carbon yield, the corresponding total organic carbon removal, per mass of wood feed, increased by a factor of 1.2 thus improving the overall process efficiency. PMID:28787817
Feau, Nicolas; Vialle, Agathe; Allaire, Mathieu; Tanguay, Philippe; Joly, David L; Frey, Pascal; Callan, Brenda E; Hamelin, Richard C
2009-01-01
Wide variation and overlap in morphological characters have led to confusion in species identification within the fungal rust genus Melampsora. The Melampsora species with uredinial-telial stages on white poplar and aspens are especially prone to misidentification. This group includes the Melampsora populnea species complex and the highly destructive pine twisting rust, Melampsora pinitorqua, which alternates between hosts in Populus section Populus and Pinus. Our objective was to compare morphologically based identification to genetic material extracted from Melampsora species pathogenic to aspen and white poplar. We compared morphometric traits and DNA barcodes obtained from internal transcribed spacer (ITS), large ribosomal RNA subunit (28S), and mitochondrial cytochrome oxidase 1 (CO1) sequences to delimit within this taxonomically difficult group. Eight different Melampsora species were initially defined based on host specificity and morphometric data. DNA barcodes were then overlaid on these initial species definitions. The DNA barcodes, specifically those defined on ITS and 28S sequences, provided a highly accurate means of identifying and resolving Melampsora taxa. We highlighted species misidentification in specimens from Canadian herbaria related to either Melampsora medusae f. sp. tremuloidae or Melampsora aecidioides. Finally, we evidenced that the north-American species found on Populus alba, M. aecidioides is closely related but distinct from the four species of the M. populnea complex (Melampsora larici-tremulae, Melampsora magnusiana, Melampsora pinitorqua, and Melampsora rostrupii) found in Eurasia.
AmeriFlux US-UMd UMBS Disturbance
Curtis, Peter [Ohio State University; Gough, Christopher [Virginia Commonwealth University
2016-01-01
This is the AmeriFlux version of the carbon flux data for the site US-UMd UMBS Disturbance. Site Description - The UMBS Disturbance site is an artificial disturbance site that has recently been created as part of the Forest Accelerate Succession ExperimenT (FASET). In Spring 2008, every aspen and birch tree (>6,700, ~35% canopy LAI), the dominant early successional trees, were girdled over 39 ha of the FASET treatment plot to stimulate a disturbance that will move the forest into a later successional stage, dominated by maples, oaks, and white pine. This treatment caused aspen and birch mortality within 2 - 3 years. As a result of the changed canopy structure, there is a divergence in net ecosystem exchange between the control plot (enhanced carbon uptake) and the treatment plot (reduced carbon uptake).
Manuwoto, S; Scriber, J M; Hsia, M T; Sunarjo, P
1985-08-01
Previous studies have shown leaves of tulip tree, Liriodendron tulipifera L. (of the Magnoliaceae) and of Populus tremuloides Michx. (of the Salicaceae) to be antixenotic/antibiotic to many Lepidoptera, including one of the most polyphagous of all phytophagous insects, the southern armyworm, Spodoptera eridania Cramer (Noctuidae). We investigated the physiological responses to this phytochemical activity on neonate and late instar armyworm larvae in controlled environments with particular emphasis upon the leaf extracts containing condensed tannins and hydrolysable tannins. These tannin-containing extracts of tulip tree leaves and quaking aspen leaves were generally toxic to neonate larvae. For later instars, growth suppression was not due to digestibility-reduction, but instead to suppressed consumption rates and greatly increased metabolic (respiratory) costs as reflected in reduced biomass conversion efficiencies.
Aspen Global Change Institute: 25 Years of Interdisciplinary Global Change Science
DOE Office of Scientific and Technical Information (OSTI.GOV)
Meehl, Gerald A.; Moss, Richard
Global environmental changes such as climate change result from the interaction of human and natural systems. Research to understand these changes and options for addressing them requires the physical, environmental, and social sciences, as well as engineering and other applied fields. In this essay, we describe how the Aspen Global Change Institute (AGCI) has provided leadership in global change science over the past 25 years—in particular how it has contributed to the integration of the natural and social sciences needed to research the drivers of change, Earth system response, natural and human system impacts, and options for risk management. Wemore » illustrate the ways the history of AGCI has been intertwined with the evolution of global change science as it has become an increasingly interdisciplinary endeavor.« less
Sirdaarta, Joseph; Maen, Anton; Rayan, Paran; Matthews, Ben; Cock, Ian Edwin
2016-01-01
Background: High antioxidant capacities have been linked to the treatment and prevention of several cancers. Recent reports have identified several native Australian fruits with high antioxidant capacities. Despite this, several of these species are yet to be tested for anticancer activity. Materials and Methods: Solvent extracts prepared from high antioxidant native Australian fruits were analyzed for antioxidant capacity by the di (phenyl)-(2,4,6-trinitrophenyl) iminoazanium free radical scavenging assay. Antiproliferative activities against CaCo2 and HeLa cancer cells were determined by a multicellular tumor spheroid-based cell proliferation assay. Toxicity was determined by Artemia franciscana bioassay. Results: Methanolic extracts of all plant species displayed high antioxidant contents (equivalent to approximately 7–16 mg of vitamin C per gram of fruit extracted). Most aqueous extracts also contained relatively high antioxidant capacities. In contrast, the ethyl acetate, chloroform, and hexane extracts of most species (except lemon aspen and bush tomato) had lower antioxidant contents (below 1.5 mg of vitamin C equivalents per gram of plant material extracted). The antioxidant contents correlated with the ability of the extracts to inhibit proliferation of CaCo2 and HeLa cancer cell lines. The high antioxidant methanolic extracts of all species were potent inhibitors of cell proliferation. The methanolic lemon aspen extract was particularly effective, with IC50 values of 480 and 769 μg/mL against HeLa and CaCo2 cells, respectively. In contrast, the lower antioxidant ethyl acetate and hexane extracts (except the lemon aspen ethyl acetate extract) generally did not inhibit cancer cell proliferation or inhibited to only a minor degree. Indeed, most of the ethyl acetate and hexane extracts induced potent cell proliferation. The native tamarind ethyl acetate extract displayed low-moderate toxicity in the A. franciscana bioassay (LC50 values below 1000 μg/mL). All other extracts were nontoxic. A total of 145 unique mass signals were detected in the lemon aspen methanolic and aqueous extracts by nonbiased high-performance liquid chromatography-mass spectrometry analysis. Of these, 20 compounds were identified as being of particular interest due to their reported antioxidant and/or anticancer activities. Conclusions: The lack of toxicity and antiproliferative activity of the high antioxidant plant extracts against HeLa and CaCo2 cancer cell lines indicates their potential in the treatment and prevention of some cancers. SUMMARY Australian fruit extracts with high antioxidant contents were potent inhibitors of CaCo2 and HeLa carcinoma cell proliferationMethanolic lemon aspen extract was particularly potent, with IC50 values of 480 μg/mL (HeLa) and 769 μg/mL (CaCo2)High-performance liquid chromatography-mass spectrometry-quadrupole time-of-flight analysis highlighted and putatively identified 20 compounds in the antiproliferative lemon aspen extractsIn contrast, lower antioxidant content extracts stimulated carcinoma cell proliferationAll extracts with antiproliferative activity were nontoxic in the Artemia nauplii assay. Abbreviations used: DPPH: di (phenyl)- (2,4,6-trinitrophenyl) iminoazanium, HPLC: High-performance liquid chromatography, IC50: The concentration required to inhibit by 50%, LC50: The concentration required to achieve 50% mortality, MS: Mass spectrometry. PMID:27279705