Gross Basal Area Growth of Northern White-Cedar is Independent of Stand Density Over A Wide Range
Bruce W. Foltz; William F. Johnston
1968-01-01
A 20-year study in Wisconsin swamp stand of northern white-cedar indicates that after a second thinning gross basal area growth is independent of stand densities ranging from 90 to 225 square feet of basal area per acre. Mortality, net growth, and ingrowth were also independent of density in the thinned plots.
Effects of low-density thinning in a declining white pine stand in Maine
William B. Leak; Mariko. Yamasaki
2013-01-01
Low-density (32 ft2/acre residual basal area) and medium-low density (60 ft2/acre residual basal area) thinnings were studied over a 4-year period in a declining white pine stand on the Massabesic Experimental Forest in southern Maine. Gross basal area growth at 60 ft2 was about three-fourths the rate...
Norghauer, Julian M; Newbery, David M
2016-05-01
Although negative density dependence (NDD) can facilitate tree species coexistence in forests, the underlying mechanisms can differ, and rarely are the dynamics of seedlings and saplings studied together. Herein we present and discuss a novel mechanism based on our investigation of NDD predictions for the large, grove-forming ectomycorrhizal mast fruiting tree, Microberlinia bisulcata (Caesalpiniaceae), in an 82.5-ha plot at Korup, Cameroon. We tested whether juvenile density, size, growth and survival decreases with increasing conspecific adult basal area for 3245 'new' seedlings and 540 'old' seedlings (< 75-cm tall) during an approximately 4-year study period (2008-2012) and for 234 'saplings' (≥ 75-cm tall) during an approximately 6-year study period (2008-2014). We found that the respective densities of new seedlings, old seedlings and saplings were positively, not and negatively related to increasing BA. Maximum leaf numbers and heights of old seedlings were negatively correlated with increasing basal areas, as were sapling heights and stem diameters. Whereas survivorship of new seedlings decreased by more than one-half with increasing basal area over its range in 2010-2012, that of old seedlings decreased by almost two-thirds, but only in 2008-2010, and was generally unrelated to conspecific seedling density. In 2010-2012 relative growth rates in new seedlings' heights decreased with increasing basal area, as well as with increasing seedling density, together with increasing leaf numbers, whereas old seedlings' growth was unrelated to either conspecific density or basal area. Saplings of below-average height had reduced survivorship with increasing basal area (probability decreasing from approx. 0.4 to 0.05 over the basal area range tested), but only sapling growth in terms of leaf numbers decreased with increasing basal area. These static and dynamic results indicate that NDD is operating within this system, possibly stabilizing the M. bisulcata population. However, these NDD patterns are unlikely to be caused by symmetric competition or by consumers. Instead, an alternative mechanism for conspecific adult-juvenile negative feedback is proposed, one which involves the interaction between tree phenology and ectomycorrhizal linkages.
Growth of ponderosa pine thinned to different stocking levels in central Oregon: 30-year results.
P.H. Cochran; James W. Barrett
1999-01-01
Periodic annual increments (PAI) for survivor diameters decreased curvilinearly with increasing stand density. Gross volume and basal areas PAIs increased linearly with increasing stand density. Growth of basal area and volume for the 20 largest trees per acre were reduced curvilinearly with increasing stand density. Bark beetles were the primary cause of mortality. No...
Predicting oak density with ecological, physical, and soil indicators
Callie Jo Schweitzer; Adrian A. Lesak; Yong Wang
2006-01-01
We predicted density of oak species in the mid-Cumberland Plateau region of northeastern Alabama on the basis of basal area of tree associations based on light tolerances, physical site characteristics, and soil type. Tree basal area was determined for four species groups: oaks (Quercus spp.), hickories (Carya spp.), yellow-poplar...
Zhang, Hongmin; He, Siyu; Liu, Susu; Xie, Yanting; Chen, Guoming; Zhang, Junjie; Sun, Shengtao; Liang, David; Wang, Liya
2016-04-01
To measure the cell size and cell density in five layers of the central cornea in the widely used inbred C57BL/6 mouse strain using in vivo three-dimensional (3D) two-photon (2PH) imaging. Corneas were scanned using a 2PH laser scanning fluorescence microscope after staining with plasma membrane stain and Hoechst 33342. Good quality 3D images were selected for the cell density and cell size analysis. Cell density was determined by counting the cell nuclei in a predefined cube of 3D images. Cell size measurements, including cell surface area, cell volume, nuclear surface area and nuclear volume, were automatically quantified using the Imaris software. The cell and nuclear surface-area-to-volume ratio (S:V ratio) and the cell nuclear-cytoplasmic ratio (N:C ratio) were calculated. The highest cell density was observed in the basal epithelium and the lowest in the posterior stroma. The highest cell surface area was found in the anterior stroma, and the highest cell volume was observed in the superficial epithelium. The lowest cell surface area and cell volume were both found in the basal epithelium. The highest S:V ratio was observed in the basal epithelium and the lowest in the superficial epithelium. The highest cell nuclear surface area and volume were both observed in the superficial epithelium and the lowest in the basal epithelium. The highest cell nuclear S:V ratio was observed in the basal epithelium and the lowest in the superficial epithelium. The highest N:C ratio was found in the basal epithelial cells and the lowest in the posterior keratocytes. We are the first to quantify the cell density and size parameters, including cell surface area and volume, cell nuclear surface area and volume, and the S:V ratio, in the five layers of the central cornea. These data provide important cell morphology features for the study of corneal physiology, pathology and disease in mice, particularly in C57BL/6 mice.
Homeostatic maintenance of ponderosa pine gas exchange in response to stand density changes.
McDowell, Nate G; Adams, Henry D; Bailey, John D; Hess, Marcey; Kolb, Thomas E
2006-06-01
Homeostatic maintenance of gas exchange optimizes carbon gain per water loss. Homeostasis is regulated by short-term physiological and long-term structural mechanisms, both of which may respond to changes in resource availability associated with competition. Therefore, stand density regulation via silvicultural manipulations may facilitate growth and survival through mechanisms operating at both short and long timescales. We investigated the responses of ponderosa pine (Pinus ponderosa) to stand basal area manipulations in Arizona, USA. Stand basal area was manipulated to seven replicated levels in 1962 and was maintained for four decades by decadal thinning. We measured basal area increment (BAI) to assess the response and sustainability of wood growth, carbon isotope discrimination (A) inferred from annual rings to assess the response of crown gas exchange, and ratios of leaf area to sapwood area (A(l):A(s)) to assess longer term structural acclimation. Basal area treatments increased soil water potential (r2 = 0.99) but did not affect photosynthetic capacity. BAI increased within two years of thinning, and the 40-year mean BAI was negatively correlated with stand basal area (r2 = 0.98). delta was negatively correlated with stand basal area for years 5 through 12 after thinning (r2 = 0.90). However, delta was relatively invariant with basal area for the period 13-40 years after initial thinning despite maintenance of treatment basal areas via repeated decadal thinnings. Independent gas exchange measurements verified that the ratio of photosynthesis to stomatal conductance was invariant with basal area, but absolute values of both were elevated at lower basal areas. A(l):A(s) was negatively correlated with basal area (r2 = 0.93). We hypothesize that increased A(l):A(s) is a homeostatic response to increased water availability that maximizes water-use efficiency and whole-tree carbon uptake. Elevated A(l):A(s) of trees at low basal areas was associated with greater resilience to climate, i.e., greater absolute BAI during drought; however, trees with high A(l):A(s) in low basal area stands also exhibited the greatest sensitivity to drought, i.e., greater relative decline in BAI.
A density management diagram for even-aged ponderosa pine stands
James N. Long; John D. Shaw
2005-01-01
We developed a density management diagram (DMD) for ponderosa pine using Forest Inventory and Analysis (FIA) data. Analysis plots were drawn from all FIA plots in the western United States on which ponderosa pine occurred. A total of 766 plots met the criteria for analysis. Selection criteria were for purity, defined as ponderosa pine basal area 80% of plot basal area...
The Influence of Stand Density and Structure on Growth of Northern Hardwoods in New England
Dale S. Solomon
1977-01-01
Growth of northern hardwoods over a 10-year period was studied in plots that were treated to produce residual densities of 40, 60, 80, and 100 square feet of basal area per acre with stand structures of 30, 45 and 60 percent sawtimber. Both diameter and basal area growth are tabulated by treatment and species.
Ivan L. Sander
1986-01-01
Results from a shortleaf pine thinning study in Missouri show that continually thinning a stand to the same basal area will eventually create an understocked stand and reduce yields. Using stocking percent to control thinning intensity allows basal area to increase as stands get older. The best yield should occur when shortleaf pine is repeatedly thinned to 60 percent...
Andrew T. Hudak; Nicholas L. Crookston; Jeffrey S. Evans; Michael K. Falkowski; Alistair M. S. Smith; Paul E. Gessler; Penelope Morgan
2006-01-01
We compared the utility of discrete-return light detection and ranging (lidar) data and multispectral satellite imagery, and their integration, for modeling and mapping basal area and tree density across two diverse coniferous forest landscapes in north-central Idaho. We applied multiple linear regression models subset from a suite of 26 predictor variables derived...
Curtis, Robert O.; Bansal, Sheel; Harrington, Constance A.
2016-01-01
This report presents updated information on a 1981 Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco var. menziesii) plantation spacing trial at 33 years from planting. Stand statistics at the most recent measurement were compared for initial spacing of 1 through 6 meters and associated relative densities. There was no clear relationship of spacing to top height. Diameter, live crown ratio, and percent survival increased with spacing; basal area and relative density decreased with increase in spacing. Volume in trees ≥ 4 cm diameter was greatest at 2 m spacing, while utilizable volume (trees ≥20 cm dbh) was greatest at 4 m spacing. Live crown ratio decreased and total crown projectional area increased with increasing relative density indices. Total crown projectional area was more closely related to relative density than to basal area.
Fu, Xiaoli; Wang, Jianlei; Di, Yuebao; Wang, Huimin
2015-01-01
Variation of total fine-root biomass among types of tree stands has previously been attributed to the characteristics of the stand layers. The effects of the understory vegetation on total fine-root biomass are less well studied. We examined the variation of total fine-root biomass in subtropical tree stands at two sites of Datian and Huitong in China. The two sites have similar humid monsoon climate but different soil organic carbon. One examination compared two categories of basal areas (high vs. low basal area) in stands of single species. A second examination compared single-species and mixed stands with comparable basal areas. Low basal area did not correlate with low total fine-root biomass in the single-species stands. The increase in seedling density but decrease in stem density for the low basal area stands at Datian and the quite similar stand structures for the basal-area contrast at Huitong helped in the lack of association between basal area and total fine-root biomass at the two sites, respectively. The mixed stands also did not yield higher total fine-root biomasses. In addition to the lack of niche complementarity between tree species, the differences in stem and seedling densities and the belowground competition between the tree and non-tree species also contributed to the similarity of the total fine-root biomasses in the mixed and single-species stands. Across stand types, the more fertile site Datian yielded higher tree, non-tree and total fine-root biomasses than Huitong. However, the contribution of non-tree fine-root biomass to the total fine-root biomass was higher at Huitong (29.4%) than that at Datian (16.7%). This study suggests that the variation of total fine-root biomass across stand types not only was associated with the characteristics of trees, but also may be highly dependent on the understory layer. PMID:26047358
Fu, Xiaoli; Wang, Jianlei; Di, Yuebao; Wang, Huimin
2015-01-01
Variation of total fine-root biomass among types of tree stands has previously been attributed to the characteristics of the stand layers. The effects of the understory vegetation on total fine-root biomass are less well studied. We examined the variation of total fine-root biomass in subtropical tree stands at two sites of Datian and Huitong in China. The two sites have similar humid monsoon climate but different soil organic carbon. One examination compared two categories of basal areas (high vs. low basal area) in stands of single species. A second examination compared single-species and mixed stands with comparable basal areas. Low basal area did not correlate with low total fine-root biomass in the single-species stands. The increase in seedling density but decrease in stem density for the low basal area stands at Datian and the quite similar stand structures for the basal-area contrast at Huitong helped in the lack of association between basal area and total fine-root biomass at the two sites, respectively. The mixed stands also did not yield higher total fine-root biomasses. In addition to the lack of niche complementarity between tree species, the differences in stem and seedling densities and the belowground competition between the tree and non-tree species also contributed to the similarity of the total fine-root biomasses in the mixed and single-species stands. Across stand types, the more fertile site Datian yielded higher tree, non-tree and total fine-root biomasses than Huitong. However, the contribution of non-tree fine-root biomass to the total fine-root biomass was higher at Huitong (29.4%) than that at Datian (16.7%). This study suggests that the variation of total fine-root biomass across stand types not only was associated with the characteristics of trees, but also may be highly dependent on the understory layer.
Partial denervation of sub-basal axons persists following debridement wounds to the mouse cornea.
Pajoohesh-Ganji, Ahdeah; Pal-Ghosh, Sonali; Tadvalkar, Gauri; Kyne, Briana M; Saban, Daniel R; Stepp, Mary Ann
2015-11-01
Although sensory reinnervation occurs after injury in the peripheral nervous system, poor reinnervation in the elderly and those with diabetes often leads to pathology. Here we quantify sub-basal axon density in the central and peripheral mouse cornea over time after three different types of injury. The mouse cornea is highly innervated with a dense array of sub-basal nerves that form a spiral called the vortex at the corneal center or apex; these nerves are readily detected within flat mounted corneas. After anesthesia, corneal epithelial cells were removed using either a dulled blade or a rotating burr within an area demarcated centrally with a 1.5 mm trephine. A third wound type, superficial trephination, involved demarcating the area with the 1.5 mm trephine but not removing cells. By 7 days after superficial trephination, sub-basal axon density returns to control levels; by 28 days the vortex reforms. Although axon density is similar to control 14 days after dulled blade and rotating burr wounding, defects in axon morphology at the corneal apex remain. After 14 days, axons retract from the center leaving the sub-basal axon density reduced by 37.2 and 36.8% at 28 days after dulled blade and rotating burr wounding, respectively, compared with control. Assessment of inflammation using flow cytometry shows that persistent inflammation is not a factor in the incomplete reinnervation. Expression of mRNAs encoding 22 regeneration-associated genes involved in axon targeting assessed by QPCR reveals that netrin-1 and ephrin signaling are altered after wounding. Subpopulations of corneal epithelial basal cells at the corneal apex stop expressing ki67 as early as 7 days after injury and by 14 and 28 days after wounding, many of these basal cells undergo apoptosis and die. Although sub-basal axons are restored to their normal density and morphology after superficial trephination, sub-basal axon recovery is partial after debridement wounds. The increase in corneal epithelial basal cell apoptosis at the apex observed at 14 days after corneal debridement may destabilize newly reinnervated sub-basal axons and lead to their retraction toward the periphery.
Verity Salmon; Colleen Iversen; Peter Thornton; Ma
2017-03-01
Transect data is from point center quarter surveys for shrub density performed in July 2016 at the Kougarok hill slope located at Kougarok Road, Mile Marker 64. For each sample point along the transects, moving averages for shrub density and shrub basal area are provided along with GPS coordinates, average shrub height and active layer depth. The individual height, basal area, and species of surveyed shrubs are also included. Data upload will be completed January 2017.
NASA Astrophysics Data System (ADS)
Varo-Martínez, Mª Ángeles; Navarro-Cerrillo, Rafael M.; Hernández-Clemente, Rocío; Duque-Lazo, Joaquín
2017-04-01
Traditionally, forest-stand delineation has been assessed based on orthophotography. The application of LiDAR has improved forest management by providing high-spatial-resolution data on the vertical structure of the forest. The aim of this study was to develop and test a semi-automated algorithm for stands delineation in a plantation of Pinus sylvestris L. using LiDAR data. Three specific objectives were evaluated, i) to assess two complementary LiDAR metrics, Assmann dominant height and basal area, for the characterization of the structure of P. sylvestris Mediterranean forests based on object-oriented segmentation, ii) to evaluate the influence of the LiDAR pulse density on forest-stand delineation accuracy, and iii) to investigate the algorithmś effectiveness in the delineation of P. sylvestris stands for map prediction of Assmann dominant height and basal area. Our results show that it is possible to generate accurate P. sylvestris forest-stand segmentations using multiresolution or mean shift segmentation methods, even with low-pulse-density LiDAR - which is an important economic advantage for forest management. However, eCognition multiresolution methods provided better results than the OTB (Orfeo Tool Box) for stand delineation based on dominant height and basal area estimations. Furthermore, the influence of pulse density on the results was not statistically significant in the basal area calculations. However, there was a significant effect of pulse density on Assmann dominant height [F2,9595 = 5.69, p = 0.003].for low pulse density. We propose that the approach shown here should be considered for stand delineation in other large Pinus plantations in Mediterranean regions with similar characteristics.
Scott, Michael L.; Reynolds, Elizabeth W.
2007-01-01
Compared to 5-m by 20-m tree quadrats, belt transects were shown to provide similar estimates of stand structure (stem density and stand basal area) in less than 30 percent of the time. Further, for the streams sampled, there were no statistically significant differences in stem density and basal area estimates between 10-m and 20-m belt transects and the smaller belts took approximately half the time to sample. There was, however, high variance associated with estimates of stand structure for infrequently occurring stems, such as large, relict or legacy riparian trees. Legacy riparian trees occurred in limited numbers at all sites sampled. A reachscale population census of these trees indicated that the 10-m belt transects tended to underestimate both stem density and basal area for these riparian forest elements and that a complete reach-scale census of legacy trees averaged less than one hour per site.
The Relationship Between Basal Area and Hard Mast Production in the Ouachita Mountains
Roger W. Perry; Ronald E. Thill; Philip A. Tappe; David G. Peitz
2004-01-01
Abstract - Because the relationship between stand density and hard mast production is not clear, we investigated the effects of varying total overstory basal area (BA) on acorn and hickory nut production in the Ouachita Mountains. We used Whitehead visual surveys to estimate mast production in oaks (Quercus spp.) and hickories (...
Initial thinning in red pine plantations.
John H. Cooley
1969-01-01
Response to thinning in six red pine plantations in Lower Michigan supported previous findings by showing basal area growth to vary little over a wide range of residual densities. Furthermore, the method of selecting trees for removal in the first thinning had little or no influence on basal area growth. However, the immediate effect of thinning on average stand...
Shortleaf pine reproduction abundance and growth in pine-oak stands in the Missouri Ozarks
Elizabeth M. Blizzard; Doyle Henken; John M. Kabrick; Daniel C. Dey; David R. Larsen; David Gwaze
2007-01-01
We conducted an operational study to evaluate effect of site preparation treatments on pine reproduction density and the impact of overstory basal area and understory density on pine reproduction height and basal diameter in pine-oak stands in the Missouri Ozarks. Stands were harvested to or below B-level stocking, but patchiness of the oak decline lead to some plots...
Moore, Georgianne W; Bond, Barbara J; Jones, Julia A; Phillips, Nathan; Meinzer, Federick C
2004-05-01
Large areas of forests in the Pacific Northwest are being transformed to younger forests, yet little is known about the impact this may have on hydrological cycles. Previous work suggests that old trees use less water per unit leaf area or sapwood area than young mature trees of the same species in similar environments. Do old forests, therefore, use less water than young mature forests in similar environments, or are there other structural or compositional components in the forests that compensate for tree-level differences? We investigated the impacts of tree age, species composition and sapwood basal area on stand-level transpiration in adjacent watersheds at the H.J. Andrews Forest in the western Cascades of Oregon, one containing a young, mature (about 40 years since disturbance) conifer forest and the other an old growth (about 450 years since disturbance) forest. Sap flow measurements were used to evaluate the degree to which differences in age and species composition affect water use. Stand sapwood basal area was evaluated based on a vegetation survey for species, basal area and sapwood basal area in the riparian area of two watersheds. A simple scaling exercise derived from estimated differences in water use as a result of differences in age, species composition and stand sapwood area was used to estimate transpiration from late June through October within the entire riparian area of these watersheds. Transpiration was higher in the young stand because of greater sap flux density (sap flow per unit sapwood area) by age class and species, and greater total stand sapwood area. During the measurement period, mean daily sap flux density was 2.30 times higher in young compared with old Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) trees. Sap flux density was 1.41 times higher in young red alder (Alnus rubra Bong.) compared with young P. menziesii trees, and was 1.45 times higher in old P. menziesii compared with old western hemlock (Tsuga heterophylla (Raf.) Sarg.) trees. Overall, sapwood basal area was 21% higher in the young stand than in the old stand. In the old forest, T. heterophylla is an important co-dominant, accounting for 58% of total sapwood basal area, whereas P. menziesii is the only dominant conifer in the young stand. Angiosperms accounted for 36% of total sapwood basal area in the young stand, but only 7% in the old stand. For all factors combined, we estimated 3.27 times more water use by vegetation in the riparian area of the young stand over the measurement period. Tree age had the greatest effect on stand differences in water use, followed by differences in sapwood basal area, and finally species composition. The large differences in transpiration provide further evidence that forest management alters site water balance via elevated transpiration in vigorous young stands.
Design Difficulties in Stand Density Studies
Frank A. Bennett
1969-01-01
Designing unbiased stand density studies is difficult. An acceptable sample requires stratification of the plots of age, site, and density. When basal area, percent stocking, or Reineke's stand density index is used as the density measure, this stratification forces a high negative correlation between site and number of trees per acre. Mortality in trees per acre...
A simple index of stand density for Douglas-fir.
R.O. Curtis
1982-01-01
The expression RD = G/(Dg½), where G is basal area and Dg is quadratic mean stand diameter, provides a simple and convenient scale of relative stand density for Douglas-fir, equivalent to other generally accepted diameter-based stand density measures.
Douglas J. Stevenson; Thomas B. Lynch; James M. Guldin
2010-01-01
Shortleaf pine grows in association with many other species, particularly understory hardwoods, which compete with it, limiting its productivity (Bower and Ferguson 1968, Cain 1988). Which species are the most competitive varies with site quality, density of the pine overstory and years since thinning. Basal area and site index closely approximate the principle...
Estimating historical snag density in dry forests east of the Cascade Range
Richy J. Harrod; William L. Gaines; William E. Hartl; Ann. Camp
1998-01-01
Estimating snag densities in pre-European settlement landscapes (i.e., historical conditions) provides land managers with baseline information for comparing current snag densities. We propose a method for determining historical snag densities in the dry forests east of the Cascade Range. Basal area increase was calculated from tree ring measurements of old ponderosa...
Species composition, diversity and structure of novel forests of Castilla elastica in Puerto Rico
J. Fonseca da Silva; NO-VALUE
2014-01-01
Attributes of novel forests (secondary forests containing introduced species) were compared with those of native secondary forests of similar age. The study area was the biological reserve called El Tallonal, in Puerto Rico. Species composition, tree density, basal area and soil bulk density were characterized; Importance value index (IVI), Shannonâs diversity index (H...
P.H. Cochran; Walter G. Dahms
2000-01-01
Concave curvilinear decreases in diameter growth occurred with increasing stand densities. A convex curvilinear increase in gross growth of basal area and total cubic volume took place with increasing stand density. Maximum cumulative net cubic (total and merchantable) and board-foot yields varied curvilinearly with stand density. These yields peaked at intermediate...
Boris Zeide
2004-01-01
Estimation of stand density is based on a relationship between number of trees and their average diameter in fully stocked stands. Popular measures of density (Reinekeâs stand density index and basal area) assume that number of trees decreases as a power function of diameter. Actually, number of trees drops faster than predicted by the power function because the number...
Best density and structure for uneven-aged northern hardwood management in New England
William B. Leak
2003-01-01
Choice of the best residual density (basal area per acre) and structure (diameter distribution) for uneven-aged management of northern hardwoods is a complex decision that depends on the manager's decision rules, product objectives, site conditions, and - perhaps most important - current stand conditions. In contrast to other recommendations on residual density...
NASA Astrophysics Data System (ADS)
Mohammadi, Jahangir; Shataee, Shaban; Namiranian, Manochehr; Næsset, Erik
2017-09-01
Inventories of mixed broad-leaved forests of Iran mainly rely on terrestrial measurements. Due to rapid changes and disturbances and great complexity of the silvicultural systems of these multilayer forests, frequent repetition of conventional ground-based plot surveys is often cost prohibitive. Airborne laser scanning (ALS) and multispectral data offer an alternative or supplement to conventional inventories in the Hyrcanian forests of Iran. In this study, the capability of a combination of ALS and UltraCam-D data to model stand volume, tree density, and basal area using random forest (RF) algorithm was evaluated. Systematic sampling was applied to collect field plot data on a 150 m × 200 m sampling grid within a 1100 ha study area located at 36°38‧- 36°42‧N and 54°24‧-54°25‧E. A total of 308 circular plots (0.1 ha) were measured for calculation of stand volume, tree density, and basal area per hectare. For each plot, a set of variables was extracted from both ALS and multispectral data. The RF algorithm was used for modeling of the biophysical properties using ALS and UltraCam-D data separately and combined. The results showed that combining the ALS data and UltraCam-D images provided a slight increase in prediction accuracy compared to separate modeling. The RMSE as percentage of the mean, the mean difference between observed and predicted values, and standard deviation of the differences using a combination of ALS data and UltraCam-D images in an independent validation at 0.1-ha plot level were 31.7%, 1.1%, and 84 m3 ha-1 for stand volume; 27.2%, 0.86%, and 6.5 m2 ha-1 for basal area, and 35.8%, -4.6%, and 77.9 n ha-1 for tree density, respectively. Based on the results, we conclude that fusion of ALS and UltraCam-D data may be useful for modeling of stand volume, basal area, and tree density and thus gain insights into structural characteristics in the complex Hyrcanian forests.
Longleaf pine cone production in relation to site index, stand age, and stand density
Thomas Croker
1973-01-01
Few cones were produced in stands less than 30 years old. In stands 30 to 70 years in age, production seemed best at timber densities of about 30 square feet of basal area per acre, and tended to increase with increasing site index.
Influence of residual stand density on regeneration of northern hardwoods
W. B. Leak; Dale S. Solomon
1975-01-01
In a study of hardwood regeneration, experimental plots were treated to produce residual densities of 40, 60, 80, and 100 square feet of basal area per acre with 30, 45, and 60 percent sawtimber. After 9 years, numbers of stems and percent stocking were estimated for each species and they were rated for reproductive potential under different residual densities. Data...
Woody encroachment and its consequences on hydrological processes in the savannah
2016-01-01
Woody encroachment due to changes in climate or in the disturbance regimes (fire and herbivory) has been observed throughout the savannah biome over the last century with ecological, hydrological and socioeconomic consequences. We assessed changes in tree density and basal area and estimated changes in rain interception by the canopies across a 5-year period over a biomass gradient in Cerrado vegetation protected from fire. We modelled throughfall, stemflow and net rainfall on the basis of tree basal area (TBA). Tree density increased by an average annual rate of 6.7%, basal area at 5.7% and rain interception by the canopies at 0.6% of the gross rainfall. Independent of the vegetation structure, we found a robust relationship of 0.9% less rainfall reaching the ground as TBA increases by 1 m2 ha−1. Increases in tree biomass with woody encroachment may potentially result in less water available for uptake by plants and to recharge rivers and groundwater reserves. Given that water is a seasonally scarce resource in all savannahs, woody encroachment may threaten the ecosystem services related to water resources. This article is part of the themed issue ‘Tropical grassy biomes: linking ecology, human use and conservation’. PMID:27502378
Influence of stand density on stem quality in pole-size northern hardwoods.
Richard M. Godman; David J. Books
1971-01-01
Relates the type and frequency of limbs and limb-related defects in the first two logs of five hardwood species to residual basal area 15 years after initial cutting. Also discusses other tree characteristics influenced by stand density and the applicability of present silvicultural guides to improve stem quality.
John D. Shaw; James N. Long
2007-01-01
We developed a density management diagram (DMD) for longleaf pine (Pinus palustris P. Mill.) using data from Forest Inventory and Analysis plots. Selection criteria were for purity, defined as longleaf pine basal area (BA) that is 90% or more of plot BA, and even-agedness, as defined by a ratio between two calculations of stand density index. The...
Results after 20 years from a western larch levels-of-growing-stock study.
K.W. Seidel
1987-01-01
The 20-year growth response from a levels-of-growing-stock study in an even-aged western larch stand in eastern Oregon, first thinned at age 33, showed that trees growing at low stand densities grew more rapidly in diameter than trees in high-density plots. Height growth was relatively uniform among density levels. Both basal-area and total cubic-volume increment...
Natural seed fall in white pine (Pinus strobes L.) stands of varying density
Raymond E. Graber
1970-01-01
Seed fall was observed in three stands of mature white pines at stand basal-area densities of 80, 120, and 187 square feet per acre. It was found that the intermediate-density stand produced nearly 50 percent more seed than the stands of other densities. During a good seed year this stand produced 59 pounds of dry sound seed per acre. Most of the seeds were dispersed...
Cayuela, Luis; González-Caro, Sebastián; Aldana, Ana M.; Stevenson, Pablo R.; Phillips, Oliver; Cogollo, Álvaro; Peñuela, Maria C.; von Hildebrand, Patricio; Jiménez, Eliana; Melo, Omar; Londoño-Vega, Ana Catalina; Mendoza, Irina; Velásquez, Oswaldo; Fernández, Fernando; Serna, Marcela; Velázquez-Rua, Cesar; Benítez, Doris; Rey-Benayas, José M.
2017-01-01
Understanding and predicting the likely response of ecosystems to climate change are crucial challenges for ecology and for conservation biology. Nowhere is this challenge greater than in the tropics as these forests store more than half the total atmospheric carbon stock in their biomass. Biomass is determined by the balance between biomass inputs (i.e., growth) and outputs (mortality). We can expect therefore that conditions that favor high growth rates, such as abundant water supply, warmth, and nutrient-rich soils will tend to correlate with high biomass stocks. Our main objective is to describe the patterns of above ground biomass (AGB) stocks across major tropical forests across climatic gradients in Northwestern South America. We gathered data from 200 plots across the region, at elevations ranging between 0 to 3400 m. We estimated AGB based on allometric equations and values for stem density, basal area, and wood density weighted by basal area at the plot-level. We used two groups of climatic variables, namely mean annual temperature and actual evapotranspiration as surrogates of environmental energy, and annual precipitation, precipitation seasonality, and water availability as surrogates of water availability. We found that AGB is more closely related to water availability variables than to energy variables. In northwest South America, water availability influences carbon stocks principally by determining stand structure, i.e. basal area. When water deficits increase in tropical forests we can expect negative impact on biomass and hence carbon storage. PMID:28301482
Álvarez-Dávila, Esteban; Cayuela, Luis; González-Caro, Sebastián; Aldana, Ana M; Stevenson, Pablo R; Phillips, Oliver; Cogollo, Álvaro; Peñuela, Maria C; von Hildebrand, Patricio; Jiménez, Eliana; Melo, Omar; Londoño-Vega, Ana Catalina; Mendoza, Irina; Velásquez, Oswaldo; Fernández, Fernando; Serna, Marcela; Velázquez-Rua, Cesar; Benítez, Doris; Rey-Benayas, José M
2017-01-01
Understanding and predicting the likely response of ecosystems to climate change are crucial challenges for ecology and for conservation biology. Nowhere is this challenge greater than in the tropics as these forests store more than half the total atmospheric carbon stock in their biomass. Biomass is determined by the balance between biomass inputs (i.e., growth) and outputs (mortality). We can expect therefore that conditions that favor high growth rates, such as abundant water supply, warmth, and nutrient-rich soils will tend to correlate with high biomass stocks. Our main objective is to describe the patterns of above ground biomass (AGB) stocks across major tropical forests across climatic gradients in Northwestern South America. We gathered data from 200 plots across the region, at elevations ranging between 0 to 3400 m. We estimated AGB based on allometric equations and values for stem density, basal area, and wood density weighted by basal area at the plot-level. We used two groups of climatic variables, namely mean annual temperature and actual evapotranspiration as surrogates of environmental energy, and annual precipitation, precipitation seasonality, and water availability as surrogates of water availability. We found that AGB is more closely related to water availability variables than to energy variables. In northwest South America, water availability influences carbon stocks principally by determining stand structure, i.e. basal area. When water deficits increase in tropical forests we can expect negative impact on biomass and hence carbon storage.
The role of red alder in riparian forest structure along headwater streams in southeastern Alaska
Orlikowska, E.H.; Deal, R.L.; Hennon, P.E.; Wipfli, M.S.
2004-01-01
We assessed the influence of red alder on tree species composition, stand density, tree size distribution, tree mortality, and potential for producing large conifers, in 38-42 yr old riparian forests along 13 headwater streams in the Maybeso and Harris watersheds on Prince of Wales Island, Alaska. Red alder ranged from 0 to 53% of the total live basal area of the stands. Tree density, basal area of live and dead trees, and mean diameter of live conifers were not significantly related to the percent of alder as a proportion of total stand live basal area within these riparian forests. The mean diameter of the 100 largest conifers per hectare (the largest trees) was similar among different sites and appeared unrelated to the amount of alder in the stands. The mean diameter of dead conifers increased slightly with increasing proportion of red alder. Most dead trees were small and died standing. Red alder was much more concentrated immediately along stream margins (within 0-1 m distance from the stream bank vs. > 1 m). The presence of red alder did not inhibit the production of large-diameter conifers, and both alder and conifers provided small woody debris for fishless headwater streams in southeastern Alaska. Red alder is an important structural component of young-growth riparian stands.
Woody encroachment and its consequences on hydrological processes in the savannah.
Honda, Eliane A; Durigan, Giselda
2016-09-19
Woody encroachment due to changes in climate or in the disturbance regimes (fire and herbivory) has been observed throughout the savannah biome over the last century with ecological, hydrological and socioeconomic consequences. We assessed changes in tree density and basal area and estimated changes in rain interception by the canopies across a 5-year period over a biomass gradient in Cerrado vegetation protected from fire. We modelled throughfall, stemflow and net rainfall on the basis of tree basal area (TBA). Tree density increased by an average annual rate of 6.7%, basal area at 5.7% and rain interception by the canopies at 0.6% of the gross rainfall. Independent of the vegetation structure, we found a robust relationship of 0.9% less rainfall reaching the ground as TBA increases by 1 m(2) ha(-1) Increases in tree biomass with woody encroachment may potentially result in less water available for uptake by plants and to recharge rivers and groundwater reserves. Given that water is a seasonally scarce resource in all savannahs, woody encroachment may threaten the ecosystem services related to water resources.This article is part of the themed issue 'Tropical grassy biomes: linking ecology, human use and conservation'. © 2016 The Author(s).
John M. Kabrick; David R. Larsen; Stephen R. Shifley
1997-01-01
We conducted a study to identify pre-treatment trends in woody species density, diameter, and basal area among MOFEP sites, blocks, and treatment areas; relate woody species differences among sites, blocks, and treatment areas to differences in environmental conditions; and identify potential treatment response differences based upon our fmdings. Sites 2 through 5 had...
Andrew T. Hudak; Benjamin C. Bright; Scott M. Pokswinski; E. Louise Loudermilk; Joseph J. O' Brien; Benjamin S. Hornsby; Carine Klauberg; Carlos A. Silva
2016-01-01
Eglin Air Force Base (AFB) in Florida, in the United States, conserves a large reservoir of native longleaf pine (Pinus palustris Mill.) stands that land managers maintain by using frequent fires. We predicted tree density, basal area, and dominant tree species from 195 forest inventory plots, low-density airborne LiDAR, and Landsat data available across the entirety...
Does initial spacing influence crown and hydraulic architecture of Eucalyptus marginata?
Grigg, A H; Macfarlane, C; Evangelista, C; Eamus, D; Adams, M A
2008-05-01
Long-term declines in rainfall in south-western Australia have resulted in increased interest in the hydraulic characteristics of jarrah (Eucalyptus marginata Donn ex Smith) forest established in the region's drinking water catchments on rehabilitated bauxite mining sites. We hypothesized that in jarrah forest established on rehabilitated mine sites: (1) leaf area index (L) is independent of initial tree spacing; and (2) more densely planted trees have less leaf area for the same leaf mass, or the same sapwood area, and have denser sapwood. Initial stand densities ranged from about 600 to 9000 stems ha(-1), and trees were 18 years old at the time of sampling. Leaf area index was unaffected by initial stand density, except in the most sparsely stocked stands where L was 1.2 compared with 2.0-2.5 in stands at other spacings. The ratio of leaf area to sapwood area (A(l):A(s)) was unaffected by tree spacing or tree size and was 0.2 at 1.3 m height and 0.25 at the crown base. There were small increases in sapwood density and decreases in leaf specific area with increased spacing. Tree diameter or basal area was a better predictor of leaf area than sapwood area. At the stand scale, basal area was a good predictor of L (r(2) = 0.98, n = 15) except in the densest stands. We conclude that the hydraulic attributes of this forest type are largely independent of initial tree spacing, thus simplifying parameterization of stand and catchment water balance models.
Maximum growth potential in loblolly pine: results from a 47-year-old spacing study in Hawaii
Lisa J. Samuelson; Thomas L. Eberhardt; John R. Butnor; Tom A. Stokes; Kurt H. Johnsen
2010-01-01
Growth, allocation to woody root biomass, wood properties, leaf physiology, and shoot morphology were examined in a 47-year-old loblolly pine (Pinus taeda L.) density trial located in Maui, Hawaii, to determine if stands continued to carry the high density, basal area, and volume reported at younger ages and to identify potential factors controlling...
Required sample size for monitoring stand dynamics in strict forest reserves: a case study
Diego Van Den Meersschaut; Bart De Cuyper; Kris Vandekerkhove; Noel Lust
2000-01-01
Stand dynamics in European strict forest reserves are commonly monitored using inventory densities of 5 to 15 percent of the total surface. The assumption that these densities guarantee a representative image of certain parameters is critically analyzed in a case study for the parameters basal area and stem number. The required sample sizes for different accuracy and...
Individual tree growth response to variable-density thinning in coastal Pacific Northwest forests.
Scott D.s Roberts; Constance A. Harrington
2008-01-01
We examined 5-year basal area growth of nearly 2600 trees in stem-mapped plots at five locations differing in site characteristics, species composition, and management history on the Olympic Peninsula in Western Washington, USA. Our objectives were to determine if internal edges, the boundaries within the stand between components of the variable-density thinning,...
Forest plantations in the Midsouth, U.S.A.
James F. Rosson
1995-01-01
Timberland that has been artificially regenerated in the seven Midsouth States was analyzed by ownership, forest type, stocking class, age, tree density, basal area, site class, and volume. Growing-stock volumes of natural stands and plantations were compared.
Edward F. Loewenstein; Kenneth R. Davidson
2002-01-01
In 1998, a project was initiated to demonstrate techniques and evaluate the efficacy of reducing overstory tree density and reintroducing fire in order to develop the tree composition, structure, and herbaceous complex typical of a savanna. On three study areas, two dominated by oak and one by shortleaf pine, the total basal area of all trees = 1.6 inches DBH was...
NASA Astrophysics Data System (ADS)
Rack, Wolfgang; Haas, Christian; Langhorne, Pat J.
2013-11-01
We present airborne measurements to investigate the thickness of the western McMurdo Ice Shelf in the western Ross Sea, Antarctica. Because of basal accretion of marine ice and brine intrusions conventional radar systems are limited in detecting the ice thickness in this area. In November 2009, we used a helicopter-borne laser and electromagnetic induction sounder (EM bird) to measure several thickness and freeboard profiles across the ice shelf. The maximum electromagnetically detectable ice thickness was about 55 m. Assuming hydrostatic equilibrium, the simultaneous measurement of ice freeboard and thickness was used to derive bulk ice densities ranging from 800 to 975 kg m-3. Densities higher than those of pure ice can be largely explained by the abundance of sediments accumulated at the surface and present within the ice shelf, and are likely to a smaller extent related to the overestimation of ice thickness by the electromagnetic induction measurement related to the presence of a subice platelet layer. The equivalent thickness of debris at a density of 2800 kg m-3 is found to be up to about 2 m thick. A subice platelet layer below the ice shelf, similar to what is observed in front of the ice shelf below the sea ice, is likely to exist in areas of highest thickness. The thickness and density distribution reflects a picture of areas of basal freezing and supercooled Ice Shelf Water emerging from below the central ice shelf cavity into McMurdo Sound.
K.W. Seidel
1980-01-01
The 10-year growth of a 55-year-old, even-aged western larch (Larix occidentalis Nutt.) stand after it was thinned from above and below to a wide range of stocking levels was measured in eastern Oregon. Basal area and volume growth increased as stand density increased for both thinning methods. Despite heavy infestations of the larch casebearer...
Tara L. Keyser; Stanley J. Zarnoch
2014-01-01
Much about stump sprout dynamics of upland hardwood trees species has been obtained in clearcuts. Information on the response of stump sprouts to alternative silvicultural treatments, including treatments that manipulate stand density and stand structure is lacking. In this study we examined the influence of harvest season and levels of basal area reduction on the...
K.W. Seidel
1986-01-01
The 15-year growth response from a levels-of-growing-stock study in an even-aged western larch (Larix occidentalis Nutt.) stand, first thinned from above and below at age 55, was measured in northeastern Oregon. Basal area and volume growth increased with stand density for both thinning methods, whereas diameter growth decreased. Attacks of the...
Laser scanning in vivo confocal microscopy of the normal human corneoscleral limbus.
Patel, Dipika V; Sherwin, Trevor; McGhee, Charles N J
2006-07-01
To elucidate the structure of the human corneoscleral limbus by in vivo laser scanning confocal microscopy and to correlate limbal epithelial dimensions and density with the central epithelium and in relation to age. Fifty adult subjects were recruited into one of two age groups: younger (age<45 years) and older (age>or=45 years). Fifty left eyes of these 50 healthy subjects were examined by laser scanning in vivo confocal microscopy, to assess the basal epithelium of the central cornea and inferior limbus. Mean epithelial cell diameter, area, and density were calculated for the central basal epithelium, limbus-corneal basal epithelium, and limbus-palisade epithelium. Data were analyzed in relation to the two age groups, group A, 30+/-6 years (n=25; mean+/-SD), and group B, 60+/-11 years (n=25; P<0.01). Mean epithelial density in the limbus-cornea and limbus-palisade regions decreased significantly with age: limbus-cornea group A=7253+/-1077 cells/mm2 group B=6614+/-987 cells/mm2, P=0.03; limbus palisade group A=5409+/-799 cells/mm2, group B=5055+/-722 cells/mm2, P=0.03). Central corneal epithelial density did not change with age: group A=6162+/-503 cells/mm2, group B=6362+/-614 cells/mm2, P=0.08. Mean epithelial density was greatest at the limbus-cornea (7010+/-1081 cells/mm2) and lowest at the limbus-palisades (5289+/-847 cells/mm2). The mean width of palisade ridges was 25.0+/-6.3 microm. This is the first study to image clearly the living human corneal limbus by laser scanning in vivo confocal microscopy and to demonstrate quantitative changes in the basal epithelium with age.
Leaf-on canopy closure in broadleaf deciduous forests predicted during winter
Twedt, Daniel J.; Ayala, Andrea J.; Shickel, Madeline R.
2015-01-01
Forest canopy influences light transmittance, which in turn affects tree regeneration and survival, thereby having an impact on forest composition and habitat conditions for wildlife. Because leaf area is the primary impediment to light penetration, quantitative estimates of canopy closure are normally made during summer. Studies of forest structure and wildlife habitat that occur during winter, when deciduous trees have shed their leaves, may inaccurately estimate canopy closure. We estimated percent canopy closure during both summer (leaf-on) and winter (leaf-off) in broadleaf deciduous forests in Mississippi and Louisiana using gap light analysis of hemispherical photographs that were obtained during repeat visits to the same locations within bottomland and mesic upland hardwood forests and hardwood plantation forests. We used mixed-model linear regression to predict leaf-on canopy closure from measurements of leaf-off canopy closure, basal area, stem density, and tree height. Competing predictive models all included leaf-off canopy closure (relative importance = 0.93), whereas basal area and stem density, more traditional predictors of canopy closure, had relative model importance of ≤ 0.51.
Memiaghe, Hervé R; Lutz, James A; Korte, Lisa; Alonso, Alfonso; Kenfack, David
2016-01-01
Tropical forests have long been recognized for their biodiversity and ecosystem services. Despite their importance, tropical forests, and particularly those of central Africa, remain understudied. Until recently, most forest inventories in Central Africa have focused on trees ≥10 cm in diameter, even though several studies have shown that small-diameter tree population may be important to demographic rates and nutrient cycling. To determine the ecological importance of small-diameter trees in central African forests, we used data from a 25-ha permanent plot that we established in the rainforest of Gabon to study the diversity and dynamics of these forests. Within the plot, we censused 175,830 trees ≥1 cm dbh from 54 families, 192 genera, and 345 species. Average tree density was 7,026 trees/ha, basal area 31.64 m2/ha, and above-ground biomass 369.40 Mg/ha. Fabaceae, Ebenaceae and Euphorbiaceae were the most important families by basal area, density and above-ground biomass. Small-diameter trees (1 cm ≥ dbh <10 cm) comprised 93.7% of the total tree population, 16.5% of basal area, and 4.8% of the above-ground biomass. They also had diversity 18% higher at family level, 34% higher at genus level, and 42% higher at species level than trees ≥10 cm dbh. Although the relative contribution of small-diameter trees to biomass was comparable to other forests globally, their contribution to forest density, and diversity was disproportionately higher. The high levels of diversity within small-diameter classes may give these forests high levels of structural resilience to anthropogenic/natural disturbance and a changing climate.
Memiaghe, Hervé R.; Lutz, James A.; Korte, Lisa; Alonso, Alfonso; Kenfack, David
2016-01-01
Tropical forests have long been recognized for their biodiversity and ecosystem services. Despite their importance, tropical forests, and particularly those of central Africa, remain understudied. Until recently, most forest inventories in Central Africa have focused on trees ≥10 cm in diameter, even though several studies have shown that small-diameter tree population may be important to demographic rates and nutrient cycling. To determine the ecological importance of small-diameter trees in central African forests, we used data from a 25-ha permanent plot that we established in the rainforest of Gabon to study the diversity and dynamics of these forests. Within the plot, we censused 175,830 trees ≥1 cm dbh from 54 families, 192 genera, and 345 species. Average tree density was 7,026 trees/ha, basal area 31.64 m2/ha, and above-ground biomass 369.40 Mg/ha. Fabaceae, Ebenaceae and Euphorbiaceae were the most important families by basal area, density and above-ground biomass. Small-diameter trees (1 cm ≥ dbh <10 cm) comprised 93.7% of the total tree population, 16.5% of basal area, and 4.8% of the above-ground biomass. They also had diversity 18% higher at family level, 34% higher at genus level, and 42% higher at species level than trees ≥10 cm dbh. Although the relative contribution of small-diameter trees to biomass was comparable to other forests globally, their contribution to forest density, and diversity was disproportionately higher. The high levels of diversity within small-diameter classes may give these forests high levels of structural resilience to anthropogenic/natural disturbance and a changing climate. PMID:27186658
Mortality among Seed Trees in Longleaf Pine Shelterwood Stands
William D. Boyer
1970-01-01
Mortality of longieaf pine (Pinus palustris Mill.) seed trees was recorded in 27 regeneration areas ranging from North Carolina to Louisiana. Annual mortality averaged 0.7 percent before, and 1.9 percent after a seed cut reduced stand density to about 30 square feet of basal area per acre. On a per-acre basis, however, annual losses averaged 0....
Growth and yield of western larch: 15-year results of a levels-of-growing-stock study.
K.W. Seidel
1982-01-01
The 15-year growth response from a levels-of-growing-stock study in an even-aged western larch (Larix occidentalis Nutt.) stand in northeastern Oregon, first thinned at age 33, showed that trees growing at lower stand densities grew more rapidly in diameter but did not grow faster in height than trees in high density plots. Both basal area and...
Modeling abundance effects in distance sampling
Royle, J. Andrew; Dawson, D.K.; Bates, S.
2004-01-01
Distance-sampling methods are commonly used in studies of animal populations to estimate population density. A common objective of such studies is to evaluate the relationship between abundance or density and covariates that describe animal habitat or other environmental influences. However, little attention has been focused on methods of modeling abundance covariate effects in conventional distance-sampling models. In this paper we propose a distance-sampling model that accommodates covariate effects on abundance. The model is based on specification of the distance-sampling likelihood at the level of the sample unit in terms of local abundance (for each sampling unit). This model is augmented with a Poisson regression model for local abundance that is parameterized in terms of available covariates. Maximum-likelihood estimation of detection and density parameters is based on the integrated likelihood, wherein local abundance is removed from the likelihood by integration. We provide an example using avian point-transect data of Ovenbirds (Seiurus aurocapillus) collected using a distance-sampling protocol and two measures of habitat structure (understory cover and basal area of overstory trees). The model yields a sensible description (positive effect of understory cover, negative effect on basal area) of the relationship between habitat and Ovenbird density that can be used to evaluate the effects of habitat management on Ovenbird populations.
Quantitative analysis of American woodcock nest and brood habitat
Bourgeois, A.; Keppie, Daniel M.; Owen, Ray B.
1977-01-01
Sixteen nest and 19 brood sites of American woodcock (Philohela minoI) were examined in northern lower Michigan between 15 April and 15 June 1974 to determine habitat structure associated with these sites. Woodcock hens utilized young, second-growth forest stands which were similar in species composition for both nesting and brood rearing. A multi-varIate discriminant function analysis revealed a significant (P< 0.05) difference, however, in habitat structure. Nest habitat was characterized by lower tree density (2176 trees/ha) and basal area (8.6 m2/ha), by being close to forest openings (7 m) and by being situated on dry, relatively well drained sites. In contrast, woodcock broods were located in sites that had nearly twice the tree density (3934 trees/hal and basal area (16.5 m2/ha), was located over twice as far from forest openings (18 m) and generally occurred on damp sites, near (8 m) standing water. Importance of the habitat features to the species and possible management implications are discussed.
Surface and Basal Roughness in Radar Sounding Data: Obstacle and Opportunity
NASA Astrophysics Data System (ADS)
Schroeder, D. M.; Grima, C.; Haynes, M.
2015-12-01
The surface and basal roughness of glaciers, ice sheets, and ice shelves can pose a significant obstacle to the visual interpretation and quantitative analysis of radar sounding data. Areas of high surface roughness - including grounding zones, shear margins, and crevasse fields - can produce clutter and side-lobe signals that obscure the interpretation of englacial and subglacial features. These areas can also introduce significant variation in bed echo strength profiles as a result of losses from two-way propagation through rough ice surfaces. Similarly, reflections from rough basal interfaces beneath ice sheets and ice shelves can also result in large, spatially variable losses in bed echo power. If unmitigated and uncorrected, these effects can degrade or prevent the definitive interpretation of material and geometric properties at the base of ice sheets and ice shelves using radar reflectivity and bed echo character. However, these effects also provide geophysical signatures of surface and basal interface character - including surface roughness, firn density, subglacial bedform geometry, ice shelf basal roughness, marine-ice/brine detection, and crevasse geometry - that can be observed and constrained by exploiting roughness effects in radar sounding data. We present a series of applications and approaches for characterizing and correcting surface and basal roughness effects for airborne radar sounding data collected in Antarctica. We also present challenges, insights, and opportunities for extending these techniques to the orbital radar sounding of Europa's ice shell.
Ten-year growth of planted slash pine after thinnings
Hans G. Enghardt; W.F. Mann
1972-01-01
volume growth of slash pine between ages 17 and 27 years was directly related to residual basal area per acre after thinning. Diameter growth was inversely related to stand density, and very heavy cutting was required to attain a rate of 3 inches in 10 years.
Rodney E. Will; Thomas C. Hennessey; Thomas B. Lynch; Robert Heinemann; Randal Holeman; Dennis Wilson; Keith Anderson; Gregory Campbell
2013-01-01
We determined the effects of planting density (4- by 4-, 6- by 6-, 8- by 8-, and 10- by 10-foot spacing) on stand-level height, diameter at breast height, stem volume, basal area, and periodic annual increment for two loblolly pine (Pinus taeda L.) seed sources. Seed sources for the 25-year-old stands were a North Carolina seed source (NCC 8-01) and...
Fifteen-year results from six cutting methods in second-growth northern hardwoods.
Gayne G. Erdmann; Robert R. Oberg
1973-01-01
Presents and compares stand growth and yield information from three single-tree selection cuts, a crop-tree release treatment, an 8-inch diameter limit cut, and an uncut control. Discusses the influence of stand density on basal area growth, cubic volume growth, and board-foot volume growth.
Glendon W. Smalley; Robert L. Bailey
1974-01-01
Detailed schedules of trees per acre, basal area, mean tree height, and cubic-foot yields in eight volume categories by l-inch diameter classes are presented for all combinations of four site indexes, seven ages from seed, and nine planting densities.
Vegetation and floristics of a lowland tropical rainforest in northeast Australia
Apgaua, Deborah M. G.; Campbell, Mason J; Cox, Casey J; Crayn, Darren M; Ishida, Françoise Y; Laidlaw, Melinda J; Liddell, Michael J; Seager, Michael; Laurance, Susan G. W.
2016-01-01
Abstract Background Full floristic data, tree demography, and biomass estimates incorporating non-tree lifeforms are seldom collected and reported for forest plots in the tropics. Established research stations serve as important repositories of such biodiversity and ecological data. With a canopy crane setup within a tropical lowland rainforest estate, the 42-ha Daintree Rainforest Observatory (DRO) in Cape Tribulation, northern Australia is a research facility of international significance. We obtained an estimate of the vascular plant species richness for the site, by surveying all vascular plant species from various mature-phase, remnant and open vegetation patches within the site. We also integrate and report the demography and basal areas of trees ≥ 10 cm diameter at breast height (dbh) in a new 1-ha core plot, an extension to the pre-existing forest 1-ha plot under the canopy crane. In addition, we report for the canopy crane plot new demography and basal areas for smaller-size shrubs and treelets subsampled from nine 20 m2 quadrats, and liana basal area and abundance from the whole plot. The DRO site has an estimated total vascular plant species richness of 441 species, of which 172 species (39%) are endemic to Australia, and 4 species are endemics to the Daintree region. The 2 x 1-ha plots contains a total of 262 vascular plant species of which 116 (1531 individuals) are tree species ≥ 10 cm dbh. We estimate a stem basal area of 34.9 m2 ha-1, of which small stems (tree saplings and shrubs <10cm dbh) and lianas collectively contribute c.4.2%. Comparing the stem density-diversity patterns of the DRO forest with other tropical rainforests globally, our meta-analysis shows that DRO forests has a comparatively high stem density and moderate species diversity, due to the influence of cyclones. These data will provide an important foundation for ecological and conservation studies in lowland tropical forest. New information We present a floristic checklist, a lifeform breakdown, and demography data from two 1-ha rainforest plots from a lowland tropical rainforest study site. We also present a meta-analysis of stem densities and species diversity from comparable-sized plots across the tropics. PMID:27099552
Doolin, Paul F.; Birge, Wesley J.
1966-01-01
Ultrastructural studies were performed on normal and abnormal cilia and basal bodies associated with the choroidal epithelium of the chick embryo. Tissues were prepared in each of several fixatives including: 1% osmium tetroxide, in both phosphate and veronal acetate buffers; 2% glutaraldehyde, followed by postfixation in osmium tetroxide; 1% potassium permanganate in veronal acetate buffer. Normal cilia display the typical pattern of 9 peripheral doublets and 2 central fibers, as well as a system of 9 secondary fibers. The latter show distinct interconnections between peripheral and central fibers. Supernumerary fibers were found to occur in certain abnormal cilia. The basal body is complex, bearing 9 transitional fibers at the distal end and numerous cross-striated rootlets at the proximal end. The distal end of the basal body is delimited by a basal plate of moderate density. The tubular cylinder consists of 9 triple fibers. The C subfibers end at the basal plate, whereas subfibers A and B continue into the shaft of the cilium. The 9 transitional fibers radiate out from the distal end of the basal body, ending in bulblike terminal enlargements which are closely associated with the cell membrane in the area of the basal cup. One or 2 prominent basal feet project laterally from the basal body. These structures characteristically show several dense cross-bands and, on occasion, are found associated with microtubules. PMID:5335827
Response of old-growth conifers to reduction in stand density in western Oregon forests
Latham, P.; Tappeiner, J. C.
2002-01-01
The positive growth response of healthy young trees to density reduction is well known. In contrast, large old trees are usually thought to be intrinsically limited in their ability to respond to increased growing space; therefore, density reduction is seldom used in stands of old-growth trees. We tested the null hypothesis that old-growth trees are incapable of responding with increased growth following density reduction. The diameter growth response of 271 Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco), ponderosa pine (Pinus ponderosa Dougl. ex Laws) and sugar pine (Pinus lambertiana Dougl.) trees ranging in age from 158 to 650 years was examined 20 to 50 years after density reduction. Density reduction involved either light thinning with removal of less vigorous trees, or shelterwood treatments in which overstory trees were not removed. Ratios of basal area growth after treatment to basal area growth before treatment, and several other measures of growth, all indicated that the old trees sometimes benefited and were not harmed by density reduction. Growth increased by 10% or more for 68% of the trees in treated stands, and nearly 30% of trees increased growth by over 50%. This growth response persisted for at least 20 years. During this 20-year period, only three trees in treated stands (1.5%) exhibited a rapid decrease in growth, whereas growth decreased in 64% of trees in untreated stands. The length of time before a growth response to density reduction occurred varied from 5 to 25 years, with the greatest growth response often occurring 20 to 25 years after treatment. These results have important implications both for the basic biology of aging in woody plants as well as for silvicultural practices in forests with old-growth trees.
Allen L. Lundgren
1981-01-01
Describes an analysis of initial density and subsequent thinning options for red pine (Pinus resinosa Ait.) plantations in the Lake States. Results showed that the initial number of established trees per acres has a major impact on the amount and quality of timber product yields, with 200 trees per acre (500/ha) thinned to 120 square feet of basal area per acre (27.5...
Three-dimensional structure of basal body triplet revealed by electron cryo-tomography
Li, Sam; Fernandez, Jose-Jesus; Marshall, Wallace F; Agard, David A
2012-01-01
Basal bodies and centrioles play central roles in microtubule (MT)-organizing centres within many eukaryotes. They share a barrel-shaped cylindrical structure composed of nine MT triplet blades. Here, we report the structure of the basal body triplet at 33 Å resolution obtained by electron cryo-tomography and 3D subtomogram averaging. By fitting the atomic structure of tubulin into the EM density, we built a pseudo-atomic model of the tubulin protofilaments at the core of the triplet. The 3D density map reveals additional densities that represent non-tubulin proteins attached to the triplet, including a large inner circular structure in the basal body lumen, which functions as a scaffold to stabilize the entire basal body barrel. We found clear longitudinal structural variations along the basal body, suggesting a sequential and coordinated assembly mechanism. We propose a model in which δ-tubulin and other components participate in the assembly of the basal body. PMID:22157822
Habitat characteristics of American woodcock nest sites on a managed area in Maine
McAuley, D.G.; Longcore, J.R.; Sepik, G.F.; Pendleton, G.W.
1996-01-01
We measured characteristics of habitat near 89 nests of American woodcock (Scolopax minor) and 100 randomly selected points on Moosehorn National Wildlife Refuge, Calais, Maine, an area managed for woodcock. At nest sites, basal area was lower (P 0.05) or between sites of successful nests and nests destroyed by predators, although the large variances of the variables reduced our power to detect differences. Habitat around sites of renests differed from sites of first nests. Sites around first nests had lower basal area of dead trees (P = 0.05) and higher stem densities of aspen (P = 0.03) and cherry saplings (P = 0.001), and viburnum (P = 0.05), while renest sites had taller trees (P = 0.02). The change from nest sites in areas dominated by alders and tree-size gray birch used in 1977-80 to sites dominated by sapling trees, especially aspen, used during 1987-90 suggests that woodcock in the expanding population at the refuge are selecting nest sites created by habitat management since 1979.
Harvesting small trees for bio-energy
John Klepac; Robert Rummer; Jason Thompson
2011-01-01
A conventional whole-tree logging operation consisting of 4-wheeled and 3-wheeled saw-head feller-bunchers, two grapple skidders and a chipper that produces dirty chips was monitored across several stands and machine performance evaluated. Stands were inventoried to determine density, volume, and basal area per acre and will be used to relate machine performance to...
Wayne C. Zipperer
2002-01-01
Regenerated and remnant forest patches were inventoried in Syracuse, New York, USA to determine differences in structure, species composition, human disturbances, and landscape context. Patches had similar mean stem diameter, total stem density, and total basal areas, but differed with respect to diameter distribution, disturbance regime, landscape context, and...
Structure of Pine Stands in the Southeast
William A. Bechtold; Gregory A. Ruark
1988-01-01
Distributional and statistical information associated with stand age, site index, basal area per acre, number of stems per acre, and stand density index is reported for major pine cover types of the Southeastern United States. Means, standard deviations, and ranges of these variables are listed by State and physiographic region for loblolly, slash, longleaf, pond,...
Growth response of managed uneven-aged northern conifer stands
Dale S. Solomon; Robert M. Frank
1983-01-01
The growth response of trees in spruce-fir-hemlock stands was recorded from plots that were managed to control stand density, species composition, length of harvest interval, and salvage of mortality. Basal area, volume, and diameter increment are presented by species and size classification for harvesting intervals of 5, 10, and 20 years.
Lisa A. Schulte; David J. Mladenoff; Erik V. Nordheim
2002-01-01
We developed a quantitative and replicable classification system to improve understanding of historical composition and structure within northern Wisconsin's forests. The classification system was based on statistical cluster analysis and two forest metrics, relative dominance (% basal area) and relative importance (mean of relative dominance and relative density...
A generalized system of models forecasting Central States tree growth.
Stephen R. Shifley
1987-01-01
Describes the development and testing of a system of individual tree-based growth projection models applicable to species in Indiana, Missouri, and Ohio. Annual tree basal area growth is estimated as a function of tree size, crown ratio, stand density, and site index. Models are compatible with the STEMS and TWIGS Projection System.
Michael D. Cain; Michael G. Shelton
1994-01-01
In 1992 we analyzed the composition of a 32-ha pine-hardwood forest that originated from the partial cutting of the existing virgin forest around 1915. The area has been reserved from timber management since 1935. Pines >9 cm in diameter at a height of 1.37 m accounted for 61% of overstory and midstory basal area but only 21% of density. Of those trees that had...
Breeding birds in managed forests on public conservation lands in the Mississippi Alluvial Valley
Twedt, Daniel J.; Wilson, R. Randy
2017-01-01
Managers of public conservation lands in the Mississippi Alluvial Valley have implemented forest management strategies to improve bottomland hardwood habitat for target wildlife species. Through implementation of various silvicultural practices, forest managers have sought to attain forest structural conditions (e.g., canopy cover, basal area, etc.) within values postulated to benefit wildlife. We evaluated data from point count surveys of breeding birds on 180 silviculturally treated stands (1049 counts) that ranged from 1 to 20 years post-treatment and 134 control stands (676 counts) that had not been harvested for >20 years. Birds detected during 10-min counts were recorded within four distance classes and three time intervals. Avian diversity was greater on treated stands than on unharvested stands. Of 42 commonly detected species, six species including Prothonotary Warbler (Prothonotaria citrea) and Acadian Flycatcher (Empidonax virescens) were indicative of control stands. Similarly, six species including Indigo Bunting (Passerina cyanea) and Yellow-breasted Chat (Icteria virens) were indicative of treated stands. Using a removal model to assess probability of detection, we evaluated occupancy of bottomland forests at two spatial scales (stands and points within occupied stands). Wildlife-forestry treatment improved predictive models of species occupancy for 18 species. We found years post treatment (range = 1–20), total basal area, and overstory canopy were important species-specific predictors of occupancy, whereas variability in basal area was not. In addition, we used a removal model to estimate species-specific probability of availability for detection, and a distance model to estimate effective detection radius. We used these two estimated parameters to derive species densities and 95% confidence intervals for treated and unharvested stands. Avian densities differed between treated and control stands for 16 species, but only Common Yellowthroat (Geothlypis trichas) and Yellow-breasted Chat had greater densities on treated stands.
Evaluation of Sentinel-1A Data For Above Ground Biomass Estimation in Different Forests in India
NASA Technical Reports Server (NTRS)
Vadrevu, Krishna Prasad
2017-01-01
Use of remote sensing data for mapping and monitoring of forest biomass across large spatial scales can aid in addressing uncertainties in carbon cycle. Earlier, several researchers reported on the use of Synthetic Aperture Radar (SAR) data for characterizing forest structural parameters and the above ground biomass estimation. However, these studies cannot be generalized and the algorithms cannot be applied to all types of forests without additional information on the forest physiognomy, stand structure and biomass characteristics. The radar backscatter signal also saturates as forest parameters such as biomass and the tree height increase. It is also not clear how different polarizations (VV versus VH) impact the backscatter retrievals in different forested regions. Thus, it is important to evaluate the potential of SAR data in different landscapes for characterizing forest structural parameters. In this study, the SAR data from Sentinel-1A has been used to characterize forest structural parameters including the above ground biomass from tropical forests of India. Ground based data on tree density, basal area and above ground biomass data from thirty-eight different forested sites has been collected to relate to SAR data. After the pre-processing of Sentinel 1-A data for radiometric calibration, geo-correction, terrain correction and speckle filtering, the variability in the backscatter signal in relation tree density, basal area and above biomass density has been investigated. Results from the curve fitting approach suggested exponential model between the Sentinel-1A backscatter versus tree density and above ground biomass whereas the relationship was almost linear with the basal area in the VV polarization mode. Of the different parameters, tree density could explain most of the variations in backscatter. Both VV and VH backscatter signals could explain only thirty and thirty three percent of variation in above biomass in different forest sites of India. Results also suggested saturation of the Sentinel-1A backscatter signal around hundred tonnes per hectare for VV polarization and one hundred and forty five tonnes per hectare for VH polarization. The presentation will highlight the above results in addition to potentials and limitations of Sentinel-1A data for retrieving forest structural parameters. Also, background information on different forest types of India, biomass variations and forest type mapping efforts in the region will be presented.
Western spruce budworm as related to stand characteristics in the bitterroot national forest
Carroll B. Williams; Patrick J. Shea; Gerald S. Walton
1971-01-01
Relation of population density to certain stand conditions and damage indicators was analyzed in four drainages on the Bitterroot National Forest of Montana. Western spruce budworm (Choristoneura occidentalis Freeman) populations were strongly related to plot basal area, tree species, and tree crown levels, and also to current and past levels of tree defoliation....
Jianwei Zhang; Martin W. Ritchie; Douglas A. Maguire; William W. Oliver
2013-01-01
We analyzed 45-yr data collected from three ponderosa pine (Pinus ponderosa Lawson & C. Lawson) levels-of-growing-stock installations in Oregon (OR) and northern California (CA), USA, to determine the effect of stand density regimes on stand productivity and mortality. We found that periodic annual increment (PAI) of diameter, basal area (BA...
Nearest neighbor imputation of species-level, plot-scale forest structure attributes from LiDAR data
Andrew T. Hudak; Nicholas L. Crookston; Jeffrey S. Evans; David E. Hall; Michael J. Falkowski
2008-01-01
Meaningful relationships between forest structure attributes measured in representative field plots on the ground and remotely sensed data measured comprehensively across the same forested landscape facilitate the production of maps of forest attributes such as basal area (BA) and tree density (TD). Because imputation methods can efficiently predict multiple response...
Forty years later at Taylor Woods: Merging the old and new
John D. Bailey
2008-01-01
The Taylor Woods "Levels-of-Growing-Stock" study was established in 1962 to create a replicated ponderosa pine density experiment for the Southwest, making a valuable addition to research in the Fort Valley Experimental Forest. Basal area treatments ranged from 5-20 m2/ha (19-80 ft2/ac) when installed, designed as...
A frontier shortleaf pine stand in the old-growth Cross Timbers of Oklahoma
K. Chris Cerny; David W. Stahle; Don C. Bragg
2016-01-01
We investigated an old-growth oak-shortleaf pine (Pinus echinata) stand of high ecological integrity in east-central Oklahoma located west of the continuous native distribution of shortleaf pine. With the exception of an abundance of shortleaf pine, the basal area (17.2 m2/ha), density (559.6 trees/ha), and species...
Dinamica de mediano plazo del bosque nuboso de Volcan Mombacho, Nicaragua
Fabricio Diaz Santos; Peter L. Weaver; Fabio Gabriel Diaz Santos
2010-01-01
The structure and dynamics of the cloud foreston Mombacho Volcano were evaluated 11 years after the establishment of six permanent plots (PPMs). The data collected during this period showed the following trends: stem density per hectare had been decreasing, basal area increasing, recruitment staying constant, and mortality diminishing on the most of the PPMs. Although...
Stand dynamics of mixed red alder-conifer forests of southeast Alaska.
Robert L. Deal; Paul E. Hennon; Ewa H. Orlikowska; David V. D' Amore
2004-01-01
Stand structure and dynamics were evaluated in mixed red alder (Alnus rubra Bong.) - conifer forests of southeast Alaska. We assessed stand development, tree density, total basal area, diameter distribution of live and dead trees, height distribution of live trees, and mean diameter of all and largest conifers in 40-year-old red alder - conifer...
Changing stand structure and regional growth reductions in Georgia's natural pine stands
W.A. Bechtold; G.A. Ruark; F.T. Lloyd
1991-01-01
Forest Inventory and Analysis (FIA) data indicate reductions in the growth of naturally regenerated pines in Georgia between the two latest measurement periods (1961-1972 vs. 1972-1982). Analysis of Covariance was used to adjust stand-level basal area growth rates for differences between periods in stand age, stand density, site index, mortality, and hardwood...
The isolated red spruce communities of Virginia and West Virginia
Harold S. Adams; Steven Stephenson; Adam W. Rollins; Mary Beth Adams
2010-01-01
Quantitative data on the composition and structure of coniferous forests containing red spruce in the mountains of central and southwestern Virginia and eastern central West Virginia, based on sampling carried out in 67 stands during the 1982 to 1984 field seasons, are provided. The average importance value ([relative basal area + relative density/2]) of red spruce was...
Partial Denervation of Subbasal Axons Persists Following Debridement Wounds to the Mouse Cornea
Pajoohesh-Ganji, Ahdeah; Pal-Ghosh, Sonali; Tadvalkar, Gauri; Kyne, Briana M.; Saban, Daniel R.; Stepp, Mary Ann
2015-01-01
Although sensory reinnervation occurs after injury in the PNS, poor reinnervation in the elderly and those with diabetes often leads to pathology. Here we quantify subbasal axon density in the central and peripheral mouse cornea over time after three different types of injury. The mouse cornea is highly innervated with a dense array of subbasal nerves that form a spiral called the vortex at the corneal center or apex; these nerves are readily detected within flat mounted corneas. After anesthesia, corneal epithelial cells were removed using either a dulled blade or a rotating burr within an area demarcated centrally with a 1.5 mm trephine. A third wound type, superficial trephination, involved demarcating the area with the 1.5 mm trephine but not removing cells. By 7d after superficial trephination, subbasal axon density returns to control levels; by 28d the vortex reforms. Although axon density is similar to control 14d after dulled blade and rotating burr wounding, defects in axon morphology at the corneal apex remain. After 14d, axons retract from the center leaving the subbasal axon density reduced by 37.2% and 36.8% at 28d after dulled blade and rotating burr wounding, respectively, compared to control. Assessment of inflammation using flow cytometry shows that persistent inflammation is not a factor in the incomplete reinnervation. Expression of mRNAs encoding 22 regeneration associated genes (RAGs) involved in axon targeting assessed by QPCR reveals that netrin-1 and ephrin signaling are altered after wounding. Subpopulations of corneal epithelial basal cells at the corneal apex stop expressing ki67 as early as 7d after injury and by 14d and 28d after wounding, many of these basal cells undergo apoptosis and die. While subbasal axons are restored to their normal density and morphology after superficial trephination, subbasal axon recovery is partial after debridement wounds. The increase in corneal epithelial basal cell apoptosis at the apex observed at 14d after corneal debridement may destabilize newly reinnervated subbasal axons and lead to their retraction towards the periphery. PMID:26280222
Post-fire regeneration in seasonally dry tropical forest fragments in southeastern Brazil.
Costa, Mayke B; Menezes, Luis Fernando T DE; Nascimento, Marcelo T
2017-01-01
Seasonally dry tropical forest is one of the highly threatened biome. However, studies on the effect of fire on these tree communities are still scarce. In this context, a floristic and structural survey in three forest areas in the southeast of Brazil that were affected by fire between 14 and 25 years ago was performed with the objective of evaluating post-fire regeneration. In each site, five systematically placed plots (25 m x 25 m each) were established. The more recently burnt site had significantly lower values of richness and diversity than the other two sites. However, the sites did not differ in density and basal area. Annona dolabripetala, Astronium concinnum, Joannesia princeps and Polyandrococos caudescens were within the 10 most important species for the three sites. Comparing these data with adjacent mature forests, the results indicated differences both in structural and floristic aspects, suggesting that the time after fire was not sufficient for recuperation of these areas. The recovery process indicate at least 190 years for areas return to basal area values close to those observed in mature forests nearby.
Crop-tree release thinning in 65-year-old commercial cherry-maple stands (5-year results)
H. Clay Smith; Gary W. Miller; Neil I. Lamson
1994-01-01
Crop-tree release was applied to a 65-year-old cherry-maple stand in north central West Virginia. Criteria were developed for selecting crop trees for high quality sawtimber and veneer products. Five-year stand growth, mortality, and ingrowth using basal areas, volume, relative density, and number of trees were discussed for the treatments.
Forest biomass and tree planting for fossil fuel offsets in the Colorado Front Range
Mike A. Battaglia; Kellen Nelson; Dan Kashian; Michael G. Ryan
2010-01-01
This study estimates the amount of carbon available for removal in fuel reduction and reforestation treatments in montane forests of the Colorado Front Range based on site productivity, pre-treatment basal area, and planting density. Thinning dense stands will yield the greatest offsets for biomass fuel. However, this will also yield the greatest carbon losses, if the...
Potential fire behavior is reduced following forest restoration treatments
Peter Z. Fule; Charles McHugh; Thomas A. Heinlein; W. Wallace Covington
2001-01-01
Potential fire behavior was compared under dry, windy weather conditions in 12 ponderosa pine stands treated with alternative thinning prescriptions in the wildland/urban interface of Flagstaff, Arizona. Prior to thinning, stands averaged 474 trees/ acre, 158 ft2/acre basal area, crown bulk density 0.0045 lb/ft3, and crown base height 19.2 ft. Three thinning treatments...
Dynamic programming for optimization of timber production and grazing in ponderosa pine
Kurt H. Riitters; J. Douglas Brodie; David W. Hann
1982-01-01
Dynamic programming procedures are presented for optimizing thinning and rotation of even-aged ponderosa pine by using the four descriptors: age, basal area, number of trees, and time since thinning. Because both timber yield and grazing yield are functions of stand density, the two outputs-forage and timber-can both be optimized. The soil expectation values for single...
Characteristics of Five Climax Stands in New Hampshire
W.B. Leak; W.B. Leak
1987-01-01
Analysis of species composition, stand density, and diameter distribution in five climax or old-growth stands in New Hampshire indicates that the northern hardwood climax is characterized by at least 65 to 70 percent tolerant hardwoods, 130 square feet basal area per acre, and "q" ratios (2-inch diameter classes) of 1.3 to 1.4. Climax sprucehemlock have at...
Christine E. Hura; Thomas R. Crow
2004-01-01
We examined the effects of management on coarse woody debris, both standing and downed, in thinned and unthinned northern hardwood forests in upper Michigan. The unthinned conditions included old growth and second growth, while the thinned conditions included both even- and uneven-aged management. The structural features analyzed were stem diameter, density, basal area...
Thinning increases growth of 60-year-old cherry-maple stands in West Virginia
Neil I. Lamson; Neil I. Lamson
1985-01-01
In north-central West Virginia, previously unmanaged 60-year-old cherrymaple stands were thinned to 60 percent relative stand density. Thinning reduced mortality, redistributed growth onto fewer, larger stems, and increased individual tree growth. Five-year periodic basal-area growth per acre was 1.2 times greater in thinned stands than in unthinned stands. Periodic...
Thinning cherry-maple stands in West Virginia: 5-year results
Neil I. Lamson; H. Clay. Smith; H. Clay. Smith
1988-01-01
In northern West Virginia, 60-year-old cherry-maple stands were thinned to 75,60, and 45 percent relative stand density. Analysis of 5-year growth data showed that basal-area growth was not reduced by thinning. Cubic-foot and board-foot volume growth decreased slightly. Individual-tree growth of all trees, dominant/codominant trees, and the 50 largest diameter trees...
Liana abundance, diversity, and distribution on Barro Colorado Island, Panama.
Schnitzer, Stefan A; Mangan, Scott A; Dalling, James W; Baldeck, Claire A; Hubbell, Stephen P; Ledo, Alicia; Muller-Landau, Helene; Tobin, Michael F; Aguilar, Salomon; Brassfield, David; Hernandez, Andres; Lao, Suzanne; Perez, Rolando; Valdes, Oldemar; Yorke, Suzanne Rutishauser
2012-01-01
Lianas are a key component of tropical forests; however, most surveys are too small to accurately quantify liana community composition, diversity, abundance, and spatial distribution - critical components for measuring the contribution of lianas to forest processes. In 2007, we tagged, mapped, measured the diameter, and identified all lianas ≥1 cm rooted in a 50-ha plot on Barro Colorado Island, Panama (BCI). We calculated liana density, basal area, and species richness for both independently rooted lianas and all rooted liana stems (genets plus clones). We compared spatial aggregation patterns of liana and tree species, and among liana species that varied in the amount of clonal reproduction. We also tested whether liana and tree densities have increased on BCI compared to surveys conducted 30-years earlier. This study represents the most comprehensive spatially contiguous sampling of lianas ever conducted and, over the 50 ha area, we found 67,447 rooted liana stems comprising 162 species. Rooted lianas composed nearly 25% of the woody stems (trees and lianas), 35% of woody species richness, and 3% of woody basal area. Lianas were spatially aggregated within the 50-ha plot and the liana species with the highest proportion of clonal stems more spatially aggregated than the least clonal species, possibly indicating clonal stem recruitment following canopy disturbance. Over the past 30 years, liana density increased by 75% for stems ≥1 cm diameter and nearly 140% for stems ≥5 cm diameter, while tree density on BCI decreased 11.5%; a finding consistent with other neotropical forests. Our data confirm that lianas contribute substantially to tropical forest stem density and diversity, they have highly clumped distributions that appear to be driven by clonal stem recruitment into treefall gaps, and they are increasing relative to trees, thus indicating that lianas will play a greater role in the future dynamics of BCI and other neotropical forests.
NASA Astrophysics Data System (ADS)
Hayashi, Shohei; Yamashita, Tamotsu; Senzaki, Junji; Miyazato, Masaki; Ryo, Mina; Miyajima, Masaaki; Kato, Tomohisa; Yonezawa, Yoshiyuki; Kojima, Kazutoshi; Okumura, Hajime
2018-04-01
The origin of expanded single Shockley-type stacking faults in forward-current degradation of 4H-SiC p-i-n diodes was investigated by the stress-current test. At a stress-current density lower than 25 A cm-2, triangular stacking faults were formed from basal-plane dislocations in the epitaxial layer. At a stress-current density higher than 350 A cm-2, both triangular and long-zone-shaped stacking faults were formed from basal-plane dislocations that converted into threading edge dislocations near the interface between the epitaxial layer and the substrate. In addition, the conversion depth of basal-plane dislocations that expanded into the stacking fault was inside the substrate deeper than the interface. These results indicate that the conversion depth of basal-plane dislocations strongly affects the threshold stress-current density at which the expansion of stacking faults occurs.
Carlisle, L; Steel, K; Forge, A
1990-11-01
Deafness in the viable dominant spotting mouse mutant is due to a primary defect of the stria vascularis which results in absence of the positive endocochlear potential in scala media. Endocochlear potentials were measured and the structure of stria vascularis of mutants with potentials close to zero was compared with that in normal littermate controls by use of morphometric methods. The stria vascularis was significantly thinner in mutants. Marginal cells were not significantly different from controls in terms of volume density or intramembrane particle density but the network density of tight junctions was significantly reduced in the mutants. A virtual absence of gap junctions between basal cells and marginal or intermediate cells was observed, but intramembrane particle density and junctional complexes between adjacent basal cells were not different from controls. The volume density of basal cells was significantly greater in mutants. Intermediate cells accounted for a significantly smaller volume density of the stria vascularis in mutants and had a lower density of intramembrane particles than controls. Melanocytes were not identified in the stria vascularis of mutants. These results suggest that communication between marginal, intermediate and basal cells might be important to the normal function of the stria vascularis.
Analysis of forest structure using thematic mapper simulator data
NASA Technical Reports Server (NTRS)
Peterson, D. L.; Westman, W. E.; Brass, J. A.; Stephenson, N. J.; Ambrosia, V. G.; Spanner, M. A.
1986-01-01
The potential of Thematic Mapper Simulator (TMS) data for sensing forest structure information has been explored by principal components and feature selection techniques. In a survey of forest structural properties conducted for 123 field sites of the Sequoia National Park, the canopy closure could be well estimated (r = 0.62 to 0.69) by a variety of channel bands and band ratios, without reference to the forest type. Estimation of the basal area was less successful (r = 0.51 or less) on the average, but could be improved for certain forest types when data were stratified by floristic composition. To achieve such a stratification, individual sites were ordinated by a detrended correspondence analysis based on the canopy of dominant species. The analysis of forest structure in the Sequoia data suggests that total basal area can be best predicted in stands of lower density, and in younger even-aged managed stands.
Successional dynamics in Neotropical forests are as uncertain as they are predictable
Norden, Natalia; Angarita, Héctor A.; Bongers, Frans; Martínez-Ramos, Miguel; Granzow-de la Cerda, Iñigo; van Breugel, Michiel; Lebrija-Trejos, Edwin; Meave, Jorge A.; Vandermeer, John; Williamson, G. Bruce; Finegan, Bryan; Mesquita, Rita; Chazdon, Robin L.
2015-01-01
Although forest succession has traditionally been approached as a deterministic process, successional trajectories of vegetation change vary widely, even among nearby stands with similar environmental conditions and disturbance histories. Here, we provide the first attempt, to our knowledge, to quantify predictability and uncertainty during succession based on the most extensive long-term datasets ever assembled for Neotropical forests. We develop a novel approach that integrates deterministic and stochastic components into different candidate models describing the dynamical interactions among three widely used and interrelated forest attributes—stem density, basal area, and species density. Within each of the seven study sites, successional trajectories were highly idiosyncratic, even when controlling for prior land use, environment, and initial conditions in these attributes. Plot factors were far more important than stand age in explaining successional trajectories. For each site, the best-fit model was able to capture the complete set of time series in certain attributes only when both the deterministic and stochastic components were set to similar magnitudes. Surprisingly, predictability of stem density, basal area, and species density did not show consistent trends across attributes, study sites, or land use history, and was independent of plot size and time series length. The model developed here represents the best approach, to date, for characterizing autogenic successional dynamics and demonstrates the low predictability of successional trajectories. These high levels of uncertainty suggest that the impacts of allogenic factors on rates of change during tropical forest succession are far more pervasive than previously thought, challenging the way ecologists view and investigate forest regeneration. PMID:26080411
Successional dynamics in Neotropical forests are as uncertain as they are predictable.
Norden, Natalia; Angarita, Héctor A; Bongers, Frans; Martínez-Ramos, Miguel; Granzow-de la Cerda, Iñigo; van Breugel, Michiel; Lebrija-Trejos, Edwin; Meave, Jorge A; Vandermeer, John; Williamson, G Bruce; Finegan, Bryan; Mesquita, Rita; Chazdon, Robin L
2015-06-30
Although forest succession has traditionally been approached as a deterministic process, successional trajectories of vegetation change vary widely, even among nearby stands with similar environmental conditions and disturbance histories. Here, we provide the first attempt, to our knowledge, to quantify predictability and uncertainty during succession based on the most extensive long-term datasets ever assembled for Neotropical forests. We develop a novel approach that integrates deterministic and stochastic components into different candidate models describing the dynamical interactions among three widely used and interrelated forest attributes--stem density, basal area, and species density. Within each of the seven study sites, successional trajectories were highly idiosyncratic, even when controlling for prior land use, environment, and initial conditions in these attributes. Plot factors were far more important than stand age in explaining successional trajectories. For each site, the best-fit model was able to capture the complete set of time series in certain attributes only when both the deterministic and stochastic components were set to similar magnitudes. Surprisingly, predictability of stem density, basal area, and species density did not show consistent trends across attributes, study sites, or land use history, and was independent of plot size and time series length. The model developed here represents the best approach, to date, for characterizing autogenic successional dynamics and demonstrates the low predictability of successional trajectories. These high levels of uncertainty suggest that the impacts of allogenic factors on rates of change during tropical forest succession are far more pervasive than previously thought, challenging the way ecologists view and investigate forest regeneration.
Naupaka Zimmerman; Flint 1 Hughes; Patrick Hart; Heather Kalei Chang; David Perez; Ryan Kaipoalohaakala Like; Rebecca Ostertag
2008-01-01
The majority of Hawaii's lowland wet forests no longer exist, with many of the last remaining patches found on the eastern, windward sides of the largest islands. To better understand successional patterns and invasion in these native systems, we quantified basal area (BA) and densities of woody species and understory cover at nine sites in the Puna district on...
John Hogland; Nathaniel Anderson; Joseph St. Peter; Jason Drake; Paul Medley
2018-01-01
Accurate information is important for effective management of natural resources. In the field of forestry, field measurements of forest characteristics such as species composition, basal area, and stand density are used to inform and evaluate management activities. Quantifying these metrics accurately across large landscapes in a meaningful way is extremely important...
Changes in tree species diversity after successive clearcuts in the Southern Appalachians
Katherine J. Elliott; Wayne T. Swank
1994-01-01
A 16 ha watershed in the Coweeta Basin was clearcut in 1939 and 1962. Vegetation was inventoried in 1934 and at about 7-year intervals to 1991. After the first clearcut, tree diversity remained high until after the second cut. Diversity based on density and basal area decreased significantly 14 years after the second clearcut and remained low through 1991. Diversity...
Site factors influencing oak decline in the interior highlands of Arkansas, Missouri, and Oklahoma
Edward A. Poole; Eric Heitzman; James M. Guldin
2006-01-01
Oak decline is affecting the forests in the Interior Highlands of Arkansas, Missouri, and Oklahoma. In 2002 and 2003, field plots were established throughout the region to evaluate the influence of topographic position and aspect on oak decline. Density and basal area of dead and dying oaks did not significantly differ by either topographic position or aspect. Lack of...
Composition and structure of an old-growth floodplain forest of the lower Kaskaskia River
John B. Taft
2003-01-01
Compositional and structural properties of canopy, shrub/sapling, and ground-cover strata were measured within an old-growth floodplain forest bordering the lower Kaskaskia River in southwestern Illinois. Basal area for trees was estimated at 31.8 m²/ha, tree density was 398 trees/ha with 27 species recorded in the canopy stratum. The dominant tree species...
Andrew T. Hudak; Jeffrey S. Evans; Michael J. Falkowski; Nicholas L. Crookston; Paul E. Gessler; Penelope Morgan; Alistair M. S. Smith
2005-01-01
Multispectral satellite imagery are appealing for their relatively low cost, and have demonstrated utility at the landscape level, but are typically limited at the stand level by coarse resolution and insensitivity to variation in vertical canopy structure. In contrast, lidar data are less affected by these difficulties, and provide high structural detail, but are less...
J.D. McIver; R. Ottmar
2006-01-01
Stand structure and downed woody fuel mass were measured in four replicate units for each of three treatments (unlogged control, commercial harvest, and fuel reduction harvest) following the 1996 Summit Wildfire in northeastern Oregon. Commercial and fuel-reduction harvest resulted in a significant decrease in tree density and tree basal areas. The total downed woody...
Tejpal, C S; Sumitha, E B; Pal, A K; Shivananda Murthy, H; Sahu, N P; Siddaiah, G M
2014-04-01
A 60 day feeding trial was conducted to study the effect of dietary l-tryptophan on thermal tolerance and oxygen consumption rate of freshwater fish, mrigala, Cirrhinus mrigala reared under ambient temperature at low and high stocking density. Four hundred eighty fingerlings were distributed into eight experimental groups. Four groups each of low density group (10 fishes/75L water) and higher density group (30 fishes/75L water) were fed a diet containing 0, 0.68, 1.36 or 2.72% l-tryptophan in the diet, thus forming eight experimental groups namely, Low density control (LC) (basal feed +0% l-tryptophan); LT1 (basal feed+0.68% l-tryptophan); LT2 (basal feed+1.36% l-tryptophan); LT3 (basal feed+2.72% l-tryptophan); high density control (HC) (basal feed+0% l-tryptophan); HT1 (basal feed+0.68% l-tryptophan); HT2 (basal feed+1.36% l-tryptophan); and HT3 (basal feed+2.72% l-tryptophan) were fed at 3% of the body weight. The test diets having crude protein 34.33±0.23 to 35.81±0.18% and lipid 423.49±1.76 to 425.85±0.31KCal/100g were prepared using purified ingredients. The possible role of dietary l-tryptophan on thermal tolerance and oxygen consumption rate was assessed in terms of critical thermal maxima (CTMax), critical thermal minima (CTMin), lethal thermal maxima (LTMax) and lethal thermal minima (LTMin). The CTMax, CTMin, LTMax and LTMin values were found to be significantly higher (p<0.05) in the treatment groups with CTMax 42.94±0.037 (LT2); LT Max 43.18±0.070 (LT2); CTMin 10.47±0.088 (LT2) and LTMin 9.42±0.062 (LT3), whereas the control group showed a lower tolerance level. The same trend was observed in the high density group (CTMax 42.09±0.066 (LT3); LTMax 43 23±0.067 (HT3); CTMin 10.98±0.040 (HT3) and LTMin 9.74±0.037 (HT3). However, gradual supplementation of dietary l-tryptophan in the diet significantly reduced the oxygen consumption rate in both the low density group (Y=-26.74x+222.4, r²=0.915) and the high density group (Y=-32.96x+296.5, r²=0.8923). Dietary supplementation of l-tryptophan at a level of 1.36% improved the thermal tolerance level and reduced the oxygen consumption rate in C. mrigala fingerlings. Copyright © 2014 Elsevier Ltd. All rights reserved.
Shekhar, Abhishek; Das, Samiran; Bhattacharyya, Jayanta; Goel, Preeti; Majumdar, Sayan; Ghosh, Soumitra
2018-01-01
This study aims to find the effect of change in different salivary factors before and after complete denture insertion and to measure the maxillary denture retention in different arch forms. Thirty completely edentulous individuals (10 each of square, tapered, and ovoid arch form of maxilla) belonging to the age group of 40-70 years were selected. Salivary factors (flow, density, pH, viscosity, and total protein) were evaluated before and after denture insertion. Retention of maxillary denture was measured in all the different arch forms. Student's independent sample's t -test was applied. The correlation was analyzed by Pearson's correlation analysis. While mean flow rate and pH of saliva increased, mean viscosity, total protein, and density of saliva decreased after maxillary complete denture insertion. A positive correlation was found between retention and total maxillary basal surface area. Retention value was found to be greatest in square type and least in tapered type. Complete denture acts as a mechanical stimulant thus increasing flow rate and pH immediately after complete denture insertion. Density, total protein, and viscosity of saliva decreased after complete denture insertion which may be due to increase in water content of saliva. The retention of maxillary complete denture does not seem to depend on the rate of change of the salivary factors, before and after complete denture insertion. Total basal surface area and maxillary denture retention values were highest in square arch form and least in tapered arch form.
Analysis of ecological thresholds in a temperate forest undergoing dieback
Newton, Adrian C.; Cantarello, Elena; Evans, Paul M.
2017-01-01
Positive feedbacks in drivers of degradation can cause threshold responses in natural ecosystems. Though threshold responses have received much attention in studies of aquatic ecosystems, they have been neglected in terrestrial systems, such as forests, where the long time-scales required for monitoring have impeded research. In this study we explored the role of positive feedbacks in a temperate forest that has been monitored for 50 years and is undergoing dieback, largely as a result of death of the canopy dominant species (Fagus sylvatica, beech). Statistical analyses showed strong non-linear losses in basal area for some plots, while others showed relatively gradual change. Beech seedling density was positively related to canopy openness, but a similar relationship was not observed for saplings, suggesting a feedback whereby mortality in areas with high canopy openness was elevated. We combined this observation with empirical data on size- and growth-mediated mortality of trees to produce an individual-based model of forest dynamics. We used this model to simulate changes in the structure of the forest over 100 years under scenarios with different juvenile and mature mortality probabilities, as well as a positive feedback between seedling and mature tree mortality. This model produced declines in forest basal area when critical juvenile and mature mortality probabilities were exceeded. Feedbacks in juvenile mortality caused a greater reduction in basal area relative to scenarios with no feedback. Non-linear, concave declines of basal area occurred only when mature tree mortality was 3–5 times higher than rates observed in the field. Our results indicate that the longevity of trees may help to buffer forests against environmental change and that the maintenance of old, large trees may aid the resilience of forest stands. In addition, our work suggests that dieback of forests may be avoidable providing pressures on mature and juvenile trees do not pass critical thresholds. PMID:29240842
Multiple browsers structure tree recruitment in logged temperate forests
Faison, Edward K.; DeStefano, Stephen; Foster, David R.; Rapp, Joshua M.; Compton, Justin A.
2016-01-01
Historical extirpations have resulted in depauperate large herbivore assemblages in many northern forests. In eastern North America, most forests are inhabited by a single wild ungulate species, white-tailed deer (Odocoileus virginianus), and relationships between deer densities and impacts on forest regeneration are correspondingly well documented. Recent recolonizations by moose (Alces americanus) in northeastern regions complicate established deer density thresholds and predictions of browsing impacts on forest dynamics because size and foraging differences between the two animals suggest a lack of functional redundancy. We asked to what extent low densities of deer + moose would structure forest communities differently from that of low densities of deer in recently logged patch cuts of Massachusetts, USA. In each site, a randomized block with three treatment levels of large herbivores–no-ungulates (full exclosure), deer (partial exclosure), and deer + moose (control) was established. After 6–7 years, deer + moose reduced stem densities and basal area by 2-3-fold, Prunus pensylvanica and Quercus spp. recruitment by 3–6 fold, and species richness by 1.7 species (19%). In contrast, in the partial exclosures, deer had non-significant effects on stem density, basal area, and species composition, but significantly reduced species richness by 2.5 species on average (28%). Deer browsing in the partial exclosure was more selective than deer + moose browsing together, perhaps contributing to the decline in species richness in the former treatment and the lack of additional decline in the latter. Moose used the control plots at roughly the same frequency as deer (as determined by remote camera traps), suggesting that the much larger moose was the dominant browser species in terms of animal biomass in these cuts. A lack of functional redundancy with respect to foraging behavior between sympatric large herbivores may explain combined browsing effects that were both large and complex.
Hall, S. A.; Burke, I.C.; Box, D. O.; Kaufmann, M. R.; Stoker, Jason M.
2005-01-01
The ponderosa pine forests of the Colorado Front Range, USA, have historically been subjected to wildfires. Recent large burns have increased public interest in fire behavior and effects, and scientific interest in the carbon consequences of wildfires. Remote sensing techniques can provide spatially explicit estimates of stand structural characteristics. Some of these characteristics can be used as inputs to fire behavior models, increasing our understanding of the effect of fuels on fire behavior. Others provide estimates of carbon stocks, allowing us to quantify the carbon consequences of fire. Our objective was to use discrete-return lidar to estimate such variables, including stand height, total aboveground biomass, foliage biomass, basal area, tree density, canopy base height and canopy bulk density. We developed 39 metrics from the lidar data, and used them in limited combinations in regression models, which we fit to field estimates of the stand structural variables. We used an information–theoretic approach to select the best model for each variable, and to select the subset of lidar metrics with most predictive potential. Observed versus predicted values of stand structure variables were highly correlated, with r2 ranging from 57% to 87%. The most parsimonious linear models for the biomass structure variables, based on a restricted dataset, explained between 35% and 58% of the observed variability. Our results provide us with useful estimates of stand height, total aboveground biomass, foliage biomass and basal area. There is promise for using this sensor to estimate tree density, canopy base height and canopy bulk density, though more research is needed to generate robust relationships. We selected 14 lidar metrics that showed the most potential as predictors of stand structure. We suggest that the focus of future lidar studies should broaden to include low density forests, particularly systems where the vertical structure of the canopy is important, such as fire prone forests.
David G. Ray
2013-01-01
Restoring natural fire regimes and diverse ground cover to planted or old-field origin southern pine stands typically requires a substantial reduction in overstory density. While maintaining full canopy cover (CC) is consistent with maximizing fiber production, this approach does not allow sufficient light to reach the forest floor to accomplish a broader set of...
Long-term development of regeneration under longleaf pine seedtree and shelterwood stands
William D. Boyer
1993-01-01
Well-stocked mature longleaf pine (Pinus palustris Mill.) stands were cut tofive residual basal areas in 1957, namely 9,18,2 7.36, and 45 ft2 per ac, to observe the effect of stand density on seed production and seedling establishment. Seedlings, mainly from the 195.5 or 1961 seed crops, were established in treated stands. All pines on net 0.9 ac...
Justin L. Pszwaro; Anthony W. D' Amato; Thomas E. Burk; Matthew B. Russell; Brian J. Palik; Terry F. Strong
2016-01-01
Red maple (Acer rubrum L.), historically a common but not abundant tree species in North America, has increased in abundance throughout its range over the last several decades; however, it has received little attention in growth and yield studies. The objectives of this study were to (i) evaluate the effects of stocking level and stand density on...
Changes in a Missouri Ozark oak-hickory forest during 40 years of uneven-aged management
Edward F. Loewenstein; Harold E. Garrett; John P. Dwyer
1995-01-01
Changes in basal area, density and average diameter were examined on a 156,000-acre privately owned oak-hickory forest in the Missouri Ozarks. The forest has been managed since 1954 using the single-tree selection method. Trees greater than five inches dbh were monitored on 486 one-fifth-acre permanent plots at five year intervals from 1962 through 1992. Seven species/...
Andrew R. Neill; Klaus J. Puettmann; Adrian Ares
2013-01-01
To understand the impacts of overstory density reductions on resilience of forest ecosystems (i.e., the capacity of an ecosystem to maintain desired ecosystem functions in a fl uctuating environment), we examined overstory basal area and understory vegetation cover and richness collected 6 years after thinning in seven 40- to 60-year-old forests dominated by Douglas-fi...
Forty years later at Taylor Woods: Merging the old and new (P-53)
John D. Bailey
2008-01-01
The Taylor Woods "Levels-of-Growing-Stock" study was established in 1962 to create a replicated ponderosa pine density experiment for the Southwest, making a valuable addition to research in the Fort Valley Experimental Forest. Basal area treatments ranged from 5-20 m2/ha (19-80 ft2/ac) when installed, designed as growing stock levels 30/40, 60, 80, 100, 120...
Growth and yield for a 7-year-old yellow-poplar plantation in northern West Virginia
John R. Brooks
2013-01-01
Results for several major stand level variables from a 7-year-old yellow-poplar (Liriodendron tulipifera L.) plantation established in a converted pasture in northern West Virginia were summarized based on initial planting densities of 1,517 trees/ac, 972 trees/ac, 765 trees/ac, and 602 trees/ac. Stand basal area/acre at age 7 was greatest (54.7 ft...
James M. Guldin; Thomas Foti
2004-01-01
Abstract - Baseline tree data were collected in four watersheds in the eastern Ouachita Mountains during 1996-98. By watershed, average basal area ranged from 71 to 102 square feet per acre, average tree density ranged from 234 to 295 trees per acre, and quadratic mean diameter of trees ranged from 7.41 to 8.22 inches. Variables for which the largest...
K.W. Seidel; J. Michael Geist; Gerald S. Strickler
1990-01-01
Natural regeneration was abundant, regardless of grazing and grass seeding treatments,after shelterwood cutting to three overstory densities (27, 73, and 119 square feet of basal area per acre) in mixed conifer stands in the Starkey Experimental Forest and Range in eastern Oregon. After 6 years, the number of tree seedlings ranged from about 3,800 per acre on the low-...
Jianwei Zhang; William W. Oliver; Martin W. Ritchie; Donald L. Neal
2013-01-01
We periodically measured overstory ponderosa pine (Pinus ponderosa) growth and understory cover and abundance in a long-term study on the west slope of the Sierra Nevada, California, USA. The study was established in 1969 in a 20-year-old plantation, thinned to basal areas of 9, 16, 23, 30, and 37 m2 ha-1...
Spatial and Working Memory Is Linked to Spine Density and Mushroom Spines
Aher, Yogesh D.; Sase, Ajinkya; Gröger, Marion; Mokhtar, Maher; Höger, Harald; Lubec, Gert
2015-01-01
Background Changes in synaptic structure and efficacy including dendritic spine number and morphology have been shown to underlie neuronal activity and size. Moreover, the shapes of individual dendritic spines were proposed to correlate with their capacity for structural change. Spine numbers and morphology were reported to parallel memory formation in the rat using a water maze but, so far, there is no information on spine counts or shape in the radial arm maze (RAM), a frequently used paradigm for the evaluation of complex memory formation in the rodent. Methods 24 male Sprague-Dawley rats were divided into three groups, 8 were trained, 8 remained untrained in the RAM and 8 rats served as cage controls. Dendritic spine numbers and individual spine forms were counted in CA1, CA3 areas and dentate gyrus of hippocampus using a DIL dye method with subsequent quantification by the Neuronstudio software and the image J program. Results Working memory errors (WME) and latency in the RAM were decreased along the training period indicating that animals performed the task. Total spine density was significantly increased following training in the RAM as compared to untrained rats and cage controls. The number of mushroom spines was significantly increased in the trained as compared to untrained and cage controls. Negative significant correlations between spine density and WME were observed in CA1 basal dendrites and in CA3 apical and basal dendrites. In addition, there was a significant negative correlation between spine density and latency in CA3 basal dendrites. Conclusion The study shows that spine numbers are significantly increased in the trained group, an observation that may suggest the use of this method representing a morphological parameter for memory formation studies in the RAM. Herein, correlations between WME and latency in the RAM and spine density revealed a link between spine numbers and performance in the RAM. PMID:26469788
Tree species and size structure of old-growth Douglas-fir forests in central western Oregon, USA
Poage, Nathan; Tappeiner, J. C.
2005-01-01
We characterized the structure of 91 old-growth forests dominated by Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco), using inventory data from recent (1985a??1991) old-growth timber sales in western Oregon. The data were complete counts (i.e., censuses) of all live trees >20 cm diameter at breast height (dbh, measured at 1.4 m above the ground) over a mean area of 17.1 ha at each site. Across all sites, Douglas-fir accounted for 79% of the total basal area (m2/ha) of all species. The average density of trees >100 cm dbh was 19 trees/ha and 90% of these trees were Douglas-fir. Species other than Douglas-fir constituted only about 20% of the total basal area at each old-growth site, on average, but largely accounted for the structural variation between sites. We used multivariate techniques such as cluster analysis, indicator species analysis, and ordination with non-metric multidimensional scaling (NMS) to identify and characterize six structural groups in terms of basal area in different speciesa??diameter classes. Almost 97% of the structural information was captured by the first (r2 = 0.841) and second (r2 = 0.128) NMS ordination axis. Geographic information systems (GIS) analysis and NMS indicated that the structural differences among groups of sites were associated with moisture, temperature, and elevation gradients within the study area. This type of analysis can be used to help define differences among old-growth forests and to set local structural goals for growing forests with old-growth characteristics.
Urban climate modifies tree growth in Berlin
NASA Astrophysics Data System (ADS)
Dahlhausen, Jens; Rötzer, Thomas; Biber, Peter; Uhl, Enno; Pretzsch, Hans
2017-12-01
Climate, e.g., air temperature and precipitation, differs strongly between urban and peripheral areas, which causes diverse life conditions for trees. In order to compare tree growth, we sampled in total 252 small-leaved lime trees (Tilia cordata Mill) in the city of Berlin along a gradient from the city center to the surroundings. By means of increment cores, we are able to trace back their growth for the last 50 to 100 years. A general growth trend can be shown by comparing recent basal area growth with estimates from extrapolating a growth function that had been fitted with growth data from earlier years. Estimating a linear model, we show that air temperature and precipitation significantly influence tree growth within the last 20 years. Under consideration of housing density, the results reveal that higher air temperature and less precipitation led to higher growth rates in high-dense areas, but not in low-dense areas. In addition, our data reveal a significantly higher variance of the ring width index in areas with medium housing density compared to low housing density, but no temporal trend. Transferring the results to forest stands, climate change is expected to lead to higher tree growth rates.
Urban climate modifies tree growth in Berlin
NASA Astrophysics Data System (ADS)
Dahlhausen, Jens; Rötzer, Thomas; Biber, Peter; Uhl, Enno; Pretzsch, Hans
2018-05-01
Climate, e.g., air temperature and precipitation, differs strongly between urban and peripheral areas, which causes diverse life conditions for trees. In order to compare tree growth, we sampled in total 252 small-leaved lime trees ( Tilia cordata Mill) in the city of Berlin along a gradient from the city center to the surroundings. By means of increment cores, we are able to trace back their growth for the last 50 to 100 years. A general growth trend can be shown by comparing recent basal area growth with estimates from extrapolating a growth function that had been fitted with growth data from earlier years. Estimating a linear model, we show that air temperature and precipitation significantly influence tree growth within the last 20 years. Under consideration of housing density, the results reveal that higher air temperature and less precipitation led to higher growth rates in high-dense areas, but not in low-dense areas. In addition, our data reveal a significantly higher variance of the ring width index in areas with medium housing density compared to low housing density, but no temporal trend. Transferring the results to forest stands, climate change is expected to lead to higher tree growth rates.
Urban climate modifies tree growth in Berlin.
Dahlhausen, Jens; Rötzer, Thomas; Biber, Peter; Uhl, Enno; Pretzsch, Hans
2018-05-01
Climate, e.g., air temperature and precipitation, differs strongly between urban and peripheral areas, which causes diverse life conditions for trees. In order to compare tree growth, we sampled in total 252 small-leaved lime trees (Tilia cordata Mill) in the city of Berlin along a gradient from the city center to the surroundings. By means of increment cores, we are able to trace back their growth for the last 50 to 100 years. A general growth trend can be shown by comparing recent basal area growth with estimates from extrapolating a growth function that had been fitted with growth data from earlier years. Estimating a linear model, we show that air temperature and precipitation significantly influence tree growth within the last 20 years. Under consideration of housing density, the results reveal that higher air temperature and less precipitation led to higher growth rates in high-dense areas, but not in low-dense areas. In addition, our data reveal a significantly higher variance of the ring width index in areas with medium housing density compared to low housing density, but no temporal trend. Transferring the results to forest stands, climate change is expected to lead to higher tree growth rates.
Selective logging: does the imprint remain on tree structure and composition after 45 years?
Osazuwa-Peters, Oyomoare L; Chapman, Colin A; Zanne, Amy E
2015-01-01
Selective logging of tropical forests is increasing in extent and intensity. The duration over which impacts of selective logging persist, however, remains an unresolved question, particularly for African forests. Here, we investigate the extent to which a past selective logging event continues to leave its imprint on different components of an East African forest 45 years later. We inventoried 2358 stems ≥10 cm in diameter in 26 plots (200 m × 10 m) within a 5.2 ha area in Kibale National Park, Uganda, in logged and unlogged forest. In these surveys, we characterized the forest light environment, taxonomic composition, functional trait composition using three traits (wood density, maximum height and maximum diameter) and forest structure based on three measures (stem density, total basal area and total above-ground biomass). In comparison to unlogged forests, selectively logged forest plots in Kibale National Park on average had higher light levels, different structure characterized by lower stem density, lower total basal area and lower above-ground biomass, and a distinct taxonomic composition driven primarily by changes in the relative abundance of species. Conversely, selectively logged forest plots were like unlogged plots in functional composition, having similar community-weighted mean values for wood density, maximum height and maximum diameter. This similarity in functional composition irrespective of logging history may be due to functional recovery of logged forest or background changes in functional attributes of unlogged forest. Despite the passage of 45 years, the legacy of selective logging on the tree community in Kibale National Park is still evident, as indicated by distinct taxonomic and structural composition and reduced carbon storage in logged forest compared with unlogged forest. The effects of selective logging are exerted via influences on tree demography rather than functional trait composition.
Selective logging: does the imprint remain on tree structure and composition after 45 years?
Osazuwa-Peters, Oyomoare L.; Chapman, Colin A.; Zanne, Amy E.
2015-01-01
Selective logging of tropical forests is increasing in extent and intensity. The duration over which impacts of selective logging persist, however, remains an unresolved question, particularly for African forests. Here, we investigate the extent to which a past selective logging event continues to leave its imprint on different components of an East African forest 45 years later. We inventoried 2358 stems ≥10 cm in diameter in 26 plots (200 m × 10 m) within a 5.2 ha area in Kibale National Park, Uganda, in logged and unlogged forest. In these surveys, we characterized the forest light environment, taxonomic composition, functional trait composition using three traits (wood density, maximum height and maximum diameter) and forest structure based on three measures (stem density, total basal area and total above-ground biomass). In comparison to unlogged forests, selectively logged forest plots in Kibale National Park on average had higher light levels, different structure characterized by lower stem density, lower total basal area and lower above-ground biomass, and a distinct taxonomic composition driven primarily by changes in the relative abundance of species. Conversely, selectively logged forest plots were like unlogged plots in functional composition, having similar community-weighted mean values for wood density, maximum height and maximum diameter. This similarity in functional composition irrespective of logging history may be due to functional recovery of logged forest or background changes in functional attributes of unlogged forest. Despite the passage of 45 years, the legacy of selective logging on the tree community in Kibale National Park is still evident, as indicated by distinct taxonomic and structural composition and reduced carbon storage in logged forest compared with unlogged forest. The effects of selective logging are exerted via influences on tree demography rather than functional trait composition. PMID:27293697
NASA Astrophysics Data System (ADS)
Van Liefferinge, Brice; Pattyn, Frank; Cavitte, Marie G. P.; Young, Duncan A.; Roberts, Jason L.
2017-04-01
The quest for oldest ice in Antarctica has recently been launched through an EU H2020 project (Beyond EPICA - Oldest Ice) and aims at identifying suitable areas for a potential future drilling. Retrieving an ice core of such age is essential to understand the relation between orbital changes and atmospheric composition during the mid-Pliocene transition. However, sites for a potential undisturbed record of 1.5 million-year old ice in Antarctica are difficult to find and require slow-moving ice (preferably an ice divide) and basal conditions that are not disturbed by large topographic variations. Furthermore, ice should be sufficiently thick but cold basal conditions should still prevail, since basal melting would destroy the bottom layers. Therefore, ice-flow conditions and thermodynamic characteristics are crucial for identifying potential locations of undisturbed ice. Van Liefferinge and Pattyn (2013) identified suitable areas based on a pan-Antarctic simplified thermodynamic ice sheet model and demonstrated that uncertainty in geothermal conditions remain a major unknown. In order to refine these estimates, and provide uncertainties, we employ a full thermo-mechanically coupled higher-order ice sheet model (Pattyn, 2003; Pattyn et al., 2004). Initial conditions for the calculations are based on an inversion of basal slipperiness, based on observed surface topography (Pollard and DeConto, 2012; Pattyn, in prep.). Uncertainties in geothermal conditions are introduced using the convolution of two Gaussian probability density functions: (a) the reconstruction of the Antarctic ice sheet geometry and testing ice thickness variability over the last 2 million years (Pollard and DeConto, 2009) and (b) the surface temperature reconstruction over the same period (Snyder et al., 2016). The standard deviation, the skewness and the kurtosis of the whole Antarctic ice sheet are analyzed to observe likely probable melt conditions. Finally, we focus on model results in the divide area between Dome Concordia and Dome Fuji, and compare to newly acquired radar data in the region (OIA survey).
Behzad, A R; Chu, F; Walker, D C
1996-05-01
Previous findings have shown that pulmonary fibroblasts are associated with preexisting holes in the endothelial and epithelial basal laminae through which neutrophils appear to enter and leave the interstitium as they migrate from capillaries to alveoli. To determine their role in neutrophil migration, fibroblast organization within the interstitium was assessed by transmission electron microscope observations of serial-sectioned rabbit lung tissue. Interstitial fibroblasts were found to physically interconnect the endothelial basal lamina holes to epithelial basal lamina holes. Morphometric assessment of rabbit lung tissue instilled with Streptococcus pneumoniae revealed that approximately 70% of the surface area density of migrating neutrophils is in close contact (15 nm or less) with interstitial fibroblasts and extracellular matrix elements (30 and 40%, respectively). Although migrating neutrophils were close enough to adhere to both fibroblasts and extracellular elements, the interstitial fibroblasts are organized in a manner that would allow them to provide directional information to the neutrophils. A model illustrating this process is proposed.
Attenuated frontal and sensory inputs to the basal ganglia in cannabis users.
Blanco-Hinojo, Laura; Pujol, Jesus; Harrison, Ben J; Macià, Dídac; Batalla, Albert; Nogué, Santiago; Torrens, Marta; Farré, Magí; Deus, Joan; Martín-Santos, Rocío
2017-07-01
Heavy cannabis use is associated with reduced motivation. The basal ganglia, central in the motivation system, have the brain's highest cannabinoid receptor density. The frontal lobe is functionally coupled to the basal ganglia via segregated frontal-subcortical circuits conveying information from internal, self-generated activity. The basal ganglia, however, receive additional influence from the sensory system to further modulate purposeful behaviors according to the context. We postulated that cannabis use would impact functional connectivity between the basal ganglia and both internal (frontal cortex) and external (sensory cortices) sources of influence. Resting-state functional connectivity was measured in 28 chronic cannabis users and 29 controls. Selected behavioral tests included reaction time, verbal fluency and exposition to affective pictures. Assessments were repeated after one month of abstinence. Cannabis exposure was associated with (1) attenuation of the positive correlation between the striatum and areas pertaining to the 'limbic' frontal-basal ganglia circuit, and (2) attenuation of the negative correlation between the striatum and the fusiform gyrus, which is critical in recognizing significant visual features. Connectivity alterations were associated with lower arousal in response to affective pictures. Functional connectivity changes had a tendency to normalize after abstinence. The results overall indicate that frontal and sensory inputs to the basal ganglia are attenuated after chronic exposure to cannabis. This effect is consistent with the common behavioral consequences of chronic cannabis use concerning diminished responsiveness to both internal and external motivation signals. Such an impairment of the fine-tuning in the motivation system notably reverts after abstinence. © 2016 Society for the Study of Addiction.
Zhu, Xiang-cheng; Zhang, Zhen-ping; Zhang, Jun; Deng, Ai-xing; Zhang, Wei-jian
2016-02-01
The traditional rice growing practice has to change to save resource and protect environment, and it' s necessary to develop new technology in rice cultivation. Therefore, a two-year field experiment of Japonica rice (Liaoxing 1) was conducted in Northeast China in 2012 and 2013 to investigate the integrated effects of dense planting with less basal nitrogen (N) and unchanged top-dressing N (IR) on rice yield, N use efficiency (NUE) and greenhouse gas emissions. Compared with traditional practice (CK), we increased the rice seedling density by 33.3% and reduced the basal N rate by 20%. The results showed that the average N agronomy efficiency and partial factor productivity were improved by 49.6% (P<0.05) and 20.4% (P<0.05), respectively, while the area and yield-scaled greenhouse gas emissions were reduced by 9.9% and 12.7% (P<0.05), respectively. Although IR cropping mode decreased panicle number and biomass production, it significantly enhanced rice seed setting rate and harvest index, resulting in an unchanged or even highei yield. NH4+-N and NO3(-)-N concentrations in rice rhizosphere soil were reduced, resulting in an increment of N recovery efficiency. Generally, proper dense planting with less basal N applicatior could be a good approach for the trade-off between rice yield, NUE and greenhouse gas emission.
1983-10-01
sites should be collected and correlated to vegetational response through ordination or factor analysis techniques. 3. If site-specific methods are...specified number of samples is collected . If the 40 ."l ’ - - - .. -’-. - : -. .~ -, .4 .- . -.- , - ...... rb. . o* . -.% o - .- * .-.- v- . J...only, although the data collected were sufficient for determination of such parameters as density, cover, frequency, basal area, and importance value
Plume dispersion in four pine thinning scenarios: development of a simple pheromone dispersion model
Holly Peterson; Harold Thistle; Brian Lamb; Gene Allwine; Steve Edburg; Brian Strom
2010-01-01
A unique field campaign was conducted in 2004 to examine how changes in stand density may affect dispersion of insect pheromones in forest canopies. Over a l4-day period, 126 tracer tests were performed, and conditions ranged from an unthinned loblolly pine (Pinus taeda) canopy through a series of thinning scenarios with basal areas of32.l, 23.0, and 16.1 m2ha-l.ln...
R.G. Ribe
2009-01-01
Tensions between amenity- and timber-based economies in the U.S. and Canadian Pacific Northwest motivated a study of scenic beauty inside mature forests and timber harvests. A diverse sample of regional forests, measures of forest structure, and large, representative samples of photographs and public judges were employed to measure scenic beauty inside unharvested...
Wang, Zheng Q.; Newton, M.; Tappeiner, J. C.
1995-01-01
A large area of Pacific Coast forests is characterized by shallow soil, with negligible rainfall in the growing season. This study explores water-seeking strategy on such a site. We studied availability of bedrock water and its effects on growth and ecophysiology of 11-yr-old planted Douglas-fir (Pseudotsuga menziesii [Mirb.] Franco) and sprouting Pacific madrone (Arbutus menziesii Pursh). The study was carried out at three regulated densities of madrone sprouts on shallow ( 0.05) among three densities of madrone. However, cover in plots with the highest density of madrone (1322 sprout clumps/ha) depleted 50 mm of water from the 1.5 m layer by June, whereas vegetation on lower density treatments withdrew 15-28 mm by June, with later withdrawal distributed more uniformly through the growing season. Madrone density significantly affected basal diameter (P a?? 0.0001) and height growth (P a?? 0.002) of Douglas-fir. Madrone was consistently taller than Douglas-fir in all plots. The height of 11-yr-old madrone sprout clumps (424-465 cm) did not differ significantly among densities. Madrone leaf area index and biomass were higher at the high density of madrone than at medium density (P a?? 0.045, LAI; P a?? 0.001, biomass). Physiological advantages and rooting habits of madrone give it a competitive advantage over Douglas-fir in this area that it might not have if bedrock did not provide the principal water reservoir for summer growth.
Basal area growth, carbon isotope discrimination, and intrinsic ...
Many hectares of intensively managed Douglas-fir (Pseudotsuga menziesii Mirb. Franco) stands in western North America are fertilized with nitrogen to increase growth rates. Understanding the mechanisms of response facilitates prioritization of stands for treatment. The objective of this study was to test the hypothesis that the short-term basal area growth response to a single application of 224 kg N ha-1 as urea was associated with reduced stable carbon isotope discrimination (∆13C) and increased intrinsic water use efficiency (iWUE) in a 20-yr-old plantation of Douglas-fir in the Oregon Coast Range, USA. Increment cores were measured to estimate earlywood, latewood, and total basal area increment over a time series from 1997 to 2015. Stable carbon isotope discrimination and iWUE were estimated using earlywood and latewood stable carbon isotope concentrations in tree-ring holocellulose starting seven years before fertilization in early 2009 and ending seven years after treatment. A highly significant interaction effect between fertilization treatment and year was found for total basal area growth and earlywood basal area increment. Fertilized trees showed significant total basal area growth and earlywood basal area increment in the first (2009) and second (2010) growing seasons after fertilization in 2009. A marginally significant fertilization effect was found for latewood basal area increment only in the first growing season after treatment. A significant i
Excavation of red squirrel middens by grizzly bears in the whitebark pine zone
Mattson, D.J.; Reinhart, Daniel P.
1997-01-01
Whitebark pine seeds Pinus albicaulis are an important food of grizzly Ursus arctos horribilis bears wherever whitebark pine is abundant in the contiguous United States of America; availability of seeds affects the distribution of bears, and the level of conflict between bears and humans. Almost all of the seeds consumed by bears are excavated from middens where red squirrels Tamiasciurus hudsonicus have cached whitebark pine cones.Relationships among the occupancy of middens by squirrels, the excavation of middens by bears, and site features were investigated in this study. Data were collected from radio-marked bears and from middens located from line transects on two study sites in the Yellowstone ecosystem.Densities of active middens were positively related to lodgepole pine Pinus contorta basal area and negatively related to steepness of slope.The probability that a midden was occupied by a squirrel (i.e. active) was positively related to lodgepole pine basal area in the surrounding stand, size of the midden and size of the whitebark pine cone crop, and negatively related to elevation and to bear excavation during the previous 2-12 months.The probability that a midden had been excavated by a bear during the previous 12 months was positively related to size of the midden, and to whitebark pine basal area and cone crop, and negatively related to nearness of roads and town sites.The influence of midden size on bear use was attributable to a positive relationship with the number of excavated cones. The positive association between bear excavations and whitebark pine basal area or cone crops was attributable to availability of pine seeds.Grizzly bears would benefit from the minimization of roads and other human facilities in the whitebark pine zone and from increases in the availability of whitebark pine seeds, potentially achieved by increasing the numbers of cone-producing whitebark pine trees, especially in lower elevations of the whitebark pine zone where red squirrels are more abundant.
Prado-Junior, Jamir; de Oliveira-Neto, Norberto Emídio; Santana, Lucas Dezidério; do Vale, Vagner Santiago; Jacobson, Tamiel Baiocchi; de Oliveira, Paulo Eugênio Alves Macedo; Carvalho, Fabrício Alvim
2018-01-01
Understanding the relationships between Coffea arabica L. and the native tree community of secondary forests regrowing after the abandonment of coffee plantations is important because, as a non-native species in the Neotropics, coffee can outcompete native species, reducing diversity and forests ecosystem services. We aimed to answer three questions: 1) Does coffee regeneration in secondary forests differ between shaded and unshaded abandoned plantations?; 2) How is coffee basal area related to structural attributes, species diversity and composition of the native community?; and 3) Do the relationships between coffee and native community differ between tree and sapling components? We sampled the tree and sapling components in a seasonal tropical dry forest that were previously used as shaded and unshaded coffee plantations. Coffee was the most important species in the sapling component of shaded systems, but was almost absent in unshaded ones. Coffee basal area was negatively related with the native density and absolute species richness of the sapling component; and was negatively related with tree density, and positively related with the percentage of pioneer individuals of the native tree component. Our results indicate that coffee persists in secondary forest communities even after more than 70 years of shaded-coffee plantations were abandoned, potentially reducing density and diversity of native species. Despite limitations, which hinder more general conclusions on coffee invasiveness in Brazilian secondary tropical forests, our results indicate that coffee is a strong competitor in the studied secondary forests and provide important insights for future research on this topic. PMID:29538468
NASA Astrophysics Data System (ADS)
Barcheck, C. Grace; Tulaczyk, Slawek; Schwartz, Susan Y.; Walter, Jacob I.; Winberry, J. Paul
2018-03-01
The Whillans Ice Plain (WIP) is unique among Antarctic ice streams because it moves by stick-slip. The conditions allowing stick-slip and its importance in controlling ice dynamics remain uncertain. Local basal seismicity previously observed during unstable slip is a clue to the mechanism of ice stream stick-slip and a window into current basal conditions, but the spatial extent and importance of this basal seismicity are unknown. We analyze data from a 2010-2011 ice-plain-wide seismic and GPS network to show that basal micro-seismicity correlates with large-scale patterns in ice stream slip behavior: Basal seismicity is common where the ice moves the least between unstable slip events, with small discrete basal micro-earthquakes happening within 10s of km of the central stick-slip nucleation area and emergent basal tremor occurring downstream of this area. Basal seismicity is largely absent in surrounding areas, where inter-slip creep rates are high. The large seismically active area suggests that a frictional sliding law that can accommodate stick-slip may be appropriate for ice stream beds on regional scales. Variability in seismic behavior over inter-station distances of 1-10 km indicates heterogeneity in local bed conditions and frictional complexity. WIP unstable slips may nucleate when stick-slip basal earthquake patches fail over a large area. We present a conceptual model in which basal seismicity results from slip-weakening frictional failure of over-consolidated till as it is eroded and mobilized into deforming till.
Michael G. Shelton; Paul A. Murphy
1997-01-01
Growth was monitored for 4 years in a thinned stand in southern Arkansas with three pine basal areas (70, 85, and 100 ft2/ac) and three hardwood basal areas (0, 15, and 30 ft2/ac); pretreatment basal areasaveraged 119 and 33 ft2/ac for pines and hardwoods, respectively.Treatments were arranged in a 3 x 3 factorial...
Highly Enhanced Gas Adsorption Properties in Vertically Aligned MoS2 Layers.
Cho, Soo-Yeon; Kim, Seon Joon; Lee, Youhan; Kim, Jong-Seon; Jung, Woo-Bin; Yoo, Hae-Wook; Kim, Jihan; Jung, Hee-Tae
2015-09-22
In this work, we demonstrate that gas adsorption is significantly higher in edge sites of vertically aligned MoS2 compared to that of the conventional basal plane exposed MoS2 films. To compare the effect of the alignment of MoS2 on the gas adsorption properties, we synthesized three distinct MoS2 films with different alignment directions ((1) horizontally aligned MoS2 (basal plane exposed), (2) mixture of horizontally aligned MoS2 and vertically aligned layers (basal and edge exposed), and (3) vertically aligned MoS2 (edge exposed)) by using rapid sulfurization method of CVD process. Vertically aligned MoS2 film shows about 5-fold enhanced sensitivity to NO2 gas molecules compared to horizontally aligned MoS2 film. Vertically aligned MoS2 has superior resistance variation compared to horizontally aligned MoS2 even with same surface area exposed to identical concentration of gas molecules. We found that electrical response to target gas molecules correlates directly with the density of the exposed edge sites of MoS2 due to high adsorption of gas molecules onto edge sites of vertically aligned MoS2. Density functional theory (DFT) calculations corroborate the experimental results as stronger NO2 binding energies are computed for multiple configurations near the edge sites of MoS2, which verifies that electrical response to target gas molecules (NO2) correlates directly with the density of the exposed edge sites of MoS2 due to high adsorption of gas molecules onto edge sites of vertically aligned MoS2. We believe that this observation extends to other 2D TMD materials as well as MoS2 and can be applied to significantly enhance the gas sensor performance in these materials.
Estimating Volume, Biomass, and Carbon in Hedmark County, Norway Using a Profiling LiDAR
NASA Technical Reports Server (NTRS)
Nelson, Ross; Naesset, Erik; Gobakken, T.; Gregoire, T.; Stahl, G.
2009-01-01
A profiling airborne LiDAR is used to estimate the forest resources of Hedmark County, Norway, a 27390 square kilometer area in southeastern Norway on the Swedish border. One hundred five profiling flight lines totaling 9166 km were flown over the entire county; east-west. The lines, spaced 3 km apart north-south, duplicate the systematic pattern of the Norwegian Forest Inventory (NFI) ground plot arrangement, enabling the profiler to transit 1290 circular, 250 square meter fixed-area NFI ground plots while collecting the systematic LiDAR sample. Seven hundred sixty-three plots of the 1290 plots were overflown within 17.8 m of plot center. Laser measurements of canopy height and crown density are extracted along fixed-length, 17.8 m segments closest to the center of the ground plot and related to basal area, timber volume and above- and belowground dry biomass. Linear, nonstratified equations that estimate ground-measured total aboveground dry biomass report an R(sup 2) = 0.63, with an regression RMSE = 35.2 t/ha. Nonstratified model results for the other biomass components, volume, and basal area are similar, with R(sup 2) values for all models ranging from 0.58 (belowground biomass, RMSE = 8.6 t/ha) to 0.63. Consistently, the most useful single profiling LiDAR variable is quadratic mean canopy height, h (sup bar)(sub qa). Two-variable models typically include h (sup bar)(sub qa) or mean canopy height, h(sup bar)(sub a), with a canopy density or a canopy height standard deviation measure. Stratification by productivity class did not improve the nonstratified models, nor did stratification by pine/spruce/hardwood. County-wide profiling LiDAR estimates are reported, by land cover type, and compared to NFI estimates.
Reach-scale effects of riparian forest cover on urban stream ecosystems
Roy, A.H.; Faust, C.L.; Freeman, Mary C.; Meyer, J.L.
2005-01-01
We compared habitat and biota between paired open and forested reaches within five small streams (basin area 10?20 km2) in suburban catchments (9%?49% urban land cover) in the Piedmont of Georgia, USA. Stream reaches with open canopies were narrower than forested reaches (4.1 versus 5.0 m, respectively). There were no differences in habitat diversity (variation in velocity, depth, or bed particle size) between open and forested reaches. However, absence of local forest cover corresponded to decreased large wood and increased algal chlorophyll a standing crop biomass. These differences in basal food resources translated into higher densities of fishes in open (9.0 individuals?m?2) versus forested (4.9 individuals?m?2) reaches, primarily attributed to higher densities of the herbivore Campostoma oligolepis. Densities of terrestrial invertebrate inputs were higher in open reaches; however, trends suggested higher biomass of terrestrial inputs in forested reaches and a corresponding higher density of terrestrial prey consumed by water column feeding fishes. Reach-scale biotic integrity (macroinvertebrates, salamanders, and fishes) was largely unaffected by differences in canopy cover. In urbanizing areas where catchment land cover drives habitat and biotic quality, management practices that rely exclusively on forested riparian areas for stream protection are unlikely to be effective at maintaining ecosystem integrity.
Relation of biomass to basal area and site index on an Appalachian watershed
Harry V., Jr. Wiant; Robert Knight; John E. Baumgras
1984-01-01
The biomass of 50-year-old cove hardwood and upland oak stands on an Appalachian watershed was more strongly related to basal area than to site index. Equations are presented for predicting the green and dry weight per acre of biomass components with basal area as the independent variable.
Eren, U; Kum, S; Sandikçi, M; Eren, V; Ilhan, F
2009-08-01
The aim of this study was to determine and examine the distribution of major frequency MHC II+ cells in the oviduct and vagina of cows during the oestrous and dioestrus phases. Right oviduct (ampulla, isthmus) and vaginal samples taken from a total of twenty seven multiparous cows were used. Tissue samples were processed to obtain both cryostat and paraffin sections. Sections were stained immunocytochemically using StreptABC method using a specific monoclonal antibody to MHC II+ cell population. Intra-epithelial and subepithelial areas along with lamina propria, muscularis mucosae and serosa of both ampulla and isthmus and intra-epithelial/subepithelial areas and mucosae of vagina were examined for the presence of MHC II+ cells. The density of immune positive cells was determined using a subjective scoring system. MHC II+ cells were demonstrated in all areas examined in both oestrus and dioestrus. In oestrus, the density of MHC II+ cells decreased in subepithelial areas (in between the epithelial cells and the basal membrane) of isthmus, whereas the density of immune positive cells was increased in muscularis mucosae of isthmus (P < 0.05), lamina propria and muscularis mucosae of ampulla (P < 0.05) as well as in the mucosae of vagina (P
A whole stand basal area projection model for Appalachian hardwoods
John R. Brooks; Lichun Jiang; Matthew Perkowski; Benktesh Sharma
2008-01-01
Two whole-stand basal area projection models were developed for Appalachian hardwood stands. The proposed equations are an algebraic difference projection form based on existing basal area and the change in age, trees per acre, and/or dominant height. Average equation error was less than 10 square feet per acre and residuals exhibited no irregular trends.
Samuelsson, Kristin; Osman, Ayman A. M.; Angeria, Maria; Risling, Mårten; Mohseni, Simin; Press, Rayomand
2016-01-01
Twenty-five percent of polyneuropathies are idiopathic. Microangiopathy has been suggested to be a possible pathogenic cause of chronic idiopathic axonal polyneuropathy (CIAP). Dysfunction of the autophagy pathway has been implicated as a marker of neurodegeneration in the central nervous system, but the autophagy process is not explored in the peripheral nervous system. In the current study, we examined the presence of microangiopathy and autophagy-related structures in sural nerve biopsies of 10 patients with CIAP, 11 controls with inflammatory neuropathy and 10 controls without sensory polyneuropathy. We did not find any significant difference in endoneurial microangiopathic markers in patients with CIAP compared to normal controls, though we did find a correlation between basal lamina area thickness and age. Unexpectedly, we found a significantly larger basal lamina area thickness in patients with vasculitic neuropathy. Furthermore, we found a significantly higher density of endoneurial autophagy-related structures, particularly in patients with CIAP but also in patients with inflammatory neuropathy, compared to normal controls. It is unclear if the alteration in the autophagy pathway is a consequence or a cause of the neuropathy. Our results do not support the hypothesis that CIAP is primarily caused by a microangiopathic process in endoneurial blood vessels in peripheral nerves. The significantly higher density of autophagy structures in sural nerves obtained from patients with CIAP and inflammatory neuropathy vs. controls indicates the involvement of this pathway in neuropathy, particularly in CIAP, since the increase in density of autophagy-related structures was more pronounced in patients with CIAP than those with inflammatory neuropathy. To our knowledge this is the first report investigating signs of autophagy process in peripheral nerves in patients with CIAP and inflammatory neuropathy. PMID:27662650
Schwilk, D.W.; Keeley, J.E.; Knapp, E.E.; Mciver, J.; Bailey, J. D.; Fettig, C.J.; Fiedler, C.E.; Harrod, R.J.; Moghaddas, J.J.; Outcalt, K.W.; Skinner, C.N.; Stephens, S.L.; Waldrop, T.A.; Yaussy, D.A.; Youngblood, A.
2009-01-01
Changes in vegetation and fuels were evaluated from measurements taken before and after fuel reduction treatments (prescribed fire, mechanical treatments, and the combination of the two) at 12 Fire and Fire Surrogate (FFS) sites located in forests with a surface fire regime across the conterminous United States. To test the relative effectiveness of fuel reduction treatments and their effect on ecological parameters we used an informationtheoretic approach on a suite of 12 variables representing the overstory (basal area and live tree, sapling, and snag density), the understory (seedling density, shrub cover, and native and alien herbaceous species richness), and the most relevant fuel parameters for wildfire damage (height to live crown, total fuel bed mass, forest floor mass, and woody fuel mass). In the short term (one year after treatment), mechanical treatments were more effective at reducing overstory tree density and basal area and at increasing quadratic mean tree diameter. Prescribed fire treatments were more effective at creating snags, killing seedlings, elevating height to live crown, and reducing surface woody fuels. Overall, the response to fuel reduction treatments of the ecological variables presented in this paper was generally maximized by the combined mechanical plus burning treatment. If the management goal is to quickly produce stands with fewer and larger diameter trees, less surface fuel mass, and greater herbaceous species richness, the combined treatment gave the most desirable results. However, because mechanical plus burning treatments also favored alien species invasion at some sites, monitoring and control need to be part of the prescription when using this treatment. ?? 2009 by the Ecological Society of America.
Coexistence and community structure of tropical trees in a Hawaiian montane rain forest
Hatfield, J.S.; Link, W.A.; Dawson, D.K.; Lindquist, E.L.
1996-01-01
We measured the diameter at breast height of all trees and shrubs > 5 meters in height, including standing dead trees, on 68 0.04-hectare study plots in a montane, subtropical rain forest on Mauna Loa, Hawai`i. The canopy species consisted of 88 percent Metrosideros polymorpha (ohia) and 12 percent Acacia koa (koa). Negative associations were found between the densities of koa and ohia, the density of koa and the total basal area of ohia, and the total basal areas of koa and ohia. The two-species lottery competition model, a stochastic model in which the coexistence of two species in a space-limited community results from temporal variation in recruitment and death rates, predicts a quadratic-beta distribution for the proportion of space occupied by each species. A discrete version of the quadratic-beta distribution, the quadratic-beta binomial distribution, was fit to the live koa and ohia densities and assessed with goodness-of-fit tests. Likelihood ratio tests provided evidence that the mean adult death rates of the two species were equal but that the relative competitive abilities of the two species favored ohia. These tests were corroborated by a contingency table analysis of death rates based on standing dead trees and growth rate studies which report that koa grows much faster than ohia. The lottery model predicts a positive covariance between death rates and ohia recruitment when mean death rates are equal and koa has a higher growth rate than ohia. We argue that the competitive advantage of ohia is due to its superior dispersal ability into large gaps, which would yield the positive covariance described above, and it is this positive covariance term that skews the occupation of space in favor of ohia.
Boyd E. Wickman
1988-01-01
Twenty-year postoutbreak growth was compared in thinned and unthinned, severely defoliated stands. Basal area of unthinned white fir has declined 37 percent and pine basal area has increased 32 percent since 1964. The stand thinned in 1960 has the lowest basal area in the study area, but the greatest tree growth before and after the outbreak. All defoliated fir are...
Effects of chronic ammonium sulfate treatment on the forest at the Bear Brook Watershed in Maine.
Elvir, Jose Alexander; Wiersma, G Bruce; Bethers, Suzanne; Kenlan, Peter
2010-12-01
At the Bear Brook Watershed in Maine (BBWM), the forest tree composition was characterized and the effects of the chronic ammonium sulfate ((NH(4))(2)SO(4)) treatment on basal area growth, foliar chemistry, and gas exchange were investigated on forest species. The BBWM is a paired watershed forest ecosystem study with one watershed, West Bear (WB), treated since 1989 with 26.6 kg N ha(-1) year(-1) and 30 kg S ha(-1) year(-1)applied bimonthly as (NH(4))(2)SO(4), while the other watershed, East Bear (EB), serves as a reference. Tree species richness, density, and mortality were found to be similar between watersheds. Basal area increment was estimated from red spruce and sugar maple, showing that, for the first 7 years of treatment, it was significantly higher for sugar maple growing in WB compared to EB, but no differences were observed for red spruce between watersheds. However, the initial higher sugar maple basal area growth in WB subsequently decreased after 8 years of treatment. Foliar chemical analysis performed in trees, saplings, and ground flora showed higher N concentrations in the treated WB compared to the reference EB. But, foliar cation concentrations, especially Ca and Mg, were significantly lower for most of the species growing in WB compared with those growing in EB. For sugar maple, foliar N was higher on WB, but there were no differences in foliar Ca and Mg concentrations between treated and reference watersheds. In addition, only sugar maple trees in the treated WB showed significantly higher photosynthetic rates compared to reference EB trees.
Uçakhan, Ömür Ö; Bayraktutar, Betül
2017-01-01
To evaluate the morphological features of the corneal limbus as measured by in vivo confocal microscopy (IVCM) following standard and accelerated corneal collagen cross-linking (CXL) for keratoconus. Patients with progressive keratoconus scheduled to undergo standard CXL (group 1; 31 patients, 3 mW/cm, 370 nm, 30 minutes), or accelerated CXL (group 2; 20 patients, 9 mW/cm, 370 nm, 10 minutes) in the worse eye were included in this prospective study. Thirty eyes of 30 age-matched patients served as controls (group 3). All patient eyes underwent IVCM scanning of the central cornea and the inferior limbal area at baseline and 1, 3, and 6 months after CXL. After CXL, epithelial regrowth was complete by day 4 in both groups 1 and 2. There were no statistically significant differences between the baseline mean central corneal wing or basal cell density, limbus-palisade middle or basal cell densities of groups 1, 2, or 3. At postoperative months 1, 3, and 6, there were no statistically significant differences in either central or limbus-palisade epithelial cell densities or diameters in keratoconic eyes that underwent standard or accelerated CXL (P > 0.05). The morphology of the limbal cells was preserved as well. The morphology of limbus structures seems to be preserved following standard and accelerated CXL in short-term follow-up, as measured using IVCM.
Forest structure in low-diversity tropical forests: a study of Hawaiian wet and dry forests.
Ostertag, Rebecca; Inman-Narahari, Faith; Cordell, Susan; Giardina, Christian P; Sack, Lawren
2014-01-01
The potential influence of diversity on ecosystem structure and function remains a topic of significant debate, especially for tropical forests where diversity can range widely. We used Center for Tropical Forest Science (CTFS) methodology to establish forest dynamics plots in montane wet forest and lowland dry forest on Hawai'i Island. We compared the species diversity, tree density, basal area, biomass, and size class distributions between the two forest types. We then examined these variables across tropical forests within the CTFS network. Consistent with other island forests, the Hawai'i forests were characterized by low species richness and very high relative dominance. The two Hawai'i forests were floristically distinct, yet similar in species richness (15 vs. 21 species) and stem density (3078 vs. 3486/ha). While these forests were selected for their low invasive species cover relative to surrounding forests, both forests averaged 5->50% invasive species cover; ongoing removal will be necessary to reduce or prevent competitive impacts, especially from woody species. The montane wet forest had much larger trees, resulting in eightfold higher basal area and above-ground biomass. Across the CTFS network, the Hawaiian montane wet forest was similar to other tropical forests with respect to diameter distributions, density, and aboveground biomass, while the Hawai'i lowland dry forest was similar in density to tropical forests with much higher diversity. These findings suggest that forest structural variables can be similar across tropical forests independently of species richness. The inclusion of low-diversity Pacific Island forests in the CTFS network provides an ∼80-fold range in species richness (15-1182 species), six-fold variation in mean annual rainfall (835-5272 mm yr(-1)) and 1.8-fold variation in mean annual temperature (16.0-28.4°C). Thus, the Hawaiian forest plots expand the global forest plot network to enable testing of ecological theory for links among species diversity, environmental variation and ecosystem function.
Forest Structure in Low-Diversity Tropical Forests: A Study of Hawaiian Wet and Dry Forests
Ostertag, Rebecca; Inman-Narahari, Faith; Cordell, Susan; Giardina, Christian P.; Sack, Lawren
2014-01-01
The potential influence of diversity on ecosystem structure and function remains a topic of significant debate, especially for tropical forests where diversity can range widely. We used Center for Tropical Forest Science (CTFS) methodology to establish forest dynamics plots in montane wet forest and lowland dry forest on Hawai‘i Island. We compared the species diversity, tree density, basal area, biomass, and size class distributions between the two forest types. We then examined these variables across tropical forests within the CTFS network. Consistent with other island forests, the Hawai‘i forests were characterized by low species richness and very high relative dominance. The two Hawai‘i forests were floristically distinct, yet similar in species richness (15 vs. 21 species) and stem density (3078 vs. 3486/ha). While these forests were selected for their low invasive species cover relative to surrounding forests, both forests averaged 5–>50% invasive species cover; ongoing removal will be necessary to reduce or prevent competitive impacts, especially from woody species. The montane wet forest had much larger trees, resulting in eightfold higher basal area and above-ground biomass. Across the CTFS network, the Hawaiian montane wet forest was similar to other tropical forests with respect to diameter distributions, density, and aboveground biomass, while the Hawai‘i lowland dry forest was similar in density to tropical forests with much higher diversity. These findings suggest that forest structural variables can be similar across tropical forests independently of species richness. The inclusion of low-diversity Pacific Island forests in the CTFS network provides an ∼80-fold range in species richness (15–1182 species), six-fold variation in mean annual rainfall (835–5272 mm yr−1) and 1.8-fold variation in mean annual temperature (16.0–28.4°C). Thus, the Hawaiian forest plots expand the global forest plot network to enable testing of ecological theory for links among species diversity, environmental variation and ecosystem function. PMID:25162731
Siegel, Jessica A.; Park, Byung S.; Raber, Jacob
2013-01-01
Children exposed to methamphetamine during brain development as a result of maternal drug use have long-term hippocampus-dependent cognitive impairments, but the mechanisms underlying these impairments are not understood. The acetylcholine system plays an important role in cognitive function and potential methamphetamine-induced acetylcholine alterations may be related to methamphetamine-induced cognitive impairments. In this study, we investigated the potential long-term effects of methamphetamine exposure during hippocampal development on the acetylcholine system in adolescence mice on postnatal day 30 and in adult mice on postnatal day 90. Methamphetamine exposure increased the density of acetylcholine neurons in regions of the basal forebrain and the area occupied by acetylcholine axons in the hippocampus in adolescent female mice. In contrast, methamphetamine exposure did not affect the density of GABA cells or total neurons in the basal forebrain. Methamphetamine exposure also increased the number of muscarinic acetylcholine receptors in the hippocampus of adolescent male and female mice. Our results demonstrate for the first time that methamphetamine exposure during hippocampal development affects the acetylcholine system in adolescent mice and that these changes are more profound in females than males. PMID:21824143
Basal Area Growth Estimators for Survivor Component: A Quality Control Application
Charles E. Thomas; Francis A. Roesch
1990-01-01
Several possible estimators are available for basal area growth of survivor trees, when horizontal prism (or point) plots (HPP) are remeasured. This study's comparison of three estimators not only provides a check for the estimate of basal area growth but suggests that they can provide a quality control indicator for yield procedures. An example is derived from...
Akiyama, Yoshikatsu; Kikuchi, Akihiko; Yamato, Masayuki; Okano, Teruo
2014-08-01
A double polymeric nanolayer consisting of poly(N-isopropylacrylamide) (PIPAAm) and hydrophilic polyacrylamide (PAAm) was deposited on tissue culture polystyrene (TCPS) surfaces using electron beam irradiation to form a new temperature-responsive cell culture surface in which the basal hydrophilic PAAm component in the double polymeric layer promotes the hydration of the upper PIPAAm layer and induces rapid cell detachment compared to a conventional temperature-responsive cell culture surface, PIPAAm-grafted TCPS (PIPAAm-TCPS). Take-off angle-dependent X-ray photoelectron spectroscopy spectral analysis demonstrated that the grafted PIPAAm and PAAm components were located in the upper and basal regions of the double polymeric layer, respectively, suggesting that the double polymeric layer forms an inter-penetrating-network-like structure with PAAm at the basal portion of the PIPAAm grafted chains. The wettability of the temperature-responsive cell culture surfaces with the double polymeric layer tended to be more hydrophilic, with an increase in the basal PAAm graft density at a constant PIPAAm graft density. However, when the graft densities of the upper PIPAAm and basal PAAm were optimized, the resulting temperature-responsive cell culture surface with the double polymeric layer exhibited rapid cell detachment while maintaining cell adhesive character comparable to that of PIPAAm-TCPS. The cell adhesive character was altered from cell-adhesive to cell-repellent with increasing PAAm or PIPAAm graft density. The cell adhesive character of the temperature-responsive cell culture surfaces was relatively consistent with their contact angles. These results strongly suggest that the basal PAAm surface properties affect the degree of hydration and dehydration of the subsequently grafted PIPAAm. In addition, the roles of the hydrophilic component in accelerating cell detachment are further discussed in terms of the mobility of the grafted PIPAAm chains. Applications of this insight might be useful for designing temperature-responsive cell culture surfaces for achieving efficient cell culture and quick target cell detachment. Copyright © 2014 Acta Materialia Inc. Published by Elsevier Ltd. All rights reserved.
Tree Productivity Enhanced with Conversion from Forest to Urban Land Covers.
Briber, Brittain M; Hutyra, Lucy R; Reinmann, Andrew B; Raciti, Steve M; Dearborn, Victoria K; Holden, Christopher E; Dunn, Allison L
2015-01-01
Urban areas are expanding, changing the structure and productivity of landscapes. While some urban areas have been shown to hold substantial biomass, the productivity of these systems is largely unknown. We assessed how conversion from forest to urban land uses affected both biomass structure and productivity across eastern Massachusetts. We found that urban land uses held less than half the biomass of adjacent forest expanses with a plot level mean biomass density of 33.5 ± 8.0 Mg C ha(-1). As the intensity of urban development increased, the canopy cover, stem density, and biomass decreased. Analysis of Quercus rubra tree cores showed that tree-level basal area increment nearly doubled following development, increasing from 17.1 ± 3.0 to 35.8 ± 4.7 cm(2) yr(-1). Scaling the observed stem densities and growth rates within developed areas suggests an aboveground biomass growth rate of 1.8 ± 0.4 Mg C ha(-1) yr(-1), a growth rate comparable to nearby, intact forests. The contrasting high growth rates and lower biomass pools within urban areas suggest a highly dynamic ecosystem with rapid turnover. As global urban extent continues to grow, cities consider climate mitigation options, and as the verification of net greenhouse gas emissions emerges as critical for policy, quantifying the role of urban vegetation in regional-to-global carbon budgets will become ever more important.
Tree Productivity Enhanced with Conversion from Forest to Urban Land Covers
Briber, Brittain M.; Hutyra, Lucy R.; Reinmann, Andrew B.; Raciti, Steve M.; Dearborn, Victoria K.; Holden, Christopher E.; Dunn, Allison L.
2015-01-01
Urban areas are expanding, changing the structure and productivity of landscapes. While some urban areas have been shown to hold substantial biomass, the productivity of these systems is largely unknown. We assessed how conversion from forest to urban land uses affected both biomass structure and productivity across eastern Massachusetts. We found that urban land uses held less than half the biomass of adjacent forest expanses with a plot level mean biomass density of 33.5 ± 8.0 Mg C ha-1. As the intensity of urban development increased, the canopy cover, stem density, and biomass decreased. Analysis of Quercus rubra tree cores showed that tree-level basal area increment nearly doubled following development, increasing from 17.1 ± 3.0 to 35.8 ± 4.7 cm2 yr-1. Scaling the observed stem densities and growth rates within developed areas suggests an aboveground biomass growth rate of 1.8 ± 0.4 Mg C ha-1 yr-1, a growth rate comparable to nearby, intact forests. The contrasting high growth rates and lower biomass pools within urban areas suggest a highly dynamic ecosystem with rapid turnover. As global urban extent continues to grow, cities consider climate mitigation options, and as the verification of net greenhouse gas emissions emerges as critical for policy, quantifying the role of urban vegetation in regional-to-global carbon budgets will become ever more important. PMID:26302444
Bradford, John B.; Bell, David M.
2017-01-01
Increasing aridity as a result of climate change is expected to exacerbate tree mortality. Reducing forest basal area – the cross-sectional area of tree stems within a given ground area – can decrease tree competition, which may reduce drought-induced tree mortality. However, neither the magnitude of expected mortality increases, nor the potential effectiveness of basal area reduction, has been quantified in dryland forests such as those of the drought-prone Southwest US. We used thousands of repeatedly measured forest plots to show that unusually warm and dry conditions are related to high tree mortality rates and that mortality is positively related to basal area. Those relationships suggest that while increasing high temperature extremes forecasted by climate models may lead to elevated tree mortality during the 21st century, future tree mortality might be partly ameliorated by reducing stand basal area. This adaptive forest management strategy may provide a window of opportunity for forest managers and policy makers to guide forest transitions to species and/or genotypes more suited to future climates.
Large-scale wind disturbances promote tree diversity in a Central Amazon forest.
Marra, Daniel Magnabosco; Chambers, Jeffrey Q; Higuchi, Niro; Trumbore, Susan E; Ribeiro, Gabriel H P M; Dos Santos, Joaquim; Negrón-Juárez, Robinson I; Reu, Björn; Wirth, Christian
2014-01-01
Canopy gaps created by wind-throw events, or blowdowns, create a complex mosaic of forest patches varying in disturbance intensity and recovery in the Central Amazon. Using field and remote sensing data, we investigated the short-term (four-year) effects of large (>2000 m(2)) blowdown gaps created during a single storm event in January 2005 near Manaus, Brazil, to study (i) how forest structure and composition vary with disturbance gradients and (ii) whether tree diversity is promoted by niche differentiation related to wind-throw events at the landscape scale. In the forest area affected by the blowdown, tree mortality ranged from 0 to 70%, and was highest on plateaus and slopes. Less impacted areas in the region affected by the blowdown had overlapping characteristics with a nearby unaffected forest in tree density (583 ± 46 trees ha(-1)) (mean ± 99% Confidence Interval) and basal area (26.7 ± 2.4 m(2) ha(-1)). Highly impacted areas had tree density and basal area as low as 120 trees ha(-1) and 14.9 m(2) ha(-1), respectively. In general, these structural measures correlated negatively with an index of tree mortality intensity derived from satellite imagery. Four years after the blowdown event, differences in size-distribution, fraction of resprouters, floristic composition and species diversity still correlated with disturbance measures such as tree mortality and gap size. Our results suggest that the gradients of wind disturbance intensity encompassed in large blowdown gaps (>2000 m(2)) promote tree diversity. Specialists for particular disturbance intensities existed along the entire gradient. The existence of species or genera taking an intermediate position between undisturbed and gap specialists led to a peak of rarefied richness and diversity at intermediate disturbance levels. A diverse set of species differing widely in requirements and recruitment strategies forms the initial post-disturbance cohort, thus lending a high resilience towards wind disturbances at the community level.
Large-Scale Wind Disturbances Promote Tree Diversity in a Central Amazon Forest
Marra, Daniel Magnabosco; Chambers, Jeffrey Q.; Higuchi, Niro; Trumbore, Susan E.; Ribeiro, Gabriel H. P. M.; dos Santos, Joaquim; Negrón-Juárez, Robinson I.; Reu, Björn; Wirth, Christian
2014-01-01
Canopy gaps created by wind-throw events, or blowdowns, create a complex mosaic of forest patches varying in disturbance intensity and recovery in the Central Amazon. Using field and remote sensing data, we investigated the short-term (four-year) effects of large (>2000 m2) blowdown gaps created during a single storm event in January 2005 near Manaus, Brazil, to study (i) how forest structure and composition vary with disturbance gradients and (ii) whether tree diversity is promoted by niche differentiation related to wind-throw events at the landscape scale. In the forest area affected by the blowdown, tree mortality ranged from 0 to 70%, and was highest on plateaus and slopes. Less impacted areas in the region affected by the blowdown had overlapping characteristics with a nearby unaffected forest in tree density (583±46 trees ha−1) (mean±99% Confidence Interval) and basal area (26.7±2.4 m2 ha−1). Highly impacted areas had tree density and basal area as low as 120 trees ha−1 and 14.9 m2 ha−1, respectively. In general, these structural measures correlated negatively with an index of tree mortality intensity derived from satellite imagery. Four years after the blowdown event, differences in size-distribution, fraction of resprouters, floristic composition and species diversity still correlated with disturbance measures such as tree mortality and gap size. Our results suggest that the gradients of wind disturbance intensity encompassed in large blowdown gaps (>2000 m2) promote tree diversity. Specialists for particular disturbance intensities existed along the entire gradient. The existence of species or genera taking an intermediate position between undisturbed and gap specialists led to a peak of rarefied richness and diversity at intermediate disturbance levels. A diverse set of species differing widely in requirements and recruitment strategies forms the initial post-disturbance cohort, thus lending a high resilience towards wind disturbances at the community level. PMID:25099118
Large-Scale Wind Disturbances Promote Tree Diversity in a Central Amazon Forest
DOE Office of Scientific and Technical Information (OSTI.GOV)
Marra, Daniel Magnabosco; Chambers, Jeffrey Q.; Higuchi, Niro
Canopy gaps created by wind-throw events, or blowdowns, create a complex mosaic of forest patches varying in disturbance intensity and recovery in the Central Amazon. Using field and remote sensing data, we investigated the short-term (four-year) effects of large (>2000 m 2) blowdown gaps created during a single storm event in January 2005 near Manaus, Brazil, to study (i) how forest structure and composition vary with disturbance gradients and (ii) whether tree diversity is promoted by niche differentiation related to wind-throw events at the landscape scale. In the forest area affected by the blowdown, tree mortality ranged from 0 tomore » 70%, and was highest on plateaus and slopes. Less impacted areas in the region affected by the blowdown had overlapping characteristics with a nearby unaffected forest in tree density (583±46 trees ha -1) (mean±99% Confidence Interval) and basal area (26.7±2.4 m 2 ha -1). Highly impacted areas had tree density and basal area as low as 120 trees ha -1 and 14.9 m 2 ha -1, respectively. In general, these structural measures correlated negatively with an index of tree mortality intensity derived from satellite imagery. Four years after the blowdown event, differences in size-distribution, fraction of resprouters, floristic composition and species diversity still correlated with disturbance measures such as tree mortality and gap size. Our results suggest that the gradients of wind disturbance intensity encompassed in large blowdown gaps (>2000 m 2) promote tree diversity. Specialists for particular disturbance intensities existed along the entire gradient. The existence of species or genera taking an intermediate position between undisturbed and gap specialists led to a peak of rarefied richness and diversity at intermediate disturbance levels. A diverse set of species differing widely in requirements and recruitment strategies forms the initial post-disturbance cohort, thus lending a high resilience towards wind disturbances at the community level.« less
Large-Scale Wind Disturbances Promote Tree Diversity in a Central Amazon Forest
Marra, Daniel Magnabosco; Chambers, Jeffrey Q.; Higuchi, Niro; ...
2014-08-06
Canopy gaps created by wind-throw events, or blowdowns, create a complex mosaic of forest patches varying in disturbance intensity and recovery in the Central Amazon. Using field and remote sensing data, we investigated the short-term (four-year) effects of large (>2000 m 2) blowdown gaps created during a single storm event in January 2005 near Manaus, Brazil, to study (i) how forest structure and composition vary with disturbance gradients and (ii) whether tree diversity is promoted by niche differentiation related to wind-throw events at the landscape scale. In the forest area affected by the blowdown, tree mortality ranged from 0 tomore » 70%, and was highest on plateaus and slopes. Less impacted areas in the region affected by the blowdown had overlapping characteristics with a nearby unaffected forest in tree density (583±46 trees ha -1) (mean±99% Confidence Interval) and basal area (26.7±2.4 m 2 ha -1). Highly impacted areas had tree density and basal area as low as 120 trees ha -1 and 14.9 m 2 ha -1, respectively. In general, these structural measures correlated negatively with an index of tree mortality intensity derived from satellite imagery. Four years after the blowdown event, differences in size-distribution, fraction of resprouters, floristic composition and species diversity still correlated with disturbance measures such as tree mortality and gap size. Our results suggest that the gradients of wind disturbance intensity encompassed in large blowdown gaps (>2000 m 2) promote tree diversity. Specialists for particular disturbance intensities existed along the entire gradient. The existence of species or genera taking an intermediate position between undisturbed and gap specialists led to a peak of rarefied richness and diversity at intermediate disturbance levels. A diverse set of species differing widely in requirements and recruitment strategies forms the initial post-disturbance cohort, thus lending a high resilience towards wind disturbances at the community level.« less
Charles E. Rose; Thomas B. Lynch
2001-01-01
A method was developed for estimating parameters in an individual tree basal area growth model using a system of equations based on dbh rank classes. The estimation method developed is a compromise between an individual tree and a stand level basal area growth model that accounts for the correlation between trees within a plot by using seemingly unrelated regression (...
Relative maxima of diameter and basal area
Thomas B. Lynch; Difei Zhang
2012-01-01
It has often been observed that maximum dbh growth occurs at an earlier age than maximum individual tree basal area growth. This can be deduced from the geometry of the tree stem, by observing that a dbh increment at a given radius will be associated with a larger basal area increment than an equal dbh increment occurring at a shorter radius from the stem center. Thus...
Relation of snowpack Accumulation to Red Pine Stocking
Edward A. Hansen
1969-01-01
A snow accumulation study was conducted in a 33-year-old red pine plantation thinned to different stocking levels. Snowpack water content increased an average of 2 percent for each 10 square feet of basal area reduction, within the range of 60 to 180 square feet of basal area. Reducing plantation stocking from 180 to 60 square feet of basal area per acre would result...
Basal erosion: barrier to earthquake propagation? Insight from the northern chilean forearc
NASA Astrophysics Data System (ADS)
Cubas, N.
2017-12-01
Subducted topographic features have often been suspected as barriers to large earthquake propagation. These features would induce basal erosion, leading to a large network of fractures impeding large nucleation or shear localization. Looking for correlation between basal erosion and megathrust ruptures is thus critical nowadays to understand earthquake mechanics and infer rupture scenarios. In this study, we propose to seek possible location of basal erosion from the forearc morphology by applying the critical taper theory. We focus on the North Chile subduction zone that has experienced four major earthquakes during the last two decades and where basal erosion and seamount subduction have already been suspected. Basal erosion should occur when the basal friction approaches the internal friction. We thus seek what part of the forearc is at critical state and select areas for which the two frictions are almost equal. We find a large band, located at 25km depth, from the Mejillones peninsula to the Iquique region at critical state with very high basal friction. The critical areas seem to surround the Tocopilla 2007 Mw 7.7 and the Iquique 2014 Mw 8.1 ruptures. When compared with the interseismic coupling, except for the Tocopilla segment, the critical areas are located in low-coupled zones. More interestingly, the reported normal faults of the forearc do not appear above the erosional areas but rather between them. These normal faults are systematically located above locked patches and seismic asperities. These areas are actually at extensional critical state and characterized by a very low effective friction. We thus suspect the extensional features to be related to earthquakes rather than basal erosion. We then look for similar relationships along the Sumatra subduction zone to see if basal erosion is a common process. The Tocopilla and Iquique earthquakes ruptured only part of the northern Chile seismic gap although the full segment was ready for a new large rupture. Areas of basal erosion might have impeded the propagation of these ruptures and could as well delineate the future mega-earthquake.
[Structural recovering in Andean successional forests from Porce (Antioquia, Colombia)].
Yepes, Adriana P; del Valle, Jorge I; Jaramillo, Sandra L; Orrego, Sergio A
2010-03-01
Places subjected to natural or human disturbance can recover forest through an ecological process called secondary succession. Tropical succession is affected by factors such as disturbances, distance from original forest, surface configuration and local climate. These factors determine the composition of species and the time trend of the succession itself. We studied succession in soils used for cattle ranching over various decades in the Porce Region of Colombia (Andean Colombian forests). A set of twenty five permanent plots was measured, including nine plots (20 x 50 m) in primary forests and sixteen (20 x 25 m) in secondary forests. All trees with diameter > or =1.0 cm were measured. We analyzed stem density, basal area, above-ground biomass and species richness, in a successional process of ca. 43 years, and in primary forests. The secondary forests' age was estimated in previous studies, using radiocarbon dating, aerial photographs and a high-resolution satellite image analysis (7 to >43 years). In total, 1,143 and 1,766 stems were measured in primary and secondary forests, respectively. Basal area (5.7 to 85.4 m2 ha(-1)), above-ground biomass (19.1 to 1,011.5 t ha(-1)) and species richness (4 to 69) directly increased with site age, while steam density decreased (3,180 to 590). Diametric distributions were "J-inverted" for primary forests and even-aged size-class structures for secondary forests. Three species of palms were abundant and exclusive in old secondary forests and primary forests: Oenocarpus mapora, Euterpe precatoria and Oenocarpus bataua. These palms happened in cohorts after forest disturbances. Secondary forest structure was 40% in more than 43 years of forest succession and indicate that many factors are interacting and affecting the forests succession in the area (e.g. agriculture, cattle ranching, mining, etc.).
NASA Astrophysics Data System (ADS)
Rolim, Samir Gonçalves; Sambuichi, Regina Helena Rosa; Schroth, Götz; Nascimento, Marcelo Trindade; Gomes, José Manoel Lucio
2017-03-01
Cocoa agroforests like the cabrucas of Brazil's Atlantic forest are among the agro-ecosystems with greatest potential for biodiversity conservation. Despite a global trend for their intensification, cocoa agroforests are also being abandoned for socioeconomic reasons especially on marginal sites, because they are incorporated in public or private protected areas, or are part of mandatory set-asides under Brazilian environmental legislation. However, little is known about phylogenetic structure, the processes of forest regeneration after abandonment and the conservation value of former cabruca sites. Here we compare the vegetation structure and composition of a former cabruca 30-40 years after abandonment with a managed cabruca and mature forest in the Atlantic forest region of Espirito Santo, Brazil. The forest in the abandoned cabruca had recovered a substantial part of its original structure. Abandoned cabruca have a higher density (mean ± CI95 %: 525.0 ± 40.3 stems per ha), basal area (34.0 ± 6.5 m2 per ha) and species richness (148 ± 11.5 species) than managed cabruca (96.0 ± 17.7; 24.15 ± 3.9 and 114.5 ± 16.0, respectively) but no significant differences to mature forest in density (581.0 ± 42.2), basal area (29.9.0 ± 3.3) and species richness (162.6 ± 15.5 species). Thinning (understory removal) changes phylogenetic structure from evenness in mature forest to clustering in managed cabruca, but after 30-40 years abandoned cabruca had a random phylogenetic structure, probably due to a balance between biotic and abiotic filters at this age. We conclude that abandoned cocoa agroforests present highly favorable conditions for the regeneration of Atlantic forest and could contribute to the formation of an interconnected network of forest habitat in this biodiversity hotspot.
Veronica I. Emhart; Timothy A. Martin; Timothy L. White; Dudley A. Huber
2006-01-01
We quantified basal area increment phenology over a 2-year period in one loblolly pine (Pinus taeda L.) and four slash pine (Pinus elliottii Engelm. var. elliottii) full-sib families propagated as rooting cuttings. In 2002, basal area growth started in March and stopped in October for both species, while in 2003, initiation and cessation occurred 2...
A novel form of epithelial wound healing of the embryonic epidermis.
Armstrong, Margaret T; Turlo, Kirsten; Elges, Chris J; Dayton, Sarah M; Lee, Janet; Armstrong, Peter B
2006-08-01
The embryonic epidermis of amniotes is a two-cell layer sheet with a periderm positioned superficial to the basal cell layer which, itself, attaches apically to the basal surface of the periderm and basally to the basal lamina. The presence of the periderm is essential to maintain the basal layer as a two-dimensional monolayer. Wounds to the epidermis that remove selectively just the periderm are healed by a stacking of the basal layer cells that constitute the wound bed. Basal cell stacking involves the desertion of the basal lamina by many of the cells so as to increase their contact area with other basal layer cells. This suggests that a preferential adhesion to the planar basal lamina is not important for the monolayered organization of the basal layer but, instead, association with inner surface of the planar periderm is the principal process that maintains the basal layer as a monolayer. The conversion of the basal layer from monolayer to multilayer during wound healing diminishes its planar area, resulting in movement of the wound borders toward the center of the wound. This is a novel scenario for wound healing.
Shakeri, M; Zulkifli, I; Soleimani, A F; O'Reilly, E L; Eckersall, P D; Anna, A A; Kumari, S; Abdullah, F F J
2014-11-01
A study was conducted to determine whether supplementing AminoGut (a commercial dietary supplement containing a mixture of l-glutamine and l-glutamic acid) to broiler chickens stocked at 2 different densities affected performance, physiological stress responses, foot pad dermatitis incidence, and intestinal morphology and microflora. A randomized design in a factorial arrangement with 4 diets [basal diet, basal diet + 0.5% AminoGut from d 1 to 21, basal diet + 0.5% AminoGut from d 1 to 42, and basal diet + virginiamycin (0.02%) for d 1 to 42] and 2 stocking densities [0.100 m(2)/bird (23 birds/pen; LD) or 0.067 m(2)/bird (35 birds/pen; HD)]. Results showed that villi length and crypt depth were not changed by different dietary treatments. However, birds in the HD group had smaller villi (P = 0.03) compared with those of the LD group. Regardless of diet, HD consistently increased the serum concentrations of ceruloplasmin, α-1 acid glycoprotein, ovotransferin, and corticosterone (P = 0.0007), and elevated heterophil to lymphocyte ratio (0.0005). Neither AminoGut supplementation nor stocking density affected cecal microflora counts. In conclusion, under the conditions of this study, dietary supplementation of AminoGut, irrespective of stocking density, had no beneficial effect on growth performance, intestinal morphology, and physiological adaptive responses of broiler chickens raised under hot and humid tropical conditions. However, AminoGut supplementation from d 1 to 42 was beneficial in reducing mortality rate. Also, the increased serum concentrations of a wide range of acute phase proteins together with elevated corticosterone and heterophil to lymphocyte ratio suggested that high stocking density induced an acute phase response either indirectly as a result of increased incidence of inflammatory diseases such as foot pad dermatitis or possibly as a direct physiological response to the stress of high stocking density. ©2014 Poultry Science Association Inc.
NASA Astrophysics Data System (ADS)
Casas Planes, Á.; Garcia, M.; Siegel, R.; Koltunov, A.; Ramirez, C.; Ustin, S.
2015-12-01
Occupancy and habitat suitability models for snag-dependent wildlife species are commonly defined as a function of snag basal area. Although critical for predicting or assessing habitat suitability, spatially distributed estimates of snag basal area are not generally available across landscapes at spatial scales relevant for conservation planning. This study evaluates the use of airborne laser scanning (ALS) to 1) identify individual conifer snags and map their basal area across a recently burned forest, and 2) map habitat suitability for a wildlife species known to be dependent on snag basal area, specifically the black-backed woodpecker (Picoides arcticus). This study focuses on the Rim Fire, a megafire that took place in 2013 in the Sierra Nevada Mountains of California, creating large patches of medium- and high-severity burned forest. We use forest inventory plots, single-tree ALS-derived metrics and Gaussian processes classification and regression to identify conifer snags and estimate their stem diameter and basal area. Then, we use the results to map habitat suitability for the black-backed woodpecker using thresholds for conifer basal area from a previously published habitat suitability model. Local maxima detection and watershed segmentation algorithms resulted in 75% detection of trees with stem diameter larger than 30 cm. Snags are identified with an overall accuracy of 91.8 % and conifer snags are identified with an overall accuracy of 84.8 %. Finally, Gaussian process regression reliably estimated stem diameter (R2 = 0.8) using height and crown area. This work provides a fast and efficient methodology to characterize the extent of a burned forest at the tree level and a critical tool for early wildlife assessment in post-fire forest management and biodiversity conservation.
NASA Astrophysics Data System (ADS)
Fatma, N. A. H.; Wan Juliana, W. A.; Shaharuddin, M. I.; Wickneswari, R.
2016-11-01
A descriptive study of species composition, community structure and biomass was conducted in compartment 107, which is a rehabilitated area at Kenaboi Forest Reserve, Jelebu, Negeri Sembilan. The objective is to determine the forest structure and species composition in a rehabilitated area of Kenaboi FR since enrichment planting had done. A sample plot of 1 hectare was censused and a total of 395 trees with diameter ≥ 5 cm DBH were recorded. A total of 285 individual trees were identified belonging to 20 families and the commonest family was Dipterocarpaceae with 193 individuals. The highest tree density per ha was Shorea acuminata at 33% followed by S. parvifolia, 10% and S. leprosula, 6%. The biggest tree was Artocarpus elasticus Reinw. ex Blume with a diameter of 101 cm. The total basal area was 34.48 m2/ha, whereby the highest basal area was between 45 - 54.9 cm DBH class that contributed 10.21 m2/ha (30%). The total biomass estimation (above ground and below ground) was 792.57 t/ha. Dipterocarpaceae contributed the highest total biomass at 545.14 t/ha with S. acuminata contributed the highest total biomass of 330.45 t/ha. This study will contribute to the knowledge of regeneration forest especially on how the ecological process restoring the biodiversity and ecosystem functioning in rehabilitated forest by practicing the enrichment planting of native species.
A system for predicting the amount of Phellinus (Fomes) igniarius rot in trembling aspen stands
Robert L. Anderson; Arthur L. Jr. Schipper
1978-01-01
The occurrence of Phellinus (Fomes) igniarius white trunk rot in 45- to 50-year-old trembling aspen stands can be predicted by applying a constant to the stand basal area with P. igniarius conks to estimate the total basal area with P. igniarius rot. Future decay projections can be made by reapplying the basal area of hidden decay for each 6 years projected. This paper...
Timothy B. Harrington; M. Boyd Edwards
1997-01-01
Twelve years after various methods and intensities of site preparation in the Georgia Piedmont, diameter distributions and basal area (BA) of pines and hardwoods varied considerably among treatments. Site preparation reduced hardwood basal area to 36 percent of that observed in clearcut-only plots. As a result, planted-pine BA in the presence of site preparation was 2...
Niemiec, Joanna; Adamczyk, Agnieszka; Ambicka, Aleksandra; Mucha-Małecka, Anna; Wysocki, Wojciech; Mituś, Jerzy; Ryś, Janusz
2012-11-01
Lymphangiogenesis is a potential indicator of cancer patients' survival. However, there is no standardisation of methodologies applied to the assessment of lymphatic vessel density. In 156 invasive ductal breast cancers (T 1/N+/M0), lymphatic and blood vessels were visualised using podoplanin and CD34, respectively. Based on these markers expression, four parameters were assessed: (i) distribution of podoplanin-stained vessels (DPV) - the percentage of fields with at least one lymphatic vessel (a simple method proposed by us), (ii) lymphatic vessel density (LVD), (iii) LVD to microvessel density ratio (LVD/MVD) and (iv) the expression of podoplanin in cancer-associated fibroblasts. Next, we estimated relations between the above-mentioned parameters and: (i) breast cancer subtype, (ii) tumour grade, and (iii) basal markers expression. We found that intensive lymphangiogenesis, assessed using all studied methods, is positively related to high tumour grade, triple negative or HER2 subtype and expression of basal markers. Whereas, the absence of podoplanin expression in fibroblasts of cancer stroma is related to luminal A subtype, low tumour grade or lack of basal markers expression. Distribution of podoplanin-stained vessels, assessed by a simple method proposed by us (indicating the percentage of fields with at least one lymphatic vessel), might be used instead of the "hot-spot" method.
Modeling the Effects of Fire Frequency and Severity on Forests in the Northwestern United States
Busing, Richard T.; Solomon, Allen M.
2006-01-01
This study used a model of forest dynamics (FORCLIM) and actual forest survey data to demonstrate the effects of various fire regimes on different forest types in the Pacific Northwest. We examined forests in eight ecoregions ranging from wet coastal forests dominated by Pseudotsuga menziesii and other tall conifers to dry interior forests dominated by Pinus ponderosa. Fire effects simulated as elevated mortality of trees based on their species and size did alter forest structure and species composition. Low frequency fires characteristic of wetter forests (return interval >200 yr) had minor effects on composition. When fires were severe, they tended to reduce total basal area with little regard to species differences. High frequency fires characteristic of drier forests (return interval <30 yr) had major effects on species composition and on total basal area. Typically, they caused substantial reductions in total basal area and shifts in dominance toward highly fire tolerant species. With the addition of fire, simulated basal areas averaged across ecoregions were reduced to levels approximating observed basal areas.
Novel characterization of landscape-level variability in historical vegetation structure.
Collins, Brandon M; Lydersen, Jamie M; Everett, Richard G; Fry, Danny L; Stephens, Scott L
2015-07-01
We analyzed historical timber inventory data collected systematically across a large mixed-conifer-dominated landscape to gain insight into the interaction between disturbances and vegetation structure and composition prior to 20th century land management practices. Using records from over 20 000 trees, we quantified historical vegetation structure and composition for nine distinct vegetation groups. Our findings highlight some key aspects of forest structure under an intact disturbance regime: (1) forests were low density, with mean live basal area and tree density ranging from 8-30 m2 /ha and 25-79 trees/ha, respectively; (2) understory and overstory structure and composition varied considerably across the landscape; and (3) elevational gradients largely explained variability in forest structure over the landscape. Furthermore, the presence of large trees across most of the surveyed area suggests that extensive stand-replacing disturbances were rare in these forests. The vegetation structure and composition characteristics we quantified, along with evidence of largely elevational control on these characteristics, can provide guidance for restoration efforts in similar forests.
Iversen, C. M. [Oak Ridge National Laboratory, U.S. Department of Energy, Oak Ridge, Tennessee, U.S.A; Garrett, A. [Oak Ridge National Laboratory, U.S. Department of Energy, Oak Ridge, Tennessee, U.S.A; Martin, A. [Oak Ridge National Laboratory, U.S. Department of Energy, Oak Ridge, Tennessee, U.S.A; Turetsky, M. R. [Oak Ridge National Laboratory, U.S. Department of Energy, Oak Ridge, Tennessee, U.S.A; Norby, R. J. [Oak Ridge National Laboratory, U.S. Department of Energy, Oak Ridge, Tennessee, U.S.A; Childs, J. [Oak Ridge National Laboratory, U.S. Department of Energy, Oak Ridge, Tennessee, U.S.A; Ontl, T. A. [Oak Ridge National Laboratory, U.S. Department of Energy, Oak Ridge, Tennessee, U.S.A
2017-01-01
The composition and cover of woody and understory species, along with the timing of stem growth, were characterized near minirhizotron installations at the south and north ends of the S1 bog in order to understand the relationship between root dynamics and the surrounding plant communities. There are three data streams: (1) Woody species basal area -- the number, basal area, and distance from the center point of all trees within a 5-m radius of each minirhizotron pair, which were quantified in 2011, (2) Understory cover -- the understory vascular plant community composition and percent cover, which were surveyed in 1 m2 areas at four cardinal directions adjacent to each pair of minirhizotrons in June 2011, and (3) Stem growth -- the basal area increment (stem growth) of nearby trees, which was quantified using automated or manual dendrobands in 2011 and 2012, respectively.
Goring, Simon; Mladenoff, David J.; Cogbill, Charles; Record, Sydne; Paciorek, Christopher J.; Dietze, Michael C.; Dawson, Andria; Matthes, Jaclyn; McLachlan, Jason S.; Williams, John W.
2016-01-01
EuroAmerican land-use and its legacies have transformed forest structure and composition across the United States (US). More accurate reconstructions of historical states are critical to understanding the processes governing past, current, and future forest dynamics. Here we present new gridded (8x8km) reconstructions of pre-settlement (1800s) forest composition and structure from the upper Midwestern US (Minnesota, Wisconsin, and most of Michigan), using 19th Century Public Land Survey System (PLSS), with estimates of relative composition, above-ground biomass, stem density, and basal area for 28 tree types. This mapping is more robust than past efforts, using spatially varying correction factors to accommodate sampling design, azimuthal censoring, and biases in tree selection.
Effect of the environment on the dendritic morphology of the rat auditory cortex
Bose, Mitali; Muñoz-Llancao, Pablo; Roychowdhury, Swagata; Nichols, Justin A.; Jakkamsetti, Vikram; Porter, Benjamin; Byrapureddy, Rajasekhar; Salgado, Humberto; Kilgard, Michael P.; Aboitiz, Francisco; Dagnino-Subiabre, Alexies; Atzori, Marco
2010-01-01
The present study aimed to identify morphological correlates of environment-induced changes at excitatory synapses of the primary auditory cortex (A1). We used the Golgi-Cox stain technique to compare pyramidal cells dendritic properties of Sprague-Dawley rats exposed to different environmental manipulations. Sholl analysis, dendritic length measures, and spine density counts were used to monitor the effects of sensory deafness and an auditory version of environmental enrichment (EE). We found that deafness decreased apical dendritic length leaving basal dendritic length unchanged, whereas EE selectively increased basal dendritic length without changing apical dendritic length. On the contrary, deafness decreased while EE increased spine density in both basal and apical dendrites of A1 layer 2/3 (LII/III) neurons. To determine whether stress contributed to the observed morphological changes in A1, we studied neural morphology in a restraint-induced model that lacked behaviorally relevant acoustic cues. We found that stress selectively decreased apical dendritic length in the auditory but not in the visual primary cortex. Similar to the acoustic manipulation, stress-induced changes in dendritic length possessed a layer specific pattern displaying LII/III neurons from stressed animals with normal apical dendrites but shorter basal dendrites, while infragranular neurons (layers V and VI) displayed shorter apical dendrites but normal basal dendrites. The same treatment did not induce similar changes in the visual cortex, demonstrating that the auditory cortex is an exquisitely sensitive target of neocortical plasticity, and that prolonged exposure to different acoustic as well as emotional environmental manipulation may produce specific changes in dendritic shape and spine density. PMID:19771593
De Jager, N. R.; Pastor, J.
2009-01-01
Ungulate herbivores create patterns of forage availability, plant species composition, and soil fertility as they range across large landscapes and consume large quantities of plant material. Over time, herbivore populations fluctuate, producing great potential for spatio-temporal landscape dynamics. In this study, we extend the spatial and temporal extent of a long-term investigation of the relationship of landscape patterns to moose foraging behavior at Isle Royale National Park, MI. We examined how patterns of browse availability and consumption, plant basal area, and soil fertility changed during a recent decline in the moose population. We used geostatistics to examine changes in the nature of spatial patterns in two valleys over 18 years and across short-range and long-range distance scales. Landscape patterns of available and consumed browse changed from either repeated patches or randomly distributed patches in 1988-1992 to random point distributions by 2007 after a recent record high peak followed by a rapid decline in the moose population. Patterns of available and consumed browse became decoupled during the moose population low, which is in contrast to coupled patterns during the earlier high moose population. Distributions of plant basal area and soil nitrogen availability also switched from repeated patches to randomly distributed patches in one valley and to random point distributions in the other valley. Rapid declines in moose population density may release vegetation and soil fertility from browsing pressure and in turn create random landscape patterns. ?? Springer Science+Business Media B.V. 2009.
McCoy, John W.; Draugelis-Dale, Rassa O.; Keeland, Bobby D.; Darville, Roy
2010-01-01
Aboveground primary productivity for cypress forests was assessed from measurements of litter production in two age groups and in two hydrological regimes (standing water and free-flowing). Caddo Lake, located in northeast Texas on the Texas-Louisiana border, offered a unique study site since it is dominated by extensive stands composed entirely of Taxodium distichum (L.) Rich, (baldcypress) in different age groups. Young stands (approximately 100 years old) are found along the shoreline and on shallow flooded islands. Old stands (-150 to 300 years old) are found in deeper water where they were continuously flooded. Litter production over three years from October 1998 to September 2001 was measured. Litter consisting of leaves, twigs, bark, reproductive parts, and Tillandsia usneoides (L.) L. (Spanish moss) was collected monthly using 0.5 m2 floating traps. Tree diameters were measured within 200 m2 circular plots in each stand. The young stands supported densities greater than 2,000 stems/ha and a mean stand basal area of 72.3 m2/ha, whereas old stands supported lower densities of about 500 stems/ha but with a similar mean stand basal area of 73.3 m2/ha. There was a significant difference between old and young stands for overall yearly litter production, averaging about 670 g/m2/yr in the young stands and 460 g/m2/yr in the old stands. Leaves and twigs were significantly greater in the young stands, while reproductive parts were higher in old stands. Litter collections between years or hydrological regimes were not significantly different.
Cognitive-motor interactions of the basal ganglia in development
Leisman, Gerry; Braun-Benjamin, Orit; Melillo, Robert
2014-01-01
Neural circuits linking activity in anatomically segregated populations of neurons in subcortical structures and the neocortex throughout the human brain regulate complex behaviors such as walking, talking, language comprehension, and other cognitive functions associated with frontal lobes. The basal ganglia, which regulate motor control, are also crucial elements in the circuits that confer human reasoning and adaptive function. The basal ganglia are key elements in the control of reward-based learning, sequencing, discrete elements that constitute a complete motor act, and cognitive function. Imaging studies of intact human subjects and electrophysiologic and tracer studies of the brains and behavior of other species confirm these findings. We know that the relation between the basal ganglia and the cerebral cortical region allows for connections organized into discrete circuits. Rather than serving as a means for widespread cortical areas to gain access to the motor system, these loops reciprocally interconnect a large and diverse set of cerebral cortical areas with the basal ganglia. Neuronal activity within the basal ganglia associated with motor areas of the cerebral cortex is highly correlated with parameters of movement. Neuronal activity within the basal ganglia and cerebellar loops associated with the prefrontal cortex is related to the aspects of cognitive function. Thus, individual loops appear to be involved in distinct behavioral functions. Damage to the basal ganglia of circuits with motor areas of the cortex leads to motor symptoms, whereas damage to the subcortical components of circuits with non-motor areas of the cortex causes higher-order deficits. In this report, we review some of the anatomic, physiologic, and behavioral findings that have contributed to a reappraisal of function concerning the basal ganglia and cerebellar loops with the cerebral cortex and apply it in clinical applications to attention deficit/hyperactivity disorder (ADHD) with biomechanics and a discussion of retention of primitive reflexes being highly associated with the condition. PMID:24592214
Caro-Magdaleno, Manuel; Alfaro-Juárez, Asunción; Montero-Iruzubieta, Jesús; Fernández-Palacín, Ana; Muñoz-Morales, Ana; Castilla-Martino, Manuel Alberto; Spínola-Muñoz, Consuelo; Rodríguez-de-la-Rúa, Enrique
2018-05-18
Limbal stem cell deficiency (LSCD) is characterised by a marked decrease in limbal stem cells. It is classified primarily using subjective slit-lamp observations. In vivo confocal microscopy (IVCM) can non-invasively provide objective information on the condition of the limbal niche, the corneal epithelial basal cell density and the corneal sub-basal nerve plexus density (SND). We here used IVCM to evaluate changes in SND to improve LSCD classification. We evaluated and classified 38 patients (76 eyes, 44 with LSC and 32 control eyes) using the Rama, López-García and Deng (clinical and confocal) classifications and evaluated the concordance of the confocal and clinical classifications. We constructed a logistic regression model using multivariate analysis to correlate different degrees of conjunctivalisation with IVCM parameters and used receiver operating characteristic (ROC) curve analysis to establish the SND cut-off value with maximum diagnostic sensitivity and specificity. The classification systems correlated moderately at best (kappa, 0.449). The corneal SND of cases (6469±6295 µm/mm 2 ) was less (p<0.001) than in controls (20911±4142 µm/mm 2 ). The SND, but not basal cell density, played a protective role against conjunctivalisation (OR, 0.069; 95% CI 0.008-0.619; p=0.01). An SND cut-off value of 17 215 µm/mm 2 yielded a sensitivity and specificity of 95.5% and 90.6%, respectively, for LSCD diagnosis. The density of the corneal sub-basal nerve plexus was inversely related to conjunctivalisation in LSCD. Further studies are needed to verify this and to elucidate the directionality between these factors. © Article author(s) (or their employer(s) unless otherwise stated in the text of the article) 2018. All rights reserved. No commercial use is permitted unless otherwise expressly granted.
NASA Astrophysics Data System (ADS)
Elshaafi, Abdelsalam; Gudmundsson, Agust
2017-05-01
The Al Haruj Volcanic Province (AHVP) consists of two distinct volcanic systems. In the north is the system of Al Haruj al Aswad, covering an area of 34,200 km2, while in the south the system of Al Haruj al Abyad, covering an area of 7,850 km2. The systems have produced some 432 monogenetic volcanoes, primarily scoria (cinder) cones, lava shields, and maars. The density distribution of the volcanoes in each system, plotted as eruption points or sites, has a roughly elliptical surface expression, suggesting similar plan-view geometry of the magma sources, here suggested as deep-seated reservoirs. More specifically, the Al Haruj al Aswad magma reservoir has major and minor axes of 210 km and 119 km, respectively, and an area of 19,176 km2, the corresponding figures for the Haruj al Abyad reservoir being 108 km and 74 km, for the axes, and 6209 km2 for the area. We measured 55 lava shields on the AHVP. They are mostly restricted to the northern and southern parts of AHVP and date from late Miocene to (at least) the end of Pleistocene, while some may have been active into Holocene. In fact, although primarily monogenetic, some of the lava shields show evidence of (possibly Holocene) fissure eruptions in the summit parts. The early lava shields tend to be located at the edges of volcanic systems and with greater volumes than later (more central) shields. The average lava shield basal diameter is 4.5 km and height 63 m. There is strong linear correlation between lava shield volume and basal area, the coefficient of determination (R2) being about 0.75. When 22 Holocene Icelandic lava shields are added to the dataset, for comparison, the correlation between volume and basal area becomes R2 = 0.95. Numerical models suggest that the local stress fields favoured rupture and dyke injection at the margins of the source reservoirs during late Miocene - early Pliocene, in agreement with the distribution of the early, large-volume shields.
Helmisaari, Heljä-Sisko; Derome, John; Nöjd, Pekka; Kukkola, Mikko
2007-10-01
Variations in fine root biomass of trees and understory in 16 stands throughout Finland were examined and relationships to site and stand characteristics determined. Norway spruce fine root biomass varied between 184 and 370 g m(-2), and that of Scots pine ranged between 149 and 386 g m(-2). In northern Finland, understory roots and rhizomes (< 2 mm diameter) accounted for up to 50% of the stand total fine root biomass. Therefore, the fine root biomass of trees plus understory was larger in northern Finland in stands of both tree species, resulting in a negative relationship between fine root biomass and the temperature sum and a positive relationship between fine root biomass and the carbon:nitrogen ratio of the soil organic layer. The foliage:fine root ratio varied between 2.1 and 6.4 for Norway spruce and between 0.8 and 2.2 for Scots pine. The ratio decreased for both Norway spruce and Scots pine from south to north, as well as from fertile to more infertile site types. The foliage:fine root ratio of Norway spruce was related to basal area and stem surface area. The strong positive correlations of these three parameters with fine root nitrogen concentration implies that more fine roots are needed to maintain a certain amount of foliage when nutrient availability is low. No significant relationships were found between stand parameters and fine root biomass at the stand level, but the relationships considerably improved when both fine root biomass and stand parameters were calculated for the mean tree in the stand. When the northern and southern sites were analyzed separately, fine root biomass per tree of both species was significantly correlated with basal area and stem surface area per tree. Basal area, stem surface area and stand density can be estimated accurately and easily. Thus, our results may have value in predicting fine root biomass at the tree and stand level in boreal Norway spruce and Scots pine forests.
Reduction of stand density as a management tool to mitigate the effect of drought
NASA Astrophysics Data System (ADS)
Giuggiola, A.; Rigling, A.; Dobbertin, M.
2012-04-01
An increasing frequency and severity of drought combined with increased competition due to reduced forest management practices are putting many Scots pine forest (Pinus sylvestris) under increased drought pressure. Declining pines are already been observed in many drought exposed regions as in southern Europe or in the dry inner Alpine valleys. Thus, forest management practices oriented at reducing competition for water should increase pines tolerance to climate change and thus enhancing their long-term mitigation potential. In this study, we are testing the beneficial effect of thinning and understory removal as possible management practices. As a first study object we selected a trial with 3 thinning intensities (basal area reduction of 15%, 46% and 70%) and one control (unmanaged forest). The second experiment consisted in removing the understory layer in a radius of 5 meter from 6 mature pine trees. Water-related indicators, such as soil water content, sapflow, point dendrometer and ring width measurements over the growing season were then compared with control trees. Both objects belongs to the pine forests from the dry Rhone valley. Our results indicates that over the 10 years following the thinning performed in 1965 (when the stand was 45 years old) doubled and quadrupled the basal area increment in the medium and heavy treatments compared to the control. The annual mortality rates for the period 1978-1990 ranged between 2.9% for the control and 0.8% for the heavy thinned stand. An increasing mortality rates during the period 1991-2009 (up to 3.3%), with consequent decline in basal area and carbon sequestration, has been observed in relation to high remaining stand density. The removal of understory performed in April 2010 increased soil water content at 30 cm and 65 cm depth reducing trees drought stress. The transpiration and the predawn leaf water potential of overstory trees were higher in the trees with removal of the understory vegetation. The same trees also showed delayed stomata closure during drought periods. This advantage eventually resulted in a significant increase in tree growth, which was observed in the second year after removal. This combining approach highlight the positive effect of competition removal of growth and vitality of Scots pine trees. Our results suggest that in dry forests of the inner-Alps or the Mediterranean region, a strong reduction of basal area (15-25 m2/ha) is suitable to preserve enough water availability to maintain and healthy state. If thinning lead to a major increase of the understory layer, understory removal would prolong the initial positive effect of thinning. Thinning and understory removal enhance tree vitality and increase tree resistance against drought, insects and pathogens. These results simultaneously suggests that pine decline is not driven only by the increase of temperature, but is also a consequence of a change of forest use, specifically for the Valais, to a reduction of goat grazing pressure and wood harvesting.
Ozcelik, Ramazan; Gul, Altay Ugur; Merganic, Jan; Merganicova, Katarina
2008-05-01
We studied the effects of stand parameters (crown closure, basal area, stand volume, age, mean stand diameter number of trees, and heterogeneity index) and geomorphology features (elevation, aspect and slope) on tree species diversity in an example of untreated natural mixed forest stands in the eastern Black Sea region of Turkey. Tree species diversity and basal area heterogeneity in forest ecosystems are quantified using the Shannon-Weaver and Simpson indices. The relationship between tree species diversity basal area heterogeneity stand parameters and geomorphology features are examined using regression analysis. Our work revealed that the relationship between tree species diversity and stand parameters is loose with a correlation coefficient between 0.02 and 0.70. The correlation of basal area heterogeneity with stand parameters fluctuated between 0.004 and 0.77 (R2). According to our results, stands with higher tree species diversity are characterised by higher mean stand diameter number of diameter classes, basal area and lower homogeneity index value. Considering the effect of geomorphology features on tree species or basal area heterogeneity we found that all investigated relationships are loose with R < or = 0.24. A significant correlation was detected only between tree species diversity and aspect. Future work is required to verify the detected trends in behaviour of tree species diversity if it is to estimate from the usual forest stand parameters and topography characteristics.
Schulze, E-D; Wirth, C; Mollicone, D; Ziegler, W
2005-11-01
The dark taiga of Siberia is a boreal vegetation dominated by Picea obovata, Abies sibirica, and Pinus sibirica during the late succession. This paper investigates the population and age structure of 18 stands representing different stages after fire, wind throw, and insect damage. To our knowledge, this is the first time that the forest dynamics of the Siberian dark taiga is described quantitatively in terms of succession, and age after disturbance, stand density, and basal area. The basis for the curve-linear age/diameter relation of trees is being analyzed. (1) After a stand-replacing fire Betula dominates (4,000 trees) for about 70 years. Although tree density of Betula decreases rapidly, basal area (BA) reached >30 m2/ha after 40 years. (2) After fire, Abies, Picea, and Pinus establish at the same time as Betula, but grow slower, continue to gain height and eventually replace Betula. Abies has the highest seedling number (about 1,000 trees/ha) and the highest mortality. Picea establishes with 100-400 trees/ha, it has less mortality, but reached the highest age (>350 years, DBH 51 cm). Picea is the most important indicator for successional age after disturbance. Pinus sibirica is an accompanying species. The widely distributed "mixed boreal forest" is a stage about 120 years after fire reaching a BA of >40 m2/ha. (3) Wind throw and insect damage occur in old conifer stands. Betula does not establish. Abies initially dominates (2,000-6,000 trees/ha), but Picea becomes dominant after 150-200 years since Abies is shorter lived. (4) Without disturbance the forest develops into a pure coniferous canopy (BA 40-50 m2/ha) with a self-regenerating density of 1,000 coniferous canopy trees/ha. There is no collapse of old-growth stands. The dark taiga may serve as an example in which a limited set to tree species may gain dominance under certain disturbance conditions without ever getting monotypic.
Nevoid Basal Cell Carcinoma Syndrome
... develops. Some individuals may have thousands of basal cell cancers in areas of skin that are exposed to ... have been approved to treat people with basal cell cancers that have spread in the body or that ...
Computational insight into the capacitive performance of graphene edge planes
Zhan, Cheng; Zhang, Yu; Cummings, Peter T.; ...
2017-02-01
Recent experiments have shown that electric double-layer capacitors utilizing electrodes consisting of graphene edge plane exhibit higher capacitance than graphene basal plane. However, theoretical understanding of this capacitance enhancement is still limited. Here we applied a self-consistent joint density functional theory calculation on the electrode/electrolyte interface and found that the capacitance of graphene edge plane depends on the edge type: zigzag edge has higher capacitance than armchair edge due to the difference in their electronic structures. We further examined the quantum, dielectric, and electric double-layer (EDL) contributions to the total capacitance of the edge-plane electrodes. Classical molecular dynamics simulation foundmore » that the edge planes have higher EDL capacitance than the basal plane due to better adsorption of counter-ions and higher solvent accessible surface area. Finally, our work therefore has elucidated the capacitive energy storage in graphene edge planes that take into account both the electrode's electronic structure and the EDL structure.« less
Physiological activation of Akt by PHLPP1 deletion protects against pathological hypertrophy.
Moc, Courtney; Taylor, Amy E; Chesini, Gino P; Zambrano, Cristina M; Barlow, Melissa S; Zhang, Xiaoxue; Gustafsson, Åsa B; Purcell, Nicole H
2015-02-01
To examine the role of physiological Akt signalling in pathological hypertrophy through analysis of PHLPP1 (PH domain leucine-rich repeat protein phosphatase) knock-out (KO) mice. To investigate the in vivo requirement for 'physiological' control of Akt activation in cardiac growth, we examined the effect of deleting the Akt phosphatase, PHLPP, on the induction of cardiac hypertrophy. Basal Akt phosphorylation increased nearly two-fold in the cardiomyocytes from PHLPP1 KO mice and physiological hypertrophy induced by swimming exercise was accentuated as assessed by increased heart size and myocyte cell area. In contrast, the development of pathophysiological hypertrophy induced by pressure overload and assessed by increases in heart size, myocyte cell area, and hypertrophic gene expression was attenuated. This attenuation coincided with decreased fibrosis and cell death in the KO mice. Cast moulding revealed increased capillary density basally in the KO hearts, which was further elevated relative to wild-type mouse hearts in response to pressure overload. In vitro studies with isolated myocytes in co-culture also demonstrated that PHLPP1 deletion in cardiomyocytes can enhance endothelial tube formation. Expression of the pro-angiogenic factor VEGF was also elevated basally and accentuated in response to transverse aortic constriction in hearts from KO mice. Our data suggest that enhancing Akt activity by inhibiting its PHLPP1-mediated dephosphorylation promotes processes associated with physiological hypertrophy that may be beneficial in attenuating the development of pathological hypertrophy. Published on behalf of the European Society of Cardiology. All rights reserved. © The Author 2014. For permissions please email: journals.permissions@oup.com.
The influence of badland surfaces and erosion processes on vegetation cover
NASA Astrophysics Data System (ADS)
Hardenbicker, Ulrike; Matheis, Sarah
2014-05-01
To assess the links between badland geomorphology and vegetation cover, we used detailed mapping in the Avonlea badlands, 60 km southwest of Regina, Saskatchewan Canada. Three badlands surfaces are typical in the study area: a basal pediment surface, a mid-slope of bentonitic mudstone with typical popcorn surface, and an upper slope with mud-cemented sandstone. Badland development was triggered by rapid post Pleistocene incision of a meltwater channel in Upper Cretaceous marine and lagoonal sediments. After surveying and mapping of a test area, sediment samples were taken to analyze geophysical parameters. A detailed geomorphic map and vegetation map (1:1000) were compared and analyzed in order to determine the geomorphic environment for plant colonization. The shrink-swell capacity of the bentonitic bedrock, slaking potential and dispersivity are controlled by soil texture, clay mineralogy and chemistry, strongly influencing the timing and location of runoff and the relative significance of surface and subsurface erosional processes. The absence of shrink-swell cracking of the alluvial surfaces of the pediments indicates a low infiltration capacity and sheetflow. The compact lithology of the sandstone is responsible for its low permeability and high runoff coefficient. Slope drainage of steep sandstone slopes is routed through a deep corrasional pipe network. Silver sagebrush (Artemisia cana) is the only species growing on the popcorn surface of the mudrock, which is in large parts vegetation free. The basal pediment shows a distinct 2 m band surrounding the mudrock outcrop without vegetation as a result of high sedimentation rate due to slope wash. Otherwise the typical pioneer vegetation of this basal pediment are grasses. In the transition zone below the steep sandstone cliffs and above the gentle bentonitic mudrock surfaces patches of short-grass vegetation are found, marking slumped blocks with intact vegetation and soil cover. These patches are surrounded by less dense pioneer vegetation consisting of grasses and sage bushes indicating minimal surface erosion or sedimentation. Geomorphic mapping documented a high density of active pipes in this area, transporting silt and fine sand from the sandstone cliffs to lower and basal pediments. Vegetation cover alone is a poor indicator of badland surfaces and erosion processes because of the three-dimensional nature of badland erosion processes, and the shrink-swell capacity of the bentonitic bedrock. A combination of geomorphic and vegetation mapping is needed to identify badland surfaces and processes in the study area.
Hernández-Ruedas, Manuel A; Arroyo-Rodríguez, Víctor; Morante-Filho, José Carlos; Meave, Jorge A; Martínez-Ramos, Miguel
2018-05-04
Understanding the patterns and processes driving biodiversity maintenance in fragmented tropical forests is urgently needed for conservation planning, especially in species-rich forest reserves. Of particular concern are the effects that habitat modifications at the landscape scale may have on forest regeneration and ecosystem functioning: a topic that has received limited attention. Here, we assessed the effects of landscape structure (i.e., forest cover, open area matrices, forest fragmentation, and mean inter-patch isolation distance) on understory plant assemblages in the Los Tuxtlas Biosphere Reserve, Mexico. Previous studies suggest that the demographic burst of the strong competitor palm Astrocaryum mexicanum in the core area of this reserve limits plant recruitment and imperils biodiversity conservation within this protected area. Yet, the local and landscape predictors of this palm, and its impact on tree recruitment at a regional scale are unknown. Thus, we used structural equation modeling to assess the direct and cascading effects of landscape structure on stem and species density in the understory of 20 forest sites distributed across this biodiversity hotspot. Indirect paths included the effect of landscape structure on tree basal area (a proxy of local disturbance), and the effects of these variables on A. mexicanum. Density of A. mexicanum mainly increased with decreasing both fragmentation and open areas in the matrix (matrix contrast, hereafter), and such an increase in palm density negatively affected stem and species density in the understory. The negative direct effect of matrix contrast on stem density was overridden by the indirect positive effects (i.e., through negative cascading effects on A. mexicanum), resulting in a weak effect of matrix contrast on stem density. These findings suggest that dispersal limitation and negative edge effects in more fragmented landscapes dominated by open areas prevent the proliferation of this palm species, enhancing the diversity and abundance of understory trees. This "positive" news adds to an increasing line of evidence suggesting that fragmentation may have some positive effects on biodiversity, in this case by preventing the proliferation of species that can jeopardize biodiversity conservation within tropical reserves. © 2018 by the Ecological Society of America.
Unoki, H; Fan, J; Watanabe, T
1999-01-01
We investigated the structural and functional properties of human umbilical vein endothelial cells (HUVECs) cultured on a two-chamber culture model system using an amnion membrane. Compared to HUVECs cultured on a plastic dish, HUVECs cultured on the model system exhibited several features similar to those of in vivo vessels, including formation of the intercellular junctional devices and expression of tight junction-associated protein ZO-1 and adherence junction-associated protein alpha-catenin. Furthermore, we found that HUVECs had a property of polar secretion of endothelin-1 (ET-1). About 90% of the total amount of synthesized ET-1 was found in the lower well, designated as the basal side. When HUVECs were incubated with either native low-density lipoproteins (nLDLs) or oxidized LDLs (oxLDLs) at a concentration of 100 microgram/ml, ET-1 secretion was significantly increased, dependent on the cell side (apical vs basal) on which the nLDLs or oxLDLs were loaded. When the LDLs were loaded on the apical side, the secretion of ET-1 from HUVECs on the apical side was increased by 48% (nLDL) and 61% (oxLDL), whereas it was accompanied by a concomitant decrease of ET-1 on the basal side (45% by nLDLs and 38% by oxLDLs). When loaded on the basal side, however, ET-1 was increased by 23% (nLDLs) and 53% (oxLDLs) on the basal side, with a 26% simultaneous decrease of ET-1 on the opposite side for both nLDLs and oxLDLs. On the contrary, high-density lipoproteins (HDLs) inhibited ET-1 secretion from HUVECs on the opposite side of the well on which HDLs were loaded; there was a 57% decrease on the basal side when HDLs were loaded on the apical side, and a 46% decrease on the apical side when loaded on the basal side. These results indicate that modulation of ET-1 secretion from ECs by lipoproteins is virtually dependent on the place (apical vs basal) where these proteins are present. The finding that nLDLs and oxLDLs enhance ET-1 secretion by ECs in a polarized pattern suggests that ET-1 may be involved in pathophysiological processes such as atherogenesis.
Structure and energetics of extended defects in ice Ih
NASA Astrophysics Data System (ADS)
Silva Junior, Domingos L.; de Koning, Maurice
2012-01-01
We consider the molecular structure and energetics of extended defects in proton-disordered hexagonal ice Ih. Using plane-wave density functional theory (DFT) calculations, we compute the energetics of stacking faults and determine the structure of the 30∘ and 90∘ partial dislocations on the basal plane. Consistent with experimental data, the formation energies of all fully reconstructed stacking faults are found to be very low. This is consistent with the idea that basal-plane glide dislocations in ice Ih are dissociated into partial dislocations separated by an area of stacking fault. For both types of partial dislocation we find a strong tendency toward core reconstruction through pairwise hydrogen-bond reformation. In the case of the 30∘ dislocation, the pairwise hydrogen-bond formation leads to a period-doubling core structure equivalent to that seen in zinc-blende semiconductor crystals. For the 90∘ partial we consider two possible core reconstructions, one in which the periodicity of the structure along the core remains unaltered and another in which it is doubled. The latter is preferred, although the energy difference between both is rather small, so that a coexistence of both reconstructions appears plausible. Our results imply that a mobility theory for dislocations on the basal plane in ice Ih should be based on the idea of reconstructed partial dislocations.
Historical harvests reduce neighboring old-growth basal area across a forest landscape.
Bell, David M; Spies, Thomas A; Pabst, Robert
2017-07-01
While advances in remote sensing have made stand, landscape, and regional assessments of the direct impacts of disturbance on forests quite common, the edge influence of timber harvesting on the structure of neighboring unharvested forests has not been examined extensively. In this study, we examine the impact of historical timber harvests on basal area patterns of neighboring old-growth forests to assess the magnitude and scale of harvest edge influence in a forest landscape of western Oregon, USA. We used lidar data and forest plot measurements to construct 30-m resolution live tree basal area maps in lower and middle elevation mature and old-growth forests. We assessed how edge influence on total, upper canopy, and lower canopy basal area varied across this forest landscape as a function of harvest characteristics (i.e., harvest size and age) and topographic conditions in the unharvested area. Upper canopy, lower canopy, and total basal area increased with distance from harvest edge and elevation. Forests within 75 m of harvest edges (20% of unharvested forests) had 4% to 6% less live tree basal area compared with forest interiors. An interaction between distance from harvest edge and elevation indicated that elevation altered edge influence in this landscape. We observed a positive edge influence at low elevations (<800 m) and a negative edge influence at moderate to high elevations (>800 m). Surprisingly, we found no or weak effects of harvest age (13-60 yr) and harvest area (0.2-110 ha) on surrounding unharvested forest basal area, implying that edge influence was relatively insensitive to the scale of disturbance and multi-decadal recovery processes. Our study indicates that the edge influence of past clearcutting on the structure of neighboring uncut old-growth forests is widespread and persistent. These indirect and diffuse legacies of historical timber harvests complicate forest management decision-making in old-growth forest landscapes by broadening the traditional view of stand boundaries. Furthermore, the consequences of forest harvesting may reach across ownership boundaries, highlighting complex governance issues surrounding landscape management of old-growth forests. © 2017 by the Ecological Society of America.
Windthrow and salvage logging in an old-growth hemlock-northern hardwoods forest
Lang, K.D.; Schulte, L.A.; Guntenspergen, G.R.
2009-01-01
Although the initial response to salvage (also known as, post-disturbance or sanitary) logging is known to vary among system components, little is known about longer term forest recovery. We examine forest overstory, understory, soil, and microtopographic response 25 years after a 1977 severe wind disturbance on the Flambeau River State Forest in Wisconsin, USA, a portion of which was salvage logged. Within this former old-growth hemlock-northern hardwoods forest, tree dominance has shifted from Eastern hemlock (Tsuga canadensis) to broad-leaf deciduous species (Ulmus americana, Acer saccharum, Tilia americana, Populus tremuloides, and Betula alleghaniensis) in both the salvaged and unsalvaged areas. While the biological legacies of pre-disturbance seedlings, saplings, and mature trees were initially more abundant in the unsalvaged area, regeneration through root suckers and stump sprouts was common in both areas. After 25 years, tree basal area, sapling density, shrub layer density, and seedling cover had converged between unsalvaged and salvaged areas. In contrast, understory herb communities differed between salvaged and unsalvaged forest, with salvaged forest containing significantly higher understory herb richness and cover, and greater dominance of species benefiting from disturbance, especially Solidago species. Soil bulk density, pH, organic carbon content, and organic nitrogen content were also significantly higher in the salvaged area. The structural legacy of tip-up microtopography remains more pronounced in the unsalvaged area, with significantly taller tip-up mounds and deeper pits. Mosses and some forest herbs, including Athyrium filix-femina and Hydrophyllum virginianum, showed strong positive responses to this tip-up microrelief, highlighting the importance of these structural legacies for understory biodiversity. In sum, although the pathways of recovery differed, this forest appeared to be as resilient to the compound disturbances of windthrow plus salvage logging as to wind disturbance alone, by most vegetative measures.
Parkinson's disease as a system-level disorder.
Caligiore, Daniele; Helmich, Rick C; Hallett, Mark; Moustafa, Ahmed A; Timmermann, Lars; Toni, Ivan; Baldassarre, Gianluca
2016-01-01
Traditionally, the basal ganglia have been considered the main brain region implicated in Parkinson's disease. This single area perspective gives a restricted clinical picture and limits therapeutic approaches because it ignores the influence of altered interactions between the basal ganglia and other cerebral components on Parkinsonian symptoms. In particular, the basal ganglia work closely in concert with cortex and cerebellum to support motor and cognitive functions. This article proposes a theoretical framework for understanding Parkinson's disease as caused by the dysfunction of the entire basal ganglia-cortex-cerebellum system rather than by the basal ganglia in isolation. In particular, building on recent evidence, we propose that the three key symptoms of tremor, freezing, and impairments in action sequencing may be explained by considering partially overlapping neural circuits including basal ganglia, cortical and cerebellar areas. Studying the involvement of this system in Parkinson's disease is a crucial step for devising innovative therapeutic approaches targeting it rather than only the basal ganglia. Possible future therapies based on this different view of the disease are discussed.
Wallace, K J; Laughlin, Daniel C; Clarkson, Bruce D
2017-06-01
Restoring forest structure and composition is an important component of urban land management, but we lack clear understanding of the mechanisms driving restoration success. Here we studied two indicators of restoration success in temperate rainforests: native tree regeneration and epiphyte colonization. We hypothesized that ecosystem properties such as forest canopy openness, abundance of exotic herbaceous weeds, and the microclimate directly affect the density and diversity of native tree seedlings and epiphytes. Relationships between environmental conditions and the plant community were investigated in 27 restored urban forests spanning 3-70 years in age and in unrestored and remnant urban forests. We used structural equation modelling to determine the direct and indirect drivers of native tree regeneration and epiphyte colonization in the restored forests. Compared to remnant forest, unrestored forest had fewer native canopy tree species, significantly more light reaching the forest floor annually, and higher exotic weed cover. Additionally, epiphyte density was lower and native tree regeneration density was marginally lower in the unrestored forests. In restored forests, light availability was reduced to levels found in remnant forests within 20 years of restoration planting, followed shortly thereafter by declines in herbaceous exotic weeds and reduced fluctuation of relative humidity and soil temperatures. Contrary to expectations, canopy openness was only an indirect driver of tree regeneration and epiphyte colonization, but it directly regulated weed cover and microclimatic fluctuations, both of which directly drove the density and richness of regeneration and epiphyte colonization. Epiphyte density and diversity were also positively related to forest basal area, as large trees provide physical habitat for colonization. These results imply that ecosystem properties change predictably after initial restoration plantings, and that reaching critical thresholds in some ecosystem properties makes conditions suitable for the regeneration of late successional species, which is vital for restoration success and long-term ecosystem sustainability. Abiotic and biotic conditions that promote tree regeneration and epiphyte colonization will likely be present in forests with a basal area ≥27 m 2 /ha. We recommend that urban forest restoration plantings be designed to promote rapid canopy closure to reduce light availability, suppress herbaceous weeds, and stabilize the microclimate. © 2017 by the Ecological Society of America.
Mahelkova, Gabriela; Jirsova, Katerina; Seidler Stangova, Petra; Palos, Michalis; Vesela, Viera; Fales, Ivan; Jiraskova, Nada; Dotrelova, Dagmar
2017-05-01
In vivo corneal confocal microscopy allows the examination of each layer of the cornea in detail and the identification of pathological changes at the cellular level. The purpose of this study was to identify the possible effects of a three-month treatment with autologous serum eye-drops in different corneal layers of patients with severe dry eye disease using corneal confocal microscopy. Twenty-six patients with dry eye disease were included in the study. Corneal fluorescein staining was performed. The corneas of the right eyes were examined using in vivo corneal confocal microscopy before and after a three-month treatment with autologous serum drops. The densities of superficial and basal epithelial cells, Langerhans cells, the keratocytes and activated keratocytes, the density of endothelial cells and the status of the sub-basal nerve plexus fibres were evaluated. A significant decrease in corneal fluorescein staining was found after the three-month autologous serum treatment (p = 0.0006). The basal epithelial cell density decreased significantly (p = 0.001), while the density of superficial epithelial cells did not change significantly (p = 0.473) nor did the number of Langerhans cells or activated keratocytes (p = 0.223; p = 0.307, respectively). There were no differences in the other corneal cell layers or in the status of the nerve fibres. The results demonstrate the ability of corneal confocal microscopy to evaluate an improvement in the basal epithelial cell layer of the cornea after autologous serum treatment in patients with dry eye disease. More studies with longer follow-up periods are needed to elucidate the suitability of corneal confocal microscopy to follow the effect of autologous serum treatment on nerve fibres or other corneal layers in dry eye disease patients. © 2016 Optometry Australia.
Effects of oyster harvest activities on Louisiana reef habitat and resident nekton communities
Beck, Steve; LaPeyre, Megan K.
2015-01-01
Oysters are often cited as “ecosystem engineers” because they modify their environment. Coastal Louisiana contains extensive oyster reef areas that have been harvested for decades, and whether differences in habitat functions exist between those areas and nonharvested reefs is unclear. We compared reef physical structure and resident community metrics between these 2 subtidal reef types. Harvested reefs were more fragmented and had lower densities of live eastern oysters (Crassostrea virginica) and hooked mussels (Ischadium recurvum) than the nonharvested reefs. Stable isotope values (13C and 15N) of dominant nekton species and basal food sources were used to compare food web characteristics. Nonpelagic source contributions and trophic positions of dominant species were slightly elevated at harvested sites. Oyster harvesting appeared to have decreased the number of large oysters and to have increased the percentage of reefs that were nonliving by decreasing water column filtration and benthopelagic coupling. The differences in reef matrix composition, however, had little effect on resident nekton communities. Understanding the thresholds of reef habitat areas, the oyster density or oyster size distribution below which ecosystem services may be compromised, remains key to sustainable management.
Jucker, Tommaso; Sanchez, Aida Cuni; Lindsell, Jeremy A; Allen, Harriet D; Amable, Gabriel S; Coomes, David A
2016-06-01
Tropical forests currently play a key role in regulating the terrestrial carbon cycle and abating climate change by storing carbon in wood. However, there remains considerable uncertainty as to whether tropical forests will continue to act as carbon sinks in the face of increased pressure from expanding human activities. Consequently, understanding what drives productivity in tropical forests is critical. We used permanent forest plot data from the Gola Rainforest National Park (Sierra Leone) - one of the largest tracts of intact tropical moist forest in West Africa - to explore how (1) stand basal area and tree diversity, (2) past disturbance associated with past logging, and (3) underlying soil nutrient gradients interact to determine rates of aboveground wood production (AWP). We started by statistically modeling the diameter growth of individual trees and used these models to estimate AWP for 142 permanent forest plots. We then used structural equation modeling to explore the direct and indirect pathways which shape rates of AWP. Across the plot network, stand basal area emerged as the strongest determinant of AWP, with densely packed stands exhibiting the fastest rates of AWP. In addition to stand packing density, both tree diversity and soil phosphorus content were also positively related to productivity. By contrast, historical logging activities negatively impacted AWP through the removal of large trees, which contributed disproportionately to productivity. Understanding what determines variation in wood production across tropical forest landscapes requires accounting for multiple interacting drivers - with stand structure, tree diversity, and soil nutrients all playing a key role. Importantly, our results also indicate that logging activities can have a long-lasting impact on a forest's ability to sequester and store carbon, emphasizing the importance of safeguarding old-growth tropical forests.
Composition and structure of Pinus koraiensis mixed forest respond to spatial climatic changes.
Zhang, Jingli; Zhou, Yong; Zhou, Guangsheng; Xiao, Chunwang
2014-01-01
Although some studies have indicated that climate changes can affect Pinus koraiensis mixed forest, the responses of composition and structure of Pinus koraiensis mixed forests to climatic changes are unknown and the key climatic factors controlling the composition and structure of Pinus koraiensis mixed forest are uncertain. Field survey was conducted in the natural Pinus koraiensis mixed forests along a latitudinal gradient and an elevational gradient in Northeast China. In order to build the mathematical models for simulating the relationships of compositional and structural attributes of the Pinus koraiensis mixed forest with climatic and non-climatic factors, stepwise linear regression analyses were performed, incorporating 14 dependent variables and the linear and quadratic components of 9 factors. All the selected new models were computed under the +2°C and +10% precipitation and +4°C and +10% precipitation scenarios. The Max Temperature of Warmest Month, Mean Temperature of Warmest Quarter and Precipitation of Wettest Month were observed to be key climatic factors controlling the stand densities and total basal areas of Pinus koraiensis mixed forest. Increased summer temperatures and precipitations strongly enhanced the stand densities and total basal areas of broadleaf trees but had little effect on Pinus koraiensis under the +2°C and +10% precipitation scenario and +4°C and +10% precipitation scenario. These results show that the Max Temperature of Warmest Month, Mean Temperature of Warmest Quarter and Precipitation of Wettest Month are key climatic factors which shape the composition and structure of Pinus koraiensis mixed forest. Although the Pinus koraiensis would persist, the current forests dominated by Pinus koraiensis in the region would all shift and become broadleaf-dominated forests due to the dramatic increase of broadleaf trees under the future global warming and increased precipitation.
Giolli, R A; Gregory, K M; Suzuki, D A; Blanks, R H; Lui, F; Betelak, K F
2001-01-01
Anatomical findings are presented that identify cortical and subcortical sources of afferents to the nucleus reticularis tegmenti pontis (NRTP) and basal pontine nuclei. Projections from the middle temporal visual area (MT), medial superior temporal visual area (MST), lateral intraparietal area (LIP), and areas 7a and 7b to the basal pontine nuclei were studied using 3H-leucine autoradiography. The results complemented a parallel study of retrograde neuronal labeling attributable to injecting WGA-HRP into NRTP and neighboring pontine nuclei. Small 3H-leucine injections confined to MT, MST, LIP, area 7a, or area 7b, produced multiple patches of pontine terminal label distributed as follows: (1) An injection within MT produced terminal label limited to the dorsolateral and lateral pontine nuclei. (2) Injections restricted to MST or LIP showed patches of terminal label in the dorsal, dorsolateral, lateral, and peduncular pontine nuclei. (3) Area 7a targets the dorsal, dorsolateral, lateral, peduncular, and ventral pontine nuclei, whereas area 7b projects, additionally, to the dorsomedial and paramedian pontine nuclei. Notably, no projections were seen to NRTP from any of these cortical areas. In contrast, injections made by other investigators into cortical areas anterior to the central sulcus revealed cerebrocortical afferents to NRTP, in addition to nuclei of the basal pontine gray. With our pontine WGA-HRP injections, retrograde neuronal labeling was observed over a large extent of the frontal cortex continuing onto the medial surface which included the lining of the cingulate sulcus and cingulate gyrus. Significant subcortical sources for afferents to the NRTP and basal pontine nuclei were the zona incerta, ventral mesencephalic tegmentum, dorsomedial hypothalamic area, rostral interstitial nucleus of the medial longitudinal fasciculus, red nucleus, and subthalamic nucleus. The combined anterograde and retrograde labeling data indicated that visuo-motor cortico-pontine pathways arising from parietal cortices target only the basal pontine gray, whereas the NRTP, together with select pontine nuclei, is a recipient of afferents from frontal cortical areas. The present findings implicate the existence of parallel direct and indirect cortico-pontine pathways from frontal motor-related cortices to NRTP and neighboring pontine nuclei.
Within-plant distribution of onion thrips (Thysanoptera: Thripidae) in onions.
Mo, Jianhua; Munro, Scott; Boulton, Alan; Stevens, Mark
2008-08-01
Two aspects of the within-plant distribution of Thrips tabaci Lindeman (Thysanoptera: Thripidae) on onion, Allium cepa L., plants were investigated: 1) diurnal variations in the distribution of adults and larvae between basal and upper sections of onion leaves, and 2) between-leaf and within-leaf distribution of the eggs. The diurnal investigations showed that higher proportions of larvae than of adults congregated at the basal sections of plants, particularly when plants were young and thrips density was low. As plants matured and thrips density increased, the larvae became more dispersed. Regardless of plant size, there were always more adults in the upper than basal plant sections. There were no clear time-windows during the 24-h diurnal cycle when more thrips were in the upper plant parts. T. tabaci eggs were laid everywhere in the plant. Leaves of intermediate ages had more eggs than older or younger leaves. Within leaves, the white leaf sheath received the least eggs and leaf tips received slightly more eggs than leaf sheaths. The highest egg density was found between the green leaf base and the leaf tips. Regardless of plant size, more than half of all eggs were laid above the basal sections. The percentage increased to >95% in mature plants. Except when plants were small the outer leaves were preferred over inner leaves and upper leaf sections preferred over lower leaf sections as egg-laying sites by adults. Implications of the results in the management of T. tabaci are discussed.
Rolim, Samir Gonçalves; Sambuichi, Regina Helena Rosa; Schroth, Götz; Nascimento, Marcelo Trindade; Gomes, José Manoel Lucio
2017-03-01
Cocoa agroforests like the cabrucas of Brazil's Atlantic forest are among the agro-ecosystems with greatest potential for biodiversity conservation. Despite a global trend for their intensification, cocoa agroforests are also being abandoned for socioeconomic reasons especially on marginal sites, because they are incorporated in public or private protected areas, or are part of mandatory set-asides under Brazilian environmental legislation. However, little is known about phylogenetic structure, the processes of forest regeneration after abandonment and the conservation value of former cabruca sites. Here we compare the vegetation structure and composition of a former cabruca 30-40 years after abandonment with a managed cabruca and mature forest in the Atlantic forest region of Espirito Santo, Brazil. The forest in the abandoned cabruca had recovered a substantial part of its original structure. Abandoned cabruca have a higher density (mean ± CI95 %: 525.0 ± 40.3 stems per ha), basal area (34.0 ± 6.5 m 2 per ha) and species richness (148 ± 11.5 species) than managed cabruca (96.0 ± 17.7; 24.15 ± 3.9 and 114.5 ± 16.0, respectively) but no significant differences to mature forest in density (581.0 ± 42.2), basal area (29.9.0 ± 3.3) and species richness (162.6 ± 15.5 species). Thinning (understory removal) changes phylogenetic structure from evenness in mature forest to clustering in managed cabruca, but after 30-40 years abandoned cabruca had a random phylogenetic structure, probably due to a balance between biotic and abiotic filters at this age. We conclude that abandoned cocoa agroforests present highly favorable conditions for the regeneration of Atlantic forest and could contribute to the formation of an interconnected network of forest habitat in this biodiversity hotspot.
Neural correlates underlying micrographia in Parkinson’s disease
Zhang, Jiarong; Hallett, Mark; Feng, Tao; Hou, Yanan; Chan, Piu
2016-01-01
Micrographia is a common symptom in Parkinson’s disease, which manifests as either a consistent or progressive reduction in the size of handwriting or both. Neural correlates underlying micrographia remain unclear. We used functional magnetic resonance imaging to investigate micrographia-related neural activity and connectivity modulations. In addition, the effect of attention and dopaminergic administration on micrographia was examined. We found that consistent micrographia was associated with decreased activity and connectivity in the basal ganglia motor circuit; while progressive micrographia was related to the dysfunction of basal ganglia motor circuit together with disconnections between the rostral supplementary motor area, rostral cingulate motor area and cerebellum. Attention significantly improved both consistent and progressive micrographia, accompanied by recruitment of anterior putamen and dorsolateral prefrontal cortex. Levodopa improved consistent micrographia accompanied by increased activity and connectivity in the basal ganglia motor circuit, but had no effect on progressive micrographia. Our findings suggest that consistent micrographia is related to dysfunction of the basal ganglia motor circuit; while dysfunction of the basal ganglia motor circuit and disconnection between the rostral supplementary motor area, rostral cingulate motor area and cerebellum likely contributes to progressive micrographia. Attention improves both types of micrographia by recruiting additional brain networks. Levodopa improves consistent micrographia by restoring the function of the basal ganglia motor circuit, but does not improve progressive micrographia, probably because of failure to repair the disconnected networks. PMID:26525918
Briggs, Jenny S.; Hawbaker, Todd J.; Vandendriesche, Don
2015-01-01
After causing widespread mortality in lodgepole pine forests in North America, the mountain pine beetle (MPB) has recently also affected ponderosa pine, an alternate host species that may have different levels of resilience to this disturbance. We collected field data in ponderosa pine- and lodgepole pine-dominated forests attacked by MPB in Colorado and then simulated stand growth over 200 years using the Forest Vegetation Simulator. We compared scenarios of no disturbance with scenarios of MPB-caused mortality, both with and without regeneration. Results indicated that basal area and tree density recovered to predisturbance levels relatively rapidly (within 1‐8 decades) in both forest types. However, convergence of the disturbed conditions with simulated undisturbed conditions took longer (12‐20+ decades) and was delayed by the absence of regeneration. In MPB-affected ponderosa pine forests without regeneration, basal area did not converge with undisturbed conditions within 200 years, implying lower resilience in this ecosystem. Surface fuels accumulated rapidly in both forest types after MPB-induced mortality, remaining high for 3‐6 decades in simulations. Our results suggest that future patterns of succession, regeneration, fuel loading, climate, and disturbance interactions over long time periods should be considered in management strategies addressing MPB effects in either forest type, but particularly in ponderosa pine.
Huang, Jian-Guo; Stadt, Kenneth J; Dawson, Andria; Comeau, Philip G
2013-01-01
We examined the effect of competition on stem growth of Picea glauca and Populus tremuloides in boreal mixedwood stands during the stem exclusion stage. We combined traditional approaches of collecting competition data with dendrochronology to provide retrospective measurements of stem diameter growth. Several competition indices including stand basal area (BA), the sum of stem diameter at breast height (SDBH), and density (N) for the broadleaf and coniferous species, as well as similar indices considering only trees with diameters greater than each subject (BAGR, SDBHGR, and NGR), were evaluated. We used a nonlinear mixed model to characterize the basal area increment over the past 5, 10, 15, 20, 25, 30, and 35 years as a function of growth of nearby dominant trees, the size of the subject trees, deciduous and coniferous competition indices, and ecoregions. SDBHGR and BAGR were better predictors for spruce, and SDBHGR and NGR were better for aspen, respectively, than other indices. Results showed strongest correlations with long-term stem growth, as the best models integrated growth for 10-25 years for aspen and ≥ 25 for spruce. Our model demonstrated a remarkable capability (adjusted R(2)>0.67) to represent this complex variation in growth as a function of site, size and competition.
Huang, Jian-Guo; Stadt, Kenneth J.; Dawson, Andria; Comeau, Philip G.
2013-01-01
We examined the effect of competition on stem growth of Picea glauca and Populus tremuloides in boreal mixedwood stands during the stem exclusion stage. We combined traditional approaches of collecting competition data with dendrochronology to provide retrospective measurements of stem diameter growth. Several competition indices including stand basal area (BA), the sum of stem diameter at breast height (SDBH), and density (N) for the broadleaf and coniferous species, as well as similar indices considering only trees with diameters greater than each subject (BAGR, SDBHGR, and NGR), were evaluated. We used a nonlinear mixed model to characterize the basal area increment over the past 5, 10, 15, 20, 25, 30, and 35 years as a function of growth of nearby dominant trees, the size of the subject trees, deciduous and coniferous competition indices, and ecoregions. SDBHGR and BAGR were better predictors for spruce, and SDBHGR and NGR were better for aspen, respectively, than other indices. Results showed strongest correlations with long-term stem growth, as the best models integrated growth for 10–25 years for aspen and ≥25 for spruce. Our model demonstrated a remarkable capability (adjusted R2>0.67) to represent this complex variation in growth as a function of site, size and competition. PMID:24204891
A local basal area adjustment for crown width prediction
Don C. Bragg
2001-01-01
Nonlinear crown width regressive equations were developed for 24 species common to the upper Lake States of Michigan, Minnesota, and Wisconsin. Of the species surveyed, 15 produced statistically significant (P 0.05) local basal area effect coefficients showing a reduction in crown...
Rain forest fragmentation and the proliferation of successional trees.
Laurance, William F; Nascimento, Henrique E M; Laurance, Susan G; Andrade, Ana C; Fearnside, Philip M; Ribeiro, José E L; Capretz, Robson L
2006-02-01
The effects of habitat fragmentation on diverse tropical tree communities are poorly understood. Over a 20-year period we monitored the density of 52 tree species in nine predominantly successional genera (Annona, Bellucia, Cecropia, Croton, Goupia, Jacaranda, Miconia, Pourouma, Vismia) in fragmented and continuous Amazonian forests. We also evaluated the relative importance of soil, topographic, forest dynamic, and landscape variables in explaining the abundance and species composition of successional trees. Data were collected within 66 permanent 1-ha plots within a large (approximately 1000 km2) experimental landscape, with forest fragments ranging from 1 to 100 ha in area. Prior to forest fragmentation, successional trees were uncommon, typically comprising 2-3% of all trees (> or =10 cm diameter at breast height [1.3 m above the ground surface]) in each plot. Following fragmentation, the density and basal area of successional trees increased rapidly. By 13-17 years after fragmentation, successional trees had tripled in abundance in fragment and edge plots and constituted more than a quarter of all trees in some plots. Fragment age had strong, positive effects on the density and basal area of successional trees, with no indication of a plateau in these variables, suggesting that successional species could become even more abundant in fragments over time. Nonetheless, the 52 species differed greatly in their responses to fragmentation and forest edges. Some disturbance-favoring pioneers (e.g., Cecropia sciadophylla, Vismia guianensis, V. amazonica, V. bemerguii, Miconia cf. crassinervia) increased by >1000% in density on edge plots, whereas over a third (19 of 52) of all species remained constant or declined in numbers. Species responses to fragmentation were effectively predicted by their median growth rate in nearby intact forest, suggesting that faster-growing species have a strong advantage in forest fragments. An ordination analysis revealed three main gradients in successional-species composition across our study area. Species gradients were most strongly influenced by the standlevel rate of tree mortality on each plot and by the number of nearby forest edges. Species-composition also varied significantly among different cattle ranches, which differed in their surrounding matrices and disturbance histories. These same variables were also the best predictors of total successional-tree abundance and species richness. Successional-tree assemblages in fragment interior plots (>150 m from edge), which are subjected to fragment area effects but not edge effects, did not differ significantly from those in intact forest, indicating that area effects per se had little influence on successional trees. Soils and topography also had little discernable effect on these species. Collectively, our results indicate that successional-tree species proliferate rapidly in fragmented Amazonian forests, largely as a result of chronically elevated tree mortality near forest edges and possibly an increased seed rain from successional plants growing in nearby degraded habitats. The proliferation of fast-growing successional trees and correlated decline of old-growth trees will have important effects on species composition, forest dynamics, carbon storage, and nutrient cycling in fragmented forests.
NASA Astrophysics Data System (ADS)
Manuri, Solichin; Andersen, Hans-Erik; McGaughey, Robert J.; Brack, Cris
2017-04-01
The airborne lidar system (ALS) provides a means to efficiently monitor the status of remote tropical forests and continues to be the subject of intense evaluation. However, the cost of ALS acquisition can vary significantly depending on the acquisition parameters, particularly the return density (i.e., spatial resolution) of the lidar point cloud. This study assessed the effect of lidar return density on the accuracy of lidar metrics and regression models for estimating aboveground biomass (AGB) and basal area (BA) in tropical peat swamp forests (PSF) in Kalimantan, Indonesia. A large dataset of ALS covering an area of 123,000 ha was used in this study. This study found that cumulative return proportion (CRP) variables represent a better accumulation of AGB over tree heights than height-related variables. The CRP variables in power models explained 80.9% and 90.9% of the BA and AGB variations, respectively. Further, it was found that low-density (and low-cost) lidar should be considered as a feasible option for assessing AGB and BA in vast areas of flat, lowland PSF. The performance of the models generated using reduced return densities as low as 1/9 returns per m2 also yielded strong agreement with the original high-density data. The use model-based statistical inferences enabled relatively precise estimates of the mean AGB at the landscape scale to be obtained with a fairly low-density of 1/4 returns per m2, with less than 10% standard error (SE). Further, even when very low-density lidar data was used (i.e., 1/49 returns per m2) the bias of the mean AGB estimates were still less than 10% with a SE of approximately 15%. This study also investigated the influence of different DTM resolutions for normalizing the elevation during the generation of forest-related lidar metrics using various return densities point cloud. We found that the high-resolution digital terrain model (DTM) had little effect on the accuracy of lidar metrics calculation in PSF. The accuracy of low-density lidar metrics in PSF was more influenced by the density of aboveground returns, rather than the last return. This is due to the flat topography of the study area. The results of this study will be valuable for future economical and feasible assessments of forest metrics over large areas of tropical peat swamp ecosystems.
Differential serotonergic innervation of the amygdala in bonobos and chimpanzees
Barger, Nicole; Taglialatela, Jared P.; Gendron-Fitzpatrick, Annette; Hof, Patrick R.; Hopkins, William D.; Sherwood, Chet C.
2016-01-01
Humans’ closest living relatives are bonobos (Pan paniscus) and chimpanzees (Pan troglodytes), yet these great ape species differ considerably from each other in terms of social behavior. Bonobos are more tolerant of conspecifics in competitive contexts and often use sexual behavior to mediate social interactions. Chimpanzees more frequently employ aggression during conflicts and actively patrol territories between communities. Regulation of emotional responses is facilitated by the amygdala, which also modulates social decision-making, memory and attention. Amygdala responsiveness is further regulated by the neurotransmitter serotonin. We hypothesized that the amygdala of bonobos and chimpanzees would differ in its neuroanatomical organization and serotonergic innervation. We measured volumes of regions and the length density of serotonin transporter-containing axons in the whole amygdala and its lateral, basal, accessory basal and central nuclei. Results showed that accessory basal nucleus volume was larger in chimpanzees than in bonobos. Of particular note, the amygdala of bonobos had more than twice the density of serotonergic axons than chimpanzees, with the most pronounced differences in the basal and central nuclei. These findings suggest that variation in serotonergic innervation of the amygdala may contribute to mediating the remarkable differences in social behavior exhibited by bonobos and chimpanzees. PMID:26475872
NASA Astrophysics Data System (ADS)
Soria, Ana R.; Liesa, Carlos L.; Mata, Maria Pilar; Arz, José A.; Alegret, Laia; Arenillas, Ignacio; Meléndez, Alfonso
2001-03-01
Slumps affecting uppermost Méndez Formation marls, as well as the spherulitic layer and basal part of the sandy deposits of the Cretaceous-Tertiary (K-T) boundary clastic unit, are described at the new K-T El Tecolote section (northeastern Mexico). These K-T clastic deposits represent sedimentation at middle-bathyal water depths in channel and nonchannel or levee areas of reworked materials coming from environments ranging from outer shelf to shallower slope via a unidirectional, high- to low-density turbidite flow. We emphasize the development and accretion of a lateral bar in a channel area from a surging low-density turbidity current and under a high-flow regime. The slumps discovered on land and the sedimentary processes of the K-T clastic unit reflect destabilization and collapse of the continental margin, support the mechanism of gravity flows in the deep sea, and represent important and extensive evidence for the impact effects in the Gulf of México triggered by the Chicxulub event.
Parkinson’s disease as a system-level disorder
Caligiore, Daniele; Helmich, Rick C; Hallett, Mark; Moustafa, Ahmed A; Timmermann, Lars; Toni, Ivan; Baldassarre, Gianluca
2016-01-01
Traditionally, the basal ganglia have been considered the main brain region implicated in Parkinson’s disease. This single area perspective gives a restricted clinical picture and limits therapeutic approaches because it ignores the influence of altered interactions between the basal ganglia and other cerebral components on Parkinsonian symptoms. In particular, the basal ganglia work closely in concert with cortex and cerebellum to support motor and cognitive functions. This article proposes a theoretical framework for understanding Parkinson’s disease as caused by the dysfunction of the entire basal ganglia–cortex–cerebellum system rather than by the basal ganglia in isolation. In particular, building on recent evidence, we propose that the three key symptoms of tremor, freezing, and impairments in action sequencing may be explained by considering partially overlapping neural circuits including basal ganglia, cortical and cerebellar areas. Studying the involvement of this system in Parkinson’s disease is a crucial step for devising innovative therapeutic approaches targeting it rather than only the basal ganglia. Possible future therapies based on this different view of the disease are discussed. PMID:28725705
John B Bradford; David M Bell
2016-01-01
Increasing aridity as a result of climate change is expected to exacerbate tree mortality. Reducing forest basal area â the cross-sectional area of tree stems within a given ground area â can decrease tree competition, which may reduce drought-induced tree mortality. However, neither the magnitude of expected mortality increases, nor the potential effectiveness of...
NASA Astrophysics Data System (ADS)
Mayes, Marc; Mustard, John; Melillo, Jerry; Neill, Christopher; Nyadzi, Gerson
2017-08-01
In sub-Saharan Africa (SSA), tropical dry forests and savannas cover over 2.5 million km2 and support livelihoods for millions in fast-growing nations. Intensifying land use pressures have driven rapid changes in tree cover structure (basal area, biomass) that remain poorly characterized at regional scales. Here, we posed the hypothesis that tree cover structure related strongly to senesced and non-photosynthetic (NPV) vegetation features in a SSA tropical dry forest landscape, offering improved means for satellite remote sensing of tree cover structure compared to vegetation greenness-based methods. Across regrowth miombo woodland sites in Tanzania, we analyzed relationships among field data on tree structure, land cover, and satellite indices of green and NPV features based on spectral mixture analyses and normalized difference vegetation index calculated from Landsat 8 data. From satellite-field data relationships, we mapped regional basal area and biomass using NPV and greenness-based metrics, and compared map performances at landscape scales. Total canopy cover related significantly to stem basal area (r 2 = 0.815, p < 0.01) and biomass (r 2 = 0.635, p < 0.01), and NPV dominated ground cover (> 60%) at all sites. From these two conditions emerged a key inverse relationship: skyward exposure of NPV ground cover was high at sites with low tree basal area and biomass, and decreased with increasing stem basal area and biomass. This pattern scaled to Landsat NPV metrics, which showed strong inverse correlations to basal area (Pearson r = -0.85, p < 0.01) and biomass (r = -0.86, p < 0.01). Biomass estimates from Landsat NPV-based maps matched field data, and significantly differentiated landscape gradients in woody biomass that greenness metrics failed to track. The results suggest senesced vegetation metrics at Landsat scales are a promising means for improved monitoring of tree structure across disturbance and ecological gradients in African and other tropical dry forests.
Determination of the Thermal Noise Limit of Graphene Biotransistors.
Crosser, Michael S; Brown, Morgan A; McEuen, Paul L; Minot, Ethan D
2015-08-12
To determine the thermal noise limit of graphene biotransistors, we have measured the complex impedance between the basal plane of single-layer graphene and an aqueous electrolyte. The impedance is dominated by an imaginary component but has a finite real component. Invoking the fluctuation-dissipation theorem, we determine the power spectral density of thermally driven voltage fluctuations at the graphene/electrolyte interface. The fluctuations have 1/f(p) dependence, with p = 0.75-0.85, and the magnitude of fluctuations scales inversely with area. Our results explain noise spectra previously measured in liquid-gated suspended graphene devices and provide realistic targets for future device performance.
Evidence for a basal temporal visual language center: cortical stimulation producing pure alexia.
Mani, J; Diehl, B; Piao, Z; Schuele, S S; Lapresto, E; Liu, P; Nair, D R; Dinner, D S; Lüders, H O
2008-11-11
Dejerine and Benson and Geschwind postulated disconnection of the dominant angular gyrus from both visual association cortices as the basis for pure alexia, emphasizing disruption of white matter tracts in the dominant temporooccipital region. Recently functional imaging studies provide evidence for direct participation of basal temporal and occipital cortices in the cognitive process of reading. The exact location and function of these areas remain a matter of debate. To confirm the participation of the basal temporal region in reading. Extraoperative electrical stimulation of the dominant hemisphere was performed in three subjects using subdural electrodes, as part of presurgical evaluation for refractory epilepsy. Pure alexia was reproduced during cortical stimulation of the dominant posterior fusiform and inferior temporal gyri in all three patients. Stimulation resulted in selective reading difficulty with intact auditory comprehension and writing. Reading difficulty involved sentences and words with intact letter by letter reading. Picture naming difficulties were also noted at some electrodes. This region is located posterior to and contiguous with the basal temporal language area (BTLA) where stimulation resulted in global language dysfunction in visual and auditory realms. The location corresponded with the visual word form area described on functional MRI. These observations support the existence of a visual language area in the dominant fusiform and occipitotemporal gyri, contiguous with basal temporal language area. A portion of visual language area was exclusively involved in lexical processing while the other part of this region processed both lexical and nonlexical symbols.
Ground-water status report, Pearl Harbor area, Hawaii, 1978
Soroos, Ronald L.; Ewart, Charles J.
1979-01-01
Increasing demand for freshwater in Hawaii has placed heavy stress on many of the State 's basal aquifer systems. The most heavily stressed of these systems is the Pearl Harbor on Oahu. The Pearl Harbor basal aquifer supplies as much as 277 million gallons per day. Since early in this century, spring discharge has been declining while pumpage has been increasing. Total ground-water discharge has remained steady despite short-term fluctuations. Some wells show general increases in chloride concentration while others remain steady. Chloride concentrations throughout the area show no apparent increase since 1970. Basal water head maps of the Pearl Harbor area clearly reflect the natural discharge points, which are the springs located along the shore near the center of Pearl Harbor. Basal-water hydrographs show a general decline of about 0.09 foot per year. This implies depletion of storage at a rate of about 25 million gallons per day. (USGS).
Busing, Richard T.; Solomon, Allen M.
2005-01-01
An individual-based model of forest dynamics (FORCLIM) was tested for its ability to simulate forest composition and structure in the Pacific Northwest region of North America. Simulation results across gradients of climate and disturbance were compared to forest survey data from several vegetation zones in western Oregon. Modelled patterns of tree species composition, total basal area and stand height across climate gradients matched those in the forest survey data. However, the density of small stems (<50 cm DBH) was underestimated by the model. Thus actual size-class structure and other density-based parameters of stand structure were not simulated with high accuracy. The addition of partial-stand disturbances at moderate frequencies (<0.01 yr-1) often improved agreement between simulated and actual results. Strengths and weaknesses of the FORCLIM model in simulating forest dynamics and structure in the Pacific Northwest are discussed.
NASA Astrophysics Data System (ADS)
Roche, J. W.; Goulden, M.; Bales, R. C.
2017-12-01
Increased forest evapotranspiration (ET) coupled with snowpack decreases in a warming climate is likely to decrease runoff and increase forest drought stress. Field experiments and modeling suggest that forest thinning can reduce ET and thus increase potential runoff relative to untreated areas. We investigated the potential magnitude and duration of ET decreases resulting from forest-thinning treatments and fire using a robust empirical relation between Landsat-derived mean-annual normalized difference vegetation index (NDVI) and annual ET measured at flux towers. Among forest treatments, the minimum observed NDVI change required to produce a significant departure from control plots with NDVI of about 0.70 was -0.07 units, corresponding to a basal-area reduction of 3.1 m2 ha-1, and equivalent to an estimated ET reduction of -102 mm yr-1. Intensive thinning in highly productive forests that approached pre-fire-exclusion densities reduced basal area by 40-50%, generating estimated ET reductions of 152-216 mm yr-1 over five years following treatment. Between 1990 and 2008, fires in the American River basin generated more than twice the ET reduction per unit area than those in the Kings River basin, corresponding to greater water and energy limitations in the latter and greater fire severity in the former. A rough extrapolation of these results to the entire American River watershed, much of which would have burned naturally during this 19-year period, could result in ET reductions that approach 10% of full natural flows for drought years and 5% averaged over all years. This work demonstrates the potential utility to estimate forest ET change at the patch scale, which in turn may allow managers to estimate thinning benefits in areas lacking detailed hydrologic measurements.
Smitley, D R; Rebek, E J; Royalty, R N; Davis, T W; Newhouse, K F
2010-02-01
We conducted field trials at five different locations over a period of 6 yr to investigate the efficacy of imidacloprid applied each spring as a basal soil drench for protection against emerald ash borer, Agrilus planipennis Fairmaire (Coleoptera: Buprestidae). Canopy thinning and emerald ash borer larval density were used to evaluate efficacy for 3-4 yr at each location while treatments continued. Test sites included small urban trees (5-15 cm diameter at breast height [dbh]), medium to large (15-65 cm dbh) trees at golf courses, and medium to large street trees. Annual basal drenches with imidacloprid gave complete protection of small ash trees for three years. At three sites where the size of trees ranged from 23 to 37 cm dbh, we successfully protected all ash trees beginning the test with <60% canopy thinning. Regression analysis of data from two sites reveals that tree size explains 46% of the variation in efficacy of imidacloprid drenches. The smallest trees (<30 cm dbh) remained in excellent condition for 3 yr, whereas most of the largest trees (>38 cm dbh) declined to a weakened state and undesirable appearance. The five-fold increase in trunk and branch surface area of ash trees as the tree dbh doubles may account for reduced efficacy on larger trees, and suggests a need to increase treatment rates for larger trees.
Basal area increment and growth efficiency as functions of canopy dynamics and stem mechanics
Thomas J. Dean
2004-01-01
Crown and canopy structurecorrelate with growth efficiency and also determine stem size and taper as described by the uniform stress principle of stem formation. A regression model was derived from this principle that expresses basal area increment in terms of the amount and vertical distribution of leaf area and change in these variables during a growth period. This...
Modeled distributions of 12 tree species in New York
Rachel I. Riemann; Barry T. Wilson; Andrew J. Lister; Oren Cook; Sierra Crane-Murdoch
2014-01-01
These maps depict the distribution of 12 tree species across the state of New York. The maps show where these trees do not occur (gray), occasionally occur (pale green), are a minor component (medium green), are a major component (dark green), or are the dominant species (black) in the forest, as determined by that species' total basal area. Basal area is the area...
Thompson, Christopher K.; Schwabe, Fabian; Schoof, Alexander; Mendoza, Ezequiel; Gampe, Jutta; Rochefort, Christelle; Scharff, Constance
2013-01-01
FOXP2 is a transcription factor functionally relevant for learned vocalizations in humans and songbirds. In songbirds, FoxP2 mRNA expression in the medium spiny neurons of the basal ganglia song nucleus Area X is developmentally regulated and varies with singing conditions in different social contexts. How individual neurons in Area X change FoxP2 expression across development and in social contexts is not known, however. Here we address this critical gap in our understanding of FoxP2 as a link between neuronal networks and behavior. We used a statistically unbiased analysis of FoxP2-immunoreactivity (FoxP2-IR) on a neuron-by-neuron basis and found a bimodal distribution of FoxP2-IR neurons in Area X: weakly-stained and intensely-stained. The density of intensely-stained FoxP2-IR neurons was 10 times higher in juveniles than in adults, exponentially decreased with age, and was negatively correlated with adult song stability. Three-week old neurons labeled with BrdU were more than five times as likely to be intensely-stained than weakly-stained. The density of FoxP2-IR putative migratory neurons with fusiform-shaped nuclei substantially decreased as birds aged. The density of intensely-stained FoxP2-IR neurons was not affected by singing whereas the density of weakly-stained FoxP2-IR neurons was. Together, these data indicate that young Area X medium spiny neurons express FoxP2 at high levels and decrease expression as they become integrated into existing neural circuits. Once integrated, levels of FoxP2 expression correlate with singing behavior. Together, these findings raise the possibility that FoxP2 levels may orchestrate song learning and song stereotypy in adults by a common mechanism. PMID:23450800
Thompson, Christopher K; Schwabe, Fabian; Schoof, Alexander; Mendoza, Ezequiel; Gampe, Jutta; Rochefort, Christelle; Scharff, Constance
2013-01-01
FOXP2 is a transcription factor functionally relevant for learned vocalizations in humans and songbirds. In songbirds, FoxP2 mRNA expression in the medium spiny neurons of the basal ganglia song nucleus Area X is developmentally regulated and varies with singing conditions in different social contexts. How individual neurons in Area X change FoxP2 expression across development and in social contexts is not known, however. Here we address this critical gap in our understanding of FoxP2 as a link between neuronal networks and behavior. We used a statistically unbiased analysis of FoxP2-immunoreactivity (FoxP2-IR) on a neuron-by-neuron basis and found a bimodal distribution of FoxP2-IR neurons in Area X: weakly-stained and intensely-stained. The density of intensely-stained FoxP2-IR neurons was 10 times higher in juveniles than in adults, exponentially decreased with age, and was negatively correlated with adult song stability. Three-week old neurons labeled with BrdU were more than five times as likely to be intensely-stained than weakly-stained. The density of FoxP2-IR putative migratory neurons with fusiform-shaped nuclei substantially decreased as birds aged. The density of intensely-stained FoxP2-IR neurons was not affected by singing whereas the density of weakly-stained FoxP2-IR neurons was. Together, these data indicate that young Area X medium spiny neurons express FoxP2 at high levels and decrease expression as they become integrated into existing neural circuits. Once integrated, levels of FoxP2 expression correlate with singing behavior. Together, these findings raise the possibility that FoxP2 levels may orchestrate song learning and song stereotypy in adults by a common mechanism.
Hu, Ya-Lin; Hu, Li-Le; Zeng, De-Hui
2014-01-01
Afforestation of former croplands has been proposed as a promising way to mitigate rising atmospheric CO2 concentration in view of the commitment to the Kyoto Protocol. Central to this C sequestration is the dynamics of soil organic C (SOC) storage and stability with the development of afforested plantations. Our previous study showed that SOC storage was not changed after afforestation except for the 0–10 cm layer in a semi-arid region of Keerqin Sandy Lands, northeast China. In this study, soil organic C was further separated into light and heavy fractions using the density fractionation method, and their organic C concentration and 13C signature were analyzed to investigate the turnover of old vs. new SOC in the afforested soils. Surface layer (0–10 cm) soil samples were collected from 14 paired plots of poplar (Populus × xiaozhuanica W. Y. Hsu & Liang) plantations with different stand basal areas (the sum of the cross-sectional area of all live trees in a stand), ranging from 0.2 to 32.6 m2 ha−1, and reference maize (Zea mays L.) croplands at the same sites as our previous study. Soil ΔC stocks (ΔC refers to the difference in SOC content between a poplar plantation and the paired cropland) in bulk soil and light fraction were positively correlated with stand basal area (R 2 = 0.48, p<0.01 and R 2 = 0.40, p = 0.02, respectively), but not for the heavy fraction. SOCcrop (SOC derived from crops) contents in the light and heavy fractions in poplar plantations were significantly lower as compared with SOC contents in croplands, but tree-derived C in bulk soil, light and heavy fraction pools increased gradually with increasing stand basal area after afforestation. Our study indicated that cropland afforestation could sequester new C derived from trees into surface mineral soil, but did not enhance the stability of SOC due to a fast turnover of SOC in this semi-arid region. PMID:24466183
Bechard, Allison R.; Cacodcar, Nadia; King, Michael A.; Lewis, Mark H.
2015-01-01
Repetitive motor behaviors are observed in many neurodevelopmental and neurological disorders (e.g. autism spectrum disorders, Tourette syndrome, fronto-temporal dementia). Despite their clinical importance, the neurobiology underlying these highly stereotyped, apparently functionless behaviors is poorly understood. Identification of mechanisms that mediate the development of repetitive behaviors will aid in the discovery of new therapeutic targets and treatment development. Using a deer mouse model, we have shown that decreased indirect basal ganglia pathway activity is associated with high levels of repetitive behavior. Environmental enrichment (EE) markedly attenuates the development of such aberrant behaviors in mice, although mechanisms driving this effect are unknown. We hypothesized that EE would reduce repetitive motor behaviors by increasing indirect basal ganglia pathway function. We assessed neuronal activation and dendritic spine density in basal ganglia of adult deer mice reared in EE and standard housing. Significant increases in neuronal activation and dendritic spine densities were observed only in the subthalamic nucleus (STN) and globus pallidus (GP), and only for those mice that exhibited an EE-induced decrease in repetitive motor behavior. As the STN and GP lie within the indirect pathway, these data suggest that EE-induced attenuation of repetitive motor behaviors is associated with increased functional activation of the indirect basal ganglia pathway. These results are consistent with our other findings highlighting the importance of the indirect pathway in mediating repetitive motor behaviors. PMID:26620495
Swenson, J.B.; Person, M.; Raffensperger, Jeff P.; Cannon, W.F.; Woodruff, L.G.; Berndt, M.E.
2004-01-01
This paper presents a suite of two-dimensional mathematical models of basin-scale groundwater flow and heat transfer for the middle Proterozoic Midcontinent Rift System. The models were used to assess the hydrodynamic driving mechanisms responsible for main-stage stratiform copper mineralization of the basal Nonesuch Formation during the post-volcanic/pre-compressional phase of basin evolution. Results suggest that compaction of the basal aquifer (Copper Harbor Formation), in response to mechanical loading during deposition of the overlying Freda Sandstone, generated a pulse of marginward-directed, compaction-driven discharge of cupriferous brines from within the basal aquifer. The timing of this pulse is consistent with the radiometric dates for the timing of mineralization. Thinning of the basal aquifer near White Pine, Michigan, enhanced stratiform copper mineralization. Focused upward leakage of copper-laden brines into the lowermost facies of the pyrite-rich Nonesuch Formation resulted in copper sulfide mineralization in response to a change in oxidation state. Economic-grade mineralization within the White Pine ore district is a consequence of intense focusing of compaction-driven discharge, and corresponding amplification of leakage into the basal Nonesuch Formation, where the basal aquifer thins dramatically atop the Porcupine Mountains volcanic structure. Equilibrium geochemical modeling and mass-balance calculations support this conclusion. We also assessed whether topography and density-driven flow systems could have caused ore genesis at White Pine. Topography-driven flow associated with the Ottawan orogeny was discounted because it post-dates main-stage ore genesis and because recent seismic interpretations of basin inversion indicates that basin geometry would not be conductive to ore genesis. Density-driven flow systems did not produce focused discharge in the vicinity of the White Pine ore district.
Kinze, S; Schöneberg, T; Meyer, R; Martin, H; Kaufmann, R
1996-10-11
In this paper, cholecystokinin (CCK) B-type binding sites were characterized with receptor binding studies in different human brain regions (various parts of cerebral cortex, basal ganglia, hippocampus, thalamus, cerebellar cortex) collected from 22 human postmortem brains. With the exception of the thalamus, where no specific CCK binding sites were found, a pharmacological characterization demonstrated a single class of high affinity CCK sites in all brain areas investigated. Receptor densities ranged from 0.5 fmol/mg protein (hippocampus) to 8.4 fmol/mg protein (nucleus caudatus). These CCK binding sites displayed a typical CCKA binding profile as shown in competition studies by using different CCK-related compounds and non peptide CCK antagonists discriminating between CCKA and CCKB sites. The rank order of agonist or antagonist potency in inhibiting specific sulphated [propionyl-3H]cholecystokinin octapeptide binding was similar and highly correlated for the brain regions investigated as demonstrated by a computer-assisted analysis. Therefore it is concluded that CCKB binding sites in human cerebral cortex, basal ganglia, cerebellar cortex share identical ligand binding characteristics.
Niemiec, Joanna A; Adamczyk, Agnieszka; Małecki, Krzysztof; Majchrzyk, Kaja; Ryś, Janusz
2012-12-01
There is still a lack of complete consensus on immunohistochemical surrogate markers for luminal A (LA) and luminal B (LB), HER2, and basal-like subtypes of breast carcinomas and their correlation with cancer cell adhesion and invasion-promoting factors. Therefore, early-stage invasive ductal breast cancer patients (N=209) were recruited to the study and divided into 4 subtypes, on the basis of the expression of the estrogen/progesterone receptor and HER2 (LA: 74.4% of cases; LB: 7.8%; HER2: 5.6%; and triple-negative phenotype: 12.2%). Regardless of the above-mentioned classification, we divided all carcinomas into 2 groups: carcinomas expressing at least 1 basal marker [cytokeratine (CK)5/6, CK5, vimentin, epidermal growth factor receptor, or aberrant CK8/18 expression-membranous or in <10% of cells] versus carcinomas negative for basal markers. Then we studied the relationships between the above subtypes (2 classifications) and (i) the expression of adhesion molecules (Ep-CAM, P-cadherin), (ii) matrix metalloproteinases (MMP)-2, (iii) the proliferation index (MIB-1 LI), and (iv) the microvascular density. We confirmed that triple-negative phenotypes are characterized by basal marker expression, a high tumor grade, and high MIB-1 LI. In this subtype, we found MMP-2 expression in stromal leukocytes less frequently. Both LA carcinomas and carcinomas negative for basal markers were more often negative for epithelial cell adhesion molecule (Ep-CAM) and P-cadherin. Moreover, we noted a higher mean value of microvascular density in CK5/6 and Ep-CAM-immunopositive tumors, carcinomas with aberrant CK8/18 expression, and carcinomas with no or strong expression of MMP-2 in stromal fibroblast-like cells. These results might suggest that mechanisms of stroma remodeling and carcinogenesis (Ep-CAM is the suggested marker of breast progenitors) may differ between breast cancer subtypes.
Wallesch, C W; Brunner, R J; Seemüller, E
1983-12-01
Repetitive phenomena in spontaneous speech were investigated in 30 patients with chronic infarctions of the left hemisphere which included Broca's and/or Wernicke's area and/or the basal ganglia. Perseverations, stereotypies, and echolalias occurred with all types of brain lesions, automatisms and recurring utterances only with those patients, whose infarctions involved Wernicke's area and basal ganglia. These patients also showed more echolalic responses. The results are discussed in view of the role of the basal ganglia as motor program generators.
2010-10-17
conditions between active, recently active and randomly selected non-burrow locations at this site. Field surveys were completed in three study areas ...at the installation. On average, burrow sites had a much higher overall occurrence of longleaf pine and significantly lower total basal area as...East Area burrow and non-burrow locations, Camp Shelby, MS 2007 .................. 10 2 Mean basal area per acre for pine and hardwood on T-44, Mars
Coleman, Leon G.; He, Jun; Lee, Joohwi; Styner, Martin; Crews, Fulton T.
2013-01-01
Background Binge-drinking is common in human adolescents. The adolescent brain is undergoing structural maturation and has a unique sensitivity to alcohol neurotoxicity. Therefore, adolescent binge ethanol may have long-term effects on the adult brain that alter brain structure and behaviors that are relevant to alcohol use disorders. Methods In order to determine if adolescent ethanol binge drinking alters the adult brain, male C57BL/6 mice were treated with either water or ethanol during adolescence (5g/kg/day i.g., post-natal days P28-37) and assessed during adulthood (P60-P88). An array of neurotransmitter-specific genes, behavioral tests (i.e. reversal learning, prepulse inhibition, and open field), and post-mortem brain structure using MRI and immunohistochemistry, were employed to assess persistent alterations in adult brain. Results At P38, 24 hours after adolescent ethanol (AE) binge, many neurotransmitter genes, particularly cholinergic and dopaminergic, were reduced by ethanol treatment. Interestingly, dopamine receptor type 4 mRNA was reduced and confirmed using immunohistochemistry. Normal control maturation (P38-P88) resulted in decreased neurotransmitter mRNA, e.g. an average decrease of 56%. Following adolescent ethanol treatment, adults showed greater gene expression reductions than controls, averaging 73%. Adult spatial learning assessed in the Morris water maze was not changed by adolescent ethanol treatment, but reversal learning experiments revealed deficits. Assessment of adult brain region volumes using MRI indicated that the olfactory bulb and basal forebrain were smaller in adults following adolescent ethanol. Immunohistochemical analyses found reduced basal forebrain area and fewer basal forebrain cholinergic neurons. Conclusions Adolescent binge ethanol treatment reduces adult neurotransmitter gene expression, particularly cholinergic genes, reduces basal forebrain and olfactory bulb volumes, and causes a reduction in the density of basal forebrain acetylcholine neurons. Loss of cholinergic neurons and forebrain structure could underlie adult reversal learning deficits following adolescent binge drinking. PMID:21223304
Differential serotonergic innervation of the amygdala in bonobos and chimpanzees.
Stimpson, Cheryl D; Barger, Nicole; Taglialatela, Jared P; Gendron-Fitzpatrick, Annette; Hof, Patrick R; Hopkins, William D; Sherwood, Chet C
2016-03-01
Humans' closest living relatives are bonobos (Pan paniscus) and chimpanzees (Pan troglodytes), yet these great ape species differ considerably from each other in terms of social behavior. Bonobos are more tolerant of conspecifics in competitive contexts and often use sexual behavior to mediate social interactions. Chimpanzees more frequently employ aggression during conflicts and actively patrol territories between communities. Regulation of emotional responses is facilitated by the amygdala, which also modulates social decision-making, memory and attention. Amygdala responsiveness is further regulated by the neurotransmitter serotonin. We hypothesized that the amygdala of bonobos and chimpanzees would differ in its neuroanatomical organization and serotonergic innervation. We measured volumes of regions and the length density of serotonin transporter-containing axons in the whole amygdala and its lateral, basal, accessory basal and central nuclei. Results showed that accessory basal nucleus volume was larger in chimpanzees than in bonobos. Of particular note, the amygdala of bonobos had more than twice the density of serotonergic axons than chimpanzees, with the most pronounced differences in the basal and central nuclei. These findings suggest that variation in serotonergic innervation of the amygdala may contribute to mediating the remarkable differences in social behavior exhibited by bonobos and chimpanzees. © The Author (2015). Published by Oxford University Press. For Permissions, please email: journals.permissions@oup.com.
Arbib, Michael A.; Baldassarre, Gianluca
2017-01-01
Motor tics are a cardinal feature of Tourette syndrome and are traditionally associated with an excess of striatal dopamine in the basal ganglia. Recent evidence increasingly supports a more articulated view where cerebellum and cortex, working closely in concert with basal ganglia, are also involved in tic production. Building on such evidence, this article proposes a computational model of the basal ganglia-cerebellar-thalamo-cortical system to study how motor tics are generated in Tourette syndrome. In particular, the model: (i) reproduces the main results of recent experiments about the involvement of the basal ganglia-cerebellar-thalamo-cortical system in tic generation; (ii) suggests an explanation of the system-level mechanisms underlying motor tic production: in this respect, the model predicts that the interplay between dopaminergic signal and cortical activity contributes to triggering the tic event and that the recently discovered basal ganglia-cerebellar anatomical pathway may support the involvement of the cerebellum in tic production; (iii) furnishes predictions on the amount of tics generated when striatal dopamine increases and when the cortex is externally stimulated. These predictions could be important in identifying new brain target areas for future therapies. Finally, the model represents the first computational attempt to study the role of the recently discovered basal ganglia-cerebellar anatomical links. Studying this non-cortex-mediated basal ganglia-cerebellar interaction could radically change our perspective about how these areas interact with each other and with the cortex. Overall, the model also shows the utility of casting Tourette syndrome within a system-level perspective rather than viewing it as related to the dysfunction of a single brain area. PMID:28358814
Caligiore, Daniele; Mannella, Francesco; Arbib, Michael A; Baldassarre, Gianluca
2017-03-01
Motor tics are a cardinal feature of Tourette syndrome and are traditionally associated with an excess of striatal dopamine in the basal ganglia. Recent evidence increasingly supports a more articulated view where cerebellum and cortex, working closely in concert with basal ganglia, are also involved in tic production. Building on such evidence, this article proposes a computational model of the basal ganglia-cerebellar-thalamo-cortical system to study how motor tics are generated in Tourette syndrome. In particular, the model: (i) reproduces the main results of recent experiments about the involvement of the basal ganglia-cerebellar-thalamo-cortical system in tic generation; (ii) suggests an explanation of the system-level mechanisms underlying motor tic production: in this respect, the model predicts that the interplay between dopaminergic signal and cortical activity contributes to triggering the tic event and that the recently discovered basal ganglia-cerebellar anatomical pathway may support the involvement of the cerebellum in tic production; (iii) furnishes predictions on the amount of tics generated when striatal dopamine increases and when the cortex is externally stimulated. These predictions could be important in identifying new brain target areas for future therapies. Finally, the model represents the first computational attempt to study the role of the recently discovered basal ganglia-cerebellar anatomical links. Studying this non-cortex-mediated basal ganglia-cerebellar interaction could radically change our perspective about how these areas interact with each other and with the cortex. Overall, the model also shows the utility of casting Tourette syndrome within a system-level perspective rather than viewing it as related to the dysfunction of a single brain area.
Thinning regimes and initial spacing for Eucalyptus plantations in Brazil.
Ferraz Filho, Antonio C; Mola-Yudego, Blas; González-Olabarria, José R; Scolforo, José Roberto S
2018-01-01
This study focuses on the effects of different thinning regimes on clonal Eucalyptus plantations growth. Four different trials, planted in 1999 and located in Bahia and Espírito Santo States, were used. Aside from thinning, initial planting density, and post thinning fertilization application were also evaluated. Before canopy closure, and therefore before excessive competition between trees took place, it was found that stands planted under low densities (667 trees per hectare) presented a lower mortality proportion when compared to stand planted under higher densities (1111 trees per hectare). However, diameter growth prior to thinning operations was not statistically different between these two densities, presenting an overall mean of 4.9 cm/year. After canopy closure and the application of the thinning treatments, it was found that thinning regimes beginning early in the life of the stand and leaving a low number of residual trees presented the highest diameter and height growth. Unthinned treatments and thinning regimes late in the life of the stand (after 5.5 years), leaving a large number of residual trees presented the highest values of basal area production. The choice of the best thinning regime for Eucalyptus clonal material will vary according to the plantation objective.
Basal area growth impacts of dwarf misletoe on western hemlock in an old-growth forest
David C. Shaw; Huso Manuela; Howard Bruner
2008-01-01
We investigated the effect of western hemlock dwarf mistletoe (Arceuthobium tsugense (Rosend.) G.N. Jones ssp. tsugense) on the 13-year basal area growth of large (>45.7 cm diameter at breast height) western hemlock (Tsuga heterophylla (Raf.) Sarg.) trees in an old-growth forest in southwestern Washington...
Impacts of Oak Decline on Forest Structure in Arkansas and Oklahoma: Preliminary Results
Eric Heitzman; James M. Guldin
2004-01-01
We established field plots in the Ouachita and Ozark Mountains of Arkansas and Oklahoma to quantify the impacts of oak decline on forest structure. Plots were identified as either high risk (red oak basal area > 20 square feet per acre) or low risk (red oak basal area
Hamberg, Leena; Lehvävirta, Susanna; Kotze, D Johan; Heikkinen, Juha
2015-03-15
Recent studies have shown a considerable increase in the abundance of rowan (Sorbus aucuparia) saplings in urban forests in Finland, yet the reasons for this increase are not well understood. Here we investigated whether canopy cover or tree species composition, i.e., the basal areas of different tree species in Norway spruce dominated urban forests, affects the abundances of rowan seedlings, saplings and trees. Altogether 24 urban forest patches were investigated. We sampled the number of rowan and other saplings, and calculated the basal areas of trees. We showed that rowan abundance was affected by tree species composition. The basal area of rowan trees (≥ 5 cm in diameter at breast height, dbh) decreased with increasing basal area of Norway spruce, while the cover of rowan seedlings increased with an increase in Norway spruce basal area. However, a decrease in the abundance of birch (Betula pendula) and an increase in the broad-leaved tree group (Acer platanoides, Alnus glutinosa, Alnus incana, Amelanchier spicata, Prunus padus, Quercus robur, Rhamnus frangula and Salix caprea) coincided with a decreasing number of rowans. Furthermore, rowan saplings were scarce in the vicinity of mature rowan trees. Although it seems that tree species composition has an effect on rowan, the relationship between rowan saplings and mature trees is complex, and therefore we conclude that regulating tree species composition is not an easy way to keep rowan thickets under control in urban forests in Finland. Copyright © 2015 Elsevier Ltd. All rights reserved.
Growth phenology of coast Douglas-fir seed sources planted in diverse environments.
Gould, Peter J; Harrington, Constance A; St Clair, J Bradley
2012-12-01
The timing of periodic life cycle events in plants (phenology) is an important factor determining how species and populations will react to climate change. We evaluated annual patterns of basal-area and height growth of coast Douglas-fir (Pseudotusga menziesii var. menziesii (Mirb.) Franco) seedlings from four seed sources that were planted in four diverse environments as part of the Douglas-fir Seed-Source Movement Trial. Stem diameters and heights were measured periodically during the 2010 growing season on 16 open-pollinated families at each study installation. Stem diameters were measured on a subset of trees with electronic dendrometers during the 2010 and 2011 growing seasons. Trees from the four seed sources differed in phenology metrics that described the timing of basal-area and height-growth initiation, growth cessation and growth rates. Differences in the height-growth metrics were generally larger than differences in the basal-area growth metrics and differences among installations were larger than differences among seed sources, highlighting the importance of environmental signals on growth phenology. Variations in the height- and basal-area growth metrics were correlated with different aspects of the seed-source environments: precipitation in the case of height growth and minimum temperature in the case of basal-area growth. The detailed dendrometer measurements revealed differences in growth patterns between seed sources during distinct periods in the growing season. Our results indicate that multiple aspects of growth phenology should be considered along with other traits when evaluating adaptation of populations to future climates.
The scaling of maximum and basal metabolic rates of mammals and birds
NASA Astrophysics Data System (ADS)
Barbosa, Lauro A.; Garcia, Guilherme J. M.; da Silva, Jafferson K. L.
2006-01-01
Allometric scaling is one of the most pervasive laws in biology. Its origin, however, is still a matter of dispute. Recent studies have established that maximum metabolic rate scales with an exponent larger than that found for basal metabolism. This unpredicted result sets a challenge that can decide which of the concurrent hypotheses is the correct theory. Here, we show that both scaling laws can be deduced from a single network model. Besides the 3/4-law for basal metabolism, the model predicts that maximum metabolic rate scales as M, maximum heart rate as M, and muscular capillary density as M, in agreement with data.
Zhao, Dong; Yan, Ting-mei; Qiao, Jun; Yang, Lin-zhang; Tang, Fang; Song, Yun-fei
2015-06-01
This study focused on the nitrogen loss via runoff, change of nitrogen in different forms in surface water in paddy field, and grain yield, through further reduction of nitrogen fertilizer application rate under green manuring without basal dressing. Results showed that with 150 kg · hm(-2) inorganic N fertilizer input after return of green manure to soil, no basal dressing could not only sharply reduce N concentration in surface water and decrease 17.2% of N loss, but also increase 2.8% of grain yield in comparison with basal dressing. It was a worthwhile farming method that inorganic nitrogen fertilizer was not used for basal dressing but for topdressing after return of green ma- nure to soil in Taihu Area. However, the grain yield would decrease if the rate of topdressing nitro- gen was excessively reduced or increased. After all, it was feasible to realize harmonization of grain yield and environmental benefits in Taihu Area, with 133 kg · hm(-2) inorganic N fertilizer input after return of green manure to soil as well as no application of basal dressing, which could greatly reduce N fertilizer input and N loss as well as ensure rice yield.
Caligiore, Daniele; Pezzulo, Giovanni; Baldassarre, Gianluca; Bostan, Andreea C; Strick, Peter L; Doya, Kenji; Helmich, Rick C; Dirkx, Michiel; Houk, James; Jörntell, Henrik; Lago-Rodriguez, Angel; Galea, Joseph M; Miall, R Chris; Popa, Traian; Kishore, Asha; Verschure, Paul F M J; Zucca, Riccardo; Herreros, Ivan
2017-02-01
Despite increasing evidence suggesting the cerebellum works in concert with the cortex and basal ganglia, the nature of the reciprocal interactions between these three brain regions remains unclear. This consensus paper gathers diverse recent views on a variety of important roles played by the cerebellum within the cerebello-basal ganglia-thalamo-cortical system across a range of motor and cognitive functions. The paper includes theoretical and empirical contributions, which cover the following topics: recent evidence supporting the dynamical interplay between cerebellum, basal ganglia, and cortical areas in humans and other animals; theoretical neuroscience perspectives and empirical evidence on the reciprocal influences between cerebellum, basal ganglia, and cortex in learning and control processes; and data suggesting possible roles of the cerebellum in basal ganglia movement disorders. Although starting from different backgrounds and dealing with different topics, all the contributors agree that viewing the cerebellum, basal ganglia, and cortex as an integrated system enables us to understand the function of these areas in radically different ways. In addition, there is unanimous consensus between the authors that future experimental and computational work is needed to understand the function of cerebellar-basal ganglia circuitry in both motor and non-motor functions. The paper reports the most advanced perspectives on the role of the cerebellum within the cerebello-basal ganglia-thalamo-cortical system and illustrates other elements of consensus as well as disagreements and open questions in the field.
Effects of thinning and herbicide application on vertebrate communities in longleaf pine plantations
DOE Office of Scientific and Technical Information (OSTI.GOV)
Brunjes, Kristina J.; Miller, Karl V.; Ford, Mark W.
Currently, nearly 98% of the land area once dominated by longleaf pine ecosystems has been converted to other uses. The U.S. Forest Service is replanting logged areas with longleaf pine at the Savannah River Site, New Ellenton, South Carolina, in an effort to restore these ecosystems. To ascertain the effects of various silvicultural management techniques on the vertebrate communities, we surveyed small mammal, herpetofaunal, and avian communities in six 10- to 13-year-old longleaf pine plantations subjected to various thinning and herbicide regimes. Areas within each plantation were randomly assigned one of four treatments: thinning, herbicide spraying, thinning and herbicide, andmore » an untreated control. For all vertebrate groups, abundance and species diversity tended to be less in the controls than treated areas. Birds and small mammals were most abundant and diverse in thinned treatments versus spray only and control. Herpetofauna capture rates were low and, thus, we were unable to detect treatment-related differences. Silvicultural treatments that reduce hardwood stem density and pine basal area can enhance habitat conditions for numerous vertebrate species.« less
Pradip Saud; Thomas B. Lynch; Duncan S. Wilson; John Stewart; James M. Guldin; Bob Heinemann; Randy Holeman; Dennis Wilson; Keith Anderson
2015-01-01
An individual-tree basal area growth model previously developed for even-aged naturally occurring shortleaf pine trees (Pinus echinata Mill.) in western Arkansas and southeastern Oklahoma did not include weather variables. Individual-tree growth and yield modeling of shortleaf pine has been carried out using the remeasurements of over 200 plots...
A Growth and Yield Model for Thinned Stands of Yellow-Poplar
Bruce R. Knoebel; Harold E. Burkhart; Donald E. Beck
1986-01-01
Simultaneous growth and yield equations were developed for predicting basal area growth and cubic-foot volume growth and yield in thinned stands of yellow-poplar. A joint loss function involving both volume and basal area was used to estimate the coefficients in the system of equations. The estimates obtained were analytically compatible, invariant for projection...
Experience with basal area estimation by prisms in lodgepole pine.
James M. Trappe
1957-01-01
Estimation of basal area by prisms offers intriguing possibilities for reducing time and effort in making stand inventories. Increased inventory efficiency is a particular need in stands that are relatively low in value due to small stems, predominance of low value species or heavy defect. In the Pacific Northwest, lodgepole pine characteristically forms dense low-...
Nonlinear mixed modeling of basal area growth for shortleaf pine
Chakra B. Budhathoki; Thomas B. Lynch; James M. Guldin
2008-01-01
Mixed model estimation methods were used to fit individual-tree basal area growth models to tree and stand-level measurements available from permanent plots established in naturally regenerated shortleaf pine (Pinus echinata Mill.) even-aged stands in western Arkansas and eastern Oklahoma in the USA. As a part of the development of a comprehensive...
Differentiation of anchoring junctions in tracheal basal cells in the growing rat.
Evans, M J; Cox, R A; Burke, A S; Moller, P C
1992-02-01
A function of airway basal cells is to attach ciliated and nonciliated columnar cells to the basal lamina. The significance of the basal cell in attachment is related to the height of the columnar epithelium. In taller epithelia, basal cells are more numerous and differentiated with respect to anchoring junctional adhesion mechanisms (desmosomes, hemidesmosomes, and the cytoskeleton) than in shorter epithelia. In this study, we determined if basal cell anchoring junctional adhesion mechanisms differentiated during growth of the airway. Tracheas from five 3-day-old, five 30-day-old, and five 90-day-old rats were prepared for electron microscopy and morphometrically studied by standard techniques. The circumference of the trachea increased from 2.5 +/- 0.2 to 7.5 +/- 0.4 mm during growth. The height of the columnar cell increased from 13.4 +/- 1.5 to 24.6 +/- 3.9 microns, and the number of basal cells per millimeter increased from 3.2 +/- 0.7 to 9.6 +/- 1.8 during growth. The number of desmosomes per basal cell profile increased significantly from 1.5 +/- 0.1 to 2.1 +/- 0.1, as did keratin filament volume density from 0.046 +/- 0.05 to 0.098 +/- 0.032. The amount of hemidesmosome attachment per basal cell did not increase significantly during growth of the airway. These data demonstrate that as tracheas grow in circumference, the columnar cells increase in height, basal cells increase in number, and anchoring junctional adhesion mechanisms differentiate in the basal cells. These changes are closely related to the height of the epithelium and result in maintaining a constant amount of attachment between the columnar epithelium and the basal lamina as the epithelium increases in height.
Serum Fetuin-A Levels in Patients with Bilateral Basal Ganglia Calcification.
Demiryurek, Bekir Enes; Gundogdu, Asli Aksoy
2018-02-14
The idiopathic basal ganglia calcification (Fahr syndrome) may occur due to senility. Fetuin-A is a negative acute phase reactant which inhibits calcium-phosphorus precipitation and vascular calcification. In this study, we aimed to evaluate whether serum fetuin-A levels correlate with bilateral basal ganglia calcification. Forty-five patients who had bilateral basal ganglia calcification on brain CT were selected according to the inclusion and exclusion criteria, and 45 age and gender-matched subjects without basal ganglia calcification were included for the control group. Serum fetuin-A levels were measured from venous blood samples. All participants were divided into two groups; with and without basal ganglia calcification. These groups were divided into subgroups regarding age (18-32 and 33-45 years of age) and gender (male, female). We detected lower levels of serum fetuin-A in patients with basal ganglia calcification compared with the subjects without basal ganglia calcification. In all subgroups (female, male, 18-32 years and 33-45 years), mean fetuin-A levels were significantly lower in patients with basal ganglia calcification (p = 0.017, p = 0.014, p = 0.024, p = 0.026, p = 0.01 respectively). And statistically significantly lower levels of fetuin-A was found to be correlated with the increasing densities of calcification in the calcified basal ganglia group (p-value: <0.001). Considering the role of fetuin-A in tissue calcification and inflammation, higher serum fetuin-A levels should be measured in patients with basal ganglia calcification. We believe that the measurement of serum fetuin-A may play a role in the prediction of basal ganglia calcification as a biomarker. Copyright © 2017 Elsevier B.V. All rights reserved.
NASA Astrophysics Data System (ADS)
De Ridder, Maaike; De Haulleville, Thalès; Kearsley, Elizabeth; Van den Bulcke, Jan; Van Acker, Joris; Beeckman, Hans
2014-05-01
It is commonly acknowledged that allometric equations for aboveground biomass and carbon stock estimates are improved significantly if density is included as a variable. However, not much attention is given to this variable in terms of exact, measured values and density profiles from pith to bark. Most published case-studies obtain density values from literature sources or databases, this way using large ranges of density values and possible causing significant errors in carbon stock estimates. The use of one single fixed value for density is also not recommended if carbon stock increments are estimated. Therefore, our objective is to measure and analyze a large number of tree species occurring in two Biosphere Reserves (Luki and Yangambi). Nevertheless, the diversity of tree species in these tropical forests is too high to perform this kind of detailed analysis on all tree species (> 200/ha). Therefore, we focus on the most frequently encountered tree species with high abundance (trees/ha) and dominance (basal area/ha) for this study. Increment cores were scanned with a helical X-ray protocol to obtain density profiles from pith to bark. This way, we aim at dividing the tree species with a distinct type of density profile into separate groups. If, e.g., slopes in density values from pith to bark remain stable over larger samples of one tree species, this slope could also be used to correct for errors in carbon (increment) estimates, caused by density values from simplified density measurements or density values from literature. In summary, this is most likely the first study in the Congo Basin that focuses on density patterns in order to check their influence on carbon stocks and differences in carbon stocking based on species composition (density profiles ~ temperament of tree species).
Lloyd, O.B.; Davis, R.W.
1989-01-01
Preliminary interpretation of available hydrogeologic data suggests that some areas underlying eastern Indiana, north-central Kentucky, and western Ohio might be worthy of further study regarding the disposal of high-level radioactive waste in Precambrian crystalline rocks buried beneath Paleozoic sedimentary rocks in the area. The data indicate that (1) largest areas of deepest potential burial and thickest sedimentary rock cover occur in eastern Indiana; (2) highest concentrations of dissolved solids in the basal sandstone aquifer, suggesting the most restricted circulation, are found in the southern part of the area near the Kentucky-Ohio State line and in southeastern Indiana; (3) largest areas of lowest porosity in the basal sandstone aquifer, low porosity taken as an indicator of the lowest groundwater flow velocity and contaminant migration, are found in northeastern Indiana and northwestern Ohio, central and southeastern Indiana, and central Kentucky; (4) the thickest confining units that directly overlie the basal sandstone aquifer are found in central Kentucky and eastern Indiana where their thickness exceeds 500 ft; (5) steeply dipping faults that form potential hydraulic connections between crystalline rock, the basal sandstone aquifer, and the freshwater circulation system occur on the boundaries of the study area mainly in central Kentucky and central Indiana. Collectively, these data indicate that the hydrogeology of the sedimentary rocks in the western part of the study area is more favorably suited than that in the remainder of the area for the application of the buried crystalline-rock concept. (USGS)
Depboylu, Candan; Weihe, Eberhard; Eiden, Lee E.
2011-01-01
The simian immunodeficiency virus (SIV) macaque model resembles human HIV-AIDS and associated brain dysfunction. Altered expression of synaptic markers and transmitters in neuro-AIDS has been reported, but limited data exist for the cholinergic system and lipid mediators such as prostaglandins. Here, we analyzed cholinergic basal forebrain neurons with their telencephalic projections and the rate-limiting enzymes for prostaglandin synthesis, cyclooxygenases 1 and 2 (COX1 and 2) in brains of SIV-infected macaques with and without encephalitis and antiretroviral therapy, and uninfected controls. COX1 but not COX2 was co-expressed with markers of cholinergic phenotype, i.e. choline acetyltransferase and vesicular acetylcholine transporter (VAChT), in basal forebrain neurons of monkey, as well as human samples. COX1 was decreased in basal forebrain neurons in macaques with AIDS vs. uninfected and asymptomatic SIV-infected macaques. VAChT-positive fiber density was reduced in frontal, parietal and hippocampal-entorhinal cortex. Although brain SIV burden and associated COX1- and COX2-positive mononuclear and endothelial inflammatory reactions were mostly reversed in AIDS-diseased macaques that received 6-chloro-2′,3′-dideoxyguanosine treatment, decreased VAChT-positive terminal density and reduced cholinergic COX1 expression were not. Thus, COX1 expression is a feature of primate cholinergic basal forebrain neurons; it may be functionally important and a critical biomarker of cholinergic dysregulation accompanying lentiviral encephalopathy. These results imply that insufficiently prompt initiation of antiretroviral therapy in lentiviral infection may lead to neurostructurally unremarkable but neurochemically prominent, irreversible brain damage. PMID:22157616
NASA Astrophysics Data System (ADS)
Benyon, Richard G.; Lane, Patrick N. J.; Jaskierniak, Dominik; Kuczera, George; Haydon, Shane R.
2015-07-01
Mean sapwood thickness, measured in fifteen 73 year old Eucalyptus regnans and E. delegatensis stands, correlated strongly with forest overstorey stocking density (R2 0.72). This curvilinear relationship was used with routine forest stocking density and basal area measurements to estimate sapwood area of the forest overstorey at various times in 15 research catchments in undisturbed and disturbed forests located in the Great Dividing Range, Victoria, Australia. Up to 45 years of annual precipitation and streamflow data available from the 15 catchments were used to examine relationships between mean annual loss (evapotranspiration estimated as mean annual precipitation minus mean annual streamflow), and sapwood area. Catchment mean sapwood area correlated strongly (R2 0.88) with catchment mean annual loss. Variation in sapwood area accounted for 68% more variation in mean annual streamflow than precipitation alone (R2 0.90 compared with R2 0.22). Changes in sapwood area accounted for 96% of the changes in mean annual loss observed after forest thinning or clear-cutting and regeneration. We conclude that forest inventory data can be used reliably to predict spatial and temporal variation in catchment annual losses and streamflow in response to natural and imposed disturbances in even-aged forests. Consequently, recent advances in mapping of sapwood area using airborne light detection and ranging will enable high resolution spatial and temporal mapping of mean annual loss and mean annual streamflow over large areas of forested catchment. This will be particularly beneficial in management of water resources from forested catchments subject to disturbance but lacking reliable long-term (years to decades) streamflow records.
Large slope instabilities in Northern Chile and Southern Peru
NASA Astrophysics Data System (ADS)
Crosta, Giovanni B.; Hermanns, Reginald L.; Valbuzzi, Elena; Frattini, Paolo; Valagussa, Andrea
2014-05-01
Deep canyon incision into Tertiary paleosurfaces and large slope instabilities along the canyon flanks characterize the landscape of western slope of the Andes of northern Chile and South Peru. This area belongs to the Coastal Escarpment and Precordillera and is formed by coarse-grained clastic and volcanoclastic formations. The area is characterized by intense seismicity and long-term hyperaridity (Atacama Desert). Landslides along the canyon flanks affect volumes generally up to 1 km3 and locally evolved in large rock avalanches. We prepared a landslide inventory covering an area of about 30,000 km2, extending from Iquique (Chile) to the South and Tacna (Peru) to the North. A total of 606 landslides have been mapped in the area by use of satellite images and direct field surveys, prevalently including large phenomena. The landslides range from 1 10-3 km2 to 464 km2 (Lluta landslide). The total landslide area, inclusive of the landslide scarp and of the deposit, amounts to about 2,130 km2 (about 7% of the area). The mega landslides can be classified as large block slides that can evolve in large rock avalanches (e.g. Minimini landslide). Their initiation seems to be strongly associated to the presence of secondary faults and large fractures transversal to the slope. These landslides show evidence suggesting a re-incision by the main canyon network. This seems particularly true for the Lluta collapse where the main 'landslide' mass is masked or deleted by the successive erosion. Other landslides have been mapped along the Coastal Escarpment and some of the major tectonic escarpments with an E-W trend. We examined area-frequency distributions of landslides by developing logarithmically binned, non-cumulative size frequency distributions that report frequency density as a function of landslide planar area A. The size frequency distribution presents a strong undersampling for smaller landslides, due to the extremely old age of the inventory. For landslides larger than 2 000 m2, the distribution exhibits a power-law behaviour with scaling exponent, β, equal to -2.24. For comparison, we analysed the power-law behaviour of other earthquake-induced landslide inventories, obtaining similar results, although the geological and seismic conditions may have been very different (Buller, New Zealand, β = -2.42; Iningahua, New Zealand, β = -2.53; Northridge, USA, β = -2.39; Chi-Chi, Taiwan, β = -2.30; Wenchuan Earthquake, China, β = -2.19). Volume estimates and slope stability modelling have been completed to characterize the phenomena and the possible triggering mechanisms. For volume estimate, we reconstructed the pre-failure surface for tens of landslides, in order to characterize the area-volume relationship. By using this relationship, we assigned a volume to all landslides of the inventory. The study area is subject to a high seismicity associated to earthquakes of different type: interplate (superficial and intermediate depth), subduction zone earthquakes, and earthquake along the Coastal Escarpment. By analysing the frequency size relationships for earthquake-induced landslides from literature, it is possible to observe that the higher the earthquake Magnitude, the higher the frequency density curve. To quantify this observation, we used the power-law relationships derived for each inventory to calculate the frequency density associated to selected areas, and we plotted these frequencies as a function of the magnitude of the respective earthquakes. By fitting these values, we derived the expected Magnitude required to generate the landslide distribution of the study area. In conclusion, we argue that the evolution of these landslides is controlled by: deep valley incision, canyon walls undercutting and lateral migration of the river controlled by valley flank instabilities, the Presence of weak lithologies and weak basal layers, the river incision debuttressing the slope toe and especially brings to daylighting the weak basal layers observed at some landslide sites, the possible deep groundwater flow above the deep impermeable formations and clay layers, the movement along sub-horizontal basal shear zones which can be locally extruded at the slope toe, the river damming because of the strong lateral components of displacement and successive reincision by the river with dam failure, the possible sequence of reactivations by reincision of the deposit, and the occurrence of high magnitude (8-9) earthquakes.
Adrenal steroids in post-menopausal women: relation to obesity and to bone mineral content.
Brody, S; Carlström, K; Lagrelius, A; Lunell, N O; Möllerström, G
1987-04-01
Basal levels and ACTH-induced increments of serum 17 alpha-hydroxyprogesterone (170HP), cortisol, 4-androstene-3,17-dione (A-4), dehydroepiandrosterone (DHA), and dehydroepiandrosterone sulphate (DHAS) were related to the degree of obesity and to trabecular bone mineral density in 29 postmenopausal women. The ACTH-induced increment of 170HP (delta 170HP) was negatively correlated to basal DHA and delta DHA. Positive correlations were found between obesity, expressed as Broca's index, and delta DHA and the delta DHA/delta 170HP ratio. Bone mineral density was positively correlated to basal DHAS, delta DHA, delta DHAS and the delta DHA/delta 170HP ratio, and negatively correlated to delta 170HP. DHA and 170HP represent a crossroad in adrenocortical steroid biosynthesis, leading to delta 5-androgens and glucocorticoids as main products. Besides age, obesity may also influence the intra-adrenal distribution between these two main steroidogenic pathways. The results suggest that differences at a very early stage of the adrenal steroidogenesis may influence calcium homeostasis in the post-menopausal woman.
Parresol, B. R.; Scott, D. A.; Zarnoch, S. J.; ...
2017-12-15
Spatially explicit mapping of forest productivity is important to assess many forest management alternatives. We assessed the relationship between mapped variables and site index of forests ranging from southern pine plantations to natural hardwoods on a 74,000-ha landscape in South Carolina, USA. Mapped features used in the analysis were soil association, land use condition in 1951, depth to groundwater, slope and aspect. Basal area, species composition, age and height were the tree variables measured. Linear modelling identified that plot basal area, depth to groundwater, soils association and the interactions between depth to groundwater and forest group, and between land usemore » in 1951 and forest group were related to site index (SI) (R 2 =0.37), but this model had regression attenuation. We then used structural equation modeling to incorporate error-in-measurement corrections for basal area and groundwater to remove bias in the model. We validated this model using 89 independent observations and found the 95% confidence intervals for the slope and intercept of an observed vs. predicted site index error-corrected regression included zero and one, respectively, indicating a good fit. With error in measurement incorporated, only basal area, soil association, and the interaction between forest groups and land use were important predictors (R2 =0.57). Thus, we were able to develop an unbiased model of SI that could be applied to create a spatially explicit map based primarily on soils as modified by past (land use and forest type) and recent forest management (basal area).« less
DOE Office of Scientific and Technical Information (OSTI.GOV)
Parresol, B. R.; Scott, D. A.; Zarnoch, S. J.
Spatially explicit mapping of forest productivity is important to assess many forest management alternatives. We assessed the relationship between mapped variables and site index of forests ranging from southern pine plantations to natural hardwoods on a 74,000-ha landscape in South Carolina, USA. Mapped features used in the analysis were soil association, land use condition in 1951, depth to groundwater, slope and aspect. Basal area, species composition, age and height were the tree variables measured. Linear modelling identified that plot basal area, depth to groundwater, soils association and the interactions between depth to groundwater and forest group, and between land usemore » in 1951 and forest group were related to site index (SI) (R 2 =0.37), but this model had regression attenuation. We then used structural equation modeling to incorporate error-in-measurement corrections for basal area and groundwater to remove bias in the model. We validated this model using 89 independent observations and found the 95% confidence intervals for the slope and intercept of an observed vs. predicted site index error-corrected regression included zero and one, respectively, indicating a good fit. With error in measurement incorporated, only basal area, soil association, and the interaction between forest groups and land use were important predictors (R2 =0.57). Thus, we were able to develop an unbiased model of SI that could be applied to create a spatially explicit map based primarily on soils as modified by past (land use and forest type) and recent forest management (basal area).« less
Cohen, Philip R
2014-08-17
Basal cell carcinoma of the axilla, an area that is not usually exposed to the sun, is rare. Individuals with basal cell nevus syndrome, a disorder associated with a mutation in the patch 1 (PTCH1) gene, develop numerous basal cell carcinomas. To describe a woman with basal cell nevus syndrome who developed a pigmented basal cell carcinoma in each of her axilla and to review the features of axillary basal cell carcinoma patients with Goltz-Gorlin syndrome. Pubmed was used to search the following terms: axillary basal cell carcinoma and basal cell nevus syndrome. The papers and their citations were evaluated. Basal cell nevus syndrome patients with basal cell carcinoma of the axilla were observed in two women; this represents 2.5% (2 of 79) of the patients with axillary basal cell carcinoma. Both women had pigmented tumors that were histologically nonaggressive. The cancers did not recur after curettage or excision. Basal cell carcinoma of the axilla has only been described in 79 individuals; two of the patients were women with pigmented tumors who had basal cell nevus syndrome. Similar to other patients with axillary basal cell carcinoma, the tumors were histologically nonaggressive and did not recur following treatment. Whether PTCH1 gene mutation predisposes basal cell nevus patients to develop axillary basal cell carcinomas remains to be determined.
McCullough, Deborah G; Poland, Therese M; Anulewicz, Andrea C; Lewis, Phillip; Cappaert, David
2011-10-01
Effective methods are needed to protect ash trees (Fraxinus spp.) from emerald ash borer, Agrilus planipennis Fairmaire (Coleoptera: Buprestidae), an invasive buprestid that has killed millions of North American ash (Fraxinus spp.) trees. We randomly assigned 175 ash trees (11.5-48.1 cm in diameter) in 25 blocks located in three study sites in Michigan to one of seven insecticide treatments in May 2007. Treatments included 1) trunk-injected emamectin benzoate; 2) trunk-injected imidacloprid; 3) basal trunk spray of dinotefuran with or 4) without Pentra-Bark, an agricultural surfactant; 5) basal trunk spray of imidacloprid with or 6) without Pentra-Bark; or (7) control. Foliar insecticide residues (enzyme-linked immunosorbent assay) and toxicity of leaves to adult A. planipennis (4-d bioassays) were quantified at 3-4-wk intervals posttreatment. Seven blocks of trees were felled and sampled in fall 2007 to quantify A. planipennis larval density. Half of the remaining blocks were retreated in spring 2008. Bioassays and residue analyses were repeated in summer 2008, and then all trees were sampled to assess larval density in winter. Foliage from emamectin benzoate-treated trees was highly toxic to adult A. planipennis, and larval density was < 1% of that in comparable control trees, even two seasons posttreatment. Larval densities in trees treated with trunk-injected imidacloprid in 2007 + 2008 were similar to control trees. Dinotefuran and imidacloprid were effectively translocated within trees treated with the noninvasive basal trunk sprays; the surfactant did not appreciably enhance A. planipennis control. In 2008, larval densities were 57-68% lower in trees treated with dinotefuran or imidacloprid trunk sprays in 2007 + 2008 than on controls, but densities in trees treated only in 2007 were similar to controls. Highly effective control provided by emamectin benzoate for > or = 2 yr may reduce costs or logistical issues associated with treatment.
Compatible Basal Area and Number of Trees Estimators from Remeasured Horizontal Point Samples
Francis A. Roesch; Edwin J. Green; Charles T. Scott
1989-01-01
Compatible groups of estimators for total value at time 1 (V1), survivor growth (S), and ingrowth (I) for use with permanent horizontal point samples are evaluated for the special cases of estimating the change in both the number of trees and basal area. Caveats which should be observed before any one compatible grouping of estimators is chosen...
Basal area growth for 15 tropical trees species in Puerto Rico. Forest
B. R. Parresol
1995-01-01
The tabonuco forest of Puerto Rico support a diverse population of tree species valued for timber, fuel, food, wildlife food and cover, and erosion control among other use. tree basal area growth data spanning 39 years are avaible on 15 species from eigth permanent plots in Luquillo Experimental Forest. The complexity of the rain forest challeges current forest...
Darren A. Miller; Bruce D. Leopold; L. Mike Conner; Michael G. Shelton
1999-01-01
Uneven-aged management (UEAM) is becoming increasingly popular in the southeastern United States. However, effects of UEAM on wildlife habitat have not been adequately documented. We examined response of habitat within stands of varying levels of pine and hardwood basal area under an uneven-aged managegement regime in southern Mississippi. Summer and winter trends in...
Coast Live Oak Thinning Study in the Central Coast of California
Norman H. Pillsbury; Michael J. DeLasaux; Timothy R. Plumb
1987-01-01
Abstract: Along-term thinning study was established in ten stands of coast live oak (Quercus agrifolia N in the Central Coast of California. Information about diameter, basal area, and volume growth and yield is being obtained from unthinned control plots and from plots thinned to 50 and 100 square feet of basal area per acre. Descriptive information was also collected...
Yield Tables and Stand Structure for Unthinned Longleaf Pine Plantations in Louisiana and Texas
Richard E. Longrey; Robert L. Bailey
1977-01-01
A system of equations is developed to forecast number of trees per acre, basal area, and cubic foot yields in eight volume categories by l-inch diameter classes for several combinations of site index, age from planting, and either number of trees planted, number of trees surviving, or basal area at a given age.
Growth and Yield of Appalachian Mixed Hardwoods After Thinning
Wade C. Harrison; Harold E. Burkhart; Thomas E. Burk; Donald E. Beckand
1986-01-01
G-RAT (Growth of Hardwoods After Thinning) is a system of computer programs used to predict growth and yield of Appalachian mixed hardwoods after thinning. Given a tree list or stand table, along with inputs of stand age, site index, and stand basal area before thinning, G-RAT software uses species-specific individual tree equations to predict tree basal area...
Basal area growth for 15 tropical tree species in Puerto Rico
Bernard R. Parresol
1995-01-01
The tabonuco forests of Puerto Rico support a diverse population of tree species valued for timber, fuel, food, wildlife food and cover, and erosion control among other uses. Tree basal area growth data spanning 39 years are available on 15 species from eight permanent plots in the Luquillo Experimental Forest. The complexity of the rain forest challenges current...
Estimation of crown cover in interior ponderosa pine stands: Effects of thinning and prescribed fire
Nicholas Vaughn; Martin W. Ritchie
2005-01-01
We evaluated the relationship between crown cover measured with a vertical sight tube and stand basal area per acre in treated (thinned, burned, and thinned and burned) and untreated interior ponderosa pine (Pinus ponderosa P. & C. Lawson) stands in northeastern California. Crown cover was significantly related to basal area at the plot level and...
Stand basal-area and tree-diameter growth in red spruce-fir forests in Maine, 1960-80
S.J. Zarnoch; D.A. Gansner; D.S. Powell; T.A. Birch; T.A. Birch
1990-01-01
Stand basal-area change and individual surviving red spruce d.b.h. growth from 1960 to 1980 were analyzed for red spruce-fir stands in Maine. Regression modeling was used to relate these measures of growth to stand and tree conditions and to compare growth throughout the period. Results indicate a decline in growth.
Tree basal area as an index of thermal cover for elk.
J. Edward Dealy
1985-01-01
The relationship of basal area to crown closure was studied in five major forest types of the Blue Mountains of Oregon and Washington. The regressions developed give wildlife and forest managers a tool for estimating the amount of crown closure if data are not available from stand examinations. Information is used in determining quantity and quality of elk thermal...
Forage production after thinning a natural loblolly pine-hardwoocl stand to clifferent basal areas
David G. Peitz; Michael G. Shelton; Philip A. Tappe
2001-01-01
Mixed pine (Pinus spp.)-hardwood forests are common in the southern United States (U.S.), but little quantitative information exists on the response of understory forage to reductions in basal area from thinning. We determined understory forage characteristics before thinning and 2 and 4 years after thinning a 35-year-old natural loblolly pine (
Heavy thinning of ponderosa pine stands: An Arizona case study
Peter F. Ffolliott; Jr. Baker; Gerald J. Gottfried
2000-01-01
Growth and structural changes in a mosaic of even-aged ponderosa pine (Pinus ponderosa) stands were studied for 25 years to determine the long-term impacts of a heavy thinning treatment to a basal-area level of 25 ft2/acre. Basal area and volume growth of these stands has increased since thinning and likely will continue to...
Historical harvests reduce neighboring old-growth basal area across a forest landscape
David M. Bell; Thomas A. Spies; Robert Pabst
2017-01-01
While advances in remote sensing have made stand, landscape, and regional assessments of the direct impacts of disturbance on forests quite common, the edge influence of timber harvesting on the structure of neighboring unharvested forests has not been examined extensively. In this study, we examine the impact of historical timber harvests on basal area patterns of...
Bravo, Felipe; Hann, D.W.; Maguire, Douglas A.
2001-01-01
Mixed conifer and hardwood stands in southwestern Oregon were studied to explore the hypothesis that competition effects on individual-tree growth and survival will differ according to the species comprising the competition measure. Likewise, it was hypothesized that competition measures should extrapolate best if crown-based surrogates are given preference over diameter-based (basal area based) surrogates. Diameter growth and probability of survival were modeled for individual Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) trees growing in pure stands. Alternative models expressing one-sided and two-sided competition as a function of either basal area or crown structure were then applied to other plots in which Douglas-fir was mixed with other conifers and (or) hardwood species. Crown-based variables outperformed basal area based variables as surrogates for one-sided competition in both diameter growth and survival probability, regardless of species composition. In contrast, two-sided competition was best represented by total basal area of competing trees. Surrogates reflecting differences in crown morphology among species relate more closely to the mechanics of competition for light and, hence, facilitate extrapolation to species combinations for which no observations are available.
The role of basal cells in adhesion of columnar epithelium to airway basement membrane.
Evans, M J; Plopper, C G
1988-08-01
In this report, we present a new concept of the role of the basal cell in airway epithelium. Previously, the basal cell was thought to be the progenitor cell for the columnar epithelium. However, several studies have shown that this concept may not be correct. The morphologic aspects of the basal cell suggest that it could play a role in adhesion of the columnar epithelium to the basement membrane. Basal cells form attachments with columnar cells (desmosomes) and with the basement membrane (hemidesmosomes). Columnar cells do not form hemidesmosome attachments with the basement membrane. Basal cells could strengthen the adhesion of columnar cells to the basement membrane by forming hemidesmosome attachments to the basement membrane and desmosome attachments with adjacent columnar cells. Incidental evidence from 2 existing publications concerning airway microanatomy support this concept. As columnar cells grow taller, the proportion of the cell surface in contact with the basement membrane becomes progressively smaller, and thus the cell surface area related to adhesion also becomes smaller. It was found that the number of basal cells per millimeter of basement membrane was closely related to the height of the columnar cell epithelium (r = 0.98), but not to the number of columnar cells (r = 0.42). The consistency of the relationship between increased columnar cell height (and thus decreased surface area for adhesion) and the number of basal cells present (r = 0.98) supports the concept that the basal cell plays a role in adhesion of columnar cells to the basement membrane.(ABSTRACT TRUNCATED AT 250 WORDS)
A novel method of basal crevasse height estimation and subsequent rifting
NASA Astrophysics Data System (ADS)
Logan, L.; Catania, G. A.; Lavier, L. L.; Choi, E.
2012-12-01
Basal crevasses may play an important precursory role in the location and propagation of rifts and in ice shelf disintegration. Here we develop a novel method for estimating the locations and heights of basal crevasses formed at the grounding line of ice shelves and ice streams. We assume a thin-elastic beam formulation (TEB) with a tensional plastic yielding criterion to capture the physics of a tidally flexed grounding line. Observations of basal crevasses in the Siple Coast area match well with predictions produced by this method. Areas with large misfit can be delineated by examining the strain rate field; indeed, in our estimations those crevasses which deviate most from the TEB prediction lie directly in a shear margin. We test the method against other areas in the Larsen Ice Shelf, and find again a good match. Thus we suggest the TEB as an alternative to other crevasse estimation methods, as it produces a good fit in predominantly tensile regions, requires no tuning or prior information, and is computationally free to implement into large scale ice models which aim at physically simulating calving and fracture processes. We pursue modeling basal crevasses as they evolve with a thermomechanical finite-difference 3-dimensional model called SNAC. Viscoelastoplastic ice follows Mohr-Coulomb tension failure with Glen's flow law. We examine the conditions necessary for a basal crevasse formed on the downstream side of an ice rise to propagate the full thickness of the ice, developing into a rift.
Andó, Rómeó D; Adori, Csaba; Kirilly, Eszter; Molnár, Eszter; Kovács, Gábor G; Ferrington, Linda; Kelly, Paul A T; Bagdy, György
2010-03-05
To assess the functional state of the serotonergic system, the acute behavioural and brain metabolic effect of SSRI antidepressants were studied during the recovery period after MDMA-induced neuronal damage. The effects of the SSRI fluoxetine and the serotonin receptor agonist meta-chloro-phenylpiperazine (m-CPP) were investigated in the social interaction test in Dark Agouti rats, 6 months after treatment with a single dose of MDMA (15 or 30 mg kg(-1), i.p.). At earlier time points these doses of MDMA have been shown to cause 30-60% loss in axonal densities in several brain regions. Densities of the serotonergic axons were assessed using serotonin-transporter and tryptophan-hydroxylase immunohistochemistry. In a parallel group of animals, brain function was examined following an acute challenge with either fluoxetine or citalopram, using 2-deoxyglucose autoradiographic imaging. Six months after MDMA treatment the densities of serotonergic axons were decreased in only a few brain areas including hippocampus and thalamus. Basal anxiety was unaltered in MDMA-treated animals. However, the acute anxiogenic effects of fluoxetine, but not m-CPP, were attenuated in animals pretreated with MDMA. The metabolic response to both citalopram and fluoxetine was normal in most of the brain areas examined with the exception of ventromedial thalamus and hippocampal sub-fields where the response was attenuated. These data provide evidence that 6 months after MDMA-induced damage serotonergic axons show recovery in most brain areas, but serotonergic functions to challenges with SSRIs including anxiety and aggression remain altered. Copyright 2009 Elsevier B.V. All rights reserved.
Morello, Noemi; Plicato, Ornella; Piludu, Maria Antonietta; Poddighe, Laura; Serra, Maria Pina; Quartu, Marina; Corda, Maria Giuseppa; Giorgi, Osvaldo; Giustetto, Maurizio
2017-01-01
Stressful events evoke molecular adaptations of neural circuits through chromatin remodeling and regulation of gene expression. However, the identity of the molecular pathways activated by stress in experimental models of depression is not fully understood. We investigated the effect of acute forced swimming (FS) on the phosphorylation of the extracellular signal-regulated kinase (ERK)1/2 (pERK) and histone H3 (pH3) in limbic brain areas of genetic models of vulnerability (RLA, Roman low-avoidance rats) and resistance (RHA, Roman high-avoidance rats) to stress-induced depression-like behavior. We demonstrate that FS markedly increased the density of pERK-positive neurons in the infralimbic (ILCx) and the prelimbic area (PrLCx) of the prefrontal cortex (PFCx), the nucleus accumbens, and the dorsal blade of the hippocampal dentate gyrus to the same extent in RLA and RHA rats. In addition, FS induced a significant increase in the intensity of pERK immunoreactivity (IR) in neurons of the PFCx in both rat lines. However, RHA rats showed stronger pERK-IR than RLA rats in the ILCx both under basal and stressed conditions. Moreover, the density of pH3-positive neurons was equally increased by FS in the PFCx of both rat lines. Interestingly, pH3-IR was higher in RHA than RLA rats in PrLCx and ILCx, either under basal conditions or upon FS. Finally, colocalization analysis showed that in the PFCx of both rat lines, almost all pERK-positive cells express pH3, whereas only 50% of the pH3-positive neurons is also pERK-positive. Moreover, FS increased the percentage of neurons that express exclusively pH3, but reduced the percentage of cells expressing exclusively pERK. These results suggest that (i) the distinctive patterns of FS-induced ERK and H3 phosphorylation in the PFCx of RHA and RLA rats may represent molecular signatures of the behavioural traits that distinguish the two lines and (ii) FS-induced H3 phosphorylation is, at least in part, ERK-independent.
Piludu, Maria Antonietta; Poddighe, Laura; Serra, Maria Pina; Quartu, Marina; Corda, Maria Giuseppa; Giorgi, Osvaldo
2017-01-01
Stressful events evoke molecular adaptations of neural circuits through chromatin remodeling and regulation of gene expression. However, the identity of the molecular pathways activated by stress in experimental models of depression is not fully understood. We investigated the effect of acute forced swimming (FS) on the phosphorylation of the extracellular signal-regulated kinase (ERK)1/2 (pERK) and histone H3 (pH3) in limbic brain areas of genetic models of vulnerability (RLA, Roman low-avoidance rats) and resistance (RHA, Roman high-avoidance rats) to stress-induced depression-like behavior. We demonstrate that FS markedly increased the density of pERK-positive neurons in the infralimbic (ILCx) and the prelimbic area (PrLCx) of the prefrontal cortex (PFCx), the nucleus accumbens, and the dorsal blade of the hippocampal dentate gyrus to the same extent in RLA and RHA rats. In addition, FS induced a significant increase in the intensity of pERK immunoreactivity (IR) in neurons of the PFCx in both rat lines. However, RHA rats showed stronger pERK-IR than RLA rats in the ILCx both under basal and stressed conditions. Moreover, the density of pH3-positive neurons was equally increased by FS in the PFCx of both rat lines. Interestingly, pH3-IR was higher in RHA than RLA rats in PrLCx and ILCx, either under basal conditions or upon FS. Finally, colocalization analysis showed that in the PFCx of both rat lines, almost all pERK-positive cells express pH3, whereas only 50% of the pH3-positive neurons is also pERK-positive. Moreover, FS increased the percentage of neurons that express exclusively pH3, but reduced the percentage of cells expressing exclusively pERK. These results suggest that (i) the distinctive patterns of FS-induced ERK and H3 phosphorylation in the PFCx of RHA and RLA rats may represent molecular signatures of the behavioural traits that distinguish the two lines and (ii) FS-induced H3 phosphorylation is, at least in part, ERK-independent. PMID:28107383
Heuner, Maike; Silinski, Alexandra; Schoelynck, Jonas; Bouma, Tjeerd J; Puijalon, Sara; Troch, Peter; Fuchs, Elmar; Schröder, Boris; Schröder, Uwe; Meire, Patrick; Temmerman, Stijn
2015-01-01
In hydrodynamically stressful environments, some species--known as ecosystem engineers--are able to modify the environment for their own benefit. Little is known however, about the interaction between functional plant traits and ecosystem engineering. We studied the responses of Scirpus tabernaemontani and Scirpus maritimus to wave impact in full-scale flume experiments. Stem density and biomass were used to predict the ecosystem engineering effect of wave attenuation. Also the drag force on plants, their bending angle after wave impact and the stem biomechanical properties were quantified as both responses of stress experienced and effects on ecosystem engineering. We analyzed lignin, cellulose, and silica contents as traits likely effecting stress resistance (avoidance, tolerance). Stem density and biomass were strong predictors for wave attenuation, S. maritimus showing a higher effect than S. tabernaemontani. The drag force and drag force per wet frontal area both differed significantly between the species at shallow water depths (20 cm). At greater depths (35 cm), drag forces and bending angles were significantly higher for S. maritimus than for S. tabernaemontani. However, they do not differ in drag force per wet frontal area due to the larger plant surface of S. maritimus. Stem resistance to breaking and stem flexibility were significantly higher in S. tabernaemontani, having a higher cellulose concentration and a larger cross-section in its basal stem parts. S. maritimus had clearly more lignin and silica contents in the basal stem parts than S. tabernaemontani. We concluded that the effect of biomass seems more relevant for the engineering effect of emergent macrophytes with leaves than species morphology: S. tabernaemontani has avoiding traits with minor effects on wave attenuation; S. maritimus has tolerating traits with larger effects. This implies that ecosystem engineering effects are directly linked with traits affecting species stress resistance and responding to stress experienced.
Schoelynck, Jonas; Bouma, Tjeerd J.; Puijalon, Sara; Troch, Peter; Fuchs, Elmar; Schröder, Boris; Schröder, Uwe; Meire, Patrick; Temmerman, Stijn
2015-01-01
In hydrodynamically stressful environments, some species—known as ecosystem engineers—are able to modify the environment for their own benefit. Little is known however, about the interaction between functional plant traits and ecosystem engineering. We studied the responses of Scirpus tabernaemontani and Scirpus maritimus to wave impact in full-scale flume experiments. Stem density and biomass were used to predict the ecosystem engineering effect of wave attenuation. Also the drag force on plants, their bending angle after wave impact and the stem biomechanical properties were quantified as both responses of stress experienced and effects on ecosystem engineering. We analyzed lignin, cellulose, and silica contents as traits likely effecting stress resistance (avoidance, tolerance). Stem density and biomass were strong predictors for wave attenuation, S. maritimus showing a higher effect than S. tabernaemontani. The drag force and drag force per wet frontal area both differed significantly between the species at shallow water depths (20 cm). At greater depths (35 cm), drag forces and bending angles were significantly higher for S. maritimus than for S. tabernaemontani. However, they do not differ in drag force per wet frontal area due to the larger plant surface of S. maritimus. Stem resistance to breaking and stem flexibility were significantly higher in S. tabernaemontani, having a higher cellulose concentration and a larger cross-section in its basal stem parts. S. maritimus had clearly more lignin and silica contents in the basal stem parts than S. tabernaemontani. We concluded that the effect of biomass seems more relevant for the engineering effect of emergent macrophytes with leaves than species morphology: S. tabernaemontani has avoiding traits with minor effects on wave attenuation; S. maritimus has tolerating traits with larger effects. This implies that ecosystem engineering effects are directly linked with traits affecting species stress resistance and responding to stress experienced. PMID:26367004
Neoplasms treatment by diode laser with and without real time temperature control on operation zone
NASA Astrophysics Data System (ADS)
Belikov, Andrey V.; Gelfond, Mark L.; Shatilova, Ksenia V.; Sosenkova, Svetlana A.; Lazareva, Anastasia A.; Semyashkina, Yulia V.
2016-04-01
Results of nevus, papilloma, dermatofibroma, and basal cell skin cancer in vivo removal by a 980+/-10 nm diode laser with "blackened" tip operating in continuous (CW) mode and automatic power control (APC) mode are presented. The collateral damage width and width of graze wound area around the collateral damage area were demonstrated. The total damage area width was calculated as sum of collateral damage width and graze wound area width. The mean width of total damage area reached 1.538+/-0.254 mm for patient group with nevus removing by 980 nm diode laser operating in CW mode, papilloma - 0.586+/-0.453 mm, dermatofibroma - 1.568+/-0.437 mm, and basal cell skin cancer - 1.603+/-0.613 mm. The mean width of total damage area reached 1.201+/-0.292 mm for patient group with nevus removing by 980 nm diode laser operating in APC mode, papilloma - 0.413+/-0.418 mm, dermatofibroma - 1.240+/-0.546 mm, and basal cell skin cancer - 1.204+/-0.517 mm. It was found that using APC mode decreases the total damage area width at removing of these nosological neoplasms of human skin, and decreases the width of graze wound area at removing of nevus and basal cell skin cancer. At the first time, the dynamic of output laser power and thermal signal during laser removal of nevus in CW and APC mode is presented. It was determined that output laser power during nevus removal for APC mode was 1.6+/-0.05 W and for CW mode - 14.0+/-0.1 W. This difference can explain the decrease of the total damage area width and width of graze wound area for APC mode in comparison with CW mode.
Maximum size-density relationships for mixed-hardwood forest stands in New England
Dale S. Solomon; Lianjun Zhang
2000-01-01
Maximum size-density relationships were investigated for two mixed-hardwood ecological types (sugar maple-ash and beech-red maple) in New England. Plots meeting type criteria and undergoing self-thinning were selected for each habitat. Using reduced major axis regression, no differences were found between the two ecological types. Pure species plots (the species basal...
Benjamin C. Bright; Andrew T. Hudak; Robert E. Kennedy; Arjan J. H. Meddens
2014-01-01
Bark beetle-caused tree mortality affects important forest ecosystem processes. Remote sensing methodologies that quantify live and dead basal area (BA) in bark beetle-affected forests can provide valuable information to forest managers and researchers. We compared the utility of light detection and ranging (lidar) and the Landsat-based detection of trends in...
Comparing basal area growth models, consistency of parameters, and accuracy of prediction
J.J. Colbert; Michael Schuckers; Desta Fekedulegn
2002-01-01
We fit alternative sigmoid growth models to sample tree basal area historical data derived from increment cores and disks taken at breast height. We examine and compare the estimated parameters for these models across a range of sample sites. Models are rated on consistency of parameters and on their ability to fit growth data from four sites that are located across a...
Nathan J. Poage
2008-01-01
To better understand the long-term basal area and diameter growth response of young, well-stocked, even-aged, mixed-species stands of western hemlock (Tsuga heterophylla (Raf.) Sarg.) and Sitka spruce (Picea sitchensis (Bong.) Carr.) to a range of thinning intensities (heavy, moderate, light, and unthinned), 20 years of...
Ferhat Kara; Edward F. Loewenstein
2015-01-01
Even-aged silvicultural methods have been successfully used to manage longleaf pine (Pinus palustris Mill.) forests for wood production; however, successful use of uneven-aged methods to manage this ecosystem is less well documented. In this study, the effects of varying levels of residual basal area (RBA) (9.2, 13.8, and 18.4 m2...
Marcos Barrio Anta; Fernando Castedo Dorado; Ulises Dieguez-Aranda; Juan G. Alvarez Gonzalez; Bernard R. Parresol; Roque Rodriguez Soalleiro
2006-01-01
A basal area growth system for single-species, even-aged maritime pine (Pinus pinaster Ait.) stands in Galicia (northwestern Spain) was developed from data of 212 plots measured between one and four times. Six dynamic equations were considered for analysis, and both numerical and graphical methods were used to compare alternative models. The double...
An individual-tree basal area growth model for loblolly pine stands
Paul A. Murphy; Michael G. Shelton
1996-01-01
Tree basal area growth has been modeled as a combination of a potential growth function and a modifier function, in which the potential function is fitted separately from open-grown tree data or a subset of the data and the modifier function includes stand and site variables. We propose a modification of this by simultaneously fitting both a growth component and a...
Individual tree basal-area growth parameter estimates for four models
J.J. Colbert; Michael Schuckers; Desta Fekedulegn; James Rentch; Mairtin MacSiurtain; Kurt Gottschalk
2004-01-01
Four sigmoid growth models are fit to basal-area data derived from increment cores and disks taken at breast height from oak trees. Models are rated on their ability to fit growth data from five datasets that are obtained from 10 locations along a longitudinal gradient across the states of Delaware, Pennsylvania, West Virginia, and Ohio in the USA. We examine and...
Comparing Basal Area Growth Rates in Repeated Inventories: Simpson's Paradox in Forestry
Charles E. Thomas; Bernard R. Parresol
1989-01-01
Recent analyses of radial growth rates in southern commercial forests have shown that current rates are lower than past rates when compared diameter class by diameter class. These results have been interpreted as an indication that the growth rate of the forest is declining. In this paper, growth rates of forest populations in Alabama are studied. Basal area growth (a...
Revised white pine stocking guide for managed stands
William B. Leak; Neil I. Lamson
1999-01-01
Stocking guides are basic tools for forest managers. They provide estimates of the range in acceptable stocking for full occupancy of the site. The first stocking guide for white pine was developed by Philbrook et al (1973). It was of conventional format: showing trees and basal area per acre (in the main crown canopy) by mean stand dbh (the tree of average basal area...
Dale G. Brockway; Edward F. Loewenstein; Kenneth W. Outcalt
2014-01-01
Proportional basal area (Pro-B) was developed as an accurate, easy-to-use method for making uneven-aged silviculture a practical management option. Following less than 3 h of training, forest staff from a range of professional backgrounds used Pro-B in an operational-scale field study to apply single-tree selection and group selection systems in longleaf pine (Pinus...
Diameter Growth of Loblolly Pine Trees as Affected by Soil-Moisture Availibility
John R. Bassett
1964-01-01
In a 30-year-old even-aged stand of loblolly pine on a site 90 loessial soil in southeast Arkansas during foul growing seasons, most trees on plots thinned to 125 square feet of basal area per acre increased in basal area continuously when, under the crown canopy, available water in the surface foot remained above 65 percent. Measurable diameter growth ceased when...
Jose F. Negron; Willis C. Schaupp; Kenneth E. Gibson; John Anhold; Dawn Hansen; Ralph Thier; Phil Mocettini
1999-01-01
Data collected from Douglas-fir stands infected by the Douglas-fir beetle in Wyoming, Montana, Idaho, and Utah, were used to develop models to estimate amount of mortality in terms of basal area killed. Models were built using stepwise linear regression and regression tree approaches. Linear regression models using initial Douglas-fir basal area were built for all...
Patterned basal seismicity shows sub-ice stream bedforms
NASA Astrophysics Data System (ADS)
Barcheck, C. G.; Tulaczyk, S. M.; Schwartz, S. Y.
2017-12-01
Patterns in seismicity emanating from the bottom of fast-moving ice streams and glaciers may indicate localized patches of higher basal resistance— sometimes called 'sticky spots', or otherwise varying basal properties. These seismogenic basal areas resist an unknown portion of the total driving stress of the Whillans Ice Plain (WIP), in West Antarctica, but may play an important role in the WIP stick-slip cycle and ice stream slowdown. To better understand the mechanism and importance of basal seismicity beneath the WIP, we analyze seismic data collected by a small aperture (< 3km) network of 8 surface and 5 borehole seismometers installed in the main central sticky spot of the WIP. We use a network beamforming technique to detect and roughly locate thousands of small (magnitude < 0), local basal micro-earthquakes in Dec 2014, and we compare the resulting map of seismicity to ice bottom depth measured by airborne radar. The number of basal earthquakes per area within the network is spatially heterogeneous, but a pattern of two 400m wide streaks of high seismicity rates is evident, with >50-500 earthquakes detected per 50x50m grid cell in 2 weeks. These seismically active streaks are elongated approximately in the ice flow direction with a spacing of 750m. Independent airborne radar measurements of ice bottom depth from Jan 2013 show a low-amplitude ( 5m) undulation in the basal topography superposed on a regional gradient in ice bottom depth. The flow-perpendicular wavelength of these low-amplitude undulations is comparable to the spacing of the high seismicity bands, and the streaks of high seismicity intersect local lows in the undulating basal topography. We interpret these seismic and radar observations as showing seismically active sub-ice stream bedforms that are low amplitude and elongated in the direction of ice flow, comparable to the morphology of mega scale glacial lineations (MSGLs), with high basal seismicity rates observed in the MSGL troughs. These results have implications for understanding the formation mechanism of MSGLS and well as understanding the interplay between basal topographic roughness, spatially varying basal till and hydrologic properties, basal resistance to fast ice flow, and ice stream stick-slip.
Micro-Scale Genomic DNA Copy Number Aberrations as Another Means of Mutagenesis in Breast Cancer
Chao, Hann-Hsiang; He, Xiaping; Parker, Joel S.; Zhao, Wei; Perou, Charles M.
2012-01-01
Introduction In breast cancer, the basal-like subtype has high levels of genomic instability relative to other breast cancer subtypes with many basal-like-specific regions of aberration. There is evidence that this genomic instability extends to smaller scale genomic aberrations, as shown by a previously described micro-deletion event in the PTEN gene in the Basal-like SUM149 breast cancer cell line. Methods We sought to identify if small regions of genomic DNA copy number changes exist by using a high density, gene-centric Comparative Genomic Hybridizations (CGH) array on cell lines and primary tumors. A custom tiling array for CGH (244,000 probes, 200 bp tiling resolution) was created to identify small regions of genomic change, which was focused on previously identified basal-like-specific, and general cancer genes. Tumor genomic DNA from 94 patients and 2 breast cancer cell lines was labeled and hybridized to these arrays. Aberrations were called using SWITCHdna and the smallest 25% of SWITCHdna-defined genomic segments were called micro-aberrations (<64 contiguous probes, ∼ 15 kb). Results Our data showed that primary tumor breast cancer genomes frequently contained many small-scale copy number gains and losses, termed micro-aberrations, most of which are undetectable using typical-density genome-wide aCGH arrays. The basal-like subtype exhibited the highest incidence of these events. These micro-aberrations sometimes altered expression of the involved gene. We confirmed the presence of the PTEN micro-amplification in SUM149 and by mRNA-seq showed that this resulted in loss of expression of all exons downstream of this event. Micro-aberrations disproportionately affected the 5′ regions of the affected genes, including the promoter region, and high frequency of micro-aberrations was associated with poor survival. Conclusion Using a high-probe-density, gene-centric aCGH microarray, we present evidence of small-scale genomic aberrations that can contribute to gene inactivation. These events may contribute to tumor formation through mechanisms not detected using conventional DNA copy number analyses. PMID:23284754
Stohlgren, Thomas J.
1992-01-01
The Big Stump Grove of giant sequoia (Sequoiadendron giganteum (Lindl.) Buchholz) was heavily logged between 1883 and 1889 and the stand naturally regenerated from seed following logging. In 1968, as part of a 100% sequoia tree inventory, all living sequoias (n = 3587) and dead trees and stumps (n=588) were measured (diameter at breast height, dbh) and mapped. A comparison of pre- to post-logging (85 years later in 1968) stand characteristics showed the estimated basal area of 56.7 m2 ha−1 in the pre-cut 1883 Big Stump Grove was very similar to the population mean basal area of 30 other giant sequoia groves (with more than 30 trees) in Sequoia and Kings Canyon National Parks. Sequoia density in 1968 was 1.5 times higher than the population mean, and over 45% of the basal area had been recovered after only 85 years. Assuming most re-establishment occurred over roughly a 9 year period (1883–1892), the diameter growth rate of trees less than 1.95 m dbh, averaged 6.1–6.8 mm year−1 but greatly varied as the 24 trees in the 1.8 m size class had a mean diameter growth rate of 21–24 mm year−1. Data generated by dividing the grove into 0.25 ha contiguous plots indicated that only about 3.3 ha of the pre-cut 1883 grove did not have sequoia regeneration whereas 16.5 ha of the 1968 grove had sequoia regeneration but no sign of logs or stumps. The proportion of only-regeneration plots was significantly greater (Pt=0; 1968) stand, overrepresentation of 0.3–1.2 m dbh trees may produce a bimodal size distribution lasting perhaps 800 years or more into the future. Giant sequoia stand characteristics such as age and size structure are not highly resilient and may take several centuries to approach the ‘domain’ of age or size structure typical of old-growth sequoia forests. Grove boundaries may be less stable following a major disturbance.
Omega-3 supplementation is neuroprotective to corneal nerves in dry eye disease: a pilot study.
Chinnery, Holly R; Naranjo Golborne, Cecilia; Downie, Laura E
2017-07-01
To investigate whether oral, long-chain omega-3 (ω-3) essential fatty acid (EFA) supplementation, for 3 months, induces changes to the central corneal sub-basal nerve plexus in dry eye disease and whether nerve alterations correlate with clinical findings. This prospective, comparative study involved the final 12 participants enrolled in a randomised, double-masked, placebo-controlled clinical trial of 60 participants with moderate dry eye disease. Participants received either placebo (olive oil 1500 mg/day; n = 4) or ω-3 EFA supplements (~1000 mg/day eicosapentaenoic acid + ~500 mg/day docosahexaenoic acid; n = 8) for 90 days. The main outcome measure was the mean change in central corneal sub-basal plexus nerve parameters between days one and 90, quantified using in vivo confocal microscopy. Secondary outcomes included mean change in tear osmolarity, corneal dendritic cell density and basal epithelial cell density. Compared with baseline, the reduction in OSDI score and tear osmolarity at day 90 were greater in the ω-3 EFA group than the placebo group (OSDI: ω-3 EFA, mean ± SEM: -15.6 ± 2.8 vs placebo: -2.8 ± 4.1 units, t 5 = 2.6, p = 0.04; tearosmolarity: ω-3 EFA: -22.63 ± 5.7 vs placebo: -8 ± 2.7 mOsmol/L, t 9 = 2.3, p = 0.04). At day 90, corneal total nerve branch density (CTBD: 91.1 ± 8.6 vs 45.1 ± 13.4 branches/mm 2 , F 1,10 = 14, p = 0.004) and corneal nerve branch density on the main fibre (CNBD: 63.4 ± 6.5 vs 27.9 ± 11.5 branches/mm 2 , F 1,10 = 6, p = 0.03) were higher in the ω-3 EFA group compared with placebo. Relative to day 1, CNBD (branches/mm 2 ) increased at day 90 in the ω-3 EFA group (+20.0 ± 9.2, t 8 = 3.2 p = 0.01) compared with placebo (-10.8 ± 3.2). Similar changes were evident for corneal nerve fibre length (CNFL, mm/mm 2 ), which increased from baseline at day 90 in the omega-3 EFA group (+2.9 ± 1.6, t 8 = 3.4 p = 0.01) compared with placebo (-2.7 ± 0.5). There was a negative correlation between CTBD and tear osmolarity (r 10 = -0.70, p = 0.01). No significant changes were observed for basal epithelial cell or corneal dendritic cell density. These pilot study findings suggest that ω-3 EFA supplementation imparts neuroprotective effects in the corneal sub-basal plexus that correlate with the extent of tear osmolarity normalisation. © 2017 The Authors Ophthalmic & Physiological Optics © 2017 The College of Optometrists.
Vegetation ecology and carbon sequestration potential of shrubs in tropics of Chhattisgarh, India.
Jhariya, Manoj Kumar
2017-09-25
Tropical forests are well known to have great species diversity and contribute substantial share in terrestrial carbon (C) stocks worldwide. Shrubs are long-neglected life form in the forest ecosystem, playing many roles in the forest and human life. Shrub has great impact on vegetation attributes which in turn modify the C storage and capture. In the present investigation, an attempt has been made to explore the dynamics of shrub species in four fire regimes, viz. high, medium, low, and no fire zones of Bhoramdeo Wildlife Sanctuary of Kawardha forest division (Chhattisgarh), India. The variations in structure, diversity, biomass, productivity, and C sequestration potential in all the sites were quantified. The density and basal area of shrub varied from 1250 to 3750 individuals ha -1 and 2.79 to 4.92 m 2 ha -1 , respectively. The diversity indices showed that the value of Shannon index was highest in medium fire zone (3.77) followed by high, low, and no fire zones as 3.25, 3.12, and 2.32, respectively. The value of Simpson's index or concentration of dominance (Cd) ranged from 0.08 to 0.20, species richness from 0.56 to 1.58, equitability from 1.41 to 1.44, and beta diversity from 1.50 to 4.20, respectively. The total biomass and C storage ranged from 6.82 to 15.71 and from 2.93 to 6.76 t ha -1 , respectively. The shrub density, importance value index (IVI), and abundance to frequency ratio (A/F) significantly correlated between high fire and medium fire zone. The basal area was found to be significantly positively correlated between high fire and medium fire, and low and no fire zones, respectively. Two-way cluster analysis reflected various patterns of clustering due to influence of the forest fire which showed that some species have distant clustering while some have smaller cluster. Principal component analysis (PCA) reflects variable scenario with respect to shrub layer. Ventilago calyculata and Zizyphus rotundifolia showed higher correlation between themselves in terms of basal area (BA). The total shrub production was 1.59-3.53 t ha -1 year -1 while the C sequestration potential of 0.71-1.57 t ha -1 year -1 under different fire regimes. Shrub community in the medium fire zone reflected higher productivity and higher C sequestration in comparison to other fire zone. Among the different plant parts, the biomass accumulation ratio was highest in the root of shrub community among various fire regimes. Screening of species for restoration and different land-use pattern on the basis of biomass accumulation and carbon sequestering potential would be an effective strategy for decision-making in sustainable forest management.
Depboylu, Candan; Weihe, Eberhard; Eiden, Lee E
2012-01-01
The simian immunodeficiency virus (SIV) macaque model resembles human immunodeficiency virus-acquired immunodeficiency syndrome (AIDS) and associated brain dysfunction. Altered expression of synaptic markers and transmitters in neuro-AIDS has been reported, but limited data exist for the cholinergic system and lipid mediators such as prostaglandins. Here, we analyzed cholinergic basal forebrain neurons with their telencephalic projections and the rate-limiting enzymes for prostaglandin synthesis, cyclooxygenase isotypes 1 and 2 (COX1 and COX2) in the brains of SIV-infected macaques with or without encephalitis and antiretroviral therapy and uninfected controls.Cyclooxygenase isotype 1, but not COX2, was coexpressed with markers of cholinergic phenotype, that is, choline acetyltransferase and vesicular acetylcholine transporter (VAChT), in basal forebrain neurons of monkey, as well as human, brain. Cyclooxygenase isotype 1 was decreased in basal forebrain neurons in macaques with AIDS versus uninfected and asymptomatic SIV-infected macaques. The VAChT-positive fiber density was reduced in frontal, parietal, and hippocampal-entorhinal cortex. Although brain SIV burden and associated COX1- and COX2-positive mononuclear and endothelial inflammatory reactions were mostly reversed in AIDS-diseased macaques that received 6-chloro-2',3'-dideoxyguanosine treatment, decreased VAChT-positive terminal density and reduced cholinergic COX1 expression were not. Thus, COX1 expression is a feature of primate cholinergic basal forebrain neurons; it may be functionally important and a critical biomarker of cholinergic dysregulation accompanying lentiviral encephalopathy. These results further imply that insufficiently prompt initiation of antiretroviral therapy in lentiviral infection may lead to neurostructurally unremarkable but neurochemically prominent irreversible brain damage.
MR-DTI and PET multimodal imaging of dopamine release within subdivisions of basal ganglia
NASA Astrophysics Data System (ADS)
Tziortzi, A.; Searle, G.; Tsoumpas, C.; Long, C.; Shotbolt, P.; Rabiner, E.; Jenkinson, M.; Gunn, R. N.
2011-09-01
The basal ganglia is a group of anatomical nuclei, functionally organised into limbic, associative and sensorimotor regions, which plays a central role in dopamine related neurological and psychiatric disorders. In this study, we combine two imaging modalities to enable the measurement of dopamine release in functionally related subdivisions of the basal ganglia. [11C]-(+)-PHNO Positron Emission Tomography (PET) measurements in the living human brain pre- and post-administration of amphetamine allow for the estimation of regional dopamine release. Combined Magnetic Resonance Diffusion Tensor Imaging (MR-DTI) data allows for the definition of functional territories of the basal ganglia from connectivity information. The results suggest that there is a difference in dopamine release among the connectivity derived functional subdivisions. Dopamine release is highest in the limbic area followed by the sensorimotor and then the associative area with this pattern reflected in both striatum and pallidum.
Loehman, Rachel A.; Keane, Robert E.; Holsinger, Lisa M.; Wu, Zhiwei
2016-01-01
ContextInteractions among disturbances, climate, and vegetation influence landscape patterns and ecosystem processes. Climate changes, exotic invasions, beetle outbreaks, altered fire regimes, and human activities may interact to produce landscapes that appear and function beyond historical analogs.ObjectivesWe used the mechanistic ecosystem-fire process model FireBGCv2 to model interactions of wildland fire, mountain pine beetle (Dendroctonus ponderosae), and white pine blister rust (Cronartium ribicola) under current and future climates, across three diverse study areas.MethodsWe assessed changes in tree basal area as a measure of landscape response over a 300-year simulation period for the Crown of the Continent in north-central Montana, East Fork of the Bitterroot River in western Montana, and Yellowstone Central Plateau in western Wyoming, USA.ResultsInteracting disturbances reduced overall basal area via increased tree mortality of host species. Wildfire decreased basal area more than beetles or rust, and disturbance interactions modeled under future climate significantly altered landscape basal area as compared with no-disturbance and current climate scenarios. Responses varied among landscapes depending on species composition, sensitivity to fire, and pathogen and beetle suitability and susceptibility.ConclusionsUnderstanding disturbance interactions is critical for managing landscapes because forest responses to wildfires, pathogens, and beetle attacks may offset or exacerbate climate influences, with consequences for wildlife, carbon, and biodiversity.
Fisher, Simon D.; Reynolds, John N. J.
2014-01-01
Anatomical investigations have revealed connections between the intralaminar thalamic nuclei and areas such as the superior colliculus (SC) that receive short latency input from visual and auditory primary sensory areas. The intralaminar nuclei in turn project to the major input nucleus of the basal ganglia, the striatum, providing this nucleus with a source of subcortical excitatory input. Together with a converging input from the cerebral cortex, and a neuromodulatory dopaminergic input from the midbrain, the components previously found necessary for reinforcement learning in the basal ganglia are present. With this intralaminar sensory input, the basal ganglia are thought to play a primary role in determining what aspect of an organism’s own behavior has caused salient environmental changes. Additionally, subcortical loops through thalamic and basal ganglia nuclei are proposed to play a critical role in action selection. In this mini review we will consider the anatomical and physiological evidence underlying the existence of these circuits. We will propose how the circuits interact to modulate basal ganglia output and solve common behavioral learning problems of agency determination and action selection. PMID:24765070
Duursma, Remko A; Falster, Daniel S
2016-10-01
Here, we aim to understand differences in biomass distribution between major woody plant functional types (PFTs) (deciduous vs evergreen and gymnosperm vs angiosperm) in terms of underlying traits, in particular the leaf mass per area (LMA) and leaf area per unit stem basal area. We used a large compilation of plant biomass and size observations, including observations of 21 084 individuals on 656 species. We used a combination of semiparametric methods and variance partitioning to test the influence of PFT, plant height, LMA, total leaf area, stem basal area and climate on above-ground biomass distribution. The ratio of leaf mass to above-ground woody mass (MF /MS ) varied strongly among PFTs. We found that MF /MS at a given plant height was proportional to LMA across PFTs. As a result, the PFTs did not differ in the amount of leaf area supported per unit above-ground biomass or per unit stem basal area. Climate consistently explained very little additional variation in biomass distribution at a given plant size. Combined, these results demonstrate consistent patterns in above-ground biomass distribution and leaf area relationships among major woody PFTs, which can be used to further constrain global vegetation models. © 2016 The Authors. New Phytologist © 2016 New Phytologist Trust.
Clark, Jason A.; Loehman, Rachel A.; Keane, Robert E.
2017-01-01
We present landscape simulation results contrasting effects of changing climates on forest vegetation and fire regimes in Yellowstone National Park, USA, by mid-21st century. We simulated potential changes to fire dynamics and forest characteristics under three future climate projections representing a range of potential future conditions using the FireBGCv2 model. Under the future climate scenarios with moderate warming (>2°C) and moderate increases in precipitation (3–5%), model simulations resulted in 1.2–4.2 times more burned area, decreases in forest cover (10–44%), and reductions in basal area (14–60%). In these same scenarios, lodgepole pine (Pinus contorta) decreased in basal area (18–41%), while Douglas-fir (Pseudotsuga menziesii) basal area increased (21–58%). Conversely, mild warming (<2°C) coupled with greater increases in precipitation (12–13%) suggested an increase in forest cover and basal area by mid-century, with spruce and subalpine fir increasing in abundance. Overall, we found changes in forest tree species compositions were caused by the climate-mediated changes in fire regime (56–315% increase in annual area burned). Simulated changes in forest composition and fire regime under warming climates portray a landscape that shifts from lodgepole pine to Douglas-fir caused by the interaction between the magnitude and seasonality of future climate changes, by climate-induced changes in the frequency and intensity of wildfires, and by tree species response.
Remote Characterization of Ice Shelf Surface and Basal Processes: Examples from East Antarctica
NASA Astrophysics Data System (ADS)
Greenbaum, J. S.; Blankenship, D. D.; Grima, C.; Schroeder, D. M.; Soderlund, K. M.; Young, D. A.; Kempf, S. D.; Siegert, M. J.; Roberts, J. L.; Warner, R. C.; van Ommen, T. D.
2017-12-01
The ability to remotely characterize surface and basal processes of ice shelves has important implications for conducting systematic, repeatable, and safe evaluations of their stability in the context of atmospheric and oceanic forcing. Additionally, techniques developed for terrestrial ice shelves can be adapted to orbital radar sounding datasets planned for forthcoming investigations of icy moons. This has been made possible through recent advances in radar signal processing that enable these data to be used to test hypotheses derived from conceptual and numerical models of ice shelf- and ice shell-ocean interactions. Here, we present several examples of radar sounding-derived characterizations of surface and basal processes underway on ice shelves in East Antarctica. These include percolation of near-surface meltwater in warm austral summers, brine infiltration along ice shelf calving fronts, basal melt rate and distribution, and basal freeze distribution. On Europa, near-surface brines and their migration may impact local geological variability, while basal processes likely control the distribution of melt and freeze. Terrestrially, we emphasize radar-sounding records of the Totten Glacier Ice Shelf which hosts each of these processes as well as the highest known density of basal melt channels of any terrestrial ice shelf. Further, with a maximum floating ice thickness of over 2.5 km, the pressure at Totten's basal interface may be similar to that at Europa's ice-ocean interface; therefore, evaluating surface and basal processes of Totten Glacier and other ice shelves could serve as analogs for understanding melting processes of Europa's ice shell.
Pyramidal neurons in the septal and temporal CA1 field of the human and hedgehog tenrec hippocampus.
Liagkouras, Ioannis; Michaloudi, Helen; Batzios, Christos; Psaroulis, Dimitrios; Georgiadis, Marios; Künzle, Heinz; Papadopoulos, Georgios C
2008-07-07
The present study examines comparatively the cellular density of disector-counted/Nissl-stained CA1 pyramidal neurons and the morphometric characteristics (dendritic number/length, spine number/density and Sholl-counted dendritic branch points/20 microm) of the basal and apical dendritic systems of Golgi-impregnated CA1 neurons, in the septal and temporal hippocampus of the human and hedgehog tenrec brain. The obtained results indicate that in both hippocampal parts the cellular density of the CA1 pyramidal neurons is lower in human than in tenrec. However, while the human pyramidal cell density is higher in the septal hippocampal part than in the temporal one, in the tenrec the density of these cells is higher in the temporal part. The dendritic tree of the CA1 pyramidal cells, more developed in the septal than in temporal hippocampus in both species studied, is in general more complex in the human hippocampus. The basal and the apical dendritic systems exhibit species related morphometric differences, while dendrites of different orders exhibit differences in their number and length, and in their spine density. Finally, in both species, as well as hippocampal parts and dendritic systems, changes of dendritic morphometric features along ascending dendritic orders fluctuate in a similar way, as do the number of dendritic branch points in relation to the distance from the neuron soma.
Fertilization Increases Growth of Sawlog-Size Yellow-Poplar and Red Oak in West Virginia
N. I. Lamson
1978-01-01
Sawlog-size even-aged hardwood stands in north-central West Virginia were fertilized with N, P, and K, singly and in combinations. Applications of N alone increased the annual basal area growth of yellow-poplar more than that of red oak during the first 7 years after fertilization, whereas P alone increased the annual basal area growth of red oak more than that of...
Costs of reducing sapling basal area in thinned cherry-maple stands in West Virginia
Gary W. Miller
1984-01-01
Unmanaged 60-year-old cherry-maple stands in West Virginia were thinned to three levels of stocking according to the Allegheny hardwoods stocking guide. After the merchantable timber was removed, the basal area in saplings was reduced to less than 10 ft² per acre (2.3 m²/ha), as the guide recommends for stands with dense understories. A detailed time...
William W. Oliver
1979-01-01
Growth was analyzed after one thinning in a plantation of pole-size ponderosa and Jeffrey pines on land having a site index of 50 feet at 50 years. Periodic annual increment was determined for each of three 5-year periods. On this basis, increment in diameter and cubic volume were found to he related closely to stand basal area only. Basal area and height increment,...
Climate-tree growth models in relation to long-term growth trends of white oak in Pennsylvania
D. D. Davis; R. P. Long
2003-01-01
We examined long-term growth trends of white oak by comparing tree-ring chronologies developed from an old-growth stand, where the average tree age was 222 years, with a second-growth stand where average tree age was 78 years. Evaluation of basal area growth trends suggested that an anomalous decrease in basal area increment trend occurred in both stands during the...
Michael G. Shelton
2004-01-01
Abstract - Treatments were the following hardwood basal areas (square feet per acre) and spatial arrangements: 0, 15-grouped, 15-scattered, 30-scattered, and an untreated control. Pine basal area was reduced by harvesting to 60 square feet per acre in all treatments except the control. After six growing seasons, pine regeneration ranged from 8,890...
K. Tshireletso; J. C. Malechek; D. L. Bartos
2010-01-01
The objective of the study was to determine the effects of season and intensity of clipping using simulated browsing on suckers' (Populus tremulaides Michx.) basal area growth on Cedar Mountain, Southern Utah, Western United States of America. Three randomly selected stands measuring 70 m x 70 m were clear-felled in mid-July, 2005, and fenced. Simulated browsing...
J.D. Burton
1980-01-01
Second-growth even-aged loblolly-shortleaf pine stands on good and medium sites were thinned from above or below to a basal area of 70 ft2, 85 ft2, and 100 ft2/acre, to an increasing basal area, or according to the judgment of a committee. Treatments began at age 20 for original plots and at age 25...
Response of a 110-year-old Douglas-fir stand to urea and ammonium nitrate fertilization
Constance A. Harrington; Richard E. Miller
1979-01-01
Basal area response to 150 pounds of nitrogen per acre applied as urea or ammonium nitrate was monitored on 1/5-acre plots for 4 years in a recently thinned, 110-year-old, site II, Douglas-fir stand. Nitrogen fertilization significantly increased growth. Basal area increment was increased 59 percent over the controls by ammonium nitrate and 37 percent by urea. The...
Paul A. Murphy; David L. Graney
1988-01-01
Models were developed for individual-tree basal area growth, survival, and total heights for different species of upland hardwoods in the Boston Mountains of north Arkansas. Data used were from 87 permanent plots located in an array of different sites and stand ages; the plots were thinned to different stocking levels and included unthinned controls. To test these...
Jeffrey H. Gove
2003-01-01
This note seeks to extend the utility of size-biased distribution theory as applied to forestry through two relationships regarding the quadratic mean stand diameter. First, the quadratic mean stand diameter's relationship to the harmonic mean basal area for horizontal point sampling, which has been known algebraically from early on, is proved under size-biased...
Site condition, structure, and growth of baldcypress along tidal/non-tidal salinity gradients
Krauss, K.W.; Duberstein, J.A.; Doyle, T.W.; Conner, W.H.; Day, Richard H.; Inabinette, L.W.; Whitbeck, J.L.
2009-01-01
This report documents changes in forest structure and growth potential of dominant trees in salt-impacted tidal and non-tidal baldcypress wetlands of the southeastern United States. We inventoried basal area and tree height, and monitored incremental growth (in basal area) of codominant baldcypress (Taxodium distichum) trees monthly, for over four years, to examine the inter-relationships among growth, site fertility, and soil physico-chemical characteristics. We found that salinity, soil total nitrogen (TN), flood duration, and flood frequency affected forest structure and growth the greatest. While mean annual site salinity ranged from 0.1 to 3.4 ppt, sites with salinity concentrations of 1.3 ppt or greater supported a basal area of less than 40 m2/ha. Where salinity was < 0.7 ppt, basal area was as high as 87 m2/ha. Stand height was also negatively affected by higher salinity. However, salinity related only to soil TN concentrations or to the relative balance between soil TN and total phosphorus (TP), which reached a maximum concentration between 1.2 and 2.0 ppt salinity. As estuarine influence shifts inland with sea-level rise, forest growth may become more strongly linked to salinity, not only due to salt effects but also as a consequence of site nitrogen imbalance.
Forest discrimination with multipolarization imaging radar
NASA Technical Reports Server (NTRS)
Ford, J. P.; Wickland, D. E.
1985-01-01
The use of radar polarization diversity for discriminating forest canopy variables on airborne synthetic-aperture radar (SAR) images is evaluated. SAR images were acquired at L-Band (24.6 cm) simultaneously in four linear polarization states (HH, HV, VH, and VV) in South Carolina on March 1, 1984. In order to relate the polarization signatures to biophysical properties, false-color composite images were compared to maps of forest stands in the timber compartment. In decreasing order, the most useful correlative forest data are stand basal area, forest age, site condition index, and forest management type. It is found that multipolarization images discriminate variation in tree density and difference in the amount of understory, but do not discriminate between evergreen and deciduous forest types.
The corneal nerve density in the sub-basal plexus decreases with increasing myopia: a pilot study.
Harrison, Wendy W; Putnam, Nicole M; Shukis, Christine; Nguyen, Evelyn; Reinard, Kristen; Hundelt, Elizabeth; Vardanyan, Galina; Gabai, Celine; Yevseyenkov, Vladimir
2017-07-01
Myopia can cause many changes in the health of the eye. As it becomes more prevalent worldwide, more patients seek correction in the form of glasses, contact lenses and refractive surgery. In this study we explore the impact that high myopia has on central corneal nerve density by comparing sub basal nerve plexus density measured by confocal microscopy in a variety of refractive errors. Seventy healthy adult subjects between the ages of 21-50 years participated in this study. The study took place in two phases with no overlapping subjects (n = 30 phase 1 and n = 40 phase 2). In both phases an autorefraction, keratometry reading, corneal thickness measure and confocal corneal scan of the sub basal nerve plexus were performed for both eyes. There were 11 hyperopes (+0.50 to +3.50DS), six emmetropes (-0.25 to +0.50DS), 30 low myopes (-5.50 to -0.50DS), and 23 high myopes (-5.50DS and above). In the second phase of the study additional tests were performed including an axial length, additional corneal scans, and a questionnaire that asked about age of first refractive correction and contact lens wear. Corneal nerves were imaged over the central cornea with a Nidek CS4 confocal microscope (460 × 345 μm field). Nerves were evaluated using the NeuronJ program for density calculation. One eye was selected for inclusion based on image quality and higher refractive error (more myopic or hyperopic). As myopia increased, nerve density decreased (t 1 = 3.86, p < 0.001). We also note a decrease in data scatter above -7 D. The relationship between axial length values and nerve density was also significant and the slope was not as robust as refractive error (t 1 = 2.4, p < 0.04). As expected there was a significant difference between the four groups in axial length (F 3 = 19.9, p < 0.001) and age of first refractive correction of the myopic groups (14.9 vs 11.5 years; t 46 = 2.99 p < 0.01). There was no difference in keratometry readings or corneal thickness between the groups (F 3 = 0.6, p = 0.66 and F 3 = 1.2, p = 0.33 respectively). Corneal nerve density in the sub-basal plexus decreased with increasing myopia. This could have implications for corneal surgery and contact lens wear in this patient population. © 2017 The Authors Ophthalmic & Physiological Optics © 2017 The College of Optometrists.
Avecillas-Chasin, Josué M; Rascón-Ramírez, Fernando; Barcia, Juan A
2016-05-01
The cortico-basal ganglia and corticothalamic projections have been extensively studied in the context of neurological and psychiatric disorders. Deep brain stimulation (DBS) is known to modulate many of these pathways to produce the desired clinical effect. The aim of this work is to describe the anatomy of the main circuits of the basal ganglia using tractography in a surgical planning station. We used imaging studies of 20 patients who underwent DBS for movement and psychiatric disorders. We segmented the putamen, caudate nucleus (CN), thalamus, and subthalamic nucleus (STN), and we also segmented the cortical areas connected with these subcortical areas. We used tractography to define the subdivisions of the basal ganglia and thalamus through the generation of fibers from the cortical areas to the subcortical structures. We were able to generate the corticostriatal and corticothalamic connections involved in the motor, associative and limbic circuits. Furthermore, we were able to reconstruct the hyperdirect pathway through the corticosubthalamic connections and we found subregions in the STN. Finally, we reconstructed the cortico-subcortical connections of the ventral intermediate nucleus, the nucleus accumbens and the CN. We identified a feasible delineation of the basal ganglia and thalamus connections using tractography. These results could be potentially useful in DBS if the parcellations are used as targets during surgery. © 2016 Wiley Periodicals, Inc.
2010-01-01
Background Drought is a common stressor in many regions of the world and current climatic global circulation models predict further increases in warming and drought in the coming decades in several of these regions, such as the Mediterranean basin. The changes in leaf water content, distribution and dynamics in plant tissues under different soil water availabilities are not well known. In order to fill this gap, in the present report we describe our study withholding the irrigation of the seedlings of Quercus ilex, the dominant tree species in the evergreen forests of many areas of the Mediterranean Basin. We have monitored the gradual changes in water content in the different leaf areas, in vivo and non-invasively, by 1H magnetic resonance imaging (MRI) using proton density weighted (ρw) images and spin-spin relaxation time (T2) maps. Results ρw images showed that the distal leaf area lost water faster than the basal area and that after four weeks of similar losses, the water reduction was greater in leaf veins than in leaf parenchyma areas and also in distal than in basal leaf area. There was a similar tendency in all different areas and tissues, of increasing T2 values during the drought period. This indicates an increase in the dynamics of free water, suggesting a decrease of cell membranes permeability. Conclusions The results indicate a non homogeneous leaf response to stress with a differentiated capacity to mobilize water between its different parts and tissues. This study shows that the MRI technique can be a useful tool to follow non-intrusively the in vivo water content changes in the different parts of the leaves during drought stress. It opens up new possibilities to better characterize the associated physiological changes and provides important information about the different responses of the different leaf areas what should be taken into account when conducting physiological and metabolic drought stress studies in different parts of the leaves during drought stress. PMID:20735815
Sardans, Jordi; Peñuelas, Josep; Lope-Piedrafita, Silvia
2010-08-24
Drought is a common stressor in many regions of the world and current climatic global circulation models predict further increases in warming and drought in the coming decades in several of these regions, such as the Mediterranean basin. The changes in leaf water content, distribution and dynamics in plant tissues under different soil water availabilities are not well known. In order to fill this gap, in the present report we describe our study withholding the irrigation of the seedlings of Quercus ilex, the dominant tree species in the evergreen forests of many areas of the Mediterranean Basin. We have monitored the gradual changes in water content in the different leaf areas, in vivo and non-invasively, by 1H magnetic resonance imaging (MRI) using proton density weighted (rhow) images and spin-spin relaxation time (T2) maps. Rhow images showed that the distal leaf area lost water faster than the basal area and that after four weeks of similar losses, the water reduction was greater in leaf veins than in leaf parenchyma areas and also in distal than in basal leaf area. There was a similar tendency in all different areas and tissues, of increasing T2 values during the drought period. This indicates an increase in the dynamics of free water, suggesting a decrease of cell membranes permeability. The results indicate a non homogeneous leaf response to stress with a differentiated capacity to mobilize water between its different parts and tissues. This study shows that the MRI technique can be a useful tool to follow non-intrusively the in vivo water content changes in the different parts of the leaves during drought stress. It opens up new possibilities to better characterize the associated physiological changes and provides important information about the different responses of the different leaf areas what should be taken into account when conducting physiological and metabolic drought stress studies in different parts of the leaves during drought stress.
Activity propagation in an avian basal ganglia-thalamo-cortical circuit essential for vocal learning
Kojima, Satoshi; Doupe, Allison J.
2009-01-01
In mammalian basal ganglia-thalamo-cortical circuits, GABAergic pallidal neurons are thought to ‘gate’ or modulate excitation in thalamus with their strong inhibitory inputs, and thus signal to cortex by pausing and permitting thalamic neurons to fire in response to excitatory drive. In contrast, in a homologous circuit specialized for vocal learning in songbirds, evidence suggests that pallidal neurons signal by eliciting postinhibitory rebound spikes in thalamus, which could occur even without any excitatory drive to thalamic neurons. To test whether songbird pallidal neurons can also communicate with thalamus by gating excitatory drive, as well as by postinhibitory rebound, we examined the activity of thalamic relay neurons in response to acute inactivation of the basal ganglia structure Area X; Area X contains the pallidal neurons that project to thalamus. Although inactivation of Area X should eliminate rebound-mediated spiking in thalamus, this manipulation tonically increases the firing rate of thalamic relay neurons, providing evidence that songbird pallidal neurons can gate tonic thalamic excitatory drive. We also found that the increased thalamic activity was fed forward to its target in the avian equivalent of cortex, which includes neurons that project to the vocal premotor area. These data raise the possibility that basal ganglia circuits can signal to cortex through thalamus both by generating postinhibitory rebound and by gating excitatory drive, and may switch between these modes depending on the statistics of pallidal firing. Moreover, these findings provide insight into the strikingly different disruptive effects of basal ganglia and ‘cortical’ lesions on songbird vocal learning. PMID:19369547
Zhang, Peng; Shang, Qingli; Ma, Jingxue; Hao, Yuhua; Ye, Cunxi
2017-03-20
To determine the correlation between the preoperative basal diameter of macular hole, the postoperative area of high autofluorescence (AF) in macula, and visual acuity in full-thickness macular hole. Forty-nine patients with full-thickness macular hole who underwent vitrectomy and C3F8 filling were reviewed. The preoperative diameter of macular hole, the 6 months postoperative area of high AF in macula if it existed, the length of inner segment/outer segment (IS/OS) defect, and visual acuity were obtained. The correlation between them was determined. At postoperative 6 months, the rate of high AF in macula was 63.3%. There were statistical differences between with and without high AF groups in postoperative best-corrected visual acuity (BCVA) (t = -2.751, p = 0.008), preoperative basal diameter of macular hole (t = -4.946, p = 0.00001), and postoperative length of IS/OS defect (t = -8.351, p<0.00001). Simple linear regression analysis showed high positive correlations between preoperative basal diameter of macular hole and area of high AF (p<0.00001, r = 0.893), postoperative length of IS/OS defect and area of high fundus AF (FAF) (p<0.00001, r = 0.779), and negative correlations between area of high AF and postoperative BCVA (p = 0.037, r = 0.375). There was low correlation between diameter of macular hole and postoperative BCVA (p = 0.112). The preoperative basal diameter of macular hole and postoperative length of IS/OS defect decides the postoperative area of high AF in macula to some degree, and the postoperative area of high AF in macula can be an evaluating indicator for poor macular function recovery.
Advancement of tree species across ecotonal borders into non-forested ecosystems
NASA Astrophysics Data System (ADS)
Hanberry, Brice B.; Hansen, Mark H.
2015-10-01
Woody species are increasing in density, causing transition to more densely wooded vegetation states, and encroaching across ecotonal borders into non-forested ecosystems. We examined USDA Forest Service Forest Inventory and Analysis data to identify tree species that have expanded longitudinally in range, particularly into the central United States. We analyzed compositional differences within ecological regions (i.e., subsections) in eastern and western ranges of species using repeated measures ANOVA. We considered differences in outer ranges to indicate range expansion or contraction. We also estimated the shift in forest area and basal area relative to the center of the US and compared change in deciduous forest land cover. Out of 80 candidate species, 22 species expanded to the west, seven species expanded to the east, and five species expanded in both directions. During the survey interval, eastern tree species advanced into the predominantly non-forested ecosystems of central United States. Eastern cottonwood, eastern hophornbeam, eastern redbud, honeylocust, Osage-orange, pecan, red mulberry, and Shumard oak represent some of the species that are advancing eastern forest boundaries across forest-grassland ecotones into the central United States. Forest land has shifted towards the center of the continent, as has the center of mean tree basal area, and a simple comparison of deciduous cover change also displayed forest advancement into the central United States from eastern forests. The expanding species may spread along riparian migration corridors that provide protection from drought. Humans use the advancing tree species for windbreaks, fencerows, and ornamental landscaping, while wildlife spread fruit seeds, which results in unintentional assisted migration, or translocation, to drier sites across the region.
Michael D. Cain
1993-01-01
Abstract.The effects of burning cycles and pine basal area levels were assessed on natural pine regeneration and hardwood development in uneven-aged stands of loblolly andshortleafpines (Pinw taeda L. and P. echinata Mill.). The treatments included an unburned control and prescribed winter burrs at 3-, 6-, and 9-yr intervals. Basal area treatments were 9, 14, 18, and...
Influence of managed pine stands and mixed pine/hardwood stands on well being of deer
Lowell K. Halls; Charles E. Boyd
1982-01-01
A 172-acre enclosure where all the hardwood trees were removed or deadened and a 167 -acre enclosure where hardwoods comprised 25 percent of the tree basal area were each stocked in 1965 with 3 white-tailed deer (1 buck and 2 does). In 1963, before any timber cutting practices were imposed, tree basal area averaged 111 sq. ft. per acre and forage yields averaged 260...
Cohen, Philip R
2017-03-22
Red dot basal cell carcinoma is a unique variant of basal cell carcinoma. Including the three patients described in this report, red dot basal cell carcinoma has only been described in seven individuals. This paper describes the features of two males and one female with red dot basal cell carcinoma and reviews the characteristics of other patients with this clinical subtype of basal cell carcinoma. A 70-year-old male developed a pearly-colored papule with a red dot in the center on his nasal tip. A 71-year-old male developed a red dot surrounded by a flesh-colored papule on his left nostril. Lastly, a 74-year-old female developed a red dot within an area of erythema on her left mid back. Biopsy of the lesions all showed nodular and/or superficial basal cell carcinoma. Correlation of the clinical presentation and pathology established the diagnosis of red dot basal cell carcinoma. The tumors were treated by excision using the Mohs surgical technique. Pubmed was searched with the keyword: basal, cell, cancer, carcinoma, dot, red, and skin. The papers generated by the search and their references were reviewed. Red dot basal cell carcinoma has been described in three females and two males; the gender was not reported in two patients. The tumor was located on the nose (five patients), back (one patient) and thigh (one patient). Cancer presented as a solitary small red macule or papule; often, the carcinoma was surrounded by erythema or a flesh-colored papule. Although basal cell carcinomas usually do not blanch after a glass microscope slide is pressed against them, the red dot basal cell carcinoma blanched after diascopy in two of the patients, resulting in a delay of diagnosis in one of these individuals. Dermoscopy may be a useful non-invasive modality for evaluating skin lesions when the diagnosis of red dot basal cell carcinoma is considered. Mohs surgery is the treatment of choice; in some of the patients, the ratio of the area of the postoperative wound to that of the preoperative cancer was greater than 12:1, demonstrating a significant lateral spread of the tumor beyond the observed clinical margins of the neoplasm. In conclusion, in a patient with a personal history of actinic keratosis or nonmelanoma skin cancer, the appearance of a new red dot in a sun-exposed site should prompt additional evaluation of the skin lesion to exclude or establish the diagnosis of red dot basal cell carcinoma.
2017-01-01
Red dot basal cell carcinoma is a unique variant of basal cell carcinoma. Including the three patients described in this report, red dot basal cell carcinoma has only been described in seven individuals. This paper describes the features of two males and one female with red dot basal cell carcinoma and reviews the characteristics of other patients with this clinical subtype of basal cell carcinoma. A 70-year-old male developed a pearly-colored papule with a red dot in the center on his nasal tip. A 71-year-old male developed a red dot surrounded by a flesh-colored papule on his left nostril. Lastly, a 74-year-old female developed a red dot within an area of erythema on her left mid back. Biopsy of the lesions all showed nodular and/or superficial basal cell carcinoma. Correlation of the clinical presentation and pathology established the diagnosis of red dot basal cell carcinoma. The tumors were treated by excision using the Mohs surgical technique. Pubmed was searched with the keyword: basal, cell, cancer, carcinoma, dot, red, and skin. The papers generated by the search and their references were reviewed. Red dot basal cell carcinoma has been described in three females and two males; the gender was not reported in two patients. The tumor was located on the nose (five patients), back (one patient) and thigh (one patient). Cancer presented as a solitary small red macule or papule; often, the carcinoma was surrounded by erythema or a flesh-colored papule. Although basal cell carcinomas usually do not blanch after a glass microscope slide is pressed against them, the red dot basal cell carcinoma blanched after diascopy in two of the patients, resulting in a delay of diagnosis in one of these individuals. Dermoscopy may be a useful non-invasive modality for evaluating skin lesions when the diagnosis of red dot basal cell carcinoma is considered. Mohs surgery is the treatment of choice; in some of the patients, the ratio of the area of the postoperative wound to that of the preoperative cancer was greater than 12:1, demonstrating a significant lateral spread of the tumor beyond the observed clinical margins of the neoplasm. In conclusion, in a patient with a personal history of actinic keratosis or nonmelanoma skin cancer, the appearance of a new red dot in a sun-exposed site should prompt additional evaluation of the skin lesion to exclude or establish the diagnosis of red dot basal cell carcinoma. PMID:28465868
Seyfori, Hossein; Ghasemi, Hossein Ali; Hajkhodadadi, Iman; Nazaran, Mohammad Hassan; Hafizi, Maryam
2018-05-01
The effects of water supplementation of chelated trace minerals (CTM, which is named Bonzaplex designed with chelate compounds technology) on growth performance, apparent total tract digestibility (ATTD) of minerals, and some blood metabolites, TM, and antioxidant enzyme values in African ostriches were investigated from 8 to 12 months of age. A total of 20 8-month-old ostriches (five birds in five replicate pens) was randomly allocated into one of the following four treatments: (1) control (basal diet + tap water), (2) low CTM (basal diet +100 mg/bird/day CTM powder in tap water), (3) medium CTM (basal diet +1 g/bird/day CTM powder in tap water), and (4) high CTM (basal diet +2 g/bird/day CTM powder in tap water). Compared with control, medium CTM improved (P < 0.05) daily weight gain and ATTD of phosphorous, zinc, and copper in 12-month-old ostriches. Furthermore, the feed conversion ratio was lower, and ATTD of magnesium was higher in the medium- and high-CTM groups than that in the control group (P < 0.05). At the end of the trial, ostriches receiving high-CTM treatment exhibited the lower (P < 0.05) serum triglyceride and very low-density lipoprotein cholesterol concentrations and higher copper levels compared to those of the control treatment. Supplementation of higher amounts of CTM (medium and high CTM) also increased the activity of serum superoxide dismutase (P < 0.05). No differences were detected for other blood parameters including glucose, total protein, albumin, cholesterol, high-density lipoprotein cholesterol, low-density lipoprotein cholesterol, iron, magnesium, and glutathione peroxidase values. In conclusion, supplementation of CTM at the level of 1 g/bird/day to the drinking water can be recommended for improving growth performance, mineral absorption, and antioxidant status of ostriches fed diets containing the recommended levels of inorganic TM.
Repeated wildfires alter forest recovery of mixed-conifer ecosystems.
Stevens-Rumann, Camille; Morgan, Penelope
2016-09-01
Most models project warmer and drier climates that will contribute to larger and more frequent wildfires. However, it remains unknown how repeated wildfires alter post-fire successional patterns and forest structure. Here, we test the hypothesis that the number of wildfires, as well as the order and severity of wildfire events interact to alter forest structure and vegetation recovery and implications for vegetation management. In 2014, we examined forest structure, composition, and tree regeneration in stands that burned 1-18 yr before a subsequent 2007 wildfire. Three important findings emerged: (1) Repeatedly burned forests had 15% less woody surface fuels and 31% lower tree seedling densities compared with forests that only experienced one recent wildfire. These repeatedly burned areas are recovering differently than sites burned once, which may lead to alternative ecosystem structure. (2) Order of burn severity (high followed by low severity compared with low followed by high severity) did influence forest characteristics. When low burn severity followed high, forests had 60% lower canopy closure and total basal area with 92% fewer tree seedlings than when high burn severity followed low. (3) Time between fires had no effect on most variables measured following the second fire except large woody fuels, canopy closure and tree seedling density. We conclude that repeatedly burned areas meet many vegetation management objectives of reduced fuel loads and moderate tree seedling densities. These differences in forest structure, composition, and tree regeneration have implications not only for the trajectories of these forests, but may reduce fire intensity and burn severity of subsequent wildfires and may be used in conjunction with future fire suppression tactics. © 2016 by the Ecological Society of America.
Shimizu, Michiru; Ishida, Atsushi; Tange, Takeshi; Yagi, Hisayoshi
2006-04-01
We tested the hypothesis that sapling growth following a sudden increase in solar irradiance is related to recovery from photoinhibition and the balance between rate of production of new leaves and rate of abscision of old leaves. Leaf gas exchange, chlorophyll fluorescence and relative growth rate (RGR) of stem basal area were measured following the sudden exposure of shade-grown (7% of full sunlight) saplings of four Shorea species to full sunlight. Sudden exposure to full sunlight resulted in an immediate and substantial reduction in dark-adapted quantum yield of photosystem II (Fv/Fm), followed by a gradual recovery in all species. Near light-saturated net assimilation rate (A max) and area-based leaf chlorophyll concentration ([Chl area]) also declined immediately after exposure. Eleven days after exposure, A max had recovered to pre-exposure values in all species, whereas [Chl area] had not recovered. Across species, RGR of stem basal area increased with increasing RGR of the number of leaves following exposure to full sunlight. The interspecific variations in RGR of stem basal area suggest that new leaf production is crucial for determining the potential growth of saplings following gap formation.
Radiogenic heat production in sedimentary rocks of the Gulf of Mexico Basin, south Texas
McKenna, T.E.; Sharp, J.M.
1998-01-01
Radiogenic heat production within the sedimentary section of the Gulf of Mexico basin is a significant source of heat. Radiogenic heat should be included in thermal models of this basin (and perhaps other sedimentary basins). We calculate that radiogenic heat may contribute up to 26% of the overall surface heat-flow density for an area in south Texas. Based on measurements of the radioactive decay rate of ??-particles, potassium concentration, and bulk density, we calculate radiogenic heat production for Stuart City (Lower Cretaceous) limestones, Wilcox (Eocene) sandstones and mudrocks, and Frio (Oligocene) sandstones and mudrocks from south Texas. Heat production rates range from a low of 0.07 ?? 0.01 ??W/m3 in clean Stuart City limestones to 2.21 ?? 0.24??W/m3 in Frio mudrocks. Mean heat production rates for Wilcox sandstones, Frio sandstones, Wilcox mudrocks, and Frio mudrocks are 0.88, 1.19, 1.50, and 1.72 ??W/m3, respectively. In general, the mudrocks produce about 30-40% more heat than stratigraphically equivalent sandstones. Frio rocks produce about 15% more heat than Wilcox rocks per unit volume of clastic rock (sandstone/mudrock). A one-dimensional heat-conduction model indicates that this radiogenic heat source has a significant effect on subsurface temperatures. If a thermal model were calibrated to observed temperatures by optimizing basal heat-flow density and ignoring sediment heat production, the extrapolated present-day temperature of a deeply buried source rock would be overestimated.Radiogenic heat production within the sedimentary section of the Gulf of Mexico basin is a significant source of heat. Radiogenic heat should be included in thermal models of this basin (and perhaps other sedimentary basins). We calculate that radiogenic heat may contribute up to 26% of the overall surface heat-flow density for an area in south Texas. Based on measurements of the radioactive decay rate of ??-particles, potassium concentration, and bulk density, we calculate radiogenic heat production for Stuart City (Lower Cretaceous) limestones, Wilcox (Eocene) sandstones and mudrocks, and Frio (Oligocene) sandstones and mudrocks from south Texas. Heat production rates range from a low of 0.07??0.01 ??W/m3 in clean Stuart City limestones to 2.21??0.24 ??W/m3 in Frio mudrocks. Mean heat production rates for Wilcox sandstones, Frio sandstones, Wilcox mudrocks, and Frio mudrocks are 0.88, 1.19, 1.50, and 1.72 ??W/m3, respectively. In general, the mudrocks produce about 30-40% more heat than stratigraphically equivalent sandstones. Frio rocks produce about 15% more heat than Wilcox rocks per unit volume of clastic rock (sandstone/mudrock). A one-dimensional heat-conduction model indicates that this radiogenic heat source has a significant effect on subsurface temperatures. If a thermal model were calibrated to observed temperatures by optimizing basal heat-flow density and ignoring sediment heat production, the extrapolated present-day temperature of a deeply buried source rock would be overestimated.
Larrañaga, Pedro; Benavides-Piccione, Ruth; Fernaud-Espinosa, Isabel; DeFelipe, Javier; Bielza, Concha
2017-01-01
We modeled spine distribution along the dendritic networks of pyramidal neurons in both basal and apical dendrites. To do this, we applied network spatial analysis because spines can only lie on the dendritic shaft. We expanded the existing 2D computational techniques for spatial analysis along networks to perform a 3D network spatial analysis. We analyzed five detailed reconstructions of adult human pyramidal neurons of the temporal cortex with a total of more than 32,000 spines. We confirmed that there is a spatial variation in spine density that is dependent on the distance to the cell body in all dendrites. Considering the dendritic arborizations of each pyramidal cell as a group of instances of the same observation (the neuron), we used replicated point patterns together with network spatial analysis for the first time to search for significant differences in the spine distribution of basal dendrites between different cells and between all the basal and apical dendrites. To do this, we used a recent variant of Ripley’s K function defined to work along networks. The results showed that there were no significant differences in spine distribution along basal arbors of the same neuron and along basal arbors of different pyramidal neurons. This suggests that dendritic spine distribution in basal dendritic arbors adheres to common rules. However, we did find significant differences in spine distribution along basal versus apical networks. Therefore, not only do apical and basal dendritic arborizations have distinct morphologies but they also obey different rules of spine distribution. Specifically, the results suggested that spines are more clustered along apical than in basal dendrites. Collectively, the results further highlighted that synaptic input information processing is different between these two dendritic domains. PMID:28662210
Anton-Sanchez, Laura; Larrañaga, Pedro; Benavides-Piccione, Ruth; Fernaud-Espinosa, Isabel; DeFelipe, Javier; Bielza, Concha
2017-01-01
We modeled spine distribution along the dendritic networks of pyramidal neurons in both basal and apical dendrites. To do this, we applied network spatial analysis because spines can only lie on the dendritic shaft. We expanded the existing 2D computational techniques for spatial analysis along networks to perform a 3D network spatial analysis. We analyzed five detailed reconstructions of adult human pyramidal neurons of the temporal cortex with a total of more than 32,000 spines. We confirmed that there is a spatial variation in spine density that is dependent on the distance to the cell body in all dendrites. Considering the dendritic arborizations of each pyramidal cell as a group of instances of the same observation (the neuron), we used replicated point patterns together with network spatial analysis for the first time to search for significant differences in the spine distribution of basal dendrites between different cells and between all the basal and apical dendrites. To do this, we used a recent variant of Ripley's K function defined to work along networks. The results showed that there were no significant differences in spine distribution along basal arbors of the same neuron and along basal arbors of different pyramidal neurons. This suggests that dendritic spine distribution in basal dendritic arbors adheres to common rules. However, we did find significant differences in spine distribution along basal versus apical networks. Therefore, not only do apical and basal dendritic arborizations have distinct morphologies but they also obey different rules of spine distribution. Specifically, the results suggested that spines are more clustered along apical than in basal dendrites. Collectively, the results further highlighted that synaptic input information processing is different between these two dendritic domains.
NASA Astrophysics Data System (ADS)
Passalacqua, Olivier; Ritz, Catherine; Parrenin, Frédéric; Urbini, Stefano; Frezzotti, Massimo
2017-09-01
Basal melt rate is the most important physical quantity to be evaluated when looking for an old-ice drilling site, and it depends to a great extent on the geothermal flux (GF), which is poorly known under the East Antarctic ice sheet. Given that wet bedrock has higher reflectivity than dry bedrock, the wetness of the ice-bed interface can be assessed using radar echoes from the bedrock. But, since basal conditions depend on heat transfer forced by climate but lagged by the thick ice, the basal ice may currently be frozen whereas in the past it was generally melting. For that reason, the risk of bias between present and past conditions has to be evaluated. The objective of this study is to assess which locations in the Dome C area could have been protected from basal melting at any time in the past, which requires evaluating GF. We used an inverse approach to retrieve GF from radar-inferred distribution of wet and dry beds. A 1-D heat model is run over the last 800 ka to constrain the value of GF by assessing a critical ice thickness, i.e. the minimum ice thickness that would allow the present local distribution of basal melting. A regional map of the GF was then inferred over a 80 km × 130 km area, with a N-S gradient and with values ranging from 48 to 60 mW m-2. The forward model was then emulated by a polynomial function to compute a time-averaged value of the spatially variable basal melt rate over the region. Three main subregions appear to be free of basal melting, two because of a thin overlying ice and one, north of Dome C, because of a low GF.
Miguel, Magalhaes Amade
2015-01-01
Shallow basal root growth angle (BRGA) increases phosphorus acquisition efficiency by enhancing topsoil foraging because in most soils, phosphorus is concentrated in the topsoil. Root hair length and density (RHL/D) increase phosphorus acquisition by expanding the soil volume subject to phosphorus depletion through diffusion. We hypothesized that shallow BRGA and large RHL/D are synergetic for phosphorus acquisition, meaning that their combined effect is greater than the sum of their individual effects. To evaluate this hypothesis, phosphorus acquisition in the field in Mozambique was compared among recombinant inbred lines of common bean (Phaseolus vulgaris) having four distinct root phenotypes: long root hairs and shallow basal roots, long root hairs and deep basal roots, short root hairs and shallow basal roots, and short root hairs and deep basal roots. The results revealed substantial synergism between BRGA and RHL/D. Compared with short-haired, deep-rooted phenotypes, long root hairs increased shoot biomass under phosphorus stress by 89%, while shallow roots increased shoot biomass by 58%. Genotypes with both long root hairs and shallow roots had 298% greater biomass accumulation than short-haired, deep-rooted phenotypes. Therefore, the utility of shallow basal roots and long root hairs for phosphorus acquisition in combination is twice as large as their additive effects. We conclude that the anatomical phene of long, dense root hairs and the architectural phene of shallower basal root growth are synergetic for phosphorus acquisition. Phene synergism may be common in plant biology and can have substantial importance for plant fitness, as shown here. PMID:25699587
Passive seismic monitoring of the Bering Glacier during its last surge event
NASA Astrophysics Data System (ADS)
Zhan, Z.
2017-12-01
The physical causes behind glacier surges are still unclear. Numerous evidences suggest that they probably involve changes in glacier basal conditions, such as switch of basal water system from concentrated large tunnels to a distributed "layer" as "connected cavities". However, most remote sensing approaches can not penetrate to the base to monitor such changes continuously. Here we apply seismic interferometry using ambient noise to monitor glacier seismic structures, especially to detect possible signatures of the hypothesized high-pressure water "layer". As an example, we derive an 11-year long history of seismic structure of the Bering Glacier, Alaska, covering its latest surge event. We observe substantial drops of Rayleigh and Love wavespeeds across the glacier during the surge event, potentially caused by changes in crevasse density, glacier thickness, and basal conditions.
Movement of basal plane dislocations in GaN during electron beam irradiation
DOE Office of Scientific and Technical Information (OSTI.GOV)
Yakimov, E. B.; National University of Science and Technology MISiS, Leninskiy pr. 4, Moscow 119049; Vergeles, P. S.
The movement of basal plane segments of dislocations in low-dislocation-density GaN films grown by epitaxial lateral overgrowth as a result of irradiation with the probing beam of a scanning electron microscope was detected by means of electron beam induced current. Only a small fraction of the basal plane dislocations was susceptible to such changes and the movement was limited to relatively short distances. The effect is explained by the radiation enhanced dislocation glide for dislocations pinned by two different types of pinning sites: a low-activation-energy site and a high-activation-energy site. Only dislocation segments pinned by the former sites can bemore » moved by irradiation and only until they meet the latter pinning sites.« less
Sustained High Basal Motion of the Greenland Ice Sheet Revealed by Borehole Deformation
NASA Technical Reports Server (NTRS)
Ryser, Claudia; Luthi, Martin P.; Andrews, Lauren C.; Hoffman, Matthew, J.; Catania, Ginny A.; Hawley, Robert L.; Neumann, Thomas A.; Kristensen, Steen S.
2014-01-01
Ice deformation and basal motion characterize the dynamical behavior of the Greenland ice sheet (GrIS). We evaluate the contribution of basal motion from ice deformation measurements in boreholes drilled to the bed at two sites in the western marginal zone of the GrIS. We find a sustained high amount of basal motion contribution to surface velocity of 44-73 percent in winter, and up to 90 percent in summer. Measured ice deformation rates show an unexpected variation with depth that can be explained with the help of an ice-flow model as a consequence of stress transfer from slippery to sticky areas. This effect necessitates the use of high-order ice-flow models, not only in regions of fast-flowing ice streams but in all temperate-based areas of the GrIS. The agreement between modeled and measured deformation rates confirms that the recommended values of the temperature-dependent flow rate factor A are a good choice for ice-sheet models.
Eagles-Smith, Collin A.; Herring, Garth; Johnson, Branden L.; Graw, Rick
2016-01-01
Remote high-elevation lakes represent unique environments for evaluating the bioaccumulation of atmospherically deposited mercury through freshwater food webs, as well as for evaluating the relative importance of mercury loading versus landscape influences on mercury bioaccumulation. The increase in mercury deposition to these systems over the past century, coupled with their limited exposure to direct anthropogenic disturbance make them useful indicators for estimating how changes in mercury emissions may propagate to changes in Hg bioaccumulation and ecological risk. We evaluated mercury concentrations in resident fish from 28 high-elevation, sub-alpine lakes in the Pacific Northwest region of the United States. Fish total mercury (THg) concentrations ranged from 4 to 438 ng/g wet weight, with a geometric mean concentration (±standard error) of 43 ± 2 ng/g ww. Fish THg concentrations were negatively correlated with relative condition factor, indicating that faster growing fish that are in better condition have lower THg concentrations. Across the 28 study lakes, mean THg concentrations of resident salmonid fishes varied as much as 18-fold among lakes. We used a hierarchal statistical approach to evaluate the relative importance of physiological, limnological, and catchment drivers of fish Hg concentrations. Our top statistical model explained 87% of the variability in fish THg concentrations among lakes with four key landscape and limnological variables: catchment conifer density (basal area of conifers within a lake's catchment), lake surface area, aqueous dissolved sulfate, and dissolved organic carbon. Conifer density within a lake's catchment was the most important variable explaining fish THg concentrations across lakes, with THg concentrations differing by more than 400 percent across the forest density spectrum. These results illustrate the importance of landscape characteristics in controlling mercury bioaccumulation in fish.
Eagles-Smith, Collin A; Herring, Garth; Johnson, Branden; Graw, Rick
2016-05-01
Remote high-elevation lakes represent unique environments for evaluating the bioaccumulation of atmospherically deposited mercury through freshwater food webs, as well as for evaluating the relative importance of mercury loading versus landscape influences on mercury bioaccumulation. The increase in mercury deposition to these systems over the past century, coupled with their limited exposure to direct anthropogenic disturbance make them useful indicators for estimating how changes in mercury emissions may propagate to changes in Hg bioaccumulation and ecological risk. We evaluated mercury concentrations in resident fish from 28 high-elevation, sub-alpine lakes in the Pacific Northwest region of the United States. Fish total mercury (THg) concentrations ranged from 4 to 438 ng/g wet weight, with a geometric mean concentration (±standard error) of 43 ± 2 ng/g ww. Fish THg concentrations were negatively correlated with relative condition factor, indicating that faster growing fish that are in better condition have lower THg concentrations. Across the 28 study lakes, mean THg concentrations of resident salmonid fishes varied as much as 18-fold among lakes. We used a hierarchal statistical approach to evaluate the relative importance of physiological, limnological, and catchment drivers of fish Hg concentrations. Our top statistical model explained 87% of the variability in fish THg concentrations among lakes with four key landscape and limnological variables: catchment conifer density (basal area of conifers within a lake's catchment), lake surface area, aqueous dissolved sulfate, and dissolved organic carbon. Conifer density within a lake's catchment was the most important variable explaining fish THg concentrations across lakes, with THg concentrations differing by more than 400 percent across the forest density spectrum. These results illustrate the importance of landscape characteristics in controlling mercury bioaccumulation in fish. Published by Elsevier Ltd.
Robert L. Deal; Troy Heithecker; Eric K. Zenner
2010-01-01
The effects of partial cutting on tree size structure and stand growth were evaluated in 52 plots in 13 stands in southeast Alaska that were partially harvested 53 to 96 years ago and compared with 50-year-old even-aged stands that developed after clearcutting. The net basal-area growth was greater in the partially cut plots than in the uncut plots, and basal-area...
Yanagihara, Shin; Hessler, Neal A.
2011-01-01
Reactivations of waking experiences during sleep have been considered fundamental neural processes for memory consolidation. In songbirds, evidence suggests the importance of sleep-related neuronal activity in song system motor pathway nuclei for both juvenile vocal learning and maintenance of adult song. Like those in singing motor nuclei, neurons in the basal ganglia nucleus Area X, part of the basal ganglia-thalamocortical circuit essential for vocal plasticity, exhibit singing-related activity. It is unclear, however, whether Area X neurons show any distinctive spiking activity during sleep similar to that during singing. Here we demonstrate that, during sleep, Area X pallidal neurons exhibit phasic spiking activity, which shares some firing properties with activity during singing. Shorter interspike intervals that almost exclusively occurred during singing in awake periods were also observed during sleep. The level of firing variability was consistently higher during singing and sleep than during awake non-singing states. Moreover, deceleration of firing rate, which is considered to be an important firing property for transmitting signals from Area X to the thalamic nucleus DLM, was observed mainly during sleep as well as during singing. These results suggest that songbird basal ganglia circuitry may be involved in the off-line processing potentially critical for vocal learning during sensorimotor learning phase. PMID:21991379
Regional distribution of neuropeptide Y mRNA in postmortem human brain.
Brené, S; Lindefors, N; Kopp, J; Sedvall, G; Persson, H
1989-12-01
The distribution of messenger RNA encoding neuropeptide Y (NPY) was studied in 11 different postmortem human brain regions using in situ hybridization histochemistry, and RNA blot analysis. In situ hybridization data revealed that the highest numerical density of labeled cells corresponded to neurons in accumbens area, caudate nucleus, putamen, and substantia innominata. Significantly fewer NPY mRNA-containing neurons were found in frontal and parietal cortex, amygdaloid body and dentate gyrus. No NPY mRNA-containing cells were found in substantia nigra. NPY mRNA-positive neurons from all regions studied showed relatively similar labeling, as revealed by computerized image analysis. Blot analysis showed an approximately 0.8 kb NPY mRNA in all brain regions studied, except in substantia nigra and cerebellum. Densitometric scanning of the autoradiograms revealed levels of NPY mRNA in the following order: putamen greater than caudate nucleus greater than frontal cortex (Brodmann areas 4 and 6) greater than temporal cortex (Brodmann area 38) greater than parietal cortex (Brodmann areas 5 and 7) greater than frontal cortex (Brodmann area 11). Hence, although NPY mRNA is widely distributed in neurons of the human brain large regional variation exists, with the highest expression in accumbens area and parts of the basal ganglia.
Erkan Turan, Kadriye; Kocabeyoglu, Sibel; Bekircan-Kurt, Can Ebru; Bezci, Figen; Erdem-Ozdamar, Sevim; Irkec, Murat
2018-03-01
To evaluate ocular surface alterations and characteristics of corneal basal epithelium and subbasal nerves in patients with myasthenia gravis. Myasthenia gravis patients (n = 21) and healthy controls (n = 20) were enrolled. All participants underwent ocular surface testing in the following order: tear break-up time, lissamine green staining, Schirmer I test with anesthesia, and Ocular Surface Disease Index questionnaire. The Cochet-Bonnet esthesiometer was used to measure corneal sensitivity. Basal epithelial cells and subbasal nerves were evaluated using in vivo confocal microscopy. Myasthenia gravis patients had higher Ocular Surface Disease Index score (13.9 ± 15.0 vs 1.4 ± 2.2, p < 0.001) and lissamine green staining score (0.6 ± 0.4 vs 0.2 ± 0.4, p = 0.007). Break-up time score (9.3 ± 3.0 vs 9.9 ± 1.9, p = 0.481) and Schirmer I test score (16.5 ± 9.2 vs 19.3 ± 8.4, p = 0.323) did not differ significantly. Corneal sensation was 0.4 g/mm 2 in all eyes. Patients with myasthenia gravis had lower basal epithelial cell density (3775.7 ± 938.1 vs 4983.1 ± 608.5, p < 0.001) and total nerve density (1956.1 ± 373.3 vs 2277.9 ± 405.0, p = 0.012) and higher subbasal nerve tortuosity (1.9 ± 0.8 vs 1.6 ± 0.7, p = 0.007) than controls. A significant increase in Ocular Surface Disease Index scores was found with decreasing basal epithelial cell density (rho = -0.518, p = 0.001). There was a significantly moderate negative correlation between the duration of myasthenia gravis and the number of corneal nerves (rho = -0.497, p = 0.022). Significant alterations of basal epithelial cells and subbasal nerves were demonstrated in myasthenia gravis patients although there was no difference of corneal sensitivity between myasthenia gravis patients and healthy controls. Thus, it should be borne in mind that myasthenia gravis patients deserve further evaluation with regard to ocular surface disease.
Rolle, Teresa; Spinetta, Roberta; Nuzzi, Raffaele
2017-08-03
The effects of preservatives of antiglaucoma medications on corneal surface and tear function have been widely shown in literature; it's not the same as regards the active compounds themselves. The purpose of our study was to compare Ocular Surface Disease (OSD) signs and symptoms of Tafluprost 0.0015% versus preservative free (PF) Timolol 0.1% eyedrops in ocular hypertensive (OH) and in primary open-angle glaucoma (POAG) patients. A cross-sectional study included patients in monotherapy for at least 36 months with Tafluprost 0.0015% (27) or PF Timolol 0.1% (24) and 20 healthy age and sex-matched volunteers. All subjects underwent clinical tests (Schirmer I and break-up time), in vivo confocal microscopy (IVCM) and were surveyed using Ocular Surface Disease Index (OSDI) and Glaucoma Symptoms Scale (GSS) questionnaires. The groups were compared with ANOVA, Kruskal-Wallis test, t-test, Mann-Whitney test and Bonferroni's adjustment of p-values. No significant differences were found in questionnaires scores, clinical tests, IVCM variables between therapy groups. Tafluprost 0.0015% group showed significantly higher OSDI score, basal epithelial cells density, stromal reflectivity, sub-basal nerves tortuosity (p = 0.0000, 0.037, 0.006, 0.0000) and less GSS score, number of sub-basal nerves (p = 0.0000, 0.037) than controls but similar clinical tests results (p > 0.05). PF Timolol group had significantly higher OSDI score, basal epithelial cells density, stromal reflectivity and sub-basal nerve tortuosity (p = 0.000, 0.014, 0.008, 0.002), less GSS score, BUT and number of sub-basal nerves (p = 0.0000, 0.026, 0.003) than controls. Compared to PF Timolol 0.1%, Tafluprost 0.0015% showed similar safety with regards to tear function and corneal status and a similar tolerability profile. Both therapy groups show some alterations in corneal microstructure but no side effects on tear function except for an increased tear instability in PF Timolol 0.1% group. Ophtalmologists should be aware that even PF formulations may lead to a mild ocular surface impairment.
Field measurement of basal forces generated by erosive debris flows
McCoy, S.W.; Tucker, G.E.; Kean, J.W.; Coe, J.A.
2013-01-01
It has been proposed that debris flows cut bedrock valleys in steeplands worldwide, but field measurements needed to constrain mechanistic models of this process remain sparse due to the difficulty of instrumenting natural flows. Here we present and analyze measurements made using an automated sensor network, erosion bolts, and a 15.24 cm by 15.24 cm force plate installed in the bedrock channel floor of a steep catchment. These measurements allow us to quantify the distribution of basal forces from natural debris‒flow events that incised bedrock. Over the 4 year monitoring period, 11 debris‒flow events scoured the bedrock channel floor. No clear water flows were observed. Measurements of erosion bolts at the beginning and end of the study indicated that the bedrock channel floor was lowered by 36 to 64 mm. The basal force during these erosive debris‒flow events had a large‒magnitude (up to 21 kN, which was approximately 50 times larger than the concurrent time‒averaged mean force), high‒frequency (greater than 1 Hz) fluctuating component. We interpret these fluctuations as flow particles impacting the bed. The resulting variability in force magnitude increased linearly with the time‒averaged mean basal force. Probability density functions of basal normal forces were consistent with a generalized Pareto distribution, rather than the exponential distribution that is commonly found in experimental and simulated monodispersed granular flows and which has a lower probability of large forces. When the bed sediment thickness covering the force plate was greater than ~ 20 times the median bed sediment grain size, no significant fluctuations about the time‒averaged mean force were measured, indicating that a thin layer of sediment (~ 5 cm in the monitored cases) can effectively shield the subjacent bed from erosive impacts. Coarse‒grained granular surges and water‒rich, intersurge flow had very similar basal force distributions despite differences in appearance and bulk‒flow density. These results demonstrate that debris flows can have strong control on rates of steepland evolution and contribute to a foundation needed for modeling debris‒flow incision stochastically.
NASA Astrophysics Data System (ADS)
Keller, M. M.; d'Oliveira, M. N.; Takemura, C. M.; Vitoria, D.; Araujo, L. S.; Morton, D. C.
2012-12-01
Selective logging, the removal of several valuable timber trees per hectare, is an important land use in the Brazilian Amazon and may degrade forests through long term changes in structure, loss of forest carbon and species diversity. Similar to deforestation, the annual area affected by selected logging has declined significantly in the past decade. Nonetheless, this land use affects several thousand km2 per year in Brazil. We studied a 1000 ha area of the Antimary State Forest (FEA) in the State of Acre, Brazil (9.304 ○S, 68.281 ○W) that has a basal area of 22.5 m2 ha-1 and an above-ground biomass of 231 Mg ha-1. Logging intensity was low, approximately 10 to 15 m3 ha-1. We collected small-footprint airborne lidar data using an Optech ALTM 3100EA over the study area once each in 2010 and 2011. The study area contained both recent and older logging that used both conventional and technologically advanced logging techniques. Lidar return density averaged over 20 m-2 for both collection periods with estimated horizontal and vertical precision of 0.30 and 0.15 m. A relative density model comparing returns from 0 to 1 m elevation to returns in 1-5 m elevation range revealed the pattern of roads and skid trails. These patterns were confirmed by ground-based GPS survey. A GIS model of the road and skid network was built using lidar and ground data. We tested and compared two pattern recognition approaches used to automate logging detection. Both segmentation using commercial eCognition segmentation and a Frangi filter algorithm identified the road and skid trail network compared to the GIS model. We report on the effectiveness of these two techniques.
Widespread increase of tree mortality rates in the Western United States
van Mantgem, P.J.; Stephenson, N.L.; Byrne, J.C.; Daniels, L.D.; Franklin, J.F.; Fule, P.Z.; Harmon, M.E.; Larson, A.J.; Smith, Joseph M.; Taylor, A.H.; Veblen, T.T.
2009-01-01
Persistent changes in tree mortality rates can alter forest structure, composition, and ecosystem services such as carbon sequestration. Our analyses of longitudinal data from unmanaged old forests in the western United States showed that background (noncatastrophic) mortality rates have increased rapidly in recent decades, with doubling periods ranging from 17 to 29 years among regions. Increases were also pervasive across elevations, tree sizes, dominant genera, and past fire histories. Forest density and basal area declined slightly, which suggests that increasing mortality was not caused by endogenous increases in competition. Because mortality increased in small trees, the overall increase in mortality rates cannot be attributed solely to aging of large trees. Regional warming and consequent increases in water deficits are likely contributors to the increases in tree mortality rates.
In vivo hyperspectral imaging and differentiation of skin cancer
NASA Astrophysics Data System (ADS)
Zherdeva, Larisa A.; Bratchenko, Ivan A.; Myakinin, Oleg O.; Moryatov, Alexander A.; Kozlov, Sergey V.; Zakharov, Valery P.
2016-10-01
Results of hyperspectral imaging analysis for in vivo visualization of skin neoplasms are presented. 16 melanomas, 19 basal cell carcinomas and 10 benign tumors with different stages of neoplasm growth were tested. The HSI system provide skin tissue images with 5 nm spectral resolution in the range of 450-750 nm with automatic stabilization of each frame compensating displacement of the scanning area due to spontaneous macro-movements of the patient. The integrated optical densities in 530-600 and 600-670 nm ranges are used for real-time hemoglobin and melanin distribution imaging in skin tissue. It was shown that the total accuracy of skin cancer identification exceeds 90% and 70% for differentiation of melanomas from BCC and begihn tumors. It was demonstrated the possibility for HSI classification of melanomas of different stages.
Basal cell carcinomas in elderly patients treated by cryotherapy
Chiriac, Anca; Mihaila, Doina; Foia, Liliana; Solovan, Caius
2013-01-01
Basal cell carcinoma is a malignant skin tumor with high incidence in our country, especially in rural areas, on sun-exposed skin (particularly on the face) in elderly patients. We present three cases of basal cell carcinoma with good results with cryotherapy. This report aims to outline and to prove that in some difficult situations, a simple, inexpensive, easy-to-perform procedure with no contraindications and with minimal side effects (erythema, mild pain) can be applied and resolve such cases. PMID:23569366
Müller glial cells of the primate foveola: An electron microscopical study.
Syrbe, Steffen; Kuhrt, Heidrun; Gärtner, Ulrich; Habermann, Gunnar; Wiedemann, Peter; Bringmann, Andreas; Reichenbach, Andreas
2018-02-01
Previous studies on the ultrastructure of the primate foveola suggested the presence of an inverted cone-like structure which is formed by 25-35 specialized Müller cells overlying the area of high photoreceptor density. We investigated the ultrastructure of the Müller cells in the foveola of a human and macaque retina. Sections through the posterior poles of an eye of a 40 years-old human donor and an eye of an adult cynomolgus monkey (Macaca fascicularis) were investigated with transmission electron microscopy. The foveola consisted of an inner layer (thickness, 5.5-12 μm) which mainly contained somata (including nuclei) and inner processes of Müller cells; this layer overlaid the central Henle fibers and outer nuclear layer. The inner layer contained numerous watery cysts and thin lamelliform and tubular Müller cell processes which spread along the inner limiting membrane (ILM). The cytoplasm of the outer Müller cell processes became increasingly dispersed and electron-lucent in the course towards the outer limiting membrane. The ILM of the foveola was formed by a very thin basal lamina (thickness, <40 nm) while the basal lamina of the parafovea was thick (0.9-1 μm). The data show that there are various conspicuous features of foveolar Müller cells. The numerous thin Müller cell processes below the ILM may smooth the inner surface of the foveola (to minimize image distortion resulting from varying light refraction angles at an uneven retinal surface), create additional barriers to the vitreous cavity (compensating the thinness of the ILM), and provide mechanical stability to the tissue. The decreasing density of the outer process cytoplasm may support the optical function of the foveola. Copyright © 2017. Published by Elsevier Ltd.
Stopa, E G; Koh, E T; Svendsen, C N; Rogers, W T; Schwaber, J S; King, J C
1991-06-01
Immunocytochemistry performed on 80-microns unembedded tissue sections was used to study the localization of GnRH-containing neurons and fibers in the basal forebrain and amygdala of six adult (four male, two female) human brains. Sections from one of the female brains were subjected to computer-assisted microscopic mapping to generate a three-dimensional analysis of immunoreactive structures. In all six brains examined, cell bodies were concentrated in the preoptic area and basal hypothalamus, but were also evident in the septal region, anterior olfactory area, and cortical and medial amygdaloid nuclei. GnRH-containing fibers were observed within the hypothalamus (predominantly infundibular region and preoptic area), septum, stria terminalis, ventral pallidum, dorsomedial thalamus, olfactory stria, and anterior olfactory area. Many fibers could also be seen coursing along the base of the brain between the hypothalamus and cortical and medial amygdaloid nuclei. The localization of GnRH-containing cells and fibers in several of these areas represents new observations in the human brain and suggests a role for the amygdaloid complex in the regulation of gonadotropin secretion. The comprehensive view provided by these data may be useful in the clinical application of novel transplantation strategies.
Johnny L. Boggs; Steven G. McNulty; Linda H. Pardo
2007-01-01
We evaluated foliar and forest floor chemistry across a gradient of N deposition in the Northeast at 11 red spruce (Picea rubens Sarg.) sites in 1987/1988 and foliar and forest floor chemistry and basal area growth at six paired spruce and deciduous sites in 1999. The six red spruce plots were a subset of the original 1987/1988 spruce sites. In 1999...
Edge effects in vertically-oriented graphene based electric double-layer capacitors
NASA Astrophysics Data System (ADS)
Yang, Huachao; Yang, Jinyuan; Bo, Zheng; Zhang, Shuo; Yan, Jianhua; Cen, Kefa
2016-08-01
Vertically-oriented graphenes (VGs) have been demonstrated as a promising active material for electric double-layer capacitors (EDLCs), partially due to their edge-enriched structure. In this work, the 'edge effects', i.e., edges as the promoters of high capacitance, in VG based EDLCs are investigated with experimental research and numerical simulations. VGs with diverse heights (i.e., edge-to-basal ratios) and edge densities are prepared with varying the plasma-enabled growth time and employing different plasma sources. Electrochemical measurements show that the edges play a predominant role on the charge storage behavior of VGs. A simulation is further conducted to unveil the roles of the edges on the separation and adsorption of ions within VG channels. The initial charge distribution of a VG plane is obtained with density functional theory (DFT) calculations, which is subsequently applied to a molecular dynamics (MD) simulation system to gain the insights into the microscope EDLC structures. Compared with the basal planes, the edges present higher initial charge density (by 4.2 times), higher ion packing density (by 2.6 times), closer ion packing location (by 0.8 Å), and larger ion separation degree (by 14%). The as-obtained findings will be instructive in designing the morphology and structure of VGs for enhanced capacitive performances.
Ran, Chao; Huang, Lu; Hu, Jun; Tacon, Philippe; He, Suxu; Li, Zhimin; Wang, Yibing; Liu, Zhi; Xu, Li; Yang, Yalin; Zhou, Zhigang
2016-09-01
In this study, the effects of baker's yeast as probiotics was evaluated in Nile tilapia reared at high density. Juvenile tilapia were distributed to tanks at high density (436 fish/m(3)) and fed with basal diet (CK) or diets supplemented with live (LY) or heat-inactivated yeast (HIY). Another group of fish reared at low density (218 fish/m(3)) and fed with basal diet was also included (LowCK). After 8 weeks of feeding, growth, feed utilization, gut microvilli morphology, digestive enzymes, and expressions of hsp70 and inflammation-related cytokines in the intestine were assessed. Intestinal microbiota was investigated using 16S rRNA gene pyrosequencing. Fish were challenged with Aeromonas hydrophila to evaluate disease resistance. High rearing density significantly decreased the growth, feed utilization, microvilli length, and disease resistance of fish (CK versus LowCK). Moreover, the intestinal hsp70 expression was increased in fish reared at high density, supporting a stress condition. Compared to CK group, supplementation of live yeast significantly increased gut microvilli length and trypsin activity, decreased intestinal hsp70 expression, and enhanced resistance of fish against A. hydrophila (reflected by reduced intestinal alkaline phosphatase activity 24 h post infection). The gut microbiota was not markedly influenced by either rearing density or yeast supplementation. Heat-inactivated yeast (HIY) didn't display the beneficial effects observed in LY except an increase in gut trypsin activity, suggesting the importance of yeast viability and thus secretory metabolites of yeast. In conclusion, live baker's yeast may alleviate the negative effects induced by crowding stress, and has the potential to be used as probiotics for tilapia reared at high density. Copyright © 2016 Elsevier Ltd. All rights reserved.
HINCKLEY, Thomas M.; CHI, Phillip; HAGMANN, Keala; HARRELL, Stevan; SCHMIDT, Amanda Henck; URGENSON, Lauren; ZENG, Zong-yong
2015-01-01
This paper examines human impact on stands and individual trees of Pinus yunnanensis growing near the small mountain villages of Pianshui and Yangjuan in southwestern Sichuan Province, China. In an effort to assess whether use of these forests was sustainable, we examined the effects of human use in two ways. First, we directly measured the effect of cutting branches, for fuel and fodder, on tree growth. We hypothesized that branch cutting would negatively impact tree growth. We established 12 plots on four hills and compared 14 pairs of trees, one tree in each pair with an apparently full crown and the other with a considerable portion of the crown removed. Second, we assessed stand and tree properties over a 500 m elevation gradient above the villages where we hypothesized that as elevation increases, stand and tree properties should show fewer human impacts. Although extensive branch cutting reduced the live crown, tree height and diameter, compensatory processes likely enabled trees to recover and to add basal area increments (BAIs) similar to those added by trees with full crowns. Trees and stands close to villages showed less growth and lower basal areas, respectively, than stands and trees at intermediate or distant elevations from villages. Areas relatively close to the villages showed considerable effects of human-related disturbances such as branch cutting, grazing, tree and shrub removal, losses of litter, and human and animal trails. Such areas had increased soil erosion and often loss of the ‘A’ horizon. Stands close to villages had younger trees, lower stand basal areas, smaller basal area increments, and more stumps. Our results suggest an increasingly vulnerable interface between occupants of these two villages and their surrounding forests. PMID:26478727
Wu, Qian; Pagès, Loïc; Wu, Jie
2016-01-01
Background and Aims Root diameter, especially apical diameter, plays an important role in root development and function. The variation in diameter between roots, and along roots, affects root structure and thus the root system’s overall foraging performance. However, the effect of diameter variation on root elongation, branching and topological connections has not been examined systematically in a population of high-order roots, nor along the roots, especially for mature plants grown in the field. Methods A method combining both excavation and analysis was applied to extract and quantify root architectural traits of adult, field-grown maize plants. The relationships between root diameter and other root architectural characteristics are analysed for two maize cultivars. Key Results The basal diameter of the lateral roots (orders 1–3) was highly variable. Basal diameter was partly determined by the diameter of the bearing segment. Basal diameter defined a potential root length, but the lengths of most roots fell far short of this. This was explained partly by differences in the pattern of diameter change along roots. Diameter tended to decrease along most roots, with the steepness of the gradient of decrease depending on basal diameter. The longest roots were those that maintained (or sometimes increased) their diameters during elongation. The branching density (cm–1) of laterals was also determined by the diameter of the bearing segment. However, the location of this bearing segment along the mother root was also involved – intermediate positions were associated with higher densities of laterals. Conclusions The method used here allows us to obtain very detailed records of the geometry and topology of a complex root system. Basal diameter and the pattern of diameter change along a root were associated with its final length. These relationships are especially useful in simulations of root elongation and branching in source–sink models. PMID:26744490
Fung, Thomas K; Law, Clayton S; Leung, L Stan
2016-06-01
Spike timing-dependent plasticity in the hippocampus has rarely been studied in vivo. Using extracellular potential and current source density analysis in urethane-anesthetized adult rats, we studied synaptic plasticity at the basal dendritic excitatory synapse in CA1 after excitation-spike (ES) pairing; E was a weak basal dendritic excitation evoked by stratum oriens stimulation, and S was a population spike evoked by stratum radiatum apical dendritic excitation. We hypothesize that positive ES pairing-generating synaptic excitation before a spike-results in long-term potentiation (LTP) while negative ES pairing results in long-term depression (LTD). Pairing (50 pairs at 5 Hz) at ES intervals of -10 to 0 ms resulted in significant input-specific LTP of the basal dendritic excitatory sink, lasting 60-120 min. Pairing at +10- to +20-ms ES intervals, or unpaired 5-Hz stimulation, did not induce significant basal dendritic or apical dendritic LTP or LTD. No basal dendritic LTD was found after stimulation of stratum oriens with 200 pairs of high-intensity pulses at 25-ms interval. Pairing-induced LTP was abolished by pretreatment with an N-methyl-d-aspartate receptor antagonist, 3-(2-carboxypiperazin-4-yl)-propyl-1-phosphonic acid (CPP), which also reduced spike bursting during 5-Hz pairing. Pairing at 0.5 Hz did not induce spike bursts or basal dendritic LTP. In conclusion, ES pairing at 5 Hz resulted in input-specific basal dendritic LTP at ES intervals of -10 ms to 0 ms but no LTD at ES intervals of -20 to +20 ms. Associative LTP likely occurred because of theta-rhythmic coincidence of subthreshold excitation with a backpropagated spike burst, which are conditions that can occur naturally in the hippocampus. Copyright © 2016 the American Physiological Society.
Fung, Thomas K.; Law, Clayton S.
2016-01-01
Spike timing-dependent plasticity in the hippocampus has rarely been studied in vivo. Using extracellular potential and current source density analysis in urethane-anesthetized adult rats, we studied synaptic plasticity at the basal dendritic excitatory synapse in CA1 after excitation-spike (ES) pairing; E was a weak basal dendritic excitation evoked by stratum oriens stimulation, and S was a population spike evoked by stratum radiatum apical dendritic excitation. We hypothesize that positive ES pairing—generating synaptic excitation before a spike—results in long-term potentiation (LTP) while negative ES pairing results in long-term depression (LTD). Pairing (50 pairs at 5 Hz) at ES intervals of −10 to 0 ms resulted in significant input-specific LTP of the basal dendritic excitatory sink, lasting 60–120 min. Pairing at +10- to +20-ms ES intervals, or unpaired 5-Hz stimulation, did not induce significant basal dendritic or apical dendritic LTP or LTD. No basal dendritic LTD was found after stimulation of stratum oriens with 200 pairs of high-intensity pulses at 25-ms interval. Pairing-induced LTP was abolished by pretreatment with an N-methyl-d-aspartate receptor antagonist, 3-(2-carboxypiperazin-4-yl)-propyl-1-phosphonic acid (CPP), which also reduced spike bursting during 5-Hz pairing. Pairing at 0.5 Hz did not induce spike bursts or basal dendritic LTP. In conclusion, ES pairing at 5 Hz resulted in input-specific basal dendritic LTP at ES intervals of −10 ms to 0 ms but no LTD at ES intervals of −20 to +20 ms. Associative LTP likely occurred because of theta-rhythmic coincidence of subthreshold excitation with a backpropagated spike burst, which are conditions that can occur naturally in the hippocampus. PMID:27052581
Moisen, Gretchen G.; Freeman, E.A.; Blackard, J.A.; Frescino, T.S.; Zimmermann, N.E.; Edwards, T.C.
2006-01-01
Many efforts are underway to produce broad-scale forest attribute maps by modelling forest class and structure variables collected in forest inventories as functions of satellite-based and biophysical information. Typically, variants of classification and regression trees implemented in Rulequest's?? See5 and Cubist (for binary and continuous responses, respectively) are the tools of choice in many of these applications. These tools are widely used in large remote sensing applications, but are not easily interpretable, do not have ties with survey estimation methods, and use proprietary unpublished algorithms. Consequently, three alternative modelling techniques were compared for mapping presence and basal area of 13 species located in the mountain ranges of Utah, USA. The modelling techniques compared included the widely used See5/Cubist, generalized additive models (GAMs), and stochastic gradient boosting (SGB). Model performance was evaluated using independent test data sets. Evaluation criteria for mapping species presence included specificity, sensitivity, Kappa, and area under the curve (AUC). Evaluation criteria for the continuous basal area variables included correlation and relative mean squared error. For predicting species presence (setting thresholds to maximize Kappa), SGB had higher values for the majority of the species for specificity and Kappa, while GAMs had higher values for the majority of the species for sensitivity. In evaluating resultant AUC values, GAM and/or SGB models had significantly better results than the See5 models where significant differences could be detected between models. For nine out of 13 species, basal area prediction results for all modelling techniques were poor (correlations less than 0.5 and relative mean squared errors greater than 0.8), but SGB provided the most stable predictions in these instances. SGB and Cubist performed equally well for modelling basal area for three species with moderate prediction success, while all three modelling tools produced comparably good predictions (correlation of 0.68 and relative mean squared error of 0.56) for one species. ?? 2006 Elsevier B.V. All rights reserved.
Metastatic Basal cell carcinoma accompanying gorlin syndrome.
Bilir, Yeliz; Gokce, Erkan; Ozturk, Banu; Deresoy, Faik Alev; Yuksekkaya, Ruken; Yaman, Emel
2014-01-01
Gorlin-Goltz syndrome or basal cell nevus syndrome is an autosomal dominant syndrome characterized by skeletal anomalies, numerous cysts observed in the jaw, and multiple basal cell carcinoma of the skin, which may be accompanied by falx cerebri calcification. Basal cell carcinoma is the most commonly skin tumor with slow clinical course and low metastatic potential. Its concomitance with Gorlin syndrome, resulting from a mutation in a tumor suppressor gene, may substantially change morbidity and mortality. A 66-year-old male patient with a history of recurrent basal cell carcinoma was presented with exophthalmus in the left eye and the lesions localized in the left lateral orbita and left zygomatic area. His physical examination revealed hearing loss, gapped teeth, highly arched palate, and frontal prominence. Left orbital mass, cystic masses at frontal and ethmoidal sinuses, and multiple pulmonary nodules were detected at CT scans. Basal cell carcinoma was diagnosed from biopsy of ethmoid sinus. Based on the clinical and typical radiological characteristics (falx cerebri calcification, bifid costa, and odontogenic cysts), the patient was diagnosed with metastatic skin basal cell carcinoma accompanied by Gorlin syndrome. Our case is a basal cell carcinoma with aggressive course accompanying a rarely seen syndrome.
Metastatic Basal Cell Carcinoma Accompanying Gorlin Syndrome
Bilir, Yeliz; Gokce, Erkan; Ozturk, Banu; Deresoy, Faik Alev; Yuksekkaya, Ruken; Yaman, Emel
2014-01-01
Gorlin-Goltz syndrome or basal cell nevus syndrome is an autosomal dominant syndrome characterized by skeletal anomalies, numerous cysts observed in the jaw, and multiple basal cell carcinoma of the skin, which may be accompanied by falx cerebri calcification. Basal cell carcinoma is the most commonly skin tumor with slow clinical course and low metastatic potential. Its concomitance with Gorlin syndrome, resulting from a mutation in a tumor suppressor gene, may substantially change morbidity and mortality. A 66-year-old male patient with a history of recurrent basal cell carcinoma was presented with exophthalmus in the left eye and the lesions localized in the left lateral orbita and left zygomatic area. His physical examination revealed hearing loss, gapped teeth, highly arched palate, and frontal prominence. Left orbital mass, cystic masses at frontal and ethmoidal sinuses, and multiple pulmonary nodules were detected at CT scans. Basal cell carcinoma was diagnosed from biopsy of ethmoid sinus. Based on the clinical and typical radiological characteristics (falx cerebri calcification, bifid costa, and odontogenic cysts), the patient was diagnosed with metastatic skin basal cell carcinoma accompanied by Gorlin syndrome. Our case is a basal cell carcinoma with aggressive course accompanying a rarely seen syndrome. PMID:25506011
Ozaki, Mitsunori; Sano, Hiromi; Sato, Shigeki; Ogura, Mitsuhiro; Mushiake, Hajime; Chiken, Satomi; Nakao, Naoyuki; Nambu, Atsushi
2017-12-01
To understand how information from different cortical areas is integrated and processed through the cortico-basal ganglia pathways, we used optogenetics to systematically stimulate the sensorimotor cortex and examined basal ganglia activity. We utilized Thy1-ChR2-YFP transgenic mice, in which channelrhodopsin 2 is robustly expressed in layer V pyramidal neurons. We applied light spots to the sensorimotor cortex in a grid pattern and examined neuronal responses in the globus pallidus (GP) and entopeduncular nucleus (EPN), which are the relay and output nuclei of the basal ganglia, respectively. Light stimulation typically induced a triphasic response composed of early excitation, inhibition, and late excitation in GP/EPN neurons. Other response patterns lacking 1 or 2 of the components were also observed. The distribution of the cortical sites whose stimulation induced a triphasic response was confined, whereas stimulation of the large surrounding areas induced early and late excitation without inhibition. Our results suggest that cortical inputs to the GP/EPN are organized in a "local inhibitory and global excitatory" manner. Such organization seems to be the neuronal basis for information processing through the cortico-basal ganglia pathways, that is, releasing and terminating necessary information at an appropriate timing, while simultaneously suppressing other unnecessary information. © The Author 2017. Published by Oxford University Press.
NASA Astrophysics Data System (ADS)
Gong, Yongmei; Zwinger, Thomas; Åström, Jan; Gladstone, Rupert; Schellenberger, Thomas; Altena, Bas; Moore, John
2017-04-01
The outlet glacier at Basin 3, Austfonna ice-cap entered its active surge phase in autumn 2012. We assess the evolution of the basal friction during the surge through inverse modelling of basal friction coefficients using recent velocity observation from 2012 to 2014 in a continuum ice dynamic model Elmer/ice. The obtained basal friction coefficient distributions at different time instances are further used as a boundary condition in a discrete element model (HiDEM) that is capable of computing fracturing of ice. The inverted basal friction coefficient evolution shows a gradual 'unplugging' of the stagnant frontal area and northwards and inland expansion of the fast flowing region in the southern basin. The validation between the modeled crevasses distribution and the satellite observation in August 2013 shows a good agreement in shear zones inland and at the frontal area. Crevasse distributions of the summer before and after the glacier reached its maximum velocity in January 2013 (August 2012 and August 2014, respectively) are also evaluated. Previous studies suggest the triggering and development of the surge are linked to surface melt water penetrating through ice to form an efficient basal hydrology system thereby triggering a hydro- thermodynamic feedback. This preliminary offline coupling between a continuum ice dynamic model and a discrete element model will give a hint on future model development of linking supra-glacial to sub-glacial hydrology system.
Duinkerken, Charlotte W; Lohuis, Peter J F M; Crijns, Marianne B; Navran, Arash; Haas, Rick L M; Hamming-Vrieze, Olga; Klop, W Martin C; van den Brekel, Michiel W M; Al-Mamgani, Abrahim
Surgery is the golden standard for treating basal cell carcinomas. In case of positive tumor margins or recurrent disease, postoperative adjuvant or salvaging therapy is suggested to achieve good local control. To retrospectively report on local control and toxicity of postoperative radiotherapy by means of orthovoltage X-rays for residual or recurrent basal cell carcinoma after surgery in the head and neck area. Sixty-six surgically resected residual or recurrent basal cell carcinomas of the head and neck region were irradiated postoperatively by means of orthovoltage X-rays at the Netherlands Cancer Institute between January 2000 and February 2015. After a median follow-up duration of 30.5 months, only 5 recurrences were reported. The 5-year local control rates at 1, 3, and 5 years were 100%, 87%, and 87%, respectively. The 5-year local control rate was 92% for immediate postoperative radiotherapy of incompletely resected basal cell carcinomas, 90% for recurrences after 1 previously performed excision, and 71% for multiple recurrences, namely, a history of more than 1 excision ( P = .437). Acute toxicity healed spontaneously within 3 months. Late toxicities were mild. Radiotherapy by means of orthovoltage X-ray is an excellent alternative for re-excision in case of incompletely resected or recurrent basal cell carcinomas that are at risk of serious functional and cosmetic impairments after re-excision, with a 5-year local control rate of 87% and a low toxicity profile.
2016-01-01
Basal ganglia circuits are critical for the modulation of motor performance across behavioral states. In zebra finches, a cortical-basal ganglia circuit dedicated to singing is necessary for males to adjust their song performance and transition between spontaneous singing, when they are alone (“undirected” song), and a performance state, when they sing to a female (“female-directed” song). However, we know little about the role of different basal ganglia cell types in this behavioral transition or the degree to which behavioral context modulates the activity of different neuron classes. To investigate whether interneurons in the songbird basal ganglia encode information about behavioral state, I recorded from two interneuron types, fast-spiking interneurons (FSI) and external pallidal (GPe) neurons, in the songbird basal ganglia nucleus area X during both female-directed and undirected singing. Both cell types exhibited higher firing rates, more frequent bursting, and greater trial-by-trial variability in firing when male zebra finches produced undirected songs compared with when they produced female-directed songs. However, the magnitude and direction of changes to the firing rate, bursting, and variability of spiking between when birds sat silently and when they sang undirected and female-directed song varied between FSI and GPe neurons. These data indicate that social modulation of activity important for eliciting changes in behavioral state is present in multiple cell types within area X and suggests that social interactions may adjust circuit dynamics during singing at multiple points within the circuit. PMID:27628208
Low thinning and crown thinning of two severities as restoration tools at Redwood National Park
Teraoka, Jason R; van Mantgem, Phillip J.; Keyes, Christopher R.
2017-01-01
Interest in the restoration of second-growth forests has continued to increase in the redwood region, which has further increased the importance of evaluating restoration-based silvicultural strategies. This study assessed the short-term effectiveness of four silvicultural treatments (two silvicultural thinning methods, low thinning and crown thinning, and two basal area retentions, 80 percent and 45 percent) as forest restoration tools via analysis of relative basal area growth at Redwood National Park. Prior to treatment, the second-growth stand had more than 1,600 trees ha-1 and 70.0 m2 ha-1 basal area and consisted primarily of two species, Douglas-fir (Pseudotsuga menziesii(Mirb.) Franco) (the dominant species) and redwood (Sequoia sempervirens (D. Don) Endl.). Growth was enhanced for all treatments with 5-year net basal area gains of 28.4 percent for the lowretention crown thinning, 28.1 percent for the low-retention low thinning, 23.3 percent for the high-retention crown thinning, 19.1 percent for high-retention low thinning, and only 14.2 percent for the control. We conclude that all four thinning treatments improved tree growth; but among them, the low-retention treatments were most effective in accomplishing restoration objectives, while the high-retention low thinning was least effective. Increasing the array of silvicultural tools that Redwood National Park can use may prove helpful in accomplishing restoration goals in future projects.
Brain correlates of constituent structure in sign language comprehension.
Moreno, Antonio; Limousin, Fanny; Dehaene, Stanislas; Pallier, Christophe
2018-02-15
During sentence processing, areas of the left superior temporal sulcus, inferior frontal gyrus and left basal ganglia exhibit a systematic increase in brain activity as a function of constituent size, suggesting their involvement in the computation of syntactic and semantic structures. Here, we asked whether these areas play a universal role in language and therefore contribute to the processing of non-spoken sign language. Congenitally deaf adults who acquired French sign language as a first language and written French as a second language were scanned while watching sequences of signs in which the size of syntactic constituents was manipulated. An effect of constituent size was found in the basal ganglia, including the head of the caudate and the putamen. A smaller effect was also detected in temporal and frontal regions previously shown to be sensitive to constituent size in written language in hearing French subjects (Pallier et al., 2011). When the deaf participants read sentences versus word lists, the same network of language areas was observed. While reading and sign language processing yielded identical effects of linguistic structure in the basal ganglia, the effect of structure was stronger in all cortical language areas for written language relative to sign language. Furthermore, cortical activity was partially modulated by age of acquisition and reading proficiency. Our results stress the important role of the basal ganglia, within the language network, in the representation of the constituent structure of language, regardless of the input modality. Copyright © 2017 Elsevier Inc. All rights reserved.
Busing, Richard T.; Solomon, Allen M.
2004-01-01
Two forest dynamics simulators are compared along climatic gradients in the Pacific Northwest. The ZELIG and FORCLIM models are tested against forest survey data from western Oregon. Their ability to generate accurate patterns of forest basal area and species composition is evaluated for series of sites with contrasting climate. Projections from both models approximate the basal area and composition patterns for three sites along the elevation gradient at H.J. Andrews Experimental Forest in the western Cascade Range. The ZELIG model is somewhat more accurate than FORCLIM at the two low-elevation sites. Attempts to project forest composition along broader climatic gradients reveal limitations of ZELIG, however. For example, ZELIG is less accurate than FORCLIM at projecting the average composition of a west Cascades ecoregion selected for intensive analysis. Also, along a gradient consisting of several sites on an east to west transect at 44.1oN latitude, both the FORCLIM model and the actual data show strong changes in composition and total basal area, but the ZELIG model shows a limited response. ZELIG does not simulate the declines in forest basal area and the diminished dominance of mesic coniferous species east of the Cascade crest. We conclude that ZELIG is suitable for analyses of certain sites for which it has been calibrated. FORCLIM can be applied in analyses involving a range of climatic conditions without requiring calibration for specific sites.
Pompa-García, Marín; Venegas-González, Alejandro
2016-01-01
Forest ecosystems play an important role in the global carbon cycle. Therefore, understanding the dynamics of carbon uptake in forest ecosystems is much needed. Pinus cooperi is a widely distributed species in the Sierra Madre Occidental in northern Mexico and future climatic variations could impact these ecosystems. Here, we analyze the variations of trunk carbon in two populations of P. cooperi situated at different elevational gradients, combining dendrochronological techniques and allometry. Carbon sequestration (50% biomass) was estimated from a specific allometric equation for this species based on: (i) variation of intra-annual wood density and (ii) diameter reconstruction. The results show that the population at a higher elevation had greater wood density, basal area, and hence, carbon accumulation. This finding can be explained by an ecological response of trees to adverse weather conditions, which would cause a change in the cellular structure affecting the within-ring wood density profile. The influence of variations in climate on the maximum density of chronologies showed a positive correlation with precipitation and the Multivariate El Niño Southern Oscillation Index during the winter season, and a negative correlation with maximum temperature during the spring season. Monitoring previous conditions to growth is crucial due to the increased vulnerability to extreme climatic variations on higher elevational sites. We concluded that temporal variability of wood density contributes to a better understanding of environmental historical changes and forest carbon dynamics in Northern Mexico, representing a significant improvement over previous studies on carbon sequestration. Assuming a uniform density according to tree age is incorrect, so this method can be used for environmental mitigation strategies, such as for managing P. cooperi, a dominant species of great ecological amplitude and widely used in forest industries. PMID:27272519
Pompa-García, Marín; Venegas-González, Alejandro
2016-01-01
Forest ecosystems play an important role in the global carbon cycle. Therefore, understanding the dynamics of carbon uptake in forest ecosystems is much needed. Pinus cooperi is a widely distributed species in the Sierra Madre Occidental in northern Mexico and future climatic variations could impact these ecosystems. Here, we analyze the variations of trunk carbon in two populations of P. cooperi situated at different elevational gradients, combining dendrochronological techniques and allometry. Carbon sequestration (50% biomass) was estimated from a specific allometric equation for this species based on: (i) variation of intra-annual wood density and (ii) diameter reconstruction. The results show that the population at a higher elevation had greater wood density, basal area, and hence, carbon accumulation. This finding can be explained by an ecological response of trees to adverse weather conditions, which would cause a change in the cellular structure affecting the within-ring wood density profile. The influence of variations in climate on the maximum density of chronologies showed a positive correlation with precipitation and the Multivariate El Niño Southern Oscillation Index during the winter season, and a negative correlation with maximum temperature during the spring season. Monitoring previous conditions to growth is crucial due to the increased vulnerability to extreme climatic variations on higher elevational sites. We concluded that temporal variability of wood density contributes to a better understanding of environmental historical changes and forest carbon dynamics in Northern Mexico, representing a significant improvement over previous studies on carbon sequestration. Assuming a uniform density according to tree age is incorrect, so this method can be used for environmental mitigation strategies, such as for managing P. cooperi, a dominant species of great ecological amplitude and widely used in forest industries.
Inundation and Fire Shape the Structure of Riparian Forests in the Pantanal, Brazil
Arruda, Wellinton de Sá; Oldeland, Jens; Paranhos Filho, Antonio Conceição; Pott, Arnildo; Cunha, Nicolay L.; Ishii, Iria Hiromi; Damasceno-Junior, Geraldo Alves
2016-01-01
Inundation and fire can affect the structure of riparian vegetation in wetlands. Our aim was to verify if there are differences in richness, abundance, basal area, composition and topographic preference of woody species in riparian forests related to the fire history, flooding duration, or the interaction between both. The study was conducted in the riparian forests of the Paraguay River some of which were burned three times between 2001 and 2011. We sampled trees with a girth of at least 5 cm at breast height in 150 5 × 10 m plots (79 burned and 71 unburned). We also measured height of the flood mark and estimated the flooding duration of each plot. We performed Generalized Linear Mixed Models to verify differences in richness, basal area, and abundance of individuals associated to interaction of fire and inundation. We used an analysis of similarity (ANOSIM) and indicator species analysis to identify differences in composition of species and the association with burned and unburned area according to different levels of inundation. Finally, we used a hierarchical set of Generalized Linear Models (GLM), the so-called HOF models, to analyse each species’ specific response to inundation based on topography and to determine their preferred optimal topographic position for both burned as well as unburned areas. Richness was positively associated with elevation only in burned areas while abundance was negatively influenced by inundation only in burned areas. Basal area was negatively associated with time of inundation independent of fire history. There were 15 species which were significant indicators for at least one combination of the studied factors. We found nine species in burned areas and 15 in unburned areas, with response curves in HOF models along the inundation gradient. From these, five species shifted their optimal position along the inundation gradient in burned areas. The interaction of fire and inundation did not appear to affect the basal area, but it did affect the richness, number of individuals, success of some species, and seemed to shape the boundary of these forests as shown by the difference in the positioning of these species along the inundation gradient. PMID:27280879
Inundation and Fire Shape the Structure of Riparian Forests in the Pantanal, Brazil.
Arruda, Wellinton de Sá; Oldeland, Jens; Paranhos Filho, Antonio Conceição; Pott, Arnildo; Cunha, Nicolay L; Ishii, Iria Hiromi; Damasceno-Junior, Geraldo Alves
2016-01-01
Inundation and fire can affect the structure of riparian vegetation in wetlands. Our aim was to verify if there are differences in richness, abundance, basal area, composition and topographic preference of woody species in riparian forests related to the fire history, flooding duration, or the interaction between both. The study was conducted in the riparian forests of the Paraguay River some of which were burned three times between 2001 and 2011. We sampled trees with a girth of at least 5 cm at breast height in 150 5 × 10 m plots (79 burned and 71 unburned). We also measured height of the flood mark and estimated the flooding duration of each plot. We performed Generalized Linear Mixed Models to verify differences in richness, basal area, and abundance of individuals associated to interaction of fire and inundation. We used an analysis of similarity (ANOSIM) and indicator species analysis to identify differences in composition of species and the association with burned and unburned area according to different levels of inundation. Finally, we used a hierarchical set of Generalized Linear Models (GLM), the so-called HOF models, to analyse each species' specific response to inundation based on topography and to determine their preferred optimal topographic position for both burned as well as unburned areas. Richness was positively associated with elevation only in burned areas while abundance was negatively influenced by inundation only in burned areas. Basal area was negatively associated with time of inundation independent of fire history. There were 15 species which were significant indicators for at least one combination of the studied factors. We found nine species in burned areas and 15 in unburned areas, with response curves in HOF models along the inundation gradient. From these, five species shifted their optimal position along the inundation gradient in burned areas. The interaction of fire and inundation did not appear to affect the basal area, but it did affect the richness, number of individuals, success of some species, and seemed to shape the boundary of these forests as shown by the difference in the positioning of these species along the inundation gradient.
Basal Ganglia Circuits as Targets for Neuromodulation in Parkinson Disease.
DeLong, Mahlon R; Wichmann, Thomas
2015-11-01
The revival of stereotactic surgery for Parkinson disease (PD) in the 1990s, with pallidotomy and then with high-frequency deep brain stimulation (DBS), has led to a renaissance in functional surgery for movement and other neuropsychiatric disorders. To examine the scientific foundations and rationale for the use of ablation and DBS for treatment of neurologic and psychiatric diseases, using PD as the primary example. A summary of the large body of relevant literature is presented on anatomy, physiology, pathophysiology, and functional surgery for PD and other basal ganglia disorders. The signs and symptoms of movement disorders appear to result largely from signature abnormalities in one of several parallel and largely segregated basal ganglia thalamocortical circuits (ie, the motor circuit). The available evidence suggests that the varied movement disorders resulting from dysfunction of this circuit result from propagated disruption of downstream network activity in the thalamus, cortex, and brainstem. Ablation and DBS act to free downstream networks to function more normally. The basal ganglia thalamocortical circuit may play a key role in the expression of disordered movement, and the basal ganglia-brainstem projections may play roles in akinesia and disturbances of gait. Efforts are under way to target circuit dysfunction in brain areas outside of the traditionally implicated basal ganglia thalamocortical system, in particular, the pedunculopontine nucleus, to address gait disorders that respond poorly to levodopa and conventional DBS targets. Deep brain stimulation is now the treatment of choice for many patients with advanced PD and other movement disorders. The success of DBS and other forms of neuromodulation for neuropsychiatric disorders is the result of the ability to modulate circuit activity in discrete functional domains within the basal ganglia circuitry with highly focused interventions, which spare uninvolved areas that are often disrupted with drugs.
Forest reproduction along a climatic gradient in the Sierra Nevada, California
van Mantgem, Phillip J.; Stephenson, Nathan L.; Keeley, Jon E.
2006-01-01
To elucidate broad-scale environmental controls of coniferous forest reproduction in the Sierra Nevada, California, we monitored reproduction for 5 years in 47 plots arrayed across a steep elevational (climatic) gradient. We found that both absolute seedling densities (stems < 1.37 m) and seedling densities relative to overstory parent tree basal area declined sharply with elevation. Rates of seedling turnover (the average of birth and death rates) also declined with elevation. In contrast, seed production was not predicted by elevation and was highly variable from year to year. During a mast year of seed production, the intensity of masting was uneven among plots. Seedling densities were elevated only during the single year immediately following the mast year, suggesting reproduction in our forests may be primarily limited by abiotic factors such as the availability of suitable sites and weather. Disturbance also clearly affected reproduction; plots that had recently burned had significantly higher seedling to parent tree ratios for Abies species, suggesting that even though established Abies concolor may be relatively susceptible to fire, the species can recover rapidly through prolific reproduction. Since reproductive failures may be our earliest signal of changing forest conditions, seedling dynamics could provide a sensitive, if variable, indicator of environmental changes.
Basal-like Breast Cancers: From Pathology to Biology and Back Again.
Gusterson, Barry; Eaves, Connie J
2018-06-05
Human breast cancers referred to as "basal-like" are of interest because they lack effective therapies and their biology is poorly understood. The term basal-like derives from studies demonstrating tumor gene expression profiles that include some transcripts characteristic of the basal cells of the normal adult human mammary gland and others associated with a subset of normal luminal cells. Elucidating the mechanisms responsible for the profiles of basal-like tumors is an active area of investigation. More refined molecular analysis of patients' samples and genetic strategies to produce breast cancers de novo from defined populations of normal mouse mammary cells have served as complementary approaches to identify relevant pathway alterations. However, both also have limitations. Here, we review some of the underlying reasons, including the unifying concept that some normal luminal cells have both luminal and basal features, as well as some emerging new avenues of investigation. Copyright © 2018 The Author(s). Published by Elsevier Inc. All rights reserved.
[Soil and forest structure in the Colombian Amazon].
Calle-Rendón, Bayron R; Moreno, Flavio; Cárdenas López, Dairon
2011-09-01
Forests structural differences could result of environmental variations at different scales. Because soils are an important component of plant's environment, it is possible that edaphic and structural variables are associated and that, in consequence, spatial autocorrelation occurs. This paper aims to answer two questions: (1) are structural and edaphic variables associated at local scale in a terra firme forest of Colombian Amazonia? and (2) are these variables regionalized at the scale of work? To answer these questions we analyzed the data of a 6ha plot established in a terra firme forest of the Amacayacu National Park. Structural variables included basal area and density of large trees (diameter > or = 10cm) (Gdos and Ndos), basal area and density of understory individuals (diameter < 10cm) (Gsot and Nsot) and number of species of large trees (sp). Edaphic variables included were pH, organic matter, P, Mg, Ca, K, Al, sand, silt and clay. Structural and edaphic variables were reduced through a principal component analysis (PCA); then, the association between edaphic and structural components from PCA was evaluated by multiple regressions. The existence of regionalization of these variables was studied through isotropic variograms, and autocorrelated variables were spatially mapped. PCA found two significant components for structure, corresponding to the structure of large trees (G, Gdos, Ndos and sp) and of small trees (N, Nsot and Gsot), which explained 43.9% and 36.2% of total variance, respectively. Four components were identified for edaphic variables, which globally explained 81.9% of total variance and basically represent drainage and soil fertility. Regression analyses were significant (p < 0.05) and showed that the structure of both large and small trees is associated with greater sand contents and low soil fertility, though they explained a low proportion of total variability (R2 was 4.9% and 16.5% for the structure of large trees and small tress, respectively). Variables with spatial autocorrelation were the structure of small trees, Al, silt, and sand. Among them, Nsot and sand content showed similar patterns of spatial distribution inside the plot.
West, Daniel R.; Briggs, Jennifer S.; Jacobi, William R.; Negrón, José F.
2014-01-01
Eruptive mountain pine beetle (Dendroctonus ponderosae, MPB) populations have caused widespread mortality of pines throughout western North America since the late 1990s. Early work by A.D. Hopkins suggested that when alternate host species are available, MPB will prefer to breed in the host to which it has become adapted. In Colorado, epidemic MPB populations that originated in lodgepole pine expanded into mixed-conifer stands containing ponderosa pine, a related host. We evaluated the susceptibility of both hosts to successful MPB colonization in a survey of 19 sites in pine-dominated mixed-conifer stands spanning 140 km of the Front Range, CO, USA. In each of three 0.2-ha plots at each site, we (1) assessed trees in the annual flights of 2008–2011 to compare MPB-caused mortality between lodgepole and ponderosa pine; (2) recorded previous MPB-caused tree mortality from 2004–2007 to establish baseline mortality levels; and (3) measured characteristics of the stands (e.g. tree basal area) and sites (e.g. elevation, aspect) that might be correlated with MPB colonization. Uninfested average live basal area of lodgepole and ponderosa pine was 74% of total basal area before 2004. We found that for both species, annual percent basal area of attacked trees was greatest in one year (2009), and was lower in all other years (2004–2007, 2008, 2010, and 2011). Both pine species had similar average total mortality of 38–39% by 2011. Significant predictors of ponderosa pine mortality in a given year were basal area of uninfested ponderosa pine and the previous year’s mortality levels in both ponderosa and lodgepole pine. Lodgepole pine mortality was predicted by uninfested basal areas of both lodgepole and ponderosa pine, and the previous year’s lodgepole pine mortality. These results indicate host selection by MPB from lodgepole pine natal hosts into ponderosa pine the following year, but not the reverse. In both species, diameters of attacked trees within each year were similar, and were progressively smaller the last four years of the study period. Our results suggest that, in contrast to previous reports, ponderosa and lodgepole pine were equally susceptible to MPB infestation in the CO Front Range during our study period. This suggests that forest managers may anticipate similar impacts in both hosts during similar environmental conditions when epidemic-level MPB populations are active in mixed-pine stands.
NASA Astrophysics Data System (ADS)
Navelot, Vivien; Favier, Alexiane; Géraud, Yves; Diraison, Marc; Corsini, Michel; Verati, Chrystèle; Lardeaux, Jean-Marc; Mercier de Lépinay, Jeanne; Munschy, Marc
2017-04-01
The GEOTREF project (high enthalpy geothermal energy in fractured reservoirs), supported by the French government program, "Investissements d'avenir" develops a sustainable geothermal resource in the Vieux Habitants area, 8-km south of the currently exploited Bouillante geothermal field. The Basse Terre Island is a recent volcanic arc (< 3 Myr) belonging to the Lesser Antilles subduction zone. It is composed of arc typical calc-alkaline volcanic rocks. Outcrops of the studied area consist either of andesitic lava flows, volcanic sedimentary facies or dikes. Field studies allow to propose a structural framework and highlight three major directions N000˚ E, N050˚ E and N090˚ E, which are consistent with the regional tectonic trends of the arc. Petrographical and petrophysical studies displayed that the major part of outcropping facies in the Vieux-Habitants area are not altered. Andesitic lava flows have poor reservoir properties with porosity and permeability lower than 5 % and 10-15m2 respectively. These results are in contrast with measurements performed in volcano-sedimentary rocks, which have heterogeneous petrophysical properties ranging from 15 to 50 % for porosity and from 10-15to 10-9m2 for permeability. Such surface data would probably change and decrease when depth increases. As there is a lack of underground data under the Vieux-Habitants area (wireline, drill core), exhumated rocks outcropping in the northern part of Basse-Terre Island (Basal Complex) have been studied. Such rocks have been identified in the Basal Complex (2.5 - 3 Myr) located in the northern part of the Basse-Terre Island. Previous works have demonstrated a 1000 m/Myr erosional rate, which corresponds at least to a 2 - 3 km exhumation. The petrography study of the Basal Complex reveals sub-greenschist type mineralogical transformations (chlorite, white mica, quartz...) changing the andesitic protolith in a meta-andesite. This metamorphism forms cleavage plans thanks to a pressure-solution mechanism. Mineralogical transformations associated with these cleavage planes have an impact on petrophysical properties. The solid phase density and porosity decrease. An anisotropy of permeability develops due to cleavage plans. Thermodynamics modelling based on the rock chemical composition and petrography observations emphasizes a steady-state mineral assemblage between 1.5 - 2 kbar and 280 - 320˚ C. This is consistent with an in situ measured volcanic arc conductive geothermal gradient of 70 ˚ C/km.
García-Cordero, Indira; Esteves, Sol; Mikulan, Ezequiel P.; Hesse, Eugenia; Baglivo, Fabricio H.; Silva, Walter; García, María del Carmen; Vaucheret, Esteban; Ciraolo, Carlos; García, Hernando S.; Adolfi, Federico; Pietto, Marcos; Herrera, Eduar; Legaz, Agustina; Manes, Facundo; García, Adolfo M.; Sigman, Mariano; Bekinschtein, Tristán A.; Ibáñez, Agustín; Sedeño, Lucas
2017-01-01
Interoception, the monitoring of visceral signals, is often presumed to engage attentional mechanisms specifically devoted to inner bodily sensing. In fact, most standardized interoceptive tasks require directing attention to internal signals. However, most studies in the field have failed to compare attentional modulations between internally- and externally-driven processes, thus probing blind to the specificity of the former. Here we address this issue through a multidimensional approach combining behavioral measures, analyses of event-related potentials and functional connectivity via high-density electroencephalography, and intracranial recordings. In Study 1, 50 healthy volunteers performed a heartbeat detection task as we recorded modulations of the heartbeat-evoked potential (HEP) in three conditions: exteroception, basal interoception (also termed interoceptive accuracy), and post-feedback interoception (sometimes called interoceptive learning). In Study 2, to evaluate whether key interoceptive areas (posterior insula, inferior frontal gyrus, amygdala, and somatosensory cortex) were differentially modulated by externally- and internally-driven processes, we analyzed human intracranial recordings with depth electrodes in these regions. This unique technique provides a very fine grained spatio-temporal resolution compared to other techniques, such as EEG or fMRI. We found that both interoceptive conditions in Study 1 yielded greater HEP amplitudes than the exteroceptive one. In addition, connectivity analysis showed that post-feedback interoception, relative to basal interoception, involved enhanced long-distance connections linking frontal and posterior regions. Moreover, results from Study 2 showed a differentiation between oscillations during basal interoception (broadband: 35–110 Hz) and exteroception (1–35 Hz) in the insula, the amygdala, the somatosensory cortex, and the inferior frontal gyrus. In sum, this work provides convergent evidence for the specificity and dynamics of attentional mechanisms involved in interoception. PMID:28769749
García-Cordero, Indira; Esteves, Sol; Mikulan, Ezequiel P; Hesse, Eugenia; Baglivo, Fabricio H; Silva, Walter; García, María Del Carmen; Vaucheret, Esteban; Ciraolo, Carlos; García, Hernando S; Adolfi, Federico; Pietto, Marcos; Herrera, Eduar; Legaz, Agustina; Manes, Facundo; García, Adolfo M; Sigman, Mariano; Bekinschtein, Tristán A; Ibáñez, Agustín; Sedeño, Lucas
2017-01-01
Interoception, the monitoring of visceral signals, is often presumed to engage attentional mechanisms specifically devoted to inner bodily sensing. In fact, most standardized interoceptive tasks require directing attention to internal signals. However, most studies in the field have failed to compare attentional modulations between internally- and externally-driven processes, thus probing blind to the specificity of the former. Here we address this issue through a multidimensional approach combining behavioral measures, analyses of event-related potentials and functional connectivity via high-density electroencephalography, and intracranial recordings. In Study 1, 50 healthy volunteers performed a heartbeat detection task as we recorded modulations of the heartbeat-evoked potential (HEP) in three conditions: exteroception, basal interoception (also termed interoceptive accuracy), and post-feedback interoception (sometimes called interoceptive learning). In Study 2, to evaluate whether key interoceptive areas (posterior insula, inferior frontal gyrus, amygdala, and somatosensory cortex) were differentially modulated by externally- and internally-driven processes, we analyzed human intracranial recordings with depth electrodes in these regions. This unique technique provides a very fine grained spatio-temporal resolution compared to other techniques, such as EEG or fMRI. We found that both interoceptive conditions in Study 1 yielded greater HEP amplitudes than the exteroceptive one. In addition, connectivity analysis showed that post-feedback interoception, relative to basal interoception, involved enhanced long-distance connections linking frontal and posterior regions. Moreover, results from Study 2 showed a differentiation between oscillations during basal interoception (broadband: 35-110 Hz) and exteroception (1-35 Hz) in the insula, the amygdala, the somatosensory cortex, and the inferior frontal gyrus. In sum, this work provides convergent evidence for the specificity and dynamics of attentional mechanisms involved in interoception.
Glaciation and regional groundwater flow in the Fennoscandian shield
Provost, A.M.; Voss, C.I.; Neuzil, C.E.
2012-01-01
Regional-scale groundwater flow modeling of the Fennoscandian shield suggests that groundwater flow can be strongly affected by future climate change and glaciation. We considered variable-density groundwater flow in a 1500-km-long and approximately 10-km-deep cross-section through southern Sweden. Groundwater flow and shield brine transport in the cross-sectional model were analyzed under projected surface conditions for the next 140 ka. Simulations suggest that blockage of recharge and discharge by low-permeability permafrost or cold-based ice causes sinking of brine and consequent freshening of near-surface water in areas of natural discharge. Although recharge of basal meltwater is limited by the requirement that water pressure at the base of the ice sheet not exceed the pressure exerted by the weight of the ice, warm-based ice with basal melting creates a potential for groundwater recharge rates much larger than those of present, ice-free conditions. In the simulations, regional-scale redistribution of recharged water by subsurface flow is minor over the duration of a glacial advance (approximately 10 ka). During glacial retreat, significant upward flow of groundwater may occur below the ice sheet owing to pressure release. If the mechanical loading efficiency of the rocks is high, both subsurface penetration of meltwater during glacial advance and up-flow during glacial retreat are reduced because of loading-induced pressure changes. The maximum rate of groundwater discharge in the simulations occurs at the receding ice margin, and some discharge occurs below incursive postglacial seas. Recharge of basal meltwater could decrease the concentration of dissolved solids significantly below present-day levels at depths of up to several kilometers and may bring oxygenated conditions to an otherwise reducing chemical environment for periods exceeding 10 ka.
NASA Astrophysics Data System (ADS)
McCracken, R. G.; Iverson, N. R.; Benediktsson, Í. Ö.; Schomacker, A.; Johnson, M. D.; Zoet, L.; Hooyer, T.
2015-12-01
The forefield of Múlajökull—a warm-based, surge-type glacier in central Iceland—comprises the only known active drumlin field, with drumlins of similar morphometry to their Pleistocene counterparts but formed under better-known glaciological conditions. Study of till magnetic and till fabrics, till densities and preconsolidation stresses, and drumlin stratigraphy indicates that drumlin relief reflects both erosion and deposition: 1) the package of basal tills that constitutes the drumlins thickens where the forefield has experienced more surging, but uniformities are common on drumlin flanks, 2) attitudes of till layers and patterns of deformation within them indicate till deposition occurred on drumlin slopes rather than prior to drumlinization, and 3) past effective stresses during quiescent periods were highest in interdrumlin areas. These data suggest that erosion occurred during quiescence, rather than during surging, with erosion rates that increased under increasing effective stress. Stratigraphic evidence connecting specific till layers to surge moraines (Johnson et al., 2010, Geology 38, 943-6), on the other hand, indicates that deposition occurred during surges. Such deposition could have resulted from negative flux divergence in a shearing bed, but till fabrics provide no evidence for longitudinally compressive strain. A more likely origin for the basal till is that rapid, uniform bed shear during surging—consistent with the low and relatively uniform effective stresses expected during such periods—generated basal melt rates sufficient to release debris from ice and lodge it onto the bed. This conceptual model of alternating erosion and deposition implies drumlins formed by slow flowing ice elsewhere will be dominantly erosional, whereas drumlins with evidence of deposition may have experienced fast ice flow.
Ramírez, Manuel; Martínez-Martínez, Laura-Aline; Hernández-Quintela, Everardo; Velazco-Casapía, Jorge; Vargas, Angélica; Martínez-Lavín, Manuel
2015-10-01
A consistent line of investigation suggests that fibromyalgia is a neuropathic pain syndrome. This outlook has been recently reinforced by several controlled studies that describe decreased small nerve fiber density in skin biopsies of patients with fibromyalgia. The cornea receives the densest small fiber innervation of the body. Corneal confocal bio-microscopy is a new noninvasive method to evaluate small nerve fiber morphology. Our objective was to assess corneal small nerve fiber morphology in patients with fibromyalgia, and to associate corneal nerve microscopic features with neuropathic pain descriptors and other fibromyalgia symptoms. We studied 17 female patients with fibromyalgia and 17 age-matched healthy control subjects. All the participants completed different questionnaires regarding the symptoms of fibromyalgia, including a neuropathic pain survey. A central corneal thickness scan was obtained with a confocal microscope. Nerve measurements were made by a single ophthalmologist without knowledge of the clinical diagnosis. Stromal nerve thickness was defined as the mean value between the widest and the narrowest portion of each analyzed stromal nerve. Corneal sub-basal plexus nerve density was also assessed. Patients with fibromyalgia had stromal nerve thickness of 5.0 ± 1.0 µm (mean ± standard deviation) significantly different from that of control's values (6.1 ± 1.3) p = 0.01. Patients also had decreased sub-basal plexus nerve density per square millimeter (85 ± 29) vs. 107 ± 26 of controls p = 0.02. When controls and patients were grouped together, there was an association between stromal nerve slenderness and neuropathic pain descriptors (Fisher's exact test p = 0.007). Women suffering from fibromyalgia have thinner corneal stromal nerves and diminished sub-basal plexus nerve density when compared to healthy controls. Nerve scarcity is associated with neuropathic pain descriptors. Small fiber neuropathy may play a role in the pathogenesis of fibromyalgia pain. Corneal confocal microscopy could become a useful test in the study of patients with fibromyalgia. Copyright © 2015. Published by Elsevier Inc.
Barcelos, Amanda; Tibirica, Eduardo; Lamas, Cristiane
2018-07-01
To evaluate the systemic microcirculation of patients with infective endocarditis (IE). This is a comparative study of patients with definite IE by the modified Duke criteria admitted to our center for treatment. A reference group of sex- and age-matched healthy volunteers was included. Microvascular flow was evaluated in the forearm using a laser speckle contrast imaging system, for noninvasive measurement of cutaneous microvascular perfusion, in combination with skin iontophoresis of acetylcholine (ACh) and sodium nitroprusside (SNP) to test microvascular reactivity. Microvascular density was evaluated using skin video-capillaroscopy. We studied 22 patients with IE; 15 were male and seven female. The mean age and standard deviation (SD) were 45.5 ± 17.3 years. Basal skin microvascular conductance was significantly increased in patients with IE, compared with healthy individuals (0.36 ± 0.13 versus 0.21 ± 0.08 APU/mmHg; P < 0.0001). The increase in microvascular conductance induced by ACh in patients was 0.21 ± 0.17 and in the reference group, it was 0.37 ± 0.14 APU/mmHg (P = 0.0012). The increase in microvascular conductance induced by SNP in patients was 0.18 ± 0.14 and it was 0.29 ± 0.15 APU/mmHg (P = 0.0140) in the reference group. The basal mean skin capillary density of patients (135 ± 24 capillaries/mm 2 ) was significantly higher, compared with controls (97 ± 21 capillaries/mm 2 ; P < 0.0001). The main findings in the microcirculation of patients with IE were greater basal vasodilation and a reduction of the endothelium-dependent and -independent microvascular reactivity, as well as greater functional skin capillary density compared to healthy individuals. Copyright © 2018 Elsevier Inc. All rights reserved.
The deformation mechanisms and size effects of single-crystal magnesium
NASA Astrophysics Data System (ADS)
Byer, Cynthia M.
In this work, we seek to understand the deformation mechanisms and size effects of single-crystal magnesium at the micrometer scale through both microcompression experiments and finite element simulations. Microcompression experiments are conducted to investigate the impact of initial dislocation density and orientation on size effects. Micropillars are fabricated using a focused ion beam and tested in a Nanoindenter using a diamond fiat tip as a compression platen. Two different initial dislocation densities are examined for [0001] oriented micropillars. Our results demonstrate that decreasing the initial dislocation density results in an increased size effect in terms of increased strength and stochasticity. Microcompression along the [23¯14] axis results in much lower strengths than for [0001] oriented samples. Post-mortem analysis reveals basal slip in both [0001] and [23¯14] micropillars. The application of a stochastic probability model shows good agreement between theoretical predictions and experimental results for size effects with our values of initial dislocation density and micropillar dimensions. Size effects are then incorporated into a single-crystal plasticity model (modified from Zhang and Joshi [1]) implemented in ABAQUS/STANDARD as a user-material subroutine. The model successfully captures the phenomena typically associated with size effects of increasing stochasticity and strength with decreasing specimen size and also accounts for the changing trends resulting from variations in initial dislocation density that we observe in the experiments. Finally, finite element simulations are performed with the original (traditional, without size effects) crystal plasticity model [1] to investigate the relative activities of the deformation modes of single-crystal magnesium for varying degrees of misalignment in microcompression. The simulations reveal basal activity in all micropillars, even for perfectly aligned compression along the [0001] axis. Pyramidal < c + a > activity dominates until the misalignment increases to 2°, when basal slip takes over as the dominant mode. The stress-strain curves for the case of 0° misalignment agrees well with experimental curves, indicating that good alignment was achieved during the experiments. Through this investigation, we gain a better understanding of how to control the size effects, as well as the deformation mechanisms operating at the small scale in magnesium.
Hughes, A. Randall; Rooker, Kelly; Murdock, Meagan; Kimbro, David L.
2012-01-01
Predators can influence prey abundance and traits by direct consumption, as well as by non-consumptive effects of visual, olfactory, or tactile cues. The strength of these non-consumptive effects (NCEs) can be influenced by a variety of factors, including predator foraging mode, temporal variation in predator cues, and the density of competing prey. Testing the relative importance of these factors for determining NCEs is critical to our understanding of predator-prey interactions in a variety of settings. We addressed this knowledge gap by conducting two mesocosm experiments in a tri-trophic intertidal oyster reef food web. More specifically, we tested how a predatory fish (hardhead catfish, Ariopsis felis) directly influenced their prey (mud crabs, Panopeus spp.) and indirectly affected basal resources (juvenile oysters, Crassostrea virginica), as well as whether these direct and indirect effects changed across a density gradient of competing prey. Per capita crab foraging rates were inversely influenced by crab density, but they were not affected by water-borne predator cues. As a result, direct consumptive effects on prey foraging rates were stronger than non-consumptive effects. In contrast, predator cue and crab density interactively influenced indirect predator effects on oyster mortality in two experiments, with trait-mediated and density-mediated effects of similar magnitude operating to enhance oyster abundance. Consistent differences between a variable predator cue environment and other predator cue treatments (no cue and constant cue) suggests that an understanding of the natural risk environment experienced by prey is critical to testing and interpreting trait-mediated indirect interactions. Further, the prey response to the risk environment may be highly dependent on prey density, particularly in prey populations with strong intra-specific interactions. PMID:22970316
Shear, C R; Bloch, R J
1985-07-01
Using immunocytochemical methods we have studied the distribution of vinculin in the anterior and posterior latissimus dorsi skeletal (ALD and PLD, respectively) muscles of the adult chicken. The ALD muscle is made up of both tonic (85%) and twitch (15%) myofibers, and the PLD muscle is made up entirely of twitch myofibers. In indirect immunofluorescence, antivinculin antibodies stained specific regions adjacent to the sarcolemma of the ALD and PLD muscles. In the central and myotendinous regions of the ALD, staining of the tonic fibers was intense all around the fiber periphery. Staining of the twitch fibers of both ALD and PLD muscles was intense only at neuromuscular junctions and myotendinous regions. Electron microscopy revealed subsarcolemmal, electron-dense plaques associated with the membrane only in those regions where vinculin was localized by immunofluorescence. Using antivinculin antibody and protein A conjugated to colloidal gold, we found that the electron-dense subsarcolemmal densities in the tonic fibers of the ALD contain vinculin; no other structures were labeled. The basal lamina overlying the densities appeared to be connected to the sarcolemma by fine, filamentous structures, more enriched at these sites than elsewhere along the muscle fiber. Increased amounts of endomysial connective tissue were often found just outside the basal lamina near the densities. In tonic ALD muscle fibers, the subsarcolemmal densities were present preferentially over the I-bands. In partially contracted ALD muscle, subsarcolemmal densities adjacent to the Z-disk appeared to be connected to that structure by short filaments. We propose that in the ALD muscle, through their association with the extracellular matrix, the densities stabilize the muscle membrane and perhaps assist in force transmission.
Mohania, Dheeraj; Kansal, Vinod Kumar; Shah, Dilip; Nagpal, Ravinder; Kumar, Manoj; Gautam, Sanjeev Kumar; Singh, Birbal; Behare, Pradip Vishnu
2013-09-01
This study examined the effects of probiotic dahi prepared by Lactobacillus plantarum Lp9 and dahi culture in buffalo milk on lowering cholesterol in rats fed a hypercholesterolemic basal diet. Male Wistar rats were divided into 3 groups and fed with probiotic dahi, dahi, or buffalo milk for 120 days. Following the consumption of supplements (probiotic dahi, dahi or buffalo milk), the animals were fed a basal hypercholesterolemic diet. Plasma total cholesterol and triglycerides (TAGs) were decreased by 35% and 72% in rats fed with probiotic dahi group, while cholesterol levels increased by 70% and TAGs increased by 97% in buffalo milk and 59% in dahi fed groups. Supplementation of probiotic dahi further lowered plasma low-density lipoprotein (LDL) + very-low-density lipoprotein (VLDL)- cholesterol by 59%, while it elevated plasma high-density lipoprotein (HDL)-cholesterol by 116%. As a result, atherogenic index, the ratio of HDL to LDL + VLDL was markedly improved. Deposition of cholesterol and TAGs in liver and aorta were significantly reduced in rats fed with probiotic dahi. These observations suggest that probiotic dahi may have therapeutic potential to decrease plasma, hepatic and aortic lipid profile, and attenuate diet-induced hypercholesterolemia.
Powers, Brian E.; Velazquez, Ramon; Kelley, Christy M.; Ash, Jessica A.; Strawderman, Myla S.; Alldred, Melissa J.; Ginsberg, Stephen D.; Mufson, Elliott J.
2016-01-01
Individuals with Down syndrome (DS) exhibit intellectual disability and develop Alzheimer's disease-like neuropathology during the third decade of life. The Ts65Dn mouse model of DS exhibits key features of both disorders, including impairments in learning, attention and memory, as well as atrophy of basal forebrain cholinergic neurons (BFCNs). The present study evaluated attentional function in relation to BFCN morphology in young (3 months) and middle-aged (12 months) Ts65Dn mice and disomic (2N) controls. Ts65Dn mice exhibited attentional dysfunction at both ages, with greater impairment in older trisomics. Density of BFCNs was significantly lower for Ts65Dn mice independent of age, which may contribute to attentional dysfunction since BFCN density was positively associated with performance on an attention task. BFCN volume decreased with age in 2N but not Ts65Dn mice. Paradoxically, BFCN volume was greater in older trisomic mice, suggestive of a compensatory response. In sum, attentional dysfunction occurred in both young and middle-aged Ts65Dn mice, which may in part reflect reduced density and/or phenotypic alterations in BFCNs. PMID:26719290
Powers, Brian E; Velazquez, Ramon; Kelley, Christy M; Ash, Jessica A; Strawderman, Myla S; Alldred, Melissa J; Ginsberg, Stephen D; Mufson, Elliott J; Strupp, Barbara J
2016-12-01
Individuals with Down syndrome (DS) exhibit intellectual disability and develop Alzheimer's disease-like neuropathology during the third decade of life. The Ts65Dn mouse model of DS exhibits key features of both disorders, including impairments in learning, attention and memory, as well as atrophy of basal forebrain cholinergic neurons (BFCNs). The present study evaluated attentional function in relation to BFCN morphology in young (3 months) and middle-aged (12 months) Ts65Dn mice and disomic (2N) controls. Ts65Dn mice exhibited attentional dysfunction at both ages, with greater impairment in older trisomics. Density of BFCNs was significantly lower for Ts65Dn mice independent of age, which may contribute to attentional dysfunction since BFCN density was positively associated with performance on an attention task. BFCN volume decreased with age in 2N but not Ts65Dn mice. Paradoxically, BFCN volume was greater in older trisomic mice, suggestive of a compensatory response. In sum, attentional dysfunction occurred in both young and middle-aged Ts65Dn mice, which may in part reflect reduced density and/or phenotypic alterations in BFCNs.
The Evaluation of Basal Respiration for Various Soil Textures in Ecologically Sensitive Area
NASA Astrophysics Data System (ADS)
Huličová, P.; Kotorová, D.; Fazekašová, D.; Hynšt, J.
2017-10-01
The present contribution was focused on monitoring changes in the soil basal respiration in different textures of soil in the dry polder Beša. The research was conducted between 2012 and 2014 on soil type Fluvisol locations on three soil textures: clay - loam soil, clayey soil and clay soil in three soil depths. The basal respiration (BR) has been determine by soil CO2 production measuring from incubated soil samples in serum bottles in laboratory condition. Release Co2 has been analysed by gas chromatography. Content of clay particles were in the range 52.18 % to 81.31%, indicating the high difference between the minimum and maximum content. By using of multiple LSD-test we recorded statistically significant impact of clay on basal respiration. Results confirm the values of basal respiration with the depth of the soil profile decreased.
NASA Astrophysics Data System (ADS)
Guthoff, Rudolf F.; Zhivov, Andrey; Stachs, Oliver
2010-02-01
The aim of the study was to produce two-dimensional reconstruction maps of the living corneal sub-basal nerve plexus by in vivo laser scanning confocal microscopy in real time. CLSM source data (frame rate 30Hz, 384x384 pixel) were used to create large-scale maps of the scanned area by selecting the Automatic Real Time (ART) composite mode. The mapping algorithm is based on an affine transformation. Microscopy of the sub-basal nerve plexus was performed on normal and LASIK eyes as well as on rabbit eyes. Real-time mapping of the sub-basal nerve plexus was performed in large-scale up to a size of 3.2mm x 3.2mm. The developed method enables a real-time in vivo mapping of the sub-basal nerve plexus which is stringently necessary for statistically firmed conclusions about morphometric plexus alterations.
Goldberg, Jesse H.; Adler, Avital; Bergman, Hagai; Fee, Michale S.
2010-01-01
The songbird area X is a basal ganglia homologue that contains two pallidal cell types—local neurons that project within the basal ganglia and output neurons that project to the thalamus. Based on these projections, it has been proposed that these classes are structurally homologous to the primate external (GPe) and internal (GPi) pallidal segments. To test the hypothesis that the two area X pallidal types are functionally homologous to GPe and GPi neurons, we recorded from neurons in area X of singing juvenile male zebra finches, and directly compare their firing patterns to neurons recorded in the primate pallidus. In area X, we find two cell classes that exhibited high firing (HF) rates (>60Hz) characteristic of pallidal neurons. HF-1 neurons, like most GPe neurons we examined, exhibited large firing rate modulations, including bursts and long pauses. In contrast, HF-2 neurons, like GPi neurons, discharged continuously without bursts or long pauses. To test if HF-2 neurons were the output neurons that project to the thalamus, we next recorded directly from pallidal axon terminals in thalamic nucleus DLM, and found that all terminals exhibited singing-related firing patterns indistinguishable from HF-2 neurons. Our data show that singing-related neural activity distinguishes two putative pallidal cell types in area X: thalamus-projecting neurons that exhibit activity similar to the primate GPi, and non-thalamus-projecting neurons that exhibit activity similar to the primate GPe. These results suggest that song learning in birds and motor learning in mammals employ conserved basal ganglia signaling strategies. PMID:20484651
Higuchi, P; Silva, A C; Louzada, J N C; Machado, E L M
2010-05-01
The objectives of this study were to evaluate the influence of propagules source and the implication of tree size class on the spatial pattern of Xylopia brasiliensis Spreng. individuals growing under the canopy of an experimental plantation of eucalyptus. To this end, all individuals of Xylopia brasiliensis with diameter at soil height (dsh) > 1 cm were mapped in the understory of a 3.16 ha Eucalyptus spp. and Corymbia spp. plantation, located in the municipality of Lavras, SE Brazil. The largest nearby mature tree of X. brasiliensis was considered as the propagules source. Linear regressions were used to assess the influence of the distance of propagules source on the population parameters (density, basal area and height). The spatial pattern of trees was assessed through the Ripley K function. The overall pattern showed that the propagules source distance had strong influence over spatial distribution of trees, mainly the small ones, indicating that the closer the distance from the propagules source, the higher the tree density and the lower the mean tree height. The population showed different spatial distribution patterns according to the spatial scale and diameter class considered. While small trees tended to be aggregated up to around 80 m, the largest individuals were randomly distributed in the area. A plausible explanation for observed patterns might be limited seed rain and intra-population competition.
Alterations of cortical pyramidal neurons in mice lacking high-affinity nicotinic receptors
Ballesteros-Yáñez, Inmaculada; Benavides-Piccione, Ruth; Bourgeois, Jean-Pierre; Changeux, Jean-Pierre; DeFelipe, Javier
2010-01-01
The neuronal nicotinic acetylcholine receptors (nAChRs) are allosteric membrane proteins involved in multiple cognitive processes, including attention, learning, and memory. The most abundant form of heterooligomeric nAChRs in the brain contains the β2- and α4- subunits and binds nicotinic agonists with high affinity. In the present study, we investigated in the mouse the consequences of the deletion of one of the nAChR components: the β2-subunit (β2−/−) on the microanatomy of cortical pyramidal cells. Using an intracellular injection method, complete basal dendritic arbors of 650 layer III pyramidal neurons were sampled from seven cortical fields, including primary sensory, motor, and associational areas, in both β2−/− and WT animals. We observed that the pyramidal cell phenotype shows significant quantitative differences among different cortical areas in mutant and WT mice. In WT mice, the density of dendritic spines was rather similar in all cortical fields, except in the prelimbic/infralimbic cortex, where it was significantly higher. In the absence of the β2-subunit, the most significant reduction in the density of spines took place in this high-order associational field. Our data suggest that the β2-subunit is involved in the dendritic morphogenesis of pyramidal neurons and, in particular, in the circuits that contribute to the high-order functional connectivity of the cerebral cortex. PMID:20534523
NASA Astrophysics Data System (ADS)
Grima, C.; Koch, I.; Greenbaum, J. S.; Soderlund, K. M.; Blankenship, D. D.; Young, D. A.; Fitzsimons, S.
2017-12-01
The McMurdo ice shelves (northern and southern MIS), adjacent to the eponymous station and the Ross Ice Shelf, Antarctica, are known for large gradients in surface snow accumulation and snow/ice impurities. Marine ice accretion and melting are important contributors to MIS's mass balance. Due to erosive winds, the southern MIS (SMIS) shows a locally negative surface mass balance. Thus, marine ice once accreted at the ice shelf base crops out at the surface. However, the exact processes that exert primary control on SMIS mass balance have remained elusive. Radar statistical reconnaissance (RSR) is a recent technique that has been used to characterize the surface properties of the Earth's cryosphere, Mars, and Titan from the stochastic character of energy scattered by the surface. Here, we apply RSR to map the surface density and roughness of the SMIS and extend the technique to derive the basal reflectance and scattering coefficients of the ice-ocean interface. We use an airborne radar survey grid acquired over the SMIS in the 2014-2015 austral summer by the University of Texas Institute for Geophysics with the High Capability Radar Sounder (HiCARS2; 60-MHz center frequency and 15-MHz bandwidth). The RSR-derived snow density values and patterns agree with directly -measured ice shelf surface accumulation rates. We also compare the composition of SMIS ice surface samples to test the ability of RSR to discriminate ices with varying dielectric properties (e.g., marine versus meteoric ice) and hypothesize relationships between the RSR-derived basal reflectance/scattered coefficients and accretion or melting at the ice-ocean interface. This improved knowledge of air-ice and ice-ocean boundaries provides a new perspective on the processes governing SMIS surface and basal mass balance.
Crespo, N; Esteve-Garcia, E
2003-07-01
An experiment was conducted to study the effect of different dietary fatty acid profiles on plasma levels of insulin, very low density lipoproteins (VLDL), cholesterol, and glucose. Diets with four types of fat (tallow, olive, sunflower, and linseed oils) at an inclusion level of 10% and a basal diet without additional fat were administered to female broiler chickens. Serum insulin, cholesterol, and plasma VLDL were affected by the different treatments; however, glucose concentrations were similar among treatments. In the fasted state, broilers fed diets with sunflower or linseed oil presented lower levels of insulin and cholesterol with respect to those fed tallow or olive oil (P < 0.05). VLDL in the fasted state was reduced in broilers fed sunflower and linseed oils (P < 0.05) with respect to those fed tallow, olive oil, or the basal diet. Plasma levels of VLDL were only significantly correlated with abdominal fat in birds fed the basal diet, in the fed and in the fasted state, and in those fed linseed oil in the fed state (P < 0.05). Results of this experiment suggest that higher insulin levels in broilers fed diets rich in saturated fatty acids could be related to higher fat deposition. Fat deposition in birds fed high fat diets was not correlated with circulating VLDL, which suggested direct dietary fat deposition, except for birds fed linseed oil diets. Although birds fed linseed oil diets presented lower levels of VLDL than those fed tallow, olive oil, or the basal diet, the higher correlation with abdominal fat suggests that in these birds, fat deposition is more dependent on hepatic VLDL secretion, despite the high dietary fat level.
Computer simulation of fat and muscle burn in long-distance bird migration
Pennycuick
1998-03-07
The mechanical power required from a bird's flight muscles was recalculated at regular intervals (default 6 min), and the energy consumed in the interval was accounted for by reducing fuel reserves, which also reduced the all-up mass and the body cross-sectional area. Part of the energy requirement was met by consuming flight muscle tissue, according to one of three alternative "muscle burn criteria". These were (1) specific work held constant, (2) power density held constant and (3) muscle mass held constant, i.e. no muscle consumed. Holding the specific work constant produced results in the best agreement with the results of other studies. This criterion was therefore selected to compare simulated flights of three very different species whose flight and migrations have been extensively studied, (1) Thrush Nightingale (Luscinia luscinia), (2) Knot (Calidris canutus) and (3) Whooper Swan (Cygnus cygnus). The ratio of protein to fat consumed ranged from 0.19 to 0.36, depending mainly on the starting value assumed for the muscle fraction. Specific work and starting power density were much higher for the Whooper Swan than for the two smaller species, suggesting that the latter have power to spare for climbing to high cruising altitudes, whereas the swan has not. If all three species were able to reach high cruising altitudes, the result would be a large reduction in journey time, which in turn would result in a small increase in range, due to a saving of energy required for basal metabolism. On current assumptions, the proportion of the fuel energy spent on basal metabolism would be eight times higher in the Thrush Nightingale than in the Whooper Swan, consequently the gain in range due to flying high would be greater in the smaller bird. In order to run the simulation, assumptions have been made at the primary physical level, to calculate the mechanical power required, and also at the secondary physiological level, to convert this into fuel consumption. The physical assumptions mostly take the form of variables whose existence is not in doubt, but whose values are poorly known, whereas in the case of some of the most important physiological variables, even the principles are unknown. Attention is drawn to a number of problems in need of attention, including (1) the mass and energy requirements of respiratory and circulatory organs required to sustain aerobically a given level of mechanical power; (2) the capabilities of bird lungs at high altitudes; (3) the possibility that heart muscle and lung tissue may be consumed in flight; (4) the two "biological constants", isometric force per myosin fibril and inverse power density of mitochondria; (5) the energy density of different fuels, and the conversion efficiency of the flight muscles; and (6) the manner in which basal metabolism combines with other demands for power in an exercising animal. Copyright 1998 Academic Press Limited
Scasta, John Derek; Engle, David M; Harr, Ryan N; Debinski, Diane M
2014-12-01
Symphoricarpos, a genus of the Caprifoliaceae family, consists of about 15 species of clonal deciduous shrubs in North America and 1 species endemic to China. In North American tallgrass prairie, Symphoricarpos orbiculatus (buckbrush) is the dominant shrub often forming large colonies via sexual and asexual reproductive mechanisms. Symphoricarpos shrubs, in particular S. orbiculatus, use a unique sexual reproductive mechanism known as layering where vertical stems droop and the tips root upon contact with the soil. Because of conflicting societal values of S. orbiculatus for conservation and agriculture and the current attempt to restore historical fire regimes, there is a need for basic research on the biological response of S. orbiculatus to anthropogenic burning regimes. From 2007 through 2013 we applied prescribed fires in the late dormant season on grazed pastures in the Grand River Grasslands of Iowa. From 2011 to 2013, we measured how S. orbiculatus basal resprouting and layering stems were affected by patchy fires on grazed pastures, complete pasture fires on grazed pastures or fire exclusion without grazing for more than three years. We measured ramet height, ramet canopy diameter, stems per ramet, ramets per 100 m 2 , and probability of new layering stems 120 days after fire. Height in burned plots was lower than unburned plots but S. orbiculatus reached ~ 84% of pre-burn height 120 days after fire. Stems per ramet were 2x greater in the most recently burned plots due to basal re-sprouting. Canopy diameter and density of ramets was not affected by time since fire, but burned pastures had marginally lower densities than plots excluded from fire (P = 0.07). Fire triggered new layering stems and no new layering stems were found in plots excluded from fire. The mechanisms of both basal sprouting and aerial layering after fire suggest S. orbiculatus is tolerant to dormant season fires. Furthermore, dormant season fires, regardless if they were patchy fires or complete pasture fires, did not result in mortality of S. orbiculatus. Dormant season fires can reduce S. orbiculatus structural dominance and maintain lower ramet densities but also trigger basal resprouting and layering.
Solute effect on basal and prismatic slip systems of Mg.
Moitra, Amitava; Kim, Seong-Gon; Horstemeyer, M F
2014-11-05
In an effort to design novel magnesium (Mg) alloys with high ductility, we present a first principles data based on the Density Functional Theory (DFT). The DFT was employed to calculate the generalized stacking fault energy curves, which can be used in the generalized Peierls-Nabarro (PN) model to study the energetics of basal slip and prismatic slip in Mg with and without solutes to calculate continuum scale dislocation core widths, stacking fault widths and Peierls stresses. The generalized stacking fault energy curves for pure Mg agreed well with other DFT calculations. Solute effects on these curves were calculated for nine alloying elements, namely Al, Ca, Ce, Gd, Li, Si, Sn, Zn and Zr, which allowed the strength and ductility to be qualitatively estimated based on the basal dislocation properties. Based on our multiscale methodology, a suggestion has been made to improve Mg formability.
Kobayashi, Katsumi; Yokoi, Yota; Masuzawa, Takehiro
2011-05-01
To determine the main benefits of clonal expansion of Miscanthus sinensis patches (monoclones), we observed the annual pattern of the areal expansion of a number of M. sinensis patches and examined how the quantity of rhizomes in such patches is related to changes in their basal area. To forage for nutriments, a patch must continuously widen its habitat. Patches annually expanded centrifugally by sympodial branching of short rhizomes, which originated in tillering that occurred more than once a year. However, the basal area of the patches approached a ceiling as the patches aged. Both the number and the weight of rhizomes in the patches continued to increase as long as the basal area expanded. The mean weight of rhizomes in patches also initially increased quickly, but then reached a ceiling as the clones expanded. Similarly, the amount of reserve substance per shoot in the patches increased asymptotically along with the clonal expansion, depending on the rhizome mass allotted to each shoot. These results suggest that, in the clonal growth of M. sinensis patches, the accumulation of reserve matter in the rhizomes is more important than foraging in new areas.
de Paula, Felipe Rossetti; Ferraz, Silvio Frosini de Barros; Gerhard, Pedro; Vettorazzi, Carlos Alberto; Ferreira, Anderson
2011-10-01
Riparian forests are important for the structure and functioning of stream ecosystems, providing structural components such as large woody debris (LWD). Changes in these forests will cause modifications in the LWD input to streams, affecting their structure. In order to assess the influence of riparian forests changes in LWD supply, 15 catchments (third and fourth order) with riparian forests at different conservation levels were selected for sampling. In each catchment we quantified the abundance, volume and diameter of LWD in stream channels; the number, area and volume of pools formed by LWD and basal area and tree diameter of riparian forest. We found that riparian forests were at a secondary successional stage with predominantly young trees (diameter at breast height <10 cm) in all studied streams. Results showed that basal area and diameter of riparian forest differed between the stream groups (forested and non-forested), but tree density did not differ between groups. Differences were also observed in LWD abundance, volume, frequency of LWD pools with subunits and area and volume of LWD pools. LWD diameter, LWD that form pools diameter and frequency of LWD pools without subunits did not differ between stream groups. Regression analyses showed that LWD abundance and volume, and frequency of LWD pools (with and without subunits) were positively related with the proportion of riparian forest. LWD diameter was not correlated to riparian tree diameter. The frequency of LWD pools was correlated to the abundance and volume of LWD, but characteristics of these pools (area and volume) were not correlated to the diameter of LWD that formed the pools. These results show that alterations in riparian forest cause modifications in the LWD abundance and volume in the stream channel, affecting mainly the structural complexity of these ecosystems (reduction in the number and structural characteristics of LWD pools). Our results also demonstrate that riparian forest conservation actions must consider not only its extension, but also successional stage to guarantee the quantity and quality of LWD necessary to enable the structuring of stream channels.
NASA Technical Reports Server (NTRS)
Chakrabarti, A.; Schatten, H.; Mitchell, K. D.; Crosser, M.; Taylor, M.
1998-01-01
The mitotic inhibitor, chloral hydrate, induces ciliary loss in the early embryo phase of Lytechinus pictus. It causes a breakdown of cilia at the junction of the cilium and the basal body known as the basal plate. This leaves the plasma membrane temporarily unsealed. The basal apparatus accessory structures, consisting of the basal body, basal foot, basal foot cap, striated side arm, and striated rootlet, are either misaligned or disintegrated by treatment with chloral hydrate. Furthermore, microtubules which are associated with the basal apparatus are disassembled. Mitochondria accumulate at the base of cilia - underneath the plasma membrane - and show alterations in their structural organization. The accumulation of mitochondria is observed in 40% of all electron micrograph sections while 60% show the areas mostly devoid of mitochondria. The microvilli surrounding a cilium and striated rootlet remain intact in the presence of chloral hydrate. These results suggest that deciliation in early sea urchin embryos by chloral hydrate is caused by combined effects on the ciliary membrane and on microtubules in the cilia. Furthermore, it is suggested that chloral hydrate can serve as a tool to explore the cytoskeletal mechanisms that are involved in cilia motility in the developing sea urchin embryo.
USDA-ARS?s Scientific Manuscript database
Basal stalk rot (BSR) caused by the ascomycete fungus Sclerotinia sclerotiorum (Lib.) de Bary is a serious disease of sunflower (Helianthus annuus L.) in the cool and humid production areas of the world. Quantitative trait loci (QTL) for BSR resistance were identified in a sunflower recombinant inbr...
Junior High Basals: Effective Hi/Lo Materials for Remedial High School Readers.
ERIC Educational Resources Information Center
Alvermann, Donna E.
1981-01-01
Discusses the results of an analysis of the appropriateness of eighth-grade basal reading materials for remedial instruction of ninth- and tenth-grade students who read two to three years below grade level. Readability, interest appeal, and representation of content areas are considered. Three data tables and a 14-item reference list are included.…
Extensive Lesions of Cholinergic Basal Forebrain Neurons Do Not Impair Spatial Working Memory
ERIC Educational Resources Information Center
Vuckovich, Joseph A.; Semel, Mara E.; Baxter, Mark G.
2004-01-01
A recent study suggests that lesions to all major areas of the cholinergic basal forebrain in the rat (medial septum, horizontal limb of the diagonal band of Broca, and nucleus basalis magnocellularis) impair a spatial working memory task. However, this experiment used a surgical technique that may have damaged cerebellar Purkinje cells. The…
USDA-ARS?s Scientific Manuscript database
Basal endosperm transfer cells (BETCs) constitute one of the four cell types in an endosperm with a major role in solute acquisition and transport functions from the mother plant. The BETCs with their wall-in-growth (WIG) feature that greatly increase plasma membrane area of each cell are critical f...
NASA Astrophysics Data System (ADS)
McEathron, K. M.; Mitchell, M. J.; Zhang, L.
2013-07-01
Grass Pond watershed is located within the southwestern Adirondack Mountain region of New York State, USA. This region receives some of the highest rates of acidic deposition in North America and is particularly sensitive to acidic inputs due to many of its soils having shallow depths and being generally base poor. Differences in soil chemistry and tree species between seven subwatersheds were examined in relation to acid-base characteristics of the seven major streams that drain into Grass Pond. Mineral soil pH, stream water BCS (base-cation surplus) and pH exhibited a positive correlation with sugar maple basal area (p = 0.055; 0.48 and 0.39, respectively). Black cherry basal area was inversely correlated with stream water BCS, ANC (acid neutralizing capacity)c and NO3- (p = 0.23; 0.24 and 0.20, respectively). Sugar maple basal areas were positively associated with watershed characteristics associated with the neutralization of atmospheric acidic inputs while in contrast, black cherry basal areas showed opposite relationships to these same watershed characteristics. Canonical correspondence analysis indicated that black cherry had a distinctive relationship with forest floor chemistry apart from the other tree species, specifically a strong positive association with forest floor NH4, while sugar maple had a distinctive relationship with stream chemistry variables, specifically a strong positive association with stream water ANCc, BCS and pH. Our results provide evidence that sugar maple is acid-intolerant or calciphilic tree species and also demonstrate that black cherry is likely an acid-tolerant tree species.
Isolation, separation, and characterization of epithelial and connective cells from rat palate
DOE Office of Scientific and Technical Information (OSTI.GOV)
Terranova, Victor Paul
1979-01-01
Epithelial and connective tissue cells were isolated from rat palate by sequential collagenase, hyaluronidase and trypsin digestion of the extracellular matrix. Differences between the two populations were noted with respect to total cell protein, total cell water, proline uptake and incorporation, percent collagen synthesized, effects of parathyroid hormone, metabolism of D-valine and cell density. Basal epithelial cells were subsequently separated from the heterogeneous epithelial cell population on shallow linear density gradients by velocity centrifugation. The type of collagen synthesized by the basal epithelial cells was compared to the type of collagen synthesized by the connective tissue cells by means ofmore » labeled amino acid incorporation ratios. Cells isolated from the epithelial and connective tissue were compared. From these studies it can be concluded that epithelial and connective tissue cells can be isolated from rat palate as viable and distinct populations with respect to the biochemical parameters examined. Furthermore, subpopulations can be separated and biochemically characterized.« less
Acoustic plane wave preferential orientation of metal oxide superconducting materials
Tolt, Thomas L.; Poeppel, Roger B.
1991-01-01
A polycrystalline metal oxide such as YBa.sub.2 Cu.sub.3 O.sub.7-X (where 0
He, Shaojun; Zhao, Shujing; Dai, Sifa; Liu, Deyi; Bokhari, Shehla Gul
2015-10-01
We evaluated the effects of supplementing betaine on growth performance, fat deposition and lipid metabolism status in broilers kept under chronic heat stress. Five hundred and forty chicks were randomly divided into six groups and the two normal temperature groups were held at normal ambient temperature and fed the basal diet (CONT) and basal diet plus 0.1% betaine, respectively. Heat stressed (HS) broilers were held at 32 ± 1°C from days 22 to 42 and fed the basal diet containing variable levels of betaine. Broilers were examined at days 28, 35 and 42 for body weight, feed consumption, fat deposition and serum lipids. The CONT and betaine-supplemented groups showed higher (P < 0.01 or P < 0.05) feed consumption, body weight gain, and lower feed : gain ratio compared with the HS-CONT group. Meanwhile, heat stress increased abdominal, intermuscular and subcutaneous fat deposition, whereas the supplemental betaine significantly decreased those compared with the HS-CONT group. Additionally, betaine supplementation significantly decreased triglyceride, free fatty acids, low-density lipoprotein cholesterol and high-density lipoprotein cholesterol compared with HS-CONT. Chronic HS reduces broiler production performance. However, betaine can reverse these negative effects partially and thus improve carcass composition by changing lipid metabolism. © 2015 Japanese Society of Animal Science.
Basal ganglia function, stuttering, sequencing, and repair in adult songbirds.
Kubikova, Lubica; Bosikova, Eva; Cvikova, Martina; Lukacova, Kristina; Scharff, Constance; Jarvis, Erich D
2014-10-13
A pallial-basal-ganglia-thalamic-pallial loop in songbirds is involved in vocal motor learning. Damage to its basal ganglia part, Area X, in adult zebra finches has been noted to have no strong effects on song and its function is unclear. Here we report that neurotoxic damage to adult Area X induced changes in singing tempo and global syllable sequencing in all animals, and considerably increased syllable repetition in birds whose song motifs ended with minor repetitions before lesioning. This stuttering-like behavior started at one month, and improved over six months. Unexpectedly, the lesioned region showed considerable recovery, including immigration of newly generated or repaired neurons that became active during singing. The timing of the recovery and stuttering suggest that immature recovering activity of the circuit might be associated with stuttering. These findings indicate that even after juvenile learning is complete, the adult striatum plays a role in higher level organization of learned vocalizations.
Glenday, Julia
2008-07-01
Concerns about rapid tropical deforestation, and its contribution to rising atmospheric concentrations of greenhouse gases, increase the importance of monitoring terrestrial carbon storage in changing landscapes. Emerging markets for carbon emission offsets may offer developing nations needed incentives for reforestation, rehabilitation, and avoided deforestation. However, relatively little empirical data exists regarding carbon storage in African tropical forests, particularly for those in arid or semi-arid regions. Kenya's 416 km(2) Arabuko-Sokoke Forest (ASF) is the largest remaining fragment of East African coastal dry forest and is considered a global biodiversity hotspot (Myers et al. 2000), but has been significantly altered by past commercial logging and ongoing extraction. Forest carbon storage for ASF was estimated using allometric equations for tree biomass, destructive techniques for litter and herbaceous vegetation biomass, and spectroscopy for soils. Satellite imagery was used to assess land cover changes from 1992 to 2004. Forest and thicket types (Cynometra webberi dominated, Brachystegia spiciformis dominated, and mixed species forest) had carbon densities ranging from 58 to 94 Mg C/ha. The ASF area supported a 2.8-3.0 Tg C carbon stock. Although total forested area in ASF did not change over the analyzed time period, ongoing disturbances, quantified by the basal area of cut tree stumps per sample plot, correlated with decreased carbon densities. Madunguni Forest, an adjoining forest patch, lost 86% of its forest cover and at least 76% of its terrestrial carbon stock in the time period. Improved management of wood harvesting in ASF and rehabilitation of Madunguni Forest could substantially increase terrestrial carbon sequestration in the region.
Large-scale patterns of turnover and Basal area change in Andean forests.
Báez, Selene; Malizia, Agustina; Carilla, Julieta; Blundo, Cecilia; Aguilar, Manuel; Aguirre, Nikolay; Aquirre, Zhofre; Álvarez, Esteban; Cuesta, Francisco; Duque, Álvaro; Farfán-Ríos, William; García-Cabrera, Karina; Grau, Ricardo; Homeier, Jürgen; Linares-Palomino, Reynaldo; Malizia, Lucio R; Cruz, Omar Melo; Osinaga, Oriana; Phillips, Oliver L; Reynel, Carlos; Silman, Miles R; Feeley, Kenneth J
2015-01-01
General patterns of forest dynamics and productivity in the Andes Mountains are poorly characterized. Here we present the first large-scale study of Andean forest dynamics using a set of 63 permanent forest plots assembled over the past two decades. In the North-Central Andes tree turnover (mortality and recruitment) and tree growth declined with increasing elevation and decreasing temperature. In addition, basal area increased in Lower Montane Moist Forests but did not change in Higher Montane Humid Forests. However, at higher elevations the lack of net basal area change and excess of mortality over recruitment suggests negative environmental impacts. In North-Western Argentina, forest dynamics appear to be influenced by land use history in addition to environmental variation. Taken together, our results indicate that combinations of abiotic and biotic factors that vary across elevation gradients are important determinants of tree turnover and productivity in the Andes. More extensive and longer-term monitoring and analyses of forest dynamics in permanent plots will be necessary to understand how demographic processes and woody biomass are responding to changing environmental conditions along elevation gradients through this century.
Large-Scale Patterns of Turnover and Basal Area Change in Andean Forests
Blundo, Cecilia; Aguilar, Manuel; Aguirre, Nikolay; Aquirre, Zhofre; Álvarez, Esteban; Cuesta, Francisco; Farfán-Ríos, William; García-Cabrera, Karina; Grau, Ricardo; Linares-Palomino, Reynaldo; Malizia, Lucio R.; Cruz, Omar Melo; Osinaga, Oriana; Reynel, Carlos; Silman, Miles R.
2015-01-01
General patterns of forest dynamics and productivity in the Andes Mountains are poorly characterized. Here we present the first large-scale study of Andean forest dynamics using a set of 63 permanent forest plots assembled over the past two decades. In the North-Central Andes tree turnover (mortality and recruitment) and tree growth declined with increasing elevation and decreasing temperature. In addition, basal area increased in Lower Montane Moist Forests but did not change in Higher Montane Humid Forests. However, at higher elevations the lack of net basal area change and excess of mortality over recruitment suggests negative environmental impacts. In North-Western Argentina, forest dynamics appear to be influenced by land use history in addition to environmental variation. Taken together, our results indicate that combinations of abiotic and biotic factors that vary across elevation gradients are important determinants of tree turnover and productivity in the Andes. More extensive and longer-term monitoring and analyses of forest dynamics in permanent plots will be necessary to understand how demographic processes and woody biomass are responding to changing environmental conditions along elevation gradients through this century. PMID:25973977
Mathis, Kristian; Capek, J.; Clausen, Bjorn; ...
2015-04-20
Influence of aluminium content on the deformation mechanisms in Mg–Al binary alloys has been studied using in-situ neutron diffraction and acoustic emission technique. Here, it is shown that the addition of the solute increases the critical resolved shear stress for twinning. Further, the role of aluminium on the solid solution hardening of the basal plane and softening of non-basal planes are discussed using results of the convolutional multiple peak profile analysis of diffraction patterns. In conclusion, the results indicate that the density of both prismatic and pyramidal dislocations increases with increasing alloying content.
NASA Astrophysics Data System (ADS)
Gu, H.; Townsend, P. A.; Singh, A.
2014-12-01
Urban forests provide important ecosystem services related to climate, nutrients, runoff and aesthetics. Assessment of variations in urban forest growth is critical to urban management and planning, as well as to identify responses to climate and other environmental changes. We estimated annual relative basal area increment by tree rings from 37 plots in Madison, Wisconsin and neighboring municipalities. We related relative basal area growth to variables of vegetation traits derived from remote sensing, including structure (aboveground biomass, diameter, height, basal area, crown width and crown length) from discrete-return airborne lidar, and biochemical variables (foliar nitrogen, carbon, lignin, cellulose, fiber and LMA), spectral indices (NDVI, NDWI, PRI, NDII etc.) and species composition from AVIRIS hyperspectral imagery. Variations in tree growth was mainly correlated with tree species composition (R2 = 0.29, RMSE = 0.004) with coniferous stands having a faster growth rate than broadleaf plots. Inclusion of stand basal area improved model prediction from R2 = 0.29 to 0.35, with RMSE = 0.003. Then, we assessed the growth by functional type, we found that foliar lignin concentration and the proportion of live coniferous trees explained 57% variance in the growth of conifer stands. In contrast, broadleaf forest growth was more strongly correlated with species composition and foliar carbon (R2 = 0.59, RMSE = 0.003). Finally, we compared the relative basal area growth by species. In our study area, red pine and white pine exhibited higher growth rates than other species, while white oak plots grew slowest. There is a significant negative relationship between tree height and the relative growth in red pine stands (r = -0.95), as well as a strong negative relationship between crown width and the relative growth in white pine stands (r = -0.87). Growth declines as trees grow taller and wider may partly be the result of reduced photosynthesis and water availability. We also found that canopy cellulose content was negatively correlated with growth in white oak (r = -0.59), which could be caused by trade off of carbon allocation from shoot storage to leaves. These results demonstrate the potential of lidar and hyperspectral imagery to characterize important traits associated with biomass accumulation in urban forests.
Puszta, András; Katona, Xénia; Bodosi, Balázs; Pertich, Ákos; Nyujtó, Diána; Braunitzer, Gábor; Nagy, Attila
2018-01-01
The computer-based Rutgers Acquired Equivalence test (RAET) is a widely used paradigm to test the function of subcortical structures in visual associative learning. The test consists of an acquisition (pair learning) and a test (rule transfer) phase, associated with the function of the basal ganglia and the hippocampi, respectively. Obviously, such a complex task also requires cortical involvement. To investigate the activity of different cortical areas during this test, 64-channel EEG recordings were recorded in 24 healthy volunteers. Fast-Fourier and Morlet wavelet convolution analyses were performed on the recordings. The most robust power changes were observed in the theta (4–7 Hz) and gamma (>30 Hz) frequency bands, in which significant power elevation was observed in the vast majority of the subjects, over the parieto-occipital and temporo-parietal areas during the acquisition phase. The involvement of the frontal areas in the acquisition phase was remarkably weaker. No remarkable cortical power elevations were found in the test phase. In fact, the power of the alpha and beta bands was significantly decreased over the parietooccipital areas. We conclude that the initial acquisition of the image pairs requires strong cortical involvement, but once the pairs have been learned, neither retrieval nor generalization requires strong cortical contribution. PMID:29867412
Puszta, András; Katona, Xénia; Bodosi, Balázs; Pertich, Ákos; Nyujtó, Diána; Braunitzer, Gábor; Nagy, Attila
2018-01-01
The computer-based Rutgers Acquired Equivalence test (RAET) is a widely used paradigm to test the function of subcortical structures in visual associative learning. The test consists of an acquisition (pair learning) and a test (rule transfer) phase, associated with the function of the basal ganglia and the hippocampi, respectively. Obviously, such a complex task also requires cortical involvement. To investigate the activity of different cortical areas during this test, 64-channel EEG recordings were recorded in 24 healthy volunteers. Fast-Fourier and Morlet wavelet convolution analyses were performed on the recordings. The most robust power changes were observed in the theta (4-7 Hz) and gamma (>30 Hz) frequency bands, in which significant power elevation was observed in the vast majority of the subjects, over the parieto-occipital and temporo-parietal areas during the acquisition phase. The involvement of the frontal areas in the acquisition phase was remarkably weaker. No remarkable cortical power elevations were found in the test phase. In fact, the power of the alpha and beta bands was significantly decreased over the parietooccipital areas. We conclude that the initial acquisition of the image pairs requires strong cortical involvement, but once the pairs have been learned, neither retrieval nor generalization requires strong cortical contribution.
Evaluation of basal sex hormone levels for activation of the hypothalamic-pituitary-gonadal axis.
Ding, Yu; Li, Juan; Yu, Yongguo; Yang, Peirong; Li, Huaiyuan; Shen, Yongnian; Huang, Xiaodong; Liu, Shijian
2018-03-28
This study aimed to identify the predictive value of basal sex hormone levels for activation of the hypothalamic-pituitary-gonadal (HPG) axis in girls. Gonadotropin-releasing hormone (GnRH) stimulation tests were performed and evaluated in a total of 1750 girls with development of secondary sex characteristics. Correlation analyses were conducted between basal sex hormones and peak luteinizing hormone (LH) levels ≥5 IU/L during the GnRH stimulation test. Receiver operating characteristic (ROC) curves for basal levels of LH, follicle-stimulating hormone (FSH), LH/FSH, and estradiol (E2) before the GnRH stimulation test were plotted. The area under the curve (AUC) and 95% confidence intervals (CIs) were measured for each curve. The maximum AUC value was observed for basal LH levels (0.77, 95% CI: 0.74-0.79), followed by basal FSH levels (0.73, 95% CI: 0.70-0.75), the basal LH/FSH ratio (0.68, 95% CI: 0.65-0.71), and basal E2 levels (0.61, 95% CI: 0.59-0.64). The appropriate cutoff value of basal LH levels associated with a positive response of the GnRH stimulation test was 0.35 IU/L, with a sensitivity of 63.96% and specificity of 76.3% from the ROC curves when Youden's index showed the maximum value. When 100% of patients had peak LH levels ≥5 IU/L, basal LH values were >2.72 IU/L, but the specificity was only 5.45%. Increased basal LH levels are a significant predictor of a positive response during the GnRH stimulation test for assessing activation of the HPG axis in most girls with early pubertal signs.
Application of Lidar remote sensing to the estimation of forest canopy and stand structure
NASA Astrophysics Data System (ADS)
Lefsky, Michael Andrew
A new remote sensing instrument, SLICER (Scanning Lidar Imager of Canopies by Echo Recovery), has been applied to the problem of remote sensing the canopy and stand structure of two groups of deciduous forests, Tulip Poplar-Oak stands in the vicinity of Annapolis, MD. and bottomland hardwood stands near Williamston, NC. The ability of the SLICER instrument to remotely sense the vertical distribution of canopy structure (Canopy Height Profile), bulk canopy transmittance, and several indices of canopy height has been successfully validated using twelve stands with coincident field and SLICER estimates of canopy structure. Principal components analysis has been applied to canopy height profiles from both field sites, and three significant factors were identified, each closely related to the amount of foliage in a recognizable layer of the forest, either understory, midstory, or overstory. The distribution of canopy structure to these layers is significantly correlated with the size and number of stems supporting them. The same layered structure was shown to apply to both field and SLICER remotely sensed canopy height profiles, and to apply to SLICER remotely sensed canopy profiles from both the bottomland hardwood stands in the coastal plain of North Carolina, and to mesic Tulip-Poplars stands in the upland coastal plain of Maryland. Linear regressions have demonstrated that canopy and stand structure are correlated to both a statistically significant and useful degree. Stand age and stem density is more highly correlated to stand height, while stand basal area and aboveground biomass are more closely related to a new measure of canopy structure, the quadratic mean canopy height. A geometric model of canopy structure has been shown to explain the differing relationships between canopy structure and stand basal area for stands of Eastern Deciduous Forest and Douglas Fir Forest.
NASA Astrophysics Data System (ADS)
Wood, Jeffrey D.; Knapp, Benjamin O.; Muzika, Rose-Marie; Stambaugh, Michael C.; Gu, Lianhong
2018-01-01
Forests are expected to become more vulnerable to drought-induced tree mortality owing to rising temperatures and changing precipitation patterns that amplify drought lethality. There is a crucial knowledge gap regarding drought-pathogen interactions and their effects on tree mortality. The objectives of this research were to examine whether stand dynamics and ‘background’ mortality rates were affected by a severe drought in 2012; and to evaluate the importance of drought-pathogen interactions within the context of a mortality event that killed 10.0% and 26.5% of white (Quercus alba L.) and black (Q. velutina Lam.) oak stems, respectively, in a single year. We synthesized (i) forest inventory data (24 years), (ii) 11 years of ecosystem flux data with supporting biological data including predawn leaf water potential and annual forest inventories, (iii) tree-ring analyses of individual white oaks that were alive and ones that died in 2013, and (iv) documentation of a pathogen infection. This forest displayed stand dynamics consistent with expected patterns of decreasing tree density and increasing basal area. Continued basal area growth outpaced mortality implying a net accumulation of live biomass, which was supported by eddy covariance ecosystem carbon flux observations. Individual white and black oaks that died in 2013 displayed historically lower growth with the majority of dead trees exhibiting Biscogniauxia cankers. Our observations point to the importance of event-based oak mortality and that drought-Biscogniauxia interactions are important in shaping oak stand dynamics in this region. Although forest function has not been significantly impaired, these drought-pathogen interactions could amplify mortality under future climate conditions and thus warrant further investigation.
Ramírez, David A; Balaguer, Luis; Mancilla, Rosa; González, Virginia; Coaguila, Daniel; Talavera, Carmelo; Villegas, Luis; Ortega, Aldo; Jiménez, Percy; Moreno, José M
2012-01-01
Myrcianthes ferreyrae is an endemic, endangered species, with a small number of individuals located only in hyperarid, fog-oases known as lomas along the Peruvian desert in southern Peru, where fog is the main source of water. Following centuries of severe deforestation, reforestation with this native species was conducted in the Atiquipa lomas, Arequipa-Perú. On five slopes, five 2-year-old seedlings were irrigated monthly with water trapped by raschel-mesh fog collectors, supplementing natural rainfall with 0, 20, 40, 60 and 80 mm month(-1) from February to August 2008. We measured plant growth, increment in basal diameter, height and five leaf traits: leaf mass area (LMA), leaf carbon isotope composition (δ(13)C), nitrogen per leaf area, total leaf carbon and stomatal density; which are indicative of the physiological changes resulting from increased water supply. Plant growth rates, estimated from the variation of either shoot basal diameter or maximum height, were highly correlated with total biomass. Only LMA and δ(13)C were higher in irrigated than in control plants, but we found no further differences among irrigation treatments. This threshold response suggests an on-off strategy fitted to exploit pulses of fog water, which are always limited in magnitude in comparison with natural rain. The absence of a differential response to increased water supply is in agreement with the low phenotypic plasticity expected in plants from very stressful environments. Our results have practical implications for reforestation projects, since irrigating with 20 mm per month is sufficient to achieve the full growth capacity of this species.
Acker, S.A.; Beechie, T.J.; Shafroth, P.B.
2008-01-01
Ephemeral dams caused by landslides have been observed around the world, yet little is known about the effects of their failure on landforms and vegetation. In 1967, a landslide-dam-break flood in a pristine reach of the Elwha River valley filled the former channel and diverted the river. The reach is a reference site for restoration following the planned removal of dams on the river. We identified five surfaces on the 25 ha debris fan deposited by the flood. Based on tree ages and historic air photos, three of the surfaces formed in 1967, while two formed later. The surfaces varied in substrate (silt and sand, to boulders), and height above the river channel. Tree mortality resulted from tree removal and burial by sediment, the latter leaving snags and some surviving trees. Tree species composition was generally consistent within each surface. Dominant species included red alder (Alnus rubra) and Sitka willow (Salix sitchensis), alone or in combination, a combination of Douglas-fir (Pseudotsuga menziesii) and black cottonwood (Populus balsamifera ssp. trichocarpa), or a combination of alder and Cottonwood. There were significant differences between surfaces in stem density, basal area, and rate of basal area growth. The large degree of heterogeneity in forest structure, composition, and productivity within a relatively small floodplain feature is in part due to spatial variability in the intensity of a single disturbance event, and in part due to the occurrence of subsequent, smaller events. To recreate natural diversity of riparian forests may require mimicking the variety of physical and biotic habitats that a single, complex disturbance event may create.
Wood, Jeffrey D.; Knapp, Benjamin O.; Muzika, Rose-Marie; ...
2017-10-20
Forests are expected to become more vulnerable to drought-induced tree mortality owing to rising temperatures and changing precipitation patterns that amplify drought lethality. There is a crucial knowledge gap regarding drought–pathogen interactions and their effects on tree mortality. The objectives of this research were to examine whether stand dynamics and 'background' mortality rates were affected by a severe drought in 2012; and to evaluate the importance of drought–pathogen interactions within the context of a mortality event that killed 10.0% and 26.5% of white (Quercus alba L.) and black (Q. velutina Lam.) oak stems, respectively, in a single year. We synthesizedmore » (i) forest inventory data (24 years), (ii) 11 years of ecosystem flux data with supporting biological data including predawn leaf water potential and annual forest inventories, (iii) tree-ring analyses of individual white oaks that were alive and ones that died in 2013, and (iv) documentation of a pathogen infection. This forest displayed stand dynamics consistent with expected patterns of decreasing tree density and increasing basal area. Continued basal area growth outpaced mortality implying a net accumulation of live biomass, which was supported by eddy covariance ecosystem carbon flux observations. Individual white and black oaks that died in 2013 displayed historically lower growth with the majority of dead trees exhibiting Biscogniauxia cankers. Our observations point to the importance of event-based oak mortality and that drought–Biscogniauxia interactions are important in shaping oak stand dynamics in this region. Although forest function has not been significantly impaired, these drought–pathogen interactions could amplify mortality under future climate conditions and thus warrant further investigation.« less
Villa, Paola; Costantini, Barbara; Suriano, Rosanna; Perri, Concetta; Macrì, Francesca; Ricciardi, Luigi; Panunzi, Simona; Lanzone, Antonio
2009-02-01
The wide family of the phytoestrogens has become an alternative to the classical hormonal therapy in menopause; nevertheless, some findings are still conflicting. To examine the effect of genistein administration on metabolic parameters and vascular reactivity considering the basal endocrine status of the patients. A randomized placebo controlled study was conducted at a university hospital. Fifty postmenopausal women participated. Thirty subjects (group A) were randomized to receive 54 mg/d genistein while 20 subjects (group B) were treated with the placebo for 24 wk. In group A, we distinguish two subgroups: 14 normoinsulinemic and 12 hyperinsulinemic patients. Anthropometric measures, hormonal and lipid assays, oral glucose tolerance test with glycemic, insulin, and C-peptide evaluation, indexes of insulin sensitivity and endothelial function, and euglycemic-hyperinsulinemic clamps were performed. The insulin basal values significantly decreased in group A, whereas the homeostasis model index of insulin sensitivity and the fasting glucose levels significantly improved compared with placebo group. The genistein administration decreased fasting glucose and area under the curve glucose levels in the normoinsulinemic patients after treatment. In the hyperinsulinemic patients, a significant reduction in fasting insulin, fasting C-peptide, and area under the curve insulin levels as well as an increase in fractional hepatic insulin extraction was shown. In these patients, high-density lipoprotein cholesterol levels were significantly improved. The endothelium-dependent and -independent dilatation improved in the treated group. Normoinsulinemic patients showed both a significantly enhanced flow-mediated and nitrate-mediated dilatation, whereas no significant changes were found in the hyperinsulinemic group. The glycoinsulinemic metabolism and the endothelial function were significantly influenced by genistein. In particular, normoinsulinemic patients showed an improvement in glycemic and vascular reactivity indexes. Conversely, an improvement in the insulin sensitivity indexes was noted in hyperinsulinemic patients.
DOE Office of Scientific and Technical Information (OSTI.GOV)
Wood, Jeffrey D.; Knapp, Benjamin O.; Muzika, Rose-Marie
Forests are expected to become more vulnerable to drought-induced tree mortality owing to rising temperatures and changing precipitation patterns that amplify drought lethality. There is a crucial knowledge gap regarding drought–pathogen interactions and their effects on tree mortality. The objectives of this research were to examine whether stand dynamics and 'background' mortality rates were affected by a severe drought in 2012; and to evaluate the importance of drought–pathogen interactions within the context of a mortality event that killed 10.0% and 26.5% of white (Quercus alba L.) and black (Q. velutina Lam.) oak stems, respectively, in a single year. We synthesizedmore » (i) forest inventory data (24 years), (ii) 11 years of ecosystem flux data with supporting biological data including predawn leaf water potential and annual forest inventories, (iii) tree-ring analyses of individual white oaks that were alive and ones that died in 2013, and (iv) documentation of a pathogen infection. This forest displayed stand dynamics consistent with expected patterns of decreasing tree density and increasing basal area. Continued basal area growth outpaced mortality implying a net accumulation of live biomass, which was supported by eddy covariance ecosystem carbon flux observations. Individual white and black oaks that died in 2013 displayed historically lower growth with the majority of dead trees exhibiting Biscogniauxia cankers. Our observations point to the importance of event-based oak mortality and that drought–Biscogniauxia interactions are important in shaping oak stand dynamics in this region. Although forest function has not been significantly impaired, these drought–pathogen interactions could amplify mortality under future climate conditions and thus warrant further investigation.« less
On the relations between cratonic lithosphere thickness, plate motions, and basal drag
Artemieva, I.M.; Mooney, W.D.
2002-01-01
An overview of seismic, thermal, and petrological evidence on the structure of Precambrian lithosphere suggests that its local maximum thickness is highly variable (140-350 km), with a bimodal distribution for Archean cratons (200-220 km and 300-350 km). We discuss the origin of such large differences in lithospheric thickness, and propose that the lithospheric base can have large depth variations over short distances. The topography of Bryce Canyon (western USA) is proposed as an inverted analog of the base of the lithosphere. The horizontal and vertical dimensions of Archean cratons are strongly correlated: larger cratons have thicker lithosphere. Analysis of the bimodal distribution of lithospheric thickness in Archean cratons shows that the "critical" surface area for cratons to have thick (>300 km) keels is >6-8 ?? 106 km2 . Extrapolation of the linear trend between Archean lithospheric thickness and cratonic area to zero area yields a thickness of 180 km. This implies that the reworking of Archean crust should be accompanied by thinning and reworking of the entire lithospheric column to a thickness of 180 km in accord with thickness estimates for Proterozoic lithosphere. Likewise, extrapolation of the same trend to the size equal to the total area of all Archean cratons implies that the lithospheric thickness of a hypothesized early Archean supercontinent could have been 350-450 km decreasing to 280-400 km for Gondwanaland. We evaluate the basal drag model as a possible mechanism that may thin the cratonic lithosphere. Inverse correlations are found between lithospheric thickness and (a) fractional subduction length and (b) the effective ridge length. In agreement with theoretical predictions, lithospheric thickness of Archean keels is proportional to the square root of the ratio of the craton length (along the direction of plate motion) to the plate velocity. Large cratons with thick keels and low plate velocities are less eroded by basal drag than small fast-moving cratons. Basal drag may have varied in magnitude over the past 4 Ga. Higher mantle temperatures in the Archean would have resulted in lower mantle viscosity. This in turn would have reduced basal drag and basal erosion, and promoted the preservation of thick (>300 km) Archean keels, even if plate velocities were high during the Archean. ?? 2002 Elsevier Science B.V. All rights reserved.
Wichmann, Thomas; Bergman, Hagai; DeLong, Mahlon R
2018-03-01
Studies in non-human primates (NHPs) have led to major advances in our understanding of the function of the basal ganglia and of the pathophysiologic mechanisms of hypokinetic movement disorders such as Parkinson's disease and hyperkinetic disorders such as chorea and dystonia. Since the brains of NHPs are anatomically very close to those of humans, disease states and the effects of medical and surgical approaches, such as deep brain stimulation (DBS), can be more faithfully modeled in NHPs than in other species. According to the current model of the basal ganglia circuitry, which was strongly influenced by studies in NHPs, the basal ganglia are viewed as components of segregated networks that emanate from specific cortical areas, traverse the basal ganglia, and ventral thalamus, and return to the frontal cortex. Based on the presumed functional domains of the different cortical areas involved, these networks are designated as 'motor', 'oculomotor', 'associative' and 'limbic' circuits. The functions of these networks are strongly modulated by the release of dopamine in the striatum. Striatal dopamine release alters the activity of striatal projection neurons which, in turn, influences the (inhibitory) basal ganglia output. In parkinsonism, the loss of striatal dopamine results in the emergence of oscillatory burst patterns of firing of basal ganglia output neurons, increased synchrony of the discharge of neighboring basal ganglia neurons, and an overall increase in basal ganglia output. The relevance of these findings is supported by the demonstration, in NHP models of parkinsonism, of the antiparkinsonian effects of inactivation of the motor circuit at the level of the subthalamic nucleus, one of the major components of the basal ganglia. This finding also contributed strongly to the revival of the use of surgical interventions to treat patients with Parkinson's disease. While ablative procedures were first used for this purpose, they have now been largely replaced by DBS of the subthalamic nucleus or internal pallidal segment. These procedures are not only effective in the treatment of parkinsonism, but also in the treatment of hyperkinetic conditions (such as chorea or dystonia) which result from pathophysiologic changes different from those underlying Parkinson's disease. Thus, these interventions probably do not counteract specific aspects of the pathophysiology of movement disorders, but non-specifically remove the influence of the different types of disruptive basal ganglia output from the relatively intact portions of the motor circuitry downstream from the basal ganglia. Knowledge gained from studies in NHPs remains critical for our understanding of the pathophysiology of movement disorders, of the effects of DBS on brain network activity, and the development of better treatments for patients with movement disorders and other neurologic or psychiatric conditions.
Pandit, Maharaj K; Grumbine, R Edward
2012-12-01
Indian Himalayan basins are earmarked for widespread dam building, but aggregate effects of these dams on terrestrial ecosystems are unknown. We mapped distribution of 292 dams (under construction and proposed) and projected effects of these dams on terrestrial ecosystems under different scenarios of land-cover loss. We analyzed land-cover data of the Himalayan valleys, where dams are located. We estimated dam density on fifth- through seventh-order rivers and compared these estimates with current global figures. We used a species-area relation model (SAR) to predict short- and long-term species extinctions driven by deforestation. We used scatter plots and correlation studies to analyze distribution patterns of species and dams and to reveal potential overlap between species-rich areas and dam sites. We investigated effects of disturbance on community structure of undisturbed forests. Nearly 90% of Indian Himalayan valleys would be affected by dam building and 27% of these dams would affect dense forests. Our model projected that 54,117 ha of forests would be submerged and 114,361 ha would be damaged by dam-related activities. A dam density of 0.3247/1000 km(2) would be nearly 62 times greater than current average global figures; the average of 1 dam for every 32 km of river channel would be 1.5 times higher than figures reported for U.S. rivers. Our results show that most dams would be located in species-rich areas of the Himalaya. The SAR model projected that by 2025, deforestation due to dam building would likely result in extinction of 22 angiosperm and 7 vertebrate taxa. Disturbance due to dam building would likely reduce tree species richness by 35%, tree density by 42%, and tree basal cover by 30% in dense forests. These results, combined with relatively weak national environmental impact assessment and implementation, point toward significant loss of species if all proposed dams in the Indian Himalaya are constructed. ©2012 Society for Conservation Biology.
Bryson-Morrison, Nicola; Matsuzawa, Tetsuro; Humle, Tatyana
2016-12-01
Many primate populations occur outside protected areas in fragmented anthropogenic landscapes. Empirical data on the ecological characteristics that define an anthropogenic landscape are urgently required if conservation initiatives in such environments are to succeed. The main objective of our study was to determine the composition and availability of chimpanzee (Pan troglodytes verus) food resources across fine spatial scales in the anthropogenic landscape of Bossou, Guinea, West Africa. We examined food resources in all habitat types available in the chimpanzees' core area. We surveyed resource composition, structure and heterogeneity (20 m × 20 m quadrats, N = 54) and assessed temporal availability of food from phenology trails (total distance 5951 m; 1073 individual trees) over 1 year (2012-2013). Over half of Bossou consists of regenerating forest and is highly diverse in terms of chimpanzee food species; large fruit bearing trees are rare and confined to primary and riverine forest. Moraceae (mulberries and figs) was the dominant family, trees of which produce drupaceous fruits favored by chimpanzees. The oil palm occurs at high densities throughout and is the only species found in all habitat types except primary forest. Our data suggest that the high densities of oil palm and fig trees, along with abundant terrestrial herbaceous vegetation and cultivars, are able to provide the chimpanzees with widely available resources, compensating for the scarcity of large fruit trees. A significant difference was found between habitat types in stem density/ha and basal area m 2 /ha of chimpanzee food species. Secondary, young secondary, and primary forest emerged as the most important habitat types for availability of food tree species. Our study emphasizes the importance of examining ecological characteristics of an anthropogenic landscape as each available habitat type is unlikely to be equally important in terms of spatial and temporal availability of resources. Am. J. Primatol. 78:1237-1249, 2016. © 2016 Wiley Periodicals, Inc. © 2016 Wiley Periodicals, Inc.
Estimating soil turnover rate from tree uprooting during hurricanes in Puerto Rico
Lenart, M.T.; Falk, D.A.; Scatena, F.N.; Osterkamp, W.R.
2010-01-01
Soil turnover by tree uprooting in primary and secondary forests on the island of Puerto Rico was measured in 42 study plots in the months immediately after the passage of a Category 3 hurricane. Trunk basal area explained 61% of the variability of mound volume and 53% of the variability of mound area. The proportion of uprooted trees, the number of uprooted trees, or the proportion of uprooted basal area explained 84-85% of the variation in hurricane-created mound area. These same variables explain 79-85% of the variation in mound volume. The study indicates that the soil turnover period from tree uprooting by Puerto Rican hurricanes is between 1600 and 4800 years. These rates are faster than soil turnover by landslides and background treefall in the same area and provide a useful age constraint on soil profile development and soil carbon sequestration in these dynamic landscapes. ?? 2009 Elsevier B.V.
Hasheimi, S R; Zulkifli, I; Somchit, M N; Zunita, Z; Loh, T C; Soleimani, A F; Tang, S C
2013-08-01
The present study was conducted to assess the effects of dietary supplementation of Zingiber officinale and Zingiber zerumbet and to heat-stressed broiler chickens on heat shock protein (HSP) 70 density, plasma corticosterone concentration (CORT), heterophil to lymphocyte ratio (HLR) and body temperature. Beginning from day 28, chicks were divided into five dietary groups: (i) basal diet (control), (ii) basal diet +1%Z. zerumbet powder (ZZ1%), (iii) basal diet +2%Z. zerumbet powder (ZZ2%), (iv) basal diet +1%Z. officinale powder (ZO1%) and (v) basal diet +2%Z. officinale powder (ZO2%). From day 35-42, heat stress was induced by exposing birds to 38±1°C and 80% RH for 2 h/day. Irrespective of diet, heat challenge elevated HSP70 expression, CORT and HLR on day 42. On day 42, following heat challenge, the ZZ1% birds showed lower body temperatures than those of control, ZO1% and ZO2%. Neither CORT nor HLR was significantly affected by diet. The ZO2% and ZZ2% diets enhanced HSP70 expression when compared to the control groups. We concluded that dietary supplementation of Z. officinale and Z. zerumbet powder may induce HSP70 reaction in broiler chickens exposed to heat stress. © 2012 Blackwell Verlag GmbH.
Okamoto, Mayumi; Shinoda, Tomoyasu; Kawaue, Takumi; Nagasaka, Arata; Miyata, Takaki
2014-09-01
The thick outer subventricular zone (OSVZ) is characteristic of the development of human neocortex. How this region originates from the ventricular zone (VZ) is largely unknown. Recently, we showed that over-proliferation-induced acute nuclear densification and thickening of the VZ in neocortical walls of mice, which lack an OSVZ, causes reactive delamination of undifferentiated progenitors and invasion by these cells of basal areas outside the VZ. In this study, we sought to determine how VZ cells behave in non-rodent animals that have an OSVZ. A comparison of mid-embryonic mice and ferrets revealed: (1) the VZ is thicker and more pseudostratified in ferrets. (2) The soma and nuclei of VZ cells were horizontally and apicobasally denser in ferrets. (3) Individual endfeet were also denser on the apical (ventricular) surface in ferrets. (4) In ferrets, apicalward nucleokinesis was less directional, whereas basalward nucleokinesis was more directional; consequently, the nuclear density in the periventricular space (within 16 μm of the apical surface) was smaller in ferrets than in mice, despite the nuclear densification seen basally in ferrets. These results suggest that species-specific differences in nucleokinesis strategies may have evolved in close association with the magnitudes and patterns of nuclear stratification in the VZ. Copyright © 2014 The Authors. Published by Elsevier Ireland Ltd.. All rights reserved.
Gao, Daqiang; Xia, Baorui; Wang, Yanyan; Xiao, Wen; Xi, Pinxian; Xue, Desheng; Ding, Jun
2018-04-01
Although transition metal dichalcogenide MoSe 2 is recognized as one of the low-cost and efficient electrocatalysts for the hydrogen evolution reaction (HER), its thermodynamically stable basal plane and semiconducting property still hamper the electrocatalytic activity. Here, it is demonstrated that the basal plane and edges of 2H-MoSe 2 toward HER can be activated by introducing dual-native vacancy. The first-principle calculations indicate that both the Se and Mo vacancies together activate the electrocatalytic sites in the basal plane and edges of MoSe 2 with the optimal hydrogen adsorption free energy (ΔG H* ) of 0 eV. Experimentally, 2D MoSe 2 nanosheet arrays with a large amount of dual-native vacancies are fabricated as a catalytic working electrode, which possesses an overpotential of 126 mV at a current density of 100 mV cm -2 , a Tafel slope of 38 mV dec -1 , and an excellent long-term durability. The findings pave a rational pathway to trigger the activity of inert MoSe 2 toward HER and also can be extended to other layered dichalcogenide. © 2018 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim.
Yang, Wenjie; Guo, Fengling; Wan, Zhengjie
2013-10-01
Oyster mushroom (Pleurotus ostreatus) was cultivated on rice straw basal substrate, wheat straw basal substrate, cotton seed hull basal substrate, and wheat straw or rice straw supplemented with different proportions (15%, 30%, and 45% in rice straw substrate, 20%, 30%, and 40% in wheat straw substrate) of cotton seed hull to find a cost effective substrate. The effect of autoclaved sterilized and non-sterilized substrate on growth and yield of oyster mushroom was also examined. Results indicated that for both sterilized substrate and non-sterilized substrate, oyster mushroom on rice straw and wheat basal substrate have faster mycelial growth rate, comparatively poor surface mycelial density, shorter total colonization period and days from bag opening to primordia formation, lower yield and biological efficiency, lower mushroom weight, longer stipe length and smaller cap diameter than that on cotton seed hull basal substrate. The addition of cotton seed hull to rice straw and wheat straw substrate slowed spawn running, primordial development and fruit body formation. However, increasing the amount of cotton seed hull can increase the uniformity and white of mycelium, yield and biological efficiency, and increase mushroom weight, enlarge cap diameter and shorten stipe length. Compared to the sterilized substrate, the non-sterilized substrate had comparatively higher mycelial growth rate, shorter total colonization period and days from bag opening to primordia formation. However, the non-sterilized substrate did not gave significantly higher mushroom yield and biological efficiency than the sterilized substrate, but some undesirable characteristics, i.e. smaller mushroom cap diameter and relatively long stipe length.
Anthropogenic effects on a tropical forest according to the distance from human settlements.
Popradit, Ananya; Srisatit, Thares; Kiratiprayoon, Somboon; Yoshimura, Jin; Ishida, Atsushi; Shiyomi, Masae; Murayama, Takehiko; Chantaranothai, Pranom; Outtaranakorn, Somkid; Phromma, Issara
2015-10-05
The protection of tropical forests is one of the most urgent issues in conservation biology because of the rapid deforestation that has occurred over the last 50 years. Even in protected forests, the anthropogenic effects from newly expanding villages such as harvesting of medicinal plants, pasturing cattle and forest fires can induce environmental modifications, especially on the forest floor. We evaluated the anthropogenic effects of the daily activities of neighboring residents on natural forests in 12 plots extending from the village boundary into a natural forest in Thailand. The basal area per unit land area did not present a significant trend; however, the species diversity of woody plants decreased linearly towards the village boundary, which caused a loss of individual density because of severe declines in small saplings compared with adult trees and large saplings in proximity to the village. An analysis of tree-size categories indicates a lack of small samplings near the village boundary. The current forest appears to be well protected based on the adult tree canopy, but regeneration of the present-day forests is unlikely because of the loss of seedlings.
A Microstructure Study on an AZ31 Magnesium Alloy Tube after Hot Metal Gas Forming Process
NASA Astrophysics Data System (ADS)
Liu, Yi; Wu, Xin
2007-06-01
An AZ31 magnesium alloy tube has been deformed by the hot metal gas forming (HMGF) technique. Microstructures before and after deformation have been investigated by using Electron Backscattered Diffraction (EBSD) and Electron Microscopy. Due to the inhomogeneous distribution by induction heating, there is a temperature gradient distribution along the tube axis. Accordingly, the deformation mechanism is also different. In the middle area of deformation zone where the temperature is ˜410 °C, almost no twinning has been found, whereas at the edge areas of deformation zone where the temperature is ˜200 °C, a high density of twins has been found. EBSD experiments show a weak (0001) fiber texture along the radial direction of the tube before and after deformation in the high-temperature zone. EBSD experiments on the low temperature deformation region were not successful due to the high stored energy. Schmid factor analysis on the EBSD data shows that, despite the (0001) fiber texture, there are still many grains favoring basal slip along both the axis direction and hoop direction.
Ashida, Terunao; Takato, Tetsuya; Matsuzaki, Gen; Seko, Yoshinori; Fujii, Jun; Kawai, Sachio
2014-01-01
We have recently demonstrated that basal cardiomyopathy develops in rabbits with ventricular tachyarrhythmias that have been induced by electrical stimulation of the cervical vagus. This study investigated whether similar basal cardiomyopathy would develop in rabbits with ventricular tachyarrhythmias induced by a single injection of adrenaline. Adrenaline was intravenously infused for 10-360 seconds in anesthetized rabbits. Colloidal carbon was injected after adrenaline infusion. Wall movement velocity of the left ventricular base was assessed by tissue Doppler echocardiography. Animals were killed either 1 week or 3-4 weeks later. Pathological lesions were identified by deposits of carbon particles. Animals were divided into two groups according to the infused dose of adrenaline. The small-dose group (group S, n = 15) received 1-10 μg and the large-dose group (group L, n = 23) received 15-60 μg of adrenaline. Adrenaline infusion induced premature ventricular contractions followed by monomorphic ventricular tachycardias in 22 of 23 animals in group L, but in only 1 of 15 animals in group S. Wall movement velocity of the left ventricular base decreased just after adrenaline infusion, remained low after 1 week, and recovered to near-baseline levels after 3-4 weeks in group L. Unique cardiac lesions identified by deposits of carbon particles were frequently observed on the left ventricular basal portion, almost always associated with the mitral valve and papillary muscles, but were never observed in the apical area. Lesions involving all areas of the left ventricular basal portion were observed in 22 of 23 animals in group L, but in only 2 of 15 animals in group S. Basal cardiomyopathy developed in rabbits with ventricular tachycardias induced by a single injection of adrenaline.
Emergence of context-dependent variability across a basal ganglia network.
Woolley, Sarah C; Rajan, Raghav; Joshua, Mati; Doupe, Allison J
2014-04-02
Context dependence is a key feature of cortical-basal ganglia circuit activity, and in songbirds the cortical outflow of a basal ganglia circuit specialized for song, LMAN, shows striking increases in trial-by-trial variability and bursting when birds sing alone rather than to females. To reveal where this variability and its social regulation emerge, we recorded stepwise from corticostriatal (HVC) neurons and their target spiny and pallidal neurons in Area X. We find that corticostriatal and spiny neurons both show precise singing-related firing across both social settings. Pallidal neurons, in contrast, exhibit markedly increased trial-by-trial variation when birds sing alone, created by highly variable pauses in firing. This variability persists even when recurrent inputs from LMAN are ablated. These data indicate that variability and its context sensitivity emerge within the basal ganglia network, suggest a network mechanism for this emergence, and highlight variability generation and regulation as basal ganglia functions. Copyright © 2014 Elsevier Inc. All rights reserved.
Emergence of context-dependent variability across a basal ganglia network
Woolley, Sarah C.; Rajan, Raghav; Joshua, Mati; Doupe, Allison J.
2014-01-01
Summary Context-dependence is a key feature of cortical-basal ganglia circuit activity, and in songbirds, the cortical outflow of a basal ganglia circuit specialized for song, LMAN, shows striking increases in trial-by-trial variability and bursting when birds sing alone rather than to females. To reveal where this variability and its social regulation emerge, we recorded stepwise from cortico-striatal (HVC) neurons and their target spiny and pallidal neurons in Area X. We find that cortico-striatal and spiny neurons both show precise singing-related firing across both social settings. Pallidal neurons, in contrast, exhibit markedly increased trial-by-trial variation when birds sing alone, created by highly variable pauses in firing. This variability persists even when recurrent inputs from LMAN are ablated. These data indicate that variability and its context-sensitivity emerge within the basal ganglia network, suggest a network mechanism for this emergence, and highlight variability generation and regulation as basal ganglia functions. PMID:24698276
Dermatoscopy-guided therapy of pigmented basal cell carcinoma with imiquimod*
Husein-ElAhmed, Husein; Fernandez-Pugnaire, Maria Antonia
2016-01-01
BACKGROUND Dermatoscopy is a non-invasive diagnostic tool used to examine skin lesions with an optical magnification. It has been suggested as a useful tool for monitoring therapeutic response in lentigo maligna patients treated with imiquimod. OBJECTIVE To examine the accuracy of dermatoscopy as a tool to monitor the therapeutic response of pigmented basal cell carcinoma treated with imiquimod. METHOD The authors designed a prospective study. Patients with pigmented basal cell carcinoma were included and data regarding the dermatoscopy features were collected following the Menzies criteria, prior to initiating the imiquimod treatment. Subsequent dermatoscopic evaluations were performed at weeks 4 and 8, following imiquimod discontinuation. RESULTS Twenty lesions were included. The most common pigmented dermatoscopy features were large blue-grey ovoid nests (80%), followed by blue-grey globules (50%) and leaf-like areas (30%). No spoke wheel areas were observed. In 17 out of 20 patients, a response was noted during the first evaluation at 4 weeks, while the clearance was noted at the second check-up after 8 weeks. In two patients, the clearance was found at the initial evaluation at 4 weeks, while in one patient, the response remained unchanged. Blue-grey globules were the fastest to exhibit clearance (50% at week 4), followed by leaf-like areas (15%) and large blue-grey ovoid nests (6.25%). CONCLUSION According to our results, dermatoscopic evaluation enhances the accuracy in the assessment of the clinical response to imiquimod in pigmented basal cell carcinoma. PMID:28099598
Listening to Rhythmic Music Reduces Connectivity within the Basal Ganglia and the Reward System.
Brodal, Hans P; Osnes, Berge; Specht, Karsten
2017-01-01
Music can trigger emotional responses in a more direct way than any other stimulus. In particular, music-evoked pleasure involves brain networks that are part of the reward system. Furthermore, rhythmic music stimulates the basal ganglia and may trigger involuntary movements to the beat. In the present study, we created a continuously playing rhythmic, dance floor-like composition where the ambient noise from the MR scanner was incorporated as an additional instrument of rhythm. By treating this continuous stimulation paradigm as a variant of resting-state, the data was analyzed with stochastic dynamic causal modeling (sDCM), which was used for exploring functional dependencies and interactions between core areas of auditory perception, rhythm processing, and reward processing. The sDCM model was a fully connected model with the following areas: auditory cortex, putamen/pallidum, and ventral striatum/nucleus accumbens of both hemispheres. The resulting estimated parameters were compared to ordinary resting-state data, without an additional continuous stimulation. Besides reduced connectivity within the basal ganglia, the results indicated a reduced functional connectivity of the reward system, namely the right ventral striatum/nucleus accumbens from and to the basal ganglia and auditory network while listening to rhythmic music. In addition, the right ventral striatum/nucleus accumbens demonstrated also a change in its hemodynamic parameter, reflecting an increased level of activation. These converging results may indicate that the dopaminergic reward system reduces its functional connectivity and relinquishing its constraints on other areas when we listen to rhythmic music.
Listening to Rhythmic Music Reduces Connectivity within the Basal Ganglia and the Reward System
Brodal, Hans P.; Osnes, Berge; Specht, Karsten
2017-01-01
Music can trigger emotional responses in a more direct way than any other stimulus. In particular, music-evoked pleasure involves brain networks that are part of the reward system. Furthermore, rhythmic music stimulates the basal ganglia and may trigger involuntary movements to the beat. In the present study, we created a continuously playing rhythmic, dance floor-like composition where the ambient noise from the MR scanner was incorporated as an additional instrument of rhythm. By treating this continuous stimulation paradigm as a variant of resting-state, the data was analyzed with stochastic dynamic causal modeling (sDCM), which was used for exploring functional dependencies and interactions between core areas of auditory perception, rhythm processing, and reward processing. The sDCM model was a fully connected model with the following areas: auditory cortex, putamen/pallidum, and ventral striatum/nucleus accumbens of both hemispheres. The resulting estimated parameters were compared to ordinary resting-state data, without an additional continuous stimulation. Besides reduced connectivity within the basal ganglia, the results indicated a reduced functional connectivity of the reward system, namely the right ventral striatum/nucleus accumbens from and to the basal ganglia and auditory network while listening to rhythmic music. In addition, the right ventral striatum/nucleus accumbens demonstrated also a change in its hemodynamic parameter, reflecting an increased level of activation. These converging results may indicate that the dopaminergic reward system reduces its functional connectivity and relinquishing its constraints on other areas when we listen to rhythmic music. PMID:28400717
Availability of streamflow for recharge of the basal aquifer in the Pearl Harbor area, Hawaii
Hirashima, George Tokusuke
1971-01-01
The Pearl Harbor area is underlain by an extensive basal aquifer that contains large supplies of fresh water. Because of the presence of a cap rock composed of sedimentary material that is less permeable than the basaltic lava of the basal aquifer, seaward movement of ground water is retarded. The cap rock causes the basal water to stand at a high level; thus, the lens of fresh water that floats on sea water is thick. Discharge from the basal ground-water body, which includes pumpage from wells and shafts, averaged 250 million gallons per day during 1931-65. Because the water level in the basal aquifer did not decline progressively, recharge to the ground-water body must have been approximately equal to discharge. Although pumping for agricultural use has decreased since 1931, net ground-water discharge has increased because of a large increase in pumping for urban use. Substitution of ground water for surface water in the irrigation of sugarcane has also contributed to a net increase in ground-water discharge. The development of Mililani Town will further increase discharge. The increase in ground-water discharge may cause an increase in chloride content of the water pumped from wells near the shore of Pearl Harbor unless the increased discharge is balanced by increased recharge to the local aquifer. The aquifer is recharged by direct infiltration and deep percolation of rain, principally in the high forested area, by infiltration and percolation of irrigation water applied in excess of plant requirements, by seepage of water through streambeds, and possibly by ground-water inflow from outside the area. Recharge is greatest in the uplands, where rainfall is heavy and where much infiltration takes place before rainwater collects in the middle and lower reaches of stream channels. Once water collects in and saturates the alluvium of stream channels, additional inflow to the streams will flow out to sea, only slightly decreased by seepage. Average annual direct runoff from the 90-square-mile Pearl Harbor area is 47.27 million gallons per day, or 11.1 inches; this is 13.3 percent of the average annual rainfall (83.3 in.) over the area. Average annual direct runoff in streams at the 800- and 400-foot altitudes is 29 and 38 million gallons per day, respectively. Kipapa Stream has the largest average annual direct runoff at those altitudes--6 and 9 million gallons per day, respectively. Because streams are flashy and have a wide range in discharge, only 60 percent of the average annual runoff can be economically diverted through ditches to recharge areas. The diversion may be increased slightly if reservoirs are used in conjunction with ditches to temporarily detain flows in excess of ditch capacity. The planned irrigation use of some of the perennial flow available in Waikele Stream near sea level will decrease pumping from and increase recharge to the basal aquifer. Suspended-sediment load is mainly silt and clay, and it increases rapidly with increased discharge. Thus, the use of streamflow for artificial recharge poses problems. High flows must be used if recharge is to be effective, but flows must not be so high as to cause clogging of recharge facilities with sediment or woodland debris. Practical tests are needed to determine the advantages and disadvantages of different types of recharge structures, such as a reservoir or basin, large-diameter deep shafts, deep wells, or combinations of all these structures.
A synthesis of the basal thermal state of the Greenland Ice Sheet
MacGregor, Joseph A.; Fahnestock, Mark A.; Catania, Ginny A.; Aschwanden, Andy; Clow, Gary D.; Colgan, William T.; Gogineni, S. Prasad; Morlighem, Mathieu; Nowicki, Sophie M. J.; Paden, John D.; Price, Stephen F.; Seroussi, Hélène
2017-01-01
The basal thermal state of an ice sheet (frozen or thawed) is an important control upon its evolution, dynamics and response to external forcings. However, this state can only be observed directly within sparse boreholes or inferred conclusively from the presence of subglacial lakes. Here we synthesize spatially extensive inferences of the basal thermal state of the Greenland Ice Sheet to better constrain this state. Existing inferences include outputs from the eight thermomechanical ice-flow models included in the SeaRISE effort. New remote-sensing inferences of the basal thermal state are derived from Holocene radiostratigraphy, modern surface velocity and MODIS imagery. Both thermomechanical modeling and remote inferences generally agree that the Northeast Greenland Ice Stream and large portions of the southwestern ice-drainage systems are thawed at the bed, whereas the bed beneath the central ice divides, particularly their west-facing slopes, is frozen. Elsewhere, there is poor agreement regarding the basal thermal state. Both models and remote inferences rarely represent the borehole-observed basal thermal state accurately near NorthGRIP and DYE-3. This synthesis identifies a large portion of the Greenland Ice Sheet (about one third by area) where additional observations would most improve knowledge of its overall basal thermal state. PMID:28163988
A synthesis of the basal thermal state of the Greenland Ice Sheet.
MacGregor, Joseph A; Fahnestock, Mark A; Catania, Ginny A; Aschwanden, Andy; Clow, Gary D; Colgan, William T; Gogineni, S Prasad; Morlighem, Mathieu; Nowicki, Sophie M J; Paden, John D; Price, Stephen F; Seroussi, Hélène
2016-08-10
The basal thermal state of an ice sheet (frozen or thawed) is an important control upon its evolution, dynamics and response to external forcings. However, this state can only be observed directly within sparse boreholes or inferred conclusively from the presence of subglacial lakes. Here we synthesize spatially extensive inferences of the basal thermal state of the Greenland Ice Sheet to better constrain this state. Existing inferences include outputs from the eight thermomechanical ice-flow models included in the SeaRISE effort. New remote-sensing inferences of the basal thermal state are derived from Holocene radiostratigraphy, modern surface velocity and MODIS imagery. Both thermomechanical modeling and remote inferences generally agree that the Northeast Greenland Ice Stream and large portions of the southwestern ice-drainage systems are thawed at the bed, whereas the bed beneath the central ice divides, particularly their west-facing slopes, is frozen. Elsewhere, there is poor agreement regarding the basal thermal state. Both models and remote inferences rarely represent the borehole-observed basal thermal state accurately near NorthGRIP and DYE-3. This synthesis identifies a large portion of the Greenland Ice Sheet (about one third by area) where additional observations would most improve knowledge of its overall basal thermal state.
A synthesis of the basal thermal state of the Greenland Ice Sheet
MacGregor, Joseph A; Fahnestock, Mark A; Catania, Ginny A; Aschwanden, Andy; Clow, Gary D.; Colgan, William T.; Gogineni, Prasad S.; Morlighem, Mathieu; Nowicki, Sophie M .J.; Paden, John D; Price, Stephen F.; Seroussi, Helene
2016-01-01
The basal thermal state of an ice sheet (frozen or thawed) is an important control upon its evolution, dynamics and response to external forcings. However, this state can only be observed directly within sparse boreholes or inferred conclusively from the presence of subglacial lakes. Here we synthesize spatially extensive inferences of the basal thermal state of the Greenland Ice Sheet to better constrain this state. Existing inferences include outputs from the eight thermomechanical ice-flow models included in the SeaRISE effort. New remote-sensing inferences of the basal thermal state are derived from Holocene radiostratigraphy, modern surface velocity and MODIS imagery. Both thermomechanical modeling and remote inferences generally agree that the Northeast Greenland Ice Stream and large portions of the southwestern ice-drainage systems are thawed at the bed, whereas the bed beneath the central ice divides, particularly their west-facing slopes, is frozen. Elsewhere, there is poor agreement regarding the basal thermal state. Both models and remote inferences rarely represent the borehole-observed basal thermal state accurately near NorthGRIP and DYE-3. This synthesis identifies a large portion of the Greenland Ice Sheet (about one third by area) where additional observations would most improve knowledge of its overall basal thermal state.
The Effect of Autologous Platelet Lysate Eye Drops: An In Vivo Confocal Microscopy Study
Fea, Antonio M.; Testa, Valeria; Machetta, Federica; Parisi, Simone; D'Antico, Sergio; Spinetta, Roberta; Fusaro, Enrico; Grignolo, Federico M.
2016-01-01
Purpose. To determine the effectiveness of autologous platelet lysate (APL) eye drops in patients with primary Sjögren syndrome (SS) dry eye, refractory to standard therapy, in comparison with patients treated with artificial tears. We focused on the effect of APL on cornea morphology with the in vivo confocal microscopy (IVCM). Methods. Patients were assigned to two groups: group A used autologous platelet lysate QID, and group B used preservative-free artificial tears QID, for 90 days. Ophthalmological assessments included ocular surface disease index (OSDI), best corrected visual acuity (BCVA), Schirmer test, fluorescein score, and breakup time (BUT). A subgroup of patients in group A underwent IVCM: corneal basal epithelium, subbasal nerves, Langerhans cells, anterior stroma activated keratocytes, and reflectivity were evaluated. Results. 60 eyes of 30 patients were enrolled; in group A (n = 20 patients) mean OSDI, fluorescein score, and BUT showed significant improvement compared with group B (n = 10 patients). The IVCM showed a significant increase in basal epithelium cells density and subbasal nerve plexus density and number and a decrease in Langerhans cells density (p < 0.05). Conclusion. APL was found effective in the treatment of SS dry eye. IVCM seems to be a useful tool to visualize cornea morphologic modifications. PMID:27200376
The Effect of Autologous Platelet Lysate Eye Drops: An In Vivo Confocal Microscopy Study.
Fea, Antonio M; Aragno, Vittoria; Testa, Valeria; Machetta, Federica; Parisi, Simone; D'Antico, Sergio; Spinetta, Roberta; Fusaro, Enrico; Grignolo, Federico M
2016-01-01
Purpose. To determine the effectiveness of autologous platelet lysate (APL) eye drops in patients with primary Sjögren syndrome (SS) dry eye, refractory to standard therapy, in comparison with patients treated with artificial tears. We focused on the effect of APL on cornea morphology with the in vivo confocal microscopy (IVCM). Methods. Patients were assigned to two groups: group A used autologous platelet lysate QID, and group B used preservative-free artificial tears QID, for 90 days. Ophthalmological assessments included ocular surface disease index (OSDI), best corrected visual acuity (BCVA), Schirmer test, fluorescein score, and breakup time (BUT). A subgroup of patients in group A underwent IVCM: corneal basal epithelium, subbasal nerves, Langerhans cells, anterior stroma activated keratocytes, and reflectivity were evaluated. Results. 60 eyes of 30 patients were enrolled; in group A (n = 20 patients) mean OSDI, fluorescein score, and BUT showed significant improvement compared with group B (n = 10 patients). The IVCM showed a significant increase in basal epithelium cells density and subbasal nerve plexus density and number and a decrease in Langerhans cells density (p < 0.05). Conclusion. APL was found effective in the treatment of SS dry eye. IVCM seems to be a useful tool to visualize cornea morphologic modifications.
Microstructures and mechanical behavior of magnesium processed by ECAP at ice-water temperature
NASA Astrophysics Data System (ADS)
Zuo, Dai; Li, Taotao; Liang, Wei; Wen, Xiyu; Yang, Fuqian
2018-05-01
Magnesium of high purity is processed by equal channel angular pressing (ECAP) up to eight passes at the ice-water temperature, in which a core–shell-like structure is used. The core–shell-like structure consists of pure iron (Fe) of 1.5 mm in thickness as the shell and magnesium (Mg) as the core. The microstructure, texture and mechanical behavior of the ECAP-processed Mg are studied. The ECAP processing leads to the formation of fine and equiaxed grains of ~1.1 µm. The basal planes initially parallel to the extrusion direction evolve to slanted basal planes with the tilting angle in a range of 25°–45° to the extrusion direction. Increasing the number of the extrusion passes leads to the decreasing of twins and dislocation density in grains, while individual grains after eight passes still have high dislocation density. The large decreases of twins and the dislocation density make dynamic recrystallization (DRX) difficult, resulting in the decrease of the degree of DRX. Tension test reveals that the mechanical behavior of the ECAP-processed Mg is dependent on grain refinement and textures. The yield strength of the ECAP-extruded Mg first increases with the decrease of the grain size, and then decreases with further decrease of the grain size.
Clark, David B.; Hurtado, Johanna; Saatchi, Sassan S.
2015-01-01
Rapid biological changes are expected to occur on tropical elevational gradients as species migrate upslope or go extinct in the face of global warming. We established a series of 9 1-ha plots in old-growth tropical rainforest in Costa Rica along a 2700 m relief elevational gradient to carry out long-term monitoring of tropical rain forest structure, dynamics and tree growth. Within each plot we mapped, identified, and annually measured diameter for all woody individuals with stem diameters >10 cm for periods of 3-10 years. Wood species diversity peaked at 400-600 m and decreased substantially at higher elevations. Basal area and stem number varied by less than two-fold, with the exception of the 2800 m cloud forest summit, where basal area and stem number were approximately double that of lower sites. Canopy gaps extending to the forest floor accounted for <3% of microsites at all elevations. Height of highest crowns and the coefficient of variation of crown height both decreased with increasing elevation. Rates of turnover of individuals and of stand basal area decreased with elevation, but rates of diameter growth and stand basal area showed no simple relation to elevation. We discuss issues encountered in the design and implementation of this network of plots, including biased sampling, missing key meteorological and biomass data, and strategies for improving species-level research. Taking full advantage of the major research potential of tropical forest elevational transects will require sustaining and extending ground based studies, incorporation of new remotely-sensed data and data-acquisition platforms, and new funding models to support decadal research on these rapidly-changing systems. PMID:25856163
The Importance of Large-Diameter Trees to Forest Structural Heterogeneity
Lutz, James A.; Larson, Andrew J.; Freund, James A.; Swanson, Mark E.; Bible, Kenneth J.
2013-01-01
Large-diameter trees dominate the structure, dynamics and function of many temperate and tropical forests. However, their attendant contributions to forest heterogeneity are rarely addressed. We established the Wind River Forest Dynamics Plot, a 25.6 ha permanent plot within which we tagged and mapped all 30,973 woody stems ≥1 cm dbh, all 1,966 snags ≥10 cm dbh, and all shrub patches ≥2 m2. Basal area of the 26 woody species was 62.18 m2/ha, of which 61.60 m2/ha was trees and 0.58 m2/ha was tall shrubs. Large-diameter trees (≥100 cm dbh) comprised 1.5% of stems, 31.8% of basal area, and 17.6% of the heterogeneity of basal area, with basal area dominated by Tsuga heterophylla and Pseudotsuga menziesii. Small-diameter subpopulations of Pseudotsuga menziesii, Tsuga heterophylla and Thuja plicata, as well as all tree species combined, exhibited significant aggregation relative to the null model of complete spatial randomness (CSR) up to 9 m (P≤0.001). Patterns of large-diameter trees were either not different from CSR (Tsuga heterophylla), or exhibited slight aggregation (Pseudotsuga menziesii and Thuja plicata). Significant spatial repulsion between large-diameter and small-diameter Tsuga heterophylla suggests that large-diameter Tsuga heterophylla function as organizers of tree demography over decadal timescales through competitive interactions. Comparison among two forest dynamics plots suggests that forest structural diversity responds to intermediate-scale environmental heterogeneity and disturbances, similar to hypotheses about patterns of species richness, and richness- ecosystem function. Large mapped plots with detailed within-plot environmental spatial covariates will be required to test these hypotheses. PMID:24376579
The importance of large-diameter trees to forest structural heterogeneity.
Lutz, James A; Larson, Andrew J; Freund, James A; Swanson, Mark E; Bible, Kenneth J
2013-01-01
Large-diameter trees dominate the structure, dynamics and function of many temperate and tropical forests. However, their attendant contributions to forest heterogeneity are rarely addressed. We established the Wind River Forest Dynamics Plot, a 25.6 ha permanent plot within which we tagged and mapped all 30,973 woody stems ≥ 1 cm dbh, all 1,966 snags ≥ 10 cm dbh, and all shrub patches ≥ 2 m(2). Basal area of the 26 woody species was 62.18 m(2)/ha, of which 61.60 m(2)/ha was trees and 0.58 m(2)/ha was tall shrubs. Large-diameter trees (≥ 100 cm dbh) comprised 1.5% of stems, 31.8% of basal area, and 17.6% of the heterogeneity of basal area, with basal area dominated by Tsuga heterophylla and Pseudotsuga menziesii. Small-diameter subpopulations of Pseudotsuga menziesii, Tsuga heterophylla and Thuja plicata, as well as all tree species combined, exhibited significant aggregation relative to the null model of complete spatial randomness (CSR) up to 9 m (P ≤ 0.001). Patterns of large-diameter trees were either not different from CSR (Tsuga heterophylla), or exhibited slight aggregation (Pseudotsuga menziesii and Thuja plicata). Significant spatial repulsion between large-diameter and small-diameter Tsuga heterophylla suggests that large-diameter Tsuga heterophylla function as organizers of tree demography over decadal timescales through competitive interactions. Comparison among two forest dynamics plots suggests that forest structural diversity responds to intermediate-scale environmental heterogeneity and disturbances, similar to hypotheses about patterns of species richness, and richness- ecosystem function. Large mapped plots with detailed within-plot environmental spatial covariates will be required to test these hypotheses.
Above-ground biomass and structure of 260 African tropical forests
Lewis, Simon L.; Sonké, Bonaventure; Sunderland, Terry; Begne, Serge K.; Lopez-Gonzalez, Gabriela; van der Heijden, Geertje M. F.; Phillips, Oliver L.; Affum-Baffoe, Kofi; Baker, Timothy R.; Banin, Lindsay; Bastin, Jean-François; Beeckman, Hans; Boeckx, Pascal; Bogaert, Jan; De Cannière, Charles; Chezeaux, Eric; Clark, Connie J.; Collins, Murray; Djagbletey, Gloria; Djuikouo, Marie Noël K.; Droissart, Vincent; Doucet, Jean-Louis; Ewango, Cornielle E. N.; Fauset, Sophie; Feldpausch, Ted R.; Foli, Ernest G.; Gillet, Jean-François; Hamilton, Alan C.; Harris, David J.; Hart, Terese B.; de Haulleville, Thales; Hladik, Annette; Hufkens, Koen; Huygens, Dries; Jeanmart, Philippe; Jeffery, Kathryn J.; Kearsley, Elizabeth; Leal, Miguel E.; Lloyd, Jon; Lovett, Jon C.; Makana, Jean-Remy; Malhi, Yadvinder; Marshall, Andrew R.; Ojo, Lucas; Peh, Kelvin S.-H.; Pickavance, Georgia; Poulsen, John R.; Reitsma, Jan M.; Sheil, Douglas; Simo, Murielle; Steppe, Kathy; Taedoumg, Hermann E.; Talbot, Joey; Taplin, James R. D.; Taylor, David; Thomas, Sean C.; Toirambe, Benjamin; Verbeeck, Hans; Vleminckx, Jason; White, Lee J. T.; Willcock, Simon; Woell, Hannsjorg; Zemagho, Lise
2013-01-01
We report above-ground biomass (AGB), basal area, stem density and wood mass density estimates from 260 sample plots (mean size: 1.2 ha) in intact closed-canopy tropical forests across 12 African countries. Mean AGB is 395.7 Mg dry mass ha−1 (95% CI: 14.3), substantially higher than Amazonian values, with the Congo Basin and contiguous forest region attaining AGB values (429 Mg ha−1) similar to those of Bornean forests, and significantly greater than East or West African forests. AGB therefore appears generally higher in palaeo- compared with neotropical forests. However, mean stem density is low (426 ± 11 stems ha−1 greater than or equal to 100 mm diameter) compared with both Amazonian and Bornean forests (cf. approx. 600) and is the signature structural feature of African tropical forests. While spatial autocorrelation complicates analyses, AGB shows a positive relationship with rainfall in the driest nine months of the year, and an opposite association with the wettest three months of the year; a negative relationship with temperature; positive relationship with clay-rich soils; and negative relationships with C : N ratio (suggesting a positive soil phosphorus–AGB relationship), and soil fertility computed as the sum of base cations. The results indicate that AGB is mediated by both climate and soils, and suggest that the AGB of African closed-canopy tropical forests may be particularly sensitive to future precipitation and temperature changes. PMID:23878327
2015-01-01
Non- and semipolar GaN have great potential to improve the efficiency of light emitting devices due to much reduced internal electric fields. However, heteroepitaxial GaN growth in these crystal orientations suffers from very high dislocation and stacking faults densities. Here, we report a facile method to obtain low defect density non- and semipolar heteroepitaxial GaN via selective area epitaxy using self-assembled multilayers of silica nanospheres (MSN). Nonpolar (11–20) and semipolar (11–22) GaN layers with high crystal quality have been achieved by epitaxial integration of the MSN and a simple one-step overgrowth process, by which both dislocation and basal plane stacking fault densities can be significantly reduced. The underlying defect reduction mechanisms include epitaxial growth through the MSN covered template, island nucleation via nanogaps in the MSN, and lateral overgrowth and coalescence above the MSN. InGaN/GaN multiple quantum wells structures grown on a nonpolar GaN/MSN template show more than 30-fold increase in the luminescence intensity compared to a control sample without the MSN. This self-assembled MSN technique provides a new platform for epitaxial growth of nitride semiconductors and offers unique opportunities for improving the material quality of GaN grown on other orientations and foreign substrates or heteroepitaxial growth of other lattice-mismatched materials. PMID:27065755
Brianza-Padilla, Malinalli; Bonilla-Jaime, Herlinda; Almanza-Pérez, Julio César; López-López, Ana Laura; Sánchez-Muñoz, Fausto; Vázquez-Palacios, Gonzalo
2016-03-01
Sleep has a fundamental role in the regulation of energy balance, and it is an essential and natural process whose precise impacts on health and disease have not yet been fully elucidated. The aim of this study was to assess the consequences of different periods of paradoxical sleep deprivation (PSD) and recovery from PSD on lipid profile, oral glucose tolerance test (OGTT) results, and changes in insulin, corticosterone, ghrelin, and leptin concentrations. Three-month-old male Wistar rats weighing 250-350 g were submitted to 24, 96, or 192 h of PSD or 192 h of PSD with 480 h of recovery. The PSD was induced by the multiple platforms method. Subsequently, the animals were submitted to an OGTT. One day later, the animals were killed and the levels of triglycerides, total cholesterol, lipoproteins (low-density lipoprotein, very-low-density lipoprotein, and high-density lipoprotein), insulin, ghrelin, leptin, and corticosterone in plasma were quantified. There was a progressive decrease in body weight with increasing duration of PSD. The PSD induced basal hypoglycemia over all time periods evaluated. Evaluation of areas under the curve revealed progressive hypoglycemia only after 96 and 192 h of PSD. There was an increase in corticosterone levels after 192 h of PSD. We conclude that PSD induces alterations in metabolism that are reversed after a recovery period of 20 days.
Relating P-band AIRSAR backscatter to forest stand parameters
NASA Technical Reports Server (NTRS)
Wang, Yong; Melack, John M.; Davis, Frank W.; Kasischke, Eric S.; Christensen, Norman L., Jr.
1993-01-01
As part of research on forest ecosystems, the Jet Propulsion Laboratory (JPL) and collaborating research teams have conducted multi-season airborne synthetic aperture radar (AIRSAR) experiments in three forest ecosystems including temperate pine forest (Duke, Forest, North Carolina), boreal forest (Bonanza Creek Experimental Forest, Alaska), and northern mixed hardwood-conifer forest (Michigan Biological Station, Michigan). The major research goals were to improve understanding of the relationships between radar backscatter and phenological variables (e.g. stand density, tree size, etc.), to improve radar backscatter models of tree canopy properties, and to develop a radar-based scheme for monitoring forest phenological changes. In September 1989, AIRSAR backscatter data were acquired over the Duke Forest. As the aboveground biomass of the loblolly pine forest stands at Duke Forest increased, the SAR backscatter at C-, L-, and P-bands increased and saturated at different biomass levels for the C-band, L-band, and P-band data. We only use the P-band backscatter data and ground measurements here to study the relationships between the backscatter and stand density, the backscatter and mean trunk dbh (diameter at breast height) of trees in the stands, and the backscatter and stand basal area.
Widespread Moulin Formation During Supraglacial Lake Drainages in Greenland
DOE Office of Scientific and Technical Information (OSTI.GOV)
Hoffman, Matthew J.; Perego, Mauro; Andrews, Lauren C.
Moulins permit access of surface meltwater to the glacier bed, causing basal lubrication and ice speedup in the ablation zone of western Greenland during summer. In spite of the substantial impact of moulins on ice dynamics, the conditions under which they form are poorly understood. We assimilate a time series of ice surface velocity from a network of eleven Global Positioning System receivers into an ice sheet model to estimate ice sheet stresses during winter, spring, and summer in a ~30 × 10 km region. Surface-parallel von Mises stress increases slightly during spring speedup and early summer, sufficient to allowmore » formation of 16% of moulins mapped in the study area. Conversely, 63% of moulins experience stresses over the tensile strength of ice during a short (hours) supraglacial lake drainage event. Lake drainages appear to control moulin density, which is itself a control on subglacial drainage efficiency and summer ice velocities.« less
Widespread Moulin Formation During Supraglacial Lake Drainages in Greenland
Hoffman, Matthew J.; Perego, Mauro; Andrews, Lauren C.; ...
2018-01-17
Moulins permit access of surface meltwater to the glacier bed, causing basal lubrication and ice speedup in the ablation zone of western Greenland during summer. In spite of the substantial impact of moulins on ice dynamics, the conditions under which they form are poorly understood. We assimilate a time series of ice surface velocity from a network of eleven Global Positioning System receivers into an ice sheet model to estimate ice sheet stresses during winter, spring, and summer in a ~30 × 10 km region. Surface-parallel von Mises stress increases slightly during spring speedup and early summer, sufficient to allowmore » formation of 16% of moulins mapped in the study area. Conversely, 63% of moulins experience stresses over the tensile strength of ice during a short (hours) supraglacial lake drainage event. Lake drainages appear to control moulin density, which is itself a control on subglacial drainage efficiency and summer ice velocities.« less
Widespread Moulin Formation During Supraglacial Lake Drainages in Greenland
NASA Astrophysics Data System (ADS)
Hoffman, Matthew J.; Perego, Mauro; Andrews, Lauren C.; Price, Stephen F.; Neumann, Thomas A.; Johnson, Jesse V.; Catania, Ginny; Lüthi, Martin P.
2018-01-01
Moulins permit access of surface meltwater to the glacier bed, causing basal lubrication and ice speedup in the ablation zone of western Greenland during summer. Despite the substantial impact of moulins on ice dynamics, the conditions under which they form are poorly understood. We assimilate a time series of ice surface velocity from a network of eleven Global Positioning System receivers into an ice sheet model to estimate ice sheet stresses during winter, spring, and summer in a ˜30 × 10 km region. Surface-parallel von Mises stress increases slightly during spring speedup and early summer, sufficient to allow formation of 16% of moulins mapped in the study area. In contrast, 63% of moulins experience stresses over the tensile strength of ice during a short (hours) supraglacial lake drainage event. Lake drainages appear to control moulin density, which is itself a control on subglacial drainage efficiency and summer ice velocities.
NASA Technical Reports Server (NTRS)
Rich, Paul M.; Fournier, Robert; Hall, Forrest G. (Editor); Papagno, Andrea (Editor)
2000-01-01
The Boreal Ecosystem-Atmospheric Study (BOREAS) TE-23 (Terrestrial Ecology) team collected map plot data in support of its efforts to characterize and interpret information on canopy architecture and understory cover at the BOREAS tower flux sites and selected auxiliary sites from May to August 1994. Mapped plots (typical dimensions 50 m x 60 m) were set up and characterized at all BOREAS forested tower flux and selected auxiliary sites. Detailed measurement of the mapped plots included: (1) stand characteristics (location, density, basal area); (2) map locations diameter at breast height (DBH) of all trees; (3) detailed geometric measures of a subset of trees (height, crown dimensions); and (4) understory cover maps. The data are stored in tabular ASCII files. The data files are available on a CD-ROM (see document number 20010000884), or from the Oak Ridge National Laboratory (ORNL) Distributed Active Archive Center (DAAC).
Electrochemical Ultracapacitors Using Graphitic Nanostacks
NASA Technical Reports Server (NTRS)
Marotta, Christopher
2012-01-01
Electrochemical ultracapacitors (ECs) have been developed using graphitic nanostacks as the electrode material. The advantages of this technology will be the reduction of device size due to superior power densities and relative powers compared to traditional activated carbon electrodes. External testing showed that these materials display reduced discharge response times compared to state-of-the-art materials. Such applications are advantageous for pulsed power applications such as burst communications (satellites, cell phones), electromechanical actuators, and battery load leveling in electric vehicles. These carbon nanostructures are highly conductive and offer an ordered mesopore network. These attributes will provide more complete electrolyte wetting, and faster release of stored charge compared to activated carbon. Electrochemical capacitor (EC) electrode materials were developed using commercially available nanomaterials and modifying them to exploit their energy storage properties. These materials would be an improvement over current ECs that employ activated carbon as the electrode material. Commercially available graphite nanofibers (GNFs) are used as precursor materials for the synthesis of graphitic nanostacks (GNSs). These materials offer much greater surface area than graphite flakes. Additionally, these materials offer a superior electrical conductivity and a greater average pore size compared to activated carbon electrodes. The state of the art in EC development uses activated carbon (AC) as the electrode material. AC has a high surface area, but its small average pore size inhibits electrolyte ingress/egress. Additionally, AC has a higher resistivity, which generates parasitic heating in high-power applications. This work focuses on fabricating EC from carbon that has a very different structure by increasing the surface area of the GNF by intercalation or exfoliation of the graphitic basal planes. Additionally, various functionalities to the GNS surface will be added that can exhibit pseudocapacitance. This pseudocapacitance exhibits faradaic (charge transfer) properties that can further increase the overall relative and volumetric capacitance of the material. A process is also proposed to use GNF as a precursor material to fabricate GNS that will be used as EC electrodes. This results in much better electrical conductivity than activated carbon. This is advantageous for high-pulsed-power applications to reduce parasitic heating. Larger average pore size allows more complete electrolyte wetting (faster charge transfer kinetics). These properties contribute to a lowered equivalent series resistance (ESR), increased specific power, shorter charging times, and decreased parasitic heating. The high density of basal plane edges provides nucleation sites for activation (addition of hydrophilic functional groups) that facilitate electrolyte wetting, and will contribute to pseudocapacitance.
Compositionally graded relaxed AlGaN buffers on semipolar GaN for mid-ultraviolet emission
DOE Office of Scientific and Technical Information (OSTI.GOV)
Young, Erin C.; Wu Feng; Haeger, Daniel A.
In this Letter, we report on the growth and properties of relaxed, compositionally graded Al{sub x}Ga{sub 1-x}N buffer layers on freestanding semipolar (2021) GaN substrates. Continuous and step compositional grades with Al concentrations up to x = 0.61 have been achieved, with emission wavelengths in the mid-ultraviolet region as low as 265 nm. Coherency stresses were relaxed progressively throughout the grades by misfit dislocation generation via primary (basal) slip and secondary (non-basal) slip systems. Threading dislocation densities in the final layers of the grades were less than 10{sup 6}/cm{sup 2} as confirmed by plan-view transmission electron microscopy and cathodoluminescence studies.
Compositionally graded relaxed AlGaN buffers on semipolar GaN for mid-ultraviolet emission
NASA Astrophysics Data System (ADS)
Young, Erin C.; Wu, Feng; Romanov, Alexey E.; Haeger, Daniel A.; Nakamura, Shuji; Denbaars, Steven P.; Cohen, Daniel A.; Speck, James S.
2012-10-01
In this Letter, we report on the growth and properties of relaxed, compositionally graded AlxGa1 - xN buffer layers on freestanding semipolar (202¯1) GaN substrates. Continuous and step compositional grades with Al concentrations up to x = 0.61 have been achieved, with emission wavelengths in the mid-ultraviolet region as low as 265 nm. Coherency stresses were relaxed progressively throughout the grades by misfit dislocation generation via primary (basal) slip and secondary (non-basal) slip systems. Threading dislocation densities in the final layers of the grades were less than 106/cm2 as confirmed by plan-view transmission electron microscopy and cathodoluminescence studies.
Density- and trait-mediated effects of a parasite and a predator in a tri-trophic food web
Banerji, Aabir; Duncan, Alison B; Griffin, Joanne S; Humphries, Stuart; Petchey, Owen L; Kaltz, Oliver
2015-01-01
1. Despite growing interest in ecological consequences of parasitism in food webs, relatively little is known about effects of parasites on long-term population dynamics of non-host species or about whether such effects are density or trait mediated. 2. We studied a tri-trophic food chain comprised of (i) a bacterial basal resource (Serratia fonticola), (ii) an intermediate consumer (Paramecium caudatum), (iii) a top predator (Didinium nasutum) and (iv) a parasite of the intermediate consumer (Holospora undulata). A fully factorial experimental manipulation of predator and parasite presence/absence was combined with analyses of population dynamics, modelling and analyses of host (Paramecium) morphology and behaviour. 3. Predation and parasitism each reduced the abundance of the intermediate consumer (Paramecium), and parasitism indirectly reduced the abundance of the basal resource (Serratia). However, in combination, predation and parasitism had non-additive effects on the abundance of the intermediate consumer, as well as on that of the basal resource. In both cases, the negative effect of parasitism seemed to be effaced by predation. 4. Infection of the intermediate consumer reduced predator abundance. Modelling and additional experimentation revealed that this was most likely due to parasite reduction of intermediate host abundance (a density-mediated effect), as opposed to changes in predator functional or numerical response. 5. Parasitism altered morphological and behavioural traits, by reducing host cell length and increasing the swimming speed of cells with moderate parasite loads. Additional tests showed no significant difference in Didinium feeding rate on infected and uninfected hosts, suggesting that the combination of these modifications does not affect host vulnerability to predation. However, estimated rates of encounter with Serratia based on these modifications were higher for infected Paramecium than for uninfected Paramecium. 6. A mixture of density-mediated and trait-mediated indirect effects of parasitism on non-host species creates rich and complex possibilities for effects of parasites in food webs that should be included in assessments of possible impacts of parasite eradication or introduction. PMID:25382389
Zhou, Xin; Fu, Xin; Lin, Chun; Zhou, Xiaojuan; Liu, Jin; Wang, Li; Zhang, Xinwen; Zuo, Mingxue; Fan, Xiaolong; Li, Dapeng; Sun, Yingyu
2017-05-01
Deafening elicits a deterioration of learned vocalization, in both humans and songbirds. In songbirds, learned vocal plasticity has been shown to depend on the basal ganglia-cortical circuit, but the underlying cellular basis remains to be clarified. Using confocal imaging and electron microscopy, we examined the effect of deafening on dendritic spines in avian vocal motor cortex, the robust nucleus of the arcopallium (RA), and investigated the role of the basal ganglia circuit in motor cortex plasticity. We found rapid structural changes to RA dendritic spines in response to hearing loss, accompanied by learned song degradation. In particular, the morphological characters of RA spine synaptic contacts between 2 major pathways were altered differently. However, experimental disruption of the basal ganglia circuit, through lesions in song-specialized basal ganglia nucleus Area X, largely prevented both the observed changes to RA dendritic spines and the song deterioration after hearing loss. Our results provide cellular evidence to highlight a key role of the basal ganglia circuit in the motor cortical plasticity that underlies learned vocal plasticity. © The Author 2016. Published by Oxford University Press. All rights reserved. For Permissions, please e-mail: journals.permissions@oup.com.
Climate limits across space and time on European forest structure
NASA Astrophysics Data System (ADS)
Moreno, A. L. S.; Neumann, M.; Hasenauer, H.
2017-12-01
The impact climate has on forests has been extensively studied. However, the large scale effect climate has on forest structures, such as average diameters, heights and basal area are understudied in a spatially explicit manner. The limits, tipping points and thresholds that climate places on forest structures dictate the services a forest may provide, the vulnerability of a forest to mortality and the potential value of the timber there within. The majority of current research either investigates climate impacts on forest pools and fluxes, on a tree physiological scale or on case studies that are used to extrapolate results and potential impacts. A spatially explicit study on how climate affects forest structure over a large region would give valuable information to stakeholders who are more concerned with ecosystem services that cannot be described by pools and fluxes but require spatially explicit information - such as biodiversity, habitat suitability, and market values. In this study, we quantified the limits that climate (maximum, minimum temperature and precipitation) places on 3 forest structures, diameter at breast height, height, and basal area throughout Europe. Our results show clear climatic zones of high and low upper limits for each forest structure variable studied. We also spatially analyzed how climate restricts the potential bio-physical upper limits and creates tipping points of each forest structure variable and which climate factors are most limiting. Further, we demonstrated how the climate change has affected 8 individual forests across Europe and then the continent as a whole. We find that diameter, height and basal area are limited by climate in different ways and that areas may have high upper limits in one structure and low upper limits in another limitted by different climate variables. We also found that even though individual forests may have increased their potential upper limit forest structure values, European forests as a whole have lost, on average, 5.0%, 1.7% and 6.5% in potential mean forest diameter, height and basal area, respectively.
Fine structure of the ependyma and intercellular junctions in the area postrema of the rat.
Gotow, T; Hashimoto, P H
1979-09-03
Ependymal cells and their junctional complexes in the area postrema of the rat were studied in detail by tracer experiments using horseradish peroxidase (HRP) and colloidal lanthanum and by freeze-etch techniques, in addition to routine electron microscopy. The ependyma of the area postrema is characterized as flattened cells possessing very few cilia, a moderate amount of microvilli, a well-developed Golgi apparatus and rough endoplasmic reticulum. Numerous vesicles or tubular formations with internal dense content were found to accumulate in the basal processes of ependymal cells; the basal process makes contact with the perivascular basal lamina. It is suggested that the dense material in the tubulovesicular formations is synthesized within the ependymal cell and discharged into the perivascular space. The apical junctions between adjacent ependymal cells display very close apposition, with a gap of 2--3 nm, but no fusion of adjacent plasma membranes; they thus represent a transitional form between the zonulae adhaerentes present in the ordinary mural ependyma and the zonulae occludentes in the choroidal epithelium. A direct intercommunication between the ventricular cerebrospinal fluid (CSF) and the blood vascular system indicates that a region exists lacking a blood-ventricular CSF barrier.
NASA Astrophysics Data System (ADS)
Lane, Patrick
2016-04-01
Estimating the water balance of ungauged catchments has been the subject of decades of research. An extension of the fundamental problem of estimating the hydrology is then understanding how do changes in catchment attributes affect the water balance component? This is a particular issue in forest hydrology where vegetation exerts such a strong influence on evapotranspiration (ET), and consequent streamflow (Q). Given the primacy of trees in the water balance, and the potential for change to species and density through logging, fire, pests and diseases and drought, methods that directly relate ET/Q to vegetation structure, species, and stand density are very powerful. Plot studies on tree water use routinely use sapwood area (SA) to calculate transpiration and upscale to the stand/catchment scale. Recent work in south eastern Australian forests have found stand-wide SA to be linearly correlated (R2 = 0.89) with long term mean annual loss (P-Q), and hence, long term mean annual catchment streamflow. Robust relationships can be built between basal area (BA), tree density and stand SA. BA and density are common forest inventory measurements. Until now, no research has related the fundamental stand attribute of SA to streamflow. The data sets include catchments that have been thinned and with varying age classes. Thus far these analyses have been for energy limited systems in wetter forest types. SA has proven to be a more robust biometric than leaf area index which varies seasonally. That long term ET/Q is correlated with vegetation conforms to the Budyko framework. Use of a downscaled (20 m) Aridity Index (AI) has shown distinct correlations with stand SA, and therefore T. Structural patterns at a the hillslope scale not only correlate with SA and T, but also with interception (I) and forest floor evaporation (Es). These correlations between AI and I and Es have given R2 > 0.8. The result of these studies suggest an ability to estimate mean annual ET fluxes at sub hillslope scale using mappable attributes (AI, forest inventory data). Advances in forest inventory techniques, including LiDAR, mean stand attributes can increasingly be mapped over large areas. If combined with process measurements, these mapped attributes provide a powerful platform for simple but robust modelling at the sub-hillslope scale, including exploring hinge points of stand vulnerability to the drier, hotter climate predicted for SE Australia where energy limited systems may face water limitation.
Mendivelso, Hooz A; Camarero, J Julio; Royo Obregón, Oriol; Gutiérrez, Emilia; Toledo, Marisol
2013-01-01
A seasonal period of water deficit characterizes tropical dry forests (TDFs). There, sympatric tree species exhibit a diversity of growth rates, functional traits, and responses to drought, suggesting that each species may possess different strategies to grow under different conditions of water availability. The evaluation of the long-term growth responses to changes in the soil water balance should provide an understanding of how and when coexisting tree species respond to water deficit in TDFs. Furthermore, such differential growth responses may be linked to functional traits related to water storage and conductance. We used dendrochronology and climate data to retrospectively assess how the radial growth of seven coexisting deciduous tree species responded to the seasonal soil water balance in a Bolivian TDF. Linear mixed-effects models were used to quantify the relationships between basal area increment and seasonal water balance. We related these relationships with wood density and sapwood production to assess if they affect the growth responses to climate. The growth of all species responded positively to water balance during the wet season, but such responses differed among species as a function of their wood density. For instance, species with a strong growth response to water availability averaged a low wood density which may facilitate the storage of water in the stem. By contrast, species with very dense wood were those whose growth was less sensitive to water availability. Coexisting tree species thus show differential growth responses to changes in soil water balance during the wet season. Our findings also provide a link between wood density, a trait related to the ability of trees to store water in the stem, and wood formation in response to water availability.
Mendivelso, Hooz A.; Camarero, J. Julio; Royo Obregón, Oriol; Gutiérrez, Emilia; Toledo, Marisol
2013-01-01
A seasonal period of water deficit characterizes tropical dry forests (TDFs). There, sympatric tree species exhibit a diversity of growth rates, functional traits, and responses to drought, suggesting that each species may possess different strategies to grow under different conditions of water availability. The evaluation of the long-term growth responses to changes in the soil water balance should provide an understanding of how and when coexisting tree species respond to water deficit in TDFs. Furthermore, such differential growth responses may be linked to functional traits related to water storage and conductance. We used dendrochronology and climate data to retrospectively assess how the radial growth of seven coexisting deciduous tree species responded to the seasonal soil water balance in a Bolivian TDF. Linear mixed-effects models were used to quantify the relationships between basal area increment and seasonal water balance. We related these relationships with wood density and sapwood production to assess if they affect the growth responses to climate. The growth of all species responded positively to water balance during the wet season, but such responses differed among species as a function of their wood density. For instance, species with a strong growth response to water availability averaged a low wood density which may facilitate the storage of water in the stem. By contrast, species with very dense wood were those whose growth was less sensitive to water availability. Coexisting tree species thus show differential growth responses to changes in soil water balance during the wet season. Our findings also provide a link between wood density, a trait related to the ability of trees to store water in the stem, and wood formation in response to water availability. PMID:24116001
Cykowski, Matthew D; Takei, Hidehiro; Van Eldik, Linda J; Schmitt, Frederick A; Jicha, Gregory A; Powell, Suzanne Z; Nelson, Peter T
2016-05-01
Transactivating responsive sequence (TAR) DNA-binding protein 43-kDa (TDP-43) pathology has been described in various brain diseases, but the full anatomical distribution and clinical and biological implications of that pathology are incompletely characterized. Here, we describe TDP-43 neuropathology in the basal forebrain, hypothalamus, and adjacent nuclei in 98 individuals (mean age, 86 years; median final mini-mental state examination score, 27). On examination blinded to clinical and pathologic diagnoses, we identified TDP-43 pathology that most frequently involved the ventromedial basal forebrain in 19 individuals (19.4%). As expected, many of these brains had comorbid pathologies including those of Alzheimer disease (AD), Lewy body disease (LBD), and/or hippocampal sclerosis of aging (HS-Aging). The basal forebrain TDP-43 pathology was strongly associated with comorbid HS-Aging (odds ratio = 6.8, p = 0.001), whereas there was no significant association between basal forebrain TDP-43 pathology and either AD or LBD neuropathology. In this sample, there were some cases with apparent preclinical TDP-43 pathology in the basal forebrain that may indicate that this is an early affected area in HS-Aging. We conclude that TDP-43 pathology in the basal forebrain is strongly associated with HS-Aging. These results raise questions about a specific pathogenetic relationship between basal forebrain TDP-43 and non-HS-Aging comorbid diseases (AD and LBD). © 2016 American Association of Neuropathologists, Inc. All rights reserved.
Takei, Hidehiro; Van Eldik, Linda J.; Schmitt, Frederick A.; Jicha, Gregory A.; Powell, Suzanne Z.; Nelson, Peter T.
2016-01-01
Transactivating responsive sequence (TAR) DNA-binding protein 43-kDa (TDP-43) pathology has been described in various brain diseases, but the full anatomical distribution and clinical and biological implications of that pathology are incompletely characterized. Here, we describe TDP-43 neuropathology in the basal forebrain, hypothalamus, and adjacent nuclei in 98 individuals (mean age, 86 years; median final mini-mental state examination score, 27). On examination blinded to clinical and pathologic diagnoses, we identified TDP-43 pathology that most frequently involved the ventromedial basal forebrain in 19 individuals (19.4%). As expected, many of these brains had comorbid pathologies including those of Alzheimer disease (AD), Lewy body disease (LBD), and/or hippocampal sclerosis of aging (HS-Aging). The basal forebrain TDP-43 pathology was strongly associated with comorbid HS-Aging (odds ratio = 6.8, p = 0.001), whereas there was no significant association between basal forebrain TDP-43 pathology and either AD or LBD neuropathology. In this sample, there were some cases with apparent preclinical TDP-43 pathology in the basal forebrain that may indicate that this is an early affected area in HS-Aging. We conclude that TDP-43 pathology in the basal forebrain is strongly associated with HS-Aging. These results raise questions about a specific pathogenetic relationship between basal forebrain TDP-43 and non-HS-Aging comorbid diseases (AD and LBD). PMID:26971127
Basal ganglia function, stuttering, sequencing, and repair in adult songbirds
Kubikova, Lubica; Bosikova, Eva; Cvikova, Martina; Lukacova, Kristina; Scharff, Constance; Jarvis, Erich D.
2014-01-01
A pallial-basal-ganglia-thalamic-pallial loop in songbirds is involved in vocal motor learning. Damage to its basal ganglia part, Area X, in adult zebra finches has been noted to have no strong effects on song and its function is unclear. Here we report that neurotoxic damage to adult Area X induced changes in singing tempo and global syllable sequencing in all animals, and considerably increased syllable repetition in birds whose song motifs ended with minor repetitions before lesioning. This stuttering-like behavior started at one month, and improved over six months. Unexpectedly, the lesioned region showed considerable recovery, including immigration of newly generated or repaired neurons that became active during singing. The timing of the recovery and stuttering suggest that immature recovering activity of the circuit might be associated with stuttering. These findings indicate that even after juvenile learning is complete, the adult striatum plays a role in higher level organization of learned vocalizations. PMID:25307086
Mao, Zhun; Saint-André, Laurent; Bourrier, Franck; Stokes, Alexia; Cordonnier, Thomas
2015-01-01
Background and Aims In mountain ecosystems, predicting root density in three dimensions (3-D) is highly challenging due to the spatial heterogeneity of forest communities. This study presents a simple and semi-mechanistic model, named ChaMRoots, that predicts root interception density (RID, number of roots m–2). ChaMRoots hypothesizes that RID at a given point is affected by the presence of roots from surrounding trees forming a polygon shape. Methods The model comprises three sub-models for predicting: (1) the spatial heterogeneity – RID of the finest roots in the top soil layer as a function of tree basal area at breast height, and the distance between the tree and a given point; (2) the diameter spectrum – the distribution of RID as a function of root diameter up to 50 mm thick; and (3) the vertical profile – the distribution of RID as a function of soil depth. The RID data used for fitting in the model were measured in two uneven-aged mountain forest ecosystems in the French Alps. These sites differ in tree density and species composition. Key Results In general, the validation of each sub-model indicated that all sub-models of ChaMRoots had good fits. The model achieved a highly satisfactory compromise between the number of aerial input parameters and the fit to the observed data. Conclusions The semi-mechanistic ChaMRoots model focuses on the spatial distribution of root density at the tree cluster scale, in contrast to the majority of published root models, which function at the level of the individual. Based on easy-to-measure characteristics, simple forest inventory protocols and three sub-models, it achieves a good compromise between the complexity of the case study area and that of the global model structure. ChaMRoots can be easily coupled with spatially explicit individual-based forest dynamics models and thus provides a highly transferable approach for modelling 3-D root spatial distribution in complex forest ecosystems. PMID:26173892
Soil Quality of Bauxite Mining Areas
NASA Astrophysics Data System (ADS)
Terezinha Gonçalves Bizuti, Denise; Dinarowski, Marcela; Casagrande, José Carlos; Silva, Luiz Gabriel; Soares, Marcio Roberto; Henrique Santin Brancalion, Pedro
2015-04-01
The study on soil quality index (SQI) aims to assess the current state of the soil after use and estimating its recovery through sustainable management practices This type of study is being used in this work in order to check the efficiency of forest recovery techniques in areas that have been deeply degraded by bauxite mining process, and compare them with the area of native forest, through the determination of SQI. Treatments were newly mined areas, areas undergoing restoration (topsoil use with planting of native forest species), areas in rehabilitation (employment of the green carpet with topsoil and planting of native forest species) and areas of native forests, with six repetitions, in areas of ALCOA, in the municipality of Poços de Caldas/MG. To this end, we used the additive pondered model, establishing three functions: Fertility, water movement and root development, based on chemical parameters (organic matter, base saturation, aluminum saturation and calcium content); physical (macroporosity, soil density and clay content); and microbiological testing (basal respiration by the emission of CO2 ). The SQIs obtained for each treatment was 41%, 56%, 63% and 71% for newly mined areas, native forest, areas in restoration and rehabilitation, respectively. The recovering technique that most approximates the degraded soil to the soil of reference is the restoration, where there was no statistically significant difference of areas restored with native forest. It was found that for the comparison of the studied areas must take into account the nutrient cycling, that disappear with plant removal in mining areas, once the soil of native forest features low fertility and high saturation by aluminum, also taking in account recovering time.
NASA Astrophysics Data System (ADS)
Coiffard, Clément; Mohr, Barbara
2014-05-01
Early Cretaceous Northern Gondwana seems to be the cradle of many early flowering plants, especially mesangiosperms that include magnoliids and monocots and basal eudicots. So far our knowledge was based mostly on dispersed pollen and small flowering structures. New fossil finds from Brazil include more complete plants with attached roots, leaves and flowers. Taxonomic studies show that these fossils belonged to clades which are, based on macroscopic characters and molecular data, also considered to be rather basal, such as several members of Nymphaeales, Piperales, Laurales, Magnoliales, monocots (Araliaceae) and Ranunculales. Various parameters can be used in order to understand the physiology and habitat of these plants. Adaptations to climate and habitat are partly mirrored in their root anatomy (evidence of tap roots), leaf size and shape, leaf anatomy including presence of glands, and distribution of stomata. An important ecophysiolocical parameter is vein density as an indicator for the plants' cabability to pump water, and the stomatal pore index, representing the proportion of stomatal pore area on the leaf surface, which is related to the water vapor resistance of the leaf epidermis. During the mid-Cretaceous leaf vein density started to surpass that of gymnosperms, one factor that made angiosperms very successful in conquering many kinds of new environments. Using data on these parameters we deduce that during the late Early to mid Cretaceous angiosperms were already diverse, being represented as both herbs, with aquatic members, such as Nymphaeles, helophytes (e.g. some monocots) and plants that may have grown in shady locations. Other life forms included shrubs and perhaps already small trees (e.g. Magnoliales). These flowering plants occupied various habitats, ranging from xeric (e.g. some Magnoliales) to mesic and shady (e.g. Piperales) or aquatic (e.g. Araceae, Nymphaeales). Overall, it seems that several of these plants clearly exhibited some mechanisms to withstand drought, which in turn let us assume that the climate was characterized by dry and wet seasons.