Sample records for early cambrian record

  1. An early Cambrian greenhouse climate.

    PubMed

    Hearing, Thomas W; Harvey, Thomas H P; Williams, Mark; Leng, Melanie J; Lamb, Angela L; Wilby, Philip R; Gabbott, Sarah E; Pohl, Alexandre; Donnadieu, Yannick

    2018-05-01

    The oceans of the early Cambrian (~541 to 509 million years ago) were the setting for a marked diversification of animal life. However, sea temperatures-a key component of the early Cambrian marine environment-remain unconstrained, in part because of a substantial time gap in the stable oxygen isotope (δ 18 O) record before the evolution of euconodonts. We show that previously overlooked sources of fossil biogenic phosphate have the potential to fill this gap. Pristine phosphatic microfossils from the Comley Limestones, UK, yield a robust δ 18 O signature, suggesting sea surface temperatures of 20° to 25°C at high southern paleolatitudes (~65°S to 70°S) between ~514 and 509 million years ago. These sea temperatures are consistent with the distribution of coeval evaporite and calcrete deposits, peak continental weathering rates, and also our climate model simulations for this interval. Our results support an early Cambrian greenhouse climate comparable to those of the late Mesozoic and early Cenozoic, offering a framework for exploring the interplay between biotic and environmental controls on Cambrian animal diversification.

  2. High-precision Temporal Calibration of the Early Cambrian Biotic and Paleoenvironmental Records: New U-Pb Geochronology from Eastern Yunnan, China.

    NASA Astrophysics Data System (ADS)

    Tsukui, K.; Ramezani, J.; Zhu, M.; Maloof, A. C.; Porter, S.; Moore, J.; Eddy, M. P.; Bowring, S. A.

    2016-12-01

    The Terreneuvian Epoch of the early Cambrian marks the global diversification of early animal life, as well as major perturbations to Earth's geochemical cycles. Understanding possible links between biotic evolution (e.g., emergence of skeletal animals) and the recognized changes in ocean chemistry requires a high fidelity chronostratigraphic framework for the early Cambrian records. One such chronostratigraphy was built by mapping local early Cambrian carbon isotope profiles onto a U-Pb age-calibrated marine carbonate δ13C record from Morocco, assuming global synchroneity of the observed δ13C trends. Here we present a direct test of this assumption using high-precision U-Pb geochronology (CA-ID-TIMS method) of ash beds from key lower Cambrian horizons throughout eastern Yunnan Province in South China. Preliminary age results from ash beds near the top of the Dengying Formation (Fm.) and the basal Daibu Member (Mb.) of the Zhujiaqing Fm. in multiple sections place the basal Cambrian negative δ13C excursion (BACE) in China at ca. 540.7-539.6 Ma. Our new U-Pb dates from the overlying Zhongyicun Mb. at the Meishucun and nearby sections improve significantly upon previous in situ U-Pb geochronology and constrain the onset of high-frequency δ13C oscillations in some sections to between 533.5 and 532.9 Ma. Most importantly, a new U-Pb date of ca. 526 Ma from the basal Shiyantou Fm. in the Xiaotan Section marks the termination of a >1 million year-long period of consistently positive (≥+4‰) δ13C values (ZHUCE) that is characteristic of many early Cambrian records worldwide. This date establishes a robust time correlation between ZHUCE in South China and its equivalent 5p excursion in Morocco and Siberia, and constrains the timing and duration of the largest positive δ13C anomalies in the Cambrian.

  3. Reconstruction of early Cambrian ocean chemistry from Mo isotopes

    NASA Astrophysics Data System (ADS)

    Wen, Hanjie; Fan, Haifeng; Zhang, Yuxu; Cloquet, Christophe; Carignan, Jean

    2015-09-01

    The Neoproterozoic-Cambrian transition was a key time interval in the history of the Earth, especially for variations in oceanic and atmospheric chemical composition. However, two conflicting views exist concerning the nature of ocean chemistry across the Precambrian-Cambrian boundary. Abundant geochemical evidence suggests that oceanic basins were fully oxygenated by the late Ediacaran, while other studies provide seemingly conflicting evidence for anoxic deep waters, with ferruginous conditions [Fe(II)-enriched] persisting into the Cambrian. Here, two early Cambrian sedimentary platform and shelf-slope sections in South China were investigated to trace early Cambrian ocean chemistry from Mo isotopes. The results reveal that early Cambrian sediments deposited under oxic to anoxic/euxinic conditions have δ98/95Mo values ranging from -0.28‰ to 2.29‰, which suggests that early Cambrian seawater may have had δ98/95Mo values of at least 2.29‰, similar to modern oceans. The heaviest and relatively homogeneous δ98/95Mo values were recorded in siltstone samples formed under completely oxic conditions, which is considered that Mn oxide-free shuttling was responsible for such heavy δ98/95Mo value. Further, combined with Fe species data and the accumulation extent of Mo and U, the variation of δ98/95Mo values in the two studied sections demonstrate a redox-stratified ocean with completely oxic shallow water and predominantly anoxic (even euxinic) deeper water having developed early on, which eventually became completely oxygenated. This suggests that oceanic circulation at the time became reorganized, and such changes in oceanic chemistry may have been responsible for triggering the "Cambrian Explosion" of biological diversity.

  4. Calibrating rates of early Cambrian evolution.

    PubMed

    Bowring, S A; Grotzinger, J P; Isachsen, C E; Knoll, A H; Pelechaty, S M; Kolosov, P

    1993-09-03

    An explosive episode of biological diversification occurred near the beginning of the Cambrian period. Evolutionary rates in the Cambrian have been difficult to quantify accurately because of a lack of high-precision ages. Currently, uranium-lead zircon geochronology is the most powerful method for dating rocks of Cambrian age. Uranium-lead zircon data from lower Cambrian rocks located in northeast Siberia indicate that the Cambrian period began at approximately 544 million years ago and that its oldest (Manykaian) stage lasted no less than 10 million years. Other data indicate that the Tommotian and Atdabanian stages together lasted only 5 to 10 million years. The resulting compression of Early Cambrian time accentuates the rapidity of both the faunal diversification and subsequent Cambrian turnover.

  5. Calibrating rates of early Cambrian evolution

    NASA Technical Reports Server (NTRS)

    Bowring, Samuel A.; Grotzinger, John P.; Isachsen, Clark E.; Knoll, Andrew H.; Pelechaty, Shane M.; Kolosov, Peter

    1993-01-01

    An explosive episode of biological diversification occurred near the beginning of the Cambrian period. Evolutionary rates in the Cambrian have been difficult to quantify accurately because of a lack of high-precision ages. Currently, uranium-lead zircon geochronology is the most powerful method for dating rocks of Cambrian age. Uranium-lead zircon data from lower Cambrian rocks located in northeast Siberia indicate that the Cambrian period began about 544 million years ago and that its oldest (Manykaian) stage lasted no less than 10 million years. Other data indicate that the Tommotian and Atdabanian stages together lasted only 5 to 10 million years. The resulting compression of Early Cambrian time accentuates the rapidity of both the faunal diversification and subsequent Cambrian turnover.

  6. Early Cambrian oxygen minimum zone-like conditions at Chengjiang

    NASA Astrophysics Data System (ADS)

    Hammarlund, Emma U.; Gaines, Robert R.; Prokopenko, Maria G.; Qi, Changshi; Hou, Xian-Guang; Canfield, Donald E.

    2017-10-01

    The early Cambrian succession at Chengjiang contains the most diverse Cambrian fossil assemblage yet described, and contributes significantly to our understanding of the diversification of metazoans in the Cambrian ;explosion;. The Cambrian Period occupies a transitional episode of global ocean chemistry, following the oxygenation of the surface ocean and of shallow marine environments during the Ediacaran Period, but prior to the establishment of a predominantly oxygenated deep ocean in the mid-Paleozoic. Despite recent attention, a detailed understanding of the chemical conditions that prevailed in early Cambrian marine settings and the relationship of those conditions to early metazoan ecosystems is still emerging. Here, we report multi-proxy geochemical data from two drill cores through the early Cambrian (Series 2) Yu'anshan Formation of Yunnan, China. Results reveal dynamic water-column chemistry within the succession, which progressively shifted from euxinic to oxic conditions during deposition of the Yu'anshan Formation. The Chengjiang biota occurs in strata that appear to have been deposited under an oxygen-depleted water column that may have supported denitrification, as in modern oxygen-minimum zones. The oxygenated benthic environments in which the Chengjiang biota thrived were proximal to, but sharply separated from, the open ocean by a persistent anoxic water mass that occupied a portion of the outer shelf. Oxygen depletion in the lower water column developed dynamically in response to nutrient availability and possibly at lower thresholds of productivity due to lower atmospheric oxygen concentrations in Cambrian. These findings suggest that the frequent development of oxygen-limiting conditions in continental margin settings provided an environmental barrier that may have affected biogeographic, ecological and evolutionary development of early metazoan communities.

  7. A suspension-feeding anomalocarid from the Early Cambrian.

    PubMed

    Vinther, Jakob; Stein, Martin; Longrich, Nicholas R; Harper, David A T

    2014-03-27

    Large, actively swimming suspension feeders evolved several times in Earth's history, arising independently from groups as diverse as sharks, rays and stem teleost fishes, and in mysticete whales. However, animals occupying this niche have not been identified from the early Palaeozoic era. Anomalocarids, a group of stem arthropods that were the largest nektonic animals of the Cambrian and Ordovician periods, are generally thought to have been apex predators. Here we describe new material from Tamisiocaris borealis, an anomalocarid from the Early Cambrian (Series 2) Sirius Passet Fauna of North Greenland, and propose that its frontal appendage is specialized for suspension feeding. The appendage bears long, slender and equally spaced ventral spines furnished with dense rows of long and fine auxiliary spines. This suggests that T. borealis was a microphagous suspension feeder, using its appendages for sweep-net capture of food items down to 0.5 mm, within the size range of mesozooplankton such as copepods. Our observations demonstrate that large, nektonic suspension feeders first evolved during the Cambrian explosion, as part of an adaptive radiation of anomalocarids. The presence of nektonic suspension feeders in the Early Cambrian, together with evidence for a diverse pelagic community containing phytoplankton and mesozooplankton, indicate the existence of a complex pelagic ecosystem supported by high primary productivity and nutrient flux. Cambrian pelagic ecosystems seem to have been more modern than previously believed.

  8. Armored kinorhynch-like scalidophoran animals from the early Cambrian.

    PubMed

    Zhang, Huaqiao; Xiao, Shuhai; Liu, Yunhuan; Yuan, Xunlai; Wan, Bin; Muscente, A D; Shao, Tiequan; Gong, Hao; Cao, Guohua

    2015-11-26

    Morphology-based phylogenetic analyses support the monophyly of the Scalidophora (Kinorhyncha, Loricifera, Priapulida) and Nematoida (Nematoda, Nematomorpha), together constituting the monophyletic Cycloneuralia that is the sister group of the Panarthropoda. Kinorhynchs are unique among living cycloneuralians in having a segmented body with repeated cuticular plates, longitudinal muscles, dorsoventral muscles, and ganglia. Molecular clock estimates suggest that kinorhynchs may have diverged in the Ediacaran Period. Remarkably, no kinorhynch fossils have been discovered, in sharp contrast to priapulids and loriciferans that are represented by numerous Cambrian fossils. Here we describe several early Cambrian (~535 million years old) kinorhynch-like fossils, including the new species Eokinorhynchus rarus and two unnamed but related forms. E. rarus has characteristic scalidophoran features, including an introvert with pentaradially arranged hollow scalids. Its trunk bears at least 20 annuli each consisting of numerous small rectangular plates, and is armored with five pairs of large and bilaterally placed sclerites. Its trunk annuli are reminiscent of the epidermis segments of kinorhynchs. A phylogenetic analysis resolves E. rarus as a stem-group kinorhynch. Thus, the fossil record confirms that all three scalidophoran phyla diverged no later than the Cambrian Period.

  9. Armored kinorhynch-like scalidophoran animals from the early Cambrian

    PubMed Central

    Zhang, Huaqiao; Xiao, Shuhai; Liu, Yunhuan; Yuan, Xunlai; Wan, Bin; Muscente, A. D.; Shao, Tiequan; Gong, Hao; Cao, Guohua

    2015-01-01

    Morphology-based phylogenetic analyses support the monophyly of the Scalidophora (Kinorhyncha, Loricifera, Priapulida) and Nematoida (Nematoda, Nematomorpha), together constituting the monophyletic Cycloneuralia that is the sister group of the Panarthropoda. Kinorhynchs are unique among living cycloneuralians in having a segmented body with repeated cuticular plates, longitudinal muscles, dorsoventral muscles, and ganglia. Molecular clock estimates suggest that kinorhynchs may have diverged in the Ediacaran Period. Remarkably, no kinorhynch fossils have been discovered, in sharp contrast to priapulids and loriciferans that are represented by numerous Cambrian fossils. Here we describe several early Cambrian (~535 million years old) kinorhynch-like fossils, including the new species Eokinorhynchus rarus and two unnamed but related forms. E. rarus has characteristic scalidophoran features, including an introvert with pentaradially arranged hollow scalids. Its trunk bears at least 20 annuli each consisting of numerous small rectangular plates, and is armored with five pairs of large and bilaterally placed sclerites. Its trunk annuli are reminiscent of the epidermis segments of kinorhynchs. A phylogenetic analysis resolves E. rarus as a stem-group kinorhynch. Thus, the fossil record confirms that all three scalidophoran phyla diverged no later than the Cambrian Period. PMID:26610151

  10. Lower Cambrian polychaete from China sheds light on early annelid evolution

    NASA Astrophysics Data System (ADS)

    Liu, Jianni; Ou, Qiang; Han, Jian; Li, Jinshu; Wu, Yichen; Jiao, Guoxiang; He, Tongjiang

    2015-06-01

    We herein report a fossilized polychaete annelid, Guanshanchaeta felicia gen. et sp. nov., from the Lower Cambrian Guanshan Biota (Cambrian Series 2, stage 4). The new taxon has a generalized polychaete morphology, with biramous parapodia (most of which preserve the evidence of chaetae), an inferred prostomium bearing a pair of appendages, and a bifid pygidium. G. felicia is the first unequivocal annelid reported from the Lower Cambrian of China. It represents one of the oldest annelids among those from other early Paleozoic Lagerstätten including Sirius Passet from Greenland (Vinther et al., Nature 451: 185-188, 2008) and Emu Bay from Kangaroo island (Parry et al., Palaeontology 57: 1091-1103, 2014), and adds to our increasing roll of present-day animal phyla recognized in the early Cambrian Guanshan Biota. This finding expands the panorama of the Cambrian `explosion' exemplified by the Guanshan Biota, suggesting the presence of many more fossil annelids in the Chengjiang Lagerstätte and the Kaili Biota. In addition, this new taxon increases our knowledge of early polychaete morphology, which suggests that polychaete annelids considerably diversified in the Cambrian.

  11. Early Cambrian origin of modern food webs: evidence from predator arrow worms.

    PubMed

    Vannier, J; Steiner, M; Renvoisé, E; Hu, S-X; Casanova, J-P

    2007-03-07

    Although palaeontological evidence from exceptional biota demonstrates the existence of diverse marine communities in the Early Cambrian (approx. 540-520 Myr ago), little is known concerning the functioning of the marine ecosystem, especially its trophic structure and the full range of ecological niches colonized by the fauna. The presence of a diverse zooplankton in Early Cambrian oceans is still an open issue. Here we provide compelling evidence that chaetognaths, an important element of modern zooplankton, were present in the Early Cambrian Chengjiang biota with morphologies almost identical to Recent forms. New information obtained from the lowermost Cambrian of China added to previous studies provide convincing evidence that protoconodont-bearing animals also belonged to chaetognaths. Chaetognaths were probably widespread and diverse in the earliest Cambrian. The obvious raptorial function of their circumoral apparatuses (grasping spines) places them among the earliest active predator metazoans. Morphology, body ratios and distribution suggest that the ancestral chaetognaths were planktonic with possible ecological preferences for hyperbenthic niches close to the sea bottom. Our results point to the early introduction of prey-predator relationships into the pelagic realm, and to the increase of trophic complexity (three-level structure) during the Precambrian-Cambrian transition, thus laying the foundations of present-day marine food chains.

  12. Early Cambrian origin of modern food webs: evidence from predator arrow worms

    PubMed Central

    Vannier, J; Steiner, M; Renvoisé, E; Hu, S.-X; Casanova, J.-P

    2006-01-01

    Although palaeontological evidence from exceptional biota demonstrates the existence of diverse marine communities in the Early Cambrian (approx. 540–520 Myr ago), little is known concerning the functioning of the marine ecosystem, especially its trophic structure and the full range of ecological niches colonized by the fauna. The presence of a diverse zooplankton in Early Cambrian oceans is still an open issue. Here we provide compelling evidence that chaetognaths, an important element of modern zooplankton, were present in the Early Cambrian Chengjiang biota with morphologies almost identical to Recent forms. New information obtained from the lowermost Cambrian of China added to previous studies provide convincing evidence that protoconodont-bearing animals also belonged to chaetognaths. Chaetognaths were probably widespread and diverse in the earliest Cambrian. The obvious raptorial function of their circumoral apparatuses (grasping spines) places them among the earliest active predator metazoans. Morphology, body ratios and distribution suggest that the ancestral chaetognaths were planktonic with possible ecological preferences for hyperbenthic niches close to the sea bottom. Our results point to the early introduction of prey–predator relationships into the pelagic realm, and to the increase of trophic complexity (three-level structure) during the Precambrian–Cambrian transition, thus laying the foundations of present-day marine food chains. PMID:17254986

  13. A highly redox-heterogeneous ocean in South China during the early Cambrian (˜529-514 Ma): Implications for biota-environment co-evolution

    NASA Astrophysics Data System (ADS)

    Jin, Chengsheng; Li, Chao; Algeo, Thomas J.; Planavsky, Noah J.; Cui, Hao; Yang, Xinglian; Zhao, Yuanlong; Zhang, Xingliang; Xie, Shucheng

    2016-05-01

    The ;Cambrian Explosion; is known for rapid increases in the morphological disparity and taxonomic diversity of metazoans. It has been widely proposed that this biological event was a consequence of oxygenation of the global ocean, but this hypothesis is still under debate. Here, we present high-resolution Fe-S-C-Al-trace element geochemical records from the Jinsha (outer shelf) and Weng'an (outer shelf) sections of the early Cambrian Yangtze Platform, integrating these results with previously published data from six correlative sections representing a range of water depths (Xiaotan, Shatan, Dingtai, Yangjiaping, Songtao, and Longbizui). The integrated iron chemistry and redox-sensitive trace element data suggest that euxinic mid-depth waters dynamically coexisted with oxic surface waters and ferruginous deep waters during the earliest Cambrian, but that stepwise expansion of oxic waters commenced during Cambrian Stage 3 (∼ 521- 514 Ma). Combined with data from lower Cambrian sections elsewhere, including Oman, Iran and Canada, we infer that the global ocean exhibited a high degree of redox heterogeneity during the early Cambrian, consistent with low atmospheric oxygen levels (∼ 10- 40% of present atmospheric level, or PAL). A large spatial gradient in pyrite sulfur isotopic compositions (δ34Spy), which vary from a mean of - 12.0 ‰ in nearshore areas to + 22.5 ‰ in distal deepwater sections in lower Cambrian marine units of South China imply low concentrations and spatial heterogeneity of seawater sulfate, which is consistent with a limited oceanic sulfate reservoir globally. By comparing our reconstructed redox chemistry with fossil records from the lower Cambrian of South China, we infer that a stepwise oxygenation of shelf and slope environments occurred concurrently with a gradual increase in ecosystem complexity. However, deep waters remained anoxic and ferruginous even as macrozooplankton and suspension-feeding mesozooplankton appeared during

  14. Modern optics in exceptionally preserved eyes of Early Cambrian arthropods from Australia

    NASA Astrophysics Data System (ADS)

    Lee, Michael S. Y.; Jago, James B.; García-Bellido, Diego C.; Edgecombe, Gregory D.; Gehling, James G.; Paterson, John R.

    2011-06-01

    Despite the status of the eye as an ``organ of extreme perfection'', theory suggests that complex eyes can evolve very rapidly. The fossil record has, until now, been inadequate in providing insight into the early evolution of eyes during the initial radiation of many animal groups known as the Cambrian explosion. This is surprising because Cambrian Burgess-Shale-type deposits are replete with exquisitely preserved animals, especially arthropods, that possess eyes. However, with the exception of biomineralized trilobite eyes, virtually nothing is known about the details of their optical design. Here we report exceptionally preserved fossil eyes from the Early Cambrian (~515 million years ago) Emu Bay Shale of South Australia, revealing that some of the earliest arthropods possessed highly advanced compound eyes, each with over 3,000 large ommatidial lenses and a specialized `bright zone'. These are the oldest non-biomineralized eyes known in such detail, with preservation quality exceeding that found in the Burgess Shale and Chengjiang deposits. Non-biomineralized eyes of similar complexity are otherwise unknown until about 85 million years later. The arrangement and size of the lenses indicate that these eyes belonged to an active predator that was capable of seeing in low light. The eyes are more complex than those known from contemporaneous trilobites and are as advanced as those of many living forms. They provide further evidence that the Cambrian explosion involved rapid innovation in fine-scale anatomy as well as gross morphology, and are consistent with the concept that the development of advanced vision helped to drive this great evolutionary event.

  15. Trace fossils and substrates of the terminal Proterozoic–Cambrian transition: Implications for the record of early bilaterians and sediment mixing

    PubMed Central

    Droser, Mary L.; Jensen, Sören; Gehling, James G.

    2002-01-01

    The trace fossil record is important in determining the timing of the appearance of bilaterian animals. A conservative estimate puts this time at ≈555 million years ago. The preservational potential of traces made close to the sediment–water interface is crucial to detecting early benthic activity. Our studies on earliest Cambrian sediments suggest that shallow tiers were preserved to a greater extent than typical for most of the Phanerozoic, which can be attributed both directly and indirectly to the low levels of sediment mixing. The low levels of sediment mixing meant that thin event beds were preserved. The shallow depth of sediment mixing also meant that muddy sediments were firm close to the sediment–water interface, increasing the likelihood of recording shallow-tier trace fossils in muddy sediments. Overall, trace fossils can provide a sound record of the onset of bilaterian benthic activity. PMID:12271130

  16. Early Cambrian sipunculan worms from southwest China.

    PubMed

    Huang, Di-Ying; Chen, Jun-Yuan; Vannier, Jean; Saiz Salinas, J I

    2004-08-22

    We report the discovery of sipunculan worms from the Lower Cambrian Maotianshan Shale, near Kunming (southwest China). Their sipunculan identity is evidenced by the general morphology of the animals (sausage-shaped body with a slender retractable introvert and a wider trunk) and by other features, both external (e.g. perioral crown of tentacles, and hooks, papillae and wrinkle rings on the body surface) and internal (U-shaped gut, and the anus opening near the introvert-trunk junction). The three fossil forms (Archaeogolfingia caudata gen. et sp. nov., Cambrosipunculus tentaculatus gen. et sp. nov. and Cambrosipunculus sp.) have striking similarities to modern sipunculans, especially the Golfingiidae to which their evolutionary relationships are discussed. This study suggests that most typical features of extant sipunculans have undergone only limited changes since the Early Cambrian, thus indicating a possible evolutionary stasis over the past 520 Myr.

  17. Arthropod eyes: The early Cambrian fossil record and divergent evolution of visual systems.

    PubMed

    Strausfeld, Nicholas J; Ma, Xiaoya; Edgecombe, Gregory D; Fortey, Richard A; Land, Michael F; Liu, Yu; Cong, Peiyun; Hou, Xianguang

    2016-03-01

    Four types of eyes serve the visual neuropils of extant arthropods: compound retinas composed of adjacent facets; a visual surface populated by spaced eyelets; a smooth transparent cuticle providing inwardly directed lens cylinders; and single-lens eyes. The first type is a characteristic of pancrustaceans, the eyes of which comprise lenses arranged as hexagonal or rectilinear arrays, each lens crowning 8-9 photoreceptor neurons. Except for Scutigeromorpha, the second type typifies Myriapoda whose relatively large eyelets surmount numerous photoreceptive rhabdoms stacked together as tiers. Scutigeromorph eyes are facetted, each lens crowning some dozen photoreceptor neurons of a modified apposition-type eye. Extant chelicerate eyes are single-lensed except in xiphosurans, whose lateral eyes comprise a cuticle with a smooth outer surface and an inner one providing regular arrays of lens cylinders. This account discusses whether these disparate eye types speak for or against divergence from one ancestral eye type. Previous considerations of eye evolution, focusing on the eyes of trilobites and on facet proliferation in xiphosurans and myriapods, have proposed that the mode of development of eyes in those taxa is distinct from that of pancrustaceans and is the plesiomorphic condition from which facetted eyes have evolved. But the recent discovery of enormous regularly facetted compound eyes belonging to early Cambrian radiodontans suggests that high-resolution facetted eyes with superior optics may be the ground pattern organization for arthropods, predating the evolution of arthrodization and jointed post-protocerebral appendages. Here we provide evidence that compound eye organization in stem-group euarthropods of the Cambrian can be understood in terms of eye morphologies diverging from this ancestral radiodontan-type ground pattern. We show that in certain Cambrian groups apposition eyes relate to fixed or mobile eyestalks, whereas other groups reveal concomitant

  18. Early Cambrian sipunculan worms from southwest China.

    PubMed Central

    Huang, Di-Ying; Chen, Jun-Yuan; Vannier, Jean; Saiz Salinas, J. I.

    2004-01-01

    We report the discovery of sipunculan worms from the Lower Cambrian Maotianshan Shale, near Kunming (southwest China). Their sipunculan identity is evidenced by the general morphology of the animals (sausage-shaped body with a slender retractable introvert and a wider trunk) and by other features, both external (e.g. perioral crown of tentacles, and hooks, papillae and wrinkle rings on the body surface) and internal (U-shaped gut, and the anus opening near the introvert-trunk junction). The three fossil forms (Archaeogolfingia caudata gen. et sp. nov., Cambrosipunculus tentaculatus gen. et sp. nov. and Cambrosipunculus sp.) have striking similarities to modern sipunculans, especially the Golfingiidae to which their evolutionary relationships are discussed. This study suggests that most typical features of extant sipunculans have undergone only limited changes since the Early Cambrian, thus indicating a possible evolutionary stasis over the past 520 Myr. PMID:15306286

  19. Early and middle(?) Cambrian metazoan and protistan fossils from West Africa

    USGS Publications Warehouse

    Culver, S.J.; Repetski, J.E.; Pojeta, J.; Hunt, D.

    1996-01-01

    Supposed Upper Proterozoic strata in the southwest Taoudeni Basin, Guinea and Senegal, and from the Mauritanide fold belt, Mauritania, have yielded mostly poorly preserved small skeletal fossils of metazoan and protistan origin. Problematic, but possible echinoderm material and spicules of the heteractinid sponge Eiffelia dominate the Taoudeni Basin assemblage. The age of the material is not certain but the paleontologic data suggest an Early Cambrian age for the stratigraphically lowest faunas, and a Middle Cambrian age is possible for the stratigraphically highest collections.

  20. Early Cambrian Pentamerous Cubozoan Embryos from South China

    PubMed Central

    Han, Jian; Kubota, Shin; Li, Guoxiang; Yao, Xiaoyong; Yang, Xiaoguang; Shu, Degan; Li, Yong; Kinoshita, Shunichi; Sasaki, Osamu; Komiya, Tsuyoshi; Yan, Gang

    2013-01-01

    Background Extant cubozoans are voracious predators characterized by their square shape, four evenly spaced outstretched tentacles and well-developed eyes. A few cubozoan fossils are known from the Middle Cambrian Marjum Formation of Utah and the well-known Carboniferous Mazon Creek Formation of Illinois. Undisputed cubozoan fossils were previously unknown from the early Cambrian; by that time probably all representatives of the living marine phyla, especially those of basal animals, should have evolved. Methods Microscopic fossils were recovered from a phosphatic limestone in the Lower Cambrian Kuanchuanpu Formation of South China using traditional acetic-acid maceration. Seven of the pre-hatched pentamerous cubozoan embryos, each of which bears five pairs of subumbrellar tentacle buds, were analyzed in detail through computed microtomography (Micro-CT) and scanning electron microscopy (SEM) without coating. Results The figured microscopic fossils are unequivocal pre-hatching embryos based on their spherical fertilization envelope and the enclosed soft-tissue that has preserved key anatomical features arranged in perfect pentaradial symmetry, allowing detailed comparison with modern cnidarians, especially medusozoans. A combination of features, such as the claustrum, gonad-lamella, suspensorium and velarium suspended by the frenula, occur exclusively in the gastrovascular system of extant cubozoans, indicating a cubozoan affinity for these fossils. Additionally, the interior anatomy of these embryonic cubozoan fossils unprecedentedly exhibits the development of many new septum-derived lamellae and well-partitioned gastric pockets unknown in living cubozoans, implying that ancestral cubozoans had already evolved highly specialized structures displaying unexpected complexity at the dawn of the Cambrian. The well-developed endodermic lamellae and gastric pockets developed in the late embryonic stages of these cubozoan fossils are comparable with extant pelagic

  1. Carbon isotopic evidence for photosynthesis in Early Cambrian oceans

    NASA Astrophysics Data System (ADS)

    Surge, Donna M.; Savarese, Michael; Dodd, J. Robert; Lohmann, Kyger C.

    1997-06-01

    Were the first metazoan reefs ecologically similar to modern tropical reefs, enabling them to persist under oligotrophic conditions? We tested the hypothesis of ecological similarity by employing a geochemical approach. Petrography, cathodoluminescence, trace elements, and stable isotope analyses of primary precipitates of the Lower Cambrian Ajax Limestone, South Australia, indicate preservation of original C isotopic composition. All primary carbonate components exhibit C isotopic values similar to the composition of inorganically precipitated fibrous marine cements, suggesting that archaeocyaths and the calcimicrobe Epiphyton precipitated skeletal carbonate in equilibrium with ambient seawater in the absence of vital effects. Such data do not support the contention that archaeocyaths possessed photosymbionts. However, a +0.55‰ shift in δ13C occurs in reefs developed under shallower-water conditions relative to deeper reefs. This shift suggests the stratification of primary production in Early Cambrian oceans. The pattern is similar to that seen in the modern ocean, whereby significant photosynthesis modulates the C isotopic composition of the photic zone.

  2. Paleomagnetism of Early Cambrian Itabaiana mafic dikes (NE Brazil) and the final assembly of Gondwana

    NASA Astrophysics Data System (ADS)

    Trindade, Ricardo I. F.; D'Agrella-Filho, Manoel S.; Epof, Igor; Brito Neves, Benjamim B.

    2006-04-01

    Paleomagnetic analysis on 15 early Cambrian mafic dikes from Itabaiana (Paraíba State) yielded a southern (northwestern) direction with steep upward (downward) inclination ( Dm = 167.5°, Im = - 63.7°, α95 = 7.3°). AF and Thermal demagnetization, thermomagnetic curves, and hysteresis results suggest that this component is dominantly carried by fine-grained SD magnetite. The high stability of this component and positive baked contact tests on three dikes indicate it represents a primary thermoremanent magnetization. Ar-Ar analysis on whole-rock samples from two sites provides a strong constraint on the age of the Itabaiana paleomagnetic pole (134.6° E, 34.9° S; A95 = 7.3, K = 28) defined by plateau ages of 525 ± 5 and 526 ± 4 Ma. This pole completely satisfies six out of the seven quality criteria proposed by Van der Voo [R. Van der Voo, The reliability of paleomagnetic data, Tectonophysics 184 (1990) 1-9.] and permits a tight constraint on the Early Cambrian sector of the Gondwana apparent polar wander path. Paleogeographic reconstructions consistent with the available paleomagnetic and geological record show that Gondwana was sutured along three major orogenies, the Mozambique (Brasilano/Pan-African) Orogeny (800-650 Ma), the Kuunga Orogeny (570-530 Ma) and the Pampean-Araguaia Orogeny (540-520 Ma). We suggest that after rifting away from Laurentia at the end of the Neoproterozoic, opening the Iapetus ocean, the Amazonian craton and minor adjoining blocks, such as Rio Apa and Pampia, collided with the proto-Gondwana by Cambrian times at ca. 530-520 Ma. Unless for small adjustments, Gondwana was completely formed by 525 Ma whose paleogeography is defined by the Itabaiana pole.

  3. The origin of crustaceans: new evidence from the Early Cambrian of China.

    PubMed Central

    Chen, J. Y.; Vannier, J.; Huang, D. Y.

    2001-01-01

    One of the smallest arthropods recently discovered in the Early Cambrian Maotianshan Shale Lagerstätte is described. Ercaia gen. nov. has an untagmatized trunk bearing serially repeated biramous appendages (long and segmented endopods and flap-like exopods), a head with an acron bearing stalked lateral eyes and a sclerite and two pairs of antennae. The position of this 520 million-year-old tiny arthropod within the Crustacea is supported by several anatomical features: (i) a head with five pairs of appendages including two pairs of antennae, (ii) highly specialized antennae (large setose fans with a possible function in feeding), and (iii) specialized last trunk appendages (segmented pediform structures fringed with setae). The segmentation pattern of Ercaia (5 head and 13 trunk) is close to that of Maxillopoda but lacks the trunk tagmosis of modern representatives of the group. Ercaia is interpreted as a possible derivative of the stem group Crustacea. Ercaia is likely to have occupied an ecological niche similar to those of some Recent meiobenthic organisms (e.g. copepods living in association with sediment). This new fossil evidence supports the remote ancestry of crustaceans well before the Late Cambrian and shows, along with other fossil data (mainly Early Cambrian in China), that a variety of body plans already coexisted among the primitive crustacean stock. PMID:11674864

  4. Preservation of Early Cambrian animals of the Chengjiang biota

    NASA Astrophysics Data System (ADS)

    Gabbott, Sarah E.; Xian-Guang, Hou; Norry, Michael J.; Siveter, David J.

    2004-10-01

    The Chengjiang biota of Yunnan, China, documents the earliest extensive radiation of the Metazoa recorded in the fossil record. Gauging preservational bias is crucial in providing an assessment of the completeness of this biota and thereby elucidating whether it represents a comprehensive depiction of Early Cambrian life. We here present a model to explain the nature of the exceptional preservation of the Chengjiang biota and details of the decay process. This study indicates that Chengjiang fossils were preserved through two taphonomic pathways that may have captured tissues of distinct compositions, and this finding should provide a foundation for the interpretation of Chengjiang fossils. Many Chengjiang fossils are preserved by pyrite (later pseudomorphed by iron oxides); the clay-rich host sediment was deficient in organic carbon but replete in available Fe, and this composition ensured that a decaying carcass acted as a local substrate for Fe- and S-reducing bacteria. Pyrite morphology probably reflects contrasts in the decay rate, and hence the H2S production rate, of different tissues in a carcass. Reactive, rapidly decaying tissues would have quickly supplied H2S, producing many pyrite nuclei, resulting in framboidal habits. More recalcitrant tissues would have produced H2S more slowly, so that crystal growth operated on fewer nuclei, resulting in larger euhedral pyrite crystals. Reflective films, especially common on Chengjiang arthropods, represent the remains of degraded carbon.

  5. Exceptionally preserved Cambrian loriciferans and the early animal invasion of the meiobenthos.

    PubMed

    Harvey, Thomas H P; Butterfield, Nicholas J

    2017-01-30

    Microscopic animals that live among and between sediment grains (meiobenthic metazoans) are key constituents of modern aquatic ecosystems, but are effectively absent from the fossil record. We describe an assemblage of microscopic fossil loriciferans (Ecdysozoa, Loricifera) from the late Cambrian Deadwood Formation of western Canada. The fossils share a characteristic head structure and minute adult body size (~300 μm) with modern loriciferans, indicating the early evolution and subsequent conservation of an obligate, permanently meiobenthic lifestyle. The unsuspected fossilization potential of such small animals in marine mudstones offers a new search image for the earliest ecdysozoans and other animals, although the anatomical complexity of loriciferans points to their evolutionary miniaturization from a larger-bodied ancestor. The invasion of animals into ecospace that was previously monopolized by protists will have contributed considerably to the revolutionary geobiological feedbacks of the Proterozoic/Phanerozoic transition.

  6. Fossils, molecules and embryos: new perspectives on the Cambrian explosion

    NASA Technical Reports Server (NTRS)

    Valentine, J. W.; Jablonski, D.; Erwin, D. H.

    1999-01-01

    The Cambrian explosion is named for the geologically sudden appearance of numerous metazoan body plans (many of living phyla) between about 530 and 520 million years ago, only 1.7% of the duration of the fossil record of animals. Earlier indications of metazoans are found in the Neoproterozic; minute trails suggesting bilaterian activity date from about 600 million years ago. Larger and more elaborate fossil burrows appear near 543 million years ago, the beginning of the Cambrian Period. Evidence of metazoan activity in both trace and body fossils then increased during the 13 million years leading to the explosion. All living phyla may have originated by the end of the explosion. Molecular divergences among lineages leading to phyla record speciation events that have been earlier than the origins of the new body plans, which can arise many tens of millions of years after an initial branching. Various attempts to date those branchings by using molecular clocks have disagreed widely. While the timing of the evolution of the developmental systems of living metazoan body plans is still uncertain, the distribution of Hox and other developmental control genes among metazoans indicates that an extensive patterning system was in place prior to the Cambrian. However, it is likely that much genomic repatterning occurred during the Early Cambrian, involving both key control genes and regulators within their downstream cascades, as novel body plans evolved.

  7. Unlocking the early fossil record of the arthropod central nervous system

    PubMed Central

    Edgecombe, Gregory D.; Ma, Xiaoya; Strausfeld, Nicholas J.

    2015-01-01

    Extant panarthropods (euarthropods, onychophorans and tardigrades) are hallmarked by stunning morphological and taxonomic diversity, but their central nervous systems (CNS) are relatively conserved. The timing of divergences of the ground pattern CNS organization of the major panarthropod clades has been poorly constrained because of a scarcity of data from their early fossil record. Although the CNS has been documented in three-dimensional detail in insects from Cenozoic ambers, it is widely assumed that these tissues are too prone to decay to withstand other styles of fossilization or geologically older preservation. However, Cambrian Burgess Shale-type compressions have emerged as sources of fossilized brains and nerve cords. CNS in these Cambrian fossils are preserved as carbon films or as iron oxides/hydroxides after pyrite in association with carbon. Experiments with carcasses compacted in fine-grained sediment depict preservation of neural tissue for a more prolonged temporal window than anticipated by decay experiments in other media. CNS and compound eye characters in exceptionally preserved Cambrian fossils predict divergences of the mandibulate and chelicerate ground patterns by Cambrian Stage 3 (ca 518 Ma), a dating that is compatible with molecular estimates for these splits. PMID:26554038

  8. Decoupling biogeochemical records, extinction, and environmental change during the Cambrian SPICE event

    PubMed Central

    Schiffbauer, James D.; Huntley, John Warren; Fike, David A.; Jeffrey, Matthew Jarrell; Gregg, Jay M.; Shelton, Kevin L.

    2017-01-01

    Several positive carbon isotope excursions in Lower Paleozoic rocks, including the prominent Upper Cambrian Steptoean Positive Carbon Isotope Excursion (SPICE), are thought to reflect intermittent perturbations in the hydrosphere-biosphere system. Models explaining these secular changes are abundant, but the synchronicity and regional variation of the isotope signals are not well understood. Examination of cores across a paleodepth gradient in the Upper Cambrian central Missouri intrashelf basin (United States) reveals a time-transgressive, facies-dependent nature of the SPICE. Although the SPICE event may be a global signal, the manner in which it is recorded in rocks should and does vary as a function of facies and carbonate platform geometry. We call for a paradigm shift to better constrain facies, stratigraphic, and biostratigraphic architecture and to apply these observations to the variability in magnitude, stratigraphic extent, and timing of the SPICE signal, as well as other biogeochemical perturbations, to elucidate the complex processes driving the ocean-carbonate system. PMID:28275734

  9. Three-dimensionally preserved minute larva of a great-appendage arthropod from the early Cambrian Chengjiang biota

    NASA Astrophysics Data System (ADS)

    Liu, Yu; Melzer, Roland R.; Haug, Joachim T.; Haug, Carolin; Briggs, Derek E. G.; Hörnig, Marie K.; He, Yu-yang; Hou, Xian-guang

    2016-05-01

    A three-dimensionally preserved 2-mm-long larva of the arthropod Leanchoilia illecebrosa from the 520-million-year-old early Cambrian Chengjiang biota of China represents the first evidence, to our knowledge, of such an early developmental stage in a short-great-appendage (SGA) arthropod. The larva possesses a pair of three-fingered great appendages, a hypostome, and four pairs of well-developed biramous appendages. More posteriorly, a series of rudimentary limb Anlagen revealed by X-ray microcomputed tomography shows a gradient of decreasing differentiation toward the rear. This, and postembryonic segment addition at the putative growth zone, are features of late-stage metanauplii of eucrustaceans. L. illecebrosa and other SGA arthropods, however, are considered representative of early chelicerates or part of the stem lineage of all euarthropods. The larva of an early Cambrian SGA arthropod with a small number of anterior segments and their respective appendages suggests that posthatching segment addition occurred in the ancestor of Euarthropoda.

  10. Ichnological evidence for meiofaunal bilaterians from the terminal Ediacaran and earliest Cambrian of Brazil.

    PubMed

    Parry, Luke A; Boggiani, Paulo C; Condon, Daniel J; Garwood, Russell J; Leme, Juliana de M; McIlroy, Duncan; Brasier, Martin D; Trindade, Ricardo; Campanha, Ginaldo A C; Pacheco, Mírian L A F; Diniz, Cleber Q C; Liu, Alexander G

    2017-10-01

    The evolutionary events during the Ediacaran-Cambrian transition (~541 Myr ago) are unparalleled in Earth history. The fossil record suggests that most extant animal phyla appeared in a geologically brief interval, with the oldest unequivocal bilaterian body fossils found in the Early Cambrian. Molecular clocks and biomarkers provide independent estimates for the timing of animal origins, and both suggest a cryptic Neoproterozoic history for Metazoa that extends considerably beyond the Cambrian fossil record. We report an assemblage of ichnofossils from Ediacaran-Cambrian siltstones in Brazil, alongside U-Pb radioisotopic dates that constrain the age of the oldest specimens to 555-542 Myr. X-ray microtomography reveals three-dimensionally preserved traces ranging from 50 to 600 μm in diameter, indicative of small-bodied, meiofaunal tracemakers. Burrow morphologies suggest they were created by a nematoid-like organism that used undulating locomotion to move through the sediment. This assemblage demonstrates animal-sediment interactions in the latest Ediacaran period, and provides the oldest known fossil evidence for meiofaunal bilaterians. Our discovery highlights meiofaunal ichnofossils as a hitherto unexplored window for tracking animal evolution in deep time, and reveals that both meiofaunal and macrofaunal bilaterians began to explore infaunal niches during the late Ediacaran.

  11. Paleo-environmental conditions of the Early Cambrian Niutitang Formation in the Fenggang area, the southwestern margin of the Yangtze Platform, southern China: Evidence from major elements, trace elements and other proxies

    NASA Astrophysics Data System (ADS)

    Li, Jin; Tang, Shuheng; Zhang, Songhang; Xi, Zhaodong; Yang, Ning; Yang, Guoqiao; Li, Lei; Li, Yanpeng

    2018-06-01

    The Precambrian/Cambrian transition was a key time in Earth history, especially for marine biological evolution and oceanic chemistry. The redox-stratification with oxic shallow water and anoxic (even euxinic) deeper water in the Early Cambrian Yangtze Sea, which gradually became completely oxygenated, has been suggested as a possible trigger for the "Cambrian explosion" of biological diversity. However, for some areas in northern Guizhou where the exploration and research are lacking, identifying this pattern of redox-stratification by paleo-environmental analysis from borehole data is still in need. Here, we report a remarkable variation range in trace elements (Mo, V, U, Ni, Th, Co, Sc, Zn and Cu), molar Corg:P ratios and pyrite morphology from 27 core samples from one new drill hole (XY1, located in the Fenggang area, northern Guizhou) on the Yangtze Platform, South China. High levels of Ba (from 3242 ppm to 33,800 ppm) and total organic carbon (TOC; from 4% to 9.36%) in 15 core samples in the Lower Member (LM) of the Niutitang Formation indicated elevated primary productivity in the study area. Redox change was recorded based on enrichment factors (EFs) for RSTEs (Mo, U, and V), redox proxies (V/(V + Ni), Ni/Co, V/Sc and Th/U), Corg:P ratios and particle size of framboidal pyrite. These signatures demonstrate that the LM was deposited under anoxic conditions with sulfidic episodes, whereas the Upper Member (UM) of the Niutitang Formation was deposited under suboxic/oxic conditions with intermittently anoxic episodes. Mo/TOC ratios (from 3.72 to 39.86, mean 18.76) suggest weak-moderate water mass restriction. Mo-U covariation patterns (strong but variable enrichment of Mo and U; MoEF ranging from 31.45 to 257.97; UEF ranging from 4.68 to 39.07) in the LM show alternation of particulate shuttling and redox conditions occurred in the Early Cambrian Yangtze Sea, whereas Mo-U covariation patterns (moderate Mo enrichment but depletion or non-enrichment of U; mean Mo

  12. The impact of deep-tier burrow systems in sediment mixing and ecosystem engineering in early Cambrian carbonate settings

    PubMed Central

    Zhang, Li-Jun; Qi, Yong-An; Buatois, Luis A.; Mángano, M. Gabriela; Meng, Yao; Li, Da

    2017-01-01

    Bioturbation plays a substantial role in sediment oxygen concentration, chemical cycling, regeneration of nutrients, microbial activity, and the rate of organic matter decomposition in modern oceans. In addition, bioturbators are ecosystem engineers which promote the presence of some organisms, while precluding others. However, the impact of bioturbation in deep time remains controversial and limited sediment mixing has been indicated for early Paleozoic seas. Our understanding of the actual impact of bioturbation early in the Phanerozoic has been hampered by the lack of detailed analysis of the functional significance of specific burrow architectures. Integration of ichnologic and sedimentologic evidence from North China shows that deep-tier Thalassinoides mazes occur in lower Cambrian nearshore carbonate sediments, leading to intense disruption of the primary fabric. Comparison with modern studies suggest that some of the effects of this style of Cambrian bioturbation may have included promotion of nitrogen and ammonium fluxes across the sediment-water interface, average deepening of the redox discontinuity surface, expansion of aerobic bacteria, and increase in the rate of organic matter decomposition and the regeneration of nutrients. Our study suggests that early Cambrian sediment mixing in carbonate settings may have been more significant than assumed in previous models. PMID:28374857

  13. The early Cambrian fossil embryo Pseudooides is a direct-developing cnidarian, not an early ecdysozoan.

    PubMed

    Duan, Baichuan; Dong, Xi-Ping; Porras, Luis; Vargas, Kelly; Cunningham, John A; Donoghue, Philip C J

    2017-12-20

    Early Cambrian Pseudooides prima has been described from embryonic and post-embryonic stages of development, exhibiting long germ-band development. There has been some debate about the pattern of segmentation, but this interpretation, as among the earliest records of ecdysozoans, has been generally accepted. Here, we show that the 'germ band' of P. prima embryos separates along its mid axis during development, with the transverse furrows between the 'somites' unfolding into the polar aperture of the ten-sided theca of Hexaconularia sichuanensis , conventionally interpreted as a scyphozoan cnidarian; co-occurring post-embryonic remains of ecdysozoans are unrelated. We recognize H. sichuanensis as a junior synonym of P. prima as a consequence of identifying these two form-taxa as distinct developmental stages of the same organism. Direct development in P. prima parallels the co-occuring olivooids Olivooides, and Quadrapyrgites and Bayesian phylogenetic analysis of a novel phenotype dataset indicates that, despite differences in their tetra-, penta- and pseudo-hexa-radial symmetry, these hexangulaconulariids comprise a clade of scyphozoan medusozoans, with Arthrochites and conulariids, that all exhibit direct development from embryo to thecate polyp. The affinity of hexangulaconulariids and olivooids to extant scyphozoan medusozoans indicates that the prevalence of tetraradial symmetry and indirect development are a vestige of a broader spectrum of body-plan symmetries and developmental modes that was manifest in their early Phanerozoic counterparts. © 2017 The Authors.

  14. The early Cambrian fossil embryo Pseudooides is a direct-developing cnidarian, not an early ecdysozoan

    PubMed Central

    2017-01-01

    Early Cambrian Pseudooides prima has been described from embryonic and post-embryonic stages of development, exhibiting long germ-band development. There has been some debate about the pattern of segmentation, but this interpretation, as among the earliest records of ecdysozoans, has been generally accepted. Here, we show that the ‘germ band’ of P. prima embryos separates along its mid axis during development, with the transverse furrows between the ‘somites’ unfolding into the polar aperture of the ten-sided theca of Hexaconularia sichuanensis, conventionally interpreted as a scyphozoan cnidarian; co-occurring post-embryonic remains of ecdysozoans are unrelated. We recognize H. sichuanensis as a junior synonym of P. prima as a consequence of identifying these two form-taxa as distinct developmental stages of the same organism. Direct development in P. prima parallels the co-occuring olivooids Olivooides, and Quadrapyrgites and Bayesian phylogenetic analysis of a novel phenotype dataset indicates that, despite differences in their tetra-, penta- and pseudo-hexa-radial symmetry, these hexangulaconulariids comprise a clade of scyphozoan medusozoans, with Arthrochites and conulariids, that all exhibit direct development from embryo to thecate polyp. The affinity of hexangulaconulariids and olivooids to extant scyphozoan medusozoans indicates that the prevalence of tetraradial symmetry and indirect development are a vestige of a broader spectrum of body-plan symmetries and developmental modes that was manifest in their early Phanerozoic counterparts. PMID:29237861

  15. Early Cambrian wave-formed shoreline deposits: the Hardeberga Formation, Bornholm, Denmark

    NASA Astrophysics Data System (ADS)

    Clemmensen, Lars B.; Glad, Aslaug C.; Pedersen, Gunver K.

    2017-09-01

    During the early Cambrian, the Danish island Bornholm was situated on the northern edge of the continent Baltica with palaeolatitudes of about 35°S. An early Cambrian (Terreneuvian) transgression inundated large areas of Baltica including Bornholm creating shallow marine and coastline environments. During this period, wave-formed shoreline sediments (the Vik Member, Hardeberga Formation) were deposited on Bornholm and are presently exposed at Strøby quarry. The sediments consist of fine- and medium-grained quartz-cemented arenites in association with a few silt-rich mudstones. The presence of well-preserved subaqueous dunes and wave ripples indicates deposition in a wave-dominated upper shoreface (littoral zone) environment, and the presence of interference ripples indicates that the littoral zone environment experienced water level fluctuations due to tides and/or changing meteorological conditions. Discoidal structures (medusoids) are present in the quarry, but due to the relative poor preservation of their fine-scale structures it is difficult to determine if the discoids represent true medusae imprints or inorganic structures. The preservation of the shallow-water bedforms as well as the possible medusae imprints is related to either the formation of thin mud layers, formed during a period of calm water when winds blew offshore for a longer period, or to the growth of bacterial mats. The orientation of the wave-formed bedforms indicates a local palaeoshoreline trending NE-SW and facing a large ocean to the north.

  16. Late Cambrian - Early Ordovician turbidites of Gorny Altai (Russia): Compositions, sources, deposition settings, and tectonic implications

    NASA Astrophysics Data System (ADS)

    Kruk, Nikolai N.; Kuibida, Yana V.; Shokalsky, Sergey P.; Kiselev, Vladimir I.; Gusev, Nikolay I.

    2018-06-01

    The Cambrian-Ordovician transition was the time of several key events in the history of Central Asia. They were the accretion of Mariana-type island arc systems to the Siberian continent, the related large-scale orogeny and intrusions of basaltic and granitic magma and the formation of a huge turbidite basin commensurate with the Bengal Gulf basin in the western part of the Central Asian orogenic belt (CAOB). The structure of the basin, as well as the sources and environments of deposition remain open to discussion. This paper presents new major- and trace-element data on Late-Cambrian-Early Ordovician turbidites from different parts of the Russian Altai and a synthesis of Nd isotope composition and ages of detrital zircons. The turbidites share chemical similarity with material shed from weathered continental arcs. Broad variations of CIA (39-73) and ICV (0.63-1.66) signatures in sandstones suggest origin from diverse sources and absence of significant sorting. Trace elements vary considerably and have generally similar patterns in rocks from different terranes. On the other hand, there are at least two provinces according to Nd isotope composition and age of detrital zircons. Samples from eastern Russian Altai contain only Phanerozoic zircons and have Nd isotope ratios similar to those in Early Cambrian island arcs (εNdt + 4.4… + 5.4; TNd(DM)-2-st = 0.8-0.9 Ga). Samples from central, western, and southern parts of Russian Altai contain Precambrian zircons (some as old as Late Archean) and have a less radiogenic Nd composition (εNdt up to -3.6; TNd(DM)-2-st up to 1.5 Ga). The chemical signatures of Late Cambrian to Early Ordovician turbidites indicate a provenance chemically more mature than the island arc rocks, and the presence of zircons with 510-490 Ma ages disproves their genetic relation with island arcs. The turbidite basin formed simultaneously with peaks of granitic and alkali-basaltic magmatism in the western Central Asian orogen and resulted from

  17. Diversity partitioning during the Cambrian radiation

    PubMed Central

    Na, Lin; Kiessling, Wolfgang

    2015-01-01

    The fossil record offers unique insights into the environmental and geographic partitioning of biodiversity during global diversifications. We explored biodiversity patterns during the Cambrian radiation, the most dramatic radiation in Earth history. We assessed how the overall increase in global diversity was partitioned between within-community (alpha) and between-community (beta) components and how beta diversity was partitioned among environments and geographic regions. Changes in gamma diversity in the Cambrian were chiefly driven by changes in beta diversity. The combined trajectories of alpha and beta diversity during the initial diversification suggest low competition and high predation within communities. Beta diversity has similar trajectories both among environments and geographic regions, but turnover between adjacent paleocontinents was probably the main driver of diversification. Our study elucidates that global biodiversity during the Cambrian radiation was driven by niche contraction at local scales and vicariance at continental scales. The latter supports previous arguments for the importance of plate tectonics in the Cambrian radiation, namely the breakup of Pannotia. PMID:25825755

  18. A Cambrian mixed carbonate-siliciclastic platform in SW Gondwana: evidence from the Western Sierras Pampeanas (Argentina) and implications for the early Paleozoic paleogeography of the proto-Andean margin

    NASA Astrophysics Data System (ADS)

    Ramacciotti, Carlos D.; Casquet, César; Baldo, Edgardo G.; Galindo, Carmen; Pankhurst, Robert J.; Verdecchia, Sebastián O.; Rapela, Carlos W.; Fanning, Mark

    2018-05-01

    The Western Sierras Pampeanas (WSP) of Argentina record a protracted geological history from the Mesoproterozoic assembly of the Rodinia supercontinent to the early Paleozoic tectonic evolution of SW Gondwana. Two well-known orogenies took place at the proto-Andean margin of Gondwana in the Cambrian and the Ordovician, i.e., the Pampean (545-520 Ma) and Famatinian (490-440 Ma) orogenies, respectively. Between them, an extensive continental platform was developed, where mixed carbonate-siliciclastic sedimentation occurred. This platform was later involved in the Famatinian orogeny when it underwent penetrative deformation and metamorphism. The platform apparently extended from Patagonia to northwestern Argentina and the Eastern Sierras Pampeanas, and has probable equivalents in SW Africa, Peru, and Bolivia. The WSP record the outer (deepest) part of the platform, where carbonates were deposited in addition to siliciclastic sediments. Detrital zircon U-Pb SHRIMP ages from clastic metasedimentary successions and Sr-isotope compositions of marbles from the WSP suggest depositional ages between ca. 525 and 490 Ma. The detrital zircon age patterns further suggest that clastic sedimentation took place in two stages. The first was sourced mainly from re-working of the underlying Neoproterozoic metasedimentary rocks and the uplifted core of the early Cambrian Pampean orogen, without input from the Paleoproterozoic Río de la Plata craton. Sediments of the second stage resulted from the erosion of the still emerged Pampean belt and the Neoproterozoic Brasiliano orogen in the NE with some contribution from the Río de la Plata craton. An important conclusion is that the WSP basement was already part of SW Gondwana in the early Cambrian, and not part of the exotic Precordillera/Cuyania terrane, as was previously thought.

  19. Proterozoic and early Cambrian protists: evidence for accelerating evolutionary tempo

    NASA Technical Reports Server (NTRS)

    Knoll, A. H.

    1994-01-01

    In rocks of late Paleoproterozoic and Mesoproterozoic age (ca. 1700-1000 million years ago), probable eukaryotic microfossils are widespread and well preserved, but assemblage and global diversities are low and turnover is slow. Near the Mesoproterozoic-Neoproterozoic boundary (1000 million years ago), red, green, and chromophytic algae diversified; molecular phylogenies suggest that this was part of a broader radiation of "higher" eukaryotic phyla. Observed diversity levels for protistan microfossils increased significantly at this time, as did turnover rates. Coincident with the Cambrian radiation of marine invertebrates, protistan microfossils again doubled in diversity and rates of turnover increased by an order of magnitude. Evidently, the Cambrian diversification of animals strongly influenced evolutionary rates, within clades already present in marine communities, implying an important role for ecology in fueling a Cambrian explosion that extends across kingdoms.

  20. Transient deep-water oxygenation in the early Cambrian Nanhua Basin, South China

    NASA Astrophysics Data System (ADS)

    Cheng, Meng; Li, Chao; Zhou, Lian; Feng, LianJun; Algeo, Thomas J.; Zhang, FeiFei; Romaniello, Stephen; Jin, ChengSheng; Ling, HongFei; Jiang, ShaoYong

    2017-08-01

    Many late Neoproterozoic to early Cambrian fossils of multicellular eukaryotes, including those of benthic animals, are found preserved under anoxic and even euxinic bottom-water conditions, which is contradictory to the consensus that oxygen is essential to eukaryotes. To investigate this conundrum, we conducted an integrated study of iron speciation, redox-sensitive trace elements, and Mo isotopes (δ98Mo) on the black shale interval of the lower Cambrian Hetang Formation (∼535-521 Ma) at Lantian, South China, in which benthic sponge fossils are abundant in the lower member (LM) but absent in the upper member (UM). Iron speciation data point to uniformly anoxic-ferruginous conditions in the LM and euxinic conditions in the UM, whereas the trace-element and δ98Mo data show greater secular variation in redox conditions. The LM shows higher mean trace element concentrations (Mo: 108 ppm, U: 36 ppm, V: 791 ppm) and lower and more variable δ98Mo (+0.13 to +1.76‰) relative to the UM (Mo: 45 ppm, U: 18 ppm, V: 265 ppm, δ98Mo: +1.59 to +1.67‰), and ratios of redox-sensitive trace element concentrations to total organic carbon are significantly more variable and higher on average in the LM relative to the UM. The appearance of sponge fossils and lower δ98Mo values correlate strongly with gray (i.e., lighter-colored) layers in the LM. These patterns can best be interpreted as recording mainly euxinic conditions throughout deposition of the study units, with more intense background euxinia in the LM relative to the UM, but also with frequent transient oxygenation events in the LM that do not appear in the UM. The transient oxygenation events of the LM led to the initial colonization of the deep Nanhua Basin by sponges, and the termination of these events in the UM caused sponge faunas to disappear until a general rise in O2 levels later in the Cambrian permitted their return to deeper-water habitats. Our study also illustrates that multiple geochemical and

  1. Cambrian lobopodians and extant onychophorans provide new insights into early cephalization in Panarthropoda

    PubMed Central

    Ou, Qiang; Shu, Degan; Mayer, Georg

    2012-01-01

    Cambrian lobopodians are important for understanding the evolution of arthropods, but despite their soft-bodied preservation, the organization of the cephalic region remains obscure. Here we describe new material of the early Cambrian lobopodian Onychodictyon ferox from southern China, which reveals hitherto unknown head structures. These include a proboscis with a terminal mouth, an anterior arcuate sclerite, a pair of ocellus-like eyes and branched, antenniform appendages associated with this ocular segment. These findings, combined with a comparison with other lobopodians, suggest that the head of the last common ancestor of fossil lobopodians and extant panarthropods comprized a single ocular segment with a proboscis and terminal mouth. The lack of specialized mouthparts in O. ferox and the involvement of non-homologous mouthparts in onychophorans, tardigrades and arthropods argue against a common origin of definitive mouth openings among panarthropods, whereas the embryonic stomodaeum might well be homologous at least in Onychophora and Arthropoda. PMID:23232391

  2. Proterozoic and early Cambrian protists: evidence for accelerating evolutionary tempo.

    PubMed Central

    Knoll, A H

    1994-01-01

    In rocks of late Paleoproterozoic and Mesoproterozoic age (ca. 1700-1000 million years ago), probable eukaryotic microfossils are widespread and well preserved, but assemblage and global diversities are low and turnover is slow. Near the Mesoproterozoic-Neoproterozoic boundary (1000 million years ago), red, green, and chromophytic algae diversified; molecular phylogenies suggest that this was part of a broader radiation of "higher" eukaryotic phyla. Observed diversity levels for protistan microfossils increased significantly at this time, as did turnover rates. Coincident with the Cambrian radiation of marine invertebrates, protistan microfossils again doubled in diversity and rates of turnover increased by an order of magnitude. Evidently, the Cambrian diversification of animals strongly influenced evolutionary rates, within clades already present in marine communities, implying an important role for ecology in fueling a Cambrian explosion that extends across kingdoms. Images PMID:8041692

  3. A superarmored lobopodian from the Cambrian of China and early disparity in the evolution of Onychophora

    PubMed Central

    Yang, Jie; Ortega-Hernández, Javier; Gerber, Sylvain; Butterfield, Nicholas J.; Hou, Jin-bo; Lan, Tian; Zhang, Xi-guang

    2015-01-01

    We describe Collinsium ciliosum from the early Cambrian Xiaoshiba Lagerstätte in South China, an armored lobopodian with a remarkable degree of limb differentiation including a pair of antenna-like appendages, six pairs of elongate setiferous limbs for suspension feeding, and nine pairs of clawed annulated legs with an anchoring function. Collinsium belongs to a highly derived clade of lobopodians within stem group Onychophora, distinguished by a substantial dorsal armature of supernumerary and biomineralized spines (Family Luolishaniidae). As demonstrated here, luolishaniids display the highest degree of limb specialization among Paleozoic lobopodians, constitute more than one-third of the overall morphological disparity of stem group Onychophora, and are substantially more disparate than crown group representatives. Despite having higher disparity and appendage complexity than other lobopodians and extant velvet worms, the specialized mode of life embodied by luolishaniids became extinct during the Early Paleozoic. Collinsium and other superarmored lobopodians exploited a unique paleoecological niche during the Cambrian explosion. PMID:26124122

  4. Origin and hydrology of a large, intact Early Cambrian paleocave system and its role in overlying fluidisation structures, Arctic Canada

    NASA Astrophysics Data System (ADS)

    Mathieu, Jordan; Turner, Elizabeth C.; Rainbird, Robert H.

    2017-04-01

    Paleokarst is most commonly expressed as subtle stratigraphic surfaces rather than large void systems penetrating deeply into the paleo-subsurface. In contrast, a regional Precambrian-Cambrian unconformity on Victoria Island (NWT, Canada), is associated with exceptional exposure of large, intact Cambrian paleocaverns (100 m diameter; tens of m high). The paleocaves are distributed along a paleo-horizontal plane, and an associated gryke network is present in the 30-60 m of Neoproterozoic dolostone between cave rooves and the base of overlying Cambrian sandstone; both are filled by Cambrian sandstone. The formation and preservation of such karst features require aggressive dissolution along a stable paleo-water-table shortly before transgression and deposition of shallow-marine sand over the dolostone. During the transgression, the karst network acted as a conduit for flowing groundwater that was discharged through overlying, unconsolidated Cambrian shallow-marine sand, producing water-escape structures (sand volcanoes and their conduits). The conduits are preserved as cylindrical remnants of the sand volcanoes' feeder pipes. Sediment fluidisation was probably caused by variations in the hydraulic-head gradient in a meteoric lens near the Cambrian coastline under a tropical climate with abundant, probably seasonally variable rainfall that caused pulses in subsurface fluid flow. Spatial distribution of the paleocaves and sand volcanoes suggests their formation on the southeast side of a recently faulted horst of Proterozoic carbonate bedrock that formed a nearshore island during early Cambrian sea-level rise. Fluidisation structures such as those reported here have generally been difficult to interpret owing to a lack of data on the fluid hydraulics of the underlying aquifer. This is the first report linking the hydraulics of a well-characterised paleokarst to development of fluid-escape structures. Such structures are widely known from sandstones overlying the sub-Cambrian

  5. A Late Cambrian Carbon Isotope Excursion Recorded in Passive Margin Dolostones of the Central Appalachian Basin, USA.

    NASA Astrophysics Data System (ADS)

    Mackey, J. E.; Stewart, B. W.

    2016-12-01

    A Late Cambrian global positive carbon isotope excursion, known as the SPICE event [1,2] is linked to possible widespread ocean anoxia and enhanced carbon burial [3,4]. We report data from the central Appalachian Conasauga Group from the upper portion of the Middle Cambrian Maryville limestone, through the Late Cambrian Nolichucky shale and Maynardville limestone members. A geochemical, macro-, and micro-scale analyses of core material from southeastern Ohio was carried out to further constrain the timing of oceanic anoxia and trace element geochemistry relative to sediment fluxes occurring at the transition of the Middle to Late Cambrian. The section represents condensed, passive margin shale deposition and carbonate ramp development on the continental shelf of Laurentia. Carbonate sediments (primarily diagenetic dolomite) record a positive δ13C (relative to V-PDB) excursion starting in the upper Nolichucky shale member, reaching its peak (+4.0) in the overlying Maynardville limestone. At this location, there is an offset between the onlap Nolichucky shale deposition and start of the C isotope excursion; this was reported as well in a carbonate section further south of this location [2], on the other side of an extensional feature (Rome Trough) that formed a deep marine basin during Cambrian time. The condensed shale package and relatively low TOC content in our samples is likely due to the combination of a shallow, upslope basin location and isostatic influence on passive margin sedimentation. However, within the Rome Trough, the Nolichucky shale is rich in organic carbon and a recent target of hydrocarbon exploration. The data suggest a possible link between deposition of this shale and the global SPICE event. The robustness of the Late Cambrian δ13C excursion in diagenetically altered sediments and association with hydrocarbon bearing units indicates its utility as a stratigraphic indicator and as a target for exploration. Ongoing geochemical work will focus

  6. Ichnologic evidence of a Cambrian age in the southern Amazon Craton: Implications for the onset of the Western Gondwana history

    NASA Astrophysics Data System (ADS)

    Santos, Hudson P.; Mángano, M. Gabriela; Soares, Joelson L.; Nogueira, Afonso C. R.; Bandeira, José; Rudnitzki, Isaac D.

    2017-07-01

    Colonization of the infaunal ecospace by burrowing bilaterians is one of the most important behavioral innovations during the Ediacaran-Cambrian transition. The establishment of vertical burrows by suspension feeders in high-energy nearshore settings during Cambrian Age 2 is reflected by the appearance of the Skolithos Ichnofacies. For the first time, unquestionable vertical burrows typical of the Skolithos Ichnofacies, such as Skolithos linearis, Diplocraterion parallelum and Arenicolites isp., are recorded from nearshore siliciclastic deposits of the Raizama Formation, southeastern Amazon Craton, Brazil. Integration of ichnologic and sedimentologic datasets suggests that these trace fossils record colonization of high-energy and well-oxygenated nearshore sandy environments. Chronostratigraphically, the presence of these vertical burrows indicates an age not older than early Cambrian for the Raizama Formation, which traditionally has been regarded as Ediacaran. Therefore, the Raizama ichnofauna illustrates the advent of modern Phanerozoic ecology marked by the Agronomic Revolution. The discovery of the Skolithos Ichnofacies in these shallow-marine strata suggests possible connections between some central Western Gondwana basins.

  7. Earth's magnetic field strength in the Early Cambrian: Thellier paleointensity estimates of Itabaiana mafic dykes, Northeast Brazil

    NASA Astrophysics Data System (ADS)

    Trindade, R. I. F.; Macouin, M.; Poitou, C.; Chauvin, A.; Hill, M.

    2012-04-01

    Thellier's paleointensity and microwave paleointensity experiments were carried out in Early Cambrian dykes from Itabaiana (NE Brazil) dated at 525 ±5 Ma. A previous paleomagnetic study on these dykes reveals a very stable characteristic component, whose thermoremanent nature is confirmed by positive baked contact tests performed in three different dykes. The main magnetic carrier is Ti-poor to pure magnetite in the PSD to SD domain state. Hysteresis parameters and first-order reversal curve (FORC) diagrams will be presented in order to apprehend the two different behaviors that characterize the samples during paleointensity experiments. From the 96 samples (from 13 dykes) analyzed in two laboratories using slightly different Thellier's experimental protocols, we have retained 12 samples (3 dykes) for paleointensity estimates. Paleointensity values range from 18.1 up to 40 μΤ. This corresponds to equivalent VDMs of 4.3 ± 0.5, 4.4 ± 1.4 and 5.3 ± 0.9 x 1022 Am2, for the three dykes respectively. These results, the first obtained for rapidly cooled Cambrian rocks, document a moderate Earth field in the Precambrian-Cambrian transition.

  8. Darwin's dilemma: the realities of the Cambrian ‘explosion’

    PubMed Central

    Conway Morris, Simon

    2006-01-01

    The Cambrian ‘explosion’ is widely regarded as one of the fulcrum points in the history of life, yet its origins and causes remain deeply controversial. New data from the fossil record, especially of Burgess Shale-type Lagerstätten, indicate, however, that the assembly of bodyplans is not only largely a Cambrian phenomenon, but can already be documented in fair detail. This speaks against a much more ancient origin of the metazoans, and current work is doing much to reconcile the apparent discrepancies between the fossil record, including the Ediacaran assemblages of latest Neoproterozoic age and molecular ‘clocks’. Hypotheses to explain the Cambrian ‘explosion’ continue to be generated, but the recurrent confusion of cause and effect suggests that the wrong sort of question is being asked. Here I propose that despite its step-like function this evolutionary event is the inevitable consequence of Earth and biospheric change. PMID:16754615

  9. The origin of decoupled carbonate and organic carbon isotope signatures in the early Cambrian (ca. 542-520 Ma) Yangtze platform

    NASA Astrophysics Data System (ADS)

    Jiang, Ganqing; Wang, Xinqiang; Shi, Xiaoying; Xiao, Shuhai; Zhang, Shihong; Dong, Jin

    2012-02-01

    The early Cambrian (ca. 542-520 Ma) strata in South China record two prominent negative carbonate carbon isotope (δ13Ccarb) excursions of early Nemakit-Daldynian (N-D) and early Tommotian ages. Across each of these excursions, carbonate and organic carbon isotopes (δ13Ccarb and δ13Corg) are strongly decoupled. Regional correlation across a shelf-to-basin transect shows lateral heterogeneity of δ13Corg during the early-middle N-D but more homogenized δ13Corg values across the basin during the late N-D and Tommotian. The temporal and lateral variations in δ13Corg suggest that decoupled δ13Ccarb and δ13Corg across the N-D δ13Ccarb excursion were possibly caused by diagenetic alteration of organic matter and/or amplification of detrital organic carbon isotope signature in low-TOC carbonates. In contrast, decoupled δ13Ccarb and δ13Corg of the upper N-D and Tommotian were likely resulted from chemoautotrophic-methanotrophic biomass contribution to TOC in organic-rich black shale and carbonates. The decoupled δ13Ccarb-δ13Corg pattern from the lower N-D strata (ca. 542 Ma) shows striking similarities with those from the basal (ca. 635 Ma) and upper (ca. 551 Ma) Doushantuo Formation. In all three cases, decoupled δ13Ccarb-δ13Corg are seen in organic-poor carbonates (TOC ≤ 0.1‰) and coupled δ13Ccarb-δ13Corg occur in organic-rich black shale and carbonates at the end of the negative δ13Ccarb excursion. These similarities suggest that the shift from decoupled to coupled δ13Ccarb-δ13Corg has no causal link with the terminal oxidation of a large oceanic DOC reservoir. Given the pervasive anoxia/euxinia in Ediacaran-early Cambrian oceans, local DOC-rich environments may have been common, but a large oceanic DOC reservoir capable of buffering the δ13C of marine organic matter requires independent evidence.

  10. Sophisticated digestive systems in early arthropods.

    PubMed

    Vannier, Jean; Liu, Jianni; Lerosey-Aubril, Rudy; Vinther, Jakob; Daley, Allison C

    2014-05-02

    Understanding the way in which animals diversified and radiated during their early evolutionary history remains one of the most captivating of scientific challenges. Integral to this is the 'Cambrian explosion', which records the rapid emergence of most animal phyla, and for which the triggering and accelerating factors, whether environmental or biological, are still unclear. Here we describe exceptionally well-preserved complex digestive organs in early arthropods from the early Cambrian of China and Greenland with functional similarities to certain modern crustaceans and trace these structures through the early evolutionary lineage of fossil arthropods. These digestive structures are assumed to have allowed for more efficient digestion and metabolism, promoting carnivory and macrophagy in early arthropods via predation or scavenging. This key innovation may have been of critical importance in the radiation and ecological success of Arthropoda, which has been the most diverse and abundant invertebrate phylum since the Cambrian.

  11. First U-Pb geochronology on detrital zircons from Early-Middle Cambrian strata of the Torgau-Doberlug Syncline (eastern Germany) and palaeogeographic implications

    NASA Astrophysics Data System (ADS)

    Abubaker, Atnisha; Hofmann, Mandy; Gärtner, Andreas; Linnemann, Ulf; Elicki, Olaf

    2017-10-01

    LA-ICP-MS U-Pb data from detrital zircons of the Ediacaran to Cambrian siliciclastic sequence of the Torgau-Doberlug Syncline (TDS, Saxo-Thuringia, Germany) are reported for the first time. The majority of 203 analysed zircon grains is Proterozoic with minor amount of Archean and Palaeozoic grains. The U-Pb ages fall into three groups: 2.8-2.4 Ga (3%), Neoarchean to earliest Palaeoproterozoic; 2.3-1.6 Ga (46%), early to late Palaeoproterozoic; 1.0-0.5 Ga (47%), Neoproterozoic to Cambrian. This age distribution is typical for the West African Craton as the source area and for Cadomian orogenic events in northwestern Gondwana. The samples show an age gap between 1.6 and 1.0 Ga, which is characteristic for West African provenance and diagnostic in distinguishing this unit from East Avalonia and Baltica. The dataset shows clusters of Palaeoproterozoic ages at 2.2-1.7 Ga, that is typical for western Gondwana, which was affected by abundant magmatic intrusions (ca. 2.2-1.8 Ga) during the Eburnean orogeny (West African craton). Neoarchean zircon ages (3%) point to recycling of magmatic rocks formed during the Liberian and Leonian orogenies. Ediacaran to earliest Cambrian rocks of the TDS originated in an active margin regime of the Gondwanan shelf. The following early Palaeozoic overstep sequence was deposited within rift settings that reflects instability of the West-Gondwanan shelf and the separation of terranes from Ordovician onward. The results of this study demonstrate distinct northwestern African provenance of the Cambrian siliciclastics of the TDS. Due to Th-U ratios from concordant zircon analysis, igneous origin from felsic melts is concluded as the source of these grains.

  12. Marine redox stratification during the early Cambrian (ca. 529-509 Ma) and its control on the development of organic-rich shales in Yangtze Platform

    NASA Astrophysics Data System (ADS)

    Zhang, Yuying; He, Zhiliang; Jiang, Shu; Gao, Bo; Liu, Zhongbao; Han, Bo; Wang, Hu

    2017-06-01

    High resolution geochemical data from nine sections representing shelf to basinal environments in the Yangtze Platform were analyzed to reconstruct the marine redox environment during early Cambrian. Based on Fe species and Mo/TOC ratios, we have supplemented marine redox stratification during Stage 4 (late Canglangpuian-Longwangmiaoan, ˜514-509 Ma) on basis of the previously studied Stage 2-Stage 3 (Meishucunian-Qiongzhusian, ˜529-514 Ma). A new proposed marine stratified redox model indicates that the middepth "euxinic wedge" developed at the base of slope during ˜514-509 Ma in contrast to that the "euxinic wedge" prevailed at the shelf margin during ˜529-514 Ma, even though these middepth euxinic waters both occurred between the oxic surface waters and ferruginous deep waters. This marine redox stratification resulted in high production and good preservation of organic matter during early Cambrian. TOC values in euxinic waters in the middle are generally higher than in ferruginous waters due to upwelling in slope. Therefore, the lower Cambrian organic-rich shales in the Yangtze Platform are inferred to be deposited under the anoxic-ferruginous and euxinic bottom waters with moderate-strong restriction.

  13. Soft-part anatomy of the Early Cambrian bivalved arthropods Kunyangella and Kunmingella: significance for the phylogenetic relationships of Bradoriida

    PubMed Central

    Hou, Xianguang; Williams, Mark; Siveter, David J.; Siveter, Derek J.; Aldridge, Richard J.; Sansom, Robert S.

    2010-01-01

    Bradoriids are small bivalved marine arthropods that are widespread in rocks of Cambrian to Early Ordovician age. They comprise seven families and about 70 genera based on shield (‘carapace’) morphology. New bradoriid specimens with preserved soft-part anatomy of Kunmingella douvillei (Kunmingellidae) are reported from the Early Cambrian Chengjiang Lagerstätte of China together with, for the first time to our knowledge, a second bradoriid species with preserved soft parts, Kunyangella cheni (Comptalutidae). Kunmingella douvillei has a 10-segmented limb-bearing body with uniramous ninth and tenth appendages and a series of homogeneous, apparently (proximal parts not preserved) unspecialized post-antennal biramous limbs with setose leaf-shaped exopods. Each endopod consists of five podomeres. A presumed penultimate instar of Ky. cheni preserves remnants of three head and two trunk appendages, and the adult is reconstructed as having four head appendages. This material allows testing of the affinity of the Bradoriida. Kunmingella is identified as a stem crustacean in character-based analyses, through both morphological comparisons and cladistic reconstructions. Global parsimony analysis recovers a monophyletic Bradoriida as the sister group to crown crustaceans. PMID:20181565

  14. The oceanic carbon cycle implicated in the d13Ccarb and the d13Corg variations from the terminal Ediacaran to the Early Cambrian

    NASA Astrophysics Data System (ADS)

    Ishikawa, T.; Ueno, Y.; Komiya, T.; Shu, D.; Li, Y.; Yoshida, N.; Maruyama, S.

    2009-04-01

    The terminal Neoproterozoic and its transition into the Cambrian witnessed major evolutionary and geochemical changes (e.g. Knoll, 1994). Evolutionary features include extinction and subsequent radiation events (e.g. Brasier, 1994; Knoll, 1994; Knoll and Carroll, 1999; Shu 2008). Geochemical changes comprise secular variations of the global carbon cycle expressed as variations of the d13C isotope records. A representative d13C curve for inorganic carbon (d13Ccarb) across the Precambrian/Cambrian boundary (Pc/C boundary) shows the existence of large fluctuations (e.g. Kirschvink et al., 1991; Narbonne et al., 1994; Kaufman et al., 1995; Amthor et al., 2003; Maloof et al., 2005, Ishikawa et al., 2008). This indicates a significant change of the oceanic carbon cycle at that time. On the other hand, the d13C values for total organic carbon (d13Corg) have rarely been reported together with the d13Ccarb across the boundary. Therefore, the precise relation between the d13Ccarb and the d13Corg and the global carbon cycle at the Pc/C boundary are still ambiguous. This work presents a first high-resolution d13Corg chemostratigraphy of drill core samples across the Pc/C boundary in the Three Gorges area, South China. Based on the results, this work additionally proposes variations of the sizes of the oceanic carbon reservoirs by a calculation of the carbon cycle model at the Pc/C boundary. The Three Gorges section extends from the uppermost Ediacaran dolostone (Dengying Formation), through the lowermost Early Cambrian muddy limestone (Yanjiahe Formation) to the middle Early Cambrian calcareous black shale (Shuijingtuo Formation). The ^13Corg values exhibit relatively invariant values averaging at -31 permil. By comparison between the d13Ccarb and d13Corg, we recognize two different terms in this period. The first term from the Pc/C boundary to the early Nemakit-Daldynian (ND) is characterized by the decoupling of d13Corg and d13Ccarb, stable d13Corg and the significant

  15. Palaeogeography of late Cambrian to early Ordovician sediments in the Amadeus Basin, central Australia

    NASA Astrophysics Data System (ADS)

    Gorter, John D.

    The depositional history of 6 sequences encompassing 18 parasequence of the Late Cambrian to Early Ordovician age in the Amadeus Basin is presented in a seried of generalized paleogeographic maps. As some of the parasequence sets are known to host large deposits of oil and gas, a thorough understanding of the potential reservoir-source rock combinations in the Amadeus Basin is essential for the discovery of further oil and gas reserves in this vast, under-explored basin. The best reservoir rocks in the Pacoota Sandstone are concentrated above the major sequence boundary between the Wallaby and Tempe Vale sequences on the Central Ridge. Poorer reservoirs occur within other sequences (e.g., parasequence set 3 and 13). Parasequence set 3 reservoirs, localized on the Central Ridge, are generally poor but owe their reservoir character to weathering at the pre-Tempe Vale sequence unconformity. Parasequence set 13 reservoirs are also concenterated along the Central Ridge, where small-scale shoaling clastic cycles are better developed. Basal Stairway Sandstone reservoirs in the Mereenie area on the Central Ridge are generally very poor, due to the cementation of the clean sandstone, but should improve to the southwest due to lesser burial-induced silicification. The source potential of the major Arenig organic-rich sediments is concentrated in the transitional zone between parasequence sets 15 and 16. East of West Waterhouse 1 well, these parasequence sets have been eroded and there is no remaining source potential. The transitional source-rich zone is better developed on the Central Ridge than in the Missionary Plain Trough. The Central Ridge is therefore of prime importance in the localization of both reservoir and source rocks in the Late Cambrian and Early Ordovician section of the Amadeus Basin.

  16. Compilation and network analyses of cambrian food webs.

    PubMed

    Dunne, Jennifer A; Williams, Richard J; Martinez, Neo D; Wood, Rachel A; Erwin, Douglas H

    2008-04-29

    A rich body of empirically grounded theory has developed about food webs--the networks of feeding relationships among species within habitats. However, detailed food-web data and analyses are lacking for ancient ecosystems, largely because of the low resolution of taxa coupled with uncertain and incomplete information about feeding interactions. These impediments appear insurmountable for most fossil assemblages; however, a few assemblages with excellent soft-body preservation across trophic levels are candidates for food-web data compilation and topological analysis. Here we present plausible, detailed food webs for the Chengjiang and Burgess Shale assemblages from the Cambrian Period. Analyses of degree distributions and other structural network properties, including sensitivity analyses of the effects of uncertainty associated with Cambrian diet designations, suggest that these early Paleozoic communities share remarkably similar topology with modern food webs. Observed regularities reflect a systematic dependence of structure on the numbers of taxa and links in a web. Most aspects of Cambrian food-web structure are well-characterized by a simple "niche model," which was developed for modern food webs and takes into account this scale dependence. However, a few aspects of topology differ between the ancient and recent webs: longer path lengths between species and more species in feeding loops in the earlier Chengjiang web, and higher variability in the number of links per species for both Cambrian webs. Our results are relatively insensitive to the exclusion of low-certainty or random links. The many similarities between Cambrian and recent food webs point toward surprisingly strong and enduring constraints on the organization of complex feeding interactions among metazoan species. The few differences could reflect a transition to more strongly integrated and constrained trophic organization within ecosystems following the rapid diversification of species, body

  17. Neoproterozoic–Cambrian stratigraphic framework of the Anti-Atlas and Ouzellagh promontory (High Atlas), Morocco

    USGS Publications Warehouse

    Alvaro, Jose Javier; Benziane, Fouad; Thomas, Robert; Walsh, Gregory J.; Yazidi, Abdelaziz

    2014-01-01

    In the last two decades, great progress has been made in the geochronological, chrono- and chemostratigraphic control of the Neoproterozoic and Cambrian from the Anti-Atlas Ranges and the Ouzellagh promontory (High Atlas). As a result, the Neoproterozoic is lithostratigraphically subdivided into: (i) the Lkest-Taghdout Group (broadly interpreted at c. 800–690 Ma) representative of rift-to-passive margin conditions on the northern West African craton; (ii) the Iriri (c. 760–740 Ma), Bou Azzer (c. 762–697 Ma) and Saghro (c. 760?–610 Ma) groups, the overlying Anezi, Bou Salda, Dadès and Tiddiline formations localized in fault-grabens, and the Ouarzazate Supergroup (c. 615–548 Ma), which form a succession of volcanosedimentary complexes recording the onset of the Pan-African orogeny and its aftermath; and (iii) the Taroudant (the Ediacaran–Cambrian boundary lying in the Tifnout Member of the Adoudou Formation), Tata, Feijas Internes and Tabanite groups that have recorded development of the late Ediacaran–Cambrian Atlas Rift. Recent discussions of Moroccan strata to select new global GSSPs by the International Subcommissions on Ediacaran and Cambrian Stratigraphy have raised the stratigraphic interest in this region. A revised and updated stratigraphic framework is proposed here to assist the tasks of both subcommissions and to fuel future discussions focused on different geological aspects of the Neoproterozoic–Cambrian time span.

  18. Antarctica, supercontinents and the palaeogeography of the Cambrian 'explosion'

    NASA Astrophysics Data System (ADS)

    Dalziel, Ian

    2014-05-01

    Laurentia is bordered by latest Precambrian-Cambrian rifted margins and must therefore have been located within a Precambrian supercontinent. Geochronologic and geochemical evidence indicates that it was attached to parts of the East Antarctic craton within the Rodinian supercontinent in the late Mesoproterozoic. The Mawson craton of Antarctica rifted from the proto-Pacific margin of Laurentia during the Neooproterozoic, colliding with the present 'southern cone' of Laurentia at ~600 Ma along the Shackleton Range suture zone as Gondwana and Laurentia amalgamated to form the ephemeral Pannotia supercontinental assembly at the end of the Precambrian. The abrupt appearance of almost all animal phyla in the fossil record is often colloquially referred to as the Cambrian 'explosion' of life on Earth. It is also named 'Darwin's dilemma,' as he appreciated that this seemingly mysterious event posed a major problem for his theory of evolution by natural selection. It coincided with a time of major marine transgression over all the continents. Although the metazoan 'explosion' is now seen as more protracted than formerly recognized, it is still regarded one of the most critical events in the history of the biosphere. One of the most striking aspects of the earliest Cambrian fossils is geographic differentiation. In particular, the first benthic trilobite faunas on Laurentia, ancestral North America, and the newly amalgamated southern supercontinent of Gondwana are distinctly different. This has led to the suggestion of an unknown vicariant event intervening between an ancestral trilobite clade and higher members that are represented in the fossil record, possibly one related to the breakup of a supercontinent. Igneous rocks along the Panthalassic margin of Gondwana, including South America, southernmost Africa and the Ellsworth-Whitmore crustal block of Antarctica, and along the proto-Appalachian margin of Laurentia indicate that final separation of Laurentia from

  19. Disc-shaped fossils resembling porpitids or eldonids from the early Cambrian (Series 2: Stage 4) of western USA

    PubMed Central

    Kurkewicz, Richard; Shinogle, Heather; Kimmig, Julien; MacGabhann, Breandán Anraoi

    2017-01-01

    The morphology and affinities of newly discovered disc-shaped, soft-bodied fossils from the early Cambrian (Series 2: Stage 4, Dyeran) Carrara Formation are discussed. These specimens show some similarity to the Ordovician Discophyllum Hall, 1847; traditionally this taxon had been treated as a fossil porpitid. However, recently it has instead been referred to as another clade, the eldonids, which includes the enigmatic Eldonia Walcott, 1911 that was originally described from the Cambrian Burgess Shale. The status of various Proterozoic and Phanerozoic taxa previously referred to porpitids and eldonids is also briefly considered. To help ascertain that the specimens were not dubio- or pseudofossils, elemental mapping using energy dispersive X-ray spectroscopy (EDS) was conducted. This, in conjunction with the morphology of the specimens, indicated that the fossils were not hematite, iron sulfide, pyrolusite, or other abiologic mineral precipitates. Instead, their status as biologic structures and thus actual fossils is supported. Enrichment in the element carbon, and also possibly to some extent the elements magnesium and iron, seems to be playing some role in the preservation process. PMID:28603667

  20. Disc-shaped fossils resembling porpitids or eldonids from the early Cambrian (Series 2: Stage 4) of western USA.

    PubMed

    Lieberman, Bruce S; Kurkewicz, Richard; Shinogle, Heather; Kimmig, Julien; MacGabhann, Breandán Anraoi

    2017-01-01

    The morphology and affinities of newly discovered disc-shaped, soft-bodied fossils from the early Cambrian (Series 2: Stage 4, Dyeran) Carrara Formation are discussed. These specimens show some similarity to the Ordovician Discophyllum Hall, 1847; traditionally this taxon had been treated as a fossil porpitid. However, recently it has instead been referred to as another clade, the eldonids, which includes the enigmatic Eldonia Walcott, 1911 that was originally described from the Cambrian Burgess Shale. The status of various Proterozoic and Phanerozoic taxa previously referred to porpitids and eldonids is also briefly considered. To help ascertain that the specimens were not dubio- or pseudofossils, elemental mapping using energy dispersive X-ray spectroscopy (EDS) was conducted. This, in conjunction with the morphology of the specimens, indicated that the fossils were not hematite, iron sulfide, pyrolusite, or other abiologic mineral precipitates. Instead, their status as biologic structures and thus actual fossils is supported. Enrichment in the element carbon, and also possibly to some extent the elements magnesium and iron, seems to be playing some role in the preservation process.

  1. Reconstruction of the Paleoenvironment of the Early Cambrian Yurtus Black Shale in the Tarim Basin, Northwestern China, and Its Control on Organic Matter Accumulation

    NASA Astrophysics Data System (ADS)

    Li, J.; Ding, W.; Dong, L.

    2017-12-01

    The black shale in the early Cambrian Yurtus Formation (>521 Ma) in the Tarim basin, northwestern China, is characterized by its high TOC value (up to 16%) andgreat lateral continuity. It has been proven to be high-quality hydrocarbon source rocks. Abundant phytoplanktons and small shelly fossils have been reported from the lower Yurtus chert. However, recent biomarker discovery of aryl isoprenoid hydrocarbons suggests the existence of green sulfur bacteria, which indicates that the water column was stratified and the photic zone was prevailingly euxinic. These seemingly contradictory observations hamper our further understanding of the paleoenvironment in which the Yurtus shale was deposited and its control on the accumulation of organic matter. In this study, we systematically collected samples from the Yurtus Formation at the Kungaikuotan Section, and measured the organic carbon and nitrogen isotopic compositions and the content of trace element Barium (Ba). The strong negative excursions of nitrogen isotope ( -13‰) in the lower and upper parts of the Yurtus Formation are likely attributed to the biological activity of green and purple sulfur bacteria, which is consistent with our organic carbon isotope data as well as previous biomarker discovery. As green sulfur bacteria can only live in euxinic photic zone, it may indicate that the water column above this euxinic zone contains prolific organic matters which consume all the dissolved oxidants in surface ocean. It is well accepted that Ba flux can be used as an indicator for surface ocean primary productivity. Significant increase of barium content (from <100 to 2000 ppm) is observed at the same horizon as where the negative excursion of δ15Norg occurs, suggesting the substantive organic matter in the early Cambrian surface ocean mainly result from extremely high primary productivity. The abundant phytoplankton fossil record from this time period also supports this interpretation. In summary, high TOC in the

  2. Extraordinary fossils reveal the nature of Cambrian life: a commentary on Whittington (1975) 'The enigmatic animal Opabinia regalis, Middle Cambrian, Burgess Shale, British Columbia'.

    PubMed

    Briggs, Derek E G

    2015-04-19

    Harry Whittington's 1975 monograph on Opabinia was the first to highlight how some of the Burgess Shale animals differ markedly from those that populate today's oceans. Categorized by Stephen J. Gould as a 'weird wonder' (Wonderful life, 1989) Opabinia, together with other unusual Burgess Shale fossils, stimulated ongoing debates about the early evolution of the major animal groups and the nature of the Cambrian explosion. The subsequent discovery of a number of other exceptionally preserved fossil faunas of Cambrian and early Ordovician age has significantly augmented the information available on this critical interval in the history of life. Although Opabinia initially defied assignment to any group of modern animals, it is now interpreted as lying below anomalocaridids on the stem leading to the living arthropods. This commentary was written to celebrate the 350th anniversary of the journal Philosophical Transactions of the Royal Society.

  3. Articulated Wiwaxia from the Cambrian Stage 3 Xiaoshiba lagerstätte.

    PubMed

    Yang, Jie; Smith, Martin R; Lan, Tian; Hou, Jin-bo; Zhang, Xi-guang

    2014-04-10

    Wiwaxia is a bizarre metazoan that has been interpreted as a primitive mollusc and as a polychaete annelid worm. Extensive material from the Burgess Shale provides a detailed picture of its morphology and ontogeny, but the fossil record outside this lagerstätte is scarce, and complete wiwaxiids are particularly rare. Here we report small articulated specimens of Wiwaxia foliosa sp. nov. from the Xiaoshiba fauna (Cambrian Stage 3, Hongjingshao Formation, Kunming, south China). Although spines are absent, the fossils' sclerites - like those of W. corrugata - are symmetrically arranged in five distinct zones. They form rows across the body, and were individually added and shed throughout growth to retain an approximately symmetrical body shape. Their development pattern suggests a molluscan affinity. The basic body plan of wiwaxiids is fundamentally conserved across two continents through Cambrian Stages 3-5 - revealing morphological stasis in the wake of the Cambrian explosion.

  4. New uppermost Cambrian U-Pb date from Avalonian Wales and age of the Cambrian-Ordovician boundary

    USGS Publications Warehouse

    Davidek, K.; Landing, E.; Bowring, S.A.; Westrop, S.R.; Rushton, A.W.A.; Fortey, R.A.; Adrain, J.M.

    1998-01-01

    A crystal-rich volcaniclastic sandatone in the lower Peltura scarabaeoides Zone at Ogof-odi near Criccieth, North Wales, yields a U-Pb zircon age of 491 ?? 1 Ma. This late Late Cambrian date indicates a remarkably young age for the Cambrian-Ordovician boundary whose age must be less than 491 Ma. Hence the revised duration of the post-Placentian (trilobite-bearing) Cambrian indicates that local trilobite zonations allow a biostratigraphic resolution comparble to that provided by Ordovician graptolites and Mesozoic ammonites.

  5. Extraordinary fossils reveal the nature of Cambrian life: a commentary on Whittington (1975) ‘The enigmatic animal Opabinia regalis, Middle Cambrian, Burgess Shale, British Columbia’

    PubMed Central

    Briggs, Derek E. G.

    2015-01-01

    Harry Whittington's 1975 monograph on Opabinia was the first to highlight how some of the Burgess Shale animals differ markedly from those that populate today's oceans. Categorized by Stephen J. Gould as a ‘weird wonder’ (Wonderful life, 1989) Opabinia, together with other unusual Burgess Shale fossils, stimulated ongoing debates about the early evolution of the major animal groups and the nature of the Cambrian explosion. The subsequent discovery of a number of other exceptionally preserved fossil faunas of Cambrian and early Ordovician age has significantly augmented the information available on this critical interval in the history of life. Although Opabinia initially defied assignment to any group of modern animals, it is now interpreted as lying below anomalocaridids on the stem leading to the living arthropods. This commentary was written to celebrate the 350th anniversary of the journal Philosophical Transactions of the Royal Society. PMID:25750235

  6. Fossilized embryos are widespread but the record is temporally and taxonomically biased

    USGS Publications Warehouse

    Donoghue, P.C.J.; Kouchinsky, A.; Waloszek, Dieter; Bengtson, S.; Dong, X.-P.; Val'Kov, A.K.; Cunningham, J.A.; Repetski, J.E.

    2006-01-01

    We report new discoveries of embryos and egg capsules from the Lower Cambrian of Siberia, Middle Cambrian of Australia and Lower Ordovician of North America. Together with existing records, embryos have now been recorded from four of the seven continents. However, the new discoveries highlight secular and systematic biases in the fossil record of embryonic stages. The temporal window within which the embryos and egg capsules are found is of relatively short duration; it ends in the Early Ordovician and is roughly coincident with that of typical "Orsten"-type faunas. The reduced occurrence of such fossils has been attributed to reducing levels of phosphate in marine waters during the early Paleozoic, but may also be owing to the increasing depth of sediment mixing by infaunal metazoans. Furthermore, most records younger than the earliest Cambrian are of a single kind - large eggs and embryos of the priapulid-like scalidophoran Markuelia. We explore alternative explanations for the low taxonomic diversity of embryos recovered thus far, including sampling, size, anatomy, ecology, and environment, concluding that the preponderance of Markuelia embryos is due to its precocious development of cuticle at an embryonic stage, predisposing it to preservation through action as a substrate on which microbially mediated precipitation of authigenic calcium phosphate may occur. The fossil record of embryos may be limited to a late Neoproterozoic to early Ordovician snapshot that is subject to dramatic systematic bias. Together, these biases must be considered seriously in attempts to use the fossil record to arbitrate between hypotheses of developmental and life history evolution implicated in the origin of metazoan clades. ?? 2006 Blackwell Publishing Ltd.

  7. Life cycle and morphology of a cambrian stem-lineage loriciferan.

    PubMed

    Peel, John S; Stein, Martin; Kristensen, Reinhardt Møbjerg

    2013-01-01

    Cycloneuralians form a rich and diverse element within Cambrian assemblages of exceptionally preserved fossils. Most resemble priapulid worms whereas other Cycloneuralia (Nematoda, Nematomorpha, Kinorhyncha, Loricifera), well known at the present day, have little or no fossil record. First reports of Sirilorica Peel, 2010 from the lower Cambrian Sirius Passet fauna of North Greenland described a tubular lorica covering the abdomen and part of a well developed introvert with a circlet of 6 grasping denticles near the lorica. The introvert is now known to terminate in a narrow mouth tube, while a conical anal field is also developed. Broad muscular bands between the plates in the lorica indicate that it was capable of movement by rhythmic expansion and contraction of the lorica. Sirilorica is regarded as a macrobenthic member of the stem-lineage of the miniaturised, interstitial, present day Loricifera. Like loriciferans, Sirilorica is now known to have grown by moulting. Evidence of the life cycle of Sirilorica is described, including a large post-larval stage and probably an initial larva similar to that of the middle Cambrian fossil Orstenoloricusshergoldii.

  8. Life Cycle and Morphology of a Cambrian Stem-Lineage Loriciferan

    PubMed Central

    Peel, John S.; Stein, Martin; Kristensen, Reinhardt Møbjerg

    2013-01-01

    Cycloneuralians form a rich and diverse element within Cambrian assemblages of exceptionally preserved fossils. Most resemble priapulid worms whereas other Cycloneuralia (Nematoda, Nematomorpha, Kinorhyncha, Loricifera), well known at the present day, have little or no fossil record. First reports of Sirilorica Peel, 2010 from the lower Cambrian Sirius Passet fauna of North Greenland described a tubular lorica covering the abdomen and part of a well developed introvert with a circlet of 6 grasping denticles near the lorica. The introvert is now known to terminate in a narrow mouth tube, while a conical anal field is also developed. Broad muscular bands between the plates in the lorica indicate that it was capable of movement by rhythmic expansion and contraction of the lorica. Sirilorica is regarded as a macrobenthic member of the stem-lineage of the miniaturised, interstitial, present day Loricifera. Like loriciferans, Sirilorica is now known to have grown by moulting. Evidence of the life cycle of Sirilorica is described, including a large post-larval stage and probably an initial larva similar to that of the middle Cambrian fossil Orstenoloricusshergoldii . PMID:23991198

  9. Latest Cambrian-Early Ordovician rift-related magmatic activity in the Kouřim Unit, Bohemian Massif

    NASA Astrophysics Data System (ADS)

    Soejono, Igor; Machek, Matej; Sláma, Jiří; Janoušek, Vojtěch

    2017-04-01

    inherited from the source, represented most likely by recycled immature arc-related material (?metagraywackes). The real tectonic setting of this Late Cambrian magmatic activity seems rather indicated by the within-plate geochemistry of the metadiorite. These results bring further evidence for the presence of the Late Cambrian-Early Ordovician extensional event documented throughout the basement of the European Variscan Belt. Together with other occurrences of bimodal magmatism, as well as metamorphic and sedimentary record, indicate an important period of lithospheric thinning. This overall Early Palaeozoic rift-related architecture is often considered as a consequence of the Rheic Ocean opening.

  10. Decoupling of body-plan diversification and ecological structuring during the Ediacaran-Cambrian transition: evolutionary and geobiological feedbacks.

    PubMed

    Mángano, M Gabriela; Buatois, Luis A

    2014-04-07

    The rapid appearance of bilaterian clades at the beginning of the Phanerozoic is one of the most intriguing topics in macroevolution. However, the complex feedbacks between diversification and ecological interactions are still poorly understood. Here, we show that a systematic and comprehensive analysis of the trace-fossil record of the Ediacaran-Cambrian transition indicates that body-plan diversification and ecological structuring were decoupled. The appearance of a wide repertoire of behavioural strategies and body plans occurred by the Fortunian. However, a major shift in benthic ecological structure, recording the establishment of a suspension-feeder infauna, increased complexity of the trophic web, and coupling of benthos and plankton took place during Cambrian Stage 2. Both phases were accompanied by different styles of ecosystem engineering, but only the second one resulted in the establishment of the Phanerozoic-style ecology. In turn, the suspension-feeding infauna may have been the ecological drivers of a further diversification of deposit-feeding strategies by Cambrian Stage 3, favouring an ecological spillover scenario. Trace-fossil information strongly supports the Cambrian explosion, but allows for a short time of phylogenetic fuse during the terminal Ediacaran-Fortunian.

  11. Decoupling of body-plan diversification and ecological structuring during the Ediacaran–Cambrian transition: evolutionary and geobiological feedbacks

    PubMed Central

    Mángano, M. Gabriela; Buatois, Luis A.

    2014-01-01

    The rapid appearance of bilaterian clades at the beginning of the Phanerozoic is one of the most intriguing topics in macroevolution. However, the complex feedbacks between diversification and ecological interactions are still poorly understood. Here, we show that a systematic and comprehensive analysis of the trace-fossil record of the Ediacaran–Cambrian transition indicates that body-plan diversification and ecological structuring were decoupled. The appearance of a wide repertoire of behavioural strategies and body plans occurred by the Fortunian. However, a major shift in benthic ecological structure, recording the establishment of a suspension-feeder infauna, increased complexity of the trophic web, and coupling of benthos and plankton took place during Cambrian Stage 2. Both phases were accompanied by different styles of ecosystem engineering, but only the second one resulted in the establishment of the Phanerozoic-style ecology. In turn, the suspension-feeding infauna may have been the ecological drivers of a further diversification of deposit-feeding strategies by Cambrian Stage 3, favouring an ecological spillover scenario. Trace-fossil information strongly supports the Cambrian explosion, but allows for a short time of phylogenetic fuse during the terminal Ediacaran–Fortunian. PMID:24523279

  12. Upper Cambrian chitons (Mollusca, polyplacophora) from Missouri, USA

    USGS Publications Warehouse

    Pojeta, J.; Vendrasco, M.J.; Darrough, G.

    2010-01-01

    Numerous new specimens reveal a greater presence of chitons in Upper Cambrian rocks than previously suspected. Evidence is presented showing that the chiton esthete sensory system is present in all chiton species in this study at the very beginning of the known polyplacophoran fossil record. The stratigraphic occurrences and paleobiogeography of Late Cambrian chitons are documented. The 14 previously-named families of Cambrian and Ordovician chitons are reviewed and analyzed. Aulochitonidae n. fam. is defined, based on Aulochiton n. gen.; A. sannerae n. sp. is also defined. The long misunderstood family Preacanthochitonidae and its type genus Preacanthochiton Bergenhayn, 1960, are placed in synonymy with Mattheviidae and Chelodes Davidson & King, 1874, respectively; Eochelodes Marek, 1962, also is placed in synonymy with Chelodes, and Elongata Stinchcomb & Darrough, 1995, is placed in synonymy with Hemithecella Ulrich & Bridge, 1941. At the species level, H. elongata Stinchcomb & Darrough, 1995, and Elongata perplexa Stinchcomb & Darrough, 1995, are placed in synonymy with H. eminensis Stinchcomb & Darrough, 1995. The Ordovician species H. abrupta Stinchcomb & Darrough, 1995, is transferred to the genus Chelodes as C. abrupta (Stinchcomb & Darrough, 1995). The Ordovician species Preacanthochiton baueri Hoare & Pojeta, 2006, is transferred to the genus Helminthochiton as H. ? baueri (Hoare & Pojeta, 2006). The Ordovician species H. marginatus Hoare & Pojeta, 2006, is transferred to the genus Litochiton as L. marginatus (Hoare & Pojeta, 2006). Matthevia walcotti Runnegar, Pojeta, Taylor, & Collins, 1979, is treated as a synonym of Hemithecella expansa Ulrich & Bridge, 1941. In addition, other multivalved Cambrian mollusks are discussed; within this group, Dycheiidae n. fam. is defined, as well as Paradycheia dorisae n. gen. and n. sp. Cladistic analysis indicates a close relationship among the genera here assigned to the Mattheviidae, and between Echinochiton Pojeta

  13. Evolution of the seawater sulfate sulfur composition through the Cambrian Period: Implications from carbonate-associated sulfate

    NASA Astrophysics Data System (ADS)

    Wotte, T.

    2012-12-01

    Cambrian carbonate successions of Australia, W-Gondwana, Kazakhstan, Laurentia, and Siberia were investigated for their sulfur isotopic composition of carbonate-associated sulfate (CAS). For a secure CAS extraction a repeated leaching procedure with NaCl solution was applied as a standard protocol with supplementary analyses of pre-leach sulfate concentrations and δ34SNaCl, and chromium-reducible sulfur (CRS) concentrations and δ34SCRS as routine checks on possible contamination. Additionally, δ13Ccarb, δ18Ocarb, and elemental concentrations (Ca, Fe, Mg, Mn, Sr) of the carbonate host rock were analyzed in order to constrain diagenetic alteration of the measured δ34SCAS. About 200 δ34SCAS values were generated using this analytical procedure which allows the most precise state of the art methodology for CAS and CRS extraction. The most primary δ34SCAS values vary between 24‰ and 33‰ for successions of the transition from Cambrian Series 2 to Cambrian Series 3 (traditional Lower-Middle Cambrian boundary) and between 25‰ and 48‰ for the Cambrian Series 3-Furongian (Upper Cambrian) interval, respectively. These new δ34SCAS data are valuable proxies to verify paleoecological and paleoenvironmental information. They further close an obvious gap of the Cambrian δ34SCAS record, composed of δ34SCAS values, ranging from about 20‰ up to 70‰ (SPICE-event). However, the individual data sets of this Cambrian δ34SCAS pool were generated using various methods of CAS extraction, thus offering the potential of significant differences in the final δ34SCAS values which are consequently not per se comparable with each other. The new δ34SCAS data of worldwide Cambrian sections will be discussed in the context of this problematic.

  14. Oxygen, ecology, and the Cambrian radiation of animals

    PubMed Central

    Sperling, Erik A.; Frieder, Christina A.; Raman, Akkur V.; Girguis, Peter R.; Levin, Lisa A.; Knoll, Andrew H.

    2013-01-01

    The Proterozoic-Cambrian transition records the appearance of essentially all animal body plans (phyla), yet to date no single hypothesis adequately explains both the timing of the event and the evident increase in diversity and disparity. Ecological triggers focused on escalatory predator–prey “arms races” can explain the evolutionary pattern but not its timing, whereas environmental triggers, particularly ocean/atmosphere oxygenation, do the reverse. Using modern oxygen minimum zones as an analog for Proterozoic oceans, we explore the effect of low oxygen levels on the feeding ecology of polychaetes, the dominant macrofaunal animals in deep-sea sediments. Here we show that low oxygen is clearly linked to low proportions of carnivores in a community and low diversity of carnivorous taxa, whereas higher oxygen levels support more complex food webs. The recognition of a physiological control on carnivory therefore links environmental triggers and ecological drivers, providing an integrated explanation for both the pattern and timing of Cambrian animal radiation. PMID:23898193

  15. Oxygen, ecology, and the Cambrian radiation of animals

    NASA Astrophysics Data System (ADS)

    Sperling, Erik A.; Frieder, Christina A.; Raman, Akkur V.; Girguis, Peter R.; Levin, Lisa A.; Knoll, Andrew H.

    2013-08-01

    The Proterozoic-Cambrian transition records the appearance of essentially all animal body plans (phyla), yet to date no single hypothesis adequately explains both the timing of the event and the evident increase in diversity and disparity. Ecological triggers focused on escalatory predator-prey "arms races" can explain the evolutionary pattern but not its timing, whereas environmental triggers, particularly ocean/atmosphere oxygenation, do the reverse. Using modern oxygen minimum zones as an analog for Proterozoic oceans, we explore the effect of low oxygen levels on the feeding ecology of polychaetes, the dominant macrofaunal animals in deep-sea sediments. Here we show that low oxygen is clearly linked to low proportions of carnivores in a community and low diversity of carnivorous taxa, whereas higher oxygen levels support more complex food webs. The recognition of a physiological control on carnivory therefore links environmental triggers and ecological drivers, providing an integrated explanation for both the pattern and timing of Cambrian animal radiation.

  16. Tropical shoreline ice in the late Cambrian: Implications for earth's climate between the Cambrian Explosion and the Great Ordovician Biodiversification Event

    USGS Publications Warehouse

    Runkel, Anthony C.; MacKey, T.J.; Cowan, Clinton A.; Fox, David L.

    2010-01-01

    Middle to late Cambrian time (ca. 513 to 488 Ma) is characterized by an unstable plateau in biodiversity, when depauperate shelf faunas suffered repeated extinctions. This poorly understood interval separates the Cambrian Explosion from the Great Ordovician Biodiversification Event and is generally regarded as a time of sustained greenhouse conditions. We present evidence that suggests a drastically different climate during this enigmatic interval: Features indicative of meteoric ice are well preserved in late Cambrian equatorial beach deposits that correspond to one of the shelf extinction events. Thus, the middle to late Cambrian Earth was at least episodically cold and might best be considered a muted analogue to the environmental extremes that characterized the Proterozoic, even though cooling in the two periods may have occurred in response to different triggers. Such later Cambrian conditions may have significantly impacted evolution preceding the Ordovician radiation.

  17. Multiple Controls on the Paleoenvironment of the Early Cambrian Marine Black Shales in the Sichuan Basin, SW China: Geochemical and Organic Carbon Isotopic Evidence

    NASA Astrophysics Data System (ADS)

    Wang, S.; Zhang, G.; Dong, D.; Wang, Y.

    2016-12-01

    In order to understand the paleoenvironment of the Early Cambrian black shale deposition in the western part of the Yangtze Block, geochemical and organic carbon isotopic studies have been performed on two wells that have drilled through the Qiongzhusi Formation in the central and southeastern parts of Sichuan Basin. It shows that the lowest part of the Qiongzhusi Formation has high TOC abundance, while the middle and upper parts display relative low TOC content. Redox-sensitive element (Mo) and trace elemental redox indices (e.g., Ni/Co, V/Cr, U/Th and V/(V+Ni)) suggest that the high-TOC layers were deposited under anoxic conditions, whereas the low-TOC layers under relatively dysoxic/oxic conditions. The relationship of the enrichment factors of Mo and U further shows a transition from suboxic low-TOC layers to euxinic high-TOC layers. On the basis of the Mo-TOC relationship, the Qiongzhusi Formation black shales were deposited in a basin under moderately restricted conditions. Organic carbon isotopes display temporal variations in the Qiongzhusi Formation, with a positive excursion of δ13Corg values in the lower part and a continuous positive shift in the middle and upper parts. All these geochemical and isotopic criteria indicate a paleoenvironmental change from bottom anoxic to middle and upper dysoxic/oxic conditions for the Qiongzhusi Formation black shales. The correlation of organic carbon isotopic data for the Lower Cambrian black shales in different regions of the Yangtze Block shows consistent positive excursion of δ13Corg values in the lower part for each section. This excursion can be ascribed to the widespread Early Cambrian transgression in the Yangtze Block, under which black shales were deposited.

  18. Gas potential of the Rome Trough in Kentucky: Results of recent Cambrian exploration

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Harris, D.C.; Drahovzal, J.A.

    1996-09-01

    A recent gas discovery in the Rome Trough suggests the need to re-evaluate the deep Cambrian potential of eastern Kentucky. A new phase of Cambrian exploration began in mid-1994 with a new pool discovery by the Carson Associates No. 1 Kazee well in Elliott County, Ky. This well blew out and initially flowed 11 MMcfd of gas from the upper Conasauga Group/Rome Formation at 6,258 to 6,270 feet. After this discovery, a second exploratory well (the Blue Ridge No. 1Greene) was drilled on a separate structure in Elliott County in late 1995. The Blue Ridge well was temporarily abandoned, butmore » had shows of gas and condensate. In early 1996, Carson Associates offset their initial discovery well with the No. 33 Lawson Heirs well. This activity follows a frustrating exploration history in the Rome Trough that is marked by numerous gas and oil shows, but rare commercial production. Only three single-well pools have produced commercial gas from the trough, including the recent Kazee well. Stratigraphic units below the Cambrian-Ordovician Knox Group in the Rome Trough are dramatically thicker than their equivalents on the shelf to the north. The interval in the trough is thought to include rocks as old as Early Cambrian, consisting of a basal sandstone, equivalents of the Shady/Tomstown Dolomite, the Rome Formation, and the Conasauga Formation. Sandstones and fractured shales have been responsible for most of the production to date, but dolostone intervals may also have potential. Limited seismic data indicate possible fan-delta and basin-floor fan deposits that may have reservoir potential.« less

  19. Minerals in the gut: scoping a Cambrian digestive system

    NASA Astrophysics Data System (ADS)

    Strang, K. M.; Armstrong, H. A.; Harper, D. A. T.

    2016-11-01

    The Sirius Passet Lagerstätte of North Greenland contains the first exceptionally preserved mat-ground community of the Cambrian, dominated, in terms of abundance, by trilobites but particularly characterized by iconic arthropods and lobopods, some also occurring in the Burgess shale. High-resolution photography, scanning electron imaging and elemental mapping have been carried out on a variety of specimens of the non-mineralized arthropod Campanamuta mantonae (Budd 2011 J. Syst. Palaeontol. 9, 217-260 (doi:10.1080/14772019.2010.492644)) which has three-dimensional gut and muscle preservation. Results show that the guts contain a high concentration of calcium phosphate (approximating to the mineral francolite), whereas the adjacent muscles are silicified. This indicates a unique, tissue-specific taphonomy for this Cambrian taxon. We hypothesize that the precipitation of calcium phosphate in the guts occurs rapidly after death by `crystal seed' processes in suboxic, slightly acidic conditions; critically, the gut wall remained intact during precipitation. We postulate that the calcium phosphate was derived from ingested cellular material. Silicification of the muscles followed as the localized water chemistry became saturated in silica, high in Fe2+, and low in oxygen and sulfate. We document here the unique occurrence of two distinct but mechanistically similar taphonomic pathways within a diverse suite of possibilities in an Early Cambrian Lagerstätte.

  20. Meiofaunal deuterostomes from the basal Cambrian of Shaanxi (China).

    PubMed

    Han, Jian; Morris, Simon Conway; Ou, Qiang; Shu, Degan; Huang, Hai

    2017-02-09

    Deuterostomes include the group we belong to (vertebrates) as well as an array of disparate forms that include echinoderms, hemichordates and more problematic groups such as vetulicolians and vetulocystids. The Cambrian fossil record is well-populated with representative examples, but possible intermediates are controversial and the nature of the original deuterostome remains idealized. Here we report millimetric fossils, Saccorhytus coronarius nov. gen., nov. sp., from an Orsten-like Lagerstätte from the earliest Cambrian period of South China, which stratigraphically are amongst the earliest of deuterostomes. The bag-like body bears a prominent mouth and associated folds, and behind them up to four conical openings on either side of the body as well as possible sensory structures. An anus may have been absent, and correspondingly the lateral openings probably served to expel water and waste material. This new form has similarities to both the vetulicolians and vetulocystids and collectively these findings suggest that a key step in deuterostome evolution was the development of lateral openings that subsequently were co-opted as pharyngeal gills. Depending on its exact phylogenetic position, the meiofaunal habit of Saccorhytus may help to explain the major gap between divergence times seen in the fossil record and estimates based on molecular clocks.

  1. Rise to modern levels of ocean oxygenation coincided with the Cambrian radiation of animals.

    PubMed

    Chen, Xi; Ling, Hong-Fei; Vance, Derek; Shields-Zhou, Graham A; Zhu, Maoyan; Poulton, Simon W; Och, Lawrence M; Jiang, Shao-Yong; Li, Da; Cremonese, Lorenzo; Archer, Corey

    2015-05-18

    The early diversification of animals (∼ 630 Ma), and their development into both motile and macroscopic forms (∼ 575-565 Ma), has been linked to stepwise increases in the oxygenation of Earth's surface environment. However, establishing such a linkage between oxygen and evolution for the later Cambrian 'explosion' (540-520 Ma) of new, energy-sapping body plans and behaviours has proved more elusive. Here we present new molybdenum isotope data, which demonstrate that the areal extent of oxygenated bottom waters increased in step with the early Cambrian bioradiation of animals and eukaryotic phytoplankton. Modern-like oxygen levels characterized the ocean at ∼ 521 Ma for the first time in Earth history. This marks the first establishment of a key environmental factor in modern-like ecosystems, where animals benefit from, and also contribute to, the 'homeostasis' of marine redox conditions.

  2. Cambrian ophiolite complexes in the Beishan area, China, southern margin of the Central Asian Orogenic Belt

    NASA Astrophysics Data System (ADS)

    Shi, Yuruo; Zhang, Wei; Kröner, Alfred; Li, Linlin; Jian, Ping

    2018-03-01

    We present zircon ages and geochemical data for Cambrian ophiolite complexes exposed in the Beishan area at the southern margin of the Central Asian Orogenic Belt (CAOB). The complexes consist of the Xichangjing-Xiaohuangshan and Hongliuhe-Yushishan ophiolites, which both exhibit complete ophiolite stratigraphy: chert, basalt, sheeted dikes, gabbro, mafic and ultramafic cumulates and serpentinized mantle peridotites. Zircon grains of gabbro samples yielded 206Pb/238U ages of 516 ± 8, 521 ± 4, 528 ± 3 and 535 ± 6 Ma that reflect the timing of gabbro emplacement. The geochemical data of the basaltic rocks show enrichment in large-ion lithophile elements and depletion in the high field strength elements relative to normal mid-oceanic ridge basalt (NMORB) in response to aqueous fluids or melts expelled from the subducting slab. The gabbro samples have higher whole-rock initial 87Sr/86Sr ratios and lower positive εNd(t) values than NMORB. These geochemical signatures resulted from processes or conditions that are unique to subduction zones, and the ophiolites are therefore likely to have formed within a supra-subduction zone (SSZ) environment. We suggest that the Cambrian ophiolite complexes in the Beishan area formed within a SSZ setting, reflecting an early Paleozoic subduction of components of the Paleo-Central Asian Ocean and recording an early Paleozoic southward subduction event in the southern CAOB along the northern margin of the Tarim and North China Cratons.

  3. Giving the early fossil record of sponges a squeeze.

    PubMed

    Antcliffe, Jonathan B; Callow, Richard H T; Brasier, Martin D

    2014-11-01

    Twenty candidate fossils with claim to be the oldest representative of the Phylum Porifera have been re-analysed. Three criteria are used to assess each candidate: (i) the diagnostic criteria needed to categorize sponges in the fossil record; (ii) the presence, or absence, of such diagnostic features in the putative poriferan fossils; and (iii) the age constraints for the candidate fossils. All three criteria are critical to the correct interpretation of any fossil and its placement within an evolutionary context. Our analysis shows that no Precambrian fossil candidate yet satisfies all three of these criteria to be a reliable sponge fossil. The oldest widely accepted candidate, Mongolian silica hexacts from c. 545 million years ago (Ma), are here shown to be cruciform arsenopyrite crystals. The oldest reliable sponge remains are siliceous spicules from the basal Cambrian (Protohertzina anabarica Zone) Soltanieh Formation, Iran, which are described and analysed here in detail for the first time. Extensive archaeocyathan sponge reefs emerge and radiate as late as the middle of the Fortunian Stage of the Cambrian and demonstrate a gradual assembly of their skeletal structure through this time coincident with the evolution of other metazoan groups. Since the Porifera are basal in the Metazoa, their presence within the late Proterozoic has been widely anticipated. Molecular clock calibration for the earliest Porifera and Metazoa should now be based on the Iranian hexactinellid material dated to c. 535 Ma. The earliest convincing fossil sponge remains appeared at around the time of the Precambrian-Cambrian boundary, associated with the great radiation events of that interval. © 2014 The Authors. Biological Reviews © 2014 Cambridge Philosophical Society.

  4. Cambrian stem-group annelids and a metameric origin of the annelid head.

    PubMed

    Parry, Luke; Vinther, Jakob; Edgecombe, Gregory D

    2015-10-01

    The oldest fossil annelids come from the Early Cambrian Sirius Passet and Guanshan biotas and Middle Cambrian Burgess Shale. While these are among the best preserved polychaete fossils, their relationship to living taxa is contentious, having been interpreted either as members of extant clades or as a grade outside the crown group. New morphological observations from five Cambrian species include the oldest polychaete with head appendages, a new specimen of Pygocirrus from Sirius Passet, and an undescribed form from the Burgess Shale. We propose that the palps of Canadia are on an anterior segment bearing neuropodia and that the head of Phragmochaeta is formed of a segment bearing biramous parapodia and chaetae. The unusual anatomy of these taxa suggests that the head is not differentiated into a prostomium and peristomium, that palps are derived from a modified parapodium and that the annelid head was originally a parapodium-bearing segment. Canadia, Phragmochaeta and the Marble Canyon annelid share the presence of protective notochaetae, interpreted as a primitive character state subsequently lost in Pygocirrus and Burgessochaeta, in which the head is clearly differentiated from the trunk. © 2015 The Authors.

  5. The oldest known digestive system consisting of both paired digestive glands and a crop from exceptionally preserved trilobites of the Guanshan Biota (Early Cambrian, China)

    PubMed Central

    Chen, Feiyang; Hu, Shixue; Zhang, Zhifei

    2017-01-01

    The early Cambrian Guanshan biota of eastern Yunnan, China, contains exceptionally preserved animals and algae. Most diverse and abundant are the arthropods, of which there are at least 11 species of trilobites represented by numerous specimens. Many trilobite specimens show soft-body preservation via iron oxide pseudomorphs of pyrite replacement. Here we describe digestive structures from two species of trilobite, Palaeolenus lantenoisi and Redlichia mansuyi. Multiple specimens of both species contain the preserved remains of an expanded stomach region (a “crop”) under the glabella, a structure which has not been observed in trilobites this old, despite numerous examples of trilobite gut traces from other Cambrian Lagerstätten. In addition, at least one specimen of Palaeolenus lantenoisi shows the preservation of an unusual combination of digestive structures: a crop and paired digestive glands along the alimentary tract. This combination of digestive structures has also never been observed in trilobites this old, and is rare in general, with prior evidence of it from one juvenile trilobite specimen from the late Cambrian Orsten fauna of Sweden and possibly one adult trilobite specimen from the Early Ordovician Fezouata Lagerstätte. The variation in the fidelity of preservation of digestive structures within and across different Lagerstätten may be due to variation in the type, quality, and point of digestion of food among specimens in addition to differences in mode of preservation. The presence and combination of these digestive features in the Guanshan trilobites contradicts current models of how the trilobite digestive system was structured and evolved over time. Most notably, the crop is not a derived structure as previously proposed, although it is possible that the relative size of the crop increased over the evolutionary history of the clade. PMID:28934290

  6. The oldest known digestive system consisting of both paired digestive glands and a crop from exceptionally preserved trilobites of the Guanshan Biota (Early Cambrian, China).

    PubMed

    Hopkins, Melanie J; Chen, Feiyang; Hu, Shixue; Zhang, Zhifei

    2017-01-01

    The early Cambrian Guanshan biota of eastern Yunnan, China, contains exceptionally preserved animals and algae. Most diverse and abundant are the arthropods, of which there are at least 11 species of trilobites represented by numerous specimens. Many trilobite specimens show soft-body preservation via iron oxide pseudomorphs of pyrite replacement. Here we describe digestive structures from two species of trilobite, Palaeolenus lantenoisi and Redlichia mansuyi. Multiple specimens of both species contain the preserved remains of an expanded stomach region (a "crop") under the glabella, a structure which has not been observed in trilobites this old, despite numerous examples of trilobite gut traces from other Cambrian Lagerstätten. In addition, at least one specimen of Palaeolenus lantenoisi shows the preservation of an unusual combination of digestive structures: a crop and paired digestive glands along the alimentary tract. This combination of digestive structures has also never been observed in trilobites this old, and is rare in general, with prior evidence of it from one juvenile trilobite specimen from the late Cambrian Orsten fauna of Sweden and possibly one adult trilobite specimen from the Early Ordovician Fezouata Lagerstätte. The variation in the fidelity of preservation of digestive structures within and across different Lagerstätten may be due to variation in the type, quality, and point of digestion of food among specimens in addition to differences in mode of preservation. The presence and combination of these digestive features in the Guanshan trilobites contradicts current models of how the trilobite digestive system was structured and evolved over time. Most notably, the crop is not a derived structure as previously proposed, although it is possible that the relative size of the crop increased over the evolutionary history of the clade.

  7. Rise to modern levels of ocean oxygenation coincided with the Cambrian radiation of animals

    PubMed Central

    Chen, Xi; Ling, Hong-Fei; Vance, Derek; Shields-Zhou, Graham A.; Zhu, Maoyan; Poulton, Simon W.; Och, Lawrence M.; Jiang, Shao-Yong; Li, Da; Cremonese, Lorenzo; Archer, Corey

    2015-01-01

    The early diversification of animals (∼630 Ma), and their development into both motile and macroscopic forms (∼575–565 Ma), has been linked to stepwise increases in the oxygenation of Earth's surface environment. However, establishing such a linkage between oxygen and evolution for the later Cambrian ‘explosion' (540–520 Ma) of new, energy-sapping body plans and behaviours has proved more elusive. Here we present new molybdenum isotope data, which demonstrate that the areal extent of oxygenated bottom waters increased in step with the early Cambrian bioradiation of animals and eukaryotic phytoplankton. Modern-like oxygen levels characterized the ocean at ∼521 Ma for the first time in Earth history. This marks the first establishment of a key environmental factor in modern-like ecosystems, where animals benefit from, and also contribute to, the ‘homeostasis' of marine redox conditions. PMID:25980960

  8. Review of the Cambrian volcanic activity in Morocco: geochemical fingerprints and geotectonic implications for the rifting of West Gondwana

    NASA Astrophysics Data System (ADS)

    Pouclet, André; El Hadi, Hassan; Álvaro, J. Javier; Bardintzeff, Jacques-Marie; Benharref, Mohammed; Fekkak, Abdelilah

    2018-03-01

    Volcanic activities related to the opening of a Cambrian rift in Morocco were widespread from the Fortunian to the Cambrian Epoch 3. Numerous data are available from northwestern volcanic sites, particularly in the western High Atlas, but they are scarce from the southeastern sites. New data are documented here from the volcanic formations exposed in the Jbel Tazoult n'Ouzina of the Tafilalt Province, eastern Anti-Atlas and dated to Cambrian Epoch 2-3. The Cambrian volcanic activities recorded in the High Atlas, Anti-Atlas, and Coastal Meseta are synthesized to refine their stratigraphic setting and to characterize their magmatic affinities and fingerprints. Six volcanic pulses are determined as tholeiitic, transitional, and alkaline suites. The tholeiitic and transitional magmas originated from primitive mantle and E-MORB-type sources with a spinel- and garnet-bearing lherzolite composition. Some of them were modified by assimilation-fractional crystallisation processes during crust-mantle interactions. The alkaline magmas fit with an OIB-type and a garnet-bearing lherzolite source. The palaeogeographic distribution of the magmatic suites was controlled by the lithospheric thinning of the Cambrian Atlas Rift and lithospheric constraints of the Pan-African metacraton and West African craton.

  9. Upper Lower Cambrian depositional sequence in Avalonian New Brunswick

    USGS Publications Warehouse

    Landing, E.; Westrop, S.R.

    1996-01-01

    The Hanford Brook Formation (emended) is a thin (up to 42+ m), upper Lower Cambrian depositional sequence that is unconformably bounded by the lower Lower Cambrian (Random Formation) and the middle Middle Cambrian (Fossil Brook Member of the Chamberlain's Brook Formation). These stratigraphic relationships of the trilobite-bearing Hanford Brook Formation indicate deposition on the Avalonian marginal platform in the Saint John, New Brunswick, region and provide more evidence for a uniform, latest Precambrian-Cambrian epeirogenic history and cover sequence in Avalon. The Hanford Brook Formation is a deepening - shoaling sequence with (i) lower, transgressive sandstone deposited in episodically high-energy environments (St. Martins Member, new); (ii) highstand-regressive, dysaerobic mudstone - fine-grained sandstone with volcanic ashes (Somerset Street Member, new); and (iii) upper, regressive, planar and hummocky cross-stratified sandstone (Long Island Member, new). Trilobites are common in the distal Somerset Street Member, and ostracodes and brachiopods dominate the St. Martins and Long Island members. Condensation of the St. Martins Member and absence of the Long Island Member where the Random Formation and Fossil Brook Member are thinnest suggest onlap of the Hanford Brook and pronounced, sub-Middle Cambrian erosion across epeirogenically active blocks in southern New Brunswick.

  10. New zircon ages on the Cambrian-Ordovician volcanism of the Southern Gemericum basement (Western Carpathians, Slovakia): SHRIMP dating, geochemistry and provenance

    NASA Astrophysics Data System (ADS)

    Vozárová, Anna; Rodionov, Nickolay; Šarinová, Katarína; Presnyakov, Sergey

    2017-09-01

    The Southern Gemericum basement in the Inner Western Carpathians, composed of low-grade volcano-sedimentary rock complexes, constitutes a record of the polyphase Cambrian-Ordovician continental volcanic arc volcanism. These metavolcanic rocks are characterized by the enrichment in K, Rb, Ba, Th and Ce and Sm relative to Ta, Nb, Hf, Zr, Y and Yb that are the characteristic features for volcanic arc magmatites. The new SHRIMP U-Pb zircon data and compilation of previously published and re-evaluated zircon ages, contribute to a new constrain of the timing of the Cambrian-Ordovician volcanism that occurred between 496 and 447 Ma. The following peaks of the volcanic activity of the Southern Gemericum basement have been recognized: (a) mid-late Furongian at 492 Ma; (b) Tremadocian at 481 Ma; (c) Darriwilian at 464 Ma prolonged to 453 Ma within the early Upper Ordovician. The metavolcanic rocks are characterized by a high zircon inheritance, composed of Ediacaran (650-550 Ma), Tonian-Stenian (1.1-0.9 Ma), and, to a lesser extent, Mesoproterozoic (1.3 Ga), Paleoproterozoic (1.9 Ga) and Archaean assemblages (2.6 Ga). Based on the acquired zircon populations, it could be deduced that Cambrian-Ordovician arc crust was generated by a partial melting of Ediacaran basement in the subduction-related setting, into which old crustal fragments were incorporated. The ascertained zircon inheritances with Meso-, Paleoproterozoic and Archaean cores indicate the similarities with the Saharan Metacraton provenance.

  11. Gut Contents as Direct Indicators for Trophic Relationships in the Cambrian Marine Ecosystem

    PubMed Central

    Vannier, Jean

    2012-01-01

    Present-day ecosystems host a huge variety of organisms that interact and transfer mass and energy via a cascade of trophic levels. When and how this complex machinery was established remains largely unknown. Although exceptionally preserved biotas clearly show that Early Cambrian animals had already acquired functionalities that enabled them to exploit a wide range of food resources, there is scant direct evidence concerning their diet and exact trophic relationships. Here I describe the gut contents of Ottoia prolifica, an abundant priapulid worm from the middle Cambrian (Stage 5) Burgess Shale biota. I identify the undigested exoskeletal remains of a wide range of small invertebrates that lived at or near the water sediment interface such as hyolithids, brachiopods, different types of arthropods, polychaetes and wiwaxiids. This set of direct fossil evidence allows the first detailed reconstruction of the diet of a 505-million-year-old animal. Ottoia was a dietary generalist and had no strict feeding regime. It fed on both living individuals and decaying organic matter present in its habitat. The feeding behavior of Ottoia was remarkably simple, reduced to the transit of food through an eversible pharynx and a tubular gut with limited physical breakdown and no storage. The recognition of generalist feeding strategies, exemplified by Ottoia, reveals key-aspects of modern-style trophic complexity in the immediate aftermath of the Cambrian explosion. It also shows that the middle Cambrian ecosystem was already too complex to be understood in terms of simple linear dynamics and unique pathways. PMID:23300612

  12. The controversial "Cambrian" fossils of the Vindhyan are real but more than a billion years older.

    PubMed

    Bengtson, Stefan; Belivanova, Veneta; Rasmussen, Birger; Whitehouse, Martin

    2009-05-12

    The age of the Vindhyan sedimentary basin in central India is controversial, because geochronology indicating early Proterozoic ages clashes with reports of Cambrian fossils. We present here an integrated paleontologic-geochronologic investigation to resolve this conundrum. New sampling of Lower Vindhyan phosphoritic stromatolitic dolomites from the northern flank of the Vindhyans confirms the presence of fossils most closely resembling those found elsewhere in Cambrian deposits: annulated tubes, embryo-like globules with polygonal surface pattern, and filamentous and coccoidal microbial fabrics similar to Girvanella and Renalcis. None of the fossils, however, can be ascribed to uniquely Cambrian or Ediacaran taxa. Indeed, the embryo-like globules are not interpreted as fossils at all but as former gas bubbles trapped in mucus-rich cyanobacterial mats. Direct dating of the same fossiliferous phosphorite yielded a Pb-Pb isochron of 1,650 +/- 89 (2sigma) million years ago, confirming the Paleoproterozoic age of the fossils. New U-Pb geochronology of zircons from tuffaceous mudrocks in the Lower Vindhyan Porcellanite Formation on the southern flank of the Vindhyans give comparable ages. The Vindhyan phosphorites provide a window of 3-dimensionally preserved Paleoproterozoic fossils resembling filamentous and coccoidal cyanobacteria and filamentous eukaryotic algae, as well as problematic forms. Like Neoproterozoic phosphorites a billion years later, the Vindhyan deposits offer important new insights into the nature and diversity of life, and in particular, the early evolution of multicellular eukaryotes.

  13. Exceptionally preserved crustaceans from western Canada reveal a cryptic Cambrian radiation

    NASA Astrophysics Data System (ADS)

    Harvey, Thomas H. P.; Vélez, Maria I.; Butterfield, Nicholas J.

    2012-01-01

    The early history of crustaceans is obscured by strong biases in fossil preservation, but a previously overlooked taphonomic mode yields important complementary insights. Here we describe diverse crustacean appendages of Middle and Late Cambrian age from shallow-marine mudstones of the Deadwood Formation in western Canada. The fossils occur as flattened and fragmentary carbonaceous cuticles but provide a suite of phylogenetic and ecological data by virtue of their detailed preservation. In addition to an unprecedented range of complex, largely articulated filtering limbs, we identify at least four distinct types of mandible. Together, these fossils provide the earliest evidence for crown-group branchiopods and total-group copepods and ostracods, extending the respective ranges of these clades back from the Devonian, Pennsylvanian, and Ordovician. Detailed similarities with living forms demonstrate the early origins and subsequent conservation of various complex food-handling adaptations, including a directional mandibular asymmetry that has persisted through half a billion years of evolution. At the same time, the Deadwood fossils indicate profound secular changes in crustacean ecology in terms of body size and environmental distribution. The earliest radiation of crustaceans is largely cryptic in the fossil record, but "small carbonaceous fossils" reveal organisms of surprisingly modern aspect operating in an unfamiliar biosphere.

  14. Growth patterns of the Cambrian microbialite: Phototropism and speciation of Epiphyton

    NASA Astrophysics Data System (ADS)

    Woo, J.; Chough, S. K.

    2010-07-01

    Microbes started constructing shallow marine stromatolitic bioherms in the Archean, but they transferred their role as a major buildup maker to metazoans in Phanerozoic. Microbial buildups often recovered their predominance in the carbonate platform when reefal metazoan communities collapsed. Epiphyton, an extinct taxon of calcified microbe that possessed branching filamentous trichomes, was an important reef builder in the shallow marine carbonate platform during Middle Cambrian, aftermath of an extinction of archaeocyath sponges which were major reef-building sessile organisms in the Early Cambrian. Here we present direct evidence of phototropism of Epiphyton, found from fossilized behavior in micro- and macro-structures of meter-scale microbial bioherms of the Zhangxia Formation (Middle Cambrian), North China Platform, Shandong Province, China. The bioherms consist of stacked growth layers with the inner and outer divisions divided by distinct boundary. The inner division of growth layers of the Epiphyton bioherm is dominated by dense uniform bush-shaped Epiphyton thalli, whereas the curved outer division has layered texture normal to the surface, comprised of elongated and chambered thalli. It suggests that photosynthetic Epiphyton reacted actively to the spatial changes in intensity of sunlight, controlled by angle of illumination on the curved growth surface of the bioherm. The inner and the outer divisions comprise different morpho-types of Epiphyton. The spatial distributions of different morpho-types in variously illuminated divisions of Epiphyton might have caused further speciation of Epiphyton.

  15. An Independent Constraint on Marine Sulfate Levels at the Ediacaran-Cambrian Transition

    NASA Astrophysics Data System (ADS)

    Blättler, C. L.; Bergmann, K.; Higgins, J. A.

    2016-12-01

    Sediments from the Ediacaran-Cambrian transition in Oman record major excursions in the isotopic systems of carbon (δ13C) and sulfur (δ34S). The significance of these geochemical signals has been the subject of much debate, focusing on their association with biotic innovations and extinctions, oxygenation of the atmosphere and oceans, and changes in seawater chemistry. The concentration of marine sulfate during this time interval remains particularly uncertain, despite being a critical variable for several hypotheses related to these excursions. We present an independent constraint on relative sulfate concentrations based on calcium-isotope data from the Ara Group evaporites (547-540 Ma) of the South Oman Salt Basin. Anhydrite samples from five boreholes span the A1 through A5 carbonate-evaporite sequences, lying on both sides of the Precambrian-Cambrian boundary. The approach developed by Blättler and Higgins (2014) is used to quantify the degree of isotopic distillation of calcium by sulfate mineral precipitation. The absence of significant distillation (expressed by δ44/40Ca variability in the anhydrite) suggests that Ca/SO4 ratios in the latest Ediacaran and earliest Cambrian were greater than during the Neogene or Permian. These results, together with previously published analyses of halite fluid inclusions and models of sulfur cycling, give a clearer estimate of how marine sulfate levels may have evolved during this dynamic period in Earth history.

  16. Morphology and ultrastructure of epilithic versus cryptic, microbial growth in lower Cambrian phosphorites from the Montagne Noire, France.

    PubMed

    Alvaro, J J; Clausen, S

    2010-03-01

    The lower Cambrian grainy phosphorites of the northern Montagne Noire occur interbedded with grey to black, laminated to massive shales and limestones deposited along the edge of a continental shelf, associated with slope-related facies and unstable substrates. The concentration of phosphate took place by repeated alternations of low sedimentation rates and condensation (hardgrounds), in situ early-diagenetic precipitation of fluorapatite, winnowing and polyphase reworking of previously phosphatized skeletons and hardground-derived clasts. The succession of repeated cycles of sedimentation, phosphate concentration, and reworking led to multi-event phosphate deposits rich in allochthonous particles. Phosphogenesis was primarily mediated by microbial activity, which is evidenced by the abundance of phosphatized putative microbial remains. These occur as smooth and segmented filaments, sheaths, and ovoid-shaped coccoids. These simple morphologies commonly form composite frameworks as a result of their aggregation and entanglement, leading to the record of biofilms, microbial mats, and complex networks. These infested the calcitic skeletonized microfossils that littered the substrate. Microbial activity evidences epilithic (anisotropic coatings on skeletons), euendolithic (perforating skeletal walls), and cryptoendolithic (lining inter- and intraparticulate pores) strategies, the latter dominated by bundles of filaments and globular clusters that grew along the cavities of helcionellids and hyoliths. According to their epilithic versus cryptic strategies, microbial populations that penetrated and dwelled inside hard skeletal substrates show different network and colonial morphologies. These early Cambrian shell concentrations were the loci of a stepwise colonization made by saprophytic to mutualistic, cyanobacterial-fungal consortia. Their euendolithic and cryptoendolithic ecological niches provided microbial refugia to manage the grazing impact mainly led by metazoans.

  17. Facies analysis of Late Proterozoic through Lower Cambrian rocks of the Death Valley regional ground-water system and surrounding areas, Nevada and California

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Sweetkind, D.S.; White, D.K.

    Late Proterozoic through Lower Cambrian rocks in the southern Great Basin form a westward-thickening wedge of predominantly clastic deposits that record deposition on the early western shelf edge of western North America (Stewart and Poole, 1974; Poole and others, 1992). Regional analyses of geologic controls on ground-water flow in the southern Great Basin typically combined lithostratigraphic units into more general hydrogeologic units that have considerable lateral extent and distinct hydrologic properties. The Late Proterozoic through Lower Cambrian rocks have been treated as a single hydrogeologic unit, named the lower clastic aquitard (Winograd and Thordarson, 1975) or the quartzite confining unitmore » (Laczniak and others, 1996), that serves as the hydrologic basement to the flow system. Although accurate in a general sense, this classification ignores well-established facies relations within these rocks that might increase bedrock permeability and locally influence ground-water flow . This report presents a facies analysis of Late Proterozoic through Lower Cambrian rocks (hereafter called the study interval) in the Death Valley regional ground-water flow system - that portion of the southern Great Basin that includes Death Valley, the Nevada Test Site, and the potential high-level nuclear waste underground repository at Yucca Mountain (fig. 1). The region discussed in this report, hereafter called the study area, covers approximately 100,000 km2 (lat 35 degrees-38 degrees 15'N., long 115 degrees-118 degrees W.). The purpose of this analysis is to provide a general documentation of facies transitions within the Late Proterozoic through Lower Cambrian rocks in order to provide an estimate of material properties (via rock type, grain size, and bedding characteristics) for specific hydrogeologic units to be included in a regional ground-water flow model.« less

  18. Late Precambrian-Cambrian sediments of Huqf group, Sultanate of Oman

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Gorin, G.E.; Racz, L.G.; Walter, M.R.

    1982-12-01

    The Huqf Group is the oldest known sedimentary sequence overlying crystalline basement in the Sultanate of Oman. It crops out on a broad regional high, the Huqf Axis, which forms a dominating structural element on the southeastern edge of the Arabian peninsula. Subsurface and outcrop evidence within and outside of Oman suggests that the sediments of the Huqf Group lie within the age span of late Precambrian to Early-Middle Cambrian. The Huqf Group is subdivided into five formations corresponding to an alternation of clastics (Abu Mahara and Shuram Formations) and carbonates (Khufai and Buah Formations) deposited in essentially shallow marinemore » to supratidal (or fluviatile) conditions and terminated by an evaporitic sequence (Ara Formation). Evaporites are absent on the Huqf Axis, but they are thickly developed to the west over a large part of southern and central Oman, where they acted as the major structure former of most of Oman's fields, and even locally pierced up to the surface. Regional correlations suggest that the predominantly carbonate-evaporitic facies of the Huqf Group was widely distributed in late Precambrian-Early Cambrian time: the Huqf basin is tentatively considered part of a belt of evaporitic basins and intervening carbonate platforms, which stretched across the Pangea landmass from the Indian subcontinent (Salt Range of Pakistan) through South Yemen, Oman, and Saudi Arabia into the gulf states and Iran (Hormuz Series and carbonate platform north of the Zagros).« less

  19. Constraints on Early Triassic carbon cycle dynamics from paired organic and inorganic carbon isotope records

    NASA Astrophysics Data System (ADS)

    Meyer, K. M.; Yu, M.; Lehrmann, D.; van de Schootbrugge, B.; Payne, J. L.

    2013-01-01

    Large δ13C excursions, anomalous carbonate precipitates, low diversity assemblages of small fossils, and evidence for marine euxinia in uppermost Permian and Lower Triassic strata bear more similarity to Neoproterozoic carbonates than to the remainders of the Permian and Triassic systems. Middle Triassic diversification of marine ecosystems coincided with the waning of anoxia and stabilization of the global carbon cycle, suggesting that environment-ecosystem linkages were important to biological recovery. However, the Earth system behavior responsible for these large δ13C excursions remains poorly constrained. Here we present a continuous Early Triassic δ13Corg record from south China and use it to test the extent to which Early Triassic excursions in δ13Ccarb record changes in the δ13C of marine dissolved inorganic carbon (DIC). Regression analysis demonstrates a significant positive correlation between δ13Corg and δ13Ccarb across multiple sections that span a paleoenvironmental gradient. Such a correlation is incompatible with diagenetic alteration because no likely mechanism will alter both δ13Corg and δ13Ccarb records in parallel and therefore strongly indicates a primary depositional origin. A simple explanation for this correlation is that a substantial portion of the preserved Corg was derived from the contemporaneous DIC pool, implying that the observed excursions reflect variation in the δ13C of the exogenic carbon reservoir (ocean, atmosphere, biomass). These findings support existing evidence that large δ13C excursions are primary and therefore strengthen the case that large-scale changes to the carbon cycle were mechanistically linked to the low diversity and small size of Early Triassic fossils. Associated sedimentary and biogeochemical phenomena further suggest that similar associations in Neoproterozoic and Cambrian strata may reflect the same underlying controls.

  20. Preservation and phylogeny of Cambrian ecdysozoans tested by experimental decay of Priapulus

    NASA Astrophysics Data System (ADS)

    Sansom, Robert S.

    2016-09-01

    The exceptionally preserved Cambrian fossil record provides unique insight into the early evolutionary history of animals. Understanding of the mechanisms of exceptional soft tissue preservation frames all interpretations of the fauna and its evolutionary significance. This is especially true for recent interpretations of preserved nervous tissues in fossil ecdysozoans. However, models of soft tissue preservation lack empirical support from actualistic studies. Here experimental decay of the priapulid Priapulus reveal consistent bias towards rapid loss of internal non-cuticular anatomy compared with recalcitrant cuticular anatomy. This is consistent with models of Burgess Shale-type preservation and indicates that internal tissues are unlikely to be preserved with fidelity if organically preserved. This pattern, along with extreme body margin distortion, is consistent with onychophoran decay, and is therefore resolved as general for early ecdysozoans. Application of these patterns to phylogenetic data finds scalidophoran taxa to be very sensitive to taphonomically informed character coding, but not panarthropodan taxa. Priapulid decay also have unexpected relevance for interpretation of myomeres in fossil chordates. The decay data presented serve not only as a test of models of preservation but also a framework with which to interpret ecdysozoan fossil anatomies, and the subsequent evolutionary inferences drawn from them.

  1. Preservation and phylogeny of Cambrian ecdysozoans tested by experimental decay of Priapulus

    PubMed Central

    Sansom, Robert S.

    2016-01-01

    The exceptionally preserved Cambrian fossil record provides unique insight into the early evolutionary history of animals. Understanding of the mechanisms of exceptional soft tissue preservation frames all interpretations of the fauna and its evolutionary significance. This is especially true for recent interpretations of preserved nervous tissues in fossil ecdysozoans. However, models of soft tissue preservation lack empirical support from actualistic studies. Here experimental decay of the priapulid Priapulus reveal consistent bias towards rapid loss of internal non-cuticular anatomy compared with recalcitrant cuticular anatomy. This is consistent with models of Burgess Shale-type preservation and indicates that internal tissues are unlikely to be preserved with fidelity if organically preserved. This pattern, along with extreme body margin distortion, is consistent with onychophoran decay, and is therefore resolved as general for early ecdysozoans. Application of these patterns to phylogenetic data finds scalidophoran taxa to be very sensitive to taphonomically informed character coding, but not panarthropodan taxa. Priapulid decay also have unexpected relevance for interpretation of myomeres in fossil chordates. The decay data presented serve not only as a test of models of preservation but also a framework with which to interpret ecdysozoan fossil anatomies, and the subsequent evolutionary inferences drawn from them. PMID:27595908

  2. 3D-Mapping of Dolomitized Structures in Lower Cambrian Phosphorites

    NASA Astrophysics Data System (ADS)

    Hippler, Dorothee; Stammeier, Jessica A.; Brunner, Roland; Rosc, Jördis; Franz, Gerhard; Dietzel, Martin

    2016-04-01

    Dolomitization is a widespread phenomenon in ancient sedimentary rocks, particularly close to the Precambrian-Cambrian boundary. Dolomite can form in synsedimentary or hydrothermal environments, preferentially via the replacement of solid carbonate precursor phases. Synsedimentary dolomite formation is often associated with microbial activity, such as bacterial sulfate reduction or methanogenesis. In this study, we investigate dolomitic phosphorites from the Lowermost Cambrian Tal Group, Mussoori Syncline, Lesser Himalaya, India, using micro-CT 3D-mapping, in order to unravel the complex diagenetic history of the rocks. The selected sample shows alternating layering of phosphatic mudstones and sparitic dolostone, in which brecciated layers of phosphorite or phosphatic mudstones are immersed in a dolomite-rich matrix. Lamination occurs on a sub-millimetre scale, with lamination sometimes wavy to crinkly. This fabric is interpreted as former microbial mats, providing the environment for early diagenetic phosphatization. Preliminary electron backscatter imaging with scanning microscopy revealed that dolomite crystals often occur in spherical to ellipsoidal structures, typically with a high porosity. This dolomite is associated with botryoidal apatite, organic matter and small amounts of calcite. Micro-CT 3D-mappings reveal that dolomite structures are cigar-shaped, elongated and up to 600 μm long. They are further arranged in a Mikado-like oriented framework spanning a layer thickness of a few millimetres. Analyses of ambient pore space, with similar elongated outlines and filled with organic matter, suggest a potential coherence of ambient pore space and shape of the dolomite structures. Allowing for other associated mineral phases, such as pyrite and silicates, and their spatial distribution, the present approach can be used to unravel distinct diagenetic reaction pathways, and might thus constrain the proxy potential of these Lower Cambrian dolomitic phosphorites

  3. Reconstructing the Avalon continent: Marginal to inner platform transition in the Cambrian of southern New Brunswick

    USGS Publications Warehouse

    Landing, E.

    1996-01-01

    A west to east, marginal to inner Avalonian platform transition, comparable to that in southeast Newfoundland and southern Britain, is present in the Cambrian of southern New Brunswick. The Saint John - Caton's Island - Hanford Brook area lay on the marginal platform, and its thick, uppermost Precambrian - lower Lower Cambrian is unconformably overlain by trilobite-bearing, upper Lower Cambrian. An inner platform remnant is preserved in the Cradle Brook outlier 60 km northeast of Saint John. In contrast to the marginal platform sequences, the Cradle Brook outlier has a very thin lower Lower Cambrian and has middle Lower Cambrian strata (Bonavista Group) not present on the marginal platform. The Cradle Brook Lower Cambrian closely resembles inner platform successions in eastern Massachusetts and Trinity and Placentia bays, southeast Newfoundland. A limestone with Camenella baltica Zone fossils on Cradle Brook seems to be the peritidal limestone cap of the subtrilobitic Lower Cambrian known in Avalonian North America (Fosters Point Formation) and England (Home Farm Member).

  4. The Cambrian Evolutionary Explosion: Novel Evidence from Fossils Studied by X-ray Tomography

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Chen, Jun-Yuan

    The Cambrian explosion (from 542 million years to 488 million years ago) is one of the greatest mysteries in evolutionary biology. It wasn't until this period that complex organisms became common and diverse. the magnitude of the event can be understood based on the contrast between the biota and the degree of diversity of the fossils from both sides. great advances have been made in Cambrian palaeontology over the past century, especially the discovery of the well-preserved soft-bodied fauna from the Middle Cambrian Burgess Shale and the Lower Cambrian Maotianshan Shale deposits. The Cambrian side of the "Cambrian explosion" ismore » richly illustrated and contrasts greatly with the Precambrian side. The study of these extraordinarily preserved fossil biota is extremely difficult. A major challenge is 3-D reconstruction and determining the patter of the cell organization in Weng'an embryos and their buried structures in Maotianshan Shale fossils. This talk will show that two recent technological approaches, propagation phase contrast synchrotron x-ray microtomography and microtomography, provide unique analytical tools that permit the nondestructive computational examination and visualization of the internal and buried characters in virtual sections in any plane, and virtual 3-D depictions of internal structures.« less

  5. Depositional Architecture of Late Cambrian-Early Ordovician Siliciclastic Barik Formation; Al Huqf Area, Oman

    NASA Astrophysics Data System (ADS)

    Abbasi, Iftikhar Ahmed

    2017-04-01

    Early Paleozoic siliciclastics sediments of the Haima Supergroup are subdivided into a number of formations and members based on lithological characteristics of various rock sequences. One of the distinct sandstone sequence, the Barik Formation (Late Cambrian-Early Ordovician) of the Andam Group is a major deep gas reservoir in central Oman. The sandstone bodies are prospective reservoir rocks while thick shale and clay interbeds act as effective seal. Part of the Barik Formation (lower and middle part) is exposed in isolated outcrops in Al Huqf area as interbedded multistoried sandstone, and green and red shale. The sandstone bodies are up to 2 meters thick and can be traced laterally for 300 m to over 1 km. Most of sandstone bodies show both lateral and vertical stacking. Two types of sandstone lithofacies are identified on the basis of field characteristics; a plane-bedded sandstone lithofacies capping thick red and green color shale beds, and a cross-bedded sandstone lithofacies overlying the plane-bedded sandstone defining coarsening upward sequences. The plane-bedded sandstone at places contains Cruziana ichnofacies and bivalve fragments indicating deposition by shoreface processes. Thick cross-bedded sandstone is interpreted to be deposited by the fluvial dominated deltaic processes. Load-casts, climbing ripples and flaser-bedding in siltstone and red shale indicate influence of tidal processes at times during the deposition of the formation. This paper summarizes results of a study carried out in Al Huqf area outcrops to analyze the characteristics of the sandstone-body geometry, internal architecture, provenance and diagenetic changes in the lower and middle part of the formation. The study shows build-up of a delta complex and its progradation over a broad, low-angle shelf where fluvial processes operate beside shoreface processes in a vegetation free setting. Keywords: Andam Group, Barik Formation, Ordovician sandstone, Al Huqf, Central Oman,

  6. Cambrian Sauk transgression in the Grand Canyon region redefined by detrital zircons

    NASA Astrophysics Data System (ADS)

    Karlstrom, Karl; Hagadorn, James; Gehrels, George; Matthews, William; Schmitz, Mark; Madronich, Lauren; Mulder, Jacob; Pecha, Mark; Giesler, Dominique; Crossey, Laura

    2018-06-01

    The Sauk transgression was one of the most dramatic global marine transgressions in Earth history. It is recorded by deposition of predominantly Cambrian non-marine to shallow marine sheet sandstones unconformably above basement rocks far into the interiors of many continents. Here we use dating of detrital zircons sampled from above and below the Great Unconformity in the Grand Canyon region to bracket the timing of the Sauk transgression at this classic location. We find that the Sixtymile Formation, long considered a Precambrian unit beneath the Great Unconformity, has maximum depositional ages that get younger up-section from 527 to 509 million years old. The unit contains angular unconformities and soft-sediment deformation that record a previously unknown period of intracratonic faulting and epeirogeny spanning four Cambrian stages. The overlying Tapeats Sandstone has youngest detrital zircon ages of 505 to 501 million years old. When linked to calibrated trilobite zone ages of greater than 500 million years old, these age constraints show that the marine transgression across a greater than 300-km-wide cratonic region took place during an interval 505 to 500 million years ago—more recently and more rapidly than previously thought. We redefine this onlap as the main Sauk transgression in the region. Mechanisms for this rapid flooding of the continent include thermal subsidence following the final breakup of Rodinia, combined with abrupt global eustatic changes driven by climate and/or mantle buoyancy modifications.

  7. The anatomy, taphonomy, taxonomy and systematic affinity of Markuelia: Early Cambrian to Early Ordovician scalidophorans

    USGS Publications Warehouse

    Dong, X.-P.; Bengtson, S.; Gostling, N.J.; Cunningham, J.A.; Harvey, T.H.P.; Kouchinsky, A.; Val'Kov, A.K.; Repetski, J.E.; Stampanoni, M.; Marone, F.; Donoghue, P.C.J.

    2010-01-01

    Markuelia is a vermiform, annulated introvertan animal known as embryonic fossils from the Lower Cambrian to Lower Ordovician. Analysis of an expanded and revised dataset for Introverta shows that the precise position of Markuelia within this clade is dependent on the taxa included. As a result, Markuelia is assigned to the scalidophoran total group to reflect uncertainty as to whether it is a stem-scalidophoran or a stem-priapulid. The taxonomy of the genus is revised to provide an improved taxonomic framework for material assigned to Markuelia. Five species are recognized: M. secunda Val'kov, M. hunanensis Dong and Donoghue, M. lauriei Haug et al., M. spinulifera sp. nov. and M. waloszeki sp. nov. Finally, the preservation of Markuelia is evaluated in the light of both the taphonomy of the fossil embryos themselves and the experimental taphonomy of the priapulid Priapulus caudatus, which has been proposed as both a close relative and an anatomical analogue of Markuelia. ?? The Palaeontological Association.

  8. High-Precision U-Pb Geochronology and Correlation: An example Using the Neoproterozic-Cambrian Transition

    NASA Astrophysics Data System (ADS)

    Bowring, S. A.; Grotzinger, J. P.; Amthor, J.; Martin, M. E.

    2001-05-01

    The precise, global correlation of Precambrian and Paleozoic sedimentary rocks can be achieved using temporally calibrated chemostratigraphic records. This approach is essential for determining rates and causes of environmental and faunal change, including mass extinctions. For example, The Neoproterozoic is marked by major environmental change, including periods of global glaciation, large fluctuations in the sequestration of carbon and major tectonic reorganization followed by the explosive diversification of animals in the earliest Cambrian. The extreme climatic change associated with these glaciations have been implicated as a possible trigger for the Cambrian explosion. The recognition of thin zircon-bearing air-fall ash in Neoproterozoic and Cambrian rocks has allowed the establishment of a high-precision temporal framework for animal evolution and is helping to untangle the history of glaciations. In some cases analytical uncertainties translate to age uncertainties of less than 1 Ma and when integrated with chemostratigraphy, the potential for global correlations at even higher resolution. Progress in the global correlation of Neoproterozoic strata has been achieved through the use of C and Sr isotope chemostratigraphy although it has been hampered by a lack of precise geochronological and faunal control. For example, the period from ca 800-580 Ma is characterized by at least two and perhaps as many as four glacial events that are interpreted by many to be global glaciations on a "Snowball Earth". A lack of precise chronological constraints on the number and duration of glaciations, multiple large excursions in the carbon isotopic record, and an absence of detailed biostratigraphy have complicated global correlation and hindered our understanding of this important period of Earth history. However, the ongoing integration of chemostratigraphic and geochronological data are improving temporal resolution and detailed correlations. These data are critical for

  9. Early animal evolution: emerging views from comparative biology and geology

    NASA Technical Reports Server (NTRS)

    Knoll, A. H.; Carroll, S. B.

    1999-01-01

    The Cambrian appearance of fossils representing diverse phyla has long inspired hypotheses about possible genetic or environmental catalysts of early animal evolution. Only recently, however, have data begun to emerge that can resolve the sequence of genetic and morphological innovations, environmental events, and ecological interactions that collectively shaped Cambrian evolution. Assembly of the modern genetic tool kit for development and the initial divergence of major animal clades occurred during the Proterozoic Eon. Crown group morphologies diversified in the Cambrian through changes in the genetic regulatory networks that organize animal ontogeny. Cambrian radiation may have been triggered by environmental perturbation near the Proterozoic-Cambrian boundary and subsequently amplified by ecological interactions within reorganized ecosystems.

  10. Upper Neoproterozoic-Lower Cambrian sedimentary successions in the Central Iberian Zone (Spain): sequence stratigraphy, petrology and chemostratigraphy. Implications for other European zones

    NASA Astrophysics Data System (ADS)

    Valladares, M. I.; Barba, P.; Ugidos, J. M.; Colmenero, J. R.; Armenteros, I.

    The Upper Neoproterozoic-Lower Cambrian sedimentary succession in the central areas of the Central Iberian Zone has been subdivided into 12 mostly siliciclastic lithostratigraphic units, ranging in thickness between 1800 and 3900m. The lithology and facies of each unit are described and the facies associations are interpreted. The facies resulted mainly from turbidity currents and debris flows and, to a lesser extent, from submarine slides and traction flows. The facies associations suggest that sedimentation took place in slope and base-of-slope environments. Two depositional sequences are recognized, separated by a type-1 unconformity. The lower sequence is of Late Neoproterozoic age (units I-IV) and exhibits lowstand, transgressive, and highstand systems tracts. Most of the upper sequence is probably of Early Cambrian age (units V-XII). It begins at the base of unit V and possibly ends with the Tamames Limestone Formation. The upper sequence records a lowstand systems tract and minor-order sea-level oscillations. In the Cambrian units there are higher amounts of feldspar and smaller quantities of intrabasinal clasts than in the Neoproterozoic units. The modal data plot close to the Q-L and Qm-Lt sides of Q-F-L and Qm-F-Lt triangular diagrams, suggesting a provenance from a recycled orogen evolving into a provenance from a craton interior towards the top of the succession. The chemical results, based mainly on Al2O3, TiO2, Zr, and Nb abundances in shales from all the units, strongly suggest a gradual compositional change within this sedimentary succession. Together with the petrological data, the chemical results do not reveal any obvious coeval volcanic contribution to the sediments. On the basis of the chemical data, a comparison is made with other European zones containing detrital sediments composed of reworked crustal components.

  11. 500 Myr of thermal history elucidated by multi-method detrital thermochronology of North Gondwana Cambrian sandstone (Eilat area, Israel)

    NASA Astrophysics Data System (ADS)

    Vermeesch, P.; Avigad, D.

    2009-04-01

    Following the Neoproterozoic Pan-African orogeny, the Arabian-Nubian Shield (ANS) of North Africa and Arabia was eroded and then covered by Cambrian sandstones that record the onset of platform sedimentation. We applied K-feldspar 40Ar/39Ar, zircon and apatite fission track and apatite (U-Th)/He thermochronology to detritus from Cambrian sandstones of southern Israel deposited at about 500 Ma. U-Pb detrital zircon ages from these sandstones predate deposition and record the earlier Neoproterozoic crustal evolution of the Pan-African orogens. 40Ar/39Ar ages from 50 single grains of K-feldspar yield a Cambrian mean of approximately 535 Ma. The 40Ar/39Ar age spectrum of a multi-grain K-feldspar aliquot displays diffusion behaviour compatible with >560 Ma cooling later affected by a heating event. Assuming that the high temperature domains of the K-feldspars have not been affected by subsequent (hydro)thermal events, and taking previously published K-Ar and Rb-Sr ages from other parts of the East African Orogen at face value, these ages apparently record Pan-African thermal resetting below a thick volcano-sedimentary pile similar to the Saramuj conglomerate in Jordan and/or the Hammamat in Egypt. Detrital zircon fission track (ZFT) ages cluster around 380 Ma, consistent with previous ZFT results from Neoproterozoic basement and sediments of the region, revealing that the Cambrian platform sequence experienced a middle Devonian thermal event and low-grade metamorphism. Regional correlation indicates that during Devonian time the sedimentary cover atop the Cambrian in Israel was never in excess of 2.5 km, requiring an abnormally steep geothermal gradient to explain the complete ZFT annealing. A basal Carboniferous unconformity can be traced from Syria to southern Saudi Arabia, suggesting that the observed Devonian ZFT ages represent a regional tectonothermal event. Similar Devonian ZFT ages were reported from ANS basement outcrops in the Eastern Desert, 500 km south of

  12. The significance of 24-norcholestanes, triaromatic steroids and dinosteroids in oils and Cambrian-Ordovician source rocks from the cratonic region of the Tarim Basin, NW China

    USGS Publications Warehouse

    Li, Meijun; Wang, T.-G.; Lillis, Paul G.; Wang, Chunjiang; Shi, Shengbao

    2012-01-01

    Two oil families in Ordovician reservoirs from the cratonic region of the Tarim Basin are distinguished by the distribution of regular steranes, triaromatic steroids, norcholestanes and dinosteroids. Oils with relatively lower contents of C28 regular steranes, C26 20S, C26 20R + C27 20S and C27 20R regular triaromatic steroids, dinosteranes, 24-norcholestanes and triaromatic dinosteroids originated from Middle–Upper Ordovician source rocks. In contrast, oils with abnormally high abundances of the above compounds are derived from Cambrian and Lower Ordovician source rocks. Only a few oils have previously been reported to be of Cambrian and Lower Ordovician origin, especially in the east region of the Tarim Basin. This study further reports the discovery of oil accumulations of Cambrian and Lower Ordovician origin in the Tabei and Tazhong Uplifts, which indicates a potential for further discoveries involving Cambrian and Lower Ordovician sourced oils in the Tarim Basin. Dinosteroids in petroleum and ancient sediments are generally thought to be biomarkers for dinoflagellates and 24-norcholestanes for dinoflagellates and diatoms. Therefore, the abnormally high abundance of these compounds in extracts from the organic-rich sediments in the Cambrian and Lower Ordovician and related oils in the cratonic region of the Tarim Basin suggests that phytoplankton algae related to dinoflagellates have appeared and might have flourished in the Tarim Basin during the Cambrian Period. Steroids with less common structural configurations are underutilized and can expand understanding of the early development history of organisms, as well as define petroleum systems.

  13. Oxygen-isotope trends and seawater temperature changes across the Late Cambrian Steptoean positive carbon-isotope excursion (SPICE event)

    USGS Publications Warehouse

    Elrick, M.; Rieboldt, S.; Saltzman, M.; McKay, R.M.

    2011-01-01

    The globally recognized Late Cambrian Steptoean positive C-isotope excursion (SPICE) is characterized by a 3???-5??? positive ??13C shift spanning <4 m.y. Existing hypotheses suggest that the SPICE represents a widespread ocean anoxic event leading to enhanced burial/preservation of organic matter (Corg) and pyrite. We analyzed ??18O values of apatitic inarticulate brachiopods from three Upper Cambrian successions across Laurentia to evaluate paleotemperatures during the SPICE. ??18O values range from ~12.5??? to 16.5???. Estimated seawater temperatures associated with the SPICE are unreasonably warm, suggesting that the brachiopod ??18O values were altered during early diagenesis. Despite this, all three localities show similar trends with respect to the SPICE ??13C curve, suggesting that the brachiopod apatite preserves a record of relative ??18O and temperature changes. The trends include relatively high ??18O values at the onset of the SPICE, decreasing and lowest values during the main event, and an increase in values at the end of the event. The higher ??18O values during the global extinction at the onset of the SPICE suggests seawater cooling and supports earlier hypotheses of upwelling of cool waters onto the shallow shelf. Decreasing and low ??18O values coincident with the rising limb of the SPICE support the hypothesis that seawater warming and associated reduced thermohaline circulation rates contributed to decreased dissolved O2 concentrations, which enhanced the preservation/burial of Corg causing the positive ??13C shift. ?? 2011 Geological Society of America.

  14. Carbonate rocks of Cambrian and Ordovician age in the Lancaster quadrangle, Pennsylvania

    USGS Publications Warehouse

    Meisler, Harold; Becher, Albert E.

    1968-01-01

    Detailed mapping has shown that the carbonate rocks of Cambrian and Ordovician age in the Lancaster quadrangle, Pennsylvania, can be divided into 14 rock-stratigraphic units. These units are defined primarily by their relative proportions of limestone and dolomite. The oldest units, the Vintage, Kinzers, and Ledger Formations of Cambrian age, and the Conestoga Limestone of Ordovician age are retained in this report. The Zooks Corner Formation, of Cambrian age, a dolomite unit overlying the Ledger Dolomite, is named here for exposures along Conestoga Creek near the village of Zooks Corner. The Conococheague (Cambrian) and Beekmantown (Ordovician) Limestones, as mapped by earlier workers, have been elevated to group rank and subdivided into formations that are correlated with and named for geologic units in Lebanon and Berks Counties, Pa. These formations, from oldest to youngest, are the Buffalo Springs, Snitz Creek, Millbach, and Richland Formations of the Conococheague Group, and the Stonehenge, Bpler, and Ontelaunee Formations of the Beekmantown Group. The Annville and Myerstown Limestones, which are named for lithologically similar units in Dauphin and Lebanon Counties, Pa., overlie the Beekmantown Group in one small area in the quadrangle.

  15. The Late Cambrian Takaka Terrane, NW Nelson, New Zealand: Accretionary-prism development and arc collision followed by extension and fan-delta deposition at the SE margin of Gondwana

    NASA Astrophysics Data System (ADS)

    Pound, K. S.

    2013-12-01

    convergence, but is reinterpreted here as a ';true' fan-delta deposit, sedimentologically similar to deposits associated with extension. Textural and compositional data for the Lockett Conglomerate indicates rapid supply of new material (including quartzite, granite, gabbro, and amphibolitic metavolcanics). The Lockett Conglomerate is proposed here to record the initiation of extension, during which basement faults in the hinterland exposed previously buried source rocks. This new interpretation of the Lockett Conglomerate places that initiation of extension and subsequent passive margin sedimentation (Mt. Ellis and Mt. Arthur Groups) earlier (late Middle Cambrian) than previous work has suggested (Late Cambrian or Early Ordovician). These new interpretations provide input useful for correlations and interpretations of the complex mosaic that preserves a record of tectonic activity and processes at the Antarctic, Tasmanian and SE Australian portions of the Cambrian Gondwana margin.

  16. Stratigraphic Framework of Cambrian and Ordovician Rocks in the Appalachian Basin from Sequatchie County, Tennessee, through Eastern Kentucky, to Mingo County, West Virginia

    USGS Publications Warehouse

    Ryder, Robert T.; Crangle, Robert D.; Repetski, John E.; Harris, Anita G.

    2008-01-01

    Cross section H-H' is the seventh in a series of restored cross sections constructed by the lead author to show the stratigraphic framework of Cambrian and Ordovician rocks in the Appalachian basin from Pennsylvania to Tennessee. The sections show complexly intertongued carbonate and siliciclastic lithofacies, marked thickness variations, key marker horizons, unconformities, stratigraphic nomenclature of the Cambrian and Ordovician sequence, and major faults that offset Proterozoic basement and overlying lower Paleozoic rocks. Several of the drill holes along the cross section have yielded a variety of whole and (or) fragmented conodont elements. The identifiable conodonts are used to differentiate strata of Late Cambrian, Early Ordovician, and Middle Ordovician age, and their conodont color alteration index (CAI) values are used to establish the thermal maturity of the sequence. Previous cross sections in this series are G-G', F-F', E-E', D-D', C-C', and B-B'. Many of these cross sections (B-B', C-C', D-D', and G-G') have been improved with the addition of gamma-ray log traces, converted to digital images, and made accessible on the Web.

  17. The origin of animals: Can molecular clocks and the fossil record be reconciled?

    PubMed

    Cunningham, John A; Liu, Alexander G; Bengtson, Stefan; Donoghue, Philip C J

    2017-01-01

    The evolutionary emergence of animals is one of the most significant episodes in the history of life, but its timing remains poorly constrained. Molecular clocks estimate that animals originated and began diversifying over 100 million years before the first definitive metazoan fossil evidence in the Cambrian. However, closer inspection reveals that clock estimates and the fossil record are less divergent than is often claimed. Modern clock analyses do not predict the presence of the crown-representatives of most animal phyla in the Neoproterozoic. Furthermore, despite challenges provided by incomplete preservation, a paucity of phylogenetically informative characters, and uncertain expectations of the anatomy of early animals, a number of Neoproterozoic fossils can reasonably be interpreted as metazoans. A considerable discrepancy remains, but much of this can be explained by the limited preservation potential of early metazoans and the difficulties associated with their identification in the fossil record. Critical assessment of both records may permit better resolution of the tempo and mode of early animal evolution. © 2016 The Authors BioEssays Published by WILEY Periodicals, Inc.

  18. Recognition of Macluritella ( Gastropoda) from the Upper Cambrian of Missouri and Nevada ( USA).

    USGS Publications Warehouse

    Yochelson, E.L.; Stinchcomb, B.L.

    1987-01-01

    Open-coiled euomphalacean gastropods have been identified for the first time in the Upper Cambrian Eminence Dolomite of Missouri. These gastropods have a triangular whorl profile and are conspecific with Hyolithes walcotti described from the Upper Cambrian of Nevada. That species is questionably reassigned to the gastropod genus Macluritella, hitherto known only from the Lower Ordovician of Colorado. -Authors Ordovician Colorado

  19. Middle to late Cambrian shallow marine trace fossils from the Imfout Syncline (Western Meseta, Morocco): Palaeoecological and palaeoenvironmental significance in NW-Gondwana

    NASA Astrophysics Data System (ADS)

    Oukassou, Mostafa; Lagnaoui, Abdelouahed; Raji, Mohammed; Michard, André; Saddiqi, Omar

    2017-05-01

    The present research provides the first evidence of invertebrate activity assigned to the ichnogenus Selenichnites occurring together with moderately diverse ichnofossils from the middle to late Cambrian of the Moroccan Meseta. The invertebrate traces occur in sandstone strata of the El Hank Formation within the Imfout Syncline, in the northern part of the Rehamna Massif (Coastal Block, western Moroccan Meseta). Bedding surfaces from the top of the El Hank Formation near the Imfout Dam show diverse forms of current ripples and distincts crescentic ichnofossils in concave epirelief scattered on the surface. In this section, the traces provide evidence of the ethology of an organism inhabiting the relatively shallow waters of the area during this time. Selenichnites co-occurs with the ichnogenera Arenicolites, Diplocraterion, Lingulichnus, Monocraterion, Skolithos and unidentified burrows, and the ichnoassemblage is referred to the Skolithos ichnofacies. These traces can be referred to arthropods (e.g. polychaete worms and amphipod crustaceans), lingulid brachiopods, annelids and/or phoronids. The Imfout Selenichnites represents the first occurrence of this ichnogenus from the Cambrian of the Moroccan Meseta, and the second from the Cambrian deposits of Morocco. The potential tracemakers are still questionable, but were most likely xiphosurans, trilobites, euthycarcinoids or crustaceans. If so, the Imfout traces could be among the oldest pieces of evidence for the presence of horseshoe crabs during the Cambrian. The combination of sedimentological and ichnological data indicates that the El Hank Formation was deposited in a sublittoral soft ground environment next to a sandy shore. It was originally part of an early Palaeozoic shallow marine epicontinental platform in west-central Morocco. In addition to the equivalent Cambrian deposits from the Anti-Atlas, the El Hank Formation constituted a part of the northern Gondwana platform domain during the transgression

  20. The origin of the animals and a 'Savannah' hypothesis for early bilaterian evolution.

    PubMed

    Budd, Graham E; Jensen, Sören

    2017-02-01

    The earliest evolution of the animals remains a taxing biological problem, as all extant clades are highly derived and the fossil record is not usually considered to be helpful. The rise of the bilaterian animals recorded in the fossil record, commonly known as the 'Cambrian explosion', is one of the most significant moments in evolutionary history, and was an event that transformed first marine and then terrestrial environments. We review the phylogeny of early animals and other opisthokonts, and the affinities of the earliest large complex fossils, the so-called 'Ediacaran' taxa. We conclude, based on a variety of lines of evidence, that their affinities most likely lie in various stem groups to large metazoan groupings; a new grouping, the Apoikozoa, is erected to encompass Metazoa and Choanoflagellata. The earliest reasonable fossil evidence for total-group bilaterians comes from undisputed complex trace fossils that are younger than about 560 Ma, and these diversify greatly as the Ediacaran-Cambrian boundary is crossed a few million years later. It is generally considered that as the bilaterians diversified after this time, their burrowing behaviour destroyed the cyanobacterial mat-dominated substrates that the enigmatic Ediacaran taxa were associated with, the so-called 'Cambrian substrate revolution', leading to the loss of almost all Ediacara-aspect diversity in the Cambrian. Why, though, did the energetically expensive and functionally complex burrowing mode of life so typical of later bilaterians arise? Here we propose a much more positive relationship between late-Ediacaran ecologies and the rise of the bilaterians, with the largely static Ediacaran taxa acting as points of concentration of organic matter both above and below the sediment surface. The breaking of the uniformity of organic carbon availability would have signalled a decisive shift away from the essentially static and monotonous earlier Ediacaran world into the dynamic and burrowing world

  1. Relationship between the Neoproterozoic snowball Earth and Cambrian explosion

    NASA Astrophysics Data System (ADS)

    Maruyama, S.; Yoshihara, A.; Isozaki, Y.

    2007-12-01

    Origin of snowball Earth has been debated in terms of greenhouse gas (e.g., Hoffman and Schrag), obliqueness of Earth's rotation axis (Williams, 1975), true polar wander (Evans, 2003), Galactic cosmic ray radiation (Shaviv and Veizer, 2003; Svensmark, 2006), or weakened geomagnetism (Maruyama and Yoshihara, 2003). A major difficulty for the greenhouse gas hypothesis is the on-off switch causing decrease and increase of appropriate amounts of CO2 by plume- and plate tectonics, and also in available amount of CO2 in atmosphere to be consistent with the observations. In contrast, the cosmic ray radiation models due to the star burst peaked at 2.5- 2.1 Ga and 1.4-0.8 Ga can explain on-off switch more easily than the greenhouse gas model. Cosmic ray radiations, however, must be modified by the geomagnetic intensity, fluctuating 150"% to < 10"% of the present-day level through geologic time. Our compilation suggests the idea of extensive glaciation appeared when the intensity decreased below 50% of the present-day value, as typically seen in the Neoproterozoic time. This proposes the idea of extensive cloudiness by increased cosmic rays in the Neoproterozoic to cause the snowball Earth. Time difference between the Neoproterozoic snowball Earth and Cambrian explosion is as large as 250 millions years, and this refuses their direct close-relationship. Role of frequent mass extinctions, i.e., 8 times during 100 m.y. from 585 Ma to 488 Ma, during the Ediacaran and Cambrian, has been proposed (Zhu et al., 2007). This frequency is one order of magnitude higher compared to that in the post-Ordovician time. Yet, the Cambrian explosion cannot be explained by mass extinction which replaced the vacant niches shortly after the mass extinction and never created a new animal with a new body plan. A new model proposed herein is derived from weakened geomagnetism and resultant extensive cosmic radiation to alter gene and genome for a long period over advancement of low magnetic

  2. Sequence stratigraphy of the Upper Cambrian (Furongian; Jiangshanian and Sunwaptan) Tunnel City Group, Upper Mississippi Valley: Transgressing assumptions of cratonic flooding

    USGS Publications Warehouse

    Eoff, Jennifer D.

    2014-01-01

    New data from detailed measured sections permit comprehensive analysis of the sequence framework of the Furongian (Upper Cambrian; Jiangshanian and Sunwaptan stages) Tunnel City Group (Lone Rock Formation and Mazomanie Formation) of Wisconsin and Minnesota. The sequence-stratigraphic architecture of the lower part of the Sunwaptan Stage at the base of the Tunnel City Group, at the contact between the Wonewoc Formation and Lone Rock Formation, records the first part of complex polyphase flooding (Sauk III) of the Laurentian craton, at a scale smaller than most events recorded by global sea-level curves. Flat-pebble conglomerate and glauconite document transgressive ravinement and development of a condensed section when creation of accommodation exceeded its consumption by sedimentation. Thinly-bedded, fossiliferous sandstone represents the most distal setting during earliest highstand. Subsequent deposition of sandstone characterized by hummocky or trough cross-stratification records progradational pulses of shallower, storm- and wave-dominated environments across the craton before final flooding of Sauk III commenced with carbonate deposition during the middle part of the Sunwaptan Stage. Comparison of early Sunwaptan flooding of the inner Laurentian craton to published interpretations from other parts of North America suggests that Sauk III was not a single, long-term accommodation event as previously proposed.

  3. Cambrian-lower Middle Ordovician passive carbonate margin, southern Appalachians: Chapter 14

    USGS Publications Warehouse

    Read, J. Fred; Repetski, John E.

    2012-01-01

    The southern Appalachian part of the Cambrian–Ordovician passive margin succession of the great American carbonate bank extends from the Lower Cambrian to the lower Middle Ordovician, is as much as 3.5 km (2.2 mi) thick, and has long-term subsidence rates exceeding 5 cm (2 in.)/k.y. Subsiding depocenters separated by arches controlled sediment thickness. The succession consists of five supersequences, each of which contains several third-order sequences, and numerous meter-scale parasequences. Siliciclastic-prone supersequence 1 (Lower Cambrian Chilhowee Group fluvial rift clastics grading up into shelf siliciclastics) underlies the passive margin carbonates. Supersequence 2 consists of the Lower Cambrian Shady Dolomite–Rome-Waynesboro Formations. This is a shallowing-upward ramp succession of thinly bedded to nodular lime mudstones up into carbonate mud-mound facies, overlain by lowstand quartzose carbonates, and then a rimmed shelf succession capped by highly cyclic regressive carbonates and red beds (Rome-Waynesboro Formations). Foreslope facies include megabreccias, grainstone, and thin-bedded carbonate turbidites and deep-water rhythmites. Supersequence 3 rests on a major unconformity and consists of a Middle Cambrian differentiated rimmed shelf carbonate with highly cyclic facies (Elbrook Formation) extending in from the rim and passing via an oolitic ramp into a large structurally controlled intrashelf basin (Conasauga Shale). Filling of the intrashelf basin caused widespread deposition of thin quartz sandstones at the base of supersequence 4, overlain by widespread cyclic carbonates (Upper Cambrian lower Knox Group Copper Ridge Dolomite in the south; Conococheague Formation in the north). Supersequence 5 (Lower Ordovician upper Knox in the south; Lower to Middle Ordovician Beekmantown Group in the north) has a basal quartz sandstone-prone unit, overlain by cyclic ramp carbonates, that grade downdip into thrombolite grainstone and then storm

  4. Evolution of oceanic molybdenum and uranium reservoir size around the Ediacaran-Cambrian transition: Evidence from western Zhejiang, South China

    NASA Astrophysics Data System (ADS)

    Xiang, Lei; Schoepfer, Shane D.; Shen, Shu-zhong; Cao, Chang-qun; Zhang, Hua

    2017-04-01

    The "Cambrian explosion" is one of the most fascinating episodes of diversification in the history of life; however, its relationship to the oxygenation of the oceans and atmosphere around the Ediacaran-Cambrian transition is not fully understood. Marine inventories of redox-sensitive trace elements reflect the relative balance of oxidative weathering on land and deposition in anoxic water masses, and can be used to explore the evolution of oceanic and atmospheric redox conditions. For this study, we conducted a series of geochemical analyses on the upper Lantian, Piyuancun, and Hetang formations in the Chunye-1 well, part of the lower Yangtze Block in western Zhejiang. Iron speciation results indicate that the entire studied interval was deposited under anoxic conditions, with three intervals of persistent euxinia occurring in the uppermost Lantian Fm., the lower Hetang Formation (Fm.), and the upper Hetang Fm. Molybdenum (Mo) and uranium (U) contents and Mo/TOC and U/TOC ratios from the anoxic/euxinic intervals of the Chunye-1 well, combined with published data from the sections in the middle and upper Yangtze Block, suggest that the oceanic Mo reservoir declined consistently from the Ediacaran to Cambrian Stage 3, while the size of the oceanic U reservoir remained relatively constant. Both metals were depleted in the ocean in lower Cambrian Stage 4, before increasing markedly at the end of Stage 4. The lack of an apparent increase in the size of the marine Mo and U reservoir from the upper Ediacaran to Cambrian Stage 3 suggests that oxic water masses did not expand until Cambrian Stage 4. The increase in marine Mo and U availability in the upper Hetang Fm. may have been due to the expansion of oxic water masses in the oceans, associated with oxygenation of the atmosphere during Cambrian Stage 4. This expansion of oxic waters in the global ocean postdates the main phase of Cambrian diversification, suggesting that pervasive oxygenation of the ocean on a large

  5. The Cambrian Ross Orogeny in northern Victoria Land (Antarctica) and New Zealand: A synthesis

    USGS Publications Warehouse

    Federico, L.; Capponi, G.; Crispini, L.; Bradshaw, J.D.

    2007-01-01

    In the Cambrian, the paleo-Pacific margin of the Gondwana supercontinent included East Antarctica, Australia, Tasmania and New Zealand and was affected by themajor Ross-Delamerian Orogeny. In Antarctica, evidence suggests that this resulted from oblique subduction and that in northern Victoria Land it was accompanied by the opening and subsequent closure of a back-arc basin. Comparison of the type and timing of sedimentary, magmatic and metamorphic events in areas noted above shows strong similarities between northern Victoria Land and New Zealand. In both regions Middle Cambrian volcanites are interpreted as arc/back-arc assemblages produced by west-directed subduction; sediments interbedded with the volcanites show provenance both from the arc and from the Gondwana margin and therefore place the basin close to the continent. Back-arc closure in the Late Cambrian was likely accomplished through a second subduction system

  6. Cambrian Evolutionary Radiation: Context, correlation, and chronostratigraphy—Overcoming deficiencies of the first appearance datum (FAD) concept

    NASA Astrophysics Data System (ADS)

    Landing, Ed; Geyer, Gerd; Brasier, Martin D.; Bowring, Samuel A.

    2013-08-01

    Use of the first appearance datum (FAD) of a fossil to define a global chronostratigraphic unit's base can lead to intractable correlation and stability problems. FADs are diachronous—they reflect species' evolutionary history, dispersal, biofacies, preservation, collection, and taxonomy. The Cambrian Evolutionary Radiation is characterised by diachronous FADs, biofacies controls, and provincialism of taxa and ecological communities that confound a stable Lower Cambrian chronostratigraphy. Cambrian series and stage definitions require greater attention to assemblage zone successions and non-biostratigraphic, particularly carbon isotope, correlation techniques such as those that define the Ediacaran System base. A redefined, basal Cambrian Trichophycus pedum Assemblage Zone lies above the highest Ediacaran-type biotas (vendobionts, putative metazoans, and calcareous problematica such as Cloudina) and the basal Asteridium tornatum-Comasphaeridium velvetum Zone (acritarchs). This definition and the likely close correspondence of evolutionary origin and local FAD of T. pedum preserves the Fortune Head, Newfoundland, GSSP of the Cambrian base and allows the presence of sub-Cambrian, branched ichnofossils. The sub-Tommotian-equivalent base of Stage 2 (a suggested "Laolinian Stage") should be defined by the I'/L4/ZHUCE δ13C positive peak, bracketed by the lower ranges of Watsonella crosbyi and Aldanella attleborensis (molluscs) and the Skiagia ornata-Fimbrioglomerella membranacea Zone (acritarchs). The W. crosbyi and A. attleborensis FADs cannot define a Stage 2 base as they are diachronous even in the Newfoundland "type" W. crosbyi Zone. The Series 2 base cannot be based on a species' FAD owing to the provincialism of skeletalised metazoans in the Terreneuvian-Series 2 boundary interval and global heterochrony of the oldest trilobites. A Series 2 and Stage 3 (a suggested "Lenaldanian Series" and "Zhurinskyan Stage," new) GSSP base is proposed at the Siberian lower

  7. Unusual seismogenic soft-sediment deformation structures in Cambrian epicratonic carbonate deposits, western Colorado, U.S.A

    NASA Astrophysics Data System (ADS)

    Myrow, P.; Chen, J.

    2013-12-01

    A wide variety of unusual penecontemporaneous deformation structures exist in grainstone and flat-pebble conglomerate beds of the Upper Cambrian strata, western Colorado, including slide scarps, thrusted beds, irregular blocks and internally deformed beds. Slide scarps are characterized by concave-up, sharp surfaces that truncate one or more underlying beds. Thrusted beds record movement of a part of a bed onto itself along a moderate to steeply inclined (generally 25°-40°) ramp. The hanging wall lenses in cases show fault-bend geometries, with either intact or mildly deformed bedding. Irregular bedded to internally deformed blocks isolated on generally flat upper bedding surfaces are similar in composition to the underlying beds. These features represent parts of beds that were detached, moved up onto, and some distances across, the laterally adjacent undisturbed bed surfaces. The blocks moved either at the sediment-water interface or intrastratally at shallow depths within overlying muddy deposits. Finally, internally deformed beds have large blocks, fitted fabrics of highly irregular fragments, and contorted lamination, which represent heterogeneous deformation, such as brecciation and liquefaction. The various deformation structures were most probably triggered by earthquakes, considering the nature of deformation (regional distribution of liquefaction structures, and the brittle segmentation and subsequent transportation of semi-consolidated beds) and the reactivation of Mesoproterozoic, crustal-scale shear zones in the central Rockies during the Late Cambrian. Features produced by initial brittle deformation are unusual relative to most reported seismites, and may represent poorly recognized to unrecognized seismogenic structures in the rock record.

  8. Small bilaterian fossils from 40 to 55 million years before the cambrian.

    PubMed

    Chen, Jun-Yuan; Bottjer, David J; Oliveri, Paola; Dornbos, Stephen Q; Gao, Feng; Ruffins, Seth; Chi, Huimei; Li, Chia-Wei; Davidson, Eric H

    2004-07-09

    Ten phosphatized specimens of a small (<180 micrometers) animal displaying clear bilaterian features have been recovered from the Doushantuo Formation, China, dating from 40 to 55 million years before the Cambrian. Seen in sections, this animal (Vernanimalcula guizhouena gen. et sp. nov.) had paired coeloms extending the length of the gut; paired external pits that could be sense organs; bilateral, anterior-posterior organization; a ventrally directed anterior mouth with thick walled pharynx; and a triploblastic structure. The structural complexity is that of an adult rather than a larval form. These fossils provide the first evidence confirming the phylogenetic inference that Bilateria arose well before the Cambrian.

  9. Sedimentary facies of the upper Cambrian (Furongian; Jiangshanian and Sunwaptan) Tunnel City Group, Upper Mississippi Valley: new insight on the old stormy debate

    USGS Publications Warehouse

    Eoff, Jennifer D.

    2014-01-01

    New data from detailed measured sections permit a comprehensive revision of the sedimentary facies of the Furongian (upper Cambrian; Jiangshanian and Sunwaptan stages) Tunnel City Group (Lone Rock Formation and Mazomanie Formation) of Wisconsin and Minnesota. Heterogeneous sandstones, comprising seven lithofacies along a depositional transect from shoreface to transitional-offshore environments, record sedimentation in a storm-dominated, shallow-marine epicontinental sea. The origin of glauconite in the Birkmose Member and Reno Member of the Lone Rock Formation was unclear, but its formation and preserved distribution are linked to inferred depositional energy rather than just net sedimentation rate. Flat-pebble conglomerate, abundant in lower Paleozoic strata, was associated with the formation of a condensed section during cratonic flooding. Hummocky cross-stratification was a valuable tool used to infer depositional settings and relative paleobathymetry, and the model describing formation of this bedform is expanded to address flow types dominant during its genesis, in particular the importance of an early unidirectional component of combined flow. The depositional model developed here for the Lone Rock Formation and Mazomanie Formation is broadly applicable to other strata common to the early Paleozoic that document sedimentation along flooded cratonic interiors or shallow shelves.

  10. Sulfur and Carbon Isotope Systematics in Middle-Upper Cambrian Port au Port Group From Western Newfoundland, Canada: Implications for Seawater Sulfate Concentrations

    NASA Astrophysics Data System (ADS)

    Hurtgen, M. T.; Pruss, S.; Knoll, A. H.

    2006-12-01

    The biogeochemical cycles of carbon and sulfur are intimately linked through a variety of feedbacks that operate on timescales of days to millions of years. For example, under anaerobic conditions, some bacteria respire organic matter by sulfate reduction, reducing sulfate to sulfide, which then reacts with iron to form iron sulfide (preserved as pyrite). On much longer timescales, increases in the fraction of total carbon buried as organic carbon can drive increases in atmospheric oxygen concentrations which then facilitate an increase in the extent to which sulfides on land are oxidatively weathered and ultimately delivered to the oceans as sulfate via rivers. Interestingly, these two processes impose very different isotope relationships between the C isotope composition of marine dissolved inorganic carbon (DIC) and the S isotope composition of seawater sulfate. The former leads to a positive correlation between δ13Ccarbonate and δ34Ssulfate whereas the latter prescribes a long-term negative correlation. Of course, the recognition of either a positive or negative correlation between δ13Ccarbonate and δ34Ssulfate depends strongly on the relative sizes of the DIC and seawater sulfate reservoirs-- neither of which is well constrained for the Cambrian Period. Here, we present a high-resolution δ34S (sulfate and pyrite) and δ13Ccarbonate record from the mixed carbonate-siliciclastic Middle-Upper Cambrian Port au Port Group in western Newfoundland, Canada. The δ34Ssulfate profile displays systematic shifts of >15‰ over relatively short stratigraphic distances (10 m, likely to represent < 1 Myr). C isotope values shift sympathetically throughout much of the composite section; however, important deviations from this relationship exist. First, in the Middle Cambrian March Point Formation, a 15‰- δ34Ssulfate decrease precedes a 3‰-δ13Ccarbonate fall suggesting that the sulfur cycle recorded the perturbation to the system before the carbon cycle did

  11. Cambro-ordovician sea-level fluctuations and sequence boundaries: The missing record and the evolution of new taxa

    USGS Publications Warehouse

    Lehnert, O.; Miller, J.F.; Leslie, Stephen A.; Repetski, J.E.; Ethington, Raymond L.

    2005-01-01

    The evolution of early Palaeozoic conodont faunas shows a clear connection to sea-level changes. One way that this connection manifests itself is that thick successions of carbonates are missing beneath major sequence boundaries due to karstification and erosion. From this observation arises the question of how many taxa have been lost from different conodont lineages in these incomplete successions. Although many taxa suffered extinction due to the environmental stresses associated with falling sea-levels, some must have survived in these extreme conditions. The number of taxa missing in the early Palaeozoic tropics always will be unclear, but it will be even more difficult to evaluate the missing record in detrital successions of higher latitudes. A common pattern in the evolution of Cambrian-Ordovician conodont lineages is appearances of new species at sea-level rises and disappearances at sea-level drops. This simple picture can be complicated by intervals that consistently have no representatives of a particular lineage, even after extensive sampling of the most complete sections. Presumably the lineages survived in undocumented refugia. In this paper, we give examples of evolution in Cambrian-Ordovician shallowmarine conodont faunas and highlight problems of undiscovered or truly missing segments of lineages. ?? The Palaeontological Association.

  12. Unraveling the redox evolution of the Yangtze Block across the Precambrian/Cambrian transition

    NASA Astrophysics Data System (ADS)

    Diamond, C. W.; Zhang, F.; Chen, Y.; Lyons, T. W.

    2016-12-01

    Rocks preserved on the South China Craton have played a critical role in refining our understanding of the co-evolution of life and Earth's surface environments in the Late Neoproterozoic and earliest Paleozoic. From the earliest metazoan embryos to the many examples of exceptional preservation throughout the Cambrian Explosion, South China has preserved an outstanding record of animal evolution across this critical transition. Similarly, rocks preserved in South China hold key insights into the changing ocean chemistry that accompanied this extraordinary time. Recent work form Sahoo and others (2016, Geobiology) used redox sensitive metal enrichments in the Ediacaran Doushantuo Formation to demonstrate that the redox state of the Latest Neoproterozoic oceans was highly dynamic, rather than stably oxygenated or anoxic as had both been suggested previously. In an attempt to follow on from this and other studies, we have examined samples from a drill core taken in eastern Guizhou capturing deep-water facies of the Liuchapo and Jiumenchong formations, which contain the Precambrian/Cambrian boundary. In addition to containing the boundary, the sampled interval contains an enigmatic, widespread horizon that is strongly enriched in Ni and Mo. We have taken a multi-proxy approach in our investigation of this layer, the possible implications it has for the strata above and below (i.e., how its presence affects their utility as archives of paleo-redox conditions), and what those strata can tell us about local and global redox conditions during this pivotal time in Earth's history. Our Fe speciation data indicate that conditions were sulfidic at this location throughout the majority of the sampled interval. While redox sensitive metal concentrations are dramatically enriched in the Ni/Mo interval, their concentrations return to modest enrichments above it and continue to decrease upward. This trend suggests that while the conditions that favored extreme enrichment during the

  13. Cambrian Series 3 carbonate platform of Korea dominated by microbial-sponge reefs

    NASA Astrophysics Data System (ADS)

    Hong, Jongsun; Lee, Jeong-Hyun; Choh, Suk-Joo; Lee, Dong-Jin

    2016-07-01

    Metazoans have been considered as negligible components of Cambrian Series 3 and Furongian microbial-dominated reefs, in contrast to their presence in earlier Terreneuvian-Cambrian Series 2 microbial-archaeocyath reefs. However, recent discoveries of sponges in Cambrian Series 3-Furongian reefs of Australia, China, Iran, USA, and Korea have raised question regarding their contribution in terms of carbonate platform development, which have never been assessed. This study examines Cambrian Series 3 deposits of the Daegi Formation, Korea to elucidate this question. The 100-m-thick middle part of the Daegi Formation is dominated by boundstone facies, which occupies 45% of the study interval, as well as bioclastic wackestone to packstone, bioclastic grainstone, and ooid packstone to grainstone facies. The Daegi reefs are primarily thrombolitic in composition, with 90% (n = 26/29) of the reefs containing an average of 9% sponges in aerial percentage calculated from thin sections. Lithistid sponges composed of peloidal fabrics, some desma spicules, and spicule networks commonly occupy the interstitial space in microbial clusters, are encrusted by mesoclots and Epiphyton, and are surrounded by micrite. Subordinate non-lithistid demosponges occur within clusters of microbial elements. The middle Daegi Formation can be largely subdivided into shoal environment dominated by grainstone to packstone facies and shallow subtidal platform interior environment located behind shoal with wackestone to packstone facies. The microbial-sponge reefs mainly developed around platform interior as patch reefs. The current study indicates that metazoans in the form of lithistid and non-lithistid demosponges are nearly ubiquitously incorporated in Daegi reefs and contributed greatly to the formation of microbial-sponge reefs as well as carbonate platform during the time. Study of these microbial-sponge reefs and their distribution within the carbonate platform may help us to understand how

  14. Multiple High-Frequency Carbon Isotope Excursions Across the Precambrian-Cambrian Boundary: Implications for Correlations and Environmental Change

    NASA Astrophysics Data System (ADS)

    Smith, E. F.; Macdonald, F. A.; Schrag, D. P.; Laakso, T.

    2014-12-01

    The GSSP Precambrian-Cambrian boundary in Newfoundland is defined by the first appearance datum (FAD) of Treptichnus pedum, which is considered to be roughly coincident with the FAD of small shelly fossils (SSFs) and a large negative carbon isotope excursion. An association between the FAD of T. pedum and a negative carbon isotope excursion has previously been documented in Northwest Canada (Narbonne et al., 1994) and Death Valley (Corsetti and Hagadorn, 2000), and since then has been used as an chronostratigraphic marker of the boundary, particularly in siliciclastic poor sections that do not preserve T. pedum. Here we present new high-resolution carbon isotope (δ13C ) chemostratigraphy from multiple sections in western Mongolia and the western United States that span the Ediacaran-Cambrian transition. High-resolution sampling (0.2-1 m) reveals that instead of one large negative excursion, there are multiple, high-frequency negative excursions with an overall negative trend during the latest Ediacaran. These data help to more precisely calibrate changes in the carbon cycle across the boundary as well as to highlight the potential problem of identifying the boundary with just a few negative δ13C values. We then use a simple carbon isotope box model to explore relationships between phosphorous delivery to the ocean, oxygenation, alkalinity, and turnovers in carbonate secreting organisms. Corsetti, F.A., and Hagadorn, J.W., 2000, Precambrian-Cambrian transition: Death Valley, United States: Geology, v. 28, no. 4, p. 299-302. Narbonne, G.M., Kaufman, A.J., and Knoll, A.H., 1994, Integrated chemostratigraphy and biostratigraphy of the Windermere Supergroup, northwestern Canada: Implications for Neoproterozoic correlations and the early evolution of animals: Geological Society of America Bulletin, v. 106, no. 10, p. 1281-1292.

  15. Paleomagnetism of a well-dated marine succession in South China: A possible Late Cambrian true polar wander (TPW)

    NASA Astrophysics Data System (ADS)

    Jiao, Wen-Jun; Li, Yong-Xiang; Yang, Zhen-Yu

    2018-04-01

    The Cambrian true polar wander (TPW) hypothesis remains controversial largely because of the uncertainties in the quality and/or fidelity of the paleomagnetic data as well as their chronological control. Testing the TPW hypothesis requires high-quality paleomagnetic data of sufficient spatial and temporal resolutions. Here, we present paleomagnetic results of a continuous Cambrian shallow marine succession from South China where available detailed biostratigraphy provides exceptional chronological constraints. Forty-three sites of paleomagnetic samples were collected from this limestone-dominated succession. Stepwise thermal demagnetization generally reveals three-component magnetizations. Low- and intermediate-temperature components can be cleaned by ∼330 °C, and the high-temperature component (HTC) was isolated typically from ∼350 to ∼450 °C. A positive fold test and the presence of reversed polarity in the strata, together with rock magnetic data as well as the scanning electron microscopic (SEM) and transmission electron microscopic (TEM) results, collectively suggest that the HTCs are likely primary. A directional shift of the HTCs occurs between the lower-middle Cambrian and the upper Cambrian strata in the succession and is tentatively interpreted to indicate a ∼57° polar wander from ∼500.5 to 494 Ma. Because the rate of polar wander is too fast to be a tectonic origin, this polar wander is interpreted to represent a Late Cambrian TPW. This TPW appears coeval with the Steptoean positive carbon isotope excursion (SPICE) and the major trilobite mass extinctions, suggesting a potential link between the TPW and the Late Cambrian biotic and climatic changes. Because the proposed TPW event is exceptionally well-dated, it should be testable through examination of other worldwide sections.

  16. Palaeomagnetism and geochemistry of Early Palaeozoic rocks of the Barrandian (Teplé-Barrandian Unit, Bohemian Massif): palaeotectonic implications

    NASA Astrophysics Data System (ADS)

    Patočka, F.; Pruner, P.; Štorch, P.

    The Barrandian area (the Teplá-Barrandian unit, Bohemian Massif) provided palaeomagnetic results on Early Palaeozoic rocks and chemical data on siliciclastic sediments of both Middle Cambrian and Early Ordovician to Middle Devonian sedimentary sequences; an outcoming interpretation defined source areas of clastic material and palaeotectonic settings of the siliciclastic rock deposition. The siliciclastic rocks of the earliest Palaeozoic sedimentation cycle, deposited in the Cambrian Příbram-Jince Basin of the Barrandian, were derived from an early Cadomian volcanic island arc developed on Neoproterozoic oceanic lithosphere and accreted to a Cadomian active margin of northwestern Gondwana. Inversion of relief terminated the Cambrian sedimentation, and a successory Prague Basin subsided nearby since Tremadocian. Source area of the Ordovician and Early Silurian shallow-marine siliciclastic sediments corresponded to progressively dissected crust of continental arc/active continental margin type of Cadomian age. Since Late Ordovician onwards both synsedimentary within-plate basic volcanics and older sediments had been contributing in recognizable proportions to the siliciclastic rocks. The siliciclastic sedimentation was replaced by deposition of carbonate rocks throughout late Early Silurian to Early Devonian period of withdrawal of the Cadomian clastic material source. Above the carbonates an early Givetian flysch-like siliciclastic suite completed sedimentation in the Barrandian. In times between Middle Cambrian and Early/Middle Devonian boundary interval an extensional tectonic setting prevailed in the Teplá-Barrandian unit. The extensional regime was related to Early Palaeozoic large-scale fragmentation of the Cadomian belt of northwestern Gondwana and origin of Armorican microcontinent assemblage. The Teplá-Barrandian unit was also engaged in a peri-equatorially oriented drift of Armorican microcontinent assemblage throughout the Early Palaeozoic: respective

  17. A new U-Pb LA-ICP-MS age of the Rumburk granite (Lausitz Block, Saxo-Thuringian Zone): constraints for a magmatic event in the Upper Cambrian

    NASA Astrophysics Data System (ADS)

    Zieger, J.; Linnemann, U.; Hofmann, M.; Gärtner, A.; Marko, L.; Gerdes, A.

    2018-04-01

    The basement of the Saxo-Thuringian Zone consists of Upper Neoproterozoic (c. 650-570 Ma) Cadomian arc sediments (Lusatian greywackes) and voluminous intrusions of Early Cambrian granitoids with ages of c. 540 Ma (Lausitz Block and Karkonosze-Izera Massif). The latter basement complexes comprise several c. 505 Ma granites, granodiorites, and gneisses emplaced during the change from a collisional tectonic setting to rift-related geotectonics. We present a new age for the Rumburk granite of 504 ± 3 Ma linking Late Cambrian plutonism at the northern margin of Gondwana with the initial phase of a Cambro-Ordovician rift event. Trace element analysis points to a linkage of the Rumburk granite with other Late Cambrian aged rocks of the Karkonosze-Izera Massif. Furthermore, geochemical data also provide evidence of a melting and recycling of Lusatian greywackes by the intrusion of the Rumburk granite. The youngest age peak of the Rumburk granite at c. 504 Ma is considered to be the age of emplacement. Older inherited age populations at c. 540 and c. 610 Ma are present and likely the result of a melting and recycling of Lusatian granitoids and greywackes. The appearance of Neoproterozoic inheritance and Lu-Hf similarities with the Rumburk granite strongly suggest the Lusatian greywackes as source rocks. There is a significant age gap of c. 35 Ma between Cambrian plutonic and volcanic rocks in Saxo-Thuringia. Hence, we consider two distinct pulses of magmatic activity during the transition from the Cadomian orogeny to the opening of the Rheic Ocean.

  18. Oxygenation history of the Neoproterozoic to early Phanerozoic and the rise of land plants

    NASA Astrophysics Data System (ADS)

    Wallace, Malcolm W.; Hood, Ashleigh vS.; Shuster, Alice; Greig, Alan; Planavsky, Noah J.; Reed, Christopher P.

    2017-05-01

    There has been extensive debate about the history of Earth's oxygenation and the role that land plant evolution played in shaping Earth's ocean-atmosphere system. Here we use the rare earth element patterns in marine carbonates to monitor the structure of the marine redox landscape through the rise and diversification of animals and early land plants. In particular, we use the relative abundance of cerium (Ceanom), the only redox-sensitive rare earth element, in well-preserved marine cements and other marine precipitates to track seawater oxygen levels. Our results indicate that there was only a moderate increase in oceanic oxygenation during the Ediacaran (average Cryogenian Ceanom = 1.1, average Ediacaran Ceanom = 0.62), followed by a decrease in oxygen levels during the early Cambrian (average Cryogenian Ceanom = 0.90), with significant ocean anoxia persisting through the early and mid Paleozoic (average Early Cambrian-Early Devonian Ceanom = 0.84). It was not until the Late Devonian that oxygenation levels are comparable to the modern (average of all post-middle Devonian Ceanom = 0.55). Therefore, this work confirms growing evidence that the oxygenation of the Earth was neither unidirectional nor a simple two-stage process. Further, we provide evidence that it was not until the Late Devonian, when large land plants and forests first evolved, that oxygen levels reached those comparable to the modern world. This is recorded with the first modern-like negative Ceanom (values <0.6) occurring at around 380 Ma (Frasnian). This suggests that land plants, rather than animals, are the 'engineers' responsible for the modern fully oxygenated Earth system.

  19. Development, triploblastism, physics of wetting and the Cambrian explosion.

    PubMed

    Fleury, Vincent

    2013-09-01

    The Cambrian explosion is characterized by the sudden outburst of organized animal plans, which occurred circa 530 M years ago. Around that time, many forms of animal life appeared, including several which have since disappeared. There is no general consensus about "why" this happened, and why it had any form of suddenness. However, all organized animal plans share a common feature: they are triploblastic, i.e., composed of 3 layers of tissue, endoderm, ectoderm and mesoderm. I show here that, within simple hypotheses, the formation of the mesoderm has intrinsically a physical exponential dynamics, leading rapidly to triploblastism, and eventually, to animal formation. A novel physico-mathematical framework including epithelium-mesenchyme transition, visco-elastic constitutive equations, and conservation laws, is presented which allows one to describe gastrulation as a self-wetting phenomenon of a soft solid onto itself. This phenomenon couples differentiation and migration during gastrulation, and leads in a closed form to an exponential scaling law for the formation of the mesoderm. Therefore, the Cambrian explosion might have started, actually, by a true viscoelastic "explosion": the exponential run-away of mesenchymal cells.

  20. A molecular and isotopic study of palaeoenvironmental conditions through the middle Cambrian in the Georgina Basin, central Australia

    NASA Astrophysics Data System (ADS)

    Pagès, Anais; Schmid, Susanne; Edwards, Dianne; Barnes, Stephen; He, Nannan; Grice, Kliti

    2016-08-01

    The Cambrian period marks an important point in Earth's history with profound changes in the ocean's biogeochemistry and the occurrence of the most significant evolutionary event in the history of life, the Cambrian explosion. The Cambrian explosion is described as a succession of complex cycles of extinctions and radiations. This study integrates biomarkers and their compound-specific stable carbon isotopes to investigate the palaeoenvironmental depositional conditions in middle Cambrian (Series 3) sedimentary rocks (Thorntonia Limestone, Inca Formation and Currant Bush Limestone) from two drillholes in the Undilla Sub-basin in the eastern Georgina Basin, central Australia. The occurrence of photic zone euxinia (PZE) was detected throughout these three formations by the identification of green sulfur bacteria Chlorobiaceae-derived biomarkers, including a series of 2,3,6-aryl isoprenoids and the intact biomarker isorenieratane. Pulses of enhanced PZE conditions were detected in two core intervals (90-110 mKB, Currant Bush Limestone and 170-200 mKB, Inca Formation) by an increase in the 2,3,6-aryl isoprenoids and C19 biphenyl concentrations. These enhanced PZE conditions were followed by blooms of phytoplankton, as demonstrated by the increase in algal-derived biomarker (i.e. pristane, phytane and the C19n-alkane) concentrations and compound-specific isotopes. These observations confirm that palaeoenvironmental conditions were similar to those reported for the Permian/Triassic and Triassic/Jurassic mass extinction events. The sterane distributions varied across the three formations reflecting possible changes in the phytoplanktonic communities through time. Although a rise in atmospheric oxygen during the Cambrian has been previously associated with the rapid evolution of metazoans, the ecological challenges related to widespread anoxia must have had a major influence on the evolution of life in Cambrian oceans.

  1. Selenium and hazardous elements distribution in plant-soil-water system and human health risk assessment of Lower Cambrian, Southern Shaanxi, China.

    PubMed

    Du, Yajun; Luo, Kunli; Ni, Runxiang; Hussain, Rahib

    2018-03-01

    The natural selenium poisoning due to toxic Se levels in food chain had been observed in humans and animals in Lower Cambrian outcrop areas in Southern Shaanxi, China. To find out the distribution pattern of selenium and other hazardous elements in the plant, soil and water of Lower Cambrian in Southern Shaanxi, China, and their possible potential health risk, a total of 30 elements were analyzed and the health risk assessment of 18 elements was calculated. Results showed that the soil, plant and natural water of Lower Cambrian all had relatively high Se levels. In Lower Cambrian, the soil was enriched with Se, As, Ba, Cu, Mo, Ni, Zn, Ga, Cd and Cr (1.68 < I geo  < 4.48, I geo ; geo-accumulation index). In same plants, the contents of Se, Cd and Zn (except Cd in corn and rice, Zn in potato and corn) of Lower Cambrian were higher than that of the other strata. Ba and Ga in natural water were higher than that of the other strata, while K and Cs were opposite. The health risk assessment results showed that the people living in outcrop areas of Lower Cambrian had both high total non-carcinogenic risk of 18 elements (HI = 16.12, acceptable range: < 1) and carcinogenic risk of As (3.98E-04, acceptable range: 10 -6 -10 -4 ). High contents of Se, As, Mo and Tl of Lower Cambrian may pose a health risk to local people, and food intake was the major pathway. For minimizing potential health risk, the local inhabitants should use the mix-imported food with local growing foods.

  2. Palaeoenvironmental and geochemical approach of Archaeocyath-rich facies from Lower Cambrian of Western Gondwana margin at Central Iberian Zone (Urda, Toledo Mountains, Spain)

    NASA Astrophysics Data System (ADS)

    Menéndez, Silvia; Rodríguez-Martínez, Marta; Moreno-Eris, Elena; Perejón, Antonio; Reitner, Joachim

    2010-05-01

    Archaeocyath-rich facies are located in a quarry close to Urda village, at Toledo Mountains, Spain. The outcrops belong to the Caliza de los Navalucillos Formation and they record a considerably high diverse archaeocyath assemblage in the Lower Cambrian successions from the Central Iberian Zone (Julivert et al. 1972 [1974]). In fact, it is first time recorded the presence of Agyrekocyathus, Dokidocyathus, and Plicocyathus in the Central Iberian Zone. Therefore Plicocyathus is no longer exclusive to biozone VI in Spain. The presence of Anthomorpha is characteristic for the early Botomian, presently early Stage 4 (ICS, 2009), and the assemblage corresponds to the biozone VII (late Ovetian, following the biozonation of Perejón & Moreno-Eiris, 2006). The fossiliferous part of the succession is formed by seven lithofacies, all of them tectonically folded and with a low grade metamorphic overprint. They are comprised by two main groups of facies: (a) mound-shaped to massive lithofacies (A1, A2, A3, A4) and (b) massive to bedded and nodular lithofacies (B1, B2, B3). Archaeocyaths occur in several facies: (A1) mound-shaped white marble with irregular to stromatactoid cavities; (A2) massive mottled white to grey limestone; (A3) massive grey limestone with slumps levels; (A4) massive archaeocyath-rich orange limestone; as well as in carbonate nodules embedded in siltstones and cherts (B1, B2 and B3). The best preserved assemblage comes from the nodule record, where fossils are partially pyritized. This type of preservation is exceptional and has never been described before. XRD and wavelength-dispersive electron microprobe analyses reveal the presence of pyrite and pyrrotine partially altered to iron oxides and hydroxides (hematite and goethite) surrounding the archaeocyath cups. In Central Iberian Zone, the development of mounds and nodular facies like those described here is unusual, although the Botomian marks the peak for Early Cambrian archaeocyathan-microbial mounds

  3. EBSD characterization of pre-Cambrian deformations in conglomerate pebbles (Sierra de la Demanda, Northern Spain)

    NASA Astrophysics Data System (ADS)

    Puelles, Pablo; Ábalos, Benito; Fernández-Armas, Sergio

    2010-05-01

    Pre-Cambrian and unconformable earliest Cambrian rocks from the Sierra de la Demanda (N Spain) exhibit field and microstructural relationships that attest to orogenic events recorded by concealed basement rocks. Neoproterozoic foliated slates ("Anguiano Schists") crop out under up to 300 m thick, unfoliated quartz-rich conglomerates ("Anguiano Conglomerates") and quartzites which are stratigraphically ca. 600 m below the oldest, paleontologically dated, pre-trilobitic Cambrian layers (likely older than 520 Ma). The Anguiano Conglomerates contain mm to cm grainsized well-rounded pebbles of various types including monocrystalline quartz, detrital zircon and tourmaline-bearing sandstones, black cherts and metamorphic poly-crystalline quartz aggregates. The undeformed matrix is made of much smaller (diagenetically overgrown) monocrystaline quartz grains and minor amounts of accesory zircon, tourmaline and mica. Black chert pebbles exhibit microstructural evidence of brittle deformation (microfaults and thin veins of syntaxial fibrous quartz). These and the fine-grained sandstone pebbles can also exhibit ductile deformations (microfolds with thickened hinges and axial planar continuous foliations), too. Polycrystalline quartz pebbles exhibit a variety of microstructures that resulted from syn-metamorphic ductile deformations. These are recognisable under the petrographic microscope and include continuous foliations, quartz shape fabrics, various types of subgrain or recrystallized new grain microtextures, and lattice preferred orientations (LPOs). Conventional characterization of quartz fabrics (after oriented structural sections) is challenged in conglomerate pebble thin sections by the difficulty of unraveling in them the complete structural reference framework provided by foliation (whose trace can be unraveled) and lineation orientation (which cannot be directly identified). Quartz in various metamorphic polycrystalline pebbles was studied with the Electron Back

  4. New paleomagnetic data from Siberia: Non-uniformitarian geomagnetic field around the Proterozoic-Phanerozoic boundary?

    NASA Astrophysics Data System (ADS)

    Pavlov, V.; Shatsillo, A.; Kouznetsov, N.; Gazieva, E.

    2017-12-01

    There is a range of evidence, mainly from sedimentary and volcanic rocks of the Laurentia and Baltica cratons, that argue for the anomalous character of the Ediacaran-Early Cambrian paleomagnetic record. This feature could be linked either to some peculiarities of the paleomagnetic record itself or to some unusual geophysical event that would have taken place around the Proterozoic-Phanerozoic boundary (e.g., true polar wander or nonuniformitarian geomagnetic field behavior). In the latter case, the traces of this event should be observed in Ediacaran-Early Cambrian rocks anywhere there is a possibility to observe a primary paleomagnetic signal. In previous work, we reported results that suggested an anomalous paleomagnetic record in Siberian Ediacaran-Lower Cambrian rocks. Here we present new Siberian data that indicate a very high geomagnetic reversal frequency during this period and the coexistence of two very different paleomagnetic directions. We speculate that these features could be due either to a near-equatorial geomagnetic dipole during the polarity transitions or to alternation between axial and near equatorial dipoles not directly linked with polarity reversals.

  5. Paragenesis of the Morgan Creek Limestone, Late Cambrian, central Texas: Constraints on the formation of glauconite

    NASA Astrophysics Data System (ADS)

    Chafetz, Henry S.

    2007-06-01

    Deposition of the Morgan Creek Limestone, a member of the Cambrian Wilberns Formation, began a few meters above a disconformable surface and displays abundant indicators of accumulation in shallow marine to tidal flat environments. These indicators include: intercalation of a lithologic variety of thin beds (e.g. biosparites, biomicrites, oosparites, intrasparites), which display rapid vertical and lateral lithologic changes, numerous stromatolitic horizons, channels filled with graded oosparites and intrasparites that cut through micrite accumulations, and finely laminated (non-burrowed) siltstones. Glauconite is a ubiquitous minor constituent throughout in the form of pellets, replaced skeletal material and mica books, and, most informatively, as an authigenic precipitate in the form of fibroradiating rims on carbonate allochems and siliciclastic grains. Fibroradiating glauconite rims were disrupted and pushed away from the pelmatozoan nuclei on which they precipitated. Combined sedimentological and paragenetic constraints indicate that the glauconite was the earliest diagenetic event to affect these sediments and occurred essentially at the sediment-water interface within these relatively high-energy, shallow marine deposits. Precipitation of glauconite was closely followed by precipitation of carbonate cement as well as dissolution of aragonitic constituents. Later diagenetic processes included selective dolomitization and silicification. In modern seas glauconite is reported to form at and below mid-shelf water depths. Thus, the constraints pertaining to the origin of modern glauconite are not valid for Late Cambrian deposits and probably are also not applicable for Late Cretaceous through Early Paleogene glauconites.

  6. A Vendian-Cambrian boundary succession from the northwestern margin of the Siberian Platform: stratigraphy, palaeontology, chemostratigraphy and correlation

    NASA Technical Reports Server (NTRS)

    Bartley, J. K.; Pope, M.; Knoll, A. H.; Semikhatov, M. A.; Grotzinger, J. (Principal Investigator)

    1998-01-01

    Siberia contains several key reference sections for studies of biological and environmental evolution across the Proterozoic-Phanerozoic transition. The Platonovskaya Formation, exposed in the Turukhansk region of western Siberia, is an uppermost Proterozoic to Cambrian succession whose trace and body fossils place broad limits on the age of deposition, but do not permit detailed correlation with boundary successions elsewhere. In contrast, a striking negative carbon isotopic excursion in the lower part of the Platonovskaya Formation permits precise chemostratigraphic correlation with upper-most Yudomian successions in Siberia, and possibly worldwide. In addition to providing a tool for correlation, the isotopic excursion preserved in the Platonovskaya and contemporaneous successions documents a major biogeochemical event, likely involving the world ocean. The excursion coincides with the palaeontological breakpoint between Ediacaran- and Cambrian-style assemblages, suggesting a role for biogeochemical change in evolutionary events near the Proterozoic Cambrian boundary.

  7. A Vendian-Cambrian boundary succession from the northwestern margin of the Siberian Platform: stratigraphy, palaeontology, chemostratigraphy and correlation.

    PubMed

    Bartley, J K; Pope, M; Knoll, A H; Semikhatov, M A; Petrov PYu

    1998-07-01

    Siberia contains several key reference sections for studies of biological and environmental evolution across the Proterozoic-Phanerozoic transition. The Platonovskaya Formation, exposed in the Turukhansk region of western Siberia, is an uppermost Proterozoic to Cambrian succession whose trace and body fossils place broad limits on the age of deposition, but do not permit detailed correlation with boundary successions elsewhere. In contrast, a striking negative carbon isotopic excursion in the lower part of the Platonovskaya Formation permits precise chemostratigraphic correlation with upper-most Yudomian successions in Siberia, and possibly worldwide. In addition to providing a tool for correlation, the isotopic excursion preserved in the Platonovskaya and contemporaneous successions documents a major biogeochemical event, likely involving the world ocean. The excursion coincides with the palaeontological breakpoint between Ediacaran- and Cambrian-style assemblages, suggesting a role for biogeochemical change in evolutionary events near the Proterozoic Cambrian boundary.

  8. Magnetic reversal frequency in the Lower Cambrian Niutitang Formation, Hunan Province, South China

    NASA Astrophysics Data System (ADS)

    Duan, Zongqi; Liu, Qingsong; Ren, Shoumai; Li, Lihui; Deng, Xiaolong; Liu, Jianxing

    2018-05-01

    The reversal frequency of the paleomagnetic field bears great information of evolution of the Earth's deep interior. However, there are still debates on the frequency pattern during the older periods of the Phanerozoic. This study investigated the Niutitang Formation (Lower Cambrian) of the Ciye 1 Hole from south China. Rock magnetic results indicate that the dominant magnetic carrier is magnetite. Characteristic remanence magnetizations have been successfully isolated for the weakly-magnetized shale rocks through stepwise alternated field demagnetization using the 2 G Enterprises Rapid System Magnetometer with a low-noise thin-walled quartz-glass sample holder. Constrained by radiometric ages, our paleomagnetic results indicated frequent polarity reversals during the period of ˜524-514 Ma, which backs up the speculation about the episode of the Ediacaran-Cambrian (˜550-500 Ma) with a character of reversal hyperactivity.

  9. Groundwater quality in the Cambrian-Ordovician aquifer system, midwestern United States

    USGS Publications Warehouse

    Stackelberg, Paul E.

    2017-12-07

    Groundwater provides nearly 50 percent of the Nation’s drinking water. To help protect this vital resource, the U.S. Geological Survey (USGS) National Water-Quality Assessment (NAWQA) Project assesses groundwater quality in aquifers that are important sources of drinking water (Burow and Belitz, 2014). The Cambrian-Ordovician aquifer system constitutes one of the important areas being evaluated.

  10. Isotopically zoned carbonate cements in Early Paleozoic sandstones of the Illinois Basin: δ18O and δ13C records of burial and fluid flow

    NASA Astrophysics Data System (ADS)

    Denny, Adam C.; Kozdon, Reinhard; Kitajima, Kouki; Valley, John W.

    2017-11-01

    SEM/SIMS imaging and analysis of δ18O and δ13C in sandstones from a transect through the Illinois Basin (USA) show systematic μm-scale isotopic zonation of up to 10‰ in both carbonate and quartz cements of the middle-Ordovician St. Peter and Cambrian Mt. Simon formations. Quartz δ18O values are broadly consistent with the model of Hyodo et al. (2014), wherein burial and heating in the Illinois Basin is recorded in systematically zoned quartz overgrowths. Observations of zoned dolomite/ankerite cements indicate that they preserve a more extended record of temperature and fluid compositions than quartz, including early diagenesis before or during shallow burial, and late carbonates formed after quartz overgrowths. Many carbonate cements show innermost dolomite with δ18O values (21-25‰ VSMOW) that are too low to have formed by deposition at low temperatures from ancient seawater (δ18O > - 3‰) and most likely reflect mixing with meteoric water. A sharp increase in Fe content is commonly observed in zoned carbonate cements to be associated with a drop in δ18O and an abrupt shift in δ13C to higher or lower values. These changes are interpreted to record the passage of hot metal-rich brines through sandstone aquifers, that was associated with Mississippi-Valley Type (MVT) Pb-Zn deposits (ca. 270 Ma) of the Upper Mississippi Valley. Local variability and individual trends in δ13C are likely controlled by the sources of carbon and the degree to which carbon is sourced from adjacent carbonate units or thermal maturation of organic matter. Quartz overgrowths in sandstones provide an excellent record of conditions during burial, heating, and pressure-solution, whereas carbonate cements in sandstones preserve a more-extended record including initial pre-burial conditions and punctuated fluid flow events.

  11. Paleoarchean and Cambrian observations of the geodynamo in light of new estimates of core thermal conductivity

    NASA Astrophysics Data System (ADS)

    Tarduno, John; Bono, Richard; Cottrell, Rory

    2015-04-01

    Recent estimates of core thermal conductivity are larger than prior values by a factor of approximately three. These new estimates suggest that the inner core is a relatively young feature, perhaps as young as 500 million years old, and that the core-mantle heat flux required to drive the early dynamo was greater than previously assumed (Nimmo, 2015). Here, we focus on paleomagnetic studies of two key time intervals important for understanding core evolution in light of the revisions of core conductivity values. 1. Hadean to Paleoarchean (4.4-3.4 Ga). Single silicate crystal paleointensity analyses suggest a relatively strong magnetic field at 3.4-3.45 Ga (Tarduno et al., 2010). Paleointenity data from zircons of the Jack Hills (Western Australia) further suggest the presence of a geodynamo between 3.5 and 3.6 Ga (Tarduno and Cottrell, 2014). We will discuss our efforts to test for the absence/presence of the geodynamo in older Eoarchean and Hadean times. 2. Ediacaran to Early Cambrian (~635-530 Ma). Disparate directions seen in some paleomagnetic studies from this time interval have been interpreted as recording inertial interchange true polar wander (IITPW). Recent single silicate paleomagnetic analyses fail to find evidence for IITPW; instead a reversing field overprinted by secondary magnetizations is defined (Bono and Tarduno, 2015). Preliminary analyses suggest the field may have been unusually weak. We will discuss our on-going tests of the hypothesis that this interval represents the time of onset of inner core growth. References: Bono, R.K. & Tarduno, J.A., Geology, in press (2015); Nimmo, F., Treatise Geophys., in press (2015); Tarduno, J.A., et al., Science (2010); Tarduno, J.A. & Cottrell, R.D., AGU Fall Meeting (2014).

  12. A New Stalked Filter-Feeder from the Middle Cambrian Burgess Shale, British Columbia, Canada

    PubMed Central

    O'Brien, Lorna J.; Caron, Jean-Bernard

    2012-01-01

    Burgess Shale-type deposits provide invaluable insights into the early evolution of body plans and the ecological structure of Cambrian communities, but a number of species, continue to defy phylogenetic interpretations. Here we extend this list to include a new soft-bodied animal, Siphusauctum gregarium n. gen. and n. sp., from the Tulip Beds (Campsite Cliff Shale Member, Burgess Shale Formation) of Mount Stephen (Yoho National Park, British Columbia). With 1,133 specimens collected, S. gregarium is clearly the most abundant animal from this locality. This stalked animal (reaching at least 20 cm in length), has a large ovoid calyx connected to a narrow bilayered stem and a small flattened or bulb-like holdfast. The calyx is enclosed by a flexible sheath with six small openings at the base, and a central terminal anus near the top encircled by indistinct openings. A prominent organ, represented by six radially symmetrical segments with comb-like elements, surrounds an internal body cavity with a large stomach, conical median gut and straight intestine. Siphusauctum gregarium was probably an active filter-feeder, with water passing through the calyx openings, capturing food particles with its comb-like elements. It often occurs in large assemblages on single bedding planes suggesting a gregarious lifestyle, with the animal living in high tier clusters. These were probably buried en masse more or less in-situ by rapid mud flow events. Siphusauctum gregarium resembles Dinomischus, another Cambrian enigmatic stalked animal. Principal points of comparison include a long stem with a calyx containing a visceral mass and bract-like elements, and a similar lifestyle albeit occupying different tiering levels. The presence in both animals of a digestive tract with a potential stomach and anus suggest a grade of organization within bilaterians, but relationships with extant phyla are not straightforward. Thus, the broader affinities of S. gregarium remain largely unconstrained

  13. Molecular clocks and the early evolution of metazoan nervous systems.

    PubMed

    Wray, Gregory A

    2015-12-19

    The timing of early animal evolution remains poorly resolved, yet remains critical for understanding nervous system evolution. Methods for estimating divergence times from sequence data have improved considerably, providing a more refined understanding of key divergences. The best molecular estimates point to the origin of metazoans and bilaterians tens to hundreds of millions of years earlier than their first appearances in the fossil record. Both the molecular and fossil records are compatible, however, with the possibility of tiny, unskeletonized, low energy budget animals during the Proterozoic that had planktonic, benthic, or meiofaunal lifestyles. Such animals would likely have had relatively simple nervous systems equipped primarily to detect food, avoid inhospitable environments and locate mates. The appearance of the first macropredators during the Cambrian would have changed the selective landscape dramatically, likely driving the evolution of complex sense organs, sophisticated sensory processing systems, and diverse effector systems involved in capturing prey and avoiding predation. © 2015 The Author(s).

  14. Phosphate Biomineralization of Cambrian Microorganisms

    NASA Technical Reports Server (NTRS)

    McKay, David S.; Rozanov, Alexei Yu.; Hoover, Richard B.; Westall, Frances

    1998-01-01

    As part of a long term study of biological markers (biomarkers), we are documenting a variety of features which reflect the previous presence of living organisms. As we study meteorites and samples returned from Mars, our main clue to recognizing possible microbial material may be the presence of biomarkers rather than the organisms themselves. One class of biomarkers consists of biominerals which have either been precipitated directly by microorganisms, or whose precipitation has been influenced by the organisms. Such microbe-mediated mineral formation may include important clues to the size, shape, and environment of the microorganisms. The process of fossilization or mineralization can cause major changes in morphologies and textures of the original organisms. The study of fossilized terrestrial organisms can help provide insight into the interpretation of mineral biomarkers. This paper describes the results of investigations of microfossils in Cambrian phosphate-rich rocks (phosphorites) that were found in Khubsugul, Northern Mongolia.

  15. The 'Orsten': more than a Cambrian Konservat-Lagerstätte yielding exceptional preservation

    USGS Publications Warehouse

    Maas, Andreas; Braun, Andreas; Dong, Xi-Ping; Donoghue, Philip C.J.; Muller, Klaus J.; Olempska, Ewa; Repetski, John E.; Siveter, David J.; Stein, Martin; Waloszek, Dieter

    2006-01-01

    In several areas of southern Sweden, limestone nodules, locally called Orsten occur within bituminous alum shales. These shales and nodules were deposited under dysoxic conditions at the bottom of what was most likely a shallow sea during the late Middle to Upper Cambrian (ca. 500 million years ago). Subsequently, the name ‘Orsten’ has been referred to particular, mainly arthropod, fossils from such nodules, and, in a wider sense, to the specific type of preservation of minute fossil through secondarily phosphatization. This preservation is exceptional in yielding uncompacted and diagenetically undeformed three-dimensional fossils. ‘Orsten’-type preservation resulted from incrustation of a thin external layer and also by impregnation by calcium phosphate and, therefore, mineralization of the surface of the former animals during early diagenesis. Primarily, this type of preservation seems to have affected only cuticle-bearing metazoans such as cycloneuralian nemathelminths and arthropods. ‘Orsten’ preservation in this sense seems to be limited by size, in having yielded no partial or complete animals larger than 2 mm. On the other end of the scale, even larvae 100 μm long are preserved, often more complete than larger specimens, and details such as setules and pores smaller than 1 μm can be observed. Fossils preserved in such a manner are almost exclusively hollow carcasses, but can be filled secondarily; less common are completely phosphatized compact specimens. The high quality of preservation makes the Swedish ‘Orsten’ a typical Konservat-Lagerstätte. Yet, its special type of preservation is more widespread in time and geographical distribution than assumed initially, and the origin of the phosphate is not necessarily restricted just to one source. Subsequent to the first discoveries of limb fragments of Cambrian arthropods in 1975, animals in this special preservational type have been discovered in several continents and across a broad

  16. Nd-Sr isotopic and geochemical systematics in Cambrian boninites and tholeiites from Victoria, Australia

    NASA Astrophysics Data System (ADS)

    Nelson, D. R.; Crawford, A. J.; McCulloch, M. T.

    1984-11-01

    Rocks with boninitic affinities have been recognised in a number of “ophiolites”, including the Cambrian Heathcote and Mt Wellington Greenstone Belts of Victoria. Boninites and high-Mg andesites from the Heathcote Greenstone Belt show a restricted range of initial ɛ Nd values of between +3.3 to +5.8. Extremely refractory boninites from the Mt Wellington Greenstone Belt have ɛ Nd ranging from +1.3 to -9. Ti/Zr is positively correlated with Sm/Nd with the Heathcote lavas generally possessing greater depletion of Ti and enrichment of Zr relative to the middle and heavy REE with increasing LREE/HREE. These data are consistent with the generation of boninites by partial melting of refractory peridotite following invasion by LREE- and Zr-enriched, low ɛ Nd fluids. Tholeiites overlying the boninites in both greenstone belts have flat REE patterns and ɛ Nd˜+5, lower than that anticipated for lavas derived from depleted MORB source reservoirs in the Cambrian, suggesting that their source was also contaminated by a LREE-enriched, low ɛ Nd component similar to that involved in the generation of the Howqua boninites. The added components have characteristics compatible with their derivation from subducted altered oceanic crust and/or from wet subducted sediments. The identification of boninites and other low-Ti lavas in the Victorian greenstone belts is strong evidence for island arc development in southeastern Australia during the Lower Cambrian and provides further support for a subduction-related origin for many ophiolites.

  17. Cambrian cinctan echinoderms shed light on feeding in the ancestral deuterostome

    PubMed Central

    Rahman, Imran A.; Zamora, Samuel; Falkingham, Peter L.; Phillips, Jeremy C.

    2015-01-01

    Reconstructing the feeding mode of the latest common ancestor of deuterostomes is key to elucidating the early evolution of feeding in chordates and allied phyla; however, it is debated whether the ancestral deuterostome was a tentaculate feeder or a pharyngeal filter feeder. To address this, we evaluated the hydrodynamics of feeding in a group of fossil stem-group echinoderms (cinctans) using computational fluid dynamics. We simulated water flow past three-dimensional digital models of a Cambrian fossil cinctan in a range of possible life positions, adopting both passive tentacular feeding and active pharyngeal filter feeding. The results demonstrate that an orientation with the mouth facing downstream of the current was optimal for drag and lift reduction. Moreover, they show that there was almost no flow to the mouth and associated marginal groove under simulations of passive feeding, whereas considerable flow towards the animal was observed for active feeding, which would have enhanced the transport of suspended particles to the mouth. This strongly suggests that cinctans were active pharyngeal filter feeders, like modern enteropneust hemichordates and urochordates, indicating that the ancestral deuterostome employed a similar feeding strategy. PMID:26511049

  18. Cambrian to Devonian evolution of alluvial systems: The sedimentological impact of the earliest land plants

    NASA Astrophysics Data System (ADS)

    Davies, Neil S.; Gibling, Martin R.

    2010-02-01

    In present-day alluvial environments, the impact of vegetation on sedimentological processes and deposits is well known. A vegetated catchment may decrease sediment yield, sediment erodibility, Hortonian overland flow, aeolian winnowing of fines, the proportion of sediment transported as bedload, and may increase bank stability, infiltration into substrates, and bed roughness. Vegetation also promotes the production of chemically-weathered clays and soils and the adoption of a meandering style. It is generally understood that, prior to the evolution of terrestrial vegetation during the Early Palaeozoic, ancient alluvial systems were markedly different from modern systems, with many systems adopting a "sheet-braided" style. This understanding has previously informed the interpretations of many Precambrian pre-vegetation alluvial successions, but there has been relatively little work regarding Early Palaeozoic alluvial successions laid down prior to and during the initial colonization of the Earth's surface by plants. A comprehensive review of 144 Cambrian to Devonian alluvial successions documented in published literature was combined with original field data from 34 alluvial successions across Europe and North America. The study was designed to identify changes in alluvial style during the period that vegetation was evolving and first colonizing alluvial environments. An increase in mudrock proportion and sandstone maturity is apparent, along with a decrease in overall sand grain size through the Early Palaeozoic. These trends suggest that primitive vegetation cover promoted the production and preservation of muds from the mid Ordovician onwards and increased the residence time of sand-grade sediment in alluvial systems. The compilation also enables the first stratigraphic occurrence of certain vegetation-dependent sedimentary features to be pinpointed and related to the evolution of specific palaeobotanical adaptations. The first markedly heterolithic alluvial

  19. Tentaculate fossils from the Cambrian of Canada (British Columbia) and China (Yunnan) interpreted as primitive deuterostomes.

    PubMed

    Caron, Jean-Bernard; Conway Morris, Simon; Shu, Degan

    2010-03-08

    Molecular and morphological evidence unite the hemichordates and echinoderms as the Ambulacraria, but their earliest history remains almost entirely conjectural. This is on account of the morphological disparity of the ambulacrarians and a paucity of obvious stem-groups. We describe here a new taxon Herpetogaster collinsi gen. et sp. nov. from the Burgess Shale (Middle Cambrian) Lagerstätte. This soft-bodied vermiform animal has a pair of elongate dendritic oral tentacles, a flexible stolon with an attachment disc, and a re-curved trunk with at least 13 segments that is directed dextrally. A differentiated but un-looped gut is enclosed in a sac suspended by mesenteries. It consists of a short pharynx, a conspicuous lenticular stomach, followed by a narrow intestine sub-equal in length. This new taxon, together with the Lower Cambrian Phlogites and more intriguingly the hitherto enigmatic discoidal eldoniids (Cambrian-Devonian), form a distinctive clade (herein the cambroernids). Although one hypothesis of their relationships would look to the lophotrochozoans (specifically the entoprocts), we suggest that the evidence is more consistent with their being primitive deuterostomes, with specific comparisons being made to the pterobranch hemichordates and pre-radial echinoderms. On this basis some of the earliest ambulacrarians are interpreted as soft-bodied animals with a muscular stalk, and possessing prominent tentacles.

  20. Climatic-oceanic forcing on the organic accumulation across the shelf during the Early Cambrian (Age 2 through 3) in the mid-upper Yangtze Block, NE Guizhou, South China

    NASA Astrophysics Data System (ADS)

    Yeasmin, Rumana; Chen, Daizhao; Fu, Yong; Wang, Jianguo; Guo, Zenghui; Guo, Chuan

    2017-02-01

    The organic-rich sediments were widely deposited over the entire Yangtze Block during the Early Cambrian (late Nemakit-Daldynian to Botomian). In the mid-upper Yangtze region, northeastern Guizhou, South China, they comprise, in ascending order, the Niutitang, Jiumenchong and lower Bianmachong formations which are dominated by black shales except the middle one characterized by interbedded shales-limestones. Three third-order depositional sequences are identified in the two studied sections located on the upper slope to basin of the open shelf. The organic-rich sediments were mostly deposited notably during transgressions on the shallower upper slope-margin (TOC up to 25 wt.%) where they are characterized by co-increases in C, P, Fe, and Ba concentrations, indicating the highest organic productivity and coupled C, P and Fe cycling there. In contrast, in the shelf basin, the concomitant organic-rich sediments yield lower organic (TOC <10 wt.% mostly) and Ba abundances, and generally show antithetic relationship of TOC with P and Fe variations, indicative of a relatively low organic productivity and transient decoupled C, P and Fe cycling in the deeper basin. This spatial difference in C, P, Fe and Ba cycling as a whole thus indicate that the highest organic productivity wedge on the shallower shelf upper slope-margin was likely induced by the oceanic upwelling from the restricted basin where the P and Fe nutrients were transiently depleted without timely renewal. Meanwhile, multiple redox proxies, including V/Al, Ni/Al and Cr/Al ratios vs. TOC contents, together with V/(V + Ni) and V/Cr ratios, generally characterize a more severe euxinic water wedge localized on the shallower upper slope-margin of shelf where the organic production culminated, but a nonsulfidic (ferruginous) anoxic state in the basin due to the relatively low sulfate level. Meanwhile, episodic depletion of trace elements in the deeper basin during the higher organic productivity also reconciles the

  1. Siliceous spicules in a vauxiid sponge (Demospongia) from the Kaili Biota(Cambrian Stage 5), Guizhou, South China

    NASA Astrophysics Data System (ADS)

    Yang, X.-L.; Zhao, Y.-L.; Babcock, L. E.; Peng, J.

    2017-02-01

    Fossils of the sponge Angulosuspongia sinensis from calcareous mudstones of the middle and upper part of the Kaili Formation (Cambrian Stage 5) in the Jianhe area of Guizhou province, South China, exhibit an apparently reticulate pattern, characteristic of the Vauxiidae. Energy Dispersive X-Ray Spectrometry (EDS) and Raman spectroscopy analysis indicate the presence of silica in the skeletal elements of these fossils, suggesting that this taxon possessed a skeleton comprised of spicules. This is the first confirmation of siliceous skeletal elements in fossils of the family Vauxiidae, and it lends support to the hypothesis that some early demosponges possessed biomineralized siliceous skeletons, which were subsequently lost and replaced by spongin later in the evolutionary history of this lineage. The new materials provide critical insight into the phylogeny and evolution of biomineralization in the Demosopongiae.

  2. Uranium in the Upper Cambrian black shale of Sweden

    USGS Publications Warehouse

    McKelvey, Vincent Ellis

    1955-01-01

    The Peltura zone of the Upper Cambrian black shales of Sweden contains about 0.02 percent uranium. Maximum amounts are present in rocks deposited in an embayment in the sea and in rocks in or closely adjacent to that part of the vertical sequence that contains maximum amounts of distillable oil, total organic matter, pyrite, and a black highly uraniferous kerogen called "kolm". Available data suggest that the precipitation of uranium is favored by a low redox potential and that the uranium in the shale matrix may be in fine-grained kolm.

  3. Cambrian intermediate-mafic magmatism along the Laurentian margin: Evidence for flood basalt volcanism from well cuttings in the Southern Oklahoma Aulacogen (U.S.A.)

    NASA Astrophysics Data System (ADS)

    Brueseke, Matthew E.; Hobbs, Jasper M.; Bulen, Casey L.; Mertzman, Stanley A.; Puckett, Robert E.; Walker, J. Douglas; Feldman, Josh

    2016-09-01

    The Southern Oklahoma Aulocogen (SOA) stretches from southern Oklahoma through the Texas panhandle and into Colorado and New Mexico, and contains mafic through silicic magmatism related to the opening of the Iapetus Ocean during the early Cambrian. Cambrian magmatic products are best exposed in the Wichita Mountains (Oklahoma), where they have been extensively studied. However, their ultimate derivation is still somewhat contentious and centers on two very different models: SOA magmatism has been suggested to occur via [1] continental rifting (with or without mantle plume emplacement) or [2] transform-fault related magmatism (e.g., leaky strike-slip faults). Within the SOA, the subsurface in and adjacent to the Arbuckle Mountains in southern Oklahoma contains thick sequences of mafic to intermediate lavas, intrusive bodies, and phreatomagmatic deposits interlayered with thick, extensive rhyolite lavas, thin localized tuffs, and lesser silicic intrusive bodies. These materials were first described in the Arbuckle Mountains region by a 1982 drill test (Hamilton Brothers Turner Falls well) and the best available age constraints from SOA Arbuckle Mountains eruptive products are 535 to 540 Ma. Well cuttings of the mafic through intermediate units were collected from that well and six others and samples from all but the Turner Falls and Morton wells are the focus of this study. Samples analyzed from the wells are dominantly subalkaline, tholeiitic, and range from basalt to andesite. Their overall bulk major and trace element chemistry, normative mineralogy, and Srsbnd Nd isotope ratios are similar to magmas erupted/emplaced in flood basalt provinces. When compared with intrusive mafic rocks that crop out in the Wichita Mountains, the SOA well cuttings are geochemically most similar to the Roosevelt Gabbros. New geochemical and isotope data presented in this study, when coupled with recent geophysical work in the SOA and the coeval relationship with rhyolites, indicates

  4. Tidal-bundle sequences in the Jordan Sandstone (Upper Cambrian), southeastern Minnesota, U.S.A.: Evidence for tides along inboard shorelines of the Sauk Epicontinental Sea

    USGS Publications Warehouse

    Tape, C.H.; Cowan, Clinton A.; Runkel, Anthony C.

    2003-01-01

    This study documents for the first time tidal bundling in a lower Paleozoic sheet sandstone from the cratonic interior of North America, providing insights into the hydrodynamics of ancient epicontinental seas. The Jordan Sandstone (Upper Cambrian) in the Upper Mississippi Valley contains large-scale planar tabular cross-sets with tidal-bundle sequences, which were analyzed in detail at an exceptional exposure. Tidal-bundle sequences (neap-spring-neap cycles) were delineated by foreset thickening-thinning patterns and composite shale drapes, the latter of which represent accumulations of mud during the neap tides of neap-spring-neap tidal cycles. Fourier analysis of the bundle thickness data from the 26 measurable bundle sequences revealed cycles ranging from 15 to 34 bundles per sequence, which suggests a semidiurnal or mixed tidal system along this part of the Late Cambrian shoreline. We extend the tidal interpretation to widespread occurrences of the same facies in outcrops of lesser quality, where the facies is recognizable but too few bundles are exposed for tidal cycles to be measured. By doing so, this study shows that tidally generated deposits have a significant geographic and temporal extent in Upper Cambrian strata of central mid-continent North America. The deposition and preservation of tidal facies was related to the intermittent development of shoreline embayments during transgressions. The tidally dominated deposits filled ravined topographies that were repeatedly developed on the updip parts of the shoreface. Resulting coastal geomorphologies, accompanied perhaps by larger-scale changes in basinal conditions and/or configuration, led to changes in depositional conditions from wave-dominated to tide-dominated. Outcrops of the Jordan Sandstone tidal facies in the Upper Mississippi Valley represent the farthest inboard recorded transmission of ocean-generated tides in the Laurentian epicontinental seas, demonstrating that tidal currents were

  5. Oxygen as a driver of early arthropod micro-benthos evolution.

    PubMed

    Williams, Mark; Vannier, Jean; Corbari, Laure; Massabuau, Jean-Charles

    2011-01-01

    We examine the physiological and lifestyle adaptations which facilitated the emergence of ostracods as the numerically dominant Phanerozoic bivalve arthropod micro-benthos. The PO(2) of modern normoxic seawater is 21 kPa (air-equilibrated water), a level that would cause cellular damage if found in the tissues of ostracods and much other marine fauna. The PO(2) of most aquatic breathers at the cellular level is much lower, between 1 and 3 kPa. Ostracods avoid oxygen toxicity by migrating to waters which are hypoxic, or by developing metabolisms which generate high consumption of O(2). Interrogation of the Cambrian record of bivalve arthropod micro-benthos suggests a strong control on ecosystem evolution exerted by changing seawater O(2) levels. The PO(2) of air-equilibrated Cambrian-seawater is predicted to have varied between 10 and 30 kPa. Three groups of marine shelf-dwelling bivalve arthropods adopted different responses to Cambrian seawater O(2). Bradoriida evolved cardiovascular systems that favoured colonization of oxygenated marine waters. Their biodiversity declined during intervals associated with black shale deposition and marine shelf anoxia and their diversity may also have been curtailed by elevated late Cambrian (Furongian) oxygen-levels that increased the PO(2) gradient between seawater and bradoriid tissues. Phosphatocopida responded to Cambrian anoxia differently, reaching their peak during widespread seabed dysoxia of the SPICE event. They lacked a cardiovascular system and appear to have been adapted to seawater hypoxia. As latest Cambrian marine shelf waters became well oxygenated, phosphatocopids went extinct. Changing seawater oxygen-levels and the demise of much of the seabed bradoriid micro-benthos favoured a third group of arthropod micro-benthos, the ostracods. These animals adopted lifestyles that made them tolerant of changes in seawater O(2). Ostracods became the numerically dominant arthropod micro-benthos of the Phanerozoic. Our work

  6. Oxygen as a Driver of Early Arthropod Micro-Benthos Evolution

    PubMed Central

    Williams, Mark; Vannier, Jean; Corbari, Laure; Massabuau, Jean-Charles

    2011-01-01

    Background We examine the physiological and lifestyle adaptations which facilitated the emergence of ostracods as the numerically dominant Phanerozoic bivalve arthropod micro-benthos. Methodology/Principal Findings The PO2 of modern normoxic seawater is 21 kPa (air-equilibrated water), a level that would cause cellular damage if found in the tissues of ostracods and much other marine fauna. The PO2 of most aquatic breathers at the cellular level is much lower, between 1 and 3 kPa. Ostracods avoid oxygen toxicity by migrating to waters which are hypoxic, or by developing metabolisms which generate high consumption of O2. Interrogation of the Cambrian record of bivalve arthropod micro-benthos suggests a strong control on ecosystem evolution exerted by changing seawater O2 levels. The PO2 of air-equilibrated Cambrian-seawater is predicted to have varied between 10 and 30 kPa. Three groups of marine shelf-dwelling bivalve arthropods adopted different responses to Cambrian seawater O2. Bradoriida evolved cardiovascular systems that favoured colonization of oxygenated marine waters. Their biodiversity declined during intervals associated with black shale deposition and marine shelf anoxia and their diversity may also have been curtailed by elevated late Cambrian (Furongian) oxygen-levels that increased the PO2 gradient between seawater and bradoriid tissues. Phosphatocopida responded to Cambrian anoxia differently, reaching their peak during widespread seabed dysoxia of the SPICE event. They lacked a cardiovascular system and appear to have been adapted to seawater hypoxia. As latest Cambrian marine shelf waters became well oxygenated, phosphatocopids went extinct. Changing seawater oxygen-levels and the demise of much of the seabed bradoriid micro-benthos favoured a third group of arthropod micro-benthos, the ostracods. These animals adopted lifestyles that made them tolerant of changes in seawater O2. Ostracods became the numerically dominant arthropod micro-benthos of

  7. Tentaculate Fossils from the Cambrian of Canada (British Columbia) and China (Yunnan) Interpreted as Primitive Deuterostomes

    PubMed Central

    Caron, Jean-Bernard; Conway Morris, Simon; Shu, Degan

    2010-01-01

    Molecular and morphological evidence unite the hemichordates and echinoderms as the Ambulacraria, but their earliest history remains almost entirely conjectural. This is on account of the morphological disparity of the ambulacrarians and a paucity of obvious stem-groups. We describe here a new taxon Herpetogaster collinsi gen. et sp. nov. from the Burgess Shale (Middle Cambrian) Lagerstätte. This soft-bodied vermiform animal has a pair of elongate dendritic oral tentacles, a flexible stolon with an attachment disc, and a re-curved trunk with at least 13 segments that is directed dextrally. A differentiated but un-looped gut is enclosed in a sac suspended by mesenteries. It consists of a short pharynx, a conspicuous lenticular stomach, followed by a narrow intestine sub-equal in length. This new taxon, together with the Lower Cambrian Phlogites and more intriguingly the hitherto enigmatic discoidal eldoniids (Cambrian-Devonian), form a distinctive clade (herein the cambroernids). Although one hypothesis of their relationships would look to the lophotrochozoans (specifically the entoprocts), we suggest that the evidence is more consistent with their being primitive deuterostomes, with specific comparisons being made to the pterobranch hemichordates and pre-radial echinoderms. On this basis some of the earliest ambulacrarians are interpreted as soft-bodied animals with a muscular stalk, and possessing prominent tentacles. PMID:20221405

  8. Detrital zircon U-Pb geochronology of Cambrian to Triassic miogeoclinal and eugeoclinal strata of Sonora, Mexico

    USGS Publications Warehouse

    Gehrels, G.E.; Stewart, John H.

    1998-01-01

    One hundred and eighty two individual detrital zircon grains from Cambrian through Permian miogeoclinal strata, Ordovician eugeoclinal rocks, and Triassic post-orogenic sediments in northwestern Sonora have been analyzed. During Cambrian, Devonian, Permian, and Triassic time, most zircons accumulating along this part of the Cordilleran margin were shed from 1.40-1.45 and 1.62-1.78 Ga igneous rocks that are widespread in the southwestern United States and northwestern Mexico. Zircons with ages of approximately 1.11 Ga are common in Cambrian strata and were apparently shed from granite bodies near the sample site. The sources of 225-280 Ma zircons in our Triassic sample are more problematic, as few igneous rocks of these ages are recognized in northwestern Mexico. Such sources may be present but unrecognized, or the grains could have been derived from igneous rocks of the appropriate ages to the northwest in the Mojave Desert region, to the east in Chihuahua and Coahuila, or to the south in accreted(?) arc-type terranes. Because the zircon grains in our Cambrian and Devonian to Triassic samples could have accumulated in proximity to basement rocks near their present position or in the Death Valley region of southern California, our data do not support or refute the existence of the Mojave-Sonora megashear. Ordovician strata of both miogeoclinal and eugeoclinal affinity are dominated by >1.77 Ga detrital zircons, which are considerably older than most basement rocks in the region. Zircon grains in the miogeoclinal sample were apparently derived from the Peace River arch area of northwestern Canada and transported southward by longshore currents. The eugeoclinal grains may also have come from the Peace River arch region, with southward transport by either sedimentary or tectonic processes, or they may have been shed from off-shelf slivers of continents (perhaps Antarctica?) removed from the Cordilleran margin during Neoproterozoic rifting. It is also possible that the

  9. Siliceous spicules in a vauxiid sponge (Demospongia) from the Kaili Biota(Cambrian Stage 5), Guizhou, South China

    PubMed Central

    Yang, X.-L.; Zhao, Y.-L.; Babcock, L. E.; Peng, J.

    2017-01-01

    Fossils of the sponge Angulosuspongia sinensis from calcareous mudstones of the middle and upper part of the Kaili Formation (Cambrian Stage 5) in the Jianhe area of Guizhou province, South China, exhibit an apparently reticulate pattern, characteristic of the Vauxiidae. Energy Dispersive X-Ray Spectrometry (EDS) and Raman spectroscopy analysis indicate the presence of silica in the skeletal elements of these fossils, suggesting that this taxon possessed a skeleton comprised of spicules. This is the first confirmation of siliceous skeletal elements in fossils of the family Vauxiidae, and it lends support to the hypothesis that some early demosponges possessed biomineralized siliceous skeletons, which were subsequently lost and replaced by spongin later in the evolutionary history of this lineage. The new materials provide critical insight into the phylogeny and evolution of biomineralization in the Demosopongiae. PMID:28220860

  10. Cambrian rivers and floodplains: the significance of microbial cementation, groundwater and aeolian sediment transport

    NASA Astrophysics Data System (ADS)

    Reesink, A. J. H.; Best, J.; Freiburg, J. T.; Nathan, W.

    2016-12-01

    Rivers that existed before land plants colonized the Earth are commonly considered to be unaffected by microbial activity on their floodplains, because the limited cementation produced by microbial activity is insufficient to stabilize the river banks. Although this assumption is likely correct, such emphasis on channel dynamics ignores the potential role of floodplain dynamics as an integral component of the river system. Detailed analysis of cores from the Cambrian Mount Simon Sandstone, Illinois, suggests that a significant proportion of the terrestrial sequence is composed of flat-bedded `crinkly' structures that provide evidence of cementation by soil crusts and microbial biofilms, and that promoted the adhesion of sediment to sticky surfaces. Wind ripples and local desert pavements were abundant. These findings highlight that sediment deposition on Cambrian floodplains was often dominated by wind in locations where the ground water table reached the surface, and was thus likely independent of sediment transport within the river channel. Erosion by wind would thus have been hindered by surface cementation and the formation of desert pavements. Such ground water control on deposition, and resistance to erosion by floodplain surface hardening, appear to have been the primary controls on Cambrian floodplain topography. Because floodplain topography poses a key control on channel and floodplain flow, these processes may have affected patterns of erosion and deposition, as well as reach-scale dynamics such as channel avulsions. The autonomous operation of wind-and-groundwater controlled floodplains makes pre-vegetated river systems more sensitive to climatic conditions such as precipitation and evaporation, and strikingly different from those that occurred after the development of land plants.

  11. Stratigraphic framework of Cambrian and Ordovician rocks in the central Appalachian Basin from Medina County, Ohio, through southwestern and south-central Pennsylvania to Hampshire County, West Virginia

    USGS Publications Warehouse

    Ryder, Robert T.; Harris, Anita G.; Repetski, John E.; revised and digitized by Crangle, Robert D.

    2003-01-01

    are identified by the following descriptive names: (1) sandstone, shale, limestone, and dolomite unit, (2) dolomite and sandstone unit, (3) limestone and black shale unit, and (4) shale and sandstone unit. Each of these units and their associated subunits thicken from west to east across the restored section to a maximum near the depositional axis of the Rome trough and then thin eastward to the end of the section. The sandstone, shale, limestone, and dolomite unit is largely confined to the asymmetric graben that marks the initial phase of the Rome trough. This unit is Early and Middle Cambrian in age and consists, in ascending order, of a basal sandstone unit (undrilled but probably present), the Tomstown Dolomite (undrilled but probably present), the Waynesboro Formation, and the Pleasant Hill Limestone and its equivalent lower one-third of the Elbrook Formation at the eastern end of the section. The dolomite and sandstone unit forms the core of the Cambrian and Ordovician sequence. In the Rome trough and on the adjoining South-central Pennsylvania arch, this unit consists, in ascending order, of the Middle and Upper Cambrian Warrior Formation and the equivalent upper two-thirds of the Elbrook Formation at the eastern end of the section, the Upper Cambrian Gatesburg Formation, and the Lower Ordovician and Middle Ordovician (Whiterockian and Chazyan) Beekmantown Group. West of the Ohio-West Virginia hinge zone, the dolomite and sandstone unit consists, in ascending order, of the Conasauga Formation of Janssens (1973), the Krysik sandstone of driller's usage, the B zone of Calvert (1964), the Knox Dolomite and the associated Rose Run Sandstone Member, and the Wells Creek Formation. The widespread Knox unconformity is located at the base of the Wells Creek Formation and at or near the top of the adjoining Beekmantown Group, except near the depositional axis of the Rome trough, where the unconformity seems to be absent. The limestone and black shale unit i

  12. Le volcanisme cambrien du Maroc central : implications géodynamiquesThe Central Morocco Cambrian volcanism: geodynamic implications

    NASA Astrophysics Data System (ADS)

    Ouali, Houssa; Briand, Bernard; Bouchardon, Jean-Luc; Capiez, Paul

    2003-05-01

    In southeastern Central Morocco, the Bou-Acila volcanic complex is considered of Cambrian age. In spite of low-grade metamorphic effect, initial volcanic texture and mineralogy can be recognized and volcanic rocks are dominated by dolerites and porphyric dolerites. The initial mineralogy is composed of plagioclases, pyroxenes and dark minerals. A secondary mineral assemblage is composed of albite, epidote, chlorite and calcite. According to their immobile elements compositions, the southeastern central Morocco metavolcanites are of within-plate continental tholeiites. This volcanism and those recognized in many other areas in Morocco confirm a Cambrian extensive episode within the Gondwana supercontinent. To cite this article: H. Ouali et al., C. R. Geoscience 335 (2003).To cite this article: H. Ouali et al., C. R. Geoscience 335 (2003).

  13. Sedimentology of SPICE (Steptoean positive carbon isotope excursion): A high-resolution trace fossil and microfabric analysis of the middle to late Cambrian Alum Shale Formation, southern Sweden

    USGS Publications Warehouse

    Egenhoff, Sven; Fishman, Neil; Ahlberg, Per; Maletz, Jorg; Jackson, Allison; Kolte, Ketki; Lowers, Heather; Mackie, James; Newby, Warren; Petrowsky, Matthew

    2015-01-01

    The Cambrian Alum Shale Formation in the Andrarum-3 core from Scania, southern Sweden, consists of black siliciclastic mudstone with minor carbonate intercalations. Four facies comprise three siliciclastic mudstones and one fine-grained carbonate. The facies reflect deposition along a transect from deep ramp to basin on a Cambrian shelf. The three mudstone facies contain abundant clay clasts and laterally variable siltstone laminae. Bed-load transport processes seem to have dominated deposition on this deep shelf. These sedimentary rocks record mainly event deposition, and only relatively few, thin laminae probably resulted from suspension settling. The Alum Shale Formation deep shelf did not show a bioturbation gradient, but fecal strings are common and Planolites burrows are rare in all mudstone facies. Evidence for biotic colonization indicates that this mudstone environment was not persistently anoxic, but rather was most likely intermittently dysoxic. The Alum Shale Formation in the Andrarum-3 core shows an overall decrease of grain size, preserved energy indicators, and carbonate content upsection interpreted to reflect a deepening upward. The succession can also be divided into four small-scale fining-upward cycles that represent deepening, and four overlying coarsening-upward cycles that represent upward shallowing.

  14. Arthropod colonization of land--linking molecules and fossils in oribatid mites (Acari, Oribatida).

    PubMed

    Schaefer, Ina; Norton, Roy A; Scheu, Stefan; Maraun, Mark

    2010-10-01

    Terrestrial fossils that document the early colonization of land are scarce for >100 my after the Cambrian explosion. This raises the question whether life on land did not exist or just did not fossilize. With a molecular dating technique, we analyzed the origin of terrestrial chelicerate microarthropods (Acari, Oribatida) which have a fossil record since the Middle Devonian that is exceptional among soil animals. Our results suggest that oribatid mites originated in the Precambrian (571+/-37 mya) and that the radiation of basal groups coincides with the gap in the terrestrial fossil record between the Cambrian explosion and the earliest fossilized records of continental ecosystems. Further, they suggest that the colonization of land started via the interstitial, approximately 150 my earlier than the oldest fossils of terrestrial ecosystems. Overall, the results imply that omnivorous and detritivorous arthropods formed a major component in early terrestrial food webs, thereby facilitating the invasion of terrestrial habitats by later colonizers of higher trophic levels. Copyright 2010 Elsevier Inc. All rights reserved.

  15. Sedimentary records on the subduction-accretion history of the Russian Altai, northwestern Central Asian Orogenic Belt

    NASA Astrophysics Data System (ADS)

    Chen, Ming; Sun, Min

    2017-04-01

    The Russian Altai, comprising the northern segment of the Altai-Mongolian terrane (AM) in the south, the Gorny Altai terrane (GA) in the north and the intervening Charysh-Terekta-Ulagan-Sayan suture zone, is a key area of the northwestern Central Asian Orogenic Belt (CAOB). A combined geochemical and detrital zircon study was conducted on the (meta-)sedimentary sequences from the Russian Altai to reveal the tectono-magmatic history of these two terranes and their amalgamation history, which in turn place constraints on the accretionary orogenesis and crustal growth in the CAOB. The Cambrian-Ordovician meta-sedimentary rocks from the northern AM are dominated by immature sediments possibly sourced from intermediate-felsic igneous rocks. Geochemical data show that the sediments were likely deposited in a continental arc-related setting. Zircons separated from these rocks are mainly 566-475 Ma and 1015-600 Ma old, comparable to the magmatic records of the Tuva-Mongolian terrane and surrounding island arcs in the western Mongolia. The similar source nature, provenance and depositional setting of these rocks to the counterparts from the Chinese Altai (i.e., the southern AM) imply that the whole AM possibly represents a coherent accretionary prism of the western Mongolia in the early Paleozoic rather than a Precambrian continental block with passive marginal deposition as previously thought. In contrast, the Cambrian to Silurian (meta-)sedimentary rocks from the GA are characterized by a unitary zircon population with ages of 640-470 Ma, which were potentially sourced from the Kuznetsk-Altai intra-oceanic island arc in the east of this terrane. The low abundance of 640-540 Ma zircons (5%) may attest that this arc was under a primitive stage in the late Neoproterozoic, when mafic igneous rocks dominated. However, the voluminous 530-470 Ma zircons (95%) suggest that this arc possibly evolved toward a mature one in the Cambrian to early Ordovician with increasing amount of

  16. Provenance of north Gondwana Cambrian-Ordovician sandstone: U-Pb SHRIMP dating of detrital zircons from Israel and Jordan

    USGS Publications Warehouse

    Kolodner, K.; Avigad, D.; McWilliams, M.; Wooden, J.L.; Weissbrod, T.; Feinstein, S.

    2006-01-01

    A vast sequence of quartz-rich sandstone was deposited over North Africa and Arabia during Early Palaeozoic times, in the aftermath of Neoproterozoic Pan-African orogeny and the amalgamation of Gondwana. This rock sequence forms a relatively thin sheet (1-3 km thick) that was transported over a very gentle slope and deposited over a huge area. The sense of transport indicates unroofing of Gondwana terranes but the exact provenance of the siliciclastic deposit remains unclear. Detrital zircons from Cambrian arkoses that immediately overlie the Neoproterozoic Arabian-Nubian Shield in Israel and Jordan yielded Neoproterozoic U-Pb ages (900-530 Ma), suggesting derivation from a proximal source such as the Arabian-Nubian Shield. A minor fraction of earliest Neoproterozoic and older age zircons was also detected. Upward in the section, the proportion of old zircons increases and reaches a maximum (40%) in the Ordovician strata of Jordan. The major earliest Neoproterozoic and older age groups detected are 0.95-1.1, 1.8-1.9 and 2.65-2.7 Ga, among which the 0.95-1.1 Ga group is ubiquitous and makes up as much as 27% in the Ordovician of Jordan, indicating it is a prominent component of the detrital zircon age spectra of northeast Gondwana. The pattern of zircon ages obtained in the present work reflects progressive blanketing of the northern Arabian-Nubian Shield by Cambrian-Ordovician sediments and an increasing contribution from a more distal source, possibly south of the Arabian-Nubian Shield. The significant changes in the zircon age signal reflect many hundreds of kilometres of southward migration of the provenance. ?? 2006 Cambridge University Press.

  17. Early rifting deposition: examples from carbonate sequences of Sardinia (Cambrian) and Tuscany (Triassic-Jurassic), Italy: an analogous tectono-sedimentary and climatic context

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Cocozza, T.; Gandin, A.

    Lower Cambrian Ceroide Limestone (Sardinia) and Lower Jurassic Massiccio Limestone (Tuscany) belong to sequences deposited in analogous tectono-sedimentary context: the former linked to the Caledonian Sardic Phase, the latter to the Alpine Orogeny. Both units consist of massive pure limestone characterized by marginal and lagoonal sequences repeatedly interfingering in the same geological structure. This distribution indicates a morphology of the platforms composed of banks (marginal facies) and shallow basins (lagoonal facies) comparable with a Bahamian complex. Dolomitization affects patchily the massive limestone bodies, and karstic features, breccias, and sedimentary dikes occur at their upper boundary. Both units overlie early dolomitemore » and evaporites (sabkha facies) containing siliciclastic intercalations in their lower and/or upper part and are unconformably covered by open-shelf red (hematitic), nodular limestone Ammonitico Rosso facies). The sedimentary evolution of the two sequences appears to have been controlled by synsedimentary tectonics whose major effects are the end of the terrigenous input, the bank-and-basin morphology of the platform, the irregular distribution of the dolomitization, and the nodular fabric of the overlying facies. The end of the Bahamian-type system is marked by the karstification of the emerged blocks and is followed by their differential sinking and burial under red-nodular facies. From a geodynamic viewpoint, sequences composed of Bahamian-like platform carbonates followed by Ammonitico Rosso facies imply deposition along continental margins subjected to block-faulting during an extensional regime connected with the beginning of continental rifting. Moreover, the variation from sabkha to Bahamian conditions suggests the drifting of the continent from arid to humid, tropical areas.« less

  18. Fossil embryos from the Middle and Late Cambrian period of Hunan, south China.

    PubMed

    Dong, Xi-Ping; Donoghue, Philip C J; Cheng, Hong; Liu, Jian-Bo

    2004-01-15

    Comparative embryology is integral to uncovering the pattern and process of metazoan phylogeny, but it relies on the assumption that life histories of living taxa are representative of their antecedents. Fossil embryos provide a crucial test of this assumption and, potentially, insight into the evolution of development, but because discoveries so far lack phylogenetic constraint, their significance is moot. Here we describe a collection of embryos from the Middle and Late Cambrian period (500 million years ago) of Hunan, south China, that preserves stages of development from cleavage to the pre-hatching embryo of a direct-developing animal comparable to living Scalidophora (phyla Priapulida, Kinorhyncha, Loricifera). The latest-stage embryos show affinity to the Lower Cambrian embryo Markuelia, whose life-history strategy contrasts both with the primitive condition inferred for metazoan phyla and with many proposed hypotheses of affinity, all of which prescribe indirect development. Phylogenetic tests based on these embryological data suggest a stem Scalidophora affinity. These discoveries corroborate, rather than contradict, the predictions of comparative embryology, providing direct historical support for the view that the life-history strategies of living taxa are representative of their stem lineages.

  19. Fuxianhuiid ventral nerve cord and early nervous system evolution in Panarthropoda.

    PubMed

    Yang, Jie; Ortega-Hernández, Javier; Butterfield, Nicholas J; Liu, Yu; Boyan, George S; Hou, Jin-Bo; Lan, Tian; Zhang, Xi-Guang

    2016-03-15

    Panarthropods are typified by disparate grades of neurological organization reflecting a complex evolutionary history. The fossil record offers a unique opportunity to reconstruct early character evolution of the nervous system via exceptional preservation in extinct representatives. Here we describe the neurological architecture of the ventral nerve cord (VNC) in the upper-stem group euarthropod Chengjiangocaris kunmingensis from the early Cambrian Xiaoshiba Lagerstätte (South China). The VNC of C. kunmingensis comprises a homonymous series of condensed ganglia that extend throughout the body, each associated with a pair of biramous limbs. Submillimetric preservation reveals numerous segmental and intersegmental nerve roots emerging from both sides of the VNC, which correspond topologically to the peripheral nerves of extant Priapulida and Onychophora. The fuxianhuiid VNC indicates that ancestral neurological features of Ecdysozoa persisted into derived members of stem-group Euarthropoda but were later lost in crown-group representatives. These findings illuminate the VNC ground pattern in Panarthropoda and suggest the independent secondary loss of cycloneuralian-like neurological characters in Tardigrada and Euarthropoda.

  20. Fuxianhuiid ventral nerve cord and early nervous system evolution in Panarthropoda

    PubMed Central

    Yang, Jie; Ortega-Hernández, Javier; Butterfield, Nicholas J.; Liu, Yu; Boyan, George S.; Hou, Jin-bo; Lan, Tian; Zhang, Xi-guang

    2016-01-01

    Panarthropods are typified by disparate grades of neurological organization reflecting a complex evolutionary history. The fossil record offers a unique opportunity to reconstruct early character evolution of the nervous system via exceptional preservation in extinct representatives. Here we describe the neurological architecture of the ventral nerve cord (VNC) in the upper-stem group euarthropod Chengjiangocaris kunmingensis from the early Cambrian Xiaoshiba Lagerstätte (South China). The VNC of C. kunmingensis comprises a homonymous series of condensed ganglia that extend throughout the body, each associated with a pair of biramous limbs. Submillimetric preservation reveals numerous segmental and intersegmental nerve roots emerging from both sides of the VNC, which correspond topologically to the peripheral nerves of extant Priapulida and Onychophora. The fuxianhuiid VNC indicates that ancestral neurological features of Ecdysozoa persisted into derived members of stem-group Euarthropoda but were later lost in crown-group representatives. These findings illuminate the VNC ground pattern in Panarthropoda and suggest the independent secondary loss of cycloneuralian-like neurological characters in Tardigrada and Euarthropoda. PMID:26933218

  1. Modern freshwater microbialite analogues for ancient dendritic reef structures

    NASA Technical Reports Server (NTRS)

    Laval, B.; Cady, S. L.; Pollack, J. C.; McKay, C. P.; Bird, J. S.; Grotzinger, J. P.; Ford, D. C.; Bohm, H. R.

    2000-01-01

    Microbialites are organosedimentary structures that can be constructed by a variety of metabolically distinct taxa. Consequently, microbialite structures abound in the fossil record, although the exact nature of the biogeochemical processes that produced them is often unknown. One such class of ancient calcareous structures, Epiphyton and Girvanella, appear in great abundance during the Early Cambrian. Together with Archeocyathids, stromatolites and thrombolites, they formed major Cambrian reef belts. To a large extent, Middle to Late Cambrian reefs are similar to Precambrian reefs, with the exception that the latter, including terminal Proterozoic reefs, do not contain Epiphyton or Girvanella. Here we report the discovery in Pavilion Lake, British Columbia, Canada, of a distinctive assemblage of freshwater calcite microbialites, some of which display microstructures similar to the fabrics displayed by Epiphyton and Girvanella. The morphologies of the modern microbialites vary with depth, and dendritic microstructures of the deep water (> 30 m) mounds indicate that they may be modern analogues for the ancient calcareous structures. These microbialites thus provide an opportunity to study the biogeochemical interactions that produce fabrics similar to those of some enigmatic Early Cambrian reef structures.

  2. High Resolution Biostratigraphy and the Origin of the Basal Cambrian Bedded Chert from the Aksu Area (Tarim Block, Northwestern China)

    NASA Astrophysics Data System (ADS)

    Wang, Z.; Liu, H.; Dong, L.

    2017-12-01

    The early Cambrian Yurtus Formation in the Aksu area (Tarim block, northwestern China) consists of two lithostratigraphic units, lower black shale with interbedded chert unit and upper siltstone/carbonate unit. This time period represents the most important Proterozoic- Phanerozoic transition in earth's history. In recent years, the black shale has been confirmed to have high hydrocarbon generation potential. However, the depositional environment of the Yurtus Formation remains controversial and the biostratigraphic constrains are rather poor. The chert that is interbedded with black shale in the Yurtus Formation provides an exceptional taphonomic window to capture the diversity of the early Cambrian microfossils. Meanwhile, the origin of the bedded chert would give us some insight into the environmental background when the source rock was deposited. Therefore, in this research, we focus on the chert in the lower Yurtus formation and our purpose is to establish high resolution biostratigraphic framework and to better understand the depositional environment of the source rock. We investigated 4 sections in the Tarim basin: Kungaikuotan, Sugaite, Kule, and Yurtus VI. Abundant acritarch fossils have been identified, including Heliosphaeridium ampliatum, Yurtusia uniformis, and Comasphaeridium annulare. The tubular fossil Megathrix longus is also very common in this formation. In addition, two new types of specimens have been discovered, sheet-like encrolled fossils ( 0.5 mm in size) and regular spindle-like double layered microfossils ( 10μm in diameter). All of these fossils have constant occurrences in the studied sections, and can be well correlated with those yielded from the equivalent interval in South China. The biostratigraphic work suggests the source rock in the lower unit of the Yurtus Formation could be correlated with the Meishucunian small shelly fossil assemblage I and II. The Gemenium/Silicon ratio of the Yurtus chert is less than 1

  3. The Origin and Early Radiation of Archosauriforms: Integrating the Skeletal and Footprint Record.

    PubMed

    Bernardi, Massimo; Klein, Hendrik; Petti, Fabio Massimo; Ezcurra, Martín D

    2015-01-01

    We present a holistic approach to the study of early archosauriform evolution by integrating body and track records. The ichnological record supports a Late Permian-Early Triassic radiation of archosauriforms not well documented by skeletal material, and new footprints from the Upper Permian of the southern Alps (Italy) provide evidence for a diversity not yet sampled by body fossils. The integrative study of body fossil and footprint data supports the hypothesis that archosauriforms had already undergone substantial taxonomic diversification by the Late Permian and that by the Early Triassic archosauromorphs attained a broad geographical distribution over most parts of Pangea. Analysis of body size, as deduced from track size, suggests that archosauriform average body size did not change significantly from the Late Permian to the Early Triassic. A survey of facies yielding both skeletal and track record indicate an ecological preference for inland fluvial (lacustrine) environments for early archosauromorphs. Finally, although more data is needed, Late Permian chirotheriid imprints suggest a shift from sprawling to erect posture in archosauriforms before the end-Permian mass extinction event. We highlight the importance of approaching palaeobiological questions by using all available sources of data, specifically through integrating the body and track fossil record.

  4. The Origin and Early Radiation of Archosauriforms: Integrating the Skeletal and Footprint Record

    PubMed Central

    Bernardi, Massimo; Klein, Hendrik; Petti, Fabio Massimo; Ezcurra, Martín D.

    2015-01-01

    We present a holistic approach to the study of early archosauriform evolution by integrating body and track records. The ichnological record supports a Late Permian–Early Triassic radiation of archosauriforms not well documented by skeletal material, and new footprints from the Upper Permian of the southern Alps (Italy) provide evidence for a diversity not yet sampled by body fossils. The integrative study of body fossil and footprint data supports the hypothesis that archosauriforms had already undergone substantial taxonomic diversification by the Late Permian and that by the Early Triassic archosauromorphs attained a broad geographical distribution over most parts of Pangea. Analysis of body size, as deduced from track size, suggests that archosauriform average body size did not change significantly from the Late Permian to the Early Triassic. A survey of facies yielding both skeletal and track record indicate an ecological preference for inland fluvial (lacustrine) environments for early archosauromorphs. Finally, although more data is needed, Late Permian chirotheriid imprints suggest a shift from sprawling to erect posture in archosauriforms before the end-Permian mass extinction event. We highlight the importance of approaching palaeobiological questions by using all available sources of data, specifically through integrating the body and track fossil record. PMID:26083612

  5. Structure and function of a compound eye, more than half a billion years old.

    PubMed

    Schoenemann, Brigitte; Pärnaste, Helje; Clarkson, Euan N K

    2017-12-19

    Until now, the fossil record has not been capable of revealing any details of the mechanisms of complex vision at the beginning of metazoan evolution. Here, we describe functional units, at a cellular level, of a compound eye from the base of the Cambrian, more than half a billion years old. Remains of early Cambrian arthropods showed the external lattices of enormous compound eyes, but not the internal structures or anything about how those compound eyes may have functioned. In a phosphatized trilobite eye from the lower Cambrian of the Baltic, we found lithified remnants of cellular systems, typical of a modern focal apposition eye, similar to those of a bee or dragonfly. This shows that sophisticated eyes already existed at the beginning of the fossil record of higher organisms, while the differences between the ancient system and the internal structures of a modern apposition compound eye open important insights into the evolution of vision. Copyright © 2017 the Author(s). Published by PNAS.

  6. Detrital Record of Phanerozoic Tectonics in Iran: Evidence From U-Pb Zircon Geochronology

    NASA Astrophysics Data System (ADS)

    Horton, B. K.; Gillis, R. J.; Stockli, D. F.; Hassanzadeh, J.; Axen, G. J.; Grove, M.

    2004-12-01

    Ion-microprobe U-Pb ages of 91 detrital zircon grains supplement ongoing investigations of the tectonic history of Iran, a critical region bridging the gap between the Alpine and Himalayan orogenic belts. These data improve understanding of the distribution of continental blocks during a complex history of Late Proterozoic (Pan-African) crustal growth, Paleozoic passive-margin sedimentation, early Mesozoic collision with Eurasia, and Cenozoic collision with Arabia. U-Pb analyses of detrital zircon grains from four sandstone samples (two Lower Cambrian, one uppermost Triassic-Lower Jurassic, one Neogene) collected from the Alborz mountains of northern Iran reveal a spectrum of ages ranging from 50 to 2900 Ma. Most analyses yield concordant to moderately discordant ages. The Lower Cambrian Lalun and Barut sandstones yield age distribution peaks at approximately 550-650, 1000, and 2500 Ma, consistent with a Gondwanan source area presently to the south and west in parts of Iran and the Arabian-Nubian shield (Saudi Arabia and northwestern Africa). The uppermost Triassic-Lower Jurassic Shemshak Formation exhibits a broad range of U-Pb ages, including peaks of approximately 200-260, 330, 430, 600, and 1900 Ma, requiring a Eurasian source area presently to the north and east in the Turan plate (Turkmenistan and southwestern Asia). Neogene strata display both the youngest and oldest ages (approximately 50 and 2900 Ma) of any samples, a result of substantial sedimentary recycling of older Phanerozoic cover rocks. Because the youngest zircon ages for three of the four samples are indistinguishable from their stratigraphic (depositional) ages, these data suggest rapid exhumation and help constrain the termination age of Late Proterozoic-Early Cambrian (Pan-African) orogenesis and the timing of the Iran-Eurasia collision.

  7. Assembling and disassembling california: A zircon and monazite geochronologic framework for proterozoic crustal evolution in southern California

    USGS Publications Warehouse

    Barth, A.P.; Wooden, J.L.; Coleman, D.S.; Vogel, M.B.

    2009-01-01

    The Mojave province in southern California preserves a comparatively complete record of assembly, postorogenic sedimentation, and rifting along the southwestern North American continental margin. The oldest exposed rocks are metasedimentary gneisses and amphibolite, enclosing intrusive suites that range from tonalite and quartz mon-zodiorite to granite with minor trondhjemite. Discrete magmatic episodes occurred at approximately 1790-1730 and 1690-1640 Ma. Evidence from detrital and premagmatic zircons indicates that recycling of 1900-1790 Ma Paleopro-terozoic crust formed the unique isotopic character of the Mojave province. Peak metamorphic conditions in the Mojave province reached middle amphibolite to granulite facies; metamorphism occurred locally from 1795 to 1640 Ma, with widespread evidence for metamorphism at 1711-1689 and 1670-1650 Ma. Structures record early, tight to isoclinal folding and penetrative west-vergent shear during the final metamorphic event in the west Mojave province. Proterozoic basement rocks are overlain by siliciclastic-carbonate sequences of Mesoproterozoic, Neoproterozoic, and Cambrian age, recording environmental change over the course of the transition from stable Mojave crust to the rifted Cordilleran margin. Neoproterozoic quartzites have diverse zircon populations inconsistent with a southwest North American source, which we infer were derived from the western conjugate rift pair within Rodinia, before establishment of the miogeocline. Neoproterozoic-Cambrian miogeoclinal clastic rocks record an end to rifting and establishment of the Cordilleran miogeocline in southern California by latest Neoproterozoic to Early Cambrian time. ?? 2009 by The University of Chicago.

  8. The Late Cambrian SPICE (δ13C) event and the Sauk II-Sauk III regression: new evidence from Laurentian basins in Utah, Iowa, and Newfoundland

    USGS Publications Warehouse

    Saltzman, Matthew R.; Cowan, Clinton A.; Runkel, Anthony C.; Runnegar, Bruce; Stewart, Michael C.; Palmer, Allison R.

    2004-01-01

    Carbon isotope data from Upper Cambrian sections in three Laurentian basins in northern Utah, central Iowa, and western Newfoundland record a large positive ??13C excursion (SPICE event) of up to + 5???. Peak ??13C ratios are well dated by trilobite collections to the middle of the Steptoean Stage (Dunderbergia Zone) and occur during maximum regression associated with formation of the Sauk II-Sauk III subsequence boundary on the North American craton. Maximum regression was marked by an influx of quartz sand into carbonate-platform settings in all three widely separated basins. In northern Utah, this quartz sand formed a thick sequence known as the Worm Creek Quartzite, which marks a conspicuous interruption of carbonate deposition during the Middle to Late Cambrian in the region. In western Newfoundland, the thickness of the quartz sand unit is much reduced but still marks a brief shutdown of the carbonate factory that is unique to the Cambrian shelf succession of the area. In the central Iowa area of the cratonic interior, an upward-shallowing carbonate succession culminates in cross-stratified trilobite grainstones at the peak of the SPICE in Dunderbergia Zone time, and the lowest point on the relative-sea-level curve is associated with the occurrence of coarse quartz sand derived from the encroaching shoreface. Although it is difficult to determine precisely the departure from baseline ??13C that marks the beginning of the SPICE excursion in the stratigraphic successions analyzed, our results are consistent with a rise and subsequent fall in ??13C tracking a major regressive-transgressive event recorded across northern Laurentia. The correlation of a major ??13C excursion with regression is similar to that described for the Late Ordovician, for which the pattern has been attributed to either increased carbonate relative to terrigenous weathering rates as ice sheets covered up organic-matter-containing silicates at high latitudes or high productivity and organic

  9. Body Size Evolution in Conodonts from the Cambrian through the Triassic

    NASA Astrophysics Data System (ADS)

    Schaal, E. K.; Morgan, D. J.; Payne, J.

    2013-12-01

    The size of an organism exercises tremendous control over its physiology, life history, and ecology, yet the factors that influence body size evolution remain poorly understood. One major limitation is the lack of appropriate datasets spanning long intervals of evolutionary time. Here, we document size trends in conodonts (tooth-like microfossils from marine chordates) because they evolved rapidly and are known to change size during intervals of environmental change. By measuring photographs from the Catalogue of Conodonts (Ziegler 1982), we compiled a database of conodont P1 element measurements for 575 species and subspecies from the Cambrian through Triassic periods. Because tooth size correlates with body size in conodont animals and their extant relatives, conodont element length can serve as a proxy for the size of the conodont animal. We find that mean and maximum size across species increased during the early Paleozoic, peaked during the Devonian-Mississippian, and then generally decreased until conodonts went extinct at the end of the Triassic. We used regression analyses to compare conodont mean size trends to potential environmental predictors, such as changing atmospheric pO2, atmospheric pCO2, and sea level. Conodont size exhibited poor correlation with these environmental factors, suggesting that conodont evolution may have been more strongly influenced by other environmental covariates or ecological variables such as predation and competition.

  10. Proterozoic structure, cambrian rifting, and younger faulting as revealed by a regional seismic reflection network in the Southern Illinois Basin

    USGS Publications Warehouse

    Potter, C.J.; Drahovzal, James A.; Sargent, M.L.; McBride, J.H.

    1997-01-01

    Four high-quality seismic reflection profiles through the southern Illinois Basin, totaling 245 km in length, provide an excellent regional subsurface stratigraphic and structural framework for evaluation of seismic risk, hydrocarbon occurrence, and other regional geologic studies. These data provide extensive subsurface information on the geometry of the intersection of the Cambrian Reelfoot and Rough Creek rifts, on extensive Proterozoic reflection sequences, and on structures (including the Fluorspar Area Fault Complex and Hicks Dome) that underlie a transitional area between the well-defined New Madrid seismic zone (to the southwest) and a more diffuse area of seismicity in the southern Illinois Basin. Our principal interpretations from these data are listed here in order of geologic age, from oldest to youngest: 1. Prominent Proterozoic layering, possibly equivalent to Proterozoic (???1 Ga) Middle Run Formation clastic strata and underlying (1.3-1.5 Ga) volcanic rocks of the East Continent rift basin, has been strongly deformed, probably as part of the Grenville foreland fold and thrust belt. 2. A well-defined angular unconformity is seen in many places between Proterozoic and Cambrian strata; a post-Grenville Proterozoic sequence is also apparent locally, directly beneath the base of the Cambrian. 3. We infer a major reversal in Cambrian rift polarity (accommodation zone) in the Rough Creek Graben in western Kentucky. 4. Seismic facies analysis suggests the presence of basin-floor fan complexes at and near the base of the Cambrian interval and within parts of a Proterozoic post-Grenville sequence in several parts of the Rough Creek Graben. 5. There is an abrupt pinchout of the Mount Simon Sandstone against crystalline basement beneath the Dale Dome (near the Texaco no. 1 Cuppy well, Hamilton County) in southeastern Illinois, and a more gradual Mount Simon pinchout to the southeast. 6. Where crossed by the seismic reflection line in southeast Illinois, some

  11. Controls on Gut Phosphatisation: The Trilobites from the Weeks Formation Lagerstätte (Cambrian; Utah)

    PubMed Central

    Lerosey-Aubril, Rudy; Hegna, Thomas A.; Kier, Carlo; Bonino, Enrico; Habersetzer, Jörg; Carré, Matthieu

    2012-01-01

    Despite being internal organs, digestive structures are frequently preserved in Cambrian Lagerstätten. However, the reasons for their fossilisation and their biological implications remain to be thoroughly explored. This is particularly true with arthropods – typically the most diverse fossilised organisms in Cambrian ecosystems – where digestive structures represent an as-yet underexploited alternative to appendage morphology for inferences on their biology. Here we describe the phosphatised digestive structures of three trilobite species from the Cambrian Weeks Formation Lagerstätte (Utah). Their exquisite, three-dimensional preservation reveals unique details on trilobite internal anatomy, such as the position of the mouth and the absence of a differentiated crop. In addition, the presence of paired pygidial organs of an unknown function is reported for the first time. This exceptional material enables exploration of the relationships between gut phosphatisation and the biology of organisms. Indeed, soft-tissue preservation is unusual in these fossils as it is restricted to the digestive structures, which indicates that the gut played a central role in its own phosphatisation. We hypothesize that the gut provided a microenvironment where special conditions could develop and harboured a source of phosphorus. The fact that gut phosphatization has almost exclusively been observed in arthropods could be explained by their uncommon ability to store ions (including phosphorous) in their digestive tissues. However, in some specimens from the Weeks Formation, the phosphatisation extends to the entire digestive system, suggesting that trilobites might have had some biological particularities not observed in modern arthropods. We speculate that one of them might have been an increased capacity for ion storage in the gut tissues, related to the moulting of their heavily-mineralised carapace. PMID:22431989

  12. The oldest known priapulid-like scalidophoran animal and its implications for the early evolution of cycloneuralians and ecdysozoans.

    PubMed

    Liu, Yunhuan; Xiao, Shuhai; Shao, Tiequan; Broce, Jesse; Zhang, Huaqiao

    2014-05-01

    Morphological phylogenetic analyses suggest that scalidophorans (priapulids, loriciferans, and kinorhynchs) and nematoids (nematodes and nematomorphs) form the ecdysozoan clade Cycloneuralia, which is a sister group to panarthropods. It has been proposed that extant priapulids and Cambrian priapulid-like scalidophorans, because of their conserved evolution, have the potential to illuminate the ancestral morphology, ecology, and developmental biology of highly derived ecdysozoans such as nematods and arthropods. As such, Cambrian fossils, particularly Markuelia and possibly olivooids, can inform the early evolution of scalidophorans, cycloneuralians, and ecdysozoans. However, the scalidophoran Markuelia is known exclusively as embryo fossils, and the olivooids have been alternatively interpreted as cnidarians or cycloneuralians. Here, we describe a post-embryonic scalidophoran fossil Eopriapulites sphinx new genus and species, which represents the oldest known scalidophoran, from the early Cambrian Period (∼535 Ma) in South China. E. sphinx is similar to modern scalidophorans in having an introvert armed with hollow scalids, a collar with coronal scalids, and a pharynx with pharyngeal teeth, but its scalids and pharyngeal teeth are arranged in a hexaradial pattern. Phylogenetically resolved as a stem-group scalidophoran, E. sphinx shares a hexaradial pattern with the hexaradial arrangement of certain anatomical structures in kinorhynchs, loriciferans, nematoids, and Cambrian fossils such as Eolympia pediculata, which could also be a scalidophoran. Thus, the bodyplan of ancestral cycloneuralians may have had a component of hexaradial symmetry (i.e., some but not necessarily all anatomical parts are hexaradially arranged). If panarthropods are nested within paraphyletic cycloneuralians, as several molecular phylogenetic analyses suggest, the ancestral ecdysozoans may have been a legless worm possibly with a component of hexaradial symmetry. © 2014 Wiley

  13. Cadomian volcanosedimentary complexes across the Ediacaran-Cambrian transition of the Eastern Pyrenees, southwestern Europe

    NASA Astrophysics Data System (ADS)

    Padel, Maxime; Álvaro, J. Javier; Casas, Josep Maria; Clausen, Sébastien; Poujol, Marc; Sánchez-García, Teresa

    2017-11-01

    The volcanism hosted by the Ediacaran-Terreneuvian Canaveilles Group of the Eastern Pyrenees displays two distinct geochemical affinities: (1) metabasites of the Nyer and Olette formations reflect the emplacement of a tholeiitic magmatism linked to extensional conditions, whereas (2) subsequent felsic and calc-alkaline magmatic rocks marking the top of the Olette Formation and forming the overlying Fabert and Finestrelles members represent Cadomian magmatic events. Based on U-Pb zircon dating constraints, palaeotopographic relationships linked to onlap geometries and distance from vent sources, three volcanosedimentary edifices can be distinguished, the so-called Tregurà (ca. 565-552 Ma), Cap de Creus (ca. 558 Ma) and Coll d'Ares (ca. 542-532 Ma) edifices. The top of their palaeoreliefs recorded locally the nucleation of centres of microbial carbonate productivity (Puig Sec Member) linked to synsedimentary tilting and karstification. Throughout West Gondwana, the presence of carbonate production across the Ediacaran-Cambrian transition is exclusively located in back-arc settings (Central-Iberian Zone) and areas far from the Cadomian subduction trench and devoid of significant terrigenous input, such as those reported in the Eastern Pyrenees and the neighbouring Montagne Noire.

  14. Reactive oxygen species may play an essential role in driving biological evolution: The Cambrian Explosion as an example.

    PubMed

    Yang, Dong; Guo, Xuejun; Xie, Tian; Luo, Xiaoyan

    2018-01-01

    The Cambrian Explosion is one of the most significant events in the history of life; essentially all easily fossilizable animal body plans first evolved during this event. Although many theories have been proposed to explain this event, its cause remains unresolved. Here, we propose that the elevated level of oxygen, in combination with the increased mobility and food intake of metazoans, led to increased cellular levels of reactive oxygen species (ROS), which drove evolution by enhancing mutation rates and providing new regulatory mechanisms. Our hypothesis may provide a unified explanation for the Cambrian Explosion as it incorporates both environmental and developmental factors and is also consistent with ecological explanations for animal radiation. Future studies should focus on testing this hypothesis, and may lead to important insights into evolution. Copyright © 2017. Published by Elsevier B.V.

  15. The Early Cretaceous Sulfur Isotope Record: New Data, Revised Ages, and Updated Modeling

    NASA Astrophysics Data System (ADS)

    Kristall, B.; Hurtgen, M.; Sageman, B. B.; Jacobson, A. D.

    2015-12-01

    The Early Cretaceous is a time of significant transformation with the continued break-up of Pangea, the emplacement of several LIPs, and a climatic shift from a cool greenhouse to a warm greenhouse. The timing of these major events and their relationship to seawater geochemistry (as recorded in isotope records) is critical for understanding changes in global biogeochemical cycles during this time. Within this context, recent revisions to the Cretaceous portion of the geologic timescale necessitate a reevaluation of the Cretaceous S isotope record as recorded in marine barite (Paytan et al., 2004). We present a revised Early Cretaceous S isotope record and present new δ34Sbarite data that extend the record further back in time and provide more detail during two major S isotope shifts of the Early Cretaceous. The new data maintain the major ~5‰ negative shift but raise questions on the timing and structure of this perturbation. Furthermore, recently updated estimates for global rates of marine microbial sulfate reduction (MSR) (Bowles et al., 2014) and sulfate burial during the Phanerozoic (Halevy et al., 2012) require notable revisions in the fluxes and isotopic values used to model the global S cycle. We present a revised global S cycle box model and reconstruct the evolution of the Early Cretaceous S isotope record primarily through perturbations in volcanic and hydrothermal fluxes (e.g., submarine LIPs). Changes to the weathering and pyrite burial fluxes and the global integrated fractionation factor for MSR are also used to modulate, balance, and smooth the LIP-driven perturbation. The massive evaporite burial during the Late Aptian post dates the major -5‰ shift and has little affect on the modeled S isotope composition of seawater sulfate, despite causing a major drop in sulfate concentration. The S cycle box model is coupled to a Sr cycle box model to provide additional constraints on the magnitude and timing of perturbations within the S isotope record.

  16. Paleoproterozoic mojaveprovince in northwestern Mexico? Isotopic and U-Pb zircon geochronologic studies of precambrian and Cambrian crystalline and sedimentary rocks, Caborca, Sonora

    USGS Publications Warehouse

    Lang, Farmer G.; Bowring, S.A.; Matzel, J.; Maldonado, G.E.; Fedo, C.; Wooden, J.

    2005-01-01

    Whole-rock Nd isotopic data and U-Pb zircon geochronology from Precambrian crystalline rocks in the Caborca area, northern Sonora, reveal that these rocks are most likely a segment of the Paleoproterozoic Mojave province. Supporting this conclusion are the observations that paragneiss from the ??? 1.75 Ga Bamori Complex has a 2.4 Ga Nd model age and contains detrital zircons ranging in age from Paleo- proterozoic (1.75 Ga) to Archean (3.2 Ga). Paragneisses with similar age and isotopic characteristics occur in the Mojave province in southern California. In addition, "A-type" granite exposed at the southern end of Cerro Rajon has ca 2.0 Ga Nd model age and a U-Pb zircon age of 1.71 Ga, which are similar to those of Paleoproterozoic granites in the Mojave province. Unlike the U.S. Mojave province, the Caborcan crust contains ca. 1.1 Ga granite (Aibo Granite), which our new Nd isotopic data suggest is largely the product of anatexis of the local Precambrian basement. Detrital zircons from Neoproterozoic to early Cambrian miogeoclinal arenites at Caborca show dominant populations ca. 1.7 Ga, ca. 1.4 Ga, and ca. 1.1 Ga, with subordinate Early Cambrian and Archean zircons. These zircons were likely derived predominately from North American crust to the east and northeast, and not from the underlying Caborcan basement. The general age and isotopic similarities between Mojave province basement and overlying miogeoclinal sedimentary rocks in Sonora and southern California is necessary, but not sufficient, proof of the hypothesis that Sonoran crust is allochthonous and was transported to its current position during the Mesozoic along the proposed Mojave-Sonora megashear. One viable alternative model is that the Caborcan Precambrian crust is an isolated, autochthonous segment of Mojave province crust that shares a similar, but not identical, Proterozoic geological history with Mojave province crust found in the southwest United States ?? 2005 Geological Society of America.

  17. Evidence of the coupled geology system and its impact on the evolution of South China during the transformation from Sinian to Cambrian

    NASA Astrophysics Data System (ADS)

    Yang, C.; Wang, T.; Chen, Z.

    2016-12-01

    Separate interpretation of the evidence on tectonic, sedimentology or climate is insufficient to reappear the dynamic process of the evolution of the Earth surface, thus, tectonic, sedimentology and climate should be considered as a coupled system. Thick carbonate succession is overlaying on the paleo-uplift which is divided into two parts by a fluted belt in the center of Sichuan Basin. Sinian carbonate rocks is commonly composed by algae dolomite, while at the top of the Sinian succession the rocks had experienced meteoric karstification. Grain dolostones, fine-grained siliciclastic sandstones with mudstone appeare as the regional sediment of Cambrian. However, extraordinary thick mudstone had settled in the fluted belt, and the succession could be divided in to siliciclastic mud of the lower and clay-carbonate mud of the upper. The geochemistry and well log synthesized profile of Z4 well indicate that the chemical condition of siliciclastic mud and clay-carbonate mud had changed from oxidation to reduction, however the siliciclastic mud only appeared within the fluted belt. The fluted belt does not exist on the map of the gravity anomaly, but it had been convinced by the seismic data. The precursor of the fluted belt might be a sag within the platform basement, while with the sea level gradually raising up, the growth of algal mound exacerbated the geomorphology difference. Then a regression had happened at the end of the Sinian, starved all the algae and caused weather crust. Meanwhile, the fluted belt became a closed lagoon, received the sediment including algal mound fragment and biosilic crystals. Afterwards, rapidly increasing of the sea level deposited thick cay-carbonate mud that could be recognized as the sediment of maximum flooding surface. Then with the sea level decreasing, siliciclastic sandstones and inorganic grain carbonate became the main petrology of the Cambrian strata. Fine-grained eolian siliciclastic sandstones within the Cambrian carbonate

  18. Composition and origin of Early Cambrian Tiantaishan phosphorite-Mn carbonate ores, Shaanxi Province, China

    USGS Publications Warehouse

    Hein, J.R.; Fan, D.; Ye, J.; Liu, T.; Yeh, H.-W.

    1999-01-01

    The Tiantaishan phosphorite-Mn carbonate ores occur in the Early Cambrian Tananpo Formation in complexly folded and faulted rocks located in southern Shaanxi Province. About 65 x 106 tonnes of 17% P2O5 ore reserves exist and Mn-ore reserves are about 8.3 x 106 tonnes of +18% Mn. The stratigraphic sequence in ascending order consists of black phyllite, black to gray phosphorite ore, black phyllite, rhodochrostone ore, Mn mixed-carbonates, and dolostone. Data are presented from microprobe mineral chemistry, whole-rock chemistry, stable isotopes of carbonates, X-ray mineralogy, petrographic and SEM observations, and statistical analysis of chemical data. The dominant ore-forming minerals are hydroxy- and carbonate fluorapatite and Ca rhodochrosite, with Mg kutnahorite and dolomite comprising the Mn mixed-carbonate section. Pyrite occurs in all rock types and alabandite (MnS) occurs throughout the rhodochrostone section. The mean P2O5 content of phosphorite is 31% and argillaceous phosphorite is 16%, while the mean MnO content of rhodochrostone ore is 37%. Phosphorite ores are massive, spheroidal, laminated, and banded, while rhodochrostone ores have oolitic, spheroidal, and granular fabrics. The most distinguishing characteristics of the ores are high total organic carbon (TOC) contents (mean 8.4%) in the phosphorite and high P2O5 contents (mean 2.7%) in the rhodochrostone ore. The atypically high TOC contents in the Tiantaishan phosphorite probably result from very strong productivity leading to high sedimentation rates accompanied by weak reworking of sediments; poor utilization of the organic matter by bacteria; and/or partial replacement of bacterial or algal mats by the apatite. The depositional setting of the ores was the margin of an epicontinental seaway created as a direct consequence of global processes that included break-up of a supercontinent, formation of narrow seaways, creation of extensive continental shelves, overturn of stagnant, metal-rich deep

  19. Geochronology and geochemistry of early Paleozoic igneous rocks of the Lesser Xing'an Range, NE China: Implications for the tectonic evolution of the eastern Central Asian Orogenic Belt

    NASA Astrophysics Data System (ADS)

    Wang, Zhi-wei; Xu, Wen-liang; Pei, Fu-ping; Wang, Feng; Guo, Peng

    2016-09-01

    This paper presents new zircon U-Pb, Hf isotope, and whole-rock major and trace element data for early Paleozoic igneous rocks of the Lesser Xing'an Range, NE China, in order to constrain the early Paleozoic tectonic evolution of the eastern Central Asian Orogenic Belt (CAOB). Zircon U-Pb dating indicates that early Paleozoic magmatic events within the northern Songnen-Zhangguangcai Range Massif (SZM) can be subdivided into four stages: Middle Cambrian ( 505 Ma), Late Cambrian ( 490 Ma), Early-Middle Ordovician ( 470 Ma), and Late Ordovician (460-450 Ma). The Middle Cambrian monzogranites are K-rich, weakly to strongly peraluminous, and characterized by pronounced heavy rare earth element (HREE) depletions, high Sr/Y ratios, low Y concentrations, low primary zircon εHf(t) values (- 6.79 to - 1.09), and ancient two-stage model (TDM2) ages (1901-1534 Ma). These results indicate derivation from partial melting of thickened ancient crustal materials that formed during the amalgamation of the northern SZM and the northern Jiamusi Massif (JM). The Late Cambrian monzonite, quartz monzonite, and monzogranite units are chemically similar to A-type granites, and contain zircons with εHf(t) values of - 2.59 to + 1.78 and TDM2 ages of 1625-1348 Ma. We infer that these rocks formed from primary magmas generated by partial melting of Mesoproterozoic accreted lower crustal materials in a post-collisional extensional environment. The Early-Middle Ordovician quartz monzodiorite, quartz monzonite, monzogranite, and rhyolite units are calc-alkaline, relatively enriched in light REEs (LREEs) and large ion lithophile elements (LILEs; e.g., Rb, Th, and U), depleted in HREEs and high field strength elements (HFSEs; e.g., Nb, Ta, and Ti), and contain zircons with εHf(t) values of - 7.33 to + 4.98, indicative of formation in an active continental margin setting. The Late Ordovician alkali-feldspar granite and rhyolite units have A-type granite affinities that suggest they formed in an

  20. Casebooks in early modern England: medicine, astrology, and written records.

    PubMed

    Kassell, Lauren

    2014-01-01

    Casebooks are the richest sources that we have for encounters between early modern medical practitioners and their patients. This article compares astrological and medical records across two centuries, focused on England, and charts developments in the ways in which practitioners kept records and reflected on their practices. Astrologers had a long history of working from particular moments, stellar configurations, and events to general rules. These practices required systematic notation. Physicians increasingly modeled themselves on Hippocrates, recording details of cases as the basis for reasoned expositions of the histories of disease. Medical records, as other scholars have demonstrated, shaped the production of medical knowledge. Instead, this article focuses on the nature of casebooks as artifacts of the medical encounter. It establishes that casebooks were serial records of practice, akin to diaries, testimonials, and registers; identifies extant English casebooks and the practices that led to their production and preservation; and concludes that the processes of writing, ordering, and preserving medical records are as important for understanding the medical encounter as the records themselves.

  1. Microbial involvement in the formation of Cambrian sea-floor silica-iron oxide deposits, Australia

    NASA Astrophysics Data System (ADS)

    Duhig, Nathan C.; Davidson, Garry J.; Stolz, Joe

    1992-06-01

    The Cambrian-Ordovician Mount Windsor volcanic belt in northern Australia is host to stratiform lenses of massive ferruginous chert that are spatially associated with volcanogenic massive sulfide occurrences, in particular the Thalanga zinc-lead-copper-silver deposit. The rocks are composed principally of Fe2O3 and SiO2, with very low concentrations of alkalic elements, and lithogenous elements such as Al, Zr, and Ti; they are interpreted as nearly pure chemical sediments. Textural evidence is documented of the integral role of filamentous bacteria (and/or fungi) in depositing iron from hydrothermal fluids, and of the inorganic precipitation of silica-iron-oxyhydroxide gels that subsequently matured to subcrystalline and crystalline silica forms. At least three distinct iron-accumulating microbial forms are distinguished: networks of septate filaments, nonseptate filament networks, and extremely coarse branching filaments that do not reconnect. Values for δ34S in disseminated pyrite are up to 50‰ lighter than those of contemporaneous Cambrian seawater, suggesting postdepositional colonization of some ironstones by sulfur-reducing bacteria. The site not only preserves the textural interplay of biological and inorganic depositional processes in exhalites, but also extends the oldest known instance of microbial mediation in vent-proximal hydrothermal iron precipitation to at least 500 Ma.

  2. The evolution of early vertebrate photoreceptors.

    PubMed

    Collin, Shaun P; Davies, Wayne L; Hart, Nathan S; Hunt, David M

    2009-10-12

    Meeting the challenge of sampling an ancient aquatic landscape by the early vertebrates was crucial to their survival and would establish a retinal bauplan to be used by all subsequent vertebrate descendents. Image-forming eyes were under tremendous selection pressure and the ability to identify suitable prey and detect potential predators was thought to be one of the major drivers of speciation in the Early Cambrian. Based on the fossil record, we know that hagfishes, lampreys, holocephalans, elasmobranchs and lungfishes occupy critical stages in vertebrate evolution, having remained relatively unchanged over hundreds of millions of years. Now using extant representatives of these 'living fossils', we are able to piece together the evolution of vertebrate photoreception. While photoreception in hagfishes appears to be based on light detection and controlling circadian rhythms, rather than image formation, the photoreceptors of lampreys fall into five distinct classes and represent a critical stage in the dichotomy of rods and cones. At least four types of retinal cones sample the visual environment in lampreys mediating photopic (and potentially colour) vision, a sampling strategy retained by lungfishes, some modern teleosts, reptiles and birds. Trichromacy is retained in cartilaginous fishes (at least in batoids and holocephalans), where it is predicted that true scotopic (dim light) vision evolved in the common ancestor of all living gnathostomes. The capacity to discriminate colour and balance the tradeoff between resolution and sensitivity in the early vertebrates was an important driver of eye evolution, where many of the ocular features evolved were retained as vertebrates progressed on to land.

  3. Self-recognition and Ca2+-dependent carbohydrate-carbohydrate cell adhesion provide clues to the cambrian explosion.

    PubMed

    Fernàndez-Busquets, Xavier; Körnig, André; Bucior, Iwona; Burger, Max M; Anselmetti, Dario

    2009-11-01

    The Cambrian explosion of life was a relatively short period approximately 540 Ma that marked a generalized acceleration in the evolution of most animal phyla, but the trigger of this key biological event remains elusive. Sponges are the oldest extant Precambrian metazoan phylum and thus a valid model to study factors that could have unleashed the rise of multicellular animals. One such factor is the advent of self-/non-self-recognition systems, which would be evolutionarily beneficial to organisms to prevent germ-cell parasitism or the introduction of deleterious mutations resulting from fusion with genetically different individuals. However, the molecules responsible for allorecognition probably evolved gradually before the Cambrian period, and some other (external) factor remains to be identified as the missing triggering event. Sponge cells associate through calcium-dependent, multivalent carbohydrate-carbohydrate interactions of the g200 glycan found on extracellular proteoglycans. Single molecule force spectroscopy analysis of g200-g200 binding indicates that calcium affects the lifetime (+Ca/-Ca: 680 s/3 s) and bond reaction length (+Ca/-Ca: 3.47 A/2.27 A). Calculation of mean g200 dissociation times in low and high calcium within the theoretical framework of a cooperative binding model indicates the nonlinear and divergent characteristics leading to either disaggregated cells or stable multicellular assemblies, respectively. This fundamental phenomenon can explain a switch from weak to strong adhesion between primitive metazoan cells caused by the well-documented rise in ocean calcium levels at the end of Precambrian time. We propose that stronger cell adhesion allowed the integrity of genetically uniform animals composed only of "self" cells, facilitating genetic constitutions to remain within the metazoan individual and be passed down inheritance lines. The Cambrian explosion might have been triggered by the coincidence in time of primitive animals

  4. Cambrian archaeocyathan metazoans: revision of morphological characters and standardization of genus descriptions to establish an online identification tool.

    PubMed Central

    Kerner, Adeline; Debrenne, Françoise; Vignes-Lebbe, Régine

    2011-01-01

    Abstract Archaeocyatha represent the oldest calcified sponges and the first metazoans to build bioconstructions in association with calcimicrobes. They are a key group in biology, evolutionary studies, biostratigraphy, paleoecology and paleogeography of the early Cambrian times. The establishing of a new standardized terminology for archaeocyathans description has permitted the creation of the first knowledge base in English including descriptions of all archaeocyathan genera. This base, using the XPER² software package, is an integral part of the -Archaeocyatha- a knowledge base website, freely available at url http://www.infosyslab.fr/archaeocyatha. The website is composed of common information about Archaeocyatha, general remarks about the knowledge base, the description of the 307 genera recognized with images of type-specimens of type-species for each genus, as well as additional morphological data, an interactive free access key and its user guide. The automatic analysis and comparison of the digitized descriptions have identified some genera with highly similar morphology. These results are a great help for future taxonomic revisions and suggest a number of possible synonymies that require further study. PMID:22207818

  5. Continuous estimates on the earthquake early warning magnitude by use of the near-field acceleration records

    NASA Astrophysics Data System (ADS)

    Li, Jun; Jin, Xing; Wei, Yongxiang; Zhang, Hongcai

    2013-10-01

    In this article, the seismic records of Japan's Kik-net are selected to measure the acceleration, displacement, and effective peak acceleration of each seismic record within a certain time after P wave, then a continuous estimation is given on earthquake early warning magnitude through statistical analysis method, and Wenchuan earthquake record is utilized to check the method. The results show that the reliability of earthquake early warning magnitude continuously increases with the increase of the seismic information, the biggest residual happens if the acceleration is adopted to fit earthquake magnitude, which may be caused by rich high-frequency components and large dispersion of peak value in acceleration record, the influence caused by the high-frequency components can be effectively reduced if the effective peak acceleration and peak displacement is adopted, it is estimated that the dispersion of earthquake magnitude obviously reduces, but it is easy for peak displacement to be affected by long-period drifting. In various components, the residual enlargement phenomenon at vertical direction is almost unobvious, thus it is recommended in this article that the effective peak acceleration at vertical direction is preferred to estimate earthquake early warning magnitude. Through adopting Wenchuan strong earthquake record to check the method mentioned in this article, it is found that this method can be used to quickly, stably, and accurately estimate the early warning magnitude of this earthquake, which shows that this method is completely applicable for earthquake early warning.

  6. Characterization of the Lower Cambrian sandstone aquifer in the Swedish Baltic Sea area - assessment regarding its potential suitability for storage of CO2

    NASA Astrophysics Data System (ADS)

    Erlström, M.; Sivhed, U.

    2012-04-01

    In the Baltic region the Cambrian sandstone is considered to have great economic value concerning its aquifer and reservoir properties. Its potential as petroleum reservoir is well known, especially from the Polish, Lithuanian and Russian sectors of the Baltic Sea where oil and gas has been found in anticline traps in the sandstone sequence. Offshore exploration in the Swedish sector has so far not encountered any significant findings of oil and gas. However, the extensive exploration has generated data, which is now being used for assessing the overall properties regarding suitability for storage of CO2. The Swedish primary industry has a great interest in finding potential sites for storage of CO2. A suitable site in the Baltic Sea would be a most favourable alternative in comparison to more remote alternatives such as deep saline aquifers in the North Sea. The Lower Cambrian is in the Swedish sector of the Baltic Sea composed of three main sandstone units varying in thickness between 5 and 50 m occurring within an up to 250 m thick Cambrian sequence dominated by fine-grained terriclastic sediments. The limit of Lower Palaeozoic sequence in the Baltic area is today defined by erosional truncation because of the gently dipping Lower Palaeozoic sequence. To the north and northwest, the limit is found in the Pre-Quaternary, whereas the erosional limit is deeply buried beneath Permian and Mesozoic sediments to the south. Here the Lower Palaeozoic limit is buried to depths reaching more than 2 km. The Cambrian sequence in the distal parts of the Swedish sector occurs at depths of c. 1300 m while it constitutes the bedrock surface in a narrow zone trending from Öland to the north of of Gotland. Sandstone beds constitute 40-60% of the total Cambrian sequence. The main sandstone units have a regional distribution of several thousands of square kilometres. The up to 50 m thick Faludden sandstone member exhibits the best reservoir properties including porosities in the

  7. Exceptionally Preserved Cambrian Trilobite Digestive System Revealed in 3D by Synchrotron-Radiation X-Ray Tomographic Microscopy

    PubMed Central

    Eriksson, Mats E.; Terfelt, Fredrik

    2012-01-01

    The Cambrian ‘Orsten’ fauna comprises exceptionally preserved and phosphatised microscopic arthropods. The external morphology of these fossils is well known, but their internal soft-tissue anatomy has remained virtually unknown. Here, we report the first non-biomineralised tissues from a juvenile polymerid trilobite, represented by digestive structures, glands, and connective strands harboured in a hypostome from the Swedish ‘Orsten’ fauna. Synchrotron-radiation X-ray tomographic microscopy enabled three-dimensional internal recordings at sub-micrometre resolution. The specimen provides the first unambiguous evidence for a J-shaped anterior gut and the presence of a crop with a constricted alimentary tract in the Trilobita. Moreover, the gut is Y-shaped in cross section, probably due to a collapsed lumen of that shape, another feature which has not previously been observed in trilobites. The combination of anatomical features suggests that the trilobite hypostome is functionally analogous to the labrum of euarthropods and that it was a sophisticated element closely integrated with the digestive system. This study also briefly addresses the preservational bias of the ‘Orsten’ fauna, particularly the near-absence of polymerid trilobites, and the taphonomy of the soft-tissue-harbouring hypostome. PMID:22558180

  8. Exceptionally preserved Cambrian trilobite digestive system revealed in 3D by synchrotron-radiation X-ray tomographic microscopy.

    PubMed

    Eriksson, Mats E; Terfelt, Fredrik

    2012-01-01

    The Cambrian 'Orsten' fauna comprises exceptionally preserved and phosphatised microscopic arthropods. The external morphology of these fossils is well known, but their internal soft-tissue anatomy has remained virtually unknown. Here, we report the first non-biomineralised tissues from a juvenile polymerid trilobite, represented by digestive structures, glands, and connective strands harboured in a hypostome from the Swedish 'Orsten' fauna. Synchrotron-radiation X-ray tomographic microscopy enabled three-dimensional internal recordings at sub-micrometre resolution. The specimen provides the first unambiguous evidence for a J-shaped anterior gut and the presence of a crop with a constricted alimentary tract in the Trilobita. Moreover, the gut is Y-shaped in cross section, probably due to a collapsed lumen of that shape, another feature which has not previously been observed in trilobites. The combination of anatomical features suggests that the trilobite hypostome is functionally analogous to the labrum of euarthropods and that it was a sophisticated element closely integrated with the digestive system. This study also briefly addresses the preservational bias of the 'Orsten' fauna, particularly the near-absence of polymerid trilobites, and the taphonomy of the soft-tissue-harbouring hypostome.

  9. Biostratigraphy and chronostratigraphy of the Cambrian-Ordovician great American carbonate bank

    USGS Publications Warehouse

    Taylor, John F.; Repetski, John E.; Loch, James D.; Leslie, Stephen A.

    2012-01-01

    The carbonate strata of the great American carbonate bank (GACB) have been subdivided and correlated with ever-increasing precision and accuracy during the past half century through use of the dominant organisms that evolved on the Laurentian platform through the Cambrian and the Ordovician. Trilobites and conodonts remain the primary groups used for this purpose, although brachiopods, both calcareous and phosphatic, and graptolites are very important in certain facies and intervals. A series of charts show the chronostratigraphic units (series and stages) currently in use for deposits of the GACB and the biostratigraphic units (zones, subzones, and biomeres) whose boundaries delineate them. Older and, in some cases obsolete, stages and faunal units are included in the figures to allow users to relate information from previous publications and/or industry databases to modern units. This chapter also provides a brief discussion on the use of biostratigraphy in the recognition and interregional correlation of supersequence boundaries within the Sauk and Tippecanoe megasequences, and the varied perspectives on the nature of biostratigraphic units and their defining taxa during the past half century. Also included are a concise update on the biomere concept, and an explanation of the biostratigraphic consequences of a profound change in the dynamics of extinction and replacement that occurred on the GACB in the Early Ordovician when the factors responsible for platformwide biomere-type extinctions faded and ultimately disappeared. A final section addresses recent and pending refinements in the genus and species taxonomy of biostratigraphically significant fossil groups, the potential they hold for greatly improved correlation, and the obstacles to be overcome for that potential to be realized.

  10. Early to mid-Miocene palaeoclimate of Antarctica based on terrestrial records

    NASA Astrophysics Data System (ADS)

    Ashworth, Allan; Lewis, Adam

    2017-04-01

    Paleontological and stratigraphic studies of sites in the Transantarctic Mountains (TAM) are advancing knowledge of the landscape, vegetation and climate that existed immediately before the growth of the modern East Antarctic Ice Sheet. The sites are located in the Friis Hills and the western Olympus Range in the McMurdo Dry Valleys. In both localities, parts of ancient landscapes are preserved on upland surfaces high above modern valley floors. The early to mid-Miocene interval is bracketed by 40Ar/39Ar ages on volcanic ashes of 19.76 ± 0.11 Ma to 13.85 ± 0.03 Ma. Like all glacial records it is discontinuous but even so several trends can be detected. The record is one of an evolving glacial system during which ice caps coalesced to form an ice sheet. Initially, small alpine glaciers flowed southwestward toward the continental interior eroding shallow troughs into granitic bedrock. By the close of the interval, large glaciers flowed eastward from the continental interior to the Ross Sea. The interval was marked by numerous glacial advances and retreats. Tills are matrix-rich, and outwash sands and gravels ripple-laminated and cross-bedded, typical of those associated with wet-based glaciation. The vegetation during the interval was in a dynamic flux retreating downslope during glacial advances and recolonizing valleys after retreats. Fossils accumulated in peat beds and organic silts representing lacustrine, fluvial and paludal environments. Fossils include diatoms, fungal ascomycetes, pollen and spores, lycopod megaspores, mosses, wood and leaves of Nothofagus (southern beech), fruits of vascular plants, and insect skeletal parts of Diptera (flies) and Coleoptera (beetles). The vegetation was a tundra, initially shrub- and later moss-dominated. During the interval there was a marked decline in biodiversity. Initially, there were 4 species of Nothofagus represented by different leaf types and at least 9 species of vascular plants by their seeds. At the close of

  11. Three Cambrian fossils assembled into an extinct body plan of cnidarian affinity.

    PubMed

    Ou, Qiang; Han, Jian; Zhang, Zhifei; Shu, Degan; Sun, Ge; Mayer, Georg

    2017-08-15

    The early Cambrian problematica Xianguangia sinica , Chengjiangopenna wangii , and Galeaplumosus abilus from the Chengjiang biota (Yunnan, China) have caused much controversy in the past and their phylogenetic placements remain unresolved. Here we show, based on exceptionally preserved material (85 new specimens plus type material), that specimens previously assigned to these three species are in fact parts of the same organism and propose that C. wangii and G. abilus are junior synonyms of X. sinica Our reconstruction of the complete animal reveals an extinct body plan that combines the characteristics of the three described species and is distinct from all known fossil and living taxa. This animal resembled a cnidarian polyp in overall morphology and having a gastric cavity partitioned by septum-like structures. However, it possessed an additional body cavity within its holdfast, an anchoring pit on the basal disk, and feather-like tentacles with densely ciliated pinnules arranged in an alternating pattern, indicating that it was a suspension feeder rather than a predatory actiniarian. Phylogenetic analyses using Bayesian inference and maximum parsimony suggest that X. sinica is a stem-group cnidarian. This relationship implies that the last common ancestor of X. sinica and crown cnidarians was probably a benthic, polypoid animal with a partitioned gastric cavity and a single mouth/anus opening. This extinct body plan suggests that feeding strategies of stem cnidarians may have been drastically different from that of their crown relatives, which are almost exclusively predators, and reveals that the morphological disparity of total-group Cnidaria is greater than previously assumed.

  12. Three Cambrian fossils assembled into an extinct body plan of cnidarian affinity

    PubMed Central

    Ou, Qiang; Han, Jian; Zhang, Zhifei; Shu, Degan; Sun, Ge; Mayer, Georg

    2017-01-01

    The early Cambrian problematica Xianguangia sinica, Chengjiangopenna wangii, and Galeaplumosus abilus from the Chengjiang biota (Yunnan, China) have caused much controversy in the past and their phylogenetic placements remain unresolved. Here we show, based on exceptionally preserved material (85 new specimens plus type material), that specimens previously assigned to these three species are in fact parts of the same organism and propose that C. wangii and G. abilus are junior synonyms of X. sinica. Our reconstruction of the complete animal reveals an extinct body plan that combines the characteristics of the three described species and is distinct from all known fossil and living taxa. This animal resembled a cnidarian polyp in overall morphology and having a gastric cavity partitioned by septum-like structures. However, it possessed an additional body cavity within its holdfast, an anchoring pit on the basal disk, and feather-like tentacles with densely ciliated pinnules arranged in an alternating pattern, indicating that it was a suspension feeder rather than a predatory actiniarian. Phylogenetic analyses using Bayesian inference and maximum parsimony suggest that X. sinica is a stem-group cnidarian. This relationship implies that the last common ancestor of X. sinica and crown cnidarians was probably a benthic, polypoid animal with a partitioned gastric cavity and a single mouth/anus opening. This extinct body plan suggests that feeding strategies of stem cnidarians may have been drastically different from that of their crown relatives, which are almost exclusively predators, and reveals that the morphological disparity of total-group Cnidaria is greater than previously assumed. PMID:28760981

  13. Origin of northern Gondwana Cambrian sandstone revealed by detrital zircon SHRIMP dating

    USGS Publications Warehouse

    Avigad, D.; Kolodner, K.; McWilliams, M.; Persing, H.; Weissbrod, T.

    2003-01-01

    Voluminous Paleozoic sandstone sequences were deposited in northern Africa and Arabia following an extended Neoproterozoic orogenic cycle that culminated in the assembly of Gondwana. We measured sensitive high-resolution ion microprobe (SHRIMP) U-Pb ages of detrital zircons separated from several Cambrian units in the Elat area of southern Israel in order to unravel their provenance. This sandstone forms the base of the widespread siliciclastic section now exposed on the periphery of the Arabian-Nubian shield in northeastern Africa and Arabia. Most of the detrital zircons we analyzed yielded Neoproterozoic concordant ages with a marked concentration at 0.55–0.65 Ga. The most likely provenance of the Neoproterozoic detritus is the Arabian-Nubian shield; 0.55–0.65 Ga was a time of posttectonic igneous activity, rift-related volcanism, and strike-slip faulting there. Of the zircons, 30% yielded pre-Neoproterozoic ages grouped at 0.9–1.1 Ga (Kibaran), 1.65–1.85 Ga, and 2.45–2.7 Ga. The majority of the pre-Neoproterozoic zircons underwent Pb loss, possibly as a consequence of the Pan-African orogeny resetting their provenance. Rocks of the Saharan metacraton and the southern Afif terrane in Saudi Arabia (∼1000 km south of Elat) are plausible sources of these zircons. Kibaran basement rocks are currently exposed more than 3000 km south of Elat (flanking the Mozambique belt), but the shape of the detrital zircons of that age and the presence of feldspar in the host sandstone are not fully consistent with such a long-distance transport. Reworking of Neoproteorozoic glacial detritus may explain the presence of Kibaran detrital zircons in the Cambrian of Elat, but the possibility that the Arabian-Nubian shield contains Kibaran rocks (hitherto not recognized) should also be explored.

  14. Permian-Early Triassic tectonics and stratigraphy of the Karoo Supergroup in northwestern Mozambique

    NASA Astrophysics Data System (ADS)

    Bicca, Marcos Müller; Philipp, Ruy Paulo; Jelinek, Andrea Ritter; Ketzer, João Marcelo Medina; dos Santos Scherer, Claiton Marlon; Jamal, Daúd Liace; dos Reis, Adriano Domingos

    2017-06-01

    The Gondwana continent was the base of great basin inception, sedimentation and magmatism throughout the Cambrian to Middle Jurassic periods. The northwestern Mozambique igneous and metamorphic basement assemblages host the NW-trending Moatize Minjova Basin, which has great economic potential for coal and gas mining. This rift basin was activated by an S-SW stress field during the Early Permian period, as constrained by regional and field scale structural data. Tectonically induced subsidence in the basin, from the reactivation of NW-SE and NNE-SSW regional structures is well recorded by faults, folds and synsedimentary fractures within the Early Late Permian Moatize Formation. NW-SE, N-S and NE-SW field structures consist of post-Karoo reactivation patterns related to a NNE-SSW extension produced by the Pangea breakup and early inception stages of the Great East African Rift System. The Early Late Permian sequences of the Moatize-Minjova Basin are composed of fluvial meandering, coal-bearing beds of the Moatize Formation, which comprises mostly floodplain, crevasse splay and fluvial channel lithofacies associations, deposited in a cyclic pattern. This sequence was overlapped by a multiple-story, braided fluvial plain sequence of the Matinde Formation (Late Permian - Early Triassic). Lithofacies associations in the Matinde Formation and its internal relationships suggest deposition of poorly channelized braided alluvial plain in which downstream and probably lateral accretion macroforms alternate with gravity flow deposits. NW paleoflow measurements suggest that Permian fluvial headwaters were located somewhere southeast of the study area, possibly between the African and Antarctic Precambrian highlands.

  15. Mechanism for Burgess Shale-type preservation

    PubMed Central

    Gaines, Robert R.; Hammarlund, Emma U.; Hou, Xianguang; Qi, Changshi; Gabbott, Sarah E.; Zhao, Yuanlong; Peng, Jin; Canfield, Donald E.

    2012-01-01

    Exceptionally preserved fossil biotas of the Burgess Shale and a handful of other similar Cambrian deposits provide rare but critical insights into the early diversification of animals. The extraordinary preservation of labile tissues in these geographically widespread but temporally restricted soft-bodied fossil assemblages has remained enigmatic since Walcott’s initial discovery in 1909. Here, we demonstrate the mechanism of Burgess Shale-type preservation using sedimentologic and geochemical data from the Chengjiang, Burgess Shale, and five other principal Burgess Shale-type deposits. Sulfur isotope evidence from sedimentary pyrites reveals that the exquisite fossilization of organic remains as carbonaceous compressions resulted from early inhibition of microbial activity in the sediments by means of oxidant deprivation. Low sulfate concentrations in the global ocean and low-oxygen bottom water conditions at the sites of deposition resulted in reduced oxidant availability. Subsequently, rapid entombment of fossils in fine-grained sediments and early sealing of sediments by pervasive carbonate cements at bed tops restricted oxidant flux into the sediments. A permeability barrier, provided by bed-capping cements that were emplaced at the seafloor, is a feature that is shared among Burgess Shale-type deposits, and resulted from the unusually high alkalinity of Cambrian oceans. Thus, Burgess Shale-type preservation of soft-bodied fossil assemblages worldwide was promoted by unique aspects of early Paleozoic seawater chemistry that strongly impacted sediment diagenesis, providing a fundamentally unique record of the immediate aftermath of the “Cambrian explosion.” PMID:22392974

  16. Cambrian palaeomagnetic data confirm a Natal Embayment location for the Ellsworth-Whitmore Mountains, Antarctica, in Gondwana reconstructions

    NASA Astrophysics Data System (ADS)

    Randall, Darren E.; Mac Niocaill, Conall

    2004-04-01

    The Ellsworth-Whitmore Mountains (EWM) are one of five terranes that form West Antarctica. Constraining the positions of these terranes in pre-break up Gondwana is crucial to understanding the history of the palaeo-Pacific Gondwana margin. We report the results of a detailed palaeomagnetic investigation of the EWM, which comprises some 150 sites in six formations, ranging in age from Cambrian to Permian. Five of the studied units yield only viscous remnant magnetizations of recent age, or unstable natural remanent magnetizations. The remaining unit, the mid-late Cambrian Frazier Ridge Formation, yielded stable magnetizations at 16 of 35 sites. This component passes a fold test at the 95 per cent confidence level, indicating that it pre-dates Permian deformation, and we argue that it is of primary origin. The resulting palaeopole (9°N 293°E A95= 5.1°) is in good agreement with two previously published palaeopoles from similarly aged rocks in the EWM. Collectively these data indicate that the EWM were located in the Natal Embayment prior to Gondwana break-up, and underwent 90° of anticlockwise rotation during break-up. All three studies, however, yield inclinations that are slightly too shallow when compared with coeval Gondwana reference poles.

  17. Detrital zircon geochronology of Neoproterozoic to Middle Cambrian miogeoclinal and platformal strata: Northwest Sonora, Mexico

    USGS Publications Warehouse

    Gross, E.L.; Stewart, John H.; Gehreis, G.E.

    2000-01-01

    Eighty-five detrital zircon grains from Mesoproterozoic and/or Neoproterozoic to Middle Cambrian sedimentary strata in northwest Sonora, Mexico, have been analyzed to determine source terranes and provide limiting depositional ages of the units. The zircon suites from the Mesoproterozoic and/or Neoproterozoic El Alamo Formation and El Aguila unit yield ages between 1.06 Ga and 2.67 Ga, with predominant ages of 1.1 to 1.2 Ga. Zircons from the Lower? and Middle Cambrian Bolsa Quartzite show age groups from 525 Ma to 1.63 Ga, with a dominant population of 1.1 to 1.2 Ga grains. Grains older than 1.2 Ga in the samples were most likely derived from basement terranes and ???1.4 Ga granitic bodies of the southwest U.S. and northwest Mexico. It is also possible that the sediments were transported from the south, although source rocks of the appropriate age are not presently exposed south of the study area in northern Mexico. Three possibilities for the dominant 1.1 to 1.2 Ga grains include derivation from: (I) exposures of the Grenville belt in southern North America, (2) local 1.1-1.2 Ga granite bodies, or (3) a southern source, such as the Oaxaca terrane, that was subsequently rifted away. Sampling of additional units in the western U.S. and northern Mexico may help resolve the ambiguity surrounding the source of the 1.1 to 1.2 Ga grains.

  18. Trace fossils and bioturbation in peritidal facies of the Potsdam-Theresa Formations (Cambrian-Ordovician), Northwest Adirondacks

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Bjerstedt, T.W.; Erickson, J.M.

    The Cambrian-Ordovician Potsdam Sandstone, Theresa Formation, and Canadian correlatives in the St. Lawrence Lowlands preserve tide-dominated facies during the basal Cambrian transgression. Low intertidal sand flats in the upper Potsdam contain a Skolithos Ichnofacies dominated by Diplocraterion parallelum in clean, herringbone cross-bedded sandstones indicative of high tidal current energy. Wind-wave-driven longshore and tidal currents along a macrotidal coastline were funneled northeast-southwest by Precambrian topographic relief of up to 65 m. This relief is now expressed as the Thousand Islands of New York and Canada. The conformably overlying Theresa Formation preserves a shoaling-upward sequence of mixed clastic-carbonate facies. Shallow subtidal andmore » peritidal facies contain a mixed Skolithos-Cruziana Ichnofacies in sharply alternating lithofacies consisting of gray, intensely bioturbated, poorly sorted calcareous sandstone, and meter-thick, white cross-bedded sandstone. The parallelism between ichnofacies and lithofacies indicates that environmental energy level and persistence rather than water depth controlled trace fossil distribution. Bioturbated sandstones contain a Cruziana ichnofacies of abundant deposit feeders including: Fustiglyphus , Gyrochorte , Neonereites uniserialis , Phycodes flabellum, Planolites beverlyensis, Rosselia socialis, and Teichichnus. Suspension feeders are represented by D. habichi, D. parallelum, Skolithos, Monocraterion, and possibly Palaeophycus tubularis. Scavenging or deposit-feeding arthropods are represented by rare Cruziana furrows. Cross-bedded sandstones contain a Skolithos Ichnofacies of shallow Skolithos and Monocraterion burrows, and an undescribed large epistratal eurypterid( ) trail.« less

  19. An early record of ball lightning: Oliva (Spain), 1619

    NASA Astrophysics Data System (ADS)

    Domínguez-Castro, Fernando

    2018-05-01

    In a primary documentary source we found an early record of ball lightning (BL), which was observed in the monastery of Pi (Oliva, southeastern Spain) on 18 October 1619. The ball lightning was observed by at least three people and was described as a rolling burning vessel and a ball of fire. The ball lightning appeared following a lightning flash, showed a mainly horizontal motion, crossed a wall, smudged an image of the Lady of Rebollet (then known as Lady of Pi) and burnt her ruff, and overturned a cross.

  20. The sophisticated visual system of a tiny Cambrian crustacean: analysis of a stalked fossil compound eye

    PubMed Central

    Schoenemann, Brigitte; Castellani, Christopher; Clarkson, Euan N. K.; Haug, Joachim T.; Maas, Andreas; Haug, Carolin; Waloszek, Dieter

    2012-01-01

    Fossilized compound eyes from the Cambrian, isolated and three-dimensionally preserved, provide remarkable insights into the lifestyle and habitat of their owners. The tiny stalked compound eyes described here probably possessed too few facets to form a proper image, but they represent a sophisticated system for detecting moving objects. The eyes are preserved as almost solid, mace-shaped blocks of phosphate, in which the original positions of the rhabdoms in one specimen are retained as deep cavities. Analysis of the optical axes reveals four visual areas, each with different properties in acuity of vision. They are surveyed by lenses directed forwards, laterally, backwards and inwards, respectively. The most intriguing of these is the putatively inwardly orientated zone, where the optical axes, like those orientated to the front, interfere with axes of the other eye of the contralateral side. The result is a three-dimensional visual net that covers not only the front, but extends also far laterally to either side. Thus, a moving object could be perceived by a two-dimensional coordinate (which is formed by two axes of those facets, one of the left and one of the right eye, which are orientated towards the moving object) in a wide three-dimensional space. This compound eye system enables small arthropods equipped with an eye of low acuity to estimate velocity, size or distance of possible food items efficiently. The eyes are interpreted as having been derived from individuals of the early crustacean Henningsmoenicaris scutula pointing to the existence of highly efficiently developed eyes in the early evolutionary lineage leading towards the modern Crustacea. PMID:22048954

  1. Precambrian Time - The Story of the Early Earth

    USGS Publications Warehouse

    Lindsey, D.A.

    2007-01-01

    The Precambrian is the least-understood part of Earth history, yet it is arguably the most important. Precambrian time spans almost nine-tenths of Earth history, from the formation of the Earth to the dawn of the Cambrian Period. It represents time so vast and long ago that it challenges all comprehension. The Precambrian is the time of big questions. How old is the Earth? How old are the oldest rocks and continents? What was the early Earth like? What was the early atmosphere like? When did life appear, and what did it look like? And, how do we know this? In recent years, remarkable progress has been made in understanding the early evolution of the Earth and life itself. Yet, the scientific story of the early Earth is still a work in progress, humankind's latest attempt to understand the planet. Like previous attempts, it too will change as we learn more about the Earth. Read on to discover what we know now, in the early 21st century.

  2. Neoproterozoic-Early Paleozoic Peri-Pacific Accretionary Evolution of the Mongolian Collage System: Insights From Geochemical and U-Pb Zircon Data From the Ordovician Sedimentary Wedge in the Mongolian Altai

    NASA Astrophysics Data System (ADS)

    Jiang, Y. D.; Schulmann, K.; Kröner, A.; Sun, M.; Lexa, O.; Janoušek, V.; Buriánek, D.; Yuan, C.; Hanžl, P.

    2017-11-01

    Neoproterozoic to early Paleozoic accretionary processes of the Central Asian Orogenic Belt have been evaluated so far mainly using the geology of ophiolites and/or magmatic arcs. Thus, the knowledge of the nature and evolution of associated sedimentary prisms remains fragmentary. We carried out an integrated geological, geochemical, and zircon U-Pb geochronological study on a giant Ordovician metasedimentary succession of the Mongolian Altai Mountains. This succession is characterized by dominant terrigenous components mixed with volcanogenic material. It is chemically immature, compositionally analogous to graywacke, and marked by significant input of felsic to intermediate arc components, pointing to an active continental margin depositional setting. Detrital zircon U-Pb ages suggest a source dominated by products of early Paleozoic magmatism prevailing during the Cambrian-Ordovician and culminating at circa 500 Ma. We propose that the Ordovician succession forms an "Altai sedimentary wedge," the evolution of which can be linked to the geodynamics of the margins of the Mongolian Precambrian Zavhan-Baydrag blocks. This involved subduction reversal from southward subduction of a passive continental margin (Early Cambrian) to the development of the "Ikh-Mongol Magmatic Arc System" and the giant Altai sedimentary wedge above a north dipping subduction zone (Late Cambrian-Ordovician). Such a dynamic process resembles the tectonic evolution of the peri-Pacific accretionary Terra Australis Orogen. A new model reconciling the Baikalian metamorphic belt along the southern Siberian Craton with peri-Pacific Altai accretionary systems fringing the Mongolian microcontinents is proposed to explain the Cambro-Ordovician geodynamic evolution of the Mongolian collage system.

  3. Early Life on Earth: the Ancient Fossil Record

    NASA Astrophysics Data System (ADS)

    Westall, F.

    2004-07-01

    The evidence for early life and its initial evolution on Earth is lin= ked intimately with the geological evolution of the early Earth. The environment of the early Earth would be considered extreme by modern standards: hot (50-80=B0C), volcanically and hydrothermally active, a= noxic, high UV flux, and a high flux of extraterrestrial impacts. Habitats = for life were more limited until continent-building processes resulted in= the formation of stable cratons with wide, shallow, continental platforms= in the Mid-Late Archaean. Unfortunately there are no records of the first appearance of life and the earliest isotopic indications of the exist= ence of organisms fractionating carbon in ~3.8 Ga rocks from the Isua greenst= one belt in Greenland are tenuous. Well-preserved microfossils and micro= bial mats (in the form of tabular and domical stromatolites) occur in 3.5-= 3.3 Ga, Early Archaean, sedimentary formations from the Barberton (South Afri= ca) and Pilbara (Australia) greenstone belts. They document life forms that = show a relatively advanced level of evolution. Microfossil morphology inclu= des filamentous, coccoid, rod and vibroid shapes. Colonial microorganism= s formed biofilms and microbial mats at the surfaces of volcaniclastic = and chemical sediments, some of which created (small) macroscopic microbi= alites such as stromatolites. Anoxygenic photosynthesis may already have developed. Carbon, nitrogen and sulphur isotopes ratios are in the r= ange of those for organisms with anaerobic metabolisms, such as methanogenesi= s, sulphate reduction and photosynthesis. Life was apparently distribute= d widely in shallow-water to littoral environments, including exposed, evaporitic basins and regions of hydrothermal activity. Biomass in t= he early Archaean was restricted owing to the limited amount of energy t= hat could be produced by anaerobic metabolisms. Microfossils resembling o= xygenic photosynthesisers, such as cyanobacteria, probably first occurred in

  4. Early to middle Miocene climate evolution: benthic oxygen and carbon isotope records from Walvis Ridge Site 1264.

    NASA Astrophysics Data System (ADS)

    Lourens, L. J.; Beddow, H.; Liebrand, D.; Schrader, C.; Hilgen, F. J.

    2016-12-01

    Across the early to middle Miocene, high-resolution records from the Pacific Ocean indicate a dynamic climate system, encompassing a 2 Myr global warming event from 17 Ma to 14.7 Ma, followed by a major Cenozoic cooling step at 14.2 Ma -13.8 Ma. Currently, no high-resolution benthic record from the Atlantic Ocean exists covering both events, limiting global coverage of this intriguing period in Cenozoic climate evolution. Here, we present the first early to middle Miocene high-resolution from the Atlantic basin. These records, from Site 1264 on the Walvis Ridge, span a 5.5 Myr long interval (13.24-18.90 ma) in high temporal resolution ( 4 kyr) and are tuned to eccentricity. The d18O record shows a sudden (high-latitude) warming/deglaciation on Antarctica at 17.1 Ma, a rapid cooling/glaciation of Antarctica at 13.8 Ma, and high-amplitude ( 1‰) variability on astronomical time-scales throughout this interval. Together with other records from this time interval located in the Pacific, which show similar features, the data strongly suggests a highly dynamic global climate system. We find cooling steps in d18O at 14.7, 14.2 and 13.8 Ma, suggesting concurrent cooling in the Pacific and Atlantic deep waters during the MMCT. The benthic foraminiferal stable isotope records reveal that the dominant astronomical frequencies present at ODP Site 1264 during the early to middle Miocene interval are the 405 kyr and 110 kyr eccentricity periodicities. This is a contrast to other early to middle Miocene records from drill-sites in the Pacific and South China Sea, which show a strong expression of obliquity in particular between 14.2 and 14.7 Ma.

  5. Gene Regulatory Networks, Homology, and the Early Panarthropod Fossil Record.

    PubMed

    Tweedt, Sarah M

    2017-09-01

    The arthropod body plan is widely believed to have derived from an ancestral form resembling Cambrian-aged fossil lobopodians, and interpretations of morphological and molecular data have long favored this hypothesis. It is possible, however, that appendages and other morphologies observed in extinct and living panarthropods evolved independently. The key to distinguishing between morphological homology and homoplasy lies in the study of developmental gene regulatory networks (GRNs), and specifically, in determining the unique genetic circuits that construct characters. In this study, I discuss character identity and panarthropod appendage evolution within a developmental GRN framework, with a specific focus on potential limb character identity networks ("ChINs"). I summarize recent molecular studies, and argue that current data do not rule out the possibility of independent panarthropod limb evolution. The link between character identity and GRN architecture has broad implications for homology assessment, and this genetic framework offers alternative approaches to fossil character coding, phylogenetic analyses, and future research into the origin of the arthropod body plan. © The Author 2017. Published by Oxford University Press on behalf of the Society for Integrative and Comparative Biology. All rights reserved. For permissions please email: journals.permissions@oup.com.

  6. Age Validation in the Long Life Family Study Through a Linkage to Early-Life Census Records

    PubMed Central

    2013-01-01

    Objectives. Studies of health and longevity require accurate age reporting. Age misreporting among older adults in the United States is common. Methods. Participants in the Long Life Family Study (LLFS) were matched to early-life census records. Age recorded in the census was used to evaluate age reporting in the LLFS. The study population was 99% non-Hispanic white. Results. About 88% of the participants were matched to 1910, 1920, or 1930 U.S. censuses. Match success depended on the participant’s education, place of birth, and the number of censuses available to be searched. Age at the time of the interview based on the reported date of birth and early-life census age were consistent for about 89% of the participants, and age consistency within 1 year was found for about 99% of the participants. Discussion. It is possible to match a high fraction of older study participants to their early-life census records when detailed information is available on participants’ family of origin. Such record linkage can provide an important source of information for evaluating age reporting among the oldest old participants. Our results are consistent with recent studies suggesting that age reporting among older whites in the United States appears to be quite good. PMID:23704206

  7. Constraints on early events in Martian history as derived from the cratering record

    NASA Technical Reports Server (NTRS)

    Barlow, Nadine G.

    1990-01-01

    Constrains on early events in Martian history are derived using the planet's cratering record. Variations in the shapes of the crater size-frequency distribution curves are interpreted as indicative of the size-frequency distribution of the production populations, thus providing information about the age of the unit relative to the end of the heavy bombardment period. Results from the analysis of craters superposed on heavily cratered units across the Martian surface provide constraints on the hemispheric dichotomy and the early erosional conditions on Mars.

  8. Discriminating signal from noise in the fossil record of early vertebrates reveals cryptic evolutionary history

    PubMed Central

    Sansom, Robert S.; Randle, Emma; Donoghue, Philip C. J.

    2015-01-01

    The fossil record of early vertebrates has been influential in elucidating the evolutionary assembly of the gnathostome bodyplan. Understanding of the timing and tempo of vertebrate innovations remains, however, mired in a literal reading of the fossil record. Early jawless vertebrates (ostracoderms) exhibit restriction to shallow-water environments. The distribution of their stratigraphic occurrences therefore reflects not only flux in diversity, but also secular variation in facies representation of the rock record. Using stratigraphic, phylogenetic and palaeoenvironmental data, we assessed the veracity of the fossil records of the jawless relatives of jawed vertebrates (Osteostraci, Galeaspida, Thelodonti, Heterostraci). Non-random models of fossil recovery potential using Palaeozoic sea-level changes were used to calculate confidence intervals of clade origins. These intervals extend the timescale for possible origins into the Upper Ordovician; these estimates ameliorate the long ghost lineages inferred for Osteostraci, Galeaspida and Heterostraci, given their known stratigraphic occurrences and stem–gnathostome phylogeny. Diversity changes through the Silurian and Devonian were found to lie within the expected limits predicted from estimates of fossil record quality indicating that it is geological, rather than biological factors, that are responsible for shifts in diversity. Environmental restriction also appears to belie ostracoderm extinction and demise rather than competition with jawed vertebrates. PMID:25520359

  9. Early Terrestrial Mantle Differentiation Recorded in Paleoarchean Komatiites

    NASA Astrophysics Data System (ADS)

    Puchtel, I. S.; Blichert-Toft, J.; Touboul, M.; Horan, M. F.; Walker, R. J.

    2016-12-01

    Geochmical signatures generated in the manle as a result of radioactive decay of short- and long-lived nuclides can be used to constrain the timing of formation and the nature of now mostly vanished early terrestrial reservoirs. The 3.55 Ga komatiites from the Schapenburg Greenstone Remnant (SGR) located in the Barberton Greenstone Belt in South Africa have a unique combination of trace element abundances and isotopic compositions that place strong constraints on the origin of these reservoirs. The SGR komatiites define a Re-Os isochron with an age of 3550±87 Ma and an initial γ187Os = +3.7±0.2 (2SD). The absolute HSE abundances in the mantle source of the SGR komatiite system are estimated to be only 29±5% of those in the present-day bulk silicate Earth (BSE) estimates. The SGR komatiites show coupled depletion, relative to the modern mantle, in 142Nd and 182W (μ142Nd = -5.0±2.8, μ182W = -8.4±4.5), the decay products of the short-lived 146Sm and 182Hf nuclides, respectively, indicating derivation from a mantle domain that was enriched in incompatible elements 30 Ma after Solar System formation. Early Hadean contributors to this mantle domain could include high-pressure fractionates from a primordial magma ocean. By contrast, the long-lived Sm-Nd and Lu-Hf isotope systems (ɛ143Nd = +2.4±0.1, ɛ176Hf = +5.7±0.3) indicate that the mantle domain that the SGR komatiites were ultimately derived from underwent additional processing after the early Hadean, including melt depletion at lower pressures. The preservation of early-formed 182W and 142Nd anomalies in the mantle until at least 3.55 Ga indicates that the products of early planetary differentiation survived both later planetary accretion and convective mantle mixing during the Hadean. This study lends further support to the notion that variable late accretion, by itself, cannot account for all of the observed W isotope and absolute and relative HSE abundance variations in the Archean mantle recorded by

  10. Late-stage sulfides and sulfarsenides in Lower Cambrian black shale (stone coal) from the Huangjiawan mine, Guizhou Province, People's Republic of China

    USGS Publications Warehouse

    Belkin, H.E.; Luo, K.

    2008-01-01

    The Ni-Mo Huangjiawan mine, Guizhou Province, People's Republic of China, occurs in Lower Cambrian black shale (stone coal) in an area where other mines have recently extracted ore from the same horizon. Detailed electron microprobe (EMPA) and scanning electron microscope (SEM) analyses of representative thin sections have revealed a complex assemblage of sulfides and sulfarsenides. Early sulfidic and phosphatic nodules and host matrix have been lithified, somewhat fractured, and then mineralized with later-stage sulfides and sulfarsenides. Gersdorffite, millerite, polydymite, pyrite, sphalerite, chalcopyrite, galena, and clausthalite have been recognized. EMPA data are given for the major phases. Pyrite trace-element distributions and coeval Ni-, As-sulfides indicate that in the main ore layer, the last sulfide deposition was Ni-As-Co-rich. Mo and V deposition were early in the petrogenesis of these rocks. The assemblages gersdorffite-millerite-polydymite (pyrite) and millerite-gersdorffite (pyrite) and the composition of gersdorffite indicate a formation temperature of between 200?? and 300??C suggesting that the last solutions to infiltrate and mineralize the samples were related to hydrothermal processes. Environmentally sensitive elements such as As, Cd, and Se are hosted by sulfides and sulfarsenides and are the main source of these elements to residual soil. Crops grown on them are enriched in these elements, and they may be hazardous for animal and human consumption. ?? Springer-Verlag 2007.

  11. Metamorphosis Is Ancestral for Crown Euarthropods, and Evolved in the Cambrian or Earlier.

    PubMed

    Wolfe, Joanna M

    2017-09-01

    Macroevolutionary developmental biology employs fossilized ontogenetic data and phylogenetic comparative methods to probe the evolution of development at ancient nodes. Despite the prevalence of ecologically differentiated larval forms in marine invertebrates, it has been frequently presumed that the ancestors of arthropods were direct developers, and that metamorphosis may not have evolved until the Ordovician or later. Using fossils and new dated phylogenies, I infer that metamorphosis was likely ancestral for crown arthropods, contradicting this assumption. Based on a published morphological dataset encompassing 217 exceptionally preserved fossil and 96 extant taxa, fossils were directly incorporated into both the topology and age estimates, as in "tip dating" analyses. Using data from post-embryonic fossils representing 25 species throughout stem and crown arthropod lineages (as well as most of the 96 extant taxa), characters for metamorphosis were assigned based on inferred ecological changes in development (e.g., changes in habitat and adaptive landscape). Under all phylogenetic hypotheses, metamorphosis was supported as most likely ancestral to both ecdysozoans and euarthropods. Care must be taken to account for potential drastic post-embryonic morphological changes in evolutionary analyses. Many stem group euarthrpods may have had ecologically differentiated larval stages that did not preserve in the fossil record. Moreover, a complex life cycle and planktonic ecology may have evolved in the Ediacaran or earlier, and may have typified the pre-Cambrian explosion "wormworld" prior to the origin of crown group euarthropods. © The Author 2017. Published by Oxford University Press on behalf of the Society for Integrative and Comparative Biology. All rights reserved. For permissions please email: journals.permissions@oup.com.

  12. The characteristics and controlling factor of the multi-stage eogenetic karst in the Longwangmiao Foramtion in Lower Cambrian, western Central Yangtze Block, SW China

    NASA Astrophysics Data System (ADS)

    Lu, C.

    2017-12-01

    This study utilized field outcrops, thin sections, geochemical data, and GR logging curves to investigate the development model of paleokarst within the Longwangmiao Formation in the Lower Cambrian, western Central Yangtze Block, SW China. The Longwangmiao Formation, which belongs to a third-order sequence, consists of four forth-order sequences and is located in the uppermost part of the Lower Cambrian. The vertical variations of the δ13C and δ18O values indicate the existence of multi-stage eogenetic karst events. The eogenetic karst event in the uppermost part of the Longwangmiao Formation is recognized by the dripstones developed within paleocaves, vertical paleoweathering crust with four zones (bedrock, a weak weathering zone, an intense weathering zone and a solution collapsed zone), two generations of calcsparite cement showing bright luminescence and a zonation from nonluminescent to bright to nonluminescent, two types breccias (matrix-rich clast-supported chaotic breccia and matrix-supported chaotic breccia) and rundkarren. The episodic variations of stratiform dissolution vugs and breccias in vertical, and facies-controlled dissolution and filling features indicated the development of multi-stages eogenetic karst. The development of the paleokarst model is controlled by multi-level sea-level changes. The long eccentricity cycle dictates the fluctuations of the forth-order sea-level, generating multi-stage eogenetic karst events. The paleokarst model is an important step towards better understanding the link between the probably orbitally forced sea-level oscillations and eogenetic karst in the Lower Cambrian. According to this paleokarst model, hydrocarbon exploration should focus on both the karst highlands and the karst transitional zone.

  13. Trilobite-based biostratigraphy (arthropoda-trilobita) and related faunas of the Cambrian from Sonora, Mexico

    NASA Astrophysics Data System (ADS)

    Cuen-Romero, F. J.; Valdez-Holguín, J. E.; Buitrón-Sánchez, B. E.; Monreal, R.; Enríquez-Ocaña, L. F.; Aguirre-Hinojosa, E.; Ochoa-Granillo, J. A.; Palafox-Reyes, J. J.

    2018-04-01

    A biostratigraphic analysis based on trilobites of the main Cambrian outcrops from Sonora, Mexico is performed. The data are based on a combination of field work and published sources, including four previously studied locations from northern and eastern Sonora (Caborca, Cananea, Mazatán, and Arivechi) as well as a new location in the central part of the state of Sonora (San José de Gracia). Chronostratigraphic positions are assigned to the Cambrian outcrops according to Peng et al., 2012 and Cohen et al., 2017. In the Caborca area, the Puerto Blanco, Proveedora, Buelna, Cerro Prieto, Arrojos and El Tren formations comprise a wide range of biozones, which starts from the Fritzaspis Zone until the Glossopleura walcotti Zone (Begadean-Lincolnian Series, global Stage 3-Stage 5, Series 2-Series 3). The Bolsa Quartzite and the Abrigo Limestone exposed in Cananea area are assigned to the Cedaria/Cedarina dakotaensis Zone until the Crepicephalus Zone (Lincolnian Series-Marjuman Stage, global Series 3-Guzhangian). In the San José de Gracia area, The Proveedora, Buelna, Cerro Prieto and El Gavilán formations range from the ?Bristolia mohavensis or ?Bristolia insolens zones until the upper part of Mexicella mexicana Zone, Albertella highlandensis Subzone (Series 2-Series 3, Stage 4-Stage 5). In the Arivechi area, the La Sata, El Mogallón, La Huerta and the Milpillas formations range from Poliella denticulata Zone to the Elvinia Zone (Lincolnian-Millardan, Delamaran-Steptoean, global Series 3-Furongian, Stage 5-Paibian). Paleozoic marine fauna distribution in northwest Mexico and the southwest United States of America, suggest that during this time an extensive faunal province existed, containing a great variety of marine invertebrates with notorious intraspecific affinity. The biotic association includes poriferans, archaeocyathids, brachiopods, mollusks, arthropods and echinoderms as predominant elements.

  14. Early tectonic evolution of the Thomson Orogen in Queensland inferred from constrained magnetic and gravity data

    NASA Astrophysics Data System (ADS)

    Spampinato, Giovanni P. T.; Betts, Peter G.; Ailleres, Laurent; Armit, Robin J.

    2015-05-01

    The crustal architecture as well as the kinematic evolution of the Thomson Orogen in Queensland is poorly resolved because the region is concealed under thick Phanerozoic sedimentary basins and the basement geology is known from limited drill holes. Combined potential field and seismic interpretation indicates that the Thomson Orogen is characterized by prominent regional NE- and NW-trending structural grain defined by long wavelength and low amplitude geophysical anomalies. The 'smooth' magnetic signature is interpreted to reflect deeply buried source bodies in the mid- to lower crust. Short wavelength positive magnetic features that correlate with negative gravity anomalies are interpreted to represent shallower granitic intrusions. They appear to be focused along major fault zones that might have controlled the locus for magmatism. The eastern Thomson Orogen is characterized by a prominent NE structural grain and orthogonal faults and fold interference patterns resulting in a series of troughs and highs. The western Thomson Orogen consists of a series of NW-trending structures interpreted to reflect reverse faults. Sedimentation and basin development are interpreted to have initiated in the Neoproterozoic to Early Cambrian during E-W- to ENE-WSW extension, possibly related to the Rodinia break-up. This extensional event was followed by Late Cambrian shortening recorded in the Maneroo Platform and the Diamantina River Domain which possibly correlates with the Delamerian Orogeny. Renewed deposition and volcanism occurred during the Ordovician and may have continued until Late Silurian, resulting in thinned Proterozoic basement crust and extensive basin systems that formed in a distal continental back-arc environment. Our interpretation places the Thomson Orogen to the west of the Neoproterozoic passive margin preserved in the Anakie Inlier. The region is likely to represent the internal extensional architecture during the Rodinia break-up that has been

  15. Evidence for gill slits and a pharynx in Cambrian vetulicolians: implications for the early evolution of deuterostomes

    PubMed Central

    2012-01-01

    Background Vetulicolians are a group of Cambrian metazoans whose distinctive bodyplan continues to present a major phylogenetic challenge. Thus, we see vetulicolians assigned to groups as disparate as deuterostomes and ecdysozoans. This divergence of opinions revolves around a strikingly arthropod-like body, but one that also bears complex lateral structures on its anterior section interpreted as pharyngeal openings. Establishing the homology of these structures is central to resolving where vetulicolians sit in metazoan phylogeny. Results New material from the Chengjiang Lagerstätte helps to resolve this issue. Here, we demonstrate that these controversial structures comprise grooves with a series of openings. The latter are oval in shape and associated with a complex anatomy consistent with control of their opening and closure. Remains of what we interpret to be a musculature, combined with the capacity for the grooves to contract, indicate vetulicolians possessed a pumping mechanism that could process considerable volumes of seawater. Our observations suggest that food captured in the anterior cavity was transported to dorsal and ventral gutters, which then channeled material to the intestine. This arrangement appears to find no counterpart in any known fossil or extant arthropod (or any other ecdysozoan). Anterior lateral perforations, however, are diagnostic of deuterostomes. Conclusions If the evidence is against vetulicolians belonging to one or other group of ecdysozoan, then two phylogenetic options seem to remain. The first is that such features as vetulicolians possess are indicative of either a position among the bilaterians or deuterostomes but apart from the observation that they themselves form a distinctive and recognizable clade current evidence can permit no greater precision as to their phylogenetic placement. We argue that this is too pessimistic a view, and conclude that evidence points towards vetulicolians being members of the stem

  16. Experimental taphonomy and the anatomy and diversity of the earliest fossil vertebrates (Chengjiang Biota, Cambrian, China)

    NASA Astrophysics Data System (ADS)

    Purnell, Mark; Gabbott, Sarah; Murdock, Duncan; Cong, Peiyun

    2016-04-01

    The oldest fossil vertebrates are from the Lower Cambrian Chengjiang biota of China, which contains four genera of fish-like, primitive vertebrates: Haikouichthys, Myllokunmingia, Zhongjianichthys and Zhongxiniscus. These fossils play key roles in calibrating molecular clocks and informing our view of the anatomy of animals close to the origin of vertebrates, potentially including transitional forms between vertebrates and their nearest relatives. Despite the evident importance of these fossils, the degree to which taphonomic processes have affected their anatomical completeness has not been investigated. For example, some or all might have been affected by stemward slippage - the pattern observed in experimental decay of non-biomineralised chordates in which preferential decay of synapomorphies and retention of plesiomorphic characters would cause fossil taxa to erroneously occupy more basal positions than they should. This hypothesis is based on experimental data derived from decay of non-biomineralised chordates under laboratory conditions. We have expanded this analysis to include a broader range of potentially significant environmental variables; we have also compared and combined the results of experiments from several taxa to identify general patterns of chordate decay. Examination of the Chengjiang vertebrates in the light of these results demonstrates that, contrary to some assertions, experimentally derived models of phylogenetic bias are applicable to fossils. Anatomical and phylogenetic interpretations of early vertebrates that do not take taphonomic biases into account risk overestimating diversity and the evolutionary significance of differences between fossil specimens.

  17. [Electroencephalography (EEG) recording techniques and artifact detection in early premature babies].

    PubMed

    Wallois, F; Vecchierini, M-F; Héberlé, C; Walls-Esquivel, E

    2007-01-01

    EEG recording techniques in early premature babies are not very different from those used for full-term neonates. Here, we emphasise the most important points: asepsis precautions, full knowledge of the clinical data and drug therapies, the fundamental role of a well-trained technician in supervising the EEG recording and monitoring the baby. The best electrode positions, the most informative montages and their standardisation between neurophysiological laboratories, are suggested. Artifact detection constitutes an important aspect of EEG signal analysis in preterm babies of less than 30 weeks. It is obviously necessary to discriminate between meaningful information and artefacts. The complexity of the signal in neonates makes artifact detection difficult. We present some characteristic features and describe some methods for eliminating them. We underline the positive aspect of some artifacts and their clinical use. We emphasise the crucial role of the technicians.

  18. Early cave art and ancient DNA record the origin of European bison

    PubMed Central

    Soubrier, Julien; Gower, Graham; Chen, Kefei; Richards, Stephen M.; Llamas, Bastien; Mitchell, Kieren J.; Ho, Simon Y. W.; Kosintsev, Pavel; Lee, Michael S. Y.; Baryshnikov, Gennady; Bollongino, Ruth; Bover, Pere; Burger, Joachim; Chivall, David; Crégut-Bonnoure, Evelyne; Decker, Jared E.; Doronichev, Vladimir B.; Douka, Katerina; Fordham, Damien A.; Fontana, Federica; Fritz, Carole; Glimmerveen, Jan; Golovanova, Liubov V.; Groves, Colin; Guerreschi, Antonio; Haak, Wolfgang; Higham, Tom; Hofman-Kamińska, Emilia; Immel, Alexander; Julien, Marie-Anne; Krause, Johannes; Krotova, Oleksandra; Langbein, Frauke; Larson, Greger; Rohrlach, Adam; Scheu, Amelie; Schnabel, Robert D.; Taylor, Jeremy F.; Tokarska, Małgorzata; Tosello, Gilles; van der Plicht, Johannes; van Loenen, Ayla; Vigne, Jean-Denis; Wooley, Oliver; Orlando, Ludovic; Kowalczyk, Rafał; Shapiro, Beth; Cooper, Alan

    2016-01-01

    The two living species of bison (European and American) are among the few terrestrial megafauna to have survived the late Pleistocene extinctions. Despite the extensive bovid fossil record in Eurasia, the evolutionary history of the European bison (or wisent, Bison bonasus) before the Holocene (<11.7 thousand years ago (kya)) remains a mystery. We use complete ancient mitochondrial genomes and genome-wide nuclear DNA surveys to reveal that the wisent is the product of hybridization between the extinct steppe bison (Bison priscus) and ancestors of modern cattle (aurochs, Bos primigenius) before 120 kya, and contains up to 10% aurochs genomic ancestry. Although undetected within the fossil record, ancestors of the wisent have alternated ecological dominance with steppe bison in association with major environmental shifts since at least 55 kya. Early cave artists recorded distinct morphological forms consistent with these replacement events, around the Last Glacial Maximum (LGM, ∼21–18 kya). PMID:27754477

  19. The nature and location of the suture zone in the Rokelide orogen, Sierra Leone: Geochemical evidence

    NASA Astrophysics Data System (ADS)

    Lytwyn, Jennifer; Burke, Kevin; Culver, Stephen

    2006-12-01

    The boundaries of the West African Craton mark the location of a continuous suture zone that records Neoproterozoic to Early Cambrian oceanic closure. The western part of the circum-West African suture zone extends through the line of outcrop of the Mauritanide, Bassaride and Rokelide mountain belts. Our geochemical analyses are consistent with the idea that igneous and metamorphic rocks of the Rokelide and Southern Mauritanide mountain belts of West Africa occupy a suture zone that records the closing of a Neoproterozoic to Early Cambrian ocean basin during the Pan-African orogeny and final assembly of Gondwana. The closing of that basin was marked by the collision between Archean rocks of the Leo massif of the West African Craton and reactivated Archean and Paleoproterozoic rocks that now outcrop nearer to the coast of Africa in Sierra Leone and Liberia. Within the Rokelides, the geochemistry of the Kasewe Hills volcanic rocks and Marampa amphibolite indicate that remnants of an arc system are caught up in the suture zone. The geochemistry of Guingan schists that outcrop along strike of the Rokelides is compatible with the idea that the metamorphosed equivalents of the Marampa and Kasewe Hills arc volcanic rocks extend through the Bassarides and into the Southern Mauritanides.

  20. Baltica from the Late Precambrian to the Early Ordovician

    NASA Astrophysics Data System (ADS)

    Torsvik, T. H.; Cocks, L. R.

    2004-05-01

    Current thinking supports the existence of the Rodinia supercontinent which consolidated at perhaps 1100 to 1000 million years ago and most probably disintegrated somewhere before 800 Ma. Within the Rodinian collage, Baltica was adjacent to, and probably welded to, Laurentia, with the modern eastern (Uralian) part of Baltica conjugate with the north of Laurentia. Laurentia was in turn attached to the South American terranes of Rio Plata and Amazonia, and possibly also West Africa. Baltica became an independent terrane when it split off from Laurentia, leaving a widening Iapetus Ocean between the two. When this rifting actually commenced is a matter of dispute due to very conflicting palaeomagnetic data from both Laurentia and Baltica; however, we favor that the southern part of the Iapetus, between Laurentia and South America, opened first at about 570 Ma and that this rifting spread gradually northwards until Baltica separated from Laurentia at approximately 550 Ma, near the end of Precambrian time at 543 Ma. New palaeomagnetic and geochronological data from the 616-610 Ma Egersund Dykes (SW Baltica) place Baltica at the south pole whereas subsequent Late Precambrian to Early Cambrian poles places Baltica at lower latitudes. During the late and middle Vendian, the NW margin of Baltica changed from an extensional tectonic regime to an active margin (Timanian Orogeny) in which microcontinental blocks in the Timan-Pechora, northern Ural and Novaya Zemlya areas were united with Baltica at 550-560 Ma. Largely between Middle Cambrian and Middle Ordovician times, the whole large terrane of Baltica underwent a very substantial rotation of about 120°, the maximum rate of this rotation occurred in late Cambrian and early Ordovician times. Much of the craton of Baltica appears to have been submerged under shelf seas for long parts of this time, which lasted from 544 to 490 Ma. As a consequence the olenid trilobite fauna represent a fauna living largely in niches which were

  1. Mapping the Early Language Environment Using All-Day Recordings and Automated Analysis.

    PubMed

    Gilkerson, Jill; Richards, Jeffrey A; Warren, Steven F; Montgomery, Judith K; Greenwood, Charles R; Kimbrough Oller, D; Hansen, John H L; Paul, Terrance D

    2017-05-17

    This research provided a first-generation standardization of automated language environment estimates, validated these estimates against standard language assessments, and extended on previous research reporting language behavior differences across socioeconomic groups. Typically developing children between 2 to 48 months of age completed monthly, daylong recordings in their natural language environments over a span of approximately 6-38 months. The resulting data set contained 3,213 12-hr recordings automatically analyzed by using the Language Environment Analysis (LENA) System to generate estimates of (a) the number of adult words in the child's environment, (b) the amount of caregiver-child interaction, and (c) the frequency of child vocal output. Child vocalization frequency and turn-taking increased with age, whereas adult word counts were age independent after early infancy. Child vocalization and conversational turn estimates predicted 7%-16% of the variance observed in child language assessment scores. Lower socioeconomic status (SES) children produced fewer vocalizations, engaged in fewer adult-child interactions, and were exposed to fewer daily adult words compared with their higher socioeconomic status peers, but within-group variability was high. The results offer new insight into the landscape of the early language environment, with clinical implications for identification of children at-risk for impoverished language environments.

  2. The evolution of the passive continental margin of Norway and its adjacent mainland - using the sub-Cambrian peneplain as a reference surface.

    NASA Astrophysics Data System (ADS)

    Gabrielsen, R. H.; Faleide, J. I.; Jarsve, E. M.

    2012-04-01

    The structuring, uplift and subsidence of the passive margin and shelf of Norway and its adjacent mainland were affected by several profound geological processes, including inherited basement structural grain related to the Proterozoic and Caledonian orogens and also including the extensional collapse of the Caledonides (Gabrielsen et al. 2000). This has been followed by several stages of late Palaoezoic - Cenozoic rifting and associated thermotectonic activity, Cenozoic accelerated uplift of uncertain origin of the hinterland, creating an irregular pattern of upheaval and, finally Pleistocene - Holocene glacial loading and unloading (Gabrielsen et al. 2010). These processes have strongly influenced the topography of the hinterland, thus causing and acting in concert with climate fluctuations (Nielsen et al. 2009). The correlation of erosional surfaces of regional significance on the shelf and on the mainland is a key to the evaluating the total topography of the margin. Because of the lack of datable surfaces on the mainland, this is problematic. The so-called Paleic surface has been used in this context, but its age and nature is not well constrained and the absence of post-Caledonian rocks in the western and central mainland of southern Norway adds to this complexity. In contrast, the sub-Cambrian peneplain, which is found in larger parts of Scandinavia, is well established when it comes to dating and development (e.g. Strøm 1948). It is generally accepted that this surface had only a minor topography, if any, throughout Scandinavia at the earliest Cambrian. Hence, its present relief is the result of the accumulated vertical displacements from the Caledonian to the Present. Still, even though it was well established through regional mapping already in the late 19th century, much remains in the detailed documentation of this important surface. To improve the topographic accuracy in its characterization, fieldwork has been initiated to establish a detailed WNW

  3. Alxa Block Provenance of Ediacaran (Sinian) Sediments in the Helanshan Area: Constraints from Hf Isotopes and U-Pb Geochronology of Detrital Zircons

    NASA Astrophysics Data System (ADS)

    Xiaopeng, D.

    2016-12-01

    The tectonic relationship between the Alxa Block and the North China Craton has long been controversial. The Helanshan area lies at the western margin of the Ordos Block and east of the Alxa Block (Fig.a), and it contains rocks of the lower Zhengmuguan and upper Tuerkeng formations that belong to the Ediacaran system. The Zhengmuguan Formation is made up of abyssal facies rocks including dolomite and glacial conglomerate with dropstones, and the Tuerkeng Formation consists of silty slate of the neritic facies. A discontinuity marks the boundary between the Tuerkeng Formation and the Early Cambrian Suyukou Formation, which is composed mainly of pebbly sandstone towards the base and sandstone towards the top, representing a change in sedimentary facies from terrestrial to littoral.The Neoproterozoic U-Pb ages of zircons from the Ediacaran and Early Cambrian sediments peak at 818 ± 4 Ma (n = 88) and 905 ± 8 Ma (n = 20), consistent with the Neoproterozoic age peaks found in the Precambrian basement of the Alxa Block(Fig.b). There are few Neoproterozoic zircons in the Neoproterozoic strata of the Langshan area, and there are no reports of Neoproterozoic zircons in the Zhuozishan area, northwest of Helanshan, or in the western margin of the neighboring Ordos Basin. A number of Neoproterozoic zircons are found in the Middle Cambrian to Middle Ordovician strata of the Niushoushan area. And while Niushoushan is part of the Hexi Corridor, it did not amalgamate with the NCC before the Early-Middle Cambrian. Therefore, the Neoproterozoic and Early Cambrian sediments in Helanshan record information about Neoproterozoic magmatic events in the Alxa Block, and indicate an Alxa Block provenance(Fig.c).The Hf isotopic characteristics of the Neoproterozoic zircons from the Ediacaran Zhengmuguan Formation in the Helanshan area (eHf(t) = -7.812 to 3.274, TDMC = 2211-1578 Ma, n = 10) are similar to those Neoproterozoic igneous zircons from the Langshan area (eHf(t) = -1.105 to 5

  4. Cambrian nepheline syenite complex at Jabal Sawda, Midyan region, Kingdom of Saudi Arabia

    USGS Publications Warehouse

    Liddicoat, W.K.; Ramsay, C.R.; Hedge, C.E.

    1986-01-01

    The only nepheline syenite complex presently known in the Arabian Shield is at Jabal Sawda, about 30 km S of Haql in the extreme NW of Saudi Arabia. It is a post-tectonic, composite intrusion with a crudely concentric structure. A core of leuco-nepheline syenite, a partial ring of mela-nepheline syenite, and an almost complete outer ring of alkali-feldspar syenite are the main rock units. Several mega-inclusions of porphyritic nepheline syenite, nepheline monzosyenite, malignite and ijolite are present in the leuco-nepheline syenite. The chemical composition is notable for very high values of Al2O3, Na2O, Ba, La, Nb, Sr and Zr. U{single bond}Pb isotope dating indicates an emplacement age of 553 ?? 4 Ma, one of an increasing number of reliable Cambrian isotope dates in the northern Red Sea region. ?? 1986.

  5. Cambrian nepheline syenite complex at Jabal Sawda, Midyan region, Kingdom of Saudi Arabia

    NASA Astrophysics Data System (ADS)

    Liddicoat, W. K.; Ramsay, C. R.; Hedge, C. E.

    The only nepheline syenite complex presently known in the Arabian Shield is at Jabal Sawda, about 30 km S of Haql in the extreme NW of Saudi Arabia. It is a post-tectonic, composite intrusion with a crudely concentric structure. A core of leuco-nepheline syenite, a partial ring of mela-nepheline syenite, and an almost complete outer ring of alkali-feldspar syenite are the main rock units. Several mega-inclusions of porphyritic nepheline syenite, nepheline monzosyenite, malignite and ijolite are present in the leuco-nepheline syenite. The chemical composition is notable for very high values of Al 2O 3, Na 2O, Ba, La, Nb, Sr and Zr. U sbnd Pb isotope dating indicates an emplacement age of 553 ± 4 Ma, one of an increasing number of reliable Cambrian isotope dates in the northern Red Sea region.

  6. Orogen styles in the East African Orogen: A review of the Neoproterozoic to Cambrian tectonic evolution.

    PubMed

    Fritz, H; Abdelsalam, M; Ali, K A; Bingen, B; Collins, A S; Fowler, A R; Ghebreab, W; Hauzenberger, C A; Johnson, P R; Kusky, T M; Macey, P; Muhongo, S; Stern, R J; Viola, G

    2013-10-01

    The East African Orogen, extending from southern Israel, Sinai and Jordan in the north to Mozambique and Madagascar in the south, is the world́s largest Neoproterozoic to Cambrian orogenic complex. It comprises a collage of individual oceanic domains and continental fragments between the Archean Sahara-Congo-Kalahari Cratons in the west and Neoproterozoic India in the east. Orogen consolidation was achieved during distinct phases of orogeny between ∼850 and 550 Ma. The northern part of the orogen, the Arabian-Nubian Shield, is predominantly juvenile Neoproterozoic crust that formed in and adjacent to the Mozambique Ocean. The ocean closed during a protracted period of island-arc and microcontinent accretion between ∼850 and 620 Ma. To the south of the Arabian Nubian Shield, the Eastern Granulite-Cabo Delgado Nappe Complex of southern Kenya, Tanzania and Mozambique was an extended crust that formed adjacent to theMozambique Ocean and experienced a ∼650-620 Ma granulite-facies metamorphism. Completion of the nappe assembly around 620 Ma is defined as the East African Orogeny and was related to closure of the Mozambique Ocean. Oceans persisted after 620 Ma between East Antarctica, India, southern parts of the Congo-Tanzania-Bangweulu Cratons and the Zimbabwe-Kalahari Craton. They closed during the ∼600-500 Ma Kuungan or Malagasy Orogeny, a tectonothermal event that affected large portions of southern Tanzania, Zambia, Malawi, Mozambique, Madagascar and Antarctica. The East African and Kuungan Orogenies were followed by phases of post-orogenic extension. Early ∼600-550 Ma extension is recorded in the Arabian-Nubian Shield and the Eastern Granulite-Cabo Delgado Nappe Complex. Later ∼550-480 Ma extension affected Mozambique and southern Madagascar. Both extension phases, although diachronous,are interpreted as the result of lithospheric delamination. Along the strike of the East African Orogen, different geodynamic settings resulted in the evolution

  7. Orogen styles in the East African Orogen: A review of the Neoproterozoic to Cambrian tectonic evolution

    NASA Astrophysics Data System (ADS)

    Fritz, H.; Abdelsalam, M.; Ali, K. A.; Bingen, B.; Collins, A. S.; Fowler, A. R.; Ghebreab, W.; Hauzenberger, C. A.; Johnson, P. R.; Kusky, T. M.; Macey, P.; Muhongo, S.; Stern, R. J.; Viola, G.

    2013-10-01

    The East African Orogen, extending from southern Israel, Sinai and Jordan in the north to Mozambique and Madagascar in the south, is the world´s largest Neoproterozoic to Cambrian orogenic complex. It comprises a collage of individual oceanic domains and continental fragments between the Archean Sahara-Congo-Kalahari Cratons in the west and Neoproterozoic India in the east. Orogen consolidation was achieved during distinct phases of orogeny between ∼850 and 550 Ma. The northern part of the orogen, the Arabian-Nubian Shield, is predominantly juvenile Neoproterozoic crust that formed in and adjacent to the Mozambique Ocean. The ocean closed during a protracted period of island-arc and microcontinent accretion between ∼850 and 620 Ma. To the south of the Arabian Nubian Shield, the Eastern Granulite-Cabo Delgado Nappe Complex of southern Kenya, Tanzania and Mozambique was an extended crust that formed adjacent to theMozambique Ocean and experienced a ∼650-620 Ma granulite-facies metamorphism. Completion of the nappe assembly around 620 Ma is defined as the East African Orogeny and was related to closure of the Mozambique Ocean. Oceans persisted after 620 Ma between East Antarctica, India, southern parts of the Congo-Tanzania-Bangweulu Cratons and the Zimbabwe-Kalahari Craton. They closed during the ∼600-500 Ma Kuungan or Malagasy Orogeny, a tectonothermal event that affected large portions of southern Tanzania, Zambia, Malawi, Mozambique, Madagascar and Antarctica. The East African and Kuungan Orogenies were followed by phases of post-orogenic extension. Early ∼600-550 Ma extension is recorded in the Arabian-Nubian Shield and the Eastern Granulite-Cabo Delgado Nappe Complex. Later ∼550-480 Ma extension affected Mozambique and southern Madagascar. Both extension phases, although diachronous,are interpreted as the result of lithospheric delamination. Along the strike of the East African Orogen, different geodynamic settings resulted in the evolution of

  8. Cause of Cambrian Explosion - Terrestrial or Cosmic?

    PubMed

    Steele, Edward J; Al-Mufti, Shirwan; Augustyn, Kenneth A; Chandrajith, Rohana; Coghlan, John P; Coulson, S G; Ghosh, Sudipto; Gillman, Mark; Gorczynski, Reginald M; Klyce, Brig; Louis, Godfrey; Mahanama, Kithsiri; Oliver, Keith R; Padron, Julio; Qu, Jiangwen; Schuster, John A; Smith, W E; Snyder, Duane P; Steele, Julian A; Stewart, Brent J; Temple, Robert; Tokoro, Gensuke; Tout, Christopher A; Unzicker, Alexander; Wainwright, Milton; Wallis, Jamie; Wallis, Daryl H; Wallis, Max K; Wetherall, John; Wickramasinghe, D T; Wickramasinghe, J T; Wickramasinghe, N Chandra; Liu, Yongsheng

    2018-08-01

    We review the salient evidence consistent with or predicted by the Hoyle-Wickramasinghe (H-W) thesis of Cometary (Cosmic) Biology. Much of this physical and biological evidence is multifactorial. One particular focus are the recent studies which date the emergence of the complex retroviruses of vertebrate lines at or just before the Cambrian Explosion of ∼500 Ma. Such viruses are known to be plausibly associated with major evolutionary genomic processes. We believe this coincidence is not fortuitous but is consistent with a key prediction of H-W theory whereby major extinction-diversification evolutionary boundaries coincide with virus-bearing cometary-bolide bombardment events. A second focus is the remarkable evolution of intelligent complexity (Cephalopods) culminating in the emergence of the Octopus. A third focus concerns the micro-organism fossil evidence contained within meteorites as well as the detection in the upper atmosphere of apparent incoming life-bearing particles from space. In our view the totality of the multifactorial data and critical analyses assembled by Fred Hoyle, Chandra Wickramasinghe and their many colleagues since the 1960s leads to a very plausible conclusion - life may have been seeded here on Earth by life-bearing comets as soon as conditions on Earth allowed it to flourish (about or just before 4.1 Billion years ago); and living organisms such as space-resistant and space-hardy bacteria, viruses, more complex eukaryotic cells, fertilised ova and seeds have been continuously delivered ever since to Earth so being one important driver of further terrestrial evolution which has resulted in considerable genetic diversity and which has led to the emergence of mankind. Copyright © 2018 The Authors. Published by Elsevier Ltd.. All rights reserved.

  9. Les microbialites de l'Anti-Atlas occidental (Maroc) : marqueurs stratigraphiques et témoins des changements environnementaux au Cambrien inférieur Stratigraphic and environmental significance of the Lower-Cambrian western Anti-Atlasic microbialites (Morocco)

    NASA Astrophysics Data System (ADS)

    Benssaou, Mohammed; Hamoumi, Naima

    2004-02-01

    Three microbialite forms are recognized in the Lower-Cambrian succession of Irherm area in the western Anti-Atlas (Morocco). Stromatolites, which correspond to non-calcified shallow marine laminated microbialites, are well developed in the basal Lower-Cambrian succession. Occurrence of calcified microbial thrombolites, in the middle part of this succession, reflects an increasing sea level from the peritidal zone to the subtidal environment. In the upper part of this succession, a second increasing water depth event and the development of branching archaeocyathan reefal framework lead to dendritic microbialite emergence. To cite this article: M. Benssaou, N. Hamoumi, C. R. Geoscience 336 (2004).

  10. Les marqueurs structuraux et magmatiques de l'extension crustale au Protérozoïque terminal-Cambrien basal autour du massif de Kerdous (Anti-Atlas occidental, Maroc)

    NASA Astrophysics Data System (ADS)

    Soulaimani, Abderrahmane; Essaifi, Abderrahim; Youbi, Nassrddine; Hafid, Ahmid

    2004-12-01

    During the Late Precambrian-Early Cambrian times, the borders of the Kerdous inlier were affected by normal faults where thick conglomerates (Ouarzazate Group: PIII), grading progressively upwards into Cambrian marine sediments, were accumulated along their hanging walls. This tectonic activity persisted during the Early Cambrian and was accompanied by a magmatic activity resulting mainly in the emplacement of continental tholeiitic basalts. These tectono-sedimentary and magmatic events are related to the crustal extensional episode that affected the northwestern Gondwana margin during the opening of the Iapetus Ocean during Late Proterozoic times. To cite this article: A. Soulaimani et al., C. R. Geoscience 336 (2004).

  11. Black shale source rocks and oil generation in the Cambrian and Ordovician of the central Appalachian Basin, USA

    USGS Publications Warehouse

    Ryder, R.T.; Burruss, R.C.; Hatch, J.R.

    1998-01-01

    Nearly 600 million bbl of oil (MMBO) and 1 to 1.5 trillion ft3 (tcf) of gas have been produced from Cambrian and Ordovician reservoirs (carbonate and sandstone) in the Ohio part of the Appalachian basin and on adjoining arches in Ohio, Indiana, and Ontario, Canada. Most of the oil and gas is concentrated in the giant Lima-Indiana field on the Findlay and Kankakee arches and in small fields distributed along the Knox unconformity. Based on new geochemical analyses of oils, potential source rocks, bitumen extracts, and previously published geochemical data, we conclude that the oils in both groups of fields originated from Middle and Upper Ordovician blcak shale (Utica and Antes shales) in the Appalachian basin. Moroever, we suggest that approximately 300 MMBO and many trillions of cubic feet of gas in the Lower Silurian Clinton sands of eastern Ohio originated in the same source rocks. Oils from the Cambrian and Ordovician reservoirs have similar saturated hydrocarbon compositions, biomarker distributions, and carbon isotope signatures. Regional variations in the oils are attributed to differences in thermal maturation rather than to differences in source. Total organic carbon content, genetic potential, regional extent, and bitument extract geochemistry identify the balck shale of the Utica and Antes shales as the most plausible source of the oils. Other Cambrian and Ordovician shale and carbonate units, such as the Wells Creek formation, which rests on the Knox unconformity, and the Rome Formation and Conasauga Group in the Rome trough, are considered to be only local petroleum sources. Tmax, CAI, and pyrolysis yields from drill-hole cuttings and core indicate that the Utica Shale in eastern and central Ohio is mature with respect to oil generation. Burial, thermal, and hydrocarbon-generation history models suggest that much of the oil was generated from the Utica-Antes source in the late Paleozoic during the Alleghanian orogeny. A pervasive fracture network

  12. Geohydrologic systems in Kansas with emphasis on systems in Upper Cambrian through Lower Cretaceous rocks

    USGS Publications Warehouse

    Wolf, R.J.; Hansen, C.V.; McGovern, H.E.; Spinazola, J.M.

    1990-01-01

    This Hydrologic Investigations Atlas, which consists of a series of chapters, presents a description of (1) the physical frameworks and (2) the geohydrology of the principal aquifers and confining systems in Kansas. The report is the result of an investigation that has been made as part of the Central Midwest Regional Aquifer System Analysis (CMRASA), one of several major investigations by the U.S. Geological Survey to define regional aquifer systems. These regional analyses are designed to increase knowledge of major flow regimes and provide data for assessing, developing, and managing water supplies. The CMRASA is an investigation of water in Upper Cambrian through Lower Cretaceous rocks in parts of 10 Central Midwestern States, as shown by the map on the envelope cover.

  13. "Soft-shelled" monothalamid foraminifers as a modern analogue of early life

    NASA Astrophysics Data System (ADS)

    Kitazato, Hiroshi; Ohkawara, Nina; Gooday, Andrew

    2017-04-01

    According to the fossil record, the earliest undoubted foraminifers are found in the Early Cambrian, where they are represented by tubular agglutinated forms, thought to be the most primitive foraminiferal morphotypes. The numerous foraminifers with single-chambered, organic-walled tests (i.e. 'soft-shelled' monothalamids) exist in the deep sea and are difficult to preserve as fossils. Molecular phylogenetic data tell us that these 'primitive' taxa include the deepest foraminiferal clades, originating around 600 - 900 Ma. We found many soft-shelled monothalamids in sediment samples from deep trenches, including the Challenger Deep (Marianas Trench) and the Horizon Deep (Tonga Trench). Both deeps exceed 10,000 m water depth, well below the carbonate compensation depth, which represents an environmental barrier for calcareous foraminifera. The foraminifera at these extreme hadal sites include tubular and globular forms with organic walls, among which species of the genera Nodellum and Resigella are particularly abundant. Some forms selectively agglutinate minute flakes of clay minerals on the surface of the organic test. Many soft-shelled monothalamids, including most of those in deep tranches, contain stercomata, the function of which is currently unknown. Gromiids (a rhizarian group related to foraminifera) also accumulate stercomata in their sack-shaped tests. This suggests the possibility that the function of these waste particles is to add bulk, like the filling of soft bags or pillows. We suggest that the monothalamid foraminifera that dominate small-sized eukaryotes in extreme hadal settings may provide clues to understanding the biology and ecology of early life in Neoproterozoic sedimented habitats.

  14. The fossil record of ecdysis, and trends in the moulting behaviour of trilobites.

    PubMed

    Daley, Allison C; Drage, Harriet B

    2016-03-01

    Ecdysis, the process of moulting an exoskeleton, is one of the key characters uniting arthropods, nematodes and a number of smaller phyla into Ecdysozoa. The arthropod fossil record, particularly trilobites, eurypterids and decapod crustaceans, yields information on moulting, although the current focus is predominantly descriptive and lacks a broader evolutionary perspective. We here review literature on the fossil record of ecdysis, synthesising research on the behaviour, evolutionary trends, and phylogenetic significance of moulting throughout the Phanerozoic. Approaches vary widely between taxonomic groups, but an overall theme uniting these works suggests that identifying moults in the palaeontological record must take into account the morphology, taphonomy and depositional environment of fossils. We also quantitatively analyse trends in trilobite ecdysis based on a newly generated database of published incidences of moulting behaviour. This preliminary work reveals significant taxonomic and temporal signal in the trilobite moulting fossil record, with free cheek moulting being prevalent across all Orders and throughout the Phanerozoic, and peaks of cephalic moulting in Phacopida during the Ordovician and rostral plate moulting in Redlichiida during the Cambrian. This study and a review of the literature suggest that it is feasible to extract large-scale evolutionary information from the fossil record of moulting. Copyright © 2015 Elsevier Ltd. All rights reserved.

  15. Precambrian Continent Arctida: A New Kinematic Reconstruction of Late Precambrian - Early Paleozoic Arctida U Europe (baltia) Collision

    NASA Astrophysics Data System (ADS)

    Borisova, T. P.; Guertseva, M. V.; Egorov, A. Ju.; Kononov, M. V.; Kouznetsov, N. B.

    In according to L.P.Zonenshain and L.M.Natapov (1988, 1990), different size conti- nental blocks locating at the margins and inside of present-day Arctic ocean composed the hypothetical early Paleozoic paleocontinent Arctida. The blocks are Kara block (north part of Taymir peninsula, Severnaja Zemlja archipelago and Franz Joseph Land archipelago), north part of Alaska (northward Bruks ridge), Chukchi block, Novosi- birsky block (Novosibirskiye islands together their shelves), several fragments north- ward to the Innuitian orogen (north parts of Peary Land and Ellesmere Island), and Lomonosov ridgeSs block. In the previous kinematic reconstruction it was believed that Arctida as a whole collided with north flanks of Laurentia (Innuitian margin) and Europe (Baltia, Barentsia margin) in middle Paleozoic time. Later, the Arctida (been a fragment of supercontinent Pangea) was fragmented due to a spreading in the Arctic ocean and north part of Atlantic ocean in late Mesozoic and Cenozoic times. Then ArctidaSs fragments were accreted to the Eurasia and North America conti- nents. During the last decade "AEROGEOLOGIA" company has been gathered new data (geologic, stratigraphical, paleomagnetic, and others) of Russian Arctic sector and Svalbard. The data were summarized into "Paleogeographical Atlas for the Rus- sian Arctic sector and Svalbard from Vendian to Jurassic times" (see Abstact SE1.04, ID-NR: EGS02-A-02453). An analyzing of the maps for Vend and Cambrian times allows us to reconsider a few stages of kinematic scenario of late Precambrian - early Paleozoic Arctida U Europe collision. 1) Old interpretation: Arctida was considered as an isolated paleocontinent during early Paleozoic time. New interpretation: during the early Paleozoic Arctida together Europe (Baltia) were assembled into a paleo- continent named us Arcteurope. This conclusion is based on excellent coincidence of Paleozoic paleomagnetic poles of the Kara block (which is a part of Arctida) and Europe

  16. Water-quality assessment of the Cambrian-Ordovician aquifer system in the northern Midwest, United States

    USGS Publications Warehouse

    Wilson, John T.

    2012-01-01

    This report provides a regional assessment of groundwater quality of the Cambrian-Ordovician aquifer system, based primarily on raw water samples collected by the NAWQA Program during 1995 through 2007. The NAWQA Program has published findings in local study-unit reports encompassing parts of the Cambrian-Ordovician aquifer system. Data collected from the aquifer system were used in national synthesis reports on selected topics such as specific water-quality constituent classes, well type, or aquifer material; however, a synthesis of groundwater quality at the principal aquifer scale has not been completed and is therefore the major purpose of this report. Water samples collected by the NAWQA Program were analyzed for various classes of characteristics including physical properties, major ions, trace elements, nutrients and dissolved organic carbon, radionuclides (tritium, radon, and radium), pesticides, and volatile organic compounds. Subsequent sections of this report provide discussions on these classes of characteristics. The assessment objectives of this report are to (1) summarize constituent concentrations and compare them to human-health benchmarks and non-health guidelines; (2) determine the geographic distribution of constituent concentrations and relate them to various factors such as confining conditions, well type, land use, and groundwater age; and (3) evaluate near-decadal-scale changes in nitrate concentrations and pesticide detections. The most recent sample collected from each well by the NAWQA Program was used for most analyses. Near-decadal-scale changes in nitrate concentrations and pesticide detections were evaluated for selected well networks by using the most recent sample from each well and comparing it to the results from a sample collected 7 or 11 years earlier. Because some of the NAWQA well networks provide a limited areal coverage of the aquifer system, data for raw water samples from other USGS sources and state agencies were included

  17. The Early Miocene Climatic Optimum (18-16 Ma): Stable Isotope and Mg/Ca Records from ODP Leg 189 Site 1168.

    NASA Astrophysics Data System (ADS)

    Syed, S.; Pekar, S.

    2008-12-01

    Ice volume estimates for the late early Miocene (~18-16 Ma) were derived from paired oxygen isotope records and Mg/Ca ratios from ODP Site 1168, which is located on the southwest slope of Tasmania. These records indicate the presence of a dynamic ice sheet in Antarctica, with ice-volume estimates up to present day levels occurring with relatively warm bottom water temperatures during isotope events Mi1b (17.9-17.6 Ma) and Mi2 (16.2 Ma). These records also indicate ice-volume decreased significantly during the Early Miocene Climatic Optimum ~17.2 to 16.4 Ma suggesting a near complete collapse of the East Antarctic Ice Sheet, based on an approximately 1‰ decrease in oxygen isotope value of seawater. Bottom water temperatures (BWT) derived from Mg/Ca ratios indicate temperature varied from ~8°C to 3°C, during the early Miocene, with the warmest BWT's occurring during glacial maxima and lowest occurring during glacial minima. Mg/Ca records from other records also indicate ice-volume increases coinciding with deep sea warming. These records suggest Antarctic glaciation may have been influenced by the moisture input by warm saline deep waters (WSDW) originating from the Indian Ocean/Tethys Sea. These WSDW would become entrained and ultimately upwell near Antarctica, resulting in delivering increased moisture/snowfall and therefore increased ice volume on the Antarctic continent. However, an alternative interpretation of the records could be that temperature estimates derived from Mg/Ca ratios may be over estimating the magnitude of temperature changes, thus resulting in an overestimation of ice-volume changes.

  18. Early Holocene humidity patterns in the Iberian Peninsula reconstructed from lake, pollen and speleothem records

    NASA Astrophysics Data System (ADS)

    Morellón, Mario; Aranbarri, Josu; Moreno, Ana; González-Sampériz, Penélope; Valero-Garcés, Blas L.

    2018-02-01

    Comparison of selected, well-dated, lacustrine, speleothem and terrestrial pollen records spanning the Holocene onset and the Early Holocene (ca. 11.7-8 cal kyrs BP) in the Iberian Peninsula shows large hydrological fluctuations and landscape changes with a complex regional pattern in timing and intensity. Marine pollen records from Alboran, the Mediterranean and off shore Atlantic sites show a step-wise increase in moisture and forest during this transition. However, available continental records point to two main patterns of spatial and temporal hydrological variability: i) Atlantic-influenced sites located at the northwestern areas (Enol, Sanabria, Lucenza, PRD-4), characterized by a gradual increase in humidity from the end of the Younger Dryas to the Mid Holocene, similarly to most North Atlantic records; and ii) continental and Mediterranean-influenced sites (Laguna Grande, Villarquemado, Fuentillejo, Padul, Estanya, Banyoles, Salines), with prolonged arid conditions of variable temporal extension after the Younger Dryas, followed by an abrupt increase in moisture at 10-9 cal kyrs BP. Different local climate conditions influenced by topography or the variable sensitivity (gradual versus threshold values) of the proxies analyzed in each case are evaluated. Vegetation composition (conifers versus mesothermophilous taxa) and resilience would explain a subdued response of vegetation in central continental areas while in Mediterranean sites, insufficient summer moisture availability could not maintain high lake levels and promote mesophyte forest, in contrast to Atlantic-influenced areas. Comparison with available climate models, Greenland ice cores, North Atlantic marine sequences and continental records from Central and Northern Europe and the whole Mediterranean region underlines the distinctive character of the hydrological changes occurred in inner Iberia throughout the Early Holocene. The persistent arid conditions might be explained by the intensification

  19. Long distance transport of eclogite and blueschist during early Pacific Ocean subduction rollback

    NASA Astrophysics Data System (ADS)

    Tamblyn, Renee; Hand, Martin; Kelsey, David; Phillips, Glen; Anczkiewicz, Robert

    2017-04-01

    The Tasmanides in eastern Australia represent a period of continental crustal growth on the western margin of the Pacific Ocean associated with slab rollback from the Cambrian until the Triassic. During rollback numerical models predict that subduction products can become trapped in the forearc (Geyra et al., 2002), and can migrate with the trench as it retreats. In a long-lived subduction controlled regime such as the Tasmanides, this should result in an accumulation of subduction products with protracted geochronological and metamorphic histories. U-Pb, Lu-Hf, Sm-Nd and Ar-Ar geochronology and phase equilibria modelling of lawsonite-eclogite and garnet blueschist in the Southern New England Fold Belt in Australia demonstrate that high-P low-T rocks remained within a subduction setting for c. 40 Ma, from c. 500 to 460 Ma. High-P metamorphic rocks initially formed close to the Australian cratonic margin during the late Cambrian, and were subsequently transported over 1500 Ma oceanward, during which time subducted material continued to accumulate, resulting in the development of complex mélange which records eclogite and blueschist metamorphism and partial exhumation over 40 Ma. The duration of refrigerated metamorphism approximates the extensional evolution of the upper plate which culminated in the development of the Lachlan Fold Belt. The protracted record of eclogite and blueschist metamorphism indicates that rapid exhumation is not necessarily required for preservation of high-pressure metamorphic rocks from subduction systems. Reference: Gerya, T. V., Stockhert, B., & Perchuk, A. L. (2002). Exhumation of high-pressure metamorphic rocks in a subduction channel: A numerical simulation. Tectonics, 21(6), 6-1-6-19. doi:10.1029/2002tc001406

  20. Uranium isotopes distinguish two geochemically distinct stages during the later Cambrian SPICE event

    PubMed Central

    Dahl, Tais W.; Boyle, Richard A.; Canfield, Donald E.; Connelly, James N.; Gill, Benjamin C.; Lenton, Timothy M.; Bizzarro, Martin

    2015-01-01

    Anoxic marine zones were common in early Paleozoic oceans (542–400 Ma), and present a potential link to atmospheric pO2 via feedbacks linking global marine phosphorous recycling, primary production and organic carbon burial. Uranium (U) isotopes in carbonate rocks track the extent of ocean anoxia, whereas carbon (C) and sulfur (S) isotopes track the burial of organic carbon and pyrite sulfur (primary long-term sources of atmospheric oxygen). In combination, these proxies therefore reveal the comparative dynamics of ocean anoxia and oxygen liberation to the atmosphere over million-year time scales. Here we report high-precision uranium isotopic data in marine carbonates deposited during the Late Cambrian ‘SPICE’ event, at ca. 499 Ma, documenting a well-defined −0.18‰ negative δ238U excursion that occurs at the onset of the SPICE event’s positive δ13C and δ34S excursions, but peaks (and tails off) before them. Dynamic modelling shows that the different response of the U reservoir cannot be attributed solely to differences in residence times or reservoir sizes - suggesting that two chemically distinct ocean states occurred within the SPICE event. The first ocean stage involved a global expansion of euxinic waters, triggering the spike in U burial, and peaking in conjunction with a well-known trilobite extinction event. During the second stage widespread euxinia waned, causing U removal to tail off, but enhanced organic carbon and pyrite burial continued, coinciding with evidence for severe sulfate depletion in the oceans (Gill et al., 2011). We discuss scenarios for how an interval of elevated pyrite and organic carbon burial could have been sustained without widespread euxinia in the water column (both non-sulfidic anoxia and/or a more oxygenated ocean state are possibilities). Either way, the SPICE event encompasses two different stages of elevated organic carbon and pyrite burial maintained by high nutrient fluxes to the ocean, and potentially

  1. Orogen styles in the East African Orogen: A review of the Neoproterozoic to Cambrian tectonic evolution☆

    PubMed Central

    Fritz, H.; Abdelsalam, M.; Ali, K.A.; Bingen, B.; Collins, A.S.; Fowler, A.R.; Ghebreab, W.; Hauzenberger, C.A.; Johnson, P.R.; Kusky, T.M.; Macey, P.; Muhongo, S.; Stern, R.J.; Viola, G.

    2013-01-01

    The East African Orogen, extending from southern Israel, Sinai and Jordan in the north to Mozambique and Madagascar in the south, is the world́s largest Neoproterozoic to Cambrian orogenic complex. It comprises a collage of individual oceanic domains and continental fragments between the Archean Sahara–Congo–Kalahari Cratons in the west and Neoproterozoic India in the east. Orogen consolidation was achieved during distinct phases of orogeny between ∼850 and 550 Ma. The northern part of the orogen, the Arabian–Nubian Shield, is predominantly juvenile Neoproterozoic crust that formed in and adjacent to the Mozambique Ocean. The ocean closed during a protracted period of island-arc and microcontinent accretion between ∼850 and 620 Ma. To the south of the Arabian Nubian Shield, the Eastern Granulite–Cabo Delgado Nappe Complex of southern Kenya, Tanzania and Mozambique was an extended crust that formed adjacent to theMozambique Ocean and experienced a ∼650–620 Ma granulite-facies metamorphism. Completion of the nappe assembly around 620 Ma is defined as the East African Orogeny and was related to closure of the Mozambique Ocean. Oceans persisted after 620 Ma between East Antarctica, India, southern parts of the Congo–Tanzania–Bangweulu Cratons and the Zimbabwe–Kalahari Craton. They closed during the ∼600–500 Ma Kuungan or Malagasy Orogeny, a tectonothermal event that affected large portions of southern Tanzania, Zambia, Malawi, Mozambique, Madagascar and Antarctica. The East African and Kuungan Orogenies were followed by phases of post-orogenic extension. Early ∼600–550 Ma extension is recorded in the Arabian–Nubian Shield and the Eastern Granulite–Cabo Delgado Nappe Complex. Later ∼550–480 Ma extension affected Mozambique and southern Madagascar. Both extension phases, although diachronous,are interpreted as the result of lithospheric delamination. Along the strike of the East African Orogen, different geodynamic settings

  2. Precambrian-Cambrian provenance of Matinde Formation, Karoo Supergroup, northwestern Mozambique, constrained from detrital zircon U-Pb age and Lu-Hf isotope data

    NASA Astrophysics Data System (ADS)

    Bicca, Marcos Müller; Jelinek, Andrea Ritter; Philipp, Ruy Paulo; de Carvalho Lana, Cristiano; Alkmim, Ana Ramalho

    2018-02-01

    The Permian-Triassic time interval was a period of high sedimentation rates in the intracontinental Karoo rift basin of northwestern Mozambique, reflecting high exhumation rates in the surrounding high ground Precambrian-Cambrian basement and juxtaposed nappes. U-Pb LA-MC-ICPMS dating and Lu-Hf isotopic analysis of detrital zircons from the Late Permian-Early Triassic Matinde Formation of the Karoo Supergroup is used as a reliable proxy to map denudation patterns of source regions. Data allow discrimination of U-Pb age populations of ca. 1250-900 Ma, a secondary population between ca. 900-700 and a major contribution of ages around ca. 700-490 Ma. Zircon grains of the Mesoproterozoic age population present Mesoproterozoic (1000-1500 Ma) to Paleoproterozoic (1800-2300 Ma) Hf TDM ages, with positive (0 to +11) and negative εHf values (-3 to -15), respectively. The younger U-Pb age population also presents two different groups of zircon grains according to Lu-Hf isotopes. The first group comprise Paleoproterozoic (1800-2300 Ma) ages, with highly negative εHf values, between -10 and -22, and the second group exhibits Mesoproterozoic ages (1200-1500 Ma), with increased juvenile εHf values (ca. 0 to -5). These Hf isotopes reinforce the presence of unexposed ancient crust in this region. The oldest U-Pb age population resembles the late stages of Grenville Orogeny and the Rodinia Supercontinent geotectonic activity mostly represented by magmatic rocks, which are widely present in the basement of northern Mozambique. The juvenile Hf-isotope signature with an older age component is associated to rocks generated from subduction processes with crust assimilation by continental arcs, which we correlate to rocks of the Nampula Complex, south and east of the Moatize-Minjova Basin. The U-Pb ages between 900 and 700 Ma were correlated to the calc-alkaline magmatism registered in the Guro Suite, related to the breakup phase of Rodinia, and mark the western limit of the Moatize

  3. The Moon as a recorder of organic evolution in the early solar system: a lunar regolith analog study.

    PubMed

    Matthewman, Richard; Court, Richard W; Crawford, Ian A; Jones, Adrian P; Joy, Katherine H; Sephton, Mark A

    2015-02-01

    The organic record of Earth older than ∼3.8 Ga has been effectively erased. Some insight is provided to us by meteorites as well as remote and direct observations of asteroids and comets left over from the formation of the Solar System. These primitive objects provide a record of early chemical evolution and a sample of material that has been delivered to Earth's surface throughout the past 4.5 billion years. Yet an effective chronicle of organic evolution on all Solar System objects, including that on planetary surfaces, is more difficult to find. Fortunately, early Earth would not have been the only recipient of organic matter-containing objects in the early Solar System. For example, a recently proposed model suggests the possibility that volatiles, including organic material, remain archived in buried paleoregolith deposits intercalated with lava flows on the Moon. Where asteroids and comets allow the study of processes before planet formation, the lunar record could extend that chronicle to early biological evolution on the planets. In this study, we use selected free and polymeric organic materials to assess the hypothesis that organic matter can survive the effects of heating in the lunar regolith by overlying lava flows. Results indicate that the presence of lunar regolith simulant appears to promote polymerization and, therefore, preservation of organic matter. Once polymerized, the mineral-hosted newly formed organic network is relatively protected from further thermal degradation. Our findings reveal the thermal conditions under which preservation of organic matter on the Moon is viable.

  4. Near-Record Early Snowmelt and Signs of Environmental Change in Barrow, Alaska

    NASA Astrophysics Data System (ADS)

    Stanitski, D.; Cox, C.; Sweeney, C.; Divoky, G.; George, C.; Stone, R.

    2015-12-01

    The 2015 spring transition in Barrow, AK, was notable with the second earliest date of snow melt on record (JD148, May 28) and earliest ice free conditions on a local lagoon (JD178, June 27). The 73-year time series from the NOAA Global Monitoring Division's Barrow Observatory (BRW) has shown a trend toward earlier spring snowmelt, reinforced in 2015. Anomalous early snowmelt was also observed at nearby Cooper Island where a colony of sea birds, the Black Guillemot, nests each year once snow disappears. The appearance of "first egg" is well correlated with the date of snowmelt at BRW (Fig. 1), as is the ice-out date at the Isaktoak Lagoon (ISK). In 2015, the first egg was observed on JD159 (June 8), the earliest in the 40-year record (source: Friends of Cooper Island, http://cooperisland.org/). Each day of advance in the melt date at BRW results in an annual net radiation increase at the surface of about 1%. The documented changes can influence biogeochemical cycles, permafrost temperatures, and potentially the release of stored carbon. By mid July 2015, a 1°C increase in soil temperature at 0.5-m depth was measured compared to prior years; therefore, the active layer is expected to be unusually deep by autumn. The anomalous warmth that prevailed during spring 2015 can be attributed, in part, to atmospheric circulation, influenced by two typhoons in the North Pacific and the onset of El Niño. Warming was likely amplified locally as the early melting of snow increased absorption of solar radiation. Key factors influencing the trend toward earlier spring snowmelt will be presented as well as those contributing to the anomalous 2015 spring at BRW (e.g., winter snowfall, cloud cover, advection, local sea ice extent), and the impact early melt had on the 2015 summer surface radiation budget. Analysis of interactions underlying this anomaly will aid in developing strategies for improving predictability of interannual variability of the melt season and long-term change.

  5. Records of auroral candidates and sunspots in Rikkokushi, chronicles of ancient Japan from early 7th century to 887

    NASA Astrophysics Data System (ADS)

    Hayakawa, Hisashi; Iwahashi, Kiyomi; Tamazawa, Harufumi; Ebihara, Yusuke; Kawamura, Akito Davis; Isobe, Hiroaki; Namiki, Katsuko; Shibata, Kazunari

    2017-12-01

    We present the results of the surveys on sunspots and auroral candidates in Rikkokushi, Japanese official histories from the early 7th century to 887, to review the solar and auroral activities. In total, we found one sunspot record and 13 auroral candidates in Rikkokushi. We then examine the records of the sunspots and auroral candidates, compare the auroral candidates with the lunar phase to estimate their reliability, and compare the records of the sunspots and auroral candidates with the contemporary total solar irradiance reconstructed from radioisotope data. We also identify the locations of the observational sites to review possible equatorward expansion of the auroral oval. These discussions suggest a major gap in auroral candidates from the late 7th to early 9th centuries, which includes the candidate of the grand minimum reconstructed from the radioisotope data, a similar tendency as the distributions of sunspot records in contemporary China, and a relatively high magnetic latitude of observational sites with a higher potential for observing aurorae more frequently than at present.

  6. Accuracy and Efficiency of Recording Pediatric Early Warning Scores Using an Electronic Physiological Surveillance System Compared With Traditional Paper-Based Documentation

    PubMed Central

    Sefton, Gerri; Lane, Steven; Killen, Roger; Black, Stuart; Lyon, Max; Ampah, Pearl; Sproule, Cathryn; Loren-Gosling, Dominic; Richards, Caitlin; Spinty, Jean; Holloway, Colette; Davies, Coral; Wilson, April; Chean, Chung Shen; Carter, Bernie; Carrol, E.D.

    2017-01-01

    Pediatric Early Warning Scores are advocated to assist health professionals to identify early signs of serious illness or deterioration in hospitalized children. Scores are derived from the weighting applied to recorded vital signs and clinical observations reflecting deviation from a predetermined “norm.” Higher aggregate scores trigger an escalation in care aimed at preventing critical deterioration. Process errors made while recording these data, including plotting or calculation errors, have the potential to impede the reliability of the score. To test this hypothesis, we conducted a controlled study of documentation using five clinical vignettes. We measured the accuracy of vital sign recording, score calculation, and time taken to complete documentation using a handheld electronic physiological surveillance system, VitalPAC Pediatric, compared with traditional paper-based charts. We explored the user acceptability of both methods using a Web-based survey. Twenty-three staff participated in the controlled study. The electronic physiological surveillance system improved the accuracy of vital sign recording, 98.5% versus 85.6%, P < .02, Pediatric Early Warning Score calculation, 94.6% versus 55.7%, P < .02, and saved time, 68 versus 98 seconds, compared with paper-based documentation, P < .002. Twenty-nine staff completed the Web-based survey. They perceived that the electronic physiological surveillance system offered safety benefits by reducing human error while providing instant visibility of recorded data to the entire clinical team. PMID:27832032

  7. Accuracy and Efficiency of Recording Pediatric Early Warning Scores Using an Electronic Physiological Surveillance System Compared With Traditional Paper-Based Documentation.

    PubMed

    Sefton, Gerri; Lane, Steven; Killen, Roger; Black, Stuart; Lyon, Max; Ampah, Pearl; Sproule, Cathryn; Loren-Gosling, Dominic; Richards, Caitlin; Spinty, Jean; Holloway, Colette; Davies, Coral; Wilson, April; Chean, Chung Shen; Carter, Bernie; Carrol, E D

    2017-05-01

    Pediatric Early Warning Scores are advocated to assist health professionals to identify early signs of serious illness or deterioration in hospitalized children. Scores are derived from the weighting applied to recorded vital signs and clinical observations reflecting deviation from a predetermined "norm." Higher aggregate scores trigger an escalation in care aimed at preventing critical deterioration. Process errors made while recording these data, including plotting or calculation errors, have the potential to impede the reliability of the score. To test this hypothesis, we conducted a controlled study of documentation using five clinical vignettes. We measured the accuracy of vital sign recording, score calculation, and time taken to complete documentation using a handheld electronic physiological surveillance system, VitalPAC Pediatric, compared with traditional paper-based charts. We explored the user acceptability of both methods using a Web-based survey. Twenty-three staff participated in the controlled study. The electronic physiological surveillance system improved the accuracy of vital sign recording, 98.5% versus 85.6%, P < .02, Pediatric Early Warning Score calculation, 94.6% versus 55.7%, P < .02, and saved time, 68 versus 98 seconds, compared with paper-based documentation, P < .002. Twenty-nine staff completed the Web-based survey. They perceived that the electronic physiological surveillance system offered safety benefits by reducing human error while providing instant visibility of recorded data to the entire clinical team.

  8. The Moon as a Recorder of Organic Evolution in the Early Solar System: A Lunar Regolith Analog Study

    PubMed Central

    Court, Richard W.; Crawford, Ian A.; Jones, Adrian P.; Joy, Katherine H.; Sephton, Mark A.

    2015-01-01

    Abstract The organic record of Earth older than ∼3.8 Ga has been effectively erased. Some insight is provided to us by meteorites as well as remote and direct observations of asteroids and comets left over from the formation of the Solar System. These primitive objects provide a record of early chemical evolution and a sample of material that has been delivered to Earth's surface throughout the past 4.5 billion years. Yet an effective chronicle of organic evolution on all Solar System objects, including that on planetary surfaces, is more difficult to find. Fortunately, early Earth would not have been the only recipient of organic matter–containing objects in the early Solar System. For example, a recently proposed model suggests the possibility that volatiles, including organic material, remain archived in buried paleoregolith deposits intercalated with lava flows on the Moon. Where asteroids and comets allow the study of processes before planet formation, the lunar record could extend that chronicle to early biological evolution on the planets. In this study, we use selected free and polymeric organic materials to assess the hypothesis that organic matter can survive the effects of heating in the lunar regolith by overlying lava flows. Results indicate that the presence of lunar regolith simulant appears to promote polymerization and, therefore, preservation of organic matter. Once polymerized, the mineral-hosted newly formed organic network is relatively protected from further thermal degradation. Our findings reveal the thermal conditions under which preservation of organic matter on the Moon is viable. Key Words: Moon—Regolith—Organic preservation—Biomarkers. Astrobiology 15, 154–168. PMID:25615648

  9. High resolution windows into early Holocene climate: Sr/(Ca) coral records from the Huon Peninsula

    NASA Astrophysics Data System (ADS)

    McCulloch, Malcolm; Mortimer, Graham; Esat, Tezer; Xianhua, Li; Pillans, Brad; Chappell, John

    1996-02-01

    High-precision measurements of Sr/Ca ratios are reported for Porites corals from the uplifted Holocene coral terraces at Huon Peninsula, Papua New Guinea. The early Holocene Porites have UTh mass spectrometric ages of 8920 ± 60 yr and 7370 ± 50 yr, and δ 234U(t) values of 145 ± 2, similar to modern seawater. The Sr/Ca coral records provide 5-6 year high resolution (near weekly) time windows into early Holocene sea surface temperatures. Seasonal temperature fluctuations are generally in the range of ± 1°C, with occasional excursions of ± 2°C, which may indicate the more frequent recurrence of very strong ENSO (El Niño-Southern Oscillation) events. Mean annual Sr/Ca temperatures of 24.2 ± 1.1°C and 22.9 ± 0.8°C have been obtained, which are ˜ 2-3°C cooler than that exhibited by a modern Porites. These results indicate that, during the early Holocene, the equatorial western Pacific ocean was at least several degrees cooler than present-day temperatures. This is consistent with late glacial coral records from the Caribbean that indicate lower (˜ 6°C) sea surface temperatures for the equatorial oceans. The Huon Peninsula corals also indicate that SSTs were several degrees cooler than those in the Caribbean during the early Holocene. Thus, although the northern hemisphere summer radiation maximum occurred at ˜ 10 ka, there appears to have been a significant lag in the response of the equatorial western Pacific ocean to this warming. Cooler early Holocene sea surface temperatures in the western Pacific may have been due to changing patterns of ocean-atmosphere circulation, resulting from the exposure of large areas of continental shelf in the southeast Asia region, a consequence of lower glacial sea levels. It is likely that ocean temperatures in the Huon Peninsula were influenced by the opening at ˜ 7 ka of the Torres Strait, that now separates New Guinea from the Australian mainland.

  10. A stem-group cnidarian described from the mid-Cambrian of China and its significance for cnidarian evolution.

    PubMed

    Park, Tae-yoon; Woo, Jusun; Lee, Dong-Jin; Lee, Dong-Chan; Lee, Seung-bae; Han, Zuozhen; Chough, Sung Kwun; Choi, Duck K

    2011-08-23

    Palaeontological data of extinct groups often sheds light on the evolutionary sequences leading to extant groups, but has failed to resolve the basal metazoan phylogeny including the origin of the Cnidaria. Here we report the occurrence of a stem-group cnidarian, Cambroctoconus orientalis gen. et sp. nov., from the mid-Cambrian of China, which is a colonial organism with calcareous octagonal conical cup-shaped skeletons. It bears cnidarian features including longitudinal septa arranged in octoradial symmetry and colonial occurrence, but lacks a jelly-like mesenchyme. Such morphological characteristics suggest that the colonial occurrence with polyps of octoradial symmetry is the plesiomorphic condition of the Cnidaria and appeared earlier than the jelly-like mesenchyme during the course of evolution. © 2011 Macmillan Publishers Limited. All rights reserved.

  11. A stem-group cnidarian described from the mid-Cambrian of China and its significance for cnidarian evolution

    PubMed Central

    Park, Tae-yoon; Woo, Jusun; Lee, Dong-Jin; Lee, Dong-Chan; Lee, Seung-bae; Han, Zuozhen; Chough, Sung Kwun; Choi, Duck K.

    2011-01-01

    Palaeontological data of extinct groups often sheds light on the evolutionary sequences leading to extant groups, but has failed to resolve the basal metazoan phylogeny including the origin of the Cnidaria. Here we report the occurrence of a stem-group cnidarian, Cambroctoconus orientalis gen. et sp. nov., from the mid-Cambrian of China, which is a colonial organism with calcareous octagonal conical cup-shaped skeletons. It bears cnidarian features including longitudinal septa arranged in octoradial symmetry and colonial occurrence, but lacks a jelly-like mesenchyme. Such morphological characteristics suggest that the colonial occurrence with polyps of octoradial symmetry is the plesiomorphic condition of the Cnidaria and appeared earlier than the jelly-like mesenchyme during the course of evolution. PMID:21863009

  12. Cambrian-Ordovician Knox production in Ohio: Three case studies of structural-stratigraphic traps

    USGS Publications Warehouse

    Riley, R.A.; Wicks, J.; Thomas, Joan

    2002-01-01

    The Knox Dolomite (Cambrian-Ordovician) in Ohio consists of a mixed carbonate-siliciclastic sequence deposited in a tidal-flat to shallow-marine environment along a broad continental shelf. Knox hydrocarbon production occurs in porous sandstone and dolomite reservoirs in the Copper Ridge dolomite, Rose Run sandstone, and Beekmantown dolomite. In Ohio, historical Knox exploration and development have been focused on paleogeomorphic traps within the prolific Morrow Consolidated field, and more recently, within and adjacent to the Rose Run subcrop. Although these paleogeomorphic traps have yielded significant Knox production, structural and stratigraphic traps are being largely ignored. Three Knox-producing pools demonstrate structural and stratigraphic traps: the Birmingham-Erie pool in southern Erie and southwestern Lorain counties, the South Canaan pool in northern Wayne County, and the East Randolph pool in south-central Portage County. Enhanced porosity and permeability from fractures, as evident in the East Randolph pool, are also an underexplored mechanism for Knox hydrocarbon accumulation. An estimated 800 bcf of gas from undiscovered Knox resources makes the Knox one of the most attractive plays in the Appalachian basin.

  13. Eukaryotic organisms in Proterozoic oceans

    PubMed Central

    Knoll, A.H; Javaux, E.J; Hewitt, D; Cohen, P

    2006-01-01

    The geological record of protists begins well before the Ediacaran and Cambrian diversification of animals, but the antiquity of that history, its reliability as a chronicle of evolution and the causal inferences that can be drawn from it remain subjects of debate. Well-preserved protists are known from a relatively small number of Proterozoic formations, but taphonomic considerations suggest that they capture at least broad aspects of early eukaryotic evolution. A modest diversity of problematic, possibly stem group protists occurs in ca 1800–1300 Myr old rocks. 1300–720 Myr fossils document the divergence of major eukaryotic clades, but only with the Ediacaran–Cambrian radiation of animals did diversity increase within most clades with fossilizable members. While taxonomic placement of many Proterozoic eukaryotes may be arguable, the presence of characters used for that placement is not. Focus on character evolution permits inferences about the innovations in cell biology and development that underpin the taxonomic and morphological diversification of eukaryotic organisms. PMID:16754612

  14. Oxygen and Early Animal Evolution

    NASA Astrophysics Data System (ADS)

    Xiao, S.

    2012-12-01

    It is often hypothesized that the rise of animals was triggered by an increase in O2 levels in the atmosphere and oceans. However, this hypothesis is remarkably difficult to test, because the timing of animal divergences is poorly resolved, the physiology of early animals is often unknown, estimates of past pO2 levels come with large error bars, and causal relationships between oxygenation and animal evolution are difficult to establish. Nonetheless, existing phylogenetic, paleontological, and geochemical data indicate that the evolution of macroscopic animals and motile macrometazoans with energetically expensive lifestyles may be temporally coupled with ocean oxygenation events in the Ediacaran Period. Thus, it is plausible that ocean oxygenation may have been a limiting factor in the early evolution of macroscopic, complex, and metabolically aggressive animals (particularly bilaterian animals). However, ocean oxygenation and animal evolution were likely engaged in two-way interactions: Ediacaran oxygenation may have initially lifted a physiological barrier for the evolution of animal size, motility, and active lifestyles, but subsequent animal diversification in the Paleozoic may have also changed oceanic redox structures. Viewed in a broader context, the early evolutionary history of animals was contingent upon a series of events, including genetic preparation (developmental genetics), environmental facilitation (oceanic oxygenation), and ecological escalation (Cambrian explosion), but the rise of animals to ecological importance also had important geobiological impacts on oceanic redox structures, sedimentary fabrics, and global geochemical cycles.

  15. Proposed stratotype for the base of the highest Cambrian stage at the first appearance datum of Cordylodus andresi, Lawson Cove section, Utah, USA

    USGS Publications Warehouse

    Miller, J.F.; Ethington, Raymond L.; Evans, K.R.; Holmer, L.E.; Loch, James D.; Popov, L.E.; Repetski, J.E.; Ripperdan, R.L.; Taylor, John F.

    2006-01-01

    We propose a candidate for the Global Standard Stratotype-section and Point (GSSP) for the base of the highest stage of the Furongian Series of the Cambrian System. The section is at Lawson Cove in the Ibex area of Millard County, Utah, USA. The marker horizon is the first appearance datum (FAD) of the conodont Cordylodus andresi Viira et Sergeyeva in Kaljo et al. [Kaljo, D., Borovko, N., Heinsalu, H., Khazanovich, K., Mens, K., Popov, L., Sergeyeva, S., Sobolevskaya, R., Viira, V., 1986. The Cambrian-Ordovician boundary in the Baltic-Ladoga clint area (North Estonia and Leningrad Region, USSR). Eesti NSV Teaduste Akadeemia Toimetised. Geologia 35, 97-108]. At this section and elsewhere this horizon also is the FAD of the trilobite Eurekia apopsis (Winston et Nicholls, 1967). This conodont characterizes the base of the Cordylodus proavus Zone, which has been recognized in many parts of the world. This trilobite characterizes the base of the Eurekia apopsis Zone, which has been recognized in many parts of North America. The proposed boundary is 46.7 m above the base of the Lava Dam Member of the Notch Peak Formation at the Lawson Cove section. Brachiopods, sequence stratigraphy, and carbon-isotope geochemistry are other tools that characterize this horizon and allow it to be recognized in other areas. ?? 2006 Nanjing Institute of Geology and Palaeontology, CAS.

  16. First data on age of metarhyolites from the Turan Group of the Bureya Terrane, eastern part of the Central Asian Foldbelt

    NASA Astrophysics Data System (ADS)

    Sorokin, A. A.; Smirnov, Yu. V.; Smirnova, Yu. N.; Kudryashov, N. M.

    2011-07-01

    The U-Pb geochronological studies showed that metarhyolites from the Turan Group of the Bureya (Turan) Terrane to the east of the Central Asian Foldbelt are Middle Cambrian (504 ± 8 Ma), not Neoproterozoic in age, as was suggested before. Metarhyolites are younger than the Early Cambrian terrigenous-carbonate sediments from this terrane characterized by the Atdabanian archaeochyatid. Considering that volcanic rocks have features of intraplate origin, it may be assumed that their formation corresponds to the breakup of the Early Paleozoic passive continental margin.

  17. Seaweed morphology and ecology during the great animal diversification events of the early Paleozoic: A tale of two floras.

    PubMed

    LoDuca, S T; Bykova, N; Wu, M; Xiao, S; Zhao, Y

    2017-07-01

    Non-calcified marine macroalgae ("seaweeds") play a variety of key roles in the modern Earth system, and it is likely that they were also important players in the geological past, particularly during critical transitions such as the Cambrian Explosion (CE) and the Great Ordovician Biodiversification Event (GOBE). To investigate the morphology and ecology of seaweeds spanning the time frame from the CE through the GOBE, a carefully vetted database was constructed that includes taxonomic and morphometric information for non-calcified macroalgae from 69 fossil deposits. Analysis of the database shows a pattern of seaweed history that can be explained in terms of two floras: the Cambrian Flora and the Ordovician Flora. The Cambrian Flora was dominated by rather simple morphogroups, whereas the Ordovician Flora, which replaced the Cambrian Flora in the Ordovician and extended through the Silurian, mainly comprised comparatively complex morphogroups. In addition to morphogroup representation, the two floras show marked differences in taxonomic composition, morphospace occupation, functional-form group representation, and life habit, thereby pointing to significant morphological and ecological changes for seaweeds roughly concomitant with the GOBE and the transition from the Cambrian to Paleozoic Evolutionary Faunas. Macroalgal changes of a similar nature and magnitude, however, are not evident in concert with the CE, as the Cambrian Flora consists largely of forms established during the Ediacaran. The cause of such a lag in macroalgal morphological diversification remains unclear, but an intriguing possibility is that it signals a previously unknown difference between the CE and GOBE with regard to the introduction of novel grazing pressures. The consequences of the establishment of the Ordovician Flora for shallow marine ecosystems and Earth system dynamics remain to be explored in detail but could have been multifaceted and potentially include impacts on the global

  18. Soft-sediment deformation structures in Cambrian Series 2 tidal deposits (NW Estonia): implications for identifying endogenic triggering mechanisms in ancient sedimentary record

    NASA Astrophysics Data System (ADS)

    Põldsaar, Kairi

    2015-04-01

    Soft-sediment deformation structures (SSDS) are documented in several horizons within silt- and sandstones of the Cambrian Series 2 (Dominopolian Stage) Tiskre Formation, and some in the below-deposited argillaceous deposits of the Lükati Formation (northern part of the Baltoscandian Palaeobasin, NW Estonia). The aim of this study was to map, describe, and analyze these deformation features, discuss their deformation mechanism and possible triggers. Load structures (simple load casts, pillows, flame structures, convoluted lamination) with varying shapes and sizes occur in the Tiskre Fm in sedimentary interfaces within medium-bedded peritidal rhythmites (siltstone-argillaceous material) as well as within up to 3 m thick slightly seaward inclined stacked sandstone sequences. Homogenized beds, dish-and-pillar structures, and severely deformed bedding are also found within these stacked units and within a large tidal runoff channel infill. Autoclastic breccias and water-escape channels are rare and occur only in small-scale -- always related to thin, horizontal tidal laminae. Profound sedimentary dykes, sand volcanoes, and thrust faults, which are often related to earthquake triggered soft sediment deformation, were not observed within the studied intervals. Deformation horizon or horizons with large flat-topped pillows often with elongated morphologies occur at or near the boundary between the Tiskre and Lükati formations. Deformation mechanisms identified in this study for the various deformation types are gravitationally unstable reversed density gradient (especially in case of load features that are related to profound sedimentary interfaces) and lateral shear stress due to sediment current drag (in case of deformation structures that not related to loading at any apparent sedimentary interface). Synsedimentary liquefaction was identified as the primary driving force in most of the observed deformation horizons. Clay thixotropy may have contributed in the

  19. Spoken Records. Third Edition.

    ERIC Educational Resources Information Center

    Roach, Helen

    Surveying 75 years of accomplishment in the field of spoken recording, this reference work critically evaluates commercially available recordings selected for excellence of execution, literary or historical merit, interest, and entertainment value. Some types of spoken records included are early recording, documentaries, lectures, interviews,…

  20. A comparative assessment of the role of anoxia during the Cambrian SPICE event

    NASA Astrophysics Data System (ADS)

    LeRoy, M.; Gill, B. C.; Sperling, E. A.

    2017-12-01

    The Cambrian SPICE (Steptoean Positive Carbon Isotope Excursion) is recognized as a major oceanographic event recorded in globally-distributed stratigraphic sections as positive isotopic excursions in marine carbonates (δ13Ccarb), organic matter (δ13Corg), sulfate (δ34SCAS) and pyrite (δ34Spy). A proposed mechanism for these observed isotopic trends is that a transient increase in the areal extent of anoxic conditions within the oceans fostered enhanced burial of organic carbon and pyrite. However, direct sedimentary (e.g., abundant black shale) and geochemical (e.g., redox proxy) evidence for such a shift is scant. While the antiquity of this event is likely responsible for loss of much of this evidence, through destruction by tectonic processes, a number of stratigraphic successions suitable for investigating this hypothesis exist. Here we explore the relationship between anoxia and the SPICE using previously published and novel data generated from core material from three sedimentary basins distributed along the margins of the Iapetus Ocean. The units studied are: the Nolichucky Formation of eastern Laurentia, the Alum Shale of Baltica, and the Outwoods Shale of Avalonia. Our iron speciation data indicate consistently oxic conditions prior to the SPICE along Laurentia, while Avalonia was oxic with intervals of anoxia, and Baltica was persistently anoxic. With the initiation of the SPICE, anoxic conditions intensified in Laurentia and Avalonia and continued to persist in Baltica. This redox shift was coupled with a sharp rise in δ34Spy at all three locations. Average total organic carbon (TOC) showed little change in relation to the SPICE at the Laurentia location ( 0.15 wt%), but showed an increase in conjunction with the excursion in both the Avalonia ( 1 to 1.5 wt%) and Baltica ( 8 to 12 wt%) sections. Large differences in nutrient availability and sedimentation rates are likely responsible for the between-site disparity in TOC, while overall, anoxia

  1. Younger Dryas to Early Holocene paleoclimate in Cantabria (N Spain): Constraints from speleothem Mg, annual fluorescence banding and stable isotope records

    NASA Astrophysics Data System (ADS)

    Rossi, Carlos; Bajo, Petra; Lozano, Rafael P.; Hellstrom, John

    2018-07-01

    The Younger Dryas (YD) stadial represents the most abrupt climate change of the Earth's recent history. Thus, understanding its causes and different local responses is relevant for Quaternary paleoclimatology. We present a speleothem high-resolution proxy record of the Lateglacial to Early Holocene paleoclimate of the Cantabrian Cordillera (N Spain), a strategic location to evaluate the influence of North Atlantic events such as the YD on South-Western Europe. Fluorescence lamination, growth-rate, stable-isotope, and [Mg] records from stalagmite SIR-1 were dated using an age-depth model constrained by U-Th dates and annual-lamina counting. The YD is recorded as a prominent positive δ13C excursion whose chronology (12.95 ± 0.14 to 11.62 ± 0.16 ka) and shape closely agree with the GS-1 stadial as defined in Greenland ice, supporting the event synchronicity in both areas. A colder and drier YD climate limited soil productivity and dripwater availability, leading to higher δ13C and [Mg], reduced growth rate, and virtually absent fluorescence lamination. The early YD record (until ∼12.5 ka) reflects increasing aridity, whereas the late YD (from ∼12.2 ka on) shows the opposite trend. At the YD boundaries, temperature changes influenced the [Mg] record by modifying the Mg partition into calcite. However, this effect was superseded by major changes in dripwater Mg/Ca linked to rainfall variations. During the Early Holocene, the Arnero Sierra was forested and had a relatively warm and humid seasonal climate, indicated in SIR-1 by higher growth rates, lower δ13C and [Mg], and well-developed fluorescent lamination. Similar to other high-resolution stalagmitic records of the Cordillera, from ∼8.5 to 8.0 ka SIR-1 reflects a temporary trend of increasing aridity.

  2. An Early Holocene Record of Cimex (Hemiptera: Cimicidae) From Western North America.

    PubMed

    Adams, Martin E; Jenkins, Dennis L

    2017-07-01

    The subfossil remains of 14 cimicids (Hemiptera: Cimicidae) were recovered during archaeological investigations of the Paisley Five Mile Point Cave site (35LK3400), an exceptionally well-dated (n = 229 radiocarbon dates) late Pleistocene-early Holocene rock shelter site in south-central Oregon. Nine of the specimens have been assigned to three modern species of Nearctic Cimicidae-Cimex antennatus Usinger & Ueshima, Cimex latipennis Usinger & Ueshima, and Cimex pilosellus (Horváth)-whereas the remaining five individuals were too fragmentary to positively identify. The chronology of the insect assemblage puts one specimen at circa 5,100 calibrated years before present (cal. yr BP), and the remaining 13 range in age from 9,400 to almost 11,000 cal. yr BP. Although fossil and subfossil cimicid remains have been recovered at other archaeological sites, the fossil record for bed bugs is largely undocumented. The Paisley Caves specimens thus far represent the oldest remains of the genus in probable contact with humans on record. © The Authors 2017. Published by Oxford University Press on behalf of Entomological Society of America. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

  3. Pine nut use in the Early Holocene and beyond: The danger cave archaeobotanical record

    USGS Publications Warehouse

    Rhode, D.; Madsen, D.B.

    1998-01-01

    Nuts of limber pine (Pinus flexilis) from Early Holocene strata in Danger Cave, Utah, are distinguishable by seed-coat sculpturing from pine nuts of single-needled pinyon (Pinus monophylla), which occur in strata dating <7000 years BP. Owls and other taphonomic agents may deposit pine nuts in archaeological sites, but the morphology of the pine nuts in Danger Cave strongly indicate they were deposited by human foragers who brought small quantities with them for food for at least the last 7500 years. Large-scale transport of pine nuts to Danger Cave from distant hinterlands is unlikely, however. The seamless transition from limber pine to pinyon pine nuts in the Danger Cave record suggests that foragers who had utilized limber pine as a food resource easily switched to using pinyon pine nuts when pinyon pine migrated into the region at the close of the Early Holocene.

  4. Solution to Darwin's dilemma: Discovery of the missing Precambrian record of life

    PubMed Central

    Schopf, J. William

    2000-01-01

    In 1859, in On the Origin of Species, Darwin broached what he regarded to be the most vexing problem facing his theory of evolution—the lack of a rich fossil record predating the rise of shelly invertebrates that marks the beginning of the Cambrian Period of geologic time (≈550 million years ago), an “inexplicable” absence that could be “truly urged as a valid argument” against his all embracing synthesis. For more than 100 years, the “missing Precambrian history of life” stood out as one of the greatest unsolved mysteries in natural science. But in recent decades, understanding of life's history has changed markedly as the documented fossil record has been extended seven-fold to some 3,500 million years ago, an age more than three-quarters that of the planet itself. This long-sought solution to Darwin's dilemma was set in motion by a small vanguard of workers who blazed the trail in the 1950s and 1960s, just as their course was charted by a few pioneering pathfinders of the previous century, a history of bold pronouncements, dashed dreams, search, and final discovery. PMID:10860955

  5. Solution to Darwin's dilemma: discovery of the missing Precambrian record of life

    NASA Technical Reports Server (NTRS)

    Schopf, J. W.

    2000-01-01

    In 1859, in On the Origin of Species, Darwin broached what he regarded to be the most vexing problem facing his theory of evolution-the lack of a rich fossil record predating the rise of shelly invertebrates that marks the beginning of the Cambrian Period of geologic time ( approximately 550 million years ago), an "inexplicable" absence that could be "truly urged as a valid argument" against his all embracing synthesis. For more than 100 years, the "missing Precambrian history of life" stood out as one of the greatest unsolved mysteries in natural science. But in recent decades, understanding of life's history has changed markedly as the documented fossil record has been extended seven-fold to some 3,500 million years ago, an age more than three-quarters that of the planet itself. This long-sought solution to Darwin's dilemma was set in motion by a small vanguard of workers who blazed the trail in the 1950s and 1960s, just as their course was charted by a few pioneering pathfinders of the previous century, a history of bold pronouncements, dashed dreams, search, and final discovery.

  6. Big data from electronic health records for early and late translational cardiovascular research: challenges and potential.

    PubMed

    Hemingway, Harry; Asselbergs, Folkert W; Danesh, John; Dobson, Richard; Maniadakis, Nikolaos; Maggioni, Aldo; van Thiel, Ghislaine J M; Cronin, Maureen; Brobert, Gunnar; Vardas, Panos; Anker, Stefan D; Grobbee, Diederick E; Denaxas, Spiros

    2018-04-21

    Cohorts of millions of people's health records, whole genome sequencing, imaging, sensor, societal and publicly available data present a rapidly expanding digital trace of health. We aimed to critically review, for the first time, the challenges and potential of big data across early and late stages of translational cardiovascular disease research. We sought exemplars based on literature reviews and expertise across the BigData@Heart Consortium. We identified formidable challenges including: data quality, knowing what data exist, the legal and ethical framework for their use, data sharing, building and maintaining public trust, developing standards for defining disease, developing tools for scalable, replicable science and equipping the clinical and scientific work force with new inter-disciplinary skills. Opportunities claimed for big health record data include: richer profiles of health and disease from birth to death and from the molecular to the societal scale; accelerated understanding of disease causation and progression, discovery of new mechanisms and treatment-relevant disease sub-phenotypes, understanding health and diseases in whole populations and whole health systems and returning actionable feedback loops to improve (and potentially disrupt) existing models of research and care, with greater efficiency. In early translational research we identified exemplars including: discovery of fundamental biological processes e.g. linking exome sequences to lifelong electronic health records (EHR) (e.g. human knockout experiments); drug development: genomic approaches to drug target validation; precision medicine: e.g. DNA integrated into hospital EHR for pre-emptive pharmacogenomics. In late translational research we identified exemplars including: learning health systems with outcome trials integrated into clinical care; citizen driven health with 24/7 multi-parameter patient monitoring to improve outcomes and population-based linkages of multiple EHR sources

  7. Fossils and the Evolution of the Arthropod Brain.

    PubMed

    Strausfeld, Nicholas J; Ma, Xiaoya; Edgecombe, Gregory D

    2016-10-24

    The discovery of fossilized brains and ventral nerve cords in lower and mid-Cambrian arthropods has led to crucial insights about the evolution of their central nervous system, the segmental identity of head appendages and the early evolution of eyes and their underlying visual systems. Fundamental ground patterns of lower Cambrian arthropod brains and nervous systems correspond to the ground patterns of brains and nervous systems belonging to three of four major extant panarthropod lineages. These findings demonstrate the evolutionary stability of early neural arrangements over an immense time span. Here, we put these fossil discoveries in the context of evidence from cladistics, as well as developmental and comparative neuroanatomy, which together suggest that despite many evolved modifications of neuropil centers within arthropod brains and ganglia, highly conserved arrangements have been retained. Recent phylogenies of the arthropods, based on fossil and molecular evidence, and estimates of divergence dates, suggest that neural ground patterns characterizing onychophorans, chelicerates and mandibulates are likely to have diverged between the terminal Ediacaran and earliest Cambrian, heralding the exuberant diversification of body forms that account for the Cambrian Explosion. Copyright © 2016 Elsevier Ltd. All rights reserved.

  8. Evidence for Late-Paleozoic brine migration in Cambrian carbonate rocks of the central and southern Appalachians: Implications for Mississippi Valley-type sulfide mineralization

    USGS Publications Warehouse

    Hearn, P.P.; Sutter, J.F.; Belkin, H.E.

    1987-01-01

    Many Lower Paleozoic limestones and dolostones in the Valley and Ridge province of the central and southern Appalachians contain 10 to 25 weight percent authigenic potassium feldspar. This was considered to be a product of early diagenesis, however, 40Ar 39Ar analyses of overgrowths on detrital K-feldspar in Cambrian carbonate rocks from Pennsylvania, Maryland, Virginia, and Tennessee yield Late Carboniferous-Early Permian ages (278-322 Ma). Simple mass balance calculations suggest that the feldspar could not have formed isochemically, but required the flux of multiple pore volumes of fluid through the rocks, reflecting regional fluid migration events during the Late-Paleozoic Alleghanian orogeny. Microthermometric measurements of fluid inclusions in overgrowths on detrital K-feldspar and quartz grains from unmineralized rocks throughout the study area indicate homogenization temperatures from 100?? to 200??C and freezing point depressions of -14?? to -18.5??C (18-21 wt.% NaCl equiv). The apparent similarity of these fluids to fluid inclusions in ore and gangue minerals of nearby Mississippi Valley-type (MVT) deposits suggests that the regional occurrences of authigenic K-feldspar and MVT mineralization may be genetically related. This hypothesis is supported by the discovery of authigenic K-feldspar intergrown with sphalerite in several mines of the Mascot-Jefferson City District, E. Tennessee. Regional potassic alteration in unmineralized carbonate rocks and localized occurrences of MVT mineralization are both explainable by a gravity-driven flow model, in which deep brines migrate towards the basin margin under a hydraulic gradient established during the Alleghanian orogeny. The authigenic K-feldspar may reflect the loss of K during disequilibrium cooling of the ascending brines. MVT deposits are probably localized manifestations of the same migrating fluids, occurring where the necessary physical and chemical traps are present. ?? 1987.

  9. Patterns of metal distribution in hypersaline microbialites during early diagenesis: Implications for the fossil record.

    PubMed

    Sforna, M C; Daye, M; Philippot, P; Somogyi, A; van Zuilen, M A; Medjoubi, K; Gérard, E; Jamme, F; Dupraz, C; Braissant, O; Glunk, C; Visscher, P T

    2017-03-01

    The use of metals as biosignatures in the fossil stromatolite record requires understanding of the processes controlling the initial metal(loid) incorporation and diagenetic preservation in living microbialites. Here, we report the distribution of metals and the organic fraction within the lithifying microbialite of the hypersaline Big Pond Lake (Bahamas). Using synchrotron-based X-ray microfluorescence, confocal, and biphoton microscopies at different scales (cm-μm) in combination with traditional geochemical analyses, we show that the initial cation sorption at the surface of an active microbialite is governed by passive binding to the organic matrix, resulting in a homogeneous metal distribution. During early diagenesis, the metabolic activity in deeper microbialite layers slows down and the distribution of the metals becomes progressively heterogeneous, resulting from remobilization and concentration as metal(loid)-enriched sulfides, which are aligned with the lamination of the microbialite. In addition, we were able to identify globules containing significant Mn, Cu, Zn, and As enrichments potentially produced through microbial activity. The similarity of the metal(loid) distributions observed in the Big Pond microbialite to those observed in the Archean stromatolites of Tumbiana provides the foundation for a conceptual model of the evolution of the metal distribution through initial growth, early diagenesis, and fossilization of a microbialite, with a potential application to the fossil record. © 2016 John Wiley & Sons Ltd.

  10. Early Paleozoic magmatic events in the eastern Klamath Mountains, northern California

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Wallin, E.T.; Mattinson, J.M.; Potter, A.W.

    1988-02-01

    New U-Pb zircon ages for nine samples of tonalite and pegmatitic trondhjemite from the Trinity ophiolite and associated melange reveal a complex history of magmatic activity extending back into the earliest Cambrian, much older than previously believed. Earlier investigations, based on limited data, recognized lower Paleozoic crustal elements in the eastern Klamath terrane (EKT) ranging in age from Middle Ordovician to Early to Middle Devonian. The new work in the Yreka-Callahan area of the EKT confirms the Ordovician (440-475 Ma) and younger ages, but reveals for the first time the presence of tonalitic rocks that crystallized during a narrow timemore » interval at about 565-570 Ma. The authors also recognize younger, Late Silurian magmatism at 412 Ma. In the context of available mapping, these ages indicate that the Trinity ophiolite is broadly polygenetic because parts of it yield crystallization ages that span approximately 150 m.y. Superjacent dismembered units of probable early Paleozoic age may be tectonostratigraphically equivalent to the Sierra City melange in the northern Sierra Nevada.« less

  11. Geology of the Sierra de Fiambala, northwestern Argentina: implications for Early Palaeozoic Andean tectonics

    USGS Publications Warehouse

    Grissom, G.C.; DeBari, S.M.; Snee, L.W.

    1998-01-01

    This paper is included in the Special Publication entitled 'The proto- Andean margin of Gondwana', edited by R.J. Pankhurst and C.W. Rapela. Field mapping in conjunction with structural, metamorphic, and geochronological data document the tectono-thermal history of exhumed deep crustal rocks in the Sierra de Fiambala, NW Argentina. The range consists of two structural blocks distinguished by different metasedimentary sequences and different grades of metamorphism. Orthogneiss and paragneiss in the northern structural block may have a Precambrian history. Greenschist- to amphibolite-facies metamorphism, intrusion, and injection magmatization affected all rocks at 540-550 Ma. A subsequent event in the Late Cambrian to Ordovician (c.515 to 470 Ma) involved amphibolite- to granulite-facies metamorphism, mafic intrusion, and deformation, followed by cooling through mid-Palaeozoic time. The emplacement of Carboniferous (325-350 Ma) post-tectonic granites caused reheating and retrogression that was strongest toward the northeast part of the range. The Cambrian, Ordovician, and Carboniferous events in the Sierra de Fiambala were of regional extent as indicated by temporal correlations with events reported for other deep crustal rocks of the northern Sierras Pampeanas. Correlations between periods of intrusion and high-grade metamorphism in the northern Sierras Pampeanas and volcanic-sedimentary events in the adjacent supracrustal exposures confirm that rocks in the northern Sierras Pampeanas formed at deep (10-25 km) structural levels in the early Palaeozoic continental margin of Gondwana.

  12. The geology of the northern tip of the Arabian-Nubian Shield

    NASA Astrophysics Data System (ADS)

    Beyth, M.; Eyal, Y.; Garfunkel, Z.

    2014-11-01

    accompanied in ∼609 Ma by rhyolite, andesite and composite dykes. The last magmatic event in the Elat area is represented by the volcano-conglomeratic series comprising rhyolites, basalts, andesites, hypabyssal intrusions of monzonite and syenite and conglomerates. The conglomerates, dated to about 590 Ma, are the products of a major erosion phase in which about 12,000 m of the section were removed. These conglomerates were intruded by 585 Ma rhyolite, andesite and composite dykes. The Neoproterozoic basement is truncated by a peneplain whose age, post 532 Ma, is constrained by the age of the youngest eroded dolerite dykes. This Early Cambrian peneplain was associated with erosion of 2000 m of the section and by chemical weathering. Three major breaks in Neoproterozoic magmatic activity are recognized: the first, occurred in Cryogenian time, lasted ∼60 million years after the amphibolite facies metamorphism and before emplacement of the calc alkaline plutons, separating the first and the second orogenic phases; the second break between the orogenic and the extensional phases occurred in early Ediacaran time, encompassed ∼20 million years between the emplacement of the calc-alkaline and alkaline plutonic rocks and rhyolite, andesite and the composite dykes; and the third, ∼50 Ma break, occurred between the emplacement of the last felsic intrusions at ∼585 Ma and intrusion of the dolerite dykes in 532 Ma, before the Early Cambrian peneplain developed. The great lateral extension of the Cambrian to Eocene sedimentary rocks and their slow facies and thickness changes suggest a stable flat platform area at the northern tip of the ANS. Early Cambrian sedimentation began with fluviatile subarkoses of the Amudei Shlomo Formation. It was overlain by an Early to Middle Cambrian transgressive-regressive lagoonal cycle of dolostones, sandstones, and siltstones of the Timna Formation. Then Middle Cambrian subarkoses and siltstones of the Shehoret Formation and the quartz arenite

  13. Clades reach highest morphological disparity early in their evolution

    PubMed Central

    Hughes, Martin; Gerber, Sylvain; Wills, Matthew Albion

    2013-01-01

    There are few putative macroevolutionary trends or rules that withstand scrutiny. Here, we test and verify the purported tendency for animal clades to reach their maximum morphological variety relatively early in their evolutionary histories (early high disparity). We present a meta-analysis of 98 metazoan clades radiating throughout the Phanerozoic. The disparity profiles of groups through time are summarized in terms of their center of gravity (CG), with values above and below 0.50 indicating top- and bottom-heaviness, respectively. Clades that terminate at one of the “big five” mass extinction events tend to have truncated trajectories, with a significantly top-heavy CG distribution overall. The remaining 63 clades show the opposite tendency, with a significantly bottom-heavy mean CG (relatively early high disparity). Resampling tests are used to identify groups with a CG significantly above or below 0.50; clades not terminating at a mass extinction are three times more likely to be significantly bottom-heavy than top-heavy. Overall, there is no clear temporal trend in disparity profile shapes from the Cambrian to the Recent, and early high disparity is the predominant pattern throughout the Phanerozoic. Our results do not allow us to distinguish between ecological and developmental explanations for this phenomenon. To the extent that ecology has a role, however, the paucity of bottom-heavy clades radiating in the immediate wake of mass extinctions suggests that early high disparity more probably results from the evolution of key apomorphies at the base of clades rather than from physical drivers or catastrophic ecospace clearing. PMID:23884651

  14. The Completeness of the Fossil Record of Mesozoic Birds: Implications for Early Avian Evolution

    PubMed Central

    Brocklehurst, Neil; Upchurch, Paul; Mannion, Philip D.; O'Connor, Jingmai

    2012-01-01

    Many palaeobiological analyses have concluded that modern birds (Neornithes) radiated no earlier than the Maastrichtian, whereas molecular clock studies have argued for a much earlier origination. Here, we assess the quality of the fossil record of Mesozoic avian species, using a recently proposed character completeness metric which calculates the percentage of phylogenetic characters that can be scored for each taxon. Estimates of fossil record quality are plotted against geological time and compared to estimates of species level diversity, sea level, and depositional environment. Geographical controls on the avian fossil record are investigated by comparing the completeness scores of species in different continental regions and latitudinal bins. Avian fossil record quality varies greatly with peaks during the Tithonian-early Berriasian, Aptian, and Coniacian–Santonian, and troughs during the Albian-Turonian and the Maastrichtian. The completeness metric correlates more strongly with a ‘sampling corrected’ residual diversity curve of avian species than with the raw taxic diversity curve, suggesting that the abundance and diversity of birds might influence the probability of high quality specimens being preserved. There is no correlation between avian completeness and sea level, the number of fluviolacustrine localities or a recently constructed character completeness metric of sauropodomorph dinosaurs. Comparisons between the completeness of Mesozoic birds and sauropodomorphs suggest that small delicate vertebrate skeletons are more easily destroyed by taphonomic processes, but more easily preserved whole. Lagerstätten deposits might therefore have a stronger impact on reconstructions of diversity of smaller organisms relative to more robust forms. The relatively poor quality of the avian fossil record in the Late Cretaceous combined with very patchy regional sampling means that it is possible neornithine lineages were present throughout this interval but

  15. The Early Toarcian Oceanic Anoxic Event: A Southern Hemisphere record from Chile

    NASA Astrophysics Data System (ADS)

    Fantasia, Alicia; Föllmi, Karl B.; Adatte, Thierry; Spangenberg, Jorge E.; Bernárdez, Enrique; Mattioli, Emanuela

    2016-04-01

    The Early Toarcian was marked by important environmental changes, marine oxygen deficiency and extensive organic-rich sediment deposition (T-OAE; ˜182 Ma, Early Jurassic). The T-OAE coincides with a marked negative carbon isotope excursion (CIE) recorded in marine carbonate, and marine and terrestrial organic carbon. This is commonly attributed to the massive release of isotopically light carbon to the atmospheric and oceanic reservoirs derived from the destabilization of methane hydrates from marine sediments and/or the emissions of thermogenic methane from the eruption of the Karoo-Ferrar LIP (e.g., Hesselbo et al., 2000; Kemp et al., 2005; Svensen et al., 2007; Mazzini et al., 2010). Moreover, in most documented marine sections, this episode is marked by a generalized crisis in carbonate production and marine invertebrate extinctions (e.g. Jenkyns, 1988; Röhl et al., 2005; Suan et al., 2001). Several studies of the T-OAE have been conducted on sediments in central and northwest Europe, but only few data are available from the Southern Hemisphere, leading to large uncertainty concerning the exact expression of this event in this part of the world. The aims of this study are to characterize the sediments deposited during the Andean equivalents of the tenuicostatum and falciferum European Zones and establish in which way the T-OAE affected this region. In the Early Jurassic, the Andean basin was in a back-arc setting with marine corridors connected to Panthalassa. In this study, we have generated new high-resolution sedimentological, geochemical and mineralogical data from the sections of El Peñon and Quebrada Asiento, located in Chile in the northeastern area of the city of Copiapó, Atacama region. The biostratigraphy of these sections has been studied by von Hillebrandt and Schidt-Effing (1981) and complemented here by a biostratigraphy based on calcareous nannofossils. The sections consist of a succession of marl, limestone and siltstone of Pliensbachian and

  16. Text mining approach to predict hospital admissions using early medical records from the emergency department.

    PubMed

    Lucini, Filipe R; S Fogliatto, Flavio; C da Silveira, Giovani J; L Neyeloff, Jeruza; Anzanello, Michel J; de S Kuchenbecker, Ricardo; D Schaan, Beatriz

    2017-04-01

    Emergency department (ED) overcrowding is a serious issue for hospitals. Early information on short-term inward bed demand from patients receiving care at the ED may reduce the overcrowding problem, and optimize the use of hospital resources. In this study, we use text mining methods to process data from early ED patient records using the SOAP framework, and predict future hospitalizations and discharges. We try different approaches for pre-processing of text records and to predict hospitalization. Sets-of-words are obtained via binary representation, term frequency, and term frequency-inverse document frequency. Unigrams, bigrams and trigrams are tested for feature formation. Feature selection is based on χ 2 and F-score metrics. In the prediction module, eight text mining methods are tested: Decision Tree, Random Forest, Extremely Randomized Tree, AdaBoost, Logistic Regression, Multinomial Naïve Bayes, Support Vector Machine (Kernel linear) and Nu-Support Vector Machine (Kernel linear). Prediction performance is evaluated by F1-scores. Precision and Recall values are also informed for all text mining methods tested. Nu-Support Vector Machine was the text mining method with the best overall performance. Its average F1-score in predicting hospitalization was 77.70%, with a standard deviation (SD) of 0.66%. The method could be used to manage daily routines in EDs such as capacity planning and resource allocation. Text mining could provide valuable information and facilitate decision-making by inward bed management teams. Copyright © 2017 Elsevier Ireland Ltd. All rights reserved.

  17. DHI evaluation by combining rock physics simulation and statistical techniques for fluid identification of Cambrian-to-Cretaceous clastic reservoirs in Pakistan

    NASA Astrophysics Data System (ADS)

    Ahmed, Nisar; Khalid, Perveiz; Shafi, Hafiz Muhammad Bilal; Connolly, Patrick

    2017-10-01

    The use of seismic direct hydrocarbon indicators is very common in exploration and reservoir development to minimise exploration risk and to optimise the location of production wells. DHIs can be enhanced using AVO methods to calculate seismic attributes that approximate relative elastic properties. In this study, we analyse the sensitivity to pore fluid changes of a range of elastic properties by combining rock physics studies and statistical techniques and determine which provide the best basis for DHIs. Gassmann fluid substitution is applied to the well log data and various elastic properties are evaluated by measuring the degree of separation that they achieve between gas sands and wet sands. The method has been applied successfully to well log data from proven reservoirs in three different siliciclastic environments of Cambrian, Jurassic, and Cretaceous ages. We have quantified the sensitivity of various elastic properties such as acoustic and extended elastic (EEI) impedances, elastic moduli ( K sat and K sat- μ), lambda-mu-rho method ( λρ and μρ), P-to-S-wave velocity ratio ( V P/ V S), and Poisson's ratio ( σ) at fully gas/water saturation scenarios. The results are strongly dependent on the local geological settings and our modeling demonstrates that for Cambrian and Cretaceous reservoirs, K sat- μ, EEI, V P/ V S, and σ are more sensitive to pore fluids (gas/water). For the Jurassic reservoir, the sensitivity of all elastic and seismic properties to pore fluid reduces due to high overburden pressure and the resultant low porosity. Fluid indicators are evaluated using two metrics: a fluid indicator coefficient based on a Gaussian model and an overlap coefficient which makes no assumptions about a distribution model. This study will provide a potential way to identify gas sand zones in future exploration.

  18. Crustal Structure, Seismic Anisotropy and Deformations of the Ediacaran/Cambrian of the Małopolska Block in SE Poland Based on Data from Two Seismic Wide-Angle Experiments

    NASA Astrophysics Data System (ADS)

    Środa, Piotr

    2017-04-01

    The area of SE Poland represents a complex contact of tectonic units of different consolidation age—from the Precambrian East European Craton, through Palaeozoic West European Platform (including Małopolska Block) to Cenozoic Carpathians and Carpathian Foredeep. In order to investigate the anisotropic properties of the upper crust of the Małopolska Block and their relation to tectonic evolution of the area, two seismic datasets were used: seismic wide-angle off-line recordings from POLCRUST-01 deep seismic reflection profile and recordings from active deep seismic experiment CELEBRATION 2000. During acquisition of deep reflection seismic profile POLCRUST-01 in 2010, a 35-km-long line of 14 recorders (PA-14), oriented perpendicularly to the profile, was deployed to record the refractions from the upper crust (Pg) at wide range of azimuths. These data were used for an analysis of the azimuthal anisotropy of the MB with the modified delay-time inversion method. The results of modelling of the off-line refractions from the MB suggest 6% HTI anisotropy of the Cambrian/Ediacaran basement, with 130º azimuth of the fast velocity axis and mean Vp of 4.9 km/s. To compare this result with previous, independent information about anisotropy at larger depth, a subset of previously modelled data from CELEBRATION 2000 experiment, recorded in the MB area, was also analysed by inversion. The recordings of Pg phase at up to 120 km offsets were analysed using anisotropic delay-time inversion, providing information down to 12 km depth. The CELEBRATION 2000 model shows 9% HTI anisotropy with 126º orientation of the fast axis. Thus, local-scale anisotropy of this part of MB confirms the large-scale anisotropy suggested by previous studies based on data from a broader area and larger depth interval. The azimuthal anisotropy (i.e. HTI symmetry of the medium) is interpreted as a result of strong compressional deformation during the accretion of terranes to the EEC margin, leading to

  19. Early Proterozoic (2.04 GA) Phoshorites of Pechenga Greenstone Belt and Their Origin

    NASA Technical Reports Server (NTRS)

    Rozanov, Alexei Yu.; Astafieva, Marina M.; Hoover, Richard B.

    2007-01-01

    No principal differences have been found between microfossils described from Cambrian and Phanerozoic and the 2000 Ma phosphorites. Numerous samples revealed diverse microbial microstructures interpreted as cyanobacterial mats consisting of filamentous (1-3 microns in diameter, 20 microns in length), coccoidal (0.8-1.0 microns) and ellipsoidal or rod-shaped microfossils (0.8 microns in diameter, around 2 microns in length) which morphologically resemble modern Microcoleus and Siphonophycus, Thiocapsa, and Rhabdoderma, respectively, reported from alkali ne or saline environment_ The sequence of the early Palaeoproterozoic events which point to a significant oxidation of the hydrosphere, including the formation of phosphorites and changes in the phosphorous cycle, mimics the sequence which was repeated at the Neoproterozoic-Cembrian transition, implying that oxidation of the terrestrial atmosphere-hydrosphere system experienced an irregular cyclic development.

  20. Running Records: Authentic Instruction in Early Childhood Education

    ERIC Educational Resources Information Center

    Shea, Mary

    2012-01-01

    The most effective way to understand what a child knows about the reading process is to take a running record. In "Running Records", Mary Shea demonstrates how teachers can use this powerful tool to design lessons that decrease reading difficulties, build on strengths, and stimulate motivation, ensuring that children develop self-sustaining…

  1. The termites of Early Eocene Cambay amber, with the earliest record of the Termitidae (Isoptera)

    PubMed Central

    Engel, Michael S.; Grimaldi, David A.; Nascimbene, Paul C.; Singh, Hukam

    2011-01-01

    Abstract The fauna of termites (Isoptera) preserved in Early Eocene amber from the Cambay Basin (Gujarat, India) are described and figured. Three new genera and four new species are recognized, all of them Neoisoptera – Parastylotermes krishnai Engel & Grimaldi, sp. n. (Stylotermitidae); Prostylotermes kamboja Engel & Grimaldi, gen. et sp. n. (Stylotermitidae?); Zophotermes Engel, gen. n., with Zophotermes ashoki Engel & Singh, sp. n. (Rhinotermitidae: Prorhinotermitinae); and Nanotermes isaacae Engel & Grimaldi, gen. et sp. n. (Termitidae: Termitinae?). Together these species represent the earliest Tertiary records of the Neoisoptera and the oldest definitive record of Termitidae, a family that comprises >75% of the living species of Isoptera. Interestingly, the affinities of the Cambay amber termites are with largely Laurasian lineages, in this regard paralleling relationships seen between the fauna of bees and some flies. Diversity of Neoisoptera in Indian amber may reflect origin of the amber deposit in Dipterocarpaceae forests formed at or near the paleoequator. PMID:22287892

  2. Pikaia gracilens Walcott: stem chordate, or already specialized in the Cambrian?

    PubMed

    Mallatt, Jon; Holland, Nicholas

    2013-06-01

    For the past 35 years, the Cambrian fossil Pikaia gracilens was widely interpreted as a typical basal chordate based on short descriptions by Conway Morris. Recently, Conway Morris and Caron (CMC) (2012, Biol Rev 87:480-512) described Pikaia extensively, as a basis for new ideas about deuterostome evolution. This new Pikaia has characters with no clear homologues in other animals, so they could be phylogenetically uninformative autapomorphies. These characters include a dorsal organ, posterior ventral area, posterior fusiform structure, and anterior dorsal unit. Yet CMC interpret most of the unusual characters as primitive for chordates, thereby interpreting Pikaia as an even more convincing stem chordate than before. Moreover, they claim that segment (myomere) shape is a reliable guide for defining a chordate and even for assigning animals to their correct place in deuterostome phylogeny. By defining sigmoidal segments as a basal chordate character, they situate Pikaia at the base of the chordates and banish fossil yunnanozoans (which have straight segments) to a position deep within the deuterostomes. In addition, they consider amphioxus, with its conspicuously chevron-shaped segments, to be so highly derived that it is of little use for reconstructing the first chordates. We question their overemphasis on the phylogenetic value of segment shape and their marginalizing of amphioxus. We deduce that Pikaia, not amphioxus, is specialized. We performed a cladistic analysis that showed the character interpretations of CMC are consistent with their wide-ranging evolutionary scenario, but that these interpretations leave unresolved the position of Pikaia within chordates. Copyright © 2013 Wiley Periodicals, Inc.

  3. The Middle Cambrian fossil Pikaia and the evolution of chordate swimming.

    PubMed

    Lacalli, Thurston

    2012-07-06

    Conway Morris and Caron (2012) have recently published an account of virtually all the available information on Pikaia gracilens, a well-known Cambrian fossil and supposed basal chordate, and propose on this basis some new ideas about Pikaia's anatomy and evolutionary significance. Chief among its chordate-like features are the putative myomeres, a regular series of vertical bands that extends the length of the body. These differ from the myomeres of living chordates in that boundaries between them (the myosepta) are gently curved, with minimal overlap, whereas amphioxus and vertebrates have strongly overlapping V- and W-shaped myomeres. The implication, on biomechanical grounds, is that myomeres in Pikaia exerted much less tension on the myosepta, so the animal would have been incapable of swimming as rapidly as living chordates operating in the fast-twitch mode used for escape and attack. Pikaia either lacked the fast-twitch fibers necessary for such speeds, having instead only slow-twitch fibers, or it had an ancestral fiber type with functional capabilities more like modern slow fibers than fast ones. The first option is supported by the sequence of development in zebrafish, where both myoseptum formation and fast fiber deployment show a dependence on slow fibers, which develop first. For Pikaia, the absence of fast fibers has both behavioral and anatomical implications, which are discussed. Among the latter is the possibility that a notochord may not have been needed as a primary stiffening device if other structures (for example, the dorsal organ) could perform that role.

  4. Deep mantle roots and continental hypsometry: implications for whole-Earth elemental cycling, long-term climate, and the Cambrian explosion

    NASA Astrophysics Data System (ADS)

    Lee, C. T.

    2016-12-01

    Most of Earth's continents today are above sea level, but the presence of thick packages of ancient sediments on top of the stable cores of continents indicates that continents must have been submerged at least once in their past. Elevations of continents are controlled by the interplay between crustal thickness, mantle root thickness and the temperature of the ambient convecting mantle. The history of a continent begins with mountain building through magmatic or tectonic crustal thickening, during which exhumation of deep-seated igneous and metamorphic rocks are highest. Mountain building is followed by a long interval of subsidence as a result of continued, but decreasing erosion and thermal relaxation, the latter in the form of a growing thermal boundary layer. Subsidence is manifest first as a boring interval in which no sedimentary record is preserved, followed by continent-scale submergence wherein sediments are deposited directly on deep-seated igneous/metamorphic basement, generating a major disconformity. The terminal resting elevation of a mature continent, however, is defined by the temperature of the ambient convecting mantle: below sea level when the mantle is hot and above sea level when the mantle is cold. Using thermobarometric constraints on secular cooling of Earth's mantle, our results suggest that Earth, for most of its history, must have been a water world, with regions of land confined to narrow orogenic belts and oceans characterized by deep basins and shallow continental seas, the latter serving as repositories of sediments and key redox-sensitive biological nutrients, such as phosphorous. Cooling of the Earth led to the gradual and irreversible rise of the continents, culminating in rapid emergence, through fits and starts and possible instabilities in climate, between 500-1000 Ma. Such emergence fundamentally altered marine biogeochemical cycling, continental weathering and the global hydrologic cycle, defining the backdrop for the

  5. Paired δ34S data from carbonate-associated sulfate and chromium-reducible sulfur across the traditional Lower-Middle Cambrian boundary of W-Gondwana

    NASA Astrophysics Data System (ADS)

    Wotte, Thomas; Strauss, Harald; Fugmann, Artur; Garbe-Schönberg, Dieter

    2012-05-01

    In this study, we present the first high-resolution data from coupled δ34S analyses of carbonate-associated sulfate (CAS) and chromium-reducible sulfur (CRS) from three Lower-Middle Cambrian sections in western Gondwana. CAS and CRS were extracted and analyzed from marine dolostone, limestone, and nodular limestone from Spanish and French successions. In parallel, carbonate samples were also analyzed for δ13Ccarb, δ18Ocarb, and major/trace element concentrations (Ca, Fe, Mg, Mn, Sr). δ34SCAS values vary between 17.6‰ and 33.2‰, with a maximum CAS concentration of ca. 900 ppm. δ34SCRS data show a similar broad range between -5.1‰ and 29.7‰, with maximal CRS contents up to ca. 3700 ppm. Notably, there is little stratigraphic variation in the δ34SCAS data in each of the sections confounding inter-basinal chemostratigraphic correlations. Nonetheless, the absolute differences in δ34SCAS between sections as well as variations in CAS and CRS concentrations are attributed to paleoenvironmental differences between proximal and distal parts of the carbonate ramp, as well as effects of subaerial exposure and riverine input. Thus, the generated δ34SCAS data deliver not only valuable paleoecological and paleoenvironmental information, they also illustrate a heterogeneity in the seawater sulfate sulfur isotopic composition of the western Gondwanan ocean. Consequently, the lack of correlation between our Gondwanan δ34SCAS data and time equivalent sections of Laurentia and Siberia is probably not only caused by the absence of an internationally accepted biostratigraphic correlation, but rather supports the view that sulfate was non-conservative anion in seawater during the Cambrian Period.

  6. Electronic health records and patient safety: co-occurrence of early EHR implementation with patient safety practices in primary care settings.

    PubMed

    Tanner, C; Gans, D; White, J; Nath, R; Pohl, J

    2015-01-01

    The role of electronic health records (EHR) in enhancing patient safety, while substantiated in many studies, is still debated. This paper examines early EHR adopters in primary care to understand the extent to which EHR implementation is associated with the workflows, policies and practices that promote patient safety, as compared to practices with paper records. Early adoption is defined as those who were using EHR prior to implementation of the Meaningful Use program. We utilized the Physician Practice Patient Safety Assessment (PPPSA) to compare primary care practices with fully implemented EHR to those utilizing paper records. The PPPSA measures the extent of adoption of patient safety practices in the domains: medication management, handoffs and transition, personnel qualifications and competencies, practice management and culture, and patient communication. Data from 209 primary care practices responding between 2006-2010 were included in the analysis: 117 practices used paper medical records and 92 used an EHR. Results showed that, within all domains, EHR settings showed significantly higher rates of having workflows, policies and practices that promote patient safety than paper record settings. While these results were expected in the area of medication management, EHR use was also associated with adoption of patient safety practices in areas in which the researchers had no a priori expectations of association. Sociotechnical models of EHR use point to complex interactions between technology and other aspects of the environment related to human resources, workflow, policy, culture, among others. This study identifies that among primary care practices in the national PPPSA database, having an EHR was strongly empirically associated with the workflow, policy, communication and cultural practices recommended for safe patient care in ambulatory settings.

  7. Early Paleozoic paleogeography of the northern Gondwana margin: new evidence for Ordovician-Silurian glaciation

    NASA Astrophysics Data System (ADS)

    Semtner, A.-K.; Klitzsch, E.

    1994-12-01

    During the Early Paleozoic, transgressions and the distribution of sedimentary facies on the northern Gondwana margin were controlled by a regional NNW-SSE to almost north-south striking structural relief. In Early Silurian times, a eustatic highstand enabled the sea to reach its maximum southward extent. The counterclockwise rotation of Gondwana during the Cambrian and Early Ordovician caused the northern Gondwana margin to shift from intertropical to southern polar latitudes in Ordovician times. Glacial and periglacial deposits are reported from many localities in Morocco, Algeria, Niger, Libya, Chad, Sudan, Jordan and Saudi Arabia. The Late Ordovician glaciation phase was followed by a period of a major glacioeustatic sea-level rise in the Early Silurian due to the retreat of the ice-cap. As a consequence of the decreasing water circulation in the basin centers (Central Arabia, Murzuk- and Ghadames basins), highly bituminous euxinic shales were deposited. These shales are considered to be the main source rock of Paleozoic oil and gas deposits in parts of Saudi Arabia, Libya and Algeria. The following regression in the southern parts of the Early Silurian sea was probably caused by a second glacial advance, which was mainly restricted to areas in Chad, Sudan and Niger. Evidence for glacial activity and fluvioglacial sedimentation is available from rocks overlying the basal Silurian shale in north-east Chad and north-west Sudan. The Early Silurian ice advance is considered to be responsible for the termination of euxinic shale deposition in the basin centers.

  8. Origin and implications of a thrust-bound gypsiferous unit along the western edge of Jabal Sumeini, northern Oman Mountains

    NASA Astrophysics Data System (ADS)

    Cooper, David J. W.; Ali, Mohammed Y.; Searle, Michael P.

    2018-04-01

    The Oman Mountains comprise a series of thrust sheets of Neo-Tethyan oceanic rocks that were emplaced onto the Arabian continental margin during obduction of the Semail Ophiolite during the Late Cretaceous. Three separate groups of anomalous gypsiferous bodies intrude the allochthonous units along faults over a distance of about 150 km in the Hawasina Window, Jabal Qumayrah and Jabal Sumeini. The bodies at Jabal Sumeini form a band about 4 km long and up to 100 m wide along a late-stage thrust that restacks the allochthon over a post-emplacement Maastrichtian-Palaeogene sedimentary succession. The gypsum shows evidence of flow-folding and contains numerous clasts and rafts of a range of quartzose sandstones, but with only a minor component from carbonates from the Neo-Tethyan Sumeini Group in the hanging-wall. Palaeogene limestones from the footwall succession are essentially absent. Strontium isotope ratios are high and intersect with the open ocean-water reference curve for the Late Cambrian-Ordovician and Late Miocene-Pliocene. They are also noticeably higher than the ratios from the two other gypsiferous outcrop areas in the Oman Mountains and from outcrops of Ediacaran-Early Cambrian salt domes in central Oman. However, the regional stratigraphy points towards a source of the gypsum from either an Ediacaran-Early Cambrian Ara Group salt basin or from the Lower Fars Formation (Early-Middle Miocene), and derivation of the sandstone clasts and rafts from thick Lower Palaeozoic clastic sequences. The discrepancy with the ages inferred from the strontium isotope data can be attributed to deposition of the gypsum in restricted conditions not in equilibrium with the prevailing ocean water. Two models are presented, for an Ediacaran-Early Cambrian and an Early-Middle Miocene source. While the latter cannot be wholly discounted, the stratigraphic and structural context point more strongly towards an Ediacaran-Early Cambrian Ara Group source of the gypsum. This was

  9. Characterizing Early Adolescent Plate Waste Using the Mobile Food Record.

    PubMed

    Panizza, Chloe E; Boushey, Carol J; Delp, Edward J; Kerr, Deborah A; Lim, Eunjung; Gandhi, Krupa; Banna, Jinan C

    2017-01-26

    This study aimed to assess the amount of plate waste and how plate waste was disposed by early adolescent girls using a mobile food record (mFR). Participants were girls nine to thirteen years residing in O'ahu, Hawai'i ( n = 93). Foods selected and leftover were estimated using a three day mFR. Each leftover food was then classified as thrown into the trash, fed to a pet, eaten later, or other (e.g., composted). Repeated measures analyses of variance (ANOVA) were conducted and Tukey's post-hoc test were used to adjust for multiple comparisons between times (breakfast, lunch, dinner, and snack) on leftover food and leftover food thrown into the trash. The percentage of food leftover and thrown into the trash was highest at lunch. The percentage of protein, grain, vegetables, fruit, and dairy leftover at lunch were unexpectedly low compared to previous studies. The median for percentage of food thrown into the trash at lunch was <5% for all food groups, and was consistently low across the day (<10%). Average energy intake was 436 kcal (±216) at lunch, and 80% of caregivers reported total household income as ≥$70,000. Studies in real-time using technology over full days may better quantify plate waste among adolescents.

  10. Characterizing Early Adolescent Plate Waste Using the Mobile Food Record

    PubMed Central

    Panizza, Chloe E.; Boushey, Carol J.; Delp, Edward J.; Kerr, Deborah A.; Lim, Eunjung; Gandhi, Krupa; Banna, Jinan C.

    2017-01-01

    This study aimed to assess the amount of plate waste and how plate waste was disposed by early adolescent girls using a mobile food record (mFR). Participants were girls nine to thirteen years residing in O’ahu, Hawai’i (n = 93). Foods selected and leftover were estimated using a three day mFR. Each leftover food was then classified as thrown into the trash, fed to a pet, eaten later, or other (e.g., composted). Repeated measures analyses of variance (ANOVA) were conducted and Tukey’s post-hoc test were used to adjust for multiple comparisons between times (breakfast, lunch, dinner, and snack) on leftover food and leftover food thrown into the trash. The percentage of food leftover and thrown into the trash was highest at lunch. The percentage of protein, grain, vegetables, fruit, and dairy leftover at lunch were unexpectedly low compared to previous studies. The median for percentage of food thrown into the trash at lunch was <5% for all food groups, and was consistently low across the day (<10%). Average energy intake was 436 kcal (±216) at lunch, and 80% of caregivers reported total household income as ≥$70,000. Studies in real-time using technology over full days may better quantify plate waste among adolescents. PMID:28134757

  11. Calcification and Silicification: Fossilization Potential of Cyanobacteria from Stromatolites of Niuafo‘ou's Caldera Lakes (Tonga) and Implications for the Early Fossil Record

    PubMed Central

    Kazmierczak, Józef; Łukomska-Kowalczyk, Maja; Kempe, Stephan

    2012-01-01

    Abstract Calcification and silicification processes of cyanobacterial mats that form stromatolites in two caldera lakes of Niuafo‘ou Island (Vai Lahi and Vai Si‘i) were evaluated, and their importance as analogues for interpreting the early fossil record are discussed. It has been shown that the potential for morphological preservation of Niuafo‘ou cyanobacteria is highly dependent on the timing and type of mineral phase involved in the fossilization process. Four main modes of mineralization of cyanobacteria organic parts have been recognized: (i) primary early postmortem calcification by aragonite nanograins that transform quickly into larger needle-like crystals and almost totally destroy the cellular structures, (ii) primary early postmortem silicification of almost intact cyanobacterial cells that leave a record of spectacularly well-preserved cellular structures, (iii) replacement by silica of primary aragonite that has already recrystallized and obliterated the cellular structures, (iv) occasional replacement of primary aragonite precipitated in the mucopolysaccharide sheaths and extracellular polymeric substances by Al-Mg-Fe silicates. These observations suggest that the extremely scarce earliest fossil record may, in part, be the result of (a) secondary replacement by silica of primary carbonate minerals (aragonite, calcite, siderite), which, due to recrystallization, had already annihilated the cellular morphology of the mineralized microbiota or (b) relatively late primary silicification of already highly degraded and no longer morphologically identifiable microbial remains. Key Words: Stromatolites—Cyanobacteria—Calcification—Silicification—Niuafo‘ou (Tonga)—Archean. Astrobiology 12, 535–548. PMID:22794297

  12. Early Middle Ordovician evidence for land plants in Argentina (eastern Gondwana).

    PubMed

    Rubinstein, C V; Gerrienne, P; de la Puente, G S; Astini, R A; Steemans, P

    2010-10-01

    • The advent of embryophytes (land plants) is among the most important evolutionary breakthroughs in Earth history. It irreversibly changed climates and biogeochemical processes on a global scale; it allowed all eukaryotic terrestrial life to evolve and to invade nearly all continental environments. Before this work, the earliest unequivocal embryophyte traces were late Darriwilian (late Middle Ordovician; c. 463-461 million yr ago (Ma)) cryptospores from Saudi Arabia and from the Czech Republic (western Gondwana). • Here, we processed Dapingian (early Middle Ordovician, c. 473-471 Ma) palynological samples from Argentina (eastern Gondwana). • We discovered a diverse cryptospore assemblage, including naked and envelope-enclosed monads and tetrads, representing five genera. • Our discovery reinforces the earlier suggestion that embryophytes first evolved in Gondwana. It indicates that the terrestrialization of plants might have begun in the eastern part of Gondwana. The diversity of the Dapingian assemblage implies an earlier, Early Ordovician or even Cambrian, origin of embryophytes. Dapingian to Aeronian (Early Silurian) cryptospore assemblages are similar, suggesting that the rate of embryophyte evolution was extremely slow during the first c. 35-45 million yr of their diversification. The Argentinean cryptospores predate other cryptospore occurrences by c. 8-12 million yr, and are currently the earliest evidence of plants on land. © The Authors (2010). Journal compilation © New Phytologist Trust (2010).

  13. Modern Microbial Ecosystems are a Key to Understanding Our Biosphere's Early Evolution and its Contributions To The Atmosphere and Rock Record

    NASA Technical Reports Server (NTRS)

    DesMarais, David J.; DeVincenzi, Donald L. (Technical Monitor)

    2000-01-01

    The survival of our early biosphere depended upon efficient coordination anion- diverse microbial populations. Microbial mats exhibit a 3.46-billion-year fossil record, thus they are the oldest known ecosystems. Photosynthetic microbial mats were key because, today, sunlight powers more than 99 percent of global primary productivity. Thus photosynthetic ecosystems have affected the atmosphere profoundly and have created the most pervasive, easily-detected fossils. Photosynthetic biospheres elsewhere will be most detectible via telescopes or spacecraft. As a part of the Astrobiology Institute, our Ames Microbial Ecosystems group examines the roles played by ecological processes in the early evolution of our biosphere, as recorded in geologic fossils and in the macromolecules of living cells: (1) We are defining the microbial mat microenvironment, which was an important milieu for early evolution. (2) We are comparing mats in contrasting environments to discern strategies of adaptation and diversification, traits that were key for long-term survival. (3) We have selected sites that mimic key environmental attributes of early Earth and thereby focus upon evolutionary adaptations to long-term changes in the global environment. (4) Our studies of gas exchange contribute to better estimates of biogenic gases in Earth's early atmosphere. This group therefore directly addresses the question: How have the Earth and its biosphere influenced each other over time Our studies strengthen the systematics for interpreting the microbial fossil record and thereby enhance astrobiological studies of martian samples. Our models of biogenic gas emissions will enhance models of atmospheres that might be detected on inhabited extrasolar planets. This work therefore also addresses the question: How can other biospheres be recogniZed" Our choice of field sites helps us explore Earth's evolving early environment. For example, modern mats that occupy thermal springs and certain freshwater

  14. Petroleum source rock evaluation of the Alum and Dictyonema Shales (Upper Cambrian-Lower Ordovician) in the Baltic Basin and Podlasie Depression (eastern Poland)

    NASA Astrophysics Data System (ADS)

    Kosakowski, Paweł; Kotarba, Maciej J.; Piestrzyński, Adam; Shogenova, Alla; Więcław, Dariusz

    2017-03-01

    We present geochemical characteristics of the Lower Palaeozoic shales deposited in the Baltic Basin and Podlasie Depression. In the study area, this strata are represented by the Upper Cambrian-Lower Ordovician Alum Shale recognized in southern Scandinavia and Polish offshore and a equivalent the Lower Tremadocian Dictyonema Shale from the northern Estonia and the Podlasie Depression in Poland. Geochemical analyses reveal that the Alum Shale and Dictyonema Shale present high contents of organic carbon. These deposits have the best source quality among the Lower Palaeozoic strata, and they are the best source rocks in the Baltic region. The bituminous shales complex has TOC contents up to ca. 22 wt%. The analysed rocks contain low-sulphur, oil-prone Type-II kerogen deposited in anoxic or sub-oxic conditions. The maturity of the Alum and Dictyonema Shales changes gradually, from the east and north-east to the west and south-west, i.e. in the direction of the Tornquist-Teisseyre Zone. Samples, located in the seashore of Estonia and in the Podlasie region, are immature and in the initial phase of "oil window". The mature shales were found in the central offshore part of the Polish Baltic Basin, and the late mature and overmature are located in the western part of the Baltic Basin. The Alum and Dictyonema Shales are characterized by a high grade of radioactive elements, especially uranium. The enrichment has a syngenetic or early diagenetic origin. The measured content of uranium reached up to 750 ppm and thorium up to 37 ppm.

  15. Unexpected Early Triassic marine ecosystem and the rise of the Modern evolutionary fauna

    PubMed Central

    Brayard, Arnaud; Krumenacker, L. J.; Botting, Joseph P.; Jenks, James F.; Bylund, Kevin G.; Fara, Emmanuel; Vennin, Emmanuelle; Olivier, Nicolas; Goudemand, Nicolas; Saucède, Thomas; Charbonnier, Sylvain; Romano, Carlo; Doguzhaeva, Larisa; Thuy, Ben; Hautmann, Michael; Stephen, Daniel A.; Thomazo, Christophe; Escarguel, Gilles

    2017-01-01

    In the wake of the end-Permian mass extinction, the Early Triassic (~251.9 to 247 million years ago) is portrayed as an environmentally unstable interval characterized by several biotic crises and heavily depauperate marine benthic ecosystems. We describe a new fossil assemblage—the Paris Biota—from the earliest Spathian (middle Olenekian, ~250.6 million years ago) of the Bear Lake area, southeastern Idaho, USA. This highly diversified assemblage documents a remarkably complex marine ecosystem including at least seven phyla and 20 distinct metazoan orders, along with algae. Most unexpectedly, it combines early Paleozoic and middle Mesozoic taxa previously unknown from the Triassic strata, among which are primitive Cambrian-Ordovician leptomitid sponges (a 200–million year Lazarus taxon) and gladius-bearing coleoid cephalopods, a poorly documented group before the Jurassic (~50 million years after the Early Triassic). Additionally, the crinoid and ophiuroid specimens show derived anatomical characters that were thought to have evolved much later. Unlike previous works that suggested a sluggish postcrisis recovery and a low diversity for the Early Triassic benthic organisms, the unexpected composition of this exceptional assemblage points toward an early and rapid post-Permian diversification for these clades. Overall, it illustrates a phylogenetically diverse, functionally complex, and trophically multileveled marine ecosystem, from primary producers up to top predators and potential scavengers. Hence, the Paris Biota highlights the key evolutionary position of Early Triassic fossil ecosystems in the transition from the Paleozoic to the Modern marine evolutionary fauna at the dawn of the Mesozoic era. PMID:28246643

  16. Unexpected Early Triassic marine ecosystem and the rise of the Modern evolutionary fauna.

    PubMed

    Brayard, Arnaud; Krumenacker, L J; Botting, Joseph P; Jenks, James F; Bylund, Kevin G; Fara, Emmanuel; Vennin, Emmanuelle; Olivier, Nicolas; Goudemand, Nicolas; Saucède, Thomas; Charbonnier, Sylvain; Romano, Carlo; Doguzhaeva, Larisa; Thuy, Ben; Hautmann, Michael; Stephen, Daniel A; Thomazo, Christophe; Escarguel, Gilles

    2017-02-01

    In the wake of the end-Permian mass extinction, the Early Triassic (~251.9 to 247 million years ago) is portrayed as an environmentally unstable interval characterized by several biotic crises and heavily depauperate marine benthic ecosystems. We describe a new fossil assemblage-the Paris Biota-from the earliest Spathian (middle Olenekian, ~250.6 million years ago) of the Bear Lake area, southeastern Idaho, USA. This highly diversified assemblage documents a remarkably complex marine ecosystem including at least seven phyla and 20 distinct metazoan orders, along with algae. Most unexpectedly, it combines early Paleozoic and middle Mesozoic taxa previously unknown from the Triassic strata, among which are primitive Cambrian-Ordovician leptomitid sponges (a 200-million year Lazarus taxon) and gladius-bearing coleoid cephalopods, a poorly documented group before the Jurassic (~50 million years after the Early Triassic). Additionally, the crinoid and ophiuroid specimens show derived anatomical characters that were thought to have evolved much later. Unlike previous works that suggested a sluggish postcrisis recovery and a low diversity for the Early Triassic benthic organisms, the unexpected composition of this exceptional assemblage points toward an early and rapid post-Permian diversification for these clades. Overall, it illustrates a phylogenetically diverse, functionally complex, and trophically multileveled marine ecosystem, from primary producers up to top predators and potential scavengers. Hence, the Paris Biota highlights the key evolutionary position of Early Triassic fossil ecosystems in the transition from the Paleozoic to the Modern marine evolutionary fauna at the dawn of the Mesozoic era.

  17. The Preboreal-like Asian monsoon climate in the early last interglacial period recorded from the Dark Cave, Southwest China

    NASA Astrophysics Data System (ADS)

    Jiang, Xiuyang; He, Yaoqi; Wang, Xiaoyan; Sun, Xiaoshuang; Hong, Hui; Liu, Juan; Yu, Tsai-Luen; Li, Zhizhong; Shen, Chuan-Chou

    2017-08-01

    Transitions of glacial-interglacial cycles are critical periods for Quaternary climate shifts. Here, we present new, decadal resolution Asian summer monsoon (ASM) record from three stalagmites obtained from the Dark Cave in southwestern China over 130-114 thousand years ago (ka, before CE 1950). Chronology was anchored by 28 230Th dates with typical uncertainties of ±0.3-1.0 kyr, allowing an assessment of timing and transition of climate changes during the onset and end of the last interglacial. An agreement between this new and previous stalagmite δ18O records supports that summer insolation predominates orbital-scale ASM evolution. A 2-3 kyr-long gradually increasing ASM period, analogous to the classical Preboreal episode in the early Holocene, follows the termination of a weak monsoon interval at 129.0 ± 0.8 ka. This finding suggests a strong influence of high-latitude ice-sheet dynamics on Asian monsoonal conditions during the early interglacial period. An abrupt end of the marine isotope stage 5e at 118.8 ± 0.6 ka was probably caused by the internal climate system threshold effects.

  18. Student nurses' recognition of early signs of abnormal vital sign recordings.

    PubMed

    Leonard, Martha M; Kyriacos, Una

    2015-09-01

    There is increasing urgency for nurses to recognize early signs of deterioration in patients and to take appropriate action to prevent serious adverse effects. To assess respondents' ability to identify abnormal recordings for respiratory and heart rate, oxygen saturation level, systolic blood pressure, level of consciousness, urinary output and normal temperature. A descriptive observational survey. A nursing college in Cape Town, South Africa. A sample of 77/212 (36.3%) fourth year students. A self-administered adapted questionnaire was employed to collect demographic data and respondents' selections of recorded physiological values for the purpose of deciding when to call for more skilled help. The median age for 62/77 (80.5%) of the respondents was 25years; 3/76 (3.9%) had a previous certificate in nursing. Most respondents were female (66/76, 85.7%). Afrikaans was the first language preference of 33 (42.9%) respondents, followed by isiXhosa (31/77, 40.3%) and English (10/77, 13.0%). Most respondents (48/77, 62.3%) recognized a normal temperature reading (35-38.4°C). However, overall there would have been delays in calling for more skilled assistance in 288/416 (69.2%) instances of critical illness for a high-score MEWS of 3 and in 226/639 (35.4%) instances at a medium-score MEWS of 2 for physiological parameters. In 96/562 (17.1%) instances, respondents would have called for assistance for a low-score MEWS of 1. Non-recognition of deterioration in patients' clinical status and delayed intervention by nurses has implications for the development of serious adverse events. The MEWS is recommended as a track-and-trigger system for nursing curricula in South Africa and for implementation in practice. Copyright © 2015. Published by Elsevier Ltd.

  19. The Early Toarcian Oceanic Anoxic Event and its sedimentary record in Switzerland

    NASA Astrophysics Data System (ADS)

    Fantasia, Alicia; Föllmi, Karl B.; Adatte, Thierry; Spangenberg, Jorge E.; Montero-Serrano, Jean-Carlos

    2015-04-01

    In the Jurassic period, the Early Toarcian Oceanic Anoxic Event (T-OAE), about 183 Ma ago, was a global perturbation of paleoclimatic and paleoenvironmental conditions. This episode was associated with a crisis in marine carbonate accumulation, climate warming, an increase in sea level, ocean acidification, enhanced continental weathering, whereas organic-rich sediments are noticeable for example in the Atlantic and in the Tethys. This episode is associated with a negative carbon excursion, which is recorded both in marine and terrestrial environments. The cause(s) of this environmental crisis remain(s) still controversial. Nevertheless, the development of negative δ13C excursions is commonly interpreted as due to the injection of isotopically-light carbon associated with gas hydrate dissociation, the thermal metamorphism of carbon-rich sediments and input of thermogenic and volcanogenic carbon related to the formation of the Karoo-Ferrar basaltic province in southern Gondwana (Hesselbo et al., 2000, 2007; Beerling et al., 2002; Cohen et al., 2004, 2007; McElwain et al., 2005, Beerling and Brentnall, 2007; Svensen et al., 2007; Hermoso et al., 2009, 2012; Mazzini et al., 2010). Several studies of the T-OAE have been conducted on sediments in central and northwest Europe, but only few data are available concerning the Swiss sedimentary records. Therefore, we focused on two sections in the Jura Plateau (canton Aargau): the Rietheim section (Montero-Serrano et al., submitted) and the Gipf section (current study). A multidisciplinary approach has been chosen and the tools to be used are based on sedimentological observations (sedimentary condensation, etc.), biostratigraphy, mineralogy (bulk-rock composition), facies and microfacies analysis (presence or absence of benthos), clay-mineralogy composition (climatic conditions), major and trace-element analyses (productivity, redox conditions, etc.), phosphorus (trophic levels, anoxia), carbon isotopes and organic

  20. Stratigraphy and depositional environment of upper Cambrian Red Lion Formation, southwestern Montana

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Hayden, L.L.; Bush, J.H.

    1987-08-01

    The Red Lion Formation was examined along a northwest-southeast transect from Missoula to Bozeman, Montana. Lateral equivalents are the Snowy Range Formation east of Bozeman and the upper Fishtrap Dolomite in northwest Montana. The basal Dry Creek Member (0-5 m) consists of shale interbedded with quartz siltstones and sandstones. The overlying Sage Member, up to 115 meters in thickness, is characterized by ribbon carbonate beds containing lime mudstone and quartzose calcisiltite couplets arranged in fining-upward sequences 1-5 cm thick. Couplets are interlayered in places with thin (1-5 cm) to medium bedded (6-70 cm) units of laminated and non-laminated calcareous siltstones,more » flat-pebble conglomerates, trilobite packstones, cryptalgal boundstones, bioturbated lime mudstones and shales. In places, the upper Sage contains columnar and domal algal features. The Red Lion Formation is considered to be one Grand Cycle with the Dry Creek representing a lower inner detrital half-cycle and the Sage an upper carbonate half-cycle. The Dry Creek formed as the result of a westward clastic pulse from the inner detrital belt across an intrashelf basin onto outer middle carbonate peritidal complexes of the underlying Pilgrim Formation. Lower Sage ribbon rocks were deposited in storm-crossed, below wave-base areas. During deposition of the upper Sage, shallowing formed discontinuous algal-peritidal complexes over much of western and central Montana. These complexes were less extensive than earlier Cambrian buildups owing to slower rates of basin subsidence and clastic input suppressing carbonate production.« less

  1. Oceanic oxygenation events in the anoxic Ediacaran ocean.

    PubMed

    Sahoo, S K; Planavsky, N J; Jiang, G; Kendall, B; Owens, J D; Wang, X; Shi, X; Anbar, A D; Lyons, T W

    2016-09-01

    The ocean-atmosphere system is typically envisioned to have gone through a unidirectional oxygenation with significant oxygen increases in the earliest (ca. 635 Ma), middle (ca. 580 Ma), or late (ca. 560 Ma) Ediacaran Period. However, temporally discontinuous geochemical data and the patchy metazoan fossil record have been inadequate to chart the details of Ediacaran ocean oxygenation, raising fundamental debates about the timing of ocean oxygenation, its purported unidirectional rise, and its causal relationship, if any, with the evolution of early animal life. To better understand the Ediacaran ocean redox evolution, we have conducted a multi-proxy paleoredox study of a relatively continuous, deep-water section in South China that was paleogeographically connected with the open ocean. Iron speciation and pyrite morphology indicate locally euxinic (anoxic and sulfidic) environments throughout the Ediacaran in this section. In the same rocks, redox sensitive element enrichments and sulfur isotope data provide evidence for multiple oceanic oxygenation events (OOEs) in a predominantly anoxic global Ediacaran-early Cambrian ocean. This dynamic redox landscape contrasts with a recent view of a redox-static Ediacaran ocean without significant change in oxygen content. The duration of the Ediacaran OOEs may be comparable to those of the oceanic anoxic events (OAEs) in otherwise well-oxygenated Phanerozoic oceans. Anoxic events caused mass extinctions followed by fast recovery in biologically diversified Phanerozoic oceans. In contrast, oxygenation events in otherwise ecologically monotonous anoxic Ediacaran-early Cambrian oceans may have stimulated biotic innovations followed by prolonged evolutionary stasis. © 2016 John Wiley & Sons Ltd.

  2. Sulfur Geochemistry of a Lacustrine Record from Taiwan Reveals Enhanced Marine Aerosol Input during the Early Holocene

    PubMed Central

    Ding, Xiaodong; Li, Dawei; Zheng, Liwei; Bao, Hongyan; Chen, Huei-Fen; Kao, Shuh-Ji

    2016-01-01

    Lacustrine record of marine aerosol input has rarely been documented. Here, we present the sulfur geochemistry during the last deglaciation and early Holocene of a sediment core retrieved from the Dongyuan Lake in southern Taiwan. An unusually high sulfur peak accompanying pyrite presence is observed at 10.5 ka BP. Such high sulfur content in lacustrine record is unusual. The δ34S of sulfur varied from +9.5 to + 17.1‰ with two significant positive shifts at 10.5 and 9.4 ka BP. The sources of sulfur and potential processes involving the sulfur isotope variation including bacterial sulfate reduction, volcanic emissions, in-catchment sulfide oxidation and marine aerosol input are discussed. Enhanced marine aerosol input is the most likely explanation for such sulfur peaks and δ34S shifts. The positive δ34S shifts appeared concurrently with the maximum landslide events over Taiwan resulted from enhanced typhoon activities. The synchronicity among records suggests that increased typhoon activities promoted sea spray, and consequently enhanced the marine aerosol input with 34S-enriched sulfate. Our sulfur geochemistry data revealed sea spray history and marine influence onto terrestrial environment at coastal regions. Wider coverage of spatial-temporal lacustrine sulfur geochemistry record is needed to validate the applicability of sulfur proxy in paleoenvironmental research. PMID:27941864

  3. A molecular palaeobiological exploration of arthropod terrestrialization

    PubMed Central

    Carton, Robert; Edgecombe, Gregory D.

    2016-01-01

    Understanding animal terrestrialization, the process through which animals colonized the land, is crucial to clarify extant biodiversity and biological adaptation. Arthropoda (insects, spiders, centipedes and their allies) represent the largest majority of terrestrial biodiversity. Here we implemented a molecular palaeobiological approach, merging molecular and fossil evidence, to elucidate the deepest history of the terrestrial arthropods. We focused on the three independent, Palaeozoic arthropod terrestrialization events (those of Myriapoda, Hexapoda and Arachnida) and showed that a marine route to the colonization of land is the most likely scenario. Molecular clock analyses confirmed an origin for the three terrestrial lineages bracketed between the Cambrian and the Silurian. While molecular divergence times for Arachnida are consistent with the fossil record, Myriapoda are inferred to have colonized land earlier, substantially predating trace or body fossil evidence. An estimated origin of myriapods by the Early Cambrian precedes the appearance of embryophytes and perhaps even terrestrial fungi, raising the possibility that terrestrialization had independent origins in crown-group myriapod lineages, consistent with morphological arguments for convergence in tracheal systems. This article is part of the themed issue ‘Dating species divergences using rocks and clocks’. PMID:27325830

  4. Paleomagnetism of Early Paleozoic Rocks from the de Long Archipelago and Tectonics of the New Siberian Islands Terrane

    NASA Astrophysics Data System (ADS)

    Metelkin, D. V.; Chernova, A. I.; Matushkin, N. Y.; Vernikovskiy, V. A.

    2017-12-01

    The De Long archipelago is located to the north of the Anjou archipelago as a part of a large group between the Laptev Sea and the East Siberian Sea - the New Siberian Islands and consists of Jeannette Island, Bennett Island and Henrietta Island. These islands have been shown to be part of a single continental terrane, whose tectonic history was independent of other continental masses at least since the Ordovician. Paleomagnetic and precise geological data for the De Long archipelago were absent until recently. Only in 2013 special international field trips to the De Long Islands could be organized and geological, isotope-geochronological and paleomagnetic studies were carried out.On Jeannette Island a volcanic-sedimentary sequence intruded by mafic dikes was described. The age of these dikes is more likely Early Ordovician, close to 480 Ma, as evidenced by the results of our 40Ar/39Ar and paleomagnetic investigations of the dolerites as well as the result from detrital zircons in the host rocks published before. On Bennett Island, there are widespread Cambrian-Ordovician mainly terrigenous rocks. Paleomagnetic results from these rocks characterize the paleogeographic position of the De Long archipelago at 465 Ma and perhaps at 530 Ma, although there is no evidence for the primary origin of magnetization for the latter. On Henrietta Island the Early Cambrian volcanic-sedimentary section was investigated. A paleomagnetic pole for 520 Ma was obtained and confirmed by new 40Ar/39Ar results. Adding to our previous paleomagnetic data for the Anjou archipelago the extended variant of the apparent polar wander path for the New Siberian Island terrane was created. The established paleolatitudes define its location in the equatorial and subtropical zone no higher than 40 degrees during the Early Paleozoic. Because there are no good confirmations for true polarity and related geographic hemisphere we present two possibilities for tectonic reconstruction. But both these

  5. An Early Middle Eocene Orbital Scale Benthic Isotope Record From IODP Site 1408, Newfoundland Rise

    NASA Astrophysics Data System (ADS)

    Wu, F.; Lawler, N.; Penman, D. E.; Zachos, J. C.; Kirtland Turner, S.; Norris, R. D.; Wilson, P. A.; Hull, P. M.

    2014-12-01

    The long-term Paleogene global cooling trend and eventual glaciation of Antarctica has been attributed to a reduction in greenhouse gas levels as well as changes in the configuration of high-latitude oceanic gateways. This major trend in climate and forcing is known to have initiated in the early middle Eocene, between 44-49 Mya, yet our understanding of the detailed evolution of climate and oceanic circulation and carbon chemistry of this critical interval has been limited for lack of high-resolution proxy climate records. Integrated Ocean Drilling Program (IODP) Expedition 342, designed in part to address this deficiency, successfully recovered highly expanded sequences of middle Eocene sediment from multiple sites in the western North Atlantic, with several sites characterized by high sedimentation rates (>2.8 cm/kyr) and pronounced lithologic cycles. Using samples from cores recovered at one of these sites, 1408, located on Southeast Newfoundland Ridge, we are reconstructing the first orbital-scale deep sea δ18O and δ13C records spanning a ~1.6 million year interval (~Chron 20r) of the middle Eocene. Based on analyses of benthic foraminifer N. truempyi, our preliminary data reveal distinct high-frequency cycles with periods matching those of the orbital cycles, particularly precession and obliquity. Cross spectral analysis of δ18O, δ13C and lithologic records reveal a high degree of coherency, implying a high sensitivity in local sediment fluxes and bottom water chemistry (and circulation) to orbital forcing. Also, given the location and depth (~2600 m at 50 Ma), Site 1408 constrains the end-member composition of northern component bathyal bottom waters so that comparison with benthic isotope records from the south Atlantic and other basins can be used to assess ocean circulation patterns in the mid-Eocene. In general, bottom water temperatures appear to have been warmer, and DIC δ13C lower than observed elsewhere. Thus, our preliminary results are

  6. Varied records of early Wisconsinan alpine glaciation in the western United States derived from weathering-rind thicknesses

    USGS Publications Warehouse

    Clark, Peter U.; Lea, P.D.

    1992-01-01

    Weathering-rind thicknesses were measured on volcanic clasts in sequences of glacial deposits in seven mountain ranges in the western United States and in the Puget lowland. Because the rate of rind development decreases with time, ratios of rind thicknesses provide limits on corresponding age ratios. In all areas studied, deposits of late Wisconsinan age are obvious; deposits of late Illinoian age (ca. 140 ka) also seem to be present in each area, although independent evidence for their numerical age is circumstantial. The weathering-rind data indicate that deposits that have intermediate ages between these two are common, and ratios of rind thicknesses suggest an early Wisconsinan age (about 60 to 70 ka) for some of the intermediate deposits. Three of the seven studied alpine areas (McCall, Idaho; Yakima Valley, Washington; and Lassen Peak, California) appear to have early Wisconsinan drift beyond the extent of late Wisconsinan ice. In addition, Mount Rainier and the Puget lowland, Washington, have outwash terraces but no moraines of early Wisconsinan age. The sequences near West Yellowstone, Montana; Truckee, California; and in the southern Olympic Mountains have no recognized moraines or outwash of this age. Many of the areas have deposits that may be of middle Wisconsinan age.Differences in the relative extents of early Wisconsinan alpine glaciers are not expected from the marine oxygen-isotope record and are not explained by any simple trend in climatic variables or proximity to oceanic moisture sources. However, alpine glaciers could have responded more quickly and more variably than continental ice sheets to intense, short-lived climatic events, and they may have been influenced by local climatic or hypsometric effects. The relative sizes of early and late Wisconsinan alpine glaciers could also reflect differences between early and late Wisconsinan continental ice sheets and their regional climatic effects.

  7. Astronomically forced paleoclimate change from middle Eocene to early Oligocene: continental conditions in central China compared with the global marine isotope record

    NASA Astrophysics Data System (ADS)

    Huang, C.; Hinnov, L. A.

    2010-12-01

    The early Eocene climatic optimum ended with a long interval of global cooling that began in the early Middle Eocene and ended at the Eocene-Oligocene transition. During this long-term cooling, a series of short-term warming reversals occurred in the marine realm. Here, we investigate corresponding continental climate conditions as revealed in the Qianjiang Formation of the Jianghan Basin in central China, which consists of more than 4000 m of saline lake sediments. The Qianjiang Formation includes, in its deepest sections, a halite-rich rhythmic sediment succession with dark mudstone, brownish-white siltstone and sandstone, and greyish-white halite. Alternating fresh water (humid/cool)—saline water (dry/hot) deposits reflect climate cycles driven by orbital forcing. High-resolution gamma ray (GR) logging from the basin center captures these pronounced lithological rhythms throughout the formation. Several halite-rich intervals are interpreted as short-term warming events within the middle Eocene to early Oligocene, and could be expressions of coeval warming events in the global marine oxygen isotope record, for example, the middle Eocene climate optimum (MECO) event around 41 Ma. The Eocene-Oligocene boundary is distinguished by a radical change from halite-rich to clastic sediments, indicating a dramatic climate change from warm to cool conditions. Power spectral analysis of the GR series indicates strong short (~100 kyr) eccentricity cycling during the warm/hot episodes. Amplitude modulation of the short eccentricity in the GR series occurs with a strong 405 kyr periodicity. This cycling is calibrated to the La2004 orbital eccentricity model. A climate reversal occurs at 36.5 Ma within the long-term marine cooling trend following MECO, which is reflected also in the Qianjiang GR series, with the latter indicating several brief warm/dry reversals within the trend. A ~2.6 Myr halite-rich warm interval occurs in the latest Eocene in the continental record; both

  8. Hallucigenia's head and the pharyngeal armature of early ecdysozoans.

    PubMed

    Smith, Martin R; Caron, Jean-Bernard

    2015-07-02

    The molecularly defined clade Ecdysozoa comprises the panarthropods (Euarthropoda, Onychophora and Tardigrada) and the cycloneuralian worms (Nematoda, Nematomorpha, Priapulida, Loricifera and Kinorhyncha). These disparate phyla are united by their means of moulting, but otherwise share few morphological characters--none of which has a meaningful fossilization potential. As such, the early evolutionary history of the group as a whole is largely uncharted. Here we redescribe the 508-million-year-old stem-group onychophoran Hallucigenia sparsa from the mid-Cambrian Burgess Shale. We document an elongate head with a pair of simple eyes, a terminal buccal chamber containing a radial array of sclerotized elements, and a differentiated foregut that is lined with acicular teeth. The radial elements and pharyngeal teeth resemble the sclerotized circumoral elements and pharyngeal teeth expressed in tardigrades, stem-group euarthropods and cycloneuralian worms. Phylogenetic results indicate that equivalent structures characterized the ancestral panarthropod and, seemingly, the ancestral ecdysozoan, demonstrating the deep homology of panarthropod and cycloneuralian mouthparts, and providing an anatomical synapomorphy for the ecdysozoan supergroup.

  9. Isotopic and geochemical characterization of fossil brines of the Cambrian Mt. Simon Sandstone and Ironton-Galesville Formation from the Illinois Basin, USA

    NASA Astrophysics Data System (ADS)

    Labotka, Dana M.; Panno, Samuel V.; Locke, Randall A.; Freiburg, Jared T.

    2015-09-01

    Geochemical and isotopic characteristics of deep-seated saline groundwater provide valuable insight into the origin and evolving composition, water-rock interaction, and mixing potential of fossil brines. Such information may yield insight into intra- and interbasinal brine movement and relationships between brine evolution and regional groundwater flow systems. This investigation reports on the δ18O and δD composition and activity values, 87Sr/86Sr ratios and Sr concentrations, and major ion concentrations of the Cambrian-hosted brines of the Mt. Simon Sandstone and Ironton-Galesville Formation and discusses the evolution of these brines as they relate to other intracontinental brines. Brines in the Illinois Basin are dominated by Na-Ca-Cl-type chemistry. The Mt. Simon and overlying Ironton-Galesville brines exhibit total dissolved solids concentrations of ∼195,000 mg/L and ∼66,270 mg/L, respectively. The δD of brine composition of the Mt. Simon ranges from -34‰ to -22‰ (V-SMOW), and the Ironton-Galesville is ∼-53.2‰ (V-SMOW). The δ18O composition of the Mt. Simon brine ranges from -5.0‰ to -2.8‰ (V-SMOW), and the Ironton-Galesville brine is ∼-6.9‰ (V-SMOW). The 87Sr/86Sr values in the Mt. Simon brine range from 0.7110 to 0.7116. The less radiogenic Ironton-Galesville brine has an average 87Sr/86Sr value of 0.7107. Evaluation of δ18O and δD composition and activities and 87Sr/86Sr ratios suggests that the Mt. Simon brine is likely connate seawater and recirculating deep-seated brines that have been diluted with meteoric water and influenced by the dissolution of evaporites with a minimal halite contribution based on Cl/Br ratios. The Ironton-Galesville brine is also likely originally connate seawater that mixed with other brines and meteoric waters, including possibly Pleistocene glacial recharge. The Ca-excess vs. Na-deficiency comparison with the Basinal Fluid Line suggests the Mt. Simon and Ironton-Galesville brines have been

  10. Suspected microbial-induced sedimentary structures (MISS) in Furongian (Upper Cambrian; Jiangshanian, Sunwaptan) strata of the Upper Mississippi Valley

    USGS Publications Warehouse

    Eoff, Jennifer D.

    2014-01-01

    The Furongian (Upper Cambrian; Jiangshanian and Sunwaptan) Tunnel City Group (Lone Rock Formation and Mazomanie Formation), exposed in Wisconsin and Minnesota, represents a shallow-marine clastic environment during a time of exceptionally high sea level. Lithofacies from shoreface to transitional-offshore settings document deposition in a wave- and storm-dominated sea. Flooding of the cratonic interior was associated with formation of a condensed section and the extensive development of microbial mats. Biolamination, mat fragments, wrinkle structures, and syneresis cracks are preserved in various sandstone facies of the Lone Rock Formation, as is evidence for the cohesive behavior of sand. These microbial-induced sedimentary structures (MISS) provide unique signals of biological–physical processes that physical structures alone cannot mimic. The MISS are associated with a trilobite extinction event in the Steptoean–Sunwaptan boundary interval. This may support recent claims that Phanerozoic microbial mats were opportunistic disaster forms that flourished during periods of faunal turnover. Further investigation of stratigraphic, taphonomic, and other potential biases, however, is needed to fully test this hypothesis.

  11. Variscan ore formation and metamorphism at the Felbertal scheelite deposit (Austria): constraining tungsten mineralisation from Re-Os dating of molybdenite

    NASA Astrophysics Data System (ADS)

    Raith, Johann G.; Stein, Holly J.

    2006-10-01

    The Felbertal scheelite deposit in the Eastern Alps has been regarded as the type locality for stratabound scheelite deposits. It is hosted by a Cambro-Ordovician metavolcanic arc sequence with minor Variscan granitoids (˜ 340 Ma) in the central Tauern Window. Re-Os model ages for molybdenite from the Felbertal tungsten deposit range between ˜ 358 and ˜ 336 Ma and record several pulses of magmatic-hydrothermal-metamorphic molybdenite formation. Molybdenite ages from the K2 orebody, a scheelite-rich quartz mylonite in the Western ore field, indicate that both mineralisation and mylonite are Variscan in age and suggest that the shear zone was active for ˜ 20 million years. Early stage tungsten mineralisation ( Scheelite 1) in quartzitic ores in the Eastern ore field, which is free of molybdenite, yielded very low to near blank levels of Re and Os and thus could not be dated. However, molybdenite from scheelite-quartz stringers, previously interpreted as a feeder stockwork to quartzitic scheelite ore of presumed Cambrian age, yielded Variscan Re-Os ages of ˜ 342 and ˜ 337 Ma. Dating of molybdenite contained in scheelite ores thus far provides no indication of a Cambrian component to the tungsten mineralisation. Our data are consistent with a model of either granite intrusion-related ore formation and coeval metamorphic overprint during the Early Carboniferous or, alternatively, molybdenite formation may be exclusively attributed to Variscan metamorphism (see Stein 2006).

  12. Relations between extensional tectonics and magmatism within the Southern Oklahoma aulacogen

    NASA Technical Reports Server (NTRS)

    Mcconnell, D. A.; Gilbert, M. C.

    1985-01-01

    Variations in the geometry, distribution and thickness of Cambrian igneous and sedimentary units within southwest Oklahoma are related to a late Proterozoic - early Paleozoic rifting event which formed the Southern Oklahoma aulacogen. These rock units are exposed in the Wichita Mountains, southwest Olkahoma, located on the northern margin of a Proterozoic basin, identified in the subsurface by COCORP reflection data. Overprinting of the Cambrian extensional event by Pennyslvanian tectonism obsured the influence of pre-existing basement structures and contrasting basement lithologies upon the initial development of the aulacogen.

  13. Early organisms in the fossil record: paleontological aspects, evolutionary and ecological impacts

    NASA Astrophysics Data System (ADS)

    Sabbatini, Anna; Negri, Alessandra; Morigi, Caterina; Bartolini, Annachiara; Lipps, Jere

    2017-04-01

    With this abstract we introduce our session whose aim is twofold: 1) to gather information on the earliest foraminifera (single- organic and agglutinated taxa) which so far are sparse and uncoordinated in order to understand their evolution and their relationship with modern single-chambered taxa, contextualizing scientific current results in the geo-biological field. 2) to explore also every other early organism trace fossils or so far overlooked organisms coated with fine sediment (i.e., bacteria, testate amoebae) to understand how and if this coating might help these creatures to fossilize. For this reason, this session will integrate many disciplines, from genomics to palaeo-environmental modelling to palaeontology and geochemistry. Our experience starts from Foraminifera which are an ecologically important group of modern heterotrophic amoeboid eukaryotes whose naked and testate ancestors are thought to have evolved 1 Ga ago. However, the single-chambered agglutinated test of these protists is hypothesized to appear in the fossil record in the Neoproterozoic, before the rise of complex animals. In addition, the difficulty of recognizing unambiguously ancestral monothalamous foraminifera in the fossil record represents the main challenge and might be related to a combination of factors, such as preservation in the sediments, adverse palaeo-environmental conditions and the absence of clear morphological characters distinguishing them from other morphologically simple testate organisms. However, recent publications have evidenced the finding of such organisms in several sedimentary successions tracing back to the Neoproterozoic. An integrate approach will result in profound insights about life—past, present, future— representing a new frontier in the palaeobiological studies. Therefore, aim of this session is to bring together specialists across all these disciplines to provide a uniquely rich and fertile intellectual environment for the pursuit of this

  14. Millennial-scale Climate Variations Recorded As Far Back As The Early Pliocene

    NASA Astrophysics Data System (ADS)

    Steenbrink, J.; Hilgen, F. J.; Lourens, L. J.

    Quaternary climate proxy records show compelling evidence for climate variability on time scales of a few thousand years. The causes for these millennial-scale or sub- Milankovitch cycles are yet poorly understood, not in the least due to the complex feedback mechanisms of large ice-sheets during the Quaternary. We present evidence of millennial-scale climate variability in Early Pliocene lacustrine sediments from the intramontane Ptolemais Basin in northwestern Greece. The sediments are well ex- posed in a series of open-pit lignite mines and exhibit a distinct m-scale sedimentary cyclicity of alternating lignites and lacustrine marl beds that result from precession- induced variations in climate. A higher-frequency cyclicity is particular prominent within the marl segment of individual cycles. A stratigraphic interval of~115 kyr, cov- ering five precession-induced sedimentary cycles, was studied in nine parallel sections from two quarries located several km apart. Colour reflectance records were used to quantify the within-cycle variability and to determine its lateral continuity. Much of the within-cycle variability could be correlated between the parallel sections, even in fine detail, which suggests that these changes reflect basin-wide variations in environ- mental conditions related to (regional) climate fluctuations. Interbedded volcanic ash beds demonstrate the synchronicity of these fluctuations and spectral analysis of the reflectance time series shows a significant concentration of variability at periods of ~11,~5.5 and~2 kyr. Their occurrence at times before the intensification of the North- ern Hemisphere glaciation suggests that they cannot solely have resulted from internal ice-sheet dynamics. Possible candidates include harmonics or combination tones of the main orbital cycles, variations in solar output or periodic motions of the Earth and moon.

  15. Climate change and tectonic activity during the early Pliocene Warm Period from the ostracod record at Lake Qinghai, northeastern Tibetan Plateau

    NASA Astrophysics Data System (ADS)

    Lu, Fengyan; An, Zhisheng; Chang, Hong; Dodson, John; Qiang, Xiaoke; Yan, Hong; Dong, Jibao; Song, Yougui; Fu, Chaofeng; Li, Xiangzhong

    2017-05-01

    The Early Pliocene Warm Period (EPWP, 5-3 Ma) is sometimes thought to be a useful analogue for a future warmer world, and thus the boundary conditions and drivers of climate in the EPWP may provide valuable lessons for understanding how a future warmer world might unfold. Lake Qinghai is located on the northeastern margin of the Tibetan Plateau (TP) and is affected by both Monsoon climate and Westerlies circulation. It is sensitive to the climate drivers of these systems. Its sediments, accumulated over the Cenozoic period, are a rich source of information for climate, tectonics and environmental changes of the period. We present a high-resolution ostracod record from a Lake Qinghai sediment core with a record of the period 5.10-2.60 Ma, thus covering the EPWP. Ostracods appear at 4.63 Ma and are most abundant until 3.58 Ma, while a body of water was present at the core site. This suggests a phase of humid climate and an intensified Asian Summer Monsoon (ASM), which is consistent with a warmer and wetter climate in the early Pliocene. Within this period the ostracod record shows some variabilities in lake level with deeper periods suggesting more intense ASM compared to those with shallower water. The disappearance of ostracods at 3.58 Ma may provide evidence for the uplift of Qinghai Nanshan (south of Qinghai Lake) since this is when the ASM intensified.

  16. Geology, structure and age of the Nahuel Niyeu Formation in the Aguada Cecilio area, North Patagonian Massif, Argentina

    NASA Astrophysics Data System (ADS)

    Greco, Gerson A.; González, Pablo D.; González, Santiago N.; Sato, Ana M.; Basei, Miguel A. S.; Tassinari, Colombo C. G.; Sato, Kei; Varela, Ricardo; Llambías, Eduardo J.

    2015-10-01

    The low-grade Nahuel Niyeu Formation in the Aguada Cecilio area (40°50‧S-65°53‧W) shows ultramafic to felsic metaigneous rocks forming a sill swarm intercalated in the metasedimentary sequence and a polyphase deformation which permit an integrated study of the magmatic and tectonometamorphic evolution of this geological unit. In this paper we present a geological characterization of the Nahuel Niyeu Formation in the Aguada Cecilio area combining mapping, structural and metamorphic analysis with a SHRIMP U-Pb age and geochemical data from the metaigneous rocks. The metasedimentary sequence consists of alternating metagreywackes and phyllites, and minor metasandstones and granule metaconglomerates. The sills are pre-kinematic intrusions and yielded one SHRIMP U-Pb, zircon crystallization age of 513.6 ± 3.3 Ma. Their injection occurred after consolidation of the sedimentary sequence. A syn-sedimentary volcanic activity is interpreted by a metaandesite lava flow interlayered in the metasedimentary sequence. Sedimentary and igneous protoliths of the Nahuel Niyeu Formation would have been formed in a continental margin basin associated with active magmatic arc during the Cambrian Epoch 2. Two main low-grade tectonometamorphic events affected the Nahuel Niyeu Formation, one during the Cambrian Epoch 2-Early Ordovician and the other probably in the late Permian at ˜260 Ma. Local late folds could belong to the final stages of the late Permian deformation or be even younger. In a regional context, the Nahuel Niyeu and El Jagüelito formations and Mina Gonzalito Complex show a comparable Cambrian-Ordovician evolution related to the Terra Australis Orogen in the south Gondwana margin. This evolution is also coeval with the late and early stages of the Pampean and Famatinian orogenies of Central Argentina, respectively. The late Permian event recorded in the Nahuel Niyeu Formation in Aguada Cecilio area is identified by comparable structures affecting the Mina Gonzalito

  17. Reinterpreting the Early Cretaceous Sulfur Isotope Records: Implications for the Evolution of Seawater Chemistry

    NASA Astrophysics Data System (ADS)

    Mills, J. V.; Gomes, M. L.; Sageman, B. B.; Jacobson, A. D.; Hurtgen, M. T.

    2013-12-01

    The geologic record of the Cretaceous is punctuated by several periods of high organic carbon burial interpreted to represent global Ocean Anoxic Events (OAEs). In addition to the short-term (<1-Myr) changes in carbon (C) cycling associated with OAEs, evidence from a number of geochemical proxies has been interpreted to represent large-scale changes in ocean chemistry during the period. Specifically, the sulfur (S) isotope composition of early Cretaceous seawater sulfate as recorded in marine barite exhibits an ~5 permil shift in d34Ssulfate that persists for ~15Myr before returning to pre-excursion values. Superimposed upon this long-term shift in S-isotopes is OAE1a, the second major anoxic event recognized in the Cretaceous. Two hypotheses have been proposed to explain this S isotope perturbation: (1) massive evaporite deposition associated with rifting during the opening of the South Atlantic and a corresponding decrease in pyrite burial rates and (2) increased inputs of volcanic-derived S due to extensive LIP-volcanism. While there is geologic evidence for both evaporite deposition and enhanced hydrothermal activity, the relative influence of these potential driving factors remains largely unconstrained. Variation in the strontium (Sr) isotope composition of marine carbonates provides a tool for distinguishing between these influences. We examine the S isotope composition of carbonate-associated sulfate (CAS) spanning the Barremian through Aptian from Resolution Guyot (ODP Site 866) and compare the S isotope record to time equivalent records of carbon and strontium isotopes. Correlative changes in the C, S, and Sr cycles are observed: an ~5 permil shift in d34Ssulfate, which begins at the onset of OAE1a and continues after the positive d13Ccarb excursion, is accompanied by a contemporaneous, parallel shift in 87Sr/86Sr to unradiogenic values. The tight coupling observed between S and Sr throughout the interval is highly suggestive of a common driving mechanism

  18. Early-Mid Holocene climatic variations in Tasmania, Australia: multi-proxy records in a stalagmite from Lynds Cave

    NASA Astrophysics Data System (ADS)

    Xia, Qikai; Zhao, Jian-xin; Collerson, K. D.

    2001-12-01

    Mass spectrometric uranium-series dating and C-O isotopic analysis of a stalagmite from Lynds Cave, northern Tasmania, Australia provide a high-resolution record of regional climate change between 5100 and 9200 yr before present (BP). Combined δ18O, δ13C, growth rate, initial 234U/238U and physical property (color, transparency and porosity) records allow recognition of seven climatic stages: Stage I (>9080 yr BP) - a relatively dry period at the beginning of stalagmite growth evidenced by elevated 234U/238U; Stage II (9080-8600 yr BP) - a period of unstable climate characterized by high-frequency variability in temperature and bio-productivity; Stage III (8600-8000 yr BP) - a period of stable and moderate precipitation and stable and high bio-productivity, with a continuously rising temperature; Stage IV (8000-7400 yr BP) - the warmest period with high evaporation and low effective precipitation (rainfall less evaporation); Stage V (7400-7000 yr BP) - the wettest period with highest stalagmite growth and enhanced but unstable bio-productivity; Stage VI (7000-6600 yr BP) - a period with a significantly reduced precipitation and bio-productivity without noticeable change in temperature; Stage VII (6600-5100 yr BP) - a period of lowest temperature and precipitation marking a significant climatic deterioration. Overall, the records suggest that the warmest climate occurred between 8000 and 7400 yr BP, followed by a wettest period between 7400 and 7000 yr BP. These are broadly correlated with the so-called 'Mid Holocene optimum' previously proposed using pollen and lake level records. However, the timing and resolution of the speleothem record from Lynds Cave are significantly higher than in both the pollen and lake level records. This allows us to correlate the abrupt change in physical property, δ18O, δ13C, growth rate, and initial 234U/238U of the stalagmite at ˜8000 yr BP with a global climatic event at Early-Mid Holocene transition.

  19. 45 CFR 149.350 - Maintenance of records.

    Code of Federal Regulations, 2012 CFR

    2012-10-01

    ... REQUIREMENTS FOR THE EARLY RETIREE REINSURANCE PROGRAM Reimbursement Methods § 149.350 Maintenance of records..., or longer if otherwise required by law. (b) The records that must be retained are as follows— (1) All...

  20. 45 CFR 149.350 - Maintenance of records.

    Code of Federal Regulations, 2013 CFR

    2013-10-01

    ... REQUIREMENTS FOR THE EARLY RETIREE REINSURANCE PROGRAM Reimbursement Methods § 149.350 Maintenance of records..., or longer if otherwise required by law. (b) The records that must be retained are as follows— (1) All...

  1. 45 CFR 149.350 - Maintenance of records.

    Code of Federal Regulations, 2014 CFR

    2014-10-01

    ... REQUIREMENTS FOR THE EARLY RETIREE REINSURANCE PROGRAM Reimbursement Methods § 149.350 Maintenance of records..., or longer if otherwise required by law. (b) The records that must be retained are as follows— (1) All...

  2. Early Pliocene anuran fossils from Kanapoi, Kenya, and the first fossil record for the African burrowing frog Hemisus (Neobatrachia: Hemisotidae).

    PubMed

    Delfino, Massimo

    2017-07-13

    Isolated amphibian bones from the early Pliocene of Kanapoi (West Turkana, Kenya) help to improve the scarce fossil record of the late Neogene and Quaternary amphibians from East Africa. All currently available 579 bones are referable exclusively to the Anura (frogs and toads). More than half of the remains (366) are identified as Hemisus cf. Hemisus marmoratus, an extant species that still inhabits Kenya, but apparently not the northwest of the country and the Turkana area in particular. The rest of the remains are identified simply as Anura indet. because of poor preservation or non congruence with the relatively few African extant taxa whose osteology is known in detail. The Hemisus material represents the first fossil record for Hemisotidae, an endemic African family of peculiar, head-first burrowing frogs, whose sister taxon relationships indicate a divergence from brevicipitids in the Late Cretaceous or early Paleocene. The ecological requirements of extant H. marmoratus suggest that the Kanapoi area surrounding the fluvial and deltaic settings, from where the fossil remains of vertebrates were buried, was likely a grassland or relatively dry, open low tree-shrub savanna. Copyright © 2017 Elsevier Ltd. All rights reserved.

  3. The origin and early evolution of Sauria: reassessing the permian Saurian fossil record and the timing of the crocodile-lizard divergence.

    PubMed

    Ezcurra, Martín D; Scheyer, Torsten M; Butler, Richard J

    2014-01-01

    Sauria is the crown-group of Diapsida and is subdivided into Lepidosauromorpha and Archosauromorpha, comprising a high percentage of the diversity of living and fossil tetrapods. The split between lepidosauromorphs and archosauromorphs (the crocodile-lizard, or bird-lizard, divergence) is considered one of the key calibration points for molecular analyses of tetrapod phylogeny. Saurians have a very rich Mesozoic and Cenozoic fossil record, but their late Paleozoic (Permian) record is problematic. Several Permian specimens have been referred to Sauria, but the phylogenetic affinity of some of these records remains questionable. We reexamine and review all of these specimens here, providing new data on early saurian evolution including osteohistology, and present a new morphological phylogenetic dataset. We support previous studies that find that no valid Permian record for Lepidosauromorpha, and we also reject some of the previous referrals of Permian specimens to Archosauromorpha. The most informative Permian archosauromorph is Protorosaurus speneri from the middle Late Permian of Western Europe. A historically problematic specimen from the Late Permian of Tanzania is redescribed and reidentified as a new genus and species of basal archosauromorph: Aenigmastropheus parringtoni. The supposed protorosaur Eorasaurus olsoni from the Late Permian of Russia is recovered among Archosauriformes and may be the oldest known member of the group but the phylogenetic support for this position is low. The assignment of Archosaurus rossicus from the latest Permian of Russia to the archosauromorph clade Proterosuchidae is supported. Our revision suggests a minimum fossil calibration date for the crocodile-lizard split of 254.7 Ma. The occurrences of basal archosauromorphs in the northern (30°N) and southern (55°S) parts of Pangea imply a wider paleobiogeographic distribution for the group during the Late Permian than previously appreciated. Early archosauromorph growth

  4. The Origin and Early Evolution of Sauria: Reassessing the Permian Saurian Fossil Record and the Timing of the Crocodile-Lizard Divergence

    PubMed Central

    Ezcurra, Martín D.; Scheyer, Torsten M.; Butler, Richard J.

    2014-01-01

    Sauria is the crown-group of Diapsida and is subdivided into Lepidosauromorpha and Archosauromorpha, comprising a high percentage of the diversity of living and fossil tetrapods. The split between lepidosauromorphs and archosauromorphs (the crocodile-lizard, or bird-lizard, divergence) is considered one of the key calibration points for molecular analyses of tetrapod phylogeny. Saurians have a very rich Mesozoic and Cenozoic fossil record, but their late Paleozoic (Permian) record is problematic. Several Permian specimens have been referred to Sauria, but the phylogenetic affinity of some of these records remains questionable. We reexamine and review all of these specimens here, providing new data on early saurian evolution including osteohistology, and present a new morphological phylogenetic dataset. We support previous studies that find that no valid Permian record for Lepidosauromorpha, and we also reject some of the previous referrals of Permian specimens to Archosauromorpha. The most informative Permian archosauromorph is Protorosaurus speneri from the middle Late Permian of Western Europe. A historically problematic specimen from the Late Permian of Tanzania is redescribed and reidentified as a new genus and species of basal archosauromorph: Aenigmastropheus parringtoni. The supposed protorosaur Eorasaurus olsoni from the Late Permian of Russia is recovered among Archosauriformes and may be the oldest known member of the group but the phylogenetic support for this position is low. The assignment of Archosaurus rossicus from the latest Permian of Russia to the archosauromorph clade Proterosuchidae is supported. Our revision suggests a minimum fossil calibration date for the crocodile-lizard split of 254.7 Ma. The occurrences of basal archosauromorphs in the northern (30°N) and southern (55°S) parts of Pangea imply a wider paleobiogeographic distribution for the group during the Late Permian than previously appreciated. Early archosauromorph growth

  5. An Intracratonic Record of North American Tectonics

    NASA Astrophysics Data System (ADS)

    Lovell, Thomas Rudolph

    demonstrates that changes in the detrital record of the Illinois Basin coincide with well-documented, major tectonic and eustatic events that altered and shaped North American plate margins. Chapter 4 presents 24 apatite (U-Th)/He (AHe) ages (3 - 423 Ma) taken from subsurface Cambrian and Pennsylvanian sandstones in the Illinois Basin. Time-temperature simulations used to reproduce these ages predict a basin thermal history with a maximum temperature of 170°C in post-Pennsylvanian time followed by Mesozoic cooling at 0.3°C/Myr. These thermal simulations suggest 3 km of additional post-Pennsylvanian burial (assuming 30°C/km geotherm) followed by subsequent Mesozoic - Cenozoic removal. This burial-exhumation history is concurrent with Late Mesozoic tectoniceustatic fluctuations, including Atlantic and Gulf of Mexico opening, rejuvenation of the Appalachian region, and Gulf of Mexico sediment influx, and the Cretaceous high sea level stand. The Geochronologic and thermochronologic evidence presented in the following chapters suggests the Illinois Basin potentially contains a more robust record of North American tectonics than previously thought. These observations provide a new perspective on the utility of intracratonic basins in understanding long term changes to continental bodies.

  6. Detection of Mild Cognitive Impairment and early stage dementia with an audio-recorded cognitive scale

    PubMed Central

    Sewell, Margaret C.; Luo, Xiaodong; Neugroschl, Judith; Sano, Mary

    2014-01-01

    BACKGROUND Physicians often miss a diagnosis of Mild Cognitive Impairment (MCI) or early dementia and screening measures can be insensitive to very mild impairments. Other cognitive assessments may take too much time or be frustrating to seniors. This study examined the ability of an audio-recorded scale, developed in Australia, to detect MCI or mild Alzheimer’s disease and compared cognitive domain specific performance on the audio-recorded scale to in-person battery and common cognitive screens. METHOD Seventy-six subjects from the Mount Sinai Alzheimer’s Disease Research Center were recruited. Subjects were 75 years or older, with clinical diagnosis of AD or MCI (n=51) or normal control (n=25). Participants underwent in-person neuropsychological testing followed by testing with the Audio-recorded Cognitive Screen (ARCS). RESULTS ARCS provided better discrimination between normal and impaired elders than either the Mini-Mental Status Exam (MMSE) or the clock drawing test. The in-person battery and ARCS analogous variables were significantly correlated, most in the .4 to .7 range, including verbal memory, executive function/attention, naming, and verbal fluency. The area under the curve generated from ROC curves indicated high and equivalent discrimination for ARCS and the in-person battery (0.972 vs. 0.988; p=0.23). CONCLUSION The ARCS demonstrated better discrimination between normal controls and those with mild deficits than typical screening measures. Performance on cognitive domains within the ARCS was well correlated with the in-person battery. Completion of the ARCS was accomplished despite mild difficulty hearing the instructions even in very elderly subjects, indicating that it may be a useful measure in primary care settings. PMID:23635663

  7. Early- to Mid-Holocene hydroclimate shifts in tropical East Africa: the multi-proxy sediment record from Lake Rutundu, Kenya

    NASA Astrophysics Data System (ADS)

    De Cort, Gijs; Creutz, Mike; Barao, Lucia; Conley, Daniel; Haug, Gerald; Bodé, Samuel; Blaauw, Maarten; Engstrom, Dan; Verschuren, Dirk

    2015-04-01

    Following the generally arid conditions of the Last Glacial Maximum (LGM), a large part of the African continent experienced the Early to Mid-Holocene as a much more humid period than today. This so-called African Humid Period (AHP) coincided with high summertime insolation over the Northern Hemisphere subtropics, causing invigorated monsoons to create moist conditions over the northern parts of the continent. Similarly, equatorial and even low-latitude southeastern Africa experienced a wetter climate due to the post-glacial increase in atmospheric greenhouse gasses ultimately leading to altered Atlantic and Indian Ocean monsoon dynamics. The timing and abruptness of the onset and ending of the AHP in the different regions of the continent have been the subject of major discussion. On the other hand, shorter-lived climate fluctuations within the AHP have received much less attention, due to a scarcity of well-dated, high-resolution African paleoclimate records spanning the entire Holocene. In this study we used the sediment record of Lake Rutundu, a high-altitude crater lake on Mount Kenya, to document multidecadal to millennial-scale hydroclimate variability on the East African equator from the LGM to the present. A multiproxy approach combining core-surface scanning techniques (magnetic susceptibility, X-ray fluorescence) and close-interval bulk-sediment analyses (organic matter and biogenic Si content, grain size, organic δ15N and δ13C) resulted in a high-resolution record firmly anchored in time by an age model based on 210Pb dating and sixteen calibrated radiocarbon ages. This new Lake Rutundu hydroclimate record confirms that moister conditions following the LGM returned to East Africa ca.16 kyr BP, and it contains a perfectly timed Younger Dryas episode (12.8-11.5 kyr BP) of intermittent drought. We find that the Early- to Mid-Holocene period, which in African records is often described as uniformly wet, was in fact punctuated by three distinct, century

  8. Reprocessing VIIRS sensor data records from the early SNPP mission

    NASA Astrophysics Data System (ADS)

    Blonski, Slawomir; Cao, Changyong

    2016-10-01

    The Visible-Infrared Imaging Radiometer Suite (VIIRS) instrument onboard the Suomi National Polar-orbiting Partnership (SNPP) satellite began acquiring Earth observations in November 2011. VIIRS data from all spectral bands became available three months after launch when all infrared-band detectors were cooled down to operational temperature. Before that, VIIRS sensor data record (SDR) products were successfully generated for the visible and near infrared (VNIR) bands. Although VIIRS calibration has been significantly improved through the four years of the SNPP mission, SDR reprocessing for this early mission phase has yet to be performed. Despite a rapid decrease in the telescope throughput that occurred during the first few months on orbit, calibration coefficients for the VNIR bands were recently successfully generated using an automated procedure that is currently deployed in the operational SDR production system. The reanalyzed coefficients were derived from measurements collected during solar calibration events that occur on every SNPP orbit since the beginning of the mission. The new coefficients can be further used to reprocess the VIIRS SDR products. In this study, they are applied to reprocess VIIRS data acquired over pseudo-invariant calibration sites Libya 4 and Sudan 1 in Sahara between November 2011 and February 2012. Comparison of the reprocessed SDR products with the original ones demonstrates improvements in the VIIRS calibration provided by the reprocessing. Since SNPP is the first satellite in a series that will form the Joint Polar Satellite System (JPSS), calibration methods developed for the SNPP VIIRS will also apply to the future JPSS measurements.

  9. AMS fabric and structural record along a strain gradient in an extrusive salt diapir (Kuh-e-Namak, Dashti, Iran)

    NASA Astrophysics Data System (ADS)

    Zavada, Prokop; Schulmann, Karel; Lexa, Ondrej; Machek, Matej; Roxerova, Zuzana; Kusbach, Vladimir

    2016-04-01

    The AMS record and the halite fabrics on meso- and micro-scale were studied in detail on a well exposed salt extrusive body in Iran. In the Kuh-e-Namak (Dashti) mountain salt diapir, the deformation structures in colored salt are displayed along longitudinal profiles across the dome and two glaciers that extend from the NE and SW edge of the dome. The profiles from the dome to the frontal parts of the glaciers reveal a continuous strain gradient associated with transposition of the domal salt fabrics by axial fold cleavage development during flow of rock salt over the ridges in the channel. The extruded salt belongs to the Hormuz sequence of Neo-Proterozoic to Early Cambrian age. From central dome towards especially the northern namakier, structural record revealed zonation from; 1) gravitational collapse related recumbent isoclinal folds in the dome, 2) flat normal shears at the edge of the dome, 3) collapsed vertical layering into flat lying transpositional fabric at the toe of the dome, 4) penetrative fold cleavage transposition of earlier fabrics above the topographical ridge in the base of the flow, locally displaying strong transversal constrictional fabrics, 5) banded mylonites with isoclinal rootless folds in subhorizontally banded frontal and marginal domain of the glacier. The AMS fabric in the rock salt is generated primarily by hematite dispersed in the recrystallized halite. The AMS exhibits three main types of fabric symmetry from clustered all directions (K1,K2,K3, orthogonal fabric) to clustered K1 directions with girdle forming K2,K3 axes and clustered K3 directions with girdle of K1 and K2 directions. The AMS fabric clearly reflects the macroscopic fabric transpositions along the entire investigated strain gradient in the rock salt. Magnetic fabrics reveal continuous trends from bimodal to semi-girdle distribution of foliations in folded and cleavage present regions, to magnetic lineation clustering perpendicular to flow in completely refolded

  10. The Sanrafaelic remagnetization revisited: Magnetic properties and magnetofabrics of Cambrian-Ordovician carbonates of the Eastern Precordillera of San Juan, Argentina

    NASA Astrophysics Data System (ADS)

    Fazzito, Sabrina Y.; Rapalini, Augusto E.; Poiré, Daniel G.

    2017-11-01

    Systematic rock-magnetic and magnetofabric studies were carried out on samples from twenty-three palaeomagnetic sites distributed on the La Laja, Zonda, La Flecha, La Silla and San Juan Formations, which constitute a thick middle Cambrian to early Ordovician carbonate sequence exposed in the Eastern Precordillera of Argentina. Previous palaeomagnetic studies on these rocks showed that this succession is characterized by a recent full overprint in the lower levels and a post-tectonic Permian remagnetization associated to the widespread Sanrafaelic event in the upper part. Our investigation revealed that the fluctuations of the magnetic properties are stratigraphically (lithologically) controlled. Anisotropy of magnetic and anhysteretic susceptibility measurements defined consistent fabrics along the entire section that switch progressively from "inverse", at the bottom, to "normal", at the top, with "intermediate" fabrics occurring mainly at medium levels. Degree of dolomitization significantly affects many rock-magnetic parameters, but appears unrelated to the presence of the Permian remagnetization, which is determined to reside in magnetite despite the complex magnetic mineralogy shown by the studied carbonates. Hysteresis cycles of rocks affected by the Sanrafaelic remagnetization are governed by ferromagnetic fractions showing a clear difference respect to those not affected and characterized by the dominance of paramagnetic or diamagnetic signals. The magnetic fabrics and mineralogical characterization rule out a thermoviscous origin and suggest a chemical remagnetization originated in the authigenic formation of magnetite for the Sanrafaelic overprint. X-ray diffraction analyses indicate that clay minerals are virtually absent in the whole succession with no traces of illite in any sample, discarding burial diagenesis of clay minerals for the origin of the remagnetization. Lack of late Palaeozoic magmatic rocks near the study area difficults correlation of

  11. Record-Breaking Early Flowering in the Eastern United States

    PubMed Central

    Ellwood, Elizabeth R.; Temple, Stanley A.; Primack, Richard B.; Davis, Charles C.

    2013-01-01

    Flowering times are well-documented indicators of the ecological effects of climate change and are linked to numerous ecosystem processes and trophic interactions. Dozens of studies have shown that flowering times for many spring-flowering plants have become earlier as a result of recent climate change, but it is uncertain if flowering times will continue to advance as temperatures rise. Here, we used long-term flowering records initiated by Henry David Thoreau in 1852 and Aldo Leopold in 1935 to investigate this question. Our analyses demonstrate that record-breaking spring temperatures in 2010 and 2012 in Massachusetts, USA, and 2012 in Wisconsin, USA, resulted in the earliest flowering times in recorded history for dozens of spring-flowering plants of the eastern United States. These dramatic advances in spring flowering were successfully predicted by historical relationships between flowering and spring temperature spanning up to 161 years of ecological change. These results demonstrate that numerous temperate plant species have yet to show obvious signs of physiological constraints on phenological advancement in the face of climate change. PMID:23342001

  12. Accessory Mineral Records of Early Earth Crust-Mantle Systematics: an Example From West Greenland

    NASA Astrophysics Data System (ADS)

    Storey, C. D.; Hawkesworth, C. J.

    2008-12-01

    Conditions for the formation and the nature of Earth's early crust are enigmatic due to poor preservation. Before c.4 Ga the only archives are detrital minerals eroded from earlier crust, such as the Jack Hills zircons in western Australia, or extinct isotope systematics. Zircons are particularly powerful since they retain precise records of their ages of crystallisation, and the Lu-Hf radiogenic isotope and O stable isotope systematics of the reservoir from which they crystallised. In principle, this allows insight into the nature of the crust, the mantle reservoir from which the melt was extracted and any reworked material incorporated into that melt. We have used in situ methods to measure U-Pb, O and Lu-Hf within single zircon crystals from tonalitic gneisses from West Greenland in the vicinity of the Isua Supracrustal Belt. They have little disturbed ages of c.3.8 Ga, mantle-like O isotope signatures and Lu-Hf isotope signatures that lie on the CHUR evolution line at 3.8 Ga. These samples have previously been subjected to Pb isotope feldspar and 142Nd whole rock analysis and have helped constrain models in which early differentiation of a proto-crust must have occurred. The CHUR-like Lu-Hf signature, along with mantle-like O signature from these zircons suggests juvenile melt production at 3.8 Ga from undifferentiated mantle, yet the other isotope systems preclude this possibility. Alternatively, this is further strong evidence for a heterogeneous mantle in the early Earth. Whilst zircons afford insight into the nature of the early crust and mantle, it is through the Sm-Nd system that the mantle has traditionally been viewed. Titanite often contains several thousand ppm Nd, making it amenable to precise analysis, and is a common accessory phase. It has a reasonably high closure temperature for Pb and O, and it can retain cores with older ages and distinct REE chemistry. It is often the main accessory phase alongside zircon, and it is the main carrier of Nd

  13. Bedrock Geology and Asbestos Deposits of the Upper Missisquoi Valley and Vicinity, Vermont

    USGS Publications Warehouse

    Cady, Wallace Martin; Albee, Arden Leroy; Chidester, A.H.

    1963-01-01

    The upper Missisquoi Valley and vicinity as described in this report covers an area of about 250 square miles at the headwaters of the Missisquoi River in north-central Vermont. About 90 percent of the area is forested and the remainder is chiefly farm land. The topography reflects the geologic structure and varied resistance of the bedrock to erosion. Most of the area is on the east limb of the Green Mountain anticlinorium, which is the principal structural feature of Vermont. The bedrock is predominantly sedimentary and volcanic rock that has been regionally metamorphosed. It was intruded before metamorphism by mafic and ultramafic igneous rocks, and after metamorphism by felsic and mafic igneous rocks. The metamorphosed sedimentary and volcanic rocks range in age from Cambrian(?) to Middle Silurian, the intrusive igneous rocks from probably Late Ordovician to probably late Permian. Metamorphism and principal folding in the region occurred in Middle Devonian time. The metamorphosed sedimentary and volcanic rocks make up a section at least 25,000 feet thick and can be divided into nine formations. The Hazens Notch formation of Cambrian(?) and Early Cambrian age is characterized by carbonaceous schist. It is succeeded in western parts of the area by the Jay Peak formation of Early Cambrian age, which is chiefly a schist that is distinguished by the general absence of carbonaceous zones; in central parts of the area the Hazens Notch formation is followed by the Belvidere Mountain amphibolite, probably the youngest of the formations of Early Cambrian age. The Ottauquechee formation, composed of carbonaceous phyllite and quartzite, and phyllitic graywacke, is of Middle Cambrian age. The Stowe formation of Late Cambrian(?) and Early(?) Ordovician age overlies the Ottauquechee and is predominantly noncarbonaceous schist, though it also contains greenstone and carbonaceous schist and phyllite. The Umbrella Hill formation of Middle Ordovician age is characteristically a

  14. Comparison of Passive Microwave-Derived Early Melt Onset Records on Arctic Sea Ice

    NASA Technical Reports Server (NTRS)

    Bliss, Angela C.; Miller, Jeffrey A.; Meier, Walter N.

    2017-01-01

    Two long records of melt onset (MO) on Arctic sea ice from passive microwave brightness temperatures (Tbs) obtained by a series of satellite-borne instruments are compared. The Passive Microwave (PMW) method and Advanced Horizontal Range Algorithm (AHRA) detect the increase in emissivity that occurs when liquid water develops around snow grains at the onset of early melting on sea ice. The timing of MO on Arctic sea ice influences the amount of solar radiation absorbed by the ice-ocean system throughout the melt season by reducing surface albedos in the early spring. This work presents a thorough comparison of these two methods for the time series of MO dates from 1979through 2012. The methods are first compared using the published data as a baseline comparison of the publically available data products. A second comparison is performed on adjusted MO dates we produced to remove known differences in inter-sensor calibration of Tbs and masking techniques used to develop the original MO date products. These adjustments result in a more consistent set of input Tbs for the algorithms. Tests of significance indicate that the trends in the time series of annual mean MO dates for the PMW and AHRA are statistically different for the majority of the Arctic Ocean including the Laptev, E. Siberian, Chukchi, Beaufort, and central Arctic regions with mean differences as large as 38.3 days in the Barents Sea. Trend agreement improves for our more consistent MO dates for nearly all regions. Mean differences remain large, primarily due to differing sensitivity of in-algorithm thresholds and larger uncertainties in thin-ice regions.

  15. Effects of maltreatment and parental schizophrenia spectrum disorders on early childhood social-emotional functioning: a population record linkage study.

    PubMed

    Matheson, S L; Kariuki, M; Green, M J; Dean, K; Harris, F; Tzoumakis, S; Tarren-Sweeney, M; Brinkman, S; Chilvers, M; Sprague, T; Carr, V J; Laurens, K R

    2017-12-01

    Childhood maltreatment and a family history of a schizophrenia spectrum disorder (SSD) are each associated with social-emotional dysfunction in childhood. Both are also strong risk factors for adult SSDs, and social-emotional dysfunction in childhood may be an antecedent of these disorders. We used data from a large Australian population cohort to determine the independent and moderating effects of maltreatment and parental SSDs on early childhood social-emotional functioning. The New South Wales Child Development Study combines intergenerational multi-agency data using record linkage methods. Multiple measures of social-emotional functioning (social competency, prosocial/helping behaviour, anxious/fearful behaviour; aggressive behaviour, and hyperactivity/inattention) on 69 116 kindergarten children (age ~5 years) were linked with government records of child maltreatment and parental presentations to health services for SSD. Multivariable analyses investigated the association between maltreatment and social-emotional functioning, adjusting for demographic variables and parental SSD history, in the population sample and in sub-cohorts exposed and not exposed to parental SSD history. We also examined the association of parental SSD history and social-emotional functioning, adjusting for demographic variables and maltreatment. Medium-sized associations were identified between maltreatment and poor social competency, aggressive behaviour and hyperactivity/inattention; small associations were revealed between maltreatment and poor prosocial/helping and anxious/fearful behaviours. These associations did not differ greatly when adjusted for parental SSD, and were greater in magnitude among children with no history of parental SSD. Small associations between parental SSD and poor social-emotional functioning remained after adjusting for demographic variables and maltreatment. Childhood maltreatment and history of parental SSD are associated independently with poor early

  16. Changes in chloride concentration in water from municipal wells that tap aquifers in rocks of Cambrian and Ordovician age in northeastern Illinois, 1915-84

    USGS Publications Warehouse

    Balding, G.O.

    1991-01-01

    During the past few decades, several municipalities in northeastern Illinois have noted increases in the salinity of water from wells that tap aquifers in rocks of Cambrian and Ordovician age. The municipalities have discontinued the use of, or sealed-off sections of, those wells. The aquifers involved include the Ancell, the Ironton-Galesville, and the Elmhurst-Mt. Simon. To define the location, magnitude, and possible causes for the salinity increases in the six northeastern counties of Illinois, 17 municipal wells and 1 deep test well were selected on the basis of their proximity to major pumping centers, the availability of water-quality data, and their documented maintenance history. Well depths ranged from about 960 to 3,475 feet. One well was finished in the middle confining unit, 2 wells were finished in the Ironton-Galesville aquifer, 4 wells were finished in the Eau Claire confining unit, and 10 wells were finished in the Elmhurst-Mt. Simon aquifer. The deep test well was finished below the Elmhurst-Mt. Simon aquifer in Precambrian-age rock. Chloride concentrations in the municipal wells ranged from less than 5 to greater than 600 milligrams per liter; in the deep test well, they ranged from 13 t o 37,000 milligrams per liter. Some changes in the chloride concentration in water from the studied municipal wells can be related to physical changes to the wells, including the partial filling in of a well, bridging within a well, the cleaning out of a well, or the deepening of a well. Some changes in chloride concentration are not related to physical changes but may be caused by increased pumpage; changes in pumping rate, frequency, or duration; cessation of pumping; improper abandonment of wells; and the upconing of highly mineralized water. The data base was inadequate for a quantitative study of the changes in chloride concentration in water from individual aquifers in rocks of Cambrian and Ordovician age.

  17. The Continuing Debate on Deep Molluscan Phylogeny: Evidence for Serialia (Mollusca, Monoplacophora + Polyplacophora)

    PubMed Central

    Stöger, I.; Sigwart, J. D.; Kano, Y.; Knebelsberger, T.; Marshall, B. A.; Schwabe, E.; Schrödl, M.

    2013-01-01

    Molluscs are a diverse animal phylum with a formidable fossil record. Although there is little doubt about the monophyly of the eight extant classes, relationships between these groups are controversial. We analysed a comprehensive multilocus molecular data set for molluscs, the first to include multiple species from all classes, including five monoplacophorans in both extant families. Our analyses of five markers resolve two major clades: the first includes gastropods and bivalves sister to Serialia (monoplacophorans and chitons), and the second comprises scaphopods sister to aplacophorans and cephalopods. Traditional groupings such as Testaria, Aculifera, and Conchifera are rejected by our data with significant Approximately Unbiased (AU) test values. A new molecular clock indicates that molluscs had a terminal Precambrian origin with rapid divergence of all eight extant classes in the Cambrian. The recovery of Serialia as a derived, Late Cambrian clade is potentially in line with the stratigraphic chronology of morphologically heterogeneous early mollusc fossils. Serialia is in conflict with traditional molluscan classifications and recent phylogenomic data. Yet our hypothesis, as others from molecular data, implies frequent molluscan shell and body transformations by heterochronic shifts in development and multiple convergent adaptations, leading to the variable shells and body plans in extant lineages. PMID:24350268

  18. New evidence on the anatomy and phylogeny of the earliest vertebrates.

    PubMed Central

    Xian-guang, Hou; Aldridge, Richard J; Siveter, David J; Siveter, Derek J; Xiang-hong, Feng

    2002-01-01

    We report the discovery of a new agnathan specimen from the Lower Cambrian Chengjiang Lagerstätte of China and thereby provide new evidence on the myomeres (V-shaped), the branchial apparatus (gill filaments and arches), the dorsal fin and the gonads (24-26) of the earliest vertebrates. The new specimen and the co-occurring Myllokunmingia fengjiaoa and Haikouichthys ercaicunensis represent a single species, which is a primitive member of the crown group craniates (vertebrates) and post-dates the origin of the myxinoids (hagfish). The origin of the vertebrate clade is at least as old as Early Cambrian. PMID:12350247

  19. Using the magmatic record to constrain the growth of continental crust-The Eoarchean zircon Hf record of Greenland

    NASA Astrophysics Data System (ADS)

    Fisher, Christopher M.; Vervoort, Jeffrey D.

    2018-04-01

    Southern West Greenland contains some of the best-studied and best-preserved magmatic Eoarchean rocks on Earth, and these provide an excellent vantage point from which to view long-standing questions regarding the growth of the earliest continental crust. In order to address the questions surrounding early crustal growth and complementary mantle depletion, we present Laser Ablation Split Stream (LASS) analyses of the U-Pb and Hf isotope compositions of zircon from eleven samples of the least-altered meta-igneous rocks from the Itsaq (Amîtsoq) Gneisses of the Isukasia and Nuuk regions of southern West Greenland. This analytical technique allows a less ambiguous approach to determining the age and Hf isotope composition of complicated zircon. Results corroborate previous findings that Eoarchean zircon from the Itsaq Gneiss (∼3.85 Ga to ∼3.63 Ga) were derived from a broadly chondritic source. In contrast to the Sm-Nd whole rock isotope record for southern West Greenland, the zircon Lu-Hf isotope record provides no evidence for early mantle depletion, nor does it suggest the presence of crust older than ∼3.85 Ga in Greenland. Utilizing LASS U-Pb and Hf data from the Greenland zircons studied here, we demonstrate the importance of focusing on the magmatic (rather than detrital) zircon record to more confidently understand early crustal growth and mantle depletion. We compare the Greenland Hf isotope data with other Eoarchean magmatic complexes such as the Acasta Gneiss Complex, Nuvvuagittuq greenstone belt, and the gneissic complexes of southern Africa, and all lack zircons with suprachondritic Hf isotope compositions. In total, these data suggest only a very modest volume of crust was produced during (or survived from) the Hadean and earliest Eoarchean. There remains no record of planet-scale early Earth mantle depletion in the Hf isotope record prior to 3.8 Ga.

  20. Orbital time scale records of Asian eolian dust from the Sea of Japan since the early Pliocene

    NASA Astrophysics Data System (ADS)

    Zhang, Wenfang; De Vleeschouwer, David; Shen, Ji; Zhang, Zeke; Zeng, Lin

    2018-05-01

    A high-resolution potassium content record of sediments from Integrated Ocean Drilling Program (IODP) Site U1422 drilled in the northern Sea of Japan was employed to yield an astronomical timescale for this ∼205 m thick sedimentary archive. The K content was quantified using the natural gamma radiation (NGR) data routinely measured on DV JOIDES Resolution. The U1422 K (wt.%) series shows an increasing trend with time, which parallels the growth of North Hemisphere ice sheets since ∼4 Ma, as revealed by the global benthic foraminifer's oxygen isotope stack (LR04). We propose that K content variations reflect changes in the relative contributions of Asian eolian dust and volcanic weathering products, in response to changes in global ice volume. Using the shipboard age model, constrained by palaeomagnetism and nanofossils datums, we tuned the U1422 K content variations to the LR04 stack. Our tuned age model for site U1422 goes back to 3.9 Ma. The studied record indicates that Asian dust generation is enhanced during periods of global cooling. The synchronous variations between the U1422 K record and the LR04 stack also suggest that global cooling played a dominant role in promoting the central Asian aridification since ∼3.9 Ma, while the uplift of Tibetan Plateau plays a secondary role, considering many disagreements still exists between the timing and amplitude of the Qinghai-Tibetan uplift and the evolution of central Asian aridity since the early Pliocene.

  1. Early Evaluation of the VIIRS Calibration, Cloud Mask and Surface Reflectance Earth Data Records

    NASA Technical Reports Server (NTRS)

    Vermote, Eric; Justice, Chris; Csiszar, Ivan

    2014-01-01

    Surface reflectance is one of the key products fromVIIRS and as withMODIS, is used in developing several higherorder land products. The VIIRS Surface Reflectance (SR) Intermediate Product (IP) is based on the heritageMODIS Collection 5 product (Vermote, El Saleous, & Justice, 2002). The quality and character of surface reflectance depend on the accuracy of the VIIRS Cloud Mask (VCM), the aerosol algorithms and the adequate calibration of the sensor. The focus of this paper is the early evaluation of the VIIRS SR product in the context of the maturity of the operational processing system, the Interface Data Processing System (IDPS). After a brief introduction, the paper presents the calibration performance and the role of the surface reflectance in calibration monitoring. The analysis of the performance of the cloud mask with a focus on vegetation monitoring (no snow conditions) shows typical problems over bright surfaces and high elevation sites. Also discussed is the performance of the aerosol input used in the atmospheric correction and in particular the artifacts generated by the use of the Navy Aerosol Analysis and Prediction System. Early quantitative results of the performance of the SR product over the AERONET sites showthatwith the fewadjustments recommended, the accuracy iswithin the threshold specifications. The analysis of the adequacy of the SR product (Land PEATE adjusted version) in applications of societal benefits is then presented. We conclude with a set of recommendations to ensure consistency and continuity of the JPSS mission with the MODIS Land Climate Data Record.

  2. Planetary biology and microbial ecology. Biochemistry of carbon and early life

    NASA Technical Reports Server (NTRS)

    Margulis, L. (Editor); Nealson, K. H. (Editor); Taylor, I. (Editor)

    1983-01-01

    Experiments made with cyanobacteria, phototrophic bacteria, and methanogenic bacteria are detailed. Significant carbon isotope fractionation data is included. Taken from well documented extant microbial communities, this data provides a basis of comparison for isotope fractionation values measured in Archean and Proterozoic (preCambrian) rocks. Media, methods, and techniques used to acquire data are also described.

  3. Phanerozoic polyphase orogenies recorded in the northeastern Okcheon Belt, Korea from SHRIMP U-Pb detrital zircon and K-Ar illite geochronologies

    NASA Astrophysics Data System (ADS)

    Jang, Yirang; Kwon, Sanghoon; Song, Yungoo; Kim, Sung Won; Kwon, Yi Kyun; Yi, Keewook

    2018-05-01

    We present the SHRIMP U-Pb detrital zircon and K-Ar illite 1Md/1M and 2M1 ages, suggesting new insight into the Phanerozoic polyphase orogenies preserved in the northeastern Okcheon Belt, Korea since the initial basin formation during Neoproterozoic rifting through several successive contractional orogens. The U-Pb detrital zircon ages from the Early Paleozoic strata of the Taebaeksan Zone suggest a Cambrian maximum deposition age, and are supported by trilobite and conodont biostratigraphy. Although the age spectra from two sedimentary groups, the Yeongwol and Taebaek Groups, show similar continuous distributions from the Late Paleoproterozoic to Early Paleozoic ages, a Grenville-age hiatus (1.3-0.9 Ga) in the continuous stratigraphic sequence from the Taebaek Group suggests the existence of different peripheral clastic sources along rifted continental margin(s). In addition, we present the K-Ar illite 1Md/1M ages of the fault gouges, which confirm fault formation/reactivation during the Late Cretaceous to Early Paleogene (ca. 82-62 Ma) and the Early Miocene (ca. 20-18 Ma). The 2M1 illite ages, at least those younger than the host rock ages, provide episodes of deformation, metamorphism and hydrothermal effects related to the tectonic events during the Devonian (ca.410 Ma) and Permo-Triassic (ca. 285-240 Ma). These results indicate that the northeastern Okcheon Belt experienced polyphase orogenic events, namely the Okcheon (Middle Paleozoic), Songrim (Late Paleozoic to Early Mesozoic), Daebo (Middle Mesozoic) and Bulguksa (Late Mesozoic to Early Cenozoic) Orogenies, reflecting the Phanerozoic tectonic evolution of the Korean Peninsula along the East Asian continental margin.

  4. Climate change and carbon-cycling during the latest Cretaceous-Early Paleogene; a new 13.5 million year-long, orbital-resolution, stable isotope record from the South Atlantic

    NASA Astrophysics Data System (ADS)

    Barnet, J.; Littler, K.; Kroon, D.; Leng, M. J.; Westerhold, T.; Roehl, U.; Zachos, J. C.

    2017-12-01

    The "greenhouse" world of the latest Cretaceous-Early Paleogene ( 70-34 Ma) was characterised by multi-million year variability in climate and the carbon-cycle. Throughout this interval the pervasive imprint of orbital-cyclicity, particularly eccentricity and precession, is visible in elemental and stable isotope data obtained from multiple deep-sea sites. Periodic "hyperthermal" events, occurring largely in-step with these orbital cycles, have proved particularly enigmatic, and may be the closest, albeit imperfect, analogues for anthropogenic climate change. This project utilises CaCO3-rich marine sediments recovered from ODP Site 1262 at a paleo-depth of 3600 m on the Walvis Ridge, South Atlantic, of late Maastrichtian-mid Paleocene age ( 67-60 Ma). We have derived high-resolution (2.5-4 kyr) carbon and oxygen isotope data from the epifaunal benthic foraminifera species Nuttallides truempyi. Combining the new record with the existing Late Paleocene-Early Eocene record generated from the same site by Littler et al. (2014), yields a single-site reference curve detailing 13.5 million years of orbital cyclicity in paleoclimate and carbon cycle from the latest Cretaceous to near the peak warmth of the Early Paleogene greenhouse. Spectral analysis of this new combined dataset allows us to identify long (405-kyr) eccentricity, short (100-kyr) eccentricity, and precession (19-23-kyr) as the principle forcing mechanisms governing pacing of the background climate and carbon-cycle during this time period, with a comparatively weak obliquity (41-kyr) signal. Cross-spectral analysis suggests that changes in climate lead the carbon cycle throughout most of the record, emphasising the role of the release of temperature-sensitive carbon stores as a positive feedback to an initial warming induced by changes in orbital configuration. The expression of comparatively understudied Early Paleocene events, including the Dan-C2 Event, Latest Danian Event, and Danian/Selandian Transition

  5. The origin of increased salinity in the Cambrian-Vendian aquifer system on the Kopli Peninsula, northern Estonia

    NASA Astrophysics Data System (ADS)

    Karro, Enn; Marandi, Andres; Vaikmäe, Rein

    Monitoring of the confined Cambrian-Vendian aquifer system utilised for industrial water supply at Kopli Peninsula in Tallinn over 24 years reveals remarkable changes in chemical composition of groundwater. A relatively fast 1.5 to 3.0-fold increase in TDS and in concentrations of major ions in ed groundwater is the consequence of heavy pumping. The main sources of dissolved load in Cambrian-Vendian groundwater are the leaching of host rock and the other geochemical processes that occur in the saturated zone. Underlying crystalline basement, which comprises saline groundwater in its upper weathered and fissured portion, and which is hydraulically connected with the overlying Cambrian-Vendian aquifer system, is the second important source of ions. The fractured basement and its clayey weathering crust host the Ca-Cl type groundwater, which is characterised by high TDS values (2-20 g/L). Intensive water ion accelerates the exchange of groundwaters and increases the area of influence of pumping. Chemical and isotopic studies of groundwater indicate an increasing contribution of old brackish water from the crystalline basement and rule out the potential implication of an intrusion of seawater into aquifer. L'origine de la salinité croissante dans le système aquifère du Cambrien-Vendien dans la péninsule de Kopli, nord de l'Estonie Le suivi à long terme du système aquifère captif du Cambrien-Vendien utilisé pour l'approvisionnement d'eaux industrielles dans la Péninsule de Kopli, nord de l'Estonie, révèle de remarquables changements dans la composition chimique des eaux souterraines. Une augmentation de facteur 1.5 à 3 de la TDS et des concentrations en ions majeurs dans l'eau souterraine est la conséquence de pompages intensifs. Les sources principales des charges dissoutes dans les eaux de l'aquifère du Cambrien-Vendien sont le lessivage des roches et d'autres phénomènes géochimiques ayant lieu dans la zone saturée. Le soubassement rocheux cristallin

  6. Late Pliocene/Early Pleistocene environments inferred from the Lake El'gygytgyn pollen record

    NASA Astrophysics Data System (ADS)

    Andreev, Andrei; Wennrich, Volker; Tarasov, Pavel; Raschke (Morozova), Elena; Brigham-Grette, Julie; Nowaczyk, Norbert; Melles, Martin

    2014-05-01

    The Arctic is known to play a crucial role within the global climate system. The mid-Pliocene (3-3.5 Ma) is considered to be the most probable scenario of the future climate changes. However, reliable climate projections are hampered by the complexity of the underlying natural variability and feedback mechanisms. An important prerequisite for the validation and improvement of the future projections is a better understanding of the long-term environmental history of the Arctic. Unfortunately, formation of continuous paleoenvironmental records in the Arctic was widely restricted due to repeated glaciations. Continuous sequences that penetrate the entire Quaternary and further into the Pliocene are highly desired and would enable to validate the temperature rise during the mid-Pliocene that was proposed by former studies. Such a record has now become available from Lake El'gygytgyn (67º30'N, 172º05E') located in a meteorite impact crater in north-eastern Siberia. The impact nearly 3.6 Ma ago formed an 18 km wide hole in the ground that then filled with water. The retrieved lake sediments have trapped pollen from a several thousand square-kilometer source area providing reliable insights into regional and over-regional millennial-scale vegetation and climate changes of the Arctic since the Pliocene. The ''El'gygytgyn Drilling Project" of ICDP has completed three holes in the center of the lake, penetrating about 318 m thick lake sediments and about 200 m of the impact rocks below. Because of its unusual origin and high-latitude setting in western Beringia, scientific drilling at Lake El'gygytgyn offered unique opportunities for paleoclimate research, allowing time-continuous climatic and environmental reconstructions back into the Pliocene. Late Pliocene and Early Pleistocene pollen assemblages can be subdivided into 55 pollen zones, which reflect the main environmental fluctuations in the region 3.55-2.15 Ma BP. Pollen-based climate reconstructions show that

  7. Paleoecology and paleoceanography of the Athel silicilyte, Ediacaran-Cambrian boundary, Sultanate of Oman.

    PubMed

    Stolper, D A; Love, G D; Bates, S; Lyons, T W; Young, E; Sessions, A L; Grotzinger, J P

    2017-05-01

    The Athel silicilyte is an enigmatic, hundreds of meters thick, finely laminated quartz deposit, in which silica precipitated in deep water (>~100-200 m) at the Ediacaran-Cambrian boundary in the South Oman Salt Basin. In contrast, Meso-Neoproterozoic sinks for marine silica were dominantly restricted to peritidal settings. The silicilyte is known to contain sterane biomarkers for demosponges, which today are benthic, obligately aerobic organisms. However, the basin has previously been described as permanently sulfidic and time-equivalent shallow-water carbonate platform and evaporitic facies lack silica. The Athel silicilyte thus represents a unique and poorly understood depositional system with implications for late Ediacaran marine chemistry and paleoecology. To address these issues, we made petrographic observations, analyzed biomarkers in the solvent-extractable bitumen, and measured whole-rock iron speciation and oxygen and silicon isotopes. These data indicate that the silicilyte is a distinct rock type both in its sedimentology and geochemistry and in the original biology present as compared to other facies from the same time period in Oman. The depositional environment of the silicilyte, as compared to the bounding shales, appears to have been more reducing at depth in sediments and possibly bottom waters with a significantly different biological community contributing to the preserved biomarkers. We propose a conceptual model for this system in which deeper, nutrient-rich waters mixed with surface seawater via episodic mixing, which stimulated primary production. The silica nucleated on this organic matter and then sank to the seafloor, forming the silicilyte in a sediment-starved system. We propose that the silicilyte may represent a type of environment that existed elsewhere during the Neoproterozoic. These environments may have represented an important locus for silica removal from the oceans. © 2017 John Wiley & Sons Ltd.

  8. Estimating the Roles of Genetic Risk, Perinatal Risk, and Marital Hostility on Early Childhood Adjustment: Medical Records and Self-Reports

    PubMed Central

    Neiderhiser, Jenae M.; Marceau, Kristine; De Araujo-Greecher, Marielena; Ganiban, Jody M.; Mayes, Linda C.; Shaw, Daniel S.; Reiss, David; Leve, Leslie D.

    2016-01-01

    A wide variety of perinatal risk factors have been linked to later developmental outcomes in children. Much of this work has relied on either birth/medical records or mothers’ self-reports collected after delivery, and there has been an ongoing debate about which strategy provides the most accurate and reliable data. This report uses a parent-offspring adoption design (N = 561 families) to (1) examine the correspondence between medical record data and self-report data, (2) examine how perinatal risk factors may influence child internalizing and externalizing behavior at age 4.5 years, and (3) explore interactions among genetic, perinatal risk, and rearing environment on child internalizing and externalizing behavior during early childhood. The agreement of self-reports and medical records data was relatively high (51–100%), although there was some variation based on the construct. There were few main effects of perinatal risk on child outcomes; however, there were several 2- and 3-way interactions suggesting that the combined influences of genetic, perinatal, and rearing environmental risks are important, particularly for predicting whether children exhibit internalizing versus externalizing symptoms at age 4.5 years. PMID:27075497

  9. Wisdom for Music Education from the Recording Studio

    ERIC Educational Resources Information Center

    Thibeault, Matthew D.

    2012-01-01

    Instead of focusing on the "stuff" of recording technology, this column explores ideas about the music that comes from the recording studio. Based on the author's experience working with Mark Rubel, four ideas about music are explored--one, the rise of alternatives to performance; two, the "get it done early" school of thought; three, recording as…

  10. Ages, geochemistry and tectonic implications of the Cambrian igneous rocks in the northern Great Xing'an Range, NE China

    NASA Astrophysics Data System (ADS)

    Feng, Zhiqiang; Liu, Yongjiang; Li, Yanrong; Li, Weimin; Wen, Quanbo; Liu, Binqiang; Zhou, Jianping; Zhao, Yingli

    2017-08-01

    The Xinlin-Xiguitu suture zone, located in the Great Xing'an Range, NE China, in the eastern segment of the Central Asian Orogenic Belt (CAOB), represents the boundary between the Erguna and Xing'an micro-continental blocks. The exact location of the Xinlin-Xiguitu suture zone has been debated, especially, the location of the northern extension of the suture zone. In this study, based on a detailed field, geochemical, geochronological and Sr-Nd-Hf isotope study, we focus our work on the Cambrian igneous rocks in the Erguna-Xing'an block. The Xinglong granitoids, mainly include ∼520 Ma diorite, ∼470 Ma monzogranite and ∼480 Ma pyroxene diorite. The granitoids show medium to high-K calc-alkaline series characteristics with post-collision granite affinity. The circa 500 Ma granitoids have low εHf (t) values (-16.6 to +2.2) and ancient two-stage model (TDM2) ages between 1317 Ma and 2528 Ma. These results indicate the primary magmas of the Xinglong granitoids were probably derived from the partial melting of a dominantly Paleo-Mesoproterozoic ;old; crustal source with possible different degrees of addition of juvenile materials, and formed in a post-collision tectonic setting after the amalgamation of the Erguna and Xing'an blocks. Compared with the Xinglong granitoids, the Duobaoshan igneous rocks are consisted of the approximately coeval rhyolitic tuffs (491 ± 5 Ma) and ultramafic intrusions (497 ± 5 Ma) within the Duobaoshan Formation. They are generally enriched in large ion lithophile elements (LILEs) and depleted in high field strength elements (HFSEs; e.g., Nb, Ta, and Ti), consistent with the geochemistry of igneous rocks from island arcs or active continental margins. The ultramafic rocks have high positive εHf (t) values (+1.3 to +15) and εNd (t) (+1.86 to +2.28), and relatively young two-stage model (TDM2) ages and low initial 87Sr/86Sr ratios (0.70628-0.70853), indicating the partial melting of a depleted mantle source from a subducted slab in

  11. Geology and lithogeochemistry of hydrothermal mudstones from the upper block near the Duck Pond volcanogenic massive sulfide (VMS) deposit, Newfoundland, Canada: evidence for low-temperature venting into oxygenated mid-Cambrian seawater

    NASA Astrophysics Data System (ADS)

    Piercey, Stephen J.; Squires, Gerry; Brace, Terry

    2018-02-01

    Pyrite- and pyrrhotite-rich mudstones are spatially associated with Cambrian ( 512-509 Ma) volcanogenic massive sulfide (VMS) deposits throughout the Tally Pond group, central Newfoundland, Canada. At the Duck Pond mine, sulfide-rich mudstones are hosted within a weakly mineralized upper block that structurally overlies the deposit but is older ( 513 versus 509 Ma). The mudstones are laminated, 10-30-cm thick, and pyrite- and pyrrhotite-rich and occur along pillow lava selvages, or in between pillow lavas, rhyolite flows, and volcaniclastic rocks. The mudstones are laterally extensive and proximal to the mudstone host rocks are hydrothermally altered to epidote-quartz-chlorite (basalt host) and sericite-quartz (rhyolite host). Lithogeochemical data for the sulfide-rich mudstones reflect the varying contributions of elements from sedimentary detritus, hydrothermal discharge, and hydrogenous scavenging from middle Cambrian seawater. The mudstones have minor detrital element abundances and significant hydrothermal element enrichments (i.e., elevated Fe2O3, S, Pb, Zn, Cu, and Ba concentrations, high Fe/Al ratios). The hydrothermal mudstones are also enriched in oxyanions (i.e., P2O5, U, V, Cr, Ni, Co, and Hg), interpreted to have been enriched via oxidative scavenging from seawater by Fe-oxide/oxyhydroxide particles. The mudstones also have REE-Y signatures similar to modern oxygenated seawater with high Y/Ho and negative Ce anomalies (Ce/Ce* = 0.40-0.86; average = 0.58), which correlate with adsorbed oxyanion concentrations. The low Eu/Eu* (1.02-1.86; average = 1.22) in the mudstones suggest that they were deposited from low-temperature (< 250 °C), Fe-rich hydrothermal fluids that likely formed a buoyant plume into an oxygenated water column. The REE-Y-oxyanion signatures suggest that the particles within the hydrothermal plume had sufficient residence time to scavenge oxyanions from seawater and inherit a middle Cambrian seawater signature. The predominant seawater

  12. Early Days of Recorder Teaching in South Australian Schools: A Personal History

    ERIC Educational Resources Information Center

    Southcott, Jane

    2016-01-01

    As a primary school student in the 1960s I learnt the recorder. This paper explores how the recorder became a staple of Australian primary school music programs. At that time recorders were comparatively recently revived Renaissance musical instruments that were adopted by music educators as a way for children and their teachers to engage in…

  13. Detrital zircon age distribution from Devonian and Carboniferous sandstone in the Southern Variscan Fold-and-Thrust belt (Montagne Noire, French Massif Central), and their bearings on the Variscan belt evolution

    NASA Astrophysics Data System (ADS)

    Lin, Wei; Faure, Michel; Li, Xian-hua; Chu, Yang; Ji, Wenbin; Xue, Zhenhua

    2016-05-01

    In the Southern French Massif Central, the Late Paleozoic sedimentary sequences of the Montagne Noire area provide clues to decipher the successive tectonic events that occurred during the evolution of the Variscan belt. Previous sedimentological studies already demonstrated that the siliciclastic deposits were supplied from the northern part of the Massif Central. In this study, detrital zircon provenance analysis has been investigated in Early Devonian (Lochkovian) conglomerate and sandstone, and in Carboniferous (Visean to Early Serpukhovian) sandstone from the recumbent folds and the foreland basin of the Variscan Southern Massif Central in Montagne Noire. The zircon grains from all of the samples yielded U-Pb age spectra ranging from Neoarchean to Late Paleozoic with several age population peaks at 2700 Ma, 2000 Ma, 980 Ma, 750 Ma, 620 Ma, 590 Ma, 560 Ma, 480 Ma, 450 Ma, and 350 Ma. The dominant age populations concentrate on the Neoproterozoic and Paleozoic. The dominant concordant detrital zircon age populations in the Lochkovian samples, the 480-445 Ma with a statistical peak around 450 Ma, are interpreted as reflecting the rifting event that separated several continental stripes, such as Armorica, Mid-German Crystalline Rise, and Avalonia from the northern part of Gondwana. However, Ediacaran and Cambrian secondary peaks are also observed. The detrital zircons with ages at 352 - 340 Ma, with a statistical peak around 350 Ma, came from the Early Carboniferous volcanic and plutonic rocks similar to those exposed in the NE part of the French Massif Central. Moreover, some Precambrian grains recorded a more complex itinerary and may have experienced a multi-recycling history: the Archean and Proterozoic grains have been firstly deposited in Cambrian or Ordovician terrigenous rocks, and secondly re-sedimented in Devonian and/or Carboniferous formations. Another possibility is that ancient grains would be inherited grains, scavenged from an underlying but not

  14. A Missing Link in the Evolution of the Cumulative Recorder

    ERIC Educational Resources Information Center

    Asano, Toshio; Lattal, Kennon A.

    2012-01-01

    A recently recovered cumulative recorder provides a missing link in the evolution of the cumulative recorder from a modified kymograph to a reliably operating, scientifically and commercially successful instrument. The recorder, the only physical evidence of such an early precommercial cumulative recorder yet found, was sent to Keio University in…

  15. Individual to Community-Level Faunal Responses to Environmental Change from a Marine Fossil Record of Early Miocene Global Warming

    PubMed Central

    Belanger, Christina L.

    2012-01-01

    Modern climate change has a strong potential to shift earth systems and biological communities into novel states that have no present-day analog, leaving ecologists with no observational basis to predict the likely biotic effects. Fossil records contain long time-series of past environmental changes outside the range of modern observation, which are vital for predicting future ecological responses, and are capable of (a) providing detailed information on rates of ecological change, (b) illuminating the environmental drivers of those changes, and (c) recording the effects of environmental change on individual physiological rates. Outcrops of Early Miocene Newport Member of the Astoria Formation (Oregon) provide one such time series. This record of benthic foraminiferal and molluscan community change from continental shelf depths spans a past interval environmental change (∼20.3-16.7 mya) during which the region warmed 2.1–4.5°C, surface productivity and benthic organic carbon flux increased, and benthic oxygenation decreased, perhaps driven by intensified upwelling as on the modern Oregon coast. The Newport Member record shows that (a) ecological responses to natural environmental change can be abrupt, (b) productivity can be the primary driver of faunal change during global warming, (c) molluscs had a threshold response to productivity change while foraminifera changed gradually, and (d) changes in bivalve body size and growth rates parallel changes in taxonomic composition at the community level, indicating that, either directly or indirectly through some other biological parameter, the physiological tolerances of species do influence community change. Ecological studies in modern and fossil records that consider multiple ecological levels, environmental parameters, and taxonomic groups can provide critical information for predicting future ecological change and evaluating species vulnerability. PMID:22558424

  16. Subdivision of the Cambrian-Ordovician Upper Sandstone Aquifer'' in the Fox Cities area, east-central Wisconsin

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Erikson, K.A.; Smith, G.L.

    The informally-named Upper Sandstone Aquifer,'' a lithologically heterogeneous unit, has been subdivided into three thinner hydrostratigraphic units, the Upper Cambrian Jordan Formation (sandstone), the Lower Ordovician Prairie du Chien Group (dolomite), and the Middle Ordovician St. Peter Formation (mostly sandstone). Reevaluation of the Upper Sandstone Aquifer'' in the Fox Cities area (Appleton, Neenah, Menasha) was based on detailed analysis of 125 geologic well logs and 200 well-construction reports. Isopach maps and geologic cross-sections of these units in the Fox Cities area show extreme variations in thickness over short lateral distances. Although the Jordan Formation has a uniform thickness of 13more » m (40 ft), the Prairie du Chien Group varies from 0--88 m (0--275 ft). The St. Peter Formation varies in thickness from 0--108 m (0--335 ft). The Prairie du Chien Group is relatively thin or absent in the Fox Cities area in locations where the overlying St. Peter Formation is relatively thick. St Peter can be further subdivided into Readstown, Tonti, and Glenwood Members. The Readstown Member is a lithologically heterogeneous unit of variable thickness and distribution. The Tonti Member is a poorly cemented, fine-medium quartz arenite with which is lumped the Glenwood Member, a conglomeratic, silty medium-coarse sandstone. Maximum St. Peter thicknesses represent filled paleokarst depressions developed on top of the Prairie du Chien Group, with profound implications for estimates of hydraulic conductivity.« less

  17. The importance of intracrustal magmatic heat advection in large hot orogens: the Ediacaran-Cambrian ultrahigh-temperature domain of southern Madagascar.

    NASA Astrophysics Data System (ADS)

    Holder, R. M.; Hacker, B. R.

    2016-12-01

    Ultrahigh temperature metamorphism (UHTM) represents the thermal extremes of crustal metamorphism and is integral to our understanding of orogenic systems and continental evolution. The relative importance of advective and radiogenic heating in UHTM is often unclear, however, due to retrogression of UHT mineral assemblages and lack of robust chronology. We report the first observation of osumilite, pseudosections, feldspar thermometry, and monazite LASS U-Th-Pb chronology from the Ediacaran-Cambrian Anosyen domain of southern Madagascar to evaluate P-T-time paths and assess the relative importance of advective and radiogenic heating. Pseudosections of an osumilite-bearing assemblage, a garnet-orthopyroxene gneiss, and a garnet-spinel leucogneiss indicate peak conditions of 930-1010 C, 4-6 kbar; 900-950 C, 6-7 kbar; and 970-1000 C, 4.5-5.5 kbar, respectively. Feldspar thermometry indicates T > 915 ± 30 C. These conditions were reached along a low-P prograde path, as implied by cordierite and sillimanite inclusions in garnet. Monazite and zircon U/Th-Pb dating indicates that prograde metamorphism began 625 Ma and lasted 60 Myr. The timing of peak metamorphism is constrained to a 16 ± 2.5 Myr period between 566 ± 2 and 550 ± 1.5 Ma, by monazite inclusions in pseudomorphs after osumilite and monazite from a leucosome in the same outcrop. Peak T was achieved immediately following emplacement of voluminous charnockites (emplaced 1000-1100 C) and related magmas of the Anosyen Batholith. Crystallization of pegmatite dikes, leucosomes, and smaller plutons during cooling lasted from 550 to 510 Ma. This broad thermal history is also recorded by metamorphic monazite Eu/Eu*, which indicate progressive melt generation and extraction prior to 550 Ma followed by melt accumulation and crystallization after 550 Ma. Although the duration of prograde metamorphism is consistent with radiogenic heating models of large hot orogens, peak T was 100-200 C higher in southern Madagascar

  18. Magmatic @d^1^8O in 4400-3900 Ma detrital zircons: A record of the alteration and recycling of crust in the Early Archean [rapid communication

    NASA Astrophysics Data System (ADS)

    Cavosie, A. J.; Valley, J. W.; Wilde, S. A.

    2005-07-01

    Ion microprobe analyses of δ 18O in 4400-3900 Ma igneous zircons from the Jack Hills, Western Australia, provide a record of the oxygen isotope composition of magmas in the earliest Archean. We have employed a detailed analysis protocol aimed at correlating spatially related micro-volumes of zircon concordant in U/Pb age with δ 18O and internal zoning. Simultaneous analysis of 18O and 16O with dual Faraday cup detectors, combined with frequent standardization, has yielded data with improved accuracy and precision over prior studies, and resulted in a narrower range of what is interpreted as magmatic δ 18O in > 3900 Ma zircons. Preserved magmatic δ 18O values from individual zircons (Zrc) range from 5.3‰ to 7.3‰ (VSMOW), and increasingly deviate from the mantle range of 5.3 ± 0.3‰ as zircons decrease in age from 4400 to 4200 Ma. Elevated δ 18O (Zrc) values up to 6.5‰ occur as early as 4325 Ma, which suggests that evolved rocks were incorporated into magmas within ˜230 Ma of Earth's accretion. Values of magmatic δ 18O (Zrc) as high as 7.3‰ are recorded in zircons by 4200 Ma, and are common thereafter. The protoliths of the magmas these zircons crystallized in were altered by low temperature interaction with liquid water near Earth's surface. These results provide the strongest evidence yet for the existence of liquid water oceans and supracrustal rocks by approximately 4200 Ma, and possibly as early as 4325 Ma. The range of magmatic δ 18O values in the 4400-3900 Ma zircons is indistinguishable from Archean igneous zircons, suggesting similar magmatic processes occurred over the first two billion years of recorded Earth history. Zircons with sub-solidus alteration histories, identified by the presence of disturbed internal zoning patterns, record δ 18O values both below (4.6‰) and above (10.3‰) the observed range for primary magmatic zircon, and are unreliable indicators of Early Archean magma chemistry.

  19. Health and safety needs in early care and education programs: what do directors, child health records, and national standards tell us?

    PubMed

    Alkon, Abbey; To, Kim; Mackie, Joanna F; Wolff, Mimi; Bernzweig, Jane

    2010-01-01

    To identify the overlapping and unique health and safety needs and concerns identified by early care and education (ECE) directors, health records, and observed compliance with national health and safety (NHS) standards. Cross-sectional study. 127 ECE programs from 5 California counties participated in the study, including 118 directors and 2,498 children's health records. Qualitative data were collected using standardized ECE directors' interviews to identify their health and safety concerns; and objective, quantitative data were collected using child health record reviews to assess regular health care, immunizations, health insurance, special health care needs, and screening tests and an observation Checklist of 66 key NHS standards collected by research assistants. The overlapping health and safety needs and concerns identified by the directors and through observations were hygiene and handwashing, sanitation and disinfection, supervision, and the safety of indoor and outdoor equipment. Some of the health and safety needs identified by only one assessment method were health and safety staff training, medical plans for children with special health care needs and follow-up on positive screening tests. Comprehensive, multimethod assessments are useful to identify health and safety needs and develop public health nursing interventions for ECE programs.

  20. Pellet microfossils: Possible evidence for metazoan life in Early Proterozoic time

    PubMed Central

    Robbins, Eleanora Iberall; Porter, Karen Glaus; Haberyan, Kurt A.

    1985-01-01

    Microfossils resembling fecal pellets occur in acid-resistant residues and thin sections of Middle Cambrian to Early Proterozoic shale. The cylindrical microfossils average 50 × 110 μm and are the size and shape of fecal pellets produced by microscopic animals today. Pellets occur in dark gray and black rocks that were deposited in the facies that also preserves sulfide minerals and that represent environments analogous to those that preserve fecal pellets today. Rocks containing pellets and algal microfossils range in age from 0.53 to 1.9 gigayears (Gyr) and include Burgess Shale, Greyson and Newland Formations, Rove Formation, and Gunflint Iron-Formation. Similar rock types of Archean age, ranging from 2.68 to 3.8 Gyr, were barren of pellets. If the Proterozoic microfossils are fossilized fecal pellets, they provide evidence of metazoan life and a complex food chain at 1.9 Gyr ago. This occurrence predates macroscopic metazoan body fossils in the Ediacaran System at 0.67 Gyr, animal trace fossils from 0.9 to 1.3 Gyr, and fossils of unicellular eukaryotic plankton at 1.4 Gyr. Images PMID:16593599

  1. Effect of recording duration on the diagnostic performance of multifocal visual-evoked potentials in high-risk ocular hypertension and early glaucoma.

    PubMed

    Fortune, Brad; Zhang, Xian; Hood, Donald C; Demirel, Shaban; Patterson, Emily; Jamil, Annisa; Mansberger, Steven L; Cioffi, George A; Johnson, Chris A

    2008-01-01

    To evaluate the effect on diagnostic performance of reducing multifocal visual-evoked potential (mfVEP) recording duration from 16 to 8 minutes per eye. Both eyes of 185 individuals with high-risk ocular hypertension or early glaucoma were studied. Two 8-minute mfVEP recordings were obtained for each eye in an ABBA order using VERIS. The first recording for each eye was compared against single run (1-Run) mfVEP normative data; the average of both recordings for each eye was compared against 2-Run normative data. Visual fields (VFs) were obtained by standard automated perimetry (SAP) within 22.3+/-27.0 days of the mfVEP. Stereo disc photographs and Heidelberg Retina Tomograph images were obtained together, within 24.8+/-50.4 days of the mfVEP and 33.1+/-62.9 days of SAP. Masked experts graded disc photographs as either glaucomatous optic neuropathy or normal. The overall Moorfields Regression Analysis result from the Heidelberg Retina Tomograph was used as a separate diagnostic classification. Thus, 4 diagnostic standards were applied in total, 2 based on optic disc structure alone and 2 others based on disc structure and SAP. Agreement between the 1-Run and 2-Run mfVEP was 90%. Diagnostic performance of the 1-Run mfVEP was similar to that of the 2-Run mfVEP for all 4 diagnostic standards. Sensitivity was slightly higher for the 2-Run mfVEP, whereas specificity was slightly higher for the 1-Run mfVEP. If higher sensitivity is sought, the 2-Run mfVEP will provide better discrimination between groups of eyes with relatively high signal-to-noise ratio (eg, early glaucoma or high-risk suspects). But if higher specificity is a more important goal, the 1-Run mfVEP provides adequate sensitivity and requires only half the test time. Considered alongside prior studies, the present results suggest that the 1-Run mfVEP is an efficient way to confirm (or refute) the extent of VF loss in patients with moderate or advanced glaucoma, particularly in those with unreliable VFs

  2. Effect of Recording Duration on the Diagnostic Performance of Multifocal Visual-evoked Potentials in High-risk Ocular Hypertension and Early Glaucoma

    PubMed Central

    Fortune, Brad; Zhang, Xian; Hood, Donald C.; Demirel, Shaban; Patterson, Emily; Jamil, Annisa; Mansberger, Steven L.; Cioffi, George A.; Johnson, Chris A.

    2010-01-01

    Purpose To evaluate the effect on diagnostic performance of reducing multifocal visual-evoked potential (mfVEP) recording duration from 16 to 8 minutes per eye. Methods Both eyes of 185 individuals with high-risk ocular hypertension or early glaucoma were studied. Two 8-minute mfVEP recordings were obtained for each eye in an ABBA order using VERIS. The first recording for each eye was compared against single run (1-Run) mfVEP normative data; the average of both recordings for each eye was compared against 2-Run normative data. Visual fields (VFs) were obtained by standard automated perimetry (SAP) within 22.3±27.0 days of the mfVEP. Stereo disc photographs and Heidelberg Retina Tomograph images were obtained together, within 24.8±50.4 days of the mfVEP and 33.1±62.9 days of SAP. Masked experts graded disc photographs as either glaucomatous optic neuropathy or normal. The overall Moorfields Regression Analysis result from the Heidelberg Retina Tomograph was used as a separate diagnostic classification. Thus, 4 diagnostic standards were applied in total, 2 based on optic disc structure alone and 2 others based on disc structure and SAP. Results Agreement between the 1-Run and 2-Run mfVEP was 90%. Diagnostic performance of the 1-Run mfVEP was similar to that of the 2-Run mfVEP for all 4 diagnostic standards. Sensitivity was slightly higher for the 2-Run mfVEP, whereas specificity was slightly higher for the 1-Run mfVEP. Conclusions If higher sensitivity is sought, the 2-Run mfVEP will provide better discrimination between groups of eyes with relatively high signal-to-noise ratio (eg, early glaucoma or high-risk suspects). But if higher specificity is a more important goal, the 1-Run mfVEP provides adequate sensitivity and requires only half the test time. Considered alongside prior studies, the present results suggest that the 1-Run mfVEP is an efficient way to confirm (or refute) the extent of VF loss in patients with moderate or advanced glaucoma, particularly

  3. Testing the apatite depletion hypothesis for early Holocene ecosystem acidification using the lake sediment record at Krâkenes, Norway.

    NASA Astrophysics Data System (ADS)

    Thrasher, I. M.; Boyle, J. F.; Chiverrell, R. C.; Plater, A. J.

    2009-04-01

    Lakes created by retreating ice at the end of the last glaciation underwent rapid acidification during the first few thousand years of their existence, a phenomenon that has been attributed in part to progressive leaching of soil bases since it was discovered more than 80 years ago. Though a role for leaching is still acknowledged, the most recent studies see this as subordinate to the effects of biological and climatic changes initiated by deglaciation, chiefly primary vegetation succession and species immigration. However, we propose a simpler alternative explanation, based on the geochemical modelling of runoff acidity. This shows that the extent and timing of early Holocene lake acidification in eight published palaeoecological records can be explained by leaching of the calcium phosphate mineral apatite from the granitic till soils of their catchments, at a rate controlled by simple dissolution kinetic factors. If confirmed, this hypothesis has important implications for our understanding of long-term lake ecosystem development. Not only does it mean that the mechanism is inherently irreversible, in contrast to the alternative ecological and climatic mechanisms which are not. Also, it reinforces the view that long-term ecosystem modelling cannot safely neglect nutrient limitation, as is currently the practice in widely used global dynamic vegetation models. Here we present a NERC-funded programme of research that uses the sediment mineral record of Kråkenes (western Norway), the best studied early Holocene lake sediment sequence in the world, to provide a simple, critical and unambiguous test of this hypothesis.

  4. Trait-based diversification shifts reflect differential extinction among fossil taxa

    PubMed Central

    Wagner, Peter J.; Estabrook, George F.

    2014-01-01

    Evolution provides many cases of apparent shifts in diversification associated with particular anatomical traits. Three general models connect these patterns to anatomical evolution: (i) elevated net extinction of taxa bearing particular traits, (ii) elevated net speciation of taxa bearing particular traits, and (iii) elevated evolvability expanding the range of anatomies available to some species. Trait-based diversification shifts predict elevated hierarchical stratigraphic compatibility (i.e., primitive→derived→highly derived sequences) among pairs of anatomical characters. The three specific models further predict (i) early loss of diversity for taxa retaining primitive conditions (elevated net extinction), (ii) increased diversification among later members of a clade (elevated net speciation), and (iii) increased disparity among later members in a clade (elevated evolvability). Analyses of 319 anatomical and stratigraphic datasets for fossil species and genera show that hierarchical stratigraphic compatibility exceeds the expectations of trait-independent diversification in the vast majority of cases, which was expected if trait-dependent diversification shifts are common. Excess hierarchical stratigraphic compatibility correlates with early loss of diversity for groups retaining primitive conditions rather than delayed bursts of diversity or disparity across entire clades. Cambrian clades (predominantly trilobites) alone fit null expectations well. However, it is not clear whether evolution was unusual among Cambrian taxa or only early trilobites. At least among post-Cambrian taxa, these results implicate models, such as competition and extinction selectivity/resistance, as major drivers of trait-based diversification shifts at the species and genus levels while contradicting the predictions of elevated net speciation and elevated evolvability models. PMID:25331898

  5. Trait-based diversification shifts reflect differential extinction among fossil taxa.

    PubMed

    Wagner, Peter J; Estabrook, George F

    2014-11-18

    Evolution provides many cases of apparent shifts in diversification associated with particular anatomical traits. Three general models connect these patterns to anatomical evolution: (i) elevated net extinction of taxa bearing particular traits, (ii) elevated net speciation of taxa bearing particular traits, and (iii) elevated evolvability expanding the range of anatomies available to some species. Trait-based diversification shifts predict elevated hierarchical stratigraphic compatibility (i.e., primitive→derived→highly derived sequences) among pairs of anatomical characters. The three specific models further predict (i) early loss of diversity for taxa retaining primitive conditions (elevated net extinction), (ii) increased diversification among later members of a clade (elevated net speciation), and (iii) increased disparity among later members in a clade (elevated evolvability). Analyses of 319 anatomical and stratigraphic datasets for fossil species and genera show that hierarchical stratigraphic compatibility exceeds the expectations of trait-independent diversification in the vast majority of cases, which was expected if trait-dependent diversification shifts are common. Excess hierarchical stratigraphic compatibility correlates with early loss of diversity for groups retaining primitive conditions rather than delayed bursts of diversity or disparity across entire clades. Cambrian clades (predominantly trilobites) alone fit null expectations well. However, it is not clear whether evolution was unusual among Cambrian taxa or only early trilobites. At least among post-Cambrian taxa, these results implicate models, such as competition and extinction selectivity/resistance, as major drivers of trait-based diversification shifts at the species and genus levels while contradicting the predictions of elevated net speciation and elevated evolvability models.

  6. Observation & Record Keeping in Early Childhood Programs. Australian Early Childhood Resource Booklets No. 1.

    ERIC Educational Resources Information Center

    Veale, Ann; Piscitelli, Barbara

    Observing children is a task teachers and caregivers perform to understand the unique characteristics of each child. Observation alone can be helpful, but in combination with record keeping it becomes a valuable aid to understanding child development and can be used as a basis for making decisions about appropriate experiences to foster each…

  7. Late Paleocene- Early Eocene paleoenvironments in the Southwest Pacific (ODP Leg 189): Revised Stratigraphy of an Antarctic PETM Record.

    NASA Astrophysics Data System (ADS)

    Bijl, P. K.; Brinkhuis, H.; Sluijs, A.; Reichart, G.; Schouten, S.; Röhl, U.

    2007-12-01

    The Late Paleocene and Early Eocene (~60 ¡V 50 Ma) were characterized by globally warm climates. Superimposed on this general warmth, several episodes of further warming occurred (so-called hyperthermals), including the Paleocene-Eocene thermal maximum (PETM, ~55 Ma) and Eocene thermal maximum 2 (ETM2, ~53 Ma). While the PETM is by now well documented from Northern Hemisphere, and some equatorial locations, southern high-latitude records are still rare. Here we present high-resolution palynological, XRF, bulk organic stable carbon isotope (Ô13CTOC), and TEX86 palaeothermometry data across Upper Paleocene through Lower Eocene pro-deltaic deposits from the Southwest Pacific Ocean, at ~65¢XS palaeolatitude (ODP Site 1172). Based on a revised integrated biomagnetostratigraphic age model and Ô13CTOC stratigraphy, we identify the southernmost marginal marine PETM ever encountered. Moreover, there is every indication that ETM2 was recovered as well at Site 1172. Despite a high latitude source of the surface waters at this site, the PETM is marked by the characteristic acme of representatives of the (sub) tropical dinoflagellate cyst (dinocyst) Apectodinium, confirming the truly global nature of this PETM event. TEX86 values indicate that surface ocean temperatures rose from ~23¢XC to ~30¢XC during the PETM at Site 1172, hence by a similar magnitude as recorded in other PETM successions globally. Before and after the CIE, mass abundances of low salinity tolerant dinocysts are recorded, taken as indicative of increased runoff. These trends are analogous to those recorded at northern high latitudes, indicating a similar climate response at both polar regions during the PETM. Yet, some distinct differences are apparent, and are discussed.

  8. The Colorado Plateau Coring Project: A Continuous Cored Non-Marine Record of Early Mesozoic Environmental and Biotic Change

    NASA Astrophysics Data System (ADS)

    Irmis, Randall; Olsen, Paul; Geissman, John; Gehrels, George; Kent, Dennis; Mundil, Roland; Rasmussen, Cornelia; Giesler, Dominique; Schaller, Morgan; Kürschner, Wolfram; Parker, William; Buhedma, Hesham

    2017-04-01

    The early Mesozoic is a critical time in earth history that saw the origin of modern ecosystems set against the back-drop of mass extinction and sudden climate events in a greenhouse world. Non-marine sedimentary strata in western North America preserve a rich archive of low latitude terrestrial ecosystem and environmental change during this time. Unfortunately, frequent lateral facies changes, discontinuous outcrops, and a lack of robust geochronologic constraints make lithostratigraphic and chronostratigraphic correlation difficult, and thus prevent full integration of these paleoenvironmental and paleontologic data into a regional and global context. The Colorado Plateau Coring Project (CPCP) seeks to remedy this situation by recovering a continuous cored record of early Mesozoic sedimentary rocks from the Colorado Plateau of the western United States. CPCP Phase 1 was initiated in 2013, with NSF- and ICDP-funded drilling of Triassic units in Petrified Forest National Park, northern Arizona, U.S.A. This phase recovered a 520 m core (1A) from the northern part of the park, and a 240 m core (2B) from the southern end of the park, comprising the entire Lower-Middle Triassic Moenkopi Formation, and most of the Upper Triassic Chinle Formation. Since the conclusion of drilling, the cores have been CT scanned at the University of Texas - Austin, and split, imaged, and scanned (e.g., XRF, gamma, and magnetic susceptibility) at the University of Minnesota LacCore facility. Subsequently, at the Rutgers University Core Repository, core 1A was comprehensively sampled for paleomagnetism, zircon geochronology, petrography, palynology, and soil carbonate stable isotopes. LA-ICPMS U-Pb zircon analyses are largely complete, and CA-TIMS U-Pb zircon, paleomagnetic, petrographic, and stable isotope analyses are on-going. Initial results reveal numerous horizons with a high proportion of Late Triassic-aged primary volcanic zircons, the age of which appears to be a close

  9. Comparisons of Lower Cambrian ( ) Chilhowee Group siliciclastics of the frontal Blue Ridge with possible equivalents in the central Blue Ridge and basement massifs in the southern Appalachians

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Aylor, J.G. Jr.; Tull, J.F.

    1993-03-01

    The Lower Cambrian Chilhowee Group (CG) along the Blue Ridge (BR) foreland boundary may also be represented in more internal parts of the orogen. The relative palinspastic positions of these more internal CG( ) sequences are poorly constrained, but they are believed to represent more outboard facies than those along the frontal BR. Although correlation with CG rocks of the frontal BR is indefinite due to metamorphism and polydeformation, correlative sequences may include the cover of the Corbin and Salem Church gneisses (Pinelog Fm.), Grandfather Mountain window, Murphy belt (MB) (Hiwassee River Group), Tallulah and Toxaway domes (TTD) (quartzite-schist membermore » of the Tallulah Falls Fm.), Pine Mountain window (Hollis Quartzite), and Sauratown Mountains window (Hogan Creek and Sauratown Fms.). The CG is generally bounded on the west by the Great Smoky fault. Siliciclastics of the CG represent stacked, coarsening upward sequences, separated by transgressive facies, and capped by a highstand of the Shady Dolomite or its equivalents. CG correlations in the Kahatchee Mountain Group of the Talladega belt and the siliciclastics in the Hiwassee River Group of the MB are supported by fossil constraints. Other units are correlated with CG based upon intimate associations with marble believed to be equivalent to the overlying Lower Cambrian Shady Dolomite, or upon presumed uncomformable relationships above Grenville basement. The CG averages about 1,260 meters thickness at the frontal BR, whereas lower siliciclastics in the MB average 1,830 meters. The Pinelog Fm. is up to 600 meters thick. The Hollis Quartzite is approximately 325 meters thick, and the estimated thickness of the quartz-schist member at Tallulah Falls is up to 900 meters. More distal siliciclastics of the central BR in the MB and distal siliciclastics overlying basement were deposited farther out on the shelf as stratigraphic, litho-facies equivalents of shallower marine and continental deposits of the

  10. Accelerating Neoproterozoic Research through Scientific Drilling

    NASA Astrophysics Data System (ADS)

    Condon, Daniel; Prave, Anthony; Boggiani, Paulo; Fike, David; Halverson, Galen; Kasemann, Simone; Knoll, Andrew; Zhu, Maoyan

    2014-05-01

    The Neoproterozoic Era (1.0 to 0.541 Ga) and earliest Cambrian (541 to ca. 520 Ma) records geologic changes unlike any other in Earth history: supercontinental tectonics of Rodinia followed by its breakup and dispersal into fragments that form the core of today's continents; a rise in oxygen that, perhaps for the first time in Earth history, resulted in the deep oceans becoming oxic; snowball Earth, which envisages a blanketing of global ice cover for millions of years; and, at the zenith of these combined biogeochemical changes, the evolutionary leap from eukaryotes to animals. Such a concentration of hallmark events in the evolution of our planet is unparalleled and many questions regarding Earth System evolution during times of profound climatic and geological changes remain to be answered. Neoproterozoic successions also offer insight into the genesis of a number of natural resources. These include banded-iron formation, organic-rich shale intervals (with demonstrated hydrocarbon source rocks already economically viable in some countries), base and precious metal ore deposits and REE occurrences, as well as industrial minerals and dimension stone. Developing our understanding of the Neoproterozoic Earth-system, combined with regional geology has the potential to impact the viability of these resources. Our understanding of the Neoproterozoic and early Cambrian, though, is overwhelmingly dependent on outcrop-based studies, which suffer from lack of continuity of outcrop and, in many instances, deep weathering profiles. A limited number of research projects study Precambrian strata have demonstrated the potential impact of scientific drilling to augment and complement ongoing outcrop based studies and advancing research. An ICDP and ECORD sponsored workshop, to be held in March 2014, has been convened to discuss the utility of scientific drilling for accelerating research of the Neoproterozoic through early Cambrian (ca. 0.9 to 0.52 Ga) rock record. The aim is to

  11. The emergence of modern type rain forests and mangroves and their traces in the palaeobotanical record during the Late Cretaceous and early Tertiary

    NASA Astrophysics Data System (ADS)

    Mohr, Barbara; Coiffard, Clément

    2014-05-01

    The origin of modern rain forests is still very poorly known. This ecosystem could have potentially fully evolved only after the development of relatively high numbers of flowering plant families adapted to rain forest conditions. During the early phase of angiosperm evolution in the early Cretaceous the palaeo-equatorial region was located in a seasonally dry climatic belt, so that during this phase, flowering plants often show adaptations to drought, rather than to continuously wet climate conditions. Therefore it is not surprising that except for the Nymphaeales, the most basal members of extant angiosperm families have members that do not necessarily occur in the continuously wet tropics today. However, during the late Early Cretaceous several clades emerged that later would give rise to families that are typically found today mostly in (shady) moist places in warmer regions. This is especially seen among the monocotyledons, a group of the mesangiosperms, that developed in many cases large leaves often with very specific venation patterns that make these leaves very unique and well recognizable. Especially members of three groups are here of interest: the arum family (Araceae), the palms (Arecaceae) and the Ginger and allies (Zingiberales). The earliest fossil of Araceae are restricted to low latitudes during the lower Cretaceous. Arecaceae and Zingiberales do not appear in the fossil record before the early late Cretaceous and occur at mid latitudes. During the Late Cretaceous, Araceae are represented at mid latitudes by non-tropical early diverging members and at low latitudes by derived rainforest members. Palms became widespread during the Late Cretataceous and also Nypa, a typical element of tropical to subtropical mangrove environments evolved during this time period. During the Paleocene Arecaceae appear to be restricted to lower latitudes as well as Zingiberales. All three groups are again widespread during the Eocene, reaching higher latitudes and

  12. Uniqueness Of docosahexaenoic acid: A master Of DNA and A Quantum gate

    USDA-ARS?s Scientific Manuscript database

    The fossil record displays the sudden appearance of intracellular detail and the 32 phyla in what is known as the “Cambrian Explosion” at about 600 million years ago. The intracellular structures were made with membrane lipids which provided for organisation and specialisation. Oxidative metabolism...

  13. Olivia Records: The Production of a Movement.

    PubMed

    Morris, Bonnie

    2015-01-01

    This article looks at the early years of Olivia Records, setting the context for the historic release of the album Where Would I Be Without You. From its origins as a Washington, D.C.-based activist collective in 1973, Olivia became a hugely successful recording company, marketing radical lesbian recordings and performances that soon defined the "women's music" movement. Both artistically and politically, Olivia's woman-identified albums became the soundtrack for a generation awakening to lesbian activism. Pat Parker and Judy Grahn's 1976 spoken-word recording is a unique demonstration of Olivia's radical production values and expanding catalog.

  14. Geochronology and thermochronology of the gneisses of the Brasiliano/Pan-African Araguaia Belt: Records of exhumation of West Gondwana and Pangea break up

    NASA Astrophysics Data System (ADS)

    Dias, A. N. C.; Moura, C. A. V.; Milhomem Neto, J. M.; Chemale, F., Jr.; Girelli, T. J.; Masuyama, K. M.

    2017-12-01

    The 1200 km long Araguaia Belt, central-north Brazil, was formed as a result of the Neoproterozoic (Ediacaran) to Early Cambrian collage of West Gondwana and has several dome structures in whose nuclei crop out basement inliers of Archean and Paleoproterozoic orthogneisses. Thus, the orthogneisses from Lontra, Cocalândia, Colméia and Cantão dome structures were selected for fission-track thermochronology and U-Pb double dating. Three Archean magmatic episodes ranging from 2830.9 ± 6.9 Ma (Colméia dome) to 2905 ± 5.1 Ma (Lontra dome) and one Paleoproterozoic magmatic event at 1835 ± 31 Ma (Cantão dome) are recognized by U-Pb LA-MC-ICP-MS zircon dating. Fission-track thermochronology in zircon permitted to identify three well-defined age populations, which correspond to major regional geological events. The older population (489 ± 15 Ma to 498±8Ma) records the exhumation process and the orogenetic collapse of the Araguaia Belt. The intermediate population (331 ± 8 Ma to 345 ± 13 Ma) is related to the Gondwanides orogeny which affected the western margin of Gondwana and therefore the interior of South American. Finally, the younger fission track ages (197 ± 3 Ma, 197 ± 4 Ma, 198 ± 2 Ma and 208 ± 10 Ma) reflect crustal heating around 200 Ma, which is associated with the Lower Jurassic Mosquito tholeiitic magmatism and extended the influence of the Central Atlantic Magmatic Province to the Araguaia Belt. This mafic magmatism has been recognized in different regions of the South American and was an important source of heat for hydrocarbon generation in some Paleozoic intracratonic basins.

  15. Sharing Medical Data for Health Research: The Early Personal Health Record Experience

    PubMed Central

    Kaci, Liljana; Mandl, Kenneth D

    2010-01-01

    Background Engaging consumers in sharing information from personally controlled health records (PCHRs) for health research may promote goals of improving care and advancing public health consistent with the federal Health Information Technology for Economic and Clinical Health (HITECH) Act. Understanding consumer willingness to share data is critical to advancing this model. Objective The objective was to characterize consumer willingness to share PCHR data for health research and the conditions and contexts bearing on willingness to share. Methods A mixed method approach integrating survey and narrative data was used. Survey data were collected about attitudes toward sharing PCHR information for health research from early adopters (n = 151) of a live PCHR populated with medical records and self-reported behavioral and social data. Data were analyzed using descriptive statistics and logistic regression to characterize willingness, conditions for sharing, and variations by sociodemographic factors. Narrative data were collected through semistructured focus group and one-on-one interviews with a separate sample of community members (n = 30) following exposure to PCHR demonstrations. Two independent analysts coded narrative data for major and minor themes using a shared rubric of a priori defined codes and an iterative inductive process. Findings were triangulated with survey results to identify patterns. Results Of PHCR users, 138 out of 151 (91%) were willing to share medical information for health research with 89 (59%) favoring an opt-in sharing model. Willingness to share was conditioned by anonymity, research use, engagement with a trusted intermediary, transparency around PCHR access and use, and payment. Consumer-determined restrictions on content and timing of sharing may be prerequisites to sharing. Select differences in support for sharing under different conditions were observed across social groups. No gender differences were observed; however differences

  16. Haootia quadriformis n. gen., n. sp., interpreted as a muscular cnidarian impression from the Late Ediacaran period (approx. 560 Ma).

    PubMed

    Liu, Alexander G; Matthews, Jack J; Menon, Latha R; McIlroy, Duncan; Brasier, Martin D

    2014-10-22

    Muscle tissue is a fundamentally eumetazoan attribute. The oldest evidence for fossilized muscular tissue before the Early Cambrian has hitherto remained moot, being reliant upon indirect evidence in the form of Late Ediacaran ichnofossils. We here report a candidate muscle-bearing organism, Haootia quadriformis n. gen., n. sp., from approximately 560 Ma strata in Newfoundland, Canada. This taxon exhibits sediment moulds of twisted, superimposed fibrous bundles arranged quadrilaterally, extending into four prominent bifurcating corner branches. Haootia is distinct from all previously published contemporaneous Ediacaran macrofossils in its symmetrically fibrous, rather than frondose, architecture. Its bundled fibres, morphology, and taphonomy compare well with the muscle fibres of fossil and extant Cnidaria, particularly the benthic Staurozoa. Haootia quadriformis thus potentially provides the earliest body fossil evidence for both metazoan musculature, and for Eumetazoa, in the geological record.

  17. Depositional and diagenetic variability within the Cambrian Mount Simon Sandstone: Implications for carbon dioxide sequestration

    USGS Publications Warehouse

    Bowen, B.B.; Ochoa, R.I.; Wilkens, N.D.; Brophy, J.; Lovell, T.R.; Fischietto, N.; Medina, C.R.; Rupp, J.A.

    2011-01-01

    The Cambrian Mount Simon Sandstone is the major target reservoir for ongoing geologic carbon dioxide (CO2) sequestration demonstrations throughout the midwest United States. The potential CO2 reservoir capacity, reactivity, and ultimate fate of injected CO2 depend on textural and compositional properties determined by depositional and diagenetic histories that vary vertically and laterally across the formation. Effective and efficient prediction and use of the available pore space requires detailed knowledge of the depositional and diagenetic textures and mineralogy, how these variables control the petrophysical character of the reservoir, and how they vary spatially. Here, we summarize the reservoir characteristics of the Mount Simon Sandstone based on examination of geophysical logs, cores, cuttings, and analysis of more than 150 thin sections. These samples represent different parts of the formation and depth ranges of more than 9000 ft (>2743 m) across the Illinois Basin and surrounding areas. This work demonstrates that overall reservoir quality and, specifically, porosity do not exhibit a simple relationship with depth, but vary both laterally and with depth because of changes in the primary depositional facies, framework composition (i.e., feldspar concentration), and diverse diagenetic modifications. Diagenetic processes that have been significant in modifying the reservoir include formation of iron oxide grain coatings, chemical compaction, feldspar precipitation and dissolution, multiple generations of quartz overgrowth cementation, clay mineral precipitation, and iron oxide cementation. These variables provide important inputs for calculating CO2 capacity potential, modeling reactivity, and are also an important baseline for comparisons after CO2 injection. Copyright ??2011. The American Association of Petroleum Geologists/Division of Environmental Geosciences. All rights reserved.

  18. Scientific interpretation of historical auroral records

    NASA Astrophysics Data System (ADS)

    Willis, D. M.; Stephenson, F. R.

    The available historical auroral records from both Europe and East Asia are examined critically for their relevance in the investigation of long-term variations in both solar activity and the Earth's magnetic field. The early oriental records are sufficiently numerous to allow scientific studies of variations on several time scales. Special attention is paid to the seasonal and secular variations of the early oriental auroral observations. In addition, the oriental auroral records exhibit a clear 27-day recurrence tendency at particular periods of time. A search has been made for examples of strictly simultaneous and indisputably independent observations of the aurora from spatially separated sites in East Asia. This search has yielded nine observations of mid-latitude auroral displays at more than one site in East Asia on the same night. A particular geomagnetic storm that occurred during December in AD 1128 is investigated in detail. Five days after the observation of two large sunspots in England, a red auroral display was observed from Korea. In addition, between the middle of AD 1127 and the middle of AD 1129, five Chinese and five Korean auroral observations were recorded. These provide evidence for recurrent auroral activity on a timescale almost exactly equal to the synodic-solar-rotation period (approximately 27 days). Finally, a new attempt is made to use the oriental historical auroral records to determine the location of the north geomagnetic pole during the European Middle Ages.

  19. New palaeomagnetic data from Argun terrane. Testing its association with Amuria and the Mongol-Okhotsk Ocean

    NASA Astrophysics Data System (ADS)

    Gordienko, I. V.; Metelkin, D. V.; Vetluzhskikh, L. I.; Mikhaltsov, N. E.; Kulakov, E. V.

    2018-06-01

    In this study, we present new palaeomagnetic and geological data obtained from Ediacaran and Cambrian sedimentary rocks of Argun terrane, which is traditionally considered a key element of the hypothetical Amuria composite continent. Since 1990, when Amuria was first proposed in palaeogeographic reconstructions, it became one of the principle members in the global palaeotectonic schemes. A scenario when collision of Amuria with Siberian margin resulted in formation of the Mongol-Okhotsk Ocean is universally accepted and supported by majority of researchers. However, time of Amuria's final assembly and relative position of the blocks within Amuria before the collision with Siberia is still a topic of debate. Questions about principal allocation of Argun terrane and its relation to Amuria during the late Neoroterozoic-Cambrian are addressed in this study. Palaeomagnetic poles for the Ediacaran-early Cambrian rocks of Argun terrane differ within an error from coeval poles from Siberia indicating that Argun terrane could have been located similar to its present-day position with respect to Siberia already at 560-525 Ma. This observation calls into question association of Argun terrane with Amuria, which in classic reconstructions is usually placed close to the North China Craton. It also questions our current understanding of the Amuria palaeocontinent and consequently, accuracy of global palaeogeographic reconstructions for the late Neoproterozoic-Cambrian in general, and those of the eastern part of the Central Asia in particular.

  20. Early evolution of multifocal optics for well-focused colour vision in vertebrates.

    PubMed

    Gustafsson, O S E; Collin, S P; Kröger, R H H

    2008-05-01

    Jawless fishes (Agnatha; lampreys and hagfishes) most closely resemble the earliest stage in vertebrate evolution and lamprey-like animals already existed in the Lower Cambrian [about 540 million years ago (MYA)]. Agnathans are thought to have separated from the main vertebrate lineage at least 500 MYA. Hagfishes have primitive eyes, but the eyes of adult lampreys are well-developed. The southern hemisphere lamprey, Geotria australis, possesses five types of opsin genes, three of which are clearly orthologous to the opsin genes of jawed vertebrates. This suggests that the last common ancestor of all vertebrate lineages possessed a complex colour vision system. In the eyes of many bony fishes and tetrapods, well-focused colour images are created by multifocal crystalline lenses that compensate for longitudinal chromatic aberration. To trace the evolutionary origins of multifocal lenses, we studied the optical properties of the lenses in four species of lamprey (Geotria australis, Mordacia praecox, Lampetra fluviatilis and Petromyzon marinus), with representatives from all three of the extant lamprey families. Multifocal lenses are present in all lampreys studied. This suggests that the ability to create well-focused colour images with multifocal optical systems also evolved very early.

  1. Support Vector Feature Selection for Early Detection of Anastomosis Leakage From Bag-of-Words in Electronic Health Records.

    PubMed

    Soguero-Ruiz, Cristina; Hindberg, Kristian; Rojo-Alvarez, Jose Luis; Skrovseth, Stein Olav; Godtliebsen, Fred; Mortensen, Kim; Revhaug, Arthur; Lindsetmo, Rolv-Ole; Augestad, Knut Magne; Jenssen, Robert

    2016-09-01

    The free text in electronic health records (EHRs) conveys a huge amount of clinical information about health state and patient history. Despite a rapidly growing literature on the use of machine learning techniques for extracting this information, little effort has been invested toward feature selection and the features' corresponding medical interpretation. In this study, we focus on the task of early detection of anastomosis leakage (AL), a severe complication after elective surgery for colorectal cancer (CRC) surgery, using free text extracted from EHRs. We use a bag-of-words model to investigate the potential for feature selection strategies. The purpose is earlier detection of AL and prediction of AL with data generated in the EHR before the actual complication occur. Due to the high dimensionality of the data, we derive feature selection strategies using the robust support vector machine linear maximum margin classifier, by investigating: 1) a simple statistical criterion (leave-one-out-based test); 2) an intensive-computation statistical criterion (Bootstrap resampling); and 3) an advanced statistical criterion (kernel entropy). Results reveal a discriminatory power for early detection of complications after CRC (sensitivity 100%; specificity 72%). These results can be used to develop prediction models, based on EHR data, that can support surgeons and patients in the preoperative decision making phase.

  2. A record of reversed polarity carries by the iron sulphide greigite in British early Pleistocene sediments

    NASA Astrophysics Data System (ADS)

    Hallam, D. F.; Maher, B. A.

    1994-01-01

    Palaeomagnetic measurements were made on samples extracted from a short sequence of early Pleistocene estuarine clays, now exposed in a sea cliff near Sheringham on the north Norfolk coast, UK. On the basis of earlier palynological work, these clays had been ascribed a Pastonian (late Tiglian) age. The clays show marked changes in colour, from reddish-brown at the top of the unit, to blue-grey in the middle, and grey-brown at the base. The palaeomagnetic data vary in close association with these colour changes. The top and basal brown clays show scattered normal directions of low intensity, while the middle blue clays show strongly clustered reversed directions, of much higher intensities. Some samples taken from the boundary between the middle blue clays and upper red clays show upon demagnetisation a normal overprint on a stable reversed polarity. Using high-gradient magnetic extraction, magnetic concentrates have been obtained from the strongly magnetic middle blue clays. The presence of iron sulphide minerals in these concentrates was identified using energy-dispersive X-ray analysis during scanning electron microscopy. More specifically, X-ray diffraction identifies greigite as the only detectable ferrimagnetic mineral in the magnetic concentrates. Rock magnetic measurements show clear qualitative differences in the magnetic mineralogies of the three clay subunits, but absolute identification of the magnetic mineralogy of the weakly magnetic upper and basal brown clays has not yet been possible. We interpret the sequence as a primary reversed polarity record. This record is carried by the iron sulphide greigate as a chemical remanence acquired during `syn'-depositional reduction of iron via the decomposition of organic material in these anoxic tidal clays. Subsequently, the upper and basl subunits of the clay have been oxidised by permeation of groundwater from the adjacent coarse-grained sediments. Most of the griegite in the oxidised margins of the clay has

  3. The early Barremian warm pulse and the late Barremian cooling: a high resolution geochemical record of the Boreal Realm

    NASA Astrophysics Data System (ADS)

    Mutterlose, J.; Malkoc, M.

    2009-04-01

    A high resolution geochemical study of seven Barremian (lower Cretaceous) sections in NW Europe documents warming and cooling periods throughout the investigated interval. A total of 289 belemnite guards from localities in NE England and NW Germany were analysed for their trace elements (Mg, Sr, Fe and Mn) and their stable isotope composition (δ13C, δ18O) in order to reconstruct the palaeotemperature record and to better understand different palaeotemperature proxies (δ18O, Mg/Ca). The material has been collected bed-by-bed from bio- and lithostratigraphically well studied outcrops with a palaeolatitude of 36-38°N. The carbon isotope curve of the Barremian is described by a long term (5my) increase from 1‰ in the early Barremian to 3‰ in the late Barremian. This long term trend is thought to reflect the global signal of the carbon budget of the Barremian oceans. Superimposed on this signature is a short termed peak (~ 105kyrs) of 2.5‰ in the late early Barremian Aulacoteuthis zone. This "Aulacoteuthis peak", which goes along with the widespread deposition of TOC rich mudstones, was caused by increased productivity. It is seen as a regional signal related to warm-humid conditions and palaeoceanographically a quasi lagoonal setting. δ13C shifts of 1.5‰ from specimens collected from one horizon may reflect high resolution seasonal changes of run-off and nutrient supply. The oxygen isotope signature clearly shows similar palaeotemperature variations of the approx. 5my lasting Barremian for all sections investigated. The climatic evolution allows to recognize three different phases. An earliest Barremian cool phase (Praeoxyteuthis pugio zone) is characterized by δ18O values of - 1.2‰ to 0.3‰ (16°C to 10°C) with most data plotting around - 0.2‰ (12°C). This "P. pugio cool phase", which extends into the underlying Hauterivian, is followed by a distinctive warming event in the late early Barremian Aulacoteuthis zone. This "Aulacoteuthis warm pulse

  4. Exploring for early bombardments on Earth from pre-3.85 Fa thermal effects recorded in Hadean zircons - a status report

    NASA Astrophysics Data System (ADS)

    Mojzsis, S. J.; Abramov, O.; Harrison, T. M.; Kring, D. A.; Levison, H. F.; Trail, D.; Watson, E. B.

    2008-12-01

    We report on our progress with high-resolution ion microprobe U-Th-Pb depth profiles and Ti+REEs spot analysis which show that subsequent to their crystallization in melts under typical crustal conditions on Earth, some Hadean (pre-3.85 Ga) zircons record common age domains with unusual chemical and isotopic characteristics consistent with a high-temperature (possibly impact) origin. We have found evidence for later overprints caused by intense thermal alteration between 3.94-3.97 Ga in six of eight studied grains but no evidence for older events. These findings alert us to two fundamental things we did not know before about the probiotic potential of the Earth in the earliest solar system: (i) that the bombardment epoch did not result in complete 'Doomsday' scale destruction of the Earth's crust since the Moon-forming event at ca. 4.5 Ga; and (ii) age constraints on both sides of the ther-mally altered 3.94-3.97 Ga zircon domains are very good and so far our data show that no detectable thermal events are recorded by the zircons before ~3.97 Ga up to about 4.3 Ga. This observation is consistent with the output of new classes of dynamical models that successfully re-create the decay of impactor populations in the early solar system as recorded on the Moon and in meteorites.

  5. A palaeoequatorial ornithischian and new constraints on early dinosaur diversification.

    PubMed

    Barrett, Paul M; Butler, Richard J; Mundil, Roland; Scheyer, Torsten M; Irmis, Randall B; Sánchez-Villagra, Marcelo R

    2014-09-22

    Current characterizations of early dinosaur evolution are incomplete: existing palaeobiological and phylogenetic scenarios are based on a fossil record dominated by saurischians and the implications of the early ornithischian record are often overlooked. Moreover, the timings of deep phylogenetic divergences within Dinosauria are poorly constrained owing to the absence of a rigorous chronostratigraphical framework for key Late Triassic-Early Jurassic localities. A new dinosaur from the earliest Jurassic of the Venezuelan Andes is the first basal ornithischian recovered from terrestrial deposits directly associated with a precise radioisotopic date and the first-named dinosaur from northern South America. It expands the early palaeogeographical range of Ornithischia to palaeoequatorial regions, an area sometimes thought to be devoid of early dinosaur taxa, and offers insights into early dinosaur growth rates, the evolution of sociality and the rapid tempo of the global dinosaur radiation following the end-Triassic mass extinction, helping to underscore the importance of the ornithischian record in broad-scale discussions of early dinosaur history. © 2014 The Author(s) Published by the Royal Society. All rights reserved.

  6. The impact of childhood residential mobility on mental health outcomes in adolescence and early adulthood: a record linkage study.

    PubMed

    Tseliou, Foteini; Maguire, Aideen; Donnelly, Michael; O'Reilly, Dermot

    2016-03-01

    Understanding the causes of poor mental health in early childhood and adolescence is important as this can be a significant determinant of mental well-being in later years. One potential and relatively unexplored factor is residential mobility in formative years. Previous studies have been relatively small and potentially limited due to methodological issues. The main aim of this study was to investigate the relationship between early residential instability and poor mental health among adolescents and young adults in Northern Ireland. A Census-based record linkage study of 28% of children aged 0-8 years in 2001 in Northern Ireland (n=49,762) was conducted, with six monthly address change assessments from health registration data and self-reported mental health status from the 2011 Census. Logistic regression models were built adjusting for socioeconomic status (SES), household composition and marital dissolution. There was a graded relationship between the number of address changes and mental ill-health (adjusted OR 3.67, 95% CIs 2.11 to 6.39 for 5 or more moves). This relationship was not modified by SES or household composition. Marital dissolution was associated with poor mental health but did not modify the relationship between address change and mental health (p=0.206). There was some indication that movement after the age of five was associated with an increased likelihood of poor mental health. This large study clearly confirms the close relationship between address change in early years and later poor mental health. Residential mobility may be a useful marker for children at risk of poorer mental health in adolescence and early adulthood. Published by the BMJ Publishing Group Limited. For permission to use (where not already granted under a licence) please go to http://www.bmj.com/company/products-services/rights-and-licensing/

  7. Le graben de l'Anti-Atlas occidental (Maroc) : contrôle tectonique de la paléogéographie et des séquences au Cambrien inférieurThe Lower-Cambrian western Anti-Atlasic graben: tectonic control of palaeogeography and sequential organisation

    NASA Astrophysics Data System (ADS)

    Benssaou, Mohammed; Hamoumi, Naı̈ma

    2003-03-01

    In the Moroccan western Anti-Atlas, the combined extensive tectonic events with a long-term sea-level rise is the main factor on building vertical stacking transgressive-regressive sequences. In the Ait Abdallah-Boussafene axis, the subsidence processes, relayed by a brutal platform tilting generated an elongated NE-SW graben. This is an evidence of the persistence of the Anti-Atlasic rifting process during the last part of the Lower-Cambrian succession.

  8. Early Eocene biota (ostracoda, foraminifera) and paleoenvironment of the Blue Marls in the Corbieres Hills (Aude, France): building a framework for the identification of early Eocene hyperthermals in continental margin records.

    NASA Astrophysics Data System (ADS)

    Pirkenseer, C.; King, C.; Steurbaut, E.; Speijer, R.

    2009-04-01

    The Corbières Foreland Basin represents the southeastern-most extension of the Aquitanian Basin and is thus palaeo(bio)geographically related to the West-European Cenozoic Basin. During the Ypresian (‘Ilerdien') a succession of marine carbonates (e.g., Calcaires blancs à Alvéolines), marine marls (Blue Marls, Marnes à Térebratulides), brackish marls to sandstones and subsequent fluvio-lacustrine sediments (e.g., Montlaur Molasse) were deposited in the Corbières Hills (Aude, France) area in several depositional sequences. The present study focuses on the upper part of the open marine Blue Marls and the overlying brackish marls and sandstones spanning about 120m thickness close to the village Pradelles-en-Val. Over one hundred samples were collected in 1m intervals in order to document the early Eocene biogeographical and paleoenvironmental evolution of this open marine sequence, through a quantitative analysis of the ostracod assemblages. Furthermore, we aim at identifying anomalous environmental conditions that might be expected to be associated with the early Eocene hyperthermals known as Elmo- (ETM2) and X-event (ETM3). These events are subordinate to the best known hyperthermal, the Paleocene-Eocene thermal maximum, which has been recorded in deep-sea to non-marine depositional settings. ETM2 and ETM3, however, have until now only been demonstrated in deep-sea sequences, not in shelf deposits. In accordance with biostratigraphical data derived from other outcrops in the region, the sampled succession is attributed to the interval of calcareous nannofossil zones NP10-NP12. The occurrences of planktonic foraminifera of the Morozovella subbotinae-group are in agreement with this stratigraphic position (P6-7) for the lower part of the profile. Recorded fossil groups include generally abundant marine ostracoda, bryozoa, benthic and planktonic foraminifera, fragments of echinoderms including ophiuroidea, moulds of gastropods (often pyritised), large dinocysts

  9. Recording of electrohysterogram laplacian potential.

    PubMed

    Alberola-Rubio, J; Garcia-Casado, J; Ye-Lin, Y; Prats-Boluda, G; Perales, A

    2011-01-01

    Preterm birth is the main cause of the neonatal morbidity. Noninvasive recording of uterine myoelectrical activity (electrohysterogram, EHG) could be an alternative to the monitoring of uterine dynamics which are currently based on tocodynamometers (TOCO). The analysis of uterine electromyogram characteristics could help the early diagnosis of preterm birth. Laplacian recordings of other bioelectrical signals have proved to enhance spatial selectivity and to reduce interferences in comparison to monopolar and bipolar surface recordings. The main objective of this paper is to check the feasibility of the noninvasive recording of uterine myoelectrical activity by means of laplacian techniques. Four bipolar EHG signals, discrete laplacian obtained from five monopolar electrodes and the signals picked up by two active concentric-ringed-electrodes were recorded on 5 women with spontaneous or induced labor. Intrauterine pressure (IUP) and TOCO were also simultaneously recorded. To evaluate the uterine contraction detectability of the different noninvasive methods in comparison to IUP the contractions consistency index (CCI) was calculated. Results show that TOCO is less consistent (83%) than most EHG bipolar recording channels (91%, 83%, 87%, and 76%) to detect the uterine contractions identified in IUP. Moreover laplacian EHG signals picked up by ringed-electrodes proved to be as consistent (91%) as the best bipolar recordings in addition to significantly reduce ECG interference.

  10. Estimation of divergence times in cnidarian evolution based on mitochondrial protein-coding genes and the fossil record.

    PubMed

    Park, Eunji; Hwang, Dae-Sik; Lee, Jae-Seong; Song, Jun-Im; Seo, Tae-Kun; Won, Yong-Jin

    2012-01-01

    The phylum Cnidaria is comprised of remarkably diverse and ecologically significant taxa, such as the reef-forming corals, and occupies a basal position in metazoan evolution. The origin of this phylum and the most recent common ancestors (MRCAs) of its modern classes remain mostly unknown, although scattered fossil evidence provides some insights on this topic. Here, we investigate the molecular divergence times of the major taxonomic groups of Cnidaria (27 Hexacorallia, 16 Octocorallia, and 5 Medusozoa) on the basis of mitochondrial DNA sequences of 13 protein-coding genes. For this analysis, the complete mitochondrial genomes of seven octocoral and two scyphozoan species were newly sequenced and combined with all available mitogenomic data from GenBank. Five reliable fossil dates were used to calibrate the Bayesian estimates of divergence times. The molecular evidence suggests that cnidarians originated 741 million years ago (Ma) (95% credible region of 686-819), and the major taxa diversified prior to the Cambrian (543 Ma). The Octocorallia and Scleractinia may have originated from radiations of survivors of the Permian-Triassic mass extinction, which matches their fossil record well. Copyright © 2011 Elsevier Inc. All rights reserved.

  11. Evidence for Cambrian petroleum source rocks in the Rome trough of West Virginia and Kentucky, Appalachian basin: Chapter G.8 in Coal and petroleum resources in the Appalachian basin: distribution, geologic framework, and geochemical character

    USGS Publications Warehouse

    Ryder, Robert T.; Harris, David C.; Gerome, Paul; Hainsworth, Timothy J.; Burruss, Robert A.; Lillis, Paul G.; Jarvie, Daniel M.; Pawlewicz, Mark J.; Ruppert, Leslie F.; Ryder, Robert T.

    2014-01-01

    The bitumen extract from the Rogersville Shale compares very closely with oils or condensates from Cambrian reservoirs in the Carson Associates No. 1 Kazee well, Homer gas field, Elliott County, Ky.; the Inland No. 529 White well, Boyd County, Ky.; and the Miller No. 1 well, Wolfe County, Ky. These favorable oil-source rock correlations suggest a new petroleum system in the Appalachian basin that is characterized by a Conasauga Group source rock and Rome Formation and Conasauga Group reservoirs. This petroleum system probably extends along the Rome trough from eastern Kentucky to at least central West Virginia.

  12. An outline of the palaeogeographic evolution of the Australasian region since the beginning of the Neoproterozoic

    NASA Astrophysics Data System (ADS)

    Li, Z. X.; Powell, C. McA.

    2001-04-01

    In the last 1000 million years, Australia has been part of two supercontinents: Palaeozoic Gondwanaland and Neoproterozoic Rodinia. Neoproterozoic Australia was covered by shallow epicontinental seas, and, in the late Neoproterozoic, by low-latitude glaciers. The breakup of Rodinia along the Tasman Line occurred at the end of the Sturtian glaciation (760 Ma) giving rise to the Palaeo-Pacific Ocean. Gondwanaland formed in the Early Cambrian, at the same time as the Tarim block broke away from northwestern Australia. Westward subduction of the Palaeo-Pacific Ocean along the eastern margin of Australia-Antarctica commenced during the Early Cambrian in northern Victoria Land and in the Middle Cambrian in South Australia, and culminated to the Late Cambrian-Early Ordovician Ross-Delamerian Mountains. In the Ordovician, the magmatic arc retreated from Australia's then-eastern continental margin, forming a marginal sea and offshore island arc. A shallow seaway across Australia in the Late Cambrian and Ordovician gradually gave way to desert-like conditions in Central Australia and the adjacent Canning Basin by Silurian time. The Silurian to mid-Devonian was an interval of rapidly changing palaeogeography in eastern Australia with deep volcanogenic troughs formed in a dextral transtensional tectonic setting. Widespread deformation in the Tasman orogenic zone in the Middle Devonian to Early Carboniferous, was accompanied by the development of an Andean-style magmatic arc along the Pacific continental margin of Australia. The most widespread Phanerozoic mountain-building stage in Central Australia occurred in the Late Devonian to mid-Carboniferous, as part of a world-wide Variscan orogenic episode associated with the collision of Gondwanaland with Laurussia to form Pangea. In the late Visean, Australia drifted rapidly southward from previous low latitudes to a near-polar position. Glacial conditions dominated the Late Carboniferous and earliest Permian. Transtensional basins

  13. New Early Triassic trace fossil records from South China: implications for biotic recovery following the end-Permian mass extinction

    NASA Astrophysics Data System (ADS)

    Luo, M.; George, A. D.; Chen, Z.; Zhang, Y.

    2013-12-01

    New Early Triassic trace fossil assemblages are documented from the Susong and Tianshengqiao areas in South China to evaluate the mode and tempo of biotic recovery of epifaunal and infaunal organisms following the end-Permian mass extinction. The Susong succession is exposed in Anhui area of the Lower Yangtze region and comprises mudstone and carbonate facies that record overall shallowing from offshore to supratidal settings. The Tianshengqiao succession crops out in the Luoping area, Yuannan Province of the Upper Yangtze region, and consists of mixed carbonate and siliciclastic facies which were deposited in shallow marine to offshore settings. Bivalve and conodont biostratigraphy helps constrain the chronostratigraphic framework of the Lower Triassic successions in these two sections. Griesbachian to Dieneria ichnological records in both successions are characterized by low ichnodiversity, low ichnofabric indices (ii=1-2) and low bedding plane bioturbation indices (bpbi=1-2). Higher ii (ii= 3 and 4) corresponding to densely populated diminutive Skolithos in the Tianshengqiao succession suggest an opportunistic strategy during earliest Triassic deposition. Ichnological data from the Susong succession show an increase in ichnodiversity during the Smithian. A total of 12 ichnogenera including Arenicolites, Chondrites, Gyrochorte, Laevicyclus, Monocraterion, Palaeophycus, Phycodes, Plaolites, Thalassinoides, Treptichnus, Trichichnus and one problematic trace are identified. Ichnofabric indices (ii) and bpbi increase to moderate to high levels (ii = 4-5, bpbi= 3-5). Although complex traces such as Rhizocorallium are in Spathian strata in this section, the low levels of ichnodiversity, ichnofabric indices and diminutive Planolites suggest a decline in recovery. In the Tianshengqiao succession, ichnofabric indices exhibit a moderate to high value (ii= 3 to 5), however, only six ichnogenera are found and Planolites burrows are consistently small (average diameter at 3

  14. Fruits and wood of Parinari from the early Miocene of Panama and the fossil record of Chrysobalanaceae.

    PubMed

    Jud, Nathan A; Nelson, Chris W; Herrera, Fabiany

    2016-02-01

    Chrysobalanaceae are woody plants with over 500 species in 20 genera. They are among the most common trees in tropical forests, but a sparse fossil record has limited our ability to test evolutionary and biogeographic hypotheses, and several previous reports of Chrysobalanaceae megafossils are doubtful. We prepared fossil endocarps and wood collected from the lower Miocene beds along the Panama Canal using the cellulose acetate peel technique and examined them using light microscopy. We compared the fossil endocarps with previously published fossils and with fruits from herbarium specimens. We compared the fossil wood with photographs and descriptions of extant species. Parinari endocarps can be distinguished from other genera within Chrysobalanaceae by a suite of features, i.e., thick wall, a secondary septum, seminal cavities lined with dense, woolly trichomes, and two ovate to lingulate basal germination plugs. Fossil endocarps from the Cucaracha, Culebra, and La Boca Formations confirm that Parinari was present in the neotropics by the early Miocene. The earliest unequivocal evidence of crown-group Chrysobalanaceae is late Oligocene-early Miocene, and the genus Parinari was distinct by at least 19 million years ago. Parinari and other Chrysobalanaceae likely reached the neotropics via long-distance dispersal rather than vicariance. The presence of Parinari in the Cucaracha flora supports the interpretation of a riparian, moist tropical forest environment. Parinari was probably a canopy-dominant tree in the Cucaracha forest and took advantage of the local megafauna for seed dispersal. © 2016 Botanical Society of America.

  15. Cambrian origin of the CYP27C1-mediated vitamin A1-to-A2 switch, a key mechanism of vertebrate sensory plasticity

    USGS Publications Warehouse

    Morshedian, Ala; Toomery, Matthew B.; Pollock, Gabriel E.; Frederiksen, Rikard; Enright, Jennifer; McCormick, Stephen; Cornwall, M. Carter; Fain, Gordon L.; Corbo, Joseph C.

    2017-01-01

    The spectral composition of ambient light varies across both space and time. Many species of jawed vertebrates adapt to this variation by tuning the sensitivity of their photoreceptors via the expression of CYP27C1, an enzyme that converts vitamin A1 into vitamin A2, thereby shifting the ratio of vitamin A1-based rhodopsin to red-shifted vitamin A2-based porphyropsin in the eye. Here, we show that the sea lamprey (Petromyzon marinus), a jawless vertebrate that diverged from jawed vertebrates during the Cambrian period (approx. 500 Ma), dynamically shifts its photoreceptor spectral sensitivity via vitamin A1-to-A2 chromophore exchange as it transitions between photically divergent aquatic habitats. We further show that this shift correlates with high-level expression of the lamprey orthologue of CYP27C1, specifically in the retinal pigment epithelium as in jawed vertebrates. Our results suggest that the CYP27C1-mediated vitamin A1-to-A2 switch is an evolutionarily ancient mechanism of sensory plasticity that appeared not long after the origin of vertebrates.

  16. Paleoclimatic and paleoceanographic records through Marine Isotope Stage 19 at the Chiba composite section, central Japan: A key reference for the Early-Middle Pleistocene Subseries boundary

    NASA Astrophysics Data System (ADS)

    Suganuma, Yusuke; Haneda, Yuki; Kameo, Koji; Kubota, Yoshimi; Hayashi, Hiroki; Itaki, Takuya; Okuda, Masaaki; Head, Martin, J.; Sugaya, Manami; Nakazato, Hiroomi; Igarashi, Atsuo; Shikoku, Kizuku; Hongo, Misao; Watanabe, Masami; Satoguchi, Yasufumi; Takeshita, Yoshihiro; Nishida, Naohisa; Izumi, Kentaro; Kawamura, Kenji; Kawamata, Moto; Okuno, Jun'ichi; Yoshida, Takeshi; Ogitsu, Itaru; Yabusaki, Hisashi; Okada, Makoto

    2018-07-01

    Marine Isotope Stage (MIS) 19 is an important analogue for the present interglacial because of its similar orbital configuration, especially the phasing of the obliquity maximum to precession minimum. However, sedimentary records suitable for capturing both terrestrial and marine environmental changes are limited, and thus the climatic forcing mechanisms for MIS 19 are still largely unknown. The Chiba composite section, east-central Japanese archipelago, is a continuous and expanded marine sedimentary succession well suited to capture terrestrial and marine environmental changes through MIS 19. In this study, a detailed oxygen isotope chronology is established from late MIS 20 to early MIS 18, supported by a U-Pb zircon age and the presence of the Matuyama-Brunhes boundary. New pollen, marine microfossil, and planktonic foraminiferal δ18O and Mg/Ca paleotemperature records reveal the complex interplay of climatic influences. Our pollen data suggest that the duration of full interglacial conditions during MIS 19 extends from 785.0 to 775.1 ka (9.9 kyr), which offers an important natural baseline in predicting the duration of the present interglacial. A Younger Dryas-type cooling event is present during Termination IX, suggesting that such events are linked to this orbital configuration. Millennial- to multi-millennial-scale variations in our δ18O and Mg/Ca records imply that the Subarctic Front fluctuated in the northwestern Pacific Ocean during late MIS 19, probably in response to East Asian winter monsoon variability. The climatic setting at this time appears to be related to less severe summer insolation minima at 65˚N and/or high winter insolation at 50˚N. Our records do not support a recently hypothesized direct coupling between variations in the geomagnetic field intensity and global/regional climate change. Our highly resolved paleoclimatic and paleoceanographic records, coupled with a well-defined Matuyama-Brunhes boundary (772.9 ka; duration 1.9 kyr

  17. Selenium speciation in Lower Cambrian Se-enriched strata in South China and its geological implications

    NASA Astrophysics Data System (ADS)

    Fan, Haifeng; Wen, Hanjie; Hu, Ruizhong; Zhao, Hui

    2011-12-01

    To understand the impact of Selenium (Se) into the biogeochemical cycle and implications for palaeo-redox environment, a sequential extraction method was utilized for samples including black shales, cherts, a Ni-Mo-Se sulfide layer, K-bentonite and phosphorite from Lower Cambrian Se-enriched strata in southern China. Seven species (water-soluble, phosphate exchangeable, base-soluble, acetic acid-soluble, sulfide/selenide associated, residual Se) and different oxidation states (selenate Se(VI), selenite Se(IV), organic Se, Se (0) and mineral Se(-II)) were determinated in this study. We found that the Ni-Mo-Se sulfide layer contained a significantly greater amount of Se(-II) associated with sulfides/selenides than those in host black shales and cherts. Furthermore, a positive correlation between the degree of sulfidation of iron (DOS) and the percentage of the sulfide/selenide-associated Se(-II) was observed for samples, which suggests the proportion of sulfide/selenide-associated Se(-II) could serve as a proxy for palaeo-redox conditions. In addition, the higher percentage of Se(IV) in K-bentonite and phosphorite was found and possibly attributed to the adsorption of Se by clay minerals, iron hydroxide surfaces and organic particles. Based on the negative correlations between the percentage of Se(IV) and that of Se(-II) in samples, we propose that the K-bentonite has been altered under the acid oxic conditions, and the most of black shale (and cherts) and the Ni-Mo-Se sulfide layer formed under the anoxic and euxinic environments, respectively. Concerning Se accumulation in the Ni-Mo-Se sulfide layer, the major mechanism can be described by (1) biotic and abiotic adsorption and further dissimilatory reduction from oxidized Se(VI) and Se(IV) to Se(-II), through elemental Se, (2) contribution of hydrothermal fluid with mineral Se(-II).

  18. The global record of local iron geochemical data from Proterozoic through Paleozoic basins

    NASA Astrophysics Data System (ADS)

    Sperling, E. A.; Wolock, C.; Johnston, D. T.; Knoll, A. H.

    2013-12-01

    Iron-based redox proxies represent one of the most mature tools available to sedimentary geochemists. These techniques, which benefit from decades of refinement, are based on the fact that rocks deposited under anoxic conditions tend to be enriched in highly-reactive iron. However, there are myriad local controls on the development of anoxia, and no local section is an exemplar for the global ocean. The global signal must thus be determined using techniques like those developed to solve an analogous problem in paleobiology: the inference of global diversity patterns through time from faunas seen in local stratigraphic sections. Here we analyze a dataset of over 4000 iron speciation measurements (including over 600 de novo analyses) to better understand redox changes from the Proterozoic through the Paleozoic Era. Preliminary database analyses yield interesting observations. We find that although anoxic water columns in the middle Proterozoic were dominantly ferruginous, there was a statistical tendency towards euxinia not seen in early Neoproterozoic or Ediacaran data. Also, we find that in the Neoproterozoic oceans, oxic depositional environments-the likely home for early animals-have exceptionally low pyrite contents, and by inference low levels of porewater sulfide. This runs contrary to notions of sulfide stress on early metazoans. Finally, the current database of iron speciation data does not support an Ediacaran or Cambrian oxygenation event. This conclusion is of course only as sharp as the ability of the Fe-proxy database to track dissolved oxygen and does not rule out the possibility of a small-magnitude change in oxygen. It does suggest, however, that if changing pO2 facilitated animal diversification it did so by a limited rise past critical ecological thresholds, such as seen in the modern Oxygen Minimum Zones benthos. Oxygen increase to modern levels thus becomes a Paleozoic problem, and one in need of better sampling if a database approach is to be

  19. Late-Glacial to Early Holocene Climate Changes from a Central Appalachians Pollen and Macrofossil Record

    NASA Technical Reports Server (NTRS)

    Kneller, Margaret; Peteet, Dorothy

    1997-01-01

    A Late-glacial to early Holocene record of pollen, plant macrofossils and charcoal, based on two cores, is presented for Browns Pond in the central Appalachians of Virginia. An AMS radiocarbon chronology defines the timing of moist and cold excursions, superimposed upon the overall warming trend from 14,200 to 7,500 C-14 yr B.P. This site shows cold, moist conditions from approximately 14,200 to 12,700 C-14 yr B.P., with warming at 12,730, 11,280 and 10,050 C-14 yr B.P. A decrease in deciduous broad-leaved tree taxa and Pinus strobus (haploxylon) pollen, simultaneous with a re-expansion of Abies denotes a brief, cold reversal from 12,260 to 12,200 C-14 yr B.P. A second cold reversal, inferred from increases in montane conifers, is centered at 7,500 C-14 yr B.P. The cold reversals at Browns Pond may be synchronous with climate change in Greenland, and northwestern Europe. Warming at 11,280 C-14 yr B.P. shows the complexity of regional climate responses during the Younger Dryas chronozone.

  20. Evidence for Water-Mass Changes on the Tasmanian Slope during the Early Miocene (19-16.5 Ma): Stable Isotope and Mg/Ca Records from ODP Leg 189 Site 1168

    NASA Astrophysics Data System (ADS)

    Pekar, S. F.; Marchitto, T. M.; Lynch-Steiglitz, J.

    2002-12-01

    High-resolution stable isotope (4-10 k.y. resolution) and moderately low-resolution Mg/Ca ratio records were constructed for the late early Miocene (19-16.5 Ma) from ODP Leg 189 Site 1168, located on the southwest slope of Tasmania. These records evaluated paleoceanographic changes that took place during isotopic excursions Mi1b (18.2-17.8 Ma) and Mi2 (16.5 Ma), and the First Climatic Optimum (17.7-16.7 Ma), a time of increased global warmth. Evidence exists that supports the idea for the development of warm saline deep waters (WSDW) originating from the eastern end of the Tethys Sea during the early Miocene. However, questions remain regarding the extent and strength of the WSDW and the possible role it played in the warming that took place during the First Climatic Optimum. Site 1168 is ideally located on the lower slope (estimates place it in lower bathyal waters during the early Miocene) to evaluate the potential penetration of WSDW and into the Southern Ocean. Large fluctuations in the isotope and Mg/Ca ratio records from Site 1168 suggest changes in the water masses that bathed the Tasmanian slope during the early Miocene. Temperature estimates based on Mg/Ca ratios contain a surprisingly high range, from 4° to 10° C. Low temperatures (4°-6° C) are associated with high carbon isotope values (>1.4‰ ) and are interpreted represent Southern Component Waters (SCW). The high carbon isotope values also suggest a proximal source for SCW. High water temperatures (7°-10° C) indicate a warm-water mass and are interpreted to be due to the penetration of WSDW into this area, replacing SCW at various times. Large high-frequency isotopic excursions (low oxygen and carbon isotope values) occurred between 18.7 and 18.4 Ma and were originally thought to be due to either localized effects (e.g., disassociation of hydrates) or possible diagensis. However, a recently published high-resolution isotopic record from the Southern Ocean (Site 1090) also contains large

  1. The controversial “Cambrian” fossils of the Vindhyan are real but more than a billion years older

    PubMed Central

    Bengtson, Stefan; Belivanova, Veneta; Rasmussen, Birger; Whitehouse, Martin

    2009-01-01

    The age of the Vindhyan sedimentary basin in central India is controversial, because geochronology indicating early Proterozoic ages clashes with reports of Cambrian fossils. We present here an integrated paleontologic–geochronologic investigation to resolve this conundrum. New sampling of Lower Vindhyan phosphoritic stromatolitic dolomites from the northern flank of the Vindhyans confirms the presence of fossils most closely resembling those found elsewhere in Cambrian deposits: annulated tubes, embryo-like globules with polygonal surface pattern, and filamentous and coccoidal microbial fabrics similar to Girvanella and Renalcis. None of the fossils, however, can be ascribed to uniquely Cambrian or Ediacaran taxa. Indeed, the embryo-like globules are not interpreted as fossils at all but as former gas bubbles trapped in mucus-rich cyanobacterial mats. Direct dating of the same fossiliferous phosphorite yielded a Pb–Pb isochron of 1,650 ± 89 (2σ) million years ago, confirming the Paleoproterozoic age of the fossils. New U–Pb geochronology of zircons from tuffaceous mudrocks in the Lower Vindhyan Porcellanite Formation on the southern flank of the Vindhyans give comparable ages. The Vindhyan phosphorites provide a window of 3-dimensionally preserved Paleoproterozoic fossils resembling filamentous and coccoidal cyanobacteria and filamentous eukaryotic algae, as well as problematic forms. Like Neoproterozoic phosphorites a billion years later, the Vindhyan deposits offer important new insights into the nature and diversity of life, and in particular, the early evolution of multicellular eukaryotes. PMID:19416859

  2. Late Pan-African and early Mesozoic brittle compressions in East and Central Africa: lithospheric deformation within the Congo-Tanzania Cratonic area

    NASA Astrophysics Data System (ADS)

    Delvaux, D.; Kipata, M. L.; Macheyeki, A. S.

    2012-04-01

    Tectonic reconstructions leading to the formation of the Central-African part of Gondwana have so far not much taken into account constraints provided by the evolution of brittle structures and related stress field. This is largely because little is known on continental brittle deformation in Equatorial Africa before the onset of the Mesozoic Central-African and Late Cenozoic East-African rifts. We present a synthesis of fault-kinematic data and paleostress inversion results from field surveys covering parts of Tanzania, Zambia and the Democratic Republic of Congo. It is based on investigations along the eastern margin of the Tanzanian craton, in the Ubendian belt between the Tanzanian craton and Bangweulu block, in the Lufilian Arc between the Kalahari and Congo cratons and along the Congo intracratonic basin. Paleostress tensors were computed for a substantial database by interactive stress tensor inversion and data subset separation, and the relative succession of major brittle events established. Two of them appear to be of regional importance and could be traced from one region to the other. The oldest one is the first brittle event recorded after the paroxysm of the Terminal Pan-African event that led to the amalgamation Gondwana at the Precambrian-Cambrian transition. It is related to compressional deformation with horizontal stress trajectories fluctuating from an E-W compression in Central Tanzania to NE-SW in the Ubende belt and Lufilian Arc. The second event is a transpressional inversion with a consistent NW-SE compression that we relate to the far-field effects of the active margin south of Gondwana during the late Triassic - early Jurassic.

  3. A new Cheirolepidiaceae (Coniferales) from the Early Jurassic of Patagonia (Argentina): Reconciling the records of impression and permineralized fossils.

    PubMed

    Escapa, Ignacio; Leslie, Andrew

    2017-02-01

    Plants preserved in different fossil modes provide complementary data concerning the paleobiology and evolutionary relationships among plant groups. New material from the Early Jurassic of Patagonia shows the importance of combining these sources of information, as we describe the first compression/impression fossils of Pararaucaria , a genus of the extinct conifer family Cheirolepidiaceae previously known from permineralized fossils. These fossils extend the temporal range of this genus and may allow its wider recognition in the fossil record. We studied fossil plants from the Early Jurassic (Pleinsbachian-Toarcian) locality of Taquetrén in Patagonia, Argentina using standard paleobotanical preparation and description techniques. Pararaucaria taquetrensis consists of isolated ovuliferous scales and small seed cones with helically arranged bract-scale complexes attached to scale-leaf foliage. Bract-scale complexes consist of separated bracts and ovuliferous scales with two seeds and three broad distal lobes. Pararaucaria taquetrensis represents the oldest known Cheirolepidiaceae seed cones from the Southern Hemisphere, and this material highlights the importance of compression and impression fossils in understanding the distribution of fossil taxa. This material also suggests that Cheirolepidiaceae cone scales can be easily confused with those of another common conifer family, the Araucariaceae, which has important implications for accurately understanding Mesozoic conifer diversity and paleoecology. © 2017 Botanical Society of America.

  4. A diploblastic radiate animal at the dawn of cambrian diversification with a simple body plan: distinct from Cnidaria?

    PubMed

    Yasui, Kinya; Reimer, James D; Liu, Yunhuan; Yao, Xiaoyong; Kubo, Daisuke; Shu, Degan; Li, Yong

    2013-01-01

    Microfossils of the genus Punctatus include developmental stages such as blastula, gastrula, and hatchlings, and represent the most complete developmental sequence of animals available from the earliest Cambrian. Despite the extremely well-preserved specimens, the evolutionary position of Punctatus has relied only on their conical remains and they have been tentatively assigned to cnidarians. We present a new interpretation of the Punctatus body plan based on the developmental reconstruction aided by recent advances in developmental biology. Punctatus developed from a rather large egg, gastrulated in a mode of invagination from a coeloblastura, and then formed a mouth directly from the blastopore. Spiny benthic hatchlings were distinguishable from swimming or crawling ciliate larvae found in cnidarians and sponges. A mouth appeared at the perihatching embryonic stage and was renewed periodically during growth, and old mouths transformed into the body wall, thus elongating the body. Growing animals retained a small blind gut in a large body cavity without partitioning by septa and did not form tentacles, pedal discs or holdfasts externally. A growth center at the oral pole was sufficient for body patterning throughout life, and the body patterning did not show any bias from radial symmetry. Contrary to proposed cnidarian affinity, the Punctatus body plan has basic differences from that of cnidarians, especially concerning a spacious body cavity separating ectoderm from endoderm. The lack of many basic cnidarian characters in the body patterning of Punctatus leads us to consider its own taxonomic group, potentially outside of Cnidaria.

  5. Precambrian animal life: probable developmental and adult cnidarian forms from Southwest China

    NASA Technical Reports Server (NTRS)

    Chen, Jun-Yuan; Oliveri, Paola; Gao, Feng; Dornbos, Stephen Q.; Li, Chia-Wei; Bottjer, David J.; Davidson, Eric H.

    2002-01-01

    The evolutionary divergence of cnidarian and bilaterian lineages from their remote metazoan ancestor occurred at an unknown depth in time before the Cambrian, since crown group representatives of each are found in Lower Cambrian fossil assemblages. We report here a variety of putative embryonic, larval, and adult microfossils deriving from Precambrian phosphorite deposits of Southwest China, which may predate the Cambrian radiation by 25-45 million years. These are most probably of cnidarian affinity. Large numbers of fossilized early planula-like larvae were observed under the microscope in sections. Though several forms are represented, the majority display remarkable conformity, which is inconsistent with the alternative that they are artifactual mineral inclusions. Some of these fossils are preserved in such high resolution that individual cells can be discerned. We confirm in detail an earlier report of the presence in the same deposits of tabulates, an extinct crown group anthozoan form. Other sections reveal structures that most closely resemble sections of basal modern corals. A large number of fossils similar to modern hydrozoan gastrulae were also observed. These again displayed great morphological consistency. Though only a single example is available, a microscopic animal remarkably similar to a modern adult hydrozoan is also presented. Taken together, the new observations reported in this paper indicate the existence of a diverse and already differentiated cnidarian fauna, long before the Cambrian evolutionary event. It follows that at least stem group bilaterians must also have been present at this time.

  6. Hurricane Mountain Formation melange: history of Cambro-Ordovician accretion of the Boundary Mountains terrane within the northern Appalachian orthotectonic zone

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Boone, G.M.; Boudette, E.L.

    1985-01-01

    The Hurricane Mountain Formation (HMF) melange and associated ophiolitic and volcanogenic formations of Cambrian and lowermost Ordovician age bound the SE margin of the Precambrian Y (Helikian) Chain Lakes Massif in western Maine. HMF melange matrix, though weakly metamorphosed, contains a wide variety of exotic greenschist to amphibolite facies blocks as components of its polymictic assemblage, but blocks of high-grade cratonal rocks such as those of Chain Lakes or Grenville affinity are lacking. Formations of melange exposed in structural culminations of Cambrian and Ordovician rocks NE of the HMF in Maine and in the Fournier Group in New Brunswick aremore » lithologically similar and probably tectonically correlative with the HMF; taken together, they may delineate a common pre-Middle Ordovician tectonic boundary. The authors infer that the Hurricane Mountain and St. Daniel melange belts define the SE and NW margins of the Boundary Mountains accreted terrane (BMT), which may consist of cratonal basement of Chain Lakes affinity extending from eastern Gaspe (deBroucker and St. Julien, 1985) to north-central New Hampshire. The Laurentian continental margin, underlain by Grenville basement, underplated the NW margin of this terrane, marked by the SDF suture zone, in late Cambrian to early Ordovician time, while terranes marked by Cambrian to Tremadocian (.) lithologies dissimilar to the Boundary Mountains terrane were accreted to its outboard margin penecontemporaneously. The docking of the Boundary Mountains terrane and the initiation of its peripheral melanges are equated to the Penobscottian disturbance.« less

  7. Arsenic stress after the Proterozoic glaciations

    NASA Astrophysics Data System (ADS)

    Chi Fru, Ernest; Arvestål, Emma; Callac, Nolwenn; El Albani, Abderrazak; Kilias, Stephanos; Argyraki, Ariadne; Jakobsson, Martin

    2015-12-01

    Protection against arsenic damage in organisms positioned deep in the tree of life points to early evolutionary sensitization. Here, marine sedimentary records reveal a Proterozoic arsenic concentration patterned to glacial-interglacial ages. The low glacial and high interglacial sedimentary arsenic concentrations, suggest deteriorating habitable marine conditions may have coincided with atmospheric oxygen decline after ~2.1 billion years ago. A similar intensification of near continental margin sedimentary arsenic levels after the Cryogenian glaciations is also associated with amplified continental weathering. However, interpreted atmospheric oxygen increase at this time, suggests that the marine biosphere had widely adapted to the reorganization of global marine elemental cycles by glaciations. Such a glacially induced biogeochemical bridge would have produced physiologically robust communities that enabled increased oxygenation of the ocean-atmosphere system and the radiation of the complex Ediacaran-Cambrian life.

  8. Haootia quadriformis n. gen., n. sp., interpreted as a muscular cnidarian impression from the Late Ediacaran period (approx. 560 Ma)

    PubMed Central

    Liu, Alexander G.; Matthews, Jack J.; Menon, Latha R.; McIlroy, Duncan; Brasier, Martin D.

    2014-01-01

    Muscle tissue is a fundamentally eumetazoan attribute. The oldest evidence for fossilized muscular tissue before the Early Cambrian has hitherto remained moot, being reliant upon indirect evidence in the form of Late Ediacaran ichnofossils. We here report a candidate muscle-bearing organism, Haootia quadriformis n. gen., n. sp., from approximately 560 Ma strata in Newfoundland, Canada. This taxon exhibits sediment moulds of twisted, superimposed fibrous bundles arranged quadrilaterally, extending into four prominent bifurcating corner branches. Haootia is distinct from all previously published contemporaneous Ediacaran macrofossils in its symmetrically fibrous, rather than frondose, architecture. Its bundled fibres, morphology, and taphonomy compare well with the muscle fibres of fossil and extant Cnidaria, particularly the benthic Staurozoa. Haootia quadriformis thus potentially provides the earliest body fossil evidence for both metazoan musculature, and for Eumetazoa, in the geological record. PMID:25165764

  9. Arsenic stress after the Proterozoic glaciations.

    PubMed

    Fru, Ernest Chi; Arvestål, Emma; Callac, Nolwenn; El Albani, Abderrazak; Kilias, Stephanos; Argyraki, Ariadne; Jakobsson, Martin

    2015-12-04

    Protection against arsenic damage in organisms positioned deep in the tree of life points to early evolutionary sensitization. Here, marine sedimentary records reveal a Proterozoic arsenic concentration patterned to glacial-interglacial ages. The low glacial and high interglacial sedimentary arsenic concentrations, suggest deteriorating habitable marine conditions may have coincided with atmospheric oxygen decline after ~2.1 billion years ago. A similar intensification of near continental margin sedimentary arsenic levels after the Cryogenian glaciations is also associated with amplified continental weathering. However, interpreted atmospheric oxygen increase at this time, suggests that the marine biosphere had widely adapted to the reorganization of global marine elemental cycles by glaciations. Such a glacially induced biogeochemical bridge would have produced physiologically robust communities that enabled increased oxygenation of the ocean-atmosphere system and the radiation of the complex Ediacaran-Cambrian life.

  10. Climate variability and volcanic history of the Eastern Romanian Carpathians since early MIS 3 recorded in sediments from Mohoş crater

    NASA Astrophysics Data System (ADS)

    Bormann, M.; Veres, D.; Wulf, S.; Papadopoulou, M.; Panagiotopoulos, K.; Schaebitz, F.

    2015-12-01

    We present a 30m long sediment record covering the last ca. 50,000 years from the in-filled Mohoş crater (46°05' N; 25°55' E) located on Ciomadul volcano (Romania) that was retrieved in 2014. The record consists of bog and lacustrine sediments that are inter-bedded with tephra deposits. Ciomadul volcano, hosting the superimposed craters of Mohoş and Sf. Ana, is the youngest volcanic edifice in the Carpathian-Balkan region. Thus, tephra-analysis on the Mohoş sediments gives valuable insights into the volcanic history of that region, mainly arising from the younger crater of Sf Ana and several secondary domes. For investigations into the past climate history, the Mohoş sediment sequence has been analysed using a multi-proxy approach including geophysical, geochemical and sedimentological parameters. Multi-Sensor core logging and ITRAX X-ray fluorescence scanning have been performed at high-resolution, whereas grain size analysis, TOC and C/N ratios supplement the geophysical and geochemical data. Chronological control is based on radiocarbon and luminescence dating. We also present first results of the tephra-analysis on the Mohoş sediment record and their correlation to medium-distal pyroclastic deposits originating in this volcanic field. We further discuss responses of this mid-altitude site (1050 m a.s.l.) to past climate oscillations since early MIS 3. To date, the Mohoş core record provides the longest time series from the Carpathian region. This study is part of the Collaborative Research Centre 806 "Our Way To Europe; Culture-Environment Interaction and Human Mobility in the Late Quaternary" (www.sfb806.de); subproject B2.

  11. Earth’s oldest ‘Bobbit worm’ – gigantism in a Devonian eunicidan polychaete

    PubMed Central

    Eriksson, Mats E.; Parry, Luke A.; Rudkin, David M.

    2017-01-01

    Whilst the fossil record of polychaete worms extends to the early Cambrian, much data on this group derive from microfossils known as scolecodonts. These are sclerotized jaw elements, which generally range from 0.1–2 mm in size, and which, in contrast to the soft-body anatomy, have good preservation potential and a continuous fossil record. Here we describe a new eunicidan polychaete, Websteroprion armstrongi gen. et sp. nov., based primarily on monospecific bedding plane assemblages from the Lower-Middle Devonian Kwataboahegan Formation of Ontario, Canada. The specimens are preserved mainly as three-dimensional moulds in the calcareous host rock, with only parts of the original sclerotized jaw walls occasionally present. This new taxon has a unique morphology and is characterized by an unexpected combination of features seen in several different Palaeozoic polychaete families. Websteroprion armstrongi was a raptorial feeder and possessed the largest jaws recorded in polychaetes from the fossil record, with maxillae reaching over one centimetre in length. Total body length of the species is estimated to have reached over one metre, which is comparable to that of extant ‘giant eunicid’ species colloquially referred to as ‘Bobbit worms’. This demonstrates that polychaete gigantism was already a phenomenon in the Palaeozoic, some 400 million years ago. PMID:28220886

  12. Magneto- and litho-stratigraphic records of the Oligocene-Early Miocene climatic changes from deep drilling in the Linxia Basin, Northeast Tibetan Plateau

    NASA Astrophysics Data System (ADS)

    Wu, Fuli; Fang, Xiaomin; Meng, Qingquan; Zhao, Yan; Tang, Fenjun; Zhang, Tao; Zhang, Weilin; Zan, Jinbo

    2017-11-01

    The East Asian monsoon is generally regarded to have initiated at the transition from the Late Oligocene to the Early Miocene. However, little is known about this process because of a lack of continuous strata across the boundary between the Late Oligocene and the Early Miocene in Asia. Based on previous drilling (core HZ-1) in the Miocene sediments in the southern Linxia Basin in NW China, we drilled a new 620 m core (HZ-2) into the Late Oligocene strata and obtained 206 m of continuous new core. The detailed paleomagnetism of the new core reveals eleven pairs of normal and reversed polarity zones that can be readily correlated with chrons 6Bn-9n of the geomagnetic polarity time scale (GPTS), define an age interval of 21.6-26.5 Ma and indicate continuity from the Late Oligocene to Early Miocene. The core is characterized by the remarkable occurrence of brownish-red paleosols of luvic cambisols (brown to luvic drab soils) above reddish-brown floodplain siltstones and mudstones, which suggest that the East Asian monsoon likely began by 26.5 Ma. In contrast to the siltstone and mudstone of the Late Oligocene strata, the Miocene strata begin with a thick fine sandstone bed, which marks sudden increases in erosion and loading that most likely reflect a response to tectonic uplift. The hematite content and redness index records of the core further demonstrate that the monsoonal climate in the Late Oligocene to Early Miocene in this area was mainly controlled by global temperature trends and events.

  13. Large-scale recording of neuronal ensembles.

    PubMed

    Buzsáki, György

    2004-05-01

    How does the brain orchestrate perceptions, thoughts and actions from the spiking activity of its neurons? Early single-neuron recording research treated spike pattern variability as noise that needed to be averaged out to reveal the brain's representation of invariant input. Another view is that variability of spikes is centrally coordinated and that this brain-generated ensemble pattern in cortical structures is itself a potential source of cognition. Large-scale recordings from neuronal ensembles now offer the opportunity to test these competing theoretical frameworks. Currently, wire and micro-machined silicon electrode arrays can record from large numbers of neurons and monitor local neural circuits at work. Achieving the full potential of massively parallel neuronal recordings, however, will require further development of the neuron-electrode interface, automated and efficient spike-sorting algorithms for effective isolation and identification of single neurons, and new mathematical insights for the analysis of network properties.

  14. Implementation and adoption of nationwide electronic health records in secondary care in England: final qualitative results from prospective national evaluation in “early adopter” hospitals

    PubMed Central

    Cornford, Tony; Barber, Nicholas; Avery, Anthony; Takian, Amirhossein; Lichtner, Valentina; Petrakaki, Dimitra; Crowe, Sarah; Marsden, Kate; Robertson, Ann; Morrison, Zoe; Klecun, Ela; Prescott, Robin; Quinn, Casey; Jani, Yogini; Ficociello, Maryam; Voutsina, Katerina; Paton, James; Fernando, Bernard; Jacklin, Ann; Cresswell, Kathrin

    2011-01-01

    Objectives To evaluate the implementation and adoption of the NHS detailed care records service in “early adopter” hospitals in England. Design Theoretically informed, longitudinal qualitative evaluation based on case studies. Setting 12 “early adopter” NHS acute hospitals and specialist care settings studied over two and a half years. Data sources Data were collected through in depth interviews, observations, and relevant documents relating directly to case study sites and to wider national developments that were perceived to impact on the implementation strategy. Data were thematically analysed, initially within and then across cases. The dataset consisted of 431 semistructured interviews with key stakeholders, including hospital staff, developers, and governmental stakeholders; 590 hours of observations of strategic meetings and use of the software in context; 334 sets of notes from observations, researchers’ field notes, and notes from national conferences; 809 NHS documents; and 58 regional and national documents. Results Implementation has proceeded more slowly, with a narrower scope and substantially less clinical functionality than was originally planned. The national strategy had considerable local consequences (summarised under five key themes), and wider national developments impacted heavily on implementation and adoption. More specifically, delays related to unrealistic expectations about the capabilities of systems; the time needed to build, configure, and customise the software; the work needed to ensure that systems were supporting provision of care; and the needs of end users for training and support. Other factors hampering progress included the changing milieu of NHS policy and priorities; repeatedly renegotiated national contracts; different stages of development of diverse NHS care records service systems; and a complex communication process between different stakeholders, along with contractual arrangements that largely excluded NHS

  15. A new model for early Earth: heat-pipe cooling

    NASA Astrophysics Data System (ADS)

    Webb, A. G.; Moore, W. B.

    2013-12-01

    In the study of heat transport and lithospheric dynamics of early Earth, current models depend upon plate tectonic and vertical tectonic concepts. Plate tectonic models adequately account for regions with diverse lithologies juxtaposed along ancient shear zones, as seen at the famous Eoarchean Isua supracrustal belt of West Greenland. Vertical tectonic models to date have involved volcanism, sub- and intra-lithospheric diapirism, and sagduction, and can explain the geology of the best-preserved low-grade ancient terranes, such as the Paleoarchean Barberton and Pilbara greenstone belts. However, these models do not offer a globally-complete framework consistent with the geologic record. Plate tectonics models suggest that paired metamorphic belts and passive margins are among the most likely features to be preserved, but the early rock record shows no evidence of these terranes. Existing vertical tectonics models account for the >300 million years of semi-continuous volcanism and diapirism at Barberton and Pilbara, but when they explain the shearing record at Isua, they typically invoke some horizontal motion that cannot be differentiated from plate motion and is not a salient feature of the lengthy Barberton and Pilbara records. Despite the strengths of these models, substantial uncertainty remains about how early Earth evolved from magma ocean to plate tectonics. We have developed a new model, based on numerical simulations and analysis of the geologic record, that provides a coherent, global geodynamic framework for Earth's evolution from magma ocean to subduction tectonics. We hypothesize that heat-pipe cooling offers a viable mechanism for the lithospheric dynamics of early Earth. Our numerical simulations of heat-pipe cooling on early Earth indicate that a cold, thick, single-plate lithosphere developed as a result of frequent volcanic eruptions that advected surface materials downward. The constant resurfacing and downward advection caused compression as the

  16. Seeking new potential in the early-late Permian Gharif Play, West Central Oman

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Guit, F.; Al-Lawati, M.; Nederlof, P.

    1995-08-01

    West Central Oman is a relatively underexplored area where the hydrocarbons found to date occur mainly within the Early-Late Permian Gharif Formation. Structural definition of the low relief closures is hampered by seismic velocity variations caused by dune terrain. Recent exploration activity resulted in several Gharif discoveries, but highlighted reservoir distribution problems. The Gharif Formation, which consists of fluvio-marine sediments, conformably overlies the glacio-lacustine sediments of the Early Permian Al Khlata Formation. It is overlain by shallow marine carbonates of the Late Permian Khuff Formation, the main regional seal. The area is located distally from the main sediment sources tomore » the east. Reservoir development and lateral continuity are seen as the main risk. Most reservoirs are beyond seismic resolution, only the stacked sandstones of the incised valley fills could provide sufficient acoustic contrast to be recognized on seismic. Geochemical typing indicates that the hydrocarbons in the Gharif can be grouped in two main families: the Huqf and Q-hydrocarbons, which are believed to originate from Cambrian to Precambrian source rocks. Although the two hydrocarbon families are sometimes found in one well, they have very different spatial distributions. The Q-oils form continuous strings of accumulations below the main regional seal, whereas the Huqf hydrocarbons occur scattered throughout the area. Mixed accumulations are found where cross-faults or salt domes intercept a Q-oil fairway. Future exploration activities will be guided by refined sedimentological, stratigraphical and hydrocarbon migration models and by the continued efforts to recognize incised valley fills on seismic.« less

  17. Penetrative trace fossils from the late Ediacaran of Mongolia: early onset of the agronomic revolution.

    PubMed

    Oji, Tatsuo; Dornbos, Stephen Q; Yada, Keigo; Hasegawa, Hitoshi; Gonchigdorj, Sersmaa; Mochizuki, Takafumi; Takayanagi, Hideko; Iryu, Yasufumi

    2018-02-01

    The Cambrian radiation of complex animals includes a dramatic increase in the depth and intensity of bioturbation in seafloor sediment known as the 'agronomic revolution'. This bioturbation transition was coupled with a shift in dominant trace fossil style from horizontal surficial traces in the late Precambrian to vertically penetrative trace fossils in the Cambrian. Here we show the existence of the first vertically penetrative trace fossils from the latest Ediacaran: dense occurrences of the U-shaped trace fossil Arenicolites from late Precambrian marine carbonates of Western Mongolia. Their Ediacaran age is established through stable carbon isotope chemostratigraphy and their occurrence stratigraphically below the first appearance of the trace fossil Treptichnus pedum . These Arenicolites are large in diameter, penetrate down to at least 4 cm into the sediment, and were presumably formed by the activity of bilaterian animals. They are preserved commonly as paired circular openings on bedding planes with maximum diameters ranging up to almost 1 cm, and as U- and J-shaped tubes in vertical sections of beds. Discovery of these complex penetrative trace fossils demonstrates that the agronomic revolution started earlier than previously considered.

  18. Penetrative trace fossils from the late Ediacaran of Mongolia: early onset of the agronomic revolution

    PubMed Central

    Dornbos, Stephen Q.; Yada, Keigo; Hasegawa, Hitoshi; Gonchigdorj, Sersmaa; Mochizuki, Takafumi; Takayanagi, Hideko; Iryu, Yasufumi

    2018-01-01

    The Cambrian radiation of complex animals includes a dramatic increase in the depth and intensity of bioturbation in seafloor sediment known as the ‘agronomic revolution’. This bioturbation transition was coupled with a shift in dominant trace fossil style from horizontal surficial traces in the late Precambrian to vertically penetrative trace fossils in the Cambrian. Here we show the existence of the first vertically penetrative trace fossils from the latest Ediacaran: dense occurrences of the U-shaped trace fossil Arenicolites from late Precambrian marine carbonates of Western Mongolia. Their Ediacaran age is established through stable carbon isotope chemostratigraphy and their occurrence stratigraphically below the first appearance of the trace fossil Treptichnus pedum. These Arenicolites are large in diameter, penetrate down to at least 4 cm into the sediment, and were presumably formed by the activity of bilaterian animals. They are preserved commonly as paired circular openings on bedding planes with maximum diameters ranging up to almost 1 cm, and as U- and J-shaped tubes in vertical sections of beds. Discovery of these complex penetrative trace fossils demonstrates that the agronomic revolution started earlier than previously considered. PMID:29515908

  19. Penetrative trace fossils from the late Ediacaran of Mongolia: early onset of the agronomic revolution

    NASA Astrophysics Data System (ADS)

    Oji, Tatsuo; Dornbos, Stephen Q.; Yada, Keigo; Hasegawa, Hitoshi; Gonchigdorj, Sersmaa; Mochizuki, Takafumi; Takayanagi, Hideko; Iryu, Yasufumi

    2018-02-01

    The Cambrian radiation of complex animals includes a dramatic increase in the depth and intensity of bioturbation in seafloor sediment known as the `agronomic revolution'. This bioturbation transition was coupled with a shift in dominant trace fossil style from horizontal surficial traces in the late Precambrian to vertically penetrative trace fossils in the Cambrian. Here we show the existence of the first vertically penetrative trace fossils from the latest Ediacaran: dense occurrences of the U-shaped trace fossil Arenicolites from late Precambrian marine carbonates of Western Mongolia. Their Ediacaran age is established through stable carbon isotope chemostratigraphy and their occurrence stratigraphically below the first appearance of the trace fossil Treptichnus pedum. These Arenicolites are large in diameter, penetrate down to at least 4 cm into the sediment, and were presumably formed by the activity of bilaterian animals. They are preserved commonly as paired circular openings on bedding planes with maximum diameters ranging up to almost 1 cm, and as U- and J-shaped tubes in vertical sections of beds. Discovery of these complex penetrative trace fossils demonstrates that the agronomic revolution started earlier than previously considered.

  20. Magnitude Estimation for Large Earthquakes from Borehole Recordings

    NASA Astrophysics Data System (ADS)

    Eshaghi, A.; Tiampo, K. F.; Ghofrani, H.; Atkinson, G.

    2012-12-01

    We present a simple and fast method for magnitude determination technique for earthquake and tsunami early warning systems based on strong ground motion prediction equations (GMPEs) in Japan. This method incorporates borehole strong motion records provided by the Kiban Kyoshin network (KiK-net) stations. We analyzed strong ground motion data from large magnitude earthquakes (5.0 ≤ M ≤ 8.1) with focal depths < 50 km and epicentral distances of up to 400 km from 1996 to 2010. Using both peak ground acceleration (PGA) and peak ground velocity (PGV) we derived GMPEs in Japan. These GMPEs are used as the basis for regional magnitude determination. Predicted magnitudes from PGA values (Mpga) and predicted magnitudes from PGV values (Mpgv) were defined. Mpga and Mpgv strongly correlate with the moment magnitude of the event, provided sufficient records for each event are available. The results show that Mpgv has a smaller standard deviation in comparison to Mpga when compared with the estimated magnitudes and provides a more accurate early assessment of earthquake magnitude. We test this new method to estimate the magnitude of the 2011 Tohoku earthquake and we present the results of this estimation. PGA and PGV from borehole recordings allow us to estimate the magnitude of this event 156 s and 105 s after the earthquake onset, respectively. We demonstrate that the incorporation of borehole strong ground-motion records immediately available after the occurrence of large earthquakes significantly increases the accuracy of earthquake magnitude estimation and the associated improvement in earthquake and tsunami early warning systems performance. Moment magnitude versus predicted magnitude (Mpga and Mpgv).

  1. The Skeleton Forming Proteome of an Early Branching Metazoan: A Molecular Survey of the Biomineralization Components Employed by the Coralline Sponge Vaceletia Sp.

    PubMed Central

    Wörheide, Gert; Jackson, Daniel John

    2015-01-01

    The ability to construct a mineralized skeleton was a major innovation for the Metazoa during their evolution in the late Precambrian/early Cambrian. Porifera (sponges) hold an informative position for efforts aimed at unraveling the origins of this ability because they are widely regarded to be the earliest branching metazoans, and are among the first multi-cellular animals to display the ability to biomineralize in the fossil record. Very few biomineralization associated proteins have been identified in sponges so far, with no transcriptome or proteome scale surveys yet available. In order to understand what genetic repertoire may have been present in the last common ancestor of the Metazoa (LCAM), and that may have contributed to the evolution of the ability to biocalcify, we have studied the skeletal proteome of the coralline demosponge Vaceletia sp. and compare this to other metazoan biomineralizing proteomes. We bring some spatial resolution to this analysis by dividing Vaceletia’s aragonitic calcium carbonate skeleton into “head” and “stalk” regions. With our approach we were able to identify 40 proteins from both the head and stalk regions, with many of these sharing some similarity to previously identified gene products from other organisms. Among these proteins are known biomineralization compounds, such as carbonic anhydrase, spherulin, extracellular matrix proteins and very acidic proteins. This report provides the first proteome scale analysis of a calcified poriferan skeletal proteome, and its composition clearly demonstrates that the LCAM contributed several key enzymes and matrix proteins to its descendants that supported the metazoan ability to biocalcify. However, lineage specific evolution is also likely to have contributed significantly to the ability of disparate metazoan lineages to biocalcify. PMID:26536128

  2. Early scattering of the solar protoplanetary disk recorded in meteoritic chondrules

    PubMed Central

    Marrocchi, Yves; Chaussidon, Marc; Piani, Laurette; Libourel, Guy

    2016-01-01

    Meteoritic chondrules are submillimeter spherules representing the major constituent of nondifferentiated planetesimals formed in the solar protoplanetary disk. The link between the dynamics of the disk and the origin of chondrules remains enigmatic. Collisions between planetesimals formed at different heliocentric distances were frequent early in the evolution of the disk. We show that the presence, in some chondrules, of previously unrecognized magnetites of magmatic origin implies the formation of these chondrules under impact-generated oxidizing conditions. The three oxygen isotopes systematic of magmatic magnetites and silicates can only be explained by invoking an impact between silicate-rich and ice-rich planetesimals. This suggests that these peculiar chondrules are by-products of the early mixing in the disk of populations of planetesimals from the inner and outer solar system. PMID:27419237

  3. The evolutionary and genetic origins of consciousness in the Cambrian Period over 500 million years ago

    PubMed Central

    Feinberg, Todd E.; Mallatt, Jon

    2013-01-01

    Vertebrates evolved in the Cambrian Period before 520 million years ago, but we do not know when or how consciousness arose in the history of the vertebrate brain. Here we propose multiple levels of isomorphic or somatotopic neural representations as an objective marker for sensory consciousness. All extant vertebrates have these, so we deduce that consciousness extends back to the group's origin. The first conscious sense may have been vision. Then vision, coupled with additional sensory systems derived from ectodermal placodes and neural crest, transformed primitive reflexive systems into image forming brains that map and perceive the external world and the body's interior. We posit that the minimum requirement for sensory consciousness and qualia is a brain including a forebrain (but not necessarily a developed cerebral cortex/pallium), midbrain, and hindbrain. This brain must also have (1) hierarchical systems of intercommunicating, isomorphically organized, processing nuclei that extensively integrate the different senses into representations that emerge in upper levels of the neural hierarchy; and (2) a widespread reticular formation that integrates the sensory inputs and contributes to attention, awareness, and neural synchronization. We propose a two-step evolutionary history, in which the optic tectum was the original center of multi-sensory conscious perception (as in fish and amphibians: step 1), followed by a gradual shift of this center to the dorsal pallium or its cerebral cortex (in mammals, reptiles, birds: step 2). We address objections to the hypothesis and call for more studies of fish and amphibians. In our view, the lamprey has all the neural requisites and is likely the simplest extant vertebrate with sensory consciousness and qualia. Genes that pattern the proposed elements of consciousness (isomorphism, neural crest, placodes) have been identified in all vertebrates. Thus, consciousness is in the genes, some of which are already known. PMID

  4. The evolutionary and genetic origins of consciousness in the Cambrian Period over 500 million years ago.

    PubMed

    Feinberg, Todd E; Mallatt, Jon

    2013-01-01

    Vertebrates evolved in the Cambrian Period before 520 million years ago, but we do not know when or how consciousness arose in the history of the vertebrate brain. Here we propose multiple levels of isomorphic or somatotopic neural representations as an objective marker for sensory consciousness. All extant vertebrates have these, so we deduce that consciousness extends back to the group's origin. The first conscious sense may have been vision. Then vision, coupled with additional sensory systems derived from ectodermal placodes and neural crest, transformed primitive reflexive systems into image forming brains that map and perceive the external world and the body's interior. We posit that the minimum requirement for sensory consciousness and qualia is a brain including a forebrain (but not necessarily a developed cerebral cortex/pallium), midbrain, and hindbrain. This brain must also have (1) hierarchical systems of intercommunicating, isomorphically organized, processing nuclei that extensively integrate the different senses into representations that emerge in upper levels of the neural hierarchy; and (2) a widespread reticular formation that integrates the sensory inputs and contributes to attention, awareness, and neural synchronization. We propose a two-step evolutionary history, in which the optic tectum was the original center of multi-sensory conscious perception (as in fish and amphibians: step 1), followed by a gradual shift of this center to the dorsal pallium or its cerebral cortex (in mammals, reptiles, birds: step 2). We address objections to the hypothesis and call for more studies of fish and amphibians. In our view, the lamprey has all the neural requisites and is likely the simplest extant vertebrate with sensory consciousness and qualia. Genes that pattern the proposed elements of consciousness (isomorphism, neural crest, placodes) have been identified in all vertebrates. Thus, consciousness is in the genes, some of which are already known.

  5. Cambrian–Ordovician of the central Appalachians:Correlations and event stratigraphy of carbonate platform andadjacent deep-water deposits

    USGS Publications Warehouse

    Brezinski, David K.; Taylor, John F.; Repetski, John E.; Loch, James D.

    2015-01-01

    deposited within the Pennsylvania and Maryland portion of the Great American Carbonate Bank. From the Early Cambrian (Dyeran) through Late Ordovician (Turinan), the Laurentian paleocontinent was rimmed by an extensive carbonate platform. During this protracted period of time, a succession of carbonate rock, more than two miles thick, was deposited in Maryland and Pennsylvania. These strata are now exposed in the Nittany arch of central Pennsylvania; the Great Valley of Pennsylvania, Maryland, and Virginia; and the Conestoga and Frederick Valleys of eastern Pennsylvania and Maryland. This fi eld trip will visit key outcrops that illustrate the varied depositional styles and environmental settings that prevailed at different times within the Pennsylvania reentrant portion of the Great American Carbonate Bank. In particular, we will contrast the timing and pattern of sedimentation in off-shelf (Frederick Valley), outer-shelf (Great Valley), and inner-shelf (Nittany arch) deposits. The deposition was controlled primarily by eustasy through the Cambrian and Early Ordovician (within the Sauk megasequence), but was strongly infl uenced later by the onset of Taconic orogenesis during deposition of the Tippecanoe megasequence.

  6. The Seungjeongwon Ilgi as a Major Source of Korean Astronomical Records

    NASA Astrophysics Data System (ADS)

    Stephenson, F. Richard

    The importance of early Korean records of supernovae, comets, meteors and aurorae in modern astronomy is well-known. However, the most extensive Korean source of such data, the Seungjeongwon Ilgi (Daily records of the Office of Royal Secretariat), has received relatively little attention among historians of astronomy. Written in Chinese (Hanmun), the Seungjeongwon Ilgi is a day-to-day chronicle of important events. The main emphasis is on matters of court and state, but observations of a wide variety of astronomical phenomena are regularly included. Although maintenance of the chronicle began early in the Joseon Dynasty (AD 1392-1910), due to wars and rebellions only the records from AD 1623 to 1894 now survive. Nevertheless, the remaining text is substantial, containing more than 3,000 chapters. In this paper, the general format of the astronomical records in the Seungjeongwon Ilgi is discussed, together with examples of the various types of celestial observations which this huge compilation contains.

  7. Collision of the Tacheng block with the Mayile-Barleik-Tangbale accretionary complex in Western Junggar, NW China: Implication for Early-Middle Paleozoic architecture of the western Altaids

    NASA Astrophysics Data System (ADS)

    Zhang, Ji'en; Xiao, Wenjiao; Luo, Jun; Chen, Yichao; Windley, Brian F.; Song, Dongfang; Han, Chunming; Safonova, Inna

    2018-06-01

    Western Junggar in NW China, located to the southeast of the Boshchekul-Chingiz (BC) Range and to the north of the Chu-Balkhash-Yili microcontinent (CBY), played a key role in the architectural development of the western Altaids. However, the mutual tectonic relationships have been poorly constrained. In this paper, we present detailed mapping, field structural geology, and geochemical data from the Barleik-Mayile-Tangbale Complex (BMTC) in Western Junggar. The Complex is divisible into Zones I, II and III, which are mainly composed of Cambrian-Silurian rocks. Zone I contains pillow lava, siliceous shale, chert, coral-bearing limestone, sandstone and purple mudstone. Zone II consists of basaltic lava, siliceous shale, chert, sandstone and mudstone. Zone III is characterized by basalt, chert, sandstone and mudstone. These rocks represent imbricated ocean plate stratigraphy, which have been either tectonically juxtaposed by thrusting or form a mélange with a block-in-matrix structure. All these relationships suggest that the BMTC is an Early-Middle Paleozoic accretionary complex in the eastern extension of the BC Range. These Early Paleozoic oceanic rocks were thrust onto Silurian sediments forming imbricate thrust stacks that are unconformably overlain by Devonian limestone, conglomerate and sandstone containing fossils of brachiopoda, crinoidea, bryozoa, and plant stems and leaves. The tectonic vergence of overturned folds in cherts, drag-related curved cleavages and σ-type structures on the main thrust surface suggests top-to-the-NW transport. Moreover, the positive εNd(t) values of volcanic rocks from the Tacan-1 drill-core, and the positive εHf(t) values and post-Cambrian ages of detrital zircons from Silurian and Devonian strata to the south of the Tacheng block indicate that its basement is a depleted and juvenile lithosphere. And there was a radial outward transition from coral-bearing shallow marine (shelf) to deep ocean (pelagic) environments, and from

  8. Differentiating pedogenesis from diagenesis in early terrestrial paleoweathering surfaces formed on granitic composition parent materials

    USGS Publications Warehouse

    Driese, S.G.; Medaris, L.G.; Ren, M.; Runkel, Anthony C.; Langford, R.P.

    2007-01-01

    Unconformable surfaces separating Precambrian crystalline basement and overlying Proterozoic to Cambrian sedimentary rocks provide an exceptional opportunity to examine the role of primitive soil ecosystems in weathering and resultant formation of saprolite (weathered rock retaining rock structure) and regolith (weathered rock without rock structure), but many appear to have been affected by burial diagenesis and hydrothermal fluid flow, leading some researchers to discount their suitability for such studies. We examine one modern weathering profile (Cecil series), four Cambrian paleoweathering profiles from the North American craton (Squaw Creek, Franklin Mountains, Core SQ-8, and Core 4), one Neoproterozoic profile (Sheigra), and one late Paleoproterozoic profile (Baraboo), to test the hypothesis that these paleoweathering profiles do provide evidence of primitive terrestrial weathering despite their diagenetic and hydrothermal overprinting, especially additions of potassium. We employ an integrated approach using (1) detailed thin-section investigations to identify characteristic pedogenic features associated with saprolitization and formation of well-drained regoliths, (2) electron microprobe analysis to identify specific weathered and new mineral phases, and (3) geochemical mass balance techniques to characterize volume changes during weathering and elemental gains and losses of major and minor elements relative to the inferred parent materials. There is strong pedogenic evidence of paleoweathering, such as clay illuviation, sepic-plasmic fabrics, redoximorphic features, and dissolution and alteration of feldspars and mafic minerals to kaolinite, gibbsite, and Fe oxides, as well as geochemical evidence, such as whole-rock losses of Na, Ca, Mg, Si, Sr, Fe, and Mn greater than in modern profiles. Evidence of diagenesis includes net additions of K, Ba, and Rb determined through geochemical mass balance, K-feldspar overgrowths in overlying sandstone sections, and

  9. Early Childhood Numeracy in a Multiage Setting

    ERIC Educational Resources Information Center

    Wood, Karen; Frid, Sandra

    2005-01-01

    This research is a case study examining numeracy teaching and learning practices in an early childhood multiage setting with Pre-Primary to Year 2 children. Data were collected via running records, researcher reflection notes, and video and audio recordings. Video and audio transcripts were analysed using a mathematical discourse and social…

  10. A Diploblastic Radiate Animal at the Dawn of Cambrian Diversification with a Simple Body Plan: Distinct from Cnidaria?

    PubMed Central

    Yasui, Kinya; Reimer, James D.; Liu, Yunhuan; Yao, Xiaoyong; Kubo, Daisuke; Shu, Degan; Li, Yong

    2013-01-01

    Background Microfossils of the genus Punctatus include developmental stages such as blastula, gastrula, and hatchlings, and represent the most complete developmental sequence of animals available from the earliest Cambrian. Despite the extremely well-preserved specimens, the evolutionary position of Punctatus has relied only on their conical remains and they have been tentatively assigned to cnidarians. We present a new interpretation of the Punctatus body plan based on the developmental reconstruction aided by recent advances in developmental biology. Results Punctatus developed from a rather large egg, gastrulated in a mode of invagination from a coeloblastura, and then formed a mouth directly from the blastopore. Spiny benthic hatchlings were distinguishable from swimming or crawling ciliate larvae found in cnidarians and sponges. A mouth appeared at the perihatching embryonic stage and was renewed periodically during growth, and old mouths transformed into the body wall, thus elongating the body. Growing animals retained a small blind gut in a large body cavity without partitioning by septa and did not form tentacles, pedal discs or holdfasts externally. A growth center at the oral pole was sufficient for body patterning throughout life, and the body patterning did not show any bias from radial symmetry. Conclusions Contrary to proposed cnidarian affinity, the Punctatus body plan has basic differences from that of cnidarians, especially concerning a spacious body cavity separating ectoderm from endoderm. The lack of many basic cnidarian characters in the body patterning of Punctatus leads us to consider its own taxonomic group, potentially outside of Cnidaria. PMID:23840375

  11. Chorioamnionitis and Culture-Confirmed, Early-Onset Neonatal Infections

    PubMed Central

    Hansen, Nellie I.; Schrag, Stephanie J.; Hale, Ellen; Van Meurs, Krisa; Sánchez, Pablo J.; Cantey, Joseph B.; Faix, Roger; Poindexter, Brenda; Goldberg, Ronald; Bizzarro, Matthew; Frantz, Ivan; Das, Abhik; Benitz, William E.; Shane, Andi L.; Higgins, Rosemary; Stoll, Barbara J.

    2016-01-01

    BACKGROUND: Current guidelines for prevention of neonatal group B streptococcal disease recommend diagnostic evaluations and empirical antibiotic therapy for well-appearing, chorioamnionitis-exposed newborns. Some clinicians question these recommendations, citing the decline in early-onset group B streptococcal disease rates since widespread intrapartum antibiotic prophylaxis implementation and potential antibiotic risks. We aimed to determine whether chorioamnionitis-exposed newborns with culture-confirmed, early-onset infections can be asymptomatic at birth. METHODS: Multicenter, prospective surveillance for early-onset neonatal infections was conducted during 2006–2009. Early-onset infection was defined as isolation of a pathogen from blood or cerebrospinal fluid collected ≤72 hours after birth. Maternal chorioamnionitis was defined by clinical diagnosis in the medical record or by histologic diagnosis by placental pathology. Hospital records of newborns with early-onset infections born to mothers with chorioamnionitis were reviewed retrospectively to determine symptom onset. RESULTS: Early-onset infections were diagnosed in 389 of 396 586 live births, including 232 (60%) chorioamnionitis-exposed newborns. Records for 229 were reviewed; 29 (13%) had no documented symptoms within 6 hours of birth, including 21 (9%) who remained asymptomatic at 72 hours. Intrapartum antibiotic prophylaxis exposure did not differ significantly between asymptomatic and symptomatic infants (76% vs 69%; P = .52). Assuming complete guideline implementation, we estimated that 60 to 1400 newborns would receive diagnostic evaluations and antibiotics for each infected asymptomatic newborn, depending on chorioamnionitis prevalence. CONCLUSIONS: Some infants born to mothers with chorioamnionitis may have no signs of sepsis at birth despite having culture-confirmed infections. Implementation of current clinical guidelines may result in early diagnosis, but large numbers of uninfected

  12. Late Eocene to early Oligocene quantitative paleotemperature record: Evidence from continental halite fluid inclusions

    PubMed Central

    Zhao, Yan-jun; Zhang, Hua; Liu, Cheng-lin; Liu, Bao-kun; Ma, Li-chun; Wang, Li-cheng

    2014-01-01

    Climate changes within Cenozoic extreme climate events such as the Paleocene–Eocene Thermal Maximum and the First Oligocene Glacial provide good opportunities to estimate the global climate trends in our present and future life. However, quantitative paleotemperatures data for Cenozoic climatic reconstruction are still lacking, hindering a better understanding of the past and future climate conditions. In this contribution, quantitative paleotemperatures were determined by fluid inclusion homogenization temperature (Th) data from continental halite of the first member of the Shahejie Formation (SF1; probably late Eocene to early Oligocene) in Bohai Bay Basin, North China. The primary textures of the SF1 halite typified by cumulate and chevron halite suggest halite deposited in a shallow saline water and halite Th can serve as an temperature proxy. In total, one-hundred-twenty-one Th data from primary and single-phase aqueous fluid inclusions with different depths were acquired by the cooling nucleation method. The results show that all Th range from 17.7°C to 50.7°C,with the maximum homogenization temperatures (ThMAX) of 50.5°C at the depth of 3028.04 m and 50.7°C at 3188.61 m, respectively. Both the ThMAX presented here are significantly higher than the highest temperature recorded in this region since 1954and agree with global temperature models for the year 2100 predicted by the Intergovernmental Panel on Climate Change. PMID:25047483

  13. Records of our Early Biosphere Illuminate our Origins and Guide our Search for Life Beyond Earth

    NASA Technical Reports Server (NTRS)

    DesMarais, David J.

    2003-01-01

    A scientific "mission of exploration to early Earth" will help us chart the distribution of life elsewhere. We must discriminate between attributes of biospheres that are universal versus those attributes that represent principally the outcomes of long-term survival specifically on Earth. In addition to the basic physics and chemistry of matter, the geologic evolution of rocky habitable planets and their climates might be similar elsewhere in the Universe. Certain key agents that drive long-term environmental change (e.g., stellar evolution, impacts, geothermal heat flow, tectonics, etc.) can help us to reconstruct ancient climates and to compare their evolution among populations of Earth- like planets. Early Earth was tectonically more active than today and therefore it exhaled reduced chemical species into the more oxidized surface environment at greater rates. This tectonic activity thus sustained oxidation-reduction reactions that provided the basis for the development of biochemical pathways that harvest chemical energy ("bioenergetics"). Most examples of bioenergetics today that extract energy by reacting oxidized and reduced chemicals in the environment were likely more pervasive among our microbial ancestors than are the presently known examples of photosynthesis. The geologic rock record indicates that, as early as 3.5 billion years ago (3.5 Ga), microbial biofilms were widespread within the coastal environments of small continents and tectonically unstable volcanic islands. Non oxygen-producing (non-oxygenic) photosynthesis preceded oxygenic photosynthesis, but all types of photosynthesis contributed substantially to the long-term increase in global primary biological productivity. Evidence of photosynthesis is tentative by 3.5 Ga and compelling by 2.7 Ga. Evidence of oxygenic photosynthesis is strong by 2.7 Ga and compelling by 2.3 Ga. These successive innovations transformed life from local communities that survived principally by catalyzing chemical

  14. A missing link in the evolution of the cumulative recorder.

    PubMed

    Asano, Toshio; Lattal, Kennon A

    2012-09-01

    A recently recovered cumulative recorder provides a missing link in the evolution of the cumulative recorder from a modified kymograph to a reliably operating, scientifically and commercially successful instrument. The recorder, the only physical evidence of such an early precommercial cumulative recorder yet found, was sent to Keio University in Tokyo, Japan, in 1952 at the behest of B. F. Skinner at Harvard University. Last used in research in the late 1960s, the cumulative recorder remained locked in a storage room until 2007, when it was found again. A historical context for the recorder is followed by a description of the recorder and a comparison between it and the commercially successful Gerbrands Model C-1 recorder. Labeled the Keio recorder, it is a testament to Skinner's persistence in developing a reliable means of quantifying the behavior of living organisms in real time.

  15. New Zircon U-Pb Age Constrain of the Origin of Devil's River Uplift (SW Texas) and Insights into the Late Proterozoic and Paleozoic Evolution of the Southern Margin of Laurentia

    NASA Astrophysics Data System (ADS)

    Rodriguez, E.; Dickerson, P. W.; Stockli, D. F.

    2017-12-01

    The Devils River Uplift (DRU) in SW Texas records the evolution of the southern Laurentian margin from Grenvillian orogenesis and assembly of Rodinia, to its fragmentation by rifting, and to the amalgamation of Pangaea. It was cored by a well (Shell No. 1 Stewart), penetrating Precambrian gneisses and Cambrian metasediments and sandstones. New zircon LA-ICP-MS data from a total of 10 samples elucidate the crystallization and depositional ages, as well as the detrital provenance, of Precambrian and Cambrian rocks from the DRU. Zircons from five Precambrian crystalline basement samples (6000-9693') yield uniform U-Pb crystallization ages of 1230 Ma that are similar to ages for young gneisses of the Valley Spring Domain (Llano uplift) in central Texas, where they mark the cessation of arc magmatism within the Grenville orogenic belt. The 1230 Ma igneous basement is overlain by L.-M. Cambrian metasedimentary rocks ( 4000-6000') with maximum depositional ages of 533-545 Ma. Detrital zircons from Cambrian strata are dominated by a 1070-1080 Ma population, likely derived from basement units exposed in Texas (Llano uplift, Franklin Mts.), with minor contributions from local 1230 Ma Precambrian basement and the 1380-1500 Ma Granite Rhyolite Province. The L.-M. Cambrian interval is dominated (>80%) by Neoproterozoic detrital magmatic zircons with two major distinct age clusters at 570-700 Ma and 780-820 Ma, supporting a two-stage Rodinia rift model and providing strong evidence for major Cryogenian-Eocambrian intraplate magmatism along the southern margin of Rodinia. Moreover, detrital zircon signatures for L.-M. and U. Cambrian strata strongly correlate with those from the Cuyania terrane of W. Argentina - notably the W. Sierras Pampeanas (Sa. Pie de Palo, Sa. de Maz): 1230 Ma from metasandstones (PdP); 1081-1038 Ma from metasiliciclastics (PdP, SdM); Cryogenian-Eocambrian [774 & 570 Ma] plutons (SdM, PdP). In summary, these new zircon U-Pb data from DRU in SW Texas show

  16. Do Jack Hills Detrital Zircons Contain Records of the Early Geodynamo?

    NASA Astrophysics Data System (ADS)

    Weiss, B. P.; Maloof, A. C.; Tailby, N. D.; Ramezani, J.; Fu, R. R.; Glenn, D. R.; Kehayias, P.; Walsworth, R. L.; Hanus, V.; Trail, D.; Watson, E. B.; Harrison, T. M.; Bowring, S. A.; Kirschvink, J. L.; Swanson-Hysell, N.; Coe, R. S.; Einsle, J. F.; Harrison, R. J.

    2015-12-01

    It is unknown when Earth's dynamo magnetic field originated. With crystallization ages ranging from 3.0-4.38 Ga, detrital zircon crystals found in the Jack Hills of Western Australia might preserve a record of the missing first billion years of Earth's magnetic field history. Recently, Tarduno et al. (2015) argued that magnetization in Jack Hills zircons provides evidence for a substantial geomagnetic field dating back to their U/Pb formation ages (3.3 and 4.2 Ga). However, the identification of such ancient field records requires establishing that the zircons have avoided remagnetization since their formation. At a minimum, it should be demonstrated that they have not been remagnetized since being deposited at ~3.0 Ga. To establish the timing and intensity of the metamorphic and alteration events experienced by the zircon, we conducted 12 paleomagnetic field tests in combination with U-Pb geochronology on their host rocks (see Weiss et al. 2015, EPSL). Our data show that the Hadean zircon-bearing rocks and surrounding region have been pervasively remagnetized, with the final major overprinting likely from emplacement of the Warakurna large igneous province at 1.1 Ga (see Figure). Even if some Jack Hills zircons do record a pre-depositional magnetization, they still could have been remagnetized sometime during the 1.4 Gy between their crystallization and their deposition. First, the temperatures capable of remagnetizing magnetite inclusions are well below those that could reset a U-Pb date or result in significant discordance. Therefore, thermal events capable of completely remagnetizing Jack Hills zircons could be undetected by the techniques reported by Tarduno at al. (2015). Second, the zircons' magnetization might be dominated by secondary ferromagnetic inclusions or contamination. To address the latter possibility, we are conducting electron microscopy, x-ray tomography, and magnetic field mapping on the zircons. Our initial quantum diamond magnetometry high

  17. Classroom Technology Rollouts

    ERIC Educational Resources Information Center

    Blow, Deborah; McConnell, Sandy

    2012-01-01

    At Cambrian School District, educators are trying a new approach as they investigate new technologies. Rather than roll out a one-size-fits-all mandate that one will use this new "box," which may also sit gathering dust, they have implemented a new model. Early in the last school year they announced the Digital Media Academy, and in…

  18. The oldest African bat from the early Eocene of El Kohol (Algeria).

    PubMed

    Ravel, Anthony; Marivaux, Laurent; Tabuce, Rodolphe; Adaci, Mohammed; Mahboubi, Mohammed; Mebrouk, Fateh; Bensalah, Mustapha

    2011-05-01

    The Afro-Arabian Paleogene fossil record of Chiroptera is very poor. In North Africa and Arabia, this record is limited, thus far, to a few localities mainly in Tunisia (Chambi, late early Eocene), Egypt (Fayum, late Eocene to early Oligocene), and Sultanate of Oman (Taqah, early Oligocene). It consists primarily of isolated teeth or mandible fragments. Interestingly, these African fossil bats document two modern groups (Vespertilionoidea and Rhinolophoidea) from the early Eocene, while the bat fossil record of the same epoch of North America, Eurasia, and Australia principally includes members of the "Eochiroptera." This paraphyletic group contains all primitive microbats excluding modern families. In Algeria, the region of Brezina, southeast of the Atlas Mountains, is famous for the early Eocene El Kohol Formation, which has yielded one of the earliest mammalian faunas of the African landmass. Recent fieldwork in the same area has led to the discovery of a new vertebrate locality, including isolated teeth of Chiroptera. These fossils represent the oldest occurrence of Chiroptera in Africa, thus extending back the record of the group to the middle early Eocene (Ypresian) on that continent. The material consists of an upper molar and two fragments of lower molars. The dental character association matches that of "Eochiroptera." As such, although very fragmentary, the material testifies to the first occurrence of "Eochiroptera" in Algeria, and by extension in Africa. This discovery demonstrates that this basal group of Chiroptera had a worldwide distribution during the early Paleogene.

  19. The oldest African bat from the early Eocene of El Kohol (Algeria)

    NASA Astrophysics Data System (ADS)

    Ravel, Anthony; Marivaux, Laurent; Tabuce, Rodolphe; Adaci, Mohammed; Mahboubi, Mohammed; Mebrouk, Fateh; Bensalah, Mustapha

    2011-05-01

    The Afro-Arabian Paleogene fossil record of Chiroptera is very poor. In North Africa and Arabia, this record is limited, thus far, to a few localities mainly in Tunisia (Chambi, late early Eocene), Egypt (Fayum, late Eocene to early Oligocene), and Sultanate of Oman (Taqah, early Oligocene). It consists primarily of isolated teeth or mandible fragments. Interestingly, these African fossil bats document two modern groups (Vespertilionoidea and Rhinolophoidea) from the early Eocene, while the bat fossil record of the same epoch of North America, Eurasia, and Australia principally includes members of the "Eochiroptera." This paraphyletic group contains all primitive microbats excluding modern families. In Algeria, the region of Brezina, southeast of the Atlas Mountains, is famous for the early Eocene El Kohol Formation, which has yielded one of the earliest mammalian faunas of the African landmass. Recent fieldwork in the same area has led to the discovery of a new vertebrate locality, including isolated teeth of Chiroptera. These fossils represent the oldest occurrence of Chiroptera in Africa, thus extending back the record of the group to the middle early Eocene (Ypresian) on that continent. The material consists of an upper molar and two fragments of lower molars. The dental character association matches that of "Eochiroptera." As such, although very fragmentary, the material testifies to the first occurrence of "Eochiroptera" in Algeria, and by extension in Africa. This discovery demonstrates that this basal group of Chiroptera had a worldwide distribution during the early Paleogene.

  20. Phase III of Early Restoration | NOAA Gulf Spill Restoration

    Science.gov Websites

    information about this phase of Early Restoration, including fact sheets on each project. The final Phase III 44 projects are documented in a final Record of Decision. Information about Phase III of Early Archive Home Phase III of Early Restoration Phase III of Early Restoration Beach habitat would be restored

  1. Sequence stratigraphy of the ANDRILL Southern McMurdo Sound (SMS) project drillcore, Antarctica: an expanded, near-field record of Antarctic Early to Middle Miocene climate and relative sea-level change

    NASA Astrophysics Data System (ADS)

    Fielding, C. R.; Browne, G. H.; Field, B.; Florindo, F.; Harwood, D. M.; Krissek, L. A.; Levy, R. H.; Panter, K.; Passchier, S.; Pekar, S. F.; SMS Science Team

    2008-12-01

    Present understanding of Antarctic climate change during the Early to Middle Miocene, including definition of major cycles of glacial expansion and contraction, relies in large part on stable isotope proxy records from Ocean Drilling Program cores. Here, we present a sequence stratigraphic analysis of the Southern McMurdo Sound drillcore (AND-2A), which was acquired during the Austral Spring of 2007. This core offers a hitherto unavailable ice-proximal stratigraphic archive of the Early to Middle Miocene from a high-accommodation Antarctic continental margin setting, and provides clear evidence of repeated fluctuations in climate, ice expansion/contraction and attendant sea-level change over the period 20-14 Ma, with a more fragmentary record of the post-14 Ma period. A succession of seventy sequences is recognized, each bounded by a significant facies dislocation (sequence boundary), composed internally of deposits of glacimarine to open shallow marine environments, and each typically dominated by the transgressive systems tract. From changes in facies abundances and sequence character, a series of long-term (m.y.) changes in climate and relative sea-level is identified. The lithostratigraphy can be correlated confidently to glacial events Mi1b and Mi2, to the Miocene Climatic Optimum, and to the global eustatic sea-level curve. SMS provides a detailed, direct, ice-proximal reference point from which to evaluate stable isotope proxy records for Neogene Antarctic paleoclimate.

  2. Exploration in Ordovician of central Michigan Basin

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Fisher, J.H.; Barratt, M.W.

    1985-12-01

    Deep wells in the central Michigan basin have provided sufficient data to define two new mappable formations - the Foster Formation and the Bruggers Formation. Recent conodont studies have corrected the age assignments of the strata containing these formations. Previously, the lower section (Foster) was classified as mostly Cambrian, and the upper unit (Bruggers) was identified as Early Ordovician. Conodont identifications indicate an Early and Middle Ordovician age for the Foster Formation and a Middle Ordovician age for the Bruggers Formation. The Michigan basin existed in embryonic form in the Late Cambrian, but the full outline of the present-day basinmore » did not develop until Early Ordovician. Gas and condensate are produced from the Bruggers Formation as deep as 11,252 ft (3429 m). Geothermal investigations suggest that gas production is possible to the base of the Paleozoic section in the central basin (17,000 ft or 5181 m). Paleotemperatures were higher during the Paleozoic owing to 3000-4000 ft (914-1291 m) of additional sedimentary cover. Five wells are producing from the Bruggers Formation. All are deeper tests in anticlines producing from Devonian reservoirs discovered earlier. The structures are the result of vertical movements of basement fault blocks activated by regional stresses. 12 figures, 2 tables.« less

  3. A new record of late Pliocene-early Pleistocene aeolian loess-red clay deposits from the western Chinese Loess Plateau and its palaeoenvironmental implications

    NASA Astrophysics Data System (ADS)

    Zan, Jinbo; Fang, Xiaomin; Zhang, Weilin; Yan, Maodu; Zhang, Dawen

    2018-04-01

    The loess-red clay sequences in northern China provide high-resolution terrestrial records of Asian monsoon evolution and aridification of the Asian interior. To date, however, aeolian deposits of late Pliocene-early Pleistocene age (3.5-2.4 Ma) have only rarely been reported from the western Chinese Loess Plateau (CLP), which significantly hinders our understanding of the distribution of aeolian deposits and the palaeoenvironmental evolution of the region. Here, we present magnetostratigraphic, lithologic and magnetic susceptibility results for two recently-drilled boreholes from the north bank of Baxie River, central Linxia Basin, which are highly correlative with those of the loess-red clay deposits spanning the interval from 3.6 to 2.4 Ma in the eastern CLP. Our results provide the first direct evidence for the occurrence of late Pliocene-early Pleistocene aeolian deposits in the western CLP and provide new insights into the distribution of aeolian deposits in northern China. The spatial coherence of the magnetic susceptibility fluctuations further indicates that magnetic susceptibility is a powerful tool for stratigraphic correlation of late Pliocene aeolian deposits in the western CLP. In addition, our results demonstrate that erosional events may have occurred in the early or middle Pleistocene, and they may provide new insights into the reasons for the absence of loess-red clay deposits from 3.5 to 2.4 Ma in most parts of the western CLP.

  4. Early Tertiary Exhumation, Erosion, and Sedimentation in the Central Andes, NW Argentina

    NASA Astrophysics Data System (ADS)

    Carrapa, B.; Decelles, P. G.; Gerhels, G.; Mortimer, E.; Strecker, M. R.

    2006-12-01

    Timing of deformation and resulting sedimentation patterns in the Altiplano-Puna Plateau-Eastern Cordillera of the southern Central Andes are the subject of ongoing controversial debate. In the Bolivian Altiplano, sedimentation into a foreland basin system commenced during the Paleocene. Farther south in the Puna and Eastern Cordillera of NW Argentina, a lack of data has precluded a similar interpretation. Early Tertiary non-marine sedimentary rocks are preserved within the present day Puna Plateau and Eastern Cordillera of NW Argentina. The Salar de Pastos Grandes basin in the Puna Plateau contains more than 2 km of Eocene alluvial and fluvial strata in the Geste Formation, deposited in close proximity to orogenic source terrains. Sandstone and conglomerate petrographic data document Ordovician quartzites and minor phyllites and schists as the main source rocks. Detrital zircon U-Pb ages from both the Geste Formation and from underlying Ordovician quartzite cluster in the 900-1200 Ma (Grenville) and late Precambrian-Cambrian (Panafrican) ranges. Sparse late Eocene (~37-34 Ma) grains are also present; their large size, euhedral shape, and decreasing mean ages upsection suggest that these grains are volcanogenic (i.e. ash fall contamination), derived from an inferred magmatic arc to the west. The Eocene ages corroborate mammalian paleontological dates, defining the approximate begin of deposition of the Geste Formation. Alternatively, these young zircons could be of plutonic origin; however, no Eocene plutons are present in the surrounding source rocks and this interpretation is not likely. From W to E, fluvial rocks of the Quebrada de los Colorados Formation show similar sedimentological features as those observed for the Geste Formation, suggesting a genetic link between the two. Detrital zircon U-Pb data show mainly Panafrican ages, with sparse ages in the 860-935 Ma range and a few mid-Proterozoic ages. More importantly, a significant number of late Eocene

  5. 470-Million-year-old black corals from China

    NASA Astrophysics Data System (ADS)

    Baliński, Andrzej; Sun, Yuanlin; Dzik, Jerzy

    2012-08-01

    Phosphatic (possibly secondarily phosphatised) remains of antipatharian coralla, previously unknown in the fossil record, occur abundantly in the early Ordovician Fenxiang Formation in the Hubei Province, southern China. Probably two species (and genera) are represented, which differ in spinosity of branches. The more spinose one, Sinopathes reptans, has its lateral spines bearing regular, longitudinally arranged costellae. The early Floian geological age of this finding, about 470 Ma, supports predictions on the timing of anthozoan phylogeny derived from the molecular phylogenetic evidence. Black corals (Antipatharia) are basal to the scleractinians in the Hexacorallia clade, being more derived than sea anemones and the Zoantharia. Based on calibration of the molecular clock with Mesozoic data, the first split of lineages within the scleractinian hexacorals was proposed to take place approximately 425 million years ago. This implies that the origin of Antipatharia should precede this date. They have not been known in the fossil record because of unmineralised skeleton composed primarily of laminar chitin complexed with a protein. Unlike all recent species, the encrusting basal part of the colony dominated in the Ordovician ones and only occasionally erect branches developed, rather chaotically ramified. This presumably plesiomorphic trait seems consistent with ancient geological age and suggests that some problematic fossils from the Late Cambrian may be their, even less-derived, relatives.

  6. Early Paleozoic tectonic reconstruction of Iran: Tales from detrital zircon geochronology

    NASA Astrophysics Data System (ADS)

    Moghadam, Hadi Shafaii; Li, Xian-Hua; Griffin, William L.; Stern, Robert J.; Thomsen, Tonny B.; Meinhold, Guido; Aharipour, Reza; O'Reilly, Suzanne Y.

    2017-01-01

    In this study we use detrital zircons to probe the Early Paleozoic history of NE Iran and evaluate the link between sediment sources and Gondwanan pre-Cadomian, Cadomian and younger events. U-Pb zircon ages and Hf isotopic compositions are reported for detrital zircons from Ordovician and Early Devonian sedimentary rocks from NE Iran. These clastic rocks are dominated by zircons with major age populations at 2.5 Ga, 0.8-0.6 Ga, 0.5 Ga and 0.5-0.4 Ga as well as a minor broad peak at 1.0 Ga. The source of 2.5 Ga detrital zircons is enigmatic; they may have been supplied from the Saharan Metacraton (or West African Craton) to the southwest or Afghanistan-Tarim to the east. The detrital zircons with age populations at 0.8-0.6 Ga probably originated from Cryogenian-Ediacaran juvenile igneous rocks of the Arabian-Nubian Shield; this inference is supported by their juvenile Hf isotopes, although some negative εHf (t) values suggest that other sources (such as the West African Craton) were also involved. The age peak at ca 0.5 Ga correlates with Cadomian magmatism reported from Iranian basement and elsewhere in north Gondwana. The variable εHf (t) values of Cadomian detrital zircons, resembling the εHf (t) values of zircons in magmatic Cadomian rocks from Iran and Taurides (Turkey), suggest an Andean-type margin and the involvement of reworked older crust in the generation of the magmatic rocks. The youngest age population at 0.5-0.4 Ga is interpreted to represent Gondwana rifting and the opening of Paleotethys, which probably started in Late Cambrian-Ordovician time. A combination of U-Pb dating and Hf-isotope data from Iran, Turkey and North Gondwana confirms that Iran and Turkey were parts of Gondwana at least until late Paleozoic time.

  7. Early anaerobic metabolisms

    PubMed Central

    Canfield, Don E; Rosing, Minik T; Bjerrum, Christian

    2006-01-01

    Before the advent of oxygenic photosynthesis, the biosphere was driven by anaerobic metabolisms. We catalogue and quantify the source strengths of the most probable electron donors and electron acceptors that would have been available to fuel early-Earth ecosystems. The most active ecosystems were probably driven by the cycling of H2 and Fe2+ through primary production conducted by anoxygenic phototrophs. Interesting and dynamic ecosystems would have also been driven by the microbial cycling of sulphur and nitrogen species, but their activity levels were probably not so great. Despite the diversity of potential early ecosystems, rates of primary production in the early-Earth anaerobic biosphere were probably well below those rates observed in the marine environment. We shift our attention to the Earth environment at 3.8 Gyr ago, where the earliest marine sediments are preserved. We calculate, consistent with the carbon isotope record and other considerations of the carbon cycle, that marine rates of primary production at this time were probably an order of magnitude (or more) less than today. We conclude that the flux of reduced species to the Earth surface at this time may have been sufficient to drive anaerobic ecosystems of sufficient activity to be consistent with the carbon isotope record. Conversely, an ecosystem based on oxygenic photosynthesis was also possible with complete removal of the oxygen by reaction with reduced species from the mantle. PMID:17008221

  8. Lithium in Brachiopods - proxy for seawater evolution?

    NASA Astrophysics Data System (ADS)

    Gaspers, Natalie; Magna, Tomas; Tomasovych, Adam; Henkel, Daniela

    2017-04-01

    Marine biogenic carbonates have the potential to serve as a proxy for evolution of seawater chemistry. In order to compile a record of the past and recent δ7Li in the oceans, foraminifera shells, scleractinian corals and belemnites have been used. However, only a foraminifera-based record appears to more accurately reflect the Li isotope composition of ocean water. At present, this record is available for the Cenozoic with implications for major events during this period of time, including K/T event [1]. A record for the entire Phanerozoic has not yet been obtained. In order to extend this record to the more distant past, Li elemental/isotope systematics of brachiopods were investigated because these marine animals were already present in Early Cambrian oceans and because they are less sensitive to diagenesis-induced modifications due to their shell mineralogy (low-Mg calcite). The preliminary data indicates a species-, temperature- and salinity-independent behavior of Li isotopes in brachiopod shells. Also, no vital effects have been observed for different shell parts. The consistent offset of -4‰ relative to modern seawater is in accordance with experimental data [2]. Further data are now being collected for Cenozoic specimens to more rigorously test brachiopods as possible archives of past seawater in comparison to the existing foraminiferal records. [1] Misra & Froelich (2012) Science 335, 818-823 [2] Marriott et al. (2004) Chem Geol 212, 5-15

  9. Casebooks in Early Modern England:

    PubMed Central

    Kassell, Lauren

    2014-01-01

    summary Casebooks are the richest sources that we have for encounters between early modern medical practitioners and their patients. This article compares astrological and medical records across two centuries, focused on England, and charts developments in the ways in which practitioners kept records and reflected on their practices. Astrologers had a long history of working from particular moments, stellar configurations, and events to general rules. These practices required systematic notation. Physicians increasingly modeled themselves on Hippocrates, recording details of cases as the basis for reasoned expositions of the histories of disease. Medical records, as other scholars have demonstrated, shaped the production of medical knowledge. Instead, this article focuses on the nature of casebooks as artifacts of the medical encounter. It establishes that casebooks were serial records of practice, akin to diaries, testimonials, and registers; identifies extant English casebooks and the practices that led to their production and preservation; and concludes that the processes of writing, ordering, and preserving medical records are as important for understanding the medical encounter as the records themselves. PMID:25557513

  10. Consensus in a Precambrian garden

    NASA Astrophysics Data System (ADS)

    Maggs, William Ward

    At the Precambrian-Cambrian boundary, the course of life on Earth underwent a dramatic change that culminated in the rise of predators and other complex animals, a group of paleontologists agreed at a conferece last week.Just prior to 590 million years ago, the ecology of life in the oceans was very simple; soft-shelled multicellular animals called Ediacara lived in apparent harmony with vast mats o f bacteria and algae that covered the seafloor, dependent on the photosynthesis or chemosynthesis of their one-celled hosts for their existence. According to the consensus reached by the scientists, this symbiotic and apparently global “Garden of Ediacara” fell early in the Cambrian Period, as the mats declined and food chains multiplied with new animals that, for the first time in Earth's history, preyed on other living things.

  11. Earliest Phanerozoic or latest Proterozoic fossils from the Arabian Shield

    USGS Publications Warehouse

    Cloud, P.; Awramik, S.M.; Morrison, K.; Hadley, D.G.

    1979-01-01

    We report here the first biologically definable fossils from pre-Saq (pre-Middle Cambrian) rocks of the Arabian Shield. They include the distinctive helically coiled tubular filaments of the oscillatorialean blue-green alga Obruchevella parva as well as two size classes of spheroidal unicells of uncertain affinity. Also present is the conical stromatolite Conophyton and unidentified stromatolites. All occur in cherty limestones of the Jubaylah Group, northern Saudi Arabia, a nonmarine to locally marine taphrogeosynclinal sequence that fills depressions along the northwest-trending Najd faults. Conophyton has heretofore been found only in strata older than about 680 Ma (except for puzzling records in modern hot springs) while Obruchevella is so far known only from rocks between about 680 and 470 Ma old. Thus it appears that the Jubaylah Group is close to the Proterozoic-Phanerozoic transition. The simple spheroidal nanno-fossils are not diagnostic as to age. Their relationships within what appears to be early diagenetic chert suggest a classical algal-mat association. The brecciated and microchanneled appearance of much of the fossiliferous rock, its locally dolomitic nature, and the prevalence of cryptalgalaminate favors a very shallow, locally turbulent, and perhaps episodically exposed marine or marginal marine setting. The Jubaylah Group lies unconformably beneath the Siq Sandstone (basal member of the Saq Sandstone) of medial Cambrian age, rests nonconformably on crystalline basement, and has yielded a K-Ar whole-rock age (on andesitic basalt) of ???540 Ma. To judge from the fossils, however, that age may be as much as 100 Ma or more too young. ?? 1979.

  12. Comparison of organic geochemistry and metal enrichment in two black shales: Cambrian Alum Shale of Sweden and Devonian Chattanooga Shale of United States

    USGS Publications Warehouse

    Leventhal, J.S.

    1991-01-01

    In most black shales, such as the Chattanooga Shale and related shales of the eastern interior United States, increased metal and metalloid contents are generally related to increased organic carbon content, decreased sedimentation rate, organic matter type, or position in the basin. In areas where the stratigraphic equivalents of the Chattanooga Shale are deeply buried and and the organic material is thermally mature, metal contents are essentially the same as in unheated areas and correlate with organic C or S contents. This paradigm does not hold for the Cambrian Alum Shale Formation of Sweden where increased metal content does not necessarily correlate with organic matter content nor is metal enrichment necessarily related to land derived humic material because this organic matter is all of marine source. In southcentral Sweden the elements U, Mo, V, Ni, Zn, Cd and Pb are all enriched relative to average black shales but only U and Mo correlate to organic matter content. Tectonically disturbed and metamorphosed allochthonous samples of Alum Shale on the Caledonian front in western Sweden have even higher amounts for some metals (V, Ni, Zn and Ba) relative to the autochthonous shales in this area and those in southern Sweden. ?? 1991 Springer-Verlag.

  13. A reassessment of the early archaeological record at Leang Burung 2, a Late Pleistocene rock-shelter site on the Indonesian island of Sulawesi

    PubMed Central

    Hakim, Budianto; Ramli, Muhammad; Aubert, Maxime; van den Bergh, Gerrit D.; Li, Bo; Burhan, Basran; Saiful, Andi Muhammad; Siagian, Linda; Sardi, Ratno; Jusdi, Andi; Abdullah; Mubarak, Andi Pampang; Moore, Mark W.; Roberts, Richard G.; Zhao, Jian-xin; McGahan, David; Jones, Brian G.; Perston, Yinika; Szabó, Katherine; Mahmud, M. Irfan; Westaway, Kira; Jatmiko; Saptomo, E. Wahyu; van der Kaars, Sander; Grün, Rainer; Wood, Rachel; Dodson, John

    2018-01-01

    This paper presents a reassessment of the archaeological record at Leang Burung 2, a key early human occupation site in the Late Pleistocene of Southeast Asia. Excavated originally by Ian Glover in 1975, this limestone rock-shelter in the Maros karsts of Sulawesi, Indonesia, has long held significance in our understanding of early human dispersals into ‘Wallacea’, the vast zone of oceanic islands between continental Asia and Australia. We present new stratigraphic information and dating evidence from Leang Burung 2 collected during the course of our excavations at this site in 2007 and 2011–13. Our findings suggest that the classic Late Pleistocene modern human occupation sequence identified previously at Leang Burung 2, and proposed to span around 31,000 to 19,000 conventional 14C years BP (~35–24 ka cal BP), may actually represent an amalgam of reworked archaeological materials. Sources for cultural materials of mixed ages comprise breccias from the rear wall of the rock-shelter–remnants of older, eroded deposits dated to 35–23 ka cal BP–and cultural remains of early Holocene antiquity. Below the upper levels affected by the mass loss of Late Pleistocene deposits, our deep-trench excavations uncovered evidence for an earlier hominin presence at the site. These findings include fossils of now-extinct proboscideans and other ‘megafauna’ in stratified context, as well as a cobble-based stone artifact technology comparable to that produced by late Middle Pleistocene hominins elsewhere on Sulawesi. PMID:29641524

  14. A reassessment of the early archaeological record at Leang Burung 2, a Late Pleistocene rock-shelter site on the Indonesian island of Sulawesi.

    PubMed

    Brumm, Adam; Hakim, Budianto; Ramli, Muhammad; Aubert, Maxime; van den Bergh, Gerrit D; Li, Bo; Burhan, Basran; Saiful, Andi Muhammad; Siagian, Linda; Sardi, Ratno; Jusdi, Andi; Abdullah; Mubarak, Andi Pampang; Moore, Mark W; Roberts, Richard G; Zhao, Jian-Xin; McGahan, David; Jones, Brian G; Perston, Yinika; Szabó, Katherine; Mahmud, M Irfan; Westaway, Kira; Jatmiko; Saptomo, E Wahyu; van der Kaars, Sander; Grün, Rainer; Wood, Rachel; Dodson, John; Morwood, Michael J

    2018-01-01

    This paper presents a reassessment of the archaeological record at Leang Burung 2, a key early human occupation site in the Late Pleistocene of Southeast Asia. Excavated originally by Ian Glover in 1975, this limestone rock-shelter in the Maros karsts of Sulawesi, Indonesia, has long held significance in our understanding of early human dispersals into 'Wallacea', the vast zone of oceanic islands between continental Asia and Australia. We present new stratigraphic information and dating evidence from Leang Burung 2 collected during the course of our excavations at this site in 2007 and 2011-13. Our findings suggest that the classic Late Pleistocene modern human occupation sequence identified previously at Leang Burung 2, and proposed to span around 31,000 to 19,000 conventional 14C years BP (~35-24 ka cal BP), may actually represent an amalgam of reworked archaeological materials. Sources for cultural materials of mixed ages comprise breccias from the rear wall of the rock-shelter-remnants of older, eroded deposits dated to 35-23 ka cal BP-and cultural remains of early Holocene antiquity. Below the upper levels affected by the mass loss of Late Pleistocene deposits, our deep-trench excavations uncovered evidence for an earlier hominin presence at the site. These findings include fossils of now-extinct proboscideans and other 'megafauna' in stratified context, as well as a cobble-based stone artifact technology comparable to that produced by late Middle Pleistocene hominins elsewhere on Sulawesi.

  15. Early Childhood Education and Care Policy in England under the Coalition Government

    ERIC Educational Resources Information Center

    Lloyd, Eva

    2015-01-01

    This paper reviews developments in policy on early childhood education and care--early years--under the Coalition Government in England. Three factors came to define the Coalition's performance and record in this area: ambivalence about the rationales for the two areas of early education and childcare; a disconnect between early years and other…

  16. The record of Tethyan planktonic foraminifera at the early Paleogene hyperthermal events and Middle Eocene Climatic Optimum in northeastern Italy: are they comparable?

    NASA Astrophysics Data System (ADS)

    Luciani, Valeria; Giusberti, Luca; Agnini, Claudia; Fornaciari, Eliana; Rio, Domenico

    2010-05-01

    The early Paleogene is one of the more climatically and evolutionary dynamic periods in the Earth history that records a pronounced warming trend peaking in the Early Eocene, and a successive composite transition towards the modern icehouse world. Ever increasingly scientific attention is dedicated to definitely comprehend timing, nature and characters of the complex, non-linear evolution of the Paleogene climate. Several complete and expanded Paleogene successions (Forada, Possagno, Alano, Farra), with a sound magneto-biochronostratigraphic and stable isotope record crop out in the Venetian Southern Alps (Northeast Italy). Recent studies (Giusberti et. al., 2007; Luciani et al., 2007; Agnini et al., 2008) and unpublished data document the presence in these section of the main short-lived warming events (hyperthermals) of the Eocene (Paleocene-Eocene Thermal Maximum, PETM, ca 55 Ma, Eocene Layer of Mysterious Origin (ELMO, ca 53,6 Ma), X-event (ca 52.5 Ma), of the Early Eocene Climatic Optimum (EECO, ca 50-52 Ma) and of the Middle Eocene Climatic Optimum (MECO, ca 40 Ma; Zachos et al., 2001. 2008). All these events are typified by marked negative shifts in δ13C curves that correspond to carbonate decrease related to rise of the carbonate compensation depth in turn induced by large introduction in the ocean-atmosphere system of CO2. Common features to the warming events are pronounced and complex changes in planktonic foraminiferal assemblages, indicating strong environmental perturbations that perfectly parallel the variations of the stable isotope curves in all the examined events. These strict correspondences indicate close cause-effect relationships between changes in environmental conditions and modifications of the assemblages. Our analysis shows that the most striking variations are recorded by the PETM and MECO assemblages that reflect highly perturbed environments. The ELMO, X-event and EECO exhibit planktic foraminiferal responses that are similar to

  17. Recording signs of deterioration in acute patients: The documentation of vital signs within electronic health records in patients who suffered in-hospital cardiac arrest.

    PubMed

    Stevenson, Jean E; Israelsson, Johan; Nilsson, Gunilla C; Petersson, Göran I; Bath, Peter A

    2016-03-01

    Vital sign documentation is crucial to detecting patient deterioration. Little is known about the documentation of vital signs in electronic health records. This study aimed to examine documentation of vital signs in electronic health records. We examined the vital signs documented in the electronic health records of patients who had suffered an in-hospital cardiac arrest and on whom cardiopulmonary resuscitation was attempted between 2007 and 2011 (n = 228), in a 372-bed district general hospital. We assessed the completeness of vital sign data compared to VitalPAC™ Early Warning Score and the location of vital signs within the electronic health records. There was a noticeable lack of completeness of vital signs. Vital signs were fragmented through various sections of the electronic health records. The study identified serious shortfalls in the representation of vital signs in the electronic health records, with consequential threats to patient safety. © The Author(s) 2014.

  18. Zoophycos macroevolution since 541 Ma.

    PubMed

    Zhang, Li-Jun; Fan, Ruo-Ying; Gong, Yi-Ming

    2015-10-09

    Zoophycos is one of the most complex and enigmatic trace fossils recorded in marine strata from Cambrian to Quaternary worldwide, which is invaluable for the study of Phanerozoic development of organism-environment interactions. Here we address and demonstrate the macroevolution of Phanerozoic Zoophycos by assembling 448 points in constructing the Phanerozoic Zoophycos database based on 291 papers from 1821 to 2015 and 180 specimens from Cambrian to Palaeogene. The comprehensive dataset reveals, for the first time, five peaks and six depressions in Phanerozoic Zoophycos occurrence frequency. Secondly, the palaeogeographical distribution of Zoophycos is closely associated with the supercontinent Pangaea shifting, independent of the latitude. Our data also attest that the bathymetrical shift of Zoophycos from the littoral-neritic to bathyal environments is synchronized with the tiering shift from shallow to deep. By detailed comparison with body fossils, geochemical and palaeogeographical records, we conclude that the macroevolution of Phanerozoic Zoophycos is multi-affected by the global biodiversity expansion, benthic nutrient enhancement, and the biotic macroevolution of the Zoophycos-producers. The macroevolution of development evidenced by the morphological changes of Zoophycos and other trace fossils, may have great implications on the behavioural and physiological adaptation of ancient animals to the environments.

  19. Zoophycos macroevolution since 541 Ma

    NASA Astrophysics Data System (ADS)

    Zhang, Li-Jun; Fan, Ruo-Ying; Gong, Yi-Ming

    2015-10-01

    Zoophycos is one of the most complex and enigmatic trace fossils recorded in marine strata from Cambrian to Quaternary worldwide, which is invaluable for the study of Phanerozoic development of organism-environment interactions. Here we address and demonstrate the macroevolution of Phanerozoic Zoophycos by assembling 448 points in constructing the Phanerozoic Zoophycos database based on 291 papers from 1821 to 2015 and 180 specimens from Cambrian to Palaeogene. The comprehensive dataset reveals, for the first time, five peaks and six depressions in Phanerozoic Zoophycos occurrence frequency. Secondly, the palaeogeographical distribution of Zoophycos is closely associated with the supercontinent Pangaea shifting, independent of the latitude. Our data also attest that the bathymetrical shift of Zoophycos from the littoral-neritic to bathyal environments is synchronized with the tiering shift from shallow to deep. By detailed comparison with body fossils, geochemical and palaeogeographical records, we conclude that the macroevolution of Phanerozoic Zoophycos is multi-affected by the global biodiversity expansion, benthic nutrient enhancement, and the biotic macroevolution of the Zoophycos-producers. The macroevolution of development evidenced by the morphological changes of Zoophycos and other trace fossils, may have great implications on the behavioural and physiological adaptation of ancient animals to the environments.

  20. Evidence of orbital forcing in 510 to 530 million year old shallow marine cycles, Utah and western Canada

    NASA Technical Reports Server (NTRS)

    Bond, Gerard C.; Beavan, John; Kominz, Michelle A.; Devlin, William

    1992-01-01

    Spectral analyses of two sequences of shallow marine sedimentary cycles that were deposited between 510 and 530 million years ago were completed. One sequence is from Middle Cambrian rocks in southern Utah and the other is from Upper Cambrian rocks in the southern Canadian Rockies. In spite of the antiquity of these strata, and even though there are differences in the age, location, and cycle facies between the two sequences, both records have distinct spectral peaks with surprisingly similar periodicities. A null model constructed to test for significance of the spectral peaks and circulatory in the methodology indicates that all but one of the spectral peaks are significant at the 90 percent confidence level. When the ratios between the statistically significant peaks are measured, we find a consistent relation to orbital forcing; specifically, the spectral peak ratios in both the Utah and Canadian examples imply that a significant amount of the variance in the cyclic records is driven by the short eccentricity (approximately 109 ky) and by the precessional (approximately 21 ky) components of the Earth's orbital variations. Neither section contains a significant component of variance at the period of the obliquity cycle, however.

  1. Microbial mats in playa lakes and other saline habitats: Early Mars analog?

    NASA Technical Reports Server (NTRS)

    Bauld, John

    1989-01-01

    Microbial mats are cohesive benthic microbial communities which inhabit various Terra (Earth-based) environments including the marine littoral and both permanent and ephemeral (playa) saline lakes. Certain geomorphological features of Mars, such as the Margaritifer Sinus, were interpreted as ancient, dried playa lakes, presumably formed before or during the transition to the present Mars climate. Studies of modern Terran examples suggest that microbial mats on early Mars would have had the capacity to survive and propagate under environmental constraints that would have included irregularly fluctuating regimes of water activity and high ultraviolet flux. Assuming that such microbial communities did indeed inhabit early Mars, their detection during the Mars Rover Sample Return (MRSR) mission depends upon the presence of features diagnostic of the prior existence of these communities or their component microbes or, as an aid to choosing suitable landing, local exploration or sampling sites, geomorphological, sedimentological or chemical features characteristic of their playa lake habitats. Examination of modern Terran playas (e.g., the Lake Eyre basin) shows that these features span several orders of magnitude in size. While stromatolites are commonly centimeter-meter scale features, bioherms or fields of individuals may extend to larger scales. Preservation of organic matter (mats and microbes) would be favored in topographic lows such as channels or ponds of high salinity, particularly those receiving silica-rich groundwaters. These areas are likely to be located near former zones of groundwater emergence and/or where flood channels entered the paleo-playa. Fossil playa systems which may aid in assessing the applicability of this particular Mars analog include the Cambrian Observatory Hill Beds of the Officer Basin and the Eocene Wilkins Peak Member of the Green River Formation.

  2. Geologic Map of the Gold Creek Gold District, Elko County, Nevada

    USGS Publications Warehouse

    Ketner, Keith B.

    2007-01-01

    The Gold Creek, Nev. area displays important stratigraphic and structural relationships between Paleozoic and early Tertiary sedimentary strata in an area dominated by large intrusive bodies of Mesozoic age and extensive volcanic fields of middle to late Tertiary age. An autochthonous sequence includes the Cambrian and Proterozoic(?) Prospect Mountain Quartzite and the overlying Cambrian and Ordovician Tennessee Mountain Formation. This autochthon is overlain by three allochthonous plates each composed of a distinctive sequence of strata and having a distinctive internal structure. The structurally lowest plate is composed of the Havallah sequence, locally of Mississippian and Pennsylvanian age, which is folded on north-south trending axes. The next higher plate is composed of somewhat younger Pennsylvanian and Permian strata cut by east-west trending low-angle faults. The highest plate is composed of early Tertiary non-marine sedimentary and igneous rocks folded on varied but mainly north-south trending axes. The question of whether the allochthonous plates were emplaced by contractional or extensional forces is indeterminate from the local evidence. Mineral deposits include gold placers of moderate size and small pockets of base metals, none of which is currently being exploited.

  3. Astronomical pacing of the global silica cycle recorded in Mesozoic bedded cherts

    NASA Astrophysics Data System (ADS)

    Ikeda, Masayuki; Tada, Ryuji; Ozaki, Kazumi

    2017-06-01

    The global silica cycle is an important component of the long-term climate system, yet its controlling factors are largely uncertain due to poorly constrained proxy records. Here we present a ~70 Myr-long record of early Mesozoic biogenic silica (BSi) flux from radiolarian chert in Japan. Average low-mid-latitude BSi burial flux in the superocean Panthalassa is ~90% of that of the modern global ocean and relative amplitude varied by ~20-50% over the 100 kyr to 30 Myr orbital cycles during the early Mesozoic. We hypothesize that BSi in chert was a major sink for oceanic dissolved silica (DSi), with fluctuations proportional to DSi input from chemical weathering on timescales longer than the residence time of DSi (<~100 Kyr). Chemical weathering rates estimated by the GEOCARBSULFvolc model support these hypotheses, excluding the volcanism-driven oceanic anoxic events of the Early-Middle Triassic and Toarcian that exceed model limits. We propose that the Mega monsoon of the supercontinent Pangea nonlinearly amplified the orbitally paced chemical weathering that drove BSi burial during the early Mesozoic greenhouse world.

  4. Facies patterns and conodont biogeography in Arctic Alaska and the Canadian Arctic Islands: Evidence against juxtaposition of these areas during early Paleozoic time

    USGS Publications Warehouse

    Dumoulin, Julie A.; Harris, A.G.; Bradley, D.C.; De Freitas, T. A.

    2000-01-01

    Differences in lithofacies and biofacies suggest that lower Paleozoic rocks now exposed in Arctic Alaska and the Canadian Arctic Islands did not form as part of a single depositional system. Lithologic contrasts are noted in shallow- and deep-water strata and are especially marked in Ordovician and Silurian rocks. A widespread intraplatform basin of Early and Middle Ordovician age in northern Alaska has no counterpart in the Canadian Arctic, and the regional drowning and backstepping of the Silurian shelf margin in Canada has no known parallel in northern Alaska. Lower Paleozoic basinal facies in northern Alaska are chiefly siliciclastic, whereas resedimented carbonates are volumetrically important in Canada. Micro- and macrofossil assemblages from northern Alaska contain elements typical of both Siberian and Laurentian biotic provinces; coeval Canadian Arctic assemblages contain Laurentian forms but lack Siberian species. Siberian affinities in northern Alaskan biotas persist from at least Middle Cambrian through Mississippian time and appear to decrease in intensity from present-day west to east. Our lithologic and biogeographic data are most compatible with the hypothesis that northern Alaska-Chukotka formed a discrete tectonic block situated between Siberia and Laurentia in early Paleozoic time. If Arctic Alaska was juxtaposed with the Canadian Arctic prior to opening of the Canada basin, biotic constraints suggest that such juxtaposition took place no earlier than late Paleozoic time.

  5. Provenance, tectonic setting and source-area weathering of the lower Cambrian sediments of the Parahio valley in the Spiti basin, India

    NASA Astrophysics Data System (ADS)

    Pandey, Shivani; Parcha, Suraj K.

    2017-03-01

    The geochemical study of siliciclastic rocks from the Lower Cambrian of Parahio Valley has been studied to describe the provenance, chemical weathering and tectonic setting. The K2O/Al2O3 ratio and positive correlation of Co ( r=0.85), Ni ( r=0.86), Zn ( r=0.82), Rb ( r=0.98) with K2O reflects that the presence of clay minerals control the abundances of these elements and suggests a warm and humid climate for this region. The chondrite normalized REE pattern of the samples is equivalent to upper continental crust, which reflects enriched LREE and flat HREE with negative Eu anomaly. The tectonic setting discriminant diagram log[K2O/Na2O] vs. SiO2; [SiO2/Al2O3] vs. log[K2O/Na2O]; [SiO2/20] - [K2O+Na2O] - [TiO2+Fe2O3+MgO] indicates transitional tectonic setting from an active continental margin to a passive margin. The discriminant function plot indicates quartzose sedimentary provenance, and to some extent, the felsic igneous provenance, derived from weathered granite, gneissic terrain and/or from pre-existing sedimentary terrain. The CIA value indicates low to moderate degree of chemical weathering and the average ICV values suggests immature sediments deposited in tectonically active settings. The A-CN-K diagram indicates that these sediments were generated from source rocks of the upper continental crust.

  6. Sporadic sampling, not climatic forcing, drives observed early hominin diversity.

    PubMed

    Maxwell, Simon J; Hopley, Philip J; Upchurch, Paul; Soligo, Christophe

    2018-05-08

    The role of climate change in the origin and diversification of early hominins is hotly debated. Most accounts of early hominin evolution link observed fluctuations in species diversity to directional shifts in climate or periods of intense climatic instability. None of these hypotheses, however, have tested whether observed diversity patterns are distorted by variation in the quality of the hominin fossil record. Here, we present a detailed examination of early hominin diversity dynamics, including both taxic and phylogenetically corrected diversity estimates. Unlike past studies, we compare these estimates to sampling metrics for rock availability (hominin-, primate-, and mammal-bearing formations) and collection effort, to assess the geological and anthropogenic controls on the sampling of the early hominin fossil record. Taxic diversity, primate-bearing formations, and collection effort show strong positive correlations, demonstrating that observed patterns of early hominin taxic diversity can be explained by temporal heterogeneity in fossil sampling rather than genuine evolutionary processes. Peak taxic diversity at 1.9 million years ago (Ma) is a sampling artifact, reflecting merely maximal rock availability and collection effort. In contrast, phylogenetic diversity estimates imply peak diversity at 2.4 Ma and show little relation to sampling metrics. We find that apparent relationships between early hominin diversity and indicators of climatic instability are, in fact, driven largely by variation in suitable rock exposure and collection effort. Our results suggest that significant improvements in the quality of the fossil record are required before the role of climate in hominin evolution can be reliably determined. Copyright © 2018 the Author(s). Published by PNAS.

  7. Maps showing petroleum exploration intensity and production in major Cambrian to Ordovician reservoir rocks in the Anadarko Basin

    USGS Publications Warehouse

    Henry, Mitch; Hester, Tim

    1996-01-01

    The Anadarko basin is a large, deep, two-stage Paleozoic basin (Feinstein, 1981) that is petroleum rich and generally well explored. The Anadarko basin province, a geogrphic area used here mostly for the convenience of mapping and data management, is defined by political boundaries that include the Anadarko basin proper. The boundaries of the province are identical to those used by the U.S. Geological Survey (USGS) in the 1995 National Assessment of United Stated Oil and Gas Resources. The data in this report, also identical to those used in the national assessment, are from several computerized data bases including Nehring Research Group (NRG) Associates Inc., Significant Oil and Gas Fields of the United States (1992); Petroleum Information (PI), Inc., Well History Control System (1991); and Petroleum Information (PI), Inc., Petro-ROM: Production data on CD-ROM (1993). Although generated mostly in response to the national assessment, the data presented here arc grouped differently and arc displayed and described in greater detail. In addition, the stratigraphic sequences discussed may not necessarily correlate with the "plays" of the 1995 national assessment. This report uses computer-generated maps to show drilling intensity, producing wells, major fields, and other geologic information relevant to petroleum exploration and production in the lower Paleozoic part of the Anadarko basin province as defined for the U.S. Geological Survey's 1995 national petroleum assessment. Hydrocarbon accumulations must meet a minimum standard of 1 million barrels of oil (MMBO) or 6 billion cubic feet of gas (BCFG) estimated ultimate recovery to be included in this report as a major field or revoir. Mapped strata in this report include the Upper Cambrian to Lower Ordovician Arbuckle and Low Ordovician Ellenburger Groups, the Middle Ordovician Simpson Group, and the Middle to Upper Ordovician Viola Group.

  8. Saline-water resources of Texas

    USGS Publications Warehouse

    Winslow, Allen George; Kister, Lester Ray

    1956-01-01

    Most of the aquifers in Texas contain saline water in some parts, and a few are capable of producing large quantities of saline water. Of the early Paleozoic formations, the Hickory sandstone member of the Riley formation of Cambrian age and the Ellenburger group of Ordovician age are potential sources of small to moderate supplies of saline water in parts of central and west-central Texas.

  9. Stenian Estuarine System and Early Neoproterozoic Microbial Records of Capiru Formation, Southern Ribeira Belt.

    NASA Astrophysics Data System (ADS)

    Cury, L. F.; Santos, L. D. R.; Leandro, R.; Lange, L.; Bahniuk Rumbelsperger, A.

    2017-12-01

    The Capiru formation is a low-grade metasedimentary sequence composed by slates, rhythmic phyllites, quartzites and marbles, disposed and disrupted in tectonic blocks delimited by thrust and strike-slip faults related to oblique collisions in the southern Ribeira Belt, Curitiba terrane, southern Brazil. The rocks of the Capiru formation crops out as a thrust-folded belt, delimited on the north by the transcurrent faults of Lancinha Shear Zone (LSZ), and to the south by thrust faults with large isograde variation. Three lithological sequences are recognized mainly by their compositional and stratigraphic records, including a (i) ferruginous sequence with quartzites, metasandstones and metaconglomerates with goethite/hematite cements and phyllites with magnetite; ii) metadolomites with stromatolites, interbeded with pelitic layers and iii) a metapelitic sequence with metarhythmites and metasandstones with well preserved organic-rich material. The records of two tectonic-metamorphic events related to thrust and transpressive tectonics are heterogeneously developed in all sequences, still been recognized sections with the original stratigraphic succession. The stratigraphic record suggests an estuarine environment with rising sea level developing tidal flats and tidal channels. U-Pb detrital zircon analyses characterizes Rhyacian ages (between 2.2-2.1 Ga) as the main sources, and Stenian ages (between 1.08-1.20 Ga) as maximum age for sedimentation. The metapelites mineral assemblage is composed by quartz, muscovite, sericite, illite, kaolinite, sepiolite, magnetite, goethite, hematite and carbonaceous material with bulk organic carbon content (BOC) ranging from 0.09 to 1.21 (%), a precambrian microbial activity record. The metadolomites are characterized by the presence of stromatolites in different types and dimensions, with microbial activity records supported by SEM-EDS (up to 91% C), with EPS-like morphologies within microporosity, NaCl compounds and clay minerals

  10. An Impaired View of Earth's Early History

    NASA Astrophysics Data System (ADS)

    Vervoort, J. D.; Kemp, A. I.; Bauer, A.; Bowring, S. A.; Fisher, C.

    2014-12-01

    The Hf and Nd isotope records of Earth's early history are sparse, difficult to interpret, and controversial, much like the few remnants of crust older than 4 Ga. New analytical techniques have been brought to bear on this problem but despite this recent work­-or, perhaps, because of it-the record is no clearer than it was 15 years ago. Several studies, based on highly variable calculated initial isotopic compositions, have argued for highly heterogeneous crust and mantle reservoirs in the early Earth1,2 and an ultra-depleted Eoarchean mantle3. These data come mostly from two sources: Hf-Nd isotope analyses of ultramafic rocks and Hf isotope analyses of zircons by solution or laser ablation. An important question for understanding the chemical evolution of the early Earth is: Do these data offer a unique window into the early Earth or are they artefacts not representative of crust/mantle evolution, giving an impaired view of the Earth's early history? In complex samples, measured isotopic compositions can result from open-system behavior in easily altered ultramafic compositions, in multicomponent, polymetamorphic gneisses, or in zircons with multiple generations of growth. Perhaps most importantly, accurate age assignment is often lacking, compromised, or impossible in these rocks, making calculation of initial epsilon Hf and Nd values ambiguous at best. In order to gain insight into crust mantle evolution in the early Earth we need, above all, a robust and unambiguous isotopic record to work with. This can be achieved by integrating zircon U-Pb and Hf and whole-rock Hf and Nd isotope compositions in relatively undisturbed igneous rocks with well-constrained ages. When this approach is used apparent isotopic heterogeneity decreases and a simpler model for crust-mantle evolution in the early Earth emerges. Careful screening of geological relationships, petrology, and geochemistry of samples from the early Earth should be done before interpreting isotopic data

  11. Arthropod visual predators in the early pelagic ecosystem: evidence from the Burgess Shale and Chengjiang biotas.

    PubMed

    Vannier, J; García-Bellido, D C; Hu, S-X; Chen, A-L

    2009-07-22

    Exceptional fossil specimens with preserved soft parts from the Maotianshan Shale (ca 520 Myr ago) and the Burgess Shale (505 Myr ago) biotas indicate that the worldwide distributed bivalved arthropod Isoxys was probably a non-benthic visual predator. New lines of evidence come from the functional morphology of its powerful prehensile frontal appendages that, combined with large spherical eyes, are thought to have played a key role in the recognition and capture of swimming or epibenthic prey. The swimming and steering of this arthropod was achieved by the beating of multiple setose exopods and a flap-like telson. The appendage morphology of Isoxys indicates possible phylogenetical relationships with the megacheirans, a widespread group of assumed predator arthropods characterized by a pre-oral 'great appendage'. Evidence from functional morphology and taphonomy suggests that Isoxys was able to migrate through the water column and was possibly exploiting hyperbenthic niches for food. Although certainly not unique, the case of Isoxys supports the idea that off-bottom animal interactions such as predation, associated with complex feeding strategies and behaviours (e.g. vertical migration and hunting) were established by the Early Cambrian. It also suggests that a prototype of a pelagic food chain had already started to build-up at least in the lower levels of the water column.

  12. Perception of music performance on historical and modern commercial recordings.

    PubMed

    Timmers, Renee

    2007-11-01

    Performing styles as well as recording styles have changed considerably within the 20th century. To what extent do the age of a recording, the unfamiliarity with performing style, and the quality of a reproduction of a recording systematically influence how we perceive performances on record? Four exploratory experiments were run to formulate an answer to this question. Each experiment examined a different aspect of the perception of performance, including judgments of quality, perceived emotion, and dynamics. Fragments from Die junge Nonne sung by famous singers from the start, middle, and second half of the 20th century were presented in a noisy and clean version to musically trained participants. The results show independence of perception of emotional activity from recording date, strong dependence of perceived quality and emotional impact on recording date, and only limited effects of reproduction quality. Standards have clearly changed, which influence judgments of quality and age. Additionally, changes restrict the communication between early recorded performers and modern listeners to some extent as shown by systematically smaller variations in communicated dynamics and emotional valence for older recordings.

  13. Biostratigraphy, taxonomic diversity and patterns of morphological evolution of Ordovician acritarchs (organic-walled microphytoplankton) from the northern Gondwana margin in relation to palaeoclimatic and palaeogeographic changes

    NASA Astrophysics Data System (ADS)

    Vecoli, Marco; Le Hérissé, Alain

    2004-10-01

    Acritarchs, the fossilizable, resting cysts of phytoplanktonic algal protists, are the dominant component of marine organic-walled microfossils in the Palaeozoic. The majority of acritarchs show strong similarities with dinoflagellate cysts in morphological and biogeochemical features, as well as distributional patterns in the sediments. The production of these organic-walled microfossils and their distribution and survivorship in the sediments were controlled by differences in ecological tolerances and life cycle (autecology) of the planktonic parent organisms. Calculation of evolutionary rates and development of a detailed diversity curve at specific level, form the basis for discussing the influence of global palaeoenvironmental perturbations on the evolution of organic-walled microphytoplankton in northern Gondwana during latest Cambrian through Ordovician times. The potential of acritarchs for biostratigraphic correlation at the regional scale (northern Gondwana domain) is much improved by our detailed revision of distributional patterns of 245 acritarch taxa. The most important Cambro-Ordovician acritarch bio-events are short periods of diversification, which also correspond to introduction of morphological innovations, observed in latest Cambrian and earliest Tremadoc, late Tremadoc, early Arenig, basal Llanvirn, and latest Ashgill, and an important extinction phase in the early Caradoc. Overall, acritarch diversity increased from the basal Ordovician up to the middle Llanvirn, then declined in the early and middle Caradoc. During Ashgill times, the assemblages are poorly diversified at the generic level as a result of a combined effect of sea level drawdown and onset of glacial conditions, but no major extinction event is observed in connection with the end-Ordovician biotic crisis. The peak in acritarch diversity during Middle Ordovician times appears to be correlated to maximum spread of palaeogeographical assembly. Acritarch dynamics appear largely

  14. Astronomical pacing of the global silica cycle recorded in Mesozoic bedded cherts

    PubMed Central

    Ikeda, Masayuki; Tada, Ryuji; Ozaki, Kazumi

    2017-01-01

    The global silica cycle is an important component of the long-term climate system, yet its controlling factors are largely uncertain due to poorly constrained proxy records. Here we present a ∼70 Myr-long record of early Mesozoic biogenic silica (BSi) flux from radiolarian chert in Japan. Average low-mid-latitude BSi burial flux in the superocean Panthalassa is ∼90% of that of the modern global ocean and relative amplitude varied by ∼20–50% over the 100 kyr to 30 Myr orbital cycles during the early Mesozoic. We hypothesize that BSi in chert was a major sink for oceanic dissolved silica (DSi), with fluctuations proportional to DSi input from chemical weathering on timescales longer than the residence time of DSi (<∼100 Kyr). Chemical weathering rates estimated by the GEOCARBSULFvolc model support these hypotheses, excluding the volcanism-driven oceanic anoxic events of the Early-Middle Triassic and Toarcian that exceed model limits. We propose that the Mega monsoon of the supercontinent Pangea nonlinearly amplified the orbitally paced chemical weathering that drove BSi burial during the early Mesozoic greenhouse world. PMID:28589958

  15. Astronomical pacing of the global silica cycle recorded in Mesozoic bedded cherts.

    PubMed

    Ikeda, Masayuki; Tada, Ryuji; Ozaki, Kazumi

    2017-06-07

    The global silica cycle is an important component of the long-term climate system, yet its controlling factors are largely uncertain due to poorly constrained proxy records. Here we present a ∼70 Myr-long record of early Mesozoic biogenic silica (BSi) flux from radiolarian chert in Japan. Average low-mid-latitude BSi burial flux in the superocean Panthalassa is ∼90% of that of the modern global ocean and relative amplitude varied by ∼20-50% over the 100 kyr to 30 Myr orbital cycles during the early Mesozoic. We hypothesize that BSi in chert was a major sink for oceanic dissolved silica (DSi), with fluctuations proportional to DSi input from chemical weathering on timescales longer than the residence time of DSi (<∼100 Kyr). Chemical weathering rates estimated by the GEOCARBSULFvolc model support these hypotheses, excluding the volcanism-driven oceanic anoxic events of the Early-Middle Triassic and Toarcian that exceed model limits. We propose that the Mega monsoon of the supercontinent Pangea nonlinearly amplified the orbitally paced chemical weathering that drove BSi burial during the early Mesozoic greenhouse world.

  16. The early evolution of eukaryotes - A geological perspective

    NASA Technical Reports Server (NTRS)

    Knoll, Andrew H.

    1992-01-01

    This paper examines the goodness of fit between patterns of biological and environmental history implied by molecular phylogenies of eukaryotic organisms and the geological records of early eukaryote evolution. It was found that Precambrian geological records show evidence that episodic increases in biological diversity roughly coincided with episodic environmental changes and by sharp increases in atmospheric oxygen concentrations which significantly changed the earth surface environments. Although the goodness of fit among physical and biological changes is gratifyingly high, the records of these changes do not always coincide in time. The additional information in these fields that is needed for complete integration of geological and phylogenic records is suggested.

  17. Holocene East Asian summer monsoon records in northern China and their inconsistency with Chinese stalagmite δ18O records

    NASA Astrophysics Data System (ADS)

    Liu, Jianbao; Chen, Jianhui; Zhang, Xiaojian; Chen, Fahu

    2016-04-01

    Monsoon precipitation over China exhibits large spatial differences. It has been found that a significantly enhanced East Asian summer monsoon (EASM) is characterized by increased rainfall in northern China and by reduced rainfall in southern China, and this relationship occurs on different time scales during the Holocene. This study presents results from a diverse range of proxy paleoclimatic records from northern China where precipitation variability is traditionally considered as an EASM proxy. Our aim is to evaluate the evolution of the EASM during the Holocene and to compare it with all of the published stalagmite δ18O records from the Asian Monsoon region in order to explore the potential mechanism(s) controlling the Chinese stalagmite δ18O. We found that the intensity of the EASM during the Holocene recorded by the traditional EASM proxy of moisture (or precipitation) records from northern China are significantly different from the Chinese stalagmite δ18O records. The EASM maximum occurred during the mid-Holocene, challenging the prevailing view of an early Holocene EASM maximum mainly inferred from stalagmite δ18O records in eastern China. In addition, all of the well-dated Holocene stalagmite δ18O records, covering a broad geographical region, exhibit a remarkably similar trend of variation and are statistically well-correlated on different time scales, thus indicating a common signal. However, in contrast with the clear consistency in the δ18O values in all of the cave records, both instrumental and paleoclimatic records exhibit significant spatial variations in rainfall on decadal-to- centennial time scales over eastern China. In addition, both paleoclimatic records and modeling results suggest that Holocene East Asian summer monsoon precipitation reached a maximum at different periods in different regions of China. Thus the stalagmite δ18O records from the EASM region should not be regarded as a reliable indicator of the strength of the East

  18. Integrated approaches to terminal Proterozoic stratigraphy: an example from the Olenek Uplift, northeastern Siberia

    NASA Technical Reports Server (NTRS)

    Knoll, A. H.; Grotzinger, J. P.; Kaufman, A. J.; Kolosov, P.

    1995-01-01

    In the Olenek Uplift of northeastern Siberia, the Khorbusuonka Group and overlying Kessyusa and Erkeket formations preserve a significant record of terminal Proterozoic and basal Cambrian Earth history. A composite section more than 350 m thick is reconstructed from numerous exposures along the Khorbusuonka River. The Khorbusuonka Group comprises three principal sedimentary sequences: peritidal dolomites of the Mastakh Formation, which are bounded above and below by red beds; the Khatyspyt and most of the overlying Turkut formations, which shallow upward from relatively deep-water carbonaceous micrites to cross-bedded dolomitic grainstones and stromatolites; and a thin upper Turkut sequence bounded by karst surfaces. The overlying Kessyusa Formation is bounded above and below by erosional surfaces and contains additional parasequence boundaries internally. Ediacaran metazoans, simple trace fossils, and vendotaenids occur in the Khatyspyt Formation; small shelly fossils, more complex trace fossils, and acritarchs all appear near the base of the Kessyusa Formation and diversify upward. The carbon-isotopic composition of carbonates varies stratigraphically in a pattern comparable to that determined for other terminal Proterozoic and basal Cambrian successions. In concert, litho-, bio-, and chemostratigraphic data indicate the importance of the Khorbusuonka Group in the global correlation of terminal Proterozoic sedimentary rocks. Stratigraphic data and a recently determined radiometric date on basal Kessyusa volcanic breccias further underscore the significance of the Olenek region in investigations of the Proterozoic-cambrian boundary.

  19. Integrated approaches to terminal Proterozoic stratigraphy: an example from the Olenek Uplift, northeastern Siberia.

    PubMed

    Knoll, A H; Grotzinger, J P; Kaufman, A J; Kolosov, P

    1995-01-01

    In the Olenek Uplift of northeastern Siberia, the Khorbusuonka Group and overlying Kessyusa and Erkeket formations preserve a significant record of terminal Proterozoic and basal Cambrian Earth history. A composite section more than 350 m thick is reconstructed from numerous exposures along the Khorbusuonka River. The Khorbusuonka Group comprises three principal sedimentary sequences: peritidal dolomites of the Mastakh Formation, which are bounded above and below by red beds; the Khatyspyt and most of the overlying Turkut formations, which shallow upward from relatively deep-water carbonaceous micrites to cross-bedded dolomitic grainstones and stromatolites; and a thin upper Turkut sequence bounded by karst surfaces. The overlying Kessyusa Formation is bounded above and below by erosional surfaces and contains additional parasequence boundaries internally. Ediacaran metazoans, simple trace fossils, and vendotaenids occur in the Khatyspyt Formation; small shelly fossils, more complex trace fossils, and acritarchs all appear near the base of the Kessyusa Formation and diversify upward. The carbon-isotopic composition of carbonates varies stratigraphically in a pattern comparable to that determined for other terminal Proterozoic and basal Cambrian successions. In concert, litho-, bio-, and chemostratigraphic data indicate the importance of the Khorbusuonka Group in the global correlation of terminal Proterozoic sedimentary rocks. Stratigraphic data and a recently determined radiometric date on basal Kessyusa volcanic breccias further underscore the significance of the Olenek region in investigations of the Proterozoic-cambrian boundary.

  20. Altered carbon cycling and coupled changes in Early Cretaceous weathering patterns: Evidence from integrated carbon isotope and sandstone records of the western Tethys

    NASA Astrophysics Data System (ADS)

    Wortmann, Ulrich Georg; Herrle, Jens Olaf; Weissert, Helmut

    2004-03-01

    In this study we investigate if a major perturbation of the Early Cretaceous carbon cycle was accompanied by altered weathering and erosion rates. The large Aptian carbon isotope anomaly records the response of the biosphere to widespread volcanic activity and probably resulting changes in atmospheric pCO2 levels. Elevated pCO2 levels should also result in an accelerated hydrological cycle and increased silicate weathering, creating a negative feedback loop removing CO2 from the atmosphere. We propose to interpret the widespread occurrence of quartz sandstones in the Tethys-Atlantic seaway as a result of altered weathering and erosion rates in the wake of the Aptian carbon cycle excursion. We challenge the traditional notion that these are 'flysch' deposits associated with Early Cretaceous orogenic movements in the western Tethys. We propose that these sandstones were most likely part of a large conveyor belt system, acting along the Iberian and European margin of the Tethys seaway. Using chemostratigraphic correlations, we show that the activity of this system was only short-lived and coeval with changes in coastal ecology and the Aptian carbon cycle perturbations. We tentatively relate the existence of this system to a transient climate regime, characterized by fluctuating pCO2 levels.