Thompson, Laura A.; Romano, Tracy A.
2016-01-01
Increased pressure, associated with diving, can alter cell function through several mechanisms and has been shown to impact immune functions performed by peripheral blood mononuclear cells (PBMC) in humans. While marine mammals possess specific adaptations which protect them from dive related injury, it is unknown how their immune system is adapted to the challenges associated with diving. The purpose of this study was to measure PBMC activation (IL2R expression) and Concanavalin A induced lymphocyte proliferation (BrdU incorporation) in belugas following in vitro pressure exposures during baseline, Out of Water Examination (OWE) and capture/release conditions. Beluga blood samples (n = 4) were obtained from animals at the Mystic Aquarium and from free ranging animals in Alaska (n = 9). Human blood samples (n = 4) (Biological Specialty Corporation) were run for comparison. In vivo catecholamines and cortisol were measured in belugas to characterize the neuroendocrine response. Comparison of cellular responses between controls and pressure exposed cells, between conditions in belugas, between belugas and humans as well as between dive profiles, were run using mixed generalized linear models (α = 0.05). Cortisol was significantly higher in Bristol Bay belugas and OWE samples as compared with baseline for aquarium animals. Both IL2R expression and proliferation displayed significant pressure induced changes, and these responses varied between conditions in belugas. Both belugas and humans displayed increased IL2R expression, while lymphocyte proliferation decreased for aquarium animals and increased for humans and Bristol Bay belugas. Results suggest beluga PBMC function is altered during diving and changes may represent dive adaptation as the response differs from humans, a non-dive adapted mammal. In addition, characteristics of a dive (i.e., duration, depth) as well as neuroendocrine activity can alter the response of beluga cells, potentially impacting the ability of animals to fight infection or avoid dive related pathologies. PMID:27746745
Thompson, Laura A; Romano, Tracy A
2016-01-01
Increased pressure, associated with diving, can alter cell function through several mechanisms and has been shown to impact immune functions performed by peripheral blood mononuclear cells (PBMC) in humans. While marine mammals possess specific adaptations which protect them from dive related injury, it is unknown how their immune system is adapted to the challenges associated with diving. The purpose of this study was to measure PBMC activation (IL2R expression) and Concanavalin A induced lymphocyte proliferation (BrdU incorporation) in belugas following in vitro pressure exposures during baseline, Out of Water Examination (OWE) and capture/release conditions. Beluga blood samples ( n = 4) were obtained from animals at the Mystic Aquarium and from free ranging animals in Alaska ( n = 9). Human blood samples ( n = 4) (Biological Specialty Corporation) were run for comparison. In vivo catecholamines and cortisol were measured in belugas to characterize the neuroendocrine response. Comparison of cellular responses between controls and pressure exposed cells, between conditions in belugas, between belugas and humans as well as between dive profiles, were run using mixed generalized linear models (α = 0.05). Cortisol was significantly higher in Bristol Bay belugas and OWE samples as compared with baseline for aquarium animals. Both IL2R expression and proliferation displayed significant pressure induced changes, and these responses varied between conditions in belugas. Both belugas and humans displayed increased IL2R expression, while lymphocyte proliferation decreased for aquarium animals and increased for humans and Bristol Bay belugas. Results suggest beluga PBMC function is altered during diving and changes may represent dive adaptation as the response differs from humans, a non-dive adapted mammal. In addition, characteristics of a dive (i.e., duration, depth) as well as neuroendocrine activity can alter the response of beluga cells, potentially impacting the ability of animals to fight infection or avoid dive related pathologies.
Diving bradycardia: a mechanism of defence against hypoxic damage.
Alboni, Paolo; Alboni, Marco; Gianfranchi, Lorella
2011-06-01
A feature of all air-breathing vertebrates, diving bradycardia is triggered by apnoea and accentuated by immersion of the face or whole body in cold water. Very little is known about the afferents of diving bradycardia, whereas the efferent part of the reflex circuit is constituted by the cardiac vagal fibres. Diving bradycardia is associated with vasoconstriction of selected vascular beds and a reduction in cardiac output. The diving response appears to be more pronounced in mammals than in birds. In humans, the bradycardic response to diving varies greatly from person to person; the reduction in heart rate generally ranges from 15 to 40%, but a small proportion of healthy individuals can develop bradycardia below 20 beats/min. During prolonged dives, bradycardia becomes more pronounced because of activation of the peripheral chemoreceptors by a reduction in the arterial partial pressure of oxygen (O2), responsible for slowing of heart rate. The vasoconstriction is associated with a redistribution of the blood flow, which saves O2 for the O2-sensitive organs, such as the heart and brain. The results of several investigations carried out both in animals and in humans show that the diving response has an O2-conserving effect, both during exercise and at rest, thus lengthening the time to the onset of serious hypoxic damage. The diving response can therefore be regarded as an important defence mechanism for the organism.
Genetic determination of the vascular reactions in humans in response to the diving reflex.
Baranova, Tatiana I; Berlov, Dmitrii N; Glotov, Oleg S; Korf, Ekaterina A; Minigalin, Alexey D; Mitrofanova, Alla V; Ahmetov, Ildus I; Glotov, Andrey S
2017-03-01
The purpose of this study was to investigate the genetic mechanisms of the defense vascular reactions in response to the diving reflex in humans with polymorphisms in the genes ADBR2, ACE, AGTR1, BDKRB2 , and REN We hypothesized that protective vascular reactions, in response to the diving reflex, are genetically determined and are distinguished in humans with gene polymorphisms of the renin-angiotensin and kinin-bradykinin system. A total of 80 subjects (19 ± 1.4 yr) participated in the study. The intensity of the vascular response was estimated using photoplethysmogram. The I/D polymorphism (rs4340) of ACE was analyzed by PCR. REN (G/A, rs2368564), AGTR1 (A/C, rs5186), BDKRB2 (T/C, rs1799722), and ADBR2 (A/G, rs1042713) polymorphisms were examined using the two-step multiplex PCR followed by carrying allele hybridization on the biochip. Subjects with the BDKRB2 (C/C), ACE (D/D), and ADBR2 (G/G, G/A) genotypes exhibited the strongest peripheral vasoconstriction in response to diving. In subjects with a combination of the BDKRB2 (C/C) plus ACE (D/D) genotypes, we observed the lowest pulse wave amplitude and pulse transit time values and the highest arterial blood pressure during face immersion compared with the heterozygous individuals, suggesting that these subjects are more susceptible to diving hypoxia. This study observed that humans with gene polymorphisms of the renin-angiotensin and kinin-bradykinin systems demonstrate various expressions of protective vascular reactions in response to the diving reflex. The obtained results might be used in estimation of resistance to hypoxia of any origin in human beings or in a medical practice. NEW & NOTEWORTHY Our study demonstrates that the vascular reactions in response to the diving reflex are genetically determined and depend on gene polymorphisms of the kinin-bradykinin and the renin-angiotensin systems. Copyright © 2017 the American Physiological Society.
Thompson, Laura A.; Romano, Tracy A.
2015-01-01
While it is widely known that marine mammals possess adaptations which allow them to make repetitive and extended dives to great depths without suffering ill effects seen in humans, the response of marine mammal immune cells to diving is unknown. Renewed interest in marine mammal dive physiology has arisen due to reports of decompression sickness-like symptoms and embolic damage in stranded and by-caught animals, and there is concern over whether anthropogenic activities can impact marine mammal health by disrupting adaptive dive responses and behavior. This work addresses the need for information concerning marine mammal immune function during diving by evaluating granulocyte and monocyte phagocytosis, and granulocyte activation in belugas (n = 4) in comparison with humans (n = 4), with and without in vitro pressure exposures. In addition, the potential for additional stressors to impact immune function was investigated by comparing the response of beluga cells to pressure between baseline and stressor conditions. Granulocyte and monocyte phagocytosis, as well as granulocyte activation, were compared between pressure exposed and non-exposed cells for each condition, between different pressure profiles and between conditions using mixed generalized linear models (α = 0.05). The effects of pressure varied between species as well by depth, compression/decompression rates, and length of exposures, and condition for belugas. Pressure induced changes in granulocyte and monocyte function in belugas could serve a protective function against dive-related pathologies and differences in the response between humans and belugas could reflect degrees of dive adaptation. The alteration of these responses during physiologically challenging conditions may increase the potential for dive-related in jury and disease in marine mammals. PMID:25999860
Thompson, Laura A; Romano, Tracy A
2015-01-01
While it is widely known that marine mammals possess adaptations which allow them to make repetitive and extended dives to great depths without suffering ill effects seen in humans, the response of marine mammal immune cells to diving is unknown. Renewed interest in marine mammal dive physiology has arisen due to reports of decompression sickness-like symptoms and embolic damage in stranded and by-caught animals, and there is concern over whether anthropogenic activities can impact marine mammal health by disrupting adaptive dive responses and behavior. This work addresses the need for information concerning marine mammal immune function during diving by evaluating granulocyte and monocyte phagocytosis, and granulocyte activation in belugas (n = 4) in comparison with humans (n = 4), with and without in vitro pressure exposures. In addition, the potential for additional stressors to impact immune function was investigated by comparing the response of beluga cells to pressure between baseline and stressor conditions. Granulocyte and monocyte phagocytosis, as well as granulocyte activation, were compared between pressure exposed and non-exposed cells for each condition, between different pressure profiles and between conditions using mixed generalized linear models (α = 0.05). The effects of pressure varied between species as well by depth, compression/decompression rates, and length of exposures, and condition for belugas. Pressure induced changes in granulocyte and monocyte function in belugas could serve a protective function against dive-related pathologies and differences in the response between humans and belugas could reflect degrees of dive adaptation. The alteration of these responses during physiologically challenging conditions may increase the potential for dive-related in jury and disease in marine mammals.
DiNovo, Karyn. M.; Connolly, Tiffanny M.
2010-01-01
The mammalian diving response, consisting of apnea, bradycardia, and increased total peripheral resistance, can be modified by conscious awareness, fear, and anticipation. We wondered whether swim and dive training in rats would 1) affect the magnitude of the cardiovascular responses during voluntary and forced diving, and 2) whether this training would reduce or eliminate any stress due to diving. Results indicate Sprague-Dawley rats have a substantial diving response. Immediately upon submersion, heart rate (HR) decreased by 78%, from 453 ± 12 to 101 ± 8 beats per minute (bpm), and mean arterial pressure (MAP) decreased 25%, from 143 ± 1 to 107 ± 5 mmHg. Approximately 4.5 s after submergence, MAP had increased to a maximum 174 ± 3 mmHg. Blood corticosterone levels indicate trained rats find diving no more stressful than being held by a human, while untrained rats find swimming and diving very stressful. Forced diving is stressful to both trained and untrained rats. The magnitude of bradycardia was similar during both voluntary and forced diving, while the increase in MAP was greater during forced diving. The diving response of laboratory rats, therefore, appears to be dissimilar from that of other animals, as most birds and mammals show intensification of diving bradycardia during forced diving compared with voluntary diving. Rats may exhibit an accentuated antagonism between the parasympathetic and sympathetic branches of the autonomic nervous system, such that in the autonomic control of HR, parasympathetic activity overpowers sympathetic activity. Additionally, laboratory rats may lack the ability to modify the degree of parasympathetic outflow to the heart during an intense cardiorespiratory response (i.e., the diving response). PMID:19923359
NASA Astrophysics Data System (ADS)
Roche, Ronan C.; Harvey, Chloe V.; Harvey, James J.; Kavanagh, Alan P.; McDonald, Meaghan; Stein-Rostaing, Vivienne R.; Turner, John R.
2016-07-01
Recreational diving on coral reefs is an activity that has experienced rapidly growing levels of popularity and participation. Despite providing economic activity for many developing coastal communities, the potential role of dive impacts in contributing to coral reef damage is a concern at heavily dived locations. Management measures to address this issue increasingly include the introduction of programmes designed to encourage environmentally responsible practices within the dive industry. We examined diver behaviour at several important coral reef dive locations within the Philippines and assessed how diver characteristics and dive operator compliance with an environmentally responsible diving programme, known as the Green Fins approach, affected reef contacts. The role of dive supervision was assessed by recording dive guide interventions underwater, and how this was affected by dive group size. Of the 100 recreational divers followed, 88 % made contact with the reef at least once per dive, with a mean (±SE) contact rate of 0.12 ± 0.01 per min. We found evidence that the ability of dive guides to intervene and correct diver behaviour in the event of a reef contact decreases with larger diver group sizes. Divers from operators with high levels of compliance with the Green Fins programme exhibited significantly lower reef contact rates than those from dive operators with low levels of compliance. The successful implementation of environmentally responsible diving programmes, which focus on influencing dive industry operations, can contribute to the management of human impacts on coral reefs.
Roche, Ronan C; Harvey, Chloe V; Harvey, James J; Kavanagh, Alan P; McDonald, Meaghan; Stein-Rostaing, Vivienne R; Turner, John R
2016-07-01
Recreational diving on coral reefs is an activity that has experienced rapidly growing levels of popularity and participation. Despite providing economic activity for many developing coastal communities, the potential role of dive impacts in contributing to coral reef damage is a concern at heavily dived locations. Management measures to address this issue increasingly include the introduction of programmes designed to encourage environmentally responsible practices within the dive industry. We examined diver behaviour at several important coral reef dive locations within the Philippines and assessed how diver characteristics and dive operator compliance with an environmentally responsible diving programme, known as the Green Fins approach, affected reef contacts. The role of dive supervision was assessed by recording dive guide interventions underwater, and how this was affected by dive group size. Of the 100 recreational divers followed, 88 % made contact with the reef at least once per dive, with a mean (±SE) contact rate of 0.12 ± 0.01 per min. We found evidence that the ability of dive guides to intervene and correct diver behaviour in the event of a reef contact decreases with larger diver group sizes. Divers from operators with high levels of compliance with the Green Fins programme exhibited significantly lower reef contact rates than those from dive operators with low levels of compliance. The successful implementation of environmentally responsible diving programmes, which focus on influencing dive industry operations, can contribute to the management of human impacts on coral reefs.
Fitz-Clarke, John R
2018-03-25
Breath-hold diving is practiced by recreational divers, seafood divers, military divers, and competitive athletes. It involves highly integrated physiology and extreme responses. This article reviews human breath-hold diving physiology beginning with an historical overview followed by a summary of foundational research and a survey of some contemporary issues. Immersion and cardiovascular adjustments promote a blood shift into the heart and chest vasculature. Autonomic responses include diving bradycardia, peripheral vasoconstriction, and splenic contraction, which help conserve oxygen. Competitive divers use a technique of lung hyperinflation that raises initial volume and airway pressure to facilitate longer apnea times and greater depths. Gas compression at depth leads to sequential alveolar collapse. Airway pressure decreases with depth and becomes negative relative to ambient due to limited chest compliance at low lung volumes, raising the risk of pulmonary injury called "squeeze," characterized by postdive coughing, wheezing, and hemoptysis. Hypoxia and hypercapnia influence the terminal breakpoint beyond which voluntary apnea cannot be sustained. Ascent blackout due to hypoxia is a danger during long breath-holds, and has become common amongst high-level competitors who can suppress their urge to breathe. Decompression sickness due to nitrogen accumulation causing bubble formation can occur after multiple repetitive dives, or after single deep dives during depth record attempts. Humans experience responses similar to those seen in diving mammals, but to a lesser degree. The deepest sled-assisted breath-hold dive was to 214 m. Factors that might determine ultimate human depth capabilities are discussed. © 2018 American Physiological Society. Compr Physiol 8:585-630, 2018. Copyright © 2018 American Physiological Society. All rights reserved.
Using Stimulation of the Diving Reflex in Humans to Teach Integrative Physiology
ERIC Educational Resources Information Center
Choate, Julia K.; Denton, Kate M.; Evans, Roger G.; Hodgson, Yvonne
2014-01-01
During underwater submersion, the body responds by conserving O[subscript 2] and prioritizing blood flow to the brain and heart. These physiological adjustments, which involve the nervous, cardiovascular, and respiratory systems, are known as the diving response and provide an ideal example of integrative physiology. The diving reflex can be…
Using stimulation of the diving reflex in humans to teach integrative physiology.
Choate, Julia K; Denton, Kate M; Evans, Roger G; Hodgson, Yvonne
2014-12-01
During underwater submersion, the body responds by conserving O2 and prioritizing blood flow to the brain and heart. These physiological adjustments, which involve the nervous, cardiovascular, and respiratory systems, are known as the diving response and provide an ideal example of integrative physiology. The diving reflex can be stimulated in the practical laboratory setting using breath holding and facial immersion in water. Our undergraduate physiology students complete a laboratory class in which they investigate the effects of stimulating the diving reflex on cardiovascular variables, which are recorded and calculated with a Finapres finger cuff. These variables include heart rate, cardiac output, stroke volume, total peripheral resistance, and arterial pressures (mean, diastolic, and systolic). Components of the diving reflex are stimulated by 1) facial immersion in cold water (15°C), 2) breathing with a snorkel in cold water (15°C), 3) facial immersion in warm water (30°C), and 4) breath holding in air. Statistical analysis of the data generated for each of these four maneuvers allows the students to consider the factors that contribute to the diving response, such as the temperature of the water and the location of the sensory receptors that initiate the response. In addition to providing specific details about the equipment, protocols, and learning outcomes, this report describes how we assess this practical exercise and summarizes some common student misunderstandings of the essential physiological concepts underlying the diving response. Copyright © 2014 The American Physiological Society.
Effects of successive air and nitrox dives on human vascular function.
Marinovic, Jasna; Ljubkovic, Marko; Breskovic, Toni; Gunjaca, Grgo; Obad, Ante; Modun, Darko; Bilopavlovic, Nada; Tsikas, Dimitrios; Dujic, Zeljko
2012-06-01
SCUBA diving is regularly associated with asymptomatic changes in cardiac, pulmonary and vascular function. The aim of this study was to evaluate the changes in vascular/endothelial function following SCUBA diving and to assess the potential difference between two breathing gases: air and nitrox 36 (36% oxygen and 64% nitrogen). Ten divers performed two 3-day diving series (no-decompression dive to 18 m with 47 min bottom time with air and nitrox, respectively), with 2 weeks pause in between. Arterial/endothelial function was assessed using SphygmoCor and flow-mediated dilation measurements, and concentration of nitrite before and after diving was determined in venous blood. Production of nitrogen bubbles post-dive was assessed by ultrasonic determination of venous gas bubble grade. Significantly higher bubbling was found after all air dives as compared to nitrox dives. Pulse wave velocity increased slightly (~6%), significantly after both air and nitrox diving, indicating an increase in arterial stiffness. However, augmentation index became significantly more negative after diving indicating smaller wave reflection. There was a trend for post-dive reduction of FMD after air dives; however, only nitrox diving significantly reduced FMD. No significant differences in blood nitrite before and after the dives were found. We found that nitrox diving affects systemic/vascular function more profoundly than air diving by reducing FMD response, most likely due to higher oxygen load. Both air and nitrox dives increased arterial stiffness, but decreased wave reflection suggesting a decrease in peripheral resistance due to exercise during diving. These effects of nitrox and air diving were not followed by changes in plasma nitrite.
2016-03-21
Ocean Acoustics Human Effects Analysis NMFS National Marine Fisheries Service NOAA National Oceanic and Atmospheric Administration ODBA...dive basis, switch effectively between different diving states, and do so while maintaining foraging efficiency and social cohesion. Benefits...adverse effects on some species of odontocete cetaceans under certain environmental conditions. Some beaked whales appeared to be particularly sensitive
Decompression sickness in breath-hold divers: a review.
Lemaitre, Frederic; Fahlman, Andreas; Gardette, Bernard; Kohshi, Kiyotaka
2009-12-01
Although it has been generally assumed that the risk of decompression sickness is virtually zero during a single breath-hold dive in humans, repeated dives may result in a cumulative increase in the tissue and blood nitrogen tension. Many species of marine mammals perform extensive foraging bouts with deep and long dives interspersed by a short surface interval, and some human divers regularly perform repeated dives to 30-40 m or a single dive to more than 200 m, all of which may result in nitrogen concentrations that elicit symptoms of decompression sickness. Neurological problems have been reported in humans after single or repeated dives and recent necropsy reports in stranded marine mammals were suggestive of decompression sickness-like symptoms. Modelling attempts have suggested that marine mammals may live permanently with elevated nitrogen concentrations and may be at risk when altering their dive behaviour. In humans, non-pathogenic bubbles have been recorded and symptoms of decompression sickness have been reported after repeated dives to modest depths. The mechanisms implicated in these accidents indicate that repeated breath-hold dives with short surface intervals are factors that predispose to decompression sickness. During deep diving, the effect of pulmonary shunts and/or lung collapse may play a major role in reducing the incidence of decompression sickness in humans and marine mammals.
Cardiac changes induced by immersion and breath-hold diving in humans.
Marabotti, Claudio; Scalzini, Alessandro; Cialoni, Danilo; Passera, Mirko; L'Abbate, Antonio; Bedini, Remo
2009-01-01
To evaluate the separate cardiovascular response to body immersion and increased environmental pressure during diving, 12 healthy male subjects (mean age 35.2 +/- 6.5 yr) underwent two-dimensional Doppler echocardiography in five different conditions: out of water (basal); head-out immersion while breathing (condition A); fully immersed at the surface while breathing (condition B) and breath holding (condition C); and breath-hold diving at 5-m depth (condition D). Heart rate, left ventricular volumes, stroke volume, and cardiac output were obtained by underwater echocardiography. Early (E) and late (A) transmitral flow velocities, their ratio (E/A), and deceleration time of E (DTE) were also obtained from pulsed-wave Doppler, as left ventricular diastolic function indexes. The experimental protocol induced significant reductions in left ventricular volumes, left ventricular stroke volume (P < 0.05), cardiac output (P < 0.001), and heart rate (P < 0.05). A significant increase in E peak (P < 0.01) and E/A (P < 0.01) and a significant reduction of DTE (P < 0.01) were also observed. Changes occurring during diving (condition D) accounted for most of the changes observed in the experimental series. In particular, cardiac output at condition D was significantly lower compared with each of the other experimental conditions, E/A was significantly higher during condition D than in conditions A and C. Finally, DTE was significantly shorter at condition D than in basal and condition C. This study confirms a reduction of cardiac output in diving humans. Since most of the changes were observed during diving, the increased environmental pressure seems responsible for this hemodynamic rearrangement. Left ventricular diastolic function changes suggest a constrictive effect on the heart, possibly accounting for cardiac output reduction.
Cole, A J; Griffiths, D; Lavender, S; Summers, P; Rich, K
2006-05-01
To test the hypothesis that artefact caused by postmortem off-gassing is at least partly responsible for the presence of gas within the vascular system and tissues of the cadaver following death associated with compressed air diving. Controlled experiment sacrificing sheep after a period of simulated diving in a hyperbaric chamber and carrying out sequential postmortem computed tomography (CT) on the cadavers. All the subject sheep developed significant quantities of gas in the vascular system within 24 hours, as demonstrated by CT and necropsy, while the control animals did not. The presence of gas in the vascular system of human cadavers following diving associated fatalities is to be expected, and is not necessarily connected with gas embolism following pulmonary barotrauma, as has previously been claimed.
The diving paradox: new insights into the role of the dive response in air-breathing vertebrates.
Davis, Randall W; Polasek, Lori; Watson, Rebecca; Fuson, Amanda; Williams, Terrie M; Kanatous, Shane B
2004-07-01
When aquatic reptiles, birds and mammals submerge, they typically exhibit a dive response in which breathing ceases, heart rate slows, and blood flow to peripheral tissues is reduced. The profound dive response that occurs during forced submergence sequesters blood oxygen for the brain and heart while allowing peripheral tissues to become anaerobic, thus protecting the animal from immediate asphyxiation. However, the decrease in peripheral blood flow is in direct conflict with the exercise response necessary for supporting muscle metabolism during submerged swimming. In free diving animals, a dive response still occurs, but it is less intense than during forced submergence, and whole-body metabolism remains aerobic. If blood oxygen is not sequestered for brain and heart metabolism during normal diving, then what is the purpose of the dive response? Here, we show that its primary role may be to regulate the degree of hypoxia in skeletal muscle so that blood and muscle oxygen stores can be efficiently used. Paradoxically, the muscles of diving vertebrates must become hypoxic to maximize aerobic dive duration. At the same time, morphological and enzymatic adaptations enhance intracellular oxygen diffusion at low partial pressures of oxygen. Optimizing the use of blood and muscle oxygen stores allows aquatic, air-breathing vertebrates to exercise for prolonged periods while holding their breath.
Cole, A J; Griffiths, D; Lavender, S; Summers, P; Rich, K
2006-01-01
Aims To test the hypothesis that artefact caused by postmortem off‐gassing is at least partly responsible for the presence of gas within the vascular system and tissues of the cadaver following death associated with compressed air diving. Methods Controlled experiment sacrificing sheep after a period of simulated diving in a hyperbaric chamber and carrying out sequential postmortem computed tomography (CT) on the cadavers. Results All the subject sheep developed significant quantities of gas in the vascular system within 24 hours, as demonstrated by CT and necropsy, while the control animals did not. Conclusions The presence of gas in the vascular system of human cadavers following diving associated fatalities is to be expected, and is not necessarily connected with gas embolism following pulmonary barotrauma, as has previously been claimed. PMID:16489175
The rat: a laboratory model for studies of the diving response
Gan, Qi; Juric, Rajko
2010-01-01
Underwater submersion in mammals induces apnea, parasympathetically mediated bradycardia, and sympathetically mediated peripheral vasoconstriction. These effects are collectively termed the diving response, potentially the most powerful autonomic reflex known. Although these physiological responses are directed by neurons in the brain, study of neural control of the diving response has been hampered since 1) it is difficult to study the brains of animals while they are underwater, 2) feral marine mammals are usually large and have brains of variable size, and 3) there are but few references on the brains of naturally diving species. Similar responses are elicited in anesthetized rodents after stimulation of their nasal mucosa, but this nasopharyngeal reflex has not been compared directly with natural diving behavior in the rat. In the present study, we compared hemodynamic responses elicited in awake rats during volitional underwater submersion with those of rats swimming on the water's surface, rats involuntarily submerged, and rats either anesthetized or decerebrate and stimulated nasally with ammonia vapors. We show that the hemodynamic changes to voluntary diving in the rat are similar to those of naturally diving marine mammals. We also show that the responses of voluntary diving rats are 1) significantly different from those seen during swimming, 2) generally similar to those elicited in trained rats involuntarily “dunked” underwater, and 3) generally different from those seen from dunking naive rats underwater. Nasal stimulation of anesthetized rats differed most from the hemodynamic variables of rats trained to dive voluntarily. We propose that the rat trained to dive underwater is an excellent laboratory model to study neural control of the mammalian diving response, and also suggest that some investigations may be done with nasal stimulation of decerebrate preparations to decipher such control. PMID:20093670
Effects of depth and chest volume on cardiac function during breath-hold diving.
Marabotti, Claudio; Scalzini, Alessandro; Cialoni, Danilo; Passera, Mirko; Ripoli, Andrea; L'Abbate, Antonio; Bedini, Remo
2009-07-01
Cardiac response to breath-hold diving in human beings is primarily characterized by the reduction of both heart rate and stroke volume. By underwater Doppler-echocardiography we observed a "restrictive/constrictive" left ventricular filling pattern compatible with the idea of chest squeeze and heart compression during diving. We hypothesized that underwater re-expansion of the chest would release heart constriction and normalize cardiac function. To this aim, 10 healthy male subjects (age 34.2 +/- 10.4) were evaluated by Doppler-echocardiography during breath-hold immersion at a depth of 10 m, before and after a single maximal inspiration from a SCUBA device. During the same session, all subjects were also studied at surface (full-body immersion) and at 5-m depth in order to better characterize the relationship of echo-Doppler pattern with depth. In comparison to surface immersion, 5-m deep diving was sufficient to reduce cardiac output (P = 0.042) and increase transmitral E-peak velocity (P < 0.001). These changes remained unaltered at a 10-m depth. Chest expansion at 10 m decreased left ventricular end-systolic volume (P = 0.024) and increased left ventricular stroke volume (P = 0.024). In addition, it decreased transmitral E-peak velocity (P = 0.012) and increased deceleration time of E-peak (P = 0.021). In conclusion the diving response, already evident during shallow diving (5 m) did not progress during deeper dives (10 m). The rapid improvement in systolic and diastolic function observed after lung volume expansion is congruous with the idea of a constrictive effect on the heart exerted by chest squeeze.
Responses of blind fish to gravitational changes as achieved in parabolic flight
NASA Technical Reports Server (NTRS)
Vonbaumgarten, R. J.; Shillinger, G. L., Jr.; Baldright, G.
1972-01-01
Blind fish, during parabolic flight, display a measurable and consistent behavior. The most spectacular new behavioral response is the forward looping of blind fish in or near weightlessness. This response shows no measurable adaptation during the entire period of weightlessness of about 30 sec. During the entrance and exit of weightless parabolas (pushover and pullout) respectively, the fish assumes a forward tilted diving position. Parabolic flight with negative g in the range between 0g and -1g causes similar diving responses of the fish with the only difference being that the dive is directed toward the top of the fish tank. When the response to a g value less than 1g is compared to the response to increased g load on the ground (escape of darting response) an essential difference is seen: higher horizontal acceleration or jerk on the ground causes fish to swim, or even dart, against the direction of inertial force; fish during weightless parabolas move into the direction of the inertial or gravitational force. Since the vestibular system of fish is homologous to that of man, the observed behavior of fish in weightless flight could help to better understand human performance and sensations in comparable situations.
Deadly diving? Physiological and behavioural management of decompression stress in diving mammals
Hooker, S. K.; Fahlman, A.; Moore, M. J.; Aguilar de Soto, N.; Bernaldo de Quirós, Y.; Brubakk, A. O.; Costa, D. P.; Costidis, A. M.; Dennison, S.; Falke, K. J.; Fernandez, A.; Ferrigno, M.; Fitz-Clarke, J. R.; Garner, M. M.; Houser, D. S.; Jepson, P. D.; Ketten, D. R.; Kvadsheim, P. H.; Madsen, P. T.; Pollock, N. W.; Rotstein, D. S.; Rowles, T. K.; Simmons, S. E.; Van Bonn, W.; Weathersby, P. K.; Weise, M. J.; Williams, T. M.; Tyack, P. L.
2012-01-01
Decompression sickness (DCS; ‘the bends’) is a disease associated with gas uptake at pressure. The basic pathology and cause are relatively well known to human divers. Breath-hold diving marine mammals were thought to be relatively immune to DCS owing to multiple anatomical, physiological and behavioural adaptations that reduce nitrogen gas (N2) loading during dives. However, recent observations have shown that gas bubbles may form and tissue injury may occur in marine mammals under certain circumstances. Gas kinetic models based on measured time-depth profiles further suggest the potential occurrence of high blood and tissue N2 tensions. We review evidence for gas-bubble incidence in marine mammal tissues and discuss the theory behind gas loading and bubble formation. We suggest that diving mammals vary their physiological responses according to multiple stressors, and that the perspective on marine mammal diving physiology should change from simply minimizing N2 loading to management of the N2 load. This suggests several avenues for further study, ranging from the effects of gas bubbles at molecular, cellular and organ function levels, to comparative studies relating the presence/absence of gas bubbles to diving behaviour. Technological advances in imaging and remote instrumentation are likely to advance this field in coming years. PMID:22189402
McCulloch, Paul F.; Warren, Erik A.; DiNovo, Karyn M.
2016-01-01
This research was designed to investigate the role of the anterior ethmoidal nerve (AEN) during repetitive trained diving in rats, with specific attention to activation of afferent and efferent brainstem nuclei that are part of this reflexive response. The AEN innervates the nose and nasal passages and is thought to be an important component of the afferent limb of the diving response. Male Sprague-Dawley rats (N = 24) were trained to swim and dive through a 5 m underwater maze. Some rats (N = 12) had bilateral sectioning of the AEN, others a Sham surgery (N = 12). Twelve rats (6 AEN cut and 6 Sham) had 24 post-surgical dive trials over 2 h to activate brainstem neurons to produce Fos, a neuronal activation marker. Remaining rats were non-diving controls. Diving animals had significantly more Fos-positive neurons than non-diving animals in the caudal pressor area, ventral medullary dorsal horn, ventral paratrigeminal nucleus, nucleus tractus solitarius, rostral ventrolateral medulla, Raphe nuclei, A5, Locus Coeruleus, and Kölliker-Fuse area. There were no significant differences in brainstem Fos labeling in rats diving with and without intact AENs. Thus, the AENs are not required for initiation of the diving response. Other nerve(s) that innervate the nose and nasal passages, and/or suprabulbar activation of brainstem neurons, may be responsible for the pattern of neuronal activation observed during repetitive trained diving in rats. These results help define the central neuronal circuitry of the mammalian diving response. PMID:27148082
Edmund Goodwyn and the first description of diving bradycardia.
Vega, Jose L
2017-08-01
Diving bradycardia is a primordial oxygen-conserving reflex by which the heart rate of air-breathing vertebrates, including humans, slows down in response to water immersion. Its discovery is attributed to Paul Bert, whose seminal observation was published in 1870 as part of a series of experiments that examined physiological adaptations to asphyxia in ducks and other animals. However, Edmund Goodwyn, a British physician who studied medicine at the University of Edinburgh, had already described this reflex in his doctoral thesis, which was originally published in Latin in 1786 and again in English in 1788. Ironically, even though Goodwyn's work has yet to be recognized in the diving physiology literature, it was referenced in the very publication that contains Bert's original observation. Thus this article brings Goodwyn's work and its historical context to light and argues that he should be credited with the first description of diving bradycardia. Copyright © 2017 the American Physiological Society.
The trigeminocardiac reflex – a comparison with the diving reflex in humans
Lemaitre, Frederic; Schaller, Bernhard
2015-01-01
The trigeminocardiac reflex (TCR) has previously been described in the literature as a reflexive response of bradycardia, hypotension, and gastric hypermotility seen upon mechanical stimulation in the distribution of the trigeminal nerve. The diving reflex (DR) in humans is characterized by breath-holding, slowing of the heart rate, reduction of limb blood flow and a gradual rise in the mean arterial blood pressure. Although the two reflexes share many similarities, their relationship and especially their functional purpose in humans have yet to be fully elucidated. In the present review, we have tried to integrate and elaborate these two phenomena into a unified physiological concept. Assuming that the TCR and the DR are closely linked functionally and phylogenetically, we have also highlighted the significance of these reflexes in humans. PMID:25995761
Ridgway, S H; Carder, D A; Kamolnick, T; Smith, R R; Schlundt, C E; Elsberry, W R
2001-11-01
Hearing is attenuated in the aerial ear of humans and other land mammals tested in pressure chambers as a result of middle ear impedance changes that result from increased air density. We tested the hypothesis, based on recent middle ear models, that increasing the density of middle ear air at depth might attenuate whale hearing. Two white whales Delphinapterus leucas made dives to a platform at a depth of 5, 100, 200 or 300 m in the Pacific Ocean. During dives to station on the platform for up to 12 min, the whales whistled in response to 500 ms tones projected at random intervals to assess their hearing threshold at each depth. Analysis of response whistle spectra, whistle latency in response to tones and hearing thresholds showed that the increased hydrostatic pressure at depth changed each whale's whistle response at depth, but did not attenuate hearing overall. The finding that whale hearing is not attenuated at depth suggests that sound is conducted through the head tissues of the whale to the ear without requiring the usual ear drum/ossicular chain amplification of the aerial middle ear. These first ever hearing tests in the open ocean demonstrate that zones of audibility for human-made sounds are just as great throughout the depths to which these whales dive, or at least down to 300 m.
The impact of consecutive freshwater trimix dives at altitude on human cardiovascular function.
Lozo, Mislav; Madden, Dennis; Gunjaca, Grgo; Ljubkovic, Marko; Marinovic, Jasna; Dujic, Zeljko
2015-03-01
Self-contained underwater breathing apparatus (SCUBA) diving is regularly associated with numerous asymptomatic changes in cardiovascular function. Freshwater SCUBA diving presents unique challenges compared with open sea diving related to differences in water density and the potential for dive locations at altitude. The aim of this study was to evaluate the impact of freshwater trimix diving at altitude on human cardiovascular function. Ten divers performed two dives in consecutive days at 294 m altitude with the surface interval of 24 h. Both dives were at a depth of 45 m with total dive time 29 and 26 min for the first and second dive, respectively. Assessment of venous gas embolization, hydration status, cardiac function and arterial stiffness was performed. Production of venous gas emboli was low, and there were no significant differences between the dives. After the first dive, diastolic blood pressure was significantly reduced, which persisted up to 24 h. Left ventricular stroke volume decreased, and heart rate increased after both dives. Pulse wave velocity was unchanged following the dives. However, the central and peripheral augmentation index became more negative after both dives, indicating reduced wave reflection. Ejection duration and round trip travel time were prolonged 24 h after the first dive, suggesting longer-lasting suppression of cardiac and endothelial function. This study shows that freshwater trimix dives with conservative profiles and low venous gas bubble loads can result in multiple asymptomatic acute cardiovascular changes some of which were present up to 24 h after dive. © 2014 Scandinavian Society of Clinical Physiology and Nuclear Medicine. Published by John Wiley & Sons Ltd.
Chotiyanonta, Jill S; DiNovo, Karyn M; McCulloch, Paul F
2013-01-01
The diving response is characterized by bradycardia, apnea, and increased peripheral resistance. This reflex response is initiated by immersing the nose in water. Because the anterior ethmoidal nerve (AEN) innervates the nose, our hypothesis was that intact AENs are essential for initiating the diving response in voluntarily diving rats. Heart rate (HR) and arterial blood pressure (BPa) were monitored using implanted biotransmitters. Sprague-Dawley rats were trained to voluntarily swim 5 m underwater. During diving, HR decreased from 480 ± 15 to 99 ± 5 bpm and BPa increased from 136 ± 2 to 187 ± 3 mmHg. Experimental rats (N = 9) then received bilateral AEN sectioning, while Sham rats (N = 8) did not. During diving in Experimental rats 7 days after AEN surgery, HR decreased from 478 ± 13 to 76 ± 4 bpm and BPa increased from 134 ± 3 to 186 ± 4 mmHg. Responses were similar in Sham rats. Then, during nasal stimulation with ammonia vapors in urethane-anesthetized Experimental rats, HR decreased from 368 ± 7 to 83 ± 4 bpm, and BPa increased from 126 ± 7 to 175 ± 4 mmHg. Responses were similar in Sham rats. Thus, 1 week after being sectioned the AENs are not essential for initiating a full cardiorespiratory response during both voluntary diving and nasal stimulation. We conclude that other nerve(s) innervating the nose are able to provide an afferent signal sufficient to initiate the diving response, although neuronal plasticity within the medullary dorsal horn may be necessary for this to occur. PMID:24400143
The marine mammal dive response is exercise modulated to maximize aerobic dive duration.
Davis, Randall W; Williams, Terrie M
2012-08-01
When aquatically adapted mammals and birds swim submerged, they exhibit a dive response in which breathing ceases, heart rate slows, and blood flow to peripheral tissues and organs is reduced. The most intense dive response occurs during forced submersion which conserves blood oxygen for the brain and heart, thereby preventing asphyxiation. In free-diving animals, the dive response is less profound, and energy metabolism remains aerobic. However, even this relatively moderate bradycardia seems diametrically opposed to the normal cardiovascular response (i.e., tachycardia and peripheral vasodilation) during physical exertion. As a result, there has been a long-standing paradox regarding how aquatic mammals and birds exercise while submerged. We hypothesized based on cardiovascular modeling that heart rate must increase to ensure adequate oxygen delivery to active muscles. Here, we show that heart rate (HR) does indeed increase with flipper or fluke stroke frequency (SF) during voluntary, aerobic dives in Weddell seals (HR = 1.48SF - 8.87) and bottlenose dolphins (HR = 0.99SF + 2.46), respectively, two marine mammal species with different evolutionary lineages. These results support our hypothesis that marine mammals maintain aerobic muscle metabolism while swimming submerged by combining elements of both dive and exercise responses, with one or the other predominating depending on the level of exertion.
29 CFR 1910.423 - Post-dive procedures.
Code of Federal Regulations, 2012 CFR
2012-07-01
... human occupancy; (ii) A built-in-breathing-system with a minimum of one mask per occupant; (iii) A two-way voice communication system between occupants and a dive team member at the dive location; (iv) A... location. (6) A dive team member shall be available at the dive location during and for at least one hour...
29 CFR 1910.423 - Post-dive procedures.
Code of Federal Regulations, 2013 CFR
2013-07-01
... human occupancy; (ii) A built-in-breathing-system with a minimum of one mask per occupant; (iii) A two-way voice communication system between occupants and a dive team member at the dive location; (iv) A... location. (6) A dive team member shall be available at the dive location during and for at least one hour...
29 CFR 1910.423 - Post-dive procedures.
Code of Federal Regulations, 2010 CFR
2010-07-01
... human occupancy; (ii) A built-in-breathing-system with a minimum of one mask per occupant; (iii) A two-way voice communication system between occupants and a dive team member at the dive location; (iv) A... location. (6) A dive team member shall be available at the dive location during and for at least one hour...
29 CFR 1910.423 - Post-dive procedures.
Code of Federal Regulations, 2014 CFR
2014-07-01
... human occupancy; (ii) A built-in-breathing-system with a minimum of one mask per occupant; (iii) A two-way voice communication system between occupants and a dive team member at the dive location; (iv) A... location. (6) A dive team member shall be available at the dive location during and for at least one hour...
29 CFR 1910.423 - Post-dive procedures.
Code of Federal Regulations, 2011 CFR
2011-07-01
... human occupancy; (ii) A built-in-breathing-system with a minimum of one mask per occupant; (iii) A two-way voice communication system between occupants and a dive team member at the dive location; (iv) A... location. (6) A dive team member shall be available at the dive location during and for at least one hour...
Diving Response in Rats: Role of the Subthalamic Vasodilator Area
Golanov, Eugene V.; Shiflett, James M.; Britz, Gavin W.
2016-01-01
Diving response (DR) is a powerful integrative response targeted toward survival of the hypoxic/anoxic conditions. Being present in all animals and humans, it allows to survive adverse conditions like diving. Earlier, we discovered that forehead stimulation affords neuroprotective effect, decreasing infarction volume triggered by permanent occlusion of the middle cerebral artery in rats. We hypothesized that cold stimulation of the forehead induces DR in rats, which, in turn, exerts neuroprotection. We compared autonomic [AP, heart rate (HR), cerebral blood flow (CBF)] and EEG responses to the known DR-triggering stimulus, ammonia stimulation of the nasal mucosa, cold stimulation of the forehead, and cold stimulation of the glabrous skin of the tail base in anesthetized rats. Responses in AP, HR, CBF, and EEG to cold stimulation of the forehead and ammonia vapors instillation into the nasal cavity were comparable and differed significantly from responses to the cold stimulation of the tail base. Excitotoxic lesion of the subthalamic vasodilator area (SVA), which is known to participate in CBF regulation and to afford neuroprotection upon excitation, failed to affect autonomic components of the DR evoked by forehead cold stimulation or nasal mucosa ammonia stimulation. We conclude that cold stimulation of the forehead triggers physiological response comparable to the response evoked by ammonia vapor instillation into nasal cavity, which is considered as stimulus triggering protective DR. These observations may explain the neuroprotective effect of the forehead stimulation. Data demonstrate that SVA does not directly participate in the autonomic adjustments accompanying DR; however, it is involved in diving-evoked modulation of EEG. We suggest that forehead stimulation can be employed as a stimulus capable of triggering oxygen-conserving DR and can be used for neuroprotective therapy. PMID:27708614
Running, swimming and diving modifies neuroprotecting globins in the mammalian brain
Williams, Terrie M; Zavanelli, Mary; Miller, Melissa A; Goldbeck, Robert A; Morledge, Michael; Casper, Dave; Pabst, D. Ann; McLellan, William; Cantin, Lucas P; Kliger, David S
2007-01-01
The vulnerability of the human brain to injury following just a few minutes of oxygen deprivation with submergence contrasts markedly with diving mammals, such as Weddell seals (Leptonychotes weddellii), which can remain underwater for more than 90 min while exhibiting no neurological or behavioural impairment. This response occurs despite exposure to blood oxygen levels concomitant with human unconsciousness. To determine whether such aquatic lifestyles result in unique adaptations for avoiding ischaemic–hypoxic neural damage, we measured the presence of circulating (haemoglobin) and resident (neuroglobin and cytoglobin) oxygen-carrying globins in the cerebral cortex of 16 mammalian species considered terrestrial, swimming or diving specialists. Here we report a striking difference in globin levels depending on activity lifestyle. A nearly 9.5-fold range in haemoglobin concentration (0.17–1.62 g Hb 100 g brain wet wt−1) occurred between terrestrial and deep-diving mammals; a threefold range in resident globins was evident between terrestrial and swimming specialists. Together, these two globin groups provide complementary mechanisms for facilitating oxygen transfer into neural tissues and the potential for protection against reactive oxygen and nitrogen groups. This enables marine mammals to maintain sensory and locomotor neural functions during prolonged submergence, and suggests new avenues for averting oxygen-mediated neural injury in the mammalian brain. PMID:18089537
Kvadsheim, P H; Miller, P J O; Tyack, P L; Sivle, L D; Lam, F P A; Fahlman, A
2012-01-01
Naval sonar has been accused of causing whale stranding by a mechanism which increases formation of tissue N(2) gas bubbles. Increased tissue and blood N(2) levels, and thereby increased risk of decompression sickness (DCS), is thought to result from changes in behavior or physiological responses during diving. Previous theoretical studies have used hypothetical sonar-induced changes in both behavior and physiology to model blood and tissue N(2) tension [Formula: see text], but this is the first attempt to estimate the changes during actual behavioral responses to sonar. We used an existing mathematical model to estimate blood and tissue N(2) tension [Formula: see text] from dive data recorded from sperm, killer, long-finned pilot, Blainville's beaked, and Cuvier's beaked whales before and during exposure to Low- (1-2 kHz) and Mid- (2-7 kHz) frequency active sonar. Our objectives were: (1) to determine if differences in dive behavior affects risk of bubble formation, and if (2) behavioral- or (3) physiological responses to sonar are plausible risk factors. Our results suggest that all species have natural high N(2) levels, with deep diving generally resulting in higher end-dive [Formula: see text] as compared with shallow diving. Sonar exposure caused some changes in dive behavior in both killer whales, pilot whales and beaked whales, but this did not lead to any increased risk of DCS. However, in three of eight exposure session with sperm whales, the animal changed to shallower diving, and in all these cases this seem to result in an increased risk of DCS, although risk was still within the normal risk range of this species. When a hypothetical removal of the normal dive response (bradycardia and peripheral vasoconstriction), was added to the behavioral response during model simulations, this led to an increased variance in the estimated end-dive N(2) levels, but no consistent change of risk. In conclusion, we cannot rule out the possibility that a combination of behavioral and physiological responses to sonar have the potential to alter the blood and tissue end-dive N(2) tension to levels which could cause DCS and formation of in vivo bubbles, but the actually observed behavioral responses of cetaceans to sonar in our study, do not imply any significantly increased risk of DCS.
Hooker, Sascha K; Baird, Robin W; Fahlman, Andreas
2009-07-31
A mathematical model, based on current knowledge of gas exchange and physiology of marine mammals, was used to predict blood and tissue tension N2 (P(N2)) using field data from three beaked whale species: northern bottlenose whales, Cuvier's beaked whales, and Blainville's beaked whales. The objective was to determine if physiology (body mass, diving lung volume, dive response) or dive behaviour (dive depth and duration, changes in ascent rate, diel behaviour) would lead to differences in P(N2) levels and thereby decompression sickness (DCS) risk between species. Diving lung volume and extent of the dive response had a large effect on end-dive P(N2). The dive profile had a larger influence on end-dive P(N2) than body mass differences between species. Despite diel changes in dive behaviour, P(N2) levels showed no consistent trend. Model output suggested that all three species live with tissue P(N2) levels that would cause a significant proportion of DCS cases in terrestrial mammals. Cuvier's beaked whale diving behaviour appears to put them at higher risk than the other species, which may explain their prevalence in strandings after the use of mid-frequency sonar.
46 CFR 197.404 - Responsibilities of the diving supervisor.
Code of Federal Regulations, 2010 CFR
2010-10-01
... planning and execution of the diving operation including the responsibility for the safety and health of... SAFETY AND HEALTH STANDARDS GENERAL PROVISIONS Commercial Diving Operations Operations § 197.404... provisions of this subpart; (2) Be fully cognizant of the provisions of the operations manual required by...
NASA Astrophysics Data System (ADS)
Joyce, T. W.; Durban, J. W.; Fearnbach, H. H.; Claridge, D. E.; Ballance, L. T.
2016-02-01
Diving and spatial distribution data from small (55g) satellite transmitter tags attached to five species of deep-diving toothed whales were used to examine the physiological and ecological tradeoffs influencing vertical foraging ranges in the Bahamas. These tradeoffs have important consequences in terms of the ecological impacts of different toothed whale predators on meso- and bathypelagic prey populations, and also on relative vulnerabilities to human impacts (e.g., noise, vessel-strike). Within this assemblage, larger toothed-whales were hypothesized to more efficiently access deeper prey by having the capacity to sustain longer dives, based on a divergence of metabolic rates from oxygen storage capacity as mass increases. However, the observed vertical foraging ranges of melon-headed whales (Peponocephala electra, n=13), short-finned pilot whales (Globicephala macrorhynchus, n=15), Blainville's beaked whales (Mesoplodon densirostris, n=12), Cuvier's beaked whales (Ziphius cavirostris, n=7), and sperm whales (Physeter macrocephalus, n=27), only weakly support hypothesized increases in dive duration and depth as power law functions body mass (R2=0.36 & 0.23). In particular, the relatively small beaked whales (M.d. 853kg; Z.c. 1557kg) performed extremely long and deep foraging dives (M.d. max. 67mins & 1888m; Z.c. max. 103mins & 1888m) relative to expectations of simple allometric scaling. Based on foraging dive durations and post-foraging dive recovery patterns, both beaked whales appear to exceed aerobic dive limits, which enabled access to bathypelagic niches but at the cost of significantly longer recovery periods between foraging dives and comparatively low foraging time efficiency (<29% of time in foraging strata). The inclusion of aerobic and anaerobic dive strategies in allometric models of dive duration and depth yielded considerably greater explanatory power (R2=0.96 & 0.90), providing an improved framework for interpreting the tradeoffs between body size, diving efficiency, and access to different prey layers. Vertical foraging ranges in turn had important implications in terms of responses to diurnal variation in light intensity, and the relative affinities of different species to deep-scattering and benthic boundary layers of prey.
Balestra, C; Cimino, F; Theunissen, S; Snoeck, T; Provyn, S; Canali, R; Bonina, A; Virgili, F
2016-09-01
Nutritional antioxidants have been proposed as an expedient strategy to counter the potentially deleterious effects of scuba diving on endothelial function, flow-mediated dilation (FMD) and heart function. Sixteen volunteers performing a single standard dive (20 min at 33 m) according to US Navy diving procedures were randomly assigned to two groups: one was administered with two doses of 200 mg of an anthocyanins (AC)-rich extract from red oranges, 12 and 4 h before diving. Anthocyanins supplementation significantly modulated the effects of diving on haematocrit, body water distribution and FMD. AC administration significantly reduces the potentially harmful endothelial effects of a recreational single dive. The lack of any significant effect on the most common markers of plasma antioxidant capacity suggests that the mechanism underlying this protective activity is independent of the putative antioxidant effect of AC and possibly involves cellular signalling modulation of the response to high oxygen.
Kvadsheim, P. H.; Miller, P. J. O.; Tyack, P. L.; Sivle, L. D.; Lam, F. P. A.; Fahlman, A.
2012-01-01
Naval sonar has been accused of causing whale stranding by a mechanism which increases formation of tissue N2 gas bubbles. Increased tissue and blood N2 levels, and thereby increased risk of decompression sickness (DCS), is thought to result from changes in behavior or physiological responses during diving. Previous theoretical studies have used hypothetical sonar-induced changes in both behavior and physiology to model blood and tissue N2 tension PN2, but this is the first attempt to estimate the changes during actual behavioral responses to sonar. We used an existing mathematical model to estimate blood and tissue N2 tension PN2 from dive data recorded from sperm, killer, long-finned pilot, Blainville’s beaked, and Cuvier’s beaked whales before and during exposure to Low- (1–2 kHz) and Mid- (2–7 kHz) frequency active sonar. Our objectives were: (1) to determine if differences in dive behavior affects risk of bubble formation, and if (2) behavioral- or (3) physiological responses to sonar are plausible risk factors. Our results suggest that all species have natural high N2 levels, with deep diving generally resulting in higher end-dive PN2 as compared with shallow diving. Sonar exposure caused some changes in dive behavior in both killer whales, pilot whales and beaked whales, but this did not lead to any increased risk of DCS. However, in three of eight exposure session with sperm whales, the animal changed to shallower diving, and in all these cases this seem to result in an increased risk of DCS, although risk was still within the normal risk range of this species. When a hypothetical removal of the normal dive response (bradycardia and peripheral vasoconstriction), was added to the behavioral response during model simulations, this led to an increased variance in the estimated end-dive N2 levels, but no consistent change of risk. In conclusion, we cannot rule out the possibility that a combination of behavioral and physiological responses to sonar have the potential to alter the blood and tissue end-dive N2 tension to levels which could cause DCS and formation of in vivo bubbles, but the actually observed behavioral responses of cetaceans to sonar in our study, do not imply any significantly increased risk of DCS. PMID:22590458
Song, Mengmeng; Cheng, Mengjiao; Ju, Guannan; Zhang, Yajun; Shi, Feng
2014-11-05
A smart device that can dive or surface in aqueous medium has been developed by combining a pH-responsive surface with acid-responsive magnesium. The diving-surfacing cycles can be used to convert chemical energy into electricity. During the diving-surfacing motion, the smart device cuts magnetic flux lines and produces a current, demonstrating that motional energy can be realized by consuming chemical energy of magnesium, thus producing electricity. © 2014 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim.
Parameter estimation of the copernicus decompression model with venous gas emboli in human divers.
Gutvik, Christian R; Dunford, Richard G; Dujic, Zeljko; Brubakk, Alf O
2010-07-01
Decompression Sickness (DCS) may occur when divers decompress from a hyperbaric environment. To prevent this, decompression procedures are used to get safely back to the surface. The models whose procedures are calculated from, are traditionally validated using clinical symptoms as an endpoint. However, DCS is an uncommon phenomenon and the wide variation in individual response to decompression stress is poorly understood. And generally, using clinical examination alone for validation is disadvantageous from a modeling perspective. Currently, the only objective and quantitative measure of decompression stress is Venous Gas Emboli (VGE), measured by either ultrasonic imaging or Doppler. VGE has been shown to be statistically correlated with DCS, and is now widely used in science to evaluate decompression stress from a dive. Until recently no mathematical model has existed to predict VGE from a dive, which motivated the development of the Copernicus model. The present article compiles a selection experimental dives and field data containing computer recorded depth profiles associated with ultrasound measurements of VGE. It describes a parameter estimation problem to fit the model with these data. A total of 185 square bounce dives from DCIEM, Canada, 188 recreational dives with a mix of single, repetitive and multi-day exposures from DAN USA and 84 experimentally designed decompression dives from Split Croatia were used, giving a total of 457 dives. Five selected parameters in the Copernicus bubble model were assigned for estimation and a non-linear optimization problem was formalized with a weighted least square cost function. A bias factor to the DCIEM chamber dives was also included. A Quasi-Newton algorithm (BFGS) from the TOMLAB numerical package solved the problem which was proved to be convex. With the parameter set presented in this article, Copernicus can be implemented in any programming language to estimate VGE from an air dive.
Cardiac responses of grey seals during diving at sea.
Thompson, D; Fedak, M A
1993-01-01
Heart rate, swimming speed and diving depth data were collected from free-ranging grey seals, Halichoerus grypus, as they foraged and travelled in the sea around the Hebrides Islands off western Scotland. Information was collected on a tracking yacht using a combination of sonic and radio telemetry. Diving heart rate declined as a function of dive duration. In long dives, grey seals employed extreme bradycardia, with heart rates falling to 4 beats min-1 for extended periods, despite the animal being free to breath at will. This extreme dive response is part of the normal foraging behaviour. Seals spent 89% of the time submerged during bouts of long dives; swimming was restricted to ascent and descent. Dive durations exceeded estimated aerobic dive limit, even assuming resting metabolic rates. These results indicate that behavioural, and possibly cellular, energy-sparing mechanisms play an important role in diving behaviour of grey seals. This has implications not only for studies of mammalian energetics but also for our understanding of the foraging tactics and prey selection of marine mammals. If some seals are using energy-sparing mechanisms to reduce metabolic costs while at depth, they may be forced to wait for and ambush prey rather than to search for and chase it.
Potentially conflicting metabolic demands of diving and exercise in seals.
Castellini, M A; Murphy, B J; Fedak, M; Ronald, K; Gofton, N; Hochachka, P W
1985-02-01
Metabolic replacement rates (Ra) for glucose and free fatty acids (FFA) were determined during rest, exercise, and diving conditions in the gray seal using bolus injections of radiotracers. In the exercise experiments the seal swam at a metabolic rate elevated twofold over resting Ra for glucose and FFA while resting were similar to values found in terrestrial mammals and other marine mammal species. During exercise periods glucose turnover increased slightly while FFA turnover changes were variable. However, the energetic demands of exercise could not be met by the increase in the replacement rates of glucose or FFA even if both were completely oxidized. Under diving conditions the tracer pool displayed radically different specific activity curves indicative of the changes in perfusion and metabolic rate associated with a strong dive response. Since the radiotracer curves during exercise and diving differed qualitatively and quantitatively, it is possible that similar studies on freely diving animals can be used to assess the role of the diving response during underwater swimming in nature.
Avoidance responses of minke whales to 1-4kHz naval sonar.
Kvadsheim, Petter H; DeRuiter, Stacy; Sivle, Lise D; Goldbogen, Jeremy; Roland-Hansen, Rune; Miller, Patrick J O; Lam, Frans-Peter A; Calambokidis, John; Friedlaender, Ari; Visser, Fleur; Tyack, Peter L; Kleivane, Lars; Southall, Brandon
2017-08-15
Minke whales are difficult to study and little information exists regarding their responses to anthropogenic sound. This study pools data from behavioural response studies off California and Norway. Data are derived from four tagged animals, of which one from each location was exposed to naval sonar signals. Statistical analyses were conducted using Mahalanobis distance to compare overall changes in parameters summarising dive behaviour, avoidance behaviour, and potential energetic costs of disturbance. Our quantitative analysis showed that both animals initiated avoidance behaviour, but responses were not associated with unusual dive behaviour. In one exposed animal the avoidance of the sonar source included a 5-fold increase in horizontal speed away from the source, implying a significant increase in metabolic rate. Despite the different environmental settings and exposure contexts, clear changes in behaviour were observed providing the first insights into the nature of responses to human noise for this wide-ranging species. Copyright © 2017. Published by Elsevier Ltd.
Effects of Long-term Diving Training on Cortical Gyrification.
Zhang, Yuanchao; Zhao, Lu; Bi, Wenwei; Wang, Yue; Wei, Gaoxia; Evans, Alan; Jiang, Tianzi
2016-06-20
During human brain development, cortical gyrification, which is believed to facilitate compact wiring of neural circuits, has been shown to follow an inverted U-shaped curve, coinciding with the two-stage neurodevelopmental process of initial synaptic overproduction with subsequent pruning. This trajectory allows postnatal experiences to refine the wiring, which may manifest as endophenotypic changes in cortical gyrification. Diving experts, typical elite athletes who commence intensive motor training at a very young age in their early childhood, serve ideal models for examining the gyrification changes related to long-term intensive diving training. Using local gyrification index (LGI), we compared the cortical gyrification between 12 diving experts and 12 controls. Compared with controls, diving experts showed widespread LGI reductions in regions relevant to diving performance. Negative correlations between LGIs and years of diving training were also observed in diving experts. Further exploratory network efficiency analysis of structural cortical networks, inferred from interregional correlation of LGIs, revealed comparable global and local efficiency in diving experts relative to controls. These findings suggest that gyrification reductions in diving experts may be the result of long-term diving training which could refine the neural circuitry (via synaptic pruning) and might be the anatomical substrate underlying their extraordinary diving performance.
Decompression sickness ('the bends') in sea turtles.
García-Párraga, D; Crespo-Picazo, J L; de Quirós, Y Bernaldo; Cervera, V; Martí-Bonmati, L; Díaz-Delgado, J; Arbelo, M; Moore, M J; Jepson, P D; Fernández, Antonio
2014-10-16
Decompression sickness (DCS), as clinically diagnosed by reversal of symptoms with recompression, has never been reported in aquatic breath-hold diving vertebrates despite the occurrence of tissue gas tensions sufficient for bubble formation and injury in terrestrial animals. Similarly to diving mammals, sea turtles manage gas exchange and decompression through anatomical, physiological, and behavioral adaptations. In the former group, DCS-like lesions have been observed on necropsies following behavioral disturbance such as high-powered acoustic sources (e.g. active sonar) and in bycaught animals. In sea turtles, in spite of abundant literature on diving physiology and bycatch interference, this is the first report of DCS-like symptoms and lesions. We diagnosed a clinico-pathological condition consistent with DCS in 29 gas-embolized loggerhead sea turtles Caretta caretta from a sample of 67. Fifty-nine were recovered alive and 8 had recently died following bycatch in trawls and gillnets of local fisheries from the east coast of Spain. Gas embolization and distribution in vital organs were evaluated through conventional radiography, computed tomography, and ultrasound. Additionally, positive response following repressurization was clinically observed in 2 live affected turtles. Gas embolism was also observed postmortem in carcasses and tissues as described in cetaceans and human divers. Compositional gas analysis of intravascular bubbles was consistent with DCS. Definitive diagnosis of DCS in sea turtles opens a new era for research in sea turtle diving physiology, conservation, and bycatch impact mitigation, as well as for comparative studies in other air-breathing marine vertebrates and human divers.
Quantification of HTLV-1 Clonality and TCR Diversity
Laydon, Daniel J.; Melamed, Anat; Sim, Aaron; Gillet, Nicolas A.; Sim, Kathleen; Darko, Sam; Kroll, J. Simon; Douek, Daniel C.; Price, David A.; Bangham, Charles R. M.; Asquith, Becca
2014-01-01
Estimation of immunological and microbiological diversity is vital to our understanding of infection and the immune response. For instance, what is the diversity of the T cell repertoire? These questions are partially addressed by high-throughput sequencing techniques that enable identification of immunological and microbiological “species” in a sample. Estimators of the number of unseen species are needed to estimate population diversity from sample diversity. Here we test five widely used non-parametric estimators, and develop and validate a novel method, DivE, to estimate species richness and distribution. We used three independent datasets: (i) viral populations from subjects infected with human T-lymphotropic virus type 1; (ii) T cell antigen receptor clonotype repertoires; and (iii) microbial data from infant faecal samples. When applied to datasets with rarefaction curves that did not plateau, existing estimators systematically increased with sample size. In contrast, DivE consistently and accurately estimated diversity for all datasets. We identify conditions that limit the application of DivE. We also show that DivE can be used to accurately estimate the underlying population frequency distribution. We have developed a novel method that is significantly more accurate than commonly used biodiversity estimators in microbiological and immunological populations. PMID:24945836
Lucrezi, Serena; Egi, Salih Murat; Pieri, Massimo; Burman, Francois; Ozyigit, Tamer; Cialoni, Danilo; Thomas, Guy; Marroni, Alessandro; Saayman, Melville
2018-01-01
Introduction: Scuba diving is an important marine tourism sector, but requires proper safety standards to reduce the risks and increase accessibility to its market. To achieve safety goals, safety awareness and positive safety attitudes in recreational scuba diving operations are essential. However, there is no published research exclusively focusing on scuba divers’ and dive centres’ perceptions toward safety. This study assessed safety perceptions in recreational scuba diving operations, with the aim to inform and enhance safety and risk management programmes within the scuba diving tourism industry. Materials and Methods: Two structured questionnaire surveys were prepared by the organisation Divers Alert Network and administered online to scuba diving operators in Italy and scuba divers in Europe, using a mixture of convenience and snowball sampling. Questions in the survey included experience and safety offered at the dive centre; the buddy system; equipment and accessories for safe diving activities; safety issues in the certification of new scuba divers; incidents/accidents; and attitudes toward safety. Results: 91 scuba diving centres and 3,766 scuba divers participated in the study. Scuba divers gave importance to safety and the responsiveness of service providers, here represented by the dive centres. However, they underestimated the importance of a personal emergency action/assistance plan and, partly, of the buddy system alongside other safety procedures. Scuba divers agreed that some risks, such as those associated with running out of gas, deserve attention. Dive centres gave importance to aspects such as training and emergency action/assistance plans. However, they were limitedly involved in safety campaigning. Dive centres’ perceptions of safety in part aligned with those of scuba divers, with some exceptions. Conclusion: Greater responsibility is required in raising awareness and educating scuba divers, through participation in prevention campaigns and training. The study supports the introduction of programmes aiming to create a culture of safety among dive centres and scuba divers. Two examples, which are described in this paper, include the Hazard Identification and Risk Assessment protocol for dive centres and scuba divers, and the Diving Safety Officer programme to create awareness, improve risk management, and mitigate health and safety risks. PMID:29628904
Lucrezi, Serena; Egi, Salih Murat; Pieri, Massimo; Burman, Francois; Ozyigit, Tamer; Cialoni, Danilo; Thomas, Guy; Marroni, Alessandro; Saayman, Melville
2018-01-01
Introduction: Scuba diving is an important marine tourism sector, but requires proper safety standards to reduce the risks and increase accessibility to its market. To achieve safety goals, safety awareness and positive safety attitudes in recreational scuba diving operations are essential. However, there is no published research exclusively focusing on scuba divers' and dive centres' perceptions toward safety. This study assessed safety perceptions in recreational scuba diving operations, with the aim to inform and enhance safety and risk management programmes within the scuba diving tourism industry. Materials and Methods: Two structured questionnaire surveys were prepared by the organisation Divers Alert Network and administered online to scuba diving operators in Italy and scuba divers in Europe, using a mixture of convenience and snowball sampling. Questions in the survey included experience and safety offered at the dive centre; the buddy system; equipment and accessories for safe diving activities; safety issues in the certification of new scuba divers; incidents/accidents; and attitudes toward safety. Results: 91 scuba diving centres and 3,766 scuba divers participated in the study. Scuba divers gave importance to safety and the responsiveness of service providers, here represented by the dive centres. However, they underestimated the importance of a personal emergency action/assistance plan and, partly, of the buddy system alongside other safety procedures. Scuba divers agreed that some risks, such as those associated with running out of gas, deserve attention. Dive centres gave importance to aspects such as training and emergency action/assistance plans. However, they were limitedly involved in safety campaigning. Dive centres' perceptions of safety in part aligned with those of scuba divers, with some exceptions. Conclusion: Greater responsibility is required in raising awareness and educating scuba divers, through participation in prevention campaigns and training. The study supports the introduction of programmes aiming to create a culture of safety among dive centres and scuba divers. Two examples, which are described in this paper, include the Hazard Identification and Risk Assessment protocol for dive centres and scuba divers, and the Diving Safety Officer programme to create awareness, improve risk management, and mitigate health and safety risks.
2015-09-30
differentiate krill from larger fish targets, as krill have greater backscatter at 120kHz than 38kHz. Figure 3. Clover leaf sampling design...response in dive axis 1 (dive time, surface time, breaths , dive depth, etc.) showed a significant before-after effect including potential changes in...acoustic instruments for fish density estimation: a practical guide. International Council for the Exploration of the Sea (ICES) Cooperative
A no-decompression air dive and ultrasound lung comets.
Dujic, Zeljko; Marinovic, Jasna; Obad, Ante; Ivancev, Vladimir; Breskovic, Toni; Jovovic, Pavle; Ljubkovic, Marko
2011-01-01
Increased accumulation of extravascular lung water after repetitive deep trimix dives was recently reported. This effect was evident 40 min post-dive, but in subsequent studies most signs of this lung congestion were not evident 2-3 h post-dive, indicating no major negative effects on respiratory gas exchange following deep dives. Whether this response is unique for trimix dives or also occurs in more frequent air dives is presently unknown. A single no-decompression field dive to 33 m with 20 min bottom time was performed by 12 male divers. Multiple ultrasound lung comets (ULC), bubble grade (BG), and single-breath lung diffusing capacity (DLCO) measurements were made before and up to 120 min after the dive. Median BG was rather high with maximal values observed at 40 min post-dive [median 4 (4-4)]. Arterialization of bubbles from the venous side was observed only in one diver lasting up to 60 min post-dive. Despite high BG, no DCS symptoms were noted. DLCO and ULC were unchanged after the dive at any time point (DLCO(corr) was 33.6 +/- 1.9 ml x min(-1) mmHg(-1) pre-dive, 32.7 +/- 3.8 ml x min(-1) x mmHg(-1) at 60 min post-dive, and 33.2 +/- 5.3 ml x min(-1) x mmHg(-1) at 120 min post-dive; ULC count was 4.1 +/- 1.9 pre-dive, 4.9 +/- 3.3 at 20 min post-dive, and 3.3 +/- 1.9 at 60 min post-dive. These preliminary findings show no evidence of increased accumulation of extravascular lung water in male divers after a single no-decompression air dive at the limits of accepted Norwegian diving tables.
Diurnal variation in the diving bradycardia response in young men.
Konishi, Masayuki; Kawano, Hiroshi; Xiang, Mi; Kim, Hyeon-Ki; Ando, Karina; Tabata, Hiroki; Nishimaki, Mio; Sakamoto, Shizuo
2016-04-01
The present study aimed to examine diurnal variation of the diving bradycardia responses on the same day. Eighteen young men (age 26 ± 2 years; height 174.2 ± 6.0 cm; body mass 70.2 ± 8.1 kg; body fat 18.0 ± 3.8 %; mean ± standard deviation) participated in this study. Oral temperature, heart rate variability (HRV) from 5-min of electrocardiogram data, and diving bradycardia responses were measured at 0900, 1300, and 1700 hours daily. All participants performed diving reflex tests twice in the sitting position with the face immersed in cold water (1.9-3.1 °C) and apnea at midinspiration for a minimum of 30 s and as long as possible, in consecutive order. Oral temperature was found to be less in the morning (0900) than in the afternoon (1300) and evening (1700). In the frequency domain parameters of heart rate variability, the natural logarithms of high-frequency power were higher in the morning than in the evening. All participants showed bradycardia response to the two diving reflex tests. The peak values of R-R interval during the diving reflex test both for as long as possible and 30 s were longer in the morning than in the afternoon and evening. Our results indicated that the maximal bradycardia during the diving reflex test exhibits a diurnal variation, with peak levels at morning and gradual decrease towards the evening. The HRV indexes show the same variation.
Rodgers, Essie M.; Schwartz, Jonathon J.; Franklin, Craig E.
2015-01-01
Air-breathing, diving ectotherms are a crucial component of the biodiversity and functioning of aquatic ecosystems, but these organisms may be particularly vulnerable to the effects of climate change on submergence times. Ectothermic dive capacity is thermally sensitive, with dive durations significantly reduced by acute increases in water temperature; it is unclear whether diving performance can acclimate/acclimatize in response to long-term exposure to elevated water temperatures. We assessed the thermal sensitivity and plasticity of ‘fright-dive’ capacity in juvenile estuarine crocodiles (Crocodylus porosus; n = 11). Crocodiles were exposed to one of three long-term thermal treatments, designed to emulate water temperatures under differing climate change scenarios (i.e. current summer, 28°C; ‘moderate’ climate warming, 31.5°C; ‘high’ climate warming, 35°C). Dive trials were conducted in a temperature-controlled tank across a range of water temperatures. Dive durations were independent of thermal acclimation treatment, indicating a lack of thermal acclimation response. Acute increases in water temperature resulted in significantly shorter dive durations, with mean submergence times effectively halving with every 3.5°C increase in water temperature (Q10 0.17, P < 0.001). Maximal dive performances, however, were found to be thermally insensitive across the temperature range of 28–35°C. These results suggest that C. porosus have a limited or non-existent capacity to thermally acclimate sustained ‘fright-dive’ performance. If the findings here are applicable to other air-breathing, diving ectotherms, the functional capacity of these organisms will probably be compromised under climate warming. PMID:27293738
2013-09-30
target species (such as melon-headed whales and false killer whales ) for some of these other projects were unusually low in 2013, so that there were...Science Center is supporting research on false killer whale movements in Hawaiian waters, and the Naval Postgraduate School (with funding from N45) is...determine the normal cardiovascular dive response of deep-diving odontocetes like beaked whales , and to examine how that response might be altered
The Mammalian Diving Response: An Enigmatic Reflex to Preserve Life?
2013-01-01
The mammalian diving response is a remarkable behavior that overrides basic homeostatic reflexes. It is most studied in large aquatic mammals but is seen in all vertebrates. Pelagic mammals have developed several physiological adaptations to conserve intrinsic oxygen stores, but the apnea, bradycardia, and vasoconstriction is shared with those terrestrial and is neurally mediated. The adaptations of aquatic mammals are reviewed here as well as the neural control of cardiorespiratory physiology during diving in rodents. PMID:23997188
The oxygen-conserving potential of the diving response: A kinetic-based analysis.
Costalat, Guillaume; Coquart, Jeremy; Castres, Ingrid; Joulia, Fabrice; Sirost, Olivier; Clua, Eric; Lemaître, Frédéric
2017-04-01
We investigated the oxygen-conserving potential of the human diving response by comparing trained breath-hold divers (BHDs) to non-divers (NDs) during simulated dynamic breath-holding (BH). Changes in haemodynamics [heart rate (HR), stroke volume (SV), cardiac output (CO)] and peripheral muscle oxygenation [oxyhaemoglobin ([HbO 2 ]), deoxyhaemoglobin ([HHb]), total haemoglobin ([tHb]), tissue saturation index (TSI)] and peripheral oxygen saturation (SpO 2 ) were continuously recorded during simulated dynamic BH. BHDs showed a breaking point in HR kinetics at mid-BH immediately preceding a more pronounced drop in HR (-0.86 bpm.% -1 ) while HR kinetics in NDs steadily decreased throughout BH (-0.47 bpm.% -1 ). By contrast, SV remained unchanged during BH in both groups (all P > 0.05). Near-infrared spectroscopy (NIRS) results (mean ± SD) expressed as percentage changes from the initial values showed a lower [HHb] increase for BHDs than for NDs at the cessation of BH (+24.0 ± 10.1 vs. +39.2 ± 9.6%, respectively; P < 0.05). As a result, BHDs showed a [tHb] drop that NDs did not at the end of BH (-7.3 ± 3.2 vs. -3.0 ± 4.7%, respectively; P < 0.05). The most striking finding of the present study was that BHDs presented an increase in oxygen-conserving efficiency due to substantial shifts in both cardiac and peripheral haemodynamics during simulated BH. In addition, the kinetic-based approach we used provides further credence to the concept of an "oxygen-conserving breaking point" in the human diving response.
NASA Technical Reports Server (NTRS)
Smith, Scott M.; Davis-Street, Janis E.; Fesperman, J. Vernell; Smith, Myra D.; Rice, Barbara L.; Zwart, Sara R.
2004-01-01
Ground-based analogs of spaceflight are an important means of studying physiologic and nutritional changes associated with space travel, and the NASA Extreme Environment Mission Operations V (NEEMO) is such an analog. To determine whether saturation diving has nutrition-related effects similar to those of spaceflight, we conducted a clinical nutritional assessment of the NEEMO crew (4 men, 2 women) before, during, and after their 14-d saturation dive. Blood and urine samples were collected before, during, and after the dive. The foods consumed by the crew were typical of the spaceflight food system. A number of physiologic changes were observed, during and after the dive, that are also commonly observed during spaceflight. Hemoglobin and hematocrit were lower (P < 0.05) after the dive. Transferrin receptors were significantly lower immediately after the dive. Serum ferritin increased significantly during the dive. There was also evidence indicating that oxidative damage and stress increased during the dive. Glutathione peroxidase and superoxide dismutase decreased during and after the dive (P < 0.05). Decreased leptin during the dive (P < 0.05) may have been related to the increased stress. Subjects had decreased energy intake and weight loss during the dive, similar to what is observed during spaceflight. Together, these similarities to spaceflight provide a model to use in further defining the physiologic effects of spaceflight and investigating potential countermeasures.
Risk of Neurological Insult in Competitive Deep Breath-Hold Diving.
Tetzlaff, Kay; Schöppenthau, Holger; Schipke, Jochen D
2017-02-01
It has been widely believed that tissue nitrogen uptake from the lungs during breath-hold diving would be insufficient to cause decompression stress in humans. With competitive free diving, however, diving depths have been ever increasing over the past decades. A case is presented of a competitive free-diving athlete who suffered stroke-like symptoms after surfacing from his last dive of a series of 3 deep breath-hold dives. A literature and Web search was performed to screen for similar cases of subjects with serious neurological symptoms after deep breath-hold dives. A previously healthy 31-y-old athlete experienced right-sided motor weakness and difficulty speaking immediately after surfacing from a breathhold dive to a depth of 100 m. He had performed 2 preceding breath-hold dives to that depth with surface intervals of only 15 min. The presentation of symptoms and neuroimaging findings supported a clinical diagnosis of stroke. Three more cases of neurological insults were retrieved by literature and Web search; in all cases the athletes presented with stroke-like symptoms after single breath-hold dives of depths exceeding 100 m. Two of these cases only had a short delay to recompression treatment and completely recovered from the insult. This report highlights the possibility of neurological insult, eg, stroke, due to cerebral arterial gas embolism as a consequence of decompression stress after deep breath-hold dives. Thus, stroke as a clinical presentation of cerebral arterial gas embolism should be considered another risk of extreme breath-hold diving.
NASA Astrophysics Data System (ADS)
Jung, Sunghwan; Chang, Brian; Croson, Matt; Straker, Lorian; Dove, Carla
2015-03-01
Diving is the activity of falling from air into water, which is somewhat dangerous due to the impact. Humans dive for entertainments less than 20 meters high, however seabirds dive as a hunting mechanism from more than 20 meters high. Moreover, most birds including seabirds have a slender and long neck compared to many other animals, which can potentially be the weakest part of the body upon axial impact compression. Motivated by the diving dynamics, we investigate the effect of surface and geometric configurations on structures consisting of a beak-like cone and a neck-like elastic beam. A transition from non-buckling to buckling is characterized and understood through physical experiments and an analytical model.
Madden, Dennis; Thom, Stephen R; Yang, Ming; Bhopale, Veena M; Ljubkovic, Marko; Dujic, Zeljko
2014-09-01
Venous gas emboli (VGE) have traditionally served as a marker for decompression stress after SCUBA diving and a reduction in bubble loads is a target for precondition procedures. However, VGE can be observed in large quantities with no negative clinical consequences. The effect of exercise before diving on VGE has been evaluated with mixed results. Microparticle (MP) counts and sub-type expression serve as indicators of vascular inflammation and DCS in mice. The goal of the present study is to evaluate the effect of anaerobic cycling (AC) on VGE and MP following SCUBA diving. Ten male divers performed two dives to 18 m for 41 min, one dive (AC) was preceded by a repeated-Wingate cycling protocol; a control dive (CON) was completed without exercise. VGE were analyzed at 15, 40, 80, and 120 min post-diving. Blood for MP analysis was collected before exercise (AC only), before diving, 15 and 120 min after surfacing. VGE were significantly lower 15 min post-diving in the AC group, with no difference in the remaining measurements. MPs were elevated by exercise and diving, however, post-diving elevations were attenuated in the AC dive. Some markers of neutrophil elevation (CD18, CD41) were increased in the CON compared to the AC dive. The repeated-Wingate protocol resulted in an attenuation of MP counts and sub-types that have been related to vascular injury and DCS-like symptoms in mice. Further studies are needed to determine if MPs represent a risk factor or marker for DCS in humans.
How seabirds plunge-dive without injuries
NASA Astrophysics Data System (ADS)
Chang, Brian; Croson, Matthew; Straker, Lorian; Gart, Sean; Dove, Carla; Gerwin, John; Jung, Sunghwan
In nature, several seabirds (e.g., gannets and boobies) dive into water at up to 24 m/s as a hunting mechanism; furthermore, gannets and boobies have a slender neck, which is potentially the weakest part of the body under compression during high-speed impact. In this work, we investigate the stability of the bird's neck during plunge-diving by understanding the interaction between the fluid forces acting on the head and the flexibility of the neck. First, we use a salvaged bird to identify plunge-diving phases. Anatomical features of the skull and neck were acquired to quantify the effect of beak geometry and neck musculature on the stability during a plunge-dive. Second, physical experiments using an elastic beam as a model for the neck attached to a skull-like cone revealed the limits for the stability of the neck during the bird's dive as a function of impact velocity and geometric factors. We find that the neck length, neck muscles, and diving speed of the bird predominantly reduce the likelihood of injury during the plunge-dive. Finally, we use our results to discuss maximum diving speeds for humans to avoid injury.
Murphy, F Gregory; Hada, Ethan A; Doolette, David J; Howle, Laurens E
2017-07-01
Decompression sickness (DCS) is a disease caused by gas bubbles forming in body tissues following a reduction in ambient pressure, such as occurs in scuba diving. Probabilistic models for quantifying the risk of DCS are typically composed of a collection of independent, perfusion-limited theoretical tissue compartments which describe gas content or bubble volume within these compartments. It has been previously shown that 'pharmacokinetic' gas content models, with compartments coupled in series, show promise as predictors of the incidence of DCS. The mechanism of coupling can be through perfusion or diffusion. This work examines the application of five novel pharmacokinetic structures with compartments coupled by perfusion to the prediction of the probability and time of onset of DCS in humans. We optimize these models against a training set of human dive trial data consisting of 4335 exposures with 223 DCS cases. Further, we examine the extrapolation quality of the models on an additional set of human dive trial data consisting of 3140 exposures with 147 DCS cases. We find that pharmacokinetic models describe the incidence of DCS for single air bounce dives better than a single-compartment, perfusion-limited model. We further find the U.S. Navy LEM-NMRI98 is a better predictor of DCS risk for the entire training set than any of our pharmacokinetic models. However, one of the pharmacokinetic models we consider, the CS2T3 model, is a better predictor of DCS risk for single air bounce dives and oxygen decompression dives. Additionally, we find that LEM-NMRI98 outperforms CS2T3 on the extrapolation data. Copyright © 2017 Elsevier Ltd. All rights reserved.
Williams, Terrie M; Fuiman, Lee A; Kendall, Traci; Berry, Patrick; Richter, Beau; Noren, Shawn R; Thometz, Nicole; Shattock, Michael J; Farrell, Edward; Stamper, Andy M; Davis, Randall W
2015-01-16
Unlike their terrestrial ancestors, marine mammals routinely confront extreme physiological and physical challenges while breath-holding and pursuing prey at depth. To determine how cetaceans and pinnipeds accomplish deep-sea chases, we deployed animal-borne instruments that recorded high-resolution electrocardiograms, behaviour and flipper accelerations of bottlenose dolphins (Tursiops truncatus) and Weddell seals (Leptonychotes weddellii) diving from the surface to >200 m. Here we report that both exercise and depth alter the bradycardia associated with the dive response, with the greatest impacts at depths inducing lung collapse. Unexpectedly, cardiac arrhythmias occurred in >73% of deep, aerobic dives, which we attribute to the interplay between sympathetic and parasympathetic drivers for exercise and diving, respectively. Such marked cardiac variability alters the common view of a stereotypic 'dive reflex' in diving mammals. It also suggests the persistence of ancestral terrestrial traits in cardiac function that may help explain the unique sensitivity of some deep-diving marine mammals to anthropogenic disturbances.
Immune responses in humans after 60 days of confinement
NASA Technical Reports Server (NTRS)
Schmitt, D. A.; Peres, C.; Sonnenfeld, G.; Tkackzuk, J.; Arquier, M.; Mauco, G.; Ohayon, E.
1995-01-01
A confinement experiment in a normobaric diving chamber was undertaken to better understand the effect of confinement and isolation on human psychology and physiology. Pre- and postconfinement blood samples were obtained from four test subjects and control donors to analyze immune responses. No modification in the levels of CD2+, CD3+, CD4+, CD8+, CD19+, and CD56+ cells was observed after confinement. Mitogen-induced T-lymphocyte proliferation and interleukin-2 receptor expression were not altered significantly. Whole blood interferon-alpha and gamma-induction and plasma cortisol levels were also unchanged, as was natural killer cell activity. These data suggest that in humans, no specific components of the immune response are affected by a 2-month isolation and confinement of a small group.
NASA Astrophysics Data System (ADS)
Cox, S. L.; Miller, P. I.; Embling, C. B.; Scales, K. L.; Bicknell, A. W. J.; Hosegood, P. J.; Morgan, G.; Ingram, S. N.; Votier, S. C.
2016-09-01
Oceanic fronts are key habitats for a diverse range of marine predators, yet how they influence fine-scale foraging behaviour is poorly understood. Here, we investigated the dive behaviour of northern gannets Morus bassanus in relation to shelf-sea fronts. We GPS (global positioning system) tracked 53 breeding birds and examined the relationship between 1901 foraging dives (from time-depth recorders) and thermal fronts (identified via Earth Observation composite front mapping) in the Celtic Sea, Northeast Atlantic. We (i) used a habitat-use availability analysis to determine whether gannets preferentially dived at fronts, and (ii) compared dive characteristics in relation to fronts to investigate the functional significance of these oceanographic features. We found that relationships between gannet dive probabilities and fronts varied by frontal metric and sex. While both sexes were more likely to dive in the presence of seasonally persistent fronts, links to more ephemeral features were less clear. Here, males were positively correlated with distance to front and cross-front gradient strength, with the reverse for females. Both sexes performed two dive strategies: shallow V-shaped plunge dives with little or no active swim phase (92% of dives) and deeper U-shaped dives with an active pursuit phase of at least 3 s (8% of dives). When foraging around fronts, gannets were half as likely to engage in U-shaped dives compared with V-shaped dives, independent of sex. Moreover, V-shaped dive durations were significantly shortened around fronts. These behavioural responses support the assertion that fronts are important foraging habitats for marine predators, and suggest a possible mechanistic link between the two in terms of dive behaviour. This research also emphasizes the importance of cross-disciplinary research when attempting to understand marine ecosystems.
User settings on dive computers: reliability in aiding conservative diving.
Sayer, Martin D J; Azzopardi, Elaine; Sieber, Arne
2016-06-01
Divers can make adjustments to diving computers when they may need or want to dive more conservatively (e.g., diving with a persistent (patent) foramen ovale). Information describing the effects of these alterations or how they compare to other methods, such as using enriched air nitrox (EANx) with air dive planning tools, is lacking. Seven models of dive computer from four manufacturers (Mares, Suunto, Oceanic and UWATEC) were subjected to single square-wave compression profiles (maximum depth: 20 or 40 metres' sea water, msw), single multi-level profiles (maximum depth: 30 msw; stops at 15 and 6 msw), and multi-dive series (two dives to 30 msw followed by one to 20 msw). Adjustable settings were employed for each dive profile; some modified profiles were compared against stand-alone use of EANx. Dives were shorter or indicated longer decompression obligations when conservative settings were applied. However, some computers in default settings produced more conservative dives than others that had been modified. Some computer-generated penalties were greater than when using EANx alone, particularly at partial pressures of oxygen (PO₂) below 1.40 bar. Some computers 'locked out' during the multi-dive series; others would continue to support decompression with, in some cases, automatically-reduced levels of conservatism. Changing reduced gradient bubble model values on Suunto computers produced few differences. The range of possible adjustments and the non-standard computer response to them complicates the ability to provide accurate guidance to divers wanting to dive more conservatively. The use of EANx alone may not always generate satisfactory levels of conservatism.
A Machine Reading System for Assembling Synthetic Paleontological Databases
Peters, Shanan E.; Zhang, Ce; Livny, Miron; Ré, Christopher
2014-01-01
Many aspects of macroevolutionary theory and our understanding of biotic responses to global environmental change derive from literature-based compilations of paleontological data. Existing manually assembled databases are, however, incomplete and difficult to assess and enhance with new data types. Here, we develop and validate the quality of a machine reading system, PaleoDeepDive, that automatically locates and extracts data from heterogeneous text, tables, and figures in publications. PaleoDeepDive performs comparably to humans in several complex data extraction and inference tasks and generates congruent synthetic results that describe the geological history of taxonomic diversity and genus-level rates of origination and extinction. Unlike traditional databases, PaleoDeepDive produces a probabilistic database that systematically improves as information is added. We show that the system can readily accommodate sophisticated data types, such as morphological data in biological illustrations and associated textual descriptions. Our machine reading approach to scientific data integration and synthesis brings within reach many questions that are currently underdetermined and does so in ways that may stimulate entirely new modes of inquiry. PMID:25436610
Perović, Antonija; Sobočanec, Sandra; Dabelić, Sanja; Balog, Tihomir; Dumić, Jerka
2018-02-01
The aim of this study was to examine the effects of scuba diving on oxidative damage markers in erythrocytes and plasma, antioxidant system in peripheral blood mononuclear cells (PBMCs), as well as sirtuin 1 (SIRT1) and sirtuin 3 (SIRT3) gene expressions in recreational divers after a winter nondive period (at least 5 months). For that purpose, 17 male recreational divers performed an immersion at a depth of 30 m for 30 min. Blood samples were collected immediately before and after diving, 3 and 6 h after diving. Erythrocyte lipid peroxidation measured by thiobarbituric-reactive substances (TBARS) method was significantly increased immediately after diving, but returned to the baseline 6 h after diving, while no significant change was found for plasma TBARS and protein carbonyl derivates in both plasma and erythrocytes. Diving-induced catalase (CAT), superoxide dismutase 2 (SOD2), and consequently total superoxide dismutase (SOD) activities in the PBMC samples (significantly increased immediately after diving, reached the maximum activities 3 h after diving, while 6 h after diving only CAT activity remained significantly increased). No significant change was observed for SOD1 activity and gene expression, as well as SOD2 expression, while CAT and SIRT1 expressions were slightly decreased immediately after diving and 3 h after diving. Interestingly, SIRT3 expression was significantly increased 6 h after diving. In conclusion, after the first dive to 30 m after a nondive season, activation of antioxidant defence was not sufficient to prevent oxidative damage, while SIRT3 upregulation could be a step towards an adaptive response to the diving.
Rosen, David A S; Hindle, Allyson G; Gerlinsky, Carling D; Goundie, Elizabeth; Hastie, Gordon D; Volpov, Beth L; Trites, Andrew W
2017-01-01
Marine mammals are characterized as having physiological specializations that maximize the use of oxygen stores to prolong time spent under water. However, it has been difficult to undertake the requisite controlled studies to determine the physiological limitations and trade-offs that marine mammals face while diving in the wild under varying environmental and nutritional conditions. For the past decade, Steller sea lions (Eumetopias jubatus) trained to swim and dive in the open ocean away from the physical confines of pools participated in studies that investigated the interactions between diving behaviour, energetic costs, physiological constraints, and prey availability. Many of these studies measured the cost of diving to understand how it varies with behaviour and environmental and physiological conditions. Collectively, these studies show that the type of diving (dive bouts or single dives), the level of underwater activity, the depth and duration of dives, and the nutritional status and physical condition of the animal affect the cost of diving and foraging. They show that dive depth, dive and surface duration, and the type of dive result in physiological adjustments (heart rate, gas exchange) that may be independent of energy expenditure. They also demonstrate that changes in prey abundance and nutritional status cause sea lions to alter the balance between time spent at the surface acquiring oxygen (and offloading CO 2 and other metabolic by-products) and time spent at depth acquiring prey. These new insights into the physiological basis of diving behaviour further our understanding of the potential scope for behavioural responses of marine mammals to environmental changes, the energetic significance of these adjustments, and the consequences of approaching physiological limits.
Antioxidant pretreatment and reduced arterial endothelial dysfunction after diving.
Obad, Ante; Valic, Zoran; Palada, Ivan; Brubakk, Alf O; Modun, Darko; Dujić, Zeljko
2007-12-01
We have recently shown that a single air dive leads to acute arterial vasodilation and impairment of endothelium-dependent vasodilatation in humans. Additionally we have found that predive antioxidants at the upper recommended daily allowance partially prevented some of the negative effects of the dive. In this study we prospectively evaluated the effect of long-term antioxidants at a lower RDA dose on arterial endothelial function. Eight professional male divers performed an open sea air dive to 30 msw. Brachial artery flow-mediated dilation (FMD) was assessed before and after diving. The first dive, without antioxidants, caused significant brachial arterial diameter increase from 3.85 +/- 0.55 to 4.04 +/- 0.5 mm and a significant reduction of FMD from 7.6 +/- 2.7 to 2.8 +/- 2.1%. The second dive, with antioxidants, showed unchanged arterial diameter and significant reduction of FMD from 8.11 +/- 2.4 to 6.8 +/- 1.4%. The FMD reduction was significantly less with antioxidants. Vascular smooth muscle function, assessed by nitroglycerine (endothelium-independent dilation), was unaffected by diving. This study shows that long-term antioxidant treatment at a lower RDA dose ending 3-4 h before a dive reduces the endothelial dysfunction in divers. Since the scuba dive was of a similar depth and duration to those practiced by numerous recreational divers, this study raises the possibility of routine predive supplementation with antioxidants.
Diving behaviour of Cuvier's beaked whales exposed to two types of military sonar.
Falcone, Erin A; Schorr, Gregory S; Watwood, Stephanie L; DeRuiter, Stacy L; Zerbini, Alexandre N; Andrews, Russel D; Morrissey, Ronald P; Moretti, David J
2017-08-01
Cuvier's beaked whales ( Ziphius cavirostris ) have stranded in association with mid-frequency active sonar (MFAS) use, and though the causative mechanism linking these events remains unclear, it is believed to be behaviourally mediated. To determine whether MFAS use was associated with behavioural changes in this species, satellite tags were used to record the diving and movements of 16 Cuvier's beaked whales for up to 88 days in a region of frequent MFAS training off the coast of Southern California. Tag data were combined with summarized records of concurrent bouts of high-power, surface-ship and mid-power, helicopter-deployed MFAS use, along with other potential covariates, in generalized additive mixed-effects models. Deep dives, shallow dives and surface intervals tended to become longer during MFAS use, with some variation associated with the total amount of overlapping MFAS during the behaviour. These changes in dives and surface intervals contributed to a longer interval between deep dives, a proxy for foraging disruption in this species. Most responses intensified with proximity and were more pronounced during mid-power than high-power MFAS use at comparable distances within approximately 50 km, despite the significantly lower source level of mid-power MFAS. However, distance-mediated responses to high-power MFAS, and increased deep dive intervals during mid-power MFAS, were evident up to approximately 100 km away.
Diving behaviour of Cuvier's beaked whales exposed to two types of military sonar
Schorr, Gregory S.; Watwood, Stephanie L.; DeRuiter, Stacy L.; Zerbini, Alexandre N.; Andrews, Russel D.; Morrissey, Ronald P.; Moretti, David J.
2017-01-01
Cuvier's beaked whales (Ziphius cavirostris) have stranded in association with mid-frequency active sonar (MFAS) use, and though the causative mechanism linking these events remains unclear, it is believed to be behaviourally mediated. To determine whether MFAS use was associated with behavioural changes in this species, satellite tags were used to record the diving and movements of 16 Cuvier's beaked whales for up to 88 days in a region of frequent MFAS training off the coast of Southern California. Tag data were combined with summarized records of concurrent bouts of high-power, surface-ship and mid-power, helicopter-deployed MFAS use, along with other potential covariates, in generalized additive mixed-effects models. Deep dives, shallow dives and surface intervals tended to become longer during MFAS use, with some variation associated with the total amount of overlapping MFAS during the behaviour. These changes in dives and surface intervals contributed to a longer interval between deep dives, a proxy for foraging disruption in this species. Most responses intensified with proximity and were more pronounced during mid-power than high-power MFAS use at comparable distances within approximately 50 km, despite the significantly lower source level of mid-power MFAS. However, distance-mediated responses to high-power MFAS, and increased deep dive intervals during mid-power MFAS, were evident up to approximately 100 km away. PMID:28879004
2018-01-01
Skin lesion which develops after deep sea diving is termed as coral dermatitis. The corals are known to produce a toxic substance which when comes in to contact with human skin may elicit hypersensitive reactions. Most previous reports highlight the allergic reactions caused by deep sea diving. This is a rare case of staphylococcal skin infection in a second-year medical student caused by Staphylococcus aureus; he reported a history of deep sea diving before being presented to the hospital with skin rashes. This case highlights the importance of considering infectious aetiology in cases of coral dermatitis. PMID:29666774
Extracting Databases from Dark Data with DeepDive.
Zhang, Ce; Shin, Jaeho; Ré, Christopher; Cafarella, Michael; Niu, Feng
2016-01-01
DeepDive is a system for extracting relational databases from dark data : the mass of text, tables, and images that are widely collected and stored but which cannot be exploited by standard relational tools. If the information in dark data - scientific papers, Web classified ads, customer service notes, and so on - were instead in a relational database, it would give analysts a massive and valuable new set of "big data." DeepDive is distinctive when compared to previous information extraction systems in its ability to obtain very high precision and recall at reasonable engineering cost; in a number of applications, we have used DeepDive to create databases with accuracy that meets that of human annotators. To date we have successfully deployed DeepDive to create data-centric applications for insurance, materials science, genomics, paleontologists, law enforcement, and others. The data unlocked by DeepDive represents a massive opportunity for industry, government, and scientific researchers. DeepDive is enabled by an unusual design that combines large-scale probabilistic inference with a novel developer interaction cycle. This design is enabled by several core innovations around probabilistic training and inference.
NASA Technical Reports Server (NTRS)
Smith, S. M.; Davis-Street, J. E.; Fesperman, J. V.; Smith, M. D.; Rice, B. L.; Zwart, S. R.
2006-01-01
Ground-based analogs of spaceflight are an important means of studying physiological and nutritional changes associated with space travel, particularly since exploration missions are anticipated, and flight research opportunities are limited. A clinical nutritional assessment of the NASA Extreme Environment Mission Operation V (NEEMO) crew (4 M, 2 F) was conducted before, during, and after the 14-d saturation dive. Blood and urine samples were collected before (D-12 and D-1), during (MD 7 and MD 12), and after (R + 0 and R + 7) the dive. The foods were typical of the spaceflight food system. A number of physiological changes were reported both during the dive and post dive that are also commonly observed during spaceflight. Serum hemoglobin and hematocrit were decreased (P less than 0.05) post dive. Serum ferritin and ceruloplasmin significantly increased during the dive, while transferring receptors tended to go down during the dive and were significantly decreased by the last day (R + 0). Along with significant hematological changes, there was also evidence for increased oxidative damage and stress during the dive. 8-hydroxydeoxyguanosine was elevated (P less than 0.05) during the dive, while glutathione peroxidase and superoxide disrnutase activities were decreased (P less than 0.05) during the dive. Serum C-reactive protein (CRP) concentration also tended to increase during the dive, suggesting the presence of a stress-induced inflammatory response, Decreased leptin during the dive (P less than 0.05) may also be related to the increased stress. Similar to what is observed during spaceflight, subjects had decreased energy intake and weight loss during the dive. Together, these similarities to spaceflight provide a model to further define the physiological effects of spaceflight and investigate potential countermeasures.
Obad, Ante; Palada, Ivan; Valic, Zoran; Ivančev, Vladimir; Baković, Darija; Wisløff, Ulrik; Brubakk, Alf O; Dujić, Željko
2007-01-01
Diving-induced acute alterations in cardiovascular function such as arterial endothelial dysfunction, increased pulmonary artery pressure (PAP) and reduced heart function have been recently reported. We tested the effects of acute antioxidants on arterial endothelial function, PAP and heart function before and after a field dive. Vitamins C (2 g) and E (400 IU) were given to subjects 2 h before a second dive (protocol 1) and in a placebo-controlled crossover study design (protocol 2). Seven experienced divers performed open sea dives to 30 msw with standard decompression in a non-randomized protocol, and six of them participated in a randomized trial. Before and after the dives ventricular volumes and function and pulmonary and brachial artery function were assessed by ultrasound. The control dive resulted in a significant reduction in flow-mediated dilatation (FMD) and heart function with increased mean PAP. Twenty-four hours after the control dive FMD was still reduced 37% below baseline (8.1 versus 5.1%, P = 0.005), while right ventricle ejection fraction (RV-EF), left ventricle EF and endocardial fractional shortening were reduced much less (∼2–3%). At the same time RV end-systolic volume was increased by 9% and mean PAP by 5%. Acute antioxidants significantly attenuated only the reduction in FMD post-dive (P < 0.001), while changes in pulmonary artery and heart function were unaffected by antioxidant ingestion. These findings were confirmed by repeating the experiments in a randomized study design. FMD returned to baseline values 72 h after the dive with pre-dive placebo, whereas for most cardiovascular parameters this occurred earlier (24–48 h). Right ventricular dysfunction and increased PAP lasted longer. Acute antioxidants attenuated arterial endothelial dysfunction after diving, while reduction in heart and pulmonary artery function were unchanged. Cardiovascular changes after diving are not fully reversed up to 3 days after a dive, suggesting longer lasting negative effects. PMID:17110413
Obad, Ante; Palada, Ivan; Valic, Zoran; Ivancev, Vladimir; Baković, Darija; Wisløff, Ulrik; Brubakk, Alf O; Dujić, Zeljko
2007-02-01
Diving-induced acute alterations in cardiovascular function such as arterial endothelial dysfunction, increased pulmonary artery pressure (PAP) and reduced heart function have been recently reported. We tested the effects of acute antioxidants on arterial endothelial function, PAP and heart function before and after a field dive. Vitamins C (2 g) and E (400 IU) were given to subjects 2 h before a second dive (protocol 1) and in a placebo-controlled crossover study design (protocol 2). Seven experienced divers performed open sea dives to 30 msw with standard decompression in a non-randomized protocol, and six of them participated in a randomized trial. Before and after the dives ventricular volumes and function and pulmonary and brachial artery function were assessed by ultrasound. The control dive resulted in a significant reduction in flow-mediated dilatation (FMD) and heart function with increased mean PAP. Twenty-four hours after the control dive FMD was still reduced 37% below baseline (8.1 versus 5.1%, P = 0.005), while right ventricle ejection fraction (RV-EF), left ventricle EF and endocardial fractional shortening were reduced much less (approximately 2-3%). At the same time RV end-systolic volume was increased by 9% and mean PAP by 5%. Acute antioxidants significantly attenuated only the reduction in FMD post-dive (P < 0.001), while changes in pulmonary artery and heart function were unaffected by antioxidant ingestion. These findings were confirmed by repeating the experiments in a randomized study design. FMD returned to baseline values 72 h after the dive with pre-dive placebo, whereas for most cardiovascular parameters this occurred earlier (24-48 h). Right ventricular dysfunction and increased PAP lasted longer. Acute antioxidants attenuated arterial endothelial dysfunction after diving, while reduction in heart and pulmonary artery function were unchanged. Cardiovascular changes after diving are not fully reversed up to 3 days after a dive, suggesting longer lasting negative effects.
'Diving reflex' in man - Its relation to isometric and dynamic exercise.
NASA Technical Reports Server (NTRS)
Bergman, S. A., Jr.; Campbell, J. K.; Wildenthal, K.
1972-01-01
To test the influence of physical activity on the diving reflex, 10 normal men held their breath with their faces immersed in 15 C water during rest, bicycle exercise, and sustained isometric handgrip contraction. At all conditions, a slight but statistically significant elevation of blood pressure and a marked decrease in heart rate occurred during each dive. During moderate bicycle exercise heart rate fell more rapidly than at rest and the final level of bradycardia approached that achieved at rest, despite the fact that predive heart rates were much higher during exercise. When diving occurred in combination with isometric exercise, bradycardia was less severe than during resting dives and final heart rates could be represented as the sum of the expected responses to each intervention alone. In all conditions apnea without face immersion caused bradycardia that was less severe than during wet dives.
Titus, Benjamin M; Daly, Marymegan; Exton, Dan A
2015-01-01
Contact between humans and the marine environment is increasing, but the capacity of communities to adapt to human presence remains largely unknown. The popularization of SCUBA diving has added a new dimension to human impacts in aquatic systems and, although individual-level impacts have been identified, cumulative effects on ecosystem function and community-wide responses are unclear. In principle, habituation may mitigate the consequences of human presence on the biology of an individual and allow the quick resumption of its ecological roles, but this has not been documented in aquatic systems. Here, we investigate the short-term impact of human presence and the long-term habituation potential of reef-fish communities to recreational SCUBA divers by studying symbiotic cleaning interactions on coral reefs with differing levels of historical contact with divers. We show that incidences of human contact result in a smaller decline in ecosystem function and more rapid resumption of baseline services on a reef in Utila, Honduras that has heavy historical levels of SCUBA diver presence, compared to an un-dived reef site in the Cayos Cochinos Marine Protected Area (CCMPA). Nonetheless, despite the generally smaller change in ecosystem function and decades of regular contact with divers, cleaning behavior is suppressed by >50% at Utila when divers are present. We hypothesize that community-wide habituation of reef fish is not fully achievable and may be biologically restricted to only partial habituation. Differential responses to human presence impacts the interpretation and execution of behavioral research where SCUBA is the predominant means of data collection, and provides an important rationale for future research investigating the interplay between human presence, ecosystem function, and community structure on coral reefs.
NASA Astrophysics Data System (ADS)
Ortega-Martinez, Antonio; Goenka, Chhavi; Booker, Marloes; Grange, Robert M. H.; Hindle, Allyson G.; Franco, Walfre
2018-02-01
Marine mammals possess impressive breath-holding capabilities made possible by physiological adjustments during dives. Studying marine mammals in their natural environment unravels vital information about these physiological adjustments particularly when we can monitor altered dive behavior in response to stressful situations such as human-induced oceanic disturbances, presence of predators and altered prey distributions. An important indicator of physiological status during submergence is the change in oxygen saturation in the muscles and blood of these mammals. In this work, we aim to investigate oxygen storage and consumption in the muscles of free-diving elephant seals when exposed to disturbances such as sonar or predator sounds while they are at sea. Optical oxygen sensors are a mature technology with multiple medical applications that provide a way to measure oxygenation changes in biological tissues in a minimally invasive manner. While these sensors are well calibrated and readily available for humans, they are still inadequate for marine mammals primarily due to a very small number of test candidates and therefore little data is available for validation and calibration. We propose a probe geometry and associated mathematical model for measuring muscle oxygenation in seals based on near infrared diffuse transport with no need for calibration. A prototype based on this concept has been designed and tested on humans and rats. We use the test results to discuss the advantages and limitations of the approach. We also detail the constraints on size, sensor location, electronics, light source properties and detector characteristics posed by the unique biology of seals.
López-Cruz, Roberto I.; Crocker, Daniel E.; Gaxiola-Robles, Ramón; Bernal, Jaime A.; Real-Valle, Roberto A.; Lugo-Lugo, Orlando; Zenteno-Savín, Tania
2016-01-01
Marine mammals are exposed to ischemia/reperfusion and hypoxia/reoxygenation during diving. During oxygen deprivation, adenosine triphosphate (ATP) breakdown implies purine metabolite accumulation, which in humans is associated with pathological conditions. Purine recycling in seals increases in response to prolonged fasting and ischemia. Concentrations of metabolites and activities of key enzymes in purine metabolism were examined in plasma and red blood cells from bottlenose dolphins (Tursiops truncatus) and humans. Hypoxanthine and inosine monophosphate concentrations were higher in plasma from dolphins than humans. Plasma hypoxanthine-guanine phosphoribosyl transferase (HGPRT) activity in dolphins suggests an elevated purine recycling rate, and a mechanism for avoiding accumulation of non-recyclable purines (xanthine and uric acid). Red blood cell concentrations of hypoxanthine, adenosine diphosphate, ATP and guanosine triphosphate were lower in dolphins than in humans; adenosine monophosphate and nicotinamide adenine dinucleotide concentrations were higher in dolphins. HGPRT activity in red blood cells was higher in humans than in dolphins. The lower concentrations of purine catabolism and recycling by-products in plasma from dolphins could be beneficial in providing substrates for recovery of ATP depleted during diving or vigorous swimming. These results suggest that purine salvage in dolphins could be a mechanism for delivering nucleotide precursors to tissues with high ATP and guanosine triphosphate requirements. PMID:27375492
López-Cruz, Roberto I; Crocker, Daniel E; Gaxiola-Robles, Ramón; Bernal, Jaime A; Real-Valle, Roberto A; Lugo-Lugo, Orlando; Zenteno-Savín, Tania
2016-01-01
Marine mammals are exposed to ischemia/reperfusion and hypoxia/reoxygenation during diving. During oxygen deprivation, adenosine triphosphate (ATP) breakdown implies purine metabolite accumulation, which in humans is associated with pathological conditions. Purine recycling in seals increases in response to prolonged fasting and ischemia. Concentrations of metabolites and activities of key enzymes in purine metabolism were examined in plasma and red blood cells from bottlenose dolphins (Tursiops truncatus) and humans. Hypoxanthine and inosine monophosphate concentrations were higher in plasma from dolphins than humans. Plasma hypoxanthine-guanine phosphoribosyl transferase (HGPRT) activity in dolphins suggests an elevated purine recycling rate, and a mechanism for avoiding accumulation of non-recyclable purines (xanthine and uric acid). Red blood cell concentrations of hypoxanthine, adenosine diphosphate, ATP and guanosine triphosphate were lower in dolphins than in humans; adenosine monophosphate and nicotinamide adenine dinucleotide concentrations were higher in dolphins. HGPRT activity in red blood cells was higher in humans than in dolphins. The lower concentrations of purine catabolism and recycling by-products in plasma from dolphins could be beneficial in providing substrates for recovery of ATP depleted during diving or vigorous swimming. These results suggest that purine salvage in dolphins could be a mechanism for delivering nucleotide precursors to tissues with high ATP and guanosine triphosphate requirements.
Reactive Oxygen Species, Mitochondria, and Endothelial Cell Death during In Vitro Simulated Dives.
Wang, Qiong; Guerrero, François; Mazur, Aleksandra; Lambrechts, Kate; Buzzacott, Peter; Belhomme, Marac; Theron, Michaël
2015-07-01
Excessive reactive oxygen species (ROS) is considered a consequence of hyperoxia and a major contributor to diving-derived vascular endothelial damage and decompression sickness. The aims of this work were: 1) to directly observe endothelial ROS production during simulated air dives as well as its relation with both mitochondrial activity and cell survival; and 2) to determine which ambient factor during air diving (hydrostatic pressure or oxygen and/or nitrogen partial pressure) is responsible for the observed modifications. In vitro diving simulation was performed with bovine arterial endothelial cells under real-time observation. The effects of air diving, hydrostatic, oxygen and nitrogen pressures, and N-acetylcysteine (NAC) treatment on mitochondrial ROS generation, mitochondrial membrane potential and cellular survival during simulation were investigated. Vascular endothelial cells performing air diving simulation suffered excessive mitochondrial ROS, mitochondrial depolarization, and cell death. These effects were prevented by NAC: after NAC treatment, the cells presented no difference in damage from nondiving cells. Oxygen diving showed a higher effect on ROS generation but lower impacts on mitochondrial depolarization and cell death than hydrostatic or nitrogen diving. Nitrogen diving had no effect on the inductions of ROS, mito-depolarization, or cell death. This study is the first direct observation of mitochondrial ROS production, mitochondrial membrane potential and cell survival during diving. Simulated air SCUBA diving induces excessive ROS production, which leads to mitochondrial depolarization and endothelial cell death. Oxygen partial pressure plays a crucial role in the production of ROS. Deleterious effects of hyperoxia-induced ROS are potentiated by hydrostatic pressure. These findings hold new implications for the pathogenesis of diving-derived endothelial dysfunction.
A field study of the ventilatory response to ambient temperature and pressure in sport diving.
Muller, F L
1995-01-01
This study reports on the relationship between minute ventilation (VE) and environmental variables of temperature (T) and pressure (P) during open water diving. The author conducted a total of 38 dives involving either a light (20 dives) or a moderate (18 dives) level of physical activity. Within each of these groups, P and T taken together accounted for about two thirds of the variance in the VE data. A very significant increase in VE was observed as T decreased (1 < T(degrees C) < 22), and the magnitude of this increase at a given pressure level was similar in the 'light' and the 'moderate' data sets. A second order observation, particularly notable at lower temperature, was the decrease in VE with increasing pressure under conditions of light work. Empirical functions of the from VE = A+B/P n[1 + exp(T - 8)/10], where A, B, and n are adjustable variables, could accommodate both data sets over the whole range of T and P. These results are the first obtained under actual diving conditions to provide evidence for interactions between P, T, and VE. Understanding the physiological mechanisms by which these interactions occur would assist in appreciation of the limitations imposed on scuba divers by the environmental conditions as they affect their ventilatory responses. PMID:8800853
Behavioral responses of big brown bats to dives by praying mantises.
Ghose, Kaushik; Triblehorn, Jeffrey D; Bohn, Kari; Yager, David D; Moss, Cynthia F
2009-03-01
Insectivorous echolocating bats face a formidable array of defenses employed by their airborne prey. One such insect defense is the ultrasound-triggered dive, which is a sudden, rapid drop in altitude, sometimes all the way to the ground. Although many previous studies have investigated the dynamics of such dives and their effect on insect survival rate, there has been little work on how bats may adapt to such an insect defense employed in the middle of pursuit. In this study we investigated how big brown bats (Eptesicus fuscus) adjust their pursuit strategy when flying praying mantises (Parasphendale agrionina) execute evasive, ultrasound-triggered dives. Although the mantis dive occasionally forced the bat to completely abort its chase (25% trials), in a number of cases (75% trials) the bat followed the mantis into the dive. In such cases the bat kept its sonar beam locked onto the target and maneuvered to maintain the same time efficient strategy it adopted during level flight pursuit, though it was ultimately defeated by the dive. This study suggests that although the mantis dive can be effective in evading the bat, it does not always deter the bat from continuing pursuit and, given enough altitude, the bat can potentially capture diving prey using the same flight strategy it employs to intercept prey in level flight.
Yang, Ming; Barak, Otto F; Dujic, Zeljko; Madden, Dennis; Bhopale, Veena M; Bhullar, Jasjeet; Thom, Stephen R
2015-08-15
Predicated on evidence that diving-related microparticle generation is an oxidative stress response, this study investigated the role that oxygen plays in augmenting production of annexin V-positive microparticles associated with open-water SCUBA diving and whether elevations can be abrogated by ascorbic acid. Following a cross-over study design, 14 male subjects ingested placebo and 2-3 wk later ascorbic acid (2 g) daily for 6 days prior to performing either a 47-min dive to 18 m of sea water while breathing air (∼222 kPa N2/59 kPa O2) or breathing a mixture of 60% O2/balance N2 from a tight-fitting face mask at atmospheric pressure for 47 min (∼40 kPa N2/59 kPa O2). Within 30 min after the 18-m dive in the placebo group, neutrophil activation, and platelet-neutrophil interactions occurred, and the total number of microparticles, as well as subgroups bearing CD66b, CD41, CD31, CD142 proteins or nitrotyrosine, increased approximately twofold. No significant elevations occurred among divers after ingesting ascorbic acid, nor were elevations identified in either group after breathing 60% O2. Ascorbic acid had no significant effect on post-dive intravascular bubble production quantified by transthoracic echocardiography. We conclude that high-pressure nitrogen plays a key role in neutrophil and microparticle-associated changes with diving and that responses can be abrogated by dietary ascorbic acid supplementation. Copyright © 2015 the American Physiological Society.
Extracting Databases from Dark Data with DeepDive
Zhang, Ce; Shin, Jaeho; Ré, Christopher; Cafarella, Michael; Niu, Feng
2016-01-01
DeepDive is a system for extracting relational databases from dark data: the mass of text, tables, and images that are widely collected and stored but which cannot be exploited by standard relational tools. If the information in dark data — scientific papers, Web classified ads, customer service notes, and so on — were instead in a relational database, it would give analysts a massive and valuable new set of “big data.” DeepDive is distinctive when compared to previous information extraction systems in its ability to obtain very high precision and recall at reasonable engineering cost; in a number of applications, we have used DeepDive to create databases with accuracy that meets that of human annotators. To date we have successfully deployed DeepDive to create data-centric applications for insurance, materials science, genomics, paleontologists, law enforcement, and others. The data unlocked by DeepDive represents a massive opportunity for industry, government, and scientific researchers. DeepDive is enabled by an unusual design that combines large-scale probabilistic inference with a novel developer interaction cycle. This design is enabled by several core innovations around probabilistic training and inference. PMID:28316365
Breath-hold duration in man and the diving response induced by face immersion.
Sterba, J A; Lundgren, C E
1988-09-01
The objective of this study in 5 selected volunteer subjects was to see whether the circulatory diving response which is elicited by breath holding and by cold water on the face would affect the duration of maximal-effort breath holds. Compared to control measurements (breath holding during resting, breathing with 35 degrees C water on the face) breath holding with the face cooled by 20 degrees C water caused a 12% reduction of heart rate, 6% reduction of cardiac output, 33% reduction in [corrected] forearm blood flow, and 9% rise in mean arterial blood pressure, but there was no difference in breath-hold duration (control and experimental both 94 s). There were also no differences in time of appearance of the first involuntary respiratory efforts during breath holding, in alveolar gas exchange, or in breaking-point alveolar O2 and CO2 tensions. When the diving response was magnified by a brief bout of exercise so that there was a 19% [corrected] reduction in heart rate, 23% reduction in cardiac output, and 48% reduction in forearm blood flow, breath-hold duration was still unaffected by face cooling. Compared to intermittent immersions, continuous exposure of the face to cold water abolished the diving response, probably by a cold adaptation of facial thermal receptors. These results with cooling of the face only are consistent with our earlier finding that there was a negative correlation between the duration of a maximal-effort breath hold and the diving response during whole-body submersion in cold water.
Weingartner, Gundula M.; Thornton, Sheila J.; Andrews, Russel D.; Enstipp, Manfred R.; Barts, Agnieszka D.; Hochachka, Peter W.
2012-01-01
Many phocid seals are expert divers that remain submerged longer than expected based on estimates of oxygen storage and utilization. This discrepancy is most likely due to an overestimation of diving metabolic rate. During diving, a selective redistribution of blood flow occurs, which may result in reduced metabolism in the hypoperfused tissues and a possible decline in whole-body metabolism to below the resting level (hypometabolism). Thyroid hormones are crucial in regulation of energy metabolism in vertebrates and therefore their control might be an important part of achieving a hypometabolic state during diving. To investigate the effect of thyroid hormones on diving physiology of phocid seals, we measured oxygen consumption, heart rate, and post-dive lactate concentrations in five harbor seals (Phoca vitulina) conducting 5 min dives on command, in both euthyroid and experimentally induced hyperthyroid states. Oxygen consumption during diving was significantly reduced (by 25%) in both euthyroid and hyperthyroid states, confirming that metabolic rate during diving falls below resting levels. Hyperthyroidism increased oxygen consumption (by 7–8%) when resting in water and during diving, compared with the euthyroid state, illustrating the marked effect of thyroid hormones on metabolic rate. Consequently, post-dive lactate concentrations were significantly increased in the hyperthyroid state, suggesting that the greater oxygen consumption rates forced seals to make increased use of anaerobic metabolic pathways. During diving, hyperthyroid seals also exhibited a more profound decline in heart rate than seals in the euthyroid state, indicating that these seals were pushed toward their aerobic limit and required a more pronounced cardiovascular response. Our results demonstrate the powerful role of thyroid hormones in metabolic regulation and support the hypothesis that thyroid hormones play a role in modulating the at-sea metabolism of phocid seals. PMID:23060807
Effect of recreational diving on Patagonian rocky reefs.
Bravo, Gonzalo; Márquez, Federico; Marzinelli, Ezequiel M; Mendez, María M; Bigatti, Gregorio
2015-03-01
Tourism has grown considerably in the last decades, promoting activities such as recreational SCUBA diving that may affect marine benthic communities. In Puerto Madryn, Patagonia Argentina, sub-aquatic tourism areas (STA) receive about 7,000 divers per year. Diving is concentrated on a few small rocky reefs and 50% of the dives occur in summer. In this work, we evaluated the effect of recreational diving activities on benthic communities and determined whether diving causes a press (long-term) or a pulse (short-term) response. We quantified the percentage cover of benthic organisms and compared benthic assemblage structure and composition between two sites with contrasting usage by divers, 'highly disturbed' and 'moderately disturbed' sites, and two 'control' sites with similar physical characteristics but no diving activity, twice before and after the diving peak in summer. We found differences in benthic assemblage structure (identity and relative abundance of taxa) and composition (identity only) among diving sites and controls. These differences were consistent before and after the peak of diving in summer, suggesting that recreational diving may produce a press impact on overall benthic assemblage structure and composition in these STA. At the moderately disturbed site, however, covers of specific taxa, such as some key habitat-forming or highly abundant species, usually differed from those in controls only immediately after summer, after which they begun to resemble controls, suggesting a pulse impact. Thus, STA in Golfo Nuevo seem to respond differently to disturbances of diving depending on the usage of the sites. This information is necessary to develop sound management strategies in order to preserve local biodiversity. Copyright © 2014 Elsevier Ltd. All rights reserved.
Evaluating the Effects of Stressors on Immune Function during Simulated Dives in Marine Mammals
2015-04-24
differed the most from all other exposures. In contrast, human cells tended to display either smaller decreases, or increases, in phagocytic activity...phagocytosis similar to baseline samples; decreased phagocytic activity for the dive periods of pressure exposures, with increased activity following the
Physiological Monitoring in Diving Mammals
2014-09-30
currently developing two different units, one based upon the succesful construction of an oximeter used in Weddel seals (Guyton, Stanek et al. 1995), and...Picolas, Germany) is the only commercial driver with variable pulse duration, an output current up to 500 mA, 3 V forward voltage, frequency response...calibrate the oximeter sensor using human blood. Aim 2: Following the first field season in April, 2013, implantation and data collection using
Diving response after a one-week diet and overnight fasting.
Ghiani, Giovanna; Marongiu, Elisabetta; Olla, Sergio; Pinna, Marco; Pusceddu, Matteo; Palazzolo, Girolamo; Sanna, Irene; Roberto, Silvana; Crisafulli, Antonio; Tocco, Filippo
2016-01-01
We hypothesized that overnight fasting after a short dietary period, especially with carbohydrates, could allow performing breath-hold diving with no restraint for diaphragm excursion and blood shift and without any increase of metabolism, and in turn improve the diving response. During two separate sessions, 8 divers carried out two trials: (A) a 30-m depth dive, three hours after a normal breakfast and (B) a dive to the same depth, but after following a diet and fasting overnight. Each test consisted of 3 apnea phases: descent, static and ascent whose durations were measured by a standard chronometer. An impedance cardiograph, housed in an underwater torch, provided data on trans-thoracic fluid index (TFI), stroke volume (SV), heart rate (HR) and cardiac output (CO). Mean blood pressure (MBP), arterial O2 saturation (SaO2), blood glucose (Glu) and blood lactate (BLa) were also collected. In condition B, duration of the static phase of the dive was longer than A (37.8 ± 7.4 vs. 27.3 ± 8.4 s respectively, P < 0.05). In static phases, mean ∆ SV value (difference between basal and nadir values) during fasting was lower than breakfast one (-2.6 ± 5.1 vs. 5.7 ± 7.6 ml, P < 0.05). As a consequence, since mean ∆ HR values were equally decreased in both metabolic conditions, mean ∆ CO value during static after fasting was lower than the same phase after breakfast (-0.4 ± 0.5 vs. 0.4 ± 0.5 L · min(-1) respectively, P < 0.05). At emersion, despite the greater duration of dives during fasting, SaO2 was higher than A (92.0 ± 2.7 vs. 89.4 ± 2.9 % respectively, P < 0.05) and BLa was lower in the same comparison (4.2 ± 0.7 vs. 5.3 ± 1.1 mmol∙L(-1), P < 0.05). An adequate balance between metabolic and splancnic status may improve the diving response during a dive at a depth of 30 m, in safe conditions for the athlete's health.
Behavioral responses of big brown bats to dives by praying mantises
Ghose, Kaushik; Triblehorn, Jeffrey D.; Bohn, Kari; Yager, David D.; Moss, Cynthia F.
2009-01-01
Summary Insectivorous echolocating bats face a formidable array of defenses employed by their airborne prey. One such insect defense is the ultrasound-triggered dive, which is a sudden, rapid drop in altitude, sometimes all the way to the ground. Although many previous studies have investigated the dynamics of such dives and their effect on insect survival rate, there has been little work on how bats may adapt to such an insect defense employed in the middle of pursuit. In this study we investigated how big brown bats (Eptesicus fuscus) adjust their pursuit strategy when flying praying mantises (Parasphendale agrionina) execute evasive, ultrasound-triggered dives. Although the mantis dive occasionally forced the bat to completely abort its chase (25% trials), in a number of cases (75% trials) the bat followed the mantis into the dive. In such cases the bat kept its sonar beam locked onto the target and maneuvered to maintain the same time efficient strategy it adopted during level flight pursuit, though it was ultimately defeated by the dive. This study suggests that although the mantis dive can be effective in evading the bat, it does not always deter the bat from continuing pursuit and, given enough altitude, the bat can potentially capture diving prey using the same flight strategy it employs to intercept prey in level flight. PMID:19218521
Souday, Vincent; Koning, Nick J; Perez, Bruno; Grelon, Fabien; Mercat, Alain; Boer, Christa; Seegers, Valérie; Radermacher, Peter; Asfar, Pierre
2016-01-01
To test the hypothesis whether enriched air nitrox (EAN) breathing during simulated diving reduces decompression stress when compared to compressed air breathing as assessed by intravascular bubble formation after decompression. Human volunteers underwent a first simulated dive breathing compressed air to include subjects prone to post-decompression venous gas bubbling. Twelve subjects prone to bubbling underwent a double-blind, randomized, cross-over trial including one simulated dive breathing compressed air, and one dive breathing EAN (36% O2) in a hyperbaric chamber, with identical diving profiles (28 msw for 55 minutes). Intravascular bubble formation was assessed after decompression using pulmonary artery pulsed Doppler. Twelve subjects showing high bubble production were included for the cross-over trial, and all completed the experimental protocol. In the randomized protocol, EAN significantly reduced the bubble score at all time points (cumulative bubble scores: 1 [0-3.5] vs. 8 [4.5-10]; P < 0.001). Three decompression incidents, all presenting as cutaneous itching, occurred in the air versus zero in the EAN group (P = 0.217). Weak correlations were observed between bubble scores and age or body mass index, respectively. EAN breathing markedly reduces venous gas bubble emboli after decompression in volunteers selected for susceptibility for intravascular bubble formation. When using similar diving profiles and avoiding oxygen toxicity limits, EAN increases safety of diving as compared to compressed air breathing. ISRCTN 31681480.
Johnson, Rebecca
2016-09-01
Some people with well-managed insulin-dependent diabetes can dive safely. Those cleared to participate should control tightly the variables that impact blood glucose levels, including activity, timing, food and insulin. Honest self-assessment is critical. A diabetic diver should cancel a dive if seasick, unusually anxious, or following significant high or low blood glucose levels in the preceding 24 hours. The diver should enter the water with a blood glucose level above 8.3 mmol·L⁻¹ and below 14 mmol·L⁻¹ with a stable or rising trend in blood glucose established with glucose tests at 90, 60, and 30 minutes prior to a dive. The diver should carry emergency glucose at all times and brief dive buddies about hypoglycaemia procedures. This is a personal account of the author's experience diving with type 1 diabetes and details how the UHMS/DAN recommendations are put into practice on dive days. Key elements of the self-assessment process, long- and rapid-acting insulin adjustments, meal timing, responses to blood glucose trends, handling hypoglycaemia and approaching multi-dive days are described. Some considerations for people using insulin pumps are also briefly discussed.
Young, Cole; Reinkensmeyer, David J
2014-08-01
Athletes rely on subjective assessment of complex movements from coaches and judges to improve their motor skills. In some sports, such as diving, snowboard half pipe, gymnastics, and figure skating, subjective scoring forms the basis for competition. It is currently unclear whether this scoring process can be mathematically modeled; doing so could provide insight into what motor skill is. Principal components analysis has been proposed as a motion analysis method for identifying fundamental units of coordination. We used PCA to analyze movement quality of dives taken from USA Diving's 2009 World Team Selection Camp, first identifying eigenpostures associated with dives, and then using the eigenpostures and their temporal weighting coefficients, as well as elements commonly assumed to affect scoring - gross body path, splash area, and board tip motion - to identify eigendives. Within this eigendive space we predicted actual judges' scores using linear regression. This technique rated dives with accuracy comparable to the human judges. The temporal weighting of the eigenpostures, body center path, splash area, and board tip motion affected the score, but not the eigenpostures themselves. These results illustrate that (1) subjective scoring in a competitive diving event can be mathematically modeled; (2) the elements commonly assumed to affect dive scoring actually do affect scoring (3) skill in elite diving is more associated with the gross body path and the effect of the movement on the board and water than the units of coordination that PCA extracts, which might reflect the high level of technique these divers had achieved. We also illustrate how eigendives can be used to produce dive animations that an observer can distort continuously from poor to excellent, which is a novel approach to performance visualization. Copyright © 2014 Elsevier B.V. All rights reserved.
Suzuki, Miwa; Tomoshige, Mika; Ito, Miki; Koga, Sotaro; Yanagisawa, Makio; Bungo, Takashi; Makiguchi, Yuya
2017-07-01
In cetaceans, diving behavior immediately induces a change in blood circulation to favor flow to the brain and heart; this is achieved by intense vasoconstriction of the blood vessels that serve other organs. This blood circulation response is allied to a decrease in heart rate in order to optimize oxygen usage during diving. Vasoconstrictors are present in all mammals and stimulate the contraction of the smooth muscle in the walls of blood vessels. The most important of these vasoconstrictors are the hormones adrenaline (A), noradrenaline (NA), and angiotensin II (ANG II). At present, the contribution of these hormones to vasoconstriction during diving in cetaceans is unclear. To elucidate their possible roles, changes in serum levels of A, NA and ANG II were monitored together with heart rate in the Indo-Pacific bottlenose dolphin Tursiops aduncus during 90 and 180s dives. Both brief diving periods induced an increase in serum NA concentration and a decrease in heart rate; however, no changes were detected in serum levels of A or ANG II. These data indicate that NA may play a role in diving-induced vasoconstriction. Copyright © 2017 Elsevier Inc. All rights reserved.
Smith, Kirby; Scarr, Mark; Scarpaci, Carol
2010-11-01
Humans can dive with critically endangered grey nurse sharks (Carcharias taurus) along the east coast of Australia. This study investigated both compliance of tourist divers to a code of conduct and legislation and the behaviour of grey nurse sharks in the presence of divers. A total of 25 data collection dives were conducted from December 2008 to January 2009. Grey nurse shark and diver behaviour were documented using 2-min scan samples and continuous observation. The proportion of time spent observing human-shark interactions was 9.4% of total field time and mean human-shark interaction time was 15.0 min. Results were used to gauge the effectiveness of current management practices for the grey nurse shark dive industry at Fish Rock in New South Wales, Australia. Grey nurse shark dive tourists were compliant to stipulations in the code of conduct and legislation (compliance ranged from 88 to 100%). The research detailed factors that may promote compliance in wildlife tourism operations such as the clarity of the stipulations, locality of the target species and diver perceptions of sharks. Results indicated that grey nurse sharks spent the majority of their time milling (85%) followed by active swimming (15%). Milling behaviour significantly decreased in the presence of more than six divers. Distance between sharks and divers, interaction time and number of sharks were not significantly correlated with grey nurse shark school behaviour. Jaw gaping, rapid withdrawal and stiff or jerky movement were the specific behaviours of grey nurse sharks that occurred most frequently and were associated with distance between divers and sharks and the presence of six or more divers. Revision of the number of divers allowed per interaction with a school of grey nurse sharks and further research on the potential impacts that shark-diving tourism may pose to grey nurse sharks is recommended.
NASA Astrophysics Data System (ADS)
Smith, Kirby; Scarr, Mark; Scarpaci, Carol
2010-11-01
Humans can dive with critically endangered grey nurse sharks ( Carcharias taurus) along the east coast of Australia. This study investigated both compliance of tourist divers to a code of conduct and legislation and the behaviour of grey nurse sharks in the presence of divers. A total of 25 data collection dives were conducted from December 2008 to January 2009. Grey nurse shark and diver behaviour were documented using 2-min scan samples and continuous observation. The proportion of time spent observing human-shark interactions was 9.4% of total field time and mean human-shark interaction time was 15.0 min. Results were used to gauge the effectiveness of current management practices for the grey nurse shark dive industry at Fish Rock in New South Wales, Australia. Grey nurse shark dive tourists were compliant to stipulations in the code of conduct and legislation (compliance ranged from 88 to 100%). The research detailed factors that may promote compliance in wildlife tourism operations such as the clarity of the stipulations, locality of the target species and diver perceptions of sharks. Results indicated that grey nurse sharks spent the majority of their time milling (85%) followed by active swimming (15%). Milling behaviour significantly decreased in the presence of more than six divers. Distance between sharks and divers, interaction time and number of sharks were not significantly correlated with grey nurse shark school behaviour. Jaw gaping, rapid withdrawal and stiff or jerky movement were the specific behaviours of grey nurse sharks that occurred most frequently and were associated with distance between divers and sharks and the presence of six or more divers. Revision of the number of divers allowed per interaction with a school of grey nurse sharks and further research on the potential impacts that shark-diving tourism may pose to grey nurse sharks is recommended.
NASA Astrophysics Data System (ADS)
German, C. R.; Fornari, D. J.; Fryer, P.; Girguis, P. R.; Humphris, S. E.; Kelley, D. S.; Tivey, M.; Van Dover, C. L.; Von Damm, K.
2012-12-01
In 2013, Alvin returns to service after significant observational and operational upgrades supported by the NSF, NAVSEA & NOAA. Here we review highlights of the first half-century of deep submergence science conducted by Alvin, describe some of the most significant improvements for the new submarine and discuss the importance of these new capabilities for 21st century ocean science and education. Alvin has a long history of scientific exploration, discovery and intervention at the deep seafloor: in pursuit of hypothesis-driven research and in response to human impacts. One of Alvin's earliest achievements, at the height of the Cold War, was to help locate & recover an H-bomb in the Mediterranean, while the last dives completed, just ahead of the current refit, were to investigate the impacts of the Deep Water Horizon oil spill. Alvin has excelled in supporting a range of Earth & Life Science programs including, in the late 1970s, first direct observations and sampling of deep-sea hydrothermal vents and the unusual fauna supported by microbial chemosynthesis. The 1980s saw expansion of Alvin's dive areas to newly discovered hot-springs in the Atlantic & NE Pacific, Alvin's first dives to the wreck of RMS Titanic and its longest excursions away from WHOI yet, via Loihi Seamount (Hawaii) to the Mariana Trench. The 1990s saw Alvin's first event-response dives to sites where volcanic eruptions had just occurred at the East Pacific Rise & Juan de Fuca Ridge while the 2000s saw Alvin discover novel off-axis venting at Lost City. Observations from these dives fundamentally changed our views of volcanic and microbial processes within young ocean crust and even the origins of life! In parallel, new deep submergence capabilities, including manipulative experiments & sensor development, relied heavily on testing using Alvin. Recently, new work has focused on ocean margins where fluid flow from the seafloor results in the release of hydrocarbons and other chemical species that can sustain chemosynthetic seep ecosystems comparable to, and sometimes sharing species with, hot vents. What will Alvin's next 50 years discover? During 2011-12, Alvin has undergone a transformation, including a larger personnel sphere with more & larger viewports to provide improved overlapping fields of view for the pilot & observers. The new Alvin will be certified for operations to 4500m depth initially, but the new sphere will be 6500m-rated and planned future upgrades will ultimately allow the vehicle to dive that deep, enabling human access to 98% of the global ocean floor. This will allow the study of processes and dynamics of Earth's largest ecosystem (the abyssal plains) as well as margin and ridge environments and the overlying water column. Meantime, the current upgrades to Alvin already include a suite of scientific enhancements including new HD video & still imaging, sophisticated data acquisition systems for seafloor observations and mapping, a new work platform with greater payload capacity and improved observer ergonomics. The new Alvin is poised to play important roles in core Earth and Life science programs and to serve large-scale programs such as the Ocean Observatory Initiative (OOI) and the International Ocean Discovery Program (IODP). It will continue to attract, engage and inspire a new generation of scientists & students to explore and study the largest ecosystem on Earth, just as it has done throughout its first half century.
Wright, Brianna M; Ford, John K B; Ellis, Graeme M; Deecke, Volker B; Shapiro, Ari Daniel; Battaile, Brian C; Trites, Andrew W
2017-01-01
We sought to quantitatively describe the fine-scale foraging behavior of northern resident killer whales ( Orcinus orca ), a population of fish-eating killer whales that feeds almost exclusively on Pacific salmon ( Oncorhynchus spp.). To reconstruct the underwater movements of these specialist predators, we deployed 34 biologging Dtags on 32 individuals and collected high-resolution, three-dimensional accelerometry and acoustic data. We used the resulting dive paths to compare killer whale foraging behavior to the distributions of different salmonid prey species. Understanding the foraging movements of these threatened predators is important from a conservation standpoint, since prey availability has been identified as a limiting factor in their population dynamics and recovery. Three-dimensional dive tracks indicated that foraging ( N = 701) and non-foraging dives ( N = 10,618) were kinematically distinct (Wilks' lambda: λ 16 = 0.321, P < 0.001). While foraging, killer whales dove deeper, remained submerged longer, swam faster, increased their dive path tortuosity, and rolled their bodies to a greater extent than during other activities. Maximum foraging dive depths reflected the deeper vertical distribution of Chinook (compared to other salmonids) and the tendency of Pacific salmon to evade predators by diving steeply. Kinematic characteristics of prey pursuit by resident killer whales also revealed several other escape strategies employed by salmon attempting to avoid predation, including increased swimming speeds and evasive maneuvering. High-resolution dive tracks reconstructed using data collected by multi-sensor accelerometer tags found that movements by resident killer whales relate significantly to the vertical distributions and escape responses of their primary prey, Pacific salmon.
NASA Astrophysics Data System (ADS)
Kokubun, Nobuo; Yamamoto, Takashi; Sato, Nobuhiko; Watanuki, Yutaka; Will, Alexis; Kitaysky, Alexander S.; Takahashi, Akinori
2016-04-01
Subarctic environmental changes are expected to affect the foraging ecology of marine top predators, but the response to such changes may vary among species if they use food resources differently. We examined the characteristics of foraging behavior of two sympatric congeneric diving seabird: common (Uria aalge: hereafter COMUs) and thick-billed (U. lomvia: hereafter TBMUs) murres breeding on St. George Island, located in the seasonal sea-ice region of the Bering Sea. We investigated their foraging trip and flight durations, diel patterns of dive depth, and underwater wing strokes, along with wing morphology and blood stable isotope signatures and stress hormones. Acceleration-temperature-depth loggers were attached to chick-guarding birds, and data were obtained from 7 COMUs and 12 TBMUs. Both species showed similar mean trip duration (13.2 h for COMUs and 10.5 h for TBMUs) and similar diurnal patterns of diving (frequent dives to various depths in the daytime and less frequent dives to shallow depths in the nighttime). During the daytime, the dive depths of COMUs had two peaks in shallow (18.1 m) and deep (74.2 m) depths, while those of TBMUs were 20.2 m and 59.7 m. COMUs showed more frequent wing strokes during the bottom phase of dives (1.90 s-1) than TBMUs (1.66 s-1). Fish occurred more frequently in the bill loads of COMUs (85 %) than those of TBMUs (56 %). The δ15N value of blood was significantly higher in COMUs (14.5 ‰) than in TBMUs (13.1 ‰). The relatively small wing area (0.053 m2) of COMUs compared to TBMUs (0.067 m2) may facilitate their increased agility while foraging and allow them to capture more mobile prey such as larger fishes that inhabit deeper depths. These differences in food resource use may lead to the differential responses of the two murre species to marine environmental changes in the Bering Sea.
Glen, S
2004-01-01
Background: The need for routine medical examinations of sport divers in the Scottish Sub-Aqua Club (Scot-SAC) was revised in March 2000, and a new system using a self administered screening questionnaire was developed to allow divers to be assessed when necessary by doctors with diving medicine experience. Objective: To assess the effect of the new medical system on medical referee workload, diver exclusion rates, and diving incident frequency. Methods: All divers were required to complete a questionnaire to screen for conditions that might affect fitness to dive. Divers answering "Yes" to any of the questions had their medical background assessed by a diving doctor, and, if necessary, received a clinical examination or investigation. The rate of diver exclusions based on the questionnaire response was recorded in conjunction with analysis of the incident reports. Results: The number of forms requiring review by diving doctors increased from 1.2% to 5.7% (p<0.0001, 95% confidence interval (CI) –0.06 to –0.03) in the year after the introduction of the new medical system and gradually increased in subsequent years to 7.7% (p<0.0001, 95% CI –0.08 to –0.05). The number of divers failing to be certified fit to dive increased slightly from 0.7% to 1.0% after one year (p = 0.26, 95% CI –0.01 to 0.00) and subsequently to 2.0% (p = 0.0003, 95% CI 0.02 to –0.01) after three years. Most divers were certified fit to dive on the basis of the questionnaire alone, and only 0.9% required objective investigation (such as exercise testing or echocardiography). Analysis of the incidents during three years of follow up confirmed that no incident occurred because of an undetected pre-existing medical condition. Two incidents involved divers with hypertension, but both had received medical examinations and investigation based on their responses to the questionnaire. Conclusion: The new self administered questionnaire system appears to be an effective screening tool for the detection of divers requiring detailed assessment by doctors with diving medicine experience. PMID:15562174
ERIC Educational Resources Information Center
Braus, Judy, Ed.
1992-01-01
Ranger Rick's NatureScope is a creative education series dedicated to inspiring in children an understanding and appreciation of the natural world while developing the skills they will need to make responsible decisions about the environment. The topic of this issue is "Diving Into Oceans." Contents are organized into the following…
Del Castillo Velasco-Martínez, Iris; Hernández-Camacho, Claudia J; Méndez-Rodríguez, Lía C; Zenteno-Savín, Tania
2016-01-01
In mammalian tissues under hypoxic conditions, ATP degradation results in accumulation of purine metabolites. During exercise, muscle energetic demand increases and oxygen consumption can exceed its supply. During breath-hold diving, oxygen supply is reduced and, although oxygen utilization is regulated by bradycardia (low heart rate) and peripheral vasoconstriction, tissues with low blood flow (ischemia) may become hypoxic. The goal of this study was to evaluate potential differences in the circulating levels of purine metabolism components between diving and exercise in bottlenose dolphins (Tursiops truncatus). Blood samples were taken from captive dolphins following a swimming routine (n=8) and after a 2min dive (n=8). Activity of enzymes involved in purine metabolism (hypoxanthine guanine phosphoribosyl transferase (HGPRT), inosine monophosphate deshydrogenase (IMPDH), xanthine oxidase (XO), purine nucleoside phosphorylase (PNP)), and purine metabolite (hypoxanthine (HX), xanthine (X), uric acid (UA), inosine monophosphate (IMP), inosine, nicotinamide adenine dinucleotide (NAD(+)), adenosine, adenosine monophosphate (AMP), adenosine diphosphate (ADP), ATP, guanosine diphosphate (GDP), guanosine triphosphate (GTP)) concentrations were quantified in erythrocyte and plasma samples. Enzymatic activity and purine metabolite concentrations involved in purine synthesis and degradation, were not significantly different between diving and exercise. Plasma adenosine concentration was higher after diving than exercise (p=0.03); this may be related to dive-induced ischemia. In erythrocytes, HGPRT activity was higher after diving than exercise (p=0.007), suggesting an increased capacity for purine recycling and ATP synthesis from IMP in ischemic tissues of bottlenose dolphins during diving. Purine recycling and physiological adaptations may maintain the ATP concentrations in bottlenose dolphins after diving and exercise. Copyright © 2015 Elsevier Inc. All rights reserved.
Aerobic exercise before diving reduces venous gas bubble formation in humans
Dujić, Željko; Duplančic, Darko; Marinovic-Terzić, Ivana; Baković, Darija; Ivančev, Vladimir; Valic, Zoran; Eterović, Davor; Petri, Nadan M; Wisløff, Ulrik; Brubakk, Alf O
2004-01-01
We have previously shown in a rat model that a single bout of high-intensity aerobic exercise 20h before a simulated dive reduces bubble formation and after the dive protects from lethal decompression sickness. The present study investigated the importance of these findings in man. Twelve healthy male divers were compressed in a hyperbaric chamber to 280kPa at a rate of 100kPamin−1 breathing air and remaining at pressure for 80min. The ascent rate was 9mmin−1 with a 7min stop at 130kPa. Each diver underwent two randomly assigned simulated dives, with or without preceding exercise. A single interval exercise performed 24h before the dive consisted of treadmill running at 90% of maximum heart rate for 3min, followed by exercise at 50% of maximum heart rate for 2min; this was repeated eight times for a total exercise period of 40min. Venous gas bubbles were monitored with an ultrasonic scanner every 20min for 80min after reaching surface pressure. The study demonstrated that a single bout of strenuous exercise 24h before a dive to 18 m of seawater significantly reduced the average number of bubbles in the pulmonary artery from 0.98 to 0.22 bubbles cm−2(P= 0.006) compared to dives without preceding exercise. The maximum bubble grade was decreased from 3 to 1.5 (P= 0.002) by pre-dive exercise, thereby increasing safety. This is the first report to indicate that pre-dive exercise may form the basis for a new way of preventing serious decompression sickness. PMID:14755001
Mechanics of airway and alveolar collapse in human breath-hold diving.
Fitz-Clarke, John R
2007-11-15
A computational model of the human respiratory tract was developed to study airway and alveolar compression and re-expansion during deep breath-hold dives. The model incorporates the chest wall, supraglottic airway, trachea, branched airway tree, and elastic alveoli assigned time-dependent surfactant properties. Total lung collapse with degassing of all alveoli is predicted to occur around 235 m, much deeper than estimates for aquatic mammals. Hysteresis of the pressure-volume loop increases with maximum diving depth due to progressive alveolar collapse. Reopening of alveoli occurs stochastically as airway pressure overcomes adhesive and compressive forces on ascent. Surface area for gas exchange vanishes at collapse depth, implying that the risk of decompression sickness should reach a plateau beyond this depth. Pulmonary capillary transmural stresses cannot increase after local alveolar collapse. Consolidation of lung parenchyma might provide protection from capillary injury or leakage caused by vascular engorgement due to outward chest wall recoil at extreme depths.
Project Tektite 1: A multiagency 60-day saturated dive
NASA Technical Reports Server (NTRS)
Pauli, D. C. (Editor); Cole, H. A. (Editor)
1970-01-01
The history, organization, and accomplishments of Project Tektite are presented. The project demonstrated that it is possible for men to live and work successfully at underwater depths of 49 feet. The biomedical responses of the aquanauts to the 60 day dive are described. The behavioral and psychological responses to each other, the work, and the isolated environment are reported. The experiments conducted in the fields of marine science, psychological sciences, and biomedical sciences are discussed.
Probability of Decompression Sickness in No-Stop Air Diving
2004-12-01
21 Figure 10. VVal-1 8 and StandAir Models .......................................... 22 Figure 11. Comparisons for...recommendations appear in heavy boxes. Information outside the heavy boxes allows comparisons between models. The recommendations are essentially arbitrary and...N2-0 2 Saturation Dives in Humans: DCS Risk and Evidence of a Threshold," Undersea Hyperbaric Medicine, In Press. 15. S. S. Survanshi, E. D. Parker, E
Viviant, Morgane; Monestiez, Pascal; Guinet, Christophe
2014-01-01
Predicting how climatic variations will affect marine predator populations relies on our ability to assess foraging success, but evaluating foraging success in a marine predator at sea is particularly difficult. Dive metrics are commonly available for marine mammals, diving birds and some species of fish. Bottom duration or dive duration are usually used as proxies for foraging success. However, few studies have tried to validate these assumptions and identify the set of behavioral variables that best predict foraging success at a given time scale. The objective of this study was to assess if foraging success in Antarctic fur seals could be accurately predicted from dive parameters only, at different temporal scales. For this study, 11 individuals were equipped with either Hall sensors or accelerometers to record dive profiles and detect mouth-opening events, which were considered prey capture attempts. The number of prey capture attempts was best predicted by descent and ascent rates at the dive scale; bottom duration and descent rates at 30-min, 1-h, and 2-h scales; and ascent rates and maximum dive depths at the all-night scale. Model performances increased with temporal scales, but rank and sign of the factors varied according to the time scale considered, suggesting that behavioral adjustment in response to prey distribution could occur at certain scales only. The models predicted the foraging intensity of new individuals with good accuracy despite high inter-individual differences. Dive metrics that predict foraging success depend on the species and the scale considered, as verified by the literature and this study. The methodology used in our study is easy to implement, enables an assessment of model performance, and could be applied to any other marine predator. PMID:24603534
Chimienti, Marianna; Cornulier, Thomas; Owen, Ellie; Bolton, Mark; Davies, Ian M; Travis, Justin M J; Scott, Beth E
2017-12-01
Detailed information acquired using tracking technology has the potential to provide accurate pictures of the types of movements and behaviors performed by animals. To date, such data have not been widely exploited to provide inferred information about the foraging habitat. We collected data using multiple sensors (GPS, time depth recorders, and accelerometers) from two species of diving seabirds, razorbills ( Alca torda , N = 5, from Fair Isle, UK) and common guillemots ( Uria aalge , N = 2 from Fair Isle and N = 2 from Colonsay, UK). We used a clustering algorithm to identify pursuit and catching events and the time spent pursuing and catching underwater, which we then used as indicators for inferring prey encounters throughout the water column and responses to changes in prey availability of the areas visited at two levels: individual dives and groups of dives. For each individual dive ( N = 661 for guillemots, 6214 for razorbills), we modeled the number of pursuit and catching events, in relation to dive depth, duration, and type of dive performed (benthic vs. pelagic). For groups of dives ( N = 58 for guillemots, 156 for razorbills), we modeled the total time spent pursuing and catching in relation to time spent underwater. Razorbills performed only pelagic dives, most likely exploiting prey available at shallow depths as indicated by the vertical distribution of pursuit and catching events. In contrast, guillemots were more flexible in their behavior, switching between benthic and pelagic dives. Capture attempt rates indicated that they were exploiting deep prey aggregations. The study highlights how novel analysis of movement data can give new insights into how animals exploit food patches, offering a unique opportunity to comprehend the behavioral ecology behind different movement patterns and understand how animals might respond to changes in prey distributions.
Spisni, Enzo; Marabotti, Claudio; De Fazio, Luigia; Valerii, Maria Chiara; Cavazza, Elena; Brambilla, Stefano; Hoxha, Klarida; L'Abbate, Antonio; Longobardi, Pasquale
2017-03-01
The aim of this study was to compare two decompression procedures commonly adopted by technical divers: the ZH-L16 algorithm modified by 30/85 gradient factors (compartmental decompression model, CDM) versus the 'ratio decompression strategy' (RDS). The comparison was based on an analysis of changes in diver circulating inflammatory profiles caused by decompression from a single dive. Fifty-one technical divers performed a single trimix dive to 50 metres' sea water (msw) for 25 minutes followed by enriched air (EAN50) and oxygen decompression. Twenty-three divers decompressed according to a CDM schedule and 28 divers decompressed according to a RDS schedule. Peripheral blood for detection of inflammatory markers was collected before and 90 min after diving. Venous gas emboli were measured 30 min after diving using 2D echocardiography. Matched groups of 23 recreational divers (dive to 30 msw; 25 min) and 25 swimmers were also enrolled as control groups to assess the effects of decompression from a standard air dive or of exercise alone on the inflammatory profile. Echocardiography at the single 30 min observation post dive showed no significant differences between the two decompression procedures. Divers adopting the RDS showed a worsening of post-dive inflammatory profile compared to the CDM group, with significant increases in circulating chemokines CCL2 (P = 0.001) and CCL5 (P = 0.006) levels. There was no increase in chemokines following the CDM decompression. The air scuba group also showed a statistically significant increase in CCL2 (P < 0.001) and CCL5 (P = 0.003) levels post dive. No cases of decompression sickness occurred. The ratio deco strategy did not confer any benefit in terms of bubbles but showed the disadvantage of increased decompression-associated secretion of inflammatory chemokines involved in the development of vascular damage.
Determinants of arterial gas embolism after scuba diving.
Ljubkovic, Marko; Zanchi, Jaksa; Breskovic, Toni; Marinovic, Jasna; Lojpur, Mihajlo; Dujic, Zeljko
2012-01-01
Scuba diving is associated with breathing gas at increased pressure, which often leads to tissue gas supersaturation during ascent and the formation of venous gas emboli (VGE). VGE crossover to systemic arteries (arterialization), mostly through the patent foramen ovale, has been implicated in various diving-related pathologies. Since recent research has shown that arterializations frequently occur in the absence of cardiac septal defects, our aim was to investigate the mechanisms responsible for these events. Divers who tested negative for patent foramen ovale were subjected to laboratory testing where agitated saline contrast bubbles were injected in the cubital vein at rest and exercise. The individual propensity for transpulmonary bubble passage was evaluated echocardiographically. The same subjects performed a standard air dive followed by an echosonographic assessment of VGE generation (graded on a scale of 0-5) and distribution. Twenty-three of thirty-four subjects allowed the transpulmonary passage of saline contrast bubbles in the laboratory at rest or after a mild/moderate exercise, and nine of them arterialized after a field dive. All subjects with postdive arterialization had bubble loads reaching or exceeding grade 4B in the right heart. In individuals without transpulmonary passage of saline contrast bubbles, injected either at rest or after an exercise bout, no postdive arterialization was detected. Therefore, postdive VGE arterialization occurs in subjects that meet two criteria: 1) transpulmonary shunting of contrast bubbles at rest or at mild/moderate exercise and 2) VGE generation after a dive reaches the threshold grade. These findings may represent a novel concept in approach to diving, where diving routines will be tailored individually.
Bender, Renato; Bender, Nicole
2013-09-01
Extant hominoids, including humans, are well known for their inability to swim instinctively. We report swimming and diving in two captive apes using visual observation and video recording. One common chimpanzee and one orangutan swam repeatedly at the water surface over a distance of 2-6 m; both individuals submerged repeatedly. We show that apes are able to overcome their negative buoyancy by deliberate swimming, using movements which deviate from the doggy-paddle pattern observed in other primates. We suggest that apes' poor swimming ability is due to behavioral, anatomical, and neuromotor changes related to an adaptation to arboreal life in their early phylogeny. This strong adaptive focus on arboreal life led to decreased opportunities to interact with water bodies and consequently to a reduction of selective pressure to maintain innate swimming behavior. As the doggy paddle is associated with quadrupedal walking, a deviation from terrestrial locomotion might have interfered with the fixed rhythmic action patterns responsible for innate swimming. Copyright © 2013 Wiley Periodicals, Inc.
Prediction of human adaptation and performance in underwater environments.
Colodro Plaza, Joaquín; Garcés de los Fayos Ruiz, Enrique J; López García, Juan J; Colodro Conde, Lucía
2014-01-01
Environmental stressors require the professional diver to undergo a complex process of psychophysiological adaptation in order to overcome the demands of an extreme environment and carry out effective and efficient work under water. The influence of cognitive and personality traits in predicting underwater performance and adaptation has been a common concern for diving psychology, and definitive conclusions have not been reached. In this ex post facto study, psychological and academic data were analyzed from a large sample of personnel participating in scuba diving courses carried out in the Spanish Navy Diving Center. In order to verify the relevance of individual differences in adaptation to a hostile environment, we evaluated the predictive validity of general mental ability and personality traits with regression techniques. The data indicated the existence of psychological variables that can predict the performance ( R² = .30, p <.001) and adaptation ( R²(N) = .51, p <.001) of divers in underwater environment. These findings support the hypothesis that individual differences are related to the probability of successful adaptation and effective performance in professional diving. These results also verify that dispositional traits play a decisive role in diving training and are significant factors in divers' psychological fitness.
2016-11-01
ORGANIZATION NAME(S) AND ADDRESS(ES) 8. PERFORMING ORGANIZATION Biology & Marine Biology , UNCW REPORT NUMBER 601 S. College Road Wi lmington, NC...Information PI: Dr. Heather Koopman Institution: Biology & Marine Biology , University of North Carolina Wilmington Award #: N000141210442 Title...both graduate and undergraduate) • 1 paper published in Journal of Experimental Biology (and picked up by the media; see X. G. below); 3 additional
Do recreational activities affect coastal biodiversity?
NASA Astrophysics Data System (ADS)
Riera, Rodrigo; Menci, Cristiano; Sanabria-Fernández, José Antonio; Becerro, Mikel A.
2016-09-01
Human activities are largely affecting coastal communities worldwide. Recreational perturbations have been overlooked in comparison to other perturbations, yet they are potential threats to marine biodiversity. They affect coastal communities in different ways, underpinning consistent shifts in fish and invertebrates assemblages. Several sites were sampled subjected to varying effects by recreational fishermen (low and high pressure) and scuba divers (low and high) in an overpopulated Atlantic island. Non-consistent differences in ecological, trophic and functional diversity were found in coastal communities, considering both factors (;diving; and ;fishing;). Multivariate analyses only showed significant differences in benthic invertebrates between intensively-dived and non-dived sites. The lack of clear trends may be explained by the depletion of coastal resources in the study area, an extensively-affected island by overfishing.
Naito, Yasuhiko; Costa, Daniel P; Adachi, Taiki; Robinson, Patrick W; Peterson, Sarah H; Mitani, Yoko; Takahashi, Akinori
2017-08-01
Little is known about the foraging behavior of top predators in the deep mesopelagic ocean. Elephant seals dive to the deep biota-poor oxygen minimum zone (OMZ) (>800 m depth) despite high diving costs in terms of energy and time, but how they successfully forage in the OMZ remains largely unknown. Assessment of their feeding rate is the key to understanding their foraging behavior, but this has been challenging. Here, we assessed the feeding rate of 14 female northern elephant seals determined by jaw motion events (JME) and dive cycle time to examine how feeding rates varied with dive depth, particularly in the OMZ. We also obtained video footage from seal-mounted videos to understand their feeding in the OMZ. While the diel vertical migration pattern was apparent for most depths of the JME, some very deep dives, beyond the normal diel depth ranges, occurred episodically during daylight hours. The midmesopelagic zone was the main foraging zone for all seals. Larger seals tended to show smaller numbers of JME and lower feeding rates than smaller seals during migration, suggesting that larger seals tended to feed on larger prey to satisfy their metabolic needs. Larger seals also dived frequently to the deep OMZ, possibly because of a greater diving ability than smaller seals, suggesting their dependency on food in the deeper depth zones. Video observations showed that seals encountered the rarely reported ragfish ( Icosteus aenigmaticus ) in the depths of the OMZ, which failed to show an escape response from the seals, suggesting that low oxygen concentrations might reduce prey mobility. Less mobile prey in OMZ would enhance the efficiency of foraging in this zone, especially for large seals that can dive deeper and longer. We suggest that the OMZ plays an important role in structuring the mesopelagic ecosystem and for the survival and evolution of elephant seals.
Williams, Terrie M; Kendall, Traci L; Richter, Beau P; Ribeiro-French, Courtney R; John, Jason S; Odell, Kim L; Losch, Barbara A; Feuerbach, David A; Stamper, M Andrew
2017-03-15
Exponential increases in hydrodynamic drag and physical exertion occur when swimmers move quickly through water, and underlie the preference for relatively slow routine speeds by marine mammals regardless of body size. Because of this and the need to balance limited oxygen stores when submerged, flight (escape) responses may be especially challenging for this group. To examine this, we used open-flow respirometry to measure the energetic cost of producing a swimming stroke during different levels of exercise in bottlenose dolphins ( Tursiops truncatus ). These data were then used to model the energetic cost of high-speed escape responses by other odontocetes ranging in mass from 42 to 2738 kg. The total cost per stroke during routine swimming by dolphins, 3.31±0.20 J kg -1 stroke -1 , was doubled during maximal aerobic performance. A comparative analysis of locomotor costs (LC; in J kg -1 stroke -1 ), representing the cost of moving the flukes, revealed that LC during routine swimming increased with body mass ( M ) for odontocetes according to LC=1.46±0.0005 M ; a separate relationship described LC during high-speed stroking. Using these relationships, we found that continuous stroking coupled with reduced glide time in response to oceanic noise resulted in a 30.5% increase in metabolic rate in the beaked whale, a deep-diving odontocete considered especially sensitive to disturbance. By integrating energetics with swimming behavior and dive characteristics, this study demonstrates the physiological consequences of oceanic noise on diving mammals, and provides a powerful tool for predicting the biological significance of escape responses by cetaceans facing anthropogenic disturbances. © 2017. Published by The Company of Biologists Ltd.
Biochemical and clinical responses of Common Eiders to implanted satellite transmitters
Latty, Christopher J.; Hollmen, Tuula E.; Petersen, Margaret; Powell, Abby; Andrews, Russel D.
2016-01-01
Implanted biologging devices, such as satellite-linked platform transmitter terminals (PTTs), have been used widely to delineate populations and identify movement patterns of sea ducks. Although in some cases these ecological studies could reveal transmitter effects on behavior and mortality, experiments conducted under controlled conditions can provide valuable information to understand the influence of implanted tags on health and physiology. We report the clinical, mass, biochemical, and histological responses of captive Common Eiders (Somateria mollissima) implanted with PTTs with percutaneous antennas. We trained 6 individuals to dive 4.9 m for their food, allowed them to acclimate to this dive depth, and implanted them with PTTs. We collected data before surgery to establish baselines, and for 3.5 mo after surgery. The first feeding dive took place 22 hr after surgery, with 5 of 6 birds diving to the bottom within 35 hr of surgery. Plumage waterproofing around surgical sites was reduced ≤21 days after surgery. Mass; albumin; albumin:globulin ratio; aspartate aminotransferase; β1-, β2-, and γ-globulins; creatine kinase; fecal glucocorticoid metabolites; heterophil:lymphocyte ratio; and packed cell volume changed from baseline on one or more of the postsurgery sampling dates, and some changes were still evident 3.5 mo after surgery. Our findings show that Common Eiders physiologically responded for up to 3.5 mo after surgical implantation of a PTT, with the greatest response occurring within the first few weeks of implantation. These responses support the need for postsurgery censor periods for satellite telemetry data and should be considered when designing studies and analyzing information from PTTs in sea ducks.
Gut fermentation seems to promote decompression sickness in humans.
de Maistre, Sébastien; Vallee, Nicolas; Gempp, Emmanuel; Louge, Pierre; Duchamp, Claude; Blatteau, Jean-Eric
2016-10-01
Massive bubble formation after diving can lead to decompression sickness (DCS) that can result in neurological disorders. In experimental dives using hydrogen as the diluent gas, decreasing the body's H 2 burden by inoculating hydrogen-metabolizing microbes into the gut reduces the risk of DCS. In contrast, we have shown that gut bacterial fermentation in rats on a standard diet promotes DCS through endogenous hydrogen production. Therefore, we set out to test these experimental results in humans. Thirty-nine divers admitted into our hyperbaric center with neurological DCS (Affected Divers) were compared with 39 healthy divers (Unaffected Divers). Their last meal time and composition were recorded. Gut fermentation rate was estimated by measuring breath hydrogen 1-4 h after the dive. Breath hydrogen concentrations were significantly higher in Affected Divers (15 ppm [6-23] vs. 7 ppm [3-12]; P = 0.0078). With the use of a threshold value of 16.5 ppm, specificity was 87% [95% confidence interval (CI) 73-95] for association with neurological DCS onset. We observed a strong association between hydrogen values above this threshold and an accident occurrence (odds ratio = 5.3, 95% CI 1.8-15.7, P = 0.0025). However, high fermentation potential foodstuffs consumption was not different between Affected and Unaffected Divers. Gut fermentation rate at dive time seemed to be higher in Affected Divers. Hydrogen generated by fermentation diffuses throughout the body and could increase DCS risk. Prevention could be helped by excluding divers who are showing a high fermentation rate, by eliminating gas produced in gut, or even by modifying intestinal microbiota to reduce fermentation rate during a dive. Copyright © 2016 the American Physiological Society.
Increasing the probability of surviving loss of consciousness underwater when using a rebreather.
Haynes, Paul
2016-12-01
Re-circulating underwater breathing apparatus (rebreathers) have become increasingly popular amongst sport divers. In comparison to open-circuit scuba, rebreathers are complex life support equipment that incorporates many inherent failure modes and potential for human error. This individually or in combination can lead to an inappropriate breathing gas. Analysis of rebreather diving incidents suggests that inappropriate breathing gas is the most prevalent disabling agent. This can result in spontaneous loss of consciousness (LoC), water aspiration and drowning. Protecting the airway by maintaining the diver/rebreather oral interface may delay water aspiration following LoC underwater; the possibility of a successful rescue is, thus, increased. One means of protecting the airway following LoC underwater is the use of a full-face mask (FFM). However, such masks are complex and expensive; therefore, they have not been widely adopted by the sport diving community. An alternative to the FFM used extensively throughout the global military diving community is the mouthpiece retaining strap (MRS). A recent study documented 54 LoC events in military rebreather diving with only three consequent drownings; all divers were reported to be using a MRS. Even allowing for the concomitant use of a tethered diving partner system in most cases, the low number of fatalities in this large series is circumstantially supportive of the efficacy of the MRS. Despite drowning featuring as a final common pathway in the vast majority of rebreather fatalities, the MRS has not been widely adopted by the sport rebreather diving community.
A comparison of auditory brainstem responses across diving bird species
Crowell, Sara E.; Berlin, Alicia; Carr, Catherine E.; Olsen, Glenn H.; Therrien, Ronald E.; Yannuzzi, Sally E.; Ketten, Darlene R.
2015-01-01
There is little biological data available for diving birds because many live in hard-to-study, remote habitats. Only one species of diving bird, the black-footed penguin (Spheniscus demersus), has been studied in respect to auditory capabilities (Wever et al., Proc Natl Acad Sci USA 63:676–680, 1969). We, therefore, measured in-air auditory threshold in ten species of diving birds, using the auditory brainstem response (ABR). The average audiogram obtained for each species followed the U-shape typical of birds and many other animals. All species tested shared a common region of the greatest sensitivity, from 1000 to 3000 Hz, although audiograms differed significantly across species. Thresholds of all duck species tested were more similar to each other than to the two non-duck species tested. The red-throated loon (Gavia stellata) and northern gannet (Morus bassanus) exhibited the highest thresholds while the lowest thresholds belonged to the duck species, specifically the lesser scaup (Aythya affinis) and ruddy duck (Oxyura jamaicensis). Vocalization parameters were also measured for each species, and showed that with the exception of the common eider (Somateria mollisima), the peak frequency, i.e., frequency at the greatest intensity, of all species' vocalizations measured here fell between 1000 and 3000 Hz, matching the bandwidth of the most sensitive hearing range.
A comparison of auditory brainstem responses across diving bird species
Crowell, Sara E.; Wells-Berlin, Alicia M.; Carr, Catherine E.; Olsen, Glenn H.; Therrien, Ronald E.; Yannuzzi, Sally E.; Ketten, Darlene R.
2015-01-01
There is little biological data available for diving birds because many live in hard-to-study, remote habitats. Only one species of diving bird, the black-footed penguin (Spheniscus demersus), has been studied in respect to auditory capabilities (Wever et al. 1969). We therefore measured in-air auditory threshold in ten species of diving birds, using the auditory brainstem response (ABR). The average audiogram obtained for each species followed the U-shape typical of birds and many other animals. All species tested shared a common region of greatest sensitivity, from 1000 to 3000 Hz, although audiograms differed significantly across species. Thresholds of all duck species tested were more similar to each other than to the two non-duck species tested. The red-throated loon (Gavia stellata) and northern gannet (Morus bassanus) exhibited the highest thresholds while the lowest thresholds belonged to the duck species, specifically the lesser scaup (Aythya affinis) and ruddy duck (Oxyura jamaicensis). Vocalization parameters were also measured for each species, and showed that with the exception of the common eider (Somateria mollisima), the peak frequency, i.e. frequency at the greatest intensity, of all species’ vocalizations measured here fell between 1000 and 3000 Hz, matching the bandwidth of the most sensitive hearing range. PMID:26156644
1988-08-01
of the equipment, but NCSC may have non-obstructive life cycle instrumentation attached. NEDU will provide a suitable vehicle to transport the LWDS...CF b. Diving Supervisor - QMCS( MDV ) T.R. Griggs, USN c. Project Medical Department Representative - LCDR M. T. Wallick, MSC, USNU d. Duty Medical...NEDU using a vehicle . Notes on transportability will be made. Per the program of ANNEX A, NCSC will instruct the NEDU team in the operation of the LWDS
1977-11-01
in oceanography, marine sL veying and marine engineering. The elective subjects cover such topics as hydrographic sur- veying, sounding , underwater...and Diving in Mr. J. Tomlinson D.F.E. Department of Fisheries and Marine Services L.Portable System for Ultrasonic Mr. B. EAtock, DCIEM Doppler...semi-tropical and temperate environments (1). As with all homeothermic mammals , the human body has a deep internal temperattre of approximately 37 0
Effects of growth, diving history, and high altitude on blood oxygen capacity in harbor seals
NASA Technical Reports Server (NTRS)
Kodama, A. M.; Elsner, R.; Pace, N.
1977-01-01
Blood volume and body composition for diving and nondiving harbor seals were measured at six-week intervals during a 10-month period of captitivity. Whole body hematocrit, red cell volume per kg of lean body mass, and total circulating hemoglobin per kg lean body mass were significantly higher in the diving group, but relatively large blood volumes expressed in terms of body weight (11-12%) were found in both groups. A pair of harbor seals exposed to high altitude for about three months registered significant increases in red cell volume, blood hemoglobin levels, and blood volume expressed in terms of body weight; results of alveolar gas analyses indicate that hyperventilation also occurred. These typical mammalian responses to hypoxia suggest that the harbor seal's large blood volume and high hemoglobin content are an expression of phylogenetic control, and that in spite of its adaptability to apnea during its diving life, the animal cannot be considered preacclimatized to high altitude.
T-dependence of human B lymphocyte proliferative response to mitogens.
Brochier, J; Samarut, C; Gueho, J P; Revillard, J P
1976-01-01
Human peripheral blood and tonsil lymphocytes were fractionated on anti-Ig-coated Sephadex columns or by centrifugation after rosetting with native sheep erythrocytes. Both methods allowed the recovery of B and T-enriched populations the purity of which was checked by fluorescein-labelled anti-Ig serum, E and EAC rosette formation, and heterologous antisera specific for B or T lymphocytes. The proliferative response of T cells to PHA, Con A, PWM, and ALS was not found different from that of unfractionated cells, whereas no response of the B cells could be observed to these mitogens providing that no contaminating T cells were present. Addition of T lymphocytes to these unresponsive B cells allowed them to respond to phytomitogens, but not to ALS. X-irradiated T cells could, to some extent, replace the diving T lymphocytes; no T-replacing factor could be found in cell-free supernatants from T cells, whether or not they had been activated by mitrogens. This model of B-T cooperation appears useful for studying the differentiation and maturation of human B lymphocytes.
Depletion of deep marine food patches forces divers to give up early.
Thums, Michele; Bradshaw, Corey J A; Sumner, Michael D; Horsburgh, Judy M; Hindell, Mark A
2013-01-01
Many optimal foraging models for diving animals examine strategies that maximize time spent in the foraging zone, assuming that prey acquisition increases linearly with search time. Other models have considered the effect of patch quality and predict a net energetic benefit if dives where no prey is encountered early in the dive are abandoned. For deep divers, however, the energetic benefit of giving up is reduced owing to the elevated energy costs associated with descending to physiologically hostile depths, so patch residence time should be invariant. Others consider an asymptotic gain function where the decision to leave a patch is driven by patch-depletion effects - the marginal value theorem. As predator behaviour is increasingly being used as an index of marine resource density and distribution, it is important to understand the nature of this gain function. We investigated the dive behaviour of the world's deepest-diving seal, the southern elephant seal Mirounga leonina, in response to patch quality. Testing these models has largely been limited to controlled experiments on captive animals. By integrating in situ measurements of the seal's relative lipid content obtained from drift rate data (a measure of foraging success) with area-restricted search behaviour identified from first-passage time analysis, we identified regions of high- and low-quality patches. Dive durations and bottom times were not invariant and did not increase in regions of high quality; rather, both were longer when patches were of relatively low quality. This is consistent with the predictions of the marginal value theorem and provides support for a nonlinear relationship between search time and prey acquisition. We also found higher descent and ascent rates in high-quality patches suggesting that seals minimized travel time to the foraging patch when quality was high; however, this was not achieved by increasing speed or dive angle. Relative body lipid content was an important predictor of dive behaviour. Seals did not schedule their diving to maximize time spent in the foraging zone in higher-quality patches, challenging the widely held view that maximizing time in the foraging zone translates to greater foraging success. © 2012 The Authors. Journal of Animal Ecology © 2012 British Ecological Society.
46 CFR 54.01-17 - Pressure vessel for human occupancy (PVHO).
Code of Federal Regulations, 2013 CFR
2013-10-01
... 46 Shipping 2 2013-10-01 2013-10-01 false Pressure vessel for human occupancy (PVHO). 54.01-17... PRESSURE VESSELS General Requirements § 54.01-17 Pressure vessel for human occupancy (PVHO). Pressure vessels for human occupancy (PVHO's) must meet the requirements of subpart B (Commercial Diving Operations...
46 CFR 54.01-17 - Pressure vessel for human occupancy (PVHO).
Code of Federal Regulations, 2014 CFR
2014-10-01
... 46 Shipping 2 2014-10-01 2014-10-01 false Pressure vessel for human occupancy (PVHO). 54.01-17... PRESSURE VESSELS General Requirements § 54.01-17 Pressure vessel for human occupancy (PVHO). Pressure vessels for human occupancy (PVHO's) must meet the requirements of subpart B (Commercial Diving Operations...
46 CFR 54.01-17 - Pressure vessel for human occupancy (PVHO).
Code of Federal Regulations, 2010 CFR
2010-10-01
... 46 Shipping 2 2010-10-01 2010-10-01 false Pressure vessel for human occupancy (PVHO). 54.01-17... PRESSURE VESSELS General Requirements § 54.01-17 Pressure vessel for human occupancy (PVHO). Pressure vessels for human occupancy (PVHO's) must meet the requirements of subpart B (Commercial Diving Operations...
46 CFR 54.01-17 - Pressure vessel for human occupancy (PVHO).
Code of Federal Regulations, 2012 CFR
2012-10-01
... 46 Shipping 2 2012-10-01 2012-10-01 false Pressure vessel for human occupancy (PVHO). 54.01-17... PRESSURE VESSELS General Requirements § 54.01-17 Pressure vessel for human occupancy (PVHO). Pressure vessels for human occupancy (PVHO's) must meet the requirements of subpart B (Commercial Diving Operations...
46 CFR 54.01-17 - Pressure vessel for human occupancy (PVHO).
Code of Federal Regulations, 2011 CFR
2011-10-01
... 46 Shipping 2 2011-10-01 2011-10-01 false Pressure vessel for human occupancy (PVHO). 54.01-17... PRESSURE VESSELS General Requirements § 54.01-17 Pressure vessel for human occupancy (PVHO). Pressure vessels for human occupancy (PVHO's) must meet the requirements of subpart B (Commercial Diving Operations...
Distribution, abundance and habitat use of deep diving cetaceans in the North-East Atlantic
NASA Astrophysics Data System (ADS)
Rogan, Emer; Cañadas, Ana; Macleod, Kelly; Santos, M. Begoña; Mikkelsen, Bjarni; Uriarte, Ainhize; Van Canneyt, Olivier; Vázquez, José Antonio; Hammond, Philip S.
2017-07-01
In spite of their oceanic habitat, deep diving cetacean species have been found to be affected by anthropogenic activities, with potential population impacts of high intensity sounds generated by naval research and oil prospecting receiving the most attention. Improving the knowledge of the distribution and abundance of this poorly known group is an essential prerequisite to inform mitigation strategies seeking to minimize their spatial and temporal overlap with human activities. We provide for the first time abundance estimates for five deep diving cetacean species (sperm whale, long-finned pilot whale, northern bottlenose whale, Cuvier's beaked whale and Sowerby's beaked whale) using data from three dedicated cetacean sighting surveys that covered the oceanic and shelf waters of the North-East Atlantic. Density surface modelling was used to obtain model-based estimates of abundance and to explore the physical and biological characteristics of the habitat used by these species. Distribution of all species was found to be significantly related to depth, distance from the 2000m depth contour, the contour index (a measure of variability in the seabed) and sea surface temperature. Predicted distribution maps also suggest that there is little spatial overlap between these species. Our results represent the best abundance estimates for deep-diving whales in the North-East Atlantic, predict areas of high density during summer and constitute important baseline information to guide future risk assessments of human activities on these species, evaluate potential spatial and temporal trends and inform EU Directives and future conservation efforts.
Jodice, Patrick G.R.; Collopy, Michael W.
1999-01-01
The diving behavior of Marbled Murrelets (Brachyramphus marmoratus) was studied using telemetry along the Oregon coast during the 1995 and 1996 breeding seasons and examined in relation to predictions from optimal-breathing models. Duration of dives, pauses, dive bouts, time spent under water during dive bouts, and nondiving intervals between successive dive bouts were recorded. Most diving metrics differed between years but not with oceanographic conditions or shore type. There was no effect of water depth on mean dive time or percent time spent under water even though dive bouts occurred in depths from 3 to 36 m. There was a significant, positive relationship between mean dive time and mean pause time at the dive-bout scale each year. At the dive-cycle scale, there was a significant positive relationship between dive time and preceding pause time in each year and a significant positive relationship between dive time and ensuing pause time in 1996. Although it appears that aerobic diving was the norm, there appeared to be an increase in anaerobic diving in 1996. The diving performance of Marbled Murrelets in this study appeared to be affected by annual changes in environmental conditions and prey resources but did not consistently fit predictions from optimal-breathing models.
Diving Medicine: Frequently Asked Questions
... after diving In-Water Recompression Inter-Island Flights Massage & Diving Return to Diving After DCI Subcutaneous Emphysema ... Donating Blood Flu-like Symptoms Following a Dive Foot Pain After Diving Fin Foot Frontal Headaches Hand & ...
Sayer, Martin Dj; Akroyd, Jim; Williams, Guy D
2008-06-01
Wreck diving at Bikini Atoll consists of a relatively standard series of decompression dives with maximum depths in the region of 45-55 metres' sea water (msw). In a typical week of diving at Bikini, divers can perform up to 12 decompression dives to these depths over seven days; on five of those days, divers can perform two decompression dives per day. All the dives employ multi-level, staged decompression schedules using air and surface-supplied nitrox containing 80% oxygen. Bikini is serviced by a single diving operator and so a relatively precise record exists both of the actual number of dives undertaken and of the decompression illness incidents both for customer divers and the dive guides. The dive guides follow exactly the dive profiles and decompression schedules of the customers. Each dive guide will perform nearly 400 decompression dives a year, with maximum depths mostly around 50 msw, compared with an average of 10 (maximum of 12) undertaken typically by each customer diver in a week. The incidence of decompression illness for the customer population (presumed in the absence of medical records) is over ten times higher than that for the dive guides. The physiological reasons for such a marked difference are discussed in terms of customer demographics and dive-guide acclimatization to repetitive decompression stress. The rates of decompression illness for a range of diving populations are reviewed.
1981-09-09
oxygen. The last paper of Session 1 was given by Dr. P. Bennett (Duke Univ. Medical Center, NC), and described the Duke "Atlantis" series of deep ...Medicine, Scotland) led off with a presentation on thermal comfort and deep -core temperatures in Antarctic scientific divers. During air diving under ice...than 300 msw (for humans) and deeper than 800 msw (for animals, including several recent dives using baboons to as deep as 1,030 msw) call for a new
ERIC Educational Resources Information Center
Glickstein, Neil
2000-01-01
Introduces an integrated unit on scuba science. Studies oxygen in kinetic theory, Boyle's law, Charles's law, Dalton's law, human circulatory and respiratory systems, and diving dangers such as decompression sickness. (YDS)
Volpov, Beth L; Rosen, David A S; Trites, Andrew W; Arnould, John P Y
2015-08-01
We tested the ability of overall dynamic body acceleration (ODBA) to predict the rate of oxygen consumption ([Formula: see text]) in freely diving Steller sea lions (Eumetopias jubatus) while resting at the surface and diving. The trained sea lions executed three dive types-single dives, bouts of multiple long dives with 4-6 dives per bout, or bouts of multiple short dives with 10-12 dives per bout-to depths of 40 m, resulting in a range of activity and oxygen consumption levels. Average metabolic rate (AMR) over the dive cycle or dive bout calculated was calculated from [Formula: see text]. We found that ODBA could statistically predict AMR when data from all dive types were combined, but that dive type was a significant model factor. However, there were no significant linear relationships between AMR and ODBA when data for each dive type were analyzed separately. The potential relationships between AMR and ODBA were not improved by including dive duration, food consumed, proportion of dive cycle spent submerged, or number of dives per bout. It is not clear whether the lack of predictive power within dive type was due to low statistical power, or whether it reflected a true absence of a relationship between ODBA and AMR. The average percent error for predicting AMR from ODBA was 7-11 %, and standard error of the estimated AMR was 5-32 %. Overall, the extensive range of dive behaviors and physiological conditions we tested indicated that ODBA was not suitable for estimating AMR in the field due to considerable error and the inconclusive effects of dive type.
Kot, J; Sicko, Z; Zyszkowski, M; Brajta, M
2014-01-01
When going to high altitude (higher than 2,400 meters above mean sea level [about 8,200 feet]), human physiology is strongly affected by changes in atmospheric conditions, including decreased ambient pressure and hypobaric hypoxia, which can lead to severe hypoxemia, brain and/or pulmonary edema, negative changes in body and blood composition, as well as disturbances in regional microcirculation. When adding other factors, such as dehydration, physical exercise and exposure to low temperature, it is likely that nitrogen desaturation after diving at such environmental conditions is far from optimal, There are only single reports on diving at high alti-tudes. In 2007 a Polish team of climbers and divers participated in the Tilicho Lake and Peak Expedition to the Himalaya Mountains in Nepal. During this expedition, four divers conducted six dives in the Tilicho Lake at altitude of 4,919 meters above mean sea level equivalent (16,138 feet) to a maximum depth of 15 meters of fresh water (mfw) (equivalent to 28 mfw at sea level by the Cross Correction method) and 30 mfw (equivalent to 57 mfw at sea level "by Cross correction). Decompression debt was calculated using Cross Correction with some additional safety add-ons. Precordial Doppler recordings were taken every 15 minutes until 90 minutes after surfacing. No signs or symptoms of decompression sickness were observed after diving but in one diver, very high bubble grade Doppler signals were recorded. It can be concluded that diving at high altitude should be accompanied by additional safety precautions as well as taking into account personal sensitivity for such conditions.
Johnston, Daniel T; Furness, Robert W; Robbins, Alexandra M C; Tyler, Glen; Taggart, Mark A; Masden, Elizabeth A
2018-03-01
The black guillemot Cepphus grylle has been identified as a species likely to interact with marine renewable energy devices, specifically tidal turbines, with the potential to experience negative impacts. This likelihood is primarily based on the species being a diving seabird, and an inshore, benthic forager often associating with tidal streams. These behavioural properties may bring them into contact with turbine blades, or make them susceptible to alterations to tidal current speed, and/or changes in benthic habitat structure. We examine the knowledge currently available to assess the potential impacts of tidal stream turbines on black guillemot ecology, highlight knowledge gaps and make recommendations for future research. The key ecological aspects investigated include: foraging movements, diving behaviour, seasonal distribution, other sources of disturbance and colony recovery. Relating to foraging behaviour, between studies there is heterogeneity in black guillemot habitat use in relation to season, tide, diurnal cycles, and bathymetry. Currently, there is also little knowledge regarding the benthic habitats associated with foraging. With respect to diving behaviour, there is currently no available research regarding how black guillemots orientate and manoeuvre within the water column. Black guillemots are considered to be a non-migratory species, however little is known about their winter foraging range and habitat. The effect of human disturbance on breeding habitat and the metapopulation responses to potential mortalities are unknown. It is clear further understanding of black guillemot foraging habitat and behaviour is needed to provide renewable energy developers with the knowledge to sustainably locate tidal turbines and mitigate their impacts. Copyright © 2018 Elsevier Ltd. All rights reserved.
[Decompression problems in diving in mountain lakes].
Bühlmann, A A
1989-08-01
The relationship between tolerated high-pressure tissue nitrogen and ambient pressure is practically linear. The tolerated nitrogen high pressure decreases at altitude, as the ambient pressure is lower. Additionally, tissues with short nitrogen half-times have a higher tolerance than tissues which retain nitrogen for longer duration. For the purpose of determining safe decompression routines, the human body can be regarded as consisting of 16 compartments with half-times from 4 to 635 minutes for nitrogen. The coefficients for calculation of the tolerated nitrogen-high pressure in the tissues can be deduced directly from the half-times for nitrogen. We show as application the results of 573 simulated air dives in the pressure-chamber and 544 real dives in mountain lakes in Switzerland (1400-2600 m above sea level) and in Lake Titicaca (3800 m above sea level). They are in accordance with the computed limits of tolerance.
Diving at altitude: from definition to practice.
Egi, S Murat; Pieri, Massimo; Marroni, Alessandro
2014-01-01
Diving above sea level has different motivations for recreational, military, commercial and scientific activities. Despite the apparently wide practice of inland diving, there are three major discrepancies about diving at altitude: threshold elevation that requires changes in sea level procedures; upper altitude limit of the applicability of these modifications; and independent validation of altitude adaptation methods of decompression algorithms. The first problem is solved by converting the normal fluctuation in barometric pressure to an altitude equivalent. Based on the barometric variations recorded from a meteorological center, it is possible to suggest 600 meters as a threshold for classifying a dive as an "altitude" dive. The second problem is solved by proposing the threshold altitude of aviation (2,400 meters) to classify "high" altitude dives. The DAN (Divers Alert Network) Europe diving database (DB) is analyzed to solve the third problem. The database consists of 65,050 dives collected from different dive computers. A total of 1,467 dives were found to be classified as altitude dives. However, by checking the elevation according to the logged geographical coordinates, 1,284 dives were disqualified because the altitude setting had been used as a conservative setting by the dive computer despite the fact that the dive was made at sea level. Furthermore, according to the description put forward in this manuscript, 72 dives were disqualified because the surface level elevation is lower than 600 meters. The number of field data (111 dives) is still very low to use for the validation of any particular method of altitude adaptation concerning decompression algorithms.
Career perspective: Alf O. Brubakk-looking back to see ahead.
Brubakk, Alf O
2015-01-01
The following describes my professional life up till today, but it also describes what I think lies ahead. I have led an interesting professional life and been lucky enough to be at the centre of some of the important development in modern medicine and diving, namely ultrasound in cardiology and the mechanisms of decompression. I therefore should be able to see some of the most challenging and exciting problems ahead. Ultrasound in cardiology has developed from simply listening to the Doppler signal to determine the velocity of blood flow to the complicated description of images presented today. Diving, in addition to being an important commercial and environmental activity, exposes the individual to intermittent hyperoxia and pressure reductions. These challenges evoke the production of radical oxygen species (ROS) and microparticles (MP) that also are central to many pathophysiologic mechanisms that are involved in a number of severe human diseases. Thus, diving can be regarded as an important model of disease and allows us to study their effects on healthy young individuals. The future thus points towards an integration of environmental physiology with detailed physiological and pathophysiological mechanisms and makes diving physiology a potentially very important field of study.
Dive patterns of tagged right whales in the Great South Channel
NASA Astrophysics Data System (ADS)
Winn, Howard E.; Goodyear, Jeffrey D.; Kenney, Robert D.; Petricig, Richard O.
Right whales were tagged in 1988 and 1989 with radio and sonic telemetry tags as part of a multidisciplinary investigation of right whales and their habitat in the Great South Channel region east of Cape Cod. The tags yielded data on the durations of 6456 dives and 6482 surfacings, as well as 23,538 measurements of the depth of a diving whale. Log-survivorship analysis of the 1988 data showed a clear separation between the durations of dives between blows within a single surfacing sequence or bout (intea-bout dives) and longer dives between surfacing sequences (interbout dives) at 27 s, which was also applied to the 1989 data. Inter-bout dives averaged 127.3 s, and were significantly longer in 1988 than in 1989. Inter-bout dives were significantly longer during the day than night in 1988, and longer at night in 1989. The average intea-bout dive duration was 11.8 s, with 1989 dives longer than those in 1988. Surface durations averaged 6.2 s, and were also significantly longer in 1989. Dive depths were recorded only in 1989. Mean dive depth was 7.3 m, and only 12 dives went deeper than 30 m. The typical right whale dive pattern in 1988 included relatively short surfacings, long dives during the day, and shorter dives at night. This correlated with strong diel vertical migration by the dense zooplankton patches on which they were presumed to be feeding based on indirect evidence-from near the surface at night to near the bottom during the day. The 1989 pattern included longer dives during the night, as well as some exceptionally long surfacings. Zooplankton in 1989 did not migrate vertically, and remained near the surface day and night in right whale feeding areas. Right whale dive patterns in the Great South Channel are closely correlated with the horizontal and vertical distributions and movements of dense patches of their zooplankton prey.
Meir, Jessica U; Robinson, Patrick W; Vilchis, L Ignacio; Kooyman, Gerald L; Costa, Daniel P; Ponganis, Paul J
2013-01-01
Although energetics is fundamental to animal ecology, traditional methods of determining metabolic rate are neither direct nor instantaneous. Recently, continuous blood oxygen (O2) measurements were used to assess energy expenditure in diving elephant seals (Mirounga angustirostris), demonstrating that an exceptional hypoxemic tolerance and exquisite management of blood O2 stores underlie the extraordinary diving capability of this consummate diver. As the detailed relationship of energy expenditure and dive behavior remains unknown, we integrated behavior, ecology, and physiology to characterize the costs of different types of dives of elephant seals. Elephant seal dive profiles were analyzed and O2 utilization was classified according to dive type (overall function of dive: transit, foraging, food processing/rest). This is the first account linking behavior at this level with in vivo blood O2 measurements in an animal freely diving at sea, allowing us to assess patterns of O2 utilization and energy expenditure between various behaviors and activities in an animal in the wild. In routine dives of elephant seals, the blood O2 store was significantly depleted to a similar range irrespective of dive function, suggesting that all dive types have equal costs in terms of blood O2 depletion. Here, we present the first physiological evidence that all dive types have similarly high blood O2 demands, supporting an energy balance strategy achieved by devoting one major task to a given dive, thereby separating dive functions into distinct dive types. This strategy may optimize O2 store utilization and recovery, consequently maximizing time underwater and allowing these animals to take full advantage of their underwater resources. This approach may be important to optimizing energy expenditure throughout a dive bout or at-sea foraging trip and is well suited to the lifestyle of an elephant seal, which spends > 90% of its time at sea submerged making diving its most "natural" state.
46 CFR 197.434 - Surface-supplied mixed-gas diving.
Code of Federal Regulations, 2010 CFR
2010-10-01
... 46 Shipping 7 2010-10-01 2010-10-01 false Surface-supplied mixed-gas diving. 197.434 Section 197... HEALTH STANDARDS GENERAL PROVISIONS Commercial Diving Operations Specific Diving Mode Procedures § 197.434 Surface-supplied mixed-gas diving. The diving supervisor shall insure that— (a) When mixed-gas...
NASA Astrophysics Data System (ADS)
Lee, Joo-Young; Park, Joonhee; Koh, Eunsook; Cha, Seongwon
2017-07-01
The aim of the present study was to evaluate the local cold tolerance of older Korean female divers, haenyeo ( N = 22) in terms of cold acclimatization and ageing. As control groups, older non-diving females ( N = 25) and young females from a rural area ( N = 15) and an urban area ( N = 51) participated in this study. To evaluate local cold tolerance, finger cold-induced vasodilation (CIVD) during finger immersion of 4 °C water was examined. As a result, older haenyeos showed greater minimum finger temperature and recovery finger temperature than older non-diving females ( P < 0.05), but similar responses in onset time, peak time, maximum finger temperature, frequency of CIVD, heart rate, blood pressure, and thermal and pain sensations as those of older non-diving females. Another novel finding was that young urban females showed more vulnerable responses to local cold in CIVD variables and subjective sensations when compared to older females, whereas young rural females had the most excellent cold tolerance in terms of maximum temperature and frequency of CIVD among the four groups ( P < 0.05). The present results imply that older haenyeos still retain cold acclimatized features on the periphery even though they changed their cotton diving suits to wet suits in the early 1980s. However, cardiovascular responses and subjective sensations to cold reflect aging effects. In addition, we suggest that young people who have been adapted to highly insulated clothing and indoor heating systems in winter should be distinguished from young people who were exposed to less modern conveniences when compared to the aged in terms of cold tolerance.
Lee, Joo-Young; Park, Joonhee; Koh, Eunsook; Cha, Seongwon
2017-07-01
The aim of the present study was to evaluate the local cold tolerance of older Korean female divers, haenyeo (N = 22) in terms of cold acclimatization and ageing. As control groups, older non-diving females (N = 25) and young females from a rural area (N = 15) and an urban area (N = 51) participated in this study. To evaluate local cold tolerance, finger cold-induced vasodilation (CIVD) during finger immersion of 4 °C water was examined. As a result, older haenyeos showed greater minimum finger temperature and recovery finger temperature than older non-diving females (P < 0.05), but similar responses in onset time, peak time, maximum finger temperature, frequency of CIVD, heart rate, blood pressure, and thermal and pain sensations as those of older non-diving females. Another novel finding was that young urban females showed more vulnerable responses to local cold in CIVD variables and subjective sensations when compared to older females, whereas young rural females had the most excellent cold tolerance in terms of maximum temperature and frequency of CIVD among the four groups (P < 0.05). The present results imply that older haenyeos still retain cold acclimatized features on the periphery even though they changed their cotton diving suits to wet suits in the early 1980s. However, cardiovascular responses and subjective sensations to cold reflect aging effects. In addition, we suggest that young people who have been adapted to highly insulated clothing and indoor heating systems in winter should be distinguished from young people who were exposed to less modern conveniences when compared to the aged in terms of cold tolerance.
Boyd, Charlotte; Castillo, Ramiro; Hunt, George L; Punt, André E; VanBlaricom, Glenn R; Weimerskirch, Henri; Bertrand, Sophie
2015-11-01
Understanding the ecological processes that underpin species distribution patterns is a fundamental goal in spatial ecology. However, developing predictive models of habitat use is challenging for species that forage in marine environments, as both predators and prey are often highly mobile and difficult to monitor. Consequently, few studies have developed resource selection functions for marine predators based directly on the abundance and distribution of their prey. We analysed contemporaneous data on the diving locations of two seabird species, the shallow-diving Peruvian Booby (Sula variegata) and deeper diving Guanay Cormorant (Phalacrocorax bougainvilliorum), and the abundance and depth distribution of their main prey, Peruvian anchoveta (Engraulis ringens). Based on this unique data set, we developed resource selection functions to test the hypothesis that the probability of seabird diving behaviour at a given location is a function of the relative abundance of prey in the upper water column. For both species, we show that the probability of diving behaviour is mostly explained by the distribution of prey at shallow depths. While the probability of diving behaviour increases sharply with prey abundance at relatively low levels of abundance, support for including abundance in addition to the depth distribution of prey is weak, suggesting that prey abundance was not a major factor determining the location of diving behaviour during the study period. The study thus highlights the importance of the depth distribution of prey for two species of seabird with different diving capabilities. The results complement previous research that points towards the importance of oceanographic processes that enhance the accessibility of prey to seabirds. The implications are that locations where prey is predictably found at accessible depths may be more important for surface foragers, such as seabirds, than locations where prey is predictably abundant. Analysis of the relative importance of abundance and accessibility is essential for the design and evaluation of effective management responses to reduced prey availability for seabirds and other top predators in marine systems. © 2015 The Authors. Journal of Animal Ecology © 2015 British Ecological Society.
NASA Astrophysics Data System (ADS)
Kokubun, N.; Yamamoto, T.,; Sato, N.; Watanuki, Y.; Will, A.; Kitaysky, A. S.; Takahashi, A.
2015-11-01
Sub-arctic environmental changes are expected to affect the ecology of marine top predators. We examined the characteristics of foraging behavior of two sympatric congeneric diving seabirds, common (Uria aalge: hereafter COMU) and thick-billed (U. lomvia: hereafter TBMU) murres breeding on St. George Island located in the seasonal sea-ice region of the Bering Sea. We investigated their flight duration, diel patterns of dive depth, and underwater wing strokes, along with morphology and blood stable isotopes. Acceleration-temperature-depth data loggers were attached to chick-guarding birds, and behavioral data were obtained from 7 COMU and 12 TBMU. Both species showed similar trip duration (13.21 ± 4.79 h for COMU and 10.45 ± 7.09 h for TBMU) and similar diurnal patterns of diving (frequent dives to various depths in the daytime and less frequent dives to shallow depths in the nighttime). During the daytime, dive depths of COMU had two peaks in shallow (18.1 ± 6.0 m) and deep (74.2 ± 8.7 m) depths, while those of TBMU were 20.2 ± 7.4 m and 59.7 ± 7.9 m. COMU showed more frequent wing strokes during the bottom phase of dives (1.90 ± 0.11 s-1) than TBMU (1.66 ± 0.15 s-1). Fishes occurred with higher proportion in the bill-loads brought back to chicks in COMU (85 %) than in TBMU (56 %). δ15N value of blood was significantly higher in COMU (14.47 ± 0.27 ‰) than in TBMU (13.14 ± 0.36 ‰). Relatively small wing area (0.053 ± 0.007 m2) of COMU compared to TBMU (0.067 ± 0.007 m2) may make them more agile underwater and thus enable them to target more mobile prey including larger fishes that inhabit deeper depths. These differences in foraging behavior between COMU and TBMU might explain the differences in their responses to long-term marine environmental changes.
The effect of pre-dive ingestion of dark chocolate on endothelial function after a scuba dive.
Theunissen, Sigrid; Balestra, Costantino; Boutros, Antoine; De Bels, David; Guerrero, François; Germonpré, Peter
2015-03-01
The aim of the study was to observe the effects of dark chocolate on endothelial function after scuba diving. Forty-two male scuba divers were divided into two groups: a control (n=21) and a chocolate group (n=21). They performed a 33-metres deep scuba-air dive for 20 minutes in a diving pool (Nemo 33, Brussels). Water temperature was 33⁰C. The chocolate group ingested 30 g of dark chocolate (86% cocoa) 90 minutes before the dive. Flow-mediated dilatation (FMD), digital photoplethysmography and nitric oxide (NO) and peroxynitrites (ONOO-) levels were measured before and after the scuba dive in both groups. A significant decrease in FMD was observed in the control group after the dive (91±7% (mean±95% confidence interval) of pre-dive values; P<0.001) while it was increased in the chocolate group (105±5% of pre-dive values; P<0.001). No difference in digital photoplethysmography was observed between before and after the dives. No variation of circulating NO level was observed in the control group whereas an increase was shown in the chocolate group (154±73% of pre-dive values; P=0.04). A significant reduction in ONOO- was observed in the control group (84±12% of pre-dive values; P=0.003) whereas no variation was shown after the dive with chocolate intake (100±28% of pre-dive values; ns). Ingestion of 30 g of dark chocolate 90 minutes before scuba diving prevented post-dive endothelial dysfunction, as the antioxidants contained in dark chocolate probably scavenge free radicals.
Stimpert, A. K.; DeRuiter, S. L.; Southall, B. L.; Moretti, D. J.; Falcone, E. A.; Goldbogen, J. A.; Friedlaender, A.; Schorr, G. S.; Calambokidis, J.
2014-01-01
Beaked whales are hypothesized to be particularly sensitive to anthropogenic noise, based on previous strandings and limited experimental and observational data. However, few species have been studied in detail. We describe the underwater behavior of a Baird's beaked whale (Berardius bairdii) from the first deployment of a multi-sensor acoustic tag on this species. The animal exhibited shallow (23 ± 15 m max depth), intermediate (324 ± 49 m), and deep (1138 ± 243 m) dives. Echolocation clicks were produced with a mean inter-click interval of approximately 300 ms and peak frequency of 25 kHz. Two deep dives included presumed foraging behavior, with echolocation pulsed sounds (presumed prey capture attempts) associated with increased maneuvering, and sustained inverted swimming during the bottom phase of the dive. A controlled exposure to simulated mid-frequency active sonar (3.5–4 kHz) was conducted 4 hours after tag deployment, and within 3 minutes of exposure onset, the tagged whale increased swim speed and body movement, and continued to show unusual dive behavior for each of its next three dives, one of each type. These are the first data on the acoustic foraging behavior in this largest beaked whale species, and the first experimental demonstration of a response to simulated sonar. PMID:25391309
Diving down the reefs? Intensive diving tourism threatens the reefs of the northern Red Sea.
Hasler, Harald; Ott, Jörg A
2008-10-01
Intensive recreational SCUBA diving threatens coral reef ecosystems. The reefs at Dahab, South Sinai, Egypt, are among the world's most dived (>30,000 dives y(-1)). We compared frequently dived sites to sites with no or little diving. Benthic communities and condition of corals were examined by the point intercept sampling method in the reef crest zone (3m) and reef slope zone (12 m). Additionally, the abundance of corallivorous and herbivorous fish was estimated based on the visual census method. Sediments traps recorded the sedimentation rates caused by SCUBA divers. Zones subject to intensive SCUBA diving showed a significantly higher number of broken and damaged corals and significantly lower coral cover. Reef crest coral communities were significantly more affected than those of the reef slope: 95% of the broken colonies were branching ones. No effect of diving on the abundance of corallivorous and herbivorous fish was evident. At heavily used dive sites, diver-related sedimentation rates significantly decreased with increasing distance from the entrance, indicating poor buoyancy regulation at the initial phase of the dive. The results show a high negative impact of current SCUBA diving intensities on coral communities and coral condition. Corallivorous and herbivorous fishes are apparently not yet affected, but are endangered if coral cover decline continues. Reducing the number of dives per year, ecologically sustainable dive plans for individual sites, and reinforcing the environmental education of both dive guides and recreational divers are essential to conserve the ecological and the aesthetic qualities of these dive sites.
Diving injuries of the cervical spine in amateur divers.
Korres, Demetrios S; Benetos, Ioannis S; Themistocleous, George S; Mavrogenis, Andreas F; Nikolakakos, Leonidas; Liantis, Panagiotis T
2006-01-01
Diving injuries are the cause of potentially devastating trauma, primarily affecting the cervical spine. Our purpose was to describe our experience with diving injuries treatment. Retrospective review. Twenty patients with diving injuries. Using the American Spinal Injury Association (ASIA) impairment scales as the primary outcome measure, the patients' neurological status before and after treatment was assessed. In this way we were able to draw conclusions about neurological improvement or deterioration in response to conservative or operative treatment. We retrospectively reviewed 20 patients with diving injuries of the cervical spine who were admitted to our institute over a 34-year period from 1970 until 2004. The typical patient profile was of a young, healthy, athletic male who suffered an injury to the cervical spine after diving into shallow water. The number of cases corresponds to 2.6% of all admitted cervical spine injuries. All injures occurred between May and September. The most commonly fractured vertebrae were C5 and C6. Four patients were treated operatively and 16 conservatively. The indications for surgical treatment were posttraumatic instability and persistent neurological deficit. The mean follow-up of the patients was 17 years. Five patients died within the first month of their hospitalization and 1 patient died 1 year after his injury. Of the 14 patients who were available for follow-up 5 years past injury time, 6 improved neurologically and 8 remained unchanged in relation to their neurology upon admission. Of the 11 patients who were available for follow-up 10 years past injury time, 9 remained neurologically unchanged, 1 deteriorated, and 1 improved in relation to their neurology in the 5-year follow-up. Diving injuries of the cervical spine demonstrate high mortality and morbidity rates. Recovery depends on the severity of the initial neurological damage. Conservative treatment is justified in specific patients and can lead to improvement of the initial neurological deficit.
A Software Tool for the Annotation of Embolic Events in Echo Doppler Audio Signals
Pierleoni, Paola; Maurizi, Lorenzo; Palma, Lorenzo; Belli, Alberto; Valenti, Simone; Marroni, Alessandro
2017-01-01
The use of precordial Doppler monitoring to prevent decompression sickness (DS) is well known by the scientific community as an important instrument for early diagnosis of DS. However, the timely and correct diagnosis of DS without assistance from diving medical specialists is unreliable. Thus, a common protocol for the manual annotation of echo Doppler signals and a tool for their automated recording and annotation are necessary. We have implemented original software for efficient bubble appearance annotation and proposed a unified annotation protocol. The tool auto-sets the response time of human “bubble examiners,” performs playback of the Doppler file by rendering it independent of the specific audio player, and enables the annotation of individual bubbles or multiple bubbles known as “showers.” The tool provides a report with an optimized data structure and estimates the embolic risk level according to the Extended Spencer Scale. The tool is built in accordance with ISO/IEC 9126 on software quality and has been projected and tested with assistance from the Divers Alert Network (DAN) Europe Foundation, which employs this tool for its diving data acquisition campaigns. PMID:29242701
Hidden Markov models reveal complexity in the diving behaviour of short-finned pilot whales
Quick, Nicola J.; Isojunno, Saana; Sadykova, Dina; Bowers, Matthew; Nowacek, Douglas P.; Read, Andrew J.
2017-01-01
Diving behaviour of short-finned pilot whales is often described by two states; deep foraging and shallow, non-foraging dives. However, this simple classification system ignores much of the variation that occurs during subsurface periods. We used multi-state hidden Markov models (HMM) to characterize states of diving behaviour and the transitions between states in short-finned pilot whales. We used three parameters (number of buzzes, maximum dive depth and duration) measured in 259 dives by digital acoustic recording tags (DTAGs) deployed on 20 individual whales off Cape Hatteras, North Carolina, USA. The HMM identified a four-state model as the best descriptor of diving behaviour. The state-dependent distributions for the diving parameters showed variation between states, indicative of different diving behaviours. Transition probabilities were considerably higher for state persistence than state switching, indicating that dive types occurred in bouts. Our results indicate that subsurface behaviour in short-finned pilot whales is more complex than a simple dichotomy of deep and shallow diving states, and labelling all subsurface behaviour as deep dives or shallow dives discounts a significant amount of important variation. We discuss potential drivers of these patterns, including variation in foraging success, prey availability and selection, bathymetry, physiological constraints and socially mediated behaviour. PMID:28361954
77 FR 25963 - Marine Mammals; File No. 14325
Federal Register 2010, 2011, 2012, 2013, 2014
2012-05-02
... attachment of scientific instruments to investigate foraging ecology, diving behavior and habitat use. The... the permit would not have a significant adverse impact on the human environment. Concurrent with the...
78 FR 53732 - Marine Mammals; File No. 14325
Federal Register 2010, 2011, 2012, 2013, 2014
2013-08-30
... attachment of scientific instruments to investigate foraging ecology, diving behavior and habitat use. The... issuance of the permit would not have a significant adverse impact on the human environment. Concurrent...
Kwiatkowski, M; Wurlitzer, M; Krutilin, A; Kiani, P; Nimer, R; Omidi, M; Mannaa, A; Bussmann, T; Bartkowiak, K; Kruber, S; Uschold, S; Steffen, P; Lübberstedt, J; Küpker, N; Petersen, H; Knecht, R; Hansen, N O; Zarrine-Afsar, A; Robertson, W D; Miller, R J D; Schlüter, H
2016-02-16
Posttranslational modifications and proteolytic processing regulate almost all physiological processes. Dysregulation can potentially result in pathologic protein species causing diseases. Thus, tissue species proteomes of diseased individuals provide diagnostic information. Since the composition of tissue proteomes can rapidly change during tissue homogenization by the action of enzymes released from their compartments, disease specific protein species patterns can vanish. Recently, we described a novel, ultrafast and soft method for cold vaporization of tissue via desorption by impulsive vibrational excitation (DIVE) using a picosecond-infrared-laser (PIRL). Given that DIVE extraction may provide improved access to the original composition of protein species in tissues, we compared the proteome composition of tissue protein homogenates after DIVE homogenization with conventional homogenizations. A higher number of intact protein species was observed in DIVE homogenates. Due to the ultrafast transfer of proteins from tissues via gas phase into frozen condensates of the aerosols, intact protein species were exposed to a lesser extent to enzymatic degradation reactions compared with conventional protein extraction. In addition, total yield of the number of proteins is higher in DIVE homogenates, because they are very homogenous and contain almost no insoluble particles, allowing direct analysis with subsequent analytical methods without the necessity of centrifugation. Enzymatic protein modifications during tissue homogenization are responsible for changes of the in-vivo protein species composition. Cold vaporization of tissues by PIRL-DIVE is comparable with taking a snapshot at the time of the laser irradiation of the dynamic changes that occur continuously under in-vivo conditions. At that time point all biomolecules are transferred into an aerosol, which is immediately frozen. Copyright © 2016 The Authors. Published by Elsevier B.V. All rights reserved.
Kwiatkowski, M.; Wurlitzer, M.; Krutilin, A.; Kiani, P.; Nimer, R.; Omidi, M.; Mannaa, A.; Bussmann, T.; Bartkowiak, K.; Kruber, S.; Uschold, S.; Steffen, P.; Lübberstedt, J.; Küpker, N.; Petersen, H.; Knecht, R.; Hansen, N.O.; Zarrine-Afsar, A.; Robertson, W.D.; Miller, R.J.D.; Schlüter, H.
2016-01-01
Posttranslational modifications and proteolytic processing regulate almost all physiological processes. Dysregulation can potentially result in pathologic protein species causing diseases. Thus, tissue species proteomes of diseased individuals provide diagnostic information. Since the composition of tissue proteomes can rapidly change during tissue homogenization by the action of enzymes released from their compartments, disease specific protein species patterns can vanish. Recently, we described a novel, ultrafast and soft method for cold vaporization of tissue via desorption by impulsive vibrational excitation (DIVE) using a picosecond-infrared-laser (PIRL). Given that DIVE extraction may provide improved access to the original composition of protein species in tissues, we compared the proteome composition of tissue protein homogenates after DIVE homogenization with conventional homogenizations. A higher number of intact protein species was observed in DIVE homogenates. Due to the ultrafast transfer of proteins from tissues via gas phase into frozen condensates of the aerosols, intact protein species were exposed to a lesser extent to enzymatic degradation reactions compared with conventional protein extraction. In addition, total yield of the number of proteins is higher in DIVE homogenates, because they are very homogenous and contain almost no insoluble particles, allowing direct analysis with subsequent analytical methods without the necessity of centrifugation. Biological significance Enzymatic protein modifications during tissue homogenization are responsible for changes of the in-vivo protein species composition. Cold vaporization of tissues by PIRL-DIVE is comparable with taking a snapshot at the time of the laser irradiation of the dynamic changes that occur continuously under in-vivo conditions. At that time point all biomolecules are transferred into an aerosol, which is immediately frozen. PMID:26778141
Peery, M.Z.; Newman, S.H.; Storlazzi, C.D.; Beissinger, S.R.
2009-01-01
Seabirds maintain plasticity in their foraging behavior to cope with energy demands and foraging constraints that vary over the reproductive cycle, but behavioral studies comparing breeding and nonbreeding individuals are rare. Here we characterize how Marbled Murrelets (Brachyramphus marmoratus) adjust their foraging effort in response to changes in reproductive demands in an upwelling system in central California. We radio-marked 32 murrelets of known reproductive status (9 breeders, 12 potential breeders, and 11 nonbreeders) and estimated both foraging ranges and diving rates during the breeding season. Murrelets spent more time diving during upwelling than oceanographic relaxation, increased their foraging ranges as the duration of relaxation grew longer, and reduced their foraging ranges after transitions to upwelling. When not incubating, murrelets moved in a circadian pattern, spending nighttime hours resting near flyways used to reach nesting habitat and foraging during the daytime an average of 5.7 km (SD 6.7 km) from nighttime locations. Breeders foraged close to nesting habitat once they initiated nesting and nest attendance was at a maximum, and then resumed traveling longer distances following the completion of nesting. Nonbreeders had similar nighttime and daytime distributions and tended to be located farther from inland flyways. Breeders increased the amount of time they spent diving by 71-73% when they had an active nest by increasing the number of dives rather than by increasing the frequency of anaerobiosis. Thus, to meet reproductive demands during nesting, murrelets adopted a combined strategy of reducing energy expended commuting to foraging sites and increasing aerobic dive rates. ?? 2009 by The Cooper Ornithological Society. All rights reserved.
Accident rates at a busy diving centre.
Davis, Michael; Malcolm, Kate
2008-06-01
Dear Editor, The Poor Knights Islands in Northland, New Zealand, is a world-famous, temperate-water, diving tourism destination, popularised many years ago by Jacques Cousteau. By far the largest dive operator there is Dive! Tutukaka, with five vessels carrying up to 30 divers, operating on a regular basis throughout the year. Dive! Tutukaka is required to keep a detailed, daily vessel manifest. Thus, the number of divers is known accurately and all incidents are recorded by the Skipper or the Chief Divemaster on board. Although all dives are logged (time in, time out and maximum depth for every diver) and kept permanently, these data were not utilised for this brief report. Each customer does two dives on a trip and there are between one and four divemasters on board who may do one, two or more dives a day (van der Hulst G, unpublished observations). Thus the accident rate per diver is known, and it is assumed that the rate per dive is very close to half this figure. In addition, under health and safety regulations all non-diving injuries both on shore and on board are documented, but these will include some non-divers. For the three financial years between July 2005 and 14 June 2008, 32,302 customers dived with Dive! Tutukaka, approximately 63,000 dives (a small minority did only one dive). Over the same period, there were an estimated 7,600 dives conducted by the divemasters. The injuries documented during this time are shown in Table 1. There were seven cases of decompression illness (DCI), a rate of about 1 per 10,000 divers (0.5 per 10,000 dives). Two of the seven DCI cases involved serious neurological injury. There was one further possible case of DCI who did not seek medical advice. If this diver is included then the rate is 1.14 per 10,000 divers. More minor diving injuries and incidents occurred at a rate of approximately 2 per 10,000 divers. Non-diving injuries occurred rarely, the most common being various musculo-skeletal injuries to staff, requiring time off work. Many of these were secondary to lifting and carrying heavy diving equipment, particularly dive tanks. This indicates an area where improved practices by staff could be achieved. We believe these injury data are robust and provide an accurate picture of a single, mainstream, international tourism diving centre in temperate waters, and indicate a low rate of injury, comparable to the international literature.
NASA Astrophysics Data System (ADS)
Sasaki, T.; Azuma, S.; Matsuda, S.; Nagayama, A.; Ogido, M.; Saito, H.; Hanafusa, Y.
2016-12-01
The Japan Agency for Marine-Earth Science and Technology (JAMSTEC) archives a large amount of deep-sea research videos and photos obtained by JAMSTEC's research submersibles and vehicles with cameras. The web site "JAMSTEC E-library of Deep-sea Images : J-EDI" (http://www.godac.jamstec.go.jp/jedi/e/) has made videos and photos available to the public via the Internet since 2011. Users can search for target videos and photos by keywords, easy-to-understand icons, and dive information at J-EDI because operating staffs classify videos and photos as to contents, e.g. living organism and geological environment, and add comments to them.Dive survey data including videos and photos are not only valiant academically but also helpful for education and outreach activities. With the aim of the improvement of visibility for broader communities, we added new functions of 3-dimensional display synchronized various dive survey data with videos in this year.New Functions Users can search for dive survey data by 3D maps with plotted dive points using the WebGL virtual map engine "Cesium". By selecting a dive point, users can watch deep-sea videos and photos and associated environmental data, e.g. water temperature, salinity, rock and biological sample photos, obtained by the dive survey. Users can browse a dive track visualized in 3D virtual spaces using the WebGL JavaScript library. By synchronizing this virtual dive track with videos, users can watch deep-sea videos recorded at a point on a dive track. Users can play an animation which a submersible-shaped polygon automatically traces a 3D virtual dive track and displays of dive survey data are synchronized with tracing a dive track. Users can directly refer to additional information of other JAMSTEC data sites such as marine biodiversity database, marine biological sample database, rock sample database, and cruise and dive information database, on each page which a 3D virtual dive track is displayed. A 3D visualization of a dive track makes users experience a virtual dive survey. In addition, by synchronizing a virtual dive track with videos, it is easy to understand living organisms and geological environments of a dive point. Therefore, these functions will visually support understanding of deep-sea environments in lectures and educational activities.
36 CFR 3.18 - May I snorkel or underwater dive in park waters?
Code of Federal Regulations, 2011 CFR
2011-07-01
..., DEPARTMENT OF THE INTERIOR BOATING AND WATER USE ACTIVITIES § 3.18 May I snorkel or underwater dive in park... open to the use of vessels, a diver must prominently display a dive flag during dive operations. A dive...
36 CFR 3.18 - May I snorkel or underwater dive in park waters?
Code of Federal Regulations, 2010 CFR
2010-07-01
..., DEPARTMENT OF THE INTERIOR BOATING AND WATER USE ACTIVITIES § 3.18 May I snorkel or underwater dive in park... open to the use of vessels, a diver must prominently display a dive flag during dive operations. A dive...
Diving bradycardia of elderly Korean women divers, haenyeo, in cold seawater: a field report
LEE, Joo-Young; LEE, Hyo-Hyun; KIM, Siyeon; JANG, Young-Joon; BAEK, Yoon-Jeong; KANG, Kwon-Yong
2015-01-01
The purpose of the present field study was to explore diving patterns and heart rate of elderly Korean women divers (haenyeo) while breath-hold diving in cold seawater. We hypothesized that the decreasing rate in heart rate of elderly haenyeos during breath-hold diving was greater and total diving time was shorter than those of young haenyeos from previous studies. Nine haenyeos participated in a field study [68 ± 10 yr in age, ranged from 56 to 83 yr] at a seawater temperature of 10 to 13 °C. Average total diving time including surface swimming time between dives was 253 ± 73 min (155–341 min). Total frequency of dives was 97 ± 28 times and they dived 23 ± 8 times per hour. All haenyeos showed diving bradycardia with a decreased rate of 20 ± 8% at the bottom time (101 ± 20 bpm) when compared to surface swimming time (125 ± 16 bpm) in the sea. Older haenyeos among the nine elderly haenyeos had shorter diving time, less diving frequencies, and lower heart rate at work (p<0.05). These reductions imply that haenyeos voluntarily adjust their workload along with advancing age and diminished cardiovascular functions. PMID:26632118
Diving bradycardia of elderly Korean women divers, haenyeo, in cold seawater: a field report.
Lee, Joo-Young; Lee, Hyo-Hyun; Kim, Siyeon; Jang, Young-Joon; Baek, Yoon-Jeong; Kang, Kwon-Yong
2016-01-01
The purpose of the present field study was to explore diving patterns and heart rate of elderly Korean women divers (haenyeo) while breath-hold diving in cold seawater. We hypothesized that the decreasing rate in heart rate of elderly haenyeos during breath-hold diving was greater and total diving time was shorter than those of young haenyeos from previous studies. Nine haenyeos participated in a field study [68 ± 10 yr in age, ranged from 56 to 83 yr] at a seawater temperature of 10 to 13 °C. Average total diving time including surface swimming time between dives was 253 ± 73 min (155-341 min). Total frequency of dives was 97 ± 28 times and they dived 23 ± 8 times per hour. All haenyeos showed diving bradycardia with a decreased rate of 20 ± 8% at the bottom time (101 ± 20 bpm) when compared to surface swimming time (125 ± 16 bpm) in the sea. Older haenyeos among the nine elderly haenyeos had shorter diving time, less diving frequencies, and lower heart rate at work (p<0.05). These reductions imply that haenyeos voluntarily adjust their workload along with advancing age and diminished cardiovascular functions.
Gordine, Samantha Alex; Fedak, Michael; Boehme, Lars
2015-01-01
ABSTRACT In southern elephant seals (Mirounga leonina), fasting- and foraging-related fluctuations in body composition are reflected by buoyancy changes. Such buoyancy changes can be monitored by measuring changes in the rate at which a seal drifts passively through the water column, i.e. when all active swimming motion ceases. Here, we present an improved knowledge-based method for detecting buoyancy changes from compressed and abstracted dive profiles received through telemetry. By step-wise filtering of the dive data, the developed algorithm identifies fragments of dives that correspond to times when animals drift. In the dive records of 11 southern elephant seals from South Georgia, this filtering method identified 0.8–2.2% of all dives as drift dives, indicating large individual variation in drift diving behaviour. The obtained drift rate time series exhibit that, at the beginning of each migration, all individuals were strongly negatively buoyant. Over the following 75–150 days, the buoyancy of all individuals peaked close to or at neutral buoyancy, indicative of a seal's foraging success. Independent verification with visually inspected detailed high-resolution dive data confirmed that this method is capable of reliably detecting buoyancy changes in the dive records of drift diving species using abstracted data. This also affirms that abstracted dive profiles convey the geometric shape of drift dives in sufficient detail for them to be identified. Further, it suggests that, using this step-wise filtering method, buoyancy changes could be detected even in old datasets with compressed dive information, for which conventional drift dive classification previously failed. PMID:26486362
The tourism carrying capacity of underwater trails in Isabel Island National Park, Mexico.
Ríos-Jara, Eduardo; Galván-Villa, Cristian Moisés; Rodríguez-Zaragoza, Fabián Alejandro; López-Uriarte, Ernesto; Muñoz-Fernández, Vicente Teófilo
2013-08-01
The popularity of ecotourism in the marine protected areas of Mexico has increased over the last 10 years; in particular there is a large development of a SCUBA diving industry in the Mexican Pacific including Isabel Island. Given the risks associated with human activity in the marine environments around this island, we propose two ecotourism management strategies: (1) the creation and use of underwater trails, and (2) the estimation of the specific tourism carrying capacity (TCC) for each trail. Six underwater trails were selected in sites that presented elements of biological, geological, and scenic interest, using information obtained during field observations. The methodology used to estimate the TCC was based upon the physical and biological conditions of each site, the infrastructure and equipment available, and the characteristics of the service providers and the administrators of the park. Correction factors of the TCC included elements of the quality of the visit and the threat and vulnerability of the marine environment of each trail (e.g., divers' expertise, size and distance between groups of divers, accessibility, wind, coral coverage). The TCC values ranged between 1,252 and 1,642 dives/year/trail, with a total of 8,597 dives/year for all six trails. Although these numbers are higher than the actual number of recreational visitors to the island (~1,000 dives per year), there is a need for adequate preventive management if the diving sites are to maintain their esthetic appeal and biological characteristics. Such management might be initially directed toward using only the sites and the TCC proposed here.
Fago, Angela; Parraga, Daniel Garcia; Petersen, Elin E; Kristensen, Niels; Giouri, Lea; Jensen, Frank B
2017-03-01
The ability of marine mammals to hunt prey at depth is known to rely on enhanced oxygen stores and on selective distribution of blood flow, but the molecular mechanisms regulating blood flow and oxygen transport remain unresolved. To investigate the molecular mechanisms that may be important in regulating blood flow, we measured concentration of nitrite and S-nitrosothiols (SNO), two metabolites of the vasodilator nitric oxide (NO), in the blood of 5 species of marine mammals differing in their dive duration: bottlenose dolphin, South American sea lion, harbor seal, walrus and beluga whale. We also examined oxygen affinity, sensitivity to 2,3-diphosphoglycerate (DPG) and nitrite reductase activity of the hemoglobin (Hb) to search for possible adaptive variations in these functional properties. We found levels of plasma and red blood cells nitrite similar to those reported for terrestrial mammals, but unusually high concentrations of red blood cell SNO in bottlenose dolphin, walrus and beluga whale, suggesting enhanced SNO-dependent signaling in these species. Purified Hbs showed similar functional properties in terms of oxygen affinity and sensitivity to DPG, indicating that reported large variations in blood oxygen affinity among diving mammals likely derive from phenotypic variations in red blood cell DPG levels. The nitrite reductase activities of the Hbs were overall slightly higher than that of human Hb, with the Hb of beluga whale, capable of longest dives, having the highest activity. Taken together, these results underscore adaptive variations in circulatory NO metabolism in diving mammals but not in the oxygenation properties of the Hb. Copyright © 2016 Elsevier Inc. All rights reserved.
The Tourism Carrying Capacity of Underwater Trails in Isabel Island National Park, Mexico
NASA Astrophysics Data System (ADS)
Ríos-Jara, Eduardo; Galván-Villa, Cristian Moisés; Rodríguez-Zaragoza, Fabián Alejandro; López-Uriarte, Ernesto; Muñoz-Fernández, Vicente Teófilo
2013-08-01
The popularity of ecotourism in the marine protected areas of Mexico has increased over the last 10 years; in particular there is a large development of a SCUBA diving industry in the Mexican Pacific including Isabel Island. Given the risks associated with human activity in the marine environments around this island, we propose two ecotourism management strategies: (1) the creation and use of underwater trails, and (2) the estimation of the specific tourism carrying capacity (TCC) for each trail. Six underwater trails were selected in sites that presented elements of biological, geological, and scenic interest, using information obtained during field observations. The methodology used to estimate the TCC was based upon the physical and biological conditions of each site, the infrastructure and equipment available, and the characteristics of the service providers and the administrators of the park. Correction factors of the TCC included elements of the quality of the visit and the threat and vulnerability of the marine environment of each trail (e.g., divers' expertise, size and distance between groups of divers, accessibility, wind, coral coverage). The TCC values ranged between 1,252 and 1,642 dives/year/trail, with a total of 8,597 dives/year for all six trails. Although these numbers are higher than the actual number of recreational visitors to the island (~1,000 dives per year), there is a need for adequate preventive management if the diving sites are to maintain their esthetic appeal and biological characteristics. Such management might be initially directed toward using only the sites and the TCC proposed here.
NASA Astrophysics Data System (ADS)
Ferrini, V. L.; Grange, B.; Morton, J. J.; Soule, S. A.; Carbotte, S. M.; Lehnert, K.
2016-12-01
The National Deep Submergence Facility (NDSF) operates the Human Occupied Vehicle (HOV) Alvin, the Remotely Operated Vehicle (ROV) Jason, and the Autonomous Underwater Vehicle (AUV) Sentry. These vehicles are deployed throughout the global oceans to acquire sensor data and physical samples for a variety of interdisciplinary science programs. As part of the EarthCube Integrative Activity Alliance Testbed Project (ATP), new web services were developed to improve access to existing online NDSF data and metadata resources. These services make use of tools and infrastructure developed by the Interdisciplinary Earth Data Alliance (IEDA) and enable programmatic access to metadata and data resources as well as the development of new service-driven user interfaces. The Alvin Frame Grabber and Jason Virtual Van enable the exploration of frame-grabbed images derived from video cameras on NDSF dives. Metadata available for each image includes time and vehicle position, data from environmental sensors, and scientist-generated annotations, and data are organized and accessible by cruise and/or dive. A new FrameGrabber web service and service-driven user interface were deployed to offer integrated access to these data resources through a single API and allows users to search across content curated in both systems. In addition, a new NDSF Dive Metadata web service and service-driven user interface was deployed to provide consolidated access to basic information about each NDSF dive (e.g. vehicle name, dive ID, location, etc), which is important for linking distributed data resources curated in different data systems.
Reconstructing the evolution of first-row transition metal minerals by GeoDeepDive
NASA Astrophysics Data System (ADS)
Liu, C.; Peters, S. E.; Ross, I.; Golden, J. J.; Downs, R. T.; Hazen, R. M.
2016-12-01
Terrestrial mineralogy evolves as a consequence of a range of physical, chemical, and biological processes [1]. Evolution of the first-row transition metal minerals could mirror the evolution of Earth's oxidation state and life, since these elements mostly are redox-sensitive and/or play critical roles in biology. The fundamental building blocks to reconstruct mineral evolution are the mineral species, locality, and age data, which are typically dispersed in sentences in scientific and technical publications. These data can be tracked down in a brute-force way, i.e., human retrieval, reading, and recording all relevant literature. Alternatively, they can be extracted automatically by GeoDeepDive. In GeoDeepDive, scientific and technical articles from publishers, including Elsevier, Wiley, USGS, SEPM, GSA and Canada Science Publishing, have been parsed into a Javascript database with NLP tags. Sentences containing data of mineral names, locations, and ages can be recognized and extracted by user-developed applications. In a preliminary search for cobalt mineral ages, we successfully extracted 678 citations with >1000 mentions of cobalt minerals, their locations, and ages. The extracted results are in agreement with brute-force search results. What is more, GeoDeepDive provides 40 additional data points that were not recovered by the brute-force approach. The extracted mineral locality-age data suggest that the evolution of Co minerals is controlled by global supercontinent cycles, i.e., more Co minerals form during episodes of supercontinent assembly. Mineral evolution of other first-row transition elements is being investigated through GeoDeepDive. References: [1] Hazen et al. (2008) Mineral evolution. American Mineralogist, 93, 1693-1720
29 CFR 1910.426 - Mixed-gas diving.
Code of Federal Regulations, 2014 CFR
2014-07-01
... 29 Labor 5 2014-07-01 2014-07-01 false Mixed-gas diving. 1910.426 Section 1910.426 Labor... Mixed-gas diving. (a) General. Employers engaged in mixed-gas diving shall comply with the following requirements, unless otherwise specified. (b) Limits. Mixed-gas diving shall be conducted only when: (1) A...
29 CFR 1910.426 - Mixed-gas diving.
Code of Federal Regulations, 2012 CFR
2012-07-01
... 29 Labor 5 2012-07-01 2012-07-01 false Mixed-gas diving. 1910.426 Section 1910.426 Labor... Mixed-gas diving. (a) General. Employers engaged in mixed-gas diving shall comply with the following requirements, unless otherwise specified. (b) Limits. Mixed-gas diving shall be conducted only when: (1) A...
29 CFR 1910.426 - Mixed-gas diving.
Code of Federal Regulations, 2013 CFR
2013-07-01
... 29 Labor 5 2013-07-01 2013-07-01 false Mixed-gas diving. 1910.426 Section 1910.426 Labor... Mixed-gas diving. (a) General. Employers engaged in mixed-gas diving shall comply with the following requirements, unless otherwise specified. (b) Limits. Mixed-gas diving shall be conducted only when: (1) A...
29 CFR 1910.426 - Mixed-gas diving.
Code of Federal Regulations, 2011 CFR
2011-07-01
... 29 Labor 5 2011-07-01 2011-07-01 false Mixed-gas diving. 1910.426 Section 1910.426 Labor... Mixed-gas diving. (a) General. Employers engaged in mixed-gas diving shall comply with the following requirements, unless otherwise specified. (b) Limits. Mixed-gas diving shall be conducted only when: (1) A...
46 CFR 197.334 - Open diving bells.
Code of Federal Regulations, 2012 CFR
2012-10-01
... 46 Shipping 7 2012-10-01 2012-10-01 false Open diving bells. 197.334 Section 197.334 Shipping... GENERAL PROVISIONS Commercial Diving Operations Equipment § 197.334 Open diving bells. Each open diving... the open bottom and his head in the bubble; (b) Have lifting equipment capable of returning the...
46 CFR 197.334 - Open diving bells.
Code of Federal Regulations, 2014 CFR
2014-10-01
... 46 Shipping 7 2014-10-01 2014-10-01 false Open diving bells. 197.334 Section 197.334 Shipping... GENERAL PROVISIONS Commercial Diving Operations Equipment § 197.334 Open diving bells. Each open diving... the open bottom and his head in the bubble; (b) Have lifting equipment capable of returning the...
46 CFR 197.334 - Open diving bells.
Code of Federal Regulations, 2013 CFR
2013-10-01
... 46 Shipping 7 2013-10-01 2013-10-01 false Open diving bells. 197.334 Section 197.334 Shipping... GENERAL PROVISIONS Commercial Diving Operations Equipment § 197.334 Open diving bells. Each open diving... the open bottom and his head in the bubble; (b) Have lifting equipment capable of returning the...
46 CFR 197.334 - Open diving bells.
Code of Federal Regulations, 2010 CFR
2010-10-01
... 46 Shipping 7 2010-10-01 2010-10-01 false Open diving bells. 197.334 Section 197.334 Shipping... GENERAL PROVISIONS Commercial Diving Operations Equipment § 197.334 Open diving bells. Each open diving... the open bottom and his head in the bubble; (b) Have lifting equipment capable of returning the...
46 CFR 197.334 - Open diving bells.
Code of Federal Regulations, 2011 CFR
2011-10-01
... 46 Shipping 7 2011-10-01 2011-10-01 false Open diving bells. 197.334 Section 197.334 Shipping... GENERAL PROVISIONS Commercial Diving Operations Equipment § 197.334 Open diving bells. Each open diving... the open bottom and his head in the bubble; (b) Have lifting equipment capable of returning the...
29 CFR 1910.426 - Mixed-gas diving.
Code of Federal Regulations, 2010 CFR
2010-07-01
... 29 Labor 5 2010-07-01 2010-07-01 false Mixed-gas diving. 1910.426 Section 1910.426 Labor... OCCUPATIONAL SAFETY AND HEALTH STANDARDS Commercial Diving Operations Specific Operations Procedures § 1910.426 Mixed-gas diving. (a) General. Employers engaged in mixed-gas diving shall comply with the following...
Rozali, A; Khairuddin, H; Sherina, M S; Halim, M Abd; Zin, B Mohd; Sulaiman, A
2008-06-01
This paper describes the pattern of diving accidents treated in a military hospital-based recompression chamber facility in Peninsular Malaysia. A retrospective study was carried out to utilize secondary data from the respective hospital medical records from 1st January 1996 to 31st December 2004. A total of 179 cases categorized as diving accidents received treatment with an average of 20 cases per year. Out of 179 cases, 96.3% (n = 173) received recompression treatment. Majority were males (93.3%), civilians (87.2%) and non-Malaysian citizens (59.2%). Commercial diving activities contributed the highest percentage of diving accidents (48.0%), followed by recreational (39.2%) and military (12.8%). Diving accidents due to commercial diving (n = 86) were mainly contributed by underwater logging activities (87.2%). The most common cases sustained were decompression illness (DCI) (96.1%). Underwater logging and recreational diving activities which contribute to a significant number of diving accidents must be closely monitored. Notification, centralised data registration, medical surveillance as well as legislations related to diving activities in Malaysia are essential to ensure adequate monitoring of diving accidents in the future.
Sivle, L D; Kvadsheim, P H; Fahlman, A; Lam, F P A; Tyack, P L; Miller, P J O
2012-01-01
Anthropogenic underwater sound in the environment might potentially affect the behavior of marine mammals enough to have an impact on their reproduction and survival. Diving behavior of four killer whales (Orcinus orca), seven long-finned pilot whales (Globicephala melas), and four sperm whales (Physeter macrocephalus) was studied during controlled exposures to naval sonar [low frequency active sonar (LFAS): 1-2 kHz and mid frequency active sonar (MFAS): 6-7 kHz] during three field seasons (2006-2009). Diving behavior was monitored before, during and after sonar exposure using an archival tag placed on the animal with suction cups. The tag recorded the animal's vertical movement, and additional data on horizontal movement and vocalizations were used to determine behavioral modes. Killer whales that were conducting deep dives at sonar onset changed abruptly to shallow diving (ShD) during LFAS, while killer whales conducting deep dives at the onset of MFAS did not alter dive mode. When in ShD mode at sonar onset, killer whales did not change their diving behavior. Pilot and sperm whales performed normal deep dives (NDD) during MFAS exposure. During LFAS exposures, long-finned pilot whales mostly performed fewer deep dives and some sperm whales performed shallower and shorter dives. Acoustic recording data presented previously indicates that deep diving (DD) is associated with feeding. Therefore, the observed changes in dive behavior of the three species could potentially reduce the foraging efficiency of the affected animals.
Sivle, L. D.; Kvadsheim, P. H.; Fahlman, A.; Lam, F. P. A.; Tyack, P. L.; Miller, P. J. O.
2012-01-01
Anthropogenic underwater sound in the environment might potentially affect the behavior of marine mammals enough to have an impact on their reproduction and survival. Diving behavior of four killer whales (Orcinus orca), seven long-finned pilot whales (Globicephala melas), and four sperm whales (Physeter macrocephalus) was studied during controlled exposures to naval sonar [low frequency active sonar (LFAS): 1–2 kHz and mid frequency active sonar (MFAS): 6–7 kHz] during three field seasons (2006–2009). Diving behavior was monitored before, during and after sonar exposure using an archival tag placed on the animal with suction cups. The tag recorded the animal's vertical movement, and additional data on horizontal movement and vocalizations were used to determine behavioral modes. Killer whales that were conducting deep dives at sonar onset changed abruptly to shallow diving (ShD) during LFAS, while killer whales conducting deep dives at the onset of MFAS did not alter dive mode. When in ShD mode at sonar onset, killer whales did not change their diving behavior. Pilot and sperm whales performed normal deep dives (NDD) during MFAS exposure. During LFAS exposures, long-finned pilot whales mostly performed fewer deep dives and some sperm whales performed shallower and shorter dives. Acoustic recording data presented previously indicates that deep diving (DD) is associated with feeding. Therefore, the observed changes in dive behavior of the three species could potentially reduce the foraging efficiency of the affected animals. PMID:23087648
Flying after diving: in-flight echocardiography after a scuba diving week.
Cialoni, Danilo; Pieri, Massimo; Balestra, Costantino; Marroni, Alessandro
2014-10-01
Flying after diving may increase decompression sickness risk (DCS), but strong evidence indicating minimum preflight surface intervals (PFSI) is missing. On return flights after a diving week on a live-aboard, 32 divers were examined by in-flight echocardiography with the following protocol: 1) outgoing flight, no previous dive; 2) during the diving week; 3) before the return flight after a 24-h PFSI; and 4) during the return flight. All divers completed similar multiple repetitive dives during the diving week. All dives were equivalent as to inert gas load and gradient factor upon surfacing. No bubbles in the right heart were found in any diver during the outgoing flight or at the preflight control after a 24-h PFSI following the diving week. A significant increase in the number and grade of bubbles was observed during the return flight. However, bubbles were only observed in 6 of the 32 divers. These six divers were the same ones who developed bubbles after every dive. Having observed a 24-h preflight interval, the majority of divers did not develop bubbles during altitude exposure; however, it is intriguing to note that the same subjects who developed significant amounts of bubbles after every dive showed equally significant bubble grades during in-flight echocardiography notwithstanding a correct PFSI. This indicates a possible higher susceptibility to bubble formation in certain individuals, who may need longer PFSI before altitude exposure after scuba diving.
Chambault, Philippine; Pinaud, David; Vantrepotte, Vincent; Kelle, Laurent; Entraygues, Mathieu; Guinet, Christophe; Berzins, Rachel; Bilo, Karin; Gaspar, Philippe; de Thoisy, Benoît; Le Maho, Yvon; Chevallier, Damien
2015-01-01
In response to seasonality and spatial segregation of resources, sea turtles undertake long journeys between their nesting sites and foraging grounds. While satellite tracking has made it possible to outline their migration routes, we still have little knowledge of how they select their foraging grounds and adapt their migration to dynamic environmental conditions. Here, we analyzed the trajectories and diving behavior of 19 adult green turtles (Chelonia mydas) during their post-nesting migration from French Guiana and Suriname to their foraging grounds off the coast of Brazil. First Passage Time analysis was used to identify foraging areas located off Ceará state of Brazil, where the associated habitat corresponds to favorable conditions for seagrass growth, i.e. clear and shallow waters. The dispersal and diving patterns of the turtles revealed several behavioral adaptations to the strong hydrodynamic processes induced by both the North Brazil current and the Amazon River plume. All green turtles migrated south-eastward after the nesting season, confirming that they coped with the strong counter North Brazil current by using a tight corridor close to the shore. The time spent within the Amazon plume also altered the location of their feeding habitats as the longer individuals stayed within the plume, the sooner they initiated foraging. The green turtles performed deeper and shorter dives while crossing the mouth of the Amazon, a strategy which would help turtles avoid the most turbulent upper surface layers of the plume. These adjustments reveal the remarkable plasticity of this green turtle population when reducing energy costs induced by migration. PMID:26398528
Chambault, Philippine; Pinaud, David; Vantrepotte, Vincent; Kelle, Laurent; Entraygues, Mathieu; Guinet, Christophe; Berzins, Rachel; Bilo, Karin; Gaspar, Philippe; de Thoisy, Benoît; Le Maho, Yvon; Chevallier, Damien
2015-01-01
In response to seasonality and spatial segregation of resources, sea turtles undertake long journeys between their nesting sites and foraging grounds. While satellite tracking has made it possible to outline their migration routes, we still have little knowledge of how they select their foraging grounds and adapt their migration to dynamic environmental conditions. Here, we analyzed the trajectories and diving behavior of 19 adult green turtles (Chelonia mydas) during their post-nesting migration from French Guiana and Suriname to their foraging grounds off the coast of Brazil. First Passage Time analysis was used to identify foraging areas located off Ceará state of Brazil, where the associated habitat corresponds to favorable conditions for seagrass growth, i.e. clear and shallow waters. The dispersal and diving patterns of the turtles revealed several behavioral adaptations to the strong hydrodynamic processes induced by both the North Brazil current and the Amazon River plume. All green turtles migrated south-eastward after the nesting season, confirming that they coped with the strong counter North Brazil current by using a tight corridor close to the shore. The time spent within the Amazon plume also altered the location of their feeding habitats as the longer individuals stayed within the plume, the sooner they initiated foraging. The green turtles performed deeper and shorter dives while crossing the mouth of the Amazon, a strategy which would help turtles avoid the most turbulent upper surface layers of the plume. These adjustments reveal the remarkable plasticity of this green turtle population when reducing energy costs induced by migration.
Kotzerka, J.; Hatch, Shyla A.; Garthe, S.
2011-01-01
The Pelagic Cormorant (Phalacrocorax pelagicus) is the most widespread cormorant in the North Pacific, but little is known about its foraging and diving behavior. However, knowledge of seabirds' foraging behavior is important to understanding their function in the marine environment. In 2006, using GPS dataloggers, we studied the foraging behavior of 14 male Pelagic Cormorants rearing chicks on Middleton Island, Alaska. For foraging, the birds had high fidelity to a small area 8 km north of the colony. Within that area, the cormorants' diving activity was of two distinct kinds-near-surface dives (1-6 m) and benthic dives (28-33 m). Individuals were consistent in the depths of their dives, either mostly shallow or mostly deep. Few showed no depth preference. Dive duration, time at maximum depth, and pauses at the water surface between consecutive dives were shorter for shallow dives than for deep dives. The cormorants made dives of both types throughout the day, but the frequency of deep dives increased toward evening. Maximum foraging range was 9 km; maximum total distance traveled per trip was 43.4 km. Trip durations ranged from 0.3 to 7.7 hr. Maximum depth of a dive was 42.2 m, and duration of dives ranged from 4 to 120 sec. We found that Pelagic Cormorants at Middleton Island were faithful to one particular foraging area and individuals dived in distinct patterns. Distinct, specialized foraging behavior may be advantageous in reducing intra- and interspecific competition but may also render the species vulnerable to changing environmental conditions. Copyright ?? The Cooper Ornithological Society 2011.
29 CFR 1910.402 - Definitions.
Code of Federal Regulations, 2011 CFR
2011-07-01
... a specific depth-time exposure or exposures. Dive-guiding operations means leading groups of sports...). Recreational diving instruction means training diving students in the use of recreational diving procedures and...
Code of Federal Regulations, 2013 CFR
2013-10-01
... 43 Public Lands: Interior 1 2013-10-01 2013-10-01 false Skin diving. 15.8 Section 15.8 Public Lands: Interior Office of the Secretary of the Interior KEY LARGO CORAL REEF PRESERVE § 15.8 Skin diving. Diving with camera, or diving for observation and pleasure is permitted and encouraged within the...
Code of Federal Regulations, 2014 CFR
2014-10-01
... 43 Public Lands: Interior 1 2014-10-01 2014-10-01 false Skin diving. 15.8 Section 15.8 Public Lands: Interior Office of the Secretary of the Interior KEY LARGO CORAL REEF PRESERVE § 15.8 Skin diving. Diving with camera, or diving for observation and pleasure is permitted and encouraged within the...
Code of Federal Regulations, 2012 CFR
2012-10-01
... 43 Public Lands: Interior 1 2012-10-01 2011-10-01 true Skin diving. 15.8 Section 15.8 Public Lands: Interior Office of the Secretary of the Interior KEY LARGO CORAL REEF PRESERVE § 15.8 Skin diving. Diving with camera, or diving for observation and pleasure is permitted and encouraged within the Preserve. ...
Code of Federal Regulations, 2010 CFR
2010-10-01
... 43 Public Lands: Interior 1 2010-10-01 2010-10-01 false Skin diving. 15.8 Section 15.8 Public Lands: Interior Office of the Secretary of the Interior KEY LARGO CORAL REEF PRESERVE § 15.8 Skin diving. Diving with camera, or diving for observation and pleasure is permitted and encouraged within the...
Code of Federal Regulations, 2011 CFR
2011-10-01
... 43 Public Lands: Interior 1 2011-10-01 2011-10-01 false Skin diving. 15.8 Section 15.8 Public Lands: Interior Office of the Secretary of the Interior KEY LARGO CORAL REEF PRESERVE § 15.8 Skin diving. Diving with camera, or diving for observation and pleasure is permitted and encouraged within the...
46 CFR 197.432 - Surface-supplied air diving.
Code of Federal Regulations, 2012 CFR
2012-10-01
... 46 Shipping 7 2012-10-01 2012-10-01 false Surface-supplied air diving. 197.432 Section 197.432...-supplied air diving. The diving supervisor shall insure that— (a) Surface-supplied air diving is conducted... space; and (f) The surface-supplied air diver has the equipment required by § 197.346 (b) or (d). ...
46 CFR 197.432 - Surface-supplied air diving.
Code of Federal Regulations, 2013 CFR
2013-10-01
... 46 Shipping 7 2013-10-01 2013-10-01 false Surface-supplied air diving. 197.432 Section 197.432...-supplied air diving. The diving supervisor shall insure that— (a) Surface-supplied air diving is conducted... space; and (f) The surface-supplied air diver has the equipment required by § 197.346 (b) or (d). ...
29 CFR 1910.425 - Surface-supplied air diving.
Code of Federal Regulations, 2013 CFR
2013-07-01
... 29 Labor 5 2013-07-01 2013-07-01 false Surface-supplied air diving. 1910.425 Section 1910.425... Procedures § 1910.425 Surface-supplied air diving. (a) General. Employers engaged in surface-supplied air...-supplied air diving shall not be conducted at depths deeper than 190 fsw, except that dives with bottom...
29 CFR 1910.425 - Surface-supplied air diving.
Code of Federal Regulations, 2012 CFR
2012-07-01
... 29 Labor 5 2012-07-01 2012-07-01 false Surface-supplied air diving. 1910.425 Section 1910.425... Procedures § 1910.425 Surface-supplied air diving. (a) General. Employers engaged in surface-supplied air...-supplied air diving shall not be conducted at depths deeper than 190 fsw, except that dives with bottom...
29 CFR 1910.425 - Surface-supplied air diving.
Code of Federal Regulations, 2010 CFR
2010-07-01
... 29 Labor 5 2010-07-01 2010-07-01 false Surface-supplied air diving. 1910.425 Section 1910.425... Procedures § 1910.425 Surface-supplied air diving. (a) General. Employers engaged in surface-supplied air...-supplied air diving shall not be conducted at depths deeper than 190 fsw, except that dives with bottom...
46 CFR 197.432 - Surface-supplied air diving.
Code of Federal Regulations, 2010 CFR
2010-10-01
... 46 Shipping 7 2010-10-01 2010-10-01 false Surface-supplied air diving. 197.432 Section 197.432...-supplied air diving. The diving supervisor shall insure that— (a) Surface-supplied air diving is conducted... space; and (f) The surface-supplied air diver has the equipment required by § 197.346 (b) or (d). ...
29 CFR 1910.425 - Surface-supplied air diving.
Code of Federal Regulations, 2011 CFR
2011-07-01
... 29 Labor 5 2011-07-01 2011-07-01 false Surface-supplied air diving. 1910.425 Section 1910.425... Procedures § 1910.425 Surface-supplied air diving. (a) General. Employers engaged in surface-supplied air...-supplied air diving shall not be conducted at depths deeper than 190 fsw, except that dives with bottom...
46 CFR 197.432 - Surface-supplied air diving.
Code of Federal Regulations, 2011 CFR
2011-10-01
... 46 Shipping 7 2011-10-01 2011-10-01 false Surface-supplied air diving. 197.432 Section 197.432...-supplied air diving. The diving supervisor shall insure that— (a) Surface-supplied air diving is conducted... space; and (f) The surface-supplied air diver has the equipment required by § 197.346 (b) or (d). ...
29 CFR 1910.410 - Qualifications of dive team.
Code of Federal Regulations, 2013 CFR
2013-07-01
... 29 Labor 5 2013-07-01 2013-07-01 false Qualifications of dive team. 1910.410 Section 1910.410... Requirements § 1910.410 Qualifications of dive team. (a) General. (1) Each dive team member shall have the experience or training necessary to perform assigned tasks in a safe and healthful manner. (2) Each dive team...
29 CFR 1910.410 - Qualifications of dive team.
Code of Federal Regulations, 2014 CFR
2014-07-01
... 29 Labor 5 2014-07-01 2014-07-01 false Qualifications of dive team. 1910.410 Section 1910.410... Requirements § 1910.410 Qualifications of dive team. (a) General. (1) Each dive team member shall have the experience or training necessary to perform assigned tasks in a safe and healthful manner. (2) Each dive team...
29 CFR 1910.410 - Qualifications of dive team.
Code of Federal Regulations, 2011 CFR
2011-07-01
... 29 Labor 5 2011-07-01 2011-07-01 false Qualifications of dive team. 1910.410 Section 1910.410... Requirements § 1910.410 Qualifications of dive team. (a) General. (1) Each dive team member shall have the experience or training necessary to perform assigned tasks in a safe and healthful manner. (2) Each dive team...
29 CFR 1910.410 - Qualifications of dive team.
Code of Federal Regulations, 2012 CFR
2012-07-01
... 29 Labor 5 2012-07-01 2012-07-01 false Qualifications of dive team. 1910.410 Section 1910.410... Requirements § 1910.410 Qualifications of dive team. (a) General. (1) Each dive team member shall have the experience or training necessary to perform assigned tasks in a safe and healthful manner. (2) Each dive team...
Provisional report on diving-related fatalities in Australian waters 2004.
Walker, Douglas; Lippmann, John; Lawrence, Chris; Huston, John; Fock, Andrew
2009-09-01
An individual case review of diving-related deaths reported to have occurred in Australia in 2004 was conducted as part of the DAN Asia-Pacific dive fatality reporting project. The case studies were compiled using reports from witnesses, the police and coroners. In each case, the particular circumstances of the accident are provided, and also details from the post-mortem examination, where available. In total, there were 22 reported fatalities, all male. Nine deaths occurred while snorkelling and/or breath-hold diving, 10 while scuba diving, one just prior to scuba diving, one while using surface-supply breath apparatus and one while diving with a rebreather. In this series, cardiac-related issues were thought to have contributed to the deaths of five snorkel divers and three scuba divers, and in one person who was about to go diving. Three of the deaths in breath-hold divers were likely to have been associated with apnoeic hypoxia blackout. Pre-existing medical conditions, inexperience, time away from diving, inadequate supervision, and diving without appropriate training were features in several scuba deaths in this series.
Advanced instrumentation for research in diving and hyperbaric medicine.
Sieber, Arne; L'Abbate, Antonio; Kuch, Benjamin; Wagner, Matthias; Benassi, Antonio; Passera, Mirko; Bedini, Remo
2010-01-01
Improving the safety of diving and increasing knowledge about the adaptation of the human body to underwater and hyperbaric environment require specifically developed underwater instrumentation for physiological measurements. In fact, none of the routine clinical devices for health control is suitable for in-water and/or under-pressure operation. The present paper addresses novel technological acquisitions and the development of three dedicated devices: * an underwater data logger for recording O2 saturation (reflective pulsoxymetry), two-channel ECG, depth and temperature; * an underwater blood pressure meter based on the oscillometric method; and * an underwater echography system. Moreover, examples of recordings are presented and discussed.
Sperm whale dive behavior characteristics derived from intermediate-duration archival tag data.
Irvine, Ladd; Palacios, Daniel M; Urbán, Jorge; Mate, Bruce
2017-10-01
Here, we describe the diving behavior of sperm whales ( Physeter macrocephalus ) using the Advanced Dive Behavior (ADB) tag, which records depth data at 1-Hz resolution and GPS-quality locations for over 1 month, before releasing from the whale for recovery. A total of 27 ADB tags were deployed on sperm whales in the central Gulf of California, Mexico, during spring 2007 and 2008, of which 10 were recovered for data download. Tracking durations of all tags ranged from 0 to 34.5 days (median = 2.3 days), and 0.6 to 26.6 days (median = 5.0 days) for recovered tags. Recovered tags recorded a median of 50.8 GPS-quality locations and 42.6 dives per day. Dive summary metrics were generated for archived dives and were subsequently classified into six categories using hierarchical cluster analysis. A mean of 77% of archived dives per individual were one of four dive categories with median Maximum Dive Depth >290 m (V-shaped, Mid-water, Benthic, or Variable), likely associated with foraging. Median Maximum Dive Depth was <30 m for the other two categories (Short- and Long-duration shallow dives), likely representing socializing or resting behavior. Most tagged whales remained near the tagging area during the tracking period, but one moved north of Isla Tiburón, where it appeared to regularly dive to, and travel along the seafloor. Three whales were tagged on the same day in 2007 and subsequently traveled in close proximity (<1 km) for 2 days. During this period, the depth and timing of their dives were not coordinated, suggesting they were foraging on a vertically heterogeneous prey field. The multiweek dive records produced by ADB tags enabled us to generate a robust characterization of the diving behavior, activity budget, and individual variation for an important predator of the mesopelagos over temporal and spatial scales not previously possible.
Estimating the risk of a scuba diving fatality in Australia.
Lippmann, John; Stevenson, Christopher; McD Taylor, David; Williams, Jo
2016-12-01
There are few data available on which to estimate the risk of death for Australian divers. This report estimates the risk of a scuba diving fatality for Australian residents, international tourists diving in Queensland, and clients of a large Victorian dive operator. Numerators for the estimates were obtained from the Divers Alert Network Asia-Pacific dive fatality database. Denominators were derived from three sources: Participation in Exercise, Recreation and Sport Surveys, 2001-2010 (Australian resident diving activity data); Tourism Research Australia surveys of international visitors to Queensland 2006-2014 and a dive operator in Victoria 2007-2014. Annual fatality rates (AFR) and 95% confidence intervals (95% CI) were calculated using an exact binomial test. Estimated AFRs were: 0.48 (0.37-0.59) deaths per 100,000 dives, or 8.73 (6.85-10.96) deaths per 100,000 divers for Australian residents; 0.12 (0.05-0.25) deaths per 100,000 dives, or 0.46 (0.20-0.91) deaths per 100,000 divers for international visitors to Queensland; and 1.64 (0.20-5.93) deaths per 100,000 dives for the dive operator in Victoria. On a per diver basis, Australian residents are estimated to be almost twenty times more likely to die whilst scuba diving than are international visitors to Queensland, or to lower than fourfold on a per dive basis. On a per dive basis, divers in Victoria are fourteen times more likely to die than are Queensland international tourists. Although some of the estimates are based on potentially unreliable denominator data extrapolated from surveys, the diving fatality rates in Australia appear to vary by State, being considerably lower in Queensland than in Victoria. These estimates are similar to or lower than comparable overseas estimates, although reliability of all such measurements varies with study size and accuracy of the data available.
Blood Oxygen Conservation in Diving Sea Lions: How Low Does Oxygen Really Go?
2014-09-30
arterial saturations during dives as long as 10 min (Meir and Ponganis 2009). It is also notable that a severe bradycardia during descent occurs in deep...are not completely depleted in even the longest of sea lion dives. The severe bradycardia during deep dives contributes to the preservation of the...3332-3341. McDonald, B.I., and P.J. Ponganis. 2014. Deep-diving sea lions exhibit extreme bradycardia in long- duration dives. Journal of Experimental
2015-05-28
Diver Characteristics Appendix E Diving Schedule Appendix F Medical Incidents Appendix G UBA Gas Compositions iv ACKNOWLEDGEMENTS The...experimental dives (median = 3). The schedule of each diver’s participation in experimental dives is given in Appendix E . Divers were required to avoid any...divers’ participation on each test schedule is given in Appendix E . The numbers of completed man-dives on the two schedules are not multiples of the
Board Diving Regulations in Public Swimming Pools and Risk of Injury.
Williams, David; Odin, Louise
2016-06-01
Public session access to diving boards is one of the stepping stones for those wishing to develop their skills in the sport of diving. The extent to which certain dive forms are considered risky (forward/backward/rotations) and therefore not permitted is a matter for local pool managers. In Study 1, 20 public pools with diving facilities responded to a U.K. survey concerning their diving regulation policy and related injury incidence in the previous year. More restrictive regulation of dive forms was not associated with a decrease in injuries (rs [42] = -0.20, p = 0.93). In Study 2, diving risk perception and attitudes towards regulation were compared between experienced club divers (N = 22) and nondivers (N = 22). Risk was perceived to be lower for those with experience, and these people favored less regulation. The findings are interpreted in terms of a risk thermostat model, where for complex physical performance activities such as diving, individuals may exercise caution in proportion to their ability and previous experience of success and failure related to the activity. Though intuitively appealing, restrictive regulation of public pool diving may be ineffective in practice because risk is not simplistically associated with dive forms, and divers are able to respond flexibly to risk by exercising caution where appropriate. © 2015 Society for Risk Analysis.
Sustaining Statewide Disaster Response Capabilities from a Fire Service Perspective
2013-03-01
2010). These increased capabilities can be vital to the success of response operations that involve automobile accidents, train wrecks , boat...Trench Rescue Subterranean Rescue Dive Rescue Wilderness Rescue 2. 1670 Operations and Training for Technical Search and Rescue Incidents The
Uhlig, Florian; Muth, Claus-Martin; Tetzlaff, Kay; Koch, Andreas; Leberle, Richard; Georgieff, Michael; Winkler, Bernd E
2014-06-01
Full-face-masks (FFM) prevent the diver's face from cold and can support nasal breathing underwater. The aim of the study was to evaluate the effect of the use of FFMs on lung function and wellbeing. Twenty-one, healthy, non-asthmatic divers performed two cold-water dives (4⁰C, 25 min, 10 metres' depth) - one with a FFM and the other with a standard scuba regulator (SSR). Spirometry was performed before and after each dive and well-being and cold sensation were assessed after the dives. Significant decreases in forced vital capacity (FVC), forced expiratory volume in one second (FEV₁) and midexpiratory flow at 75% of FVC (MEF₇₅) occurred after both FFM and SSR dives. Changes in FVC and FEV₁ did not differ significantly between FFM and SSR dives. However, the mid-expiratory flows measured at 50% and 25% of FVC (MEF₅₀ and MEF₂₅) were significantly lower 10 minutes after the FFM dive compared to 10 minutes after the SSR dive. The wellbeing and cold sensation of the divers were significantly improved with FFM dives compared to SSR dives. Cold-water dives during wintertime can be associated with airway narrowing. During cold-water dives, the use of a FFM appears to reduce the cold sensation and enhance the well-being of the divers. However, a FFM does not appear to prevent airway narrowing in healthy, non-asthmatic subjects.
Li, H; Wang, Y; Dou, R; Wang, L; Xu, L L; Li, X J; Zhang, J M
2017-01-11
Objective: To evaluate the effects of side-cut angles on corneal biomechanical properties after femtosecond laser assisted-laser in situ keratomileusis (FS-LASIK). Methods: In this clinical control study, 97 right eyes of 97 patients with myopia and myopic astigmatism undergoing FS-LASIK surgery were included randomly. Two types of side-cut angles were designed for the flap, 90° in 48 eyes and 130° in 49 eyes. The values of corneal resistance factor (CRF), cornea hysteresis (CH) and 37 biomechanical waveform parameters were measured using the ocular response analyzer preoperatively and at 1 and 3 months postoperatively. The comparison between the two groups at different follow-up time points was made with one-way analysis of variance, and the comparison among different follow-up time points in each group was performed with the repeated measures analysis of variance. Results: The mean CRF, CH, p area, h, dive1, dive2, w11 and w21 in both groups showed significant reduction at 1 month postoperatively (group 90°: CRF 10.49 vs. 6.85, CH 10.02 vs. 7.55, p1 area 3 537.54 vs. 1 918.07, h1 404.74 vs. 283.25, dive1 350.63 vs. 243.33, w11 10.50 vs. 7.58. group 130°: CRF 10.14 vs. 6.38, CH 9.93 vs. 7.13, p1 area 3 498.93 vs. 2 038.74, h1 411.93 vs. 304.49, dive1 352.51 vs. 265.12, w11 10.55 vs. 7.78. P< 0.001) . The mean CRF, CH, p area, w, h, dive1 and dive2 in both groups showed significant reduction at 3 months postoperatively (group 90°: CRF 10.49 vs. 6.60, CH 10.02 vs. 7.65, p1 area 3537.54 vs. 2042.91, h1 404.74 vs. 307.77, dive1 350.63 vs. 263.33, w11 10.50 vs. 7.58. group 130°: CRF 10.14 vs. 6.45, CH 9.93 vs. 7.67, p1 area 3 498.93 vs. 2 187.97, h1 411.93 vs. 327.39, dive1 352.51 vs. 284.26, w11 10.55 vs. 7.61. P< 0.001). The mean path in both groups showed significant increase at 1 and 3 months postoperatively (group 90°: path1 23.14 vs. 30.50 vs. 30.79. group 130°: path1 24.12 vs. 32.18 vs. 31.49. P< 0.001). The mean aplhf, bindex, dive2 and path2 between both groups at 1 month postoperatively showed significant difference ( F= 8.609, 7.482, 5.238, 4.885, P <0.05). The mean path11 between both groups at 3 months postoperatively showed significant difference ( F= 6.160, P <0.05). Conclusions: Corneal biomechanical properties significantly decrease after FS-LASIK, and different side-cut angles have some effect on corneal biomechanical properties after FS-LASIK. The corneal deformation is more stable after flap creation with an obtuse side-cut angle making the cornea under stress. Combined with waveform parameters of the ocular response analyzer, more subtle changes of corneal biomechanical properties can be found. (Chin J Ophthalmol, 2017, 53:23-32) .
Irgens, Ågot; Troland, Kari; Grønning, Marit
2017-01-01
The aim of the present study was to explore the potential differences between female and male professional divers with regards to demographics, diving certificates, areas of diving, diving activity and health effects. The Norwegian Labour Inspection Authority's Diving certificate register contains data on all professional inshore divers who have held a certificate at any time since 1980. Forty nine per cent of these divers responded to the "Norwegian diver 2011" questionnaire. Of these divers 64 female and 1327 male divers completed the questionnaire about their professional diving career, certificate, year of onset and the year they stopped diving professionally if they were not still active in the diving industry. The level of general education was higher among female divers. More males than females were fully certified in diving. The mean age was lower among female than male fully certified divers. Fully certified female divers reported a lower total number of dives, shallower dives and diving for a shorter period of time than the male divers. They also had a lower percentage of work within the quay/construction sector and more often worked as teachers/instructors. A lower percentage of fully certified females than males had experienced decompression sickness (16.7% vs. 26.9%). Life-threatening events and psychologically challenging events were less common among females, as were adverse health effects. No such gender differences were seen for divers with a restricted certificate. The fully certified, female professional divers in our study had a very short diving career, reported fewer and shallower dives, and chose less physically demanding jobs than their male counterparts. They also had a higher level of education, reported less health problems and a better quality of life. The health effects seem to be related to the type of work rather than to gender.
Flipper stroke rate and venous oxygen levels in free-ranging California sea lions.
Tift, Michael S; Hückstädt, Luis A; McDonald, Birgitte I; Thorson, Philip H; Ponganis, Paul J
2017-04-15
The depletion rate of the blood oxygen store, development of hypoxemia and dive capacity are dependent on the distribution and rate of blood oxygen delivery to tissues while diving. Although blood oxygen extraction by working muscle would increase the blood oxygen depletion rate in a swimming animal, there is little information on the relationship between muscle workload and blood oxygen depletion during dives. Therefore, we examined flipper stroke rate, a proxy of muscle workload, and posterior vena cava oxygen profiles in four adult female California sea lions ( Zalophus californianus ) during foraging trips at sea. Flipper stroke rate analysis revealed that sea lions minimized muscle metabolism with a stroke-glide strategy when diving, and exhibited prolonged glides during the descent of deeper dives (>100 m). During the descent phase of these deep dives, 55±21% of descent was spent gliding, with the longest glides lasting over 160 s and covering a vertical distance of 340 m. Animals also consistently glided to the surface from 15 to 25 m depth during these deeper dives. Venous hemoglobin saturation ( S O 2 ) profiles were highly variable throughout dives, with values occasionally increasing during shallow dives. The relationship between S O 2 and flipper stroke rate was weak during deeper dives, while this relationship was stronger during shallow dives. We conclude that (1) the depletion of oxygen in the posterior vena cava in deep-diving sea lions is not dependent on stroke effort, and (2) stroke-glide patterns during dives contribute to a reduction of muscle metabolic rate. © 2017. Published by The Company of Biologists Ltd.
High-affinity hemoglobin and blood oxygen saturation in diving emperor penguins.
Meir, Jessica U; Ponganis, Paul J
2009-10-01
The emperor penguin (Aptenodytes forsteri) thrives in the Antarctic underwater environment, diving to depths greater than 500 m and for durations longer than 23 min. To examine mechanisms underlying the exceptional diving ability of this species and further describe blood oxygen (O2) transport and depletion while diving, we characterized the O2-hemoglobin (Hb) dissociation curve of the emperor penguin in whole blood. This allowed us to (1) investigate the biochemical adaptation of Hb in this species, and (2) address blood O2 depletion during diving, by applying the dissociation curve to previously collected partial pressure of O2 (PO2) profiles to estimate in vivo Hb saturation (SO2) changes during dives. This investigation revealed enhanced Hb-O2 affinity (P50=28 mmHg, pH 7.5) in the emperor penguin, similar to high-altitude birds and other penguin species. This allows for increased O2 at low blood PO2 levels during diving and more complete depletion of the respiratory O2 store. SO2 profiles during diving demonstrated that arterial SO2 levels are maintained near 100% throughout much of the dive, not decreasing significantly until the final ascent phase. End-of-dive venous SO2 values were widely distributed and optimization of the venous blood O2 store resulted from arterialization and near complete depletion of venous blood O2 during longer dives. The estimated contribution of the blood O2 store to diving metabolic rate was low and highly variable. This pattern is due, in part, to the influx of O2 from the lungs into the blood during diving, and variable rates of tissue O2 uptake.
Dark chocolate reduces endothelial dysfunction after successive breath-hold dives in cool water.
Theunissen, Sigrid; Schumacker, Julie; Guerrero, François; Tillmans, Frauke; Boutros, Antoine; Lambrechts, Kate; Mazur, Aleksandra; Pieri, Massimo; Germonpré, Peter; Balestra, Costantino
2013-12-01
The aim of this study is to observe the effects of dark chocolate on endothelial function after a series of successive apnea dives in non-thermoneutral water. Twenty breath-hold divers were divided into two groups: a control group (8 males and 2 females) and a chocolate group (9 males and 1 female). The control group was asked to perform a series of dives to 20 m adding up to 20 min in the quiet diving pool of Conflans-Ste-Honorine (Paris, France), water temperature was 27 °C. The chocolate group performed the dives 1 h after ingestion of 30 g of dark chocolate. Flow-mediated dilatation (FMD), digital photoplethysmography, nitric oxide (NO), and peroxynitrite ONOO−) levels were measured before and after each series of breath-hold dives. A significant decrease in FMD was observed in the control group after the dives (95.28 ± 2.9 % of pre-dive values, p < 0.001) while it was increased in the chocolate group (104.1 ± 2.9 % of pre-dive values, p < 0.01). A decrease in the NO level was observed in the control group (86.76 ± 15.57 %, p < 0.05) whereas no difference was shown in the chocolate group (98.44 ± 31.86 %, p > 0.05). No differences in digital photoplethysmography and peroxynitrites were observed between before and after the dives. Antioxidants contained in dark chocolate scavenge free radicals produced during breath-hold diving. Ingestion of 30 g of dark chocolate 1 h before the dive can thus prevent endothelial dysfunction which can be observed after a series of breath-hold dives.
Pelland, Noel A; Sterling, Jeremy T; Lea, Mary-Anne; Bond, Nicholas A; Ream, Rolf R; Lee, Craig M; Eriksen, Charles C
2014-01-01
Behavioral responses by top marine predators to oceanographic features such as eddies, river plumes, storms, and coastal topography suggest that biophysical interactions in these zones affect predators' prey, foraging behaviors, and potentially fitness. However, examining these pathways is challenged by the obstacles inherent in obtaining simultaneous observations of surface and subsurface environmental fields and predator behavior. In this study, migratory movements and, in some cases, diving behavior of 40 adult female northern fur seals (NFS; Callorhinus ursinus) were quantified across their range and compared to remotely-sensed environmental data in the Gulf of Alaska and California Current ecosystems, with a particular focus off the coast of Washington State (USA)--a known foraging ground for adult female NFS and where autonomous glider sampling allowed opportunistic comparison of seal behavior to subsurface biophysical measurements. The results show that in these ecosystems, adult female habitat utilization was concentrated near prominent coastal topographic, riverine, or inlet features and within 200 km of the continental shelf break. Seal dive depths, in most ecosystems, were moderated by surface light level (solar or lunar), mirroring known behaviors of diel vertically-migrating prey. However, seal dives differed in the California Current ecosystem due to a shift to more daytime diving concentrated at or below the surface mixed layer base. Seal movement models indicate behavioral responses to season, ecosystem, and surface wind speeds; individuals also responded to mesoscale eddies, jets, and the Columbia River plume. Foraging within small scale surface features is consistent with utilization of the inner coastal transition zone and habitats near coastal capes, which are known eddy and filament generation sites. These results contribute to our knowledge of NFS migratory patterns by demonstrating surface and subsurface behavioral responses to a spatially and temporally dynamic ocean environment, thus reflecting its influence on associated NFS prey species.
Pelland, Noel A.; Sterling, Jeremy T.; Lea, Mary-Anne; Bond, Nicholas A.; Ream, Rolf R.; Lee, Craig M.; Eriksen, Charles C.
2014-01-01
Behavioral responses by top marine predators to oceanographic features such as eddies, river plumes, storms, and coastal topography suggest that biophysical interactions in these zones affect predators' prey, foraging behaviors, and potentially fitness. However, examining these pathways is challenged by the obstacles inherent in obtaining simultaneous observations of surface and subsurface environmental fields and predator behavior. In this study, migratory movements and, in some cases, diving behavior of 40 adult female northern fur seals (NFS; Callorhinus ursinus) were quantified across their range and compared to remotely-sensed environmental data in the Gulf of Alaska and California Current ecosystems, with a particular focus off the coast of Washington State (USA) – a known foraging ground for adult female NFS and where autonomous glider sampling allowed opportunistic comparison of seal behavior to subsurface biophysical measurements. The results show that in these ecosystems, adult female habitat utilization was concentrated near prominent coastal topographic, riverine, or inlet features and within 200 km of the continental shelf break. Seal dive depths, in most ecosystems, were moderated by surface light level (solar or lunar), mirroring known behaviors of diel vertically-migrating prey. However, seal dives differed in the California Current ecosystem due to a shift to more daytime diving concentrated at or below the surface mixed layer base. Seal movement models indicate behavioral responses to season, ecosystem, and surface wind speeds; individuals also responded to mesoscale eddies, jets, and the Columbia River plume. Foraging within small scale surface features is consistent with utilization of the inner coastal transition zone and habitats near coastal capes, which are known eddy and filament generation sites. These results contribute to our knowledge of NFS migratory patterns by demonstrating surface and subsurface behavioral responses to a spatially and temporally dynamic ocean environment, thus reflecting its influence on associated NFS prey species. PMID:25153524
Andersen, Julie M.; Skern-Mauritzen, Mette; Boehme, Lars; Wiersma, Yolanda F.; Rosing-Asvid, Aqqalu; Hammill, Mike O.; Stenson, Garry B.
2013-01-01
With the exception of relatively brief periods when they reproduce and moult, hooded seals, Cystophora cristata, spend most of the year in the open ocean where they undergo feeding migrations to either recover or prepare for the next fasting period. Valuable insights into habitat use and diving behaviour during these periods have been obtained by attaching Satellite Relay Data Loggers (SRDLs) to 51 Northwest (NW) Atlantic hooded seals (33 females and 18 males) during ice-bound fasting periods (2004−2008). Using General Additive Models (GAMs) we describe habitat use in terms of First Passage Time (FPT) and analyse how bathymetry, seasonality and FPT influence the hooded seals’ diving behaviour described by maximum dive depth, dive duration and surface duration. Adult NW Atlantic hooded seals exhibit a change in diving activity in areas where they spend >20 h by increasing maximum dive depth, dive duration and surface duration, indicating a restricted search behaviour. We found that male and female hooded seals are spatially segregated and that diving behaviour varies between sexes in relation to habitat properties and seasonality. Migration periods are described by increased dive duration for both sexes with a peak in May, October and January. Males demonstrated an increase in dive depth and dive duration towards May (post-breeding/pre-moult) and August–October (post-moult/pre-breeding) but did not show any pronounced increase in surface duration. Females dived deepest and had the highest surface duration between December and January (post-moult/pre-breeding). Our results suggest that the smaller females may have a greater need to recover from dives than that of the larger males. Horizontal segregation could have evolved as a result of a resource partitioning strategy to avoid sexual competition or that the energy requirements of males and females are different due to different energy expenditure during fasting periods. PMID:24282541
Argon used as dry suit insulation gas for cold-water diving.
Vrijdag, Xavier Ce; van Ooij, Pieter-Jan Am; van Hulst, Robert A
2013-06-03
Cold-water diving requires good thermal insulation because hypothermia is a serious risk. Water conducts heat more efficiently compared to air. To stay warm during a dive, the choice of thermal protection should be based on physical activity, the temperature of the water, and the duration of exposure. A dry suit, a diving suit filled with gas, is the most common diving suit in cold water. Air is the traditional dry suit inflation gas, whereas the thermal conductivity of argon is approximately 32% lower compared to that of air. This study evaluates the benefits of argon, compared to air, as a thermal insulation gas for a dry suit during a 1-h cold-water dive by divers of the Royal Netherlands Navy. Seven male Special Forces divers made (in total) 19 dives in a diving basin with water at 13 degrees C at a depth of 3 m for 1 h in upright position. A rubber dry suit and woollen undergarment were used with either argon (n = 13) or air (n = 6) (blinded to the divers) as suit inflation gas. Core temperature was measured with a radio pill during the dive. Before, halfway, and after the dive, subjective thermal comfort was recorded using a thermal comfort score. No diver had to abort the test due to cold. No differences in core temperature and thermal comfort score were found between the two groups. Core temperature remained unchanged during the dives. Thermal comfort score showed a significant decrease in both groups after a 60-min dive compared to baseline. In these tests the combination of the dry suit and undergarment was sufficient to maintain core temperature and thermal comfort for a dive of 1h in water at 13 degrees C. The use of argon as a suit inflation gas had no added value for thermal insulation compared to air for these dives.
Field Management of Accidental Hypothermia during Diving
1990-01-01
diving operations. 4 3. Diving after tending for prolonged periods leading to hypothermia before diving. 4. Dry suit undergarments that are wet due to...dives, reducing insulation of the undergarment, and then remaining at rest for prolonged decompressiofi stops. 6. Inadequate thermal insulation: wet ...suit instead of dry suit, undergarment selection too thin, too compressible or of poor insulation when wet (63-64), inadequate thermal insulation of the
Code of Federal Regulations, 2014 CFR
2014-07-01
... (i.e., commercially pre-packed), disposable scrubber cartridge containing a CO2-sorbent material that... permit a diver to use a dive-decompression computer designed to regulate decompression when the dive...-activity test; (ii) The RoTap shaker and nested-sieves test; (iii) The Navy Experimental Diving Unit (“NEDU...
Code of Federal Regulations, 2010 CFR
2010-07-01
... (i.e., commercially pre-packed), disposable scrubber cartridge containing a CO2-sorbent material that... permit a diver to use a dive-decompression computer designed to regulate decompression when the dive...-activity test; (ii) The RoTap shaker and nested-sieves test; (iii) The Navy Experimental Diving Unit (“NEDU...
Code of Federal Regulations, 2013 CFR
2013-07-01
... (i.e., commercially pre-packed), disposable scrubber cartridge containing a CO2-sorbent material that... permit a diver to use a dive-decompression computer designed to regulate decompression when the dive...-activity test; (ii) The RoTap shaker and nested-sieves test; (iii) The Navy Experimental Diving Unit (“NEDU...
Code of Federal Regulations, 2011 CFR
2011-07-01
... (i.e., commercially pre-packed), disposable scrubber cartridge containing a CO2-sorbent material that... permit a diver to use a dive-decompression computer designed to regulate decompression when the dive...-activity test; (ii) The RoTap shaker and nested-sieves test; (iii) The Navy Experimental Diving Unit (“NEDU...
Code of Federal Regulations, 2012 CFR
2012-07-01
... (i.e., commercially pre-packed), disposable scrubber cartridge containing a CO2-sorbent material that... permit a diver to use a dive-decompression computer designed to regulate decompression when the dive...-activity test; (ii) The RoTap shaker and nested-sieves test; (iii) The Navy Experimental Diving Unit (“NEDU...
Gerlinsky, Carling D; Haulena, Martin; Trites, Andrew W; Rosen, David A S
2018-03-01
Decreased health may have lowered the birth and survival rates of Steller sea lions ( Eumetopias jubatus) in the Gulf of Alaska and Aleutian Islands over the past 30 yr. Reference ranges for clinical hematology and serum chemistry parameters needed to assess the health of wild sea lion populations are limited. Here, blood parameters were serially measured in 12 captive female Steller sea lions ranging in age from 3 wk to 16 yr to establish baseline values and investigate age-related changes. Whether diving activity affects hematology parameters in animals swimming in the ocean compared with animals in a traditional aquarium setting was also examined. Almost all blood parameters measured exhibited significant changes with age. Many of the age-related changes reflected developmental life history changes, including a change in diet during weaning, an improvement of diving capacity, and the maturity of the immune system. Mean corpuscular hemoglobin and mean corpuscular volume were also higher in the ocean diving group compared with the aquarium group, likely reflecting responses to increased exercise regimes. These data provide ranges of hematology and serum chemistry values needed to evaluate and compare the health and nutritional status of captive and wild Steller sea lions.
Testing optimal foraging theory in a penguin-krill system.
Watanabe, Yuuki Y; Ito, Motohiro; Takahashi, Akinori
2014-03-22
Food is heterogeneously distributed in nature, and understanding how animals search for and exploit food patches is a fundamental challenge in ecology. The classic marginal value theorem (MVT) formulates optimal patch residence time in response to patch quality. The MVT was generally proved in controlled animal experiments; however, owing to the technical difficulties in recording foraging behaviour in the wild, it has been inadequately examined in natural predator-prey systems, especially those in the three-dimensional marine environment. Using animal-borne accelerometers and video cameras, we collected a rare dataset in which the behaviour of a marine predator (penguin) was recorded simultaneously with the capture timings of mobile, patchily distributed prey (krill). We provide qualitative support for the MVT by showing that (i) krill capture rate diminished with time in each dive, as assumed in the MVT, and (ii) dive duration (or patch residence time, controlled for dive depth) increased with short-term, dive-scale krill capture rate, but decreased with long-term, bout-scale krill capture rate, as predicted from the MVT. Our results demonstrate that a single environmental factor (i.e. patch quality) can have opposite effects on animal behaviour depending on the time scale, emphasizing the importance of multi-scale approaches in understanding complex foraging strategies.
High-altitude diving in river otters: coping with combined hypoxic stresses.
Crait, Jamie R; Prange, Henry D; Marshall, Noah A; Harlow, Henry J; Cotton, Clark J; Ben-David, Merav
2012-01-15
River otters (Lontra canadensis) are highly active, semi-aquatic mammals indigenous to a range of elevations and represent an appropriate model for assessing the physiological responses to diving at altitude. In this study, we performed blood gas analyses and compared blood chemistry of river otters from a high-elevation (2357 m) population at Yellowstone Lake with a sea-level population along the Pacific coast. Comparisons of oxygen dissociation curves (ODC) revealed no significant difference in hemoglobin-oxygen (Hb-O(2)) binding affinity between the two populations - potentially because of demands for tissue oxygenation. Instead, high-elevation otters had greater Hb concentrations (18.7 g dl(-1)) than sea-level otters (15.6 g dl(-1)). Yellowstone otters displayed higher levels of the vasodilator nitric oxide (NO), and half the concentration of the serum protein albumin, possibly to compensate for increased blood viscosity. Despite compensation in several hematological and serological parameters, theoretical aerobic dive limits (ADL) were similar between high-elevation and sea-level otters because of the lower availability of O(2) at altitude. Our results suggest that recent disruptions to the Yellowstone Lake food web could be detrimental to otters because at this high elevation, constraints on diving may limit their ability to switch to prey in a deep-water environment.
Smart, David R; Van den Broek, Cory; Nishi, Ron; Cooper, P David; Eastman, David
2014-09-01
Tasmania's aquaculture industry produces over 40,000 tonnes of fish annually, valued at over AUD500M. Aquaculture divers perform repetitive, short-duration bounce dives in fish pens to depths up to 21 metres' sea water (msw). Past high levels of decompression illness (DCI) may have resulted from these 'yo-yo' dives. This study aimed to assess working divers, using Doppler ultrasonic bubble detection, to determine if yo-yo diving was a risk factor for DCI, determine dive profiles with acceptable risk and investigate productivity improvement. Field data were collected from working divers during bounce diving at marine farms near Hobart, Australia. Ascent rates were less than 18 m·min⁻¹, with routine safety stops (3 min at 3 msw) during the final ascent. The Kisman-Masurel method was used to grade bubbling post dive as a means of assessing decompression stress. In accordance with Defence Research and Development Canada Toronto practice, dives were rejected as excessive risk if more than 50% of scores were over Grade 2. From 2002 to 2008, Doppler data were collected from 150 bounce-dive series (55 divers, 1,110 bounces). Three series of bounce profiles, characterized by in-water times, were validated: 13-15 msw, 10 bounces inside 75 min; 16-18 msw, six bounces inside 50 min; and 19-21 msw, four bounces inside 35 min. All had median bubble grades of 0. Further evaluation validated two successive series of bounces. Bubble grades were consistent with low-stress dive profiles. Bubble grades did not correlate with the number of bounces, but did correlate with ascent rate and in-water time. These data suggest bounce diving was not a major factor causing DCI in Tasmanian aquaculture divers. Analysis of field data has improved industry productivity by increasing the permissible number of bounces, compared to earlier empirically-derived tables, without compromising safety. The recommended Tasmanian Bounce Diving Tables provide guidance for bounce diving to a depth of 21 msw, and two successive bounce dive series in a day's diving.
Garcia Párraga, Daniel; Moore, Michael; Fahlman, Andreas
2018-04-25
Hydrostatic lung compression in diving marine mammals, with collapsing alveoli blocking gas exchange at depth, has been the main theoretical basis for limiting N 2 uptake and avoiding gas emboli (GE) as they ascend. However, studies of beached and bycaught cetaceans and sea turtles imply that air-breathing marine vertebrates may, under unusual circumstances, develop GE that result in decompression sickness (DCS) symptoms. Theoretical modelling of tissue and blood gas dynamics of breath-hold divers suggests that changes in perfusion and blood flow distribution may also play a significant role. The results from the modelling work suggest that our current understanding of diving physiology in many species is poor, as the models predict blood and tissue N 2 levels that would result in severe DCS symptoms (chokes, paralysis and death) in a large fraction of natural dive profiles. In this review, we combine published results from marine mammals and turtles to propose alternative mechanisms for how marine vertebrates control gas exchange in the lung, through management of the pulmonary distribution of alveolar ventilation ([Formula: see text]) and cardiac output/lung perfusion ([Formula: see text]), varying the level of [Formula: see text] in different regions of the lung. Man-made disturbances, causing stress, could alter the [Formula: see text] mismatch level in the lung, resulting in an abnormally elevated uptake of N 2 , increasing the risk for GE. Our hypothesis provides avenues for new areas of research, offers an explanation for how sonar exposure may alter physiology causing GE and provides a new mechanism for how air-breathing marine vertebrates usually avoid the diving-related problems observed in human divers. © 2018 The Authors.
Diving through the thermal window: implications for a warming world
Campbell, Hamish A.; Dwyer, Ross G.; Gordos, Matthew; Franklin, Craig E.
2010-01-01
Population decline and a shift in the geographical distribution of some ectothermic animals have been attributed to climatic warming. Here, we show that rises in water temperature of a few degrees, while within the thermal window for locomotor performance, may be detrimental to diving behaviour in air-breathing ectotherms (turtles, crocodilians, marine iguanas, amphibians, snakes and lizards). Submergence times and internal and external body temperature were remotely recorded from freshwater crocodiles (Crocodylus johnstoni) while they free-ranged throughout their natural habitat in summer and winter. During summer, the crocodiles' mean body temperature was 5.2 ± 0.1°C higher than in winter and the largest proportion of total dive time was composed of dive durations approximately 15 min less than in winter. Diving beyond 40 min during summer required the crocodiles to exponentially increase the time they spent on the surface after the dive, presumably to clear anaerobic debt. The relationship was not as significant in winter, even though a greater proportion of dives were of a longer duration, suggesting that diving lactate threshold (DLT) was reduced in summer compared with winter. Additional evidence for a reduced DLT in summer was derived from the stronger influence body mass exerted upon dive duration, compared to winter. The results demonstrate that the higher summer body temperature increased oxygen demand during the dive, implying that thermal acclimatization of the diving metabolic rate was inadequate. If the study findings are common among air-breathing diving ectotherms, then long-term warming of the aquatic environment may be detrimental to behavioural function and survivorship. PMID:20610433
Scanning sonar of rolling porpoises during prey capture dives.
Akamatsu, T; Wang, D; Wang, K; Li, S; Dong, S
2010-01-01
Dolphins and porpoises have excellent biosonar ability, which they use for navigation, ranging and foraging. However, the role of biosonar in free-ranging small cetaceans has not been fully investigated. The biosonar behaviour and body movements of 15 free-ranging finless porpoises (Neophocaena phocaenoides) were observed using electronic tags attached to the animals. The porpoises often rotated their bodies more than 60 deg., on average, around the body axis in a dive bout. This behaviour occupied 31% of the dive duration during 186 h of effective observation time. Rolling dives were associated with extensive searching effort, and 23% of the rolling dive time was phonated, almost twice the phonation ratio of upright dives. Porpoises used short inter-click interval sonar 4.3 times more frequently during rolling dives than during upright dives. Sudden speed drops, which indicated that an individual turned around, occurred 4.5 times more frequently during rolling dives than during upright dives. Together, these data suggest that the porpoises searched extensively for targets and rolled their bodies to enlarge the search area by changing the narrow beam axis of the biosonar. Once a possible target was detected, porpoises frequently produced short-range sonar sounds. Continuous searching for prey and frequent capture trials appeared to occur during rolling dives of finless porpoises. In contrast, head movements ranging +/-2 cm, which can also change the beam axis, were regularly observed during both dives. Head movements might assist in instant assessment of the arbitrary direction by changing the beam axis rather than prey searching and pursuit.
Venous gas embolism after an open-water air dive and identical repetitive dive.
Schellart, N A M; Sterk, W
2012-01-01
Decompression tables indicate that a repetitive dive to the same depth as a first dive should be shortened to obtain the same probability of occurrence of decompression sickness (pDCS). Repetition protocols are based on small numbers, a reason for re-examination. Since venous gas embolism (VGE) and pDCS are related, one would expect a higher bubble grade (BG) of VGE after the repetitive dive without reducing bottom time. BGs were determined in 28 divers after a first and an identical repetitive air dive of 40 minutes to 20 meters of sea water. Doppler BG scores were transformed to log number of bubbles/cm2 (logB) to allow numerical analysis. With a previously published model (Model2), pDCS was calculated for the first dive and for both dives together. From pDCS, theoretical logBs were estimated with a pDCS-to-logB model constructed from literature data. However, pDCS the second dive was provided using conditional probability. This was achieved in Model2 and indirectly via tissue saturations. The combination of both models shows a significant increase of logB after the second dive, whereas the measurements showed an unexpected lower logB. These differences between measurements and model expectations are significant (p-values < 0.01). A reason for this discrepancy is uncertain. The most likely speculation would be that the divers, who were relatively old, did not perform physical activity for some days before the first dive. Our data suggest that, wisely, the first dive after a period of no exercise should be performed conservatively, particularly for older divers.
Measurement of fatigue following 18 msw dry chamber dives breathing air or enriched air nitrox.
Harris, R J D; Doolette, D J; Wilkinson, D C; Williams, D J
2003-01-01
Many divers report less fatigue following diving breathing oxygen rich N2-O2 mixtures compared with breathing air. In this double blinded, randomized controlled study 11 divers breathed either air or Enriched Air Nitrox 36% (oxygen 36%, nitrogen 64%) during an 18 msw (281 kPa(a)) dry chamber dive for a bottom time of 40 minutes. Two periods of exercise were performed during the dive. Divers were assessed before and after each dive using the Multidimensional Fatigue Inventory-20, a visual analogue scale, Digit Span Tests, Stroop Tests, and Divers Health Survey (DHS). Diving to 18m produced no measurable difference in fatigue, attention levels, ability to concentrate or DHS scores, following dives using either breathing gas.
[Scuba diving in children: Physiology, risks and recommendations].
Cilveti, R; Osona, B; Peña, J A; Moreno, L; Asensio, O
2015-12-01
The increase in recreational scuba diving in recent years, including children, involves risks and the possibility of accidents. While legislation, conditions and risks of scuba diving are well documented in adults, scientific evidence in scuba diving by children and adolescents is sparse and isolated. Furthermore, existing guidelines and recommendations for adults cannot be transferred directly to children. These circumstances have led to the Group on Techniques of the Spanish Society of Pediatric Pulmonology (SENP) to perform a literature search to review and update the knowledge about scuba diving in children. Physiological adaptations of the body are examined during the dive, as well as the anatomical and physiological characteristics of children that should be taken into account in scuba diving. The most common types of accidents and its causes, as well as the risks of scuba diving practice in children with previous diseases are discussed, along with details of the medical and psychological requirements for scuba diving to be considered in the assessment of child and adolescent. A list of recommendations for scuba diving with compressed air in children is presented by a group of experts. Copyright © 2015 Asociación Española de Pediatría. Published by Elsevier España, S.L.U. All rights reserved.
Wang, Qiong; Belhomme, Marc; Guerrero, François; Mazur, Aleksandra; Lambrechts, Kate; Theron, Michaël
2013-06-01
How underwater diving effects the function of the arterial wall and the activities of endothelial cells is the focus of recent studies on decompression sickness. Here we describe an in vitro diving system constructed to achieve real-time monitoring of cell activity during simulated dives under fluorescent microscopy and confocal microscopy. A 1-mL chamber with sapphire windows on both sides and located on the stage of an inverted microscope was built to allow in vitro diving simulation of isolated cells or arteries in which activities during diving are monitored in real-time via fluorescent microscopy and confocal microscopy. Speed of compression and decompression can range from 20 to 2000 kPa/min, allowing systemic pressure to range up to 6500 kPa. Diving temperature is controlled at 37°C. During air dive simulation oxygen partial pressure is optically monitored. Perfusion speed can range from 0.05 to 10 mL/min. The system can support physiological viability of in vitro samples for real-time monitoring of cellular activity during diving. It allows regulations of pressure, speeds of compression and decompression, temperature, gas saturation, and perfusion speed. It will be a valuable tool for hyperbaric research.
Okuyama, Junichi; Tabata, Runa; Nakajima, Kana; Arai, Nobuaki; Kobayashi, Masato; Kagawa, Shiro
2014-11-22
Air-breathing divers are assumed to have evolved to apportion their time between surface and underwater periods to maximize the benefit gained from diving activities. However, whether they change their time allocation depending on the aim of the dive is still unknown. This may be particularly crucial for 'surfacers' because they dive for various purposes in addition to foraging. In this study, we counted breath events at the surface and estimated oxygen consumption during resting, foraging and other dives in 11 green turtles (Chelonia mydas) in the wild. Breath events were counted by a head-mounted acceleration logger or direct observation based on an animal-borne video logger, and oxygen consumption was estimated by measuring overall dynamic body acceleration. Our results indicate that green turtles maximized their submerged time, following this with five to seven breaths to replenish oxygen for resting dives. However, they changed their dive tactic during foraging and other dives; they surfaced without depleting their estimated stores of oxygen, followed by only a few breaths for effective foraging and locomotion. These dichotomous surfacing tactics would be the result of behavioural modifications by turtles depending on the aim of each dive. © 2014 The Author(s) Published by the Royal Society. All rights reserved.
Okuyama, Junichi; Tabata, Runa; Nakajima, Kana; Arai, Nobuaki; Kobayashi, Masato; Kagawa, Shiro
2014-01-01
Air-breathing divers are assumed to have evolved to apportion their time between surface and underwater periods to maximize the benefit gained from diving activities. However, whether they change their time allocation depending on the aim of the dive is still unknown. This may be particularly crucial for ‘surfacers’ because they dive for various purposes in addition to foraging. In this study, we counted breath events at the surface and estimated oxygen consumption during resting, foraging and other dives in 11 green turtles (Chelonia mydas) in the wild. Breath events were counted by a head-mounted acceleration logger or direct observation based on an animal-borne video logger, and oxygen consumption was estimated by measuring overall dynamic body acceleration. Our results indicate that green turtles maximized their submerged time, following this with five to seven breaths to replenish oxygen for resting dives. However, they changed their dive tactic during foraging and other dives; they surfaced without depleting their estimated stores of oxygen, followed by only a few breaths for effective foraging and locomotion. These dichotomous surfacing tactics would be the result of behavioural modifications by turtles depending on the aim of each dive. PMID:25297856
A case-control study evaluating relative risk factors for decompression sickness: a research report.
Suzuki, Naoko; Yagishita, Kazuyosi; Togawa, Seiichiro; Okazaki, Fumihiro; Shibayama, Masaharu; Yamamoto, Kazuo; Mano, Yoshihiro
2014-01-01
Factors contributing to the pathogenesis of decompression sickness (DCS) in divers have been described in many studies. However, relative importance of these factors has not been reported. In this case-control study, we compared the diving profiles of divers experiencing DCS with those of a control group. The DCS group comprised 35 recreational scuba divers who were diagnosed by physicians as having DCS. The control group consisted of 324 apparently healthy recreational divers. All divers conducted their dives from 2009 to 2011. The questionnaire consisted of 33 items about an individual's diving profile, physical condition and activities before, during and just after the dive. To simplify dive parameters, the dive site was limited to Izu Osezaki. Odds ratios and multiple logistic regression were used for the analysis. Odds ratios revealed several items as dive and health factors associated with DCS. The major items were as follows: shortness of breath after heavy exercise during the dive (OR = 12.12), dehydration (OR = 10.63), and maximum dive depth > 30 msw (OR = 7.18). Results of logistic regression were similar to those by odds ratio analysis. We assessed the relative weights of the surveyed dive and health factors associated with DCS. Because results of several factors conflict with previous studies, future studies are needed.
NASA Astrophysics Data System (ADS)
Kerby, T.; Smith, J. R.; Shackelford, R.; Wiltshire, J. C.; Malahoff, A.
2005-12-01
The Hawaii Undersea Research Laboratory (HURL) recently completed an internationally partnered 5-month, 14,500 nautical mile multiple leg expedition to the South Pacific that included 21 study sites in the waters of American Samoa, New Zealand, Tonga, and the U.S. Line Islands to commemorate its 25th anniversary of supporting deep submergence science in the Pacific Ocean. During this voyage, HURL successfully operated its two human occupied vehicles ( Pisces IV and Pisces V) each capable of diving to 2000 m from their support ship, the R/V Ka'imikai-o-Kanaloa ( KoK). In addition, a remotely operated vehicle ( RCV-150) with a nearly 1000-m depth limit was utilized alternately with the Pisces HOV's. The size and organized placement of these vehicles on the compact but efficiently run KoK (70-m length, 2000-tons displacement, 14 crew) allowed for deployment of a CTD rosette system and recovery of instrument package moorings during the same cruise leg. The Pisces submersibles are 20-ft long, 13-ton, 3-person vehicles with 7-10 hours duration, up to 350-lb payload capacities, and three forward looking viewports. The small size of the Pisces' relative to much larger deeper diving HOV's increases their agility, thus allowing maneuvering into more difficult sampling site terrain. The smaller package also facilitates rapid launch (8 min avg, stdev=1) and recovery (12 min avg, stdev=2) in heavier seas (up to sea state 5), as routinely experienced in the South Pacific during the austral winter. In addition to the enhanced safety aspect of having two compatible submersibles aboard, scientific efficiency has benefited by allowing the rotation of vehicles on extended deployments prior to battery servicing, thus maintaining an overall dive time average of 7.1 hr (stdev=1.52) for an average dive depth of 891 m (stdev=431) in 2005. Having the two fully operational submersibles also provides a contingency for equipment malfunction while on site that saved 7 dive days in 2005 alone. The final dive count resulted in 61 out of 56 scheduled Pisces science dives completed; made possible by careful planning and on-site cruise management and aided by our can-do crew. The RCV-150 was also used on 17 dives when depth and sea conditions allowed, typically in a scouting mode for potential HOV dives to increase the efficiency of the entire operation. The total package described here was made available for less than $30K per dive day in 2005. Standard equipment on the Pisces pair and the RCV-150 are listed at the URL below. In addition, science team equipment was interfaced including the NOAA VENTS fluid sampler and gas tight bottles, Imagenex pencil beam bathymetric sonar system, and the WHOI TowCam external digital still camera. Tracking and navigation accuracy to within 20 m at 1500-1800 m depth was provided by a TrackLink 5000HA ultra-short baseline system interfaced with a MAHRS motion sensor to provide pitch and roll correction. Additional vehicle details, operational results, and statistics for the 2005 South Pacific expedition will be presented.
Venous gas emboli are involved in post-dive macro, but not microvascular dysfunction.
Lambrechts, Kate; Balestra, Costantino; Theron, Michaël; Henckes, Anne; Galinat, Hubert; Mignant, Fanny; Belhomme, Marc; Pontier, Jean-Michel; Guerrero, François
2017-02-01
Previous studies have shown vascular dysfunction of main conductance arteries and microvessels after diving. We aim to evaluate the impact of bubble formation on vascular function and haemostasis. To achieve this, we used a vibration preconditioning to influence bubble levels without changing any other parameters linked to the dive. Twentty-six divers were randomly assigned to one of three groups: (1) the "vibrations-dive" group (VD; n = 9) was exposed to a whole-body vibration session 30 min prior the dive; (2) the "diving" group (D; n = 9) served as a control for the effect of the diving protocol; (3) The "vibration" protocol (V; n = 8) allowed us to assess the effect of vibrations without diving. Macro- and microvascular function was assessed for each subject before and after the dive, subsequently. Bubble grades were monitored with Doppler according to the Spencer grading system. Blood was taken before and after the protocol to assess any change of platelets or endothelial function. Bubble formation was lower in the VD than the diving group. The other measured parameters remained unchanged after the "vibration" protocol alone. Diving alone induced macrovascular dysfunction, and increased PMP and thrombin generation. Those parameters were no longer changed in the VD group. Conversely, a microvascular dysfunction persists despite a significant decrease of circulating bubbles. Finally, the results of this study suggest that macro- but not microvascular impairment results at least partly from bubbles, possibly related to platelet activation and generation of pro-coagulant microparticles.
Goldman, Saul; Solano-Altamirano, J M
2015-04-01
We solved the Laplace equation for the radius of an arterial gas embolism (AGE), during and after breath-hold diving. We used a simple three-region diffusion model for the AGE, and applied our results to two types of breath-hold dives: single, very deep competitive-level dives and repetitive shallower breath-hold dives similar to those carried out by indigenous commercial pearl divers in the South Pacific. Because of the effect of surface tension, AGEs tend to dissolve in arterial blood when arteries remote from supersaturated tissue. However if, before fully dissolving, they reach the capillary beds that perfuse the brain and the inner ear, they may become inflated with inert gas that is transferred into them from these contiguous temporarily supersaturated tissues. By using simple kinetic models of cerebral and inner ear tissue, the nitrogen tissue partial pressures during and after the dive(s) were determined. These were used to theoretically calculate AGE growth and dissolution curves for AGEs lodged in capillaries of the brain and inner ear. From these curves it was found that both cerebral and inner ear decompression sickness are expected to occur occasionally in single competitive-level dives. It was also determined from these curves that for the commercial repetitive dives considered, the duration of the surface interval (the time interval separating individual repetitive dives from one another) was a key determinant, as to whether inner ear and/or cerebral decompression sickness arose. Our predictions both for single competitive-level and repetitive commercial breath-hold diving were consistent with what is known about the incidence of cerebral and inner ear decompression sickness in these forms of diving. Copyright © 2015 Elsevier Inc. All rights reserved.
Teilmann, Jonas; Hermannsen, Line; Galatius, Anders; Sveegaard, Signe; Balle, Jeppe Dalgaard; Dietz, Rune; Nabe-Nielsen, Jacob
2018-01-01
Knowledge about the impact of anthropogenic disturbances on the behavioural responses of cetaceans is constrained by lack of data on fine-scale movements of individuals. We equipped five free-ranging harbour porpoises (Phocoena phocoena) with high-resolution location and dive loggers and exposed them to a single 10 inch3 underwater airgun producing high-intensity noise pulses (2–3 s intervals) for 1 min. All five porpoises responded to capture and tagging with longer, faster and more directed movements as well as with shorter, shallower, less wiggly dives immediately after release, with natural behaviour resumed in less than or equal to 24 h. When we exposed porpoises to airgun pulses at ranges of 420–690 m with noise level estimates of 135–147 dB re 1 µPa2s (sound exposure level), one individual displayed rapid and directed movements away from the exposure site and two individuals used shorter and shallower dives compared to natural behaviour immediately after exposure. Noise-induced movement typically lasted for less than or equal to 8 h with an additional 24 h recovery period until natural behaviour was resumed. The remaining individuals did not show any quantifiable responses to the noise exposure. Changes in natural behaviour following anthropogenic disturbances may reduce feeding opportunities, and evaluating potential population-level consequences should be a priority research area. PMID:29410789
Durban, John W.; Claridge, Diane E.; Dunn, Charlotte A.; Fearnbach, Holly; Parsons, Kim M.; Andrews, Russel D.; Ballance, Lisa T.
2017-01-01
Dive capacity among toothed whales (suborder: Odontoceti) has been shown to generally increase with body mass in a relationship closely linked to the allometric scaling of metabolic rates. However, two odontocete species tagged in this study, the Blainville’s beaked whale Mesoplodon densirostris and the Cuvier’s beaked whale Ziphius cavirostris, confounded expectations of a simple allometric relationship, with exceptionally long (mean: 46.1 min & 65.4 min) and deep dives (mean: 1129 m & 1179 m), and comparatively small body masses (med.: 842.9 kg & 1556.7 kg). These two species also exhibited exceptionally long recovery periods between successive deep dives, or inter-deep-dive intervals (M. densirostris: med. 62 min; Z. cavirostris: med. 68 min). We examined competing hypotheses to explain observed patterns of vertical habitat use based on body mass, oxygen binding protein concentrations, and inter-deep-dive intervals in an assemblage of five sympatric toothed whales species in the Bahamas. Hypotheses were evaluated using dive data from satellite tags attached to the two beaked whales (M. densirostris, n = 12; Z. cavirostris, n = 7), as well as melon-headed whales Peponocephala electra (n = 13), short-finned pilot whales Globicephala macrorhynchus (n = 15), and sperm whales Physeter macrocephalus (n = 27). Body mass and myoglobin concentration together explained only 36% of the variance in maximum dive durations. The inclusion of inter-deep-dive intervals, substantially improved model fits (R2 = 0.92). This finding supported a hypothesis that beaked whales extend foraging dives by exceeding aerobic dive limits, with the extension of inter-deep-dive intervals corresponding to metabolism of accumulated lactic acid. This inference points to intriguing tradeoffs between body size, access to prey in different depth strata, and time allocation within dive cycles. These tradeoffs and resulting differences in habitat use have important implications for spatial distribution patterns, and relative vulnerabilities to anthropogenic impacts. PMID:29020021
Joyce, Trevor W; Durban, John W; Claridge, Diane E; Dunn, Charlotte A; Fearnbach, Holly; Parsons, Kim M; Andrews, Russel D; Ballance, Lisa T
2017-01-01
Dive capacity among toothed whales (suborder: Odontoceti) has been shown to generally increase with body mass in a relationship closely linked to the allometric scaling of metabolic rates. However, two odontocete species tagged in this study, the Blainville's beaked whale Mesoplodon densirostris and the Cuvier's beaked whale Ziphius cavirostris, confounded expectations of a simple allometric relationship, with exceptionally long (mean: 46.1 min & 65.4 min) and deep dives (mean: 1129 m & 1179 m), and comparatively small body masses (med.: 842.9 kg & 1556.7 kg). These two species also exhibited exceptionally long recovery periods between successive deep dives, or inter-deep-dive intervals (M. densirostris: med. 62 min; Z. cavirostris: med. 68 min). We examined competing hypotheses to explain observed patterns of vertical habitat use based on body mass, oxygen binding protein concentrations, and inter-deep-dive intervals in an assemblage of five sympatric toothed whales species in the Bahamas. Hypotheses were evaluated using dive data from satellite tags attached to the two beaked whales (M. densirostris, n = 12; Z. cavirostris, n = 7), as well as melon-headed whales Peponocephala electra (n = 13), short-finned pilot whales Globicephala macrorhynchus (n = 15), and sperm whales Physeter macrocephalus (n = 27). Body mass and myoglobin concentration together explained only 36% of the variance in maximum dive durations. The inclusion of inter-deep-dive intervals, substantially improved model fits (R2 = 0.92). This finding supported a hypothesis that beaked whales extend foraging dives by exceeding aerobic dive limits, with the extension of inter-deep-dive intervals corresponding to metabolism of accumulated lactic acid. This inference points to intriguing tradeoffs between body size, access to prey in different depth strata, and time allocation within dive cycles. These tradeoffs and resulting differences in habitat use have important implications for spatial distribution patterns, and relative vulnerabilities to anthropogenic impacts.
Characteristics of diving in radio-marked Xantus's Murrelets
Hamilton, C.D.; Golightly, R.T.; Takekawa, John Y.
2005-01-01
We monitored diving activity of radio-marked Xantus's Murrelets Synthliboramphus hypoleucus near Anacapa Island, California, during the breeding season. Thirteen radio-marked murrelets were remotely monitored from Anacapa Island with a handheld antenna and radio receiver for 29 hours in three sample periods in April and May 2003. Mean dive durations in the sample periods were 18 s ?? 2 s, 28 s ?? 2 s, and 24 s ?? 4 s, suggesting that dives were less than 21 m from the surface. Dive duration and subsequent time on the surface differed between the sample periods. Dive duration and subsequent time on the surface were not correlated in observations stratified by individual bird or by sample period. Further, dive duration and subsequent time on the surface were not correlated within foraging bouts. Dive characteristics measured near Anacapa Island suggested that Xantus's Murrelets have the ability to capture prey found at varying depths, but will feed on prey that is most available near the surface of the water.
Fossette, Sabrina; Gaspar, Philippe; Handrich, Yves; Le Maho, Yvon; Georges, Jean-Yves
2008-03-01
1. Investigating the foraging patterns of free-ranging species is essential to estimate energy/time budgets for assessing their real reproductive strategy. Leatherback turtles Dermochelys coriacea (Vandelli 1761), commonly considered as capital breeders, have been reported recently to prospect actively during the breeding season in French Guiana, Atlantic Ocean. In this study we investigate the possibility of this active behaviour being associated with foraging, by studying concurrently diving and beak movement patterns in gravid females equipped with IMASEN (Inter-MAndibular Angle SENsor). 2. Four turtles provided data for periods varying from 7.3 to 56.1 h while exhibiting continuous short and shallow benthic dives. Beak movement ('b-m') events occurred in 34% of the dives, on average 1.8 +/- 1.4 times per dive. These b-m events lasted between 1.5 and 20 s and occurred as isolated or grouped (two to five consecutive beak movements) events in 96.0 +/- 4.0% of the recorded cases, and to a lesser extent in series (> five consecutive beak movements). 3. Most b-m events occurred during wiggles at the bottom of U- and W-shaped dives and at the beginning and end of the bottom phase of the dives. W-shaped dives were associated most frequently with beak movements (65% of such dives) and in particular with grouped beak movements. 4. Previous studies proposed wiggles to be indicator of predatory activity, U- and W-shaped dives being putative foraging dives. Beak movements recorded in leatherbacks during the first hours of their internesting interval in French Guiana may be related to feeding attempts. 5. In French Guiana, leatherbacks show different mouth-opening patterns for different dive patterns, suggesting that they forage opportunistically on occasional prey, with up to 17% of the dives appearing to be successful feeding dives. 6. This study highlights the contrasted strategies adopted by gravid leatherbacks nesting on the Pacific coasts of Costa Rica, in the deep-water Caribbean Sea and in the French Guianan shallow continental shelf, and may be related to different local prey accessibility among sites. Our results may help to explain recently reported site-specific individual body size and population dynamics.
Otorhinolaryngologic disorders and diving accidents: an analysis of 306 divers.
Klingmann, Christoph; Praetorius, Mark; Baumann, Ingo; Plinkert, Peter K
2007-10-01
Diving is a very popular leisure activity with an increasing number of participants. As more than 80% of the diving related problems involve the head and neck region, every otorhinolaryngologist should be familiar with diving medical standards. We here present an analysis of more than 300 patients we have treated in the past four years. Between January 2002 and October 2005, 306 patients presented in our department with otorhinological disorders after diving, or after diving accidents. We collected the following data: name, sex, age, date of treatment, date of accident, diagnosis, special aspects of the diagnosis, number of dives, diving certification, whether and which surgery had been performed, history of acute diving accidents or follow up treatment, assessment of fitness to dive and special remarks. The study setting was a retrospective cohort study. The distribution of the disorders was as follows: 24 divers (8%) with external ear disorders, 140 divers (46%) with middle ear disorders, 56 divers (18%) with inner ear disorders, 53 divers (17%) with disorders of the nose and sinuses, 24 divers (8%) with decompression illness (DCI) and 9 divers (3%) who complained of various symptoms. Only 18% of the divers presented with acute disorders. The most common disorder (24%) was Eustachian tube dysfunction. Female divers were significantly more often affected. Chronic sinusitis was found to be associated with a significantly higher number of performed dives. Conservative treatment failed in 30% of the patients but sinus surgery relieved symptoms in all patients of this group. The middle ear is the main problem area for divers. Middle ear ventilation problems due to Eustachian tube dysfunction can be treated conservatively with excellent results whereas pathology of the tympanic membrane and ossicular chain often require surgery. More than four out of five patients visited our department to re-establish their fitness to dive. Although the treatment of acute diving-related disorders is an important field for the treatment of divers, the main need of divers seems to be assessment and recovery of their fitness to dive.
Are recreational SCUBA divers with asthma at increased risk?
Ustrup, Amalie S; Ulrik, Charlotte S
2017-10-01
Asthma has traditionally been regarded as a contraindication to self-contained underwater breathing apparatus (SCUBA) diving, although large numbers of patients with asthma dive. The aim of the review is to provide an update on current knowledge on potential disease-related hazards in SCUBA divers with asthma. Systematic literature review based on the Preferred Reporting Items for Systematic reviews and Meta-Analyses (PRISMA) guidelines. Seven studies met the criteria for inclusion in the review (comprising a total of 560 subjects). Five studies reported an increased risk for developing diving-related injuries in divers with asthma, based on case reports (n = 1), case history combined with objective assessment (n = 1), and dives and/or simulated dives (n = 3). The remaining studies (n = 2) were based on self-reported diving habits in divers suffering from asthma, obtained from anonymous questionnaires in diving magazines, reported no diving-related injuries among respondents. Due to limited evidence it is difficult to draw valid conclusions, but there are indications that recreational divers with asthma may be at increased risk for diving-related injuries compared to non-asthmatic divers. However, it is of at most importance to obtain further evidence from large-scale, well-designed studies.
USDA-ARS?s Scientific Manuscript database
Cattle are a major reservoir for Shiga toxin-producing Escherichia coli O157 (STEC O157) and harbor multiple genetic subtypes that do not all associate with human disease. STEC O157 evolved from an E. coli O55:H7 progenitor, however, a lack of genome sequence has hindered investigations on the dive...
Huchim-Lara, Oswaldo; Salas, Silvia; Chin, Walter; Montero, Jorge; Fraga, Julia
2015-01-01
An average of 209 cases of decompression sickness (DCS) have been reported every year among artisanal fishermen. divers of the Yucatan Peninsula, Mexico. DCS is a major problem among fishermen divers worldwide. This paper explores how diving behavior and fishing techniques among fishermen relate to the probability of experiencing DCS (Pdcs). Fieldwork was conducted in two communities during the 2012-2013 fishing season. Fishermen were classified into three groups (two per group) according to their fishing performance and followed during their journeys. Dive profiles were recorded using Sensus Ultra dive recorders (Reefet Inc.). Surveys were used to record fishing yields from cooperative and individual fishermen along with fishing techniques and dive behavior. 120 dives were recorded. Fishermen averaged three dives/day, with an average depth of 47 ± 2 feet of sea water (fsw) and an average total bottom time (TBT) of 95 ± 11 minutes. 24% of dives exceeded the 2008 U.S. Navy no-decompression limit. The average ascent rate was 20 fsw/minute, and 5% of those exceeded 40 fsw/minute. Inadequate decompression was observed in all fishermen. Fishermen are diving outside the safety limits of both military and recreational standards. Fishing techniques and dive behavior were important factors in Pdcs. Fishermen were reluctant to seek treatment, and symptoms were relieved with analgesics.
Novel locomotor muscle design in extreme deep-diving whales.
Velten, B P; Dillaman, R M; Kinsey, S T; McLellan, W A; Pabst, D A
2013-05-15
Most marine mammals are hypothesized to routinely dive within their aerobic dive limit (ADL). Mammals that regularly perform deep, long-duration dives have locomotor muscles with elevated myoglobin concentrations that are composed of predominantly large, slow-twitch (Type I) fibers with low mitochondrial volume densities (V(mt)). These features contribute to extending ADL by increasing oxygen stores and decreasing metabolic rate. Recent tagging studies, however, have challenged the view that two groups of extreme deep-diving cetaceans dive within their ADLs. Beaked whales (including Ziphius cavirostris and Mesoplodon densirostris) routinely perform the deepest and longest average dives of any air-breathing vertebrate, and short-finned pilot whales (Globicephala macrorhynchus) perform high-speed sprints at depth. We investigated the locomotor muscle morphology and estimated total body oxygen stores of several species within these two groups of cetaceans to determine whether they (1) shared muscle design features with other deep divers and (2) performed dives within their calculated ADLs. Muscle of both cetaceans displayed high myoglobin concentrations and large fibers, as predicted, but novel fiber profiles for diving mammals. Beaked whales possessed a sprinter's fiber-type profile, composed of ~80% fast-twitch (Type II) fibers with low V(mt). Approximately one-third of the muscle fibers of short-finned pilot whales were slow-twitch, oxidative, glycolytic fibers, a rare fiber type for any mammal. The muscle morphology of beaked whales likely decreases the energetic cost of diving, while that of short-finned pilot whales supports high activity events. Calculated ADLs indicate that, at low metabolic rates, both beaked and short-finned pilot whales carry sufficient onboard oxygen to aerobically support their dives.
NASA Astrophysics Data System (ADS)
Blumenthal, J. M.; Austin, T. J.; Bothwell, J. B.; Broderick, A. C.; Ebanks-Petrie, G.; Olynik, J. R.; Orr, M. F.; Solomon, J. L.; Witt, M. J.; Godley, B. J.
2009-03-01
As historically abundant spongivores, hawksbill turtles Eretmochelys imbricata likely played a key ecological role on coral reefs. However, coral reefs are now experiencing global declines and many hawksbill populations are critically reduced. For endangered species, tracking movement has been recognized as fundamental to management. Since movements in marine vertebrates encompass three dimensions, evaluation of diving behavior and range is required to characterize marine turtle habitat. In this study, habitat use of hawksbill turtles on a Caribbean coral reef was elucidated by quantifying diel depth utilization and movements in relation to the boundaries of marine protected areas. Time depth recorders (TDRs) and ultrasonic tags were deployed on 21 Cayman Islands hawksbills, ranging in size from 26.4 to 58.4 cm straight carapace length. Study animals displayed pronounced diel patterns of diurnal activity and nocturnal resting, where diurnal dives were significantly shorter, deeper, and more active. Mean diurnal dive depth (±SD) was 8 ± 5 m, range 2-20 m, mean nocturnal dive depth was 5 ± 5 m, range 1-14 m, and maximum diurnal dive depth was 43 ± 27 m, range 7-91 m. Larger individuals performed significantly longer dives. Body mass was significantly correlated with mean dive depth for nocturnal but not diurnal dives. However, maximum diurnal dive depth was significantly correlated with body mass, suggesting partitioning of vertical habitat by size. Thus, variable dive capacity may reduce intraspecific competition and provide resistance to degradation in shallow habitats. Larger hawksbills may also represent important predators on deep reefs, creating a broad ecological footprint over a range of depths.
Residual oxygen time model for oxygen partial pressure near 130 kPa (1.3 atm).
Shykoff, Barbara E
2015-01-01
A two-part residual oxygen time model predicts the probability of detectible pulmonary oxygen toxicity P(P[O2tox]) after dives with oxygen partial pressure (PO2) approximately 130 kPa, and provides a tool to plan dive series with selected risk of P[O2tox]. Data suggest that pulmonary oxygen injury at this PO2 is additive between dives. Recovery begins after a delay and continues during any following dive. A logistic relation expresses P(P[O2tox]) as a function of dive duration (T(dur)) [hours]: P(P[O2tox]) = 100/[1+exp (3.586-0.49 x T(dur))] This expression maps T(dur) to P(P[O2tox]) or, in the linear mid-portion of the curve, P(P[O2tox]) usefully to T(dur). For multiple dives or during recovery, it maps to an equivalent dive duration, T(eq). T(eq) was found after second dives of duration T(dur 2). Residual time from the first dive t(r) = T(eq) - T(dur2). With known t(r), t and T(dur) a recovery model was fitted. t(r) = T(dur) x exp [-k x((t-5)/T(dur)2], where t = t - 5 hours, k = 0.149 for resting, and 0.047 for exercising divers, and t represents time after surfacing. The fits were assessed for 1,352 man-dives. Standard deviations of the residuals were 8.5% and 18.3% probability for resting or exercise dives, respectively.
46 CFR 197.314 - First aid and treatment equipment.
Code of Federal Regulations, 2013 CFR
2013-10-01
... treatment equipment. (a) Each dive location must have— (1) A medical kit approved by a physician that... communications system to obtain emergency assistance except when the vessel or facility ship-to-shore, two-way communications system is readily available. (c) Each dive location supporting mixed-gas dives, dives deeper than...
46 CFR 197.314 - First aid and treatment equipment.
Code of Federal Regulations, 2014 CFR
2014-10-01
... treatment equipment. (a) Each dive location must have— (1) A medical kit approved by a physician that... communications system to obtain emergency assistance except when the vessel or facility ship-to-shore, two-way communications system is readily available. (c) Each dive location supporting mixed-gas dives, dives deeper than...
46 CFR 197.314 - First aid and treatment equipment.
Code of Federal Regulations, 2011 CFR
2011-10-01
... treatment equipment. (a) Each dive location must have— (1) A medical kit approved by a physician that... communications system to obtain emergency assistance except when the vessel or facility ship-to-shore, two-way communications system is readily available. (c) Each dive location supporting mixed-gas dives, dives deeper than...
46 CFR 197.410 - Dive procedures.
Code of Federal Regulations, 2010 CFR
2010-10-01
... location decompression chamber for at least one hour after the completion of a dive, decompression, or... corrective action taken, if necessary, to reduce the probability of recurrence. (b) The diving supervisor shall ensure that the working interval of a dive is terminated when he so directs or when— (1) A diver...
Curé, Charlotte; Antunes, Ricardo; Alves, Ana Catarina; Visser, Fleur; Kvadsheim, Petter H.; Miller, Patrick J. O.
2013-01-01
Interactions between individuals of different cetacean species are often observed in the wild. Killer whales (Orcinus orca) can be potential predators of many other cetaceans, and the interception of their vocalizations by unintended cetacean receivers may trigger anti-predator behavior that could mediate predator-prey interactions. We explored the anti-predator behaviour of five typically-solitary male sperm whales (Physeter macrocephalus) in the Norwegian Sea by playing sounds of mammal-feeding killer whales and monitoring behavioural responses using multi-sensor tags. Our results suggest that, rather than taking advantage of their large aerobic capacities to dive away from the perceived predator, sperm whales responded to killer whale playbacks by interrupting their foraging or resting dives and returning to the surface, changing their vocal production, and initiating a surprising degree of social behaviour in these mostly solitary animals. Thus, the interception of predator vocalizations by male sperm whales disrupted functional behaviours and mediated previously unrecognized anti-predator responses. PMID:23545484
Curé, Charlotte; Antunes, Ricardo; Alves, Ana Catarina; Visser, Fleur; Kvadsheim, Petter H; Miller, Patrick J O
2013-01-01
Interactions between individuals of different cetacean species are often observed in the wild. Killer whales (Orcinus orca) can be potential predators of many other cetaceans, and the interception of their vocalizations by unintended cetacean receivers may trigger anti-predator behavior that could mediate predator-prey interactions. We explored the anti-predator behaviour of five typically-solitary male sperm whales (Physeter macrocephalus) in the Norwegian Sea by playing sounds of mammal-feeding killer whales and monitoring behavioural responses using multi-sensor tags. Our results suggest that, rather than taking advantage of their large aerobic capacities to dive away from the perceived predator, sperm whales responded to killer whale playbacks by interrupting their foraging or resting dives and returning to the surface, changing their vocal production, and initiating a surprising degree of social behaviour in these mostly solitary animals. Thus, the interception of predator vocalizations by male sperm whales disrupted functional behaviours and mediated previously unrecognized anti-predator responses.
77 FR 19645 - Marine Mammals; File No. 16111
Federal Register 2010, 2011, 2012, 2013, 2014
2012-04-02
... abundance, distribution, population structure, habitat use, social structure, movement patterns, diving behavior, and diet. The proposed project would also assess the impact of human activities such as ship strikes, noise exposure, contaminants, and fishery interactions on marine mammals. Focal species are blue...
Diving-related visual loss in the setting of angioid streaks: report of two cases.
Angulo Bocco, Maria I; Spielberg, Leigh; Coppens, Greet; Catherine, Janet; Verougstraete, Claire; Leys, Anita M
2012-01-01
The purpose of this study was to report diving-related visual loss in the setting of angioid streaks. Observational case reports of two patients with angioid streaks suffering sudden visual loss immediately after diving. Two young adult male patients presented with visual loss after diving headfirst. Funduscopy revealed angioid streaks, peau d'orange, subretinal hemorrhages, and ruptures of Bruch membrane. Choroidal neovascularization developed during follow-up. Both patients had an otherwise uneventful personal and familial medical history. In patients with angioid streaks, diving headfirst can lead to subretinal hemorrhages and traumatic ruptures in Bruch membrane and increase the risk of maculopathy. Ophthalmologists should caution patients with angioid streaks against diving headfirst.
Seals map bathymetry of the Antarctic continental shelf
NASA Astrophysics Data System (ADS)
Padman, Laurie; Costa, Daniel P.; Bolmer, S. Thompson; Goebel, Michael E.; Huckstadt, Luis A.; Jenkins, Adrian; McDonald, Birgitte I.; Shoosmith, Deborah R.
2010-11-01
We demonstrate the first use of marine mammal dive-depth data to improve maps of bathymetry in poorly sampled regions of the continental shelf. A group of 57 instrumented elephant seals made on the order of 2 × 105 dives over and near the continental shelf on the western side of the Antarctic Peninsula during five seasons, 2005-2009. Maximum dive depth exceeded 2000 m. For dives made near existing ship tracks with measured water depths H<700 m, ˜30% of dive depths were to the seabed, consistent with expected benthic foraging behavior. By identifying the deepest of multiple dives within small areas as a dive to the seabed, we have developed a map of seal-derived bathymetry. Our map fills in several regions for which trackline data are sparse, significantly improving delineation of troughs crossing the continental shelf of the southern Bellingshausen Sea.
46 CFR 197.434 - Surface-supplied mixed-gas diving.
Code of Federal Regulations, 2012 CFR
2012-10-01
... 46 Shipping 7 2012-10-01 2012-10-01 false Surface-supplied mixed-gas diving. 197.434 Section 197....434 Surface-supplied mixed-gas diving. The diving supervisor shall insure that— (a) When mixed-gas... supply meeting the requirements of § 197.340; and (k) The surface-supplied mixed-gas diver has the...
46 CFR 197.434 - Surface-supplied mixed-gas diving.
Code of Federal Regulations, 2011 CFR
2011-10-01
... 46 Shipping 7 2011-10-01 2011-10-01 false Surface-supplied mixed-gas diving. 197.434 Section 197....434 Surface-supplied mixed-gas diving. The diving supervisor shall insure that— (a) When mixed-gas... supply meeting the requirements of § 197.340; and (k) The surface-supplied mixed-gas diver has the...
46 CFR 197.434 - Surface-supplied mixed-gas diving.
Code of Federal Regulations, 2014 CFR
2014-10-01
... 46 Shipping 7 2014-10-01 2014-10-01 false Surface-supplied mixed-gas diving. 197.434 Section 197....434 Surface-supplied mixed-gas diving. The diving supervisor shall insure that— (a) When mixed-gas... supply meeting the requirements of § 197.340; and (k) The surface-supplied mixed-gas diver has the...
46 CFR 197.434 - Surface-supplied mixed-gas diving.
Code of Federal Regulations, 2013 CFR
2013-10-01
... 46 Shipping 7 2013-10-01 2013-10-01 false Surface-supplied mixed-gas diving. 197.434 Section 197....434 Surface-supplied mixed-gas diving. The diving supervisor shall insure that— (a) When mixed-gas... supply meeting the requirements of § 197.340; and (k) The surface-supplied mixed-gas diver has the...
Federal Register 2010, 2011, 2012, 2013, 2014
2011-11-01
...] Standard on Commercial Diving Operations; Extension of the Office of Management and Budget's (OMB) Approval... Commercial Diving Operations Standard (29 CFR part 1910, subpart T). DATES: Comments must be submitted... existing Standard on Commercial Diving Operations (29 CFR part 1910, Subpart [[Page 67481
Evidence for behavioural thermoregulation by the world's largest fish
Thums, Michele; Meekan, Mark; Stevens, John; Wilson, Steven; Polovina, Jeff
2013-01-01
Many fishes make frequent ascents to surface waters and often show prolonged surface swimming following descents to deep water. This affinity for the surface is thought to be related to the recovery of body heat lost at depth. We tested this hypothesis using data from time–depth recorders deployed on four whale sharks (Rhincodon typus). We summarized vertical movements into bouts of dives and classified these into three main types, using cluster analysis. In addition to day and night ‘bounce’ dives where sharks rapidly descended and ascended, we found a third type: single deep (mean: 340 m), long (mean: 169 min) dives, occurring in daytime with extremely long post-dive surface durations (mean: 146 min). Only sharks that were not constrained by shallow bathymetry performed these dives. We found a negative relationship between the mean surface duration of dives in the bout and the mean minimum temperature of dives in the bout that is consistent with the hypothesis that thermoregulation was a major factor driving use of the surface. The relationship broke down when sharks were diving in mean minimum temperatures around 25°C, suggesting that warmer waters did not incur a large metabolic cost for diving and that other factors may also influence surface use. PMID:23075547
Functional properties of myoglobins from five whale species with different diving capacities.
Helbo, Signe; Fago, Angela
2012-10-01
Whales show an exceptionally wide range of diving capabilities and many express high amounts of the O(2) carrier protein myoglobin (Mb) in their muscle tissues, which increases their aerobic diving capacity. Although previous studies have mainly focused on the muscle Mb concentration and O(2) carrying capacity as markers of diving behavior in whales, it still remains unexplored whether whale Mbs differ in their O(2) affinities and nitrite reductase and peroxidase enzymatic activities, all functions that could contribute to differences in diving capacities. In this study, we have measured the functional properties of purified Mbs from five toothed whales and two baleen whales and have examined their correlation with average dive duration. Results showed that some variation in functional properties exists among whale Mbs, with toothed whale Mbs having higher O(2) affinities and nitrite reductase activities (similar to those of horse Mb) compared with baleen whale Mbs. However, these differences did not correlate with average dive duration. Instead, a significant correlation was found between whale Mb concentration and average duration and depth of dives, and between O(2) affinity and nitrite reductase activity when including horse Mb. Despite the fact that the functional properties showed little species-specific differences in vitro, they may still contribute to enhancing diving capacity as a result of the increased muscle Mb concentration found in extreme divers. In conclusion, Mb concentration rather than specific functional reactivities may support whale diving performance.
The use of drugs by UK recreational divers: illicit drugs.
Dowse, Marguerite St Leger; Shaw, Steve; Cridge, Christine; Smerdon, Gary
2011-03-01
Anecdotal observations suggest the use of illicit drugs takes place amongst recreational divers but, to date, there has been little open debate within the diving community concerning possible prevalence. This study investigated the prevalence and type of illicit drugs used by recreational divers in the United Kingdom (UK). Anonymous questionnaires were circulated via UK dive clubs, dive schools, dive shows and conferences. Questions incorporated diver and diving demographics and general health, which included anxiety, depression and panic attacks, alcohol use, smoking and illicit drug use since learning to dive and closest time to a dive. Questions pertaining to over-the-counter and prescription drug use were also asked. 479 divers responded (66% males and 34% females) in the age range 16 to 59 years. Of the respondents, 22% had used one or more illicit drug since learning to dive, reporting benzodiazepines, amphetamines, cocaine, ecstasy, LSD, cannabis, heroin, and 'magic mushrooms'. Illicit drugs had been used by 3.5% of respondents in the last 12 months, and 3% in the last month. Cannabis, cocaine and ecstasy use was reported within 6 hours of a dive. Logistic regression confirmed a relationship between illicit drug use and depression (P = 0.014), and also between illicit drug use and anxiety (P = 0.024). These data support anecdotal reports that recreational divers use a range of illicit drugs. The significant relationship between illicit drug use and depression and anxiety supports the literature in non-diving populations.
Sayer, Martin D J; Azzopardi, Elaine; Sieber, Arne
2014-12-01
Dive computers are used in some occupational diving sectors to manage decompression but there is little independent assessment of their performance. A significant proportion of occupational diving operations employ single square-wave pressure exposures in support of their work. Single examples of 43 models of dive computer were compressed to five simulated depths between 15 and 50 metres' sea water (msw) and maintained at those depths until they had registered over 30 minutes of decompression. At each depth, and for each model, downloaded data were used to collate the times at which the unit was still registering "no decompression" and the times at which various levels of decompression were indicated or exceeded. Each depth profile was replicated three times for most models. Decompression isopleths for no-stop dives indicated that computers tended to be more conservative than standard decompression tables at depths shallower than 30 msw but less conservative between 30-50 msw. For dives requiring decompression, computers were predominantly more conservative than tables across the whole depth range tested. There was considerable variation between models in the times permitted at all of the depth/decompression combinations. The present study would support the use of some dive computers for controlling single, square-wave diving by some occupational sectors. The choice of which makes and models to use would have to consider their specific dive management characteristics which may additionally be affected by the intended operational depth and whether staged decompression was permitted.
Yet More Visualized JAMSTEC Cruise and Dive Information
NASA Astrophysics Data System (ADS)
Tomiyama, T.; Hase, H.; Fukuda, K.; Saito, H.; Kayo, M.; Matsuda, S.; Azuma, S.
2014-12-01
Every year, JAMSTEC performs about a hundred of research cruises and numerous dive surveys using its research vessels and submersibles. JAMSTEC provides data and samples obtained during these cruises and dives to international users through a series of data sites on the Internet. The "DARWIN (http://www.godac.jamstec.go.jp/darwin/e)" data site disseminates cruise and dive information. On DARWIN, users can search interested cruises and dives with a combination search form or an interactive tree menu, and find lists of observation data as well as links to surrounding databases. Document catalog, physical sample databases, and visual archive of dive surveys (e. g. in http://www.godac.jamstec.go.jp/jmedia/portal/e) are directly accessible from the lists. In 2014, DARWIN experienced an update, which was arranged mainly for enabling on-demand data visualization. Using login users' functions, users can put listed data items into the virtual basket and then trim, plot and download the data. The visualization tools help users to quickly grasp the quality and characteristics of observation data. Meanwhile, JAMSTEC launched a new data site named "JDIVES (http://www.godac.jamstec.go.jp/jdives/e)" to visualize data and sample information obtained by dive surveys. JDIVES shows tracks of dive surveys on the "Google Earth Plugin" and diagrams of deep-sea environmental data such as temperature, salinity, and depth. Submersible camera images and links to associated databases are placed along the dive tracks. The JDVIES interface enables users to perform so-called virtual dive surveys, which can help users to understand local geometries of dive spots and geological settings of associated data and samples. It is not easy for individual researchers to organize a huge amount of information recovered from each cruise and dive. The improved visibility and accessibility of JAMSTEC databases are advantageous not only for second-hand users, but also for on-board researchers themselves.
Blanchet, Marie-Anne; Lydersen, Christian; Ims, Rolf A.; Kovacs, Kit M.
2015-01-01
The harbour seal (Phoca vitulina) population in Svalbard marks the northernmost limit of the species’ range. This small population experiences environmental extremes in sea and air temperatures, sea ice cover and also in light regime for this normally temperate species. This study deployed Conductivity Temperature Depth Satellite Relay Data Loggers (CTD-SRDLs) on 30 adult and juvenile harbour seals in 2009 and 2010 to study their foraging behaviour across multiple seasons. A total of 189,104 dives and 16,640 CTD casts (mean depth 72 m ± 59) were recorded. Individuals dove to a mean depth of 41 m ± 24 with a maximum dive depth range of 24 – 403 m. Dives lasted on average 204 sec ± 120 with maximum durations ranging between 240 – 2,220 sec. Average daily depth and duration of dives, number of dives, time spent diving and dive time/surface time were influenced by date, while sex, age, sea-ice concentration and their interactions were not particularly influential. Dives were deeper (~150 m), longer (~480 sec), less numerous (~250 dives/day) and more pelagic during the winter/early spring compared to the fall and animals spent proportionally less time at the bottom of their dives during the winter. Influxes of warm saline water, corresponding to Atlantic Water characteristics, were observed intermittently at depths ~100 m during both winters in this study. The seasonal changes in diving behaviour were linked to average weekly wind stresses from the north or north-east, which induced upwelling events onto the shelf through offshore Ekman transport. During these events the shelf became flooded with AW from the West Spitsbergen Current, which presumably brought Atlantic fish species close to shore and within the seals’ foraging depth-range. Predicted increased in the influx of AW in this region are likely going to favour the growth and geographic expansion of this harbour seal population in the future. PMID:26196289
Cialoni, Danilo; Pieri, Massimo; Balestra, Costantino; Marroni, Alessandro
2017-01-01
Introduction: The popularity of SCUBA diving is steadily increasing together with the number of dives and correlated diseases per year. The rules that govern correct decompression procedures are considered well known even if the majority of Decompression Sickness (DCS) cases are considered unexpected confirming a bias in the "mathematical ability" to predict DCS by the current algorithms. Furthermore, little is still known about diving risk factors and any individual predisposition to DCS. This study provides an in-depth epidemiological analysis of the diving community, to include additional risk factors correlated with the development of circulating bubbles and DCS. Materials and Methods: An originally developed database (DAN DB) including specific questionnaires for data collection allowed the statistical analysis of 39,099 electronically recorded open circuit dives made by 2,629 European divers (2,189 males 83.3%, 440 females 16.7%) over 5 years. The same dive parameters and risk factors were investigated also in 970 out of the 39,099 collected dives investigated for bubble formation, by 1-min precordial Doppler, and in 320 sea-level dives followed by DCS symptoms. Results: Mean depth and GF high of all the recorded dives were 27.1 m, and 0.66, respectively; the average ascent speed was lower than the currently recommended "safe" one (9-10 m/min). We found statistically significant relationships between higher bubble grades and BMI, fat mass, age, and diving exposure. Regarding incidence of DCS, we identified additional non-bubble related risk factors, which appear significantly related to a higher DCS incidence, namely: gender, strong current, heavy exercise, and workload during diving. We found that the majority of the recorded DCS cases were not predicted by the adopted decompression algorithm and would have therefore been defined as "undeserved." Conclusion: The DAN DB analysis shows that most dives were made in a "safe zone," even if data show an evident "gray area" in the "mathematical" ability to predict DCS by the current algorithms. Some other risk factors seem to influence the possibility to develop DCS, irrespective of their effect on bubble formation, thus suggesting the existence of some factors influencing or enhancing the effects of bubbles.
Acoustic Studies of the Effects of Environmental Stresses on Marine Mammals in Large Ocean Basins
NASA Astrophysics Data System (ADS)
Sidorovskaia, N.; Ma, B.; Ackleh, A. S.; Tiemann, C.; Ioup, G. E.; Ioup, J. W.
2014-12-01
Effects of environmental stresses on deep-diving marine mammal populations have not been studied systematically. Long-term regional passive acoustic monitoring of phonating marine mammals opens opportunities for such studies. This paper presents a unique multi-year study conducted by the Littoral Acoustic Demonstration Center (LADC) in the Northern Gulf of Mexico to understand short-term and long-term effects of anthropogenic stresses on resident populations of endangered sperm and elusive beaked whales. Both species spend many hours each day in deep dives which last about one hour each, so any visual observations for population estimates and behavioral responses are very limited. However, much more cost-efficient acoustic recordings of the phonations during dives on bottom-mounted hydrophones are not skewed by weather conditions or daylight requirements. Broadband passive acoustic data were collected by LADC in 2007 and 2010 at three ranges, 15, 40, and 80 km away from the 2010 Deep Water Horizon oil spill site. Pre-spill and post-spill data processing and comparison allow observing responses of both species to local short-term environmental condition changes and long-term responses to the spill. The short-term effects are studied by correlating daily activity cycles with anthropogenic noise curve daily and weekly cycles at different sites. The strong correlation between the decrease in overall daily activity and the increase in anthropogenic noise level associated with seismic exploration signals can be seen. After streaming raw acoustic data through detection algorithms and detailed assessment of false detection rates, the temporal densities of acoustic phonations are passed into statistical algorithms for resident population estimations. The statistically significant results have shown different regional abundance trends, associated with long-term responses to environmental stresses, for these two species.
Crowell, Sara E.; Wells-Berlin, Alicia M.; Therrien, Ronald E.; Yannuzzi, Sally E.; Carr, Catherine E.
2016-01-01
Auditory sensitivity was measured in a species of diving duck that is not often kept in captivity, the lesser scaup. Behavioral (psychoacoustics) and electrophysiological [the auditory brainstem response (ABR)] methods were used to measure in-air auditory sensitivity, and the resulting audiograms were compared. Both approaches yielded audiograms with similar U-shapes and regions of greatest sensitivity (2000−3000 Hz). However, ABR thresholds were higher than psychoacoustic thresholds at all frequencies. This difference was least at the highest frequency tested using both methods (5700 Hz) and greatest at 1000 Hz, where the ABR threshold was 26.8 dB higher than the behavioral measure of threshold. This difference is commonly reported in studies involving many different species. These results highlight the usefulness of each method, depending on the testing conditions and availability of the animals.
Crowell, Sara E; Wells-Berlin, Alicia M; Therrien, Ronald E; Yannuzzi, Sally E; Carr, Catherine E
2016-05-01
Auditory sensitivity was measured in a species of diving duck that is not often kept in captivity, the lesser scaup. Behavioral (psychoacoustics) and electrophysiological [the auditory brainstem response (ABR)] methods were used to measure in-air auditory sensitivity, and the resulting audiograms were compared. Both approaches yielded audiograms with similar U-shapes and regions of greatest sensitivity (2000-3000 Hz). However, ABR thresholds were higher than psychoacoustic thresholds at all frequencies. This difference was least at the highest frequency tested using both methods (5700 Hz) and greatest at 1000 Hz, where the ABR threshold was 26.8 dB higher than the behavioral measure of threshold. This difference is commonly reported in studies involving many different species. These results highlight the usefulness of each method, depending on the testing conditions and availability of the animals.
[The research progress of diving medicine in China].
Fang, Yi-Qun; Bao, Xiao-Chen; Li, Ci; Meng, Miao; Yuan, Heng-Rong; Ma, Jun; Wang, Yan
2012-11-01
Diving medicine is one of the branches of military medicine, and plays an important role in naval development. This review introduces the progress of researches on undersea and hyperbaric physiology and medicine in the past few years in China. The article describes our research achievement in conventional diving and its medical support, researches on saturation diving and its medical support, submarine escape and its medical support, effects of hyperbaric environments and fast buoyancy ascent on immunological and cardiological functions. Diving disorders (including decompression sickness and oxygen toxicity) are also introduced.
Cialoni, Danilo; Pieri, Massimo; Balestra, Costantino; Marroni, Alessandro
2015-03-01
Inert gas accumulated after multiple recreational dives can generate tissue supersaturation and bubble formation when ambient pressure decreases. We hypothesized that this could happen even if divers respected the currently recommended 24-hour pre-flight surface interval (PFSI). We performed transthoracic echocardiography (TTE) on a group of 56 healthy scuba divers (39 male, 17 female) as follows: first echo--during the outgoing flight, no recent dives; second echo--before boarding the return flight, after a multiday diving week in the tropics and a 24-hour PFSI; third echo--during the return flight at 30, 60 and 90 minutes after take-off. TTE was also done after every dive during the week's diving. Divers were divided into three groups according to their 'bubble-proneness': non-bubblers, occasional bubblers and consistent bubblers. During the diving, 23 subjects never developed bubbles, 17 only occasionally and 16 subjects produced bubbles every day and after every dive. Bubbles on the return flight were observed in eight of the 56 divers (all from the 'bubblers' group). Two subjects who had the highest bubble scores during the diving were advised not to make the last dive (increasing their PFSI to approximately 36 hours), and did not demonstrate bubbles on the return flight. Even though a 24-hour PFSI is recommended on the basis of clinical trials showing a low risk of decompression sickness (DCS), the presence of venous gas bubbles in-flight in eight of 56 divers leads us to suspect that in real-life situations DCS risk after such a PFSI is not zero.
Clark, Natalie J; Gordos, Matthew A; Franklin, Craig E
2008-01-01
Locomotion is a common measure of performance used in studies of thermal acclimation because of its correlation with predator escape and prey capture. However, for sedentary animals such as freshwater turtles, we propose that diving behavior may be a more ecologically relevant measure of performance. Increasing dive duration in hatchling turtles reduces predator exposure and therefore functions as an ecological benefit. Diving behavior is thermally dependent, and in some species of freshwater turtles, it is also reliant on aquatic respiration. This study examined the influence of thermal acclimation on diving behavior, aquatic respiration, and locomotor performance in the endangered, bimodally respiring Mary River turtle Elusor macrurus. Diving behavior was found to partially acclimate at 17 degrees C, with turtles acclimated to a cold temperature (17 degrees C) having a significantly longer dive duration than hatchlings acclimated to a warm temperature (28 degrees C). This increase in dive duration at 17 degrees C was not a result of physiological alterations in metabolic rate but was due instead to an increase in aquatic oxygen consumption. Increasing aquatic oxygen consumption permitted cold-acclimated hatchlings to remain submerged for significantly longer periods, with one turtle undertaking a dive of over 2.5 d. When burst-swimming speed was used as the measure of performance, thermal acclimation was not detected. Overall, E. macrurus demonstrated a partial ability to acclimate to changes in environmental temperature.
NASA Technical Reports Server (NTRS)
Mattson, Axel T.
1946-01-01
The results of tests made to determine the aerodynamic characteristics of a solid brake, a slotted brake, and a dive-recovery flap mounted on a high aspect ratio wing at high Mach numbers are presented. The data were obtained in the Langley 8-foot high-speed tunnel for corrected Mach numbers up to 0.940. The results have been analyzed with regard to the suitability of dive-control devices for a proposed high-speed airplane in limiting the airplane terminal Mach number by the use of dive brakes and in achieving favorable dive-recovery characteristics by the use of a dive-recovery flap. The analysis of the results indicated that the slotted brake would limit the proposed airplane terminal Mach number to values below 0.880 for altitudes up to 35,000 feet and a wing loading of 80 pounds per square foot and the dive-recovery flap would produce trim changes required for controlled pull-outs at 25,000 feet for a Mach number range from 0.800 to 0.900. Basic changes in spanwise loading are presented to aid in the evaluation of the wing strength requirements.
Hays, G C; Akesson, S; Broderick, A C; Glen, F; Godley, B J; Luschi, P; Martin, C; Metcalfe, J D; Papi, F
2001-12-01
Satellite telemetry was used to record the submergence duration of green turtles (Chelonia mydas) as they migrated from Ascension Island to Brazil (N=12 individuals) while time/depth recorders (TDRs) were used to examine the depth distribution and dive profiles of individuals returning to Ascension Island to nest after experimental displacement (N=5 individuals). Satellite telemetry revealed that most submergences were short (<5 min) but that some submergences were longer (>20 min), particularly at night. TDRs revealed that much of the time was spent conducting short (2-4 min), shallow (approximately 0.9-1.5 m) dives, consistent with predictions for optimisation of near-surface travelling, while long (typically 20-30 min), deep (typically 10-20 m) dives had a distinctive profile found in other marine reptiles. These results suggest that green turtles crossing the Atlantic do not behave invariantly, but instead alternate between periods of travelling just beneath the surface and diving deeper. These deep dives may have evolved to reduce silhouetting against the surface, which would make turtles more susceptible to visual predators such as large sharks.
Sports-related lung injury during breath-hold diving.
Mijacika, Tanja; Dujic, Zeljko
2016-12-01
The number of people practising recreational breath-hold diving is constantly growing, thereby increasing the need for knowledge of the acute and chronic effects such a sport could have on the health of participants. Breath-hold diving is potentially dangerous, mainly because of associated extreme environmental factors such as increased hydrostatic pressure, hypoxia, hypercapnia, hypothermia and strenuous exercise.In this article we focus on the effects of breath-hold diving on pulmonary function. Respiratory symptoms have been reported in almost 25% of breath-hold divers after repetitive diving sessions. Acutely, repetitive breath-hold diving may result in increased transpulmonary capillary pressure, leading to noncardiogenic oedema and/or alveolar haemorrhage. Furthermore, during a breath-hold dive, the chest and lungs are compressed by the increasing pressure of water. Rapid changes in lung air volume during descent or ascent can result in a lung injury known as pulmonary barotrauma. Factors that may influence individual susceptibility to breath-hold diving-induced lung injury range from underlying pulmonary or cardiac dysfunction to genetic predisposition.According to the available data, breath-holding does not result in chronic lung injury. However, studies of large populations of breath-hold divers are necessary to firmly exclude long-term lung damage. Copyright ©ERS 2016.
A forensic diving medicine examination of a highly publicised scuba diving fatality.
Edmonds, Carl
2012-12-01
A high-profile diving death occurred in 2003 at the site of the wreck of the SS Yongala off the Queensland coast. The victim's buddy, her husband, was accused of her murder and found guilty of manslaughter in an Australian court. A detailed analysis of all the evidence concerning this fatality suggests alternative medical reasons for her death. The value of decompression computers in determining the diving details and of CT scans in clarifying autopsy findings is demonstrated. The victim was medically, physically and psychologically unfit to undertake the fatal dive. She was inexperienced and inadequately supervised. She was over-weighted and exposed for the first time to difficult currents. The analysis of the dive demonstrates how important it is to consider the interaction of all factors and to not make deductions from individual items of information. It also highlights the importance of early liaison between expert divers, technicians, diving clinicians and pathologists, if inappropriate conclusions are to be avoided.
Strategic Acoustic Control of a Hummingbird Courtship Dive.
Clark, Christopher J; Mistick, Emily A
2018-04-23
Male hummingbirds court females with a high-speed dive in which they "sing" with their tail feathers. The male's choice of trajectory provides him strategic control over acoustic frequency and pressure levels heard by the female. Unlike related species, male Costa's hummingbirds (Calypte costae) choose to place their dives to the side of females. Here we show that this minimizes an audible Doppler curve in their dive sound, thereby depriving females of an acoustic indicator that would otherwise reveal male dive speed. Wind-tunnel experiments indicate that the sounds produced by their feathers are directional; thus, males should aim their tail toward females. High-speed video of dives reveal that males twist half of their tail vertically during the dive, which acoustic-camera video shows effectively aims this sound sideways, toward the female. Our results demonstrate that male animals can strategically modulate female perception of dynamic aspects of athletic motor displays, such as their speed. Copyright © 2018 Elsevier Ltd. All rights reserved.
The cardiovascular system and diving risk.
Bove, Alfred A
2011-01-01
Recreational scuba diving is a sport that requires a certain physical capacity, in addition to consideration of the environmental stresses produced by increased pressure, low temperature and inert gas kinetics in tissues of the body. Factors that may influence ability to dive safely include age, physical conditioning, tolerance of cold, ability to compensate for central fluid shifts induced by water immersion, and ability to manage exercise demands when heart disease might compromise exercise capacity. Patients with coronary heart disease, valvular heart disease, congenital heart disease and cardiac arrhythmias are capable of diving, but consideration must be given to the environmental factors that might interact with the cardiac disorder. Understanding of the interaction of the diving environment with various cardiac disorders is essential to providing a safe diving environment to individual divers with known heart disease.
Gliding flight: speed and acceleration of ideal falcons during diving and pull out.
Tucker
1998-01-14
Some falcons, such as peregrines (Falco peregrinus), attack their prey in the air at the end of high-speed dives and are thought to be the fastest of animals. Estimates of their top speed in a dive range up to 157 m s-1, although speeds this high have never been accurately measured. This study investigates the aerodynamic and gravitational forces on 'ideal falcons' and uses a mathematical model to calculate speed and acceleration during diving. Ideal falcons have body masses of 0.5-2.0 kg and morphological and aerodynamic properties based on those measured for real falcons. The top speeds reached during a dive depend on the mass of the bird and the angle and duration of the dive. Given enough time, ideal falcons can reach top speeds of 89-112 m s-1 in a vertical dive, the higher speed for the heaviest bird, when the parasite drag coefficient has a value of 0.18. This value was measured for low-speed flight, and it could plausibly decline to 0.07 at high speeds. Top speeds then would be 138-174 m s-1. An ideal falcon diving at angles between 15 and 90 degrees with a mass of 1 kg reaches 95 % of top speed after travelling approximately 1200 m. The time and altitude loss to reach 95 % of top speed range from 38 s and 322 m at 15 degrees to 16 s and 1140 m at 90 degrees, respectively. During pull out at top speed from a vertical dive, the 1 kg ideal falcon can generate a lift force 18 times its own weight by reducing its wing span, compared with a lift force of 1.7 times its weight at full wing span. The falcon loses 60 m of altitude while pulling out of the dive, and lift and loss of altitude both decrease as the angle of the dive decreases. The 1 kg falcon can slow down in a dive by increasing its parasite drag and the angle of attack of its wings. Both lift and drag increase with angle of attack, but the falcon can cancel the increased lift by holding its wings in a cupped position so that part of the lift is directed laterally. The increased drag of wings producing maximum lift is great enough to decelerate the falcon at -1.5 times the acceleration of gravity at a dive angle of 45 degrees and a speed of 41 m s-1 (0.5 times top speed). Real falcons can control their speeds in a dive by changing their drag and by choosing the length of the dive. They would encounter both advantages and disadvantages by diving at the top speeds of ideal falcons, and whether they achieve those speeds remains to be investigated.
Analysis and flight evaluation of a small, fixed-wing aircraft equipped with hinged plate spoilers
NASA Technical Reports Server (NTRS)
Olcott, J. W.; Sackel, E.; Ellis, D. R.
1981-01-01
The results of a four phase effort to evaluate the application of hinged plate spoilers/dive brakes to a small general aviation aircraft are presented. The test vehicle was a single engine light aircraft modified with an experimental set of upper surface spoilers and lower surface dive brakes similar to the type used on sailplanes. The lift, drag, stick free stability, trim, and dynamic response characteristics of four different spoiler/dive brake configurations were determined. Tests also were conducted, under a wide range of flight conditions and with pilots of various experience levels, to determine the most favorable methods of spoiler control and to evaluate how spoilers might best be used during the approach and landing task. The effects of approach path angle, approach airspeed, and pilot technique using throttle/spoiler integrated control were investigated for day, night, VFR, and IFR approaches and landings. The test results indicated that spoilers offered significant improvements in the vehicle's performance and flying qualities for all elements of the approach and landing task, provided a suitable method of control was available.
Diving accidents in sports divers in Orkney waters.
Trevett, A J; Forbes, R; Rae, C K; Sheehan, C; Ross, J; Watt, S J; Stephenson, R
2001-12-01
Scapa Flow in Orkney is one of the major world centres for wreck diving. Because of the geography of Orkney and the nature of the diving, it is possible to make relatively accurate estimates of the number of dives taking place. The denominator of dive activity allows the unusual opportunity of precise calculation of accident rates. In 1999, one in every 178 sports divers visiting Orkney was involved in a significant accident, in 2000 the figure was one in 102. Some of these accidents appear to have been predictable and could be avoided by better education and preparation of visiting divers.
Penguin lungs and air sacs: implications for baroprotection, oxygen stores and buoyancy.
Ponganis, P J; St Leger, J; Scadeng, M
2015-03-01
The anatomy and volume of the penguin respiratory system contribute significantly to pulmonary baroprotection, the body O2 store, buoyancy and hence the overall diving physiology of penguins. Therefore, three-dimensional reconstructions from computerized tomographic (CT) scans of live penguins were utilized to measure lung volumes, air sac volumes, tracheobronchial volumes and total body volumes at different inflation pressures in three species with different dive capacities [Adélie (Pygoscelis adeliae), king (Aptenodytes patagonicus) and emperor (A. forsteri) penguins]. Lung volumes scaled to body mass according to published avian allometrics. Air sac volumes at 30 cm H2O (2.94 kPa) inflation pressure, the assumed maximum volume possible prior to deep dives, were two to three times allometric air sac predictions and also two to three times previously determined end-of-dive total air volumes. Although it is unknown whether penguins inhale to such high volumes prior to dives, these values were supported by (a) body density/buoyancy calculations, (b) prior air volume measurements in free-diving ducks and (c) previous suggestions that penguins may exhale air prior to the final portions of deep dives. Based upon air capillary volumes, parabronchial volumes and tracheobronchial volumes estimated from the measured lung/airway volumes and the only available morphometry study of a penguin lung, the presumed maximum air sac volumes resulted in air sac volume to air capillary/parabronchial/tracheobronchial volume ratios that were not large enough to prevent barotrauma to the non-collapsing, rigid air capillaries during the deepest dives of all three species, and during many routine dives of king and emperor penguins. We conclude that volume reduction of airways and lung air spaces, via compression, constriction or blood engorgement, must occur to provide pulmonary baroprotection at depth. It is also possible that relative air capillary and parabronchial volumes are smaller in these deeper-diving species than in the spheniscid penguin of the morphometry study. If penguins do inhale to this maximum air sac volume prior to their deepest dives, the magnitude and distribution of the body O2 store would change considerably. In emperor penguins, total body O2 would increase by 75%, and the respiratory fraction would increase from 33% to 61%. We emphasize that the maximum pre-dive respiratory air volume is still unknown in penguins. However, even lesser increases in air sac volume prior to a dive would still significantly increase the O2 store. More refined evaluations of the respiratory O2 store and baroprotective mechanisms in penguins await further investigation of species-specific lung morphometry, start-of-dive air volumes and body buoyancy, and the possibility of air exhalation during dives. © 2015. Published by The Company of Biologists Ltd.
29 CFR Appendix B to Subpart T of... - Guidelines for Scientific Diving
Code of Federal Regulations, 2014 CFR
2014-07-01
... project using scientific diving is the advancement of science; therefore, information and data resulting... data gatherer. Construction and trouble-shooting tasks traditionally associated with commercial diving...
29 CFR Appendix B to Subpart T to... - Guidelines for Scientific Diving
Code of Federal Regulations, 2012 CFR
2012-07-01
... project using scientific diving is the advancement of science; therefore, information and data resulting... data gatherer. Construction and trouble-shooting tasks traditionally associated with commercial diving...
29 CFR Appendix B to Subpart T of... - Guidelines for Scientific Diving
Code of Federal Regulations, 2013 CFR
2013-07-01
... project using scientific diving is the advancement of science; therefore, information and data resulting... data gatherer. Construction and trouble-shooting tasks traditionally associated with commercial diving...
29 CFR Appendix B to Subpart T to... - Guidelines for Scientific Diving
Code of Federal Regulations, 2011 CFR
2011-07-01
... project using scientific diving is the advancement of science; therefore, information and data resulting... data gatherer. Construction and trouble-shooting tasks traditionally associated with commercial diving...
The relative risk of decompression sickness during and after air travel following diving.
Freiberger, J J; Denoble, P J; Pieper, C F; Uguccioni, D M; Pollock, N W; Vann, R D
2002-10-01
Decompression sickness (DCS) can be provoked by post-dive flying but few data exist to quantify the risk of different post-dive, preflight surface intervals (PFSI). We conducted a case-control study using field data from the Divers Alert Network to evaluate the relative risk of DCS from flying after diving. The PFSI and the maximum depths on the last day of diving (MDLD) were analyzed from 627 recreational dive profiles. The data were divided into quartiles based on surface interval and depth. Injured divers (cases) and uninjured divers (controls) were compared using logistic regression to determine the association of DCS with time and depth while controlling for diver and dive profiles characteristics. These included PFSI, MDLD, gender, height, weight, age, and days of diving. The means (+/-SD) for cases and controls were as follows: PFSI, 20.7 +/- 9.6 h vs. 27.1 +/- 6.7 h; MDLD, 22.5 +/- 14 meters sea water (msw) vs. 19 +/- 11.3 msw; male gender, 60% vs. 70%; weight, 75.8 +/- 18 kg vs. 77.6 +/- 16 kg; height, 173 +/- 16 cm vs. 177 +/- 9 cm; age, 36.8 +/- 10 yr vs. 42.9 +/- 11 yr; diving > or = 3 d, 58% vs. 97%. Relative to flying > 28 h after diving, the odds of DCS (95% CI) were: 1.02 (0.61, 1.7) 24-28 h; 1.84 (1.0, 3.3) 20-24 h; and 8.5 (3.85, 18.9) < 20 h. Relative to a depth of < 14.7 msw, the odds of DCS (95% CI) were: 1.2 (0.6, 1.7) 14.7-18.5 msw; 2.9 (1.65, 5.3) 18.5-26 msw; and 5.5 (2.96, 1 0.0) > 26 msw. Odds ratios approximate relative risk in rare diseases such as DCS. This study demonstrated an increase in relative risk from flying after diving following shorter PFSIs and/or greater dive depths on the last day. The relative risk increases geometrically as the PFSI becomes smaller.
Aoki, Kagari; Sato, Katsufumi; Isojunno, Saana; Narazaki, Tomoko; Miller, Patrick J O
2017-10-15
To maximize foraging duration at depth, diving mammals are expected to use the lowest cost optimal speed during descent and ascent transit and to minimize the cost of transport by achieving neutral buoyancy. Here, we outfitted 18 deep-diving long-finned pilot whales with multi-sensor data loggers and found indications that their diving strategy is associated with higher costs than those of other deep-diving toothed whales . Theoretical models predict that optimal speed is proportional to (basal metabolic rate/drag) 1/3 and therefore to body mass 0.05 The transit speed of tagged animals (2.7±0.3 m s -1 ) was substantially higher than the optimal speed predicted from body mass (1.4-1.7 m s -1 ). According to the theoretical models, this choice of high transit speed, given a similar drag coefficient (median, 0.0035) to that in other cetaceans, indicated greater basal metabolic costs during diving than for other cetaceans. This could explain the comparatively short duration (8.9±1.5 min) of their deep dives (maximum depth, 444±85 m). Hydrodynamic gliding models indicated negative buoyancy of tissue body density (1038.8±1.6 kg m -3 , ±95% credible interval, CI) and similar diving gas volume (34.6±0.6 ml kg -1 , ±95% CI) to those in other deep-diving toothed whales. High diving metabolic rate and costly negative buoyancy imply a 'spend more, gain more' strategy of long-finned pilot whales, differing from that in other deep-diving toothed whales, which limits the costs of locomotion during foraging. We also found that net buoyancy affected the optimal speed: high transit speeds gradually decreased during ascent as the whales approached neutral buoyancy owing to gas expansion. © 2017. Published by The Company of Biologists Ltd.
Individual Foraging Strategies Reveal Niche Overlap between Endangered Galapagos Pinnipeds
Villegas-Amtmann, Stella; Jeglinski, Jana W. E.; Costa, Daniel P.; Robinson, Patrick W.; Trillmich, Fritz
2013-01-01
Most competition studies between species are conducted from a population-level approach. Few studies have examined inter-specific competition in conjunction with intra-specific competition, with an individual-based approach. To our knowledge, none has been conducted on marine top predators. Sympatric Galapagos fur seals (Arctocephalus galapagoensis) and sea lions (Zalophus wollebaeki) share similar geographic habitats and potentially compete. We studied their foraging niche overlap at Cabo Douglas, Fernandina Island from simultaneously collected dive and movement data to examine spatial and temporal inter- and intra-specific competition. Sea lions exhibited 3 foraging strategies (shallow, intermediate and deep) indicating intra-specific competition. Fur seals exhibited one foraging strategy, diving predominantly at night, between 0–80 m depth and mostly at 19–22 h. Most sea lion dives also occurred at night (63%), between 0–40 m, within fur seals' diving depth range. 34% of sea lions night dives occurred at 19–22 h, when fur seals dived the most, but most of them occurred at dawn and dusk, when fur seals exhibited the least amount of dives. Fur seals and sea lions foraging behavior overlapped at 19 and 21 h between 0–30 m depths. Sea lions from the deep diving strategy exhibited the greatest foraging overlap with fur seals, in time (19 h), depth during overlapping time (21–24 m), and foraging range (37.7%). Fur seals foraging range was larger. Cabo Douglas northwest coastal area, region of highest diving density, is a foraging “hot spot” for both species. Fur seals and sea lions foraging niche overlap occurred, but segregation also occurred; fur seals primarily dived at night, while sea lions exhibited night and day diving. Both species exploited depths and areas exclusive to their species. Niche breadth generally increases with environmental uncertainty and decreased productivity. Potential competition between these species could be greater during warmer periods when prey availability is reduced. PMID:23967096
Pontier, J-M; Lambrechts, K
2014-06-01
We highlighted a relationship between decompression-induced bubble formation and platelet micro-particle (PMP) release after a scuba air-dive. It is known that decompression protocol using oxygen-stop accelerates the washout of nitrogen loaded in tissues. The aim was to study the effect of oxygen deco-stop on bubble formation and cell-derived MP release. Healthy experienced divers performed two scuba-air dives to 30 msw for 30 min, one with an air deco-stop and a second with 100% oxygen deco-stop at 3 msw for 9 min. Bubble grades were monitored with ultrasound and converted to the Kisman integrated severity score (KISS). Blood samples for cell-derived micro-particle analysis (AnnexinV for PMP and CD31 for endothelial MP) were taken 1 h before and after each dive. Mean KISS bubble score was significantly lower after the dive with oxygen-decompression stop, compared to the dive with air-decompression stop (4.3 ± 7.3 vs. 32.7 ± 19.9, p < 0.001). After the dive with an air-breathing decompression stop, we observed an increase of the post-dive mean values of PMP (753 ± 245 vs. 381 ± 191 ng/μl, p = 0.003) but no significant change in the oxygen-stop decompression dive (329 ± 215 vs. 381 +/191 ng/μl, p = 0.2). For the post-dive mean values of endothelial MP, there was no significant difference between both the dives. The Oxygen breathing during decompression has a beneficial effect on bubble formation accelerating the washout of nitrogen loaded in tissues. Secondary oxygen-decompression stop could reduce bubble-induced platelet activation and the pro-coagulant activity of PMP release preventing the thrombotic event in the pathogenesis of decompression sickness.
The Effect of a Diving Mask on Intraocular Pressure in a Healthy Population.
Goenadi, Catherina Josephine; Law, David Zhiwei; Lee, Jia Wen; Ong, Ee Lin; Chee, Wai Kitt; Cheng, Jason
2016-01-01
Swimming goggles increase the intraocular pressure (IOP) via the periorbital frame pressure and suction effect. In comparison, diving masks have a larger frame rim and incorporate the nose. The exact effect(s) of diving masks on IOP is unknown. This study evaluates the influence of diving masks on IOP in normal, healthy subjects. Tonometry was performed in both eyes of all subjects with an AVIA(®)Tono-Pen by a single investigator. Measurements were taken at baseline without the diving mask and with the subjects wearing a small-volume, double-window diving mask, but with the mask lenses removed. Two IOP readings in each eye were measured, and an additional reading was measured if the difference between the initial 2 was ≥2 mm Hg. Central corneal thickness (CCT) was also measured in each eye, using a contact pachymeter (OcuScan(®)Alcon). Forty eyes of 20 healthy volunteers (age 29.7 ± 9.3 years; range 21-52) were included. The mean CCT was 544.4 ± 43.5 µm. The mean IOP before the diving mask was worn had been 17.23 ± 2.18 mm Hg (n = 40). The IOP decreased by 0.43 mm Hg (p $1003c; 0.05) to 16.80 ± 2.57 mm Hg after the diving mask had been put on. There was no correlation between IOP change and age (r = 0.143, p = 0.337), gender (r = -0.174, p = 0.283) or CCT (r = -0.123, p = 0.445). There was no increase in IOP after the diving mask had been worn. A small but statistically significant decrease in IOP was observed. This study demonstrates that unlike swimming goggles, the strap tension and frame pressure on the periorbital tissue from a diving mask does not increase IOP. Diving masks may be a suitable alternative to swimming goggles for patients with advanced glaucoma or glaucoma filtration surgery.
The Effect of a Diving Mask on Intraocular Pressure in a Healthy Population
Goenadi, Catherina Josephine; Law, David Zhiwei; Lee, Jia Wen; Ong, Ee Lin; Chee, Wai Kitt; Cheng, Jason
2016-01-01
Purpose Swimming goggles increase the intraocular pressure (IOP) via the periorbital frame pressure and suction effect. In comparison, diving masks have a larger frame rim and incorporate the nose. The exact effect(s) of diving masks on IOP is unknown. This study evaluates the influence of diving masks on IOP in normal, healthy subjects. Methods Tonometry was performed in both eyes of all subjects with an AVIA®Tono-Pen by a single investigator. Measurements were taken at baseline without the diving mask and with the subjects wearing a small-volume, double-window diving mask, but with the mask lenses removed. Two IOP readings in each eye were measured, and an additional reading was measured if the difference between the initial 2 was ≥2 mm Hg. Central corneal thickness (CCT) was also measured in each eye, using a contact pachymeter (OcuScan®Alcon). Results Forty eyes of 20 healthy volunteers (age 29.7 ± 9.3 years; range 21–52) were included. The mean CCT was 544.4 ± 43.5 µm. The mean IOP before the diving mask was worn had been 17.23 ± 2.18 mm Hg (n = 40). The IOP decreased by 0.43 mm Hg (p $1003c; 0.05) to 16.80 ± 2.57 mm Hg after the diving mask had been put on. There was no correlation between IOP change and age (r = 0.143, p = 0.337), gender (r = −0.174, p = 0.283) or CCT (r = −0.123, p = 0.445). Conclusion There was no increase in IOP after the diving mask had been worn. A small but statistically significant decrease in IOP was observed. This study demonstrates that unlike swimming goggles, the strap tension and frame pressure on the periorbital tissue from a diving mask does not increase IOP. Diving masks may be a suitable alternative to swimming goggles for patients with advanced glaucoma or glaucoma filtration surgery. PMID:27462262
Cook, Timothée R.; Kato, Akiko; Tanaka, Hideji; Ropert-Coudert, Yan; Bost, Charles-André
2010-01-01
Background Because they have air stored in many body compartments, diving seabirds are expected to exhibit efficient behavioural strategies for reducing costs related to buoyancy control. We study the underwater locomotor activity of a deep-diving species from the Cormorant family (Kerguelen shag) and report locomotor adjustments to the change of buoyancy with depth. Methodology/Principal Findings Using accelerometers, we show that during both the descent and ascent phases of dives, shags modelled their acceleration and stroking activity on the natural variation of buoyancy with depth. For example, during the descent phase, birds increased swim speed with depth. But in parallel, and with a decay constant similar to the one in the equation explaining the decrease of buoyancy with depth, they decreased foot-stroke frequency exponentially, a behaviour that enables birds to reduce oxygen consumption. During ascent, birds also reduced locomotor cost by ascending passively. We considered the depth at which they started gliding as a proxy to their depth of neutral buoyancy. This depth increased with maximum dive depth. As an explanation for this, we propose that shags adjust their buoyancy to depth by varying the amount of respiratory air they dive with. Conclusions/Significance Calculations based on known values of stored body oxygen volumes and on deep-diving metabolic rates in avian divers suggest that the variations of volume of respiratory oxygen associated with a respiration mediated buoyancy control only influence aerobic dive duration moderately. Therefore, we propose that an advantage in cormorants - as in other families of diving seabirds - of respiratory air volume adjustment upon diving could be related less to increasing time of submergence, through an increased volume of body oxygen stores, than to reducing the locomotor costs of buoyancy control. PMID:20352122
Wallace, Bryan P; Williams, Cassondra L; Paladino, Frank V; Morreale, Stephen J; Lindstrom, R Todd; Spotila, James R
2005-10-01
Physiology, environment and life history demands interact to influence marine turtle bioenergetics and activity. However, metabolism and diving behavior of free-swimming marine turtles have not been measured simultaneously. Using doubly labeled water, we obtained the first field metabolic rates (FMRs; 0.20-0.74 W kg(-1)) and water fluxes (16-30% TBW day(-1), where TBW=total body water) for free-ranging marine turtles and combined these data with dive information from electronic archival tags to investigate the bioenergetics and diving activity of reproductive adult female leatherback turtles Dermochelys coriacea. Mean dive durations (7.8+/-2.4 min (+/-1 s.d.), bottom times (2.7+/-0.8 min), and percentage of time spent in water temperatures (Tw) < or =24 degrees C (9.5+/-5.7%) increased with increasing mean maximum dive depths (22.6+/-7.1 m; all P< or =0.001). The FMRs increased with longer mean dive durations, bottom times and surface intervals and increased time spent in Tw< or =24 degrees C (all r2> or =0.99). This suggests that low FMRs and activity levels, combined with shuttling between different water temperatures, could allow leatherbacks to avoid overheating while in warm tropical waters. Additionally, internesting leatherback dive durations were consistently shorter than aerobic dive limits calculated from our FMRs (11.7-44.3 min). Our results indicate that internesting female leatherbacks maintained low FMRs and activity levels, thereby spending relatively little energy while active at sea. Future studies should incorporate data on metabolic rate, dive patterns, water temperatures, and body temperatures to develop further the relationship between physiological and life history demands and marine turtle bioenergetics and activity.
Schorr, Gregory S; Falcone, Erin A; Moretti, David J; Andrews, Russel D
2014-01-01
Cuvier's beaked whales (Ziphius cavirostris) are known as extreme divers, though behavioral data from this difficult-to-study species have been limited. They are also the species most often stranded in association with Mid-Frequency Active (MFA) sonar use, a relationship that remains poorly understood. We used satellite-linked tags to record the diving behavior and locations of eight Ziphius off the Southern California coast for periods up to three months. The effort resulted in 3732 hr of dive data with associated regional movements--the first dataset of its kind for any beaked whale--and included dives to 2992 m depth and lasting 137.5 min, both new mammalian dive records. Deep dives had a group mean depth of 1401 m (s.d. = 137.8, n = 1142) and duration of 67.4 min (s.d. = 6.9). The group mean time between deep dives was 102.3 min (s.d. = 30.8, n = 783). While the previously described stereotypic pattern of deep and shallow dives was apparent, there was considerable inter- and intra-individual variability in most parameters. There was significant diel behavioral variation, including increased time near the surface and decreased shallow diving at night. However, maximum depth and the proportion of time spent on deep dives (presumed foraging), varied little from day to night. Surprisingly, tagged whales were present within an MFA sonar training range for 38% of days locations were received, and though comprehensive records of sonar use during tag deployments were not available, we discuss the effects frequent acoustic disturbance may have had on the observed behaviors. These data better characterize the true behavioral range of this species, and suggest caution should be exercised when drawing conclusions about behavior using short-term datasets.
Open water scuba diving accidents at Leicester: five years' experience.
Hart, A J; White, S A; Conboy, P J; Bodiwala, G; Quinton, D
1999-01-01
OBJECTIVES: The aim of this study was to determine the incidence, type, outcome, and possible risk factors of diving accidents in each year of a five year period presenting from one dive centre to a large teaching hospital accident and emergency (A&E) department. METHODS: All patients included in this study presented to the A&E department at a local teaching hospital in close proximity to the largest inland diving centre in the UK. Our main outcome measures were: presenting symptoms, administration of recompression treatment, mortality, and postmortem examination report where applicable. RESULTS: Overall, 25 patients experienced a serious open water diving accident at the centre between 1992 and 1996 inclusive. The percentage of survivors (n = 18) with symptoms of decompression sickness receiving recompression treatment was 52%. All surviving patients received medical treatment for at least 24 hours before discharge. The median depth of diving accidents was 24 metres (m) (range 7-36 m). During the study period, 1992-96, the number of accidents increased from one to 10 and the incidence of diving accidents increased from four per 100,000 to 15.4 per 100,000. Over the same time period the number of deaths increased threefold. CONCLUSIONS: The aetiology of the increase in the incidence of accidents is multifactorial. Important risk factors were thought to be: rapid ascent (in 48% of patients), cold water, poor visibility, the number of dives per diver, and the experience of the diver. It is concluded that there needs to be an increased awareness of the management of diving injuries in an A&E department in close proximity to an inland diving centre. PMID:10353047
Goldbogen, Jeremy A; Calambokidis, John; Croll, Donald A; Harvey, James T; Newton, Kelly M; Oleson, Erin M; Schorr, Greg; Shadwick, Robert E
2008-12-01
Lunge feeding in rorqual whales is a drag-based feeding mechanism that is thought to entail a high energetic cost and consequently limit the maximum dive time of these extraordinarily large predators. Although the kinematics of lunge feeding in fin whales supports this hypothesis, it is unclear whether respiratory compensation occurs as a consequence of lunge-feeding activity. We used high-resolution digital tags on foraging humpback whales (Megaptera novaengliae) to determine the number of lunges executed per dive as well as respiratory frequency between dives. Data from two whales are reported, which together performed 58 foraging dives and 451 lunges. During one study, we tracked one tagged whale for approximately 2 h and examined the spatial distribution of prey using a digital echosounder. These data were integrated with the dive profile to reveal that lunges are directed toward the upper boundary of dense krill aggregations. Foraging dives were characterized by a gliding descent, up to 15 lunges at depth, and an ascent powered by steady swimming. Longer dives were required to perform more lunges at depth and these extended apneas were followed by an increase in the number of breaths taken after a dive. Maximum dive durations during foraging were approximately half of those previously reported for singing (i.e. non-feeding) humpback whales. At the highest lunge frequencies (10 to 15 lunges per dive), respiratory rate was at least threefold higher than that of singing humpback whales that underwent a similar degree of apnea. These data suggest that the high energetic cost associated with lunge feeding in blue and fin whales also occurs in intermediate sized rorquals.
Recreational technical diving part 1: an introduction to technical diving methods and activities.
Mitchell, Simon J; Doolette, David J
2013-06-01
Technical divers use gases other than air and advanced equipment configurations to conduct dives that are deeper and/or longer than typical recreational air dives. The use of oxygen-nitrogen (nitrox) mixes with oxygen fractions higher than air results in longer no-decompression limits for shallow diving, and faster decompression from deeper dives. For depths beyond the air-diving range, technical divers mix helium, a light non-narcotic gas, with nitrogen and oxygen to produce 'trimix'. These blends are tailored to the depth of intended use with a fraction of oxygen calculated to produce an inspired oxygen partial pressure unlikely to cause cerebral oxygen toxicity and a nitrogen fraction calculated to produce a tolerable degree of nitrogen narcosis. A typical deep technical dive will involve the use of trimix at the target depth with changes to gases containing more oxygen and less inert gas during the decompression. Open-circuit scuba may be used to carry and utilise such gases, but this is very wasteful of expensive helium. There is increasing use of closed-circuit 'rebreather' devices. These recycle expired gas and potentially limit gas consumption to a small amount of inert gas to maintain the volume of the breathing circuit during descent and the amount of oxygen metabolised by the diver. This paper reviews the basic approach to planning and execution of dives using these methods to better inform physicians of the physical demands and risks.
To what extent might N2 limit dive performance in king penguins?
Fahlman, A; Schmidt, A; Jones, D R; Bostrom, B L; Handrich, Y
2007-10-01
A mathematical model was used to explore if elevated levels of N2, and risk of decompression sickness (DCS), could limit dive performance (duration and depth) in king penguins (Aptenodytes patagonicus). The model allowed prediction of blood and tissue (central circulation, muscle, brain and fat) N2 tensions (P(N2)) based on different cardiac outputs and blood flow distributions. Estimated mixed venous P(N2) agreed with values observed during forced dives in a compression chamber used to validate the assumptions of the model. During bouts of foraging dives, estimated mixed venous and tissue P(N2) increased as the bout progressed. Estimated mean maximum mixed venous P(N2) upon return to the surface after a dive was 4.56+/-0.18 atmospheres absolute (ATA; range: 4.37-4.78 ATA). This is equivalent to N2 levels causing a 50% DCS incidence in terrestrial animals of similar mass. Bout termination events were not associated with extreme mixed venous N2 levels. Fat P(N2) was positively correlated with bout duration and the highest estimated fat P(N2) occurred at the end of a dive bout. The model suggested that short and shallow dives occurring between dive bouts help to reduce supersaturation and thereby DCS risk. Furthermore, adipose tissue could also help reduce DCS risk during the first few dives in a bout by functioning as a sink to buffer extreme levels of N2.
Federal Register 2010, 2011, 2012, 2013, 2014
2011-12-09
....--Principal Investigator) has applied for an amendment to Scientific Research Permit No. 14534-01. DATES... during studies of sound production, diving, responses to sound, and other behavior. The research is... significant environmental impacts could result from issuance of the proposed scientific research permit. The...
NASA Astrophysics Data System (ADS)
Kitazato, Hiroshi; Fujikura, Katsunori; Sumida, Paulo G. Y.; Pellizari, Vivian H.; Perez, Jose Angel
2017-12-01
The deep sea comprises a series of extreme environments, characterized by low temperatures, high hydraulic pressure, the virtual absence of sunlight, and the scarcity of organic nutrients. High hydraulic pressure prevents human access to these environments without using human-occupied submersibles (HOVs), landers or other instruments. Unlike the extreme outer space environments, visited by hundreds of astronauts, only three humans have accessed to the deepest point of the planet.
First aid kits for recreational dive boats, what should they contain?
Pye, Jacqueline; Greenhalgh, Trisha
2010-09-01
Well-equipped first-aid kits are necessary but not always provided on recreational dive boats. We aimed to review the types of illness and injury likely to be encountered on such boats and inform a content list for such kits. We conducted a 3-round Delphi study by email using a volunteer panel of 18 experts drawn from diving, dive medicine and nursing. In round 1, panellists shared examples of illnesses and injuries they had come across personally. These scenarios were circulated along with findings from a literature review, including existing recommendations. In rounds 2 and 3, the list of kit for dive boats in different settings was iteratively refined through online discussion and feedback. Passengers and crew on recreational dive boats may encounter a range of medical problems from minor injuries to serious accidents and non-dive-related illnesses. Recommended kit varied depending on context and setting (e.g. distance from land, qualifications and experience of crew). Consensus was quickly reached on key first-aid items but experts' views on emergency medicines differed. The study highlights the diversity of medical problems encountered on recreational dive boats. We offer preliminary guidance on the content of suitable first-aid kits and suggest areas for further research. Copyright © 2010 Elsevier Ltd. All rights reserved.
[The heart and underwater diving].
Lafay, V
2006-11-01
Cardiovascular examination of a certain number of candidates for underwater diving raises justifiable questions of aptitude. An indicative list of contraindications has been proposed by the French Federation of Underwater Studies and Sports but a physiopathological basis gives a better understanding of what is involved. During diving, the haemodynamic changes due not only to the exercise but also to cold immersion, hyperoxaemia and decompression impose the absence of any symptomatic cardiac disease. Moreover, the vasoconstriction caused by the cold and hyperoxaemia should incite great caution in both coronary and hypertensive patients. The contraindication related to betablocker therapy is controversial and the debate has not been settled in France. The danger of drowning makes underwater diving hazardous in all pathologies carrying a risk of syncope. Pacemaker patients should be carefully assessed and the depth of diving limited. Finally, the presence of right-to-left intracardiac shunts increases the risk of complications during decompressionand contraindicates underwater diving. Patent foramen ovale is a special case but no special investigation is required for its detection. The cardiologist examining candidates for underwater diving should take all these factors into consideration because, although underwater diving is a sport associated with an increased risk, each year there are more and more people, with differing degrees of aptitude, who wish to practice it.
Aerospace Physiology, AFSC 911X0
1988-11-01
EXPERIMENTAL HYPOBARIC OR HYPERBARIC CHAMBERS 50 E149 REVIEW RESEARCH SUBJECT RECORDS FOR COMPLIANCE WITH HUMAN USE COMMITTEE DIRECTIVES 38 AS -. - t...5 Career Ladder Structure .. .. ....... ...... ...... 5 Overview. ........................... 5 Comparison to...30 MAJCOM Comparison .. .. ...... ....... ......... 30 Overlap with AFSC 122X0, Aircrew Life Support .. .. ........ 36 Number of flights, dives, and
Decompression syndrome and the evolution of deep diving physiology in the Cetacea
NASA Astrophysics Data System (ADS)
Beatty, Brian Lee; Rothschild, Bruce M.
2008-09-01
Whales repetitively dive deep to feed and should be susceptible to decompression syndrome, though they are not known to suffer the associated pathologies. Avascular osteonecrosis has been recognized as an indicator of diving habits of extinct marine amniotes. Vertebrae of 331 individual modern and 996 fossil whales were subjected to macroscopic and radiographic examination. Avascular osteonecrosis was found in the Oligocene basal odontocetes (Xenorophoidea) and in geologically younger mysticetes, such as Aglaocetus [a sister taxon to Balaenopteridae + (Balaenidae + Eschrichtiidae) clade]. These are considered as early “experiments” in repetitive deep diving, indicating that they independently converged on their similar specialized diving physiologies.
Shallow Water Diving - The NASA Experience
NASA Technical Reports Server (NTRS)
Fitzpatrick, Daniel; Kelsey-Seybold
2010-01-01
This slide presentation reviews some of the problems and solutions that personnel have experienced during sessions in the Neutral Bu0yancy Lab (NBL). It reviews the standard dive that occurs at the NBL, Boyles and Henry's laws as they relate to the effects of diving. It then reviews in depth some of the major adverse physiologic events that happen during a diving session: Ear and Sinus Barotrauma, Decompression Sickness, (DCS), Pulmonary Barotrauma (i.e., Arterial Gas Embolism (AGE). Mediastinal Emphysema, Subcutaneous Emphysema, and Pneumothorax) Oxygen Toxicity and Hypothermia. It includes information about the pulmonary function in NBL divers. Also included is recommendations about flying after diving.
Decompression syndrome and the evolution of deep diving physiology in the Cetacea.
Beatty, Brian Lee; Rothschild, Bruce M
2008-09-01
Whales repetitively dive deep to feed and should be susceptible to decompression syndrome, though they are not known to suffer the associated pathologies. Avascular osteonecrosis has been recognized as an indicator of diving habits of extinct marine amniotes. Vertebrae of 331 individual modern and 996 fossil whales were subjected to macroscopic and radiographic examination. Avascular osteonecrosis was found in the Oligocene basal odontocetes (Xenorophoidea) and in geologically younger mysticetes, such as Aglaocetus [a sister taxon to Balaenopteridae + (Balaenidae + Eschrichtiidae) clade]. These are considered as early "experiments" in repetitive deep diving, indicating that they independently converged on their similar specialized diving physiologies.
29 CFR 1910.422 - Procedures during dive.
Code of Federal Regulations, 2011 CFR
2011-07-01
... applicable to each diving operation unless otherwise specified. (b) Water entry and exit. (1) A means capable... from a dive team member; (3) Communications are lost and can not be quickly re-established between the...
29 CFR 1910.422 - Procedures during dive.
Code of Federal Regulations, 2012 CFR
2012-07-01
... applicable to each diving operation unless otherwise specified. (b) Water entry and exit. (1) A means capable... from a dive team member; (3) Communications are lost and can not be quickly re-established between the...
29 CFR 1910.422 - Procedures during dive.
Code of Federal Regulations, 2010 CFR
2010-07-01
... applicable to each diving operation unless otherwise specified. (b) Water entry and exit. (1) A means capable... from a dive team member; (3) Communications are lost and can not be quickly re-established between the...
29 CFR 1910.422 - Procedures during dive.
Code of Federal Regulations, 2014 CFR
2014-07-01
... applicable to each diving operation unless otherwise specified. (b) Water entry and exit. (1) A means capable... from a dive team member; (3) Communications are lost and can not be quickly re-established between the...
29 CFR 1910.422 - Procedures during dive.
Code of Federal Regulations, 2013 CFR
2013-07-01
... applicable to each diving operation unless otherwise specified. (b) Water entry and exit. (1) A means capable... from a dive team member; (3) Communications are lost and can not be quickly re-established between the...
Effects of oxygen-enriched air on cognitive performance during SCUBA-diving - an open-water study.
Brebeck, Anne-Kathrin; Deussen, Andreas; Schmitz-Peiffer, Henning; Range, Ursula; Balestra, Costantino; Cleveland, Sinclair; Schipke, Jochen D
2017-01-01
Backround: Nitrogen narcosis impairs cognitive function, a fact relevant during SCUBA-diving. Oxygen-enriched air (nitrox) became popular in recreational diving, while evidence of its advantages over air is limited. Compare effects of nitrox28 and air on two psychometric tests. In this prospective, double-blind, open-water study, 108 advanced divers (38 females) were randomized to an air or a nitrox-group for a 60-min dive to 24 m salt water. Breathing gas effects on cognitive performance were assessed during the dive using a short- and long-term memory test and a number connection test. Nitrox28 divers made fewer mistakes only on the long-term memory test (p = 0.038). Female divers remembered more items than male divers (p < 0.001). There were no significant differences in the number connection test between the groups. Likely owing to the comparatively low N 2 reduction and the conservative dive, beneficial nitrox28 effects to diver performance were moderate but could contribute to diving safety.
Comparison of functional aspects of the coagulation cascade in human and sea turtle plasmas.
Soslau, Gerald; Wallace, Bryan; Vicente, Catherine; Goldenberg, Seth J; Tupis, Todd; Spotila, James; George, Robert; Paladino, Frank; Whitaker, Brent; Violetta, Gary; Piedra, Rotney
2004-08-01
Functional hemostatic pathways are critical for the survival of all vertebrates and have been evolving for more than 400 million years. The overwhelming majority of studies of hemostasis in vertebrates have focused on mammals with very sparse attention paid to reptiles. There have been virtually no studies of the coagulation pathway in sea turtles whose ancestors date back to the Jurassic period. Sea turtles are often exposed to rapidly altered environmental conditions during diving periods. This may reduce their blood pH during prolonged hypoxic dives. This report demonstrates that five species of turtles possess only one branch of the mammalian coagulation pathway, the extrinsic pathway. Mixing studies of turtle plasmas with human factor-deficient plasmas indicate that the intrinsic pathway factors VIII and IX are present in turtle plasma. These two factors may play a significant role in supporting the extrinsic pathway by feedback loops. The intrinsic factors, XI and XII are not detected which would account for the inability of reagents to induce coagulation via the intrinsic pathway in vitro. The analysis of two turtle factors, factor II (prothrombin) and factor X, demonstrates that they are antigenically/functionally similar to the corresponding human factors. The turtle coagulation pathway responds differentially to both pH and temperature relative to each turtle species and relative to human samples. The coagulation time (prothrombin time) increases as the temperature decreases between 37 and 15 degrees C. The increased time follows a linear relationship, with similar slopes for loggerhead, Kemps ridley and hawksbill turtles as well as for human samples. Leatherback turtle samples show a dramatic nonlinear increased time below 23 degrees C, and green turtle sample responses were similar but less dramatic. All samples also showed increased prothrombin times as the pH decreased from 7.8 to 6.4, except for three turtle species. The prothrombin times decreased, to varying extents, in a linear fashion relative to reduced pH with the rate of change greatest in leatherbacks>green>loggerhead turtles. All studies were conducted with reagents developed for human samples which would impact on the quantitative results with the turtle samples, but are not likely to alter the qualitative results. These comparative studies of the coagulation pathway in sea turtles and humans could enhance our knowledge of structure/function relationships and evolution of coagulation factors.
Pilla, R; Held, H E; Landon, C S; Dean, J B
2013-08-29
Pseudoephedrine (PSE) salts (hydrochloride and sulfate) are commonly used as nasal and paranasal decongestants by scuba divers. Anecdotal reports from the Divers Alert Network suggest that taking PSE prior to diving while breathing pure O₂ increases the risk for CNS oxygen toxicity (CNS-OT), which manifests as seizures. We hypothesized that high doses of PSE reduce the latency time to seizure (LS) in unanesthetized rats breathing 5 atmospheres absolute (ATA) of hyperbaric oxygen. Sixty-three male rats were implanted with radio-transmitters that recorded electroencephalogram activity and body temperature. After ≥7-day recovery, and 2 h before "diving", each rat was administered either saline solution (control) or PSE hydrochloride intragastrically at the following doses (mg PSE/kg): 0, 40, 80, 100, 120, 160, and 320. Rats breathed pure O₂ and were dived to 5ATA until the onset of behavioral seizures coincident with neurological seizures. LS was the time elapsed between reaching 5ATA and exhibiting seizures. We observed a significant dose-dependent decrease in the LS at doses of 100-320 mg/kg, whereas no significant differences in LS from control value were observed at doses ≤80 mg/kg. Our findings showed that high doses of PSE accelerate the onset of CNS-OT seizures in unanesthetized rats breathing 5ATA of poikilocapnic hyperoxia. Extrapolating our findings to humans, we conclude that the recommended daily dose of PSE should not be abused prior to diving with oxygen-enriched gas mixes or pure O₂. Copyright © 2013 IBRO. Published by Elsevier Ltd. All rights reserved.
Underwater and Dive Station Work-Site Noise Surveys
2008-03-14
A) octave band noise measurements, dB (A) correction factors, dB ( A ) levels , MK-21 diving helmet attenuation correction factors, overall in-helmet...band noise measurements, dB (A) correction factors, dB ( A ) levels , MK-21 diving helmet attenuation correction factors, overall in-helmet dB (A...noise measurements, dB (A) correction factors, dB ( A ) levels , MK-21 diving helmet attenuation correction factors, overall in-helmet dB (A) level, and
Sharon L. Todd; Tiffany Cooper; Alan R. Graefe
2001-01-01
This study examined SCUBA divers' level of development in relationship to environmental beliefs, ascriptions of responsibility, and management preferences concerning the use and management of New York's Great Lakes' underwater cultural resources. More than 850 New York State divers were surveyed during the fall of 1999, ranging from novices to experts...
Gagnon, Dominique D; Pretorius, Thea; McDonald, Gerren; Kenny, Glen P; Giesbrecht, Gordon G
2013-06-01
Facial cooling can regulate reflexes of the dive response whereas further body cooling generally induces the cold-shock response. We examined the cardiovascular and ventilatory parameters of these responses during 3-min immersions of the head dorsum, face, and whole head in 17 degrees C water while breathing was maintained. From a horizontal position, the head was inserted into a temperature controlled immersion tank in which the water level could be changed rapidly. On four occasions, either the head dorsum, face or whole head (prone and supine) were exposed to water. Mean decrease in heart rate (14%) and increases in systolic (9%) and diastolic (5%) blood pressures were seen during immersion. Relative mean finger skin blood flow had an early transient decrease (31%) for 90 s and then returned to baseline values. A strong transient increase was seen in minute ventilation (92%) at 20 s of immersion via tidal volume (85%). There were no consistent differences between the head dorsum, face, and whole head for all variables in response to immersion. The cold-shock response (increased minute ventilation and tidal volume) predominated over the dive response in the initial moments of immersion only. The order of emergence of these responses provides further recommendation to avoid head submersion upon cold water entry. It is important to protect the face, with a facemask, and the head dorsum, with an insulative hood, in cold water.
Variability in circulating gas emboli after a same scuba diving exposure.
Papadopoulou, V; Germonpré, P; Cosgrove, D; Eckersley, R J; Dayton, P A; Obeid, G; Boutros, A; Tang, M-X; Theunissen, S; Balestra, C
2018-06-01
A reduction in ambient pressure or decompression from scuba diving can result in ultrasound-detectable venous gas emboli (VGE). These environmental exposures carry a risk of decompression sickness (DCS) which is mitigated by adherence to decompression schedules; however, bubbles are routinely observed for dives well within these limits and significant inter-personal variability in DCS risk exists. Here, we assess the variability and evolution of VGE for 2 h post-dive using echocardiography, following a standardized pool dive in calm warm conditions. 14 divers performed either one or two (with a 24 h interval) standardized scuba dives to 33 mfw (400 kPa) for 20 min of immersion time at NEMO 33 in Brussels, Belgium. Measurements were performed at 21, 56, 91 and 126 min post-dive: bubbles were counted for all 68 echocardiography recordings and the average over ten consecutive cardiac cycles taken as the bubble score. Significant inter-personal variability was demonstrated despite all divers following the same protocol in controlled pool conditions: in the detection or not of VGE, in the peak VGE score, as well as time to VGE peak. In addition, intra-personal differences in 2/3 of the consecutive day dives were seen (lower VGE counts or faster clearance). Since VGE evolution post-dive varies between people, more work is clearly needed to isolate contributing factors. In this respect, going toward a more continuous evaluation, or developing new means to detect decompression stress markers, may offer the ability to better assess dynamic correlations to other physiological parameters.
29 CFR 1910.410 - Qualifications of dive team.
Code of Federal Regulations, 2010 CFR
2010-07-01
... Labor Regulations Relating to Labor (Continued) OCCUPATIONAL SAFETY AND HEALTH ADMINISTRATION, DEPARTMENT OF LABOR OCCUPATIONAL SAFETY AND HEALTH STANDARDS Commercial Diving Operations Personnel... control the exposure of others to hyperbaric conditions shall be trained in diving-related physics and...
NASA Astrophysics Data System (ADS)
Weiland, C.; Chadwick, W. W.; Hanshumaker, W.; Osis, V.; Hamilton, C.
2002-12-01
We have created a new interactive exhibit in which the user can sit down and simulate that they are making a dive to the seafloor with the remotely operated vehicle (ROV) named ROPOS. The exhibit immerses the user in an interactive experience that is naturally fun but also educational. This new public display is located at the Hatfield Marine Science Visitor Center in Newport, Oregon. The exhibit is designed to look like the real ROPOS control console and includes three video monitors, a PC, a DVD player, an overhead speaker, graphic panels, buttons, lights, dials, and a seat in front of a joystick. The dives are based on real seafloor settings at Axial seamount, an active submarine volcano on the Juan de Fuca Ridge (NE Pacific) that is also the location of a seafloor observatory called NeMO. The user can choose between 1 of 3 different dives sites in the caldera of Axial Volcano. Once a dive is chosen, then the user watches ROPOS being deployed and then arrives into a 3-D computer-generated seafloor environment that is based on the real world but is easier to visualize and navigate. Once on the bottom, the user is placed within a 360 degree panorama and can look in all directions by manipulating the joystick. By clicking on markers embedded in the scene, the user can then either move to other panorama locations via movies that travel through the 3-D virtual environment, or they can play video clips from actual ROPOS dives specifically related to that scene. Audio accompanying the video clips informs the user where they are going or what they are looking at. After the user is finished exploring the dive site they end the dive by leaving the bottom and watching the ROV being recovered onto the ship at the surface. The user can then choose a different dive or make the same dive again. Within the three simulated dives there are a total of 6 arrival and departure movies, 7 seafloor panoramas, 12 travel movies, and 23 ROPOS video clips. The exhibit software was created with Macromedia Director using Apple Quicktime and Quicktime VR. The exhibit is based on the NeMO Explorer web site (http://www.pmel.noaa.gov/vents/nemo/explorer.html).
Time Variation of the Distance Separating Bomb and Dive Bomber Subsequent to Bomb Release
NASA Technical Reports Server (NTRS)
Mathews, Charles W.
1952-01-01
A study has been made of the variation of the distance separating bomb and aircraft with time after release as applied to dive-bombing operations, Separation distances determined from this study are presented in terms of two variables only, dive angle and maximum airplane accelerometer reading; the values of separation distance include the effects of delay in initiation of the pull-out and lag in attainment of the maximum normal acceleration.Contains analysis and calculations of the separation distances between bomb and dive bomber following bomb release, Separation distances as determined by the dive angle and the maximum airplane accelerometer reading are presented in a single chart.
Evaluating the Effects of Stressors on Immune Function during Simulated Dives in Marine Mammals
2014-09-30
Physiology and Stressors on Immune Cell Function in a Deep Diving Monodontid and Three Shallow Diving Phocid Species. PhD Dissertation, University...Research Permit No. 14245). Blood samples were initially processed in the field and shipped back to Mystic Aquarium in LN dry shippers for hormone...of damage from inflammatory processes . Values were returned to control levels suggesting the effects of a dive are not long lasting. That results for
De Brauwer, Maarten; Saunders, Benjamin J; Ambo-Rappe, Rohani; Jompa, Jamaluddin; McIlwain, Jennifer L; Harvey, Euan S
2018-07-15
Scuba diving tourism is a sustainable source of income for many coastal communities, but can have negative environmental impacts if not managed effectively. Diving on soft sediment habitats, typically referred to as 'muck diving', is a growing multi-million dollar industry with a strong focus on photographing cryptobenthic fauna. We assessed how the environmental impacts of scuba divers are affected by the activity they are engaged in while diving and the habitat they dive in. To do this, we observed 66 divers on coral reefs and soft sediment habitats in Indonesia and the Philippines. We found diver activity, specifically interacting with and photographing fauna, causes greater environmental disturbances than effects caused by certification level, gender, dive experience or age. Divers touched the substrate more often while diving on soft sediment habitats than on coral reefs, but this did not result in greater environmental damage on soft sediment sites. Divers had a higher impact on the substrate and touch animals more frequently when observing or photographing cryptobenthic fauna. When using dSLR-cameras, divers spent up to five times longer interacting with fauna. With the unknown, long-term impacts on cryptobenthic fauna or soft sediment habitats, and the increasing popularity of underwater photography, we argue for the introduction of a muck diving code of conduct. Copyright © 2018 Elsevier Ltd. All rights reserved.
Powell, Abby N.; Latty, Christopher J.; Hollmén, Tuula E.; Petersen, Margaret R.; Andrews, Russel D.
2010-01-01
Implanted transmitters have become an important tool for studying the ecology of sea ducks, but their effects remain largely undocumented. To address this, we assessed how abdominally implanted transmitters with percutaneous antennas affect the vertical dive speeds, stroke frequencies, bottom time, and dive duration of captive Common Eiders (Somateria mollissima). To establish baselines, we recorded video of six birds diving 4.9 m prior to surgery, implanted them with 38- to 47-g platform transmitter terminals, and then recorded their diving for 3.5 months after surgery to determine effects. Descent speeds were 16–25% slower and ascent speeds were 17–44% slower after surgery, and both remained below baseline at the end of the study. Dive durations were longer than baseline until day 22. On most days between 15 and 107 days after surgery, foot-stroke frequencies of birds foraging on the bottom were slower. Foot- and wing-stroke frequencies during descent and bottom time did not differ across the time series. If birds that rely on benthic invertebrates for sustenance dive slower and stay submerged longer after being implanted with a satellite transmitter, their foraging energetics may be affected. Researchers considering use of implanted transmitters with percutaneous antennas should be mindful of these effects and the possibility of concomitant alterations in diving behavior, foraging success, and migratory behavior compared to those of unmarked conspecifics.
Jones, D. R.
1973-01-01
1. In the duck systemic arterial baroreceptors which cause bradycardia in response to induced hypertension are located in the walls of the ascending aorta, innervated by the depressor nerves. 2. The location of the baroreceptors was confirmed both histologically and by recording activity from the depressor nerve. Stimulation of the central cut end of a depressor nerve caused transient bradycardia and a fall in blood pressure which was maintained throughout the period of stimulation. 3. Cardiovascular adjustments to submergence of 2 min duration were monitored in intact, sham-operated and denervated ducks. The sham-operated and denervated ducks were used in the experiments some 20-50 days post-operation. The denervations were checked at post-mortem. 4. In the first series of experiments on young ducks mean arterial pressure during a 2 min dive fell by 30% in intact, 17·5% in sham-operated, and 48% in denervated ducks. In all ducks heart rate was reduced by 84-85%. 5. In a second series of experiments on older ducks sciatic artery blood flow was also recorded and mean arterial blood pressure fell by 9·2% in intact and by 53% in denervated animals, although there were no significant differences in heart rate during the 2 min dives. In normal animals sciatic vascular resistance increased after 2 min submergence by 7·86 ± 1·7 times, whereas in denervated ducks it increased by only 2·32 ± 0·5 times. 6. The role of systemic arterial baroreceptors in generation of the cardiovascular responses to submergence in ducks is discussed in terms of the input supplied by the baroreceptors to the central nervous system. ImagesPlate 1 PMID:4764429
Caputo, M P; Alphin, R L; Pritchett, E; Hougentogler, D P; Johnson, A L; Benson, E R; Patil, C
2013-02-01
The mass depopulation of production birds remains an effective means of controlling fast-moving, highly infectious diseases such as avian influenza and virulent Newcastle disease. Water-based fire-fighting foam is a conditionally approved method of depopulating floor-reared gallinaceous poultry such as chickens and turkeys; however, ducks have physiological mechanisms that may make them more resistant to this method of depopulation. The following experiment was designed to assess the physiological responses of White Pekin ducks to nonterminal submersion in water-based foam compared with water. The hypothesis of this experiment was that submersion of ducks in water or water-based foam would trigger the diving reflex and lead to bradycardia. All treatments led to pronounced bradycardia. Heart rate was not significantly different between treatments during the final 30 s of the 60-s treatment period. Heart rate dropped significantly faster for the water dip and foam dip treatments and rose significantly faster than the foam pour treatment after the termination of the 60-s treatment period. Duration of bradycardia approached significance for the foam pour treatment, leading to a longer duration of bradycardia compared with the water pour, water dip, and foam dip treatments. The results of this experiment demonstrated that apnea and bradycardia as a result of the diving reflex can occur as a result of submersion in foam, which may have an impact on the time it takes White Pekin ducks to reach unconsciousness and death during water-based foam depopulation.
Ansai, Satoshi; Hosokawa, Hiroshi; Maegawa, Shingo; Kinoshita, Masato
2016-04-15
Medaka (Oryzias latipes) is a small freshwater teleost that is an emerging model system for neurobehavioral research and toxicological testing. The selective serotonin reuptake inhibitor class of antidepressants such as fluoxetine is one of the widely prescribed drugs, but little is known about the effects of these drugs on medaka behaviors. To assess the behavioral effects of fluoxetine, we chronically administrated fluoxetine to medaka adult fish and analyzed the anxiety-related and social behaviors using five behavioral paradigms (diving, open-field, light-dark transition, mirror-biting, and social interaction) with an automated behavioral testing system. Fish chronically treated with fluoxetine exhibited anxiolytic responses such as an overall increased time spent in the top area in the diving test and an increased time spent in center area in the open-field test. Analysis of socially evoked behavior showed that chronic fluoxetine administration decreased the number of mirror biting times in the mirror-biting test and increased latency to first contact in the social interaction test. Additionally, chronic fluoxetine administration reduced the horizontal locomotor activity in the open-field test but not the vertical activity in the diving test. These investigations are mostly consistent with previous reports in the other teleost species and rodent models. These results indicate that behavioral assessment in medaka adult fish will become useful for screening of effects of pharmaceutical and toxicological compounds in animal behaviors. Copyright © 2016 Elsevier B.V. All rights reserved.
29 CFR 1910.424 - SCUBA diving.
Code of Federal Regulations, 2010 CFR
2010-07-01
... OCCUPATIONAL SAFETY AND HEALTH STANDARDS Commercial Diving Operations Specific Operations Procedures § 1910.424... physically confining spaces. (4) A diver-carried reserve breathing gas supply shall be provided for each... breathing gas supply shall be in the closed position prior to the dive. ...
29 CFR 1926.1086 - Mixed-gas diving.
Code of Federal Regulations, 2010 CFR
2010-07-01
... Regulations Relating to Labor (Continued) OCCUPATIONAL SAFETY AND HEALTH ADMINISTRATION, DEPARTMENT OF LABOR (CONTINUED) SAFETY AND HEALTH REGULATIONS FOR CONSTRUCTION Diving Specific Operations Procedures § 1926.1086 Mixed-gas diving. Note: The requirements applicable to construction work under this section are...
29 CFR 1926.1084 - SCUBA diving.
Code of Federal Regulations, 2010 CFR
2010-07-01
... Relating to Labor (Continued) OCCUPATIONAL SAFETY AND HEALTH ADMINISTRATION, DEPARTMENT OF LABOR (CONTINUED) SAFETY AND HEALTH REGULATIONS FOR CONSTRUCTION Diving Specific Operations Procedures § 1926.1084 SCUBA diving. Note: The requirements applicable to construction work under this section are identical to those...
29 CFR 1926.1081 - Pre-dive procedures.
Code of Federal Regulations, 2010 CFR
2010-07-01
... Regulations Relating to Labor (Continued) OCCUPATIONAL SAFETY AND HEALTH ADMINISTRATION, DEPARTMENT OF LABOR (CONTINUED) SAFETY AND HEALTH REGULATIONS FOR CONSTRUCTION Diving General Operations Procedures § 1926.1081 Pre-dive procedures. Note: The requirements applicable to construction work under this section are...
29 CFR 1926.1083 - Post-dive procedures.
Code of Federal Regulations, 2010 CFR
2010-07-01
... Regulations Relating to Labor (Continued) OCCUPATIONAL SAFETY AND HEALTH ADMINISTRATION, DEPARTMENT OF LABOR (CONTINUED) SAFETY AND HEALTH REGULATIONS FOR CONSTRUCTION Diving General Operations Procedures § 1926.1083 Post-dive procedures. Note: The requirements applicable to construction work under this section are...
29 CFR 1926.1082 - Procedures during dive.
Code of Federal Regulations, 2010 CFR
2010-07-01
... Regulations Relating to Labor (Continued) OCCUPATIONAL SAFETY AND HEALTH ADMINISTRATION, DEPARTMENT OF LABOR (CONTINUED) SAFETY AND HEALTH REGULATIONS FOR CONSTRUCTION Diving General Operations Procedures § 1926.1082 Procedures during dive. Note: The requirements applicable to construction work under this section are...
Wind-Tunnel Investigations of Diving Brakes
NASA Technical Reports Server (NTRS)
Fucha, D.
1942-01-01
Unduly high diving speeds can be effectively controlled by diving brakes but their employment involves at the same time a number of disagreeable features: namely, rotation of zero lift direction, variation of diviving moment, and, the creation of a potent dead air region.
Schorr, Gregory S.; Falcone, Erin A.; Moretti, David J.; Andrews, Russel D.
2014-01-01
Cuvier’s beaked whales (Ziphius cavirostris) are known as extreme divers, though behavioral data from this difficult-to-study species have been limited. They are also the species most often stranded in association with Mid-Frequency Active (MFA) sonar use, a relationship that remains poorly understood. We used satellite-linked tags to record the diving behavior and locations of eight Ziphius off the Southern California coast for periods up to three months. The effort resulted in 3732 hr of dive data with associated regional movements – the first dataset of its kind for any beaked whale – and included dives to 2992 m depth and lasting 137.5 min, both new mammalian dive records. Deep dives had a group mean depth of 1401 m (s.d. = 137.8, n = 1142) and duration of 67.4 min (s.d. = 6.9). The group mean time between deep dives was 102.3 min (s.d. = 30.8, n = 783). While the previously described stereotypic pattern of deep and shallow dives was apparent, there was considerable inter- and intra-individual variability in most parameters. There was significant diel behavioral variation, including increased time near the surface and decreased shallow diving at night. However, maximum depth and the proportion of time spent on deep dives (presumed foraging), varied little from day to night. Surprisingly, tagged whales were present within an MFA sonar training range for 38% of days locations were received, and though comprehensive records of sonar use during tag deployments were not available, we discuss the effects frequent acoustic disturbance may have had on the observed behaviors. These data better characterize the true behavioral range of this species, and suggest caution should be exercised when drawing conclusions about behavior using short-term datasets. PMID:24670984
Optimal diving behaviour and respiratory gas exchange in birds.
Halsey, Lewis G; Butler, Patrick J
2006-11-01
This review discusses the advancements in our understanding of the physiology and behaviour of avian diving that have been underpinned by optimal foraging theory and the testing of optimal models. To maximise their foraging efficiency during foraging periods, diving birds must balance numerous factors that are directly or indirectly related to the replenishment of the oxygen stores and the removal of excess carbon dioxide. These include (1) the time spent underwater (which diminishes the oxygen supply, increases carbon dioxide levels and may even include a build up of lactate due to anaerobic metabolism), (2) the time spent at the surface recovering from the previous dive and preparing for the next (including reloading their oxygen supply, decreasing their carbon dioxide levels and possibly also metabolising lactate) and (3) the trade-off between maximising oxygen reserves for consumption underwater by taking in more air to the respiratory system, and minimising the energy costs of positive buoyancy caused by this air, to maximise the time available underwater to forage. Due to its importance in avian diving, replenishment of the oxygen stores has become integral to models of optimal diving, which predict the time budgeting of animals foraging underwater. While many of these models have been examined qualitatively, such tests of predictive trends appear fallible and only quantifiable support affords strong evidence of their predictive value. This review describes how the quantification of certain optimal diving models, using tufted ducks, indeed demonstrates some predictive success. This suggests that replenishment of the oxygen stores and removal of excess carbon dioxide have significant influences on the duration of the surface period between dives. Nevertheless, present models are too simplistic to be robust predictors of diving behaviour for individual animals and it is proposed that they require refinement through the incorporation of other variables that also influence diving behaviour such as, perhaps, prey density and predator avoidance.
New Polish occupational health and safety regulations for underwater works.
Kot, Jacek; Sićko, Zdzisław
2007-01-01
In Poland, the new regulation of the Ministry of Health on Occupational Health for Underwater Works (dated 2007) pursuant to the Act on Underwater Works (dated 2003) has just been published. It is dedicated for commercial, non-military purposes. It defines health requirements for commercial divers and candidates for divers, medical assessment guide with a list of specific medical tests done on initial and periodical medical examination in order for a diver or a candidate for diver to be recognised fit for work, health surveillance during diving operations, compression and decompression procedures, list of content for medical equipment to be present at any diving place, formal qualifications for physicians conducting medical assessment of divers, requirements for certifications confirming the medical status of divers and candidates for divers. Decompression tables cover divings up to 120 meters of depth using compressed air, oxygen, nitrox and heliox as breathing mixtures. There are also decompression tables for repetitive diving, altitude diving and diving in the high-density waters (mud diving). It this paper, general description of health requirements for divers, as well as decompression tables that are included in the new Regulation on Occupational Health for Underwater Works are presented.
Pressure equilibration in the penguin middle ear.
Sadé, Jacob; Handrich, Yves; Bernheim, Joelle; Cohen, David
2008-01-01
King penguins have a venous structure in the form of a corpus cavernosum (CC) in their middle ear (ME) submucosa. The CC may be viewed as a special organelle that can change ME volume for pressure equilibration during deep-sea diving it is a pressure regulating organelle (PRO). A similar CC and muscles also surround the external ear (EE) and may constrict it, isolating the tympanic membrane from the outside. A CC was previously found also in the ME of marine diving mammals and can be expected to exist in other deep diving animals, such as marine turtles. Marine animals require equalization of middle ear (ME) pressure when diving hundreds or thousands of meters to catch prey. We investigated what mechanism enables king penguins to protect their ME when they dive to great depths. Biopsies and serial sections of the ME and the EE of the deep diving king penguin (Aptenodytes patagonicus) were examined microscopically. It was demonstrated that the penguin ME has an extensive network of small and large submucosal venous sinuses. This venous formation, a corpus cavernosum, can expand and potentially 'flood' the ME almost completely on diving, thus elevating ME pressure and reducing the ME space. The EE has a similar protective mechanism.
Schulze, W; Schul, J
2001-02-01
The responses of female Tettigonia viridissima to simulated bat echolocation calls were examined during tethered flight. The insects responded with three distinct behaviours, which occurred at graded stimulus intensities. At low intensities (threshold 54 dB SPL), T. viridissima responded by steering away from the sound source (negative phonotaxis). At intensities approximately 10 dB higher, beating of the hindwing was interrupted, although the insect remained in the flight posture. A diving response (cessation of the wingbeat, closure of the forewings and alignment of the legs against the body) occurred with a threshold of 76 dB SPL. Considering these thresholds, we estimate that the diving response occurs at approximately the sound amplitude at which many aerial-hawking bats first receive echoes from the insect. The other behaviours probably occur before the bat detects the insect and should therefore be interpreted as early avoidance behaviours. The repertoire of startle responses in T. viridissima, with directional and non-directional components, is similar to those of crickets and moths, but quite different from those described for another bushcricket (Neoconocephalus ensiger), which shows only a non-directional response. This supports the conclusion that bat-evasive behaviours are not conserved within the Tettigoniidae, but instead are shaped by the ecological constraints of the insects.
Deep-Diving California Sea Lions: Are They Pushing Their Physiological Limit?
2014-09-30
were 55 + 8, 51 + 6, and 40 + bpm. As illustrated in Figs. 1 and 2, the heart rate profile was characterized by rapid development of a bradycardia ...slow heart rate), and a gradual increase in heart rate during the bottom phase of the dive and during ascent. The degree of bradycardia was more...deeper dives of longer duration, and the degree of bradycardia increases with maximum depth and duration of dives. 4 0 20 40 60 80 100 120 140 0
A Simple Probabilistic Model for Estimating the Risk of Standard Air Dives
2004-12-01
Decompression Models Table Al. Decompression Table Based on the StandAir Model and Comparison with the VVaI-1 8 Algorithm. A-l-A-4 Table A2. The VVaI-1 8...cannot be as strong as might be desired - especially for dives with long TDTs. Comparisons of the positions of the dive-outcome symbols with the... comparisons for several depth/bottom-time combinations. The three left-hand panels, for dives with short bottom times, show that the crossover point
NASA Astrophysics Data System (ADS)
Clanet, Christophe; Guillet, Thibault; Coux, Martin; Quéré, David
2017-11-01
Many seabirds (gannets, pelicans, gulls, albatrosses) dive into water at high speeds (25 m/s) in order to capture underwater preys. Diving depths of 20 body lengths are reported in the literature. This value is much larger than the one achieved by men, which is typically of the order of 3. We study this difference by comparing the vertical impact of slender vs bluff bodies. We quantify the influence of wetting and of the geometry on the trajectory and discuss the different laws that govern the diving depth.
Deep-Diving California Sea Lions: Are They Pushing Their Physiological Limit
2015-09-30
resting heart rate (70 bpm ) (Ponganis et al. 1997) is reached, and e) duration of and heart rate during the ascent tachycardia. If possible, heart rate...Resting heart rates were 54 + 6 beats min-1 ( bpm ), and in dives of 1-3 min, 3-5 min, and > 5 min, dive heart rates (number of beats/dive duration...were 55 + 8, 51 + 6, and 40 + bpm . As illustrated in Figs. 1 and 2, the heart rate profile was characterized by rapid development of a bradycardia
2015-05-28
true, trimix would be an attractive alternative to heliox for U. S. Navy MK 16 MOD 1 underwater breathing apparatus (UBA) diving. However, there is no...for the semi-closed circuit Canadian Underwater Mine-countermeasures Apparatus (CUMA) that are shorter than...diving15-17 and by the U.S. Navy for the MK 6 semi- closed circuit underwater breathing apparatus (UBA);18,19 each of these programs tested trimix schedules
NASA Technical Reports Server (NTRS)
Lattanzi, Bernardino; Bellante, Erno
1949-01-01
The present report is concerned with a series of tests on a model airplane fitted with four types of dive flaps of various shapes, positions, and incidence located near the leading edge of the wing (from 5 to 20 percent of the wing chord). Tests were also made on a stub airfoil fitted with a ventral dive (located at 8 percent of the wing chord). The hinge moments of the dive flaps were measured.
Germonpré, Peter; Balestra, Costantino; Hemelryck, Walter; Buzzacott, Peter; Lafère, Pierre
2017-05-01
Divers try to limit risks associated with their sport, for instance by breathing enriched air nitrox (EANx) instead of air. This double blinded, randomized trial was designed to see if the use of EANx could effectively improve cognitive performance while diving. Eight volunteers performed two no-decompression dry dives breathing air or EANx for 20 min at 0.4 MPa. Cognitive functions were assessed with a computerized test battery, including MathProc and Ptrail. Measurements were taken before the dive, upon arrival and after 15 min at depth, upon surfacing, and at 30 min postdive. After each dive subjects were asked to identify the gas they had just breathed. Identification of the breathing gas was not possible on subjective assessment alone, while cognitive assessments showed significantly better performance while breathing EANx. Before the dives, breathing air, mean time to complete the task was 1795 ms for MathProc and 1905 ms for Ptrail. When arriving at depth MathProc took 1616 ms on air and 1523 ms on EANx, and Ptrail took 1318 ms on air and and 1356 ms on EANx, followed 15 min later by significant performance inhibition while breathing air during the ascent and the postdive phase, supporting the concept of late dive/postdive impairment. The results suggest that EANx could protect against decreased neuro-cognitive performance induced by inert gas narcosis. It was not possible for blinded divers to identify which gas they were breathing and differences in postdive fatigue between air and EANx diving deserve further investigation.Germonpré P, Balestra C, Hemelryck W, Buzzacott P, Lafère P. Objective vs. subjective evaluation of cognitive performance during 0.4-MPa dives breathing air or nitrox. Aerosp Med Hum Perform. 2017; 88(5):469-475.
Preconditioning to Reduce Decompression Stress in Scuba Divers.
Germonpré, Peter; Balestra, Costantino
2017-02-01
Using ultrasound imaging, vascular gas emboli (VGE) are observed after asymptomatic scuba dives and are considered a key element in the potential development of decompression sickness (DCS). Diving is also accompanied with vascular dysfunction, as measured by flow-mediated dilation (FMD). Previous studies showed significant intersubject variability to VGE for the same diving exposure and demonstrated that VGE can be reduced with even a single pre-dive intervention. Several preconditioning methods have been reported recently, seemingly acting either on VGE quantity or on endothelial inflammatory markers. Nine male divers who consistently showed VGE postdive performed a standardized deep pool dive (33 m/108 ft, 20 min in 33°C water temperature) to investigate the effect of three different preconditioning interventions: heat exposure (a 30-min session of dry infrared sauna), whole-body vibration (a 30-min session on a vibration mattress), and dark chocolate ingestion (30 g of chocolate containing 86% cocoa). Dives were made one day per week and interventions were administered in a randomized order. These interventions were shown to selectively reduce VGE, FMD, or both compared to control dives. Vibration had an effect on VGE (39.54%, SEM 16.3%) but not on FMD postdive. Sauna had effects on both parameters (VGE: 26.64%, SEM 10.4%; FMD: 102.7%, SEM 2.1%), whereas chocolate only improved FMD (102.5%, SEM 1.7%). This experiment, which had the same subjects perform all control and preconditioning dives in wet but completely standardized diving conditions, demonstrates that endothelial dysfunction appears to not be solely related to VGE.Germonpré P, Balestra C. Preconditioning to reduce decompression stress in scuba divers. Aerosp Med Hum Perform. 2017; 88(2):114-120.
Improved pulmonary function in working divers breathing nitrox at shallow depths
NASA Technical Reports Server (NTRS)
Fitzpatrick, Daniel T.; Conkin, Johnny
2003-01-01
INTRODUCTION: There is limited data about the long-term pulmonary effects of nitrox use in divers at shallow depths. This study examined changes in pulmonary function in a cohort of working divers breathing a 46% oxygen enriched mixture while diving at depths less than 12 m. METHODS: A total of 43 working divers from the Neutral Buoyancy Laboratory (NBL), NASA-Johnson Space Center completed a questionnaire providing information on diving history prior to NBL employment, diving history outside the NBL since employment, and smoking history. Cumulative dive hours were obtained from the NBL dive-time database. Medical records were reviewed to obtain the diver's height, weight, and pulmonary function measurements from initial pre-dive, first year and third year annual medical examinations. RESULTS: The initial forced vital capacity (FVC) and forced expiratory volume in 1 s (FEV1) were greater than predicted, 104% and 102%, respectively. After 3 yr of diving at the NBL, both the FVC and FEV1 showed a significant (p < 0.01) increase of 6.3% and 5.5%, respectively. There were no significant changes in peak expiratory flow (PEF), forced mid-expiratory flow rate (FEF(25-75%)), and forced expiratory flow rates at 25%, 50%, and 75% of FVC expired (FEF25%, FEF50%, FEF75%). Cumulative NBL dive hours was the only contributing variable found to be significantly associated with both FVC and FEV1 at 1 and 3 yr. CONCLUSIONS: NBL divers initially belong to a select group with larger than predicted lung volumes. Regular diving with nitrox at shallow depths over a 3-yr period did not impair pulmonary function. Improvements in FVC and FEV1 were primarily due to a training effect.
Wienke, B R; O'Leary, T R
2008-05-01
Linking model and data, we detail the LANL diving reduced gradient bubble model (RGBM), dynamical principles, and correlation with data in the LANL Data Bank. Table, profile, and meter risks are obtained from likelihood analysis and quoted for air, nitrox, helitrox no-decompression time limits, repetitive dive tables, and selected mixed gas and repetitive profiles. Application analyses include the EXPLORER decompression meter algorithm, NAUI tables, University of Wisconsin Seafood Diver tables, comparative NAUI, PADI, Oceanic NDLs and repetitive dives, comparative nitrogen and helium mixed gas risks, USS Perry deep rebreather (RB) exploration dive,world record open circuit (OC) dive, and Woodville Karst Plain Project (WKPP) extreme cave exploration profiles. The algorithm has seen extensive and utilitarian application in mixed gas diving, both in recreational and technical sectors, and forms the bases forreleased tables and decompression meters used by scientific, commercial, and research divers. The LANL Data Bank is described, and the methods used to deduce risk are detailed. Risk functions for dissolved gas and bubbles are summarized. Parameters that can be used to estimate profile risk are tallied. To fit data, a modified Levenberg-Marquardt routine is employed with L2 error norm. Appendices sketch the numerical methods, and list reports from field testing for (real) mixed gas diving. A Monte Carlo-like sampling scheme for fast numerical analysis of the data is also detailed, as a coupled variance reduction technique and additional check on the canonical approach to estimating diving risk. The method suggests alternatives to the canonical approach. This work represents a first time correlation effort linking a dynamical bubble model with deep stop data. Supercomputing resources are requisite to connect model and data in application.
29 CFR 1926.1085 - Surface-supplied air diving.
Code of Federal Regulations, 2013 CFR
2013-07-01
... 29 Labor 8 2013-07-01 2013-07-01 false Surface-supplied air diving. 1926.1085 Section 1926.1085 Labor Regulations Relating to Labor (Continued) OCCUPATIONAL SAFETY AND HEALTH ADMINISTRATION... Procedures § 1926.1085 Surface-supplied air diving. Note: The requirements applicable to construction work...
29 CFR 1926.1086 - Mixed-gas diving.
Code of Federal Regulations, 2011 CFR
2011-07-01
... 29 Labor 8 2011-07-01 2011-07-01 false Mixed-gas diving. 1926.1086 Section 1926.1086 Labor Regulations Relating to Labor (Continued) OCCUPATIONAL SAFETY AND HEALTH ADMINISTRATION, DEPARTMENT OF LABOR... Mixed-gas diving. Note: The requirements applicable to construction work under this section are...
29 CFR 1926.1086 - Mixed-gas diving.
Code of Federal Regulations, 2013 CFR
2013-07-01
... 29 Labor 8 2013-07-01 2013-07-01 false Mixed-gas diving. 1926.1086 Section 1926.1086 Labor Regulations Relating to Labor (Continued) OCCUPATIONAL SAFETY AND HEALTH ADMINISTRATION, DEPARTMENT OF LABOR... Mixed-gas diving. Note: The requirements applicable to construction work under this section are...
29 CFR 1926.1086 - Mixed-gas diving.
Code of Federal Regulations, 2014 CFR
2014-07-01
... 29 Labor 8 2014-07-01 2014-07-01 false Mixed-gas diving. 1926.1086 Section 1926.1086 Labor Regulations Relating to Labor (Continued) OCCUPATIONAL SAFETY AND HEALTH ADMINISTRATION, DEPARTMENT OF LABOR... Mixed-gas diving. Note: The requirements applicable to construction work under this section are...
29 CFR 1926.1086 - Mixed-gas diving.
Code of Federal Regulations, 2012 CFR
2012-07-01
... 29 Labor 8 2012-07-01 2012-07-01 false Mixed-gas diving. 1926.1086 Section 1926.1086 Labor Regulations Relating to Labor (Continued) OCCUPATIONAL SAFETY AND HEALTH ADMINISTRATION, DEPARTMENT OF LABOR... Mixed-gas diving. Note: The requirements applicable to construction work under this section are...
29 CFR 1926.1085 - Surface-supplied air diving.
Code of Federal Regulations, 2010 CFR
2010-07-01
... 29 Labor 8 2010-07-01 2010-07-01 false Surface-supplied air diving. 1926.1085 Section 1926.1085 Labor Regulations Relating to Labor (Continued) OCCUPATIONAL SAFETY AND HEALTH ADMINISTRATION... Procedures § 1926.1085 Surface-supplied air diving. Note: The requirements applicable to construction work...
29 CFR 1926.1076 - Qualifications of dive team.
Code of Federal Regulations, 2011 CFR
2011-07-01
... 29 Labor 8 2011-07-01 2011-07-01 false Qualifications of dive team. 1926.1076 Section 1926.1076 Labor Regulations Relating to Labor (Continued) OCCUPATIONAL SAFETY AND HEALTH ADMINISTRATION... § 1926.1076 Qualifications of dive team. Note: The requirements applicable to construction work under...
29 CFR 1926.1076 - Qualifications of dive team.
Code of Federal Regulations, 2010 CFR
2010-07-01
... 29 Labor 8 2010-07-01 2010-07-01 false Qualifications of dive team. 1926.1076 Section 1926.1076 Labor Regulations Relating to Labor (Continued) OCCUPATIONAL SAFETY AND HEALTH ADMINISTRATION... § 1926.1076 Qualifications of dive team. Note: The requirements applicable to construction work under...
29 CFR 1926.1076 - Qualifications of dive team.
Code of Federal Regulations, 2013 CFR
2013-07-01
... 29 Labor 8 2013-07-01 2013-07-01 false Qualifications of dive team. 1926.1076 Section 1926.1076 Labor Regulations Relating to Labor (Continued) OCCUPATIONAL SAFETY AND HEALTH ADMINISTRATION... § 1926.1076 Qualifications of dive team. Note: The requirements applicable to construction work under...
29 CFR 1926.1076 - Qualifications of dive team.
Code of Federal Regulations, 2014 CFR
2014-07-01
... 29 Labor 8 2014-07-01 2014-07-01 false Qualifications of dive team. 1926.1076 Section 1926.1076 Labor Regulations Relating to Labor (Continued) OCCUPATIONAL SAFETY AND HEALTH ADMINISTRATION... § 1926.1076 Qualifications of dive team. Note: The requirements applicable to construction work under...
29 CFR 1926.1076 - Qualifications of dive team.
Code of Federal Regulations, 2012 CFR
2012-07-01
... 29 Labor 8 2012-07-01 2012-07-01 false Qualifications of dive team. 1926.1076 Section 1926.1076 Labor Regulations Relating to Labor (Continued) OCCUPATIONAL SAFETY AND HEALTH ADMINISTRATION... § 1926.1076 Qualifications of dive team. Note: The requirements applicable to construction work under...
Wind and water tunnel testing of a morphing aquatic micro air vehicle.
Siddall, Robert; Ortega Ancel, Alejandro; Kovač, Mirko
2017-02-06
Aerial robots capable of locomotion in both air and water would enable novel mission profiles in complex environments, such as water sampling after floods or underwater structural inspections. The design of such a vehicle is challenging because it implies significant propulsive and structural design trade-offs for operation in both fluids. In this paper, we present a unique Aquatic Micro Air Vehicle (AquaMAV), which uses a reconfigurable wing to dive into the water from flight, inspired by the plunge diving strategy of water diving birds in the family Sulidae . The vehicle's performance is investigated in wind and water tunnel experiments, from which we develop a planar trajectory model. This model is used to predict the dive behaviour of the AquaMAV, and investigate the efficacy of passive dives initiated by wing folding as a means of water entry. The paper also includes first field tests of the AquaMAV prototype where the folding wings are used to initiate a plunge dive.
NASA Astrophysics Data System (ADS)
Wijaya, Putranto; Putra, Tri; Hidayat, Fatra; Levraeni, Chandra; Rizmaadi, Mada; Ambariyanto, Ambariyanto
2018-02-01
Indonesian government currently has policies to improve the performance of the tourism sector, including marine tourism. One of the attractions of marine tourism is the coral ecosystem especially through scuba diving activities. The purpose of this study was to determine the suitability of the coral ecosystem on Saebus Island, East Java, to find appropriate locations for scuba diving activities. Purposive samplings were done around the island to determine four stations which will be assessed through suitability analysis. Tourism Suitability Index was used to assess all stations for scuba diving activities. The result showed that all four stations were categorized as very suitable with the score: 85%, 85%, 85% and 83%, respectively. Several aspects that need to be improved and anticipated for diving at all stations are coral coverage and water current. These results suggest that there are several spots around Saebus Island that are suitable for diving site, and can be promoted as marine tourism destination.
Provisional report on diving-related fatalities in Australian waters 2003.
Walker, Douglas; Lippmann, John
2009-03-01
An individual case review of the diving-related deaths that were reported to have occurred in Australia in 2003 was conducted as part of the combined Project Stickybeak/DAN Asia-Pacific dive fatality reporting project. The case studies were compiled using reports from witnesses, the police and coroners. In each case, the particular circumstances of the accident are provided, as well as details from the post mortem examination, where available. In total there were 22 reported fatalities, 18 men and four women. Twelve deaths occurred while snorkelling and/or breath-hold diving, nine while scuba diving and one while using surface-supply breathing apparatus. Cardiac-related issues were thought to have contributed to the deaths of six snorkel divers (50%) and four scuba divers (44%) in this series. There were three deaths in breath-hold divers likely to have been associated with apnoeic hypoxia blackout. Inexperience, time away from diving and lack of common sense were features in several scuba deaths.
Provisional report on diving-related fatalities in Australian waters 2005.
Walker, Douglas; Lippmann, John; Lawrence, Chris; Fock, Andrew; Wodak, Thomas; Jamieson, Scott
2010-09-01
An individual case review of diving-related deaths reported as occurring in Australia in 2005 was conducted as part of the DAN Asia-Pacific dive fatality reporting project. The case studies were compiled using reports from witnesses, the police and coroners. In each case, the particular circumstances of the accident and details from the post-mortem examination, where available, are provided. In total, there were 24 reported fatalities, comprising four females and 20 males. Fourteen deaths occurred while snorkelling and/or breath-hold diving, nine while scuba diving, and one while using surface-supply breathing apparatus. Four deaths from large marine animal attacks were recorded. Once again, cardiac-related issues were thought to have contributed to some deaths: five snorkel divers and at least two but possibly up to four scuba divers. Three of the deaths in breath-hold divers were likely to have been associated with apnoeic hypoxia blackout. Pre-existing medical conditions, trauma from marine creatures and snorkelling or diving alone were features in several deaths in this series.
Wind and water tunnel testing of a morphing aquatic micro air vehicle
Ortega Ancel, Alejandro; Kovač, Mirko
2017-01-01
Aerial robots capable of locomotion in both air and water would enable novel mission profiles in complex environments, such as water sampling after floods or underwater structural inspections. The design of such a vehicle is challenging because it implies significant propulsive and structural design trade-offs for operation in both fluids. In this paper, we present a unique Aquatic Micro Air Vehicle (AquaMAV), which uses a reconfigurable wing to dive into the water from flight, inspired by the plunge diving strategy of water diving birds in the family Sulidae. The vehicle's performance is investigated in wind and water tunnel experiments, from which we develop a planar trajectory model. This model is used to predict the dive behaviour of the AquaMAV, and investigate the efficacy of passive dives initiated by wing folding as a means of water entry. The paper also includes first field tests of the AquaMAV prototype where the folding wings are used to initiate a plunge dive. PMID:28163877
Foraging depths of sea otters and implications to coastal marine communities
Bodkin, James L.; Esslinger, George G.; Monson, Daniel H.
2004-01-01
We visually observed 1,251 dives, of 14 sea otters instrumented with TDRs in southeast Alaska, and used attribute values from observed dives to classify 180,848 recorded dives as foraging (0.64), or traveling (0.36). Foraging dives were significantly deeper, with longer durations, bottom times, and postdive surface intervals, and greater descent and ascent rates, compared to traveling dives. Most foraging occurred in depths between 2 and 30 m (0.84), although 0.16 of all foraging was between 30 and 100 m. Nine animals, including all five males, demonstrated bimodal patterns in foraging depths, with peaks between 5 and 15 m and 30 and 60 m, whereas five of nine females foraged at an average depth of 10 m. Mean shallow foraging depth was 8 m, and mean deep foraging depth was 44 m. Maximum foraging depths averaged 61 m (54 and 82 for females and males, respectively) and ranged from 35 to 100 m. Female sea otters dove to depths ≤20 m on 0.85 of their foraging dives while male sea otters dove to depths ≥45 m on 0.50 of their foraging dives. Less than 0.02 of all foraging dives were >55 m, suggesting that effects of sea otter foraging on nearshore marine communities should diminish at greater depths. However, recolonization of vacant habitat by high densities of adult male sea otters may result in initial reductions of some prey species at depths >55 m.
Deep Diving Cetacean Behavioral Response Study MED 09
2009-09-30
distribution may be affected by anthropogenic noise. The role of the SPAWAR Systems Center (SSC) Pacific team was to intergrate the interdisiplinary...over 100 hours of data was collected on production sonobuoys for post test ambient noise data. CTD casts were taken at 56 sites, collecting
Nest visits and capture events affect breeding success of Yellow-billed and Pacific loons
Uher-Koch, Brian D.; Schmutz, Joel A.; Wright, Kenneth G.
2015-01-01
Accurate estimates of breeding success are essential for understanding population dynamics and for managing populations. Unfortunately, research activities to collect these data can negatively impact the breeding success of the study species and bias estimates of breeding success. Despite the potential for negative impacts, few studies have documented the effect of capturing incubating adults on nest survival or compared nest survival following different capture methods. In this study we evaluate the impacts of investigator disturbance associated with captures and nest visits on nest survival of Yellow-billed Loons (Gavia adamsii) and Pacific Loons (Gavia pacifica) in the National Petroleum Reserve-Alaska (NPR-A), an area of conservation concern, in 2011–2013. In an effort to reduce capture-related nest failures, we developed a new suspended dive net technique to catch territorial aquatic birds while off their nests. We then compared nest survival following suspended dive net captures to bow-net trap captures of breeding adult loons. Daily nest survival following bow-net trap or suspended dive net capture was about 30% lower than when adults were not captured. The effect of captures on nest survival was similar between bow-net trap and suspended dive net capture methods. Nest visits without captures also negatively impacted nest survival, although less than captures. If not accounted for, nest visitation biased daily survival rates of nests downward 6%. Effects of investigator disturbance did not differ by species or between years. Our results suggest that any source of disturbance that displaces incubating adult loons could potentially reduce nest survival. To maximize breeding success, human disturbance factors should be limited near loon nests.
Association of microparticles and neutrophil activation with decompression sickness.
Thom, Stephen R; Bennett, Michael; Banham, Neil D; Chin, Walter; Blake, Denise F; Rosen, Anders; Pollock, Neal W; Madden, Dennis; Barak, Otto; Marroni, Alessandro; Balestra, Costantino; Germonpre, Peter; Pieri, Massimo; Cialoni, Danilo; Le, Phi-Nga Jeannie; Logue, Christopher; Lambert, David; Hardy, Kevin R; Sward, Douglas; Yang, Ming; Bhopale, Veena B; Dujic, Zeljko
2015-09-01
Decompression sickness (DCS) is a systemic disorder, assumed due to gas bubbles, but additional factors are likely to play a role. Circulating microparticles (MPs)--vesicular structures with diameters of 0.1-1.0 μm--have been implicated, but data in human divers have been lacking. We hypothesized that the number of blood-borne, Annexin V-positive MPs and neutrophil activation, assessed as surface MPO staining, would differ between self-contained underwater breathing-apparatus divers suffering from DCS vs. asymptomatic divers. Blood was analyzed from 280 divers who had been exposed to maximum depths from 7 to 105 meters; 185 were control/asymptomatic divers, and 90 were diagnosed with DCS. Elevations of MPs and neutrophil activation occurred in all divers but normalized within 24 h in those who were asymptomatic. MPs, bearing the following proteins: CD66b, CD41, CD31, CD142, CD235, and von Willebrand factor, were between 2.4- and 11.7-fold higher in blood from divers with DCS vs. asymptomatic divers, matched for time of sample acquisition, maximum diving depth, and breathing gas. Multiple logistic regression analysis documented significant associations (P < 0.001) between DCS and MPs and for neutrophil MPO staining. Effect estimates were not altered by gender, body mass index, use of nonsteroidal anti-inflammatory agents, or emergency oxygen treatment and were modestly influenced by divers' age, choice of breathing gas during diving, maximum diving depth, and whether repetitive diving had been performed. There were no significant associations between DCS and number of MPs without surface proteins listed above. We conclude that MP production and neutrophil activation exhibit strong associations with DCS. Copyright © 2015 the American Physiological Society.
The hydrothermal exploration system on the 'Qianlong2' AUV
NASA Astrophysics Data System (ADS)
Tao, W.; Tao, C.; Jinhui, Z.; Cai, L.; Guoyin, Z.
2016-12-01
ABSTRACT: Qianlong2, is a fully Autonomous Underwater Vehicle (AUV) designed for submarine resources research, especially for polymetallic sulphides, and the survey depths of is up to 4500 m. Qianlong2 had successfully explored hydrothermal vent field on the Southwest Indian Ridge (SWIR), and collected conductance, temperature and depth (CTD), turbidity, and Oxidation-Reduction Potential (ORP) data. It also had mapped precise topography by high resolution side scan sonar (HRBSSS) during every dive; and obtained photographs of sulfide deposits during some dives. Here, we detailedly described the implementation of investigation, data administration, and fast mapping of hydrothermal exploration system by Qianlong2. Giving a description of how to remove the platform magnetic interference by using magnetic data during Qianlong2 spin. Based on comprehensive hydrochemical anomalies, we get a rapid method for finding the localization of hydrothermal vents. Taking one dive as an example, we systemically showed the process about how to analyse hydrothermal survey data and acquire the location results of hydrothermal vents. Considering that this method is effective and can be used in other deep-submergence assets such as human occupied vehicles (HOVs) and remotely operated vehicles (ROVs) during further studies. Finally, we discussed how to promote and optimize the installation and application of those sensors and how to improve Qianlong2's autonomy of investigation.
Castelblanco-Martínez, Delma Nataly; Morales-Vela, Benjamin; Slone, Daniel H.; Padilla-Saldívar, Janneth Adriana; Reid, James P.; Hernández-Arana, Héctor Abuid
2015-01-01
Diving or respiratory behavior in aquatic mammals can be used as an indicator of physiological activity and consequently, to infer behavioral patterns. Five Antillean manatees, Trichechus manatus manatus, were captured in Chetumal Bay and tagged with GPS tracking devices. The radios were equipped with a micropower saltwater sensor (SWS), which records the times when the tag assembly was submerged. The information was analyzed to establish individual fine-scale behaviors. For each fix, we established the following variables: distance (D), sampling interval (T), movement rate (D/T), number of dives (N), and total diving duration (TDD). We used logic criteria and simple scatterplots to distinguish between behavioral categories: ‘Travelling’ (D/T ≥ 3 km/h), ‘Surface’ (↓TDD, ↓N), ‘Bottom feeding’ (↑TDD, ↑N) and ‘Bottom resting’ (↑TDD, ↓N). Habitat categories were qualitatively assigned: Lagoon, Channels, Caye shore, City shore, Channel edge, and Open areas. The instrumented individuals displayed a daily rhythm of bottom activities, with surfacing activities more frequent during the night and early in the morning. More investigation into those cycles and other individual fine-scale behaviors related to their proximity to concentrations of human activity would be informative
Scuba Diving and Kinesiology: Development of an Academic Program
ERIC Educational Resources Information Center
Kovacs, Christopher R.; Walter, Daniel
2015-01-01
The use of scuba diving as a recreational activity within traditional university instructional programs has been well established. Departments focusing on kinesiology, physical education, or exercise science have often provided scuba diving lessons as part of their activity-based course offerings. However, few departments have developed an…
Federal Register 2010, 2011, 2012, 2013, 2014
2010-06-24
... Conservation Seat and Diving Operations Seat for the Flower Garden Banks National Marine Sanctuary Advisory... Flower Garden Banks National Marine Sanctuary Advisory Council: Conservation and Diving Operations... Jennifer Morgan, NOAA--Flower Garden Banks National Marine Sanctuary, 4700 Avenue U, Bldg. 216, Galveston...
46 CFR 197.450 - Breathing gas tests.
Code of Federal Regulations, 2013 CFR
2013-10-01
... 46 Shipping 7 2013-10-01 2013-10-01 false Breathing gas tests. 197.450 Section 197.450 Shipping COAST GUARD, DEPARTMENT OF HOMELAND SECURITY (CONTINUED) MARINE OCCUPATIONAL SAFETY AND HEALTH STANDARDS GENERAL PROVISIONS Commercial Diving Operations Periodic Tests and Inspections of Diving Equipment § 197.450 Breathing gas tests. The diving...
Spinal Injuries in the Aquatics Environment, Part I: Prevention.
ERIC Educational Resources Information Center
Dworkin, Gerald M.
1987-01-01
Water-related activities are the number one cause of spinal cord injuries resulting from sports and recreation activities. This article discusses principles of safe diving; principles of safe water sliding; ways to reduce springboard diving accidents; factors contributing to springboard diving accidents; and safety recommendations for open water…
46 CFR 197.346 - Diver's equipment.
Code of Federal Regulations, 2012 CFR
2012-10-01
... breathing gas supply with a cylinder pressure gage readable by the diver during the dive; and (ii) A diver... (8) A depth gage. (b) Each diver using a heavyweight diving outfit must— (1) Have a helmet group consisting of helmet, breastplate, and associated valves and connections; (2) Have a diving dress group...
46 CFR 197.346 - Diver's equipment.
Code of Federal Regulations, 2013 CFR
2013-10-01
... breathing gas supply with a cylinder pressure gage readable by the diver during the dive; and (ii) A diver... (8) A depth gage. (b) Each diver using a heavyweight diving outfit must— (1) Have a helmet group consisting of helmet, breastplate, and associated valves and connections; (2) Have a diving dress group...
Insulin-dependent diabetes mellitus and recreational scuba diving in Australia.
Johnson, Rebecca
2016-09-01
Dive medicine bodies worldwide recognise that, with comprehensive screening and careful management, people with insulin-dependent diabetes (IDDM) can dive safely. Despite this, people with IDDM in Australia are generally denied access to dive training, an out-dated status quo that is not acceptable to the Australian diabetes community. This paper reflects upon the important advocacy work that has been done to progress this issue, and what is still required to open up access and bring Australia into line with more flexible and supportive international standards.
Evaluating the risk of decompression sickness for a yo-yo dive using a rat model.
Ofir, Dror; Yanir, Yoav; Abramovich, Amir; Bar, Ronen; Arieli, Yehuda
2016-01-01
The frequent ascents made during yo-yo diving may contribute to gas bubble clearance but paradoxically may also increase the risk of central nervous system decompression illness (DCI). We evaluated the risk of DCI due to yo-yo dives with very short surface intervals, using a controlled animal model. Dives were conducted on air to a depth of 90 meters (10 atmospheres absolute) for 32 minutes of bottom time, at a descent/ascent rate of 10 meters/ minute. Sprague-Dawley rats weighing ~ 300 grams were divided randomly into three groups. Group A performed a square dive protocol without any surface intervals, Group B conducted a protocol that included two surface intervals during the dive, and Group C performed a protocol with three surface intervals. Ascent/descent rate for surface intervals, each lasting one minute, was also 10 meters/minute. Manifestations of DCI were observed in 13 of 16 animals in Group A (81.3%), six of 12 in Group B (58.3%), and two of 12 in Group C (16.7%). Mortality rates were similar in all groups. Surface intervals during dives breathing air significantly reduced DCI risk in the rat. Further studies are required using a larger animal model to reinforce the results of the present investigation.
Ecological carrying capacity assessment of diving site: A case study of Mabul Island, Malaysia.
Zhang, Li-Ye; Chung, Shan-Shan; Qiu, Jian-Wen
2016-12-01
Despite considered a non-consumptive use of the marine environment, diving-related activities can cause damages to coral reefs. It is imminent to assess the maximum numbers of divers that can be accommodated by a diving site before it is subject to irreversible deterioration. This study aimed to assess the ecological carrying capacity of a diving site in Mabul Island, Malaysia. Photo-quadrat line transect method was used in the benthic survey. The ecological carrying capacity was assessed based on the relationship between the number of divers and the proportion of diver damaged hard corals in Mabul Island. The results indicated that the proportion of diver damaged hard corals occurred exponentially with increasing use. The ecological carrying capacity of Mabul Island is 15,600-16,800 divers per diving site per year at current levels of diver education and training with a quarterly threshold of 3900-4200 per site. Our calculation shows that management intervention (e.g. limiting diving) is justified at 8-14% of hard coral damage. In addition, the use of coral reef dominated diving sites should be managed according to their sensitivity to diver damage and the depth of the reefs. Copyright © 2016 Elsevier Ltd. All rights reserved.
Bruton, Melissa J; Cramp, Rebecca L; Franklin, Craig E
2012-05-01
The presumption that organisms benefit from thermal acclimation has been widely debated in the literature. The ability to thermally acclimate to offset temperature effects on physiological function is prevalent in ectotherms that are unable to thermoregulate year-round to maintain performance. In this study we examined the physiological and behavioural consequences of long-term exposure to different water temperatures in the aquatic snake Acrochordus arafurae. We hypothesised that long dives would benefit this species by reducing the likelihood of avian predation. To achieve longer dives at high temperatures, we predicted that thermal acclimation of A. arafurae would reduce metabolic rate and increase use of aquatic respiration. Acrochordus arafurae were held at 24 or 32°C for 3 months before dive duration and physiological factors were assessed (at both 24 and 32°C). Although filesnakes demonstrated thermal acclimation of metabolic rate, use of aquatic respiration was thermally independent and did not acclimate. Mean dive duration did not differ between the acclimation groups at either temperature; however, warm-acclimated animals increased maximum and modal dive duration, demonstrating a longer dive duration capacity. Our study established that A. arafurae is capable of thermal acclimation and this confers a benefit to the diving abilities of this snake.
Allometric scaling of lung volume and its consequences for marine turtle diving performance.
Hochscheid, Sandra; McMahon, Clive R; Bradshaw, Corey J A; Maffucci, Fulvio; Bentivegna, Flegra; Hays, Graeme C
2007-10-01
Marine turtle lungs have multiple functions including respiration, oxygen storage and buoyancy regulation, so lung size is an important indicator of dive performance. We determined maximum lung volumes (V(L)) for 30 individuals from three species (Caretta caretta n=13; Eretmochelys imbricata n=12; Natator depressus n=5) across a range of body masses (M(b)): 0.9 to 46 kg. V(L) was 114 ml kg(-1) and increased with M(b) with a scaling factor of 0.92. Based on these values for V(L) we demonstrated that diving capacities (assessed via aerobic dive limits) of marine turtles were potentially over-estimated when the V(L)-body mass effect was not considered (by 10 to 20% for 5 to 25 kg turtles and by >20% for turtles > or =25 kg). While aerobic dive limits scale with an exponent of 0.6, an analysis of average dive durations in free-ranging chelonian marine turtles revealed that dive duration increases with a mass exponent of 0.51, although there was considerable scatter around the regression line. While this highlights the need to determine more parameters that affect the duration-body mass relationship, our results provide a reference point for calculating oxygen storage capacities and air volumes available for buoyancy control.
DOE Office of Scientific and Technical Information (OSTI.GOV)
Nie, Song; Shi, Tujin; Fillmore, Thomas L.
Mass spectrometry-based targeted proteomics (e.g., selected reaction monitoring, SRM) is emerging as an attractive alternative to immunoassays for protein quantification. Recently we have made significant progress in SRM sensitivity for enabling quantification of low ng/mL to sub-ng/mL level proteins in nondepleted human blood plasma/serum without affinity enrichment. However, precise quantification of extremely low abundant but biologically important proteins (e.g., ≤100 pg/mL in blood plasma/serum) using targeted proteomics approaches still remains challenging. To address this need, we have developed an antibody-independent Deep-Dive SRM (DD-SRM) approach that capitalizes on multidimensional high-resolution reversed-phase liquid chromatography (LC) separation for target peptide enrichment combined withmore » precise selection of target peptide fractions of interest, significantly improving SRM sensitivity by ~5 orders of magnitude when compared to conventional LC-SRM. Application of DD-SRM to human serum and tissue has been demonstrated to enable precise quantification of endogenous proteins at ~10 pg/mL level in nondepleted serum and at <10 copies per cell level in tissue. Thus, DD-SRM holds great promise for precisely measuring extremely low abundance proteins or protein modifications, especially when high-quality antibody is not available.« less
Some new cave diving exploration results from Croatian karst area
NASA Astrophysics Data System (ADS)
Garasic, Davor; Garasic, Mladen
2017-04-01
In the recent years, several international cave diving expeditions took place in the Dinaric karst of Croatia. The objectives were conducting a new research of previously known karstic springs and also exploring new ones. The deepest karst cave in Croatia filled with water is Crveno jezero (lake) near Imotski town, with water depth of 281 meters and total cave depth of 528 meters. Volume of water in this cave is about 16 millions m3. Diving expeditions were held in 1997 and 1998.The deepest karst spring in the Dinaric karst of Croatia is Vrelo of Una River (with max discharge about 100 m3/s), where divers measured depth of -248 meters. Explorations were made in 2007 and 2016. Sinac spring in Pla\\vsko Polje has been dived to the depth of -203 meters. Cave diving was done in 1984, 1999, 2003, 2007 - 2016. Furthermore, very popular springs of the river Kupa (-155 m) in Gorski Kotar (explored since1995 till 2015), river Gacka (-105 in depth, 1150m in length) in Lika, explored from 1992 to 2016, river Cetina (-110 m in depth, 1300 m in length), cave diving explored from 2000 to 2016 in the Dalmatinska Zagora, Rumin Veliki spring (- 150 m in depth) in the Sinjska Krajina (explored and dived in 2006 and 2010), than rivers Krnjeza and Krupa in Ravni kotari with diving depths of over 100 meters (in 2004 and 2005) and so on. Along the Adriatic coast in Croatia there are many deep and long submarine springs (vrulje), ie. caves under seawater springs. called - vruljas for example Vrulja Zecica with over 900 meters ine length and Vrulja Modrič with completely flooded cave channels that extend over 2300 meters in length. Cave diving was conducted from 2010 to 2016. Vrulja Dubci is also worth mentioning (dived and explored in 2000), 161 meters deep and so on. Tectonic activity plays a dominant role in the creation and function of these caves. Geological, hydrogeological and lithostratigraphic conditions are also very important in speleogenesis of these caves in Croatian karst system.
Diving accidents: a cohort study from the Netherlands.
Smithuis, J W; Gips, E; van Rees Vellinga, T P; Gaakeer, M I
2016-12-01
Diving is, besides professional reasons, an increasingly popular leisure activity. Whilst statistically compared to other sports safe, diving accidents can result in serious complications. In order to treat this specific patient category adequately, early diagnosis is important. In this study, we explore various medical aspects of diving accidents. By sharing our experiences, we intend to create awareness and enhance urgent medical care for this specific category of patients. We conducted a retrospective cohort study using anonymized patient records from the emergency department (ED) of the Admiraal De Ruyter Hospital (ADRZ) and affiliated Medical Centre Hyperbaric Oxygen Therapy (MCHZ1) both in Goes, Netherlands. We evaluated all patients that presented to our ED as a diving accident from 1 November 2011 to 30 August 2015. In the selected period, 43 patients presented to our ED with complaints after diving; 84 % were male and 49 % older than 40 years, and they came by ambulance or referred by a general practitioner or other medical centres in the area; 70 % presented the same date as their dive, 21 % 1 to 3 days and 9 % later than 3 days after having dived. Pain was the most frequently reported symptom (44 %), followed by constitutional symptoms (42 %). Numbness or paraesthesia was reported in 33 %. Respiratory symptoms, dizziness, a change in mental status (e.g. apathy, confused or restlessness) and problems with coordination were present in 10-21 % of the cases. Symptoms that were apparent in less than 10 % of the cases were cutis marmorata, visual or auditory complaints, muscle weakness, cardiovascular symptoms or a malfunction of the anal sphincter or urinary bladder. Most of our patients exhibited more than one symptom; 70 % of all patients received hyperbaric oxygen recompression therapy. The limited number of patients presenting with complaints after a diving incident, the difficulty of recognition and the (potential) huge impact if not recognized and treated adequately make us believe that every diving accident should be discussed with a centre of expertise.
The role of infrequent and extraordinary deep dives in leatherback turtles (Dermochelys coriacea).
Houghton, Jonathan D R; Doyle, Thomas K; Davenport, John; Wilson, Rory P; Hays, Graeme C
2008-08-01
Infrequent and exceptional behaviours can provide insight into the ecology and physiology of a particular species. Here we examined extraordinarily deep (300-1250 m) and protracted (>1h) dives made by critically endangered leatherback turtles (Dermochelys coriacea) in the context of three previously suggested hypotheses: predator evasion, thermoregulation and exploration for gelatinous prey. Data were obtained via satellite relay data loggers attached to adult turtles at nesting beaches (N=11) and temperate foraging grounds (N=2), constituting a combined tracking period of 9.6 years (N=26,146 dives) and spanning the entire North Atlantic Ocean. Of the dives, 99.6% (N=26,051) were to depths <300 m with only 0.4% (N=95) extending to greater depths (subsequently termed ;deep dives'). Analysis suggested that deep dives: (1) were normally distributed around midday; (2) may exceed the inferred aerobic dive limit for the species; (3) displayed slow vertical descent rates and protracted durations; (4) were much deeper than the thermocline; and (5) occurred predominantly during transit, yet ceased once seasonal residence on foraging grounds began. These findings support the hypothesis that deep dives are periodically employed to survey the water column for diurnally descending gelatinous prey. If a suitable patch is encountered then the turtle may cease transit and remain within that area, waiting for prey to approach the surface at night. If unsuccessful, then migration may continue until a more suitable site is encountered. Additional studies using a meta-analytical approach are nonetheless recommended to further resolve this matter.
Performance of life support breathing apparatus for under-ice diving operations.
Lang, Michael A; Clarke, John R
2017-01-01
Single-hose scuba regulators dived in very cold water may suffer first- or second-stage malfunction, yielding complete occlusion of air flow or massive freeflow that rapidly expends a diver's air supply. This study, conducted in Antarctica, evaluated the under-ice performance of a sampling of commercially available regulators. Seventeen science divers logged a total of 305 dives in -1.86°C seawater under 6-meter-thick Antarctic fast-ice over two field seasons in 2008 and 2009. Dive profiles had an average depth of 30 msw and dive time of 29 minutes, including a mandatory three-minute safety stop at 6 msw. Sixty-nine unmodified regulator units (17 models) from 12 different manufacturers underwent standardized pre-dive regulator care and were randomly assigned to divers. Depths and times of onset of second-stage regulator freeflow were recorded. In 305 dives, there were 65 freeflows. The freeflows were not evenly distributed across the regulator brands. Regulator failure rates fell into two categories (⟨ 11% and ⟩ 26%). The regulators classified for the purpose of the test as "acceptable" (⟨ 11% failure rate: Dive-Rite Jetstream, Sherwood Maximus SRB3600, Poseidon Xstream Deep, Poseidon Jetstream, Sherwood Maximus SRB7600, Poseidon Cyklon, Mares USN22 Abyss) experienced only nine freeflows out of 146 exposures for a 6% overall freeflow incidence. Those classified as "unacceptable" (⟨ 26% failure rate) suffered 56 freeflows out of 159 exposures (35% freeflow incidence.). Contrary to expectations, the pooled incidences for the seven best performing regulators was significantly different by Chi-square test from the 10 remaining regulators (P ⟨ 0.001).
Behaviour and buoyancy regulation in the deepest-diving reptile: the leatherback turtle.
Fossette, Sabrina; Gleiss, Adrian C; Myers, Andy E; Garner, Steve; Liebsch, Nikolai; Whitney, Nicholas M; Hays, Graeme C; Wilson, Rory P; Lutcavage, Molly E
2010-12-01
In the face of the physical and physiological challenges of performing breath-hold deep dives, marine vertebrates have evolved different strategies. Although behavioural strategies in marine mammals and seabirds have been investigated in detail, little is known about the deepest-diving reptile - the leatherback turtle (Dermochelys coriacea). Here, we deployed tri-axial accelerometers on female leatherbacks nesting on St Croix, US Virgin Islands, to explore their diving strategy. Our results show a consistent behavioural pattern within dives among individuals, with an initial period of active swimming at relatively steep descent angles (∼-40 deg), with a stroke frequency of 0.32 Hz, followed by a gliding phase. The depth at which the gliding phase began increased with the maximum depth of the dives. In addition, descent body angles and vertical velocities were higher during deeper dives. Leatherbacks might thus regulate their inspired air-volume according to the intended dive depth, similar to hard-shelled turtles and penguins. During the ascent, turtles actively swam with a stroke frequency of 0.30 Hz but with a low vertical velocity (∼0.40 ms(-1)) and a low pitch angle (∼+26 deg). Turtles might avoid succumbing to decompression sickness ('the bends') by ascending slowly to the surface. In addition, we suggest that the low body temperature of this marine ectotherm compared with that of endotherms might help reduce the risk of bubble formation by increasing the solubility of nitrogen in the blood. This physiological advantage, coupled with several behavioural and physical adaptations, might explain the particular ecological niche the leatherback turtle occupies among marine reptiles.
PHYSIOLOGICAL EVALUATION OF A FREE-FLOODING DIVER HEAT REPLACEMENT GARMENT.
The general capabilities of a free-flooding heat replacement garment in maintaining thermal comfort in 40F water, at both surface and deep diving...recorded. Suit inlet temperatures which produce a subjective response of thermal comfort by the diver (Comfort Zone Inlet Temperature) at various flow
Federal Register 2010, 2011, 2012, 2013, 2014
2013-01-15
... MONITOR, NOAA has determined that enhancing recreational diving access to the wreck is a priority. NOAA... diving near the USS MONITOR; and (2) fireworks displays. The remaining two special use permit categories..., etc. Specific examples of such events involve the promotion of SCUBA diving; an annual underwater...
Federal Register 2010, 2011, 2012, 2013, 2014
2010-12-27
... Recreational Diving, Oil and Gas Operations and Commercial Fishing Seats for the Flower Garden Banks National... seats on the Flower Garden Banks National Marine Sanctuary Advisory Council: Recreational Diving, Oil... February 4, 2010. ADDRESSES: Application kits may be obtained from Jennifer Morgan, NOAA--Flower Garden...
2006-09-01
also been reported for northern bottlenose whales (Hooker and Baird 1999), and for deep- diving beluga whales, Delphinapterus leucas (Martin and...Smith. 1992. Deep diving in wild, free-ranging beluga whales, Delphinapterus leucas . Canadian Journal of Fisheries and Aquatic Sciences 49:462-466
29 CFR 1910.421 - Pre-dive procedures.
Code of Federal Regulations, 2010 CFR
2010-07-01
... transportation; and (5) The nearest U.S. Coast Guard Rescue Coordination Center. (c) First aid supplies. (1) A first aid kit appropriate for the diving operation and approved by a physician shall be available at the dive location. (2) When used in a decompression chamber or bell, the first aid kit shall be suitable...
Smart, David R; Sage, Martin; Davis, F Michael
2014-06-01
Immersion pulmonary oedema (IPE) is being increasingly recognized in swimmers, snorkellers and scuba divers presenting with acute symptoms of respiratory distress following immersion, but fatal case reports are uncommon. We report two fatal cases of probable IPE in middle-aged women, one whilst snorkelling and the other associated with a scuba dive. In the snorkeller's case, an episode of exercise-related chest tightness and shortness of breath that occurred 10 months previously was investigated but this proved negative, and she was on no medications. However, at autopsy, moderate left ventricular hypertrophy was noted. The scuba diver had suffered several previous episodes of severe shortness of breath following dives, one being so severe it led to cyanosis and impaired consciousness. At inquest, the pathologist's diagnosis was given as drowning and IPE was not mentioned. Expert input from doctors trained in diving medicine should be compulsory in the investigation of diving deaths, and forensic pathologists should be properly trained in and have guidelines for the conduct of post-immersion and post-diving autopsies.
Diving Simulation concerning Adélie Penguin
NASA Astrophysics Data System (ADS)
Ito, Shinichiro; Harada, Masanori
Penguins are sea birds that swim using lift and drag forces by flapping their wings like other birds. Although diving data can be obtained using a micro-data logger which has improved in recent years, all the necessary diving conditions for analysis cannot be acquired. In order to determine all these hard-to-get conditions, the posture and lift and drag forces of penguins were theoretically calculated by the technique used in the analysis of the optimal flight path of aircrafts. In this calculation, the actual depth and speed of the dive of an Adélie penguin (Pygoscelis adeliae) were utilized. Then, the calculation result and experimental data were compared, and found to be in good agreement. Thus, it is fully possible to determine the actual conditions of dive by this calculation, even those that cannot be acquired using a data logger.
The impact of recreational boat traffic on Marbled Murrelets (Brachyramphus marmoratus).
Bellefleur, Danielle; Lee, Philip; Ronconi, Robert A
2009-01-01
This study evaluated the impact of small boat traffic on reaction distances of Marbled Murrelets (Brachyramphus marmoratus), in the marine waters of Pacific Rim National Park Reserve, British Columbia, Canada. Observers on moving boats recorded the minimum distance the boat came to murrelets on the water, and any disturbance reaction (fly, dive, no reaction). Out of the 7500 interactions 11.7% flew, 30.8% dove and 58.1% exhibited no flushing reaction. Using a product-limit analysis, we developed curves for the proportion of Marbled Murrelets flushing (dive or flight) as a function of reaction distance. Overall, the majority of Marbled Murrelets waited until boats were within 40 m before reacting, with 25% of the population reacting at 29.2m. A stepwise Cox regression indicated that age, boat speed, and boat density (loaded in that order), significantly affected flushing response. More juveniles flushed than adults (70.1 versus 51.7%), but at closer distances. Faster boats caused a greater proportion of birds to flush, and at further distances (25% of birds flushed at 40 m at speeds > 29 kph versus 28m at speeds <12kph). A stepwise logistic regression on diving and flight responses indicated that birds tended to fly completely out of feeding areas at the approach of boats travelling >28.8 kph and later in the season (July and August). Other secondary variables included; boat density and time of day. Discussion focused on possible management actions such as the application of speed limits, set back distances, and exclusion of boat traffic to protect Marbled Murrelets.
Respiration and heart rate at the surface between dives in northern elephant seals.
Le Boeuf, B J; Crocker, D E; Grayson, J; Gedamke, J; Webb, P M; Blackwell, S B; Costa, D P
2000-11-01
All underwater activities of diving mammals are constrained by the need for surface gas exchange. Our aim was to measure respiratory rate (fb) and heart rate (fh) at the surface between dives in free-ranging northern elephant seals Mirounga angustirostris. We recorded fb and fh acoustically in six translocated juveniles, 1.8-2. 4 years old, and three migrating adult males from the rookery at Año Nuevo, California, USA. To each seal, we attached a diving instrument to record the diving pattern, a satellite tag to track movements and location, a digital audio tape recorder or acoustic datalogger with an external hydrophone to record the sounds of respiration and fh at the surface, and a VHF transmitter to facilitate recovery. During surface intervals averaging 2.2+/-0.4 min, adult males breathed a mean of 32.7+/-5.4 times at a rate of 15. 3+/-1.8 breaths min(-)(1) (means +/- s.d., N=57). Mean fh at the surface was 84+/-3 beats min(-)(1). The fb of juveniles was 26 % faster than that of adult males, averaging 19.2+/-2.2 breaths min(-)(1) for a mean total of 41.2+/-5.0 breaths during surface intervals lasting 2.6+/-0.31 min. Mean fh at the surface was 106+/-3 beats min(-)(1). fb and fh did not change significantly over the course of surface intervals. Surface fb and fh were not clearly associated with levels of exertion, such as rapid horizontal transit or apparent foraging, or with measures of immediately previous or subsequent diving performance, such as diving duration, diving depth or swimming speed. Together, surface respiration rate and the duration of the preceding dive were significant predictors of surface interval duration. This implies that elephant seals minimize surface time spent loading oxygen depending on rates of oxygen uptake and previous depletion of stores.
Can Foraging Ecology Drive the Evolution of Body Size in a Diving Endotherm?
Cook, Timothée R.; Lescroël, Amélie; Cherel, Yves; Kato, Akiko; Bost, Charles-André
2013-01-01
Within a single animal species, different morphs can allow for differential exploitation of foraging niches between populations, while sexual size dimorphism can provide each sex with access to different resources. Despite being potentially important agents of evolution, resource polymorphisms, and the way they operate in wild populations, remain poorly understood. In this study, we examine how trophic factors can select for different body sizes between populations and sexes in a diving endotherm. Dive depth and duration are positively related to body size in diving birds and mammals, a relationship explained by a lower mass-specific metabolic rate and greater oxygen stores in larger individuals. Based on this allometry, we predict that selection for exploiting resources situated at different depths can drive the evolution of body size in species of diving endotherms at the population and sexual level. To test this prediction, we studied the foraging ecology of Blue-eyed Shags, a group of cormorants with male-biased sexual size dimorphism from across the Southern Ocean. We found that mean body mass and relative difference in body mass between sexes varied by up to 77% and 107% between neighbouring colonies, respectively. Birds from colonies with larger individuals dived deeper than birds from colonies with smaller individuals, when accounting for sex. In parallel, males dived further offshore and deeper than females and the sexual difference in dive depth reflected the level of sexual size dimorphism at each colony. We argue that body size in this group of birds is under intense selection for diving to depths of profitable benthic prey patches and that, locally, sexual niche divergence selection can exaggerate the sexual size dimorphism of Blue-eyed Shags initially set up by sexual selection. Our findings suggest that trophic resources can select for important geographic micro-variability in body size between populations and sexes. PMID:23409169
Can foraging ecology drive the evolution of body size in a diving endotherm?
Cook, Timothée R; Lescroël, Amélie; Cherel, Yves; Kato, Akiko; Bost, Charles-André
2013-01-01
Within a single animal species, different morphs can allow for differential exploitation of foraging niches between populations, while sexual size dimorphism can provide each sex with access to different resources. Despite being potentially important agents of evolution, resource polymorphisms, and the way they operate in wild populations, remain poorly understood. In this study, we examine how trophic factors can select for different body sizes between populations and sexes in a diving endotherm. Dive depth and duration are positively related to body size in diving birds and mammals, a relationship explained by a lower mass-specific metabolic rate and greater oxygen stores in larger individuals. Based on this allometry, we predict that selection for exploiting resources situated at different depths can drive the evolution of body size in species of diving endotherms at the population and sexual level. To test this prediction, we studied the foraging ecology of Blue-eyed Shags, a group of cormorants with male-biased sexual size dimorphism from across the Southern Ocean. We found that mean body mass and relative difference in body mass between sexes varied by up to 77% and 107% between neighbouring colonies, respectively. Birds from colonies with larger individuals dived deeper than birds from colonies with smaller individuals, when accounting for sex. In parallel, males dived further offshore and deeper than females and the sexual difference in dive depth reflected the level of sexual size dimorphism at each colony. We argue that body size in this group of birds is under intense selection for diving to depths of profitable benthic prey patches and that, locally, sexual niche divergence selection can exaggerate the sexual size dimorphism of Blue-eyed Shags initially set up by sexual selection. Our findings suggest that trophic resources can select for important geographic micro-variability in body size between populations and sexes.
Naranjo Madrigall, Helven; Salas Marquez, Silvia
2014-12-01
Artisanal diving fisheries are a source of income, employment and food security of coastal areas in many countries. Understanding the dynamics of these fisheries, including the spatial and temporal dynamics of fishing effort, gears and species can help to address the challenges involved in fisheries management. We aimed to analyze the differences in fishing strategies undertaken by fishers that use two different diving methods (hookah and free diving), the conditions and their potential impacts on catches when adjustments to those strategies are applied over time. For this, detailed information of fishing operations from artisanal boats in the North Pacific coast of Costa Rica was analyzed in two fishing seasons (2007-2008 and 2011-2012). Data were collected by onboard observers (fishing site, fishing time, species composition, depth and visibility). Additionally, interviews with divers were applied to obtain information of price per species, species volume and fishing operations. From the total number of trips during both seasons, hookah diving was represented by a sample size of 69.3%, while free diving, with a sample of 41.9%. More than 15 species were identified in each fishing season. Nevertheless, three categories had substantial contributions in both seasons with differences in the proportions for each case: green lobster (Panulirus gracilis), octopus (Octopus sp.) and parrotfish (Scarus perrico and S. ghobban). It is worth noting that an important proportion of catch was retained by fishers for personal consumption purposes, including species of high commercial value. Additional night diving activity, increased the number of dives from one season to another. Besides, cooperation processes in free diving fishing operations, and changes in fishing effort between seasons, defined important changes in fishing strategies. Potential causes of changes in fishing strategies and the implications for management to ensure the sustainability of these fisheries in the long term are discussed.
Dive and discover: Expeditions to the seafloor
NASA Astrophysics Data System (ADS)
Lawrence, Lisa Ayers
The Dive and Discover Web site is a virtual treasure chest of deep sea science and classroom resources. The goals of Dive and Discover are to engage students, teachers, and the general public in the excitement of ocean disco very through an interactive educational Web site. You can follow scientists on oceanographic research cruises by reading their daily cruise logs, viewing photos and video clips of the discoveries, and even e-mailing questions to the scientists and crew. WHOI has also included an “Educator's Companion” section with teaching strategies, activities, and assessments, making Dive and Discover an excellent resource for the classroom.
Dive and discover: Expeditions to the seafloor
NASA Astrophysics Data System (ADS)
Ayers Lawrence, Lisa
The Dive and Discover Web site is a virtual treasure chest of deep sea science and classroom resources. The goals of Dive and Discover are to engage students, teachers, and the general public in the excitement of ocean disco very through an interactive educational Web site. You can follow scientists on oceanographic research cruises by reading their daily cruise logs, viewing photos and video clips of the discoveries, and even e-mailing questions to the scientists and crew. WHOI has also included an "Educator's Companion" section with teaching strategies, activities, and assessments, making Dive and Discover an excellent resource for the classroom.
Antioxidants may Attenuate Plasma Erythropoietin Decline after Hyperbaric Oxygen Diving.
Mutzbauer, T S; Schneider, M; Neubauer, B; Weiss, M; Tetzlaff, K
2015-11-01
According to previous studies, plasma erythropoietin (EPO) may decrease after hyperbaric oxygen exposure due to oxidative stress. It is hypothesized that the decrease of EPO can be attenuated by oxygen free radical scavengers.The aim of the present study was to evaluate whether EPO plasma levels can be influenced by oral application of vitamin C and E before repeated hyperbaric oxygen exposure during diving. 16 healthy male police task force divers performed 3 morning dives on oxygen within a regular diving schedule on 3 consecutive days. They were randomized into either the placebo group or the vitamin group, receiving 1 g ascorbic acid and 600 IU D-α-tocopherol orally 60 min before the dive. Blood samples for EPO measurement were taken on days 1, 2, and 3 at T1, T3 and T5 60 min before and at T2, T4 and T6 60 min after each dive, respectively. A moderate decrease of EPO was observed beginning at T3 until T6 in the placebo group. The EPO concentrations in the vitamin group did not show relevant variations compared to baseline. Radical scavenging vitamins C and D may counteract hyperbaric oxygen related mechanisms reducing EPO production in hyperbaric oxygen exposure during diving. © Georg Thieme Verlag KG Stuttgart · New York.
Flying After Conducting an Aircraft Excessive Cabin Leakage Test.
Houston, Stephen; Wilkinson, Elizabeth
2016-09-01
Aviation medical specialists should be aware that commercial airline aircraft engineers may undertake a 'dive equivalent' operation while conducting maintenance activities on the ground. We present a worked example of an occupational risk assessment to determine a minimum safe preflight surface interval (PFSI) for an engineer before flying home to base after conducting an Excessive Cabin Leakage Test (ECLT) on an unserviceable aircraft overseas. We use published dive tables to determine the minimum safe PFSI. The estimated maximum depth acquired during the procedure varies between 10 and 20 fsw and the typical estimated bottom time varies between 26 and 53 min for the aircraft types operated by the airline. Published dive tables suggest that no minimum PFSI is required for such a dive profile. Diving tables suggest that no minimum PFSI is required for the typical ECLT dive profile within the airline; however, having conducted a risk assessment, which considered peak altitude exposure during commercial flight, the worst-case scenario test dive profile, the variability of interindividual inert gas retention, and our existing policy among other occupational groups within the airline, we advised that, in the absence of a bespoke assessment of the particular circumstances on the day, the minimum PFSI after conducting ECLT should be 24 h. Houston S, Wilkinson E. Flying after conducting an aircraft excessive cabin leakage test. Aerosp Med Hum Perform. 2016; 87(9):816-820.
Machovsky-Capuska, Gabriel E.; Howland, Howard C.; Raubenheimer, David; Vaughn-Hirshorn, Robin; Würsig, Bernd; Hauber, Mark E.; Katzir, Gadi
2012-01-01
Australasian gannets (Morus serrator), like many other seabird species, locate pelagic prey from the air and perform rapid plunge dives for their capture. Prey are captured underwater either in the momentum (M) phase of the dive while descending through the water column, or the wing flapping (WF) phase while moving, using the wings for propulsion. Detection of prey from the air is clearly visually guided, but it remains unknown whether plunge diving birds also use vision in the underwater phase of the dive. Here we address the question of whether gannets are capable of visually accommodating in the transition from aerial to aquatic vision, and analyse underwater video footage for evidence that gannets use vision in the aquatic phases of hunting. Photokeratometry and infrared video photorefraction revealed that, immediately upon submergence of the head, gannet eyes accommodate and overcome the loss of greater than 45 D (dioptres) of corneal refractive power which occurs in the transition between air and water. Analyses of underwater video showed the highest prey capture rates during WF phase when gannets actively pursue individual fish, a behaviour that very likely involves visual guidance, following the transition after the plunge dive's M phase. This is to our knowledge the first demonstration of the capacity for visual accommodation underwater in a plunge diving bird while capturing submerged prey detected from the air. PMID:22874749
Effect of Diving and Diving Hoods on the Bacterial Flora of the External Ear Canal and Skin
1982-05-01
in parentheses itidicate number of sites tested. b One strain isolated from skin laceration exposed to water. "Diver developed external otitis media 5... otitis media (11), skin infections skin of wearing diving hoods in and out of the (6), and diarrheal diseases (10). One aspect of water. We
29 CFR Appendix B to Subpart Y of... - Guidelines for Scientific Diving
Code of Federal Regulations, 2010 CFR
2010-07-01
... 29 Labor 8 2010-07-01 2010-07-01 false Guidelines for Scientific Diving B Appendix B to Subpart Y of Part 1926 Labor Regulations Relating to Labor (Continued) OCCUPATIONAL SAFETY AND HEALTH..., Subpt. Y, App. B Appendix B to Subpart Y of Part 1926—Guidelines for Scientific Diving Note: The...
NASA Technical Reports Server (NTRS)
Behar, Alberto; Matthews, Jaret; Venkateswaran, Kasthuri; Bruckner, James; Basic, Goran; So, Edmond; Rivadeneyra, Cesar
2005-01-01
This paper provides a physical description of the current system, as well as a summary of the preliminary tests conducted in 2005: a pressure chamber test, a dive test in a 30 foot dive pool, and a dive operation at a hydrothermal vent off the northern coast of Iceland.
Scientific Diving Training Course. Red Sea & Gulf of Aden Programme (PERSGA).
ERIC Educational Resources Information Center
Arab Organization for Education and Science, Cairo (Egypt).
This document presents the scientific diving training course organized by the Arab League Educational, Cultural and Scientific Organization (ALECSO) for the Program for Environmental Studies, Red Sea and Gulf of Aden (PERSGA). This course of six weeks duration aims to produce a person who is capable of carrying out scientific diving tasks in the…
Injury survey in scuba divers of British Sub-Aqua Club: A retrospective study
Hyun, Gwang-Suk; Jee, Yong-Seok; Park, Jung-Min; Cho, Nam-Heung; Cha, Jun-Youl
2015-01-01
Scuba diving itself is generally known as a safe sports. However, various injury accidents can happen, and the incidences vary depending on divers’ education grade levels about the risks. Therefore, the study set out to identify and analyze the causes and patterns of injuries depending on the divers’ safety education grade levels through a questionnaire survey targeting ocean divers (n=12), sports divers (n=16), and dive leaders (n=15), all of whom belong to the British Sub-Aqua Club. After conducting a frequency analysis on the collected questionnaires, the conclusions are made as follows. First, in terms of diving depth, the most frequent diving depth was 15–20 m among ocean divers, 20–25 m among sports divers, and 15–20 m in case of dive leaders. Second, with regard to the causes of injuries, the most frequently answered causes are ‘overtension’ and ‘low skill’ among ocean divers; ‘low skill’ among sports divers; ‘overaction’ among dive leaders. Third, in terms of injury patterns, the most frequently answered injury patterns are ‘ear’ injuries among ocean divers; ‘ankle’ injuries among sports divers; ‘ankle’ and ‘calf’ injuries among dive leaders. Fourth, with regard to who performed first-aid when an injury accident happened, the most frequent answers are ‘instructor’ among ocean divers; ‘instructor’ and ‘self’ among sports divers; ‘self’ among dive leaders. We might suggest that more efforts need to be made to improve divers’ low dependence on specialists for treatment and consultation so that we can prevent an injury from leading to the second injury accident. PMID:26730384
Richard, Gaëtan; Vacquié-Garcia, Jade; Jouma'a, Joffrey; Picard, Baptiste; Génin, Alexandre; Arnould, John P Y; Bailleul, Frédéric; Guinet, Christophe
2014-07-15
Mature female southern elephant seals (Mirounga leonina) come ashore only in October to breed and in January to moult, spending the rest of the year foraging at sea. Mature females may lose as much as 50% of their body mass, mostly in lipid stores, during the breeding season due to fasting and lactation. When departing to sea, post-breeding females are negatively buoyant, and the relative change in body condition (i.e. density) during the foraging trip has previously been assessed by monitoring the descent rate during drift dives. However, relatively few drift dives are performed, resulting in low resolution of the temporal reconstruction of body condition change. In this study, six post-breeding females were equipped with time-depth recorders and accelerometers to investigate whether changes in active swimming effort and speed could be used as an alternative method of monitoring density variations throughout the foraging trip. In addition, we assessed the consequences of density change on the swimming efforts of individuals while diving and investigated the effects on dive duration. Both descent swimming speed and ascent swimming effort were found to be strongly correlated to descent rate during drift dives, enabling the fine-scale monitoring of seal density change over the whole trip. Negatively buoyant seals minimized swimming effort during descents, gliding down at slower speeds, and reduced their ascent swimming effort to maintain a nearly constant swimming speed as their buoyancy increased. One per cent of seal density variation over time was found to induce a 20% variation in swimming effort during dives with direct consequences on dive duration. © 2014. Published by The Company of Biologists Ltd.
Antioxidants, endothelial dysfunction, and DCS: in vitro and in vivo study.
Wang, Qiong; Mazur, Aleksandra; Guerrero, François; Lambrechts, Kate; Buzzacott, Peter; Belhomme, Marc; Theron, Michaël
2015-12-15
Reactive oxygen species (ROS) production is a well-known effect in individuals after an undersea dive. This study aimed to delineate the links between ROS, endothelial dysfunction, and decompression sickness (DCS) through the use of antioxidants in vitro and in vivo. The effect of N-acetylcysteine (NAC) on superoxide and peroxynitrite, nitric oxide (NO) generation, and cell viability during in vitro diving simulation were analyzed. Also analyzed was the effect of vitamin C and NAC on plasma glutathione thiol and thiobarbituric acid reactive substances (TBARS), plasma angiotensin-converting enzyme (ACE) activity, and angiotensin-II and DCS morbidity during in vivo diving simulation. During an in vitro diving simulation, vascular endothelial cells showed overproduction of superoxide and peroxynitrite, obvious attenuation of NO generation, and promotion of cell death, all of which were reversed by NAC treatment. After in vivo diving simulation, plasma ACE activity and angiotensin-II level were not affected. The plasma level of glutathione thiol was downregulated after the dive, which was attenuated partially by NAC treatment. Plasma TBARS level was upregulated; however, either NAC or vitamin C treatment failed to prevent DCS morbidity. During in vitro simulation, endothelial superoxide and peroxynitrite-mediated oxidative stress were involved in the attenuation of NO availability and cell death. This study is the first attempt to link oxidative stress and DCS occurrence, and the link could not be confirmed in vivo. Even in the presence of antioxidants, ROS and bubbles generated during diving and/or decompression might lead to embolic or biochemical stress and DCS. Diving-induced oxidative stress might not be the only trigger of DCS morbidity. Copyright © 2015 the American Physiological Society.
Gait switches in deep-diving beaked whales: biomechanical strategies for long-duration dives.
Martín López, Lucía Martina; Miller, Patrick J O; Aguilar de Soto, Natacha; Johnson, Mark
2015-05-01
Diving animals modulate their swimming gaits to promote locomotor efficiency and so enable longer, more productive dives. Beaked whales perform extremely long and deep foraging dives that probably exceed aerobic capacities for some species. Here, we use biomechanical data from suction-cup tags attached to three species of beaked whales (Mesoplodon densirostris, N=10; Ziphius cavirostris, N=9; and Hyperoodon ampullatus, N=2) to characterize their swimming gaits. In addition to continuous stroking and stroke-and-glide gaits described for other diving mammals, all whales produced occasional fluke-strokes with distinctly larger dorso-ventral acceleration, which we termed 'type-B' strokes. These high-power strokes occurred almost exclusively during deep dive ascents as part of a novel mixed gait. To quantify body rotations and specific acceleration generated during strokes we adapted a kinematic method combining data from two sensors in the tag. Body rotations estimated with high-rate magnetometer data were subtracted from accelerometer data to estimate the resulting surge and heave accelerations. Using this method, we show that stroke duration, rotation angle and acceleration were bi-modal for these species, with B-strokes having 76% of the duration, 52% larger body rotation and four times more surge than normal strokes. The additional acceleration of B-strokes did not lead to faster ascents, but rather enabled brief glides, which may improve the overall efficiency of this gait. Their occurrence towards the end of long dives leads us to propose that B-strokes may recruit fast-twitch fibres that comprise ∼80% of swimming muscles in Blainville's beaked whales, thus prolonging foraging time at depth. © 2015. Published by The Company of Biologists Ltd.
Myoglobin oxygen affinity in aquatic and terrestrial birds and mammals.
Wright, Traver J; Davis, Randall W
2015-07-01
Myoglobin (Mb) is an oxygen binding protein found in vertebrate skeletal muscle, where it facilitates intracellular transport and storage of oxygen. This protein has evolved to suit unique physiological needs in the muscle of diving vertebrates that express Mb at much greater concentrations than their terrestrial counterparts. In this study, we characterized Mb oxygen affinity (P50) from 25 species of aquatic and terrestrial birds and mammals. Among diving species, we tested for correlations between Mb P50 and routine dive duration. Across all species examined, Mb P50 ranged from 2.40 to 4.85 mmHg. The mean P50 of Mb from terrestrial ungulates was 3.72±0.15 mmHg (range 3.70-3.74 mmHg). The P50 of cetaceans was similar to terrestrial ungulates ranging from 3.54 to 3.82 mmHg, with the exception of the melon-headed whale, which had a significantly higher P50 of 4.85 mmHg. Among pinnipeds, the P50 ranged from 3.23 to 3.81 mmHg and showed a trend for higher oxygen affinity in species with longer dive durations. Among diving birds, the P50 ranged from 2.40 to 3.36 mmHg and also showed a trend of higher affinities in species with longer dive durations. In pinnipeds and birds, low Mb P50 was associated with species whose muscles are metabolically active under hypoxic conditions associated with aerobic dives. Given the broad range of potential globin oxygen affinities, Mb P50 from diverse vertebrate species appears constrained within a relatively narrow range. High Mb oxygen affinity within this range may be adaptive for some vertebrates that make prolonged dives. © 2015. Published by The Company of Biologists Ltd.
The Risks of Scuba Diving: A Focus on Decompression Illness
2014-01-01
Decompression Illness includes both Decompression Sickness (DCS) and Pulmonary Overinflation Syndrome (POIS), subsets of diving-related injury related to scuba diving. DCS is a condition in which gas bubbles that form while diving do not have adequate time to be resorbed or “off-gassed,” resulting in entrapment in specific regions of the body. POIS is due to an overly rapid ascent to the surface resulting in the rupture of alveoli and subsequent extravasation of air bubbles into tissue planes or even the cerebral circulation. Divers must always be cognizant of dive time and depth, and be trained in the management of decompression. A slow and controlled ascent, plus proper control of buoyancy can reduce the dangerous consequences of pulmonary barotrauma. The incidence of adverse effects can be diminished with safe practices, allowing for the full enjoyment of this adventurous aquatic sport. PMID:25478296
Bio-logging of physiological parameters in higher marine vertebrates
NASA Astrophysics Data System (ADS)
Ponganis, Paul J.
2007-02-01
Bio-logging of physiological parameters in higher marine vertebrates had its origins in the field of bio-telemetry in the 1960s and 1970s. The development of microprocessor technology allowed its first application to bio-logging investigations of Weddell seal diving physiology in the early 1980s. Since that time, with the use of increased memory capacity, new sensor technology, and novel data processing techniques, investigators have examined heart rate, temperature, swim speed, stroke frequency, stomach function (gastric pH and motility), heat flux, muscle oxygenation, respiratory rate, diving air volume, and oxygen partial pressure (P) during diving. Swim speed, heart rate, and body temperature have been the most commonly studied parameters. Bio-logging investigation of pressure effects has only been conducted with the use of blood samplers and nitrogen analyses on animals diving at isolated dive holes. The advantages/disadvantages and limitations of recording techniques, probe placement, calibration techniques, and study conditions are reviewed.
By the Light of the Moon: North Pacific Dolphins Optimize Foraging with the Lunar Cycle
NASA Astrophysics Data System (ADS)
Simonis, Anne Elizabeth
The influence of the lunar cycle on dolphin foraging behavior was investigated in the productive, southern California Current Ecosystem and the oligotrophic Hawaiian Archipelago. Passive acoustic recordings from 2009 to 2015 were analyzed to document the presence of echolocation from four dolphin species that demonstrate distinct foraging preferences and diving abilities. Visual observations of dolphins, cloud coverage, commercial landings of market squid (Doryteuthis opalescens) and acoustic backscatter of fish were also considered in the Southern California Bight. The temporal variability of echolocation is described from daily to annual timescales, with emphasis on the lunar cycle as an established behavioral driver for potential dolphin prey. For dolphins that foraged at night, the presence of echolocation was reduced during nights of the full moon and during times of night that the moon was present in the night sky. In the Southern California Bight, echolocation activity was reduced for both shallow- diving common dolphins (Delphinus delphis) and deeper-diving Risso's dolphins (Grampus griseus) during times of increased illumination. Seasonal differences in acoustic behavior for both species suggest a geographic shift in dolphin populations, shoaling scattering layers or prey switching behavior during warm months, whereby dolphins target prey that do not vertically migrate. In the Hawaiian Archipelago, deep-diving short-finned pilot whales (Globicephala macrorhynchus) and shallow-diving false killer whales (Pseudorca crassidens) also showed reduced echolocation behavior during periods of increased lunar illumination. In contrast to nocturnal foraging in the northwestern Hawaiian Islands, false killer whales in the main Hawaiian Islands mainly foraged during the day and the lunar cycle showed little influence on their nocturnal acoustic behavior. Different temporal patterns in false killer whale acoustic behavior between the main and northwestern Hawaiian Islands can likely be attributed to the presence of distinct populations or social clusters with dissimilar foraging strategies. Consistent observations of reduced acoustic activity during times of increased lunar illumination show that the lunar cycle is an important predictor for nocturnal dolphin foraging behavior. The result of this research advances the scientific understanding of how dolphins optimize their foraging behavior in response to the changing distribution and abundance of their prey.
ERIC Educational Resources Information Center
Flemming, N. C., Ed.; Max, M. D., Ed.
This publication has been prepared to provide scientific divers with guidance on safe practice under varying experimental and environmental conditions. The Code offers advice and recommendations on administrative practices, insurance, terms of employment, medical standards, training standards, dive planning, safety with different breathing gases…
ERIC Educational Resources Information Center
Thapa, Brijesh; Graefe, Alan R.; Meyer, Louisa A.
2005-01-01
Given the growth in scuba diving activities and the importance of environmental education programs to alleviate the potential impacts on coral reef ecosystems, there is a need to better understand the diving community, its environmental knowledge, and subsequent behavioral actions. The purpose of this study was to explore the role or influence of…
29 CFR Appendix B to Subpart Y of... - Guidelines for Scientific Diving
Code of Federal Regulations, 2013 CFR
2013-07-01
... 29 Labor 8 2013-07-01 2013-07-01 false Guidelines for Scientific Diving B Appendix B to Subpart Y..., Subpt. Y, App. B Appendix B to Subpart Y of Part 1926—Guidelines for Scientific Diving Note: The requirements applicable to construction work under this appendix B are identical to those set forth at appendix...
29 CFR Appendix B to Subpart Y of... - Guidelines for Scientific Diving
Code of Federal Regulations, 2012 CFR
2012-07-01
... 29 Labor 8 2012-07-01 2012-07-01 false Guidelines for Scientific Diving B Appendix B to Subpart Y..., Subpt. Y, App. B Appendix B to Subpart Y of Part 1926—Guidelines for Scientific Diving Note: The requirements applicable to construction work under this appendix B are identical to those set forth at appendix...
Dembert, M L
1977-08-01
The principal scuba diving medical problems of barotrauma, air embolism and decompression sickness have as their pathophysiologic basis the Ideal Gas Law and Boyle's Law. Hyperbaric chamber recompression therapy is the only definitive treatment of air embolism and decompression sickness. However, with a basic knowledge of diving medicine, the family physician can provide effective supportive care to the patient prior to initiation of hyperbaric therapy.
29 CFR Appendix B to Subpart T to... - Guidelines for Scientific Diving
Code of Federal Regulations, 2010 CFR
2010-07-01
... 29 Labor 5 2010-07-01 2010-07-01 false Guidelines for Scientific Diving B Appendix B to Subpart T.... 1910, Subpt. T, App. B Appendix B to Subpart T to Part 1910—Guidelines for Scientific Diving This appendix contains guidelines that will be used in conjunction with § 1910.401(a)(2)(iv) to determine those...
The neuropsychology of repeated 1- and 3-meter springboard diving among college athletes.
Zillmer, Eric A
2003-01-01
This study examined the neuropsychological effects of repeated springboard diving. It was hypothesized that the impact velocity, which can range from 20 to 30 mph, and accompanying deceleration in the water may lead to concussions and affect the diver's cognitive function. Six varsity National Collegiate Athletic Association Division 1 springboard divers participated in the study. Each diver performed a total of 50 practice dives from either the 1- or 3-m springboard. After each set of 10 dives, the participants were immediately evaluated at poolside using the Symbol Digit Modalities Test, the Stroop Color Word Test, and the Trail Making Test B. Baseline testing revealed, consistent with their athletic specialty, clear neurocognitive strengths among the divers on tests sensitive to proprioception, motor speed, and visual-spatial organization. Results from the serial assessments indicated no detectable neuropsychological deficits among competitive divers compared to baseline testing. Skilled diving at the collegiate level appears to be a safe sport and water appears to present the perfect medium for gradual deceleration. More studies, however, are warranted for 5-, 7.5-, and 10-m platform diving since the impact velocity of the diver from these heights is higher.
Provisional report on diving-related fatalities in Australian waters 2008.
Lippmann, John; Walker, Douglas; Lawrence, Chris; Fock, Andrew; Wodak, Thomas; Harris, Richard; Jamieson, Scott
2013-03-01
An individual case review of diving-related deaths, reported as occurring in Australia in 2008, was conducted as part of the DAN Asia-Pacific dive fatality reporting project. The case studies were compiled using reports from witnesses, the police and coroners. In each case, the particular circumstances of the accident and details from the post-mortem examination, where available, are provided. In total, there were 19 reported fatalities (the same as for 2007), 17 involving males. Twelve deaths occurred while snorkelling and/or breath-hold diving,and six while scuba diving. One diver died while using surface-supply breathing apparatus. Two breath-hold divers appear to have died as a result of apnoeic hypoxia, at least one case likely associated with hyperventilation. Two deaths resulted from trauma: one from impact with a boat and the other from an encounter with a great white shark. Cardiac-related issues were thought to have contributed to the deaths of five snorkellers and at least two, possibly three, scuba divers. Trauma from a marine creature, snorkelling or diving alone, apnoeic hypoxia and pre-existing medical conditions were once again features in several deaths in this series.
Gas exchange and dive characteristics of the free-swimming backswimmer Anisops deanei.
Jones, Karl K; Snelling, Edward P; Watson, Amy P; Seymour, Roger S
2015-11-01
Many aquatic insects utilise air bubbles on the surface of their bodies to supply O2 while they dive. The bubbles can simply store O2, as in the case of an 'air store', or they can act as a physical 'gas gill', extracting O2 from the water. Backswimmers of the genus Anisops augment their air store with O2 from haemoglobin cells located in the abdomen. The O2 release from the haemoglobin helps stabilise bubble volume, enabling backswimmers to remain near neutrally buoyant for a period of the dive. It is generally assumed that the backswimmer air store does not act as a gas gill and that gas exchange with the water is negligible. This study combines measurements of dive characteristics under different exotic gases (N2, He, SF6, CO) with mathematical modelling, to show that the air store of the backswimmer Anisops deanei does exchange gases with the water. Our results indicate that approximately 20% of O2 consumed during a dive is obtained directly from the water. Oxygen from the water complements that released from the haemoglobin, extending the period of near-neutral buoyancy and increasing dive duration. © 2015. Published by The Company of Biologists Ltd.
Model Based Design and Verification of a Rapid Dive Controller for an Autonomous Underwater Vehicle
1989-03-01
S,NS,0.O5): YHA’r=FILP(SD,U,XO): / OBIAS IS ADDED TO RESPONSE OF PITCH RATE OF MAGNITUDE DELTA QBIAS=DELTAO*ONES(250,1); ZB1AS=DELTAZ*ONES(250,1); YHAT...BB;CC DD]; V=EIG(AA); SD=DISCRETlZE(S,NSO0.OS); YHAT=FILP(SD,U,XO); // QBIAS IS ADDED TO RESPONSE OF PITCH RATE //OF MAGNITUDE DELTA OBIAS =DELTAQ*ONES
The development of an intermediate-duration tag to characterize the diving behavior of large whales.
Mate, Bruce R; Irvine, Ladd M; Palacios, Daniel M
2017-01-01
The development of high-resolution archival tag technologies has revolutionized our understanding of diving behavior in marine taxa such as sharks, turtles, and seals during their wide-ranging movements. However, similar applications for large whales have lagged behind due to the difficulty of keeping tags on the animals for extended periods of time. Here, we present a novel configuration of a transdermally attached biologging device called the Advanced Dive Behavior (ADB) tag. The ADB tag contains sensors that record hydrostatic pressure, three-axis accelerometers, magnetometers, water temperature, and light level, all sampled at 1 Hz. The ADB tag also collects Fastloc GPS locations and can send dive summary data through Service Argos, while staying attached to a whale for typical periods of 3-7 weeks before releasing for recovery and subsequent data download. ADB tags were deployed on sperm whales ( Physeter macrocephalus; N = 46), blue whales ( Balaenoptera musculus; N = 8), and fin whales ( B. physalus; N = 5) from 2007 to 2015, resulting in attachment durations from 0 to 49.6 days, and recording 31 to 2,539 GPS locations and 27 to 2,918 dives per deployment. Archived dive profiles matched well with published dive shapes of each species from short-term records. For blue and fin whales, feeding lunges were detected using peaks in accelerometer data and matched corresponding vertical excursions in the depth record. In sperm whales, rapid orientation changes in the accelerometer data, often during the bottom phase of dives, were likely related to prey pursuit, representing a relative measure of foraging effort. Sperm whales were documented repeatedly diving to, and likely foraging along, the seafloor. Data from the temperature sensor described the vertical structure of the water column in all three species, extending from the surface to depths >1,600 m. In addition to providing information needed to construct multiweek time budgets, the ADB tag is well suited to studying the effects of anthropogenic sound on whales by allowing for pre- and post-exposure monitoring of the whale's dive behavior. This tag begins to bridge the gap between existing long-duration but low-data throughput tags, and short-duration, high-resolution data loggers.
2014-09-30
mammals face potentially dramatic changes in the environment, as well as continued disturbances of their ocean habitat from shipping, sonar, fisheries... Cheetahs of the deep sea: deep foraging sprints in short-finned pilot whales off Tenerife (Canary Islands). Journal of Animal Ecology 77:936-947.
A portable device to assess underwater changes of cardio dynamic variables by impedance cardiography
NASA Astrophysics Data System (ADS)
Tocco, F.; Crisafulli, A.; Marongiu, E.; Milia, R.; Kalb, A.; Concu, A.
2012-12-01
Data concerning heart rate (HR), stroke volume (SV), and cardiac output (CO) during dynamic apnoea (DA) were collected from 10 healthy male, elite divers by means of an impedance cardiograph adapted to the underwater environment (C. O. Re., from 2C Technologies Inc, Italy). Three trials were performed by the divers in a 3-m-deep pool with a water temperature of 25°C: 3-minute head-out immersion during normal breathing (A), till exhaustion immersed at the surface (B) and at 3m depth (C). Both B and C conditions did not led to changes in HR, SV and CO compared to A. Data indicate that typical diving response consisting in a reduction of HR, SV and CO was not present during DA, probably due to sympathetic activation induced by exercise during DA, which partially obscured the effects of the diving response. Moreover, this study highlights the innovative role of our portable, impedance cardiography device, i.e. the C. O. Re., in easily assessing cardiodynamic changes in subjects engaged in exercise schedules including phases of underwater, dynamic apnoea.
Poor flight performance in deep-diving cormorants.
Watanabe, Yuuki Y; Takahashi, Akinori; Sato, Katsufumi; Viviant, Morgane; Bost, Charles-André
2011-02-01
Aerial flight and breath-hold diving present conflicting morphological and physiological demands, and hence diving seabirds capable of flight are expected to face evolutionary trade-offs regarding locomotory performances. We tested whether Kerguelen shags Phalacrocorax verrucosus, which are remarkable divers, have poor flight capability using newly developed tags that recorded their flight air speed (the first direct measurement for wild birds) with propeller sensors, flight duration, GPS position and depth during foraging trips. Flight air speed (mean 12.7 m s(-1)) was close to the speed that minimizes power requirement, rather than energy expenditure per distance, when existing aerodynamic models were applied. Flights were short (mean 92 s), with a mean summed duration of only 24 min day(-1). Shags sometimes stayed at the sea surface without diving between flights, even on the way back to the colony, and surface durations increased with the preceding flight durations; these observations suggest that shags rested after flights. Our results indicate that their flight performance is physiologically limited, presumably compromised by their great diving capability (max. depth 94 m, duration 306 s) through their morphological adaptations for diving, including large body mass (enabling a large oxygen store), small flight muscles (to allow for large leg muscles for underwater propulsion) and short wings (to decrease air volume in the feathers and hence buoyancy). The compromise between flight and diving, as well as the local bathymetry, shape the three-dimensional foraging range (<26 km horizontally, <94 m vertically) in this bottom-feeding cormorant.
Chan-Seng, E; Perrin, F E; Segnarbieux, F; Lonjon, N
2013-09-01
Ninety percent of the lesions resulting from diving injuries affect the cervical spine and are potentially associated with spinal cord injuries. The objective is to determine the most frequent lesion mechanisms. Evaluate the therapeutic alternatives and the biomechanical evolution (kyphotic deformation) of diving-induced cervical spine injuries. Define epidemiological characteristics of diving injuries. A retrospective analysis over a period of 10 years was undertaken for patients admitted to the Department of Neurosurgery of Montpellier, France, with cervical spinal injuries due to a diving accident. Patients were re-evaluated and clinical and radiological evaluation follow-ups were done. This study included 64 patients. Cervical spine injuries resulting from diving predominantly affect young male subjects. They represent 9.5% of all the cervical spine injuries. In 22% of cases, patients presented severe neurological troubles (ASIA A, B, C) at the time of admission. A surgical treatment was done in 85% of cases, mostly using an anterior cervical approach. This is a retrospective study (type IV) with some limitations. The incidence of diving injuries in our region is one of the highest as compared to reports in the literature. Despite an increase of our surgical indications, 55% of these cases end up with a residual kyphotic deformation but there is no relationship between the severity of late vertebral deformity and high Neck Pain and Disability Scale (NPDS) scores. Level IV, retrospective study. Copyright © 2013 Elsevier Masson SAS. All rights reserved.
Foraging patterns and prey selection in an increasing and expanding sea otter population
Laidre, K.L.; Jameson, R.J.
2006-01-01
Focal observations of sea otter (Enhydra lutris kenyoni) foraging patterns and prey selection were collected in coastal Washington between 1993 and 1999. Records consisted of 13,847 individual dives from 841 feeding bouts ranging from 1 min to >4 h. Average dive time was 55 s ?? 0.9 SE and average surface time was 45 s ?? 2.3 SE, irrespective of dive success. At least 77% of all dives (n = 10,636) were successful prey captures (dives in low light or of undetermined success were excluded). Prey capture success was significantly lower for subadults (63% ?? 5 SE) than adults (82% ?? 1 SE; P 60% red urchins (Strongylocentrotus franciscanus), with only 2 other prey species comprising >10% of their diet. Prey size and prey category were dominant predictor variables in generalized linear models of dive duration and postdive surface duration on successful dives. Significant increases in areal extent of surface canopy of giant kelp (Macrocystis integrifolia) and bull kelp (Nereocystis leutkeana) were found both in the outer coast and the Strait of Juan de Fuca (0.4-0.5 km2 per year, P < 0.05) and suggest increasing suitable habitat for a growing population. The growth and expansion of a small and isolated sea otter population provides a unique opportunity to examine the relationship between dietary diversity and population status and explore similarities and differences between trophic paradigms established for sea otter populations at other localities. ?? 2006 American Society of Mammalogists.
Presumed Arterial Gas Embolism After Breath-Hold Diving in Shallow Water.
Harmsen, Stefani; Schramm, Dirk; Karenfort, Michael; Christaras, Andreas; Euler, Michael; Mayatepek, Ertan; Tibussek, Daniel
2015-09-01
Dive-related injuries are relatively common, but almost exclusively occur in recreational or scuba diving. We report 2 children with acute central nervous system complications after breath-hold diving. A 12-year-old boy presented with unilateral leg weakness and paresthesia after diving beneath the water surface for a distance of ∼25 m. After ascent, he suddenly felt extreme thoracic pain that resolved spontaneously. Neurologic examination revealed right leg weakness and sensory deficits with a sensory level at T5. Spinal MRI revealed a nonenhancing T2-hyperintense lesion in the central cord at the level of T1/T2 suggesting a spinal cord edema. A few weeks later, a 13-year-old girl was admitted with acute dizziness, personality changes, confusion, and headache. Thirty minutes before, she had practiced diving beneath the water surface for a distance of ∼25 m. After stepping out, she felt sudden severe thoracic pain and lost consciousness. Shortly later she reported headache and vertigo, and numbness of the complete left side of her body. Neurologic examination revealed reduced sensibility to all modalities, a positive Romberg test, and vertigo. Cerebral MRI revealed no pathologic findings. Both children experienced a strikingly similar clinical course. The chronology of events strongly suggests that both patients were suffering from arterial gas embolism. This condition has been reported for the first time to occur in children after breath-hold diving beneath the water surface without glossopharyngeal insufflation. Copyright © 2015 by the American Academy of Pediatrics.
Left ventricle changes early after breath-holding in deep water in elite apnea divers.
Pingitore, Alessandro; Gemignani, Angelo; Menicucci, Danilo; Passera, Mirko; Frassi, Francesca; Marabotti, Claudio; Piarulli, Andrea; Benassi, Antonio; L'Abbate, Antonio; Bedini, Remo
2010-01-01
To study by ultrasounds cardiac morphology and function early after breath-hold diving in deep water in elite athletes. Fifteen healthy male divers (age 28 +/- 3 years) were studied using Doppler-echocardiography, immediately before (basal condition, BC) and two minutes after breath-hold diving (40 meters, acute post-apnea condition, APAC). Each subject performed a series of three consecutive breath-hold dives (20-30 and 40 m depth). End-diastolic left ventricular (LV) diameter (EDD) and end-diastolic LV volume (EDV) increased significantly (p < 0.01). Stroke volume (SV), cardiac index (CI), septal and posterior systolic wall-thickening (SWT) also significantly increased after diving (p < 0.01). No wall motion abnormalities were detected, and wall motion score index was unchanged between BC and APAC. Doppler mitral E wave increased significantly (p < 0.01), whereas the A wave was unchanged. Systemic vascular resistance (SVR) decreased significantly after diving (p < 0.05). In the factor analysis, filtering out the absolute values smaller than 0.7 in the loading matrix, it resulted that factor I consists of EDV, posterior SWT, SV and CI, factor II of diastolic blood pressure, waves A and E and factor III of heart rate and SVR. Systo-diastolic functions were improved in the early period after deep breath-hold diving due to favorable changes in loading conditions relative to pre-diving, namely the recruitment of left ventricular preload reserve and the reduction in afterload.
NASA Astrophysics Data System (ADS)
Labrousse, Sara; Vacquié-Garcia, Jade; Heerah, Karine; Guinet, Christophe; Sallée, Jean-Baptiste; Authier, Matthieu; Picard, Baptiste; Roquet, Fabien; Bailleul, Frédéric; Hindell, Mark; Charrassin, Jean-Benoit
2015-09-01
Understanding the responses of animals to the environment is crucial for identifying critical foraging habitat. Elephant seals (Mirounga leonina) from the Kerguelen Islands (49°20‧S, 70°20‧E) have several different foraging strategies. Why some individuals undertake long trips to the Antarctic continent while others utilize the relatively close frontal zones is poorly understood. Here, we investigate how physical properties within the sea ice zone are linked to foraging activities of southern elephant seals (SES). To do this, we first developed a new approach using indices of foraging derived from high temporal resolution dive and accelerometry data to predict foraging behaviour in an extensive, low resolution dataset from CTD-Satellite Relay Data Loggers (CTD-SRDLs). A sample of 37 post-breeding SES females were used to construct a predictive model applied to demersal and pelagic dive strategies relating prey encounter events (PEE) to dive parameters (dive duration, bottom duration, hunting-time, maximum depth, ascent speed, descent speed, sinuosity, and horizontal speed) for each strategy. We applied these models to a second sample of 35 seals, 20 males and 15 females, during the post-moult foraging trip to the Antarctic continental shelf between 2004 and 2013, which did not have fine-scale behavioural data. The females were widely distributed with important foraging activity south of the Southern Boundary Front, while males predominately travelled to the south-eastern part of the East Antarctica region. Combining our predictions of PEE with environmental features (sea ice concentration, water masses at the bottom phase of dives, bathymetry and slope index) we found higher foraging activity for females over shallower seabed depths and at the boundary between the overlying Antarctic Surface Water (AASW) and the underlying Modified Circumpolar Deep Water (MCDW). Increased biological activity associated with the upper boundary of MCDW, may provide overwintering areas for SES prey. Male foraging activity was strongly associated with pelagic dives within the Antarctic Slope Front where upwelling of nutrient rich Circumpolar Deep Water onto surface water may enhance and concentrate resources. A positive association between sea ice and foraging activity was found for both sexes where increased biological activity may sustain an under-ice ecosystem. Variability of the East Antarctic sea ice season duration is likely a crucial element to allow air-breathing predators to benefit from profitable prey patches within the pack ice habitat.
Domoto, Hideharu; Iwaya, Keiichi; Ikomi, Fumitaka; Matsuo, Hirotaka; Tadano, Yutaka; Fujii, Shigenori; Tachi, Kazuyoshi; Itoh, Yoshiyuki; Sato, Michiya; Inoue, Kimitoshi; Shinomiya, Nariyoshi
2016-01-01
Saturation diving (SD) is one of the safest techniques for tolerating hyperbaric conditions for long durations. However, the changes in the human plasma protein profile that occur during SD are unknown. To identify differential protein expression during or after SD, 65 blood samples from 15 healthy Japanese men trained in SD were analyzed by two-dimensional fluorescence difference gel electrophoresis. The expression of two proteins, one 32.4 kDa with an isoelectric point (pI) of 5.8 and the other 44.8 kDa with pI 4.0, were elevated during SD to 60, 100, and 200 meters sea water (msw). The expression of these proteins returned to pre-diving level when the SD training was completed. The two proteins were identified using in-gel digestion and mass spectrometric analysis; the 32.4 kDa protein was transthyretin and the 44.8 kDa protein was alpha-1-acid glycoprotein 1. Oxidation was detected at methionine 13 of transthyretin and at methionine 129 of alpha-1-acid glycoprotein 1 by tandem mass spectrometry. Moreover, haptoglobin was up-regulated during the decompression phase of 200 msw. These plasma proteins up-regulated during SD have a common function as anti-oxidants. This suggests that by coordinating their biological effects, these proteins activate a defense mechanism to counteract the effects of hyperbaric-hyperoxic conditions during SD.
Domoto, Hideharu; Iwaya, Keiichi; Ikomi, Fumitaka; Matsuo, Hirotaka; Tadano, Yutaka; Fujii, Shigenori; Tachi, Kazuyoshi; Itoh, Yoshiyuki; Sato, Michiya; Inoue, Kimitoshi; Shinomiya, Nariyoshi
2016-01-01
Saturation diving (SD) is one of the safest techniques for tolerating hyperbaric conditions for long durations. However, the changes in the human plasma protein profile that occur during SD are unknown. To identify differential protein expression during or after SD, 65 blood samples from 15 healthy Japanese men trained in SD were analyzed by two-dimensional fluorescence difference gel electrophoresis. The expression of two proteins, one 32.4 kDa with an isoelectric point (pI) of 5.8 and the other 44.8 kDa with pI 4.0, were elevated during SD to 60, 100, and 200 meters sea water (msw). The expression of these proteins returned to pre-diving level when the SD training was completed. The two proteins were identified using in-gel digestion and mass spectrometric analysis; the 32.4 kDa protein was transthyretin and the 44.8 kDa protein was alpha-1-acid glycoprotein 1. Oxidation was detected at methionine 13 of transthyretin and at methionine 129 of alpha-1-acid glycoprotein 1 by tandem mass spectrometry. Moreover, haptoglobin was up-regulated during the decompression phase of 200 msw. These plasma proteins up-regulated during SD have a common function as anti-oxidants. This suggests that by coordinating their biological effects, these proteins activate a defense mechanism to counteract the effects of hyperbaric-hyperoxic conditions during SD. PMID:27741252
Experiments and Phase-field Modeling of Hydrate Growth at the Interface of Migrating Gas Fingers
NASA Astrophysics Data System (ADS)
Fu, X.; Jimenez-Martinez, J.; Porter, M. L.; Cueto-Felgueroso, L.; Juanes, R.
2016-12-01
The first indisputable observation of a large expanse of intact seamount exposed in the inner slope of any convergent plate margin was in June 2016. The only other potential evidence for an exposed subducted seamount was observations from a series of Nautile submersible dives in the 1980's. On these dives, brecciated boulders of Cretaceous reefal debris lay on the deepest 30 m of the inner slope of the Japan Trench near Daiichi-Kashima Seamount. Because the subducting plate within 60 to 120 km outboard of a trench is usually heavily faulted, it has been suggested that seamounts impinging on a forearc region should be heavily deformed. This is not what we observed in the inner Mariana Trench during the third leg of the NOAA ship Okeanos Explorer's expedition to the Mariana subduction region. In June 2016 we recorded 275 m of exposed reef on Dive 4 (at 20.5°N) with the NOAA "Deep Discoverer" remotely operated vehicle (D-2 ROV), starting at 5,995 m on the inner slope of the Mariana Trench. The deposits are morphologically identical to observations on Dive 16 on a summit escarpment of the Cretaceous Fryer Guyot ( 20.5°N) just east of the trench. We interpret the inner trench slope exposure to be part of a Cretaceous reef complex of a seamount partially subducted beneath the overriding plate edge. Large-scale differences in the two exposures are the prevalence of vertical debris chutes between steep ridges seen in Dive 4 versus smoother, steeper slopes on Dive 16. The reefal sequences on Dive 16 show numerous fossils including bivalves in place, and layers with rudist morphology (S. Stanley, 2017, pers. comm.) in alternating tan and white bands. Similar sequences were observed on Dive 4. Slump scars observed on Dive 4 indicate mass wasting, but there is no indication of shearing or large-scale deformation. Thus, we interpret the exposure to reveal a large section of the reef complex that is partially subducted and largely intact beneath the overriding Philippine Sea Plate edge.
A Subducted Seamount Revealed: 2016, NOAA OER Deepwater Exploration of the Marianas
NASA Astrophysics Data System (ADS)
Fryer, P. B.; Kelley, C.; Pomponi, S. A.; Glickson, D.; Amon, D.
2017-12-01
The first indisputable observation of a large expanse of intact seamount exposed in the inner slope of any convergent plate margin was in June 2016. The only other potential evidence for an exposed subducted seamount was observations from a series of Nautile submersible dives in the 1980's. On these dives, brecciated boulders of Cretaceous reefal debris lay on the deepest 30 m of the inner slope of the Japan Trench near Daiichi-Kashima Seamount. Because the subducting plate within 60 to 120 km outboard of a trench is usually heavily faulted, it has been suggested that seamounts impinging on a forearc region should be heavily deformed. This is not what we observed in the inner Mariana Trench during the third leg of the NOAA ship Okeanos Explorer's expedition to the Mariana subduction region. In June 2016 we recorded 275 m of exposed reef on Dive 4 (at 20.5°N) with the NOAA "Deep Discoverer" remotely operated vehicle (D-2 ROV), starting at 5,995 m on the inner slope of the Mariana Trench. The deposits are morphologically identical to observations on Dive 16 on a summit escarpment of the Cretaceous Fryer Guyot ( 20.5°N) just east of the trench. We interpret the inner trench slope exposure to be part of a Cretaceous reef complex of a seamount partially subducted beneath the overriding plate edge. Large-scale differences in the two exposures are the prevalence of vertical debris chutes between steep ridges seen in Dive 4 versus smoother, steeper slopes on Dive 16. The reefal sequences on Dive 16 show numerous fossils including bivalves in place, and layers with rudist morphology (S. Stanley, 2017, pers. comm.) in alternating tan and white bands. Similar sequences were observed on Dive 4. Slump scars observed on Dive 4 indicate mass wasting, but there is no indication of shearing or large-scale deformation. Thus, we interpret the exposure to reveal a large section of the reef complex that is partially subducted and largely intact beneath the overriding Philippine Sea Plate edge.
Persistent (patent) foramen ovale (PFO): implications for safe diving.
Germonpré, Peter
2015-06-01
Diving medicine is a peculiar specialty. There are physicians and scientists from a wide variety of disciplines with an interest in diving and who all practice 'diving medicine': the study of the complex whole-body physiological changes and interactions upon immersion and emersion. To understand these, the science of physics and molecular gas and fluid movements comes into play. The ultimate goal of practicing diving medicine is to preserve the diver's health, both during and after the dive. Good medicine starts with prevention. For most divers, underwater excursions are not a professional necessity but a hobby; avoidance of risk is generally a much better option than risk mitigation or cure. However, prevention of diving illnesses seems to be even more difficult than treating those illnesses. The papers contained in this issue of DHM are a nice mix of various aspects of PFO that divers are interested in, all of them written by specialist doctors who are avid divers themselves. However, diving medicine should also take advantage of research from the "non-diving" medicine community, and PFO is a prime example. Cardiology and neurology have studied PFO for as long, or even longer than divers have been the subjects of PFO research, and with much greater numbers and resources. Unexplained stroke has been associated with PFO, as has severe migraine with aura. As the association seems to be strong, investigating the effect of PFO closure was a logical step. Devices have been developed and perfected, allowing now for a relatively low-risk procedure to 'solve the PFO problem'. However, as with many things in science, the results have not been as spectacular as hoped for: patients still get recurrences of stroke, still have migraine attacks. The risk-benefit ratio of PFO closure for these non-diving diseases is still debated. For diving, we now face a similar problem. Let there be no doubt that PFO is a pathway through which venous gas emboli (VGE) can arterialize, given sufficiently favourable circumstances (such as: a large quantity of VGE, size of the PFO, straining or provocation manoeuvres inducing increased right atrial pressure, delayed tissue desaturation so that seeding arterial gas emboli (AGE) grow instead of shrink, and there may be other, as yet unknown factors). There is no doubt that closing a PFO, either surgically or using a catheter-delivered device, can reduce the number of VGE becoming AGE. There is also no doubt that the procedure itself carries some health risks which are, at 1% or higher risk of serious complications, an order of magnitude greater than the risk for decompression illness (DCI) in recreational diving. Scientists seek the 'truth', but the truth about how much of a risk PFO represents for divers is not likely to be discovered nor universally accepted. First of all, the exact prevalence of PFO in divers is not known. As it has been pointed out in the recent literature, a contrast echocardiography (be it transthoracic or transoesophageal) or Doppler examination is only reliable if performed according to a strict protocol, taking into account the very many pitfalls yielding false negative results. The optimal procedure for injection of contrast medium was described several years ago, but has not received enough attention. Indeed, it is our and others' experience that many divers presenting with PFO-related DCI symptoms initially are declared "PFO-negative" by eminent, experienced cardiologists! Failing a prospective study, the risks of diving with a right-to left vascular shunt can only be expressed as an 'odds ratio', which is a less accurate measure than is 'relative risk'. The DAN Europe Carotid Doppler Study, started in 2001, is nearing completion and will provide more insight into the actual risks of DCI for recreational divers. The degree of DCI risk reduction from closing a PFO is thus not only dependent on successful closure but also (mostly?) on how the diver manages his/her dive and decompression in order to reduce the incidence of VGE. It has been convincingly shown that conservative dive profiles reduce DCI incidence even in divers with large PFOs, just as PFO closure does not protect completely from DCI if the dive profiles are aggressive. Prospective studies should not only focus on the reduction of DCI incidence after closure, but should take into account the costs and side effects of the procedure, as has been done in the cardiology and neurology studies. Imagine lung transplants becoming a routine operation, costly but with a high success rate; imagine also a longterm smoker suffering from a mild form of obstructive lung disease and exercise-limiting dyspnoea. Which of two options would you recommend: having a lung transplant and continue smoking as before, or quit smoking and observe a progressive improvement of pulmonary and cardiac pathology? As opposed to patients with thrombotic disease and migraine, divers can choose to reduce DCI risk. In fact, all it takes is acceptance that some types of diving carry too high a health risk - whether it is because of a PFO or another 'natural' factor. It would be unethical to promote PFO closure in divers solely on the basis of its efficacy of shunt reduction. Unfortunately, at least one device manufacturer has already done so in the past, citing various publications to specifically target recreational divers. Some technical diving organizations even have recommended preventive PFO closure in order to undertaking high-risk dive training. As scientists, we must not allow ourselves to be drawn into intuitive diver fears and beliefs. Nor should we let ourselves be blinded by the ease and seemingly low risk of the procedure. With proper and objective information provided by their diving medicine specialist, divers could make an informed decision, rather than focus on the simplistic idea that they need 'to get it fixed' in order to continue diving. A significant relationship between PFO and cerebral damage, in the absence of high-risk diving or DCI, has yet to be confirmed. Studying PFO-related DCI provides us with unique opportunities to learn more about the effect of gas bubbles in various tissues, including the central vascular bed and neurological tissue. It may also serve to educate divers that safe diving is something that needs to be learned, not something that can be implanted.
NASA Astrophysics Data System (ADS)
Weiland, C.; Chadwick, W. W.
2004-12-01
Several years ago we created an exciting and engaging multimedia exhibit for the Hatfield Marine Science Center that lets visitors simulate making a dive to the seafloor with the remotely operated vehicle (ROV) named ROPOS. The exhibit immerses the user in an interactive experience that is naturally fun but also educational. The public display is located at the Hatfield Marine Science Visitor Center in Newport, Oregon. We are now completing a revision to the project that will make this engaging virtual exploration accessible to a much larger audience. With minor modifications we will be able to put the exhibit onto the world wide web so that any person with internet access can view and learn about exciting volcanic and hydrothermal activity at Axial Seamount on the Juan de Fuca Ridge. The modifications address some cosmetic and logistic ISSUES confronted in the museum environment, but will mainly involve compressing video clips so they can be delivered more efficiently over the internet. The web version, like the museum version, will allow users to choose from 1 of 3 different dives sites in the caldera of Axial Volcano. The dives are based on real seafloor settings at Axial seamount, an active submarine volcano on the Juan de Fuca Ridge (NE Pacific) that is also the location of a seafloor observatory called NeMO. Once a dive is chosen, then the user watches ROPOS being deployed and then arrives into a 3-D computer-generated seafloor environment that is based on the real world but is easier to visualize and navigate. Once on the bottom, the user is placed within a 360 degree panorama and can look in all directions by manipulating the computer mouse. By clicking on markers embedded in the scene, the user can then either move to other panorama locations via movies that travel through the 3-D virtual environment, or they can play video clips from actual ROPOS dives specifically related to that scene. Audio accompanying the video clips informs the user where they are going or what they are looking at. After the user is finished exploring the dive site they end the dive by leaving the bottom and watching the ROV being recovered onto the ship at the surface. Within the three simulated dives there are a total of 6 arrival and departure movies, 7 seafloor panoramas, 12 travel movies, and 23 ROPOS video clips. This virtual exploration is part of the NeMO web site and will be at this URL http://www.pmel.noaa.gov/vents/dive.html
Foraging dives by post-breeding northern pintails
Miller, Michael R.
1983-01-01
Dabbling ducks (Anatini), including Northern Pintails (Anas acuta), typically feed by “tipping-up” (Bellrose, Ducks, Geese, and Swans of North America, Stackpole Books, Harrisburg, Pennsylvania, 1976) in shallow water. Pintails are not as adapted for diving as members of Aythyini or Oxyurini (Catlett and Johnston, Comp. Biochem. Physiol. 47A:925-931, 1974); however, incidents of foraging dives by small numbers of pintails have been reported (Chapman et al., Br. Birds 52:60, 1959; Bourget and Chapdelaine, Wildfowl 26:55-57, 1975). This paper reports on forage diving by a flock of several hundred pintails. Ecological explanations are suggested to account for the behavior and comparisons with tip-up feeding are presented.
DIVE: A Graph-based Visual Analytics Framework for Big Data
Rysavy, Steven J.; Bromley, Dennis; Daggett, Valerie
2014-01-01
The need for data-centric scientific tools is growing; domains like biology, chemistry, and physics are increasingly adopting computational approaches. As a result, scientists must now deal with the challenges of big data. To address these challenges, we built a visual analytics platform named DIVE: Data Intensive Visualization Engine. DIVE is a data-agnostic, ontologically-expressive software framework capable of streaming large datasets at interactive speeds. Here we present the technical details of the DIVE platform, multiple usage examples, and a case study from the Dynameomics molecular dynamics project. We specifically highlight our novel contributions to structured data model manipulation and high-throughput streaming of large, structured datasets. PMID:24808197
Reduced taxonomic richness of lice (Insecta: Phthiraptera) in diving birds.
Felsõ, B; Rózsa, L
2006-08-01
Avian lice occupy different habitats in the host plumage that the physical environment outside the host body may affect in several ways. Interactions between host plumage and water may be an important source of such effects. Here, we use a comparative approach to examine the effect of a host's diving behavior on the taxonomic richness of its lice. Louse genera richness was significantly lower in clades of diving birds than on their nondiving sister clades. Species richness of host and body mass did not differ significantly between these clades; thus, these factors did not bias our results. This study suggests that the hosts' diving behavior can effectively influence ectoparasite communities.
Prevalence of health promotion policies in sports clubs in Victoria, Australia.
Dobbinson, Suzanne Jane; Hayman, Jane Amanda; Livingston, Patricia Mary
2006-06-01
In recent years, some health agencies offered sponsorship to sporting associations to promote healthy environments by encouraging clubs to develop health-related policies. However, the extent to which these sponsorship contracts reach their stated aims is of concern. This study aimed to quantify levels of policy development and practice in sports clubs for each of five key health areas, namely smoke-free facilities, sun protection, healthy catering, responsible serving of alcohol and sports injury prevention. Representatives from 932 Victorian sports clubs were contacted by telephone with 640 clubs (69%) participating in the survey. Results suggested that the establishment of written policies on the key health areas by sports clubs varied widely by affiliated sport and health area: 70% of all clubs with bar facilities had written policies on responsible serving of alcohol, ranging from 58% of tennis clubs to 100% of diving and surfing clubs. In contrast, approximately one-third of sports clubs had a smoke-free policy, with 36% of tennis, 28% of country football and 28% of men's cricket clubs having policy. Moreover, 34% of clubs overall had established sun protection policy, whereas clubs competing outside during summer months, [diving (86%) and life-saving (81%)] were most likely to have a written sun protection policy. Injury prevention policies were established in 30% of sports clubs, and were most common among football (56%), diving (43%) and life-saving (41%). This study suggests that policy development for health promotion can be achieved in sports clubs when it is well supported by health agencies and consideration is given to the appropriateness of the specific behaviours to be encouraged for a given sport. Communication between associations and clubs needs to be monitored by health agencies to ensure support and resources for policy development to reach the club level.
Browaldh, Nanna; Bautista, Tara G; Dutschmann, Mathias; Berkowitz, Robert G
2016-11-01
To review the scientific literature on the relationship between Kölliker-Fuse nucleus (KF) and cranial nerve function in animal models, with view to evaluating the potential role of KF maturation in explaining age-related normal physiologic parameters and developmental and acquired impairment of cranial nerve function in humans. Medical databases (Medline and PubMed). Studies investigating evidence of KF activity responsible for a specific cranial nerve function that were based on manipulation of KF activity or the use of neural markers were included. Twenty studies were identified that involved the trigeminal (6 studies), vagus (9), and hypoglossal nerves (5). These pertained specifically to a role of the KF in mediating the dive reflex, laryngeal adductor control, swallowing function and upper airway tone. The KF acts as a mediator of a number of important functions that relate primarily to laryngeal closure, upper airway tone and swallowing. These areas are characterized by a variety of disorders that may present to the otolaryngologist, and hence the importance of understanding the role played by the KF in maintaining normal function.
Sakamoto, Kentaro Q; Takahashi, Akinori; Iwata, Takashi; Trathan, Philip N
2009-10-07
Albatrosses fly many hundreds of kilometers across the open ocean to find and feed upon their prey. Despite the growing number of studies concerning their foraging behaviour, relatively little is known about how albatrosses actually locate their prey. Here, we present our results from the first deployments of a combined animal-borne camera and depth data logger on free-ranging black-browed albatrosses (Thalassarche melanophrys). The still images recorded from these cameras showed that some albatrosses actively followed a killer whale (Orcinus orca), possibly to feed on food scraps left by this diving predator. The camera images together with the depth profiles showed that the birds dived only occasionally, but that they actively dived when other birds or the killer whale were present. This association with diving predators or other birds may partially explain how albatrosses find their prey more efficiently in the apparently 'featureless' ocean, with a minimal requirement for energetically costly diving or landing activities.
Sakamoto, Kentaro Q.; Takahashi, Akinori; Iwata, Takashi; Trathan, Philip N.
2009-01-01
Albatrosses fly many hundreds of kilometers across the open ocean to find and feed upon their prey. Despite the growing number of studies concerning their foraging behaviour, relatively little is known about how albatrosses actually locate their prey. Here, we present our results from the first deployments of a combined animal-borne camera and depth data logger on free-ranging black-browed albatrosses (Thalassarche melanophrys). The still images recorded from these cameras showed that some albatrosses actively followed a killer whale (Orcinus orca), possibly to feed on food scraps left by this diving predator. The camera images together with the depth profiles showed that the birds dived only occasionally, but that they actively dived when other birds or the killer whale were present. This association with diving predators or other birds may partially explain how albatrosses find their prey more efficiently in the apparently ‘featureless’ ocean, with a minimal requirement for energetically costly diving or landing activities. PMID:19809497
Brünnich's guillemots (Uria lomvia) maintain high temperature in the body core during dives.
Niizuma, Yasuaki; Gabrielsen, Geir W; Sato, Katsufumi; Watanuki, Yutaka; Naito, Yasuhiko
2007-06-01
A major challenge for diving birds, reptiles, and mammals is regulating body temperature while conserving oxygen through a reduction in metabolic processes. To gain insight into how these needs are met, we measured dive depth and body temperatures at the core or periphery between the skin and abdominal muscles simultaneously in freely diving Brünnich's guillemots (Uria lomvia), an arctic seabird, using an implantable data logger (16-mm diameter, 50-mm length, 14-g mass, Little Leonardo Ltd., Tokyo). Guillemots exhibited increased body core temperatures, but decreased peripheral temperatures, during diving. Heat conservation within the body core appeared to result from the combined effect of peripheral vasoconstriction and a high wing beat frequency that generates heat. Conversely, the observed tissue hypothermia in the periphery should reduce metabolic processes as well as heat loss to the water. These physiological effects are likely one of the key physiological adaptations that makes guillemots to perform as an efficient predator in arctic waters.
Click rates and silences of sperm whales at Kaikoura, New Zealand
NASA Astrophysics Data System (ADS)
Douglas, Lesley A.; Dawson, Stephen M.; Jaquet, Nathalie
2005-07-01
Analysis of the usual click rates of sperm whales (Physeter macrocephalus) at Kaikoura, New Zealand, confirms the potential for assessing abundance via ``click counting.'' Usual click rates over three dive cycles each of three photographically identified whales showed that 5 min averages of usual click rate did not differ significantly within dives, among dives of the same whale or among whales. Over the nine dives (n=13 728 clicks) mean usual click rate was 1.272 clicks s-1 (95% CI=0.151). On average, individual sperm whales at Kaikoura spent 60% of their time usual clicking in winter and in summer. There was no evidence that whale identity or stage of the dive recorded affects significantly the percentage of time spent usual clicking. Differences in vocal behavior among sperm whale populations worldwide indicate that estimates of abundance that are based on click rates need to based on data from the population of interest, rather than from another population or some global average.
Villegas-Amtmann, Stella; Atkinson, Shannon; Paras-Garcia, Alberto; Costa, Daniel P
2012-08-01
Survival depends on an animal's ability to find and acquire prey. In diving vertebrates, this ability is directly related to their physiological capability (e.g. oxygen stores). We studied the seasonal variation in oxygen stores, body temperature and body condition in California sea lions (Zalophus californianus) (CSL) as a function of seasonal variation in temperature, primary productivity, diving behavior and reproductive stage. During summer, blood oxygen stores were significantly greater and muscle oxygen stores were significantly lower than in winter. Total oxygen stores, body condition and body temperature did not change between seasons but variations in body temperature were greater during summer. Changes in oxygen stores are partly attributed to diving behavior, temperature and pregnancy that could increase oxygen consumption. Blood and muscle oxygen stores appear to be influenced by reproductive state. Blood oxygen stores are more likely influenced by diving behavior and temperature than muscle oxygen stores. Copyright © 2012 Elsevier Inc. All rights reserved.
Rozali, A; Khairuddin, H; Sherina, M S; Zin, B Mohd; Sulaiman, A
2008-06-01
Occupational divers are exposed to hazards which contribute to the risk of developing decompression illnesses (DCI). DCI consists of Type I decompression sickness (DCS), Type II DCS and arterial gas embolism (AGE), developed from formation of bubbles in the tissues or circulation as a result of inadequate elimination of inert gas (nitrogen) after a dive. In Malaysia, DCI is one of the significant contributions to mortality and permanent residual morbidity in diving accidents. This is a case of a diver who suffered from Type II DCS with neurological complications due to an occupational diving activity. This article mentions the clinical management of the case and makes several recommendations based on current legislations and practise implemented in Malaysia in order to educate medical and health practitioners on the current management of DCI from the occupational perspective. By following these recommendations, hopefully diving accidents mainly DCI and its sequalae among occupational divers can be minimized and prevented, while divers who become injured receive the proper compensation for their disabilities.
The Impact of Experience, Exposure and Support on Emergency Worker Health
1996-02-26
resources (Lazarus & Folkman, 1984). One method of coping with stress reflects attempts to manage emotional responses during stress o r after...contribution of the factors under study to response outcomes, more methodologica l ly sound studies need to be conducted. Some of the probl ems with...t le r estaurants whose wi ndows vaunted e~quisite feeding; taverns, dives, oyster and wine ba r s , ham and beef shops ; and small shops marketing
Convolutional Architecture Exploration for Action Recognition and Image Classification
2015-01-01
that has 200 videos taken in 720x480 resolution of 9 different sporting activities: diving, golf , swinging , kicking, lifting, horseback riding, running...sporting activities: diving, golf swinging , kicking, lifting, horseback riding, running, skateboarding, swinging (various gymnastics), and walking. In this...Testing Videos Diving 13 3 Golf Swinging 21 4 Horseback Riding 11 3 Kicking 21 4 Lifting 12 3 Running 12 3 Skateboarding 12 3 Swinging (Gymnastics) 28
Blood Oxygen Conservation in Diving Sea Lions: How Low Does Oxygen Really Go
2015-09-30
5 6 7 0 20 40 60 80 100 % H b Sa tu ra tio n (S O2 ) Time into Dive (min) Arterial SO2 PostVenaCava SO2 AntVenaCava SO2 3 Figure 2. Rate of...change in posterior venacaval hemoglobin saturation ( SO2 ) in relation to stroke rate during descent, bottom phase, and ascent of all dives of sea
Lung Mechanics in Marine Mammals
2014-09-30
468. 13. Fahlman, A., et al., Estimating the effect of lung collapse and pulmonary shunt on gas exchange during breath -hold diving: the Scholander...vital to understand how diving mammals manage inert and metabolic gases during diving and will help determine what behavioral and physiological...N2 levels, and that they use both physiological and behavioral means to avoid DCS [1, 2]. But what physiological variables are the most important to
Buzzacott, P; Denoble, P J
2017-01-01
The first diver certification programme for recreational 'enriched air nitrox' (EAN) diving was released in 1985. Concerns were expressed that many EAN divers might suffer central nervous system (CNS) oxygen toxicity seizures and drown. US fatalities on open-circuit scuba occurring between 2004-2013, where the breathing gas was either air or EAN, were identified. Causes of death and preceding circumstances were examined by a medical examiner experienced in diving autopsies. Case notes were searched for witnessed seizures at elevated partial pressures of oxygen. The dataset comprised 344 air divers (86%) and 55 divers breathing EAN (14%). EAN divers' fatal dives were deeper than air divers' (28 msw vs 18 msw, p < 0.0001). Despite this, of the 249 cases where a cause of death was established, only three EAN divers were considered to have possibly died following CNS oxygen toxicity seizures at depth (ppO2 132, 142 and 193 kPa). The analysis of recreational diving fatalities in the US over 10 years found just one death likely from CNS oxygen toxicity among EAN divers. A further two possible, although unlikely, cases were also found. Fears of commonplace CNS oxygen toxicity seizures while EAN diving have not apparently been realized.
Biosonar, diving and movements of two tagged white-beaked dolphin in Icelandic waters
NASA Astrophysics Data System (ADS)
Rasmussen, M. H.; Akamatsu, T.; Teilmann, J.; Vikingsson, G.; Miller, L. A.
2013-04-01
For the first time bio-logging tags were attached to free-ranging white-beaked dolphins, Lagenorhynchus albirostris. A satellite tag was attached to one animal while an acoustic A-tag, a time-depth recorder and a VHF transmitter complex was attached to a second dolphin with a suction cup. The satellite tag transmitted for 201 day, during which time the dolphin stayed in the coastal waters of western Iceland. The acoustic tag complex was on the second animal for 13 h and 40 min and provided the first insight into the echolocation behaviour of a free-ranging white-beaked dolphin. The tag registered 162 dives. The dolphin dove to a maximum depth of 45 m, which is about the depth of the bay in which the dolphin was swimming. Two basic types of dives were identified; U-shaped and V-shaped dives. The dolphin used more time in U-shaped dives, more clicks and sonar signals with shorter click intervals compared to those it used in V-shaped dives. The dolphin was in acoustic contact with other dolphins about five hours after it was released and stayed with these for the rest of the tagging time. Possible foraging attempts were found based on the reduction of click intervals from about 100 ms to 2-3 ms, which suggests a prey capture attempt. We found 19 punitive prey capture attempts and of these 53% occurred at the maximum dive depth. This suggests that more than half of the possible prey capture events occurred at or near the sea bed.
A subjective evaluation of a drinking system for saturation divers.
Hope, Arvid; Brekken, Rudolf
2010-03-01
Studies have shown that divers may lose large volumes of body fluids in hot water suit (HWS) dives lasting for four hours or longer, and that this dehydration is mainly caused by sweating. Body fluid balance may be impaired and the diver's alertness and power of judgement could be influenced by such imbalance. The main objective of the present study was to obtain a subjective judgement of a drinking system for divers (DSFD) and to obtain information related to body fluid loss during long saturation lock-out dives. Via a suction pipe imbedded in the microphone unit in the oronasal mask, the DSFD makes it possible for the diver to drink while in the water. Ten divers tested the drinking system during 12 saturation lock-out dives lasting on average for 5.5 h. A questionnaire was answered after each dive. The divers drank 21 times (range 5-30 times) during the dives, and the average drinking volume was 1.4 litre (range 1.0-1.5 litre) but only drank 0.04 litre (range 0-0.3 litre) in the bell after diving. The system was easy to operate and preparation and clothing did not cause any delay. The suction pipe did not intrude and the microphone performed excellently. The work in water was not hindered by DSFD and all divers were very satisfied with the drinking system. It was obvious that the need for fluid intake after a dive with DSFD was markedly reduced; another good indication of maintained body fluid balance.
Sponge divers of the Aegean and medical consequences of risky compressed-air dive profiles.
Toklu, Akin Savas; Cimsit, Maide
2009-04-01
Historically, Turkey once had a substantial number of professional sponge divers, a population known for a relatively high incidence of diving-related conditions such as decompression sickness (DCS) and dysbaric osteonecrosis (DON). Sponge diving ended in the mid-1980s when nearly all of the sponges in the Aegean and Mediterranean Seas contracted a bacterial disease and the occupation became unprofitable. We reviewed the records of Turkish sponge divers for information on their level of knowledge, diving equipment, dive profiles, and occupational health problems. Information was collected by: 1) interviewing former sponge divers near Bodrum, where most of them had settled; 2) reviewing the relevant literature; and 3) examining the medical records of sponge divers who underwent recompression treatment. These divers used three types of surface-supplied equipment, including hard helmets, Fernez apparatus, and hookahs; the latter were preferred because they allowed divers the greatest freedom of movement while harvesting sponges underwater. These divers used profiles that we now know involved a high risk for DCS and DON. We were able to access the records of 58 divers who had received recompression treatment. All of the cases involved severe DCS and delays from dive to recompression that averaged 72 h. Complete resolution of symptoms occurred in only 11 cases (19%). Thus, we were able to document the several factors that contributed to the risks in this occupational group, including unsafe dive profiles, resistance to seeking treatment, long delays before recompression, and the fact that recompression treatment used air rather than oxygen.
Perry, Matthew C.; Wells-Berlin, Alicia M.; Kidwell, David M.; Osenton, Peter C.
2007-01-01
Population and trophic relationships among diving ducks in Chesapeake Bay are diverse and complex as they include five species of bay ducks (Aythya spp.), nine species of seaducks (Tribe Mergini), and the Ruddy Duck (Oxyura jamaicensis). Here we considered the relationships between population changes and diet over the past half century to assess the importance of prey changes to wintering waterfowl in the Bay. Food habits of 643 diving ducks collected from Chesapeake Bay during 1999-2006 were determined by analyses of their gullet (esophagus and proventriculus) and gizzard contents and compared to historical data (1885-1979) of 1,541 diving ducks. Aerial waterfowl surveys, in general, suggest that six species of seaducks were more commonly located in the meso- to polyhaline areas of the Bay, whereas five species of bay ducks and Ruddy Ducks were in the oligo- to mesohaline areas. Seaducks fed on a molluscan diet of Hooked Mussel (Ischadium recurvum), Amethyst Gemclam (Gemma gemma), and Dwarf Surfclarn (Mulinia lateralis). Bay ducks and Ruddy Ducks fed more on Baltic Macoma (Macoma balthica), the adventive Atlantic Rangia (Rangia cuneata), and submerged aquatic vegetation (SAV). Mergansers were found over the widest salinity range in the Bay, probably because of their piscivorous diet. Each diving duck species appears to fill a unique foraging niche, although there is much overlap of selected prey. When current food habits are compared to historic data, only the Canvasback (Aythya valisineria) has had major diet changes, although SAV now accounts for less food volume for all diving duck species, except the Redhead (Aythya americana). Understanding the trophic-habitat relationships of diving ducks in coastal wintering areas will give managers a better understanding of the ecological effects of future environmental changes. Intensive restoration efforts on SAV and oyster beds should greatly benefit diving duck populations.
Jia, Xinghua; Zhang, Mingjun
2012-01-01
Whirligig beetles (Coleoptera, Gyrinidae) can fly through the air, swiftly swim on the surface of water, and quickly dive across the air-water interface. The propulsive efficiency of the species is believed to be one of the highest measured for a thrust generating apparatus within the animal kingdom. The goals of this research were to understand the distinctive biological mechanisms that allow the beetles to swim and dive, while searching for potential bio-inspired robotics applications. Through static and dynamic measurements obtained using a combination of microscopy and high-speed imaging, parameters associated with the morphology and beating kinematics of the whirligig beetle's legs in swimming and diving were obtained. Using data obtained from these experiments, dynamics models of both swimming and diving were developed. Through analysis of simulations conducted using these models it was possible to determine several key principles associated with the swimming and diving processes. First, we determined that curved swimming trajectories were more energy efficient than linear trajectories, which explains why they are more often observed in nature. Second, we concluded that the hind legs were able to propel the beetle farther than the middle legs, and also that the hind legs were able to generate a larger angular velocity than the middle legs. However, analysis of circular swimming trajectories showed that the middle legs were important in maintaining stable trajectories, and thus were necessary for steering. Finally, we discovered that in order for the beetle to transition from swimming to diving, the legs must change the plane in which they beat, which provides the force required to alter the tilt angle of the body necessary to break the surface tension of water. We have further examined how the principles learned from this study may be applied to the design of bio-inspired swimming/diving robots. PMID:23209398
Carter, Matt Ian Daniel; Bennett, Kimberley A; Embling, Clare B; Hosegood, Philip J; Russell, Debbie J F
2016-01-01
In the last thirty years, the emergence and progression of biologging technology has led to great advances in marine predator ecology. Large databases of location and dive observations from biologging devices have been compiled for an increasing number of diving predator species (such as pinnipeds, sea turtles, seabirds and cetaceans), enabling complex questions about animal activity budgets and habitat use to be addressed. Central to answering these questions is our ability to correctly identify and quantify the frequency of essential behaviours, such as foraging. Despite technological advances that have increased the quality and resolution of location and dive data, accurately interpreting behaviour from such data remains a challenge, and analytical methods are only beginning to unlock the full potential of existing datasets. This review evaluates both traditional and emerging methods and presents a starting platform of options for future studies of marine predator foraging ecology, particularly from location and two-dimensional (time-depth) dive data. We outline the different devices and data types available, discuss the limitations and advantages of commonly-used analytical techniques, and highlight key areas for future research. We focus our review on pinnipeds - one of the most studied taxa of marine predators - but offer insights that will be applicable to other air-breathing marine predator tracking studies. We highlight that traditionally-used methods for inferring foraging from location and dive data, such as first-passage time and dive shape analysis, have important caveats and limitations depending on the nature of the data and the research question. We suggest that more holistic statistical techniques, such as state-space models, which can synthesise multiple track, dive and environmental metrics whilst simultaneously accounting for measurement error, offer more robust alternatives. Finally, we identify a need for more research to elucidate the role of physical oceanography, device effects, study animal selection, and developmental stages in predator behaviour and data interpretation.
Hedger, R D; Rikardsen, A H; Thorstad, E B
2017-01-01
The effects of large, externally attached pop-up satellite archival tags (PSATs) were compared with those of small implanted data storage tags (DSTs) on adult Atlantic salmon Salmo salar during their ocean migration in regards to depth utilization, diving depth, diving rate, diving speed and temperatures experienced. Additionally the return rate and growth of individuals tagged with PSATs was compared with those of small acoustic tags and DSTs. Overall, the depth distribution of individuals tagged with PSATs was similar to that of those tagged with DSTs, reflecting the pelagic nature of S. salar at sea. Individuals tagged with PSATs, however, dived less frequently and to shallower depths, and dived and surfaced at slower velocities. Sea surface temperatures experienced by individuals tagged with PSATs were similar to those experienced by those tagged with DSTs for the same time of year, suggesting that there were no large differences in the ocean migration. Return rates did not depend on whether individuals were tagged with PSATs or not, indicating that survival at sea was not impacted by PSATs in comparison to small internal tags. Individuals tagged with PSATs, however, had a smaller increase in body mass than those tagged with acoustic tags or DSTs. It was concluded that PSATs are suitable for use in researching large-scale migratory behaviour of adult S. salar at sea, but that some effects on their behaviour from tagging must be expected. Effects of PSATs may be largest in the short term when S. salar are swimming in bursts at high speeds. Even though individuals tagged with PSATs performed deep and frequent dives, the results of this study suggest that untagged individuals would perform even deeper and more frequent dives than tagged individuals. © 2016 The Fisheries Society of the British Isles.
Ludynia, Katrin; Dehnhard, Nina; Poisbleau, Maud; Demongin, Laurent; Masello, Juan F.; Quillfeldt, Petra
2012-01-01
Logger technology has revolutionised our knowledge of the behaviour and physiology of free-living animals but handling and logger attachments may have negative effects on the behaviour of the animals and their welfare. We studied southern rockhopper penguin (Eudyptes chrysocome) females during the guard stage in three consecutive breeding seasons (2008/09−2010/11) to evaluate the effects of handling and logger attachment on foraging trip duration, dive behaviour and physiological parameters. Smaller dive loggers (TDRs) were used in 2010/11 for comparison to larger GPS data loggers used in all three seasons and we included two categories of control birds: handled controls and PIT control birds that were previously marked with passive integrative transponders (PITs), but which had not been handled during this study. Increased foraging trip duration was only observed in GPS birds during 2010/11, the breeding season in which we also found GPS birds foraging further away from the colony and travelling longer distances. Compared to previous breeding seasons, 2010/11 may have been a period with less favourable environmental conditions, which would enhance the impact of logger attachments. A comparison between GPS and TDR birds showed a significant difference in dive depth frequencies with birds carrying larger GPS data loggers diving shallower. Mean and maximum dive depths were similar between GPS and TDR birds. We measured little impact of logger attachments on physiological parameters (corticosterone, protein, triglyceride levels and leucocyte counts). Overall, handling and short-term logger attachments (1–3 days) showed limited impact on the behaviour and physiology of the birds but care must be taken with the size of data loggers on diving seabirds. Increased drag may alter their diving behaviour substantially, thus constraining them in their ability to catch prey. Results obtained in this study indicate that data recorded may also not represent their normal dive behaviour. PMID:23185623
Human Bone Matrix Changes During Deep Saturation Dives
2008-08-08
of theories have been investigated to elucidate this 2 separate factor. They include fat embolism , decompression triggered intravascular...arterial gas embolism . In bone, bubbles can also cause a mechanical blockage of blood flow that result in tissue damage and necrosis. Blood flow is...exposures, but the longest follow-up to date is only 10 years.11 If lesions progress and cause femoral fractures or osteoarthritis, the typical
M. Zachariah Peery; Scott H. Newman; Curt D. Storlazzi; Steven R. Beissinger
2009-01-01
Seabirds maintain plasticity in their foraging behavior to cope with energy demands and foraging constraints that vary over the reproductive cycle, but behavioral studies comparing breeding and nonbreeding individuals are rare. Here we characterize how Marbled Murrelets (Brachyramphus marmoratus) adjust their foraging effort in response to changes...
ERIC Educational Resources Information Center
Freifeld, Lorri
2012-01-01
Many employees claim they learn best while doing, so they prefer to dive right in and learn as they go when they get a new job or take on new responsibilities. But the most successful and quickest learning on the job takes place when there is a formal on-the-job training (OJT) program--a fact many organizations and managers fail to take into…
NASA Astrophysics Data System (ADS)
Galvin, C.
2012-12-01
Cyril Galvin, Coastal Engineer Springfield, Virginia 22150 USA Since 1911, the Steel Pier at Atlantic City, New Jersey, has been the site of the Atlantic City tide gauge, except for two intervals: 1911-1921 when the gauge was at the Million Dollar Pier in Atlantic City, and 1985-1991 when the gauge was at the Ventnor Fishing Pier (see Table 2, Zervos, 2009). By design, the Steel Pier was an amusement pier, and its most famous amusement was the Diving Horses: they dove bareback with a woman rider from a platform about 40 feet above sea level. They did that between 1929 and 1978, except for seven years - a post-war period, 1945 to 1953, when diving was suspended. The popularity of the diving horses is recorded on photos of crowds which occupied the bleachers at the seaward end of the pier to view the diving horses. By my count, the crowd pictured in the end papers of the book by Steve Liebowitz (2009) was about 4000 people. Typically, there were multiple shows daily. The weight of the crowd, estimated from the count of the crowd, was about 150 tons. This weight was loaded down on the piles by the crowd of spectators, and unloaded between shows of the diving horses. Most of the piles supporting the pier deck were imbedded in sand newly deposited since 1850. Using Atlantic City sea levels from the PSMSL data base and historical facts from Liebowitz (2009), and beginning with a 1912 start of the tide gauge, the apparent sea level rose at a rate of 3.1mm/yr until 1929 when the horses began diving. With the 1929 start of diving, the apparent sea level rise tripled, averaging 9.4 mm/yr until the act was suspended in 1945. In the 1945-1953 interval, when the horses did not dive (no crowds on the pier), apparent sea level fell (sea level FELL) at a rate of -1.6 mm/yr. The horses resumed diving in 1953, when the apparent sea level resumed at a rate of 4.0mm/yr. This 4.0 mm/yr is identical to the longtime sea level trend (1911-2006) from Zervos (2009) of 3.99mm/yr The history of apparent sea level rise at Steel Pier is consistent with increases caused by loading the pier deck with crowds, and the absence of apparent sea level rise when the pier deck was not loaded by spectators. CG/08Aug 2012
Cognitive symptoms and welding fume exposure.
Ross, John A S; Macdiarmid, Jennifer I; Semple, Sean; Watt, Stephen J; Moir, Gill; Henderson, George
2013-01-01
Prevalence of moderate to severe cognitive symptoms is markedly higher in UK professional divers who have also worked as a welder (28%) than in either divers who have not welded (18%) or offshore workers who have worked neither as a diver nor as a welder (6%). To determine whether cognitive symptoms are related to welding fume exposure or diving. Three age-matched groups of male workers were studied using postal questionnaire: professional divers who had worked as a welder (PDW, n = 361), professional welders who had not dived (NDW, n = 352), and offshore oil field workers who had neither dived nor welded (NDNW, n =503). Health-related quality of life was assessed by the Short Form 12 questionnaire (SF12). Cognitive symptomatology was assessed using the Cognitive Failures Questionnaire (CFQ). A single variable for welding fume exposure (mg m(-3) days) was calculated, incorporating welding experience in different environments and using different welding techniques and respiratory protective equipment. The level of fume exposure during hyperbaric welding operations was measured during such work as ambient PM(10) (particles of 10 µm or less). Diving exposure was assessed as the number of dives performed plus the number of days spent working during saturation diving. Questionnaires were returned by 153 PDW, 108 NDW, and 252 NDNW. SF12 scores were the same in all groups and fell within normative values. Mean (95% CI) CFQ scores were higher in PDW [40.3 (37.7-42.9)] than in both NDW [34.6 (31.6-37.7)] and NDNW [32.1 (30.4-33.9)], but the scores in no groups fell outside the normative range. The mean PM(10) exposure during hyperbaric welding operations was 2.58 mg m(-3). The geometric mean mg m(-3) days (95% CI) for welding fume exposure in NDW [33 128 (24 625-44 567) n = 85] was higher than for that in PDW [10 904 (8103-14 673) n = 112]. For PDW the geometric mean (95% CI) diving exposure was 1491 [(1192-1866) n = 94] dives and days in saturation. In the general linear model regression analyses adjusted for age, alcohol consumption, and somatization, there was no signification association of CFQ score with either welding fume exposure (F = 0.072, P = 0.79, n = 152) or diving exposure (F = 0.042, P = 0.84, n = 74). In conclusion, cognitive sympomatology was not related to retrospectively assessed measures of welding fume exposure or diving experience. In addition, the levels of cognitive symptomatology, even in PDW, did not exceed normative values.
Virtual Diving in the Underwater Archaeological Site of Cala Minnola
NASA Astrophysics Data System (ADS)
Bruno, F.; Lagudi, A.; Barbieri, L.; Muzzupappa, M.; Mangeruga, M.; Pupo, F.; Cozza, M.; Cozza, A.; Ritacco, G.; Peluso, R.; Tusa, S.
2017-02-01
The paper presents the application of the technologies and methods defined in the VISAS project for the case study of the underwater archaeological site of Cala Minnola located in the island of Levanzo, in the archipelago of the Aegadian Islands (Sicily, Italy). The VISAS project (http://visas-project.eu) aims to improve the responsible and sustainable exploitation of the Underwater Cultural Heritage by means the development of new methods and technologies including an innovative virtual tour of the submerged archaeological sites. In particular, the paper describes the 3D reconstruction of the underwater archaeological site of Cala Minnola and focus on the development of the virtual scene for its visualization and exploitation. The virtual dive of the underwater archaeological site allows users to live a recreational and educational experience by receiving historical, archaeological and biological information about the submerged exhibits, the flora and fauna of the place.
Global population trends and human use patterns of Manta and Mobula rays.
Ward-Paige, Christine A; Davis, Brendal; Worm, Boris
2013-01-01
Despite being the world's largest rays and providing significant revenue through dive tourism, little is known about the population status, exploitation and trade volume of the Mobulidae (mobulids; Manta and Mobula spp.). There is anecdotal evidence, however, that mobulid populations are declining, largely due to the recent emergence of a widespread trade for their gill rakers, which is reflected in increasing Food and Agriculture Organization landings trends. Here, we present results from two dedicated diver surveys, one from the eManta project, which includes summary observations from ninety 10°x10° regions with ∼200-62,000 dives per region, and the other from the Reef Environmental Education Foundation, which includes spatially more detailed observations from 3 regions with ∼4,000-118,000 dives per region. We show that mobulids as a group, which includes eleven species, have globally and regionally restricted distributions, typically have low sighting frequency (<1% of dives) and aggregate in only a few locations. Of the regions surveyed by divers, almost half (47%) report declining mobulid sightings over the last decade. Divers indicate that although mobulid ecotourism occurs in many regions (45% of those reported, n = 41) they are considered protected in only 32% of the regions. Mobulids being fished or sold in local markets were reported from 16% and 12% of regions, respectively, with most being adjacent to mobulid abundance hotspot and ecotourism regions (e.g. Sri Lanka, Indonesia, east Africa). Identification of regions where ecotourism and exploitation are at odds could help prioritize conservation efforts. Vulnerability analysis, using life history characteristics, indicates that Manta spp. are vulnerable to exploitation, tolerating only low fishing mortality rates; data limitations prohibited such analysis for Mobula spp. Our analyses support previous studies in showing the need for improved conservation and monitoring efforts, and suggest that international and enforceable management policies are required to prevent further population decline.
Global Population Trends and Human Use Patterns of Manta and Mobula Rays
Ward-Paige, Christine A.; Davis, Brendal; Worm, Boris
2013-01-01
Despite being the world’s largest rays and providing significant revenue through dive tourism, little is known about the population status, exploitation and trade volume of the Mobulidae (mobulids; Manta and Mobula spp.). There is anecdotal evidence, however, that mobulid populations are declining, largely due to the recent emergence of a widespread trade for their gill rakers, which is reflected in increasing Food and Agriculture Organization landings trends. Here, we present results from two dedicated diver surveys, one from the eManta project, which includes summary observations from ninety 10°x10° regions with ∼200–62,000 dives per region, and the other from the Reef Environmental Education Foundation, which includes spatially more detailed observations from 3 regions with ∼4,000–118,000 dives per region. We show that mobulids as a group, which includes eleven species, have globally and regionally restricted distributions, typically have low sighting frequency (<1% of dives) and aggregate in only a few locations. Of the regions surveyed by divers, almost half (47%) report declining mobulid sightings over the last decade. Divers indicate that although mobulid ecotourism occurs in many regions (45% of those reported, n = 41) they are considered protected in only 32% of the regions. Mobulids being fished or sold in local markets were reported from 16% and 12% of regions, respectively, with most being adjacent to mobulid abundance hotspot and ecotourism regions (e.g. Sri Lanka, Indonesia, east Africa). Identification of regions where ecotourism and exploitation are at odds could help prioritize conservation efforts. Vulnerability analysis, using life history characteristics, indicates that Manta spp. are vulnerable to exploitation, tolerating only low fishing mortality rates; data limitations prohibited such analysis for Mobula spp. Our analyses support previous studies in showing the need for improved conservation and monitoring efforts, and suggest that international and enforceable management policies are required to prevent further population decline. PMID:24040348
Scientific diving plays an important role in helping EPA protect our oceans and waterways. EPA's divers set a high standard for safety and operational procedures in dangerous polluted water conditions.