Sample records for high-elevation subalpine forest
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Larson, Gary L.; McIntire, C.D.; Truitt, R.E.; Liss, W.J.; Hoffman, Robert L.; Deimling, E.; Lomnicky, G.A.
1998-01-01
Phytoplankton assemblages in high-elevation lakes of North Cascades National Park Service Complex were studied during the open-water period in 1989. Collectively, 93 taxa were identified in 55 samples from 51 lakes. Based on cell densities, cyanobacteria had the highest relative abundance (36.7 %), followed by chlorophytes (29.8 %), and chrysophytes (19.6 %). Aphanocapsa delicatissima had the highest proportional abundance (14.0 %). Only 15.1 % percent of the taxa occurred in more than 20 samples. Phytoplankton cell densities increased following a gradient of increasing lake-water temperature, alkalinity, and concentration of total Kjeldahl-N with decreasing lake elevation. Chrysophytes and cyanobacteria were quantitatively (relative abundance) the most important taxa in alpine and subalpine lakes, whereas cyanobacteria had the highest relative abundances in high-forest and low-forest lakes. Chlorophytes had their highest relative abundance in high-forest lakes. Although low in relative abundance, diatoms and dinoflagellates were most abundant in alpine lakes. An ordination by correspondence analysis indicated that most alpine, subalpine, and high-forest lakes had similar floras. Although a few subalpine lakes exhibited deviations from this pattern, the main differences in phytoplankton composition were found in a group of low-forest and high-forest lakes. Canonical correspondence analysis (CCA) provided evidence that the distribution of samples and taxa in ordination space was correlated with a gradient of decreasing lake elevation and increasing water temperature, alkalinity, and concentration of nitrogen. When CCA was used to examine relationships among phytoplankton taxa and vegetation zones, a continuous distribution of taxa was found from the low-forest zone to the subalpine zone, with a large number of taxa occurring primarily in the subalpine and high-forest zones. Three phytoplankton taxa occurred primarily in alpine lakes, whereas five taxa co-occurred in alpine, subalpine, and high forest zones. Collectively, lake elevation and associated changes in water quality and concentrations of nutrients, especially nitrogen, appeared to be the primary physical and chemical factors influencing the taxonomic structures of phytoplankton assemblages.
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Carbon storage in subalpine forests and meadows of the Olympic Mountains, Washington
DOE Office of Scientific and Technical Information (OSTI.GOV)
Prichard, S.J.; Peterson, D.L.
1995-06-01
We investigated carbon storage in high elevation ecosystems of the Olympic Mountains. A sharp precipitation gradient created by the Olympic mountain range allows for comparison of carbon storage in different climatic regimes and vegetation types. Carbon in soils, vegetation, and woody debris was examined in subalpine forests and meadows of the northeast (dry) and southwest (wet) Olympics. Soil carbon storage in high elevation sites appears to be considerably greater than most low elevation forests. Above-ground carbon storage is generally greater in southwest sites. Meadow soils contained high carbon concentrations in upper horizons, while forests also stored a substantial amount ofmore » carbon in lower horizons. Information gained from this study will provide a better understanding of soil-vegetation relationships in subalpine ecosystems, especially with respect to potential climatic change impacts.« less
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Kueppers, Lara M.; Conlisk, Erin; Castanha, Cristina; Moyes, Andrew B.; Germino, Matthew; de Valpine, Perry; Torn, Margaret S.; Mitton, Jeffry B.
2017-01-01
Climate niche models project that subalpine forest ranges will extend upslope with climate warming. These projections assume that the climate suitable for adult trees will be adequate for forest regeneration, ignoring climate requirements for seedling recruitment, a potential demographic bottleneck. Moreover, local genetic adaptation is expected to facilitate range expansion, with tree populations at the upper forest edge providing the seed best adapted to the alpine. Here, we test these expectations using a novel combination of common gardens, seeded with two widely distributed subalpine conifers, and climate manipulations replicated at three elevations. Infrared heaters raised temperatures in heated plots, but raised temperatures more in the forest than at or above treeline because strong winds at high elevation reduced heating efficiency. Watering increased season-average soil moisture similarly across sites. Contrary to expectations, warming reduced Engelmann spruce recruitment at and above treeline, as well as in the forest. Warming reduced limber pine first-year recruitment in the forest, but had no net effect on fourth-year recruitment at any site. Watering during the snow-free season alleviated some negative effects of warming, indicating that warming exacerbated water limitations. Contrary to expectations of local adaptation, low-elevation seeds of both species initially recruited more strongly than high-elevation seeds across the elevation gradient, although the low-provenance advantage diminished by the fourth year for Engelmann spruce, likely due to small sample sizes. High- and low-elevation provenances responded similarly to warming across sites for Engelmann spruce, but differently for limber pine. In the context of increasing tree mortality, lower recruitment at all elevations with warming, combined with lower quality, high-provenance seed being most available for colonizing the alpine, portends range contraction for Engelmann spruce. The lower sensitivity of limber pine to warming indicates a potential for this species to become more important in subalpine forest communities in the coming centuries.
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Kueppers, Lara M; Conlisk, Erin; Castanha, Cristina; Moyes, Andrew B; Germino, Matthew J; de Valpine, Perry; Torn, Margaret S; Mitton, Jeffry B
2017-06-01
Climate niche models project that subalpine forest ranges will extend upslope with climate warming. These projections assume that the climate suitable for adult trees will be adequate for forest regeneration, ignoring climate requirements for seedling recruitment, a potential demographic bottleneck. Moreover, local genetic adaptation is expected to facilitate range expansion, with tree populations at the upper forest edge providing the seed best adapted to the alpine. Here, we test these expectations using a novel combination of common gardens, seeded with two widely distributed subalpine conifers, and climate manipulations replicated at three elevations. Infrared heaters raised temperatures in heated plots, but raised temperatures more in the forest than at or above treeline because strong winds at high elevation reduced heating efficiency. Watering increased season-average soil moisture similarly across sites. Contrary to expectations, warming reduced Engelmann spruce recruitment at and above treeline, as well as in the forest. Warming reduced limber pine first-year recruitment in the forest, but had no net effect on fourth-year recruitment at any site. Watering during the snow-free season alleviated some negative effects of warming, indicating that warming exacerbated water limitations. Contrary to expectations of local adaptation, low-elevation seeds of both species initially recruited more strongly than high-elevation seeds across the elevation gradient, although the low-provenance advantage diminished by the fourth year for Engelmann spruce, likely due to small sample sizes. High- and low-elevation provenances responded similarly to warming across sites for Engelmann spruce, but differently for limber pine. In the context of increasing tree mortality, lower recruitment at all elevations with warming, combined with lower quality, high-provenance seed being most available for colonizing the alpine, portends range contraction for Engelmann spruce. The lower sensitivity of limber pine to warming indicates a potential for this species to become more important in subalpine forest communities in the coming centuries. © 2016 John Wiley & Sons Ltd.
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DOE Office of Scientific and Technical Information (OSTI.GOV)
Kueppers, Lara M.; Conlisk, Erin; Castanha, Cristina
Climate niche models project that subalpine forest ranges will extend upslope with climate warming. These projections assume that the climate suitable for adult trees will be adequate for forest regeneration, ignoring climate requirements for seedling recruitment, a potential demographic bottleneck. Moreover, local genetic adaptation is expected to facilitate range expansion, with tree populations at the upper forest edge providing the seed best adapted to the alpine. Here, we test these expectations using a novel combination of common gardens, seeded with two widely distributed subalpine conifers, and climate manipulations replicated at three elevations. Infrared heaters raised temperatures in heated plots, butmore » raised temperatures more in the forest than at or above treeline because strong winds at high elevation reduced heating efficiency. Watering increased season-average soil moisture similarly across sites. Contrary to expectations, warming reduced Engelmann spruce recruitment at and above treeline, as well as in the forest. Warming reduced limber pine first-year recruitment in the forest, but had no net effect on fourth-year recruitment at any site. Watering during the snow-free season alleviated some negative effects of warming, indicating that warming exacerbated water limitations. Contrary to expectations of local adaptation, low-elevation seeds of both species initially recruited more strongly than high-elevation seeds across the elevation gradient, although the low-provenance advantage diminished by the fourth year for Engelmann spruce, likely due to small sample sizes. High- and low-elevation provenances responded similarly to warming across sites for Engelmann spruce, but differently for limber pine. In the context of increasing tree mortality, lower recruitment at all elevations with warming, combined with lower quality, high-provenance seed being most available for colonizing the alpine, portends range contraction for Engelmann spruce. The lower sensitivity of limber pine to warming indicates a potential for this species to become more important in subalpine forest communities in the coming centuries.« less
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Kueppers, Lara M.; Conlisk, Erin; Castanha, Cristina; ...
2016-12-15
Climate niche models project that subalpine forest ranges will extend upslope with climate warming. These projections assume that the climate suitable for adult trees will be adequate for forest regeneration, ignoring climate requirements for seedling recruitment, a potential demographic bottleneck. Moreover, local genetic adaptation is expected to facilitate range expansion, with tree populations at the upper forest edge providing the seed best adapted to the alpine. Here, we test these expectations using a novel combination of common gardens, seeded with two widely distributed subalpine conifers, and climate manipulations replicated at three elevations. Infrared heaters raised temperatures in heated plots, butmore » raised temperatures more in the forest than at or above treeline because strong winds at high elevation reduced heating efficiency. Watering increased season-average soil moisture similarly across sites. Contrary to expectations, warming reduced Engelmann spruce recruitment at and above treeline, as well as in the forest. Warming reduced limber pine first-year recruitment in the forest, but had no net effect on fourth-year recruitment at any site. Watering during the snow-free season alleviated some negative effects of warming, indicating that warming exacerbated water limitations. Contrary to expectations of local adaptation, low-elevation seeds of both species initially recruited more strongly than high-elevation seeds across the elevation gradient, although the low-provenance advantage diminished by the fourth year for Engelmann spruce, likely due to small sample sizes. High- and low-elevation provenances responded similarly to warming across sites for Engelmann spruce, but differently for limber pine. In the context of increasing tree mortality, lower recruitment at all elevations with warming, combined with lower quality, high-provenance seed being most available for colonizing the alpine, portends range contraction for Engelmann spruce. The lower sensitivity of limber pine to warming indicates a potential for this species to become more important in subalpine forest communities in the coming centuries.« less
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Anderson, R. Scott; Allen, Craig D.; Toney, J.L.; Jass, R.B.; Bair, A.N.
2008-01-01
Our understanding of the present forest structure of western North America hinges on our ability to determine antecedent forest conditions. Sedimentary records from lakes and bogs in the southern Rocky Mountains of Colorado and New Mexico provide information on the relationships between climate and vegetation change, and fire history since deglaciation. We present a new pollen record from Hunters Lake (Colorado) as an example of a high-elevation vegetation history from the southern Rockies. We then present a series of six sedimentary records from ???2600 to 3500-m elevation, including sites presently at the alpine?subalpine boundary, within the Picea engelmannii?Abies lasiocarpa forest and within the mixed conifer forest, to determine the history of fire in high-elevation forests there. High Artemisia and low but increasing percentages of Picea and Pinus suggest vegetation prior to 13 500 calendar years before present (cal yr BP) was tundra or steppe, with open spruce woodland to ???11 900 cal yr BP. Subalpine forest (Picea engelmannii, Abies lasiocarpa) existed around the lake for the remainder of the Holocene. At lower elevations, Pinus ponderosa and/or contorta expanded 11 900 to 10 200 cal yr BP; mixed conifer forest expanded ???8600 to 4700 cal yr BP; and Pinus edulis expanded after ???4700 cal yr BP. Sediments from lake sites near the alpine?subalpine transition contained five times less charcoal than those entirely within subalpine forests, and 40 times less than bog sites within mixed conifer forest. Higher fire episode frequencies occurred between ???12 000 and 9000 cal yr BP (associated with the initiation or expansion of south-west monsoon and abundant lightning, and significant biomass during vegetation turnover) and at ???2000?1000 cal yr BP (related to periodic droughts during the long-term trend towards wetter conditions and greater biomass). Fire episode frequencies for subalpine?alpine transition and subalpine sites were on average 5 to 10 fire events/1000 years over the Holocene, corresponding to one fire event every ???100 to 200 years. (5) Our Holocene-length sedimentary charcoal records provide additional evidence for the anomalous nature of the 20th-century fire regime, where fires were largely suppressed as a national policy. ?? IAWF 2008.
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Ozone and modeled stomatal conductance at a high elevation subalpine site in southeastern Wyoming
Robert C. Musselman; Karl F. Zeller; Nedialko T. Nikolov
1998-01-01
Ozone concentrations have been monitored at the Glacier Lakes Ecosystem Experiment Site (GLEES) in the Snowy Range of the Medicine Bow Mountains 55 km west of Laramie, Wyoming, USA. The site is located at 3,186 m elevation in a large subalpine meadow of a mature subalpine forest near timberline. Continuous ozone and meteorological monitoring are a part of the GLEES...
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Ashton, L A; Nakamura, A; Burwell, C J; Tang, Y; Cao, M; Whitaker, T; Sun, Z; Huang, H; Kitching, R L
2016-05-23
South-western China is widely acknowledged as a biodiversity 'hotspot': there are high levels of diversity and endemism, and many environments are under significant anthropogenic threats not least climate warming. Here, we explore diversity and compare response patterns of moth assemblages among three elevational gradients established within different climatic bioregions - tropical rain forest, sub-tropical evergreen broad-leaved forest and sub-alpine coniferous forest in Yunnan Province, China. We hypothesised that tropical assemblages would be more elevationally stratified than temperate assemblages, and tropical species would be more elevationally restricted than those in the temperate zone. Contrary to our hypothesis, the moth fauna was more sensitive to elevational differences within the temperate transect, followed by sub-tropical and tropical transects. Moths in the cooler and more seasonal temperate sub-alpine gradient showed stronger elevation-decay beta diversity patterns, and more species were restricted to particular elevational ranges. Our study suggests that moth assemblages are under threat from future climate change and sub-alpine rather than tropical faunas may be the most sensitive to climate change. These results improve our understanding of China's biodiversity and can be used to monitor future changes to herbivore assemblages in a 'hotspot' of biodiversity.
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Ashton, L. A.; Nakamura, A.; Burwell, C. J.; Tang, Y.; Cao, M.; Whitaker, T.; Sun, Z.; Huang, H.; Kitching, R. L.
2016-01-01
South-western China is widely acknowledged as a biodiversity ‘hotspot’: there are high levels of diversity and endemism, and many environments are under significant anthropogenic threats not least climate warming. Here, we explore diversity and compare response patterns of moth assemblages among three elevational gradients established within different climatic bioregions - tropical rain forest, sub-tropical evergreen broad-leaved forest and sub-alpine coniferous forest in Yunnan Province, China. We hypothesised that tropical assemblages would be more elevationally stratified than temperate assemblages, and tropical species would be more elevationally restricted than those in the temperate zone. Contrary to our hypothesis, the moth fauna was more sensitive to elevational differences within the temperate transect, followed by sub-tropical and tropical transects. Moths in the cooler and more seasonal temperate sub-alpine gradient showed stronger elevation-decay beta diversity patterns, and more species were restricted to particular elevational ranges. Our study suggests that moth assemblages are under threat from future climate change and sub-alpine rather than tropical faunas may be the most sensitive to climate change. These results improve our understanding of China’s biodiversity and can be used to monitor future changes to herbivore assemblages in a ‘hotspot’ of biodiversity. PMID:27211989
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NASA Astrophysics Data System (ADS)
Ashton, L. A.; Nakamura, A.; Burwell, C. J.; Tang, Y.; Cao, M.; Whitaker, T.; Sun, Z.; Huang, H.; Kitching, R. L.
2016-05-01
South-western China is widely acknowledged as a biodiversity ‘hotspot’: there are high levels of diversity and endemism, and many environments are under significant anthropogenic threats not least climate warming. Here, we explore diversity and compare response patterns of moth assemblages among three elevational gradients established within different climatic bioregions - tropical rain forest, sub-tropical evergreen broad-leaved forest and sub-alpine coniferous forest in Yunnan Province, China. We hypothesised that tropical assemblages would be more elevationally stratified than temperate assemblages, and tropical species would be more elevationally restricted than those in the temperate zone. Contrary to our hypothesis, the moth fauna was more sensitive to elevational differences within the temperate transect, followed by sub-tropical and tropical transects. Moths in the cooler and more seasonal temperate sub-alpine gradient showed stronger elevation-decay beta diversity patterns, and more species were restricted to particular elevational ranges. Our study suggests that moth assemblages are under threat from future climate change and sub-alpine rather than tropical faunas may be the most sensitive to climate change. These results improve our understanding of China’s biodiversity and can be used to monitor future changes to herbivore assemblages in a ‘hotspot’ of biodiversity.
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R. C. Musselman; W. J. Massman; J. M. Frank; J. L. Korfmacher
2005-01-01
Carbon dioxide (CO2) concentration under snow was examined through two winter seasons at a 3100 m elevation subalpine site in the Snowy Range of Wyoming. CO2 was monitored every half hour at the soil/snow interface, and at about 25 cm soil depth the second year, in a meadow and in an adjacent forest. CO2 under snow in the meadow was significantly higher than that in...
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Conlisk, Erin; Castanha, Cristina; Germino, Matthew J.; Veblen, Thomas T; Smith, Jeremy M.; Kueppers, Lara M.
2017-01-01
Species distribution shifts in response to climate change require that recruitment increase beyond current range boundaries. For trees with long life spans, the importance of climate-sensitive seedling establishment to the pace of range shifts has not been demonstrated quantitatively.Using spatially explicit, stochastic population models combined with data from long-term forest surveys, we explored whether the climate-sensitivity of recruitment observed in climate manipulation experiments was sufficient to alter populations and elevation ranges of two widely distributed, high-elevation North American conifers.Empirically observed, warming-driven declines in recruitment led to rapid modelled population declines at the low-elevation, ‘warm edge’ of subalpine forest and slow emergence of populations beyond the high-elevation, ‘cool edge’. Because population declines in the forest occurred much faster than population emergence in the alpine, we observed range contraction for both species. For Engelmann spruce, this contraction was permanent over the modelled time horizon, even in the presence of increased moisture. For limber pine, lower sensitivity to warming may facilitate persistence at low elevations – especially in the presence of increased moisture – and rapid establishment above tree line, and, ultimately, expansion into the alpine.Synthesis. Assuming 21st century warming and no additional moisture, population dynamics in high-elevation forests led to transient range contractions for limber pine and potentially permanent range contractions for Engelmann spruce. Thus, limitations to seedling recruitment with warming can constrain the pace of subalpine tree range shifts.
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Conlisk, Erin; Castanha, Cristina; Germino, Matthew J.; ...
2017-02-08
Species distribution shifts in response to climate change require that recruitment increase beyond current range boundaries. For trees with long life spans, the importance of climate-sensitive seedling establishment to the pace of range shifts has not been demonstrated quantitatively. Using spatially explicit, stochastic population models combined with data from long-term forest surveys, we explored whether the climate-sensitivity of recruitment observed in climate manipulation experiments was sufficient to alter populations and elevation ranges of two widely distributed, high-elevation North American conifers. Empirically observed, warming-driven declines in recruitment led to rapid modelled population declines at the low-elevation, ‘warm edge’ of subalpine forestmore » and slow emergence of populations beyond the high-elevation, ‘cool edge’. Because population declines in the forest occurred much faster than population emergence in the alpine, we observed range contraction for both species. For Engelmann spruce, this contraction was permanent over the modelled time horizon, even in the presence of increased moisture. For limber pine, lower sensitivity to warming may facilitate persistence at low elevations – especially in the presence of increased moisture – and rapid establishment above tree line, and, ultimately, expansion into the alpine. Synthesis. Assuming 21st century warming and no additional moisture, population dynamics in high-elevation forests led to transient range contractions for limber pine and potentially permanent range contractions for Engelmann spruce. Thus, limitations to seedling recruitment with warming can constrain the pace of subalpine tree range shifts.« less
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DOE Office of Scientific and Technical Information (OSTI.GOV)
Conlisk, Erin; Castanha, Cristina; Germino, Matthew J.
Species distribution shifts in response to climate change require that recruitment increase beyond current range boundaries. For trees with long life spans, the importance of climate-sensitive seedling establishment to the pace of range shifts has not been demonstrated quantitatively. Using spatially explicit, stochastic population models combined with data from long-term forest surveys, we explored whether the climate-sensitivity of recruitment observed in climate manipulation experiments was sufficient to alter populations and elevation ranges of two widely distributed, high-elevation North American conifers. Empirically observed, warming-driven declines in recruitment led to rapid modelled population declines at the low-elevation, ‘warm edge’ of subalpine forestmore » and slow emergence of populations beyond the high-elevation, ‘cool edge’. Because population declines in the forest occurred much faster than population emergence in the alpine, we observed range contraction for both species. For Engelmann spruce, this contraction was permanent over the modelled time horizon, even in the presence of increased moisture. For limber pine, lower sensitivity to warming may facilitate persistence at low elevations – especially in the presence of increased moisture – and rapid establishment above tree line, and, ultimately, expansion into the alpine. Synthesis. Assuming 21st century warming and no additional moisture, population dynamics in high-elevation forests led to transient range contractions for limber pine and potentially permanent range contractions for Engelmann spruce. Thus, limitations to seedling recruitment with warming can constrain the pace of subalpine tree range shifts.« less
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Factors influencing avian communities in high-elevation southern Allegheny mountain forests
Harry A. Kahler; James T. Anderson
2010-01-01
Myriad factors may influence bird community characteristics among subalpine, central, and northern hardwood forest cover types of the southern Allegheny Mountains. Differences in forest cover types may result from natural characteristics, such as tree species composition, topography, or elevation, as well as from past influences, such as poor logging practices. Our...
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C.I. Millar; P.W. Rundel
2016-01-01
The subalpine forests of California comprise the highest elevation ecosystems that are dominated by upright trees. They are defined as a zone influenced primarily by abiotic controls, including persistent snowpack, desiccating winds, acute and chronic extreme temperatures, soil moisture and evapotranspirative stresses, and short growing seasons. Bounded at the...
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DOE Office of Scientific and Technical Information (OSTI.GOV)
Kueppers, Lara; Faist, Akasha; Ferrenberg, Scott
Accurately predicting upslope shifts in subalpine tree ranges with warming requires understanding how future forest populations will be affected by climate change, as these are the seed sources for new tree line and alpine populations. Early life history stages are particularly sensitive to climate and are also influenced by genetic variation among populations. Here, we tested the climate sensitivity of germination and initial development for two widely distributed subalpine conifers, using controlled-environment growth chambers with one temperature regime from subalpine forest in the Colorado Rocky Mountains and one 5 °C warmer, and two soil moisture levels. We also tracked germinationmore » rate and timing, rate of seedling development, and seedling morphology for two seed provenances separated by ~300 m elevation. Warming advanced germination timing and initial seedling development by a total of ~2 weeks, advances comparable to mean differences between provenances. Advances were similar for both provenances and species; however, warming reduced the overall germination rate, as did low soil moisture, only for Picea engelmannii. A three-year field warming and watering experiment planted with the same species and provenances yielded responses qualitatively consistent with the lab trials. Altogether these experiments indicate that in a warmer, drier climate, P. engelmannii germination, and thus regeneration, could decline, which could lead to declining subalpine forest populations, while Pinus flexilis forest populations could remain robust as a seed source for upslope range shifts.« less
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Kueppers, Lara; Faist, Akasha; Ferrenberg, Scott; ...
2017-11-11
Accurately predicting upslope shifts in subalpine tree ranges with warming requires understanding how future forest populations will be affected by climate change, as these are the seed sources for new tree line and alpine populations. Early life history stages are particularly sensitive to climate and are also influenced by genetic variation among populations. Here, we tested the climate sensitivity of germination and initial development for two widely distributed subalpine conifers, using controlled-environment growth chambers with one temperature regime from subalpine forest in the Colorado Rocky Mountains and one 5 °C warmer, and two soil moisture levels. We also tracked germinationmore » rate and timing, rate of seedling development, and seedling morphology for two seed provenances separated by ~300 m elevation. Warming advanced germination timing and initial seedling development by a total of ~2 weeks, advances comparable to mean differences between provenances. Advances were similar for both provenances and species; however, warming reduced the overall germination rate, as did low soil moisture, only for Picea engelmannii. A three-year field warming and watering experiment planted with the same species and provenances yielded responses qualitatively consistent with the lab trials. Altogether these experiments indicate that in a warmer, drier climate, P. engelmannii germination, and thus regeneration, could decline, which could lead to declining subalpine forest populations, while Pinus flexilis forest populations could remain robust as a seed source for upslope range shifts.« less
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DOE Office of Scientific and Technical Information (OSTI.GOV)
Wang, Wenzhi; Jia, Min; Wang, Genxu
Tree radial growth is expected to increase at higher elevations under climate warming, while lower elevation tree growth is expected to decline. However, numerous studies have found tree radial growth responds consistently to climate along elevational gradients. Here, we sampled five plots across the subalpine Abies fabri forest belt on Gongga Mountain in the eastern Tibetan Plateau to determine tree radial growth trends and responses to climate. Three commonly used detrending methods all consistently showed that tree radial growth at high elevation (> 3100 m) increased, while tree growth declined at the lower elevations (2700 m–2900 m) over the lastmore » three decades. Increasing late-growing season temperature positively (p < 0.05) correlated to tree radial growth at higher elevations, but the sign of this relationship reversed to become negative at lower elevations. Moving-window correlation analyses indicated the difference between high and low elevations response to temperature variation increased strongly with warming. Placing our result into the global context, 62% of 39 published studies found that trees along elevation gradients respond divergently to warming, and that these are located in warmer and wetter regions of the Earth. Notably, 28% of studies found non-significant responses to temperature at both high and low elevations. Our findings in the subalpine mountain forest in the eastern Tibetan Plateau were consistent with the majority of published datasets, and imply increasing temperature benefit for tree populations at higher elevation, while warming dampens growth at lower elevations.« less
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Host-environment mismatches associated with subalpine fir decline in Colorado
Robin M. Reich; John E. Lundquist; Kristina Hughes
2016-01-01
Subalpine fir decline (SFD) has killed more trees in Coloradoâs high elevation forests than any other insect or disease problem. The widespread nature of this disorder suggests that the cause involves climatic factors. We examined the influence of varying combinations of average annual temperature and precipitation on the incidence and distribution of SFD. Climatic...
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D. R. Bowling; W. J. Massman; S. M. Schaeffer; S. P. Burns; R. K. Monson; M. W. Williams
2009-01-01
Considerable research has recently been devoted to understanding biogeochemical processes under winter snow cover, leading to enhanced appreciation of the importance of many winter ecological processes. In this study, a comprehensive investigation of the stable carbon isotope composition (δ 13C) of CO2 within a high-elevation subalpine...
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Ozone concentration characteristics at a high-elevation forest site
G. Wooldridge; K. Zeller; R. Musselman
1997-01-01
Atmospheric ozone concentrations have been monitored at a subalpine forest ecosystem site, 3180m above mean sea level (msl), and at a 2680m msl forest-steppe ecotone site 15km to the southeast. Ozone concentrations were monitored at three heights above the ground on a 30m tower at the higher elevation site, and on a 10m tower in a large meadow downwind of this site....
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Sparkle L. Malone; Anna W. Schoettle; Jonathan D. Coop
2018-01-01
Like many other high elevation alpine tree species, Rocky Mountain bristlecone pine (Pinus aristata Engelm.) may be particularly vulnerable to climate change. To evaluate its potential vulnerability to shifts in climate, we defined the suitable climate space for each of four genetic lineages of bristlecone pine and for other subalpine tree species in close proximity to...
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Robert R. Ziemer
1963-01-01
Abstract - The quantity of summer soil moisture loss from logged forest openings was related to the length of time since the creation of the opening in a study made in the subalpine forest zone of the Sierra Nevada west-side near the Central Sierra Snow Laboratory, California, within the elevational range of 6,000 to 7,000 feet. Soil moisture depletion was measured in...
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AmeriFlux US-CZ4 Sierra Critical Zone, Sierra Transect, Subalpine Forest, Shorthair
DOE Office of Scientific and Technical Information (OSTI.GOV)
Goulden, Michael
This is the AmeriFlux version of the carbon flux data for the site US-CZ4 Sierra Critical Zone, Sierra Transect, Subalpine Forest, Shorthair. Site Description - Half hourly data are available at https://www.ess.uci.edu/~california/. This site is one of four Southern Sierra Critical Zone Observatory flux towers operated along an elevation gradient (sites are USCZ1, USCZ2, USCZ3 and USCZ4). This site is a lodgepole pine subalpine woodland with no recent disturbance.
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Andrew J. Friedland; Eric K. Miller
1996-01-01
Patterns and trends in forest elemental cycling can become more apparent in the presence of atmospheric perturbations. High-elevation forests of the northeastern United States have received large amounts of atmospheric deposition of pollutants, which have altered natural elemental cycling and retention rates in a variety of ways. This study examined atmospheric...
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Constance I. Millar; Robert D. Westfall; Diane L. Delany; Matthew J. Bokach; Alan L. Flint; Lorraine E. Flint
2012-01-01
Whitebark pine (Pinus albicaulis Engelm.) in subalpine zones of eastern California experienced significant mortality from 2007 to 2010. Dying stands were dense (mean basal area 47.5 m2/ha), young (mean 176 years), and even-age; mean stand mortality was 70%. Stands were at low elevations (mean 2993 m), on northerly aspects, and...
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Chapter 16: Inland Habitat Associations of Marbled Murrelets in British Columbia
Alan E. Burger
1995-01-01
Most Marbled Murrelets (Brachyramphus marmoratus) in British Columbia nest in the Coastal Western Hemlock biogeoclimatic zone. In this zone, detection frequencies were highest in the moister ecosections and in low elevation forests. Nests and moderately high levels of activity were also found in some forest patches in the subalpine Mountain Hemlock...
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Processes regulating watershed chemical export during snowmelt, fraser experimental forest, Colorado
Stottlemyer, R.
2001-01-01
In the Central Rocky Mountains, snowfall dominates precipitation. Airborne contaminants retained in the snowpack can affect high elevation surface water chemistry during snowmelt. At the Fraser Experimental Forest (FEF), located west of the Continental Divide in Central Colorado, snowmelt dominates the annual hydrograph, and accounts for >95% of annual stream water discharge. During the winters of 1989-1993, we measured precipitation inputs, snowpack water equivalent (SWE) and ion content, and stream water chemistry every 7-10 days along a 3150-3500 m elevation gradient in the subalpine and alpine Lexen Creek watershed. The study objectives were to (1) quantify the distribution of SWE and snowpack chemical content with elevation and aspect, (2) quantify snowmelt rates, temperature of soil, snowpack, and air with elevation and aspect, and (3) use change in upstream-downstream water chemistry during snowmelt to better define alpine and subalpine flowpaths. The SWE increased with elevation (P - 3??C) temperatures throughout winter which resulted in significant snowpack ion loss. By snowpack PWE in mid May, the snowpack had lost almost half the cumulative precipitation H+, NH4+, and SO42- inputs and a third of the NO3- input. Windborne soil particulate inputs late in winter increased snowpack base cation content. Variation in subalpine SWE and snowpack ion content with elevation and aspect, and wind redistribution of snowfall in the alpine resulted in large year-to-year differences in the timing and magnitude of SWE, PWE, and snowpack ion content. The alpine stream water ion concentrations changed little during snowmelt indicating meltwater passed quickly through surface porous soils and was well mixed before entering the stream. Conversely, subalpine stream water chemistry was diluted during snowmelt suggesting much melt water moved to the stream as shallow subsurface lateral flow. Published by Elsevier Science B.V.
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The forest fire season at different elevations in Idaho
J. A. Larsen
1925-01-01
In any fire-ridden forest region, such as north Idaho, there is great need for a tangible basis by which to judge the length and the intensity of the fire season in different forest types and at different elevations. The major and natural forest types, such as the western yellow pine forests, the western white-pine forests, and the subalpine forests occur in...
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Earle F. Layser
1992-01-01
Onion Park is a floristically rich naturally occurring mountain meadow and wetland complex which is surrounded by subalpine forest. The grass- and wetlands comprising the Park contribute biological diversity to an otherwise predominantly lodgepole pine-forested, subalpine setting. Onion Park is located at 7400' elevation in the Little Belt Mountains, five miles...
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Taking the pulse of mountains: Ecosystem responses to climatic variability
Fagre, Daniel B.; Peterson, David L.; Hessl, Amy E.
2003-01-01
An integrated program of ecosystem modeling and field studies in the mountains of the Pacific Northwest (U.S.A.) has quantified many of the ecological processes affected by climatic variability. Paleoecological and contemporary ecological data in forest ecosystems provided model parameterization and validation at broad spatial and temporal scales for tree growth, tree regeneration and treeline movement. For subalpine tree species, winter precipitation has a strong negative correlation with growth; this relationship is stronger at higher elevations and west-side sites (which have more precipitation). Temperature affects tree growth at some locations with respect to length of growing season (spring) and severity of drought at drier sites (summer). Furthermore, variable but predictable climate-growth relationships across elevation gradients suggest that tree species respond differently to climate at different locations, making a uniform response of these species to future climatic change unlikely. Multi-decadal variability in climate also affects ecosystem processes. Mountain hemlock growth at high-elevation sites is negatively correlated with winter snow depth and positively correlated with the winter Pacific Decadal Oscillation (PDO) index. At low elevations, the reverse is true. Glacier mass balance and fire severity are also linked to PDO. Rapid establishment of trees in subalpine ecosystems during this century is increasing forest cover and reducing meadow cover at many subalpine locations in the western U.S.A. and precipitation (snow depth) is a critical variable regulating conifer expansion. Lastly, modeling potential future ecosystem conditions suggests that increased climatic variability will result in increasing forest fire size and frequency, and reduced net primary productivity in drier, east-side forest ecosystems. As additional empirical data and modeling output become available, we will improve our ability to predict the effects of climatic change across a broad range of climates and mountain ecosystems in the northwestern U.S.A.
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Pansing, Elizabeth R; Tomback, Diana F; Wunder, Michael B; French, Joshua P; Wagner, Aaron C
2017-11-01
Tree recruitment is a spatially structured process that may undergo change over time because of variation in postdispersal processes. We examined seed pilferage, seed germination, and seedling survival in whitebark pine to determine whether 1) microsite type alters the initial spatial pattern of seed caches, 2) higher abiotic stress (i.e. higher elevations) exacerbates spatial distribution changes, and 3) these postdispersal processes are spatially clustered. At two study areas, we created a seed distribution pattern by burying seed caches in microsite types frequently used by whitebark pine's avian seed disperser (Clark's nutcracker) in upper subalpine forest and at treeline, the latter characterized by high abiotic environmental stress. We monitored caches for two years for pilferage, germination, and seedling survival. Odds of pilferage (both study areas), germination (northern study area), and survival (southern study area) were higher at treeline relative to subalpine forest. At the southern study area, we found higher odds of 1) pilferage near rocks and trees relative to no object in subalpine forest, 2) germination near rocks relative to trees within both elevation zones, and 3) seedling survival near rocks and trees relative to no object at treeline. No microsite effects were detected at the northern study area. Findings indicated that the microsite distribution of seed caches changes with seed/seedling stage. Higher odds of seedling survival near rocks and trees were observed at treeline, suggesting abiotic stress may limit safe site availability, thereby shifting the spatial distribution toward protective microsites. Higher odds of pilferage at treeline, however, suggest rodents may limit treeline recruitment. Further, odds of pilferage were higher near rocks and trees relative to no object in subalpine forest but did not differ among microsites at treeline, suggesting pilferage can modulate the spatial structure of regeneration, a finding supported by limited clustering of postdispersal processes.
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Dead wood biomass and turnover time, measured by radiocarbon, along a subalpine elevation gradient.
Kueppers, Lara M; Southon, John; Baer, Paul; Harte, John
2004-12-01
Dead wood biomass can be a substantial fraction of stored carbon in forest ecosystems, and coarse woody debris (CWD) decay rates may be sensitive to climate warming. We used an elevation gradient in Colorado Rocky Mountain subalpine forest to examine climate and species effects on dead wood biomass, and on CWD decay rate. Using a new radiocarbon approach, we determined that the turnover time of lodgepole pine CWD (340+/-130 years) was roughly half as long in a site with 2.5-3 degrees C warmer air temperature, as that of pine (630+/-400 years) or Engelmann spruce CWD (800+/-960 and 650+/-410 years) in cooler sites. Across all sites and both species, CWD age ranged from 2 to 600 years, and turnover time was 580+/-180 years. Total standing and fallen dead wood biomass ranged from 4.7+/-0.2 to 54+/-1 Mg ha(-1), and from 2.8 to 60% of aboveground live tree biomass. Dead wood biomass increased 75 kg ha(-1) per meter gain in elevation and decreased 13 Mg ha(-1) for every degree C increase in mean air temperature. Differences in biomass and decay rates along the elevation gradient suggest that climate warming will lead to a loss of dead wood carbon from subalpine forest.
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Baron, Jill S.; Ojima, Dennis S.; Holland, Elisabeth A.; Parton, William J.
1994-01-01
We employed grass and forest versions of the CENTURY model under a range of N deposition values (0.02–1.60 g N m−2 y−1) to explore the possibility that high observed lake and stream N was due to terrestrial N saturation of alpine tundra and subalpine forest in Loch Vale Watershed, Rocky Mountain National Park, Colorado. Model results suggest that N is limiting to subalpine forest productivity, but that excess leachate from alpine tundra is sufficient to account for the current observed stream N. Tundra leachate, combined with N leached from exposed rock surfaces, produce high N loads in aquatic ecosystems above treeline in the Colorado Front Range. A combination of terrestrial leaching, large N inputs from snowmelt, high watershed gradients, rapid hydrologic flushing and lake turnover times, and possibly other nutrient limitations of aquatic organisms constrain high elevation lakes and streams from assimilating even small increases in atmospheric N. CENTURY model simulations further suggest that, while increased N deposition will worsen the situation, nitrogen saturation is an ongoing phenomenon.
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AmeriFlux US-NR1 Niwot Ridge Forest (LTER NWT1)
Blanken, Peter [University of Colorado
2016-01-01
This is the AmeriFlux version of the carbon flux data for the site US-NR1 Niwot Ridge Forest (LTER NWT1). Site Description - The Niwot Ridge AmeriFlux site is located in a subalpine forest ecosystem just below the Continental Divide near Nederland, CO. The site is located at 3050 m elevation, within 600m of the NOAA C1 long-term monitoring station, approximately 8 km east of the Continental Divide. The surrounding subalpine forest is ~97 years old and in a state of aggradation, having recovered from early twentieth century logging (Monson, et al. Global Change Biology (2002), 8 459-478).
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Decay of subalpine fir in Colorado
Thomas E. Hinds; Frank G. Hawksworth; Ross W. Davidson
1960-01-01
Spruce-fir is one of the major forest types in the central Rocky Mountains. Engelmann spruce, Picea engelmanni Parry, is usually the predominant species with subalpine fir, Abies lasiocarpa (Hook. ) Nutt., making up one-fourth or less of the total volume. Lodgepole pine, Pinus contorta Dougl. ex Loud., is frequently present at the lower elevations of the spruce-fir...
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Constance I. Millar; Robert D. Westfall; Diane L. Delany; Alan L. Flint; Lorraine E. Flint
2015-01-01
Over the period 1883â2013, recruitment of subalpine limber pine (Pinus flexilis E. James) and Great Basin bristlecone pine (Pinus longaeva D.K. Bailey) above the upper tree line, below the lower tree line, and across middle-elevation forest borders occurred at localized sites across four mountain ranges in the western Great...
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Anita C. Risch; Martin F. Jurgensen; Deborah S. Page-Dumroese; Otto Wildi; Martin Schultz
2008-01-01
Vegetation changes following agricultural land abandonment at high elevation - which is frequent in Europe - could have a major impact on carbon (C) sequestration. However, most information on the effects of vegetation changes on ecosystem C stocks originates from low-elevation studies on reforestation or early successional forests, and little is known about how these...
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Bleaching of leaf litter and associated microfungi in subboreal and subalpine forests.
Hagiwara, Yusuke; Matsuoka, Shunsuke; Hobara, Satoru; Mori, Akira S; Hirose, Dai; Osono, Takashi
2015-10-01
Fungal decomposition of lignin leads to the whitening, or bleaching, of leaf litter, especially in temperate and tropical forests, but less is known about such bleaching in forests of cooler regions, such as boreal and subalpine forests. The purposes of the present study were to examine the extent of bleached area on the surface of leaf litter and its variation with environmental conditions in subboreal and subalpine forests in Japan and to examine the microfungi associated with the bleaching of leaf litter by isolating fungi from the bleached portions of the litter. Bleached area accounted for 21.7%-32.7% and 2.0%-10.0% of total leaf area of Quercus crispula and Betula ermanii, respectively, in subboreal forests, and for 6.3% and 18.6% of total leaf area of B. ermanii and Picea jezoensis var. hondoensis, respectively, in a subalpine forest. In subboreal forests, elevation, C/N ratio and pH of the FH layer, and slope aspect were selected as predictor variables for the bleached leaf area. Leaf mass per area and lignin content were consistently lower in the bleached area than in the nonbleached area of the same leaves, indicating that the selective decomposition of acid unhydrolyzable residue (recalcitrant compounds such as lignin, tannins, and cutins) enhanced the mass loss of leaf tissues in the bleached portions. Isolates of a total of 11 fungal species (6 species of Ascomycota and 5 of Basidiomycota) exhibited leaf-litter-bleaching activity under pure culture conditions. Two fungal species (Coccomyces sp. and Mycena sp.) occurred in both subboreal and subalpine forests, which were separated from each other by approximately 1100 km.
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Brian V. Smithers; Malcolm P. North; Constance I. Millar; Andrew M. Latimer
2017-01-01
In response to climate warming, subalpine treelines are expected to move up in elevation since treelines are generally controlled by growing season temperature. Where treeline is advancing, dispersal differences and early life stage environmental tolerances are likely to affect how species expand their ranges. Species with an establishment advantage will...
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Bird-habitat relationships in subalpine riparian shrublands of the Central Rocky Mountains
Deborah M. Finch
1987-01-01
Breeding birds were counted in 1982, 1983, and 1984 using the spot-map method on seven 8.1-ha plots In the Medicine Bow National Forest, Wyoming. At elevations of 2,280 to 3,000 m, riparian habitats were structurally simple, dominated by one or more bush willow species. Subalpine riparian avifaunas were depauperate with only four abundant species - song sparrow, white-...
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2018-01-01
Like many other high elevation alpine tree species, Rocky Mountain bristlecone pine (Pinus aristata Engelm.) may be particularly vulnerable to climate change. To evaluate its potential vulnerability to shifts in climate, we defined the suitable climate space for each of four genetic lineages of bristlecone pine and for other subalpine tree species in close proximity to bristlecone pine forests. Measuring changes in the suitable climate space for lineage groups is an important step beyond models that assume species are genetically homogenous. The suitable climate space for bristlecone pine in the year 2090 is projected to decline by 74% and the proportional distribution of suitable climate space for genetic lineages shifts toward those associated with warmer and wetter conditions. The 2090 climate space for bristlecone pine exhibits a bimodal distribution along an elevation gradient, presumably due to the persistence of the climate space in the Southern Rocky Mountains and exclusion at mid-elevations by conditions that favor the climate space of other species. These shifts have implications for changes in fire regimes, vulnerability to pest and pathogens, and altered carbon dynamics across the southern Rockies, which may reduce the likelihood of bristlecone pine trees achieving exceptional longevity in the future. The persistence and expansion of climate space for southern bristlecone pine genetic lineage groups in 2090 suggests that these sources may be the least vulnerable in the future. While these lineages may be more likely to persist and therefore present opportunities for proactive management (e.g., assisted migration) to maintain subalpine forest ecosystem services in a warmer world, our findings also imply heighted conservation concern for vulnerable northern lineages facing range contractions. PMID:29554097
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Long-term landscape changes in a subalpine spruce-fir forest in central Utah, USA
Jesse L. Morris; R. Justin DeRose; Andrea R. Brunelle
2015-01-01
In Western North America, increasing wildfire and outbreaks of native bark beetles have been mediated by warming climate conditions. Bioclimatic models forecast the loss of key high elevation species throughout the region. This study uses retrospective vegetation and fire history data to reconstruct the drivers of past disturbance and environmental change....
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Heart Rots of Engelmann Spruce and Subalpine Fir in the Central Rocky Mountain Region (FIDL)
T.E. Hinds
1977-01-01
Engelmann spruce (Picea engelmannii)-subalpine fir (Abies lasiocarpa) forests are widely distributed in western North America--from the northern Rocky Mountains of British Columbia and Alberta southward into Arizona and New Mexico. They occur at elevations of 2,000 to 7,000 feet in their northern range whereas they are found from about 8,000 to 12,000 feet in the south...
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Chadwick A. Moore; Ward W. McCaughey
1997-01-01
Snow accumulation in forested watersheds is controlled by climate, elevation, topographic factors and vegetation structure. Conifers affect snow accumulation principally by intercepting snow with the canopy which may later be sublimated. Various tree, stand, species and canopy densities of a subalpine fir habitat (ALBANASC) in central Montana were studied to determine...
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Fire disturbance effects in subalpine forests of north central Washington.
R. Schellhaas; D. Spurbeck; P. Ohlson; et al.
2001-01-01
Assessment of the historical disturbance effects across landscapes can provide an important foundation for land management decisions aimed at ecosystem restoration and maintenance. There have been numerous investigations regarding the historic disturbance patterns and resulting stand characteristics in low-elevation, dry forests of the inland west (Arno 1995, Everett...
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Establishment patterns of whitebark pine following fire in the Canadian Rockies
Brendan Wilson
2011-01-01
I examined the regeneration of whitebark pine (Pinus albicaulis) and four other high elevation conifers in young subalpine forest following two stand replacing fires in the Canadian Rockies. These were the Vermilion Pass fire of 1968, located in Kootenay and Banff national parks, and the Rock Canyon Creek fire of 1960, located approximately 125 km further southeast in...
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W. Lopushlnsky; D. Zabowskl; T.D. Anderson
1992-01-01
Logging residues were broadcast burned, piled and burned, removed, or left in place after clearcutting in a high-elevation subalpine fir (Abies lasiocarpa (Hook.) Nutt.) lodgepole pine (Pinus contorta Dougl. ex Loud.) forest in north-central Washington. Survival, height growth and foliar nutrient content of planted Douglas-fir...
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Summer evapotranspiration trends as related to time after logging of forests in Sierra Nevada
Robert R. Ziemer
1964-01-01
Abstract - The quantity of summer soil moisture lost from logged forest openings was related to the length of time since the creation of the opening, in the subalpine forest zone of the Sierra Nevada west side, near the Central Sierra Snow Laboratory, California, at an elevation of 6000 to 7000 feet. Soil moisture depletion was measured in forest openings which were...
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Robert C. Musselman; John L. Korfmacher
2007-01-01
A study was begun in the winter of 2000-2001 and continued through the winter of 2001-2002 to examine air quality at the Green Rock snowmobile staging area at 2,985 m elevation in the Snowy Range of Wyoming. The study was designed to evaluate the effects of winter recreation snowmobile activity on air quality at this high elevation site by measuring levels of...
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Increasing elevation of fire in the Sierra Nevada and implications for forest change
Schwartz, Mark W.; Butt, Nathalie; Dolanc, Christopher R.; Holguin, Andrew; Moritz, Max A.; North, Malcolm P.; Safford, Hugh D.; Stephenson, Nathan L.; Thorne, James H.; van Mantgem, Phillip J.
2015-01-01
Fire in high-elevation forest ecosystems can have severe impacts on forest structure, function and biodiversity. Using a 105-year data set, we found increasing elevation extent of fires in the Sierra Nevada, and pose five hypotheses to explain this pattern. Beyond the recognized pattern of increasing fire frequency in the Sierra Nevada since the late 20th century, we find that the upper elevation extent of those fires has also been increasing. Factors such as fire season climate and fuel build up are recognized potential drivers of changes in fire regimes. Patterns of warming climate and increasing stand density are consistent with both the direction and magnitude of increasing elevation of wildfire. Reduction in high elevation wildfire suppression and increasing ignition frequencies may also contribute to the observed pattern. Historical biases in fire reporting are recognized, but not likely to explain the observed patterns. The four plausible mechanistic hypotheses (changes in fire management, climate, fuels, ignitions) are not mutually exclusive, and likely have synergistic interactions that may explain the observed changes. Irrespective of mechanism, the observed pattern of increasing occurrence of fire in these subalpine forests may have significant impacts on their resilience to changing climatic conditions.
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DOE Office of Scientific and Technical Information (OSTI.GOV)
Zolbrod, A.N.; Peterson, D.L.
1995-06-01
Subalpine and alpine vegetation may be particularly sensitive to climatic change, such as expected temperature increases and altered precipitation patterns with global warming. The gap replacement model ZELIG was modified and used to examine transient and steady-state changes in altitudinal treeline, tree species distribution, and forest structure and composition along elevation gradients in the Olympic Mountains, Washington, under a range of temperature and precipitation changes. Changes in vegetation pattern were examined for north vs. south aspects, and wet (southwest) vs. dry (northeast) regions of the mountains. The seedling establishment subroutine in ZELIG was improved to specifically model the complexities ofmore » tree invasion in subalpine meadows and include empirical data. A function allowing for stand replacement fire was also added in order to examine the role of altered disturbance regimes on vegetation change. Results indicate that distribution of tree species will change under various climate change scenarios, but future elevation of treeline depends greatly on precipitation levels, disturbance frequency, and aspect.« less
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Restoring the subalpine mosaic
Ward McCaughey; Cathy Stewart; Colin Hardy
1996-01-01
Fire dependent lodgepole pine communities are an important component of upper elevation forests of the Bitterroot Mountains and the Northern Rockies in generaL A study of 1937 aerial photos from the Bitterroot National Forest is describing the pre- 1900 disturbances that shaped lodgepole pine mosaics, The 1937 photos will be compared with stand history information...
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NASA Astrophysics Data System (ADS)
Castanha, C.; Germino, M. J.; Torn, M. S.; Ferrenberg, S.; Harte, J.; Kueppers, L. M.
2010-12-01
The effect of climate change on future ranges of treeline species is poorly understood. For example, it is not known whether trees will recruit into the alpine, above the current treeline, and whether population-level differences in trees will mediate range shifts. At Niwot Ridge, Colorado, we used common gardens and climate manipulations to test predictions that warming will lead to greater recruitment at and beyond the cold edge of these species ranges, and will reduce recruitment at the warm edge. Seed from local populations of limber pine and Englemann spruce was harvested and reciprocally planted in 3 experimental sites spanning an elevation gradient from lower subalpine forest (10,000’), to the upper subalpine treeline ecotone (11,000’), to the alpine tundra (11,300’). In Fall 2009 seeds were sown into 20 plots at each site. Overhead infrared heaters targeted increases in growing season surface soil temperature of 4-5°C. The heating treatment, which began in October 2009, was crossed with manual watering, which was initiated following snowmelt in 2010. Over the 2010 growing season, we surveyed seedling germination and mortality weekly. Germination began in early May at the forest site, in early June at the krummholz site, and in early July at the alpine site. Depending on the site and plot, heating accelerated germination by 1 to 4 weeks. Seed source elevation, species, and site all affected germination, with effects for the two species also depending on site. At all sites, lower elevation, warm-edge populations had higher germination rates than high-elevation, cool-edge populations, indicating a potential bottleneck for germination of the high elevation seed sources in the adjacent alpine tundra. At all sites, survival was generally higher for pine than for spruce. Watering tended to enhance pine germinant survival while heating tended to depress spruce germinant survival. Our results indicate that the alpine tundra, generally considered an inhospitable environment, was not favorable for Englemann spruce, even with warming. In contrast, once seeds were introduced, the alpine tundra proved favorable to limber pine germination, irrespective of the climate manipulation.
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Instream wood loads in montane forest streams of the Colorado Front Range, USA
NASA Astrophysics Data System (ADS)
Jackson, Karen J.; Wohl, Ellen
2015-04-01
Although several studies examine instream wood loads and associated geomorphic effects in streams of subalpine forests in the U.S. Southern Rocky Mountains, little is known of instream wood loads in lower elevation, montane forests of the region. We compare instream wood loads and geomorphic effects between streams draining montane forest stands of differing age (old growth versus younger) and disturbance history (healthy versus infested by mountain pine beetles). We examined forest stand characteristics, instream wood load, channel geometry, pool volume, and sediment storage in 33 pool-riffle or plane-bed stream reaches with objectives of determining whether (i) instream wood and geomorphic effects differed significantly among old-growth, younger, healthy, and beetle-infested forest stands and (ii) wood loads correlated with valley and channel characteristics. Wood loads were standardized to drainage area, stream gradient, reach length, bankfull width, and floodplain area. Streams flowing through old-growth forests had significantly larger wood loads and logjam volumes (pairwise t-tests), as well as logjam frequencies (Kruskal-Wallis test), residual pool volume, and fine sediment storage around wood than streams flowing through younger forests. Wood loads in streams draining beetle-infested forest did not differ significantly from those in healthy forest stands, but best subset regression models indicated that elevation, stand age, and beetle infestation were the best predictors of wood loads in channels and on floodplains, suggesting that beetle infestation is affecting instream wood characteristics. Wood loads are larger than values from subalpine streams in the same region and jams are larger and more closely spaced. We interpret these differences to reflect greater wood piece mobility in subalpine zone streams. Stand age appears to exert the dominant influence on instream wood characteristics within pool-riffle streams in the study area rather than beetle infestation, although this may reflect the relatively recent nature (< 10 years) of the infestation.
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David L. Peterson; Darren R. Anderson
1990-01-01
The wood of lodgepole pines and whitebark pines from a high elevation site in the east central Sierra Nevada of California was analyzed for chemical content to determine whether there were any temporal patterns of chemical distribution in tree rings. Cores were taken from 10 trees of each species and divided into 5-year increments for chemical analysis. Correlation...
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Bianca N.I. Eskelson; Vicente J. Monleon
2018-01-01
Whitebark pine (Pinus albicaulis Engelm.) is a keystone species that provides a variety of ecosystem services, such as soil stabilization and protracted snowmelts. Found in many upper subalpine forests in Western North America (Arno and Hoff 1989), whitebark pine is an important high-elevation food source for grizzly bears (Bjoernlie and others 2014) and numerous other...
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Response of high-elevation forests in the Olympic Mountains to climatic change
Zolbrod, A.N.; Peterson, D.L.
1999-01-01
The gap model ZELIG was used to examine the effects of increased temperature (2°C) and altered precipitation on high-elevation ecosystems of the Olympic Mountains, Washington, U.S.A. Changes in tree species distribution and abundance, as well as stand biomass, were examined on north and south aspects in the dry northeast (NE) and wet southwest (SW) regions of the Olympics for (i) warmer, (ii) warmer and 20% wetter, and (iii) warmer and 20% drier climatic-change scenarios. Dominant tree species shift upwards 300-600 m in elevation in the SW, with subalpine meadows and Tsuga mertensiana (Bong.) Carr. forests being replaced by Abies amabilis (Dougl.) Forbes forests at higher elevations and A. amabilis forests being replaced by Tsuga heterophylla (Raf.) Sarg. forests at lower elevations. In the NE, drought-tolerant species become dominant approximately 200 m lower than present, with A. lasiocarpa dominating the north aspect and Pinus contorta Dougl. ex Loud. the south aspect. Biomass increases in the SW and generally decreases in the NE, depending on aspect and precipitation regime. This study suggests that species and site-specific responses at mesoscale (e.g., wet vs. dry climatic regime) and microscale (e.g., north vs. south aspect) resolutions must be characterized to quantify the variation in potential effects of climatic change on forest vegetation in mountainous regions.
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Ravindran, Anita; Yang, Shang-Shyng
2015-08-01
Microbial biomass plays an important role in nutrient transformation and conservation of forest and grassland ecosystems. The objective of this study was to determine the microbial biomass among three vegetation types in subalpine mountain forest soils of Taiwan. Tatachia is a typical high-altitude subalpine temperate forest ecosystem in Taiwan with an elevation of 1800-3952 m and consists of three vegetation types: spruce, hemlock, and grassland. Three plots were selected in each vegetation type. Soil samples were collected from the organic layer, topsoil, and subsoil. Microbial biomass carbon (Cmic) was determined by the chloroform fumigation-extraction method, and microbial biomass nitrogen (Nmic) was determined from the total nitrogen (Ntot) released during fumigation-extraction. Bacteria, actinomycetes, fungi, cellulolytic microbes, phosphate-solubilizing microbes, and nitrogen-fixing microbes were also counted. The Cmic and Nmic were highest in the surface soil and declined with the soil depth. These were also highest in spruce soils, followed by in hemlock soils, and were lowest in grassland soils. Cmic and Nmic had the highest values in the spring season and the lowest values in the winter season. Cmic and Nmic had significantly positive correlations with total organic carbon (Corg) and Ntot. Contributions of Cmic and Nmic, respectively, to Corg and Ntot indicated that the microbial biomass was immobilized more in spruce and hemlock soils than in grassland soils. Microbial populations of the tested vegetation types decreased with increasing soil depth. Cmic and Nmic were high in the organic layer and decreased with the depth of layers. These values were higher for spruce and hemlock soils than for grassland soils. Positive correlations were observed between Cmic and Nmic and between Corg and Ntot. Copyright © 2014. Published by Elsevier B.V.
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Area burned in alpine treeline ecotones reflects region-wide trends
C. Alina Cansler; Donald McKenzie; Charles B. Halpern
2016-01-01
The direct effects of climate change on alpine treeline ecotones â the transition zones between subalpine forest and non-forested alpine vegetation â have been studied extensively, but climate-induced changes in disturbance regimes have received less attention. To determine if recent increases in area burned extend to these higher-elevation landscapes, we analysed...
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Luo, Ya-Huang; Liu, Jie; Tan, Shao-Lin; Cadotte, Marc William; Wang, Yue-Hua; Xu, Kun; Li, De-Zhu; Gao, Lian-Ming
2016-01-01
Understanding how communities respond to environmental variation is a central goal in ecology. Plant communities respond to environmental gradients via intraspecific and/or interspecific variation in plant functional traits. However, the relative contribution of these two responses to environmental factors remains poorly tested. We measured six functional traits (height, leaf thickness, specific leaf area (SLA), leaf carbon concentration (LCC), leaf nitrogen concentration (LNC) and leaf phosphorus concentration (LPC)) for 55 tree species occurring at five elevations across a 1200 m elevational gradient of subalpine forests in Yulong Mountain, Southwest China. We examined the relative contribution of interspecific and intraspecific traits variability based on community weighted mean trait values and functional diversity, and tested how different components of trait variation respond to different environmental axes (climate and soil variables). Species turnover explained the largest amount of variation in leaf morphological traits (leaf thickness and SLA) across the elevational gradient. However, intraspecific variability explained a large amount of variation (49.3%-76.3%) in three other traits (height, LNC and LPC) despite high levels of species turnover. The detection of limiting similarity in community assembly was improved when accounting for both intraspecific and interspecific variability. Different components of trait variation respond to different environmental axes, especially soil water content and climatic variables. Our results indicate that intraspecific variation is critical for understanding community assembly and evaluating community response to environmental change.
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Luo, Ya-Huang; Liu, Jie; Tan, Shao-Lin; Cadotte, Marc William; Wang, Yue-Hua; Xu, Kun; Li, De-Zhu; Gao, Lian-Ming
2016-01-01
Understanding how communities respond to environmental variation is a central goal in ecology. Plant communities respond to environmental gradients via intraspecific and/or interspecific variation in plant functional traits. However, the relative contribution of these two responses to environmental factors remains poorly tested. We measured six functional traits (height, leaf thickness, specific leaf area (SLA), leaf carbon concentration (LCC), leaf nitrogen concentration (LNC) and leaf phosphorus concentration (LPC)) for 55 tree species occurring at five elevations across a 1200 m elevational gradient of subalpine forests in Yulong Mountain, Southwest China. We examined the relative contribution of interspecific and intraspecific traits variability based on community weighted mean trait values and functional diversity, and tested how different components of trait variation respond to different environmental axes (climate and soil variables). Species turnover explained the largest amount of variation in leaf morphological traits (leaf thickness and SLA) across the elevational gradient. However, intraspecific variability explained a large amount of variation (49.3%–76.3%) in three other traits (height, LNC and LPC) despite high levels of species turnover. The detection of limiting similarity in community assembly was improved when accounting for both intraspecific and interspecific variability. Different components of trait variation respond to different environmental axes, especially soil water content and climatic variables. Our results indicate that intraspecific variation is critical for understanding community assembly and evaluating community response to environmental change. PMID:27191402
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Integrating limnological characteristics of high mountain lakes into the landscape of a natural area
Larson, Gary L.; Wones, A.; McIntire, C.D.; Samora, B.
1994-01-01
A general conceptual watershed-lake model of the complex interactions among climatic conditions, watershed location and characteristics, lake morphology, and fish predation was used to evaluate limnological characteristics of high mountain lakes. Our main hypothesis was that decreasing elevation in mountainous terrain corresponds to an increase in diversity of watershed size and lake area, depth, temperature, nutrient concentrations, and productivity. A second hypothesis was that watershed location and aspect relative to climatic gradients within mountainous terrain influences the limnological characteristics of the lakes. We evaluated these hypotheses by examining watershed location, aspect and size; lake morphology; water quality; and phytoplankton and zooplankton community characteristics among high mountain forest and subalpine lakes in Mount Rainier National Park. Although many of the comparisons between all forest and subalpine lakes were statistically insignificant, the results revealed trends that were consistent with our hypotheses. The forest lake group included more lakes with larger watersheds, larger surface areas, greater depths, higher concentrations of nutrients, and higher algal biovolumes than did the group of subalpine lakes. Deep lakes, which were mostly of the forest lake type, exhibited thermal stratification and relatively high values of some of the water-quality variables near the lake bottoms. However, the highest near-surface water temperatures and phytoplankton densities and the taxonomic structures of the phytoplankton and zooplankton assemblages were more closely related to geographical location, which corresponded to a west-east climate gradient in the park, than to lake type. Some crustacean and rotifer taxa, however, were limited in distribution by lake type. Fish predation did not appear to play an important role in the structure of the crustacean zooplankton communities at the genus level with the exception of Mowich Lake, where crustacean taxa were absent from the zooplankton community. This was the only lake inhabited by a true zooplanktivourous species of fish.
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[Spatial pattern of sub-alpine forest restoration in west Sichuan].
Zhang, Yuandong; Liu, Shirong; Zhao, Changming
2005-09-01
West Sichuan sub-alpine is an extension of Qinghai-Tibet Plateau to southeast China, which is covered mainly with dark coniferous forest. As a result of long-term large scale over-logging, the forests have been greatly reduced and degraded. Nowadays, the forest restoration and regeneration in the region are being highlighted. Selecting Miyaluo as a case study area and employing the methods of plot investigation, ETM image interpretation, and overlaying vegetation map with digital topography, this paper analyzed the relations between the appearance and origin of four forest vegetation types, along with their topography differentiation and spatial patterns after a large scale logging and regeneration. The results showed that the appearance of forest vegetations was significantly correlated with their origin. Old coniferous forests (OC) were primitive ones, middle-aged and young coniferous forests (MYC) were from artificial regeneration, deciduous broadleaf forests (DB) were natural secondary ones, while mixed coniferous and deciduous forests (MCD) were partly from natural secondary ones and others from the conjunct action of artificial and natural regeneration. The main cut area in Miyaluo located in the sites with elevation from 2 800 to 3 600 m, where forest restoration appeared difference among different aspects. MYC was mainly distributed on sunny and half-sunny slope, DB and MCD were distributed on shady and half-shady slope, and OC were reserved on the sites with elevation more than 3 600 m. In the process of forest restoration, the four forest vegetation types were in mosaic pattern, and the landscape was seriously fragmentized.
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NASA Astrophysics Data System (ADS)
Konno, A.
2015-12-01
The coniferous forest (largely composed of Abies mariesii) is presently the typical vegetation of the subalpine zone in Japan. Pollen analysis revealed that few A. mariesii were present during the last glacial period, and the species began to expand to the subalpine zone during the Holocene (Morita, 1992). However, on Mt. Akita-Komagatake in northern Japan, the expected predominance of A. mariesii is not extensively observed, and the predominant vegetation is instead the dwarf bamboo (Sasa kurilensis). It is unknown why the area under coniferous forest is small in this region. Therefore, I examined this issue from the perspectives of (1) distribution of vegetation, (2) geomorphology, (3) soil moisture, and (4) vegetation history. (1) Precise digital elevation model data and photographic interpretation showed that this coniferous forest was densely distributed in a flat segment considered to be formed by a landslide; (2) this landslide is thought to have occurred up to 3,699 ± 26 yr BP because a boring-core specimen from the landslide included the AK-3 tephra layer (2,300-2,800 yr BP: Wachi et al, 1997) and the radiocarbon date of the lowermost humic soil layer was 3,699 ± 26 yr BP; (3) the soil in the forest area had higher volumetric water content than that in the non-forest area; and (4) phytolith analysis revealed that the main species in the study site was initially dwarf bamboo, but coniferous forest replaced it after the Towada-a tephra (1035 cal. BP, Machida and Arai, 1992) layer fell. These results suggest that soil water conditions changed because of the formation of the flat segment by the landslide, and the coniferous forest was consequently established. However, the landslide only indirectly affected the formation of the coniferous forest, because the forest developed over several thousand years after the landslide occurred. In other words, more direct reasons for the establishment of the coniferous forest may involve changes in soil moisture. This unresolved issue warrants further investigation of the vegetation on the subalpine zone.
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NASA Astrophysics Data System (ADS)
Bowling, D. R.; Blanken, P.; Burns, S. P.; Frankenberg, C.; Grossman, K.; Lin, J. C.; Logan, B. A.; Magney, T. S.; Richardson, A. D.; Stutz, J.; Aubrecht, D.
2017-12-01
Temperate and boreal conifer forests are dormant for many months during the cold season, during which they continue to absorb solar radiation. Thus they exhibit a marked seasonal change in light-use efficiency, challenging our ability to monitor gross primary productivity (GPP) from remote sensing platforms. We are studying the factors limiting the seasonality of photosynthesis of a high-elevation subalpine forest in Colorado. Using in-situ thermal imagery, we find that foliage in winter is sometimes near the optimum temperature for photosynthesis, but photosynthesis is shut down for most of the cold season. Water transport is limited by blockage of sap transport by frozen boles, but not by frozen soils. Foliar carotenoid content exhibits strong upregulation during winter, driven largely by increase in the pool size of the photoprotective xanthophyll cycle, but with no seasonal change in chlorophyll content. The seasonality of GPP is strongly linked to xanthophyll cycle conversion state and thawing of boles. Ongoing research includes examination of leaf-level chlorophyll fluorescence emission and gas exchange, combined with measurement of canopy-level spectral reflectance and solar-induced fluorescence (SIF) at high spatio-temporal resolution using a custom tower-based PhotoSpec scanning spectrometer system. These results will be synthesized in the context of using SIF as a metric for GPP.
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Warming-induced upslope advance of subalpine forest is severely limited by geomorphic processes
Macias-Fauria, Marc; Johnson, Edward A.
2013-01-01
Forests are expected to expand into alpine areas because of climate warming, causing land-cover change and fragmentation of alpine habitats. However, this expansion will only occur if the present upper treeline is limited by low-growing season temperatures that reduce plant growth. This temperature limitation has not been quantified at a landscape scale. Here, we show that temperature alone cannot realistically explain high-elevation tree cover over a >100-km2 area in the Canadian Rockies and that geologic/geomorphic processes are fundamental to understanding the heterogeneous landscape distribution of trees. Furthermore, upslope tree advance in a warmer scenario will be severely limited by availability of sites with adequate geomorphic/topographic characteristics. Our results imply that landscape-to-regional scale projections of warming-induced, high-elevation forest advance into alpine areas should not be based solely on temperature-sensitive, site-specific upper-treeline studies but also on geomorphic processes that control tree occurrence at long (centuries/millennia) timescales. PMID:23569221
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NASA Astrophysics Data System (ADS)
Schwörer, C.; Fisher, D. M.; Gavin, D. G.; Temperli, C.; Bartlein, P. J.
2015-12-01
Mountain forest composition and distribution is strongly affected by temperature and is expected to shift to higher elevations with climate change. However, warmer winters will also lead to an upward shift of the snowline and a decrease in snowpack at lower and intermediate elevations. In the mountain ranges of Western North America, snowpack plays an important role in providing additional moisture during the dry summer months. It is therefore unclear if the projected climate change will lead to a rise of subalpine forest due to a longer growing season or a contraction due to drought stress. Since forest succession processes take place over decades and centuries we use LandClim, a dynamic vegetation model, to assess the impact of climate change on mountain forests on the Olympic Peninsula (Washington, USA). As a reality check we first simulate vegetation dynamics since the last Ice Age and compare model output with paleobotanical data from five natural archives that span the topographic and climatic gradients on the Peninsula. LandClim produces realistic present-day species compositions with respect to elevation and precipitation gradients. Moreover, the simulations of forest dynamics for the last 16,000 years generally agree with the pollen and macrofossil data. We then simulated mountain forests under future climate projections. As a result, our model indicates drastic changes in species composition with a replacement of mountain hemlock (Tsuga mertensiana) by more drought-resistant species such as subalpine fir (Abies lasiocarpa). On the drier, eastern side of the Peninsula, the model even suggests a lowering of timberline due to insufficient moisture availability in shallow alpine soils. Our results have important implications for ecosystem managers and stress the urgency of climate change mitigation.
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Subalpine Forest Carbon Cycling Short- and Long-Term Influence ofClimate and Species
DOE Office of Scientific and Technical Information (OSTI.GOV)
Kueppers, L.; Harte, J.
2005-08-23
Ecosystem carbon cycle feedbacks to climate change comprise one of the largest remaining sources of uncertainty in global model predictions of future climate. Both direct climate effects on carbon cycling and indirect effects via climate-induced shifts in species composition may alter ecosystem carbon balance over the long term. In the short term, climate effects on carbon cycling may be mediated by ecosystem species composition. We used an elevational climate and tree species composition gradient in Rocky Mountain subalpine forest to quantify the sensitivity of all major ecosystem carbon stocks and fluxes to these factors. The climate sensitivities of carbon fluxesmore » were species-specific in the cases of relative above ground productivity and litter decomposition, whereas the climate sensitivity of dead wood decay did not differ between species, and total annual soil CO2 flux showed no strong climate trend. Lodge pole pine relative productivity increased with warmer temperatures and earlier snowmelt, while Engelmann spruce relative productivity was insensitive to climate variables. Engelmann spruce needle decomposition decreased linearly with increasing temperature(decreasing litter moisture), while lodgepole pine and subalpine fir needle decay showed a hump-shaped temperature response. We also found that total ecosystem carbon declined by 50 percent with a 2.88C increase in mean annual temperature and a concurrent 63 percent decrease ingrowing season soil moisture, primarily due to large declines in mineral soil and dead wood carbon. We detected no independent effect of species composition on ecosystem C stocks. Overall, our carbon flux results suggest that, in the short term, any change in subalpine forest net carbon balance will depend on the specific climate scenario and spatial distribution of tree species. Over the long term, our carbon stock results suggest that with regional warming and drying, Rocky Mountain subalpine forest will be a net source of carbon to the atmosphere.« less
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Spatial and temporal patterns in water chemistry of two high elevation lakes in southeast Wyoming
Robert C. Musselman
1995-01-01
The Glacier Lakes Ecosystem Experiments Site (GLEES) was established to examine the effects of atmospheric deposition and climate change on alpine and subalpine ecosystems. The site contains East Glacier Lake (3282 m elevation) and West Glacier Lake (3276 m elevation), and their watersheds. These two small lakes are located 120m from each other at the alpine/subalpine...
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Falandysz, Jerzy; Saba, Martyna; Liu, Hong-Gao; Li, Tao; Wang, Ji-Peng; Wiejak, Anna; Zhang, Ji; Wang, Yuan-Zhong; Zhang, Dan
2016-12-01
This study aimed to investigate and discuss the occurrence and accumulation of mercury in the fruiting bodies of wild-growing fungi (Macromycetes) collected from montane forests in two regions of southwestern China with differences in soil geochemistry, climate and geographical conditions. Fungal mycelia in soils of the subalpine region of the Minya Konka (Gongga Mountain) in Sichuan and in the highlands of Yunnan efficiently accumulated mercury in fruiting bodies (mushrooms). The examined sites in Yunnan with highly mineralized red and yellow soils showed Hg contents ranging from 0.066 to 0.28 mg kg -1 dry biomass (db) which is roughly similar to the results obtained for samples collected from sites with dark soils relatively rich in organic matter from a remote, the subalpine region of Minya Konka. Due to the remoteness of the subalpine section of Minya Konka, as well as its elevation and climate, airborne mercury from long-range transport could be deposited preferentially on the topsoil and the Hg levels determined in soil samples taken beneath the fruiting bodies were up to 0.48 mg kg -1 dry matter. In Yunnan, with polymetallic soils (Circum-Pacific Mercuriferous Belt), Amanita mushrooms showed mercury in caps of fruiting bodies of up to 7.3 mg kg -1 dry biomass. Geogenic Hg from the mercuriferous belt seems to be the overriding source of mercury accumulated in mushrooms foraged in the regions of Yunnan, while long-range atmospheric transport and subsequent deposition are the mercury sources for specimens foraged in the region of Minya Konka.
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Rundel, Philip W.; Keeley, Jon E.
2016-01-01
Patterns of elevational distribution of alien plant species in the southern Sierra Nevada of California were used to test the hypothesis that alien plant species invading high elevations around the world are typically climate generalists capable of growing across a wide elevational range. The Sierra Nevada has been heavily impacted for more than a century and a half, first by heavy grazing up into high elevation meadows, followed by major logging, and finally, by impacts associated with recreational use. The comparative elevational patterns of distribution and growth form were compared for native and alien plant species in the four families (Asteraceae, Brassicaceae, Fabaceae, and Poaceae) that contribute the majority of naturalized aliens in the study area. The distribution of realized climatic niche breadth, as measured by elevational range of occurrence, was virtually identical for alien and native species, with both groups showing a roughly Gaussian distribution peaking with species whose range covers a span of 1500–1999 m. In contrast to alien species, which only rarely occurred at higher elevations, native species showed a distribution of upper elevation limits peaking at 3000–3499 m, an elevation that corresponds to the zone of upper montane and subalpine forests. Consistent with a hypothesis of abiotic limitations, only a few alien species have been ecologically successful invaders at subalpine and alpine elevations above 2500 m. The low diversity of aliens able to become established in these habitats is unlikely due to dispersal limitations, given the long history of heavy grazing pressure at high elevations across this region. Instead, this low diversity is hypothesized to be a function of life history traits and multiple abiotic stresses that include extremes of cold air and soil temperature, heavy snowfall, short growing seasons, and low resource availability. These findings have significant implications for resource managers.
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Boot, Claudia M.; Hall, Ed K.; Denef, Karolien; Baron, Jill S.
2016-01-01
Elevated nitrogen (N) deposition due to increased fossil fuel combustion and agricultural practices has altered global carbon (C) cycling. Additions of reactive N to N-limited environments are typically accompanied by increases in plant biomass. Soil C dynamics, however, have shown a range of different responses to the addition of reactive N that seem to be ecosystem dependent. We evaluated the effect of N amendments on biogeochemical characteristics and microbial responses of subalpine forest organic soils in order to develop a mechanistic understanding of how soils are affected by N amendments in subalpine ecosystems. We measured a suite of responses across three years (2011–2013) during two seasons (spring and fall). Following 17 years of N amendments, fertilized soils were more acidic (control mean 5.09, fertilized mean 4.68), and had lower %C (control mean 33.7% C, fertilized mean 29.8% C) and microbial biomass C by 22% relative to control plots. Shifts in biogeochemical properties in fertilized plots were associated with an altered microbial community driven by reduced arbuscular mycorrhizal (control mean 3.2 mol%, fertilized mean 2.5 mol%) and saprotrophic fungal groups (control mean 17.0 mol%, fertilized mean 15.2 mol%), as well as a decrease in N degrading microbial enzyme activity. Our results suggest that decreases in soil C in subalpine forests were in part driven by increased microbial degradation of soil organic matter and reduced inputs to soil organic matter in the form of microbial biomass.
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Castanha, C.; Torn, M.S.; Germino, M.J.; Weibel, Bettina; Kueppers, L.M.
2013-01-01
Background: Seedling germination and survival is a critical control on forest ecosystem boundaries, such as at the alpine–treeline ecotone. In addition, while it is known that species respond individualistically to the same suite of environmental drivers, the potential additional effect of local adaptation on seedling success has not been evaluated. Aims: To determine whether local adaptation may influence the position and movement of forest ecosystem boundaries, we quantified conifer seedling recruitment in common gardens across a subalpine forest to alpine tundra gradient at Niwot Ridge, Colorado, USA. Methods: We studied Pinus flexilis and Picea engelmannii grown from seed collected locally at High (3400 m a.s.l.) and Low (3060 m a.s.l.) elevations. We monitored emergence and survival of seeds sown directly into plots and survival of seedlings germinated indoors and transplanted after snowmelt. Results: Emergence and survival through the first growing season was greater for P. flexilis than P. engelmannii and for Low compared with High provenances. Yet survival through the second growing season was similar for both species and provenances. Seedling emergence and survival tended to be greatest in the subalpine forest and lowest in the alpine tundra. Survival was greater for transplants than for field-germinated seedlings. Conclusions: These results suggest that survival through the first few weeks is critical to the establishment of natural germinants. In addition, even small distances between seed sources can have a significant effect on early demographic performance – a factor that has rarely been considered in previous studies of tree recruitment and species range shifts.
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Camarero, J Julio; Gazol, Antonio; Galván, Juan Diego; Sangüesa-Barreda, Gabriel; Gutiérrez, Emilia
2015-02-01
Theory predicts that the postindustrial rise in the concentration of CO2 in the atmosphere (c(a)) should enhance tree growth either through a direct fertilization effect or indirectly by improving water use efficiency in dry areas. However, this hypothesis has received little support in cold-limited and subalpine forests where positive growth responses to either rising ca or warmer temperatures are still under debate. In this study, we address this issue by analyzing an extensive dendrochronological network of high-elevation Pinus uncinata forests in Spain (28 sites, 544 trees) encompassing the whole biogeographical extent of the species. We determine if the basal area increment (BAI) trends are linked to climate warming and increased c(a) by focusing on region- and age-dependent responses. The largest improvement in BAI over the past six centuries occurred during the last 150 years affecting young trees and being driven by recent warming. Indeed, most studied regions and age classes presented BAI patterns mainly controlled by temperature trends, while growing-season precipitation was only relevant in the driest sites. Growth enhancement was linked to rising ca in mature (151-300 year-old trees) and old-mature trees (301-450 year-old trees) from the wettest sites only. This finding implies that any potential fertilization effect of elevated c(a) on forest growth is contingent on tree features that vary with ontogeny and it depends on site conditions (for instance water availability). Furthermore, we found widespread growth decline in drought-prone sites probably indicating that the rise in ca did not compensate for the reduction in water availability. Thus, warming-triggered drought stress may become a more important direct driver of growth than rising ca in similar subalpine forests. We argue that broad approaches in biogeographical and temporal terms are required to adequately evaluate any effect of rising c(a) on forest growth. © 2014 John Wiley & Sons Ltd.
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Climate and landscape drive the pace and pattern of conifer encroachment into subalpine meadows.
Lubetkin, Kaitlin C; Westerling, Anthony LeRoy; Kueppers, Lara M
2017-09-01
Mountain meadows have high biodiversity and help regulate stream water release following the snowmelt pulse. However, many meadows are experiencing woody plant encroachment, threatening these ecosystem services. While there have been field surveys of individual meadows and remote sensing-based landscape-scale studies of encroachment, what is missing is a broad-scale, ground-based study to understand common regional drivers, especially at high elevations, where land management has often played a less direct role. With this study, we ask: What are the climate and landscape conditions conducive to woody plant encroachment at the landscape scale, and how has historical climate variation affected tree recruitment in subalpine meadows over time? We measured density of encroaching trees across 340 subalpine meadows in the central Sierra Nevada, California, USA, and used generalized additive models (GAMs) to determine the relationship between landscape-scale patterns of encroachment and meadow environmental properties. We determined ages of trees in 30 survey meadows, used observed climate and GAMs to model the relationship between timing of recruitment and climate since the early 1900s, and extrapolated recruitment patterns into the future using downscaled climate scenarios. Encroachment was high among meadows with lodgepole pine (Pinus contorta Douglas ex Loudon var. murrayana (Balf.) Engelm.) in the immediate vicinity, at lower elevations, with physical conditions favoring strong soil drying, and with maximum temperatures above or below average. Climatic conditions during the year of germination were unimportant, with tree recruitment instead depending on a 3-yr seed production period prior to germination and a 6-yr seedling establishment period following germination. Recruitment was high when the seed production period had high snowpack, and when the seedling establishment period had warm summer maximum temperatures, high summer precipitation, and high snowpack. Applying our temporal model to downscaled output from four global climate models indicated that the average meadow will shift to forest by the end of the 21st century. Sierra Nevada meadow encroachment by conifers is ubiquitous and associated with climate conditions increasingly favorable for tree recruitment, which will lead to substantial changes in subalpine meadows and the ecosystem services they provide. © 2017 by the Ecological Society of America.
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Deep Undercooling of Tissue Water and Winter Hardiness Limitations in Timberline Flora 1
Becwar, Michael R.; Rajashekar, Channa; Bristow, Katherine J. Hansen; Burke, Michael J.
1981-01-01
Deep undercooled tissue water, which froze near −40 C, was found in winter collected stem and leaf tissue of the dominant timberline tree species of the Colorado Rocky Mountains, Engelmann spruce (Picea engelmannii (Parry) Engelm.) and subalpine fir (Abies lasiocarpa (Hook.) Nutt.), and in numerous other woody species in and below the subalpine vegetation zone. Previous work on numerous woody plants indicates that deep undercooling in xylem makes probable a −40 C winter hardiness limit in stem tissue. Visual injury determinations and electrolyte loss measurements on stem tissue revealed injury near −40 C associated with the freezing of the deep undercooled stem tissue water. These results suggest that the winter hardiness limit of this woody flora is near −40 C. The relevance of deep undercooling in relation to timberline, the upper elevational limit of the subalpine forest, is discussed. PMID:16661852
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Chapter 10. Dynamics of subalpine forests
Dennis H. Knight
1994-01-01
The boreal owl's fairly specific habitat requirements restrict its range in the conterminous U.S. to subalpine forests (see Chapter 9). These forests provide tree cavities, uncrusted snow that facilitates preying on small mammals, and cool microclimates essential for summer roosting. Such forests also provide habitat for the owl's prey which consists...
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Biogeochemical impacts of wildfires over four millennia in a Rocky Mountain subalpine watershed.
Dunnette, Paul V; Higuera, Philip E; McLauchlan, Kendra K; Derr, Kelly M; Briles, Christy E; Keefe, Margaret H
2014-08-01
Wildfires can significantly alter forest carbon (C) storage and nitrogen (N) availability, but the long-term biogeochemical legacy of wildfires is poorly understood. We obtained a lake-sediment record of fire and biogeochemistry from a subalpine forest in Colorado, USA, to examine the nature, magnitude, and duration of decadal-scale, fire-induced ecosystem change over the past c. 4250 yr. The high-resolution record contained 34 fires, including 13 high-severity events within the watershed. High-severity fires were followed by increased sedimentary N stable isotope ratios (δ15N) and bulk density, and decreased C and N concentrations--reflecting forest floor destruction, terrestrial C and N losses, and erosion. Sustained low sediment C : N c. 20-50 yr post-fire indicates reduced terrestrial organic matter subsidies to the lake. Low sedimentary δ15N c. 50-70 yr post-fire, coincident with C and N recovery, suggests diminishing terrestrial N availability during stand development. The magnitude of post-fire changes generally scaled directly with inferred fire severity. Our results support modern studies of forest successional C and N accumulation and indicate pronounced, long-lasting biogeochemical impacts of wildfires in subalpine forests. However, even repeated high-severity fires over millennia probably did not deplete C or N stocks, because centuries between high-severity fires allowed for sufficient biomass recovery. © 2014 The Authors. New Phytologist © 2014 New Phytologist Trust.
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High-mountain lakes as a hotspot of dissolved organic matter production in a changing climate
NASA Astrophysics Data System (ADS)
Abood, P. H.; Williams, M. W.; McKnight, D. M.; Hood, E. H.
2004-12-01
Changes in climate may adversely affect mountain environments before downstream ecosystems are affected. Steep topography, thin soils with limited extent, sparse vegetation, short growing seasons, and climatic extremes (heavy snowfalls, cold temperatures, high winds), all contribute to the sensitivity of high mountain environments to perturbations. Here we evaluate the role of oligatrophic high-elevation lakes as "hot spots" of aquatic production that may respond to changes in temperature, precipitation amount, and pollution deposition faster and more directly than co-located terrestrial ecosystems. Our research was conducted in the Rocky Mountains, USA. Water samples were collected for dissolved organic carbon (DOC), other solutes, and water isotopes over the course of the runoff season along a longitudinal transect of North Boulder Creek in the Colorado Front Range from the continental divide and alpine areas to downstream forested systems. Sources of DOC were evaluated using chemical fractionation with XAD-8 resins and fluorescence spectroscopy. There was net DOC production in the two alpine lakes but not for the forested subalpine lake. Oxygen-18 values showed that water residence times in lakes increased dramatically in late summer compared to snowmelt. Chemical fractionation of DOC showed there was a increase in the non-humic acid content across the summer of 2003 at all elevations, with alpine waters showing greater increases than subalpine waters. The fluorescence properties of DOC and water isotopes suggested that DOC in aquatic systems was primarily derived from terrestrial precursor material during snowmelt. However, fluorescence properties of DOC in high-elevation lakes on the recession limb of the hydrograph suggest DOC derived from algal and microbial biomass in the lakes was a more important source of DOC in late summer and fall. Alpine lakes produced 14 times more DOC on unit area basis compared to the surrounding terrestrial ecosystems. We hypothesize that much of the authochthonous production is a result of algal growth in alpine lakes caused by the increases in nitrogen deposition from wetfall.
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Aaron F. Wells
2006-01-01
This guide presents a classification of the deep canyon and subalpine riparian and wetland vegetation types of the Malheur, Umatilla, and Wallowa-Whitman National Forests. A primary goal of the deep canyon and subalpine riparian and wetland classification was a seamless linkage with the midmontane northeastern Oregon riparian and wetland classification provided by...
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DOE Office of Scientific and Technical Information (OSTI.GOV)
Tuskan, Gerald A; Yin, Tongming
Lack of aspen (Populus tremuloides) recruitment and canopy replacement of aspen stands that grow on the edges of grasslands on the low-elevation elk (Cervus elaphus) winter range of Rocky Mountain National Park (RMNP) in Colorado has been a cause of concern for more than 70 years (Packard, 1942; Olmsted, 1979; Stevens, 1980; Hess, 1993; R.J. Monello, T.L. Johnson, and R.G. Wright, Rocky Mountain National Park, 2006, written commun.). These aspen stands are a significant resource since they are located close to the park's road system and thus are highly visible to park visitors. Aspen communities are integral to the ecologicalmore » structure of montane and subalpine landscapes because they contain high native species richness of plants, birds, and butterflies (Chong and others, 2001; Simonson and others, 2001; Chong and Stohlgren, 2007). These low-elevation, winter range stands also represent a unique component of the park's plant community diversity since most (more than 95 percent) of the park's aspen stands grow in coniferous forest, often on sheltered slopes and at higher elevations, while these winter range stands are situated on the low-elevation ecotone between the winter range grasslands and some of the park's drier coniferous forests.« less
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Sensitivity of a high-elevation Rocky Mountain watershed to altered climate and CO2
Baron, Jill S.; Hartman, Melannie D.; Band, L.E.; Lammers, R.B.
2000-01-01
We explored the hydrologic and ecological responses of a headwater mountain catchment, Loch Vale watershed, to climate change and doubling of atmospheric CO2 scenarios using the Regional Hydro-Ecological Simulation System (RHESSys). A slight (2°C) cooling, comparable to conditions observed over the past 40 years, led to greater snowpack and slightly less runoff, evaporation, transpiration, and plant productivity. An increase of 2°C yielded the opposite response, but model output for an increase of 4°C showed dramatic changes in timing of hydrologic responses. The snowpack was reduced by 50%, and runoff and soil water increased and occurred 4–5 weeks earlier with 4°C warming. Alpine tundra photosynthetic rates responded more to warmer and wetter conditions than subalpine forest, but subalpine forest showed a greater response to doubling of atmospheric CO2 than tundra. Even though water use efficiency increased with the double CO2 scenario, this had little effect on basin-wide runoff because the catchment is largely unvegetated. Changes in winter and spring climate conditions were more important to hydrologic and vegetation dynamics than changes that occurred during summer.
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K. R. Sherrill; M. A. Lefsky; J. B. Bradford; M. G. Ryan
2008-01-01
This study evaluates the relative ability of simple light detection and ranging (lidar) indices (i.e., mean and maximum heights) and statistically derived canonical correlation analysis (CCA) variables attained from discrete-return lidar to estimate forest structure and forest biomass variables for three temperate subalpine forest sites. Both lidar and CCA explanatory...
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K.R. Sherrill; M.A. Lefsky; J.B. Bradford; M.G. Ryan
2008-01-01
This study evaluates the relative ability of simple light detection and ranging (lidar) indices (i.e., mean and maximum heights) and statistically derived canonical correlation analysis (CCA) variables attained from discrete-return lidar to estimate forest structure and forest biomass variables for three temperate subalpine forest sites. Both lidar and CCA explanatory...
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Moyes, Andrew B.; Castanha, Cristina; Germino, Matthew J.; Kueppers, Lara M.
2013-01-01
Continued changes in climate are projected to alter the geographic distributions of plant species, in part by affecting where individuals can establish from seed. We tested the hypothesis that warming promotes uphill redistribution of subalpine tree populations by reducing cold limitation at high elevation and enhancing drought stress at low elevation. We seeded limber pine (Pinus flexilis) into plots with combinations of infrared heating and water addition treatments, at sites positioned in lower subalpine forest, the treeline ecotone, and alpine tundra. In 2010, first-year seedlings were assessed for physiological performance and survival over the snow-free growing season. Seedlings emerged in midsummer, about 5–8 weeks after snowmelt. Low temperature was not observed to limit seedling photosynthesis or respiration between emergence and October, and thus experimental warming did not appear to reduce cold limitation at high elevation. Instead, gas exchange and water potential from all sites indicated a prevailing effect of summer moisture stress on photosynthesis and carbon balance. Infrared heaters raised soil growing degree days (base 5 °C, p p 3 m-3 consistently corresponded with moderate and severe indications of drought stress in midday stem water potential, stomatal conductance, photosynthesis, and respiration. Seedling survival was greater in watered plots than in heated plots (p = 0.01), and negatively related to soil growing degree days and duration of exposure to θ 3 m-3 in a stepwise linear regression model (p < 0.0001). We concluded that seasonal moisture stress and high soil surface temperature imposed a strong limitation to limber pine seedling establishment across a broad elevation gradient, including at treeline, and that these limitations are likely to be enhanced by further climate warming.
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Li, Guixiang; Xu, Guorui; Shen, Congcong; Tang, Yong; Zhang, Yuxin; Ma, Keming
2016-11-01
Above- and below-ground organisms are closely linked, but how elevational distribution pattern of soil microbes shifting across the treeline still remains unknown. Sampling of 140 plots with transect, we herein investigated soil bacterial distribution pattern from a temperate forest up to a subalpine meadow along an elevational gradient using Illumina sequencing. Our results revealed distinct elevational patterns of bacterial diversity above and below the treeline in responding to changes in soil conditions: a hollow elevational pattern in the forest (correlated with soil temperature, pH, and C:N ratio) and a significantly decreasing pattern in the meadow (correlated with soil pH, and available phosphorus). The bacterial community structure was also distinct between the forest and meadow, relating to soil pH in the forest and soil temperature in the meadow. Soil bacteria did not follow the distribution pattern of herb diversity, but bacterial community structure could be predicted by herb community composition. These results suggest that plant communities have an important influence on soil characteristics, and thus change the elevational distribution of soil bacteria. Our findings are useful for future assessments of climate change impacts on microbial community.
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N(2)O emissions and source processes in snow-covered soils in the Swiss Alps.
Mohn, Joachim; Steinlin, Christine; Merbold, Lutz; Emmenegger, Lukas; Hagedorn, Frank
2013-01-01
Nitrous oxide (N2O) emissions from snow-covered soils represent a significant fraction of the annual flux from alpine, subalpine or cold-temperate regions. In winter 2010-2011, we investigated the temporal variability of N2O emissions and source processes from a subalpine valley in the Swiss Alps. The study included regular measurements of N2O snow profiles at a fixed location and an intensive sampling campaign along a transversal cut through the valley with grassland at the bottom and coniferous forest at the slopes. During the intensive campaign, recently developed laser spectroscopy was employed for high-precision N2O isotopomer analysis. Maximum N2O fluxes (0.77±0.64 nmol m(-2) h(-1)) were found for periods with elevated air temperature and, in contrast to our expectations, were higher from forest than from grassland in mid-February. At maximum snow height (63 cm) the main N2O source processes were heterotrophic denitrification and nitrifier denitrification. The reduction of N2O by heterotrophic denitrifiers was much more pronounced for the grassland compared with the forest soil, as indicated by the (15)N site preferences of 16.4±11.5 ‰ (grassland) and-1.6±2.1 ‰ (forest). This illustrates the potential of laser spectroscopic N2O isotopomer analysis for the identification of source processes even at low emission rates in nutrient poor ecosystems.
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Constance I. Millar; Robert D. Westfall; Diane L. Delany; John C. King; Lisa J. Graumlich
2004-01-01
Four independent studies of conifer growth between 1880 and 2002 in upper elevation forests of the central Sierra Nevada, California, U.S.A., showed correlated multidecadal and century-long responses associated with climate. Using tree-ring and ecological plot analysis, we studied annual branch growth of krummholz Pinus albicaulis; invasion by P....
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Diana F. Tomback; Lynn M. Resler; Robert E. Keane; Elizabeth R. Pansing; Andrew J. Andrade; Aaron C. Wagner
2016-01-01
Whitebark pine (Pinus albicaulis) has the largest and most northerly distribution of any white pine (Subgenus Strobus) in North America, encompassing 18° latitude and 21° longitude in western mountains. Within this broad range, however, whitebark pine occurs within a narrow elevational zone, including upper subalpine and treeline forests, and functions...
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NASA Astrophysics Data System (ADS)
Yager, D. B.; Burchell, A.; Robinson, R.; Odell, S.; Dick, R. P.; Johnson, C. A.; Hidinger, J.; Rathke, D.
2007-12-01
There is now widespread agreement that, if the climate is to be stabilized, then net greenhouse gas emissions must be greatly reduced (IPCC, 2007). The need to reduce net CO2 emissions plus the possible economic and environmental ramifications of not addressing climate change have stimulated important atmospheric carbon mitigation actions, as well as, studies to understand and quantify potential carbon sinks. Soils represent a potentially large and environmentally significant natural carbon reservoir. Increasing the natural terrestrial sequestration potential (NTS) of soils is among the seven, "Sokolow CO2 stabilization wedges' or carbon management strategies needed to thwart doubling of atmospheric CO2. Additionally, high plains to subalpine temperate soils tend to be less susceptible to baseline C pool declines due to global warming than are warmer regions and are important ecosystems in which to quantify soil carbon storage capacity. To examine the potential of magnesium silicate-bearing soils to sequester additional carbon, we sampled 60 high plains prairie to subalpine forest soil horizons derived from weathering of Tertiary-age dacite-andesite- basalt compositions in Colorado, U.S.A.: the San Luis Valley, San Juan Volcanic Field, Grand Mesa, White River- Roan Plateau (Flat Tops), Rocky Mountain National Park, Front Range and propylitically-altered terrain in the western San Juan Volcanic field containing secondary magnesium silicates (chlorite-species). Data for C, N, O (total conc., isotopes), metals, major and trace elements, Hg, S, microbial enzymes (β-glucosidase, arylsulfatase, acid neutralizing capacity (ANC), and 14C radiocarbon dates are reported. Samples demonstrate variable but elevated C relative to average global soil C. In particular, the propylitically-altered rocks have a high instantaneous ANC in laboratory tests (> 20 kg/ton CaCO3 equivalent) and derivative forest soils containing low-temperature charcoal "burn" horizons have high total organic carbon contents (12-14 Wt.% in the A-B horizons; 0 to 30 cm). These data are important to understanding the carbon sequestration potential that soils derived from intermediate to mafic igneous rocks can provide. Additionally, for range or forest management and mine waste remediation scenarios, this data suggests C mitigation efforts may be augmented by 'geomimicry' scenarios whereby projects model and enhance natural processes that support CO2 sequestration.
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Evidence for foliar endophytic nitrogen fixation in a widely distributed subalpine conifer
DOE Office of Scientific and Technical Information (OSTI.GOV)
Moyes, Andrew B.; Kueppers, Lara M.; Pett-Ridge, Jennifer
Coniferous forest nitrogen (N) budgets indicate unknown sources of N. A consistent association between limber pine ( Pinus flexilis) and potential N 2-fixing acetic acid bacteria (AAB) indicates that native foliar endophytes may supply subalpine forests with N.
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Evidence for foliar endophytic nitrogen fixation in a widely distributed subalpine conifer
Moyes, Andrew B.; Kueppers, Lara M.; Pett-Ridge, Jennifer; ...
2016-02-01
Coniferous forest nitrogen (N) budgets indicate unknown sources of N. A consistent association between limber pine ( Pinus flexilis) and potential N 2-fixing acetic acid bacteria (AAB) indicates that native foliar endophytes may supply subalpine forests with N.
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Late Quaternary environments and biogeography in the Great Basin
NASA Astrophysics Data System (ADS)
Thompson, R. S.; Mead, J. I.
1982-01-01
Plant and animal remains found in packrat ( Neotoma spp.) middens and cave fill from the eastern and southern Great Basin region reveal the presence of subalpine conifers and boreal mammals at relatively low elevations during the Late Wisconsin. Limber pine ( Pinus flexilis) and bristlecone pine ( P. longaeva) were important in the late Pleistocene plant communities throughout this region. Spruce ( Picea cf. engelmannii) and common juniper ( Juniperus communis) were present in some of the more northerly localities, and Douglas fir ( Pseudotsuga menziesii) and white fir ( Abies concolor) were present in southern and eastern localities. Single needle pinyon pine ( Pinus monophylla), common across this region today, was apparently not present north of the Sheep Range of southern Nevada during the Late Wisconsin. Pikas ( Ochotona cf. princeps), small boreal mammals present in only a few Great Basin mountain ranges today, were common throughout the region. Heather voles ( Phenacomys cf. intermedius) have been found in two cave fill deposits in Nevada, though they are unknown in the Great Basin today. Limber and bristlecone pines are generally restricted to rocky substrates in modern subalpine habitats in the Great Basin, and this may also have been the case when these plants grew at lower elevations during the Late Wisconsin. Subalpine conifers were present on the rock outcrops sampled by the packrat middens, but shrub communities, perhaps dominated by sagebrush ( Artemisia spp.), may have been present on alluvial valley-bottom substrates. Forested habitats would thus have been isolated habitat islands, as they are today. Boreal small mammals, including pikas and heather voles, were able to colonize the Great Basin mountain ranges during the late Pleistocene. We suggest that these mammals were able to survive in the intervening valley-bottoms under a cool-summer climatic regime, and that continuous forest or woodland corridors were not necessary for migration.
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Comparison of the abundance and composition of litter fauna in tropical and subalpine forests
G. Gonzalez; T.R. Seastedt
2000-01-01
In this study, we quantify the abundance and composition of the litter fauna in dry and wet tropical forests and north- and south-facing subalpine forests. We used the same litter species contained in litterbags across study sites to standardize for substrate conditions, and a single method of fauna extraction from the litter (Tullgren method). Fauna densities were...
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Space-time modelling of lightning-caused ignitions in the Blue Mountains, Oregon
Diaz-Avalos, Carlos; Peterson, D.L.; Alvarado, Ernesto; Ferguson, Sue A.; Besag, Julian E.
2001-01-01
Generalized linear mixed models (GLMM) were used to study the effect of vegetation cover, elevation, slope, and precipitation on the probability of ignition in the Blue Mountains, Oregon, and to estimate the probability of ignition occurrence at different locations in space and in time. Data on starting location of lightning-caused ignitions in the Blue Mountains between April 1986 and September 1993 constituted the base for the analysis. The study area was divided into a pixela??time array. For each pixela??time location we associated a value of 1 if at least one ignition occurred and 0 otherwise. Covariate information for each pixel was obtained using a geographic information system. The GLMMs were fitted in a Bayesian framework. Higher ignition probabilities were associated with the following cover types: subalpine herbaceous, alpine tundra, lodgepole pine (Pinus contorta Dougl. ex Loud.), whitebark pine (Pinus albicaulis Engelm.), Engelmann spruce (Picea engelmannii Parry ex Engelm.), subalpine fir (Abies lasiocarpa (Hook.) Nutt.), and grand fir (Abies grandis (Dougl.) Lindl.). Within each vegetation type, higher ignition probabilities occurred at lower elevations. Additionally, ignition probabilities are lower in the northern and southern extremes of the Blue Mountains. The GLMM procedure used here is suitable for analysing ignition occurrence in other forested regions where probabilities of ignition are highly variable because of a spatially complex biophysical environment.
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Provenance variability in nursery growth of subalpine fir
Charlie Cartwright; Cheng Ying
2011-01-01
Subalpine fir (Abies lasiocarpa [Hook] Nutt.) is a wide-ranging, high-elevation species in the interior of British Columbia. It is commonly harvested for lumber, but replanting of it is limited. Some reticence is based upon wood quality and rate of growth, but there are also seed and nursery culturing difficulties. This study investigated seedling growth traits of 111...
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James K. Agee
2000-01-01
Disturbance dynamics differ in the three subregions of the North American boreal forest (taiga, western United States, and eastern United States) where lynx are found, resulting in a range of potential effects on lynx populations. Fire severity tends to be high in most of the forest types where lynx habitat occurs, although subsequent succession will differ...
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Zheng, Bao-Qiang; Zou, Long-Hai; Li, Kui; Wan, Xiao; Wang, Yan
2017-01-01
Cypripedium tibeticum, a subalpine orchid species, inhabits various habitats of subalpine forests, mainly including the forest edge (FE), forest gap (FG), and understory (UST), which have significantly different light intensities (FE > FG > UST). However, the ecological and physiological influences caused by different light regimes in this species are still poorly understood. In the present study, photosynthetic, morphological, and reproductive characteristics were comprehensively studied in plants of C. tibeticum grown in three types of habitats. The photosynthetic capacities, such as the net photosynthetic rate, light-saturated photosynthesis (Pmax), and dry mass per unit leaf area (LMA), were higher in FE and FG than in UST according to light availability. Compared with FG, the populations in FE and UST suffer from excessively strong and inadequate radiation, respectively, which was further corroborated by the low Fv/Fm in FE and high apparent quantum yield (AQY) in FG. The leaves of the orchids had various proportions of constituents, such as the leaf area, thickness and (or) epidermal hair, to reduce damage from high radiation (including ultraviolet-b radiation) in FE and capture more light in FG and UST. Although the flower rate (FR) was positively correlated to both Pmax and the daily mean PAR, fruit-set only occurred in the populations in FG. The failures in FE and UST might be ascribed to changes in the floral functional structure and low biomass accumulation, respectively. Moreover, analysis of the demographic statistics showed that FG was an advantageous habitat for the orchid. Thus, C. tibeticum reacted to photosynthetic and morphological changes to adapt to different subalpine forest habitats, and neither full (under FE) nor low (UST) illumination was favorable for population expansion. These findings could serve as a guide for the protection and reintroduction of C. tibeticum and emphasize the importance of specific habitats for Cypripedium spp.
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Observation of Snow cover glide on Sub-Alpine Coniferous Forests in Mount Zao, Northeastern Japan
NASA Astrophysics Data System (ADS)
Sasaki, A.; Suzuki, K.
2017-12-01
This is the study to clarify the snow cover glide behavior in the sub-alpine coniferous forests on Mount Zao, Northeastern Japan, in the winter of 2014-2015. We installed the glide-meter which is sled type, and measured the glide motion on the slope of Abies mariesii forest and its surrounding slope. In addition, we observed the air temperature, snow depth, density of snow, and snow temperature to discuss relationship between weather conditions and glide occurrence. The snow cover of the 2014-15 winter started on November 13th and disappeared on April 21st. The maximum snow depth was 242 cm thick, it was recorded at February 1st. The snow cover glide in the surrounding slope was occurred first at February 10th, although maximum snow depth recorded on February 1st. The glide motion in the surrounding slope is continuing and its velocity was 0.4 cm per day. The glide in the surrounding slope stopped at March 16th. The cumulative amount of the glide was 21.1 cm. The snow cover glide in the A. mariesii forest was even later occurred first at February 21st. The glide motion of it was intermittent and extremely small. On sub-alpine zone of Mount Zao, snow cover glide intensity is estimated to be 289 kg/m2 on March when snow water equivalent is maximum. At same period, maximum snow cover glide intensity is estimated to be about 1000 kg/m2 at very steep slopes where the slope angle is about 35 degree. Although potential of snow cover glide is enough high, the snow cover glide is suppressed by stem of A. mariesii trees, in the sub-alpine coniferous forest.
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Seasonal patterns in soil surface CO2 flux under snow cover in 50 and 300 year old subalpine forests
Robert M. Hubbard; Michael G. Ryan; Kelly Elder; Charles C. Rhoades
2005-01-01
Soil CO2 flux can contribute as much as 60-80% of total ecosystem respiration in forests. Although considerable research has focused on quantifying this flux during the growing season, comparatively little effort has focused on non-growing season fluxes. We measured soil CO2 efflux through snow in 50 and 300 year old subalpine forest stands near Fraser CO. Our...
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Esther M. Cole; Malcolm North
2014-01-01
Many high-elevation regions in the western USA are protected public lands that remain relatively undisturbed by human impact. Over the last century, however, nonnative trout and cattle have been introduced to subalpine wetland habitats used by sensitive amphibian species. Our study compares the relative importance of cattle and trout impact on amphibian assemblages,...
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Restoration of plant cover on campsites in subalpine forests: Sawtooth Wilderness, Idaho
David N. Cole; Liese Dean; Debarah Taylor; Troy E. Hall
2012-01-01
This study assessed the effectiveness of restoration treatments in enhancing the growth of Vaccinium scoparium transplants and plants established from seed on six closed campsites in subalpine forests in the Sawtooth Wilderness, Idaho. In the primary experiment, the soil on all plots was scarified and amended with organic matter; plots varied regarding the type and...
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Drought-driven disturbance history characterizes a southern Rocky Mountain subalpine forest
R. Justin DeRose; James N. Long
2012-01-01
The view that subalpine forest vegetation dynamics in western North America are "driven" by a particular disturbance type (i.e., fire) has shaped our understanding of their disturbance regimes. In the wake of a recent (1990s) landscape- extent spruce beetle (Dendroctonus rufipennis Kirby) outbreak in the southern Rocky Mountains, we re-examined the temporal...
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Averett, Joshua P.; McCune, Bruce; Parks, Catherine G.; Naylor, Bridgett J.; DelCurto, Tim; Mata-González, Ricardo
2016-01-01
Mountain environments are currently among the ecosystems least invaded by non-native species; however, mountains are increasingly under threat of non-native plant invasion. The slow pace of exotic plant invasions in mountain ecosystems is likely due to a combination of low anthropogenic disturbances, low propagule supply, and extreme/steep environmental gradients. The importance of any one of these factors is debated and likely ecosystem dependent. We evaluated the importance of various correlates of plant invasions in the Wallowa Mountain Range of northeastern Oregon and explored whether non-native species distributions differed from native species along an elevation gradient. Vascular plant communities were sampled in summer 2012 along three mountain roads. Transects (n = 20) were evenly stratified by elevation (~70 m intervals) along each road. Vascular plant species abundances and environmental parameters were measured. We used indicator species analysis to identify habitat affinities for non-native species. Plots were ordinated in species space, joint plots and non-parametric multiplicative regression were used to relate species and community variation to environmental variables. Non-native species richness decreased continuously with increasing elevation. In contrast, native species richness displayed a unimodal distribution with maximum richness occurring at mid–elevations. Species composition was strongly related to elevation and canopy openness. Overlays of trait and environmental factors onto non-metric multidimensional ordinations identified the montane-subalpine community transition and over-story canopy closure exceeding 60% as potential barriers to non-native species establishment. Unlike native species, non-native species showed little evidence for high-elevation or closed-canopy specialization. These data suggest that non-native plants currently found in the Wallowa Mountains are dependent on open canopies and disturbance for establishment in low and mid elevations. Current management objectives including restoration to more open canopies in dry Rocky Mountain forests, may increase immigration pressure of non-native plants from lower elevations into the montane and subalpine zones. PMID:26824750
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Averett, Joshua P; McCune, Bruce; Parks, Catherine G; Naylor, Bridgett J; DelCurto, Tim; Mata-González, Ricardo
2016-01-01
Mountain environments are currently among the ecosystems least invaded by non-native species; however, mountains are increasingly under threat of non-native plant invasion. The slow pace of exotic plant invasions in mountain ecosystems is likely due to a combination of low anthropogenic disturbances, low propagule supply, and extreme/steep environmental gradients. The importance of any one of these factors is debated and likely ecosystem dependent. We evaluated the importance of various correlates of plant invasions in the Wallowa Mountain Range of northeastern Oregon and explored whether non-native species distributions differed from native species along an elevation gradient. Vascular plant communities were sampled in summer 2012 along three mountain roads. Transects (n = 20) were evenly stratified by elevation (~70 m intervals) along each road. Vascular plant species abundances and environmental parameters were measured. We used indicator species analysis to identify habitat affinities for non-native species. Plots were ordinated in species space, joint plots and non-parametric multiplicative regression were used to relate species and community variation to environmental variables. Non-native species richness decreased continuously with increasing elevation. In contrast, native species richness displayed a unimodal distribution with maximum richness occurring at mid-elevations. Species composition was strongly related to elevation and canopy openness. Overlays of trait and environmental factors onto non-metric multidimensional ordinations identified the montane-subalpine community transition and over-story canopy closure exceeding 60% as potential barriers to non-native species establishment. Unlike native species, non-native species showed little evidence for high-elevation or closed-canopy specialization. These data suggest that non-native plants currently found in the Wallowa Mountains are dependent on open canopies and disturbance for establishment in low and mid elevations. Current management objectives including restoration to more open canopies in dry Rocky Mountain forests, may increase immigration pressure of non-native plants from lower elevations into the montane and subalpine zones.
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NASA Astrophysics Data System (ADS)
Kueppers, L. M.
2010-12-01
Niche models and paleoecological studies indicate that future climate change will alter the geographic distributions of plant species. Changes in temperature, snowmelt timing, or moisture conditions at one edge of a species’ range may have different consequences for recruitment, carbon exchange, phenology, and survival than changes at another edge. Similarly, local genetic adaptation may constrain species and community responses to climate change. We have established a new experiment to investigate potential shifts in the distribution of subalpine tree species, and the alpine species they might replace. We are asking how tree species recruitment and alpine species growth and reproduction vary within their current ranges, and in response to temperature and soil moisture manipulations. We are also examining whether genetic provenance and ecosystem processes constrain tree seedling and alpine herb responses. Our Alpine Treeline Warming Experiment is located across three sites at Niwot Ridge, CO, ranging from near the lower limit of subalpine forest to alpine tundra. We use infrared heaters to raise growing season surface soil temperatures by 4-5°C, and to lengthen the growing season. The warming treatment is crossed with a soil moisture manipulation to distinguish effects due to higher temperatures from those due to drier soil. Each plot is a common garden sown with high and low elevation provenances of limber pine (Pinus flexilis) and Engelmann spruce (Picea engelmannii). We established an additional set of experimental plots to examine treatment effects on alpine species phenology, growth and reproduction. Under ambient conditions in 2009, tree seedling germination rate, lifespan, and first season survival was higher within the species’ current range than in the alpine, and for Engelmann spruce, was higher at the low elevation limit than the high elevation limit. Source population (low vs. high elevation) was a significant factor explaining natural variation in germination rates and timing, seedling physiology, and seedling survival. In 2010, the first season with experimental effects data, the timing of germination was substantially advanced with warming for both species, and warming appeared to increase germination rates for limber pine, but to depress rates for Engelmann spruce at treeline. Seedling carbon balance was negative at the warmest leaf temperatures and there is some indication that the low elevation provenance has a higher total assimilation rate and net carbon gain than the high elevation provenance. Water availability was an important driver of variation in carbon assimilation through the growing season. Our early results suggest that with higher germination rates and lower mortality rates, limber pine is better able to recruit into the alpine than Engelmann spruce, and that lower elevation provenances of limber pine are better at assimilating carbon for growth regardless of site. Ultimate success in seedling establishment may be more contingent on water availability than temperature, even at these high elevations.
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Berryman, E. M.; Barnard, H. R.; Adams, H. R.; ...
2015-02-10
Forest soil respiration is a major carbon (C) flux that is characterized by significant variability in space and time. In this paper, we quantified growing season soil respiration during both a drought year and a nondrought year across a complex landscape to identify how landscape and climate interact to control soil respiration. We asked the following questions: (1) How does soil respiration vary across the catchments due to terrain-induced variability in moisture availability and temperature? (2) Does the relative importance of moisture versus temperature limitation of respiration vary across space and time? And (3) what terrain elements are important formore » dictating the pattern of soil respiration and its controls? Moisture superseded temperature in explaining watershed respiration patterns, with wetter yet cooler areas higher up and on north facing slopes yielding greater soil respiration than lower and south facing areas. Wetter subalpine forests had reduced moisture limitation in favor of greater seasonal temperature limitation, and the reverse was true for low-elevation semiarid forests. Coincident climate poorly predicted soil respiration in the montane transition zone; however, antecedent precipitation from the prior 10 days provided additional explanatory power. A seasonal trend in respiration remained after accounting for microclimate effects, suggesting that local climate alone may not adequately predict seasonal variability in soil respiration in montane forests. Finally, soil respiration climate controls were more strongly related to topography during the drought year highlighting the importance of landscape complexity in ecosystem response to drought.« less
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DOE Office of Scientific and Technical Information (OSTI.GOV)
Berryman, E. M.; Barnard, H. R.; Adams, H. R.
Forest soil respiration is a major carbon (C) flux that is characterized by significant variability in space and time. In this paper, we quantified growing season soil respiration during both a drought year and a nondrought year across a complex landscape to identify how landscape and climate interact to control soil respiration. We asked the following questions: (1) How does soil respiration vary across the catchments due to terrain-induced variability in moisture availability and temperature? (2) Does the relative importance of moisture versus temperature limitation of respiration vary across space and time? And (3) what terrain elements are important formore » dictating the pattern of soil respiration and its controls? Moisture superseded temperature in explaining watershed respiration patterns, with wetter yet cooler areas higher up and on north facing slopes yielding greater soil respiration than lower and south facing areas. Wetter subalpine forests had reduced moisture limitation in favor of greater seasonal temperature limitation, and the reverse was true for low-elevation semiarid forests. Coincident climate poorly predicted soil respiration in the montane transition zone; however, antecedent precipitation from the prior 10 days provided additional explanatory power. A seasonal trend in respiration remained after accounting for microclimate effects, suggesting that local climate alone may not adequately predict seasonal variability in soil respiration in montane forests. Finally, soil respiration climate controls were more strongly related to topography during the drought year highlighting the importance of landscape complexity in ecosystem response to drought.« less
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Berryman, Erin Michele; Barnard, H.R.; Adams, H.R.; Burns, M.A.; Gallo, E.; Brooks, P.D.
2015-01-01
Forest soil respiration is a major carbon (C) flux that is characterized by significant variability in space and time. We quantified growing season soil respiration during both a drought year and a nondrought year across a complex landscape to identify how landscape and climate interact to control soil respiration. We asked the following questions: (1) How does soil respiration vary across the catchments due to terrain-induced variability in moisture availability and temperature? (2) Does the relative importance of moisture versus temperature limitation of respiration vary across space and time? And (3) what terrain elements are important for dictating the pattern of soil respiration and its controls? Moisture superseded temperature in explaining watershed respiration patterns, with wetter yet cooler areas higher up and on north facing slopes yielding greater soil respiration than lower and south facing areas. Wetter subalpine forests had reduced moisture limitation in favor of greater seasonal temperature limitation, and the reverse was true for low-elevation semiarid forests. Coincident climate poorly predicted soil respiration in the montane transition zone; however, antecedent precipitation from the prior 10 days provided additional explanatory power. A seasonal trend in respiration remained after accounting for microclimate effects, suggesting that local climate alone may not adequately predict seasonal variability in soil respiration in montane forests. Soil respiration climate controls were more strongly related to topography during the drought year highlighting the importance of landscape complexity in ecosystem response to drought.
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Jonathan D. Coop; Robert T. Massatti; Anna W. Schoettle
2010-01-01
These subalpine wildfires generated considerable, persistent increases in plant species richness at local and landscape scales, and a diversity of plant communities. The findings suggest that fire suppression in such systems must lead to reduced diversity. Concerns about post-fire invasion by exotic plants appear unwarranted in high-elevation wilderness settings.
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Forest Vegetation Monitoring Protocol for National Parks in the North Coast and Cascades Network
Woodward, Andrea; Hutten, Karen M.; Boetsch, John R.; Acker, Steven A.; Rochefort, Regina M.; Bivin, Mignonne M.; Kurth, Laurie L.
2009-01-01
Plant communities are the foundation for terrestrial trophic webs and animal habitat, and their structure and species composition are an integrated result of biological and physical drivers (Gates, 1993). Additionally, they have a major role in geologic, geomorphologic and soil development processes (Jenny, 1941; Stevens and Walker, 1970). Throughout most of the Pacific Northwest, environmental conditions support coniferous forests as the dominant vegetation type. In the face of anthropogenic climate change, forests have a global role as potential sinks for atmospheric carbon (Goodale and others, 2002). Consequently, knowledge of the status of forests in the three large parks of the NCCN [that is, Mount Rainier (MORA), North Cascades (NOCA), and Olympic (OLYM) National Parks] is fundamental to understanding the condition of Pacific Northwest ecosystems. Diverse climate and soil properties across the Pacific Northwest result in a variety of forest types (Franklin and Dyrness, 1973; Franklin and others, 1988; Henderson and others, 1989, 1992). The mountainous terrain of Mount Rainier, North Cascades, and Olympic National Parks create steep elevational and precipitation gradients within and among the parks: collectively, these parks span from sea level to more than 4,200 m; and include areas with precipitation from 90 to more than 500 cm. The resulting forests range from coastal rainforests with dense understories and massive trees draped with epiphytes; to areas with drought-adapted Ponderosa pines; to high-elevation subalpine fir forests interspersed with meadows just below treeline (table 1). These forests, in turn, are the foundation for other biotic communities constituting Pacific Northwest ecosystems.
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McKinney, Shawn T; Fiedler, Carl E; Tomback, Diana F
2009-04-01
Human-caused disruptions to seed-dispersal mutualisms increase the extinction risk for both plant and animal species. Large-seeded plants can be particularly vulnerable due to highly specialized dispersal systems and no compensatory regeneration mechanisms. Whitebark pine (Pinus albicaulis), a keystone subalpine species, obligately depends upon the Clark's Nutcracker (Nucifraga columbiana) for dispersal of its large, wingless seeds. Clark's Nutcracker, a facultative mutualist with whitebark pine, is sensitive to rates of energy gain, and emigrates from subalpine forests during periods of cone shortages. The invasive fungal pathogen Cronartium ribicola, which causes white pine blister rust, reduces whitebark pine cone production by killing cone-bearing branches and trees. Mortality from blister rust reaches 90% or higher in some whitebark pine forests in the Northern Rocky Mountains, USA, and the rust now occurs nearly rangewide in whitebark pine. Our objectives were to identify the minimum level of cone production necessary to elicit seed dispersal by nutcrackers and to determine how cone production is influenced by forest structure and health. We quantified forest conditions and ecological interactions between nutcrackers and whitebark pine in three Rocky Mountain ecosystems that differ in levels of rust infection and mortality. Both the frequency of nutcracker occurrence and probability of seed dispersal were strongly related to annual whitebark pine cone production, which had a positive linear association with live whitebark pine basal area, and negative linear association with whitebark pine tree mortality and rust infection. From our data, we estimated that a threshold level of approximately 1000 cones/ha is needed for a high likelihood of seed dispersal by nutcrackers (probability > or = 0.7), and that this level of cone production can be met by forests with live whitebark pine basal area > 5.0 m2/ha. The risk of mutualism disruption is greatest in northern most Montana (USA), where three-year mean cone production and live basal area fell below predicted threshold levels. There, nutcracker occurrence, seed dispersal, and whitebark pine regeneration were the lowest of the three ecosystems. Managers can use these threshold values to differentiate between restoration sites requiring planting of rust-resistant seedlings and sites where nutcracker seed dispersal can be expected.
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Banko, P.C.; Oboyski, P.T.; Slotterback, J.W.; Dougill, Steve J.; Goltz, Dan M.; Johnson, L.; Laut, M.E.; Murray, T.C.
2002-01-01
Aim: We evaluated how an elevation gradient affects: (1) the availability of food required by a specialist seed-eater, Loxioides bailleui Oustalet (Drepanidinae), or pallia, and hence the distribution of this endangered Hawaiian bird, and (2) the distribution of alien threats to Loxioides populations, their primary foods, and their dry-forest habitat, and hence strategies for their conservation. Location: We worked throughout the subalpine forest that encircles Mauna Kea Volcano, Hawai'i Island, but we focused our studies mainly on the western slope between 2000 and 3000 m elevation, where the gradient of elevation was greatest and palila were most abundant. Methods: We determined phenology and productivity patterns of the endemic dry-forest tree species, Sophora chrysophylla (Salisb.) Seem. (Fabaceae), or ma??mane, which provides Loxioides with most of their food, and another common endemic tree, Myoporum sandwicense A. Gray (Myoporaceae), or naio, which provides some resources, along a 786-m elevation gradient at monthly intervals for 10 years (Sophora only). We also determined the availability each month of moth larvae (Lepidoptera) for that were important in the diet of nestling and adult palila. In addition, we documented the incidence of parasitism on moth larvae by several wasp (Hymenoptera) and fly (Diptera) species, and we determined the distribution of predatory wasps and ants (Hymenoptera), which potentially threaten insect prey of birds. Percentage cover of alien grass species that pose fire threats in palila habitat and other weeds were assessed during one survey. Small mammal abundance and distribution were determined by trapping during three (rodent) or five (carnivore) surveys. Results: Sophora flower and seed (pod) availability varied predictably along the elevation gradient, with about 4 months separating peaks in reproduction at high and low elevations. This, together with highly variable production of flowers and pods within elevation strata, resulted in Sophora resources being available to Loxioides throughout the year on the western slope of Mauna Kea. Sophora produced flowers and pods more seasonally where gradients of elevation were short; thus, resources were available less consistently. In contrast, Myoporum produced flowers and fruits with little variation with respect to season or elevation. The availability of important insect prey of Loxioides was also related to elevation, in part because threats to Lepidoptera larvae from parasitic wasps were generally less at higher elevations. Threats to insect prey from predatory ants was also less at higher elevations but the abundance of predatory wasps was not related to elevation. Several weeds that pose the most serious threats to Loxioides habitat were more abundant at mid and low elevations, and alien grass cover was somewhat greater at mid elevation, thereby increasing fire risks in the centre of Loxioides habitat. Predatory mammals, in particular Felis catus Linnaeus, were common throughout the subalpine forest of Mauna Kea. However, Rattus rattus Linnaeus was rare, especially at higher elevations, whereas Mus musculus Linnaeus was more abundant at lower elevations. Main conclusions: Loxioides are concentrated in habitat that is distributed along a substantial gradient of elevation at least in part because food is available throughout the year and threats to food resources are less concentrated. To recover Loxioides elsewhere in its former range, habitats must be restored and alien threats reduced along extensive elevation gradients. Conservation along environmental gradients will likely benefit other Hawaiian birds that track the availability of food across landscapes or that have been stranded in the higher portions of their original ranges because of the greater impacts of alien diseases, predators, food competitors, and habitat stressors at lower elevations.
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Energy budget above a high-elevation subalpine forest in complex topography
Turnipseed, A.A.; Blanken, P.D.; Anderson, D.E.; Monson, Russell K.
2002-01-01
Components of the energy budget were measured above a subalpine coniferous forest over two complete annual cycles. Sensible and latent heat fluxes were measured by eddy covariance. Bowen ratios ranged from 0.7 to 2.5 in the summer (June-September) depending upon the availability of soil water, but were considerably higher (???3-6) during winter (December-March). Energy budget closure averaged better than 84% on a half-hourly basis in both seasons with slightly greater closure during the winter months. The energy budget showed a dependence on friction velocity (u*), approaching complete closure at u* values greater than 1 m s-1. The dependence of budget closure on u* explained why energy balance was slightly better in the winter as opposed to summer, since numerous periods of high turbulence occur in winter. It also explained the lower degree of energy closure (???10% less) during easterly upslope flow since these periods were characterized by low wind speeds (U < 4 m s-1) and friction velocities (u* < 0.5 m s-1). Co-spectral analysis suggests a shift of flux density towards higher frequencies under conditions where closure was obtained. It is suggested that low frequency contributions to the flux and advection were responsible for the lack of day-time energy budget closure. These effects were reduced at high friction velocities observed at our site. Our ability to close the energy budget at night was also highly dependent on friction velocity, approaching near closure (???90%) at u* values between 0.7 and 1.1 m s-1. Below this range, the airflow within the canopy becomes decoupled with the flow above. Above this range, insufficient temperature resolution of the sonic anemometer obscured the small temperature fluctuations, rendering measurements intractable. ?? 2002 Elsevier Science B.V. All rights reserved.
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Li, Liguang; Vogel, Jason; He, Zhenli; Zou, Xiaoming; Ruan, Honghua; Huang, Wei; Wang, Jiashe; Bianchi, Thomas S
2016-01-01
Forest soils play a critical role in the sequestration of atmospheric CO2 and subsequent attenuation of global warming. The nature and properties of organic matter in soils have an influence on the sequestration of carbon. In this study, soils were collected from representative forestlands, including a subtropical evergreen broad-leaved forest (EBF), a coniferous forest (CF), a subalpine dwarf forest (DF), and alpine meadow (AM) along an elevation gradient on Wuyi Mountain, which is located in a subtropical area of southeastern China. These soil samples were analyzed in the laboratory to examine the distribution and speciation of organic carbon (OC) within different size fractions of water-stable soil aggregates, and subsequently to determine effects on carbon sequestration. Soil aggregation rate increased with increasing elevation. Soil aggregation rate, rather than soil temperature, moisture or clay content, showed the strongest correlation with OC in bulk soil, indicating soil structure was the critical factor in carbon sequestration of Wuyi Mountain. The content of coarse particulate organic matter fraction, rather than the silt and clay particles, represented OC stock in bulk soil and different soil aggregate fractions. With increasing soil aggregation rate, more carbon was accumulated within the macroaggregates, particularly within the coarse particulate organic matter fraction (250-2000 μm), rather than within the microaggregates (53-250μm) or silt and clay particles (< 53μm). In consideration of the high instability of macroaggregates and the liability of SOC within them, further research is needed to verify whether highly-aggregated soils at higher altitudes are more likely to lose SOC under warmer conditions.
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Li, Liguang; Vogel, Jason; He, Zhenli; Zou, Xiaoming; Ruan, Honghua; Huang, Wei; Wang, Jiashe; Bianchi, Thomas S.
2016-01-01
Forest soils play a critical role in the sequestration of atmospheric CO2 and subsequent attenuation of global warming. The nature and properties of organic matter in soils have an influence on the sequestration of carbon. In this study, soils were collected from representative forestlands, including a subtropical evergreen broad-leaved forest (EBF), a coniferous forest (CF), a subalpine dwarf forest (DF), and alpine meadow (AM) along an elevation gradient on Wuyi Mountain, which is located in a subtropical area of southeastern China. These soil samples were analyzed in the laboratory to examine the distribution and speciation of organic carbon (OC) within different size fractions of water-stable soil aggregates, and subsequently to determine effects on carbon sequestration. Soil aggregation rate increased with increasing elevation. Soil aggregation rate, rather than soil temperature, moisture or clay content, showed the strongest correlation with OC in bulk soil, indicating soil structure was the critical factor in carbon sequestration of Wuyi Mountain. The content of coarse particulate organic matter fraction, rather than the silt and clay particles, represented OC stock in bulk soil and different soil aggregate fractions. With increasing soil aggregation rate, more carbon was accumulated within the macroaggregates, particularly within the coarse particulate organic matter fraction (250–2000 μm), rather than within the microaggregates (53–250μm) or silt and clay particles (< 53μm). In consideration of the high instability of macroaggregates and the liability of SOC within them, further research is needed to verify whether highly-aggregated soils at higher altitudes are more likely to lose SOC under warmer conditions. PMID:26964101
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Fire severity unaffected by spruce beetle outbreak in spruce-fir forests in southwestern Colorado.
Andrus, Robert A; Veblen, Thomas T; Harvey, Brian J; Hart, Sarah J
2016-04-01
Recent large and severe outbreaks of native bark beetles have raised concern among the general public and land managers about potential for amplified fire activity in western North America. To date, the majority of studies examining bark beetle outbreaks and subsequent fire severity in the U.S. Rocky Mountains have focused on outbreaks of mountain pine beetle (MPB; Dendroctonus ponderosae) in lodgepole pine (Pinus contorta) forests, but few studies, particularly field studies, have addressed the effects of the severity of spruce beetle (Dendroctonus rufipennis Kirby) infestation on subsequent fire severity in subalpine Engelmann spruce (Picea engelmannii) and subalpine fir (Abies lasiocarpa) forests. In Colorado, the annual area infested by spruce beetle outbreaks is rapidly rising, while MPB outbreaks are subsiding; therefore understanding this relationship is of growing importance. We collected extensive field data in subalpine forests in the eastern San Juan Mountains, southwestern Colorado, USA, to investigate whether a gray-stage (< 5 yr from outbreak to time of fire) spruce beetle infestation affected fire severity. Contrary to the expectation that bark beetle infestation alters subsequent fire severity, correlation and multivariate generalized linear regression analysis revealed no influence of pre-fire spruce beetle severity on nearly all field or remotely sensed measurements of fire severity. Findings were consistent across moderate and extreme burning conditions. In comparison to severity of the pre-fire beetle outbreak, we found that topography, pre-outbreak basal area, and weather conditions exerted a stronger effect on fire severity. Our finding that beetle infestation did not alter fire severity is consistent with previous retrospective studies examining fire activity following other bark beetle outbreaks and reiterates the overriding influence of climate that creates conditions conducive to large, high-severity fires in the subalpine zone of Colorado. Both bark beetle outbreaks and wildfires have increased autonomously due to recent climate variability, but this study does not support the expectation that post-beetle outbreak forests will alter fire severity, a result that has important implications for management and policy decisions.
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Yin, Xiuqin; Qiu, Lili; Jiang, Yunfeng; Wang, Yeqiao
2017-06-01
The understanding of patterns of vertical variation and diversity of flora and fauna along elevational change has been well established over the past century. However, it is unclear whether there is an elevational distribution pattern for soil fauna. This study revealed the diversity and spatial-temporal distribution of soil macrofauna communities in different vegetation zones from forest to alpine tundra along elevation of the Changbai Mountain, China. The abundance, richness, and Shannon-Wiener diversity index of soil macrofauna communities were compared in four distinguished vegetation zones including the coniferous and broadleaved mixed forest zone, the coniferous forest zone, the subalpine dwarf birch (Betula ermanii) forest zone, and the alpine tundra zone. Soil macrofauna were extracted in May, July, and September of 2009. In each season, the abundance and richness of the soil macrofauna decreased with the ascending elevation. The Shannon-Wiener diversity indices of the soil macrofauna were higher in the vegetation zones of lower elevation than of higher elevation. Significant differences were observed in the abundance, richness, and Shannon-Wiener diversity index for the studied vegetation zones. Soil macrofauna congregated mainly to the litter layer in the low-elevation areas and in the 0-5 cm soil layer of the higher elevation areas. The results emphasized that the diversity of soil macrofauna communities decreased as the elevation increased and possess the distinct characteristics of zonation in the mountain ecosystem. The diversity and distribution of soil macrofauna communities were influenced by mean annual precipitation, altitude, annual radiation quantity, and mean annual temperature. © The Authors 2017. Published by Oxford University Press on behalf of Entomological Society of America. All rights reserved. For Permissions, please email: journals.permissions@oup.com.
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Wildfires and geochemical change in a subalpine forest over the past six millennia
NASA Astrophysics Data System (ADS)
Leys, Bérangère; Higuera, Philip E.; McLauchlan, Kendra K.; Dunnette, Paul V.
2016-12-01
The frequency of large wildfires in western North America has been increasing in recent decades, yet the geochemical impacts of these events are poorly understood. The multidecadal timescales of both disturbance-regime variability and ecosystem responses make it challenging to study the effects of fire on terrestrial nutrient cycling. Nonetheless, disturbance-mediated changes in nutrient concentrations could ultimately limit forest productivity over centennial to millennial time scales. Here, we use a novel approach that combines quantitative elemental analysis of lake sediments using x-ray fluorescence to assess the geochemical impacts of high-severity fires in a 6200 year long sedimentary record from a small subalpine lake in Rocky Mountain National Park, Colorado, USA. Immediately after 17 high-severity fires, the sedimentary concentrations of five elements increased (Ti, Ca, K, Al, and P), but returned to pre-fire levels within three decades. Multivariate analyses indicate that erosion of weathered mineral material from the catchment is a primary mechanism though which high-severity fires impact element cycling. A longer-term trend in sediment geochemistry was also identified over millennial time scales. This decrease in the concentrations of six elements (Al, Si, K, Ti, Mn, and Fe) over the past 6200 years may have been due to a decreased rate of high-severity fires, long-term ecosystem development, or changes in precipitation regime. Our results indicate that high-severity fire events can determine elemental concentrations in subalpine forests. The degree of variability in geochemical response across time scales suggests that shifting rates of high-severity burning can cause significant changes in key rock-derived nutrients. To our knowledge, these results are the first to reveal repeated loss of rock-derived nutrients from the terrestrial ecosystem due to high-severity fires. Understanding the future of fire-prone coniferous forests requires further documentation and quantification of this important mechanism linking fire regimes and biogeochemical cycles.
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Soil propagule banks of ectomycorrhizal fungi share many common species along an elevation gradient.
Miyamoto, Yumiko; Nara, Kazuhide
2016-04-01
We conducted bioassay experiments to investigate the soil propagule banks of ectomycorrhizal (EM) fungi in old-growth forests along an elevation gradient and compared the elevation pattern with the composition of EM fungi on existing roots in the field. In total, 150 soil cores were collected from three forests on Mt. Ishizuchi, western Japan, and subjected to bioassays using Pinus densiflora and Betula maximowicziana. Using molecular analyses, we recorded 23 EM fungal species in the assayed propagule banks. Eight species (34.8 %) were shared across the three sites, which ranged from a warm-temperate evergreen mixed forest to a subalpine conifer forest. The elevation pattern of the assayed propagule banks differed dramatically from that of EM fungi on existing roots along the same gradient, where only a small proportion of EM fungal species (3.5 %) were shared across sites. The EM fungal species found in the assayed propagule banks included many pioneer fungal species and composition differed significantly from that on existing roots. Furthermore, only 4 of 23 species were shared between the two host species, indicating a strong effect of bioassay host identity in determining the propagule banks of EM fungi. These results imply that the assayed propagule bank is less affected by climate compared to EM fungal communities on existing roots. The dominance of disturbance-dependent fungal species in the assayed propagule banks may result in higher ecosystem resilience to disturbance even in old-growth temperate forests.
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Quantifying cambial activity of high-elevation conifers in the Great Basin, Nevada, USA
NASA Astrophysics Data System (ADS)
Ziaco, E.; Biondi, F.; Rossi, S.; Deslauriers, A.
2013-12-01
Understanding the physiological mechanisms that control the formation of tree rings provides the necessary biological basis for developing dendroclimatic reconstructions and dendroecological histories. Studies of wood formation in the Great Basin are now being conducted in connection with the Nevada Climate-ecohydrological Assessment Network (NevCAN), a recently established transect of valley-to-mountaintop instrumented stations in the Snake and Sheep Ranges of the Great Basin. Automated sensors record meteorological, soil, and vegetational variables at these sites, providing unique opportunities for ecosystem science, and are being used to investigate the ecological implications of xylogenesis. We present here an initial study based on microcores collected during summer 2013 from mountain and subalpine conifers (including Great Basin bristlecone pine, Pinus longaeva) growing on the west slope of Mt. Washington. Samples were taken from the mountain west (SM; 2810 m elevation) and the subalpine west (SS, 3355 m elevation) NevCAN sites on June 16th and 27th, 2013. The SS site was further subdivided in a high (SSH) and a low (SSL) group of trees, separated by about 10 m in elevation. Microscopic analyses showed the effect of elevation on cambial activity, as annual ring formation was more advanced at the lower (mountain) site compared to the higher (subalpine) one. At all sites cambium size showed little variations between the two sampling dates. The number of xylem cells in the radial enlargement phase decreased between the two sampling dates at the mountain site but increased at the subalpine site, confirming a delayed formation of wood at the higher elevations. Despite relatively high within-site variability, a general trend of increasing number of cells in the lignification phase was found at all sites. Mature cells were present only at the mountain site on June 27th. Spatial differences in the xylem formation process emerged at the species level and, within species, at different locations. In particular, bristlecone pines growing at SSL showed slightly more advanced tree-ring formation compared to those growing at SSH. These results highlight differences in xylogenesis related to elevation, a clear proxy for climate in this region, and further analyses will focus on understanding the climatic factors that drive wood formation in Great Basin conifers. Fig.1 Boxplot summary of cell types found at the study sites during two sampling dates.
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NASA Astrophysics Data System (ADS)
Macias-Fauria, M.; Johnson, E. A.; Forbes, B. C.; Willis, K. J.
2013-12-01
In cold ecosystems such as sub-alpine forests and forest-tundra, vegetation geographical ranges are expected to expand upward/northward in a warmer world. Such moving fronts have been predicted to 1) decrease the remaining alpine area in mountain systems, increasing fragmentation and extinction risk of many alpine taxa, and 2) fundamentally modify the energy budget of newly afforested areas, enhancing further regional warming due to a reduction in albedo. The latter is particularly significant in the forest-tundra, where changes over large regions can have regional-to-global effects on climate. An integral part of the expected range shifts is their velocity. Whereas range shifts across thermal gradients can theoretically be fast in an elevation gradient relative to climate velocity (i.e. rate of climate change) due to the short distances involved, large lags are expected over the flat forest-tundra. Mountain regions have thus been identified as buffer areas where species can track climate change, in opposition to flat terrain where climate velocity is faster. Thus, much shorter time-to-equilibrium are expected for advancing upslope sub-alpine forest than for advancing northern boreal forest. We contribute to this discussion by showing two mechanisms that might largely alter the above predictions in opposite directions: 1) In mountain regions, terrain heterogeneity not only allows for slower climate velocities, but slope processes largely affect the advance of vegetation. Indeed, such mechanisms can potentially reduce the climatic signal in vegetation distribution limits (e.g. treeline), precluding it from migrating to climatically favourable areas - since these areas occur in geologically unfavourable ones. Such seemingly local control to species range shifts was found to reduce the climate-sensitive treeline areas in the sub-alpine forest of the Canadian Rocky Mountains to ~5% at a landscape scale, fundamentally altering the predictions of vegetation response to climate warming in the region (Macias-Fauria & Johnson 20013, PNAS). 2) In the low arctic tundra, un-treed to treed landscapes have sprouted in several parts of the tundra in a matter of decades, as opposed to the previously predicted response times of several centuries for boreal forest to advance to its new climate optimum (migrational lags). This takes place not through very rapid moving fronts, but through phenotypic responses of extant vegetation with highly flexible life forms, such as woody deciduous shrubs (Salix, Alnus, Betula). The resulting vegetation response creates strong energy feedbacks while at the same time potentially further reduces the speed of northward displacement of the boreal forest, that has to compete with a new treed ecosystem (Macias-Fauria et al. 2012, Nature Climate Change). In conclusion, control of rates of migration by factors other than climate in mountain systems can largely reduce the ability of vegetation to track climate change, and emergence of structurally novel ecosystems in low arctic tundra might largely alter current predictions based on climate response of vegetation, by accelerating ecosystem change and reducing migrational rates simultaneously.
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Berryman, Erin Michele; Ryan, Michael G.; Bradford, John B.; Hawbaker, Todd J.; Birdsey, R.
2016-01-01
In forests, total belowground carbon (C) flux (TBCF) is a large component of the C budget and represents a critical pathway for delivery of plant C to soil. Reducing uncertainty around regional estimates of forest C cycling may be aided by incorporating knowledge of controls over soil respiration and TBCF. Photosynthesis, and presumably TBCF, declines with advancing tree size and age, and photosynthesis increases yet C partitioning to TBCF decreases in response to high soil fertility. We hypothesized that these causal relationships would result in predictable patterns of TBCF, and partitioning of C to TBCF, with natural variability in leaf area index (LAI), soil nitrogen (N), and tree height in subalpine forests in the Rocky Mountains, USA. Using three consecutive years of soil respiration data collected from 22 0.38-ha locations across three 1-km2 subalpine forested landscapes, we tested three hypotheses: (1) annual soil respiration and TBCF will show a hump-shaped relationship with LAI; (2) variability in TBCF unexplained by LAI will be related to soil nitrogen (N); and (3) partitioning of C to TBCF (relative to woody growth) will decline with increasing soil N and tree height. We found partial support for Hypothesis 1 and full support for Hypotheses 2 and 3. TBCF, but not soil respiration, was explained by LAI and soil N patterns (r2 = 0.49), and the ratio of annual TBCF to TBCF plus aboveground net primary productivity (ANPP) was related to soil N and tree height (r2 = 0.72). Thus, forest C partitioning to TBCF can vary even within the same forest type and region, and approaches that assume a constant fraction of TBCF relative to ANPP may be missing some of this variability. These relationships can aid with estimates of forest soil respiration and TBCF across landscapes, using spatially explicit forest data such as national inventories or remotely sensed data products.
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Woody encroachment and soil carbon stocks in subalpine areas in the Central Spanish Pyrenees.
Nadal-Romero, E; Otal-Laín, I; Lasanta, T; Sánchez-Navarrete, P; Errea, P; Cammeraat, E
2018-05-01
Woody encroachment has been an ongoing process in the subalpine belt of Mediterranean mountains, after land abandonment, the disappearance of the transhumant system and the decrease of the livestock number. The main objectives of this study were: (i) to identify land use/land cover (LULC) changes from 1956 to 2015, and (ii) to investigate the effects of LULC changes in physical and chemical soil properties and soil organic carbon (SOC) and nitrogen (N) stocks. It is hypothesized that woody encroachment in the subalpine belt may lead to significant changes in soil properties, and will generate an increase in the SOC stocks. A land use gradient was identified in the subalpine belt of the Central Spanish Pyrenees: (i) subalpine grasslands, (ii) shrublands, (iii) young forests, and (iv) old forests. Mineral soil samples were collected every 10 cm, down to 40 cm, at three points per each LULC and a total of 48 samples were analyzed. The results showed that (i) woody encroachment has occurred from 1956 to 2015 due to the expansion of coniferous forests and shrublands (at the expense of grasslands), (ii) land cover and soil depth had significant effects on soil properties (except for pH), being larger in the uppermost 0-10 cm depth, (iii) SOC and N contents and stocks were higher in the grassland sites, and (iv) the woody encroachment process initially produced a decrease in the SOC stocks (shrublands), but no differences were observed considering the complete soil profile between grasslands and young and old forests. Further studies, describing SOC stabilization and quantifying above-ground carbon (shrub and tree biomass) are required. Copyright © 2018 Elsevier B.V. All rights reserved.
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Community structure and elevational diversity patterns of soil Acidobacteria.
Zhang, Yuguang; Cong, Jing; Lu, Hui; Li, Guangliang; Qu, Yuanyuan; Su, Xiujiang; Zhou, Jizhong; Li, Diqiang
2014-08-01
Acidobacteria is one of the most dominant and abundant phyla in soil, and was believed to have a wide range of metabolic and genetic functions. Relatively little is known about its community structure and elevational diversity patterns. We selected four elevation gradients from 1000 to 2800 m with typical vegetation types of the northern slope of Shennongjia Mountain in central China. The vegetation types were evergreen broadleaved forest, deciduous broadleaved forest, coniferous forest and sub-alpine shrubs. We analyzed the soil acidobacterial community composition, elevational patterns and the relationship between Acidobacteria subdivisions and soil enzyme activities by using the 16S rRNA meta-sequencing technique and multivariate statistical analysis. The result found that 19 known subdivisions as well as an unclassified phylotype were presented in these forest sites, and Subdivision 6 has the highest number of detectable operational taxonomic units (OTUs). A significant single peak distribution pattern (P<0.05) between the OTU number and the elevation was observed. The Jaccard and Bray-Curtis index analysis showed that the soil Acidobacteria compositional similarity significantly decreased (P<0.01) with the increase in elevation distance. Mantel test analysis showed the most of the soil Acidobacteria subdivisions had the significant relationship (P<0.01) with different soil enzymes. Therefore, soil Acidobacteria may be involved in different ecosystem functions in global elemental cycles. Partial Mantel tests and CCA analysis showed that soil pH, soil temperature and plant diversity may be the key factors in shaping the soil Acidobacterial community structure. Copyright © 2014. Published by Elsevier B.V.
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Silvics of whitebark pine (Pinus albicaulis)
Stephen F. Arno; Raymond J. Hoff
1989-01-01
Whitebark pine (Pinus albicaulis) is a long-lived tree inhabiting the upper subalpine forest and timberline zone on high mountains of Western North America. The species' habitat, life history, growth and yield, mortality factors, special uses, and genetics are described.
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NASA Astrophysics Data System (ADS)
Dailey, K. R.; Hughes, H.; Williams, M. W.
2015-12-01
Geochemical surface and groundwater data were used to examine groundwater recharge dynamics and hydrologic connectivity in the dominantly subalpine Como Creek headwater catchment within the Boulder Creek Watershed in the Colorado Front Range. Streamwater chemistry along an elevational gradient of Como Creek showed strong responses to variations in precipitation inputs spanning 2011-2014. Elevation effects on δ18O were apparent, with more depleted values indicative of snowmelt influence observed at the higher elevation sites. Results from one-way ANOVA indicated that the highest elevation stream site, situated right below treeline, was significantly different from the lower three sites with regards to DOC, δ18O, and Ca2+ (p < 0.05) over May-October 2011-2014. Additionally, the second highest site in elevation was found to be significantly different from all other sites with respect to Ca2+ concentrations. Soil moisture sensor and geochemical data from soil tension lysimeters co-located with subalpine groundwater wells revealed a disconnect between soil and surface water chemistry during snowmelt and that of deeper, underlying groundwater. The initial results of this study provide insight on where groundwater recharge and discharge may be occurring in the catchment and help us to evaluate the large September 2013 rain event in the Colorado Front Range, a once in a 1000 year event. Water isotopes were enriched, Ca2+ decreased, and DOC was enriched, showing that new event water had flowed through near-surface soils but not deeper, recharging groundwater, with all values returning to normal within six months. The event was also observable compared to a long-term geochemical and stream stage record at the stream site near the catchment outlet, with the most enriched δ18O value on record of -13.41‰ corresponding to the flood. Remaining questions concerning groundwater dynamics in the catchment include constraining the tradeoffs between forest ET, groundwater recharge, and streamflow under a warming climate in order to forecast future water yield from the catchment. Further research quantifying where and when groundwater is recharging in the catchment will be conducted to determine how hydrological resilience of the catchment will change under varying climatic conditions.
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Calder, W John; Horn, Kevin J; St Clair, Samuel B
2011-06-01
Disturbance patterns strongly influence plant community structure. What remains less clear, particularly at a mechanistic level, is how changes in disturbance cycles alter successional outcomes in plant communities. There is evidence that fire suppression is resulting in longer fire return intervals in subalpine forests and that these lengthened intervals increase competitive interactions between aspen and conifer species. We conducted a field and greenhouse study to compare photosynthesis, growth and defense responses of quaking aspen and subalpine fir regeneration under light reductions and shifts in soil chemistry that occur as conifers increase in dominance. The studies demonstrated that aspen regeneration was substantially more sensitive to light and soil resource limitations than that of subalpine fir. For aspen, light reductions and/or shifts in soil chemistry limited height growth, biomass gain, photosynthesis and the production of defense compounds (phenolic glycosides and condensed tannins). Biomass gain and phenolic glycoside concentrations were co-limited by light reduction and changes in soil chemistry. In contrast, subalpine fir seedlings tended to be more tolerant of low light conditions and showed no sensitivity to changes in soil chemistry. Unlike aspen, subalpine fir increased its root to shoot ratio on conifer soils, which may partially explain its maintenance of growth and defense. The results suggest that increasing dominance of conifers in subalpine forests alters light conditions and soil chemistry in a way that places greater physiological and growth constraints on aspen than subalpine fir, with a likely outcome being more successful recruitment of conifers and losses in aspen cover.
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Wang, Guobing; Zhou, Yan; Xu, Xia; Ruan, Honghua; Wang, Jiashe
2013-01-01
Soil organic carbon (SOC) actively participates in the global carbon (C) cycle. Despite much research, however, our understanding of the temperature sensitivity of soil organic carbon (SOC) mineralization is still very limited. To investigate the responses of SOC mineralization to temperature, we sampled surface soils (0-10 cm) from evergreen broad-leaf forest (EBF), coniferous forest (CF), sub-alpine dwarf forest (SDF), and alpine meadow (AM) along an elevational gradient in the Wuyi Mountains, China. The soil samples were incubated at 5, 15, 25, and 35°C with constant soil moisture for 360 days. The temperature sensitivity of SOC mineralization (Q(10)) was calculated by comparing the time needed to mineralize the same amount of C at any two adjacent incubation temperatures. Results showed that the rates of SOC mineralization and the cumulative SOC mineralized during the entire incubation significantly increased with increasing incubation temperatures across the four sites. With the increasing extent of SOC being mineralized (increasing incubation time), the Q(10) values increased. Moreover, we found that both the elevational gradient and incubation temperature intervals significantly impacted Q(10) values. Q(10) values of the labile and recalcitrant organic C linearly increased with elevation. For the 5-15, 15-25, and 25-35°C intervals, surprisingly, the overall Q(10) values for the labile C did not decrease as the recalcitrant C did. Generally, our results suggest that subtropical forest soils may release more carbon than expected in a warmer climate.
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Solar dimming above temperate forests and its impact on local climate
NASA Astrophysics Data System (ADS)
Tudoroiu, M.; Genesio, L.; Gioli, B.; Schume, H.; Knohl, A.; Brümmer, C.; Miglietta, F.
2018-06-01
Vegetation has a substantial impact on the local climate. Land cover changes through afforestation or deforestation can amplify or mitigate climate warming by changes in biophysical and biogeochemical mechanisms. In the montane to subalpine area of the Eastern Alps in Europe, where forests have constantly expanded in the last four decades, data of meteorological stations show a consistent reduction in incoming global radiation for the period 2000–2015. To assess the potential role of forests in contributing to such a reduction, three site pairs in Central Europe with neighbouring forest and non-forest sites were analysed. In all the pairs, a lower amount of incoming radiation was recorded at the forest site. When biophysical mechanisms such as albedo, surface roughness and Bowen ratio changes were modelled together with changes in global radiation, the total radiative forcing accounted for a rate of change in air temperature was equal to 0.032 °C ± 0.01 °C per Wm‑2. These results suggest that local climate is influenced by land cover change through afforestation both via albedo and radiation feedbacks but also by means of indirect biophysical and species-dependent mechanisms. The data obtained for the site pairs in Central Europe are finally discussed to infer the occurrence of similar forest-driven effects in the Eastern Alps which may explain part of the solar dimming observed in high elevation weather stations.
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Kubo, Joshua S.; Torgersen, Christian E.; Bolton, Susan M.; Weekes, Anne A.; Gara, Robert I.
2013-01-01
1. Aquatic habitats and biotic assemblages in subalpine headwaters are sensitive to climate and human impacts. Understanding biotic responses to such perturbations and the contribution of high-elevation headwaters to riverine biodiversity requires the assessment of assemblage composition among habitat types. We compared aquatic insect assemblages among headwater stream segment types in relict glaciated subalpine basins in Mt. Rainier National Park, Washington, USA. 2. Aquatic insects were collected during summer and autumn in three headwater basins. In each basin, three different stream segment types were sampled: colluvial groundwater sources, alluvial lake inlets, and cascade-bedrock lake outlets. Ward's hierarchical cluster analysis revealed high β diversity in aquatic insect assemblages, and non-metric multidimensional scaling indicated that spatial and temporal patterns in assemblage composition differed among headwater stream segment types. Aquatic insect assemblages showed more fidelity to stream segment types than to individual basins, and the principal environmental variables associated with assemblage structure were temperature and substrate. 3. Indicator species analyses identified specific aquatic insects associated with each stream segment type. Several rare and potentially endemic aquatic insect taxa were present, including the recently described species, Lednia borealis (Baumann and Kondratieff). 4. Our results indicate that aquatic insect assemblages in relict glaciated subalpine headwaters were strongly differentiated among stream segment types. These results illustrate the contribution of headwaters to riverine biodiversity and emphasise the importance of these habitats for monitoring biotic responses to climate change. Monitoring biotic assemblages in high-elevation headwaters is needed to prevent the potential loss of unique and sensitive biota.
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Eagles-Smith, Collin A.; Herring, Garth; Johnson, Branden L.; Graw, Rick
2016-01-01
Remote high-elevation lakes represent unique environments for evaluating the bioaccumulation of atmospherically deposited mercury through freshwater food webs, as well as for evaluating the relative importance of mercury loading versus landscape influences on mercury bioaccumulation. The increase in mercury deposition to these systems over the past century, coupled with their limited exposure to direct anthropogenic disturbance make them useful indicators for estimating how changes in mercury emissions may propagate to changes in Hg bioaccumulation and ecological risk. We evaluated mercury concentrations in resident fish from 28 high-elevation, sub-alpine lakes in the Pacific Northwest region of the United States. Fish total mercury (THg) concentrations ranged from 4 to 438 ng/g wet weight, with a geometric mean concentration (±standard error) of 43 ± 2 ng/g ww. Fish THg concentrations were negatively correlated with relative condition factor, indicating that faster growing fish that are in better condition have lower THg concentrations. Across the 28 study lakes, mean THg concentrations of resident salmonid fishes varied as much as 18-fold among lakes. We used a hierarchal statistical approach to evaluate the relative importance of physiological, limnological, and catchment drivers of fish Hg concentrations. Our top statistical model explained 87% of the variability in fish THg concentrations among lakes with four key landscape and limnological variables: catchment conifer density (basal area of conifers within a lake's catchment), lake surface area, aqueous dissolved sulfate, and dissolved organic carbon. Conifer density within a lake's catchment was the most important variable explaining fish THg concentrations across lakes, with THg concentrations differing by more than 400 percent across the forest density spectrum. These results illustrate the importance of landscape characteristics in controlling mercury bioaccumulation in fish.
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Eagles-Smith, Collin A; Herring, Garth; Johnson, Branden; Graw, Rick
2016-05-01
Remote high-elevation lakes represent unique environments for evaluating the bioaccumulation of atmospherically deposited mercury through freshwater food webs, as well as for evaluating the relative importance of mercury loading versus landscape influences on mercury bioaccumulation. The increase in mercury deposition to these systems over the past century, coupled with their limited exposure to direct anthropogenic disturbance make them useful indicators for estimating how changes in mercury emissions may propagate to changes in Hg bioaccumulation and ecological risk. We evaluated mercury concentrations in resident fish from 28 high-elevation, sub-alpine lakes in the Pacific Northwest region of the United States. Fish total mercury (THg) concentrations ranged from 4 to 438 ng/g wet weight, with a geometric mean concentration (±standard error) of 43 ± 2 ng/g ww. Fish THg concentrations were negatively correlated with relative condition factor, indicating that faster growing fish that are in better condition have lower THg concentrations. Across the 28 study lakes, mean THg concentrations of resident salmonid fishes varied as much as 18-fold among lakes. We used a hierarchal statistical approach to evaluate the relative importance of physiological, limnological, and catchment drivers of fish Hg concentrations. Our top statistical model explained 87% of the variability in fish THg concentrations among lakes with four key landscape and limnological variables: catchment conifer density (basal area of conifers within a lake's catchment), lake surface area, aqueous dissolved sulfate, and dissolved organic carbon. Conifer density within a lake's catchment was the most important variable explaining fish THg concentrations across lakes, with THg concentrations differing by more than 400 percent across the forest density spectrum. These results illustrate the importance of landscape characteristics in controlling mercury bioaccumulation in fish. Published by Elsevier Ltd.
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Smithers, Brian V; North, Malcolm P; Millar, Constance I; Latimer, Andrew M
2018-02-01
In response to climate warming, subalpine treelines are expected to move up in elevation since treelines are generally controlled by growing season temperature. Where treeline is advancing, dispersal differences and early life stage environmental tolerances are likely to affect how species expand their ranges. Species with an establishment advantage will colonize newly available habitat first, potentially excluding species that have slower establishment rates. Using a network of plots across five mountain ranges, we described patterns of upslope elevational range shift for the two dominant Great Basin subalpine species, limber pine and Great Basin bristlecone pine. We found that the Great Basin treeline for these species is expanding upslope with a mean vertical elevation shift of 19.1 m since 1950, which is lower than what we might expect based on temperature increases alone. The largest advances were on limber pine-dominated granitic soils, on west aspects, and at lower latitudes. Bristlecone pine juveniles establishing above treeline share some environmental associations with bristlecone adults. Limber pine above-treeline juveniles, in contrast, are prevalent across environmental conditions and share few environmental associations with limber pine adults. Strikingly, limber pine is establishing above treeline throughout the region without regard to site characteristic such as soil type, slope, aspect, or soil texture. Although limber pine is often rare at treeline where it coexists with bristlecone pine, limber pine juveniles dominate above treeline even on calcareous soils that are core bristlecone pine habitat. Limber pine is successfully "leap-frogging" over bristlecone pine, probably because of its strong dispersal advantage and broader tolerances for establishment. This early-stage dominance indicates the potential for the species composition of treeline to change in response to climate change. More broadly, it shows how species differences in dispersal and establishment may result in future communities with very different specific composition. © 2017 John Wiley & Sons Ltd.
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Ecological transition in Arizona's subalpine and montane grasslands
Michael R. White
2000-01-01
Important components of Southwest forest ecosystem are subalpine and montane grassland communities, Grassland communities provide habitat diversity for wildlife, forage for domestic livestock and wildlife, and contribute to the visual quality of an area. The objectives of this research were to determine if: 1) vegetation attributes and soil-surface cover variables of...
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A half century of change in alpine treeline patterns at Glacier National Park, Montana, U.S.A.
Klasner, F.L.; Fagre, D.B.
2002-01-01
Using sequential aerial photography, we identified changes in the spatial distribution of subalpine fir (Abies lasiocarpa) habitat at the alpine treeline ecotone. Six 40-ha study sites in the McDonald Creek drainage of Glacier National Park contained subalpine fir forests that graded into alpine tundra. Over a 46-yr period, altitudinal changes in the location of alpine treeline ecotone were not observed. However, over this 46-yr period the area of krummholz, patch-forest, and continuous canopy forest increased by 3.4%, and tree density increased within existing patches of krummholz and patch-forest. Change in subalpine fir vegetation patterns within 100 m of trails was also compared to areas without trails. Within 100 m of trails, the number of small, discrete krummholz stands increased compared to areas without trails, but there was no significant change in total krummholz area. We used historical terrestrial photography to expand the period (to 70 yr) considered. This photography supported the conclusions that a more abrupt ecotone transition developed from forest to tundra at alpine treeline, that tree density within forested areas increased, and that krummholz became fragmented along trails. This local assessment of fine-grained change in the alpine treeline ecotone provides a comparative base for looking at ecotone change in other mountain regions throughout the world.
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Air-Pollution-Mediated Changes in Alpine Ecosystems and Ecotones.
Rusek, Josef
1993-08-01
Soil biological parameters (e.g., Collembola), soil types, soil chemical parameters (pH, humus substances), and plant communities were studied in different ecosystems and ecotones in alpine, subalpine, and spruce forest zones in the Tatra National Park, Slovak Republic. The preliminary, selected data, based on a long-term research program, showed a high sensitivity of some alpine ecotones and ecosystems to long-distance transported acid deposits. The changes in different ecosystem parameters since 1977 were more extensive in alpine grasslands on limestone than on granite. The greatest soil pH decrease was in the plant communities Festucetum versicoloris (-1.5 pH), Geranio-Alchemilletum crinitae (-1.32 pH), and Saxifragetum perdurantis (-1.25 pH), which are restricted to places with snow accumulation and water runoff gullies. In these ecosystems the greatest changes occurred in the leaching of humus substances. Some formerly less abundant and rare soil animals restricted to acid bedrock became dominant in some ecosystems on limestone as well as on granite; other formerly dominant species disappeared from the entire study area (e.g., Folsomia alpina). The aerial extent of some ecosystems changed substantially since 1977, and their surrounding ecotones moved into the space formerly occupied by one of the adjacent ecosystems. These changes are detectable by remote-sensing methods. In Central European mountains, strongly affected by global and regional industrial air pollution (e.g., Krusne Hory, Krkonose, Beskydy), spruce forests started to die back from higher to lower mountain elevations. The effects of air pollution on alpine and subalpine vegetation were not studied there. Strong alterations in alpine ecosystems and ecotones were detected by the author during long-term studies in the High Tatra Mountains, and I suggest that subalpine and mountain forest belts will be affected here in the near future as they were in the more polluted Central European mountains. The ecosystems and ecotones in higher alpine zones are likely to be affected earlier than the ecosystems at lower altitudes. Detection of ecosystem alteration in the alpine zone may be used for prediction of acidification processes and global change in ecosystems at lower altitudes. The consequences of global climate change are predictable by monitoring changes in the extent of some ecosystems located in discrete mountain geomorphological units (e.g., karstic sinkholes, water runoff gullies, wind shadows, ridges exposed to wind, etc.) and ecotones among them because of their dependence on duration of snow cover, water supply, wind and frost exposure, and other abiotic and biotic factors. © 1993 by the Ecological Society of America.
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Chen, Ya; Yang, Wan Qin; Wu, Fu Zhong; Yang, Fan; Lan, Li Ying; Liu, Yu Wei; Guo, Cai Hong; Tan, Bo
2017-10-01
In order to understand the diversity of soil nematodes in the subalpine/alpine forests of the eastern Qinghai-Tibet Plateau, soil nematodes in the primary forest, mixed forest and secondary forest of Abies faxoniana were extracted by elutriation and sugar-centrifugation method in July 2015, and the composition and structure characteristics of soil nematode communities were studied in the three forests at different altitudes. A total of 37950 soil nematodes were collected, which belonged to 20 families and 27 genera, and the mean density was 4217 ind·100 g -1 dry soil. Filenchus was the dominant genus in the primary forest, and Filenchus and Pararotylenchus in the mixed forest and secondary forest, respectively. The individual number of each dominant genus was significantly affected by forest type. All nematode individuals were classified into the four trophic groups of bacterivores, fungivores, plant-parasites and omnivore-predators. The fungivores were dominant in the primary and secondary forest and the bacterivores in the mixed forest. The number of soil nematode c-p (colonizer-persister) groups of c-p 1, c-p 2, c-p 3 and c-p 4 accounted for 6.1%, 51.1%, 30.0% and 12.7% of the total nematode abundance, respectively. The maturity index (MI), the total maturity index (∑MI) and the plant parasitic index (PPI) of soil nematodes decreased gradually with the increase of altitude. The nematode channel ratio in the mixed forest was higher than 0.5, but that in the primary forest and secondary forest was below 0.5. The forest type significantly affected the soil nematode maturity index and channel ratio, but the forest type, soil layer and their interaction had no significant effect on the diversity index. There were obvious diffe-rences in the composition, nutrient structure and energy flow channel of soil nematodes in the subalpine/alpine forests of western Sichuan, providing an important reference for understanding the function of soil nematodes in soil processes of this region.
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Pan, Xin-li; Lin, Bo; Liu, Qing
2008-08-01
To investigate the effects of elevated temperature on the soil organic carbon content, soil respiration rate, and soil enzyme activities in subalpine Picea asperata plantations in western Sichuan Province of China, a simulation study was conducted in situ with open-top chambers from November 2005 to July 2007. The results showed that under elevated temperature, the mean air temperature and soil temperature were 0.42 degrees C and 0.25 degrees C higher than the control, respectively. In the first and the second year, the increased temperature had somewhat decreasing effects on the soil organic carbon and the C/N ratio at the soil depths of 0-10 cm and 10-20 cm. In the first year the soil organic carbon and the C/N ratio in 0-10 cm soil layer decreased by 8.69%, and 8.52%, respectively; but in the second year, the decrements were lesser. Soil respiration rate was significantly enhanced in the first year of warming, but had no significant difference with the control in the second year. In the first year of warming, the activities of soil invertase, polyphenol oxidase, catalase, protease, and urease increased, and the invertase and polyphenol oxidase activities in 0-10 cm soil layer were significantly higher than the control. In the second year of warming, the activities of invertase, protease and urease still had an increase, but those of catalase and polyphenol oxidase had a downtrend, compared with the control.
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NASA Astrophysics Data System (ADS)
Dobrowski, S. Z.; Greenberg, J. A.; Schladow, G.
2006-12-01
There is evidence from the Sierra Nevada that sub-alpine and alpine environments are currently experiencing landscape-mediated changes in growth and recruitment due to recent climate change. Understanding the biophysical controls of forest structure, growth, and recruitment in these environments is critical for interpreting and predicting the direction and magnitude of biotic responses to climate shift. We examined the abiotic controls of forest biomass within a 305 km2 region of the Carson Range on the eastern shore of Lake Tahoe, CA USA using estimates of forest structure and biophysical drivers developed continuously over the landscape. The study area ranged from 1900 m to 3400 m a.s.l. and encompassed montane, sub-alpine, and alpine environments. From hyperspatial optical imagery (IKONOS), we derived per-tree positions and crown sizes using a template matching approach applied to a pre-classified image of sunlit and shadowed vegetation pixels. From this remote sensing derived stem map, we calculated plot-level estimates of stem density, tree cover and average crown size. Additionally, we developed high resolution (30 m) estimates of climate variables within the study area using meteorological station data, topographic data, and a combination of empirical and mechanistic modeling approaches. From these climate surfaces, digital elevation data, and soil survey data, we derived estimates of direct and indirect biophysical drivers including heat loading, reference evapotranspiration, water deficit, solar radiation, topographic convergence, soil depth, and soil water holding capacity. Using these data sets, we conducted a regression tree analysis with stem density, tree cover, and average tree size as response and biophysical drivers as predictors. Trees were fit using half of the dataset randomly sampled (168,000 samples) and pruned using cost-complexity pruning based on 10-fold cross- validation. Predictions from pruned trees were then assessed against the hold-out data. Preliminary results from this analysis suggest that: 1) the relative importance and dependencies of biophysical drivers on forest structure are contingent upon the position of these forests along gradients of a limiting resource, 2) stem density shows a stronger dependence on water availability than tree size and 3) that the predictive power of abiotic variables are limited with our best models accounting for only 36-40 percent of the variance in the response. These results suggest that the response of forest structure to climate change may be highly idiosyncratic and difficult to predict using abiotic drivers alone.
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Forest dynamics in a forest-tundra ecotone, Medicine Bow Mountains, Wyoming
Christopher J. Earle
1993-01-01
The alpine timberline in much of western North America is characterized by a structurally complex transition from subalpine forest to alpine tundra, the forest-tundra ecotone. Trees within the ecotone are typically arrayed across the landscape within clumps or "ribbon forests," elongated strips oriented perpendicular to the prevailing winds. This study...
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Glacial and Quaternary geology of the northern Yellowstone area, Montana and Wyoming
Pierce, Kenneth L.; Licciardi, Joseph M.; Krause, Teresa R.; Whitlock, Cathy
2014-01-01
This field guide focuses on the glacial geology and paleoecology beginning in the Paradise Valley and progressing southward into northern Yellowstone National Park. During the last (Pinedale) glaciation, the northern Yellowstone outlet glacier flowed out of Yellowstone Park and down the Yellowstone River Valley into the Paradise Valley. The field trip will traverse the following Pinedale glacial sequence: (1) deposition of the Eightmile terminal moraines and outwash 16.5 ± 1.4 10Be ka in the Paradise Valley; (2) glacial recession of ~8 km and deposition of the Chico moraines and outwash 16.1 ± 1.7 10Be ka; (3) glacial recession of 45 km to near the northern Yellowstone boundary and moraine deposition during the Deckard Flats readjustment 14.2 ± 1.2 10Be ka; and (4) glacial recession of ~37 km and deposition of the Junction Butte moraines 15.2 ± 1.3 10Be ka (this age is a little too old based on the stratigraphic sequence). Yellowstone's northern range of sagebrush-grasslands and bison, elk, wolf, and bear inhabitants is founded on glacial moraines, sub-glacial till, and outwash deposited during the last glaciation. Floods released from glacially dammed lakes and a landslide-dammed lake punctuate this record. The glacial geologic reconstruction was evaluated by calculation of basal shear stress, and yielded the following values for flow pattern in plan view: strongly converging—1.21 ± 0.12 bars (n = 15); nearly uniform—1.04 ± 0.16 bars (n = 11); and strongly diverging—0.84 ± 0.14 bars (n = 16). Reconstructed mass balance yielded accumulation and ablation each of ~3 km3/yr, with glacial movement near the equilibrium line altitude dominated by basal sliding. Pollen and charcoal records from three lakes in northern Yellowstone provide information on the postglacial vegetation and fire history. Following glacial retreat, sparsely vegetated landscapes were colonized first by spruce parkland and then by closed subalpine forests. Regional fire activity increased significantly with the development of closed subalpine forests as a result of increased fuel biomass and warmer summers. Warm dry conditions prevailed at low elevations during the early Holocene, as indicated by the presence of steppe and open mixed conifer forest. At the same time, closed subalpine forests with low fire frequency were present at higher elevations, suggesting relatively wet summer conditions. Douglas fir populations expanded throughout northern Yellowstone in the middle Holocene as a result of effectively drier conditions than before, and a decline of mesophytic plant taxa during the late Holocene imply continued drying, even though fire frequency decreased in recent millennia.
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Zhu, Jianxiao; Yang, Wanqin; He, Xinhua
2013-01-01
Relationships between abiotic (soil temperature and number of freeze-thaw cycles) or biotic factors (chemical elements, microbial biomass, extracellular enzymes, and decomposer communities in litter) and litter decomposition rates were investigated over two years in subalpine forests close to the Qinghai-Tibet Plateau in China. Litterbags with senescent birch, fir, and spruce leaves were placed on the forest floor at 2,704 m, 3,023 m, 3,298 m, and 3,582 m elevation. Results showed that the decomposition rate positively correlated with soil mean temperature during the plant growing season, and with the number of soil freeze-thaw cycles during the winter. Concentrations of soluble nitrogen (N), phosphorus (P) and potassium (K) had positive effects but C:N and lignin:N ratios had negative effects on the decomposition rate (k), especially during the winter. Meanwhile, microbial biomass carbon (MBC), N (MBN), and P (MBP) were positively correlated with k values during the first growing season. These biotic factors accounted for 60.0% and 56.4% of the variation in decomposition rate during the winter and the growing season in the first year, respectively. Specifically, litter chemistry (C, N, P, K, lignin, C:N and lignin:N ratio) independently explained 29.6% and 13.3%, and the microbe-related factors (MBC, MBN, MBP, bacterial and fungal biomass, sucrase and ACP activity) explained 22.9% and 34.9% during the first winter and the first growing season, respectively. We conclude that frequent freeze-thaw cycles and litter chemical properties determine the winter decomposition while microbe-related factors play more important roles in determining decomposition in the subsequent growing season. PMID:23620803
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Qing Xu; Harbin Li; Jiquan Chen; Jiquan Cheng; Xiaoli Cheng; Shirong Liu; Shuqing An
2011-01-01
Determination of water sources of plant species in a community is critical for understanding the hydrological processes and their importance in ecosystem functions. Such partitioning of plant xylem water into specific sources (i.e. precipitation, groundwater) can be achieved by analyzing deuterium isotopic composition (δD) values for source waters. A subalpine dark...
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Historical land-cover classification for conservation and management in Hawaiian subalpine drylands
James R. Kellner; Gregory P. Asner; Susan Cordell; Jarrod M. Thaxton; Kealoha M. Kinney; Ty Kennedy-Bowdoin; David E. Knapp; Erin F. Questad; Stephen Ambagis
2012-01-01
We used aerial photography from 1954 and airborne LiDAR and imaging spectroscopy from 2008 to infer changes in extent and location of tallstature woody vegetation in 127 km2 of subalpine dry forest on the island of Hawaii (Pohakuloa Training Area), and to identify 25.8 km2 of intact woody vegetation for restoration and...
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NASA Astrophysics Data System (ADS)
Griepentrog, Marco; Bodé, Samuel; Boudin, Mathieu; Dercon, Gerd; Doetterl, Sebastian; Matulanya, Machibya; Msigwa, Anna; Vermeir, Pieter; Boeckx, Pascal
2017-04-01
Terrestrial ecosystems are strongly influenced by climate change and soils are key compartments of the global carbon (C) cycle in terms of their potential to store or release significant amounts of C. This study is part of the interregional IAEA Technical Cooperation Project ``Assessing the Impact of Climate Change and its Effects on Soil and Water Resources in Polar and Mountainous Regions (INT5153)'' aiming to improve the understanding of climate change impacts on soil organic carbon (SOC) in fragile polar and high mountainous ecosystems at local and global scale for their better management and conservation. The project includes 13 benchmark sites situated around the world. Here we present novel data from altitudinal transects of three different mountain regions (Mount Kilimanjaro, Tanzania; Mount Gongga, China; Cordillera Blanca, Peru). All altitudinal transects cover a wide range of natural ecosystems under different climates and soil geochemistry. Bulk soil samples (four field replicates per ecosystem) were subjected to a combination of aggregate and particle-size fractionation followed by organic C, total nitrogen, stable isotope (13C, 15N) and radiocarbon (14C) analyses of all fractions. Bulk soils were further characterized for their geochemistry (Na, K, Ca, Mg, Al, Fe, Mn, Si, P) and incubated for 63 days to assess greenhouse gas emissions (CO2, CH4, NO, N2O). Further, stable C isotopic signature of CO2 was measured to determine the isotopic signature of soil respiration (using Keeling plots) and to estimate potential respiration sources. The following four ecosystems were sampled at an altitudinal transect on the (wet) southern slopes of Mount Kilimanjaro: savannah (920m), lower montane rain forests with angiosperm trees (2020m), upper montane cloud forest with gymnosperm trees (2680m), subalpine heathlands (3660m). Both forests showed highest C contents followed by subalpine and savannah. The largest part of SOC was found in particulate organic matter followed by microaggregates, except for the subalpine ecosystem which had most SOC stored in microaggregates. Silt and clay fractions stored the smallest fraction of SOC for all ecosystems. Cumulative soil CO2 emissions (normalized to SOC, gCO_2-C kgSOC-1) after 63 days of incubation were highest for savannah (15.2 ± 1.4) followed by subalpine (7.9 ± 0.5), upper forest (6.9 ± 1.0) and lower forest (4.8 ± 0.4). CO2 emissions were negatively correlated with soil C contents, showing that soils with lower C contents loose higher relative amounts of their SOC through soil respiration. Keeling plot intercept is a measure for the isotopic signature of respired CO2 and high offsets between Keeling plot intercepts and the isotopic signature of bulk SOC point towards labile (13C-depleted) SOC fractions as respiration sources. Highest offsets (and thus most labile respiration sources) were observed for savannah followed by subalpine, lower forest and upper forest and these were positively correlated with cumulative CO2 emissions, showing that in savannah soils, which have lowest C contents and respire highest amounts of CO2, mainly labile SOC is used as respiration source. Results from the other two altitudinal transects are currently under investigation and will be presented in conjunction with climatic and geochemical data.
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Long-term tree inventory data from mountain forest plots in France.
Fuhr, Marc; Cordonnier, Thomas; Courbaud, Benoît; Kunstler, Georges; Mermin, Eric; Riond, Catherine; Tardif, Pascal
2017-04-01
We present repeated tree measurement data from 63 permanent plots in mountain forests in France. Plot elevations range from 800 (lower limit of the montane belt) to 1942 m above sea level (subalpine belt). Forests mainly consist of pure or mixed stands dominated by European beech (Fagus sylvatica), Silver fir (Abies alba), and Norway spruce (Picea abies), in association with various broadleaved species at low elevation and with Arolla pine (Pinus cembra) at high elevation. The plot network includes 23 plots in stands that have not been managed for the last 40 years (at least) and 40 plots in plots managed according to an uneven-aged system with single-tree or small-group selection cutting. Plot sizes range from 0.2 to 1.9 ha. Plots were installed from 1994 to 2004 and remeasured two to five times during the 1994-2015 period. During the first census (installation), living trees more than 7.5 cm in dbh were identified, their diameter at breast height (dbh) was measured and their social status (strata) noted. Trees were spatially located, either with x, y, and z coordinates (40 plots) or within 0.25-ha square subplots (23 plots). In addition, in a subset of plots (58 plots), tree heights and tree crown dimensions were measured on a subset of trees and dead standing trees and stumps were included in the census. Remeasurements after installation include live tree diameters (including recruited trees), tree status (living, damaged, dead, stump), and for a subset of trees, height. At the time of establishment of the plots, plot densities range from 181 to 1328 stems/ha and plot basal areas range from 13.6 to 81.3 m 2 /ha. © 2017 by the Ecological Society of America.
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Phillip E. Farnes; Raymond C. Shearer; Ward W. McCaughey; Katherine J. Hansen
1995-01-01
There are two experimental forests in Montana established by the U.S. Department of Agriculture, Forest Service, Intermountain Research Station (INT). Both experimental forests are administered by INT's Research work Unit, RWU-4151, Silviculture of Subalpine Forest Ecosystems. Tenderfoot Creek Experimental Forest (TCEF) is east of the continental Divide and is...
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Vegetation and soil restoration on highly impacted campsites in the Eagle Cap Wilderness, Oregon
David N. Cole; David R. Spildie
2007-01-01
We assessed the effectiveness of planting techniques (seeding and transplanting) and restoration treatments designed to improve the physical, biological, and chemical properties of soils and ameliorate microclimatic conditions on six closed campsites in subalpine forests. Restoration treatments included scarification, soil amendment with organic matter, compost and...
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Anderson, Lesleigh; Brunelle, Andrea; Thompson, Robert S.
2015-01-01
Apparent changes in vegetation distribution, fire, and other disturbance regimes throughout western North America have prompted investigations of the relative importance of human activities and climate change as potential causal mechanisms. Assessing the effects of Euro-American settlement is difficult because climate changes occur on multi-decadal to centennial time scales and require longer time perspectives than historic observations can provide. Here, we report vegetation and environmental changes over the past ~13,000 years as recorded in a sediment record from Bison Lake, a subalpine lake on a high plateau in northwestern Colorado. Results are based on multiple independent proxies, which include pollen, charcoal, and elemental geochemistry, and are compared with previously reported interpretations of hydroclimatic changes from oxygen isotope ratios. The pollen data indicate a slowly changing vegetation sequence from sagebrush steppe during the late glacial to coniferous forest through the late Holocene. The most dramatic vegetation changes of the Holocene occurred during the ‘Medieval Climate Anomaly’ (MCA) and ‘Little Ice Age’ (LIA) with rapid replacement of conifer forest by grassland followed by an equally rapid return to conifer forest. Late Holocene vegetation responses are mirrored by changes in fire, lake biological productivity, and watershed erosion. These combined records indicate that subsequent disturbance related to Euro-American settlement, although perhaps significant, had acted upon a landscape that was already responding to MCA-LIA hydroclimatic change. Results document both rapid and long-term subalpine grassland ecosystem dynamics driven by agents of change that can be anticipated in the future and simulated by ecosystem models.
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Braking effect of climate and topography on global change-induced upslope forest expansion.
Alatalo, Juha M; Ferrarini, Alessandro
2017-03-01
Forests are expected to expand into alpine areas due to global climate change. It has recently been shown that temperature alone cannot realistically explain this process and that upslope tree advance in a warmer scenario may depend on the availability of sites with adequate geomorphic/topographic characteristics. Here, we show that, besides topography (slope and aspect), climate itself can produce a braking effect on the upslope advance of subalpine forests and that tree limit is influenced by non-linear and non-monotonic contributions of the climate variables which act upon treeline upslope advance with varying relative strengths. Our results suggest that global climate change impact on the upslope advance of subalpine forests should be interpreted in a more complex way where climate can both speed up and slow down the process depending on complex patterns of contribution from each climate and non-climate variable.
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Tree age, disturbance history, and carbon stocks and fluxes in subalpine Rocky Mountain forests
J.B. Bradford; R.A. Birdsey; L.A. Joyce; M.G. Ryan
2008-01-01
Forest carbon stocks and fluxes vary with forest age, and relationships with forest age are often used to estimate fluxes for regional or national carbon inventories. Two methods are commonly used to estimate forest age: observed tree age or time since a known disturbance. To clarify the relationships between tree age, time since disturbance and forest carbon storage...
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Fire history of southeastern Glacier National Park: Missouri River Drainage
Barrett, Stephen W.
1993-01-01
In 1982, Glacier National Park (GNP) initiated long-term studies to document the fire history of all forested lands in the 410,000 ha. park. To date, studies have been conducted for GNP west of the Continental Divide (Barrett et al. 1991), roughly half of the total park area. These and other fire history studies in the Northern Rockies (Arno 1976, Sneck 1977, Arno 1980, Romme 1982, Romme and Despain 1989, Barrett and Arno 1991, Barrett 1993a, Barrett 1993b) have shown that fire history data can be an integral element of fire management planning, particularly wen natiral fire plans are being developed for parks and wilderness. The value of site specific fire history data is apparent when considering study results for lodgepole pin (Pinus contorta var. latifolia) forests. Lodgepole pine is a major subalpine type in the Northern Rockies and such stands experiences a wide range of presettlement fire patterns. On relatively warm-dry sites at lower elevations, such as in GNP's North Fork drainage (Barrett et al. 1991), short to moderately long interval (25-150 yr) fires occurred in a mixed severity pattern ranging from non-lethal underburns to total stand replacement (Arno 1976, Sneck 1977, Barrett and Arno 1991). Markedly different fire history occurred at high elevation lodgepole pine stands on highly unproductive sites, such as on Yellowstone National Park's (YNP) subalpine plateau. Romme (1982) found that, on some sites, stand replacing fires recurred after very long intervals (300-400 yr), and that non-lethal surface fires were rare. For somewhat more productive sites in the Absaroka Mountains in YNP, Barrett (1993a) estimated a 200 year mean replacement interval, in a pattern similar to that found in steep mountain terrain elsewhere, such as in the Middle Fork Flathead River drainage (Barrett et al. 1991, Sneck 1977). Aside from post-1900 written records (ayres 1900; fire atlas data on file, GNP Archives Div. and GNP Resources Mgt. Div.), little fire history information existed for GNP's east-side forests, which are dominated primarily by lodgepole pine. In fall 1992, the park initiated a study to determine the fire history of the Missouri River drainage portion of southeastern GNP. Given the known variation in pre-1900 fire patterns for lodgepole pine, this study was seen as a potentially important contribution to GNP's Fire Management Plan, and to the expanding data base of fire history studies in the region. Resource managers sought this information to assist their development of appropriate fire management strategies for the east-side forests, and the fire history data also would be a useful interactive component of the park's Geographic Information System (GIS). Primary objectives were to: 1) determine pre-1900 fire periodicities, severities, burning patterns, and post-fire succession for major forest types, and 2) document and map the forest age class mosaic, reflecting the history of stand replacing fires at the landscape level of analysis. Secondary objectives were to interpret the possible effects of modern fire suppression on area forests, and to determine fire regime patterns relative to other lodgepole pine ecosystems in the Northern Rockies.
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Tan, Bo; Wu, Fu-Zhong; Yang, Wan-Qin; Yu, Sheng; Yang, Yu-Lian; Wang, Ao
2011-05-01
Late soil-thawing period is a critical stage connecting winter and growth season. The significant temperature fluctuation at this stage might have strong effects on soil ecological processes. In order to understand the soil biochemical processes at this stage in the subalpine/alpine forests of west Sichuan, soil samples were collected from the representative forests including primary fir forest, fir and birch mixed forest, and secondary fir forest in March 5-April 25, 2009, with the activities of soil invertase, urease, and phosphatase (neutral, acid and alkaline phosphatases) measured. In soil frozen period, the activities of the three enzymes in test forests still kept relatively higher. With the increase of soil temperature, the activities of hydrolases at the early stage of soil-thawing decreased rapidly after a sharp increase, except for neutral phosphatease. Thereafter, there was an increase in the activities of urease and phosphatase. Relative to soil mineral layer, soil organic layer had higher hydrolase activity in late soil-thawing period, and showed more obvious responses to the variation of soil temperature.
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Chapter 14. Review of technical knowledge: Great gray owls
James R. Duncan; Patricia H. Hayward
1994-01-01
The great gray owl (Strix nebulosa) is the longest, but not heaviest, of the northern forest owls. Distributed holarctically across the boreal forests of North America and Eurasia, the great gray owl extends its range southward into the contiguous states by inhabiting forests other than the boreal type. The subalpine and montane forests of the...
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Shawn T. McKinney; Carl E. Fiedler; Diana F. Tomback
2009-01-01
Human-caused disruptions to seed-dispersal mutualisms increase the extinction risk for both plant and animal species. Large-seeded plants can be particularly vulnerable due to highly specialized dispersal systems and no compensatory regeneration mechanisms. Whitebark pine (Pinus albicaulis), a keystone subalpine species, obligately depends upon the Clark's...
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Fire, Fuel Composition and Resilience Threshold in Subalpine Ecosystem
Blarquez, Olivier; Carcaillet, Christopher
2010-01-01
Background Forecasting the effects of global changes on high altitude ecosystems requires an understanding of the long-term relationships between biota and forcing factors to identify resilience thresholds. Fire is a crucial forcing factor: both fuel build-up from land-abandonment in European mountains, and more droughts linked to global warming are likely to increase fire risks. Methods To assess the vegetation response to fire on a millennium time-scale, we analyzed evidence of stand-to-local vegetation dynamics derived from sedimentary plant macroremains from two subalpine lakes. Paleobotanical reconstructions at high temporal resolution, together with a fire frequency reconstruction inferred from sedimentary charcoal, were analyzed by Superposed Epoch Analysis to model plant behavior before, during and after fire events. Principal Findings We show that fuel build-up from arolla pine (Pinus cembra) always precedes fires, which is immediately followed by a rapid increase of birch (Betula sp.), then by ericaceous species after 25–75 years, and by herbs after 50–100 years. European larch (Larix decidua), which is the natural co-dominant species of subalpine forests with Pinus cembra, is not sensitive to fire, while the abundance of Pinus cembra is altered within a 150-year period after fires. A long-term trend in vegetation dynamics is apparent, wherein species that abound later in succession are the functional drivers, loading the environment with fuel for fires. This system can only be functional if fires are mainly driven by external factors (e.g. climate), with the mean interval between fires being longer than the minimum time required to reach the late successional stage, here 150 years. Conclusion Current global warming conditions which increase drought occurrences, combined with the abandonment of land in European mountain areas, creates ideal ecological conditions for the ignition and the spread of fire. A fire return interval of less than 150 years would threaten the dominant species and might override the resilience of subalpine forests. PMID:20814580
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Home range and diet of feral cats in Hawaii forests
Smucker, T.D.; Lindsey, G.D.; Mosher, S.M.
2000-01-01
Feral cat Felis catus home range in a Hawaiian montane wet forest and their diet in three habitats - montane wet forest, subalpine dry forest, and lowland dry forest - were determined to provide baseline ecological data and to assess potential impacts to native terrestrial fauna. Seven cats (three males and four females) were captured in 624 trap nights. Mean weight of adult cats was 2.85 ?? 0.27 (SE) Kg for males and 1.87 ?? 0.03 kg for females. Mean diumal home range using the adaptive kernel method was 5.74 ?? 2.73 km2 for three males and 2.23 ?? 0.44 km2 for two females. Daytime locations were always within the montane wet forest with the borders on one or more sides of the home ranges of all cats defined by open grassland pastures. Rodents comprised the majority of the cat diets in all three habitats, with the frequencies of occurence between 0.88 and 0.91. Bird remains were a regular component of the diet of cats, with montane wet forest having the highest frequency of occurence (0.68), followed by subalpine dry forest (0.53), and lowland dry forest (0.21).
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NASA Astrophysics Data System (ADS)
Millar, C. I.; Westfall, R. D.; Delany, D. L.
2010-12-01
Widespread forest mortality in high-elevation forests has been increasing across western North American mountains in recent years, with climate, insects, and disease the primary causes. Subalpine forests in the eastern Sierra Nevada, by contrast, have experienced far less mortality than other ranges, and mortality events have been patchy and episodic. This situation, and lack of significant effect of non-native white-pine blister rust, enable investigation of fine-scale response of two subalpine Sierran species, whitebark pine (Pinus albicaulis, PiAl) and limber pine (P. flexilis, PiFl), to climate variability. We report similarities and differences between the two major mortality events in these pines in the last 150 years: 1988-1992 for PiFl and 2006-ongoing for PiAl. In both species, the events occurred within monotypic, closed-canopy, relatively young stands (< 200 yrs PiAl, < 300 yrs in PiFl); were localized to central-eastern Sierra Nevada; and occurred at 2740-2840 m along the eastern edge of the escarpment on north/northeast aspects with slopes > 40%. Mortality patches averaged 40-80 ha in both species, with mean stand mortality of trees > 10 cm diameter 91% in PiAl and 60% in PiFl. The ultimate cause of tree death was mountain pine beetle (Dendroctonus ponderosae) in both species, with increasing 20th/21st C minimum temperatures combined with drought the pre-conditioning factors. Overall growth in the past 150 years suggests that PiFl is more drought hardy than PiAl but responds sensitively to the combined effects of drought and increasing warmth. After the 1988-1992 drought, surviving PiFl recovered growth. PiAl trees grew very poorly during that drought, and continued poor growth in the years until 2006 when the mortality event occurred in PiAl. A significant species effect is the apparent difference in levels of within-stand genetic diversity for climate factors. Differential growth between 19th C (cool, wet) and 20th/21st C (warming, drying) of PiFl trees that died versus survivors indicates that considerable within-stand genetic diversity for climate existed in PiFl. For PiFl, the late 20th C mortality event acted as strong natural selection to improve within-stand fitness for warmer and drier conditions. PiFl trees that survived the 1988-1992 drought remained healthy through subsequent droughts, including the drought that is currently causing PiAl mortality. By contrast, the PiAl stands do not appear to have contained adaptive genetic diversity for drought and warmth, and PiAl trees growth behavior over the past 150 years was similar in pattern to the PiFl trees that died. As a result, the mortality event in PiAl is creating forest openings, with unknown future stand conditions, rather than rapid within-species adaptation that occurred in PiFl.
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Sara A. Goeking; Deborah K. Izlar
2015-01-01
Whitebark pine (Pinus albicaulis) is an ecologically important species in high-altitude, mid-latitude areas of western North America due to the habitat and food source it provides for many wildlife species. Recent concerns about the long-term viability of whitebark pine stands have arisen in the face of high mortality due to a combination of fire...
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1995-05-01
In wind-swept areas, cushion-like plants with small leaves dominate. The flora of the mountain range encompassing the meadow was listed by Nelson...listed the flora . Elevation effects on sky IR Sky thermal changes across an elevational gradient were measured using a hemispherical, all-wave...in subalpine native flora , different mechanisms exist for protecting active photosynthetic tissues from cold stress than those observed in less frost
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The magnificent high-elevation five-needle white pines: Ecological roles and future outlook
Diana F. Tomback; Peter Achuff; Anna W. Schoettle; John W. Schwandt; Ron J. Mastrogiuseppe
2011-01-01
The High Five symposium is devoted to exchanging information about a small group of pines with little commercial value but great importance to the ecology of high-mountain ecosystems of the West. These High Five pines include the subalpine and treeline species - whitebark (Pinus albicaulis), Rocky Mountain bristlecone (P. aristata), Great Basin bristlecone (P. longaeva...
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Amanda R. Carlson; Jason S. Sibold; Timothy J. Assal; Jose F. Negron
2017-01-01
Spruce beetle (Dendroctonus rufipennis) outbreaks are rapidly spreading throughout subalpine forests of the Rocky Mountains, raising concerns that altered fuel structures may increase the ecological severity of wildfires. Although many recent studies have found no conclusive link between beetle outbreaks and increased fire size or canopy mortality, few studies have...
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Habitat selection by the American marten in northeastern Oregon.
Evelyn L. Bull; Thad W. Heater; Jay F. Shepherd
2005-01-01
Habitat used by 20 adult radio-collared American martens was investigated in northeastern Oregon between 1993 and 1997 to provide land managers with information on habitat management for this species. Martens showed a strong preference for old structure, unlogged stands in subalpine fir and spruce forests with canopy closures 2508, a high density of dead trees and logs...
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Wintertime ozone fluxes and profiles above a subalpine spruce-fir forest
Karl Zeller
2000-01-01
High rural concentrations of ozone (O3) are thought to be stratospheric in origin, advected from upwind urban sources, or photochemically generated locally by natural trace gas emissions. Ozone is known to be transported vertically downward from the above-canopy atmospheric surface layer and destroyed within stomata or on other biological and mineral surfaces. However...
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Ecological contingency in the effects of climatic warming on forest herb communities
Harrison, S.; Damschen, E.I.; Grace, J.B.
2010-01-01
Downscalingfromthe predictions ofgeneral climatemodels is critical to current strategies for mitigating species loss caused by climate change. A key impediment to this downscaling is that we lack a fully developed understanding of howvariation in physical, biological, or land-use characteristics mediates the effects of climate change on ecological communities within regions. We analyzed change in understory herb communities over a 60-y period (1949/1951-2007/ 2009) in a complexmontane landscape (the SiskiyouMountains, Oregon) where mean temperatures have increased 2 ??C since 1948, similar to projections for other terrestrial communities. Our 185 sites included primary and secondary-growth lower montane forests (500-1.200 m above sea level) and primary upper montane to subalpine forests (1,500-2,100 m above sea level). In lower montane forests, regardless of land-use history, we found multiple herbcommunity changes consistent with an effectively drier climate, including lower mean specific leaf area, lower relative cover by species of northern biogeographic affinity, and greater compositional resemblance to communities in southerly topographic positions. At higher elevations we found qualitatively different andmoremodest changes, including increases in herbs of northern biogeographic affinity and in forest canopy cover. Our results provide communitylevel validation of predicted nonlinearities in climate change effects.
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Guidelines for whitebark pine planting prescriptions
Glenda L. Scott; Ward W. McCaughey; Kay Izlar
2011-01-01
Whitebark pine (Pinus albicaulis) is a keystone species in high-elevation ecosystems of the western United States. Unfortunately many fragile subalpine ecosystems are losing whitebark pine as a functional community component due to the combined effects of an introduced disease, insects and succession. Planting whitebark pine is one part of a multifaceted restoration...
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NASA Astrophysics Data System (ADS)
O'Leary, Donal S.; Kellermann, Jherime L.; Wayne, Chris
2018-02-01
Anthropogenic climate change is having significant impacts on montane and high-elevation areas globally. Warmer winter temperatures are driving reduced snowpack in the western USA with broad potential impacts on ecosystem dynamics of particular concern for protected areas. Vegetation phenology is a sensitive indicator of ecological response to climate change and is associated with snowmelt timing. Human monitoring of climate impacts can be resource prohibitive for land management agencies, whereas remotely sensed phenology observations are freely available at a range of spatiotemporal scales. Little work has been done in regions dominated by evergreen conifer cover, which represents many mountain regions at temperate latitudes. We used moderate resolution imaging spectroradiometer (MODIS) data to assess the influence of snowmelt timing and elevation on five phenology metrics (green up, maximum greenness, senescence, dormancy, and growing season length) within Crater Lake National Park, Oregon, USA from 2001 to 2012. Earlier annual mean snowmelt timing was significantly correlated with earlier onset of green up at the landscape scale. Snowmelt timing and elevation have significant explanatory power for phenology, though with high variability. Elevation has a moderate control on early season indicators such as snowmelt timing and green up and less on late-season variables such as senescence and growing season length. PCA results show that early season indicators and late season indicators vary independently. These results have important implications for ecosystem dynamics, management, and conservation, particularly of species such as whitebark pine ( Pinus albicaulis) in alpine and subalpine areas.
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NASA Astrophysics Data System (ADS)
Hansen, W. D.; Braziunas, K. H.; Rammer, W.; Seidl, R.; Turner, M. G.
2017-12-01
Twenty-first century forests will experience increased stress as environmental conditions and disturbance regimes change. Whether forests retain their structure or transitions to alternate states, particularly when affected by multiple stressors, remains unresolved. Subalpine forests in Yellowstone National Park, WY experience large severe wildfires, and postfire-tree regeneration is necessary to assure resilience. Drying is projected, causing frequent larger wildfires that could reduce seed supply and drought that could constrain postfire-seedling establishment. We asked what combinations of warming-drying conditions, increased fire frequency, and increased burned-patch size cause postfire tree-regeneration failure in Yellowstone? We conducted a simulation experiment to identify combinations of fire frequency, fire size, postfire climate, substrate type, and elevation where lodgepole-pine and Douglas-fir regeneration failed. We expected postfire densities to be reduced if burned-patch sizes exceeded effective dispersal distance, sequential fires burned before trees reached reproductive maturity, or drought occurred after fire. We also expected regeneration failure only where multiple stressors occurred simultaneously at low elevation or on poor substrates.Douglas-fir stands were most vulnerable to regeneration failure. 98% of simulated Douglas-fir stands located in the middle of large burned patches failed to regenerate 30 years post fire. Lodgepole-pine stands in the middle of large burned patches failed to regenerate if they were also located at low elevations (93%) or at higher elevations on soils with poor water retention (73%). Stands of serotinous lodgepole (i.e., trees with closed cones that open when heated) also failed to regenerate if fire recurred before trees were reproductively mature (82%). Drought constrained postfire regeneration, yet, enhanced establishment due to release from cold-temperatures during mid-to-late 21st century often outweighed drought effects. Postfire tree regeneration arises from the interplay between multiple factors; some constraining establishment and others enhancing it. To understand 21st-century climate and fire effects on postfire tree regeneration and forest resilience, a reductionist approach is insufficient.
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Jennifer D. Scott; Diana F. Tomback; Michael B. Wunder
2011-01-01
Whitebark pine (Pinus albicaulis), one of five stone pines worldwide, is found at treeline and subalpine elevations in the mountains of western North America (McCaughey and Schmidt 2001). Considered a keystone species, it helps maintain subalpine biodiversity, protects watersheds and promotes post-fire regeneration (Tomback and others 2001). The Clark's nutcracker...
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Lazarus, Brynne E.; Castanha, Cristina; Germino, Matthew; Kueppers, Lara M.; Moyes, Andrew B.
2018-01-01
1.Predictions of upslope range shifts for tree species with warming are based on assumptions of moisture stress at lower elevation limits and low temperature stress at high elevation limits. However, recent studies have shown that warming can reduce tree seedling establishment across the entire gradient from subalpine forest to alpine via moisture limitation. Warming effects also vary with species, potentially resulting in community shifts in high elevation forests. 2.We examined the growth and physiology underlying effects of warming on seedling demographic patterns. We evaluated dry mass (DM), root length, allocation above- and belowground, and relative growth rate (RGR) of whole seedlings, and their ability to avoid or endure water stress via water-use efficiency and resisting turgor loss, for Pinus flexilis, Picea engelmannii and Pinus contorta seeded below, at, and above treeline in experimentally warmed, watered, and control plots in the Rocky Mountains, USA. We expected that growth and allocation responses to warming would relate to moisture status and that variation in drought tolerance traits would explain species differences in survival rates. 3.Across treatments and elevations, seedlings of all species had weak turgor-loss resistance, and growth was marginal with negative RGR in the first growth phase (-0.01 to -0.04 g/g/d). Growth was correlated with soil moisture, particularly in the relatively small-seeded P. contorta and P. engelmannii. P. flexilis, known to have the highest survivorship, attained the greatest DM and longest root but was also the slowest growing and most water-use-efficient. This was likely due to its greater reliance on seed reserves. Seedlings developed 15% less total DM, 25% less root DM, and 11% shorter roots in heated compared to unheated plots. Higher temperatures slightly increased DM, root length and RGR where soils were wettest, but more strongly decreased these variables under drier conditions. 4.Synthesis: The surprising heat-inhibition of tree seedling establishment at the cold edge of forests appears to have a physiological basis: newly germinated seedlings have poor moisture stress tolerance, which appears related to marginal initial growth and heavy reliance on seed reserves. Variation in these attributes among tree species at treeline helps explain their different climate responses.
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NASA Technical Reports Server (NTRS)
Bujakiewicz, A.
1984-01-01
The role and value of fungi in forest associations of Mt. Babia Gora massif were determined. The general physiographic characteristics of the research terrain, the distribution of the fungi sites, a list of the 618 taxons noted in the subalpine forests of Mt. Babia Gora, and the initial characteristics of the forest mycoflora of this massif are presented.
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Water chemistry of Rocky Mountain Front Range aquatic ecosystems
Robert C. Musselman; Laura Hudnell; Mark W. Williams; Richard A. Sommerfeld
1996-01-01
A study of the water chemistry of Colorado Rocky Mountain Front Range alpine/subalpine lakes and streams in wilderness ecosystems was conducted during the summer of 1995 by the USDA Forest Service Arapaho and Roosevelt National Forests and Rocky Mountain Forest and Range Experiment Station, and the University of Colorado Institute of Alpine and Arctic Research. Data...
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Measurements of upward turbulent ozone fluxes above a subalpine spruce-fir forest
Karl Zeller; Ted Hehn
1996-01-01
High rural concentrations of ozone (O3) are thought to be either stratospheric in origin, advected from upwind urban sources, or photochemically generated locally as a result of natural trace gas emissions. Ozone is known to be transported vertically downward from the above-canopy atmospheric surface layer and destroyed within stomata or on other biological and mineral...
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Subalpine bumble bee foraging distances and densities in relation to flower availability.
Elliott, Susan E
2009-06-01
Bees feed almost exclusively on nectar and pollen from flowers. However, little is known about how food availability limits bee populations, especially in high elevation areas. Foraging distances and relationships between forager densities and resource availability can provide insights into the potential for food limitation in mobile consumer populations. For example, if floral resources are limited, bee consumers should fly farther to forage, and they should be more abundant in areas with more flowers. I estimated subalpine bumble bee foraging distances by calculating forager recapture probabilities at increasing distances from eight marking locations. I measured forager and flower densities over the flowering season in six half-hectare plots. Because subalpine bumble bees have little time to build their colonies, they may forage over short distances and forager density may not be constrained by flower density. However, late in the season, when floral resources dwindle, foraging distances may increase, and there may be stronger relationships between forager and flower densities. Throughout the flowering season, marked bees were primarily found within 100 m (and never >1,000 m) from their original marking location, suggesting that they typically did not fly far to forage. Although the density of early season foraging queens increased with early-season flower density, the density of mid- and late-season workers and males did not vary with flower density. Short foraging distances and no relationships between mid- and late-season forager and flower densities suggest that high elevation bumble bees may have ample floral resources for colony growth reproduction.
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NASA Astrophysics Data System (ADS)
Pugh, E. T.; Small, E. E.
2010-12-01
The high-elevation forests that are a primary source for Colorado’s domestic and agricultural water needs are changing rapidly due to an infestation by the mountain pine beetle (MPB). MPB are native to Colorado’s high elevation forests. However, the frequency of MPB infestation and resulting tree death has increased dramatically over the past 15 years. In Colorado, over 8,000 km2 of Lodgepole (Pinus contorta) and Ponderosa Pine (Pinus ponderosa) forest have been infested by MPB since 1996. It is predicted that the current epidemic will kill most of the pines in these areas; MPB are very destructive to forest canopies, often killing all of the overhead trees within lodgepole pine stands. Current widespread MPB outbreaks are not limited to Colorado; they are also impacting forests in much of the Western US and British Columbia, Canada. This study is focused on quantifying the impacts of widespread tree death on Colorado’s mountain snowpack. The data were collected one to three years after beetle infestation, at various stages of tree mortality. During the winters of 2009 and 2010, snowpack and meteorological properties were measured at eight pairs of dead and living lodgepole pine stands. All stands are located at an elevation of 2720 ± 32m, in a subalpine region along the headwaters of the Colorado River. Trees in living stands were generally smaller in diameter and more densely populated than trees in dead stands. In the red phase of tree death, snowpack accumulated equally beneath living and dead tree stands. Additionally, snow under all tree stands became isothermal on the same date regardless of mortality. However, the snow was depleted as much as one week earlier beneath red phase dead stands. Canopy transmission of solar radiation was not consistently different between living and red phase dead stands. We noted more ground litter in red phase dead stands which would decrease snowpack albedo and lead to the snowmelt differences observed. We also performed an albedo experiment to quantify the impact of surface litter on snow albedo. Results are also reported for more advanced grey phase dead stands. Lastly, we present a conceptual model of how the primary snow processes change with time as tree mortality progresses through various stages and introduce future work.
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[Soil macropore characteristics under typical vegetations in Liupan Mountains].
Shi, Zhong-Jie; Wang, Yan-Hui; Xu, Li-Hong; Yu, Peng-Tao; Xiong, Wei; Xu, Da-Ping
2007-12-01
The radius and density of soil macropores under eight typical vegetations in Liupan Mountains of Northwest China were studied by using water breakthrough curves and Poiseuille equation. The results indicated that the radii of soil macropores ranged from 0.4 mm to 2.3 mm, and the weighted mean radii ranged from 0.57 mm to 1.21 mm, with a mean of 0.89 mm. The density of soil macropores ranged from 57 individuals per dm2 to 1 117 individuals per dm2, with a mean of 408 individuals per dm2. The macropores with radii bigger than 1.4 mm had a lower density, accounting for only 6.86% of the total. The area proportion of soil macropores ranged from 0.76% to 31.26%, with a mean of 10.82%. In study area, the density of soil macropores was higher in broadleaf forest than in coniferous forest, but basically the same in sub-alpine meadow and in broadleaf forest, as well as in shrubs and in coniferous forest. As for the area proportion of soil macropores, it was also higher in broadleaf forest than in coniferous forest, but basically the same in shrubs and in broadleaf forest soil, as well as in sub-alpine meadow and in coniferous forest.
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Fish resources within the Tenderfoot Experimental Forest, Montana: 1991-95
Bradley B. Shepard; Susan C. Ireland; Robert G. White
1997-01-01
The Tenderfoot Creek Experimental Forest (TCEF) was established in 1961 to investigate effects of timber harvest on water yield and quality in a subalpine forest of the Northern Rockies draining east of the Continental Divide (Fames et al. 1995; Figure 1). Prior to 1991 the only management activity within the TCEF was the construction of an access road to install...
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Managing coarse woody debris in forests of the Rocky Mountains
Russell T. Graham; Alan E. Harvey; Martin F. Jurgensen; Theresa B. Jain; Jonalea R. Tonn; Deborah S. Page-Dumroese
1994-01-01
Recommendations for managing coarse woody debris after timber harvest were developed for 14 habitat types, ranging from ponderosa pine (Pinus ponderosa) habitat types of Arizona to subalpine fir (Abies lasiocarpa) habitat types of western Montana. Ectomycorrhizae were used as a bioindicator of healthy, productive forest soils....
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William C. Fischer
1981-01-01
Two series of color photographs show different levels of downed woody material resulting from natural processes in two forest cover types in Montana. Each photo is supplemented by fuel inventory data and potential fire· behavior ratings.
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Chapter 9. Review of technical knowledge: Boreal owls
Gregory D. Hayward
1994-01-01
The boreal owl (Aegolius funereus), known as Tengmalm's owl in Eurasia, occurs throughout the holarctic in boreal climatic zones. This medium-size owl (100-170 g) occupies boreal and subalpine forests in an almost continuous circumboreal distribution that extends from Scandinavia eastward across the northern forests of Siberia and from Alaska...
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Todd, Christopher M.; Pinzari, Corinna A.; Bonaccorso, Frank
2016-01-01
The Kahikinui Forest Reserve and the adjoining Nakula Natural Area Reserve (KFR-NNAR) was established in 2011 as a conservation area on the leeward slope of Haleakalā Volcano on the island of Maui to protect unique natural features and endangered species including the Hawaiian hoary bat, Lasiurus cinereus semotus. We recorded bat vocalizations from July 2012 to November 2014 using automated echolocation detectors at 14 point locations in the KFRNNAR. Our study area included remnants of recovering mesic montane forest with interspersed grasses (1,250‒1,850 m elevation, hereafter called “forest”) and xeric subalpine shrubland plant communities (1,860‒2,800 m, hereafter called “shrubland”). Monthly detections of Hawaiian hoary bats, Lasiurus cinereus semotus, within the KFR-NNAR identified areas of high and low detection probability as well as foraging activity. Sixty per cent of all detector-nights had confirmed bat vocalizations and included detections in every month of the study. Monthly detection probability values were highest from July to November 2012; these values were significantly greater than values measured in any month thereafter. Pooled values of detection probabilities, mean pulses/night, percentage of nights with feeding activity, and acoustic detections all were greater in the recovering forest zone than corresponding values from the shrublands. Our data provide baseline levels of hoary bat echolocation activity that may be compared with future studies in the KFR-NNAR relative to success criteria for Hawaiian hoary bat habitat restoration.
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Preface [to special section on recent Loch Vale Watershed research
Baron, Jill S.; Williams, Mark W.
2000-01-01
Catchment-scale intensive and extensive research conducted over the last decade shows that our understanding of the biogeochemical and hydrologic processes in subalpine and alpine basins is not yet sufficiently mature to model and predict how biogeochemical transformations and surface water quality will change in response to climatic or human-driven changes in energy, water, and chemicals. A better understanding of these processes is needed for input to decision-making regulatory agencies and federal land managers. In recognition of this problem the National Research Council [1998] has identified as a critical research need an improved understanding of how global change will affect biogeochemical interactions with the hydrologic cycle and biogeochemical controls over the transport of water, nutrients, and materials from land to freshwater ecosystems. Improved knowledge of alpine and subalpine ecosystems is particularly important since high-elevation catchments are very sensitive to small changes in the flux of energy, chemicals, and water. Furthermore, alpine ecosystems may act as early warning indicators for ecosystem changes at lower elevations.
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R. Scott Anderson; Susan J. Smith; Ann M. Lynch; Brian W. Geils
2010-01-01
The frequency and intensity of ecosystem disturbance, including outbreaks of forest insects and forest fires, is expected to increase in the future as a result of higher temperatures and prolonged drought. While many studies have concentrated on the future climatic impacts on fire, little is known about the impact of future climate on insect infestation....
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E. Berryman; Michael Ryan; J. B. Bradford; T. J. Hawbaker; R. Birdsey
2016-01-01
In forests, total belowground carbon (C) flux (TBCF) is a large component of the C budget and represents a critical pathway for delivery of plant C to soil. Reducing uncertainty around regional estimates of forest C cycling may be aided by incorporating knowledge of controls over soil respiration and TBCF. Photosynthesis, and presumably TBCF, declines with...
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The contribution of red wood ants to soil C and N pools and CO2 emissions in subalpine forests
Anita C. Risch; Martin F. Jurgensen; Martin Schutz; Deborah S. Page-Dumroese
2005-01-01
Little information is available regarding red wood ant (RWA; Formica rufa group) impacts on soil carbon (C) and nitrogen (N) cycling in forest ecosystems. We found that RWA mound density (number per ha) was linked to forest tree species composition, slope aspect, and canopy closure. The size of RWA mounds was positively correlated with successional...
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Xia, Lei; Wu, Fu-Zhong; Yang, Wan-Qin; Tan, Bo
2012-02-01
In order to quantify the contribution of soil fauna to the decomposition of birch (Betula albosinensis) leaf litter in subalpine forests in western Sichuan of Southwest China during freeze-thaw season, a field experiment with different mesh sizes (0.02, 0.125, 1 and 3 mm) of litterbags was conducted in a representative birch-fir (Abies faxoniana) forest to investigate the mass loss rate of the birch leaf litter from 26 October, 2010 to 18 April, 2011, and the contributions of micro-, meso- and macro-fauna to the decomposition of the leaf litter. Over the freeze-thaw season, 11.8%, 13.2%, 15.4% and 19.5% of the mass loss were detected in the litterbags with 0.02, 0. 125, 1 and 3 mm mesh sizes, respectively. The total contribution of soil fauna to the litter decomposition accounted for 39.5% of the mass loss, and the taxa and individual relative density of the soil fauna in the litterbags had the similar variation trend with that of the mass loss rate. The contribution rate of soil fauna to the leaf litter mass loss showed the order of micro- < meso- < macro-fauna, with the highest contribution of micro-fauna (7.9%), meso-fauna (11.9%), and macro-fauna (22.7%) at the onset of freezing stage, deeply frozen stage, and thawing stage, respectively. The results demonstrated that soil fauna played an important role in the litter decomposition in subalpine forests of western Sichuan during freeze-thaw season.
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Ecological contingency in the effects of climatic warming on forest herb communities.
Harrison, Susan; Damschen, Ellen I; Grace, James B
2010-11-09
Downscaling from the predictions of general climate models is critical to current strategies for mitigating species loss caused by climate change. A key impediment to this downscaling is that we lack a fully developed understanding of how variation in physical, biological, or land-use characteristics mediates the effects of climate change on ecological communities within regions. We analyzed change in understory herb communities over a 60-y period (1949/1951-2007/2009) in a complex montane landscape (the Siskiyou Mountains, Oregon) where mean temperatures have increased 2 °C since 1948, similar to projections for other terrestrial communities. Our 185 sites included primary and secondary-growth lower montane forests (500-1.200 m above sea level) and primary upper montane to subalpine forests (1,500-2,100 m above sea level). In lower montane forests, regardless of land-use history, we found multiple herb-community changes consistent with an effectively drier climate, including lower mean specific leaf area, lower relative cover by species of northern biogeographic affinity, and greater compositional resemblance to communities in southerly topographic positions. At higher elevations we found qualitatively different and more modest changes, including increases in herbs of northern biogeographic affinity and in forest canopy cover. Our results provide community-level validation of predicted nonlinearities in climate change effects.
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Ecological contingency in the effects of climatic warming on forest herb communities
Harrison, Susan; Damschen, Ellen Ingman; Grace, James B.
2010-01-01
Downscaling from the predictions of general climate models is critical to current strategies for mitigating species loss caused by climate change. A key impediment to this downscaling is that we lack a fully developed understanding of how variation in physical, biological, or land-use characteristics mediates the effects of climate change on ecological communities within regions. We analyzed change in understory herb communities over a 60-y period (1949/1951–2007/2009) in a complex montane landscape (the Siskiyou Mountains, Oregon) where mean temperatures have increased 2 °C since 1948, similar to projections for other terrestrial communities. Our 185 sites included primary and secondary-growth lower montane forests (500–1.200 m above sea level) and primary upper montane to subalpine forests (1,500–2,100 m above sea level). In lower montane forests, regardless of land-use history, we found multiple herb-community changes consistent with an effectively drier climate, including lower mean specific leaf area, lower relative cover by species of northern biogeographic affinity, and greater compositional resemblance to communities in southerly topographic positions. At higher elevations we found qualitatively different and more modest changes, including increases in herbs of northern biogeographic affinity and in forest canopy cover. Our results provide community-level validation of predicted nonlinearities in climate change effects.
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Recreation in whitebark pine ecosystems: Demand, problems, and management strategies
David N. Cole
1990-01-01
Whitebark pine ecosystems are an important element of many of the most spectacular high-elevation landscapes in the western United States. They occupy upper subalpine and timberline zones in the prime recreation lands of the Cascades, the Sierra Nevada, and the Northern Rocky Mountains. This paper explores the nature of the recreational opportunities that the whitebark...
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Wind and ecosystem response at the GLEES
Robert C. Musselman; Gene L. Wooldridge; William J. Massman; Richard A. Sommerfeld
1995-01-01
Research was conducted to determine wind patterns and snow deposition at a high elevation alpine/subalpine ecotone site using deformation response of trees to prevailing winds. The research has provided detailed maps of wind speed, wind direction, and snow depth as determined from tree deformation. The effects of prevailing wind on tree blowdown at the site have also...
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Hydrology and landscape structure control subalpine catchment carbon export
Vincent Jerald Pacific
2009-01-01
Carbon export from high elevation ecosystems is a critical component of the global carbon cycle. Ecosystems in northern latitudes have become the focus of much research due to their potential as large sinks of carbon in the atmosphere. However, there exists limited understanding of the controls of carbon export from complex mountain catchments due to strong spatial and...
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Low Elevation Riparian Environments: Warm-Climate Refugia for Conifers in the Great Basin, USA?
NASA Astrophysics Data System (ADS)
Millar, C.; Charlet, D. A.; Westfall, R. D.; Delany, D.
2015-12-01
The Great Basin, USA, contains hundreds of small to large mountain ranges. Many reach alpine elevations, which are separated from each other by low-elevation basins currently inhospitable to conifer growth. Many of these ranges support montane and subalpine conifer species that have affinities to the Sierra Nevada or Rocky Mountains, and from which these conifers migrated during cool periods of the Pleistocene. Under Holocene climates, the Great Basin geography became a terrestrial island-archipelago, wherein conifer populations are isolated among ranges, and inter-range migration is highly limited. During warm intervals of the Holocene, conifers would be expected to have migrated upslope following favorable conditions, and extirpation would be assumed to result from continued warming. Independent patterns, repeating across multiple species' distributions, however, suggest that refugia were present in these ranges during warm periods, and that low elevation environments below the current main distributions acted as climatic refugia. We hypothesize that cool, narrow, and north-aspect ravines, which during cool climates support persistent or seasonal creeks and deciduous riparian communities, become available as conifer habitat when warming climates desiccate creeks and deplete riparian species. We further speculate that cold-air drainage, reduced solar insolation, lower wind exposure, and higher water tables in these topographic positions support populations of montane and subalpine conifers even during warm climate intervals when high elevations are unfavorable for conifer persistence. On return to cool climates, low elevation refugia become sources for recolonizing higher slopes, and/or continue to persist as relictual populations. We present several lines of evidence supporting this hypothesis, and speculate that low-elevation, extramarginal riparian environments might act as climate refugia for Great Basin conifers in the future as well.
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Spectral evidence of early-stage spruce beetle infestation in Engelmann spruce
Adrianna C. Foster; Jonathan A. Walter; Herman H. Shugart; Jason Sibold; Jose Negron
2017-01-01
Spruce beetle (Dendroctonus rufipennis (Kirby)) outbreaks cause widespread mortality of Engelmann spruce (Picea engelmannii (Parry ex Engelm)) within the subalpine forests of the western United States. Early detection of infestations could allow forest managers to mitigate outbreaks or anticipate a response to tree mortality and the potential effects on ecosystem...
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Raymond C. Shearer
2008-01-01
The larches - Larix P. Mill. - of the world are usually grouped into 10 species that are widely distributed over much of the mountainous, cooler regions of the Northern Hemisphere (Hora 1981; Krussmann 1985; Ostenfeld and Larsen 1930; Rehder 1940; Schmidt 1995). Some species dominate at the northern limits of boreal forests and others occur above subalpine forests (...
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Altered species interactions and implications for natural regeneration in whitebark pine communities
Shawn T. McKinney; Diana F. Tomback; Carl E. Fiedler
2011-01-01
Whitebark pine (Pinus albicaulis) decline has altered trophic interactions and led to changes in community dynamics in many Rocky Mountain subalpine forests (McKinney and Tomback 2007). Here we discuss how altered species interactions, driven by disproportionate whitebark pine mortality, constrain the capability of whitebark pine forests to contribute genetic material...
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Calvin A. Farris; Ellis Q. Margolis; John A. Kupfer
2008-01-01
We compared the spatial characteristics of fire severity patches within individual fire ârunsâ (contiguous polygons burned during a given day) resulting from a 72,000 ha fire in centralIdaho in 1994. Our hypothesis was that patch characteristics of four fire severity classes (high, moderate, low, and unburned), as captured by five landscape metrics, would...
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NASA Astrophysics Data System (ADS)
Burns, Sean P.; Maclean, Gordon D.; Blanken, Peter D.; Oncley, Steven P.; Semmer, Steven R.; Monson, Russell K.
2016-09-01
The Niwot Ridge Subalpine Forest AmeriFlux site (US-NR1) has been measuring eddy-covariance ecosystem fluxes of carbon dioxide, heat, and water vapor since 1 November 1998. Throughout this 17-year period there have been changes to the instrumentation and improvements to the data acquisition system. Here, in Part 1 of this three-part series of papers, we describe the hardware and software used for data-collection and metadata documentation. We made changes to the data acquisition system that aimed to reduce the system complexity, increase redundancy, and be as independent as possible from any network outages. Changes to facilitate these improvements were (1) switching to a PC/104-based computer running the National Center for Atmospheric Research (NCAR) In-Situ Data Acquisition Software (NIDAS) that saves the high-frequency data locally and over the network, and (2) time-tagging individual 10 Hz serial data samples using network time protocol (NTP) coupled to a GPS-based clock, providing a network-independent, accurate time base. Since making these improvements almost 2 years ago, the successful capture of high-rate data has been better than 99.98 %. We also provide philosophical concepts that shaped our design of the data system and are applicable to many different types of environmental data collection.
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NASA Astrophysics Data System (ADS)
Martin, Y. E.; Johnson, E. A.; Chaikina, O.
2013-10-01
During the cycle of forest disturbance, regeneration, and maturity, tree mortality leading to topple is a regular occurrence. When tree topple occurs relatively soon after mortality and if the tree has attained some threshold diameter at breast height (dbh) at the time of death, then notable amounts of soil may be upheaved along with the root wad. This upheaval may result in sediment transfers and soil production. A combination of field evidence and numerical modeling is used herein to gain insights regarding the temporal dynamics of tree topple, associated root throw processes, and pit-mound microtopography. Results from our model of tree population dynamics demonstrate temporal patterns in root throw processes in subalpine forests of the Canadian Rockies, a region in which forests are affected largely by wildfire disturbance. As the forest regenerates after disturbance, the new cohort of trees has to reach a critical dbh before significant root plate upheaval can occur; in the subalpine forests of the Canadian Rockies, this may take up to ~ 102 years. Once trees begin to reach this critical dbh for root plate upheaval, a period of sporadic root throw arises that is caused by mortality of trees during competition. In due course, another wildfire will occur on the landscape and a period of much increased root throw activity then takes place for the next several decades; tree sizes and, therefore, the amount of sediment disturbance will be greater the longer the time period since the previous fire. Results of previous root throw studies covering a number of regional settings are used to guide an exercise in diffusion modeling with the aim of defining a range of reasonable diffusion coefficients for pit-mound degradation; the most appropriate values to fit the field data ranged from 0.01 m2 y- 1 to 0.1 m2 y- 1. A similar exercise is then undertaken that is guided by our field observations in subalpine forests of the Canadian Rockies. For these forests, the most appropriate range of diffusion coefficients is in the range 0.001 m2 y- 1 to 0.01 m2 y- 1. Finally, the model of tree population dynamics is combined with the model of pit-mound degradation to demonstrate the integration of these combined processes on the appearance of pit-mound microtopography and soil bioturbation in subalpine forests of the Canadian Rockies. We conclude that the appearance of notable pit-mound microtopography is limited to very specific time periods and is not visible for much of the time. Most of the hillslope plot is affected by root throw during the 1000-year model run time.
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Buckley, Thomas N; Turnbull, Tarryn L; Pfautsch, Sebastian; Adams, Mark A
2011-01-01
We measured sap flux (S) and environmental variables in four monospecific stands of alpine ash (Eucalyptus delegatensis R. Baker, AA) and snowgum (E. pauciflora Sieb. ex Spreng., SG) in Australia's Victorian Alps. Nocturnal S was 11.8 ± 0.8% of diel totals. We separated transpiration (E) and refilling components of S using a novel modeling approach based on refilling time constants. The nocturnal fraction of diel water loss (fn) averaged 8.6 ± 0.6% for AA and 9.8 ± 1.7% for SG; fn differed among sites but not species. Evaporative demand (D) was the strongest driver of nocturnal E (En). The ratio En/D (Gn) was positively correlated to soil moisture in most cases, whereas correlations between wind speed and Gn varied widely in sign and strength. Our results suggest (1) the large, mature trees at our subalpine sites have greater fn than the few Australian native tree species that have been studied at lower elevations, (2) AA and SG exhibit similar fn despite very different size and life history, and (3) fn may differ substantially among sites, so future work should be replicated across differing sites. Our novel approach to quantifying fn can be applied to S measurements obtained by any method. PMID:22393512
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David W. Peterson; Becky K. Kerns; Erich Kyle Dodson
2014-01-01
The purpose of this study was to review scientifi c knowledge and model projections on vegetation vulnerability to climatic and other environmental changes in the Pacifi c Northwest, with emphasis on fi ve major biome types: subalpine forests and alpine meadows, maritime coniferous forests, dry coniferous forests, savannas and woodlands (oak and juniper), and interior...
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Xian, Jun-Ren; Hu, Ting-Xing; Zhang, Yuan-Bin; Wang, Kai-Yun
2007-04-01
By the method of strip transect sampling, the density, height, basal diameter, and components biomass of Abies faxoniana seedlings (H < or = 100 cm) lived in the forest gap (FG) and under the forest canopy (FC) of subalpine natural coniferous forest in West Sichuan were investigated, and the relationships among different components biomass were analyzed. The results indicated that the density and average height (H) of A. faxoniana seedlings were significantly different in FG and under FC, with the values being 12 903 and 2 017 per hectare, and 26.6 cm and 24.3 cm, respectively, while no significant differences were found in average basal diameter (D) and biomass. The biomass allocation in seedling's components was markedly affected by forest gap. In FG, the biomass ratio of branch to trunk (BRBT) reached the maximum (1.54) at 12th year, and then, declined and fluctuated at 0. 69. Under FC, the BRBT was increased with seedlings growth, and exceeded 1.0 at about 15th year. The total biomass and the biomass of leaf, stem, shoot and root grown in FG and under FC were significantly linearly correlated with D2H. There were significant positive correlations among the biomass of different seedling's components.
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NASA Astrophysics Data System (ADS)
Molotch, Noah P.; Barnard, David M.; Burns, Sean P.; Painter, Thomas H.
2016-09-01
The distribution of forest cover exerts strong controls on the spatiotemporal distribution of snow accumulation and snowmelt. The physical processes that govern these controls are poorly understood given a lack of detailed measurements of snow states. In this study, we address one of many measurement gaps by using contact spectroscopy to measure snow optical grain size at high spatial resolution in trenches dug between tree boles in a subalpine forest. Trenches were collocated with continuous measurements of snow depth and vertical profiles of snow temperature and supplemented with manual measurements of snow temperature, geometric grain size, grain type, and density from trench walls. There was a distinct difference in snow optical grain size between winter and spring periods. In winter and early spring, when facetted snow crystal types were dominant, snow optical grain size was 6% larger in canopy gaps versus under canopy positions; a difference that was smaller than the measurement uncertainty. By midspring, the magnitude of snow optical grain size differences increased dramatically and patterns of snow optical grain size became highly directional with 34% larger snow grains in areas south versus north of trees. In winter, snow temperature gradients were up to 5-15°C m-1 greater under the canopy due to shallower snow accumulation. However, in canopy gaps, snow depths were greater in fall and early winter and therefore more significant kinetic growth metamorphism occurred relative to under canopy positions, resulting in larger snow grains in canopy gaps. Our findings illustrate the novelty of our method of measuring snow optical grain size, allowing for future studies to advance the understanding of how forest and meteorological conditions interact to impact snowpack evolution.
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P.F. Hessburg; R.G. Mitchell; G.M. Filip
1994-01-01
This paper examines by climax conifer series, historical and current roles of many important pathogens and insects of interior Northwest coniferous forests, and their unique responses to changing successional conditions resulting from management. Insects and pathogens of the subalpine fir and mountain hemlock series historically reduced inter-tree competition for site...
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Marcus V. Warwell; Gerald E. Rehfeldt; Nicholas L. Crookston
2010-01-01
The Random Forests multiple regression tree was used to develop an empirically based bioclimatic model of the presence-absence of species occupying small geographic distributions in western North America. The species assessed were subalpine larch (Larix lyallii), smooth Arizona cypress (Cupressus arizonica ssp. glabra...
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Management of subalpine forests: Building on 50 years of research
Charles A. Troendle; Merrill R. Kaufmann; R. H. Hamre; Robert P. Winokur
1987-01-01
This summer marked the 50th anniversary of the Fraser Experimental Forest. These proceedings are the tangible product of the technical conference called to summarize, discuss, and transfer the knowledge learned over these 50 years. Eighteen formal papers describe the status of our knowledge about the interactions among timber, water, and wildlife. These papers were...
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NASA Astrophysics Data System (ADS)
Williams, C.; Silins, U.; Wagner, M. J.; Bladon, K. D.; Martens, A. M.; Anderson, A.; Stone, M.; Emelko, M. B.
2014-12-01
Interception of precipitation in sub-alpine forests is likely to be strongly reduced after wildfire, potentially producing large increases in net precipitation. Objectives of this study were to describe changes in rainfall and snow interception, and net precipitation after the severe 2003 Lost Creek wildfire as part of the Southern Rockies Watershed Project in the south-west Rocky Mountains of Alberta, Canada. Throughfall troughs and stemflow gauges were used to explore relationships between throughfall, stemflow, and net rainfall with variation in gross rainfall in burned and undisturbed stands during the summers of 2006-2008. These relationships were used to scale the effects of the wildfire on net rainfall for the first decade after the wildfire (2004-2013) using a 10 year rainfall record in the watershed. Annual snowpack surveys (5 snow courses in each of burned and reference stands) measured peak snowpack depth, density, and snow water equivalent (SWE) for this same period. Mean annual P was 1140 mm (684-1519 mm) during the first 10 years after the wildfire, with 61% falling as snow. Throughfall and stemflow in the burned forest accounted for 86% and 7% of gross rainfall, respectively, compared with 53% and 0.002% in the unburned stands in the summers of 2006-2008. Scaled rainfall interception relationships (=f(rainfall event size)) indicated annual increases in net rainfall were 192 mm/yr (133-347 mm) for 10 years after the fire. Similarly, mean increases in peak SWE were 134 mm/yr (93-216 mm). Collectively, the mean increase in net precipitation was 325 mm/yr (226-563 mm; 29%) for the first decade after the wildfire. Hydrologic forcing by increased net precipitation may be a particularly important element of wildfire impacts on sub-alpine watersheds. Furthermore, because of the very slow growth rates of sub-alpine forests, increases in net precipitation are likely to persist and affect precipitation-runoff relationships for decades in these environments.
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Harold S.J. Zald; Thomas A. Spies; Manuela Huso; Demetrios Gatziolis
2012-01-01
Tree invasions have been documented throughout Northern Hemisphere high elevation meadows, as well as globally in many grass and forb-dominated ecosystems. Tree invasions are often associated with large-scale changes in climate or disturbance regimes, but are fundamentally driven by regeneration processes influenced by interactions between climatic, topographic, and...
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Paul E. Trusty; Cathy L. Cripps
2011-01-01
Whitebark pine (Pinus albicaulis) is a threatened keystone species in subalpine zones of Western North America that plays a role in watershed dynamics and maintenance of high elevation biodiversity (Schwandt, 2006). Whitebark pine has experienced significant mortality due to white pine blister rust, mountain pine beetle outbreaks and successional replacement possibly...
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Nefedieva, Julia S.; Nefediev, Pavel S.; Sakhnevich, Miroslava B.; Dyachkov, Yuri V.
2015-01-01
Abstract The distribution of millipedes along an altitudinal gradient in the south of Lake Teletskoye, Altai, Russia based on new samples from the Kyga Profile sites, as well as on partly published and freshly revised material (Mikhaljova et al. 2007, 2008, 2014, Nefedieva and Nefediev 2008, Nefediev and Nefedieva 2013, Nefedieva et al. 2014), is established. The millipede diversity is estimated to be at least 15 species and subspecies from 10 genera, 6 families and three orders. The bulk of species diversity is confined both to low- and mid-mountain chern taiga forests and high-mountain shrub tundras, whereas the highest numbers, reaching up to 130 ind./m², is shown in subalpine Pinus sibirica sparse growths. Based on clustering studied localities on species diversity similarity two groups of sites are defined: low-mountain sites and subalpine sparse growths of Pinus sibirica ones. PMID:26257540
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[Maples at the sub-Alpine vegetation belt: a long history].
David, F; Barbero, M
2001-02-01
Pollen analysis was carried out on lacustrine sediment of a small hollow (15 m x 25 m) at the treeless sub-Alpine belt (202 m) of the inner Maurienne valley in the northern French Alps. A 2,500-year-long maple settlement was demonstrared. Three AMS dates of terrestrial plant macroremains support the chronology. First, Betula and Salix spread prior to 9,000 C14 BP. The first pollen grains of Acer, Abies and Pinus cembra are quoted at 8,600 C14 BP. High frequencies of Alnus glutinosa/incana (20%) and Acer (10%) show that mixed communities of Acer and Alnus persisted above the mountainous Abies forest between 7,490 and 5,850 C14 BP. After 5,850 C14 BP, the decrease in Acer stands could be attributed to fire as suggested by the strong increase in Betula and by the delayed expansion of Pinus cembra.
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Kristen Pelz; C. C. Rhoades; R. M. Hubbard; M. A. Battaglia; F. W. Smith
2015-01-01
Mountain pine beetle outbreaks have killed lodgepole pine on more than one million hectares of Colorado and southern Wyoming forest during the last decade and have prompted harvest operations throughout the region. In northern Colorado, lodgepole pine commonly occurs in mixed stands with subalpine fir, Engelmann spruce, and aspen. Variation in tree species composition...
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James M. Lenihan; Dominique Bachelet; Ronald P. Neilson; Raymond Drapeck
2008-01-01
The response of vegetation distribution, carbon, and fire to three scenarios of future climate change was simulated for California using the MC1 Dynamic General Vegetation Model. Under all three scenarios, Alpine/Subalpine Forest cover declined, and increases in the productivity of evergreen hardwoods led to the displacement of Evergreen Conifer Forest by Mixed...
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Christy E. Briles; Cathy Whitlock; Carl N. Skinner; Jerry Mohr
2011-01-01
The influence of substrate on long-term vegetation dynamics has received little attention, and yet nutrient-limited ecosystems have some of the highest levels of endemism in the world. The diverse geology of the Klamath Mountains of northern California (USA) allows examination of the long-term influence of edaphic constraints in subalpine forests through a comparison...
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Grizelle Gonzalez; Timothy R. Seastedt; Zugeily Donato
2003-01-01
We compared the abundance and community composition of earthworms, soil macroarthropods, and litter microarthropods to test faunal effects on plant litter decomposition rates in two forests in the subalpine in Colorado, USA. Litterbags containing recently senesced litter of Populus tremuloides (aspen) and Pinus contorta (lodgepole pine) were placed in aspen and pine...
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Jon M. Skovlin; Gerald S. Strickler; Jesse L. Peterson; Arthur W. Sampson
2001-01-01
We compared 45 photographs taken before 1925 to photographs taken as late as 1999 and documented landscape changes above 5,000 feet elevation in the Wallowa, Elkhorn, and Greenhorn Mountains of northeastern Oregon. We noted the following major changes from these comparisons: (1) the expansion of subalpine fir into mountain grasslands, (2) the invasion of moist and wet...
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Anna W. Schoettle; Richard A. Sniezko; Angelia Kegley; Kelly S. Burns
2011-01-01
Limber pine ( Pinus flexilis James) and Rocky Mountain bristlecone pine (P. aristata Engelm.; hereafter referred to as bristlecone pine) are the dominant pines that occupy high elevation habitats of the southern Rockies. Bristlecone pine is primarily a subalpine and tree-line species while limber pine in the southern Rocky Mountains grows from 1600 m in the short grass...
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NASA Astrophysics Data System (ADS)
Bunte, Kristin; Swingle, Kurt W.; Turowski, Jens M.; Abt, Steven R.; Cenderelli, Daniel A.
2016-08-01
Coarse particulate organic matter (CPOM) provides a food source for benthic organisms, and the fluvial transport of CPOM is one of the forms in which carbon is exported from a forested basin. However, little is known about transport dynamics of CPOM, its relation to discharge, and its annual exports from mountain streams. Much of this knowledge gap is due to sampling difficulties. In this study, CPOM was sampled over one-month snowmelt high flow seasons in two high-elevation, subalpine, streams in the Rocky Mountains. Bedload traps developed for sampling gravel bedload were found to be suitable samplers for CPOM transport. CPOM transport rates were well related to flow in consecutive samples but showed pronounced hysteresis over the diurnal fluctuations of flow, between consecutive days, and over the rising and falling limbs of the high-flow season. In order to compute annual CPOM load, hysteresis effects require intensive sampling and establishing separate rating curves for all rising and falling limbs. Hysteresis patterns of CPOM transport relations identified in the well-sampled study streams may aid with estimates of CPOM transport and export in less well-sampled Rocky Mountain streams. Transport relations for CPOM were similar among three high elevation mountain stream with mainly coniferous watersheds. Differences among streams can be qualitatively attributed to differences in CPOM contributions from litter fall, from the presence of large woody debris, its grinding into CPOM sized particles by gravel-cobble bedload transport, hillslope connectivity, drainage density, and biological consumption. CPOM loads were 3.6 and 3.2 t/yr for the two Rocky Mountain streams. Adjusted to reflect decadal averages, values increased to 11.3 and 10.2 t/yr. CPOM yields related to the entire watershed were 2.7 and 4 kg/ha/yr for the years studied, but both streams exported similar amounts of 6.5 and 6.6 kg/ha/yr when taking the forested portion of the watershed into account. To reflect decadal averages, CPOM yields per basin area were adjusted to 8.6 and 12.6 kg/ha/yr and to 21 kg/ha/yr for the forested watershed parts. CPOM yield may be more meaningfully characterized if annual CPOM loads are normalized by the area of a seam along the stream banks together with the stream surface area rather than by the forested or total watershed area.
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NASA Astrophysics Data System (ADS)
Reinhardt, K.; Castanha, C.; Germino, M. J.; Kueppers, L. M.
2011-12-01
The elevation limit of tree growth (alpine treeline) is considered to be constrained by environmental (i.e., thermal) and genetic (i.e., inability to adapt to climatic conditions) limitations to growth. Warming conditions due to climate change are predicted to cause upward shifts in the elevation of alpine treelines, through relief of cold-induced physiological limitations on seedling recruitment beyond current treeline boundaries. To determine how genetics and climate may interact to affect seedling establishment, we transplanted recently germinated seedlings from high- and low-elevation provenances (HI and LO, respectively) of Pinus flexilis in common gardens arrayed along an elevation and canopy gradient from subalpine forest into the alpine zone at Niwot Ridge, CO. We compared differences in microclimate and seedling ecophysiology among sites and between provenances. During the first summer of growth, frequently cloudy skies resulted in similar solar radiation incidence and air and soil temperatures among sites, despite nearly a 500 m-span in elevation across all sites. Preliminary findings suggest that survival of seedlings was similar between the lowest and highest elevations, with greater survival of LO (60%) compared to HI (40%) seedlings at each of these sites. Photosynthesis, carbon balance (photosynthesis/respiration), and conductance increased more than 2X with elevation for both provenances, and were 35-77% greater in LO seedlings compared to HI seedlings. There were no differences in dark-adapted chlorophyll fluorescence (Fv/Fm) among sites or between provenances. However, in a common-garden study at low elevation, we observed no differences in carbon or water relations between two naturally-germinated mitochondrial haplotypes of P. flexilis (of narrow and wide-ranging distributions). We did observe water-related thresholds on seedling carbon balance and survival that occurred when soil volumetric water content dropped below 10% and seedling water potentials went below -4 MPa. Our preliminary results suggest that for high-elevation conifer seedlings such as P. flexilis: 1) individuals can survive and even have enhanced physiological performance at and above treeline when/where clouds or other conditions minimize factors like cold-induced photoinhibition; 2) in the field, provenances selected for aboveground growth may out-perform those selected for stress-resistance in the absence of harsh climatic conditions, even well above the species' range limits in the alpine; 3) water, and not thermal, limitations might explain treeline altitude in this particular mountain range; 4) forest genetics may be important to understanding and managing species' range adjustments due to climate change.
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Burns, Sean P.; Maclean, Gordon D.; Blanken, Peter D.; ...
2016-09-29
The Niwot Ridge Subalpine Forest AmeriFlux site (US-NR1) has been measuring eddy-covariance ecosystem fluxes of carbon dioxide, heat, and water vapor since 1 November 1998. Throughout this 17-year period there have been changes to the instrumentation and improvements to the data acquisition system. Here, in Part 1 of this three-part series of papers, we describe the hardware and software used for data-collection and metadata documentation. We made changes to the data acquisition system that aimed to reduce the system complexity, increase redundancy, and be as independent as possible from any network outages. Changes to facilitate these improvements were (1) switching to a PC/104-based computer runningmore » the National Center for Atmospheric Research (NCAR) In-Situ Data Acquisition Software (NIDAS) that saves the high-frequency data locally and over the network, and (2) time-tagging individual 10 Hz serial data samples using network time protocol (NTP) coupled to a GPS-based clock, providing a network-independent, accurate time base. Since making these improvements almost 2 years ago, the successful capture of high-rate data has been better than 99.98 %. Here, we also provide philosophical concepts that shaped our design of the data system and are applicable to many different types of environmental data collection.« less
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DOE Office of Scientific and Technical Information (OSTI.GOV)
Burns, Sean P.; Maclean, Gordon D.; Blanken, Peter D.
The Niwot Ridge Subalpine Forest AmeriFlux site (US-NR1) has been measuring eddy-covariance ecosystem fluxes of carbon dioxide, heat, and water vapor since 1 November 1998. Throughout this 17-year period there have been changes to the instrumentation and improvements to the data acquisition system. Here, in Part 1 of this three-part series of papers, we describe the hardware and software used for data-collection and metadata documentation. We made changes to the data acquisition system that aimed to reduce the system complexity, increase redundancy, and be as independent as possible from any network outages. Changes to facilitate these improvements were (1) switching to a PC/104-based computer runningmore » the National Center for Atmospheric Research (NCAR) In-Situ Data Acquisition Software (NIDAS) that saves the high-frequency data locally and over the network, and (2) time-tagging individual 10 Hz serial data samples using network time protocol (NTP) coupled to a GPS-based clock, providing a network-independent, accurate time base. Since making these improvements almost 2 years ago, the successful capture of high-rate data has been better than 99.98 %. Here, we also provide philosophical concepts that shaped our design of the data system and are applicable to many different types of environmental data collection.« less
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Controls on nitrogen flux in alpine/subalpine watersheds of Colorado
Campbell, Donald H.; Baron, Jill S.; Tonnessen, Kathy A.; Brooks, Paul D.; Schuster, Paul F.
2000-01-01
High‐altitude watersheds in the Front Range of Colorado show symptoms of advanced stages of nitrogen excess, despite having less nitrogen in atmospheric deposition than other regions where watersheds retain nitrogen. In two alpine/subalpine subbasins of the Loch Vale watershed, atmospheric deposition of NO3− plus NH4+ was 3.2–5.5 kg N ha−1, and watershed export was 1.8–3.9 kg N ha−1 for water years 1992–1997. Annual N export increased in years with greater input of N, but most of the additional N was retained in the watershed, indicating that parts of the ecosystem are nitrogen‐limited. Dissolved inorganic nitrogen (DIN) concentrations were greatest in subsurface water of talus landscapes, where mineralization and nitrification augment high rates of atmospheric deposition of N. Tundra landscapes had moderately high DIN concentrations, whereas forest and wetland landscapes had low concentrations, indicating little export of nitrogen from these landscapes. Between the two subbasins the catchment of Icy Brook had greater retention of nitrogen than that of Andrews Creek because of landscape and hydrologic characteristics that favor greater N assimilation in both the terrestrial and aquatic ecosystems. These results suggest that export of N from alpine/subalpine watersheds is caused by a combination of direct flushing of N from atmospheric deposition and release of N from ecosystem biogeochemical processes (N cycling). Sensitivity of alpine ecosystems in the western United States to atmospheric deposition of N is a function of landscape heterogeneity, hydrologic flow paths, and climatic extremes that limit primary productivity and microbial activity, which, in turn, control retention and release of nitrogen. Conceptual and mechanistic models of N excess that have been developed for forested ecosystems need to be modified in order to predict the response of alpine ecosystems to future changes in climate and atmospheric deposition of N.
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Carbon storage in mountainous headwater streams: The role of old-growth forest and logjams
NASA Astrophysics Data System (ADS)
Beckman, Natalie D.; Wohl, Ellen
2014-03-01
We measured wood piece characteristics and particulate organic matter (POM) in stored sediments in 30 channel-spanning logjams along headwater streams in the Colorado Front Range, USA. Logjams are on streams flowing through old-growth (>200 years), disturbed (<200 years, natural disturbance), or altered (<200 years, logged) subalpine conifer forest. We examined how channel-spanning logjams influence riverine carbon storage (measured as the total volatile carbon fraction of stored sediment and instream wood). Details of carbon storage associated with logjams reflect age and disturbance history of the adjacent riparian forest. A majority of the carbon within jams is stored as wood. Wood volume is significantly larger in old-growth and disturbed reaches than in altered reaches. Carbon storage also differs in relation to forest characteristics. Sediment from old-growth streams has significantly higher carbon content than altered streams. Volume of carbon stored in jam sediment correlates with jam wood volume in old-growth and disturbed forests, but not in altered forests. Forest stand age and wood volume within a jam explain 43% of the variation of carbon stored in jam sediment. First-order estimates of the amount of carbon stored within a stream reach show an order of magnitude difference between disturbed and altered reaches. Our first-order estimates of reach-scale riverine carbon storage suggest that the carbon per hectare stored in streams is on the same order of magnitude as the carbon stored as dead biomass in terrestrial subalpine forests of the region. Of particular importance, old-growth forest correlates with more carbon storage in rivers.
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NASA Astrophysics Data System (ADS)
Foster, A.; Shuman, J. K.; Shugart, H. H., Jr.; Dwire, K. A.; Fornwalt, P.; Sibold, J.; Negrón, J. F.
2016-12-01
Forests in the Rocky Mountains are a crucial part of the North American carbon budget, but increases in disturbances such as insect outbreaks and fire, in conjunction with climate change, threaten their vitality. Mean annual temperatures in the western United States have increased by 2°C since 1950 and the higher elevations are warming faster than the rest of the landscape. It is predicted that this warming trend will continue, and that by the end of this century, nearly 50% of the western US landscape will have climate profiles with no current analog within that region. Individual tree-based modeling allows various climate change scenarios and their effects on forest dynamics to be tested. We use an updated individual-based gap model, the University of Virginia Forest Model Enhanced (UVAFME) at a subalpine site in the southern Rocky Mountains. UVAFME has been quantitatively and qualitatively validated in the southern Rocky Mountains, and results show that UVAFME-output on size structure, biomass, and species composition compares reasonably to inventory data and descriptions of vegetation zonation and successional dynamics for the region. We perform a climate sensitivity test in which temperature is first increased linearly by 2°C over 100 years, stabilized for 200 years, cooled back to present climate values over 100 years, and again stabilized for 200 years. This test is conducted to determine what effect elevated temperatures may have on vegetation zonation, and how persistent the changes may be if the climate is brought back to its current state. Results show that elevated temperatures within the southern Rocky Mountains may lead to decreases in biomass and changes in species composition as species migrate upslope. These changes are also likely to be fairly persistent for at least one- to two-hundred years. The results from this study suggest that UVAFME and other individual-based gap models can be used to inform forest management and climate mitigation strategies for this vitally important region.
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NASA Astrophysics Data System (ADS)
Miller, M.; McKnight, D.; Alexander, K.
2006-12-01
We studied the impact of a sustained high elevation rain event in mid-summer on the biogeochemistry of dissolved organic material (DOM) in an aquatic ecosystem in the Colorado Front Range. In the Green Lakes Valley, an alpine-subalpine catchment, the hydrology is typically defined by two distinct periods: snowmelt and baseflow. Similarly, characterization of DOM by fluorescence spectroscopy and other methods shows that the source and chemical character of the DOM changes with the hydrology. Surface water samples were collected from the outlet of a small alpine lake as well as a stream site downstream of a larger subalpine lake from the initiation of snowmelt through late summer. Beginning on July 7th and ending on July 9th 2006 a continuous low intensity rain event produced approximately 9 cm of precipitation. The rain event increased discharge at the two sites to flow rates that were 2.5 fold greater than those observed during peak snowmelt. The fluorescence characteristics of the DOM as well as the percent fulvic acid contribution to the sample were reset to values similar to those observed during snowmelt at the alpine site but were relatively unaffected at the subalpine site. These results suggest that alpine ecosystems are more sensitive to hydrologic changes than subalpine ecosystems and residence times of the lakes in these systems may play an important role in regulating stream chemistry.
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Annual wet and dry deposition of sulfur and nitrogen in the Snowy Range, Wyoming
Karl Zeller; Debra Harrington; Al Riebau; Evgeny Donev
2000-01-01
The collocation of three national networked programs NADP, EPA's CASTNET, and the Forest Service's IMPROVE Module A, within a few hundred meters of each other in the pristine Medicine Bow forest of Wyoming has made it possible to assess the total amount of sulfur and nitrogen deposition, both wet and dry for this alpine/subalpine ecosystem. Additional sites...
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Kevin M. O' Neill; Jessica E. Fultz; Michael A. Ivie
2008-01-01
We examined the distribution of adult Buprestidae and Cerambycidae in the Tenderfoot Creek Experimental Forest in the Little Belt Mountains of central Montana, U.S.A., using pan traps and sweep samples on differnt species of flowering plants. Using several methods during multi-year (2001-2004), summer-long surveys, we documented the presence of adults of three species...
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Paula J. Fornwalt; Charles C. Rhoades; Robert M. Hubbard; Rebecca L. Harris; Akasha M. Faist; William D. Bowman
2018-01-01
Recent bark beetle outbreaks in western North American subalpine forests have prompted managers to salvage log some beetle-affected stands. We examined the short-term (i.e., two to three years post-treatment) consequences of such salvage logging on vascular understory plant (i.e., graminoid, forb, and shrub) communities. At 24 lodgepole pine (Pinus contorta) sites in...
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Yang, Fan; Yang, Wan Qin; Wu, Fu Zhong; Wang, Hui; Lan, Li Ying; Liu, Yu Wei; Guo, Cai Hong; Tan, Bo
2017-06-18
As a biocide to reduce soil and litter faunal populations in field experiments, naphthalene has been widely used in the study of ecological functions of soil fauna, but the non-target effects of naphthalene bring about enormous uncertainty to its application. In order to understand whether there were non-target effects of naphthalene in subalpine forest soil, soil in the subalpine forests of west Qinghai-Tibet Plateau was taken as study object. The short-term responses of soil respiration rate, nutrient content and enzyme activity to naphthalene were studied in microcosms. The results showed that soil respiration rate was significantly suppressed by application of naphthalene within 0-10 days, and then showed a significant promotion effect. Naphthalene significantly affected the dynamics of soil NH 4 + -N and NO 3 - -N contents. With application of naphthalene, the highest contents of NH 4 + -N and NO 3 - -N occurred at the 3rd and 7th day, respectively. But they were observed at the 45th and 52nd day with no-naphthalene, respectively. Moreover, soil dissolved carbon content in the naphthalene microcosms showed a sharp increase and then decrease dynamic at the 3rd day, while small change was detected in the no-naphthalene microcosms. Dissolved nitrogen content in both the naphthalene and no-naphthalene microcosms showed an increase at first and then decreased subsequently during the study period. Similar dynamics were found for the soil enzyme activities in both the naphthalene and no-naphthalene microcosms. The highest activities of urease, nitrate reductase and nitrite reductase in both the naphthalene and no-naphthalene microcosms were at the 45th, 38th and 10th day, respectively. In addition, the interaction of naphthalene treatment and sampling time had significant effects on soil respiration rate, the contents of NH 4 + -N, NO 3 - -N and dissolved nitrogen, but had no significant effects on soil dissolved carbon content, and the activities of invertase, nitrate reductase and nitrite reductase. In a short time, the non-target effect of naphthalene as a biocide to reduce soil fauna abundance might have an important influence on the soil nitrogen cycling in subalpine forest of western Sichuan.
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Takahashi, Koichi; Tanaka, Saeka
2016-11-01
This study examined how habitat filtering and limiting similarity affect species assemblages of alpine and subalpine plant communities along a slope gradient on Mt. Norikura in central Japan. Plant traits (plant height, individual leaf area, specific leaf area (SLA), leaf linearity, leaf nitrogen and chlorophyll concentrations) and abiotic environmental factors (elevation, slope inclination, ground surface texture, soil water, soil pH, soil nutrient concentrations of NH 4 -N and NO 3 -N) were examined. The metrics of variance, range, kurtosis and the standard deviation of neighbor distance divided by the range of traits present (SDNDr) were calculated for each plant trait to measure trait distribution patterns. Limiting similarity was detected only for chlorophyll concentration. By contrast, habitat filtering was detected for individual leaf area, SLA, leaf linearity, chlorophyll concentration. Abiotic environmental factors were summarized by the principal component analysis (PCA). The first PCA axis positively correlated with elevation and soil pH, and negatively correlated with sand cover, soil water, NH 4 -N and NO 3 -N concentrations. High values of the first PCA axis represent the wind-exposed upper slope with lower soil moisture and nutrient availabilities. Plant traits changed along the first PCA axis. Leaf area, SLA and chlorophyll concentration decreased, and leaf linearity increased with the first PCA axis. This study showed that the species assemblage of alpine and subalpine plants was determined mainly by habitat filtering, indicating that abiotic environmental factors are more important for species assemblage than interspecific competition. Therefore, only species adapting to abiotic environments can distribute to these environments.
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Yi, C.; Monson, Russell K.; Zhai, Z.; Anderson, D.E.; Lamb, B.; Allwine, G.; Turnipseed, A.A.; Burns, Sean P.
2005-01-01
The nocturnal drainage flow of air causes significant uncertainty in ecosystem CO2, H2O, and energy budgets determined with the eddy covariance measurement approach. In this study, we examined the magnitude, nature, and dynamics of the nocturnal drainage flow in a subalpine forest ecosystem with complex terrain. We used an experimental approach involving four towers, each with vertical profiling of wind speed to measure the magnitude of drainage flows and dynamics in their occurrence. We developed an analytical drainage flow model, constrained with measurements of canopy structure and SF6 diffusion, to help us interpret the tower profile results. Model predictions were in good agreement with observed profiles of wind speed, leaf area density, and wind drag coefficient. Using theory, we showed that this one-dimensional model is reduced to the widely used exponential wind profile model under conditions where vertical leaf area density and drag coefficient are uniformly distributed. We used the model for stability analysis, which predicted the presence of a very stable layer near the height of maximum leaf area density. This stable layer acts as a flow impediment, minimizing vertical dispersion between the subcanopy air space and the atmosphere above the canopy. The prediction is consistent with the results of SF6 diffusion observations that showed minimal vertical dispersion of nighttime, subcanopy drainage flows. The stable within-canopy air layer coincided with the height of maximum wake-to-shear production ratio. We concluded that nighttime drainage flows are restricted to a relatively shallow layer of air beneath the canopy, with little vertical mixing across a relatively long horizontal fetch. Insight into the horizontal and vertical structure of the drainage flow is crucial for understanding the magnitude and dynamics of the mean advective CO2 flux that becomes significant during stable nighttime conditions and are typically missed during measurement of the turbulent CO2 flux. The model and interpretation provided in this study should lead to research strategies for the measurement of these advective fluxes and their inclusion in the overall mass balance for CO2 at this site with complex terrain. Copyright 2005 by the American Geophysical Union.
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NASA Astrophysics Data System (ADS)
Yi, Chuixiang; Monson, Russell K.; Zhai, Zhiqiang; Anderson, Dean E.; Lamb, Brian; Allwine, Gene; Turnipseed, Andrew A.; Burns, Sean P.
2005-11-01
The nocturnal drainage flow of air causes significant uncertainty in ecosystem CO2, H2O, and energy budgets determined with the eddy covariance measurement approach. In this study, we examined the magnitude, nature, and dynamics of the nocturnal drainage flow in a subalpine forest ecosystem with complex terrain. We used an experimental approach involving four towers, each with vertical profiling of wind speed to measure the magnitude of drainage flows and dynamics in their occurrence. We developed an analytical drainage flow model, constrained with measurements of canopy structure and SF6 diffusion, to help us interpret the tower profile results. Model predictions were in good agreement with observed profiles of wind speed, leaf area density, and wind drag coefficient. Using theory, we showed that this one-dimensional model is reduced to the widely used exponential wind profile model under conditions where vertical leaf area density and drag coefficient are uniformly distributed. We used the model for stability analysis, which predicted the presence of a very stable layer near the height of maximum leaf area density. This stable layer acts as a flow impediment, minimizing vertical dispersion between the subcanopy air space and the atmosphere above the canopy. The prediction is consistent with the results of SF6 diffusion observations that showed minimal vertical dispersion of nighttime, subcanopy drainage flows. The stable within-canopy air layer coincided with the height of maximum wake-to-shear production ratio. We concluded that nighttime drainage flows are restricted to a relatively shallow layer of air beneath the canopy, with little vertical mixing across a relatively long horizontal fetch. Insight into the horizontal and vertical structure of the drainage flow is crucial for understanding the magnitude and dynamics of the mean advective CO2 flux that becomes significant during stable nighttime conditions and are typically missed during measurement of the turbulent CO2 flux. The model and interpretation provided in this study should lead to research strategies for the measurement of these advective fluxes and their inclusion in the overall mass balance for CO2 at this site with complex terrain.
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Patricia E. Maloney; Detlev R. Vogler; Camille E. Jensen; Annette Delfino Mix
2012-01-01
For over a century, white pine blister rust (WPBR), caused by the introduced fungal pathogen, Cronartium ribicola J.C. Fisch., has affected white pine (Subgenus Strobus) individuals, populations, and associated forest communities in North America. We surveyed eight populations of whitebark pine (Pinus albicaulis Engelm.) across a range of environmental conditions in...
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NASA Astrophysics Data System (ADS)
Raczka, B. M.; Bowling, D. R.; Lin, J. C.; Lee, J. E.; Yang, X.; Duarte, H.; Zuromski, L.
2017-12-01
Forests of the Western United States are prone to drought, temperature extremes, forest fires and insect infestation. These disturbance render carbon stocks and land-atmosphere carbon exchanges highly variable and vulnerable to change. Regional estimates of carbon exchange from terrestrial ecosystem models are challenged, in part, by a lack of net ecosystem exchange observations (e.g. flux towers) due to the complex mountainous terrain. Alternatively, carbon estimates based on light use efficiency models that depend upon remotely-sensed greenness indices are challenged due to a weak relationship with GPP during the winter season. Recent advances in the retrieval of remotely sensed solar induced fluorescence (SIF) have demonstrated a strong seasonal relationship between GPP and SIF for deciduous, grass and, to a lesser extent, conifer species. This provides an important opportunity to use remotely-sensed SIF to calibrate terrestrial ecosystem models providing a more accurate regional representation of biomass and carbon exchange across mountainous terrain. Here we incorporate both leaf-level fluorescence and leaf-to-canopy radiative transfer represented by the SCOPE model into CLM 4.5 (CLM-SIF). We simulate canopy level fluorescence at a sub-alpine forest site (Niwot Ridge, Colorado) and test whether these simulations reproduce remotely-sensed SIF from a satellite (GOME2). We found that the average peak SIF during the growing season (yrs 2007-2013) was similar between the model and satellite observations (within 15%); however, simulated SIF during the winter season was significantly greater than the satellite observations (5x higher). This implies that the fluorescence yield is overestimated by the model during the winter season. It is important that the modeled representation of seasonal fluorescence yield is improved to provide an accurate seasonal representation of SIF across the Western United States.
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NASA Technical Reports Server (NTRS)
Potter, Christopher
2015-01-01
Landsat imagery was analyzed to understand changes in subalpine forest stands since the mid-1980s in the Sierra-Nevada region of California. At locations where long-term plot measurements have shown that stands are becoming denser in the number of small tree stems (compared to the early 1930s), the 30-year analysis of Landsat greenness index (NDVI) indicated that no consistent increases in canopy leaf cover have occurred at these same locations since the mid-1980s. Interannual variations in stand NDVI closely followed snow accumulation amounts recorded at nearby stations. In contrast, at eastern Sierra whitebark pine stand locations where it has been observed that widespread tree mortality has occurred, decreasing NDVI trends over the past 5-10 years were consistent with rapid loss of forest canopy cover. Landsat imagery was further analyzed to understand patterns of post-wildfire vegetation recovery, focusing on high burn severity (HBS) patches within burned areas dating from the late 1940s. Analysis of landscape metrics showed that the percentage of total HBS area comprised by the largest patch of recovered woody cover was relatively small in all fires that occurred since 1995, but increased rapidly with time since fire. Patch complexity of recovered woody cover decreased notably after more than 50 years of regrowth, but was not readily associated with time for fires that occurred since the mid 1990s. The aggregation level of patches with recovery of woody cover increased steadily with time since fire. The study approach using satellite remote sensing can be expanded to assess the consequences of stand-replacing wildfires in all forests of the region.
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Calder, W John; Shuman, Bryan
2017-10-01
Ecosystems may shift abruptly when the effects of climate change and disturbance interact, and landscapes with regularly patterned vegetation may be especially vulnerable to abrupt shifts. Here we use a fossil pollen record from a regularly patterned ribbon forest (alternating bands of forests and meadows) in Colorado to examine whether past changes in wildfire and climate produced abrupt vegetation shifts. Comparing the percentages of conifer pollen with sedimentary δ 18 O data (interpreted as an indicator of temperature or snow accumulation) indicates a first-order linear relationship between vegetation composition and climate change with no detectable lags over the past 2,500 yr (r = 0.55, P < 0.001). Additionally, however, we find that the vegetation changed abruptly within a century of extensive wildfires, which were recognized in a previous study to have burned approximately 80% of the surrounding 1,000 km 2 landscape 1,000 yr ago when temperatures rose ~0.5°C. The vegetation change was larger than expected from the effects of climate change alone. Pollen assemblages changed from a composition associated with closed subalpine forests to one similar to modern ribbon forests. Fossil pollen assemblages then remained like those from modern ribbon forests for the following ~1,000 yr, providing a clear example of how extensive disturbances can trigger persistent new vegetation states and alter how vegetation responds to climate. © 2017 by the Ecological Society of America.
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Oboyski, P.T.; Slotterback, J.W.; Banko, P.C.
2004-01-01
Alien parasitic wasps, including accidental introductions and purposefully released biological control agents, have been implicated in the decline of native Hawaiian Lepidoptera. Understanding the potential impacts of alien wasps requires knowledge of ecological parameters that influence parasitism rates for species in their new environment. Sophora seed-feeding Cydia spp. (Lepidoptera: Tortricidae) were surveyed for larval parasitoids to determine how native and alien wasps are partitioned over an elevation gradient (2200-2800 m) on Hawaii Island, Hawaii. Parasitism rate of native Euderus metallicus (Eulophidae) increased with increased elevation, while parasitism rate by immigrant Calliephialtes grapholithae (Ichneumonidae) decreased. Parasitism by Pristomerus hawaiiensis (Ichneumonidae), origins uncertain, also decreased with increased elevation. Two other species, Diadegma blackburni (Ichneumonidae), origins uncertain, and Brasema cushmani (Eupelmidae), a purposefully introduced biological control agent for pepper weevil, did not vary significantly with elevation. Results are contrasted with a previous study of this system with implications for the conservation of an endangered bird species that feed on Cydia larvae. Interpretation of results is hindered by lack of knowledge of autecology of moths and wasps, origins, phylogeny, systematics, competitive ability, and physiological limitations of each wasp species. These factors should be incorporated into risk analysis for biological control introductions and invasive species programs. ?? 2004 Kluwer Academic Publishers.
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[Canopy interception of sub-alpine dark coniferous communities in western Sichuan, China].
Lü, Yu-liang; Liu, Shi-rong; Sun, Peng-sen; Liu, Xing-liang; Zhang, Rui-pu
2007-11-01
Based on field measurements of throughfall and stemflow in combination with climatic data collected from the meteorological station adjacent to the studied sub-alpine dark coniferous forest in Wolong, Sichuan Province, canopy interception of sub-alpine dark coniferous forests was analyzed and modeled at both stand scale and catchment scale. The results showed that monthly interception rate of Fargesia nitida, Bashania fangiana--Abies faxoniana old-growth ranged from 33% Grass to 72%, with the average of 48%. In growing season, there was a linear or powerful or exponential relationship between rainfall and interception an. a negative exponential relationship between rainfall and interception rate. The mean maximum canopy interception by the vegetation in the catchment of in.44 km was 1.74 ment and the significant differences among the five communities occurred in the following sequence: Moss-Fargesia nitida, Bashan afanglana-A. faxoniana stand > Grass-F. nitida, B. fangiana-A. faxoniana stand > Moss-Rhododendron spp.-A. faxoniana stand > Grass-Rh. spp.-A. faxoniana stand > Rh. spp. shrub. In addition, a close linear relationship existed between leaf area index (LAI) and maximum canopy interception. The simulated value of canopy interception rate, maximum canopy interception rate and addition interception rate of the vegetation in the catchment were 39%, 25% and 14%, respectively. Simulation of the canopy interception model was better at the overall growing season scale, that the mean relative error was 9%-14%.
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NASA Astrophysics Data System (ADS)
Liang, S.; Hurteau, M. D.; Westerling, A. L.
2014-12-01
The Sierra Nevada Mountains are occupied by a diversity of forest types that sort by elevation. The interaction of changing climate and altered disturbance regimes (e.g. fire) has the potential to drive changes in forest distribution as a function of species-specific response. Quantifying the effects of these drivers on species distributions and productivity under future climate-fire interactions is necessary for informing mitigation and adaptation efforts. In this study, we assimilated forest inventory and soil survey data and species life history traits into a landscape model, LANDIS-II, to quantify the response of forest dynamics to the interaction of climate change and large wildfire frequency in the Sierra Nevada. We ran 100-year simulations forced with historical climate and climate projections from three models (GFDL, CNRM and CCSM3) driven by the A2 emission scenario. We found that non-growing season NPP is greatly enhanced by 15%-150%, depending on the specific climate projection. The greatest increase occurs in subalpine forests. Species-specific response varied as a function of life history characteristics. The distribution of drought and fire-tolerant species, such as ponderosa pine, expanded by 7.3-9.6% from initial conditions, while drought and fire-intolerant species, such as white fir, showed little change in the absence of fire. Changes in wildfire size and frequency influence species distributions by altering the successional stage of burned patches. The range of responses to different climate models demonstrates the sensitivity of these forests to climate variability. The scale of climate projections relative to the scale of forest simulations presents a source of uncertainty, particularly at the ecotone between forest types and for identifying topographically mediated climate refugia. Improving simulations will likely require higher resolution climate projections.
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Tree root dynamics in montane and sub-alpine mixed forest patches.
Wang, Y; Kim, J H; Mao, Z; Ramel, M; Pailler, F; Perez, J; Rey, H; Tron, S; Jourdan, C; Stokes, A
2018-02-28
The structure of heterogeneous forests has consequences for their biophysical environment. Variations in the local climate significantly affect tree physiological processes. We hypothesize that forest structure also alters tree root elongation and longevity through temporal and spatial variations in soil temperature and water potential. We installed rhizotrons in paired vegetation communities of closed forest (tree islands) and open patches (canopy gaps), along a soil temperature gradient (elevations of 1400, 1700 and 2000 m) in a heterogeneous mixed forest. We measured the number of growing tree roots, elongation and mortality every month over 4 years. The results showed that the mean daily root elongation rate (RER) was not correlated with soil water potential but was significantly and positively correlated with soil temperature between 0 and 8 °C only. The RER peaked in spring, and a smaller peak was usually observed in the autumn. Root longevity was dependent on altitude and the season in which roots were initiated, and root diameter was a significant factor explaining much of the variability observed. The finest roots usually grew faster and had a higher risk of mortality in gaps than in closed forest. At 2000 m, the finest roots had a higher risk of mortality compared with the lower altitudes. The RER was largely driven by soil temperature and was lower in cold soils. At the treeline, ephemeral fine roots were more numerous, probably in order to compensate for the shorter growing season. Differences in soil climate and root dynamics between gaps and closed forest were marked at 1400 and 1700 m, but not at 2000 m, where canopy cover was more sparse. Therefore, heterogeneous forest structure and situation play a significant role in determining root demography in temperate, montane forests, mostly through impacts on soil temperature.
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Zhang, Jin-Tun; Xiang, ChunLing; Li, Min
2012-02-01
Subalpine meadows in the Dongling Mountains (located at E115º26'-115º40', N40º00'-40º05') of Beijing, China are important for tourism and the provision of ecosystem services. However, because of poor management serious degradation has occurred on these subalpine meadows. The aim of this paper is to present a quantitative analysis of effects of tourism disturbance and topography on the status and diversity of montane meadow communities and to provide direction for improved management. Sixty quadrats of 2 × 2 m(2) along 10 transects were set up to collect data on site characteristics and vegetation status. The relationships between community composition and structure, species diversity, and tourism disturbance and topographic variables were analyzed by multivariate methods (TWINSPAN and CCA). The results showed that eight meadow communities were identified by TWINSPAN. Most of them were seriously degraded. The first CCA axis identified an elevation and tourism disturbance intensity gradient, which illustrated that tourism disturbance and elevation were most important factors influencing meadow types, composition and structure. Some resistant species and response species to tourism disturbance were identified and can be used as indicator species of tourism disturbance. Species richness, heterogeneity and evenness were closely related to tourism disturbance and elevation. It is concluded that tourism disturbance must be controlled to enable grassland rehabilitation to occur in the meadows. Measures of effective management of the meadows were discussed.
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Extreme climate events counteract the effects of climate and land-use changes in Alpine treelines
Barros, Ceres; Guéguen, Maya; Douzet, Rolland; Carboni, Marta; Boulangeat, Isabelle; Zimmermann, Niklaus E.; Münkemüller, Tamara; Thuiller, Wilfried
2017-01-01
Summary 1. Climate change and extreme events, such as drought, threaten ecosystems worldwide and in particular mountain ecosystems, where species often live at their environmental tolerance limits. In the European Alps, plant communities are also influenced by land-use abandonment leading to woody encroachment of subalpine and alpine grasslands. 2. In this study, we explored how the forest–grassland ecotone of Alpine treelines will respond to gradual climate warming, drought events and land-use change in terms of forest expansion rates, taxonomic diversity and functional composition. We used a previously validated dynamic vegetation model, FATE-HD, parameterised for plant communities in the Ecrins National Park in the French Alps. 3. Our results showed that intense drought counteracted the forest expansion at higher elevations driven by land-use abandonment and climate change, especially when combined with high drought frequency (occurring every 2 or less than 2 years). 4. Furthermore, intense and frequent drought accelerated the rates of taxonomic change and resulted in overall higher taxonomic spatial heterogeneity of the ecotone than would be expected under gradual climate and land-use changes only. 5. Synthesis and applications. The results from our model show that intense and frequent drought counteracts forest expansion driven by climate and land-use changes in the forest–grassland ecotone of Alpine treelines. We argue that land-use planning must consider the effects of extreme events, such as drought, as well as climate and land-use changes, since extreme events might interfere with trends predicted under gradual climate warming and agricultural abandonment. PMID:28670002
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Effect of prior disturbances on the extent and severity of wildfire in Colorado subalpine forests.
Kulakowski, Dominik; Veblen, Thomas T
2007-03-01
Disturbances are important in creating spatial heterogeneity of vegetation patterns that in turn may affect the spread and severity of subsequent disturbances. Between 1997 and 2002 extensive areas of subalpine forests in northwestern Colorado were affected by a blowdown of trees, bark beetle outbreaks, and salvage logging. Some of these stands were also affected by severe fires in the late 19th century. During a severe drought in 2002, fires affected extensive areas of these subalpine forests. We evaluated and modeled the extent and severity of the 2002 fires in relation to these disturbances that occurred over the five years prior to the fires and in relation to late 19th century stand-replacing fires. Occurrence of disturbances prior to 2002 was reconstructed using a combination of tree-ring methods, aerial photograph interpretation, field surveys, and geographic information systems (GIS). The extent and severity of the 2002 fires were based on the normalized difference burn ratio (NDBR) derived from satellite imagery. GIS and classification trees were used to analyze the effects of prefire conditions on the 2002 fires. Previous disturbance history had a significant influence on the severity of the 2002 fires. Stands that were severely blown down (> 66% trees down) in 1997 burned more severely than other stands, and young (approximately 120 year old) postfire stands burned less severely than older stands. In contrast, prefire disturbances were poor predictors of fire extent, except that young (approximately 120 years old) postfire stands were less extensively burned than older stands. Salvage logging and bark beetle outbreaks that followed the 1997 blowdown (within the blowdown as well as in adjacent forest that was not blown down) did not appear to affect fire extent or severity. Conclusions regarding the influence of the beetle outbreaks on fire extent and severity are limited, however, by spatial and temporal limitations associated with aerial detection surveys of beetle activity. Thus, fire extent in these forests is largely independent of prefire disturbance history and vegetation conditions. In contrast, fire severity, even during extreme fire weather and in conjunction with a multiyear drought, is influenced by prefire stand conditions, including the history of previous disturbances.
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Holz, Andrés; Wood, Sam W; Veblen, Thomas T; Bowman, David M J S
2015-01-01
Athrotaxis cupressoides is a slow-growing and long-lived conifer that occurs in the subalpine temperate forests of Tasmania, a continental island to the south of Australia. In 1960-1961, human-ignited wildfires occurred during an extremely dry summer that killed many A. cupressoides stands on the high plateau in the center of Tasmania. That fire year, coupled with subsequent regeneration failure, caused a loss of ca. 10% of the geographic extent of this endemic Tasmanian forest type. To provide historical context for these large-scale fire events, we (i) collected dendroecological, floristic, and structural data, (ii) documented the postfire survival and regeneration of A. cupressoides and co-occurring understory species, and (iii) assessed postfire understory plant community composition and flammability. We found that fire frequency did not vary following the arrival of European settlers, and that A. cupressoides populations were able to persist under a regime of low-to-mid severity fires prior to the 1960 fires. Our data indicate that the 1960 fires were (i) of greater severity than previous fires, (ii) herbivory by native marsupials may limit seedling survival in both burned and unburned A. cupressoides stands, and (iii) the loss of A. cupressoides populations is largely irreversible given the relatively high fuel loads of postfire vegetation communities that are dominated by resprouting shrubs. We suggest that the feedback between regeneration failure and increased flammability will be further exacerbated by a warmer and drier climate causing A. cupressoides to contract to the most fire-proof landscape settings. © 2014 John Wiley & Sons Ltd.
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NASA Astrophysics Data System (ADS)
Barnhart, T. B.; Vukomanovic, J.; Bourgeron, P.; Molotch, N. P.
2017-12-01
Land-cover change at the alpine-subalpine interface has the potential to change the water balance of mountainous, snow-dominated catchments due to the influence of vegetation on blowing snow, effective precipitation, evapotranspiration, and other processes. Understanding how land-cover change will impact water resources in snow-dominated regions is of critical importance as these locations produce a disproportionate amount of runoff relative to their land area. We coupled the LANdscape DIsturbance and Succession (LANDIS-II) model with a spatially explicit, physics-based, watershed process model, the Regional Hydro-Ecologic Simulation System (RHESSys), to simulate land-cover change and its impact on the water balance in a 6.6 km2 headwater catchment that spans the alpine-subalpine transition on the Colorado Front Range. We simulated two potential futures of air temperature warming (+4 °C/century) to 2100: a) increased precipitation (+15%, MP) and b) decreased precipitation (-15%, LP). As the LANDIS-II model simulates forest succession in a stochastic manner, we use three LANDIS-II model runs each for the MP and LP future forcing conditions. For both MP and LP, the RHESSys forcing data set was updated to reflect the changes in precipitation and temperature used to generate the land-cover futures. Forest cover in the catchment increased from 72% in 2000 to 84% and 83% in 2050 and to 95% and 92% in 2100 for MP and LP, respectively. Somewhat surprisingly, this increase in forest cover led to mean increases in streamflow production of 9% for MP and 3% for LP in 2050. In 2100, mean streamflow production increased by 15% and 6% for the MP and LP scenarios, respectively. This is likely due to increases in effective precipitation as the catchment forested and blowing snow decreased. Indeed, catchment effective precipitation increased from 94% in 2000 to 97% and 99% in 2050 and 2100, respectively, for both MP and LP conditions. This result counters previous work as runoff production increased with forested area, highlighting the need to better understand the impacts of forest expansion on blowing snow and effective precipitation. Identifying the hydrologic response of mountainous areas to climate warming induced land-cover change is of critical importance due to the potential water resources impacts in downstream regions.
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Eagles-Smith, Collin A.; Herring, Garth; Johnson, Branden L.; Graw, Rick
2013-01-01
Mercury (Hg) is a globally distributed pollutant that poses considerable risks to human and wildlife health. Over the past 150 years since the advent of the industrial revolution, approximately 80 percent of global emissions have come from anthropogenic sources, largely fossil fuel combustion. As a result, atmospheric deposition of Hg has increased by up to 4-fold above pre-industrial times. Because of their isolation, remote high-elevation lakes represent unique environments for evaluating the bioaccumulation of atmospherically deposited Hg through freshwater food webs, as well as for evaluating the relative importance of Hg loading versus landscape influences on Hg bioaccumulation. The increase in Hg deposition to these systems over the past century, coupled with their limited exposure to direct anthropogenic disturbance make them useful indicators for estimating how changes in Hg emissions may propagate to changes in Hg bioaccumulation and ecological risk. In this study, we evaluated Hg concentrations in fishes of high-elevation, sub-alpine lakes in the Wallowa-Whitman National Forest in northeastern Oregon and western Idaho. Our goals were to (1) assess the magnitude of Hg contamination in small-catchment lakes to evaluate the risk of atmospheric Hg to human and wildlife health, (2) quantify the spatial variability in fish Hg concentrations, and (3) determine the ecological, limnological, and landscape factors that are best correlated with fish total mercury (THg) concentrations in these systems. Across the 28 study lakes, mean THg concentrations of resident salmonid fishes varied as much as 18-fold among lakes. Importantly, our top statistical model explained 87 percent of the variability in fish THg concentrations among lakes with four key landscape and limnological variables— catchment conifer density (basal area of conifers within a lake’s catchment), lake surface area, aqueous dissolved sulfate, and dissolved organic carbon. The basal area of conifers within a lake’s catchment was by far the most important variable explaining fish THg concentrations, with an increase in THg concentrations of more than 400 percent across the forest density spectrum. Across all study lakes, fish THg concentrations ranged from 0.004 to 0.438 milligrams per kilogram wet weight (mg/kg ww). Only a single individual fish sample exceeded the U.S. Environmental Protection Agency’s (USEPA) human health tissue residue criteria of 0.3 mg/kg ww. However, 54 percent of fish (N=177) exceeded the more stringent tissue residue criteria (0.04 mg/kg ww) adopted by the Oregon Department of Environmental Quality to better protect subsistence fishers. Additionally, 2 and 10 percent of fish exceeded levels associated with reduced common loon reproduction and behavior, respectively. Whereas 25 and 68 percent of fish sampled exceeded concentrations deemed protective of mink and kingfisher, respectively. These results suggest that THg concentrations may be present in these lakes at levels associated with ecological risk. It is important to note however, that accurate inference on potential impairment cannot be made within the context of this study design and further research is needed to better quantify these risks.
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NASA Astrophysics Data System (ADS)
Liew, P. M.; Lee, C. Y.; Kuo, C. M.
2006-10-01
The East Asian monsoon Holocene optimal period has been debated both about duration and whether conditions were a maximum in thermal conditions or in precipitation. In this study we show Holocene climate variability inferred by a forest reconstruction of a subalpine pollen sequence from peat bog deposits in central Taiwan, based on modern analogues of various altitudinal biomes in the region. A warmer interval occurred between 8 and 4 ka BP (calibrated 14C years) when the subtropical forests were more extensive. The Holocene thermal optimum is represented by an altitudinal tropical forest at 6.1-5.9 ka BP and 6.9 ka BP and only the latter was accompanied by wet conditions, indicating decoupling of thermal and precipitation mechanism in the middle Holocene. Abrupt and relative severe cold phases, shown by biome changes, occurred at about 11.2-11.0 ka BP; 7.5 ka BP; 7.2 ka BP; 7.1 ka BP; 5.2 ka BP, 5.0 ka BP and 4.9 ka BP. A spectral analysis of pollen of a relatively cold taxon — Salix, reveals that the time series is dominated by a 1500 yr periodicity and similar to the cold cycle reported in the marine records of Indian and western Pacific Oceans. The cold-warm conditions inferred by the change of forests show close relationship to solar energy in comparison with the production rate of Be-10.
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Gophers as geomorphic agents in the Colorado Front Range subalpine zone
NASA Astrophysics Data System (ADS)
Winchell, Eric W.; Anderson, Robert S.; Lombardi, Elizabeth M.; Doak, Daniel F.
2016-07-01
Gophers are significant geomorphic agents in many landscapes. We document activity of the northern pocket gopher (Thomomys talpoides) in two small subalpine meadows (1050-1800 m2) of the Front Range, Colorado, USA. We tracked locations and volumes of mounds and subnivean infilled tunnels over one year and probed the thickness of the biomantle within one meadow. We infer that only 5-7 gophers occupied each meadow, implying a gopher density of 28-67 ha- 1. Fractional areal coverage of the meadows by diggings suggests that within 49-95 years gophers would fully resurface the meadows. Annual volumes of excavated soil correspond to the equivalent of 1 mm of material spread evenly over the meadows. Probed meadow resistance depths reveal a pattern we interpret to be stone lines at roughly 15 cm depths; implied vertical turnover times are therefore roughly 150 years. These spatial and temporal patterns imply that gophers should be able to churn the biomantle on approximately century timescales and should fully resurface the meadow areas in similar timescales. These field data also contribute to an investigation of lateral sediment transport; given the local slope of the landscape, gopher-driven sediment transport within our two study sites suggests a landscape diffusivity of 0.008 m2y- 1. At no time do gophers occupy the forest. As evidenced by subnivean infilled tunnels, winter activity is restricted to the upslope (and hence upwind) meadow edges, which correspond to high snow cover and warm (> 0 °C) shallow subsurface soil temperatures. Subsequent activity expands downhill into the meadows and shows a distinct pulse of mound activity in late summer through early fall prior to snowfall. Local forest fire history has led to much more extensive meadows in the past, suggesting that the geomorphic influence of gophers in the landscape is much more widespread than the present distribution of meadows and may cover the entire subalpine region of the Front Range on millennial timescales.
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Beyond CO2 - Tackling the full greenhouse gas budget of a sub-alpine forest ecosystem
NASA Astrophysics Data System (ADS)
Burri, Susanne; Merbold, Lutz; Meier, Philip; Eugster, Werner; Hörtnagl, Lukas; Buchmann, Nina
2017-04-01
In order to tackle the full greenhouse gas (GHG) budgets of forest ecosystems, it is desirable but challenging to quantify the three major GHGs, i.e. CO2, CH4 and N2O simultaneously in-situ. At the long-term forest research site Davos (Candidate Class I Ecosystem Station within the Integrated Carbon Observation System - ICOS), we have recently installed a state-of-the-art measuring system simultaneously to observe the three GHGs on a high temporal resolution and both within and above the forest canopy. Thereby, we combine above-canopy eddy covariance flux measurements and forest floor chamber flux measurements (using five custom-made fully automated chambers). Both systems are connected to a quantum cascade laser absorption spectrometer (QCL, Aerodyne) and measurements are switched between three hours of above-canopy and one hour of forest floor GHG flux measurements. Using this approach, we will be able to study the full GHG budget as well as the dynamics of the individual fluxes on two vertical levels within the forest using a single instrument. The first results presented here will highlight the suitability of this promising tool for quantifying the full GHG budget of forest ecosystems.
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Tsujino, Riyou; Yumoto, Takakazu
2013-03-01
In order to clarify how vegetation types change along the environmental gradients in a cool temperate to sub-alpine mountainous zone and the determinant factors that define plant species richness, we established 360 plots (each 4 × 10 m) within which the vegetation type, species richness, elevation, topographic position index (TPI), slope inclination, and ground light index (GLI) of the natural vegetation were surveyed. Mean elevation, TPI, slope inclination, and GLI differed across vegetation types. Tree species richness was negatively correlated with elevation, whereas fern and herb species richness were positively correlated. Tree species richness was greater in the upper slope area than the lower slope area, whereas fern and herb species richness were greater in the lower slope area. Ferns and trees species richness were smaller in the open canopy, whereas herb species richness was greater in the open canopy. Vegetation types were determined firstly by elevation and secondary by topographic configurations, such as topographic position, and slope inclination. Elevation and topography were the most important factors affecting plant richness, but the most influential variables differed among plant life-form groups. Moreover, the species richness responses to these environmental gradients greatly differed among ferns, herbs, and trees.
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Huang, Wei; McDowell, William H.; Zou, Xiaoming; Ruan, Honghua; Wang, Jiashe; Li, Liguang
2013-01-01
Stream water dissolved organic carbon (DOC) correlates positively with soil organic carbon (SOC) in many biomes. Does this relationship hold in a small geographic region when variations of temperature, precipitation and vegetation are driven by a significant altitudinal gradient? We examined the spatial connectivity between concentrations of DOC in headwater stream and contents of riparian SOC and water-soluble soil organic carbon (WSOC), riparian soil C:N ratio, and temperature in four vegetation types along an altitudinal gradient in the Wuyi Mountains, China. Our analyses showed that annual mean concentrations of headwater stream DOC were lower in alpine meadow (AM) than in subtropical evergreen broadleaf forest (EBF), coniferous forest (CF), and subalpine dwarf forest (SDF). Headwater stream DOC concentrations were negatively correlated with riparian SOC as well as WSOC contents, and were unrelated to riparian soil C:N ratio. Our findings suggest that DOC concentrations in headwater streams are affected by different factors at regional and local scales. The dilution effect of higher precipitation and adsorption of soil DOC to higher soil clay plus silt content at higher elevation may play an important role in causing lower DOC concentrations in AM stream of the Wuyi Mountains. Our results suggest that upscaling and downscaling of the drivers of DOC export from forested watersheds when exploring the response of carbon flux to climatic change or other drivers must done with caution. PMID:24265737
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Talbot, S. S.; Shasby, M.B.; Bailey, T.N.
1985-01-01
A Landsat-based vegetation map was prepared for Kenai National Wildlife Refuge and adjacent lands, 2 million and 2.5 million acres respectively. The refuge lies within the middle boreal sub zone of south central Alaska. Seven major classes and sixteen subclasses were recognized: forest (closed needleleaf, needleleaf woodland, mixed); deciduous scrub (lowland and montane, subalpine); dwarf scrub (dwarf shrub tundra, lichen tundra, dwarf shrub and lichen tundra, dwarf shrub peatland, string bog/wetlands); herbaceous (graminoid meadows and marshes); scarcely vegetated areas ; water (clear, moderately turbid, highly turbid); and glaciers. The methodology employed a cluster-block technique. Sample areas were described based on a combination of helicopter-ground survey, aerial photo interpretation, and digital Landsat data. Major steps in the Landsat analysis involved: preprocessing (geometric connection), spectral class labeling of sample areas, derivation of statistical parameters for spectral classes, preliminary classification of the entree study area using a maximum-likelihood algorithm, and final classification through ancillary information such as digital elevation data. The vegetation map (scale 1:250,000) was a pioneering effort since there were no intermediate-sclae maps of the area. Representative of distinctive regional patterns, the map was suitable for use in comprehensive conservation planning and wildlife management.
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Curtin, G.C.; King, H.D.; Mosier, E.L.
1974-01-01
Exudates from conifer trees, presumably consisting largely of volatile materials, were sampled at 19 subalpine localitites in Colorado and Idaho where anomalous amounts of several metals were determined in vegetation and mull during previous geochemical testing. The trees sampled were lodgepole pine (Pinus contorta), Engelmann spruce (Picea engelmannii) and Douglas fir (Pseudotsuga menziesii). The condensed exudates were passed through No. 40 Whatman filters, and through 5-micron, 0.45-micron, and 0.05-micron average-pore-diameter membrane filters, evaporated to dryness, and each residue was ashed and analyzed by a semiquantitative spectrographic method. The ashed residues of the exudates contain lithium, beryllium, boron, sodium, magnesium, titanium, vanadium, chromium, manganese, iron, cobalt, nickel, copper, zinc, gallium, arsenic, strontium, yttrium, zirconium, molybdenum, silver, lead, bismuth, cadmium, tin, antimony, barium, and lanthanum. The presence of these elements suggests that volatile exudates from vegetation are a medium for the transport of elements in the biogeochemical cycle in subalpine environments. Thus, air sampling and analysis of aerosols derived from volatile exudates may be a useful tool in geochemical exploration. ?? 1974.
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Similarities in riparian bird communities among elevational zones in southeastern Wyoming
Deborah M. Finch
1986-01-01
I examined trends in bird species richness and overall bird abundance in riparian habitats among elevations varying from 6740 ft. to 9800 ft. in southeastern Wyoming. Bird species diversity ranged from a low of three bird species and 23 pairs in subalpine shrub willow habitat to a maximum of 21 species and 101 pairs in lowland cottonwood habitat. Bird communities were...
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Modeling and measuring snow for assessing climate change impacts in Glacier National Park, Montana
Fagre, Daniel B.; Selkowitz, David J.; Reardon, Blase; Holzer, Karen; Mckeon, Lisa L.
2002-01-01
A 12-year program of global change research at Glacier National Park by the U.S. Geological Survey and numerous collaborators has made progress in quantifying the role of snow as a driver of mountain ecosystem processes. Spatially extensive snow surveys during the annual accumulation/ablation cycle covered two mountain watersheds and approximately 1,000 km2 . Over 7,000 snow depth and snow water equivalent (SWE) measurements have been made through spring 2002. These augment two SNOTEL sites, 9 NRCS snow courses, and approximately 150 snow pit analyses. Snow data were used to establish spatially-explicit interannual variability in snowpack SWE. East of the Continental Divide, snowpack SWE was lower but also less variable than west of the Divide. Analysis of snowpacks suggest downward trends in SWE, a reduction in snow cover duration, and earlier melt-out dates during the past 52 years. Concurrently, high elevation forests and treelines have responded with increased growth. However, the 80 year record of snow from 3 NRCS snow courses reflects a strong influence from the Pacific Decadal Oscillation, resulting in 20-30 year phases of greater or lesser mean SWE. Coupled with the fine-resolution spatial snow data from the two watersheds, the ecological consequences of changes in snowpack can be empirically assessed at a habitat patch scale. This will be required because snow distribution models have had varied success in simulating snowpack accumulation/ablation dynamics in these mountain watersheds, ranging from R2=0.38 for individual south-facing forested snow survey routes to R2=0.95 when aggregated to the watershed scale. Key ecological responses to snowpack changes occur below the watershed scale, such as snow-mediated expansion of forest into subalpine meadows, making continued spatially-explicit snow surveys a necessity.
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Fire ecology and management of the major ecosystems of southern Utah
Sharon M. Hood; Melanie Miller
2007-01-01
This document provides managers with a literature synthesis of the historical conditions, current conditions, fire regime condition classes (FRCC), and recommended treatments for the major ecosystems in southern Utah. Sections are by ecosystems and include: 1) coniferous forests (ponderosa pine, mixed conifer, and Engelmann spruce-subalpine fir), 2) aspen, 3) pinyon-...
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Soil fauna and plant litter decomposition in tropical and subalpine forests
G. Gonzalez; T.R. Seastedt
2001-01-01
The decomposition of plant residues is influenced by their chemical composition, the physical-chemical environment, and the decomposer organisms. Most studies interested in latitudinal gradients of decomposition have focused on substrate quality and climate effects on decomposition, and have excluded explicit recognition of the soil organisms involved in the process....
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The importance of wilderness to whitebark pine research and management
Robert E. Keane
2000-01-01
Whitebark pine is a keystone species in upper subalpine forests of the northern Rocky Mountains, Cascades, and Sierra Nevada that has been declining because of recent mountain pine beetle and exotic blister rust epidemics, coupled with advancing succession resulting from fire exclusion. Whitebark pine and Wilderness have a mutually beneficial relationship because 1)...
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Seasonal differences in space use by Clark's Nutcrackers in the Cascade Range
Teresa J. Lorenz; Kimberly A. Sullivan
2009-01-01
Clark's Nutcrackers (Nucifraga columbiana) are important seed dispersers for at least ten species of conifer in western North America and are obligate mutualists for the whitebark pine (Pinus albicaulis), a subalpine tree. Despite the important role they play in forest regeneration, space use by nutcrackers has not been...
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The perspectives, information and conclusions conveyed in research project abstracts, progress reports, final reports, journal abstracts and journal publications convey the viewpoints of the principal investigator and may not represent the views and policies of ORD and EPA. Concl...
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Wood decay fungi of subalpine conifer forests
Jessie A. Glaeser; Kevin T. Smith
2016-01-01
One of the fundamental skills needed for hazard tree assessment is the evaluation of decay. This may be a difficult task as we usually only use external symptoms (wounds, basal swellings, decayed branch stubs), signs (mushrooms, fungal crusts or brackets) or mechanical/indirect sampling methods (drilling, electrical or sonic resistance) to estimate the amount of sound...
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Tree squirrel habitat selection and predispersal seed predation in a declining subalpine conifer
Shawn T. McKinney; Carl E. Fiedler
2009-01-01
Differential responses by species to modern perturbations in forest ecosystems may have undesirable impacts on plant-animal interactions. If such disruptions cause declines in a plant species without corresponding declines in a primary seed predator, the effects on the plant could be exacerbated. We examined one such interaction between Pinus...
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Spatial patterns of recreational impact on experimental campsites
David N. Cole; Christopher A. Monz
2004-01-01
Management of camping impacts in protected areas worldwide is limited by inadequate understanding of spatial patterns of impact and attention to spatial management strategies. Spatial patterns of campsite impact were studied in two subalpine plant communities in the Wind River Mountains, Wyoming, USA (a forest and a meadow). Response to chronic disturbance and recovery...
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NASA Astrophysics Data System (ADS)
Gray, Christopher M.; Monson, Russell K.; Fierer, Noah
2014-04-01
Nonmethane biogenic volatile organic compounds (BVOCs) play key roles in the atmosphere, where they can influence a wide range of chemical processes, and in soils, where they can alter the rates of biogeochemical cycles and impact the growth of plants and soil organisms. However, the diversity and quantities of BVOCs released from or taken up by soils remain poorly characterized as do the biotic and abiotic controls on these fluxes. Here we used proton transfer reaction mass spectrometry to quantify BVOC flux rates from soils with and without active root systems in a subalpine coniferous forest. The total measured BVOC flux averaged 102 nmol m-2 h-1 (an estimated 2.0 µg-C m-2 h-1). The individual BVOCs with the highest net emissions from soil included monoterpenes and methanol (averaging 646 and 641 ng-C m-2 h-1, respectively) while soil represented a net sink of isoprene (-98 ng-C m-2 h-1) and formaldehyde (-37 ng-C m-2 h-1). Tree roots, directly or indirectly, contributed an average of 53% of the total carbon emitted from the soil as BVOCs, with methanol and acetaldehyde among those BVOCs most strongly associated with active root presence. The fluxes of most of the dominant BVOCs emitted from soil, including methanol, increased linearly with increasing temperature. Together the fluxes of certain BVOCs into or out of the forest floor (particularly methanol, isoprene, and monoterpenes) are likely relevant to ecosystem-level processes and belowground ecology, but these fluxes are highly variable and are strongly controlled by both root presence and soil abiotic conditions.
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NASA Astrophysics Data System (ADS)
Barber, A.; Millar, C.
2014-12-01
Documenting plant response to global climate change in sensitive zones, such as the alpine, is a major goal for global change biology. Basic information on alpine plant distribution by elevation and substrate provides a basis for anticipating which species may decline in a warming climate. The Global Observation Research Initiative in Alpine Environments (GLORIA) is a worldwide effort to document vegetation changes over time in alpine settings using permanent multi-summit plots. The California/Nevada group currently monitors seven permanent GLORIA target regions, composed of 29 summits in alpine and subalpine zones. Summits range in elevations from 2918m to 4325m on substrates including dolomite, granite, quartzite, and volcanics. High-resolution plant occurrence and cover data from the upper 10 meters of each summit are presented. Plants from our target regions can be divided into three groups: summit specialists found only on the highest peaks, alpine species found predominantly within the alpine zone, and broadly distributed species found in the alpine zone and below. Rock substrate and microsite soil development have a strong influence on plant communities and species richness. We present the first set of five-year resurvey and temperature data from 18 summits. We have documented some annual variation in species presence/absence at almost all sites, but no dramatic changes in total diversity. Consistent with the expectation of rising global temperatures, our soil temperature loggers have documented temperature increases at most of our sites. These data are a baseline for assessing bioclimatic shifts and future plant composition in California and Nevada's alpine zone.
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Shiau, Yo-Jin; Chen, Jenn-Shing; Chung, Tay-Lung; Tian, Guanglong; Chiu, Chih-Yu
2017-12-01
Soil organic carbon (SOC) and carbon (C) functional groups in different particle-size fractions are important indicators of microbial activity and soil decomposition stages under wildfire disturbances. This research investigated a natural Tsuga forest and a nearby fire-induced grassland along a sampling transect in Central Taiwan with the aim to better understand the effect of forest wildfires on the change of SOC in different soil particle scales. Soil samples were separated into six particle sizes and SOC was characterized by solid-state 13 C nuclear magnetic resonance spectroscopy in each fraction. The SOC content was higher in forest than grassland soil in the particle-size fraction samples. The O-alkyl-C content (carbohydrate-derived structures) was higher in the grassland than the forest soils, but the alkyl-C content (recalcitrant substances) was higher in forest than grassland soils, for a higher humification degree (alkyl-C/O-alkyl-C ratio) in forest soils for all the soil particle-size fractions. High humification degree was found in forest soils. The similar aromaticity between forest and grassland soils might be attributed to the fire-induced aromatic-C content in the grassland that offsets the original difference between the forest and grassland. High alkyl-C content and humification degree and low C/N ratios in the fine particle-size fractions implied that undecomposed recalcitrant substances tended to accumulate in the fine fractions of soils.
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[Effects of altitudes on soil microbial biomass and enzyme activity in alpine-gorge regions.
Cao, Rui; Wu, Fu Zhong; Yang, Wan Qin; Xu, Zhen Feng; Tani, Bo; Wang, Bin; Li, Jun; Chang, Chen Hui
2016-04-22
In order to understand the variations of soil microbial biomass and soil enzyme activities with the change of altitude, a field incubation was conducted in dry valley, ecotone between dry valley and mountain forest, subalpine coniferous forest, alpine forest and alpine meadow from 1563 m to 3994 m of altitude in the alpine-gorge region of western Sichuan. The microbial biomass carbon and nitrogen, and the activities of invertase, urease and acid phosphorus were measured in both soil organic layer and mineral soil layer. Both the soil microbial biomass and soil enzyme activities showed the similar tendency in soil organic layer. They increased from 2158 m to 3028 m, then decreased to the lowest value at 3593 m, and thereafter increased until 3994 m in the alpine-gorge region. In contrast, the soil microbial biomass and soil enzyme activities in mineral soil layer showed the trends as, the subalpine forest at 3028 m > alpine meadow at 3994 m > montane forest ecotone at 2158 m > alpine forest at 3593 m > dry valley at 1563 m. Regardless of altitudes, soil microbial biomass and soil enzyme activities were significantly higher in soil organic layer than in mineral soil layer. The soil microbial biomass was significantly positively correlated with the activities of the measured soil enzymes. Moreover, both the soil microbial biomass and soil enzyme activities were significantly positively correlated with soil water content, organic carbon, and total nitrogen. The activity of soil invertase was significantly positively correlated with soil phosphorus content, and the soil acid phosphatase was so with soil phosphorus content and soil temperature. In brief, changes in vegetation and other environmental factors resulting from altitude change might have strong effects on soil biochemical properties in the alpine-gorge region.
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Xiong, Li; Xu, Zhen-Feng; Wu, Fu-Zhong; Yang, Wan-Qin; Yin, Rui; Li, Zhi-Ping; Gou, Xiao-Lin; Tang, Shi-Shan
2014-05-01
This study characterized the dynamics of the activities of urease, nitrate reductase and nitrite reductase in both soil organic layer and mineral soil layer under three depths of snow pack (deep snowpack, moderate snowpack and shallow snowpack) over the three critical periods (snow formed period, snow stable period, and snow melt period) in the subalpine Abies faxoniana forest of western Sichuan in the winter of 2012 and 2013. Throughout the winter, soil temperature under deep snowpack increased by 46.2% and 26.2%, respectively in comparison with moderate snowpack and shallow snowpack. In general, the three nitrogen-related soil enzyme activities under shallow snowpack were 0.8 to 3.9 times of those under deep snowpack during the winter. In the beginning and thawing periods of seasonal snow pack, shallow snowpack significantly increased the activities of urease, nitrate and nitrite reductase enzyme in both soil organic layer and mineral soil layer. Although the activities of the studied enzymes in soil organic layer and mineral soil layer were observed to be higher than those under deep- and moderate snowpacks in deep winter, no significant difference was found under the three snow packs. Meanwhile, the effects of snowpack on the activities of the measured enzymes were related with season, soil layer and enzyme type. Significant variations of the activities of nitrogen-related enzymes were found in three critical periods over the winter, and the three measured soil enzymes were significantly higher in organic layer than in mineral layer. In addition, the activities of the three measured soil enzymes were closely related with temperature and moisture in soils. In conclusion, the decrease of snow pack induced by winter warming might increase the activities of soil enzymes related with nitrogen transformation and further stimulate the process of wintertime nitrogen transformation in soils of the subalpine forest.
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Factors related to the recovery of subalpine woodland on Mauna Kea, Hawaii
Hess, Steven C.; Banko, Paul C.; Brenner, Gregory J.; Jacobi, James D.
1999-01-01
We measured mature tree and sapling density, tree associations, crown size, age structure, recovery from ungulate browsing, and grass cover at four study sites in two types of subalpine woodland on Mauna Kea volcano, island of Hawaii. Beginning in 1981, introduced ungulates were reduced in number to allow regeneration of Sophora chrysophylla (mamane) in habitat supporting the endangered Hawaiian finch, Loxioides bailleui (palila). We found Sophora regeneration at all four study sites, but regeneration was higher in mixed species woodland with codominant Myoporum sandwicense (naio) than in areas where Sophora dominated. Regeneration of Myoporum was uniformly very low in comparison. Invasive grass cover, which suppresses Sophora germination, was highest in mid-elevation woodland where Sophora dominated. The distribution of mature and sapling Sophora were both related to study site, reflecting previous ungulate browsing and uneven recovery due to grasses. Densities of Sophora snags were not different among any of the sites, suggesting a more even distribution in the past. Selective browsing before ungulate reduction may have favored Myoporum over Sophora, leading to high densities of mature Myoporum in codominant woodland. After ungulate reduction, however, we found no pattern of competitive inhibition by Myoporum on regeneration of Sophora. Reduction of Myoporum is not likely to enhance habitat for Loxioides as much as supplemental planting of Sophora, grass control, and continued ungulate eradication. Mid-elevation Sophora woodland areas, where Loxioides forage and nest in high densities, would benefit the most from these management actions.
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NASA Astrophysics Data System (ADS)
Roth, T. R.; Nolin, A. W.
2016-12-01
Temperate forests modify snow evolution patterns both spatially and temporally relative to open areas. Dense, warm forests both impede snow accumulation through increased canopy snow interception and increase sub-canopy longwave energy inputs onto the snow surface. These process modifications vary in magnitude and duration depending on climatic, topographic and forest characteristics. Here we present results from a four year study of paired forested and open sites at three elevations, Low - 1150 m, Mid - 1325 m and High - 1465 m. Snowpacks are deeper and last up to 3-4 weeks longer at the Low and Mid elevation Open sites relative to the adjacent Forest sites. Conversely, at the High Forest site, snow is retained 2-4 weeks longer than the Open site. This change in snowpack depth and persistence is attributed to deposition patterns at higher elevations and forest structure differences that alter the canopy interception efficiency and the sub-canopy energy balance. Canopy interception efficiency (CIE) in the Low and Mid Forest sites, over the duration of the study were 79% and 76% of the total event snowfall, whereas CIE was 31% at the High Forest site. Longwave radiation in forested environments is the primary energy component across each elevation band due to the warm winter environment and forest presence, accounting for 82%, 88%, and 59% of the energy balance at the Low, Mid, and High Forest sites, respectively. High wind speeds in the High elevation Open site significantly increases the turbulent energy and creates preferential snowfall deposition in the nearby Forest site. These results show the importance of understanding the effects of forest cover on sub-canopy snowpack evolution and highlight the need for improved forest cover model representation to accurately predict water resources in maritime forests.
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Aiba, Shin-ichiro; Akutsu, Kosuke; Onoda, Yusuke
2013-01-01
Background and Aims Globally, conifer dominance is restricted to nutient-poor habitats in colder, drier or waterlogged environments, probably due to competition with angiosperms. Analysis of canopy structure is important for understanding the mechanism of plant coexistence in relation to competition for light. Most conifers are shade intolerant, and often have narrow, deep, conical crowns. In this study it is predicted that conifer-admixed forests have less distinct upper canopies and more undulating canopy surfaces than angiosperm-dominated forests. Methods By using a ground-based, portable light detection and ranging (LIDAR) system, canopy structure was quantified for old-growth evergreen rainforests with varying dominance of conifers along altitudinal gradients (200–3100 m a.s.l.) on tropical and sub-tropical mountains (Mount Kinabalu, Malaysian Borneo and Yakushima Island, Japan) that have different conifer floras. Key Results Conifers dominated at higher elevations on both mountains (Podocarpaceae and Araucariaceae on Kinabalu and Cupressaceae and Pinaceae on Yakushima), but conifer dominance also varied with soil/substrate conditions on Kinabalu. Conifer dominance was associated with the existence of large-diameter conifers. Forests with higher conifer dominance showed a canopy height profile (CHP) more skewed towards the understorey on both Kinabalu and Yakushima. In contrast, angiosperm-dominated forests had a CHP skewed towards upper canopy, except for lowland dipterocarp forests and a sub-alpine scrub dominated by small-leaved Leptospermum recurvum (Myrtaceae) on Kinabalu. Forests with a less dense upper canopy had more undulating outer canopy surfaces. Mixed conifer–angiosperm forests on Yakushima and dipterocarp forests on Kinabalu showed similar canopy structures. Conclusions The results generally supported the prediction, suggesting that lower growth of angiosperm trees (except L. recurvum on Kinabalu) in cold and nutrient-poor environments results in a sparser upper canopy, which allows shade-intolerant conifers to co-occur with angiosperm trees either as emergents or as codominants in the open canopy. PMID:24197751
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Aiba, Shin-ichiro; Akutsu, Kosuke; Onoda, Yusuke
2013-12-01
Globally, conifer dominance is restricted to nutient-poor habitats in colder, drier or waterlogged environments, probably due to competition with angiosperms. Analysis of canopy structure is important for understanding the mechanism of plant coexistence in relation to competition for light. Most conifers are shade intolerant, and often have narrow, deep, conical crowns. In this study it is predicted that conifer-admixed forests have less distinct upper canopies and more undulating canopy surfaces than angiosperm-dominated forests. By using a ground-based, portable light detection and ranging (LIDAR) system, canopy structure was quantified for old-growth evergreen rainforests with varying dominance of conifers along altitudinal gradients (200-3100 m a.s.l.) on tropical and sub-tropical mountains (Mount Kinabalu, Malaysian Borneo and Yakushima Island, Japan) that have different conifer floras. Conifers dominated at higher elevations on both mountains (Podocarpaceae and Araucariaceae on Kinabalu and Cupressaceae and Pinaceae on Yakushima), but conifer dominance also varied with soil/substrate conditions on Kinabalu. Conifer dominance was associated with the existence of large-diameter conifers. Forests with higher conifer dominance showed a canopy height profile (CHP) more skewed towards the understorey on both Kinabalu and Yakushima. In contrast, angiosperm-dominated forests had a CHP skewed towards upper canopy, except for lowland dipterocarp forests and a sub-alpine scrub dominated by small-leaved Leptospermum recurvum (Myrtaceae) on Kinabalu. Forests with a less dense upper canopy had more undulating outer canopy surfaces. Mixed conifer-angiosperm forests on Yakushima and dipterocarp forests on Kinabalu showed similar canopy structures. The results generally supported the prediction, suggesting that lower growth of angiosperm trees (except L. recurvum on Kinabalu) in cold and nutrient-poor environments results in a sparser upper canopy, which allows shade-intolerant conifers to co-occur with angiosperm trees either as emergents or as codominants in the open canopy.
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NASA Astrophysics Data System (ADS)
Beverly, D.; Speckman, H. N.; Ewers, B. E.
2017-12-01
Ecosystem-scale models often rely on root vulnerability or whole-plant conductance for simulating seasonal evapotranspiration declines via constraints of water uptake and vegetation mortality. Further, many of these ecosystem models rely on single, unvarying, hydraulic parameter estimates for modeling large areas. Ring-porous species have shown seasonal variability in root vulnerability (percent loss of conductivity; PLC) and whole-plant conductance (Kw) but simulations of coniferous forest typically rely on point measurements. This assumption for coniferous forest is not likely true because of seasonal variability caused by phenology and environmental stresses and the potential for cavitation fatigue is not considered. Moreover, many of these dynamics have only been considered for stems even though roots are often the most vulnerable segments of the pathway for conifers. We hypothesized that seasonally dynamic whole-plant conductance along hillslope gradients in coniferous forests are regulated by cavitation fatigue within the roots resulting in seasonal increases in vulnerability. To test the hypothesis, a subalpine mixed forest (3000 m.a.s.l) and montane forest (2550 m.a.s.l.) were monitored between 2015-2017 to quantify PLC and Kw along the hillslope gradients of 300 m and 50 m, respectively. Forest plots were instrumented with 35 Granier-type sapflow sensors. Seasonal sampling campaigns occurred to quantify PLC through centrifuge techniques and Kw through Darcy's law approximations with pre-dawn and diurnal leaf water potentials. Downslope roots exhibit a 33% decrease in maximal conductivity corresponding to the approximately 50% decrease in whole-plant conductance suggesting seasonal soil dry-down limitations within the downslope stands. Upslope stands had no to little change in root vulnerability or decrease in whole-plant conductance as soil water limitations occur immediately following snowmelt, thus limiting hydraulic conductance throughout the growing season. Integrating temporal and topographical variation for dynamic root vulnerability and whole-plant conductance estimates into ecosystem-scale models can decrease the uncertainty of evapotranspiration estimates in seasonally varying forests.
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NASA Astrophysics Data System (ADS)
Krofcheck, D. J.; Morillas, L.; Litvak, M. E.
2014-12-01
Drylands and semi-arid ecosystems cover over 45% of the global landmass. These biomes have been shown to be extremely sensitive to changes in climate, specifically decreases in precipitation and increases in air temperature. Therefore, inter-annual variability in climate has the potential to dramatically impact the carbon budget at regional and global scales. In the Southwestern US, we are in a unique position to investigate these relationships by leveraging eight years of data from the New Mexico Elevation Gradient (NMEG), eight flux towers that span six representative biomes across the semi-arid Southwest. From C4 desert grasslands to subalpine mixed conifer forests, the NMEG flux towers use identical instrumentsand processing, and afford a unique opportunity to explore patterns in biome-specific ecosystem processes and climate sensitivity. Over the last eight years the gradient has experienced climatic variability that span from wet years to an episodic megadrought. Here we report the effects of this extreme inter-annual variability in climate on the ability of semi-arid ecosystems to cycle and store energy and carbon. We also investigated biome-specific patterns of ecosystem light and water use efficiency during a series of wet and dry years, and how these vary in response to air temperature, vapor pressure deficit, evaporative fraction, and precipitation. Our initial results suggest that significant drought reduced the maximum ecosystem assimilation of carbon most at the C4 grasslands, creosote shrublands, juniper savannas, and ponderosa pine forests, with 60%, 50%, 35%, and 50% reduction respectively, relative to a wet year. Ecosystem light use efficiency tends to show the highest maximum values at the low elevation sites as a function of water availability, with the highest annual values consistently at the middle elevation and ponderosa pine sites. Water use efficiency was strongly biome dependent with the middle elevation sites showing the highest efficiencies, and the greatest within year variability at the lower elevation sites, with strong sensitivities to vapor pressure deficit. By quantifying the biome-specific ecosystem processes and functional responses, this network provides valuable insight about how vulnerable this range of semi-arid ecosystems is to future climate scenarios.
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Soil ecological interactions: comparisons between tropical and subalpine forests
Grizelle Gonzalez; Ruth E. Ley; Steven K. Schmidt; Xiaoming Zou; Timothy R. Seastedt
2001-01-01
Soil fauna can influence soil processes through interactions with the microbial community. Due to the complexity of the functional roles of fauna and their effects on microbes, little consensus has been reached on the extent to which soil fauna can regulate microbial activities. We quantified soil microbial biomass and maximum growth rates in control and fauna-excluded...
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Long-term effectiveness of restoration treatments on closed wilderness campsites
David N. Cole
2013-01-01
This study assessed long-term recovery of vegetation on six wilderness campsites in subalpine forests in Oregon that were closed to use and that received common restoration treatments (scarification, soil amendments, mulch, transplanting, and seeding). Vegetation cover was assessed every year for the first 7 years following treatment, as well as 10 and 15 years after...
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Whitebark pine ecosystem restoration in western Montana
Robert E. Keane; Stephen F. Arno
1996-01-01
Whitebark pine (Pinus albicaulis) is a major tree species of upper subalpine forests of the northern Rocky Mountains (Schmidt and McDonald 1990). It is an important nutritional and structural component of wildlife habitat (Arno and Hoff 1990; Schmidt and McDonald 1990). Its large, nutlike seeds are a major food source for many birds and mammals (...
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NASA Astrophysics Data System (ADS)
Caldwell, M. K.; Sloan, J.; Mladinich, C. S.; Wessman, C. A.
2013-12-01
Unmanned Aerial Systems (UAS) can provide detailed, fine spatial resolution imagery for ecological uses not otherwise obtainable through standard methods. The use of UAS imagery for ecology is a rapidly -evolving field, where the study of forest landscape ecology can be augmented using UAS imagery to scale and validate biophysical data from field measurements to spaceborne observations. High resolution imagery provided by UAS (30 cm2 pixels) offers detailed canopy cover and forest structure data in a time efficient and inexpensive manner. Using a GoPro Hero2 (2 mm focal length) camera mounted in the nose cone of a Raven unmanned system, we collected aerial and thermal data monthly during the summer 2013, over two subalpine forests in the Southern Rocky Mountains in Colorado. These forests are dominated by lodgepole pine (Pinus ponderosae) and have experienced insect-driven (primarily mountain pine beetle; MPB, Dendroctonus ponderosae) mortality. Objectives of this study include observations of forest health variables such as canopy water content (CWC) from thermal imagery and leaf area index (LAI), biomass and forest productivity from the Normalized Difference Vegetation Index (NDVI) from UAS imagery. Observations were, validated with ground measurements. Images were processed using a combination of AgiSoft Photoscan professional software and ENVI remote imaging software. We utilized the software Leaf Area Index Calculator (LAIC) developed by Córcoles et al. (2013) for calculating LAI from digital images and modified to conform to leaf area of needle-leaf trees as in Chen and Cihlar (1996) . LAIC uses a K-means cluster analysis to decipher the RGB levels for each pixel and distinguish between green aboveground vegetation and other materials, and project leaf area per unit of ground surface area (i.e. half total needle surface area per unit area). Preliminary LAIC UAS data shows summer average LAI was 3.8 in the most dense forest stands and 2.95 in less dense stands. These data correspond to 4.8 and 2.2 respectively from in situ Licor LAI 2200 measurements (Wilcoxon signed rank p value of 0.25, indicating there is no significant difference between LAIC and field measurements). Imagery over plots indicates about 12% canopy cover from standing dead vegetation within plots, which corresponds to about a 10% estimate of standing dead measured in the field. The next steps for analysis include calculating NDVI and CWC for plot-level vegetation, and scaling to the surrounding forested landscape. These high resolution estimates from UAS imagery will provide forest stand-to- landscape level biophysical data for forest health assessments, management, drought and disturbance monitoring and climate change modeling. Chen, J. M., and J. Cihlar. 1996. Retrieving LAI of boreal conifer forests using Landsat TM images. RSE 55:153-162. Córcoles, J., J. Ortega, D. Hernández, and M. Moreno. 2013. Use of digital photography from unmanned aerial vehicles for estimation of LAI in onion. EJoA. 45:96-104.
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NASA Astrophysics Data System (ADS)
Jennings, K. S.; Molotch, N. P.
2016-12-01
In cold, high-elevation sites, snowpack cold content acts as a buffer against climate warming by resisting snowmelt during periods of positive energy fluxes. To test the climate sensitivity of cold content and snowmelt, we employed the physical SNOWPACK snow model, forced with a 23-year, hourly, quality-controlled, gap-filled meteorological dataset from the Niwot Ridge Long Term Ecological Research (LTER) site in the Front Range mountains of Colorado. SNOWPACK was run at two points with seasonal snowpacks within the LTER, one in the alpine (3528 m) and one in the subalpine (3022 m). Model output was validated using snow water equivalent (SWE), snowpack temperature, and cold content data from snow pits dug near the met stations and automated SWE data from nearby SNOTEL snow pillows. Cold content accumulates primarily through additions of new snow, while negative energy fluxes—cooling through longwave emission and sublimation—play a lesser role, particularly in the deeper snowpack of the alpine. On average, the snowpack energy balance becomes positive on April 1 in the alpine and March 8 in the subalpine. Peak SWE occurs after these dates and its timing is primarily determined by the amount of precipitation received after peak cold content, with persistent snowfall delaying the main snowmelt pulse. Years with lower cold content, due to reduced precipitation and/or increased air temperature, experience an earlier positive energy balance with more melt events occurring before the date of peak SWE, which has implications for soil moisture, streamflow volume and timing, water uptake by vegetation, and microbial respiration. Synthetic warming experiments show significant cold content reductions and increased late-winter/early-spring melt as positive energy balances occur earlier in the snow season (a forward shift between 5.1 and 21.0 days per °C of warming). These results indicate cold, high-elevation sites, which are critical for water resources in the western United States, may lose their cold content buffering capacity and begin to experience stronger negative trends in SWE with increased climate warming, even as the majority of winter precipitation continues to fall as snow.
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Twentieth-century decline of large-diameter trees in Yosemite National Park, California, USA
Lutz, J.A.; van Wagtendonk, J.W.; Franklin, J.F.
2009-01-01
Studies of forest change in western North America often focus on increased densities of small-diameter trees rather than on changes in the large tree component. Large trees generally have lower rates of mortality than small trees and are more resilient to climate change, but these assumptions have rarely been examined in long-term studies. We combined data from 655 historical (1932-1936) and 210 modern (1988-1999) vegetation plots to examine changes in density of large-diameter trees in Yosemite National Park (3027 km2). We tested the assumption of stability for large-diameter trees, as both individual species and communities of large-diameter trees. Between the 1930s and 1990s, large-diameter tree density in Yosemite declined 24%. Although the decrease was apparent in all forest types, declines were greatest in subalpine and upper montane forests (57.0% of park area), and least in lower montane forests (15.3% of park area). Large-diameter tree densities of 11 species declined while only 3 species increased. Four general patterns emerged: (1) Pinus albicaulis, Quercus chrysolepis, and Quercus kelloggii had increases in density of large-diameter trees occur throughout their ranges; (2) Pinus jeffreyi, Pinus lambertiana, and Pinus ponderosa, had disproportionately larger decreases in large-diameter tree densities in lower-elevation portions of their ranges; (3) Abies concolor and Pinus contorta, had approximately uniform decreases in large-diameter trees throughout their elevational ranges; and (4) Abies magnifica, Calocedrus decurrens, Juniperus occidentalis, Pinus monticola, Pseudotsuga menziesii, and Tsuga mertensiana displayed little or no change in large-diameter tree densities. In Pinus ponderosa-Calocedrus decurrens forests, modern large-diameter tree densities were equivalent whether or not plots had burned since 1936. However, in unburned plots, the large-diameter trees were predominantly A. concolor, C. decurrens, and Q. chrysolepis, whereas P. ponderosa dominated the large-diameter component of burned plots. Densities of large-diameter P. ponderosa were 8.1 trees ha-1 in plots that had experienced fire, but only 0.5 trees ha-1 in plots that remained unburned. ?? 2009 Elsevier B.V. All rights reserved.
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Water and solute mass balance of five small, relatively undisturbed watersheds in the U.S.
Peters, N.E.; Shanley, J.B.; Aulenbach, Brent T.; Webb, R.M.; Campbell, D.H.; Hunt, R.; Larsen, M.C.; Stallard, R.F.; Troester, J.; Walker, J.F.
2006-01-01
Geochemical mass balances were computed for water years 1992-1997 (October 1991 through September 1997) for the five watersheds of the U.S. Geological Survey Water, Energy, and Biogeochemical Budgets (WEBB) Program to determine the primary regional controls on yields of the major dissolved inorganic solutes. The sites, which vary markedly with respect to climate, geology, physiography, and ecology, are: Allequash Creek, Wisconsin (low-relief, humid continental forest); Andrews Creek, Colorado (cold alpine, taiga/tundra, and subalpine boreal forest); Ri??o Icacos, Puerto Rico (lower montane, wet tropical forest); Panola Mountain, Georgia (humid subtropical piedmont forest); and Sleepers River, Vermont (humid northern hardwood forest). Streamwater output fluxes were determined by constructing empirical multivariate concentration models including discharge and seasonal components. Input fluxes were computed from weekly wet-only or bulk precipitation sampling. Despite uncertainties in input fluxes arising from poorly defined elevation gradients, lack of dry-deposition and occult-deposition measurements, and uncertain sea-salt contributions, the following was concluded: (1) for solutes derived primarily from rock weathering (Ca, Mg, Na, K, and H4SiO4), net fluxes (outputs in streamflow minus inputs in deposition) varied by two orders of magnitude, which is attributed to a large gradient in rock weathering rates controlled by climate and geologic parent material; (2) the net flux of atmospherically derived solutes (NH4, NO3, SO4, and Cl) was similar among sites, with SO4 being the most variable and NH4 and NO3 generally retained (except for NO 3 at Andrews); and (3) relations among monthly solute fluxes and differences among solute concentration model parameters yielded additional insights into comparative biogeochemical processes at the sites. ?? 2005 Elsevier B.V. All rights reserved.
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Krömer, Thorsten; Kreft, Holger; Gerold, Gerhard; Carvajal-Hernández, César Isidro; Heitkamp, Felix
2017-01-01
Terrestrial herbs are important elements of tropical forests; however, there is a lack of research on their diversity patterns and how they respond to different intensities of forest-use. The aim of this study was to analyze the diversity of herbaceous angiosperms along gradients of elevation (50 m to 3500 m) and forest-use intensity on the eastern slopes of the Cofre de Perote, Veracruz, Mexico. We recorded the occurrence of all herbaceous angiosperm species within 120 plots of 20 m x 20 m each. The plots were located at eight study locations separated by ~500 m in elevation and within three different habitats that differ in forest-use intensity: old-growth, degraded, and secondary forest. We analyzed species richness and floristic composition of herb communities among different elevations and habitats. Of the 264 plant species recorded, 31 are endemic to Mexico. Both α- and γ-diversity display a hump-shaped relation to elevation peaking at 2500 m and 3000 m, respectively. The relative contribution of between-habitat β-diversity to γ-diversity also showed a unimodal hump whereas within-habitat β-diversity declined with elevation. Forest-use intensity did not affect α-diversity, but β-diversity was high between old-growth and secondary forests. Overall, γ-diversity peaked at 2500 m (72 species), driven mainly by high within- and among-habitat β-diversity. We infer that this belt is highly sensitive to anthropogenic disturbance and forest-use intensification. At 3100 m, high γ-diversity (50 species) was driven by high α- and within-habitat β-diversity. There, losing a specific forest area might be compensated if similar assemblages occur in nearby areas. The high β-diversity and endemism suggest that mixes of different habitats are needed to sustain high γ-richness of terrestrial herbs along this elevational gradient. PMID:28792536
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Slatyer, Rachel A; Nash, Michael A; Miller, Adam D; Endo, Yoshinori; Umbers, Kate D L; Hoffmann, Ary A
2014-10-02
Mountain landscapes are topographically complex, creating discontinuous 'islands' of alpine and sub-alpine habitat with a dynamic history. Changing climatic conditions drive their expansion and contraction, leaving signatures on the genetic structure of their flora and fauna. Australia's high country covers a small, highly fragmented area. Although the area is thought to have experienced periods of relative continuity during Pleistocene glacial periods, small-scale studies suggest deep lineage divergence across low-elevation gaps. Using both DNA sequence data and microsatellite markers, we tested the hypothesis that genetic partitioning reflects observable geographic structuring across Australia's mainland high country, in the widespread alpine grasshopper Kosciuscola tristis (Sjösted). We found broadly congruent patterns of regional structure between the DNA sequence and microsatellite datasets, corresponding to strong divergence among isolated mountain regions. Small and isolated mountains in the south of the range were particularly distinct, with well-supported divergence corresponding to climate cycles during the late Pliocene and Pleistocene. We found mixed support, however, for divergence among other mountain regions. Interestingly, within areas of largely contiguous alpine and sub-alpine habitat around Mt Kosciuszko, microsatellite data suggested significant population structure, accompanied by a strong signature of isolation-by-distance. Consistent patterns of strong lineage divergence among different molecular datasets indicate genetic breaks between populations inhabiting geographically distinct mountain regions. Three primary phylogeographic groups were evident in the highly fragmented Victorian high country, while within-region structure detected with microsatellites may reflect more recent population isolation. Despite the small area of Australia's alpine and sub-alpine habitats, their low topographic relief and lack of extensive glaciation, divergence among populations was on the same scale as that detected in much more extensive Northern hemisphere mountain systems. The processes driving divergence in the Australian mountains might therefore differ from their Northern hemisphere counterparts.
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The Glacier Lakes Ecosystem Experiments Site
R. C. Musselman
1994-01-01
The Glacier Lakes Ecosystem Experiment Site (GLEES), a 600 ha research watershed at 3200-3400 m elevation in the Snowy Range of SE Wyoming, has been established to examine the effects of atmospheric deposition on alpine and subalpine ecosystems. This document provides preliminary data on the landscape habitats, floristics, geology, soils, aquatics, atmospheric...
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William C. Hunter
2010-01-01
The highest elevations in North America east of the Mississippi River are in the southern Blue Ridge of Virginia, North Carolina, and Tennessee. This area supports fauna and flora more characteristic of Canada than anywhere else in the southeast United States. The high-elevation forests are within the High Peaks Region to distinguish them from similar forests of the...
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Wang, Qing-Wei; Qi, Lin; Zhou, Wangming; Liu, Cheng-Gang; Yu, Dapao; Dai, Limin
2018-01-01
The growth limitation hypothesis (GLH) and carbon limitation hypothesis (CLH) are two dominant explanations for treeline formation. The GLH proposes that low temperature drives the treeline through constraining C sinks more than C sources, and it predicts that non-structural carbohydrate (NSC) levels are static or increase with elevation. Although the GLH has received strong support globally for evergreen treelines, there is still no consensus for deciduous treelines, which experience great asynchrony between supply and demand throughout the year. We investigated growth and the growing-season C dynamics in a common deciduous species, Erman's birch (Betula ermanii), along an elevational gradient from the closed forest to the treeline on Changbai Mountain, Northeast China. Samples were collected from developing organs (leaves and twigs) and main storage organs (stems and roots) for NSC analysis. Tree growth decreased with increasing elevation, and NSC concentrations differed significantly among elevations, organs, and sampling times. In particular, NSC levels varied slightly during the growing season in leaves, peaked in the middle of the growing season in twigs and stems, and increased continuously throughout the growing season in roots. NSCs also tended to increase or vary slightly in developing organs but decreased significantly in mature organs with increasing elevation. The decrease in NSCs with elevation in main storage organs indicates support for the CLH, while the increasing or static trends in new developing organs indicate support for the GLH. Our results suggest that the growth limitation theory may be less applicable to deciduous species' growth than to that of evergreen species. © 2018 Botanical Society of America.
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Fire-climate interactions in the Selway-Bitterroot Wilderness area
Kurt F. Kipfmueller; Thomas W. Swetnam
2000-01-01
Tree-ring reconstructed summer drought was examined in relation to the occurrence of 15 fires in the Selway-Bitterroot Wilderness Area (SBW). The ten largest fire years between 1880 and 1995 were selected from historical fire atlas data; five additional fire years were selected from a fire history completed in a subalpine forest within the SBW. Results of the analysis...
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K. F. Zeller; N. T. Nikolov
2000-01-01
Assessing the long-term exchange of trace gases and energy between terrestrial ecosystems and the atmosphere is an important priority of the current climate change research. In this regard, it is particularly significant to provide valid data on simultaneous fluxes of carbon, water vapor and pollutants over representative ecosystems. Eddy covariance measurements and...
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M.J. Trappe; K., Jr. Cromack; J.M. Trappe; J. Wilson; M.C. Rasmussen; M.A. Castellano; S.L. Miller
2009-01-01
Intensive recreational use of subalpine forests can create localized areas of concentrated disturbance where vegetation is altered, soils compacted, and surface fuels depleted. Many aspects of this disturbance type have been studied, but no research has focused on the effects of recreational use on mycorrhizal fungus sporocarp production. We measured the effects of...
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Effects of post-harvest treatments on high-elevation forests in the North Cascade Range, Washington.
Ella Elman; David L. Peterson
2005-01-01
We studied the effects of post-harvest treatments on regeneration and forest composition 13-27 years following harvest in high-elevation forests of the North Cascade Range, Washington. Eighteen sites encompassing three common post-harvest treatments were examined at elevations ranging from 830 m to 1460 m. Treatments included: (1) sites broadcast burned and planted...
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A molecular investigation of soil organic carbon composition across a subalpine catchment
Hsu, Hsiao-Tieh; Lawrence, Corey R.; Winnick, Matthew J.; Bargar, John R.; Maher, Katharine
2018-01-01
The dynamics of soil organic carbon (SOC) storage and turnover are a critical component of the global carbon cycle. Mechanistic models seeking to represent these complex dynamics require detailed SOC compositions, which are currently difficult to characterize quantitatively. Here, we address this challenge by using a novel approach that combines Fourier transform infrared spectroscopy (FT-IR) and bulk carbon X-ray absorption spectroscopy (XAS) to determine the abundance of SOC functional groups, using elemental analysis (EA) to constrain the total amount of SOC. We used this SOC functional group abundance (SOC-fga) method to compare variability in SOC compositions as a function of depth across a subalpine watershed (East River, Colorado, USA) and found a large degree of variability in SOC functional group abundances between sites at different elevations. Soils at a lower elevation are predominantly composed of polysaccharides, while soils at a higher elevation have more substantial portions of carbonyl, phenolic, or aromatic carbon. We discuss the potential drivers of differences in SOC composition between these sites, including vegetation inputs, internal processing and losses, and elevation-driven environmental factors. Although numerical models would facilitate the understanding and evaluation of the observed SOC distributions, quantitative and meaningful measurements of SOC molecular compositions are required to guide such models. Comparison among commonly used characterization techniques on shared reference materials is a critical next step for advancing our understanding of the complex processes controlling SOC compositions.
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Moore, Peggy E.; Alvarez, Otto; McKinney, Shawn T.; Li, Wenkai; Brooks, Matthew L.; Guo, Qinghua
2017-01-01
Whitebark pine and foxtail pine serve foundational roles in the subalpine zone of the Sierra Nevada. They provide the dominant structure in tree-line forests and regulate key ecosystem processes and community dynamics. Climate change models suggest that there will be changes in temperature regimes and in the timing and magnitude of precipitation within the current distribution of these species, and these changes may alter the species’ distributional limits. Other stressors include the non-native pathogen white pine blister rust and mountain pine beetle, which have played a role in the decline of whitebark pine throughout much of its range. The National Park Service is monitoring status and trends of these species. This report provides complementary information in the form of habitat suitability models to predict climate change impacts on the future distribution of these species within Sierra Nevada national parks.We used maximum entropy modeling to build habitat suitability models by relating species occurrence to environmental variables. Species occurrence was available from 328 locations for whitebark pine and 244 for foxtail pine across the species’ distributions within the parks. We constructed current climate surfaces for modeling by interpolating data from weather stations. Climate surfaces included mean, minimum, and maximum temperature and total precipitation for January, April, July, and October. We downscaled five general circulation models for the 2050s and the 2090s from ~125 km2 to 1 km2 under both an optimistic and an extreme climate scenario to bracket potential climatic change and its influence on projected suitable habitat. To describe anticipated changes in the distribution of suitable habitat, we compared, for each species, climate scenario, and time period, the current models with future models in terms of proportional change in habitat size, elevation distribution, model center points, and where habitat is predicted to expand or contract.Overall, models indicated that suitable habitats for whitebark and foxtail pine are more likely to shift geographically within the parks by 2100 rather than decline precipitously. This implies park managers might focus conservation efforts on stressors other than climate change, working toward species resilience in the face of threats from introduced disease and elevated native insect damage. More specifically, further understanding of the incidence and severity of white pine blister rust and other stressors in high elevation white pines would help assess vulnerability from threats other than climate change.
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NASA Astrophysics Data System (ADS)
Mihai, Bogdan; Savulescu, Ionut
2014-05-01
Forest fires in Romanian Carpathians became a frequent phenomenon during the last decade, although local climate and other environmental features did not create typical conditions. From 2004, forest fires affect in Romania more than 100 hectares/year of different forest types (deciduous and coniferous). Their magnitude and frequency are not known, since a historical forest fire inventory does not exist (only press papers and local witness for some selected events). Forest fires features the summer dry periods but there are dry autumns and early winter periods with events of different magnitudes. The application we propose is based on an empirical modeling of forest fire susceptibility in a typical mountain area from the Southern Carpathians, the Iezer Mountains (2462 m). The study area features almost all the altitudinal vegetation zones of the European temperate mountains, from the beech zone, to the coniferous zone, the subalpine and the alpine zones (Mihai et al., 2007). The analysis combines GIS and remote sensing models (Chuvieco et al., 2012), starting from the ideas that forest fires are featured by the ignition zones and then by the fire propagation zones. The first data layer (ignition zones) is the result of the crossing between the ignition factors: lightning - points of multitemporal occurence and anthropogenic activities (grazing, tourism and traffic) and the ignition zones (forest fuel zonation - forest stands, soil cover and topoclimatic factor zonation). This data is modelled from different sources: the MODIS imagery fire product (Hantson et al., 2012), detailed topographic maps, multitemporal orthophotos at 0.5 m resolution, Landsat multispectral imagery, forestry cadastre maps, detailed soil maps, meteorological data (the WorldClim digital database) as well as the field survey (mapping using GPS and local observation). The second data layer (fire propagation zones) is the result of the crossing between the forest fuel zonation, obtained with the help of forestry data, the wind regime data and the topographic features of the mountain area (elevation, slope declivity, slope aspect). The analysis also consider the insolation degree of mountain slopes, that creates favourable conditions for fire propagation between different canopies. These data layers are integrated within a simple GIS analysis in order to intersect the ignition zones with the fire propagation zones in order to obtain the potential areas to be affected by fire. The digital map show three levels of forest fire susceptibility, differenced on the basis of expert knowledge. The map can be validated from the statistical point of view with the polygons of the forest fire affected areas mapped from Landsat TM, ETM+ and OLI satellite imagery. The mapping results could be integrated within the forest management strategies and especially within the forest cadastre and development maps (updated every ten years). The result can confirm that the data gap in terms of forest fire events can be filled with expert knowledge. References Chuvieco, E, Aguado, I., Jurdao, S., Pettinari, M., Yebra, M., Salas, J., Hantson, S., de la Riva, J., Ibarra, P., Rodrigues, M., Echeverria, M., Azqueta, D., Roman, M., Bastarrika, A., Martinez, S., Recondo, C., Zapico, E., Martinez-Vega F.J. (2012) Integrating geospatial information into fire risk assessment, International Journal of Wildland Fire, 2,2, 69-86. Hantson, S., Padilla, M., Corti., D, Chuvieco, E. (2013) Strenghts and weaknesses of MODIS hotspots to characterize Global fire occurence, Remote Sensing of Environment, 131, 1, 152-159. Mihai, B., Savulescu, I.,Sandric, I. (2007) Change detection analysis (1986/2002) for the alpine, subalpine and forest landscape in Iezer Mountains (Southern Carpathians, Romania), Mountain Research and Development, 27, 250-258.
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Randall S. Morin; Richard H. Widmann
2010-01-01
Red spruce (Picea rubens Sarg.) is the most important component of the high-elevation forest ecosystems of the southern and central Appalachian Mountains. These communities are characterized by mixed hardwood/coniferous forests often with overstory dominance by red spruce. Due to their restricted geographic and elevation ranges, all community types...
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Phylogenetic structure of arbuscular mycorrhizal fungal communities along an elevation gradient.
Egan, Cameron P; Callaway, Ragan M; Hart, Miranda M; Pither, Jason; Klironomos, John
2017-04-01
Despite the importance of arbuscular mycorrhizal (AM) fungi within terrestrial ecosystems, we know little about how natural AM fungal communities are structured. To date, the majority of studies examining AM fungal community diversity have focused on single habitats with similar environmental conditions, with relatively few studies having assessed the diversity of AM fungi over large-scale environmental gradients. In this study, we characterized AM fungal communities in the soil along a high-elevation gradient in the North American Rocky Mountains. We focused on phylogenetic patterns of AM fungal communities to gain insight into how AM fungal communities are naturally assembled. We found that alpine AM fungal communities had lower phylogenetic diversity relative to lower elevation communities, as well as being more heterogeneous in composition than either treeline or subalpine communities. AM fungal communities were phylogenetically clustered at all elevations sampled, suggesting that environmental filtering, either selection by host plants or fungal niches, is the primary ecological process structuring communities along the gradient.
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Song, Xiaoyang; Li, Jieqiong; Zhang, Wenfu; Tang, Yong; Sun, Zhenhua; Cao, Min
2016-01-01
Seasonal drought is a common phenomenon in many forests predominated by monsoon climate. The impact of seasonal drought, however, may vary with elevations, and tree species of forests hence may differ in their response to elevations. In this study, we monitored the seasonal variation of seedling species composition, and their relative growth rate (RGR) along an elevational transect in tropical forests of southwest China for two years. We found tree seedling species richness declined with rising elevation. Seedling abundance and species richness increased significantly from dry season to rainy season. In dry season, RGR declined progressively from low to high elevational bands, while positive RGR occurred in each elevation in rainy season. We grouped seedling species into low and high elevation specialists based on their elevational distributions. Seasonal variance in soil moisture may lead to seasonal dynamics of seedling community in this area. Our results suggest that the observed change in local climate over the last 40 years tends to allow the tree species from high elevation to expand their distribution to the lower elevation, while the ranges of those at low elevations could be compressed or at the risk of extinction. PMID:27819274
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Johnny L. Boggs; Steven G. McNulty
2010-01-01
The objective of this study is to describe winter and summer surface air and forest floor temperature patterns and diurnal fluctuations in high-elevation red spruce (Picea rubens Sarg.) forests with different levels of canopy cover. In 1988, a series of 10- x 10-meter plots (control, low nitrogen [N] addition, and high nitrogen addition) were...
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W. J. Massman; J. M. Frank
2006-01-01
Meadow and forest CO2 amounts sampled beneath an approximately meter deep (steady state) snowpack at a subalpine site in southern Rocky Mountains of Wyoming are observed to vary by nearly 200 ppm over periods ranging from 4 to 15 days. This work employs the model of periodic, pressure-induced, advective transport in permeable media developed in...
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Paul C. Banko; Steven C. Hess; Paul G. Scowcroft; Chris Farmer; James D. Jacobi; Robert M. Stephens; Richard J. Camp; David L. Leonard; Kevin W. Brinck; J. O. Juvik; S. P. Juvik
2014-01-01
Under the multiple-use paradigm, conflicts may arise when protection of an endangered species must compete with other management objectives. To resolve such a conflict in the Critical Habitat of the endangered Hawaiian honeycreeper, palila (Loxioides bailleui), federal courts ordered the eradication of introduced ungulates responsible for damaging...
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Tang, Shi-shan; Yang, Wan-qin; Xiong, Li; Yin, Rui; Wang, Hai-peng; Zhang, Yan; Xu, Zhen-feng
2015-02-01
Fine root order was classified according to Pregitzer's method. This study measured carbon (C), nitrogen (N) and phosphorus (P) concentrations of the 1-5 root orders (diameter < 2 mm) in three dominant subalpine tree species (Betula albosinensis, Abies faxoniana and Picea asperata) of western Sichuan. Their stoichiometric ratios of different root orders were also calculated. The results showed that C concentration, C/N and C/P increased, but N and P concentrations decreased from the first to fifth order of fine root for all tree species. No significant changes in N/P among root orders were detected in each species. There were significant differences in C, N, P concentrations and their stoichiometric ratios among the tree species. The species-associated differences were dependent on root order. There were significant correlations between C, N, P concentrations and their stoichiometric ratios in the three tree species.
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NASA Astrophysics Data System (ADS)
Mackay, D. S.; Frank, J.; Reed, D.; Whitehouse, F.; Ewers, B. E.; Pendall, E.; Massman, W. J.; Sperry, J. S.
2012-04-01
In woody plant systems transpiration is often the dominant component of total evapotranspiration, and so it is key to understanding water and energy cycles. Moreover, transpiration is tightly coupled to carbon and nutrient fluxes, and so it is also vital to understanding spatial variability of biogeochemical fluxes. However, the spatial variability of transpiration and its links to biogeochemical fluxes, within- and among-ecosystems, has been a challenge to constrain because of complex feedbacks between physical and biological controls. Plant hydraulics provides an emerging theory with the rigor needed to develop testable hypotheses and build useful models for scaling these coupled fluxes from individual plants to regional scales. This theory predicts that vegetative controls over water, energy, carbon, and nutrient fluxes can be determined from the limitation of plant water transport through the soil-xylem-stomata pathway. Limits to plant water transport can be predicted from measurable plant structure and function (e.g., vulnerability to cavitation). We present a next-generation coupled transpiration-biogeochemistry model based on this emerging theory. The model, TREEScav, is capable of predicting transpiration, along with carbon and nutrient flows, constrained by plant structure and function. The model incorporates tightly coupled mechanisms of the demand and supply of water through the soil-xylem-stomata system, with the feedbacks to photosynthesis and utilizable carbohydrates. The model is evaluated by testing it against transpiration and carbon flux data along an elevation gradient of woody plants comprising sagebrush steppe, mid-elevation lodgepole pine forests, and subalpine spruce/fir forests in the Rocky Mountains. The model accurately predicts transpiration and carbon fluxes as measured from gas exchange, sap flux, and eddy covariance towers. The results of this work demonstrate that credible spatial predictions of transpiration and related biogeochemical fluxes will be possible at regional scales using relatively easily obtained vegetation structural and functional information.
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Laura Lusk; Matt Mutel; Elaine S. Walker; Foster. Levy
2010-01-01
The Black Mountain range of western North Carolina supports some of the most extensive but threatened high-elevation forests in the southern Appalachians. Of particular note, the insect pathogen, balsam woolly adelgid (Adelges piceae Ratzeburg), has been present on Mt. Mitchell for more than 50 years. In anticipation of potential changes in forest...
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Burns, S. P.; Blanken, P. D.; Turnipseed, A. A.; ...
2015-12-15
Precipitation changes the physical and biological characteristics of an ecosystem. Using a precipitation-based conditional sampling technique and a 14 year data set from a 25 m micrometeorological tower in a high-elevation subalpine forest, we examined how warm-season precipitation affected the above-canopy diel cycle of wind and turbulence, net radiation R net, ecosystem eddy covariance fluxes (sensible heat H, latent heat LE, and CO 2 net ecosystem exchange NEE) and vertical profiles of scalars (air temperature T a, specific humidity q, and CO 2 dry mole fraction χ c). This analysis allowed us to examine how precipitation modified these variables frommore » hourly (i.e., the diel cycle) to multi-day time-scales (i.e., typical of a weather-system frontal passage). During mid-day we found the following: (i) even though precipitation caused mean changes on the order of 50–70 % to R net, H, and LE, the surface energy balance (SEB) was relatively insensitive to precipitation with mid-day closure values ranging between 90 and 110 %, and (ii) compared to a typical dry day, a day following a rainy day was characterized by increased ecosystem uptake of CO 2 (NEE increased by ≈ 10 %), enhanced evaporative cooling (mid-day LE increased by ≈ 30 W m –2), and a smaller amount of sensible heat transfer (mid-day H decreased by ≈ 70 W m –2). Based on the mean diel cycle, the evaporative contribution to total evapotranspiration was, on average, around 6 % in dry conditions and between 15 and 25 % in partially wet conditions. Furthermore, increased LE lasted at least 18 h following a rain event. At night, even though precipitation (and accompanying clouds) reduced the magnitude of R net, LE increased from ≈ 10 to over 20 W m –2 due to increased evaporation. Any effect of precipitation on the nocturnal SEB closure and NEE was overshadowed by atmospheric phenomena such as horizontal advection and decoupling that create measurement difficulties. Above-canopy mean χ c during wet conditions was found to be about 2–3 μmol mol –1 larger than χ c on dry days. This difference was fairly constant over the full diel cycle suggesting that it was due to synoptic weather patterns (different air masses and/or effects of barometric pressure). Lastly, the effect of clouds on the timing and magnitude of daytime ecosystem fluxes is described.« less
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[Variations of soil labile organic carbon along an altitude gradient in Wuyi Mountain].
Xu, Xia; Chen, Yue-Qin; Wang, Jia-She; Fang, Yan-Hong; Quan, Wei; Ruan, Hong-Hua; Xu, Zi-Kun
2008-03-01
By using sequential fumigation-incubation method, this paper determined the soil labile organic carbon (LOC) content under evergreen broadleaf forest, coniferous forest, sub-alpine dwarf forest, and alpine meadow along an altitude gradient in Wuyi Mountain National Nature Reserve in Fujian Province of China, with its relations to soil microbial biomass carbon (MBC), total organic carbon (TOC), total nitrogen (TN), and fine root biomass (FRB) analyzed. The results showed that soil LOC occupied 3.40%-7.46% of soil TOC, and soil MBC occupied 26.87%-80.38% of the LOC. The LOC under different forest stands increased significantly with altitude, and decreased with soil depth. Soil LOC had very significant correlations with soil MBC, TOC, TN and FRB, and its content was obviously higher at higher altitudes than at lower altitudes.
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Growth and demography of Pinaleno high elevation forests
Christopher O' Connor; Donald A. Falk; Ann M. Lynch; Craig P. Wilcox; Thomas W. Swetnam; Tyson L. Swetnam
2010-01-01
The project goal is to understand how multiple disturbance events including fire, insect outbreaks, and climate variability interact in space and time, and how they combine to influence forest species composition, spatial structure, and tree population dynamics in high elevation forests of the Pinaleno Mountains. Information from each of these components is needed in...
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NASA Astrophysics Data System (ADS)
Kovda, Irina; Morgun, Evgeny; Golubeva, Natalia
2010-05-01
Stable isotopic composition of carbon in plant species and soil organic matter was investigated along the bioclimatic transect in the North Caucasus. The aim of this research was to find the possible shift of stable isotopic composition of carbon reflecting the gradual successive changes of landscapes connected with the changes of climatic conditions (temperature, precipitation, air humidity) at a various absolute heights above sea level i.e. along the vertical zonation. The study site was located in the North Caucasus near Arkhiz, Big Zelentchuk valley (43o33-40'N; 41o16-27'E). Soil and vegetation samples were collected along Precaucasus and Caucasus slopes at an absolute heights interval of 1280 - 2065 m. Soils are formed at eluvo-deluvium of noncarbonated silicate massive rocks under warm temperate climate with MAT 4-5oC and MAP ~ 860 mm and more. Samples of vegetation (whole grasses and tree leaves) represented several altitudinal vegetation belts including forest and subalpine belts with coniferous (pine, spruce, fir), mixed coniferous-deciduous (fir + beech), broad-leaved (beech, maple), small-leaved (birch, aspen) forests, elfin birch wood and subalpine meadows. Stable isotope composition of carbon was determined using IRMS Finnigan Delta V+. Stable isotopic composition of vegetal species ranges from -33.04 to -27.29 o/oo with the general trend of lighter δ13С with the decreasing of absolute heights. The most heavier δ13С = ~ -27o/oo were found in subalpine meadow plants, while at a smaller altitudes in the forest belt δ13С shifts to ~ 30-31, and up to ~ - 32-33o/oo. More clear regularities were found for vegetation specimens grouped into three categories such as "trees", "grasses" and "litter". δ13С of each category clearly shifts to the lighter values with the decrease of absolute heights i.e from subalpine meadows to spruce-broad-leaved forests. δ13С shift is about 2,49o/oo for trees, 1,75-4,92 o/oo for grasses and ~ 1,8 o/oo for the litter. The complimentary determination of δ15N have shown large variability from -3,59 to + 18,24o/oo which corresponds with the expected large variation of δ15N of plant debris and general trend of lighter δ15N during the decomposition. δ13С of soil organic matter varies from -28.92 to -24.99 o/oo. The similar trend for lighter stable isotopic composition of carbon at a lower absolute heights i.e. in forest belt was found for soil organic matter. The whole picture of δ13С values in soil samples is quite complicated and needs further investigation. As a result, the isotopic characteristic of carbon and nitrogen in vegetal specimens, and carbon in soil organic matter were obtained along a bioclimatic topocatena. The general trend of lighter C isotopic composition was found. It is known that lighter isotopic composition correlates with cooler and/or wetter climate. While the temperature gradually decreases upward, the changes of precipitation are not linear. Further research is needed to better explain the variation of stable isotopic compositions in dependence of those climatic factors. This research was supported by the Presidium of Russian Academy of Sciences, Program 14
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Small bugs with big impacts: Ecosystem and watershed-level responses to the MPB epidemic [Chapter 7
Rob Hubbard; Kelly Elder; Chuck Rhoades; Polly Hays; Bruce Sims
2014-01-01
Mountain pine beetle (MPB) outbreaks have the potential for prolonged impacts on the delivery of clean water from infested subalpine watersheds throughout the West. Sixty-five percent of the Westâs water supply originates on forested land (Brown and others 2008), much of which has been affected by an unprecedented MPB epidemic over the past decade. Some lodgepole pine...
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Monique LaPerriere Nelson; Charles C. Rhoades; Kathleen A. Dwire
2011-01-01
We characterized the water chemistry of nine slope wetlands and adjacent headwater streams in Colorado subalpine forests and compared sites in basins formed on crystalline bedrock with those formed in basins with a mixture of crystalline and sedimentary bedrock. The pH, Ca2+, Mg2+, NH4 +, acid neutralizing capacity, and electrical conductivity of wetland porewater and...
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Anita C. Risch; Martin F. Jurgensen; Deborah S. Page-Dumroese; Martin Schutz
2013-01-01
Forest cover has increased in mountainous areas of Europe over the past decades because of the abandonment of agricultural areas (land-use change). For this reason, understanding how land-use change affects carbon (C) source-sink strength is of great importance. However, most studies have assessed mountainous systems C stocks, and less is known about C turnover rates,...
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Ozone and carbon dioxide fluxes in a subalpine spruce-fir forest ecosystem
Karl Zeller
1995-01-01
RMFRES RWU 4452 has made several ozone (O3) and carbon dioxide (CO2) trace gas flux measurements in the Snowy Range, WY GLEES research area over the past few years. These measurements were made using the micrometeorological eddy correlation technique at two sites: one 6 m above tree canopy height on the Brooklyn tower (ozone only); and the other below canopy height, 1-...
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James M. Lenihan; Dominique Bachelet; Raymond Drapek; Ronald P. Neilson
2006-01-01
The objective of this study was to dynamically simulate the response of vegetation distribution, carbon, and fire to three scenarios of future climate change for California using the MAPSS-CENTURY (MCI) dynamic general vegetation model. Under all three scenarios, Alpine/Subalpine Forest cover declined with increased growing season length and warmth, and increases in...
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Charles F. Leaf
1975-01-01
Summarizes a series of comprehensive reports on watershed management in five major vegetation zones: (1) the coniferous forest subalpine zone; (2) the Front Range ponderosa pine zone; (3) the Black Hills ponderosa pine zone; (4) the alpine zone; and (5) the big sagebrush zone. Includes what is known about the hydrology of these lands, what hydrologic principles are...
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Hu, Jia; Moore, David J P; Riveros-Iregui, Diego A; Burns, Sean P; Monson, Russell K
2010-03-01
*Understanding controls over plant-atmosphere CO(2) exchange is important for quantifying carbon budgets across a range of spatial and temporal scales. In this study, we used a simple approach to estimate whole-tree CO(2) assimilation rate (A(Tree)) in a subalpine forest ecosystem. *We analysed the carbon isotope ratio (delta(13)C) of extracted needle sugars and combined it with the daytime leaf-to-air vapor pressure deficit to estimate tree water-use efficiency (WUE). The estimated WUE was then combined with observations of tree transpiration rate (E) using sap flow techniques to estimate A(Tree). Estimates of A(Tree) for the three dominant tree species in the forest were combined with species distribution and tree size to estimate and gross primary productivity (GPP) using an ecosystem process model. *A sensitivity analysis showed that estimates of A(Tree) were more sensitive to dynamics in E than delta(13)C. At the ecosystem scale, the abundance of lodgepole pine trees influenced seasonal dynamics in GPP considerably more than Engelmann spruce and subalpine fir because of its greater sensitivity of E to seasonal climate variation. *The results provide the framework for a nondestructive method for estimating whole-tree carbon assimilation rate and ecosystem GPP over daily-to weekly time scales.
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Robert E. Keane; Anna W. Schoettle
2011-01-01
Many ecologically important, five-needle white pine forests that historically dominated the high elevation landscapes of western North America are now being heavily impacted by mountain pine beetle (Dendroctonus spp.) outbreaks, the exotic disease white pine blister rust (WPBR), and altered high elevation fire regimes. Management intervention using specially designed...
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O'Donnell, Frances C; Flatley, William T; Springer, Abraham E; Fulé, Peter Z
2018-06-25
Climate change and wildfire are interacting to drive vegetation change and potentially reduce water quantity and quality in the southwestern United States, Forest restoration is a management approach that could mitigate some of these negative outcomes. However, little information exists on how restoration combined with climate change might influence hydrology across large forest landscapes that incorporate multiple vegetation types and complex fire regimes. We combined spatially explicit vegetation and fire modeling with statistical water and sediment yield models for a large forested landscape (335,000 ha) on the Kaibab Plateau in northern Arizona, USA. Our objective was to assess the impacts of climate change and forest restoration on the future fire regime, forest vegetation, and watershed outputs. Our model results predict that the combination of climate change and high-severity fire will drive forest turnover, biomass declines, and compositional change in future forests. Restoration treatments may reduce the area burned in high-severity fires and reduce conversions from forested to non-forested conditions. Even though mid-elevation forests are the targets of restoration, the treatments are expected to delay the decline of high-elevation spruce-fir, aspen, and mixed conifer forests by reducing the occurrence of high-severity fires that may spread across ecoregions. We estimate that climate-induced vegetation changes will result in annual runoff declines of up to 10%, while restoration reduced or reversed this decline. The hydrologic model suggests that mid-elevation forests, which are the targets of restoration treatments, provide around 80% of runoff in this system and the conservation of mid- to high-elevation forests types provides the greatest benefit in terms of water conservation. We also predict that restoration treatments will conserve water quality by reducing patches of high-severity fire that are associated with high sediment yield. Restoration treatments are a management strategy that may reduce undesirable outcomes for multiple ecosystem services. © 2018 by the Ecological Society of America.
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Vegetation mapping of Nowitna National Wildlife Reguge, Alaska using Landsat MSS digital data
Talbot, S. S.; Markon, Carl J.
1986-01-01
A Landsat-derived vegetation map was prepared for Nowitna National Wildlife Refuge. The refuge lies within the middle boreal subzone of north central Alaska. Seven major vegetation classes and sixteen subclasses were recognized: forest (closed needleleaf, open needleleaf, needleleaf woodland, mixed, and broadleaf); broadleaf scrub (lowland, alluvial, subalpine); dwarf scrub (prostrate dwarf shrub tundra, dwarf shrub-graminoid tussock peatland); herbaceous (graminoid bog, marsh and meadow); scarcely vegetated areas (scarcely vegetated scree and floodplain); water (clear, turbid); and other areas (mountain shadow). The methodology employed a cluster-block technique. Sample areas were described based on a combination of helicopter-ground survey, aerial photointerpretation, and digital Landsat data. Major steps in the Landsat analysis involved preprocessing (geometric correction), derivation of statistical parameters for spectral classes, spectral class labeling of sample areas, preliminary classification of the entire study area using a maximum-likelihood algorithm, and final classification utilizing ancillary information such as digital elevation data. The final product is a 1:250,000-scale vegetation map representative of distinctive regional patterns and suitable for use in comprehensive conservation planning.
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Nydick, K.R.; LaFrancois, B.M.; Baron, Jill S.; Johnson, B.M.
2003-01-01
We explored variability among subalpine lakes sharing very similar climate and atmospheric conditions, but differing in watershed characteristics, hydrology, and food web structure. Special attention was given to nitrogen (N) dynamics because the study area receives some of the highest levels of atmospheric N deposition in the Rocky Mountains. We asked if the effect of regional N deposition would be manifested uniformly among neighboring lakes both in terms of ambient conditions and responses to greater nutrient inputs. Catchment vegetation appeared to be the main determinant of ambient nitrate (NO3), phosphate (PO4), and dissolved organic carbon (DOC) concentrations, although in-lake differences in recycling produced variable and contrasting NH4 levels. Phytoplankton chlorophyll a temporarily responded to early season NO3 peaks in the lakes with rocky watersheds, but chlorophyll means over the ice-free season were remarkably similar among lakes despite differences in both nutrient supply and zooplankton grazing. In most cases, phosphorus was limiting to phytoplankton growth, although the importance of N deficiencies was greater in lakes with forested watersheds and fringing wetlands.
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Nydick, K.R.; LaFrancois, B.M.; Baron, Jill S.; Johnson, B.M.
2003-01-01
We explored variability among subalpine lakes sharing very similar climate and atmospheric conditions, but differing in watershed characteristics, hydrology, and food web structure. Special attention was given to nitrogen (N) dynamics because the study area receives some of the highest levels of atmospheric N deposition in the Rocky Mountains. We asked if the effect of regional N deposition would be manifested uniformly among neighboring lakes both in terms of ambient conditions and responses to greater nutrient inputs. Catchment vegetation appeared to be the main determinant of ambient nitrate (NO3), phosphate (PO4), and dissolved organic carbon (DOC) concentrations, although in-lake differences in recycling produced variable and contrasting NH4 levels. Phytoplankton chlorophyll atemporarily responded to early season NO3 peaks in the lakes with rocky watersheds, but chlorophyll means over the ice-free season were remarkably similar among lakes despite differences in both nutrient supply and zooplankton grazing. In most cases, phosphorus was limiting to phytoplankton growth, although the importance of N deficiencies was greater in lakes with forested watersheds and fringing wetlands.
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Conner, Lafe G; Gill, Richard A.; Belnap, Jayne
2016-01-01
Soil moisture in seasonally snow-covered environments fluctuates seasonally between wet and dry states. Climate warming is advancing the onset of spring snowmelt and may lengthen the summer-dry state and ultimately cause drier soil conditions. The magnitude of either response may vary across elevation and vegetation types. We situated our study at the lower boundary of persistent snow cover and the upper boundary of subalpine forest with paired treatment blocks in aspen forest and open meadow. In treatments plots, we advanced snowmelt timing by an average of 14 days by adding dust to the snow surface during spring melt. We specifically wanted to know whether early snowmelt would increase the duration of the summer-dry period and cause soils to be drier in the early-snowmelt treatments compared with control plots. We found no difference in the onset of the summer-dry state and no significant differences in soil moisture between treatments. To better understand the reasons soil moisture did not respond to early snowmelt as expected, we examined the mediating influences of soil organic matter, texture, temperature, and the presence or absence of forest. In our study, late-spring precipitation may have moderated the effects of early snowmelt on soil moisture. We conclude that landscape characteristics, including soil, vegetation, and regional weather patterns, may supersede the effects of snowmelt timing in determining growing season soil moisture, and efforts to anticipate the impacts of climate change on seasonally snow-covered ecosystems should take into account these mediating factors.
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NASA Astrophysics Data System (ADS)
Herring, Erin M.; Gavin, Daniel G.
2015-06-01
There are very few terrestrial sediment records from North America that contain a nearly continuous sequence spanning from the Last Interglacial period to the present. We present stratigraphic records of pollen and several other proxies from a Carex-dominated wetland, Star Meadows, located 140 km south of the maximum extent of the Cordilleran Ice Sheet and near the current southern extent of interior mesic forests in northern Idaho. Many species in this region are disjunct by 160 km of arid steppe and dry forest from their more extensive distribution along the Pacific Northwest coast and may have survived in an interior refugium. The chronology for the upper 251 cm was determined by six radiocarbon dates and one tephra deposit, and the age of the remainder of the core (251-809 cm) was estimated by correlation with SPECMAP δ18O. Fluctuating water levels were inferred from alternating peat, biogenic silica, and aquatic pollen types. During MIS 5e the region was warmer and drier than today and was dominated by Pinus (likely Pinus contorta) mixed conifer forest surrounding a Carex meadow. A cool-moist climate (MIS 5b-5d) soon developed, and the site was inundated with deep water. Pollen indicated wetland vegetation (Betula glandulosa, Typhaceae, and Salix) developed around a lake with a Pseudotsuga/Larix and Picea forest on the surrounding slopes. During MIS 5a, a warmer climate supported a Pseudotsuga/Larix, Abies, and Picea forest on the surrounding hillsides and a Carex-dominated environment within a dry meadow. From MIS 4 to MIS 3, a cool and wet Pinus and Picea forest predominated. Water levels rose, enabling Nuphar to persist within a perennial lake while a sedge fen established along the lake margin. As climate transitioned into MIS 2, a cooler and drier climate supported a Pinus and Picea subalpine parkland, though water levels remained high enough to support Nuphar. During the Last Glacial Maximum the sediment was mainly silt and clay with high Artemisia and very poor pollen preservation. Glaciers descended to 500 m elevation above Star Meadows in adjacent drainages suggesting a periglacial environment occurred at the site. Lake level decreased through the Pleistocene-Holocene transition (ca. 11.7 ka) and the site returned to a sedge peatland surrounded by an open Pinus forest. The most striking vegetation change occurred in the middle to late Holocene with the first occurrence and then later dominance of Cupressaceae pollen, most likely Thuja plicata, which is a dominant species in modern interior mesic forests. The late Holocene vegetation was uniquely mesic in the context of the last 120,000 years, casting doubt on this region serving as a glacial refugium.
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Mass tree mortality leads to mangrove peat collapse at Bay Islands, Honduras after Hurricane Mitch
Cahoon, D.R.; Hensel, P.; Rybczyk, J.; McKee, K.L.; Proffitt, C.E.; Perez, B.C.
2003-01-01
We measured sediment elevation and accretion dynamics in mangrove forests on the islands of Guanaja and Roatan, Honduras, impacted by Hurricane Mitch in 1998 to determine if collapse of underlying peat was occurring as a result of mass tree mortality. Little is known about the balance between production and decomposition of soil organic matter in the maintenance of sediment elevation of mangrove forests with biogenic soils. Sediment elevation change measured with the rod surface elevation table from 18 months to 33 months after the storm differed significantly among low, medium and high wind impact sites. Mangrove forests suffering minimal to partial mortality gained elevation at a rate (5 mm yeara??1) greater than vertical accretion (2 mm yeara??1) measured from artificial soil marker horizons, suggesting that root production contributed to sediment elevation. Basin forests that suffered mass tree mortality experienced peat collapse of about 11 mm yeara??1 as a result of decomposition of dead root material and sediment compaction. Low soil shear strength and lack of root growth accompanied elevation decreases. Model simulations using the Relative Elevation Model indicate that peat collapse in the high impact basin mangrove forest would be 37 mm yeara??1 for the 2 years immediately after the storm, as root material decomposed. In the absence of renewed root growth, the model predicts that peat collapse will continue for at least 8 more years at a rate (7 mm yeara??1) similar to that measured (11 mm yeara??1). Mass tree mortality caused rapid elevation loss. Few trees survived and recovery of the high impact forest will thus depend primarily on seedling recruitment. Because seedling establishment is controlled in large part by sediment elevation in relation to tide height, continued peat collapse could further impair recovery rates.
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Valuing the forest for the trees: Willingness to pay for white pine blister rust management
James R. Meldrum; Patricia A. Champ; Craig A. Bond
2011-01-01
The nearly two million acres of high elevation forests in the Western United States are not an important source of timber or any other market products. However, that does not mean that the forests are not highly valuable. Visitors and nonvisitors alike value the unique five-needle pine trees found in these high elevation ecosystems. In this study, we estimate the...
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Campbell, John L.; Shinneman, Douglas
2017-01-01
IntroductionClimate change is expected to impose significant tension on the geographic distribution of tree species. Yet, tree species range shifts may be delayed by their long life spans, capacity to withstand long periods of physiological stress, and dispersal limitations. Wildfire could theoretically break this biological inertia by killing forest canopies and facilitating species redistribution under changing climate. We investigated the capacity of wildfire to modulate climate-induced tree redistribution across a montane landscape in the central Rocky Mountains under three climate scenarios (contemporary and two warmer future climates) and three wildfire scenarios (representing historical, suppressed, and future fire regimes).MethodsDistributions of four common tree species were projected over 90 years by pairing a climate niche model with a forest landscape simulation model that simulates species dispersal, establishment, and mortality under alternative disturbance regimes and climate scenarios.ResultsThree species (Douglas-fir, lodgepole pine, subalpine fir) declined in abundance over time, due to climate-driven contraction in area suitable for establishment, while one species (ponderosa pine) was unable to exploit climate-driven expansion of area suitable for establishment. Increased fire frequency accelerated declines in area occupied by Douglas-fir, lodgepole pine, and subalpine fir, and it maintained local abundance but not range expansion of ponderosa pine.ConclusionsWildfire may play a larger role in eliminating these conifer species along trailing edges of their distributions than facilitating establishment along leading edges, in part due to dispersal limitations and interspecific competition, and future populations may increasingly depend on persistence in locations unfavorable for their establishment.
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NASA Astrophysics Data System (ADS)
Fairbanks, D.; Green, K.; Murphy, M. A.; Shepard, C.; Chorover, J.; Rich, V. I.; Gallery, R. E.
2015-12-01
Wildfires are increasing in size and severity across the western United States with impacts on regional biogeochemical cycling. The resiliency of resident soil microbial communities determines rates of nutrient transformations as well as forest structure and recovery. Redox conditions in soil determine metabolic activities of microorganisms, which first consume oxygen and a succession of alternative terminal electron acceptors to support growth and metabolism using a variety of carbon sources. Controls on redox zonation are largely unknown in dominantly oxic soils, and microbial community adaptation and response to fluctuations in redox potential in a sub-alpine forested post-disturbance catchment has not been studied. Previous work has shown that fluctuating or rising water tables result in redox-dynamic sites, which can be 'hot spots' of biogeochemical activity depending on landscape position. Fire-induced tree mortality results in altered hydrologic flow paths and decreased evapotranspiration, leading to potential for intensified hot spot activity. We are testing such coupling of microbial activity with fluctuations in redox status using field measurements and laboratory incubation experiments. The 2013 Thompson Ridge Fire in the Jemez River Basin (NM) Critical Zone Observatory provides a highly-contextualized opportunity to examine how disturbance regime affects changes in soil microbial community dynamics and fluctuations in reduction-oxidation potential (as quantified by continuous CZO measurements of O2, CO2 and Eh as a function of soil depth and landscape location). We hypothesize that areas of depositional convergence in the catchment, which have been shown to exhibit more reducing conditions, will host microbial communities that are better adapted to fluctuating redox conditions and exhibit a greater diversity in functional capabilities. In these mixed conifer forests we find shifts in redox potential status in relation to depth and topography where more reducing conditions typically occur in convergent zones and at depth. These results highlight the significance of fluctuating oxygen-depleted zones in aerobic soils on microbial community activity and structure, linking community response to larger scale ecosystem processes.
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K. S. Burns; J. W. Hanna; Ned Klopfenstein; M.-S. Kim
2016-01-01
In July 2014, mycelial fans (isolates CO104F, CO106F, and CO108F) of Armillaria sp. were collected from forest trees in Colorado. These isolates were all identified as A. sinapina based on a somatic pairing test against 18 tester isolates representing six North American Armillaria spp. and nucleotide sequences of the translation elongation factor 1alpha (tef-...
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NASA Astrophysics Data System (ADS)
Gonzalez, P.; Eigenbrod, F.; Early, R.; Wang, F.; Notaro, M.; Williams, J. W.
2016-12-01
U.S. national parks conserve globally unique biodiversity. Yet, historical impacts of climate change and future vulnerabilities threaten species and ecosystems across this system of protected areas. Spatial analyses of historical climate and downscaled future climate projections show climate trends across the system. Spatial analyses of vegetation and wildfire (using a dynamic global vegetation model), habitat fragmentation (using remote sensing-derived land cover), and invasive species introduction and establishment show patterns of future vulnerability across the 50 U.S. states and 412 U.S. national parks. Results reveal high historical and projected temperature increases and precipitation changes, projected increases of wildfire across western U.S. national parks, high vulnerability to biome shifts and habitat fragmentation of up to one-third of National Park System area, and high vulnerability to invasive species of one-ninth of National Park System area. Ecosystems in the Sierra Nevada, Cascade Range, desert Southwest, and Laurentian Great Lakes are highly vulnerable to upslope and poleward shifts of the North America sequence of biomes: temperate shrubland - temperate broadleaf forest - temperate mixed forest - temperate conifer forest - subalpine and boreal forest - alpine and tundra. These areas include Grand Canyon, Mount Rainier, and Yosemite National Parks. The southwestern U.S., including Grand Canyon and Sequoia National Parks, is vulnerable to increases in wildfire. The eastern and midwestern U.S., including Great Smokey Mountains and Voyageurs National Parks, are highly vulnerable to invasive species. These results identify vulnerable areas and potential refugia to help prioritize areas for future natural resource management actions and biodiversity conservation in U.S. national parks.
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[Distribution characteristics of heavy metals along an elevation gradient of montane forest].
Wan, Jia-rong; Nie, Ming; Zou, Qin; Hu, Shao-chang; Chen, Jia-kuan
2011-12-01
In the present paper, the concentrations of fourteen heavy metals (Fe, Al, Ti, Cu, Cr, Mn, V, Zn, Ni, Co, Pb, Se, Cd and As) were determined by ICP-AES and atomic absorption spectroscopy along an elevation gradient of montane forest. The results show that the elevation gradient had significant effects on the concentrations of Fe, Al, Ti, V, Pb and As. And the concentrations of Cu, Cr, Mn, Zn, Ni, Co, Se and Cd were not significantly affected by the elevation gradient. Because the studying area is red soil, the elevation gradient had significant effects on the concentrations of Fe, Al and Ti which are characteristic heavy metals of red soil, suggesting that the red soil at different elevations has different intensities of weathering desilication and bioaccumulation. Other heavy metals have different relationships with the elevation gradient, such as the concentrations of Cr, Zn and Cd were high at relatively high elevation and Pb and As were high at relatively low elevation. These results suggest that the different elevations of montane forest soils were polluted by differently types of heavy metals.
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Increasing elevation of fire in the Sierra Nevada and implications for forest change
Mark W. Schwartz; Nathalie Butt; Christopher R. Dolanc; Andrew Holguin; Max A. Moritz; Malcolm P. North; Hugh D. Safford; Nathan L. Stephenson; James H. Thorne; Phillip J. van Mantgem
2015-01-01
Fire in high-elevation forest ecosystems can have severe impacts on forest structure, function and biodiversity. Using a 105-year data set, we found increasing elevation extent of fires in the Sierra Nevada, and pose five hypotheses to explain this pattern. Beyond the recognized pattern of increasing fire frequency in the Sierra Nevada since the late 20th century, we...
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NASA Astrophysics Data System (ADS)
Beverly, D.; Ewers, B. E.; Hyde, K.; Ohara, N.; Speckman, H. N.
2015-12-01
High elevation watersheds of the Rocky Mountains region contribute over 70% of the streamflow needed for infrastructure, agriculture, and ecological processes. Snow-water yields are heterogeneous in space and time and are driven by a multitude of snow distribution processes, including snowpack evolution driven by physical and biological factors. Quantifying heterogeneity of snowpack is further complicated by vegetation perturbations; much of the Rocky Mountains have experienced significant tree mortality due to bark beetle outbreaks. Reduction of living crown area decreases canopy interception while increasing radiation to snow surfaces, which alters snowpack distribution throughout the catchment. We hypothesize that, in a complex watershed, topographic variation (i.e., slope and aspect) will have a greater effect on snowpack evolution and distribution than densities of canopy mortality due to beetle infestation. The 120 ha No Name watershed, located in southern Wyoming at 3000 m elevation was divided into twenty-one 175 m2 parcels, in which plots were randomly assigned within each parcel. Peak snow was measured in April; in the 50 m2 plots, depths were measured every 2 m along north-south and east-west transects. Twenty-one snow pits were excavated to quantify snow densities in 10 cm increments throughout the pit profile. Forest inventories occurred the following summer. Peak snowpack levels occurred in April with mean depth of 92.3 ± 2.4 cm and peak SWE of 34.0 ± 0.84 cm. Binary decision trees accounted for 63% of the variability after including topographic indices, beetle condition of the trees, LAI, and basal area. Snow depth showed a slight positive relationship with increased in beetle mortality on slopes less than 11 degrees. Overall, topographic indices are greater drivers for snow distributions compared to effects of tree mortality.
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Incidental captures of Eastern Spotted Skunk in a high-elevation Red Spruce forest in Virginia
Diggins, Corinne A.; Jachowski, David S.; Martin, Jay; Ford, W. Mark
2015-01-01
Spilogale putorius (Eastern Spotted Skunk) is considered rare in the southern Appalachian Mountains and throughout much of its range. We report incidental captures of 6 Eastern Spotted Skunks in a high-elevation Picea rubens (Red Spruce) forest in southwestern Virginia during late February and March 2014. At 1520 m, these observations are the highest-elevation records for Eastern Spotted Skunk in the Appalachian Mountains. They are also the first known records of this species using Red Spruce forests in the southern Appalachians.
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The ecology of mixed severity fire regimes in Washington, Oregon, and Northern California
David Perry; Paul Hessburg; Carl Skinner; Thomas Spies; Scott Stephens; Alan Henry Taylor; Jerry Franklin; Brenda McComb; Greg Riegel
2011-01-01
Forests characterized by mixed-severity fires occupy a broad moisture gradient between lower elevation forests typified by low-severity fires and higher elevation forests in which high-severity, stand replacing fires are the norm. Mixed-severity forest types are poorly documented and little understood but likely occupy significant areas in the western United States. By...
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Soil carbon in arid and semiarid forest ecosystems [Chapter 18
Daniel G. Neary; Steven T. Overby; Stephen C. Hart
2002-01-01
Forests of the semiarid and arid zones of the interior western United States (US) are some of the most unique in North America. They occupy 11 to 34% of the landscape at mostly higher elevations (USDA Forest Service, 1981). These forests are characterized by a high diversity of flora, fauna, climates, elevations, soils, geology, hydrology, and productivity. Within the...
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Koo, Kyung Ah; Kong, Woo-Seok; Nibbelink, Nathan P; Hopkinson, Charles S; Lee, Joon Ho
2015-01-01
Climate change has caused shifts in species' ranges and extinctions of high-latitude and altitude species. Most cold-tolerant evergreen broadleaved woody plants (shortened to cold-evergreens below) are rare species occurring in a few sites in the alpine and subalpine zones in the Korean Peninsula. The aim of this research is to 1) identify climate factors controlling the range of cold-evergreens in the Korean Peninsula; and 2) predict the climate change effects on the range of cold-evergreens. We used multimodel inference based on combinations of climate variables to develop distribution models of cold-evergreens at a physiognomic-level. Presence/absence data of 12 species at 204 sites and 6 climatic factors, selected from among 23 candidate variables, were used for modeling. Model uncertainty was estimated by mapping a total variance calculated by adding the weighted average of within-model variation to the between-model variation. The range of cold-evergreens and model performance were validated by true skill statistics, the receiver operating characteristic curve and the kappa statistic. Climate change effects on the cold-evergreens were predicted according to the RCP 4.5 and RCP 8.5 scenarios. Multimodel inference approach excellently projected the spatial distribution of cold-evergreens (AUC = 0.95, kappa = 0.62 and TSS = 0.77). Temperature was a dominant factor in model-average estimates, while precipitation was minor. The climatic suitability increased from the southwest, lowland areas, to the northeast, high mountains. The range of cold-evergreens declined under climate change. Mountain-tops in the south and most of the area in the north remained suitable in 2050 and 2070 under the RCP 4.5 projection and 2050 under the RCP 8.5 projection. Only high-elevations in the northeastern Peninsula remained suitable under the RCP 8.5 projection. A northward and upper-elevational range shift indicates change in species composition at the alpine and subalpine ecosystems in the Korean Peninsula.
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Koo, Kyung Ah; Kong, Woo-Seok; Nibbelink, Nathan P.; Hopkinson, Charles S.; Lee, Joon Ho
2015-01-01
Climate change has caused shifts in species’ ranges and extinctions of high-latitude and altitude species. Most cold-tolerant evergreen broadleaved woody plants (shortened to cold-evergreens below) are rare species occurring in a few sites in the alpine and subalpine zones in the Korean Peninsula. The aim of this research is to 1) identify climate factors controlling the range of cold-evergreens in the Korean Peninsula; and 2) predict the climate change effects on the range of cold-evergreens. We used multimodel inference based on combinations of climate variables to develop distribution models of cold-evergreens at a physiognomic-level. Presence/absence data of 12 species at 204 sites and 6 climatic factors, selected from among 23 candidate variables, were used for modeling. Model uncertainty was estimated by mapping a total variance calculated by adding the weighted average of within-model variation to the between-model variation. The range of cold-evergreens and model performance were validated by true skill statistics, the receiver operating characteristic curve and the kappa statistic. Climate change effects on the cold-evergreens were predicted according to the RCP 4.5 and RCP 8.5 scenarios. Multimodel inference approach excellently projected the spatial distribution of cold-evergreens (AUC = 0.95, kappa = 0.62 and TSS = 0.77). Temperature was a dominant factor in model-average estimates, while precipitation was minor. The climatic suitability increased from the southwest, lowland areas, to the northeast, high mountains. The range of cold-evergreens declined under climate change. Mountain-tops in the south and most of the area in the north remained suitable in 2050 and 2070 under the RCP 4.5 projection and 2050 under the RCP 8.5 projection. Only high-elevations in the northeastern Peninsula remained suitable under the RCP 8.5 projection. A northward and upper-elevational range shift indicates change in species composition at the alpine and subalpine ecosystems in the Korean Peninsula. PMID:26262755
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CO-OCCURRENCE OF OZONE AND ACIDIC CLOUD WATER IN HIGH-ELEVATION FORESTS
A chemical climatology for high-elevation forests was estimated from ozone and cloudwater acidity data collected in the eastern United States. esides frequent ozone-only and pH-only single-pollutant episodes, both simultaneous and sequential co-occurrence of ozone and acidic clou...
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Miao, Ning; Liu, Shi-Rong; Shi, Zuo-Min; Yu, Hong; Liu, Xing-Liang
2009-06-01
Based on the investigation in a 4 hm2 Betula-Abies forest plot in sub-alpine area in West Sichuan of China, and by using point pattern analysis method in terms of O-ring statistics, the spatial patterns of dominant species Betula albo-sinensis and Abies faxoniana in different age classes in study area were analyzed, and the intra- and inter-species associations between these age classes were studied. B. albo-sinensis had a unimodal distribution of its DBH frequency, indicating a declining population, while A. faxoniana had a reverse J-shaped pattern, showing an increasing population. All the big trees of B. albo-sinensis and A. faxoniana were spatially in random at all scales, while the medium age and small trees were spatially clumped at small scales and tended to be randomly or evenly distributed with increasing spatial scale. The maximum aggregation degree decreased with increasing age class. Spatial association mainly occurred at small scales. A. faxoniana generally showed positive intra-specific association, while B. albo-sinensis generally showed negative intra-specific association. For the two populations, big and small trees had no significant spatial association, but middle age trees had negative spatial association. Negative inter-specific associations of the two populations were commonly found in different age classes. The larger the difference of age class, the stronger the negative inter-specific association.
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NASA Astrophysics Data System (ADS)
Baron, J.; Advani, S. M.; Allen, J.; Boot, C.; Denef, K.; Denning, S.; Hall, E.; Moore, J. C.; Reuth, H.; Ryan, M. G.; Shaw, E.
2016-12-01
Long-term field experiments can reveal changes in ecosystem processes that may not be evident in short-term studies. Short-term measurements or experiments may have narrower objectives or unrealistic treatments in order to see a change, whereas long-term studies can reveal complex interactions that take longer to manifest. We report results from a long-term experiment (1996 to present) in subalpine forests to simulate the consequences of sustained atmospheric nitrogen (N) deposition. Loch Vale watershed in Rocky Mountain National Park, the location of the experiment, has received an order of magnitude greater atmospheric N deposition than estimated background since mid-20th Century. Augmenting that, in 1996 we began adding 25 kg NH4NO3 ha-1 yr-1 to three 30m x 30m old-growth Engelmann spruce and subalpine fir plots. Treated stands were matched by nearby controls. N addition caused rapid leaching of nitrate and cations from soils, and increased N mineralization and nitrification rates. These observations in the fertilized plots have been sustained over time. Soluble aluminum concentrations do not differ significantly between fertilized and control plots, but treated soils are now markedly more acidic (pH of 4.7) than original soil and controls (pH of 5.1); further acidification might increase aluminum leaching. Effects on soil carbon were complex, mediated by reductions in total microbial biomass, decreases in arbuscular mychorrizal and saprotropic fungi, and increased potential rates of N enzyme degrading activities. Initial soil C:N of 24 was lower than similar soils in low N deposition stands (C:N of 36). The C:N declined to 22 with treatment. Fertilized plots lost 11% soil C, but the mechanism is unclear. We did not measure changes in C inputs from litter, microbial biomass, or plant uptake, but there was no change in summer CO2 flux, measured in 2003, 2004, and 2014. Leaching of DOC from fertilized plots was elevated throughout the experiment, providing one pathway for C loss. The soil microfauna was dominated by nematodes; plant parasites and bacterial and fungal feeders were more abundant in fertilized plots than in controls, with fewer predaceous and omnivorous nematodes. Overall, N fertilization altered soil biogeochemical characteristics, soil food webs, and C cycling.
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Snowmelt in a High Latitude Mountain Catchment: Effect of Vegetation Cover and Elevation
NASA Astrophysics Data System (ADS)
Pomeroy, J. W.; Essery, R. L.; Ellis, C. R.; Hedstrom, N. R.; Janowicz, R.; Granger, R. J.
2004-12-01
The energetics and mass balance of snowpacks in the premelt and melt period were compared from three elevation bands in a high latitude mountain catchment, Wolf Creek Research Basin, Yukon. Elevation is strongly correlated with vegetation cover and in this case the three elevation bands (low, middle, high) correspond to mature spruce forest, dense shrub tundra and sparse tundra (alpine). Measurements of radiation, ground heat flux, snow depth, snowfall, air temperature, wind speed were made on a half-hourly basis at the three elevations for a 10 year period. Sondes provided vertical gradients of air temperature, humidity, wind speed and air pressure. Snow depth and density surveys were conducted monthly. Comparisons of wind speed, air temperature and humidity at three elevations show that the expected elevational gradients in the free atmosphere were slightly enhanced just above the surface canopies, but that the climate at the snow surface was further influenced by complex canopy effects. Premelt snow accumulation was strongly affected by intercepted snow in the forest and blowing snow sublimation in the sparse tundra but not by the small elevational gradients in snowfall. As a result the maximum premelt SWE was found in the mid-elevation shrub tundra and was roughly double that of the sparse tundra or forest. Minimum variability of SWE was observed in the forest and shrub tundra (CV=0.25) while in the sparse tundra variability doubled (CV=0.5). Snowmelt was influenced by differences in premelt accumulation as well as differences in the net energy fluxes to snow. Elevation had a strong effect on the initiation of melt with the forest melt starting on average 16 days before the shrub tundra and 19 days before the sparse tundra. Mean melt rates showed a maximum in middle elevations and increased from 860 kJ/day in the forest to 1460 kJ/day in the sparse tundra and 2730 kJ/day in the shrub tundra. The forest canopy reduced melt while the shrub canopy enhanced it relative to the sparsely vegetated tundra. Duration of melt was similar in the forest and shrub tundra at 20 days while the sparse tundra was shorter at 13 days; the differences due to differing snow accumulation and melt rates. The greatest variability in the timing and rate of melt was found in the shrub tundra, where the effect of the shrub canopy over snow depends on snow depth and insolation and is reduced in years with high snow accumulation or extensive cloudy periods in spring. The results show that it is necessary to consider the combination of elevation and vegetation effects on snow microclimate and melt processes in high latitude mountain catchments, but that weather patterns induce substantial variability on the effect these factors.
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Samuel, M.D.; Hobbelen, P.H.F.; Decastro, F.; Ahumada, J.A.; Lapointe, D.A.; Atkinson, C.T.; Woodworth, B.L.; Hart, P.J.; Duffy, D.C.
2011-01-01
We developed an epidemiological model of avian malaria (Plasmodium relictum) across an altitudinal gradient on the island of Hawaii that includes the dynamics of the host, vector, and parasite. This introduced mosquito-borne disease is hypothesized to have contributed to extinctions and major shifts in the altitudinal distribution of highly susceptible native forest birds. Our goal was to better understand how biotic and abiotic factors influence the intensity of malaria transmission and impact on susceptible populations of native Hawaiian forest birds. Our model illustrates key patterns in the malaria-forest bird system: high malaria transmission in low-elevation forests with minor seasonal or annual variation in infection;episodic transmission in mid-elevation forests with site-to-site, seasonal, and annual variation depending on mosquito dynamics;and disease refugia in high-elevation forests with only slight risk of infection during summer. These infection patterns are driven by temperature and rainfall effects on parasite incubation period and mosquito dynamics across an elevational gradient and the availability of larval habitat, especially in mid-elevation forests. The results from our model suggest that disease is likely a key factor in causing population decline or restricting the distribution of many susceptible Hawaiian species and preventing the recovery of other vulnerable species. The model also provides a framework for the evaluation of factors influencing disease transmission and alternative disease control programs, and to evaluate the impact of climate change on disease cycles and bird populations. ??2011 by the Ecological Society of America.
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DOE Office of Scientific and Technical Information (OSTI.GOV)
Mohnen, V.A.; Aneja, V.; Bailey, B.
The report summarizes the results of the four year field measurement and data analysis program of MCCP. The MCCP is sponsored by the U.S. Environmental Protection Agency as part of the joint U.S. Forest Service-EPA Spruce-Fir Research Cooperative. The objectives of the project have been met and the result is an assessment of principal atmospheric constituents as they impact the high elevation forests of the Eastern U.S. Deposition of SO4(-2), NO3(-), H(+), and NH(4+) in cloud water represents a significant input to forest canopies with elevations greater than 1000m. Cloud water deposition can exceed wet (rain) deposition and may bemore » the dominant process for input of sulfate and nitrate compounds during the growing season in high elevation forests frequently exposed to clouds. Cloud water pH concentrations may be as much as 0.6pH units lower than pH in rain. SO(4+) and NO3(-) concentrations are also higher in cloud water than in precipitation. Ozone data reveals that significant differences exist between ozone concentrations at high elevation and low elevation sites. The primary effect of the difference is to produce higher mean ozone concentrations and longer episodes at the higher elevation sites.« less
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Mitigating Future Avian Malaria Threats to Hawaiian Forest Birds from Climate Change.
Liao, Wei; Atkinson, Carter T; LaPointe, Dennis A; Samuel, Michael D
2017-01-01
Avian malaria, transmitted by Culex quinquefasciatus mosquitoes in the Hawaiian Islands, has been a primary contributor to population range limitations, declines, and extinctions for many endemic Hawaiian honeycreepers. Avian malaria is strongly influenced by climate; therefore, predicted future changes are expected to expand transmission into higher elevations and intensify and lengthen existing transmission periods at lower elevations, leading to further population declines and potential extinction of highly susceptible honeycreepers in mid- and high-elevation forests. Based on future climate changes and resulting malaria risk, we evaluated the viability of alternative conservation strategies to preserve endemic Hawaiian birds at mid and high elevations through the 21st century. We linked an epidemiological model with three alternative climatic projections from the Coupled Model Intercomparison Project to predict future malaria risk and bird population dynamics for the coming century. Based on climate change predictions, proposed strategies included mosquito population suppression using modified males, release of genetically modified refractory mosquitoes, competition from other introduced mosquitoes that are not competent vectors, evolved malaria-tolerance in native honeycreepers, feral pig control to reduce mosquito larval habitats, and predator control to improve bird demographics. Transmission rates of malaria are predicted to be higher than currently observed and are likely to have larger impacts in high-elevation forests where current low rates of transmission create a refuge for highly-susceptible birds. As a result, several current and proposed conservation strategies will be insufficient to maintain existing forest bird populations. We concluded that mitigating malaria transmission at high elevations should be a primary conservation goal. Conservation strategies that maintain highly susceptible species like Iiwi (Drepanis coccinea) will likely benefit other threatened and endangered Hawai'i species, especially in high-elevation forests. Our results showed that mosquito control strategies offer potential long-term benefits to high elevation Hawaiian honeycreepers. However, combined strategies will likely be needed to preserve endemic birds at mid elevations. Given the delay required to research, develop, evaluate, and improve several of these currently untested conservation strategies we suggest that planning should begin expeditiously.
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Mitigating future avian malaria threats to Hawaiian forest birds from climate change
Liao, Wei; Atkinson, Carter T.; LaPointe, Dennis; Samuel, Michael D.
2017-01-01
Avian malaria, transmitted by Culex quinquefasciatus mosquitoes in the Hawaiian Islands, has been a primary contributor to population range limitations, declines, and extinctions for many endemic Hawaiian honeycreepers. Avian malaria is strongly influenced by climate; therefore, predicted future changes are expected to expand transmission into higher elevations and intensify and lengthen existing transmission periods at lower elevations, leading to further population declines and potential extinction of highly susceptible honeycreepers in mid- and high-elevation forests. Based on future climate changes and resulting malaria risk, we evaluated the viability of alternative conservation strategies to preserve endemic Hawaiian birds at mid and high elevations through the 21st century. We linked an epidemiological model with three alternative climatic projections from the Coupled Model Intercomparison Project to predict future malaria risk and bird population dynamics for the coming century. Based on climate change predictions, proposed strategies included mosquito population suppression using modified males, release of genetically modified refractory mosquitoes, competition from other introduced mosquitoes that are not competent vectors, evolved malaria-tolerance in native honeycreepers, feral pig control to reduce mosquito larval habitats, and predator control to improve bird demographics. Transmission rates of malaria are predicted to be higher than currently observed and are likely to have larger impacts in high-elevation forests where current low rates of transmission create a refuge for highly-susceptible birds. As a result, several current and proposed conservation strategies will be insufficient to maintain existing forest bird populations. We concluded that mitigating malaria transmission at high elevations should be a primary conservation goal. Conservation strategies that maintain highly susceptible species like Iiwi (Drepanis coccinea) will likely benefit other threatened and endangered Hawai’i species, especially in high-elevation forests. Our results showed that mosquito control strategies offer potential long-term benefits to high elevation Hawaiian honeycreepers. However, combined strategies will likely be needed to preserve endemic birds at mid elevations. Given the delay required to research, develop, evaluate, and improve several of these currently untested conservation strategies we suggest that planning should begin expeditiously.
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Yu, Xinxiao; Zhao, Yutao; Zhang, Zhiqiang; Cheng, Genwei
2003-01-01
Dark coniferous forest is the predominant type of vegetation in the upper reaches of Yangtze River. Difference among different types of soil exists. The sand content of soil is higher and the soil texture is coarser in the early stage of forest succession. The sand content of soil decreases with the advancement of the forest succession while that of soil in Abies fabri over-mature forest is the lowest. In slope wash soil, the sand content of soil decreases with the increasing soil depth. The soil porosity and soil water-holding capacity increases and soil bulk density decreases with the advancement of forest succession and decrease of soil depth. The deeper soil depth or the smaller soil water content are, the smaller the unsaturated hydraulic conductivity of soil measured by CGA method. Moreover, the correlation of soil water content with unsaturated hydraulic conductivity of soil can be simulated by an exponential function. The saturated hydraulic conductivity of soil decreases exponentially with the increasing soil depth. The time to attain the stable infiltration rate is different among different soil depth, while the deeper the soil depth is, the longer the time needs. The variation in soil texture, soil physical properties and the high infiltration rate of soil there implicated that there are scarce surface runoff, but abundant in subsurface flow, return flow and seepage, which is the result of regulation by dark coniferous forest on hydrological processes.
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Rediscovery and uncertain future of high-elevation Haleakala carabid beetles (Coleoptera)
Krushelnycky, Paul D.; Gillespie, Rosemary G.; Loope, Lloyd L.; Liebherr, James K.
2005-01-01
Recent biotic surveys in subalpine shrubland on Haleakala¯ Volcano, Maui, Hawai‘i, have resulted in rediscovery of several species of carabid beetles previously known only from their nineteenth-century type specimens. Blackburnia lenta (Sharp), described from specimens collected just below Haleakala¯ summit in 1894, was found at lower elevational sites ranging from 2,400 to 2,750 m. Mecyclothorax rusticus Sharp, last seen in 1896, and M. nubicola (Blackburn), collected only in 1878, were also rediscovered in that vicinity. Recent collections of B. lenta contradict the U.S. Fish and Wildlife Service’s previous classification of this species as one likely to be extinct. Nevertheless, B. lenta’s known distribution comprises only 145 ha within an elevational zone that is bounded above and below by unicolonial populations of the invasive alien Argentine ant, Linepithema humile (Mayr). The known recent collections of M. rusticus and M. nubicola also occurred outside the distributional range of the Argentine ant. Mature eggs held in the lateral oviducts of B. lenta females averaged 1.4 the volume of the largest eggs previously reported among 13 species of Blackburnia. We hypothesize that the giant eggs of B. lenta result from selective forces favoring large, well-nourished developing and hatched first-instar larvae, consistent with a patchy distribution of suitable microhabitat and prey in the subalpine Haleakala¯ landscape. The specialized life history of B. lenta, and coincidence of distributional limits of the three rediscovered carabid species with range limits of the Argentine ant populations suggest that all would be jeopardized by future distributional expansion of Argentine ant. These intersecting phenomena compel us to conclude that B. lenta, M. nubicola, and M. rusticus are appropriate candidates for I.U.C.N. threatened species designation, pending further studies of their geographic ranges and historical trends in abundance.
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NASA Astrophysics Data System (ADS)
Berryman, E.; Frank, J. M.; Speckman, H. N.; Bradford, J. B.; Ryan, M. G.; Massman, W. J.; Hawbaker, T. J.
2017-12-01
Bark beetle outbreaks in Western North American forests are often considered a high-severity disturbance from a carbon (C) cycling perspective, but field measurements that quantify impacts on C dynamics are very limited. Often, factors out of the researcher's control complicate the separation of beetle impacts from other drivers of C cycling variability and restrict statistical inference. Fortuitously, we had four years of pre-spruce beetle outbreak C cycle measurements in a subalpine forest in southeastern Wyoming (Glacier Lakes Ecosystem Experiments Site, or GLEES) and sustained intermittent monitoring for nearly a decade after the outbreak. Here, we synthesize published and unpublished pre- and post-outbreak measurements of key C cycle stocks and fluxes at GLEES. Multiple lines of evidence, including chamber measurements, eddy covariance measurements, and tracking of soil and forest floor C pools over time, point to the GLEES outbreak as a moderate-severity disturbance for C loss to the atmosphere, despite 70% to 80% of overstory tree death. Reductions in NEE were short-lived and the forest quickly returned to a carbon-neutral state, likely driven by an uptick in understory growth. Effect of mortality on the C cycle was asymmetrical, with a 50% reduction in net carbon uptake (NEE) two years into the outbreak, yet no measureable change in either ecosystem or growing season soil respiration. A small pulse in soil respiration occurred but was only detectable during the winter and amounted to < 10% of NEE. Possible reasons for the lack of measureable respiration response are discussed with emphasis on lessons learned for monitoring and modeling future outbreaks. We suggest a comprehensive assessment and definition of "moderate-severity" disturbances for Western forests and suggest that all tree mortality events may not be high-severity when it comes to C fluxes.
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NASA Astrophysics Data System (ADS)
Poinsatte, J. P.; Rochefort, R.; Evans, R. D.
2014-12-01
Elevated anthropogenic nitrogen (N) emissions result in higher rates of atmospheric N deposition (Ndep) that can saturate sensitive ecosystems. Consequences of increased Ndep include higher emissions of greenhouse gases, eutrophication of watersheds, and deterioration of vegetation communities. Most of the annual N deposition at higher elevations in the Cascades is stored in snowpack until spring snowmelt when it is released as a pulse that can be assimilated by plant and microbial communities, or lost as gaseous emissions or leachate. The relative magnitude of these fluxes is unknown, particularly with accelerated rates of snowpack loss due to climate change. We quantified storage of Ndep in winter snowpack and determined impacts of Ndep on biogeochemical processes in a lush-herbaceous community characterized by Valeriana sitchensis and Lupinus latifolius, a heath-shrub community characterized by Phyllodoce empetriformis and Cassiope mertensiana, and a wet-sedge community dominated by Carex nigricans. These communities were selected to represent early, mid, and late snowmelt vegetation regimes prevalent throughout the Cascades. Ammonium (NH4+) was the dominant form of Ndep in winter snowpack and Ndep rates were higher than anticipated based on nearby National Atmospheric Deposition Program (NADP) measurements. Vegetation N uptake was the dominant N sink in the ecosystem, with the highest growing season uptake occurring in the lush-herbaceous community, while soil N leaching was the dominant N loss, with the lush-herbaceous also having the highest rates. Microbial biomass N fluctuated substantially across the growing season, with high biomass N immediately after snowmelt and again 30 days following snow release. Soil nitrous oxide (N2O) emissions peaked 30 days following snowmelt for all three communities and were greatest in the wet sedge community. These results indicate that subalpine communities have unique responses to Ndep that vary throughout the growing season. Thus, biogeochemical modeling efforts to simulate ecosystem responses to Ndep should be parameterized at a community-specific level to fully capture this variability. Ultimately, this study will provide insight to land managers on the fate of N emissions and how Ndep affects ecosystem services in high-elevation ecosystems.
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Yin, Huajun; Li, Yufei; Xiao, Juan; Xu, Zhenfeng; Cheng, Xinyin; Liu, Qing
2013-07-01
Despite the perceived importance of exudation to forest ecosystem function, few studies have attempted to examine the effects of elevated temperature and nutrition availability on the rates of root exudation and associated microbial processes. In this study, we performed an experiment in which in situ exudates were collected from Picea asperata seedlings that were transplanted in disturbed soils exposed to two levels of temperature (ambient temperature and infrared heater warming) and two nitrogen levels (unfertilized and 25 g N m(-2) a(-1) ). Here, we show that the trees exposed to an elevated temperature increased their exudation rates I (μg C g(-1) root biomass h(-1) ), II (μg C cm(-1) root length h(-1) ) and III (μg C cm(-2) root area h(-1) ) in the unfertilized plots. The altered morphological and physiological traits of the roots exposed to experimental warming could be responsible for this variation in root exudation. Moreover, these increases in root-derived C were positively correlated with the microbial release of extracellular enzymes involved in the breakdown of organic N (R(2) = 0.790; P = 0.038), which was coupled with stimulated microbial activity and accelerated N transformations in the unfertilized soils. In contrast, the trees exposed to both experimental warming and N fertilization did not show increased exudation rates or soil enzyme activity, indicating that the stimulatory effects of experimental warming on root exudation depend on soil fertility. Collectively, our results provide preliminary evidence that an increase in the release of root exudates into the soil may be an important physiological adjustment by which the sustained growth responses of plants to experimental warming may be maintained via enhanced soil microbial activity and soil N transformation. Accordingly, the underlying mechanisms by which plant root-microbe interactions influence soil organic matter decomposition and N cycling should be incorporated into climate-carbon cycle models to determine reliable estimates of long-term C storage in forests. © 2013 Blackwell Publishing Ltd.
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Bai, Fan; Sang, WeiGuo; Li, GuangQi; Liu, RuiGang; Chen, LingZhi; Wang, Kun
2008-10-01
The Changbai Mountain Nature Reserve (CNR) was established in 1960 to protect the virgin Korean pine mixed hardwood forest, a typical temperate forest of northeast China. We conducted systematic studies of vascular diversity patterns on the north slope of the CNR mountainside forests (800-1700 m a.s.l.) in 1963 and 2006 respectively. The aim of this comparison is to assess the long-term effects of the protection on plant biodiversity of CNR during the interval 43 years. The research was carried out in three types of forests: mixed coniferous and broad-leaved forest (MCBF), mixed coniferous forest (MCF), and sub-alpine coniferous forest (SCF), characterized by different dominant species. The alpha diversity indicted by species richness and the Shannon-Wiener index were found different in the same elevations and forest types during the 43-year interval. The floral composition and the diversity of vascular species were generally similar along altitudinal gradients before and after the 43-year interval, but some substantial changes were evident with the altitude gradient. In the tree layers, the dominant species in 2006 were similar to those of 1963, though diversity declined with altitude. The indices in the three forest types did not differ significantly between 1963 and 2006, and these values even increased in the MCBF and MCF from 1963 to 2006. However, originally dominant species, P. koraiensis for example, tended to decline, while the proportion of broad-leaved trees increased, and the species turnover in the succession layers trended to shift to higher altitudes. The diversity pattern of the under canopy fluctuated along the altitudinal gradient due to micro-environmental variations. Comparison of the alpha diversity in the three forests shows that the diversity of the shrub and herb layer decreased with time. During the process of survey, we also found some rare and medicinal species disappeared. Analysis indicates that the changes of the diversity pattern in this region are caused by both nature and human factors. Meteorological records revealed that climate has changed significantly in the past 43 years. We also found the most severe human disturbance to the CNR forests in the process of another field survey that is the exploitation of herb medicines and Korean pine nuts. We hope this research would give some guidance to the future reserve management in Changbai Mountain area.
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NASA Astrophysics Data System (ADS)
Webb, Ryan W.
2017-09-01
Snow is an important environmental variable in headwater systems that controls hydrological processes such as streamflow, groundwater recharge, and evapotranspiration. These processes will be affected by both the amount of snow available for melt and the rate at which it melts. Snow water equivalent (SWE) and snowmelt are known to vary within complex subalpine terrain due to terrain and canopy influences. This study assesses this variability during the melt season using ground penetrating radar to survey multiple plots in northwestern Colorado near a snow telemetry (SNOTEL) station. The plots include south aspect and flat aspect slopes with open, coniferous (subalpine fir, Abies lasiocarpa and engelman spruce, Picea engelmanii), and deciduous (aspen, populous tremuooides) canopy cover. Results show the high variability for both SWE and loss of SWE during spring snowmelt in 2014. The coefficient of variation for SWE tended to increase with time during snowmelt whereas loss of SWE remained similar. Correlation lengths for SWE were between two and five meters with melt having correlation lengths between two and four meters. The SNOTEL station regularly measured higher SWE values relative to the survey plots but was able to reasonably capture the overall mean loss of SWE during melt. Ground Penetrating Radar methods can improve future investigations with the advantage of non-destructive sampling and the ability to estimate depth, density, and SWE.
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Predicting forest road surface erosion and storm runoff from high-elevation sites
J. M. Grace III
2017-01-01
Forest roads are a concern in management because they represent areas of elevated risks associated with soil erosion and storm runoff connectivity to stream systems. Storm runoff emanating from forest roads and their connectivity to downslope resources can be influenced by a myriad of factors, including storm characteristics, management practices, and the interaction...
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Heather Lessig; William J. McShea; Jeffrey R. Walters
2010-01-01
The southern Appalachians support a unique forest ecosystem at higher elevations in which the breeding distribution of several bird species of conservation concern extends to unusually southern latitudes. The dual threats of rising global temperatures and potential wind energy development may impact these forests by reducing or fragmenting preferred habitat.
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Forest Species Diversity in Upper Elevation Hardwood Forests in the Southern Appalachian Mountains
Katherine J. Elliott; Deidre Hewitt
1997-01-01
Overstory, shrub-layer, and herb-layer flora composition and abundance patterns in eleven forest sites were studied to evaluate species diversity and richness before implementing three types of harvest treat- ments. The sites were within the Wine Spring Creek Watershed and were classified as high elevation, dry, Quercus rubra-Rhododendron calendulaceum based on...
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Forest habitat associations of the golden-mantled ground squirrel: Implications for fuels management
Katharine R. Shick; Dean E. Pearson; Leonard F. Ruggiero
2006-01-01
Golden-mantled ground squirrels are commonly associated with high-elevation habitats near or above upper timberline. This species also occurs in fire-adapted, low-elevation forests that are targeted for forest health restoration (FHR) treatments intended to remove encroaching understory trees and thin overstory trees. Hence, the golden-mantled ground squirrel...
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Deposition of Mercury in Forests along a Montane Elevation Gradient.
Blackwell, Bradley D; Driscoll, Charles T
2015-05-05
Atmospheric mercury (Hg) deposition varies along elevation gradients and is influenced by both orographic and biological factors. We quantified total Hg deposition over a 2 year period at 24 forest sites at Whiteface Mountain, NY, USA, that ranged from 450 to 1450 m above sea level and covered three distinct forest types: deciduous/hardwood forest (14.1 μg/m2-yr), spruce/fir forest (33.8 μg/m2-yr), and stunted growth alpine/fir forest (44.0 μg/m2-yr). Atmospheric Hg deposition increased with elevation, with the dominant deposition pathways shifting from litterfall in low-elevation hardwoods to throughfall in midelevation spruce/fir to cloudwater in high-elevation alpine forest. Soil Hg concentrations (ranging from 69 to 416 ng/g for the Oi/Oe and 72 to 598 ng/g for the Oa horizons) were correlated with total Hg deposition, but the weakness of the correlations suggests that additional factors such as climate and tree species also contribute to soil Hg accumulation. Meteorological conditions influenced Hg deposition pathways, as cloudwater Hg diminished in 2010 (dry conditions) compared to 2009 (wet conditions). However, the dry conditions in 2010 led to increased Hg dry deposition and subsequent significant increases in throughfall Hg fluxes compared to 2009. These findings suggest that elevation, forest characteristics, and meteorological conditions are all important drivers of atmospheric Hg deposition to montane forests.
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Climate-induced changes in high elevation stream nitrate dynamics
Baron, Jill S.; Schmidt, T.M.; Hartman, M.D.
2009-01-01
Mountain terrestrial and aquatic ecosystems are responsive to external drivers of change, especially climate change and atmospheric deposition of nitrogen (N). We explored the consequences of a temperature-warming trend on stream nitrate in an alpine and subalpine watershed in the Colorado Front Range that has long been the recipient of elevated atmospheric N deposition. Mean annual stream nitrate concentrations since 2000 are higher by 50% than an earlier monitoring period of 1991-1999. Mean annual N export increased by 28% from 2.03 kg N ha-1yr-1 before 2000 to 2.84 kg N ha-1yr-1 in Loch Vale watershed since 2000. The substantial increase in N export comes as a surprise, since mean wet atmospheric N deposition from 1991 to 2006 (3.06 kg N ha-1 yr-1) did not increase. There has been a period of below average precipitation from 2000 to 2006 and a steady increase in summer and fall temperatures of 0.12??C yr-1 in both seasons since 1991. Nitrate concentrations, as well as the weathering products calcium and sulfate, were higher for the period 2000-2006 in rock glacier meltwater at the top of the watershed above the influence of alpine and subalpine vegetation and soils. We conclude the observed recent N increases in Loch Vale are the result of warmer summer and fall mean temperatures that are melting ice in glaciers and rock glaciers. This, in turn, has exposed sediments from which N produced by nitrification can be flushed. We suggest a water quality threshold may have been crossed around 2000. The phenomenon observed in Loch Vale may be indicative of N release from ice features such as rock glaciers worldwide as mountain glaciers retreat. ?? 2009 Blackwell Publishing Ltd.
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Schaeffer, Sean M.; Anderson, Dean E.; Burns, Sean P.; Monson, Russell K.; Sun, Jielun; Bowling, David R.
2008-01-01
Stable isotopes provide insight into ecosystem carbon cycling, plant physiological processes, atmospheric boundary-layer dynamics, and are useful for the integration of processes over multiple scales. Of particular interest is the carbon isotope content (δ13C) of nocturnal ecosystem-respired CO2 (δR). Recent advances in technology have made it possible to continuously examine the variation in δR within a forest canopy over relatively long time-scales (months–years). We used tunable diode laser spectroscopy to examine δR at within- and below-canopy spatial locations in a Colorado subalpine forest (the Niwot Ridge AmeriFlux site). We found a systematic pattern of increased δR within the forest canopy (δR-c) compared to that near the ground (δR-g). Values of δR-c were weakly correlated with the previous day's mean maximum daytime vapor pressure deficit (VPD). Conversely, there was a negative but still weak correlation between δR-g and time-lagged (0–5 days) daily mean soil moisture. The topography and presence of sustained nightly drainage flows at the Niwot Ridge forest site suggests that, on nights with stable atmospheric conditions, there is little mixing of air near the ground with that in the canopy. Atmospheric stability was assessed using thresholds of friction velocity, stability above the canopy, and bulk Richardson number within the canopy. When we selectively calculated δR-g and δR-c by removing time periods when ground and canopy air were well mixed, we found stronger correlations between δR-c and VPD, and δR-g and soil moisture. This suggests that there may be fundamental differences in the environmental controls on δR at sub-canopy spatial scales. These results may help explain the wide variance observed in the correlation of δR with different environmental parameters in other studies.
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Lithosequence of soils and associated vegetation on subalpine range of the Wasatch Plateau, Utah.
James O. Klemmedson; Arthur R. Tiedemann
1998-01-01
On degraded subalpine range in Utah, the authors examined the role of soil and parent material nutrients and organic carbon (Corg) in the development of soil and plants on a transect across six strata that formed visible concentric alternating bands of high and low productivity. Relations for soil and parent material phosphorus (P) and sulfur (S) were of particular...
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Buma, Brian
2012-06-01
Forest disturbances around the world have the potential to alter forest type and cover, with impacts on diversity, carbon storage, and landscape composition. These disturbances, especially fire, are common and often large, making ground investigation of forest recovery difficult. Remote sensing offers a means to monitor forest recovery in real time, over the entire landscape. Typically, recovery monitoring via remote sensing consists of measuring vegetation indices (e.g., NDVI) or index-derived metrics, with the assumption that recovery in NDVI (for example) is a meaningful measure of ecosystem recovery. This study tests that assumption using MODIS 16-day imagery from 2000 to 2010 in the area of the Colorado's Routt National Forest Hinman burn (2002) and seedling density counts taken in the same area. Results indicate that NDVI is rarely correlated with forest recovery, and is dominated by annual and perennial forb cover, although topography complicates analysis. Utility of NDVI as a means to delineate areas of recovery or non-recovery are in doubt, as bootstrapped analysis indicates distinguishing power only slightly better than random. NDVI in revegetation analyses should carefully consider the ecology and seasonal patterns of the system in question.
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Binkley, Dan; Olsson, U.; Rochelle, R.; Stohlgren, T.; Nikolov, N.
2003-01-01
Old-growth forests of Engelmann spruce (Picea engelmannii Parry ex. Engelm.) and subalpine fir (Abies lasiocarpa (Hook.) Nutt.) dominate much of the landscape of the Rocky Mountains. We characterized the structure, biomass and production of 18 old-growth (200-450-year-old) spruce/fir forests in Rocky Mountain National Park, Colorado, as well as the stand-level supply and use of light and nitrogen. Stands were chosen to span a broad range of elevation, aspect, and topography. Aboveground tree biomass in these old-growth forests averaged 253 Mg/ha (range 130-488 Mg/ha), with aboveground net primary production of 3700 kg ha-1 yr-1 (range from 2700 to 5200 kg ha-1 yr-1). Within stands, trees >35 cm in diameter accounted for 70% of aboveground biomass, but trees <35 cm contributed 70% of the production of woody biomass. Differences in slope and aspect among sites resulted in a range of incoming light from 58 to 74 TJ ha-1 yr-1, and tree canopies intercepted an average of 71% of incoming light (range 50-90%). Aboveground net primary production (ANPP) of trees did not relate to the supply of light or N, but ANPP correlated strongly with the amount of light and N used (r2 = 0.45-0.54, P < 0.01). Uptake of 1 kg of N was associated with about 260 kg of ANPP, and one TJ of intercepted shortwave radiation produced about 78 kg of ANPP. Across these old-growth stands, stands with greater biomass showed higher rates of both ANPP and resource use; variation in aboveground biomass was associated with 24% of the variation in N use (P = 0.04), 44% of the light use (P = 0.003), and 45% of the ANPP (P = 0.002). ?? 2002 Elsevier Science B.V. All rights reserved.
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Talbot, Stephen S.; Markon, Carl J.
1988-01-01
A Landsat-derived vegetation map was prepared for lnnoko National Wildlife Refuge. The refuge lies within the northern boreal subzone of northwestern central Alaska. Six major vegetation classes and 21 subclasses were recognized: forest (closed needleleaf, open needleleaf, needleleaf woodland, mixed, and broadleaf); broadleaf scrub (lowland, upland burn regeneration, subalpine); dwarf scrub (prostrate dwarf shrub tundra, erect dwarf shrub heath, dwarf shrub-graminoid peatland, dwarf shrub-graminoid tussock peatland, dwarf shrub raised bog with scattered trees, dwarf shrub-graminoid marsh); herbaceous (graminoid bog, graminoid marsh, graminoid tussock-dwarf shrub peatland); scarcely vegetated areas (scarcely vegetated scree and floodplain); and water (clear, sedimented). The methodology employed a cluster-block technique. Sample areas were described based on a combination of helicopter-ground survey, aerial photo-interpretation, and digital Landsat data. Major steps in the Landsat analysis involved preprocessing (geometric correction), derivation of statistical parameters for spectral classes, spectral class labeling of sample areas, preliminary classification of the entire study area using a maximum-likelihood algorithm, and final classification utilizing ancillary information such as digital elevation data. The final product is 1:250,000-scale vegetation map representative of distinctive regional patterns and suitable for use in comprehensive conservation planning.
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NASA Astrophysics Data System (ADS)
Mercer, J.; Liefert, D. T.; Shuman, B. N.; Befus, K. M.; Williams, D. G.; Kraushaar, B.
2017-12-01
Alpine and subalpine lakes are important components of the hydrologic cycle in mountain ecosystems. These lakes are also highly sensitive to small shifts in temperature and precipitation. Mountain lake volumes and their contributions to mountain hydrology may change in response to even minor declines in snowpack or increases in temperature. However, it is still not clear to what degree non-climatic factors, such as geomorphic setting and lake geometry, play in shaping the sensitivity of high elevation lakes to climate change. We investigated the importance of lake geometry and groundwater connectivity to mountain lakes in the Snowy Range, Wyoming using a combination of hydrophysical and hydrochemical methods, including stable water isotopes, to better understand the role these factors play in controlling lake volume. Water isotope values in open lakes were less sensitive to evaporation compared to those in closed basin lakes. Lake geometry played an important role, with wider, shallower lakes being more sensitive to evaporation over time. Groundwater contributions appear to play only a minor role in buffering volumetric changes to lakes over the growing season. These results confirm that mountain lakes are sensitive to climate factors, but also highlight a significant amount of variability in that sensitivity. This research has implications for water resource managers concerned with downstream water quantity and quality from mountain ecosystems, biologists interested in maintaining aquatic biodiversity, and paleoclimatologists interested in using lake sedimentary information to infer past climate regimes.
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de Wit, Heleen A; Bryn, Anders; Hofgaard, Annika; Karstensen, Jonas; Kvalevåg, Maria M; Peters, Glen P
2014-07-01
Expanding high-elevation and high-latitude forest has contrasting climate feedbacks through carbon sequestration (cooling) and reduced surface reflectance (warming), which are yet poorly quantified. Here, we present an empirically based projection of mountain birch forest expansion in south-central Norway under climate change and absence of land use. Climate effects of carbon sequestration and albedo change are compared using four emission metrics. Forest expansion was modeled for a projected 2.6 °C increase in summer temperature in 2100, with associated reduced snow cover. We find that the current (year 2000) forest line of the region is circa 100 m lower than its climatic potential due to land-use history. In the future scenarios, forest cover increased from 12% to 27% between 2000 and 2100, resulting in a 59% increase in biomass carbon storage and an albedo change from 0.46 to 0.30. Forest expansion in 2100 was behind its climatic potential, forest migration rates being the primary limiting factor. In 2100, the warming caused by lower albedo from expanding forest was 10 to 17 times stronger than the cooling effect from carbon sequestration for all emission metrics considered. Reduced snow cover further exacerbated the net warming feedback. The warming effect is considerably stronger than previously reported for boreal forest cover, because of the typically low biomass density in mountain forests and the large changes in albedo of snow-covered tundra areas. The positive climate feedback of high-latitude and high-elevation expanding forests with seasonal snow cover exceeds those of afforestation at lower elevation, and calls for further attention of both modelers and empiricists. The inclusion and upscaling of these climate feedbacks from mountain forests into global models is warranted to assess the potential global impacts. © 2013 John Wiley & Sons Ltd.
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NASA Astrophysics Data System (ADS)
Roth, Travis R.; Nolin, Anne W.
2017-11-01
Forest cover modifies snow accumulation and ablation rates via canopy interception and changes in sub-canopy energy balance processes. However, the ways in which snowpacks are affected by forest canopy processes vary depending on climatic, topographic and forest characteristics. Here we present results from a 4-year study of snow-forest interactions in the Oregon Cascades. We continuously monitored snow and meteorological variables at paired forested and open sites at three elevations representing the Low, Mid, and High seasonal snow zones in the study region. On a monthly to bi-weekly basis, we surveyed snow depth and snow water equivalent across 900 m transects connecting the forested and open pairs of sites. Our results show that relative to nearby open areas, the dense, relatively warm forests at Low and Mid sites impede snow accumulation via canopy snow interception and increase sub-canopy snowpack energy inputs via longwave radiation. Compared with the Forest sites, snowpacks are deeper and last longer in the Open site at the Low and Mid sites (4-26 and 11-33 days, respectively). However, we see the opposite relationship at the relatively colder High sites, with the Forest site maintaining snow longer into the spring by 15-29 days relative to the nearby Open site. Canopy interception efficiency (CIE) values at the Low and Mid Forest sites averaged 79 and 76 % of the total event snowfall, whereas CIE was 31 % at the lower density High Forest site. At all elevations, longwave radiation in forested environments appears to be the primary energy component due to the maritime climate and forest presence, accounting for 93, 92, and 47 % of total energy inputs to the snowpack at the Low, Mid, and High Forest sites, respectively. Higher wind speeds in the High Open site significantly increase turbulent energy exchanges and snow sublimation. Lower wind speeds in the High Forest site create preferential snowfall deposition. These results show the importance of understanding the effects of forest cover on sub-canopy snowpack evolution and highlight the need for improved forest cover model representation to accurately predict water resources in maritime forests.
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Malhi, Yadvinder; Girardin, Cécile A J; Goldsmith, Gregory R; Doughty, Christopher E; Salinas, Norma; Metcalfe, Daniel B; Huaraca Huasco, Walter; Silva-Espejo, Javier E; Del Aguilla-Pasquell, Jhon; Farfán Amézquita, Filio; Aragão, Luiz E O C; Guerrieri, Rossella; Ishida, Françoise Yoko; Bahar, Nur H A; Farfan-Rios, William; Phillips, Oliver L; Meir, Patrick; Silman, Miles
2017-05-01
Why do forest productivity and biomass decline with elevation? To address this question, research to date generally has focused on correlative approaches describing changes in woody growth and biomass with elevation. We present a novel, mechanistic approach to this question by quantifying the autotrophic carbon budget in 16 forest plots along a 3300 m elevation transect in Peru. Low growth rates at high elevations appear primarily driven by low gross primary productivity (GPP), with little shift in either carbon use efficiency (CUE) or allocation of net primary productivity (NPP) between wood, fine roots and canopy. The lack of trend in CUE implies that the proportion of photosynthate allocated to autotrophic respiration is not sensitive to temperature. Rather than a gradual linear decline in productivity, there is some limited but nonconclusive evidence of a sharp transition in NPP between submontane and montane forests, which may be caused by cloud immersion effects within the cloud forest zone. Leaf-level photosynthetic parameters do not decline with elevation, implying that nutrient limitation does not restrict photosynthesis at high elevations. Our data demonstrate the potential of whole carbon budget perspectives to provide a deeper understanding of controls on ecosystem functioning and carbon cycling. © 2016 The Authors. New Phytologist © 2016 New Phytologist Trust.
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NASA Astrophysics Data System (ADS)
Millar, C.; Westfall, R. D.; Delany, D.
2016-12-01
In the Great Basin of southwestern USA, limber pine (Pinus flexilis) is a common subalpine species, often forming the upper treeline of the central to northern mountain ranges in this region. Multiple rainshadows, created by successive mountain ranges inland from Pacific-dominated storm tracks, leave interior ranges arid. Combined with cool climate, minimal alpine and subalpine herbaceous vegetation, and lack of landscape-scale fires or biotic disturbance, dead wood of limber pine persists for millennia across the mountain slopes. Using dendroecological methods, we studied distribution and ages of live and relict wood in the Wassuk Range (summit elevation, 3440m), west-central Great Basin. Currently live limber pines grow sparsely on north slopes, whereas relict wood, with stem diameters to 1 m and lengths to 10 m, is widely distributed. We cross-dated 440 limber pine stems and relict wood from 9 sites; taken altogether, limber pines have grown without gap across the last 3597 years in this mountain range. The mean elevation range of live trees is 3078m to 2821m, which is not significantly different from the mean elevation range of relict wood, which is 3096m to 2816m. Relict wood occurred on all main aspects, with age pulses of colonization and extirpation over time. Colonization periods related to the ends of centennial-scale dry periods, known from other proxies in the region. Extirpations, by corollary, roughly coincided with these long droughts, demonstrating successive diminishment of limber pine from all but north slopes over four millennia. The last gasp of pines on non-north aspects was during the warm, dry Medieval Climate Anomaly, ca 1000 yrs ago. That pines did not shift upward during warm or dry historic periods, given 340m available above uppermost tree distribution, suggests that significant climate variability was met by shifting aspect rather than elevation in this range.
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Nutrient status in soil of Ski runs in the sub-alpine belt of Uludag Mountain, Bursa, Turkey.
Guleryuz, Gurcan; Kirmizi, Serap; Arslan, Hulya
2010-01-01
Large areas of land are disturbed in sensitive bio-diverse mountain environments by Skiruns. Restoration of vegetation on such disturbed mountain sites may be hampered by soil degradation but the severity and nature of the constraints is not well understood. This study was designed to compare the water holding and nutritional status of soil in three Ski runs which had different construction dates and disturbance levels, and the adjacent undisturbed site in the Abies bommuelleriana forest community in the sub-alpine belt of Uludag Mountain (Bithynian Olympus). The values of soil parameters were depressed in proportion to the disturbance level. Water holding capacity (WHC), total nitrogen (N), organic carbon (C) and calcium (Ca2+), magnesium (Mg2+) and potassium (K+) contents (mg kg(-1) dry weight) of soils in the Ski run which had the highest disturbance level were lower than that of the undisturbed adjacent sites. However the results indicated that the soil parameters were less degraded when secondary vegetation was growing on the disturbed areas.
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Remote sensing for conservation of tropical moist forests: A study in Indonesia
NASA Astrophysics Data System (ADS)
Warwick-Smith, Robert Myles
The Indonesian archipelago extends in a great 6000km arc from the northern tip of Sumatra to the eastern border of Irian Jaya. It includes a wide diversity of ecosystems ranging from the floristically rich and economically important lowland tropical rain forests to the 'moss' and sub-alpine meadows of the higher mountains and from fresh-water swamp forest to the dry monsoon forest and savanna woodlands of the lesser Sunda islands. These forests are of importance for the protection of watersheds and catchment areas, for the maintenance of water supplies, and for their general and local influence upon climate. They are the habitat of a large number of rare, endangered and endemic plant and animal species; also many other birds, mammals, reptiles and insects which form a colourful, scientifically valuable and irreplaceable part of the national heritage and world genetic resources. This study examines an area of great ecological importance in Sulawesi, and an attempt is made to map a number of ecosystems in the area. Landsat multispectral imagery (1972) was the basis of the mapping and field work was completed in 1980. The satellite imagery proved to be a satisfactory mapping tool in these tropical moist forest conditions.
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NASA Astrophysics Data System (ADS)
Porinchu, D.; Reinemann, S.; Potito, A.; Moser, K.; MacDonald, G.; Munroe, J.; Mark, B.; Box, J.
2007-12-01
Subfossil midge analyses have been used to develop high-resolution (sub-decadal) reconstructions of 20th century temperature change in the Sierra Nevada, CA with success. Expansion of this earlier work to additional sites in the western United States suggests that a widespread increase in lake water temperatures has occurred in this region during the late 20th and early 21st centuries. Inference models for summer surface water temperature (SSWT) were developed combining midge abundance data from 56 lakes in the eastern Sierra Nevada, California, with subfossil midge remains from the Uinta Mountains, UT. The newly merged Sierra Nevada-Uinta Mountains calibration set contains a greater diversity of chironomid assemblages and spans a wider SSWT range than the previously published Sierra Nevada calibration set. The lakes in the merged calibration set spanned elevation, depth, and SSWT temperature ranges of 900 m, 12.7 m, and 11.3 °C, respectively. A robust inference model for SSWT (3-component WA-PLS), based on 90 lakes, had a high coefficient of determination (r2jack = 0.66) and a low RMSEP (1.4 °C). The midge-based SSWT inference model was applied to subfossil chironomid remains extracted from well-dated sediment sequences recovered from alpine and subalpine lakes in the Sierra Nevada, CA, Snake Range, NV and Uinta Mountains, UT. A close correspondence exists between the chironomid-inferred temperature profiles for the 20th and 21st centuries and mean July or summer temperatures measured at nearby meteorological stations. Application of this midge-based SSWT inference model to other intact, late Quaternary sedimentary sequences found in subalpine and alpine lakes in the Great Basin will help resolve the impact of late Quaternary and recent climate change in this region, improve our understanding of regional climate and aquatic ecosystem variability, and can be used to monitor the effects of climate change on aquatic ecosystems and establish 'baseline' conditions against which future biotic changes can be compared.
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Impact of snow gliding on soil redistribution for a sub-alpine area in Switzerland
NASA Astrophysics Data System (ADS)
Meusburger, K.; Leitinger, G.; Mabit, L.; Mueller, M. H.; Alewell, C.
2013-07-01
The aim of this study is to assess the importance of snow gliding as soil erosion agent for four different land use/land cover types in a sub-alpine area in Switzerland. The 14 investigated sites are located close to the valley bottom at approximately 1500 m a.s.l., while the elevation of the surrounding mountain ranges is about 2500 m a.s.l. We used two different approaches to estimate soil erosion rates: the fallout radionuclide 137Cs and the Revised Universal Soil Loss Equation (RUSLE). The RUSLE model is suitable to estimate soil loss by water erosion, while the 137Cs method integrates soil loss due to all erosion agents involved. Thus, we hypothesise that the soil erosion rates determined with the 137Cs method are higher and that the observed discrepancy between the erosion rate of RUSLE and the 137Cs method is related to snow gliding. Cumulative snow glide distance was measured for the sites in the winter 2009/2010 and modelled for the surrounding area with the Spatial Snow Glide Model (SSGM). Measured snow glide distance range from 0 to 189 cm with lower values for the north exposed slopes. We observed a reduction of snow glide distance with increasing surface roughness of the vegetation, which is an important information with respect to conservation planning and expected land use changes in the Alps. Our hypothesis was confirmed, the difference of RUSLE and 137Cs erosion rates was correlated to the measured snow glide distance (R2 = 0.73; p < 0.005). A high difference (lower proportion of water erosion compared to total net erosion) was observed for high snow glide rates and vice versa. The SSGM reproduced the relative difference of the measured snow glide values between different land use/land cover types. The resulting map highlights the relevance of snow gliding for large parts of the investigated area. Based on these results, we conclude that snow gliding is a key process impacting soil erosion pattern and magnitude in sub-alpine areas with similar topographic and climatic conditions.
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Tang, Shi-shan; Yang, Wan-qin; Wang, Hai-peng; Xiong, Li; Nie, Fu-yu; Xu, Shen-feng
2015-10-01
In this study, a buried bag experiment was used to investigate mass loss and C, N and P release patterns of fine (≤2 mm), medium (2-5 mm) and coarse (≥ 5 mm) roots of 3 subalpine dominant trees, i. e., Betula albosinensis, Abies faxoniana and Picea asperata in the growing and non-growing seasons. In general, the remaining mass of B. albosinensis was lower than that of A. faxoniana and P. asperata. In addition, root remaining mass increased with the increase of root diameter for the same species. The mass losing rate in the non-growing season was 52.1%-64.4% of a year. The C release of B. albosinensis was the highest, but that of A. faxoniana was the lowest. Also, C release decreased with the increase of root diameter. N of A. faxoniana and P. asperata were enriched in the non-growing season but released in the growing season. However, the opposite pattern was found for B. albosinensis. During the non-growing season, the amount of N enrichment increased with the increase of root diameter. The P release of 3 species was characterized as the enrichment-release pattern. P enrichment of A. faxoniana was significantly greater than that of P. asperata and B. albosinensis. Nevertheless, no significant difference was observed between diameter sizes. In conclusion, diameter size had significant effect on root decomposition in the subalpine forests of western Sichuan, and the diameter effect was dependent on tree species and season.
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John D. Lloyd; Julie Hart; J. Dan Lambert
2010-01-01
Birds occupying high-elevation forests in the northeast are perceived to be at risk from a variety of external forces, most notably the potential loss and alteration of habitat associated with global climate change and the increased deployment of wind-energy facilities. However, the Breeding Bird Survey (BBS), a standardized national monitoring scheme widely used to...
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Hrinkevich, Kathryn H; Progar, Robert A; Shaw, David C
2016-01-01
The balsam woolly adelgid (Adelges piceae (Ratzeburg) (Homoptera: Adelgidae)) (BWA) is a nonnative, invasive insect that threatens Abies species throughout North America. It is well established in the Pacific Northwest, but continues to move eastward through Idaho and into Montana and potentially threatens subalpine fir to the south in the central and southern Rocky Mountains. We developed a climatic risk model and map that predicts BWA impacts to subalpine fir using a two-step process. Using 30-year monthly climate normals from sites with quantitatively derived BWA damage severity index values, we built a regression model that significantly explained insect damage. The sites were grouped into two distinct damage categories (high damage and mortality versus little or no mortality and low damage) and the model estimates for each group were used to designate distinct value ranges for four climatic risk categories: minimal, low, moderate, and high. We then calculated model estimates for each cell of a 4-kilometer resolution climate raster and mapped the risk categories over the entire range of subalpine fir in the western United States. The spatial variation of risk classes indicates a gradient of climatic susceptibility generally decreasing from the Olympic Peninsula in Washington and the Cascade Range in Oregon and Washington moving eastward, with the exception of some high risk areas in northern Idaho and western Montana. There is also a pattern of decreasing climatic susceptibility from north to south in the Rocky Mountains. Our study provides an initial step for modeling the relationship between climate and BWA damage severity across the range of subalpine fir. We showed that September minimum temperature and a metric calculated as the maximum May temperature divided by total May precipitation were the best climatic predictors of BWA severity. Although winter cold temperatures and summer heat have been shown to influence BWA impacts in other locations, these variables were not as predictive as spring and fall conditions in the Pacific Northwest.
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NASA Astrophysics Data System (ADS)
Kerkez, B.; Rice, R.; Glaser, S. D.; Bales, R. C.; Saksa, P. C.
2010-12-01
A 100-node wireless sensor network (WSN) was designed for the purpose of monitoring snow depth in two watersheds, spanning 3 km2 in the American River basin, in the central Sierra Nevada of California. The network will be deployed as a prototype project that will become a core element of a larger water information system for the Sierra Nevada. The site conditions range from mid-elevation forested areas to sub-alpine terrain with light forest cover. Extreme temperature and humidity fluctuations, along with heavy rain and snowfall events, create particularly challenging conditions for wireless communications. We show how statistics gathered from a previously deployed 60-node WSN, located in the Southern Sierra Critical Zone Observatory, were used to inform design. We adapted robust network hardware, manufactured by Dust Networks for highly demanding industrial monitoring, and added linear amplifiers to the radios to improve transmission distances. We also designed a custom data-logging board to interface the WSN hardware with snow-depth sensors. Due to the large distance between sensing locations, and complexity of terrain, we analyzed network statistics to select the location of repeater nodes, to create a redundant and reliable mesh. This optimized network topology will maximize transmission distances, while ensuring power-efficient network operations throughout harsh winter conditions. At least 30 of the 100 nodes will actively sense snow depth, while the remainder will act as sensor-ready repeaters in the mesh. Data from a previously conducted snow survey was used to create a Gaussian Process model of snow depth; variance estimates produced by this model were used to suggest near-optimal locations for snow-depth sensors to measure the variability across a 1 km2 grid. We compare the locations selected by the sensor placement algorithm to those made through expert opinion, and offer explanations for differences resulting from each approach.
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Samuel, Michael D.; Woodworth, Bethany L.; Atkinson, Carter T.; Hart, P. J.; LaPointe, Dennis
2015-01-01
Wildlife diseases can present significant threats to ecological systems and biological diversity, as well as domestic animal and human health. However, determining the dynamics of wildlife diseases and understanding the impact on host populations is a significant challenge. In Hawai‘i, there is ample circumstantial evidence that introduced avian malaria (Plasmodium relictum) has played an important role in the decline and extinction of many native forest birds. However, few studies have attempted to estimate disease transmission and mortality, survival, and individual species impacts in this distinctive ecosystem. We combined multi-state capture-recapture (longitudinal) models with cumulative age-prevalence (cross-sectional) models to evaluate these patterns in Apapane, Hawai‘i Amakihi, and Iiwi in low-, mid-, and high-elevation forests on the island of Hawai‘i based on four longitudinal studies of 3–7 years in length. We found species-specific patterns of malaria prevalence, transmission, and mortality rates that varied among elevations, likely in response to ecological factors that drive mosquito abundance. Malaria infection was highest at low elevations, moderate at mid elevations, and limited in high-elevation forests. Infection rates were highest for Iiwi and Apapane, likely contributing to the absence of these species in low-elevation forests. Adult malaria fatality rates were highest for Iiwi, intermediate for Amakihi at mid and high elevations, and lower for Apapane; low-elevation Amakihi had the lowest malaria fatality, providing strong evidence of malaria tolerance in this low-elevation population. Our study indicates that hatch-year birds may have greater malaria infection and/or fatality rates than adults. Our study also found that mosquitoes prefer feeding on Amakihi rather than Apapane, but Apapane are likely a more important reservoir for malaria transmission to mosquitoes. Our approach, based on host abundance and infection rates, may be an effective alternative to mosquito blood meal analysis for determining vector-host contacts when mosquito densities are low and collection of blood-fed mosquitoes is impractical. Our study supports the hypothesis that avian malaria has been a primary factor influencing the elevational distribution and abundance of these three species, and likely limits other native Hawaiian species that are susceptible to malaria.
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Effects of coffee management on deforestation rates and forest integrity.
Hylander, Kristoffer; Nemomissa, Sileshi; Delrue, Josefien; Enkosa, Woldeyohannes
2013-10-01
Knowledge about how forest margins are utilized can be crucial for a general understanding of changes in forest cover, forest structure, and biodiversity across landscapes. We studied forest-agriculture transitions in southwestern Ethiopia and hypothesized that the presence of coffee (Coffea arabica)decreases deforestation rates because of coffee's importance to local economies and its widespread occurrence in forests and forest margins. Using satellite images and elevation data, we compared changes in forest cover over 37 years (1973-2010) across elevations in 2 forest-agriculture mosaic landscapes (1100 km(2) around Bonga and 3000 km(2) in Goma-Gera). In the field in the Bonga area, we determined coffee cover and forest structure in 40 forest margins that differed in time since deforestation. Both the absolute and relative deforestation rates were lower at coffee-growing elevations compared with at higher elevations (-10/20% vs. -40/50% comparing relative rates at 1800 m asl and 2300-2500 m asl, respectively). Within the coffee-growing elevation, the proportion of sites with high coffee cover (>20%) was significantly higher in stable margins (42% of sites that had been in the same location for the entire period) than in recently changed margins (0% of sites where expansion of annual crops had changed the margin). Disturbance level and forest structure did not differ between sites with 30% or 3% coffee. However, a growing body of literature on gradients of coffee management in Ethiopia reports coffee's negative effects on abundances of forest-specialist species. Even if the presence of coffee slows down the conversion of forest to annual-crop agriculture, there is a risk that an intensification of coffee management will still threaten forest biodiversity, including the genetic diversity of wild coffee. Conservation policy for Ethiopian forests thus needs to develop strategies that acknowledge that forests without coffee production may have higher deforestation risks than forests with coffee production and that forests with coffee production often have lower biodiversity value. © 2013 Society for Conservation Biology.
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NASA Astrophysics Data System (ADS)
Hu, Jia; Jencso, Kelsey; Looker, Nathaniel; Martin, Justin; Hoylman, Zachary
2015-04-01
Across the Western U.S., declining snowpacks have resulted in increased water limitation, leading to reduced productivity in high elevation forests. While our current understanding of how forests respond to climate change is typically focused on measuring/modeling the physiological responses and climate feedbacks, our study aims to combine physiology with hydrology to examine how landscape topography modulates the sensitivity of forests to climate. In a forested watershed in Western Montana, we linked climate variability to the physical watershed characteristics and the physiological response of vegetation to examine forest transpiration and productivity rates. Across the entire watershed, we found a strong relationship between productivity and the topographic wetness index, a proxy for soil moisture storage. However, this relationship was highly dependent on the intensity of solar radiation, suggesting that at high elevations productivity was limited by temperature, while at low elevations productivity was limited by moisture. In order to identify the mechanisms responsible for this relationship, we then examined how different coniferous species respond to changing environmental and hydrologic regimes. We first examined transpiration and productivity rates at the hillslope scale at four plots, ranging in elevation and aspect across the watershed. We found trees growing in the hollows had higher transpiration and productivity rates than trees growing in the side slope, but that these differences were more pronounced at lower elevations. We then used oxygen isotope to examine water source use by different species across the watershed. We found that trees growing in the hollows used snowmelt for a longer period. This was most likely due to upslope subsidies of snowmelt water to the hollow areas. However, we found that trees growing at lower elevations used proportionally more snowmelt than trees at the higher elevations. This was most likely due to the trees at lower elevation depending on deeper, more reliable water when the upper soils dried down during midsummer. These observations suggest that landscape topography influences the availability of soil water, which influences tree transpiration and productivity rates, thereby leading to watershed patterns of productivity.
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Historical harvests reduce neighboring old-growth basal area across a forest landscape.
Bell, David M; Spies, Thomas A; Pabst, Robert
2017-07-01
While advances in remote sensing have made stand, landscape, and regional assessments of the direct impacts of disturbance on forests quite common, the edge influence of timber harvesting on the structure of neighboring unharvested forests has not been examined extensively. In this study, we examine the impact of historical timber harvests on basal area patterns of neighboring old-growth forests to assess the magnitude and scale of harvest edge influence in a forest landscape of western Oregon, USA. We used lidar data and forest plot measurements to construct 30-m resolution live tree basal area maps in lower and middle elevation mature and old-growth forests. We assessed how edge influence on total, upper canopy, and lower canopy basal area varied across this forest landscape as a function of harvest characteristics (i.e., harvest size and age) and topographic conditions in the unharvested area. Upper canopy, lower canopy, and total basal area increased with distance from harvest edge and elevation. Forests within 75 m of harvest edges (20% of unharvested forests) had 4% to 6% less live tree basal area compared with forest interiors. An interaction between distance from harvest edge and elevation indicated that elevation altered edge influence in this landscape. We observed a positive edge influence at low elevations (<800 m) and a negative edge influence at moderate to high elevations (>800 m). Surprisingly, we found no or weak effects of harvest age (13-60 yr) and harvest area (0.2-110 ha) on surrounding unharvested forest basal area, implying that edge influence was relatively insensitive to the scale of disturbance and multi-decadal recovery processes. Our study indicates that the edge influence of past clearcutting on the structure of neighboring uncut old-growth forests is widespread and persistent. These indirect and diffuse legacies of historical timber harvests complicate forest management decision-making in old-growth forest landscapes by broadening the traditional view of stand boundaries. Furthermore, the consequences of forest harvesting may reach across ownership boundaries, highlighting complex governance issues surrounding landscape management of old-growth forests. © 2017 by the Ecological Society of America.
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Carbon stocks and fluxes in fire disturbed landscapes of Colorado, U.S.A.
NASA Astrophysics Data System (ADS)
Barnes, R. T.; Wolf, K.; Whittinghill, K. A.; Gilbertson, A.; Buma, B.
2016-12-01
In terrestrial ecosystems, ecological disturbances can strongly regulate material and energy flows. This often results from the reduction in biomass and associated ecological relationships and physiological processes. Researchers have noted an increase in the size and severity of disturbances, such as wildfire, in recent decades. While there is significant research examining post-disturbance carbon stocks and recovery, there is less known about the fate and quality of post-disturbance carbon pools. In an effort to understand the recovery and resilience of forest carbon stocks to severe wildfire we examined the carbon and black carbon (pyrogenic) stocks (e.g. above ground biomass, coarse woody debris, charcoal, soils) and export fluxes (stream export, soil leachate, soil respiration) within the burn scars and nearby reference sites of five 2002 Colorado fires. The fires encompass large precipitation and ecosystem gradients (relatively dry montane Ponderosa forests to relatively wet subalpine Lodgepole forests), allowing us to control for various state factors in our analyses. With the exception of the Hinman fire (subalpine, Lodgepole dominated), there is little forest regrowth more than a decade later, with only a few saplings found in burned plots; instead forbes and grasses dominate. Fire also reduces soil C stocks (by 16 to 68%) across all sites. In addition, with the shifts in vegetation we hypothesize that there will be corresponding changes in soil organic matter (SOM), altering the residence time of C in soil. Soil incubation experiments reveal that organic matter bioavailability is significantly greater in three of the burned sites, suggesting that the new sources of SOM are more bioavailable. Stable isotopic analyses of SOM and the evolved CO2 from the incubation studies will allow us to test this hypothesis. Fire also affects the amount and nature of dissolved and particulate organic matter (DOM and POM, respectively) leaving the watershed. For example, the dissolved organic matter exported from burned watersheds is less aromatic and has lower C:N than DOM exported from reference watersheds. In addition to measuring these fluxes, we assessed the fate of leached DOM via laboratory bioassays to determine the likelihood of additional CO2 losses to the atmosphere.
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The Use of Sun Elevation Angle for Stereogrammetric Boreal Forest Height in Open Canopies
NASA Technical Reports Server (NTRS)
Montesano, Paul M.; Neigh, Christopher; Sun, Guoqing; Duncanson, Laura Innice; Van Den Hoek, Jamon; Ranson, Kenneth Jon
2017-01-01
Stereogrammetry applied to globally available high resolution spaceborne imagery (HRSI; less than 5 m spatial resolution) yields fine-scaled digital surface models (DSMs) of elevation. These DSMs may represent elevations that range from the ground to the vegetation canopy surface, are produced from stereoscopic image pairs (stereo pairs) that have a variety of acquisition characteristics, and have been coupled with lidar data of forest structure and ground surface elevation to examine forest height. This work explores surface elevations from HRSI DSMs derived from two types of acquisitions in open canopy forests. We (1) apply an automated mass-production stereogrammetry workflow to along-track HRSI stereo pairs, (2) identify multiple spatially coincident DSMs whose stereo pairs were acquired under different solar geometry, (3) vertically co-register these DSMs using coincident spaceborne lidar footprints (from ICESat-GLAS) as reference, and(4) examine differences in surface elevations between the reference lidar and the co-registered HRSI DSMs associated with two general types of acquisitions (DSM types) from different sun elevation angles. We find that these DSM types, distinguished by sun elevation angle at the time of stereo pair acquisition, are associated with different surface elevations estimated from automated stereogrammetry in open canopy forests. For DSM values with corresponding reference ground surface elevation from spaceborne lidar footprints in open canopy northern Siberian Larix forests with slopes less than10, our results show that HRSI DSM acquired with sun elevation angles greater than 35deg and less than 25deg (during snow-free conditions) produced characteristic and consistently distinct distributions of elevation differences from reference lidar. The former include DSMs of near-ground surfaces with root mean square errors less than 0.68 m relative to lidar. The latter, particularly those with angles less than 10deg, show distributions with larger differences from lidar that are associated with open canopy forests whose vegetation surface elevations are captured. Terrain aspect did not have a strong effect on the distribution of vegetation surfaces. Using the two DSM types together, the distribution of DSM-differenced heights in forests (6.0 m, sigma = 1.4 m) was consistent with the distribution of plot-level mean tree heights (6.5m, sigma = 1.2 m). We conclude that the variation in sun elevation angle at time of stereo pair acquisition can create illumination conditions conducive for capturing elevations of surfaces either near the ground or associated with vegetation canopy. Knowledge of HRSI acquisition solar geometry and snow cover can be used to understand and combine stereogrammetric surface elevation estimates to co-register rand difference overlapping DSMs, providing a means to map forest height at fine scales, resolving the vertical structure of groups of trees from spaceborne platforms in open canopy forests.
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Forest diversity and disturbance: changing influences and the future of Virginia's Forests
Christine J. Small; James L. Chamberlain
2015-01-01
The Virginia landscape supports a remarkable diversity of forests, from maritime dunes, swamp forests, and pine savannas of the Atlantic coastal plain, to post-agricultural pine-hardwood forests of the piedmont, to mixed oak, mixed-mesophytic, northern hardwood, and high elevation conifer forests in Appalachian mountain provinces. Virginiaâs forests also have been...
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Patterns of reptile and amphibian species richness along elevational gradients in Mt. Kenya
MALONZA, Patrick Kinyatta
2015-01-01
Faunal species richness is traditionally assumed to decrease with increasing elevation and decreasing primary productivity. Species richness is reported to peak at mid-elevation. This survey examines the herpetofaunal diversity and distribution in Mt. Kenya (central Kenya) by testing the hypothesis that changes in species richness with elevation relate to elevation-dependent changes in climate. Sampling along transects from an elevation of approximately 1 700 m in Chogoria forest block (wind-ward side) and approximately 2 600 m in Sirimon block (rain shadow zone) upwards in March 2009. This starts from the forest to montane alpine zones. Sampling of reptiles and amphibians uses pitfall traps associated with drift fences, time-limited searches and visual encounter surveys. The results show that herpetofaunal richness differs among three vegetation zones along the elevation gradient. Chogoria has higher biodiversity than Sirimon. More species occur at low and middle elevations and few exist at high elevations. The trends are consistent with expected optimum water and energy variables. The lower alpine montane zone has high species richness but low diversity due to dominance of some high elevations species. Unambiguous data do not support a mid-domain effect (mid-elevation peak) because the observed trend better fits a model in which climatic variables (rainfall and temperature) control species richness, which indirectly measures productivity. It is important to continue protection of all indigenous forests, especially at low to mid elevations. These areas are vulnerable to human destruction yet are home to some endemic species. Firebreaks can limit the spread of the perennial wildfires, especially on the moorlands. PMID:26646571
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The South's forestland - on the hot seat to provide more
Raymond M. Sheffield; James G. Dickson
1998-01-01
Forests of the Southern United States range from tropical/subtropical forests on the southern extremities of the region, oak savanna forests on the western fringe, to central hardwood forests, and high elevation boreal forests in the north. Upland and bottomland hardwood, southern pine, and mixed pine-hardwood forests are found on the more moderate sites between these...
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Jia, Shuzheng; Nakano, Takashi; Hattori, Masahira; Nara, Kazuhide
2017-11-01
Pyroleae species are perennial understory shrubs, many of which are partial mycoheterotrophs. Most fungi colonizing Pyroleae roots are ectomycorrhizal (ECM) and share common mycobionts with their Pyroleae hosts. However, such mycobiont sharing has neither been examined in depth before nor has the interspecific variation in sharing among Pyroleae species. Here, we examined root-associated fungal communities in three co-existing Pyroleae species, including Pyrola alpina, Pyrola incarnata, and Orthilia secunda, with reference to co-existing ECM fungi on the surrounding trees in the same soil blocks in subalpine coniferous forests. We identified 42, 75, and 18 fungal molecular operational taxonomic units in P. alpina, P. incarnata, and O. secunda roots, respectively. Mycobiont sharing with surrounding trees, which was defined as the occurrence of the same mycobiont between Pyroleae and surrounding trees in each soil block, was most frequent among P. incarnata (31 of 44 plants). In P. alpina, sharing was confirmed in 12 of 37 plants, and the fungal community was similar to that of P. incarnata. Mycobiont sharing was least common in O. secunda, found in only 5 of 32 plants. Root-associated fungi of O. secunda were dominated by Wilcoxina species, which were absent from the surrounding ECM roots in the same soil blocks. These results indicate that mycobiont sharing with surrounding trees does not equally occur among Pyroleae plants, some of which may develop independent mycorrhizal associations with ECM fungi, as suggested in O. secunda at our research sites.
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Emissions pathways, climate change, and impacts on California
Hayhoe, K.; Cayan, D.; Field, C.B.; Frumhoff, P.C.; Maurer, E.P.; Miller, N.L.; Moser, S.C.; Schneider, S.H.; Cahill, K.N.; Cleland, E.E.; Dale, L.; Drapek, R.; Hanemann, R.M.; Kalkstein, L.S.; Lenihan, J.; Lunch, C.K.; Neilson, R.P.; Sheridan, S.C.; Verville, J.H.
2004-01-01
The magnitude of future climate change depends substantially on the greenhouse gas emission pathways we choose. Here we explore the implications of the highest and lowest Intergovernmental Panel on Climate Change emissions pathways for climate change and associated impacts in California. Based on climate projections from two state-of-the-art climate models with low and medium sensitivity (Parallel Climate Model and Hadley Centre Climate Model, version 3, respectively), we find that annual temperature increases nearly double from the lower B1 to the higher A1fi emissions scenario before 2100. Three of four simulations also show greater increases in summer temperatures as compared with winter. Extreme heat and the associated impacts on a range of temperature-sensitive sectors are substantially greater under the higher emissions scenario, with some interscenario differences apparent before midcentury. By the end of the century under the B1 scenario, heatwaves and extreme heat in Los Angeles quadruple in frequency while heat-related mortality increases two to three times; alpine/subalpine forests are reduced by 50-75%; and Sierra snowpack is reduced 30-70%. Under A1fi, heatwaves in Los Angeles are six to eight times more frequent, with heat-related excess mortality increasing five to seven times; alpine/subalpine forests are reduced by 75-90%; and snowpack declines 73-90%, with cascading impacts on runoff and streamflow that, combined with projected modest declines in winter precipitation, could fundamentally disrupt California's water rights system. Although interscenario differences in climate impacts and costs of adaptation emerge mainly in the second half of the century, they are strongly dependent on emissions from preceding decades.
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Mountain landscapes offer few opportunities for high-elevation tree species migration
Bell, David M.; Bradford, John B.; Lauenroth, William K.
2014-01-01
Climate change is anticipated to alter plant species distributions. Regional context, notably the spatial complexity of climatic gradients, may influence species migration potential. While high-elevation species may benefit from steep climate gradients in mountain regions, their persistence may be threatened by limited suitable habitat as land area decreases with elevation. To untangle these apparently contradictory predictions for mountainous regions, we evaluated the climatic suitability of four coniferous forest tree species of the western United States based on species distribution modeling (SDM) and examined changes in climatically suitable areas under predicted climate change. We used forest structural information relating to tree species dominance, productivity, and demography from an extensive forest inventory system to assess the strength of inferences made with a SDM approach. We found that tree species dominance, productivity, and recruitment were highest where climatic suitability (i.e., probability of species occurrence under certain climate conditions) was high, supporting the use of predicted climatic suitability in examining species risk to climate change. By predicting changes in climatic suitability over the next century, we found that climatic suitability will likely decline, both in areas currently occupied by each tree species and in nearby unoccupied areas to which species might migrate in the future. These trends were most dramatic for high elevation species. Climatic changes predicted over the next century will dramatically reduce climatically suitable areas for high-elevation tree species while a lower elevation species, Pinus ponderosa, will be well positioned to shift upslope across the region. Reductions in suitable area for high-elevation species imply that even unlimited migration would be insufficient to offset predicted habitat loss, underscoring the vulnerability of these high-elevation species to climatic changes.
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McKnight, Diane M.; Harnish, R.; Wershaw, R. L.; Baron, Jill S.; Schiff, S.
1997-01-01
The chemical relationships among particulate and colloidal organic material and dissolved fulvic acid were examined in an alpine and subalpine lake and two streams in Loch Vale Watershed, Rocky Mountain National Park. The alpine lake, Sky Pond, had the lowest dissolved organic carbon (DOC) (0.37 mgC/L), the highest particulate carbon (POC) (0.13 mgC/L), and high algal biomass. The watershed of Sky Pond is primarily talus slope, and DOC and POC may be autochthonous. Both Andrews Creek and Icy Brook gain DOC as they flow through wet sedge meadows. The subalpine lake, The Loch, receives additional organic material from the surrounding forest and had a higher DOC (0.66 mgC/L). Elemental analysis, stable carbon isotopic compositon, and 13C-NMR characterization showed that: 1) particulate material had relatively high inorganic contents and was heterogeneous in compositon, 2) colloidal material was primarily carbohydrate material with a low inorganic content at all sites; and 3) dissolved fulvic acid varied in compositon among sites. The low concentration and carbohydrate-rich character of the colloidal material suggests that this fraction is labile to microbial degradation and may be turning over more rapidly than particulate fractions or dissolved fulvic acid. Fulvic acid from Andrews Creek had the lowest N content and aromaticity, whereas Sky Pond fulvic acid had a higher N content and lower aromaticity than fulvic acid from The Loch. The UV-visible spectra of the fulvic acids demonstrate that variation in characteristics with sources of organic carbon can explain to some extent the observed nonlinear relationship between UV-B extinction coefficients and DOC concentrations in lakes.
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NASA Astrophysics Data System (ADS)
Zhang, Shengrui; Xiao, Jule; Xu, Qinghai; Wen, Ruilin; Fan, Jiawei; Huang, Yun; Yamagata, Hideki
2018-06-01
The response of vegetation to extreme cold events during the last deglaciation is important for assessing the impact of possible extreme climatic events on terrestrial ecosystems under future global warming scenarios. Here, we present a detailed record of the development of regional vegetation in the northern margin of Asian summer monsoon during the last deglaciation (16,500-11,000 cal yr BP) based on a radiocarbon-dated high-resolution pollen record from Hulun Lake, northeast China. The results show that the regional vegetation changed from subalpine meadow-desert steppe to mixed coniferous and deciduous forest-typical steppe during the last deglaciation. However, its responses to the Heinrich event 1 (H1) and the Younger Dryas event (YD) were significantly different: during the H1 event, scattered sparse forest was present in the surrounding mountains, while within the lake catchment the vegetation cover was poor and was dominated by desert steppe. In contrast, during the YD event, deciduous forest developed and the proportion of coniferous forest increased in the mountains, the lake catchment was occupied by typical steppe. We suggest that changes in Northern Hemisphere summer insolation and land surface conditions (ice sheets and sea level) caused temperature and monsoonal precipitation variations that contributed to the contrasting vegetation response during the two cold events. We conclude that under future global warming scenarios, extreme climatic events may cause a deterioration of the ecological environment of the Hulun Lake region, resulting in increased coniferous forest and decreased total forest cover in the surrounding mountains, and a reduction in typical steppe in the lake catchment.
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Host and geographic structure of endophytic and endolichenic fungi at a continental scale.
U'Ren, Jana M; Lutzoni, François; Miadlikowska, Jolanta; Laetsch, Alexander D; Arnold, A Elizabeth
2012-05-01
Endophytic and endolichenic fungi occur in healthy tissues of plants and lichens, respectively, playing potentially important roles in the ecology and evolution of their hosts. However, previous sampling has not comprehensively evaluated the biotic, biogeographic, and abiotic factors that structure their communities. Using molecular data we examined the diversity, composition, and distributions of 4154 endophytic and endolichenic Ascomycota cultured from replicate surveys of ca. 20 plant and lichen species in each of five North American sites (Madrean coniferous forest, Arizona; montane semideciduous forest, North Carolina; scrub forest, Florida; Beringian tundra and forest, western Alaska; subalpine tundra, eastern central Alaska). Endolichenic fungi were more abundant and diverse per host species than endophytes, but communities of endophytes were more diverse overall, reflecting high diversity in mosses and lycophytes. Endophytes of vascular plants were largely distinct from fungal communities that inhabit mosses and lichens. Fungi from closely related hosts from different regions were similar in higher taxonomy, but differed at shallow taxonomic levels. These differences reflected climate factors more strongly than geographic distance alone. Our study provides a first evaluation of endophytic and endolichenic fungal associations with their hosts at a continental scale. Both plants and lichens harbor abundant and diverse fungal communities whose incidence, diversity, and composition reflect the interplay of climatic patterns, geographic separation, host type, and host lineage. Although culture-free methods will inform future work, our study sets the stage for empirical assessments of ecological specificity, metabolic capability, and comparative genomics.
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The last millennium of Aleutian low variability based on dendrochonolgy and water isotope proxies
NASA Astrophysics Data System (ADS)
Gaglioti, B.; Mann, D. H.; Andreu-Hayles, L.; Wiles, G. C.; Streverler, G.; Williams, P.; Field, R. D.; D'Arrigo, R.
2017-12-01
How the wintertime Aleutian Low pressure system (AL) will respond to climate forcing is germane to the forest resources, glaciers, and fisheries in the North Pacific region. Recent work suggests the AL has strengthened over the last few centuries, and new, high-resolution paleo-records could help evaluate this trend. Namely, when it started, whether it has any historical precedents, and what it means for high-latitude climate feedbacks? Our study area is Southeast Alaska and the Southwest Yukon Territory, where the winters are warmer and wetter when the AL is stronger (deeper) and positioned further east. First, we use newly developed winter-sensitive tree-ring records to determine how AL variability has changed over the past 1000 years. During winters with a stronger AL, meridional flow brings: (i) heavy ice storms and snow loads to the coastal mountains that can injure sub-alpine trees; (ii) thaw events that can compromise growth in coastal forests; and (iii) changes in seasonality that can moderate the climate sensitivity of mid-elevation trees. Therefore, a time series of changing tree architecture in the mountains, tree ring widths in the lowlands, and running inter-series correlation in mid-slope chronologies collectively provide a landscape-level view of paleo-AL variability. Second, we compare and complement this dendro perspective with new proxy records based on stable oxygen isotope ratios measured in tree-ring and peat cellulose (d18O cell). To interpret how these proxies reflect the AL, we analyze the precipitation isotopes in Gustavus, Alaska over the past 16 months. We then use d18Ocell time series along a coastal to inland transect that records the degree of heavy-isotope rainout over the St. Elias Mountains, a factor that depends on the sources and pathways of the dominant storm tracks, which are influenced by the AL strength. We will present these new data in the context of existing AL reconstructions, and discuss the implications for the ongoing trends, and environmental corollaries of AL strength.
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Devlaeminck, Rebecca; De Schrijver, An; Hermy, Martin
2005-01-20
Throughfall deposition and canopy exchange of acidifying and eutrophying compounds and major base cations were studied by means of throughfall analysis in a deciduous beech (Fagus sylvatica L.) forest edge in Belgium over a period of 1 year. Throughfall fluxes of Cl(-), NH(4)(+) and Na(+) were significantly elevated at the forest edge compared to the forest interior. As no edge effect on throughfall water volume could be detected, the observed edge enhancement effects were mainly due to dry deposition and canopy exchange patterns. Indeed, there was an elevated dry deposition of Cl(-), Na(+), K(+), Ca(2+) and Mg(2+) up to 50 m from the field/forest border. Within the forest, throughfall and dry deposition of SO(4)(2-) were highly variable and no significant differences were observed between the forest edge and the forest interior. Leaching of K(+) and Ca(2+) was reduced in the forest edge up to a distance of 30 m from the border. The measured nitrogen and acidic depositions far exceeded the current Flemish critical loads with respect to the protection of biodiversity in forests, especially at the forest edge. This points to an urgent need for controlling emissions as well as the need to consider the elevated deposition load in forest edges when calculating the critical loads in forests.
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Mihai, Bogdan; Săvulescu, Ionuț; Rujoiu-Mare, Marina; Nistor, Constantin
2017-12-01
The paper explores the dynamics of the forest cover change in the Iezer Mountains, part of Southern Carpathians, in the context of the forest ownership recovery and deforestation processes, combined with the effects of biotic and abiotic disturbances. The aim of the study is to map and evaluate the typology and the spatial extension of changes in the montane forest cover between 700 and 2462m a.s.l., sampling all the representative Carpathian ecosystems, from the European beech zone up to the spruce-fir zone and the subalpine-alpine pastures. The methodology uses a change detection analysis of satellite imagery with Landsat ETM+/OLI and Sentinel-2 MSI data. The workflow started with a complete calibration of multispectral data from 2002, before the massive forest restitution to private owners, after the Law 247/2005 empowerment, and 2015, the intensification of deforestation process. For the data classification, a Maximum Likelihood supervised classification algorithm was utilized. The forest change map was developed after combining the classifications in a unitary formula using image difference. The principal outcome of the research identifies the type of forest cover change using a quantitative formula. This information can be integrated in the future decision-making strategies for forest stand management and sustainable development. Copyright © 2017 Elsevier B.V. All rights reserved.
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Palila Restoration: Lessons from Long-term Research
,
2006-01-01
BACKGROUND The palila (Loxioides bailleui) is a member of the Hawaiian honeycreeper family of birds (Drepanidinae), which is renowned for the profusion of species - many with bizarre bills and specialized feeding habits - that radiated from a single ancestral type. Most of the 57 or so honeycreeper species are extinct, and the palila is endangered because of its high degree of dependence on the mamane tree (Sophora chrysophylla) (Figure 1) and its restricted distribution on the upper slopes of Mauna Kea (Figure 2). Three decades of research have revealed many important facts about palila, providing the foundation and impetus for conservation programs in the wild and captivity. Additionally, an ambitious public conservation campaign arose due to the land-use conflicts on Mauna Kea. Here we summarize progress in palila conservation biology and outline steps that might overcome the remaining major challenges to its recovery. We also highlight lessons learned from palila research that may help the recovery of other Hawaiian forest birds. Palila and two closely-related species on the tiny islands of Nihoa and Laysan are the last of the seed-eating honeycreeper species in the Hawaiian Islands. About a quarter of the honeycreeper species known from living and fossil specimens had finch-like bills suited mainly for eating seeds and fruits. Because of their dietary specialization, palila are vulnerable to changes in forest size and quality, as was also likely the case for extinct species of seed specialists. Palila and many other forest bird species were once distributed in dry, lowland forests. Fossil records indicate that palila also occurred in the lowlands of O`ahu and Kaua`i until human settlement of those islands. However, because lowland habitats have been highly modified by humans and because mamane occurs today primarily at high elevation, palila are the only native bird species found exclusively in dry, subalpine habitat (2000?2850 m). Similar to other feeding specialists, palila lay few eggs, raise few young each year, and take a relatively long time to complete the nesting cycle. Low rates of reproduction result in low rates of population growth and low potential for recovery from disturbances. Long-term studies of palila offer important insights into the conservation biology of all Hawaiian forest bird species, particularly feeding specialists like the palila. Palila face many challenges common to both generalist and specialist Hawaiian honeycreeper species. Habitat loss and degradation, as well as introduced avian diseases, have reduced their numbers and limited their distribution to a very small portion of their historic range. Introduced mammals prey on palila, while alien insects reduce caterpillars that are particularly important in the diet of nestlings. Securing legal protection and funding for palila restoration has been challenging. Understanding how the palila has avoided extinction can help managers plan its recovery, and better design recovery plans for species with different feeding strategies in other habitats.
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Small-scale topography modulates elevational α-, β- and γ-diversity of Andean leaf beetles.
Thormann, Birthe; Ahrens, Dirk; Espinosa, Carlos Iván; Armijos, Diego Marín; Wagner, Thomas; Wägele, Johann W; Peters, Marcell K
2018-05-01
Elevational diversity gradients are typically studied without considering the complex small-scale topography of large mountains, which generates habitats of strongly different environmental conditions within the same elevational zones. Here we analyzed the importance of small-scale topography for elevational diversity patterns of hyperdiverse tropical leaf beetles (Coleoptera: Chrysomelidae). We compared patterns of elevational diversity and species composition of beetles in two types of forests (on mountain ridges and in valleys) and analyzed whether differences in the rate of species turnover among forest habitats lead to shifts in patterns of elevational diversity when scaling up from the local study site to the elevational belt level. We sampled beetle assemblages at 36 sites in the Podocarpus National Park, Ecuador, which were equally distributed over two forest habitats and three elevational levels. DNA barcoding and Poisson tree processes modelling were used to delimitate putative species. On average, local leaf beetle diversity showed a clear hump-shaped pattern. However, only diversity in forests on mountain ridges peaked at mid-elevation, while beetle diversity in valleys was similarly high at low- and mid-elevation and only declined at highest elevations. A higher turnover of species assemblages at lower than at mid-elevations caused a shift from a hump-shaped diversity pattern found at the local level to a low-elevation plateau pattern (with similar species numbers at low and mid-elevation) at the elevational belt level. Our study reveals an important role of small-scale topography and spatial scale for the inference on gradients of elevational species diversity.
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Effects of elevated CO2 and temperature on forest floor litter decomposition and chemistry
Forest floor can be a major component of the carbon held in forested soils. In mature forests it represents the balance between additions and decomposition under current climate conditions. Because of its position at the soil surface, this reservoir of C is highly susceptible...
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James S. Rentch; Thomas M. Schuler
2010-01-01
The proceedings includes 18 peer-reviewed papers and 41 abstracts pertaining to acid deposition and nutrient cycling, ecological classification, forest dynamics, avifauna, wildlife and fisheries, forests pests, climate change, old-growth forest structure, regeneration, and restoration.
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Development of partial rock veneers by root throw in a subalpine setting
Osterkamp, W.R.; Toy, T.J.; Lenart, M.T.
2006-01-01
Rock veneers stabilize hillslope surfaces, occur especially in areas of immature soil, and form through a variety of process sets that includes root throw. Near Westcliffe, Colorado, USA, data were collected from a 20 ?? 500 m transect on the east slope of the Sangre de Cristo Mountains. Ages of pit/mound complexes with rock fragments exposed at the surface by root throw ranged from recent (freshly toppled tree) to unknown (complete tree decay). Calculations based on dimensions of the pit/mound complexes, estimated time of free topppling, sizes of exposed rock fragments, and percentage rock covers at pit/mound complexes, as well as within the transect area, indicate that recent rates of root throw have resulted in only partial rock veneering since late Pleistocene deglaciation. Weathering of rock fragments prevent development of an extensive rock veneer and causes a balance, achieved within an estimated 700 years, between the rates of rock-fragment exposure by root throw and clast disintegration by chemical reduction. The estimated rate of rock-fragment reduction accounts for part of the fluvial sediment yields observed for forested subalpine areas of western North America. Copyright ?? 2005 John Wiley & Sons, Ltd.
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Wang, Hsueh-Ching; Chou, Chiao-Ying; Chiou, Chyi-Rong; Tian, Guanglong; Chiu, Chih-Yu
2016-01-01
Studying the influence of climatic and/or site-specific factors on soil organic matter (SOM) along an elevation gradient is important for understanding the response of SOM to global warming. We evaluated the composition of SOM and structure of humic acids along an altitudinal gradient from 600 to 1400 m in moso bamboo (Phyllostachys edulis) plantations in central Taiwan using NMR spectroscopy and photometric analysis. Total organic C and total nitrogen (N) content increased with increasing elevation. Aromaticity decreased and ΔlogK (the logarithm of the absorbance ratio of humic acids at 400 and 600 nm) increased with increasing elevation, which suggests that SOM humification decreased with increasing elevation. High temperature at low elevations seemed to enhance the decomposition (less accumulation of total organic C and N) and humification (high aromaticity and low ΔlogK). The alkyl-C/O-alkyl-C (A/O-A) ratio of humic acids increased with increasing elevation, which suggests that SOM humification increased with increasing elevation; this finding was contrary to the trend observed for ΔlogK and aromaticity. Such a discrepancy might be due to the relatively greater remaining of SOM derived from high alkyl-C broadleaf litter of previous forest at high elevations. The ratio of recalcitrant C to total organic C was low at low elevations, possibly because of enhanced decomposition of recalcitrant SOM from the previous broadleaf forest during long-term intensive cultivation and high temperature. Overall, the change in SOM pools and in the rate of humification with elevation was primarily affected by changes in climatic conditions along the elevation gradient in these bamboo plantations. However, when the composition of SOM, as assessed by NMR spectroscopy and photometric analysis was considered, site-specific factors such as residual SOM from previous forest and intensive cultivation history could also have an important effect on the humic acid composition and humification of SOM.
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Wang, Hsueh-Ching; Chou, Chiao-Ying; Chiou, Chyi-Rong; Tian, Guanglong
2016-01-01
Studying the influence of climatic and/or site-specific factors on soil organic matter (SOM) along an elevation gradient is important for understanding the response of SOM to global warming. We evaluated the composition of SOM and structure of humic acids along an altitudinal gradient from 600 to 1400 m in moso bamboo (Phyllostachys edulis) plantations in central Taiwan using NMR spectroscopy and photometric analysis. Total organic C and total nitrogen (N) content increased with increasing elevation. Aromaticity decreased and ΔlogK (the logarithm of the absorbance ratio of humic acids at 400 and 600 nm) increased with increasing elevation, which suggests that SOM humification decreased with increasing elevation. High temperature at low elevations seemed to enhance the decomposition (less accumulation of total organic C and N) and humification (high aromaticity and low ΔlogK). The alkyl-C/O-alkyl-C (A/O-A) ratio of humic acids increased with increasing elevation, which suggests that SOM humification increased with increasing elevation; this finding was contrary to the trend observed for ΔlogK and aromaticity. Such a discrepancy might be due to the relatively greater remaining of SOM derived from high alkyl-C broadleaf litter of previous forest at high elevations. The ratio of recalcitrant C to total organic C was low at low elevations, possibly because of enhanced decomposition of recalcitrant SOM from the previous broadleaf forest during long-term intensive cultivation and high temperature. Overall, the change in SOM pools and in the rate of humification with elevation was primarily affected by changes in climatic conditions along the elevation gradient in these bamboo plantations. However, when the composition of SOM, as assessed by NMR spectroscopy and photometric analysis was considered, site-specific factors such as residual SOM from previous forest and intensive cultivation history could also have an important effect on the humic acid composition and humification of SOM. PMID:27583451
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Mathiasen, Paula; Premoli, Andrea C
2016-06-01
Current climate change affects the competitive ability and reproductive success of many species, leading to local extinctions, adjustment to novel local conditions by phenotypic plasticity or rapid adaptation, or tracking their optima through range shifts. However, many species have limited ability to expand to suitable areas. Altitudinal gradients, with abrupt changes in abiotic conditions over short distances, represent "natural experiments" for the evaluation of ecological and evolutionary responses under scenarios of climate change. Nothofagus pumilio is the tree species which dominates as pure stands the montane forests of Patagonia. We evaluated the adaptive value of variation in quantitative traits of N. pumilio under contrasting conditions of the altitudinal gradient with a long-term reciprocal transplant experimental design. While high-elevation plants show little response in plant, leaf, and phenological traits to the experimental trials, low-elevation ones show greater plasticity in their responses to changing environments, particularly at high elevation. Our results suggest a relatively reduced potential for evolutionary adaptation of high-elevation genotypes, and a greater evolutionary potential of low-elevation ones. Under global warming scenarios of forest upslope migration, high-elevation variants may be outperformed by low-elevation ones during this process, leading to the local extinction and/or replacement of these genotypes. These results challenge previous models and predictions expected under global warming for altitudinal gradients, on which the leading edge is considered to be the upper treeline forests.
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Chapter 6: Creating a basis for watershed management in high elevation forests
Gerald J. Gottfried; Leonard F. DeBano; Peter F. Ffolliott
1999-01-01
Higher mountains and plateaus in the Central Arizona Highlands generally support southwestern mixed conifer forests, associated aspen and spruce-fir forests, and a small acreage of grasslands interspersed among the forested areas. Most of the major rivers in the region originate on headwater watersheds that support mixed conifer forests where annual precipitation,...
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Landscape-moderated bird nest predation in hedges and forest edges
NASA Astrophysics Data System (ADS)
Ludwig, Martin; Schlinkert, Hella; Holzschuh, Andrea; Fischer, Christina; Scherber, Christoph; Trnka, Alfréd; Tscharntke, Teja; Batáry, Péter
2012-11-01
Landscape-scale agricultural intensification has caused severe declines in biodiversity. Hedges and forest remnants may mitigate biodiversity loss by enhancing landscape heterogeneity and providing habitat to a wide range of species, including birds. However, nest predation, the major cause of reproductive failure of birds, has been shown to be higher in forest edges than in forest interiors. Little is known about how spatial arrangement (configuration) of hedges affects the avian nest predation. We performed an experiment with artificial ground and elevated nests (resembling yellowhammer and whitethroat nests) baited with quail and plasticine eggs. Nests were placed in three habitat types with different degrees of isolation from forests: forest edges, hedges connected to forests and hedges isolated from forests. Nest predation was highest in forest edges, lowest in hedges connected to forests and intermediate in isolated hedges. In the early breeding season, we found similar nest predation on ground and elevated nests, but in the late breeding season nest predation was higher on ground nests than on elevated nests. Small mammals were the main predators of ground nests and appeared to be responsible for the increase in predation from early to late breeding season, whereas the elevated nests were mainly depredated by small birds and small mammals. High predation pressure at forest edges was probably caused by both forest and open-landscape predators. The influence of forest predators may be lower at hedges, leading to lower predation pressure than in forest edges. Higher predation pressure in isolated than connected hedges might be an effect of concentration of predators in these isolated habitats. We conclude that landscape configuration of hedges is important in nest predation, with connected hedges allowing higher survival than isolated hedges and forest edges.
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Pringle, C.M.; Triska, F.J.; Browder, G.
1990-01-01
Spatial variability in selected chemical, physical and biological parameters was examined in waters draining relatively pristine tropical forests spanning elevations from 35 to 2600 meters above sea level in a volcanic landscape on Costa Rica's Caribbean slope. Waters were sampled within three different vegetative life zones and two transition zones. Water temperatures ranged from 24-25 ??C in streams draining lower elevations (35-250 m) in tropical wet forest, to 10 ??C in a crater lake at 2600 m in montane forest. Ambient phosphorus levels (60-300 ??g SRP L-1; 66-405 ??g TP L-1) were high at sites within six pristine drainages at elevations between 35-350 m, while other undisturbed streams within and above this range in elevation were low (typically <30.0 ??g SRP L-1). High ambient phosphorus levels within a given stream were not diagnostic of riparian swamp forest. Phosphorus levels (but not nitrate) were highly correlated with conductivity, Cl, Na, Ca, Mg and SO4. Results indicate two major stream types: 1) phosphorus-poor streams characterized by low levels of dissolved solids reflecting local weathering processes; and 2) phosphorus-rich streams characterized by relatively high Cl, SO4, Na, Mg, Ca and other dissolved solids, reflecting dissolution of basaltic rock at distant sources and/or input of volcanic brines. Phosphorus-poor streams were located within the entire elevation range, while phosphorus-rich streams were predominately located at the terminus of Pleistocene lava flows at low elevations. Results indicate that deep groundwater inputs, rich in phosphorus and other dissolved solids, surface from basaltic aquifers at breaks in landform along faults and/or where the foothills of the central mountain range merge with the coastal plain. ?? 1990 Kluwer Academic Publishers.
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Vertical accretion and shallow subsidence in a mangrove forest of southwestern Florida, U.S.A
Cahoon, D.R.; Lynch, J.C.
1997-01-01
Simultaneous measurements of vertical accretion from artificial soil marker horizons and soil elevation change from sedimentation-erosion table (SET) plots were used to evaluate the processes related to soil building in range, basin, and overwash mangrove forests located in a low-energy lagoon which recieves minor inputs of terregenous sediments. Vertical accretion measures reflect the contribution of surficial sedimentation (sediment deposition and surface root growth). Measures of elevation change reflect not only the contributions of vertical accretion but also those of subsurface processes such as compaction, decomposition and shrink-swell. The two measures were used to calculate amounts of shallow subsidence (accretion minus elevation change) in each mangrove forest. The three forest types represent different accretionary envrionments. The basin forest was located behind a natural berm. Hydroperiod here was controlled primarily by rainfall rather than tidal exchange, although the basin flooded during extreme tidal events. Soil accretion here occurred primarily by autochthonous organic matter inputs, and elevation was controlled by accretion and shrink-swell of the substrate apparently related to cycles of flooding-drying and/or root growth-decomposition. This hydrologically-restricted forest did not experience an accretion or elevation deficit relative to sea-level rise. The tidally dominated fringe and overwash island forests accreted through mineral sediment inputs bound in place by plant roots. Filamentous turf algae played an important role in stabilizing loose muds in the fringe forest where erosion was prevalent. Elevation in these high-energy environments was controlled not only by accretion but also by erosion and/or shallow subsidence. The rate of shallow subsidence was consistently 3-4 mm y-1 in the fringe and overwash island forests but was negligible in the basin forest. Hence, the vertical development of mangrove soils was influenced by both surface and subsurface processes and the procces controlling soil elevation differed among forest types. The mangrove ecosystem at Rookery Bay has remained stable as sea level has risen during the past 70 years. Yet, lead-210 accretion data suggest a substantial accretion deficit has occurred in the past century (accretion was 10-20 cm < sea-level rise from 1930 to 1990) in the fringe and island forests at Rookery Bay. In contrast, our measures of elevation change mostly equalled the estimates of sea-level rise and shallow subsidence. These data suggest that (1) vertical accretion in this system is driven by local sea-level rise and shallow subsidence, and (2) the mangrove forests are mostly keeping pace with sea-level rise. Thus, the vulnerability of this mangrove ecosystem to sea-level rise is best described in terms of an elevation deficit (elevation change minus sea-level rise) based on annual measures rather than an accretion deficit (accretion minus sea-level rise) based on decadal measures.
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Forest biomass variation in Southernmost Brazil: the impact of Araucaria trees.
Rosenfield, Milena Fermina; Souza, Alexandre F
2014-03-01
A variety of environmental and biotic factors determine vegetation growth and affect plant biomass accumulation. From temperature to species composition, aboveground biomass storage in forest ecosystems is influenced by a number of variables and usually presents a high spatial variability. With this focus, the aim of the study was to evaluate the variables affecting live aboveground forest biomass (AGB) in Subtropical Moist Forests of Southern Brazil, and to analyze the spatial distribution of biomass estimates. Data from a forest inventory performed in the State of Rio Grande do Sul, Southern Brazil, was used in the present study. Thirty-eight 1-ha plots were sampled and all trees with DBH > or = 9.5cm were included for biomass estimation. Values for aboveground biomass were obtained using published allometric equations. Environmental and biotic variables (elevation, rainfall, temperature, soils, stem density and species diversity) were obtained from the literature or calculated from the dataset. For the total dataset, mean AGB was 195.2 Mg/ha. Estimates differed between Broadleaf and Mixed Coniferous-Broadleaf forests: mean AGB was lower in Broadleaf Forests (AGB(BF)=118.9 Mg/ha) when compared to Mixed Forests (AGB(MF)=250.3 Mg/ha). There was a high spatial and local variability in our dataset, even within forest types. This condition is normal in tropical forests and is usually attributed to the presence of large trees. The explanatory multiple regressions were influenced mainly by elevation and explained 50.7% of the variation in AGB. Stem density, diversity and organic matter also influenced biomass variation. The results from our study showed a positive relationship between aboveground biomass and elevation. Therefore, higher values of AGB are located at higher elevations and subjected to cooler temperatures and wetter climate. There seems to be an important contribution of the coniferous species Araucaria angustifolia in Mixed Forest plots, as it presented significantly higher biomass than angiosperm species. In Brazil, this endangered species is part of a high diversity forest (Araucaria Forest) and has the potential for biomass storage. The results of the present study show the spatial and local variability in aboveground biomass in subtropical forests and highlight the importance of these ecosystems in global carbon stock, stimulating the improvement of future biomass estimates.
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NASA Astrophysics Data System (ADS)
Litvak, M. E.; Krofcheck, D. J.; Maurer, G.
2015-12-01
Semi-arid biomes in many parts of the Southwestern U.S. have experienced a range of precipitation over the last decade, ranging from wetter than average years 2006-2010 (relative to the 40-year PRISM mean), extreme drought years (2010-2011) and slightly dry-average precipitation years (2013-2015). While annual carbon uptake in semi-arid biomes of the Southwestern US is relatively low, compared to more temperate ecosystems, collectively these biomes store a significant amount of carbon on a regional scale. It is therefore of great interest to understand what impact this range in precipitation variability has on inter- and intra- annual variability in regional carbon dynamics. We use an 9 year record from 2007-2015 of continuous measurements of net ecosystem exchange of carbon (NEE) and its components (gross primary productivity (GPP) and ecosystem respiration (Re), made across a network of flux towers along an elevation/aridity gradient in New Mexico, the New Mexico Elevation Gradient (NMEG), to quantify biome-specific responses of carbon dynamics to climate variability over this time period. Biomes include a desert grassland, creosote shrubland, juniper savanna, piñon-juniper woodland, and ponderosa pine and subalpine mixed conifer forests. We compared daily, seasonal and annual NEP, GPP and Re means between pre-drought (2007-2010), drought (2011-2012), and post-drought years (2013-2015). All biomes sequestered less carbon in the drought years, compared to the pre-drought years (~30-40, 270 and 60 g C/m2 less in low and middle elevation biomes, ponderosa pine, and mixed conifer forest, respectively), as GPP in all biomes was more sensitive to the drought than Re. In the post-drought years, GPP was still only 80-90% what it was in the pre-drought years. Re, however, in all biomes except for the creosote shrubland, was 5-15% higher in the post-drought years compared to pre-drought. As a result, carbon sequestration in these biomes was 20-75% lower in the post-drought years compared to what we observed before the drought, suggesting, even 3 years post-drought, none of these biomes has recovered from the extreme drought. The large implications of this lack of resilience for the carbon sink strength in this region are discussed.
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Effects of Resource Chemistry on the Composition and Function of Stream Hyporheic Biofilms
Hall, E. K.; Besemer, K.; Kohl, L.; Preiler, C.; Riedel, K.; Schneider, T.; Wanek, W.; Battin, T. J.
2012-01-01
Fluvial ecosystems process large quantities of dissolved organic matter as it moves from the headwater streams to the sea. In particular, hyporheic sediments are centers of high biogeochemical reactivity due to their elevated residence time and high microbial biomass and activity. However, the interaction between organic matter and microbial dynamics in the hyporheic zone remains poorly understood. We evaluated how variance in resource chemistry affected the microbial community and its associated activity in experimentally grown hyporheic biofilms. To do this we fed beech leaf leachates that differed in chemical composition to a series of bioreactors filled with sediment from a sub-alpine stream. Differences in resource chemistry resulted in differences in diversity and phylogenetic origin of microbial proteins, enzyme activity, and microbial biomass stoichiometry. Specifically, increased lignin, phenolics, and manganese in a single leachate resulted in increased phenoloxidase and peroxidase activity, elevated microbial biomass carbon:nitrogen ratio, and a greater proportion of proteins of Betaproteobacteria origin. We used this model system to attempt to link microbial form (community composition and metaproteome) with function (enzyme activity) in order to better understand the mechanisms that link resource heterogeneity to ecosystem function in stream ecosystems. PMID:22347877
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Effects of resource chemistry on the composition and function of stream hyporheic biofilms.
Hall, E.K.; Besemer, K.; Kohl, L.; Preiler, C.; Reidel, K.; Schneider, T.; Wanek, W.; Battin, T.J.
2012-01-01
Fluvial ecosystems process large quantities of dissolved organic matter as it moves from the headwater streams to the sea. In particular, hyporheic sediments are centers of high biogeochemical reactivity due to their elevated residence time and high microbial biomass and activity. However, the interaction between organic matter and microbial dynamics in the hyporheic zone remains poorly understood. We evaluated how variance in resource chemistry affected the microbial community and its associated activity in experimentally grown hyporheic biofilms. To do this we fed beech leaf leachates that differed in chemical composition to a series of bioreactors filled with sediment from a sub-alpine stream. Differences in resource chemistry resulted in differences in diversity and phylogenetic origin of microbial proteins, enzyme activity, and microbial biomass stoichiometry. Specifically, increased lignin, phenolics, and manganese in a single leachate resulted in increased phenoloxidase and peroxidase activity, elevated microbial biomass carbon:nitrogen ratio, and a greater proportion of proteins of Betaproteobacteria origin. We used this model system to attempt to link microbial form (community composition and metaproteome) with function (enzyme activity) in order to better understand the mechanisms that link resource heterogeneity to ecosystem function in stream ecosystems.
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Drought effects on evapotransiration and subsurface water storage in the southern Sierra Nevada
NASA Astrophysics Data System (ADS)
Bales, R. C.; Goulden, M.; Hunsaker, C. T.; Conklin, M. H.; Hartsough, P. C.; O'Geen, T. T.; Hopmans, J. W.; Safeeq, M.
2015-12-01
Multi-year measurements of evapotranspiration (ET) at three elevations in the southern Sierra Nevada show the extent to which subsurface water storage in the regolith provides a buffer against multi-year dry periods. ET in a 2000-m elevation mixed-conifer forest showed a 24% decrease in ET in water-year 2014, the third dry year, as compared to the wet year of 2011. This decrease reflected reduced transpiration for the July to September period. Over half of the annual ET in both wet and dry years came from below the 1-m depth mapped soil, and with come coming from below the 2.5 m depth of our soil-moisture measurements. The ability of trees to access water from these depths does provide a 2-3 year buffer for ET, which also depends on forest density and the balance between perennial overstory and annual understory vegetation. An equally dense lower-elevation pine-oak forest (1160 m) showed nearly a 50% decrease in ET during the third year of drought, with significant visible effects on vegetation. While this lower elevation forest may have as much or more subsurface storage as does that at 2000-m elevation, the combination of lower precipitation as one goes down in elevation and very high forest density provides only a one-year buffer for ET in dry years. Regaining resiliency in this forest will only occur with significant reductions in biomass and commensurate lowering of ET. In a 400-m elevation oak savannah ET responds to annual precipitation, with essentially no multi-year buffer provided by subsurface storage.
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NASA Astrophysics Data System (ADS)
Rodenhouse, N.; Gebauer, R.; Lowe, W.; McFarland, K.; Bank, M. S.
2015-12-01
The soils and foods webs associated with mid to high elevation, forested, headwater streams are potential hotspots for mercury methylation and bioaccumulation but are not well studied. We tested the hypothesis that spatial variation in mercury bioaccumulation in upland taxa associated with headwater streams can be explained by variation in soil conditions promoting Hg methylation such as soil moisture, pH, and sulfur and organic matter content. We sampled at high (c. 700m) and mid elevation (c. 500m) in northern hardwood forest adjacent to and away from (75m) replicate headwater streams in the Hubbard Brook and Jeffers Brook watersheds of the White Mountains region, New Hampshire, USA. These forested watersheds differed primarily in soil calcium content and pH. We measured and assessed spatial variation in total Hg (THg) and methyl Hg (MeHg) concentrations in soils, insects, spiders, salamanders and birds. We also tested whether trophic position, as determined by nitrogen stable isotopes, was a major predictor of Hg bioaccumulation across these riparian and upland forest taxa. We found elevated levels of THg in all measured components of the food web, and conditions for methylation were better in the upland forest sites compared to the riparian sites located adjacent to headwater streams. Both THg and MeHg in biota were positively correlated with trophic position as indicated by 15N enrichment. In fact, trophic position was a better predictor of THg and MeHg content than spatial location, but the spatial patterning of bioaccumulation differed among taxa. Our data show that that significant Hg bioaccumulation and biomagnification can occur in soils and food webs of mid to high elevation temperate deciduous forests of the Northeast. They also suggest that mercury methylation in forested watersheds is a widespread phenomenon and not limited to areas with high soil moisture, such as lotic environments.
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Johnny L. Boggs; Steven G. McNulty; Michael J. Gavazzi; Jennifer Moore Myers
2005-01-01
The declining health of high-elevation red spruce (Picea rubens Sarg.) and Fraser fir (Abies fraseri (Pursh) Poir.) in the southern Appalachian region has long been linked to nitrogen (N)deposition. Recently, N deposition has also been proposed as a source of negative health impacts in lower elevation deciduous forests. In 1998 we...
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Zech, Michael; Bimüller, Carolin; Hemp, Andreas; Samimi, Cyrus; Broesike, Christina; Hörold, Claudia; Zech, Wolfgang
2011-09-01
Population pressure increasingly endangers high-mountain ecosystems such as the pastures in the Eastern Pamirs and the mountain forests on Mt. Kilimanjaro. At the same time, these ecosystems constitute the economic basis for millions of people living there. In our study, we, therefore, aimed at characterising the land-use effects on soil degradation and N-cycling by determining the natural abundance of (15)N. A short review displays that δ(15)N of plant-soil systems may often serve as an integrated indicator of N-cycles with more positive δ(15)N values pointing towards N-losses. Results for the high-mountain pastures in the Eastern Pamirs show that intensively grazed pastures are significantly enriched in (15)N compared to the less-exploited pastures by 3.5 ‰, on average. This can be attributed to soil organic matter degradation, volatile nitrogen losses, nitrogen leaching and a general opening of the N-cycle. Similarly, the intensively degraded savanna soils, the cultivated soils and the soils under disturbed forests on the foothill of Mt. Kilimanjaro reveal very positive δ(15)N values around 6.5 ‰. In contrast, the undisturbed forest soils in the montane zone are more depleted in (15)N, indicating that here the N-cycle is relatively closed. However, significantly higher δ(15)N values characterise the upper montane forest zone at the transition to the subalpine zone. We suggest that this reflects N-losses by the recently monitored and climate change and antropogenically induced increasing fire frequency pushing the upper montane rainforest boundary rapidly downhill. Overall, we conclude that the analysis of the (15)N natural abundance in high-mountain ecosystems is a purposeful tool for detecting land-use- or climate change-induced soil degradation and N-cycle opening.
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Thermal ecology of montane Atelopus (Anura: Bufonidae): A study of intrageneric diversity.
Rueda Solano, Luis Alberto; Navas, Carlos A; Carvajalino-Fernández, Juan Manuel; Amézquita, Adolfo
2016-05-01
Harlequin frogs (Bufonidae: Atelopus) are among the most threatened frog genus in the world and reach very high elevations in the tropical Andes and the Sierra Nevada de Santa Marta (SNSM). Learning about their thermal ecology is essential to infer sensitivity to environmental changes, particularly climate warming. We report on the activity temperature and thermoregulatory behavior of three high-elevation species of harlequin frogs, Atelopus nahumae, Atelopus laetissimus and Atelopus carrikeri. The first two mentioned live in streams in Andean rain forests, whereas A. carrikeri inhabits paramo streams in the SNSM. We studied the thermal ecology of these species in tree localities differing in altitude, and focused on activity body, operative, substrate and air temperature. A main trend was lower body temperature as elevation increased, so that differences among species were largely explained by differences in substrate temperature. However, this temperature variation was much lower in forest species than paramo species. The Atelopus species included in this work proved to be thermoconformers, a trend that not extended to all congenerics at high elevation. This diversity in thermal ecology poses important questions when discussing the impact of climate warming for high-elevation harlequin frogs. For example, forest species show narrow thermal ranges and, if highly specialized, may be more susceptible to temperature change. Paramo species such as A. carrikeri, in contrast, may be more resilient to temperature change. Copyright © 2016 Elsevier Ltd. All rights reserved.
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Evaluation of storage and filtration protocols for alpine/subalpine lake water quality samples
John L. Korfmacher; Robert C. Musselman
2007-01-01
Many government agencies and other organizations sample natural alpine and subalpine surface waters using varying protocols for sample storage and filtration. Simplification of protocols would be beneficial if it could be shown that sample quality is unaffected. In this study, samples collected from low ionic strength waters in alpine and subalpine lake inlets...
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Landscape and plant physiological controls on water dynamics within a watershed
NASA Astrophysics Data System (ADS)
Hu, J.; Looker, N. T.; Martin, J. T.; Hoylman, Z. H.; Jencso, K. G.
2014-12-01
Across the Western U.S., declining snowpacks have resulted in increased water limitation, leading to reduced productivity in high elevation forests. While our current understanding of how forests respond to climate change is typically focused on measuring/modeling the physiological responses and climate feedbacks, our study aims to combine physiology with hydrology to examine how landscape topography modulates the sensitivity of forests to climate. In a forested watershed in Western Montana, we linked climate variability to the physical watershed characteristics and the physiological response of vegetation to examine forest transpiration and productivity rates. Across the entire watershed, we found a strong relationship between productivity and the topographic wetness index, a proxy for soil moisture storage. However, this relationship was highly dependent on the intensity of solar radiation, suggesting that at high elevations productivity was limited by temperature, while at low elevations productivity was limited by moisture. In order to identify the mechanisms responsible for this relationship, we then examined how different coniferous species respond to changing environmental and hydrologic regimes. We first examined transpiration and productivity rates at the hillslope scale at four plots, ranging in elevation and aspect across the watershed. We found trees growing in the hollows had higher transpiration and productivity rates than trees growing in the side slope, but that these differences were more pronounced at lower elevations. We then used oxygen isotope to examine water source use by different species across the watershed. We found that trees growing in the hollows used snowmelt for a longer period. This was most likely due to upslope subsidies of snowmelt water to the hollow areas. However, we found that trees growing at lower elevations used proportionally more snowmelt than trees at the higher elevations. This was most likely due to the trees at lower elevation depending on deeper, more reliable water when the upper soils dried down during midsummer. These observations suggest that landscape topography influences the availability of soil water, which influences tree transpiration and productivity rates, thereby leading to watershed patterns of productivity.
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Sharon M. Hood; Stephen Baker; Anna Sala
2016-01-01
Fire frequency in low-elevation coniferous forests in western North America has greatly declined since the late 1800s. In many areas, this has increased tree density and the proportion of shade-tolerant species, reduced resource availability, and increased forest susceptibility to forest insect pests and high-severity wildfire. In response, treatments are...
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Simulations of snow distribution and hydrology in a mountain basin
Hartman, Melannie D.; Baron, Jill S.; Lammers, Richard B.; Cline, Donald W.; Band, Larry E.; Liston, Glen E.; Tague, Christina L.
1999-01-01
We applied a version of the Regional Hydro-Ecologic Simulation System (RHESSys) that implements snow redistribution, elevation partitioning, and wind-driven sublimation to Loch Vale Watershed (LVWS), an alpine-subalpine Rocky Mountain catchment where snow accumulation and ablation dominate the hydrologic cycle. We compared simulated discharge to measured discharge and the simulated snow distribution to photogrammetrically rectified aerial (remotely sensed) images. Snow redistribution was governed by a topographic similarity index. We subdivided each hillslope into elevation bands that had homogeneous climate extrapolated from observed climate. We created a distributed wind speed field that was used in conjunction with daily measured wind speeds to estimate sublimation. Modeling snow redistribution was critical to estimating the timing and magnitude of discharge. Incorporating elevation partitioning improved estimated timing of discharge but did not improve patterns of snow cover since wind was the dominant controller of areal snow patterns. Simulating wind-driven sublimation was necessary to predict moisture losses.
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Detlev R. Vogler; Annette Delfino-Mix; Anna W. Schoettle
2006-01-01
Recent concern about survival and recovery of high-elevation white pine ecosystems has returned white pine blister rust (caused by Cronartiurn ribicola) to prominence as a significant threat to forest health in the western U.S. (Sainman et al., 2003). This, in turn, has spurred new research into potential rust-resistance mechanisms in high-elevation...
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Paxton, Eben H.; Burgett, Jeff; McDonald-Fadden, Eve; Bean, Ellen; Atkinson, Carter T.; Ball, Donna; Cole, Colleen; Crampton, Lisa H.; Kraus, Jim; LaPointe, Dennis A.; Mehrhoff, Loyal; Samuel, Michael D.; Brewer, Donna; Converse, Sarah J.; Morey, Steve
2011-01-01
This report is a product of a one-week workshop on using Structured Decision Making to identify and prioritize conservation actions to address the threat of climate change on Hawaii‟s native forest bird community. Specifically, t his report addresses the issue of global warming ‟s likely role in increasing disease prevalence in upper elevation forests of Hawaii, negatively impacting native bird populations susceptible to the disease but currently disease - free because of the cooler temperatures at high elevations.
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Seasonal Snow Cold Content Dynamics in the Alpine and Sub-Alpine, Niwot Ridge, Colorado, USA
NASA Astrophysics Data System (ADS)
Jennings, K. S.; Molotch, N. P.
2015-12-01
Cold content represents the energy required to warm a sub-freezing snowpack to an isothermal 0°C. Across daily and seasonal time scales it is a dynamic interplay between the forces of snowpack accumulation/cooling and warming. Cold content determines snowmelt timing and is an important component of the annual energy budget of mountain sites with seasonal snowpacks. However, little is understood about seasonal snowpack cold content dynamics as calculating cold content requires depth-weighted snowpack temperature and snow water equivalent (SWE) measurements, which are scarce. A spatially distributed network of snow pits has been sampled since 1993 at the Niwot Ridge Long Term Ecological Research site on the eastern slope of the Continental Divide in Colorado's Front Range mountains. This study uses data from 3 pit sites that have at least 8 years of observations and represent alpine and sub-alpine environments. For these pits, cold content is strongly related to SWE during the cold content accumulation phase, here defined as December, January, and February. Average peak cold content ranges between -2.5 MJ m-2 and -9.2 MJ m-2 for the three sites and is strongly related to peak SWE. On average, cold content reaches its maximum on February 26, which is 61 days before the average date of peak SWE (i.e., the snowpack's cold content is satisfied over an average of 61 days). At the alpine site, later peak cold content and SWE was observed relative to the lower elevation sub-alpine sites. Interestingly, the alpine site had a smaller gap between peak cold content and SWE (55 days versus 67 days for the alpine and sub-alpine sites, respectively). The gap between peak cold content and peak SWE is primarily a function of the increase in SWE between the two dates. Hence, persistent snowfall after the date of peak cold content can delay the onset of snowmelt even if peak cold content was relatively low. Improving our understanding of seasonal cold content dynamics in mountain environments will enable us to better model the future effects of climate change on snowmelt timing and associated hydrologic response.
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Spruce-fir forest changes during a 30-year nitrogen saturation experiment
Steven G. McNulty; Johnny L. Boggs; John D. Aber; Lindsey E. Rustad
2017-01-01
A field experiment was established in a high elevation red spruce (Picea rubens Sarg.) â balsam fir (Abies balsamea) forest on Mount Ascutney Vermont, USA in 1988 to test the nitrogen (N) saturation hypothesis, and to better understand the mechanisms causing forest decline at the time. The study established replicate control, lowand high dose nitrogen addition plots (i...
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DOE Office of Scientific and Technical Information (OSTI.GOV)
Woodward, A.; Silsbee, D.G.; Schreiner, E.G.
1995-08-01
Noticeable changes in vegetation distribution have occurred in the Pacific Northwest during the last century as trees have established in some subalpine meadows. To study the relationship of this process to climate, recently established trees were aged in six subalpine meadows in the Olympic Mountains, Washington. The sites represent three points along a steep precipitation gradient. Subalpine fir (Abies lasiocarpa) has been establishing at the dry end of the gradient, mountain hemlock (Tsuga mertensiana) at the wet end, and both species in the center. Establishment patterns were compared with deviations from the century-long average for these weather variables: winter precipitation,more » Palmer Drought Severity Index, and winter, October and May temperatures. Results show that establishment occurred in dry areas when weather conditions were wetter than average, and in wet areas under drier than average conditions. Establishment at central sites did not show consistent relationships with climate. If future climatic conditions continue to warm, establishment of subalpine fir in subalpine meadows in dry areas may cease and mountain hemlock may resume in wet areas. 34 refs., 5 figs., 3 tabs.« less
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Water relations and microclimate around the upper limit of a cloud forest in Maui, Hawai'i.
Gotsch, Sybil G; Crausbay, Shelley D; Giambelluca, Thomas W; Weintraub, Alexis E; Longman, Ryan J; Asbjornsen, Heidi; Hotchkiss, Sara C; Dawson, Todd E
2014-07-01
The goal of this study was to determine the effects of atmospheric demand on both plant water relations and daily whole-tree water balance across the upper limit of a cloud forest at the mean base height of the trade wind inversion in the tropical trade wind belt. We measured the microclimate and water relations (sap flow, water potential, stomatal conductance, pressure-volume relations) of Metrosideros polymorpha Gaudich. var. polymorpha in three habitats bracketing the cloud forest's upper limit in Hawai'i to understand the role of water relations in determining ecotone position. The subalpine shrubland site, located 100 m above the cloud forest boundary, had the highest vapor pressure deficit, the least amount of rainfall and the highest levels of nighttime transpiration (EN) of all three sites. In the shrubland site, on average, 29% of daily whole-tree transpiration occurred at night, while on the driest day of the study 50% of total daily transpiration occurred at night. While EN occurred in the cloud forest habitat, the proportion of total daily transpiration that occurred at night was much lower (4%). The average leaf water potential (Ψleaf) was above the water potential at the turgor loss point (ΨTLP) on both sides of the ecotone due to strong stomatal regulation. While stomatal closure maintained a high Ψleaf, the minimum leaf water potential (Ψleafmin) was close to ΨTLP, indicating that drier conditions may cause drought stress in these habitats and may be an important driver of current landscape patterns in stand density. © The Author 2014. Published by Oxford University Press. All rights reserved. For Permissions, please email: journals.permissions@oup.com.
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Banko, Paul C.; Camp, Richard J.; Farmer, Chris; Brinck, Kevin W.; Leonard, David L.; Stephens, Robert M.
2013-01-01
Extinction has claimed half of all historically-known Hawaiian passerines, and today many extant species are increasingly threatened due to the combined effects of invasive species and climate change. Habitat disturbance has affected populations of feeding specialists most profoundly, and our results indicate that specialists continue to be most vulnerable, although even some abundant, introduced, generalist species also may be affected. Surveys of passerines during 1998–2011 in subalpine woodland habitat on Mauna Kea Volcano, Island of Hawai′i, revealed that the abundance of the critically endangered palila (Loxioides bailleui), a seed specialist, declined by 79% after 2003. The ′akiapōlā′au (Hemignathus munroi), an endangered specialist insectivore, was not detected in the survey area after 1998. The Hawai′i ′amakihi (Hemignathus virens virens), a generalist feeder and the most abundant species on Mauna Kea, was the only native species to maintain a stable population. The Japanese white-eye (Zosterops japonicus), a well-entrenched generalist and one of the three most common introduced species, declined. Drought prevailed in 74% of months during 2000–2011, and dry conditions contributed to the recent decline of the palila by reducing the annual māmane (Sophora chrysophylla) seed pod crop, which influences palila breeding and survival. Sustained browsing by introduced ungulates also lowered habitat carrying capacity, and their elimination should reduce the effects of drought and promote forest restoration. Our results illustrate how the feeding ecology of a species can influence its response to interacting environmental perturbations, and they underscore the value of long-term monitoring to detect population trends of sensitive species.
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NASA Astrophysics Data System (ADS)
Burns, Sean P.; Swenson, Sean C.; Wieder, William R.; Lawrence, David M.; Bonan, Gordon B.; Knowles, John F.; Blanken, Peter D.
2018-03-01
Precipitation changes the physiological characteristics of an ecosystem. Because land-surface models are often used to project changes in the hydrological cycle, modeling the effect of precipitation on the latent heat flux λE is an important aspect of land-surface models. Here we contrast conditionally sampled diel composites of the eddy-covariance fluxes from the Niwot Ridge Subalpine Forest AmeriFlux tower with the Community Land Model (CLM, version 4.5). With respect to measured λE during the warm season: for the day following above-average precipitation, λE was enhanced at midday by ≈40 W m-2 (relative to dry conditions), and nocturnal λE increased from ≈10 W m-2 in dry conditions to over 20 W m-2 in wet conditions. With default settings, CLM4.5 did not successfully model these changes. By increasing the amount of time that rainwater was retained by the canopy/needles, CLM was able to match the observed midday increase in λE on a dry day following a wet day. Stable nighttime conditions were problematic for CLM4.5. Nocturnal CLM λE had only a small (≈3 W m-2) increase during wet conditions, CLM nocturnal friction velocity u∗ was smaller than observed u∗, and CLM canopy air temperature was 2°C less than those measured at the site. Using observed u∗ as input to CLM increased λE; however, this caused CLM λE to be increased during both wet and dry periods. We suggest that sloped topography and the ever-present drainage flow enhanced nocturnal u∗ and λE. Such phenomena would not be properly captured by topographically blind land-surface models, such as CLM.
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Emissions pathways, climate change, and impacts on California
Hayhoe, Katharine; Cayan, Daniel; Field, Christopher B.; Frumhoff, Peter C.; Maurer, Edwin P.; Miller, Norman L.; Moser, Susanne C.; Schneider, Stephen H.; Cahill, Kimberly Nicholas; Cleland, Elsa E.; Dale, Larry; Drapek, Ray; Hanemann, R. Michael; Kalkstein, Laurence S.; Lenihan, James; Lunch, Claire K.; Neilson, Ronald P.; Sheridan, Scott C.; Verville, Julia H.
2004-01-01
The magnitude of future climate change depends substantially on the greenhouse gas emission pathways we choose. Here we explore the implications of the highest and lowest Intergovernmental Panel on Climate Change emissions pathways for climate change and associated impacts in California. Based on climate projections from two state-of-the-art climate models with low and medium sensitivity (Parallel Climate Model and Hadley Centre Climate Model, version 3, respectively), we find that annual temperature increases nearly double from the lower B1 to the higher A1fi emissions scenario before 2100. Three of four simulations also show greater increases in summer temperatures as compared with winter. Extreme heat and the associated impacts on a range of temperature-sensitive sectors are substantially greater under the higher emissions scenario, with some interscenario differences apparent before midcentury. By the end of the century under the B1 scenario, heatwaves and extreme heat in Los Angeles quadruple in frequency while heat-related mortality increases two to three times; alpine/subalpine forests are reduced by 50–75%; and Sierra snowpack is reduced 30–70%. Under A1fi, heatwaves in Los Angeles are six to eight times more frequent, with heat-related excess mortality increasing five to seven times; alpine/subalpine forests are reduced by 75–90%; and snowpack declines 73–90%, with cascading impacts on runoff and streamflow that, combined with projected modest declines in winter precipitation, could fundamentally disrupt California's water rights system. Although interscenario differences in climate impacts and costs of adaptation emerge mainly in the second half of the century, they are strongly dependent on emissions from preceding decades. PMID:15314227
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Albert, Loren P; Keenan, Trevor F; Burns, Sean P; Huxman, Travis E; Monson, Russell K
2017-05-01
Eddy covariance (EC) datasets have provided insight into climate determinants of net ecosystem productivity (NEP) and evapotranspiration (ET) in natural ecosystems for decades, but most EC studies were published in serial fashion such that one study's result became the following study's hypothesis. This approach reflects the hypothetico-deductive process by focusing on previously derived hypotheses. A synthesis of this type of sequential inference reiterates subjective biases and may amplify past assumptions about the role, and relative importance, of controls over ecosystem metabolism. Long-term EC datasets facilitate an alternative approach to synthesis: the use of inductive data-based analyses to re-examine past deductive studies of the same ecosystem. Here we examined the seasonal climate determinants of NEP and ET by analyzing a 15-year EC time-series from a subalpine forest using an ensemble of Artificial Neural Networks (ANNs) at the half-day (daytime/nighttime) time-step. We extracted relative rankings of climate drivers and driver-response relationships directly from the dataset with minimal a priori assumptions. The ANN analysis revealed temperature variables as primary climate drivers of NEP and daytime ET, when all seasons are considered, consistent with the assembly of past studies. New relations uncovered by the ANN approach include the role of soil moisture in driving daytime NEP during the snowmelt period, the nonlinear response of NEP to temperature across seasons, and the low relevance of summer rainfall for NEP or ET at the same daytime/nighttime time step. These new results offer a more complete perspective of climate-ecosystem interactions at this site than traditional deductive analyses alone.
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The vulnerability of Indo-Pacific mangrove forests to sea-level rise
Lovelock, Catherine E.; Cahoon, Donald R.; Friess, Daniel A.; Guntenspergen, Glenn R.; Krauss, Ken W.; Reef, Ruth; Rogers, Kerrylee; Saunders, Megan L.; Sidik, Frida; Swales, Andrew; Saintilan, Neil; Thuyen, Le Xuan; Triet, Tran
2015-01-01
Sea-level rise can threaten the long-term sustainability of coastal communities and valuable ecosystems such as coral reefs, salt marshes and mangroves. Mangrove forests have the capacity to keep pace with sea-level rise and to avoid inundation through vertical accretion of sediments, which allows them to maintain wetland soil elevations suitable for plant growth. The Indo-Pacific region holds most of the world’s mangrove forests, but sediment delivery in this region is declining, owing to anthropogenic activities such as damming of rivers. This decline is of particular concern because the Indo-Pacific region is expected to have variable, but high, rates of future sea-level rise. Here we analyse recent trends in mangrove surface elevation changes across the Indo-Pacific region using data from a network of surface elevation table instruments. We find that sediment availability can enable mangrove forests to maintain rates of soil-surface elevation gain that match or exceed that of sea-level rise, but for 69 per cent of our study sites the current rate of sea-level rise exceeded the soil surface elevation gain. We also present a model based on our field data, which suggests that mangrove forests at sites with low tidal range and low sediment supply could be submerged as early as 2070.
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The vulnerability of Indo-Pacific mangrove forests to sea-level rise.
Lovelock, Catherine E; Cahoon, Donald R; Friess, Daniel A; Guntenspergen, Glenn R; Krauss, Ken W; Reef, Ruth; Rogers, Kerrylee; Saunders, Megan L; Sidik, Frida; Swales, Andrew; Saintilan, Neil; Thuyen, Le Xuan; Triet, Tran
2015-10-22
Sea-level rise can threaten the long-term sustainability of coastal communities and valuable ecosystems such as coral reefs, salt marshes and mangroves. Mangrove forests have the capacity to keep pace with sea-level rise and to avoid inundation through vertical accretion of sediments, which allows them to maintain wetland soil elevations suitable for plant growth. The Indo-Pacific region holds most of the world's mangrove forests, but sediment delivery in this region is declining, owing to anthropogenic activities such as damming of rivers. This decline is of particular concern because the Indo-Pacific region is expected to have variable, but high, rates of future sea-level rise. Here we analyse recent trends in mangrove surface elevation changes across the Indo-Pacific region using data from a network of surface elevation table instruments. We find that sediment availability can enable mangrove forests to maintain rates of soil-surface elevation gain that match or exceed that of sea-level rise, but for 69 per cent of our study sites the current rate of sea-level rise exceeded the soil surface elevation gain. We also present a model based on our field data, which suggests that mangrove forests at sites with low tidal range and low sediment supply could be submerged as early as 2070.
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Changes versus homeostasis in alpine and sub-alpine vegetation over three decades in the sub-arctic.
Hedenås, Henrik; Carlsson, Bengt A; Emanuelsson, Urban; Headley, Alistair D; Jonasson, Christer; Svensson, Brita M; Callaghan, Terry V
2012-01-01
Plant species distributions are expected to shift and diversity is expected to decline as a result of global climate change, particularly in the Arctic where climate warming is amplified. We have recorded the changes in richness and abundance of vascular plants at Abisko, sub-Arctic Sweden, by re-sampling five studies consisting of seven datasets; one in the mountain birch forest and six at open sites. The oldest study was initiated in 1977-1979 and the latest in 1992. Total species number increased at all sites except for the birch forest site where richness decreased. We found no general pattern in how composition of vascular plants has changed over time. Three species, Calamagrostis lapponica, Carex vaginata and Salix reticulata, showed an overall increase in cover/frequency, while two Equisetum taxa decreased. Instead, we showed that the magnitude and direction of changes in species richness and composition differ among sites.
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Soils characterisation along ecological forest zones in the Eastern Himalayas
NASA Astrophysics Data System (ADS)
Simon, Alois; Dhendup, Kuenzang; Bahadur Rai, Prem; Gratzer, Georg
2017-04-01
Elevational gradients are commonly used to characterise vegetation patterns and, to a lesser extent, also to describe soil development. Furthermore, interactions between vegetation cover and soil characteristics are repeatedly observed. Combining information on soil development and easily to distinguish forest zones along elevational gradients, creates an added value for forest management decisions especially in less studied mountain regions. For this purpose, soil profiles along elevational gradients in the temperate conifer forests of Western and Central Bhutan, ranging from 2600-4000m asl were investigated. Thereby, 82 soil profiles were recorded and classified according to the World Reference Base for Soil Resources. Based on 19 representative profiles, genetic horizons were sampled and analysed. We aim to provide fundamental information on forest soil characteristics along these elevational transects. The results are presented with regard to ecological forest zones. The elevational distribution of the reference soil groups showed distinct distribution ranges for most of the soils. Cambisols were the most frequently recorded reference soil group with 58% of the sampled profiles, followed by Podzols in higher elevations, and Stagnosols, at intermediate elevations. Fluvisols occurred only at the lower end of the elevational transects and Phaeozems only at drier site conditions in the cool conifer dry forest zone. The humus layer thickness differs between forest zones and show a shift towards increased organic layer (O-layer) with increasing elevation. The reduced biomass productivity with increasing elevation and subsequently lower litter input compensates for the slow decomposition rates. The increasing O-layer thickness is an indicator of restrained intermixing of organic and mineral components by soil organisms at higher elevation. Overall, the soil types and soil characteristics along the elevational gradient showed a continuous and consistent change, instead of abrupt changes. We interpret these as manifestations of changes of temperature and precipitation with elevation which also drives forest zonation in these least anthropogenically influenced forest ecosystems. The elevational distribution of forest zones is correlated with the distribution of soil types and thus also reflects soil characteristics.
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NASA Astrophysics Data System (ADS)
Liu, B.; Wang, Y.; Zhu, H.; Liang, E.; Camarero, J. J.
2016-10-01
The Tibetan Plateau holds some of the world's highest undisturbed natural treelines and timberlines. Such extreme environments constitute potentially valuable monitoring sites of the effects of climate warming on high-elevation forests. Here, we analyze a network of 21 Smith fir forests situated in the Sygera Mountains, southeastern Tibetan Plateau, using tree-ring width (TRW) and basal area increment (BAI) chronologies. Sampled sites encompassed a wide elevation gradient, from 3600 to 4400 m, including some treeline sites and diverse aspects and tree ages. In comparison with TRW series, BAI series better capture the long-term warming signal. Previous November and current April and summer temperatures are the dominant climatic factors controlling Smith fir radial growth. The mean inter-series correlations of TRW increased upwards, but the forest limit presented the highest potential to reconstruct past temperature variability. Moreover, the growth responses of young trees were less stable than those of trees older than 100 years. Climate warming is accelerating radial growth of Smith fir forest subjected to mesic conditions. Collectively, these findings confirm that the effects of site elevation and tree age should be considered when quantifying climate-growth relationships. The type of tree-ring data (BAI vs. TRW) is also relevant since BAI indices seem to be a better climatic proxy of low-frequency temperature signals than TRW indices. Therefore, site (e.g., elevation) and tree (e.g., age) features should be considered to properly evaluate the effects of climate warming on growth of high-elevation forests.
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Wet and Occult Ion Deposition To An Elevated Forest Ecosystem In Switzerland
NASA Astrophysics Data System (ADS)
Buetzberger, P.; Burkard, R.; Eugster, W.
Due to much higher ion concentrations in fogwater compared to rainwater, critical deposition levels of nutrients such as sulfate, nitrate or ammonium can be achieved in areas with high fog frequency. From summer 2001 until spring 2002 a measuring campaign of the FINIMSAS project (Fog Interception and Nutrient Inputs to Montane- Subalpine Areas in Switzerland) is being conducted at Laegeren (690m asl) on the Swiss Plateau. Fog frequency was high during our campaign. Similar measurements were carried out in 1986/87 at the exact same location, providing a reference data set for comparison. Because the deposition flux was measured differently during 1986/87, direct compar- isons are only possible for ion concentrations. Preliminary results show a significant decrease of sulfate and ammonium median concentrations of more than 50 % over this 15 year period, whereas nitrate decrease is relatively small. This corresponds well with the large-scale evolution of the air pollutant emissions of SO2 (major decrease) and NOx (relatively small decrease). The strong reduction of ammonium is probably due to the reduced use of fertilizer in the area. Chloride shows the largest decrease which can be attributed to the improvement of filtering technique of waste incinerations. In order to achieve maximum comparability, similar event types (e.g. advection fog vs. radiation fog) with similar meteorological conditions were interpreted individually. Analysis of fog nutrient input with respect to wind direction, wind speed, and origin of air mass will help to understand the influence of local and large-scale emissions on fog water concentrations in Switzerland. Computations based on half-hourly mean wind direction revealed significantly lower fog water input but higher median concentra- tions of all measured components if the dominating wind sector was East. Event-based wind field analyses were also carried out and compared to computed trajectories. In order to assess the influence of fog and rain water nutrient deposition on vegetation and soil, we measured throughfall precipitation close to the forest floor. Whereas fog water showed pH values as low as 3, throughfall water was between pH 6 and 7, indicating an important buffering capacity of this ecosystem mainly due to potassium leaching and probably calcium compounds. High ionic concentrations and low pH values seem to act mostly on the leaves.
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Clow, David W.; Sueker, Julie K.
2000-01-01
Relations between stream water chemistry and topographic, vegetative, and geologic characteristics of basins were evaluated for nine alpine/subalpine basins in Rocky Mountain National Park, Colorado, to identify controlling parameters and to better understand processes governing patterns in stream water chemistry. Fractional amounts of steep slopes (≥30°), unvegetated terrain, and young surficial debris within each basin were positively correlated to each other. These terrain features, which commonly occur on steep valley side slopes underlain by talus, were negatively correlated with concentrations of base cations, silica, and alkalinity and were positively correlated with nitrate, acidity, and runoff. These relations might result from the short residence times of water and limited soil development in the talus environment, which limit chemical weathering and nitrogen uptake. Steep, unvegetated terrains also tend to promote high Ca/Na ratios in stream water, probably because physical weathering rates in those areas are high. Physical weathering exposes fresh bedrock that contains interstitial calcite, which weathers relatively quickly. The fractional amounts of subalpine meadow and, to a lesser extent, old surficial debris in the basins were positively correlated to concentrations of weathering products and were negatively correlated to nitrate and acidity. These relations may reflect more opportunities for silicate weathering and nitrogen uptake in the lower‐energy environments of the valley floor, where soils are finer‐grained, older, and better developed and slopes are relatively flat. These results indicate that in alpine/subalpine basins, slope, vegetation (or lack thereof), and distribution and age of surficial materials are interrelated and can have major effects on stream water chemistry.
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Briner, Simon; Elkin, Ché; Huber, Robert
2013-11-15
Provisioning of ecosystem services (ES) in mountainous regions is predicted to be influenced by i) the direct biophysical impacts of climate change, ii) climate mediated land use change, and iii) socioeconomic driven changes in land use. The relative importance and the spatial distribution of these factors on forest and agricultural derived ES, however, is unclear, making the implementation of ES management schemes difficult. Using an integrated economic-ecological modeling framework, we evaluated the impact of these driving forces on the provision of forest and agricultural ES in a mountain region of southern Switzerland. Results imply that forest ES will be strongly influenced by the direct impact of climate change, but that changes in land use will have a comparatively small impact. The simulation of direct impacts of climate change affects forest ES at all elevations, while land use changes can only be found at high elevations. In contrast, changes to agricultural ES were found to be primarily due to shifts in economic conditions that alter land use and land management. The direct influence of climate change on agriculture is only predicted to be substantial at high elevations, while socioeconomic driven shifts in land use are projected to affect agricultural ES at all elevations. Our simulation results suggest that policy schemes designed to mitigate the negative impact of climate change on forests should focus on suitable adaptive management plans, accelerating adaptation processes for currently forested areas. To maintain provision of agricultural ES policy needs to focus on economic conditions rather than on supporting adaptation to new climate. Copyright © 2013 Elsevier Ltd. All rights reserved.
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Phosphorus translocation by red deer on a subalpine grassland in the central European Alps
Martin Schutz; Anita C. Risch; Gerald Achermann; Conny Thiel-Egenter; Deborah Page-Dumroese; Martin F. Jurgensen; Peter J. Edward
2006-01-01
We examined the role of red deer (Cervus elaphus L.) in translocating phosphorus (P) from their preferred grazing sites (short-grass vegetation on subalpine grasslands) to their wider home range in a subalpine grassland ecosystem in the Central European Alps. Phosphorus was used because it is the limiting nutrient in these grasslands. When we compared P removal of...
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Sosa, Victoria; Ornelas, Juan Francisco; Ramírez-Barahona, Santiago; Gándara, Etelvina
2016-01-01
Cloud forests, characterized by a persistent, frequent or seasonal low-level cloud cover and fragmented distribution, are one of the most threatened habitats, especially in the Neotropics. Tree ferns are among the most conspicuous elements in these forests, and ferns are restricted to regions in which minimum temperatures rarely drop below freezing and rainfall is high and evenly distributed around the year. Current phylogeographic data suggest that some of the cloud forest-adapted species remained in situ or expanded to the lowlands during glacial cycles and contracted allopatrically during the interglacials. Although the observed genetic signals of population size changes of cloud forest-adapted species including tree ferns correspond to predicted changes by Pleistocene climate change dynamics, the observed patterns of intraspecific lineage divergence showed temporal incongruence. Here we combined phylogenetic analyses, ancestral area reconstruction, and divergence time estimates with climatic and altitudinal data (environmental space) for phenotypic traits of tree fern species to make inferences about evolutionary processes in deep time. We used phylogenetic Bayesian inference and geographic and altitudinal distribution of tree ferns to investigate ancestral area and elevation and environmental preferences of Mesoamerican tree ferns. The phylogeny was then used to estimate divergence times and ask whether the ancestral area and elevation and environmental shifts were linked to climatic events and historical climatic preferences. Bayesian trees retrieved Cyathea, Alsophyla, Gymnosphaera and Sphaeropteris in monophyletic clades. Splits for species in these genera found in Mesoamerican cloud forests are recent, from the Neogene to the Quaternary, Australia was identified as the ancestral area for the clades of these genera, except for Gymnosphaera that was Mesoamerica. Climate tolerance was not divergent from hypothesized ancestors for the most significant variables or elevation. For elevational shifts, we found repeated change from low to high elevations. Our data suggest that representatives of Cyatheaceae main lineages migrated from Australia to Mesoamerican cloud forests in different times and have persisted in these environmentally unstable areas but extant species diverged recentrly from their ancestors.
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2016-01-01
Background Cloud forests, characterized by a persistent, frequent or seasonal low-level cloud cover and fragmented distribution, are one of the most threatened habitats, especially in the Neotropics. Tree ferns are among the most conspicuous elements in these forests, and ferns are restricted to regions in which minimum temperatures rarely drop below freezing and rainfall is high and evenly distributed around the year. Current phylogeographic data suggest that some of the cloud forest-adapted species remained in situ or expanded to the lowlands during glacial cycles and contracted allopatrically during the interglacials. Although the observed genetic signals of population size changes of cloud forest-adapted species including tree ferns correspond to predicted changes by Pleistocene climate change dynamics, the observed patterns of intraspecific lineage divergence showed temporal incongruence. Methods Here we combined phylogenetic analyses, ancestral area reconstruction, and divergence time estimates with climatic and altitudinal data (environmental space) for phenotypic traits of tree fern species to make inferences about evolutionary processes in deep time. We used phylogenetic Bayesian inference and geographic and altitudinal distribution of tree ferns to investigate ancestral area and elevation and environmental preferences of Mesoamerican tree ferns. The phylogeny was then used to estimate divergence times and ask whether the ancestral area and elevation and environmental shifts were linked to climatic events and historical climatic preferences. Results Bayesian trees retrieved Cyathea, Alsophyla, Gymnosphaera and Sphaeropteris in monophyletic clades. Splits for species in these genera found in Mesoamerican cloud forests are recent, from the Neogene to the Quaternary, Australia was identified as the ancestral area for the clades of these genera, except for Gymnosphaera that was Mesoamerica. Climate tolerance was not divergent from hypothesized ancestors for the most significant variables or elevation. For elevational shifts, we found repeated change from low to high elevations. Conclusions Our data suggest that representatives of Cyatheaceae main lineages migrated from Australia to Mesoamerican cloud forests in different times and have persisted in these environmentally unstable areas but extant species diverged recentrly from their ancestors. PMID:27896030
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Beckage, Brian; Osborne, Ben; Gavin, Daniel G; Pucko, Carolyn; Siccama, Thomas; Perkins, Timothy
2008-03-18
Detecting latitudinal range shifts of forest trees in response to recent climate change is difficult because of slow demographic rates and limited dispersal but may be facilitated by spatially compressed climatic zones along elevation gradients in montane environments. We resurveyed forest plots established in 1964 along elevation transects in the Green Mountains (Vermont) to examine whether a shift had occurred in the location of the northern hardwood-boreal forest ecotone (NBE) from 1964 to 2004. We found a 19% increase in dominance of northern hardwoods from 70% in 1964 to 89% in 2004 in the lower half of the NBE. This shift was driven by a decrease (up to 76%) in boreal and increase (up to 16%) in northern hardwood basal area within the lower portions of the ecotone. We used aerial photographs and satellite imagery to estimate a 91- to 119-m upslope shift in the upper limits of the NBE from 1962 to 2005. The upward shift is consistent with regional climatic change during the same period; interpolating climate data to the NBE showed a 1.1 degrees C increase in annual temperature, which would predict a 208-m upslope movement of the ecotone, along with a 34% increase in precipitation. The rapid upward movement of the NBE indicates little inertia to climatically induced range shifts in montane forests; the upslope shift may have been accelerated by high turnover in canopy trees that provided opportunities for ingrowth of lower elevation species. Our results indicate that high-elevation forests may be jeopardized by climate change sooner than anticipated.
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Tracing the Fate of Atmospheric Nitrate in a Subalpine Watershed Using Δ17O.
Bourgeois, Ilann; Savarino, Joël; Caillon, Nicolas; Angot, Hélène; Barbero, Albane; Delbart, Franck; Voisin, Didier; Clément, Jean-Christophe
2018-05-15
Nitrogen is an essential nutrient for life on Earth, but in excess, it can lead to environmental issues (e.g., N saturation, loss of biodiversity, acidification of lakes, etc.). Understanding the nitrogen budget (i.e., inputs and outputs) is essential to evaluate the prospective decay of the ecosystem services (e.g., freshwater quality, erosion control, loss of high patrimonial-value plant species, etc.) that subalpine headwater catchments provide, especially as these ecosystems experience high atmospheric nitrogen deposition. Here, we use a multi-isotopic tracer (Δ 17 O, δ 15 N and δ 18 O) of nitrate in aerosols, snow, and streams to assess the fate of atmospherically deposited nitrate in the subalpine watershed of the Lautaret Pass (French Alps). We show that atmospheric N deposition contributes significantly to stream nitrate pool year-round, either by direct inputs (up to 35%) or by in situ nitrification of atmospheric ammonium (up to 35%). Snowmelt in particular leads to high exports of atmospheric nitrate, most likely fast enough to impede assimilation by surrounding ecosystems. Yet, in a context of climate change, with shorter snow seasons, and increasing nitrogen emissions, our results hint at possibly stronger ecological consequences of nitrogen atmospheric deposition in the close future.
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Restoring fire to wilderness: Sequoia and Kings Canyon National Parks
Manley, Jeffrey; Keifer, MaryBeth; Stephenson, Nathan L.; Kaage, William
2001-01-01
Sequoia and Kings Canyon National Parks, established in 1890, consist of 863,741 acres (349,551 ha) of Sierra Nevada foothills, mid-elevation conifer forest, and high-elevation alpine environment. The parks contain 36 giant sequoia (Sequoiadendron giganteum) groves, including the largest known tree, the General Sherman. Ninety-four percent of the parklands is in designated or proposed wilderness (fig. 1), with conditions resembling roadless areas in national forests.
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Krushelnycky, P.D.; Joe, S.M.; Medeiros, A.C.; Daehler, C.C.; Loope, L.L.
2005-01-01
Analysis of long-term patterns of invasion can reveal the importance of abiotic factors in influencing invasion dynamics, and can help predict future patterns of spread. In the case of the invasive Argentine ant (Linepithema humile), most prior studies have investigated this species' limitations in hot and dry climates. However, spatial and temporal patterns of spread involving two ant populations over the course of 30 years at a high elevation site in Hawaii suggest that cold and wet conditions have influenced both the ant's distribution and its rate of invasion. In Haleakala National Park on Maui, we found that a population invading at lower elevation is limited by increasing rainfall and presumably by associated decreasing temperatures. A second, higher elevation population has spread outward in all directions, but rates of spread in different directions appear to have been strongly influenced by differences in elevation and temperature. Patterns of foraging activity were strongly tied to soil temperatures, supporting the hypothesis that variation in temperature can influence rates of spread. Based on past patterns of spread, we predicted a total potential range that covers nearly 50% of the park and 75% of the park's subalpine habitats. We compared this rough estimate with point predictions derived from a degree-day model for Argentine ant colony reproduction, and found that the two independent predictions match closely when soil temperatures are used in the model. The cold, wet conditions that have influenced Argentine ant invasion at this site are likely to be influential at other locations in this species' current and future worldwide distribution. ?? 2005 Blackwell Publishing Ltd.
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A comparison of bat activity at low and high elevations in the Black Hills of western Washington
Erickson, J.; Adams, Michael J.
2003-01-01
We examined the differences in activity patterns and community structure of bats between low (<150 m) and high ( ! 575 m) elevation sites in two habitats of the Capitol State Forest, Washington. Total bat activity averaged four times higher at low elevation sites than at high elevation sites. Feeding activity was almost 20 times higher at low elevation sites. However, the non-myotis group had similar activity levels at high and low elevation, whereas myotis group activity decreased at higher elevations. Different levels of activity between elevations could be the result of differences in insect availability, climatic conditions, and morphology of the bat species.
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Todd Allen Bowers; Robert I. Bruck
2010-01-01
Decline in high elevation red spruce (Picea rubens Sarg.) and Fraser fir (Abies fraseri (Pursh) Poir.) forests throughout the southern Appalachians was shown following extensive surveys conducted during the 1980s.
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Habitat characteristics at marten subnivean access sites
Corn, Janelle G.; Raphael, Martin G.
1992-01-01
The occurrence of coarse woody debris (CWD) at sites of subnivean (under snow) access by martens (Martes americana) has not been quantified adequately, and must be better understood to provide suitable winter habitat management for the species. Consequently, we studied subnivean activity of martens in a subalpine forest in southern Wyoming to determine how subnivean space was accessed, and to examine microhabitat characteristics around entry sites. Martens used existing openings in snow, created primarily by logs at low snow depths and by small live spruce and fir trees at greater snow depths. Sites of marten subnivean entry had greater percent cover (P ≤ 0.01) and total volume of CWD (P ≤ 0.01), greater numbers of log layers (all P ≤ 0.02), greater volume of undecayed (P ≤ 0.05) and moderately decayed logs (P ≤ 0.02), less volume of very decayed logs (P ≤ 0.001), and fewer small root masses (P ≤ 0.001) than surrounding forest stands. Provision of sufficient CWD in winter habitat of martens may require specific effort, particularly in managed forests of the central Rocky Mountains.
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Sorenson, Daniel G.
2012-01-01
The Cascades Ecoregion (Omernik, 1987; U.S. Environmental Protection Agency, 1997) covers approximately 46,787 km2 (18,064 mi2) in Washington, Oregon, and California (fig. 1). The main body of the ecoregion extends from Snoqualmie Pass, Washington, in the north, to Hayden Mountain, near State Highway 66 in southern Oregon. Also included in the ecoregion is a small isolated section south of Bend, Oregon, as well as a larger one around Mount Shasta, California. The ecoregion is bounded on the west by the Klamath Mountains, Willamette Valley, and Puget Lowland Ecoregions; on the north by the North Cascades Ecoregion; and on the east by the Eastern Cascades Slopes and Foothills Ecoregion. The Cascades Ecoregion is a forested, mountainous ecoregion, and it contains a large amount of Cenozoic volcanic rock and many active and inactive volcanoes, especially in the east (McNab and Avers, 1994). Elevations range from near sea level at the Columbia River to 4,390 m at Mount Rainier in Washington, with most of the ecoregion between 645 and 2,258 m. The west side of the ecoregion is characterized by long, steep ridges and wide river valleys. Subalpine meadows are present at higher elevations, and alpine glaciers have left till and outwash deposits (McNab and Avers, 1994). Precipitation in the Cascades Ecoregion ranges from 1,300 to 3,800 mm, falling mostly as rain and snow from October to June. Average annual temperatures range from –1ºC to 11ºC. The length of the growing season varies from less than 30 days to 240 days (McNab and Avers, 1994).
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Hartwell H. Welsh Jr; Karen L. Pope; Daniel Boiano
2006-01-01
The goal of this study was to examine how introduced trout influence the distributions and abundances of a sub-alpine amphibian assemblage whose members display a variety of different life-history and defence strategies. Our study was conducted in the sub-alpine lentic habitats of three wilderness areas that form the core of the Klamath-Siskiyou Bioregion of northern...
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Airflow patterns in a small subalpine basin
G. Wooldridge; R. Musselman; B. Connell; D. Fox
1992-01-01
A study of mean wind speeds and directions has been completed in the Snowy Range of Southern Wyoming, U.S.A. It was conducted in a subalpine ecosystem at an altitude of 3 200 m to 3 400 m above sea level during the summers of 1988 and 1989. Indexes of deformation and axes of asymmetry due to wind shaping of Engelmann spruce (Picea engelmannii) and subalpine fir (Abies...
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Biogeography of soil organic matter molecular structure across multiple soil size fractions
NASA Astrophysics Data System (ADS)
Meier, C. L.; Neff, J.
2009-12-01
Recent work suggests that there is a common soil decomposition sequence whereby plant inputs are metabolized into a physiologically constrained set of compounds originating from microbes that may persist in soil over relatively long time-scales. Plant inputs tend to be found in coarse particulate fractions (>180 μm) with relatively fast turnover times, while microbially derived compounds tend to accrue in the finer silt + clay fractions (<53 μm) with relatively long turnover times. To investigate whether a common decomposition sequence exists, we used pyrolysis gas chromatography/mass spectrometry (py-GC/MS) to characterize the molecular structure of soil organic matter (SOM) in three size fractions (590-180 μm, 180-53 μm, and <53 μm), using soils sampled from multiple biomes (alpine tundra, sub-alpine forest, boreal forest, temperate coniferous, temperate deciduous, dry desert/savannah, and tropical forest). We hypothesized that: 1) regardless of biome, fractions >180 μm would be chemically similar, and would be characterized by lignin and other plant-derived compounds; and 2) fractions <53 μm would also be similar across biomes but would be dominated by microbially-derived compounds like polysaccharides. Across all biomes, we found that there was significantly less lignin in <53 μm fractions compared to >180 μm fractions (p<0.0001), providing some support for the idea that plant material is not incorporated into soil C pools with relatively long turnover times. However, a principal components analysis (PCA) showed that the >180 μm coarse particulate fractions also contained compounds associated with microbial origins, indicating that microbial C is not limited to <53 μm size fractions. The PCA also revealed that samples within each of the three size fractions did not cluster together (i.e. they did not share a common molecular structure), but we did note that: 1) cold alpine and sub-alpine sites were unique and chemically similar; and 2) tropical forest soils were unique and chemically similar. Moreover, we observed large differences in molecular structure for dry desert/savannah sites with varying vegetation types (trees vs. grass) and varying geologic substrates. Taken together, these observations argue that temperature, vegetation, and underlying geology influence soil molecular structure, but support for a common decomposition sequence is mixed.
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Monitoring Carbon Fluxes from Shallow Surface Soils in the Critical Zone
NASA Astrophysics Data System (ADS)
Stielstra, C. M.; Brooks, P. D.; Chorover, J.
2011-12-01
The critical zone (CZ) is the earth's porous near-surface layer, characterized by the integrated processes that occur between the bedrock and the atmospheric boundary layer. Within this area water, atmosphere, ecosystems, and soils interact on a geomorphic and geologic template. We hypothesize that CZ systems organize and evolve in response to open system fluxes of energy and mass, including meteoric inputs of radiation, water, and carbon, which can be quantified at point to watershed scales. The goal of this study is to link above-ground and below-ground carbon processes by quantifying carbon pools and fluxes from near surface soils. Soil CO2 efflux and dissolved organic carbon (DOC) are monitored over a two year period across bedrock type and vegetation type at two seasonally snow covered subalpine catchments in Arizona and New Mexico. We measure the amount of DOC present in surface soils, and install ion exchange resins at the A/B soil horizon interface to capture DOC leachate mobilized during snowmelt and summer rainfall. Throughout the summer rain and spring snowmelt seasons we monitor soil respiration of CO2. Preliminary results show that rates of gaseous carbon flux are significantly higher (p<0.05) from soils with schist bedrock (2.5 ± 0.2 gC/m2/d )than from granite bedrock (1.3 ± 0.1 gC/m2/d), and higher from healthy mixed conifer forests (1.9 ± 0.3 gC/m2/d) than from mixed conifer forests impacted by spruce budworm (1.4 ± 0.1 gC/m2/d). DOC leached from soil samples does not vary significantly with bedrock type; however, spruce budworm impacted forests have significantly higher levels of leachable DOC in surface soils (22.8 ± 4.5 gC/m2) than are found in the soils of healthy forests (10.0 ± 1.5 gC/m2) or subalpine meadows (9.1 ± 0.5 gC/m2). The results of this study will allow us to evaluate the variability of carbon fluxes with vegetation and soil type within a shallow soil carbon pool and help constrain the contributions of soil organic carbon to net carbon balance in CZO catchments with seasonal precipitation regimes.
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Das, Arundhati; Nagendra, Harini; Anand, Madhur; Bunyan, Milind
2015-01-01
The objective of this analysis was to identify topographic and bioclimatic factors that predict occurrence of forest and grassland patches within tropical montane forest-grassland mosaics. We further investigated whether interactions between topography and bioclimate are important in determining vegetation pattern, and assessed the role of spatial scale in determining the relative importance of specific topographic features. Finally, we assessed the role of elevation in determining the relative importance of diverse explanatory factors. The study area consists of the central and southern regions of the Western Ghats of Southern India, a global biodiversity hotspot. Random forests were used to assess prediction accuracy and predictor importance. Conditional inference classification trees were used to interpret predictor effects and examine potential interactions between predictors. GLMs were used to confirm predictor importance and assess the strength of interaction terms. Overall, topographic and bioclimatic predictors classified vegetation pattern with approximately 70% accuracy. Prediction accuracy was higher for grassland than forest, and for mosaics at higher elevations. Elevation was the most important predictor, with mosaics above 2000m dominated largely by grassland. Relative topographic position measured at a local scale (within a 300m neighbourhood) was another important predictor of vegetation pattern. In high elevation mosaics, northness and concave land surface curvature were important predictors of forest occurrence. Important bioclimatic predictors were: dry quarter precipitation, annual temperature range and the interaction between the two. The results indicate complex interactions between topography and bioclimate and among topographic variables. Elevation and topography have a strong influence on vegetation pattern in these mosaics. There were marked regional differences in the roles of various topographic and bioclimatic predictors across the range of study mosaics, indicating that the same pattern of grass and forest seems to be generated by different sets of mechanisms across the region, depending on spatial scale and elevation. PMID:26121353
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Das, Arundhati; Nagendra, Harini; Anand, Madhur; Bunyan, Milind
2015-01-01
The objective of this analysis was to identify topographic and bioclimatic factors that predict occurrence of forest and grassland patches within tropical montane forest-grassland mosaics. We further investigated whether interactions between topography and bioclimate are important in determining vegetation pattern, and assessed the role of spatial scale in determining the relative importance of specific topographic features. Finally, we assessed the role of elevation in determining the relative importance of diverse explanatory factors. The study area consists of the central and southern regions of the Western Ghats of Southern India, a global biodiversity hotspot. Random forests were used to assess prediction accuracy and predictor importance. Conditional inference classification trees were used to interpret predictor effects and examine potential interactions between predictors. GLMs were used to confirm predictor importance and assess the strength of interaction terms. Overall, topographic and bioclimatic predictors classified vegetation pattern with approximately 70% accuracy. Prediction accuracy was higher for grassland than forest, and for mosaics at higher elevations. Elevation was the most important predictor, with mosaics above 2000 m dominated largely by grassland. Relative topographic position measured at a local scale (within a 300 m neighbourhood) was another important predictor of vegetation pattern. In high elevation mosaics, northness and concave land surface curvature were important predictors of forest occurrence. Important bioclimatic predictors were: dry quarter precipitation, annual temperature range and the interaction between the two. The results indicate complex interactions between topography and bioclimate and among topographic variables. Elevation and topography have a strong influence on vegetation pattern in these mosaics. There were marked regional differences in the roles of various topographic and bioclimatic predictors across the range of study mosaics, indicating that the same pattern of grass and forest seems to be generated by different sets of mechanisms across the region, depending on spatial scale and elevation.
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75. Southeast elevation of Forest Hills station looking Northwest ...
75. Southeast elevation of Forest Hills station - looking Northwest from junction of Washington and Walk Hill Streets. At left is the beginning of Section F-7 the exposed steel portion of elevated structure leading to the Forest Hills storage yard (demolished in 1985). - Boston Elevated Railway, Elevated Mainline, Washington Street, Boston, Suffolk County, MA
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NASA Astrophysics Data System (ADS)
Lin, Y.; Gross, A.; Silver, W. L.
2017-12-01
Elemental stoichiometry of microorganisms is intimately related to ecosystem carbon and nutrient fluxes and is ultimately controlled by the chemical (plant tissue, soil, redox) and physical (temperature, moisture, aeration) environment. Previous meta-analyses have shown that the C:P ratio of soil microbial biomass exhibits significant variations among and within biomes. Little is known about the underlying causes of this variability. We examined soil microbial C:P ratios along an elevation gradient in the Luquillo Experimental Forest in Puerto Rico. We analyzed soils from mixed forest paired with monodominant palm forest every 100 m from 300 m to 1000 m a.s.l.. Mean annual precipitation increased with increasing elevation, resulting in stronger reducing conditions and accumulation of soil Fe(II) at higher elevations. The mean value and variability of soil microbial C:P ratios generally increased with increasing elevation except at 1000 m. At high elevations (600-900 m), the average value of microbial C:P ratio (108±10:1) was significantly higher than the global average ( 55:1). We also found that soil organic P increased with increasing elevation, suggesting that an inhibition of organic P mineralization, not decreased soil P availability, may cause the high microbial C:P ratio. The soil microbial C:P ratio was positively correlated with soil HCl-extractable Fe(II), suggesting that reducing conditions may be responsible for the elevational changes observed. In a follow-up experiment, soils from mixed forests at four elevation levels (300, 500, 700, and 1000 m) were incubated under aerobic and anaerobic conditions for two weeks. We found that anaerobic incubation consistently increased the soil microbial C:P ratio relative to the aerobic incubation. Overall, our results indicate that redox conditions can shift the elemental composition of microbial biomass. The high microbial C:P ratios induced under anoxic conditions may reflect inhibition of microbial P mineralization and/or immobilization under reducing conditions. These results will help us to better understand the patterns of nutrient cycling and microbial activity across macro-scale environmental gradients.
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Xu, Zhen-feng; Hu, Ting-xing; Zhang, Li; Zhang, Yuan-bin; Xian, Jun-ren; Wang, Kai-yun
2009-01-01
By using open-top chamber (OTC), the effects of simulated warming on the growth, leaf phenology, and leaf traits of Salix eriostachya in sub-alpine timberline ecotone of Western Sichuan were studied. The results showed that comparing with the control, the mean air temperature at 1.2 m above the ground throughout S. eriostachya growth season in OTC increased by 2.9 degrees C, while the soil temperature at the depth of 5 cm only increased by 0.4 degrees C. The temperature increase in OTC made S. eriostachya budding advanced and defoliation postponed obviously, and the leaf life-span longer. The leaf and branch growth rates as well as the specific leaf area in OTC increased obviously, whereas the leaf nitrogen concentration decreased significantly. In OTC, the stomata conductance, net photosynthetic rate, photorespiration, and dark respiration rate of S. eriostachya all exhibited an increasing trend. It was suggested that S. eriostachya had stronger capability to adapt to warming, and, under the background of future global climate change, the elevation of S. eriostachya distribution in the timberline ecotone would be likely to ascend.
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Clark, Jason A.; Loehman, Rachel A.; Keane, Robert E.
2017-01-01
We present landscape simulation results contrasting effects of changing climates on forest vegetation and fire regimes in Yellowstone National Park, USA, by mid-21st century. We simulated potential changes to fire dynamics and forest characteristics under three future climate projections representing a range of potential future conditions using the FireBGCv2 model. Under the future climate scenarios with moderate warming (>2°C) and moderate increases in precipitation (3–5%), model simulations resulted in 1.2–4.2 times more burned area, decreases in forest cover (10–44%), and reductions in basal area (14–60%). In these same scenarios, lodgepole pine (Pinus contorta) decreased in basal area (18–41%), while Douglas-fir (Pseudotsuga menziesii) basal area increased (21–58%). Conversely, mild warming (<2°C) coupled with greater increases in precipitation (12–13%) suggested an increase in forest cover and basal area by mid-century, with spruce and subalpine fir increasing in abundance. Overall, we found changes in forest tree species compositions were caused by the climate-mediated changes in fire regime (56–315% increase in annual area burned). Simulated changes in forest composition and fire regime under warming climates portray a landscape that shifts from lodgepole pine to Douglas-fir caused by the interaction between the magnitude and seasonality of future climate changes, by climate-induced changes in the frequency and intensity of wildfires, and by tree species response.
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NASA Astrophysics Data System (ADS)
Alados, Concepción L.; Gartzia, Maite; Nuche, Paloma; Saiz, Hugo; Pueyo, Yolanda
2017-04-01
Anthropogenic activities have modified vegetation in subalpine belts for long time, lowering treeline ecotone and influencing landscape mainly through grazing and fire. During the last decades the abandonment of traditional land use practices and global warming are contributing largely to the colonization of woody species in subalpine grasslands causing irreversible changes in ecosystem functioning. To prevent those changes a variety of management strategies are carried out to stop the expansion of the highly encroaching shrubs, which require the use of fire and/or clear-cutting, particularly for the conservation of grasslands in subhumid high productive ecosystems. However, it is still poorly understood how different management strategies affect the recovery of subalpine grasslands. Using a field experiment we tested the impact of management treatments on soil properties and vegetation characteristics, including species richness, community structure, interspecies interaction, and complexity of network association. Vegetation was monitored during four years in eight stands (two stands per treatment) where the vegetation was removed by prescribed fire (Burnt treatment), or by mechanical removal (Clear-cut treatment). Two undisturbed E. horridum stands were used as a control (C-Erizón) and two grassland communities regularly grazed (C-Grass) were used as a control for subalpine grassland. Soils nutrients declined in Burnt treatment 3 years after fire, but not differences between Clear-cut and C-Erizón were observed. Species richness and diversity were larger in C-Grass and lower in C-Erizón. Burnt and Clear-cut treatments increased species diversity and richness gradually after 4 year treatment. The proportion of legume forbs, grasses and non-legume forbs did not reach the levels of C-Grass after 4 years of Clear-cut or Burnt treatments. Shrubs and sub-shrubs increased faster after 4 years of burning than after 4 years of clearing, although they did not reach the level of the C-Erizón. Network connectivity and number of association was larger in well preserved grasslands (C-Grass) in comparison with any of the other treatments, mainly because of the abundance of negative plant-plant associations. On the contrary, the ratio of positive to negative associations was higher in C-Erizón and also, to a lesser extent in the Burnt treatment. This study shows that grazing favors plant diversity and community complexity in subalpine grasslands, and that clearing is a better strategy than burning to recover grasslands after shrub encroachment because burning entails deeper soil degradation and a faster recovery of the pyrophyte shrub, E. horridum.
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Paxton, Eben H.; Gorresen, P. Marcos; Camp, Richard J.
2013-01-01
Naturalists in the 1800s described the ʻIʻiwi (Vestiaria coccinea) as one of the most abundant forest birds, detected in forested areas from sea level to tree line across all the major Hawaiian Islands. However, in the late 1800s, ʻIʻiwi began to disappear from low elevation forests, and by the mid-1900s, the species was largely absent from low- and mid-elevation areas. Today, ʻIʻiwi are restricted to high-elevation forests on the islands of Hawaiʻi, east Maui, and Kauaʻi, with a few birds apparently persisting on Oʻahu, Molokaʻi, and west Maui. ʻIʻiwi are highly vulnerable to introduced disease, and the prevalence of avian malaria in low and mid-elevations is believed to be the cause of ʻIʻiwi being restricted to high elevations where temperatures are too cold for the development of the disease and its mosquito vector. With global warming, it is feared that the disease will move quickly into the high-elevation forests where the last ʻIʻiwi reside, threatening their viability. The U.S. Fish and Wildlife Service was petitioned to list the ʻIʻiwi as an Endangered Species in 2010, and this report provides a comprehensive review of the abundance, distribution, and trends using historical survey data as well as the most recently available survey information (up to 2012). We estimate the total population size of ‘I‘iwi at 550,972–659,864 (mean = 605,418) individuals. Of these, 90 percent are on the island of Hawaiʻi, followed by east Maui (about 10 percent), with less than 1 percent on Kauaʻi. ʻIʻiwi population trends vary across the islands. ʻIʻiwi population in Kauaʻi has experienced sharp declines, with a projected trend of 92 percent decline over a 25 year period based on the 2000–2012 surveys. On East Maui, the northeastern region has experienced declines (34 percent over a 25 year period), while the southeastern region has been stable to moderately increasing. On the island of Hawaiʻi, population trends are mixed. On the windward side, populations are largely declining, although the northern section (Hakalau Forest) has stable populations. On the leeward side, results suggest a strongly increasing population, with estimates of as much as a 147 percent increase over a 25 year period from the Puʻu Waʻawaʻa region. However, it is unclear how much these results from the leeward side of Hawaiʻi show a population trend contrary to population trends in all other areas or are an artifact of a sparsely sampled area. Trends by elevation suggest a large decrease in numbers of ʻIʻiwi at elevations below 1,200 meters on Kauaʻi and northeast Maui. Low elevation ʻIʻiwi populations also appear to have decreased in other regions, although low-elevation areas are not surveyed as often as other areas because of their lack of native forest birds. An exception to this pattern was the lower portions of the Hakalau Forest National Wildlife Refuge Kona Unit in the central leeward part of the island of Hawaiʻi, where populations appeared stable at the lower elevations. Based on the most recent surveys (up to 2012), approximately 50 percent of ʻIʻiwi live in a narrow, 500-meter band at elevations of 1,200–1,700 meters, suggesting that ʻIʻiwi are vulnerable to future shifts in climate.
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Can lowland dry forests represent a refuge from avian malaria for native Hawaiian birds?
Tucker-Mohl, Katherine; Hart, Patrick; Atkinson, Carter T.
2010-01-01
Hawaii's native birds have become increasingly threatened over the past century. Introduced mosquito borne diseases such as avian malaria may be responsible for the near absence of endemic Hawaiian forest birds in low-elevation habitats. The recent recognition that some native Hawaiian forest birds may be repopulating moist lowland habitats as a result of evolved resistance to this disease has increased the conservation value of these areas. Here, we investigate whether remnant low elevation dry forests on Hawaii Island provide natural 'refuges' from mosquito-transmitted malaria by nature of their low rainfall and absence of suitable natural sources of water for mosquito breeding. Unlike lowland wet forests where high rates of disease transmission may be selecting for disease resistance, lowland dry forests may provide some refuge for native forest birds without natural resistance to malaria. We mistnetted forest birds in two lowland dry forests and tested all native birds by microscopy and serology for avian malaria caused by the Plasmodium relictum parasite. We also conducted surveys for standing water and mosquito larvae. Overall prevalence of infections with Plasmodium relictum in the Hawaii Amakihi Hemignathus virens virens was 15%. Most infected birds had lowlevel parasitemias, suggesting chronic infections. Although avian malaria is present in these lowland dry forest Amakihi populations, infection rates are significantly lower than in wet forest populations at similar elevations. Sources of breeding mosquitoes in these forests appeared to be largely anthropogenic; thus, there is potential to manage dry forests as mosquito-free habitat for Hawaii Amakihi and other Hawaiian forest birds.
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Space and habitat use by black bears in the Elwha valley prior to dam removal
Sager-Fradkin, K.A.; Jenkins, K.J.; Happe, P.J.; Beecham, J.J.; Wright, R.G.; Hoffman, R.A.
2008-01-01
Dam removal and subsequent restoration of salmon to the Elwha River is expected to cause a shift in nutrient dynamics within the watershed. To document how this influx of nutrients and energy may affect black bear (Ursus americanus) ecology, we used radio-telemetry to record movements of 11 male and two female black bears in the Elwha Valley from 2002-06. Our objective was to collect baseline data on bear movements prior to dam removal. We calculated annual home ranges, described seasonal timing of den entry and emergence, and described seasonal patterns of distribution and habitat use. Adaptive kernel home ranges were larger formales (mean = 151.1 km2, SE = 21.4) than females (mean = 38.8 km2, SE = 13.0). Males ranged widely and frequently left the watershed during late summer. Further, they exhibited predictable and synchronous patterns of elevation change throughout each year. Bears entered their winter dens between 8 October and 15 December and emerged from dens between 10 March and 9 May. Male bears used low-elevation conifer and hardwood forests along the Elwha floodplain during spring, mid- to high-elevation forests and meadows during early summer, high-elevation forests, meadows and shrubs during late summer, and mid-elevation forests, shrubs and meadows during fall. Data acquired during this study provide important baseline information for comparison after dam removal, when bears may alter their late summer and fall movement and denning patterns to take advantage of energy-rich spawning salmon.
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Andersen, Douglas C.; MacMahon, James A.
1981-01-01
Studies of the bioenergetics of the northern pocket gopher, Thomomys talpoides, are coupled with data on demography, activity budgets, and microclimates to model the energy requirements of individuals and populations in the Wasatch Mountains of Utah during 1976-1979. Metabolic rates during rest increased linearly with decreasing ambient temperature, but burrowing metabolic rates (16.3 mL O2 • h-1 • g-9.75) were independent of both temperature and physical properties of the soil. Radio-telemetry studies indicated that free-ranging gophers are active =50% of each day. Conservative estimates of true energy consumption were calculated using estimates of habitat-specific minimum daily burrowing requirements. Rates of burrowing measured in the laboratory were either ∞ 0.0 or ∞ 2.0 cm/min. The low burrowing rate was observed when the soil was frozen or saturated with water, as would occur in the field in early winter and in spring, respectively. Gophers burrowed through soil at the study site at an average rate of ∞ 1.5 cm/min. Belowground food energy densities at gopher foraging depth declined from 24.6 to 3.2 J/cm3 along a successional gradient (subalpine forb meadow to Engelmann spruce dominated forest). We conclude that individual gophers are food limited within the climax spruce seral stage. Further, daily energy costs associated with reproduction in females may exceed the belowground energy supply available in intermediate seral stages (aspen and subalpine fir). Reduction of burrowing rates for any reason will affect gophers in the late seral stages proportionately more than those resident in the meadow. The peak gopher densities recorded (from 62 individuals/ha in the meadow to 2 individuals/ha in spruce forest) support these inferences. Detailed demographic information was obtained only in the meadow seral stage. Adult survivorship was lower in winter than in summer and varied greatly between years (0.18-0.70 yr-1). Juvenile survivorship from weaning through the first year was comparable to adult annual rates. The fertility rate was 3.75 young • female-1 • yr-1. The energy supply and demand analyses indicate that the growth of Thomomys talpoides populations in the early seral stages is seldom directly limited by the amount of food present. From our demographic, environmental, and autecological studies we conclude that stochastic events associated with weather affect energy acquisition (burrowing) rates, and thus survivorship. In montane environments, such events may prevent populations from attaining sizes at which territorial behavior would hypothetically limit further increases. The energy flow through the meadow population at moderate to high )1976-1977) densities (at least 1100 MJ • ha-1 • yr-1) indicates that pocket gophers are proficient energy movers relative to non-fossorial small mammals. Subalpine T. talpoides populations appear commonly to attain such densities. More than 30% of the annual primary productivity allocated to belowground parts of meadow forbs may be consumed by gophers.
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Drought-induced weakening of growth-temperature associations in high-elevation Iberian pines
NASA Astrophysics Data System (ADS)
Diego Galván, J.; Büntgen, Ulf; Ginzler, Christian; Grudd, Håkan; Gutiérrez, Emilia; Labuhn, Inga; Julio Camarero, J.
2015-01-01
The growth/climate relationship of theoretically temperature-controlled high-elevation forests has been demonstrated to weaken over recent decades. This is likely due to new tree growth limiting factors, such as an increasing drought risk for ecosystem functioning and productivity across the Mediterranean Basin. In addition, declining tree growth sensitivity to spring temperature may emerge in response to increasing drought stress. Here, we evaluate these ideas by assessing the growth/climate sensitivity of 1500 tree-ring width (TRW) and 102 maximum density (MXD) measurement series from 711 and 74 Pinus uncinata trees, respectively, sampled at 28 high-elevation forest sites across the Pyrenees and two relict populations of the Iberian System. Different dendroclimatological standardization and split period approaches were used to assess the high- to low-frequency behavior of 20th century tree growth in response to temperature means, precipitation totals and drought indices. Long-term variations in TRW track summer temperatures until about 1970 but diverge afterwards, whereas MXD captures the recent temperature increase in the low-frequency domain fairly well. On the other hand summer drought has increasingly driven TRW along the 20th century. Our results suggest fading temperature sensitivity of Iberian high-elevation P. uncinata forest growth, and reveal the importance of summer drought that is becoming the emergent limiting factor of tree ring width formation in many parts of the Mediterranean Basin.
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Soil microbial metabolic quotient (qCO2) of twelve ecosystems of Mt. Kilimanjaro
NASA Astrophysics Data System (ADS)
Pabst, Holger; Gerschlauer, Friederike; Kiese, Ralf; Kuzyakov, Yakov
2014-05-01
Soil organic carbon, microbial biomass carbon (MBC) and the metabolic quotient qCO2 - as sensitive and important parameters for soil fertility and C turnover - are strongly affected by land-use changes all over the world. These effects are particularly distinct upon conversion of natural to agricultural ecosystems due to very fast carbon (C) and nutrient cycles and high vulnerability, especially in the tropics. In this study, we used an elevational gradient on Mt. Kilimanjaro to investigate the effects of land-use change and elevation on Corg, MBC and qCO2. Down to a soil depth of 18 cm we compared 4 natural (Helichrysum, Erica forest, Podocarpus forest, Ocotea forest), 5 seminatural (disturbed Podocarpus forest, disturbed Ocotea forest, lower montane forest, grassland, savannah), 1 sustainably used (homegarden) and 2 intensively used ecosystems (coffee plantation, maize field) on an elevation gradient from 950 to 3880 m a.s.l.. Using an incubation device, soil CO2-efflux of 18 cm deep soil cores was measured under field moist conditions and mean annual temperature. MBC to Corg ratios varied between 0.7 and 2.3%. qCO2 increased with magnitude of the disturbance, albeit this effect decreased with elevation. Following the annual precipitation of the ecosystems, both, Corg and MBC showed a hum-shaped distribution with elevation, whereas their maxima were between 2500 and 3000 m a.s.l.. Additionaly, Corg and MBC contents were significantly reduced in intensively used agricultural systems. We conclude that the soil microbial biomass and its activity in Mt. Kilimanjaro ecosystems are strongly altered by land-use. This effect is more distinct in lower than in higher elevated ecosystems and strongly dependent on the magnitude of disturbance.
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Species abundances, guild dominance patterns, and community structure of breeding riparian birds
Deborah M. Finch
1989-01-01
I investigated trends in bird species populations, guild structure, and bird communities along a riparian altitudinal cline in the Medicine Bow National Forest of southeastern Wyoming. Streamside habitats were divided into three elevational zones: low-elevation (2050 to 2260 m) cottonwood zone, mid-elevation (2290 to 2530 m) mixed shrub willow zone, and high elevation...
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Norman H. Miner; James M. Trappe
1957-01-01
Lodgepole pine (Pinus contorta) forests in the Blue Mountains of eastern Oregon occupy important water-producing lands. These forests generally occur at middle to high elevations on north slopes, where a substantial portion of the precipitation is snow. To learn more about the influence of lodgepole pine forests on accumulation of mow and rate of...
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David Rogers Tilley; Wayne T. Swank
2003-01-01
Emergy (with an 'm') synthesis was used to assess the balance between nature and humanity and the equity among forest outcomes of a US Forest Service ecosystem management demonstration project on the Wine Spring Creek watershed, a high-elevation (1600 m), temperate forest located in the southern Appalachian mountains of North Carolina, USA. EM embraces a...
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Dwarf forest recovery after disturbances in the Luquillo Mountains of Puerto Rico
P.L. Weaver
2008-01-01
Dwarf forest in Puerto Ricoâs Luquillo Mountains varies according to substrate and topography with very short, dense forest growing on exposed, rocky sites. High elevation level sites suffered considerable damage during past hurricanes whereas the trees on certain lower slopes were protected by ridges or spurs. Post-disturbance recovery of dwarf forest on two types of...
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Gavazov, Konstantin; Spiegelberger, Thomas; Buttler, Alexandre
2014-04-01
Climate change could impact strongly on cold-adapted mountain ecosystems, but little is known about its interaction with traditional land-use practices. We used an altitudinal gradient to simulate a year-round warmer and drier climate for semi-natural subalpine grasslands across a landscape of contrasting land-use management. Turf mesocosms from three pasture-woodland land-use types-unwooded pasture, sparsely wooded pasture, and densely wooded pasture-spanning a gradient from high to low management intensity were transplanted downslope to test their resistance to two intensities of climate change. We found strong overall effects of intensive (+4 K) experimental climate change (i.e., warming and reduced precipitation) on plant community structure and function, while moderate (+2 K) climate change did not substantially affect the studied land-use types, thus indicating an ecosystem response threshold to moderate climate perturbation. The individual land-use types were affected differently under the +4 K scenario, with a 60% decrease in aboveground biomass (AGB) in unwooded pasture turfs, a 40% decrease in sparsely wooded pasture turfs, and none in densely wooded ones. Similarly, unwooded pasture turfs experienced a 30% loss of species, advanced (by 30 days) phenological development, and a mid-season senescence due to drought stress, while no such effects were recorded for the other land-use types. The observed contrasting effects of climate change across the pasture-woodland landscape have important implications for future decades. The reduced impact of climate change on wooded pastures as compared to unwooded ones should promote the sustainable land use of wooded pastures by maintaining low management intensity and a sparse forest canopy, which buffer the immediate impacts of climate change on herbaceous vegetation.
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Forest Types in the Lower Suwannee River Floodplain, Florida?-A Report and Interactive Map
Darst, M.R.; Light, H.M.; Lewis, L.J.; Sepulveda, A.A.
2003-01-01
A map of forest types in the lower Suwannee River floodplain, Florida, was created during a study conducted from 1996 to 2000 by the U.S. Geological Survey in cooperation with the Suwannee River Water Management District. The map is presented with this report on a compact disc with interactive viewing software. The forest map can be used by scientists for ecological studies in the floodplain based on land cover types and by landowners and management personnel making land use decisions. The study area is the 10-year floodplain of the lower Suwannee River from its confluence with the Santa Fe River to the lower limit of forests near the Gulf of Mexico. The floodplain is divided into three reaches: riverine (non-tidal), upper tidal, and lower tidal, due to changes in hydrology, vegetation, and soils with proximity to the coast. The 10-year floodplain covers about 21,170 hectares; nearly 88 percent of this area (18,580 hectares) is mapped as 14 major forest types. Approximately 29 percent (5,319 hectares) of these forests have been altered by agriculture or development. About 75 percent of the area of major forest types (13,994 hectares) is wetland forests and about 25 percent (4,586 hectares) is upland forests. Tidal wetland forests (8,955 hectares) cover a much greater area than riverine wetland forests (5,039 hectares). Oak/pine upland forests are present in the riverine and upper tidal reaches of the floodplain on elevations that are inundated only briefly during the highest floods. High bottomland hardwoods are present on the higher levees, ridges, and flats of the riverine reach where soils are usually sandy. Low bottomland hardwood forests are present in the riverine reach on swamp margins and low levees and flats that are flooded continuously for several weeks or longer every 1 to 3 years. Riverine swamps are present in the lowest and wettest areas of the non-tidal floodplain that are either inundated or saturated most of the time. Upper tidal bottomland hardwood forests are present on sandy soils on high flats and in transitional areas between upland forests and swamps. Upper tidal mixed forests are found on low levees or between swamps and higher forest types. Upper tidal swamps are present at elevations below median monthly high stage and usually have surface soils that are permanently saturated mucks. Lower tidal hammocks are found on higher elevations that do not receive regular tidal inundation but have a high water table and are briefly inundated by storm surges several times a decade. Lower tidal mixed forests include swamps with numerous small hummocks or less common larger hummocks. Lower tidal swamps are found on deep muck soils that are below the elevation of the median daily or monthly high stage. Seven additional land cover types (2,590 hectares) are mapped. Water in the main channel of the lower Suwannee River (1,767 hectares) was mapped separately from open water in the floodplain (239 hectares). Other land cover types are: seepage slopes (70 hectares), isolated forested wetlands (19 hectares), marshes upstream of the tree line (505 hectares), beds of emergent aquatic vegetation (21 hectares), and floodplain glades (46 hectares)
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Ecology and conservation biology of avian malaria.
Lapointe, Dennis A; Atkinson, Carter T; Samuel, Michael D
2012-02-01
Avian malaria is a worldwide mosquito-borne disease caused by Plasmodium parasites. These parasites occur in many avian species but primarily affect passerine birds that have not evolved with the parasite. Host pathogenicity, fitness, and population impacts are poorly understood. In contrast to continental species, introduced avian malaria poses a substantial threat to naive birds on Hawaii, the Galapagos, and other archipelagoes. In Hawaii, transmission is maintained by susceptible native birds, competence and abundance of mosquitoes, and a disease reservoir of chronically infected native birds. Although vector habitat and avian communities determine the geographic distribution of disease, climate drives transmission patterns ranging from continuous high infection in warm lowland forests, seasonal infection in midelevation forests, and disease-free refugia in cool high-elevation forests. Global warming is expected to increase the occurrence, distribution, and intensity of avian malaria across this elevational gradient and threaten high-elevation refugia, which is the key to survival of many susceptible Hawaiian birds. Increased temperatures may have already increased global avian malaria prevalence and contributed to an emergence of disease in New Zealand. © 2012 New York Academy of Sciences.
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Ecology and conservation biology of avian malaria
LaPointe, Dennis A.; Atkinson, Carter T.; Samuel, Michael D.
2012-01-01
Avian malaria is a worldwide mosquito-borne disease caused by Plasmodium parasites. These parasites occur in many avian species but primarily affect passerine birds that have not evolved with the parasite. Host pathogenicity, fitness, and population impacts are poorly understood. In contrast to continental species, introduced avian malaria poses a substantial threat to naive birds on Hawaii, the Galapagos, and other archipelagoes. In Hawaii, transmission is maintained by susceptible native birds, competence and abundance of mosquitoes, and a disease reservoir of chronically infected native birds. Although vector habitat and avian communities determine the geographic distribution of disease, climate drives transmission patterns ranging from continuous high infection in warm lowland forests, seasonal infection in midelevation forests, and disease-free refugia in cool high-elevation forests. Global warming is expected to increase the occurrence, distribution, and intensity of avian malaria across this elevational gradient and threaten high-elevation refugia, which is the key to survival of many susceptible Hawaiian birds. Increased temperatures may have already increased global avian malaria prevalence and contributed to an emergence of disease in New Zealand.
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Aluminium concentrations in Swedish forest streams and co-variations with catchment characteristics.
Löfgren, Stefan; Cory, Neil; Zetterberg, Therese
2010-07-01
The negative effects of elevated concentrations of inorganic aluminium on aquatic organisms are well documented. Acid deposition is often cited as a main driver behind the mobilisation and speciation of aluminium in soils and surface waters. In the study, we tested the hypothesis that sulphur deposition is the main driver for elevated concentrations of inorganic aluminium in 114 base poor, boreal Swedish streams. However, the deposition of anthropogenic sulphate has decreased substantially since it peaked in the 1970s, and at the current deposition levels, we hypothesise that local site parameters play an important role in determining vulnerability to elevated concentrations of inorganic aluminium in boreal stream waters. Presented here are the results of a principal components analysis of stream water chemistry, acid deposition data and local site variables, including forest composition and stem volume. It is shown that the concentrations of both organic and inorganic aluminium are not explained by either historical or current acid deposition, but are instead explained by a combination of local site characteristics. Sites with elevated concentrations of inorganic aluminium were characterised by small catchments (<500 ha) dominated by mature stands of Norway spruce with high stem volume. Using data from the Swedish National Forest Inventory the area of productive forest land in Sweden with a higher vulnerability for elevated inorganic aluminium concentrations in forests streams is approximately 1.5 million hectares or 7% of the total productive forest area; this is higher in the south of Sweden (10%) and lower in the north (2%). A better understanding of the effects of natural processes and forest management in controlling aquatic inorganic aluminium concentrations is therefore important in future discussions about measures against surface water acidification.
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[Effects of Different Altitudes on Soil Microbial PLFA and Enzyme Activity in Two Kinds of Forests].
Zeng, Qing-ping; He, Bing-hui; Mao, Qiao-zhi; Wu, Yao-peng; Huang, Qi; Li, Yuan
2015-12-01
The soil microbial community is an important part in soil ecosystem, and it is sensitive to the ecological environment. Phospholipid-derived fatty acids ( PLFA ) analysis was used to examine variations in soil microbial community diversity and its influencing factors. The results showed that: there existed 48 PLFAs that were significant in the soil samples from six altitudes. The PLFAs of six altitudes with the highest contents were i16:0, 10Me17:0, 10Me18:0 TBSA. The citrus forest exhibited richer soil PLFAs distribution both in type and amount than those in masson pine. The microbial activity and functional diversity of masson pine were increased with increasing altitudes, and citrus forest gradually decreased, the PLFA content of different microbial groups in each altitude were significantly different. The richness index, Shannon-Wiener index and Pielou evenness index of masson pine in low elevation were holistically higher than those in high elevation. However, the highest richness index of citrus forest was in low altitude, the highest Shannon-Wiener index and Pielou evenness index were in high altitude. The PLFAs content of different microbial groups were closely correlated to the soil enzyme activities and environmental factors. The PLFAs of bacteria, actinomycetes, G⁻ (Gram- positive), G⁺ (Gram-negative) were positively correlated with Ure(urease) , Ive(invertase) , CAT( catalase activity) and forest type, the PLFAs of fungi was significantly correlated with Ure, Ive, CAT, the PLFAs of bacteria, fungi, actinomycetes, G⁻ , G⁺ were significantly negatively or less correlated with elevation. Ure, Ive, CAT, forest type and elevation are the pivotal factors controlling the soil microbial biomass and activities.
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Beckage, Brian; Osborne, Ben; Gavin, Daniel G.; Pucko, Carolyn; Siccama, Thomas; Perkins, Timothy
2008-01-01
Detecting latitudinal range shifts of forest trees in response to recent climate change is difficult because of slow demographic rates and limited dispersal but may be facilitated by spatially compressed climatic zones along elevation gradients in montane environments. We resurveyed forest plots established in 1964 along elevation transects in the Green Mountains (Vermont) to examine whether a shift had occurred in the location of the northern hardwood–boreal forest ecotone (NBE) from 1964 to 2004. We found a 19% increase in dominance of northern hardwoods from 70% in 1964 to 89% in 2004 in the lower half of the NBE. This shift was driven by a decrease (up to 76%) in boreal and increase (up to 16%) in northern hardwood basal area within the lower portions of the ecotone. We used aerial photographs and satellite imagery to estimate a 91- to 119-m upslope shift in the upper limits of the NBE from 1962 to 2005. The upward shift is consistent with regional climatic change during the same period; interpolating climate data to the NBE showed a 1.1°C increase in annual temperature, which would predict a 208-m upslope movement of the ecotone, along with a 34% increase in precipitation. The rapid upward movement of the NBE indicates little inertia to climatically induced range shifts in montane forests; the upslope shift may have been accelerated by high turnover in canopy trees that provided opportunities for ingrowth of lower elevation species. Our results indicate that high-elevation forests may be jeopardized by climate change sooner than anticipated. PMID:18334647
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NASA Astrophysics Data System (ADS)
Stahle, Laura N.; Chin, Hahjung; Haberle, Simon; Whitlock, Cathy
2017-12-01
Fire activity was reconstructed at five sites and vegetation history at three sites in northwest Tasmania, Australia in order to examine the climate and human drivers of environmental change in the region. Watershed-scale reconstructions of fire were compared to regional vegetation history. Fire activity was very low until ca. 12,000 cal yr BP. An early-Holocene fire maximum, ca. 11,800-9800 cal yr BP, occurred during the warmest interval of the Holocene as recorded by regional paleoclimate proxy records. This period of elevated burning was also coincident with an increase in arboreal sclerophyll plant taxa. A maximum in rainforest taxa occurred at ca. 8500-5800 cal yr BP concurrent with sharply diminished biomass burning compared with the early Holocene. The increase in rainforest taxa is attributed to elevated effective moisture during this period. Conditions were drier and variable in the late Holocene as compared with earlier periods. A rise in fire activity at ca. 4800-3200 cal yr BP was accompanied by an increase in sclerophyll taxa and decline of rainforest and subalpine taxa. Elevated palynological richness during the late Holocene co-occurred with high levels of charcoal suggesting that fires promoted high floristic diversity. At Cradle Mountain, there is no clear evidence that fire regimes or vegetation were extensively modified by humans prior to European settlement. Climate was the primary driver of fire activity over millennial timescales as explained by the close relationship between charcoal and climate proxy data.
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NASA Astrophysics Data System (ADS)
Patterson, V. M.; Bormann, K.; Deems, J. S.; Painter, T. H.
2017-12-01
The NASA SnowEx campaign conducted in 2016 and 2017 provides a rich source of high-resolution Lidar data from JPL's Airborne Snow Observatory (ASO - http://aso.jpl.nasa.gov) combined with extensive in-situ measurements in two key areas in Colorado: Grand Mesa and Senator Beck. While the uncertainty in the 50m snow depth retrievals from NASA's ASO been estimated at 1-2cm in non-vegetated exposed areas (Painter et al., 2016), the impact of forest cover and point-cloud density on ASO snow lidar depth retrievals is relatively unknown. Dense forest canopies are known to reduce lidar penetration and ground strikes thus affecting the elevation surface retrieved from in the forest. Using high-resolution lidar point cloud data from the ASO SnowEx campaigns (26pt/m2) we applied a series of data decimations (up to 90% point reduction) to the point cloud data to quantify the relationship between vegetation, ground point density, resulting snow-off and snow-on surface elevations and finally snow depth. We observed non-linear reductions in lidar ground point density in forested areas that were strongly correlated to structural forest cover metrics. Previously, the impacts of these data decimations on a small study area in Grand Mesa showed a sharp increase in under-canopy surface elevation errors of -0.18m when ground point densities were reduced to 1.5pt/m2. In this study, we expanded the evaluation to the more topographically challenging Senator Beck basin, have conducted analysis along a vegetation gradient and are considering snow the impacts of snow depth rather than snow-off surface elevation. Preliminary analysis suggest that snow depth retrievals inferred from airborne lidar elevation differentials may systematically underestimate snow depth in forests where canopy density exceeds 1.75 and where tree heights exceed 5m. These results provide a basis from which to identify areas that may suffer from vegetation-induced biases in surface elevation models and snow depths derived from airborne lidar data, and help quantify expected spatial distributions of errors in the snow depth that can be used to improve the accuracy of ASO basin-scale depth and water equivalent products.
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Spruce aphid in high elevation habitats in the Southwest U.S.
Ann M. Lynch
2003-01-01
Spruce aphid, Elatobium abietinum (Walker) (Homoptera: Aphididae), is a new invasive pest in the interior Southwestern United States. This insect is causing extensive and severe damage on dormant Engelmann spruce, Picea engelmannii Parry, and Colorado blue spruce, P. pungens Engelm., in high elevation forests in...
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NASA Astrophysics Data System (ADS)
Borsato, Andrea; Frisia, Silvia; Miorandi, Renza
2015-04-01
Carbon dioxide concentration in soils controls carbonate dissolution, soil CO2 efflux to the atmosphere, and CO2 transfer to the subsurface that lead, ultimately, to speleothem precipitation. Systematic studies on CO2 concentration variability in soil and caves at regional scale are, however, few. Here, the systematic investigation of CO2 concentration in caves and soils in a temperate, Alpine region along a 2,100 m altitudinal range transect, which corresponds to a mean annual temperature (MAT) range of 12°C is presented. Soil pCO2 is controlled by the elevation and MAT and exhibits strong seasonality, which follows surface air temperature with a delay of about a month. The aquifer pCO2, by contrast, is fairly constant throughout the year, and it is primarily influenced by summer soil pCO2. Cave CO2 concentration is a balance between the CO2 influx and CO2 efflux, where the efflux is controlled by the cave ventilation, which is responsible for low pCO2 values recorded in most of the caves with respect to soil levels. Carbon dioxide in the innermost part of the studied caves exhibits a clear seasonal pattern. Thermal convection is the most common mechanism causing higher ventilation and low cave air pCO2 levels during the winter season: this promotes CO2 degassing and higher supersaturation in the drip water and, eventually, higher speleothem growth rates during winter. The combined influence of three parameters - dripwater pCO2, dripwater Ca content, and cave air pCO2 - all related to the infiltration elevation and MAT directly controls calcite supersaturation in dripwater. Four different altitudinal belts are then defined, which reflect temperature-dependent saturation state of dripwaters. These belts broadly correspond to vegetation zones: the lower montane (100 to 800 m asl), the upper montane (800 to 1600 m asl), the subalpine (1600 to 2200 m asl) and the Alpine (above 2200 m asl). Each altitudinal belt is characterised by different calcite fabrics, which can shift upward/downward in elevation as a response to temperature increase/decrease through time. In the lower and upper montane zones the columnar types (compact, open, fascicular optic) are the most common fabrics, with the microcrystalline type most typical of the upper montane zone. The dendritic fabric becomes predominant in the higher upper montane and lower subalpine zones. The higher subalpine to lower alpine zones the only speleothem actually forming is moonmilk. Eventually, the occurrence of "altitudinal" fabrics within the vertical growth axis of a stalagmite is indicative of changes in the MAT through time Therefore, fabric changes in fossil speleothems in temperate climate settings can be potentially used to reconstruct regional MAT changes in the past.
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Terrence J. Rogers
1996-01-01
This paper describes life histories, defoliation problems and other activities of insects associated with forest tree species growing along high elevation streams and river banks. In addition, examples of insects and diseases associated with lower elevation riparian areas are given.
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What mediates tree mortality during drought in the southern Sierra Nevada?
Paz-Kagan, Tarin; Brodrick, Philip; Vaughn, Nicholas R.; Das, Adrian J.; Stephenson, Nathan L.; Nydick, Koren R.; Asner, Gregory P.
2017-01-01
Severe drought has the potential to cause selective mortality within a forest, thereby inducing shifts in forest species composition. The southern Sierra Nevada foothills and mountains of California have experienced extensive forest dieback due to drought stress and insect outbreak. We used high-fidelity imaging spectroscopy (HiFIS) and light detection and ranging (LiDAR) from the Carnegie Airborne Observatory (CAO) to estimate the effect of forest dieback on species composition in response to drought stress in Sequoia National Park. Our aims were: (1) to quantify site-specific conditions that mediate tree mortality along an elevation gradient in the southern Sierra Nevada Mountains; (2) to assess where mortality events have a greater probability of occurring; and (3) to estimate which tree species have a greater likelihood of mortality along the elevation gradient. A series of statistical models were generated to classify species composition and identify tree mortality, and the influences of different environmental factors were spatially quantified and analyzed to assess where mortality events have a greater likelihood of occurring. A higher probability of mortality was observed in the lower portion of the elevation gradient, on southwest and west-facing slopes, in areas with shallow soils, on shallower slopes, and at greater distances from water. All of these factors are related to site water balance throughout the landscape. Our results also suggest that mortality is species-specific along the elevation gradient, mainly affecting Pinus ponderosa and Pinus lambertiana at lower elevations. Selective mortality within the forest may drive long-term shifts in community composition along the elevation gradient.
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What mediates tree mortality during drought in the southern Sierra Nevada?
Paz-Kagan, Tarin; Brodrick, Philip G; Vaughn, Nicholas R; Das, Adrian J; Stephenson, Nathan L; Nydick, Koren R; Asner, Gregory P
2017-12-01
Severe drought has the potential to cause selective mortality within a forest, thereby inducing shifts in forest species composition. The southern Sierra Nevada foothills and mountains of California have experienced extensive forest dieback due to drought stress and insect outbreak. We used high-fidelity imaging spectroscopy (HiFIS) and light detection and ranging (LiDAR) from the Carnegie Airborne Observatory (CAO) to estimate the effect of forest dieback on species composition in response to drought stress in Sequoia National Park. Our aims were (1) to quantify site-specific conditions that mediate tree mortality along an elevation gradient in the southern Sierra Nevada Mountains, (2) to assess where mortality events have a greater probability of occurring, and (3) to estimate which tree species have a greater likelihood of mortality along the elevation gradient. A series of statistical models were generated to classify species composition and identify tree mortality, and the influences of different environmental factors were spatially quantified and analyzed to assess where mortality events have a greater likelihood of occurring. A higher probability of mortality was observed in the lower portion of the elevation gradient, on southwest- and west-facing slopes, in areas with shallow soils, on shallower slopes, and at greater distances from water. All of these factors are related to site water balance throughout the landscape. Our results also suggest that mortality is species-specific along the elevation gradient, mainly affecting Pinus ponderosa and Pinus lambertiana at lower elevations. Selective mortality within the forest may drive long-term shifts in community composition along the elevation gradient. © 2017 by the Ecological Society of America.
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Morphological and functional diversity of ectomycorrhizal fungi on Roan Mountain (NC/TN)
Claire Bird; Coleman McCleneghan
2005-01-01
A comparison of ectomycorrhizal morphotypes and hypogeous fungi (truffles and false-truffles) in northern hardwood and spruce-fir forests on Roan Mountain (NC/TN) was performed to increase our knowledge of the fungal communities in the Southern Appalachian high elevation forests. These forests are home to an endangered subspecies and mycophagist, the Carolina northern...
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Spruce budworm returns to Northeast
Lloyd Irland; William H. McWilliams
2014-01-01
Thinking of the Northern Forest brings to mind spruce/fir (S/F) forests, cool climates, and high elevations: not to mention fishing and canoe trips: however, spruce and fir are also very important to the northern timber economy and rural development. Considering new concerns over the spruce budworm, an update on the status of this critically important forest resource...
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A case study of nitrogen saturation in western U.S. forests
Mark E. Fenn; Mark A. Poth
2001-01-01
Virtually complete nitrification of the available ammonium in soil and nitrification activity in the forest floor are important factors predisposing forests in the San Bernardino Mountains of southern California to nitrogen (N) saturation. As a result, inorganic N in the soil solution is dominated by nitrate. High nitrification rates also generate elevated nitric oxide...
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Forest reproduction along a climatic gradient in the Sierra Nevada, California
van Mantgem, Phillip J.; Stephenson, Nathan L.; Keeley, Jon E.
2006-01-01
To elucidate broad-scale environmental controls of coniferous forest reproduction in the Sierra Nevada, California, we monitored reproduction for 5 years in 47 plots arrayed across a steep elevational (climatic) gradient. We found that both absolute seedling densities (stems < 1.37 m) and seedling densities relative to overstory parent tree basal area declined sharply with elevation. Rates of seedling turnover (the average of birth and death rates) also declined with elevation. In contrast, seed production was not predicted by elevation and was highly variable from year to year. During a mast year of seed production, the intensity of masting was uneven among plots. Seedling densities were elevated only during the single year immediately following the mast year, suggesting reproduction in our forests may be primarily limited by abiotic factors such as the availability of suitable sites and weather. Disturbance also clearly affected reproduction; plots that had recently burned had significantly higher seedling to parent tree ratios for Abies species, suggesting that even though established Abies concolor may be relatively susceptible to fire, the species can recover rapidly through prolific reproduction. Since reproductive failures may be our earliest signal of changing forest conditions, seedling dynamics could provide a sensitive, if variable, indicator of environmental changes.
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Mark E. Fenn; Jeremy S. Fried; Haiganoush K. Preisler; Andrzej Bytnerowicz; Susan Schilling; Sarah Jovan; Olaf Kuegler
2015-01-01
The air in Californiaâs forests spans a broad range of purity, from virtually no locally generated pollutants to highly elevated levels of pollutants in forests downwind of urban and agricultural source areas. Ten-year remeasurement data from Forest Inventory and Analysis (FIA) plots in California were used in combination with modelled atmospheric nitrogen (N)...
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Paul G. Scowcroft; Jack Jeffrey
1999-01-01
Beginning in the 1850s, logging, land clearing, and burning were used to convert high elevation Hawaiian forests to cattle pasture. Recently, declining pro®ts from ranching, the need to expand habitat for endangered species, and diminishing supplies of native saw-timber have prompted interest in restoring native forests. The Forest Service, in cooperation with the...
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NASA Technical Reports Server (NTRS)
Ranson, Jon K.; Kovacs, Katalin; Kharuk, Viatcheslav; Burke, Erin
2006-01-01
Fires are a common occurrence in the Siberian boreal forest. The MOD14 Thermal anomalies product of the Terra MODIS Moderate Resolution Spectroradiometer) product set is designed to detect thermal anomalies (i.e. hotspots or fires) on the Earth's surface. Recent field studies showed a dependence of fire occurrence on topography. In this study MODIS thermal anomaly data and SRTM topography data were merged and analyzed to evaluate if forest fires are more likely to occur at certain combinations of elevation, slope and aspect. Using the satellite data over a large area can lead to better understanding how topography and forest fires are related. The study area covers a 2.5 Million krn(exp 2) portion of the Central Siberian southern taiga from 72 deg to 110 deg East and from 50 deg to 60 deg North. About 57% of the study area is forested and 80% of the forest grows between 200 and 1000 m. Forests with pine (Pinus sylvestris), larch (Larix sibirica, L. gmelinii), Siberian pine (Pinus sibirica), spruce (Picea obovata.) and fir (Abies sibirica) cover most of the landscape. Deciduous stands with birch (Betula pendula, B. pubescens) and aspen (Populus tremula) cover the areas of lower elevation in this region. The climate of this area is distinctly continental with long, cold winters and short hot summers. The tree line in this part of the world is around 1500 m in elevation with alpine tundra, snow and ice fields and rock outcrops extending up to over 3800 m. A 500 m resolution landcover map was developed using 2001 MODIS MOD13 Normalized Vegetation Index (NDVI) and Middle Infrared (MIR) products for seven 16-day periods. The classification accuracy was over 87%. The SRTM version 2 data, which is distributed in 1 degree by 1 degree tiles were mosaiced using the ENVI software. In this study, only those MODIS pixels were used that were flagged as "nominal or high confidence fire" by the MODIS fire product team. Using MODIS data from the years 2000 to 2005 along with the improved Shuttle Radar Topographic Mission (SRTM) version 2 data at 100 m resolution, the distribution of hot spots was examined by elevation, slope and aspect as well as by forest type. The results show that more forest area burns at lower elevations but a larger percentage of the available forest area burns at higher elevations. This is probably because steep slopes occur at higher elevations. Fires are only more common on slopes with a southern exposure if the slope is steeper than 15 degrees. The next step in this study will be to monitor areas where the risk of fire is high (steep slopes with a southern exposure) and to refine this method by incorporating anthropogenic features for more accurate fire disturbance monitoring.
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Simulation of Longwave Enhancement beneath Montane and Boreal Forests in CLM4.5
NASA Astrophysics Data System (ADS)
Todt, M.; Rutter, N.; Fletcher, C. G.; Wake, L. M.; Loranty, M. M.
2017-12-01
CMIP5 models have been shown to underestimate both trend and variability in northern hemisphere spring snow cover extent. A substantial fraction of this area is covered by boreal forests, in which the snow energy balance is dominated by radiation. Forest coverage impacts the surface radiation budget by shading the ground and enhancing longwave radiation. Longwave enhancement in boreal forests is a potential mechanism that contributes to uncertainty in snowmelt modelling, however, its impact on snowmelt in global land models has not been analysed yet. This study assesses the simulation of sub-canopy longwave radiation and longwave enhancement by CLM4.5, the land component of the NCAR Community Earth System Model, in which boreal forests are represented by three plant functional types (PFT): evergreen needleleaf trees (ENT), deciduous needleleaf trees (DNT), and deciduous broadleaf trees (DBT). Simulation of sub-canopy longwave enhancement is evaluated at boreal forest sites covering the three boreal PFT in CLM4.5 to assess the dependence of simulation errors on meteorological forcing, vegetation type and vegetation density. ENT are evaluated over a total of six snowmelt seasons in Swiss alpine and subalpine forests, as well as a single season at a Finnish arctic site with varying vegetation density. A Swedish artic site features varying vegetation density for DBT for a single winter, and two sites in Eastern Siberia are included covering a total of four snowmelt seasons in DNT forests. CLM4.5 overestimates the diurnal range of sub-canopy longwave radiation and consequently longwave enhancement, overestimating daytime values and underestimating nighttime values. Simulation errors result mainly from clear sky conditions, due to high absorption of shortwave radiation during daytime and radiative cooling during nighttime. Using recent improvements to the canopy parameterisations of SNOWPACK as a guideline, CLM4.5 simulations of sub-canopy longwave radiation improved through the implementation of a heat mass parameterisation, i.e. including thermal inertia due to biomass. However, this improvement does not substantially reduce the amplitude of the diurnal cycle, a result also found during the development of SNOWPACK.
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Adams, Henry D; Kolb, Thomas E
2004-07-01
We sought to understand differences in tree response to meteorological drought among species and soil types at two ecotone forests in northern Arizona, the pinyon-juniper woodland/ponderosa pine ecotone, and the higher elevation, wetter, ponderosa pine/mixed conifer ecotone. We used two approaches that provide different information about drought response: the ratio of standardized radial growth in wet years to dry years (W:D) for the period between years 1950 and 2000 as a measure of growth response to drought, and delta13C in leaves formed in non-drought (2001) and drought (2002) years as a measure of change in water use efficiency (WUE) in response to drought. W:D and leaf delta13C response to drought for Pinus edulis and P. ponderosa did not differ for trees growing on coarse-texture soils derived from cinders compared with finer textured soils derived from flow basalts or sedimentary rocks. P. ponderosa growing near its low elevation range limit at the pinyon-juniper woodland/ponderosa pine ecotone had a greater growth response to drought (higher W:D) and a larger increase in WUE in response to drought than co-occurring P. edulis growing near its high elevation range limit. P. flexilis and Pseudotsuga menziesii growing near their low elevation range limit at the ponderosa pine/mixed conifer ecotone had a larger growth response to drought than co-occurring P. ponderosa growing near its high elevation range limit. Increases in WUE in response to drought were similar for all species at the ponderosa pine/mixed conifer ecotone. Low elevation populations of P. ponderosa had greater growth response to drought than high-elevation populations, whereas populations had a similar increase in WUE in response to drought. Our findings of different responses to drought among co-occurring tree species and between low- and high-elevation populations are interpreted in the context of drought impacts on montane coniferous forests of the southwestern USA.
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Surface elevation dynamics in a regenerating mangrove forest at Homebush Bay, Australia
Rogers, K.; Saintilan, N.; Cahoon, D.
2005-01-01
Following the dieback of an interior portion of a mangrove forest at Homebush Bay, Australia, surface elevation tables and feldspar marker horizons were installed in the impacted, intermediate and control forest to measure vertical accretion, elevation change, and shallow subsidence. The objectives of the study were to determine current vertical accretion and elevation change rates as a guide to understanding mangrove dieback, ascertain the factors controlling surface elevation change, and investigate the sustainability of the mangrove forest under estimated sea-level rise conditions. The study demonstrates that the influences on surface dynamics are more complex than soil accretion and soil autocompaction alone. During strong vegetative regrowth in the impacted forest, surface elevation increase exceeded vertical accretion apparently as a result of belowground biomass production. In addition, surface elevation in all forest zones was correlated with total monthly rainfall during a severe El Ni?o event, highlighting the importance of rainfall to groundwater recharge and surface elevation. Surface elevation increase for all zones exceeded the 85-year sea level trend for Sydney Harbour. Since mean sea-level also decreased during the El Ni?o event, the decrease in surface elevation did not translate to an increase in inundation frequency or influence the sustainability of the mangrove forest. These findings indicate that subsurface soil processes such as organic matter accumulation and groundwater flux can significantly influence mangrove surface elevation, and contribute to the long-term sustainability of mangrove systems under a scenario of rising sea levels.
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Fire-induced erosion and millennial-scale climate change in northern ponderosa pine forests.
Pierce, Jennifer L; Meyer, Grant A; Jull, A J Timothy
2004-11-04
Western US ponderosa pine forests have recently suffered extensive stand-replacing fires followed by hillslope erosion and sedimentation. These fires are usually attributed to increased stand density as a result of fire suppression, grazing and other land use, and are often considered uncharacteristic or unprecedented. Tree-ring records from the past 500 years indicate that before Euro-American settlement, frequent, low-severity fires maintained open stands. However, the pre-settlement period between about ad 1500 and ad 1900 was also generally colder than present, raising the possibility that rapid twentieth-century warming promoted recent catastrophic fires. Here we date fire-related sediment deposits in alluvial fans in central Idaho to reconstruct Holocene fire history in xeric ponderosa pine forests and examine links to climate. We find that colder periods experienced frequent low-severity fires, probably fuelled by increased understory growth. Warmer periods experienced severe droughts, stand-replacing fires and large debris-flow events that comprise a large component of long-term erosion and coincide with similar events in sub-alpine forests of Yellowstone National Park. Our results suggest that given the powerful influence of climate, restoration of processes typical of pre-settlement times may be difficult in a warmer future that promotes severe fires.
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Huang, Minyi; Duan, Renyan; Wang, Shixiong; Wang, Zhigao; Fan, Weiyi
2016-01-01
Forest communities are mosaic systems composed of patches classified into four different developmental patch types: gap patch (G), building patch (B), mature patch (M) and degenerate patch (D). To study the mechanisms maintaining diversity in subalpine coniferous forests, species presence frequency and diversity in the four distinct patch types (G, B, M and D) of Larix chinensis conifer forests at three altitudinal gradients in the Qinling Mountains were analyzed. Our results were as follows: (1) Different species (or functional groups) had distinct presence frequencies in the four different patch types along the altitudinal gradient; (2) Some species or functional groups (species groups sharing similar traits and responses to the environment) only occurred in some specific patches. For seed dispersal, species using wind mainly occurred in G and D, while species using small animals mainly occurred in B and M; (3) Species composition of adjacent patch types was more similar than non-adjacent patch types, based on the lower β diversity index of the former; (4) The maximum numbers of species and two diversity indices (D' and H') were found in the middle altitudes. Various gap-forming processes and dispersal limitation may be the two major mechanisms determining species diversity in Larix chinensis coniferous forests at the patch scale.
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Dynamics and pattern of a managed coniferous forest landscape in Oregon
NASA Technical Reports Server (NTRS)
Spies, Thomas A.; Ripple, William J.; Bradshaw, G. A.
1995-01-01
We examined the process of fragmentation in a managed forest landscape by comparing rates and patterns of disturbance (primarily clear-cutting) and regrowth between 1972 and 1988 using Landsat imagery. A 2589-km(exp 2) managed forest landscape in western Oregon was classified into two forest types, closed-canopy conifer forest (CF) (typically, greater than 60% conifer cover) and other forest and nonforest types (OT) (typically, less than 40 yr old or deciduous forest). The percentage of CF declined from 71 to 58% between 1972 and 1988. Declines were greatest on private land, least in wilderness, and intermediate in public nonwilderness. High elevations (greater than 914 m) maintained a greater percentage of CF than lower elevations (less than 914 m). The percentage of the area at the edge of the two cover types increased on all ownerships and in both elevational zones, whereas the amount of interior habitat (defined as CF at least 100 m from OT) decreased on all ownerships and elevational zones. By 1988 public lands contained approximately 45% interior habitat while private lands had 12% interior habitat. Mean interior patch area declined from 160 to 62 ha. The annual rate of disturbance (primarily clear-cutting) for the entire area including the wilderness was 1.19%, which corresponds to a cutting rotation of 84 yr. The forest landscape was not in a steady state or regulated condition which is not projected to occur for at least 40 yr under current forest plans. Variability in cutting rates within ownerships was higher on private land than on nonreserve public land. However, despite the use of dispersed cutting patterns on public land, spatial patterns of cutting and remnant forest patches were nonuniform across the entire public ownership. Large remaining patches (less than 5000 ha) of contiguous interior forest were restricted to public lands designated for uses other than timber production such as wilderness areas and research natural areas.
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NASA Astrophysics Data System (ADS)
Wason, J. W., III; Dovciak, M.; Bevilacqua, E.
2015-12-01
Climate change in the northeastern United States is expected to shift climatic (temperature) envelopes for spruce-fir forests upslope and northward decreasing their area in the region by 2100. Coarse scale landscape models however, may not incorporate heterogeneity in climatic conditions in mountains that can create climatic refugia for species in high-elevation spruce-fir forests. To determine spatial and temporal trends in climate of mountain spruce-fir forests we measured microclimate at 98 forest plots in 2012 and 2013 on 12 mountains in New York, Vermont, New Hampshire, and Maine. By linking regional climate trends with our spatial climate data we calculated elevational shifts in temperature envelopes during the last 50 years. Additionally we linked our spatial dataset to a range of future climate conditions for 2100 based on Representative Concentration Pathways (1 to 5°C warming). We hypothesized that climates have already changed to an extent that spruce-fir forests should begin to respond and that future climate conditions may shift suitable habitat for spruce-fir forests beyond their current range. We found that regional climate change over the last 50 years has resulted in warming of 0.66 and 1.62°C for average annual daily maximum (Tmax) and minimum (Tmin) temperatures in the region. When linked to our spatial microclimate model, this warming results in a 100 (Tmax) and 312m (Tmin) upslope shift in temperature envelopes. Future climate projections suggest that by 2100 Tmax may shift upslope between 152 and 758m for the 1 and 5°C scenarios respectively, while Tmin may shift upslope between 192 and 962m. Spruce-fir forests typically occupy an elevation range of ~500m suggesting that the climate experienced in these forests 50 years ago may not be found within their elevation range by 2100. These results are discussed in the context of responses of tree populations and growth rates observed along the elevation gradients of northeastern United States.
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Muths, Erin L.; Scherer, R. D.; Amburgey, S. M.; Matthews, T.; Spencer, A. W.; Corn, P.S.
2016-01-01
In an era of shrinking budgets yet increasing demands for conservation, the value of existing (i.e., historical) data are elevated. Lengthy time series on common, or previously common, species are particularly valuable and may be available only through the use of historical information. We provide first estimates of the probability of survival and longevity (0.67–0.79 and 5–7 years, respectively) for a subalpine population of a small-bodied, ostensibly common amphibian, the Boreal Chorus Frog (Pseudacris maculata (Agassiz, 1850)), using historical data and contemporary, hypothesis-driven information–theoretic analyses. We also test a priori hypotheses about the effects of color morph (as suggested by early reports) and of drought (as suggested by recent climate predictions) on survival. Using robust mark–recapture models, we find some support for early hypotheses regarding the effect of color on survival, but we find no effect of drought. The congruence between early findings and our analyses highlights the usefulness of historical information in providing raw data for contemporary analyses and context for conservation and management decisions.
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Denning, A. Scott; Baron, Jill S.; Mast, M. Alisa; Arthur, Mary
1991-01-01
Intensive sampling of a stream draining an alpine-subalpine basin revealed that depressions in pH and acid neutralizing capacity (ANC) of surface water at the beginning of the spring snowmelt in 1987 and 1988 were not accompanied by increases in strong acid anions, and that surface waters did not become acidic (ANC<0). Samples of meltwater collected at the base of the snowpack in 1987 were acidic and exhibited distinct ‘pulses’ of nitrate and sulfate. Solutions collected with lysimeters in forest soils adjacent to the stream revealed high levels of dissolved organic carbon (DOC) and total Al. Peaks in concentration of DOC, Al, and nutrient species in the stream samples indicate a flush of soil solution into the surface water at the beginning of the melt. Infiltration of meltwater into soils and spatial heterogeneity in the timing of melting across the basin prevented stream and lake waters from becoming acidic.
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Diverse Soil Carbon Dynamics Expressed at the Molecular Level
NASA Astrophysics Data System (ADS)
van der Voort, T. S.; Zell, C. I.; Hagedorn, F.; Feng, X.; McIntyre, C. P.; Haghipour, N.; Graf Pannatier, E.; Eglinton, T. I.
2017-12-01
The stability and potential vulnerability of soil organic matter (SOM) to global change remain incompletely understood due to the complex processes involved in its formation and turnover. Here we combine compound-specific radiocarbon analysis with fraction-specific and bulk-level radiocarbon measurements in order to further elucidate controls on SOM dynamics in a temperate and subalpine forested ecosystem. Radiocarbon contents of individual organic compounds isolated from the same soil interval generally exhibit greater variation than those among corresponding operationally defined fractions. Notably, markedly older ages of long-chain plant leaf wax lipids (n-alkanoic acids) imply that they reflect a highly stable carbon pool. Furthermore, marked 14C variations among shorter- and longer-chain n-alkanoic acid homologues suggest that they track different SOM pools. Extremes in SOM dynamics thus manifest themselves within a single compound class. This exploratory study highlights the potential of compound-specific radiocarbon analysis for understanding SOM dynamics in ecosystems potentially vulnerable to global change.