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Sample records for eastern spruce structural

  1. Characterization of eastern US spruce-fir soils. Book chapter

    SciTech Connect

    Fernandez, I.J.

    1992-01-01

    The spruce-fir forest of the eastern United States encompasses a diverse range of edaphic conditions due to differences in surficial geology, mineralogy, elevation, and climate. This chapter describes the characteristics of soils supporting eastern spruce-fir ecosystems, including soil properties that are important in understanding forest function and the consequences of atmospheric deposition to forested ecosystems. Chapter 1 describes the silvical characteristics of the spruce-fir forest. The Spruce-Fir Research Cooperative included six intensive study sites; five were high-elevation research sites located from western North Carolina to New Hampshire, with one low-elevation site in Maine. Information gained from research at these sites, and other relevant research from these regions, provides the basis for this description of eastern U. S. spruce-fir soils.

  2. AmeriFlux CA-Qfo Quebec - Eastern Boreal, Mature Black Spruce

    SciTech Connect

    Margolis, Hank A.

    2016-01-01

    This is the AmeriFlux version of the carbon flux data for the site CA-Qfo Quebec - Eastern Boreal, Mature Black Spruce. Site Description - 49.69247° N / 74.34204° W, elevation of 387 mm, 90 - 100 yr old Black Spruce, Jack Pine, feather moss

  3. Impact of eastern dwarf mistletoe (Arceuthobium pusillum) infection on the needles of red spruce (Picea rubens) and white spruce (Picea glauca): oxygen exchange, morphology and composition.

    PubMed

    Reblin, Jaret S; Logan, Barry A; Tissue, David T

    2006-10-01

    Eastern dwarf mistletoe (Arceuthobium pusillum Peck) is a hemiparasitic angiosperm that infects white spruce (Picea glauca (Moench) Voss) and red spruce (P. rubens Sarg.) in northeastern North America. The effects of mistletoe infection differ substantially between white and red spruce, with white spruce suffering greater infection-induced mortality. In the present study, we sought to determine the role that species-specific differences in needle-scale responses to parasitism may play in the observed differences in the effect of infection on host tree health. Based on the measurements made, the most apparent effect of parasitism was a reduction in needle size distal to infections. The magnitude of this reduction was greater in white spruce than in red spruce. Eastern dwarf mistletoe was a sink for host photosynthate in red spruce and white spruce; however, there were no adjustments in needle photosynthetic capacities in either host to accommodate the added sink demands of the parasite. Needle total nonstructural carbohydrate concentrations (TNC) were also unaltered by infection. Red spruce needles had higher TNC concentrations despite having lower overall photosynthetic capacities, suggesting that red spruce may be more sink limited and therefore better able to satisfy the added sink demands of parasitic infection. However, if carbon availability limits the growth of the mistletoe, one may expect that the extent of the parasitic infection would be greater in red spruce. Yet in the field, the extent of infection is generally greater in white spruce. Taken together, these results suggest that dwarf mistletoe may not substantially perturb the carbon balance of either host spruce species and that species-specific differences in needle-scale responses to the parasite cannot explain the contrasting effects of infection on white spruce and red spruce.

  4. Structural changes in spruce and fir needles.

    PubMed

    Schmitt, U; Ruetze, M

    1990-01-01

    Needles from spruce and fir trees were analyzed for histological changes induced by long-term exposure in open-top chambers to SO(2) and/or O(3) combined with acid rain. Light and electron microscopical evaluation revealed initial structural changes in the vascular bundle of fir needles, with an increased number of crushed sieve cells in the phloem. In addition the walls of young, adaxial sieve cells lacked the typical thickening usually observed in naturally aged needles. These findings may indicate restricted assimilate translocation. The presence of SO(2) in any treatment led to thylakoidal swellings and membrane reductions in the chloroplasts of mesophyll cells near the vascular bundle. This damage pattern resembled alterations caused by nutrient deficiency rather than by the direct action of gaseous pollutants. In general, fir appears to be more sensitive to environmental stress than spruce; this substantiates the findings of previous studies.

  5. Availability of Ectomycorrhizal Fungi to Black Spruce above the Present Treeline in Eastern Labrador

    PubMed Central

    Reithmeier, Laura; Kernaghan, Gavin

    2013-01-01

    Ectomycorrhizal fungi (ECMF) are an important biotic factor in the survival of conifer seedlings under stressful conditions and therefore have the potential to facilitate conifer establishment into alpine and tundra habitats. In order to assess patterns of ectomycorrhizal availability and community structure above treeline, we conducted soil bioassays in which Picea mariana (black spruce) seedlings were grown in field-collected soils under controlled conditions. Soils were collected from distinct alpine habitats, each dominated by a different ectomycorrhizal host shrub: Betula glandulosa, Arctostaphylos alpina or Salix herbacaea. Within each habitat, half of the soils collected contained roots of ectomycorrhizal shrubs (host+) and the other half were free of host plants (host−). Forest and glacial moraine soils were also included for comparison. Fungi forming ectomycorrhizae during the bioassays were identified by DNA sequencing. Our results indicate that ECMF capable of colonizing black spruce are widespread above the current tree line in Eastern Labrador and that the level of available inoculum has a significant influence on the growth of seedlings under controlled conditions. Many of the host− soils possessed appreciable levels of ectomycorrhizal inoculum, likely in the form of spore banks. Inoculum levels in these soils may be influenced by spore production from neighboring soils where ectomycorrhizal shrubs are present. Under predicted temperature increases, ectomycorrhizal inoculum in soils with host shrubs as well as in nearby soils without host shrubs have the potential to facilitate conifer establishment above the present tree line. PMID:24204858

  6. Olfactory response of eastern spruce budworm larvae to red spruce needles exposed to acid rain and elevated levels of ozone.

    PubMed

    Cannon, W N

    1990-12-01

    Second-instar eastern spruce budworm larvae,Choristoneura fumiferana (Clem.), were tested in a two-choice, Y-type, wind-tunnel olfactometer for preferences for red spruce,Picea rubens Sarg., needles exposed for eight weeks to charcoal-filtered air (CFA) or 0.15 ppm ozone (O3) in combination with acidified rainfall at pH 4.2 or 3.0. Volatiles from needles treated with CFA plus pH 4.2 rain (control) were preferred over those from needles exposed to pH 3.0 rain + O3 or O3 alone; O3-treated needles were chosen over those exposed to pH 3.0 rain + O3. No preference was shown between the pH 4.2 and 3.0 rain treatments. Larvae chose purified air flowing through the olfactometer in preference to needle volatiles from the pH 3.0 rain + O3 treatment.

  7. AmeriFlux CA-Qcu Quebec - Eastern Boreal, Black Spruce/Jack Pine Cutover

    SciTech Connect

    Margolis, Hank A.

    2016-01-01

    This is the AmeriFlux version of the carbon flux data for the site CA-Qcu Quebec - Eastern Boreal, Black Spruce/Jack Pine Cutover. Site Description - The ground is gently rolling with a weak slope (<5%). In mesic areas (designated as well to moderately well drained areas, according to the Canadian System of Soil Classification (Agriculture Canada Expert Committee on Soil Survey, 1983)), the soil is a ferro-humic to humic podzol covered by an organic layer having an average depth of 26 cm (Fig. 1). In humid areas, the soil is organic (imperfectly to poorly drained) with an average organic layer of 125 cm. Mesic areas accounted for approximately 75% of the total surface area of the footprint and humid areas accounted for 25%. Full-time continuous measurements eneded in 2011. Intermittent measurements are on-going as resources permit.

  8. Total and pyrogenic carbon stocks in black spruce forest floors from eastern Canada

    NASA Astrophysics Data System (ADS)

    Soucemarianadin, Laure; Quideau, Sylvie; MacKenzie, M. Derek; Munson, Alison; Boiffin, Juliette; Bernard, Guy; Wasylishen, Roderick

    2016-04-01

    In boreal forests, pyrogenic carbon (PyC), a by-product of recurrent wildfires, is an important component of the global soil C pool, although precise assessment of boreal PyC stock is scarce. In this study including 14 fire sites spreading over 600 km in the Quebec province, our aim was to better estimate total C stock and PyC stock in forest floors of Eastern Canada boreal forests. We also investigated the environmental conditions controlling the stocks and characterized the composition of the various forest floor layers. We analyzed the forest floor samples that were collected from mesic black spruce sites recently affected by fire (3-5 years) using elemental analysis and solid state 13C nuclear magnetic resonance (NMR) spectroscopy. PyC content was further estimated using a molecular mixing model on the 13C NMR data. Total C stock in forest floors averaged 5.7 ± 2.9 kg C/m2 and PyC stock 0.6 ± 0.3 kg C/m2. Total C stock was under control of the position in the landscape, with a greater accumulation of organic material on northern aspects and lower slope positions. In addition, total stock was significantly higher in spruce-dominated forest floors than in stands where jack pine was dominant. The PyC stock was significantly related to the atomic H/C ratio (R2 = 0.84) of the different organic layers. 13C NMR spectroscopy revealed a large increase in aromatic carbon in the deepest forest floor layer (humified H horizon) at the organic-mineral soil interface. The majority of the PyC stock was located in this horizon and had been formed during past high severity fires rather than during the most recent fire event. Conversely, the superficial "fresh" PyC layer, produced by early-season wildfires in 2005-2007, had NMR spectra fairly similar to unburned forest floors and comparatively low PyC stocks.

  9. Soil-mediated effects of atmospheric deposition on eastern US spruce-fir forests. Book chapter

    SciTech Connect

    Johnson, D.W.; Fernandez, I.J.

    1992-01-01

    The coincident observation of Waldsterben in Germany and red spruce decline in the northeastern U.S. has naturally led to some speculation that similar mechanisms may be involved. In the German situation, soil-mediated hypotheses played (and still play) a major role; namely, soil acidification and aluminum toxicity and base cation deficiencies. In the red spruce case, there has been much concern that cation deficiencies and/or aluminum toxicity may also play a major role. The purpose of this chapter is to: (1) review some of the basic properties of soils, nutrition, and nutrient cycling in spruce-fir and fir ecosystems, both in the polluted and in the relatively unpolluted regions of the U.S. and Canada, and (2) to evaluate several soil acidity-related hypotheses for red spruce decline.

  10. Impact of eastern dwarf mistletoe (Arceuthobium pusillum) on host white spruce (Picea glauca) development, growth and performance across multiple scales.

    PubMed

    Logan, Barry A; Reblin, Jaret S; Zonana, David M; Dunlavey, Ryan F; Hricko, Carolyn R; Hall, Adam W; Schmiege, Stephanie C; Butschek, Ross A; Duran, Kristy L; Emery, R J Neil; Kurepin, Leonid V; Lewis, James D; Pharis, Richard P; Phillips, Nathan G; Tissue, David T

    2013-04-01

    Infection by eastern dwarf mistletoe (Arceuthobium pusillum) modifies needle and branch morphology and hastens white spruce (Picea glauca) mortality. We examined potential causal mechanisms and assessed the impacts of infection-induced alterations to host development and performance across scales ranging from needle hormone contents to bole expansion. Needles on infected branches (IBs) possessed higher total cytokinin (CK) and lower abscisic acid contents than needles on uninfected branches (UBs). IBs exhibited greater xylem growth than same-aged UBs, which is consistent with the promotive effect of CKs on vascular differentiation and organ sink strength. Elevated CK content may also explain the dense secondary and tertiary branching observed at the site of infection, i.e. the formation of 'witches' brooms' with significantly lower light capture efficiencies. Observed hormone perturbations were consistent with higher rates of transpiration, lower water use efficiencies (WUEs) and more negative needle carbon isotope ratios observed for IBs. Observed reductions in needle size allowed IBs to compensate for reduced hydraulic conductivity. Severe infections resulted in dramatically decreased diameter growth of the bole. It seems likely that the modifications to host hormone contents by eastern dwarf mistletoe infection led white spruce trees to dedicate a disproportionate fraction of their photoassimilate and other resources to self-shaded branches with low WUE. This would have decreased the potential for fixed carbon accumulation, generating a decline in the whole-tree resource pool. As mistletoe infections grew in size and the number of IBs increased, this burden was manifested as increasingly greater reductions in bole growth.

  11. Climatic control of tracheid production of black spruce in dense mesic stands of eastern Canada.

    PubMed

    Dufour, Boris; Morin, Hubert

    2013-02-01

    Inferences on climate change effects are reliable only if they are based on a causal relationship rather than simple statistical predictive capacity. To assess for causal links between climate and mature black spruce (Picea mariana (Mills.) BSP) radial growth, we combined the use of wood anatomy, cambium phenology, climate and soil measurements (air temperature and humidity, precipitations, soil temperature and water content, photosynthetically active radiation), and a model selection approach proceeding backwards from a full model. Results show that the number of tracheids is responsible for 88% of the variation in ring width whereas mean tracheid diameter accounts for the remaining 12%. The number of tracheids produced depends on factors related to photosynthesis during tracheid production, i.e., daily light intensity and maximum temperature between the day of initiation and the day of cessation of tracheid production, plus soil temperature during August of the previous year which is an important period for determining the number of new needles produced. It is also important to consider duration of the period for tracheid production. These results imply that short-term climate change should increase black spruce radial growth. They also suggest that the typical use of post-growth ring width sampling individually linked to air temperature and precipitations is not sufficient to infer climate change effects accurately on radial growth where there is no strong single climatic limitation but multiple limitations instead. PMID:23355635

  12. BVOC emission in Norway spruce: the effect of stand structure, high temperature and ozone levels.

    NASA Astrophysics Data System (ADS)

    Pallozzi, Emanuele; Guidolotti, Gabriele; Večeřová, Kristýna; Esposito, Raffaela; Lusini, Ilaria; Juráň, Stanislav; Urban, Otmar; Calfapietra, Carlo

    2015-04-01

    Norway spruce (Picea abies L.) is a widely distributed conifer species in the boreal zone and mountain areas of central Europe and is a moderate emitter of volatile organic compounds (BVOC). Although the vaporization and diffusion processes from resin ducts were generally considered to be the main processes for monoterpene emissions in conifers, recently it has been showed that a significant portion (up to one third) of monoterpene emissions of Norway spruce can originate from novel biosynthesis, thus depending on photosynthetic processes. For this reason, both biosynthesis and emission are strongly influenced by the environment and the stand structure. They increase with both increasing light and temperature during the warmer periods, although those are the periods with the higher ozone concentration that usually act as an inhibitor of both assimilation and isoprenoids synthesis and emission. On the other hand, stand structure can play an important role, because the photosynthetic capacity is influenced by temperature and light conditions through the canopy. In order to assess the effects of stand structure, temperature and ozone on isoprenoids emission of Norway spruce we carried out field and laboratory experiments. In the experimental field campaigns we measured: assimilation and BVOC emission from needles of sun and shade layers within the canopy of the spruce forest present at the Bily Kriz experimental research site (Moravian-Silesian Beskydy Mountains, 49° 33' N, 18° 32' E, NE of Czech Republic, 908 m a.s.l.). Moreover in the same layers we measured continuously concentration of BVOCs in the air using a PTR-TOF-MS. In laboratory we analyzed the effects of short-term exposure to high temperature and high ozone concentrations on branches of spruce trees collected at the Bily Kriz experimental research site. Preliminary results show that in Norway spruce both stand structure and environmental conditions influenced the gas exchange and BVOC emission rates

  13. Response of aluminum solubility to elevated nitrification in soil of a red spruce stand in eastern Maine

    USGS Publications Warehouse

    Lawrence, G.B.; David, M.B.

    1997-01-01

    Elevated concentrations of soluble Al can impair tree growth and be toxic to aquatic biota, but effects of acidic deposition on Al solubility in forest soils are only partially understood because of complex interactions with H+ and organic matter. We therefore evaluated Al solubility in two red spruce stands in eastern Maine, one of which received dry (NH4)2SO4 at a rate of 1800 equiv ha-1 yr-1 during 19891995. Samples of soil (Spodosol Oa and Bh horizons) and soil solution were collected on five dates from 1992 to 1995. The treatment elevated nitrification, causing an increase in acid input that led to inorganic Al concentrations of greater than 60 ??mol L-1 in both the Oa and Bh horizons. Solubility of Al was also lower in the Bh horizon of the treated stand than in the reference stand, a response related to higher DOC concentrations in the treated stand. Concentrations of CuCl2 and pyrophosphate-extractable Al were higher in the Oa horizon of the treated watershed than the reference stand, a result of accelerated weathering of mineral particles caused by lower solution pH in the treated stand (3.47) than in the reference stand (3.69). Dissolved Al concentrations in these soils are the result of complex mechanisms through which mineral matter, organic matter, and pH interact to control Al solubility; mechanisms that are not incorporated in current Al solubility models.

  14. SABATH methyltransferases from white spruce (Picea glauca): gene cloning, functional characterization and structural analysis.

    PubMed

    Zhao, Nan; Boyle, Brian; Duval, Isabelle; Ferrer, Jean-Luc; Lin, Hong; Seguin, Armand; MacKay, John; Chen, Feng

    2009-07-01

    Known members of the plant SABATH family of methyltransferases have important biological functions by methylating hormones, signalling molecules and other metabolites. While all previously characterized SABATH genes were isolated from angiosperms, in this article, we report on the isolation and functional characterization of SABATH genes from white spruce (Picea glauca [Moench] Voss), a gymnosperm. Through EST database search, three genes that encode proteins significantly homologous to known SABATH proteins were identified from white spruce. They were named PgSABATH1, PgSABATH2 and PgSABATH3, respectively. Full length cDNAs of these three genes were cloned and expressed in Escherichia coli. The E. coli-expressed recombinant proteins were tested for methyltransferase activity with a large number of compounds. While no activity was detected for PgSABATH2 and PgSABATH3, PgSABATH1 displayed the highest level of catalytic activity with indole-3-acetic acid (IAA). PgSABATH1 was, therefore, renamed PgIAMT1. Under steady-state conditions, PgIAMT1 exhibited apparent Km values of 18.2 microM for IAA. Homology-based structural modelling of PgIAMT1 revealed that the active site of PgIAMT1 is highly similar to other characterized IAMTs from angiosperms. PgIAMT1 showed expression in multiple tissues, with the highest level of expression detected in embryonic tissues. During somatic embryo maturation, a significant reduction in PgIAMT1 transcript levels was observed when developing cotyledons become apparent which is indicative of mature embryos. The biological roles of white spruce SABATH genes, especially those of PgIAMT1, and the evolution of the SABATH family are discussed.

  15. Structure and Carbon Distribution of Well- and Poorly-Drained Black Spruce Fire Chronosequences

    NASA Astrophysics Data System (ADS)

    Wang, C.; Bond-Lamberty, B.; Gower, S. T.

    2002-12-01

    The objectives of this study were to: (1) examine structure changes along a boreal black spruce (Picea mariana (Mill.) BSP) fire chronosequence in northern Manitoba, Canada; (2) quantify temporal dynamics in C content across the chronosequence; and (3) compare the C distribution for these stands on contrasting soil drainages. The experimental design was a well-drained (dry) and poorly-drained (wet) black spruce fire chronosequences comprised of seven stands that burned between 1998 and 1870. Tree species diversity was greater in the younger stands, and by 71 years after fire black spruce comprised >88% and 83% of total basal area for the dry and wet stands, respectively. Feather moss replaced sphagnum during succession in the dry stands, whereas there were no clear patterns in the wet chronosequence. Total vegetation C content (aboveground + belowground) increased with forest succession, ranging from 1.3 to 83.3 t C ha-1 for the dry stands and 0.6 to 37.4 t C ha-1 for the wet stands. Mean annual aboveground C accumulation rates (Δ CA) peaked in the 71- and 36-year-old dry and wet stands, respectively. Overstory and live moss C contents increased with stand age. The total root : shoot ratio was fairly consistent (0.235 +/- 0.046) after 37 years following fire. The basal area, vegetation C content, and Δ CA were significantly greater (α = 0.05) in the dry than wet stands, but the C contents in bryophyte and forest floor were significantly less in the dry stands. Our results emphasize the need to incorporate disturbance and soil drainage in large-scale boreal forest C models.

  16. Crustal structure beneath Eastern Greenland

    NASA Astrophysics Data System (ADS)

    Reiche, S.; Thybo, H.; Kaip, G.; Skjoth Bruun, A.; Reid, I.; Chemia, Z.; Greschke, B.

    2011-12-01

    The conjugate Atlantic passive margins of western Norway and eastern Greenland are characterized by the presence of coast-parallel mountain ranges with peak elevations of more than 3.5 km close to Scoresby Sund in Eastern Greenland. Knowledge about crustal thickness and composition below these mountain belts is needed for assessing the isostatic balance of the crust and to gain insight into possible links between crustal composition, rifting history and present-day topography of the North Atlantic Region. However, the acquisition of geophysical data onshore Greenland is logistically complicated by the presence of an up to 4 km thick ice sheet, permanently covering the largest part of the land mass. Hence previous seismic surveys have only been carried out offshore and near the coast of Greenland, where little information about the continental part of the crust could be gained. To get insight into crustal thickness and composition below the Greenland ice sheet, the TopoGreenland project collects the first ever seismic data onshore Greenland. Wide-angle data was acquired along an EW-trending profile, extending 350 km inland from the approximate edge of the stable ice cap near Scoresby Sund. Data is recorded by 350 Reftek Texan receivers for 10 equidistant shot points along the profile. We use forward ray tracing modelling to construct a two-dimensional velocity model from the observed travel times. These results show the first images of the subsurface velocity structure beneath the Greenland ice sheet and provide a link between the composition of the crust and the present-day topography of Greenland.

  17. The effects of partial cutting on stand structure and growth of western hemlock-Sitka spruce stands in southeast Alaska

    USGS Publications Warehouse

    Deal, R.L.; Tappeiner, J.C.

    2002-01-01

    The effects of partial cutting on species composition, new and residual-tree cohorts, tree size distribution, and tree growth was evaluated on 73 plots in 18 stands throughout southeast Alaska. These partially cut stands were harvested 12-96 years ago, when 16-96% of the former stand basal area was removed. Partial cutting maintained stand structures similar to uncut old-growth stands, and the cutting had no significant effects on tree species composition. The establishment of new-tree cohorts was positively related to the proportion of basal-area cut. The current stand basal area, tree species composition, and stand growth were significantly related to trees left after harvest (p < 0.001). Trees that were 20-80 cm dbh at the time of cutting had the greatest tree-diameter and basal-area growth and contributed the most to stand growth. Diameter growth of Sitka spruce and western hemlock was similar, and the proportion of stand basal-area growth between species was consistent for different cutting intensities. Concerns about changing tree species composition, lack of spruce regeneration, and greatly reduced stand growth and vigor with partial cuts were largely unsubstantiated. Silvicultural systems based on partial cutting can provide rapidly growing trees for timber production while maintaining complex stand structures with mixtures of spruce and hemlock trees similar to oldgrowth stands.

  18. Diurnal changes in the dielectric properties and water status of eastern hemlock and red spruce from Howland, ME

    NASA Technical Reports Server (NTRS)

    Salas, W. A.; Ranson, K. J.; Rock, B. N.; Moss, D. M.

    1991-01-01

    The diurnal characteristics of microwave dielectric properties and water potential of two conifer species were investigated in July and September, 1990. P-band and C-band radial dielectric profiles of hemlock and red spruce, as well as hemlock diurnal water potential and dielectric profiles, are presented. The resulting radial dielectric profiles matched the regions of the functional sapwood (water transport component of the active xylem) in both species such that the sapwood was characterized by a higher dielectric than the bark and heartwood tissues. This is probably due to characteristic differences in the water content of each tissue. As the hemlocks progressed through their diurnal water potential pattern, the dielectric profile remained static until mid-afternoon. As the tension in the water column relaxed (2 to 3 bars) the dielectric constant decreased by 30 to 40 percent. There are several possible explanations for this phenomenon, and these may relate to the dependency of the dielectric measurements on temperature, salinity, and volumetric water content.

  19. Organised Motion in a Tall Spruce Canopy: Temporal Scales, Structure Spacing and Terrain Effects

    NASA Astrophysics Data System (ADS)

    Thomas, Christoph; Foken, Thomas

    2007-01-01

    This study investigates the organised motion near the canopy-atmosphere interface of a moderately dense spruce forest in heterogeneous, complex terrain. Wind direction is used to assess differences in topography and surface properties. Observations were obtained at several heights above and within the canopy using sonic anemometers and fast-response gas analysers over the course of several weeks. Analysed variables include the three-dimensional wind vector, the sonic temperature, and the concentration of carbon dioxide. Wavelet analysis was used to extract the organised motion from time series and to derive its temporal scales. Spectral Fourier analysis was deployed to compute power spectra and phase spectra. Profiles of temporal scales of ramp-like coherent structures in the vertical and longitudinal wind components showed a reversed variation with height and were of similar size within the canopy. Temporal scales of scalar fields were comparable to those of the longitudinal wind component suggesting that the lateral scalar transport dominates. The existence of a 1 power law in the longitudinal power spectra was confirmed for a few cases only, with a majority showing a clear 5/3 decay. The variation of effective scales of organised motion in the longitudinal velocity and temperature were found to vary with atmospheric stability, suggesting that both Kelvin-Helmholtz instabilities and attached eddies dominate the flow with increasing convectional forcing. The canopy mixing-layer analogy was observed to be applicable for ramp-like coherent structures in the vertical wind component for selected wind directions only. Departures from the prediction of m = Λ w L {/s -1} = 8 10 (where Λ w is the streamwise spacing of coherent structures in the vertical wind w and L s is a canopy shear length scale) were caused by smaller shear length scales associated with large-scale changes in the terrain as well as the vertical structure of the canopy. The occurrence of linear

  20. Effects of warming on the structure and function of a boreal black spruce forest

    SciTech Connect

    Stith T.Gower

    2010-03-03

    A strong argument can be made that there is a greater need to study the effect of warming on boreal forests more than on any other terrestrial biome. Boreal forests, the second largest forest biome, are predicted to experience the greatest warming of any forest biome in the world, but a process-based understanding of how warming will affect the structure and function of this economically and ecologically important forest biome is lacking. The effects of warming on species composition, canopy structure and biogeochemical cycles are likely to be complex; elucidating the underlying mechanisms will require long-term whole-ecosystem manipulation to capture all the complex feedbacks (Shaver et al. 2000, Rustad et al. 2001, Stromgren 2001). The DOE Program for Ecosystem Research funded a three year project (2002-2005) to use replicated heated chambers on soil warming plots in northern Manitoba to examine the direct effects of whole-ecosystem warming. We are nearing completion of our first growing season of measurements (fall 2004). In spite of the unforeseen difficulty of installing the heating cable, our heating and irrigation systems worked extremely well, maintaining environmental conditions within 5-10% of the specified design 99% of the time. Preliminary data from these systems, all designed and built by our laboratory at the University of Wisconsin, support our overall hypothesis that warming will increase the carbon sink strength of upland boreal black spruce forests. I request an additional three years of funding to continue addressing the original objectives: (1) Examine the effect of warming on phenology of overstory, understory and bryophyte strata. Sap flux systems and dendrometer bands, monitored by data loggers, will be used to quantify changes in phenology and water use. (2) Quantify the effects of warming on nitrogen and water use by overstory, understory and bryophytes. (3) Compare effects of warming on autotrophic respiration and above- and belowground

  1. Can nutrient limitations explain low and declining white spruce growth near the Arctic treeline in the eastern Brooks Range, Alaska?

    NASA Astrophysics Data System (ADS)

    Ellison, S.; Sullivan, P. F.

    2014-12-01

    The position of the Arctic treeline is of critical importance for global carbon cycling and surface energy budgets. However, controls on tree growth at treeline remain uncertain. In the Alaskan Brooks Range, 20th century warming has caused varying growth responses among treeline trees, with trees in the west responding positively, while trees in the east have responded negatively. The prevailing explanation of this trend ascribes the negative growth response to warming-induced drought stress in the eastern Brooks Range. However, recent measurements of carbon isotope discrimination in tree rings, xylem sap flow and needle gas exchange suggest that drought stress cannot explain these regional growth declines. Additionally, evidence from the western Brooks Range suggests that nutrient availability, rather than drought stress, may be the proximate control on tree growth. In this study, we investigated the hypothesis that low and declining growth of eastern Brooks Range trees is due to low and declining soil nutrient availability, which may continue to decrease with climate change as soils become drier and microbial activity declines. We compared microclimate, tree performance, and a wide range of proxies for soil nutrient availability in four watersheds along a west-east transect in the Brooks Range during the growing seasons of 2013 and 2014. We hypothesized that soil nutrient availability would track closely with the strong west-east precipitation gradient, with higher rainfall and greater soil nutrient availability in the western Brooks Range. We expected to find that soil water contents in the west are near optimum for nitrogen mineralization, while those in the east are below optimum. Needle nitrogen concentration, net photosynthesis, branch extension growth, and growth in the main stem are expected to decline with the hypothesized decrease in soil nutrient availability. The results of our study will elucidate the current controls on growth of trees near the

  2. Crustal Structure in Central-Eastern Greenland

    NASA Astrophysics Data System (ADS)

    Shulgin, A.; Thybo, H.

    2013-12-01

    We present the seismic structure in the interior of Greenland based on the first measurements by the seismic refraction/wide angle reflection method. Previous seismic surveys have only been carried out offshore and near the coast of Greenland, where the crustal structure is affected by oceanic break-up and may not be representative of the interior of the island. Acquisition of geophysical data onshore Greenland is logistically complicated by the presence of an up to 3.4 km thick ice sheet, permanently covering most of the land mass. The seismic data was acquired by a team of six people during a two-month long experiment in summer of 2011 on the ice cap in the interior of central-eastern Greenland. The EW-trending profile extends 310 km inland from the approximate edge of the stable ice cap near Scoresby Sund across the centre of the ice cap. The planned extension of the profile by use of OBSs and air gun shooting in Scoresbysund Fjord to the east coast of Greenland was unfortunately cancelled, because navigation was prevented by ice drift. 350 Reftek Texan receivers recorded high-quality seismic data from 8 equidistant shots along the profile. Explosive charge sizes were 1 ton at the ends and ca. 500 kg along the profile, loaded with about 100 kg at 35-85 m depth in individual boreholes. Two-dimensional velocity model based on forward ray tracing and tomography modelling shows a decrease of crustal thickness from 47 km below the centre of Greenland in the western part to 40 km in the eastern part of the profile. Earlier studies show that crustal thickness further decreases eastward to ca. 30 km below the fjord system, but details of the changes are unknown. Relatively high lower crustal velocities (Vp 6.8 - 7.3) in the western part of the TopoGreenland profile may indicate past collision tectonics or may be related or to the passage of the Iceland mantle plume. The origin of the pronounced circum-Atlantic mountain ranges in Norway and eastern Greenland, which have

  3. Norway spruce embryogenesis: changes in carbohydrate profile, structural development and response to polyethylene glycol.

    PubMed

    Hudec, Lukáš; Konrádová, Hana; Hašková, Anna; Lipavská, Helena

    2016-05-01

    Two unrelated, geographically distinct, highly embryogenic lines of Norway spruce (Picea abies (L.) Karst.) were analysed to identify metabolic traits characteristic for lines with good yields of high-quality embryos. The results were compared with corresponding characteristics of a poorly productive line (low embryo yield, scarce high-quality embryos). The following carbohydrate profiles and spectra during maturation, desiccation and germination were identified as promising characteristics for line evaluation: a gradual decrease in total soluble carbohydrates with an increasing sucrose : hexose ratio during maturation; accumulation of raffinose family oligosaccharides resulting from desiccation and their rapid degradation at the start of germination; and a decrease in sucrose, increase in hexoses and the appearance of pinitol with proceeding germination. We propose that any deviation from this profile in an embryonic line is a symptom of inferior somatic embryo development. We further propose that a fatty acid spectrum dominated by linoleic acid (18 : 2) was a common feature of healthy spruce somatic embryos, although it was quite different from zygotic embryos mainly containing oleic acid (18 : 1). The responses of the lines to osmotic stress were evaluated based on comparison of control (without osmoticum) and polyethylene glycol (PEG)-exposed (PEG 4000) variants. Although genetically distinct, both highly embryogenic lines responded in a very similar manner, with the only difference being sensitivity to high concentrations of PEG. At an optimum PEG concentration (3.75 and 5%), which was line specific, negative effects of PEG on embryo germination were compensated for by a higher maturation efficiency so that the application of PEG at an appropriate concentration improved the yield of healthy germinants per gram of initial embryonal mass and accelerated the process. Polyethylene glycol application, however, resulted in no improvement of the poorly

  4. Geologic and structural map of eastern Asia

    SciTech Connect

    Letouzey, J.; Sage, L.

    1986-07-01

    A synthesis of the onshore and offshore geologic data of eastern Asia, prepared by the Institut Francais du Petrole (IFP), has allowed the construction of geologic and structural maps for this region. These maps include three color sheets (scale = 1:2.5 million) and three plates of geologic and structural cross sections. Located between lat. 4/sup 0/ and 35/sup 0/N, and long. 106/sup 0/ and 132/sup 0/E, the maps cover the following geographic areas: East and South China Sea, Sulu Sea, West Philippine basin and onshore neighboring terrains, Kyushu and Ryukyu Islands, the China margin, Taiwan Island, Vietnam, North West Borneo, and the Philippines. The maps synthesize seismic interpretations, oil well data, geologic work in south Japan, Taiwan, Borneo, and the Philippines, and recent data published between 1976 and 1985. Twenty-four geologic cross sections (scale = 1:1.25 million, vertical exaggeration x 6) intersect ocean margins, important basins, and the different structural domains. They are based on seismic profiles, well data, and available onshore and offshore geologic data. These cross sections show basement composition and structures, different tectonic and sedimentary domains, and the structure and thickness of different sedimentary deposits (such as age, unconformities, and geologic structures). Maps and cross sections will be published in early 1987.

  5. High Resolution Velocity Structure in Eastern Turkey

    SciTech Connect

    Pasyanos, M; Gok, R; Zor, E; Walter, W

    2004-09-03

    We investigate the crustal and upper mantle structure of eastern Turkey where the Anatolian, Arabian and Eurasian Plates meet and form a complex tectonic structure. The Bitlis suture is a continental collision zone between the Anatolian plateau and the Arabian plate. Broadband data available through the Eastern Turkey Seismic Experiment (ETSE) provided a unique opportunity for studying the high resolution velocity structure. Zor et al. found an average 46 km thick crust in Anatolian plateau using six-layered grid search inversion of the ETSE receiver functions. Receiver functions are sensitive to the velocity contrast of interfaces and the relative travel time of converted and reverberated waves between those interfaces. The interpretation of receiver function alone with many-layered parameterization may result in an apparent depth-velocity tradeoff. In order to improve previous velocity model, we employed the joint inversion method with many layered parameterization of Julia et al. (2000) to the ETSE receiver functions. In this technique, the receiver function and surface-wave observations are combined into a single algebraic equation and each data set is weighted by an estimate of the uncertainty in the observations. We consider azimuthal changes of receiver functions and have stacked them into different groups. We calculated the receiver functions using iterative time-domain deconvolution technique and surface wave group velocity dispersion curves between 10-100 sec. We are making surface wave dispersion measurements at the ETSE stations and have incorporated them into a regional group velocity model. Preliminary results indicate a strong trend in the long period group velocity in the northeast. This indicates slow upper mantle velocities in the region consistent with Pn, Sn and receiver function results. We started with both the 1-D model that is obtained with the 12 tones dam explosion shot data recorded by ETSE network and the existing receiver function

  6. Specific impacts of beech and Norway spruce on the structure and diversity of the rhizosphere and soil microbial communities

    PubMed Central

    Uroz, S.; Oger, P.; Tisserand, E.; Cébron, A.; Turpault, M.-P.; Buée, M.; De Boer, W.; Leveau, J. H. J.; Frey-Klett, P.

    2016-01-01

    The impacts of plant species on the microbial communities and physico-chemical characteristics of soil are well documented for many herbs, grasses and legumes but much less so for tree species. Here, we investigate by rRNA and ITS amplicon sequencing the diversity of microorganisms from the three domains of life (Archaea, Bacteria and Eukaryota:Fungi) in soil samples taken from the forest experimental site of Breuil-Chenue (France). We discovered significant differences in the abundance, composition and structure of the microbial communities associated with two phylogenetically distant tree species of the same age, deciduous European beech (Fagus sylvatica) and coniferous Norway spruce (Picea abies Karst), planted in the same soil. Our results suggest a significant effect of tree species on soil microbiota though in different ways for each of the three microbial groups. Fungal and archaeal community structures and compositions are mainly determined according to tree species, whereas bacterial communities differ to a great degree between rhizosphere and bulk soils, regardless of the tree species. These results were confirmed by quantitative PCR, which revealed significant enrichment of specific bacterial genera, such as Burkholderia and Collimonas, known for their ability to weather minerals within the tree root vicinity. PMID:27302652

  7. Specific impacts of beech and Norway spruce on the structure and diversity of the rhizosphere and soil microbial communities.

    PubMed

    Uroz, S; Oger, P; Tisserand, E; Cébron, A; Turpault, M-P; Buée, M; De Boer, W; Leveau, J H J; Frey-Klett, P

    2016-01-01

    The impacts of plant species on the microbial communities and physico-chemical characteristics of soil are well documented for many herbs, grasses and legumes but much less so for tree species. Here, we investigate by rRNA and ITS amplicon sequencing the diversity of microorganisms from the three domains of life (Archaea, Bacteria and Eukaryota:Fungi) in soil samples taken from the forest experimental site of Breuil-Chenue (France). We discovered significant differences in the abundance, composition and structure of the microbial communities associated with two phylogenetically distant tree species of the same age, deciduous European beech (Fagus sylvatica) and coniferous Norway spruce (Picea abies Karst), planted in the same soil. Our results suggest a significant effect of tree species on soil microbiota though in different ways for each of the three microbial groups. Fungal and archaeal community structures and compositions are mainly determined according to tree species, whereas bacterial communities differ to a great degree between rhizosphere and bulk soils, regardless of the tree species. These results were confirmed by quantitative PCR, which revealed significant enrichment of specific bacterial genera, such as Burkholderia and Collimonas, known for their ability to weather minerals within the tree root vicinity.

  8. Specific impacts of beech and Norway spruce on the structure and diversity of the rhizosphere and soil microbial communities.

    PubMed

    Uroz, S; Oger, P; Tisserand, E; Cébron, A; Turpault, M-P; Buée, M; De Boer, W; Leveau, J H J; Frey-Klett, P

    2016-01-01

    The impacts of plant species on the microbial communities and physico-chemical characteristics of soil are well documented for many herbs, grasses and legumes but much less so for tree species. Here, we investigate by rRNA and ITS amplicon sequencing the diversity of microorganisms from the three domains of life (Archaea, Bacteria and Eukaryota:Fungi) in soil samples taken from the forest experimental site of Breuil-Chenue (France). We discovered significant differences in the abundance, composition and structure of the microbial communities associated with two phylogenetically distant tree species of the same age, deciduous European beech (Fagus sylvatica) and coniferous Norway spruce (Picea abies Karst), planted in the same soil. Our results suggest a significant effect of tree species on soil microbiota though in different ways for each of the three microbial groups. Fungal and archaeal community structures and compositions are mainly determined according to tree species, whereas bacterial communities differ to a great degree between rhizosphere and bulk soils, regardless of the tree species. These results were confirmed by quantitative PCR, which revealed significant enrichment of specific bacterial genera, such as Burkholderia and Collimonas, known for their ability to weather minerals within the tree root vicinity. PMID:27302652

  9. Crustal structure beneath Central-Eastern Greenland

    NASA Astrophysics Data System (ADS)

    Reiche, S.; Thybo, H.; Reid, I.; Shulgin, A.

    2012-04-01

    Acquisition of geophysical data onshore Greenland is logistically complicated by the presence of an up to 3.4 km thick ice sheet, permanently covering most of the land mass. Previous seismic surveys have only been carried out offshore and near the coast of Greenland, where little information about the continental part of the crust could be gained. Aiming to improve our understanding about crustal thickness and composition below the Greenland ice cap, the TopoGreenland project was initiated to collect the first ever controlled source seismic data onshore Greenland. Wide-angle seismic data were acquired along an EW-trending profile, extending 310 km inland from the approximate edge of the stable ice cap near Scoresby Sund across the centre of the ice cap. In total, 348 Reftek Texan receivers recorded high-quality data from 8 equidistant shot points along the profile. Based on forward ray tracing modelling, a two-dimensional velocity model provides the first insight into the velocity structure beneath the Greenland ice sheet. Modelling results indicate a decrease of crustal thickness from 50 km below the centre of Greenland to 42 km in the eastern part of the profile. Relatively high lower crustal velocities (Vp 6.8 - 7.3) in the western part of the model may be related to past collisional tectonic events or to the passage of the Iceland mantle plume.

  10. Red spruce dynamics in an old southern Appalachian forest

    USGS Publications Warehouse

    Busing, R.T.

    2004-01-01

    By the late 1980s the composition and structure of forest stands in the southern Appalachian spruce-fir zone were altered by insect infestations to Fraser fir. The response of red spruce, the sole remaining coniferous forest dominant, to this disturbance was followed over twenty years (1983-2003) in an old spruce-fir forest at Mt. Collins, Great Smoky Mountains National Park. Although diameter growth of canopy red spruce (>30 cm dbh) at six plot sites was considerable (mean 10-yr increment 2.1 cm; 1993-2003), red spruce mortality increased sharply (mean 4% yr-1; 1993-2003). Wind-related mortality of canopy red spruce was substantial after the loss of Fraser fir from the canopy circa 1985 (>70% of the dead spruce had broken or uprooted boles; 1983-2003). Wind damage to red spruce was observed at most plot sites, but it was most pronounced on exposed topographic positions, where canopy gap expansion was extensive. The elevated mortality of red spruce at Mt. Collins was not associated with reduced diameter growth. Altered canopy structure has left large red spruce vulnerable to high winds. With the loss of canopy fir and the subsequent increase in mortality of canopy spruce, total live basal area has declined to about half of its pre-disturbance level.

  11. [Species composition and community structure of a spruce-fir forest and a larch forest on the northern slope of Changbai Mountains, Northeast China].

    PubMed

    Kuang, Xu; Xing, Ding-Liang; Zhang, Zhao-Chen; Song, Hou-Juan; Wang, Yun-Yun; Fang, Shuai; Yuan, Zuo-Qiang; Ye, Ji; Lin, Fei; Wang, Xu-Gao; Hao, Zhan-Qing

    2014-08-01

    Spruce-fir forest is the best protected forest vegetation, while larch forest is intrazonal vegetation on the northern slope of Changbai Mountains. To further understand their species composition and community structure, we established a 4 hm2 forest permanent plot in each of these two forests in 2010. All free-standing plant species with DBH (diameter at breast height) ≥ 1 cm were mapped, tagged, and identified to species. The results showed that there were 9257 stems belonging to 8640 genotype individuals, 22 species, 6 genera and 12 families in the spruce-fir forest plot, while 4060 stems belonging to 3696 genotype individuals, 22 species, 8 genera and 16 families in the larch forest plot. Species composition in the two plots was very similar. Most of the species belonged to the Changbai Mountains plant flora. The analysis of species' importance values showed that there were dominant species in both communities. The spruce-fir forest was dominated by Abies nephrolepis and Larix olgensis, whose importance values accounted for 38.7% and 23.9% of the sum of importance values over all species in the plot, respectively. The larch forest was dominated solely by L. olgensis, whose importance value accounted for 61.9% of the sum of importance values over all species in the plot. Both forests were in good condition of regeneration and showed a reversed 'J' type in tree size distributions, at community level. However, different species showed different shapes in size distribution in the two forests. A. nephrolepis showed a reversed 'J' type size distribution in the spruce-fir forest, while L. olgensis with DBH ≥ 10 cm showed a hump-shaped distribution in the larch forest. Spatial distribution patterns of the main species changed differently with size class and spatial scales. Common species had different spatial distribution patterns in the two plots. PMID:25509062

  12. Identifying the pollen of an extinct spruce species in the Late Quaternary sediments of the Tunica Hills region, south-eastern United States

    USGS Publications Warehouse

    Luke Mander,; Jacklyn Rodriguez,; Pietra G. Mueller,; Jackson, Stephen T.; Surangi W. Punyasena,

    2014-01-01

    Late Quaternary fluvial deposits in the Tunica Hills region of Louisiana and Mississippi are rich in spruce macrofossils of the extinct species Picea critchfieldii, the one recognized plant extinction of the Late Quaternary. However, the morphology of P. critchfieldii pollen is unknown, presenting a barrier to the interpretation of pollen spectra from the last glacial of North America. To address this issue, we undertook a morphometric study of Picea pollen from Tunica Hills. Morphometric data, together with qualitative observations of pollen morphology using Apotome fluorescence microscopy, indicate that Picea pollen from Tunica Hills is morphologically distinct from the pollen of P. glauca, P. mariana and P. rubens. Measurements of grain length, corpus width and corpus height indicate that Picea pollen from Tunica Hills is larger than the pollen of P. mariana and P. rubens, and is slightly larger than P. glauca pollen. We argue that the morphologically distinctive Tunica Hills Picea pollen was probably produced by the extinct spruce species P. critchfieldii. These morphological differences could be used to identify P. critchfieldii in existing and newly collected pollen records, which would refine its paleoecologic and biogeographic history and clarify the nature and timing of its extinction in the Late Quaternary.

  13. Legacy of Pre-Disturbance Spatial Pattern Determines Early Structural Diversity following Severe Disturbance in Montane Spruce Forests

    PubMed Central

    Bače, Radek; Svoboda, Miroslav; Janda, Pavel; Morrissey, Robert C.; Wild, Jan; Clear, Jennifer L.; Čada, Vojtěch; Donato, Daniel C.

    2015-01-01

    Background Severe canopy-removing disturbances are native to many temperate forests and radically alter stand structure, but biotic legacies (surviving elements or patterns) can lend continuity to ecosystem function after such events. Poorly understood is the degree to which the structural complexity of an old-growth forest carries over to the next stand. We asked how pre-disturbance spatial pattern acts as a legacy to influence post-disturbance stand structure, and how this legacy influences the structural diversity within the early-seral stand. Methods Two stem-mapped one-hectare forest plots in the Czech Republic experienced a severe bark beetle outbreak, thus providing before-and-after data on spatial patterns in live and dead trees, crown projections, down logs, and herb cover. Results Post-disturbance stands were dominated by an advanced regeneration layer present before the disturbance. Both major species, Norway spruce (Picea abies) and rowan (Sorbus aucuparia), were strongly self-aggregated and also clustered to former canopy trees, pre-disturbance snags, stumps and logs, suggesting positive overstory to understory neighbourhood effects. Thus, although the disturbance dramatically reduced the stand’s height profile with ~100% mortality of the canopy layer, the spatial structure of post-disturbance stands still closely reflected the pre-disturbance structure. The former upper tree layer influenced advanced regeneration through microsite and light limitation. Under formerly dense canopies, regeneration density was high but relatively homogeneous in height; while in former small gaps with greater herb cover, regeneration density was lower but with greater heterogeneity in heights. Conclusion These findings suggest that pre-disturbance spatial patterns of forests can persist through severe canopy-removing disturbance, and determine the spatial structure of the succeeding stand. Such patterns constitute a subtle but key legacy effect, promoting structural

  14. Eastern Boundary Effects on General Circulation Structure

    NASA Technical Reports Server (NTRS)

    Dewar, William K.

    1997-01-01

    My NASA proposal included plans to examine the dynamics of the eastern oceanic boundary, with a view towards those processes important to the interior. Several relevant tasks have been completed and either have appeared or will appear soon in the refereed literature.

  15. Genetic structure of Sakhalin spruce (Picea glehnii) in northern Japan and adjacent regions revealed by nuclear microsatellites and mitochondrial gene sequences.

    PubMed

    Aizawa, Mineaki; Yoshimaru, Hiroshi; Takahashi, Makoto; Kawahara, Takayuki; Sugita, Hisashi; Saito, Hideyuki; Sabirov, Renat N

    2015-01-01

    The genetic structure of Sakhalin spruce (Picea glehnii) was studied across the natural range of the species, including two small isolated populations in south Sakhalin and Hayachine, by using six microsatellite loci and maternally inherited mitochondrial gene sequences. We also analyzed P. jezoensis, a sympatric spruce in the range. Genetic diversity of P. glehnii was higher in central Hokkaido and the lowest in the Hayachine. Bayesian clustering and principal coordinate analysis by using the microsatellites indicated that the Hayachine was clearly distinct from other populations, implying that it had undergone strong genetic drift since the last glacial period. P. glehnii harbored four mitochondrial haplotypes, two of which were shared with P. jezoensis. One of the two was observed without geographical concentration, suggesting its derivation from ancestral polymorphism. Another was observed in south Sakhalin and in P. jezoensis across Sakhalin. The Bayesian clustering--by using four microsatellite loci, including P. jezoensis populations--indicated unambiguous species delimitation, but with possible admixture of P. jezoensis genes into P. glehnii in south Sakhalin, where P. glehnii is abundantly overwhelmed by P. jezoensis; this might explain the occurrence of introgression of the haplotype of P. jezoensis into P. glehnii.

  16. Crustal structure and active tectonics in the Eastern Alps

    NASA Astrophysics Data System (ADS)

    Brückl, E.; Behm, M.; Decker, K.; Grad, M.; Guterch, A.; Keller, G. R.; Thybo, H.

    2010-04-01

    During the last decade, a series of controlled source seismic experiments brought new insight into the crustal and lithospheric structure of the Eastern Alps and their adjacent tectonic provinces. A fragmentation of the lithosphere into three blocks, Europe (EU), Adria (AD), and the new Pannonian fragment (PA), was interpreted and a triple junction was inferred. The goal of this study has been to relate these deep crustal structures to active tectonics. We used elastic plate modeling to reconsider the Moho fragmentation. We interpret subduction of EU below AD and PA from north to south and underthusting of AD mantle below PA from southwest to northeast. The Moho fragmentation correlates well with major upper crustal structures and is supported by gravity, seismic, and geodetic data. An analysis of crustal thickening suggests that active convergence is associated with continued thrusting and lateral extrusion in the central Eastern Alps and thickening of the Adriatic indenter under the Southern Alps. According to the velocity relations at the triple junction, PA moves relative to EU and AD along ENE and SE striking faults, mainly by strike slip. An eastward directed extensional component is compensated by the lateral extrusion of the central Eastern Alps. The Periadriatic (Insubric) line east of the triple junction and the mid-Hungarian fault zone have relatively recently lost their role as first-order active structures. We favor the idea that the Pannonian fragment and the TISZA block merged to a "soft" microplate surrounded by the Eastern and Southern Alpine, Carpathian, and Dinaric orogens.

  17. Non-structural carbohydrate status in Norway spruce buds in the context of annual bud structural development as affected by acidic pollution.

    PubMed

    Svobodová; Lipavská; Albrechtová

    2000-06-01

    The present study focused on changes in the annual dynamics of the contents of non-structural saccharides (NSS) of Norway spruce vegetative buds related to their structural development under the effect of acidic pollution during the year 1995. Two types of material were analysed: (1) 4-year-old trees treated for 2 years by simulated acid rain (SAR; pH 2.9 and 3.9), and (2) 40-60-year-old trees growing in natural mountain stands exhibiting different degrees of macroscopic damage. Our study revealed that the dynamics of the NSS content reflected the major morphogenetic and developmental changes occurring during the annual bud developmental cycle. No systematic changes in the annual dynamics of NSS content were observed in buds from both mountain sites, or as a consequence of the SAR. The total sugar content of bud tissues was composed of a combination of five main sugar components: sucrose, glucose, fructose, raffinose family oligosaccharides (RFO; combination of raffinose and stachyose), and a pinitol fraction (PF) probably of cyclitols with pinitol as a main member. The dynamics of individual sugar components also reflected possible carbohydrate mediated bud frost protection. Interesting results were obtained from buds in dormant state. In dormant buds of the SAR experiment the higher value of the ratio PF:RFO of the pinitol fraction and raffinose family oligosaccharides followed the higher dose of SAR treatment. When evaluating the ratio from both types of material we assumed that changes in PF:RFO ratio corresponded to early stages of damage or acute metabolic reaction. Thus, we suggest the ratio PF:RFO as a possible non-specific metabolic marker of early bud stress reaction which is, among other stress factors, sensitive to increasing load of acidic pollutants.

  18. Cell structural changes in the mesophyll of Norway spruce needles by elevated ozone and elevated temperature in open-field exposure during cold acclimation.

    PubMed

    Kivimäenpää, Minna; Riikonen, Johanna; Sutinen, Sirkka; Holopainen, Toini

    2014-04-01

    The effects of elevated ozone (1.4× ambient) and temperature (ambient +1.3 °C) alone and in combination were studied on the needle cell structure of soil-grown Norway spruce seedlings in the late growing season and winter. Temperature treatment continued over winter and lengthened the snow-free period. Elevated temperature caused microscopic changes related to photosynthesis (decreased chloroplast size and increased number), carbon storage (reduced starch and increased cytoplasmic lipids) and defence (decreased mitochondrial size and proportion per cytoplasm, increased peroxisomes and plastoglobuli, altered appearance of tannins). The results suggest increased oxidative stress by elevated temperature and altered allocation of limited carbon reserve to defence. The number of peroxisomes and plastoglobuli remained high in the outer cells of needles of ozone-exposed seedlings but decreased in the inner cells. This may indicate defence allocation to cells close to the stomata and surface, which are experiencing more oxidative stress. Ozone reduced winter hardiness based on seasonal changes in chloroplast shape and location in the cells. The effects of ozone became evident at the end of the growing season, indicating the effect of cumulative ozone dose or that the seedlings were vulnerable to ozone at the later phases of winter hardening. Elevated temperature increased cellular damage in early winter and visible damage in spring, and the damage was enhanced by ozone. In conclusion, the study suggests that modest air temperature elevation increases stress at the cell structural level in spruce seedlings and is enhanced by low ozone elevation. Future climatic conditions where snow cover is formed later or is lacking but temperatures are low can increase the risk of severe seedling damage, and current and future predicted ozone concentrations increase this risk.

  19. Cell structural changes in the mesophyll of Norway spruce needles by elevated ozone and elevated temperature in open-field exposure during cold acclimation.

    PubMed

    Kivimäenpää, Minna; Riikonen, Johanna; Sutinen, Sirkka; Holopainen, Toini

    2014-04-01

    The effects of elevated ozone (1.4× ambient) and temperature (ambient +1.3 °C) alone and in combination were studied on the needle cell structure of soil-grown Norway spruce seedlings in the late growing season and winter. Temperature treatment continued over winter and lengthened the snow-free period. Elevated temperature caused microscopic changes related to photosynthesis (decreased chloroplast size and increased number), carbon storage (reduced starch and increased cytoplasmic lipids) and defence (decreased mitochondrial size and proportion per cytoplasm, increased peroxisomes and plastoglobuli, altered appearance of tannins). The results suggest increased oxidative stress by elevated temperature and altered allocation of limited carbon reserve to defence. The number of peroxisomes and plastoglobuli remained high in the outer cells of needles of ozone-exposed seedlings but decreased in the inner cells. This may indicate defence allocation to cells close to the stomata and surface, which are experiencing more oxidative stress. Ozone reduced winter hardiness based on seasonal changes in chloroplast shape and location in the cells. The effects of ozone became evident at the end of the growing season, indicating the effect of cumulative ozone dose or that the seedlings were vulnerable to ozone at the later phases of winter hardening. Elevated temperature increased cellular damage in early winter and visible damage in spring, and the damage was enhanced by ozone. In conclusion, the study suggests that modest air temperature elevation increases stress at the cell structural level in spruce seedlings and is enhanced by low ozone elevation. Future climatic conditions where snow cover is formed later or is lacking but temperatures are low can increase the risk of severe seedling damage, and current and future predicted ozone concentrations increase this risk. PMID:24718738

  20. Structural interpretation of the Eastern Sulaiman Foldbelt and Foredeep, Pakistan

    NASA Astrophysics Data System (ADS)

    Humayon, Mansoor; Lillie, Robert J.; Lawrence, Robert D.

    1991-04-01

    The Sulaiman foldbelt is an active tectonic feature on the northwestern margin of the Indian plate. Seismic reflection data have been combined with surface geological mapping and drill hole data to interpret the structural style and tectonic shortening of the eastern Sulaiman foldbelt and its adjacent foredeep. The data show that the basement is more than 8 km deep near the deformation front and that it deepens towards the west. The foredeep adjacent to the eastern Sulaiman foldbelt is a broad synclinal structure with a steep western limb; the more gentle eastern limb has monoclinal dips from near zero to 2.5°. Several salt-cored anticlines are observed on the eastern part of the monocline, but the salt structures are lacking in the western foredeep and in the frontal part of the Sulaiman foldbelt. The basal decollement under the Sulaiman lobe lies either in ductile Eocambrian salt or within a deep zone of other material that undergoes ductile behavior. Near the deformation front, ramps from the basal detachment become flats in Lower Triassic and Lower Cretaceous shales. A steep and highly elevated zone immediately to the west of the frontal folds is interpreted to be underlain by a passive-roof duplex. The culmination wall of this oblique duplex is separated from the overlying roof sequence by a passive-roof thrust in Lower Cretaceous shales. The roof-thrust has a backthrust sense of movement, relative to the forelandward propagating duplex, and may extend obliquely 100 km into the interior of the foldbelt. Fault-propagation and fault-bend folds are present in the frontal portion of the eastern Sulaiman foldbelt. Palinspastic restoration of a balanced cross section shows 108 km of shortening in the eastern Sulaiman foldbelt, east of the Kingri strike-slip fault. The presence of more than 11 km of pre-Neogene strata on top of basement suggests that the Mesozoic rifted margin of the Indian subcontinent may lie beneath the Sulaiman foldbelt. This interpretation is

  1. Structure, production and resource use in some old-growth spruce/fir forests in the front range of the Rocky Mountains, USA

    USGS Publications Warehouse

    Binkley, D.; Olsson, U.; Rochelle, R.; Stohlgren, T.; Nikolov, N.

    2003-01-01

    Old-growth forests of Engelmann spruce (Picea engelmannii Parry ex. Engelm.) and subalpine fir (Abies lasiocarpa (Hook.) Nutt.) dominate much of the landscape of the Rocky Mountains. We characterized the structure, biomass and production of 18 old-growth (200-450-year-old) spruce/fir forests in Rocky Mountain National Park, Colorado, as well as the stand-level supply and use of light and nitrogen. Stands were chosen to span a broad range of elevation, aspect, and topography. Aboveground tree biomass in these old-growth forests averaged 253 Mg/ha (range 130-488 Mg/ha), with aboveground net primary production of 3700 kg ha-1 yr-1 (range from 2700 to 5200 kg ha-1 yr-1). Within stands, trees >35 cm in diameter accounted for 70% of aboveground biomass, but trees <35 cm contributed 70% of the production of woody biomass. Differences in slope and aspect among sites resulted in a range of incoming light from 58 to 74 TJ ha-1 yr-1, and tree canopies intercepted an average of 71% of incoming light (range 50-90%). Aboveground net primary production (ANPP) of trees did not relate to the supply of light or N, but ANPP correlated strongly with the amount of light and N used (r2 = 0.45-0.54, P < 0.01). Uptake of 1 kg of N was associated with about 260 kg of ANPP, and one TJ of intercepted shortwave radiation produced about 78 kg of ANPP. Across these old-growth stands, stands with greater biomass showed higher rates of both ANPP and resource use; variation in aboveground biomass was associated with 24% of the variation in N use (P = 0.04), 44% of the light use (P = 0.003), and 45% of the ANPP (P = 0.002). ?? 2002 Elsevier Science B.V. All rights reserved.

  2. Upper Mantle Anisotropy Structures Beneath Eastern Tibet and Northeast Asia

    NASA Astrophysics Data System (ADS)

    Wang, Y.; Wen, L.

    2015-12-01

    Tibetan plateau and eastern Asia subduction zone are tectonic active regions. Understanding the evolution and dynamics process of the two regions is important for us to understand mantle dynamics. A lot of work has been done on the velocity structures beneath the two regions, and several tectonic models are proposed to explain their dynamic process. But due to the absence of the detailed upper mantle anisotropy structures, those models are still under debate. Fine upper mantle velocity and anisotropy structures can help us understand the dynamic process of the two regions. Waveform modeling of upper mantle triplication phases can provide a good vertical resolution of upper mantle velocity structures, but present methods for calculating synthetic seismograms cannot process anisotropic media. We develop a method based on the generalized reflection and transmission method (GRTM) to calculate synthetic seismograms for wave propagating in stratified VTI media, so we can waveform model upper mantle triplications propagating in anisotropic media. In this study, we waveform model the tangential and radial seismic triplication data recorded in Chinese digital seismic stations at a epicentral distance of 10-30 degree for one event occurring in middle Tibet and one event occurring in Japan, to constrain fine upper mantle velocity and anisotropy structures beneath eastern Tibet and northeast Asia. The result shows that beneath eastern Tibet, horizontal S wave velocity is larger than vertical S wave velocity in the upper mantle; beneath northeast Asia, horizontal S wave velocity is larger than vertical S wave velocity above the depth of 190 km, and is smaller than vertical S wave velocity below the depth of 190 km. We also build a mineral physics modeling method, which can calculate upper mantle anisotropy structures based on mantle temperatures, compositions and directions of mantle flow, and use this method to explore compositional and dynamic models that would explain the

  3. Investigation of upper crustal structure beneath eastern Java

    NASA Astrophysics Data System (ADS)

    Martha, Agustya Adi; Widiyantoro, Sri; Cummnins, Phil; Saygin, Erdinc; Masturyono

    2016-05-01

    The complexity of geology structure in eastern Java causes this region has many potential resources as much as the disasters. Therefore, the East Java province represents an interesting area to be explored, especially regarding its upper crustal structure. To investigate this structure, we employ the Ambient Noise Tomography (ANT) method. We have used seismic waveform data from 25 Meteorological, Climatological and Geophysical Agency (BMKG) stationary seismographic stations and 26 portable seismographs installed for 2 to 8 weeks. Inter-station cross-correlation produces more than 800 Rayleigh wave components, which depict the structure beneath eastern Java. Based on the checkerboard resolution test, we found that the optimal grid size is 0.25ox0.25o. Our inversion results for the periods of 1 to 10 s indicate a good agreement with geological and Bouguer anomaly maps. Rembang high depression, most of the southern mountains zone, the northern part of Rembang zone and the central part of the Madura Island, the area of high gravity anomaly and areas dominated with igneous rocks are associated with high velocity zones. On the other hand, Kendeng zone and most of the basin in the Rembang zone are associated with low velocity zones.

  4. Composition of the Spruce Budworm (Choristoneura fumiferana) Midgut Microbiota as Affected by Rearing Conditions.

    PubMed

    Landry, Mathieu; Comeau, André M; Derome, Nicolas; Cusson, Michel; Levesque, Roger C

    2015-01-01

    The eastern spruce budworm (Choristoneura fumiferana) is one of the most destructive forest insect pests in Canada. Little is known about its intestinal microbiota, which could play a role in digestion, immune protection, communication and/or development. The present study was designed to provide a first characterization of the effects of rearing conditions on the taxonomic diversity and structure of the C. fumiferana midgut microbiota, using a culture-independent approach. Three diets and insect sources were examined: larvae from a laboratory colony reared on a synthetic diet and field-collected larvae reared on balsam fir or black spruce foliage. Bacterial DNA from the larval midguts was extracted to amplify and sequence the V6-V8 region of the 16S rRNA gene, using the Roche 454 GS-FLX technology. Our results showed a dominance of Proteobacteria, mainly Pseudomonas spp., in the spruce budworm midgut, irrespective of treatment group. Taxonomic diversity of the midgut microbiota was greater for larvae reared on synthetic diet than for those collected and reared on host plants, a difference that is likely accounted for by several factors. A greater proportion of bacteria from the phylum Bacteroidetes in insects fed artificial diet constituted the main difference between this group and those reared on foliage; within the phylum Proteobacteria, the presence of the genus Bradyrhizobium was also unique to insects reared on artificial diet. Strikingly, a Bray-Curtis analysis showed important differences in microbial diversity among the treatment groups, pointing to the importance of diet and environment in defining the spruce budworm midgut microbiota. PMID:26636571

  5. Composition of the Spruce Budworm (Choristoneura fumiferana) Midgut Microbiota as Affected by Rearing Conditions

    PubMed Central

    Landry, Mathieu; Comeau, André M.; Derome, Nicolas; Cusson, Michel; Levesque, Roger C.

    2015-01-01

    The eastern spruce budworm (Choristoneura fumiferana) is one of the most destructive forest insect pests in Canada. Little is known about its intestinal microbiota, which could play a role in digestion, immune protection, communication and/or development. The present study was designed to provide a first characterization of the effects of rearing conditions on the taxonomic diversity and structure of the C. fumiferana midgut microbiota, using a culture-independent approach. Three diets and insect sources were examined: larvae from a laboratory colony reared on a synthetic diet and field-collected larvae reared on balsam fir or black spruce foliage. Bacterial DNA from the larval midguts was extracted to amplify and sequence the V6-V8 region of the 16S rRNA gene, using the Roche 454 GS-FLX technology. Our results showed a dominance of Proteobacteria, mainly Pseudomonas spp., in the spruce budworm midgut, irrespective of treatment group. Taxonomic diversity of the midgut microbiota was greater for larvae reared on synthetic diet than for those collected and reared on host plants, a difference that is likely accounted for by several factors. A greater proportion of bacteria from the phylum Bacteroidetes in insects fed artificial diet constituted the main difference between this group and those reared on foliage; within the phylum Proteobacteria, the presence of the genus Bradyrhizobium was also unique to insects reared on artificial diet. Strikingly, a Bray-Curtis analysis showed important differences in microbial diversity among the treatment groups, pointing to the importance of diet and environment in defining the spruce budworm midgut microbiota. PMID:26636571

  6. Crustal Structure in Central-Eastern Greenland: TOPOGREENLAND Refraction Profile

    NASA Astrophysics Data System (ADS)

    Shulgin, A. A.; Thybo, H.

    2013-05-01

    We present the seismic structure in the interior of Greenland based on the first measurements by the seismic refraction/wide angle reflection method. Previous seismic surveys have only been carried out offshore and near the coast of Greenland, where the crustal structure is affected by oceanic break-up and may not be representative of the interior of the island. Acquisition of geophysical data onshore Greenland is logistically complicated by the presence of an up to 3.4 km thick ice sheet, permanently covering most of the land mass. The seismic data was acquired by a team of six people during a two-month long experiment in summer of 2011 on the ice cap in the interior of central-eastern Greenland. The EW-trending profile extends 310 km inland from the approximate edge of the stable ice cap near Scoresby Sund across the centre of the ice cap. The planned extension of the profile by use of OBSs and air gun shooting in Scoresbysund Fjord to the east coast of Greenland was unfortunately cancelled, because navigation was prevented by ice drift. 350 Reftek Texan receivers recorded high-quality seismic data from 8 equidistant shots along the profile. Explosive charge sizes were 1 ton at the ends and ca. 500 kg along the profile, loaded with about 100 kg at 35-85 m depth in individual boreholes. Two-dimensional velocity model based on forward ray tracing and tomography modelling shows a decrease of crustal thickness from 47 km below the centre of Greenland in the western part to 40 km in the eastern part of the profile. Earlier studies show that crustal thickness further decreases eastward to ca. 30 km below the fjord system, but details of the changes are unknown. Relatively high lower crustal velocities (Vp 6.8 - 7.3) in the western part of the TopoGreenland profile may indicate past collision tectonics or may be related or to the passage of the Iceland mantle plume. The origin of the pronounced circum-Atlantic mountain ranges in Norway and eastern Greenland, which have

  7. Profiling functions of ectomycorrhizal diversity and root structuring in seedlings of Norway spruce (Picea abies) with fast- and slow-growing phenotypes.

    PubMed

    Velmala, Sannakajsa M; Rajala, Tiina; Heinonsalo, Jussi; Taylor, Andy F S; Pennanen, Taina

    2014-01-01

    We studied the role of taxonomical and functional ectomycorrhizal (ECM) fungal diversity in root formation and nutrient uptake by Norway spruce (Picea abies) seedlings with fast- and slow-growing phenotypes. Seedlings were grown with an increasing ECM fungal diversity gradient from one to four species and sampled before aboveground growth differences between the two phenotypes were apparent. ECM fungal colonization patterns were determined and functional diversity was assayed via measurements of potential enzyme activities of eight exoenzymes probably involved in nutrient mobilization. Phenotypes did not vary in their receptiveness to different ECM fungal species. However, seedlings of slow-growing phenotypes had higher fine-root density and thus more condensed root systems than fast-growing seedlings, but the potential enzyme activities of ectomycorrhizas did not differ qualitatively or quantitatively. ECM species richness increased host nutrient acquisition potential by diversifying the exoenzyme palette. Needle nitrogen content correlated positively with high chitinase activity of ectomycorrhizas. Rather than fast- and slow-growing phenotypes exhibiting differing receptiveness to ECM fungi, our results suggest that distinctions in fine-root structuring and in the belowground growth strategy already apparent at early stages of seedling development may explain later growth differences between fast- and slow-growing families.

  8. SPRUCE experiment data infrastructure

    NASA Astrophysics Data System (ADS)

    Krassovski, M.; Hanson, P. J.; Boden, T.; Riggs, J.; Nettles, W. R.; Hook, L. A.

    2013-12-01

    The Carbon Dioxide Information Analysis Center (CDIAC) at Oak Ridge National Laboratory (ORNL), USA has provided scientific data management support for the US Department of Energy and international climate change science since 1982. Among the many data activities CDIAC performs are design and implementation of the data systems. One current example is the data system and network for SPRUCE experiment. The SPRUCE experiment (http://mnspruce.ornl.gov) is the primary component of the Terrestrial Ecosystem Science Scientific Focus Area of ORNL's Climate Change Program, focused on terrestrial ecosystems and the mechanisms that underlie their responses to climatic change. The experimental work is to be conducted in a bog forest in northern Minnesota, 40 km north of Grand Rapids, in the USDA Forest Service Marcell Experimental Forest (MEF). The site is located at the southern margin of the boreal peatland forest. Experimental work in the 8.1-ha S1 bog will be a climate change manipulation focusing on the combined responses to multiple levels of warming at ambient or elevated CO2 (eCO2) levels. The experiment provides a platform for testing mechanisms controlling the vulnerability of organisms, biogeochemical processes and ecosystems to climatic change (e.g., thresholds for organism decline or mortality, limitations to regeneration, biogeochemical limitations to productivity, the cycling and release of CO2 and CH4 to the atmosphere). The manipulation will evaluate the response of the existing biological communities to a range of warming levels from ambient to +9°C, provided via large, modified open-top chambers. The ambient and +9°C warming treatments will also be conducted at eCO2 (in the range of 800 to 900 ppm). Both direct and indirect effects of these experimental perturbations will be analyzed to develop and refine models needed for full Earth system analyses. SPRUCE provides wide range continuous and discrete measurements. To successfully manage SPRUCE data flow

  9. Genetic Structure of Daphnia galeata Populations in Eastern China

    PubMed Central

    Wolinska, Justyna; Ma, Xiaolin; Yang, Zhong; Hu, Wei; Yin, Mingbo

    2015-01-01

    This study presents the first examination of the genetic structure of Daphnia longispina complex populations in Eastern China. Only one species, D. galeata, was present across the eight investigated lakes; as identified by taxon assignment using allelic variation at 15 microsatellite loci. Three genetically differentiated D. galeata subgroups emerged independent of the type of statistical analysis applied. Thus, Bayesian clustering, discriminant analysis based on results from factorial correspondence analysis, and UPGMA clustering consistently showed that populations from two neighbouring lakes were genetically separated from a mixture of genotypes found in other lakes, which formed another two subgroups. Clonal diversity was high in all D. galeata populations, and most samples showed no deviation from Hardy-Weinberg equilibrium, indicating that clonal selection had little effect on the genetic diversity. Overall, populations did not cluster by geographical origin. Further studies will show if the observed pattern can be explained by natural colonization processes or by recent anthropogenic impact on predominantly artificial lakes. PMID:25768727

  10. Structural evolution of the Llanos foothills, Eastern Cordillera, Colombia

    NASA Astrophysics Data System (ADS)

    Martinez, Jaime A.

    2006-09-01

    The Llanos foothills are located in the frontal thrust zone of the Eastern Cordillera in Colombia in a complex environment that BP has been exploring actively since 1988. This exploration has resulted in the discovery of several fields with a variety of hydrocarbon fluids (gas condensate and volatile oil) in very tight quartz-arenites. The structural style and complexity of this fold-and-thrust belt changes along the trend from single frontal structures to an imbricate of up to five thrust sheets in a triangle zone. In highly complex environments, the seismic image quality is poor, and interpretation becomes very challenging. The structural models of the area have evolved as more data have been acquired. The initial structural model required inversion of the basin at the end of the Andean orogeny. The structural style changed to an in-sequence imbricate thrust stack with very long, trailing back limbs that return to regional elevation and finalize in a tighter structures with short back limbs. The concept of early deformation and multiple phases has been introduced. Three main phases have been distinguished: (1) an early event during the deposition of the Lower Carbonera (39-29 Ma), with incipient structures formed to create syntectonic deposition; (2) a phase of steady subsidence that increased notably at the end of the period (29-7 Ma); and (3) the latest phase (7-0 Ma), when most deformation and uplifting occurred. The migration of hydrocarbons happened simultaneously with the deformation, and its final distribution, amount, and variation in composition is related to the structural evolution of the area.

  11. Climate-induced mortality of spruce stands in Belarus

    NASA Astrophysics Data System (ADS)

    Kharuk, Viacheslav I.; Im, Sergei T.; Dvinskaya, Maria L.; Golukov, Alexei S.; Ranson, Kenneth J.

    2015-12-01

    The aim of this work is an analysis of the causes of spruce (Picea abies L.) decline and mortality in Belarus. The analysis was based on forest inventory and Landsat satellite (land cover classification, climate variables (air temperature, precipitation, evaporation, vapor pressure deficit, SPEI drought index)), and GRACE-derived soil moisture estimation (equivalent of water thickness anomalies, EWTA). We found a difference in spatial patterns between dead stands and all stands (i.e., before mortality). Dead stands were located preferentially on relief features with higher water stress risk (i.e., higher elevations, steeper slopes, south and southwestern exposure). Spruce mortality followed a series of repeated droughts between 1990 and 2010. Mortality was negatively correlated with air humidity (r = -0.52), and precipitation (r = -0.57), and positively correlated with the prior year vapor pressure deficit (r = 0.47), and drought increase (r = 0.57). Mortality increased with the increase in occurrence of spring frosts (r = 0.5), and decreased with an increase in winter cloud cover (r = -0.37). Spruce mortality was negatively correlated with snow water accumulation (r = -0.81) and previous year anomalies in water soil content (r = -0.8). Weakened by water stress, spruce stands were attacked by pests and phytopathogens. Overall, spruce mortality in Belarussian forests was caused by drought episodes and drought increase in synergy with pest and phytopathogen attacks. Vast Picea abies mortality in Belarus and adjacent areas of Russia and Eastern Europe is a result of low adaptation of that species to increased drought. This indicates the necessity of spruce replacement by drought-tolerant indigenous (e.g., Pinus sylvestris, Querqus robur) or introduced (e.g., Larix sp. or Pseudotsuga menzieslii) species to obtain sustainable forest growth management.

  12. Structural Model of the Tucupita Field, Eastern Venezuela Basin

    NASA Astrophysics Data System (ADS)

    Arteaga, L. A.

    2013-05-01

    The Tucupita Field has an area of 73,51 Km2, is located between the states of Monagas and Delta Amacuro, geologically is located at the greater Temblador area in the Eastern Venezuela Basin, where the Oficina Formation's sands represent the main hydrocarbons reservoirs. From the results of the seismic reprocessing realized by Fusion Petroleum Technologies, Inc., the structural model of this field was done as initial step to the geocellular model of the Oficina-40 Reservoir, which was defined as a Faulted Relay Ramp, where the normal faults are dominant with NE-SW orientation Introduction The Tucupita Field is a mature oilfield at the greater Temblador area, however most of the wells were completed in the upper sands, therefore the main study is focused in the geological characterization of the Oficina-40 Reservoir's lower sands, starting by the structural model Previous Studies 1. Proyecto Tucupita 3D The seismic data of the Tucupita Field were adquired in 1996 by Western Atlas of Venezuela for the Benton Vinccler Company. The UTM coordinates of the wells used in this project, have been taken to make this geological model 2. Soporte Geofísico Integrado The seismic project Tucupita was processed by Fusion Petroleum Technologies Inc., in Houston and consisted of reprocessing and pre-stack migration in time (PSTM) and pre-stack migration in depth (PSDM), this data belong to the Petrodelta Company Based on the regional stratigraphy, were validated the "picks" to make the structural sections to support research with hard data. After, it proceeded to interpret the structural style of the field from the seismic amplitude cube. Then, it was done the faults modelling and the stratigraphic horizons to carry out the geocellular model Three structural sections were realized, which was interpreted like a faulted monocline, whose peak is located southward, where justly the wells are located. The contact oil-water was interpreted to -5648'. Echelon faults were interpreted in a

  13. Genetic diversity and population structure of rice landraces from Eastern and North Eastern States of India

    PubMed Central

    2013-01-01

    Background Adaptations to different habitats across the globe and consequent genetic variation within rice have resulted in more than 120,000 diverse accessions including landraces, which are vital genetic resources for agronomic and quality traits. In India the rice landraces of the states West Bengal, Assam, Mizoram, Manipur and Nagaland are worthy candidates for genetic assessment. Keeping the above in view, the present study was conducted with the aim to (i) calculate the genetic distances among the accessions of 83 landraces collected from these states along with 8 check accessions (total 91 accessions) using 23 previously mapped SSR markers and (ii) examine the population structure among the accessions using model-based clustering approach. Results Among the 91 accessions, 182 alleles were identified which included 51 rare and 27 null alleles. The average PIC value was 0.7467/marker. The non-aromatic landraces from West Bengal was most diverse with 154 alleles and an average PIC value of 0.8005/marker, followed by the aromatic landraces from West Bengal with 118 alleles and an average PIC value of 0.6524/marker, while the landraces from North East ranked third with 113 alleles and an average PIC value of 0.5745/marker. In the dendrogram distinct clusters consisting of predominantly aromatic landraces and predominantly North East Indian landraces were observed. The non-aromatic landraces from West Bengal were interspersed within these two clusters. The accessions were moderately structured, showing four sub-populations (A-D) with an Fst value of 0.398, 0.364, 0.206 and 0.281, respectively. The assigned clustering of accessions was well in agreement in both distance-based and model-based approaches. Conclusions Each of the accessions could be identified unequivocally by the SSR profiles. Genetically the non aromatic landraces from West Bengal were most diverse followed by the aromatic landraces from the same state. The North Eastern accessions ranked third

  14. Densities of breeding birds and changes in vegetation in an alaskan boreal forest following a massive disturbance by spruce beetles

    USGS Publications Warehouse

    Matsuoka, S.M.; Handel, C.M.; Ruthrauff, D.R.

    2001-01-01

    We examined bird and plant communities among forest stands with different levels of spruce mortality following a large outbreak of spruce beetles (Dendroctonus rufipennis (Kirby)) in the Copper River Basin, Alaska. Spruce beetles avoided stands with black spruce (Picea mariana) and selectively killed larger diameter white spruce (Picea glauca), thereby altering forest structure and increasing the dominance of black spruce in the region. Alders (Alnus sp.) and crowberry (Empetrum nigrum) were more abundant in areas with heavy spruce mortality, possibly a response to the death of overstory spruce. Grasses and herbaceous plants did not proliferate as has been recorded following outbreaks in more coastal Alaskan forests. Two species closely tied to coniferous habitats, the tree-nesting Ruby-crowned Kinglet (Regulus calendula) and the red squirrel (Tamiasciurus hudsonicus), a major nest predator, were less abundant in forest stands with high spruce mortality than in low-mortality stands. Understory-nesting birds as a group were more abundant in forest stands with high levels of spruce mortality, although the response of individual bird species to tree mortality was variable. Birds breeding in stands with high spruce mortality likely benefited reproductively from lower squirrel densities and a greater abundance of shrubs to conceal nests from predators.

  15. 1. 20472009 SPRUCE ST. RUNS FROM LEFT TO RIGHT. SOUTH ...

    Library of Congress Historic Buildings Survey, Historic Engineering Record, Historic Landscapes Survey

    1. 2047-2009 SPRUCE ST. RUNS FROM LEFT TO RIGHT. SOUTH (FRONT) FACADES. VIEW LOOKING NORTHEAST - Spruce Street Area Study, 2009-2045 Spruce Street (Houses), Spruce Street, north side, Philadelphia, Philadelphia County, PA

  16. The 3D geological model of the Eastern Romania tectonics and structure

    NASA Astrophysics Data System (ADS)

    Necula, Nicusor; Sorin Baciu, Dorin; Niculita, Mihai; Dumitriu, Tony-Cristian

    2016-04-01

    3D geologic modelling is a modern tool which allow the conceptualization of geologic relations in an interactive environment, strengthening the ability to understand and present tectonic and structural geologic models. We integrated the data available in the literature (wells, maps, cross-sections) for the geological structure of the Eastern Romania, comprising the Eastern Carpathians Orogen and its foreland. The subducting East European plate generated the Eastern Carphatians thrusts. Under the Eastern Carpathians, beside East European plate, the Tornquist-Teysseire zone is caught. East European Craton (Proterozoic), Scythian Platform (Paleozoic), North Dobrogean Orogen (Paleozoic) and Moesian Platform (Paleozoic), all neighbor Tornquist-Teysseire zone (Paleozoic), playing the role of foreland for the Eastern Carpathian Orogen. The Eastern Carphatians Orogen has two flysch belts, the Inner Carpathian called Dacides formed in Cretacic deformations and the Outer Carpathian called Moldavides and formed in Late Badenian to Sarmatian deformations. The modelling was performed in Midland Valley's Move software. The boundaries of all the structural units presented above were modelled, together with the faults which are represented on the various osurces used. The created 3D geological model is seen as a tool to better understand and represent the tectonic and structural model of the Eastern ROmania and will also allow a better quantification of the relations between geology and landforms in Eastern Romania.

  17. Pulpwood, pesticides, and people. Controlling spruce budworm in northeastern North America

    NASA Astrophysics Data System (ADS)

    Irland, Lloyd C.

    1980-09-01

    The eastern spruce budworm is a major forest pest over the continental range of the spruce-fir forest ecosystem and its southern ecotonal fringes in Canada and the northeastern United States. The current budworm outbreak illustrates the difficulty of arriving at economically sound and publicly acceptable forest pest control policies. Policies ranging from no use of chemical control to annual widespread crop protection have been adopted. There is no single all-around “best” policy for spruce budworm control. Chemical spray programs have demonstrably slowed the normal progress of mortality due to budworm, but have not eradicated the pest. Where industry remains heavily dependent on a fully utilized spruce-fir forest, no easy, low-cost solutions to the budworm problem exist. Reliance on spraying will have to be reduced and plans made to utilize higher levels of tree mortality and to manage the forest for lower future vulnerability.

  18. Fire severity unaffected by spruce beetle outbreak in spruce-fir forests in southwestern Colorado.

    PubMed

    Andrus, Robert A; Veblen, Thomas T; Harvey, Brian J; Hart, Sarah J

    2016-04-01

    Recent large and severe outbreaks of native bark beetles have raised concern among the general public and land managers about potential for amplified fire activity in western North America. To date, the majority of studies examining bark beetle outbreaks and subsequent fire severity in the U.S. Rocky Mountains have focused on outbreaks of mountain pine beetle (MPB; Dendroctonus ponderosae) in lodgepole pine (Pinus contorta) forests, but few studies, particularly field studies, have addressed the effects of the severity of spruce beetle (Dendroctonus rufipennis Kirby) infestation on subsequent fire severity in subalpine Engelmann spruce (Picea engelmannii) and subalpine fir (Abies lasiocarpa) forests. In Colorado, the annual area infested by spruce beetle outbreaks is rapidly rising, while MPB outbreaks are subsiding; therefore understanding this relationship is of growing importance. We collected extensive field data in subalpine forests in the eastern San Juan Mountains, southwestern Colorado, USA, to investigate whether a gray-stage (< 5 yr from outbreak to time of fire) spruce beetle infestation affected fire severity. Contrary to the expectation that bark beetle infestation alters subsequent fire severity, correlation and multivariate generalized linear regression analysis revealed no influence of pre-fire spruce beetle severity on nearly all field or remotely sensed measurements of fire severity. Findings were consistent across moderate and extreme burning conditions. In comparison to severity of the pre-fire beetle outbreak, we found that topography, pre-outbreak basal area, and weather conditions exerted a stronger effect on fire severity. Our finding that beetle infestation did not alter fire severity is consistent with previous retrospective studies examining fire activity following other bark beetle outbreaks and reiterates the overriding influence of climate that creates conditions conducive to large, high-severity fires in the subalpine zone of Colorado

  19. Fire severity unaffected by spruce beetle outbreak in spruce-fir forests in southwestern Colorado.

    PubMed

    Andrus, Robert A; Veblen, Thomas T; Harvey, Brian J; Hart, Sarah J

    2016-04-01

    Recent large and severe outbreaks of native bark beetles have raised concern among the general public and land managers about potential for amplified fire activity in western North America. To date, the majority of studies examining bark beetle outbreaks and subsequent fire severity in the U.S. Rocky Mountains have focused on outbreaks of mountain pine beetle (MPB; Dendroctonus ponderosae) in lodgepole pine (Pinus contorta) forests, but few studies, particularly field studies, have addressed the effects of the severity of spruce beetle (Dendroctonus rufipennis Kirby) infestation on subsequent fire severity in subalpine Engelmann spruce (Picea engelmannii) and subalpine fir (Abies lasiocarpa) forests. In Colorado, the annual area infested by spruce beetle outbreaks is rapidly rising, while MPB outbreaks are subsiding; therefore understanding this relationship is of growing importance. We collected extensive field data in subalpine forests in the eastern San Juan Mountains, southwestern Colorado, USA, to investigate whether a gray-stage (< 5 yr from outbreak to time of fire) spruce beetle infestation affected fire severity. Contrary to the expectation that bark beetle infestation alters subsequent fire severity, correlation and multivariate generalized linear regression analysis revealed no influence of pre-fire spruce beetle severity on nearly all field or remotely sensed measurements of fire severity. Findings were consistent across moderate and extreme burning conditions. In comparison to severity of the pre-fire beetle outbreak, we found that topography, pre-outbreak basal area, and weather conditions exerted a stronger effect on fire severity. Our finding that beetle infestation did not alter fire severity is consistent with previous retrospective studies examining fire activity following other bark beetle outbreaks and reiterates the overriding influence of climate that creates conditions conducive to large, high-severity fires in the subalpine zone of Colorado

  20. Afghanistan structure, greater India concept and Eastern Tethys evolution

    NASA Astrophysics Data System (ADS)

    Boulin, Jean

    1981-02-01

    The large and complex area between Central Asia and the Arabia—India block includes, from north to south, the Tien Shan and Hindu Kush—Kun Lun Hercynian belts, the Turkman—Farah Rud and Tanghla Shan Neokimmerian belt, the Zagros—Makran—Baluch—Himalayan Alpine belt and relics of corresponding oceanic spaces. The Angarian or Gondwanian affinities of the different blocks separated by these successive peri-Indian belts are discussed. An interpretation is proposed for the paleogeographic and structural evolution of this area, from Early Paleozoic to Present. This interpretation is based on geologic and paleomagnetic data and on global tectonic considerations. The Hindu Kush—Kun Lun pre-Hercynian ocean probably separated Laurasia from Gondwanaland. It was an Early Paleozoic Tethys (Tethys 1). The Tien Shan ocean may have been a marginal basin for the Hindu Kush—Kun Lun ocean. The Turkman-Farah Rud—Tanghla Shan pre-Neokimmerian ocean resulted from the break-up of the Gondwanaland margin. Its opening seems to have been related to the pre-Hercynian ocean closure resulting in the formation of a new Tethys (Tethys 2). Its subsequent closure may also have been related to the opening of the Zagros—Baluch—Indus—Tsang Po pre-Alpine ocean (Tethys 3). According to this interpretation, Eurasia appears to have increased from Early Paleozoic to Cenozoic by successive accretion of cratonic blocks which came from the south through the Tethys. This evolution can be compared to the similar Mediterranean evolution. This interpretation is in agreement with the Greater India concept. However, the northern border of Greater India differs according to the stage of the Gondwanaland evolution which is considered. Actually, Greatest India extended north to the Caledonian belt and consequently has recorded the whole history of the Eastern Tethys.

  1. Structure and tectonic setting of the eastern Tehachapi Range, California

    SciTech Connect

    Wood, D.J.; Saleeby, J.B.; Silver, L.T. . Division of Geological and Planetary Sciences)

    1993-04-01

    In the easternmost Tehachapi mountains a moderately SE dipping fault zone with an irregular trace juxtaposes high grade ductilely deformed footwall gneisses against a cataclastically deformed granitic hanging wall complex from Tehachapi Valley to the Garlock FZ. The fault zone exhibits evidence of both ductile and brittle deformation. Moderately SE plunging stretching lineations and sense of shear indicators in the footwall suggest normal displacement and the sole of the hanging wall consists of a chloritic breccia. The footwall consists of garnet-amphibolite grade orthogneiss, paragneiss, and marble, locally protomylonitic, folded into presently SW vergent isoclinal folds with a penetrative, NW trending, moderately NE dipping foliation. This gneiss complex hosts a 0.2 to 0.8 km wide, NW tending, shallow NE dipping ductile shear zone, called here the eastern Tehachapi shear zone (ETSZ). The ETSZ appears to post-date much of the isoclinal folding in the gneisses. The ETSZ is inferred to continue north of Tehachapi valley (Ross, 1989) where it steepens and swings to a more northerly trend into the Lake Isabella region where it has been called the proto-Kern Canyon fault zone. A post ETSZ deformation refolded the gneisses into open, upright folds with moderately NE plunging axes. Juxtaposition of the gneisses and the cataclastically deformed granitic hanging wall rocks occurred during and/or after this post ETSZ folding event. In the structurally lowest levels of the gneiss complex exposed along the Garlock fault, quartz diorite orthogneiss becomes increasingly deformed downward with an NE striking, moderately NW dipping mylonitic fabric. The youngest deformation in the areas is a series of WNW trending, regional scale antiforms and synforms which exert a first order control on the topography of the region.

  2. ASSOCIATIONS OF EASTERN REDCEDAR AND COMMUNITY STRUCTURE OF SMALL MAMMALS

    EPA Science Inventory

    Increased abundance of eastern redcedar (Juniperus virginianas), a native but invasive species in the Great Plains, has been associated with several changes in ecosystem function. It can lead to a reduction in herbaceous biomass in the canopy zone, alter species composition, and...

  3. Social Structure and Social Change in Eastern Europe.

    ERIC Educational Resources Information Center

    Fischer, George; Schenkel, Walter

    This specialized bibliography of scholarly writings since 1945 on Eastern Europe covers the countries of Albania, Bulgaria, Czechoslovakia, Greece, Hungary, Poland, Romania, and Yugoslavia. Distinct entries number about 700 and cover works published in English in the United States and Great Britain and also sources in French and German published…

  4. The Crustal Structure and Seismicity of Eastern Venezuela

    NASA Astrophysics Data System (ADS)

    Schmitz, M.; Martins, A.; Sobiesiak, M.; Alvarado, L.; Vasquez, R.

    2001-12-01

    Eastern Venezuela is characterized by a moderate to high seismicity, evidenced recently by the 1997 Cariaco earthquake located on the El Pilar Fault, a right lateral strike slip fault which marks the plate boundary between the Caribbean and South-American plates in this region. Recently, the seismic activity seems to migrate towards the zone of subduction of the Lesser Antilles in the northeast, where a mb 6.0 earthquake occurred in October 2000 at 120 km of depth. Periodical changes in the seismic activity are related to the interaction of the stress fields of the strike-slip and the subduction regimes. The seismic activity decreases rapidly towards to the south with some disperse events on the northern edge of the Guayana Shield, related to the Guri fault system. The crustal models used in the region are derived from the information generated by the national seismological network since 1982 and by microseismicity studies in northeastern Venezuela, coinciding in a crustal thickness of about 35 km in depth. Results of seismic refraction measurements for the region were obtained during field campains in 1998 (ECOGUAY) for the Guayana Shield and the Cariaco sedimentary basin and in 2001 (ECCO) for the Oriental Basin. The total crustal thickness decreases from about 45 km on the northern edge of the Guayana Shield to some 36 km close to El Tigre in the center of the Oriental Basin. The average crustal velocity decreases in the same sense from 6.5 to 5.8 km/s. In the Cariaco sedimentary basin a young sedimentary cover of 1 km thickness with a seismic velocity of 2 km/s was derived. Towards the northern limit of the South-American plate, no deep seismic refraction data are available up to now. The improvement of the crustal models used in that region would constitute a step forward in the analysis of the seismic hazard. Seismic refraction studies funded by CONICIT S1-97002996 and S1-2000000685 projects and PDVSA (additional drilling and blasting), recording equipment

  5. Growth declines in red spruce

    SciTech Connect

    McLaughlin, S.B. ); Adams, H.S. )

    1987-10-01

    In this letter, the authors take issue with Zedaker, Hyink, and Smith who have indicated that observed red spruce growth declines can be expected based on growth trends for even-aged stands of red spruce as documented in Meyer (1929). Recently, an examination was made of stand stocking levels at 750 sites where red spruce were cored and neither the rate of growth decline nor the extent of mortality were found to be related to stand stocking levels or previous disturbance history. The authors conclude that the Meyer data do not represent an appropriate model for stand dynamics of old-growth, high-elevation stands and no not adequately explain the growth declines observed at many of those sites.

  6. Clinal variation at phenology-related genes in spruce: parallel evolution in FTL2 and Gigantea?

    PubMed

    Chen, Jun; Tsuda, Yoshiaki; Stocks, Michael; Källman, Thomas; Xu, Nannan; Kärkkäinen, Katri; Huotari, Tea; Semerikov, Vladimir L; Vendramin, Giovanni G; Lascoux, Martin

    2014-07-01

    Parallel clines in different species, or in different geographical regions of the same species, are an important source of information on the genetic basis of local adaptation. We recently detected latitudinal clines in SNPs frequencies and gene expression of candidate genes for growth cessation in Scandinavian populations of Norway spruce (Picea abies). Here we test whether the same clines are also present in Siberian spruce (P. obovata), a close relative of Norway spruce with a different Quaternary history. We sequenced nine candidate genes and 27 control loci and genotyped 14 SSR loci in six populations of P. obovata located along the Yenisei river from latitude 56°N to latitude 67°N. In contrast to Scandinavian Norway spruce that both departs from the standard neutral model (SNM) and shows a clear population structure, Siberian spruce populations along the Yenisei do not depart from the SNM and are genetically unstructured. Nonetheless, as in Norway spruce, growth cessation is significantly clinal. Polymorphisms in photoperiodic (FTL2) and circadian clock (Gigantea, GI, PRR3) genes also show significant clinal variation and/or evidence of local selection. In GI, one of the variants is the same as in Norway spruce. Finally, a strong cline in gene expression is observed for FTL2, but not for GI. These results, together with recent physiological studies, confirm the key role played by FTL2 and circadian clock genes in the control of growth cessation in spruce species and suggest the presence of parallel adaptation in these two species.

  7. Growth declines in red spruce

    SciTech Connect

    Zedaker, S.M.; Hyink, D.M.; Smith, D.W.

    1987-01-01

    Over the past two decades second-growth red spruce stands in the Northeast have demonstrated declines in radial increment. Some observers are implicating air pollution as a primary cause of the declines, based on recently acquired increment cores from dominant trees. Various forms of air pollution (O/sub 3/, NO/sub x/, SO/sub 2/, and trace metals) are known to reduce growth and development of tree species, but few studies have provided concrete evidence of regional pollution-caused declines in forest ecosystems. Recently published evidence of a synchronous, consistent, and unprecedented regional decline in red spruce should be weighed against the realization that radial increment in red spruce declines naturally as stands age. Separating anthropogenic stress-caused growth patterns from natural stand dynamics requires an in-depth knowledge of forest growth and yield, tree silvics, and forest ecosystem processes. Detailed analyses of growth by stand characteristics - site index, density, elevation, stand history - will be necessary to implicate air pollution as a primary cause of red spruce decline.

  8. Red spruce decline---Winter injury and air pollutants

    SciTech Connect

    Roberts, T.M. )

    1989-10-01

    There has been a widespread decline in growth of red spruce (Picea rubens Sarg.) since 1960 in the eastern United States. There is evidence that this decline is at least partly attributable to age- and density-related growth patterns, particularly at lower elevations. Mortality has been severe at high elevation sites where similar episodes have occasionally occurred in the last 100 years. At these sites, periods of low growth preceding 1960 were related to periods with warm late summers and cold early winters. Since 1960, this relationship no longer holds, although there is an association with unusual deviations from mean temperatures. There are field reports that one of the main causes of reduced growth and mortality is apical dieback induced by severe winter conditions. Preliminary observations suggest that high elevation red spruce may not be sufficiently hardened to tolerate low autumn temperatures. However, appearance of injury in the spring, association of injury with wind exposure and correlation of provenance susceptibility with cuticular transpiration rates, including the importance of desiccation injury. Sensitivity to both types of winter injury may be increased by air pollutants (particularly ozone and less probably, acid mist or excess nitrogen deposition). Nutrient deficiency (particularly magnesium and to a lesser extent potassium) may also increase cold sensitivity. The nature and extent of these interactions are being actively researched for red spruce. 48 refs.

  9. Effects of artificial defoliation of pines on the structure and physiology of the soil fungal community of a mixed pine-spruce forest

    NASA Technical Reports Server (NTRS)

    Cullings, Ken; Raleigh, Christopher; New, Michael H.; Henson, Joan

    2005-01-01

    Loss of photosynthetic area can affect soil microbial communities by altering the availability of fixed carbon. We used denaturing gradient gel electrophoresis (DGGE) and Biolog filamentous-fungus plates to determine the effects of artificial defoliation of pines in a mixed pine-spruce forest on the composition of the fungal community in a forest soil. As measured by DGGE, two fungal species were affected significantly by the defoliation of pines (P < 0.001); the frequency of members of the ectomycorrhizal fungus genus Cenococcum decreased significantly, while the frequency of organisms of an unidentified soil fungus increased. The decrease in the amount of Cenococcum organisms may have occurred because of the formation of extensive hyphal networks by species of this genus, which require more of the carbon fixed by their host, or because this fungus is dependent upon quantitative differences in spruce root exudates. The defoliation of pines did not affect the overall composition of the soil fungal community or fungal-species richness (number of species per core). Biolog filamentous-fungus plate assays indicated a significant increase (P < 0.001) in the number of carbon substrates utilized by the soil fungi and the rate at which these substrates were used, which could indicate an increase in fungal-species richness. Thus, either small changes in the soil fungal community give rise to significant increases in physiological capabilities or PCR bias limits the reliability of the DGGE results. These data indicate that combined genetic and physiological assessments of the soil fungal community are needed to accurately assess the effect of disturbance on indigenous microbial systems.

  10. Improved white spruce (Picea glauca) genome assemblies and annotation of large gene families of conifer terpenoid and phenolic defense metabolism.

    PubMed

    Warren, René L; Keeling, Christopher I; Yuen, Macaire Man Saint; Raymond, Anthony; Taylor, Greg A; Vandervalk, Benjamin P; Mohamadi, Hamid; Paulino, Daniel; Chiu, Readman; Jackman, Shaun D; Robertson, Gordon; Yang, Chen; Boyle, Brian; Hoffmann, Margarete; Weigel, Detlef; Nelson, David R; Ritland, Carol; Isabel, Nathalie; Jaquish, Barry; Yanchuk, Alvin; Bousquet, Jean; Jones, Steven J M; MacKay, John; Birol, Inanc; Bohlmann, Joerg

    2015-07-01

    White spruce (Picea glauca), a gymnosperm tree, has been established as one of the models for conifer genomics. We describe the draft genome assemblies of two white spruce genotypes, PG29 and WS77111, innovative tools for the assembly of very large genomes, and the conifer genomics resources developed in this process. The two white spruce genotypes originate from distant geographic regions of western (PG29) and eastern (WS77111) North America, and represent elite trees in two Canadian tree-breeding programs. We present an update (V3 and V4) for a previously reported PG29 V2 draft genome assembly and introduce a second white spruce genome assembly for genotype WS77111. Assemblies of the PG29 and WS77111 genomes confirm the reconstructed white spruce genome size in the 20 Gbp range, and show broad synteny. Using the PG29 V3 assembly and additional white spruce genomics and transcriptomics resources, we performed MAKER-P annotation and meticulous expert annotation of very large gene families of conifer defense metabolism, the terpene synthases and cytochrome P450s. We also comprehensively annotated the white spruce mevalonate, methylerythritol phosphate and phenylpropanoid pathways. These analyses highlighted the large extent of gene and pseudogene duplications in a conifer genome, in particular for genes of secondary (i.e. specialized) metabolism, and the potential for gain and loss of function for defense and adaptation. PMID:26017574

  11. Improved white spruce (Picea glauca) genome assemblies and annotation of large gene families of conifer terpenoid and phenolic defense metabolism.

    PubMed

    Warren, René L; Keeling, Christopher I; Yuen, Macaire Man Saint; Raymond, Anthony; Taylor, Greg A; Vandervalk, Benjamin P; Mohamadi, Hamid; Paulino, Daniel; Chiu, Readman; Jackman, Shaun D; Robertson, Gordon; Yang, Chen; Boyle, Brian; Hoffmann, Margarete; Weigel, Detlef; Nelson, David R; Ritland, Carol; Isabel, Nathalie; Jaquish, Barry; Yanchuk, Alvin; Bousquet, Jean; Jones, Steven J M; MacKay, John; Birol, Inanc; Bohlmann, Joerg

    2015-07-01

    White spruce (Picea glauca), a gymnosperm tree, has been established as one of the models for conifer genomics. We describe the draft genome assemblies of two white spruce genotypes, PG29 and WS77111, innovative tools for the assembly of very large genomes, and the conifer genomics resources developed in this process. The two white spruce genotypes originate from distant geographic regions of western (PG29) and eastern (WS77111) North America, and represent elite trees in two Canadian tree-breeding programs. We present an update (V3 and V4) for a previously reported PG29 V2 draft genome assembly and introduce a second white spruce genome assembly for genotype WS77111. Assemblies of the PG29 and WS77111 genomes confirm the reconstructed white spruce genome size in the 20 Gbp range, and show broad synteny. Using the PG29 V3 assembly and additional white spruce genomics and transcriptomics resources, we performed MAKER-P annotation and meticulous expert annotation of very large gene families of conifer defense metabolism, the terpene synthases and cytochrome P450s. We also comprehensively annotated the white spruce mevalonate, methylerythritol phosphate and phenylpropanoid pathways. These analyses highlighted the large extent of gene and pseudogene duplications in a conifer genome, in particular for genes of secondary (i.e. specialized) metabolism, and the potential for gain and loss of function for defense and adaptation.

  12. Magnetic structure of polluted soil profiles from Eastern Ukraine

    NASA Astrophysics Data System (ADS)

    Jeleńska, Maria; Hasso-Agopsowicz, Agata; Kądziałko-Hofmokl, Magdalena; Kopcewicz, Barbara; Sukhorada, Anatoliy; Bondar, Ksenija; Matviishina, Zhanna

    2008-12-01

    Our study concerned magnetic properties of soil profiles taken from polluted regions of Eastern Ukraine around the industrial centres Krivyj Rig, Mariupol and Komsomolsk. Soils represent chernozem and podzoluvisol. The low-field magnetic susceptibility shows enhancement in the topsoil caused by contamination by coarse-grain magnetite connected with industrial pollution. Magnetic mineralogy was determined by means of thermal demagnetisation of SIRM, monitoring of susceptibility changes during warming from -196°C to room temperature and heating to 700°C, and Mössbauer analysis. Granulometry of magnetic particles was investigated by determination of hysteresis parameters, susceptibility, M s, SIRM and ARM ratios and frequency dependence of susceptibility. The chemical parameters, namely pH, organic carbon and iron content in different pedogenic and lithogenic minerals, measured for particular horizons determined pedogenic characteristic of soil profiles. Our study showed that differences in magnetic parameters of non-polluted and polluted soil profiles are not limited to the topsoil, but reach deep layers of the parent material. Industrial pollution promotes formation of the so-called "pedogenic" SP and SD particles in these layers.

  13. Red spruce physiology and growth in response to elevated CO[sub 2], water stress and nutrient limitations

    SciTech Connect

    Samuelson, L.J.

    1992-01-01

    Spruce-fir ecosystems of the eastern United States interest scientists because of reported changes in population growth. This research examined the growth and physical responses of red spruce seedlings (Picea rubens Sarg.) to change in atmospheric CO[sub 2], water and nutrient availability to determine the response of this species to potential climatic changes. Red spruce seedlings were grown from seed for 1 year in ambient (374 ppm) or elevated (713 ppm) CO[sub 2] in combination with low or high soil fertility treatment, and well-watered or water-stressed conditions. Red spruce seedlings grown with limited nutrient and water availability increased growth in elevated CO[sub 2] as did seedlings grown with high soil fertility treatment and ample water. At 12 months of age, elevated CO[sub 2]-grown seedlings had greater dry weight, height, diameter and specific leaf weight than ambient CO[sub 2[minus

  14. Ammonium assmilation in spruce ectomycorrhizas

    SciTech Connect

    Chalot, M.; Brun, A.; Botton, B. ); Stewart, G. )

    1990-05-01

    Assimilation of labelled NH{sub 4}{sup +} into amino acids has been followed in ectomycorrhizal roots of spruce. Over an 18 h period of NH{sub 4}{sup +} feeding, Gln, Glu and Ala became the most abundant amino acids. Gln was also the most highly labelled amino acid during the experiment, followed by Glu and Ala. This result indicates that Gln synthesis is an important ammonium utilization reaction in spruce mycorrhizas. Addition of MSX to NH{sub 4}{sup +} fed mycorrhizas caused an inhibition of Gln accumulation with a corresponding increase in Glu, Ala and Asn levels. The supply of MSX induced a sharp diminution of {sup 15}N enrichment in both amino and amido groups of glutamine. In contrast, the {sup 15}N incorporation into Glu and derivatives (Ala and Asp) remained very high. This study demonstrates that the fungal glutamate dehydrogenase is quite operative in spruce ectomycorrhizas since it is able to sustain ammonium assimilation when glutamine synthetase is inhibited.

  15. Space sequestration below ground in old-growth spruce-beech forests—signs for facilitation?

    PubMed Central

    Bolte, Andreas; Kampf, Friederike; Hilbrig, Lutz

    2013-01-01

    Scientists are currently debating the effects of mixing tree species for the complementary resource acquisition in forest ecosystems. In four unmanaged old-growth spruce-beech forests in strict nature reserves in southern Sweden and northern Germany we assessed forest structure and fine rooting profiles and traits (≤2 mm) by fine root sampling and the analysis of fine root morphology and biomass. These studies were conducted in selected tree groups with four different interspecific competition perspectives: (1) spruce as a central tree, (2) spruce as competitor, (3) beech as a central tree, and (4) beech as competitor. Mean values of life fine root attributes like biomass (FRB), length (FRL), and root area index (RAI) were significantly lower for spruce than for beech in mixed stands. Vertical profiles of fine root attributes adjusted to one unit of basal area (BA) exhibited partial root system stratification when central beech is growing with spruce competitors. In this constellation, beech was able to raise its specific root length (SRL) and therefore soil exploration efficiency in the subsoil, while increasing root biomass partitioning into deeper soil layers. According to relative values of fine root attributes (rFRA), asymmetric below-ground competition was observed favoring beech over spruce, in particular when central beech trees are admixed with spruce competitors. We conclude that beech fine rooting is facilitated in the presence of spruce by lowering competitive pressure compared to intraspecific competition whereas the competitive pressure for spruce is increased by beech admixture. Our findings underline the need of spatially differentiated approaches to assess interspecific competition below ground. Single-tree approaches and simulations of below-ground competition are required to focus rather on microsites populated by tree specimens as the basic spatial study area. PMID:24009616

  16. Space sequestration below ground in old-growth spruce-beech forests-signs for facilitation?

    PubMed

    Bolte, Andreas; Kampf, Friederike; Hilbrig, Lutz

    2013-01-01

    Scientists are currently debating the effects of mixing tree species for the complementary resource acquisition in forest ecosystems. In four unmanaged old-growth spruce-beech forests in strict nature reserves in southern Sweden and northern Germany we assessed forest structure and fine rooting profiles and traits (≤2 mm) by fine root sampling and the analysis of fine root morphology and biomass. These studies were conducted in selected tree groups with four different interspecific competition perspectives: (1) spruce as a central tree, (2) spruce as competitor, (3) beech as a central tree, and (4) beech as competitor. Mean values of life fine root attributes like biomass (FRB), length (FRL), and root area index (RAI) were significantly lower for spruce than for beech in mixed stands. Vertical profiles of fine root attributes adjusted to one unit of basal area (BA) exhibited partial root system stratification when central beech is growing with spruce competitors. In this constellation, beech was able to raise its specific root length (SRL) and therefore soil exploration efficiency in the subsoil, while increasing root biomass partitioning into deeper soil layers. According to relative values of fine root attributes (rFRA), asymmetric below-ground competition was observed favoring beech over spruce, in particular when central beech trees are admixed with spruce competitors. We conclude that beech fine rooting is facilitated in the presence of spruce by lowering competitive pressure compared to intraspecific competition whereas the competitive pressure for spruce is increased by beech admixture. Our findings underline the need of spatially differentiated approaches to assess interspecific competition below ground. Single-tree approaches and simulations of below-ground competition are required to focus rather on microsites populated by tree specimens as the basic spatial study area.

  17. Structural framework across the Bastar craton - the Eastern Ghats Granulite Belt interface: Implications for making of eastern Gondwana

    NASA Astrophysics Data System (ADS)

    Patole, Vishal; Nasipuri, Pritam

    2015-04-01

    The transformation of palaeo-continents involve breakup, dispersal and reassembly of cratonic blocks by collisional suturing that develop a network of orogenic (mobile) belts around the periphery of the stable cratons. During the collision, partial melting of the different crustal blocks produces migmatites at the craton-mobile belt interface. Thus, migmatites at the craton-mobile belt contact can provide valuable information regarding the pressure-temperature conditions of the melting of lower crust during supercontinent building processes. In this contribution, we document the structural framework across the Bastar craton- Eastern Ghats Granulite Belt (EGGB) interface that developed during the accretion of EGGB over Bastar craton. Near Bhawanipatna, Orissa, Eastern India, the granulites of the mobile belt are juxtaposed against the granitic rocks of the Bastar craton. Away from the contact domain, the cratonic granite is non-migmatitic and blasto-porphyritic in nature that gradually transforms to migmatitic variety towards the contact domain. In the non-migmatitic variety, the E-W trending stromatic leucozomes and biotite-hornblende rich fabric (S1) wraps around recrystallized K-feldspar augens. In the migmatitic variety towards the contact domain, NNE-SSW trending diatexite leucozomes (S2) are prominent and the intensity of melting and tightness of folding increases towards the contact domain. Structural measurements indicate that the S1 fabric is folded with the development of NNE-SSW axial plane with easterly plunging fold axis (50 -> 050N). To correlate the geological history of EGGB in the context of supercontinent reconstruction, the existence of a cratonic block consisting of India - Madagascar - Sri Lanka - Enderby Land-Kalahari ("IMSLEK") from 3000 Ma upto 750 Ma has been invoked by several authors. The apparent continuity of the Grenvillian metamorphic orogen along the East Antarctica-Australia-India margin has been taken as conclusive evidence for the

  18. The Social Consequences of Postcommunist Structural Change: An Analysis of Suicide Trends in Eastern Europe

    ERIC Educational Resources Information Center

    Minagawa, Yuka

    2013-01-01

    Guided by Durkheim's classic theory of suicide, this article examines suicide trends and determinants in Eastern European countries for the period of 1989-2006, with particular attention given to the association between postcommunist social change and suicide mortality. I find that countries characterized by more drastic structural change…

  19. Virulence structure of the eastern U.S. wheat powdery mildew population

    Technology Transfer Automated Retrieval System (TEKTRAN)

    Little is known about the population structure of wheat powdery mildew in the eastern 2 U.S., and the most recent report on virulence in this pathogen population involved isolates 3 collected in 1993-94. In the present study, wheat leaves naturally infected with powdery mildew 4 were collected from ...

  20. Selection effects of air pollution on gene pools of Norway spruce, European silver fir and European beech.

    PubMed

    Longauer, R; Gömöry, D; Paule, L; Karnosky, D F; Manikovská, B; Müller-Starck, G; Percy, K; Szaro, R

    2001-01-01

    The effects of industrial pollution on allelic and genotypic structures of Norway spruce. European silver fir and European beech were investigated by means of isozyme analysis. In a mixed Norway spruce-silver fir forest stand in an area heavily polluted by sulphur dioxide and heavy metals in the region of Spis (eastern Slovakia), pairs of neighbouring damaged and apparently healthy trees were selected in two replicates (44 and 69 pairs in a heavily and moderately damaged stand, respectively). Pairwise sampling of trees with contrasting vitality was applied to reduce potential effects of site heterogeneity on the vitality of sampled trees. No significant differences in allelic and genotypic frequencies were found between sets of healthy and declining trees. There were differences in the single-locus heterozygosities, but these were not consistent between the replicates. However, the set of damaged trees exhibited higher levels of genetic multiplicity and diversity, possibly due to the deleterious effect of rare alleles under the conditions of air pollution. Consequently. following the decline of pollutant-sensitive trees, the remaining stand will be depleted of a part of alleles with unknown adaptive value to future selection pressures.

  1. Organic matter characteristics in boreal forest soils under stands of silver birch, Norway spruce, and Norway spruce with a mixture of silver birch

    NASA Astrophysics Data System (ADS)

    Smolander, A.; Kitunen, V.

    2012-04-01

    The aim was to study how tree species and a tree species mixture affect microbial C and N transformations and two major plant secondary compound groups, terpenes and phenolic compounds in soil. The study site was a tree-species experiment in middle-eastern part of Finland containing plots of 43-year-old silver birch, Norway spruce and Norway spruce with a mixture of silver birch (22 and 37 % birch of the total stem number). Soil was podzol and humus type mor. Samples were taken from the organic layer. C and N in the microbial biomass, rates of C mineralization (CO2 evolution), net N mineralization and nitrification, and concentrations of total water-soluble phenolic compounds, condensed tannins and different kind of terpenes were measured. Amounts of C and N in the microbial biomass and the rates of C mineralization and net N mineralization were all lower under spruce than birch, and particularly net N mineralization was stimulated by birch mixture. Concentrations of total water-soluble phenolic compounds were on a similar level, irrespective of tree species. However, there were less low-molecular-weight phenolics and more high-molecular-weight phenolics under spruce than birch. Concentrations of condensed tannins and both sesqui- and diterpenes were all higher under spruce than birch but the concentrations of triterpenes were similar in all soils. The difference between tree species was greatest with monoterpenes which were measured from both organic layer and soil atmosphere: high concentrations under spruce and negligible under birch. Birch mixture tended to decrease the concentrations of condensed tannins and mono-, sesqui- and diterpenes.

  2. SPRUCE: Spruce and Peatland Responses under Climatic and Environmental Change

    DOE Data Explorer

    SPRUCE is an experiment to assess the response of northern peatland ecosystems to increases in temperature and exposures to elevated atmospheric CO2 concentrations. It is the primary component of the Terrestrial Ecosystem Science Scientific Focus Area of ORNL's Climate Change Program, focused on terrestrial ecosystems and the mechanisms that underlie their responses to climatic change. The experimental work is to be conducted in a Picea mariana [black spruce] - Sphagnum spp. bog forest in northern Minnesota, 40 km north of Grand Rapids, in the USDA Forest Service Marcell Experimental Forest (MEF). The site is located at the southern margin of the boreal peatland forest. It is an ecosystem considered especially vulnerable to climate change, and anticipated to be near its tipping point with respect to climate change. Responses to warming and interactions with increased atmospheric CO2 concentration are anticipated to have important feedbacks on the atmosphere and climate, because of the high carbon stocks harbored by such ecosystems.[copied from http://mnspruce.ornl.gov/] While some data files are restricted to access by project members only, others are available for public download now, even as research is being actively conducted.

  3. Stand and landscape level effects of a major outbreak of spruce beetles on forest vegetation in the Copper River Basin, Alaska

    USGS Publications Warehouse

    Allen, J.L.; Wesser, S.; Markon, C.J.; Winterberger, K.C.

    2006-01-01

    From 1989 to 2003, a widespread outbreak of spruce beetles (Dendroctonus rufipennis) in the Copper River Basin, Alaska, infested over 275,000 ha of forests in the region. During 1997 and 1998, we measured forest vegetation structure and composition on one hundred and thirty-six 20-m ?? 20-m plots to assess both the immediate stand and landscape level effects of the spruce beetle infestation. A photo-interpreted vegetation and infestation map was produced using color-infrared aerial photography at a scale of 1:40,000. We used linear regression to quantify the effects of the outbreak on forest structure and composition. White spruce (Picea glauca) canopy cover and basal area of medium-to-large trees [???15 cm diameter-at-breast height (1.3 m, dbh)] were reduced linearly as the number of trees attacked by spruce beetles increased. Black spruce (Picea mariana) and small diameter white spruce (<15 cm dbh) were infrequently attacked and killed by spruce beetles. This selective attack of mature white spruce reduced structural complexity of stands to earlier stages of succession and caused mixed tree species stands to lose their white spruce and become more homogeneous in overstory composition. Using the resulting regressions, we developed a transition matrix to describe changes in vegetation types under varying levels of spruce beetle infestations, and applied the model to the vegetation map. Prior to the outbreak, our study area was composed primarily of stands of mixed white and black spruce (29% of area) and pure white spruce (25%). However, the selective attack on white spruce caused many of these stands to transition to black spruce dominated stands (73% increase in area) or shrublands (26% increase in area). The post-infestation landscape was thereby composed of more even distributions of shrubland and white, black, and mixed spruce communities (17-22% of study area). Changes in the cover and composition of understory vegetation were less evident in this study

  4. Integrating remote sensing and magnetic data for structural geology investigation in pegmatite areas in eastern Afghanistan

    NASA Astrophysics Data System (ADS)

    Salehi, Ratib; Saadi, Nureddin M.; Khalil, Ahmed; Watanabe, Koichiro

    2015-01-01

    This study used an integrated approach to investigate pegmatite areas in eastern Afghanistan. The analysis of surface data, including a digital elevation model (DEM), and Landsat Enhanced Thematic Mapper Plus (ETM+) images, was combined with airborne magnetic data to better understand three-dimensional geology in the area. The ETM+ and DEM data were used to map geological structures at the surface, which indicate that the area consists of two main fault systems that trend NNE and E-W. The two trends represent the remnants of reactivated structures that formed under the stress regimes generated during the tectonic evolution of eastern Afghanistan. Magnetic data indicate an NE-SW trending basin. A two-dimensional schematic model shows that the basin gradually deepens toward the SW with depths to the magnetic basement ranging between 2 and 11.5 km. The integration of the results gave new insight into the tectonic evolution and structure patterns near the pegmatites area.

  5. The extent and meaning of hybridization and introgression between Siberian spruce (Picea obovata) and Norway spruce (Picea abies): cryptic refugia as stepping stones to the west?

    PubMed

    Tsuda, Yoshiaki; Chen, Jun; Stocks, Michael; Källman, Thomas; Sønstebø, Jørn Henrik; Parducci, Laura; Semerikov, Vladimir; Sperisen, Christoph; Politov, Dmitry; Ronkainen, Tiina; Väliranta, Minna; Vendramin, Giovanni Giuseppe; Tollefsrud, Mari Mette; Lascoux, Martin

    2016-06-01

    Boreal species were repeatedly exposed to ice ages and went through cycles of contraction and expansion while sister species alternated periods of contact and isolation. The resulting genetic structure is consequently complex, and demographic inferences are intrinsically challenging. The range of Norway spruce (Picea abies) and Siberian spruce (Picea obovata) covers most of northern Eurasia; yet their geographical limits and histories remain poorly understood. To delineate the hybrid zone between the two species and reconstruct their joint demographic history, we analysed variation at nuclear SSR and mitochondrial DNA in 102 and 88 populations, respectively. The dynamics of the hybrid zone was analysed with approximate Bayesian computation (ABC) followed by posterior predictive structure plot reconstruction and the presence of barriers across the range tested with estimated effective migration surfaces. To estimate the divergence time between the two species, nuclear sequences from two well-separated populations of each species were analysed with ABC. Two main barriers divide the range of the two species: one corresponds to the hybrid zone between them, and the other separates the southern and northern domains of Norway spruce. The hybrid zone is centred on the Urals, but the genetic impact of Siberian spruce extends further west. The joint distribution of mitochondrial and nuclear variation indicates an introgression of mitochondrial DNA from Norway spruce into Siberian spruce. Overall, our data reveal a demographic history where the two species interacted frequently and where migrants originating from the Urals and the West Siberian Plain recolonized northern Russia and Scandinavia using scattered refugial populations of Norway spruce as stepping stones towards the west. PMID:27087633

  6. The extent and meaning of hybridization and introgression between Siberian spruce (Picea obovata) and Norway spruce (Picea abies): cryptic refugia as stepping stones to the west?

    PubMed

    Tsuda, Yoshiaki; Chen, Jun; Stocks, Michael; Källman, Thomas; Sønstebø, Jørn Henrik; Parducci, Laura; Semerikov, Vladimir; Sperisen, Christoph; Politov, Dmitry; Ronkainen, Tiina; Väliranta, Minna; Vendramin, Giovanni Giuseppe; Tollefsrud, Mari Mette; Lascoux, Martin

    2016-06-01

    Boreal species were repeatedly exposed to ice ages and went through cycles of contraction and expansion while sister species alternated periods of contact and isolation. The resulting genetic structure is consequently complex, and demographic inferences are intrinsically challenging. The range of Norway spruce (Picea abies) and Siberian spruce (Picea obovata) covers most of northern Eurasia; yet their geographical limits and histories remain poorly understood. To delineate the hybrid zone between the two species and reconstruct their joint demographic history, we analysed variation at nuclear SSR and mitochondrial DNA in 102 and 88 populations, respectively. The dynamics of the hybrid zone was analysed with approximate Bayesian computation (ABC) followed by posterior predictive structure plot reconstruction and the presence of barriers across the range tested with estimated effective migration surfaces. To estimate the divergence time between the two species, nuclear sequences from two well-separated populations of each species were analysed with ABC. Two main barriers divide the range of the two species: one corresponds to the hybrid zone between them, and the other separates the southern and northern domains of Norway spruce. The hybrid zone is centred on the Urals, but the genetic impact of Siberian spruce extends further west. The joint distribution of mitochondrial and nuclear variation indicates an introgression of mitochondrial DNA from Norway spruce into Siberian spruce. Overall, our data reveal a demographic history where the two species interacted frequently and where migrants originating from the Urals and the West Siberian Plain recolonized northern Russia and Scandinavia using scattered refugial populations of Norway spruce as stepping stones towards the west.

  7. Mantle Structures between Guatemala and the Eastern USArray

    NASA Astrophysics Data System (ADS)

    Chu, R.; Helmberger, D. V.; Wei, S.

    2015-12-01

    While cratons are stable features based on their ages, they apparently can be disturbed by flattened slabs and hidden hotspots. When these slabs break up they provide a host of volatiles which interact with the old lithosphere producing complicated structures. These have been imaged by a host of techniques involving modeling regional waveforms and an assortment of tomography models. Here we test these models against a deep March 25th, 2013 Guatemala event at a depth of 190 km, which produced an excellent set of TA record sections that provide dense sampling of the structure, spanning the ranges from 15 to 35 degrees. Because the structure in the southern Gulf of Mexico is uncertain, we first concentrated on deriving a pure-path model for this region. Both P and SH triplication data is well sampled by stations located in Florida from 13 degrees and beyond. The best fitting model is modified from the SH model ATL (Grand and Helmberger, 1984) with a P-model in Zhao and Helmberger (1993). These models contain a high-velocity lid structure along with a low-velocity gradient at depth. This allows the up-going S wave (direct) to remain the first arrival traveling in the fast lid to beyond 180 degrees. The modeling of the direct S along with the diving phases and depth phase triplications, pP, sP, sS allows for a well constrained model. This model ATLm and the earlier craton model CR along with mixture of the two are tested with the Cut-and-Paste (CAP) code on whole 3 component records. Over large paths of data produced by the southern US support the so-called X-phase arrivals which are produced by a 300 km transition. This feature breas down at large distance (>25 degree) along a northwest-southeast boundary. Data at larger distances indicate a complexity in the 660 transition which appears to be caused by a fast slab located at the top of the lower mantle. Variation in the EF-CD branches of pP and sP display changes of up to 5 s and sS up to 10 s, associated with the

  8. Hierarchical analysis of genetic structure in the habitat-specialist Eastern Sand Darter (Ammocrypta pellucida)

    PubMed Central

    Ginson, Robert; Walter, Ryan P; Mandrak, Nicholas E; Beneteau, Courtney L; Heath, Daniel D

    2015-01-01

    Quantifying spatial genetic structure can reveal the relative influences of contemporary and historic factors underlying localized and regional patterns of genetic diversity and gene flow – important considerations for the development of effective conservation efforts. Using 10 polymorphic microsatellite loci, we characterize genetic variation among populations across the range of the Eastern Sand Darter (Ammocrypta pellucida), a small riverine percid that is highly dependent on sandy substrate microhabitats. We tested for fine scale, regional, and historic patterns of genetic structure. As expected, significant differentiation was detected among rivers within drainages and among drainages. At finer scales, an unexpected lack of within-river genetic structure among fragmented sandy microhabitats suggests that stratified dispersal resulting from unstable sand bar habitat degradation (natural and anthropogenic) may preclude substantial genetic differentiation within rivers. Among-drainage genetic structure indicates that postglacial (14 kya) drainage connectivity continues to influence contemporary genetic structure among Eastern Sand Darter populations in southern Ontario. These results provide an unexpected contrast to other benthic riverine fish in the Great Lakes drainage and suggest that habitat-specific fishes, such as the Eastern Sand Darter, can evolve dispersal strategies that overcome fragmented and temporally unstable habitats. PMID:25691991

  9. Clinal Variation at Phenology-Related Genes in Spruce: Parallel Evolution in FTL2 and Gigantea?

    PubMed Central

    Chen, Jun; Tsuda, Yoshiaki; Stocks, Michael; Källman, Thomas; Xu, Nannan; Kärkkäinen, Katri; Huotari, Tea; Semerikov, Vladimir L.; Vendramin, Giovanni G.; Lascoux, Martin

    2014-01-01

    Parallel clines in different species, or in different geographical regions of the same species, are an important source of information on the genetic basis of local adaptation. We recently detected latitudinal clines in SNPs frequencies and gene expression of candidate genes for growth cessation in Scandinavian populations of Norway spruce (Picea abies). Here we test whether the same clines are also present in Siberian spruce (P. obovata), a close relative of Norway spruce with a different Quaternary history. We sequenced nine candidate genes and 27 control loci and genotyped 14 SSR loci in six populations of P. obovata located along the Yenisei river from latitude 56°N to latitude 67°N. In contrast to Scandinavian Norway spruce that both departs from the standard neutral model (SNM) and shows a clear population structure, Siberian spruce populations along the Yenisei do not depart from the SNM and are genetically unstructured. Nonetheless, as in Norway spruce, growth cessation is significantly clinal. Polymorphisms in photoperiodic (FTL2) and circadian clock (Gigantea, GI, PRR3) genes also show significant clinal variation and/or evidence of local selection. In GI, one of the variants is the same as in Norway spruce. Finally, a strong cline in gene expression is observed for FTL2, but not for GI. These results, together with recent physiological studies, confirm the key role played by FTL2 and circadian clock genes in the control of growth cessation in spruce species and suggest the presence of parallel adaptation in these two species. PMID:24814465

  10. Eastern rim of the Chesapeake Bay impact crater: Morphology, stratigraphy, and structure

    USGS Publications Warehouse

    Poag, C.W.

    2005-01-01

    This study reexamines seven reprocessed (increased vertical exaggeration) seismic reflection profiles that cross the eastern rim of the Chesapeake Bay impact crater. The eastern rim is expressed as an arcuate ridge that borders the crater in a fashion typical of the "raised" rim documented in many well preserved complex impact craters. The inner boundary of the eastern rim (rim wall) is formed by a series of raterfacing, steep scarps, 15-60 m high. In combination, these rim-wall scarps represent the footwalls of a system of crater-encircling normal faults, which are downthrown toward the crater. Outboard of the rim wall are several additional normal-fault blocks, whose bounding faults trend approximately parallel to the rim wall. The tops of the outboard fault blocks form two distinct, parallel, flat or gently sloping, terraces. The innermost terrace (Terrace 1) can be identified on each profile, but Terrace 2 is only sporadically present. The terraced fault blocks are composed mainly of nonmarine, poorly to moderately consolidated, siliciclastic sediments, belonging to the Lower Cretaceous Potomac Formation. Though the ridge-forming geometry of the eastern rim gives the appearance of a raised compressional feature, no compelling evidence of compressive forces is evident in the profiles studied. The structural mode, instead, is that of extension, with the clear dominance of normal faulting as the extensional mechanism. ?? 2005 Geological Society of America.

  11. Structural interpretation of seismic reflection data from eastern Salt Range and Potwar Plateau, Pakistan

    SciTech Connect

    Pennock, E.S.; Lillie, R.J.; Zaman, A.S.H.; Yousaf, M.

    1989-07-01

    Approximately 1600 km of seismic reflection profiles from the eastern Salt Range and Potwar Plateau (SR/PP) of Pakistan is integrated with available magnetostratigraphic, surface geologic, and well data to classify structural styles, determine the timing of deformation, and estimate the amount of telescoping of the sedimentary cover. The eastern SR/PP are similar to other fold-and-thrust belts underlain by evaporites in that (1) it is part of a zone of overthrusting that extends considerably farther over the Himalayan foreland than adjacent areas not underlain by evaporites, (2) the overall thrust wedge has a narrow cross-sectional taper, (3) structures verge toward the hinterland as well as toward the foreland, and (4) fold trends are long and continuous, consisting of tight salt-cored anticlines separated by broad synclines. 11 figures, 1 table.

  12. Reconstructing the landscape structure of the Lena-Angara interfluve (south part of Eastern Siberia)

    NASA Astrophysics Data System (ADS)

    Atutova, Zhanna

    2015-04-01

    Historical-geographical reconstructions of the landscape structure of territories developed in the remote past constitute the necessary element in the chain of research into the dynamics and the degree of transformation of geosystems caused by the influence of the natural regularities and anthropogenic factors. The objective of this study is to determine the specific features of the territory of the Lena-Angara interfluve in the late 19th - early 20th centuries in the interest of a subsequent different-time comparative analysis of the landscape situation. An analysis of the features inherent in the functioning of the geosystems of the Lena-Angara interfluve was made by using, as an example, an elevated plateau with the sources of the Kuda river as well as of the Ilga and Kuda rivers. The relief is represented by a tableland with narrow crests of the watersheds, heavily dissected by a dense network of the valleys of rivers. The denudation processes created planate table-shaped elevations and plateaus whose range of absolute altitudes varies between 400 and 1000 m. The analysis of the landscape structure showed that the study territory was the home for mountain-taiga dark-coniferous and deciduous classes of facies. Larch, spruce-larch and, in places, pine-larch subshrub-grass-moss forests grew within the basins of the Ilga and Kulenga rivers. The watershed spaces of the Ilga-Kuda interfluve, and also the slopes of the upper reaches of the Kuda river were occupied by Siberian stone pine and larch-spruce subshrub-moss groups of facies. In spite of the ubiquitous occurrence of taiga-forest ranges, most of them transformed to derivative groups of facies. Forest fires gave impetus to a widespread occurrence of coniferous/small-leaved complexes in burned-over areas. The study area was poorly populated at the period under investigation; therefore, cultivated lands occupied very small territories. The upper reaches of the Kulenga river included small tracts of arable land

  13. Uppermost mantle structure beneath eastern China and its surroundings from Pn and Sn tomography

    NASA Astrophysics Data System (ADS)

    Sun, Weijia; Kennett, B. L. N.

    2016-04-01

    The Pn and Sn residuals from regional events provide strong constraints on the structure and lithological characteristics of the uppermost mantle beneath eastern China and its surroundings. With the dense Chinese Digital Seismic Network in eastern China, separate Pn and Sn tomographic inversions have been exploited to obtain P and S velocities at a resolution of 2° × 2° or better. The patterns of P velocities are quite consistent with the S velocities at depth of 50 and 60 km, but the amplitude of P wave speed anomalies are a little larger than those of S wave speed. The low P wave speed, high S wave speed, and low Vp/Vs ratio beneath the northern part of Ordos Basin are related to upwelling hot material. Abrupt changes in material properties are indicated from the rapid variations in the Vp/Vs ratio.

  14. Crustal structure beneath western and eastern Iceland from surface waves and receiver functions

    USGS Publications Warehouse

    Du, Z.; Foulger, G.R.; Julian, B.R.; Allen, R.M.; Nolet, G.; Morgan, W.J.; Bergsson, B.H.; Erlendsson, P.; Jakobsdottir, S.; Ragnarsson, S.; Stefansson, R.; Vogfjord, K.

    2002-01-01

    We determine the crustal structures beneath 14 broad-band seismic stations, deployed in western, eastern, central and southern Iceland, using surface wave dispersion curves and receiver functions. We implement a method to invert receiver functions using constraints obtained from genetic algorithm inversion of surface waves. Our final models satisfy both data sets. The thickness of the upper crust, as defined by the velocity horizon Vs = 3.7 km s-1, is fairly uniform at ???6.5-9 km beneath the Tertiary intraplate areas of western and eastern Iceland, and unusually thick at 11 km beneath station HOT22 in the far south of Iceland. The depth to the base of the lower crust, as defined by the velocity horizon Vs = 4.1 km s-1 is ???20-26 km in western Iceland and ???27-33 km in eastern Iceland. These results agree with those of explosion profiles that detect a thinner crust beneath western Iceland than beneath eastern Iceland. An earlier report of a substantial low-velocity zone beneath the Middle Volcanic Zone in the lower crust is confirmed by a similar observation beneath an additional station there. As was found in previous receiver function studies, the most reliable feature of the results is the clear division into an upper sequence that is a few kilometres thick where velocity gradients are high, and a lower, thicker sequence where velocity gradients are low. The transition to typical mantle velocities is variable, and may range from being very gradational to being relatively sharp and clear. A clear Moho, by any definition, is rarely seen, and there is thus uncertainty in estimates of the thickness of the crust in many areas. Although a great deal of seismic data are now available constraining the structures of the crust and upper mantle beneath Iceland, their geological nature is not well understood.

  15. Structural and tectonic evolution of the eastern Cayman Trough (Caribbean Sea) from seismic reflection data

    SciTech Connect

    Leroy, S.; Mauffret, A.; Pubellier, M.

    1996-02-01

    The eastern Cayman Trough preserves a record of the Late Cretaceous to Paleogene Caribbean history that is largely affected by Neogene strike-slip tectonics of the current plate boundary. We conducted an analysis of seismic data within the eastern Cayman Trough, based upon single and multi-channel seismic reflection profiles collected during the Seacarib II cruise in 1987 and the Casis cruise in 1992. These data show that the basement of the eastern Cayman Trough can be divided into four domains from east to west, with distinct morphologic and sedimentary character and inferred older to younger ages: (1) a province of rifted Mesozoic continental crust exhibiting seven parallel horst blocks striking northeast-southwest; (2) a continent-ocean transition between provinces 1 and 3 that exhibits seamounts, small hills, and sedimentary basins; (3) an Eocene oceanic crust with rough basement but smoother relief than the rifted crust; basement trends are roughly north-south and oblique to the northwest trend in domain 1, and (4) the northern Jamaica slope, which forms an east-west-trending slope, with northward-dipping strata that flank the three deeper water domains of the Cayman Trough. The domains are interpreted to be the product of the Eocene east-west opening of the Cayman Trough as a pull-apart basin in a left-lateral strike-slip setting. Closure of the 1100 km of Eocene and younger oceanic crust of the Cayman Trough places the fault-block province adjacent to the Belize margin of Central America. A Neogene phase of transpression has reactivated structures in the four domains, along with on-land structures described by previous authors in Jamaica. The proximity of the eastern margin of the Cayman Trough to petroliferous, continental rocks in Central America suggests an improved possibility of hydrocarbon potential. Unfortunately, sediment thicknesses of less than 1 km probably are not conducive to hydrocarbon formation.

  16. Factors affecting spruce establishment and recruitment near western treeline, Alaska

    NASA Astrophysics Data System (ADS)

    Miller, A. E.; Sherriff, R.; Wilson, T. L.

    2015-12-01

    Regional warming and increases in tree growth are contributing to increased productivity near the western forest margin in Alaska. The effects of warming on seedling recruitment has received little attention, in spite of forecasted forest expansion near western treeline. Here, we used stand structure and environmental data from white spruce (Picea glauca) stands (n = 95) sampled across a longitudinal gradient to explore factors influencing white spruce growth, establishment and recruitment in southwest Alaska. Using tree-ring chronologies developed from a subset of the plots (n = 30), we estimated establishment dates and basal area increment (BAI) for trees of all age classes across a range of site conditions. We used GLMs (generalized linear models) to explore the relationship between tree growth and temperature in undisturbed, low elevation sites along the gradient, using BAI averaged over the years 1975-2000. In addition, we examined the relationship between growing degree days (GDD) and seedling establishment over the previous three decades. We used total counts of live seedlings, saplings and live and dead trees, representing four cohorts, to evaluate whether geospatial, climate, and measured plot covariates predicted abundance of the different size classes. We hypothesized that the relationship between abundance and longitude would vary by size class, and that this relationship would be mediated by growing season temperature. We found that mean BAI for trees in undisturbed, low elevation sites increased with July maximum temperature, and that the slope of the relationship with temperature changed with longitude (interaction significant with 90% confidence). White spruce establishment was positively associated with longer summers and/or greater heat accumulation, as inferred from GDD. Seedling, sapling and tree abundance were also positively correlated with temperature across the study area. The response to longitude was mixed, with smaller size classes

  17. White Spruce Regeneration Following a Major Spruce Beetle Outbreak in Forests on the Kenai Peninsula, Alaska

    Technology Transfer Automated Retrieval System (TEKTRAN)

    Between 1987 and 2000, a spruce beetle (Dendroctonus rufipennis) epidemic infested 1.19 million hectares of spruce (Picea spp.) forests in Alaska, killing most of the large diameter trees. We evaluated whether these forests would recover to their pre-outbreak density, and determined the site conditi...

  18. Growth strategy of Norway spruce under air elevated [CO2

    NASA Astrophysics Data System (ADS)

    Pokorny, R.; Urban, O.; Holisova, P.; Sprtova, M.; Sigut, L.; Slipkova, R.

    2012-04-01

    Plants will respond to globally increasing atmospheric CO2 concentration ([CO2]) by acclimation or adaptation at physiological and morphological levels. Considering the temporal onset, physiological responses may be categorized as short-term and morphological ones as long-term responses. The degree of plant growth responses, including cell division and cell expansion, is highly variable. It depends mainly on the specie's genetic predisposition, environment, mineral nutrition status, duration of CO2 enrichment, and/or synergetic effects of other stresses. Elevated [CO2] causes changes in tissue anatomy, quantity, size, shape and spatial orientation and can result in altered sink strength. Since, there are many experimental facilities for the investigation of elevated [CO2] effects on trees: i) closed systems or open top chambers (OTCs), ii) semi-open systems (for example glass domes with adjustable lamella windows - DAWs), and iii) free-air [CO2] enrichments (FACE); the results are still unsatisfactory due to: i) relatively short-term duration of experiments, ii) cultivation of young plants with different growth strategy comparing to old ones, iii) plant cultivation under artificial soil and weather conditions, and iv) in non-representative stand structure. In this contribution we are discussing the physiological and morphological responses of Norway spruce trees cultivated in DAWs during eight consecutive growing seasons in the context with other results from Norway spruce cultivation under air-elevated [CO2] conditions. On the level of physiological responses, we discuss the changes in the rate of CO2 assimilation, assimilation capacity, photorespiration, dark respiration, stomatal conductance, water potential and transpiration, and the sensitivity of these physiological processes to temperature. On the level of morphological responses, we discuss the changes in bud and growth phenology, needle and shoot morphology, architecture of crown and root system, wood

  19. Subducting slab structure below the eastern Sunda arc inferred from non-linear seismic tomographic imaging

    NASA Astrophysics Data System (ADS)

    Widiyantoro, S.; Pesicek, J. D.; Thurber, C. H.

    2011-12-01

    Detailed P-wave speed velocity structure beneath the Sunda arc has been successfully imaged by applying a non-linear approach to seismic tomography. Nearly one million compressional phases from events within the Indonesian region have been used. These include the surface-reflected depth phases pP and pwP in order to improve the sampling of the uppermantle structure, particularly below the back-arc regions. We have combined a high-resolution regional inversion with a low-resolution global inversion to minimize the mapping of distant aspherical mantle structure into the study region. In this paper, we focus our discussion on the upper mantle structure beneath the eastern part of the Sunda arc. The tomographic images confirm previous observations of a hole in the subducted slab in the upper mantle beneath eastern Java. The images also suggest that a tear in the slab exists below the easternmost part of the Sunda arc, where the down-going slab is deflected in the mantle transition zone. In good agreement with previous studies, the properties of the deflected slab show a strong bulk-sound signature.

  20. The impact of family structure and disruption on intergenerational emotional exchange in Eastern Europe.

    PubMed

    Moor, Nienke; Komter, Aafke

    2012-06-01

    Demographic trends across Europe involve a decrease in fertility and mortality rates, and an increase in divorce and stepfamily formation. Life courses and living arrangements have become less standardized and the structure of families has changed. In this article, we examine to what extent contemporary family structure and composition resulting from demographic changes affect emotional exchange between children and their parents, both from adult child to parent and from parent to child. Because the general level of well-being has been shown to be lower in Eastern Europe, thereby potentially affecting emotional exchange within families, we focus our research on Eastern Europe. We use the "conservation of resources theory" to derive hypotheses on how family structure may affect intergenerational emotional exchange. Family ties are assumed to be important resources of affection that people want to obtain and retain throughout their lives. Data from the Generations and Gender Survey (GGS) are used to test our hypotheses. In general, our data offer more support for the idea that families are resilient than for the often heard assumption that families are in decline as a consequence of the changed family structure and composition.

  1. Structurally controlled 'teleconnection' of large-scale mass wasting (Eastern Alps)

    NASA Astrophysics Data System (ADS)

    Ostermann, Marc; Sanders, Diethard

    2015-04-01

    In the Brenner Pass area (Eastern Alps) , closely ahead of the most northward outlier ('nose') of the Southern-Alpine continental indenter, abundant deep-seated gravitational slope deformations and a cluster of five post-glacial rockslides are present. The indenter of roughly triangular shape formed during Neogene collision of the Southern-Alpine basement with the Eastern-Alpine nappe stack. Compression by the indenter activated a N-S striking, roughly W-E extensional fault northward of the nose of the indenter (Brenner-normal fault; BNF), and lengthened the Eastern-Alpine edifice along a set of major strike-slip faults. These fault zones display high seismicity, and are the preferred locus of catastrophic rapid slope failures (rockslides, rock avalanches) and deep-seated gravitational slope deformations. The seismotectonic stress field, earthquake activity, and structural data all indicate that the South-Alpine indenter still - or again - exerts compression; in consequence, the northward adjacent Eastern Alps are subject mainly to extension and strike-slip. For the rockslides in the Brenner Pass area, and for the deep-seated gravitational slope deformations, the fault zones combined with high seismic activity predispose massive slope failures. Structural data and earthquakes mainly record ~W-E extension within an Eastern Alpine basement block (Oetztal-Stubai basement complex) in the hangingwall of the BNF. In the Northern Calcareous Alps NW of the Oetztal-Stubai basement complex, dextral faults provide defacement scars for large rockfalls and rockslides. Towards the West, these dextral faults merge into a NNW-SSE striking sinistral fault zone that, in turn, displays high seismic activity and is the locus of another rockslide cluster (Fern Pass cluster; Prager et al., 2008). By its kinematics dictated by the South-Alpine indenter, the relatively rigid Oetztal-Stubai basement block relays faulting and associated mass-wasting over a N-S distance of more than 60

  2. Black spruce growth forms as a record of a changing winter environment at treeline, Quebec, Canada

    SciTech Connect

    Lavoie, C.; Payette, S. )

    1992-02-01

    The environmental conditions prevailing at treeline in subarctic Quebec have been reconstructed over the past 400 yr through a comparative analysis of tree rings and growth forms of black spruce (Picea mariana (Mill.) B.S.P.). Because black spruce growth forms are closely associated with the winter environment, they are a direct response to conditions of low temperature and windblown snow abrasion affecting living tissues at the snow-air interface. The age structure of supranival shoot populations was closely associated with periods of higher stem survival in winter most likely under snowier and windless conditions. Spruce growth on slopes and in the valley revealed periods of low tree-ring growth between 1601 and 1663 and between 1700 and 1904, respectively. A long-lasting period of low radial growth 1697 and 1939 prevailed in the hilltop site. During the 20th century, spruce height increased from 0.8 to 1.6 m on slopes and in the valley, while the basal level of abrasion from windblown snow increased from 0.1 to 0.5 m, suggesting an increasing trend towards warmer and snowier conditions. Abraded spruces growing during the Little Ice Age (1570-1880) were replaced by symmetrical trees during the 20th century. Supranival skirted and whorled spruces which dominated on the hilltop site during the 16th century reverted to infranival cushion and mat growth forms during the Little Ice Age. These stunted spruces were unable to recover during the recent warming because of their inability to catch enough drifting snow to allow vertical growth.

  3. 9. LOOKING NORTH ON SPRUCE STREET, SHOWING MILLWRIGHT SHOP, FITTING ...

    Library of Congress Historic Buildings Survey, Historic Engineering Record, Historic Landscapes Survey

    9. LOOKING NORTH ON SPRUCE STREET, SHOWING MILLWRIGHT SHOP, FITTING SHOP, ADMINISTRATION BUILDING AND ERECTING SHOP - UNION WORKS IN BACKGROUND. - Rogers Locomotive & Machine Works, Spruce & Market Streets, Paterson, Passaic County, NJ

  4. Hydrodynamic and Thermohaline Seasonal Structures of Peninsular Malaysia's eastern continental shelf sea

    NASA Astrophysics Data System (ADS)

    Daryabor, Farshid; Tangang, Fredolin; Juneng, Liew

    2010-05-01

    The seasonal ocean circulation and the seasonal thermal structure in Peninsular Malaysia's eastern continental shelf sea were studied numerically using the Regional Ocean Model System (ROMS) with 18-km horizontal resolution and 50 sigma levels conforming to a realistic bottom topography. A 10-year control run was performed using climatological monthly mean wind stresses, restoring-type surface salt, temp and heat, and observational oceanic inflow/outflow at the open boundaries. The seasonally averaged effects of isolated forcing terms are presented and analyzed from the following experiments: 1) heat fluxes effects removed and 2) wind effects removed. This procedure allowed analysis of the contribution of individual parameters to the general hydrology and specific features of the Peninsular Malaysia's eastern continental shelf sea: for example, coastal jets, mesoscale topographic gyres, and countercurrents. The results show that the ROMS model has the capability of simulating seasonal variations of the Peninsular Malaysia's eastern continental shelf sea circulation and thermohaline structure. The simulated Peninsular Malaysia's eastern continental shelf sea surface circulation is generally anticyclonic (cyclonic) during the summer (winter) monsoon period with a strong western boundary current, a mean maximum speed of 0.8 m s-1 (0.5 m s-1), and extending to a depth of around 30 m (40 m). During summer, the western boundary current splits and partially leaves the coast; the bifurcation point is at 40°N in Jun and shifts north to 70°N in July. A mesoscale eddy on the Sunda shelf, the southwest of Natuna Island at (2.50°N, 1060°E) and extending to two mesoscale eddy at (20°N, 1070°E) and (2.50°N, 1090°E) also on the Gulf of Thailand (7.50°N, 1040°E), was also simulated. These eddies are cyclonic (anticyclonic) with maximum swirl velocity of 0.3 m s-1 at the peak of the summer (winter) monsoon. The simulated thermohaline structure for summer and winter are

  5. TopoGreenland: crustal structure in central-eastern Greenland along a new refraction profile

    NASA Astrophysics Data System (ADS)

    Shulgin, Alexey; Thybo, Hans; Field Team TopoGreenland

    2013-04-01

    We present the seismic structure in the interior of Greenland based on the first measurements by the seismic refraction/wide angle reflection method. Previous seismic surveys have only been carried out offshore and near the coast of Greenland, where the crustal structure is affected by oceanic break-up and may not be representative of the interior of the island. Acquisition of geophysical data in onshore Greenland is logistically complicated by the presence of an up to 3.4 km thick ice sheet, permanently covering most of the land mass. The seismic data was acquired by a team of six people during a two-month long experiment in summer of 2011 on the ice cap in the interior of central-eastern Greenland. The EW-trending profile extends 310 km inland from the approximate edge of the stable ice cap near Scoresby Sund across the center of the ice cap. The planned extension of the profile by use of OBSs and air gun shooting in Scoresbysund Fjord to the east coast of Greenland was unfortunately canceled, because navigation was prevented by ice drift. 350 Reftek Texan receivers recorded high-quality seismic data from 8 equidistant shots along the profile. Explosive charge sizes were 1 ton at the ends and ca. 500 kg along the profile, loaded with about 125 kg at 35-85 m depth in individual boreholes. Two-dimensional velocity model based on tomographic inversion and forward ray tracing modeling shows a decrease of crustal thickness from 47 km below the center of Greenland in the western part to 40 km in the eastern part of the profile. Earlier studies show that crustal thickness further decreases eastward to ca. 30 km below the fjord system, but details of the changes are unknown. Relatively high lower crustal velocities (Vp 6.8 - 7.3) in the western part of the TopoGreenland profile may indicate past collision tectonics or may be related or to the passage of the Iceland mantle plume. The origin of the pronounced circum-Atlantic mountain ranges in Norway and eastern Greenland

  6. Early Mesozoic structural evolution of the eastern West Qinling, northwest China

    NASA Astrophysics Data System (ADS)

    Wu, Guo-Li; Meng, Qing-Ren; Duan, Liang; Li, Lin

    2014-09-01

    This paper aims to reconstruct Early Mesozoic structural development of the eastern West Qinling by integrating structural and geochronologic analyses. The results show that the eastern West Qinling experienced two-phase deformations, separated by a period of tectonic quiescence. Large-scale south-directed displacement of thrust sheets in association with folding characterized the first-phase deformation in Late Triassic time, leading to the formation of the West Qinling fold-and-thrust belt that is composed primarily of Paleozoic-Triassic strata. This fold-and-thrust belt is in general south-convexing arc-shaped, with an accumulated south-directed displacement being over 100 km. The folding and thrusting ended up during the Norian of the Late Triassic Epoch and were immediately followed by widespread granite intrusions. Marked uplift and erosion occurred in the Early Jurassic, resulting in exhumation of the Late Triassic granites. Transpressional deformation took place in the eastern West Qinling in the Middle Jurassic on account of occurrences of strike-slip faulting and refolding. In the easternmost part of the West Qinling exists a Permian-Triassic turbidite wedge that is bordered by a right-slip fault on the northeast and by a left-slip fault on the south, indicating a westward movement that was accommodated by slip faulting. It is argued that collision of the North and South China blocks was responsible for formation of the West Qinling fold-and-thrust belt in the Late Triassic, whereas Middle Jurassic transgression is considered as the result of westward extrusion of Permian-Triassic turbiditic materials from the East Qinling owing to renewed intracontinental convergence between the North and South China blocks. A tectonic model is advanced for Early Mesozoic tectonic development of the West Qinling.

  7. Origin and demographic history of the endemic Taiwan spruce (Picea morrisonicola).

    PubMed

    Bodare, Sofia; Stocks, Michael; Yang, Jeng-Chuann; Lascoux, Martin

    2013-09-01

    Taiwan spruce (Picea morrisonicola) is a vulnerable conifer species endemic to the island of Taiwan. A warming climate and competition from subtropical tree species has limited the range of Taiwan spruce to the higher altitudes of the island. Using seeds sampled from an area in the central mountain range of Taiwan, 15 nuclear loci were sequenced in order to measure genetic variation and to assess the long-term genetic stability of the species. Genetic diversity is low and comparable to other spruce species with limited ranges such as Picea breweriana, Picea chihuahuana, and Picea schrenkiana. Importantly, analysis using approximate Bayesian computation (ABC) provides evidence for a drastic decline in the effective population size approximately 0.3-0.5 million years ago (mya). We used simulations to show that this is unlikely to be a false-positive result due to the limited sample used here. To investigate the phylogenetic origin of Taiwan spruce, additional sequencing was performed in the Chinese spruce Picea wilsonii and combined with previously published data for three other mainland China species, Picea purpurea, Picea likiangensis, and P. schrenkiana. Analysis of population structure revealed that P. morrisonicola clusters most closely with P. wilsonii, and coalescent analyses using the program MIMAR dated the split to 4-8 mya, coincidental to the formation of Taiwan. Considering the population decrease that occurred after the split, however, led to a much more recent origin.

  8. Microbial communities in the litter of middle taiga bilberry-spruce forests

    NASA Astrophysics Data System (ADS)

    Sizonenko, T. A.; Zagirova, S. V.; Khabibullina, F. M.

    2010-10-01

    The structure of the microbial communities in the litters of middle-taiga bilberry-spruce forests was studied. It was found that ammonifying and oligonitrophilic microorganisms predominate in these communities. Two maximums in the population density of the microorganisms were observed in June and August. The number of microorganisms increased in the direction from the spruce trunks to the periphery of the crowns. The species composition of the micromycetes in the litters under the spruce crowns and within the intercrown spaces differed. The maximum population density of the fungi was found in the litter under the periphery of the spruce crowns, whereas the maximum diversity of the micromycetes was observed within the intercrown spaces. The Trichoderma, Trichosporiella, Penicillium, Paecilomyces, and Chaetomium genera were most abundant in the litters of the bilberry spruce forests. The Penicillium genus had the maximum abundance during the entire growing period, and the amount of Mycelia sterilia increased in the fall. The maximum diversity of the fungi was observed in May and June.

  9. Distribution of Lepidopteran Larvae on Norway Spruce: Effects of Slope and Crown Aspect.

    PubMed

    Kulfan, Ján; Dvořáčková, Katarína; Zach, Peter; Parák, Michal; Svitok, Marek

    2016-04-01

    Lepidoptera associated with Norway spruce, Picea abies (L.) Karsten, play important roles in ecosystem processes, acting as plant pests, prey for predators, and hosts for parasites and parasitoids. Their distribution patterns in spruce crowns and forests are only poorly understood. We examined how slope and crown aspect affect the occurrence and abundance of moth larvae on solitary spruce trees in a montane region in Central Europe. Moth larvae were collected from southern and northern crowns of trees growing on south- and north-facing slopes (four treatments) using emergence boxes at the end of winter and by the beating method during the growing season. Species responses to slope and crown aspect were not uniform. Treatment effects on moth larvae were stronger in the winter than during the growing season. In winter, the abundance of bud-boring larvae was significantly higher in northern than in southern crowns regardless of the slope aspect, while both slope and aspect had marginally significant effects on abundance of miners. During the growing season, the occurrence of free-living larvae was similar among treatments. Emergence boxes and beating spruce branches are complementary techniques providing valuable insights into the assemblage structure of moth larvae on Norway spruce. Due to the uneven distribution of larvae detected in this study, we recommend adoption of a protocol that explicitly includes sampling of trees from contrasting slopes and branches from contrasting crown aspect in all seasons. PMID:26795212

  10. Origin and demographic history of the endemic Taiwan spruce (Picea morrisonicola)

    PubMed Central

    Bodare, Sofia; Stocks, Michael; Yang, Jeng-Chuann; Lascoux, Martin

    2013-01-01

    Taiwan spruce (Picea morrisonicola) is a vulnerable conifer species endemic to the island of Taiwan. A warming climate and competition from subtropical tree species has limited the range of Taiwan spruce to the higher altitudes of the island. Using seeds sampled from an area in the central mountain range of Taiwan, 15 nuclear loci were sequenced in order to measure genetic variation and to assess the long-term genetic stability of the species. Genetic diversity is low and comparable to other spruce species with limited ranges such as Picea breweriana, Picea chihuahuana, and Picea schrenkiana. Importantly, analysis using approximate Bayesian computation (ABC) provides evidence for a drastic decline in the effective population size approximately 0.3–0.5 million years ago (mya). We used simulations to show that this is unlikely to be a false-positive result due to the limited sample used here. To investigate the phylogenetic origin of Taiwan spruce, additional sequencing was performed in the Chinese spruce Picea wilsonii and combined with previously published data for three other mainland China species, Picea purpurea, Picea likiangensis, and P. schrenkiana. Analysis of population structure revealed that P. morrisonicola clusters most closely with P. wilsonii, and coalescent analyses using the program MIMAR dated the split to 4–8 mya, coincidental to the formation of Taiwan. Considering the population decrease that occurred after the split, however, led to a much more recent origin. PMID:24223271

  11. Mantle transition zone thinning beneath eastern Africa: Evidence for a whole-mantle superplume structure

    NASA Astrophysics Data System (ADS)

    Mulibo, Gabriel D.; Nyblade, Andrew A.

    2013-07-01

    to S conversions from the 410 and 660 km discontinuities observed in receiver function stacks reveal a mantle transition zone that is ~30-40 km thinner than the global average in a region ~200-400 km wide extending in a SW-NE direction from central Zambia, across Tanzania and into Kenya. The thinning of the transition zone indicates a ~190-300 K thermal anomaly in the same location where seismic tomography models suggest that the lower mantle African superplume structure connects to thermally perturbed upper mantle beneath eastern Africa. This finding provides compelling evidence for the existence of a continuous thermal structure extending from the core-mantle boundary to the surface associated with the African superplume.

  12. The structure of the Ishtar Terra central and eastern parts and some tectonic problems of Venus

    NASA Technical Reports Server (NTRS)

    Bazilevskiy, A. T.

    1986-01-01

    Moving from the Maxwell Montes to the eastern edge of Ishtar Terra there is a gradual shift in submeridional meandering folded ranges in the western section of the area to straight intersecting disjunctive systems of connected faults and sublatitudinal shifts. These disjunctive systems evidently transform older structures; the major axis of the stresses created by them is primarily oriented sublatitudinally. Relative to younger structures, in the western they occupy a higher hypsometric position. The reason for the formation of this entire system may be a large astenospheric flow wihch rise in the region of Lakashmi Planum and Maxwell Montes and which spreads and plunges in an easterly direction, taking with it deformed blocks of the lithosphere.

  13. A Contribution to the Understanding of the Regional Seismic Structure in the Eastern Mediterranean

    NASA Astrophysics Data System (ADS)

    Di Luccio, F.; Thio, H.; Pino, N.

    2001-12-01

    Regional earthquakes recorded by two digital broadband stations (BGIO and KEG) located in the Eastern Mediterranean have been analyzed in order to study the seismic structure in this region. The area consists of different tectonic provinces, which complicate the modeling of the seismic wave propagation. We have modeled the Pnl arrivals using the FK-integration technique (Saikia, 1994) along different paths at the two stations, at several distances, ranging from 400 to 1500 km. Comparing the synthetics obtained by using several models compiled by other authors, we have constructed a velocity model, considering the informations deriving from group velocity distribution, in order to determine the finer structure in the analyzed paths. The model has been perturbed by trial and error until a compressional velocity profile has been found producing the shape of the observed waveforms. The crustal thickness, upper mantle P-wave velocity and 410-km discontinuity determine the shape of the observed waveform portions.

  14. History of the spruce-fir forest in the Catskill Mountains of New York.

    PubMed

    Kudish, Michael

    2013-09-01

    Red spruce (Picea rubens Sarg.) and balsam fir (Abies balsamea (L.) Mill.) were present by 13,700 years B.C.E. in the Catskills Mountains of southeastern New York State. These conifers were, and still are, largely confined to the eastern and far western portions of the region. A gap in the distribution exists between these populations. Both species are absent from the intervening East Branch Delaware River watershed. No red spruce macrofossils were found in this watershed, suggesting that this conifer never colonized the gap postglacially. Rare macrofossils of balsam fir were found in only three of the 24 peatlands in this watershed, the conifer having disappeared between 11,300 and 8,200 years B.C.E.

  15. Influence of Mesozoic age structure on Miocene tectonic development in NE Anzoategui, Eastern Venezuela Basin

    SciTech Connect

    Sadler, P.; White, S.

    1996-08-01

    Structure within and surrounding the Quiamare-La Ceiba region, Eastern Venezuela Basin, is dominated by two major thrust fault systems. They were generated during Early-Middle Miocene time in response to oblique convergence of the Caribbean and South American plates. They are. respectively, the SE vergent NE-SW oriented Anaco fault system, and the SSE vergent ENE-WSW oriented Pirital fault system. The major structural feature associated with each fault system is a basement cored ramp anticline. New seismic data provides evidence that contributes to a better understanding of the sequence of tectonic development within and surrounding the Quiamare-La Ceiba region. Compressional structures in both the hanging wall and the footwall of the Pirital fault system appear to be inverted normal faults, that were previously active during Mesozoic time along the northern South America passive margin. A conjugate set of strike-slip faults is also present. They are oriented NNW-SSE, parallel to the Urica lineation, and SSW-NNE, respectively. A Mesozoic origin for these faults is suggested. Post-compressional relaxation during Plio-Pleistocene time resulted in the development of shallow, small scale normal faults. These normal faults appear to be localized by structural adjustments along the strike-slip fault sets. Existing oil and gas production within the Quiamare-La Ceiba region is from localized structural closures. Strike-slip faults dissect the prevailing structural grain, and may provide an additional hydrocarbon trapping mechanism.

  16. Structural and thermochronological evidence for Paleogene basement-involved shortening in the axial Eastern Cordillera, Colombia

    NASA Astrophysics Data System (ADS)

    Saylor, Joel E.; Horton, Brian K.; Stockli, Daniel F.; Mora, Andrés; Corredor, Jaime

    2012-11-01

    Although most recent studies regard the northern Andes primarily as a low-shortening inversion orogen, new mapping and thermochronology along the paired basement-involved Floresta massif and Floresta basin in the axial Eastern Cordillera of Colombia suggest major Paleogene shortening in a ramp-flat fold-thrust belt. Field mapping indicates that the hanging wall of the east-directed Soápaga fault system contains a series of upright thrust sheets with flat-on-flat cutoff relationships and a deformed footwall characterized by a complex triangle zone. These geometries necessitate roughly east-west shortening exceeding that of a previously mapped overturned hanging wall anticline and disharmonic footwall folds. Zircon (U-Th)/He (ZHe) ages indicate exhumation-induced cooling of the Soápaga hanging wall through the ˜180 °C closure temperature at 31-25 Ma. This cooling postdated documented shortening to the west and predated shortening to the east, suggesting an eastward progression of Paleogene deformation. Synorogenic Oligocene footwall strata of the Floresta basin contain distal fine-grained sediments and lack growth strata or Oligocene detrital ZHe ages, suggesting relatively high heave along the Soápaga fault system. These results are consistent with a rapidly eastward-propagating, basement-involved fold-thrust belt with ramp-flat structures that accommodated tens of km of shortening. Long-term stasis of the deformation front on the eastern and western flanks of the Eastern Cordillera due to localization of Neogene shortening along Mesozoic rift-bounding normal faults indicates a shift in deformational style in late Oligocene-early Miocene time. This geometric and temporal framework implies: 1) a total shortening in the northern Andes exceeding most current estimates; 2) Paleogene deformation in the Eastern Cordillera marked by rapid advances of the deformation front along a ramp-flat thrust system; and 3) focused Neogene reactivation (inversion) of master rift

  17. Patterns of Population Structure for Inshore Bottlenose Dolphins along the Eastern United States

    PubMed Central

    2013-01-01

    Globally distributed, the bottlenose dolphin (Tursiops truncatus) is found in a range of offshore and coastal habitats. Using 15 microsatellite loci and mtDNA control region sequences, we investigated patterns of genetic differentiation among putative populations along the eastern US shoreline (the Indian River Lagoon, Florida, and Charleston Harbor, South Carolina) (microsatellite analyses: n = 125, mtDNA analyses: n = 132). We further utilized the mtDNA to compare these populations with those from the Northwest Atlantic, Gulf of Mexico, and Caribbean. Results showed strong differentiation among inshore, alongshore, and offshore habitats (ФST = 0.744). In addition, Bayesian clustering analyses revealed the presence of 2 genetic clusters (populations) within the 250 km Indian River Lagoon. Habitat heterogeneity is likely an important force diversifying bottlenose dolphin populations through its influence on social behavior and foraging strategy. We propose that the spatial pattern of genetic variation within the lagoon reflects both its steep longitudinal transition of climate and also its historical discontinuity and recent connection as part of Intracoastal Waterway development. These findings have important management implications as they emphasize the role of habitat and the consequence of its modification in shaping bottlenose dolphin population structure and highlight the possibility of multiple management units existing in discrete inshore habitats along the entire eastern US shoreline. PMID:24129993

  18. Climate-mediated changes in zooplankton community structure for the eastern Bering Sea

    NASA Astrophysics Data System (ADS)

    Eisner, Lisa B.; Napp, Jeffrey M.; Mier, Kathryn L.; Pinchuk, Alexei I.; Andrews, Alexander G.

    2014-11-01

    Zooplankton are critical to energy transfer between higher and lower trophic levels in the eastern Bering Sea ecosystem. Previous studies from the southeastern Bering Sea shelf documented substantial differences in zooplankton taxa in the Middle and Inner Shelf Domains between warm and cold years. Our investigation expands this analysis into the northern Bering Sea and the south Outer Domain, looking at zooplankton community structure during a period of climate-mediated, large-scale change. Elevated air temperatures in the early 2000s resulted in regional warming and low sea-ice extent in the southern shelf whereas the late 2000s were characterized by cold winters, extensive spring sea ice, and a well-developed pool of cold water over the entire Middle Domain. The abundance of large zooplankton taxa such as Calanus spp. (C. marshallae and C. glacialis), and Parasagitta elegans, increased from warm to cold periods, while the abundance of gelatinous zooplankton (Cnidaria) and small taxa decreased. Biomass followed the same trends as abundance, except that the biomass of small taxa in the southeastern Bering Sea remained constant due to changes in abundance of small copepod taxa (increases in Acartia spp. and Pseudocalanus spp. and decreases in Oithona spp.). Statistically significant changes in zooplankton community structure and individual species were greatest in the Middle Domain, but were evident in all shelf domains, and in both the northern and southern portions of the eastern shelf. Changes in community structure did not occur abruptly during the transition from warm to cold, but seemed to begin gradually and build as the influence of the sea ice and cold water temperatures persisted. The change occurred one year earlier in the northern than the southern Middle Shelf. These and previous observations demonstrate that lower trophic levels within the eastern Bering Sea respond to climate-mediated changes on a variety of time scales, including those shorter than

  19. Shallow crustal structure of eastern-central Trans-Mexican Volcanic Belt.

    NASA Astrophysics Data System (ADS)

    Campos-Enriquez, J. O.; Ramón, V. M.; Lermo-Samaniego, J.

    2015-12-01

    Central-eastern Trans-Mexican Volcanic Belt (TMVB) is featured by large basins (i.e., Toluca, Mexico, Puebla-Tlaxcala, Libres-Oriental). It has been supposed that major crustal faults limit these basins. Sierra de Las Cruces range separates the Toluca and Mexico basins. The Sierra Nevada range separates Mexico basin from the Puebla-Tlaxcala basin. Based in gravity and seismic data we inferred the Toluca basin is constituted by the Ixtlahuaca sub-basin, to the north, and the Toluca sub-basin to the south, which are separated by a relative structural high. The Toluca depression is more symmetric and bounded by sub-vertical faults. In particular its eastern master fault controlled the emplacement of Sierra de Las Cruces range. Easternmost Acambay graben constitutes the northern and deepest part of the Ixtlahuaca depression. The Toluca-Ixtlahuaca basin is inside the Taxco-San Miguel de Allende fault system, and limited to the west by the Guerrero terrane which continues beneath the TMVB up to the Acambay graben. Mexico basin basement occupies an intermediate position and featured by a relative structural high to the north-east, as established by previous studies. This relative structural high is limited to the west by the north-south Mixhuca trough, while to the south it is bounded by the east-west Copilco-Xochimilco-Chalco sub-basin. The Puebla-Tlaxcala basin basement is the shallowest of these 3 tectonic depressions. In general, features (i.e., depth) and relationship between these basins, from west to east, are controlled by the regional behavior of the Sierra Madre Oriental fold and thrust belt basement (i.e., Oaxaca Complex?). This study indicates that an active east-west regional fault system limits to the south the TMVB (from the Nevado de Toluca volcano through the Popocatepetl volcano and eastward along southern Puebla-Tlaxcala basin). The Tenango and La Pera fault systems constituting the western part of this regional fault system coincide with northern

  20. Lenticular stretch structures in eastern Nevada - possible trapping mechanism in supposed graben

    SciTech Connect

    Walker, C.T.; Dennis, J.G.; Lumsden, W.W.

    1986-04-01

    Eastern Nevada is widely recognized as a region of tectonic extension. The dominant structures are generally agreed to be low-dipping, younger over older faults and steeper listric faults that are responsible for the basins (grabens) and ranges (horsts). In the Schell Creek-Duck Creek Range, east of Ely, and in the White Pine Range, southwest of Ely, small lenticular structures bounded by tectonic discontinuities can be clearly seen in the field. These lenticular units, or stretch structures, range in length from a few meters to more than 200 m. All lenticular stretch structures that can be clearly seen in the field are stratigraphically restricted; the stretched formations are the Eureka Quartzite, the Pilot Shale, the Joana Limestone, and the Chainman Shale. Still larger stretch structures, which may include several formations, are inferred, and the authors suggest that extension has created lenticular structures at all scales. The Duck Creek and Schell Creek Ranges east of Ely consist mostly of Devonian and older rocks. They are separated by a topographically lower area containing mostly Mississippian and Pennsylvanian rocks. This structure, which separates the ranges, has been referred to as a graben, but field evidence suggests that it is a large-scale lenticular stretch structure. Unlike a true graben, the structure does not extend downward. For example, in several places within the supposed graben, Cambrian and Ordovician rocks project through a cover of Carboniferous Chainman Shale and Ely Limestone, suggesting the Chainman-Ely is a thin sheet underlain by Cambrian-Ordovician rocks. Accordingly, they suggest that extension in the Duck Creek-Schell Creek Ranges stretched the formations into lenticular bodies. Between the Duck Creek and Schell Creek Ranges, the Cambrian-Ordovician is attenuated, and the resulting tectonic depression is occupied by a lenticular mass of Carboniferous rocks.

  1. Four New Sulfated Polar Steroids from the Far Eastern Starfish Leptasterias ochotensis: Structures and Activities

    PubMed Central

    Malyarenko, Timofey V.; Malyarenko (Vishchuk), Olesya S.; Ivanchina, Natalia V.; Kalinovsky, Anatoly I.; Popov, Roman S.; Kicha, Alla A.

    2015-01-01

    Three new sulfated steroid monoglycosides, leptaochotensosides A–C (1–3), and a new sulfated polyhydroxylated steroid (4) were isolated from the alcoholic extract of the Far Eastern starfish Leptasterias ochotensis. The structures of compounds 1–4 were established by extensive nuclear magnetic resonance (NMR) and electrospray ionization mass spectrometry (ESIMS) analyses and chemical transformations. Although the isolated compounds did not show any apparent cytotoxicity against melanoma RPMI-7951 and breast cancer T-47D cell lines, leptaochotensoside A (1) demonstrated inhibition of T-47D cell colony formation in a soft agar clonogenic assay at nontoxic doses. In addition, this compound decreased the epidermal growth factor (EGF)-induced colony formation of mouse epidermal JB6 Cl41 cells. The cancer preventive action of 1 is realized through regulation of mitogen-activated protein kinase (MAPK) signaling pathway. PMID:26193286

  2. Structure of the lithosphere beneath the Eastern Alps (southern sector of the TRANSALP transect)

    NASA Astrophysics Data System (ADS)

    Castellarin, Alberto; Nicolich, Rinaldo; Fantoni, Roberto; Cantelli, Luigi; Sella, Mattia; Selli, Luigi

    2006-02-01

    supported by any significant uplifting of the Dolomites. The total average uplift of the Dolomites during the Neogene appears to be 6-7 times smaller than that recognized in the TW. Markedly the northward dip of the PL, reaching a depth of approximately 20 km, is proposed in our interpretation; finally, the Adriatic upper crustal evolution points to the late post-collision change in the tectonic grow-up of the Eastern Alps orogenic chain. The tectonic accretion of the northern frontal zone of the Eastern and Central Alps was interrupted from the Late Miocene (Serravallian-Tortonian) onward, as documented by the Molasse basin evolution. On the contrary, the structural nucleation along the S-vergent tectonic belt of the eastern Southern Alps ( Montello-Friuli thrust belt) severely continued during the Messinian and the Plio-Pleistocene. This structural evolution can be considered to be consistent with the deep under-thrusting and wedge indentation of the Adriatic lithosphere underneath the southern side of the Eastern Alps thrust-and-fold belt. Similarly, the significance of the magmatic activity for the construction of the Southern Alps crust and for its mechanical and geological differentiation, which qualified the evolution of the thrust-and-fold belt, is highlighted, starting with the Permian-Triassic magmatism and progressing with the Paleogene occurrences along the Periadriatic Lineament and in the Venetian Magmatic Province (Lessini-Euganei Hills).

  3. Forest dynamics after successive spruce budworm outbreaks in mixedwood forests.

    PubMed

    Bouchard, Mathieu; Kneeshaw, Daniel; Bergeron, Yves

    2006-09-01

    In order to assess the long-term spatiotemporal influence of the spruce budworm in sub-boreal mixedwood forests, we studied the effect of three successive outbreaks in a region of western Quebec, Canada. We used dendrochronology to detect past outbreaks in three areas (111-185 ha), based on the recruitment age of balsam fir (Abies balsamea) and on growth patterns of white spruce (Picea glauca), the two main host species of this defoliating insect. We also used a series of aerial photographs taken between 1935 and 2003 to evaluate overstory mortality and post-outbreak succession patterns in these same areas. Individual outbreaks had a spatially homogenous impact on host species throughout the region, but successive outbreaks differed in intensity: the two outbreaks around 1910 and 1980 caused widespread mortality in the overstory, but an outbreak around 1945 had little impact, probably because the forest mosaic had not yet recuperated from the 1910 outbreak. No clear outbreak was detected in the later part of the 19th century. In portions of the study areas where the 1910 outbreak had a major impact, between 36% and 50% of the stands were reoccupied by balsam fir stands in the period up to the 1980 outbreak (cyclic succession), the rest being at least partly replaced by nonhost species such as Betula spp. Changes in forest composition after the 1910 outbreak were mostly associated with upper-slope positions in all study areas. The 1980 outbreak also had a higher impact than earlier outbreaks in lower-slope positions dominated by black spruce (Picea mariana)-balsam fir mixtures. These results suggest that, at the regional scale, the abundance of mature or over-mature balsam fir stands does not determine the outbreak cycle. When an outbreak occurs, however, its impact will be strongly constrained by forest characteristics such as stand composition and structure, which are themselves influenced by previous disturbances and slope position.

  4. Forest dynamics after successive spruce budworm outbreaks in mixedwood forests.

    PubMed

    Bouchard, Mathieu; Kneeshaw, Daniel; Bergeron, Yves

    2006-09-01

    In order to assess the long-term spatiotemporal influence of the spruce budworm in sub-boreal mixedwood forests, we studied the effect of three successive outbreaks in a region of western Quebec, Canada. We used dendrochronology to detect past outbreaks in three areas (111-185 ha), based on the recruitment age of balsam fir (Abies balsamea) and on growth patterns of white spruce (Picea glauca), the two main host species of this defoliating insect. We also used a series of aerial photographs taken between 1935 and 2003 to evaluate overstory mortality and post-outbreak succession patterns in these same areas. Individual outbreaks had a spatially homogenous impact on host species throughout the region, but successive outbreaks differed in intensity: the two outbreaks around 1910 and 1980 caused widespread mortality in the overstory, but an outbreak around 1945 had little impact, probably because the forest mosaic had not yet recuperated from the 1910 outbreak. No clear outbreak was detected in the later part of the 19th century. In portions of the study areas where the 1910 outbreak had a major impact, between 36% and 50% of the stands were reoccupied by balsam fir stands in the period up to the 1980 outbreak (cyclic succession), the rest being at least partly replaced by nonhost species such as Betula spp. Changes in forest composition after the 1910 outbreak were mostly associated with upper-slope positions in all study areas. The 1980 outbreak also had a higher impact than earlier outbreaks in lower-slope positions dominated by black spruce (Picea mariana)-balsam fir mixtures. These results suggest that, at the regional scale, the abundance of mature or over-mature balsam fir stands does not determine the outbreak cycle. When an outbreak occurs, however, its impact will be strongly constrained by forest characteristics such as stand composition and structure, which are themselves influenced by previous disturbances and slope position. PMID:16995632

  5. Unexpectedly low rangewide population genetic structure of the imperiled eastern box turtle Terrapene c. carolina.

    PubMed

    Kimble, Steven J A; Rhodes, O E; Williams, Rod N

    2014-01-01

    Rangewide studies of genetic parameters can elucidate patterns and processes that operate only over large geographic scales. Herein, we present a rangewide population genetic assessment of the eastern box turtle Terrapene c. carolina, a species that is in steep decline across its range. To inform conservation planning for this species, we address the hypothesis that disruptions to demographic and movement parameters associated with the decline of the eastern box turtle has resulted in distinctive genetic signatures in the form of low genetic diversity, high population structuring, and decreased gene flow. We used microsatellite genotype data from (n = 799) individuals from across the species range to perform two Bayesian population assignment approaches, two methods for comparing historical and contemporary migration among populations, an evaluation of isolation by distance, and a method for detecting barriers to gene flow. Both Bayesian methods of population assignment indicated that there are two populations rangewide, both of which have maintained high levels of genetic diversity (HO = 0.756). Evidence of isolation by distance was detected in this species at a spatial scale of 300-500 km, and the Appalachian Mountains were identified as the primary barrier to gene flow across the species range. We also found evidence for historical but not contemporary migration between populations. Our prediction of many, highly structured populations across the range was not supported. This may point to cryptic contemporary gene flow, which might in turn be explained by the presence of rare transients in populations. However these data may be influenced by historical signatures of genetic connectivity because individuals of this species can be long-lived. PMID:24647580

  6. The Electrical Resistivity Structure of the Eastern Anatolian Collision Zone, Northeastern Anatolia

    NASA Astrophysics Data System (ADS)

    Cengiz, Özlem; Tuǧrul Başokur, Ahmet; Tolak Çiftçi, Elif

    2016-04-01

    The Northeastern Anatolia is located at the intensely deformed Eastern Anatolian Collision Zone (EACZ), and its tectonic framework is characterized by the collision of the Arabian plate with Eurasian. Although extensive attention is given to understand the crustal and upper mantle processes at this convergent boundary, there is still an ongoing debate over the geodynamic processes of the region. In this study, we were specifically interested in the geoelectric properties and thus geodynamics of the crust beneath the EACZ. Magnetotelluric (MT) measurements were made on two profiles across the north of the EACZ in 1998 as part of a national project undertaken by the Turkish Petroleum Corporation (TPAO). MT data in the frequency range of 300-0.001 Hz were collected from 168 stations located along 78 km north to south and 47 km west to east profiles where direct convergence occurs between Arabian and Eurasian plates. Two and three-dimensional inversion algorithms were used to obtain resistivity models of the study area. According to these models, the upper crust consists of low resistivity sedimentary rocks (<30 Ωm) that are underlain by highly resistive (~500-1000 Ωm) crystalline basement rocks of the Eastern Anatolian Accretionary Complex and Pontides. While the upper and lower crustal resistivity at the northern part of the study area shows a layered structure, significant horizontal and vertical variations for the rest of the EACZ exists on resistivity models. The broad low resistivity zones (<50 Ωm) observed at mid and lower crustal levels throughout the EACZ. These fluid-rich regions along with high temperatures could indicate weak zones representing the locations of active deformation induced by continent-continent collision and correlate with volcanic centers in the region. The variation in the resistivity structure supports the southward subduction model with the resistive continental block and the deep conductive zones presumably corresponding to the

  7. Uppermost mantle P wavespeed structure beneath eastern China and its surroundings

    NASA Astrophysics Data System (ADS)

    Sun, Weijia; Kennett, B. L. N.

    2016-06-01

    Pn travel-time tomography provides a way of improving structural information on the uppermost mantle across eastern China exploiting recent developments of dense seismic networks with well recorded seismic events. We used waveforms from 2009 at Chinese stations, supplemented by bulletin arrival times. An initial P wave model was constructed using the crustal model from CRUST1.0 coupled to a P wave model in the mantle derived from the SL2013sv model to capture the broad-scale features. This starting model enables us to compensate for the large contrasts in crustal thickness across the region. All events were relocated using the initial 3-D P model, and after relocation, consistent patterns of travel-time residuals are obtained. We extract Pn as the first arrival in the distance range 1.8 ∘ to 12 ∘. We use the FMTOMO (Fast Marching TOMOgraphy) approach to invert the travel-time results to generate a P wavespeed structure with a resolution of 2 ∘× 2 ∘ down to 75 km. There are considerable variations in Pn wavespeed in the uppermost mantle across the region. The central portion of the North China craton is imaged with particularly slow P wavespeeds, whilst most of the neighbouring Ordos block is fast. Fast P wavespeeds extend through much of the uppermost mantle beneath eastern Central Asia Orogen, northeast China and beneath the Korean peninsula. In the south, the Sichuan Block and the western Yangtze craton show rather fast P wavespeeds. The Tanlu fault system appears to cut through the crust into the mantle with marked slow P wavespeed at its southern end.

  8. Structural and petrologic evolution of the Lihue basin and eastern Kauai, Hawaii

    USGS Publications Warehouse

    Reiners, P.W.; Nelson, B.K.; Izuka, S.K.

    1999-01-01

    The topography of the eastern part of the Hawaiian island of Kauai is dominated by the Lihue basin, a large (???110 km2) semicircular depression bounded by steep cliffs and partly filled by late rejuvenated-stage (or post-erosional stage) volcanic material. As with other large, semicircular basins on ocean-island volcanoes, the subsurface geology and origin (e.g., structural collapse vs. fluvial erosion) of the Lihue basin are poorly understood. New analyses of samples collected from eastern Kauai and drill holes within the basin document several important features of the late-stage geologic evolution of Kauai. First, thick (>300 m) sequences of rejuvenated-stage Koloa Volcanics in the Lihue basin show systematic, basin-wide geochemical trends of increasingly incompatible elements with time, indicating a gradual decrease in the extent of partial melting of mantle sources with time. Second, beneath the rejuvenated-stage volcanics in the basin, a thin layer of postshield alkalic stage lavas (e.g., hawaiites and mugearites) overlies older shield-stage tholeiitic lavas of the Napali Member, indicating that the Lihue basin formed by structural collapse, not fluvial erosion. Third, a large (???2-5 km3) matrix-supported breccia, interpreted as deposits of one or more debris flows, is within the rejuvenated-stage volcanics throughout the basin, and correlates with surficial exposures of the Palikea Breccia west of the basin. Isotopic compositions of the bulk breccia are similar to those of tholeiites from the east side of Kauai, and distinct from those of west Kauai tholeiites. Clasts within the breccia are dominantly hawaiite and alkali gabbro. The source region of the breccia in the steep cliffs and highlands of the central massif to the west of the basin must contain magmatic products of an extensive postshield alkalic stage, including hawaiite flows and one or more large intrusive bodies or ponded sequences of alkali gabbro.

  9. Unexpectedly Low Rangewide Population Genetic Structure of the Imperiled Eastern Box Turtle Terrapene c. carolina

    PubMed Central

    Kimble, Steven J. A.; Rhodes Jr., O. E.; Williams, Rod N.

    2014-01-01

    Rangewide studies of genetic parameters can elucidate patterns and processes that operate only over large geographic scales. Herein, we present a rangewide population genetic assessment of the eastern box turtle Terrapene c. carolina, a species that is in steep decline across its range. To inform conservation planning for this species, we address the hypothesis that disruptions to demographic and movement parameters associated with the decline of the eastern box turtle has resulted in distinctive genetic signatures in the form of low genetic diversity, high population structuring, and decreased gene flow. We used microsatellite genotype data from (n = 799) individuals from across the species range to perform two Bayesian population assignment approaches, two methods for comparing historical and contemporary migration among populations, an evaluation of isolation by distance, and a method for detecting barriers to gene flow. Both Bayesian methods of population assignment indicated that there are two populations rangewide, both of which have maintained high levels of genetic diversity (HO = 0.756). Evidence of isolation by distance was detected in this species at a spatial scale of 300 – 500 km, and the Appalachian Mountains were identified as the primary barrier to gene flow across the species range. We also found evidence for historical but not contemporary migration between populations. Our prediction of many, highly structured populations across the range was not supported. This may point to cryptic contemporary gene flow, which might in turn be explained by the presence of rare transients in populations. However these data may be influenced by historical signatures of genetic connectivity because individuals of this species can be long-lived. PMID:24647580

  10. Negative Feedbacks on Bark Beetle Outbreaks: Widespread and Severe Spruce Beetle Infestation Restricts Subsequent Infestation

    PubMed Central

    Hart, Sarah J.; Veblen, Thomas T.; Mietkiewicz, Nathan; Kulakowski, Dominik

    2015-01-01

    Understanding disturbance interactions and their ecological consequences remains a major challenge for research on the response of forests to a changing climate. When, where, and how one disturbance may alter the severity, extent, or occurrence probability of a subsequent disturbance is encapsulated by the concept of linked disturbances. Here, we evaluated 1) how climate and forest habitat variables, including disturbance history, interact to drive 2000s spruce beetle (Dendroctonus rufipennis) infestation of Engelmann spruce (Picea engelmannii) across the Southern Rocky Mountains; and 2) how previous spruce beetle infestation affects subsequent infestation across the Flat Tops Wilderness in northwestern Colorado, which experienced a severe landscape-scale spruce beetle infestation in the 1940s. We hypothesized that drought and warm temperatures would promote infestation, whereas small diameter and non-host trees, which may reflect past disturbance by spruce beetles, would inhibit infestation. Across the Southern Rocky Mountains, we found that climate and forest structure interacted to drive the 2000s infestation. Within the Flat Tops study area we found that stands infested in the 1940s were composed of higher proportions of small diameter and non-host trees ca. 60 years later. In this area, the 2000s infestation was constrained by a paucity of large diameter host trees (> 23 cm at diameter breast height), not climate. This suggests that there has not been sufficient time for trees to grow large enough to become susceptible to infestation. Concordantly, we found no overlap between areas affected by the 1940s infestation and the current infestation. These results show a severe spruce beetle infestation, which results in the depletion of susceptible hosts, can create a landscape template reducing the potential for future infestations. PMID:26000906

  11. Negative feedbacks on bark beetle outbreaks: widespread and severe spruce beetle infestation restricts subsequent infestation.

    PubMed

    Hart, Sarah J; Veblen, Thomas T; Mietkiewicz, Nathan; Kulakowski, Dominik

    2015-01-01

    Understanding disturbance interactions and their ecological consequences remains a major challenge for research on the response of forests to a changing climate. When, where, and how one disturbance may alter the severity, extent, or occurrence probability of a subsequent disturbance is encapsulated by the concept of linked disturbances. Here, we evaluated 1) how climate and forest habitat variables, including disturbance history, interact to drive 2000s spruce beetle (Dendroctonus rufipennis) infestation of Engelmann spruce (Picea engelmannii) across the Southern Rocky Mountains; and 2) how previous spruce beetle infestation affects subsequent infestation across the Flat Tops Wilderness in northwestern Colorado, which experienced a severe landscape-scale spruce beetle infestation in the 1940s. We hypothesized that drought and warm temperatures would promote infestation, whereas small diameter and non-host trees, which may reflect past disturbance by spruce beetles, would inhibit infestation. Across the Southern Rocky Mountains, we found that climate and forest structure interacted to drive the 2000s infestation. Within the Flat Tops study area we found that stands infested in the 1940s were composed of higher proportions of small diameter and non-host trees ca. 60 years later. In this area, the 2000s infestation was constrained by a paucity of large diameter host trees (> 23 cm at diameter breast height), not climate. This suggests that there has not been sufficient time for trees to grow large enough to become susceptible to infestation. Concordantly, we found no overlap between areas affected by the 1940s infestation and the current infestation. These results show a severe spruce beetle infestation, which results in the depletion of susceptible hosts, can create a landscape template reducing the potential for future infestations.

  12. Negative feedbacks on bark beetle outbreaks: widespread and severe spruce beetle infestation restricts subsequent infestation.

    PubMed

    Hart, Sarah J; Veblen, Thomas T; Mietkiewicz, Nathan; Kulakowski, Dominik

    2015-01-01

    Understanding disturbance interactions and their ecological consequences remains a major challenge for research on the response of forests to a changing climate. When, where, and how one disturbance may alter the severity, extent, or occurrence probability of a subsequent disturbance is encapsulated by the concept of linked disturbances. Here, we evaluated 1) how climate and forest habitat variables, including disturbance history, interact to drive 2000s spruce beetle (Dendroctonus rufipennis) infestation of Engelmann spruce (Picea engelmannii) across the Southern Rocky Mountains; and 2) how previous spruce beetle infestation affects subsequent infestation across the Flat Tops Wilderness in northwestern Colorado, which experienced a severe landscape-scale spruce beetle infestation in the 1940s. We hypothesized that drought and warm temperatures would promote infestation, whereas small diameter and non-host trees, which may reflect past disturbance by spruce beetles, would inhibit infestation. Across the Southern Rocky Mountains, we found that climate and forest structure interacted to drive the 2000s infestation. Within the Flat Tops study area we found that stands infested in the 1940s were composed of higher proportions of small diameter and non-host trees ca. 60 years later. In this area, the 2000s infestation was constrained by a paucity of large diameter host trees (> 23 cm at diameter breast height), not climate. This suggests that there has not been sufficient time for trees to grow large enough to become susceptible to infestation. Concordantly, we found no overlap between areas affected by the 1940s infestation and the current infestation. These results show a severe spruce beetle infestation, which results in the depletion of susceptible hosts, can create a landscape template reducing the potential for future infestations. PMID:26000906

  13. Canopy gap dynamics of second-growth red spruce-northern hardwood stands in West Virginia

    USGS Publications Warehouse

    Rentch, J.S.; Schuler, T.M.; Nowacki, G.J.; Beane, N.R.; Ford, W.M.

    2010-01-01

    Forest restoration requires an understanding of the natural disturbance regime of the target community and estimates of the historic range of variability of ecosystem components (composition, structure, and disturbance processes). Management prescriptions that support specific restoration activities should be consistent with these parameters. In this study, we describe gap-phase dynamics of even-aged, second-growth red spruce-northern hardwood stands in West Virginia that have been significantly degraded following early Twentieth Century harvesting and wildfire. In the current stage of stand development, gaps tended to be small, with mean canopy gap and extended canopy gap sizes of 53.4m2 and 199.3m2, respectively, and a canopy turnover rate of 1.4%year-1. The majority of gaps resulted from the death of one or two trees. American beech snags were the most frequent gap maker, partially due to the elevated presence of beech-bark disease in the study area. Gaps ranged in age from 1 to 28 years, had a mean of 13 years, and were unimodal in distribution. We projected red spruce to be the eventual gap filler in approximately 40% of the gaps. However, we estimated that most average-sized gaps will close within 15-20 years before red spruce canopy ascension is projected (30-60 years). Accordingly, many understory red spruce will require more than one overhead release - an observation verified by the tree-ring record and consistent with red spruce life history characteristics. Based on our observations, silvicultural prescriptions that include overhead release treatments such as thinning from above or small gap creation through selection harvesting could be an appropriate activity to foster red spruce restoration in the central Appalachians. ?? 2010 Elsevier B.V.

  14. Extension-related structural traps in fault basins of eastern Nevada

    SciTech Connect

    Walker, C.T.; Dennis, J.G.; Lumsden, W.W.

    1989-04-01

    Low-angle younger-over-older faults have been widely reported in eastern Nevada, although no general agreement exists on their origin. Three preferred models are (1) local gravity sliding, (2) mid-crustal ductile extension, and (3) a master detachment possibly extending into the mantle. None are fully supported by field evidence. Some low-angle faults in the White Pine, Duck Creek and Schell Creek Ranges involve ductile extension along incompetent sedimentary units and brittle extension of intercalated competent units to form lenticular stretch structures. Upper crustal extension may be an indirect response to midcrustal ductile extension that occurred during a Tertiary heating event. Between the Duck Creek and Schell Creek Ranges, extension attenuated the pre-carboniferous section, creating a depression in which Carboniferous rocks were preserved. Since this structure, termed a pseudograben, can be traced northward into a typical graben, basins may be initiated by attenuation, with rifting occurring later. In such basins, subsurface detachments should be expected. Some low-angle faults resemble megalandslides because fragmented competent units slid into depressions created by attenuation. With retreat of the geotherms, ductile extension was confined to deep crustal levels and rifting replaced low-angle faulting at higher levels. Potential extension traps in basins are large-scale stretch structures, porous and permeable units truncated and sealed by detachment, and tilted blocks cut by steep faults. Best prospects are likely to occur in basins subjected only to Tertiary heating because oil generated at that time would be tapped in developing extension structures.

  15. Stable isotopes provide insight into population structure and segregation in eastern North Atlantic sperm whales.

    PubMed

    Borrell, Asunción; Velásquez Vacca, Adriana; Pinela, Ana M; Kinze, Carl; Lockyer, Christina H; Vighi, Morgana; Aguilar, Alex

    2013-01-01

    In pelagic species inhabiting large oceans, genetic differentiation tends to be mild and populations devoid of structure. However, large cetaceans have provided many examples of structuring. Here we investigate whether the sperm whale, a pelagic species with large population sizes and reputedly highly mobile, shows indication of structuring in the eastern North Atlantic, an ocean basin in which a single population is believed to occur. To do so, we examined stable isotope values in sequential growth layer groups of teeth from individuals sampled in Denmark and NW Spain. In each layer we measured oxygen- isotope ratios (δ(18)O) in the inorganic component (hydroxyapatite), and nitrogen and carbon isotope ratios (δ(15)N: δ(13)C) in the organic component (primarily collagenous). We found significant differences between Denmark and NW Spain in δ(15)N and δ(18)O values in the layer deposited at age 3, considered to be the one best representing the baseline of the breeding ground, in δ(15)N, δ(13)C and δ(18)O values in the period up to age 20, and in the ontogenetic variation of δ(15)N and δ(18)O values. These differences evidence that diet composition, use of habitat and/or migratory destinations are dissimilar between whales from the two regions and suggest that the North Atlantic population of sperm whales is more structured than traditionally accepted.

  16. Stable Isotopes Provide Insight into Population Structure and Segregation in Eastern North Atlantic Sperm Whales

    PubMed Central

    Borrell, Asunción; Velásquez Vacca, Adriana; Pinela, Ana M.; Kinze, Carl; Lockyer, Christina H.; Vighi, Morgana; Aguilar, Alex

    2013-01-01

    In pelagic species inhabiting large oceans, genetic differentiation tends to be mild and populations devoid of structure. However, large cetaceans have provided many examples of structuring. Here we investigate whether the sperm whale, a pelagic species with large population sizes and reputedly highly mobile, shows indication of structuring in the eastern North Atlantic, an ocean basin in which a single population is believed to occur. To do so, we examined stable isotope values in sequential growth layer groups of teeth from individuals sampled in Denmark and NW Spain. In each layer we measured oxygen- isotope ratios (δ18O) in the inorganic component (hydroxyapatite), and nitrogen and carbon isotope ratios (δ15N: δ13C) in the organic component (primarily collagenous). We found significant differences between Denmark and NW Spain in δ15N and δ18O values in the layer deposited at age 3, considered to be the one best representing the baseline of the breeding ground, in δ15N, δ13C and δ18O values in the period up to age 20, and in the ontogenetic variation of δ15N and δ18O values. These differences evidence that diet composition, use of habitat and/or migratory destinations are dissimilar between whales from the two regions and suggest that the North Atlantic population of sperm whales is more structured than traditionally accepted. PMID:24324782

  17. Crustal structure in eastern Kazakh, U. S. S. R. from teleseismic receiver functions

    SciTech Connect

    Priestley, K.F.; Zandt, G.; Randall, G.E.

    1988-06-01

    Broadband receiver functions determined from teleseismic P-waveforms recorded at two seismic stations in eastern Kazakh, U.S.S.R., are inverted for the vertical velocity structure beneath the stations. The detailed broadband receiver functions are obtained by stacking source-equalized radial components of teleseismic P-waveforms. A time-domain inversion of the radial receiver function is used to determine the structure assuming a crustal model parameterized by flat-lying, homogeneous layers. The general features of the inversion results are: a complex shallow crust, velocities less than 6 km/s in the upper curst, a high velocity (6.9--7.5 km/s) lower crust, and a Moho that varies between 47 and 57 km depth. These results compare favorably with a composite velocity model from Deep Seismic Sounding data, but show lower velocities in the upper curst, and higher velocities in the lower crust. The results indicate that the crust in this region is relatively uniform, however the nature and depth of the Moho changes significantly across the region. The receiver function inversion structure for the Kazakh sites is similar to a published receiver function structure for a site on the Canadian Shield. copyright American Geophysical Union 1988

  18. Structural studies of TiO2/wood coatings prepared by hydrothermal deposition of rutile particles from TiCl4 aqueous solutions on spruce (Picea Abies) wood

    NASA Astrophysics Data System (ADS)

    Pori, Pavel; Vilčnik, Aljaž; Petrič, Marko; Sever Škapin, Andrijana; Mihelčič, Mohor; Šurca Vuk, Angela; Novak, Urban; Orel, Boris

    2016-05-01

    A low temperature approach was developed for the deposition of rutile TiO2 particles on a wood surface by hydrolysis of TiCl4 in aqueous solutions acidified with HCl, and crystallization at 75 and 90 °C (1 h). Prior to hydrothermal treatment, Picea Abies wood was first soaked in a 0.5 mmol/l aqueous solution containing anionic surfactant sodium dodecyl sulphate (SDS, Sigma Aldrich) for 2 h at 80 °C. The crystal structure of the hydrothermally made rutile particles was determined with XRD, while the morphology of the deposited TiO2 particles and their distribution in the wood were examined with SEM and EDX measurements. The penetration and amount of deposited rutile particles could be modified by changing the deposition conditions. Thicker layers were obtained from more concentrated aqueous TiCl4 solutions with and without added HCl, and with longer deposition times and higher temperatures of the hydrothermal treatment. The interaction of TiO2 particles with hemicellulose and lignin in wood was established from infrared attenuated total reflection (FT-IR ATR) and Raman spectra measurements, from which the spectra of wood were subtracted. Analysis of the subtraction spectra showed the presence of titania particles on the wood surface, revealing also the establishment of TiO2-wood coordinative bonds of titanium ions with hemicellulose and lignin. The red frequency shift of the OH stretching modes suggested interaction of the TiO2 particles with water molecules of wood. TiO2 deposited on wood treated with SDS became hydrophobic (water contact angles (WCA) of 150°), contrasting the properties of untreated wood with a deposited TiO2 particle coating, which remained hydrophilic.

  19. Intra-annual variability of anatomical structure and δ13C values within tree rings of spruce and pine in alpine, temperate and boreal Europe

    PubMed Central

    Vaganov, Eugene A.; Skomarkova, Marina V.; Knohl, Alexander; Brand, Willi A.; Roscher, Christiane

    2009-01-01

    Tree-ring width, wood density, anatomical structure and 13C/12C ratios expressed as δ13C-values of whole wood of Picea abies were investigated for trees growing in closed canopy forest stands. Samples were collected from the alpine Renon site in North Italy, the lowland Hainich site in Central Germany and the boreal Flakaliden site in North Sweden. In addition, Pinus cembra was studied at the alpine site and Pinus sylvestris at the boreal site. The density profiles of tree rings were measured using the DENDRO-2003 densitometer, δ13C was measured using high-resolution laser-ablation-combustion-gas chromatography-infra-red mass spectrometry and anatomical characteristics of tree rings (tracheid diameter, cell-wall thickness, cell-wall area and cell-lumen area) were measured using an image analyzer. Based on long-term statistics, climatic variables, such as temperature, precipitation, solar radiation and vapor pressure deficit, explained <20% of the variation in tree-ring width and wood density over consecutive years, while 29–58% of the variation in tree-ring width were explained by autocorrelation between tree rings. An intensive study of tree rings between 1999 and 2003 revealed that tree ring width and δ13C-values of whole wood were significantly correlated with length of the growing season, net radiation and vapor pressure deficit. The δ13C-values were not correlated with precipitation or temperature. A highly significant correlation was also found between δ13C of the early wood of one year and the late wood of the previous year, indicating a carry-over effect of the growing conditions of the previous season on current wood production. This latter effect may explain the high autocorrelation of long-term tree-ring statistics. The pattern, however, was complex, showing stepwise decreases as well as stepwise increases in the δ13C between late wood and early wood. The results are interpreted in the context of the biochemistry of wood formation and its linkage

  20. Crustal structure of the Central-Eastern Greenland: results from the TopoGreenland refraction profile

    NASA Astrophysics Data System (ADS)

    Shulgin, Alexey; Thybo, Hans

    2014-05-01

    Until present, seismic surveys have only been carried out offshore and near the coasts of Greenland, where the crustal structure is affected by oceanic break-up. We present the deep seismic structure of the crust of the interior of Greenland, based on the new and the only existing so far seismic refraction/wide-angle reflection profile. The seismic data was acquired by a team of six people during a two-month long experiment in summer of 2011 on the ice cap in the interior of central-eastern Greenland. The presence of an up to 3.4 km thick ice sheet, permanently covering most of the land mass, made acquisition of geophysical data logistically complicated. The profile extends 310 km inland in E-W direction from the approximate edge of the stable ice cap near the Scoresby Sund across the center of the ice cap. 350 Reftek Texan receivers recorded high-quality seismic data from 8 equidistant shots along the profile. Explosive charge sizes were 1 ton at the ends and ca. 500 kg along the profile, loaded with about 125 kg at 35-85 m depth in individual boreholes. Given that the data acquisition was affected by the thick ice sheet, we questioned the quality of seismic records in such experiment setup. We have developed an automatic routine to check the amplitudes and spectra of the selected seismic phases and to check the differences/challenges in making seismic experiments on ice and the effects of ice on data interpretation. Using tomographic inversion and forward ray tracing modelling we have obtained the two-dimensional velocity model down to a 50 km depth. The model shows a decrease of crustal thickness from 47 km below the centre of Greenland in the western part of the profile to 40 km in its eastern part. Relatively high lower crustal velocities (Vp 6.8 - 7.3 km/s) in the western part of the TopoGreenland profile may result from past collision tectonics or, alternatively, may be related to the speculated passage of the Iceland mantle plume. Comparison of our results

  1. Geologic framework, structure, and hydrogeologic characteristics of the Knippa Gap area in eastern Uvalde and western Medina Counties, Texas

    USGS Publications Warehouse

    Clark, Allan K.; Pedraza, Diana E.; Morris, Robert R.

    2013-01-01

    By using data that were compiled and collected for this study and previous studies, a revised map was constructed depicting the geologic framework, structure, and hydrogeologic characteristics of the Knippa Gap area in eastern Uvalde and western Medina Counties, Tex. The map also shows the interpreted structural dip directions and interpreted location of a structural low (trough) in the area known as the Knippa Gap.

  2. Carbon sequestration by young Norway spruce monoculture

    NASA Astrophysics Data System (ADS)

    Pokorny, R.; Rajsnerova, P.; Kubásek, J.

    2012-04-01

    Many studies have been focused on allometry, wood-mass inventory, carbon (C) sequestration, and biomass expansion factors as the first step for the evaluation of C sinks of different plant ecosystems. To identify and quantify these terrestrial C sinks, and evaluate CO2 human-induced emissions on the other hand, information for C balance accounting (for impletion of commitment to Kyoto protocol) are currently highly needed. Temperate forest ecosystems have recently been identified as important C sink. Carbon sink might be associated with environmental changes (elevated [CO2], air temperature, N deposition etc.) and large areas of managed fast-growing young forests. Norway spruce (Pice abies L. Karst) is the dominant tree species (35%) in Central European forests. It covers 55 % of the total forested area in the Czech Republic, mostly at high altitudes. In this contribution we present C sequestration by young (30-35 year-old) Norway spruce monocultures in highland (650-700 m a.s.l., AT- mean annual temperature: 6.9 ° C; P- annual amount of precipitation: 700 mm; GL- growing season duration: 150 days) and mountain (850-900 m a.s.l.; AT of 5.5 ° C; P of 1300 mm; and GL of 120 days) areas and an effect of a different type of thinning. However, the similar stem diameter at the breast height and biomass proportions among above-ground tree organs were obtained in the both localities; the trees highly differ in their height, above-ground organ's biomass values and total above ground biomass, particularly in stem. On the total mean tree biomass needle, branch and stem biomass participated by 22 %, 24 % and 54 % in highland, and by 19 %, 23 % and 58 % in mountain area, respectively. Silvicultural management affects mainly structure, density, and tree species composition of the stand. Therefore, dendrometric parameters of a tree resulted from genotype, growth conditions and from management history as well. Low type of thinning (LT; common in highland) stimulates rather tree

  3. Structuring and evolution of Neogene transcurrent basins in the Tellian foreland domain, north-eastern Tunisia

    NASA Astrophysics Data System (ADS)

    Melki, Fetheddine; Zouaghi, Taher; Harrab, Salah; Sainz, Antonio Casas; Bédir, Mourad; Zargouni, Fouad

    2011-07-01

    The Neogene sedimentary basins (Serravallian to Quaternary) of the Tellian tectonic foreland in north-eastern Tunisia formed within the overall NE-SW sinistral strike-slip tectonic framework of the Ras El Korane-Thibar and El Alia-Teboursouk fault systems. From stratigraphic logs, structural cross sections and interpretation of 2D seismic lines and boreholes, the pre-Neogene basement can be interpreted to be structured according to Eocene (NW-SE) compressional and Oligocene extensional phases. This basement comprises structural highs (anticlines and horsts) and subsiding areas (synclines, half-grabens and grabens) formed during the Neogene. The subsiding areas are delineated by faults striking N030E, N-S and N140E, defining (i) narrow, strongly subsiding synclines, (ii) lozenge-shaped basins and (iii) trapezoidal basins. The architecture of their fill results from the sedimentary balance between tectonics and eustatism. Halokinesis and clay diapirism (driven by Triassic and Neogene evaporites and clays) also played an important role in basin evolution, contributing to the formation of domes and diapirs along active faults.

  4. Trophic size-structure of sailfish Istiophorus platypterus in eastern Taiwan estimated by stable isotope analysis.

    PubMed

    Tsai, C-N; Chiang, W-C; Sun, C-L; Shao, K-T; Chen, S-Y; Yeh, S-Z

    2014-02-01

    To examine trophic dynamics over different size classes, an isotopic study of sailfish Istiophorus platypterus life-history stages was carried out. Samples were collected from eastern Taiwan and the South China Sea during April 2009 and February 2012. A total of 263 samples (111-245 cm, lower jaw fork length, LLJFL ) were examined for changes in trophic structure in relation to LLJFL by using stable isotope analysis of carbon (δ(13) C) and nitrogen (δ(15) N). The δ(15) N values for I. platypterus ranged from 7·51 to 14·19‰ (mean ± s.d. = 12·06 ± 1·16‰) and the δ(13) C values ranged from -22·04 to -15·48‰ (mean ± s.d. = -17·62 ± 1·10‰). The δ(15) N values were positively dependent on LLJFL (r(2)  = 0·377), whereas δ(13) C were negatively dependent on LLJFL (r(2)  = 0·063). There were significantly different seasonal changes in nitrogen and carbon isotopic concentration, but no significant differences in concentrations between eastern Taiwan and the South China Sea were reported. The trophic level (TL ) of each LLJFL class was correlated, starting from 2·84 TL for size class I (LLJFL  < 140 cm) and reaching 5·03 TL for size class VI (LLJFL > 221 cm). The mean ± s.d. TL was 4·43 ± 0·19 for all samples. The results reveal that I. platypterus occupies a wide range of trophic levels and different size classes occupy different trophic positions in the pelagic ecosystem.

  5. The uppermost mantle shear wave velocity structure of eastern Africa from Rayleigh wave tomography: constraints on rift evolution

    NASA Astrophysics Data System (ADS)

    O'Donnell, J. P.; Adams, A.; Nyblade, A. A.; Mulibo, G. D.; Tugume, F.

    2013-08-01

    An expanded model of the 3-D shear wave velocity structure of the uppermost mantle beneath eastern Africa has been developed using earthquakes recorded by the AfricaArray East African Seismic Experiment in conjunction with data from permanent stations and previously deployed temporary stations. The combined data set comprises 331 earthquakes recorded on a total of 95 seismic stations spanning Kenya, Uganda, Tanzania, Zambia and Malawi. In this study, data from 149 earthquakes were used to determine fundamental-mode Rayleigh wave phase velocities at periods ranging from 20 to 182 s using the two-plane wave method, and then combined with the similarly processed published measurements and inverted for a 3-D shear wave velocity model of the uppermost mantle. New features in the model include (1) a low-velocity region in western Zambia, (2) a high-velocity region in eastern Zambia, (3) a low-velocity region in eastern Tanzania and (4) low-velocity regions beneath the Lake Malawi rift. When considered in conjunction with mapped seismicity, these results support a secondary western rift branch striking southwestwards from Lake Tanganyika, likely exploiting the relatively weak lithosphere of the southern Kibaran Belt between the Bangweulu Block and the Congo Craton. We estimate a lithospheric thickness of ˜150-200 km for the substantial fast shear wave anomaly imaged in eastern Zambia, which may be a southward subsurface extension of the Bangweulu Block. The low-velocity region in eastern Tanzania suggests that the eastern rift branch trends southeastwards offshore eastern Tanzania coincident with the purported location of the northern margin of the proposed Ruvuma microplate. Pronounced velocity lows along the Lake Malawi rift are found beneath the northern and southern ends of the lake, but not beneath the central portion of the lake.

  6. Third year effects of cloudwater and ozone on red spruce seedlings

    SciTech Connect

    Pier, P.A.; Thornton, F.C.; McDuffie, C. Jr. )

    1991-05-01

    The reduction in growth of high elevation red spruce in the eastern US has been attributed in part to greater exposure to atmospheric pollution which occurs at high elevation. The authors objective was to evaluate the impact of ambient ozone and cloudwater deposition on the growth of red spruce seedlings at a high elevation site. Potted native and Phyton-grown (Phyton Technologies) red spruce seedlings were exposed in open-top field chambers at Whitetop Mountain, Virginia (elevation 1,680) for the third season to treatments of: (1) exclusion of clouds and 50% reduction in ambient O{sub 3} (COE), (2) O{sub 3} with clouds excluded (CO), (3) exposure to clouds and O{sub 3}, as control chambers (CC), and (4) open plots (AA). Plant biomass components and diameter increment growth for both seedling types were not affected by treatments. Photosynthesis was not enhanced by removal of cloudwater and O{sub 3}. Respiration (R{sub d}) generally was not affected by treatments; however, R{sub d} in native seedling needles of previous year and two-year previous growth was significantly greater in CC than CO and COE on several sampling dates, indicating that cloudwater and O{sub 3} may be causing higher R{sub d}.

  7. Predicting community structure in snakes on Eastern Nearctic islands using ecological neutral theory and phylogenetic methods.

    PubMed

    Burbrink, Frank T; McKelvy, Alexander D; Pyron, R Alexander; Myers, Edward A

    2015-11-22

    Predicting species presence and richness on islands is important for understanding the origins of communities and how likely it is that species will disperse and resist extinction. The equilibrium theory of island biogeography (ETIB) and, as a simple model of sampling abundances, the unified neutral theory of biodiversity (UNTB), predict that in situations where mainland to island migration is high, species-abundance relationships explain the presence of taxa on islands. Thus, more abundant mainland species should have a higher probability of occurring on adjacent islands. In contrast to UNTB, if certain groups have traits that permit them to disperse to islands better than other taxa, then phylogeny may be more predictive of which taxa will occur on islands. Taking surveys of 54 island snake communities in the Eastern Nearctic along with mainland communities that have abundance data for each species, we use phylogenetic assembly methods and UNTB estimates to predict island communities. Species richness is predicted by island area, whereas turnover from the mainland to island communities is random with respect to phylogeny. Community structure appears to be ecologically neutral and abundance on the mainland is the best predictor of presence on islands. With regard to young and proximate islands, where allopatric or cladogenetic speciation is not a factor, we find that simple neutral models following UNTB and ETIB predict the structure of island communities. PMID:26609083

  8. Predicting community structure in snakes on Eastern Nearctic islands using ecological neutral theory and phylogenetic methods.

    PubMed

    Burbrink, Frank T; McKelvy, Alexander D; Pyron, R Alexander; Myers, Edward A

    2015-11-22

    Predicting species presence and richness on islands is important for understanding the origins of communities and how likely it is that species will disperse and resist extinction. The equilibrium theory of island biogeography (ETIB) and, as a simple model of sampling abundances, the unified neutral theory of biodiversity (UNTB), predict that in situations where mainland to island migration is high, species-abundance relationships explain the presence of taxa on islands. Thus, more abundant mainland species should have a higher probability of occurring on adjacent islands. In contrast to UNTB, if certain groups have traits that permit them to disperse to islands better than other taxa, then phylogeny may be more predictive of which taxa will occur on islands. Taking surveys of 54 island snake communities in the Eastern Nearctic along with mainland communities that have abundance data for each species, we use phylogenetic assembly methods and UNTB estimates to predict island communities. Species richness is predicted by island area, whereas turnover from the mainland to island communities is random with respect to phylogeny. Community structure appears to be ecologically neutral and abundance on the mainland is the best predictor of presence on islands. With regard to young and proximate islands, where allopatric or cladogenetic speciation is not a factor, we find that simple neutral models following UNTB and ETIB predict the structure of island communities.

  9. Halokinesis and structural evolution of the major features in eastern and southern Tunisian Atlas

    NASA Astrophysics Data System (ADS)

    Hlaiem, Amor

    1999-05-01

    The eastern and southern Tunisian Atlas is marked by the presence of three major faults; the N-S Axis Fault, the NW-SE Gafsa Fault, and the E-W northern Chott Fault. Seismic lines crossing these structures show the existence of significant halokinetics that began in the Early Jurassic. The aim of this paper is to study and better understand the Triassic salt movements, which are mainly linked to periods when the major basement faults were reactivated resulting in extension, compression, transtension or/and transpression. Analysis of the seismic data allows us to conclude that the Triassic salt migration started in the Early Jurassic and continued during the Late Jurassic and the Early Cretaceous. Although these periods are characterized by an extensional tectonic regime, the synsedimentary halokinesis is accompanied by thickening and thinning of the overlying series. Salt remobilization was accentuated during the compressional deformation which occurred from the Late Tertiary to Quaternary and resulted in local diapiric extrusions along the major structural features.

  10. Inner Structure of Atmospheric Inversion Layers over Haifa Bay in the Eastern Mediterranean

    NASA Astrophysics Data System (ADS)

    Haikin, N.; Galanti, E.; Reisin, T. G.; Mahrer, Y.; Alpert, P.

    2015-09-01

    Capping inversions act as barriers to the vertical diffusion of pollutants, occasionally leading to significant low-level air pollution episodes in the lower troposphere. Here, we conducted two summer campaigns where global positioning system radiosondes were operated in Haifa Bay on the eastern Mediterranean coast, a region of steep terrain with significant pollution. The campaigns provided unique high resolution measurements related to capping inversions. It was found that the classical definition of a capping inversion was insufficient for an explicit identification of a layer; hence additional criteria are required for a complete spatial analysis of inversion evolution. Based on the vertical temperature derivative, an inner fine structure of inversion layers was explored, and was then used to track inversion layers spatially and to investigate their evolution. The exploration of the inner structure of inversion layers revealed five major patterns: symmetric peak, asymmetric peak, double peak, flat peak, and the zig-zag pattern. We found that the symmetric peak is related to the strongest inversions, double peak inversions tended to break apart into two layers, and the zig-zag pattern was related to the weakest inversions. Employing this classification is suggested for assistance in following the evolution of inversion layers.

  11. Seismic velocity structure and earthquake relocation for the magmatic system beneath Long Valley Caldera, eastern California

    NASA Astrophysics Data System (ADS)

    Lin, Guoqing

    2015-04-01

    A new three-dimensional (3-D) seismic velocity model and high-precision location catalog for earthquakes between 1984 and 2014 are presented for Long Valley Caldera and its adjacent fault zones in eastern California. The simul2000 tomography algorithm is applied to derive the 3-D Vp and Vp/Vs models using first-arrivals of 1004 composite earthquakes obtained from the original seismic data at the Northern California Earthquake Data Center. The resulting Vp model reflects geological structures and agrees with previous local tomographic studies. The simultaneously resolved Vp/Vs model is a major contribution of this study providing an important complement to the Vp model for the interpretation of structural heterogeneities and physical properties in the study area. The caldera is dominated by low Vp anomalies at shallow depths due to postcaldera fill. High Vp and low Vp/Vs values are resolved from the surface to ~ 3.4 km depth beneath the center of the caldera, corresponding to the structural uplift of the Resurgent Dome. An aseismic body with low Vp and high Vp/Vs anomalies at 4.2-6.2 km depth below the surface is consistent with the location of partial melt suggested by previous studies based on Vp models only and the inflation source locations based on geodetic modeling. The Sierran crystalline rocks outside the caldera are generally characterized with high Vp and low Vp/Vs values. The newly resolved velocity model improves absolute location accuracy for the seismicity in the study area and ultimately provides the basis for a high-precision earthquake catalog based on similar-event cluster analysis and waveform cross-correlation data. The fine-scale velocity structure and precise earthquake relocations are useful for investigating magma sources, seismicity and stress interaction and other seismological studies in Long Valley.

  12. Significant role of structural fractures in Ren-Qiu buried-block oil field, eastern China

    SciTech Connect

    Fei, Q.; Xie-Pei, W.

    1983-03-01

    Ren-qui oil field is in a buried block of Sinian (upper Proterozoic) rocks located in the Ji-zhong depression of the western Bohai Bay basin in eastern China. The main reservoir consists of Sinian dolomite rocks. It is a fault block with a large growth fault on the west side which trends north-northeast with throws of up to 1 km (0.6 mi) or more. The source rocks for the oil are Paleogene age and overlie the Sinian dolomite rocks. The structural fractures are the main factor forming the reservoir of the buried-block oil field. Three structural lines, trending northeast, north-northeast, and northwest, form the regional netted fracture system. The north-northeast growth fault controlled the structural development of the buried block. The block was raised and eroded before the Tertiary sediments were deposited. In the Eocene Epoch, the Ji-zhong depression subsided, but the deposition, faulting, and related uplift of the block happened synchronously as the block was gradually submerged. At the same time, several horizontal and vertical karst zones were formed by the karst water along the netted structural fractures. The Eocene oil source rocks lapped onto the block and so the buried block, with many developed karst fractures, was surrounded by a great thickness of source rocks. As the growth fault developed, the height of the block was increased from 400 m (1300 ft) before the Oligocene to 1300 m (4250 ft) after. As the petroleum was generated, it migrated immediately into the karst fractures of the buried block along the growth fault. The karst-fractured block reservoir has an 800-m (2600-ft) high oil-bearing closure and good connections developed between the karst fractures.

  13. Structure, intrusion, and tectonic origin of the Independence dike swarm, eastern California

    NASA Astrophysics Data System (ADS)

    Carl, Brian Stewart

    2000-10-01

    The Jurassic (˜148 Ma) Independence dike swarm is exposed over a distance of >600 km and has served as a valuable temporal and structural marker across eastern California. This study investigates new structural, field, and compositional data associated with the swarm's tectonic origin, deformation and emplacement, and proposes models to explain these data. In addition, this study explores an enigmatic relationship between deformed mafic dikes and their undeformed granitic host. In the eastern Sierra, Independence dikes within Jurassic plutons cut and are themselves deformed by NNE-striking reverse-sense mylonitic shear zones, providing strong evidence for syndeformational dike intrusion. Many dikes contain a penetrative, solid-state ductile fabric which formed during sinistral shear across the dikes. Mylonitization and sinistral shear along Sierran dikes may have occurred along a Late Jurassic transpressional Cordilleran arc. Swarm intrusion likely occurred in response to a rapid change in North American plate motion rather than during rifting of the arc or in response to a collisional event. Composite Independence dikes may indicate the mode of dike emplacement and magma interaction at depth. Most Sierran composite dikes contain subparallel mafic intrusions but a few contain both mafic and felsic phases. Along more southerly portions of the swarm, in the Spangler Hills and in the Granite and Fry mountains, composite dikes contain abundant enclaves. These features may correlate with dike compositions. In the Sierra, composite dikes are dominantly either mafic or felsic, whereas more southerly composite dikes include intermediate phases. If dikes tapped plutons emplaced at similar paleodepths, then dike features and compositional variability may reflect different degrees of mixing along dike conduits. Rock mechanics experiments indicate that mafic rocks should be more competent than quartz-rich granite. Dikes which are strongly mylonitized along their margins

  14. Boreal peatland margins as repository sites of long-term natural disturbances of balsam fir/spruce forests

    NASA Astrophysics Data System (ADS)

    Lavoie, Martin; Filion, Louise; Robert, Élisabeth C.

    2009-05-01

    A multidisciplinary, high-resolution paleoecological study (Lepidoptera and plant remains, macroscopic charcoal, pollen) was conducted on a 4000-yr peat monolith extracted from the margin of an ombrotrophic peatland on Anticosti Island (Gulf of St. Lawrence, eastern Canada) to reconstruct the long-term natural disturbances (insect outbreaks, forest fires) of a balsam fir/spruce forest. We hypothesized that an activity of insect defoliators (spruce budworm, hemlock looper) was the main disturbance factor of conifer forests during the Late Holocene. The earliest remains of spruce budworm and hemlock looper were found ca. 3220 and 2350 cal yr BP, respectively. Peaks of insect head capsules occurred from ca. 1640 to ca. 625 cal yr BP. Low balsam fir pollen concentrations during this period suggest a lengthy episode (˜ 1000 yr) of high insect activity, resulting in extensive fir dieback and mortality. The long-term dynamics of the pristine balsam fir/spruce forests were mainly governed by the activity of insect defoliators. The limited extent and possibly the low occurrence of forest fires in the maritime environment of Anticosti Island allowed the development of mature coniferous stands propitious for insect infestations. Insect head capsules appeared to be a useful and effective tool for establishing insect presence and activity during the Holocene.

  15. Stem analysis of a long-lived black spruce clone at treeline

    SciTech Connect

    Payette, S.; Delwaide, A.; Morneau, C.; Lavoie, C. )

    1994-02-01

    Because of its ability to layer and to produce different phenotypes, black spruce (Picea mariana [Mill.] BSP.) develops a complex clonal structure ensuring its survival and longevity. Here we report tree stem development and demise of a black spruce clone at treeline over the last 500 yr. Since the 16th century, the apical meristems of the clonal spruce experienced three periods of stem development associated with brief warmings and two periods of stem decline corresponding to known cold spells of the Little Ice Age. Ortet development was particularly vigorous in the 16th century, while the two layered stems slowly developed in the late 17th century and in the 20th century, respectively. Stem decline appeared as a progressive process lasting for several decades in the form of a basipetal death-gradient along the bole amplified by the above/below snow-pack position. Stem elongation was possibly facilitated by lesser winter-snow abrasion and/or thicker snowpack. Clonal stem development may have important implications for spruce spread in the arctic tundra in a warmer world. 26 refs. 2 figs.

  16. Detailed structure and stratigraphy of the eastern Marble Mountain terrane, Klamath Mountains, CA

    SciTech Connect

    Miller, D.E.; Hacker, B.R. . Dept. of Geology)

    1993-04-01

    Amphibolite-grade rocks in the eastern Marble Mountains (MM), N. California, consist of several fault-bounded, SSE-dipping lithotectonic units. Each unit is ca. 2 km thick and is characterized by differences in rock type, metamorphism, and structural style. The lowermost unit composed of well-foliated and lineated epidote amphibolite grading upward into clinopyroxene-bearing amphibolite with a consistent NE-SW lineation. Structurally overlying these rocks are andalusite- and staurolite-bearing, epidote-amphibolite facies rocks (Wright Lake assemblage (WLa)) that have variable foliation and lineation orientations. The WLa consists of meta-supracrustal rocks with well-preserved relict textures, and massive, meta-ultramafic rock. Supracrustal rocks include polymict conglomerate and breccia, fine- to medium-grained clastic rocks, alkalic pillow basalt, chert, and carbonate. Conglomerate clasts include partially recrystallized granitoids and quartzite. Previous studies have interpreted the WLa to represent a fragment of oceanic crust, but coeval coarse-grained sedimentation and alkalic volcanism, small volume of mafic volcanics, conglomerate composition, and lack of oceanic plutonic and hypabyssal rocks suggest deposition in an arc-related rift or transtensional basin. Previous studies have also described the terrane as melange, but recognition of local pseudostratigraphy allows mapping of multiply folded, isoclinal, nappe-like structures. Small-scale nappes are generally 100+ m thick and are imbricated with massive meta-ultramafic rocks along gently to steeply east-dipping shear zones. Shear zones are characterized by metamorphosed ultramafic fault rocks that suggest a range of brittle to ductile behavior. Regionally distributed, Ar/Ar hornblende ages of 149.9[+-]0.4, 150.3[+-]0.6, 152.1[+-]4.7, 152.5[+-]2.5 Ma and Ar/Ar biotite ages of 148.8[+-]2.6 and 149.9[+-]0.4 Ma indicate the MM terrane cooled rapidly through ca. 500--300 C in the Late Jurassic.

  17. Crustal structure of the Eastern Sierras Pampeanas of Argentina using high frequency local receiver functions

    NASA Astrophysics Data System (ADS)

    Perarnau, Marcelo; Gilbert, Hersh; Alvarado, Patricia; Martino, Roberto; Anderson, Megan

    2012-12-01

    The Eastern Sierras Pampeanas are basement cored outcrops uplifted in the Andean foreland where the easternmost segment of the Pampean flat slab segment starts dipping more steeply into the deeper mantle. These ranges of central Argentina known as the Sierras de Córdoba have an enriched-quartz composition and are bounded by a series of reverse faults. Different models have been suggested to represent the style of the thick-skinned deformation in this area. However the overall structure linking the exposed faults and terrane boundaries with their probable continuation at depth is unknown. In this paper we present images of the crustal structure beneath the Sierras de Córdoba using the common conversion point stacking method of high frequency local receiver functions recorded by the ESP broadband seismic array. The work consists of two transects located around 31°S and 32°S across the Sierras de Córdoba. The results show a consistent sharp Moho signal associated with a high contrast in seismic velocities in good agreement with the granitic character of the crust lying above the mafic upper mantle. The Moho morphology varies exhibiting thicknesses of 38 km in the west to 35 km in the east with a vertical shifting under the Sierra Chica. We relate this variable character to the presence of the boundary between the Pampia terrane and the Rio de La Plata craton. Our results for the intra-crustal structure indicate the presence of three discontinuities in the northern transect and at least two discontinuities in the southern transect. These discontinuities appear vertically displaced beneath the surface traces of the major range bounding faults providing evidence for the continuation to mid-crustal depths of the exposed reverse faults. Thus, the reverse faults seem to have displaced several horizontal intra-crustal interfaces. Finally in the lower crust we found a region which seems to be aseismic.

  18. Contemporary Genetic Structure, Phylogeography and Past Demographic Processes of Wild Boar Sus scrofa Population in Central and Eastern Europe

    PubMed Central

    Kusza, Szilvia; Podgórski, Tomasz; Scandura, Massimo; Borowik, Tomasz; Jávor, András; Sidorovich, Vadim E.; Bunevich, Aleksei N.; Kolesnikov, Mikhail; Jędrzejewska, Bogumiła

    2014-01-01

    The wild boar (Sus scrofa) is one of the most widely distributed mammals in Europe. Its demography was affected by various events in the past and today populations are increasing throughout Europe. We examined genetic diversity, structure and population dynamics of wild boar in Central and Eastern Europe. MtDNA control region (664 bp) was sequenced in 254 wild boar from six countries (Poland, Hungary, Belarus, Ukraine, Moldova and the European part of Russia). We detected 16 haplotypes, all known from previous studies in Europe; 14 of them belonged to European 1 (E1) clade, including 13 haplotypes from E1-C and one from E1-A lineages. Two haplotypes belonged respectively to the East Asian and the Near Eastern clade. Both haplotypes were found in Russia and most probably originated from the documented translocations of wild boar. The studied populations showed moderate haplotype (0.714±0.023) and low nucleotide diversity (0.003±0.002). SAMOVA grouped the genetic structuring of Central and Eastern European wild boar into three subpopulations, comprising of: (1) north-eastern Belarus and the European part of Russia, (2) Poland, Ukraine, Moldova and most of Belarus, and (3) Hungary. The multimodal mismatch distribution, Fu's Fs index, Bayesian skyline plot and the high occurrence of shared haplotypes among populations did not suggest strong demographic fluctuations in wild boar numbers in the Holocene and pre-Holocene times. This study showed relatively weak genetic diversity and structure in Central and Eastern European wild boar populations and underlined gaps in our knowledge on the role of southern refugia and demographic processes shaping genetic diversity of wild boar in this part of Europe. PMID:24622149

  19. Influences of forest structure, climate and species composition on tree mortality across the eastern US.

    PubMed

    Lines, Emily R; Coomes, David A; Purves, Drew W

    2010-10-13

    Few studies have quantified regional variation in tree mortality, or explored whether species compositional changes or within-species variation are responsible for regional patterns, despite the fact that mortality has direct effects on the dynamics of woody biomass, species composition, stand structure, wood production and forest response to climate change. Using bayesian analysis of over 430,000 tree records from a large eastern US forest database we characterised tree mortality as a function of climate, soils, species and size (stem diameter). We found (1) mortality is U-shaped vs. stem diameter for all 21 species examined; (2) mortality is hump-shaped vs. plot basal area for most species; (3) geographical variation in mortality is substantial, and correlated with several environmental factors; and (4) individual species vary substantially from the combined average in the nature and magnitude of their mortality responses to environmental variation. Regional variation in mortality is therefore the product of variation in species composition combined with highly varied mortality-environment correlations within species. The results imply that variation in mortality is a crucial part of variation in the forest carbon cycle, such that including this variation in models of the global carbon cycle could significantly narrow uncertainty in climate change predictions.

  20. Detectability of temporal changes in fine structures near the inner core boundary beneath the eastern hemisphere

    NASA Astrophysics Data System (ADS)

    Yu, W.

    2016-07-01

    The inner core boundary (ICB), where melting and solidification of the core occur, plays a crucial role in the dynamics of the Earth's interior. To probe temporal changes near the ICB beneath the eastern hemisphere, I analyze differential times of PKiKP (dt(PKiKP)), double differential times of PKiKP-PKPdf, and PKiKP coda waves from repeating earthquakes in the southwest Pacific subduction zones. dt(PKiKP) values are mostly within ±30 ms of one another, without systematic temporal dependence. Some observations of PKiKP coda waves have absolute time shifts of >50 ms relative to their main phases. The combination of temporal changes in PKiKP coda arrivals and negligible changes in PKiKP arrivals favors a smooth ICB with fine-scale structures in the upper inner core. dt(PKiKP) values are interpreted in the context of melting- or growth-induced ICB topography, based on dynamic models. Uncertainties in dt(PKiKP) prevent verification of ICB melting or growth on decadal time scales.

  1. Thermocline Structure and ENSO Variability in the eastern equatorial Pacific during the Last Glacial Maximum

    NASA Astrophysics Data System (ADS)

    Clark, S.; Koutavas, A.; Lynch-Stieglitz, J.; Rustic, G.

    2015-12-01

    The mean state and variability of the eastern equatorial Pacific (EEP) during the Last Glacial Maximum (LGM) are of great interest because of the region's role in the El Niño-Southern Oscillation (ENSO) and global climate. We investigated changes in thermocline structure between the Late Holocene (LH) and LGM with stable isotopes of planktonic foraminifera in sediment cores from the Galápagos. We measured δ18O in two species—Globigerinoides ruber, inhabiting the surface mixed layer, and Neogloboquadrina dutertrei, inhabiting the deep thermocline—in order to evaluate the vertical temperature contrast between the two species. We also measured δ18O of individual N. dutertrei from modern (late 20th century) and LGM sediments in order to assess thermocline temperature variability related to ENSO activity. Our data indicate a reduced vertical contrast in the upper ocean during the LGM, which is most consistent with a deeper thermocline and thicker mixed layer. Additionally, δ18O of individual N. dutertrei shells shows 2.5 times greater population variance in the LGM than in the modern sample. This large variance indicates that thermocline temperatures were more variable during the LGM than today, consistent with more active ENSO. Together, these results imply that the mean state of the EEP was characterized by a deeper thermocline and greater ENSO variability. The results further show the potential for reconstructing ENSO variability from deep-sea sediments of the EEP, where other geological archives of ENSO are currently extremely limited.

  2. Genetic structure of populations of Rhizoctonia solani AG-3 on potato in eastern North Carolina.

    PubMed

    Ceresini, Paulo C; Shew, H David; Vilgalys, Rytas J; Rosewich, U Liane; Cubeta, Marc A

    2002-01-01

    A polymerase chain reaction-restriction fragment length polymorphism (PCR-RFLP) method was developed to identify and differentiate genotypes of Rhizoctonia solani anastomosis group 3 subgroup PT (AG-3 PT), a fungal pathogen of potato. Polymorphic co-dominant single-locus PCR-RFLP markers were identified after sequencing of clones from a genomic library and digestion with restriction enzymes. Multilocus genotypes were determined by a combination of PCR product and digestion with a specific restriction enzyme for each of seven loci. A sample of 104 isolates from one commercial field in each of five counties in eastern North Carolina was analyzed, and evidence for high levels of gene flow between populations was revealed. When data were clone-corrected and samples pooled into one single North Carolina population, random associations of alleles were found for all loci or pairs of loci, indicating random mating. However, when all genotypes were analyzed, the observed genotypic diversity deviated from panmixia and alleles within and between loci were not randomly associated. These findings support a model of population structure for R. solani AG-3 PT on potato that includes both recombination and clonality.

  3. Adria-Europe crustal structure relationship in the Eastern Alps (project EASI)

    NASA Astrophysics Data System (ADS)

    Hetényi, György; Plomerová, Jaroslava; Bianchi, Irene; Kampfová Exnerová, Hana

    2016-04-01

    Project EASI is the first implemented Complementary Experiment within the AlpArray program (http://www.alparray.ethz.ch) and stands for Eastern Alpine Seismic Investigation. The seismological field experiment ran for one year, from Summer 2014 to Summer 2015, composed of 55 broadband stations deployed in zig-zag in a ca. 15 km-wide band along longitude 13.35°E, spanning 540 km from the Czech-German border to the Adriatic Sea. Here we present first results using P-to-S converted waves from teleseismic distances. The variation of Moho depth along the profile is analyzed and linked to the two colliding plates, Adria and Europe, as well as to the overlying lithospheric blocks of the Bohemian Massif. The suggested Moho "hole" between Adria and Europe is characterized. We investigate the anisotropic nature of the lower crust of both plates. We conclude on the structural relationship of Adria and Europe at the crustal level, and infer their respective positions at depth. Furthermore, preliminary S-to-P conversions illuminating the lithosphere-asthenosphere boundary test the significant depth variation of this boundary along the EASI transect and complement our receiver function study.

  4. Geological structure of the deep eastern Mediterranean Sea (east of 25 Degrees E)

    SciTech Connect

    Montadert, L.; Sage, L.; Letouzey, J. )

    1988-08-01

    The deformation fronts of the Cyprus arc and the Mediterranean ridge, extending from the Turkey-Syria boundary to north Cyrenacia, are the southernmost superficial expression of the convergence between the Eurasian and African plates. They separate the Eastern Mediterranean deep basin into two different structural units: (1) A thrust belt, northward, with the presence of Cenozoic sedimentary basins which could be considered piggy-back basins (Iskenderun, Adana, Cilicia, and Antalya basins). These basins, filled by 4,000 to 6,000 m of Cenozoic sediments, lie on a substratum composed of south-vergent nappes emplaced between the Late Cretaceous and the late Miocene. (2) A foreland area, southward, where the thick Herodotus and Levantine sedimentary basins, relatively undeformed, lie on the passive and subsident African continental margin initiated during Late Triassic or early Liassic time. Due to the still-active collision between the thrust belt and the Erathosthenes seamount, Cyprus was uplifted and today represents the emerged part of the deformation front. During the Messinian, with the isolation of the Mediterranean Sea, evaporitic deposits including a salt layer (sometimes more than 2,000 m thick) were widely distributed into the Iskenderun, Cilicia, Antalya, Levantine, and Herodotus basins. In these basins, the Messinian sedimentation was directly controlled by basin topography.

  5. ANOTHER LOOK AT THE EASTERN BANDED STRUCTURE: A STELLAR DEBRIS STREAM AND A POSSIBLE PROGENITOR

    SciTech Connect

    Grillmair, C. J.

    2011-09-01

    Using the Sloan Digital Sky Survey Data Release 7, we re-examine the Eastern Banded Structure (EBS), a stellar debris stream first discovered in Data Release 5 and more recently detected in velocity space by Schlaufman et al. The visible portion of the stream is 18{sup 0} long, lying roughly in the Galactic Anticenter direction and extending from Hydra to Cancer. At an estimated distance of 9.7 kpc, the stream is {approx}170 pc across on the sky. The curvature of the stream implies a fairly eccentric box orbit that passes close to both the Galactic center and to the Sun, making it dynamically distinct from the nearby Monoceros, Anticenter, and GD-1 streams. Within the stream is a relatively strong, 2{sup 0}-wide concentration of stars with a very similar color-magnitude distribution that we designate Hydra I. Given its prominence within the stream and its unusual morphology, we suggest that Hydra I is the last vestige of EBS's progenitor, possibly already unbound or in the final throes of tidal dissolution. Though both Hydra I and the EBS have a relatively high-velocity dispersion, given the comparatively narrow width of the stream and the high frequency of encounters with the bulge and massive constituents of the disk that such an eccentric orbit would entail, we suggest that the progenitor was likely a globular cluster and that both it and the stream have undergone significant heating over time.

  6. Application of electrical resistivity tomography to map lithological differences and subsurface structures (Eastern Sudetes, Czech Republic)

    NASA Astrophysics Data System (ADS)

    Stan, Dominika; Stan-Kłeczek, Iwona

    2014-09-01

    The aim of this study is to determine the subsurface lithological variability and thickness of each metamorphic rocks unit by means of geophysical methods, which were presented within the framework of SEDIBUD. Research was performed on the ridge and the steep eastern slope of the Orlik massif (1204 m) in the Hrubý Jesenik range of the Eastern Sudetes, the Czech Republic. To obtain a spatial image of a geological rock mass, the non-invasive and relatively rapid method of electrical resistivity tomography (ERT) was used, which resulted in the creation of six profiles using the Dipole-Dipole and Wenner-Schlumberger arrays. As complementary methods, vertical electrical sounding (VES) and seismic refraction tomography were chosen. Fifteen measuring points along two profiles obtained by vertical electrical sounding allowed the determination of the following in the vertical section layers of different resistivity and thickness at direct points in contact primarily with Devonian quartzite rocks and weathered covers. In a series of field studies, a four-layered rock mass was examined in which the first layer is of fine-grained materials with medium- and large-size clasts, the second layer is quartzite, the third is gneiss and the fourth is phyllite. The five profiles measured by seismic refraction tomography made it possible to draw the boundary between the studied phyllites, gneisses and quartzites. In addition, geophysical surveys and the results of mineralogical composition analysis allowed us to determine the nature and thickness of the weathered layers present on the massif area, which, in the upper sections, are predominantly a clayey loam partially covered with blocks. The inclination of the slope is associated with an increase in the fraction of coarse material. The majority of the thickness of these layers is related to the local flattened surfaces and the presence of numerous streams, which contributed to the accumulation of the sediments that were also

  7. Invasive symbiont bearing (and other) foraminifera altering the community structure of eastern Mediterranean rocky reefs environments

    NASA Astrophysics Data System (ADS)

    Hyams-Kaphzan, Orit; Perelis Grossowicz, Lydia; Almogi-Labin, Ahuva

    2015-04-01

    The rocky reefs of the Israeli eastern Mediterranean shelf constitute a highly diverse marine ecosystem rich in macroalgae and calcareous organisms. The benthic foraminiferal community living in this ecosystem is rapidly changing due to massive invasion of symbiont bearing foraminifera (SBF) as well as other foraminiferal species of tropical origin. This trend facilitated by the ongoing increase in temperature enables more tropical species to adjust to the eastern Mediterranean habitats. In order to document the status of the benthic foraminiferal community structure rocky reefs at Akhziv (AK) and Carmel Head (CH), northern Israel were sampled by scuba diving. Different macroalgae species, including invasive ones, accommodating the live epiphytic benthic foraminifera were sampled twice a year at AK and in each season at CH in three depth intervals between 5-20 m, during 2013-4. The numerical abundance of the group ranges between 170-3500 #/10cc (wet macroalgae volume) without any significant difference in standing stocks within regions, water depths or macroalgae preference. In total 77 benthic foraminiferal species were identified 71 in CH and only 43 at AK. Species richness per site varied between 3 and 42 with higher values at CH. 25% of all species were aliens, mostly Lessepsian, that comprise on average 70% - 84% of the numerical abundance of AK and CH respectively. Cluster analysis using benthic foraminifera relative abundance data did not correlate with the different macroalgae species, water depths or seasonality, indicating that the foraminiferal community in the two regions is quite homogenous. Amphistegina lobifera a Lessepsian migrant is by far the most common species on the Israeli rocky reefs occurring in all samples and comprising 18-93% of the foraminiferal community. Heterostegina depressa behaves similarly to A. lobifera though it occurs in lower numbers. Pararotalia calcariformata, a recently arriving SBF occupies mainly shallow water sites at CH

  8. Convection and Easterly Wave Structure Observed in the Eastern Pacific Warm-Pool during EPIC-2001

    NASA Technical Reports Server (NTRS)

    Peterson, Walter A.; Cifelli, R.; Boccippio, D.; Rutledge, S. A.; Fairall, C. W.; Arnold, James E. (Technical Monitor)

    2002-01-01

    During September-October 2001, the East Pacific Investigation of Climate Processes in the Coupled Ocean-Atmosphere System (EPIC-2001) ITCZ field campaign focused on studies of deep convection in the warm-pool region of the East Pacific. In addition to the TAO mooring array, observational platforms deployed during the field phase included the NOAA ship RN Ronald H. Brown, the NSF ship RN Horizon, and the NOAA P-3 and NCAR C-130 aircraft. This study combines C-band Doppler radar, rawinsonde, and surface heat flux data collected aboard the RN Brown to describe ITCZ convective structure and rainfall statistics in the eastern Pacific as a function of 3-5 day easterly wave phase. Three distinct easterly wave passages occurred during EPIC-2001. Wind and thermodynamic data reveal that the wave trough axes exhibited positively correlated U and V winds and a slight westward phase tilt with height. A relatively strong (weak) northeasterly deep tropospheric shear followed the trough (ridge) axis. Temperature and humidity perturbations exhibited mid-to upper level cooling (warming) and drying (moistening) in the northerly (trough and southerly) phase. At low levels warming (cooling) occurred in the northerly (southerly) phase with little change in the relative humidity, though mixed layer mixing ratios were larger during the northerly phase. When composited, radar, sounding, lightning and surface heat flux observations suggest the following systematic behavior as a function of wave phase: approximately zero to one quarter wavelength ahead of (behind) the wave trough in northerly (southerly) flow, larger (smaller) CAPE, lower (higher) CIN, weaker (stronger) tropospheric shear, higher (lower) conditional mean rain rates, higher (lower) lightning flash densities, and more (less) robust convective vertical structure occurred. Latent and sensible heat fluxes reached a minimum in the northerly phase and then increased through the trough, reaching a peak during the ridge phase

  9. Crustal structure beneath the eastern Swiss Alps derived from seismic refraction data

    NASA Astrophysics Data System (ADS)

    Ye, S.; Ansorge, J.; Kissling, E.; Mueller, St.

    1995-02-01

    The eastern part of the Swiss Central Alps is densely covered by a network of seismic refraction and wide-angle reflection profiles recorded mostly parallel to the tectonic strike of the Alps and along the newly acquired N-S-oriented European Geotraverse (EGT). To obtain a well-constrained crustal transect along the EGT an initial model was constructed from information based mainly on the coincident reflection seismic profile of the Swiss National Research Program NFP20 and short-range observations of the EGT data for the shallow structure and on the along-strike wide-angle profiles for the deep structure. The refinement of the initial model by 2-D raytracing during the subsequent interpretation of the EGT data leads to a detailed P-wave velocity distribution of the crustal cross-section beneath the Central Swiss Alps and its adjacent areas. In general, the distinctly layered crustal structure below the Alpine foreland thickens considerably as the Alps are approached, reaching a maximum thickness of nearly 60 km below the Insubric Line. The upper and middle crust has velocities between 6.0 and 6.2 km/s. Except for the area below the southern part of the Molasse Basin and the Helvetic nappes a distinct lower crust with a relatively low velocity of 6.5-6.6 km/s is found. Below the Penninic nappes the lower crust thickens remarkably, merging probably with the high-velocity zone of 6.6 km/s at a depth of about 21 km, which has been interpreted as the top of the indenting lower crust of the Adriatic promontory of the African plate. A clear vertical offset between the smoothly south-dipping European and the more rapidly rising Adriatic crust-mantle boundaries is found. The complex structures of the upper crust beneath the Alps caused by the nappe tectonics can only be partly resolved by the refraction seismic data. A south-dipping high-velocity zone within the Penninic nappe pile and a reflector beneath the northern front of the Aar Massif can possibly be interpreted as

  10. EFFECTS OF EASTERN REDCEDAR ON SMALL MAMMAL COMMUNITY STRUCTURE IN THE OKLAHOMA CROSSTIMBERS

    EPA Science Inventory

    Increased abundance of eastern redcedar (Juniperus virginianus), a native but invasive species in the Great Plains, has been associated with reduced herbaceous biomass in the canopy zone, altered plant species composition, and reduced understory light and soil water content. By ...

  11. Paleogene strata of the Eastern Los Angeles basin, California: Paleogeography and constraints on neogene structural evolution

    USGS Publications Warehouse

    McCulloh, T.H.; Beyer, L.A.; Enrico, R.J.

    2000-01-01

    Post-Paleogene dextral slip of 8-9 km is demonstrated for the southeastern part of the Whittier fault zone in the eastern Los Angeles basin area of southern California. A linear axis of greatest thickness for the combined upper Paleocene and lower to lower-middle Eocene clastic formations intersects the fault zone and is offset by it to give the new measure. Fragmentary evidence hints that the Whittier structural zone may have exerted control on bathymetric-topographic relief and sedimentation even in latest Paleocene (ca. 54 Ma). A clear topographic influence was exerted by 20-17 Ma. Strike-slip and present deformational style is younger than ca. 8 Ma. Our Paleogene isopach map extends as far west as long 117??58'W and is a foundation for companion zonal maps of predominant lithology and depositional environments. Integration of new palynological data with published biostratigraphic results and both new and published lithologic and sedimentological interpretations support the zonal maps. Reconstruction of marine-nonmarine facies and fragmented basin margins yields a model for the northeastern corner of a Paleogene coastal basin. Palinspastic adjustment for the Neogene-Quaternary Whittier fault offset and a reasoned westerly extension of the northern edge of the basin model yield a reconstruction of Paleogene paleogeography-paleoceanography. Our reconstruction is based partly on the absence of both Paleocene and Eocene deposits beneath the unconformable base of the middle Miocene Topanga Group in a region nowhere less than 15 km wide between the Raymond-Sierra Madre-Cucamonga fault zone and the northern edge of the Paleocene basin. Thus, Paleogene strata of the Santa Monica Mountains could not have been offset from the northern extension of the Santa Ana Mountains by sinistral slip on those boundary faults. Structural rearrangements needed to accommodate the clockwise rotation of the western Transverse Ranges from the early Miocene starting position are thereby

  12. Eastern Olympus Mons Basal Scarp and Aureole lobe: Recent Structural Evidence for Formation and Development

    NASA Astrophysics Data System (ADS)

    Weller, M. B.; McGovern, P. J.; Fournier, T.; Morgan, J. K.; Katz, O.

    2012-12-01

    The origin of the Olympus Mons (OM) basal scarp and the adjacent enigmatic aureole lobe deposits have long been controversial, with many hypotheses having been postulated for their linked formation, ranging from sub-glacial eruptions (e.g. Helgason, 1999), to numerous landslide and flank collapse models (e.g. Borgia, 1990; Lopes et al., 1980). While a landslide origin is compelling upon the examination of the OM basal scarp morphology, much of the controversy remains, and the subsurface structure of OM is not well understood. With the release of high resolution Digital Terrain Models (DTMs) of Mars from High Resolution Stereo Camera (HRSC) data, we examine a unique portion of the edifice, the Eastern OM Basal Scarp (EOMBS). The EOMBS exhibits two parallel bounding structures: a nearly 100 km-long up-slope extensional normal fault system, with associated offsets on the order of 10-100m, bordered by radial tear faults; and a 160-200 km-long down-slope contractional wrinkle ridge network of varying vergence, inferred to define the boundaries of a "pop-up" structure, associated offsets are on the order of 100m, and were last inferred to be active in recent geologic time at < 45 Ma (Basilevsky et al., 2006). The wrinkle ridge network extends an additional 60-100 km south of the identified normal fault system. We suggest that these fault systems, within the region of the radial tear faults, may be linked by failure surfaces, that their displacements accommodate downslope movement of the flank, and may be direct evidence linking the "East" aureole lobe to the EOMBS. Calculations based on limit equilibrium principles, and matched to the observed faulting locations and slope geometries, suggests a mechanically weak, pore-fluid saturated detachment layer (e.g. similar to phyllosilicates), at a depth of 0 to -3km below the reference datum, underlies eastern OM, consistent with models of OM as a spreading volcanic edifice (Borgia, 1990; McGovern and Morgan, 2009). Our

  13. Space-time structure of the 2003 geomagnetic jerk at Mid-Eastern Asia

    NASA Astrophysics Data System (ADS)

    Ou, Jiaming; Du, Aimin; Xu, Wenyao; Yang, Dongmei

    2015-04-01

    The 2003 jerk has an abrupt change in the geomagnetic secular variation (SV), and was recognized as a local phenomenon of internal origin from the satellite observations (Olsen and Mandea, 2007). Notable strength of the 2003 jerk is located at Mid-Eastern Asia. The temporal and spatial features at this area are important to resolve the Earth's core fluid flow dynamics at local scale (e.g. Wardinski et al., 2008). We investigate the temporal-spatial development of the 2003 jerk in more detail at Mid-Eastern Asia with the ground-based observations and CHAOS-3 core field model. We select the data in the international geomagnetic quiet days to calculate the monthly means. In order to reduce the influence of the external field, we adopt a function comprising the terms associated with the indices of the geomagnetic activity, and the terms of the periodic signals on the observatory monthly means data (Stewart and Whaler, 1992). We then use an empirical AR-2 model to represent the internal field signals in the observatory data. The extreme detection is applied to identify the jerk in the SV time series. The onset time and the strength of the 2003 jerk are obtained through the detection for geomagnetic field component, X, Y and Z. The maximum of the strength of the 2003 jerk is located under the Indian mainland. The onset time of this jerk propagates approximately southeastward. Two jerks in 2001 and 2003 for the Z component are further compared and they are confirmed as independent processes. We suggest the jerk in 2001 identical to the well known 1999 jerk in Europe (Mandea et al., 2000). Our results reveal the fine structures of the 2003 jerk that corroborate the conclusions in previous studies. The larger scale time-spatial structure given by the AR-2 model constructed from ground observatory data (monthly values) is consistent with the results from the CHAOS-3 model. This structure can be applied for further inversion of the local core surface fluid flow motions

  14. The P and S wave velocity structure of the mantle beneath eastern Africa and the African superplume anomaly

    NASA Astrophysics Data System (ADS)

    Mulibo, Gabriel D.; Nyblade, Andrew A.

    2013-08-01

    P and S relative arrival time residuals from teleseismic earthquakes recorded on over 60 temporary AfricaArray broadband seismic stations deployed in Uganda, Tanzania, and Zambia between 2007 and 2011 have been inverted, together with relative arrival time residuals from earthquakes recorded by previous deployments, for a tomographic image of mantle wave speed variations extending to a depth of 1200 km beneath eastern Africa. The image shows a low-wave speed anomaly (LWA) well developed at shallow depths (100-200 km) beneath the Eastern and Western branches of the Cenozoic East African rift system and northwestern Zambia, and a fast wave speed anomaly at depths ≤ 350 km beneath the central and northern parts of the East African Plateau and the eastern and central parts of Zambia. At depths ≥350 km the LWA is most prominent under the central and southern parts of the East African Plateau and dips to the southwest beneath northern Zambia, extending to a depth of at least 900 km. The amplitude of the LWA is consistent with a ˜150-300 K thermal perturbation, and its depth extent indicates that the African superplume, originally identified as a lower mantle anomaly, is likely a whole mantle structure. A superplume extending from the core-mantle boundary to the surface implies an origin for the Cenozoic extension, volcanism, and plateau uplift in eastern Africa rooted in the dynamics of the lower mantle.

  15. Structure and evolution of the eastern Gulf of Aden conjugate margins from seismic reflection data

    NASA Astrophysics Data System (ADS)

    d'Acremont, Elia; Leroy, Sylvie; Beslier, Marie-Odile; Bellahsen, Nicolas; Fournier, Marc; Robin, Cécile; Maia, Marcia; Gente, Pascal

    2005-03-01

    The Gulf of Aden is a young and narrow oceanic basin formed in Oligo-Miocene time between the rifted margins of the Arabian and Somalian plates. Its mean orientation, N75°E, strikes obliquely (50°) to the N25°E opening direction. The western conjugate margins are masked by Oligo-Miocene lavas from the Afar Plume. This paper concerns the eastern margins, where the 19-35 Ma breakup structures are well exposed onshore and within the sediment-starved marine shelf. Those passive margins, about 200 km distant, are non-volcanic. Offshore, during the Encens-Sheba cruise we gathered swath bathymetry, single-channel seismic reflection, gravity and magnetism data, in order to compare the structure of the two conjugate margins and to reconstruct the evolution of the thinned continental crust from rifting to the onset of oceanic spreading. Between the Alula-Fartak and Socotra major fracture zones, two accommodation zones trending N25°E separate the margins into three N110°E-trending segments. The margins are asymmetric: offshore, the northern margin is narrower and steeper than the southern one. Including the onshore domain, the southern rifted margin is about twice the breadth of the northern one. We relate this asymmetry to inherited Jurassic/Cretaceous rifts. The rifting obliquity also influenced the syn-rift structural pattern responsible for the normal faults trending from N70°E to N110°E. The N110°E fault pattern could be explained by the decrease of the influence of rift obliquity towards the central rift, and/or by structural inheritance. The transition between the thinned continental crust and the oceanic crust is characterized by a 40 km wide zone. Our data suggest that its basement is made up of thinned continental crust along the southern margin and of thinned continental crust or exhumed mantle, more or less intruded by magmatic rocks, along the northern margin.

  16. Geology of the Plumtree area, Spruce Pine district, North Carolina

    USGS Publications Warehouse

    Brobst, Donald Albert

    1953-01-01

    This report describes the results of study and geologic mapping (1:12,000) in the 70-square-mile Plumtree area in the northeastern part of the Spruce Pine pegmatite district, on the Blue Ridge upland in western North Carolina. The district has been the chief domestic source of feldspar and sheet mica. The mining belt just west of the Blue Ridge Front trends northeast and is 25 miles long and 10 miles wide. The center of the Plumtree area lies 10 miles northeast of Spruce Pine pegmatite district, on the Blue Ridge upland in western North Carolina. The district has been the chief domestic source of feldspar and sheet mica. The mining belt just west of the Blue Ridge Front trends northeast and is 25 miles long and 10 miles wide. The center of the Plumtree area lies 10 miles northeast of Spruce Pine and includes parts of Mitchell and Avery Counties shown on the portions of the 7.5-minute Spruce Pine, Linville Falls, Newland, North Carolina, and Carvers Gap, North Carolina and Tennessee quadrangle. The topography varies from rugged mountains to rounded or flat topped hills near the entrenched, meandering master streams. Old erosion surfaces are approximately 600,1,100, 1,500, and 2,500 feet above the present master stream level. The area is in late youth or early maturity after rejuvenation.. The regionally metamorphosed rocks of the amophibolite facies form three mappable units: mica gneiss, mica schist, and hornblende rock. These rocks, perhaps of Precambrian age, are intimately interlayered with thicknesses of the individual layers ranging from less than one inch to several tons of feet. Field relationships and chemical data suggest that the mica (Carolina-type) rocks were derived from sandstones, graywackes, and shales and that the hornblende-rich (Roan-type) layers were derived from impure carbonate rocks. The igneous rocks include alaskite and associated pegmatite of early Paleozoic age (?), dunite and associated soapstone of a prepegmatite age, and a few diabasic

  17. 2-D Finite Difference Modeling of the D'' Structure Beneath the Eastern Cocos Plate: Part I

    NASA Astrophysics Data System (ADS)

    Helmberger, D. V.; Song, T. A.; Sun, D.

    2005-12-01

    The discovery of phase transition from Perovskite (Pv) to Post-Perovskite (PPv) at depth nears the lowermost mantle has revealed a new view of the earth's D'' layer (Oganov et al. 2004; Murakami et al. 2004). Hernlund et al. (2004) recently pusposed that, depending on the geotherm at the core-mantle boundary (CMB), a double-crossing of the phase boundary by the geotherm at two different depths may also occur. To explore these new findings, we adopt 2-D finite difference scheme (Helmberger and Vidale, 1988) to model wave propagation in rapidly varying structure. We collect broadband waveform data recorded by several Passcal experiments, such as La Ristra transect and CDROM transect in the southwest US to constrain the lateral variations in D'' structure. These data provide fairly dense sampling (~ 20 km) in the lowermost mantle beneath the eastern Cocos plate. Since the source-receiver paths are mostly in the same azimuth, we make 2-D cross-sections from global tomography model (Grand, 2002) and compute finite difference synthetics. We modify the lowermost mantle below 2500 km with constraints from transverse-component waveform data at epicentral distances of 70-82 degrees in the time window between S and ScS, essentially foward modeling waveforms. Assuming a velocity jump of 3 % at D'', our preferred model shows that the D'' topography deepens from the north to the south by about 120 km over a lateral distance of 300 km. Such large topography jumps have been proposed by Thomas et al. (2004) using data recorded by TriNet. In addition, there is a negative velocity jump (-3 %) 100 km above the CMB in the south. This simple model compare favorably with results from a study by Sun, Song and Helmberger (2005), who follow Sidorin et al. (1999) approach and produce a thermodynamically consistent velocity model with Pv-PPv phase boundary. It appears that much of this complexity exists in Grand's tomographic maps with rapid variation in velocities just above the D''. We also

  18. Population and Demographic Structure of Ixodes scapularis Say in the Eastern United States

    PubMed Central

    Sakamoto, Joyce M.; Goddard, Jerome; Rasgon, Jason L.

    2014-01-01

    Introduction The most significant vector of tick-borne pathogens in the United States is Ixodes scapularis Say (the blacklegged tick). Previous studies have identified significant genetic, behavioral and morphological differences between northern vs. southern populations of this tick. Because tick-borne pathogens are dependent on their vectors for transmission, a baseline understanding of the vector population structure is crucial to determining the risks and epidemiology of pathogen transmission. Methods We investigated population genetic variation of I. scapularis populations in the eastern United States using a multilocus approach. We sequenced and analyzed the mitochondrial COI and 16S genes and three nuclear genes (serpin2, ixoderin B and lysozyme) from wild specimens. Results We identified a deep divergence (3–7%) in I. scapularis COI gene sequences from some southern specimens, suggesting we had sampled a different Ixodes species. Analysis of mitochondrial 16S rRNA sequences did not support this hypothesis and indicated that all specimens were I. scapularis. Phylogenetic analysis and analysis of molecular variance (AMOVA) supported significant differences between northern vs. southern populations. Demographic analysis suggested that northern populations had experienced a bottleneck/expansion event sometime in the past, possibly associated with Pleistocene glaciation events. Conclusions Similar to other studies, our data support the division of northern vs. southern I. scapularis genetic lineages, likely due to differences in the demographic histories between these geographic regions. The deep divergence identified in some COI gene sequences highlights a potential hazard of relying solely on COI for species identification (“barcoding”) and population genetics in this important vector arthropod. PMID:25025532

  19. Orogenic structure of the Eastern Alps, Europe, from TRANSALP deep seismic reflection profiling

    NASA Astrophysics Data System (ADS)

    Lüschen, Ewald; Lammerer, Bernd; Gebrande, Helmut; Millahn, Karl; Nicolich, Rinaldo; Transalp Working Group

    2004-09-01

    The TRANSALP Group, comprising of partner institutions from Italy, Austria and Germany, acquired data on a 340 km long deep seismic reflection line crossing the Eastern Alps between Munich and Venice. Although the field work was split into four campaigns, between fall 1998 and summer 2001, the project gathered for the first time a continuous profile across the Alps using consistent field acquisition and data processing parameters. These sections span the orogen itself, at its broadest width, as well as the editor Fred Davey and the two adjacent basins. Vibroseis and explosion data, complementary in their depth penetration and resolution characteristics, were obtained along with wide-angle and teleseismic data. The profile shows a bi-vergent asymmetric structure of the crust beneath the Alpine axis which reaches a maximum thickness of 55 km, and 80-100 km long transcrustal ramps, the southward dipping 'Sub-Tauern-Ramp' and the northward-dipping 'Sub-Dolomites-Ramp'. Strongly reflective patterns of these ramps can be traced towards the north to the Inn Valley and towards the south to the Valsugana thrust belt, both of which show enhanced seismicity in the brittle upper crust. The seismic sections do not reveal any direct evidence for the presence of the Periadriatic Fault system, the presumed equivalent to the Insubric Line in the Western Alps. According to our new evolutionary model, the Sub-Tauern-Ramp is linked at depth with remnants of the subducted Penninic Ocean. The 'crocodile'-type model describes an upper/lower crustal decoupling and wedging of both the European and the Adriatic-African continents.

  20. Seismic velocity structure and seismotectonics of the eastern San Francisco Bay region, California

    USGS Publications Warehouse

    Hardebeck, J.L.; Michael, A.J.; Brocher, T.M.

    2007-01-01

    The Hayward Fault System is considered the most likely fault system in the San Francisco Bay Area, California, to produce a major earthquake in the next 30 years. To better understand this fault system, we use microseismicity to study its structure and kinematics. We present a new 3D seismic-velocity model for the eastern San Francisco Bay region, using microseismicity and controlled sources, which reveals a ???10% velocity contrast across the Hayward fault in the upper 10 km, with higher velocity in the Franciscan Complex to the west relative to the Great Valley Sequence to the east. This contrast is imaged more sharply in our localized model than in previous regional-scale models. Thick Cenozoic sedimentary basins, such as the Livermore basin, which may experience particularly strong shaking during an earthquake, are imaged in the model. The accurate earthquake locations and focal mechanisms obtained by using the 3D model allow us to study fault complexity and its implications for seismic hazard. The relocated hypocenters along the Hayward Fault in general are consistent with a near-vertical or steeply east-dipping fault zone. The southern Hayward fault merges smoothly with the Calaveras fault at depth, suggesting that large earthquakes may rupture across both faults. The use of the 3D velocity model reveals that most earthquakes along the Hayward fault have near-vertical strike-slip focal mechanisms, consistent with the large-scale orientation and sense of slip of the fault, with no evidence for zones of complex fracturing acting as barriers to earthquake rupture.

  1. Functional group biodiversity in Eastern Boundary Upwelling Ecosystems questions the wasp-waist trophic structure

    NASA Astrophysics Data System (ADS)

    Fréon, Pierre; Arístegui, Javier; Bertrand, Arnaud; Crawford, Robert J. M.; Field, John C.; Gibbons, Mark J.; Tam, Jorge; Hutchings, Larry; Masski, Hicham; Mullon, Christian; Ramdani, Mohamed; Seret, Bernard; Simier, Monique

    2009-12-01

    The species diversity of the four major Eastern Boundary Upwelling Ecosystems (EBUEs) is studied and compared with the aim of better understanding their functioning. Functional groups (FGs) of organisms were defined according to their taxonomy, body size and trophic level (TL), and span from plankton to top predators. Four large sub-divisions are defined in each system: two latitudinal sub-divisions (north and south) and two zonal sub-divisions (inshore and offshore), resulting in four sub-ecosystems per EBUE. A semi-quantitative approach is used in which only the dominant species (contributing 90% of overall biomass) are considered. EBUEs are compared in regard to their species composition, dominant species richness and evenness within FGs. The data are interpreted, focusing on latitudinal, zonal and depth gradients of diversity. Trophic flows (inflow and outflow) through the small pelagic fish FG are derived from different Ecopath models. This analysis of the four ecosystems and their sub-divisions does not provide support for the expected wasp-waist food web structure and functioning, with a single or several species of small pelagic fish primarily channelling the energy flow from lower to higher TL. Instead, similar low levels of richness were observed in many FGs of intermediate TL, allowing several energy transfer pathways. The gamma diversity is high due to the geographical distance between EBUEs and the presence or absence of rivers, but not to differences in their latitudinal position. The beta diversity is also high, due to the same factors plus the variation in shelf width and the contrast between inshore and offshore sub-divisions. The differences in richness and evenness among EBUEs are minor and do not explain the higher secondary and tertiary productivity of the Humboldt ecosystem.

  2. SPRUCE Discovery Manual, 169 Investigations Indoors and Outdoors.

    ERIC Educational Resources Information Center

    Busch, Phyllis S.

    Contained are instructional materials developed by the Science Project Related to Upgrading Conservation Education ("SPRUCE"). It is designed for use with the SPRUCE "Discovery Box" and contains twenty-one sets of investigations based on the twenty-one packets of specimens in the box; three sets are recommended for each of Grades K through 6. Each…

  3. Study of Reservoir Heterogencities and Structural Features Affecting Production in the Shallow Oil Zone, Eastern Elk Hills Area, California

    SciTech Connect

    Janice Gillespie

    2004-11-01

    Late Neogene (Plio-Pleistocene) shallow marine strata of the western Bakersfield Arch and Elk Hills produce hydrocarbons from several different reservoirs. This project focuses on the shallow marine deposits of the Gusher and Calitroleum reservoirs in the Lower Shallow Oil Zone (LSOZ). In the eastern part of the study area on the Bakersfield Arch at North and South Coles Levee field and in two wells in easternmost Elk Hills, the LSOZ reservoirs produce dry (predominantly methane) gas. In structurally higher locations in western Elk Hills, the LSOZ produces oil and associated gas. Gas analyses show that gas from the eastern LSOZ is bacterial and formed in place in the reservoirs, whereas gas associated with oil in the western part of the study area is thermogenic and migrated into the sands from deeper in the basin. Regional mapping shows that the gas-bearing LSOZ sands in the Coles Levee and easternmost Elk Hills area are sourced from the Sierra Nevada to the east whereas the oil-bearing sands in western Elk Hills appear to be sourced from the west. The eastern Elk Hills area occupied the basin depocenter, farthest from either source area. As a result, it collected mainly low-permeability offshore shale deposits. This sand-poor depocenter provides an effective barrier to the updip migration of gases from east to west. The role of small, listric normal faults as migration barriers is more ambiguous. Because our gas analyses show that the gas in the eastern LSOZ reservoirs is bacterial, it likely formed in-place near the reservoirs and did not have to migrate far. Therefore, the gas could have been generated after faulting and accumulated within the fault blocks as localized pools. However, bacterial gas is present in both the eastern AND western parts of Elk Hills in the Dry Gas Zone (DGZ) near the top of the stratigraphic section even though the measured fault displacement is greatest in this zone. Bacterial gas is not present in the west in the deeper LSOZ which

  4. Black carbon characterization in Quebec black spruce forests

    NASA Astrophysics Data System (ADS)

    Soucemarianadin, Laure; Quideau, Sylvie; Wasylishen, Roderick; MacKenzie, M. Derek

    2014-05-01

    Black carbon (BC), an important component of the global soil carbon pool, is a major by-product of wildfires in Quebec black spruce forests. However, BC characteristics vary depending on the environmental conditions under which it is formed and this may further affect its resistance to degradation. The objective of this study was to characterize the chemical and physical properties of BC formed under variable fire severity to assess its potential for recalcitrance as a passive carbon pool. Samples (n = 267) of BC produced by early season wildfires in 2005-2007 were collected from the surface of black spruce forest floors to cover the range of severity encountered in these fire-affected forests. Representative samples (n = 33) were then analyzed using elemental analysis, solid-state 13C nuclear magnetic resonance (NMR) spectroscopy, scanning electron microscopy and surface area analysis (BET method). Properties of BC sampled in the field were compared with those of samples produced under a range of controlled formation conditions in the laboratory. The NMR spectra of the BC collected on sites affected by low fire severity showed a distribution of total intensity between the different spectral regions very similar to those of unburned fuels, and were dominated by peaks indicative of cellulose, while spectra for BC from higher fire severity sites were dominated by a broad peak assigned to aromatic carbons. Atomic H/C and O/C ratios decreased along the fire severity gradient, confirming that increasing severity was associated with an increase in condensation. By comparing field- to laboratory-produced samples, we concluded that the temperature of formation in the field ranged between 75 and 250 ° C. In all analyzed BC samples, the fraction of aromatic carbon:total carbon was low, suggesting that the freshly produced BC in this boreal forest environment may be susceptible to rapid physical alteration and chemical degradation. Nevertheless, it is important to highlight

  5. Deep resistivity structure along the Longmen Mountain fault zone in the eastern Tibetan plateau of China

    NASA Astrophysics Data System (ADS)

    Wang, X.; Zhang, G.; Luo, W.; Luo, H.; Cai, X.; Zhou, Y.; Zhang, W.; Qin, Q.

    2012-12-01

    1.Introduction Many researchers (e.g.,Wang et al.,2009) have proposed the relevant knowledge of tectonic evolution and dynamic characteristics of the Longmen Mountain belt as well as the Songpan-Ganzi and Yangtze blocks in the past few decades, the knowledge of shallow thrust nappe tectonic along the belt has then been generally recognized. It's, however, still difficult to image the deep crust and mantle structures and reveal the dynamic mechanism of the crustal formation under the Longmen Mountain. In this study, we carried out the MT experiments along and across the Longmen mountain region and investigated the relationships between the crust structure and seismic activity basing on the latest MT geological results. 2. Field observations We conducted three MT experiment profiles in the eastern Tibetan Plateau. One is along the Mingshan-Guangyuan profile parallel with the structural direction, and another two profiles (Maqu-Gaoliangzhen and Luqu-Hechuan) perpendicular to the Longmen Mountain fault zone. In this study, we use the conventional magnetotelluric (MT) data combine with the long-period magnetotelluric (LMT) data to observe electromagnetic response. The MT and LMT data was observed by using the V8 instrument and LEMI-417, respectively. 3. Conclusion (1) According to the results of MT inversion, we find that the high concentration of stress process along the Songpan-Ganzi block and the Yangtze block colliding zone might result from the deep crust-mantle tough shear Zone of Longmen Mountain expanded to mid-upper crust, and finally leads to a new rupture. This could be one of the focal mechanisms of the Wenchuan earthquake (Ms 8.0) generating. The deep resistivity structure along the Longmen Mountain fault zone can be divided into southern,middle and northern segments from southwest to northeast. The total resistivity of southern segment is lower than the middle and northern portions. We suggest that the upper crust of the Longmen Mountain, south of Dayi

  6. Structures, microfabrics and textures of the Cordilleran-type Rechnitz metamorphic core complex, Eastern Alps☆

    PubMed Central

    Cao, Shuyun; Neubauer, Franz; Bernroider, Manfred; Liu, Junlai; Genser, Johann

    2013-01-01

    Rechnitz window group represents a Cordilleran-style metamorphic core complex, which is almost entirely located within nearly contemporaneous Neogene sediments at the transition zone between the Eastern Alps and the Neogene Pannonian basin. Two tectonic units are distinguished within the Rechnitz metamorphic core complex (RMCC): (1) a lower unit mainly composed of Mesozoic metasediments, and (2) an upper unit mainly composed of ophiolite remnants. Both units are metamorphosed within greenschist facies conditions during earliest Miocene followed by exhumation and cooling. The internal structure of the RMCC is characterized by the following succession of structure-forming events: (1) blueschist relics of Paleocene/Eocene age formed as a result of subduction (D1), (2) ductile nappe stacking (D2) of an ophiolite nappe over a distant passive margin succession (ca. E–W to WNW–ESE oriented stretching lineation), (3) greenschist facies-grade metamorphism annealing dominant in the lower unit, and (4) ductile low-angle normal faulting (D3) (with mainly NE–SW oriented stretching lineation), and (5) ca. E to NE-vergent folding (D4). The microfabrics are related to mostly ductile nappe stacking to ductile low-angle normal faulting. Paleopiezometry in conjunction with P–T estimates yield high strain rates of 10− 11 to 10− 13 s− 1, depending on the temperature (400–350 °C) and choice of piezometer and flow law calibration. Progressive microstructures and texture analysis indicate an overprint of the high-temperature fabrics (D2) by the low-temperature deformation (D3). Phengitic mica from the Paleocene/Eocene high-pressure metamorphism remained stable during D2 ductile deformation as well as preserved within late stages of final sub-greenschist facies shearing. Chlorite geothermometry yields two temperature groups, 376–328 °C, and 306–132 °C. Chlorite is seemingly accessible to late-stage resetting. The RMCC underwent an earlier large-scale coaxial

  7. Upper Mantle Discontinuity Structure Beneath the Western Atlantic Ocean and Eastern North America from SS Precursors

    NASA Astrophysics Data System (ADS)

    Schmerr, N. C.; Beghein, C.; Kostic, D.; Baldridge, A. M.; West, J. D.; Nittler, L. R.; Bull, A. L.; Montesi, L.; Byrne, P. K.; Hummer, D. R.; Plescia, J. B.; Elkins-Tanton, L. T.; Lekic, V.; Schmidt, B. E.; Elkins, L. J.; Cooper, C. M.; ten Kate, I. L.; Van Hinsbergen, D. J. J.; Parai, R.; Glass, J. B.; Ni, J.; Fuji, N.; McCubbin, F. M.; Michalski, J. R.; Zhao, C.; Arevalo, R. D., Jr.; Koelemeijer, P.; Courtier, A. M.; Dalton, H.; Waszek, L.; Bahamonde, J.; Schmerr, B.; Gilpin, N.; Rosenshein, E.; Mach, K.; Ostrach, L. R.; Caracas, R.; Craddock, R. A.; Moore-Driskell, M. M.; Du Frane, W. L.; Kellogg, L. H.

    2015-12-01

    Seismic discontinuities within the mantle arise from a wide range of mechanisms, including changes in mineralogy, major element composition, melt content, volatile abundance, anisotropy, or a combination of the above. In particular, the depth and sharpness of upper mantle discontinuities at 410 and 660 km depth are attributed to solid-state phase changes sensitive to both mantle temperature and composition, where regions of thermal heterogeneity produce topography and chemical heterogeneity changes the impedance contrast across the discontinuity. Seismic mapping of this topography and sharpness thus provides constraint on the thermal and compositional state of the mantle. The EarthScope USArray is providing unprecedented access to a wide variety of new regions previously undersampled by the SS precursors. This includes the boundary between the oceanic plate in the western Atlantic Ocean and continental margin of eastern North America. Here we use a seismic array approach to image the depth, sharpness, and topography of the upper mantle discontinuities, as well as other possible upper mantle reflectors beneath this region. This array approach utilizes seismic waves that reflect off the underside of a mantle discontinuity and arrive several hundred seconds prior to the SS seismic phase as precursory energy. In this study, we collected high-quality broadband data SS precursors data from shallow focus (< 30 km deep), mid-Atlantic ridge earthquakes recorded by USArray seismometers in Alaska. We generated 4th root vespagrams to enhance the SS precursors and determine how they sample the mantle. Our data show detection of localized structure on the discontinuity boundaries as well as additional horizons, such as the X-discontinuity and a potential reflection from a discontinuity near the depth of the lithosphere-asthenosphere boundary. These structures are related to the transition from predominantly old ocean lithosphere to underlying continental lithosphere, as while

  8. The Aerodynamic Properties of Spruce Twig Elements.

    NASA Astrophysics Data System (ADS)

    Grant, Richard Harrison

    1982-03-01

    The drag force and eddy shedding of three-pronged spruce twigs has been measured in a low speed wind tunnel for velocities ranging from 15 to 450 cm/s. The force on the twig at a given flow velocity is affected by the needle density of the twig and the twig orientation and flexibility. Based on the eddy shedding of the twig in 380 cm/s flow, the twig diameter is identified as the important length scale. Using the Strouhal relationship, the effective diameter of the twig is calculated to be approximately equal to the measured needle-tip to needle-tip twig diameter of a single-prong. Based on the twig diameter parallel to the flow, the experimental Re ranged from 200 to 12,000. The twig drag coefficient values suggest three flow regimes over the range of Re studied. These regimes are defined by the trend of the drag coefficient with increasing Re. The onset of the flow regimes is a function of the twig incidence with the regimes shifting up the Re scale with decreasing twig incidence angle. The twig drag response is compared and likened to that of a circular cylinder. In low Re flow the twig acts as a porous circular cylinder with the twig response primarily due to skin friction. In high Re flow the twig acts as a roughened solid circular cylinder with the twig response primarily due to form drag. The difference in the effective diameter and the measured diameter is defined to be the surface roughness. Based on the twig roughness, the critical Reynolds number for spruce twigs is calculated to be about 16,000 with a critical flow velocity of 10 m/s. Using the twig diameter length scale, the drag coefficients for spruce twigs and trees are compared providing a continuum of drag response for Reynolds 200 to 40,000. Based on the circular cylinder model of flow through and around and drag on the spruce twig, whole tree drag measurements can be explained.

  9. Flattening of the Greater Himalayan Zone within the Eastern Himalaya: Insights from Pressure-Temperature-Structural Distance Trends from Central and Eastern Bhutan

    NASA Astrophysics Data System (ADS)

    Agustsson, K. S.; Gordon, S. M.; Long, S. P.; Seward, G.; Zeiger, K. J.; Penfold, M. L.

    2014-12-01

    Quantifying pressure-temperature (P-T) trends across structural thickness provides insight into the tectonic mechanisms for emplacement and exhumation of orogen-scale ductile thrust sheets in active continental collision zones. Furthermore, these trends can reveal, via metamorphic discontinuities, the presence or absence of intra-thrust sheet structures. The Greater Himalayan zone (GHZ) contains mid-crustal rocks exhumed during Neogene convergence between India and Asia; however, the mechanisms that drove their emplacement and exhumation are debated. Within the Bhutan Himalaya, an intra-GHZ structure, the Kakhtang thrust (KT) is proposed to separate the GHZ into structurally upper and lower levels, and is interpreted to extend across much of the eastern Himalaya. In this investigation, peak P-T conditions were determined for rocks from two transects through both levels of the GHZ in north-central and northeastern Bhutan, in order to evaluate the significance of the KT, and to better understand the emplacement mechanism of GHZ rocks. The composition of the phases Grt + Pl + Bt ± Ms ± St ± Amp ± Kfs from twelve metapelite and orthogneiss samples were analyzed using the electron microprobe at UC-Santa Barbara. Peak P-T estimates, determined via THERMOCALC and supplemented by Ti-in-biotite thermometry, show similar T- but differing P versus structural distance trends across each transect. A gradual decrease in temperatures structurally upsection (~750 ºC to 600 ºC) was observed on both transects, and in northeast Bhutan, a steep decreasing pressure trend (0.40 ± 0.08 kbar/km) is documented. In comparison, in north-central Bhutan, a super-lithostatic pressure gradient (0.74 ± 0.16 kbar/km) is observed across >5.5 km of structural thickness, and is interpreted to represent >60 % layer-normal flattening strain, similar in magnitude to flattening documented in GHZ rocks in south-central Bhutan. The documented gradients show no significant change in P-T conditions

  10. Analyses of Great Smoky Mountain red spruce tree-ring data. Forest Service general technical report

    SciTech Connect

    Van Deusen, P.C.

    1988-05-01

    Contents include: Southern Appalachian Red Spruce--Fraser Fir Forests; A tree ring analysis of Red Spruce in the Southern Appalachian Mountains; Utilizing time series models and spatial analysis of forecast residuals for tree-ring analysis of Red Spruce; A fractal approach to analysis of tree-ring increments; Red Spruce Tree Ring Analysis using a Kalman Filter.

  11. Gravity field and structure of the Sorong Fault Zone, eastern Indonesia

    NASA Astrophysics Data System (ADS)

    Sardjono

    Gravity surveys along coastlines of islands in the region Banggai-Sula, Eastern Sulawesi, Halmahera, Bacan and Obi were carried out as part of the Sorong Fault Zone Project. Results of the Surveys were integrated with gravity data previously acquired by other projects, including on-land gravity data from the Bird Head area Irian Jaya (Dow et al 1986), Seram Island (Milsom 1977), Buru Island (Oemar and Reminton 1993) and Central Sulawesi (Silver et al. 1983) as well as marine gravity information within and surrounding the Sorong Fault Zone (Bowin et al. 1980). Gravity expeditions of the Sorong Fault Zone Project also include measurements in Mayu Island and the island group of Talaud, situated further north in the Central Molucca Sea region. A total of one hundred and forty two gravity data were acquired in the region of Banggai-Sula islands, forty seven in eastern part of Central Sulawesi, about four hundred in Halmahera, Bacan and Obi, and seventy nine in Mayu and Talaud. Surveys in the eastern part of Central Sulawesi were carried out for the purpose of tieing the older gravity data obtained from Silver et al. (1983) and the more recent data of the Sorong Fault Zone Project. About one thousand thirty hundred and thirty gravity data were acquired as part of the Irian Jaya Geological Mapping Project (IJGMP) in the period of 1978-1983, a project commissioned by the Indonesian Geological Research and Development Centre (GRDC) and the Australian Bureau of Mineral Resources (BMR). The remoteness of the survey areas of the Sorong Fault Zone Project necessitated a careful planning for travel arrangements and provision of logistics. A wide range of magnitude of gravity field was observed in the Sorong Fault Zone, extending from values below -250 mGal recorded in the southern part of the Molucca Sea to values in excess of +320 mGal measured near to sea level in the coastal areas south of Mangole and north of Sulabesi, the two islands of the Sula Group. Steep gradients of

  12. Kinematics, crustal structure and seismotectonics of the subducting northernmost Luzon arc in eastern Taiwan

    NASA Astrophysics Data System (ADS)

    Lee, Jian-Cheng; Chen, Chia-Yu; Chen, Rou-Fei; Chen, Yue-Gau; Chen, Horng-Yue

    2015-04-01

    The Coastal Range in eastern Taiwan belongs to the northernmost Luzon arc system, which sits on the western edge of the NW-moving Philippine Sea plates (PSP). As the PSP subducting and colliding with the Eurasian continental margin in Taiwan, the northern part of the Coastal Range provides a crucial key for better understand the geological structures and processes at tectonic transition from collision to subduction at plate corner. In this study, we conducted a dense network of GPS measurements at the northern tip of the Coastal Range and compiled available geological and geophysical information, including surface geomorphic features, geological structures, seismological data, seismic tomography, in order to provide insights on kinematics, crustal structure and seismotectonics of the transition from waning collision to subduction of the northern Luzon arc system and its vicinity. Regional short-term geodetic data, including GPS and levelling, and long-term thousand-year scale geological vertical rates indicate that the Coastal Range is going down toward the north, which we interpret it as being pulling down by the north- subduction PSP. Combing with the local GPS measurements carried out at the northern tip of the Longitudinal Valley, the plate suture, which shows a significant clockwise rotation at a rate of 33° M/yr, we interpret the tectonic escape of the northernmost Longitudinal Valley as being initiated locally by the northwest indentation of the Coastal Range, which pushed the northern Longitudinal Valley to move upward and eastward to form a 2-km-wide, 8-km-long, 100-m-high Milun Tableland. No significant deformation was observed across the surface trace of the major active Milun fault on the western side of the tableland, which has been ruptured during the 1951 M=7.1 Hualien earthquake, indicating that the Milun fault is now probably locked in the near surface. As for the crustal structure, we anticipate that there exists a fore-arc basement of the Luzon

  13. Structural Dimensions of Roma School Desegregation Policies in Central and Eastern Europe

    ERIC Educational Resources Information Center

    Rostas, Iulius; Kostka, Joanna

    2014-01-01

    Scrutiny of the socio-economic exclusion of the Roma in Central and Eastern Europe has brought attention to the widespread practice of school segregation of Romani children who are automatically placed in classes for the mentally disabled or shunted into separate and inferior schools and classrooms. It is now widely recognised that such practices…

  14. The Structural Disempowerment of Eastern European Migrant Farm Workers in Norwegian Agriculture

    ERIC Educational Resources Information Center

    Rye, Johan Fredrik; Andrzejewska, Joanna

    2010-01-01

    Since the 2004 EU enlargement established one European common labour market, a large number of Eastern Europeans have taken up seasonal employment as hired farm workers in Norwegian agriculture. Much attention in the public has been given to the potential for "social dumping" of these migrating workers, as they are considered prone to exploitation…

  15. Rift Structure along the Eastern Continental Margin of India - new constraints on style of breakup of the Indian landmass from the eastern Gondwanaland

    NASA Astrophysics Data System (ADS)

    Ismaiel, M.; Krishna, K. S.; Karlapati, S.; Mishra, J.; D, S.

    2015-12-01

    The Eastern Continental Margin of India (ECMI), a classical passive margin has evolved after breakup of the Indian landmass from the East Antarctica during the Early Cretaceous. Anomalous thick sediments and lack of cohesive magnetic signatures in the Bay of Bengal hampered delineation of rift-structure and age assignment for the continental breakup between India and East Antarctica. Further, absence of lithological and geochronological information and a few seismic profiles from the margin led to put forward several competing models for the rift initiation and evolution of the ECMI. Here, we analyze long streamer seismic reflection data and deep-water drill well information from the western Bay of Bengal to infer the buried rift structure, crustal architecture and stratigraphy along the ECMI. Following the structural pattern of the margin, the region is divided into four domains as decoupled, coupled, exhumed and oceanic, which in turn helped us to demarcate the variations in rift structure from south to north along the margin. The southern segment in the vicinity of Cauvery Basin consists of steep continental shelf associated with few major normal faults, which indicates that the segment was evolved as mix shear-rifted margin. The central segment off southern part of the Krishna-Godavari Basin is controlled by a series of fault-bounded half-graben structures and presence of thinned continental crust over the exhumed mantle body, revealing that the segment was formed under hyper-rifting process. While the northern segment extends up to Mahanadi Basin shows relatively less gradient continental slope with a few major faults, suggesting that the segment was evolved by hypo-extended process. Variable crustal architecture lying along the ECMI supports each segment of the margin formed in a specific rift process. A breakup unconformity considered as important geological constraint for completion of rift process between India and East Antarctica is clearly mapped on

  16. Stand structural diversity rather than species diversity enhances aboveground carbon storage in secondary subtropical forests in Eastern China

    NASA Astrophysics Data System (ADS)

    Ali, Arshad; Yan, En-Rong; Chen, Han Y. H.; Chang, Scott X.; Zhao, Yan-Tao; Yang, Xiao-Dong; Xu, Ming-Shan

    2016-08-01

    Stand structural diversity, typically characterized by variances in tree diameter at breast height (DBH) and total height, plays a critical role in influencing aboveground carbon (C) storage. However, few studies have considered the multivariate relationships of aboveground C storage with stand age, stand structural diversity, and species diversity in natural forests. In this study, aboveground C storage, stand age, tree species, DBH and height diversity indices, were determined across 80 subtropical forest plots in Eastern China. We employed structural equation modelling (SEM) to test for the direct and indirect effects of stand structural diversity, species diversity, and stand age on aboveground C storage. The three final SEMs with different directions for the path between species diversity and stand structural diversity had a similar goodness of fit to the data. They accounted for 82 % of the variation in aboveground C storage, 55-59 % of the variation in stand structural diversity, and 0.1 to 9 % of the variation in species diversity. Stand age demonstrated strong positive total effects, including a positive direct effect (β = 0.41), and a positive indirect effect via stand structural diversity (β = 0.41) on aboveground C storage. Stand structural diversity had a positive direct effect on aboveground C storage (β = 0.56), whereas there was little total effect of species diversity as it had a negative direct association with, but had a positive indirect effect, via stand structural diversity, on aboveground C storage. The negligible total effect of species diversity on aboveground C storage in the forests under study may have been attributable to competitive exclusion with high aboveground biomass, or a historical logging preference for productive species. Our analyses suggested that stand structural diversity was a major determinant for variations in aboveground C storage in the secondary subtropical forests in Eastern China. Hence, maintaining tree DBH and

  17. Low temperature thermochronology in the Eastern Alps: Implications for structural and topographic evolution

    PubMed Central

    Wölfler, Andreas; Stüwe, Kurt; Danišík, Martin; Evans, Noreen J.

    2012-01-01

    According to new apatite fission track, zircon- and apatite (U–Th)/He data, we constrain the near-surface history of the southeastern Tauern Window and adjacent Austrolapine units. The multi-system thermochronological data demonstrate that age-elevation correlations may lead to false implications about exhumation and cooling in the upper crust. We suggest that isothermal warping in the Penninic units that are in the position of a footwall, is due to uplift, erosion and the buildup of topography. Additionally we propose that exhumation rates in the Penninic units did not increase during the Middle Miocene, thus during the time of lateral extrusion. In contrast, exhumation rates of the Austroalpine hangingwall did increase from the Paleogene to the Neogene and the isotherms in this unit were not warped. The new zircon (U–Th)/He ages as well as zircon fission track ages from the literature document a Middle Miocene exhumation pulse which correlates with a period of enhanced sediment accumulation during that time. However, enhanced sedimentation- and exhumation rates at the Miocene/Pliocene boundary, as observed in the Western- and Central Alps, cannot be observed in the Eastern Alps. This contradicts a climatic trigger for surface uplift, and makes a tectonic trigger and/or deep-seated mechanism more obvious to explain surface uplift in the Eastern Alps. In combination with already published geochronological ages, our new data demonstrate Oligocene to Late Miocene fault activity along the Möll valley fault that constitutes a major shear zone in the Eastern Alps. In this context we suggest a geometrical and temporal relationship of the Katschberg-, Polinik–Möll valley- and Mur–Mürz faults that define the extruding wedge in the eastern part of the Eastern Alps. Equal deformation- and fission track cooling ages along the Katschberg–Brenner- and Simplon normal faults demonstrate overall Middle Miocene extension in the whole alpine arc. PMID:27065501

  18. Total OH reactivity emissions from Norway spruce

    NASA Astrophysics Data System (ADS)

    Nölscher, Anke; Bourtsoukidis, Efstratios; Bonn, Boris; Kesselmeier, Jürgen; Lelieveld, Jos; Williams, Jonathan

    2013-04-01

    Forest emissions represent a strong potential sink for the main tropospheric oxidant, the hydroxyl radical (OH). In forested environments, the comparison of the directly determined overall sink of OH radicals, the total OH reactivity, and the individually measured OH sink compounds often exposes a significant gap. This "missing" OH reactivity can be high and influenced by both direct biogenic emissions and secondary photo-oxidation products. To investigate the source of the missing OH sinks in forests, total OH reactivity emission rates were determined for the first time from a Norway spruce (Picea abies) throughout spring, summer and autumn 2011. The total OH reactivity was measured inside a branch enclosure using the Comparative Reactivity Method (CRM) with a Proton Transfer Reaction-Mass Spectrometer (PTR-MS) as the detector. In parallel, separate volatile organic compounds (VOC) emission rates were monitored by a second PTR-MS, including the signal of isoprene, acetaldehyde, total monoterpenes and total sesquiterpenes. The comparison of known and PTR-MS detected OH sink compounds and the directly measured total OH reactivity emitted from Norway spruce revealed unmeasured and possibly unknown primary biogenic emissions. These were found to be highest in late summer during daytime coincident with highest temperatures and ozone levels.

  19. [Richard Spruce, botanist-South America's explorer].

    PubMed

    Seaward, M R

    2000-01-01

    Between 1849 and 1864, the English botanist and explorer Richard Spruce carried out a detailed study of the Amazon flora and the costumes of the peoples who inhabited the region. To date a large part of the existing knowledge about several botanical families in the region stems from this scientist's efforts. His comprehensive interests, his detailed and precise descriptions were outstanding: nothing seems to have been left out of his scrutiny and recording aptitude. Not only was Spruce a remarkable botanist but he was also a distinctive anthropologist, linguist (he knew French, Spanish and Portuguese), geologist and geographer, as well as an acute sociological observer of the political systems and habits of the Amazonian and Andean trips in which has has been. He could thus make a considerable contribution to the understanding of indigenous beliefs and practices, as well as to the knowledge and uses of plants within the Amazonian context. Also important was his participation in the economic exploration of local species, particularly as regards the Hevea and the Cinchona genera.

  20. Genetic population structure of spotted seatrout Cynoscion nebulosus along the south-eastern U.S.A.

    PubMed

    O'Donnell, T P; Denson, M R; Darden, T L

    2014-08-01

    Analyses of the genetic population structure of spotted seatrout Cynoscion nebulosus along the south-eastern U.S. coast using 13 microsatellites suggest significant population differentiation between fish in North Carolina (NC) compared with South Carolina (SC) and Georgia (GA), with New River, NC, serving as an area of integration between northern and southern C. nebulosus. Although there is a significant break in gene flow between these areas, the overall pattern throughout the sampling range represents a gradient in genetic diversification with the degree of geographic separation. Latitudinal distance and estuarine density appear to be main drivers in the genetic differentiation of C. nebulosus along the south-eastern U.S. coast. The isolation-by-distance gene-flow pattern creates fine-scale differences in the genetic composition of proximal estuaries and dictates that stocking must be confined to within 100 km of the location of broodstock collection in order to maintain the natural gradient of genetic variation along the south-eastern U.S. coast.

  1. Analysis of time-structure of BSRs using 3D seismic data in the Eastern Nankai Trough, Japan

    NASA Astrophysics Data System (ADS)

    Nagakubo, S.; Inamori, T.; Kobayashi, T.; Fujii, T.

    2005-12-01

    According to the result of METI Exploratory Wells "Tokai-Oki to Kumano-nada" conducted in FY2003 in Japan, it is suggested that methane hydrate bearing layers in the Eastern Nanakai Trough distribute heterogeneously above BSRs (Bottom Simulating Reflectors). To understand the heterogeneity of distribution of methane hydrate bearing layers and explore concentrated hydrate bearing layers, we conducted a detailed analysis of time-structure map of BSR using 3D seismic survey data acquired in the Eastern Nankai Trough. Since P-wave velocity of hydrate bearing layers are high, it was expected that two-way-time from sea bottom to BSR is short above concentrated hydrate bearing layers compared to hydrate bearing zones. Although significant anomalies are recognized on time-structure map, it seems that anomalies are corresponding to heterogeneous thermal-structure in preference to distribution of hydrate bearing layers around surveyed area. It should be considered that these thermal anomalies are depending on fluid migration with hydrocarbons through faults, unconformities and permeable sand layers from deeper formations. Since occurrences of methane hydrates are strongly restricted by temperature and pressure, analysis of time-structure of BSR acquired by seismic data could be helpful to understand the accumulation mechanism of methane hydrates in sediments.

  2. Status of the spruce; Fir cooperative research program

    SciTech Connect

    Hertel, G.D.; Zarnoch, S.J.; Arre, T. ); Eager, C. ); Mohnen, V. ); MedLarz, S. )

    1987-01-01

    Aside from the mixed conifer forest in the San Bernadino National Forest near the Los Angeles Basin, the only significant visible decline and mortality of a U.S. forest possibly caused by regional air pollution is found in the high elevation spruce/fir forests of the Appalachians (VA, NC, TN, W VA), Adirondacks (NY), Green Mountains (VT), and the White Mountains (NH). In January, most of the scientists that have or are currently studying Spruce-Fir conditions met in Philadelphia. They came to a consensus on the regional condition of the Spruce-Fir forests. The results of that meeting are summarized.

  3. Crustal S-wave structure beneath Eastern Black Sea Region revealed by Rayleigh-wave group velocities

    NASA Astrophysics Data System (ADS)

    Çınar, Hakan; Alkan, Hamdi

    2016-01-01

    In this study, the crustal S-wave structure beneath the Eastern Black Sea Region (including the Eastern Black Sea Basin (EBSB) and Eastern Pontides (EP)) has been revealed using inversion of single-station, fundamental-mode Rayleigh-wave group velocities in the period range of 4-40 seconds. We used digital broadband recordings of 13 regional earthquakes that recently occurred in the easternmost EBSB recorded at stations of the Kandilli Observatory and Earthquake Research Institute (KOERI). The average group-velocity-dispersion curves were generated from 26 paths for the EBSB, and 16 paths for the EP, and they were inverted to determine the average 1-D shear-wave structure of the region. We have created a pseudo-section, roughly depicting the crustal structure of the region based on the group velocity inversion results of all station-earthquake paths. The thickness of the sedimentary layer reaches 12 km in the center of EBSB (Vs = 2.5-3.1 km/s) and decreases 4 km in the EP. There is a thin sedimentary layer in the EP (Vs = 2.7 km/s). A consolidated thin crust that exists in the EBSB possesses a high seismic velocity (Vs = 3.8 km/s). While a thin (∼26 km) and transitional crust exists beneath the EBSB, a thick (about 42 km) continental crust exists beneath the EP where the Conrad is clearly seen at about a 24 km depth. Thick continental crust in the EP region is clearly distinguished from a gradational velocity change (Vs = 3.4-3.8 km/s). The Moho dips approximately southwards, and the Vs velocity (4.25-4.15 km/s) beneath the Moho discontinuity decreases from the EBSB to the EP in the N-S direction. This may be an indication of a southward subduction.

  4. 20. GROVE OF TREES PINES, MULBERRY, JUNIPER, BLUE SPRUCE ...

    Library of Congress Historic Buildings Survey, Historic Engineering Record, Historic Landscapes Survey

    20. GROVE OF TREES -- PINES, MULBERRY, JUNIPER, BLUE SPRUCE -- TRANSPLANTED FROM NEW MEXICO MANZANO MOUNTAINS, WEST OF BUILDINGS 4 AND T-59, LOOKING NORTHWEST - U. S. Veterans Administration Medical Center, 2100 Ridgecrest Southeast, Albuquerque, Bernalillo County, NM

  5. 3. View from northeast corner, Canisteo and Spruce Streets. Photo ...

    Library of Congress Historic Buildings Survey, Historic Engineering Record, Historic Landscapes Survey

    3. View from northeast corner, Canisteo and Spruce Streets. Photo shows the garage area (Building #5) with sawtooth roofline and front elevation of Buildings #6 and #1. - Merrill Silk Mill, 233 Canisteo Street, Hornell, Steuben County, NY

  6. [Variability of Allozyme and cpSSR Markers in the Populations of Siberian Spruce].

    PubMed

    Ekart, A K; Semerikova, S A; Semerikov, V L; Larionova, A Ya; Kravchenko, A N; Dymshakova, O S

    2016-03-01

    The variability of 21 allozyme and three microsatellite loci of chloroplast DNA (cpDNA) was studied in the populations of Siberian spruce (Picea obovata Ledeb.) from Irkutsk oblast, Magadan oblast, Buryatia, and Mongolia. It was demonstrated that the highest level of genetic diversity among the examined populations at both allozyme and microsatellite loci was observed in the Tulyushka population from Irkutsk oblast. The lowest level of genetic diversity was observed in marginal isolated populations of Bogd Uul and Magadan. In the relict spruce population from Olkhon Island, differing from the other populations in the lowest allelic diversity of both types of markers, no expected decline of expected heterozygosity and haplotype diversity was observed. In this population, the variability parameters mentioned were close to the population mean. The obtained intrapopulation and intraspecific variability parameters of allozyme and microsatellite loci of chloroplast DNA and the data on the population differentiation at these loci indicate that the given markers can be used for the analysis of the population structure of Siberian spruce.

  7. STRUCTURAL AND HYDROGEOLOGIC APPLICATIONS OF REMOTE SENSING DATA, EASTERN YUCATAN PENINSULA, MEXICO.

    USGS Publications Warehouse

    Southworth, C. Scott; ,

    1984-01-01

    Landsat and Seasat satellite images and aerial photographs of eastern Yucatan Peninsula, Mexico, were analyzed to delineate geologic controls of ground water. Significant interpretation results include the delineation of linear topographic swales, interpreted as fractures, extending more than 50 km along strike from the previously known limit of the Holbox fracture system; the alignment of sink holes (cenotes) and inlets (caletas) on strike with existing faults and fracture systems; and the identification of tonal anomalies in Ingles Lagoon suggesting fresh-water discharge from a submarine spring.

  8. The Genetic Structure of Wild Orobanche cumana Wallr. (Orobanchaceae) Populations in Eastern Bulgaria Reflects Introgressions from Weedy Populations

    PubMed Central

    Pineda-Martos, Rocío; Pujadas-Salvà, Antonio J.; Fernández-Martínez, José M.; Stoyanov, Kiril; Pérez-Vich, Begoña

    2014-01-01

    Orobanche cumana is a holoparasitic plant naturally distributed from central Asia to south-eastern Europe, where it parasitizes wild Asteraceae species. It is also an important parasitic weed of sunflower crops. The objective of this research was to investigate genetic diversity, population structure, and virulence on sunflower of O. cumana populations parasitizing wild plants in eastern Bulgaria. Fresh tissue of eight O. cumana populations and mature seeds of four of them were collected in situ on wild hosts. Genetic diversity and population structure were studied with SSR markers and compared to weedy populations. Two main gene pools were identified in Bulgarian populations, with most of the populations having intermediate characteristics. Cross-inoculation experiments revealed that O. cumana populations collected on wild species possessed similar ability to parasitize sunflower to those collected on sunflower. The results were explained on the basis of an effective genetic exchange between populations parasitizing sunflower crops and those parasitizing wild species. The occurrence of bidirectional gene flow may have an impact on wild populations, as new physiological races continuously emerge in weedy populations. Also, genetic variability of wild populations may favour the ability of weedy populations to overcome sunflower resistance mechanisms. PMID:25143963

  9. Population structure, effective population size and adverse effects of stocking in the endangered Australian eastern freshwater cod Maccullochella ikei.

    PubMed

    Nock, C J; Ovenden, J R; Butler, G L; Wooden, I; Moore, A; Baverstock, P R

    2011-01-01

    Microsatellite markers were used to examine spatio-temporal genetic variation in the endangered eastern freshwater cod Maccullochella ikei in the Clarence River system, eastern Australia. High levels of population structure were detected. A model-based clustering analysis of multilocus genotypes identified four populations that were highly differentiated by F-statistics (F(ST) = 0·09 - 0·49; P < 0·05), suggesting fragmentation and restricted dispersal particularly among upstream sites. Hatchery breeding programmes were used to re-establish locally extirpated populations and to supplement remnant populations. Bayesian and frequency-based analyses of hatchery fingerling samples provided evidence for population admixture in the hatchery, with the majority of parental stock sourced from distinct upstream sites. Comparison between historical and contemporary wild-caught samples showed a significant loss of heterozygosity (21%) and allelic richness (24%) in the Mann and Nymboida Rivers since the commencement of stocking. Fragmentation may have been a causative factor; however, temporal shifts in allele frequencies suggest swamping with hatchery-produced M. ikei has contributed to the genetic decline in the largest wild population. This study demonstrates the importance of using information on genetic variation and population structure in the management of breeding and stocking programmes, particularly for threatened species. PMID:21235562

  10. Pulpability of beetle-killed spruce. Forest Service research paper

    SciTech Connect

    Scott, G.M.; Bormett, D.W.; Sutherland, N.R.; Abubakr, S.; Lowell, E.

    1996-08-01

    Infestation of the Dendroctonus rufipennis beetle has resulted in large stands of dead and dying timber on the Kenai Peninsula in Alaska. Tests were conducted to evaluate the value of beetle-killed spruce as pulpwood. The results showed that live and dead spruce wood can be pulped effectively. The two least deteriorated classes and the most deteriorated class of logs had similar characteristics when pulped; the remaining class had somewhat poorer pulpability.

  11. Eastern margin of the Ross Sea Rift in western Marie Byrd Land, Antarctica: Crustal structure and tectonic development

    NASA Astrophysics Data System (ADS)

    Luyendyk, Bruce P.; Wilson, Douglas S.; Siddoway, Christine S.

    2003-10-01

    The basement rock and structures of the Ross Sea rift are exposed in coastal western Marie Byrd Land (wMBL), West Antarctica. Thinned, extended continental crust forms wMBL and the eastern Ross Sea continental shelf, where faults control the regional basin-and range-type topography at ˜20 km spacing. Onshore in the Ford Ranges and Rockefeller Mountains of wMBL, basement rocks consist of Early Paleozoic metagreywacke and migmatized equivalents, intruded by Devonian-Carboniferous and Cretaceous granitoids. Marine geophysical profiles suggest that these geological formations continue offshore to the west beneath the eastern Ross Sea, and are covered by glacial and glacial marine sediments. Airborne gravity and radar soundings over wMBL indicate a thicker crust and smoother basement inland to the north and east of the northern Ford Ranges. A migmatite complex near this transition, exhumed from mid crustal depths between 100-94 Ma, suggests a profound crustal discontinuity near the inboard limit of extended crust, ˜300 km northeast of the eastern Ross Sea margin. Near this limit, aeromagnetic mapping reveals an extensive region of high amplitude anomalies east of the Ford ranges that can be interpreted as a sub ice volcanic province. Modeling of gravity data suggests that extended crust in the eastern Ross Sea and wMBL is 8-9 km thinner than interior MBL (β = 1.35). Gravity modeling also outlines extensive regions of low-density (2300-2500 kg m-3) buried basement rock that is lighter than rock exposed at the surface. These regions are interpreted as bounded by throughgoing east-west faults with vertical separation. These buried low-density rocks are possibly a low-density facies of Early Paleozoic metagreywacke, or the low-density epizonal facies of Cretaceous granites, or felsic volcanic rocks known from moraines. These geophysical features and structures on land in the wMBL region preserve the record of middle and Late Cretaceous development of the Ross Sea rift

  12. Labeling Feral Spruce Budworm (Lepidoptera: Tortricidae) Populations With Rubidium.

    PubMed

    MacKinnon, Wayne; Eveleigh, Eldon; Silk, Peter; Forbes, Glen

    2016-04-01

    Rubidium (Rb) is a trace element that occurs naturally in low concentrations and is easily absorbed by plants, making it a useful tool for labeling insect defoliators, such as spruce budworm (Choristoneura fumiferana Clemens). Balsam fir trees (Abies balsamea (L.) Miller) injected with either 8 or 16 g per tree of rubidium chloride (RbCl) showed quick uptake and distribution throughout the crown, with no negative effects on tree shoot growth or spruce budworm survival and development. Adult spruce budworm that fed as larvae on trees injected with RbCl were clearly labeled, with significantly higher Rb concentrations than the background levels found in adults that fed as larvae on control trees. Rb concentrations in feral spruce budworm adults for both the 8 g (9 µg/g) and 16 g (25 µg/g) per tree treatments were at least five times lower than those in laboratory-reared adults on 1,000 µg/g RbCl diet (125 µg/g); survival, development, pupal weight, sex ratio, and mating status of spruce budworm were not adversely affected by Rb treatment. Egg masses laid by feral females that fed as larvae on Rb-labeled trees were also labeled with Rb. Injecting trees with RbCl is a viable technique for labeling feral spruce budworm populations to help distinguish local populations from immigrants to better evaluate the success of early intervention strategies such as mating disruption. PMID:26920559

  13. Monoterpene emissions from bark beetle infested Engelmann spruce trees

    NASA Astrophysics Data System (ADS)

    Amin, Hardik S.; Russo, Rachel S.; Sive, Barkley; Richard Hoebeke, E.; Dodson, Craig; McCubbin, Ian B.; Gannet Hallar, A.; Huff Hartz, Kara E.

    2013-06-01

    Bark beetle infestation impacts the health of coniferous forests, which are an important source of volatile organic compounds (VOCs) to the atmosphere. The types and amounts of VOCs emitted from forests can influence secondary organic aerosol (SOA) formation and impact overall air quality. In this initial work, the impact of bark beetle infestation on SOA precursors from Engelmann spruce is assessed. The VOCs emitted from the trunk of infested and healthy spruce trees were sampled using both sorbent traps and evacuated canisters that were analyzed by gas chromatography/mass spectroscopy. The samples from the infested spruce tree suggest a nine-fold enhancement in the total VOC emissions. The dominant VOCs in the infested spruce trees were 3-carene, β-pinene, and α-pinene. The increase observed in VOCs sampled at the trunk of the infested spruce was consistent with increases observed at infested lodgepole pine trunks. However, the types and amounts of VOCs emitted from Engelmann spruce and lodgepole pine are different, which suggests that additional measures of VOC emissions are needed to characterize the impact of bark beetle infestation on VOC emissions and SOA precursors.

  14. Fluoroquinolone Resistance Mechanisms and population structure of Enterobacter cloacae non-susceptible to Ertapenem in North-Eastern France

    PubMed Central

    Guillard, Thomas; Cholley, Pascal; Limelette, Anne; Hocquet, Didier; Matton, Lucie; Guyeux, Christophe; Lebreil, Anne-Laure; Bajolet, Odile; Brasme, Lucien; Madoux, Janick; Vernet-Garnier, Véronique; Barbe, Coralie; Bertrand, Xavier; de Champs on behalf of CarbaFrEst Group, Christophe

    2015-01-01

    Fluoroquinolone (FQ) agents are a potential resort to treat infection due to Enterobacteriaceae producing extended spectrum β-lactamase and susceptible to FQ. In a context of increase of non-susceptibility to carbapenems among Enterobacteriaceae, we characterized FQ resistance mechanisms in 75 Enterobacter cloacae isolates non-susceptible to ertapenem in North-Eastern France in 2012 and describe the population structure by pulsed field gel electrophoresis (PFGE) and multi-locus sequence typing (MLST). Among them, 14.7% (12/75) carried a carbapenemase-encoding gene. Except one isolate producing VIM-1, the carbapenemase-producing isolates carried the well-known IncL/M pOXA48a plasmid. Most of the isolates (59/75) harbored at least a FQ-R determinant. qnr genes were predominant (40%, 30/75). The MLST study revealed that E. cloacae isolates’ clonality was wide [24 different sequence types (STs)]. The more widespread STs were ST74, ST101, ST110, ST114, and ST133. Carbapenem MICs were higher for E. cloacae ST74 than for other E. cloacae isolates. Plasmid-mediated quinolone resistance determinants were more often observed in E. cloacae ST74 isolates. These findings showed that (i) pOXA-48a is spreading in North-Eastern France, (ii) qnr is preponderant in E. cloacae, (iii) E. cloacae comprised a large amount of lineages spreading in North-Eastern France, and (iv) FQ as an alternative to β-lactams to treat ertapenem non-susceptible Enterobacteriaceae are compromised. PMID:26557115

  15. Negative impacts of high temperatures on growth of black spruce forests intensify with the anticipated climate warming.

    PubMed

    Girardin, Martin P; Hogg, Edward H; Bernier, Pierre Y; Kurz, Werner A; Guo, Xiao Jing; Cyr, Guillaume

    2016-02-01

    An increasing number of studies conclude that water limitations and heat stress may hinder the capacity of black spruce (Picea mariana (Mill.) B.S.P.) trees, a dominant species of Canada's boreal forests, to grow and assimilate atmospheric carbon. However, there is currently no scientific consensus on the future of these forests over the next century in the context of widespread climate warming. The large spatial extent of black spruce forests across the Canadian boreal forest and associated variability in climate, demography, and site conditions pose challenges for projecting future climate change responses. Here we provide an evaluation of the impacts of climate warming and drying, as well as increasing [CO2 ], on the aboveground productivity of black spruce forests across Canada south of 60°N for the period 1971 to 2100. We use a new extensive network of tree-ring data obtained from Canada's National Forest Inventory, spatially explicit simulations of net primary productivity (NPP) and its drivers, and multivariate statistical modeling. We found that soil water availability is a significant driver of black spruce interannual variability in productivity across broad areas of the western to eastern Canadian boreal forest. Interannual variability in productivity was also found to be driven by autotrophic respiration in the warmest regions. In most regions, the impacts of soil water availability and respiration on interannual variability in productivity occurred during the phase of carbohydrate accumulation the year preceding tree-ring formation. Results from projections suggest an increase in the importance of soil water availability and respiration as limiting factors on NPP over the next century due to warming, but this response may vary to the extent that other factors such as carbon dioxide fertilization, and respiration acclimation to high temperature, contribute to dampening these limitations. PMID:26507106

  16. Negative impacts of high temperatures on growth of black spruce forests intensify with the anticipated climate warming.

    PubMed

    Girardin, Martin P; Hogg, Edward H; Bernier, Pierre Y; Kurz, Werner A; Guo, Xiao Jing; Cyr, Guillaume

    2016-02-01

    An increasing number of studies conclude that water limitations and heat stress may hinder the capacity of black spruce (Picea mariana (Mill.) B.S.P.) trees, a dominant species of Canada's boreal forests, to grow and assimilate atmospheric carbon. However, there is currently no scientific consensus on the future of these forests over the next century in the context of widespread climate warming. The large spatial extent of black spruce forests across the Canadian boreal forest and associated variability in climate, demography, and site conditions pose challenges for projecting future climate change responses. Here we provide an evaluation of the impacts of climate warming and drying, as well as increasing [CO2 ], on the aboveground productivity of black spruce forests across Canada south of 60°N for the period 1971 to 2100. We use a new extensive network of tree-ring data obtained from Canada's National Forest Inventory, spatially explicit simulations of net primary productivity (NPP) and its drivers, and multivariate statistical modeling. We found that soil water availability is a significant driver of black spruce interannual variability in productivity across broad areas of the western to eastern Canadian boreal forest. Interannual variability in productivity was also found to be driven by autotrophic respiration in the warmest regions. In most regions, the impacts of soil water availability and respiration on interannual variability in productivity occurred during the phase of carbohydrate accumulation the year preceding tree-ring formation. Results from projections suggest an increase in the importance of soil water availability and respiration as limiting factors on NPP over the next century due to warming, but this response may vary to the extent that other factors such as carbon dioxide fertilization, and respiration acclimation to high temperature, contribute to dampening these limitations.

  17. Negative impacts of high temperatures on growth of black spruce forests intensify with the anticipated climate warming

    NASA Astrophysics Data System (ADS)

    Girardin, M. P.; Hogg, T.; Kurz, W.; Bernier, P. Y.; Guo, X. J.; Cyr, G.

    2015-12-01

    An increasing number of studies conclude that water limitations and heat stress may hinder the capacity of black spruce (Picea mariana (Mill.) B.S.P.) trees, a dominant species of Canada's boreal forests, to grow and assimilate atmospheric carbon. However, there is currently no scientific consensus on the future of these forests over the next century in the context of widespread climate warming. The large spatial extent of black spruce forests across the Canadian boreal forest and associated variability in climate, demography and site conditions pose challenges for projecting future climate change responses. Here we provide an evaluation of the impacts of climate warming and drying, as well as increasing [CO2], on the aboveground productivity of black spruce forests across Canada south of 60ºN for the period 1971 to 2100. We use a new extensive network of tree-ring data obtained from Canada's National Forest Inventory, spatially-explicit simulations of Net Primary Productivity (NPP) and its drivers, and multivariate statistical modelling. We found that soil water availability is a significant driver of black spruce inter-annual variability in productivity across broad areas of the western to eastern Canadian boreal forest. Inter-annual variability in productivity was also found to be driven by autotrophic respiration in the warmest regions. In most regions, the impacts of soil water availability and respiration on inter-annual variability in productivity occurred during the phase of carbohydrate accumulation the year preceding tree ring formation. Results from projections suggest an increase in the importance of soil water availability and respiration as limiting factors on NPP over the next century due to warming, but this response may vary to the extent that other factors such as carbon dioxide fertilization, and respiration acclimation to high temperature, contribute to dampening these limitations.

  18. Climate-growth relationships for bog-grown black spruce in northern Minnesota

    SciTech Connect

    Vogel, K.J. )

    1993-06-01

    Black spruce (Picea mariana) tree-ring chronologies were derived for three bogs in northern Minnesota. Standard chronologies were highly intercorrelated (0.72 to 0.87). The ring-width variability attributable to a common signal ranged from 38.6 to 56.8 percent which is large for closed canopy eastern forests. These chronologies exhibited great serial correlation, therefore all chronologies were autoregressively modelled prior to climatic analyses. Each chronology was compared to monthly temperature and precipitation data from a nearby weather station. Strengths of linear relationships were measured by the product-moment correlation coefficient. May and August temperatures from the previous year and March precipitation of the current year were significantly correlated with ring-width indices. These data suggest that tree-ring chronologies from mid-continental peatlands may be a valuable, though presently ignored, source of paleoclimatic data.

  19. Changes of the spruce forest stand aerodynamic properties during ten growing seasons

    NASA Astrophysics Data System (ADS)

    Hurtalova, T.; Matejka, F.; Janous, D.; Czerny, R.

    2009-04-01

    Objective of this study was to quantify the influence of a young spruce forest stand on airflow and its aerodynamic characteristics during ten growing seasons. With this aim the wind speed profiles measured in and above investigated spruce stand during growing seasons, from May to October, 1998-2007 were analysed. Experimental site is situated on a mild slope with SW orientation in the locality Bílý Kříž (49o30'17'' N, 18o32'28'' E, 898-908 m a.s.l.), which is in the highest part of the Moravian-Silesian Beskydy Mts, Czech Republic. The experimental site consisting of two plots Fd and Fs with different tree density is created by the monoculture of young Norway spruce stand (Picea abies L., Karst) with age of 17 years in 1998. Each of these plots has the area of 2500 m2, density of 2600 trees/ha in Fd plot and 2400 trees/ha in Fs plot in 1998, and gradually 1652 trees/ha (Fd) and 1428 trees/ha (Fs) in 2007. The aerodynamic characteristics can be described by the roughness length (z0) and the zero plane displacement (d). The presented study aims to analyse the changes in d and z0 values for a young spruce forest stand during ten consecutive growing seasons, and to relate the aerodynamic properties of an air layer affected by this stand to its growth parameters. It is known, that the local terrain and structure of forest stand influenced the direction and power of the airflow, as well as the structure of vertical wind speed profiles. From the wind speed profile analysis it follows, that the investigated spruce stand was in an aerodynamic unsteady state and then d and z0 values vary also with the wind speed. During investigated seasons the mean seasonal z0 values ranged between 0.48 m and 1.32 m in Fd and the corresponding values in Fs plot varied between 0.41 m and 1.36 m. The mean seasonal d values varied between 0.60h and 0.76h in Fd, and 61h and 0.76h in Fs, h is mean stand height.

  20. Structural characteristic and origin of intra-continental fold belt in the eastern Sichuan basin, South China Block

    NASA Astrophysics Data System (ADS)

    Li, Chuanxin; He, Dengfa; Sun, Yanpeng; He, Jinyou; Jiang, Zaixing

    2015-11-01

    The fold-and-thrust belt in the eastern Sichuan basin is represented by a series of subparallel chevron anticlines. Under the orogenic tectonic setting within the South China Block in Meso-Cenozoic period and the influence of the multi-layer detachment fault, the deformation of the thrust belt exhibits remarkably layered and large-scale intracontinental thrusting structural characteristics. In this paper, we focus on the structural geometry and deformational mechanisms using the latest two-dimensional (2D) and three-dimensional (3D) seismic reflection data in combination with well and outcrop data. The multi-layer detachment faults, especially the upper gypsum-bearing detachment in the Middle Triassic Jialingjiang Formation and lower detachment with gypsum or shale in the Lower-Middle Cambrian system, directly control the deformational styles of the study area. Interpretation of seismic sections indicates that the fold-and-thrust belt has various deformational styles during folding, including fault-propagation fold, fault-bend fold, and detachment fold with box-fold or pop-up structural geometry. Regional location and structural boundaries play significant roles in controlling the deformational styles, and distinct differences exist among the different anticlines. The Huayingshan anticline located at the front of the thrust belt shows intense structural deformation with northwestward thrusting direction and a relatively weak opposite southeastward thrusting. In addition, the anticlines exhibit structural differences along strike and the fold-and-thrust belt in the northern segment is influenced by the North China Block.

  1. Structural models of the Eastern Alps derived from geophysical data and their relation to tectonic processes (Invited)

    NASA Astrophysics Data System (ADS)

    Brueckl, E. P.; Behm, M.

    2009-12-01

    The closure of the Penninic Ocean, the collision of the Adriatic micro-plate with the European platform, the generation of the Pannonian basin in conjunction with the roll back of the Carpathian subduction, and lateral extrusion (gravitational collapse and tectonic escape) were the main tectonic processes which have been shaping the Eastern Alps since late Cretaceous. Recent large scale seismic refraction and wide angle reflection experiments (CELEBRATION 2000 and ALP 2002) completed and refined considerably the conception of the lithospheric structure of the Eastern Alps and their surrounding tectonic provinces (Bohemian Massif, Carpathians, Dinarides, and the Molasse, Pannonian and Adriatic basins). We review this data and refer to related results achieved by other authors and working groups (e.g. TRANSALP). Elastic plate modelling supports the interpretation of plate boundaries and gravity data provide additional constraints to our model of the lithosphere. An analysis of lower crustal and upper mantle wide angle reflections yields further information on tectonic processes. The structure of the Moho boundary and systematic differences of crustal seismic velocities and densities indicate clearly a fragmentation of the lithosphere into three blocks, Europe (EU), Adria (AD) and the Pannonian fragment (PA). These three blocks form a triple junction near the eastern termination of the Tauern window. West of the triple junction AD collides directly with EU in N-S direction. Thickening of the crustal basement is confined mainly to the orogenic core of the Eastern Alps and the Southern Alps. Cross sectional balancing indicates that the exhumation of the Tauern window has an eastward directed horizontal component in agreement with the lateral extrusion process. East of the triple junction, EU borders PA at an NE-ENE oriented boundary. Steeply subducting lithospheric slabs revealed by teleseismic tomography are located nearly vertically below the EU-AD and EU-PA Moho

  2. Scientists study deep geological structure between New Hebrides Arc and Eastern Australian Margin

    NASA Astrophysics Data System (ADS)

    Lafoy, Y.; Van de Beuque, S.; Bernardel, G.; Missegue, F.; Nercessian, A.; Auzende, J. M.; Symonds, P. A.; Exon, N. F.

    French and Australian scientists hope that deep seismic profiling performed earlier this year in the southwest Pacific will improve understanding of the geological framework, crustal characteristics, and evolution of basins and ridges between the eastern Australian margin and the New Hebrides Trench system. The data already have indicated that the basements of several basins are thinned and oceanic, that the basements of several ridges are continental, and that, contrary to previous thinking, the Loyalty Basin was formed in late Cretaceous time.Submerged continental fragments separated by deep ocean basins dominate the region in the west, and the Australian Plate is subducting beneath the New Hebrides Arc in the east. Subduction started in Late Miocene time [MacFarlane et al., 1988] at a rate of 12 cm/ year with a convergence direction of WSW-ENE [Dubois et al., 1977].

  3. Soil-solution chemistry in a low-elevation spruce-fir ecosystem, Howland, Maine

    USGS Publications Warehouse

    Fernandez, Ivan J.; Lawrence, Gregory B.; Son, Yowhan

    1995-01-01

    Soil solutions were collected monthly by tension and zero-tension lysimeters in a low-elevation red spruce stand in east-central Maine from May 1987 through December 1992. Soil solutions collected by Oa tension lysimeters had higher concentrations of most constituents than the Oa zero-tension lysimeters. In Oa horizon soil solutions growing season concentrations for SO4, Ca, and Mg averaged 57, 43, and 30 μmol L−1 in tension lysimeters, and 43, 28, and 19 μmol L−1 in zero-tension lysimeters, respectively. Because tension lysimeters remove water held by the soil at tensions up to 10 kPa, solutions are assumed to have more time to react with the soil compared to freely draining solutions collected by zero-tension lysimeters. Solutions collected in the Bs horizon by both types of collectors were similar which was attributed to the frequency of time periods when the water table was above the Bs lysimeters. Concentrations of SO4 and NO3 at this site were lower than concentrations reported for most other eastern U.S. spruce-fir sites, but base cation concentrations fell in the same range. Aluminum concentrations in this study were also lower than reported for other sites in the eastern U.S. and Ca/Al ratios did not suggest inhibition of Ca uptake by roots. Concentrations of SO4, Ca, K, and Cl decreased significantly in both the Oa and Bs horizons over the 56-month sampling period, which could reflect decreasing deposition rates for sulfur and base cations, climatic influences, or natural variation. A longer record of measured fluxes will be needed to adequately define temporal trends in solution chemistry and their causes.

  4. Blue whale population structure along the eastern South Pacific Ocean: evidence of more than one population.

    PubMed

    Torres-Florez, J P; Hucke-Gaete, R; LeDuc, R; Lang, A; Taylor, B; Pimper, L E; Bedriñana-Romano, L; Rosenbaum, H C; Figueroa, C C

    2014-12-01

    Blue whales (Balaenoptera musculus) were among the most intensively exploited species of whales in the world. As a consequence of this intense exploitation, blue whale sightings off the coast of Chile were uncommon by the end of the 20th century. In 2004, a feeding and nursing ground was reported in southern Chile (SCh). With the aim to investigate the genetic identity and relationship of these Chilean blue whales to those in other Southern Hemisphere areas, 60 biopsy samples were collected from blue whales in SCh between 2003 and 2009. These samples were genotyped at seven microsatellite loci and the mitochondrial control region was sequenced, allowing us to identify 52 individuals. To investigate the genetic identity of this suspected remnant population, we compared these 52 individuals to blue whales from Antarctica (ANT, n = 96), Northern Chile (NCh, n = 19) and the eastern tropical Pacific (ETP, n = 31). No significant differentiation in haplotype frequencies (mtDNA) or among genotypes (nDNA) was found between SCh, NCh and ETP, while significant differences were found between those three areas and Antarctica for both the mitochondrial and microsatellite analyses. Our results suggest at least two breeding population units or subspecies exist, which is also supported by other lines of evidence such as morphometrics and acoustics. The lack of differences detected between SCh/NCh/ETP areas supports the hypothesis that eastern South Pacific blue whales are using the ETP area as a possible breeding area. Considering the small population sizes previously reported for the SCh area, additional conservation measures and monitoring of this population should be developed and prioritized. PMID:25492593

  5. Blue whale population structure along the eastern South Pacific Ocean: evidence of more than one population.

    PubMed

    Torres-Florez, J P; Hucke-Gaete, R; LeDuc, R; Lang, A; Taylor, B; Pimper, L E; Bedriñana-Romano, L; Rosenbaum, H C; Figueroa, C C

    2014-12-01

    Blue whales (Balaenoptera musculus) were among the most intensively exploited species of whales in the world. As a consequence of this intense exploitation, blue whale sightings off the coast of Chile were uncommon by the end of the 20th century. In 2004, a feeding and nursing ground was reported in southern Chile (SCh). With the aim to investigate the genetic identity and relationship of these Chilean blue whales to those in other Southern Hemisphere areas, 60 biopsy samples were collected from blue whales in SCh between 2003 and 2009. These samples were genotyped at seven microsatellite loci and the mitochondrial control region was sequenced, allowing us to identify 52 individuals. To investigate the genetic identity of this suspected remnant population, we compared these 52 individuals to blue whales from Antarctica (ANT, n = 96), Northern Chile (NCh, n = 19) and the eastern tropical Pacific (ETP, n = 31). No significant differentiation in haplotype frequencies (mtDNA) or among genotypes (nDNA) was found between SCh, NCh and ETP, while significant differences were found between those three areas and Antarctica for both the mitochondrial and microsatellite analyses. Our results suggest at least two breeding population units or subspecies exist, which is also supported by other lines of evidence such as morphometrics and acoustics. The lack of differences detected between SCh/NCh/ETP areas supports the hypothesis that eastern South Pacific blue whales are using the ETP area as a possible breeding area. Considering the small population sizes previously reported for the SCh area, additional conservation measures and monitoring of this population should be developed and prioritized.

  6. Trophic structure and pathways of biogenic carbon flow in the eastern North Water Polynya

    NASA Astrophysics Data System (ADS)

    Tremblay, Jean-Éric; Hattori, Hiroshi; Michel, Christine; Ringuette, Marc; Mei, Zhi-Ping; Lovejoy, Connie; Fortier, Louis; Hobson, Keith A.; Amiel, David; Cochran, Kirk

    2006-10-01

    In the eastern North Water, most of the estimated annual new and net production of carbon (C) occurred during the main diatom bloom in 1998. During the bloom, at least 30% of total and new phytoplankton production occurred as dissolved organic carbon (DOC) and was unavailable for short-term assimilation into the herbivorous food web or sinking export. Based on particle interceptor traps and 234Th deficits, 27% of the particulate primary production (PP) sank out of the upper 50 m, with only 7% and 1% of PP reaching the benthos at shallow (≈200 m) and deep (≈500 m) sites, respectively. Mass balance calculations and grazing estimates agree that ≈79% of PP was ingested by pelagic consumers between April and July. During this period, the vertical flux of biogenic silica (BioSi) at 50 m was equivalent to the total BioSi produced, indicating that all of the diatom production was removed from the euphotic zone as intact cells (direct sinking) or empty frustules (grazing or lysis). The estimated flux of empty frustules was consistent with rates of herbivory by the large, dominant copepods and appendicularians during incubations. Since the carbon demand of the dominant planktivorous bird, Alle alle, amounted to ≈2% of the biomass synthesized by its main prey, the large copepod Calanus hyperboreus, most of the secondary carbon production was available to pelagic carnivores. Stable isotopes indicated that the biomass of predatory amphipods, polar cod and marine mammals was derived from these herbivores, but corresponding carbon fluxes were not quantified. Our analysis shows that a large fraction of PP in the eastern North Water was ingested by consumers in the upper 50 m, leading to substantial carbon respiration and DOC accumulation in surface waters. An increasingly early and prolonged opening of the Artic Ocean is likely to promote the productivity of the herbivorous food web, but not the short-term efficiency of the particulate, biological CO 2 pump.

  7. Canopy-Forming Seaweeds in Urchin-Dominated Systems in Eastern Canada: Structuring Forces or Simple Prey for Keystone Grazers?

    PubMed Central

    Blain, Caitlin; Gagnon, Patrick

    2014-01-01

    Models of benthic community dynamics for the extensively studied, shallow rocky ecosystems in eastern Canada emphasize kelp-urchin interactions. These models may bias the perception of factors and processes that structure communities, for they largely overlook the possible contribution of other seaweeds to ecosystem resilience. We examined the persistence of the annual, acidic (H2SO4), brown seaweed Desmarestia viridis in urchin barrens at two sites in Newfoundland (Canada) throughout an entire growth season (February to October). We also compared changes in epifaunal assemblages in D. viridis and other conspicuous canopy-forming seaweeds, the non-acidic conspecific Desmarestia aculeata and kelp Agarum clathratum. We show that D. viridis can form large canopies within the 2-to-8 m depth range that represent a transient community state termed “Desmarestia bed”. The annual resurgence of Desmarestia beds and continuous occurrence of D. aculeata and A. clathratum, create biological structure for major recruitment pulses in invertebrate and fish assemblages (e.g. from quasi-absent gastropods to >150 000 recruits kg−1 D. viridis). Many of these pulses phase with temperature-driven mass release of acid to the environment and die-off in D. viridis. We demonstrate experimentally that the chemical makeup of D. viridis and A. clathratum helps retard urchin grazing compared to D. aculeata and the highly consumed kelp Alaria esculenta. In light of our findings and related studies, we propose fundamental changes to the study of community shifts in shallow, rocky ecosystems in eastern Canada. In particular, we advocate the need to regard certain canopy-forming seaweeds as structuring forces interfering with top-down processes, rather than simple prey for keystone grazers. We also propose a novel, empirical model of ecological interactions for D. viridis. Overall, our study underscores the importance of studying organisms together with cross-scale environmental variability

  8. Canopy-forming seaweeds in urchin-dominated systems in eastern Canada: structuring forces or simple prey for keystone grazers?

    PubMed

    Blain, Caitlin; Gagnon, Patrick

    2014-01-01

    Models of benthic community dynamics for the extensively studied, shallow rocky ecosystems in eastern Canada emphasize kelp-urchin interactions. These models may bias the perception of factors and processes that structure communities, for they largely overlook the possible contribution of other seaweeds to ecosystem resilience. We examined the persistence of the annual, acidic (H2SO4), brown seaweed Desmarestia viridis in urchin barrens at two sites in Newfoundland (Canada) throughout an entire growth season (February to October). We also compared changes in epifaunal assemblages in D. viridis and other conspicuous canopy-forming seaweeds, the non-acidic conspecific Desmarestia aculeata and kelp Agarum clathratum. We show that D. viridis can form large canopies within the 2-to-8 m depth range that represent a transient community state termed "Desmarestia bed". The annual resurgence of Desmarestia beds and continuous occurrence of D. aculeata and A. clathratum, create biological structure for major recruitment pulses in invertebrate and fish assemblages (e.g. from quasi-absent gastropods to >150,000 recruits kg(-1) D. viridis). Many of these pulses phase with temperature-driven mass release of acid to the environment and die-off in D. viridis. We demonstrate experimentally that the chemical makeup of D. viridis and A. clathratum helps retard urchin grazing compared to D. aculeata and the highly consumed kelp Alaria esculenta. In light of our findings and related studies, we propose fundamental changes to the study of community shifts in shallow, rocky ecosystems in eastern Canada. In particular, we advocate the need to regard certain canopy-forming seaweeds as structuring forces interfering with top-down processes, rather than simple prey for keystone grazers. We also propose a novel, empirical model of ecological interactions for D. viridis. Overall, our study underscores the importance of studying organisms together with cross-scale environmental variability to

  9. Structure, Aboveground Biomass, and Soil Characterization of Avicennia marina in Eastern Mangrove Lagoon National Park, Abu Dhabi

    NASA Astrophysics Data System (ADS)

    Alsumaiti, Tareefa Saad Sultan

    Mangrove forests are national treasures of the United Arab Emirates (UAE) and other arid countries with limited forested areas. Mangroves form a crucial part of the coastal ecosystem and provide numerous benefits to society, economy, and especially the environment. Mangrove trees, specifically Avicennia marina, are studied in their native habitat in order to characterize their population structure, aboveground biomass, and soil properties. This study focused on Eastern Mangrove Lagoon National Park in Abu Dhabi, which was the first mangrove protected area to be designated in UAE. In situ measurements were collected to estimate Avicennia marina status, mortality rate (%), height (m), crown spread (m), stem number, diameter at breast height (cm), basal area (m), and aboveground biomass (t ha-1 ). Small-footprint aerial light detection and ranging (LIDAR) data acquired by UAE were processed to characterize mangrove canopy height and aboveground biomass density. This included extraction of LIDAR-derived height percentile statistics, segmentation of the forest into structurally homogenous units, and development of regression relationships between in situ reference and remote sensing data using a machine learning approach. An in situ soil survey was conducted to examine the soils' physical and chemical properties, fertility status, and organic matter. The data of soil survey were used to create soil maps to evaluate key characteristics of soils, and their influence on Avicennia marina in Eastern Mangrove Lagoon National Park. The results of this study provide new insights into Avicennia marina canopy population, structure, aboveground biomass, and soil properties in Abu Dhabi, as data in such arid environments is lacking. This valuable information can help in managing and preserving this unique ecosystem.

  10. Public health research support through the European structural funds in central and eastern Europe and the Mediterranean

    PubMed Central

    2012-01-01

    Background Public health research provides evidence for practice across fields including health care, health promotion and health surveillance. Levels of public health research vary markedly across European Union (EU) countries, and are lowest in the EU's new member states (in Central and Eastern Europe and the Mediterranean). However, these countries now receive most of the EU's Structural Funds, some of which are allocated to research. Methods STEPS, an EU-funded study, sought to assess support for public health research at national and European levels. To identify support through the Structural funds, STEPS drew information from country respondents and internet searches for all twelve EU new member states. Results The EU allocates annually around €7 billion through the Structural Funds for member states' own use on research. These funds can cover infrastructure, academic employment, and direct research grants. The programmes emphasise links to business. Support for health research includes major projects in biosciences, but direct support for public health research was found in only three countries - Cyprus, Latvia and Lithuania. Conclusions Public health research is not prioritised in the EU's Structural Funds programme in comparison with biomedicine. For the research dimension of the new European programme for Structural Funds 2014-2002, ministries of health should propose public health research to strengthen the evidence-base for European public health policy and practice. PMID:22480250

  11. Throughfall chemistry in a spruce chronosequence in southern Poland.

    PubMed

    Małek, Stanisław; Astel, Aleksander

    2008-10-01

    The chemical composition of throughfall and canopy leaching, as well as the acid neutralizing capacity and alkalinity depended on the age of Norway spruce (Picea abies Karst) stands and season of the year. A higher amount of sulphur and strong acids was deposited to the soil in the older age classes. Concentrations of SO(4)(2)(-), K(+), H(+), Mn(2+), Fe(2+) and Zn(2+) in throughfall were higher than in bulk precipitation in any season. This suggests that these ions were washed out or washed from the surface of needles and/or barks. The other ions NO(3)(-), NH(4)(+), Ca(2+) and Mg(2+) were retained by the canopy, in particular Ca(2+) and Mg(2+) during the growing season in young stands. Principal component analysis identified five factors responsible for the data structure and suggested the major anthropogenic emission sources were acidic emission (SO(4)(2)(-)+NO(3)(-)), heavy metals-dust particles (Fe(2+)+Mn(2+)+Zn(2+)), ammonium (NH(4)(+)) and H(+), while the natural-origin emission was mineral dust (Na(+)+K(+)+Ca(2+)+Mg(2+)).

  12. Investigations on spruce decline in the Bavarian forest.

    PubMed

    Osswald, W F; Elstner, E F

    1987-01-01

    The primary damaging reactions in spruce needles may operate as follows: 1) Trees under "stress" produce the plant hormone ethylene. 2) Ethylene and ozone react extremely fast forming hydrogen peroxide and formaldehyde, compounds which may damage the wax layer. 3) Ozone as a very aggressive oxidant will inactivate membrane bound enzymes through oxidation of their thiol groups. Thus the translocation of sugars from the chloroplast into the phloem may be inhibited or blocked. The result will be an "over-reduction" of the electron transport chain resulting in the formation of reactive oxygen species in the light. These reactive oxygen species will induce lipid peroxidation and pigment co-oxidation. 4) The visible effects are bleached needles and an impairment of structural resistance against fungal infections. 5) In addition ozone will directly reduce the content of antifungal compounds such as p-HAP. 6) Furthermore p-HAP may be involved in the bleaching reaction after its release from picein. 7) Finally, fungi may penetrate the needles and eventually grow faster in bleached needles. Infected needles will become necrotic and abscise.

  13. Population structure of guppies in north-eastern Venezuela, the area of putative incipient speciation

    PubMed Central

    2014-01-01

    Background Geographic barriers to gene flow and divergence among populations in sexual traits are two important causes of genetic isolation which may lead to speciation. Genetic isolation may be facilitated if these two mechanisms act synergistically. The guppy from the Cumaná region (within the Cariaco drainage) of eastern Venezuela has been previously described as a case of incipient speciation driven by sexual selection, significantly differentiated in sexual colouration and body shape from the common guppy, Poecilia reticulata. The latter occurs widely in northern Venezuela, including the south-eastern side of Cordillera de la Costa, where it inhabits streams belonging to the San Juan drainage. Here, we present molecular and morphological analyses of differentiation among guppy populations in the Cariaco and San Juan drainages. Our analyses are based on a 953 bp long mtDNA fragment, a set of 15 microsatellites (519 fish from 20 populations), and four phenotypic traits. Results Both microsatellite and mtDNA data showed that guppies inhabiting the two drainages are characterised by a significant genetic differentiation, but a higher proportion of the genetic variance was distributed among populations within regions. Most guppies in the Cariaco drainage had mtDNA from a distinct lineage, but we also found evidence for widespread introgression of mtDNA from the San Juan drainage into the Cariaco drainage. Phenotypically, populations in the two regions differed significantly only in the number of black crescents. Phenotypic clustering did not support existence of two distinct groupings, but indicated a degree of distinctiveness of Central Cumaná (CC) population. However, CC population showed little differentiation at the neutral markers from the proximate populations within the Cariaco drainage. Conclusions Our findings are consistent with only partial genetic isolation between the two geographic regions and indicate that the geographic barrier of Cordillera de

  14. Cretaceous to Miocene fault zone evolution in the Eastern Alps constrained by multi-system thermochronometry and structural data.

    NASA Astrophysics Data System (ADS)

    Wölfler, Andreas; Frisch, Wolfgang; Danišík, Martin; Fritz, Harald; Wölfler, Anke

    2015-04-01

    Fault zones that display both, ductile and brittle deformation stages offer perfect sites to study the evolution of the earth's crust over a wide range of temperatures and possibly over long time spans. This study combines structural- geo- and thermochronologcial data to evaluate the tectonic evolution of a fault zone to the southeast of the Tauern Window in the Eastern Alps. This fault zone comprises a mylonitic part, the so-called "Main Mylonitic Zone" (MMZ) that has been reworked by brittle faulting, the so-called "Ragga-Teuchl fault" (RTF). Structural data of the MMZ demonstrate ductile deformation with top-to-the NW transport in the Late Cretaceous under greenschist facies conditions. Subsequent SE-directed extension occurred under semi-brittle to brittle conditions during the Late Cretaceous and Paleocene. The Polinik Block to the north of the RTF revealed Late Cretaceous Ar/Ar ages, which reflect cooling subsequent to the thermal peak of Eo-alpine metamorphism. In contrast, the Kreuzeck Block to the south of the RTF shows early Permian Ar/Ar ages that reflect cooling related to both, late Variscan collapse in the late Carboniferous and post-Variscan extension in the Permian. Zircon and apatite fission track ages and thermal history modeling results suggest that the Polinik Block cooled rapidly to near surface temperatures in the middle Miocene. The Kreuzeck Block, in contrast, cooled and exhumed to near surface conditions already in the Oligocene and early Miocene. Thermal history modeling and apatite fission track ages of 23.3±0.8 and 11.5±1.0 suggest that brittle deformation along the RTF occurred in the middle- and late Miocene. Our results demonstrate that one single fault zone may comprise information about the evolution of the Eastern Alps from Late Cretaceous to Miocene time and that low-temperature thermochronology is a viable tool to resolve the timing of brittle faulting and accompanied fluid activity.

  15. Mother-offspring distances reflect sex differences in fine-scale genetic structure of eastern grey kangaroos.

    PubMed

    King, Wendy J; Garant, Dany; Festa-Bianchet, Marco

    2015-05-01

    Natal dispersal affects life history and population biology and causes gene flow. In mammals, dispersal is usually male-biased so that females tend to be philopatric and surrounded by matrilineal kin, which may lead to preferential associations among female kin. Here we combine genetic analyses and behavioral observations to investigate spatial genetic structure and sex-biased dispersal patterns in a high-density population of mammals showing fission-fusion group dynamics. We studied eastern grey kangaroos (Macropus giganteus) over 2 years at Wilsons Promontory National Park, Australia, and found weak fine-scale genetic structure among adult females in both years but no structure among adult males. Immature male kangaroos moved away from their mothers at 18-25 months of age, while immature females remained near their mothers until older. A higher proportion of male (34%) than female (6%) subadults and young adults were observed to disperse, although median distances of detected dispersals were similar for both sexes. Adult females had overlapping ranges that were far wider than the maximum extent of spatial genetic structure found. Female kangaroos, although weakly philopatric, mostly encounter nonrelatives in fission-fusion groups at high density, and therefore kinship is unlikely to strongly affect sociality.

  16. The Corossol structure: A possible impact crater on the seafloor of the northwestern Gulf of St. Lawrence, Eastern Canada

    NASA Astrophysics Data System (ADS)

    Lajeunesse, Patrick; St-Onge, Guillaume; Locat, Jacques; Duchesne, Mathieu J.; Higgins, Michael D.; Sanfaçon, Richard; Ortiz, Joseph

    2013-12-01

    We report on a 4.1 (±0.2) km diameter and 185 m deep circular submarine structure exposed on the seabed in >40 m water depths in the northwestern Gulf of St. Lawrence (Eastern Canada) from the analysis of high-resolution multibeam bathymetric and seismic data. The presence of a circular form characterized by a central uplift and concentric rings resembles the morphology and geometry of complex meteorite impact structures. Also, other origins, such as kimberlites, intrusions, karsts, or diapirs, can be eliminated on geological criteria. A single 4 cm long breccia fragment recovered from the central uplift has numerous glassy droplets of fluorapatite composition, assumed to be impact melts, and a single quartz grain with planar intersection features thought to be shock-induced planar deformation features (PDFs). The absolute age of this possible impact structure is unknown, but its geological setting indicates that it was formed long after the Mid-Ordovician and before regional pre-Quaternary sea-level lowstands. Present results outline the need for further examination to confirm an impact origin and to precisely date the formation of the structure.

  17. Mother–offspring distances reflect sex differences in fine-scale genetic structure of eastern grey kangaroos

    PubMed Central

    King, Wendy J; Garant, Dany; Festa-Bianchet, Marco

    2015-01-01

    Natal dispersal affects life history and population biology and causes gene flow. In mammals, dispersal is usually male-biased so that females tend to be philopatric and surrounded by matrilineal kin, which may lead to preferential associations among female kin. Here we combine genetic analyses and behavioral observations to investigate spatial genetic structure and sex-biased dispersal patterns in a high-density population of mammals showing fission–fusion group dynamics. We studied eastern grey kangaroos (Macropus giganteus) over 2 years at Wilsons Promontory National Park, Australia, and found weak fine-scale genetic structure among adult females in both years but no structure among adult males. Immature male kangaroos moved away from their mothers at 18–25 months of age, while immature females remained near their mothers until older. A higher proportion of male (34%) than female (6%) subadults and young adults were observed to disperse, although median distances of detected dispersals were similar for both sexes. Adult females had overlapping ranges that were far wider than the maximum extent of spatial genetic structure found. Female kangaroos, although weakly philopatric, mostly encounter nonrelatives in fission–fusion groups at high density, and therefore kinship is unlikely to strongly affect sociality. PMID:26045958

  18. Maps Showing Geology and Shallow Structure of Eastern Rhode Island Sound and Vineyard Sound, Massachusetts

    USGS Publications Warehouse

    O'Hara, Charles J.; Oldale, Robert N.

    1980-01-01

    This report presents results of marine studies conducted by the U.S. Geological Survey (USGS) during the summers of 1975 and 1976 in eastern Rhode Island Sound and Vineyard Sound (fig. 1) located off the southeastern coast of Massachusetts. The study was made in cooperation with the Massachusetts Department of Public Works and the New England Division of the U.S. Army Corps of Engineers. It covered an area of the Atlantic Inner Continental Shelf between latitude 41 deg 12' and 41 deg 33'N, and between longitude 70 deg 37' and 71 deg 15'W (see index map). Major objectives included assessment of sand and gravel resources, environmental impact evaluation both of offshore mining of these resources and of offshore disposal of solid waste and dredge spoil material, identification and mapping of the offshore geology, and determination of the geologic history of this part of the Inner Shelf. A total of 670 kilometers (km) of closely spaced high-resolution seismic-reflection profiles, 224 km of side-scan sonar data, and 16 cores totaling 90 meters (m) of recovered sediment, were collected during the investigation. This report is companion to geologic maps published for Cape Cod Bay (Oldale and O'Hara, 1975) and Buzzards Bay, Mass. (Robb and Oldale, 1977).

  19. Hydrography and population genetic structure in brook charr (Salvelinus fontinalis, Mitchill) from eastern Canada.

    PubMed

    Hébert, C; Danzman, R G; Jones, M W; Bernatchez, L

    2000-07-01

    Despite the abundance of studies of genetic diversity in freshwater fishes, few have specifically addressed the role of habitat structure in partitioning genetic variance within and among populations. In this study, we analysed the variability of six microsatellite loci among 24 brook charr population samples in order to correlate hydrographic structure with genetic organization. These populations originated from three Canadian National parks (Kouchibouguac, Fundy and Forillon) that showed distinct hydrographic structure. Considering the general characteristics of these habitats, we formulated specific hypotheses in regard to genetic structure, which were principally based on the potential for gene flow and population size associated with each habitat. The hierarchical analysis of molecular variance and the genetic distances computed among populations revealed that habitat structure analyses constitute an important, but insufficient, predictor of genetic structure. We discuss the importance of habitat complexity on genetic structure in the context of management and conservation.

  20. Decline of red spruce in the Adirondacks, New York

    SciTech Connect

    Scott, J.T.; Siccama, T.G.; Johnson, A.H.; Breisch, A.R.

    1984-01-01

    Thirty-two stands in the spruce-fir forests of Whiteface Mountain in the Adirondacks, originally sampled from 1964-66, were resurveyed in 1982. From 10-25 Bitterlich points were used in each stand in 1982 to obtain an estimate of basal area per hectare. Data were summarized for low elevation (<900m) and high elevation (> or = 900m) forests. Red spruce declined by 40-60% in basal area for the low elevation forests and by 60-70% above 900m. Balsam fir decreased by 35% at high elevations, due to natural disturbance in several of the stands, but was unchanged when only undisturbed stands were considered. The decline of red spruce accounted for about three quarters of the total decrease in basal area for both the high- and low-elevation forests. Spruce seedling frequency for the high-elevation sample decreased by 80%, but was unchanged below 900m. The pattern of spruce decline in the Adirondacks is similar to findings for New England. The cause of the decline is speculative at the time.

  1. Mathematical Methods of Modelling the Morphology of Spruce Trees

    NASA Astrophysics Data System (ADS)

    Janoutová, Růžena; Novotný, Jan; Pivovarník, Marek; Zemek, František

    2014-05-01

    Radiative transfer (RT) models are simulation tools which can be used to quantify relationships between vegetation canopy properties and observed remotely sensed data. This study aims at creating a spruce tree growth model as a key input for use in RT models. The spruce tree model is built on data obtained from terrestrial laser scanning of spruce trees. Each tree model is unique. This uniqueness is achieved by using L-systems which are able to simulate natural randomness while complying with the given tree parameters. L-systems are established on a theory of grammar that enables rewriting a string of symbols according to specified rewriting rules. In practice, our tree models are generated in Blender visualization software, implementing an algorithm written in Python. The algorithm generates the basic parameters of the whole tree and then creates the parameters of the spruce trunk and initial branches. The parameters are generated randomly within a range that is calculated from measured data. Then each branch is grown on the basis of annual increments defined by field measurements. Tree needles are distributed with respect to the age of individual branches; therefore, the needles have different colors according to their age. Cones and faces are graphical representations of the spruce model. Branches are represented by cones and needles are represented by faces around the branches. The faces are transparent, thus simulating light transmittance in-between the needles. The whole model is highly computationally demanding, especially with respect to computer memory.

  2. Experimental warming delays autumn senescence in a boreal spruce bog: Initial results from the SPRUCE experiment

    NASA Astrophysics Data System (ADS)

    Richardson, Andrew; Furze, Morgan; Aubrecht, Donald; Milliman, Thomas; Nettles, Robert; Krassovski, Misha; Hanson, Paul

    2016-04-01

    Phenology is considered one of the most robust indicators of the biological impacts of global change. In temperate and boreal regions, long-term data show that rising temperatures are advancing spring onset (e.g. budburst and flowering) and delaying autumn senescence (e.g. leaf coloration and leaf fall) in a wide range of ecosystems. While warm and cold temperatures, day length and insolation, precipitation and water availability, and other factors, have all been shown to influence plant phenology, the future response of phenology to rising temperatures and elevated CO2 still remains highly uncertain because of the challenges associated with conducting realistic manipulative experiments to simulate future environmental conditions. At the SPRUCE (Spruce and Peatland Responses Under Climatic and Environmental Change) experiment in the north-central United States, experimental temperature (0 to +9° C above ambient) and CO2 (ambient and elevated) treatments are being applied to mature, and intact, Picea mariana-Sphagnum spp. bog communities in their native habitat through the use of ten large (approximately 12 m wide, 10 m high) open-topped enclosures. We are tracking vegetation green-up and senescence in these chambers, at both the individual and whole-community level, using repeat digital photography. Within each chamber, digital camera images are recorded every 30 minutes and uploaded to the PhenoCam (http://phenocam.sr.unh.edu) project web page, where they are displayed in near-real-time. Image processing is conducted nightly to extract quantitative measures of canopy color, which we characterize using Gcc, the green chromatic coordinate. Data from a camera mounted outside the chambers (since November 2014) indicate strong seasonal variation in Gcc for both evergreen shrubs and trees. Shrub Gcc rises steeply in May and June, and declines steeply in September and October. By comparison, tree Gcc rises gradually from March through June, and declines gradually from

  3. Crustal structure of the eastern Qinling orogenic belt and implication for reactivation since the Cretaceous

    NASA Astrophysics Data System (ADS)

    Guo, Zhen; Chen, Y. John

    2016-06-01

    A high resolution crustal model of the eastern Qinling belt and central North China Craton (central NCC) is obtained along a N-S trending profile (corridor) by joint inversion of surface wave and receiver function. The NCC is one of the oldest cratons on Earth and the Qinling belt is the suture zone between the NCC and South China block (SCB). The Qinling belt is characterized by low crustal velocity (< 3.6 km/s) as well as low bulk Vp/Vs ratio (1.66-1.8), suggesting that the mid- to lower crust is predominantly felsic in composition, which could be the consequence of removal of mafic root by delamination in the past. The quartz-rich hence mechanically weak crust beneath the Qinling belt could be responsible for the strain focusing and significant reactivation since the Cretaceous. Beneath the central NCC, a ~ 10 km thick high-velocity layer (3.9-4.1 km/s) is observed just above the Moho, consistent with the regional high bulk Vp/Vs ratio (> 1.8). The forward gravity modeling supports the presence of a high-density layer (3.05 g/cm3) at the base of the crust beneath the central NCC. We propose that the high velocity in the lowermost crust beneath the central NCC is most likely due to the repeated mafic underplating, which also results in high crustal Vp/Vs ratio and is responsible for the rapid crustal uplift during the late Mesozoic.

  4. Unusual lithospheric structure beneath the Hyderabad granitic region, eastern Dharwar craton, south India

    NASA Astrophysics Data System (ADS)

    Pandey, O. P.; Agrawal, P. K.; Chetty, T. R. K.

    2002-03-01

    Using multiparametric geological and geophysical data, the evolutionary history of the lithosphere beneath the Late Archean—early Proterozoic Hyderabad granitic region (HGR) of the eastern Dharwar craton of south Indian shield has been attempted. Our study reveals that the entire granitic region and the surroundings (˜200 km×200 km) is being neotectonically uplifted possibly due to a major intrusive body situated at sub-crustal depth, leading to changes in river courses and also erosion of several kilometers (˜10 km) from its upper crustal column. Beneath this region, the Moho is elevated (32-33 km), the reduced heat flow is of the order of 28 mW/m 2 and the asthenosphere is located at shallow depths of about 124 km, i.e. far less than 200-400 km depth found in Precambrian shield areas of the globe. The cratonic mantle lithosphere beneath HGR contains a highly conductive (15 Ωm) hydrous and seismically anisotropic metasomatic zone between depth of 90 and 100 km, where the estimated temperatures could be in the range of 900-1000 °C. The surface granitic layer, containing unusually high radioactivity (5.25 μW/m 3), appears very thin probably a kilometer or so, beneath which the entire crustal column seems to be differentiated and made up of low radioactive granulite facies (?) rocks of acid to intermediate composition. Persistent episodic thermal reactivations during the last 2.6 gega years and continuous uplifting seems to have played a major role towards making its crustal column unusual.

  5. Influence of chemosynthetic ecosystems on nematode community structure and biomass in the deep eastern Mediterranean Sea

    NASA Astrophysics Data System (ADS)

    Lampadariou, N.; Kalogeropoulou, V.; Sevastou, K.; Keklikoglou, K.; Sarrazin, J.

    2013-08-01

    Mud volcanoes are a~special type of cold seeps where life is based on chemoautotrophic processes. They are considered to be extreme environments and are characterized by unique megafaunal and macrofaunal communities. However, very few studies on mud volcanoes taking into account the smaller meiobenthic communities have been carried out. Two mud volcanoes were explored during the MEDECO (MEditerranean Deep-sea ECOsystems) cruise (2007) with the remotely operated vehicle (ROV) Victor-6000: Amsterdam, located south of Turkey between 1700 and 2000 m depth (Anaximander mud field); and Napoli, south of Crete, located along the Mediterranean Ridge at about 2000 m depth (Olimpi mud field). The major aim of this study was to describe distributional patterns of meiofaunal communities and nematode assemblages from different seep microhabitats. Meiofaunal taxa and nematode assemblages at both mud volcanoes differed significantly from other Mediterranean sites in terms of standing stocks, dominance and species diversity. Density and biomass values were significantly higher at the seep sites, particularly at Amsterdam. Patterns of nematode diversity, the dominant meiofaunal taxon, varied, displaying both very high or very low species richness and dominance, depending on the microhabitat studied. The periphery of the Lamellibrachia and bivalve shell microhabitats of Napoli exhibited the highest species richness, while the reduced sediments of Amsterdam yielded a species-poor nematode community dominated by two successful species, one belonging to the genus Aponema and the other to the genus Sabatieria. Analysis of β-diversity showed that microhabitat heterogeneity of mud volcanoes contributed substantially to the total nematode species richness in the eastern Mediterranean Sea. These observations indicate a strong influence of mud volcanoes and cold-seep ecosystems on the meiofaunal communities and nematode assemblages.

  6. Influence of chemosynthetic ecosystems on nematode community structure and biomass in the deep eastern Mediterranean Sea

    NASA Astrophysics Data System (ADS)

    Lampadariou, N.; Kalogeropoulou, V.; Sevastou, K.; Keklikoglou, K.; Sarrazin, J.

    2012-12-01

    Mud volcanoes are a special type of cold seeps where life is based on chemoautotrophic processes. They are considered as extreme environments and are characterised by unique megafaunal and macrofaunal communities. However, very few studies on mud volcanoes taking into account the smaller meiobenthic communities have been carried out. Two mud volcanoes were explored during the MEDECO cruise (2007) with the Remotely Operated Vehicle (ROV) Victor-6000; Amsterdam, located south of Turkey between 1700 and 2000 m depth (Anaximander mud field) and Napoli, south of Crete, located along the Mediterranean Ridge at about 2000 m depth (Olimpi mud field). The major aim of this study was to describe distributional patterns of meiofaunal communities and nematode assemblages from different seep microhabitats. Meiofaunal taxa and nematode assemblages at both mud volcanoes differed significantly from other Mediterranean sites in terms of standing stocks, dominance and species diversity. Density and biomass values were significantly higher at the seep sites, particularly at Amsterdam. Nematodes, the dominant meiofaunal taxon, displayed deeper penetration vertically into the sediment at the seep areas, indicating that biological rather than physicochemical factors are responsible for their vertical distribution. Patterns of nematode diversity varied, displaying both very high or very low species richness and dominance, depending on the habitat studied. The Lamellibrachia periphery and mussel bed of Napoli exhibited the highest species richness while the reduced sediments of Amsterdam yielded a species-poor nematode community, dominated by two successful species; one belonging to the genus Aponema and the other to the genus Sabatieria. Analysis of β-diversity showed that habitat heterogeneity of mud volcanoes contributed substantially to the total nematode species richness in the eastern Mediterranean Sea. These observations indicate a strong influence of mud volcanoes and cold

  7. Multi-scale variation in spatial heterogeneity for microbial community structure in an eastern Virginia agricultural field

    NASA Technical Reports Server (NTRS)

    Franklin, Rima B.; Mills, Aaron L.

    2003-01-01

    To better understand the distribution of soil microbial communities at multiple spatial scales, a survey was conducted to examine the spatial organization of community structure in a wheat field in eastern Virginia (USA). Nearly 200 soil samples were collected at a variety of separation distances ranging from 2.5 cm to 11 m. Whole-community DNA was extracted from each sample, and community structure was compared using amplified fragment length polymorphism (AFLP) DNA fingerprinting. Relative similarity was calculated between each pair of samples and compared using geostatistical variogram analysis to study autocorrelation as a function of separation distance. Spatial autocorrelation was found at scales ranging from 30 cm to more than 6 m, depending on the sampling extent considered. In some locations, up to four different correlation length scales were detected. The presence of nested scales of variability suggests that the environmental factors regulating the development of the communities in this soil may operate at different scales. Kriging was used to generate maps of the spatial organization of communities across the plot, and the results demonstrated that bacterial distributions can be highly structured, even within a habitat that appears relatively homogeneous at the plot and field scale. Different subsets of the microbial community were distributed differently across the plot, and this is thought to be due to the variable response of individual populations to spatial heterogeneity associated with soil properties. c2003 Federation of European Microbiological Societies. Published by Elsevier Science B.V. All rights reserved.

  8. Effect of thinning on anatomical adaptations of Norway spruce needles.

    PubMed

    Gebauer, Roman; Volarík, Daniel; Urban, Josef; Børja, Isabella; Nagy, Nina Elisabeth; Eldhuset, Toril Drabløs; Krokene, Paal

    2011-10-01

    Conifers and other trees are constantly adapting to changes in light conditions, water/nutrient supply and temperatures by physiological and morphological modifications of their foliage. However, the relationship between physiological processes and anatomical characteristics of foliage has been little explored in trees. In this study we evaluated needle structure and function in Norway spruce families exposed to different light conditions and transpiration regimes. We compared needle characteristics of sun-exposed and shaded current-year needles in a control plot and a thinned plot with 50% reduction in stand density. Whole-tree transpiration rates remained similar across plots, but increased transpiration of lower branches after thinning implies that sun-exposed needles in the thinned plot were subjected to higher water stress than sun-exposed needles in the control plot. In general, morphological and anatomical needle parameters increased with increasing tree height and light intensity. Needle width, needle cross-section area, needle stele area and needle flatness (the ratio of needle thickness to needle width) differed most between the upper and lower canopy. The parameters that were most sensitive to the altered needle water status of the upper canopy after thinning were needle thickness, needle flatness and percentage of stele area in needle area. These results show that studies comparing needle structure or function between tree species should consider not only tree height and light gradients, but also needle water status. Unaccounted for differences in needle water status may have contributed to the variable relationship between needle structure and irradiance that has been observed among conifers. PMID:21891783

  9. Possible red spruce decline: Contributions of tree-ring analysis

    SciTech Connect

    Van Deusen, P.C. ); Reams, G.A. ); Cook, E.R. )

    1991-01-01

    Debate continues about the cause of apparent unprecedented decreases in ring width at all elevations, and increasing levels of mortality at high elevations, in red spruce (Picea rubens) stands in the northeastern United States. These growth and mortality trends are often used as evidence of red spruce decline, but the possibility remains that they may be occurring naturally. Two hypotheses are being used to explain the causes of red spruce growth reduction across its range and increased levels of standing dead at some high-elevation sites. This article summarizes the basic evidence used by advocates of these hypotheses and discusses the strengths of their arguments. The information presented is based primarily on tree-ring studies sponsored by the Forest Response Program, which is part of the National Acid Precipitation Assessment Program.

  10. Impact of Forest Harvesting on Trophic Structure of Eastern Canadian Boreal Shield Lakes: Insights from Stable Isotope Analyses

    PubMed Central

    Glaz, Patricia; Sirois, Pascal; Archambault, Philippe; Nozais, Christian

    2014-01-01

    Perturbations on ecosystems can have profound immediate effects and can, accordingly, greatly alter the natural community. Land-use such as forestry activities in the Canadian Boreal region have increased in the last decades, raising concerns about their potential impact on aquatic ecosystems. The objective of this study was to evaluate the impact of forest harvesting on trophic structure in eastern Canadian Boreal Shield lakes. We measured carbon and nitrogen stable isotopes values for aquatic primary producers, terrestrial detritus, benthic macroinvertebrates, zooplankton and brook trout (Salvelinus fontinalis) over a three-year period in eight eastern Boreal Shield lakes. Four lakes were studied before, one and two years after forest harvesting (perturbed lakes) and compared with four undisturbed reference lakes (unperturbed lakes) sampled at the same time. Stable isotope mixing models showed leaf-litter to be the main food source for benthic primary consumers in both perturbed and unperturbed lakes, suggesting no logging impact on allochthonous subsidies to the littoral food web. Brook trout derived their food mainly from benthic predatory macroinvertebrates in unperturbed lakes. However, in perturbed lakes one year after harvesting, zooplankton appeared to be the main contributor to brook trout diet. This change in brook trout diet was mitigated two years after harvesting. Size-related diet shift were also observed for brook trout, indicating a diet shift related to size. Our study suggests that carbon from terrestrial habitat may be a significant contribution to the food web of oligotrophic Canadian Boreal Shield lakes. Forest harvesting did not have an impact on the diet of benthic primary consumers. On the other hand, brook trout diet composition was affected by logging with greater zooplankton contribution in perturbed lakes, possibly induced by darker-colored environment in these lakes one year after logging. PMID:24763366

  11. Impact of forest harvesting on trophic structure of eastern Canadian Boreal Shield lakes: insights from stable isotope analyses.

    PubMed

    Glaz, Patricia; Sirois, Pascal; Archambault, Philippe; Nozais, Christian

    2014-01-01

    Perturbations on ecosystems can have profound immediate effects and can, accordingly, greatly alter the natural community. Land-use such as forestry activities in the Canadian Boreal region have increased in the last decades, raising concerns about their potential impact on aquatic ecosystems. The objective of this study was to evaluate the impact of forest harvesting on trophic structure in eastern Canadian Boreal Shield lakes. We measured carbon and nitrogen stable isotopes values for aquatic primary producers, terrestrial detritus, benthic macroinvertebrates, zooplankton and brook trout (Salvelinus fontinalis) over a three-year period in eight eastern Boreal Shield lakes. Four lakes were studied before, one and two years after forest harvesting (perturbed lakes) and compared with four undisturbed reference lakes (unperturbed lakes) sampled at the same time. Stable isotope mixing models showed leaf-litter to be the main food source for benthic primary consumers in both perturbed and unperturbed lakes, suggesting no logging impact on allochthonous subsidies to the littoral food web. Brook trout derived their food mainly from benthic predatory macroinvertebrates in unperturbed lakes. However, in perturbed lakes one year after harvesting, zooplankton appeared to be the main contributor to brook trout diet. This change in brook trout diet was mitigated two years after harvesting. Size-related diet shift were also observed for brook trout, indicating a diet shift related to size. Our study suggests that carbon from terrestrial habitat may be a significant contribution to the food web of oligotrophic Canadian Boreal Shield lakes. Forest harvesting did not have an impact on the diet of benthic primary consumers. On the other hand, brook trout diet composition was affected by logging with greater zooplankton contribution in perturbed lakes, possibly induced by darker-colored environment in these lakes one year after logging. PMID:24763366

  12. Impact of forest harvesting on trophic structure of eastern Canadian Boreal Shield lakes: insights from stable isotope analyses.

    PubMed

    Glaz, Patricia; Sirois, Pascal; Archambault, Philippe; Nozais, Christian

    2014-01-01

    Perturbations on ecosystems can have profound immediate effects and can, accordingly, greatly alter the natural community. Land-use such as forestry activities in the Canadian Boreal region have increased in the last decades, raising concerns about their potential impact on aquatic ecosystems. The objective of this study was to evaluate the impact of forest harvesting on trophic structure in eastern Canadian Boreal Shield lakes. We measured carbon and nitrogen stable isotopes values for aquatic primary producers, terrestrial detritus, benthic macroinvertebrates, zooplankton and brook trout (Salvelinus fontinalis) over a three-year period in eight eastern Boreal Shield lakes. Four lakes were studied before, one and two years after forest harvesting (perturbed lakes) and compared with four undisturbed reference lakes (unperturbed lakes) sampled at the same time. Stable isotope mixing models showed leaf-litter to be the main food source for benthic primary consumers in both perturbed and unperturbed lakes, suggesting no logging impact on allochthonous subsidies to the littoral food web. Brook trout derived their food mainly from benthic predatory macroinvertebrates in unperturbed lakes. However, in perturbed lakes one year after harvesting, zooplankton appeared to be the main contributor to brook trout diet. This change in brook trout diet was mitigated two years after harvesting. Size-related diet shift were also observed for brook trout, indicating a diet shift related to size. Our study suggests that carbon from terrestrial habitat may be a significant contribution to the food web of oligotrophic Canadian Boreal Shield lakes. Forest harvesting did not have an impact on the diet of benthic primary consumers. On the other hand, brook trout diet composition was affected by logging with greater zooplankton contribution in perturbed lakes, possibly induced by darker-colored environment in these lakes one year after logging.

  13. Mantle structure beneath eastern Africa: Evidence for a through going-mantle anomaly and its implications for the origin of Cenozoic tectonism in eastern Africa

    NASA Astrophysics Data System (ADS)

    Mulibo, G.; Tugume, F.; Julia, J.

    2012-12-01

    In this study, teleseismic earthquakes recorded on over 60 temporary AfricaArray seismic stations deployed in Uganda, Kenya, Tanzania and Zambia between 2007 and 2011 are used to invert P and S travel time residuals, together with travel time residuals from previous deployments, for a 3D image of mantle wave speeds and for examining relief on transition zone discontinuities using receiver function stacks. Tomographic images reveal a low wave speed anomaly (LWA) that dips to the SW beneath northern Zambia, extending to a depth of at least 900 km. The anomaly appears to be continuous across the transition zone, extending into the lower mantle. Receiver function stacks reveal an average transition zone thickness (TZT) across a wide region extending from central Zambia to the NE through Tanzania and into Kenya, which is ~30-40 km thinner than the global average. These results are not easily explained by models for the origin of the Cenozoic tectonism in eastern Africa that invoke a plume head or small scale convection either by edge flow or passive stretching of the lithosphere. However, the depth extent of the LWA coincident with a thin transition zone is consistent with a model invoking a through-going mantle anomaly beneath eastern Africa that links anomalous upper mantle to the African Superplume anomaly in the lower mantle beneath southern Africa. This finding indicates that geodynamic processes deep in the lower mantle are influencing surface dynamics across the Afro-Arabian rift system.

  14. Population structure and phylogeography reveal pathways of colonization by a migratory marine reptile (Chelonia mydas) in the central and eastern Pacific.

    PubMed

    Dutton, Peter H; Jensen, Michael P; Frey, Amy; LaCasella, Erin; Balazs, George H; Zárate, Patricia; Chassin-Noria, Omar; Sarti-Martinez, Adriana Laura; Velez, Elizabeth

    2014-11-01

    Climate, behavior, ecology, and oceanography shape patterns of biodiversity in marine faunas in the absence of obvious geographic barriers. Marine turtles are an example of highly migratory creatures with deep evolutionary lineages and complex life histories that span both terrestrial and marine environments. Previous studies have focused on the deep isolation of evolutionary lineages (>3 mya) through vicariance; however, little attention has been given to the pathways of colonization of the eastern Pacific and the processes that have shaped diversity within the most recent evolutionary time. We sequenced 770 bp of the mtDNA control region to examine the stock structure and phylogeography of 545 green turtles from eight different rookeries in the central and eastern Pacific. We found significant differentiation between the geographically separated nesting populations and identified five distinct stocks (F ST = 0.08-0.44, P < 0.005). Central and eastern Pacific Chelonia mydas form a monophyletic group containing 3 subclades, with Hawaii more closely related to the eastern Pacific than western Pacific populations. The split between sampled central/eastern and western Pacific haplotypes was estimated at around 0.34 mya, suggesting that the Pacific region west of Hawaii has been a more formidable barrier to gene flow in C. mydas than the East Pacific Barrier. Our results suggest that the eastern Pacific was colonized from the western Pacific via the Central North Pacific and that the Revillagigedos Islands provided a stepping-stone for radiation of green turtles from the Hawaiian Archipelago to the eastern Pacific. Our results fit with a broader paradigm that has been described for marine biodiversity, where oceanic islands, such as Hawaii and Revillagigedo, rather than being peripheral evolutionary "graveyards", serve as sources and recipients of diversity and provide a mechanism for further radiation.

  15. Population structure and phylogeography reveal pathways of colonization by a migratory marine reptile (Chelonia mydas) in the central and eastern Pacific

    PubMed Central

    Dutton, Peter H; Jensen, Michael P; Frey, Amy; LaCasella, Erin; Balazs, George H; Zárate, Patricia; Chassin-Noria, Omar; Sarti-Martinez, Adriana Laura; Velez, Elizabeth

    2014-01-01

    Climate, behavior, ecology, and oceanography shape patterns of biodiversity in marine faunas in the absence of obvious geographic barriers. Marine turtles are an example of highly migratory creatures with deep evolutionary lineages and complex life histories that span both terrestrial and marine environments. Previous studies have focused on the deep isolation of evolutionary lineages (>3 mya) through vicariance; however, little attention has been given to the pathways of colonization of the eastern Pacific and the processes that have shaped diversity within the most recent evolutionary time. We sequenced 770 bp of the mtDNA control region to examine the stock structure and phylogeography of 545 green turtles from eight different rookeries in the central and eastern Pacific. We found significant differentiation between the geographically separated nesting populations and identified five distinct stocks (FST = 0.08–0.44, P < 0.005). Central and eastern Pacific Chelonia mydas form a monophyletic group containing 3 subclades, with Hawaii more closely related to the eastern Pacific than western Pacific populations. The split between sampled central/eastern and western Pacific haplotypes was estimated at around 0.34 mya, suggesting that the Pacific region west of Hawaii has been a more formidable barrier to gene flow in C. mydas than the East Pacific Barrier. Our results suggest that the eastern Pacific was colonized from the western Pacific via the Central North Pacific and that the Revillagigedos Islands provided a stepping-stone for radiation of green turtles from the Hawaiian Archipelago to the eastern Pacific. Our results fit with a broader paradigm that has been described for marine biodiversity, where oceanic islands, such as Hawaii and Revillagigedo, rather than being peripheral evolutionary “graveyards”, serve as sources and recipients of diversity and provide a mechanism for further radiation. PMID:25540693

  16. Population structure and phylogeography reveal pathways of colonization by a migratory marine reptile (Chelonia mydas) in the central and eastern Pacific.

    PubMed

    Dutton, Peter H; Jensen, Michael P; Frey, Amy; LaCasella, Erin; Balazs, George H; Zárate, Patricia; Chassin-Noria, Omar; Sarti-Martinez, Adriana Laura; Velez, Elizabeth

    2014-11-01

    Climate, behavior, ecology, and oceanography shape patterns of biodiversity in marine faunas in the absence of obvious geographic barriers. Marine turtles are an example of highly migratory creatures with deep evolutionary lineages and complex life histories that span both terrestrial and marine environments. Previous studies have focused on the deep isolation of evolutionary lineages (>3 mya) through vicariance; however, little attention has been given to the pathways of colonization of the eastern Pacific and the processes that have shaped diversity within the most recent evolutionary time. We sequenced 770 bp of the mtDNA control region to examine the stock structure and phylogeography of 545 green turtles from eight different rookeries in the central and eastern Pacific. We found significant differentiation between the geographically separated nesting populations and identified five distinct stocks (F ST = 0.08-0.44, P < 0.005). Central and eastern Pacific Chelonia mydas form a monophyletic group containing 3 subclades, with Hawaii more closely related to the eastern Pacific than western Pacific populations. The split between sampled central/eastern and western Pacific haplotypes was estimated at around 0.34 mya, suggesting that the Pacific region west of Hawaii has been a more formidable barrier to gene flow in C. mydas than the East Pacific Barrier. Our results suggest that the eastern Pacific was colonized from the western Pacific via the Central North Pacific and that the Revillagigedos Islands provided a stepping-stone for radiation of green turtles from the Hawaiian Archipelago to the eastern Pacific. Our results fit with a broader paradigm that has been described for marine biodiversity, where oceanic islands, such as Hawaii and Revillagigedo, rather than being peripheral evolutionary "graveyards", serve as sources and recipients of diversity and provide a mechanism for further radiation. PMID:25540693

  17. Excess growing-season water limits lowland black spruce productivity

    NASA Astrophysics Data System (ADS)

    Dymond, S.; Kolka, R. K.; Bolstad, P. V.; Gill, K.; Curzon, M.; D'Amato, A. W.

    2015-12-01

    The annual growth of many tree species is limited by water availability, with growth increasing as water becomes less scarce. In lowland bogs of northern Minnesota, however, black spruce (Picea mariana) is often exposed to excess water via high water table elevations. These trees grow in thick deposits of organic mucky peat and often have shallow rooting systems to avoid the complete submersion of roots in water. While it is generally believed that black spruce decrease growth rates with rising water table elevations, this hypothesis has not been tested in situ. We used a unique, 50-year record of daily bog water table elevations at the Marcell Experimental Forest (MEF) in northern Minnesota to investigate the relationship between climate and black spruce productivity. Nine 1/20th ha circular plots were established in five different bogs and tree height, diameter-at-breast-height (DBH), and crown class were recorded. Additionally, two perpendicular cores were collected on all trees greater than 10 cm diameter-at-breast-height. Tree cores were sanded, mounted, cross-dated, and de-trended according to standard dendrochronological procedures. Ring width measurements were correlated with precipitation, temperature, and water table elevation using package BootRes in R to determine the climatic variables most associated with stand level productivity. Across the different plots, we found that early growing season water table elevation (May and June) was negatively correlated with both individual and stand-level black spruce growth (p < 0.01), while growth was positively correlated with March temperatures (p < 0.01). No significant relationships existed between black spruce growth and monthly precipitation. If summer water table elevations in these peatland ecosystems rise as is anticipated with more extreme precipitation events due to climate change, we could see an overall decrease in the stand level productivity of black spruce.

  18. Cloud deposition to a spruce forest edge

    NASA Astrophysics Data System (ADS)

    Weathers, K. C.; Lovett, G. M.; Likens, G. E.

    Deposition from clouds to a spruce (Picea rubens Sarg.) forest edge on Hunter Mt. in the Catskill Mts of New York State was measured during 1987 and 1988 to determine whether the windward edge of forest floor receives greater deposition of water and ions via cloud water than the interior of a forest. Throughfall was used as a measure of deposition and was collected during cloud-only and mixed cloud-and-rain events along five windward-to-leeward transects in a 30 x 30 m forested area. Ambient cloud water was also collected in a passive collector and chemically analyzed. Trees at the edge of the forest received on average three times, and up to 15 times, greater deposition of ions than those in the interior of the forest. Lead content in samples from Hunter Mt. forest floor at the windward edge, relative to the interior, was enhanced as well. Using a regression of distance vs deposition, the deposition "half-distance", (i.e. the point at which the rate of cloud water deposition is 50% of the rate at the windward edge of the forest) was found to be 28 m. The cloud deposition data from this study are compared to other studies of Na particle deposition to low-elevation forest edges, which show similar deposition "half distances", ranging from ˜ 2 to 36 m into the forest. Most models of cloud deposition currently in use assume landscape homogeneity. Montane forest landscapes, however, are often highly heterogeneous, consisting of many "edges", and thus current models may seriously underestimate cloud deposition.

  19. Lithospheric structure of the eastern flank of the Rio Grande Rift via receiver function velocity analysis

    NASA Astrophysics Data System (ADS)

    Agrawal, M.; Pulliam, J.; Sen, M. K.; Grand, S.

    2015-12-01

    To better delineate a seismic anomaly beneath the eastern flank of the Rio Grande Rift identified by seismic tomography, we depth-migrated Ps and Sp receiver functions using data from the SIEDCAR (Seismic Investigation of Edge Driven Convection Association with Rio Grande Rift) and USArray Transportable Array (TA) deployments. We performed Common Conversion Point (CCP) stacking to improve the S/N ratio of receiver functions. Using an incorrect velocity model for depth migration of a stacked CCP image may generate an inaccurate picture of the subsurface. To find sufficiently accurate P- and S-velocity models for migration, we optimize the average correlation value of common receiver gathers for target features - in this case the Moho and the LAB - while perturbing the shear wave velocities in a process driven by simulated annealing. The technique simultaneously finds depths to major discontinuities (in this case the Moho and LAB) and S and P velocity profiles beneath each seismic station in a manner that is similar to velocity analysis in reflection seismology. An application to data acquired in southeastern New Mexico and west Texas, at an average station spacing of 35 km, reveals an abrupt increase in lithospheric thickness from west to east, from the Rio Grande Rift to the Great Plains craton. Previous studies found an elongated high velocity anomaly that extends to depths approaching 500 km in southeastern New Mexico and west Texas that is distinct from the thick Great Plains lithosphere. Our stacked 3-D image confirms the anomaly's existence and shows that it is more laterally extensive than was previously indicated. Recent numerical modeling suggests that an abrupt change in lithospheric thickness, which creates a step change in densities, may produce a gravitational instability that leads to thicker mantle lithosphere dripping off into the lower density asthenosphere. As the mantle deforms it alternately thickens and thins the crust, producing topographic

  20. Genetic uniformity and the decline of red spruce

    SciTech Connect

    Not Available

    1989-05-01

    This brief note examines current research that suggests genetic uniformity as a cause of red spruce decline. Although acid rain and ozone have been implicated in the decline of the species, dieback has been observed evening areas where pollution is low. In addition, the dieback is not observed in other species. Researchers have analyzed the seeds of approximately 500 red spruce trees representing 13 boreal and Montane populations from southern Appalachia to Canada. Of 42 gene loci examined, on the average only 8% of the genes were heterozygousand only 31% were polymorphic.

  1. Neotectonic stage in the formation of the Khokhlovskoe uranium deposit, eastern Transural region: Structural, hydrochemical, mineralogical, and geochemical formation conditions

    NASA Astrophysics Data System (ADS)

    Vinokurov, S. F.; Prokof'ev, V. Yu.; Malkovsky, V. I.; Dymkov, Yu. M.; Chugaev, A. V.; Nesterova, M. V.

    2013-11-01

    The structural, hydrochemical, mineralogical, and geochemical features of the Khokhlovskoe uranium deposit related to neotectonic processes are considered. The structural feature is expressed in neotectonic dislocations in the form of overall intense fragmentation of host rocks and widespread low-amplitude strike-slip faulting. The hydrochemical specificity is determined by the appearance of thermal carbonated formation water in ore-bearing aquifers. This water is similar in chemical and gas composition to hydrothermal solutions in fluid inclusions and mineral waters abundant in this district. The mineralogical and geochemical features comprise the occurrence of newly formed ferroan carbonates and late iron hydroxides in altered (bleached) pelitic rocks; the formation of silicic opal segregations in ore-bearing sand and sandstone; late sulfides, arsenides, and selenides of iron and other metals; and multiphase gel-pitchblende enriched in Zr especially typical of high-grade uranium ore. The age of high-grade ore determined by a precision uranium-ionium method coincides with the time when thermal carbonated water appeared in the host rocks. This time was estimated from a mathematical model of heat transfer and regional dynamics of underground water. This coincidence clearly indicates that the aforementioned processes are related to the late Quaternary neotectonic reactivation of the eastern Transural region.

  2. Changes in soil microbial community structure following the abandonment of agricultural terraces in mountainous areas of Eastern Spain

    PubMed Central

    Zornoza, R.; Guerrero, C.; Mataix-Solera, J.; Scow, K.M.; Arcenegui, V.; Mataix-Beneyto, J.

    2012-01-01

    In Eastern Spain, almond trees have been cultivated in terraced orchards for centuries, forming an integral part of the Mediterranean forest scene. In the last decades, orchards have been abandoned due to changes in society. This study investigates effects of changes in land use from forest to agricultural land and the posterior land abandonment on soil microbial community, and the influence of soil physico-chemical properties on the microbial community composition (assessed as abundances of phospholipids fatty acids, PLFA). For this purpose, three land uses (forest, agricultural and abandoned agricultural) at four locations in SE Spain were selected. Multivariate analysis showed a substantial level of differentiation in microbial community structure according to land use. The microbial communities of forest soils were highly associated with soil organic matter content. However, we have not found any physical or chemical soil property capable of explaining the differences between agricultural and abandoned agricultural soils. Thus, it was suggested that the cessation of the perturbation caused by agriculture and shifts in vegetation may have led to changes in the microbial community structure. PLFAs indicative of fungi and ratio of fungal to bacterial PLFAs were higher in abandoned agricultural soils, whereas the relative abundance of bacteria was higher in agricultural soils. Actinomycetes were generally lower in abandoned agricultural soils, while the proportions of vesicular-arbuscular mycorrhyzal fungi were, as a general trend, higher in agricultural and abandoned agricultural soils than in forests. Total microbial biomass and richness increased as agricultural < abandoned agricultural < forest soils. PMID:22291451

  3. Structural control on mass-movement evolution: a case study from the Vizze Valley, Italian Eastern Alps

    NASA Astrophysics Data System (ADS)

    Bistacchi, A.; dal Piaz, G. V.; Massironi, M.; Monopoli, B.; Schiavo, A.

    2003-04-01

    A large post-glacial deep-seated mass movement is described from the area east of Vipiteno-Sterzing (Alto Adige South Tyrol, Eastern Alps), mainly developed in calcschists of the ophiolitic Glockner nappe (southern antiform of SW-Tauern window). A close relationship exists between differential slope evolution, structural setting, and slope attitude. Lateral spread develops in an area where the dominant schistosity is subvertical and parallel to the slope, and is facilitated by dissolution of evaporite-bearing bodies. Sackung develops in areas with low-angle schistosity and NNE-SSW high-angle faults, parallel to the ridge segment. A huge rock-fall was controlled by the intersection of the steep N-dipping schistosity with E-W and N-S to NNE-SSW faults and joints. As a whole, this deep-seated gravitational slope deformation (DSGSD) is an outstanding example of how the mechanics of large mass movements and their potential evolution are controlled by the structural framework, which is the ultimate result of a complex tectonic history.

  4. Effectiveness of polyethylene sheeting in controlling spruce beetles ( coleoptera: scolytidae') in infested stacks of spruce firewood in Alaska. Forest Service research paper

    SciTech Connect

    Holsten, E.H.; Werner, R.A.

    1993-06-01

    The covering stacks of spruce firewood with either clear or black polyethylene sheeting does not raise log temperatures high enough to kill spruce beetle brood in the logs. Based on the results of the study, the authors do not recommend the use of polyethylene sheeting as a remedial measure for the reduction of spruce beetle brood in infested firewood or log decks in south-central Alaska.

  5. An Expanded Chesapeake Bay Impact Structure, Eastern Virginia: New Corehole and Geophysical Data

    NASA Technical Reports Server (NTRS)

    Powars, D. S.; Johnson, G. H.; Edwards, L. E.; Horton, J. W., Jr.; Gohn, G. S.; Catchings, R. D.; McFarland, E. R.; Izett, G. A.; Bruce, T. S.; Levine, J. S.

    2002-01-01

    Data from several deep coreholes, seismic reflection surveys, and surface mapping indicate that the buried Chesapeake Bay impact structure is wider (160 km, due to 35-km-wide outer fracture zone) and deeper (2 miles) than previously reported. Additional information is contained in the original extended abstract.

  6. Biological structure and dynamics of littoral fish assemblages in the eastern Finger Lakes

    USGS Publications Warehouse

    McKenna, James E.

    2001-01-01

    Fish assemblages from three of the New York Finger Lakes were examined for structure within and between lakes and over time. Species-area relationships indicated that local fish assemblages are the result of recent, lake-specific events that altered the regional species pool. Fish assemblages varied among seasons and those occupying eutrophic waters had different characteristics from those in oligotrophic waters. Bluntnose minnows (Pimephales notatus) were a persistent and important component of most assemblages, but abundance of bluegill (Lepomis macrochirus) was the most distinguishing feature. Species associations indicated that interactions among the fishes had little influence on assemblage structure. Correlations between community structure and abiotic factors were identified. Ten abiotic variables were strongly associated with the species assemblages, but could not fully explain differences between assemblages. Results indicate that the abundance and diversity of water column feeders was related to productivity of lake habitat. In general, fish populations were smaller in oligotrophic waters and water column feeders were poorly represented in those assemblages. Productivity at various trophic levels was implicated as a major factor determining lake fish assemblage structure.

  7. Transverse zones controlling the structural evolution of the Zipaquira Anticline (Eastern Cordillera, Colombia): Regional implications

    NASA Astrophysics Data System (ADS)

    García, Helbert; Jiménez, Giovanny

    2016-08-01

    We report paleomagnetic, magnetic fabric and structural results from 21 sites collected in Cretaceous marine mudstones and Paleogene continental sandstones from the limbs, hinge and transverse zones of the Zipaquira Anticline (ZA). The ZA is an asymmetrical fold with one limb completely overturned by processes like gravity and salt tectonics, and marked by several axis curvatures. The ZA is controlled by at least two (2) transverse zones known as the Neusa and Zipaquira Transverse Zones (NTZ and ZTZ, respectively). Magnetic mineralogy methods were applied at different sites and the main carriers of the magnetic properties are paramagnetic components with some sites being controlled by hematite and magnetite. Magnetic fabric analysis shows rigid-body rotation for the back-limb in the ZA, while the forelimb is subjected to internal deformation. Structural and paleomagnetic data shows the influence of the NTZ and ZTZ in the evolution of the different structures like the ZA and the Zipaquira, Carupa, Rio Guandoque, Las Margaritas and Neusa faults, controlling several factors as vergence, extension, fold axis curvature and stratigraphic detatchment. Clockwise rotations unraveled a block segmentation following a discontinuos model caused by transverse zones and one site reported a counter clockwise rotation associated with a left-lateral strike slip component for transverse faults (e.g. the Neusa Fault). We propose that diverse transverse zones have been active since Paleogene times, playing an important role in the tectonic evolution of the Cundinamarca sub-basin and controlling the structural evolution of folds and faults with block segmentation and rotations.

  8. Plasticity in variation of xylem and phloem cell characteristics of Norway spruce under different local conditions

    PubMed Central

    Gričar, Jožica; Prislan, Peter; de Luis, Martin; Gryc, Vladimír; Hacurová, Jana; Vavrčík, Hanuš; Čufar, Katarina

    2015-01-01

    There is limited information on intra-annual plasticity of secondary tissues of tree species growing under different environmental conditions. To increase the knowledge about the plasticity of secondary growth, which allows trees to adapt to specific local climatic regimes, we examined climate–radial growth relationships of Norway spruce [Picea abies (L.) H. Karst.] from three contrasting locations in the temperate climatic zone by analyzing tree-ring widths for the period 1932–2010, and cell characteristics in xylem and phloem increments formed in the years 2009–2011. Variation in the structure of xylem and phloem increments clearly shows that plasticity in seasonal dynamics of cambial cell production and cell differentiation exists on xylem and phloem sides. Anatomical characteristics of xylem and phloem cells are predominantly site-specific characteristics, because they varied among sites but were fairly uniform among years in trees from the same site. Xylem and phloem tissues formed in the first part of the growing season seemed to be more stable in structure, indicating their priority over latewood and late phloem for tree performance. Long-term climate and radial growth analyses revealed that growth was in general less dependent on precipitation than on temperature; however, growth sensitivity to local conditions differed among the sites. Only partial dependence of radial growth of spruce on climatic factors on the selected sites confirms its strategy to adapt the structure of wood and phloem increments to function optimally in local conditions. PMID:26442044

  9. Plasticity in variation of xylem and phloem cell characteristics of Norway spruce under different local conditions.

    PubMed

    Gričar, Jožica; Prislan, Peter; de Luis, Martin; Gryc, Vladimír; Hacurová, Jana; Vavrčík, Hanuš; Čufar, Katarina

    2015-01-01

    There is limited information on intra-annual plasticity of secondary tissues of tree species growing under different environmental conditions. To increase the knowledge about the plasticity of secondary growth, which allows trees to adapt to specific local climatic regimes, we examined climate-radial growth relationships of Norway spruce [Picea abies (L.) H. Karst.] from three contrasting locations in the temperate climatic zone by analyzing tree-ring widths for the period 1932-2010, and cell characteristics in xylem and phloem increments formed in the years 2009-2011. Variation in the structure of xylem and phloem increments clearly shows that plasticity in seasonal dynamics of cambial cell production and cell differentiation exists on xylem and phloem sides. Anatomical characteristics of xylem and phloem cells are predominantly site-specific characteristics, because they varied among sites but were fairly uniform among years in trees from the same site. Xylem and phloem tissues formed in the first part of the growing season seemed to be more stable in structure, indicating their priority over latewood and late phloem for tree performance. Long-term climate and radial growth analyses revealed that growth was in general less dependent on precipitation than on temperature; however, growth sensitivity to local conditions differed among the sites. Only partial dependence of radial growth of spruce on climatic factors on the selected sites confirms its strategy to adapt the structure of wood and phloem increments to function optimally in local conditions. PMID:26442044

  10. Structure and dynamics in the north Jizhong Depression, Bohai Bay Basin, Eastern China

    NASA Astrophysics Data System (ADS)

    Miao, Q.; Qi, J.; Zhang, J.

    2015-12-01

    Study of structural geology in the north Jizhong Depression, Bohai Bay has achieved great breakthroughs in recent years. However the studies of structure and dynamics still remain much controversy. Based on the 3D and partial 2D seismic data, combined with regional geology and well data, the characteristics of structures in the north Jizhong Deprssion are analyzed, the dynamics is discussed. Our analyses show that the Cenozoic structures in the north Jizhong Depression can be subdivided into extensional system and strike-slip system. Extensional system concludes series of normal faults and transfer faults. Normal faults are mainly trend NNE and NE. They control the Paleogene sediments in sub-depressions of hanging-wall, and bottom out into a sub-horizontal detachment zone in deep level shaped like listric. Transfer faults mainly adjust displacement of normal faults, yet some like Tongbozhen and Niutuozhen are transfer faults in Paleocene and Eocene but change to normal faults in Oligocene. Strike-slip system is predominantly consisted by sub-vertical right-lateral strike-slip faults such as Xin`anzhen and Maxi. From seismic profiles, Xin`anzhen and Maxi cut into basement but only influence the sediments of Ed of Paleogene and Lower Neogene. Based on the relation of sedimentary sequence and faults, the extensional system mainly develops from Paleocene to Late Oligocene, but strike-slip system predominantly develops from Late Oligocene to Miocene. From the superposed section of crustal structure and basin structure in Jizhong Depression, the Cenozoic sub-basin is just located on the thinner zones of crustal, corresponding with the location of mantle uplift. It indicates that the Paleogene development of Jizhong Depression is related to the horizontal tension caused by uplift of magma in mantle. The results illustrate that extensional deformation in Jizhong depression is caused by the uplift of magma. The right-lateral strike-slip deformation, same with Tancheng

  11. Competitive strategies in adult beech and spruce: space-related foliar carbon investment versus carbon gain.

    PubMed

    Reiter, I M; Häberle, K-H; Nunn, A J; Heerdt, C; Reitmayer, H; Grote, R; Matyssek, R

    2005-12-01

    In Central Europe, Fagus sylvatica and Picea abies represent contrasting extremes in foliage type, crown structure and length of growing season. In order to examine the competitive strategies of these two co-occurring species, we tested the following hypotheses: (1) the space occupied by the foliage of sun branches is characterized by greater foliar mass investment compared to shade branches, (2) the carbon (C) gain per unit of occupied space is greater in sun than in shade branches, and (3) annual C and water costs of the foliage for sustaining the occupied space are low, wherever C gain per unit of occupied space is low. These were investigated in a mature forest in Southern Germany. The examination was based on the annual assessment of space-related resource investments and gains of the foliage. The foliated space around branches was regarded as the relevant volume with respect to aboveground resource availability. Occupied crown space per standing foliage mass was higher in shade compared to sun branches of beech, whereas no difference existed in crown volume per foliage mass between sun and shade branches of spruce (hypothesis 1 accepted for beech but rejected for spruce). However, beech occupied more space per foliage mass than spruce. The C gain per occupied crown volume was greater in sun than in shade branches (hypothesis 2 accepted) but did not differ between species. The amount of occupied space per respiratory and transpiratory costs did not differ between species or between sun and shade branches. In beech and spruce, the proportion of foliage investment in the annual C balance of sun and shade branches remained rather stable, whereas respiratory costs distinctly increased in shade foliage. Hence, shade branches were costly structures to occupy space, achieving only low and even negative C balances (rejection of hypothesis 3), which conflicts with the claimed C autonomy of branches. Our findings suggest that competitiveness is determined by the

  12. Effects of structural complexity enhancement on eastern red-backed salamander (Plethodon cinereus) populations in northern hardwood forests

    USGS Publications Warehouse

    McKenny, H.C.; Keeton, W.S.; Donovan, T.M.

    2006-01-01

    Managing for stand structural complexity in northern hardwood forests has been proposed as a method for promoting microhabitat characteristics important to eastern red-backed salamanders (Plethodon cinereus). We evaluated the effects of alternate, structure-based silvicultural systems on red-backed salamander populations at two research sites in northwestern Vermont. Treatments included two uneven-aged approaches (single-tree selection and group-selection) and one unconventional approach, termed "structural complexity enhancement" (SCE), that promotes development of late-successional structure, including elevated levels of coarse woody debris (CWD). Treatments were applied to 2 ha units and were replicated two to four times depending on treatment. We surveyed red-backed salamanders with a natural cover search method of transects nested within vegetation plots 1 year after logging. Abundance estimates corrected for detection probability were calculated from survey data with a binomial mixture model. Abundance estimates differed between study areas and were influenced by forest structural characteristics. Model selection was conducted using Akaike Information Criteria, corrected for over-dispersed data and small sample size (QAICc). We found no difference in abundance as a response to treatment as a whole, suggesting that all of the uneven-aged silvicultural systems evaluated can maintain salamander populations after harvest. However, abundance was tied to specific structural habitat attributes associated with study plots within treatments. The most parsimonious model of habitat covariates included site, relative density of overstory trees, and density of more-decayed and less-decayed downed CWD. Abundance responded positively to the density of downed, well-decayed CWD and negatively to the density of poorly decayed CWD and to overstory relative density. CWD volume was not a strong predictor of salamander abundance. We conclude that structural complexity enhancement

  13. Subsurface structure along the eastern marginal fault zone of Yokote Basin by Seismic reflection profiling studies, Northeast Japan

    NASA Astrophysics Data System (ADS)

    Kagohara, K.; Imaizumi, T.; Echigo, T.; Miyauchi, T.; Sato, H.

    2005-12-01

    Typical reverse faults, which are known as Senya earthquake faults appeared along the western foot of the Mahiru Mountains, associated with The Rikuu Earthquake (Mj7.2) of 1896 in Northeast Japan. Eastern marginal fault zone of the Yokote Basin consist of four main surface ruptures, about 35 km long, Obonai fault, Shiraiwa fault, Ota fault and Senya fault, depending on their continuity and strike (Matsuda et al., 1980). We carried out the seismic reflection profiling across these faults (Kawaguchi03 Seismic line, Unjono04 Seismic line and Kotaki05 Seismic line) to clarify the subsurface structure of these reverse fault system based on the data of tectonic geomorphology and structural geology and furthermore, to discuss the timing of migration of the thrusting from the range front to the basin margin. The seismic source was mini-vibrator trucks, with 20seconds of 10-100Hz signals at 10m or 5m intervals. The sweep signals were recorded by a digital telemetry system (GDAPS-4a) with 10 Hz geophones. The obtained seismic reflection data were processed by conventional Common mid-point (CMP) methods, including post-stack migration and depth conversion. The resulting seismic reflection profile reveals a thrust structure beneath these areas. At the Center of Senya hills there are two thrusts and one high angle reverse fault (1997 Seismic Line). Senya fault is an active frontal emergent thrust with flat and ramp structure. Although, the high angle reverse fault, located along the foot of the range is a short-cut branching fault from the Senya fault in the central part of the Senya hill (Sato et al., 1998), in the Unjono04 seismic line, the depth of the flat and ramp structure gradually shallow in the north part of the Senya hill, where the flexure scarp accompanied with antithetic faults formed on the fluvial terraces. In the Kawaguchi03 seismic line, the concealed fault, 0.5km below the surface, branched from the master Ota fault, form a flexure scarplet on the alluvial fan

  14. Cenozoic structural history of selected areas in the eastern Great Basin, Nevada-Utah

    USGS Publications Warehouse

    Anderson, R. Ernest

    1983-01-01

    The Confusion Range structural trough (CRST) of west-central Utah predates the Oligocene rocks that are exposed along it. The northern part of the axial region of the CRST is complicated by structures that include reverse faults and associated folds, a large-amplitude mushroom fold, and belts of sharply flexed to overturned strata some of which are fault bounded. These structures, which also predate the Oligocene rocks, formed in a compressional regime that has been interpreted as resulting from thin-skinned gravitational gliding toward the axis of the CRST. Study of the sparse Tertiary rocks that are scattered along the axial region of the CRST reveals abundant evidence of Oligocene and younger deformation. The chief evidence includes (1) widespread Oligocene and Miocene coarse clastic rocks, many of which are conglomerates, that attest to local and distant tectonism, (2) faults that range from high-angle structures generally with less than 100 m of normal displacement to low-angle attenuation faults some of which may have large displacements, and (3) open asymmetric folds. Together with the distribution of sheet-form bodies of ash-flow tuffs, the Oligocene stratigraphic record allows for paleogeographic reconstruction of a lacustrine basin across what is now the northern Confusion Range and one or more basins in the southern part of the CRST. The basins are inferred to have been fault controlled by reactivation of previously formed faults or steep fold flanks. They may have been localized by differential vertical movements similar to those that produced the older systems of folds and faults. Parts of early formed basins were cannibalized as local syndepositional deformation took place in the axial region of the CRST. Both limbs of the CRST have been modified by folds that involve Oligocene rocks. Some of these folds appear to be genetically related to displacements on faults that bound them. They may record thin-skinned Neogene tectonic displacements toward the

  15. Wood Anatomy and Insect Defoliator Systems: Is there an anatomical response to sustained feeding by the western spruce budworm (Choristoneura occidentalis) on Douglas-fir (Pseudotusga menziesii)?

    NASA Astrophysics Data System (ADS)

    Axelson, Jodi; Gärtner, Holger; Alfaro, René; Smith, Dan

    2013-04-01

    The western spruce budworm (Choristoneura occidentalis Freeman) is the most widespread and destructive defoliator of coniferous forests in western North America, and has a long-term coexistence with its primary host tree, Douglas-fir (Pseudotsuga menziesii Franco). Western spruce budworm (WSB) outbreaks usually last for several years, and cause reductions in annual growth, stem defects, and regeneration delays. In British Columbia, the WSB is the second most damaging insect after the mountain pine beetle, and sustained and/or severe defoliation can result in the mortality of host trees. Numerous studies have used tree rings to reconstruct WSB outbreaks across long temporal scales, to evaluate losses in stand productivity, and examine isotope ratios. Although some studies have looked at the impacts of artificial defoliation on balsam fir in eastern North America, there has been no prior research on how WSB outbreaks affect the anatomical structure of the stem as described by intra-annual wood density and potential cell size variations. The objective of this study was to anatomically examine the response of Douglas-fir to sustained WSB outbreaks in two regions of southern British Columbia. We hypothesize that the anatomical intra-annual characteristics of the tree rings, such as cell wall thickness, latewood cell size, and/or lumen area changes during sustained WSB outbreaks. To test this hypothesis we sampled four permanent sample plots in coastal and dry interior sites, which had annually resolved defoliation data collected over a 7-12 year period. At each site diameter-at-breast height (cm), height (m), and crown position were recorded and three increment cores were extracted from 25 trees. Increment cores were prepared to permit anatomical and x-ray density analyses. For each tree, a 15µm thick micro section was cut from the radial plane. Digital images of the micro sections were captured and processed. In each annual ring, features such as cell lumen area (µm2

  16. Long-term trends in the structure of eastern Adriatic littoral fish assemblages: Consequences for fisheries management

    NASA Astrophysics Data System (ADS)

    Stagličić, N.; Matić-Skoko, S.; Pallaoro, A.; Grgičević, R.; Kraljević, M.; Tutman, P.; Dragičević, B.; Dulčić, J.

    2011-09-01

    Long-term interannual changes in abundance, biomass, diversity and structure of littoral fish assemblages were examined between 1993 and 2009 by experimental trammel net fishing up to six times per year, within the warm period - May to September, at multiple areas along the eastern Adriatic coast with the aim of testing for the consistency of patterns of change across a large spatial scale (˜600 km). The results revealed spatially consistent increasing trends of total fish abundance and biomass growing at an average rate of 15 and 14% per year, respectively. Of the diversity indices analysed, the same pattern of variability was observed for Shannon diversity, while Pielou evenness and average taxonomic distinctness measures Δ ∗ and Δ + showed spatial variability with no obvious temporal trends. Multivariate fish assemblage structure underwent a directional change displaying a similar pattern through time for all the areas. The structural change in fish assemblages generally involved most of the species present in trammel net catches. A large pool of fish species responsible for producing the temporal pattern of assemblage change was relatively different in each of the areas reflecting a large geographic range covered by the study. An analysis of 4 fish species ( Symphodus tinca, Pagellus erythrinus, Mullus surmuletus, Scorpaena porcus) common to each of the study areas as the ones driving the temporal change indicated that there were clear increasing trends of their mean catches across the years at all the study areas. A common pattern among time trajectories across the spatial scale studied implies that the factor affecting the littoral fish assemblages is not localised but regional in nature. As an underlying factor having the potential to induce such widespread and consistent improvements in littoral fish assemblages, a more restrictive artisanal fishery management that has progressively been put in place during the study period, is suggested and discussed.

  17. A new name for the western spruce budworm (Lepidoptera: Tortricidae)?

    Technology Transfer Automated Retrieval System (TEKTRAN)

    There currently is considerable confusion as to the correct application of the name Choristoneura occidentalis. This name has historically been applied to the western spruce budworm, a serious forest pest in western North America. However, Razowski (2008) transferred Archips occidentalis into Choris...

  18. Structural survey and faults kinematics analysis of the eastern Thaumasia Region, Mars

    NASA Astrophysics Data System (ADS)

    Borraccini, F.; di Achille, G.

    Thaumasia Region, Mars F. Borraccini (1), G. Di Achille (2) (1) Institute of Environmental Dynamics, University of Urbino, Italy, (2) Int'l Research School of Planetary Sciences, Pescara, Italy (borraccini@uniurb.it / Fax: +39 0722 304295 / Phone: +39 0722 304295) Thaumasia plateau is a tectonically uplifted area buonded by outward-verging ridges (Thaumasia Highlands and Coprates Rise) and by extensional features (Valles Marineris and Claritas Fossae). Since the geodynamic and tectonic evolution of Thaumasia is still debated, we have undertaken a detailed tectonic mapping of a key-sector (of about 2.400.000 km2 ) in order to provide new constraints for the reconstruction of the geodynamic evolution of the whole Thaumasia Region. The study area encompasses the Coprates Chasma and Thaumasia Planum and is bordered by Melas Dorsae and by Nectaris Fossae to the West and East, respectively. In addition to previous Martian data sets (MOC, Themis, and Viking), we also used the HRSC images which made possible a detailed structural analysis. The area shows tectonic features characterized by different kinematics, origins and ages. In particular, horst and graben structures are arranged in two main WNW-ESE and NW-SE trending sets. A further set of normal faults is E-W oriented and dissects the Coprates Rise. This is affected by intense scarp retreat processes probably associated to wind and/or fluvial activities. E-W oriented right-lateral strike-slip faults postdates the extensional features and locally shows associated pull-apart basins. A compressive tectonic phase is recorded by NNE-SSW trending wrinkle ridges and by N-S oriented, east-verging regional buckling folds. We also investigated the relationship between the kinematics of those sets of structures and of major features like Valles Marineris and Coprates Rise. The analysis of crosscutting relationships between observed tectonic trends allowed the identification of the main deformational events occurred in the

  19. The Tschigot granodiorite (Eastern Alps, Southern Tyrol): a structural and AMS study

    NASA Astrophysics Data System (ADS)

    Georgiev, Neven; Pomella, Hannah; Cifelli, Francesca; Mattei, Massimo; Fügenschuh, Bernhard

    2013-04-01

    The Ordovician Tschigot granodiorite, hosted by the Texel complex, underwent polyphase tectonometamorphic overprint during the Variscan and Alpine orogenies. The main magmatic body forms a large sigma-like clast. While the contact with the host rock is characterized by a strong and coherent mylonitic foliation the internal part of the body is almost isotropic. The mylonitic foliation (S1.2 ) dips 40-60° toward N to NW. The related stretching lineation (L1.2) plunges 5-35° toward W or E, but also with some 10-15° deviations to NW or SW. Shear sense criteria related to L1.2 indicate top-to-the W sense of shear. Towards the inner parts of the body the intensity of deformation decreases rapidly within first 200-300 meters. From the mylonites at the contact (i.e. LS tectonites) the fabric first changes to form L>S and L-tectonites with their lineation (30-40° to NW) in perfect agreement with the orientation of the crenulation lineation (L2) from the surrounding host rocks. Unfortunately the continuous transition from LS to L-tectonites is hardly outcropping. Rarely two foliations (S1.1 and S1.2) can be observed in the granodiorite with the intersection lineation shallowly dipping towards NW (i.e. parallel to the lineation from the L-tectonite). While dynamic recrystallization of quartz in the L-tectonite is dominated by grain boundary migration with subordinate subgrain rotation (SGR), SGR characterizesthe mylonite from the contact area. With respect to their orientation the analysed magnetic lineations (Lmag) and foliations (Smag) are perfectly in agreement with the structural field data. The parallelism between magnetic (Smag, Lmag) and field structures (S1.2, L1.2) in the mylonite domain shows that in the temperature range between 400-500°C and under conditions of intensive shearing preexisting structural and magnetic features are fully overprinted and reoriented. In conclusion our data demonstrate perfect overlap between magnetic and structural fabrics in

  20. A terminal Pleistocene child cremation and residential structure from eastern Beringia.

    PubMed

    Potter, Ben A; Irish, Joel D; Reuther, Joshua D; Gelvin-Reymiller, Carol; Holliday, Vance T

    2011-02-25

    The dearth of human remains and residential sites has constrained inquiry into Beringian lifeways at the transition of the late Pleistocene-early Holocene. We report on human skeletal remains and a residential structure from central Alaska dated to ~11,500 calendar years ago. The remains are from a ~3-year-old child who was cremated in a pit within a semisubterranean house. The burial-cremation and house have exceptional integrity and preservation and exhibit similarities and differences to both Siberian Upper Paleolithic and North American Paleoindian features.

  1. A terminal Pleistocene child cremation and residential structure from eastern Beringia.

    PubMed

    Potter, Ben A; Irish, Joel D; Reuther, Joshua D; Gelvin-Reymiller, Carol; Holliday, Vance T

    2011-02-25

    The dearth of human remains and residential sites has constrained inquiry into Beringian lifeways at the transition of the late Pleistocene-early Holocene. We report on human skeletal remains and a residential structure from central Alaska dated to ~11,500 calendar years ago. The remains are from a ~3-year-old child who was cremated in a pit within a semisubterranean house. The burial-cremation and house have exceptional integrity and preservation and exhibit similarities and differences to both Siberian Upper Paleolithic and North American Paleoindian features. PMID:21350175

  2. Triangle zone and displacement transfer structures in the eastern Front Ranges, southern Canadian Rocky Mountains

    SciTech Connect

    Sanderson, D.A. ); Spratt, D.A. )

    1992-06-01

    The geometry of a relict triangle zone at the boundary of the Foothills and Front Ranges in the southern Canadian Rocky Mountains is constrained by detailed surface mapping over 700 m of relief and by seismic reflection data. The geometry and progressive development of the triangle zone along a strike length of 15-20 km, in the displacement transfer zone between the Coleman and Misty thrusts, is illustrated using closely spaced balanced cross sections, palinspastic restorations, and s sequentially restored cross section. Structural geometries show that a northeast- (foreland-) verging, mainly carbonate wedge of Mississippian to Triassic rock was inserted along a major upper detachment zone in shale, near the base of the Jurassic-Cretaceous clastic package. This was accompanied by southwest- (hinterland-) verging displacements along the upper detachment zone, tectonic thickening of the clastic package exceeding 200%, and backthrusting. Later northeast-verging deformation slightly modified the triangle zone by steepening structures, tightening folds, and minor thrusting. Recognition of relict triangle zones within the fold and thrust belt may document important changes in the rate of thrust front advancement, and aid in the delineation of potential hydrocarbon traps, similar to those discovered along the present-day thrust-belt margin in the southern Canadian Rocky Mountains.

  3. Structure of the eastern Seattle fault zone, Washington state: New insights from seismic reflection data

    USGS Publications Warehouse

    Liberty, L.M.; Pratt, T.L.

    2008-01-01

    We identify and characterize the active Seattle fault zone (SFZ) east of Lake Washington with newly acquired seismic reflection data. Our results focus on structures observed in the upper 1 km below the cities of Bellevue, Sammamish, Newcastle, and Fall City, Washington. The SFZ appears as a broad zone of faulting and folding at the southern boundary of the Seattle basin and north edge of the Seattle uplift. We interpret the Seattle fault as a thrust fault that accommodates north-south shortening by forming a fault-propagation fold with a forelimb breakthrough. The blind tip of the main fault forms a synclinal growth fold (deformation front) that extends at least 8 km east of Vasa Park (west side of Lake Sammamish) and defines the south edge of the Seattle basin. South of the deformation front is the forelimb break-through fault, which was exposed in a trench at Vasa Park. The Newcastle Hills anticline, a broad anticline forming the north part of the Seattle uplift east of Lake Washington, is interpreted to lie between the main blind strand of the Seattle fault and a backthrust. Our profiles, on the northern limb of this anticline, consistently image north-dipping strata. A structural model for the SFZ east of Lake Washington is consistent with about 8 km of slip on the upper part of the Seattle fault, but the amount of motion is only loosely constrained.

  4. ERTS-1 imagery of eastern Africa: A first look at the geological structure of selected areas

    NASA Technical Reports Server (NTRS)

    Mohr, P. A. (Principal Investigator)

    1972-01-01

    The author has identified the following significant results. Imagery of the African rift system resolves the major Cainozoic faults, zones of warping, and associated volcanism. It also clearly depicts the crystal grain of the Precambrian rocks where these are exposed. New structural features, or new properties of known features such as greater extent, continuity, and linearity are revealed by ERTS-1 imagery. This applies, for example, to the NE-SW fracture zones in Yemen, the Aswa mylonite zone at the northern end of the Western Rift, the Nandi fault of western Kenya, the linear faults of the Elgeyo escarpment in the Gregory Rift, and the hemibasins of warped Tertiary lavas on the Red Sea margin of Yemen, matching those of Ethiopian plateau-Afar margin. A tentative scheme is proposed, relating the effect on the pattern of Cainozoic faulting of the degree of obliquity to Precambrian structural trend. It is particularly noteworthy that, even where the Precambrian grain determines the rift faulting to be markedly oblique to the overall trend of the rift trough, for example, in central Lake Tanganyika, the width of the trough is not significantly increased. Some ground mapped lithological boundaries are obscure on ERTS-1 imagery.

  5. South Florida Ecosystem Program: quantifying freshwater discharge for coastal hydraulic control structures in eastern Dade County, Florida

    USGS Publications Warehouse

    Kapadia, Amit; Swain, Eric D.

    1996-01-01

    The South Florida Ecosystem Restoration Program is an intergovernmental effort, involving a number of agencies, to reestablish and maintain the ecosystem of south Florida. One element of the restoration effort is the development of a firm scientific basis for resource decision making. The U.S. Geological Survey (USGS), one of the agencies, provides scientific information as part of the South Florida Ecosystem Restoration Program. The USGS began their ow program, called the South Florida Ecosystem Program, in fiscal year 1995 for the purpose of gathering hydrologic, cartographic, and geologic data that relate to the mainland of south Florida, Florida Bay, and the Florida Keys and Reef ecosystems. As part of the South Florida Ecosystem Program, the USGS, in cooperation with the South Florida Water Management District (SFWMD), has conducted a study to determine discharge ratings for 16 coastal hydraulic control structures in eastern Dade County, Fla. (fig. 1 ). Discharge data are needed to quantify water that can be made available for water supply and ecosystem restoration and to calibrate regional hydrologic models.

  6. Higher fine-scale genetic structure in peripheral than in core populations of a long-lived and mixed-mating conifer - eastern white cedar (Thuja occidentalis L.)

    PubMed Central

    2012-01-01

    Background Fine-scale or spatial genetic structure (SGS) is one of the key genetic characteristics of plant populations. Several evolutionary and ecological processes and population characteristics influence the level of SGS within plant populations. Higher fine-scale genetic structure may be expected in peripheral than core populations of long-lived forest trees, owing to the differences in the magnitude of operating evolutionary and ecological forces such as gene flow, genetic drift, effective population size and founder effects. We addressed this question using eastern white cedar (Thuja occidentalis) as a model species for declining to endangered long-lived tree species with mixed-mating system. Results We determined the SGS in two core and two peripheral populations of eastern white cedar from its Maritime Canadian eastern range using six nuclear microsatellite DNA markers. Significant SGS ranging from 15 m to 75 m distance classes was observed in the four studied populations. An analysis of combined four populations revealed significant positive SGS up to the 45 m distance class. The mean positive significant SGS observed in the peripheral populations was up to six times (up to 90 m) of that observed in the core populations (15 m). Spatial autocorrelation coefficients and correlograms of single and sub-sets of populations were statistically significant. The extent of within-population SGS was significantly negatively correlated with all genetic diversity parameters. Significant heterogeneity of within-population SGS was observed for 0-15 m and 61-90 m between core and peripheral populations. Average Sp, and gene flow distances were higher in peripheral (Sp = 0.023, σg = 135 m) than in core (Sp = 0.014, σg = 109 m) populations. However, the mean neighborhood size was higher in the core (Nb = 82) than in the peripheral (Nb = 48) populations. Conclusion Eastern white cedar populations have significant fine-scale genetic structure at short distances. Peripheral

  7. Neogene Structural History of Biak and the Biak Basin, Eastern Indonesia

    NASA Astrophysics Data System (ADS)

    Gold, D.; Hall, R.; Burgess, P.

    2014-12-01

    The Biak Basin is within a frontier region, located between the islands of Biak to the north and Yapen to the south in the Indonesian province of West Papua. The Biak Basin is underlain by a Paleogene basement sequence that originated in an intra-oceanic island arc, formed on the leading edge of the oceanic Pacific-Caroline Plate. The basin has a structural history that can be divided into three stages: 1) a compressional stage, 2) a rifting stage and 3) a strike-slip stage. The compressional stage occurred during the Early Miocene. Collision of the arc with the northern edge of the Australian continental margin caused folding and thrusting of the basement sequence which was uplifted and subaerially exposed resulting in a widespread unconformity that can be traced west through the western Bird's Head as far as Halmahera. Following this collision the Biak Basin was filled largely by carbonates from the Early Miocene. During the Middle-Late Miocene the Biak Basin underwent a period of rapid subsidence resulting from a period of rifting. Extension occurred as convergence of the Pacific Plate with the Australian margin produced a regional stress field in which the maximum compressive stress was orientated NE-SW and maximum tensional stress NW-SE. This tensional stress produced NE-SW striking horsts and grabens with a Paleogene basement overlain by Lower Miocene carbonates. A final stage involving strike-slip faulting occurred during the Pliocene-Pleistocene with the initiation of major regional faults such as those of the Yapen Fault Zone and Biak Array which bound the southern and northeastern margins of the basin respectively. The strike-slip faulting accommodated continued convergence between the Pacific and Australian plates and uplifted Miocene carbonates as pop-up structures now observed on Biak.There are parallels to the Biak Basin in basins within the Basin and Range Province of the western United States, in particular the Walker Lake Basin, Nevada and other

  8. First images of the crustal structure across the eastern Algerian margin, from deep penetrating seismic data.

    NASA Astrophysics Data System (ADS)

    Bouyahiaoui, Boualem; Abtout, Abdeslam; Sage, Françoise; Klingelhoeffer, Frauke; Collot, Jean-yves; Yelles-chaouche, Abdelkarim; Marok, Abbas; Djellit, Hamou; Galves, Audrey; Bracène, Rabah; Schnurle, Philippe; Graindorge, David; party, Scientific

    2013-04-01

    The Algerian continental margin North Africa presents one of only a few examples of a passive continental margin formed in a back-arc environment, which undergoes current compression and is proposed to be reactivated today. In the framework of the Algerian - French SPIRAL research program (Sismique Profonde et Investigation Regionale du nord de l'ALgérie), a seismic cruise was conducted on the R/V Atalante from September to November 2009. During the cruise, deep penetrating low frequency multichannel and wide-angle seismic data were acquired in order to study the deep structure of the Algerian margin. In this work, we present the preliminary results from wide-angle modeling of the North-east Algerian margin in the region of Annaba along a N-S transect using a data set of 42 OBS (ocean bottom seismometers) along a profile extending 117km, and 13 broadband seismological stations along a profile of 80 km length. Travel-time tomography and forward modeling were undertaken to model the velocity structure in this region. The resulting velocity models image the thickness of the sedimentary layers, which varies between a few hundred meters on the continental margin of more than 4 km in the basin. The crust is about 6 km thick in the basin, and thickens to 7-8 km between 40 and 60km distance from the margin toe. Crustal thickness increases to about 22 km at the continental slope over a distance of ~ 90 km. The nature of the crust was determined to be thin oceanic with abnormal velocity gradient in the basin, and thinned continental from around 30 km distance from the coast landward. Integration of the wide-angle seismic data with multichannel seismic, gravity and magnetic data will help to better understand the structure of the Algerian margin and the adjacent oceanic basin in the Annaba region, and to discuss the numerous cinematic models proposed in literature regarding the formation of the north-Algerian basin.

  9. Environmental factors structuring fish composition and assemblages in a small macrotidal estuary (eastern English Channel)

    NASA Astrophysics Data System (ADS)

    Selleslagh, Jonathan; Amara, Rachid

    2008-09-01

    The fish assemblage structure was analyzed along an estuarine gradient of a small macrotidal estuary (the Canche, France). Fishes were collected every two months between May 2006 and July 2007 from 12 sampling stations using a 1.5-m beam trawl with a 5 mm mesh size in the cod end. To complement this information, sampling was also performed using 15-m fyke nets (8 mm mesh size in the cod end). For each sample, abiotic (temperature, salinity, pH, oxygen, turbidity, river flow, wind speed and depth) and biotic (macro crustacean species abundances) were recorded. Throughout the study, 28 fish species belonging to 20 families were collected. Fish catches were dominated by juveniles, especially Young-Of-the-Year (YOY) for the majority of the species. According to the Index of Relative Importance (IRI), common goby Pomatoschistus microps, flounder Platichtys flesus, sprat Sprattus sprattus, sea-bass Dicentrarchus labrax and plaice Pleuronectes platessa were the most abundant species, together accounting for 99.2% of the total IRI. Estuarine residents (ER = 66.2%) and marine juvenile migrants species (MJ = 31.4%) were the most important ecological guilds. The structure of the fish assemblage and its relationship to environmental variables was examined using multivariate techniques. Cluster and non-metric multidimensional scaling (nMDS) analysis defined six distinct groups in the Canche estuary, which are discriminated by specific species (SIMPER). Spatio-temporal variations in fish assemblage structure reflect the density peaks of the most abundant species. Spearman rank correlations and canonical correspondence analysis (CCA) showed that among the ten environmental variables examined, temperature, salinity and Crangon crangon (a potential predator for YOY fish or prey for older ones) are the three most important factors influencing fish species richness and abundances. Our observations reinforce the idea that certain fish species may have different life history styles in

  10. Geological and structural conditions of eclogitization of Paleoproterozoic basic dikes in the eastern Belomorian Mobile Belt

    NASA Astrophysics Data System (ADS)

    Kozlovsky, V. M.; Aranovich, L. Ya.

    2008-07-01

    A new, Krasnaya Guba dike field that comprises more than 30 minor gabbronorite intrusions and dikes of “garnet” Fe-gabbro was revealed at the northeastern flank of the Belomorian Mobile Belt. Gabbronorite intrusions were emplaced along the faults that formed as a result of brittle failure under conditions of general extension. As follows from the structural relationships, dikes of Fe-gabbro are younger than gabbronorite. They were formed under compression and shearing. The mobile melt was pumped into gneisses that underwent ductile deformation. Eclogitization in the Krasnaya Guba dike field was selective and affected only dikes of Fe-gabbro and minor gabbronorite intrusions and did not develop in country rocks. The bodies of Fe-gabbro are eclogitized almost completely, whereas gabbronorite is transformed into eclogite only along shear zones. Eclogites were formed after the complete solidification of intrusive bodies, and their localization is controlled by the concentration of deformations owing to the different elastoplastic properties of gabbroic rocks and country gneisses and migmatites. The different degree of fracturing in basic dikes and country gneisses resulted in the concentration of fluid flows in fracture systems within competent rocks. Late Svecofennian pegmatites (1.86 Ga) distinctly crosscut eclogites that experienced retrograde metamorphism. The eclogitic rocks from the Krasnaya Guba are the youngest known in the Belomorye. The age of eclogitization is 2.12 1.86 Ga. A new finding of eclogitic rocks in the Belomorye, as well as previously known similar rocks near Gridino Village, emphasize tectonic activity of the northwestern Umba Collision Zone. The eclogitization of basic rocks in Krasnaya Guba is related to their desilification in shearing zones in the course of interaction of plagioclase with ortho-and clinopyroxene with the formation of garnet-omphacite aggregate and removal of silica. The currently observed texture, structure, and chemical

  11. Cloudwater and ozone effects upon high elevation red spruce: A summary of study results from Whitetop Mountain, Virginia

    SciTech Connect

    Thornton, F.C.; Joslin, J.D.; Pier, P.A.

    1994-11-01

    This paper integrates the results of a number of studies on the effects of cloudwater and ozone (O{sub 3}) on red spruce (Picea rubens Sarg.) seedlings, saplings, and mature trees at Whitetop Mountain, VA, over a 3-yr period. These investigations consisted of (1) seedling chamber exclusion studies, (2) mature tree branch chamber exclusion studies, and (3) field experiments comparing responses of seedlings, saplings, and mature trees. The studies included treatments that : (1) excluded clouds and O{sub 3} (COE), (2) excluded clouds and had ambient O{sub 3} (CE), and (3) exposed plants to both clouds and O{sub 3} either with (CC) or without (AA) chambers. Seedlings and mature branches in the various treatments were compared with respect to growth rates, gas exchange rates, foliar nutrition, and chlorophyll and wax content. Soil solution, throughfall, and foliar responses of mature trees near the summit, receiving differing amounts of cloud exposure (low cloud and high cloud sites) were also monitored. Ozone was found to have minimal effects on the parameters measured, whereas cloudwater exposure was found to have adverse effects on several response parameters. Chambered seedlings that were exposed to cloudwater (AA and CC), and mature trees at the high cloud site had significantly lower foliar Ca and Mg concentrations than their counterparts, which were protected from exposure (seedlings) or received low cloud exposure (mature). A 3 to 5{degrees}C increase in cold tolerance was also measured in seedlings form which cloudwater was excluded. These findings suggest that cloudwater-mediated effects are currently having a negative impact on the health of red spruce, and may be involved in red spruce decline in the eastern USA. 62 refs., 2 figs., 5 tabs.

  12. Deep crustal structure off Akita, eastern margin of the Japan Sea, deduced from ocean bottom seismographic measurements

    NASA Astrophysics Data System (ADS)

    Nishizawa, Azusa; Asada, Akira

    1999-06-01

    A seismic exploration was carried out to detect the heterogeneity of the seismic wavespeed structure associated with the plate convergence at the eastern margin of the Japan Sea. Two airgun-OBS (ocean bottom seismograph) profiles were shot off Akita, Japan, where a seismic gap seems to exist but the location of the plate boundary has not been confirmed. One of the profiles was 60 km long, trending NNE-SSW, named Line OBS-9, at the northeastern end of the Yamato Basin and the other was 170 km long, trending WNW-ESE, Line NT95-1, parallel to the direction of the supposed plate convergence. The crustal structure beneath Line OBS-9 consists of six layers. The uppermost layer is sediment. Three layers are identified beneath the top sedimentary layer and their P wavespeeds are estimated to be 3.3-3.4 km/s, 5.1-5.4 km/s and 5.8-6.3 km/s, which corresponds to the upper crust. Underneath these layers, a layer with P wavespeed ranges of 6.3-7.2 km/s comprises the middle and lower crust. The depth of Moho is inferred to be 19 km. These wavespeed values are comparable with those of the present Japanese island arc, while the thickness of the crust is one-half of that of the Japanese arc. The crustal model supports the scenario that the Yamato Basin is formed by extension of the island arc. The crustal model for Line NT95-1 shows a transition from the extended island arc structure beneath the Yamato Basin to a thicker crust similar to the Japanese arc. P wavespeed heterogeneity related to the plate boundary is not detected. However, a significant change in the structural model along the profile is found around the region where the largest change in the seafloor topography exists. In that region, the wavespeeds in the middle crust have lower values than those of the neighboring area and the Moho begins to deepen towards the Japanese island arc. From comparison with the relationship between P wavespeed structure and aftershock distribution of the 1993 Hokkaido

  13. Genetic Diversity and Population Structure in Aromatic and Quality Rice (Oryza sativa L.) Landraces from North-Eastern India.

    PubMed

    Roy, Somnath; Banerjee, Amrita; Mawkhlieng, Bandapkuper; Misra, A K; Pattanayak, A; Harish, G D; Singh, S K; Ngachan, S V; Bansal, K C

    2015-01-01

    The North-eastern (NE) India, comprising of Arunachal Pradesh, Assam, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim and Tripura, possess diverse array of locally adapted non-Basmati aromatic germplasm. The germplasm collections from this region could serve as valuable resources in breeding for abiotic stress tolerance, grain yield and cooking/eating quality. To utilize such collections, however, breeders need information about the extent and distribution of genetic diversity present within collections. In this study, we report the result of population genetic analysis of 107 aromatic and quality rice accessions collected from different parts of NE India, as well as classified these accessions in the context of a set of structured global rice cultivars. A total of 322 alleles were amplified by 40 simple sequence repeat (SSR) markers with an average of 8.03 alleles per locus. Average gene diversity was 0.67. Population structure analysis revealed that NE Indian aromatic rice can be subdivided into three genetically distinct population clusters: P1, joha rice accessions from Assam, tai rices from Mizoram and those from Sikkim; P2, aromatic rice accessions from Nagaland; and P3, chakhao rice germplasm from Manipur [corrected]. Pair-wise FST between three groups varied from 0.223 (P1 vs P2) to 0.453 (P2 vs P3). With reference to the global classification of rice cultivars, two major groups (Indica and Japonica) were identified in NE Indian germplasm. The aromatic accessions from Assam, Manipur and Sikkim were assigned to the Indica group, while the accessions from Nagaland exhibited close association with Japonica. The tai accessions of Mizoram along with few chakhao accessions collected from the hill districts of Manipur were identified as admixed. The results highlight the importance of regional genetic studies for understanding diversification of aromatic rice in India. The data also suggest that there is scope for exploiting the genetic diversity of aromatic and

  14. Genetic Diversity and Population Structure in Aromatic and Quality Rice (Oryza sativa L.) Landraces from North-Eastern India

    PubMed Central

    Mawkhlieng, Bandapkuper; Misra, A. K.; Pattanayak, A.; Harish, G. D.; Singh, S. K.; Ngachan, S. V.; Bansal, K. C.

    2015-01-01

    The North-eastern (NE) India, comprising of Arunachal Pradesh, Assam, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim and Tripura, possess diverse array of locally adapted non-Basmati aromatic germplasm. The germplasm collections from this region could serve as valuable resources in breeding for abiotic stress tolerance, grain yield and cooking/eating quality. To utilize such collections, however, breeders need information about the extent and distribution of genetic diversity present within collections. In this study, we report the result of population genetic analysis of 107 aromatic and quality rice accessions collected from different parts of NE India, as well as classified these accessions in the context of a set of structured global rice cultivars. A total of 322 alleles were amplified by 40 simple sequence repeat (SSR) markers with an average of 8.03 alleles per locus. Average gene diversity was 0.67. Population structure analysis revealed that NE Indian aromatic rice can be subdivided into three genetically distinct population clusters: P1, joha rice accessions from Assam, tai rices from Mizoram and those from Sikkim; P2, chakhao rice germplasm from Manipur; and P3, aromatic rice accessions from Nagaland. Pair-wise FST between three groups varied from 0.223 (P1 vs P2) to 0.453 (P2 vs P3). With reference to the global classification of rice cultivars, two major groups (Indica and Japonica) were identified in NE Indian germplasm. The aromatic accessions from Assam, Manipur and Sikkim were assigned to the Indica group, while the accessions from Nagaland exhibited close association with Japonica. The tai accessions of Mizoram along with few chakhao accessions collected from the hill districts of Manipur were identified as admixed. The results highlight the importance of regional genetic studies for understanding diversification of aromatic rice in India. The data also suggest that there is scope for exploiting the genetic diversity of aromatic and quality rice

  15. Population genetic structure of a three-host tick, Amblyomma dissimile, in eastern Venezuela.

    PubMed

    Lampo, M; Rangel, Y; Mata, A

    1998-12-01

    Patterns of genetic variation for the tick Amblyomma dissimile were analyzed from a total of 200 ticks collected on 12 toads (Bufo marinus), 14 snakes (Boa constrictor), and 8 lizards (Iguana iguana) at 11 localities. The analyses were performed on electrophoretic data from 8 isozyme loci. Mean heterozygosity per locus was 6% (+/-3.1) per population. Differences in allelic frequencies among ticks from different individual hosts were the major source of genetic variability in this study. Host species was a smaller source of genetic variation. Genetic distances between localities varied according to which host species was present in each locality, and these appeared to be related to the extent of habitat overlap between host species. The smallest genetic distances between samples from different host species were recorded for I. iguana and B. constrictor. In contrast, the genetic distances between tick samples from B. marinus and either of the reptile species were significantly larger than between tick samples from this amphibian species. Ecological variables or the geographic distance did not explain the local patterns of differentiation observed in A. dissimile. Major genetic differences between island and mainland sites (0.03702) suggested an association between genetic distances and geographic isolation. The consistency between patterns of genetic variation and those of host home range overlap suggests that host dispersion is the main force structuring the genetic variation within this tick species.

  16. Abundance, distribution and size structure of Diadema antillarum (Echinodermata: Diadematidae) in South Eastern Cuban coral reefs.

    PubMed

    Martín Blanco, F; González Sansón, G; Pina Amargós, F; Clero Alonso, L

    2010-06-01

    The 1983-1984 mass mortality event of Diadema antillarum affected more than 93% of the total Caribbean population. Although there are no records about the status of Diadema populations before and after die-off on Cuban reefs, anecdotal information suggests that populations were struck. We analyzed spatial variation in the abundance and size structure of D. antillarum in 22 reefs sites in Jardines de la Reina, from June 2004 to September 2005. Counts of Diadema were performed in five 30x2 m transects at each sampling site and sampling time, and test diameters were measured in September 2005 at the same fore reefs. Abundances were higher at reef crests (mean densities 0.08-2.18 ind./m2), while reef slope populations reached a maximum site level of 0.13 ind./m2 at only one site and showed values up to three orders of magnitude lower than those from reef crests. Highest abundance occurred at the west margin of major channels between keys where larval recruitment seems to be favored by local oceanographic features and facilitated by the abundance of Echinometra lucunter. The size frequency distribution of D. antillarum indicates that recruitment began to be noticeable three years before September 2005, suggesting these populations were depleted in the past and they are recovering now.

  17. Abundance, distribution and size structure of Diadema antillarum (Echinodermata: Diadematidae) in South Eastern Cuban coral reefs.

    PubMed

    Martín Blanco, F; González Sansón, G; Pina Amargós, F; Clero Alonso, L

    2010-06-01

    The 1983-1984 mass mortality event of Diadema antillarum affected more than 93% of the total Caribbean population. Although there are no records about the status of Diadema populations before and after die-off on Cuban reefs, anecdotal information suggests that populations were struck. We analyzed spatial variation in the abundance and size structure of D. antillarum in 22 reefs sites in Jardines de la Reina, from June 2004 to September 2005. Counts of Diadema were performed in five 30x2 m transects at each sampling site and sampling time, and test diameters were measured in September 2005 at the same fore reefs. Abundances were higher at reef crests (mean densities 0.08-2.18 ind./m2), while reef slope populations reached a maximum site level of 0.13 ind./m2 at only one site and showed values up to three orders of magnitude lower than those from reef crests. Highest abundance occurred at the west margin of major channels between keys where larval recruitment seems to be favored by local oceanographic features and facilitated by the abundance of Echinometra lucunter. The size frequency distribution of D. antillarum indicates that recruitment began to be noticeable three years before September 2005, suggesting these populations were depleted in the past and they are recovering now. PMID:20527467

  18. Fatty acids as biomarkers for food web structure in the eastern North Pacific Ocean

    NASA Astrophysics Data System (ADS)

    Behrens, J.; Aluwihare, L.; Stephens, B. M.

    2015-12-01

    Resulting from a NSF funded REU program at Scripps Institution of Oceanography in 2015, this research utilized gas chromatography-mass spectrometry (GC-MS) to analyze the fatty acid composition of suspended particulate organic matter (POM) and zooplankton (ZP; primarily copepods). Samples analyzed for this study were collected simultaneously from surface waters approximately 9 miles off the coast of San Diego in June 2015. I was testing the hypothesis that essential fatty acids in ZP should reflect their diet, in particular, distinguishing contributions from a microbial versus traditional food web. Food web structure in this region of the ocean has been shown to be sensitive to climate change on inter-annual and longer timescales. Thus, a proxy that identifies restructuring of food webs would be useful for examining the response of ocean ecosystems to future climate change. Lipids were extracted from ZP and POM using a modified Bligh and Dyer method with sonication. Following saponification free fatty acids and other lipids were further purified using column chromatography. Polar functional groups in lipids were then methylated prior to GC-MS analysis. In addition, 2-dimensional GCxGC with time of flight MS was used to distinguish polyunsaturated fatty acid isomers. My poster will present initial findings of shared fatty acids of zooplankton and POM suspended material from the Northern Pacific collection site. Further research will be focused on analyzing the hydrogen isotope composition of fatty acids in zooplankton and suspended DOM obtained at the collection site to further characterize and increase certainty on the role of microbes and phytoplankton in the region's food-web to distinguish prokaryotic and eukaryotic sources.

  19. Geophysical Investigation and Reconstruction of Lithospheric Structure and its Control on Geology, Structure and Mineralisation in the Cordillera of Northern Canada and Eastern Alaska

    NASA Astrophysics Data System (ADS)

    Hayward, N.

    2015-12-01

    A reconstruction of the Tintina fault is applied to regional gravity, aeromagnetic and topographic data, facilitating the definition of E-W trending lineaments within the lower crust and/or mantle lithosphere, oblique to the dominantly NW-trending structure of the Cordilleran terranes. The lineaments, which are continuous to the Denali fault, exhibit a range of geophysical and geological signatures. They are interpreted to be related to the Liard transfer zone that divided lower and upper plates during late Proterozoic-Cambrian rifting of the Laurentian margin. Density models derived from the 3-D inversion of reconstructed gravity data reflect a change from NW-trending structure in the upper to middle crust, to E-W trending structure below. These deeper structures are associated with a small increase in the density of the lower crust and mantle lithosphere to the north. The transfer zone also divides bimodal mantle xenolith suites, derived from late Tertiary to Recent alkaline basalts, to the south from unimodal suites to the north. These conclusions suggest that extended North American basement, related to Laurentian margin rifting, that would have brought mantle lithosphere rocks to a shallow depth, continuously underlies a thin carapace of accreted terranes in western Yukon and eastern Alaska. The interpreted continuity of North American basement reaffirms that if oroclinal bending of the Intermontane terranes occurred, then it was prior to its emplacement upon the rifted basement. Examination of the spatial relationships between mineral occurrences and post-accretionary, Cretaceous lithospheric lineaments, from their manifestation in geophysical, geological, and topographic data, suggests that the late Proterozoic lineaments influenced Mesozoic mineralization through influence on the development of the shallow crustal structure, intrusion, and exhumation and erosion.

  20. Decadal-Scale Variations in Eastern Pacific Thermocline Structure from Soledad Basin, Baja California

    NASA Astrophysics Data System (ADS)

    Levi, C.; van Geen, A.; Ortiz, J. D.; Zheng, Y.; Marchitto, T. M.; Dean, W. E.; Carriquiry, J.

    2004-12-01

    Soledad Basin, a semi-enclosed basin on the Pacific margin of southern Baja California at 25oN, is ideally located to document past variations of ocean/atmosphere interactions responding to the Pacific Decadal Oscillation (PDO) and the El Nino-Southern Oscillation (ENSO). Very high sedimentation rates (~108 cm/kyr; van Geen et al., Paleoceanography, v. 8, no. 4, 2003) combined with low bottow-water oxygen levels have prevented sediment bioturbation throughout the Holocene, setting the stage for high-resolution paleoclimatic reconstructions in this key climatic area. Current studies of this site focus on the combination of a 210Pb-dated multicore and 14C-dated gravity and piston cores. Available data include a 1-cm resolution diffuse spectral reflectance record, indicative of diagenetic processes linked to productivity (Ortiz et al., Geology, v. 32, no. 6, 2004) and Mg/Ca ratios for several planktonic foraminifera species. Mg/Ca results for two shallow-dwelling species in the multicore indicate little change in sea surface temperature (+/-1.1 deg C) over the mean value of 23 deg C observed over the past two centuries. In contrast, variations of Mg/Ca ratios for a deeper-dwelling species suggest considerably larger temperature variations of +/-1.8 deg C with respect to the mean value of 17.5 deg C at the depth habitat of this species. This suggests fluctuations in the temperature, and therefore nutrient content, of thermocline waters upwelling toward the surface at this site without any appreciable changes in upwelling. Periods of low subsurface temperatures in the water column inferred from Mg/Ca correspond to darker sediment bands, suggesting a connection between the structure of the thermocline, the supply of nutrients to the photic zone, and surface productivity. This interpretation is supported by elevated authigenic Mo concentrations in those same dark bands of 30-40 mg/kg. The presentation will include a comparison of these results with instrumental and other

  1. Structural patterns and tectonic history of the Bauer microplate, Eastern Tropical Pacific

    USGS Publications Warehouse

    Eakins, B.W.; Lonsdale, P.F.

    2003-01-01

    motion. Some structures of the Bauer microplate boundary, such as deep rift valleys and a broad zone of thrust-faulted lithosphere, are, however, similar to those observed around the smaller, active microplates. Analysis of how the Bauer microplate was captured when coupling to the Pacific plate was reduced invites speculation on why risecrest microplates eventually lose their independence. ?? Springer 2005.

  2. Crustal seismic velocity structure from Eratosthenes Seamount to Hecataeus Rise across the Cyprus Arc, eastern Mediterranean

    NASA Astrophysics Data System (ADS)

    Welford, J. Kim; Hall, Jeremy; Hübscher, Christian; Reiche, Sönke; Louden, Keith

    2015-02-01

    Wide-angle reflection/refraction seismic profiles were recorded across the Cyprus Arc, the plate boundary between the African Plate and the Aegean-Anatolian microplate, from the Eratosthenes Seamount to the Hecataeus Rise immediately south of Cyprus. The resultant models were able to resolve detail of significant lateral velocity variations, though the deepest crust and Moho are not well resolved from the seismic data alone. Conclusions from the modelling suggest that (i) Eratosthenes Seamount consists of continental crust but exhibits a laterally variable velocity structure with a thicker middle crust and thinner lower crust to the northeast; (ii) the Hecataeus Rise has a thick sedimentary rock cover on an indeterminate crust (likely continental) and the crust is significantly thinner than Eratosthenes Seamount based on gravity modelling; (iii) high velocity basement blocks, coincident with highs in the magnetic field, occur in the deep water between Eratosthenes and Hecataeus, and are separated and bounded by deep low-velocity troughs and (iv) one of the high velocity blocks runs parallel to the Cyprus Arc, while the other two appear linked based on the magnetic data and run NW-SE, parallel to the margin of the Hecataeus Rise. The high velocity block beneath the edge of Eratosthenes Seamount is interpreted as an older magmatic intrusion while the linked high velocity blocks along Hecataeus Rise are interpreted as deformed remnant Tethyan oceanic crust or mafic intrusives from the NNW-SSE oriented transform margin marking the northern boundary of Eratosthenes Seamount. Eratosthenes Seamount, the northwestern limit of rifted continental crust from the Levant Margin, is part of a jagged rifted margin transected by transform faults on the northern edge of the lower African Plate that is being obliquely subducted under the Aegean-Anatolian upper plate. The thicker crust of Eratosthenes Seamount may be acting as an asperity on the subducting slab, locally locking up

  3. Ocean-continent-transition and oceanic ridge structural evolution (eastern Gulf of Aden): Implications for rift to seafloor spreading processes

    NASA Astrophysics Data System (ADS)

    D'Acremont, E.; Leroy, S. D.; Beslier, M.; Autin, J.; Watremez, L.; Maia, M. A.; Gente, P.

    2009-12-01

    The rifting between Arabia and Somalia, which started around 35 Ma ago, is followed by oceanic accretion from at least 17.6 Ma leading to the present Gulf of Aden. The transition between the thinned continental and the oceanic crusts is characterized, in space and time, by an ocean-continent transition (OCT). Here, we use bathymetry, gravity, seismic reflection and magnetism from the Encens-Sheba and Encens cruises in order to constrain the structure and segmentation of the conjugate OCT as well as the oceanic ridge between two main fracture zones (Alula-Fartak and Socotra-Hadbeen). The segmentation of the initial oceanic spreading centers seem directly related to the margin structure. Then, magmatic processes and kinematics change strongly influenced the evolution of the segmentation. The OCT and the oceanic domain can be divided into two distinct areas in the study area. The Eastern area is characterized by an extremely thin OCT and oceanic crusts (< 4km), a ~30 km wide and tectonized OCT with isolated continental blocks and short axial segments. In the western area, thicker OCT and oceanic crusts (>5km), a ~15 km wide OCT with a volcanic ridge, and a 6 km thick underplated mafic body in the northern margin suggest a high melt supply. The magmatic supply observed in the western domain is probably due to an off-axis thermal anomaly located below the southern flank of the Sheba ridge, at 75 km east of the major Alula-Fartak transform fault. This suggests that the OCT and the axial ridge morphology of this domain are perturbed by post-rift volcanism, which is due to a combination of the spreading rate, a thermal anomaly, and the cold edge effect of the Alula-Fartak transform fault. The presence of the inherited Mesozoic basins (Jezar-Qamar-Gardafui basin) located on this western domain can also explain, the difference in both the structure and the nature of the OCT between the two domains. The nature of the OCT could be either (or both) exhumed lower crust or

  4. Fish composition and assemblage structure in three Eastern English Channel macrotidal estuaries: A comparison with other French estuaries

    NASA Astrophysics Data System (ADS)

    Selleslagh, Jonathan; Amara, Rachid; Laffargue, Pascal; Lesourd, Sandric; Lepage, Mario; Girardin, Michel

    2009-01-01

    This study has analysed for the first time fish composition and assemblage structures of three small macrotidal estuaries of the Eastern English Channel (EEC) and has explored the influences of 19 biotic and abiotic variables on the fish assemblages. Fish from Canche, Authie and Somme estuaries were collected during spring (June 2006 and May 2007) and autumn (September 2006) along the estuarine gradients using a 1.5 m beam trawl. Using identical sampling protocols, the study also analysed and compared for the first time taxonomic and functional aspects of the fish assemblages in 15 estuaries located along the Atlantic and English Channel coasts. SIMPER analysis showed high similarities in fish assemblages in the three EEC estuaries and during either spring or autumn periods. However, intra-estuary similarities were relatively low, indicating that fish assemblage structures (species richnesses or abundances) were more variable within the estuary (salinity gradient) than between estuaries and/or seasons (spring vs autumn). Although numerous environmental variables were included in the study, only 47% of the variability observed in the fish distribution was explained. Fish spatial variations in the EEC estuaries are mostly driven by abiotic variables as opposed to biological interactions. As indicated by CCA, salinity and muddy sediments were the two most important factors structuring the fish assemblages. The macrobenthos being very abundant in the EEC estuaries (580-1121 ind. m -2), the availability of potential prey is probably not a limiting factor in the utilization of estuaries by fish. Contrary to the majority of French estuaries dominated by estuarine species (ES), the fish assemblages of the EEC estuaries are clearly dominated by marine migrant (MM) species (65% on average) with high abundance of juveniles (mostly young-of-the-year). Cluster and SIMPROF's analyses distinguished the functional structure of the 15 estuarine fish assemblages into different

  5. Hydraulic properties and inner structure of a relict rock glacier in the Eastern Alps, Austria

    NASA Astrophysics Data System (ADS)

    Pauritsch, Marcus; Winkler, Gerfried; Kellerer-Pirklbauer, Andreas; Birk, Steffen

    2013-04-01

    Water economic studies in 1990s documented the importance of the springs draining relict rock glaciers for water supply and human consumption as well as for the ecosystem in alpine catchments in the Niederen Tauern Range, Austria. Recent studies confirm the hydrologic importance and show that in the easternmost subunit, the Seckauer Tauern Range, more than 40% of the area above 2000 m a.s.l. and up to 20% of the area above 1500 m a.s.l. drain through relict rock glaciers. Thus, the hydraulic properties of these alpine aquifers are considered to be important controls on the hydrology of these areas. Nevertheless their hydraulic properties and their inner structure are still poorly understood. Our hydrogeological research is carried out at the Schöneben Rock Glacier, located in Seckauer Tauern Range, Austria. This rock glacier is presumably relict although patches of permafrost might exist particularly in the upper part of the landform. The rock glacier covers an area of 0.11 km² and drains a total catchment of 0.76 km² with a maximum elevation of 2282 m a.s.l.. The rock glacier consists predominantly of gneissic sediments (mainly coarse-grained, blocky at the surface) and extends from 1720 to 1905 m a.s.l.. Discharge of the rock glacier spring is recorded since 2002. Electrical conductivity and water temperature used as natural tracers are continuously monitored since 2008. Furthermore, a tracer test with simultaneous injection of the fluorescent dyes naphthionate and fluoresceine at two injection points (one close to the front and one close to the rooting zone of the rock glacier) was performed. Recession analysis of the spring hydrograph reveals similarities to the flow dynamics of karst springs. The results exhibit on the one hand a slow base flow recession indicating a high storage capacity and on the other hand sharp discharge peaks immediately after rainfall events referring to a high hydraulic conductivity. Applying different analytic runoff models, the

  6. Active tectonics of the Devils Mountain Fault and related structures, northern Puget Lowland and eastern Strait of Juan de Fuca region, Pacific Northwest

    USGS Publications Warehouse

    Johnson, Samuel Y.; Dadisman, Shawn V.; Mosher, David C.; Blakely, Richard J.; Childs, Jonathan R.

    2001-01-01

    Information from marine high-resolution and conventional seismic-reflection surveys, aeromagnetic mapping, coastal exposures of Pleistocene strata, and lithologic logs of water wells is used to assess the active tectonics of the northern Puget Lowland and eastern Strait of Juan de Fuca region of the Pacific Northwest. These data indicate that the Devils Mountain Fault and the newly recognized Strawberry Point and Utsalady Point faults are active structures and represent potential earthquake sources.

  7. Tectolinear interpretation of a 1:5,000,000 Landsat-1 mosaic compared with the structure of central and eastern United States

    USGS Publications Warehouse

    Kutina, Jan; Carter, William D.

    1978-01-01

    The pattern of lineaments and curvilinear features interpreted from a 1:5,000,000 mosaic of satellite images (Landsat-1 was superimposed on a simplified version of the Geological Map of the United States, 1:2,500,000 scale, showing the structural scheme of Central and Eastern United States. A comparison of the above two patterns, shown in Fig. 1, is presented in this paper.

  8. The impact of disturbance and ensuing forestry practices on Collembola in spruce forest stands

    NASA Astrophysics Data System (ADS)

    Čuchta, Peter

    2016-04-01

    Soil Collembola communities were investigated in spruce forest stands of the High Tatra Mts that had been heavily damaged by a windstorm in November 2004 and subsequently by a wildfire in July 2005. The study focused on the impact of these disturbances and forestry practices on collembolan community distribution and structure four years after the disturbance. Four different treatments were selected for this study: intact forest stands (REF), non-extracted windthrown stands (NEX), clear-cut windthrown stands (EXT) and burnt windthrown stands (FIR). From a total of 7,820 individuals, 72 species were identified. The highest total abundance mean was recorded in FIR stands followed by NEX and EXT stands and, surprisingly, the lowest in REF stands. The highest total species richness was observed in REF stands, followed by NEX stands and FIR stands and the lowest in EXT stands. In REF and NEX stands the most abundant species were Folsomia penicula and Tetracanthella fjellbergi, while in heavily damaged stands the most abundant was Anurophorus laricis. The present study shows the negative impact of windthrow on Collembola communities as reflected in decreased species richness and abundance. However, disturbance by fire caused a considerable increase in collembolan abundance three years after the event. Moreover, we found out that clearing of windthrown spruce forests after a windstorm is less favourable for communities of soil collembolans and slows down the recovery process.

  9. Early to Late Cenozoic structural inheritance of Paleozoic basement structures in the northern Alpine foreland: examples from eastern France and northern Switzerland

    NASA Astrophysics Data System (ADS)

    Madritsch, Herfried

    2014-05-01

    During his time at the Geological Institute of the University of Basel, Peter Ziegler was the main initiator of the EUCOR-URGENT project, a joint multi-disciplinary research and training programme aiming at a better understanding of seismic hazard, neotectonics and evolution of the Upper Rhine Graben and surrounding areas. Throughout the duration of the programme from 1999 to 2007 the EUCOR-URGENT network embraced more than 40 Ph.D. students, 20 Post-Docs and 18 senior researchers, who were based at one of the 25 involved universities or national organizations. Peter's natural drive, networking capabilities and scientific enthusiasm were without doubt the main reasons for this success story. The Rhine-Bresse Transfer Zone (RBTZ) in eastern France, one of the natural laboratories investigated within the EUCOR-URGENT framework, is a major segment of the European Cenozoic Rift system (Ziegler, 1992) and formed by structural inheritance of the pre-existing Late Paleozoic Burgundy Trough. The Mid-Eocene to Oligocene evolution of the sinistral transtensional RBTZ was kinematically linked to crustal extension across the Upper Rhine and Bresse Grabens (Lacombe et al., 1993). From the Early Miocene onward the RBTZ further evolved under the influence of the far field effects of the Alpine collision involving Late Miocene to Pliocene NW-ward propagation of the thin-skinned Jura Thrust Belt but also thick-skinned reactivation of the Late Paleozoic and Paleogene fault systems in the RBTZ. In fact, shortening throughout the RBTZ appears to be still mildly active, as is indicated by one of the very few clearly oblique-compressive focal mechanisms in the northern Alpine foreland and evidenced by geomorphologic investigations that yielded Late Quaternary folding of fluvial meanders in the area of Besançon (Madritsch et al. 2010). The Late Paleozoic Burgundy Trough as well as the Jura Thrust Belt continue eastward into northern Switzerland. In this area, reprocessed and newly

  10. Crustal and uppermost mantle structure of the eastern margin of the Yilgarn Craton (Australia) from passive seismic data

    NASA Astrophysics Data System (ADS)

    Sippl, Christian; Tkalčić, Hrvoje; Kennett, Brian; Spaggiari, Catherine; Gessner, Klaus

    2016-04-01

    The Yilgarn Craton in Western Australia is one of the largest units of Archean lithosphere on earth. Along its southern and southeastern margin, it is bounded by the Albany-Fraser Orogen (AFO), a Paleo- to Mesoproterozoic extensioal-accretionary orogen. In this contribution, we investigate the crustal and upper mantle structure of the AFO and adjacent regions using passive seismic data collected during the recent ALFREX experiment. Since the entire region has not been significantly reactivated since the Mesoproterozoic, the old signature of craton edge modification should have been well preserved until today. From November 2013 until January 2016, we operated a temporary passive seismic network consisting of 70 stations in the eastern Albany-Fraser Orogen. The array had an average station spacing of about 40 km and was designed to fill the gap between recently acquired active seismic profiles. We present results from the analysis of P receiver functions and ambient noise tomography using the ALFREX data. Receiver functions were used to derive a Moho depth map via H-K stacking, for direct imaging (common conversion point stacking) as well as joint inversion with surface wave dispersion data to derive 1D S-velocity profiles beneath the stations. The obtained receiver functions show a marked change of character from west to east across the array. Whereas they feature clear and sharp Moho phases for stations on the Yilgarn Craton, significantly more crustal complexity and fainter Moho phases are seen throughout the AFO. Crustal thickness increases from 36-39 km for the Yilgarn Craton to values between 42 and 48 km across the AFO, decreasing to around 40 km in the east. Ambient noise cross-correlations were used to derive maps of phase and group velocities of Rayleigh waves at periods between 1 and 30 seconds. A three-dimensional model of S wavespeeds throughout the area was then computed by pixelwise inversion of dispersion curves. Obtained S wavespeeds are generally

  11. Nesting ecology of boreal forest birds following a massive outbreak of spruce beetles

    USGS Publications Warehouse

    Matsuoka, S.M.; Handel, C.M.

    2007-01-01

    We studied breeding dark-eyed juncos (Junco hyemalis), yellow-rumped warblers (Dendroica coronata), and spruce-nesting birds from 1997 to 1998 among forests with different levels of spruce (Picea spp.) mortality following an outbreak of spruce beetles (Dendroctonus rufipennis) in Alaska, USA. We identified species using live and beetle-killed spruce for nest sites and monitored nests to determine how the outbreak influenced avian habitat selection and reproduction. We tested predictions that 1) nesting success of ground-nesting juncos would increase with spruce mortality due to proliferation of understory vegetation available to conceal nests from predators, 2) nesting success of canopy-nesting warblers would decrease with spruce mortality due to fewer live spruce in which to conceal nests, and 3) both species would alter nest-site selection in response to disturbance. Juncos did not benefit from changes in understory vegetation; nesting success in highly disturbed stands (46%) was comparable to that in undisturbed habitats throughout their range. In stands with low spruce mortality, nesting success of juncos was low (5%) and corresponded with high densities of red squirrels (Tamiasciurus hudsonicus). Yellow-rumped warblers nested exclusively in spruce, but success did not vary with spruce mortality. As disturbance increased, nesting warblers switched from selecting forest patches with high densities of live white spruce (Picea glauca) to patches with beetle-killed spruce. Warblers also placed nests in large-diameter live or beetle-killed spruce, depending on which was more abundant in the stand, with no differences in nesting success. Five of the 12 other species of spruce-nesting birds also used beetle-killed spruce as nest sites. Because beetle-killed spruce can remain standing for >50 years, even highly disturbed stands provide an important breeding resource for boreal forest birds. We recommend that boreal forest managers preserve uncut blocks of infested

  12. Crustal structure from the Hecataeus Rise to the Levantine Basin, eastern Mediterranean, from seismic refraction and gravity modelling

    NASA Astrophysics Data System (ADS)

    Welford, J. Kim; Hall, J.; Rahimi, A.; Reiche, S.; Hübscher, C.; Louden, K.

    2015-12-01

    In 2010, a wide-angle seismic reflection/refraction profile was acquired along the Hecataeus Rise, an area of shallow seabed immediately south of Cyprus in the eastern Mediterranean. The profile crossed from the Hecataeus Rise, through the Cyprus Arc to the Levantine Basin beyond. Due to the short length of the profile and the corresponding lack of deep ray coverage, velocity modelling was complemented by gravity modelling to gain constraints on deep crustal structure. The resultant model reveals velocities for the Hecataeus Rise that show no evidence of shallow ophiolites like those seen on mainland Cyprus, and the velocities are not diagnostic of a unique crustal affinity. Low-velocity sediments make up at least 7 km of the upper structure of Hecataeus Rise and these sediments overlie a two-layered crust. From the gravity modelling, the combined sediments and crust of Hecataeus Rise appear to be thinner than the Eratosthenes Seamount block to the southwest. A high-velocity lower crustal block is modelled under the seaward edge of Hecataeus Rise and, based on the gravity modelling, is inferred to extend landwards beneath the Rise. Similar high-velocity blocks were identified on the southwestern edge of Hecataeus Rise along nearby refraction lines and were interpreted as remnant Tethyan oceanic crust, foundered in the Cyprus Arc, along which subduction has ceased in this area. Given the thin two-layered crust beneath a thick accumulation of sediments modelled for Hecataeus Rise, we interpret that Hecataeus Rise represents a collage of oceanic fragments, accreted together within the failed subduction zone. Outboard of the crust of Hecataeus Rise, a 5-km deep low-velocity basin, possibly an accretionary wedge, is imaged that appears to correspond with the Cyprus Arc deformation zone imaged on both coincident and along-strike seismic reflection lines. A similar and wider feature is observed on seismic refraction lines to the west and combined, these may be revealing

  13. Cumulative potential hydrologic impacts of surface coal mining in the eastern Powder River structural basin, northeastern Wyoming

    USGS Publications Warehouse

    Martin, L.J.; Naftz, D.L.; Lowham, H.W.; Rankl, J.G.

    1988-01-01

    There are 16 existing and six proposed surface coal mines in the eastern Powder River structural basin of northeastern Wyoming. Coal mining companies predict water level declines of 5 ft or more in the Wasatch aquifer to extend form about 1,000 to about 2,000 ft beyond the mine pits. The predicted 5 ft water level decline in the Wyodak coal aquifer generally extends 4-8 mi beyond the lease areas. About 3,000 wells are in the area of potential cumulative water level declines resulting from all anticipated mining. Of these 3,000 wells, about 1,200 are outside the areas of anticipated mining: about 1,000 wells supply water for domestic or livestock uses, and about 200 wells supply water for municipal, industrial, irrigation, and miscellaneous uses. The 1,800 remaining wells are used by coal mining companies. Future surface coal mining probably will result in postmining groundwater of similar quality to that currently present in the study area. By use of geochemical modeling techniques, the results of a hypothetical reaction path exercise indicate the potential for marked improvements in postmining water quality because of chemical reactions as postmining groundwater with a large dissolved solids concentration (3,540 mg/L) moves into a coal aquifer with relatively small dissolved solids concentrations (910 mg/L). Results of the modeling exercise also indicate geochemical conditions that are most ideal for large decreases in dissolved solids concentrations in coal aquifers receiving recharge from a spoil aquifer. (Lantz-PTT)

  14. An integrated structural and geochemical study of fracture aperture growth in the Campito Formation of eastern California

    NASA Astrophysics Data System (ADS)

    Doungkaew, N.; Eichhubl, P.

    2015-12-01

    Processes of fracture formation control flow of fluid in the subsurface and the mechanical properties of the brittle crust. Understanding of fundamental fracture growth mechanisms is essential for understanding fracture formation and cementation in chemically reactive systems with implications for seismic and aseismic fault and fracture processes, migration of hydrocarbons, long-term CO2 storage, and geothermal energy production. A recent study on crack-seal veins in deeply buried sandstone of east Texas provided evidence for non-linear fracture growth, which is indicated by non-elliptical kinematic fracture aperture profiles. We hypothesize that similar non-linear fracture growth also occurs in other geologic settings, including under higher temperature where solution-precipitation reactions are kinetically favored. To test this hypothesis, we investigate processes of fracture growth in quartzitic sandstone of the Campito Formation, eastern California, by combining field structural observations, thin section petrography, and fluid inclusion microthermometry. Fracture aperture profile measurements of cemented opening-mode fractures show both elliptical and non-elliptical kinematic aperture profiles. In general, fractures that contain fibrous crack-seal cement have elliptical aperture profiles. Fractures filled with blocky cement have linear aperture profiles. Elliptical fracture aperture profiles are consistent with linear-elastic or plastic fracture mechanics. Linear aperture profiles may reflect aperture growth controlled by solution-precipitation creep, with the aperture distribution controlled by solution-precipitation kinetics. We hypothesize that synkinematic crack-seal cement preserves the elliptical aperture profiles of elastic fracture opening increments. Blocky cement, on the other hand, may form postkinematically relative to fracture opening, with fracture opening accommodated by continuous solution-precipitation creep.

  15. [Community stability for spruce-fir forest at different succession stages in Changbai Mountains, Northeast China].

    PubMed

    Zhang, Meng-tao; Zhang, Qing; Kang, Xin-gang; Yang, Ying-jun; Xu, Guang; Zhang, Li-xin

    2015-06-01

    Based on the analysis of three forest communities (polar-birch secondary forest, spruce-fir mixed forest, spruce-fir near pristine forest) in Changbai Mountains, a total of 22 factors of 5 indices, including the population regeneration, soil fertility (soil moisture and soli nutrient), woodland productivity and species diversity that reflected community characteristics were used to evaluate the stability of forest community succession at different stages by calculating subordinate function values of a model based on fuzzy mathematics. The results that the indices of population regeneration, soli nutrient, woodland productivity and species diversity were the highest in the spruce-fir mixed forest, and the indices of soil moisture were the highest in the spruce-fir near-pristine forest. The stability of three forest communities was in order of natural spruce-fir mixed forest > spruce-fir near pristine forest > polar-birch secondary forest.

  16. Productivity of the spruce grouse in fragmented habitat at the edge of its range

    USGS Publications Warehouse

    Whitcomb, S.D.; O'Connell, A.F.; Servello, F.A.

    1996-01-01

    We measured productivity of the Spruce Grouse (Dendragapusc anadensicsa nadensis) in patchy black spruce (Picea mariana) habitat along the southeastern limit of its range in mid-coastal Maine. We captured grouse and attached necklace-mounted radio transmitters to hens prior to nesting. Of 19 females monitored, only 26% raised chicks to the late brood-rearing period. Predation was high on hens (37%) and five were killed before hatching eggs. Six (55%) entire broods were lost and only 30% of chicks survived to late summer. Production ( No. of chicks/female), an index of productivity, was < 1 and lower in Maine and Minnesota study areas in black spruce than areas dominated by jack pine (Pinus banksiana) or a mixture of jack pine and spruce with dense undergrowth. Where Spruce Grouse breed in patchy black spruce communities, immigration from neighboring populations or inter-patch movement by local individuals may be required to maintain viable populations.

  17. [Community stability for spruce-fir forest at different succession stages in Changbai Mountains, Northeast China].

    PubMed

    Zhang, Meng-tao; Zhang, Qing; Kang, Xin-gang; Yang, Ying-jun; Xu, Guang; Zhang, Li-xin

    2015-06-01

    Based on the analysis of three forest communities (polar-birch secondary forest, spruce-fir mixed forest, spruce-fir near pristine forest) in Changbai Mountains, a total of 22 factors of 5 indices, including the population regeneration, soil fertility (soil moisture and soli nutrient), woodland productivity and species diversity that reflected community characteristics were used to evaluate the stability of forest community succession at different stages by calculating subordinate function values of a model based on fuzzy mathematics. The results that the indices of population regeneration, soli nutrient, woodland productivity and species diversity were the highest in the spruce-fir mixed forest, and the indices of soil moisture were the highest in the spruce-fir near-pristine forest. The stability of three forest communities was in order of natural spruce-fir mixed forest > spruce-fir near pristine forest > polar-birch secondary forest. PMID:26572010

  18. Hyperspectral data for assessment of temporal changes in Norway spruce forest conditions in the mountainous region of the Czech Republic affected by long-term acidic deposition

    NASA Astrophysics Data System (ADS)

    Albrechtova, J.; Lhotakova, Z.; Misurec, J.; Kopackova, V.; Campbell, P. K. E.; Edwards-Jonasova, M.; Kupkova, L.; Cervena, L.; Potuckova, M.; Cudlin, P.

    2015-12-01

    The Ore Mts. located in the western part of the Czech Republic suffered during 1950's-1990´s heavy atmospheric pollution due to the mining activities and brown coal combustion. Acidic deposition in combination with harsh climatic conditions led there to large-scale forest decline. Although the load of SO2 has significantly decreased since 1991, tree damage was still visible in 1998 in terms of high defoliation or dead trees. Nowadays Norway spruce trees do not exhibit visible symptoms of damage but the full recovery of Norway spruce forests is not complete yet due to persisting adverse soil conditions. The temporal changes in the physiological status of Norway spruce forests in the Krušné Hory Mts. were evaluated using two sets of spectral images acquired in 1998 (ASAS) and in 2013 (APEX) and ground truth data (LAI, tree crown status, photosynthetic pigment contents, leaf spectral properties measured by spectroradiometer, soil properties - pH, contents of basic cations, heavy metals, etc.). Ground truth data were evaluated by unconstrained and constrained multivariate analyses using Canoco 5. The high resolution spectral images (ASAS and APEX) enabled the identification of a gradient of forest conditions and their comparison. In 1998 the stands exhibited different physiological status corresponding to the pollution gradient with healthier trees at the western part of the mountains. Analysis of the foliar chemistry in 2013 show a slight improvement of the Norway spruce physiological status in the eastern part of the mountains while the status of the western-located stands slightly worsened. In 2013 we also studied the differences in soil geochemical conditions, which appeared to be less favorable in the western part of the mountains characterized by a low base cation contents in the top organic horizon and a very low pH (pH<3).

  19. Genetic structure and diversity of indigenous rice (Oryza sativa) varieties in the Eastern Himalayan region of Northeast India.

    PubMed

    Choudhury, Baharul; Khan, Mohamed Latif; Dayanandan, Selvadurai

    2013-12-01

    The Eastern Himalayan region of Northeast (NE) India is home to a large number of indigenous rice varieties, which may serve as a valuable genetic resource for future crop improvement to meet the ever-increasing demand for food production. However, these varieties are rapidly being lost due to changes in land-use and agricultural practices, which favor agronomically improved varieties. A detailed understanding of the genetic structure and diversity of indigenous rice varieties is crucial for efficient utilization of rice genetic resources and for developing suitable conservation strategies. To explore the genetic structure and diversity of rice varieties in NE India, we genotyped 300 individuals of 24 indigenous rice varieties representing sali, boro, jum and glutinous types, 5 agronomically improved varieties, and one wild rice species (O. rufipogon) using seven SSR markers. A total of 85 alleles and a very high level of gene diversity (0.776) were detected among the indigenous rice varieties of the region. Considerable level of genetic variation was found within indigenous varieties whereas improved varieties were monoporphic across all loci. The comparison of genetic diversity among different types of rice revealed that sali type possessed the highest gene diversity (0.747) followed by jum (0.627), glutinous (0.602) and boro (0.596) types of indigenous rice varieties, while the lowest diversity was detected in agronomically improved varieties (0.459). The AMOVA results showed that 66% of the variation was distributed among varieties indicating a very high level of genetic differentiation in rice varieties in the region. Two major genetically defined clusters corresponding to indica and japonica groups were detected in rice varieties of the region. Overall, traditionally cultivated indigenous rice varieties in NE India showed high levels of genetic diversity comparable to levels of genetic diversity reported from wild rice populations in various parts of the

  20. Seismically imaging the structural legacy of rifting and collision events in the central and eastern U.S. crust

    NASA Astrophysics Data System (ADS)

    Schmandt, B.; Lin, F. C.; Karlstrom, K. E.

    2015-12-01

    EarthScope's USArray now provides broadband seismic data across the contiguous U.S. and southeastern Canada. We used teleseismic receiver functions and surface wave tomography to map crustal structure beneath the entire array. Crust thickness was estimated with multi-mode Ps receiver function images using <0.5 Hz Ps and <0.25 Hz 2p1s and 2s1p reverberations between the free-surface and Moho. In areas of sedimentary basins or large impedance contrasts in the middle crust the reverberations alone often provide clearer images of the Moho than the Ps mode, because of interference from conversions at shallow interfaces is reduced at greater lag times. The new results enable large-scale comparison of the structural legacy of multiple rifting and collision events in eastern North America. Some Proterozoic rift segments defined by Bouguer gravity and surface geology maintain locally thin crust while others lack correlated Moho topography or are areas of locally thicker crust. Locally thin crust is found at southern end of the mid-continent rift (MCR) in northern Kansas and southern Nebraska, along the Reelfoot rift, and beneath inferred rifts in Michigan, Indiana, and Ohio. The Oklahoma aulacogen is not associated with a coherent change in crust thickness along its length, at least at a scale resolvable by USArray data and our imaging approach. The MCR extending northeast from Nebraska to Lake Superior has locally thicker crust, consistent with other recent results. We suggest that magmatic additions to the lower crust overwhelmed extension in the northern mid-continent rift, but not the rift segments further south and east. Collision events of the Grenville orogeny and Paleozoic orogens that created the Appalachian Mountains are still associated with ~45-55 km thick crust extending from the Grenville front eastward across the Appalachian Mountains to the fall line that marks the abrupt geomorphic transition to the coastal plains. Despite the ~45-55 km crust thickness long

  1. Exome capture from the spruce and pine giga-genomes.

    PubMed

    Suren, H; Hodgins, K A; Yeaman, S; Nurkowski, K A; Smets, P; Rieseberg, L H; Aitken, S N; Holliday, J A

    2016-09-01

    Sequence capture is a flexible tool for generating reduced representation libraries, particularly in species with massive genomes. We used an exome capture approach to sequence the gene space of two of the dominant species in Canadian boreal and montane forests - interior spruce (Picea glauca x engelmanii) and lodgepole pine (Pinus contorta). Transcriptome data generated with RNA-seq were coupled with draft genome sequences to design baits corresponding to 26 824 genes from pine and 28 649 genes from spruce. A total of 579 samples for spruce and 631 samples for pine were included, as well as two pine congeners and six spruce congeners. More than 50% of targeted regions were sequenced at >10× depth in each species, while ~12% captured near-target regions within 500 bp of a bait position were sequenced to a depth >10×. Much of our read data arose from off-target regions, which was likely due to the fragmented and incomplete nature of the draft genome assemblies. Capture in general was successful for the related species, suggesting that baits designed for a single species are likely to successfully capture sequences from congeners. From these data, we called approximately 10 million SNPs and INDELs in each species from coding regions, introns, untranslated and flanking regions, as well as from the intergenic space. Our study demonstrates the utility of sequence capture for resequencing in complex conifer genomes, suggests guidelines for improving capture efficiency and provides a rich resource of genetic variants for studies of selection and local adaptation in these species. PMID:27428061

  2. Highly Informative Single-Copy Nuclear Microsatellite DNA Markers Developed Using an AFLP-SSR Approach in Black Spruce (Picea mariana) and Red Spruce (P. rubens)

    PubMed Central

    Shi, Yong-Zhong; Forneris, Natascha; Rajora, Om P.

    2014-01-01

    Background Microsatellites or simple sequence repeats (SSRs) are highly informative molecular markers for various biological studies in plants. In spruce (Picea) and other conifers, the development of single-copy polymorphic genomic microsatellite markers is quite difficult, owing primarily to the large genome size and predominance of repetitive DNA sequences throughout the genome. We have developed highly informative single-locus genomic microsatellite markers in black spruce (Picea mariana) and red spruce (Picea rubens) using a simple but efficient method based on a combination of AFLP and microsatellite technologies. Principal Findings A microsatellite-enriched library was constructed from genomic AFLP DNA fragments of black spruce. Sequencing of the 108 putative SSR-containing clones provided 94 unique sequences with microsatellites. Twenty-two of the designed 34 primer pairs yielded scorable amplicons, with single-locus patterns. Fourteen of these microsatellite markers were characterized in 30 black spruce and 30 red spruce individuals drawn from many populations. The number of alleles at a polymorphic locus ranged from 2 to 18, with a mean of 9.3 in black spruce, and from 3 to 15, with a mean of 6.2 alleles in red spruce. The polymorphic information content or expected heterozygosity ranged from 0.340 to 0.909 (mean = 0.67) in black spruce and from 0.161 to 0.851 (mean = 0.62) in red spruce. Ten SSR markers showing inter-parental polymorphism inherited in a single-locus Mendelian mode, with two cases of distorted segregation. Primer pairs for almost all polymorphic SSR loci resolved microsatellites of comparable size in Picea glauca, P. engelmannii, P. sitchensis, and P. abies. Significance The AFLP-based microsatellite-enriched library appears to be a rapid, cost-effective approach for isolating and developing single-locus informative genomic microsatellite markers in black spruce. The markers developed should be useful in black spruce, red spruce

  3. Alkaline intrusion in a granulite ensemble in the eastern ghats belt, India: Shear zone pathway and a pull-apart structure

    NASA Astrophysics Data System (ADS)

    Bhattacharya, S.; Kar, Rajib

    2004-03-01

    The alkaline complex of Koraput, Orissa, India, is one of several bodies in the high-grade Eastern Ghats belt, but this one is an integral part of the high-grade belt and remote from the western boundary against the Bastar craton. The Koraput complex forms a lozenge-shaped intrusion into the metapelitic granulites and is bounded by shear zones. The combined effect of movement along these shear zones, is a northeasterly elongated sygmoidal cavity with maximum width along the northwesterly trending Reidel shear. Thus the Koraput alkaline complex can be considered to have been emplaced in a pull-apart structure, developed in the granulitic country rocks. Moreover, in view of the fact that the western margin of the high-grade Eastern Ghats belt bears clear evidence of collisional features, rather than that of rifting or break-up, the rift-valley model for the alkaline magmatism in this high-grade belt appears untenable.

  4. Can repeated fertilizer applications to young Norway spruce enhance avian diversity in intensively managed forests?

    PubMed

    Edenius, Lars; Mikusiński, Grzegorz; Bergh, Johan

    2011-07-01

    Repeated fertilization of forests to increase biomass production is an environmentally controversial proposal, the effects of which we assessed on breeding birds in stands of young Norway spruce (Picea abies), in an intensively managed forest area in southern Sweden. Our results show that fertilized stands had 38% more species and 21% more individuals than unfertilized stands. Compared with stands under traditional management, the further intensification of forestry by repeated applications of fertilizers thus seemed to enhance species richness and abundance of forest birds. We cannot conclude at this stage whether the response in the bird community was caused by changes in food resources or increased structural complexity in the forest canopy due to the skid roads used for the application of the fertilizers. Future studies should focus on structural and compositional effects of fertilization processes during the entire rotation period and at assessing its effects in a landscape PMID:21848140

  5. Possible red spruce decline: Contributions of tree-ring analysis

    SciTech Connect

    Van Deusen, P.C.; Reams, G.A.; Cook, E.R.

    1991-01-01

    In studies of the northeastern red spruce ecosystem, several points evoke some agreement: (1) many high-elevation sites exhibit substantial post-1960 mortality that could be due to winter injury; (2) there is widespread pre-1950 growth increase with a subsequent post-1960 growth decrease; (3) many locations show none of these effects; and (4) dendro-climate models suggest that late summer and early winter temperatures of the previous year are significant determinants of red spruce year-to-year growth. It is a mistake to advocate a single cause at this time, because there is compelling evidence that both climate and stand dynamics are involved to some degree. The study of long-term forest trends based on tree-ring data is difficult and subject to interpretation, but it is unlikely that other data is available for most natural forest areas. In fact, progress made in the study of northeastern red spruce owes much to tree-ring analysis, and other ecological studies could benefit as well.

  6. Wood energy fuel cycle optimization in beech and spruce forests

    NASA Astrophysics Data System (ADS)

    Meyer, Nickolas K.; Mina, Marco

    2012-03-01

    A novel synergistic approach to reducing emissions from residential wood combustion (RWC) is presented. Wood energy fuel cycle optimization (FCO) aims to provide cleaner burning fuels through optimization of forestry and renewable energy management practices. In this work, beech and spruce forests of average and high quality were modelled and analysed to determine the volume of fuel wood and its associated bark fraction produced during typical forestry cycles. Two separate fuel wood bark production regimes were observed for beech trees, while only one production regime was observed for spruce. The single tree and stand models were combined with existing thinning parameters to replicate existing management practices. Utilizing estimates of initial seedling numbers and existing thinning patterns a dynamic model was formed that responded to changes in thinning practices. By varying the thinning parameters, this model enabled optimization of the forestry practices for the reduction of bark impurities in the fuel wood supply chain. Beech forestry cycles responded well to fuel cycle optimization with volume reductions of bark from fuel wood of between ˜10% and ˜20% for average and high quality forest stands. Spruce, on the other hand, was fairly insensitive to FCO with bark reductions of 0-5%. The responsiveness of beech to FCO further supports its status as the preferred RWC fuel in Switzerland. FCO could easily be extended beyond Switzerland and applied across continental Europe and North America.

  7. Kinematic evolution of Andean fold-thrust structures along the boundary between the Eastern Cordillera and Middle Magdalena Valley basin, Colombia

    NASA Astrophysics Data System (ADS)

    SáNchez, Javier; Horton, Brian K.; Tesón, Eliseo; Mora, AndréS.; Ketcham, Richard A.; Stockli, Daniel F.

    2012-06-01

    Surface and subsurface data support a kinematic reconstruction of Cenozoic fold-thrust deformation along the Eastern Cordillera-Magdalena Valley transition in Colombia. The La Salina fault (LSF) marks the boundary between west-vergent Eastern Cordillera structures and hinterland deposits of the Middle Magdalena Valley basin. Apatite fission track and (U-Th)/He thermochronological results for the west-directed LSF reveal initial hanging wall exhumation during middle Eocene-early Oligocene (45-30 Ma) shortening, renewed exhumation in the early middle Miocene (18-12 Ma), and accelerated late Miocene-Pliocene (12-3 Ma) exhumation. Vitrinite reflectance data suggest maximum burial of 4-6 km, helping constrain Cenozoic basin architecture. Mapping of the LSF reveals hanging wall Cretaceous-Eocene rocks in a broad anticline-syncline pair with limited faulting and footwall Eocene-Quaternary basin fill in a complex series of tight thrust-related folds. Limited displacement along the westernmost (frontal) thrust suggests that shortening is largely accommodated by east-directed thrusting within a broader triangle zone of a passive-roof duplex (and probable minor strike-slip deformation). In the preferred kinematic restoration, the most recent phase of shortening to transpressional deformation represents out-of-sequence reactivation of the LSF consistent with irregular crosscutting relationships among footwall structures. Earliest exhumation by 45-30 Ma in the Eastern Cordillera fold-thrust belt is correlated with increased sedimentary lithic fragments and high compositional maturity in sandstones of the adjacent Magdalena Valley basin. Exhumation since ˜15 Ma coincided with decreased compositional maturity and elevated accumulation rates for the Real Group. The compositional provenance shifts are attributed to westward advance of fold-thrust deformation into the proximal (eastern) segments of the Magdalena Valley basin.

  8. Numerical simulation of structural evolution from regional to local scale in the Outokumpu ore district, eastern Finland

    NASA Astrophysics Data System (ADS)

    Zhang, Yanhua; Sorjonen-Ward, Peter; Ord, Alison; Kontinen, Asko

    2015-04-01

    Numerical simulations of geological processes may be used in several ways. On the one hand there is an analytical, or forensic approach, analogous to geophysical inversion, to constrain boundary conditions and to demonstrate how a particular geological process or sequence of events is feasible, or even probable. Alternatively, or additionally, modeling of earth processes can be used in a predictive sense, where forward modeling of various scenarios representing different initial states and applied boundary conditions and processes can provide generic or specific insights - depending on model complexity - which may be applied to problems as diverse as geohazard risk assessment and mineral exploration. These two approaches are complementary, and either may be emphasized, depending on the degree of understanding or density of data in a given study area. Here we review how the results of modeling can be used to develop and test structural scenarios and hypotheses and how they can be integrated with new data sets, in this case, deep crustal and upper crustal high resolution reflection seismic data acquired in recent years in the Paleoproterozoic Outokumpu ore district in eastern Finland. A range of process models have been devised and run for the Outokumpu mineral system, including coupled convective reactive transport models, coupled thermomechanical models assessing thermal regimes in rifting, and coupled mechanical and fluid flow models, but here we focus on the results of mechanical modeling using the finite element code FLAC. Models designed at different scales have provided simple and plausible solutions that affirm the geometric and kinematic scenarios based on regional and mine-scale structural data. At regional scale, FLAC models effectively simulated the partitioning of deformation into NW-SE trending ductile shear zones and domains where coeval folding and thrusting have NE-trending axial trends. At a more detailed district scale, development of local

  9. Fire Severity Controlled Susceptibility to a 1940s Spruce Beetle Outbreak in Colorado, USA.

    PubMed

    Kulakowski, Dominik; Veblen, Thomas T; Bebi, Peter

    2016-01-01

    The frequency, magnitude, and size of forest disturbances are increasing globally. Much recent research has focused on how the occurrence of one disturbance may affect susceptibility to subsequent disturbances. While much has been learned about such linked disturbances, the strength of the interactions is likely to be contingent on the severity of disturbances as well as climatic conditions, both of which can affect disturbance intensity and tree resistance to disturbances. Subalpine forests in western Colorado were affected by extensive and severe wildfires in the late 19th century and an extensive and severe outbreak of spruce beetle (Dendroctonus rufipennis) in the 1940s. Previous research found that most, but not all, of the stands that burned and established following the late 19th century fires were not susceptible to the 1940s outbreak as beetles preferentially attack larger trees and stands in advanced stages of development. However, previous research also left open the possibility that some stands that burned and established following the 19th century fires may have been attacked during the 1940s outbreak. Understanding how strongly stand structure, as shaped by disturbances of varying severity, affected susceptibility to past outbreaks is important to provide a baseline for assessing the degree to which recent climate change may be relaxing the preferences of beetles for larger trees and for stands in latter stages of structural development and thereby changing the nature of linked disturbances. Here, dendroecological methods were used to study disturbance history and tree age of stands in the White River National Forest in Western Colorado that were identified in historical documents or remotely-sensed images as having burned in the 19th century and having been attacked by spruce beetle in the 1940s. Dendroecological reconstructions indicate that in young post-fire stands only old remnant trees that survived the otherwise stand-replacing fires were

  10. Fire Severity Controlled Susceptibility to a 1940s Spruce Beetle Outbreak in Colorado, USA

    PubMed Central

    Kulakowski, Dominik; Veblen, Thomas T.; Bebi, Peter

    2016-01-01

    The frequency, magnitude, and size of forest disturbances are increasing globally. Much recent research has focused on how the occurrence of one disturbance may affect susceptibility to subsequent disturbances. While much has been learned about such linked disturbances, the strength of the interactions is likely to be contingent on the severity of disturbances as well as climatic conditions, both of which can affect disturbance intensity and tree resistance to disturbances. Subalpine forests in western Colorado were affected by extensive and severe wildfires in the late 19th century and an extensive and severe outbreak of spruce beetle (Dendroctonus rufipennis) in the 1940s. Previous research found that most, but not all, of the stands that burned and established following the late 19th century fires were not susceptible to the 1940s outbreak as beetles preferentially attack larger trees and stands in advanced stages of development. However, previous research also left open the possibility that some stands that burned and established following the 19th century fires may have been attacked during the 1940s outbreak. Understanding how strongly stand structure, as shaped by disturbances of varying severity, affected susceptibility to past outbreaks is important to provide a baseline for assessing the degree to which recent climate change may be relaxing the preferences of beetles for larger trees and for stands in latter stages of structural development and thereby changing the nature of linked disturbances. Here, dendroecological methods were used to study disturbance history and tree age of stands in the White River National Forest in Western Colorado that were identified in historical documents or remotely-sensed images as having burned in the 19th century and having been attacked by spruce beetle in the 1940s. Dendroecological reconstructions indicate that in young post-fire stands only old remnant trees that survived the otherwise stand-replacing fires were

  11. Fire Severity Controlled Susceptibility to a 1940s Spruce Beetle Outbreak in Colorado, USA.

    PubMed

    Kulakowski, Dominik; Veblen, Thomas T; Bebi, Peter

    2016-01-01

    The frequency, magnitude, and size of forest disturbances are increasing globally. Much recent research has focused on how the occurrence of one disturbance may affect susceptibility to subsequent disturbances. While much has been learned about such linked disturbances, the strength of the interactions is likely to be contingent on the severity of disturbances as well as climatic conditions, both of which can affect disturbance intensity and tree resistance to disturbances. Subalpine forests in western Colorado were affected by extensive and severe wildfires in the late 19th century and an extensive and severe outbreak of spruce beetle (Dendroctonus rufipennis) in the 1940s. Previous research found that most, but not all, of the stands that burned and established following the late 19th century fires were not susceptible to the 1940s outbreak as beetles preferentially attack larger trees and stands in advanced stages of development. However, previous research also left open the possibility that some stands that burned and established following the 19th century fires may have been attacked during the 1940s outbreak. Understanding how strongly stand structure, as shaped by disturbances of varying severity, affected susceptibility to past outbreaks is important to provide a baseline for assessing the degree to which recent climate change may be relaxing the preferences of beetles for larger trees and for stands in latter stages of structural development and thereby changing the nature of linked disturbances. Here, dendroecological methods were used to study disturbance history and tree age of stands in the White River National Forest in Western Colorado that were identified in historical documents or remotely-sensed images as having burned in the 19th century and having been attacked by spruce beetle in the 1940s. Dendroecological reconstructions indicate that in young post-fire stands only old remnant trees that survived the otherwise stand-replacing fires were

  12. Observations of oceanic crust and mantle structures at a deep ocean seismic array in the Eastern Mid Atlantic

    NASA Astrophysics Data System (ADS)

    Hannemann, Katrin; Krüger, Frank; Dahm, Torsten

    2016-04-01

    In 2011, twelve ocean bottom stations (OBS) were installed approximately 100 km North of the Gloria Fault during the DOCTAR project (Deep OCean Test ARray). This fault marks the plate boundary between the Eurasian and African plate in the North Eastern Mid Atlantic. The experiment took place in water depth of 4-6 km, 800 km West of the Portuguese coast. The stations were equipped with broad band seismometers which recorded for ten months. We employ P and S receiver functions (RF) to have a closer look at the structure of crust and mantle. The ocean is a quite noisy environment, therefore the number of usable events is low (around 20) compared to RF studies on land. We use several quality criteria (e.g. signal to noise ratio, relative spike position) to select proper processing parameters for the calculation of the RF and carefully reviewed all later on used RF. Despite the low number of events, the usage of an array of OBS with an aperture of 75 km allows us to investigate deeper discontinuities (e.g. in 410 and 660 km depth) compared to single station approaches which are usually employed for OBS. Furthermore, we increase the number of usable events by applying array methods. We use move out corrected and stacked RF to have a closer look at the mantle transition zone, and estimate average depth values for the Moho, the lithosphere asthenosphere boundary (LAB) and the base of the asthenosphere. The Moho lies at depth of 7 km, the LAB at approximately 50 km and the asthenosphere has an approximated thickness of 110 km. We observe a slight increase in the time difference of the mantle discontinuity conversion times compared to PREM. RF give just information regarding the impedance contrast at a discontinuity instead of velocities. We additionally use P wave polarization of teleseismic events to estimate absolute S velocities beneath the single stations. All in all, we use the information gained by the RF analysis, and the analysis of the P wave polarization to

  13. Effects of NaCl on responses of ectomycorrhizal black spruce (Picea mariana), white spruce (Picea glauca) and jack pine (Pinus banksiana) to fluoride.

    PubMed

    Calvo-Polanco, Mónica; Zwiazek, Janusz J; Jones, Melanie D; MacKinnon, Michael D

    2009-01-01

    Black spruce (Picea mariana), white spruce (Picea glauca) and jack pine (Pinus banksiana) were inoculated with Suillus tomentosus and subjected to potassium fluoride (1 mM KF and 5 mM KF) in the presence and absence of 60 mM NaCl. The NaCl and KF treatments reduced total dry weights in jack pine and black spruce seedlings, but they did not affect total dry weights in white spruce seedlings. The addition of 60 mM NaCl to KF treatment solutions alleviated fluoride-induced needle injury in ectomycorrhizal (ECM) black spruce and white spruce, but had little effect in jack pine seedlings. Both KF and 60 mM NaCl treatments reduced E values compared with non-treated control seedlings. However, with the exception of small reductions of K(r) by NaCl treatments in black spruce, the applied KF and NaCl treatments had little effect on K(r) in ECM plants. Chloride tissue concentrations in NaCl-treated plants were not affected by the presence of KF in treatment solutions. However, shoot F concentrations in ECM black spruce and white spruce treated with 5 mM KF + 60 mM NaCl were significantly reduced compared with the 5 mM KF treatment. The results point to a possible competitive inhibition of F transport by Cl. We also suggest that the possibility that aquaporins may be involved in the transmembrane transport of F should be further investigated.

  14. Evaluation of genetic variability in a small, insular population of spruce grouse

    USGS Publications Warehouse

    O'Connell, A.F.; Rhymer, J.; Keppie, D.M.; Svenson, K.L.; Paigan, B.J.

    2002-01-01

    Using microsatellite markers we determined genetic variability for two populations of spruce grouse in eastern North America, one on a coastal Maine island where breeding habitat is limited and highly fragmented, the other in central New Brunswick (NB), where suitable breeding habitat is generally contiguous across the region. We examined six markers for both populations and all were polymorphic. Although the number of alleles per locus and the proportion of unique alleles were lower in the island population, and probably a result of small sample.size, heterozygosity and a breeding coefficient (Fis) indicated slightly more variability in the island population. Deviation from Hardy-Weinberg equilibrium also was more evident in loci for the mainland population. Several traits previously documented in the island population: relatively long natal dispersal distances, reproductive success, territoriality, adult survival, and longevity support the maintenance of hetrerzygosity, at least in the short-term. Sample collection from two small (500 ha), separate areas in NB, and the predicted importance of immigration density to supplement this population demonstrate the need for behavioral and ecological information when interpreting genetic variation. We discuss the relevance of these issues with respect to genetic variability and viability.

  15. Doctoring in Eastern Europe

    PubMed Central

    Wilde, Henry

    1983-01-01

    Health care in Eastern Europe has not achieved world standards nor the goals of planners of socialist societies. With luck, perseverance, bribes or good connections, it is possible to obtain good medical and surgical care in Eastern Europe for a major illness. Primary and even secondary care usually are substandard, however, and often completely unacceptable to most Western foreigners. The reasons for this are complex but mainly rooted in different attitudes of health workers towards their patients, poor physical plants, poor salary structures, inadequate advancement opportunities for health care workers, poor social status and professional recognition for nurses and almost complete isolation of the average primary care doctor from hospital medicine. PMID:6659504

  16. Magnetic fabrics and Pan-African structural evolution in the Najd Fault corridor in the Eastern Desert of Egypt

    NASA Astrophysics Data System (ADS)

    Abdeen, Mamdouh M.; Greiling, Reinhard O.; Sadek, Mohamed F.; Hamad, Sayed S.

    2014-11-01

    In order to assess the Pan-African structural evolution from early orogenic fabrics through Najd wrenching to the latest orogenic collapse/extension, the authors used field work, aided by aerial photographs and satellite images. This work is complemented by the study of the anisotropy of the magnetic susceptibility (AMS, or magnetic fabric). The Pan-African rock associations of the Um Gheig-Kadabora area can be divided into a lower tier composed mainly of amphibolite-migmatite and granitoid gneisses, and an upper tier of ophiolitic rocks, metavolcanics and their related volcaniclastics, and molasse-type Hammamat sediments. Both these units are intruded by late orogenic granitoid plutons and dykes. The lower tier is exposed in a domal structure in the El Sibai area, the upper tier forms a series of weakly to highly deformed thrust units, called Pan-African Nappes here, which are dissected by high strain shear zones. According to their age, these rock units are divided here into early and late-orogenic. The early orogenic rock association is characterized by medium-high metamorphic grades. The late orogenic rock association is characterized by low metamorphic grade. The rocks in the upper tier form a series of low angle thrust sheets, which are bounded by NW-striking high angle shear zones related to the Najd Fault System. The early orogenic rocks show a polyphase structural evolution with early folds, thrusts, and strike-slip shear zones. The late orogenic rocks show a relatively weaker deformation. The latest intrusives studied here are the dykes dissecting the late orogenic Kadabora granite. In the present work magnetic fabric data document the deformational features in detail and assess the role of the Najd Fault System in the deformational evolution. A strong variation in volume susceptibility of various rocks, due to their variations in mineral composition, is observed. Lower values are in the range of 10-6 SI units for late-orogenic alkaline granite and the

  17. Integrated well log and 2-D seismic data interpretation to image the subsurface stratigraphy and structure in north-eastern Bornu (Chad) basin

    NASA Astrophysics Data System (ADS)

    Isyaku, Aminu A.; Rust, Derek; Teeuw, Richard; Whitworth, Malcolm

    2016-09-01

    Structural and stratigraphic mapping within the Bornu Basin in north east Nigeria was commonly carried out using traditional field geological methods. However, such traditional approaches remain inadequate in the semi-arid region characterised by topographically flat areas and lack of continuous bedrock outcrops that are mostly concealed beneath sand cover. Previous studies in the north-eastern part of the basin carried out using ditch cuttings from few wells and disconnected seismic data were largely inadequate and the resulting stratigraphic analyses were more often generalised. This paper presents an integrated structural and stratigraphic study of the basin using combined subsurface geophysical datasets. A Combined Log Pattern (CLP) method is a well log analysis, which utilises various well log data including gamma ray, resistivity, bulk density and sonic logs to identify lithology and stratigraphic boundaries of subsurface formations. This method is applied to constrain the subsurface stratigraphy of the north-eastern part of the Bornu Basin bordering the Lake Chad. In addition to qualitative combined well log analysis, the time-depth relationship of the sonic log and seismic data was quantitatively determined by tying a well with an intersecting seismic section to validate the stratigraphic facies horizons identified. Four well log facies and their environments of deposition were characterised from the combined well log analysis of the different log types. It is discovered that the Cretaceous basement structural features controlled the deposition of overlying formations in the basin. Without intact core data, the shallower wells were discovered to have bottomed over subsurface horst features while deeper wells penetrated into the basal facies contained mainly within the grabens. Main subsurface structural lineaments in the area include NW-SE, NE-SW and NNW-SSE trending faults, which mainly formed the horst and graben features. Some stratigraphic formations

  18. Structural evolution and Cenozoic tectonostratigraphy of the Cairo-Suez district, north Eastern Desert of Egypt: Field-structural data from Gebel Qattamiya-Gebel Um Reheiat area

    NASA Astrophysics Data System (ADS)

    Hagag, Wael

    2016-06-01

    Detailed field mapping reveals that continental rifting is strongly deforming the Gebel Qattamiya-Gebel Um Reheiat area and the entire Cairo-Suez district, in north Eastern Desert of Egypt. Rift-related structures are predominantly represented by E to WNW, NNW and NW oriented faults. The E to WNW oriented faults are small and build up the Gebel Qattamiya en echelon fault belt, whereas the faults trending NNW and NW establish a pervasive horst and graben structural style involving some rhomb-shape horsts as Gebel Qattamiya (GQRH), Gebel Um Reheiat (GURRH) and south Gebel Um Reheiat (SGURRH). Rock units of the Eocene succession and Oligocene sediments are well exposed and highly controlled by rift-related structures. Rifting was developed through two rift-phases; initial and major ones. The initial phase (a newly recognized phase in this contribution) has been occurred in Late Eocene (Priabonian), while the main phase was prevailing during Late Oligocene-Early Miocene time and is characterized by hydrothermal veins and basaltic eruptions. Continental transtension in the Cairo-Suez district, including the study area, was probably synchronous with a major tectonic stage (Pyrenean-Atlasic movement) of continental collision between African-Arabian and Eurasian plates in Late Eocene-Oligocene time. Field investigation suggests that the transfer of displacement (slip) from the Gulf of Suez proto-rift into the E-W oriented faults ''relays'' is an important mechanism, which helps to explain the current structural framework and tectonic evolution of the Cairo-Suez district. Reactivation of such faults with right-lateral divergent wrenching with NE-SW oriented extension deformed the Cairo-Suez district with several E-W oriented en echelon fault belts (e.g. Gebel Qattamiya fault belt in the study area). Thus the Cairo-Suez district represents an accommodation or transfer zone in northeastern Egypt, intercepting the ''far-field stresses'' from the Arabian-Nubian Shield, the Red

  19. Effects of tsaoko (Fructus tsaoko) cultivating on tree diversity and canopy structure in the habitats of eastern hoolock gibbon (Hoolock leuconedys).

    PubMed

    Yuan, Sheng-Dong; Fei, Han-Lan; Zhu, Shao-Han; Cui, Liang-Wei; Ai, Huai-Sen; Fan, Peng-Fei

    2014-05-01

    In this study, the quadrat method was used to study the effects of tsaoko (Fructus tsaoko) plantation on tree diversity and canopy structure of two natural habitats of eastern hoolock gibbon (Hoolock leuconedys): Nankang (characterized by extensive tsaoko plantation) and Banchang (relatively well reserved and without tsaoko plantation). Totally, 102 tree species from 25 families and 16 woody liana species from 10 families were recorded in Nankang, whereas 108 tree species from 30 families and 17 woody liana species from 12 families were recorded in Banchang. Although the tree species between two habitats is different, both habitats are characterized by enriched food resources for eastern hoolock gibbons, sharing similar dominant plant families. Due to tsaoko plantation, tree density proportion and diversity of forest layerⅠ (>20 m) in Nankang were both significantly decreased, but the tree density of layerⅡ (10-20 m) increased. Likewise, in conjunction with these behavioral observations, we also address potential impacts of tsaoko plantation on the behavior of eastern hoolock gibbon. PMID:24866494

  20. Population structure and gene flow of the Atlantic walrus (Odobenus rosmarus rosmarus) in the eastern Atlantic Arctic based on mitochondrial DNA and microsatellite variation.

    PubMed

    Andersen, L W; Born, E W; Gjertz, I; Wiig, O; Holm, L E; Bendixen, C

    1998-10-01

    The population structure of the Atlantic walrus, Odobenus rosmarus rosmarus, was studied using 11 polymorphic microsatellites and restriction fragment length polymorphism detected in the NADH-dehydrogenase ND1, ND2 and ND3/4 segments in mtDNA. A total of 105 walrus samples were analysed from northwest (NW) Greenland, east (E) Greenland, Svalbard and Franz Joseph Land. Two of the 10 haplotypes detected in the four samples were diagnostic for the NW Greenland sample, which implied that the group of walruses in this area is evolutionary distinct from walruses in the other three areas. One individual sampled in E Greenland exhibited a Pacific haplotype, which proved a connection between the Pacific walrus and walruses in eastern Greenland. The Franz Joseph Land, Svalbard and E Greenland samples shared the most common haplotype, indicating very little differentiation at the mtDNA level. Gene flow (Nm) estimates among the four areas indicated a very restricted exchange of female genes between NW Greenland and the more eastern Atlantic Arctic samples, and a closer relationship between the three samples composing the eastern Atlantic Arctic. The genetic variation at 11 polymorphic microsatellite loci grouped individuals into three populations, NW Greenland, E Greenland and a common Franz Joseph Land-Svalbard population, which were connected by moderate gene flow.

  1. Population structure and gene flow of the Atlantic walrus (Odobenus rosmarus rosmarus) in the eastern Atlantic Arctic based on mitochondrial DNA and microsatellite variation.

    PubMed

    Andersen, L W; Born, E W; Gjertz, I; Wiig, O; Holm, L E; Bendixen, C

    1998-10-01

    The population structure of the Atlantic walrus, Odobenus rosmarus rosmarus, was studied using 11 polymorphic microsatellites and restriction fragment length polymorphism detected in the NADH-dehydrogenase ND1, ND2 and ND3/4 segments in mtDNA. A total of 105 walrus samples were analysed from northwest (NW) Greenland, east (E) Greenland, Svalbard and Franz Joseph Land. Two of the 10 haplotypes detected in the four samples were diagnostic for the NW Greenland sample, which implied that the group of walruses in this area is evolutionary distinct from walruses in the other three areas. One individual sampled in E Greenland exhibited a Pacific haplotype, which proved a connection between the Pacific walrus and walruses in eastern Greenland. The Franz Joseph Land, Svalbard and E Greenland samples shared the most common haplotype, indicating very little differentiation at the mtDNA level. Gene flow (Nm) estimates among the four areas indicated a very restricted exchange of female genes between NW Greenland and the more eastern Atlantic Arctic samples, and a closer relationship between the three samples composing the eastern Atlantic Arctic. The genetic variation at 11 polymorphic microsatellite loci grouped individuals into three populations, NW Greenland, E Greenland and a common Franz Joseph Land-Svalbard population, which were connected by moderate gene flow. PMID:9787444

  2. Effects of tsaoko (Fructus tsaoko) cultivating on tree diversity and canopy structure in the habitats of eastern hoolock gibbon (Hoolock leuconedys)

    PubMed Central

    Sheng-Dong, YUAN; Han-Lan, FEI; Shao-Han, ZHU; Liang-Wei, CUI; Huai-Sen, AI; Peng-Fei, FAN

    2014-01-01

    In this study, the quadrat method was used to study the effects of tsaoko (Fructus tsaoko) plantation on tree diversity and canopy structure of two natural habitats of eastern hoolock gibbon (Hoolock leuconedys): Nankang (characterized by extensive tsaoko plantation) and Banchang (relatively well reserved and without tsaoko plantation). Totally, 102 tree species from 25 families and 16 woody liana species from 10 families were recorded in Nankang, whereas 108 tree species from 30 families and 17 woody liana species from 12 families were recorded in Banchang. Although the tree species between two habitats is different, both habitats are characterized by enriched food resources for eastern hoolock gibbons, sharing similar dominant plant families. Due to tsaoko plantation, tree density proportion and diversity of forest layerⅠ (>20 m) in Nankang were both significantly decreased, but the tree density of layerⅡ (10−20 m) increased. Likewise, in conjunction with these behavioral observations, we also address potential impacts of tsaoko plantation on the behavior of eastern hoolock gibbon. PMID:24866494

  3. Effects of tsaoko (Fructus tsaoko) cultivating on tree diversity and canopy structure in the habitats of eastern hoolock gibbon (Hoolock leuconedys).

    PubMed

    Yuan, Sheng-Dong; Fei, Han-Lan; Zhu, Shao-Han; Cui, Liang-Wei; Ai, Huai-Sen; Fan, Peng-Fei

    2014-05-01

    In this study, the quadrat method was used to study the effects of tsaoko (Fructus tsaoko) plantation on tree diversity and canopy structure of two natural habitats of eastern hoolock gibbon (Hoolock leuconedys): Nankang (characterized by extensive tsaoko plantation) and Banchang (relatively well reserved and without tsaoko plantation). Totally, 102 tree species from 25 families and 16 woody liana species from 10 families were recorded in Nankang, whereas 108 tree species from 30 families and 17 woody liana species from 12 families were recorded in Banchang. Although the tree species between two habitats is different, both habitats are characterized by enriched food resources for eastern hoolock gibbons, sharing similar dominant plant families. Due to tsaoko plantation, tree density proportion and diversity of forest layerⅠ (>20 m) in Nankang were both significantly decreased, but the tree density of layerⅡ (10-20 m) increased. Likewise, in conjunction with these behavioral observations, we also address potential impacts of tsaoko plantation on the behavior of eastern hoolock gibbon.

  4. Disturbance and climatic effects on red spruce community dynamics at its southern continuous range margin

    PubMed Central

    2014-01-01

    Red spruce (Picea rubens) populations experienced a synchronous rangewide decline in growth and vigor starting in the 1960s, likely caused by climate change and a combination of environmental disturbances. However, it is not yet known if populations continue to decline or have recovered. Red spruce growing near its southern range margin in Massachusetts is a species of concern, in light of the vulnerability to climate change. This study uses population data from 17 permanent plots coupled with tree-ring data to examine radial growth rates, determine the growth-climate relationship, and document disturbance events. Red spruce at these plots ranged from 90 to 184 years old, and comprised 15 to 29 m2/ha basal area. Red spruce seedlings and saplings were common at plots with previously high overstory spruce abundance, indicating it could return to a more dominant position under favorable growing conditions. However, permanent plot measures over a 50 year time span did not indicate any consistent trends for changes in basal area or density for red spruce or other woody species. Climate data show that mean annual minimum, maximum, and summer temperatures have increased over the last 100 years. Dendroclimatological analyses indicated that red spruce growth was sensitive to both temperature and precipitation. Prior to the 1960s, spruce at these sites showed a positive response to precipitation; however after a multi-year drought in the 1960s showed an increasingly negative correlation with precipitation. There has been a negative growth response to regional warming, as spruce radial growth was mostly constrained by increasing temperatures, potentially coupled with the associated increasing drought-dress. I suggest the change in climate response is potentially due to a physiological threshold response to increasing temperatures, which may cause spruce to continue to decline or be lost from the lower elevation sites, while the high elevation sites has a persistent spruce

  5. Extensive Families of miRNAs and PHAS Loci in Norway Spruce Demonstrate the Origins of Complex phasiRNA Networks in Seed Plants.

    PubMed

    Xia, Rui; Xu, Jing; Arikit, Siwaret; Meyers, Blake C

    2015-11-01

    In eudicot plants, the miR482/miR2118 superfamily regulates and instigates the production of phased secondary small interfering RNAs (siRNAs) from NB-LRR (nucleotide binding leucine-rich repeat) genes that encode disease resistance proteins. In grasses, this miRNA family triggers siRNA production specifically in reproductive tissues from long noncoding RNAs. To understand this functional divergence, we examined the small RNA population in the ancient gymnosperm Norway spruce (Picea abies). As many as 41 miRNA families in spruce were found to trigger phasiRNA (phased, secondary siRNAs) production from diverse PHAS loci, with a remarkable 19 miRNA families capable of targeting over 750 NB-LRR genes to generate phasiRNAs. miR482/miR2118, encoded in spruce by at least 24 precursor loci, targets not only NB-LRR genes to trigger phasiRNA production (as in eudicots) but also noncoding PHAS loci, generating phasiRNAs preferentially in male or female cones, reminiscent of its role in the grasses. These data suggest a dual function of miR482/miR2118 present in gymnosperms that was selectively yet divergently retained in flowering plants. A few MIR482/MIR2118 precursors possess an extremely long stem-loop structure, one arm of which shows significant sequence similarity to spruce NB-LRR genes, suggestive of an evolutionary origin from NB-LRR genes through gene duplication. We also characterized an expanded miR390-TAS3 (TRANS-ACTING SIRNA GENE 3)-ARF (AUXIN RESPONSIVE FACTOR) pathway, comprising 18 TAS3 genes of diverse features. Finally, we annotated spruce miRNAs and their targets. Taken together, these data expand our understanding of phasiRNA network in plants and the evolution of plant miRNAs, particularly miR482/miR2118 and its functional diversification.

  6. Extensive Families of miRNAs and PHAS Loci in Norway Spruce Demonstrate the Origins of Complex phasiRNA Networks in Seed Plants

    PubMed Central

    Xia, Rui; Xu, Jing; Arikit, Siwaret; Meyers, Blake C.

    2015-01-01

    In eudicot plants, the miR482/miR2118 superfamily regulates and instigates the production of phased secondary small interfering RNAs (siRNAs) from NB-LRR (nucleotide binding leucine-rich repeat) genes that encode disease resistance proteins. In grasses, this miRNA family triggers siRNA production specifically in reproductive tissues from long noncoding RNAs. To understand this functional divergence, we examined the small RNA population in the ancient gymnosperm Norway spruce (Picea abies). As many as 41 miRNA families in spruce were found to trigger phasiRNA (phased, secondary siRNAs) production from diverse PHAS loci, with a remarkable 19 miRNA families capable of targeting over 750 NB-LRR genes to generate phasiRNAs. miR482/miR2118, encoded in spruce by at least 24 precursor loci, targets not only NB-LRR genes to trigger phasiRNA production (as in eudicots) but also noncoding PHAS loci, generating phasiRNAs preferentially in male or female cones, reminiscent of its role in the grasses. These data suggest a dual function of miR482/miR2118 present in gymnosperms that was selectively yet divergently retained in flowering plants. A few MIR482/MIR2118 precursors possess an extremely long stem-loop structure, one arm of which shows significant sequence similarity to spruce NB-LRR genes, suggestive of an evolutionary origin from NB-LRR genes through gene duplication. We also characterized an expanded miR390-TAS3 (TRANS-ACTING SIRNA GENE 3)-ARF (AUXIN RESPONSIVE FACTOR) pathway, comprising 18 TAS3 genes of diverse features. Finally, we annotated spruce miRNAs and their targets. Taken together, these data expand our understanding of phasiRNA network in plants and the evolution of plant miRNAs, particularly miR482/miR2118 and its functional diversification. PMID:26318183

  7. Micro- and macro-geographic scale effect on the molecular imprint of selection and adaptation in Norway spruce.

    PubMed

    Scalfi, Marta; Mosca, Elena; Di Pierro, Erica Adele; Troggio, Michela; Vendramin, Giovanni Giuseppe; Sperisen, Christoph; La Porta, Nicola; Neale, David B

    2014-01-01

    Forest tree species of temperate and boreal regions have undergone a long history of demographic changes and evolutionary adaptations. The main objective of this study was to detect signals of selection in Norway spruce (Picea abies [L.] Karst), at different sampling-scales and to investigate, accounting for population structure, the effect of environment on species genetic diversity. A total of 384 single nucleotide polymorphisms (SNPs) representing 290 genes were genotyped at two geographic scales: across 12 populations distributed along two altitudinal-transects in the Alps (micro-geographic scale), and across 27 populations belonging to the range of Norway spruce in central and south-east Europe (macro-geographic scale). At the macrogeographic scale, principal component analysis combined with Bayesian clustering revealed three major clusters, corresponding to the main areas of southern spruce occurrence, i.e. the Alps, Carpathians, and Hercynia. The populations along the altitudinal transects were not differentiated. To assess the role of selection in structuring genetic variation, we applied a Bayesian and coalescent-based F(ST)-outlier method and tested for correlations between allele frequencies and climatic variables using regression analyses. At the macro-geographic scale, the F(ST)-outlier methods detected together 11 F(ST)-outliers. Six outliers were detected when the same analyses were carried out taking into account the genetic structure. Regression analyses with population structure correction resulted in the identification of two (micro-geographic scale) and 38 SNPs (macro-geographic scale) significantly correlated with temperature and/or precipitation. Six of these loci overlapped with F(ST)-outliers, among them two loci encoding an enzyme involved in riboflavin biosynthesis and a sucrose synthase. The results of this study indicate a strong relationship between genetic and environmental variation at both geographic scales. It also suggests that an

  8. Drought-induced stomatal closure probably cannot explain divergent white spruce growth in the Brooks Range, Alaska, USA.

    PubMed

    Brownlee, Annalis H; Sullivan, Patrick F; Csank, Adam Z; Sveinbjörnsson, Bjartmar; Ellison, Sarah B Z

    2016-01-01

    Increment cores from the boreal forest have long been used to reconstruct past climates. However, in recent years, numerous studies have revealed a deterioration of the correlation between temperature and tree growth that is commonly referred to as divergence. In the Brooks Range of northern Alaska, USA, studies of white spruce (Picea glauca) revealed that trees in the west generally showed positive growth trends, while trees in the central and eastern Brooks Range showed mixed and negative trends during late 20th century warming. The growing season climate of the eastern Brooks Range is thought to be drier than the west. On this basis, divergent tree growth in the eastern Brooks Range has been attributed to drought stress. To investigate the hypothesis that drought-induced stomatal closure can explain divergence in the Brooks Range, we synthesized all of the Brooks Range white spruce data available in the International Tree Ring Data Bank (ITRDB) and collected increment cores from our primary sites in each of four watersheds along a west-to-east gradient near the Arctic treeline. For cores from our sites, we measured ring widths and calculated carbon isotope discrimination (δ13C), intrinsic water-use efficiency (iWUE), and needle intercellular CO2 concentration (C(i)) from δ13C in tree-ring alpha-cellulose. We hypothesized that trees exhibiting divergence would show a corresponding decline in δ13C, a decline in C(i), and a strong increase in iWUE. Consistent with the ITRDB data, trees at our western and central sites generally showed an increase in the strength of the temperature-growth correlation during late 20th century warming, while trees at our eastern site showed strong divergence. Divergent tree growth was not, however, associated with declining δ13C. Meanwhile, estimates of C(i) showed a strong increase at all of our study sites, indicating that more substrate was available for photosynthesis in the early 21st than in the early 20th century. Our

  9. Drought-induced stomatal closure probably cannot explain divergent white spruce growth in the Brooks Range, Alaska, USA.

    PubMed

    Brownlee, Annalis H; Sullivan, Patrick F; Csank, Adam Z; Sveinbjörnsson, Bjartmar; Ellison, Sarah B Z

    2016-01-01

    Increment cores from the boreal forest have long been used to reconstruct past climates. However, in recent years, numerous studies have revealed a deterioration of the correlation between temperature and tree growth that is commonly referred to as divergence. In the Brooks Range of northern Alaska, USA, studies of white spruce (Picea glauca) revealed that trees in the west generally showed positive growth trends, while trees in the central and eastern Brooks Range showed mixed and negative trends during late 20th century warming. The growing season climate of the eastern Brooks Range is thought to be drier than the west. On this basis, divergent tree growth in the eastern Brooks Range has been attributed to drought stress. To investigate the hypothesis that drought-induced stomatal closure can explain divergence in the Brooks Range, we synthesized all of the Brooks Range white spruce data available in the International Tree Ring Data Bank (ITRDB) and collected increment cores from our primary sites in each of four watersheds along a west-to-east gradient near the Arctic treeline. For cores from our sites, we measured ring widths and calculated carbon isotope discrimination (δ13C), intrinsic water-use efficiency (iWUE), and needle intercellular CO2 concentration (C(i)) from δ13C in tree-ring alpha-cellulose. We hypothesized that trees exhibiting divergence would show a corresponding decline in δ13C, a decline in C(i), and a strong increase in iWUE. Consistent with the ITRDB data, trees at our western and central sites generally showed an increase in the strength of the temperature-growth correlation during late 20th century warming, while trees at our eastern site showed strong divergence. Divergent tree growth was not, however, associated with declining δ13C. Meanwhile, estimates of C(i) showed a strong increase at all of our study sites, indicating that more substrate was available for photosynthesis in the early 21st than in the early 20th century. Our

  10. Hartig' net formation of Tricholoma vaccinum-spruce ectomycorrhiza in hydroponic cultures.

    PubMed

    Henke, Catarina; Jung, Elke-Martina; Kothe, Erika

    2015-12-01

    For re-forestation of metal-contaminated land, ectomycorrhizal trees may provide a solution. Hence, the study of the interaction is necessary to allow for comprehensive understanding of the mutually symbiotic features. On a structural level, hyphal mantle and the Hartig' net formed in the root apoplast are essential for plant protection and mycorrhizal functioning. As a model, we used the basidiomycete Tricholoma vaccinum and its host spruce (Picea abies). Using an optimized hydroponic cultivation system, both features could be visualized and lower stress response of the tree was obtained in non-challenged cultivation. Larger spaces in the apoplasts could be shown with high statistical significance. The easy accessibility will allow to address metal stress or molecular responses in both partners. Additionally, the proposed cultivation system will enable for other experimental applications like addressing flooding, biological interactions with helper bacteria, chemical signaling, or other biotic or abiotic challenges relevant in the natural habitat. PMID:25791268

  11. Hartig' net formation of Tricholoma vaccinum-spruce ectomycorrhiza in hydroponic cultures.

    PubMed

    Henke, Catarina; Jung, Elke-Martina; Kothe, Erika

    2015-12-01

    For re-forestation of metal-contaminated land, ectomycorrhizal trees may provide a solution. Hence, the study of the interaction is necessary to allow for comprehensive understanding of the mutually symbiotic features. On a structural level, hyphal mantle and the Hartig' net formed in the root apoplast are essential for plant protection and mycorrhizal functioning. As a model, we used the basidiomycete Tricholoma vaccinum and its host spruce (Picea abies). Using an optimized hydroponic cultivation system, both features could be visualized and lower stress response of the tree was obtained in non-challenged cultivation. Larger spaces in the apoplasts could be shown with high statistical significance. The easy accessibility will allow to address metal stress or molecular responses in both partners. Additionally, the proposed cultivation system will enable for other experimental applications like addressing flooding, biological interactions with helper bacteria, chemical signaling, or other biotic or abiotic challenges relevant in the natural habitat.

  12. In vitro fungistatic effects of natural coniferous resin from Norway spruce (Picea abies).

    PubMed

    Rautio, M; Sipponen, A; Lohi, J; Lounatmaa, K; Koukila-Kähkölä, P; Laitinen, K

    2012-08-01

    Resins (rosin, pitch) are natural products of the coniferous trees and are antimicrobial against a wide range of microbes. The antifungal effectiveness of resin, purified from Norway spruce (Picea abies), was studied against human pathogenic fungi and yeasts with the agar plate diffusion tests and electron microscopy (EM). The fungistatic effect of these resin mixtures (resin salves) was tested against a set of Candida yeasts, dermatophytes, and opportunistic fungi. Transmission and scanning EM was done from samples of fungi (Trichophyton mentagrophytes). In agar diffusion tests, the resin was strongly antifungal against all dermatophytes tested, e.g., against all fungi of the genus Trichophyton, but it was not antifungal against the Candida yeasts or against the opportunistic fungi tested. According to EM, resin caused damages in the cell hyphae and cell wall structures. We conclude that, in the agar plate diffusion test, coniferous resins are strongly fungistatic against the dermatophytic fungi only.

  13. Stratigraphy, structure and regional correlation of eastern Blue Ridge sequences in southern Virginia and northwestern North Carolina: an interim report from new USGS mapping

    USGS Publications Warehouse

    Carter, Mark W.; Merschat, Arthur J.

    2014-01-01

    The contact between eastern Blue Ridge stratified rocks above Mesoproterozoic basement rocks is mostly faulted (Gossan Lead and Red Valley). The Callaway fault juxtaposes Ashe and Lynchburg rocks above Wills Ridge Formation. Alligator Back Formation rocks overlie Ashe and Lynchburg rocks along the Rock Castle Creek fault, which juxtaposes rocks of different metamorphism. The fault separates major structural domains: rocks with one penetrative foliation in the footwall, and pin-striped recrystallized compositional layering, superposed penetrative foliations, and cleavage characterize the hanging wall. These relationships are ambiguous along strike to the southwest, where the Ashe and Alligator Back formations are recrystallized at higher metamorphic grades.

  14. Accumulation of organic air constituents by plant surfaces. Spruce needles for monitoring airborne chlorinated hydrocarbons

    SciTech Connect

    Reischl, A.; Thoma, H.; Reissinger, M.; Hutzinger, O. )

    1988-10-01

    The needles of the spruce (Picea abies) were used to monitor ambient air for organic trace substances. Analyses of spruce needles in an industrialized area demonstrated that the concentrations of these substances were much higher than those in a nonindustrialized area.

  15. Foliage responses of spruce trees to long-term low-grade sulfur dioxide deposition.

    PubMed

    Meng, F R; Bourque, C P; Belczewski, R F; Whitney, N J; Arp, P A

    1995-01-01

    Foliage on spruce trees (Picea rubens Sarg.) growing on dry SO(2) deposition zones (dry SO(2) deposition ranging from 0.5 and 8.5 S kg ha(-1) year(-1)) downwind from a SO(2) emission source was analyzed to assess chronic effects of long-term low-grade SO(2) deposition on net photosynthesis, stomatal conductance, dark respiration, stomatal antechamber wax structures, elemental concentrations in and on foliage (bulk and surficial concentrations), and types of epiphytic fungi that reside in the phylloplane. Elemental distributions on stomatal antechambers, on fungal colonies, and on smooth surfaces between stomates and fungus colonies were determined with a scanning electronic microscope (SEM) by way of X-ray scanning. It was found that net photosynthesis of newly developed spruce foliage (current-year, and 1-year-old) was not significantly affected by the local SO(2) deposition rates. Sulfur dioxide deposition, however, may have contributed to the gradual decrease in net photosynthesis with increasing needle age. Dark respiration rates were significantly higher on foliage taken from high SO(2) deposition zones. Stomatal rod-web structures deteriorated to flakes with increasing needle age and increasing SO(2) deposition. Further inspection of the needle surfaces revealed an increasing abundance of fungal colonies with increasing needle age. Many fungal taxa were isolated and identified. It was found that black yeasts responded positively, and Xylohypha pinicola responded negatively to high rates of SO(2) deposition. Surficial concentrations of elements such as P, S, K, Cl, Ca were about 10 times higher on fungal colonies than on smooth needle surfaces. Surficial Ca contents on 4 or 5-year-old needles decreased with increasing SO(2) deposition, but surficial S concentrations remained the same. In contrast, bulk foliar Ca and S concentrations increased with increasing SO(2) deposition.

  16. Foliage responses of spruce trees to long-term low-grade sulfur dioxide deposition.

    PubMed

    Meng, F R; Bourque, C P; Belczewski, R F; Whitney, N J; Arp, P A

    1995-01-01

    Foliage on spruce trees (Picea rubens Sarg.) growing on dry SO(2) deposition zones (dry SO(2) deposition ranging from 0.5 and 8.5 S kg ha(-1) year(-1)) downwind from a SO(2) emission source was analyzed to assess chronic effects of long-term low-grade SO(2) deposition on net photosynthesis, stomatal conductance, dark respiration, stomatal antechamber wax structures, elemental concentrations in and on foliage (bulk and surficial concentrations), and types of epiphytic fungi that reside in the phylloplane. Elemental distributions on stomatal antechambers, on fungal colonies, and on smooth surfaces between stomates and fungus colonies were determined with a scanning electronic microscope (SEM) by way of X-ray scanning. It was found that net photosynthesis of newly developed spruce foliage (current-year, and 1-year-old) was not significantly affected by the local SO(2) deposition rates. Sulfur dioxide deposition, however, may have contributed to the gradual decrease in net photosynthesis with increasing needle age. Dark respiration rates were significantly higher on foliage taken from high SO(2) deposition zones. Stomatal rod-web structures deteriorated to flakes with increasing needle age and increasing SO(2) deposition. Further inspection of the needle surfaces revealed an increasing abundance of fungal colonies with increasing needle age. Many fungal taxa were isolated and identified. It was found that black yeasts responded positively, and Xylohypha pinicola responded negatively to high rates of SO(2) deposition. Surficial concentrations of elements such as P, S, K, Cl, Ca were about 10 times higher on fungal colonies than on smooth needle surfaces. Surficial Ca contents on 4 or 5-year-old needles decreased with increasing SO(2) deposition, but surficial S concentrations remained the same. In contrast, bulk foliar Ca and S concentrations increased with increasing SO(2) deposition. PMID:15091479

  17. Water masses, ocean fronts, and the structure of Antarctic seabird communities: putting the eastern Bellingshausen Sea in perspective

    USGS Publications Warehouse

    Ribic, Christine A.; Ainley, David G.; Ford, R. Glenn; Fraser, William R.; Tynan, Cynthia T.; Woehler, Eric J.

    2015-01-01

    Waters off the western Antarctic Peninsula (i.e., the eastern Bellingshausen Sea) are unusually complex owing to the convergence of several major fronts. Determining the relative influence of fronts on occurrence patterns of top-trophic species in that area, therefore, has been challenging. In one of the few ocean-wide seabird data syntheses, in this case for the Southern Ocean, we analyzed ample, previously collected cruise data, Antarctic-wide, to determine seabird species assemblages and quantitative relationships to fronts as a way to provide context to the long-term Palmer LTER and the winter Southern Ocean GLOBEC studies in the eastern Bellingshausen Sea. Fronts investigated during both winter (April–September) and summer (October–March) were the southern boundary of the Antarctic Circumpolar Current (ACC), which separates the High Antarctic from the Low Antarctic water mass, and within which are embedded the marginal ice zone and Antarctic Shelf Break Front; and the Antarctic Polar Front, which separates the Low Antarctic and the Subantarctic water masses. We used clustering to determine species' groupings with water masses, and generalized additive models to relate species' densities, biomass and diversity to distance to respective fronts. Antarctic-wide, in both periods, highest seabird densities and lowest species diversity were found in the High Antarctic water mass. In the eastern Bellingshausen, seabird density in the High Antarctic water mass was lower (as low as half that of winter) than found in other Antarctic regions. During winter, Antarctic-wide, two significant species groups were evident: one dominated by Adélie penguins (Pygoscelis adeliae) (High Antarctic water mass) and the other by petrels and prions (no differentiation among water masses); in eastern Bellingshausen waters during winter, the one significant species group was composed of species from both Antarctic-wide groups. In summer, Antarctic-wide, a High Antarctic group

  18. Water masses, ocean fronts, and the structure of Antarctic seabird communities: Putting the eastern Bellingshausen Sea in perspective

    NASA Astrophysics Data System (ADS)

    Ribic, Christine A.; Ainley, David G.; Glenn Ford, R.; Fraser, William R.; Tynan, Cynthia T.; Woehler, Eric J.

    2011-07-01

    Waters off the western Antarctic Peninsula (i.e., the eastern Bellingshausen Sea) are unusually complex owing to the convergence of several major fronts. Determining the relative influence of fronts on occurrence patterns of top-trophic species in that area, therefore, has been challenging. In one of the few ocean-wide seabird data syntheses, in this case for the Southern Ocean, we analyzed ample, previously collected cruise data, Antarctic-wide, to determine seabird species assemblages and quantitative relationships to fronts as a way to provide context to the long-term Palmer LTER and the winter Southern Ocean GLOBEC studies in the eastern Bellingshausen Sea. Fronts investigated during both winter (April-September) and summer (October-March) were the southern boundary of the Antarctic Circumpolar Current (ACC), which separates the High Antarctic from the Low Antarctic water mass, and within which are embedded the marginal ice zone and Antarctic Shelf Break Front; and the Antarctic Polar Front, which separates the Low Antarctic and the Subantarctic water masses. We used clustering to determine species' groupings with water masses, and generalized additive models to relate species' densities, biomass and diversity to distance to respective fronts. Antarctic-wide, in both periods, highest seabird densities and lowest species diversity were found in the High Antarctic water mass. In the eastern Bellingshausen, seabird density in the High Antarctic water mass was lower (as low as half that of winter) than found in other Antarctic regions. During winter, Antarctic-wide, two significant species groups were evident: one dominated by Adélie penguins ( Pygoscelis adeliae) (High Antarctic water mass) and the other by petrels and prions (no differentiation among water masses); in eastern Bellingshausen waters during winter, the one significant species group was composed of species from both Antarctic-wide groups. In summer, Antarctic-wide, a High Antarctic group dominated

  19. Water masses, ocean fronts, and the structure of Antarctic seabird communities: putting the eastern Bellingshausen Sea in perspective

    USGS Publications Warehouse

    Ribic, Christine A.; Ainley, David G.; Ford, R. Glenn; Fraser, William R.; Tynan, Cynthia T.; Woehler, Eric J.

    2011-01-01

    Waters off the western Antarctic Peninsula (i.e., the eastern Bellingshausen Sea) are unusually complex owing to the convergence of several major fronts. Determining the relative influence of fronts on occurrence patterns of top-trophic species in that area, therefore, has been challenging. In one of the few ocean-wide seabird data syntheses, in this case for the Southern Ocean, we analyzed ample, previously collected cruise data, Antarctic-wide, to determine seabird species assemblages and quantitative relationships to fronts as a way to provide context to the long-term Palmer LTER and the winter Southern Ocean GLOBEC studies in the eastern Bellingshausen Sea. Fronts investigated during both winter (April–September) and summer (October–March) were the southern boundary of the Antarctic Circumpolar Current (ACC), which separates the High Antarctic from the Low Antarctic water mass, and within which are embedded the marginal ice zone and Antarctic Shelf Break Front; and the Antarctic Polar Front, which separates the Low Antarctic and the Subantarctic water masses. We used clustering to determine species' groupings with water masses, and generalized additive models to relate species' densities, biomass and diversity to distance to respective fronts. Antarctic-wide, in both periods, highest seabird densities and lowest species diversity were found in the High Antarctic water mass. In the eastern Bellingshausen, seabird density in the High Antarctic water mass was lower (as low as half that of winter) than found in other Antarctic regions. During winter, Antarctic-wide, two significant species groups were evident: one dominated by Adélie penguins (Pygoscelis adeliae) (High Antarctic water mass) and the other by petrels and prions (no differentiation among water masses); in eastern Bellingshausen waters during winter, the one significant species group was composed of species from both Antarctic-wide groups. In summer, Antarctic-wide, a High Antarctic group

  20. New structural and seismological evidence and interpretation of a lithospheric-scale shear zone at the southern edge of the Ionian subduction system (central-eastern Sicily, Italy)

    NASA Astrophysics Data System (ADS)

    Barreca, G.; Scarfı, L.; Cannavò, F.; Koulakov, I.; Monaco, C.

    2016-06-01

    Geological, gravimetric, and seismological data from the central-eastern Sicily (Italy) provide evidences of a NW-SE oriented shear zone at the southern edge of the Ionian subduction system. This structure consists of a near 100 km long lithospheric-scale structural and seismic boundary. In the near-surface, it shows Plio-Pleistocene vertical-axis structural rotations, kilometer-scale topographic imprint, progressive wrenching, and large down-faulting. All these features, together with its location south-west of the subduction system, allow us to interpret the shear zone as the upper plate expression of an abandoned Subduction Transform Edge Propagator fault, working before slab detachment, currently reactivated by elastic rebound or mantle upwelling mechanism triggered by slab detachment, to form an incipient transform belt separating compartments characterized by different motion in the modern context of Africa-Europe convergence.

  1. Midwinter needle temperature and winter injury of montane red spruce.

    PubMed

    Strimbeck, G R; Johnson, A H; Vann, D R

    1993-09-01

    To assess the role of solar warming and associated temperature fluctuations in the winter injury of sun-exposed red spruce foliage, we used fine wire thermocouples to monitor midwinter needle temperature in the upper canopy of mature red spruce trees over two winters. In 1989-1990, 15-min mean temperatures were recorded for six needles in a single tree. In 1990-1991, 10-min mean temperatures of six needles in one tree, and 1-min mean temperatures of seven needles in a second tree were recorded during rapid temperature changes. Warming was more frequent and greatest on terminal shoots of branches with a south to southwest aspect. The maximum rise above ambient air temperature exceeded 20 degrees C, and the maximum one minute decrease in temperature was 9 degrees C, with maximum rates of 0.8 and 0.6 degrees C min(-1) sustained over 10- and 15-min intervals, respectively. These data demonstrate that red spruce is subject to rapid temperature fluctuations similar to those known to produce visible injury in American aborvitae, a much hardier species. We concluded that solar warming to temperatures above the freezing point was unlikely to result in dehardening and subsequent freezing injury, because warming was infrequent, of short duration, and did not always raise needle temperature above the freezing point. Parts of branches and some individual shoots were frequently covered by snow or rime that may have prevented injury by reducing the frequency or intensity of needle temperature fluctuations. Radiation load on exposed shoots may have been increased by reflection of short wave radiation from snow and rime deposits on surrounding surfaces, which would exacerbate temperature fluctuations. PMID:14969891

  2. Sperm whale population structure in the eastern and central North Pacific inferred by the use of single-nucleotide polymorphisms, microsatellites and mitochondrial DNA.

    PubMed

    Mesnick, Sarah L; Taylor, Barbara L; Archer, Frederick I; Martien, Karen K; Treviño, Sergio Escorza; Hancock-Hanser, Brittany L; Moreno Medina, Sandra Carolina; Pease, Victoria L; Robertson, Kelly M; Straley, Janice M; Baird, Robin W; Calambokidis, John; Schorr, Gregory S; Wade, Paul; Burkanov, Vladimir; Lunsford, Chris R; Rendell, Luke; Morin, Phillip A

    2011-03-01

    We use mitochondrial DNA (mtDNA) (400 bp), six microsatellites and 36 single-nucleotide polymorphisms (SNPs), 20 of which were linked, to investigate population structure of sperm whales (Physeter macrocephalus) in the eastern and central North Pacific. SNP markers, reproducible across technologies and laboratories, are ideal for long-term studies of globally distributed species such as sperm whales, a species of conservation concern because of both historical and contemporary impacts. We estimate genetic differentiation among three strata in the temperate to tropical waters where females are found: California Current, Hawai`i and the eastern tropical Pacific. We then consider how males on sub-Arctic foraging grounds assign to these strata. The California Current stratum was differentiated from both the other strata (P < 0.05) for mtDNA, microsatellites and SNPs, suggesting that the region supports a demographically independent population and providing the first indication that males may exhibit reproductive philopatry. Comparisons between the Hawai`i stratum and the eastern tropical Pacific stratum are not conclusive at this time. Comparisons with Alaska males were statistically significant, or nearly so, from all three strata and individuals showed mixed assignment to, and few exclusions from, the three potential source strata, suggesting widespread origin of males on sub-Arctic feeding grounds. We show that SNPs have sufficient power to detect population structure even when genetic differentiation is low. There is a need for better analytical methods for SNPs, especially when linked SNPs are used, but SNPs appear to be a valuable marker for long-term studies of globally dispersed and highly mobile species.

  3. Profile Measurements of BVOC Emissions from a Norway spruce

    NASA Astrophysics Data System (ADS)

    Wang, M.

    2015-12-01

    Biogenic volatile organic compounds (BVOCs) are known as a source of secondary organic aerosols (SOA) due to their high reactivity with O3 and OH in the atmosphere [1, 2]. Norway spruce is one of the dominant boreal forest species in northern Europe, which has been considered to be high monoterpene (MT) emitters [3, 4]. BVOC emissions and compound composition vary considerably under different temperature and light conditions through growing season [5, 6]. Information of vertical profile emission patterns is indispensible for understanding chemical processes and oxidation sinks within canopy and for modelling evaluation. We characterize the in-canopy BVOC emissions from a 120 years old Norway spruce in Central Sweden (Norunda, 60°05'N, 17°29'E). Air samples were taken during the growing season (June to September 2013 & 2014) from transparent dynamic branch chambers set up in a vertical profile with 4 levels (20 m, 15 m, 11 m and 3 m agl.) on the spruce. Samples were collected every hour from the chamber with Tenax-TA adsorbent tubes and a pocket pump, and analyzed later by gas chromatography and a mass selective detector (GC-MS) to quantify each trapped terpenoid compound. The emission spectrum of Norway spruce at 20 m canopy height was found to be more complex than the emissions spectra at lower canopy levels, and included isoprene, MT and SQT from June to September, while MT was the dominating terpenoid species. The emission spectra of July and August (isoprene, 14 MT and 3 SQT) were much more complex compared to June and September at the 20 m canopy level, and mainly caused by an increase of MT species during peak season. Similarly, isoprene showed a distinctive seasonal pattern, and was detected at all the heights during noon time except the bottom 3 m level during peak summer, but only at the highest layer (20 m) during noon in early or late summer. O3 vertical profile data will be available for further chemical process analysis within canopy. References[1]M

  4. Crust Uppermost Mantle Structure beneath Eastern Asia: Progress towards a Uniform, Tightly Constrained, High Resolution 3-D Model

    NASA Astrophysics Data System (ADS)

    Shen, W.; Ritzwoller, M. H.; Zheng, Y.; Lin, F. C.; Kim, Y.; Ning, J.; Kang, D.; Feng, L.; Wiens, D. A.

    2015-12-01

    In the past decade, large and dense seismic arrays have been deployed across much of eastern Asia (e.g., the "CEArray" and the "China Array" deployed by the China Earthquake Administration (CEA), the NECESS Array deployed collaboratively by China, Japan and the US, Korean Seismic Network, KNET and other networks in Japan, and historical PASSCAL installations), which have been used to produce increasingly well resolved models of the crust and uppermost mantle at different length scales. These models, however, do not cover eastern Asia uniformly. In this presentation, we report on an effort to generate a uniform high resolution 3-D model of the crust and uppermost mantle beneath eastern Asia using state-of-art surface wave and body wave inversion techniques. Highlights of this effort include: 1) We collect ambient noise cross-correlations using more than 1,800 seismic stations from multiple seismic arrays in this area and perform uniform surface wave tomography for the study area. 2) We collect P-wave receiver functions for over 1,000 stations and Rayleigh wave H/V ratio measurements for over 200 stations in this area. 3) We adopt a Bayesian Monte Carlo inversion to the Rayleigh wave dispersion maps and produce a uniform 3-D model with uncertainties of the crust and uppermost mantle. 4) In the areas where receiver functions and/or Rayleigh wave H/V ratios are collected, we replace the surface wave inversion by a joint inversion of surface waves and these seismic observables. The resulting model displays a great variety and considerable richness of geological and tectonic features in the crust and in the uppermost mantle which we summarize and discuss with focus on the relationship between the observed crustal variations and tectonic/geological boundaries and lithospheric modifications associated with volcanism in Northeast China.

  5. The history of Seasonally Dry Tropical Forests in eastern South America: inferences from the genetic structure of the tree Astronium urundeuva (Anacardiaceae).

    PubMed

    Caetano, S; Prado, D; Pennington, R T; Beck, S; Oliveira-Filho, A; Spichiger, R; Naciri, Y

    2008-07-01

    Today, the Seasonally Dry Tropical Forests (SDTF) of eastern South America occur as large, well-defined nuclei (e.g. Caatinga in the northeast) and as smaller enclaves within other vegetations (e.g. Cerrado and Chaco). In order to infer the way the present SDTF distribution was attained, the genetic structure of Astronium urundeuva, a tree confined to SDTF, was assessed using two chloroplast spacers and nine microsatellite loci. Five haplotypes were identified, whose distribution was spatially structured. The distribution of the two most common and divergent haplotypes suggested former vicariance and progressive divergence due to isolation. More recent range expansions of these two lineages subsequently occurred, leading to a secondary contact at the southern limit of the Caatinga SDTF nucleus. The multilocus-Bayesian approach using microsatellites consistently identified three groups of populations (Northeast, Central and Southwest). Isolation by distance was found in Northeast and Southwest groups whereas admixture was detected in the Central group, located at the transition between Caatinga and Cerrado domains. All together, the results support the existence of range expansions and secondary contact in the Central group. This study provides arguments that favour the existence of a previously more continuous formation of SDTF in eastern South America.

  6. Combined structural interventions for gender equality and livelihood security: a critical review of the evidence from southern and eastern Africa and the implications for young people

    PubMed Central

    Gibbs, Andrew; Willan, Samantha; Misselhorn, Alison; Mangoma, Jaqualine

    2012-01-01

    Background Young people in southern and eastern Africa remain disproportionately vulnerable to HIV with gender inequalities and livelihood insecurities being key drivers of this. Behavioural HIV prevention interventions have had weak outcomes and a new generation of structural interventions have emerged seeking to challenge the wider drivers of the HIV epidemic, including gender inequalities and livelihood insecurities. Methods We searched key academic data bases to identify interventions that simultaneously sought to strengthen people's livelihoods and transform gender relationships that had been evaluated in southern and eastern Africa. Our initial search identified 468 articles. We manually reviewed these and identified nine interventions that met our criteria for inclusion. Results We clustered the nine interventions into three groups: microfinance and gender empowerment interventions; supporting greater participation of women and girls in primary and secondary education; and gender empowerment and financial literacy interventions. We summarise the strengths and limitations of these interventions, with a particular focus on what lessons may be learnt for young people (18–24). Conclusions Our review identified three major lessons for structural interventions that sought to transform gender relationships and strengthen livelihoods: 1) interventions have a narrow conceptualisation of livelihoods, 2) there is limited involvement of men and boys in such interventions, 3) studies have typically been done in stable populations. We discuss what this means for future interventions that target young people through these methods. PMID:22713350

  7. Impact of warming and drought on carbon balance related to wood formation in black spruce

    PubMed Central

    Deslauriers, Annie; Beaulieu, Marilène; Balducci, Lorena; Giovannelli, Alessio; Gagnon, Michel J.; Rossi, Sergio

    2014-01-01

    Background and Aims Wood formation in trees represents a carbon sink that can be modified in the case of stress. The way carbon metabolism constrains growth during stress periods (high temperature and water deficit) is now under debate. In this study, the amounts of non-structural carbohydrates (NSCs) for xylogenesis in black spruce, Picea mariana, saplings were assessed under high temperature and drought in order to determine the role of sugar mobilization for osmotic purposes and its consequences for secondary growth. Methods Four-year-old saplings of black spruce in a greenhouse were subjected to different thermal conditions with respect to the outside air temperature (T0) in 2010 (2 and 5 °C higher than T0) and 2011 (6 °C warmer than T0 during the day or night) with a dry period of about 1 month in June of each year. Wood formation together with starch, NSCs and leaf parameters (water potential and photosynthesis) were monitored from May to September. Key Results With the exception of raffinose, the amounts of soluble sugars were not modified in the cambium even if gas exchange and photosynthesis were greatly reduced during drought. Raffinose increased more than pinitol under a pre-dawn water potential of less than –1 Mpa, presumably because this compound is better suited than polyol for replacing water and capturing free radicals, and its degradation into simple sugar is easier. Warming decreased the starch storage in the xylem as well the available hexose pool in the cambium and the xylem, probably because of an increase in respiration. Conclusions Radial stem growth was reduced during drought due to the mobilization of NSCs for osmotic purposes and due to the lack of cell turgor. Thus plant water status during wood formation can influence the NSCs available for growth in the cambium and xylem. PMID:24950772

  8. A high-density genetic linkage map of a black spruce (Picea mariana) × red spruce (Picea rubens) interspecific hybrid.

    PubMed

    Kang, Bum-Yong; Major, John E; Rajora, Om P

    2011-02-01

    Genetic maps provide an important genomic resource of basic and applied significance. Spruce (Picea) has a very large genome size (between 0.85 × 1010 and 2.4 × 1010 bp; 8.5-24.0 pg/1C, a mean of 17.7 pg/1C ). We have constructed a near-saturated genetic linkage map for an interspecific backcross (BC1) hybrid of black spruce (BS; Picea mariana (Mill.) B.S.P.) and red spruce (RS; Picea rubens Sarg.), using selectively amplified microsatellite polymorphic loci (SAMPL) markers. A total of 2284 SAMPL markers were resolved using 31 SAMPL-MseI selective nucleotide primer combinations. Of these, 1216 SAMPL markers showing Mendelian segregation were mapped, whereas 1068 (46.8%) SAMPL fragments showed segregation distortion at α = 0.05. Maternal, paternal, and consensus maps consistently coalesced into 12 linkage groups, corresponding to the haploid chromosome number (1n = 1x = 12) of 12 in the genus Picea. The maternal BS map consisted of 814 markers distributed over 12 linkage groups, covering 1670 cM, with a mean map distance of 2.1 cM between adjacent markers. The paternal BS × RS map consisted of 773 markers distributed over 12 linkage groups, covering 1563 cM, with a mean map distance of 2.0 cM between adjacent markers. The consensus interspecific hybrid BC1 map consisted of 1216 markers distributed over 12 linkage groups, covering 1865 cM (98% genome coverage), with a mean map distance of 1.5 cM between adjacent markers. The genetic map reported here provides an important genomic resource in Picea, Pinaceae, and conifers.

  9. Genetic structure of the date palm (Phoenix dactylifera) in the Old World reveals a strong differentiation between eastern and western populations

    PubMed Central

    Zehdi-Azouzi, Salwa; Cherif, Emira; Moussouni, Souhila; Gros-Balthazard, Muriel; Abbas Naqvi, Summar; Ludeña, Bertha; Castillo, Karina; Chabrillange, Nathalie; Bouguedoura, Nadia; Bennaceur, Malika; Si-Dehbi, Farida; Abdoulkader, Sabira; Daher, Abdourahman; Terral, Jean-Frederic; Santoni, Sylvain; Ballardini, Marco; Mercuri, Antonio; Ben Salah, Mohamed; Kadri, Karim; Othmani, Ahmed; Littardi, Claudio; Salhi-Hannachi, Amel; Pintaud, Jean-Christophe; Aberlenc-Bertossi, Frédérique

    2015-01-01

    Background and Aims Date palms (Phoenix dactylifera, Arecaceae) are of great economic and ecological value to the oasis agriculture of arid and semi-arid areas. However, despite the availability of a large date palm germplasm spreading from the Atlantic shores to Southern Asia, improvement of the species is being hampered by a lack of information on global genetic diversity and population structure. In order to contribute to the varietal improvement of date palms and to provide new insights on the influence of geographic origins and human activity on the genetic structure of the date palm, this study analysed the diversity of the species. Methods Genetic diversity levels and population genetic structure were investigated through the genotyping of a collection of 295 date palm accessions ranging from Mauritania to Pakistan using a set of 18 simple sequence repeat (SSR) markers and a plastid minisatellite. Key Results Using a Bayesian clustering approach, the date palm genotypes can be structured into two different gene pools: the first, termed the Eastern pool, consists of accessions from Asia and Djibouti, whilst the second, termed the Western pool, consists of accessions from Africa. These results confirm the existence of two ancient gene pools that have contributed to the current date palm diversity. The presence of admixed genotypes is also noted, which points at gene flows between eastern and western origins, mostly from east to west, following a human-mediated diffusion of the species. Conclusions This study assesses the distribution and level of genetic diversity of accessible date palm resources, provides new insights on the geographic origins and genetic history of the cultivated component of this species, and confirms the existence of at least two domestication origins. Furthermore, the strong genetic structure clearly established here is a prerequisite for any breeding programme exploiting the effective polymorphism related to each gene pool. PMID

  10. Fresh-stem bending of silver fir and Norway spruce.

    PubMed

    Lundström, Tor; Stoffel, Markus; Stöckli, Veronika

    2008-03-01

    The bending and growth characteristics of large fresh stems from four silver fir (Abies alba Mill.) and three Norway spruce (Picea abies (L.) Karst.) trees were studied. Twenty logs taken from different stem heights were subjected to four-point bending tests. From the bending test records, we calculated stress-strain curves, which accounted for detailed log taper, shear deformation and self weight. From these curves we determined, among other parameters, the modulus of elasticity (MOE), the modulus of rupture (MOR) and the work absorbed in bending (W). No significant differences were found between species for the wood properties examined. Values of MOE, MOR and W generally decreased with stem height, with MOR in the range of 43 to 59 MPa and MOE ranging from 10.6 to 15.6 GPa. These MOE values are twice or more those reported for stems of young Sitka spruce (Picea sitchensis (Bong.) Carr.) trees. Based on the radial growth properties measured in discs from the logs, we calculated predicted values of MOE and MOR for the stem cross section. The predictions of MOE were precise, whereas those of MOR were approximate because of a complex combination of different failure mechanisms. Methods to test and calculate MOE, MOR and W for the stems of living trees are discussed with the aim of improving analyses of tree biomechanics and assessments of forest stability protection.

  11. Functional profile of black spruce wetlands in Alaska

    SciTech Connect

    Post, R.A.

    1996-09-01

    The profile describes the ecologic context and wetland functions of black spruce (Picea mariana) wetlands (BSWs) covering about 14 million ha of Alaska taiga. Ecologic descriptions include climate, permafrost, landforms, post-Pleistocene vegetation, fire, successional processes, black spruce community types and adaptations, and characteristics of BSWs. The profile describes human activities potentially affecting BSWs and identifies research literature and data gaps generally applicable to BSWs. Hydrologic, water quality, global biogeochemical, and ecologic functions of BSWs, as well as their socioeconomic uses, appear in the profile, along with potential functional indicators, expected sensitivities of functions to fill placement or weltand drainage, and potential mitigation strategies for impacts. Functional analysis separately considers ombrotrophic and minerotrophic BSWs where appropriate. Depending on trophic status, Alaska`s BSWs perform several low-magnitude hydrologic (groundwater discharge and recharge, flow regulation, and erosion control) and ecologic (nutrient export, nutrient cycling, and food-chain support) functions and several substantial water quality (sediment retention, nutrient transformation, nutrient uptake, and contaminant removal), global biogeochemical (carbon cycling and storage), and ecologic (avian and mammalian habitat) functions. BSWs also provide important socioeconomic uses: harvested of wetland-dependent fish, wildlife, and plant resources and active winter recreation.

  12. Morphometric analysis of pollen grains for paleoecological studies: classification of Picea from eastern North America.

    PubMed

    Lindbladh, Matts; O'Connor, Raymond; Jacobson, George L

    2002-09-01

    Little is known about the paleoecological histories of the three spruce species (white spruce, Picea glauca; black spruce, P. mariana; and red spruce P. rubens) in eastern North America, largely because of the difficulty of separating the three species in the pollen record. We describe a novel and effective classification method of distinguishing pollen grains on the basis of quantitative analysis of grain attributes. The method is illustrated by an analysis of a large sample of modern pollen grains (522 grains from 38 collections) of the three Picea species, collected from the region where the three species co-occur today. For each species X we computed a binary regression tree that classified each grain either as X or as not-X; these three determinations for each grain were then combined as Hamming codes in an error/uncertainty detection procedure. The use of Hamming codes to link multiple binary trees for error detection allowed identification and exclusion of problematic specimens, with correspondingly greater classification certainty among the remaining grains. We measured 13 attributes of 419 reference grains of the three species to construct the regression trees and classified 103 other reference grains by testing. Species-specific accuracies among the reliably classified grains were 100, 77, and 76% for P. glauca, P. mariana, and P. rubens, respectively, and 21, 30, and 22% of the grains by species, respectively, were problematic. The method is applicable to any multi-species classification problem for which a large reference sample is available.

  13. Crustal structure and rift tectonics across the Cauvery-Palar basin, Eastern Continental Margin of India based on seismic and potential field modelling

    NASA Astrophysics Data System (ADS)

    Twinkle, D.; Rao, G. Srinivasa; Radhakrishna, M.; Murthy, K. S. R.

    2016-03-01

    The Cauvery-Palar basin is a major peri-cratonic rift basin located along the Eastern Continental Margin of India (ECMI) that had formed during the rift-drift events associated with the breakup of eastern Gondwanaland (mainly India-Sri Lanka-East Antarctica). In the present study, we carry out an integrated analysis of the potential field data across the basin to understand the crustal structure and the associated rift tectonics. The composite-magnetic anomaly map of the basin clearly shows the onshore-to-offshore structural continuity, and presence of several high-low trends related to either intrusive rocks or the faults. The Curie depth estimated from the spectral analysis of offshore magnetic anomaly data gave rise to 23 km in the offshore Cauvery-Palar basin. The 2D gravity and magnetic crustal models indicate several crustal blocks separated by major structures or faults, and the rift-related volcanic intrusive rocks that characterize the basin. The crustal models further reveal that the crust below southeast Indian shield margin is ˜36 km thick and thins down to as much as 13-16 km in the Ocean Continent Transition (OCT) region and increases to around 19-21 km towards deep oceanic areas of the basin. The faulted Moho geometry with maximum stretching in the Cauvery basin indicates shearing or low angle rifting at the time of breakup between India-Sri Lanka and the East Antarctica. However, the additional stretching observed in the Cauvery basin region could be ascribed to the subsequent rifting of Sri Lanka from India. The abnormal thinning of crust at the OCT is interpreted as the probable zone of emplaced Proto-Oceanic Crust (POC) rocks during the breakup. The derived crustal structure along with other geophysical data further reiterates sheared nature of the southern part of the ECMI.

  14. Importance of the margin inheritance in the present-day crustal structure of the Longmen Shan range, Eastern border of the Tibetan plateau

    NASA Astrophysics Data System (ADS)

    Robert, A.; Pubellier, M.; de Sigoyer, J.; Billerot, A.; Vergne, J.

    2011-12-01

    The Longmen Shan range, located at the eastern border of the Tibetan plateau (Sichuan province, China), is characterized by one of the most important topographic gradient of the world whereas the present-day convergence recorded by GPS measurements is quite low (~3mm/yr). The mountain is also located above an abrupt 18km-high Moho step interpreted as the result of the confrontation of the thick Tibetan crust (~63km) against the resistant Yangtze craton (~45km). This peculiar crustal structure, as well as the thickening of the Tibetan crust in its eastern part, is often considerate as a result of the Cenozoic collision between India and Eurasia. This study highlights the importance of structural and thermal inheritance in the current crustal structure of this area. In fact, sedimentary records indicate that the Longmen Shan formed an abrupt passive margin from the beginning of the Paleozoic to the middle Triassic as a consequence of the opening of the Prototethys and the Paleotethys. The Longmen Shan correspond to a major paleogeographic boundary between the shallow water sediments of the Sichuan basin to the South-East and the deep sea mostly flysch-type sediments of the Songpan Garze terrane to the North West. Moreover, geochimical data indicate an important extension of the continental crust further west of the sedimentary transition. The oblique position of the Longmen Shan compared to the direction of opening is responsible of a trantensive passive margin structure as observed for the present-day vietnamian passive margin. This geometry is characterized by an abrupt decrease of the continental crust thickness associated with the sedimentary transition whereas the continent/ocean transition is located further away. We propose that the peculiar geometry of the old passive margin in the Longmen Shan is responsible for the important localization of the deformation during the 3 reactivations of the range (Triassic, Cretaceous and Late Tertiary).

  15. Responses of black spruce (Picea mariana) and tamarack (Larix laricina) to flooding and ethylene.

    PubMed

    Islam, M Anisul; MacDonald, S Ellen; Zwiazek, Janusz J

    2003-06-01

    Black spruce (Picea mariana (Mill.) BSP) and tamarack (Larix laricina (Du Roi) K. Koch) are the predominant tree species in the boreal peatlands of Alberta, Canada, where low nutrient availability, low soil temperature and a high water table limit their growth. Effects of flooding for 28 days on morphological and physiological responses were investigated in greenhouse-grown black spruce and tamarack seedlings in a growth chamber. Flooding reduced root hydraulic conductance, net assimilation rate and stomatal conductance, and increased water-use efficiency (WUE) and needle electrolyte leakage in both species. Although flooded black spruce seedlings maintained higher net assimilation rates and stomatal conductance than flooded tamarack seedlings, flooded tamarack seedlings were able to maintain higher root hydraulic conductance than flooded black spruce seedlings. Needles of flooded black spruce developed tip necrosis and electrolyte leakage after 14 days of flooding, and these symptoms were subsequently more prominent than in needles of flooded tamarack seedlings. Flooded tamarack seedlings exhibited no visible injury symptoms and developed hypertrophied lenticels at their stem base. Application of exogenous ethylene resulted in a significant reduction in net assimilation, stomatal conductance and root respiration, whereas root hydraulic conductivity increased in both species. Thus, although flooded black spruce seedlings maintained a higher stomatal conductance and net assimilation rate than tamarack seedlings, black spruce did not cope with the deleterious effects of prolonged soil flooding and exogenous ethylene as well as tamarack. PMID:12730046

  16. Influence of eastern hemlock (Tsuga canadensis L.) on fish community structure and function in headwater streams of the Delaware River basin

    USGS Publications Warehouse

    Ross, R.M.; Bennett, R.M.; Snyder, C.D.; Young, J.A.; Smith, D.R.; Lemarie, D.P.

    2003-01-01

    Hemlock (Tsuga canadensis) forest of the eastern U.S. are in decline due to invasion by the exotic insect hemlock woolly adelgid (Adelges tsugae). Aquatic biodiversity in hemlock ecosystems has not been documented; thus the true impact of the infestation cannot be assessed. We compared ichthyofaunal assemblages and trophic structure of streams draining hemlock and hardwood forests by sampling first- and second-order streams draining 14 paired hemlock and hardwood stands during base flows in July 1997 at the Delaware Water Gap National Recreation Area of Pennsylvania and New Jersey. Over 1400 fish of 15 species and 7 families were collected, but hemlock and hardwood streams individually harbored only one to four species. Brook trout (Salvelinus fontinalis) and brown trout (Salmo trutta) were two to three times as prevalent in hemlock than hardwood streams. Insectivorous fishes occurred in significantly higher proportion in streams of hardwood (0.90) than hemlock (0.46) stands, while piscivores occurred more often in hemlock (0.85) than hardwood (0.54) stands. Functional (trophic) diversity of fishes in hemlock and second-order streams was numerically greater than that of hardwood and first-order streams. Species composition also differed by stream order and terrain type. Biodiversity is threatened at several levels within hemlock ecosystems at risk to the hemlock woolly adelgid in eastern U.S. forests.

  17. Reproductive potential of balsam fir (Abies balsamea), white spruce (Picea glauca), and black spruce (P. mariana) at the ecotone between mixedwood and coniferous forests in the boreal zone of western Quebec.

    PubMed

    Messaoud, Yassine; Bergeron, Yves; Asselin, Hugo

    2007-05-01

    The reproductive potentials of balsam fir and white spruce (co-dominants in mixedwood forests) and black spruce (dominant in coniferous forests) were studied to explain the location of the ecotone between the two forest types in the boreal zone of Quebec. Four sites were selected along a latitudinal gradient crossing the ecotone. Cone crop, number of seeds per cone, percentage filled seeds, and percentage germination were measured for each species. Balsam fir and white spruce cone crops were significantly lower in the coniferous than in the mixedwood forest, while black spruce had greater crop constancy and regularity between both forest types. Mast years were more frequent for black spruce than for balsam fir in both forest types (mast year data not available for white spruce). The number of seeds per cone was more related to cone size than to forest type for all species. Black spruce produced more filled seeds in the coniferous forest than balsam fir or white spruce. The sum of growing degree-days and the maximum temperature of the warmest month (both for the year prior to cone production) significantly affected balsam fir cone production. The climate-related northward decrease in reproductive potential of balsam fir and white spruce could partly explain the position of the northern limit of the mixedwood forest. This could change drastically, however, as the ongoing climate warming might cancel this competitive advantage of black spruce.

  18. Monoterpene emissions from Scots pine and Norwegian spruce

    SciTech Connect

    Janson, R.W. )

    1993-02-20

    Rates of monoterpene emissions from Scots pine (Pinus sylvestris) and Norwegian spruce (Picea abies) have been measured at four sites in Sweden with a dynamic flow chamber technique. Forest floor emissions have been made in the pine forest with the static chamber technique. The compounds [Delta][sup 3]-carene and [alpha]-pinene were the predominant terpenes emitted from the crown and floor of the Scots pine forest. Alpha-pinene was the main terpene emitted from Norwegian spruce at the sites in southern and central Sweden, while [Delta][sup 3]-carene was predominant at the northern site. Emission rates, normalized to temperature, were seen to vary diurnally with a maximum at midday, and seasonally with maxima in early May and October, and a summer maximum in June-July. The possible dependence of the emission rate on needle growth rate and other plant-physiological processes is discussed. A higher emission rate and different relative composition of the emission was seen to occur when the vegetation was wet, as compared to dry vegetation. The emission from the pine forest floor was seen to have a composition different from that of the crown and a seasonality of the rate similar to that of the crown. The ground emission could not be explained by sources in the litter or ground vegetation alone, and it is suggested that the root system of the trees is also an emission source. The emission rate from the pine forest floor was of the order of 30% of the crown emission. The July rate of emission from the crown of Scots pine, normalized to 20[degrees]C and averaged over four sites in Sweden, was 0.8 [plus minus] 0.4 [mu]g (gdw (grams dry weight) h)[sup [minus]1], and for Norwegian spruce, 0.5 [plus minus] 0.7 [mu]g(gdw h)[sup [minus]1]. It would seem that previous regional and global estimates of hydrocarbon fluxes to the atmosphere have used emission factors which are too high for boreal coniferous forests. 52 refs., 8 figs., 9 tabs.

  19. Seasonal variation of BVOC emissions from Norway spruce

    NASA Astrophysics Data System (ADS)

    Wang, Min; Schurgers, Guy; Ekberg, Anna; Arneth, Almut; Holst, Thomas

    2015-04-01

    Biogenic volatile organic compounds (BVOCs) are known as a source of secondary organic aerosols (SOA) due to their high reactivity in the atmosphere [1, 2]. Dominant boreal forest species (pine, spruce and birch) have been considered to be high monoterpene (MT) emitters [3, 4], and BVOC emissions and compound composition vary considerably under different temperature and light conditions through growing season [5, 6]. We characterize the canopy BVOC emissions variation from a Norway spruce dominated boreal forest in Central Sweden (Norunda, 60°05'N, 17°29'E). Air samples were taken during growing season (June to September 2013) from transparent dynamic branch chambers set up on Norway spruce at 20m agl. using a scaffolding tower. Air samples were collected every hour from the chamber with Tenax-TA adsorbent tubes and a pocket pump, and analyzed later by gas chromatography and a mass selective detector (GC-MS) to quantify trapped terpenoid compounds. Total terpenoids emission rates in August were found to be highest even though the highest average air temperature was observed in July. Isoprene could not be detected in any sample in June and in most samples from September, but during peak season. Emissions of Isoprene, MT and sesquiterpenes (SQT) showed a clear diurnal pattern in July and August with highest emissions at noon time, however, the composition of terpenoids was slightly changing among different months. The most complex chemical composition with 13 different MT species occurred in late July, while 9 SQT species occurred in the middle of August. However, the fraction of dominant MT species (Limonene, α-Pinene, β-Pinene and Camphene) of the total terpenoids emission was almost constant throughout the whole season from June to September except for β-Pinene which showed a higher fraction in August. References [1]M.Ehn et al., 2014, Nature, 506(7489), 476-479. [2]M.Kulmala et al., 2004, Atmos. Environ., 4, 557-562. [3]J.Rinne et al., 2005, Boreal Environ

  20. Genetic signatures of natural selection in response to air pollution in red spruce (Picea rubens, Pinaceae).

    PubMed

    Bashalkhanov, Stanislav; Eckert, Andrew J; Rajora, Om P

    2013-12-01

    One of the most important drivers of local adaptation for forest trees is climate. Coupled to these patterns, however, are human-induced disturbances through habitat modification and pollution. The confounded effects of climate and disturbance have rarely been investigated with regard to selective pressure on forest trees. Here, we have developed and used a population genetic approach to search for signals of selection within a set of 36 candidate genes chosen for their putative effects on adaptation to climate and human-induced air pollution within five populations of red spruce (Picea rubens Sarg.), distributed across its natural range and air pollution gradient in eastern North America. Specifically, we used FST outlier and environmental correlation analyses to highlight a set of seven single nucleotide polymorphisms (SNPs) that were overly correlated with climate and levels of sulphate pollution after correcting for the confounding effects of population history. Use of three age cohorts within each population allowed the effects of climate and pollution to be separated temporally, as climate-related SNPs (n = 7) showed the strongest signals in the oldest cohort, while pollution-related SNPs (n = 3) showed the strongest signals in the youngest cohorts. These results highlight the usefulness of population genetic scans for the identification of putatively nonneutral evolution within genomes of nonmodel forest tree species, but also highlight the need for the development and application of robust methodologies to deal with the inherent multivariate nature of the genetic and ecological data used in these types of analyses.

  1. Combined fluorescence, reflectance, and ground measurements of a stressed Norway spruce forest for forest damage assessment

    NASA Technical Reports Server (NTRS)

    Banninger, C.

    1991-01-01

    The detection and monitoring of stress and damage in forested areas is of utmost importance to forest managers for planning purposes. Remote sensing are the most suitable means to obtain this information. This requires that remote sensing data employed in a forest survey be properly chosen and utilized for their ability to measure canopy spectral features directly related to key tree and canopy properties that are indicators of forest health and vitality. Plant reflectance in the visible to short wave IR regions (400 to 2500 nm) provides information on its biochemical, biophysical, and morphological make up, whereas plant fluorescence in the 400 to 750 nm region is more indicative of the capacity and functioning of its photosynthetic apparatus. A measure of both these spectral properties can be used to provide an accurate assessment of stress and damage within the forest canopy. Foliar chlorophyll and nitrogen are essential biochemical constituents required for the proper functioning and maintenance of a plant's biological processes. Chlorophyll-a is the prime reactive center for photosynthesis, by which a plant converts CO2 and H2O into necessary plant products. Nitrogen forms an important component of the amino-acids, enzymes, proteins, alkaloids, and cyanogenic compounds that make up a plant, including its pigments. Both chlorophyll and nitrogen have characteristic absorption features in the visible to short wave IR region. By measuring the wavelength position and depth of these features and the fluorescence response of the foliage, the health and vitality of a canopy can be ascertained. Examples for a stressed Norway spruce forest in south-eastern Austria are presented.

  2. Evidence of Variscan and Alpine tectonics in the structural and thermochronological record of the central Serbo-Macedonian Massif (south-eastern Serbia)

    NASA Astrophysics Data System (ADS)

    Antić, Milorad D.; Kounov, Alexandre; Trivić, Branislav; Spikings, Richard; Wetzel, Andreas

    2016-07-01

    The Serbo-Macedonian Massif (SMM) represents a composite crystalline belt within the Eastern European Alpine orogen, outcropping from the Pannonian basin in the north to the Aegean Sea in the south. The central parts of this massif (south-eastern Serbia) consist of the medium- to high-grade Lower Complex and the low-grade Vlasina Unit. Outcrop- and micro-scale ductile structures in this area document three major stages of ductile deformation. The earliest stage D1 is related to isoclinal folding, commonly preserved as up to decimetre-scale quartz-feldspar rootless fold hinges. D2 is associated with general south-eastward tectonic transport and refolding of earlier structures into recumbent metre- to kilometre-scale tight to isoclinal folds. Stages D1 and D2 could not be temporally separated and probably took place in close sequence. The age of these two ductile deformation stages was constrained to the Variscan orogeny based on indirect geological evidence (i.e. ca. 408-ca. 328). During this period, the SMM was involved in a transpressional amalgamation of the western and eastern parts of the Galatian super-terrane and subsequent collision with Laurussia. Outcrop-scale evidence of the final stage D3 is limited to spaced and crenulation cleavage, which are probably related to formation of large-scale open upright folds as reported previously. 40Ar/39Ar thermochronology was applied on hornblende, muscovite, and biotite samples in order to constrain the age of tectonothermal events and activity along major shear zones. These 40Ar/39Ar data reveal three major cooling episodes affecting the central SMM. Cooling below greenschist facies conditions in the western part of the Vlasina Unit took place in a post-orogenic setting (extensional or transtensional) in the early Permian (284 ± 1 Ma). The age of activity along the top-to-the-west shear zone formed within the orthogneiss in the Božica area of the Vlasina Unit was constrained to Middle Triassic (246 ± 1 Ma). This

  3. Structure and segmentation of the eastern Gulf of Aden basin and the Sheba ridge from gravity, bathymetric and magnetic anomalies: implications for accretion processes

    NASA Astrophysics Data System (ADS)

    D'Acremont, E.; Leroy, S.; Maia, M.; Gente, P.; Autin, J.

    2007-12-01

    The eastern Gulf of Aden is a key place for investigating seafloor spreading processes and the evolution in space and time of the margin and ridge segmentation. The rifting of the Gulf that separated Arabia from Somalia started around 35 Ma ago followed by oceanic accretion from at least17.6 Ma. Bathymetric, gravity and magnetic data from the Encens-Sheba cruise are used to study the structure and segmentation of the eastern part of the basin and ridge, which have strong implications for accretion processes. The segmentation of the first oceanic spreading centre, which is dated at least 17.6 Ma by the magnetic anomaly (A5d) identification, seems to be directly related to the structural geometry of the margins. Then, magmatic processes governed the evolution of the segmentation. The segmentation of the oceanic crust evolved, by eastward propagation of the western segment, from three segments (from an5d to an5) to two segments (from an5). At 6 Ma (an3a) a third segment appeared by duplication of the Socotra transform fault, maybe due to a regional kinematics change. The Encens-Sheba oceanic domain is divided in two distinct areas trending NE-SW perpendicular to the Sheba ridge. (1) The Eastern area is characterized by a shorter wavelength variation of the axial segmentation with two spreading segments 30 to 40 km long, and by a thin crust particularly on the northern and southern ends of its flanks. (2) The Western zone, whose axial segment is more than 120 km long, is characterized by a thick crust and/or a hot mantle and no axial rift valley. This abnormal volcanic activity for a slow spreading ridge is emphasized by bathymetric highs with 5-10 km wide volcanic edifices, and by a negative anomaly of the MBA. These different results support the presence of an off-axis thermal anomaly located below the southern flank of the Sheba ridge. The magnetic anomalies and spreading asymmetry reveal that the location of this thermal anomaly might be relatively recent (~ 10 Ma

  4. Simultaneous Inversion of Interpolated Receiver Functions, Surface-wave Dispersion, and Gravity Observations for Lithospheric Structure Beneath the Eastern United States

    NASA Astrophysics Data System (ADS)

    Chai, C.; Ammon, C. J.; Maceira, M.; Herrmann, R. B.

    2015-12-01

    The unprecedented high-quality seismic data from Earthscope's Transportable Array provide us a great opportunity to investigate the subsurface structure beneath North America. Even with such a fine network, tightly constraining the 3D lithospheric structure is a challenge. Integrating complementary geophysical observations in simultaneous inversions has produced promising results. We have developed a receiver-function wavefield interpolation/smoothing method to enhance the complementariness of receiver functions and surface-wave dispersion. Combining information from adjacent seismic stations suppresses poorly sampled and difficult-to-interpret back-azimuthal variations and allows the stable extraction of the key features in the receiver-function wavefield. The interpolated receiver functions are inverted simultaneously with Rayleigh-wave phase and group velocities and Bouguer gravity observations to produce a robust estimate of the broad 3D shear-wave speed variations beneath the eastern United States. P-wave velocities and density variations are related to shear-speed using empirical velocity ratios and relations. We constrain the 3D variations to be laterally and vertically smooth. Application of the same methods to the western conterminous United States resulted in velocity images that are consistent with published models on the first order. With the completion of the Transportable Array deployment in the northeast US, the seismic dataset beneath the eastern U.S. region is complete. Preliminary inversions contain expected near-surface low shear-wave speeds associated with large basins and coastal regions and thicker crust beneath the interior compared with the coastal regions. Speeds in the upper mantle are generally typical, but the model includes several regions of relatively slow mantle beneath the northern Mississippi Embayment, the east coast, and beneath New England.

  5. Physiological diagnosis of the health of spruce and fir at high elevations in the Southern Appalachians

    SciTech Connect

    McLaughlin, S.B.; Wullschleger, S.D.; Stone, A. )

    1994-06-01

    A sequence of field studies to evaluate causes of decreasing radial growth rates of red spruce at high elevations in the Great Smokey Mountains during the past 30 years has examined climatic signals, competition, xylem wood chemistry, soil chemistry, foliar nutrition and carbon allocation patterns. The resultant hypothesis that acid deposition alters red spruce growth through limiting calcium availability, and consequently net carbon assimilation, has now been tested in controlled greenhouse and field studies. Recent measurements of reduce respiration and increased photosynthesis of red spruce samplings in response to adding calcium in the field, provides additional evidence linking acid deposition to altered nutrition, physiology, and growth of red spruce. Initial data from physiological gradient analysis also support the occurrence of parallel elevational gradients in physiology of fraser fir.

  6. A novel alkaline oxidation pretreatment for spruce, birch and sugar cane bagasse.

    PubMed

    Kallioinen, Anne; Hakola, Maija; Riekkola, Tiina; Repo, Timo; Leskelä, Markku; von Weymarn, Niklas; Siika-aho, Matti

    2013-07-01

    Alkaline oxidation pretreatment was developed for spruce, birch and sugar cane bagasse. The reaction was carried out in alkaline water solution under 10 bar oxygen pressure and at mild reaction temperature of 120-140°C. Most of the lignin was solubilised by the alkaline oxidation pretreatment and an easily hydrolysable carbohydrate fraction was obtained. After 72 h hydrolysis with a 10 FPU/g enzyme dosage, glucose yields of 80%, 91%, and 97%, for spruce, birch and bagasse, respectively, were achieved. The enzyme dosage could be decreased to 4 FPU/g without a major effect in terms of the hydrolysis performance. Compared to steam explosion alkaline oxidation was found to be significantly better in the conditions tested, especially for the pretreatment of spruce. In hydrolysis and fermentation at 12% d.m. consistency an ethanol yield of 80% could be obtained with both bagasse and spruce in 1-3 days.

  7. A novel alkaline oxidation pretreatment for spruce, birch and sugar cane bagasse.

    PubMed

    Kallioinen, Anne; Hakola, Maija; Riekkola, Tiina; Repo, Timo; Leskelä, Markku; von Weymarn, Niklas; Siika-aho, Matti

    2013-07-01

    Alkaline oxidation pretreatment was developed for spruce, birch and sugar cane bagasse. The reaction was carried out in alkaline water solution under 10 bar oxygen pressure and at mild reaction temperature of 120-140°C. Most of the lignin was solubilised by the alkaline oxidation pretreatment and an easily hydrolysable carbohydrate fraction was obtained. After 72 h hydrolysis with a 10 FPU/g enzyme dosage, glucose yields of 80%, 91%, and 97%, for spruce, birch and bagasse, respectively, were achieved. The enzyme dosage could be decreased to 4 FPU/g without a major effect in terms of the hydrolysis performance. Compared to steam explosion alkaline oxidation was found to be significantly better in the conditions tested, especially for the pretreatment of spruce. In hydrolysis and fermentation at 12% d.m. consistency an ethanol yield of 80% could be obtained with both bagasse and spruce in 1-3 days. PMID:23711947

  8. Climate and red spruce growth and decline in the northern Appalachians

    PubMed Central

    Johnson, A. H.; Cook, E. R.; Siccama, T. G.

    1988-01-01

    Between the mid-1960s and mid-1980s, red spruce (Picea rubens Sarg.) died at unusual rates on the mountains of New York and western New England. We determined the relationship between standardized tree ring widths and monthly climate data for calibration and verification periods from 1856 to 1981 and found that after about 1960, there was a distinct shift in the temperature variables related to standardized ring widths in vigorous spruce. The beginning of widespread spruce mortality, regionwide growth decreases, and the shift in response to climate in the early 1960s corresponds to the onset of a decade of unusually cold winters and several consecutive years when severe winter damage was noted across the Northeast in this species. We suggest that the episodes of winter damage are an important initiating and synchronizing factor in the red spruce decline. PMID:16593962

  9. Yellowheaded spruce sawfly: Its ecology and management. Forest Service general technical report

    SciTech Connect

    Katovich, S.A.; McCullough, D.G.; Haack, R.A.

    1995-12-14

    The yellowheaded spruce sawfly (YHSS), Pikonema alaskensis (Rohwer), (Hymenoptera: Tenthredinidae), defoliates spruce, Picea sp., throughout the Northern United States and Canada. YHSS defoliation can result in substantial growth reduction and tree mortality. Young, open-grown trees, 3 to 18 feet in height and 5 to 9 years old, are more vulnerable to YHSS damage than are understory trees, older trees, or trees in dense stands. Young plantations and naturally regenerated stands of spruce YHSS defoliation, particularly in the Great Lakes region. Many Christmans trees, nursery stock, roadside and windbreak trees, and ornamental spruce are also damaged. Susceptibility to YHSS drops sharply once trees reach 10 to 12 years of age and stands reach the stage of crown closure.

  10. Red spruce decline in the northeastern US: hypotheses regarding the role of acid rain

    SciTech Connect

    Johnson, A.H.

    1983-11-01

    Red spruce have died in abnormal numbers in the high elevation forests of New York and New England during the past two decades while spruce in the southern Appalachians remain healthy. Investigations of insect damage, fungai pathogens, successional dynamics, competitive status, climate and weather patterns, and possible pollutant effects indicate that the decline was triggered by abiotic stress during the dry years of the 1960s. Tree response, as recorded in the pattern of annual rings, and the wide range of soil conditions in which spruce are declining, suggest drought or dry summers as key factors. Hypotheses regarding the role of acid deposition induced stress have been offered, but at present there is not evidence which clearly links acid deposition to spruce decline. Indirect effects of acid deposition on soils, direct effects of acid deposition on foliage, and interactions of acid deposition and drought stress are possible but unproven pathways by which acid deposition could be involved. 23 references, 6 figures, 1 table.

  11. Emission of volatile sulfur compounds from spruce trees

    SciTech Connect

    Rennenberg, H.; Huber, B.; Schroeder, P.; Stahl, K.; Haunold, W.; Georgil, H.W.; Slovik, S.; Pfanz, H. )

    1990-03-01

    Spruce (Picea abies L.) trees from the same clone were supplied with different, but low, amounts of plant available sulfate in the soil (9.7-18.1 milligrams per 100 grams of soil). Branches attached to the trees were enclosed in a dynamic gas exchange cuvette and analyzed for the emission of volatile sulfur compounds. Independent of the sulfate supply in the soil, H{sub 2}S was the predominant reduced sulfur compound continuously emitted from the branches with high rates during the day and low rates in the night. In the light, as well as in the dark, the rates of H{sub 2}S emission increased exponentially with increasing water vapor flux from the needles. Approximately 1 nanomole of H{sub 2}S was found to be emitted per mole of water. When stomata were closed completely, only minute emission of H{sub 2}S was observed. Apparently, H{sub 2}S emission from the needles is highly dependent on stromatal aperture, and permeation through the cuticle is negligible. In several experiments, small amounts of dimethylsulfide and carbonylsulfide were also detected in a portion of the samples. However, SO{sub 2} was the only sulfur compound consistently emitted from branches of spruce trees in addition to H{sub 2}S. Emission of SO{sub 2} mainly proceeded via an outburst starting before the beginning of the light period. The total amount of SO{sub 2} emitted from the needles during this outburst was correlated with the plant available sulfate in the soil. The diurnal changes in sulfur metabolism that may result in an outburst of SO{sub 2} are discussed.

  12. Spatial variability in phytoplankton community structure along the eastern Arabian Sea during the onset of south-west monsoon

    NASA Astrophysics Data System (ADS)

    Ahmed, Ayaz; Kurian, Siby; Gauns, Mangesh; Chndrasekhararao, A. V.; Mulla, Amara; Naik, Bhagyashri; Naik, Hema; Naqvi, S. W. A.

    2016-05-01

    The Arabian Sea experiences moderate to weak upwelling along the south-west coast of India, which subsequently propagates towards the north. This causes variation in plankton community composition, which is addressed in the present study. Here we report the spatial variations in distribution of phytoplankton groups along the north-south transect in the eastern Arabian Sea based on marker pigments supported with flow-cytometric and microscopic analyses. 15 phytoplankton pigments were identified using High-performance liquid chromatography (HPLC) and the chemotaxonomic software (CHEMTAX) analysis associated these to seven major group of phytoplankton. The phytoplankton biomass, chlorophyll a (Chl a) was higher in southern stations with dominance of fucoxanthin whereas, divinyl chlorophyll a (divinyl Chl a), marker pigment of Prochlorococcus was present only in the northern region. Microscopic observation revealed the dominance of larger forms; diatoms (Chaetoceros coarctatum and Nitzschia sp.) and dinoflagellates (Scrippsiella sp., Oxytoxum nanum and Oxytoxum sp.) in the southern region. Furthermore, a study of plankton size distribution showed dominance of picoplankton (fpico) followed by nanoplankton (fnano) along the northern stations with comparatively higher microplankton (fmicro) in the south. This study clearly showed the influence of different environmental conditions on the phytoplankton community as reflected in dominance of diatoms in the southern (south of 12 °N) and that of picoplankton in the northern (north of 12 °N) region.

  13. Crustal velocity structure associated with the eastern Tennessee seismic zone: Vp and Vs images based upon local earthquake tomography

    NASA Astrophysics Data System (ADS)

    Powell, Christine A.; Withers, Mitchell M.; Cox, Randel Tom; Vlahovic, Gordana; Arroucau, Pierre

    2014-01-01

    We present three-dimensional P and S wave velocity models for the active eastern Tennessee seismic zone (ETSZ) using arrival time data from more than 1000 local earthquakes. A nonlinear tomography method is used that involves sequential inversion for model and hypocenter parameters. We image several velocity anomalies that persist through most of the inversion volume. Some anomalies support the presence of known features such as an ancient rift zone in northern Tennessee. Other anomalies reveal the presence of basement features that can be correlated with regional gravity and magnetic anomalies. We image a narrow, NE-SW trending, steeply dipping zone of low velocities that extends to a depth of at least 24 km and is associated with the vertical projection of the prominent New York-Alabama magnetic lineament. The low-velocity zone may have an apparent dip to the SE at depths exceeding 15 km. The low-velocity zone is interpreted as a major basement fault juxtaposing Granite-Rhyolite basement to the NW from Grenville southern Appalachian basement to the SE. Relocated hypocenters align in near-vertical segments suggesting reactivation of a distributed zone of deformation associated with a major strike-slip fault. We suggest that the ETSZ represents reactivation of an ancient shear zone established during formation of the super continent Rodinia.

  14. A spruce gene map infers ancient plant genome reshuffling and subsequent slow evolution in the gymnosperm lineage leading to extant conifers

    PubMed Central

    2012-01-01

    Background Seed plants are composed of angiosperms and gymnosperms, which diverged from each other around 300 million years ago. While much light has been shed on the mechanisms and rate of genome evolution in flowering plants, such knowledge remains conspicuously meagre for the gymnosperms. Conifers are key representatives of gymnosperms and the sheer size of their genomes represents a significant challenge for characterization, sequencing and assembling. Results To gain insight into the macro-organisation and long-term evolution of the conifer genome, we developed a genetic map involving 1,801 spruce genes. We designed a statistical approach based on kernel density estimation to analyse gene density and identified seven gene-rich isochors. Groups of co-localizing genes were also found that were transcriptionally co-regulated, indicative of functional clusters. Phylogenetic analyses of 157 gene families for which at least two duplicates were mapped on the spruce genome indicated that ancient gene duplicates shared by angiosperms and gymnosperms outnumbered conifer-specific duplicates by a ratio of eight to one. Ancient duplicates were much more translocated within and among spruce chromosomes than conifer-specific duplicates, which were mostly organised in tandem arrays. Both high synteny and collinearity were also observed between the genomes of spruce and pine, two conifers that diverged more than 100 million years ago. Conclusions Taken together, these results indicate that much genomic evolution has occurred in the seed plant lineage before the split between gymnosperms and angiosperms, and that the pace of evolution of the genome macro-structure has been much slower in the gymnosperm lineage leading to extent conifers than that seen for the same period of time in flowering plants. This trend is largely congruent with the contrasted rates of diversification and morphological evolution observed between these two groups of seed plants. PMID:23102090

  15. Plasticity and Evolution of (+)-3-Carene Synthase and (−)-Sabinene Synthase Functions of a Sitka Spruce Monoterpene Synthase Gene Family Associated with Weevil Resistance*

    PubMed Central

    Roach, Christopher R.; Hall, Dawn E.; Zerbe, Philipp; Bohlmann, Jörg

    2014-01-01

    The monoterpene (+)-3-carene is associated with resistance of Sitka spruce against white pine weevil, a major North American forest insect pest of pine and spruce. High and low levels of (+)-3-carene in, respectively, resistant and susceptible Sitka spruce genotypes are due to variation of (+)-3-carene synthase gene copy number, transcript and protein expression levels, enzyme product profiles, and enzyme catalytic efficiency. A family of multiproduct (+)-3-carene synthase-like genes of Sitka spruce include the three (+)-3-carene synthases, PsTPS-3car1, PsTPS-3car2, PsTPS-3car3, and the (−)-sabinene synthase PsTPS-sab. Of these, PsTPS-3car2 is responsible for the relatively higher levels of (+)-3-carene in weevil-resistant trees. Here, we identified features of the PsTPS-3car1, PsTPS-3car2, PsTPS-3car3, and PsTPS-sab proteins that determine different product profiles. A series of domain swap and site-directed mutations, supported by structural comparisons, identified the amino acid in position 596 as critical for product profiles dominated by (+)-3-carene in PsTPS-3car1, PsTPS-3car2, and PsTPS-3car3, or (−)-sabinene in PsTPS-sab. A leucine in this position promotes formation of (+)-3-carene, whereas phenylalanine promotes (−)-sabinene. Homology modeling predicts that position 596 directs product profiles through differential stabilization of the reaction intermediate. Kinetic analysis revealed position 596 also plays a role in catalytic efficiency. Mutations of position 596 with different side chain properties resulted in a series of enzymes with different product profiles, further highlighting the inherent plasticity and potential for evolution of alternative product profiles of these monoterpene synthases of conifer defense against insects. PMID:25016016

  16. Distribution of long-range linkage disequilibrium and Tajima's D values in Scandinavian populations of Norway Spruce (Picea abies).

    PubMed

    Larsson, Hanna; Källman, Thomas; Gyllenstrand, Niclas; Lascoux, Martin

    2013-05-20

    The site frequency spectrum of mutations (SFS) and linkage disequilibrium (LD) are the two major sources of information in population genetics studies. In this study we focus on the levels of LD and the SFS and on the effect of sample size on summary statistics in 10 Scandinavian populations of Norway spruce. We found that previous estimates of a low level of LD were highly influenced by both sampling strategy and the fact that data from multiple loci were analyzed jointly. Estimates of LD were in fact heterogeneous across loci and increased within individual populations compared with the estimate from the total data. The variation in levels of LD among populations most likely reflects different demographic histories, although we were unable to detect population structure by using standard approaches. As in previous studies, we also found that the SFS-based test Tajima's D was highly sensitive to sample size, revealing that care should be taken to draw strong conclusions from this test when sample size is small. In conclusion, the results from this study are in line with recent studies in other conifers that have revealed a more complex and variable pattern of LD than earlier studies suggested and with studies in trees and humans that suggest that Tajima's D is sensitive to sample size. This has large consequences for the design of future association and population genetic studies in Norway spruce.

  17. Effects of prolonged drought on the anatomy of sun and shade needles in young Norway spruce trees.

    PubMed

    Gebauer, Roman; Volařík, Daniel; Urban, Josef; Børja, Isabella; Nagy, Nina Elisabeth; Eldhuset, Toril Drabløs; Krokene, Paal

    2015-11-01

    Predicted increases in the frequency and duration of drought are expected to negatively affect tree vitality, but we know little about how water shortage will influence needle anatomy and thereby the trees' photosynthetic and hydraulic capacity. In this study, we evaluated anatomical changes in sun and shade needles of 20-year-old Norway spruce trees exposed to artificial drought stress. Canopy position was found to be important for needle structure, as sun needles had significantly higher values than shade needles for all anatomical traits (i.e., cross-sectional needle area, number of tracheids in needle, needle hydraulic conductivity, and tracheid lumen area), except proportion of xylem area per cross-sectional needle area. In sun needles, drought reduced all trait values by 10-40%, whereas in shade needles, only tracheid maximum diameter was reduced by drought. Due to the relatively weaker response of shade needles than sun needles in drought-stressed trees, the difference between the two needle types was reduced by 25% in the drought-stressed trees compared to the control trees. The observed changes in needle anatomy provide new understanding of how Norway spruce adapts to drought stress and may improve predictions of how forests will respond to global climate change. PMID:26640676

  18. Effects of prolonged drought on the anatomy of sun and shade needles in young Norway spruce trees.

    PubMed

    Gebauer, Roman; Volařík, Daniel; Urban, Josef; Børja, Isabella; Nagy, Nina Elisabeth; Eldhuset, Toril Drabløs; Krokene, Paal

    2015-11-01

    Predicted increases in the frequency and duration of drought are expected to negatively affect tree vitality, but we know little about how water shortage will influence needle anatomy and thereby the trees' photosynthetic and hydraulic capacity. In this study, we evaluated anatomical changes in sun and shade needles of 20-year-old Norway spruce trees exposed to artificial drought stress. Canopy position was found to be important for needle structure, as sun needles had significantly higher values than shade needles for all anatomical traits (i.e., cross-sectional needle area, number of tracheids in needle, needle hydraulic conductivity, and tracheid lumen area), except proportion of xylem area per cross-sectional needle area. In sun needles, drought reduced all trait values by 10-40%, whereas in shade needles, only tracheid maximum diameter was reduced by drought. Due to the relatively weaker response of shade needles than sun needles in drought-stressed trees, the difference between the two needle types was reduced by 25% in the drought-stressed trees compared to the control trees. The observed changes in needle anatomy provide new understanding of how Norway spruce adapts to drought stress and may improve predictions of how forests will respond to global climate change.

  19. AmeriFlux CA-Man Manitoba - Northern Old Black Spruce (former BOREAS Northern Study Area)

    SciTech Connect

    Amiro, Brian

    2016-01-01

    This is the AmeriFlux version of the carbon flux data for the site CA-Man Manitoba - Northern Old Black Spruce (former BOREAS Northern Study Area). Site Description - 55.880° N, 98.481° W, elevation of 259 m, Boreal coniferous: Black spruce; occasional larch present in poorly-drained areas. Groundcover is moss (feathermosses and Sphagnum), Labrador Tea, Vaccinium, and willows are a main component of the understory. It was established in 1993 as a BOREAS site.

  20. Inclination distributions and size measurements of hemlock and red spruce needles

    NASA Technical Reports Server (NTRS)

    Kimes, Daniel S.; Smith, J. A.

    1991-01-01

    Needle inclination angle distributions were plotted for 5 and 10 deg intervals for two hemlock trees and one red spruce. The distributions for the hemlock and spruce were similar, with the peak normal angle occurring between 10 and 20 deg. These distributions are between two theoretical (planophile and spherical) leaf angle distributions. The results can be used as an input to radiative transfer models that require a distribution of the orientation of the scattering elements of the canopy.

  1. Beat sampling accuracy in estimating spruce spider mite (Acari: Tetranychidae) populations and injury on juniper.

    PubMed

    Shrewsbury, Paula M; Hardin, Mark R

    2004-08-01

    The use of a standardized beat sampling method for estimating spruce spider mite, Oligonychus ununguis (Jacobi) (Acari: Tetranychidae), densities on a widely used evergreen ornamental plant species, Juniperus chinensis variety 'Sargentii' A. Henry (Cupressaceae), was examined. There was a significant positive relationship between total spruce spider mite densities and spider mite densities from beat sampling on juniper. The slope and intercept of the relationship may be used by pest managers to predict total spider mite densities on plants from beat sample counts. Beat sampling dramatically underestimates the total number of spider mites on a foliage sample. The relationships between spruce spider mite feeding injury and spider mite density estimates from beat sampling juniper foliage and total spider mite counts on foliage were also examined. There was a significant positive relationship between spruce spider mite density as estimated from beat sampling and injury to the plants. There was a similar positive relationship between the total number of spruce spider mites and injury to the plants, suggesting that a pest manager could use beat sampling counts to estimate plant injury and related thresholds. These findings have important implications to decision-making for spruce spider mite control, especially as it relates to threshold levels and determining rates of predator releases. Further assessment of the effectiveness of beat and other sampling methods across multiple spider mite- host plant associations needs to be examined to enable pest managers to select sampling plans that are feasible and reliable.

  2. Genetical Genomics Identifies the Genetic Architecture for Growth and Weevil Resistance in Spruce

    PubMed Central

    Porth, Ilga; White, Richard; Jaquish, Barry; Alfaro, René; Ritland, Carol; Ritland, Kermit

    2012-01-01

    In plants, relationships between resistance to herbivorous insect pests and growth are typically controlled by complex interactions between genetically correlated traits. These relationships often result in tradeoffs in phenotypic expression. In this study we used genetical genomics to elucidate genetic relationships between tree growth and resistance to white pine terminal weevil (Pissodes strobi Peck.) in a pedigree population of interior spruce (Picea glauca, P. engelmannii and their hybrids) that was growing at Vernon, B.C. and segregating for weevil resistance. Genetical genomics uses genetic perturbations caused by allelic segregation in pedigrees to co-locate quantitative trait loci (QTLs) for gene expression and quantitative traits. Bark tissue of apical leaders from 188 trees was assayed for gene expression using a 21.8K spruce EST-spotted microarray; the same individuals were genotyped for 384 SNP markers for the genetic map. Many of the expression QTLs (eQTL) co-localized with resistance trait QTLs. For a composite resistance phenotype of six attack and oviposition traits, 149 positional candidate genes were identified. Resistance and growth QTLs also overlapped with eQTL hotspots along the genome suggesting that: 1) genetic pleiotropy of resistance and growth traits in interior spruce was substantial, and 2) master regulatory genes were important for weevil resistance in spruce. These results will enable future work on functional genetic studies of insect resistance in spruce, and provide valuable information about candidate genes for genetic improvement of spruce. PMID:22973444

  3. Food-web structure and elemental (C, N and P) fluxes in the eastern tropical North Atlantic

    NASA Astrophysics Data System (ADS)

    Agusti, Susana; Duarte, Carlos M.; Vaqué, Dolors; Hein, Mette; Gasol, Josep M.; Vidal, Montserrat

    The flow of biogenic elements (C, N and P) in the upper tropical Atlantic (29-5°N, 17-22°W) was studied are the Latitude II cruise 1995 with the aim of examining planktonic food web controls on these fluxes. The transect crossed three distinct water masses, a productive area in the NW African upwelling between 18° and 23°N, a warm water mass between 6° and 13°N, and unproductive waters at the northern range of the transect. The contrasting phytoplankton biomass and production across the eastern tropical N Atlantic appear to be driven by major changes in the supply of nutrients across the thermocline, which averaged 0.99±0.6 mmol N m -2 d -1 and 0.13±0.07 mmol P m -2 d -1, and was particularly high at the upwelling area off NW Africa. This supply was relatively low in nitrogen relative to that in phosphorus, driving the system towards nitrogen limitation. This was further enhanced by an important sinking flux of nitrogen in dissolved organic matter, which averaged 14.8±6.8 mmol N m -2 d -1 and exceeded the average internal input of nitrate by seven fold. There was, therefore, a severe nitrogen deficit that must have been offset by lateral and/or atmospheric inputs. Evaluation of the likely size of these sources indicates that lateral inputs must be important in the southern range of the area investigated, while atmospheric inputs seem to deliver the required N to balance losses in the northern area of the eastern subtropical Atlantic. Planktonic biomass was dominated by heterotrophs, and community respiration rates and bacterial carbon requirements exceeded the net primary production in the most unproductive areas. There must be significant allocthonous inputs of organic carbon and nitrogen in this region of the ocean. The distribution of primary production was uncoupled to that of the calculated CO 2 flux across the ocean surface. The productive area off the NW African upwelling acted as a source of CO 2 to the atmosphere (0.5 mmol C m -2 d -1), driven by

  4. Correlation by Rb-Sr geochronology of garnet growth histories from different structural levels within the Tauern Window, Eastern Alps

    SciTech Connect

    Christensen, John N.; Selverstone, Jane; Rosenfeld, John L.; Depaolo, Donald J.

    1993-06-01

    In order to evaluate rates of tectonometamorphic processes, growth rates of garnets from metamorphic rocks of the Tauern Window, Eastern Alps were measured using Rb-Sr isotopes. The garnet growth rates were determined from Rb-Sr isotopic zonation of single garnet crystals and the Rb-Sr isotopic compositions of their associated rock matrices. Garnets were analyzed from the Upper Schieferhulle (USH) and Lower Schieferhulle (LSH) within the Tauern Window. Two garnets from the USH grew at rates of 0.67(-0.13)+0.19 mm/million years and 0.88(-0.19)+0.34 mm/million years, respectively, indicating an average growth duration of 5.4 +- 1.7 million years. The duration of growth coupled with the amount of rotation recorded by inclusion trails in the USH garnets yields an average shear-strain rate during garnet growth of 2.7(-0.7)+1.2 x 10(-14) s-1 . Garnet growth in the sample from the USH occurred between 35.4 +- 0.6 and 30 +- 0.8 Ma. The garnet from the LSH grew at a rate of 0.23 +- 0.015 mm/mil lion years, between 62 +- 1.5 Ma and 30.2 +- 1.5 Ma. Contemporaneous cessation of garnet growth in both units at approximately 30 Ma is in accord with previous dating of the thermal peak of metamorphism in the Tauern Window. Correlation with previously published pressure-temperature paths for garnets from the USH and LSH yields approximate rates of burial, exhumation and heating during garnet growth. Assuming that these P - T paths are applicable to the garnets in this study, the contemporaneous exhumation rates recorded by garnet in the USH and LSH were approximately 4(-2)+3 mm/year and 2 +- 1 mm/year, respectively. [References: 34

  5. Structure of a Young Oceanic Basin: Results of the Encens-Sheba Cruise in the Eastern Gulf of Aden

    NASA Astrophysics Data System (ADS)

    Leroy, S.; Fournier, M.; Gente, P.; Al-Kathiri, A.; Bellahsen, N.; Beslier, M.; Blais, A.; d'Acremont, E.; Mercouriev, S.; Patriat, P.; Perrot, J.

    2001-12-01

    The Encens-Sheba cruise was carried out in July 2000 aboard R/V Marion Dufresne in the eastern part of the Gulf of Aden. It produced a complete swath bathymetry, gravity and magnetic data set between southern Oman (Dhofar) and Yemen in the north and Socotra island in the south, and between Alula-Fartak fracture zone in the west and the Socotra fracture zone in the east. The new data set shows a complete section of a young oceanic basin from conjugate passive continental margins up to the oceanic spreading center. On the conjugate passive margins, single channel seismic data have been also collected. The two conjugate margins are steep, narrow and asymmetric. Titled blocks, horsts and grabens bounded by faults dipping towards the ocean or the continent compose the northern margin, whereas a deep basin near the continental slope in the vicinity of the continent-ocean transition characterizes the southern margin. The two margins are divided by transfer faults in 3 major segments. The continent-ocean transition is marked by a negative gradient of the free-air gravity anomalies. The present Sheba ridge is divided by discontinuities in 3 segments from 20 to 100 km long. The central part of the 100-km-long western segment is characterized by an axial uplift reflecting a large magmatic activity. The two other shorter segments display axial valley more classical for a slow spreading ridge (1.1cm/a). Although the magnetic anomalies show a complex history of oceanic accretion, the evolution of the segmentation can be followed. Indeed, the segmentation of the conjugate passive margins seems to be correlated with the location of early traces of identifiable Sheba ridge offset. The oldest anomalies are identified as an5C-an5D. The Gulf of Aden in this area thus opened 16-17Ma ago, which is significantly older than the 12-13 Ma suggested by previous studies.

  6. Spatial Genetic Analyses Reveal Cryptic Population Structure and Migration Patterns in a Continuously Harvested Grey Wolf (Canis lupus) Population in North-Eastern Europe

    PubMed Central

    Hindrikson, Maris; Remm, Jaanus; Männil, Peep; Ozolins, Janis; Tammeleht, Egle; Saarma, Urmas

    2013-01-01

    Spatial genetics is a relatively new field in wildlife and conservation biology that is becoming an essential tool for unravelling the complexities of animal population processes, and for designing effective strategies for conservation and management. Conceptual and methodological developments in this field are therefore critical. Here we present two novel methodological approaches that further the analytical possibilities of STRUCTURE and DResD. Using these approaches we analyse structure and migrations in a grey wolf (Canislupus) population in north-eastern Europe. We genotyped 16 microsatellite loci in 166 individuals sampled from the wolf population in Estonia and Latvia that has been under strong and continuous hunting pressure for decades. Our analysis demonstrated that this relatively small wolf population is represented by four genetic groups. We also used a novel methodological approach that uses linear interpolation to statistically test the spatial separation of genetic groups. The new method, which is capable of using program STRUCTURE output, can be applied widely in population genetics to reveal both core areas and areas of low significance for genetic groups. We also used a recently developed spatially explicit individual-based method DResD, and applied it for the first time to microsatellite data, revealing a migration corridor and barriers, and several contact zones. PMID:24069446

  7. Seismic velocity structure of the crust and shallow mantle of the Central and Eastern United States by seismic surface wave imaging

    USGS Publications Warehouse

    Pollitz, Fred; Mooney, Walter D.

    2016-01-01

    Seismic surface waves from the Transportable Array of EarthScope's USArray are used to estimate phase velocity structure of 18 to 125 s Rayleigh waves, then inverted to obtain three-dimensional crust and upper mantle structure of the Central and Eastern United States (CEUS) down to ∼200 km. The obtained lithosphere structure confirms previously imaged CEUS features, e.g., the low seismic-velocity signature of the Cambrian Reelfoot Rift and the very low velocity at >150 km depth below an Eocene volcanic center in northwestern Virginia. New features include high-velocity mantle stretching from the Archean Superior Craton well into the Proterozoic terranes and deep low-velocity zones in central Texas (associated with the late Cretaceous Travis and Uvalde volcanic fields) and beneath the South Georgia Rift (which contains Jurassic basalts). Hot spot tracks may be associated with several imaged low-velocity zones, particularly those close to the former rifted Laurentia margin.

  8. 2D magnetotelluric imaging of the Anqing-Guichi ore district, Yangtze metallogenic belt, eastern China: An insight into the crustal structure and tectonic units

    NASA Astrophysics Data System (ADS)

    Chen, Xiangbin; Yan, Jiayong

    2016-08-01

    Two parallel NW-trending magnetotelluric (MT) profiles were placed perpendicularly to the main structures of the Anqing-Guichi ore district, one of the seven ore districts in the middle-lower Yangtze River metallogenic belt of eastern China. In October-December 2013, the MT data acquisition was carried out at 117 sites with 0.5-1 km site spacing. The MT data has a good quality in the frequency range between 320 and 0.01 Hz. The dimensionality analysis and 2D resistivity inversion results indicate that: (1) the deep of the ore district with three-dimensional structural characteristics, but two-dimensional structural characteristics for shallow; (2) there is a clear correlation between resistivity and the main geological units of the ore district, as well as correlation with mapped surface faults; (3) the Gandan deep fault (GDF) and Jiangnan deep fault (JNF) extend from the surface to 10 km deep, with dip of NW45°, and dip angles larger than 60°. A series of NE-trending acidic intrusive rocks were controlled by the GDF.

  9. Finders keepers, losers weepers - drought as a modifier of competition between European beech and Norway spruce -

    NASA Astrophysics Data System (ADS)

    Goisser, Michael; Blanck, Christian; Geppert, Uwe; Häberle, Karl-Heinz; Matyssek, Rainer; Grams, Thorsten E. E.

    2016-04-01

    Mixed stands of European beech (Fagus sylvatica L.) and Norway spruce (Picea abies (L.) Karst.) frequently reflect over-yielding, when compared to respective monospecific stands. Over-yielding is attributed to enhanced resource uptake efficiency through niche complementarity alleviating species competition. Under climate change, however, with severe and frequent summer drought, water limitation may become crucial in modifying the competitive interaction between neighboring beech and spruce trees. In view of the demands by silvicultural practice, basic knowledge from experimental field work about competitive versus facilitative interaction in maturing mixed beech-spruce forests is scarce. To this end, we investigate species-specific drought response including underlying mechanisms of species interaction in a maturing group-wise mixed beech-spruce forest, amongst 60 and 53 adult trees of beech and spruce, respectively (spruce 65 ± 2, beech 85 ± 4 years old). Severe and repeated experimental drought is being induced over several years through a stand-scale approach of rain throughfall exclusion (Kranzberg Forest Roof Experiment, KROOF). The experimental design comprises 6 roofed (E, automated, closing only during rain) and 6 control (C) plots with a total area of almost 1800 square meters. In 2015 minimum predawn potentials of -2.16 MPa and -2.26 MPa were reached in E for beech and spruce respectively. At the leaf level, spruce displayed high drought susceptibility reflected by a distinct decrease in both stomatal conductance and net CO2 uptake rate by more than 80% each, suggesting isohydric response. Beech rather displayed anisohydry indicated by less pronounced yet significant reduction of stomatal conductance and net CO2 uptake rate by more than 55% and 45%, respectively. Under the C regime, a negative species interaction effect on stomatal conductance was found in beech, contrasting with a positive effect in spruce. However, drought reversed the effect of

  10. Multi-scale integrated structural and aeromagnetic analysis to guide tectonic models: An example from the eastern Musgrave Province, Central Australia

    NASA Astrophysics Data System (ADS)

    Aitken, Alan R. A.; Betts, Peter G.

    2009-10-01

    In many polydeformed Precambrian provinces, the understanding of their tectonic history is limited by insufficient exposure, which leads to difficulty in developing links between local and regional architecture, and results in poorly constrained models of their regional-scale architecture. High-resolution aeromagnetic data allows interpretation and modelling at a scale comparable to structural mapping, and by integrating these methods, these links can be developed with high confidence. As an example, we integrate structural mapping, aeromagnetic data interpretation and 3D magnetic inversions in a multi-scale structural analysis of the polydeformed eastern Musgrave Province in central Australia. Deformation during the ca. 1200 Ma Musgravian Orogeny is characterised by an originally northeast-trending structural grain defined by shear zones and tight to isoclinal folds at all scales. 3D magnetic inversion models of syn-to-post Musgravian granites indicate that this structural grain is predominantly west dipping. This architecture suggests that a northeast-oriented fold and thrust belt developed as a result of northwest-southeast compression in central Australia during the Musgravian Orogeny. The next deformation event is characterised by an array of originally east-southeast trending shear zones that truncate and offset the Musgravian structural grain with apparent dextral displacements of up to 1 km. These shear zones are commonly co-located with mafic and granitic dykes, suggesting a dilational origin, and they are interpreted to have initiated under northeast-southwest extension during the ca. 1080 Ma Giles Event. The architecture of these two Mesoproterozoic events was later extensively overprinted by two deformation events during the Pan-African aged (ca. 550 Ma) Petermann Orogeny. The first is characterised by the widespread reactivation of Mesoproterozoic structures, resulting in a network of shear zones of variable orientation, and the second is characterised

  11. Structural development of the central Kyrenia Range (north Cyprus) in its regional setting in the eastern Mediterranean region

    NASA Astrophysics Data System (ADS)

    Robertson, A. H. F.; Kinnaird, T. C.

    2016-01-01

    A detailed structural analysis of the Mesozoic-Cenozoic geological development of the central segment of the Kyrenia Range in its regional tectonic context is given here. The structural evidence comes from five structural traverses, outcrop observations, small-scale structures and related regional evidence. The majority of the structures are fault planes, of which a subordinate number exhibit slickenlines (fault plane data, n = 2688; with kinematics, n = 537). Additional kinematic data were obtained from C-S fabrics and folds. Small-scale structures in each stratigraphic unit were `backstripped' to reveal relative chronology. Synthesis of the structural information indicates three phases of convergence-related deformation: (1) Late Cretaceous, associated with greenschist facies metamorphism, followed by exhumation that was probably associated with WNW-ESE to ENE-WSW-trending high-angle faulting; (2) Mid-Eocene, associated with southward thrusting, coupled with ~N-S strike-slip (transfer faulting) and oblique faulting in an overall sinistral transpressive stress regime; (3) Late Miocene-earliest Pliocene, involving southward thrusting and folding, localised back-thrusting, extensive fault reactivation and large-scale segmentation of the range. Intense uplift of the Kyrenia Range took place during the Plio-Pleistocene, possibly related to the collision of the Eratosthenes Seamount with the Cyprus trench to the south of the island. The three main convergent phases relate to stages of northward subduction and diachronous continental collision affecting the northerly, active continental margin of the Southern Neotethys.

  12. Modular organization of the white spruce (Picea glauca) transcriptome reveals functional organization and evolutionary signatures.

    PubMed

    Raherison, Elie S M; Giguère, Isabelle; Caron, Sébastien; Lamara, Mebarek; MacKay, John J

    2015-07-01

    Transcript profiling has shown the molecular bases of several biological processes in plants but few studies have developed an understanding of overall transcriptome variation. We investigated transcriptome structure in white spruce (Picea glauca), aiming to delineate its modular organization and associated functional and evolutionary attributes. Microarray analyses were used to: identify and functionally characterize groups of co-expressed genes; investigate expressional and functional diversity of vascular tissue preferential genes which were conserved among Picea species, and identify expression networks underlying wood formation. We classified 22 857 genes as variable (79%; 22 coexpression groups) or invariant (21%) by profiling across several vegetative tissues. Modular organization and complex transcriptome restructuring among vascular tissue preferential genes was revealed by their assignment to coexpression groups with partially overlapping profiles and partially distinct functions. Integrated analyses of tissue-based and temporally variable profiles identified secondary xylem gene networks, showed their remodelling over a growing season and identified PgNAC-7 (no apical meristerm (NAM), Arabidopsis transcription activation factor (ATAF) and cup-shaped cotyledon (CUC) transcription factor 007 in Picea glauca) as a major hub gene specific to earlywood formation. Reference profiling identified comprehensive, statistically robust coexpressed groups, revealing that modular organization underpins the evolutionary conservation of the transcriptome structure.

  13. Flux-based Water Use Efficiency over Two Larch Forests with Different Soil and Canopy Structure at Eastern Siberia

    NASA Astrophysics Data System (ADS)

    Kotani, A.; Ohta, T.; Maximov, T.

    2013-12-01

    This study investigated evapotranspiration and CO2 exchange, observed by the eddy covariance method, over two larch-dominated forests mixed with birch and willow, in the southern and middle parts of the Lena basin, eastern Siberia. One is the Spasskaya Pad station (SP) at Yakutsk (62° 15'N, 129° 14'E). The other station named Elgeeii station (EG) (60° 0'N, 133° 49'E) is located at 300 km southeast of Yakutsk. The dominant species in the forest is larch (Larix cajanderi), mixed with birch (Betula pendula), willow (Salix bebbiana) and pine (Pinus sylvesrtis). The stand density of larch trees is 720 and 1040 trees ha-1 (1800 and 2600 trees ha-1 including birch, salix and pine) at SP and EG, respectively. The mean stand height of upper canopy, which is comprised of larch trees, is around 20m at SP and 25m at EG, and plant area index measured in August 2011 is 1.4 at SP and 2.1 at EG. At SP forest, decline of some larch trees was recently observed. Observations of eddy-covariance and meteorological data in three growing seasons showed similarities in the seasonal variation of meteorological conditions and evapotranspiration (ET), while the two sites varied in gross CO2 uptake and ecosystem respiration during the growing season. An earlier onset of and a later decline of gross CO2 uptake (GPP) compared to ET at EG resulted in a seasonal time lag between ET and CO2 uptake and therefore mid-summer depletion of ecosystem WUE (defined as the ratio of net or gross CO2 uptake to ET), while relatively stable WUE was observed at SP. The total amount of ET from May to September was almost identical at the two sites, whereas the components of CO2 exchange (gross uptake and respiration) at EG were 1.31 times those recorded at SP. Consequently, ecosystem WUE was larger at EG than at SP. The difference in total GPP was explained mainly by difference in the PAI, that is, the tree biomass above ground, rather than the climate condition. It is likely that the level of forest

  14. Genetic variation and population structure of American mink Neovison vison from PCB-contaminated and non-contaminated locales in eastern North America.

    PubMed

    Wirgin, Isaac; Maceda, Lorraine; Waldman, John; Mayack, David T

    2015-11-01

    American mink Neovison vison may be particularly vulnerable to toxicities of persistent contaminants such as PCBs because of their aquatic-based diet, position near the top of the food web, and small deme sizes. Furthermore, ranched mink are sensitive to reproductive toxicities of fish diets from PCB-polluted sites. The upper Hudson River is highly contaminated with PCBs and previous studies have shown elevated hepatic burdens of total and coplanar PCBs in mink collected near the river compared with those from more distant locales in New York and elsewhere. We hypothesized that bioaccumulation of PCBs in Hudson River mink has reduced their levels of genetic diversity or altered their genetic population structure. To address this, we conducted microsatellite DNA analysis on collections made in proximity to and from more distant locales in the Hudson River watershed, elsewhere in New York State, and at other sites in eastern North America including New Brunswick, four locales in Ontario, multiple drainages in Maine, and two ecoregions in Rhode Island. We did not find reduced genetic diversity at the individual or population levels in mink collected near (<6 km) to PCB hotspots in the Hudson River nor evidence of altered population structure. Consistent with their distribution in small localized and isolated demes, we did find significant genetic population structure among many mink collections in New York State and elsewhere. Depending on the analytical approach used, genetically distinct populations numbered between 16 when using STRUCTURE to 19-20 when using Exact G tests, F ST, or AMOVA analyses. Genetically distinct population units were found among major ecoregions and minor ecoregions in New York State, among different hydrologic subunits within the Hudson River watershed, among spatially separate locales in Ontario, and among most watersheds in Maine. However, despite this localization and potential heightened impact of stressors, genetic diversity and genetic

  15. Diversity and three-dimensional structures of the alpha Mcr of the methanogenic Archaea from the anoxic region of Tucuruí Lake, in Eastern Brazilian Amazonia

    PubMed Central

    Santana, Priscila Bessa; Junior, Rubens Ghilardi; Alves, Claudio Nahum; Silva, Jeronimo Lameira; McCulloch, John Anthony; Schneider, Maria Paula Cruz; da Costa da Silva, Artur

    2012-01-01

    Methanogenic archaeans are organisms of considerable ecological and biotechnological interest that produce methane through a restricted metabolic pathway, which culminates in the reaction catalyzed by the Methyl-coenzyme M reductase (Mcr) enzyme, and results in the release of methane. Using a metagenomic approach, the gene of the α subunit of mcr (mcrα) was isolated from sediment sample from an anoxic zone, rich in decomposing organic material, obtained from the Tucuruí hydroelectric dam reservoir in eastern Brazilian Amazonia. The partial nucleotide sequences obtained were 83 to 95% similar to those available in databases, indicating a low diversity of archaeans in the reservoir. Two orders were identified - the Methanomicrobiales, and a unique Operational Taxonomic Unit (OTU) forming a clade with the Methanosarcinales according to low bootstrap values. Homology modeling was used to determine the three-dimensional (3D) structures, for this the partial nucleotide sequence of the mcrα were isolated and translated on their partial amino acid sequences. The 3D structures of the archaean Mcrα observed in the present study varied little, and presented approximately 70% identity in comparison with the Mcrα of Methanopyrus klanderi. The results demonstrated that the community of methanogenic archaeans of the anoxic C1 region of the Tucurui reservoir is relatively homogeneous. PMID:22481885

  16. Genetic structure and diversity of natural and domesticated populations of Citrus medica L. in the Eastern Himalayan region of Northeast India.

    PubMed

    Barbhuiya, Atiqur R; Khan, Mohammed L; Dayanandan, Selvadurai

    2016-06-01

    Citron (Citrus medica L.) is a medicinally important species of citrus native to India and occurs in natural forests and home gardens in the foothills of the eastern Himalayan region of northeast India. The wild populations of citron in the region have undergone rapid decline due to natural and anthropogenic disturbances and most of the remaining individuals of citron are found in fragmented natural forests and home gardens in the region. In order to assess the genetic structure and diversity of citron in wild and domesticated populations, we analyzed 219 individuals of C. medica collected from four wild and eight domesticated populations using microsatellite markers. The genetic analysis based on five polymorphic microsatellite loci revealed an average of 13.40 allele per locus. The mean observed and expected heterozygosity values ranged between 0.220-0.540 and 0.438-0.733 respectively among the wild and domesticated populations. Domesticated populations showed close genetic relationships as compared to wild populations and pairwise Nei's genetic distance ranged from 0.062 to 2.091 among wild and domesticated populations. Analysis of molecular variance (AMOVA) showed higher genetic diversity among- than within populations. The analysis of population structure revealed five groups. Mixed ancestry of few individuals of different populations revealed exchange of genetic materials among farmers in the region. Citron populations in the region show high genetic variation. The knowledge gained through this study is invaluable for devising genetically sound strategies for conservation of citron genetic resources in the region. PMID:27516853

  17. Seasonal variations of marine protist community structure based on taxon-specific traits using the eastern English Channel as a model coastal system.

    PubMed

    Genitsaris, Savvas; Monchy, Sébastien; Viscogliosi, Eric; Sime-Ngando, Télesphore; Ferreira, Stéphanie; Christaki, Urania

    2015-05-01

    Previous microscopy-based studies in the eastern English Channel have revealed it to be a productive meso-eutrophic coastal ecosystem, characterized by strong repeating patterns in microplankton succession. The present study examines the seasonal structure of the entire protistan community from March 2011 to July 2013, using tag pyrosequencing of the V2-V3 hypervariable region of the 18S rRNA gene. A total of 1242 OTUs and 28 high-level taxonomic groups, which included previously undetected taxa in the area, were identified. The detected OTUs were considered according to taxon-specific traits, which included their trophic role, abundance and specialization level. Taxa differentiation based on specialization level rather than abundance was more informative in describing community organization. While generalists were always abundant, numerous specialists that were either rare or absent in most samples, increased in abundance for short periods, appearing to be overall abundant. Statistical and network analyses showed that the protistan seasonal organization was influenced by environmental parameters. It also highlighted that in addition to grazers, fungi and parasites played potentially significant roles during phytoplankton blooms. Overall, while the protistan succession was mainly shaped by environmental variations, biotic interactions among co-occurring taxa were the main structural drivers of the temporal assemblages.

  18. Research aircraft observations of the mesoscale and microscale structure of a cold front over the eastern Pacific Ocean

    NASA Technical Reports Server (NTRS)

    Bond, Nicholas A.; Shapiro, M. A.

    1991-01-01

    The structure of an oceanic cold front is described on the basis of research aircraft observations taken during the Ocean Storms field experiment. Synoptic and mesoscale analyses compare the structure of an upper-level jet-front system observed slightly downstream from the wind speed maximum to its structure in the upstream entrance region. Stratospheric potential vorticity and ozone were found within the frontal zone down to about 800 mb. Microscale analyses of the front near the sea surface were carried out for a portion of the front having the signature of a 'rope' cloud in satellite imagery. A narrow (less than 1 km) zone of upward motion (about 4 m/s) and of horizontal shear (about 0.01/s) characterized the front near the surface. Significant alongfront variability was found, including lateral displacements in the frontal zone where there were weaker updrafts.

  19. Comparing modern and presettlement forest dynamics of a subboreal wilderness: does spruce budworm enhance fire risk?

    PubMed

    Sturtevant, Brian R; Miranda, Brian R; Shinneman, Douglas J; Gustafson, Eric J; Wolter, Peter T

    2012-06-01

    Insect disturbance is often thought to increase fire risk through enhanced fuel loadings, particularly in coniferous forest ecosystems. Yet insect disturbances also affect successional pathways and landscape structure that interact with fire disturbances (and vice-versa) over longer time scales. We applied a landscape succession and disturbance model (LANDIS-II) to evaluate the relative strength of interactions between spruce budworm (Choristoneura fumiferana) outbreaks and fire disturbances in the Boundary Waters Canoe Area (BWCA) in northern Minnesota (USA). Disturbance interactions were evaluated for two different scenarios: presettlement forests and fire regimes vs. contemporary forests and fire regimes. Forest composition under the contemporary scenario trended toward mixtures of deciduous species (primarily Betula papyrifera and Populus spp.) and shade-tolerant conifers (Picea mariana, Abies balsamea, Thuja occidentalis), with disturbances dominated by a combination of budworm defoliation and high-severity fires. The presettlement scenario retained comparatively more "big pines" (i.e., Pinus strobus, P. resinosa) and tamarack (L. laricina), and experienced less budworm disturbance and a comparatively less-severe fire regime. Spruce budworm disturbance decreased area burned and fire severity under both scenarios when averaged across the entire 300-year simulations. Contrary to past research, area burned and fire severity during outbreak decades were each similar to that observed in non-outbreak decades. Our analyses suggest budworm disturbances within forests of the BWCA have a comparatively weak effect on long-term forest composition due to a combination of characteristics. These include strict host specificity, fine-scaled patchiness created by defoliation damage, and advance regeneration of its primary host, balsam fir (A. balsamea) that allows its host to persist despite repeated disturbances. Understanding the nature of the three-way interaction between

  20. Comparing modern and presettlement forest dynamics of a subboreal wilderness: Does spruce budworm enhance fire risk?

    USGS Publications Warehouse

    Sturtevant, Brian R.; Miranda, Brian R.; Shinneman, Douglas J.; Gustafson, Eric J.; Wolter, Peter T.

    2012-01-01

    Insect disturbance is often thought to increase fire risk through enhanced fuel loadings, particularly in coniferous forest ecosystems. Yet insect disturbances also affect successional pathways and landscape structure that interact with fire disturbances (and vice-versa) over longer time scales. We applied a landscape succession and disturbance model (LANDIS-II) to evaluate the relative strength of interactions between spruce budworm (Choristoneura fumiferana) outbreaks and fire disturbances in the Boundary Waters Canoe Area (BWCA) in northern Minnesota (USA). Disturbance interactions were evaluated for two different scenarios: presettlement forests and fire regimes vs. contemporary forests and fire regimes. Forest composition under the contemporary scenario trended toward mixtures of deciduous species (primarily Betula papyrifera and Populus spp.) and shade-tolerant conifers (Picea mariana, Abies balsamea, Thuja occidentalis), with disturbances dominated by a combination of budworm defoliation and high-severity fires. The presettlement scenario retained comparatively more “big pines” (i.e., Pinus strobus, P. resinosa) and tamarack (L. laricina), and experienced less budworm disturbance and a comparatively less-severe fire regime. Spruce budworm disturbance decreased area burned and fire severity under both scenarios when averaged across the entire 300-year simulations. Contrary to past research, area burned and fire severity during outbreak decades were each similar to that observed in non-outbreak decades. Our analyses suggest budworm disturbances within forests of the BWCA have a comparatively weak effect on long-term forest composition due to a combination of characteristics. These include strict host specificity, fine-scaled patchiness created by defoliation damage, and advance regeneration of its primary host, balsam fir (A. balsamea) that allows its host to persist despite repeated disturbances. Understanding the nature of the three-way interaction

  1. Stratigraphy, petrology, and structure of the Pingston terrane, Mount Hayes C-5 and C-6 quadrangles, eastern Alaska Range, Alaska

    NASA Astrophysics Data System (ADS)

    Nokleberg, W. J.; Schwab, C. E.; Miyaoka, R. T.; Buhrmaster, C. L.

    Recent field, petrologic, and structural studies of the Pingston terrane in the Mount Hayes C-5 and C-6 quandrangles reveal that in this area the terrane: (1) has a highly distinctive stratigraphy, age, petrology (relict textures, relict minerals, and metamorphic facies), and structure; and (2) differs markedly from that described in previous studies. These more recent studies indicate that the major rock types, in order of decreasing abundance, are meta-andesite, metadacite and metarhyodacite flows and (or) tuff, metabasalt, metagabbro, metavolcanic graywacke, metagray-wacke, metasiltstone, metaquartzite or metachert, and very sparse marble. The general petrography of the major rock units in the Pingston terrane is given.

  2. Forcing, properties, structure, and antecedent synoptic climatology of the Snake River Plain Convergence Zone of eastern Idaho: Analyses of observations and numerical simulations

    NASA Astrophysics Data System (ADS)

    Andretta, Thomas A.

    The Snake River Plain Convergence Zone (SPCZ) is a convergent shear zone generated by synoptic-scale post cold-frontal winds in the planetary boundary layer (PBL) interacting with the complex topography of eastern Idaho. The SPCZ produces clouds and occasional precipitation over time scales of 6--12 hours in a significant area of mesoscale dimensions (10--50 x 10 3 km2). This meso-beta-scale feature also contributes to the precipitation climatology in a semi-arid plain. The SPCZ is climatologically linked to the passage of synoptic-scale cold fronts and typically occurs in the fall and winter months with the highest frequencies in October, November, and January. The Snake River Plain of eastern Idaho is covered by a dense surface mesonetwork of towers with sensible weather measurements, single Doppler weather radar, regional soundings, and operational model sources. The ability of numerical weather prediction models to simulate the SPCZ depends on several factors: the accuracy of the large scale flow upstream of the zone, terrain resolution, grid scale, boundary layer parameterizations of stability, cumulus parameterizations, and microphysics schemes. This dissertation explores several of these issues with the aforementioned observations and with the Weather Research and Forecasting-Advanced Research WRF (WRF-ARW) model simulations of selected SPCZ events. This dissertation first explains the conceptual models of the flow patterns related to the genesis of the SPCZ in light of other well-documented topographically-generated zones. The study then explores the links between the theoretical models and observations of the SPCZ in several episodes. With this foundation, the dissertation then tests several hypotheses relating to the horizontal and vertical zone structure, topographic sensitivity on the zone structure, and boundary layer evolution of the zone through the use of high resolution nested grid numerical simulations. The SPCZ consists of windward and leeward

  3. Characterizing ammonia emissions from swine farms in eastern North Carolina: Reduction of emissions from water-holding structures at two candidate superior technologies for waste treatment

    NASA Astrophysics Data System (ADS)

    Aneja, Viney P.; Arya, S. Pal; Rumsey, Ian C.; Kim, D.-S.; Bajwa, K. S.; Williams, C. M.

    Program OPEN (Odor, Pathogens, and Emissions of Nitrogen) was an integrated study of the emissions of ammonia (NH 3), odor and odorants, and pathogens from potential environmentally superior technologies (ESTs) for swine facilities in eastern North Carolina. This paper, as part of program OPEN, focuses on quantifying emissions of NH 3 from water-holding structures at two of the best ESTs and compares them with the projected emissions from two conventional lagoon and spray technologies (LSTs). The evaluated ESTs are: (1) Super Soils at Goshen Ridge; and (2) Environmental Technologies at Red Hill. The water-holding structures for these two ESTs contained no conventional anaerobic lagoon. A dynamic flow-through chamber was used to measure NH 3 fluxes from the water-holding structures at both the ESTs and at the conventional LST farms. In order to compare the emissions from the water-holding structures at the ESTs with those from the lagoons at the conventional sites under similar conditions, a statistical-observational model for lagoon NH 3 emissions was used. A mass-balance approach was used to quantify the emissions. All emissions were normalized by nitrogen-excretion rates. The percentage reductions relative to the conventional lagoons were calculated for the two ESTs. Results showed substantial reductions in NH 3 emissions at both ESTs. Super Soils had reductions of 94.7% for the warm season and 99.0% for the cool season. Environmental Technologies had slightly larger reductions of 99.4% and 99.98% for the cool and warm season, respectively. As a result of such large reductions in ammonia emissions, both technologies meet the criteria to be classified as ESTs for ammonia emissions.

  4. Delineating ecological regions in marine systems: Integrating physical structure and community composition to inform spatial management in the eastern Bering Sea

    NASA Astrophysics Data System (ADS)

    Baker, Matthew R.; Hollowed, Anne B.

    2014-11-01

    Characterizing spatial structure and delineating meaningful spatial boundaries have useful applications to understanding regional dynamics in marine systems, and are integral to ecosystem approaches to fisheries management. Physical structure and drivers combine with biological responses and interactions to organize marine systems in unique ways at multiple scales. We apply multivariate statistical methods to define spatially coherent ecological units or ecoregions in the eastern Bering Sea. We also illustrate a practical approach to integrate data on species distribution, habitat structure and physical forcing mechanisms to distinguish areas with distinct biogeography as one means to define management units in large marine ecosystems. We use random forests to quantify the relative importance of habitat and environmental variables to the distribution of individual species, and to quantify shifts in multispecies assemblages or community composition along environmental gradients. Threshold shifts in community composition are used to identify regions with distinct physical and biological attributes, and to evaluate the relative importance of predictor variables to determining regional boundaries. Depth, bottom temperature and frontal boundaries were dominant factors delineating distinct biological communities in this system, with a latitudinal divide at approximately 60°N. Our results indicate that distinct climatic periods will shift habitat gradients and that dynamic physical variables such as temperature and stratification are important to understanding temporal stability of ecoregion boundaries. We note distinct distribution patterns among functional guilds and also evidence for resource partitioning among individual species within each guild. By integrating physical and biological data to determine spatial patterns in community composition, we partition ecosystems along ecologically significant gradients. This may provide a basis for defining spatial management

  5. Interspecific variation in resistance of two host tree species to spruce budworm

    NASA Astrophysics Data System (ADS)

    Fuentealba, Alvaro; Bauce, Éric

    2016-01-01

    Woody plants regularly sustain biomass losses to herbivorous insects. Consequently, they have developed various resistance mechanisms to cope with insect attack. However, these mechanisms of defense and how they are affected by resource availability are not well understood. The present study aimed at evaluating and comparing the natural resistance (antibiosis and tolerance) of balsam fir (Abies balsamea [L.] Mill.) and white spruce (Picea glauca [Moench) Voss] to spruce budworm, Choristoneura fumiferana (Clem.), and how drainage site quality as a component of resource availability affects the expression of resistance over time (6 years). Our results showed that there are differences in natural resistance between the two tree species to spruce budworm, but it was not significantly affected by drainage quality. Balsam fir exhibited higher foliar toxic secondary compounds concentrations than white spruce in all drainage classes, resulting in lower male pupal mass, survival and longer male developmental time. This, however, did not prevent spruce budworm from consuming more foliage in balsam fir than in white spruce. This response suggests that either natural levels of measured secondary compounds do not provide sufficient toxicity to reduce defoliation, or spruce budworm has developed compensatory mechanisms, which allow it to utilize food resources more efficiently or minimize the toxic effects that are produced by its host's defensive compounds. Larvae exhibited lower pupal mass and higher mortality in rapidly drained and subhygric sites. Drainage class also affected the amount of foliage destroyed but its impact varied over the years and was probably influenced by climatic variables. These results demonstrate the complexity of predicting the effect of resource availability on tree defenses, especially when other confounding environmental factors can affect tree resource allocation and utilization.

  6. The spruce shoot gall midges (Diptera: Cecidomyiidae): Piceacecis, a new genus for a non-native pest of Norway spruce from Europe and its native American relative

    Technology Transfer Automated Retrieval System (TEKTRAN)

    Dasineura abietiperda (Henschel), a European pest of Norway spruce, Picea abies (Pinaceae), is reported as new to North America. Damage symptoms are illustrated and an outline of its biology is given. A new genus, Piceacecis Gagné is described to include it and its North American relative, Phytophag...

  7. Structural controls on the spatial distribution and geochemical composition of volcanism in a continental rift zone; an example from Owens Valley, eastern California

    NASA Astrophysics Data System (ADS)

    Haproff, P. J.; Yin, A.

    2014-12-01

    Bimodal volcanism is common in continental rift zones. Structural controls to the emplacement and compositions of magmas, however, are not well understood. To address this issue, we examine the location, age, and geochemistry of active volcanic centers, and geometry and kinematics of rift-related faults across the active transtensional Owens Valley rift zone. Building on existing studies, we postulate that the spatial distribution and geochemical composition of volcanism are controlled by motion along rift-bounding fault systems. Along-strike variation in fault geometry and characteristics of active volcanism allow us to divide Owens Valley into three segments: southern, northern, and central. The southern segment of Owens Valley is a simple shear, asymmetric rift bounded to the west by the east-dipping Sierra Nevada frontal fault (SNFF). Active vents of Coso volcanic field are distributed along the eastern rift shoulder and characterized by the eruption of bimodal lavas. The SNFF within this segment is low-angle and penetrates through the lithosphere and into the ductile asthenosphere, allowing for mantle-derived magma to migrate across the weakest part of the fault zone beneath the eastern rift shoulder. Magma thermally weakens wall rocks and eventually stalls in the crust where the melt develops a greater felsic component prior to eruption. The northern segment of Owens Valley displays similar structural geometry, as the west-dipping White Mountains fault (WMF) is listric at depth and offsets the crust and mantle lithosphere, allowing for vertical transport of magma and reservoir emplacement within the crust. Bimodal lavas periodically erupted in the Long Valley Caldera region along the western rift shoulder. The central segment of Owens Valley is a pure shear, symmetric graben generated by motion along the SNFF and WMF. The subvertical, right-slip Owens Valley fault (OVF) strikes along the axis of the valley and penetrates through the lithosphere into the

  8. The crustal structure of the eastern Fennoscandian Shield and central part of the East-European platform based on seismic, regional geophysic and geological data

    NASA Astrophysics Data System (ADS)

    Mints, M. V.; Berzin, R. G.; Babayants, P. S.; Konilov, A. N.; Suleimanov, A. K.; Zamozhniaya, N. G.; Zlobin, V. L.

    2003-04-01

    The 1-EU and 4B CDP transects worked out during 1998-2002 years by "Spetsgeophyzica", together with previously developed CDP profiles, have crossed most of the main tectonic units of the eastern Fennoscandian Shield and central part of the East-European platform. They provide seismic images of the Early Precambrian crust and upper mantle from the surface to about 80 km depth (25 s). The Neoarchaean granite-greenstone complexes of the Karelia craton along the 4B profile form a series of the tectonic slices descending eastward, some of which can be traced to the Moho. The Palaeoproterozoic structures presented by two main types: (1) volcano-sedimentary (VS) and (2) granulite-gneiss (GN) belts. The Pechenga-Varzuga VS belt has been identified as overthrust-underthrust southward-dipping package. Tectonic slices formed by the Palaeoproterozoic VS belts alternating with slices of the Neoarchaean granite-gneisses form the imbricated crustal unit that extends along the eastern margin of the Neoarchaean Karelia craton. The slices dip steeply northeastward flattening and partially juxtaposing at 20 km depth at the 1-EU cross-section. This level, which can be understood as the surface of main detachment, ascends westward. An imbrication and related thickening of the crust was caused by displacement of crustal slices in western and southwestern directions because of the Palaeoproterozoic collision event. The Palaeoproterozoic Onega unit comprising VS assemblages originated in a setting of the rifted passive margin forms the northwestward displaced thrust nappe complex. It is considered initially belonging to the southern edge of the Svecofennian passive margin. The Lapland GN belt has been transected by the Polar and EGGI profiles. Both cross-sections demonstrated that it constitutes thick composite crustal-scale tectonic slice. According to geophysical data, the continuation of the Lapland GN belt beneath the platform cover of the East European Craton forms an extended arch

  9. Deep Crustal Magma Conduits, Diabase Internal Structure, and Coupled Hydrothermal Processes in Mesozoic Basins of Eastern North America

    NASA Astrophysics Data System (ADS)

    Ryan, M. P.; Ingerov, A.; Daniels, D. L.; Carr, P. M.; Elliott, G.; Fox, L.; Pierce, H. A.; Sutphin, D. M.

    2004-05-01

    Integrated geophysical, geological, and hydrological studies of the Culpeper (VA) and Gettysburg (PA) basins have shown fundamental new relationships among the internal structure of their plutonic rocks and have constrained the structural controls on paleo- and contemporary subsurface fluid flow within their ancient rift zone intrusive centers. Deep sub-basin igneous structure and the cross-sectional structure of the Culpeper basin. Magnetotelluric (MT) measurements along two basin-crossing transects have been inverted to reveal the basin cross-sectional structure and the structure of the primary conduit that transported tholeiitic magma through the crust during the Lower Jurassic. In cross-section, the basin deepens abruptly east of the Bull Run and Catoctin mountains, reaching a maximum depth of 2.5 to 2.75 km. The basin then shallows gradually towards the east. Basin rocks span the range 25 to 8,000 Ohm-m and range from fluid-saturated siltstones to unsaturated diabase, respectively. The crustal conduit for the Belmont diabase sheet has been resolved to 20 km depth. At that depth, it is characterized by a vertical region of relatively low resistivity (150 to 4,000 Ohm-m) embedded in relatively resistive (~10,000+ Ohm-m) crystalline Proterozoic rocks. At 4--5 km depth, the conduit width is ~2 km, flaring outward to a mean width of 6 km at 20 km depth. The conduit is inferred to be a laminate-like resistivity ``composite,'' with a central core resistivity of ~150 Ohm-m grading outwards through zones of 103, 2×103, and finally 4×103 Ohm-m near the outer margins. Based on the phase equilibria of ultramafic rocks and the petrology of diabase, the conduit is inferred to be olivine gabbro +/- dunite, and to be compositionally-zoned from core to margin. In general, the resistivity structure of the conduit is bilaterally-symmetric. Intra-basin igneous structure of the Gettysburg basin. The structure and geometry of the diabase sheets, lopoliths, and dikes of the

  10. Steam pretreatment of spruce forest residues: optimal conditions for biogas production and enzymatic hydrolysis.

    PubMed

    Janzon, Ron; Schütt, Fokko; Oldenburg, Saskia; Fischer, Elmar; Körner, Ina; Saake, Bodo

    2014-01-16

    Steam refining of non-debarked spruce forest residues was investigated as pretreatment for enzymatic hydrolysis as well as for biogas production. Pretreatment conditions were varied in the range of 190-220 °C, 5-10 min and 0-3.7% SO₂ according to a statistical design. For both applications highest product yields were predicted at 220 °C and 2.4% SO₂, whereas the reaction time had only a minor influence. The conformity of the model results allows the conclusion that enzymatic hydrolysis is a suitable test method to evaluate the degradability of lignocellulosic biomass in the biogas process. In control experiments under optimal conditions the results of the model were verified. The yield of total monomeric carbohydrates after enzymatic hydrolysis was equivalent to 55% of all theoretically available polysaccharides. The corresponding biogas yield from the pretreated wood amounted to 304 mL/gODM. Furthermore, furans produced under optimal process conditions showed no inhibitory effect on biogas production. It can be concluded that steam refining opens the structure of wood, thus improving the enzymatic hydrolysis of the polysaccharides to fermentable monomeric sugars and subsequently enabling a higher and faster production of biogas. Anaerobic fermentation of pretreated wood is a serious alternative to alcoholic fermentation especially when low quality wood grades and residues are used. Anaerobic digestion should be further investigated in order to diversify the biorefinery options for lignocellulosic materials.

  11. Universal fractionation of lignin–carbohydrate complexes (LCCs) from lignocellulosic biomass: an example using spruce wood

    PubMed Central

    Du, Xueyu; Gellerstedt, Goran; Li, Jiebing

    2014-01-01

    SUMMARY It is of both theoretical and practical importance to develop a universally applicable approach for the fractionation and sensitive lignin characterization of lignin–carbohydrate complexes (LCCs) from all types of lignocellulosic biomass, both natively and after various types of processing. In the present study, a previously reported fractionation approach that is applicable for eucalyptus (hardwood) and flax (non-wood) was further improved by introducing an additional step of barium hydroxide precipitation to isolate the mannan-enriched LCC (glucomannan-lignin, GML), in order to suit softwood species as well. Spruce wood was used as the softwood sample. As indicated by the recovery yield and composition analysis, all of the lignin was recovered in three LCC fractions: a glucan-enriched fraction (glucan-lignin, GL), a mannan-enriched fraction (GML) and a xylan-enriched fraction (xylan-lignin, XL). All of the LCCs had high molecular masses and were insoluble or barely soluble in a dioxane/water solution. Carbohydrate and lignin signals were observed in 1H NMR, 13C CP-MAS NMR and normal- or high-sensitivity 2D HSQC NMR analyses. The carbohydrate and lignin constituents in each LCC fraction are therefore believed to be chemically bonded rather than physically mixed with one another. The three LCC fractions were found to be distinctly different from each other in terms of their lignin structures, as revealed by highly sensitive analyses by thioacidolysis-GC, thioacidolysis-SEC and pyrolysis-GC. PMID:23332001

  12. Predicting adaptive phenotypes from multilocus genotypes in Sitka spruce (Picea sitchensis) using random forest.

    PubMed

    Holliday, Jason A; Wang, Tongli; Aitken, Sally

    2012-09-01

    Climate is the primary driver of the distribution of tree species worldwide, and the potential for adaptive evolution will be an important factor determining the response of forests to anthropogenic climate change. Although association mapping has the potential to improve our understanding of the genomic underpinnings of climatically relevant traits, the utility of adaptive polymorphisms uncovered by such studies would be greatly enhanced by the development of integrated models that account for the phenotypic effects of multiple single-nucleotide polymorphisms (SNPs) and their interactions simultaneously. We previously reported the results of association mapping in the widespread conifer Sitka spruce (Picea sitchensis). In the current study we used the recursive partitioning algorithm 'Random Forest' to identify optimized combinations of SNPs to predict adaptive phenotypes. After adjusting for population structure, we were able to explain 37% and 30% of the phenotypic variation, respectively, in two locally adaptive traits--autumn budset timing and cold hardiness. For each trait, the leading five SNPs captured much of the phenotypic variation. To determine the role of epistasis in shaping these phenotypes, we also used a novel approach to quantify the strength and direction of pairwise interactions between SNPs and found such interactions to be common. Our results demonstrate the power of Random Forest to identify subsets of markers that are most important to climatic adaptation, and suggest that interactions among these loci may be widespread.

  13. Formalized classification of moss litters in swampy spruce forests of intermontane depressions of Kuznetsk Alatau

    NASA Astrophysics Data System (ADS)

    Efremova, T. T.; Avrova, A. F.; Efremov, S. P.

    2016-09-01

    The approaches of multivariate statistics have been used for the numerical classification of morphogenetic types of moss litters in swampy spruce forests according to their physicochemical properties (the ash content, decomposition degree, bulk density, pH, mass, and thickness). Three clusters of moss litters— peat, peaty, and high-ash peaty—have been specified. The functions of classification for identification of new objects have been calculated and evaluated. The degree of decomposition and the ash content are the main classification parameters of litters, though all other characteristics are also statistically significant. The final prediction accuracy of the assignment of a litter to a particular cluster is 86%. Two leading factors participating in the clustering of litters have been determined. The first factor—the degree of transformation of plant remains (quality)—specifies 49% of the total variance, and the second factor—the accumulation rate (quantity)— specifies 26% of the total variance. The morphogenetic structure and physicochemical properties of the clusters of moss litters are characterized.

  14. Vertical and horizontal water redistribution in Norway spruce (Picea abies) roots in the Moravian Upland.

    PubMed

    Nadezhdina, Nadezhda; Cermák, Jan; Gaspárek, Jan; Nadezhdin, Valeriy; Prax, Alois

    2006-10-01

    Hydraulic redistribution (HR) by roots of large Norway spruce (Picea abies (L.) Karst.) trees was investigated by means of sap flow measurements made with the heat field deformation method. Irrigation was applied to a limited portion of the root system to steepen gradients of water potential in the soil and thus enhance rates of HR. On completion of the sap flow measurements, and to aid in their interpretation, the structure of the root system of seven of the investigated trees was exposed to a depth of 30 cm with a supersonic air-stream (air-spade). Before irrigation, vertical redistribution of water was observed in large coarse roots and some adjacent small lateral roots. Immediately after localized irrigation, horizontal redistribution of water from watered roots to dry roots via the stem base was demonstrated. The amount of horizontal distribution depended on the position of the receiving roots relative to the watered roots and the absorbing area of the watered root. No redistribution from watered roots via dry soil to roots of neighboring trees was detected. Responses of sap flow to localized irrigation were more pronounced in small lateral roots than in large branching roots where release and uptake of water are integrated. Sap flow measurements with multi-point sensors along radii in large lateral roots demonstrated water extraction from different soil horizons. We conclude that synchronous measurements of sap flow in both small and large lateral roots are needed to study water absorption and transport in tree root systems.

  15. Geology of the Spruce Pine District, Avery, Mitchell, and Yancy Counties, North Carolina

    USGS Publications Warehouse

    Brobst, Donald Albert

    1962-01-01

    The Spruce Pine pegmatite district, a northeastward-trending belt 25 miles long and 10 miles wide, lies in parts of Avery, Mitchell, and Yancey Counties in the Blue Ridge Province of western North Carolina. The most abundant rocks in the district are interlayered mica and amphibole gneisses and schists, all of which are believed to be of Precambrian age. These rocks are cut by small bodies of dunite and associated rocks of Precambrian (?) age, large bodies of alaskite and associated pegmatite of early Paleozoic age, and basaltic and diabasic dikes and sills of Triassic (?) age. The rocks of the district have been weathered to saprolite that is locally 50 feet thick. The major structure in the area is a southwestward-plunging asymmetrical synclinorium that has its steeper limb on the northwest side. Feldspar, muscovite as sheet and scrap (ground) mica, and kaolin from the alaskite and associated pegmatite account for over 90 percent of the total mineral production of the district. Amounts of other pegmatite minerals, including quartz, beryl, columbite-tantalite, rare-earth and uranium minerals are an extremely small part of the mineral resources. Actual or potential products from other rocks are olivine, vermiculite, asbestos, talc, chromium and nickel, soapstone, mica schist, garnet, kyanite, dolomite marble, and construction materials.

  16. Fresh-wood bending: linking the mechanical and growth properties of a Norway spruce stem.

    PubMed

    Lundström, Tor; Heiz, Urs; Stoffel, Markus; Stöckli, Veronika

    2007-09-01

    To provide data and methods for analyzing stem mechanics, we investigated bending, density and growth characteristics of 207 specimens of fresh wood from different heights and radial positions of the stem of one mature Norway spruce (Picea abies L. Karst.) tree. From the shape of each stress-strain curve, which was calculated from bending tests that accounted for shear deformation, we determined the modulus of elasticity (MOE), the modulus of rupture (MOR), the completeness of the material, an idealized stress-strain curve and the work involved in bending. In general, all mechanical properties increased with distance from the pith, with values in the ranges of 5.7-18 GPa for MOE, 23-90 MPa for MOR and 370-630 and 430-1100 kg m(-3) for dry and fresh wood densities, respectively. The first three properties generally decreased with stem height, whereas fresh wood density increased. Multiple regression equations were calculated, relating MOR, MOE and dry wood density to growth properties. We applied these equations to the growth of the entire stem and considered the annual rings as superimposed cylindrical shells, resulting in stem-section values of MOE, MOR and dry and fresh densities as a function of stem height and cambial age. The standing tree exhibits an inner stem structure that is well designed for bending, especially at a mature stage. PMID:17545123

  17. Three-dimensional crustal structure of a craton rim: Preliminary results from passive seismic imaging of the eastern Albany-Fraser Orogen, Western Australia

    NASA Astrophysics Data System (ADS)

    Sippl, Christian; Tkalčić, Hrvoje; Kennett, Brian L. N.; Spaggiari, Catherine V.; Gessner, Klaus

    2014-05-01

    Western Australia consists of two Archaean cratons (Yilgarn and Pilbara) and a number of Proterozoic orogens surrounding them that attest to past continental collisions. While the former feature seismically fast crust of average thickness (around 35 km) and a usually well defined Moho overlying a thick mantle lithospheric keel, the latter have been significantly less well studied and appear to be less uniform in terms of their crustal architecture. Thicker crust and a more fuzzy Moho are two common characteristics of these belts. The Albany-Fraser orogen, situated at the south-eastern margin of the Yilgarn craton, has been interpreted as an old suture zone from the collision of the West Australian craton (Yilgarn and Pilbara already welded together) with the Mawson craton (southern Australia and part of Antarctica today). Newer evidence, however, might point at an original rift or backarc setting of the units. It is a complex amalgam of different structures that vary significantly along its strike, featuring heavily reworked parts of the outermost Yilgarn craton as well as younger units accreted or intruded significantly later. Two major deformation stages at 1345-1260 Ma and 1214-1140 Ma have been deduced for these, the first of which has been associated with the aforementioned collision/backarc rifting itself, while the second phase is commonly interpreted as intracratonic reworking due to a major thermal event. No large-scale tectonic overprint has occurred in the region since the second deformation phase, which means that the originally emplaced units have been unusually well preserved until the present day. However, surface outcrops of rocks are very rare, so that most knowledge about extent and geometric configuration of different rock suites comes from the interpretation of magnetic and gravity data. The eastern end of the Albany-Fraser orogen, in all likelihood corresponding to the Mawson craton's westernmost edge, is hidden beneath the limestones of the

  18. Expression of the β-glucosidase gene Pgβglu-1 underpins natural resistance of white spruce against spruce budworm.

    PubMed

    Mageroy, Melissa H; Parent, Geneviève; Germanos, Gaby; Giguère, Isabelle; Delvas, Nathalie; Maaroufi, Halim; Bauce, Éric; Bohlmann, Joerg; Mackay, John J

    2015-01-01

    Periodic outbreaks of spruce budworm (SBW) affect large areas of ecologically and economically important conifer forests in North America, causing tree mortality and reduced forest productivity. Host resistance against SBW has been linked to growth phenology and the chemical composition of foliage, but the underlying molecular mechanisms and population variation are largely unknown. Using a genomics approach, we discovered a β-glucosidase gene, Pgβglu-1, whose expression levels and function underpin natural resistance to SBW in mature white spruce (Picea glauca) trees. In phenotypically resistant trees, Pgβglu-1 transcripts were up to 1000 times more abundant than in non-resistant trees and were highly enriched in foliage. The encoded PgβGLU-1 enzyme catalysed the cleavage of acetophenone sugar conjugates to release the aglycons piceol and pungenol. These aglycons were previously shown to be active against SBW. Levels of Pgβglu-1 transcripts and biologically active acetophenone aglycons were substantially different between resistant and non-resistant trees over time, were positively correlated with each other and were highly variable in a natural white spruce population. These results suggest that expression of Pgβglu-1 and accumulation of acetophenone aglycons is a constitutive defence mechanism in white spruce. The progeny of resistant trees had higher Pgβglu-1 gene expression than non-resistant progeny, indicating that the trait is heritable. With reported increases in the intensity of SBW outbreaks, influenced by climate, variation of Pgβglu-1 transcript expression, PgβGLU-1 enzyme activity and acetophenone accumulation may serve as resistance markers to better predict impacts of SBW in both managed and wild spruce populations.

  19. Expression of the β-glucosidase gene Pgβglu-1 underpins natural resistance of white spruce against spruce budworm

    PubMed Central

    Mageroy, Melissa H; Parent, Geneviève; Germanos, Gaby; Giguère, Isabelle; Delvas, Nathalie; Maaroufi, Halim; Bauce, Éric; Bohlmann, Joerg; Mackay, John J

    2015-01-01

    Periodic outbreaks of spruce budworm (SBW) affect large areas of ecologically and economically important conifer forests in North America, causing tree mortality and reduced forest productivity. Host resistance against SBW has been linked to growth phenology and the chemical composition of foliage, but the underlying molecular mechanisms and population variation are largely unknown. Using a genomics approach, we discovered a β-glucosidase gene, Pgβglu-1, whose expression levels and function underpin natural resistance to SBW in mature white spruce (Picea glauca) trees. In phenotypically resistant trees, Pgβglu-1 transcripts were up to 1000 times more abundant than in non-resistant trees and were highly enriched in foliage. The encoded PgβGLU-1 enzyme catalysed the cleavage of acetophenone sugar conjugates to release the aglycons piceol and pungenol. These aglycons were previously shown to be active against SBW. Levels of Pgβglu-1 transcripts and biologically active acetophenone aglycons were substantially different between resistant and non-resistant trees over time, were positively correlated with each other and were highly variable in a natural white spruce population. These results suggest that expression of Pgβglu-1 and accumulation of acetophenone aglycons is a constitutive defence mechanism in white spruce. The progeny of resistant trees had higher Pgβglu-1 gene expression than non-resistant progeny, indicating that the trait is heritable. With reported increases in the intensity of SBW outbreaks, influenced by climate, variation of Pgβglu-1 transcript expression, PgβGLU-1 enzyme activity and acetophenone accumulation may serve as resistance markers to better predict impacts of SBW in both managed and wild spruce populations. PMID:25302566

  20. Xylogenesis in black spruce: does soil temperature matter?

    PubMed

    Lupi, Carlo; Morin, Hubert; Deslauriers, Annie; Rossi, Sergio

    2012-01-01

    In boreal ecosystems, an increase in soil temperature can stimulate plant growth. However, cambium phenology in trees was better explained by air than soil temperature, which suggested that soil temperature is not the main limiting factor affecting xylogenesis. Since soil temperature and snowmelt are correlated to air temperature, the ques