Sample records for ireland glacial refugia

  1. Phylogeographic insights into cryptic glacial refugia.

    PubMed

    Provan, Jim; Bennett, K D

    2008-10-01

    The glacial episodes of the Quaternary (2.6 million years ago-present) were a major factor in shaping the present-day distributions of extant flora and fauna, with expansions and contractions of the ice sheets rendering large areas uninhabitable for most species. Fossil records suggest that many species survived glacial maxima by retreating to refugia, usually at lower latitudes. Recently, phylogeographic studies have given support to the existence of previously unknown, or cryptic, refugia. Here we summarise many of these insights into the glacial histories of species in cryptic refugia gained through phylogeographic approaches. Understanding such refugia might be important as the Earth heads into another period of climate change, in terms of predicting the effects on species distribution and survival.

  2. Glacial refugia and recolonization pathways in the brown seaweed Fucus serratus.

    PubMed

    Hoarau, G; Coyer, J A; Veldsink, J H; Stam, W T; Olsen, J L

    2007-09-01

    The last glacial maximum (20,000-18,000 years ago) dramatically affected extant distributions of virtually all northern European biota. Locations of refugia and postglacial recolonization pathways were examined in Fucus serratus (Heterokontophyta; Fucaceae) using a highly variable intergenic spacer developed from the complete mitochondrial genome of Fucus vesiculosus. Over 1,500 samples from the entire range of F. serratus were analysed using fluorescent single strand conformation polymorphism. A total of 28 mtDNA haplotypes was identified and sequenced. Three refugia were recognized based on high haplotype diversities and the presence of endemic haplotypes: southwest Ireland, the northern Brittany-Hurd Deep area of the English Channel, and the northwest Iberian Peninsula. The Irish refugium was the source for a recolonization sweep involving a single haplotype via northern Scotland and throughout Scandinavia, whereas recolonization from the Brittany-Hurd Deep refugium was more limited, probably because of unsuitable soft-bottom habitat in the Bay of Biscay and along the Belgian and Dutch coasts. The Iberian populations reflect a remnant refugium at the present-day southern boundary of the species range. A generalized skyline plot suggested exponential population expansion beginning in the mid-Pleistocene with maximal growth during the Eems interglacial 128,000-67,000 years ago, implying that the last glacial maximum mainly shaped population distributions rather than demography.

  3. Glacial refugia, recolonization patterns and diversification forces in Alpine-endemic Megabunus harvestmen.

    PubMed

    Wachter, Gregor A; Papadopoulou, Anna; Muster, Christoph; Arthofer, Wolfgang; Knowles, L Lacey; Steiner, Florian M; Schlick-Steiner, Birgit C

    2016-06-01

    The Pleistocene climatic fluctuations had a huge impact on all life forms, and various hypotheses regarding the survival of organisms during glacial periods have been postulated. In the European Alps, evidence has been found in support of refugia outside the ice shield (massifs de refuge) acting as sources for postglacial recolonization of inner-Alpine areas. In contrast, evidence for survival on nunataks, ice-free areas above the glacier, remains scarce. Here, we combine multivariate genetic analyses with ecological niche models (ENMs) through multiple timescales to elucidate the history of Alpine Megabunus harvestmen throughout the ice ages, a genus that comprises eight high-altitude endemics. ENMs suggest two types of refugia throughout the last glacial maximum, inner-Alpine survival on nunataks for four species and peripheral refugia for further four species. In some geographic regions, the patterns of genetic variation are consistent with long-distance dispersal out of massifs de refuge, repeatedly coupled with geographic parthenogenesis. In other regions, long-term persistence in nunataks may dominate the patterns of genetic divergence. Overall, our results suggest that glacial cycles contributed to allopatric diversification in Alpine Megabunus, both within and at the margins of the ice shield. These findings exemplify the power of ENM projections coupled with genetic analyses to identify hypotheses about the position and the number of glacial refugia and thus to evaluate the role of Pleistocene glaciations in driving species-specific responses of recolonization or persistence that may have contributed to observed patterns of biodiversity. © 2016 John Wiley & Sons Ltd.

  4. Could brown bears (Ursus arctos) have survived in Ireland during the Last Glacial Maximum?

    PubMed Central

    Leonard, Saoirse A.; Risley, Claire L.; Turvey, Samuel T.

    2013-01-01

    Brown bears are recorded from Ireland during both the Late Pleistocene and early–mid Holocene. Although most of the Irish landmass was covered by an ice sheet during the Last Glacial Maximum (LGM), Irish brown bears are known to have hybridized with polar bears during the Late Pleistocene, and it is suggested that the Irish brown bear population did not become extinct but instead persisted in situ through the LGM in a southwestern ice-free refugium. We use historical population modelling to demonstrate that brown bears are highly unlikely to have survived through the LGM in Ireland under any combination of life-history parameters shown by living bear populations, but instead would have rapidly become extinct following advance of the British–Irish ice sheet, and probably recolonized Ireland during the end-Pleistocene Woodgrange Interstadial from a closely related nearby source population. The time available for brown bear–polar bear hybridization was therefore restricted to narrow periods at the beginning or end of the LGM. Brown bears would have been extremely vulnerable to extinction in Quaternary habitat refugia and required areas substantially larger than southwestern Ireland to survive adverse glacial conditions. PMID:23676655

  5. Could brown bears (Ursus arctos) have survived in Ireland during the Last Glacial Maximum?

    PubMed

    Leonard, Saoirse A; Risley, Claire L; Turvey, Samuel T

    2013-08-23

    Brown bears are recorded from Ireland during both the Late Pleistocene and early-mid Holocene. Although most of the Irish landmass was covered by an ice sheet during the Last Glacial Maximum (LGM), Irish brown bears are known to have hybridized with polar bears during the Late Pleistocene, and it is suggested that the Irish brown bear population did not become extinct but instead persisted in situ through the LGM in a southwestern ice-free refugium. We use historical population modelling to demonstrate that brown bears are highly unlikely to have survived through the LGM in Ireland under any combination of life-history parameters shown by living bear populations, but instead would have rapidly become extinct following advance of the British-Irish ice sheet, and probably recolonized Ireland during the end-Pleistocene Woodgrange Interstadial from a closely related nearby source population. The time available for brown bear-polar bear hybridization was therefore restricted to narrow periods at the beginning or end of the LGM. Brown bears would have been extremely vulnerable to extinction in Quaternary habitat refugia and required areas substantially larger than southwestern Ireland to survive adverse glacial conditions.

  6. Mitochondrial DNA signals of late glacial recolonization of Europe from near eastern refugia.

    PubMed

    Pala, Maria; Olivieri, Anna; Achilli, Alessandro; Accetturo, Matteo; Metspalu, Ene; Reidla, Maere; Tamm, Erika; Karmin, Monika; Reisberg, Tuuli; Hooshiar Kashani, Baharak; Perego, Ugo A; Carossa, Valeria; Gandini, Francesca; Pereira, Joana B; Soares, Pedro; Angerhofer, Norman; Rychkov, Sergei; Al-Zahery, Nadia; Carelli, Valerio; Sanati, Mohammad Hossein; Houshmand, Massoud; Hatina, Jiři; Macaulay, Vincent; Pereira, Luísa; Woodward, Scott R; Davies, William; Gamble, Clive; Baird, Douglas; Semino, Ornella; Villems, Richard; Torroni, Antonio; Richards, Martin B

    2012-05-04

    Human populations, along with those of many other species, are thought to have contracted into a number of refuge areas at the height of the last Ice Age. European populations are believed to be, to a large extent, the descendants of the inhabitants of these refugia, and some extant mtDNA lineages can be traced to refugia in Franco-Cantabria (haplogroups H1, H3, V, and U5b1), the Italian Peninsula (U5b3), and the East European Plain (U4 and U5a). Parts of the Near East, such as the Levant, were also continuously inhabited throughout the Last Glacial Maximum, but unlike western and eastern Europe, no archaeological or genetic evidence for Late Glacial expansions into Europe from the Near East has hitherto been discovered. Here we report, on the basis of an enlarged whole-genome mitochondrial database, that a substantial, perhaps predominant, signal from mitochondrial haplogroups J and T, previously thought to have spread primarily from the Near East into Europe with the Neolithic population, may in fact reflect dispersals during the Late Glacial period, ∼19-12 thousand years (ka) ago. Copyright © 2012 The American Society of Human Genetics. Published by Elsevier Inc. All rights reserved.

  7. Genome-wide set of SNPs reveals evidence for two glacial refugia and admixture from postglacial recolonization in an alpine ungulate.

    PubMed

    Sim, Zijian; Hall, Jocelyn C; Jex, Bill; Hegel, Troy M; Coltman, David W

    2016-08-01

    Past glaciation events have played a major role in shaping the genetic diversity and distribution of wild sheep in North America. The advancement of glaciers can isolate populations in ice-free refugia, where they can survive until the recession of ice sheets. The major Beringian refugium is thought to have held thinhorn sheep (Ovis dalli) populations during times of glacial advance. While isolation in the major refugium can account for much of the genetic and morphological diversity seen in extant thinhorn sheep populations, mounting evidence suggests the persistence of populations in smaller minor refugia. We investigated the refugial origins of thinhorn sheep using ~10 000 SNPs obtained via a cross-species application of the domestic sheep ovine HD BeadChip to genotype 52 thinhorn sheep and five bighorn sheep (O. canadensis) samples. Phylogenetic inference revealed a distinct lineage of thinhorn sheep inhabiting British Columbia, which is consistent with the survival of a group of thinhorn sheep in a minor refugium separate from the Beringian refugium. Isolation in separate glacial refugia probably mediated the evolution of the two thinhorn sheep subspecies, the white Dall's sheep (O. d. dalli), which persisted in Beringia, and the dark Stone's sheep (O. d. stonei), which utilized the minor refugium. We also found the first genetic evidence for admixture between sheep from different glacial refugia in south-central Yukon as a consequence of post glacial expansion and recolonization. These results show that glaciation events can have a major role in the evolution of species inhabiting previously glaciated habitats and the need to look beyond established refugia when examining the evolutionary history of such species. © 2016 John Wiley & Sons Ltd.

  8. A Complex System of Glacial Sub-Refugia Drives Endemic Freshwater Biodiversity on the Tibetan Plateau.

    PubMed

    Clewing, Catharina; Albrecht, Christian; Wilke, Thomas

    2016-01-01

    Although only relatively few freshwater invertebrate families are reported from the Tibetan Plateau, the degree of endemism may be high. Many endemic lineages occur within permafrost areas, raising questions about the existence of isolated intra-plateau glacial refugia. Moreover, if such refugia existed, it might be instructive to learn whether they were associated with lakes or with more dynamic ecosystems such as ponds, wetlands, or springs. To study these hypotheses, we used pulmonate snails of the plateau-wide distributed genus Radix as model group and the Lake Donggi Cona drainage system, located in the north-eastern part of the plateau, as model site. First, we performed plateau-wide phylogenetic analyses using mtDNA data to assess the overall relationships of Radix populations inhabiting the Lake Donggi Cona system for revealing refugial lineages. We then conducted regional phylogeographical analyses applying a combination of mtDNA and nuclear AFLP markers to infer the local structure and demographic history of the most abundant endemic Radix clade for identifying location and type of (sub-)refugia within the drainage system. Our phylogenetic analysis showed a high diversity of Radix lineages in the Lake Donggi Cona system. Subsequent phylogeographical analyses of the most abundant endemic clade indicated a habitat-related clustering of genotypes and several Late Pleistocene spatial/demographic expansion events. The most parsimonious explanation for these patterns would be a scenario of an intra-plateau glacial refugium in the Lake Donggi Cona drainage system, which might have consisted of isolated sub-refugia. Though the underlying processes remain unknown, an initial separation of lake and watershed populations could have been triggered by lake-level fluctuations before and during the Last Glacial Maximum. This study inferred the first intra-plateau refugium for freshwater animals on the Tibetan Plateau. It thus sheds new light on the evolutionary history

  9. Evidence for cryptic northern refugia in the last glacial period in Cryptomeria japonica

    PubMed Central

    Kimura, Megumi K.; Uchiyama, Kentaro; Nakao, Katsuhiro; Moriguchi, Yoshinari; San Jose-Maldia, Lerma; Tsumura, Yoshihiko

    2014-01-01

    Background and Aims Distribution shifts and natural selection during past climatic changes are important factors in determining the genetic structure of forest species. In particular, climatic fluctuations during the Quaternary appear to have caused changes in the distribution ranges of plants, and thus strongly affected their genetic structure. This study was undertaken to identify the responses of the conifer Cryptomeria japonica, endemic to the Japanese Archipelago, to past climatic changes using a combination of phylogeography and species distribution modelling (SDM) methods. Specifically, this study focused on the locations of refugia during the last glacial maximum (LGM). Methods Genetic diversity and structure were examined using 20 microsatellite markers in 37 populations of C. japonica. The locations of glacial refugia were assessed using STRUCTURE analysis, and potential habitats under current and past climate conditions were predicted using SDM. The process of genetic divergence was also examined using the approximate Bayesian computation procedure (ABC) in DIY ABC to test the divergence time between the gene pools detected by the STRUCTURE analysis. Key Results STRUCTURE analysis identified four gene pools: northern Tohoku district; from Chubu to Chugoku district; from Tohoku to Shikoku district on the Pacific Ocean side of the Archipelago; and Yakushima Island. DIY ABC analysis indicated that the four gene pools diverged at the same time before the LGM. SDM also indicated potential northern cryptic refugia. Conclusions The combined evidence from microsatellites and SDM clearly indicates that climatic changes have shaped the genetic structure of C. japonica. The gene pool detected in northern Tohoku district is likely to have been established by cryptic northern refugia on the coast of the Japan Sea to the west of the Archipelago. The gene pool in Yakushima Island can probably be explained simply by long-term isolation from the other gene pools since

  10. Extra-Mediterranean refugia: The rule and not the exception?

    PubMed Central

    2012-01-01

    Some decades ago, biogeographers distinguished three major faunal types of high importance for Europe: (i) Mediterranean elements with exclusive glacial survival in the Mediterranean refugia, (ii) Siberian elements with glacial refugia in the eastern Palearctic and only postglacial expansion to Europe and (iii) arctic and/or alpine elements with large zonal distributions in the periglacial areas and postglacial retreat to the North and/or into the high mountain systems. Genetic analyses have unravelled numerous additional refugia both of continental and Mediterranean species, thus strongly modifying the biogeographical view of Europe. This modified notion is particularly true for the so-called Siberian species, which in many cases have not immigrated into Europe during the postglacial period, but most likely have survived the last, or even several glacial phases, in extra-Mediterranean refugia in some climatically favourable but geographically limited areas of southern Central and Eastern Europe. Recently, genetic analyses revealed that typical Mediterranean species have also survived the Last Glacial Maximum in cryptic northern refugia (e.g. in the Carpathians or even north of the Alps) in addition to their Mediterranean refuge areas. PMID:22953783

  11. Two California lineages of Oxalis oregana: genetic evidence for a Pleistocene separation into northern and southern glacial refugia

    Treesearch

    Chris Brinegar

    2017-01-01

    In the Pacific Northwest, there are discontinuities in the lineages of several plant and animal species in the northern California/Oregon region that are thought to have their origins in the separation of populations into refugia during the Pleistocene glacial periods. Redwood sorrel (Oxalis oregana Nutt.), a common understory species of the...

  12. Refugia revisited: individualistic responses of species in space and time

    PubMed Central

    Stewart, John R.; Lister, Adrian M.; Barnes, Ian; Dalén, Love

    2010-01-01

    Climate change in the past has led to significant changes in species' distributions. However, how individual species respond to climate change depends largely on their adaptations and environmental tolerances. In the Quaternary, temperate-adapted taxa are in general confined to refugia during glacials while cold-adapted taxa are in refugia during interglacials. In the Northern Hemisphere, evidence appears to be mounting that in addition to traditional southern refugia for temperate species, cryptic refugia existed in the North during glacials. Equivalent cryptic southern refugia, to the south of the more conventional high-latitude polar refugia, exist in montane areas during periods of warm climate, such as the current interglacial. There is also a continental/oceanic longitudinal gradient, which should be included in a more complete consideration of the interaction between species ranges and climates. Overall, it seems clear that there is large variation in both the size of refugia and the duration during which species are confined to them. This has implications for the role of refugia in the evolution of species and their genetic diversity. PMID:19864280

  13. Pleistocene glacial refugia across the Appalachian Mountains and coastal plain in the millipede genus Narceus: Evidence from population genetic, phylogeographic, and paleoclimatic data

    PubMed Central

    Walker, Matt J; Stockman, Amy K; Marek, Paul E; Bond, Jason E

    2009-01-01

    Background Species that are widespread throughout historically glaciated and currently non-glaciated areas provide excellent opportunities to investigate the role of Pleistocene climatic change on the distribution of North American biodiversity. Many studies indicate that northern animal populations exhibit low levels of genetic diversity over geographically widespread areas whereas southern populations exhibit relatively high levels. Recently, paleoclimatic data have been combined with niche-based distribution modeling to locate possible refugia during the Last Glacial Maximum. Using phylogeographic, population, and paleoclimatic data, we show that the distribution and mitochondrial data for the millipede genus Narceus are consistent with classical examples of Pleistocene refugia and subsequent post-glacial population expansion seen in other organismal groups. Results The phylogeographic structure of Narceus reveals a complex evolutionary history with signatures of multiple refugia in southeastern North America followed by two major northern expansions. Evidence for refugial populations were found in the southern Appalachian Mountains and in the coastal plain. The northern expansions appear to have radiated from two separate refugia, one from the Gulf Coastal Plain area and the other from the mid-Atlantic coastal region. Distributional models of Narceus during the Last Glacial Maximum show a dramatic reduction from the current distribution, with suitable ecological zones concentrated along the Gulf and Atlantic coastal plain. We found a strong correlation between these zones of ecological suitability inferred from our paleo-model with levels of genetic diversity derived from phylogenetic and population estimates of genetic structuring. Conclusion The signature of climatic change, during and after the Pleistocene, on the distribution of the millipede genus Narceus is evident in the genetic data presented. Niche-based historical distribution modeling strengthens the

  14. Extra-Mediterranean glacial refugia in a Mediterranean faunal element: the phylogeography of the chalk-hill blue Polyommatus coridon (Lepidoptera, Lycaenidae)

    PubMed Central

    Kühne, Gero; Kosuch, Joachim; Hochkirch, Axel; Schmitt, Thomas

    2017-01-01

    Most warm-adapted Central European species are thought to have survived ice ages exclusively in Mediterranean refugia. During recent years, this point of view has been questioned. Therefore, we tested the hypothesis that extra-Mediterranean refugia also played a role in warm-adapted insect species and selected the chalk-hill blue, Polyommatus coridon. We sequenced two mitochondrial loci (COI, CR) in 150 individuals from 30 populations covering nearly the complete range. Minimum spanning networks and other statistical analyses concordantly revealed four genetic lineages with strong phylogeographic signal: a western group in Italy, France and western/central Germany, an eastern lineage in the Balkan Peninsula, the Carpathian Basin and eastern Central Europe, an Alpine group with populations in the Alps and southern Germany and a Pyrenean group. Our results are generally consistent with previous analyses for P. coridon based on allozymes and DNA sequences, but provide additional insights. We propose that these four lineages have evolved during allopatry in different glacial refugia, two in typical Mediterranean refugia (Apennines and Balkan Peninsulas), but two in extra-Mediterranean areas south of the Alps and Pyrenees. This supports survival of warm-adapted organisms in these regions in close geographic proximity to the refugia of high mountain species.

  15. Species-wide phylogeography of North American mule deer (Odocoileus hemionus): cryptic glacial refugia and postglacial recolonization.

    PubMed

    Latch, Emily K; Heffelfinger, James R; Fike, Jennifer A; Rhodes, Olin E

    2009-04-01

    Quaternary climatic oscillations greatly influenced the present-day population genetic structure of animals and plants. For species with high dispersal and reproductive potential, phylogeographic patterns resulting from historical processes can be cryptic, overshadowed by contemporary processes. Here we report a study of the phylogeography of Odocoileus hemionus, a large, vagile ungulate common throughout western North America. We examined sequence variation of mitochondrial DNA (control region and cytochrome b) within and among 70 natural populations across the entire range of the species. Among the 1766 individual animals surveyed, we recovered 496 haplotypes. Although fine-scale phylogenetic structure was weakly resolved using phylogenetic methods, network analysis clearly revealed the presence of 12 distinct haplogroups. The spatial distribution of haplogroups showed a strong genetic discontinuity between the two morphological types of O. hemionus, mule deer and black-tailed deer, east and west of the Cascade Mountains in the Pacific Northwest. Within the mule deer lineage, we identified several haplogroups that expanded before or during the Last Glacial Maximum, suggesting that mule deer persisted in multiple refugia south of the ice sheets. Patterns of genetic diversity within the black-tailed deer lineage suggest a single refugium along the Pacific Northwest coast, and refute the hypothesis that black-tailed deer persisted in one or more northern refugia. Our data suggest that black-tailed deer recolonized areas in accordance with the pattern of glacial retreat, with initial recolonization northward along a coastal route and secondary recolonization inland.

  16. Glacial Refugia and Future Habitat Coverage of Selected Dactylorhiza Representatives (Orchidaceae)

    PubMed Central

    2015-01-01

    The intensively discussed taxonomic complexity of the Dactylorhiza genus is probably correlated with its migration history during glaciations and interglacial periods. Previous studies on past processes affecting the current distribution of Dactylorhiza species as well as the history of the polyploid complex formation were based only on molecular data. In the present study the ecological niche modeling (ENM) technique was applied in order to describe the distribution of potential refugia for the selected Dactylorhiza representatives during the Last Glacial Maximum. Additionally, future changes in their potential habitat coverage were measured with regard to three various climatic change scenarios. The maximum entropy method was used to create models of suitable niche distribution. A database of Dactylorhiza localities was prepared on the grounds of information collected from literature and data gathered during field works. Our research indicated that the habitats of majority of the studied taxa will decrease by 2080, except for D. incarnata var. incarnata, for which suitable habitats will increase almost two-fold in the global scale. Moreover, the potential habitats of some taxa are located outside their currently known geographical ranges, e.g. the Aleutian Islands, the western slopes of the Rocky Mountains, Newfoundland, southern Greenland and Iceland. ENM analysis did not confirm that the Balkans, central Europe or central Russia served as the most important refugia for individual representatives of the Dactylorhiza incarnata/maculata complex. Our study rather indicated that the Black Sea coast, southern Apennines and Corsica were the main areas characterized by habitats suitable for most of the taxa. PMID:26599630

  17. Tracing glacial refugia of Triturus newts based on mitochondrial DNA phylogeography and species distribution modeling

    PubMed Central

    2013-01-01

    Introduction The major climatic oscillations during the Quaternary Ice Age heavily influenced the distribution of species and left their mark on intraspecific genetic diversity. Past range shifts can be reconstructed with the aid of species distribution modeling and phylogeographical analyses. We test the responses of the different members of the genus Triturus (i.e. the marbled and crested newts) as the climate shifted from the previous glacial period (the Last Glacial Maximum, ~21 Ka) to the current interglacial. Results We present the results of a dense mitochondrial DNA phylogeography (visualizing genetic diversity within and divergence among populations) and species distribution modeling (using two different climate simulations) for the nine Triturus species on composite maps. Conclusions The combined use of species distribution modeling and mitochondrial phylogeography provides insight in the glacial contraction and postglacial expansion of Triturus. The combined use of the two independent techniques yields a more complete understanding of the historical biogeography of Triturus than both approaches would on their own. Triturus newts generally conform to the ‘southern richness and northern purity’ paradigm, but we also find more intricate patterns, such as the absence of genetic variation and suitable area at the Last Glacial Maximum (T. dobrogicus), an ‘extra-Mediterranean’ refugium in the Carpathian Basin (T. cristatus), and areas where species displaced one another postglacially (e.g. T. macedonicus and western T. karelinii). We provide a biogeographical scenario for Triturus, showing the positions of glacial refugia, the regions that were postglacially colonized and the areas where species displaced one another as they shifted their ranges. PMID:23514662

  18. Tracing glacial refugia of Triturus newts based on mitochondrial DNA phylogeography and species distribution modeling.

    PubMed

    Wielstra, Ben; Crnobrnja-Isailović, Jelka; Litvinchuk, Spartak N; Reijnen, Bastian T; Skidmore, Andrew K; Sotiropoulos, Konstantinos; Toxopeus, Albertus G; Tzankov, Nikolay; Vukov, Tanja; Arntzen, Jan W

    2013-03-20

    The major climatic oscillations during the Quaternary Ice Age heavily influenced the distribution of species and left their mark on intraspecific genetic diversity. Past range shifts can be reconstructed with the aid of species distribution modeling and phylogeographical analyses. We test the responses of the different members of the genus Triturus (i.e. the marbled and crested newts) as the climate shifted from the previous glacial period (the Last Glacial Maximum, ~21 Ka) to the current interglacial. We present the results of a dense mitochondrial DNA phylogeography (visualizing genetic diversity within and divergence among populations) and species distribution modeling (using two different climate simulations) for the nine Triturus species on composite maps. The combined use of species distribution modeling and mitochondrial phylogeography provides insight in the glacial contraction and postglacial expansion of Triturus. The combined use of the two independent techniques yields a more complete understanding of the historical biogeography of Triturus than both approaches would on their own. Triturus newts generally conform to the 'southern richness and northern purity' paradigm, but we also find more intricate patterns, such as the absence of genetic variation and suitable area at the Last Glacial Maximum (T. dobrogicus), an 'extra-Mediterranean' refugium in the Carpathian Basin (T. cristatus), and areas where species displaced one another postglacially (e.g. T. macedonicus and western T. karelinii). We provide a biogeographical scenario for Triturus, showing the positions of glacial refugia, the regions that were postglacially colonized and the areas where species displaced one another as they shifted their ranges.

  19. Genetic signature of Last Glacial Maximum regional refugia in a circum-Antarctic sea spider

    PubMed Central

    Soler-Membrives, Anna; Linse, Katrin; Miller, Karen J.

    2017-01-01

    The evolutionary history of Antarctic organisms is becoming increasingly important to understand and manage population trajectories under rapid environmental change. The Antarctic sea spider Nymphon australe, with an apparently large population size compared with other sea spider species, is an ideal target to look for molecular signatures of past climatic events. We analysed mitochondrial DNA of specimens collected from the Antarctic continent and two Antarctic islands (AI) to infer past population processes and understand current genetic structure. Demographic history analyses suggest populations survived in refugia during the Last Glacial Maximum. The high genetic diversity found in the Antarctic Peninsula and East Antarctic (EA) seems related to multiple demographic contraction–expansion events associated with deep-sea refugia, while the low genetic diversity in the Weddell Sea points to a more recent expansion from a shelf refugium. We suggest the genetic structure of N. australe from AI reflects recent colonization from the continent. At a local level, EA populations reveal generally low genetic differentiation, geographically and bathymetrically, suggesting limited restrictions to dispersal. Results highlight regional differences in demographic histories and how these relate to the variation in intensity of glaciation–deglaciation events around Antarctica, critical for the study of local evolutionary processes. These are valuable data for understanding the remarkable success of Antarctic pycnogonids, and how environmental changes have shaped the evolution and diversification of Southern Ocean benthic biodiversity. PMID:29134072

  20. Genetic signature of Last Glacial Maximum regional refugia in a circum-Antarctic sea spider

    NASA Astrophysics Data System (ADS)

    Soler-Membrives, Anna; Linse, Katrin; Miller, Karen J.; Arango, Claudia P.

    2017-10-01

    The evolutionary history of Antarctic organisms is becoming increasingly important to understand and manage population trajectories under rapid environmental change. The Antarctic sea spider Nymphon australe, with an apparently large population size compared with other sea spider species, is an ideal target to look for molecular signatures of past climatic events. We analysed mitochondrial DNA of specimens collected from the Antarctic continent and two Antarctic islands (AI) to infer past population processes and understand current genetic structure. Demographic history analyses suggest populations survived in refugia during the Last Glacial Maximum. The high genetic diversity found in the Antarctic Peninsula and East Antarctic (EA) seems related to multiple demographic contraction-expansion events associated with deep-sea refugia, while the low genetic diversity in the Weddell Sea points to a more recent expansion from a shelf refugium. We suggest the genetic structure of N. australe from AI reflects recent colonization from the continent. At a local level, EA populations reveal generally low genetic differentiation, geographically and bathymetrically, suggesting limited restrictions to dispersal. Results highlight regional differences in demographic histories and how these relate to the variation in intensity of glaciation-deglaciation events around Antarctica, critical for the study of local evolutionary processes. These are valuable data for understanding the remarkable success of Antarctic pycnogonids, and how environmental changes have shaped the evolution and diversification of Southern Ocean benthic biodiversity.

  1. Hidden levels of phylodiversity in Antarctic green algae: further evidence for the existence of glacial refugia

    PubMed Central

    De Wever, Aaike; Leliaert, Frederik; Verleyen, Elie; Vanormelingen, Pieter; Van der Gucht, Katleen; Hodgson, Dominic A.; Sabbe, Koen; Vyverman, Wim

    2009-01-01

    Recent data revealed that metazoans such as mites and springtails have persisted in Antarctica throughout several glacial–interglacial cycles, which contradicts the existing paradigm that terrestrial life was wiped out by successive glacial events and that the current inhabitants are recent colonizers. We used molecular phylogenetic techniques to study Antarctic microchlorophyte strains isolated from lacustrine habitats from maritime and continental Antarctica. The 14 distinct chlorophycean and trebouxiophycean lineages observed point to a wide phylogenetic diversity of apparently endemic Antarctic lineages at different taxonomic levels. This supports the hypothesis that long-term survival took place in glacial refugia, resulting in a specific Antarctic flora. The majority of the lineages have estimated ages between 17 and 84 Ma and probably diverged from their closest relatives around the time of the opening of Drake Passage (30–45 Ma), while some lineages with longer branch lengths have estimated ages that precede the break-up of Gondwana. The variation in branch length and estimated age points to several independent but rare colonization events. PMID:19625320

  2. Phylogeography of the Patagonian otter Lontra provocax: adaptive divergence to marine habitat or signature of southern glacial refugia?

    PubMed Central

    2011-01-01

    Background A number of studies have described the extension of ice cover in western Patagonia during the Last Glacial Maximum, providing evidence of a complete cover of terrestrial habitat from 41°S to 56°S and two main refugia, one in south-eastern Tierra del Fuego and the other north of the Chiloé Island. However, recent evidence of high genetic diversity in Patagonian river species suggests the existence of aquatic refugia in this region. Here, we further test this hypothesis based on phylogeographic inferences from a semi-aquatic species that is a top predator of river and marine fauna, the huillín or Southern river otter (Lontra provocax). Results We examined mtDNA sequences of the control region, ND5 and Cytochrome-b (2151 bp in total) in 75 samples of L. provocax from 21 locations in river and marine habitats. Phylogenetic analysis illustrates two main divergent clades for L. provocax in continental freshwater habitat. A highly diverse clade was represented by haplotypes from the marine habitat of the Southern Fjords and Channels (SFC) region (43°38' to 53°08'S), whereas only one of these haplotypes was paraphyletic and associated with northern river haplotypes. Conclusions Our data support the hypothesis of the persistence of L. provocax in western Patagonia, south of the ice sheet limit, during last glacial maximum (41°S latitude). This limit also corresponds to a strong environmental change, which might have spurred L. provocax differentiation between the two environments. PMID:21356052

  3. Glacial Refugia in Pathogens: European Genetic Structure of Anther Smut Pathogens on Silene latifolia and Silene dioica

    PubMed Central

    Vercken, Elodie; Fontaine, Michael C.; Gladieux, Pierre; Hood, Michael E.; Jonot, Odile; Giraud, Tatiana

    2010-01-01

    Climate warming is predicted to increase the frequency of invasions by pathogens and to cause the large-scale redistribution of native host species, with dramatic consequences on the health of domesticated and wild populations of plants and animals. The study of historic range shifts in response to climate change, such as during interglacial cycles, can help in the prediction of the routes and dynamics of infectious diseases during the impending ecosystem changes. Here we studied the population structure in Europe of two Microbotryum species causing anther smut disease on the plants Silene latifolia and Silene dioica. Clustering analyses revealed the existence of genetically distinct groups for the pathogen on S. latifolia, providing a clear-cut example of European phylogeography reflecting recolonization from southern refugia after glaciation. The pathogen genetic structure was congruent with the genetic structure of its host species S. latifolia, suggesting dependence of the migration pathway of the anther smut fungus on its host. The fungus, however, appeared to have persisted in more numerous and smaller refugia than its host and to have experienced fewer events of large-scale dispersal. The anther smut pathogen on S. dioica also showed a strong phylogeographic structure that might be related to more northern glacial refugia. Differences in host ecology probably played a role in these differences in the pathogen population structure. Very high selfing rates were inferred in both fungal species, explaining the low levels of admixture between the genetic clusters. The systems studied here indicate that migration patterns caused by climate change can be expected to include pathogen invasions that follow the redistribution of their host species at continental scales, but also that the recolonization by pathogens is not simply a mirror of their hosts, even for obligate biotrophs, and that the ecology of hosts and pathogen mating systems likely affects recolonization

  4. The rare peat moss Sphagnum wulfianum (Sphagnaceae) did not survive the last glacial period in northern European refugia.

    PubMed

    Kyrkjeeide, Magni Olsen; Hassel, Kristian; Flatberg, Kjell I; Stenøien, Hans K

    2012-04-01

    Organisms may survive unfavorable conditions either by moving to more favorable areas by means of dispersal or by adapting to stressful environments. Pleistocene glacial periods represent extremely unfavorable conditions for the majority of life forms, especially sessile organisms. Many studies have revealed placements of refugial areas and postglacial colonization patterns of seed plants, but little is still known about areas of long-term survival and historical migration routes of bryophytes. Given overall differences in stress tolerance between seed plants and bryophytes, it is of interest to know whether bryophytes have survived periods of extreme climatic conditions better then seed plants in northern areas. The haploid and rarely spore-producing peat moss Sphagnum wulfianum is mostly found in areas that were covered by ice during the last glacial maximum. Twelve microsatellite markers were amplified from 43 populations (367 shoots) of this species, and data were analyzed using population genetic diversity statistics, Bayesian clustering methods, and coalescence-based inference tools to estimate historical and demographic parameters. Genetic diversity within populations was low, but populations were highly differentiated, with two main genetic clusters being recognized. The two main genetic groups have diverged quite recently in the Holocene, and the pattern of genetic variability and structuring gives no support for survival in Scandinavian refugia during the last glacial period in this species. The dispersal ability of this plant thus seems surprisingly high despite its infrequent spore production.

  5. Mapping Post-Glacial expansions: The Peopling of Southwest Asia

    PubMed Central

    Platt, Daniel E.; Haber, Marc; Dagher-Kharrat, Magda Bou; Douaihy, Bouchra; Khazen, Georges; Ashrafian Bonab, Maziar; Salloum, Angélique; Mouzaya, Francis; Luiselli, Donata; Tyler-Smith, Chris; Renfrew, Colin; Matisoo-Smith, Elizabeth; Zalloua, Pierre A.

    2017-01-01

    Archaeological, palaeontological and geological evidence shows that post-glacial warming released human populations from their various climate-bound refugia. Yet specific connections between these refugia and the timing and routes of post-glacial migrations that ultimately established modern patterns of genetic variation remain elusive. Here, we use Y-chromosome markers combined with autosomal data to reconstruct population expansions from regional refugia in Southwest Asia. Populations from three regions in particular possess distinctive autosomal genetic signatures indicative of likely refugia: one, in the north, centered around the eastern coast of the Black Sea, the second, with a more Levantine focus, and the third in the southern Arabian Peninsula. Modern populations from these three regions carry the widest diversity and may indeed represent the most likely descendants of the populations responsible for the Neolithic cultures of Southwest Asia. We reveal the distinct and datable expansion routes of populations from these three refugia throughout Southwest Asia and into Europe and North Africa and discuss the possible correlations of these migrations to various cultural and climatic events evident in the archaeological record of the past 15,000 years. PMID:28059138

  6. Genetic and ecological insights into glacial refugia of walnut (Juglans regia L.)

    PubMed Central

    Aradhya, Mallikarjuna; Ibrahimov, Zakir; Toktoraliev, Biimyrza; Maghradze, David; Musayev, Mirza; Bobokashvili, Zviadi; Preece, John E.

    2017-01-01

    The distribution and survival of trees during the last glacial maximum (LGM) has been of interest to paleoecologists, biogeographers, and geneticists. Ecological niche models that associate species occurrence and abundance with climatic variables are widely used to gain ecological and evolutionary insights and to predict species distributions over space and time. The present study deals with the glacial history of walnut to address questions related to past distributions through genetic analysis and ecological modeling of the present, LGM and Last Interglacial (LIG) periods. A maximum entropy method was used to project the current walnut distribution model on to the LGM (21–18 kyr BP) and LIG (130–116 kyr BP) climatic conditions. Model tuning identified the walnut data set filtered at 10 km spatial resolution as the best for modeling the current distribution and to hindcast past (LGM and LIG) distributions of walnut. The current distribution model predicted southern Caucasus, parts of West and Central Asia extending into South Asia encompassing northern Afghanistan, Pakistan, northwestern Himalayan region, and southwestern Tibet, as the favorable climatic niche matching the modern distribution of walnut. The hindcast of distributions suggested the occurrence of walnut during LGM was somewhat limited to southern latitudes from southern Caucasus, Central and South Asian regions extending into southwestern Tibet, northeastern India, Himalayan region of Sikkim and Bhutan, and southeastern China. Both CCSM and MIROC projections overlapped, except that MIROC projected a significant presence of walnut in the Balkan Peninsula during the LGM. In contrast, genetic analysis of the current walnut distribution suggested a much narrower area in northern Pakistan and the surrounding areas of Afghanistan, northwestern India, and southern Tajikistan as a plausible hotspot of diversity where walnut may have survived glaciations. Overall, the findings suggest that walnut perhaps

  7. Genetic and ecological insights into glacial refugia of walnut (Juglans regia L.).

    PubMed

    Aradhya, Mallikarjuna; Velasco, Dianne; Ibrahimov, Zakir; Toktoraliev, Biimyrza; Maghradze, David; Musayev, Mirza; Bobokashvili, Zviadi; Preece, John E

    2017-01-01

    The distribution and survival of trees during the last glacial maximum (LGM) has been of interest to paleoecologists, biogeographers, and geneticists. Ecological niche models that associate species occurrence and abundance with climatic variables are widely used to gain ecological and evolutionary insights and to predict species distributions over space and time. The present study deals with the glacial history of walnut to address questions related to past distributions through genetic analysis and ecological modeling of the present, LGM and Last Interglacial (LIG) periods. A maximum entropy method was used to project the current walnut distribution model on to the LGM (21-18 kyr BP) and LIG (130-116 kyr BP) climatic conditions. Model tuning identified the walnut data set filtered at 10 km spatial resolution as the best for modeling the current distribution and to hindcast past (LGM and LIG) distributions of walnut. The current distribution model predicted southern Caucasus, parts of West and Central Asia extending into South Asia encompassing northern Afghanistan, Pakistan, northwestern Himalayan region, and southwestern Tibet, as the favorable climatic niche matching the modern distribution of walnut. The hindcast of distributions suggested the occurrence of walnut during LGM was somewhat limited to southern latitudes from southern Caucasus, Central and South Asian regions extending into southwestern Tibet, northeastern India, Himalayan region of Sikkim and Bhutan, and southeastern China. Both CCSM and MIROC projections overlapped, except that MIROC projected a significant presence of walnut in the Balkan Peninsula during the LGM. In contrast, genetic analysis of the current walnut distribution suggested a much narrower area in northern Pakistan and the surrounding areas of Afghanistan, northwestern India, and southern Tajikistan as a plausible hotspot of diversity where walnut may have survived glaciations. Overall, the findings suggest that walnut perhaps

  8. Thriving in the Cold: Glacial Expansion and Post-Glacial Contraction of a Temperate Terrestrial Salamander (Plethodon serratus)

    PubMed Central

    Newman, Catherine E.; Austin, Christopher C.

    2015-01-01

    The dynamic geologic history of the southeastern United States has played a major role in shaping the geographic distributions of amphibians in the region. In the phylogeographic literature, the predominant pattern of distribution shifts through time of temperate species is one of contraction during glacial maxima and persistence in refugia. However, the diverse biology and ecology of amphibian species suggest that a “one-size-fits-all” model may be inappropriate. Nearly 10% of amphibian species in the region have a current distribution comprised of multiple disjunct, restricted areas that resemble the shape of Pleistocene refugia identified for other temperate taxa in the literature. Here, we apply genetics and spatially explicit climate analyses to test the hypothesis that the disjunct regions of these species ranges are climatic refugia for species that were more broadly distributed during glacial maxima. We use the salamander Plethodon serratus as a model, as its range consists of four disjunct regions in the Southeast. Phylogenetic results show that P. serratus is comprised of multiple genetic lineages, and the four regions are not reciprocally monophyletic. The Appalachian salamanders form a clade sister to all other P. serratus. Niche and paleodistribution modeling results suggest that P. serratus expanded from the Appalachians during the cooler Last Glacial Maximum and has since been restricted to its current disjunct distribution by a warming climate. These data reject the universal applicability of the glacial contraction model to temperate taxa and reiterate the importance of considering the natural history of individual species. PMID:26132077

  9. Continuity of brown bear maternal lineages in northern England through the Last-glacial period

    NASA Astrophysics Data System (ADS)

    Edwards, Ceiridwen J.; Ho, Simon Y. W.; Barnett, Ross; Coxon, Peter; Bradley, Daniel G.; Lord, Tom C.; O'Connor, Terry

    2014-07-01

    Brown bears recolonised Europe rapidly after the Last Glacial Maximum (LGM), but there has been debate about whether bear populations were confined to separate glacial refugia in southern Europe, or if there was continuous gene flow among groups. To look in more detail at recolonisation routes into the British Isles after the LGM, 16 brown bear (Ursus arctos) samples from Lateglacial Yorkshire were analysed for mitochondrial DNA survival. The resulting data were compared with earlier work on Late Pleistocene and Holocene bears from Ireland (Edwards et al., 2011), as well as with both modern and ancient bears from across continental Europe. The results highlight the temporal and spatial continuity of brown bear maternal lineages through the Lateglacial period in northern England. While this region was not a refugial area in the LGM for the Irish Clade 2 brown bears, our data suggest that populations of brown bear in England did act as refugial sources for the later colonisation of Ireland, by Clade 1-i bears, during the Holocene. Our results contribute to a wider understanding of the phylogenetic relationships of brown bears through the Late Quaternary, and lend a valuable perspective on bear migration into peripheral Europe.

  10. Glacial vicariance in Eurasia: mitochondrial DNA evidence from Scots pine for a complex heritage involving genetically distinct refugia at mid-northern latitudes and in Asia Minor

    PubMed Central

    Naydenov, Krassimir; Senneville, Sauphie; Beaulieu, Jean; Tremblay, Francine; Bousquet, Jean

    2007-01-01

    Background At the last glacial maximum, Fennoscandia was covered by an ice sheet while the tundra occupied most of the rest of northern Eurasia. More or less disjunct refugial populations of plants were dispersed in southern Europe, often trapped between mountain ranges and seas. Genetic and paleobotanical evidences indicate that these populations have contributed much to Holocene recolonization of more northern latitudes. Less supportive evidence has been found for the existence of glacial populations located closer to the ice margin. Scots pine (Pinus sylvestris L.) is a nordic conifer with a wide natural range covering much of Eurasia. Fractures in its extant genetic structure might be indicative of glacial vicariance and how different refugia contributed to the current distribution at the continental level. The population structure of Scots pine was investigated on much of its Eurasian natural range using maternally inherited mitochondrial DNA polymorphisms. Results A novel polymorphic region of the Scots pine mitochondrial genome has been identified, the intron 1 of nad7, with three variants caused by insertions-deletions. From 986 trees distributed among 54 populations, four distinct multi-locus mitochondrial haplotypes (mitotypes) were detected based on the three nad7 intron 1 haplotypes and two previously reported size variants for nad1 intron B/C. Population differentiation was high (GST = 0.657) and the distribution of the mitotypes was geographically highly structured, suggesting at least four genetically distinct ancestral lineages. A cosmopolitan lineage was widely distributed in much of Europe throughout eastern Asia. A previously reported lineage limited to the Iberian Peninsula was confirmed. A new geographically restricted lineage was found confined to Asia Minor. A new lineage was restricted to more northern latitudes in northeastern Europe and the Baltic region. Conclusion The contribution of the various ancestral lineages to the current

  11. Last Glacial-Interglacial Transition ice dynamics in the Wicklow Mountains, Ireland

    NASA Astrophysics Data System (ADS)

    Knight, Lauren; Boston, Clare; Lovell, Harold; Pepin, Nick

    2017-04-01

    Understanding of the extent and dynamics of former ice masses in the Wicklow Mountains, Ireland, during the Last Glacial-Interglacial Transition (LGIT; 15-10 ka BP) is currently unresolved. Whilst it is acknowledged that the region hosted a local ice cap within the larger British-Irish Ice Sheet at the Last Glacial Maximum (LGM; 27 ka BP), there has been little consideration of ice cap disintegration to a topographically constrained ice mass during the LGIT. This research has produced the first regional glacial geomorphological map, through remote sensing (aerial photograph and digital terrain model interrogation) and field mapping. This has allowed both the style and extent of mountain glaciation and ice recession dynamics during the LGIT to be established. This geomorphological mapping has highlighted that evidence for local glaciation in the Wicklow Mountains is more extensive than previously recognised, and that small icefields and associated outlet valley glaciers existed during the LGIT following disintegration of the Wicklow Ice Cap. A relative chronology based on morphostratigraphic principles is developed, which indicates complex patterns of ice mass oscillation characterised by periods of both sustained retreat and minor readvance. Variations in the pattern of recession across the Wicklow Mountains are evident and appear to be influenced, in part, by topographic controls (e.g. slope, aspect, glacier hypsometry). In summary, this research establishes a relative chronology of glacial events in the region during the LGIT and presents constraints on ice mass extent, dynamics and retreat patterns, offering an insight into small ice mass behaviour in a warming climate.

  12. Widespread Secondary Contact and New Glacial Refugia in the Halophilic Rotifer Brachionus plicatilis in the Iberian Peninsula

    PubMed Central

    Campillo, Sergi; Serra, Manuel; Carmona, María José; Gómez, Africa

    2011-01-01

    Small aquatic organisms harbour deep phylogeographic patterns and highly structured populations even at local scales. These patterns indicate restricted gene flow, despite these organisms' high dispersal abilities, and have been explained by a combination of (1) strong founder effects due to rapidly growing populations and very large population sizes, and (2) the development of diapausing egg banks and local adaptation, resulting in low effective gene flow, what is known as the Monopolization hypothesis. In this study, we build up on our understanding of the mitochondrial phylogeography of the halophilic rotifer Brachionus plicatilis in the Iberian Peninsula by both increasing the number of sampled ponds in areas where secondary contact is likely and doubling sample sizes. We analyzed partial mitochondrial sequences of 252 individuals. We found two deep mitochondrial DNA lineages differing in both their genetic diversity and the complexity of their phylogeographic structure. Our analyses suggest that several events of secondary contact between clades occurred after their expansion from glacial refugia. We found a pattern of isolation-by-distance, which we interpret as being the result of historical colonization events. We propose the existence of at least one glacial refugium in the SE of the Iberian Peninsula. Our findings challenge predictions of the Monopolization hypothesis, since coexistence (i.e., secondary contact) of divergent lineages in some ponds in the Iberian Peninsula is common. Our results indicate that phylogeographic structures in small organisms can be very complex and that gene flow between diverse lineages after population establishment can indeed occur. PMID:21698199

  13. Widespread secondary contact and new glacial refugia in the halophilic rotifer Brachionus plicatilis in the Iberian Peninsula.

    PubMed

    Campillo, Sergi; Serra, Manuel; Carmona, María José; Gómez, Africa

    2011-01-01

    Small aquatic organisms harbour deep phylogeographic patterns and highly structured populations even at local scales. These patterns indicate restricted gene flow, despite these organisms' high dispersal abilities, and have been explained by a combination of (1) strong founder effects due to rapidly growing populations and very large population sizes, and (2) the development of diapausing egg banks and local adaptation, resulting in low effective gene flow, what is known as the Monopolization hypothesis. In this study, we build up on our understanding of the mitochondrial phylogeography of the halophilic rotifer Brachionus plicatilis in the Iberian Peninsula by both increasing the number of sampled ponds in areas where secondary contact is likely and doubling sample sizes. We analyzed partial mitochondrial sequences of 252 individuals. We found two deep mitochondrial DNA lineages differing in both their genetic diversity and the complexity of their phylogeographic structure. Our analyses suggest that several events of secondary contact between clades occurred after their expansion from glacial refugia. We found a pattern of isolation-by-distance, which we interpret as being the result of historical colonization events. We propose the existence of at least one glacial refugium in the SE of the Iberian Peninsula. Our findings challenge predictions of the Monopolization hypothesis, since coexistence (i.e., secondary contact) of divergent lineages in some ponds in the Iberian Peninsula is common. Our results indicate that phylogeographic structures in small organisms can be very complex and that gene flow between diverse lineages after population establishment can indeed occur.

  14. Phylogeographic Analyses of American Black Bears (Ursus americanus) Suggest Four Glacial Refugia and Complex Patterns of Postglacial Admixture.

    PubMed

    Puckett, Emily E; Etter, Paul D; Johnson, Eric A; Eggert, Lori S

    2015-09-01

    Studies of species with continental distributions continue to identify intraspecific lineages despite continuous habitat. Lineages may form due to isolation by distance, adaptation, divergence across barriers, or genetic drift following range expansion. We investigated lineage diversification and admixture within American black bears (Ursus americanus) across their range using 22 k single nucleotide polymorphisms and mitochondrial DNA sequences. We identified three subcontinental nuclear clusters which we further divided into nine geographic regions: Alaskan (Alaska-East), eastern (Central Interior Highlands, Great Lakes, Northeast, Southeast), and western (Alaska-West, West, Pacific Coast, Southwest). We estimated that the western cluster diverged 67 ka, before eastern and Alaskan divergence 31 ka; these divergence dates contrasted with those from the mitochondrial genome where clades A and B diverged 1.07 Ma, and clades A-east and A-west diverged 169 ka. We combined estimates of divergence timing with hindcast species distribution models to infer glacial refugia for the species in Beringia, Pacific Northwest, Southwest, and Southeast. Our results show a complex arrangement of admixture due to expansion out of multiple refugia. The delineation of the genomic population clusters was inconsistent with the ranges for 16 previously described subspecies. Ranges for U. a. pugnax and U. a. cinnamomum were concordant with admixed clusters, calling into question how to order taxa below the species level. Additionally, our finding that U. a. floridanus has not diverged from U. a. americanus also suggests that morphology and genetics should be reanalyzed to assess taxonomic designations relevant to the conservation management of the species. © The Author 2015. Published by Oxford University Press on behalf of the Society for Molecular Biology and Evolution. All rights reserved.For permissions please email: journals.permissions@oup.com.

  15. Displaced phylogeographic signals from Gyrodactylus arcuatus, a parasite of the three-spined stickleback Gasterosteus aculeatus, suggest freshwater glacial refugia in Europe

    USGS Publications Warehouse

    Lumme, Jaakko; Mäkinen, Hannu; Ermolenko, Alexey V.; Gregg, Jacob L.; Ziętara, Marek S.

    2016-01-01

    We examined the global mitochondrial phylogeography of Gyrodactylus arcuatus, a flatworm ectoparasite of three-spined stickleback Gasterosteus aculeatus. In accordance with the suggested high divergence rate of 13%/million years, the genetic variation of the parasite was high: haplotype diversity h = 0.985 and nucleotide diversity π = 0.0161. The differentiation among the parasite populations was substantial (Φst = 0.759), with two main allopatric clades (here termed Euro and North) accounting for 54% of the total genetic variation. The diversity center of the Euro clade was in the Baltic Sea, while the North clade was spread across the Barents and White Seas. A single haplotype within the North clade was found in the western and eastern Pacific Ocean. Divergence of main clades was estimated to be circa 200 thousand years ago. Each main clade was further divided into six distinct subclades, estimated to have diverged in isolation since 135 thousand years ago. This second division corresponds approximately to the Eemian interglacial predating the last glacial maximum. A demographic expansion of the subclades is associated with colonisation of northern Europe since the last glacial maximum, circa 15–40 thousand years ago. The parasite phylogeny is most likely explained by sequential isolated bottlenecks and expansions in numerous allopatric refugia. The postglacial intermingling and high variation in the marine parasite populations, separately in the Baltic and Barents Seas, suggest low competition of divergent parasite matrilines, coupled with a large population size and high rate of dispersal of hosts. The genetic contribution of the assumed refugial fish populations maintaining the parasite during the last glacial maximum was not detected among the marine sticklebacks, which perhaps were infected after range expansion.

  16. Displaced phylogeographic signals from Gyrodactylus arcuatus, a parasite of the three-spined stickleback Gasterosteus aculeatus, suggest freshwater glacial refugia in Europe.

    PubMed

    Lumme, Jaakko; Mäkinen, Hannu; Ermolenko, Alexey V; Gregg, Jacob L; Ziętara, Marek S

    2016-08-01

    We examined the global mitochondrial phylogeography of Gyrodactylus arcuatus, a flatworm ectoparasite of three-spined stickleback Gasterosteus aculeatus. In accordance with the suggested high divergence rate of 13%/million years, the genetic variation of the parasite was high: haplotype diversity h=0.985 and nucleotide diversity π=0.0161. The differentiation among the parasite populations was substantial (Φst=0.759), with two main allopatric clades (here termed Euro and North) accounting for 54% of the total genetic variation. The diversity center of the Euro clade was in the Baltic Sea, while the North clade was spread across the Barents and White Seas. A single haplotype within the North clade was found in the western and eastern Pacific Ocean. Divergence of main clades was estimated to be circa 200 thousand years ago. Each main clade was further divided into six distinct subclades, estimated to have diverged in isolation since 135 thousand years ago. This second division corresponds approximately to the Eemian interglacial predating the last glacial maximum. A demographic expansion of the subclades is associated with colonisation of northern Europe since the last glacial maximum, circa 15-40 thousand years ago. The parasite phylogeny is most likely explained by sequential isolated bottlenecks and expansions in numerous allopatric refugia. The postglacial intermingling and high variation in the marine parasite populations, separately in the Baltic and Barents Seas, suggest low competition of divergent parasite matrilines, coupled with a large population size and high rate of dispersal of hosts. The genetic contribution of the assumed refugial fish populations maintaining the parasite during the last glacial maximum was not detected among the marine sticklebacks, which perhaps were infected after range expansion. Copyright © 2016 Australian Society for Parasitology. All rights reserved.

  17. The Connemara Fan: a major glacial grounding line fan west of Ireland

    NASA Astrophysics Data System (ADS)

    McCarron, Stephen; Praeg, Daniel; Monteys, Xavier; Scott, Gill

    2014-05-01

    Glacigenic topography on the mid-shelf (~130-350 m water depth) west of Galway, Ireland appears to have the morphological form, internal architecture and sediments associated with a large glacial grounding-line fan. Seismic data collected in 2009 and 2012 (during the GLAMAR and GATEWAYS 1 campaigns) reveal that the broad, arcuate ridges of the 'Olex moraine' form the landward part of a fan system which prograded beyond the mid-shelf break (defining the outer margin of the 'Clare Platform') westwards into the Porcupine Seabight. The topography is comparable to larger shelf-edge trough-mouth fans found further north along the same margin, however no discernible 'trough' has been identified on the Clare Platform. The ridge and fan topographic assemblage is renamed the 'Connemara Fan' in its entirety, based on its genetic relations and geographic location due west of Connemara, western Ireland. A macrofossil recovered from within a debris flow on the outer fan slope comprised of remobilised plumites dates to ~ 20 ka Cal B.P., indicating sediment reworking downslope following deglacial sediment input to at least that time. The Connemara Fan is the most southerly glacigenic fan identified along the north-east Atlantic margin. Its identification also adds to our knowledge of possibly multiple generations of ice sheets feeding onto the Irish shelf from west-central Ireland and the occurrence of ice sheet geometries and dynamics that evacuated ice, melt-water and sediment (ice streams?) westwards across the Clare Platform during past glaciations.

  18. Clew Bay, Ireland

    NASA Image and Video Library

    2016-07-26

    Clew Bay is in County Mayo, Republic of Ireland. It contains Ireland's best example of sunken glacial drumlins. Clew Bay is associated with Grace O'Malley, the Pirate Queen during Elizabethan times; and Dorinish, a private island purchased by John Lennon. The drumlins are low hills formed from glacial sediment deposited at the end of the last Ice Age. The image was acquired May 31, 2016, covers an area of 22.5 by 26.2 km, and is located at 53.9 degrees north, 9.6 degrees west. http://photojournal.jpl.nasa.gov/catalog/PIA18670

  19. A Holarctic Biogeographical Analysis of the Collembola (Arthropoda, Hexapoda) Unravels Recent Post-Glacial Colonization Patterns

    PubMed Central

    Ávila-Jiménez, María Luisa; Coulson, Stephen James

    2011-01-01

    We aimed to describe the main Arctic biogeographical patterns of the Collembola, and analyze historical factors and current climatic regimes determining Arctic collembolan species distribution. Furthermore, we aimed to identify possible dispersal routes, colonization sources and glacial refugia for Arctic collembola. We implemented a Gaussian Mixture Clustering method on species distribution ranges and applied a distance- based parametric bootstrap test on presence-absence collembolan species distribution data. Additionally, multivariate analysis was performed considering species distributions, biodiversity, cluster distribution and environmental factors (temperature and precipitation). No clear relation was found between current climatic regimes and species distribution in the Arctic. Gaussian Mixture Clustering found common elements within Siberian areas, Atlantic areas, the Canadian Arctic, a mid-Siberian cluster and specific Beringian elements, following the same pattern previously described, using a variety of molecular methods, for Arctic plants. Species distribution hence indicate the influence of recent glacial history, as LGM glacial refugia (mid-Siberia, and Beringia) and major dispersal routes to high Arctic island groups can be identified. Endemic species are found in the high Arctic, but no specific biogeographical pattern can be clearly identified as a sign of high Arctic glacial refugia. Ocean currents patterns are suggested as being an important factor shaping the distribution of Arctic Collembola, which is consistent with Antarctic studies in collembolan biogeography. The clear relations between cluster distribution and geographical areas considering their recent glacial history, lack of relationship of species distribution with current climatic regimes, and consistency with previously described Arctic patterns in a series of organisms inferred using a variety of methods, suggest that historical phenomena shaping contemporary collembolan

  20. A coherent high-precision radiocarbon chronology for the Late-glacial sequence at Sluggan Bog, Co. Antrim, Northern Ireland

    NASA Astrophysics Data System (ADS)

    Lowe, J. J.; Walker, M. J. C.; Scott, E. M.; Harkness, D. D.; Bryant, C. L.; Davies, S. M.

    2004-02-01

    Seventy-five radiocarbon dates are presented from Sluggan Bog in Co. Antrim, Northern Ireland. The Holocene peats are underlain by Late-glacial sediments, which also appear to have accumulated largely in a mire environment. The radiocarbon dates, from the Late-glacial and early Holocene part of the profile, were obtained from the humic and humin fractions of the sedimentary matrix, and from plant macrofossils. The last-named were dated by AMS and the sediment samples by radiometric (beta counting) methods. Age-depth models for the three dating series show a very high level of agreement between the two fractions and the macrofossils. No statistically significant difference is found between the beta counting and AMS results. Three tephras were located in the profile, the uppermost of which is in a stratigraphical position suggestive of the Vedde Ash, but the geochemical and radiocarbon evidence do not support this interpretation. The lower ashes are in the correct stratigraphical position for the Laacher See and Borrobol tephras, attributions substantiated by the radiocarbon evidence, but not by the geochemical data. The Sluggan sequence has generated one of the most internally consistent radiocarbon chronologies for any Late-glacial site in the British Isles, and it is suggested that in future more effort should be devoted to the search for, and analysis of, Late-glacial mire sequences, rather than the limnic records that have formed the principal focus of Late-glacial investigations hitherto. Copyright

  1. Rethinking refugia: Tree topology, divergence dates, and demographic history trace the distribution of the endangered Plymouth gentian (Sabatia kennedyana) from the Pleistocene glaciation to present day.

    PubMed

    Suarez-Gonzalez, Adriana; Sutton, Jolene T; Trant, Andrew J; Zamlynny, Elena; Good, Sara V

    2015-04-01

    Molecular population genetics is a powerful tool to infer how species responded to past environmental change. In the northern hemisphere, interest is increasing in how species responded to changes in ice coverage and temperature during the last glaciation maximum (LGM, between 18000-21000 yr ago) with a common assumption that glacial refugia were located at the southern edge of a species range. We reconstructed the glacial and postglacial phylogeography of Sabatia kennedyana, a member of the Atlantic Coastal Plains Flora with a current distribution from Nova Scotia (NS) to South Carolina, using both cpDNA and nuclear markers. We also examined clinal variation in morphological traits, in particular relative investment in asexual vs sexual growth. We find strong evidence that the species did not reside in southern glacial refugia, but rather in primary glacial refugia off the exposed continental shelf extending from Cape Cod and that this area was responsible for the founding of modern populations across the range from Nova Scotia (NS) to the United States. Additionally, based on the finding of higher cpDNA diversity and older cpDNA lineages in NS, we propose that multiple founder events occurred in NS, while only a single lineage gave rise to current populations in the United States. By understanding how S. kennedyana responded to past shifts in climate and by identifying areas of high genetic diversity in the northern range edge, we discuss the potential response of the species to future climate change scenarios. © 2015 Botanical Society of America, Inc.

  2. Phylogeography of Quercus variabilis Based on Chloroplast DNA Sequence in East Asia: Multiple Glacial Refugia and Mainland-Migrated Island Populations

    PubMed Central

    Kang, Hongzhang; Sun, Xiao; Yin, Shan; Du, Hongmei; Yamanaka, Norikazu; Gapare, Washington; Wu, Harry X.; Liu, Chunjiang

    2012-01-01

    The biogeographical relationships between far-separated populations, in particular, those in the mainland and islands, remain unclear for widespread species in eastern Asia where the current distribution of plants was greatly influenced by the Quaternary climate. Deciduous Oriental oak (Quercus variabilis) is one of the most widely distributed species in eastern Asia. In this study, leaf material of 528 Q. variabilis trees from 50 populations across the whole distribution (Mainland China, Korea Peninsular as well as Japan, Zhoushan and Taiwan Islands) was collected, and three cpDNA intergenic spacer fragments were sequenced using universal primers. A total of 26 haplotypes were detected, and it showed a weak phylogeographical structure in eastern Asia populations at species level, however, in the central-eastern region of Mainland China, the populations had more haplotypes than those in other regions, with a significant phylogeographical structure (N ST = 0.751> G ST = 0.690, P<0.05). Q. variabilis displayed high interpopulation and low intrapopulation genetic diversity across the distribution range. Both unimodal mismatch distribution and significant negative Fu’s FS indicated a demographic expansion of Q. variabilis populations in East Asia. A fossil calibrated phylogenetic tree showed a rapid speciation during Pleistocene, with a population augment occurred in Middle Pleistocene. Both diversity patterns and ecological niche modelling indicated there could be multiple glacial refugia and possible bottleneck or founder effects occurred in the southern Japan. We dated major spatial expansion of Q. variabilis population in eastern Asia to the last glacial cycle(s), a period with sea-level fluctuations and land bridges in East China Sea as possible dispersal corridors. This study showed that geographical heterogeneity combined with climate and sea-level changes have shaped the genetic structure of this wide-ranging tree species in East Asia. PMID:23115642

  3. Postglacial recolonization at a snail's pace (Trochulus villosus): confronting competing refugia hypotheses using model selection.

    PubMed

    Dépraz, A; Cordellier, M; Hausser, J; Pfenninger, M

    2008-05-01

    The localization of Last Glacial Maximum (LGM) refugia is crucial information to understand a species' history and predict its reaction to future climate changes. However, many phylogeographical studies often lack sampling designs intensive enough to precisely localize these refugia. The hairy land snail Trochulus villosus has a small range centred on Switzerland, which could be intensively covered by sampling 455 individuals from 52 populations. Based on mitochondrial DNA sequences (COI and 16S), we identified two divergent lineages with distinct geographical distributions. Bayesian skyline plots suggested that both lineages expanded at the end of the LGM. To find where the origin populations were located, we applied the principles of ancestral character reconstruction and identified a candidate refugium for each mtDNA lineage: the French Jura and Central Switzerland, both ice-free during the LGM. Additional refugia, however, could not be excluded, as suggested by the microsatellite analysis of a population subset. Modelling the LGM niche of T. villosus, we showed that suitable climatic conditions were expected in the inferred refugia, but potentially also in the nunataks of the alpine ice shield. In a model selection approach, we compared several alternative recolonization scenarios by estimating the Akaike information criterion for their respective maximum-likelihood migration rates. The 'two refugia' scenario received by far the best support given the distribution of genetic diversity in T. villosus populations. Provided that fine-scale sampling designs and various analytical approaches are combined, it is possible to refine our necessary understanding of species responses to environmental changes.

  4. Geothermal activity helps life survive glacial cycles

    PubMed Central

    Fraser, Ceridwen I.; Terauds, Aleks; Smellie, John; Convey, Peter; Chown, Steven L.

    2014-01-01

    Climate change has played a critical role in the evolution and structure of Earth’s biodiversity. Geothermal activity, which can maintain ice-free terrain in glaciated regions, provides a tantalizing solution to the question of how diverse life can survive glaciations. No comprehensive assessment of this “geothermal glacial refugia” hypothesis has yet been undertaken, but Antarctica provides a unique setting for doing so. The continent has experienced repeated glaciations that most models indicate blanketed the continent in ice, yet many Antarctic species appear to have evolved in almost total isolation for millions of years, and hence must have persisted in situ throughout. How could terrestrial species have survived extreme glaciation events on the continent? Under a hypothesis of geothermal glacial refugia and subsequent recolonization of nongeothermal regions, we would expect to find greater contemporary diversity close to geothermal sites than in nongeothermal regions, and significant nestedness by distance of this diversity. We used spatial modeling approaches and the most comprehensive, validated terrestrial biodiversity dataset yet created for Antarctica to assess spatial patterns of diversity on the continent. Models clearly support our hypothesis, indicating that geothermally active regions have played a key role in structuring biodiversity patterns in Antarctica. These results provide critical insights into the evolutionary importance of geothermal refugia and the history of Antarctic species. PMID:24616489

  5. Relative sea-level changes and crustal movements in Britain and Ireland since the Last Glacial Maximum

    NASA Astrophysics Data System (ADS)

    Shennan, Ian; Bradley, Sarah L.; Edwards, Robin

    2018-05-01

    The new sea-level database for Britain and Ireland contains >2100 data points from 86 regions and records relative sea-level (RSL) changes over the last 20 ka and across elevations ranging from ∼+40 to -55 m. It reveals radically different patterns of RSL as we move from regions near the centre of the Celtic ice sheet at the last glacial maximum to regions near and beyond the ice limits. Validated sea-level index points and limiting data show good agreement with the broad patterns of RSL change predicted by current glacial isostatic adjustment (GIA) models. The index points show no consistent pattern of synchronous coastal advance and retreat across different regions, ∼100-500 km scale, indicating that within-estuary processes, rather than decimetre- and centennial-scale oscillations in sea level, produce major controls on the temporal pattern of horizontal shifts in coastal sedimentary environments. Comparisons between the database and GIA model predictions for multiple regions provide potentially powerful constraints on various characteristics of global GIA models, including the magnitude of MWP1A, the final deglaciation of the Laurentide ice sheet and the continued melting of Antarctica after 7 ka BP.

  6. A long pollen record from lowland Amazonia: Forest and cooling in glacial times

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Colinvaux, P.A.; Moreno, J.E.; Bush, M.B.

    A continuous pollen history of more than 40,000 years was obtained from a lake in the lowland Amazon rain forest. Pollen spectra demonstrate that tropical rain forest occupied the region continuously and that savannas or grasslands were not present during the last glacial maximum. The data suggest that the western Amazon forest was not fragmented into refugia in glacial times and that the lowlands were not a source of dust. Glacial age forests were comparable to modern forests but also included species now restricted to higher evaluations by temperature, suggesting a cooling of the order of 5{degrees} to 6{degrees}C. 23more » refs., 22 tabs.« less

  7. Managing climate change refugia for climate adaptation

    USGS Publications Warehouse

    Morelli, Toni L.; Jackson, Stephen T.

    2016-01-01

    Refugia have long been studied from paleontological and biogeographical perspectives to understand how populations persisted during past periods of unfavorable climate. Recently, researchers have applied the idea to contemporary landscapes to identify climate change refugia, here defined as areas relatively buffered from contemporary climate change over time that enable persistence of valued physical, ecological, and socio-cultural resources. We differentiate historical and contemporary views, and characterize physical and ecological processes that create and maintain climate change refugia. We then delineate how refugia can fit into existing decision support frameworks for climate adaptation and describe seven steps for managing them. Finally, we identify challenges and opportunities for operationalizing the concept of climate change refugia. Managing climate change refugia can be an important option for conservation in the face of ongoing climate change.

  8. Managing Climate Change Refugia for Climate Adaptation.

    PubMed

    Morelli, Toni Lyn; Daly, Christopher; Dobrowski, Solomon Z; Dulen, Deanna M; Ebersole, Joseph L; Jackson, Stephen T; Lundquist, Jessica D; Millar, Constance I; Maher, Sean P; Monahan, William B; Nydick, Koren R; Redmond, Kelly T; Sawyer, Sarah C; Stock, Sarah; Beissinger, Steven R

    2016-01-01

    Refugia have long been studied from paleontological and biogeographical perspectives to understand how populations persisted during past periods of unfavorable climate. Recently, researchers have applied the idea to contemporary landscapes to identify climate change refugia, here defined as areas relatively buffered from contemporary climate change over time that enable persistence of valued physical, ecological, and socio-cultural resources. We differentiate historical and contemporary views, and characterize physical and ecological processes that create and maintain climate change refugia. We then delineate how refugia can fit into existing decision support frameworks for climate adaptation and describe seven steps for managing them. Finally, we identify challenges and opportunities for operationalizing the concept of climate change refugia. Managing climate change refugia can be an important option for conservation in the face of ongoing climate change.

  9. Managing Climate Change Refugia for Climate Adaptation

    PubMed Central

    Daly, Christopher; Dobrowski, Solomon Z.; Dulen, Deanna M.; Ebersole, Joseph L.; Jackson, Stephen T.; Lundquist, Jessica D.; Millar, Constance I.; Maher, Sean P.; Monahan, William B.; Nydick, Koren R.; Redmond, Kelly T.; Sawyer, Sarah C.; Stock, Sarah; Beissinger, Steven R.

    2016-01-01

    Refugia have long been studied from paleontological and biogeographical perspectives to understand how populations persisted during past periods of unfavorable climate. Recently, researchers have applied the idea to contemporary landscapes to identify climate change refugia, here defined as areas relatively buffered from contemporary climate change over time that enable persistence of valued physical, ecological, and socio-cultural resources. We differentiate historical and contemporary views, and characterize physical and ecological processes that create and maintain climate change refugia. We then delineate how refugia can fit into existing decision support frameworks for climate adaptation and describe seven steps for managing them. Finally, we identify challenges and opportunities for operationalizing the concept of climate change refugia. Managing climate change refugia can be an important option for conservation in the face of ongoing climate change. PMID:27509088

  10. Phylogeography of the South China Field Mouse (Apodemus draco) on the Southeastern Tibetan Plateau Reveals High Genetic Diversity and Glacial Refugia

    PubMed Central

    Liu, Yang; Liao, Lihuan; Zhang, Xiuyue; Yue, Bisong

    2012-01-01

    The southeastern margin of the Tibetan Plateau (SEMTP) is a particularly interesting region due to its topographic complexity and unique geologic history, but phylogeographic studies that focus on this region are rare. In this study, we investigated the phylogeography of the South China field mouse, Apodemus draco, in order to assess the role of geologic and climatic events on the Tibetan Plateau in shaping its genetic structure. We sequenced mitochondrial cytochrome b (cyt b) sequences in 103 individuals from 47 sampling sites. In addition, 23 cyt b sequences were collected from GenBank for analyses. Phylogenetic, demographic and landscape genetic methods were conducted. Seventy-six cyt b haplotypes were found and the genetic diversity was extremely high (π = 0.0368; h = 0.989). Five major evolutionary clades, based on geographic locations, were identified. Demographic analyses implied subclade 1A and subclade 1B experienced population expansions at about 0.052-0.013 Mya and 0.014-0.004 Mya, respectively. The divergence time analysis showed that the split between clade 1 and clade 2 occurred 0.26 Mya, which fell into the extensive glacial period (EGP, 0.5-0.17 Mya). The divergence times of other main clades (2.20-0.55 Mya) were congruent with the periods of the Qingzang Movement (3.6-1.7 Mya) and the Kun-Huang Movement (1.2-0.6 Mya), which were known as the most intense uplift events in the Tibetan Plateau. Our study supported the hypothesis that the SEMTP was a large late Pleistocene refugium, and further inferred that the Gongga Mountain Region and Hongya County were glacial refugia for A. draco in clade 1. We hypothesize that the evolutionary history of A. draco in the SEMTP primarily occurred in two stages. First, an initial divergence would have been shaped by uplift events of the Tibetan Plateau. Then, major glaciations in the Pleistocene added complexity to its demographic history and genetic structure. PMID:22666478

  11. Glacial history affected phenotypic differentiation in the alpine plant, Campanula thyrsoides.

    PubMed

    Scheepens, J F; Frei, Eva S; Stöcklin, Jürg

    2013-01-01

    Numerous widespread Alpine plant species show molecular differentiation among populations from distinct regions. This has been explained as the result of genetic drift during glacial survival in isolated refugia along the border of the European Alps. Since genetic drift may affect molecular markers and phenotypic traits alike, we asked whether phenotypic differentiation mirrors molecular patterns among Alpine plant populations from different regions. Phenotypic traits can be under selection, so we additionally investigated whether part of the phenotypic differentiation can be explained by past selection and/or current adaptation. Using the monocarpic Campanula thyrsoides as our study species, a common garden experiment with plants from 21 populations from four phylogeographic groups located in regions across the Alps and the Jura Mountains was performed to test for differentiation in morphological and phenological traits. Past selection was investigated by comparing phenotypic differentiation among and within regions with molecular differentiation among and within regions. The common garden results indicated regional differentiation among populations for all investigated phenotypic traits, particularly in phenology. Delayed flowering in plants from the South-eastern Alps suggested adaptation to long sub-mediterranean summers and contrasted with earlier flowering of plants experiencing shorter growing seasons in regions with higher elevation to the West. Comparisons between molecular and phenotypic differentiation revealed diversifying selection among regions in height and biomass, which is consistent with adaptation to environmental conditions in glacial refugia. Within regions, past selection acted against strong diversification for most phenotypic traits, causing restricted postglacial adaptation. Evidence consistent with post-glacial adaptation was also given by negative correlation coefficients between several phenotypic traits and elevation of the population

  12. Less Pollen-Mediated Gene Flow for More Signatures of Glacial Lineages: Congruent Evidence from Balsam Fir cpDNA and mtDNA for Multiple Refugia in Eastern and Central North America

    PubMed Central

    Cinget, Benjamin; Gérardi, Sébastien; Beaulieu, Jean; Bousquet, Jean

    2015-01-01

    The phylogeographic structure and postglacial history of balsam fir (Abies balsamea), a transcontinental North American boreal conifer, was inferred using mitochondrial DNA (mtDNA) and chloroplast DNA (cpDNA) markers. Genetic structure among 107 populations (mtDNA data) and 75 populations (cpDNA data) was analyzed using Bayesian and genetic distance approaches. Population differentiation was high for mtDNA (dispersed by seeds only), but also for cpDNA (dispersed by seeds and pollen), indicating that pollen gene flow is more restricted in balsam fir than in other boreal conifers. Low cpDNA gene flow in balsam fir may relate to low pollen production due to the inherent biology of the species and populations being decimated by recurrent spruce budworm epidemics, and/or to low dispersal of pollen grains due to their peculiar structural properties. Accordingly, a phylogeographic structure was detected using both mtDNA and cpDNA markers and population structure analyses supported the existence of at least five genetically distinct glacial lineages in central and eastern North America. Four of these would originate from glacial refugia located south of the Laurentide ice sheet, while the last one would have persisted in the northern Labrador region. As expected due to reduced pollen-mediated gene flow, congruence between the geographic distribution of mtDNA and cpDNA lineages was higher than in other North American conifers. However, concordance was not complete, reflecting that restricted but nonetheless detectable cpDNA gene flow among glacial lineages occurred during the Holocene. As a result, new cpDNA and mtDNA genome combinations indicative of cytoplasmic genome capture were observed. PMID:25849816

  13. Distribution and protection of climatic refugia in North America.

    PubMed

    Michalak, Julia L; Lawler, Joshua J; Roberts, David R; Carroll, Carlos

    2018-05-10

    As evidenced by past climatic refugia, locations projected to harbor remnants of present day climates may serve as critical refugia for current biodiversity in the face of modern climate change. Here, we map potential future climatic refugia across North America, defined as locations with increasingly rare climatic conditions. We identified these locations by tracking projected changes in the size and distribution of climate analogs over time. We used biologically-derived thresholds to define analogs and tested the impacts of dispersal limitation using four distances to limit analog searches. We identified at most 12% of North America as potential climatic refugia. Refugia extent varied depending on the analog threshold, dispersal distance, and climate projection. However, in all cases refugia were concentrated at high elevations and in topographically complex regions. Refugia identified using different climate projections were largely nested, suggesting that identified refugia were relatively robust to climate projection selection. Existing conservation areas cover approximately 10% of North America and yet protected up to 25% of identified refugia, indicating that protected areas disproportionately include refugia. Refugia located at lower latitudes (≤ 40°N) and slightly lower elevations (∼2500 m) were more likely to be unprotected. Based on our results, a 23% expansion of the protected areas network would be sufficient to protect the refugia that were present under all three of the climate projections that we explored. We propose that these refugia are high conservation priorities, due to their potential to harbor rare species in the future. However, these locations are simultaneously highly vulnerable to climate change over the long-term. We found that these refugia contracted substantially between the 2050s and the 2080s, emphasizing that the pace of climate change will strongly determine the availability and effectiveness of refugia for protecting today

  14. Managing Climate Change Refugia for Climate Adaptation ...

    EPA Pesticide Factsheets

    The concept of refugia has long been studied from theoretical and paleontological perspectives to understand how populations persisted during past periods of unfavorable climate. Recently, researchers have applied the idea to contemporary landscapes to identify climate change refugia, locations that may be unusually buffered from climate change effects so as to increase persistence of valued resources. Here we distinguish between paleoecological and contemporary viewpoints, characterize physical and ecological processes that create and maintain climate change refugia, summarize the process of identifying and mapping them, and delineate how refugia can fit into the existing framework of natural resource management. We also suggest three primary courses of action at these sites: prioritization, protection, and propagation. Although not a panacea, managing climate change refugia can be an important adaptation option for conserving valuable resources in the face of ongoing and future climate change. “In a nutshell” (100 words) • Climate change refugia are defined as areas relatively buffered from contemporary climate change, enabling persistence of valued physical, ecological, and cultural resources. • Refugia can be incorporated as key components of a climate adaptation strategy because their prioritization by management may enable their associated resources to persist locally and eventually spread to future suitable habitat. • Steps for

  15. Several Pleistocene refugia detected in the high alpine plant Phyteuma globulariifolium sternb & hoppe (Campanulaceae) in the European Alps.

    PubMed

    Schönswetter, P; Tribsch, A; Barfuss, M; Niklfeld, H

    2002-12-01

    Phyteuma globulariifolium is a high alpine plant species growing in the European Alps and the Pyrenees. In order to elucidate its glacial history, 325 individuals from 69 populations were analysed using the amplified fragment length polymorphism (AFLP) technique. A strongly hierarchical phylogeographical pattern was detected: Two major east-west vicariant groups can be separated along a gap in the distributional area. A further subdivision into at least four populational groups is in congruence with presumed peripheral glacial refugia. There is no indication for survival on unglaciated mountain tops (nunataks) in the interior of the Pleistocene ice shield covering the Alps. Our results favour glacial survival in peripheral, unglaciated or not fully glaciated areas. Populations of P. globulariifolium in the Pyrenees are the result of relatively recent long-distance dispersal. Within the Alps, there is strong differentiation among groups of populations, whereas within them the differentiation is weak. This suggests high levels of gene-flow over short to middle distances.

  16. Late-glacial recolonization and phylogeography of European red deer (Cervus elaphus L.).

    PubMed

    Meiri, Meirav; Lister, Adrian M; Higham, Thomas F G; Stewart, John R; Straus, Lawrence G; Obermaier, Henriette; González Morales, Manuel R; Marín-Arroyo, Ana B; Barnes, Ian

    2013-09-01

    The Pleistocene was an epoch of extreme climatic and environmental changes. How individual species responded to the repeated cycles of warm and cold stages is a major topic of debate. For the European fauna and flora, an expansion-contraction model has been suggested, whereby temperate species were restricted to southern refugia during glacial times and expanded northwards during interglacials, including the present interglacial (Holocene). Here, we test this model on the red deer (Cervus elaphus) a large and highly mobile herbivore, using both modern and ancient mitochondrial DNA from the entire European range of the species over the last c. 40,000 years. Our results indicate that this species was sensitive to the effects of climate change. Prior to the Last Glacial Maximum (LGM) haplogroups restricted today to South-East Europe and Western Asia reached as far west as the UK. During the LGM, red deer was mainly restricted to southern refugia, in Iberia, the Balkans and possibly in Italy and South-Western Asia. At the end of the LGM, red deer expanded from the Iberian refugium, to Central and Northern Europe, including the UK, Belgium, Scandinavia, Germany, Poland and Belarus. Ancient DNA data cannot rule out refugial survival of red deer in North-West Europe through the LGM. Had such deer survived, though, they were replaced by deer migrating from Iberia at the end of the glacial. The Balkans served as a separate LGM refugium and were probably connected to Western Asia with genetic exchange between the two areas. © 2013 John Wiley & Sons Ltd.

  17. Projected climate changes threaten ancient refugia of kelp forests in the North Atlantic.

    PubMed

    Assis, Jorge; Araújo, Miguel B; Serrão, Ester A

    2018-01-01

    Intraspecific genetic variability is critical for species adaptation and evolution and yet it is generally overlooked in projections of the biological consequences of climate change. We ask whether ongoing climate changes can cause the loss of important gene pools from North Atlantic relict kelp forests that persisted over glacial-interglacial cycles. We use ecological niche modelling to predict genetic diversity hotspots for eight species of large brown algae with different thermal tolerances (Arctic to warm temperate), estimated as regions of persistence throughout the Last Glacial Maximum (20,000 YBP), the warmer Mid-Holocene (6,000 YBP), and the present. Changes in the genetic diversity within ancient refugia were projected for the future (year 2100) under two contrasting climate change scenarios (RCP2.6 and RCP8.5). Models predicted distributions that matched empirical distributions in cross-validation, and identified distinct refugia at the low latitude ranges, which largely coincide among species with similar ecological niches. Transferred models into the future projected polewards expansions and substantial range losses in lower latitudes, where richer gene pools are expected (in Nova Scotia and Iberia for cold affinity species and Gibraltar, Alboran, and Morocco for warm-temperate species). These effects were projected for both scenarios but were intensified under the extreme RCP8.5 scenario, with the complete borealization (circum-Arctic colonization) of kelp forests, the redistribution of the biogeographical transitional zones of the North Atlantic, and the erosion of global gene pools across all species. As the geographic distribution of genetic variability is unknown for most marine species, our results represent a baseline for identification of locations potentially rich in unique phylogeographic lineages that are also climatic relics in threat of disappearing. © 2017 John Wiley & Sons Ltd.

  18. Managing climate change refugia for climate adaptation

    Treesearch

    Toni Lyn Morelli; Christopher Daly; Solomon Z. Dobrowski; Deanna M. Dulen; Joseph L. Ebersole; Stephen T. Jackson; Jessica D. Lundquist; Connie Millar; Sean P. Maher; William B. Monahan; Koren R. Nydick; Kelly T. Redmond; Sarah C. Sawyer; Sarah Stock; Steven R. Beissinger

    2016-01-01

    Refugia have long been studied from paleontological and biogeographical perspectives to understand how populations persisted during past periods of unfavorable climate. Recently, researchers have applied the idea to contemporary landscapes to identify climate change refugia, here defined as areas relatively buffered from contemporary climate change over time that...

  19. Linking landscape variables to cold water refugia in rivers.

    PubMed

    Monk, Wendy A; Wilbur, Nathan M; Curry, R Allen; Gagnon, Rolland; Faux, Russell N

    2013-03-30

    The protection of coldwater refugia within aquatic systems requires the identification of thermal habitats in rivers. These refugia provide critical thermal habitats for brook trout (Salvelinus fontinalis) and Atlantic salmon (Salmo salar) during periods of thermal stress, for example during summer high temperature events. This study aims to model these refugia using georeferenced thermal infrared images collected during late July 2008 and 2009 for a reach of the Cains River, New Brunswick, Canada. These images were paired with geospatial catchment variables to identify the driving factors for coldwater refugia located within tributaries to the main channel. Using Partial Least Square (PLS) Regression, results suggest that median temperatures of tributary catchments are driven by their position within the landscape including slope in addition to the density of wetlands and mixed forest within the upstream catchment. Similar results are presented when PLS models were developed to predict the magnitude of the cold water refugia (i.e. the difference between the mainstem water temperature and the thermal refugia). These results suggest that thermal infrared images can be used to predict critical summer habitats for coldwater fishes. Copyright © 2013 Elsevier Ltd. All rights reserved.

  20. Post-Glacial and Paleo-Environmental History of the West Coast of Vancouver Island

    NASA Astrophysics Data System (ADS)

    Dallimore, A.; Enkin, R. J.

    2005-12-01

    Annually laminated sediments in anoxic fjords are potentially ideal paleoclimate recorders, particularly once proxy measurements for atmospheric, oceanographic and sedimentological conditions have been calibrated. On the west coast of Canada, these sediments also record the changing environment as glaciers retreated from this area about 12 ka y BP. In Effingham Inlet, a 40 m core taken from the French ship the Marion Dufresne as part of the international IMAGES/PAGES program, gives evidence of an isolation basin at maximum glacial isostatic rebound and lowest paleo-sea level followed by eustatic sea level rise about 10 ka y BP. The Late Pleistocene record also marks dramatic changes in glacial sedimentary source and transport. Excellent chronological control is provided by complementary yet independent dating methods including radiocarbon dates on both plants and shells, identification of the Mazama Ash, varve counting and paleomagnetic, paleosecular variation correlations in the lower, pro-glacial section of the core which does not contain organic material. Paleoenvironmental evidence from this core provides information on immediate post-glacial conditions along the coast and rapid climatic changes throughout the Holocene, with implications for the possibility of early human migration routes and refugia.

  1. The ancient tropical rainforest tree Symphonia globulifera L. f. (Clusiaceae) was not restricted to postulated Pleistocene refugia in Atlantic Equatorial Africa.

    PubMed

    Budde, K B; González-Martínez, S C; Hardy, O J; Heuertz, M

    2013-07-01

    Understanding the history of forests and their species' demographic responses to past disturbances is important for predicting impacts of future environmental changes. Tropical rainforests of the Guineo-Congolian region in Central Africa are believed to have survived the Pleistocene glacial periods in a few major refugia, essentially centred on mountainous regions close to the Atlantic Ocean. We tested this hypothesis by investigating the phylogeographic structure of a widespread, ancient rainforest tree species, Symphonia globulifera L. f. (Clusiaceae), using plastid DNA sequences (chloroplast DNA [cpDNA], psbA-trnH intergenic spacer) and nuclear microsatellites (simple sequence repeats, SSRs). SSRs identified four gene pools located in Benin, West Cameroon, South Cameroon and Gabon, and São Tomé. This structure was also apparent at cpDNA. Approximate Bayesian Computation detected recent bottlenecks approximately dated to the last glacial maximum in Benin, West Cameroon and São Tomé, and an older bottleneck in South Cameroon and Gabon, suggesting a genetic effect of Pleistocene cycles of forest contraction. CpDNA haplotype distribution indicated wide-ranging long-term persistence of S. globulifera both inside and outside of postulated forest refugia. Pollen flow was four times greater than that of seed in South Cameroon and Gabon, which probably enabled rapid population recovery after bottlenecks. Furthermore, our study suggested ecotypic differentiation-coastal or swamp vs terra firme-in S. globulifera. Comparison with other tree phylogeographic studies in Central Africa highlighted the relevance of species-specific responses to environmental change in forest trees.

  2. Pennsylvanian tropical rain forests responded to glacial-interglacial rhythms

    NASA Astrophysics Data System (ADS)

    Falcon-Lang, Howard J.

    2004-08-01

    Pennsylvanian tropical rain forests flourished during an icehouse climate mode. Although it is well established that Milankovitch-band glacial-interglacial rhythms caused marked synchronous changes in Pennsylvanian tropical climate and sea level, little is known of vegetation response to orbital forcing. This knowledge gap has now been addressed through sequence- stratigraphic analysis of megafloral and palynofloral assemblages within the Westphalian D Cantabrian Sydney Mines Formation of eastern Canada. This succession was deposited in a low- accommodation setting where sequences can be attributed confidently to glacio-eustasy. Results show that long-lived, low-diversity peat mires dominated by lycopsids were initiated during deglaciation events, but were mostly drowned by rising sea level at maximum interglacial conditions. Only upland coniferopsid forests survived flooding without significant disturbance. Mid- to late interglacial phases witnessed delta-plain progradation and establishment of high-diversity, mineral-substrate rain forests containing lycopsids, sphenopsids, pteridosperms, cordaites, and tree ferns. Renewed glaciation resulted in sea-level fall, paleovalley incision, and the onset of climatic aridity. Glacial vegetation was dominated by cordaites, pteridosperms, and tree ferns; hydrophilic lycopsids and sphenopsids survived in paleovalley refugia. Findings clearly demonstrate the dynamic nature of Pennsylvanian tropical ecosystems and are timely given current debates about the impact of Quaternary glacial-interglacial rhythms on the biogeography of tropical rain forest.

  3. Glacial modification of granite tors in the Cairngorms, Scotland

    USGS Publications Warehouse

    Hall, A.M.; Phillips, W.M.

    2006-01-01

    A range of evidence indicates that many granite tors in the Cairngorms have been modified by the flow of glacier ice during the Pleistocene. Comparisons with SW England and the use of a space-time transformation across 38 tor groups in the Cairngorms allow a model to be developed for progressive glacial modification. Tors with deeply etched surfaces and no, or limited, block removal imply an absence of significant glacial modification. The removal of superstructure and blocks, locally forming boulder trains, and the progressive reduction of tors to stumps and basal slabs represent the more advanced stages of modification. Recognition of some slabs as tor stumps from which glacial erosion has removed all superstructure allows the original distribution of tors to be reconstructed for large areas of the Cairngorms. Unmodified tors require covers of non-erosive, cold-based ice during all of the cold stages of the Middle and Late Pleistocene. Deformation beneath cold-based glacier ice is capable of the removal of blocks but advanced glacial modification requires former wet-based glacier ice. The depth of glacial erosion at former tor sites remains limited largely to the partial or total elimination of the upstanding tor form. Cosmogenic nuclide exposure ages (Phillips et al., 2006) together with data from weathering pit depths (Hall and Phillips, 2006), from the surfaces of tors and large erratic blocks require that the glacial entrainment of blocks from tors occurred in Marine Isotope Stages (MIS) 4-2, 6 and, probably, at least one earlier phase. The occurrence of glacially modified tors on or close to, the main summits of the Cairngorms requires full ice cover over the mountains during these Stages. Evidence from the Cairngorms indicates that tor morphology can be regarded as an important indicator of former ice cover in many formerly glaciated areas, particularly where other evidence of ice cover is sparse. Recognition of the glacial modification of tors is important

  4. Glacial legacies on interglacial vegetation at the Pliocene-Pleistocene transition in NE Asia

    PubMed Central

    Herzschuh, Ulrike; Birks, H. John B.; Laepple, Thomas; Andreev, Andrei; Melles, Martin; Brigham-Grette, Julie

    2016-01-01

    Broad-scale climate control of vegetation is widely assumed. Vegetation-climate lags are generally thought to have lasted no more than a few centuries. Here our palaeoecological study challenges this concept over glacial–interglacial timescales. Through multivariate analyses of pollen assemblages from Lake El'gygytgyn, Russian Far East and other data we show that interglacial vegetation during the Plio-Pleistocene transition mainly reflects conditions of the preceding glacial instead of contemporary interglacial climate. Vegetation–climate disequilibrium may persist for several millennia, related to the combined effects of permafrost persistence, distant glacial refugia and fire. In contrast, no effects from the preceding interglacial on glacial vegetation are detected. We propose that disequilibrium was stronger during the Plio-Pleistocene transition than during the Mid-Pliocene Warm Period when, in addition to climate, herbivory was important. By analogy to the past, we suggest today's widespread larch ecosystem on permafrost is not in climate equilibrium. Vegetation-based reconstructions of interglacial climates used to assess atmospheric CO2–temperature relationships may thus yield misleading simulations of past global climate sensitivity. PMID:27338025

  5. Alternative glacial-interglacial refugia demographic hypotheses tested on Cephalocereus columna-trajani (Cactaceae) in the intertropical Mexican drylands

    PubMed Central

    Cornejo-Romero, Amelia; Aguilar-Martínez, Gustavo F.; Medina-Sánchez, Javier; Rendón-Aguilar, Beatriz; Valverde, Pedro Luis; Zavala-Hurtado, Jose Alejandro; Serrato, Alejandra; Rivas-Arancibia, Sombra; Pérez-Hernández, Marco Aurelio; López-Ortega, Gerardo; Jiménez-Sierra, Cecilia

    2017-01-01

    Historic demography changes of plant species adapted to New World arid environments could be consistent with either the Glacial Refugium Hypothesis (GRH), which posits that populations contracted to refuges during the cold-dry glacial and expanded in warm-humid interglacial periods, or with the Interglacial Refugium Hypothesis (IRH), which suggests that populations contracted during interglacials and expanded in glacial times. These contrasting hypotheses are developed in the present study for the giant columnar cactus Cephalocereus columna-trajani in the intertropical Mexican drylands where the effects of Late Quaternary climatic changes on phylogeography of cacti remain largely unknown. In order to determine if the historic demography and phylogeographic structure of the species are consistent with either hypothesis, sequences of the chloroplast regions psbA-trnH and trnT-trnL from 110 individuals from 10 populations comprising the full distribution range of this species were analysed. Standard estimators of genetic diversity and structure were calculated. The historic demography was analysed using a Bayesian approach and the palaeodistribution was derived from ecological niche modelling to determine if, in the arid environments of south-central Mexico, glacial-interglacial cycles drove the genetic divergence and diversification of this species. Results reveal low but statistically significant population differentiation (FST = 0.124, P < 0.001), although very clear geographic clusters are not formed. Genetic diversity, haplotype network and Approximate Bayesian Computation (ABC) demographic analyses suggest a population expansion estimated to have taken place in the Last Interglacial (123.04 kya, 95% CI 115.3–130.03). The species palaeodistribution is consistent with the ABC analyses and indicates that the potential area of palaedistribution and climatic suitability were larger during the Last Interglacial and Holocene than in the Last Glacial Maximum. Overall

  6. Alternative glacial-interglacial refugia demographic hypotheses tested on Cephalocereus columna-trajani (Cactaceae) in the intertropical Mexican drylands.

    PubMed

    Cornejo-Romero, Amelia; Vargas-Mendoza, Carlos Fabián; Aguilar-Martínez, Gustavo F; Medina-Sánchez, Javier; Rendón-Aguilar, Beatriz; Valverde, Pedro Luis; Zavala-Hurtado, Jose Alejandro; Serrato, Alejandra; Rivas-Arancibia, Sombra; Pérez-Hernández, Marco Aurelio; López-Ortega, Gerardo; Jiménez-Sierra, Cecilia

    2017-01-01

    Historic demography changes of plant species adapted to New World arid environments could be consistent with either the Glacial Refugium Hypothesis (GRH), which posits that populations contracted to refuges during the cold-dry glacial and expanded in warm-humid interglacial periods, or with the Interglacial Refugium Hypothesis (IRH), which suggests that populations contracted during interglacials and expanded in glacial times. These contrasting hypotheses are developed in the present study for the giant columnar cactus Cephalocereus columna-trajani in the intertropical Mexican drylands where the effects of Late Quaternary climatic changes on phylogeography of cacti remain largely unknown. In order to determine if the historic demography and phylogeographic structure of the species are consistent with either hypothesis, sequences of the chloroplast regions psbA-trnH and trnT-trnL from 110 individuals from 10 populations comprising the full distribution range of this species were analysed. Standard estimators of genetic diversity and structure were calculated. The historic demography was analysed using a Bayesian approach and the palaeodistribution was derived from ecological niche modelling to determine if, in the arid environments of south-central Mexico, glacial-interglacial cycles drove the genetic divergence and diversification of this species. Results reveal low but statistically significant population differentiation (FST = 0.124, P < 0.001), although very clear geographic clusters are not formed. Genetic diversity, haplotype network and Approximate Bayesian Computation (ABC) demographic analyses suggest a population expansion estimated to have taken place in the Last Interglacial (123.04 kya, 95% CI 115.3-130.03). The species palaeodistribution is consistent with the ABC analyses and indicates that the potential area of palaedistribution and climatic suitability were larger during the Last Interglacial and Holocene than in the Last Glacial Maximum. Overall

  7. Post-glacial sea-level history for NE Ireland (Belfast Lough) based on offshore evidence

    NASA Astrophysics Data System (ADS)

    Quinn, R.; Plets, R. M.; Callard, L.; Cooper, A.; Long, A. J.; Belknap, D. F.; Edwards, R.; Jackson, D.; Kelley, J. T.; Long, D.; Milne, G. A.; Monteys, X.

    2013-12-01

    Glacio-isostatic adjustment (GIA) models suggest a complex relative sea-level (RSL) pattern around the Irish Sea Basin after the Last Glacial Maximum (LGM), with modelled sea-level lowstands ranging from -12 m in the north to greater than -60 m in the south of the Basin. However, these GIA models are poorly constrained by observational data offshore. Belfast Lough, on the NE coast of Ireland, is one of seven sites chosen to investigate this complex RSL history as part of the project ';Late Glacial Sea level minima in the Western British Isles' (NERC NE/H024301/1). Belfast Lough was chosen as one of the candidate sites on the basis of location (at the northern end of the Irish Sea Basin), sedimentary environment (grossly depositional) and the fact that the lowstand predicted for the Belfast Lough area by a recent version of the GIA model (-16.5 m) differs significantly from the (limited) extant observational data, which interprets the lowstand at -30 m. In 2011 and 2012 we gathered new multi-beam echo-sounder data, >200 km trackline pinger- and boomer- seismic reflection data and 46 vibrocores in Belfast Lough. Radiocarbon dating and palaeoenvironmental analysis are used to constrain the interpretation of the seismic and sediment data. Five seismo-stratigraphic units are interpreted, with a distinct erosional surface between U3 and U4 interpreted as a transgressive surface associated with sea level rise post-dating a RSL lowstand. Foraminiferal evidence indicates an increase in marine species (from lagoonal/estuarine to fully marine) from U4 to U5. Integration of the seismic and core data indicate an erosional event prior to 12.7 cal yr BP resulting in a planated surface in the inner Lough and wave-eroded drumlins at the mouth of the Lough between -15 and -22 m, interpreted as a possible slowstand. On the basis of seismic evidence in the outer Lough, an as yet undated lowstand at -42 m is tentatively interpreted to pre-date this stillstand. These results will be

  8. Post-glacial sea-level history for NE Ireland (Belfast Lough) based on offshore evidence

    NASA Astrophysics Data System (ADS)

    Quinn, Rory; Plets, Ruth; Callard, Louise; Cooper, Andrew; Antony, Long; Daniel, Belknap; Robin, Edwards; Derek, Jackson; Joseph, Kelley; David, Long; Glenn, Milne; Xavier, Monteys

    2014-05-01

    Glacio-isostatic adjustment (GIA) models suggest a complex relative sea-level (RSL) pattern around the Irish Sea Basin after the Last Glacial Maximum (LGM), with modelled sea-level lowstands ranging from -12 m in the north to greater than -60 m in the south of the Basin. However, these GIA models are poorly constrained by observational data offshore. Belfast Lough, on the NE coast of Ireland, is one of seven sites chosen to investigate this complex RSL history as part of the project 'Late Glacial Sea level minima in the Western British Isles' (NERC NE/H024301/1). Belfast Lough was chosen as one of the candidate sites on the basis of location (at the northern end of the Irish Sea Basin), sedimentary environment (grossly depositional) and the fact that the lowstand predicted for the Belfast Lough area by a recent version of the GIA model (-16.5 m) differs significantly from the (limited) extant observational data, which interprets the lowstand at -30 m. In 2011 and 2012 we gathered new multi-beam echo-sounder data, >200 km trackline pinger- and boomer- seismic reflection data and 46 vibrocores in Belfast Lough. Radiocarbon dating and palaeoenvironmental analysis are used to constrain the interpretation of the seismic and sediment data. Five seismo-stratigraphic units are interpreted, with a distinct erosional surface between U3 and U4 interpreted as a transgressive surface associated with sea level rise post-dating a RSL lowstand. Foraminiferal evidence indicates an increase in marine species (from lagoonal/estuarine to fully marine) from U4 to U5. Integration of the seismic and core data indicate an erosional event prior to 12.7 cal yr BP resulting in a planated surface in the inner Lough and wave-eroded drumlins at the mouth of the Lough between -15 and -22 m, interpreted as a possible slowstand. On the basis of seismic evidence in the outer Lough, an as yet undated lowstand at -42 m is tentatively interpreted to pre-date this stillstand. These results will be used

  9. The ancient tropical rainforest tree Symphonia globulifera L. f. (Clusiaceae) was not restricted to postulated Pleistocene refugia in Atlantic Equatorial Africa

    PubMed Central

    Budde, K B; González-Martínez, S C; Hardy, O J; Heuertz, M

    2013-01-01

    Understanding the history of forests and their species' demographic responses to past disturbances is important for predicting impacts of future environmental changes. Tropical rainforests of the Guineo-Congolian region in Central Africa are believed to have survived the Pleistocene glacial periods in a few major refugia, essentially centred on mountainous regions close to the Atlantic Ocean. We tested this hypothesis by investigating the phylogeographic structure of a widespread, ancient rainforest tree species, Symphonia globulifera L. f. (Clusiaceae), using plastid DNA sequences (chloroplast DNA [cpDNA], psbA-trnH intergenic spacer) and nuclear microsatellites (simple sequence repeats, SSRs). SSRs identified four gene pools located in Benin, West Cameroon, South Cameroon and Gabon, and São Tomé. This structure was also apparent at cpDNA. Approximate Bayesian Computation detected recent bottlenecks approximately dated to the last glacial maximum in Benin, West Cameroon and São Tomé, and an older bottleneck in South Cameroon and Gabon, suggesting a genetic effect of Pleistocene cycles of forest contraction. CpDNA haplotype distribution indicated wide-ranging long-term persistence of S. globulifera both inside and outside of postulated forest refugia. Pollen flow was four times greater than that of seed in South Cameroon and Gabon, which probably enabled rapid population recovery after bottlenecks. Furthermore, our study suggested ecotypic differentiation—coastal or swamp vs terra firme—in S. globulifera. Comparison with other tree phylogeographic studies in Central Africa highlighted the relevance of species-specific responses to environmental change in forest trees. PMID:23572126

  10. Climate oscillations, glacial refugia, and dispersal ability: factors influencing the genetic structure of the least salmonfly, Pteronarcella badia (Plecoptera), in Western North America.

    PubMed

    Sproul, John S; Houston, Derek D; Nelson, C Riley; Evans, R Paul; Crandall, Keith A; Shiozawa, Dennis K

    2015-12-12

    Phylogeographic studies of aquatic insects provide valuable insights into mechanisms that shape the genetic structure of communities, yet studies that include broad geographic areas are uncommon for this group. We conducted a broad scale phylogeographic analysis of the least salmonfly Pteronarcella badia (Plecoptera) across western North America. We tested hypotheses related to mode of dispersal and the influence of historic climate oscillations on population genetic structure. In order to generate a larger mitochondrial data set, we used 454 sequencing to reconstruct the complete mitochondrial genome in the early stages of the project. Our analysis revealed high levels of population structure with several deeply divergent clades present across the sample area. Evidence from five mitochondrial genes and one nuclear locus identified a potentially cryptic lineage in the Pacific Northwest. Gene flow estimates and geographic clade distributions suggest that overland flight during the winged adult stage is an important dispersal mechanism for this taxon. We found evidence of multiple glacial refugia across the species distribution and signs of secondary contact within and among major clades. This study provides a basis for future studies of aquatic insect phylogeography at the inter-basin scale in western North America. Our findings add to an understanding of the role of historical climate isolations in shaping assemblages of aquatic insects in this region. We identified several geographic areas that may have historical importance for other aquatic organisms with similar distributions and dispersal strategies as P. badia. This work adds to the ever-growing list of studies that highlight the potential of next-generation DNA sequencing in a phylogenetic context to improve molecular data sets from understudied groups.

  11. Pleistocene range shifts, refugia and the origin of widespread species in western Palaearctic water beetles.

    PubMed

    García-Vázquez, David; Bilton, David T; Foster, Garth N; Ribera, I

    2017-09-01

    Quaternary glacial cycles drove major shifts in both the extent and location of the geographical ranges of many organisms. During glacial maxima, large areas of central and northern Europe were inhospitable to temperate species, and these areas are generally assumed to have been recolonized during interglacials by range expansions from Mediterranean refugia. An alternative is that this recolonization was from non-Mediterranean refugia, in central Europe or western Asia, but data on the origin of widespread central and north European species remain fragmentary, especially for insects. We studied three widely distributed lineages of freshwater beetles (the Platambus maculatus complex, the Hydraena gracilis complex, and the genus Oreodytes), all restricted to running waters and including both narrowly distributed southern endemics and widespread European species, some with distributions spanning the Palearctic. Our main goal was to determine the role of the Pleistocene glaciations in shaping the diversification and current distribution of these lineages. We sequenced four mitochondrial and two nuclear genes in populations drawn from across the ranges of these taxa, and used Bayesian probabilities and Maximum Likelihood to reconstruct their phylogenetic relationships, age and geographical origin. Our results suggest that all extant species in these groups are of Pleistocene origin. In the H. gracilis complex, the widespread European H. gracilis has experienced a rapid, recent range expansion from northern Anatolia, to occupy almost the whole of Europe. However, in the other two groups widespread central and northern European taxa appear to originate from central Asia, rather than the Mediterranean. These widespread species of eastern origin typically have peripherally isolated forms in the southern Mediterranean peninsulas, which may be remnants of earlier expansion-diversification cycles or result from incipient isolation of populations during the most recent Holocene

  12. The vegetation cover of New Zealand at the Last Glacial Maximum

    NASA Astrophysics Data System (ADS)

    Newnham, Rewi; McGlone, Matt; Moar, Neville; Wilmshurst, Janet; Vandergoes, Marcus

    2013-08-01

    A new reconstruction of the vegetation cover for New Zealand at the Last Glacial Maximum (LGM) is presented, based primarily on a database of 66 pollen site records and a more limited range of plant macrofossil and coleopteran records. Extensive forest is evident only from Auckland northwards. Conifer-broadleaf forest similar to that in the region today, but with Agathis australis scarce, persisted in the far north, whilst Nothofagus trees and a range of shrub taxa characterised the more open forests elsewhere in Northland. Survival of Nothofagus-dominated forest in coastal and exposed continental shelf locations to the southwest of Auckland and northwestern South Island is also indicated. Beyond these regions, vegetation cover comprised shrubland- and grassland-dominant communities, with the latter more prominent in eastern areas, to the south and presumably at higher altitudes. Nevertheless the survival of forest trees is indicated unambiguously in most regions apart from the eastern South Island. Thus the concept of 'micro glacial forest refugia' in New Zealand remains supported by this latest glacial vegetation reconstruction and we draw possible parallels with the developing but contentious concept of 'northern cryptic refugia' in Europe. Recent assertions that pollen and beetle reconstructions of the New Zealand LGM vegetation patterns diverge significantly are not supported by this analysis. Rather, the two proxies are readily reconciled if the term 'woody' as indicated by coleoptera is not restricted to tall forest trees but extended to the widespread woody shrub and small tree elements of the New Zealand flora. Regional distinctions in the LGM vegetation reconstruction concur broadly with the contemporary vegetation pattern, suggesting that, along with temperature depression and likely drier growing conditions, a zonal circulation regime with prominent southern westerly winds was important at 21 ka, as it is today. Pollen-climate modelling of the extent of

  13. Uncovering the glacial history of the Irish continental shelf (Invited)

    NASA Astrophysics Data System (ADS)

    Dunlop, P.; Benetti, S.; OCofaigh, C.

    2013-12-01

    In 1999 the Irish Government initiated a €32 million survey of its territorial waters known as the Irish National Seabed Survey (INSS). The INSS is amongst the largest marine mapping programmes ever undertaken anywhere in the world and provides high-resolution multibeam, backscatter and seismic data of the seabed around Ireland. These data have been used to provide the first clear evidence for extensive glaciation of the continental shelf west and northwest of Ireland. Streamlined drumlins on the mid to outer shelf record former offshore-directed ice flow towards the shelf edge and show that the ice sheet was grounded in a zone of confluence where ice flowing onto the shelf from northwest Ireland merged with ice flowing across the Malin Shelf from southwest Scotland. The major glacial features on the shelf are well developed nested arcuate moraine systems that mark the position of the ice sheet margin and confirm that the former British Irish Ice Sheet was grounded as far as the shelf edge around 100 km offshore of west Donegal at the last glacial maximum. Distal to the moraines, on the outermost shelf, prominent zones of iceberg plough marks give way to the Barra/Donegal fan and a well developed system of gullies and canyons which incise the continental slope. Since 2008 several scientific cruises have retrieved cores from the shelf and slope to help build a more detailed understanding of glacial events in this region. This presentation will provide an overview of the glacial history of the Irish shelf and will discuss ongoing research programmes that are building on the initial research findings to produce a better understanding of the nature and timing of ice sheet events in this region.

  14. Last Glacial Maximum cirque glaciation in Ireland and implications for reconstructions of the Irish Ice Sheet

    NASA Astrophysics Data System (ADS)

    Barth, Aaron M.; Clark, Peter U.; Clark, Jorie; McCabe, A. Marshall; Caffee, Marc

    2016-06-01

    Reconstructions of the extent and height of the Irish Ice Sheet (IIS) during the Last Glacial Maximum (LGM, ∼19-26 ka) are widely debated, in large part due to limited age constraints on former ice margins and due to uncertainties in the origin of the trimlines. A key area is southwestern Ireland, where various LGM reconstructions range from complete coverage by a contiguous IIS that extends to the continental shelf edge to a separate, more restricted southern-sourced Kerry-Cork Ice Cap (KCIC). We present new 10Be surface exposure ages from two moraines in a cirque basin in the Macgillycuddy's Reeks that provide a unique and unequivocal constraint on ice thickness for this region. Nine 10Be ages from an outer moraine yield a mean age of 24.5 ± 1.4 ka while six ages from an inner moraine yield a mean age of 20.4 ± 1.2 ka. These ages show that the northern flanks of the Macgillycuddy's Reeks were not covered by the IIS or a KCIC since at least 24.5 ± 1.4 ka. If there was more extensive ice coverage over the Macgillycuddy's Reeks during the LGM, it occurred prior to our oldest ages.

  15. Coalescent Simulation and Paleodistribution Modeling for Tabebuia rosealba Do Not Support South American Dry Forest Refugia Hypothesis

    PubMed Central

    de Melo, Warita Alves; Lima-Ribeiro, Matheus S.; Terribile, Levi Carina

    2016-01-01

    Studies based on contemporary plant occurrences and pollen fossil records have proposed that the current disjunct distribution of seasonally dry tropical forests (SDTFs) across South America is the result of fragmentation of a formerly widespread and continuously distributed dry forest during the arid climatic conditions associated with the Last Glacial Maximum (LGM), which is known as the modern-day dry forest refugia hypothesis. We studied the demographic history of Tabebuia rosealba (Bignoniaceae) to understand the disjunct geographic distribution of South American SDTFs based on statistical phylogeography and ecological niche modeling (ENM). We specifically tested the dry forest refugia hypothesis; i.e., if the multiple and isolated patches of SDTFs are current climatic relicts of a widespread and continuously distributed dry forest during the LGM. We sampled 235 individuals across 18 populations in Central Brazil and analyzed the polymorphisms at chloroplast (trnS-trnG, psbA-trnH and ycf6-trnC intergenic spacers) and nuclear (ITS nrDNA) genomes. We performed coalescence simulations of alternative hypotheses under demographic expectations from two a priori biogeographic hypotheses (1. the Pleistocene Arc hypothesis and, 2. a range shift to Amazon Basin) and other two demographic expectances predicted by ENMs (3. expansion throughout the Neotropical South America, including Amazon Basin, and 4. retraction during the LGM). Phylogenetic analyses based on median-joining network showed haplotype sharing among populations with evidence of incomplete lineage sorting. Coalescent analyses showed smaller effective population sizes for T. roseoalba during the LGM compared to the present-day. Simulations and ENM also showed that its current spatial pattern of genetic diversity is most likely due to a scenario of range retraction during the LGM instead of the fragmentation from a once extensive and largely contiguous SDTF across South America, not supporting the South

  16. Coalescent Simulation and Paleodistribution Modeling for Tabebuia rosealba Do Not Support South American Dry Forest Refugia Hypothesis.

    PubMed

    de Melo, Warita Alves; Lima-Ribeiro, Matheus S; Terribile, Levi Carina; Collevatti, Rosane G

    2016-01-01

    Studies based on contemporary plant occurrences and pollen fossil records have proposed that the current disjunct distribution of seasonally dry tropical forests (SDTFs) across South America is the result of fragmentation of a formerly widespread and continuously distributed dry forest during the arid climatic conditions associated with the Last Glacial Maximum (LGM), which is known as the modern-day dry forest refugia hypothesis. We studied the demographic history of Tabebuia rosealba (Bignoniaceae) to understand the disjunct geographic distribution of South American SDTFs based on statistical phylogeography and ecological niche modeling (ENM). We specifically tested the dry forest refugia hypothesis; i.e., if the multiple and isolated patches of SDTFs are current climatic relicts of a widespread and continuously distributed dry forest during the LGM. We sampled 235 individuals across 18 populations in Central Brazil and analyzed the polymorphisms at chloroplast (trnS-trnG, psbA-trnH and ycf6-trnC intergenic spacers) and nuclear (ITS nrDNA) genomes. We performed coalescence simulations of alternative hypotheses under demographic expectations from two a priori biogeographic hypotheses (1. the Pleistocene Arc hypothesis and, 2. a range shift to Amazon Basin) and other two demographic expectances predicted by ENMs (3. expansion throughout the Neotropical South America, including Amazon Basin, and 4. retraction during the LGM). Phylogenetic analyses based on median-joining network showed haplotype sharing among populations with evidence of incomplete lineage sorting. Coalescent analyses showed smaller effective population sizes for T. roseoalba during the LGM compared to the present-day. Simulations and ENM also showed that its current spatial pattern of genetic diversity is most likely due to a scenario of range retraction during the LGM instead of the fragmentation from a once extensive and largely contiguous SDTF across South America, not supporting the South

  17. Multilocus phylogeography and population structure of common eiders breeding in North America and Scandinavia

    USGS Publications Warehouse

    Sonsthagen, Sarah A.; Talbot, Sandra L.; Scribner, Kim T.; McCracken, Kevin G.

    2011-01-01

    Aim  Glacial refugia during the Pleistocene had major impacts on the levels and spatial apportionment of genetic diversity of species in northern latitude ecosystems. We characterized patterns of population subdivision, and tested hypotheses associated with locations of potential Pleistocene refugia and the relative contribution of these refugia to the post-glacial colonization of North America and Scandinavia by common eiders (Somateria mollissima). Specifically, we evaluated localities hypothesized as ice-free areas or glacial refugia for other Arctic vertebrates, including Beringia, the High Arctic Canadian Archipelago, Newfoundland Bank, Spitsbergen Bank and north-west Norway.Location  Alaska, Canada, Norway and Sweden.Methods  Molecular data from 12 microsatellite loci, the mitochondrial DNA (mtDNA) control region, and two nuclear introns were collected and analysed for 15 populations of common eiders (n = 716) breeding throughout North America and Scandinavia. Population genetic structure, historical population fluctuations and gene flow were inferred using F-statistics, analyses of molecular variance, and multilocus coalescent analyses.Results  Significant inter-population variation in allelic and haplotypic frequencies were observed (nuclear DNA FST = 0.004–0.290; mtDNA ΦST = 0.051–0.927). Whereas spatial differentiation in nuclear genes was concordant with subspecific designations, geographic proximity was more predictive of inter-population variance in mitochondrial DNA haplotype frequency. Inferences of historical population demography were consistent with restriction of common eiders to four geographic areas during the Last Glacial Maximum: Belcher Islands, Newfoundland Bank, northern Alaska and Svalbard. Three of these areas coincide with previously identified glacial refugia: Newfoundland Bank, Beringia and Spitsbergen Bank. Gene-flow and clustering analyses indicated that the Beringian refugium contributed little to common

  18. Use of on-site refugia to protect unionid populations from zebra mussel-induced mortality

    USGS Publications Warehouse

    Nichols, S. Jerrine; Black, M. Glen; Allen, Jeffrey D.

    2000-01-01

    Protecting unionid populations as zebra mussels spread into inland waterways has relied mainly on relocating at-risk animals into aquaculture facilities. While such relocations are the only viable management technique for some populations, facility availability is limited, leaving many unionids facing extirpation. Another management strategy is in-situ protection either by enhancing natural refugia or by creating managed refugia. We have reviewed all reports of natural refugia and found that refugia for unionids can be found in many areas. There are many habitats where zebra mussel colonization has been limited, or of a temporary nature. Within zebra mussel infested areas, unionid communities continue to survive in some shallow water sites such as estuaries, deltas, and lake-connected wetlands. Managed refugia can be created in areas where natural refugia do not exist. We present a case study on recent efforts to create refugia in an area with rapidly expanding zebra mussel populations. Preliminary analysis of unionid body condition indicates that removal of encrusted zebra mussels only once a year can improve unionid condition factors and decrease mortality. Natural and managed refugia can provide an additional conservation management option in some areas.

  19. Paleodistribution modeling suggests glacial refugia in Scandinavia and out-of-Tibet range expansion of the Arctic fox.

    PubMed

    Fuentes-Hurtado, Marcelo; Hof, Anouschka R; Jansson, Roland

    2016-01-01

    Quaternary glacial cycles have shaped the geographic distributions and evolution of numerous species in the Arctic. Ancient DNA suggests that the Arctic fox went extinct in Europe at the end of the Pleistocene and that Scandinavia was subsequently recolonized from Siberia, indicating inability to track its habitat through space as climate changed. Using ecological niche modeling, we found that climatically suitable conditions for Arctic fox were found in Scandinavia both during the last glacial maximum (LGM) and the mid-Holocene. Our results are supported by fossil occurrences from the last glacial. Furthermore, the model projection for the LGM, validated with fossil records, suggested an approximate distance of 2000 km between suitable Arctic conditions and the Tibetan Plateau well within the dispersal distance of the species, supporting the recently proposed hypothesis of range expansion from an origin on the Tibetan Plateau to the rest of Eurasia. The fact that the Arctic fox disappeared from Scandinavia despite suitable conditions suggests that extant populations may be more sensitive to climate change than previously thought.

  20. Extreme Glacial Legacies: A Synthesis of the Antarctic Springtail Phylogeographic Record

    PubMed Central

    McGaughran, Angela; Stevens, Mark I.; Hogg, Ian D.; Carapelli, Antonio

    2011-01-01

    We review current phylogeographic knowledge from across the Antarctic terrestrial landscape with a focus on springtail taxa. We describe consistent patterns of high genetic diversity and structure among populations which have persisted in glacial refugia across Antarctica over both short (<2 Mya) and long (>10 Mya) timescales. Despite a general concordance of results among species, we explain why location is important in determining population genetic patterns within bioregions. We complete our review by drawing attention to the main limitations in the field of Antarctic phylogeography, namely that the scope of geographic focus is often lacking within studies, and that large gaps remain in our phylogeographic knowledge for most terrestrial groups. PMID:26467614

  1. Surviving in mountain climate refugia: new insights from the genetic diversity and structure of the relict shrub Myrtus nivellei (Myrtaceae) in the Sahara Desert.

    PubMed

    Migliore, Jérémy; Baumel, Alex; Juin, Marianick; Fady, Bruno; Roig, Anne; Duong, Nathalie; Médail, Frédéric

    2013-01-01

    The identification of past glacial refugia has become a key topic for conservation under environmental change, since they contribute importantly to shaping current patterns of biodiversity. However, little attention has been paid so far to interglacial refugia despite their key role for the survival of relict species currently occurring in climate refugia. Here, we focus on the genetic consequences of range contraction on the relict populations of the evergreen shrub Myrtus nivellei, endemic in the Saharan mountains since at least the end of the last Green Sahara period, around 5.5 ka B.P. Multilocus genotypes (nuclear microsatellites and AFLP) were obtained from 215 individuals collected from 23 wadis (temporary rivers) in the three main mountain ranges in southern Algeria (the Hoggar, Tassili n'Ajjer and Tassili n'Immidir ranges). Identical genotypes were found in several plants growing far apart within the same wadis, a pattern taken as evidence of clonality. Multivariate analyses and Bayesian clustering revealed that genetic diversity was mainly structured among the mountain ranges, while low isolation by distance was observed within each mountain range. The range contraction induced by the last episode of aridification has likely increased the genetic isolation of the populations of M. nivellei, without greatly affecting the genetic diversity of the species as a whole. The pattern of genetic diversity observed here suggests that high connectivity may have prevailed during humid periods, which is consistent with recent paleoenvironmental reconstructions.

  2. Did debris-covered glaciers serve as pleistocene refugia for plants? A new hypothesis derived from observations of recent plant growth on glacier surfaces

    USGS Publications Warehouse

    Fickert, T.; Friend, D.; Gruninger, F.; Molnia, B.; Richter, M.

    2007-01-01

    This study proposes a new hypothesis: Debris-covered glaciers served as Pleistocene biological refugia. This is based on detailed studies of vascular plant growth on six debris-mantled glaciers, literally around the world, as well as many casual observations also across the globe. We find that such glaciers are quite common and are distributed globally. Using Carbon Glacier, Mount Rainier, U.S.A., as a type locality and case study, we show aspects of the floristic and structural diversity as well as spatial patterns of plant growth on the glacier surface. Migration strategies, root characteristics, and origin and dispersal strategies for vascular plant species are documented. Also reported are special microclimatic conditions in these areas allowing for this remarkable plant ecology. We find that alpine taxa can grow considerably below their usual altitudinal niche due to the cooler subsurface soil temperatures found on glacial debris with ice underneath, and that may have significantly altered the spatial distribution of such flora during full glacial conditions. This in turn creates previously undocumented areas from which alpine, and perhaps arctic, plant species reestablished in post-glacial time. This hypothesis is complementary to both the nunatak hypothesis and tabula rasa theory and possibly helps solve the ongoing controversy between them. ?? 2007 Regents of the University of Colorado.

  3. Reef-coral refugia in a rapidly changing ocean.

    PubMed

    Cacciapaglia, Chris; van Woesik, Robert

    2015-06-01

    This study sought to identify climate-change thermal-stress refugia for reef corals in the Indian and Pacific Oceans. A species distribution modeling approach was used to identify refugia for 12 coral species that differed considerably in their local response to thermal stress. We hypothesized that the local response of coral species to thermal stress might be similarly reflected as a regional response to climate change. We assessed the contemporary geographic range of each species and determined their temperature and irradiance preferences using a k-fold algorithm to randomly select training and evaluation sites. That information was applied to downscaled outputs of global climate models to predict where each species is likely to exist by the year 2100. Our model was run with and without a 1°C capacity to adapt to the rising ocean temperature. The results show a positive exponential relationship between the current area of habitat that coral species occupy and the predicted area of habitat that they will occupy by 2100. There was considerable decoupling between scales of response, however, and with further ocean warming some 'winners' at local scales will likely become 'losers' at regional scales. We predicted that nine of the 12 species examined will lose 24-50% of their current habitat. Most reductions are predicted to occur between the latitudes 5-15°, in both hemispheres. Yet when we modeled a 1°C capacity to adapt, two ubiquitous species, Acropora hyacinthus and Acropora digitifera, were predicted to retain much of their current habitat. By contrast, the thermally tolerant Porites lobata is expected to increase its current distribution by 14%, particularly southward along the east and west coasts of Australia. Five areas were identified as Indian Ocean refugia, and seven areas were identified as Pacific Ocean refugia for reef corals under climate change. All 12 of these reef-coral refugia deserve high-conservation status. © 2015 John Wiley & Sons Ltd.

  4. Ploidy race distributions since the Last Glacial Maximum in the North American desert shrub, Larrea tridentata

    USGS Publications Warehouse

    Hunter, K.L.; Betancourt, J.L.; Riddle, B.R.; Van Devender, T. R.; Cole, K.L.; Geoffrey, Spaulding W.

    2000-01-01

    1 A classic biogeographic pattern is the alignment of diploid, tetraploid and hexaploid races of creosote bush (Larrea tridentata) across the Chihuahuan, Sonoran and Mohave Deserts of western North America. We used statistically robust differences in guard cell size of modern plants and fossil leaves from packrat middens to map current and past distributions of these ploidy races since the Last Glacial Maximum (LGM). 2 Glacial/early Holocene (26-10 14C kyr BP or thousands of radiocarbon years before present) populations included diploids along the lower Rio Grande of west Texas, 650 km removed from sympatric diploids and tetraploids in the lower Colorado River Basin of south-eastern California/south-western Arizona. Diploids migrated slowly from lower Rio Grande refugia with expansion into the northern Chihuahuan Desert sites forestalled until after ???4.0 14C kyr BP. Tetraploids expanded from the lower Colorado River Basin into the northern limits of the Sonoran Desert in central Arizona by 6.4 14C kyr BP. Hexaploids appeared by 8.5 14C kyr BP in the lower Colorado River Basin, reaching their northernmost limits (???37??N) in the Mohave Desert between 5.6 and 3.9 14C kyr BP. 3 Modern diploid isolates may have resulted from both vicariant and dispersal events. In central Baja California and the lower Colorado River Basin, modern diploids probably originated from relict populations near glacial refugia. Founder events in the middle and late Holocene established diploid outposts on isolated limestone outcrops in areas of central and southern Arizona dominated by tetraploid populations. 4 Geographic alignment of the three ploidy races along the modern gradient of increasingly drier and hotter summers is clearly a postglacial phenomenon, but evolution of both higher ploidy races must have happened before the Holocene. The exact timing and mechanism of polyploidy evolution in creosote bush remains a matter of conjecture. ?? 2001 Blackwell Science Ltd.

  5. Ploidy race distributions since the Last Glacial Maximum in the North American desert shrub, Larea tridentata

    USGS Publications Warehouse

    Hunter, Kimberly L.; Betancourt, Julio L.; Riddle, Brett R.; Van Devender, Thomas R.; Cole, K.L.; Spaulding, W.G.

    2001-01-01

    1. A classic biogeographic pattern is the alignment of diploid, tetraploid and hexaploid races of creosote bush (Larrea tridentata) across the Chihuahuan, Sonoran and Mohave Deserts of western North America. We used statistically robust differences in guard cell size of modern plants and fossil leaves from packrat middens to map current and past distributions of these ploidy races since the Last Glacial Maximum (LGM). 2 Glacial/early Holocene (26a??10 14C kyr bp or thousands of radiocarbon years before present) populations included diploids along the lower Rio Grande of west Texas, 650 km removed from sympatric diploids and tetraploids in the lower Colorado River Basin of south-eastern California/south-western Arizona. Diploids migrated slowly from lower Rio Grande refugia with expansion into the northern Chihuahuan Desert sites forestalled until after ~4.0 14C kyr bp. Tetraploids expanded from the lower Colorado River Basin into the northern limits of the Sonoran Desert in central Arizona by 6.4 14C kyr bp. Hexaploids appeared by 8.5 14C kyr bp in the lower Colorado River Basin, reaching their northernmost limits (~37A?N) in the Mohave Desert between 5.6 and 3.9 14C kyr bp. 3 Modern diploid isolates may have resulted from both vicariant and dispersal events. In central Baja California and the lower Colorado River Basin, modern diploids probably originated from relict populations near glacial refugia. Founder events in the middle and late Holocene established diploid outposts on isolated limestone outcrops in areas of central and southern Arizona dominated by tetraploid populations. 4 Geographic alignment of the three ploidy races along the modern gradient of increasingly drier and hotter summers is clearly a postglacial phenomenon, but evolution of both higher ploidy races must have happened before the Holocene. The exact timing and mechanism of polyploidy evolution in creosote bush remains a matter of conjecture.

  6. Loss of thermal refugia near equatorial range limits.

    PubMed

    Lima, Fernando P; Gomes, Filipa; Seabra, Rui; Wethey, David S; Seabra, Maria I; Cruz, Teresa; Santos, António M; Hilbish, Thomas J

    2016-01-01

    This study examines the importance of thermal refugia along the majority of the geographical range of a key intertidal species (Patella vulgata Linnaeus, 1758) on the Atlantic coast of Europe. We asked whether differences between sun-exposed and shaded microhabitats were responsible for differences in physiological stress and ecological performance and examined the availability of refugia near equatorial range limits. Thermal differences between sun-exposed and shaded microhabitats are consistently associated with differences in physiological performance, and the frequency of occurrence of high temperatures is most probably limiting the maximum population densities supported at any given place. Topographical complexity provides thermal refugia throughout most of the distribution range, although towards the equatorial edges the magnitude of the amelioration provided by shaded microhabitats is largely reduced. Importantly, the limiting effects of temperature, rather than being related to latitude, seem to be tightly associated with microsite variability, which therefore is likely to have profound effects on the way local populations (and consequently species) respond to climatic changes. © 2015 John Wiley & Sons Ltd.

  7. Using biological data to test climate change refugia

    NASA Astrophysics Data System (ADS)

    Morelli, T. L.; Maher, S. P.

    2015-12-01

    The concept of refugia has been discussed from theoretical and paleontological perspectives to address how populations persisted during periods of unfavorable climate. Recently, several studies have applied the idea to contemporary landscapes to identify locations that are buffered from climate change effects so as to favor greater persistence of valued resources relative to other areas. Refugia are now being discussed among natural resource agencies as a potential adaptation option in the face of anthropogenic climate change. Using downscaled climate data, we identified hypothetical refugial meadows in the Sierra Nevada and then tested them using survey and genetic data from Belding's ground squirrel (Urocitellus beldingi) populations. We predicted that refugial meadows would show higher genetic diversity, higher rates of occupancy and lower rates of extirpation over time. At each step of the research, we worked with managers to ensure the largest impact. Although no panacea, identifying climate change refugia could be an important strategy for prioritizing habitats for management intervention in order to conserve populations. This research was supported by the California LCC, the Northeast Climate Science Center, and NSF.

  8. Refugia Research Coalition: A regional-scale approach for connecting refugia science to natural and cultural resource management

    EPA Science Inventory

    Background / question / methods Warmer air and water temperatures, changing precipitation patterns, and altered fire regimes associated with climate change threaten many important natural and cultural resources. Climate change refugia are areas relatively buffered from contempora...

  9. Shallow phylogeographic structuring of Vimba vimba across Europe suggests two distinct refugia during the last glaciation.

    PubMed

    Hänfling, B; Dümpelmann, C; Bogutskaya, N G; Brandl, R; Brändle, M

    2009-12-01

    Genetic variation and geographical structuring of vimba Vimba vimba were analysed across 26 sites (80 individuals) by means of mtDNA sequences (cyt b gene, mitochondrial control region) to localize hypothesized glacial refugia and to reconstruct postglacial recoloniation routes. Although genetic diversity among sequenced individuals was low, a combined analysis of the two sequenced fragments revealed a western (central and northern Europe: Danube, Elbe and lakes of Sweden) and an eastern clade (eastern Europe: Dnieper-South Bug, Don, Neman). Furthermore, a number of divergent ancestral haplotypes distributed around the Black and Caspian Seas became apparent. Mismatch analyses supported a sudden expansion model for the populations of the western clade between 50 and 10 000 bp. Overall, the study provides strong evidence for a northward and westward expansion of V. vimba from two refugial regions located in the Danubian drainage and the northern Pontic regions respectively.

  10. Science questions for implementing climate refugia for salmon as a conservation strategy

    EPA Science Inventory

    The recognition and protection of climate refugia has been proposed as a potential adaptation strategy that may be useful for protecting the biotic integrity of watersheds under a changing climate. Climate refugia are areas that are buffered from climate change effects relative t...

  11. Gridded climate data from 5 GCMs of the Last Glacial Maximum downscaled to 30 arc s for Europe

    NASA Astrophysics Data System (ADS)

    Schmatz, D. R.; Luterbacher, J.; Zimmermann, N. E.; Pearman, P. B.

    2015-06-01

    Studies of the impacts of historical, current and future global change require very high-resolution climate data (≤ 1 km) as a basis for modelled responses, meaning that data from digital climate models generally require substantial rescaling. Another shortcoming of available datasets on past climate is that the effects of sea level rise and fall are not considered. Without such information, the study of glacial refugia or early Holocene plant and animal migration are incomplete if not impossible. Sea level at the last glacial maximum (LGM) was approximately 125 m lower, creating substantial additional terrestrial area for which no current baseline data exist. Here, we introduce the development of a novel, gridded climate dataset for LGM that is both very high resolution (1 km) and extends to the LGM sea and land mask. We developed two methods to extend current terrestrial precipitation and temperature data to areas between the current and LGM coastlines. The absolute interpolation error is less than 1 and 0.5 °C for 98.9 and 87.8 %, respectively, of all pixels within two arc degrees of the current coastline. We use the change factor method with these newly assembled baseline data to downscale five global circulation models of LGM climate to a resolution of 1 km for Europe. As additional variables we calculate 19 "bioclimatic" variables, which are often used in climate change impact studies on biological diversity. The new LGM climate maps are well suited for analysing refugia and migration during Holocene warming following the LGM.

  12. A review of pipe and bamboo artificial refugia as sampling tools in anuran studies

    USGS Publications Warehouse

    Glorioso, Brad M.; Waddle, J. Hardin

    2014-01-01

    Artificial pipe-like refugia have been used for more than 40 years in anuran studies, and have captured 28 species, primarily (82%) hylid treefrogs. Early pipe-like refugia were made using cut pieces of bamboo in the tropical forests of Puerto Rico, but most recent studies have used synthetic pipes and have occurred primarily in the southeastern United States. Characteristics of artificial refugia (e.g., color, length, and diameter), and their placement in the environment have varied greatly among studies, making comparisons difficult. Here, we summarize and evaluate different pipe designs and placement, address potential concerns when using artificial pipe-like refugia, and suggest studies necessary to better interpret the data gained from this technique in anuran studies.

  13. Migration Patterns of Subgenus Alnus in Europe since the Last Glacial Maximum: A Systematic Review

    PubMed Central

    Douda, Jan; Doudová, Jana; Drašnarová, Alena; Kuneš, Petr; Hadincová, Věroslava; Krak, Karol; Zákravský, Petr; Mandák, Bohumil

    2014-01-01

    Background/Aims Recently, new palaeoecological records supported by molecular analyses and palaeodistributional modelling have provided more comprehensive insights into plant behaviour during the last Quaternary cycle. We reviewed the migration history of species of subgenus Alnus during the last 50,000 years in Europe with a focus on (1) a general revision of Alnus history since the Last Glacial Maximum (LGM), (2) evidence of northern refugia of Alnus populations during the LGM and (3) the specific history of Alnus in particular European regions. Methodology We determined changes in Alnus distribution on the basis of 811 and 68 radiocarbon-dated pollen and macrofossil sites, respectively. We compiled data from the European Pollen Database, the Czech Quaternary Palynological Database, the Eurasian Macrofossil Database and additional literature. Pollen percentage thresholds indicating expansions or retreats were used to describe patterns of past Alnus occurrence. Principal Findings An expansion of Alnus during the Late Glacial and early Holocene periods supports the presence of alders during the LGM in southern peninsulas and northerly areas in western Europe, the foothills of the Alps, the Carpathians and northeastern Europe. After glaciers withdrew, the ice-free area of Europe was likely colonized from several regional refugia; the deglaciated area of Scandinavia was likely colonized from a single refugium in northeastern Europe. In the more northerly parts of Europe, we found a scale-dependent pattern of Alnus expansion characterised by a synchronous increase of Alnus within individual regions, though with regional differences in the times of the expansion. In southern peninsulas, the Alps and the Carpathians, by contrast, it seems that Alnus expanded differently at individual sites rather than synchronously in whole regions. Conclusions Our synthesis supports the idea that northern LGM populations were important sources of postglacial Alnus expansion. The

  14. A Late-Glacial sedimentary sequence at KIlkeel, Northern Ireland: implications for the glaciation of the Irish Sea Basin

    NASA Astrophysics Data System (ADS)

    Merritt, Jon; Roberson, Sam; Cooper, Mark

    2017-04-01

    This paper re-evaluates the nature and timing of a Late-Glacial ice sheet re-advance in the north western sector of the Irish Sea basin. The sedimentary archive in the region records the collapse of the Irish Sea Ice Stream, a major outlet glacier of the British-Irish Ice Sheet. The region documents the interplay between southerly flowing Scottish ice, ice flowing southeast from Lough Neagh and locally sourced Mournes ice. We present the results of sedimentological analysis of a glacigenic sequence exposed in a modern cliff section 3 km long between Derryoge and Kilkeel, Co. Down, Northern Ireland. The interaction between an advancing ice-sheet outlet lobe and rapidly changing sea levels are examined using facies analysis and micromorphology. The section is composed of four lithofacies associations (LAs). These are, from the base, a laminated, fossiliferous and deformed silt (LA1) at least 4.5 m thick that contains lenses of diamicton and discontinuous rafts of sandy gravel. Marine shells form the axis of a fold hinge, part of a lightly tectonised channel fill within the raft. LA1 is overlain by a sandy diamict (LA2) up to 14 m thick containing mainly local clasts with some of northern provenance. Within LA2 are wide channel structures infilled by laminated clayey silts (LA2b). These form deposits up to 14 m thick and contain small-scale folds, discrete shear zones and ball-and-pillow structures. LA2b forms a lithofacies association with LA2, consisting of a lower subfacies of sheared and deformed silts, overlain by sandy diamicton, capped by a striated boulder pavement. These are interpreted to represent retreat/advance cycles of a marine terminating ice margin. Up to five such cycles are identified. LA2 is widely punctuated by fissures and conduits infilled by loose sands and gravels. These are inferred to be emplaced by subglacial meltwater during the final stages of ice sheet advance. Covering both LA2 and LA2b, LA3 is a unit of glaciofluvial outwash, composed

  15. Influence of spider silk on refugia preferences of the recluse spiders Loxosceles reclusa and Loxosceles laeta (Araneae: Sicariidae).

    PubMed

    Vetter, Richard S; Rust, Michael K

    2010-06-01

    In a previous experimental study, recluse spiders Loxosceles reclusa Gertsch and Mulaik and Loxosceles laeta (Nicolet) (Araneae: Sicariidae) preferred small cardboard refugia covered with conspecific silk compared with never-occupied refugia. Herein, we investigated some factors that might be responsible for this preference using similar cardboard refugia. When the two Loxosceles species were given choices between refugia previously occupied by their own and by the congeneric species, neither showed a species-specific preference; however, each chose refugia coated with conspecific silk rather than those previously inhabited by a distantly related cribellate spider, Metaltella simoni (Keyserling). When L. laeta spiders were offered refugia that were freshly removed from silk donors compared with heated, aged refugia from the same silk donor, older refugia were preferred. Solvent extracts of L. laeta silk were chosen approximately as often as control refugia when a range of solvents (methylene chloride:methanol, water, and hexane) were used. However, when acetone was used on similar silk, there was a statistical preference for the control, indicating that there might be a mildly repellent aspect to acetone-washed silk. Considering the inability to show attraction to chemical aspects of fresh silk, it seems that physical attributes may be more important for selection and that there might be repellency to silk of a recently vacated spider. These findings are discussed in regard to pest management strategies to control recluse spiders.

  16. Managing Climate Change Refugia for Climate Adaptation

    EPA Science Inventory

    The concept of refugia has long been studied from theoretical and paleontological perspectives to understand how populations persisted during past periods of unfavorable climate. Recently, researchers have applied the idea to contemporary landscapes to identify climate change ref...

  17. Post-glacial Paleo-oceanographic and Paleo-climatic Conditions and Linkages Along the West Coast of Canada

    NASA Astrophysics Data System (ADS)

    Dallimore, A.; Enkin, R. J.; McKechnie, I.

    2006-12-01

    Along the west coast of Canada, our continuing studies of annually laminated marine sediments in anoxic fjords illustrate the changing environment as glaciers retreated from this area about 12 ka y BP. New data from mid-coastal British Columbia expands our knowledge of the interplay between climate and ocean dynamics in the northeastern Pacific Ocean, and defines the evolution of modern climate conditions as ice receded from the coast, followed by the establishment of modern oceanographic and climatic conditions about 6,000 ky BP. The Late Pleistocene and Holocene record also marks dramatic changes in sea level, climate, coastal oceanographic dynamics and glacial sedimentary source and transport, with implications for the possibility of early human migration routes and glacial refugia. Changes in pre-historical aboriginal settlement sites and food sources also give indications of a dynamic Holocene land and seascape as modern conditions became established. Excellent chronological control is provided by complementary yet independent dating methods including radiocarbon dates on both plants and shells, identification of the Mazama Ash, varve counting and paleomagnetic/paleosecular variation correlations.

  18. Fine Dissection of Human Mitochondrial DNA Haplogroup HV Lineages Reveals Paleolithic Signatures from European Glacial Refugia

    PubMed Central

    Sarno, Stefania; Sevini, Federica; Vianello, Dario; Tamm, Erika; Metspalu, Ene; van Oven, Mannis; Hübner, Alexander; Sazzini, Marco; Franceschi, Claudio; Pettener, Davide; Luiselli, Donata

    2015-01-01

    Genetic signatures from the Paleolithic inhabitants of Eurasia can be traced from the early divergent mitochondrial DNA lineages still present in contemporary human populations. Previous studies already suggested a pre-Neolithic diffusion of mitochondrial haplogroup HV*(xH,V) lineages, a relatively rare class of mtDNA types that includes parallel branches mainly distributed across Europe and West Asia with a certain degree of structure. Up till now, variation within haplogroup HV was addressed mainly by analyzing sequence data from the mtDNA control region, except for specific sub-branches, such as HV4 or the widely distributed haplogroups H and V. In this study, we present a revised HV topology based on full mtDNA genome data, and we include a comprehensive dataset consisting of 316 complete mtDNA sequences including 60 new samples from the Italian peninsula, a previously underrepresented geographic area. We highlight points of instability in the particular topology of this haplogroup, reconstructed with BEAST-generated trees and networks. We also confirm a major lineage expansion that probably followed the Late Glacial Maximum and preceded Neolithic population movements. We finally observe that Italy harbors a reservoir of mtDNA diversity, with deep-rooting HV lineages often related to sequences present in the Caucasus and the Middle East. The resulting hypothesis of a glacial refugium in Southern Italy has implications for the understanding of late Paleolithic population movements and is discussed within the archaeological cultural shifts occurred over the entire continent. PMID:26640946

  19. Science questions for implementing climate refugia for cold-water fish as an adaptation strateby

    EPA Science Inventory

    Managing climate refugia has been proposed as a potential adaptation strategy that may be useful for protecting the biotic integrity of watersheds under a changing climate. Paleo-ecological evidence suggests that refugia allowed species to persist through prior periods of climate...

  20. The role of glacial cycles in promoting genetic diversity in the Neotropics: the case of cloud forests during the Last Glacial Maximum

    PubMed Central

    Ramírez-Barahona, Santiago; Eguiarte, Luis E

    2013-01-01

    The increasing aridity during the Last Glacial Maximum (LGM) has been proposed as a major factor affecting Neotropical species. The character and intensity of this change, however, remains the subject of ongoing debate. This review proposes an approach to test contrasting paleoecological hypotheses by way of their expected demographic and genetic effects on Neotropical cloud forest species. We reviewed 48 paleoecological records encompassing the LGM in the Neotropics. The records show contrasting evidence regarding the changes in precipitation during this period. Some regions remained fairly moist and others had a significantly reduced precipitation. Many paleoecological records within the same region show apparently conflicting evidence on precipitation and forest stability. From these data, we propose and outline two demographic/genetic scenarios for cloud forests species based on opposite precipitation regimes: the dry refugia and the moist forests hypotheses. We searched for studies dealing with the population genetic structure of cloud forest and other montane taxa and compared their results with the proposed models. To date, the few available molecular studies show insufficient genetic evidence on the predominance of glacial aridity in the Neotropics. In order to disentangle the climatic history of the Neotropics, the present study calls for a general multi-disciplinary approach to conduct future phylogeographic studies. Given the contradictory paleoecological information, population genetic data on Neotropical cloud forest species should be used to explicitly test the genetic consequences of competing paleoecological models. PMID:23531632

  1. The changing seascape of Galway Bay, Western Ireland

    NASA Astrophysics Data System (ADS)

    Mc Cullagh, D.; Benetti, S.; Plets, R. M. K.; Edwards, R.

    2016-12-01

    During the late Quaternary significant environmental and relative sea-level variations have contributed to shaping present day coastlines. This is particularly evident along formerly glaciated continental margins. Strong evidence of these changes are recorded in Galway Bay, Western Ireland. This research uses a multidisciplinary approach. Seismic and multibeam data, sedimentological, micropaleontological, geochemical analysis and 15 radiocarbon dates of sediment cores from the bay provide post last glacial maximum (LGM) sea level and environmental reconstructions for the region. The acoustic stratigraphy of the bay includes 3 seismic units: the deepest unit represents the acoustic basement, composed of limestone and granite bedrock; the middle unit is composed of the oldest preserved sediments, deposited during and after the LGM, and interpreted to be glacial till. The uppermost unit represents deposition and reworking after glacial retreat. The erosive action of the ice sheet that extended off the Irish coast is thought to be responsible for the removal and reworking of all sediments older that the LGM. In the sediment cores, three main lithofacies were identified: 1) a sandy silt and clay facies, 2) a distinct shell hash interlayer and, 3) a fine silty sand facies. These 3 facies are found within the uppermost seismic unit, and initial radiocarbon dating of shells in 4 cores, constrain these sediments and the uppermost seismic unit to the Holocene. Preliminary qualitative analysis on foraminifera from several cores shows a general trend of progression from estuarine to open marine conditions, inferred from indicator species. This trend is supported by X-ray fluorescence (XRF) analysis which shows increased ratios of Cl/Fe in younger deposits. Constraining dates on sea level variations in the region will be added to the sea level database for Ireland and possibly used to adjust the existing relative sea level models. These are important for understating past sea

  2. Origin of British and Irish mammals: disparate post-glacial colonisation and species introductions

    NASA Astrophysics Data System (ADS)

    Montgomery, W. Ian; Provan, Jim; McCabe, A. Marshal; Yalden, Derek W.

    2014-08-01

    Global climate changes during the Quaternary reveal much about broader evolutionary effects of environmental change. Detailed regional studies reveal how evolutionary lineages and novel communities and ecosystems, emerge through glacial bottlenecks or from refugia. There have been significant advances in benthic imaging and dating, particularly with respect to the movements of the British (Scottish) and Irish ice sheets and associated changes in sea level during and after the Last Glacial Maximum (LGM). Ireland has been isolated as an island for approximately twice as long as Britain with no evidence of any substantial, enduring land bridge between these islands after ca 15 kya. Recent biogeographical studies show that Britain's mammal community is akin to those of southern parts of Scandinavia, The Netherlands and Belgium, but the much lower mammal species richness of Ireland is unique and needs explanation. Here, we consider physiographic, archaeological, phylogeographical i.e. molecular genetic, and biological evidence comprising ecological, behavioural and morphological data, to review how mammal species recolonized western Europe after the LGM with emphasis on Britain and, in particular, Ireland. We focus on why these close neighbours had such different mammal fauna in the early Holocene, the stability of ecosystems after LGM subject to climate change and later species introductions. There is general concordance of archaeological and molecular genetic evidence where data allow some insight into history after the LGM. Phylogeography reveals the process of recolonization, e.g. with respect to source of colonizers and anthropogenic influence, whilst archaeological data reveal timing more precisely through carbon dating and stratigraphy. More representative samples and improved calibration of the ‘molecular clock' will lead to further insights with regards to the influence of successive glaciations. Species showing greatest morphological, behavioural and

  3. Extinction and recolonization of maritime Antarctica in the limpet Nacella concinna (Strebel, 1908) during the last glacial cycle: toward a model of Quaternary biogeography in shallow Antarctic invertebrates.

    PubMed

    González-Wevar, C A; Saucède, T; Morley, S A; Chown, S L; Poulin, E

    2013-10-01

    Quaternary glaciations in Antarctica drastically modified geographical ranges and population sizes of marine benthic invertebrates and thus affected the amount and distribution of intraspecific genetic variation. Here, we present new genetic information in the Antarctic limpet Nacella concinna, a dominant Antarctic benthic species along shallow ice-free rocky ecosystems. We examined the patterns of genetic diversity and structure in this broadcast spawner along maritime Antarctica and from the peri-Antarctic island of South Georgia. Genetic analyses showed that N. concinna represents a single panmictic unit in maritime Antarctic. Low levels of genetic diversity characterized this population; its median-joining haplotype network revealed a typical star-like topology with a short genealogy and a dominant haplotype broadly distributed. As previously reported with nuclear markers, we detected significant genetic differentiation between South Georgia Island and maritime Antarctica populations. Higher levels of genetic diversity, a more expanded genealogy and the presence of more private haplotypes support the hypothesis of glacial persistence in this peri-Antarctic island. Bayesian Skyline plot and mismatch distribution analyses recognized an older demographic history in South Georgia. Approximate Bayesian computations did not support the persistence of N. concinna along maritime Antarctica during the last glacial period, but indicated the resilience of the species in peri-Antarctic refugia (South Georgia Island). We proposed a model of Quaternary Biogeography for Antarctic marine benthic invertebrates with shallow and narrow bathymetric ranges including (i) extinction of maritime Antarctic populations during glacial periods; (ii) persistence of populations in peri-Antarctic refugia; and (iii) recolonization of maritime Antarctica following the deglaciation process. © 2013 John Wiley & Sons Ltd.

  4. First evidence of a Late Upper Palaeolithic human presence in Ireland

    NASA Astrophysics Data System (ADS)

    Dowd, Marion; Carden, Ruth F.

    2016-05-01

    The colonisation of North West Europe by humans and fauna following the Last Glacial Maximum (LGM) has been the subject of considerable discussion in recent decades and within multiple disciplines. Here we present new evidence that pushes back the date of human footfall in Ireland by up to 2500 cal BP to the Upper Palaeolithic. An assemblage of animal bones recovered from a cave in the west of Ireland during antiquarian excavations in 1903 included a butchered brown bear bone (patella) which was recently subjected to two independent radiocarbon dating processes; the resultant dates were in agreement: 12,810-12,590 cal BP and 12,810-12,685 cal BP. This find rewrites the antiquity of human occupation of Ireland and challenges the traditional paradigm that certain biota may have naturally colonised the island prior to human arrival.

  5. Outcrop Geomorphology and Sediment Characterization in Killary Harbour, Ireland

    NASA Astrophysics Data System (ADS)

    Crouse, L. E.; Sautter, L.

    2016-02-01

    Killary Harbour in western Ireland is one of three fjards found in the country. A fjard is formed by glacial carving, but differs from fjords in being shallower, shorter, and broader in profile. The Harbour is located near the border of County Mayo and County Galway. The Marine Institute of Ireland, The Geological Survey of Ireland, and the INFOMAR project, led by chief scientist Kevin Sheehan, conducted bathymetric surveys in Killary Harbour, Ireland during July and August of 2014 aboard the R/V Celtic Voyager. A Kongsberg EM2040 multi-beam sonar was utilized to acquire the raw data, which was post-processed using CARIS HIPS and SIPS 9.0. The area of study is shallow harbor seafloor depths ranging from 15 to 60 m. An abundance of rocky outcrops and narrow channels are present in the North-Western section of the study area. Backscatter data collected during acquisition were used to determine the relative hardness of seafloor sediments and rocky outcrops. Characterization of the bathymetric terrain and sediments of the harbour could prove to be useful in finding hardbottom substrate for ideal fish habitat.

  6. Post-glacial sea-level history for SW Ireland (Bantry Bay) based on offshore evidence

    NASA Astrophysics Data System (ADS)

    Plets, R. M.; Callard, L.; Cooper, A.; Long, A. J.; Belknap, D. F.; Edwards, R.; Jackson, D.; Kelley, J. T.; Long, D.; Milne, G. A.; Monteys, X.; Quinn, R.

    2013-12-01

    In recent years, progress in remote sensing techniques has helped to constrain the advance and retreat phases of the British-Irish Ice Sheet during and after the Last Glacial Maximum (LGM), both on- and offshore. However, little evidence has been collected to study the pattern of relative sea-level (RSL) change immediately after ice sheet retreat. Glacio-isostatic adjustment (GIA) models suggest a complex RSL pattern around Ireland, influenced by local and regional isostatic movements. Unfortunately, such models are poorly constrained for periods during which RSL was significantly lower than present, particularly the Late Pleistocene and Early Holocene, owing to the paucity of accurate observational data offshore. This poster presents post-LGM stratigraphic evidence from Bantry Bay (SW Ireland), one of seven areas targeted around the Irish Sea as part of a larger NERC funded project which aims to provide the first field data on the depth and age of the RSL minimum since deglaciation in the Irish Sea Basin. Data examined consists of: multibeam bathymetry and backscatter, pinger sub-bottom and vibrocores (25 sites). Notable features on the multibeam are a bluff line in the outer bay with a maximum height of 10 m in water depths of c. -80 m which forms the western edge of a large sediment lobe. The south-western boundary of this lobe is marked by a series of long (up to 22 km), parallel ridges at depths between -96 m and -131 m, with iceberg scouring evident on the offshore margin. Six seismo-stratigraphic units are interpreted from the pinger data, the most prominent of which can be traced from the inner part of the Bay to the inshore edge of the ridges. This unit sits on an erosional surface, is characterised by a turbid acoustic signature and is identified as alternating sand and clay layers with some traces of organic material and gas. Equal amounts of marine and estuarine foraminifera are present within this unit, whilst the underlying unit has a higher

  7. Caribbean mesophotic coral ecosystems are unlikely climate change refugia.

    PubMed

    Smith, Tyler B; Gyory, Joanna; Brandt, Marilyn E; Miller, William J; Jossart, Jonathan; Nemeth, Richard S

    2016-08-01

    Deeper coral reefs experience reduced temperatures and light and are often shielded from localized anthropogenic stressors such as pollution and fishing. The deep reef refugia hypothesis posits that light-dependent stony coral species at deeper depths are buffered from thermal stress and will avoid bleaching-related mass mortalities caused by increasing sea surface temperatures under climate change. This hypothesis has not been tested because data collection on deeper coral reefs is difficult. Here we show that deeper (mesophotic) reefs, 30-75 m depth, in the Caribbean are not refugia because they have lower bleaching threshold temperatures than shallow reefs. Over two thermal stress events, mesophotic reef bleaching was driven by a bleaching threshold that declines 0.26 °C every +10 m depth. Thus, the main premise of the deep reef refugia hypothesis that cooler environments are protective is incorrect; any increase in temperatures above the local mean warmest conditions can lead to thermal stress and bleaching. Thus, relatively cooler temperatures can no longer be considered a de facto refugium for corals and it is likely that many deeper coral reefs are as vulnerable to climate change as shallow water reefs. © 2015 John Wiley & Sons Ltd.

  8. Phylogeography and Post-Glacial Recolonization in Wolverines (Gulo gulo) from across Their Circumpolar Distribution

    PubMed Central

    Zigouris, Joanna; Schaefer, James A.; Fortin, Clément; Kyle, Christopher J.

    2013-01-01

    Interglacial-glacial cycles of the Quaternary are widely recognized in shaping phylogeographic structure. Patterns from cold adapted species can be especially informative - in particular, uncovering additional glacial refugia, identifying likely recolonization patterns, and increasing our understanding of species’ responses to climate change. We investigated phylogenetic structure of the wolverine, a wide-ranging cold adapted carnivore, using a 318 bp of the mitochondrial DNA control region for 983 wolverines (n = 209 this study, n = 774 from GenBank) from across their full Holarctic distribution. Bayesian phylogenetic tree reconstruction and the distribution of observed pairwise haplotype differences (mismatch distribution) provided evidence of a single rapid population expansion across the wolverine’s Holarctic range. Even though molecular evidence corroborated a single refugium, significant subdivisions of population genetic structure (0.01< ΦST <0.99, P<0.05) were detected. Pairwise ΦST estimates separated Scandinavia from Russia and Mongolia, and identified five main divisions within North America - the Central Arctic, a western region, an eastern region consisting of Ontario and Quebec/Labrador, Manitoba, and California. These data are in contrast to the nearly panmictic structure observed in northwestern North America using nuclear microsatellites, but largely support the nuclear DNA separation of contemporary Manitoba and Ontario wolverines from northern populations. Historic samples (c. 1900) from the functionally extirpated eastern population of Quebec/Labrador displayed genetic similarities to contemporary Ontario wolverines. To understand these divergence patterns, four hypotheses were tested using Approximate Bayesian Computation (ABC). The most supported hypothesis was a single Beringia incursion during the last glacial maximum that established the northwestern population, followed by a west-to-east colonization during the Holocene. This

  9. Phylogeography and post-glacial recolonization in wolverines (Gulo gulo) from across their circumpolar distribution.

    PubMed

    Zigouris, Joanna; Schaefer, James A; Fortin, Clément; Kyle, Christopher J

    2013-01-01

    Interglacial-glacial cycles of the Quaternary are widely recognized in shaping phylogeographic structure. Patterns from cold adapted species can be especially informative - in particular, uncovering additional glacial refugia, identifying likely recolonization patterns, and increasing our understanding of species' responses to climate change. We investigated phylogenetic structure of the wolverine, a wide-ranging cold adapted carnivore, using a 318 bp of the mitochondrial DNA control region for 983 wolverines (n=209 this study, n=774 from GenBank) from across their full Holarctic distribution. Bayesian phylogenetic tree reconstruction and the distribution of observed pairwise haplotype differences (mismatch distribution) provided evidence of a single rapid population expansion across the wolverine's Holarctic range. Even though molecular evidence corroborated a single refugium, significant subdivisions of population genetic structure (0.01< ΦST <0.99, P<0.05) were detected. Pairwise ΦST estimates separated Scandinavia from Russia and Mongolia, and identified five main divisions within North America - the Central Arctic, a western region, an eastern region consisting of Ontario and Quebec/Labrador, Manitoba, and California. These data are in contrast to the nearly panmictic structure observed in northwestern North America using nuclear microsatellites, but largely support the nuclear DNA separation of contemporary Manitoba and Ontario wolverines from northern populations. Historic samples (c. 1900) from the functionally extirpated eastern population of Quebec/Labrador displayed genetic similarities to contemporary Ontario wolverines. To understand these divergence patterns, four hypotheses were tested using Approximate Bayesian Computation (ABC). The most supported hypothesis was a single Beringia incursion during the last glacial maximum that established the northwestern population, followed by a west-to-east colonization during the Holocene. This pattern is

  10. Large Scale Anthropogenic Reduction of Forest Cover in Last Glacial Maximum Europe

    PubMed Central

    Pfeiffer, Mirjam; Kolen, Jan C. A.; Davis, Basil A. S.

    2016-01-01

    Reconstructions of the vegetation of Europe during the Last Glacial Maximum (LGM) are an enigma. Pollen-based analyses have suggested that Europe was largely covered by steppe and tundra, and forests persisted only in small refugia. Climate-vegetation model simulations on the other hand have consistently suggested that broad areas of Europe would have been suitable for forest, even in the depths of the last glaciation. Here we reconcile models with data by demonstrating that the highly mobile groups of hunter-gatherers that inhabited Europe at the LGM could have substantially reduced forest cover through the ignition of wildfires. Similar to hunter-gatherers of the more recent past, Upper Paleolithic humans were masters of the use of fire, and preferred inhabiting semi-open landscapes to facilitate foraging, hunting and travel. Incorporating human agency into a dynamic vegetation-fire model and simulating forest cover shows that even small increases in wildfire frequency over natural background levels resulted in large changes in the forested area of Europe, in part because trees were already stressed by low atmospheric CO2 concentrations and the cold, dry, and highly variable climate. Our results suggest that the impact of humans on the glacial landscape of Europe may be one of the earliest large-scale anthropogenic modifications of the earth system. PMID:27902716

  11. Fire and ice: volcanic and glacial impacts on the phylogeography of the New Zealand forest fern Asplenium hookerianum.

    PubMed

    Shepherd, Lara D; Perrie, Leon R; Brownsey, Patrick J

    2007-11-01

    In the Southern Hemisphere there has been little phylogeographical investigation of forest refugia sites during the last glacial. Hooker's spleenwort, Asplenium hookerianum, is a fern that is found throughout New Zealand. It is strongly associated with forest and is a proxy for the survival of woody vegetation during the last glacial maximum. DNA sequence data from the chloroplast trnL-trnF locus were obtained from 242 samples, including c. 10 individuals from each of 21 focal populations. Most populations contained multiple, and in many cases unique, haplotypes, including those neighbouring formerly glaciated areas, while the predominant inference from nested clade analysis was restricted gene flow with isolation by distance. These results suggest that A. hookerianum survived the last glacial maximum in widespread populations of sufficient size to retain the observed phylogeography, and therefore that the sheltering woody vegetation must have been similarly abundant. This is consistent with palynological interpretations for the survival in New Zealand of thermophilous forest species at considerably smaller distances from the ice sheets than recorded for the Northern Hemisphere. Eastern and central North Island populations of A. hookerianum were characterized by a different subset of haplotypes to populations from the remainder of the country. A similar east-west phylogeographical pattern has been detected in a diverse array of taxa, and has previously been attributed to recurrent vulcanism in the central North Island.

  12. Managing Climate Change Refugia for Biodiversity ...

    EPA Pesticide Factsheets

    Climate change threatens to create fundamental shifts in in the distributions and abundances of species. Given projected losses, increased emphasis on management for ecosystem resilience to help buffer fish and wildlife populations against climate change is emerging. Such efforts stake a claim for an adaptive, anticipatory planning response to the climate change threat. To be effective, approaches will need to address critical uncertainties in both the physical basis for projected landscape changes, as well as the biological responses of organisms. Recent efforts define future potential climate refugia based on air temperatures and associated microclimatic changes. These efforts reflect the relatively strong conceptual foundation for linkages between regional climate change and local responses and thermal dynamics. Yet important questions remain. Drawing on case studies, we illustrate some key uncertainties in the responses of species and their habitats to altered hydro-climatic regimes currently not well addressed by physical or ecological models. These uncertainties need not delay anticipatory planning, but rather highlight the need for identification and communication of actions with high probabilities of success, and targeted research within an adaptive management framework.In this workshop, we will showcase the latest science on climate refugia and participants will interact through small group discussions, relevant examples, and facilitated dialogue to i

  13. Paleoclimatic implications of glacial and postglacial refugia for Pinus pumila in western Beringia

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Anderson, P M; Lozhkin, A V; Solomatkina, T B

    Palynological results from Julietta Lake currently provide the most direct evidence to support the existence of a glacial refugium for Pinus pumila in mountains of southwestern Beringia. Both percentages and accumulation rates indicate the evergreen shrub survived until at least {approx}19,000 14C yr B.P. in the Upper Kolyma region. Percentage data suggest numbers dwindled into the late glaciation, whereas pollen accumulation rates point towards a more rapid demise shortly after {approx}19,000 14C yr B.P. Pinus pumila did not re-establish in any great numbers until {approx}8100 14C yr B.P., despite the local presence {approx}9800 14C yr B.P. of Larix dahurica, whichmore » shares similar summer temperature requirements. The postglacial thermal maximum (in Beringia {approx}11,000-9000 14C yr B.P.) provided Pinus pumila shrubs with equally harsh albeit different conditions for survival than those present during the LGM. Regional records indicate that in this time of maximum warmth Pinus pumila likely sheltered in a second, lower-elevation refugium. Paleoclimatic models and modern ecology suggest that shifts in the nature of seasonal transitions and not only seasonal extremes have played important roles in the history of Pinus pumila over the last {approx}21,000 14C yr B.P.« less

  14. Future distribution of tundra refugia in northern Alaska

    USGS Publications Warehouse

    Hope, Andrew G.; Waltari, Eric; Payer, David C.; Cook, Joseph A.; Talbot, Sandra L.

    2013-01-01

    Climate change in the Arctic is a growing concern for natural resource conservation and management as a result of accelerated warming and associated shifts in the distribution and abundance of northern species. We introduce a predictive framework for assessing the future extent of Arctic tundra and boreal biomes in northern Alaska. We use geo-referenced museum specimens to predict the velocity of distributional change into the next century and compare predicted tundra refugial areas with current land-use. The reliability of predicted distributions, including differences between fundamental and realized niches, for two groups of species is strengthened by fossils and genetic signatures of demographic shifts. Evolutionary responses to environmental change through the late Quaternary are generally consistent with past distribution models. Predicted future refugia overlap managed areas and indicate potential hotspots for tundra diversity. To effectively assess future refugia, variable responses among closely related species to climate change warrants careful consideration of both evolutionary and ecological histories.

  15. Dynamics, rate and nature of retreat of the British Irish Ice-Sheet offshore of NW Ireland following the Last Glacial Maximum

    NASA Astrophysics Data System (ADS)

    Weilbach, K.; O'Cofaigh, C.; Lloyd, J. M.; Benetti, S.; Dunlop, P.

    2016-12-01

    Recent studies of the British and Irish Ice Sheet (BIIS) have identified evidence of ice extending to the continental shelf edge along the western margin of the ice sheet off NW Ireland. While this advance is assumed to have occurred during the LGM, exact timing of maximum advance, and the timing and nature of the subsequent retreat is not well constrained. The location of the north-western sector of the BIIS adjacent to the North Atlantic makes this area ideal to study the ice sheet dynamics of a major marine terminating ice sheet, and the rate and nature of its retreat following the LGM. High resolution swath bathymetry and sub-bottom profiler (SBP) data along with sedimentological, micropalaeontological and geochronological investigations of sediment cores, collected across the NW Irish shelf, have been used to establish the extent, timing and nature of retreat of this sector of the BIIS. Swath bathymetry show glacial landforms on the shelf, and SBP-data along with twenty seven vibro-cores were collected in east-west oriented transects across a series of arcuate recessional moraines stretching from the shelf edge to Donegal Bay. These moraines record progressive still stands of a lobate ice margin during its retreat from the shelf edge, and are therefore ideal for the investigation of ice-sheet dynamics and chronology during retreat. Twenty two radiocarbon dates from foraminifera and macrofossils, sampled from the sediment cores, indicate that maximum ice sheet extent occurred around 26200 y cal BP, with an initial rapid retreat across the shelf. Visual logging, X-ray imagery, MSCL data and palaeoenvironmental analyses of the sediment cores, indicate that retreat happened in a glacimarine environment, and was punctuated by multiple stillstands and possible readvances across the mid and inner shelf, forming the arcuate moraines. The radiocarbon dates suggest that final retreat occurred after 17857 y. cal BP, which is consistent with onshore cosmogenic exposure

  16. Climate change refugia as a tool for climate adaptation

    EPA Science Inventory

    Climate change refugia, areas relatively buffered from contemporary climate change so as to increase persistence of valued physical, ecological, and cultural resources, are considered as potential adaptation options in the face of anthropogenic climate change. In a collaboration ...

  17. Temperature inverted haloclines provide winter warm-water refugia for manatees in southwest Florida

    USGS Publications Warehouse

    Stith, Bradley M.; Reid, James P.; Langtimm, Catherine A.; Swain, Eric D.; Doyle, Terry J.; Slone, Daniel H.; Decker, Jeremy D.; Soderqvist, Lars E.

    2010-01-01

    Florida manatees (Trichechus manatus latirostris) overwintering in the Ten Thousand Islands and western Everglades have no access to power plants or major artesian springs that provide warm-water refugia in other parts of Florida. Instead, hundreds of manatees aggregate at artificial canals, basins, and natural deep water sites that act as passive thermal refugia (PTR). Monitoring at two canal sites revealed temperature inverted haloclines, which provided warm salty bottom layers that generally remained above temperatures considered adverse for manatees. At the largest PTR, the warmer bottom layer disappeared unless significant salt stratification was maintained by upstream freshwater inflow over a persistent tidal wedge. A detailed three-dimensional hydrology model showed that salinity stratification inhibited vertical convection induced by atmospheric cooling. Management or creation of temperature inverted haloclines may be a feasible and desirable option for resource managers to provide passive thermal refugia for manatees and other temperature sensitive aquatic species.

  18. Climate extremes and predicted warming threaten Mediterranean Holocene firs forests refugia

    PubMed Central

    Camarero, J. Julio; Carrer, Marco; Gutiérrez, Emilia; Alla, Arben Q.; Andreu-Hayles, Laia; Hevia, Andrea; Koutavas, Athanasios; Martínez-Sancho, Elisabet; Nola, Paola; Papadopoulos, Andreas; Pasho, Edmond; Toromani, Ervin

    2017-01-01

    Warmer and drier climatic conditions are projected for the 21st century; however, the role played by extreme climatic events on forest vulnerability is still little understood. For example, more severe droughts and heat waves could threaten quaternary relict tree refugia such as Circum-Mediterranean fir forests (CMFF). Using tree-ring data and a process-based model, we characterized the major climate constraints of recent (1950–2010) CMFF growth to project their vulnerability to 21st-century climate. Simulations predict a 30% growth reduction in some fir species with the 2050s business-as-usual emission scenario, whereas growth would increase in moist refugia due to a longer and warmer growing season. Fir populations currently subjected to warm and dry conditions will be the most vulnerable in the late 21st century when climatic conditions will be analogous to the most severe dry/heat spells causing dieback in the late 20th century. Quantification of growth trends based on climate scenarios could allow defining vulnerability thresholds in tree populations. The presented predictions call for conservation strategies to safeguard relict tree populations and anticipate how many refugia could be threatened by 21st-century dry spells. PMID:29109266

  19. Climate extremes and predicted warming threaten Mediterranean Holocene firs forests refugia.

    PubMed

    Sánchez-Salguero, Raúl; Camarero, J Julio; Carrer, Marco; Gutiérrez, Emilia; Alla, Arben Q; Andreu-Hayles, Laia; Hevia, Andrea; Koutavas, Athanasios; Martínez-Sancho, Elisabet; Nola, Paola; Papadopoulos, Andreas; Pasho, Edmond; Toromani, Ervin; Carreira, José A; Linares, Juan C

    2017-11-21

    Warmer and drier climatic conditions are projected for the 21st century; however, the role played by extreme climatic events on forest vulnerability is still little understood. For example, more severe droughts and heat waves could threaten quaternary relict tree refugia such as Circum-Mediterranean fir forests (CMFF). Using tree-ring data and a process-based model, we characterized the major climate constraints of recent (1950-2010) CMFF growth to project their vulnerability to 21st-century climate. Simulations predict a 30% growth reduction in some fir species with the 2050s business-as-usual emission scenario, whereas growth would increase in moist refugia due to a longer and warmer growing season. Fir populations currently subjected to warm and dry conditions will be the most vulnerable in the late 21st century when climatic conditions will be analogous to the most severe dry/heat spells causing dieback in the late 20th century. Quantification of growth trends based on climate scenarios could allow defining vulnerability thresholds in tree populations. The presented predictions call for conservation strategies to safeguard relict tree populations and anticipate how many refugia could be threatened by 21st-century dry spells.

  20. Evolutionary refugia and ecological refuges: key concepts for conserving Australian arid zone freshwater biodiversity under climate change

    PubMed Central

    Davis, Jenny; Pavlova, Alexandra; Thompson, Ross; Sunnucks, Paul

    2013-01-01

    Refugia have been suggested as priority sites for conservation under climate change because of their ability to facilitate survival of biota under adverse conditions. Here, we review the likely role of refugial habitats in conserving freshwater biota in arid Australian aquatic systems where the major long-term climatic influence has been aridification. We introduce a conceptual model that characterizes evolutionary refugia and ecological refuges based on our review of the attributes of aquatic habitats and freshwater taxa (fishes and aquatic invertebrates) in arid Australia. We also identify methods of recognizing likely future refugia and approaches to assessing the vulnerability of arid-adapted freshwater biota to a warming and drying climate. Evolutionary refugia in arid areas are characterized as permanent, groundwater-dependent habitats (subterranean aquifers and springs) supporting vicariant relicts and short-range endemics. Ecological refuges can vary across space and time, depending on the dispersal abilities of aquatic taxa and the geographical proximity and hydrological connectivity of aquatic habitats. The most important are the perennial waterbodies (both groundwater and surface water fed) that support obligate aquatic organisms. These species will persist where suitable habitats are available and dispersal pathways are maintained. For very mobile species (invertebrates with an aerial dispersal phase) evolutionary refugia may also act as ecological refuges. Evolutionary refugia are likely future refugia because their water source (groundwater) is decoupled from local precipitation. However, their biota is extremely vulnerable to changes in local conditions because population extinction risks cannot be abated by the dispersal of individuals from other sites. Conservation planning must incorporate a high level of protection for aquifers that support refugial sites. Ecological refuges are vulnerable to changes in regional climate because they have little

  1. Evolutionary refugia and ecological refuges: key concepts for conserving Australian arid zone freshwater biodiversity under climate change.

    PubMed

    Davis, Jenny; Pavlova, Alexandra; Thompson, Ross; Sunnucks, Paul

    2013-07-01

    Refugia have been suggested as priority sites for conservation under climate change because of their ability to facilitate survival of biota under adverse conditions. Here, we review the likely role of refugial habitats in conserving freshwater biota in arid Australian aquatic systems where the major long-term climatic influence has been aridification. We introduce a conceptual model that characterizes evolutionary refugia and ecological refugees based on our review of the attributes of aquatic habitats and freshwater taxa (fishes and aquatic invertebrates) in arid Australia. We also identify methods of recognizing likely future refugia and approaches to assessing the vulnerability of arid-adapted freshwater biota to a warming and drying climate. Evolutionary refugia in arid areas are characterized as permanent, groundwater-dependent habitats (subterranean aquifers and springs) supporting vicariant relicts and short-range endemics. Ecological refugees can vary across space and time, depending on the dispersal abilities of aquatic taxa and the geographical proximity and hydrological connectivity of aquatic habitats. The most important are the perennial waterbodies (both groundwater and surface water fed) that support obligate aquatic organisms. These species will persist where suitable habitats are available and dispersal pathways are maintained. For very mobile species (invertebrates with an aerial dispersal phase) evolutionary refugia may also act as ecological refugees. Evolutionary refugia are likely future refugia because their water source (groundwater) is decoupled from local precipitation. However, their biota is extremely vulnerable to changes in local conditions because population extinction risks cannot be abated by the dispersal of individuals from other sites. Conservation planning must incorporate a high level of protection for aquifers that support refugial sites. Ecological refuges are vulnerable to changes in regional climate because they have

  2. Post-glacial dispersal patterns of Northern pike inferred from an 8800 year old pike (Esox cf. lucius) skull from interior Alaska

    NASA Astrophysics Data System (ADS)

    Wooller, Matthew J.; Gaglioti, Benjamin; Fulton, Tara L.; Lopez, Andres; Shapiro, Beth

    2015-07-01

    The biogeography of freshwater fish species during and after late-Pleistocene glaciations relate to how these species are genetically organized today, and the management of these often disjunct populations. Debate exists concerning the biogeography and routes of dispersal for Northern pike (Esox lucius) after the last glaciation. A hypothesis to account for the relatively low modern genetic diversity for E. lucius is post-glacial radiation from refugia, including lakes from within the un-glaciated portions of eastern Beringia. We report the remains of a Northern pike (E. cf. lucius) skull, including bones, teeth, bone collagen and ancient DNA. The remains were preserved at a depth of between 440 and 446 cm in a 670 cm long core of sediment from Quartz Lake, which initiated at ˜11,200 cal yr BP in interior Alaska. A calibrated accelerator mass spectrometer (AMS) radiocarbon age of the collagen extracted from the preserved bones indicated that the organism was dated to 8820 cal yr BP and is bracketed by AMS values from analyses of terrestrial plant macrofossils, avoiding any potential aquatic reservoir effect that could have influenced the radiocarbon age of the bones. Scanning electron microscope images of the specimen show the hinged tooth anatomy typically of E. lucius. Molar C:N (3.5, 1σ = 0.1) value of the collagen from the specimen indicated well-preserved collagen and its mean stable nitrogen isotope value is consistent with the known predatory feeding ecology of E. lucius. Ancient DNA in the bones showed that the specimen was identical to modern E. lucius. Our record of E. lucius from interior Alaska is consistent with a biogeographic scenario involving rapid dispersal of this species from glacial refugia in the northern hemisphere after the last glaciation.

  3. Comment on "Last glacial maximum cirque glaciation in Ireland and implications for reconstructions of the Irish ice sheet. Quaternary Science Reviews 141, 85-93"

    NASA Astrophysics Data System (ADS)

    Knight, Jasper

    2016-10-01

    Southwest Ireland is a critical location to examine the sensitivity of late Pleistocene glaciers to climate variability in the northeast Atlantic, because of its proximal location to Atlantic moisture sources and the presence of high mountains in the Macgillycuddy's Reeks range which acted as a focus for glacierization (Harrison et al., 2010). The extent of Last Glacial Maximum (LGM) glaciers in southwest Ireland and their link to the wider British-Irish Ice Sheet (BIIS), however, is under debate. Some models suggest that during the LGM the region was wholly inundated by ice from the larger BIIS (Warren, 1992; Sejrup et al., 2005), whereas others suggest north-flowing ice from the semi-independent Cork-Kerry Ice Cap (CKIC) was diverted around mountain peaks, resulting in exposed nunataks in the Macgillycuddy's Reeks (Anderson et al., 2001; Ballantyne et al., 2011). Cirque glaciers may also have been present on mountain slopes above this regional ice surface (Warren, 1979; Rea et al., 2004). More recently, investigations have focused on the extent and age of cirque glaciers in the Reeks, based on the mapped distribution of end moraines (Warren, 1979; Harrison et al., 2010), and on cosmogenic dates on boulders on these moraines (Harrison et al., 2010) and on associated scoured bedrock surfaces across the region (Ballantyne et al., 2011). The recent paper by Barth et al. (2016) contributes to this debate by providing nine cosmogenic 10Be ages on boulders from two moraines from one small (∼1.7 km2) and low (373 m elevation of the cirque floor) cirque basin at Alohart (52°00‧50″N, 9°40‧30″W) within the Reeks range. These dates are welcomed because they add to the lengthening list of age constraints on geomorphic activity in the region that spans the time period from the LGM to early Holocene.

  4. Brook trout use of thermal refugia and foraging habitat influenced by brown trout

    USGS Publications Warehouse

    Hitt, Nathaniel P.; Snook, Erin; Massie, Danielle L.

    2017-01-01

    The distribution of native brook trout (Salvelinus fontinalis) in eastern North America is often limited by temperature and introduced brown trout (Salmo trutta), the relative importance of which is poorly understood but critical for conservation and restoration planning. We evaluated effects of brown trout on brook trout behavior and habitat use in experimental streams across increasing temperatures (14–23 °C) with simulated groundwater upwelling zones providing thermal refugia (6–9 °C below ambient temperatures). Allopatric and sympatric trout populations increased their use of upwelling zones as ambient temperatures increased, demonstrating the importance of groundwater as thermal refugia in warming streams. Allopatric brook trout showed greater movement rates and more even spatial distributions within streams than sympatric brook trout, suggesting interference competition by brown trout for access to forage habitats located outside thermal refugia. Our results indicate that removal of introduced brown trout may facilitate native brook trout expansion and population viability in downstream reaches depending in part on the spatial configuration of groundwater upwelling zones.

  5. Hydrologic refugia, plants, and climate change.

    PubMed

    McLaughlin, Blair C; Ackerly, David D; Klos, P Zion; Natali, Jennifer; Dawson, Todd E; Thompson, Sally E

    2017-08-01

    Climate, physical landscapes, and biota interact to generate heterogeneous hydrologic conditions in space and over time, which are reflected in spatial patterns of species distributions. As these species distributions respond to rapid climate change, microrefugia may support local species persistence in the face of deteriorating climatic suitability. Recent focus on temperature as a determinant of microrefugia insufficiently accounts for the importance of hydrologic processes and changing water availability with changing climate. Where water scarcity is a major limitation now or under future climates, hydrologic microrefugia are likely to prove essential for species persistence, particularly for sessile species and plants. Zones of high relative water availability - mesic microenvironments - are generated by a wide array of hydrologic processes, and may be loosely coupled to climatic processes and therefore buffered from climate change. Here, we review the mechanisms that generate mesic microenvironments and their likely robustness in the face of climate change. We argue that mesic microenvironments will act as species-specific refugia only if the nature and space/time variability in water availability are compatible with the ecological requirements of a target species. We illustrate this argument with case studies drawn from California oak woodland ecosystems. We posit that identification of hydrologic refugia could form a cornerstone of climate-cognizant conservation strategies, but that this would require improved understanding of climate change effects on key hydrologic processes, including frequently cryptic processes such as groundwater flow. © 2017 John Wiley & Sons Ltd.

  6. Topographic and fire weather controls of fire refugia in forested ecosystems of northwestern North America

    USGS Publications Warehouse

    Krawchuk, Meg A.; Haire, Sandra L.; Coop, Jonathan D.; Parisien, Marc-Andre; Whitman, Ellen; Chong, Geneva W.; Miller, Carol

    2016-01-01

    for seven study fires that burned in conifer-dominated forested landscapes of the Western Cordillera of Canada between 2001 and 2014. We fit nine models, each for distinct levels of fire weather and terrain ruggedness. Our framework revealed that the predictability and abundance of fire refugia varied among these environmental settings. We observed highest predictability under moderate fire weather conditions and moderate terrain ruggedness (ROC-AUC = 0.77), and lowest predictability in flatter landscapes and under high fire weather conditions (ROC-AUC = 0.63–0.68). Catchment slope, local aspect, relative position, topographic wetness, topographic convergence, and local slope all contributed to discriminating where refugia occur but the relative importance of these topographic controls differed among environments. Our framework allows us to characterize the predictability of contemporary fire refugia across multiple environmental settings and provides important insights for ecosystem resilience, wildfire management, conservation planning, and climate change adaptation.

  7. Palaeoclimate change drove diversification among isolated mountain refugia in the Australian arid zone.

    PubMed

    Pepper, Mitzy; Fujita, Matthew K; Moritz, Craig; Keogh, J Scott

    2011-04-01

    Refugia featured prominently in shaping evolutionary trajectories during repeated cycles of glaciation in the Quaternary, particularly in the Northern Hemisphere. The Southern Hemisphere instead experienced cycles of severe aridification but little is known about the temporal presence and role of refugia for arid-adapted biota. Isolated mountain ranges located in the Australian arid zone likely provided refugia for many species following Mio/Pliocene (<15 Ma) aridification; however, the evolutionary consequences of the recent development of widespread sand deserts is largely unknown. To test alternative hypotheses of ancient vs. recent isolation, we generated a 10 gene data set to assess divergence history among saxicolous geckos in the genus Heteronotia that have distributions confined to major rocky ranges in the arid zone. Phylogenetic analyses show that each rocky range harbours a divergent lineage, and substantial intraspecific diversity is likely due to topographic complexity in these areas. Old divergences (~4 Ma) among lineages pre-date the formation of the geologically young sand deserts (<1 Ma), suggesting that Pliocene climate shifts fractured the distributions of biota long before the spread of the deserts. © 2011 Blackwell Publishing Ltd.

  8. Explosive ice age diversification of kiwi.

    PubMed

    Weir, Jason T; Haddrath, Oliver; Robertson, Hugh A; Colbourne, Rogan M; Baker, Allan J

    2016-09-20

    Molecular dating largely overturned the paradigm that global cooling during recent Pleistocene glacial cycles resulted in a burst of species diversification although some evidence exists that speciation was commonly promoted in habitats near the expanding and retracting ice sheets. Here, we used a genome-wide dataset of more than half a million base pairs of DNA to test for a glacially induced burst of diversification in kiwi, an avian family distributed within several hundred kilometers of the expanding and retracting glaciers of the Southern Alps of New Zealand. By sampling across the geographic range of the five kiwi species, we discovered many cryptic lineages, bringing the total number of kiwi taxa that currently exist to 11 and the number that existed just before human arrival to 16 or 17. We found that 80% of kiwi diversification events date to the major glacial advances of the Middle and Late Pleistocene. During this period, New Zealand was repeatedly fragmented by glaciers into a series of refugia, with the tiny geographic ranges of many kiwi lineages currently distributed in areas adjacent to these refugia. Estimates of effective population size through time show a dramatic bottleneck during the last glacial cycle in all but one kiwi lineage, as expected if kiwi were isolated in glacially induced refugia. Our results support a fivefold increase in diversification rates during key glacial periods, comparable with levels observed in classic adaptive radiations, and confirm that at least some lineages distributed near glaciated regions underwent rapid ice age diversification.

  9. Low Elevation Riparian Environments: Warm-Climate Refugia for Conifers in the Great Basin, USA?

    NASA Astrophysics Data System (ADS)

    Millar, C.; Charlet, D. A.; Westfall, R. D.; Delany, D.

    2015-12-01

    The Great Basin, USA, contains hundreds of small to large mountain ranges. Many reach alpine elevations, which are separated from each other by low-elevation basins currently inhospitable to conifer growth. Many of these ranges support montane and subalpine conifer species that have affinities to the Sierra Nevada or Rocky Mountains, and from which these conifers migrated during cool periods of the Pleistocene. Under Holocene climates, the Great Basin geography became a terrestrial island-archipelago, wherein conifer populations are isolated among ranges, and inter-range migration is highly limited. During warm intervals of the Holocene, conifers would be expected to have migrated upslope following favorable conditions, and extirpation would be assumed to result from continued warming. Independent patterns, repeating across multiple species' distributions, however, suggest that refugia were present in these ranges during warm periods, and that low elevation environments below the current main distributions acted as climatic refugia. We hypothesize that cool, narrow, and north-aspect ravines, which during cool climates support persistent or seasonal creeks and deciduous riparian communities, become available as conifer habitat when warming climates desiccate creeks and deplete riparian species. We further speculate that cold-air drainage, reduced solar insolation, lower wind exposure, and higher water tables in these topographic positions support populations of montane and subalpine conifers even during warm climate intervals when high elevations are unfavorable for conifer persistence. On return to cool climates, low elevation refugia become sources for recolonizing higher slopes, and/or continue to persist as relictual populations. We present several lines of evidence supporting this hypothesis, and speculate that low-elevation, extramarginal riparian environments might act as climate refugia for Great Basin conifers in the future as well.

  10. Phylogeography of a Tertiary relict plant, Meconopsis cambrica (Papaveraceae), implies the existence of northern refugia for a temperate herb.

    PubMed

    Valtueña, Francisco J; Preston, Chris D; Kadereit, Joachim W

    2012-03-01

    The perennial herb Meconopsis cambrica, a western European endemic, is the only European species of the otherwise Himalayan genus Meconopsis and has been interpreted as a Tertiary relict species. Using rbcL and ITS sequence variation, we date the split between M. cambrica and its sister clade Papaver s.str. to the Middle to Upper Miocene (12.8 Myr, 6.4-19.2 Myr HPD). Within M. cambrica, cpDNA sequence variation reveals the existence of two groups of populations with a comparable level of genetic variation: a northern group from Great Britain, the Massif Central, the western Pyrenees and the Iberian System, and a southern group from the central and eastern Pyrenees. Populations from the Cantabrian Mountains were placed in both groups. Based on ITS sequence variation, the divergence between these two groups can be dated to 1.5 Myr (0.4-2.8 Myr HPD), and the age of the British populations is estimated as 0.37 Myr (0.0-0.9 Myr HPD). Amplified fragment length polymorphism results confirm the distinctive nature of the populations from Britain, the Massif Central and the central and eastern Pyrenees. These patterns of latitudinal variation of M. cambrica differ from patterns of longitudinal differentiation found in many other temperate species and imply glacial survival of the northern populations in northerly refugia. The primary differentiation into northern and southern cpDNA groups dates to near the onset of the Quaternary and suggests that an ancient phylogeographic pattern has survived through several glacial periods. Our data provide evidence that the species has persisted for a long period with a highly fragmented and probably very localized distribution. © 2012 Blackwell Publishing Ltd.

  11. Pleistocene refugia and genetic diversity patterns in West Africa: Insights from the liana Chasmanthera dependens (Menispermaceae)

    PubMed Central

    Schmidt, Marco; Muellner-Riehl, Alexandra Nora; Ogundipe, Oluwatoyin Temitayo; Paule, Juraj

    2017-01-01

    Processes shaping the African Guineo-Congolian rain forest, especially in the West African part, are not well understood. Recent molecular studies, based mainly on forest tree species, confirmed the previously proposed division of the western African Guineo-Congolian rain forest into Upper Guinea (UG) and Lower Guinea (LG) separated by the Dahomey Gap (DG). Here we studied nine populations in the area of the DG and the borders of LG and UG of the widespread liana species, Chasmanthera dependens (Menispermaceae) by amplified fragment length polymorphism (AFLP), a chloroplast DNA sequence marker, and modelled the distribution based on current as well as paleoclimatic data (Holocene Climate Optimum, ca. 6 kyr BP and Last Glacial Maximum, ca. 22 kyr BP). Current population genetic structure and geographical pattern of cpDNA was related to present as well as historical modelled distributions. Results from this study show that past historical factors played an important role in shaping the distribution of C. dependens across West Africa. The Cameroon Volcanic Line seems to represent a barrier for gene flow in the present as well as in the past. Distribution modelling proposed refugia in the Dahomey Gap, supported also by higher genetic diversity. This is in contrast with the phylogeographic patterns observed in several rainforest tree species and could be explained by either diverging or more relaxed ecological requirements of this liana species. PMID:28301470

  12. Reply to comment received from J. C. Knight regarding "Last Glacial Maximum cirque glaciation in Ireland and implications for reconstructions of the Irish Ice Sheet" by Barth et al. (2016), Quaternary Science Reviews 141, 85-93

    NASA Astrophysics Data System (ADS)

    Barth, Aaron M.; Clark, Peter U.; Clark, Jorie; McCabe, A. Marshall; Caffee, Marc

    2016-10-01

    We concluded that our new 10Be chronology records onset of retreat of a cirque glacier within the Alohart basin of southwestern Ireland 24.5 ± 1.4 ka, placing limiting constraints on reconstructions of the Irish Ice Sheet (IIS) and Kerry-Cork Ice Cap (KCIC) during the Last Glacial Maximum (LGM) (Barth et al., 2016). Knight (2016) raises two main arguments against our interpretation: (1) the glacier in the Alohart basin was not a cirque glacier, but instead a southern-sourced ice tongue from the KCIC overtopping the MacGillycuddy's Reeks, and (2) that the boulders we sampled for 10Be exposure dating were derived from supraglacial rockfall rather than transported subglacially, experienced nuclide inheritance, and are thus too old. In the following, we address both of these arguments.

  13. Refugial persistence and postglacial recolonization of North America by the cold-tolerant herbaceous plant Orthilia secunda.

    PubMed

    Beatty, Gemma E; Provan, Jim

    2010-11-01

    Previous phylogeographical and palaeontological studies on the biota of northern North America have revealed a complex scenario of glacial survival in multiple refugia and differing patterns of postglacial recolonization. Many putative refugial regions have been proposed both north and south of the ice sheets for species during the Last Glacial Maximum, but the locations of many of these refugia remain a topic of great debate. In this study, we used a phylogeographical approach to elucidate the refugial and recolonization history of the herbaceous plant species Orthilia secunda in North America, which is found in disjunct areas in the west and east of the continent, most of which were either glaciated or lay close to the limits of the ice sheets. Analysis of 596 bp of the chloroplast trnS-trnG intergenic spacer and five microsatellite loci in 84 populations spanning the species' range in North America suggests that O. secunda persisted through the Last Glacial Maximum (LGM) in western refugia, even though palaeodistribution modelling indicated a suitable climate envelope across the entire south of the continent. The present distribution of the species has resulted from recolonization from refugia north and south of the ice sheets, most likely in Beringia or coastal regions of Alaska and British Columbia, the Washington/Oregon region in the northwest USA, and possibly from the region associated with the putative 'ice-free corridor' between the Laurentide and Cordilleran ice sheets. Our findings also highlight the importance of the Pacific Northwest as an important centre of intraspecific genetic diversity, owing to a combination of refugial persistence in the area and recolonization from other refugia. © 2010 Blackwell Publishing Ltd.

  14. Anthelmintic resistance: Management of parasite refugia for Haemonchus contortus through the replacement of resistant with susceptible populations.

    PubMed

    Muchiut, Sebastián Manuel; Fernández, Alicia Silvina; Steffan, Pedro Eduardo; Riva, Eliana; Fiel, César Alberto

    2018-04-30

    Sheep production in tropical and temperate regions is hampered by the presence of Haemonchus contortus, the blood-sucking nematode that is the major cause of economic losses in small ruminant enterprises. The most limiting factor in the control of this parasitic disease is the steady progress of anthelmintic resistance worldwide. The search for control strategies that minimise the use of anthelmintics is therefore central to various efforts worldwide. One strategy is the introduction of susceptible parasites in refugia when these refugia are at low levels. This strategy could lead to a renewed possibility anthelmintics being effective. At farm level, this management practice could recover the use of anthelmintics in flocks with high levels of resistance. This review explores the possibility of replacing resistant H. contortus populations with susceptible ones through refugia management and. highlights the experiences of on-farm research attempts carried out in different geographical areas, reaching various degrees of success. Copyright © 2018 Elsevier B.V. All rights reserved.

  15. The Pulley Ridge deep reef is not a stable refugia through time

    NASA Astrophysics Data System (ADS)

    Slattery, Marc; Moore, Steve; Boye, Lauren; Whitney, Samantha; Woolsey, Allison; Woolsey, Max

    2018-06-01

    The deep reef refugia hypothesis (DRRH) suggests that mesophotic coral ecosystems (MCEs) represent a sanctuary for various coral reef taxa from impacts of natural and anthropogenic stressors. The Pulley Ridge FL mesophotic reef was surveyed using unmanned vehicles and technical diving in 2015 and compared to vehicle surveys in 2003, to test the DRRH. The MCE sites surveyed consisted of at least 25 species of algae ( 50% cover), 18 species of scleractinian corals (< 1% cover), 67 species of sponges (1-2% cover), in addition to 83 species of fish. The percent cover data indicate a significant decline of coral and sponges, and a significant increase in algae, relative to surveys conducted about a decade earlier. While the cause of this change is unknown, our results indicate that some mesophotic reefs may not be stable refugia for coral reef biodiversity and seed banks for resilience of damaged shallow reefs.

  16. Reconstruction of full glacial environments and summer temperatures from Lago della Costa, a refugial site in Northern Italy

    NASA Astrophysics Data System (ADS)

    Samartin, Stéphanie; Heiri, Oliver; Kaltenrieder, Petra; Kühl, Norbert; Tinner, Willy

    2016-07-01

    Vegetation and climate during the last ice age and the Last Glacial Maximum (LGM, ∼23,000-19,000 cal BP) were considerably different than during the current interglacial (Holocene). Cold climatic conditions and growing ice-sheets during the last glaciation radically reduced forest extent in Europe to a restricted number of so-called ;refugia;, mostly located in the southern part of the continent. On the basis of paleobotanical analyses the Euganian Hills (Colli Euganei) in northeastern Italy have previously been proposed as one of the northernmost refugia of temperate trees (e.g. deciduous Quercus, Tilia, Ulmus, Fraxinus excelsior, Acer, Abies alba, Fagus sylvatica, Carpinus and Castanea) in Europe. In this study we provide the first quantitative, vegetation independent summer air temperature reconstruction for Northern Italy spanning the time ∼31,000-17,000 cal yr BP, which covers the coldest periods of the last glacial, including the LGM and Heinrich stadials 1 to 3. Chironomids preserved in a lake sediment core from Lago della Costa (7m a.s.l.), a small lake at the south-eastern edge of the Euganean Hills, allowed quantitative reconstruction of Full and Late Glacial summer air temperatures using a combined Swiss-Norwegian temperature inference model based on chironomid assemblages from 274 lakes. Chironomid and pollen evidence from Lago della Costa derives from finely stratified autochthonous organic gyttja sediments, which excludes major sediment mixing or reworking. After reconstructing paleo-temperatures, we address the question whether climate conditions were warm enough to permit the local survival of temperate tree species during the LGM and whether local expansions and pollen-inferred contractions of temperate tree taxa coincided with chironomid-inferred climatic changes. Our results suggest that chironomids at Lago della Costa have responded to major climatic fluctuations such as temperature decreases during the LGM and Heinrich stadials. The

  17. Effective Climate Refugia for Cold-water Fishes

    NASA Astrophysics Data System (ADS)

    Ebersole, J. L.; Morelli, T. L.; Torgersen, C.; Isaak, D.; Keenan, D.; Labiosa, R.; Fullerton, A.; Massie, J.

    2015-12-01

    Climate change threatens to create fundamental shifts in in the distributions and abundances of endothermic organisms such as cold-water salmon and trout species (salmonids). Recently published projected declines in mid-latitude salmonid distributions under future climates range from modest to severe, depending on modeling approaches, assumptions, and spatial context of analyses. Given these projected losses, increased emphasis on management for ecosystem resilience to help buffer cold-water fish populations and their habitats against climate change is emerging. Using terms such as "climate-proofing", "climate-ready", and "climate refugia", such efforts stake a claim for an adaptive, anticipatory planning response to the climate change threat. To be effective, such approaches will need to address critical uncertainties in both the physical basis for projected landscape changes in water temperature and streamflow, as well as the biological responses of organisms. Recent efforts define future potential climate refugia based on projected streamflows, air temperatures, and associated water temperature changes. These efforts reflect the relatively strong conceptual foundation for linkages between regional climate change and local hydrological responses and thermal dynamics. Yet important questions remain. Drawing on case studies throughout the Pacific Northwest, we illustrate some key uncertainties in the responses of salmonids and their habitats to altered hydro-climatic regimes currently not well addressed by physical or ecological models. Key uncertainties include biotic interactions, organismal adaptive capacity, local climate decoupling due to groundwater-surface water interactions, the influence of human engineering responses, and synergies between climatic and other stressors. These uncertainties need not delay anticipatory planning, but rather highlight the need for identification and communication of actions with high probabilities of success, and targeted

  18. Evidence for coal forest refugia in the seasonally dry Pennsylvanian tropical lowlands of the Illinois Basin, USA

    PubMed Central

    Stevenson, Robert A.; Van Hoof, Thomas B.; Mander, Luke

    2014-01-01

    The Moscovian plant macroflora at Cottage Grove southeastern Illinois, USA, is a key example of Pennsylvanian (323–299 Million years ago) dryland vegetation. There is currently no palynological data from the same stratigraphic horizons as the plant macrofossils, leaves and other vegetative and reproductive structures, at this locality. Consequently, reconstructions of the standing vegetation at Cottage Grove from these sediments lack the complementary information and a more regional perspective that can be provided by sporomorphs (prepollen, pollen, megaspores and spores). In order to provide this, we have analysed the composition of fossil sporomorph assemblages in two rock samples taken from macrofossil-bearing inter-coal shale at Cottage Grove. Our palynological data differ considerably in composition and in the dominance-diversity profile from the macrofossil vegetation at this locality. Walchian conifers and pteridosperms are common elements in the macroflora, but are absent in the sporomorph assemblages. Reversely, the sporomorph assemblages at Cottage Grove comprise 17 spore taxa (∼16% and ∼63% of the total assemblages) that are known from the lycopsid orders Isoetales, Lepidodendrales and Selaginallales, while Cottage Grove’s macrofloral record fails to capture evidence of a considerable population of coal forest lycopsids. We interpret our results as evidence that the Pennsylvanian dryland glacial landscape at Cottage Grove included fragmented populations of wetland plants living in refugia. PMID:25392752

  19. Spatial refugia mediate juvenile coral survival during coral-predator interactions

    NASA Astrophysics Data System (ADS)

    Gallagher, Clare; Doropoulos, Christopher

    2017-03-01

    Coral recruitment and juvenile growth are essential processes for coral population maintenance and recovery. A growing body of research has evaluated the influence of reef microstructure on coral settlement and post-settlement survival, showing that physical refugia enhance recruitment. These studies have evaluated coral recruit morality from competition with macroalgae and indirect predation by grazing organisms, but the impact of direct predation by corallivorous piscine species on juvenile corals and how this interacts with reef microstructure is relatively unknown. This study examined whether refugia provided by micro-crevices enhance juvenile coral survival from corallivory. Juvenile corals from two different functional groups, the slow-growing massive Porites lobata and fast-growing branching Pocillopora damicornis, with average nubbin sizes of 1.4 cm × 0.3 cm and 0.5 cm × 1.0 cm (diameter × height), respectively, were attached to experimental tiles using small (1.44 cm3) and large (8.0 cm3) crevice sizes and were monitored for 29 d on a forereef in Palau. Full crevices (four sided) enhanced coral survival compared to exposed microhabitats in both coral taxa, but crevice size did not alter survival rates. Corallivores targeted recruits within crevices regardless of crevice size; dominant predators included small triggerfish (Balistidae), butterflyfish ( Chaetodon), and wrasse ( Cheilinus). Overall, Pocillopora suffered much higher rates of mortality than Porites. All Pocillopora were consumed by day 8 of the experiment, but mortality was significantly delayed in full crevices compared to exposed and partial crevice (three sided) microhabitats. In contrast, Por. lobata located in all microhabitats survived the entire experiment up to 29 d, with high survival in full (>90%) and partial crevices (70%), but only 28% survival in exposed microhabitats. These findings show the importance of crevices as spatial refugia from predators for juvenile corals and

  20. Rapid Characterisation of Vegetation Structure to Predict Refugia and Climate Change Impacts across a Global Biodiversity Hotspot

    PubMed Central

    Schut, Antonius G. T.; Wardell-Johnson, Grant W.; Yates, Colin J.; Keppel, Gunnar; Baran, Ireneusz; Franklin, Steven E.; Hopper, Stephen D.; Van Niel, Kimberley P.; Mucina, Ladislav; Byrne, Margaret

    2014-01-01

    Identification of refugia is an increasingly important adaptation strategy in conservation planning under rapid anthropogenic climate change. Granite outcrops (GOs) provide extraordinary diversity, including a wide range of taxa, vegetation types and habitats in the Southwest Australian Floristic Region (SWAFR). However, poor characterization of GOs limits the capacity of conservation planning for refugia under climate change. A novel means for the rapid identification of potential refugia is presented, based on the assessment of local-scale environment and vegetation structure in a wider region. This approach was tested on GOs across the SWAFR. Airborne discrete return Light Detection And Ranging (LiDAR) data and Red Green and Blue (RGB) imagery were acquired. Vertical vegetation profiles were used to derive 54 structural classes. Structural vegetation types were described in three areas for supervised classification of a further 13 GOs across the region. Habitat descriptions based on 494 vegetation plots on and around these GOs were used to quantify relationships between environmental variables, ground cover and canopy height. The vegetation surrounding GOs is strongly related to structural vegetation types (Kappa = 0.8) and to its spatial context. Water gaining sites around GOs are characterized by taller and denser vegetation in all areas. The strong relationship between rainfall, soil-depth, and vegetation structure (R2 of 0.8–0.9) allowed comparisons of vegetation structure between current and future climate. Significant shifts in vegetation structural types were predicted and mapped for future climates. Water gaining areas below granite outcrops were identified as important putative refugia. A reduction in rainfall may be offset by the occurrence of deeper soil elsewhere on the outcrop. However, climate change interactions with fire and water table declines may render our conclusions conservative. The LiDAR-based mapping approach presented enables the

  1. Rapid characterisation of vegetation structure to predict refugia and climate change impacts across a global biodiversity hotspot.

    PubMed

    Schut, Antonius G T; Wardell-Johnson, Grant W; Yates, Colin J; Keppel, Gunnar; Baran, Ireneusz; Franklin, Steven E; Hopper, Stephen D; Van Niel, Kimberley P; Mucina, Ladislav; Byrne, Margaret

    2014-01-01

    Identification of refugia is an increasingly important adaptation strategy in conservation planning under rapid anthropogenic climate change. Granite outcrops (GOs) provide extraordinary diversity, including a wide range of taxa, vegetation types and habitats in the Southwest Australian Floristic Region (SWAFR). However, poor characterization of GOs limits the capacity of conservation planning for refugia under climate change. A novel means for the rapid identification of potential refugia is presented, based on the assessment of local-scale environment and vegetation structure in a wider region. This approach was tested on GOs across the SWAFR. Airborne discrete return Light Detection And Ranging (LiDAR) data and Red Green and Blue (RGB) imagery were acquired. Vertical vegetation profiles were used to derive 54 structural classes. Structural vegetation types were described in three areas for supervised classification of a further 13 GOs across the region. Habitat descriptions based on 494 vegetation plots on and around these GOs were used to quantify relationships between environmental variables, ground cover and canopy height. The vegetation surrounding GOs is strongly related to structural vegetation types (Kappa = 0.8) and to its spatial context. Water gaining sites around GOs are characterized by taller and denser vegetation in all areas. The strong relationship between rainfall, soil-depth, and vegetation structure (R(2) of 0.8-0.9) allowed comparisons of vegetation structure between current and future climate. Significant shifts in vegetation structural types were predicted and mapped for future climates. Water gaining areas below granite outcrops were identified as important putative refugia. A reduction in rainfall may be offset by the occurrence of deeper soil elsewhere on the outcrop. However, climate change interactions with fire and water table declines may render our conclusions conservative. The LiDAR-based mapping approach presented enables the

  2. What triggers the rising of an intraspecific biodiversity hotspot? Hints from the agile frog.

    PubMed

    Canestrelli, Daniele; Bisconti, Roberta; Sacco, Florinda; Nascetti, Giuseppe

    2014-05-23

    Hotspots of genetic diversity are regions of utmost importance for species survival and conservation, and their intimate link with the geographic location of glacial refugia has been well established. Nonetheless, the microevolutionary processes underlying the generation of hotspots in such regions have only recently become a fervent field of research. We investigated the phylogeographic and population genetic structure of the agile frog, Rana dalmatina, within its putative refugium in peninsular Italy. We found this region to harbour far more diversity, phylogeographic structure, and lineages of ancient origin than that by the rest of the species' range in Europe. This pattern appeared to be well explained by climate-driven microevolutionary processes that occurred during both glacial and interglacial epochs. Therefore, the inferred evolutionary history of R. dalmatina in Italy supports a view of glacial refugia as 'factories' rather than as repositories of genetic diversity, with significant implications for conservation strategies for hotspots.

  3. Phylogeography and demographic history of Lacerta lepida in the Iberian Peninsula: multiple refugia, range expansions and secondary contact zones

    PubMed Central

    2011-01-01

    Background The Iberian Peninsula is recognized as an important refugial area for species survival and diversification during the climatic cycles of the Quaternary. Recent phylogeographic studies have revealed Iberia as a complex of multiple refugia. However, most of these studies have focused either on species with narrow distributions within the region or species groups that, although widely distributed, generally have a genetic structure that relates to pre-Quaternary cladogenetic events. In this study we undertake a detailed phylogeographic analysis of the lizard species, Lacerta lepida, whose distribution encompasses the entire Iberian Peninsula. We attempt to identify refugial areas, recolonization routes, zones of secondary contact and date demographic events within this species. Results Results support the existence of 6 evolutionary lineages (phylogroups) with a strong association between genetic variation and geography, suggesting a history of allopatric divergence in different refugia. Diversification within phylogroups is concordant with the onset of the Pleistocene climatic oscillations. The southern regions of several phylogroups show a high incidence of ancestral alleles in contrast with high incidence of recently derived alleles in northern regions. All phylogroups show signs of recent demographic and spatial expansions. We have further identified several zones of secondary contact, with divergent mitochondrial haplotypes occurring in narrow zones of sympatry. Conclusions The concordant patterns of spatial and demographic expansions detected within phylogroups, together with the high incidence of ancestral haplotypes in southern regions of several phylogroups, suggests a pattern of contraction of populations into southern refugia during adverse climatic conditions from which subsequent northern expansions occurred. This study supports the emergent pattern of multiple refugia within Iberia but adds to it by identifying a pattern of refugia coincident

  4. Phylogeography and demographic history of Lacerta lepida in the Iberian Peninsula: multiple refugia, range expansions and secondary contact zones.

    PubMed

    Miraldo, Andreia; Hewitt, Godfrey M; Paulo, Octavio S; Emerson, Brent C

    2011-06-17

    The Iberian Peninsula is recognized as an important refugial area for species survival and diversification during the climatic cycles of the Quaternary. Recent phylogeographic studies have revealed Iberia as a complex of multiple refugia. However, most of these studies have focused either on species with narrow distributions within the region or species groups that, although widely distributed, generally have a genetic structure that relates to pre-Quaternary cladogenetic events. In this study we undertake a detailed phylogeographic analysis of the lizard species, Lacerta lepida, whose distribution encompasses the entire Iberian Peninsula. We attempt to identify refugial areas, recolonization routes, zones of secondary contact and date demographic events within this species. Results support the existence of 6 evolutionary lineages (phylogroups) with a strong association between genetic variation and geography, suggesting a history of allopatric divergence in different refugia. Diversification within phylogroups is concordant with the onset of the Pleistocene climatic oscillations. The southern regions of several phylogroups show a high incidence of ancestral alleles in contrast with high incidence of recently derived alleles in northern regions. All phylogroups show signs of recent demographic and spatial expansions. We have further identified several zones of secondary contact, with divergent mitochondrial haplotypes occurring in narrow zones of sympatry. The concordant patterns of spatial and demographic expansions detected within phylogroups, together with the high incidence of ancestral haplotypes in southern regions of several phylogroups, suggests a pattern of contraction of populations into southern refugia during adverse climatic conditions from which subsequent northern expansions occurred. This study supports the emergent pattern of multiple refugia within Iberia but adds to it by identifying a pattern of refugia coincident with the southern distribution

  5. The Post-Glacial Species Velocity of Picea glauca following the Last Glacial Maximum in Alaska.

    NASA Astrophysics Data System (ADS)

    Morrison, B. D.; Napier, J.; Kelly, R.; Li, B.; Heath, K.; Hug, B.; Hu, F.; Greenberg, J. A.

    2015-12-01

    Anthropogenic climate change is leading to dramatic fluctuations to Earth's biodiversity that has not been observed since past interglacial periods. There is rising concern that Earth's warming climate will have significant impacts to current species ranges and the ability of a species to persist in a rapidly changing environment. The paleorecord provides information on past species distributions in relation to climate change, which can illuminate the patterns of potential future distributions of species. Particularly in areas where there are multiple potential limiting factors on a species' range, e.g. temperature, radiation, and evaporative demand, the spatial patterns of species migrations may be particularly complex. In this study, we assessed the change in the distributions of white spruce (Picea glauca) from the Last Glacial Maxima (LGM) to present-day for the entire state of Alaska. To accomplish this, we created species distribution models (SDMs) calibrated from modern vegetation data and high-resolution, downscaled climate surfaces at 60m. These SDMs were applied to downscaled modern and paleoclimate surfaces to produce estimated ranges of white spruce during the LGM and today. From this, we assessed the "species velocity", the rate at which white spruce would need to migrate to keep pace with climate change, with the goal of determining whether the expansion from the LGM to today originated from microclimate refugia. Higher species velocities indicate locations where climate changed drastically and white spruce would have needed to migrate rapidly to persist and avoid local extinction. Conversely, lower species velocities indicated locations where the local climate was changing less rapidly or was within the center of the range of white spruce, and indicated locations where white spruce distributions were unlikely to have changed significantly. Our results indicate the importance of topographic complexity in buffering the effects of climate change

  6. Differences in Bacterial Diversity and Communities Between Glacial Snow and Glacial Soil on the Chongce Ice Cap, West Kunlun Mountains.

    PubMed

    Yang, Guang Li; Hou, Shu Gui; Le Baoge, Ri; Li, Zhi Guo; Xu, Hao; Liu, Ya Ping; Du, Wen Tao; Liu, Yong Qin

    2016-11-04

    A detailed understanding of microbial ecology in different supraglacial habitats is important due to the unprecedented speed of glacier retreat. Differences in bacterial diversity and community structure between glacial snow and glacial soil on the Chongce Ice Cap were assessed using 454 pyrosequencing. Based on rarefaction curves, Chao1, ACE, and Shannon indices, we found that bacterial diversity in glacial snow was lower than that in glacial soil. Principal coordinate analysis (PCoA) and heatmap analysis indicated that there were major differences in bacterial communities between glacial snow and glacial soil. Most bacteria were different between the two habitats; however, there were some common bacteria shared between glacial snow and glacial soil. Some rare or functional bacterial resources were also present in the Chongce Ice Cap. These findings provide a preliminary understanding of the shifts in bacterial diversity and communities from glacial snow to glacial soil after the melting and inflow of glacial snow into glacial soil.

  7. Characterization of Thermal Refugia and Biogeochemical Hotspots at Sleepers River Watershed, VT

    NASA Astrophysics Data System (ADS)

    Hwang, K.; Chandler, D. G.; Kelleher, C.; Shanley, J. B.; Shaw, S. B.

    2017-12-01

    During low flow, changes in the extent of the channel network in headwater catchments depend on groundwater-surface water interactions, and dictate thermal and biogeochemical heterogeneities. Channel reaches with low temperature may act as refugia for valued species such as brook trout, and warmer reaches with high dissolved organic matter may act as biogeochemical hotspots. Prior studies have found uniform scaling of hydrologic and biogeochemical processes above certain spatial thresholds but sizable heterogeneities in these processes below the threshold. We utilize high resolution measurements of water quality parameters including stream temperature, conductivity and fluorescent dissolved organic matter (fDOM) at tributaries in two catchments of Sleepers River Watershed, Vermont to investigate seasonal and spatial variation of water quality and scaling of stream chemistry within the intensive study area and the larger Sleepers River Watershed. This study leverages findings from various small scale regional studies to identify differences in headwater channel reach behavior in a similar climate across some dissimilar geomorphic units, to inform the identification of thermal refugia and biogeochemical hotspots.

  8. Plant growth on debris covered glacier surfaces - ecology, vegetation patterns and implications for debris mantled glaciers serving as cold and warm stage plant refugia in the past

    NASA Astrophysics Data System (ADS)

    Fickert, Thomas; Friend, Donald; Grüninger, Friederike; Molnia, Bruce; Richter, Michael

    2017-04-01

    As stated at the International Conference on Debris-Covered Glaciers in 2000, "debris-covered glaciers comprise a significant fraction of the global population of glaciers...." Given a minimum of debris thickness and sufficient stability, these surfaces host surprisingly diverse plant assemblages, both floristically and structurally. Observations of plant growth on glacier surfaces are reported from around the world - including mature forests with trees more than 50cm in diameter. Debris covered glacier surfaces are mobile habitats for plants, which migrate downhill with glacier movement, but are able to spread upward with strong anabatic valley winds. Plant growth is possible even on a very shallow debris cover. Depending on site conditions, floristic composition and structure of vegetation on debris covered glaciers represent a mosaic of environments, including subnival pioneer communities, glacier foreland early- to late-successional stages, and morainal locations. The taxa involved display a wide spectrum of adaptations to habitat conditions with particular migration and dispersal strategies. With a shallow debris cover, alpine/subnival taxa can grow considerably below their usual altitudinal niche due to the cooler subsurface soil temperatures. In contrast, a greater thickness of debris cover allows even thermophilous plants of lower elevations to grow on glacier surfaces. Employing the principle of actualism, debris covered glaciers provided important and previously undocumented refugia for plants during the Pleistocene cold stages from which alpine and arctic plant species were able to re-establish and spread in post-glacial time. This assumption is complementary to the two competing ideas to explain the fate of alpine and/or arctic taxa during the Pleistocene, the nunatak hypothesis (i.e. in-situ survival of plants on unglaciated summits) and tabula rasa theory (i.e. displacement of plants and subsequent remigration). Vice versa debris covered glaciers

  9. What triggers the rising of an intraspecific biodiversity hotspot? Hints from the agile frog

    PubMed Central

    Canestrelli, Daniele; Bisconti, Roberta; Sacco, Florinda; Nascetti, Giuseppe

    2014-01-01

    Hotspots of genetic diversity are regions of utmost importance for species survival and conservation, and their intimate link with the geographic location of glacial refugia has been well established. Nonetheless, the microevolutionary processes underlying the generation of hotspots in such regions have only recently become a fervent field of research. We investigated the phylogeographic and population genetic structure of the agile frog, Rana dalmatina, within its putative refugium in peninsular Italy. We found this region to harbour far more diversity, phylogeographic structure, and lineages of ancient origin than that by the rest of the species' range in Europe. This pattern appeared to be well explained by climate-driven microevolutionary processes that occurred during both glacial and interglacial epochs. Therefore, the inferred evolutionary history of R. dalmatina in Italy supports a view of glacial refugia as ‘factories' rather than as repositories of genetic diversity, with significant implications for conservation strategies for hotspots. PMID:24853644

  10. Using Agent-Based Modelling to Predict the Role of Wild Refugia in the Evolution of Resistance of Sea Lice to Chemotherapeutants.

    PubMed

    McEwan, Gregor F; Groner, Maya L; Fast, Mark D; Gettinby, George; Revie, Crawford W

    2015-01-01

    A major challenge for Atlantic salmon farming in the northern hemisphere is infestation by the sea louse parasite Lepeophtheirus salmonis. The most frequent method of controlling these sea louse infestations is through the use of chemical treatments. However, most major salmon farming areas have observed resistance to common chemotherapeutants. In terrestrial environments, many strategies employed to manage the evolution of resistance involve the use of refugia, where a portion of the population is left untreated to maintain susceptibility. While refugia have not been deliberately used in Atlantic salmon farming, wild salmon populations that migrate close to salmon farms may act as natural refugia. In this paper we describe an agent-based model that explores the influence of different sizes of wild salmon populations on resistance evolution in sea lice on a salmon farm. Using the model, we demonstrate that wild salmon populations can act as refugia that limit the evolution of resistance in the sea louse populations. Additionally, we demonstrate that an increase in the size of the population of wild salmon results in an increased effect in slowing the evolution of resistance. We explore the effect of a population fitness cost associated with resistance, finding that in some cases it substantially reduces the speed of evolution to chemical treatments.

  11. Using Agent-Based Modelling to Predict the Role of Wild Refugia in the Evolution of Resistance of Sea Lice to Chemotherapeutants

    PubMed Central

    McEwan, Gregor F.; Groner, Maya L.; Fast, Mark D.; Revie, Crawford W.

    2015-01-01

    A major challenge for Atlantic salmon farming in the northern hemisphere is infestation by the sea louse parasite Lepeophtheirus salmonis. The most frequent method of controlling these sea louse infestations is through the use of chemical treatments. However, most major salmon farming areas have observed resistance to common chemotherapeutants. In terrestrial environments, many strategies employed to manage the evolution of resistance involve the use of refugia, where a portion of the population is left untreated to maintain susceptibility. While refugia have not been deliberately used in Atlantic salmon farming, wild salmon populations that migrate close to salmon farms may act as natural refugia. In this paper we describe an agent-based model that explores the influence of different sizes of wild salmon populations on resistance evolution in sea lice on a salmon farm. Using the model, we demonstrate that wild salmon populations can act as refugia that limit the evolution of resistance in the sea louse populations. Additionally, we demonstrate that an increase in the size of the population of wild salmon results in an increased effect in slowing the evolution of resistance. We explore the effect of a population fitness cost associated with resistance, finding that in some cases it substantially reduces the speed of evolution to chemical treatments. PMID:26485023

  12. Glacial stages and post-glacial environmental evolution in the Upper Garonne valley, Central Pyrenees.

    PubMed

    Fernandes, M; Oliva, M; Palma, P; Ruiz-Fernández, J; Lopes, L

    2017-04-15

    The maximum glacial extent in the Central Pyrenees during the Last Glaciation is known to have occurred before the global Last Glacial Maximum, but the succession of cold events afterwards and their impact on the landscape are still relatively unknown. This study focuses on the environmental evolution in the upper valley of the Garonne River since the Last Glaciation. Geomorphological mapping allows analysis of the spatial distribution of inherited and current processes and landforms in the study area. The distribution of glacial records (moraines, till, erratic boulders, glacial thresholds) suggests the existence of four glacial stages, from the maximum expansion to the end of the glaciation. GIS modeling allows quantification of the Equilibrium Line Altitude, extent, thickness and volume of ice in each glacial stage. During the first stage, the Garonne glacier reached 460m in the Loures-Barousse-Barbazan basin, where it formed a piedmont glacier 88km from the head and extended over 960km 2 . At a second stage of glacier stabilization during the deglaciation process, the valley glaciers were 12-23km from the head until elevations of 1000-1850m, covering an area of 157km 2 . Glaciers during stage three remained isolated in the upper parts of the valley, at heights of 2050-2200m and 2.6-4.5km from the head, with a glacial surface of 16km 2 . In stage four, cirque glaciers were formed between 2260m and 2590m, with a length of 0.4-2km and a glacial area of 5.7km 2 . Also, the wide range of periglacial, slope, nival and alluvial landforms existing in the formerly glaciated environments allows reconstruction of the post-glacial environmental dynamics in the upper Garonne basin. Today, the highest lands are organized following three elevation belts: subnival (1500-1900m), nival (1900-2300m) and periglacial/cryonival (2300-2800m). Copyright © 2017 Elsevier B.V. All rights reserved.

  13. Vegetation dynamics during the Last Interglacial-Glacial cycle in the Arno coastal plain (Tuscany, western Italy): location of a new tree refuge

    NASA Astrophysics Data System (ADS)

    Lucchi, M. Ricci

    2008-12-01

    Pollen analysis of the pre-Last Glacial Maximum succession of a 105 m-long continuous core from Tirrenia (Tuscany) provides evidence for the existence of an area of relatively high ecological stability where the effects of climate change were mitigated. The chronological framework of the vegetation record, spanning the Last Interglacial-Glacial cycle, was established by (i) AMS 14C dating, (ii) correlation with well-dated pollen sequences, and (iii) local stratigraphical constraints. A high lithological and sedimentological variability, with facies associations changing from fluvial to alluvial and coastal plain, enhances the palaeoenvironmental control on pollen distribution, thus helping to discriminate the impact of local factors on vegetation history. The most remarkable evidence, however, is represented by the continuous record of temperate trees throughout the whole glacial period, which provides useful indications on the location and nature of cold stage refugia. Most of the vegetation changes recorded in the core can be compared to the vegetation history of the Last Interglacial-Glacial cycle from southern Europe as a whole. In addition, local geographic and environmental features account for a more complex and varied floristic composition. Only the last phase of the Penultimate Glacial (MIS6), which was characterized by the diffusion of an arid steppe tundra, is recorded at the base of the core. The subsequent Last Interglacial (MIS5e) interval shows a poor and scattered pollen content due to the instability of the sedimentary environment. Nevertheless, it provides evidence of both global and local controls on vegetation dynamics, as indicated by the initial expansion of thermophilous forests and the remarkably late diffusion of conifers ( Pinus-Abies-Picea forests), respectively. Similarly, the transition to the Last Glacial (MIS5b and 5a in the core) is characterized by a reduced vegetation response to the typical stadial/interstadial climate variability

  14. Geomorphology of the Southwest Coast of County Cork, Ireland: A Look into the Rocks, Folds, and Glacial Scours

    NASA Astrophysics Data System (ADS)

    Bowden, S.; Wireman, R.; Sautter, L.; Beutel, E. K.

    2015-12-01

    Bathymetric data were collected off the southwest coast of County Cork, Ireland by the joint INFOMAR project between the Marine Institute of Ireland and the Geologic Survey of Ireland. Data were collected using a Kongsberg EM2040 multibeam sonar on the R/V Celtic Voyager, in August and September 2014, and were post-processed with CARIS HIPS and SIPS 8.1 and 9.0 software to create 2D and 3D bathymetric surfaces. From the computer generated images, some of the lithologic formations were relatively aged and observed. The studied regions range in depth from 20 to 118 m, with shallower areas to the northeast. Several large rock outcrops occur, the larger of which shows a vertical rise of nearly 20 m. These outcrops are oriented in a northeast-southwest direction, and exhibit significant bed folding, regional folding, tilted beds, and cross joints. The folds studied are plunging chevron folds. These folds have a northeast-southwest fold axis orthogonal to the cross joints and are older relative to the jointing systems. The NE-SW joints are older than the NW-SE joints due to their correlation with drainage and erosion patterns. Regional folding is the youngest feature due to its superposition on the chevron folding and jointing systems. The interaction of cross jointing and folding is consistent with the geologic history of the area, and creates a unique bathymetry worthy of further study.

  15. Glacial history of Tranquilo glacier (Central Patagonia) since the Last Glacial Maximum through to the present.

    NASA Astrophysics Data System (ADS)

    Sagredo, E. A.; Araya, P. S.; Schaefer, J. M.; Kaplan, M. R.; Kelly, M. A.; Lowell, T. V.; Aravena, J. C.

    2014-12-01

    Deciphering the timing and the inter-hemispheric phasing of former glacial fluctuations is critical for understanding the mechanisms and climate signals underlying these glacial events. Here, we present a detailed chronology of glacial fluctuations for Río Tranquilo glacier (47°S), since the LGM, including up to the present. Río Tranquilo is a small glacial valley located on the northern flank of Monte San Lorenzo, an isolated granitic massif, ~70 km to the east of the southern limit of the Northern Patagonian Icefield. Although Mt. San Lorenzo is located on the leeward side of the Andes, it is one of the most glacierized mountains in the region, with an ice surface area of ~140 km2. Geomorphic evidence suggests that during past episodes of climate change several small glaciers that today occupy the headwalls of Río Tranquilo valley expanded and coalesced, depositing a series of moraines complexes along the flanks and bottom of the valley. We used two independent dating techniques to constrain the age of the glacial history of the area. 10Be surface exposure ages from boulders located atop moraine ridges reveal that Río Tranquilo valley underwent glacial expansion/stabilization during at least the LGM (late LGM?), Late glacial (ACR and Younger Dryas) and Mid-Holocene. Within the Mid-Holocene limits, tree-ring based chronology indicates that Río Tranquilo glacier expanded during the Late Holocene as well. Our results are the first detailed chronology of glacial fluctuations from a single valley glacier, spanning the entire period from the (end of the) LGM up to the present, in southern South America. By identifying different glacial episodes within a single alpine valley, this study provides baseline data for studying the relative magnitude of the climate events responsible for these glacial events.

  16. Possible refugia in the Alexander Archipelago of southeastern Alaska during the late Wisconsin glaciation

    USGS Publications Warehouse

    Carrara, P.E.; Ager, T.A.; Baichtal, J.F.

    2007-01-01

    The interpretation of the extent of late Wisconsin glaciation in southeastern Alaska has varied between geologists and biologists. Maps and reports of the region prepared by geologists commonly indicated that late Wisconsin ice extended as a large uniform front west to the edge of the continental shelf. However, the distribution of plants and animals in the region has led many biologists to suggest that there may have been ice-free areas that served as refugia during the late Wisconsin. Based on analyses of aerial photographs, topographic maps, and bathymetric charts, in conjunction with a review of previous literature and reconnaissance fieldwork throughout the region, this study presents data supporting a limited ice extent in the Alexander Archipelago during the late Wisconsin and identifies possible ice-free areas that may have served as refugia. These areas include (1) the Fairweather Ground, (2) the Herbert Graves Island area, (3) the western coast of southern Baranof Island and adjacent continental shelf, (4) Coronation Island and the adjacent continental shelf, (5) the Warren Island area, (6) the continental shelf from west of Heceta Island to Forrester Island in the south, (7) parts of the west coast of southern Dall Island, and (8) lowland areas in southern Prince of Wales Island. The identification of these possible refugia has bearing on the recolonization of the Alexander Archipelago, as they could have served as centers of biotic dispersal upon regional deglaciation and as stepping stones for early humans with a maritime tradition entering the western hemisphere from Asia. ?? 2007 NRC Canada.

  17. Genetic variability and glacial origins of yellow perch (Perca flavescens) in North America

    USGS Publications Warehouse

    Todd, Thomas N.; Hatcher, Charles O.

    1993-01-01

    Starch–gel electrophoresis was used to analyze muscle and liver tissue for variation in 13 enzymes representing 31 presumptive loci in yellow perch (Perca flavescens) from 13 localities scattered throughout the natural geographic range of the species in North America. Ten loci were polymorphic, but only three, alcohol dehydrogenase (ADH-1*), glucose-6-phosphate isomerase (GPI-1*), and phosphoglucomutase (PGM-2*), exhibited polymorphisms at relatively high frequencies across localities. Western populations were fixed for one allele at ADH-1*, eastern populations were fixed for another allele, and populations from intermediate locations in Lake Ontario and Pennsylvania had both alleles. The distributions of alleles at GPI-1* and PGM-2*were similar to that of ADH-1*, exhibiting strong differences between eastern and western populations, although the delineation was not as clear. Western populations were much less variable than eastern populations, and the distribution of alleles indicated that the two groups were derived from Mississippi and Atlantic glacial refugia. Populations near the physiographic discontinuity between the Mississippi and Atlantic drainages in western New York and Pennsylvania exhibited an admixture of typically western and eastern alleles. Such observations are consistent with the mixed faunal history of the region and limited postglacial dispersal of western and eastern populations across the boundary.

  18. Congruence between distribution modelling and phylogeographical analyses reveals Quaternary survival of a toadflax species (Linaria elegans) in oceanic climate areas of a mountain ring range.

    PubMed

    Fernández-Mazuecos, Mario; Vargas, Pablo

    2013-06-01

    · The role of Quaternary climatic shifts in shaping the distribution of Linaria elegans, an Iberian annual plant, was investigated using species distribution modelling and molecular phylogeographical analyses. Three hypotheses are proposed to explain the Quaternary history of its mountain ring range. · The distribution of L. elegans was modelled using the maximum entropy method and projected to the last interglacial and to the last glacial maximum (LGM) using two different paleoclimatic models: the Community Climate System Model (CCSM) and the Model for Interdisciplinary Research on Climate (MIROC). Two nuclear and three plastid DNA regions were sequenced for 24 populations (119 individuals sampled). Bayesian phylogenetic, phylogeographical, dating and coalescent-based population genetic analyses were conducted. · Molecular analyses indicated the existence of northern and southern glacial refugia and supported two routes of post-glacial recolonization. These results were consistent with the LGM distribution as inferred under the CCSM paleoclimatic model (but not under the MIROC model). Isolation between two major refugia was dated back to the Riss or Mindel glaciations, > 100 kyr before present (bp). · The Atlantic distribution of inferred refugia suggests that the oceanic (buffered)-continental (harsh) gradient may have played a key and previously unrecognized role in determining Quaternary distribution shifts of Mediterranean plants. © 2013 The Authors. New Phytologist © 2013 New Phytologist Trust.

  19. Evidence of a low-latitude glacial buzzsaw: Progressive hypsometry reveals height-limiting glacial erosion in tropical mountain belts

    NASA Astrophysics Data System (ADS)

    Cunningham, M.; Stark, C. P.; Kaplan, M. R.; Schaefer, J. M.; Winckler, G.

    2017-12-01

    It has been widely demonstrated that glacial erosion limits the height of mid-latitude mountain ranges—a phenomenon commonly referred to as the "glacial buzzsaw." The strength of the buzzsaw is thought to diminish, or die out completely, at lower latitudes, where glacial landscapes occupy only a small part of mountain belts affected by Pleistocene glaciation. Here we argue that glacial erosion has actually truncated the rise of many tropical orogens. To elicit signs of height-limiting glacial erosion in the tropics, we employ a new take on an old tool: we identify transient geomorphic features by tracking the evolution of (sub)catchment hypsometry with increasing elevation above base level, a method we term "progressive hypsometry." In several tropical mountain belts, including the Central Range of Taiwan, the Talamanca of Costa Rica, the Finisterres of Papua New Guinea, and the Rwenzoris of East Africa, progressive hypsometry reveals transient landscapes perched at various elevations, but the highest of these transient features are consistently glacial landscapes near the lower limit of late-Pleistocene glacial equilibrium line altitude (ELA) fluctuation. We attribute this pattern to an efficient glacial buzzsaw. In many cases, these glacial landscapes are undergoing contemporary destruction by headward propagating, fluvially-driven escarpments. We deduce that a duel between glacial buzzcutting and fluvially-driven scarp propagation has been ongoing throughout the Pleistocene in these places, and that the preservation potential of tropical glacial landscapes is low. To this end, we have identified possible remnants of glacial landscapes in the final stages of scarp consumption, and use 3He surface exposure age dating of boulders and bedrock surfaces in two of these landscapes to constrain major geomorphic activity to before the onset of the Last Glacial Maximum. Our work points to a profound climatic influence on the evolution of these warm, tectonically active

  20. Evolution of the Northern Rockweed, Fucus distichus, in a Regime of Glacial Cycling: Implications for Benthic Algal Phylogenetics.

    PubMed

    Laughinghouse, Haywood Dail; Müller, Kirsten M; Adey, Walter H; Lara, Yannick; Young, Robert; Johnson, Gabriel

    2015-01-01

    Northern hemisphere rockweeds (Fucus) are thought to have evolved in the North Pacific and then spread to the North Atlantic following the opening of the Bering Strait. They have dispersed and widely speciated in the North Atlantic and its tributary seas. Fucus distichus is likely near the ancestral member of this genus, and studies have shown that there are several species/subspecies in this complex (i.e. F. evanescens and F. gardneri). We used phylogenetic and haplotype analyses to test the phylogenetic relationships and biogeography of F. distichus. Our data and subsequent analyses demonstrate that, unlike previous studies that lacked samples from an extensive geographical area of the Arctic and Subarctic, there is a distinct Arctic haplotype that is the source of subspecies in both the North Pacific and North Atlantic. Fucus distichus occupies a low tide zone habitat, and in Arctic/Subarctic regions it is adapted to the severe stress of sea ice coverage and disturbance during many months per year. We hypothesize that the very large geographic area of Arctic and Subarctic rocky shores available to this species during interglacials, supported by large Arctic/Subarctic fringe areas as well as unglaciated refugia during glacial cycles, provided a robust population and gene pool (described by the Thermogeographic Model). This gene pool dilutes that of the more fragmented and area-limited Temperate/Boreal area populations when they are brought together during glacial cycles. We suggest that similar subspecies complexes for a variety of Arctic/Subarctic shore biota should be examined further in this context, rather than arbitrarily being split up into numerous species.

  1. Modelling the role of groundwater hydro-refugia in East African hominin evolution and dispersal

    PubMed Central

    Cuthbert, M. O.; Gleeson, T.; Reynolds, S. C.; Bennett, M. R.; Newton, A. C.; McCormack, C. J.; Ashley, G. M.

    2017-01-01

    Water is a fundamental resource, yet its spatiotemporal availability in East Africa is poorly understood. This is the area where most hominin first occurrences are located, and consequently the potential role of water in hominin evolution and dispersal remains unresolved. Here, we show that hundreds of springs currently distributed across East Africa could function as persistent groundwater hydro-refugia through orbital-scale climate cycles. Groundwater buffers climate variability according to spatially variable groundwater response times determined by geology and topography. Using an agent-based model, grounded on the present day landscape, we show that groundwater availability would have been critical to supporting isolated networks of hydro-refugia during dry periods when potable surface water was scarce. This may have facilitated unexpected variations in isolation and dispersal of hominin populations in the past. Our results therefore provide a new environmental framework in which to understand how patterns of taxonomic diversity in hominins may have developed. PMID:28556825

  2. Modelling the role of groundwater hydro-refugia in East African hominin evolution and dispersal

    NASA Astrophysics Data System (ADS)

    Cuthbert, M. O.; Gleeson, T.; Reynolds, S. C.; Bennett, M. R.; Newton, A. C.; McCormack, C. J.; Ashley, G. M.

    2017-05-01

    Water is a fundamental resource, yet its spatiotemporal availability in East Africa is poorly understood. This is the area where most hominin first occurrences are located, and consequently the potential role of water in hominin evolution and dispersal remains unresolved. Here, we show that hundreds of springs currently distributed across East Africa could function as persistent groundwater hydro-refugia through orbital-scale climate cycles. Groundwater buffers climate variability according to spatially variable groundwater response times determined by geology and topography. Using an agent-based model, grounded on the present day landscape, we show that groundwater availability would have been critical to supporting isolated networks of hydro-refugia during dry periods when potable surface water was scarce. This may have facilitated unexpected variations in isolation and dispersal of hominin populations in the past. Our results therefore provide a new environmental framework in which to understand how patterns of taxonomic diversity in hominins may have developed.

  3. A 305 year monthly rainfall series for the Island of Ireland (1711-2016)

    NASA Astrophysics Data System (ADS)

    Murphy, Conor; Burt, Tim P.; Broderick, Ciaran; Duffy, Catriona; Macdonald, Neil; Matthews, Tom; McCarthy, Mark P.; Mullan, Donal; Noone, Simon; Ryan, Ciara; Thorne, Peter; Walsh, Seamus; Wilby, Robert L.

    2017-04-01

    This paper derives a continuous 305-year monthly rainfall series for the Island of Ireland (IoI) for the period 1711-2016. Two key data sources are employed: i) a previously unpublished UK Met Office Note which compiled annual rainfall anomalies and corresponding monthly per mille amounts from weather diaries and early observational records for the period 1711-1977; and ii) a long-term, homogenised monthly IoI rainfall series for the period 1850-2016. Using estimates of long-term average precipitation sampled from the quality assured series, the full record is reconstituted and insights drawn regarding notable periods and the range of climate variability and change experienced. Consistency with other long records for the region is examined, including: the England and Wales Precipitation series (EWP; 1766-2016); the early EWP Glasspoole series (1716-1765) and the Central England Temperature series (CET; 1711-2016). Strong correspondence between all records is noted from 1780 onwards. While disparities are evident between the early EWP and Ireland series, the latter shows strong decadal consistency with CET throughout the record. In addition, independent, early observations from Cork and Dublin, along with available documentary sources, corroborate the derived series and add confidence to our reconstruction. The new IoI rainfall record reveals that the wettest decades occurred in the early 18th Century, despite the fact that IoI has experienced a long-term winter wetting trend consistent with climate model projections. These exceptionally wet winters of the 1720s and 1730s were concurrent with almost unprecedented warmth in the CET, glacial advance throughout Scandinavia, and glacial retreat in West Greenland, consistent with a wintertime NAO-type forcing. Our study therefore demonstrates the value of long-term observational records for providing insight to the natural climate variability of the North Atlantic region.

  4. The Seabed and Shallow Geology Mapping of the Porcupine Bank, West of Ireland

    NASA Astrophysics Data System (ADS)

    Thébaudeau, B.; Monteys, X.; McCarron, S. G.

    2016-02-01

    The "Porcupine Bank" is a bathymetric high of over 40,000 km2 linked to the western shelf of Ireland which lies between 51-54° N and 11-15° W approximately 100 km west of Ireland. Water depths are as shallow as 145 m over the "Porcupine Ridge". The Bank's location on the north eastern fringe of the Atlantic Ocean, in a critical position between the shelf edge and the main land and along the line of the Polar Front, means it may contain significant indications of glacial/interglacial changes in northern hemisphere climate and in North Atlantic Ocean circulation. But it also means that it consists of strategically important marine environments with very likely future developmental pressures. Peer-reviewed publications on the geology of the Bank are very limited and this current state of knowledge will hamper any marine ecosystem research and protection. This paper will describe the first results of a research project aiming at filling the gap of our understanding of the region's shallow geology and subseabed resources and characteristics. As a first step, seabed geomorphology mapping using high resolution MBES and sub bottom data have highlighted a wealth of glacially derived features such as iceberg scours and elongated ridges whose formation could be directly influenced by the presence of ice on or nearby the bank. Other features interpreted as sand waves could help understand relict or modern currents. In addition to these surface features, this paper introduces recent geological mapping of the shallow stratigraphy of the bank using 2D seismic and sub bottom profiler data collected at a high density correlated with recently collected vibro-cores. The seismic units and corresponding lithofacies (some with radiocarbon dates) are consistently described and a regional correlation built.

  5. Phylogeography of Eomecon chionantha in subtropical China: the dual roles of the Nanling Mountains as a glacial refugium and a dispersal corridor.

    PubMed

    Tian, Shuang; Kou, Yixuan; Zhang, Zhirong; Yuan, Lin; Li, Derong; López-Pujol, Jordi; Fan, Dengmei; Zhang, Zhiyong

    2018-02-09

    Mountains have not only provided refuge for species, but also offered dispersal corridors during the Neogene and Quaternary global climate changes. Compared with a plethora of studies on the refuge role of China's mountain ranges, their dispersal corridor role has received little attention in plant phylogeographic studies. Using phylogeographic data of Eomecon chionantha Hance (Papaveraceae), this study explicitly tested whether the Nanling Mountains, which spans from west to east for more than 1000 km in subtropical China, could have functioned as a dispersal corridor during the late Quaternary in addition to a glacial refugium. Our analyses revealed a range-wide lack of phylogeographic structure in E. chionantha across three kinds of molecular markers [two chloroplast intergenic spacers, nuclear ribosomal internal transcribed spacer (nrITS), and six nuclear microsatellite loci]. Demographic inferences based on chloroplast and nrITS sequences indicated that E. chionantha could have experienced a strong postglacial range expansion between 6000 and 1000 years ago. Species distribution modelling showed that the Nanling Mountains and the eastern Yungui Plateau were the glacial refugia of E. chionantha. Reconstruction of dispersal corridors indicated that the Nanling Mountains also have acted as a corridor of population connectivity for E. chionantha during the late Quaternary. Our results suggest that the Nanling Mountains may acted dual roles as a dispersal corridor in east-west direction and as a glacial refugium in subtropical China during the late Quaternary. The population connectivity mediated by the mountain range and a strong postglacial range expansion are the most likely reasons for the lack of phylogeographic structure in E. chionantha. The hypothesis of dual roles of the mountain range presented here sheds new insights into the phylogeographic patterns of organisms in subtropical China.

  6. Glacial lakes in the Indian Himalayas--from an area-wide glacial lake inventory to on-site and modeling based risk assessment of critical glacial lakes.

    PubMed

    Worni, Raphael; Huggel, Christian; Stoffel, Markus

    2013-12-01

    Glacial lake hazards and glacial lake distributions are investigated in many glaciated regions of the world, but comparably little attention has been given to these topics in the Indian Himalayas. In this study we present a first area-wide glacial lake inventory, including a qualitative classification at 251 glacial lakes >0.01 km(2). Lakes were detected in the five states spanning the Indian Himalayas, and lake distribution pattern and lake characteristics were found to differ significantly between regions. Three glacial lakes, from different geographic and climatic regions within the Indian Himalayas were then selected for a detailed risk assessment. Lake outburst probability, potential outburst magnitudes and associated damage were evaluated on the basis of high-resolution satellite imagery, field assessments and through the use of a dynamic model. The glacial lakes analyzed in the states of Jammu and Kashmir and Himachal Pradesh were found to present moderate risks to downstream villages, whereas the lake in Sikkim severely threatens downstream locations. At the study site in Sikkim, a dam breach could trigger drainage of ca. 16×10(6)m(3) water and generate maximum lake discharge of nearly 7000 m(3) s(-). The identification of critical glacial lakes in the Indian Himalayas and the detailed risk assessments at three specific sites allow prioritizing further investigations and help in the definition of risk reduction actions. Copyright © 2012 Elsevier B.V. All rights reserved.

  7. Isolated populations of the bush-cricket Pholidoptera frivaldszkyi (Orthoptera, Tettigoniidae) in Russia suggest a disjunct area of the species distribution.

    PubMed

    Kaňuch, Peter; Dorková, Martina; Mikhailenko, Andrey P; Polumordvinov, Oleg A; Jarčuška, Benjamín; Krištín, Anton

    2017-01-01

    Phylogenetic analysis and assessment of the species status of mostly isolated populations of Pholidoptera frivaldszkyi in south-western Russia occurring far beyond the accepted area of the species distribution in the Carpathian-Balkan region were performed. Using the mitochondrial DNA cytochrome c oxidase subunit I gene fragment, we found a very low level of genetic diversity in these populations. Phylogeographic reconstruction did not support recent introduction events but rather historical range fragmentation. The grouping of the Russian and Romanian haplotypes in a distinct phylogenetic clade suggests that the pre-glacial range of P. frivaldszkyi had extended towards the Ponto-Caspian region, with considerable gene flow between different refugia. However, post-glacial northward expansion of the species from supposed Caucasus refugia contributed most likely to the current disjunct distribution of this relict-like bush-cricket.

  8. Ireland, Shaded Relief and Colored Height

    NASA Technical Reports Server (NTRS)

    2005-01-01

    The island of Ireland comprises a large central lowland of limestone with a relief of hills surrounded by a discontinuous border of coastal mountains which vary greatly in geological structure. The mountain ridges of the south are composed of old red sandstone separated by limestone river valleys. Granite predominates in the mountains of Galway, Mayo and Donegal in the west and north-west and in Counties Down and Wicklow on the east coast, while a basalt plateau covers much of the north-east of the country. The central plain, which is broken in places by low hills, is extensively covered with glacial deposits of clay and sand. It has considerable areas of bog and numerous lakes. The island has seen at least two general glaciations and everywhere ice-smoothed rock, mountain lakes, glacial valleys and deposits of glacial sand, gravel and clay mark the passage of the ice.

    Two visualization methods were combined to produce this image: shading and color coding of topographic height. The shade image was derived by computing topographic slope in the northwest-southeast direction, so that northwest slopes appear bright and southeast slopes appear dark. Color coding is directly related to topographic height, with green at the lower elevations, rising through yellow and tan, to white at the highest elevations.

    Elevation data used in this image were acquired by the Shuttle Radar Topography Mission aboard the Space Shuttle Endeavour, launched on Feb. 11, 2000. SRTM used the same radar instrument that comprised the Spaceborne Imaging Radar-C/X-Band Synthetic Aperture Radar (SIR-C/X-SAR) that flew twice on the Space Shuttle Endeavour in 1994. SRTM was designed to collect 3-D measurements of the Earth's surface. To collect the 3-D data, engineers added a 60-meter (approximately 200-foot) mast, installed additional C-band and X-band antennas, and improved tracking and navigation devices. The mission is a cooperative project between NASA, the National Geospatial

  9. Morphology and palaeoenvironmental interpretation of deformed soft-sediment clasts: examples from within Late Pleistocene glacial outwash, Tempo Valley, Northern Ireland

    NASA Astrophysics Data System (ADS)

    Knight, Jasper

    1999-10-01

    Glacial outwash, deposited during deglaciation of the late Devensian ice sheet, is present as a flat-topped valley fill in the Tempo Valley on the southern flanks of the Fintona Hills, Northern Ireland. Sedimentologically, the outwash comprises well-sorted and interbedded rippled to massive sands which record distal deposition within a proglacial water body. Beds of ripple-drift cross-laminated sands contain deformed (folded and contorted) soft-sediment clasts which are composed mainly of silt and clay. The soft-sediment clasts were deformed prior to final deposition because clast a- b planes lie conformable to sand laminae which are undeformed. Morphological characteristics of the soft-sediment clasts, and their facies context, provide evidence for transport mechanisms, depositional environment, and processes of clast deformation. The soft-sediment clasts were transported into a proglacial water body by unidirectional water currents (˜1.5-2.5 m s -1). Sediment transport processes include sediment bypassing within the water column, a low bedload component, and grain flow activity during waning flow stages. The overall morphology of soft-sediment clasts records between 1 and 3 distinct phases of hydroplastic deformation prior to emplacement. The deformation phases are recognised on the basis of morphologically `unrolling' the superimposed folds of the soft-sediment clasts. Deformation structures (i.e. fold style) and direction of the principal stress axis relative to clast axes suggest that clasts were reoriented with respect to water flow direction following each deformation phase. Processes of deformation include folding-over of the clast along its b axis into two or more components, crumpling and abrasion of the outer margins of the b plane, and squashing of the clast c axis (some of which may be post-depositional deformation). The presence of silt- and clay-rich soft-sediment clasts within the outwash succession suggests that they were ripped-up from shallow

  10. Floristic similarity, diversity and endemism as indicators of refugia characteristics and needs in the West

    USGS Publications Warehouse

    Malanson, George P.; Zimmerman, Dale L.; Fagre, Daniel B.

    2015-01-01

    The floras of mountain ranges, and their similarity, beta diversity and endemism, are indicative of processes of community assembly; they are also the initial conditions for coming disassembly and reassembly in response to climate change. As such, these characteristics can inform thinking on refugia. The published floras or approximations for 42 mountain ranges in the three major mountain systems (Sierra-Cascades, Rocky Mountains and Great Basin ranges) across the western USA and southwestern Canada were analysed. The similarity is higher among the ranges of the Rockies while equally low among the ranges of the Sierra-Cascades and Great Basin. Mantel correlations of similarity with geographic distance are also higher for the Rocky Mountains. Endemism is relatively high, but is highest in the Sierra-Cascades (due to the Sierra Nevada as the single largest range) and lowest in the Great Basin, where assemblages are allochthonous. These differences indicate that the geologic substrates of the Cascade volcanoes, which are much younger than any others, play a role in addition to geographic isolation in community assembly. The pattern of similarity and endemism indicates that the ranges of the Cascades will not function well as stepping stones and the endemic species that they harbor may need more protection than those of the Rocky Mountains. The geometry of the ranges is complemented by geology in setting the stage for similarity and the potential for refugia across the West. Understanding the geographic template as initial conditions for the future can guide the forecast of refugia and related monitoring or protection efforts.

  11. Glacial flour in lacustrine sediments: Records of alpine glaciation in the western U.S.A. during the last glacial interval

    NASA Astrophysics Data System (ADS)

    Rosenbaum, J. G.; Reynolds, R. L.

    2010-12-01

    Sediments in Bear Lake (UT/ID) and Upper Klamath Lake (OR) contain glacial flour derived during the last glacial interval from the Uinta Mountains and the southern Cascade Range, respectively. Magnetic properties provide measures of glacial-flour content and, in concert with elemental and grain-size analyses, yield high-resolution records of glacial growth and decay. Creation and preservation of such records requires that (1) properties of glacial flour contrast with those of other sedimentary components and (2) magnetic minerals are neither formed nor destroyed after deposition. In the Bear Lake watershed, glaciers were confined to a small headwater area of the Bear River underlain by hematite-rich rocks of the Uinta Mountain Group (UMG), which are not exposed elsewhere in the catchment. Because UMG detritus is abundant only in Bear Lake sediments of glacial age, hard isothermal remanent magnetization (a measure of hematite content) provides a proxy for glacial flour. In contrast, the entire Upper Klamath Lake catchment, which lies to the east of the Cascade Range in southern Oregon, is underlain largely by basalt and basaltic andesite. Magnetic properties of fresh titanomagnetite-rich rock flour from glaciers on a composite volcano contrast sharply with those of detritus from unglaciated areas in which weathering destroyed some of the titanomagnetite. Ideally, well-dated records of the flux of glacial flour can be compared to ages of glacial features (e.g., moraines). For Upper Klamath Lake, quantitative measures of rock-flour content (from magnetic properties) and excellent chronology allow accurate calculation of flux. However, ages of glacial features are lacking and mafic volcanic rocks, which weather rapidly in this environment, are not well suited for cosmogenic exposure dating. At Bear Lake, estimates of glacial-flour content are less quantitative and chronology within the glacial interval must be interpolated from radiocarbon ages above and below the

  12. Eskers in Ireland, analogs for sinuous ridges on Mars

    NASA Astrophysics Data System (ADS)

    Pellicer, Xavier; Bourke, Mary

    2014-05-01

    Sinuous ridges on the surface of Mars are often inferred as putative esker ridges. Eskers cover several hundred kilometers of the Irish landscape and are one of the dominant landforms in the Irish Midlands. Well exposed stratigraphic sections and the body of existing knowledge due to extensive research carried out on these landforms make the Irish eskers an excellent analog for sinuous ridges on Mars. The Irish Eskers are sinuous ridges 0.1 - 80 km long, 20 - 500 m wide and 4 - 50 m high laid down by glacial meltwater in tunnels and crevasses in stationary or retreating ice sheets. They are commonly composed of sands and gravels with rounded boulders and cobbles. The gravels are usually bedded and the beds often slump towards the flank of the esker, indicating collapse as the confining ice walls melt. Four types of eskers have been identified in Ireland: (i) Continuous subglacial tunnel fill represents deposition within tunnels underneath or within an ice body originally used as water escape conduits; (ii) Continuous fluvial ice-channel fill deposit in channels cut into the ice on top of the glacier or down to the substrate subsequently infilled by sediments; (iii) Long beads - subglacial tunnel fill are segmented ridges, with a length-width ratio of 5:1 to 10:1, representing sequential deposition near or at the ice margin as the ice sheet retreats; (iv) Short beads are glaciolacustrine deposits interpreted as sequential deposition of ice-contact subaqueous outwash fans. Irish eskers have significant morphological similarities with those identified on Mars providing an opportunity for an insightful morphological and morphometric analysis to determine potential formative environments on Mars. Putative Martian eskers are 2-300 km long, 50-3000 m wide and 10-150 m high. The Irish eskers are similar in scale and present dimensions within these ranges. Eskers in Ireland are composed of sand and gravel with cobbles and boulders. Mars esker-like ridges observed in high

  13. How sea level change mediates genetic divergence in coastal species across regions with varying tectonic and sediment processes.

    PubMed

    Dolby, Greer A; Ellingson, Ryan A; Findley, Lloyd T; Jacobs, David K

    2018-02-01

    Plate tectonics and sediment processes control regional continental shelf topography. We examine the genetic consequences of how glacial-associated sea level change interacted with variable nearshore topography since the last glaciation. We reconstructed the size and distribution of areas suitable for tidal estuary formation from the last glacial maximum, ~20 thousand years ago, to present from San Francisco, California, USA (~38°N) to Reforma, Sinaloa, Mexico (~25°N). We assessed range-wide genetic structure and diversity of three codistributed tidal estuarine fishes (California Killifish, Shadow Goby, Longjaw Mudsucker) along ~4,600 km using mitochondrial control region and cytB sequence, and 16-20 microsatellite loci from a total of 524 individuals. Results show that glacial-associated sea level change limited estuarine habitat to few, widely separated refugia at glacial lowstand, and present-day genetic clades were sourced from specific refugia. Habitat increased during postglacial sea level rise and refugial populations admixed in newly formed habitats. Continental shelves with active tectonics and/or low sediment supply were steep and hosted fewer, smaller refugia with more genetically differentiated populations than on broader shelves. Approximate Bayesian computation favoured the refuge-recolonization scenarios from habitat models over isolation by distance and seaway alternatives, indicating isolation at lowstand is a major diversification mechanism among these estuarine (and perhaps other) coastal species. Because sea level change is a global phenomenon, we suggest this top-down physical control of extirpation-isolation-recolonization may be an important driver of genetic diversification in coastal taxa inhabiting other topographically complex coasts globally during the Mid- to Late Pleistocene and deeper timescales. © 2018 John Wiley & Sons Ltd.

  14. Relative timing of last glacial maximum and late-glacial events in the central tropical Andes

    NASA Astrophysics Data System (ADS)

    Bromley, Gordon R. M.; Schaefer, Joerg M.; Winckler, Gisela; Hall, Brenda L.; Todd, Claire E.; Rademaker, Kurt M.

    2009-11-01

    Whether or not tropical climate fluctuated in synchrony with global events during the Late Pleistocene is a key problem in climate research. However, the timing of past climate changes in the tropics remains controversial, with a number of recent studies reporting that tropical ice age climate is out of phase with global events. Here, we present geomorphic evidence and an in-situ cosmogenic 3He surface-exposure chronology from Nevado Coropuna, southern Peru, showing that glaciers underwent at least two significant advances during the Late Pleistocene prior to Holocene warming. Comparison of our glacial-geomorphic map at Nevado Coropuna to mid-latitude reconstructions yields a striking similarity between Last Glacial Maximum (LGM) and Late-Glacial sequences in tropical and temperate regions. Exposure ages constraining the maximum and end of the older advance at Nevado Coropuna range between 24.5 and 25.3 ka, and between 16.7 and 21.1 ka, respectively, depending on the cosmogenic production rate scaling model used. Similarly, the mean age of the younger event ranges from 10 to 13 ka. This implies that (1) the LGM and the onset of deglaciation in southern Peru occurred no earlier than at higher latitudes and (2) that a significant Late-Glacial event occurred, most likely prior to the Holocene, coherent with the glacial record from mid and high latitudes. The time elapsed between the end of the LGM and the Late-Glacial event at Nevado Coropuna is independent of scaling model and matches the period between the LGM termination and Late-Glacial reversal in classic mid-latitude records, suggesting that these events in both tropical and temperate regions were in phase.

  15. Ireland

    ERIC Educational Resources Information Center

    Kail, Candice

    2010-01-01

    Ireland has a rich and long history. It is a land of fable and of strife, from the legendary warrior Fionn Mac Cumhaill (anglicized as Finn McCool) and the god-like Tuatha De Danann to the potato famine and the more recent Troubles. In the last decade, Ireland has experienced an economic boom and assumed a new place in the political landscape via…

  16. Regional patterns of postglacial changes in the Palearctic mammalian diversity indicate retreat to Siberian steppes rather than extinction

    PubMed Central

    Řičánková, Věra Pavelková; Robovský, Jan; Riegert, Jan; Zrzavý, Jan

    2015-01-01

    We examined the presence of possible Recent refugia of Pleistocene mammalian faunas in Eurasia by analysing regional differences in the mammalian species composition, occurrence and extinction rates between Recent and Last Glacial faunas. Our analyses revealed that most of the widespread Last Glacial species have survived in the central Palearctic continental regions, most prominently in Altai–Sayan (followed by Kazakhstan and East European Plain). The Recent Altai–Sayan and Kazakhstan regions show species compositions very similar to their Pleistocene counterparts. The Palearctic regions have lost 12% of their mammalian species during the last 109,000 years. The major patterns of the postglacial changes in Palearctic mammalian diversity were not extinctions but rather radical shifts of species distribution ranges. Most of the Pleistocene mammalian fauna retreated eastwards, to the central Eurasian steppes, instead of northwards to the Arctic regions, considered Holocene refugia of Pleistocene megafauna. The central Eurasian Altai and Sayan mountains could thus be considered a present-day refugium of the Last Glacial biota, including mammals. PMID:26246136

  17. Phylogeography of a tough rock survivor in European dry grasslands

    PubMed Central

    Poschlod, Peter; Reisch, Christoph

    2017-01-01

    Phylogeographic analyses of plants in Europe have revealed common glacial refugia and migration routes for several trees and herbs with arctic-alpine distributions. The postglacial histories of dry grassland species in central Europe have rarely been analyzed, even though the extremely species-rich habitat is threatened. Sedum album (Crassulaceae) is a common inhabitant of rocky sites in central European dry grasslands. We inferred the phylogeographic history of S. album over its distribution range in Europe. Genetic diversity within and differentiation between 34 S. album populations was examined using AFLP markers. Population isolation was indicated based on the rarity of the fragments and by isolation-by-distance effects. We sequenced the trnL-trnF region in 32 populations and used chloroplast microsatellites to analyze chloroplast haplotype distributions. Two distinct S. album lineages were detected. One lineage was comprised of populations from eastern and central parts of central Europe, and the Apennine Peninsula. A second lineage was comprised of populations from the Iberian Peninsula and western and northern parts of central Europe. Glacial refugia were identified based on the accumulation of ancient chloroplast haplotypes, high diversity of AFLP fragments within populations, and high levels of rare fragments in Liguria, Serbia, the Apennine and Iberian peninsulas. Cryptic refugia were detected in the Czech Republic and Slovakia. Isolation by distance was present all over the distribution range, and it was separately detected in southwestern and central Europe. In western Europe, where a contact zone between the two lineages can be expected, no isolation by distance was detected. Our results suggest migration routes of S. album northeastward from glacial refugia in southern Iberia, northward from the Apennine Peninsula, and northward and westward from the southeastern parts of central Europe. Therefore, central European grasslands were recently colonized

  18. Contrasting scaling properties of interglacial and glacial climates

    PubMed Central

    Shao, Zhi-Gang; Ditlevsen, Peter D.

    2016-01-01

    Understanding natural climate variability is essential for assessments of climate change. This is reflected in the scaling properties of climate records. The scaling exponents of the interglacial and the glacial climates are fundamentally different. The Holocene record is monofractal, with a scaling exponent H∼0.7. On the contrary, the glacial record is multifractal, with a significantly higher scaling exponent H∼1.2, indicating a longer persistence time and stronger nonlinearities in the glacial climate. The glacial climate is dominated by the strong multi-millennial Dansgaard–Oeschger (DO) events influencing the long-time correlation. However, by separately analysing the last glacial maximum lacking DO events, here we find the same scaling for that period as for the full glacial period. The unbroken scaling thus indicates that the DO events are part of the natural variability and not externally triggered. At glacial time scales, there is a scale break to a trivial scaling, contrasting the DO events from the similarly saw-tooth-shaped glacial cycles. PMID:26980084

  19. Resolving Large Pre-glacial Valleys Buried by Glacial Sediment Using Electric Resistivity Imaging (ERI)

    NASA Astrophysics Data System (ADS)

    Schmitt, D. R.; Welz, M.; Rokosh, C. D.; Pontbriand, M.-C.; Smith, D. G.

    2004-05-01

    Two-dimensional electric resistivity imaging (ERI) is the most exciting and promising geological tool in geomorphology and stratigraphy since development of ground-penetrating radar. Recent innovations in 2-D ERI provides a non-intrusive mean of efficiently resolving complex shallow subsurface structures under a number of different geological scenarios. In this paper, we test the capacity of ERI to image two large pre-late Wisconsinan-aged valley-fills in central Alberta and north-central Montana. Valley-fills record the history of pre-glacial and glacial sedimentary deposits. These fills are of considerable economical value as groundwater aquifers, aggregate resources (sand and gravel), placers (gold, diamond) and sometime gas reservoirs in Alberta. Although the approximate locations of pre-glacial valley-fills have been mapped, the scarcity of borehole (well log) information and sediment exposures make accurate reconstruction of their stratigraphy and cross-section profiles difficult. When coupled with borehole information, ERI successfully imaged three large pre-glacial valley-fills representing three contrasting geological settings. The Sand Coulee segment of the ancestral Missouri River, which has never been glaciated, is filled by electrically conductive pro-glacial lacustrine deposits over resistive sandstone bedrock. By comparison, the Big Sandy segment of the ancestral Missouri River valley has a complex valley-fill composed of till units interbedded with glaciofluvial gravel and varved clays over conductive shale. The fill is capped by floodplain, paludal and low alluvial fan deposits. The pre-glacial Onoway Valley (the ancestral North Saskatchewan River valley) is filled with thick, resistive fluvial gravel over conductive shale and capped with conductive till. The cross-sectional profile of each surveyed pre-glacial valley exhibits discrete benches (terraces) connected by steep drops, features that are hard to map using only boreholes. Best quality ERI

  20. When prey provide more than food: mammalian predators appropriating the refugia of their prey

    Treesearch

    Bill Zielinski

    2015-01-01

    Some mammalian predators acquire both food and shelter from their prey, by eating them and using the refugia the prey construct. I searched the literature for examples of predators that exhibit this behavior and summarize their taxonomic affiliations, relative sizes, and distributions. I hypothesized that size ratios of species involved in this dynamic would be near 1....

  1. Isolated populations of the bush-cricket Pholidoptera frivaldszkyi (Orthoptera, Tettigoniidae) in Russia suggest a disjunct area of the species distribution

    PubMed Central

    Kaňuch, Peter; Dorková, Martina; Mikhailenko, Andrey P.; Polumordvinov, Oleg A.; Jarčuška, Benjamín; Krištín, Anton

    2017-01-01

    Abstract Phylogenetic analysis and assessment of the species status of mostly isolated populations of Pholidoptera frivaldszkyi in south-western Russia occurring far beyond the accepted area of the species distribution in the Carpathian-Balkan region were performed. Using the mitochondrial DNA cytochrome c oxidase subunit I gene fragment, we found a very low level of genetic diversity in these populations. Phylogeographic reconstruction did not support recent introduction events but rather historical range fragmentation. The grouping of the Russian and Romanian haplotypes in a distinct phylogenetic clade suggests that the pre-glacial range of P. frivaldszkyi had extended towards the Ponto-Caspian region, with considerable gene flow between different refugia. However, post-glacial northward expansion of the species from supposed Caucasus refugia contributed most likely to the current disjunct distribution of this relict-like bush-cricket. PMID:28769628

  2. Controls on subglacial patterns and depositional environments in western Ireland

    NASA Astrophysics Data System (ADS)

    Knight, J.

    2009-12-01

    In western Ireland, Late Devensian ice flow dynamics and resultant patterns of landforms and sediments reflect the interplay between internal (glaciological) forcing and external forcing by rapid climate changes centred on the adjacent Atlantic Ocean. This interplay can be best demonstrated where ice from climatically-sensitive mountain source regions flowed into surrounding lowlands, such as the Connemara region of west County Galway, western Ireland. Here, a semi-independent ice cap was present over the Twelve Bens mountains, and interacted with ice from the much larger regional ice sheet from central Ireland. Landform and sediment patterns in the flat lowland region (c. 100 km2 below 30 m asl) to the south of the Twelve Bens reflect elements of this ice interaction. In detail, landform and sediment distributions here are highly complex with marked spatial differences in patterns of sediment availability. Across much of the region, sculpted bedrock forms (whaleback and bedrock drumlin ridges, roches mountonnées, striae) reflect subglacial abrasion across the underlying igneous and metamorphic bedrock that forms a relatively flat and lake-dominated landscape. Glacigenic sediments are found only at or around ice-retreat margins, and within isolated bedrock valleys. Here, diamicton drumlins are relatively uncommon but yet must represent depositional conditions that are not reflected elsewhere in this ice sheet sector where subglacial sediments are generally absent. This paper explores the interrelationship between local and regional ice flows through their impact on spatial patterns of glacial landforms and sediments. The paper presents field data on the characteristics of bedrock forms (erosional) and diamicton drumlins (depositional). Subglacial sediments are described from drumlin outcrops at key sites around Connemara, which helps in the understanding of the evolution of the subglacial environment in response to ice interactions from different source regions.

  3. Shallow bedrock limits groundwater seepage-based headwater climate refugia

    USGS Publications Warehouse

    Briggs, Martin A.; Lane, John W.; Snyder, Craig D.; White, Eric A.; Johnson, Zachary; Nelms, David L.; Hitt, Nathaniel P.

    2018-01-01

    Groundwater/surface-water exchanges in streams are inexorably linked to adjacent aquifer dynamics. As surface-water temperatures continue to increase with climate warming, refugia created by groundwater connectivity is expected to enable cold water fish species to survive. The shallow alluvial aquifers that source groundwater seepage to headwater streams, however, may also be sensitive to seasonal and long-term air temperature dynamics. Depth to bedrock can directly influence shallow aquifer flow and thermal sensitivity, but is typically ill-defined along the stream corridor in steep mountain catchments. We employ rapid, cost-effective passive seismic measurements to evaluate the variable thickness of the shallow colluvial and alluvial aquifer sediments along a headwater stream supporting cold water-dependent brook trout (Salvelinus fontinalis) in Shenandoah National Park, VA, USA. Using a mean depth to bedrock of 2.6 m, numerical models predicted strong sensitivity of shallow aquifer temperature to the downward propagation of surface heat. The annual temperature dynamics (annual signal amplitude attenuation and phase shift) of potential seepage sourced from the shallow modeled aquifer were compared to several years of paired observed stream and air temperature records. Annual stream water temperature patterns were found to lag local air temperature by ∼8–19 d along the stream corridor, indicating that thermal exchange between the stream and shallow groundwater is spatially variable. Locations with greater annual signal phase lag were also associated with locally increased amplitude attenuation, further suggestion of year-round buffering of channel water temperature by groundwater seepage. Numerical models of shallow groundwater temperature that incorporate regional expected climate warming trends indicate that the summer cooling capacity of this groundwater seepage will be reduced over time, and lower-elevation stream sections may no longer serve as larger

  4. Contrasting scaling properties of interglacial and glacial climates

    NASA Astrophysics Data System (ADS)

    Ditlevsen, Peter; Shao, Zhi-Gang

    2017-04-01

    Understanding natural climate variability is essential for assessments of climate change. This is reflected in the scaling properties of climate records. The scaling exponents of the interglacial and the glacial climates are fundamentally different. The Holocene record is monofractal, with a scaling exponent H˜0.7. On the contrary, the glacial record is multifractal, with a significantly higher scaling exponent H˜1.2, indicating a longer persistence time and stronger nonlinearities in the glacial climate. The glacial climate is dominated by the strong multi-millennial Dansgaard-Oeschger (DO) events influencing the long-time correlation. However, by separately analysing the last glacial maximum lacking DO events, here we find the same scaling for that period as for the full glacial period. The unbroken scaling thus indicates that the DO events are part of the natural variability and not externally triggered. At glacial time scales, there is a scale break to a trivial scaling, contrasting the DO events from the similarly saw-tooth-shaped glacial cycles. Ref: Zhi-Gang Shao and Peter Ditlevsen, Nature Comm. 7, 10951, 2016

  5. Amazonian and neotropical plant communities on glacial time-scales: The failure of the aridity and refuge hypotheses

    NASA Astrophysics Data System (ADS)

    Colinvaux, P. A.; De Oliveira, P. E.; Bush, M. B.

    2000-01-01

    Plants respond to Pleistocene climatic change as species, not as associations or biomes. This has been demonstrated unequivocally by paleobotanical data for temperate latitudes. In the far richer vegetations of the tropics species populations also fluctuated independently in response to climatic forcing, from their longlasting glacial states to the patterns of brief interglacials like the present and back again. We use pollen data to reconstruct the vegetation of the Amazon basin in oxygen isotope stages 3 and 2 of the last glaciation in order to measure how the plant populations of the Amazon responded to the global warming at the onset of the Holocene. We find that plant communities of the neotropics vent copious pollen to lake sediments and that this pollen yields powerful signals for community composition. Three continuous sedimentary records reaching through oxygen isotope stage 2 are available from the Amazon lowlands, those from Carajas, Lake Pata and marine deposits off the mouth of the Amazon River. All three records yield pollen histories of remarkable constancy and stability. By comparing them with deposits of equal antiquity from the cerrado (savanna) of central Brazil, we show that most of the Amazon lowlands remained under forest throughout a glacial cycle. This forest was never fragmented by open vegetation as postulated by the refugia hypothesis. Instead the intact forest of glacial times included significant populations of plants that are now montane, suggesting that the global warming of the early Holocene resulted in the expulsion of heat intolerant plants from the lowland forest. Pollen data from the Amazonian flank of the Andes and from Pacific Panama provide evidence that populations of these heat intolerant plants survive the heat of interglacials in part by maintaining large populations at cooler montane altitudes. Our conclusion that the Amazon lowlands were forested in glacial times specifically refutes the hypothesis of Amazonian glacial

  6. Holocene glacial fluctuations in southern South America

    NASA Astrophysics Data System (ADS)

    Reynhout, S.; Sagredo, E. A.; Kaplan, M. R.; Aravena, J. C.; Martini, M. A.; Strelin, J. A.; Schaefer, J. M.

    2016-12-01

    Understanding the timing and magnitude of former glacier fluctuations is critical to decipher long-term climatic trends and to unravel both natural cycles and human impact on the current glacial behavior. Despite more than seven decades of research efforts, a unifying model of Holocene glacial fluctuations in Southern South America remains elusive. Here, we present the state-of-the-art regarding the timing of Holocene glacial fluctuation in southern Patagonia-Tierra del Fuego, with a focus on a new generation of high-resolution radiocarbon and 10Be surface exposure dating chronologies. Recently acquired evidence suggest that after receding from advanced Late Glacial positions, Patagonian glaciers were for the most part close to, or even behind, present ice margins during the Early Holocene. On the other hand, emerging chronologies indicate that in some areas there were extensive expansions (century scale?) that punctuated the warm interval. Subsequently, we have evidence of multiple millennial timescale glacial advances starting in the middle Holocene. Several glacial maxima are defined by moraines and other landforms from 7000 years ago to the 19th century, with a gap sometime between 4,500 and 2,500 years ago. The last set of advances began around 800-600 years ago. Although glacial activity is documented in Patagonia at the same time as the European Little Ice Age, the extent of these glacial events are less prominent than those of the mid-Holocene. The causes that may explain these glacial fluctuations remain elusive. Finally, we discuss ongoing efforts to better define the timing and extent of Holocene glaciations in southern South America, and to establish the basis to test competing hypothesis of regional Holocene climate variability.

  7. Light attenuation characteristics of glacially-fed lakes

    NASA Astrophysics Data System (ADS)

    Rose, Kevin C.; Hamilton, David P.; Williamson, Craig E.; McBride, Chris G.; Fischer, Janet M.; Olson, Mark H.; Saros, Jasmine E.; Allan, Mathew G.; Cabrol, Nathalie

    2014-07-01

    Transparency is a fundamental characteristic of aquatic ecosystems and is highly responsive to changes in climate and land use. The transparency of glacially-fed lakes may be a particularly sensitive sentinel characteristic of these changes. However, little is known about the relative contributions of glacial flour versus other factors affecting light attenuation in these lakes. We sampled 18 glacially-fed lakes in Chile, New Zealand, and the U.S. and Canadian Rocky Mountains to characterize how dissolved absorption, algal biomass (approximated by chlorophyll a), water, and glacial flour contributed to attenuation of ultraviolet radiation (UVR) and photosynthetically active radiation (PAR, 400-700 nm). Variation in attenuation across lakes was related to turbidity, which we used as a proxy for the concentration of glacial flour. Turbidity-specific diffuse attenuation coefficients increased with decreasing wavelength and distance from glaciers. Regional differences in turbidity-specific diffuse attenuation coefficients were observed in short UVR wavelengths (305 and 320 nm) but not at longer UVR wavelengths (380 nm) or PAR. Dissolved absorption coefficients, which are closely correlated with diffuse attenuation coefficients in most non-glacially-fed lakes, represented only about one quarter of diffuse attenuation coefficients in study lakes here, whereas glacial flour contributed about two thirds across UVR and PAR. Understanding the optical characteristics of substances that regulate light attenuation in glacially-fed lakes will help elucidate the signals that these systems provide of broader environmental changes and forecast the effects of climate change on these aquatic ecosystems.

  8. An improved active contour model for glacial lake extraction

    NASA Astrophysics Data System (ADS)

    Zhao, H.; Chen, F.; Zhang, M.

    2017-12-01

    Active contour model is a widely used method in visual tracking and image segmentation. Under the driven of objective function, the initial curve defined in active contour model will evolve to a stable condition - a desired result in given image. As a typical region-based active contour model, C-V model has a good effect on weak boundaries detection and anti noise ability which shows great potential in glacial lake extraction. Glacial lake is a sensitive indicator for reflecting global climate change, therefore accurate delineate glacial lake boundaries is essential to evaluate hydrologic environment and living environment. However, the current method in glacial lake extraction mainly contains water index method and recognition classification method are diffcult to directly applied in large scale glacial lake extraction due to the diversity of glacial lakes and masses impacted factors in the image, such as image noise, shadows, snow and ice, etc. Regarding the abovementioned advantanges of C-V model and diffcults in glacial lake extraction, we introduce the signed pressure force function to improve the C-V model for adapting to processing of glacial lake extraction. To inspect the effect of glacial lake extraction results, three typical glacial lake development sites were selected, include Altai mountains, Centre Himalayas, South-eastern Tibet, and Landsat8 OLI imagery was conducted as experiment data source, Google earth imagery as reference data for varifying the results. The experiment consequence suggests that improved active contour model we proposed can effectively discriminate the glacial lakes from complex backgound with a higher Kappa Coefficient - 0.895, especially in some small glacial lakes which belongs to weak information in the image. Our finding provide a new approach to improved accuracy under the condition of large proportion of small glacial lakes and the possibility for automated glacial lake mapping in large-scale area.

  9. Regional and Local Glacial-Earthquake Patterns in Greenland

    NASA Astrophysics Data System (ADS)

    Olsen, K.; Nettles, M.

    2016-12-01

    Icebergs calved from marine-terminating glaciers currently account for up to half of the 400 Gt of ice lost annually from the Greenland ice sheet (Enderlin et al., 2014). When large capsizing icebergs ( 1 Gt of ice) calve, they produce elastic waves that propagate through the solid earth and are observed as teleseismically detectable MSW 5 glacial earthquakes (e.g., Ekström et al., 2003; Nettles & Ekström, 2010 Tsai & Ekström, 2007; Veitch & Nettles, 2012). The annual number of these events has increased dramatically over the past two decades. We analyze glacial earthquakes from 2011-2013, which expands the glacial-earthquake catalog by 50%. The number of glacial-earthquake solutions now available allows us to investigate regional patterns across Greenland and link earthquake characteristics to changes in ice dynamics at individual glaciers. During the years of our study Greenland's west coast dominated glacial-earthquake production. Kong Oscar Glacier, Upernavik Isstrøm, and Jakobshavn Isbræ all produced more glacial earthquakes during this time than in preceding years. We link patterns in glacial-earthquake production and cessation to the presence or absence of floating ice tongues at glaciers on both coasts of Greenland. The calving model predicts glacial-earthquake force azimuths oriented perpendicular to the calving front, and comparisons between seismic data and satellite imagery confirm this in most instances. At two glaciers we document force azimuths that have recently changed orientation and confirm that similar changes have occurred in the calving-front geometry. We also document glacial earthquakes at one previously quiescent glacier. Consistent with previous work, we model the glacial-earthquake force-time function as a boxcar with horizontal and vertical force components that vary synchronously. We investigate limitations of this approach and explore improvements that could lead to a more accurate representation of the glacial earthquake source.

  10. Environmental and climatic conditions at a potential Glacial refugial site of tree species near the Southern Alpine glaciers. New insights from multiproxy sedimentary studies at Lago della Costa (Euganean Hills, Northeastern Italy)

    NASA Astrophysics Data System (ADS)

    Kaltenrieder, Petra; Belis, Claudio A.; Hofstetter, Simone; Ammann, Brigitta; Ravazzi, Cesare; Tinner, Willy

    2009-12-01

    It has been hypothesized that refugia of thermophilous tree species were located in Northern Italy very close to the Alps, though, this hypothesis has yet to be tested thoroughly. In contrast to Central and Southern Italy with its relative wealth of data, only a few fragmentary records are currently available from Northern Italy for the last Glacial (Würm, Weichselian). Our new study site Lago della Costa lies adjacent to the catchment of the megafans of the Alpine forelands and the braided rivers of the Northeastern Po Plain that have so far inhibited the recovery of continuous Glacial and Late-Glacial records. We analyze pollen, plant macrofossils, charcoal and ostracods to reconstruct the vegetation, fire and lake history for the period 33,000-16,000 cal. BP. We compare our data with Glacial records from Southern Europe to discuss similarities and dissimilarities between these potential refugial areas. A comparison with independent paleoclimatic proxies allows to assess potential linkages between environmental and climatic variability. New macrofossil and pollen data at Lago della Costa unambiguously document the local persistence of boreal tree taxa such as Larix decidua and Betula tree species around the study site during the last Glacial. The regular occurrence of pollen of temperate trees in the organic lake sediments (fine-detritus calcareous gyttja) suggests that temperate taxa such as Corylus avellana, Quercus deciduous, Tilia, Ulmus, Fraxinus excelsior, Carpinus, Abies alba and Fagus sylvatica, most likely survived the Last Glacial Maximum (LGM) at favorable sites in the Euganean Hills. The percentage values of temperate trees are comparable with those from Southern Europe (e.g. Monticchio in Southern Italy). We conclude that the Euganean Hills were one of the northernmost refugial areas of temperate taxa in Europe. However, the relative and absolute abundances of pollen of temperate trees are highly variable. Pollen-inferred declines of temperate tree

  11. Investigating the effects of Pleistocene events on genetic divergence within Richardsonius balteatus, a widely distributed western North American minnow

    PubMed Central

    2014-01-01

    Background Biogeographers seek to understand the influences of global climate shifts and geologic changes to the landscape on the ecology and evolution of organisms. Across both longer and shorter timeframes, the western North American landscape has experienced dynamic transformations related to various geologic processes and climatic oscillations, including events as recently as the Last Glacial Maximum (LGM; ~20 Ka) that have impacted the evolution of the North American biota. Redside shiner is a cyprinid species that is widely distributed throughout western North America. The species’ native range includes several well-documented Pleistocene refugia. Here we use mitochondrial DNA sequence data to assess phylogeography, and to test two biogeographic hypotheses regarding post-glacial colonization by redside shiner: 1) Redside shiner entered the Bonneville Basin at the time of the Bonneville Flood (Late Pleistocene; 14.5 Ka), and 2) redside shiner colonized British Columbia post-glacially from a single refugium in the Upper Columbia River drainage. Results Genetic diversification in redside shiner began in the mid to late Pleistocene, but was not associated with LGM. Different clades of redside shiner were distributed in multiple glacial age refugia, and each clade retains a signature of population expansion, with clades having secondary contact in some areas. Conclusions Divergence times between redside shiner populations in the Bonneville Basin and the Upper Snake/Columbia River drainage precedes the Bonneville Flood, thus it is unlikely that redside shiner invaded the Bonneville Basin during this flooding event. All but one British Columbia population of redside shiner are associated with the Upper Columbia River drainage with the lone exception being a population near the coast, suggesting that the province as a whole was colonized from multiple refugia, but the inland British Columbia redside shiner populations are affiliated with a refugium in the Upper

  12. Inferring Multiple Refugia and Phylogeographical Patterns in Pinus massoniana Based on Nucleotide Sequence Variation and DNA Fingerprinting

    PubMed Central

    Lin, Chung-Jian; Huang, Chi-Chung; Huang, Chao-Ching; Chiang, Yu-Chung; Chiang, Tzen-Yuh

    2012-01-01

    Background Pinus massoniana, an ecologically and economically important conifer, is widespread across central and southern mainland China and Taiwan. In this study, we tested the central–marginal paradigm that predicts that the marginal populations tend to be less polymorphic than the central ones in their genetic composition, and examined a founders' effect in the island population. Methodology/Principal Findings We examined the phylogeography and population structuring of the P. massoniana based on nucleotide sequences of cpDNA atpB-rbcL intergenic spacer, intron regions of the AdhC2 locus, and microsatellite fingerprints. SAMOVA analysis of nucleotide sequences indicated that most genetic variants resided among geographical regions. High levels of genetic diversity in the marginal populations in the south region, a pattern seemingly contradicting the central–marginal paradigm, and the fixation of private haplotypes in most populations indicate that multiple refugia may have existed over the glacial maxima. STRUCTURE analyses on microsatellites revealed that genetic structure of mainland populations was mediated with recent genetic exchanges mostly via pollen flow, and that the genetic composition in east region was intermixed between south and west regions, a pattern likely shaped by gene introgression and maintenance of ancestral polymorphisms. As expected, the small island population in Taiwan was genetically differentiated from mainland populations. Conclusions/Significance The marginal populations in south region possessed divergent gene pools, suggesting that the past glaciations might have low impacts on these populations at low latitudes. Estimates of ancestral population sizes interestingly reflect a recent expansion in mainland from a rather smaller population, a pattern that seemingly agrees with the pollen record. PMID:22952747

  13. Glaciers, Glacial lakes and Glacial Lake Outburst Floods in the Koshi Basin

    NASA Astrophysics Data System (ADS)

    Shrestha, F.; Gao, X.; Khanal, N. R.; Maharjan, S. B.; Bajracharya, S. R.; Shrestha, R. B.; Lizong, W.; Mool, P. K.

    2016-12-01

    Glacier is a vital water resources for mountain communities. Recession in glacier area either increased the glacial lake size or develop a new lake. The consequences of these changes in lake has become one of the major issue in the management of GLOF risk. This paper presents the distribution of, and changes in, glaciers, glacial lakes in the Koshi basin and also looks at past GLOF events that have occurred in the basin and their distance of impact. Data on the number of glaciers and glacial lakes and their areas were generated for the years 1977, 1990, 2000, and 2010 using Landsat images. The study revealed that there were a total of 845 glaciers (Nepal side) and 2,168 glacial lakes (Nepal and China side) with a total area of 1,103 km2 and 127.608 km2 in 2010. The number of glacier increased by 15% (109) and area decreased by 26% (396 km2) over 33 years. In case of glacier lakes, the number and area increased from 1,160 to 2,168 and from 94.444 km2 to 127.608 km2 during 33 years with an overall growth rates of 86.9% and 35.1%. A large number of glacial lakes are small in size (≤ 0.1 km2). End moraine dammed lakes with area ≥ 0.1 km2 were selected to analyse the change characteristics of glacial lakes. The results show that there were 134 lakes ≥ 0.1 km2 in 2010; these lakes had a total area of 43.06 km2 in 1997, increased to 64.35 km2 in 2010. The distribution of lakes on the north side of the Himalayas (in China) was three times higher than on the south side of the Himalayas (in Nepal). Comparing the mean growth rate in area and length for the 33 years, the growth rate on the north side was found to be a little slower than that on the south side. This relationship did not hold true for length change in the different periods. The study identified 42 rapidly growing large lakes that are dangerous in terms of GLOF risk. In the past, 18 GLOF events have been reported. The downstream distance impacted by those events was up to 90 km. Among them, 13 GLOF events

  14. Ireland

    NASA Image and Video Library

    2017-12-08

    On August 7, 2003, the NASA Aqua MODIS instrument acquired this image of Ireland on the first day this summer that most of the island hasn´t been completely obscured by cloud cover. Called the Emerald Isle for a good reason, Ireland is draped in vibrant shades of green amidst the blue Atlantic Ocean and Celtic (south) and Irish (east) Seas. Faint ribbons of blue-green phytoplankton drift in the waters of the Celtic Sea, just south of Dublin. Dublin itself appears as a large grayish-brown spot on the Republic of Ireland´s northeastern coast. This large capital city (population 1.12 million) sits on the River Liffey, effectively splitting the city in half. Northern Ireland´s capital city, Belfast, also sits on a river: the River Lagan. This city, though its population is only a fifth of the size of Dublin´s, is also clearly visible in the image as a grayish-brown spot on the coast of the Irish Sea. Sensor Aqua/MODIS Credit Jeff Schmaltz, MODIS Rapid Response Team, NASA/GSFC For more information go to: visibleearth.nasa.gov/view_rec.php?id=5744 NASA Goddard Space Flight Center is home to the nation's largest organization of combined scientists, engineers and technologists that build spacecraft, instruments and new technology to study the Earth, the sun, our solar system, and the universe. Follow us on Twitter Join us on Facebook

  15. Pleistocene refugia and polytopic replacement of diploids by tetraploids in the Patagonian and Subantarctic plant Hypochaeris incana (Asteraceae, Cichorieae).

    PubMed

    Tremetsberger, Karin; Urtubey, Estrella; Terrab, Anass; Baeza, Carlos M; Ortiz, María Angeles; Talavera, María; König, Christiane; Temsch, Eva M; Kohl, Gudrun; Talavera, Salvador; Stuessy, Tod F

    2009-09-01

    We report the phylogeographic pattern of the Patagonian and Subantarctic plant Hypochaeris incana endemic to southeastern South America. We applied amplified fragment length polymorphism (AFLP) and chloroplast DNA (cpDNA) analysis to 28 and 32 populations, respectively, throughout its distributional range and assessed ploidy levels using flow cytometry. While cpDNA data suggest repeated or simultaneous parallel colonization of Patagonia and Tierra del Fuego by several haplotypes and/or hybridization, AFLPs reveal three clusters corresponding to geographic regions. The central and northern Patagonian clusters (approximately 38-51 degrees S), which are closer to the outgroup, contain mainly tetraploid, isolated and highly differentiated populations with low genetic diversity. To the contrary, the southern Patagonian and Fuegian cluster (approximately 51-55 degrees S) contains mainly diploid populations with high genetic diversity and connected by high levels of gene flow. The data suggest that H. incana originated at the diploid level in central or northern Patagonia, from where it migrated south. All three areas, northern, central and southern, have similar levels of rare and private AFLP bands, suggesting that all three served as refugia for H. incana during glacial times. In southern Patagonia and Tierra del Fuego, the species seems to have expanded its populational system in postglacial times, when the climate became warmer and more humid. In central and northern Patagonia, the populations seem to have become restricted to favourable sites with increasing temperature and decreasing moisture and there was a parallel replacement of diploids by tetraploids in local populations.

  16. Predicted changes in climatic niche and climate refugia of conservation priority salamander species in the northeastern United States

    USGS Publications Warehouse

    Sutton, William B.; Barrett, Kyle; Moody, Allison T.; Loftin, Cynthia S.; deMaynadier, Phillip G.; Nanjappa, Priya

    2015-01-01

    Global climate change represents one of the most extensive and pervasive threats to wildlife populations. Amphibians, specifically salamanders, are particularly susceptible to the effects of changing climates due to their restrictive physiological requirements and low vagility; however, little is known about which landscapes and species are vulnerable to climate change. Our study objectives included, (1) evaluating species-specific predictions (based on 2050 climate projections) and vulnerabilities to climate change and (2) using collective species responses to identify areas of climate refugia for conservation priority salamanders in the northeastern United States. All evaluated salamander species were projected to lose a portion of their climatic niche. Averaged projected losses ranged from 3%–100% for individual species, with the Cow Knob Salamander (Plethodon punctatus), Cheat Mountain Salamander (Plethodon nettingi), Shenandoah Mountain Salamander (Plethodon virginia), Mabee’s Salamander (Ambystoma mabeei), and Streamside Salamander (Ambystoma barbouri) predicted to lose at least 97% of their landscape-scale climatic niche. The Western Allegheny Plateau was predicted to lose the greatest salamander climate refugia richness (i.e., number of species with a climatically-suitable niche in a landscape patch), whereas the Central Appalachians provided refugia for the greatest number of species during current and projected climate scenarios. Our results can be used to identify species and landscapes that are likely to be further affected by climate change and potentially resilient habitats that will provide consistent climatic conditions in the face of environmental change.

  17. Sources of glacial moisture in Mesoamerica

    USGS Publications Warehouse

    Bradbury, J.P.

    1997-01-01

    Paleoclimatic records from Mesoamerica document the interplay between Atlantic and Pacific sources of precipitation during the last glacial stage and Holocene. Today, and throughout much of the Holocene, the entire region receives its principal moisture in the summer from an interaction of easterly trade winds with the equatorial calms. Glacial records from sites east of 95?? W in Guatemala, Florida, northern Venezuela and Colombia record dry conditions before 12 ka, however. West of 95?? W, glacial conditions were moister than in the Holocene. For example, pollen and diatom data show that Lake Pa??tzcuaro in the central Mexican highlands was cool, deep and fresh during this time and fossil pinyon needles in packrat middens in Chihuahua, Sonora, Arizona, and Texas indicate cooler glacial climates with increased winter precipitation. Cold Gulf of Mexico sea-surface temperatures and reduced strength of the equatorial calms can explain arid full and late glacial environments east of 95?? W whereas an intensified pattern of winter, westerly air flow dominated hydrologic balances as far south as 20?? N. Overall cooler temperatures may have increased effective moisture levels during dry summer months in both areas. ?? 1997 INQUA/ Elsevier Science Ltd.

  18. Glacial lakes of the Central and Patagonian Andes

    NASA Astrophysics Data System (ADS)

    Wilson, Ryan; Glasser, Neil F.; Reynolds, John M.; Harrison, Stephan; Anacona, Pablo Iribarren; Schaefer, Marius; Shannon, Sarah

    2018-03-01

    The prevalence and increased frequency of high-magnitude Glacial Lake Outburst Floods (GLOFs) in the Chilean and Argentinean Andes suggests this region will be prone to similar events in the future as glaciers continue to retreat and thin under a warming climate. Despite this situation, monitoring of glacial lake development in this region has been limited, with past investigations only covering relatively small regions of Patagonia. This study presents new glacial lake inventories for 1986, 2000 and 2016, covering the Central Andes, Northern Patagonia and Southern Patagonia. Our aim was to characterise the physical attributes, spatial distribution and temporal development of glacial lakes in these three sub-regions using Landsat satellite imagery and image datasets available in Google Earth and Bing Maps. Glacial lake water volume was also estimated using an empirical area-volume scaling approach. Results reveal that glacial lakes across the study area have increased in number (43%) and areal extent (7%) between 1986 and 2016. Such changes equate to a glacial lake water volume increase of 65 km3 during the 30-year observation period. However, glacial lake growth and emergence was shown to vary sub-regionally according to localised topography, meteorology, climate change, rate of glacier change and the availability of low gradient ice areas. These and other factors are likely to influence the occurrence of GLOFs in the future. This analysis represents the first large-scale census of glacial lakes in Chile and Argentina and will allow for a better understanding of lake development in this region, as well as, providing a basis for future GLOF risk assessments.

  19. Postglacial recolonization in a cold climate specialist in western Europe: patterns of genetic diversity in the adder (Vipera berus) support the central-marginal hypothesis.

    PubMed

    Ursenbacher, Sylvain; Guillon, Michaël; Cubizolle, Hervé; Dupoué, Andréaz; Blouin-Demers, Gabriel; Lourdais, Olivier

    2015-07-01

    Understanding the impact of postglacial recolonization on genetic diversity is essential in explaining current patterns of genetic variation. The central-marginal hypothesis (CMH) predicts a reduction in genetic diversity from the core of the distribution to peripheral populations, as well as reduced connectivity between peripheral populations. While the CMH has received considerable empirical support, its broad applicability is still debated and alternative hypotheses predict different spatial patterns of genetic diversity. Using microsatellite markers, we analysed the genetic diversity of the adder (Vipera berus) in western Europe to reconstruct postglacial recolonization. Approximate Bayesian Computation (ABC) analyses suggested a postglacial recolonization from two routes: a western route from the Atlantic Coast up to Belgium and a central route from the Massif Central to the Alps. This cold-adapted species likely used two isolated glacial refugia in southern France, in permafrost-free areas during the last glacial maximum. Adder populations further from putative glacial refugia had lower genetic diversity and reduced connectivity; therefore, our results support the predictions of the CMH. Our study also illustrates the utility of highly variable nuclear markers, such as microsatellites, and ABC to test competing recolonization hypotheses. © 2015 John Wiley & Sons Ltd.

  20. Sub-glacial volcanic eruptions

    USGS Publications Warehouse

    White, Donald Edward

    1956-01-01

    The literature on sub-glacial volcanic eruptions and the related flood phenomena has been reviewed as a minor part of the larger problem of convective and conductive heat transfer from intrusive magma. (See Lovering, 1955, for a review of the extensive literature on this subject.) This summary of data on sub-glacial eruptions is part of a program that the U.S. Geological Survey is conducting in connection with its Investigations of Geologic Processes project on behalf of the Division of Research, U.S. Atomic Energy Commission.

  1. Late Ordovician (Ashgillian) glacial deposits in southern Jordan

    NASA Astrophysics Data System (ADS)

    Turner, Brian R.; Makhlouf, Issa M.; Armstrong, Howard A.

    2005-11-01

    The Late Ordovician (Ashgillian) glacial deposits in southern Jordan, comprise a lower and upper glacially incised palaeovalley system, occupying reactivated basement and Pan-African fault-controlled depressions. The lower palaeovalley, incised into shoreface sandstones of the pre-glacial Tubeiliyat Formation, is filled with thin glaciofluvial sandstones at the base, overlain by up to 50 m of shoreface sandstone. A prominent glaciated surface near the top of this palaeovalley-fill contains intersecting glacial striations aligned E-W and NW-SE. The upper palaeovalley-fill comprises glaciofluvial and marine sandstones, incised into the lower palaeovalley or, where this is absent, into the Tubeiliyat Formation. Southern Jordan lay close to the margin of a Late Ordovician terrestrial ice sheet in Northwest Saudi Arabia, characterised by two major ice advances. These are correlated with the lower and upper palaeovalleys in southern Jordan, interrupted by two subsidiary glacial advances during late stage filling of the lower palaeovalley when ice advanced from the west and northwest. Thus, four ice advances are now recorded from the Late Ordovician glacial record of southern Jordan. Disturbed and deformed green sandstones beneath the upper palaeovalley-fill in the Jebel Ammar area, are confined to the margins of the Hutayya graben, and have been interpreted as structureless glacial loessite or glacial rock flour. Petrographic and textural analyses of the deformed sandstones, their mapped lateral transition into undeformed Tubeiliyat marine sandstones away from the fault zone, and the presence of similar sedimentary structures to those in the pre-glacial marine Tubeiliyat Formation suggest that they are a locally deformed facies equivalent of the Tubeiliyat, not part of the younger glacial deposits. Deformation is attributed to glacially induced crustal stresses and seismic reactivation of pre-existing faults, previously weakened by epeirogenesis, triggering sediment

  2. An attempt to replace an ivermectin-resistant Cooperia spp. population by a susceptible one on grazing pastures based on epidemiological principles and refugia management.

    PubMed

    Fiel, C A; Steffan, P E; Muchiut, S M; Fernández, A S; Bernat, G; Riva, E; Lloberas, M M; Almada, A; Homer, D

    2017-11-15

    The maintenance of anthelmintic-susceptible parasite refugia to delay the onset of anthelmintic resistance is an almost impossible effort in many grazing livestock production countries given that current refugia consist of already resistant parasites. Rather, efforts could be focused on replacing the resistant parasite refugia by susceptible parasite ones and implementing sustainable parasite control measures from then on. To this purpose, a trial was conducted to attempt to establish a new population of ivermectin-susceptible Cooperia sp. on a beef cattle farm with proven problems of ivermectin-resistant Cooperia. During two consecutive years, 82 (Year 1) and 100 (Year 2) recently weaned and parasite-free heifers were inoculated with 40,000 or 30,000 susceptible Cooperia L3, respectively, at a time when levels of resistant parasite refugia were normally low. The animals were subsequently allowed to graze on the problem pastures during autumn until the end of spring. Levels of parasitism in the animals and on pasture were monitored monthly and animals were treated with levamisole when needed. The combination of parasitological monitoring and local epidemiological knowledge was essential to determine when treatments were to be administered. No clinical signs of gastrointestinal parasitosis in the herd were observed throughout the study and unnecessary treatments were avoided. Faecal egg counts reduction tests (FECRT) and controlled efficacy tests (CET) employing worm counts were carried out at different times throughout the study to determine the clinical efficacy (FECRT) and the absolute efficacy (CET) of ivermectin, respectively. The clinical efficacy of ivermectin increased from an initial 73% to 99.4%, while the absolute efficacy increased from 54.1% to 87.5% after just two animal production cycles. The switch from a resistant parasite population to a susceptible one requires knowledge of parasitological epidemiology, especially in relation to seasonal

  3. Geodesy: Modeling Earth's Post-Glacial Rebound

    NASA Astrophysics Data System (ADS)

    Spada, Giorgio; Antonioli, Andrea; Boschi, Lapo; Brandi, Valter; Cianetti, Spina; Galvani, Gabriele; Giunchi, Carlo; Perniola, Bruna; Agostinetti, Nicola Piana; Piersanti, Antonio; Stocchi, Paolo

    2004-02-01

    Efforts to mathematically model the Earth's post-glacial rebound, or, in general, long-term planetary-scale viscoelastic deformations, have been ongoing for several decades. Unfortunately, research in the post-glacial rebound community has not been characterized by much exchange of knowledge. Groups around the world have developed their code independently, sometimes with profoundly different approaches, occasionally leading to inconsistent results [e.g., Boschi et al., 1999]. Postglacial Rebound Calculator (TABOO) is a post-glacial rebound software that is being made freely available (through Samizdat Press at http://samizdat.mines.edu/taboo/)in the hope that it might become a common reference for all post-glacial rebound researchers. TABOO is portable and has been tested on Unix, Linux, and Windows systems; all it requires is a Fortran90 compiler supporting quadruple precision. The software is easy to use. It comes with a detailed guide that can work as a quick reference cookbook, and it is also accompanied by a textbook, The Theory Behind TABOO, collecting the most significant theoretical results from post-glacial rebound literature. TABOO is not a ``black-box,'' although it may easily be used as such. The entire source code is provided and should be easy to understand for intermediate-level Fortran programmers.

  4. Phylogeography and postglacial recolonization of Europe by Rhinolophus hipposideros: evidence from multiple genetic markers.

    PubMed

    Dool, Serena E; Puechmaille, Sébastien J; Dietz, Christian; Juste, Javier; Ibáñez, Carlos; Hulva, Pavel; Roué, Stéphane G; Petit, Eric J; Jones, Gareth; Russo, Danilo; Toffoli, Roberto; Viglino, Andrea; Martinoli, Adriano; Rossiter, Stephen J; Teeling, Emma C

    2013-08-01

    The demographic history of Rhinolophus hipposideros (lesser horseshoe bat) was reconstructed across its European, North African and Middle-Eastern distribution prior to, during and following the most recent glaciations by generating and analysing a multimarker data set. This data set consisted of an X-linked nuclear intron (Bgn; 543 bp), mitochondrial DNA (cytb-tRNA-control region; 1630 bp) and eight variable microsatellite loci for up to 373 individuals from 86 localities. Using this data set of diverse markers, it was possible to determine the species' demography at three temporal stages. Nuclear intron data revealed early colonization into Europe from the east, which pre-dates the Quaternary glaciations. The mtDNA data supported multiple glacial refugia across the Mediterranean, the largest of which were found in the Ibero-Maghreb region and an eastern location (Anatolia/Middle East)-that were used by R. hipposideros during the most recent glacial cycles. Finally, microsatellites provided the most recent information on these species' movements since the Last Glacial Maximum and suggested that lineages that had diverged into glacial refugia, such as in the Ibero-Maghreb region, have remained isolated. These findings should be used to inform future conservation management strategies for R. hipposideros and show the power of using a multimarker data set for phylogeographic studies. © 2013 John Wiley & Sons Ltd.

  5. Ireland (2007)

    NASA Technical Reports Server (NTRS)

    2007-01-01

    Between 51.5 and 55.5 degrees north latitude, Ireland could easily find itself buried in snow during the winter, but the island's average temperature in January is 7 degrees Celsius (45 degrees Fahrenheit). Ireland's mild climate results from the influence of the ocean current known as the North Atlantic Drift, which extends the warm waters of the Gulf Stream northward. The island enjoys mild temperatures in the summertime as well; extreme heat and cold are virtually unknown. Precipitation ranges from 78.5 centimeters (31 inches) around Dublin to 300 centimeters (118 inches) along the west coast. The mild, rainy climate is good for vegetation. In this image from the Moderate Resolution Imaging Spectroradiometer (MODIS) on NASA's Terra satellite on May 2, 2007, Ireland overwhelms the viewer with hues of green, particularly in the interior, where the vegetation overlies lowlands of limestone--the remains of marine animals from an ancient sea. Little of the island's famous greenness results from trees, however. Seventeenth-century clearing removed most of the country's forests, and despite replanting efforts, Ireland is Europe's least forested country, after Iceland. The island's mild temperatures and humidity have instead blanketed the landscape in abundant grasses. Along Ireland's west coast, bare brown rocks emerge from the plant cover. In the north, the rocks are primarily ancient, crystalline rocks deposited well over a billion years ago. In the south, the rocks are primarily sandstone deposited roughly 350 million years ago. Evidence of urbanization dots the landscape, especially along the east coast. The metropolitan area of Dublin appears as an uneven patch of gray, mingled with dots of green. Human habitation is also evident around Londonderry, Belfast, and Cork.

  6. Benchmarking care for very low birthweight infants in Ireland and Northern Ireland.

    PubMed

    Murphy, B P; Armstrong, K; Ryan, C A; Jenkins, J G

    2010-01-01

    Benchmarking is that process through which best practice is identified and continuous quality improvement pursued through comparison and sharing. The Vermont Oxford Neonatal Network (VON) is the largest international external reference centre for very low birth weight (VLBW) infants. This report from 2004-7 compares survival and morbidity throughout Ireland and benchmarks these results against VON. A standardised VON database for VLBW infants was created in 14 participating centres across Ireland and Northern Ireland. Data on 716 babies were submitted in 2004, increasing to 796 babies in 2007, with centres caring for from 10 to 120 VLBW infants per year. In 2007, mortality rates in VLBW infants varied from 4% to 19%. Standardised mortality ratios indicate that the number of deaths observed was not significantly different from the number expected, based on the characteristics of infants treated. There was no difference in the incidence of severe intraventricular haemorrhage between all-Ireland and VON groups (5% vs 6%, respectively). All-Ireland rates for chronic lung disease (CLD; 15-21%) remained lower than rates seen in the VON group (24-28%). The rates of late onset nosocomial infection in the all-Ireland group (25-26%) remained double those in the VON group (12-13%). This is the first all-Ireland international benchmarking report in any medical specialty. Survival, severe intraventricular haemorrhage and CLD compare favourably with international standards, but rates of nosocomial infection in neonatal units are concerning. Benchmarking clinical outcomes is critical for quality improvement and informing decisions concerning neonatal intensive care service provision.

  7. Dry season refugia for anopheline larvae and mapping of the seasonal distribution in mosquito larval habitats in Kandi, northeastern Benin.

    PubMed

    Govoetchan, Renaud; Gnanguènon, Virgile; Ogouwalé, Euloge; Oké-Agbo, Frédéric; Azondékon, Roseric; Sovi, Arthur; Attolou, Roseline; Badirou, Kefilath; Youssouf, Ramziyath Agbanrin; Ossè, Razaki; Akogbéto, Martin

    2014-03-31

    The dynamics of mosquito populations depends on availability of suitable surface water for oviposition. It is well known that suitable management of mosquito larval habitats in the sub-Saharan countries, particularly during droughts, could help to suppress vector densities and malaria transmission. We conducted a field survey to investigate the spatial and seasonal distribution of mosquito larval habitats and identify drought-refugia for anopheline larvae. A GIS approach was used to identify, geo-reference and follow up longitudinally from May 2012 to May 2013, all mosquito breeding sites in two rural sites (Yondarou and Thui), one urban (Kossarou), and one peri-urban (Pèdè) site at Kandi, a municipality in northeastern Benin. In Kandi, droughts are excessive with no rain for nearly six months and a lot of sunshine. A comprehensive record of mosquito larval habitats was conducted periodically in all sites for the identification of drought-refugia of anopheline larval stages. With geospatialisation data, seasonal larval distribution maps were generated for each study site with the software ArcGIS version 10.2. Overall, 187 mosquito breeding sites were identified of which 29.95% were recorded during drought. In rural, peri-urban and urban sites, most of the drought-refugia of anopheline larvae were domestic in nature (61.54%). Moreover, in rural settings, anopheline larvae were also sampled in cisterns and wells (25% of larval habitats sampled during drought in Yondarou and 20% in Thui). The mapping showed a significant decrease in the spatial distribution of mosquito larval habitats in rural, peri-urban and urban sites during drought, except in Yondarou (rural) where the aridity did not seem to influence the distribution of larval habitats. Our data showed that the main drought-refugia of anopheline larvae were of a domestic nature as well as wells and cisterns. A suitable management of mosquito larvae in sub-Saharan countries, particularly during droughts, should

  8. Evolution of the climatic tolerance and postglacial range changes of the most primitive orchids (Apostasioideae) within Sundaland, Wallacea and Sahul.

    PubMed

    Kolanowska, Marta; Mystkowska, Katarzyna; Kras, Marta; Dudek, Magdalena; Konowalik, Kamil

    2016-01-01

    The location of possible glacial refugia of six Apostasioideae representatives is estimated based on ecological niche modeling analysis. The distribution of their suitable niches during the last glacial maximum (LGM) is compared with their current potential and documented geographical ranges. The climatic factors limiting the studied species occurrences are evaluated and the niche overlap between the studied orchids is assessed and discussed. The predicted niche occupancy profiles and reconstruction of ancestral climatic tolerances suggest high level of phylogenetic niche conservatism within Apostasioideae.

  9. Climatic vs. tectonic control on glacial relief

    NASA Astrophysics Data System (ADS)

    Prasicek, Günther; Herman, Frederic; Robl, Jörg

    2017-04-01

    The limiting effect of a climatically-induced glacial buzz-saw on the height of mountain ranges has been extensively discussed in the geosciences. The buzz-saw concept assumes that solely climate controls the amount of topography present above the equilibrium line altitude (ELA), while the rock uplift rate plays no relevant role. This view is supported by analyses of hypsometric patterns in orogens worldwide. Furthermore, numerical landscape evolution models show that glacial erosion modifies the hypsometry and reduces the overall relief of mountain landscapes. However, such models often do not incorporate tectonic uplift and can only simulate glacial erosion over a limited amount of time, typically one or several glacial cycles. Constraints on glacial end-member landscapes from analytical, time-independent models are widely lacking. Here we present a steady-state solution for a glacier equilibrium profile in an active orogen modified from the mathematical conception presented by Headley et al. (2012). Our approach combines a glacial erosion law with the shallow ice approximation, specifically the formulations of ice sliding and deformation velocities and ice flux, to calculate ice surface and bed topography from prescribed specific mass balance and rock uplift rate. This solution allows the application of both linear and non-linear erosion laws and can be iteratively fitted to a predefined gradient of specific mass balance with elevation. We tested the influence of climate (fixed rock uplift rate, different ELAs) and tectonic forcing (fixed ELA, different rock uplift rates) on steady-state relief. Our results show that, similar to fluvial orogens, both climate and rock uplift rate exert a strong influence on glacial relief and that the relation among rock uplift rate and relief is governed by the glacial erosion law. This finding can provide an explanation for the presence of high relief in high latitudes. Headley, R.M., Roe, G., Hallet, B., 2012. Glacier

  10. Glacial Buzzcutting and Scarp Encroachment Limit the Height of Tropical Mountains

    NASA Astrophysics Data System (ADS)

    Cunningham, M.; Stark, C. P.; Kaplan, M. R.; Schaefer, J. M.; Winckler, G.

    2016-12-01

    In many mountain ranges hypsometric maxima occur between the glacial equilibrium line altitude (ELA) of the Last Glacial Maximum (LGM) and that of today. A common interpretation of this large-scale observation is that a "glacial buzzsaw" acting throughout the Pleistocene concentrated area within the altitudinal band of ELA fluctuation. This hypothesis remains controversial, however, as there are many examples of uplifted relict surfaces in heavily glaciated areas that occur near the ELA by coincidence. We have focused on the role of glacial erosion in the tropics, where it is spatially restricted to high elevations and temporally limited to global glacial maxima, but appears to have nevertheless truncated vertical orogen growth. Evidence of glacial buzzcutting in some of these ranges has been obscured by post-glacial destruction of glacial valleys by expanding fluvial catchments. We deduce that a duel between glacial buzzcuting and fluvially-driven scarp encroachment has proceeded throughout the Pleistocene in these places. In Costa Rica, we use 10Be and 3He surface-exposure age dating and topographic analysis to confirm that substantial glacial denudation took place at high elevations during the LGM, and employ topographic metrics there and in the Central Range of Taiwan to reveal shrinkage of glacially buzzcut surfaces driven by post-glacial scarp encroachment. These data cast new light on the buzzsaw hypothesis by showing that glacial erosion works with remarkable efficiency in the tropics, precisely where it is likely to be least effective. Our work also draws attention to landscapes with ambiguous signs of glacial erosion, as there are apparent instances of heavily modified, pre-LGM buzzcut surfaces in several tropical ranges. These perched, possibly pre-LGM landscapes may offer a window into previous phases of buzzcutting, and place speed limits on the rate of post-glacial scarp encroachment.

  11. Glacial conditions in the Red Sea

    NASA Astrophysics Data System (ADS)

    Rohling, Eelco J.

    1994-10-01

    In this paper, results from previous studies on planktonic foraminifera, δ18O, and global sea level are combined to discuss climatic conditions in the Red Sea during the last glacial maximum (18,000 B.P.). First, the influence of 120-m sea level lowering on the exchange transport through the strait of Bab-el-Mandab is considered. This strait is the only natural connection of the Red Sea to the open ocean. Next, glacial Red Sea outflow salinity is estimated (about 48 parts per thousand) from the foraminiferal record. Combined, these results yield an estimate of the glacial net water deficit, which appears to have been quite similar to the present (about 2 m yr-1). Finally, budget calculation of δ18O fluxes suggests that the glacial δ18O value of evaporation was about 50% of the present value. This is considered to have resulted from substantially increased mean wind speeds over the glacial Red Sea, which would have caused a rapid drop in the kinematic fractionation factor for 18O. The sensitivity of the calculated values for water deficit and isotopic fractionation to the various assumptions and estimates is evaluated in the discussion. Improvents are to be expected especially through research on the glacial salinity contrast between the Red Sea and Gulf of Aden. It is argued, however, that such future improvement will likely result in a worsening of the isotopic discrepancy, thus increasing the need for an additional mechanism that influenced fractionation (such as mean wind speed). This study demonstrates the need for caution when calculating paleosalinities from δ18O records under the assumption that the modern S∶δ18O relation has remained constant through time. Previously overlooked factors, such as mean wind speed, may have significantly altered that relation in the past.

  12. Automatic Temporal Tracking of Supra-Glacial Lakes

    NASA Astrophysics Data System (ADS)

    Liang, Y.; Lv, Q.; Gallaher, D. W.; Fanning, D.

    2010-12-01

    During the recent years, supra-glacial lakes in Greenland have attracted extensive global attention as they potentially play an important role in glacier movement, sea level rise, and climate change. Previous works focused on classification methods and individual cloud-free satellite images, which have limited capabilities in terms of tracking changes of lakes over time. The challenges of tracking supra-glacial lakes automatically include (1) massive amount of satellite images with diverse qualities and frequent cloud coverage, and (2) diversity and dynamics of large number of supra-glacial lakes on the Greenland ice sheet. In this study, we develop an innovative method to automatically track supra-glacial lakes temporally using the Moderate Resolution Imaging Spectroradiometer (MODIS) time-series data. The method works for both cloudy and cloud-free data and is unsupervised, i.e., no manual identification is required. After selecting the highest-quality image within each time interval, our method automatically detects supra-glacial lakes in individual images, using adaptive thresholding to handle diverse image qualities. We then track lakes across time series of images as lakes appear, change in size, and disappear. Using multi-year MODIS data during melting season, we demonstrate that this new method can detect and track supra-glacial lakes in both space and time with 95% accuracy. Attached figure shows an example of the current result. Detailed analysis of the temporal variation of detected lakes will be presented. (a) One of our experimental data. The Investigated region is centered at Jakobshavn Isbrae glacier in west Greenland. (b) Enlarged view of part of ice sheet. It is partially cloudy and with supra-glacial lakes on it. Lakes are shown as dark spots. (c) Current result. Red spots are detected lakes.

  13. Detecting the limits of northern and southern lineages of tanoak in northern California

    Treesearch

    Eduardo Sandoval-Castro; Richard S. Dodd

    2015-01-01

    Two chloroplast lineages of tanoak (Notholithocarpus densiflorus) meet between Korbel and Hoopa in the North Coast of California. Our earlier work suggests these lineages arose from southern and northern glacial refugia and this region represents their colonizing fronts. Earlier, we detected only one population of mixed lineages, suggesting that...

  14. Last Glacial loess in the conterminous USA

    USGS Publications Warehouse

    Bettis, E. Arthur; Muhs, Daniel R.; Roberts, Helen M.; Wintle, Ann G.

    2003-01-01

    The conterminous United States contains an extensive and generally well-studied record of Last Glacial loess. The loess occurs in diverse physiographic provinces, and under a wide range of climatic and ecological conditions. Both glacial and non-glacia lloess sources are present, and many properties of the loess vary systematically with distance from loess sources. United States' mid-continent Last Glacial loess is probably the thickest in the world, and our calculated mass accumulation rates (MARs) are as high as 17,500 g/m2/yr at the Bignell Hill locality in Nebraska, and many near-source localities have MARs greater than 1500 g/m2/yr. These MARs are high relative to rates calculated in other loess provinces around the world. Recent models of LastGlacial dust sources fail to predict the extent and magnitude of dust flux from the mid-continent of the United States. A better understanding of linkages between climate, ice sheet behaviour, routing of glacial meltwater, land surface processes beyond the ice margin, and vegetation is needed to improve the predictive capabilities of models simulating dust flux from this region.

  15. Glacial Lake Lind, Wisconsin and Minnesota

    USGS Publications Warehouse

    Johnson, M.D.; Addis, K.L.; Ferber, L.R.; Hemstad, C.B.; Meyer, G.N.; Komai, L.T.

    1999-01-01

    Glacial Lake Lind developed in the pre-late Wisconsinan St. Croix River valley, Minnesota and Wisconsin, and lasted more than 1000 yr during the retreat of the Superior lobe at the end of the Wisconsinan glaciation. Lake Lind sediment consists primarily of red varved silt and clay, but also includes mud-flow deposits, nearshore silt (penecontemporaneously deformed in places), nearshore rippled sand, and deltaic sand. Lake Lind varved red clay is not part of glacial Lake Grantsburg, as suggested by earlier authors, because the red varves are separated from overlying glacial Lake Grantsburg silt and clay by a unit of deltaic and fluvial sand. Furthermore, varve correlations indicate that the base of the red varves is younger to the north, showing that the basin expanded as the Superior lobe retreated and was not a lake basin dammed to the southwest by the advancing Grantsburg sublobe. Varve correlations indicate that the Superior lobe retreated at a rate of about 200 m/yr. Uniform winter-clay thickness throughout most of the varve couplets suggests thermal stratification in the lake with clay trapped in the epilimnion; some clay would exit the lake at the outlet prior to winter freeze. Zones of thicker winter-clay layers, in places associated with mud-flow layers, indicate outlet incision, lake-level fall, and shoreline erosion and resuspension of lake clay. The most likely outlet for glacial Lake Lind was in the southwest part of the lake near the present site of Minneapolis, Minnesota. Nearshore sediment indicates that the lake level of glacial Lake Lind was around 280 m. The elevation of the base of the Lake Lind sediments indicates water depth was 20 to 55 m. Evidence in the southern part of the lake basin suggests that the Superior lobe readvanced at least once during the early stages of glacial Lake Lind. Lake Lind ended not by drainage but by being filled in by prograding deltas and outwash plains composed of sand derived from the retreating Superior lobe. It

  16. Phylogeographic, ancient DNA, fossil and morphometric analyses reveal ancient and modern introductions of a large mammal: the complex case of red deer (Cervus elaphus) in Ireland

    NASA Astrophysics Data System (ADS)

    Carden, Ruth F.; McDevitt, Allan D.; Zachos, Frank E.; Woodman, Peter C.; O'Toole, Peter; Rose, Hugh; Monaghan, Nigel T.; Campana, Michael G.; Bradley, Daniel G.; Edwards, Ceiridwen J.

    2012-05-01

    The problem of how and when the island of Ireland attained its contemporary fauna has remained a key question in understanding Quaternary faunal assemblages. We assessed the complex history and origins of the red deer (Cervus elaphus) in Ireland using a multi-disciplinary approach. Mitochondrial sequences of contemporary and ancient red deer (dating from c 30,000 to 1700 cal. yr BP) were compared to decipher possible source populations of red deer in Ireland, in addition to craniometric analyses of skulls from candidate regions to distinguish between different colonization scenarios. Radiocarbon dating was undertaken on all bone fragments that were previously undated. Finally, a comprehensive review of the scientific literature, unpublished reports and other sources of data were also searched for red deer remains within Irish palaeontological and archaeological contexts. Despite being present in Ireland prior to the Last Glacial Maximum (LGM), there is a notable scarcity of red deer from the Younger Dryas stadial period until the Neolithic. The presence of red deer in Irish archaeological sites then occurs more frequently relative to other species. One population in the southwest of Ireland (Co. Kerry) shared haplotypes with the ancient Irish specimens and molecular dating and craniometric analysis suggests its persistence in Ireland since the Neolithic period. The synthesis of the results from this multi-disciplinary study all indicate that red deer were introduced by humans during the Irish Neolithic period and that one of these populations persists today. In conjunction with recent results from other species, Neolithic people from Ireland's nearest landmass, Britain, played a vital role in establishing its contemporary fauna and flora.

  17. Evolution of the climatic tolerance and postglacial range changes of the most primitive orchids (Apostasioideae) within Sundaland, Wallacea and Sahul

    PubMed Central

    Mystkowska, Katarzyna; Kras, Marta; Dudek, Magdalena

    2016-01-01

    The location of possible glacial refugia of six Apostasioideae representatives is estimated based on ecological niche modeling analysis. The distribution of their suitable niches during the last glacial maximum (LGM) is compared with their current potential and documented geographical ranges. The climatic factors limiting the studied species occurrences are evaluated and the niche overlap between the studied orchids is assessed and discussed. The predicted niche occupancy profiles and reconstruction of ancestral climatic tolerances suggest high level of phylogenetic niche conservatism within Apostasioideae. PMID:27635348

  18. Invasive non-native species' provision of refugia for endangered native species.

    PubMed

    Chiba, Satoshi

    2010-08-01

    The influence of non-native species on native ecosystems is not predicted easily when interspecific interactions are complex. Species removal can result in unexpected and undesired changes to other ecosystem components. I examined whether invasive non-native species may both harm and provide refugia for endangered native species. The invasive non-native plant Casuarina stricta has damaged the native flora and caused decline of the snail fauna on the Ogasawara Islands, Japan. On Anijima in 2006 and 2009, I examined endemic land snails in the genus Ogasawarana. I compared the density of live specimens and frequency of predation scars (from black rats [Rattus rattus]) on empty shells in native vegetation and Casuarina forests. The density of land snails was greater in native vegetation than in Casuarina forests in 2006. Nevertheless, radical declines in the density of land snails occurred in native vegetation since 2006 in association with increasing predation by black rats. In contrast, abundance of Ogasawarana did not decline in the Casuarina forest, where shells with predation scars from rats were rare. As a result, the density of snails was greater in the Casuarina forest than in native vegetation. Removal of Casuarina was associated with an increased proportion of shells with predation scars from rats and a decrease in the density of Ogasawarana. The thick and dense litter of Casuarina appears to provide refugia for native land snails by protecting them from predation by rats; thus, eradication of rats should precede eradication of Casuarina. Adaptive strategies, particularly those that consider the removal order of non-native species, are crucial to minimizing the unintended effects of eradication on native species. In addition, my results suggested that in some cases a given non-native species can be used to mitigate the impacts of other non-native species on native species.

  19. Short-term expansion of glacial lakes in the Himalayas

    NASA Astrophysics Data System (ADS)

    Nagai, H.; Tadono, T.

    2017-12-01

    A glacial lake outburst flood (GLOF) is a serious mountainous hazard that is related to glacial shrinkage. Despite technical developments in satellite-based lake expansion monitoring, small glacial lakes were collapsed in Bhutan in June 2015 and in Nepal in May 2017. Relatively heavy rainfall was reported downstream just before the floods. Does a large amount of short-term precipitation have a possibility of triggering a GLOF? To answer this question, the temporal change in the glacial lake area is assessed by means of satellite-based synthetic aperture radar, coupled with satellite-derived spatial and temporal distribution of precipitation to evaluate the contribution of rainfall in glacial lake expansion. The Phased Array type L-band Synthetic Aperture Radar-2 (PALSAR-2) observed the Mande Chu river basin in central Bhutan on Aug 11, 2016. Glacial lakes were manually delineated from the orthorectified backscatter amplitude image. They were compared with those delineated from the old satellite images of ALOS ( 2011), PALSAR-2 (2014-2016), and Landsat-8 (2016). The temporal and spatial distributions of precipitation (2010-2016) are obtained from the Global Satellite Mapping of Precipitation (GSMaP) data (10-km spatial / 60-min. temporal resolutions), calibrated by in situ rain gauges (GSMap_RNL/MVL). The outlines of 11 glacial lakes in the study site were successfully traced from 2011 to 2016; rapid expansion was recorded especially in the period between March and July 2016. In this period, exceeding 500 mm of the total amount of precipitation is recorded by GSMaP, whereas the mean precipitation amount is 300-400 mm in the previous years. This implies that relatively larger precipitation occurred in 2016, which is related to the short-term expansion of the glacial lakes. The rapid expansion of smaller lakes can be explained by their relatively shallow depths, which is sensitive to the increase in inflow water volume. This study highlights the importance of high

  20. Glacial influences on solar radiation in a subarctic sea.

    PubMed

    Barron, Mace G; Barron, Kyle J

    2005-01-01

    Understanding macroscale processes controlling solar radiation in marine systems will be important in interpreting the potential effects of global change from increasing ultraviolet radiation (UV) and glacial retreat. This study provides the first quantitative assessment of UV in the water column of Prince William Sound, a subarctic, semienclosed sea surrounded by mountains, glaciers, rivers, bays and fjords in south central Alaska. Glacial influences on diffuse attenuation coefficients (Kd) were determined along an approximate 120 km transect running NE (61 degrees 07'43''N, 146 degrees 17'1''W) to SW (60 degrees 27'25''N, 148 degrees 05'27'' W). Glacial meltwater and flour caused a 10-fold increase in Kd for visible light, UV-A and UV-B, whereas high optical clarity was present in a diversity of areas away from glacial influences. Transition areas and locations affected by calving of a tidewater glacier had intermediate Kd values. Depths at 99% attenuation ranged from less than 0.2 m near glacial streams to greater than 5 m in bays and open ocean distant from sources of glacial sediments. These results suggest that potential global change from increasing UV and glacial retreat may have heterogeneous effects on subarctic marine systems.

  1. Paleoclimatic modeling and phylogeography of least killifish, Heterandria formosa: insights into Pleistocene expansion-contraction dynamics and evolutionary history of North American Coastal Plain freshwater biota.

    PubMed

    Bagley, Justin C; Sandel, Michael; Travis, Joseph; Lozano-Vilano, María de Lourdes; Johnson, Jerald B

    2013-10-09

    Climatic and sea-level fluctuations throughout the last Pleistocene glacial cycle (~130-0 ka) profoundly influenced present-day distributions and genetic diversity of Northern Hemisphere biotas by forcing range contractions in many species during the glacial advance and allowing expansion following glacial retreat ('expansion-contraction' model). Evidence for such range dynamics and refugia in the unglaciated Gulf-Atlantic Coastal Plain stems largely from terrestrial species, and aquatic species Pleistocene responses remain relatively uninvestigated. Heterandria formosa, a wide-ranging regional endemic, presents an ideal system to test the expansion-contraction model within this biota. By integrating ecological niche modeling and phylogeography, we infer the Pleistocene history of this livebearing fish (Poeciliidae) and test for several predicted distributional and genetic effects of the last glaciation. Paleoclimatic models predicted range contraction to a single southwest Florida peninsula refugium during the Last Glacial Maximum, followed by northward expansion. We inferred spatial-population subdivision into four groups that reflect genetic barriers outside this refuge. Several other features of the genetic data were consistent with predictions derived from an expansion-contraction model: limited intraspecific divergence (e.g. mean mtDNA p-distance = 0.66%); a pattern of mtDNA diversity (mean Hd = 0.934; mean π = 0.007) consistent with rapid, recent population expansion; a lack of mtDNA isolation-by-distance; and clinal variation in allozyme diversity with higher diversity at lower latitudes near the predicted refugium. Statistical tests of mismatch distributions and coalescent simulations of the gene tree lent greater support to a scenario of post-glacial expansion and diversification from a single refugium than to any other model examined (e.g. multiple-refugia scenarios). Congruent results from diverse data indicate H. formosa fits the classic Pleistocene

  2. Evaluation of relocation of unionid mussels to in situ refugia

    USGS Publications Warehouse

    Cope, W.G.; Hove, M.C.; Waller, D.L.; Hornbach, D.J.; Bartsch, M.R.; Cunningham, L.A.; Dunn, H.L.; Kapuscinski, A.R.

    2003-01-01

    The aim of this study was to evaluate the recovery and survival of four species of unionid mussles [pimpleback, Quadrula pustulosa pustulosa (I. Lea, 1831); spike, Elliptio dilatata (Rafinesque, 1820); Higgins eye, Lampsilis higginsii (I. Lea, 1857); and pocketbook, Lampsilis cardium (Rafinesque, 1820)] that were experimentally relocated to in situ refugia in the St Croix River of Minnesota and Wisconsin, USA. In 1996, 150 mussels of each of the first three species (450 total) were relocated to three 5 x 5 m study grids (Site A), one near Lakeland, Minnesota, which served as a source-site control, and two in the experimental refuge 48 km upstream, near Franconia, Minnesota. In a second relocation in 1997, L. Cardium was substituted for L. Higginsii and 150 mussels of this and each of the other two species (450 total), were relocated to two study grids (Site B). The source site control was near Sunrise, Minnesota and the experimental refuge was 14 km downstream near Almelund, Minnesota. Mussel recovery, survival and substratum characteristics were evaluated annually at Site A for 2 years and for 3 years at Site B. Mean annual recovery of all three species ranged from 90 to 100% at Site A, and from 34 to 70% at site B. The mean annual survival of recaptured mussels ranged from 85 to 100% at Site A, and from 88 to 100% at Site B. The textural characteristics of the substratum differed significantly between the control and the two refuge locations at the beginning of the study, but did not differ from this initial status among subsequent years at Site A. At Site B, there was a significant shift in textural characteristics from large to smaller fractions over the four years. The relatively high survival of mussels during this study demonstrates the importance of proper handling and transport protocols when relocating mussels and the selection of suitable relocation habitat with stable substratum. When established correctly, in situ refugia may be a viable tool for

  3. Evaluation of relocation of unionid mussels to in situ refugia

    USGS Publications Warehouse

    Cope, W.G.; Hove, M.C.; Waller, D.L.; Hornbach, D.J.; Bartsch, M.R.; Cunningham, L.A.; Dunn, H.L.; Kapuscinski, A.R.

    2003-01-01

    The aim of this study was to evaluate the recovery and survival of four species of unionid mussles [pimpleback, Quadrula pustulosa pustulosa (I. Lea, 1831); spike, Elliptio dilatata (Rafinesque, 1820); Higgins eye, Lampsilis higginsii (I. Lea, 1857); and pocketbook, Lampsilis cardium (Rafinesque, 1820)] that were experimentally relocated to in situ refugia in the St Croix River of Minnesota and Wisconsin, USA. In 1996, 150 mussels of each of the first three species (450 total) were relocated to three 5 ?? 5 m study grids (Site A), one near Lakeland, Minnesota, which served as a source-site control, and two in the experimental refuge 48 km upstream, near Franconia, Minnesota. In a second relocation in 1997, L. cardium was substituted for L. higginsii and 150 mussels of this and each of the other two species (450 total), were relocated to two study grids (Site B). The source site control was near Sunrise, Minnesota and the experimental refuge was 14 km downstream near Almelund, Minnesota. Mussel recovery, survival and substratum characteristics were evaluated annually at Site A for 2 years and for 3 years at Site B. Mean annual recovery of all three species ranged from 90 to 100% at Site A, and from 34 to 70% at site B. The mean annual survival of recaptured mussels ranged from 85 to 100% at Site A, and from 88 to 100% at Site B. The textural characteristics of the substratum differed significantly between the control and the two refuge locations at the beginning of the study, but did not differ from this initial status among subsequent years at Site A. At Site B, there was a significant shift in textural characteristics from large to smaller fractions over the four years. The relatively high survival of mussels during this study demonstrates the importance of proper handling and transport protocols when relocating mussels and the selection of suitable relocation habitat with stable substratum. When established correctly, in situ refugia may be a viable tool for

  4. The potential drivers in forming avian biodiversity hotspots in the East Himalaya Mountains of Southwest China.

    PubMed

    Lei, Fumin; Qu, Yanhua; Song, Gang; Alström, Per; Fjeldså, Jon

    2015-03-01

    Little has been published to describe or interpret Asian biodiversity hotspots, including those in the East Himalayan Mountains of Southwest China (HMSC), thus making necessary a review of the current knowledge. The Pliocene and Pleistocene geological and glacial histories of the Asian continent differ from those of Europe and North America, suggesting different mechanisms of speciation and extinction, and, thus, different responses to climate changes during the Quaternary glaciations. This short review summarizes potential drivers in shaping and maintaining high species richness and endemism of birds in the HMSC. The geographical location at the junction of different biogeographical realms, the wide range of habitats and climates along the extensive elevational range, the complex topography and the distinct geological history of this region have probably contributed to the evolution of an exceptionally species-rich and endemic-rich, specialized montane avian fauna. The Mountain systems in the HMSC may have provided refugia where species survived during the glacial periods and barriers for preventing species dispersal after the glacial periods. More studies are required to further test this refugia hypothesis by comparing more cold-tolerent and warm-tolerent species. © 2014 International Society of Zoological Sciences, Institute of Zoology/Chinese Academy of Sciences and Wiley Publishing Asia Pty Ltd.

  5. Genetic and palaeo-climatic evidence for widespread persistence of the coastal tree species Eucalyptus gomphocephala (Myrtaceae) during the Last Glacial Maximum.

    PubMed

    Nevill, Paul G; Bradbury, Donna; Williams, Anna; Tomlinson, Sean; Krauss, Siegfried L

    2014-01-01

    Few phylogeographic studies have been undertaken of species confined to narrow, linear coastal systems where past sea level and geomorphological changes may have had a profound effect on species population sizes and distributions. In this study, a phylogeographic analysis was conducted of Eucalyptus gomphocephala (tuart), a tree species restricted to a 400 × 10 km band of coastal sand-plain in south west Australia. Here, there is little known about the response of coastal vegetation to glacial/interglacial climate change, and a test was made as to whether this species was likely to have persisted widely through the Last Glacial Maximum (LGM), or conforms to a post-LGM dispersal model of recovery from few refugia. The genetic structure over the entire range of tuart was assessed using seven nuclear (21 populations; n = 595) and four chloroplast (24 populations; n = 238) microsatellite markers designed for eucalypt species. Correlative palaeodistribution modelling was also conducted based on five climatic variables, within two LGM models. The chloroplast markers generated six haplotypes, which were strongly geographically structured (GST = 0·86 and RST = 0·75). Nuclear microsatellite diversity was high (overall mean HE 0·75) and uniformly distributed (FST = 0·05), with a strong pattern of isolation by distance (r(2) = 0·362, P = 0·001). Distribution models of E. gomphocephala during the LGM showed a wide distribution that extended at least 30 km westward from the current distribution to the palaeo-coastline. The chloroplast and nuclear data suggest wide persistence of E. gomphocephala during the LGM. Palaeodistribution modelling supports the conclusions drawn from genetic data and indicates a widespread westward shift of E. gomphocephala onto the exposed continental shelf during the LGM. This study highlights the importance of the inclusion of complementary, non-genetic data (information on geomorphology and palaeoclimate) to interpret phylogeographic patterns.

  6. Phylogeography of wolves (Canis lupus) in the Pacific Northwest

    USGS Publications Warehouse

    Weckworth, B.V.; Talbot, S.L.; Cook, J.A.

    2010-01-01

    Glacial cycles in the late Pleistocene played a dominant role in sculpting the evolutionary histories of many high-latitude organisms. The refugial hypothesis argues that populations retracted during glacial maxima and were isolated in separate refugia. One prediction of this hypothesis is that populations inhabiting different refugia diverged and then, during interglacial periods, rapidly expanded into deglaciated regions. The range of the gray wolf (Canis lupus) was modified by these expansion and contraction cycles in the late Pleistocene. Our analyses of variation of mitochondrial control region sequences corroborate previous microsatellite analyses supporting independent evolutionary histories for Coastal and Continental wolves in North America. Coastal wolves represent the remnants of a formerly widespread and diverse southern clade that expanded into coastal Southeast Alaska, likely in the early Holocene. In contrast, extant northern Continental populations appear to be admixed, composed of lineages independently arising from ancestors that persisted in either southern or northern (Beringia) refugia. This pattern of diversification suggests the possibility of 3 temporally independent colonizations of North America by wolves from Asia. Coastal wolves are the last vestige of a formerly widespread phylogroup that largely was extirpated in North America by humans during the last century. The independent phylogeographic history of these Coastal wolves has yet to be characterized. Their distinctiveness among North American wolf populations may warrant a reevaluation of their conservation status and management. ?? 2009 American Society of Mammalogists.

  7. Glacial Earthquakes: Monitoring Greenland's Glaciers Using Broadband Seismic Data

    NASA Astrophysics Data System (ADS)

    Olsen, K.; Nettles, M.

    2017-12-01

    The Greenland ice sheet currently loses 400 Gt of ice per year, and up to half of that mass loss comes from icebergs calving from marine-terminating glaciers (Enderlin et al., 2014). Some of the largest icebergs produced by Greenland's glaciers generate magnitude 5 seismic signals when they calve. These glacial earthquakes are recorded by seismic stations around the world. Full-waveform inversion and analysis of glacial earthquakes provides a low-cost tool to identify where and when gigaton-sized icebergs calve, and to track this important mass-loss mechanism in near-real-time. Fifteen glaciers in Greenland are known to have produced glacial earthquakes, and the annual number of these events has increased by a factor of six over the past two decades (e.g., Ekström et al., 2006; Olsen and Nettles, 2017). Since 2000, the number of glacial earthquakes on Greenland's west coast has increased dramatically. Our analysis of three recent years of data shows that more glacial earthquakes occurred on Greenland's west coast from 2011 - 2013 than ever before. In some cases, glacial-earthquake force orientations allow us to identify which section of a glacier terminus produced the iceberg associated with a particular event. We are able to track the timing of major changes in calving-front orientation at several glaciers around Greenland, as well as progressive failure along a single calving front over the course of hours to days. Additionally, the presence of glacial earthquakes resolves a glacier's grounded state, as glacial earthquakes occur only when a glacier terminates close to its grounding line.

  8. The impact of Pleistocene climate change on an ancient arctic–alpine plant: multiple lineages of disparate history in Oxyria digyna

    PubMed Central

    Allen, Geraldine A; Marr, Kendrick L; McCormick, Laurie J; Hebda, Richard J

    2012-01-01

    The ranges of arctic–alpine species have shifted extensively with Pleistocene climate changes and glaciations. Using sequence data from the trnH-psbA and trnT-trnL chloroplast DNA spacer regions, we investigated the phylogeography of the widespread, ancient (>3 million years) arctic–alpine plant Oxyria digyna (Polygonaceae). We identified 45 haplotypes and six highly divergent major lineages; estimated ages of these lineages (time to most recent common ancestor, TMRCA) ranged from ∼0.5 to 2.5 million years. One lineage is widespread in the arctic, a second is restricted to the southern Rocky Mountains of the western United States, and a third was found only in the Himalayan and Altai regions of Asia. Three other lineages are widespread in western North America, where they overlap extensively. The high genetic diversity and the presence of divergent major cpDNA lineages within Oxyria digyna reflect its age and suggest that it was widespread during much of its history. The distributions of individual lineages indicate repeated spread of Oxyria digyna through North America over multiple glacial cycles. During the Last Glacial Maximum it persisted in multiple refugia in western North America, including Beringia, south of the continental ice, and within the northern limits of the Cordilleran ice sheet. Our data contribute to a growing body of evidence that arctic–alpine species have migrated from different source regions over multiple glacial cycles and that cryptic refugia contributed to persistence through the Last Glacial Maximum. PMID:22822441

  9. The glacial iron cycle from source to export

    NASA Astrophysics Data System (ADS)

    Hawkings, J.; Wadham, J. L.; Tranter, M.; Raiswell, R.; Benning, L. G.; Statham, P. J.; Tedstone, A. J.; Nienow, P. W.; Telling, J.; Bagshaw, E.; Simmons, S. L.

    2014-12-01

    Nutrient availability limits primary production in large sectors of the world's oceans. Iron is the major limiting nutrient in around one third of the oceanic euphotic zone, most significantly in the Southern Ocean proximal to Antarctica. In these areas the availability of bioavailable iron can influence the amount of primary production, and thus the strength of the biological pump and associated carbon drawdown from the atmosphere. Despite experiencing widespread iron limitation, the Polar oceans are among the most productive on Earth. Due to the extreme cold, remoteness and their perceived "stasis", ice sheets have previously been though of as insignificant in global biogeochemical cycles. However, large marine algal blooms have been observed in iron-limited areas where glacial influence is large, and it is possible that these areas are stimulated by glacial bioavailable iron input. Here we discuss the importance of the Greenland and Antarctic ice sheets in the global iron cycle. Using field collected trace element data, bulk meltwater chemistry and mineralogical analysis, including photomicrographs, EELS and XANES, we present, for the first time, a conceptual model of the glacial iron cycle from source to export. Using this data we discuss the sources of iron in glacial meltwater, transportation and alteration through the glacial system, and subsequent export to downstream environments. Data collected in 2012 and 2013 from two different Greenlandic glacial catchments are shown, with the most detailed breakdown of iron speciation and concentrations in glacial areas yet reported. Furthermore, the first data from Greenlandic icebergs is presented, allowing meltwater-derived and iceberg-derived iron export to be compared, and the influence of both in marine productivity to be estimated. Using our conceptual model and flux estimates from our dataset, glacial iron delivery in both the northern and southern hemisphere is discussed. Finally, we compare our flux

  10. Lusitania revisited: a phylogeographic analysis of the natterjack toad Bufo calamita across its entire biogeographical range.

    PubMed

    Rowe, Graham; Harris, D James; Beebee, Trevor J C

    2006-05-01

    Attempts to understand the current distributions of plants and animals require both historical and ecological information. Phylogeography has proved highly effective in elucidating historical events such as postglacial colonisations in north temperate zones. However, interesting questions still await resolution. Lusitanian distributions of fauna and flora in western Europe, for example, have puzzled biogeographers for more than 150 years. Lusitanian species have highly disjunct distributions in Ireland and in Iberia, often with few or no other populations inbetween. Despite much debate, no agreed explanation for Lusitanian distributions has yet emerged. We investigated the phylogeographic structure of one Lusitanian species, the natterjack toad Bufo calamita, using mitochondrial DNA control region sequences and allelic variation at eight microsatellite loci. Our results show that this amphibian must have survived in north European refugia, as well as in Iberia, during and since the last (Weichselian) glacial maximum around 20,000 years before present (BP). Subsequent local recolonisation after the Younger Dryas cooling around 11,000 years BP best explains the Lusitanian aspect of natterjack toad distribution.

  11. Volcanic CO2 Emissions and Glacial Cycles: Coupled Oscillations

    NASA Astrophysics Data System (ADS)

    Burley, J. M.; Huybers, P. J.; Katz, R. F.

    2016-12-01

    Following the mid-Pleistocene transition, the dominant period of glacial cycles changed from 40 ka to 100 ka. It is broadly accepted that the 40 ka glacial cycles were driven by cyclical changes in obliquity. However, this forcing does not explain the 100 ka glacial cycles. Mechanisms proposed for 100 ka cycles include isostatic bed depression and proglacial lakes destabilising the Laurentide ice sheet, non-linear responses to orbital eccentricity, and Antarctic ice sheets influencing deep-ocean stratification. None of these are universally accepted. Here we investigate the hypothesis that variations in volcanic CO2 emissions can cause 100 ka glacial cycles. Any proposed mechanism for 100 ka glacial cycles must give the Earth's climate system a memory of 10^4 - 10^5years. This timescale is difficult to achieve for surface processes, however it is possible for the solid Earth. Recent work suggests volcanic CO2 emissions change in response to glacial cycles [1] and that there could be a 50 ka delay in that response [2]. Such a lagged response could drive glacial cycles from 40 ka cycles to an integer multiple of the forcing period. Under what conditions could the climate system admit such a response? To address this, we use a simplified climate model modified from Huybers and Tziperman [3]. Our version comprises three component models for energy balance, ice sheet growth and atmospheric CO2 concentration. The model is driven by insolation alone with other components varying according to a system of coupled, differential equations. The model is run for 500 ka to produce several glacial cycles and the resulting changes in global ice volume and atmospheric CO2 concentration.We obtain a switch from 40 ka to 100 ka cycles as the volcanic CO2 response to glacial cycles is increased. These 100 ka cycles are phase-locked to obliquity, lasting 80 or 120 ka. Whilst the MOR response required (in this model) is larger than plausible estimates based on [2], it illustrates the

  12. North Atlantic Deep Water Production during the Last Glacial Maximum

    PubMed Central

    Howe, Jacob N. W.; Piotrowski, Alexander M.; Noble, Taryn L.; Mulitza, Stefan; Chiessi, Cristiano M.; Bayon, Germain

    2016-01-01

    Changes in deep ocean ventilation are commonly invoked as the primary cause of lower glacial atmospheric CO2. The water mass structure of the glacial deep Atlantic Ocean and the mechanism by which it may have sequestered carbon remain elusive. Here we present neodymium isotope measurements from cores throughout the Atlantic that reveal glacial–interglacial changes in water mass distributions. These results demonstrate the sustained production of North Atlantic Deep Water under glacial conditions, indicating that southern-sourced waters were not as spatially extensive during the Last Glacial Maximum as previously believed. We demonstrate that the depleted glacial δ13C values in the deep Atlantic Ocean cannot be explained solely by water mass source changes. A greater amount of respired carbon, therefore, must have been stored in the abyssal Atlantic during the Last Glacial Maximum. We infer that this was achieved by a sluggish deep overturning cell, comprised of well-mixed northern- and southern-sourced waters. PMID:27256826

  13. A fresh look at glacial foods

    USGS Publications Warehouse

    Colman, Steven M.

    2002-01-01

    Over the last 20 years, it has become clear that ice ages are characterized by glacial as well as climatic instability on millennial time scales. In his Perspective, Colman highlights two recent papers investigating the role of glacial meltwater and continental drainage in this instability. The results suggest a fundamental instability feedback between ocean circulation and ice sheet dynamics and provides an explanation for why instability was greatest at times of intermediate ice volume.

  14. Hybridization among Arctic white-headed gulls (Larus spp.) obscures the genetic legacy of the Pleistocene

    USGS Publications Warehouse

    Sonsthagen, Sarah A.; Chesser, R. Terry; Bell, Douglas A.; Dove, Carla J.

    2012-01-01

    We studied the influence of glacial oscillations on the genetic structure of seven species of white-headed gull that breed at high latitudes (Larus argentatus, L. canus, L. glaucescens, L. glaucoides, L. hyperboreus, L. schistisagus, and L. thayeri). We evaluated localities hypothesized as ice-free areas or glacial refugia in other Arctic vertebrates using molecular data from 11 microsatellite loci, mitochondrial DNA (mtDNA) control region, and six nuclear introns for 32 populations across the Holarctic. Moderate levels of genetic structure were observed for microsatellites (FST= 0.129), introns (ΦST= 0.185), and mtDNA control region (ΦST= 0.461), with among-group variation maximized when populations were grouped based on subspecific classification. Two haplotype and at least two allele groups were observed across all loci. However, no haplotype/allele group was composed solely of individuals of a single species, a pattern consistent with recent divergence. Furthermore, northernmost populations were not well differentiated and among-group variation was maximized when L. argentatus and L. hyberboreus populations were grouped by locality rather than species, indicating recent hybridization. Four populations are located in putative Pleistocene glacial refugia and had larger t estimates than the other 28 populations. However, we were unable to substantiate these putative refugia using coalescent theory, as all populations had genetic signatures of stability based on mtDNA. The extent of haplotype and allele sharing among Arctic white-headed gull species is noteworthy. Studies of other Arctic taxa have generally revealed species-specific clusters as well as genetic structure within species, usually correlated with geography. Aspects of white-headed gull behavioral biology, such as colonization ability and propensity to hybridize, as well as their recent evolutionary history, have likely played a large role in the limited genetic structure observed.

  15. Excitation of the earth's rotational axis by recent glacial discharges

    NASA Technical Reports Server (NTRS)

    Gasperini, P.; Sabadini, R.; Yuen, D. A.

    1986-01-01

    The effects of present-day glacial discharges and the growth of the Antarctic ice sheet on exciting the earth's rotational axis are studied. Glacial forcing could cause a maximum change in J2 of about one-third of the observed amount, for the Maxwell rheology and for Burgers' body models with a long-term, lower-mantle viscosity greater than about 10 to the 23rd P. For transient rheologies the amount of excitation due to glacial melting decreases. Polar wander is not much excited by recent glacial melting for the various types of rheologies examined.

  16. Massive Freshwater discharges: an example from Glacial Lake Missoula

    NASA Astrophysics Data System (ADS)

    Lopes, C.; Mix, A. C.

    2016-12-01

    Massive inputs of freshwater into the ocean are known to disrupt climate. This has been fairly studied in the North Atlantic with freshwater inputs from the Laurentide ice sheet and glacial Lake Agassiz. The association of these discharges with global warming has lead us to look for such prints in marine sediments. Here we show the records of Glacial Lake Missoula outbursts during the warming singe the Last Glacial Maximum in two marine cores off Oregon and California that show the presence of freshwater diatoms that are linked to massive discharges of freshwater from the glacial lake Missoula. The dynamics and timing of these north Pacific mega-flood events are fairly constrained by terrestrial records, however, the consequences of such discharges of freshwater in the northeast Pacific regional circulation remains unknown. Nevertheless we were able to estimate a salinity decrease of almost 6.0 PSU more than 400 km to the south (off northern California) during the last glacial interval (from 16-31 calendar (cal) k.y. B.P.). Anomalously high abundances of freshwater diatoms in marine sediments from the region precede generally accepted dates for the existence of glacial Lake Missoula, implying that large flooding events were also common during the advance of the Cordilleran Ice Sheet.

  17. Southern westerly winds: a pacemaker of Holocene glacial fluctuations in Patagonia?

    NASA Astrophysics Data System (ADS)

    Sagredo, E. A.; Reynhout, S.; Kaplan, M. R.; Patricio, M. I.; Aravena, J. C.; Martini, M. A.; Schaefer, J. M.

    2017-12-01

    A well-resolved glacial chronology is crucial to compare sequences of glacial/climate events within and between regions, and thus, to unravel mechanisms underlying past climate changes. Important efforts have been made towards understanding the Holocene climate evolution of the Southern Andes; however, the timing, patterns and causes of glacial fluctuations during this period still remain elusive. Recent advances in terrestrial cosmogenic nuclide surface exposure dating, together with the establishment of a Patagonian 10Be production rate, have opened new possibilities for establishing high-resolution glacial chronologies at centennial/decadal scale. Here we present a 10Be surface exposure chronology of fluctuations of a small, climate-sensitive mountain glacier at Mt. Fitz Roy area (49.3°S), spanning from the last glacial termination to the present. Thirty new 10Be ages show glacial advances and moraine building events at 17.1±0.9 ka, 13.5±0.5 ka, 10.2±0.7 ka or 9.9±0.5 ka, 6.9±0.2 ka, 6.1±0.3 ka, 4.5±0.2 ka and 0.5±0.1 ka. Similar to the pattern observed in New Zealand, this sequence features progressively less extensive glacial advances during the late-glacial and early Holocene, followed by advances of roughly similar extent during the mid- to late-Holocene. We suggest that while the magnitude of Holocene glacial fluctuations in Patagonia is modulated by SH summer insolation ("modulator"), the specific timing of these glacial events is influenced by centennial-scale shifts of the Southern Westerly Winds ("pacemaker").

  18. Cosmogenic Surface-Exposure Dating of Boulders on Last-Glacial and Late-Glacial Moraines, Lago Buenos Aires, Argentina: Interpretive Strategies and Paleoclimate Implications

    NASA Astrophysics Data System (ADS)

    Douglass, D. C.; Singer, B. S.; Kaplan, M. R.; Mickelson, D. M.; Caffee, M.

    2005-12-01

    The most substantial and least quantifiable source of uncertainty in cosmogenic surface-exposure datasets is the variable exposure histories of boulders from the same landform. The development of precise and accurate chronologies requires distinguishing boulders that best reflect the age of the landform from those which are outliers. We use the Mean Square of Weighted Deviates statistic and cumulative frequency plots to identify groups of samples that have statistically similar ages based on the number of samples and the uncertainty associated with the analyses. This group of samples most likely represents the best estimate of the landform age. We use these tools to interpret 49 surface-exposure ages from six last-glacial and late-glacial moraines at Lago Buenos Aires, Argentina (LBA; 71.0W, 46.5S). Seven of the orty-nine samples are identified as anomalously young, and are interpreted to have been exhumed after moraine deposition. The remaining samples indicate that glacial advances or still-stands of the ice margin occurred at 22.7±0.9, 21.4±1.9, 19.8±1.1, 17.0±0.8, 15.7±0.6, and 14.4±0.9 ka (±2 σ). This maximum ice extent is roughly synchronous with maximum global ice volume and several of the re-advances are contemporaneous with Heinrich events and other Northern Hemisphere cold periods. The late-glacial readvance at ca. 14.4 ka is contemporaneous with the Antarctic Cold Reversal (ACR), and precedes the Younger Dryas Chronozone (YD). No evidence for a Younger Dryas glacial advance has been found in the Lago Buenos Aires basin. This precise glacial chronology indicates there were significant and important differences in climate across southern South America. The timing of maximum ice extent and onset of deglaciation at LBA occur ~4000 years later than in the Chilean Lake District (41S). Fossil pollen from the CLD area indicates cooler conditions between ca. 14.2 and 11.2, and increased silt in a nearby lake core provides indirect evidence for glacial

  19. Spotlight on VET: Ireland

    ERIC Educational Resources Information Center

    Cedefop - European Centre for the Development of Vocational Training, 2013

    2013-01-01

    This paper provides an overview of VET (vocational education and training) in Ireland. In Ireland, the main providers of VET are the national Training and Employment Authority (FAS--a non-commercial semi-State body, part of the public sector) and vocational education committees (VECs--public sector bodies at county level responsible for vocational…

  20. Characteristics of hospital-treated intentional drug overdose in Ireland and Northern Ireland

    PubMed Central

    Griffin, Eve; Corcoran, Paul; Cassidy, Linda; O'Carroll, Amanda; Perry, Ivan J; Bonner, Brendan

    2014-01-01

    Objectives This study compared the profile of intentional drug overdoses (IDOs) presenting to emergency departments in Ireland and in the Western Trust Area of Northern Ireland between 2007 and 2012. Specifically the study aimed to compare characteristics of the patients involved, to explore the factors associated with repeated IDO and to report the prescription rates of common drug types in the population. Methods We utilised data from two comparable registries which monitor the incidence of hospital-treated self-harm, recording data from deliberate self-harm presentations involving an IDO to all hospital emergency departments for the period 1 January 2007 to 31 December 2012. Results Between 2007 and 2012 the registries recorded 56 494 self-harm presentations involving an IDO. The study showed that hospital-treated IDO was almost twice as common in Northern Ireland than in Ireland (278 vs 156/100 000, respectively). Conclusions Despite the overall difference in the rates of IDO, the profile of such presentations was remarkably similar in both countries. Minor tranquillisers were the drugs most commonly involved in IDOs. National campaigns are required to address the availability and misuse of minor tranquillisers, both prescribed and non-prescribed. PMID:25079938

  1. Palaeoflood evidence on the River Nore, South East Ireland

    NASA Astrophysics Data System (ADS)

    Fleming, Ciara; Turner, Jonathan; Bourke, Mary

    2017-04-01

    Past geomorphic changes can be detected in sediment sinks, through the investigation of natural sediment archives. Since the advent of palaeoflood hydrology in the 1980s, numerous authors have demonstrated that such sediment deposits record valuable evidence of past flooding events. Many of these studies have focussed on fluvial systems in arid environments, with bedrock channels proving to be particularly successful field sites. In some districts, the collected datasets are now routinely employed to augment analyses of flood frequency and magnitude, which have traditionally relied on extrapolation of short hydrometric datasets. This study targets river reaches in a temperate humid environment, with a predominantly alluvial channel. The River Nore is one of the largest catchments draining South East Ireland. It is situated in a valley with an inherited glacial legacy and is principally a lowland river catchment. Specific morphological zones have been targeted which are optimal for flood deposit preservation, including palaeochannels, tributary junctions and floodplain overbank settings.There are a variety of anthropogenic pressures evident in this landscape. Among them are channelisation of select tributaries, a legacy of coal mining in the upland Carboniferous limestones, and the installation of man-made obstacles or modifications along the length of the river channel such as sluices and weirs. Regarding land-use, the majority of the catchment is dominated by agriculture, mainly pasture with some tillage. This study investigates palaeoflood evidence in the River Nore catchment and examines the development of the river floodplain using a variety of complementary field and desk-based methods. The sub-surface and micro-topography of river reaches are investigated using Ground Penetrating Radar (GPR) and Unmanned Aerial Vehicle (UAV) technology. Flood deposits have been characterised by examination of bank exposures and sediment cores. Installation of sediment traps

  2. Late glacial aridity in southern Rocky Mountains

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Davis, O K; Pitblado, B L

    While the slopes of the present-day Colorado Rocky Mountains are characterized by large stands of subalpine and montane conifers, the Rockies of the late glacial looked dramatically different. Specifically, pollen records suggest that during the late glacial, Artemisia and Gramineae predominated throughout the mountains of Colorado. At some point between 11,000 and 10,000 B.P., however, both Artemisia and grasses underwent a dramatic decline, which can be identified in virtually every pollen diagram produced for Colorado mountain sites, including Como Lake (Sangre de Cristo Mountains), Copley Lake and Splains; Gulch (near Crested Butte), Molas Lake (San Juan Mountains), and Redrock Lakemore » (Boulder County). Moreover, the same pattern seems to hold for pollen spectra derived for areas adjacent to Colorado, including at sites in the Chuska Mountains of New Mexico and in eastern Wyoming. The implications of this consistent finding are compelling. The closest modem analogues to the Artemisia- and Gramineae-dominated late-glacial Colorado Rockies are found in the relatively arid northern Great Basin, which suggests that annual precipitation was much lower in the late-glacial southern Rocky Mountains than it was throughout the Holocene.« less

  3. Somma-Vesuvius ground deformation over the last glacial cycle

    NASA Astrophysics Data System (ADS)

    Marturano, Aldo; Aiello, Giuseppe; Barra, Diana

    2013-04-01

    Vertical ground movements at Somma-Vesuvius during the last glacial cycle have been inferred from micropalaeontological and petrochemical analyses of rock samples from boreholes drilled at the archaeological sites of Herculaneum and Pompeii as well as on the apron of the volcano and the adjacent Sebeto and Sarno Valleys. Opposing movements occurred during the periods preceding and following the Last Glacial Maximum (LGM). The uplift began 20 ka ago with marine deposits rising several tens of metres up to 25 m a.s.l., recovering previous subsidence which occurred during the Late glacial period, suggesting a strict connection between volcano-tectonic and glacial cycles. Here we present the analysis of deposits predating the LGM, which confirms subsidence of the Campanian Plain where Mt. Somma-Vesuvius is located, shows variable surface loading effects and highlights the volcano-tectonic stages experienced by the volcano. The self-balancing mechanism of the volcanic system, evolving towards an explosive, subaerial activity 60 ka ago, is testified to by a large ground oscillation in phase with sea level change during the last glacial cycle.

  4. What would encourage blood donation in Ireland?

    PubMed

    Harrington, M; Sweeney, M R; Bailie, K; Morris, K; Kennedy, A; Boilson, A; O'Riordan, J; Staines, A

    2007-05-01

    Recent changes have resulted in the loss of 4% of the donor panel in the Republic of Ireland and 3% in Northern Ireland. In order to increase the number of donors in these two regions, it is important that transfusion service providers explore and understand the reasons, which prevent individuals from donating. The aim of this study was to explore these issues particularly in non-donors and those who had lapsed. This 7-month all-Ireland study was conducted by computer-assisted telephone interview. Data collected included sociodemographic history, donation status, as well as barriers/deterrents to donation. There were 4166 completed questionnaires (44% donors; 56% non-donors). Of the donors, 13% had donated blood within the last 2 years. Current donors cited 'awareness of patients needs' (88%), 'trust in the blood transfusion service' (70%), and 'an advertising campaign' (70%) as reasons encouraging them to donate blood. Lapsed donors and non-donors cited 'more frequent mobile clinics/sessions' (30% lapsed donors; 53% non-donors), 'if I was asked' (28% lapsed donors; 53% non-donors), and 'more flexible opening hours' (23% lapsed donors; 44% non-donors) as reasons that would encourage them to donate. The main reasons cited by non-donors for never having donated included 'medical reasons' (41% Republic of Ireland; 43% Northern Ireland), 'lack of information' (20% Republic of Ireland; 22% Northern Ireland), 'fear of needles' (15% Republic of Ireland; 17% Northern Ireland), and 'time constraints' (12% Republic of Ireland; 13% Northern Ireland). Among the non-donor group, 10% (Republic of Ireland) and 6% (Northern Ireland) claimed that they are not permitted to donate. Replacing regular donors is a major challenge for the transfusion service providers. This study shows that by facilitating the general public by introducing more mobile clinics/sessions, more flexible opening hours and having a better level of knowledge in the community about blood donation may encourage

  5. Reliable radiocarbon evidence for the maximum extent of the West Antarctic Ice Sheet in the easternmost Amundsen Sea Embayment during the Last Glacial Maximum

    NASA Astrophysics Data System (ADS)

    Hillenbrand, C. D.; Klages, J. P.; Kuhn, G.; Smith, J.; Graham, A. G. C.; Gohl, K.; Wacker, L.

    2016-02-01

    sheet reconstructions and to quantify precisely the volume of LGM ice-sheet build-up in Antarctica. Our study also alludes to the possibility that refugia for Antarctic shelf benthos may have existed in the ASE during the last glacial period.

  6. The timing and cause of glacial activity during the last glacial in central Tibet based on 10Be surface exposure dating east of Mount Jaggang, the Xainza range

    NASA Astrophysics Data System (ADS)

    Dong, Guocheng; Zhou, Weijian; Yi, Chaolu; Fu, Yunchong; Zhang, Li; Li, Ming

    2018-04-01

    Mountain glaciers are sensitive to climate change, and can provide valuable information for inferring former climates on the Tibetan Plateau (TP). The increasing glacial chronologies indicate that the timing of the local Last Glacial Maximum (LGM) recorded across the TP is asynchronous, implying different local influences of the mid-latitude westerlies and Asian Summer Monsoon in triggering glacier advances. However, the well-dated sites are still too few, especially in the transition zone between regions controlled by the two climate systems. Here we present detailed last glacial chronologies for the Mount Jaggang area, in the Xainza range, central Tibet, with forty-three apparent 10Be exposure-ages ranging from 12.4 ± 0.8 ka to 61.9 ± 3.8 ka. These exposure-ages indicate that at least seven glacial episodes occurred during the last glacial cycle east of Mount Jaggang. These include: a local LGM that occurred at ∼61.9 ± 3.8 ka, possibly corresponding to Marine Isotope Stage 4 (MIS 4); subsequent glacial advances at ∼43.2 ± 2.6 ka and ∼35.1 ± 2.1 ka during MIS 3; one glacial re-advance/standstill at MIS3/2 transition (∼29.8 ± 1.8 ka); and three glacial re-advances/standstills that occurred following MIS 3 at ∼27.9 ± 1.7 ka, ∼21.8 ± 1.3 ka, and ∼15.1 ± 0.9 ka. The timing of these glacial activities is roughly in agreement with North Atlantic millennial-scale climate oscillations (Heinrich events), suggesting the potential correlations between these abrupt climate changes and glacial fluctuations in the Mount Jaggang area. The successively reduced glacial extent might have resulted from an overall decrease in Asian Summer Monsoon intensity over this timeframe.

  7. In and out of glacial extremes by way of dust-climate feedbacks.

    PubMed

    Shaffer, Gary; Lambert, Fabrice

    2018-02-27

    Mineral dust aerosols cool Earth directly by scattering incoming solar radiation and indirectly by affecting clouds and biogeochemical cycles. Recent Earth history has featured quasi-100,000-y, glacial-interglacial climate cycles with lower/higher temperatures and greenhouse gas concentrations during glacials/interglacials. Global average, glacial maxima dust levels were more than 3 times higher than during interglacials, thereby contributing to glacial cooling. However, the timing, strength, and overall role of dust-climate feedbacks over these cycles remain unclear. Here we use dust deposition data and temperature reconstructions from ice sheet, ocean sediment, and land archives to construct dust-climate relationships. Although absolute dust deposition rates vary greatly among these archives, they all exhibit striking, nonlinear increases toward coldest glacial conditions. From these relationships and reconstructed temperature time series, we diagnose glacial-interglacial time series of dust radiative forcing and iron fertilization of ocean biota, and use these time series to force Earth system model simulations. The results of these simulations show that dust-climate feedbacks, perhaps set off by orbital forcing, push the system in and out of extreme cold conditions such as glacial maxima. Without these dust effects, glacial temperature and atmospheric CO 2 concentrations would have been much more stable at higher, intermediate glacial levels. The structure of residual anomalies over the glacial-interglacial climate cycles after subtraction of dust effects provides constraints for the strength and timing of other processes governing these cycles. Copyright © 2018 the Author(s). Published by PNAS.

  8. Circulation and oxygenation of the glacial South China Sea

    NASA Astrophysics Data System (ADS)

    Li, Dawei; Chiang, Tzu-Ling; Kao, Shuh-Ji; Hsin, Yi-Chia; Zheng, Li-Wei; Yang, Jin-Yu Terence; Hsu, Shih-Chieh; Wu, Chau-Ron; Dai, Minhan

    2017-05-01

    Degree of oxygenation in intermediate water modulates the downward transferring efficiency of primary productivity (PP) from surface water to deep water for carbon sequestration, consequently, the storage of nutrients versus the delivery and sedimentary burial fluxes of organic matter and associated biomarkers. To better decipher the PP history of the South China Sea (SCS), appreciation about the glacial-interglacial variation of the Luzon Strait (LS) throughflow, which determines the mean residence time and oxygenation of water mass in the SCS interior, is required. Based on a well-established physical model, we conducted a 3-D modeling exercise to quantify the effects of sea level drop and monsoon wind intensity on glacial circulation pattern, thus, to evaluate effects of productivity and circulation-induced oxygenation on the burial of organic matter. Under modern climatology wind conditions, a 135 m sea-level drop results in a greater basin closeness and a ∼24% of reduction in the LS intermediate westward throughflow, consequently, an increase in the mean water residence time (from 19.0 to 23.0 years). However, when the wind intensity was doubled during glacial low sea-level conditon, the throughflow restored largely to reach a similar residence time (18.4 years) as today regardless its closeness. Comparing with present day SCS, surface circulation pattern in glacial model exhibits (1) stronger upwelling at the west off Luzon Island, and (2) an intensified southwestward jet current along the western boundary of the SCS basin. Superimposed hypothetically by stronger monsoon wind, the glacial SCS conditions facilitate greater primary productivity in the northern part. Manganese, a redox sensitive indicator, in IMAGES core MD972142 at southeastern SCS revealed a relatively reducing environment in glacial periods. Considering the similarity in the mean water residence time between modern and glacial cases, the reducing environment of the glacial southeastern SCS

  9. Phylogeography of the large white-bellied rat Niviventer excelsior suggests the influence of Pleistocene glaciations in the Hengduan mountains.

    PubMed

    Chen, Weicai; Liu, Shaoying; Liu, Yang; Hao, Haibang; Zeng, Bo; Chen, Shunde; Peng, Hongyuan; Yue, Bisong; Zhang, Xiuyue

    2010-06-01

    The Hengduan Mountains, situated in the southeastern Tibetan Plateau, have undergone dramatic geological and climatic changes over the Pleistocene epoch. Several studies have revealed that the mountains served as a refugium during the ice age. The large white-bellied rat Niviventer excelsior is a rodent endemic to the Hengduan Mountains, which makes it an appropriate species for investigating the influence of glacial movements on the genetic structure of mammals. In this study, we sequenced the partial mitochondrial DNA control region from 72 N. excelsior specimens collected from 20 localities. The results revealed very high levels of haplotype diversity (h = 0.947) and nucleotide diversity (pi = 0.101) in this species. No common haplotype was found to be shared in samples from all geographic regions. Demographic analyses suggested that N. excelsior populations had not been subject to either expansion or bottleneck. The phylogenetic relationships among the haplotypes have no correlation with their geographical origins, while topology revealed two major clades. We speculate that the populations of N. excelsior may have been restricted to two separate refugia during the Last Glacial Maximum (0.60-0.17 Mya), with one west and one east of the Shaluli Mountains. Between the two major refugia, there existed a more widely distributed network subrefugia, which conserved genetic variations in N. excelsior. These results indicated that complex topographic configuration in the Hengduan Mountains provided a network of refugia to maintain the high level of genetic diversity in Pleistocene glaciations.

  10. Population structure within an alpine archipelago: strong signature of past climate change in the New Zealand rock wren (Xenicus gilviventris).

    PubMed

    Weston, K A; Robertson, B C

    2015-09-01

    Naturally subdivided populations such as those occupying high-altitude habitat patches of the 'alpine archipelago' can provide significant insight into past biogeographical change and serve as useful models for predicting future responses to anthropogenic climate change. Among New Zealand's alpine taxa, phylogenetic studies support two major radiations: the first correlating with geological forces (Pliocene uplift) and the second with climatic processes (Pleistocene glaciations). The rock wren (Xenicus gilviventris) is a threatened alpine passerine belonging to the endemic New Zealand wren family (Acanthisittidae). Rock wren constitute a widespread, naturally fragmented population, occurring in patches of suitable habitat over c. 900 m in altitude throughout the length of the South Island, New Zealand. We investigated the relative role of historical geological versus climatic processes in shaping the genetic structure of rock wren (N = 134) throughout their range. Using microsatellites combined with nuclear and mtDNA sequence data, we identify a deep north-south divergence in rock wren (3.7 ± 0.5% at cytochrome b) consistent with the glacial refugia hypothesis whereby populations were restricted in isolated refugia during the Pleistocene c. 2 Ma. This is the first study of an alpine vertebrate to test and provide strong evidence for the glacial refugia hypothesis as an explanation for the low endemicity central zone known as the biotic 'gap' in the South Island of New Zealand. © 2015 John Wiley & Sons Ltd.

  11. Explicit tests of palaeodrainage connections of southeastern North America and the historical biogeography of Orangethroat Darters (Percidae: Etheostoma: Ceasia).

    PubMed

    Bossu, Christen M; Beaulieu, Jeremy M; Ceas, Patrick A; Near, Thomas J

    2013-11-01

    The alteration in palaeodrainage river connections has shaped patterns of speciation, genetic diversity and the geographical distribution of the species-rich freshwater fauna of North America. The integration of ancestral range reconstruction methods and divergence time estimates provides an opportunity to infer palaeodrainage connectivity and test alternative palaeodrainage hypotheses. Members of the Orangethroat Darter clade, Ceasia, are endemic to southeastern North America and occur north and south of the Pleistocene glacial front, a distributional pattern that makes this clade of closely related species an ideal system to investigate the number and location of glacial refugia and compare alternative hypotheses regarding the proposed evolution of the Teays-Mahomet palaeodrainage. This study utilized time-calibrated mitochondrial and nuclear gene phylogenies and present-day geographical distributions to investigate hypothesized Teays-Mahomet River connections through time using a dispersal-extinction-cladogenesis (DEC) framework. Results of DEC ancestral area reconstructions indicate that the Teays-Mahomet River was a key dispersal route between disjunct highland regions connecting the Mississippi River tributaries to the Old-Ohio Drainage minimally at two separate occasions during the Pleistocene. There was a dynamic interplay between palaeodrainage connections through time and postglacial range expansion from three glacial refugia that shaped the current genetic structure and geographical distributions of the species that comprise Ceasia. © 2013 John Wiley & Sons Ltd.

  12. Road accident facts Ireland, 2001

    DOT National Transportation Integrated Search

    2002-10-01

    This report covers all road or traffic accidents reported : to the Garda Sochna involving fatalities, personal : injury or material damage which occurred on public : roads in Ireland (exclusive of Northern Ireland) in 2001. : Accidents on private...

  13. Road accident facts Ireland, 2000

    DOT National Transportation Integrated Search

    2001-08-01

    This report covers all road or traffic accidents reported : to the Garda Sochna involving fatalities, personal : injury or material damage which occurred on public : roads in Ireland (exclusive of Northern Ireland) in 2000. : Accidents on private...

  14. Road accident facts Ireland, 1999

    DOT National Transportation Integrated Search

    2000-12-01

    This report covers all road or traffic accidents reported to the Garda Siochana involving fatalities, personal injury, or material damage which occurred on public roads in Ireland (exclusive of Northern Ireland) in 1999. Accidents on private property...

  15. Road accident facts Ireland, 1998

    DOT National Transportation Integrated Search

    1999-10-01

    This report covers all road or traffic accidents reported to the Garda Siochana involving fatalities, personal injury or material damage which occurred on public roads in ireland (exclusive of Northern Ireland) in 1998. Accidents on private property,...

  16. The NCI All Ireland Cancer Conference.

    PubMed

    Johnston; Daly; Liu

    1999-01-01

    The National Cancer Institute (NCI) has recently decided to embark on an international partnership with the developing cancer programs on the Island of Ireland (Northern Ireland and the Republic of Ireland) in an attempt to further improve the quality and range of cancer services available for patients. This Transatlantic Partnership called the All Ireland-NCI Cancer Consortium offers exciting opportunities in cancer treatment, education and research as the cancer-caring communities from both the Republic of Ireland and Northern Ireland prepare to join with the U.S. NCI in this major endeavor. The inaugural event of the partnership will be the NCI All Ireland Cancer Conference to be held in Belfast, October 3-6, 1999. (See www.allirelandcancer.com, for information on the conference.) Cancer is a significant cause of mortality and morbidity on the Island of Ireland. There are approximately 28,000 new cases and approximately 11,000 deaths from cancer each year. Therefore, Northern Ireland and the Republic of Ireland have among the highest cancer incidence and mortality rates in the Western World. In recent years there has been a major restructuring of cancer services in both parts of the Island. This is the result of several government reports such as the Campbell Report in Northern Ireland and the National Strategy Document for Cancer in the Republic of Ireland. The National Strategy Document proposes that cancer treatment services should be centered around primary care services, regional services, supra-regional centers and a national coordinating structure whereby the supra-regional centers deliver specialist surgery, medical and radiation oncology, rehabilitation and specialist palliative care. Three supra-regional cancer centers are being established in the cities of Dublin, Cork and Galway and a National Cancer Forum, which has served as a multidisciplinary advisory board to the Government, has pushed the development and implementation of this plan. This has

  17. Late Pleistocene glacial fluctuations in Cordillera Oriental, subtropical Andes

    NASA Astrophysics Data System (ADS)

    Martini, Mateo A.; Kaplan, Michael R.; Strelin, Jorge A.; Astini, Ricardo A.; Schaefer, Joerg M.; Caffee, Marc W.; Schwartz, Roseanne

    2017-09-01

    The behavior of subtropical glaciers during Middle to Late Pleistocene global glacial maxima and abrupt climate change events, specifically in Earth's most arid low-latitude regions, remains an outstanding problem in paleoclimatology. The present-day climate of Cordillera Oriental, in arid northwestern Argentina, is influenced by shifts in subtropical climate systems, including the South American Summer Monsoon. To understand better past glacier-subtropical climates during the global Last Glacial Maximum (LGM, 26.5-19 ka) and other time periods, we combined geomorphic features with forty-two precise 10Be ages on moraine boulders and reconstructed paleo-equilibrium line altitudes (ELA) at Nevado de Chañi (24°S) in the arid subtropical Andes. We found a major glacial expansion at ∼23 ± 1.6 ka, that is, during the global LGM. Additional glacial expansions are observed before the global LGM (at ∼52-39 ka), and after, at 15 ± 0.5 and 12 ± 0.6 ka. The ∼15 ka glacial event was found on both sides of Chañi and the ∼12 ka event is only recorded on the east side. Reconstructed ELAs of the former glaciers exhibit a rise from east to west that resembles the present subtropical climate trajectory from the Atlantic side of the continent; hence, we infer that this climate pattern must have been present in the past. Based on comparison with other low-latitude paleoclimate records, such as those from lakes and caves, we infer that both temperature and precipitation influenced past glacial occurrence in this sector of the arid Andes. Our findings also imply that abrupt deglacial climate events associated with the North Atlantic, specifically curtailed meridional overturning circulation and regional cooling, may have had attendant impacts on low subtropical Southern Hemisphere latitudes, including the climate systems that affect glacial activity around Nevado de Chañi.

  18. The Southern Glacial Maximum 65,000 years ago and its Unfinished Termination

    NASA Astrophysics Data System (ADS)

    Schaefer, Joerg M.; Putnam, Aaron E.; Denton, George H.; Kaplan, Michael R.; Birkel, Sean; Doughty, Alice M.; Kelley, Sam; Barrell, David J. A.; Finkel, Robert C.; Winckler, Gisela; Anderson, Robert F.; Ninneman, Ulysses S.; Barker, Stephen; Schwartz, Roseanne; Andersen, Bjorn G.; Schluechter, Christian

    2015-04-01

    Glacial maxima and their terminations provide key insights into inter-hemispheric climate dynamics and the coupling of atmosphere, surface and deep ocean, hydrology, and cryosphere, which is fundamental for evaluating the robustness of earth's climate in view of ongoing climate change. The Last Glacial Maximum (LGM, ∼26-19 ka ago) is widely seen as the global cold peak during the last glacial cycle, and its transition to the Holocene interglacial, dubbed 'Termination 1 (T1)', as the most dramatic climate reorganization during this interval. Climate records show that over the last 800 ka, ice ages peaked and terminated on average every 100 ka ('100 ka world'). However, the mechanisms pacing glacial-interglacial transitions remain controversial and in particular the hemispheric manifestations and underlying orbital to regional driving forces of glacial maxima and subsequent terminations remain poorly understood. Here we show evidence for a full glacial maximum in the Southern Hemisphere 65.1 ± 2.7 ka ago and its 'Unfinished Termination'. Our 10Be chronology combined with a model simulation demonstrates that New Zealand's glaciers reached their maximum position of the last glacial cycle during Marine Isotope Stage-4 (MIS-4). Southern ocean and greenhouse gas records indicate coeval peak glacial conditions, making the case for the Southern Glacial Maximum about halfway through the last glacial cycle and only 15 ka after the last warm period (MIS-5a). We present the hypothesis that subsequently, driven by boreal summer insolation forcing, a termination began but remained unfinished, possibly because the northern ice sheets were only moderately large and could not supply enough meltwater to the North Atlantic through Heinrich Stadial 6 to drive a full termination. Yet the Unfinished Termination left behind substantial ice on the northern continents (about 50% of the full LGM ice volume) and after another 45 ka of cooling and ice sheet growth the earth was at inter

  19. High-frequency seismic signals associated with glacial earthquakes in Greenland

    NASA Astrophysics Data System (ADS)

    Olsen, K.; Nettles, M.

    2017-12-01

    Glacial earthquakes are magnitude 5 seismic events generated by iceberg calving at marine-terminating glaciers. They are characterized by teleseismically detectable signals at 35-150 seconds period that arise from the rotation and capsize of gigaton-sized icebergs (e.g., Ekström et al., 2003; Murray et al., 2015). Questions persist regarding the details of this calving process, including whether there are characteristic precursory events such as ice slumps or pervasive crevasse opening before an iceberg rotates away from the glacier. We investigate the high-frequency seismic signals produced before, during, and after glacial earthquakes. We analyze a set of 94 glacial earthquakes that occurred at three of Greenland's major glaciers, Jakobshavn Isbræ, Helheim Glacier, and Kangerdlugssuaq Glacier, from 2001 - 2013. We employ data from the GLISN network of broadband seismometers around Greenland and from short-term seismic deployments located close to the glaciers. These data are bandpass filtered to 3 - 10 Hz and trimmed to one-hour windows surrounding known glacial earthquakes. We observe elevated amplitudes of the 3 - 10 Hz signal for 500 - 1500 seconds spanning the time of each glacial earthquake. These durations are long compared to the 60 second glacial-earthquake source. In the majority of cases we observe an increase in the amplitude of the 3 - 10 Hz signal 200 - 600 seconds before the centroid time of the glacial earthquake and sustained high amplitudes for up to 800 seconds after. In some cases, high-amplitude energy in the 3 - 10 Hz band precedes elevated amplitudes in the 35 - 150 s band by 300 seconds. We explore possible causes for these high-frequency signals, and discuss implications for improving understanding of the glacial-earthquake source.

  20. Oceanographic gradients and seabird prey community dynamics in glacial fjords

    USGS Publications Warehouse

    Arimitsu, Mayumi L.; Piatt, John F.; Madison, Erica N.; Conaway, Jeffrey S.; Hillgruber, N.

    2012-01-01

    Glacial fjord habitats are undergoing rapid change as a result of contemporary global warming, yet little is known about how glaciers influence marine ecosystems. These ecosystems provide important feeding, breeding and rearing grounds for a wide variety of marine organisms, including seabirds of management concern. To characterize ocean conditions and marine food webs near tidewater glaciers, we conducted monthly surveys of oceanographic variables, plankton, fish and seabirds in Kenai Fjords, Alaska, from June to August of 2007 and 2008. We also measured tidal current velocities near glacial features. We found high sediment load from glacial river runoff played a major role in structuring the fjord marine ecosystem. Submerged moraines (sills) isolated cool, fresh, stratified and silt-laden inner fjord habitats from oceanic influence. Near tidewater glaciers, surface layers of turbid glacial runoff limited availability of light to phytoplankton, but macrozooplankton were abundant in surface waters, perhaps due to the absence of a photic cue for diel migration. Fish and zooplankton community structure varied along an increasing temperature gradient throughout the summer. Acoustic measurements indicated that low density patches of fish and zooplankton were available in the surface waters near glacial river outflows. This is the foraging habitat occupied most by Kittlitz's murrelet (Brachyramphus brevirostris), a rare seabird that appears to be specialized for life in glacially influenced environments. Kittlitz's murrelets were associated with floating glacial ice, and they were more likely to occur near glaciers, in deeper water, and in areas with high acoustic backscatter. Kittlitz's murrelet at-sea distribution was limited to areas influenced by turbid glacial outflows, and where prey was concentrated near the surface in waters with low light penetration. Tidewater glaciers impart unique hydrographic characteristics that influence marine plankton and fish

  1. Paleoclimatic modeling and phylogeography of least killifish, Heterandria formosa: insights into Pleistocene expansion-contraction dynamics and evolutionary history of North American Coastal Plain freshwater biota

    PubMed Central

    2013-01-01

    Background Climatic and sea-level fluctuations throughout the last Pleistocene glacial cycle (~130-0 ka) profoundly influenced present-day distributions and genetic diversity of Northern Hemisphere biotas by forcing range contractions in many species during the glacial advance and allowing expansion following glacial retreat ('expansion-contraction’ model). Evidence for such range dynamics and refugia in the unglaciated Gulf-Atlantic Coastal Plain stems largely from terrestrial species, and aquatic species Pleistocene responses remain relatively uninvestigated. Heterandria formosa, a wide-ranging regional endemic, presents an ideal system to test the expansion-contraction model within this biota. By integrating ecological niche modeling and phylogeography, we infer the Pleistocene history of this livebearing fish (Poeciliidae) and test for several predicted distributional and genetic effects of the last glaciation. Results Paleoclimatic models predicted range contraction to a single southwest Florida peninsula refugium during the Last Glacial Maximum, followed by northward expansion. We inferred spatial-population subdivision into four groups that reflect genetic barriers outside this refuge. Several other features of the genetic data were consistent with predictions derived from an expansion-contraction model: limited intraspecific divergence (e.g. mean mtDNA p-distance = 0.66%); a pattern of mtDNA diversity (mean Hd = 0.934; mean π = 0.007) consistent with rapid, recent population expansion; a lack of mtDNA isolation-by-distance; and clinal variation in allozyme diversity with higher diversity at lower latitudes near the predicted refugium. Statistical tests of mismatch distributions and coalescent simulations of the gene tree lent greater support to a scenario of post-glacial expansion and diversification from a single refugium than to any other model examined (e.g. multiple-refugia scenarios). Conclusions Congruent results from diverse data

  2. Glacial melting: an overlooked threat to Antarctic krill.

    PubMed

    Fuentes, Verónica; Alurralde, Gastón; Meyer, Bettina; Aguirre, Gastón E; Canepa, Antonio; Wölfl, Anne-Cathrin; Hass, H Christian; Williams, Gabriela N; Schloss, Irene R

    2016-06-02

    Strandings of marine animals are relatively common in marine systems. However, the underlying mechanisms are poorly understood. We observed mass strandings of krill in Antarctica that appeared to be linked to the presence of glacial meltwater. Climate-induced glacial meltwater leads to an increased occurrence of suspended particles in the sea, which is known to affect the physiology of aquatic organisms. Here, we study the effect of suspended inorganic particles on krill in relation to krill mortality events observed in Potter Cove, Antarctica, between 2003 and 2012. The experimental results showed that large quantities of lithogenic particles affected krill feeding, absorption capacity and performance after only 24 h of exposure. Negative effects were related to both the threshold concentrations and the size of the suspended particles. Analysis of the stomach contents of stranded krill showed large quantities of large particles ( > 10(6 )μm(3)), which were most likely mobilized by glacial meltwater. Ongoing climate-induced glacial melting may impact the coastal ecosystems of Antarctica that rely on krill.

  3. Glacial melting: an overlooked threat to Antarctic krill

    PubMed Central

    Fuentes, Verónica; Alurralde, Gastón; Meyer, Bettina; Aguirre, Gastón E.; Canepa, Antonio; Wölfl, Anne-Cathrin; Hass, H. Christian; Williams, Gabriela N.; Schloss, Irene R.

    2016-01-01

    Strandings of marine animals are relatively common in marine systems. However, the underlying mechanisms are poorly understood. We observed mass strandings of krill in Antarctica that appeared to be linked to the presence of glacial meltwater. Climate-induced glacial meltwater leads to an increased occurrence of suspended particles in the sea, which is known to affect the physiology of aquatic organisms. Here, we study the effect of suspended inorganic particles on krill in relation to krill mortality events observed in Potter Cove, Antarctica, between 2003 and 2012. The experimental results showed that large quantities of lithogenic particles affected krill feeding, absorption capacity and performance after only 24 h of exposure. Negative effects were related to both the threshold concentrations and the size of the suspended particles. Analysis of the stomach contents of stranded krill showed large quantities of large particles ( > 106 μm3), which were most likely mobilized by glacial meltwater. Ongoing climate-induced glacial melting may impact the coastal ecosystems of Antarctica that rely on krill. PMID:27250339

  4. Great Britain and Ireland

    NASA Image and Video Library

    2017-12-08

    NASA image acquired March 26, 2012 This nearly cloud-free view of Great Britain and Ireland was acquired by the Moderate Resolution Imaging Spectroradiometer (MODIS) aboard NASA’s Terra satellite on March 26, 2012. Just a few days into spring, most of the land appears green, although not quite as brilliant as the summertime hues that give Ireland the nickname “the Emerald Island”. The islands of Ireland (west) and Great Britain (east) are separated by the Irish Sea, which is filled with the turquoise, green and tan swirls typical of sediment, although blooming algae could also contribute some color to the waters. To the southeast, the English Channel separates the island of Great Britain from France (south) and Belgium (north). London can be seen as a gray circle situated inland on the tan-colored River Thames. The sediment from the Thames flows into the English Channel due east of London. The United Kingdom is made up of Wales, Scotland and England, all located primarily on the island of Great Britain, and of Northern Ireland, which comprises the northern section of the island of Ireland. Dublin, in the Republic of Ireland, can be seen as a gray smudge on the eastern coast of the island. Almost due west Galway can be seen as a linear gray streak on the northern coast of Galway Bay, with the blue waters of Loch Corrib to the north. Most of the United Kingdom and Ireland are part of the Celtic broadleaf forest ecoregion, where acid-loving oak and mixed oak forests abound, along with fen and swamp forests and ombrotrophic mires. A portion of the Scottish Highlands, in the north of Great Britain, are covered by the Caledon conifer forest ecoregion. The Caledonia conifers once covered a large area of Scotland, but now only about 1% of the original forest survives, mostly high in the cooler areas of the Highlands. NASA/GSFC/Jeff Schmaltz/MODIS Land Rapid Response Team NASA image use policy. NASA Goddard Space Flight Center enables NASA’s mission through four

  5. Calcareous forest seepages acting as biodiversity hotspots and refugia for woodland snail faunas

    NASA Astrophysics Data System (ADS)

    Horsák, Michal; Tajovská, Eva; Horsáková, Veronika

    2017-07-01

    Land-snail species richness has repeatedly been found to increase with the increasing site calcium content and humidity. These two factors, reported as the main drivers of land-snail assemblage diversity, are also among the main habitat characteristics of calcareous seepages. Here we explore local species richness and compositional variation of forest spring-fed patches (i.e. seepages), to test the hypothesis that these habitats might act as biodiversity hotspots and refugia of regional snail faunas. In contrast to treeless spring fens, only little is known about land snail faunas inhabiting forest seepages. Studying 25 isolated calcareous forest seepages, evenly distributed across the White Carpathians Protected Landscape Area (SE Czech Republic), we found that these sites, albeit spatially very limited, can harbour up to 66% of the shelled land-snail species known to occur in this well-explored protected area (in total 83 species). By comparing land snail assemblages of the studied seepages with those occurring in the woodland surroundings of each site as well as those previously sampled in 28 preserved forest sites within the study area, we found the seepages to be among the most species rich sites. Although the numbers of species did not statistically differ among these three systems, we found highly significant differences in species composition. Seepage faunas were composed of many species significantly associated with spring sites, in contrast to the assemblages of both surrounding and preserved forest sites. Our results highly support the hypothesis that calcareous forest seepages might serve as refugia and biodiversity hotspots of regional land snail faunas. Protection of these unique habitats challenges both conservation plans and forest management guidelines as they might act as sources for the recolonization and restoration of forest snail assemblages particularly in areas impoverished by harvesting and clearcutting.

  6. Glacial lake evolution in the southeastern Tibetan Plateau and the cause of rapid expansion of proglacial lakes linked to glacial-hydrogeomorphic processes

    NASA Astrophysics Data System (ADS)

    Song, Chunqiao; Sheng, Yongwei; Ke, Linghong; Nie, Yong; Wang, Jida

    2016-09-01

    Glacial lakes, as an important component of the cryosphere in the southeastern Tibetan Plateau (SETP) in response to climate change, pose significant threats to the downstream lives and properties of people, engineering construction, and ecological environment via outburst floods, yet we currently have limited knowledge of their distribution, evolution, and the driving mechanism of rapid expansions due to the low accessibility and harsh natural conditions. By integrating optical imagery, satellite altimetry and digital elevation model (DEM), this study presents a regional-scale investigation of glacial lake dynamics across two river basins of the SETP during 1988-2013 and further explores the glacial-hydrogeomorphic process of rapidly expanding lakes. In total 1278 and 1396 glacial lakes were inventoried in 1988 and 2013, respectively. Approximately 92.4% of the lakes in 2013 are not in contact with modern glaciers, and the remaining 7.6% includes 27 (1.9%) debris-contact lakes (in contact with debris-covered ice) and 80 (5.7%) cirque lakes. In categorizing lake variations, we found that debris-contact proglacial lakes experienced much more rapid expansions (∼75%) than cirque lakes (∼7%) and non-glacier-contact lakes (∼3%). To explore the cause of rapid expansion for these debris-contact lakes, we further investigated the mass balance of parent glaciers and elevation changes in lake surfaces and debris-covered glacier tongues using time-series Landsat images, ICESat altimetry, and DEM. Results reveal that the upstream expansion of debris-contact proglacial lakes was not directly associated with rising water levels but with a geomorphological alternation of upstream lake basins caused by melting-induced debris subsidence at glacier termini. This suggests that the hydrogeomorphic process of glacier thinning and retreat, in comparison with direct glacial meltwater alone, may have played a dominant role in the recent glacial lake expansion observed across the

  7. Ageing in Changing Community Contexts: Cross-Border Perspectives from Rural Ireland and Northern Ireland

    ERIC Educational Resources Information Center

    Walsh, Kieran; O'Shea, Eamon; Scharf, Thomas; Murray, Michael

    2012-01-01

    Ongoing demographic, social, economic and cultural changes point to the dynamic and continually changing contexts of rural areas in Ireland and Northern Ireland. However, the influence of such changes on the lives of older people remains under-explored, particularly the question of how older people perceive, connect to and engage in their…

  8. Palaeoglaciology of the Alexander Island ice cap, western Antarctic Peninsula, reconstructed from marine geophysical and core data

    NASA Astrophysics Data System (ADS)

    Graham, Alastair G. C.; Smith, James A.

    2012-03-01

    The glacial history of the continental shelf northwest of Alexander Island is not well known, due mainly to a lack of targeted marine data on Antarctica's palaeo-ice sheets in their inter-ice-stream areas. Recently it has been argued that the region was ice-free at the Last Glacial Maximum (LGM) and thus a potential site for glacial refugia. In this paper, multibeam swath bathymetry, sub-bottom profiles and sediment cores are used to map the Alexander Island sector of the Antarctic Peninsula margin, in order to reconstruct the shelf's palaeoglaciology. Sea-floor bedforms provide evidence that an independent ice cap persisted on Alexander Island through the LGM and deglaciation. We show that this ice cap drained via two major, previously-undescribed tidewater outlets (Rothschild and Charcot Glaciers) sourced from an ice dome centred over the west of the island and near-shore areas. The glaciers grounded along deep, fjord-like cross-shelf troughs to within at least ˜10-20 km of the shelf edge, and probably reached the shelf break. Only one small outer-shelf zone appears to have remained free of ice throughout an otherwise extensive LGM. During retreat, grounding-line geomorphology indicates periodic stabilisation of Charcot Glacier on the mid-shelf after 13,500 cal yrs BP, while Rothschild Glacier retreated across its mid-shelf by 14,450 cal yrs BP. The timing of these events is in phase with retreat in nearby Marguerite Trough, and we take this as evidence of a common history and forcing with the Antarctic Peninsula Ice Sheet. The fine details of ice flow documented by our new reconstruction highlight the importance of capturing complex ice flow patterns in models (e.g. in inter-stream areas), for understanding how region-specific parts of Antarctica may change in the future. Moreover, the reconstruction shows that glacial refugia, if present, cannot have been extensive on the Alexander Island shelf at the LGM as indicated by previous biological studies; instead

  9. Hydrologic and Agent-based Modelling of Hydro-refugia in East Africa, Insights into the Importance of Water Resources in Hominin Evolution and Dispersal

    NASA Astrophysics Data System (ADS)

    Ashley, G. M.; Cuthbert, M. O.; Gleeson, T. P.; Reynolds, S. R.; Bennett, M. R.; Newton, A. C.; McCormack, C. J.

    2017-12-01

    Hominin evolution and climate variability have often been linked because of the apparent coincidence of climate fluctuations and speciation or extinctions, although the cause and effect of climate on natural selection is not clear. Climate in the EARS (East African Rift System) where most hominin first occurrences are located experienced an overall drying over the last 7 myr. Superimposed on this trend, Milankovitch cycles generated wet-dry precession cycles ( 23 kyr) that changed both water and food resource availability. During dry periods, lakes became more alkaline and rivers ephemeral but, groundwater, buffered from surface climate effects, remained a potential resource during the driest of times. The possibility of widespread groundwater sources hydro-refugia, such as springs, wetlands and groundwater-fed perennial streams has received little attention with respect to the paleoenvironmental context of hominin evolution or dispersal. We demonstrate that hydrogeological modelling of the modern landscape in East Africa coupled with ABM (agent-based modelling) of hominin movement yields new insight into potential correlates of hominin survival and dispersal. Digitized hydrological mapping of present day rivers, lakes and springs along the EARS (2000 km) from northern Tanzania to Ethiopia provided the modelling framework. Present day conditions are considered analogous to past dry periods; wet period conditions are an expanded hydrologic network including all surface water bodies. Our focus was on perennial springs discharging at 1,000 m3/y (volume to sustain a small wetland). 450 such springs occur and were found to be significantly controlled by geology, not just climate. The ABM was designed to determine if it was possible for humans to walk between hydro-refugia in 3 days. Four climate scenarios were run on ABM: wet, wet-to-dry, dry and dry-to-wet. During dry periods our results suggest that groundwater availability would have been critical to supporting

  10. Paleoclimate: A fresh look at glacial floods

    USGS Publications Warehouse

    Colman, S. M.

    2002-01-01

    Over the last 20 years, it has become clear that ice ages are characterized by glacial as well as climatic instability on millennial time scales. In his Perspective, Colman highlights two recent papers investigating the role of glacial meltwater and continental drainage in this instability. The results suggest a fundamental instability feedback between ocean circulation and ice sheet dynamics and provides an explanation for why instability was greatest at times of intermediate ice volume.

  11. Breakup of last glacial deep stratification in the South Pacific

    NASA Astrophysics Data System (ADS)

    Basak, Chandranath; Fröllje, Henning; Lamy, Frank; Gersonde, Rainer; Benz, Verena; Anderson, Robert F.; Molina-Kescher, Mario; Pahnke, Katharina

    2018-02-01

    Stratification of the deep Southern Ocean during the Last Glacial Maximum is thought to have facilitated carbon storage and subsequent release during the deglaciation as stratification broke down, contributing to atmospheric CO2 rise. Here, we present neodymium isotope evidence from deep to abyssal waters in the South Pacific that confirms stratification of the deepwater column during the Last Glacial Maximum. The results indicate a glacial northward expansion of Ross Sea Bottom Water and a Southern Hemisphere climate trigger for the deglacial breakup of deep stratification. It highlights the important role of abyssal waters in sustaining a deep glacial carbon reservoir and Southern Hemisphere climate change as a prerequisite for the destabilization of the water column and hence the deglacial release of sequestered CO2 through upwelling.

  12. Magnetic Signature of Glacial Flour in Sediments From Bear Lake, Utah/Idaho

    NASA Astrophysics Data System (ADS)

    Rosenbaum, J. G.; Dean, W. E.; Colman, S. M.; Reynolds, R. L.

    2002-12-01

    Variations in magnetic properties within an interval of Bear Lake sediments correlative with oxygen isotope stage 2 (OIS 2) and OIS 3 provide a record of glacial flour production for the Uinta Mountains. Like sediments of the same age from Upper Klamath Lake (OR), these Bear Lake sediments have high magnetic susceptibilities (MS) relative to non-glacial-age sediments and contain well-defined millennial-scale variations in magnetic properties. In contrast to glacial flour derived from volcanic rocks surrounding Upper Klamath Lake, glacial flour derived from the Uinta Mountains and deposited in Bear Lake by the Bear River has low magnetite content but high hematite content. The relatively low MS values of younger and older non-glacial-age sediments are due entirely to dilution by non-magnetic endogenic carbonate and to the effects of sulfidic alteration of detrital Fe-oxides. Analysis of samples from streams entering Bear Lake and from along the course of the Bear River demonstrates that, in comparison to other areas of the catchment, sediment derived from the Uinta Mountains is rich in hematite (high HIRM) and aluminum, and poor in magnetite (low MS) and titanium. Within the glacial-age lake sediments, there are strong positive correlations among HIRM, Al/Ti, and fine sediment grain size. MS varies inversely with theses three variables. These relations indicate that the observed millennial-scale variations in magnetic and chemical properties arise from varying proportions of two detrital components: (1) very fine-grained glacial flour derived from Proterozoic metasedimentary rocks in the Uinta Mountains and characterized by high HIRM and low MS, and (2) somewhat coarser material, characterized by higher MS and lower HIRM, derived from widespread sedimentary rocks along the course of the Bear River and around Bear Lake. Measurement of glacial flour incorporated in lake sediments can provide a continuous history of alpine glaciation, because the rate of accumulation

  13. Characteristics of hospital-treated intentional drug overdose in Ireland and Northern Ireland.

    PubMed

    Griffin, Eve; Corcoran, Paul; Cassidy, Linda; O'Carroll, Amanda; Perry, Ivan J; Bonner, Brendan

    2014-07-29

    This study compared the profile of intentional drug overdoses (IDOs) presenting to emergency departments in Ireland and in the Western Trust Area of Northern Ireland between 2007 and 2012. Specifically the study aimed to compare characteristics of the patients involved, to explore the factors associated with repeated IDO and to report the prescription rates of common drug types in the population. We utilised data from two comparable registries which monitor the incidence of hospital-treated self-harm, recording data from deliberate self-harm presentations involving an IDO to all hospital emergency departments for the period 1 January 2007 to 31 December 2012. Between 2007 and 2012 the registries recorded 56 494 self-harm presentations involving an IDO. The study showed that hospital-treated IDO was almost twice as common in Northern Ireland than in Ireland (278 vs 156/100 000, respectively). Despite the overall difference in the rates of IDO, the profile of such presentations was remarkably similar in both countries. Minor tranquillisers were the drugs most commonly involved in IDOs. National campaigns are required to address the availability and misuse of minor tranquillisers, both prescribed and non-prescribed. Published by the BMJ Publishing Group Limited. For permission to use (where not already granted under a licence) please go to http://group.bmj.com/group/rights-licensing/permissions.

  14. Comparison of glacial and non-glacial-fed streams to evaluate the loading of persistent organic pollutants through seasonal snow/ice melt.

    PubMed

    Bizzotto, E C; Villa, S; Vaj, C; Vighi, M

    2009-02-01

    The release of persistent organic pollutants (PCBs, HCB, HCHs and DDTs) accumulated in Alpine glaciers, was studied during spring-summer 2006 on the Frodolfo glacial-fed stream (Italian Alps). Samples were also taken on a non-glacial stream in the same valley, to compare POP contribution from different water sources (glacier ice, recent snow and spring). In late spring and early summer (May, June) recent snow melting is the most important process. POP contamination is more affected by local emissions and transport, and comparable levels have been measured in both streams for all studied compounds. In late summer and autumn (July-October), the contribution of ice melting strongly increases. In the glacial-fed stream the concentration of chlorinated pesticides (HCHs and DDTs) is about one order of magnitude higher than in the non-glacial-fed. A different behaviour was observed for PCBs, characterised by a peak in June showing, in both streams, concentrations three orders of magnitude higher than the background levels measured in May and in October. This result should be attributed to local emissions rather than long range atmospheric transport (LRAT). This hypothesis is supported by the PCB congener profile in June strictly comparable to the most commonly used Aroclor technical mixtures. The different seasonal behaviour observed for the different groups of chemicals indicates the POP loading in glacial streams is a combined role of long range atmospheric transport and local emission.

  15. Glacial lake expansion in the central Himalayas by Landsat images, 1990-2010.

    PubMed

    Nie, Yong; Liu, Qiao; Liu, Shiyin

    2013-01-01

    Glacial lake outburst flood (GLOF) is a serious hazard in high, mountainous regions. In the Himalayas, catastrophic risks of GLOFs have increased in recent years because most Himalayan glaciers have experienced remarkable downwasting under a warming climate. However, current knowledge about the distribution and recent changes in glacial lakes within the central Himalaya mountain range is still limited. Here, we conducted a systematic investigation of the glacial lakes within the entire central Himalaya range by using an object-oriented image processing method based on the Landsat Thematic Mapper (TM) or Enhanced Thematic Mapper (ETM) images from 1990 to 2010. We extracted the lake boundaries for four time points (1990, 2000, 2005 and 2010) and used a time series inspection method combined with a consistent spatial resolution of Landsat images that consistently revealed lake expansion. Our results show that the glacial lakes expanded rapidly by 17.11% from 1990 to 2010. The pre-existing, larger glacial lakes, rather than the newly formed lakes, contributed most to the areal expansion. The greatest expansions occurred at the altitudinal zones between 4800 m and 5600 m at the north side of the main Himalayan range and between 4500 m and 5600 m at the south side, respectively. Based on the expansion rate, area and type of glacial lakes, we identified 67 rapidly expanding glacial lakes in the central Himalayan region that need to be closely monitored in the future. The warming and increasing amounts of light-absorbing constituents of snow and ice could have accelerated the melting that directly affected the glacial lake expansion. Across the main central Himalayas, glacial lakes at the north side show more remarkable expansion than those at the south side. An effective monitoring and warning system for critical glacial lakes is urgently needed.

  16. Glacial Lake Expansion in the Central Himalayas By Landsat Images, 1990-2010

    NASA Astrophysics Data System (ADS)

    Nie, Y.; Liu, Q.; Liu, S.

    2014-12-01

    Glacial lake outburst flood (GLOF) is a serious hazard in high, mountainous regions. In the Himalayas, catastrophic risks of GLOFs have increased in recent years because most Himalayan glaciers have experienced remarkable downwasting under a warming climate. However, current knowledge about the distribution and recent changes in glacial lakes within the central Himalaya mountain range is still limited. Here, we conducted a systematic investigation of the glacial lakes within the entire central Himalaya range by using an object-oriented image processing method based on the Landsat Thematic Mapper (TM) or Enhanced Thematic Mapper (ETM) images from 1990 to 2010. We extracted the lake boundaries for four time points (1990, 2000, 2005 and 2010) and used a time series inspection method combined with a consistent spatial resolution of Landsat images that consistently revealed lake expansion. Our results show that the glacial lakes expanded rapidly by 17.11% from 1990 to 2010. The pre-existing, larger glacial lakes, rather than the newly formed lakes, contributed most to the areal expansion. The greatest expansions occurred at the altitudinal zones between 4800 m and 5600 m at the north side of the main Himalayan range and between 4500 m and 5600 m at the south side, respectively. Based on the expansion rate, area and type of glacial lakes, we identified 67 rapidly expanding glacial lakes in the central Himalayan region that need to be closely monitored in the future. The warming and increasing amounts of light-absorbing constituents of snow and ice could have accelerated the melting that directly affected the glacial lake expansion. Across the main central Himalayas, glacial lakes at the north side show more remarkable expansion than those at the south side. An effective monitoring and warning system for critical glacial lakes is urgently needed.

  17. Phylogeny and diversification of mountain vipers (Montivipera, Nilson et al., 2001) triggered by multiple Plio-Pleistocene refugia and high-mountain topography in the Near and Middle East.

    PubMed

    Stümpel, Nikolaus; Rajabizadeh, Mehdi; Avcı, Aziz; Wüster, Wolfgang; Joger, Ulrich

    2016-08-01

    The Near and Middle East is a hotspot of biodiversity, but the region remains underexplored at the level of genetic biodiversity. Here, we present an extensive molecular phylogeny of the viperid snake genus Montivipera, including all known taxa. Based on nuclear and mitochondrial data, we present novel insights into the phylogeny of the genus and review the status of its constituent species. Maximum likelihood methods revealed a montane origin of Montivipera at 12.3Mya. We then analyzed factors of mountain viper diversity. Our data support substantial changes in effective population size through Plio-Pleistocene periods. We conclude that climatic oscillations were drivers of allopatric speciation, and that mountain systems of the Near and Middle East have strongly influenced the evolution and survival of taxa, because climatic and topographical heterogeneities induced by mountains have played a crucial role as filters for dispersal and as multiple refugia. The wide diversity of montane microhabitats enabled mountain vipers to retain their ecological niche during climatic pessima. In consequence the varied geological and topographical conditions between refugia favoured genetic isolation and created patterns of species richness resulting in the formation of neoendemic taxa. Our data support high concordance between geographic distributions of Montivipera haplotypes with putative plant refugia. Copyright © 2016. Published by Elsevier Inc.

  18. International trends in health science librarianship Part 8: the UK and the Republic of Ireland Northern Ireland.

    PubMed

    Latimer, Karen; Lawton, Aoife

    2013-12-01

    This is the 8th in a series of articles exploring international trends in health science librarianship with a focus on the UK and Ireland in the first decade of the 21st century. The invited authors are from Northern Ireland and the Republic of Ireland. Future issues will track trends from Scotland and Wales. © 2013 The authors. Health Information and Libraries Journal © 2013 Health Libraries Group.

  19. Genetic and palaeo-climatic evidence for widespread persistence of the coastal tree species Eucalyptus gomphocephala (Myrtaceae) during the Last Glacial Maximum

    PubMed Central

    Nevill, Paul G.; Bradbury, Donna; Williams, Anna; Tomlinson, Sean; Krauss, Siegfried L.

    2014-01-01

    Background and Aims Few phylogeographic studies have been undertaken of species confined to narrow, linear coastal systems where past sea level and geomorphological changes may have had a profound effect on species population sizes and distributions. In this study, a phylogeographic analysis was conducted of Eucalyptus gomphocephala (tuart), a tree species restricted to a 400 × 10 km band of coastal sand-plain in south west Australia. Here, there is little known about the response of coastal vegetation to glacial/interglacial climate change, and a test was made as to whether this species was likely to have persisted widely through the Last Glacial Maximum (LGM), or conforms to a post-LGM dispersal model of recovery from few refugia. Methods The genetic structure over the entire range of tuart was assessed using seven nuclear (21 populations; n = 595) and four chloroplast (24 populations; n = 238) microsatellite markers designed for eucalypt species. Correlative palaeodistribution modelling was also conducted based on five climatic variables, within two LGM models. Key Results The chloroplast markers generated six haplotypes, which were strongly geographically structured (GST = 0·86 and RST = 0·75). Nuclear microsatellite diversity was high (overall mean HE 0·75) and uniformly distributed (FST = 0·05), with a strong pattern of isolation by distance (r2 = 0·362, P = 0·001). Distribution models of E. gomphocephala during the LGM showed a wide distribution that extended at least 30 km westward from the current distribution to the palaeo-coastline. Conclusions The chloroplast and nuclear data suggest wide persistence of E. gomphocephala during the LGM. Palaeodistribution modelling supports the conclusions drawn from genetic data and indicates a widespread westward shift of E. gomphocephala onto the exposed continental shelf during the LGM. This study highlights the importance of the inclusion of complementary, non-genetic data (information on geomorphology and

  20. Polyploid evolution and Pleistocene glacial cycles: A case study from the alpine primrose Primula marginata (Primulaceae)

    PubMed Central

    2012-01-01

    Background Recent studies highlighted the role of Pleistocene climatic cycles in polyploid speciation and of southern Alpine refugia as reservoirs of diversity during glacial maxima. The polyploid Primula marginata, endemic to the southwestern Alps, includes both hexaploid and dodecaploid cytotypes that show no ecological or morphological differences. We used flow cytometry to determine variation and geographic distribution of cytotypes within and between populations and analyses of chloroplast (cp) and nuclear ribosomal (nr) DNA sequences from the Internal Transcribed Spacer (ITS) region to infer the evolutionary history of the two cytotypes and the auto- vs. allopolyploid origin of dodecaploid populations. Results We did not detect any intermediate cytotypes or variation of ploidy levels within populations. Hexaploids occur in the western and dodecaploids in the eastern part of the distributional range, respectively. The cpDNA and nrDNA topologies are in conflict, for the former supports shared ancestry between P. marginata and P. latifolia, while the latter implies common origins between at least some ITS clones of P. marginata and P. allionii. Conclusions Our results suggest an initial episode of chloroplast capture involving ancestral lineages of P. latifolia and P. marginata, followed by polyploidization between P. marginata-like and P. allionii-like lineages in a southern refugium of the Maritime Alps. The higher proportion of ITS polymorphisms in dodecaploid than in hexaploid accessions of P. marginata and higher total nucleotide diversity of ITS clones in dodecaploid vs. hexaploid individuals sequences are congruent with the allopolyploid hypothesis of dodecaploid origin. PMID:22530870

  1. A 305-year continuous monthly rainfall series for the island of Ireland (1711-2016)

    NASA Astrophysics Data System (ADS)

    Murphy, Conor; Broderick, Ciaran; Burt, Timothy P.; Curley, Mary; Duffy, Catriona; Hall, Julia; Harrigan, Shaun; Matthews, Tom K. R.; Macdonald, Neil; McCarthy, Gerard; McCarthy, Mark P.; Mullan, Donal; Noone, Simon; Osborn, Timothy J.; Ryan, Ciara; Sweeney, John; Thorne, Peter W.; Walsh, Seamus; Wilby, Robert L.

    2018-03-01

    A continuous 305-year (1711-2016) monthly rainfall series (IoI_1711) is created for the Island of Ireland. The post 1850 series draws on an existing quality assured rainfall network for Ireland, while pre-1850 values come from instrumental and documentary series compiled, but not published by the UK Met Office. The series is evaluated by comparison with independent long-term observations and reconstructions of precipitation, temperature and circulation indices from across the British-Irish Isles. Strong decadal consistency of IoI_1711 with other long-term observations is evident throughout the annual, boreal spring and autumn series. Annually, the most recent decade (2006-2015) is found to be the wettest in over 300 years. The winter series is probably too dry between the 1740s and 1780s, but strong consistency with other long-term observations strengthens confidence from 1790 onwards. The IoI_1711 series has remarkably wet winters during the 1730s, concurrent with a period of strong westerly airflow, glacial advance throughout Scandinavia and near unprecedented warmth in the Central England Temperature record - all consistent with a strongly positive phase of the North Atlantic Oscillation. Unusually wet summers occurred in the 1750s, consistent with proxy (tree-ring) reconstructions of summer precipitation in the region. Our analysis shows that inter-decadal variability of precipitation is much larger than previously thought, while relationships with key modes of climate variability are time-variant. The IoI_1711 series reveals statistically significant multi-centennial trends in winter (increasing) and summer (decreasing) seasonal precipitation. However, given uncertainties in the early winter record, the former finding should be regarded as tentative. The derived record, one of the longest continuous series in Europe, offers valuable insights for understanding multi-decadal and centennial rainfall variability in Ireland, and provides a firm basis for

  2. Glacial Cycles Influence Marine Methane Hydrate Formation

    NASA Astrophysics Data System (ADS)

    Malinverno, A.; Cook, A. E.; Daigle, H.; Oryan, B.

    2018-01-01

    Methane hydrates in fine-grained continental slope sediments often occupy isolated depth intervals surrounded by hydrate-free sediments. As they are not connected to deep gas sources, these hydrate deposits have been interpreted as sourced by in situ microbial methane. We investigate here the hypothesis that these isolated hydrate accumulations form preferentially in sediments deposited during Pleistocene glacial lowstands that contain relatively large amounts of labile particulate organic carbon, leading to enhanced microbial methanogenesis. To test this hypothesis, we apply an advection-diffusion-reaction model with a time-dependent organic carbon deposition controlled by glacioeustatic sea level variations. In the model, hydrate forms in sediments with greater organic carbon content deposited during the penultimate glacial cycle ( 120-240 ka). The model predictions match hydrate-bearing intervals detected in three sites drilled on the northern Gulf of Mexico continental slope, supporting the hypothesis of hydrate formation driven by enhanced organic carbon burial during glacial lowstands.

  3. Glacial cycles influence marine methane hydrate formation

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Malinverno, A.; Cook, A. E.; Daigle, H.

    Methane hydrates in fine-grained continental slope sediments often occupy isolated depth intervals surrounded by hydrate-free sediments. As they are not connected to deep gas sources, these hydrate deposits have been interpreted as sourced by in situ microbial methane. We investigate here the hypothesis that these isolated hydrate accumulations form preferentially in sediments deposited during Pleistocene glacial lowstands that contain relatively large amounts of labile particulate organic carbon, leading to enhanced microbial methanogenesis. To test this hypothesis, we apply an advection-diffusion-reaction model with a time-dependent organic carbon deposition controlled by glacioeustatic sea level variations. In the model, hydrate forms in sedimentsmore » with greater organic carbon content deposited during the penultimate glacial cycle (~120-240 ka). As a result, the model predictions match hydrate-bearing intervals detected in three sites drilled on the northern Gulf of Mexico continental slope, supporting the hypothesis of hydrate formation driven by enhanced organic carbon burial during glacial lowstands.« less

  4. Glacial cycles influence marine methane hydrate formation

    DOE PAGES

    Malinverno, A.; Cook, A. E.; Daigle, H.; ...

    2018-01-12

    Methane hydrates in fine-grained continental slope sediments often occupy isolated depth intervals surrounded by hydrate-free sediments. As they are not connected to deep gas sources, these hydrate deposits have been interpreted as sourced by in situ microbial methane. We investigate here the hypothesis that these isolated hydrate accumulations form preferentially in sediments deposited during Pleistocene glacial lowstands that contain relatively large amounts of labile particulate organic carbon, leading to enhanced microbial methanogenesis. To test this hypothesis, we apply an advection-diffusion-reaction model with a time-dependent organic carbon deposition controlled by glacioeustatic sea level variations. In the model, hydrate forms in sedimentsmore » with greater organic carbon content deposited during the penultimate glacial cycle (~120-240 ka). As a result, the model predictions match hydrate-bearing intervals detected in three sites drilled on the northern Gulf of Mexico continental slope, supporting the hypothesis of hydrate formation driven by enhanced organic carbon burial during glacial lowstands.« less

  5. Glacial Meltwater Contirbutions to the Bow River, Alberta, Canada

    NASA Astrophysics Data System (ADS)

    Bash, E. A.; Marshall, S. J.; White, E. C.

    2009-12-01

    Assessment of glacial melt is critical for water resource management in areas which rely on glacier-fed rivers for agricultural and municipal uses. Changes in precipitation patterns coupled with current glacial retreat are altering the glacial contribution to river flow in areas such as the Andes of South America and the high ranges of Asia, as well as the Rockies of Western Canada. Alberta’s Bow River has its headwaters in the eastern slopes of the Canadian Rockies and contributes to the Nelson drainage system feeding into Hudson Bay. The Bow River basin contains several population centers, including the City of Calgary, and is heavily taxed for agricultural use. The combined effects of rapid glacial retreat in the Canadian Rockies, higher drought frequency, and increased demand are likely to heighten water stress in Southern Alberta. However, there has been little focus to date on the extent and importance of glacial meltwater in the Bow River. The Bow River contains 74.5 km2 of glacier ice, which amounts to only 0.29% of the basin. While this number is not high compared to some glacierized areas, Hopkinson and Young (1998) report that in dry years, glacier melt can provide up to 50% of late summer flows at a station in the upper reaches of the river system. We extend this work with an assessment of monthly and annual glacial contributions to the Bow River farther downstream in Calgary. Our analysis is based on mass balance, meteorological, and hydrological data that has been collected at the Haig Glacier since 2001. This data is used in conjunction with glacier coverage and hypsometric data for the remainder of the basin to estimate seasonal snow and glacial meltwater contributions to the Bow River from the glacierized fraction of the catchment. The results of this study show the percentage of total flow attributed to glacial melt to be highly variable. Glacier runoff contributes up to an order of magnitude more water to the Bow River per unit area of

  6. Glacial Geology of Wisconsin.

    ERIC Educational Resources Information Center

    Madison Public Schools, WI.

    This publication is a teacher's resource and guidebook for the presentation of the three filmstrips in the "Glacial Geology of Wisconsin" series. The first filmstrip is subtitled, "Evidence of the Glaciers," the second "How the Glaciers Reshaped the Landscape," and the third "Fossils of the Ice Age."…

  7. Molecular data and ecological niche modelling reveal the phylogeographic pattern of Cotinus coggygria (Anacardiaceae) in China's warm-temperate zone.

    PubMed

    Wang, W; Tian, C Y; Li, Y H; Li, Y

    2014-11-01

    The phylogeography of common and widespread species helps to elucidate the history of local flora and vegetation. In this study, we selected Cotinus coggygria, a species widely distributed in China's warm-temperate zone. One chloroplast DNA (cpDNA) region and ecological niche modelling were used to examine the phylogeographic pattern of C. coggygria. The cpDNA data revealed two phylogeographic groups (Southern and Northern) corresponding to the geographic regions. Divergence time analyses revealed that divergence of the two groups occurred at approximately 147,000 years before the present (BP), which coincided with the formation of the downstream area of the Yellow River, indicating that the Yellow River was a weak phylogeographic divide for C. coggygria. The molecular data and ecological niche modelling also indicated that C. coggyria did not experience population expansion after glaciations. This study thus supports the fact that Pleistocene glacial cycles only slightly affected C. coggygria, which survived in situ and occupied multiple localised glacial refugia during glaciations. This finding is contrary to the hypothesis of large-scale range habitat contraction and retreat into a few main refugia. © 2014 German Botanical Society and The Royal Botanical Society of the Netherlands.

  8. The extent and timing of the last British-Irish Ice Sheet offshore of west Ireland-preliminary findings

    NASA Astrophysics Data System (ADS)

    Peters, Jared; Benetti, Sara; Dunlop, Paul; Cofaigh, Colm Ó.

    2014-05-01

    Recently interpreted marine geophysical data from the western Irish shelf has provided the first direct evidence that the last British-Irish Ice Sheet (BIIS) extended westwards onto the Irish continental shelf as a grounded ice mass composed of several lobes with marine-terminating margins. Marine terminating ice margins are known to be sensitive to external forcing mechanisms and currently there is concern regarding the future stability of marine based ice sheets, such as the West Antarctic Ice Sheet, in a warming world. Given its position, the glaciated western Irish continental shelf is a prime location to investigate the processes of how marine-based ice sheets responded to past climatic and oceanic events, which may in turn help us better predict the future trajectory of the marine sectors of modern Ice Sheets. However, despite the potential importance of the former Irish ice margin to our understanding of ice sheet behaviour, the timing and nature of its advance and retreat is currently poorly understood. This study aims to describe the depositional history of the last BIIS on the continental shelf west of Ireland and age-constrain the rate of retreat of two ice lobes that extended from Galway Bay and Clew Bay. This is being accomplished through a multifaceted analysis of at least 29 sediment cores gathered across the continental shelf offshore of counties Galway and Mayo, Ireland. This poster shows results from initial sedimentological descriptions of cores from the mid to outer shelf, which support previous geomorphic interpretations of BIIS history. Preliminary palaeoenvironmental results from ongoing micropaleontological analyses are also discussed and provide new data that verifies sedimentary interpretations on ice proximity. Finally, results from several radiocarbon dates are discussed, which limit these deposits to the last glacial maximum and constrain the timings of ice advance and retreat on the continental shelf west of Ireland.

  9. Revisiting the Iberian honey bee (Apis mellifera iberiensis) contact zone: maternal and genome-wide nuclear variations provide support for secondary contact from historical refugia.

    PubMed

    Chávez-Galarza, Julio; Henriques, Dora; Johnston, J Spencer; Carneiro, Miguel; Rufino, José; Patton, John C; Pinto, M Alice

    2015-06-01

    Dissecting diversity patterns of organisms endemic to Iberia has been truly challenging for a variety of taxa, and the Iberian honey bee is no exception. Surveys of genetic variation in the Iberian honey bee are among the most extensive for any honey bee subspecies. From these, differential and complex patterns of diversity have emerged, which have yet to be fully resolved. Here, we used a genome-wide data set of 309 neutrally tested single nucleotide polymorphisms (SNPs), scattered across the 16 honey bee chromosomes, which were genotyped in 711 haploid males. These SNPs were analysed along with an intergenic locus of the mtDNA, to reveal historical patterns of population structure across the entire range of the Iberian honey bee. Overall, patterns of population structure inferred from nuclear loci by multiple clustering approaches and geographic cline analysis were consistent with two major clusters forming a well-defined cline that bisects Iberia along a northeastern-southwestern axis, a pattern that remarkably parallels that of the mtDNA. While a mechanism of primary intergradation or isolation by distance could explain the observed clinal variation, our results are more consistent with an alternative model of secondary contact between divergent populations previously isolated in glacial refugia, as proposed for a growing list of other Iberian taxa. Despite current intense honey bee management, human-mediated processes have seemingly played a minor role in shaping Iberian honey bee genetic structure. This study highlights the complexity of the Iberian honey bee patterns and reinforces the importance of Iberia as a reservoir of Apis mellifera diversity. © 2015 John Wiley & Sons Ltd.

  10. Fertile fathoms: Deep reproductive refugia for threatened shallow corals

    PubMed Central

    Holstein, Daniel M.; Smith, Tyler B.; Gyory, Joanna; Paris, Claire B.

    2015-01-01

    The persistence of natural metapopulations may depend on subpopulations that exist at the edges of species ranges, removed from anthropogenic stress. Mesophotic coral ecosystems (30–150 m) are buffered from disturbance by depth and distance, and are potentially massive reservoirs of coral diversity and fecundity; yet we know little about the reproductive capabilities of their constituent species and the potential for these marginal environments to influence patterns of coral reef persistence. We investigated the reproductive performance of the threatened depth-generalist coral Orbicella faveolata over the extent of its vertical range to assess mesophotic contributions to regional larval pools. Over equal habitat area, mesophotic coral populations were found to produce over an order of magnitude more eggs than nearby shallow populations. Positive changes with depth in both population abundance and polyp fecundity contributed to this discrepancy. Relative larval pool contributions of deeper living corals will likely increase as shallow habitats further degrade due to climate change and local habitat degradation. This is a compelling example of the potential for marginal habitat to be critical to metapopulation persistence as reproductive refugia. PMID:26196243

  11. ESR Dating Research of Glacial Tills in Tibetan Plateau

    NASA Astrophysics Data System (ADS)

    Bi, W.; Yi, C.

    2016-12-01

    In recent years, Quaternary Glacial-chronology has been made remarkable progress in the Tibetan Platean(TP) with the development of several numeric dating techniques, such as cosmogenic nuclides(NC), optically stimulated luminescence(OSL) and 14C. In constrast, the dating of Quaternary glacial tills in 100,000 years even more than million-year has been a challenge, just because the techniques has defects themselves and the sediments were stransformed during the geological and geomorphology progress later. Electron Spin Resonance(ESR) has been becoming one of the key methods of Quaternary Glacial-chronology with wide range of dating, expecially for the sample older than 100,000 years up to million-year scale. The accurate measurement of equivalent dose significantly impacts on accuracy and reliability of ESR dating method. Therefore, the study of the mechanisms of resetting processes is fundamental for accurate and reliable ESR dating. To understand the mechanism and characteristics of quartz ESR signal resetting of different samples, a series of laboratory simulation and field observation studies were carried out, which made lots of important breakthrough. But the research in quartz ESR signal of moraines is less and the test of ESR dating method is still in the qualitative investigation. Therefor, we use ESR dating and study on the mechanism and characteristics of quartz ESR signals in tills in the Tibetan Platean. In the adjust method of Modern, the quartz ESR signals in Modern glacial tills represent residual values which can be adjusted signals in the older glacial tills. As a consequence, ESR dating of the quartz in moraines needs to be explored in deep with building models to adjust ages which are measured by ESR dating. Therefore, ESR dating will become the trusted one of the cross dating methods in Quaternary Glacial-chronology with the adjust mothod improving the accuracy of ESR dating ages.

  12. Interannual physiological and growth responses of glacial Juniperus to changes in atmospheric [CO2] since the Last Glacial Maximum

    NASA Astrophysics Data System (ADS)

    Gerhart, L. M.; Harris, J. M.; Ward, J. K.

    2011-12-01

    During the Last Glacial Maximum, atmospheric [CO2] was as low as 180 ppm and has currently risen to a modern value of 393 ppm as a result of fossil fuel combustion and deforestation. In order to understand how changing [CO2] influenced trees over the last 50,000 years, we analyzed carbon isotope ratios and width of individual tree rings from glacial Juniperus specimens preserved in the Rancho La Brea tar pits in southern California (aged 14-49 kyr BP). Modern trees were also analyzed to compare effects of changing precipitation, temperature and atmospheric [CO2] on physiology and growth. To assess physiological responses, we calculated ci/ca (intercellular [CO2]/atmospheric [CO2]) for each annual ring of each tree. This ratio incorporates numerous aspects of plant physiology, including stomatal conductance and photosynthetic capacity. In addition, we measured ring widths for each sample, and standardized these measurements into indices in order to compare across individuals. Mean ci/ca values remained constant throughout 50,000 years despite major environmental changes, indicating a long-term physiological set point for ci/ca in this group. Constant ci/ca ratios would be maintained through offsetting changes in stomatal conductance and photosynthetic capacity. Glacial Juniperus never experienced ci values below 90 ppm, suggesting a survival compensation point for Juniperus. In addition, glacial trees showed significantly reduced interannual variation in ci/ca, even though interannual climatic variability was as high during the LGM in this region as it is today. A lack of variability in ci/ca of glacial trees suggests that tree physiology was dominated by low [CO2], which shows low interannual variation. Modern trees showed high interannual variation in ci/ca, since water availability dominates current physiological responses and varies greatly from year to year. Interestingly, interannual variation in ring width index did not show significant differences between

  13. Biolabile ferrous iron bearing nanoparticles in glacial sediments

    NASA Astrophysics Data System (ADS)

    Hawkings, Jon R.; Benning, Liane G.; Raiswell, Rob; Kaulich, Burkhard; Araki, Tohru; Abyaneh, Majid; Stockdale, Anthony; Koch-Müller, Monika; Wadham, Jemma L.; Tranter, Martyn

    2018-07-01

    Glaciers and ice sheets are a significant source of nanoparticulate Fe, which is potentially important in sustaining the high productivity observed in the near-coastal regions proximal to terrestrial ice cover. However, the bioavailability of particulate iron is poorly understood, despite its importance in the ocean Fe inventory. We combined high-resolution imaging and spectroscopy to investigate the abundance, morphology and valence state of particulate iron in glacial sediments. Our results document the widespread occurrence of amorphous and Fe(II)-rich and Fe(II)-bearing nanoparticles in Arctic glacial meltwaters and iceberg debris, compared to Fe(III)-rich dominated particulates in an aeolian dust sample. Fe(II) is thought to be highly biolabile in marine environments. Our work shows that glacially derived Fe is more labile than previously assumed, and consequently that glaciers and ice sheets are therefore able to export potentially bioavailable Fe(II)-containing nanoparticulate material to downstream ecosystems, including those in a marine setting. Our findings provide further evidence that Greenland Ice Sheet meltwaters may provide biolabile particulate Fe that may fuel the large summer phytoplankton bloom in the Labrador Sea, and that Fe(II)-rich particulates from a region of very high productivity downstream of a polar ice sheet may be glacial in origin.

  14. Glacial geology of the Hellas region on Mars

    NASA Technical Reports Server (NTRS)

    Kargel, Jeffrey S.; Strom, Robert G.; Johnson, Natasha

    1991-01-01

    A glacial geologic interpretation was recently presented for Argyre, which is herein extended to Hellas. This glacial event is believed to constitute an important link in a global cryohydric epoch of Middle Amazonian age. At glacial maximum, ice apparently extended far beyond the regions of Argyre and Hellas, and formed what is termed as the Austral Ice Sheet, an agglomeration of several ice domes and lobes including the Hellas Lobe. It is concluded that Hellas was apparently heavily glaciated. Also glaciation was young by Martian standards (Middle Amazonian), and ancient by terrestrial standards. Glaciation appears to have occurred during the same period that other areas on Mars were experiencing glaciation and periglacial activity. Glaciation seems to have occurred as a geological brief epoch of intense geomorphic activity in an era characterized by long periods of relative inactivity.

  15. Full Mitogenomes in the Critically Endangered Kākāpō Reveal Major Post-Glacial and Anthropogenic Effects on Neutral Genetic Diversity.

    PubMed

    Dussex, Nicolas; von Seth, Johanna; Robertson, Bruce C; Dalén, Love

    2018-04-19

    Understanding how species respond to population declines is a central question in conservation and evolutionary biology. Population declines are often associated with loss of genetic diversity, inbreeding and accumulation of deleterious mutations, which can lead to a reduction in fitness and subsequently contribute to extinction. Using temporal approaches can help us understand the effects of population declines on genetic diversity in real time. Sequencing pre-decline as well as post-decline mitogenomes representing all the remaining mitochondrial diversity, we estimated the loss of genetic diversity in the critically endangered kākāpō ( Strigops habroptilus ). We detected a signal of population expansion coinciding with the end of the Pleistocene last glacial maximum (LGM). Also, we found some evidence for northern and southern lineages, supporting the hypothesis that the species may have been restricted to isolated northern and southern refugia during the LGM. We observed an important loss of neutral genetic diversity associated with European settlement in New Zealand but we could not exclude a population decline associated with Polynesian settlement in New Zealand. However, we did not find evidence for fixation of deleterious mutations. We argue that despite high pre-decline genetic diversity, a rapid and range-wide decline combined with the lek mating system, and life-history traits of kākāpō contributed to a rapid loss of genetic diversity following severe population declines.

  16. Glacial Lake Expansion in the Central Himalayas by Landsat Images, 1990–2010

    PubMed Central

    Nie, Yong; Liu, Qiao; Liu, Shiyin

    2013-01-01

    Glacial lake outburst flood (GLOF) is a serious hazard in high, mountainous regions. In the Himalayas, catastrophic risks of GLOFs have increased in recent years because most Himalayan glaciers have experienced remarkable downwasting under a warming climate. However, current knowledge about the distribution and recent changes in glacial lakes within the central Himalaya mountain range is still limited. Here, we conducted a systematic investigation of the glacial lakes within the entire central Himalaya range by using an object-oriented image processing method based on the Landsat Thematic Mapper (TM) or Enhanced Thematic Mapper (ETM) images from 1990 to 2010. We extracted the lake boundaries for four time points (1990, 2000, 2005 and 2010) and used a time series inspection method combined with a consistent spatial resolution of Landsat images that consistently revealed lake expansion. Our results show that the glacial lakes expanded rapidly by 17.11% from 1990 to 2010. The pre-existing, larger glacial lakes, rather than the newly formed lakes, contributed most to the areal expansion. The greatest expansions occurred at the altitudinal zones between 4800 m and 5600 m at the north side of the main Himalayan range and between 4500 m and 5600 m at the south side, respectively. Based on the expansion rate, area and type of glacial lakes, we identified 67 rapidly expanding glacial lakes in the central Himalayan region that need to be closely monitored in the future. The warming and increasing amounts of light-absorbing constituents of snow and ice could have accelerated the melting that directly affected the glacial lake expansion. Across the main central Himalayas, glacial lakes at the north side show more remarkable expansion than those at the south side. An effective monitoring and warning system for critical glacial lakes is urgently needed. PMID:24376778

  17. The early rise and late demise of New Zealand's last glacial maximum.

    PubMed

    Rother, Henrik; Fink, David; Shulmeister, James; Mifsud, Charles; Evans, Michael; Pugh, Jeremy

    2014-08-12

    Recent debate on records of southern midlatitude glaciation has focused on reconstructing glacier dynamics during the last glacial termination, with different results supporting both in-phase and out-of-phase correlations with Northern Hemisphere glacial signals. A continuing major weakness in this debate is the lack of robust data, particularly from the early and maximum phase of southern midlatitude glaciation (∼30-20 ka), to verify the competing models. Here we present a suite of 58 cosmogenic exposure ages from 17 last-glacial ice limits in the Rangitata Valley of New Zealand, capturing an extensive record of glacial oscillations between 28-16 ka. The sequence shows that the local last glacial maximum in this region occurred shortly before 28 ka, followed by several successively less extensive ice readvances between 26-19 ka. The onset of Termination 1 and the ensuing glacial retreat is preserved in exceptional detail through numerous recessional moraines, indicating that ice retreat between 19-16 ka was very gradual. Extensive valley glaciers survived in the Rangitata catchment until at least 15.8 ka. These findings preclude the previously inferred rapid climate-driven ice retreat in the Southern Alps after the onset of Termination 1. Our record documents an early last glacial maximum, an overall trend of diminishing ice volume in New Zealand between 28-20 ka, and gradual deglaciation until at least 15 ka.

  18. 76 FR 50476 - Application To Export Electric Energy; Glacial Energy of Texas, Inc.

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-08-15

    ... DEPARTMENT OF ENERGY [OE Docket No. EA-382] Application To Export Electric Energy; Glacial Energy of Texas, Inc. AGENCY: Office of Electricity Delivery and Energy Reliability, DOE. ACTION: Notice of Application. SUMMARY: Glacial Energy of Texas, Inc. (Glacial) has applied for authority to transmit electric...

  19. Glacial ocean circulation and stratification explained by reduced atmospheric temperature.

    PubMed

    Jansen, Malte F

    2017-01-03

    Earth's climate has undergone dramatic shifts between glacial and interglacial time periods, with high-latitude temperature changes on the order of 5-10 °C. These climatic shifts have been associated with major rearrangements in the deep ocean circulation and stratification, which have likely played an important role in the observed atmospheric carbon dioxide swings by affecting the partitioning of carbon between the atmosphere and the ocean. The mechanisms by which the deep ocean circulation changed, however, are still unclear and represent a major challenge to our understanding of glacial climates. This study shows that various inferred changes in the deep ocean circulation and stratification between glacial and interglacial climates can be interpreted as a direct consequence of atmospheric temperature differences. Colder atmospheric temperatures lead to increased sea ice cover and formation rate around Antarctica. The associated enhanced brine rejection leads to a strongly increased deep ocean stratification, consistent with high abyssal salinities inferred for the last glacial maximum. The increased stratification goes together with a weakening and shoaling of the interhemispheric overturning circulation, again consistent with proxy evidence for the last glacial. The shallower interhemispheric overturning circulation makes room for slowly moving water of Antarctic origin, which explains the observed middepth radiocarbon age maximum and may play an important role in ocean carbon storage.

  20. Tracing an invasion: landbridges, refugia, and the phylogeography of the Neotropical rattlesnake (Serpentes: Viperidae: Crotalus durissus).

    PubMed

    Wüster, Wolfgang; Ferguson, Julia E; Quijada-Mascareñas, J Adrian; Pook, Catharine E; Salomão, Maria da Graça; Thorpe, Roger S

    2005-04-01

    Abstract Pleistocene fragmentation of the Amazonian rainforest has been hypothesized to be a major cause of Neotropical speciation and diversity. However, the role and even the reality of Pleistocene forest refugia have attracted much scepticism. In Amazonia, previous phylogeographical studies have focused mostly on organisms found in the forests themselves, and generally found speciation events to have predated the Pleistocene. However, molecular studies of open-formation taxa found both north and south of the Amazonian forests, probably because of vicariance resulting from expansion of the rainforests, may provide novel insights into the age of continuous forest cover across the Amazon basin. Here, we analyse three mitochondrial genes to infer the phylogeography of one such trans-Amazonian vicariant, the Neotropical rattlesnake (Crotalus durissus), which occupies primarily seasonal formations from Mexico to Argentina, but avoids the rainforests of Central and tropical South America. The phylogeographical pattern is consistent with gradual dispersal along the Central American Isthmus, followed by more rapid dispersal into and across South America after the uplift of the Isthmus of Panama. Low sequence divergence between populations from north and south of the Amazon rainforest is consistent with mid-Pleistocene divergence, approximately 1.1 million years ago (Ma). This suggests that the Amazonian rainforests must have become fragmented or at least shrunk considerably during that period, lending support to the Pleistocene refugia theory as an important cause of distribution patterns, if not necessarily speciation, in Amazonian forest organisms. These results highlight the potential of nonforest species to contribute to an understanding of the history of the Amazonian rainforests themselves.

  1. Inventory and recently increasing GLOF susceptibility of glacial lakes in Sikkim, Eastern Himalaya

    NASA Astrophysics Data System (ADS)

    Aggarwal, Suruchi; Rai, S. C.; Thakur, P. K.; Emmer, Adam

    2017-10-01

    Climatic changes alter the climate system, leading to a decrease of glacier mass volumes and swelling glacial lakes. This study provides a new inventory of glacial and high-altitude lakes for Sikkim, Eastern Himalaya, and evaluates the susceptibility of lakes to Glacial Lake Outburst Flood (GLOF). By using satellite data of high spatial resolution (5 m), we obtain 1104 glacial and high-altitude lakes with total area 30.498 km2, of which 472 have an area > 0.01 km2. Applying pre-defined GLOF susceptibility criteria on these 472 lakes yields 21 lakes susceptible to GLOF, which all increased in area from 1972-2015. Using Analytic Hierarchy Processes (AHP), the pairwise comparison matrix further reveals that 5 of these glacial lakes have low, 14 have medium and 2 have high GLOF susceptibility. Especially these 16 glacial lakes with high and medium GLOF susceptibility may threaten downstream communities and infrastructure and need further attention.

  2. Strong and deep Atlantic meridional overturning circulation during the last glacial cycle.

    PubMed

    Böhm, E; Lippold, J; Gutjahr, M; Frank, M; Blaser, P; Antz, B; Fohlmeister, J; Frank, N; Andersen, M B; Deininger, M

    2015-01-01

    Extreme, abrupt Northern Hemisphere climate oscillations during the last glacial cycle (140,000 years ago to present) were modulated by changes in ocean circulation and atmospheric forcing. However, the variability of the Atlantic meridional overturning circulation (AMOC), which has a role in controlling heat transport from low to high latitudes and in ocean CO2 storage, is still poorly constrained beyond the Last Glacial Maximum. Here we show that a deep and vigorous overturning circulation mode has persisted for most of the last glacial cycle, dominating ocean circulation in the Atlantic, whereas a shallower glacial mode with southern-sourced waters filling the deep western North Atlantic prevailed during glacial maxima. Our results are based on a reconstruction of both the strength and the direction of the AMOC during the last glacial cycle from a highly resolved marine sedimentary record in the deep western North Atlantic. Parallel measurements of two independent chemical water tracers (the isotope ratios of (231)Pa/(230)Th and (143)Nd/(144)Nd), which are not directly affected by changes in the global cycle, reveal consistent responses of the AMOC during the last two glacial terminations. Any significant deviations from this configuration, resulting in slowdowns of the AMOC, were restricted to centennial-scale excursions during catastrophic iceberg discharges of the Heinrich stadials. Severe and multicentennial weakening of North Atlantic Deep Water formation occurred only during Heinrich stadials close to glacial maxima with increased ice coverage, probably as a result of increased fresh-water input. In contrast, the AMOC was relatively insensitive to submillennial meltwater pulses during warmer climate states, and an active AMOC prevailed during Dansgaard-Oeschger interstadials (Greenland warm periods).

  3. Cyberbullying, Schools and the Law: A Comparative Study in Northern Ireland and the Republic of Ireland

    ERIC Educational Resources Information Center

    Purdy, Noel; Mc Guckin, Conor

    2015-01-01

    Background: This study addresses the fast developing behavioural issue of cyberbullying in schools and its complex legal context. Purpose: This study set out to investigate teachers' perceptions of the extent of cyberbullying and the extent to which school leaders in Northern Ireland and the Republic of Ireland feel knowledgeable and confident…

  4. A glacial record of the last termination in the southern tropical Andes

    NASA Astrophysics Data System (ADS)

    Bromley, G. R.; Schaefer, J. M.; Winckler, G.; Hall, B. L.; Todd, C. E.; Rademaker, K.

    2012-12-01

    The last glacial termination represents the highest-magnitude climate change of the last hundred thousand years. Accurate resolution of events during the termination is vital to our understanding of how - and why - the global climate system transitions from a full glacial to interglacial state, as well as the causes of abrupt climate change during the late-glacial period. Palaeoclimate data from low latitudes, though relatively sparse, are particularly valuable, since the tropical ocean and atmosphere likely play a crucial role in Quaternary climate variability on all timescales. We present a detailed glacier record from the Andes of southern Peru (15°S), resolved with 3He surface-exposure dating and spanning the last glacial maximum and termination. Our dataset reveals that glaciers in this part of the Southern Hemisphere maintained their Late Pleistocene maxima for several millennia and that the onset of the termination may have occurred relatively late. Deglaciation was punctuated by two major advances during the late-glacial period. Following the glacial-interglacial transition, our preliminary chronologic and morphologic data suggest that, in contrast to the Northern Hemisphere, glaciers in the southern tropical Andes have experienced overall shrinkage during the Holocene.

  5. Multiscale thermal refugia and stream habitat associations of chinook salmon in northwestern Oregon

    USGS Publications Warehouse

    Torgersen, Christian E.; Price, David M.; Li, Hiram W.; McIntosh, B.A.

    1999-01-01

    We quantified distribution and behavior of adult spring chinook salmon (Oncorhynchus tshawytscha) related to patterns of stream temperature and physical habitat at channel-unit, reach-, and section-level spatial scales in a wilderness stream and a disturbed stream in the John Day River basin in northeastern Oregon. We investigated the effectiveness of thermal remote sensing for analyzing spatial patterns of stream temperature and assessed habitat selection by spring chinook salmon, evaluating whether thermal refugia might be responsible for the persistence of these stocks in rivers where water temperatures frequently exceed their upper tolerance levels (25A?C) during spawning migration. By presenting stream temperature and the ecology of chinook salmon in a historical context, we could evaluate how changes in riverine habitat and thermal spatial structure, which can be caused by land-use practices, may influence distributional patterns of chinook salmon. Thermal remote sensing provided spatially continuous maps of stream temperature for reaches used by chinook salmon in the upper subbasins of the Middle Fork and North Fork John Day River. Electivity analysis and logistic regression were used to test for associations between the longitudinal distribution of salmon and cool-water areas and stream habitat characteristics. Chinook salmon were distributed nonuniformly in reaches throughout each stream. Salmon distribution and cool water temperature patterns were most strongly related at reach-level spatial scales in the warm stream, the Middle Fork (maximum likelihood ratio: P 0.30). Pools were preferred by adult chinook salmon in both subbasins (Bonferroni confidence interval: P a?? 0.05); however, riffles were used proportionately more frequently in the North Fork than in the Middle Fork. Our observations of thermal refugia and their use by chinook salmon at multiple spatial scales reveal that, although heterogeneity in the longitudinal stream temperature profile may

  6. Aspects of agricultural land use in Ireland

    NASA Astrophysics Data System (ADS)

    Lee, J.

    1986-02-01

    Ireland's soil regions consist largely of Luvisols, Cambisols, and Gleysols. Approximately 60% of Ireland's land area is subject to varying degrees of soil limitations. Twenty-five percent of the land area comprises wet lowland mineral soils. Ninety percent of Ireland's agricultural area comprises pasture, hay, and silage. Approximately 30% of the agricultural area is devoted to dairying, and 55% to cattle production. is devoted to dairying, and 55% to cattle production. Trends in agricultural land use indicate that tillage declined substantially while livestock showed a substantial increase particularly in the decade 1965 1975. Research concludes that over 2.8 million ha has a capacity to carry at least 100 LU/40 ha (100 acres). Levels of fertilizer use in Ireland are below EEC levels. The highest fertilizer use levels are associated with the eastern and southern areas of Ireland. Tillage crops occupy only 10% of the agricultural area, while they account for 26% of tertilizer and lime use.

  7. Glacial inception during the late Holocene without carbon emissions from early agriculture: lessons from the stage-19 glacial inception

    NASA Astrophysics Data System (ADS)

    He, F.; Vavrus, S. J.; Kutzbach, J. E.; Ruddiman, W. F.; Tzedakis, P. C.

    2013-12-01

    Decreases in orbitally-forced summer insolation along with downward trends in greenhouse gases (GHG) have been precursors to incipient glaciation in the past. In the last several thousand years of the current interglacial, while summer insolation has decreased, there was a reversal of the downward trends in CH4 and CO2 concentration within the Holocene around 5,000 and 7,000 years ago. While the cause of this reversal remains unresolved, a leading hypothesis is Ruddiman's Early Anthropogenic Hypothesis that early agriculture, starting several thousand years ago, caused emissions of GHG large enough to reverse natural downward trends in CO2 and CH4 and kept Earth's climate anomalously warm, with the corollary that this may have prevented incipient glaciation during the late Holocene. Here we use the 1-degree, fully coupled Community Climate System Model version 4 (CCSM4) with climate forcings (orbital parameters and GHG) of a previous glacial inception to investigate whether glacial inception should have occurred prior to the industrial revolution if the concentrations of CH4 and CO2 had followed their natural downward trends throughout the Holocene. Tzedakis et al. [2012] show that for the previous eight interglacials, Stage 11 and Stage 19 are the best analogs of the Holocene because of their low eccentricities, and Stage 19 is a better analog than Stage 11 for the Holocene due to the in-phase relationship between obliquity and precession. Furthermore, their study suggests that 777 ka BP (777,000 years before present) is the timing of glacial inception for Stage 19, based on the occurrence of the earliest bipolar seesaw event associated with glacial melting. Not only do the orbital parameters at 777 ka BP resemble pre-industrial conditions, but the concentrations of CO2 at that time were essentially the same as their expected 'natural' pre-industrial values in the absence of anthropogenic greenhouse emissions. Our multi-millennial coupled CCSM4 simulations show

  8. Breakup of last glacial deep stratification in the South Pacific.

    PubMed

    Basak, Chandranath; Fröllje, Henning; Lamy, Frank; Gersonde, Rainer; Benz, Verena; Anderson, Robert F; Molina-Kescher, Mario; Pahnke, Katharina

    2018-02-23

    Stratification of the deep Southern Ocean during the Last Glacial Maximum is thought to have facilitated carbon storage and subsequent release during the deglaciation as stratification broke down, contributing to atmospheric CO 2 rise. Here, we present neodymium isotope evidence from deep to abyssal waters in the South Pacific that confirms stratification of the deepwater column during the Last Glacial Maximum. The results indicate a glacial northward expansion of Ross Sea Bottom Water and a Southern Hemisphere climate trigger for the deglacial breakup of deep stratification. It highlights the important role of abyssal waters in sustaining a deep glacial carbon reservoir and Southern Hemisphere climate change as a prerequisite for the destabilization of the water column and hence the deglacial release of sequestered CO 2 through upwelling. Copyright © 2018 The Authors, some rights reserved; exclusive licensee American Association for the Advancement of Science. No claim to original U.S. Government Works.

  9. Linking glacial erosion and low-relief landscapes in tropical orogens

    NASA Astrophysics Data System (ADS)

    Cunningham, M.; Stark, C. P.; Kaplan, M. R.; Schaefer, J. M.; Galewsky, J.; Yoo, J.

    2015-12-01

    One significant way that climate influences orogenic evolution is by modulating glacial erosion. At mid-latitudes it is hypothesized that this climate-tectonic interplay is so strong that a "glacial buzzsaw" acting throughout the Quaternary outpaced tectonic uplift in most mountain belts and concentrated topography in a zone defined by the bounds of ELA fluctuation. Less attention has been paid to how the buzzsaw might manifest itself at low latitudes, where many mountain belts are just high enough to have been glaciated at the LGM but today sit well below the ELA. We have focused on the glacial history of Costa Rica and Taiwan, where we find evidence of ice cap erosion coincident with low-relief landscapes near the LGM ELA. Previous attempts to understand the formation of these perched, low-relief landscapes has mostly concerned interactions between fluvial erosion and geodynamics. Our work aims instead to describe the role that glacial erosion played in the evolution of these landscapes, and how they fit in the buzzsaw paradigm. At Cerro Chirripó in Costa Rica we use 10-Be surface exposure age dating of moraine boulders and scoured bedrock, field mapping, and remote sensing to constrain the timing, areal extent, and pattern of glacial erosion. We made similar observations of ice extent at Nanhudashan in Taiwan, where surface exposure age dating has previously been applied to glacial landforms (e.g. Hebenstreit et al., 2011; Siame et al., 2007). In Costa Rica, our 10-Be dates from scoured bedrock near the highest peak and terminal/lateral moraines show signs of ice-cap erosion until 22 ka. Similar arguments for LGM ice cap erosion have been made for Nanhudashan. Regional climate simulations (WRF) further constrain the timing and spatial extent of glaciation in these places, and the combination of field data and climate modeling will inform estimates of the magnitude of glacial erosion on perched landscapes.

  10. In and out of glacial extremes by way of dust‑climate feedbacks

    NASA Astrophysics Data System (ADS)

    Shaffer, Gary; Lambert, Fabrice

    2018-03-01

    Mineral dust aerosols cool Earth directly by scattering incoming solar radiation and indirectly by affecting clouds and biogeochemical cycles. Recent Earth history has featured quasi-100,000-y, glacial‑interglacial climate cycles with lower/higher temperatures and greenhouse gas concentrations during glacials/interglacials. Global average, glacial maxima dust levels were more than 3 times higher than during interglacials, thereby contributing to glacial cooling. However, the timing, strength, and overall role of dust‑climate feedbacks over these cycles remain unclear. Here we use dust deposition data and temperature reconstructions from ice sheet, ocean sediment, and land archives to construct dust‑climate relationships. Although absolute dust deposition rates vary greatly among these archives, they all exhibit striking, nonlinear increases toward coldest glacial conditions. From these relationships and reconstructed temperature time series, we diagnose glacial‑interglacial time series of dust radiative forcing and iron fertilization of ocean biota, and use these time series to force Earth system model simulations. The results of these simulations show that dust‑climate feedbacks, perhaps set off by orbital forcing, push the system in and out of extreme cold conditions such as glacial maxima. Without these dust effects, glacial temperature and atmospheric CO2 concentrations would have been much more stable at higher, intermediate glacial levels. The structure of residual anomalies over the glacial‑interglacial climate cycles after subtraction of dust effects provides constraints for the strength and timing of other processes governing these cycles.

  11. In and out of glacial extremes by way of dust−climate feedbacks

    PubMed Central

    Lambert, Fabrice

    2018-01-01

    Mineral dust aerosols cool Earth directly by scattering incoming solar radiation and indirectly by affecting clouds and biogeochemical cycles. Recent Earth history has featured quasi-100,000-y, glacial−interglacial climate cycles with lower/higher temperatures and greenhouse gas concentrations during glacials/interglacials. Global average, glacial maxima dust levels were more than 3 times higher than during interglacials, thereby contributing to glacial cooling. However, the timing, strength, and overall role of dust−climate feedbacks over these cycles remain unclear. Here we use dust deposition data and temperature reconstructions from ice sheet, ocean sediment, and land archives to construct dust−climate relationships. Although absolute dust deposition rates vary greatly among these archives, they all exhibit striking, nonlinear increases toward coldest glacial conditions. From these relationships and reconstructed temperature time series, we diagnose glacial−interglacial time series of dust radiative forcing and iron fertilization of ocean biota, and use these time series to force Earth system model simulations. The results of these simulations show that dust−climate feedbacks, perhaps set off by orbital forcing, push the system in and out of extreme cold conditions such as glacial maxima. Without these dust effects, glacial temperature and atmospheric CO2 concentrations would have been much more stable at higher, intermediate glacial levels. The structure of residual anomalies over the glacial−interglacial climate cycles after subtraction of dust effects provides constraints for the strength and timing of other processes governing these cycles. PMID:29440407

  12. Abrupt glacial climate shifts controlled by ice sheet changes.

    PubMed

    Zhang, Xu; Lohmann, Gerrit; Knorr, Gregor; Purcell, Conor

    2014-08-21

    During glacial periods of the Late Pleistocene, an abundance of proxy data demonstrates the existence of large and repeated millennial-scale warming episodes, known as Dansgaard-Oeschger (DO) events. This ubiquitous feature of rapid glacial climate change can be extended back as far as 800,000 years before present (BP) in the ice core record, and has drawn broad attention within the science and policy-making communities alike. Many studies have been dedicated to investigating the underlying causes of these changes, but no coherent mechanism has yet been identified. Here we show, by using a comprehensive fully coupled model, that gradual changes in the height of the Northern Hemisphere ice sheets (NHISs) can alter the coupled atmosphere-ocean system and cause rapid glacial climate shifts closely resembling DO events. The simulated global climate responses--including abrupt warming in the North Atlantic, a northward shift of the tropical rainbelts, and Southern Hemisphere cooling related to the bipolar seesaw--are generally consistent with empirical evidence. As a result of the coexistence of two glacial ocean circulation states at intermediate heights of the ice sheets, minor changes in the height of the NHISs and the amount of atmospheric CO2 can trigger the rapid climate transitions via a local positive atmosphere-ocean-sea-ice feedback in the North Atlantic. Our results, although based on a single model, thus provide a coherent concept for understanding the recorded millennial-scale variability and abrupt climate changes in the coupled atmosphere-ocean system, as well as their linkages to the volume of the intermediate ice sheets during glacials.

  13. Springs as hydrologic refugia in a changing climate? A remote sensing approach

    USGS Publications Warehouse

    Cartwright, Jennifer M.; Johnson, Henry M.

    2018-01-01

    Spring‐fed wetlands are ecologically important habitats in arid and semi‐arid regions. Springs have been suggested as possible hydrologic refugia from droughts and climate change; however, springs that depend on recent precipitation or snowmelt for recharge may be vulnerable to warming and drought intensification. Springs that are expected to maintain their ecohydrologic function in a warmer, drier climate may be priorities for conservation and restoration. Identifying such springs is difficult because many springs lack hydrologic records of adequate temporal extent and resolution to assess their resilience to water cycle changes. This study demonstrates proof‐of‐concept for the assessment of certain spring types (i.e., helocrene, hypocrene, and hillslope springs) in terms of hydrologic and ecological resilience to climatic water stress using freely available remote‐sensing and climate data. We used the Normalized Difference Vegetation Index (NDVI) from 1985 through 2011 to delineate surface‐moisture zones (SMZs) associated with 39 clusters of 172 springs in a montane sage‐steppe landscape in southeastern Oregon, USA. We developed and synthesized seven NDVI‐based indicators of SMZ resilience to interannual changes in water availability: (1) mean and (2) standard deviation of July NDVI; (3) mean difference in July NDVI and (4) difference in coefficient of variation for July NDVI between each SMZ and its surrounding watershed; (5) response of SMZ July NDVI to 90‐day antecedent precipitation; (6) response of SMZ July NDVI to the previous winter's snowpack; and (7) range of NDVI values from an exceptionally wet year followed by three dry years. Because all resilience indicators were highly inter‐correlated, we derived an overall metric of SMZ resilience using principal components analysis that accounted for 66% of total variance. This overall resilience score was positively correlated with SMZ elevation, slope, mean annual precipitation, and with

  14. 20th-century glacial-marine sedimentation in Vitus Lake, Bering Glacier, Alaska, U.S.A.

    USGS Publications Warehouse

    Molnia, B.F.; Post, A.; Carlson, P.R.

    1996-01-01

    Vitus Lake, the ice-marginal basin at the southeastern edge of Bering Glacier, Alaska, U.S.A., is a site of modern, rapid, glacial-marine sedimentation. Rather than being a fresh-water lake, Vitus Lake is a tidally influenced, marine to brackish embayment connected to the Pacific Ocean by an inlet, the Seal River. Vitus Lake consists of five deep bedrock basins, separated by interbasinal highs. Glacial erosion has cut these basins as much as 250 m below sea level. High-resolution seismic reflection surveys conducted in 1991 and 1993 of four of Vitus Lake's basins reveal a complex, variable three-component acoustic stratigraphy. Although not fully sampled, the stratigraphy is inferred to be primarily glacial-marine units of (1) basal contorted and deformed glacial-marine and glacial sediments deposited by basal ice-contact processes and submarine mass-wasting; (2) acoustically well-stratified glacial-marine sediment, which unconformably overlies the basal unit and which grades upward into (3) acoustically transparent or nearly transparent glacial-marine sediment. Maximum thicknesses of conformable glacial-marine sediment exceed 100 m. All of the acoustically transparent and stratified deposits in Vitus Lake are modern in age, having accumulated between 1967 and 1993. The basins where these three-part sequences of "present-day" glacial-marine sediment are accumulating are themselves cut into older sequences of stratified glacial and glacial-marine deposits. These older units outcrop on the islands in Vitus Lake. In 1967, as the result of a major surge, glacier ice completely filled all five basins. Subsequent terminus retreat, which continued through August 1993, exposed these basins, providing new locations for glacial-marine sediment accumulation. A correlation of sediment thicknesses measured from seismic profiles at specific locations within the basins, with the year that each location became ice-free, shows that the sediment accumulation at some locations

  15. Glacial and Quaternary geology of the northern Yellowstone area, Montana and Wyoming

    USGS Publications Warehouse

    Pierce, Kenneth L.; Licciardi, Joseph M.; Krause, Teresa R.; Whitlock, Cathy

    2014-01-01

    This field guide focuses on the glacial geology and paleoecology beginning in the Paradise Valley and progressing southward into northern Yellowstone National Park. During the last (Pinedale) glaciation, the northern Yellowstone outlet glacier flowed out of Yellowstone Park and down the Yellowstone River Valley into the Paradise Valley. The field trip will traverse the following Pinedale glacial sequence: (1) deposition of the Eightmile terminal moraines and outwash 16.5 ± 1.4 10Be ka in the Paradise Valley; (2) glacial recession of ~8 km and deposition of the Chico moraines and outwash 16.1 ± 1.7 10Be ka; (3) glacial recession of 45 km to near the northern Yellowstone boundary and moraine deposition during the Deckard Flats readjustment 14.2 ± 1.2 10Be ka; and (4) glacial recession of ~37 km and deposition of the Junction Butte moraines 15.2 ± 1.3 10Be ka (this age is a little too old based on the stratigraphic sequence). Yellowstone's northern range of sagebrush-grasslands and bison, elk, wolf, and bear inhabitants is founded on glacial moraines, sub-glacial till, and outwash deposited during the last glaciation. Floods released from glacially dammed lakes and a landslide-dammed lake punctuate this record. The glacial geologic reconstruction was evaluated by calculation of basal shear stress, and yielded the following values for flow pattern in plan view: strongly converging—1.21 ± 0.12 bars (n = 15); nearly uniform—1.04 ± 0.16 bars (n = 11); and strongly diverging—0.84 ± 0.14 bars (n = 16). Reconstructed mass balance yielded accumulation and ablation each of ~3 km3/yr, with glacial movement near the equilibrium line altitude dominated by basal sliding. Pollen and charcoal records from three lakes in northern Yellowstone provide information on the postglacial vegetation and fire history. Following glacial retreat, sparsely vegetated landscapes were colonized first by spruce parkland and then by closed subalpine forests. Regional fire activity

  16. Lizards on Ice: Evidence for Multiple Refugia in Liolaemus pictus (Liolaemidae) during the Last Glacial Maximum in the Southern Andean Beech Forests

    PubMed Central

    Vera-Escalona, Iván; D'Elía, Guillermo; Gouin, Nicolás; Fontanella, Frank M.; Muñoz-Mendoza, Carla; Sites, Jack W.; Victoriano, Pedro F.

    2012-01-01

    Historical climate changes and orogenesis are two important factors that have shaped intraspecific biodiversity patterns worldwide. Although southern South America has experienced such complex events, there is a paucity of studies examining the effects on intraspecific diversification in this part of the world. Liolaemus pictus is the southernmost distributed lizard in the Chilean temperate forest, whose genetic structure has likely been influenced by Pleistocene glaciations. We conducted a phylogeographic study of L. pictus in Chile and Argentina based on one mitochondrial and two nuclear genes recovering two strongly divergent groups, Northern and Southern clades. The first group is distributed from the northernmost limit of the species to the Araucanía region while the second group is distributed throughout the Andes and the Chiloé archipelago in Southern Chile. Our results suggest that L. pictus originated 751 Kya, with divergence between the two clades occurring in the late Pleistocene. Demographic reconstructions for the Northern and Southern clades indicate a decrease in effective population sizes likely associated with Pleistocene glaciations. Surprisingly, patterns of genetic variation, clades age and historical gene flow in populations distributed within the limits of the Last Glacial Maximum (LGM) are not explained by recent colonization. We propose an “intra-Andean multiple refuge” hypothesis, along with the classical refuge hypothesis previously proposed for the biota of the Chilean Coastal range and Eastern Andean Cordillera. Our hypothesis is supported by niche modelling analysis suggesting the persistence of fragments of suitable habitat for the species within the limits of the LGM ice shield. This type of refuge hypothesis is proposed for the first time for an ectothermic species. PMID:23209552

  17. A Brief History of the Potato in Ireland.

    ERIC Educational Resources Information Center

    Social Education, 2000

    2000-01-01

    Provides historical information on the potato in Ireland focusing on how the potato arrived in Ireland and the advantages and disadvantages of the potato as a food crop. Discusses the Irish potato famine in Ireland, effects of the famine, and the government's laissez-faire response. Includes a list of questions. (CMK)

  18. Human responses and non-responses to climatic variations during the last Glacial-Interglacial transition in the eastern Mediterranean

    NASA Astrophysics Data System (ADS)

    Roberts, Neil; Woodbridge, Jessie; Bevan, Andrew; Palmisano, Alessio; Shennan, Stephen; Asouti, Eleni

    2018-03-01

    We review and evaluate human adaptations during the last glacial-interglacial climatic transition in southwest Asia. Stable isotope data imply that climatic change was synchronous across the region within the limits of dating uncertainty. Changes in vegetation, as indicated from pollen and charcoal, mirror step-wise shifts between cold-dry and warm-wet climatic conditions, but with lag effects for woody vegetation in some upland and interior areas. Palaeoenvironmental data can be set against regional archaeological evidence for human occupancy and economy from the later Epipalaeolithic to the aceramic Neolithic. Demographic change is evaluated from summed radiocarbon date probability distributions, which indicating contrasting - and in some cases opposite - population trajectories in different regions. Abrupt warming transitions at ∼14.5 and 11.7 ka BP may have acted as pacemakers for rapid cultural change in some areas, notably at the start of the Natufian and Pre-Pottery Neolithic cultures. However temporal synchroneity does not mean that climatic changes had the same environmental or societal consequences in different regions. During cold-dry time intervals, regions such as the Levant acted as refugia for plant and animal resources and human population. In areas where socio-ecological continuity was maintained through periods of adverse climate (e.g. Younger Dryas) human communities were able to respond rapidly to subsequent climatic improvement. By contrast, in areas where there was a break in settlement at these times (e.g. central Anatolia), populations were slower to react to the new opportunities provided by the interglacial world.

  19. Glacial ocean circulation and stratification explained by reduced atmospheric temperature

    NASA Astrophysics Data System (ADS)

    Jansen, Malte F.

    2017-01-01

    Earth’s climate has undergone dramatic shifts between glacial and interglacial time periods, with high-latitude temperature changes on the order of 5-10 °C. These climatic shifts have been associated with major rearrangements in the deep ocean circulation and stratification, which have likely played an important role in the observed atmospheric carbon dioxide swings by affecting the partitioning of carbon between the atmosphere and the ocean. The mechanisms by which the deep ocean circulation changed, however, are still unclear and represent a major challenge to our understanding of glacial climates. This study shows that various inferred changes in the deep ocean circulation and stratification between glacial and interglacial climates can be interpreted as a direct consequence of atmospheric temperature differences. Colder atmospheric temperatures lead to increased sea ice cover and formation rate around Antarctica. The associated enhanced brine rejection leads to a strongly increased deep ocean stratification, consistent with high abyssal salinities inferred for the last glacial maximum. The increased stratification goes together with a weakening and shoaling of the interhemispheric overturning circulation, again consistent with proxy evidence for the last glacial. The shallower interhemispheric overturning circulation makes room for slowly moving water of Antarctic origin, which explains the observed middepth radiocarbon age maximum and may play an important role in ocean carbon storage.

  20. Glacial ocean circulation and stratification explained by reduced atmospheric temperature

    PubMed Central

    Jansen, Malte F.

    2017-01-01

    Earth’s climate has undergone dramatic shifts between glacial and interglacial time periods, with high-latitude temperature changes on the order of 5–10 °C. These climatic shifts have been associated with major rearrangements in the deep ocean circulation and stratification, which have likely played an important role in the observed atmospheric carbon dioxide swings by affecting the partitioning of carbon between the atmosphere and the ocean. The mechanisms by which the deep ocean circulation changed, however, are still unclear and represent a major challenge to our understanding of glacial climates. This study shows that various inferred changes in the deep ocean circulation and stratification between glacial and interglacial climates can be interpreted as a direct consequence of atmospheric temperature differences. Colder atmospheric temperatures lead to increased sea ice cover and formation rate around Antarctica. The associated enhanced brine rejection leads to a strongly increased deep ocean stratification, consistent with high abyssal salinities inferred for the last glacial maximum. The increased stratification goes together with a weakening and shoaling of the interhemispheric overturning circulation, again consistent with proxy evidence for the last glacial. The shallower interhemispheric overturning circulation makes room for slowly moving water of Antarctic origin, which explains the observed middepth radiocarbon age maximum and may play an important role in ocean carbon storage. PMID:27994158

  1. Late-Glacial to Late-holocene Shifts in Global Precipitation Delta(sup 18)O

    NASA Technical Reports Server (NTRS)

    Jasechko, S.; Lechler, A.; Pausata, F.S.R.; Fawcett, P.J.; Gleeson, T.; Cendon, D.I.; Galewsky, J.; LeGrande, A. N.; Risi, C.; Sharp, Z. D.; hide

    2015-01-01

    Reconstructions of Quaternary climate are often based on the isotopic content of paleo-precipitation preserved in proxy records. While many paleo-precipitation isotope records are available, few studies have synthesized these dispersed records to explore spatial patterns of late-glacial precipitation delta(sup 18)O. Here we present a synthesis of 86 globally distributed groundwater (n 59), cave calcite (n 15) and ice core (n 12) isotope records spanning the late-glacial (defined as 50,000 to 20,000 years ago) to the late-Holocene (within the past 5000 years). We show that precipitation delta(sup 18)O changes from the late-glacial to the late-Holocene range from -7.1% (delta(sup 18)O(late-Holocene) > delta(sup 18)O(late-glacial) to +1.7% (delta(sup 18)O(late-glacial) > delta(sup 18)O(late-Holocene), with the majority (77) of records having lower late-glacial delta(sup 18)O than late-Holocene delta(sup 18)O values. High-magnitude, negative precipitation delta(sup 18)O shifts are common at high latitudes, high altitudes and continental interiors.

  2. Earth's glacial record and its tectonic setting

    NASA Astrophysics Data System (ADS)

    Eyles, N.

    1993-09-01

    Glaciations have occurred episodically at different time intervals and for different durations in Earth's history. Ice covers have formed in a wide range of plate tectonic and structural settings but the bulk of Earth's glacial record can be shown to have been deposited and preserved in basins within extensional settings. In such basins, source area uplift and basin subsidence fulfill the tectonic preconditions for the initiation of glaciation and the accomodation and preservation of glaciclastic sediments. Tectonic setting, in particular subsidence rates, also dictates the type of glaciclastic facies and facies successions that are deposited. Many pre-Pleistocene glaciated basins commonly contain well-defined tectonostratigraphic successions recording the interplay of tectonics and sedimentation; traditional climatostratigraphic approaches involving interpretation in terms of either ice advance/retreat cycles or glacio-eustatic sea-level change require revision. The direct record of continental glaciation in Earth history, in the form of classically-recognised continental glacial landforms and "tillites", is meagre; it is probable that more than 95% of the volume of preserved "glacial" strata are glacially-influenced marine deposits that record delivery of large amounts of glaciclastic sediment to offshore basins. This flux has been partially or completely reworked by "normal" sedimentary processes such that the record of glaciation and climate change is recorded in marine successions and is difficult to decipher. The dominant "glacial" facies in the rock record are subaqueous debris flow diamictites and turbidites recording the selective preservation of poorly-sorted glaciclastic sediment deposited in deep water basins by sediment gravity flows. However, these facies are also typical of many non-glacial settings, especially volcanically-influenced environments; numerous Archean and Proterozoic diamictites, described in the older literature as tillites, have no

  3. Quaternary Glacial Mapping in Western Wisconsin Using Soil Survey Information

    ERIC Educational Resources Information Center

    Oehlke, Betsy M.; Dolliver, Holly A. S.

    2011-01-01

    The majority of soils in the western Wisconsin have developed from glacial sediments deposited during the Quaternary Period (2.6 million years before present). In many regions, multiple advances and retreats have left a complex landscape of diverse glacial sediments and landforms. The soils that have developed on these deposits reflect the nature…

  4. Glacial lakes in Austria - Distribution and formation since the Little Ice Age

    NASA Astrophysics Data System (ADS)

    Buckel, J.; Otto, J. C.; Prasicek, G.; Keuschnig, M.

    2018-05-01

    Glacial lakes constitute a substantial part of the legacy of vanishing mountain glaciation and act as water storage, sediment traps and sources of both natural hazards and leisure activities. For these reasons, they receive growing attention by scientists and society. However, while the evolution of glacial lakes has been studied intensively over timescales tied to remote sensing-based approaches, the longer-term perspective has been omitted due a lack of suitable data sources. We mapped and analyzed the spatial distribution of glacial lakes in the Austrian Alps. We trace the development of number and area of glacial lakes in the Austrian Alps since the Little Ice Age (LIA) based on a unique combination of a lake inventory and an extensive record of glacier retreat. We find that bedrock-dammed lakes are the dominant lake type in the inventory. Bedrock- and moraine-dammed lakes populate the highest landscape domains located in cirques and hanging valleys. We observe lakes embedded in glacial deposits at lower locations on average below 2000 m a.s.l. In general, the distribution of glacial lakes over elevation reflects glacier erosional and depositional dynamics rather than the distribution of total area. The rate of formation of new glacial lakes (number, area) has continuously accelerated over time with present rates showing an eight-fold increase since LIA. At the same time the total glacier area decreased by two-thirds. This development coincides with a long-term trend of rising temperatures and a significant stepping up of this trend within the last 20 years in the Austrian Alps.

  5. Peatlands through the Last Glacial Cycle: Evidence and Model Results

    NASA Astrophysics Data System (ADS)

    Kleinen, T.; Treat, C. C.; Brovkin, V.

    2017-12-01

    The spatiotemporal distribution of peatlands prior to the last glacial maxium (LGM) is largely unknown. However, some evidence of non-extant peatlands is available in the form of buried organic-rich sediments. We have undertaken a synthesis of these "buried" peatlands from > 1000 detailed stratigraphic descriptions and combined it with data on extant peatlands to derive a first global synthesis of global peatland extent through the last glacial cycle. We present results of this synthesis in combination with modeling results where we determined peatland extents and carbon stocks from a transient simulation of the last glacial cycle with the CLIMBER2-LPJ model. We show that peat has existed in boreal latitudes at all times since the last interglacial, that evidence for tropical peatlands exists for the last 50,000 yrs, and that the model results in general agree well with the collected evidence of past peatlands, allowing a first estimate of peat carbon stock changes through the last glacial cycle. We discuss data and model limitations, with a focus on requirements for improving model-based peatland estimates.

  6. A conceptual model for glacial cycles and the middle Pleistocene transition

    NASA Astrophysics Data System (ADS)

    Daruka, István; Ditlevsen, Peter D.

    2016-01-01

    Milankovitch's astronomical theory of glacial cycles, attributing ice age climate oscillations to orbital changes in Northern-Hemisphere insolation, is challenged by the paleoclimatic record. The climatic response to the variations in insolation is far from trivial. In general the glacial cycles are highly asymmetric in time, with slow cooling from the interglacials to the glacials (inceptions) and very rapid warming from the glacials to the interglacials (terminations). We shall refer to this fast-slow dynamics as the "saw-tooth" shape of the paleoclimatic record. This is non-linearly related to the time-symmetric variations in the orbital forcing. However, the most pronounced challenge to the Milankovitch theory is the middle Pleistocene transition (MPT) occurring about one million years ago. During that event, the prevailing 41 kyr glacial cycles, corresponding to the almost harmonic obliquity cycle were replaced by longer saw-tooth shaped cycles with a time-scale around 100 kyr. The MPT must have been driven by internal changes in climate response, since it does not correspond to any apparent changes in the orbital forcing. In order to identify possible mechanisms causing the observed changes in glacial dynamics, it is relevant to study simplified models with the capability of generating temporal behavior similar to the observed records. We present a simple oscillator type model approach, with two variables, a temperature anomaly and a climatic memory term. The generalization of the ice albedo feedback is included in terms of an effective multiplicative coupling between this latter climatic memory term (representing the internal degrees of freedom) and the external drive. The simple model reproduces the temporal asymmetry of the late Pleistocene glacial cycles and suggests that the MPT can be explained as a regime shift, aided by climatic noise, from a period 1 frequency locking to the obliquity cycle to a period 2-3 frequency locking to the same obliquity

  7. Sink detection on tilted terrain for automated identification of glacial cirques

    NASA Astrophysics Data System (ADS)

    Prasicek, Günther; Robl, Jörg; Lang, Andreas

    2016-04-01

    Glacial cirques are morphologically distinct but complex landforms and represent a vital part of high mountain topography. Their distribution, elevation and relief are expected to hold information on (1) the extent of glacial occupation, (2) the mechanism of glacial cirque erosion, and (3) how glacial in concert with periglacial processes can limit peak altitude and mountain range height. While easily detectably for the expert's eye both in nature and on various representations of topography, their complicated nature makes them a nemesis for computer algorithms. Consequently, manual mapping of glacial cirques is commonplace in many mountain landscapes worldwide, but consistent datasets of cirque distribution and objectively mapped cirques and their morphometrical attributes are lacking. Among the biggest problems for algorithm development are the complexity in shape and the great variability of cirque size. For example, glacial cirques can be rather circular or longitudinal in extent, exist as individual and composite landforms, show prominent topographic depressions or can entirely be filled with water or sediment. For these reasons, attributes like circularity, size, drainage area and topology of landform elements (e.g. a flat floor surrounded by steep walls) have only a limited potential for automated cirque detection. Here we present a novel, geomorphometric method for automated identification of glacial cirques on digital elevation models that exploits their genetic bowl-like shape. First, we differentiate between glacial and fluvial terrain employing an algorithm based on a moving window approach and multi-scale curvature, which is also capable of fitting the analysis window to valley width. We then fit a plane to the valley stretch clipped by the analysis window and rotate the terrain around the center cell until the plane is level. Doing so, we produce sinks of considerable size if the clipped terrain represents a cirque, while no or only very small sinks

  8. Potentially bioavailable ferrous iron nanoparticles in glacial sediments

    NASA Astrophysics Data System (ADS)

    Hawkings, J.; Benning, L. G.; Raiswell, R.; Kaulich, B.; Araki, T.; Abyaneh, M.; Koch-Müller, M.; Stockdale, A.; Tranter, M.; Wadham, J.

    2017-12-01

    Iron (Fe) is an essential nutrient for marine phytoplankton, the primary producers of the ocean. Despite it being the fourth most abundant element in the Earth's crust, it is highly insoluble, due in part to its rapid oxidation from ferric (Fe2+) to ferrous phases (Fe3+), which often leads to the formation of nanoparticulate iron oxyhydroxide phases1. The insoluble nature of Fe in oxygenated waters means Fe limitation of primary producers is prevalent in 30-50% of the world's oceans, including areas of high biological productivity proximal to significant glacial activity (e.g., the Southern Ocean). Glaciers and ice sheets are a significant source of nanoparticulate Fe, which may be important in sustaining the high productivity observed in the near coastal regions proximal to glacial coverage. The reactivity of particulate iron is poorly understood, despite its importance in the ocean Fe inventory. Here we combined geochemical extractions, high-resolution imaging and spectroscopy to investigate the abundance, morphology and valence state of reactive iron in glacial sediments. Our results document the widespread occurrence of amorphous and Fe(II)-rich nanoparticles in glacial meltwaters and icebergs. Fe(II) is thought to be highly bioavailable in marine environments. We argue that glaciers and ice sheets are therefore able to supply potentially bioavailable Fe(II)-containing nanoparticulate material for downstream ecosystems, including those in a marine setting. The flux of bioavailable particulate iron from Arctic glaciers may increase as rising air temperatures lead to higher meltwater export.

  9. Glacial isostatic stress shadowing by the Antarctic ice sheet

    NASA Technical Reports Server (NTRS)

    Ivins, E. R.; James, T. S.; Klemann, V.

    2005-01-01

    Numerous examples of fault slip that offset late Quaternary glacial deposits and bedrock polish support the idea that the glacial loading cycle causes earthquakes in the upper crust. A semianalytical scheme is presented for quantifying glacial and postglacial lithospheric fault reactivation using contemporary rock fracture prediction methods. It extends previous studies by considering differential Mogi-von Mises stresses, in addition to those resulting from a Coulomb analysis. The approach utilizes gravitational viscoelastodynamic theory and explores the relationships between ice mass history and regional seismicity and faulting in a segment of East Antarctica containing the great Antarctic Plate (Balleny Island) earthquake of 25 March 1998 (Mw 8.1). Predictions of the failure stress fields within the seismogenic crust are generated for differing assumptions about background stress orientation, mantle viscosity, lithospheric thickness, and possible late Holocene deglaciation for the D91 Antarctic ice sheet history. Similar stress fracture fields are predicted by Mogi-von Mises and Coulomb theory, thus validating previous rebound Coulomb analysis. A thick lithosphere, of the order of 150-240 km, augments stress shadowing by a late melting (middle-late Holocene) coastal East Antarctic ice complex and could cause present-day earthquakes many hundreds of kilometers seaward of the former Last Glacial Maximum grounding line.

  10. Cyber-Bullying: The Situation in Ireland

    ERIC Educational Resources Information Center

    O'Moore, Mona

    2012-01-01

    This paper reports on the first major survey of cyber-bullying undertaken in Ireland. While preliminary results have been published they were based on a smaller and incomplete sample of 12-16 year olds living in Ireland. The preliminary results addressed the incidence level of cyber-bullying and that of the different subcategories of…

  11. Central Equatorial Pacific Sea Surface Temperatures During the Last Glacial Maximum

    NASA Astrophysics Data System (ADS)

    Monteagudo, M. M.; Lynch-Stieglitz, J.; Schmidt, M. W.

    2017-12-01

    The state of the tropical Pacific ocean-atmosphere system during the Last Glacial Maximum (LGM, 19,000-23,000 years BP) remains an area of uncertainty. Spatial patterns of tropical Pacific sea surface temperature (SST) offer insight into atmospheric circulation (i.e. Walker Circulation), however, few records exist for the Central Tropical Pacific (CTP). The few existing glacial CTP SST reconstructions indicate 1-2 °C of warming based on foraminiferal transfer functions (CLIMAP Project Members, 1976). In contrast, evidence from geochemical proxies (Mg/Ca, UK'37, TEX86) show 1-3.5 °C cooling in the eastern and western tropical Pacific (e.g. MARGO Project Members, 2009). In this study we present the first Mg/Ca estimates of glacial CTP SST from a meridional sediment core transect along the Line Islands Ridge (0-7°N, 156-162 °W). We use a time slice approach to establish the magnitude of glacial-interglacial SST change between the LGM (19,000-23,0000 years BP) and the Holocene (0-10,000 years BP) using Mg/Ca in the surface-dwelling foraminifera Globigerinoides ruber. Our results indicate cooling at all latitudes, ranging between 1.2-2.7 °C (Holocene-LGM SST). Northern cores (6.83-2.77 °N) exhibit a smaller glacial-interglacial SST difference than equatorial site 20BB at 1.27 °N. The data generated thus far suggest the glacial meridional SST gradient may have been steeper, possibly as a result of increased zonal winds, equatorial upwelling, or westward expansion of the Eastern Pacific Cold Tongue.

  12. Glacial Hazards in Chile: Processes, Assessment, Mitigation and Risk Management Strategies

    NASA Astrophysics Data System (ADS)

    Glasser, N. F.; Wilson, R.; Casassa, G., Sr.; Reynolds, J.; Harrison, S.; Shannon, S. R.; Schaefer, M.; Iribarran, P.

    2017-12-01

    Glacial Lake Outburst Floods (GLOFs) are capable of travelling considerable distances from their source and they represent one of the most important glacial hazards. In line with observations in other parts of the world, the frequency of GLOF events in Chile has increased in recent decades highlighting the need to quantify the flood risk posed to downstream areas. This poster presents the work of the `Glacial Hazards in Chile' project which aims to (1) better understand the processes that govern the development of GLOFs in Chile, (2) estimate the socio-economic effects of GLOFs in Chile, and (3) provide a GLOF risk assessment framework that can be applied to Chile and other lower income countries globally. As an initial step towards the completion of these aims, we have recently compiled the first glacial lake inventory for the central and Patagonian Andes, which details the temporal development of glacial lakes in this region over the past three decades. This analysis was used to identify two lakes of interest that were visited during a fieldwork expedition in February 2017. The first of these, Lago Chileno in Patagonia, has recently produced a large GLOF causing significant damage to the downstream floodplain, whilst the second was identified as one of the fastest growing lakes in the central Andes. Both these lakes were surveyed using aerial imagery acquired with a drone and a custom-built bathymetry boat, data from which will help to improve our understanding of the physical processes associated with glacial lake development and failure within the Chilean Andes.

  13. Plant molecular phylogeography in China and adjacent regions: Tracing the genetic imprints of Quaternary climate and environmental change in the world's most diverse temperate flora.

    PubMed

    Qiu, Ying-Xiong; Fu, Cheng-Xing; Comes, Hans Peter

    2011-04-01

    The Sino-Japanese Floristic Region (SJFR) of East Asia harbors the most diverse of the world's temperate flora, and was the most important glacial refuge for its Tertiary representatives ('relics') throughout Quaternary ice-age cycles. A steadily increasing number of phylogeographic studies in the SJFR of mainland China and adjacent areas, including the Qinghai-Tibetan-Plateau (QTP) and Sino-Himalayan region, have documented the population histories of temperate plant species in these regions. Here we review this current literature that challenges the oft-stated view of the SJFR as a glacial sanctuary for temperate plants, instead revealing profound effects of Quaternary changes in climate, topography, and/or sea level on the current genetic structure of such organisms. There are three recurrent phylogeographic scenarios identified by different case studies that broadly agree with longstanding biogeographic or palaeo-ecological hypotheses: (i) postglacial re-colonization of the QTP from (south-)eastern glacial refugia; (ii) population isolation and endemic species formation in Southwest China due to tectonic shifts and river course dynamics; and (iii) long-term isolation and species survival in multiple localized refugia of (warm-)temperate deciduous forest habitats in subtropical (Central/East/South) China. However, in four additional instances, phylogeographic findings seem to conflict with a priori predictions raised by palaeo-data, suggesting instead: (iv) glacial in situ survival of some hardy alpine herbs and forest trees on the QTP platform itself; (v) long-term refugial isolation of (warm-)temperate evergreen taxa in subtropical China; (vi) 'cryptic' glacial survival of (cool-)temperate deciduous forest trees in North China; and (vii) unexpectedly deep (Late Tertiary/early-to-mid Pleistocene) allopatric-vicariant differentiation of disjunct lineages in the East China-Japan-Korea region due to past sea transgressions. We discuss these and other consequences

  14. Postglacial migration and adaptation for dispersal in pitch pine (Pinaceae).

    PubMed

    Ledig, F Thomas; Smouse, Peter E; Hom, John L

    2015-12-01

    Variation in a species is a blend of adaptive, random, and migratory responses. Pitch pine (Pinus rigida), a highly variable eastern conifer, has occupied multiple glacial refugia, whose harsh conditions favored adaptations enhancing subsequent dispersal and recolonization of newly deglaciated sites. We assessed phenotypic diversity in long-term growth trials to elucidate both the adaptations and likely refugia. Pitch pine progeny from 31 areas were grown in common gardens in six locations, from eastern Massachusetts to Korea. Survival increased with source latitude, but seedlings from southern latitudes were tallest in the first (postplanting) year, but that advantage dissipated in later years. Progeny from northern latitudes were precocious, highly fecund, had smaller seeds, and more seeds per cone. Seed mass decreased with latitude in both parents and progeny. Serotinous cones were notably common in the New Jersey Pine Plains and Acadia National Park. Various disease agents and frost burn exhibited latitudinal trends that were nonlinear, with a break in the regression slope at about 40°N latitude. Cluster analysis identified both northern and southern groups, largely split between unglaciated and deglaciated terrain, but with Acadia and the Pine Plains as unique outliers. Within the southern group, provenances were organized into contiguous subgroups, but geographic structure was less evident in the northern group. The present range of pitch pine was colonized by migrants from at least three different refugia, including at least one on the exposed continental shelf during the Last Glacial Maximum. This article is a U.S. Government work and is in the public domain in the USA. © Botanical Society of America (outside the USA) 2015.

  15. Intervarietal and intravarietal genetic structure in Douglas-fir: nuclear SSRs bring novel insights into past population demographic processes, phylogeography, and intervarietal hybridization

    PubMed Central

    van Loo, Marcela; Hintsteiner, Wolfgang; Pötzelsberger, Elisabeth; Schüler, Silvio; Hasenauer, Hubert

    2015-01-01

    Douglas-fir (Pseudotsuga menziesii) is one of numerous wide-range forest tree species represented by subspecies/varieties, which hybridize in contact zones. This study examined the genetic structure of this North American conifer and its two hybridizing varieties, coastal and Rocky Mountain, at intervarietal and intravarietal level. The genetic structure was subsequently associated with the Pleistocene refugial history, postglacial migration and intervarietal hybridization/introgression. Thirty-eight populations from the USA and Canada were genotyped for 13 nuclear SSRs and analyzed with simulations and traditional population genetic structuring methods. Eight genetic clusters were identified. The coastal clusters embodied five refugial populations originating from five distinct refugia. Four coastal refugial populations, three from California and one from western Canada, diverged during the Pleistocene (56.9–40.1 ka). The three Rocky Mountain clusters reflected distinct refugial populations of three glacial refugia. For Canada, ice covered during the Last Glacial Maximum, we present the following three findings. (1) One refugial population of each variety was revealed in the north of the distribution range. Additional research including paleodata is required to support and determine whether both northern populations originated from cryptic refugia situated south or north of the ice-covered area. (2) An interplay between intravarietal gene flow of different refugial populations and intervarietal gene flow by hybridization and introgression was identified. (3) The Canadian hybrid zone displayed predominantly introgressants of the Rocky Mountain into the coastal variety. This study provides new insights into the complex Quaternary dynamics of this conifer essential for understanding its evolution (outside and inside the native range), adaptation to future climates and for forest management. PMID:26140197

  16. Endemism hotspots are linked to stable climatic refugia

    PubMed Central

    Noss, Reed

    2017-01-01

    Background Centres of endemism have received much attention from evolutionists, biogeographers, ecologists and conservationists. Climatic stability is often cited as a major reason for the occurrences of these geographic concentrations of species which are not found anywhere else. The proposed linkage between endemism and climatic stability raises unanswered questions about the persistence of biodiversity during the present era of rapidly changing climate. Key Questions The current status of evidence linking geographic centres of endemism to climatic stability over evolutionary time was examined. The following questions were asked. Do macroecological analyses support such an endemism–stability linkage? Do comparative studies find that endemic species display traits reflecting evolution in stable climates? Will centres of endemism in microrefugia or macrorefugia remain relatively stable and capable of supporting high biological diversity into the future? What are the implications of the endemism–stability linkage for conservation? Conclusions Recent work using the concept of climate change velocity supports the classic idea that centres of endemism occur where past climatic fluctuations have been mild and where mountainous topography or favourable ocean currents contribute to creating refugia. Our knowledge of trait differences between narrow endemics and more widely distributed species remains highly incomplete. Current knowledge suggests that centres of endemism will remain relatively climatically buffered in the future, with the important caveat that absolute levels of climatic change and species losses in these regions may still be large. PMID:28064195

  17. Are glacials "dry" - and in what sense?

    NASA Astrophysics Data System (ADS)

    Scheff, J.; Seager, R.; Coats, S.; Liu, H.

    2016-12-01

    Glacial maxima during the Pleistocene are generally thought to be arid on land, with a few regional exceptions. Recent work on future climate change, however, has found that different wetness-related variables have opposite-signed responses over large portions of the continents, belying simple ideas of local "drying" or "wetting" with global temperature change in models. Here, we show that this behavior extends to simulations of the Last Glacial Maximum as well: the continents are modeled to have generally wetter topsoils and higher values of standard climate-wetness metrics in the LGM than in the preindustrial, as well as generally lower precipitation and ubiquitously lower photosynthesis (likely driven by the low CO2), with the streamflow response falling in between. Is this model-derived view of the LGM an accurate one? Using a large community pollen and plant-fossil compilation, we confirm that LGM grasslands and open woodlands grew at many sites of present potential forest, seasonal or dry forests at many sites of present potential rain- or seasonal forests, and so forth, while changes in the opposite sense were extremely few and spatially confined. We show that this strongly resembles the simulated photosynthesis changes, but not the simulated streamflow or soil moisture changes. Meanwhile, published LGM lake-level estimates resemble the simulated streamflow changes, but not the photosynthesis changes. Thus, the last glacial does not appear to be systematically "dry" outside the high latitudes, but merely carbon-starved. Similarly, local findings of reduced or more open vegetation at the LGM (e.g. from pollen, carbon isotopes, or dustiness) do not indicate local "aridity" unless corroborating hydrological proxies are also found. Finally, this work suggests that glacial-era evidence of open vegetation with high lake levels (as in the eastern Mediterranean) is not odd or paradoxical, but entirely consistent with climate model output.

  18. The Holocene British and Irish ancient forest fossil beetle fauna: implications for forest history, biodiversity and faunal colonisation

    NASA Astrophysics Data System (ADS)

    Whitehouse, Nicki J.

    2006-08-01

    its hosts and the role played by European glacial refugia.

  19. Speciation and genetic diversity in Centaurea subsect. Phalolepis in Anatolia

    PubMed Central

    López-Pujol, Jordi; López-Vinyallonga, Sara; Susanna, Alfonso; Ertuğrul, Kuddisi; Uysal, Tuna; Tugay, Osman; Guetat, Arbi; Garcia-Jacas, Núria

    2016-01-01

    Mountains of Anatolia are one of the main Mediterranean biodiversity hotspots and their richness in endemic species amounts for 30% of the flora. Two main factors may account for this high diversity: the complex orography and its role as refugia during past glaciations. We have investigated seven narrow endemics of Centaurea subsection Phalolepis from Anatolia by means of microsatellites and ecological niche modelling (ENM), in order to analyse genetic polymorphisms and getting insights into their speciation. Despite being narrow endemics, all the studied species show moderate to high SSR genetic diversity. Populations are genetically isolated, but exchange of genes probably occurred at glacial maxima (likely through the Anatolian mountain arches as suggested by the ENM). The lack of correlation between genetic clusters and (morpho) species is interpreted as a result of allopatric diversification on the basis of a shared gene pool. As suggested in a former study in Greece, post-glacial isolation in mountains would be the main driver of diversification in these plants; mountains of Anatolia would have acted as plant refugia, allowing the maintenance of high genetic diversity. Ancient gene flow between taxa that became sympatric during glaciations may also have contributed to the high levels of genetic diversity. PMID:27886271

  20. The early rise and late demise of New Zealand’s last glacial maximum

    PubMed Central

    Rother, Henrik; Fink, David; Shulmeister, James; Mifsud, Charles; Evans, Michael; Pugh, Jeremy

    2014-01-01

    Recent debate on records of southern midlatitude glaciation has focused on reconstructing glacier dynamics during the last glacial termination, with different results supporting both in-phase and out-of-phase correlations with Northern Hemisphere glacial signals. A continuing major weakness in this debate is the lack of robust data, particularly from the early and maximum phase of southern midlatitude glaciation (∼30–20 ka), to verify the competing models. Here we present a suite of 58 cosmogenic exposure ages from 17 last-glacial ice limits in the Rangitata Valley of New Zealand, capturing an extensive record of glacial oscillations between 28–16 ka. The sequence shows that the local last glacial maximum in this region occurred shortly before 28 ka, followed by several successively less extensive ice readvances between 26–19 ka. The onset of Termination 1 and the ensuing glacial retreat is preserved in exceptional detail through numerous recessional moraines, indicating that ice retreat between 19–16 ka was very gradual. Extensive valley glaciers survived in the Rangitata catchment until at least 15.8 ka. These findings preclude the previously inferred rapid climate-driven ice retreat in the Southern Alps after the onset of Termination 1. Our record documents an early last glacial maximum, an overall trend of diminishing ice volume in New Zealand between 28–20 ka, and gradual deglaciation until at least 15 ka. PMID:25071171

  1. Glaciation Effects on the Phylogeographic Structure of Oligoryzomys longicaudatus (Rodentia: Sigmodontinae) in the Southern Andes

    PubMed Central

    Palma, R. Eduardo; Boric-Bargetto, Dusan; Torres-Pérez, Fernando; Hernández, Cristián E.; Yates, Terry L.

    2012-01-01

    The long-tailed pygmy rice rat Oligoryzomys longicaudatus (Sigmodontinae), the major reservoir of Hantavirus in Chile and Patagonian Argentina, is widely distributed in the Mediterranean, Temperate and Patagonian Forests of Chile, as well as in adjacent areas in southern Argentina. We used molecular data to evaluate the effects of the last glacial event on the phylogeographic structure of this species. We examined if historical Pleistocene events had affected genetic variation and spatial distribution of this species along its distributional range. We sampled 223 individuals representing 47 localities along the species range, and sequenced the hypervariable domain I of the mtDNA control region. Aligned sequences were analyzed using haplotype network, Bayesian population structure and demographic analyses. Analysis of population structure and the haplotype network inferred three genetic clusters along the distribution of O. longicaudatus that mostly agreed with the three major ecogeographic regions in Chile: Mediterranean, Temperate Forests and Patagonian Forests. Bayesian Skyline Plots showed constant population sizes through time in all three clusters followed by an increase after and during the Last Glacial Maximum (LGM; between 26,000–13,000 years ago). Neutrality tests and the “g” parameter also suggest that populations of O. longicaudatus experienced demographic expansion across the species entire range. Past climate shifts have influenced population structure and lineage variation of O. longicaudatus. This species remained in refugia areas during Pleistocene times in southern Temperate Forests (and adjacent areas in Patagonia). From these refugia, O. longicaudatus experienced demographic expansions into Patagonian Forests and central Mediterranean Chile using glacial retreats. PMID:22396751

  2. Have historical climate changes affected Gentoo penguin (Pygoscelis papua) populations in Antarctica?

    PubMed

    Peña M, Fabiola; Poulin, Elie; Dantas, Gisele P M; González-Acuña, Daniel; Petry, Maria Virginia; Vianna, Juliana A

    2014-01-01

    The West Antarctic Peninsula (WAP) has been suffering an increase in its atmospheric temperature during the last 50 years, mainly associated with global warming. This increment of temperature trend associated with changes in sea-ice dynamics has an impact on organisms, affecting their phenology, physiology and distribution range. For instance, rapid demographic changes in Pygoscelis penguins have been reported over the last 50 years in WAP, resulting in population expansion of sub-Antarctic Gentoo penguin (P. papua) and retreat of Antarctic Adelie penguin (P. adeliae). Current global warming has been mainly associated with human activities; however these climate trends are framed in a historical context of climate changes, particularly during the Pleistocene, characterized by an alternation between glacial and interglacial periods. During the last maximal glacial (LGM∼21,000 BP) the ice sheet cover reached its maximum extension on the West Antarctic Peninsula (WAP), causing local extinction of Antarctic taxa, migration to lower latitudes and/or survival in glacial refugia. We studied the HRVI of mtDNA and the nuclear intron βfibint7 of 150 individuals of the WAP to understand the demographic history and population structure of P. papua. We found high genetic diversity, reduced population genetic structure and a signature of population expansion estimated around 13,000 BP, much before the first paleocolony fossil records (∼1,100 BP). Our results suggest that the species may have survived in peri-Antarctic refugia such as South Georgia and North Sandwich islands and recolonized the Antarctic Peninsula and South Shetland Islands after the ice sheet retreat.

  3. Glaciation effects on the phylogeographic structure of Oligoryzomys longicaudatus (Rodentia: Sigmodontinae) in the southern Andes.

    PubMed

    Palma, R Eduardo; Boric-Bargetto, Dusan; Torres-Pérez, Fernando; Hernández, Cristián E; Yates, Terry L

    2012-01-01

    The long-tailed pygmy rice rat Oligoryzomys longicaudatus (Sigmodontinae), the major reservoir of Hantavirus in Chile and Patagonian Argentina, is widely distributed in the Mediterranean, Temperate and Patagonian Forests of Chile, as well as in adjacent areas in southern Argentina. We used molecular data to evaluate the effects of the last glacial event on the phylogeographic structure of this species. We examined if historical Pleistocene events had affected genetic variation and spatial distribution of this species along its distributional range. We sampled 223 individuals representing 47 localities along the species range, and sequenced the hypervariable domain I of the mtDNA control region. Aligned sequences were analyzed using haplotype network, bayesian population structure and demographic analyses. Analysis of population structure and the haplotype network inferred three genetic clusters along the distribution of O. longicaudatus that mostly agreed with the three major ecogeographic regions in Chile: Mediterranean, Temperate Forests and Patagonian Forests. Bayesian Skyline Plots showed constant population sizes through time in all three clusters followed by an increase after and during the Last Glacial Maximum (LGM; between 26,000-13,000 years ago). Neutrality tests and the "g" parameter also suggest that populations of O. longicaudatus experienced demographic expansion across the species entire range. Past climate shifts have influenced population structure and lineage variation of O. longicaudatus. This species remained in refugia areas during Pleistocene times in southern Temperate Forests (and adjacent areas in Patagonia). From these refugia, O. longicaudatus experienced demographic expansions into Patagonian Forests and central Mediterranean Chile using glacial retreats.

  4. Simulated Last Glacial Maximum Δ14CATM and the Deep Glacial Ocean Reservoir

    NASA Astrophysics Data System (ADS)

    Mariotti, V.; Paillard, D.; Roche, D. M.; Bouttes, N.; Bopp, L.

    2012-12-01

    Δ14Catm has been estimated at 420 ± 80‰ (INTCAL09) during the Last Glacial Maximum (LGM) compared to preindustrial times (0‰), but mechanisms explaining this difference are not yet resolved. Δ14Catm is a function of cosmogenic production in high atmosphere and of carbon cycling in the Earth system (through carbon exchange with the superficial reservoirs, ocean and continental biosphere). 10Be-based reconstructions show a contribution of the cosmogenic production term of only 200 ± 200‰ at the LGM. The remaining 220‰ of Δ14Catm variation between the LGM and preindustrial times have thus to be explained by changes in the carbon cycle. Recently, Bouttes et al. (2010) proposed to explain most of the difference in atmospheric pCO2 between glacial and interglacial times by brine-induced ocean stratification in the Southern Ocean. This mechanism involves the formation of very saline water masses that can store Dissolved Inorganic Carbon (DIC) in the deep ocean. During glacial times, the sinking of brines is enhanced and more DIC is stored in the deep ocean, lowering atmospheric pCO2. Such an isolated ocean reservoir would be characterized by a low Δ14C signature. Evidence of such 14C-depleted deep waters during the LGM has recently been found in the Southern Ocean (Skinner et al., 2010). The degassing of this carbon with low Δ14C would then reduce Δ14Catm throughout the deglaciation. We have further developed the CLIMBER-2 model to include a cosmogenic production of 14C as well as an interactive atmospheric 14C reservoir. We investigate the role of both sinking of brines and cosmogenic production, alongside iron and vertical diffusion mechanisms to explain changes in Δ14Catm during the last deglaciation. In our simulations, not only the sinking of brine mechanism is consistent with past Δ14C data but also it explains most of the differences in atmospheric pCO2 and Δ14C between LGM and preindustrial times.

  5. The Glacial BuzzSaw, Isostasy, and Global Crustal Models

    NASA Astrophysics Data System (ADS)

    Levander, A.; Oncken, O.; Niu, F.

    2015-12-01

    The glacial buzzsaw hypothesis predicts that maximum elevations in orogens at high latitudes are depressed relative to temperate latitudes, as maximum elevation and hypsography of glaciated orogens are functions of the glacial equilibrium line altitude (ELA) and the modern and last glacial maximum (LGM) snowlines. As a consequence crustal thickness, density, or both must change with increasing latitude to maintain isostatic balance. For Airy compensation crustal thickness should decrease toward polar latitudes, whereas for Pratt compensation crustal densities should increase. For similar convergence rates, higher latitude orogens should have higher grade, and presumably higher density rocks in the crustal column due to more efficient glacial erosion. We have examined a number of global and regional crustal models to see if these predictions appear in the models. Crustal thickness is straightforward to examine, crustal density less so. The different crustal models generally agree with one another, but do show some major differences. We used a standard tectonic classification scheme of the crust for data selection. The globally averaged orogens show crustal thicknesses that decrease toward high latitudes, almost reflecting topography, in both the individual crustal models and the models averaged together. The most convincing is the western hemisphere cordillera, where elevations and crustal thicknesses decrease toward the poles, and also toward lower latitudes (the equatorial minimum is at ~12oN). The elevation differences and Airy prediction of crustal thickness changes are in reasonable agreement in the North American Cordillera, but in South America the observed crustal thickness change is larger than the Airy prediction. The Alpine-Himalayan chain shows similar trends, however the strike of the chain makes interpretation ambiguous. We also examined cratons with ice sheets during the last glacial period to see if continental glaciation also thins the crust toward

  6. Long-term cost effectiveness of cardiac secondary prevention in primary care in the Republic of Ireland and Northern Ireland.

    PubMed

    Gillespie, Paddy; Murphy, Edel; Smith, Susan M; Cupples, Margaret E; Byrne, Molly; Murphy, Andrew W

    2017-04-01

    While cardiac secondary prevention in primary care is established practice, little is known about its long-term cost effectiveness. This study examines the cost effectiveness of a secondary prevention intervention in primary care in the Republic of Ireland and Northern Ireland over 6 years. An economic evaluation, based on a cluster randomised controlled trial of 903 patients with heart disease, was conducted 4.5 years after the intervention ceased to be delivered. Patients originally randomised to the control received usual practice while those randomised to the intervention received a tailored care package over the 1.5-year delivery period. Data on healthcare costs and quality adjusted life expectancy were used to undertake incremental cost utility analysis. Multilevel regression was used to estimate mean cost effectiveness and uncertainty was examined using cost effectiveness acceptability curves. At 6 years, there was a divergence in the results across jurisdictions. While the probability of the intervention being cost effective in the Republic of Ireland was 0.434, 0.232, 0.180, 0.150, 0.115 and 0.098 at selected threshold values of €5000, €15,000, €20,000, €25,000, €35,000 and €45,000, respectively, all equivalent probabilities for Northern Ireland equalled 1.000. Our findings suggest that the intervention in its current format is likely to be more cost effective than usual general practice care in Northern Ireland, but this is not the case in the Republic of Ireland.

  7. A global perspective on Glacial- to Interglacial variability change

    NASA Astrophysics Data System (ADS)

    Rehfeld, Kira; Münch, Thomas; Ho, Sze Ling; Laepple, Thomas

    2017-04-01

    Changes in climate variability are more important for society than changes in the mean state alone. While we will be facing a large-scale shift of the mean climate in the future, its implications for climate variability are not well constrained. Here we quantify changes in temperature variability as climate shifted from the Last Glacial cold to the Holocene warm period. Greenland ice core oxygen isotope records provide evidence of this climatic shift, and are used as reference datasets in many palaeoclimate studies worldwide. A striking feature in these records is pronounced millennial variability in the Glacial, and a distinct reduction in variance in the Holocene. We present quantitative estimates of the change in variability on 500- to 1500-year timescales based on a global compilation of high-resolution proxy records for temperature which span both the Glacial and the Holocene. The estimates are derived based on power spectral analysis, and corrected using estimates of the proxy signal-to-noise ratios. We show that, on a global scale, variability at the Glacial maximum is five times higher than during the Holocene, with a possible range of 3-10 times. The spatial pattern of the variability change is latitude-dependent. While the tropics show no changes in variability, mid-latitude changes are higher. A slight overall reduction in variability in the centennial to millennial range is found in Antarctica. The variability decrease in the Greenland ice core oxygen isotope records is larger than in any other proxy dataset. These results therefore contradict the view of a globally quiescent Holocene following the instable Glacial, and imply that, in terms of centennial to millennial temperature variability, the two states may be more similar than previously thought.

  8. [Primary care in Ireland].

    PubMed

    Sánchez-Sagrado, T

    Spanish doctors are still leaving the country to look for quality work. Ireland is not a country with many Spanish professionals but it is interesting to know its particular Health care system. Ireland is one of the countries with a national health care system, although it has a mixture of private health care insurance schemes. People have a right to health care if they have been living in Ireland at least for a year. Access to the primary care health system depends on age and income: free of charge for Category 1 and co-payments for the rest. This division generates great inequalities among the population. Primary Care doctors are self-employed, and they work independently. However, since 2001 they have tended to work in multidisciplinary teams in order to strengthen the Primary Care practice. Salary is gained from a combination of public and private incomes which are not differentiated. The role of the General Practitioner consists in the treatment of acute and chronic diseases, minor surgery, child care, etc. There is no coordination between Primary and Secondary care. Access to specialised medicine is regulated by the price of consultation. Primary Care doctors are not gatekeepers. To be able to work here, doctors must have three years of training after medical school. After that, Continuing Medical Education is compulsory, and the college of general practitioners monitors it annually. The Irish health care system does not fit into the European model. Lack of a clear separation between public and private health care generates great inequalities. The non-existence of coordination between primary and specialised care leads to inefficiencies, which Ireland cannot allow itself after a decade of economic crisis. Copyright © 2017 Sociedad Española de Médicos de Atención Primaria (SEMERGEN). Publicado por Elsevier España, S.L.U. All rights reserved.

  9. Development of a glacially dominated shelf-slope-fan system in tectonically active southeast Alaska: Results of IODP Expedition 341 core-log-seismic integrated studies at glacial cycle resolution

    NASA Astrophysics Data System (ADS)

    Gulick, Sean; Jaeger, John; Mix, Alan; Swartz, John; Worthington, Lindsay; Reece, Robert

    2014-05-01

    Collision of the Yakutat microplate with North American formed the St. Elias Mountains in coastal Gulf of Alaska. While the tectonic driver for orogenesis has been ongoing since the Miocene, results from the Integrated Ocean Drilling Program Expedition 341 suggests that direct climatic perturbation of active orogenesis through glacial erosion is non-linear. Geophysical studies of the glaciated continental margin, slope, and adjacent deep-sea Surveyor Fan allow examination of the glaciated orogen from source to sink. Using high-resolution and crustal-scale seismic data and through comparison with other glaciated margins, we can identify key diagnostic seismic morphologies and facies indicative of glacial proximity and sediment routing. Expedition drilling results calibrated these images suggesting a timeline for initial advances of the Cordilleran ice sheet related glacial systems onto the shelf and a further timeline for the development of ice streams that reach the shelf edge. Comparisons can be made within this single margin between evolution of the tectonic-glacial system where erosion and sediment transport are occurring within a fold and thrust belt versus on a more stable shelf region. Onshore the Bering-Bagley glacial system in the west flows across the Yakataga fold and thrust belt, allowing examination of whether glacial erosion can cause tectonic feedbacks, whereas offshore the Bering-Bagley system interacts with the Pamplona Zone thrusts in a region of significant sediment accommodation. Results from Expedition 341 imply that timing of glacial advance to the shelf edge in this region may be driven by the necessity of filling up the accommodation through aggradation followed by progradation and thus is autogenic. In contrast the Malaspina-Hubbard glacial system to the east encountered significantly less accommodation and more directly responded to climatic forcing including showing outer shelf glacial occupation since the mid-Pleistocene transition-MPT to

  10. Understanding water column and streambed thermal refugia for endangered mussels in the Delaware River

    USGS Publications Warehouse

    Briggs, Martin A.; Voytek, Emily B.; Day-Lewis, Frederick D.; Rosenberry, Donald O.; Lane, John W.

    2013-01-01

    Groundwater discharge locations along the upper Delaware River, both discrete bank seeps and diffuse streambed upwelling, may create thermal niche environments that benefit the endangered dwarf wedgemussel (Alasmidonta heterodon). We seek to identify whether discrete or diffuse groundwater inflow is the dominant control on refugia. Numerous springs and seeps were identified at all locations where dwarf wedgemussels still can be found. Infrared imagery and custom high spatial resolution fiber-optic distributed temperature sensors reveal complex thermal dynamics at one of the seeps with a relatively stable, cold groundwater plume extending along the streambed/water-column interface during mid-summer. This plume, primarily fed by a discrete bank seep, was shown through analytical and numerical heat-transport modeling to dominate temperature dynamics in the region of potential habitation by the adult dwarf wedgemussel.

  11. Rock avalanche deposits in Alai Valley, Central Asia: misinterpretation of glacial record

    NASA Astrophysics Data System (ADS)

    Reznichenko, Natalya; Davies, Tim; Robinson, Tom; De Pascale, Gregory

    2013-04-01

    The reconstruction of Quaternary glaciations has been restricted by conventional approaches with resulting contradictions in interpretation of the regional glacial record, that recently have been subjected to critical re-evaluation. Along with uncertainties in dating techniques and their applicability to particular landforms (Kirkbride and Winkler, 2012), it has recently been demonstrated that the presence of rock avalanche debris in a landform can be unequivocally detected; this allows for the first time definitive identification of and distinction between glacial moraines and landslide deposits. It also identifies moraines that have formed due to rock avalanche deposition on glaciers, possibly with no associated climatic signal (Reznichenko et al., 2012). Confusion between landslide deposits and moraines is evident for ranges in Central Asia (e.g., Hewitt, 1999) where the least-studied glacial record is selectively correlated with established glacial chronologies in Alpine ranges, which in turn masks the actual glacial extent and their responses to climate change, tectonics and landsliding activity. We describe examples in the glaciated Alai Valley, large intermountain depression between the Zaalay Range of the Northern Pamir and the Alay Range of the Southern Tien-Shan, showing that some large Quaternary deposits classically interpreted as moraines are of rock avalanche origin. Sediment from these deposits has been tested for the presence of agglomerates that are only produced under high stress conditions during rock avalanche motion, and are absent from glacial sediments (Reznichenko et al., 2012). This reveals that morphologically-similar deposits have radically different geneses: rock avalanche origin for a deposit in the Komansu river catchment and glacial origin for deposits in the Ashiktash and Kyzylart catchments. The enormous Komansu rock avalanche deposit, probably triggered by a rupture of the Main Pamir thrust, currently covers about 100 km2 with a

  12. A comparison of climate simulations for the last glacial maximum with three different versions of the ECHAM model and implications for summer-green tree refugia

    NASA Astrophysics Data System (ADS)

    Arpe, K.; Leroy, S. A. G.; Mikolajewicz, U.

    2011-02-01

    Model simulations of the last glacial maximum (21 ± 2 ka) with the ECHAM3 T42 atmosphere-only, ECHAM5-MPIOM T31 atmosphere-ocean coupled and ECHAM5 T106 atmosphere-only models are compared. The topography, land-sea mask and glacier distribution for the ECHAM5 simulations were taken from the Paleoclimate Modelling Intercomparison Project Phase II (PMIP2) data set while for ECHAM3 they were taken from PMIP1. The ECHAM5-MPIOM T31 model produced its own sea surface temperatures (SST) while the ECHAM5 T106 simulations were forced at the boundaries by this coupled model SSTs corrected from their present-day biases and the ECHAM3 T42 model was forced with prescribed SSTs provided by Climate/Long-Range Investigation, Mapping, and Prediction project (CLIMAP). The SSTs in the ECHAM5-MPIOM simulation for the last glacial maximum (LGM) were much warmer in the northern Atlantic than those suggested by CLIMAP or Overview of Glacial Atlantic Ocean Mapping (GLAMAP) while the SSTs were cooler everywhere else. This had a clear effect on the temperatures over Europe, warmer for winters in western Europe and cooler for eastern Europe than the simulation with CLIMAP SSTs. Considerable differences in the general circulation patterns were found in the different simulations. A ridge over western Europe for the present climate during winter in the 500 hPa height field remains in both ECHAM5 simulations for the LGM, more so in the T106 version, while the ECHAM3 CLIMAP-SST simulation provided a trough which is consistent with cooler temperatures over western Europe. The zonal wind between 30° W and 10° E shows a southward shift of the polar and subtropical jets in the simulations for the LGM, least obvious in the ECHAM5 T31 one, and an extremely strong polar jet for the ECHAM3 CLIMAP-SST run. The latter can probably be assigned to the much stronger north-south gradient in the CLIMAP SSTs. The southward shift of the polar jet during the LGM is supported by palaeo-data. Cyclone tracks in

  13. Asymmetric Signature of Glacial Antarctic Intermediate Water in the Central South Pacific

    NASA Astrophysics Data System (ADS)

    Tapia, R.; Nuernberg, D.; Ho, S. L.; Lamy, F.; Ullermann, J.; Gersonde, R.; Tiedemann, R.

    2017-12-01

    Southern Ocean Intermediate Waters (SOIWs) play a key role in modulating the global climate on glacial-interglacial time scales as they connect the Southern Ocean and the tropics. Despite their importance, the past evolution of the SOIWs in the central South Pacific is largely unknown due to a dearth of sedimentary archives. Here we compare Mg/Ca-temperature, stable carbon and oxygen isotope records from surface-dwelling (G. bulloides) and deep-dwelling (G. inflata) planktic foraminifera at site PS75/059-2 (54°12.9' S, 125°25.53' W; recovery 13.98 m; 3.613 m water depth), located north of the modern Subantarctic Front. Our study focuses on the temperature and salinity variability controlled by SOIWs, which were subducted at the Subantarctic Front during the Last Glacial Maximum (LGM; 29-17ka BP) and the Penultimate Glacial Maximum (PGM; 180-150ka BP). During both glacial periods conditions at the subsurface ocean were colder and fresher relative to the Holocene (<10ka) suggesting an enhanced presence of SOIWs. In spite of the comparable subsurface cooling during both glacial, the subsurface ocean during the PGM was saltier and 0.35‰ more depleted in δ13C in comparison to the LGM. Interestingly, the mean δ13C value of the PGM is comparable to the Carbon Isotope Minimum Events, which might suggests a larger contribution of "old" low δ13C deep waters to the study site during the PGM. A Latitudinal comparison of subsurface proxies suggests glacial asymmetries in the advection of SOIWs into the central Pacific, plausibly related to glacial changes in the convection depth of SOIWs at the South Antarctic Front area rather than changes in production of the SOIWs.

  14. Rapid Expansion of Glacial Lakes Caused by Climate and Glacier Retreat in the Central Himalayas

    NASA Astrophysics Data System (ADS)

    Wang, W.

    2016-12-01

    Glacial lake outburst floods are among the most serious natural hazards in the Himalayas. Such floods are of high scientific and political importance because they exert trans-boundary impacts on bordering countries. The preparation of an updated inventory of glacial lakes and the analysis of their evolution are an important first step in assessment of hazards from glacial lake outbursts. Here, we report the spatiotemporal developments of the glacial lakes in the Poiqu River basin, a trans-boundary basin in the Central Himalayas, from 1976 to 2010 based on multi-temporal Landsat images. Studied glacial lakes are classified as glacierfed lakes and non-glacier-fed lakes according to their hydrologic connection to glacial watersheds. A total of 119 glacial lakes larger than 0.01 km2 with an overall surface area of 20.22 km2 (±10.8%) were mapped in 2010, with glacier-fed lakes being predominant in both number (69, 58.0%) and area (16.22 km2, 80.2%). We found that lakes connected to glacial watersheds (glacier-fed lakes) significantly expanded (122.1%) from 1976 to 2010, whereas lakes not connected to glacial watersheds (non-glacier-fed lakes) remained stable (+2.8%) during the same period. This contrast can be attributed to the impact of glaciers. Retreating glaciers not only supply meltwater to lakes but also leave space for them to expand. Compared with other regions of the Hindu Kush Himalayas (HKH), the lake area per glacier area in the Poiqu River basin was the highest. This observation might be attributed to the different climate regimes and glacier status along the HKH. The results presented in this study confirm the significant role of glacier retreat on the evolution of glacial lakes.

  15. Influence of glacial meltwater on global seawater δ234U

    NASA Astrophysics Data System (ADS)

    Arendt, Carli A.; Aciego, Sarah M.; Sims, Kenneth W. W.; Das, Sarah B.; Sheik, Cody; Stevenson, Emily I.

    2018-03-01

    We present the first published uranium-series measurements from modern Greenland Ice Sheet (GrIS) runoff and proximal seawater, and investigate the influence of glacial melt on global seawater δ234U over glacial-interglacial (g-ig) timescales. Climate reconstructions based on closed-system uranium-thorium (U/Th) dating of fossil corals assume U chemistry of seawater has remained stable over time despite notable fluctuations in major elemental compositions, concentrations, and isotopic compositions of global seawater on g-ig timescales. Deglacial processes increase weathering, significantly increasing U-series concentrations and changing the δ234U of glacial meltwater. Analyses of glacial discharge from GrIS outlet glaciers indicate that meltwater runoff has elevated U concentrations and differing 222Rn concentrations and δ234U compositions, likely due to variations in subglacial residence time. Locations with high δ234U have the potential to increase proximal seawater δ234U. To better understand the impact of bulk glacial melt on global seawater δ234U over time, we use a simple box model to scale these processes to periods of extreme deglaciation. We account for U fluxes from the GrIS, Antarctica, and large Northern Hemisphere Continental Ice Sheets, and assess sensitivity by varying melt volumes, duration and U flux input rates based on modern subglacial water U concentrations and compositions. All scenarios support the hypothesis that global seawater δ234U has varied by more than 1‰ through time as a function of predictable perturbations in continental U fluxes during g-ig periods.

  16. Hazard and Risk of Glacial Lake Outburst Floods in the Nepal Himalayas

    NASA Astrophysics Data System (ADS)

    Rounce, David; McKinney, Daene

    2016-04-01

    As the climate changes and glaciers continue to melt, the number of glacial lakes and the size of these lakes is rapidly increasing. These glacial lakes are contained by terminal moraines composed of debris, soil, and sometimes ice, which are susceptible to fail catastrophically and cause a glacial lake outburst flood (GLOF). Understanding the hazard and risk associated with these lakes is important for downstream communities and other stakeholders, e.g., hydroelectric companies. Unfortunately, existing methods that are used to assess GLOF hazards yield conflicting classifications, which leads to confusion amongst the stakeholders who these studies are meant to assist. This study assesses existing methods on potentially dangerous glacial lakes in Nepal and uses these methods to develop an objective and holistic risk & action framework that may be used to assist and prioritize risk-mitigation actions.

  17. Cosmogenic evidence for limited local LGM glacial expansion, Denton Hills, Antarctica

    NASA Astrophysics Data System (ADS)

    Joy, Kurt; Fink, David; Storey, Bryan; De Pascale, Gregory P.; Quigley, Mark; Fujioka, Toshiyuki

    2017-12-01

    The geomorphology of the Denton Hills provides insight into the timing and magnitude of glacial retreats in a region of Antarctica isolated from the influence of the East Antarctic ice sheet. We present 26 Beryllium-10 surface exposure ages from a variety of glacial and lacustrine features in the Garwood and Miers valleys to document the glacial history of the area from 10 to 286 ka. Our data show that the cold-based Miers, Joyce and Garwood glaciers retreated little since their maximum positions at 37.2 ± 6.9 (1σ n = 4), 35.1 ± 1.5 (1σ, n = 3) and 35.6 ± 10.1 (1σ, n = 6) ka respectively. The similar timing of advance of all three glaciers and the lack of a significant glacial expansion during the global LGM suggests a local LGM for the Denton Hills between ca. 26 and 51 ka, with a mean age of 36.0 ± 7.5 (1σ, n = 13) ka. A second cohort of exposure ages provides constraints to the behaviour of Glacial Lake Trowbridge that formerly occupied Miers Valley in the late Pleistocene. These data show active modification of the landscape from ∼20 ka until the withdrawal of ice from the valley mouths, and deposition of Ross Sea Drift, at 10-14 ka.

  18. The Glacial-Interglacial Monsoon Recorded by Speleothems from Sulawesi, Indonesia

    NASA Astrophysics Data System (ADS)

    Kimbrough, A. K.; Gagan, M. K.; Dunbar, G. B.; Krause, C.; Hantoro, W. S.; Cheng, H.; Edwards, R. L.; Shen, C. C.; Sun, H.; Cai, B.; Hellstrom, J. C.; Rifai, H.

    2015-12-01

    The Indo-Pacific Warm Pool is a primary source of heat and moisture to the global atmosphere and a key player in tropical and global climate variability. There is mounting evidence that atmospheric convection and oceanic processes in the tropics can modulate global climate on orbital and sub-orbital timescales. Glacial-interglacial cycles represent the largest natural climate changes over the last 800 kyr with each cycle terminated by rapid global warming and sea level rise. Our understanding of the role and response of tropical atmospheric convection during these periods of dramatic warming is limited. We present the first speleothem paleomonsoon record for southwest Sulawesi (5ºS, 119ºE), spanning two glacial-interglacial cycles, including glacial termination IV (~340 kyr BP) and both phases of termination III (~248 and ~220 kyr BP). This unique record is constructed from multiple stalagmites from two separate caves and is based on a multi-proxy approach (δ18O, δ13C, Mg/Ca, Sr/Ca) that provides insight into the mechanisms controlling Australian-Indonesian summer monsoon variability. Speleothem δ18O and trace element data indicate a rapid increase in rainfall at glacial terminations and wet interglacials. Terminations IV, III, and I are each characterized by an abrupt 3‰ decrease in δ18O. Variability in δ18O leading-in to glacial terminations is also similar, and corresponds to October insolation. Prior to deglaciation, there is a distinct shift to higher δ18O that is synchronized with weak monsoon intervals in Chinese speleothem records. The remarkably consistent pattern among terminations implies that the response of tropical convection to changing background climates is well regulated. Furthermore, we find that speleothem δ13C leads δ18O by ~5 kyr during glacial terminations. The early decrease in speleothem δ13C may reflect the response of tropical vegetation to rising atmospheric CO2 and temperature, rather than regional changes in rainfall.

  19. Pleistocene glacial evolution of Fuentes Carrionas (Cantabrian Range, NW Spain)

    NASA Astrophysics Data System (ADS)

    Pellitero, Ramon

    2014-05-01

    Fuentes Carrionas is a massif situated at the N of Spain, between Castilla y Leon and Cantabria regions. It is the second highest mountain massif of the Cantabrian Range after Picos de Europa, with peaks over 2500 m.a.s.l. and valleys well over 1000 m.a.s.l. Fuentes Carrionas was glaciated during Quaternary, and even during the Holocene and as far as Little Ice Age the presence of glaciers, or at least permafrost is controversial. Results from glacial geomorphology analysis of Fuentes Carrionas Massif are presented. Based on the interpretation of glacial landforms, glacial evolution since the Last Glacial Maximum until Pleistocene deglaciation is described. Four different glacial equilibrium phases are identified, the last one divided into two pulsations. Deglaciation process took place between 36 ka BP and 11 ka BP. Local Last Glacial Maximum is dated back to 36-38 ka. BP, therefore earlier than LGM. Glaciers reached 15 km. long and occupied valleys down to 1250 m.a.s.l. during this phase. By European LGM (20-18 ka.BP) glaciers had substantially retreated to fronts about 1700 m.a.s.l. A final stage with two marked pulsations shows only small glaciers located at cirques above 2000 m.a.s.l. and, finally, only small cirque glaciers at North and Northeast orientation above 2200 m.a.s.l. Both these phases have been correlated to Oldest and Younger Dryas, although no dates have been done yet. A palaeoenvironmental reconstruction is proposed, based on ELA (Equilibrium Line Altitude) rise. ELA has been calculated with the AAR method and 0.67 ratio. This reconstruction shows that temperatures ranged between 9°C and 10°C lower than present ones at the end of Pleistocene, depending on a precipitations variation between 30% higher and 20% lower than current ones. Further research will focus on these retreat phases, especially on Younger Dryas identification and reconstruction for this site and the rest of Cantabrian Range.

  20. Beryllium-10 dating of the duration and retreat of the last pinedale glacial sequence

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Gosse, J.C.; Klein, J.; Evenson, E.B.

    Accurate terrestrial glacial chronologies are needed for comparison with the marine record to establish the dynamics of global climate change during transitions from glacial to interglacial regimes. Cosmogenic beryllium-10 measurements in the Wind River Range indicate that the last glacial maximum (marine oxygen isotope stage 2) was achieved there by 21,700 {+-} 700 beryllium-10 years and lasted 5900 years. Ages of a sequence of recessional moraines and striated bedrock surfaces show that the initial deglaciation was rapid and that the entire glacial system retreated 33 kilometers to the cirque basin by 12,100 {+-} 500 beryllium-10 years.

  1. Young Adolescents' Positioning of Human Rights: Findings from Colombia, Northern Ireland, Republic of Ireland and the United States

    ERIC Educational Resources Information Center

    Barton, Keith C.

    2015-01-01

    This study investigated how young adolescents thought about the location of human rights issues and the nature of violations in differing geographic regions. Open-ended, task-based interviews were conducted with 116 students in Colombia, Northern Ireland, the Republic of Ireland and the United States. Although students in each location pointed to…

  2. Poisonings and clinical toxicology: a template for Ireland.

    PubMed

    Tormey, W P; Moore, T

    2013-03-01

    Poisons information is accessed around the clock in the British Isles from six centres of which two are in Ireland at Dublin and Belfast accompanied by consultant toxicologist advisory service. The numbers of calls in Ireland are down to about 40 per day due to easy access to online data bases. Access to Toxbase, the clinical toxicology database of the National Poisons Information Service is available to National Health Service (NHS) health professionals and to Emergency Departments and Intensive Care units in the Republic of Ireland. There are 59 Toxbase users in the Republic of Ireland and 99 % of activity originates in Emergency Departments. All United States Poison Control Centres primarily use Poisindex which is a commercial database from Thomson Reuters. Information on paracetamol, diazepam, analgesics and psycho-active compounds are the commonest queries. Data from telephone and computer accesses provide an indicator of future trends in both licit and illicit drug poisons which may direct laboratory analytical service developments. Data from National Drug-Related Deaths Index is the most accurate information on toxicological deaths in Ireland. Laboratory toxicology requirements to support emergency departments are listed. Recommendations are made for a web-based open access Toxbase or equivalent; for a co-location of poisons information and laboratory clinical toxicology; for the establishment of a National Clinical Toxicology Institute for Ireland; for a list of accredited medical advisors in clinical toxicology; for multidisciplinary case conferences in complex toxicology cases for coroners; for the establishment of a national clinical toxicology referral out-patients service in Ireland.

  3. Massive remobilization of permafrost carbon during post-glacial warming

    NASA Astrophysics Data System (ADS)

    Tesi, T.; Muschitiello, F.; Smittenberg, R. H.; Jakobsson, M.; Vonk, J. E.; Hill, P.; Andersson, A.; Kirchner, N.; Noormets, R.; Dudarev, O.; Semiletov, I.; Gustafsson, Ö.

    2016-11-01

    Recent hypotheses, based on atmospheric records and models, suggest that permafrost carbon (PF-C) accumulated during the last glaciation may have been an important source for the atmospheric CO2 rise during post-glacial warming. However, direct physical indications for such PF-C release have so far been absent. Here we use the Laptev Sea (Arctic Ocean) as an archive to investigate PF-C destabilization during the last glacial-interglacial period. Our results show evidence for massive supply of PF-C from Siberian soils as a result of severe active layer deepening in response to the warming. Thawing of PF-C must also have brought about an enhanced organic matter respiration and, thus, these findings suggest that PF-C may indeed have been an important source of CO2 across the extensive permafrost domain. The results challenge current paradigms on the post-glacial CO2 rise and, at the same time, serve as a harbinger for possible consequences of the present-day warming of PF-C soils.

  4. Tectonic control on the persistence of glacially sculpted topography

    PubMed Central

    Prasicek, Günther; Larsen, Isaac J.; Montgomery, David R.

    2015-01-01

    One of the most fundamental insights for understanding how landscapes evolve is based on determining the extent to which topography was shaped by glaciers or by rivers. More than 104 years after the last major glaciation the topography of mountain ranges worldwide remains dominated by characteristic glacial landforms such as U-shaped valleys, but an understanding of the persistence of such landforms is lacking. Here we use digital topographic data to analyse valley shapes at sites worldwide to demonstrate that the persistence of U-shaped valleys is controlled by the erosional response to tectonic forcing. Our findings indicate that glacial topography in Earth's most rapidly uplifting mountain ranges is rapidly replaced by fluvial topography and hence valley forms do not reflect the cumulative action of multiple glacial periods, implying that the classic physiographic signature of glaciated landscapes is best expressed in, and indeed limited by, the extent of relatively low-uplift terrain. PMID:26271245

  5. Temporary refugia for coral reefs in a warming world

    NASA Astrophysics Data System (ADS)

    van Hooidonk, R.; Maynard, J. A.; Planes, S.

    2013-05-01

    Climate-change impacts on coral reefs are expected to include temperature-induced spatially extensive bleaching events. Bleaching causes mortality when temperature stress persists but exposure to bleaching conditions is not expected to be spatially uniform at the regional or global scale. Here we show the first maps of global projections of bleaching conditions based on ensembles of IPCC AR5 (ref. ) models forced with the new Representative Concentration Pathways (RCPs). For the three RCPs with larger CO2 emissions (RCP 4.5, 6.0 and 8.5) the onset of annual bleaching conditions is associated with ~ 510ppm CO2 equivalent; the median year of all locations is 2040 for the fossil-fuel aggressive RCP 8.5. Spatial patterns in the onset of annual bleaching conditions are similar for each of the RCPs. For RCP 8.5, 26% of reef cells are projected to experience annual bleaching conditions more than 5 years later than the median. Some of these temporary refugia include the western Indian Ocean, Thailand, the southern Great Barrier Reef and central French Polynesia. A reduction in the growth of greenhouse-gas emissions corresponding to the difference between RCP 8.5 and 6.0 delays annual bleaching in ~ 23% of reef cells more than two decades, which might conceivably increase the potential for these reefs to cope with these changes.

  6. Brook trout movement in response to temperature, flow, and thermal refugia within a complex Appalachian riverscape

    USGS Publications Warehouse

    Petty, J. Todd; Hansbarger, Jeff L.; Huntsman, Brock M.; Mazik, Patricia M.

    2012-01-01

    We quantified movements of brook trout Salvelinus fontinalis and brown trout Salmo trutta in a complex riverscape characterized by a large, open-canopy main stem and a small, closed-canopy tributary in eastern West Virginia, USA. Our objectives were to quantify the overall rate of trout movement and relate movement behaviors to variation in streamflow, water temperature, and access to coldwater refugia. The study area experienced extremely high seasonal, yearly, and among-stream variability in water temperature and flow. The relative mobility of brook trout within the upper Shavers Fork watershed varied significantly depending on whether individuals resided within the larger main stem or the smaller tributary. The movement rate of trout inhabiting the main stem during summer months (50 m/d) was an order of magnitude higher than that of tributary fish (2 m/d). Movement rates of main-stem-resident brook trout during summer were correlated with the maximum water temperature experienced by the fish and with the fish's initial distance from a known coldwater source. For main-stem trout, use of microhabitats closer to cover was higher during extremely warm periods than during cooler periods; use of microhabitats closer to cover during warm periods was also greater for main-stem trout than for tributary inhabitants. Main-stem-resident trout were never observed in water exceeding 19.5°C. Our study provides some of the first data on brook trout movements in a large Appalachian river system and underscores the importance of managing trout fisheries in a riverscape context. Brook trout conservation in this region will depend on restoration and protection of coldwater refugia in larger river main stems as well as removal of barriers to trout movement near tributary and main-stem confluences.

  7. Boulder Clusters as Flow Refugia for Juvenile Salmonids and Aquatic Invertebrates in Steep Mountain Streams, Klamath Mountains, Northern California

    NASA Astrophysics Data System (ADS)

    Cover, M. R.; May, C. L.; Dietrich, W. E.; Resh, V. H.

    2005-12-01

    The availability of flow refugia and cover is an important factor affecting habitat suitability for fish and invertebrates, especially in steep, turbulent streams. In some channels, crevices beneath and between large rocks may be the only available flow refugia that allow rainbow trout (Oncorhynchus mykiss) to conserve energy and escape from high velocity flow during large storm events. Many aquatic invertebrates, especially large or crawling taxa, require cover that is provided by unembedded crevice space underneath large stones. To investigate the influence of channel type on habitat availability, we performed intensive surveys of crevice habitat for salmonids and benthic invertebrates in 12 reaches in Walker Creek, a 25 square km basin in the Klamath Mountains of Northern California. We identified four reaches in each of three channel types: plane bed (3.1% - 3.7% slope), step-pool (5.4% - 6.5% slope), and cascade (6.3% - 8.5% slope). We used 4 realistic fish models (5, 10, 15, and 20 cm length) to assess the size of crevices and presence of flow refugia associated with all cobble (64 - 256 mm) and boulder (> 256 mm) grains within five 0.5 m-wide diagonal transects. The total abundance of crevices was similar among plane bed (6.3 +/- 1.1 m-2) (Mean +/- SD), step-pool (6.2 +/- 0.25 m-2), and cascade (6.7 +/- 1.2 m-2) reaches. Small (5 cm) crevices made up the majority of crevices in all three reach types. While the presence of 5 cm and 10 cm crevices was not significantly different between the three channel types, there were significantly more large (20 cm) crevices in cascade (0.73 +/- 0.33 m-2) and step-pool (0.68 +/- 0.1 m-2) reaches than in plane bed (0.26 +/- 0.14 m-2) reaches (AVOVA, p < 0.05). Moderately sized (15 cm) crevices were more common in step-pool reaches (0.91 +/- 0.13 m-2) than either cascade (0.54 +/- 0.15 m-2) or plane bed (0.42 +/- 0.13 m-2) reaches. Based on these results we conclude that step-pool reaches provide the most favorable habitat

  8. Sedimentological Characteristics and Classification of Depositional Processes and Deposits in the Glacial Environment,

    DTIC Science & Technology

    1981-12-01

    I characteristics and classification of depositional processes and d,4, r -%sits in the glacial environment C",. 44k (1-I J For conversion of SI metric...Discussion with Dr. John Shaw, Dr. Geoffrey Boulton, Dr. David Croot and Dr. Ross Powell helped considerably in formulating ideas presented in this report...glacial or non- glacial origins of diamictites of Precambrian and COMPARISON OF MELT-OUT other ages (e.g., Schermerhorn 1974, Edwards AND SEDIMENT FLOW

  9. Communications, Corporatism, and Dependent Development in Ireland.

    ERIC Educational Resources Information Center

    Bell, Desmond

    1995-01-01

    Explores the complex relationship between the corporate state and public service broadcasting in Europe, using a historical case study of Ireland. Discusses the Pan-European debate; public service broadcasting; critical media theory and the state; broadcasting and the corporate state; the corporate state and broadcasting in Ireland; neo-liberal…

  10. Cryptic no more: soil macrofossils uncover Pleistocene forest microrefugia within a periglacial desert.

    PubMed

    de Lafontaine, Guillaume; Amasifuen Guerra, Carlos Alberto; Ducousso, Alexis; Petit, Rémy J

    2014-11-01

    Despite their critical importance for understanding the local effects of global climate change on biodiversity, glacial microrefugia are not well studied because they are difficult to detect by using classical palaeoecological or population genetics approaches. We used soil macrofossil charcoal analysis to uncover the presence of cryptic glacial refugia for European beech (Fagus sylvatica) and other tree species in the Landes de Gascogne (southwestern France). Using botanical identification and direct radiocarbon dating (140 (14) C-dates) of macrofossil charcoal extracted from mineral soils, we reconstructed the glacial and postglacial history of all extant beech stands in the region (n = 11). Soil charcoal macrofossils were found in all sites, allowing the identification of up to at least 14 distinct fire events per site. There was direct evidence of the presence of beech during the last glacial period at three sites. Beech was detected during Heinrich stadial-1, one of the coldest and driest intervals of the last glacial period in Western Europe. Together with previous results on the genetic structure of the species in the region, these findings suggest that beech persisted in situ in several microrefugia through full glacial and interglacial periods up to the present day. © 2014 The Authors. New Phytologist © 2014 New Phytologist Trust.

  11. Assessing glacial lake outburst flood risk

    NASA Astrophysics Data System (ADS)

    Kougkoulos, Ioannis; Cook, Simon; Jomelli, Vincent; Clarke, Leon; Symeonakis, Elias

    2017-04-01

    Glaciers across the world are thinning and receding in response to atmospheric warming. Glaciers tend to erode subglacial basins and deposit eroded materials around their margins as lateral-frontal terminal moraines. Recession into these basins and behind impounding moraines causes meltwater to pond as proglacial and supraglacial lakes. Consequently, there has been a general trend of increasing number and size of these lakes associated with glacier melting in many mountainous regions around the globe, in the last 30 years. Glacial lake outburst floods (GLOFs) then may occur where the glacial lake dam (ice, rock, moraine, or combination thereof) is breached, or overtopped, and thousands of people have lost their lives to such events in the last few decades, especially in the Andes and in the Himalaya. Given the ongoing and arguably increasing risk posed to downstream communities, and infrastructure, there has been a proliferation of GLOF studies, with many seeking to estimate GLOF hazard or risk in specific regions, or to identify 'potentially dangerous glacial lakes'. Given the increased scientific interest in GLOFs, it is timely to evaluate critically the ways in which GLOF risk has been assessed previously, and whether there are improvements that can be made to the ways in which risk assessment is achieved. We argue that, whilst existing GLOF hazard and risk assessments have been extremely valuable they often suffer from a number of key shortcomings that can be addressed by using different techniques as multi-criteria decision analysis and hydraulic modelling borrowed from disciplines like engineering, remote sensing and operations research.

  12. Decadal-scale climate drivers for glacial dynamics in Glacier National Park, Montana, USA

    USGS Publications Warehouse

    Pederson, G.T.; Fagre, D.B.; Gray, S.T.; Graumlich, L.J.

    2004-01-01

    Little Ice Age (14th-19th centuries A.D.) glacial maxima and 20th century retreat have been well documented in Glacier National Park, Montana, USA. However, the influence of regional and Pacific Basin driven climate variability on these events is poorly understood. We use tree-ring reconstructions of North Pacific surface temperature anomalies and summer drought as proxies for winter glacial accumulation and summer ablation, respectively, over the past three centuries. These records show that the 1850's glacial maximum was likely produced by ???70 yrs of cool/wet summers coupled with high snowpack. Post 1850, glacial retreat coincides with an extended period (>50 yr) of summer drought and low snowpack culminating in the exceptional events of 1917 to 1941 when retreat rates for some glaciers exceeded 100 m/yr. This research highlights potential local and ocean-based drivers of glacial dynamics, and difficulties in separating the effects of global climate change from regional expressions of decadal-scale climate variability. Copyright 2004 by the American Geophysical Union.

  13. Pollen record of the penultimate glacial period in Yuchi Basin, Central Taiwan

    NASA Astrophysics Data System (ADS)

    Lai, Hsiao-Yin; Liew, Ping-Mei

    2010-05-01

    Pollen records of the penultimate glacial period are scare not only in Taiwan, but also in East Asia area. Hence, this study intends to provide a new pollen record from a site, Yuchi Basin, in central Taiwan, which may improve our knowledge of the penultimate glacial period. The sediment core, CTN6, was drilled in the northern part of Yuchi Basin. The core is 29.4 m in length and the sampling interval is 10 cm. In total, 86 samples are processed for pollen analysis. Three pollen zones (I,II and III) are determined according to the ratio of arboreal pollens (AP) and non-arboreal pollens (NAP). Because of the scarcity of dating data, pollen assemblages compared with previous pollen records at peripheral areas is utilized to estimate the ages of each pollen zone. AP dominate (60%) Zone I and III, which consist mainly of Cyclobalanopsis-Castanopsis. Thus, Zone I may mark the MIS 5 because of a Cyclobalanopsis-Castanopsis dominant condition. In Zone II, the increase in NAP and pollen of Taxodiaceae and decrease in pollens of Cyclobalanopsis-Castanopsis indicates the penultimate glacial period, i.e. MIS 6. In contrast to the evergreen broadleaved forest found there today, the herbs occupied the basin in Zone II, indicating a relatively dry climate condition than present. Furthermore, during the penultimate glacial period, the climate condition of early part is wetter, evidenced by a higher AP/NAP in Zone IIb. Finally, comparing with the last glacial period in Toushe, we suggest that the penultimate glacial period is drier due to the lower AP/NAP.

  14. Role of Southern Ocean stratification in glacial atmospheric CO2 reduction

    NASA Astrophysics Data System (ADS)

    Kobayashi, H.; Oka, A.

    2014-12-01

    Paleoclimate proxy data at the glacial period shows high salinity of more than 37.0 psu in the deep South Atlantic. At the same time, data also indicate that the residence time of the water mass was more than 3000 years. These data implies that the stratification by salinity was stronger in the deep Southern Ocean (SO) in the Last Glacial Maximum (LGM). Previous studies using Ocean General Circulation Model (OGCM) fail to explain the low glacial atmospheric carbon dioxide (CO2) concentration at LGM. The reproducibility of salinity and water mass age is considered insufficient in these OGCMs, which may in turn affect the reproducibility of the atmospheric CO2concentration. In coarse-resolution OGCMs, The deep water is formed by unrealistic open-ocean deep convection in the SO. Considering these facts, we guessed previous studies using OGCM underestimated the salinity and water mass age at LGM. This study investigate the role of the enhanced stratification in the glacial SO on the variation of atmospheric CO2 concentration by using OGCM. In order to reproduce the recorded salinity of the deep water, relaxation of salinity toward value of recorded data is introduced in our OGCM simulations. It was found that deep water formation in East Antarctica is required for explaining the high salinity in the South Atlantic. In contrast, it is difficult to explain the glacial water mass age, even if we assume the situation vertical mixing is very weak in the SO. Contrary to previous estimate, the high salinity of the deep SO resulted in increase of Antarctic Bottom water (AABW) flow and decrease the residence time of carbon in the deep ocean, which increased atmospheric CO2 concentration. On the other hand, the weakening of the vertical mixing in the SO contributed to increase the vertical gradient of dissolved inorganic carbon (DIC), which decreased atmospheric CO2 concentration. Adding the contribution of the enhanced stratification in the glacial SO, we obtained larger

  15. An overview of cleaner fish use in Ireland.

    PubMed

    Bolton-Warberg, M

    2017-11-21

    Sea lice infestations represent one of the most significant challenges facing the salmon farming industry, giving rise to lost production, additional costs of treatment and potential negative interactions with wild stocks. At present, cleaner fish, which actively remove lice from salmon, are an effective, biological, long-term option which has been adopted by many countries. In Ireland, several key studies were conducted in the 1990s on the use of wild-caught wrasse (corkwing, goldsinny and rock cook) as cleaner fish in experimental and commercial scale trials. More recently, the National University of Ireland Galway (NUIG), at their marine research facility in Carna (CRS), has undertaken applied research on ballan wrasse and lumpsucker. Currently, CRS is providing lumpsucker juveniles and research and development for the Irish salmon industry with support from BIM (Ireland's Seafood Development Agency) and Marine Harvest Ireland. There is a large amount of research currently being carried out in this area in all countries that are utilizing cleaner fish technology. The current focus in Ireland is the development of a native lumpsucker broodstock to facilitate its sustainable production. The aim of this article was to provide an overview of the research, challenges and use of cleaner fish in Ireland. © 2017 The Authors Journal of Fish Diseases Published by John Wiley & Sons Ltd.

  16. Sediment core and glacial environment reconstruction - a method review

    NASA Astrophysics Data System (ADS)

    Bakke, Jostein; Paasche, Øyvind

    2010-05-01

    Alpine glaciers are often located in remote and high-altitude regions of the world, areas that only rarely are covered by instrumental records. Reconstructions of glaciers has therefore proven useful for understanding past climate dynamics on both shorter and longer time-scales. One major drawback with glacier reconstructions based solely on moraine chronologies - by far the most common -, is that due to selective preservation of moraine ridges such records do not exclude the possibility of multiple Holocene glacier advances. This problem is true regardless whether cosmogenic isotopes or lichenometry have been used to date the moraines, or also radiocarbon dating of mega-fossils buried in till or underneath the moraines themselves. To overcome this problem Karlén (1976) initially suggested that glacial erosion and the associated production of rock-flour deposited in downstream lakes could provide a continuous record of glacial fluctuations, hence overcoming the problem of incomplete reconstructions. We want to discuss the methods used to reconstruct past glacier activity based on sediments deposited in distal glacier-fed lakes. By quantifying physical properties of glacial and extra-glacial sediments deposited in catchments, and in downstream lakes and fjords, it is possible to isolate and identify past glacier activity - size and production rate - that subsequently can be used to reconstruct changing environmental shifts and trends. Changes in average sediment evacuation from alpine glaciers are mainly governed by glacier size and the mass turnover gradient, determining the deformation rate at any given time. The amount of solid precipitation (mainly winter accumulation) versus loss due to melting during the ablation-season (mainly summer temperature) determines the mass turnover gradient in either positive or negative direction. A prevailing positive net balance will lead to higher sedimentation rates and vice versa, which in turn can be recorded in downstream

  17. Glacial/interglacial changes in subarctic north pacific stratification.

    PubMed

    Jaccard, S L; Haug, G H; Sigman, D M; Pedersen, T F; Thierstein, H R; Röhl, U

    2005-05-13

    Since the first evidence of low algal productivity during ice ages in the Antarctic Zone of the Southern Ocean was discovered, there has been debate as to whether it was associated with increased polar ocean stratification or with sea-ice cover, shortening the productive season. The sediment concentration of biogenic barium at Ocean Drilling Program site 882 indicates low algal productivity during ice ages in the Subarctic North Pacific as well. Site 882 is located southeast of the summer sea-ice extent even during glacial maxima, ruling out sea-ice-driven light limitation and supporting stratification as the explanation, with implications for the glacial cycles of atmospheric carbon dioxide concentration.

  18. Endemism hotspots are linked to stable climatic refugia.

    PubMed

    Harrison, Susan; Noss, Reed

    2017-01-01

    Centres of endemism have received much attention from evolutionists, biogeographers, ecologists and conservationists. Climatic stability is often cited as a major reason for the occurrences of these geographic concentrations of species which are not found anywhere else. The proposed linkage between endemism and climatic stability raises unanswered questions about the persistence of biodiversity during the present era of rapidly changing climate. The current status of evidence linking geographic centres of endemism to climatic stability over evolutionary time was examined. The following questions were asked. Do macroecological analyses support such an endemism-stability linkage? Do comparative studies find that endemic species display traits reflecting evolution in stable climates? Will centres of endemism in microrefugia or macrorefugia remain relatively stable and capable of supporting high biological diversity into the future? What are the implications of the endemism-stability linkage for conservation? Recent work using the concept of climate change velocity supports the classic idea that centres of endemism occur where past climatic fluctuations have been mild and where mountainous topography or favourable ocean currents contribute to creating refugia. Our knowledge of trait differences between narrow endemics and more widely distributed species remains highly incomplete. Current knowledge suggests that centres of endemism will remain relatively climatically buffered in the future, with the important caveat that absolute levels of climatic change and species losses in these regions may still be large. © The Author 2017. Published by Oxford University Press on behalf of the Annals of Botany Company. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

  19. Ocean Acidification Refugia of the Florida Reef Tract

    NASA Astrophysics Data System (ADS)

    Manzello, D.; Enochs, I.; Melo, N.; Gledhill, D. K.; Johns, E. M.

    2012-12-01

    Ocean acidification (OA) is expected to reduce the calcification rates of marine organisms, yet we have little understanding of how OA will manifest within dynamic, real-world systems. Natural CO2, alkalinity, and salinity gradients can significantly alter local carbonate chemistry, and thereby create a range of susceptibility for different ecosystems to OA. As such, there is a need to characterize this natural variability of seawater carbonate chemistry, especially within coastal ecosystems. Since 2009, carbonate chemistry data have been collected on the Florida Reef Tract (FRT). During periods of heightened productivity, there is a net uptake of total CO2 (TCO2) which increases aragonite saturation state (Ωarag) values on inshore patch reefs of the upper FRT. These waters can exhibit greater Ωarag than what has been modeled for the tropical surface ocean during preindustrial times, with mean (± std. error) Ωarag-values in spring = 4.69 (± 0.101). Conversely, Ωarag-values on offshore reefs generally represent oceanic carbonate chemistries consistent with present day tropical surface ocean conditions. This gradient is opposite from what has been reported for other reef environments. We hypothesize this pattern is caused by the photosynthetic uptake of TCO2 mainly by seagrasses and, to a lesser extent, macroalgae in the inshore waters of the FRT. These inshore reef habitats are therefore potential acidification refugia that are defined not only in a spatial sense, but also in time; coinciding with seasonal productivity dynamics. Coral reefs located within or immediately downstream of seagrass beds may find refuge from OA.

  20. Ocean Acidification Refugia of the Florida Reef Tract

    PubMed Central

    Manzello, Derek P.; Enochs, Ian C.; Melo, Nelson; Gledhill, Dwight K.; Johns, Elizabeth M.

    2012-01-01

    Ocean acidification (OA) is expected to reduce the calcification rates of marine organisms, yet we have little understanding of how OA will manifest within dynamic, real-world systems. Natural CO2, alkalinity, and salinity gradients can significantly alter local carbonate chemistry, and thereby create a range of susceptibility for different ecosystems to OA. As such, there is a need to characterize this natural variability of seawater carbonate chemistry, especially within coastal ecosystems. Since 2009, carbonate chemistry data have been collected on the Florida Reef Tract (FRT). During periods of heightened productivity, there is a net uptake of total CO2 (TCO2) which increases aragonite saturation state (Ωarag) values on inshore patch reefs of the upper FRT. These waters can exhibit greater Ωarag than what has been modeled for the tropical surface ocean during preindustrial times, with mean (± std. error) Ωarag-values in spring = 4.69 (±0.101). Conversely, Ωarag-values on offshore reefs generally represent oceanic carbonate chemistries consistent with present day tropical surface ocean conditions. This gradient is opposite from what has been reported for other reef environments. We hypothesize this pattern is caused by the photosynthetic uptake of TCO2 mainly by seagrasses and, to a lesser extent, macroalgae in the inshore waters of the FRT. These inshore reef habitats are therefore potential acidification refugia that are defined not only in a spatial sense, but also in time; coinciding with seasonal productivity dynamics. Coral reefs located within or immediately downstream of seagrass beds may find refuge from OA. PMID:22848575

  1. Ocean acidification refugia of the Florida Reef Tract.

    PubMed

    Manzello, Derek P; Enochs, Ian C; Melo, Nelson; Gledhill, Dwight K; Johns, Elizabeth M

    2012-01-01

    Ocean acidification (OA) is expected to reduce the calcification rates of marine organisms, yet we have little understanding of how OA will manifest within dynamic, real-world systems. Natural CO(2), alkalinity, and salinity gradients can significantly alter local carbonate chemistry, and thereby create a range of susceptibility for different ecosystems to OA. As such, there is a need to characterize this natural variability of seawater carbonate chemistry, especially within coastal ecosystems. Since 2009, carbonate chemistry data have been collected on the Florida Reef Tract (FRT). During periods of heightened productivity, there is a net uptake of total CO(2) (TCO(2)) which increases aragonite saturation state (Ω(arag)) values on inshore patch reefs of the upper FRT. These waters can exhibit greater Ω(arag) than what has been modeled for the tropical surface ocean during preindustrial times, with mean (± std. error) Ω(arag)-values in spring = 4.69 (±0.101). Conversely, Ω(arag)-values on offshore reefs generally represent oceanic carbonate chemistries consistent with present day tropical surface ocean conditions. This gradient is opposite from what has been reported for other reef environments. We hypothesize this pattern is caused by the photosynthetic uptake of TCO(2) mainly by seagrasses and, to a lesser extent, macroalgae in the inshore waters of the FRT. These inshore reef habitats are therefore potential acidification refugia that are defined not only in a spatial sense, but also in time; coinciding with seasonal productivity dynamics. Coral reefs located within or immediately downstream of seagrass beds may find refuge from OA.

  2. Core-seismic investigation of Surveyor Channel tributaries: Glacial history of the southern Alaskan margin

    NASA Astrophysics Data System (ADS)

    Somchat, K.; Reece, R.; Gulick, S. P. S.; Asahi, H.; Mix, A. C.

    2016-12-01

    The low angle subduction and collision of the Yakutat microplate with the North America Plate created, and continues to contribute to the uplift of the Chugach-St. Elias Range. This heavily glaciated, high topography proximal to the shoreline creates a unique source-to-sink system in which glacial sediment is transported and preserved offshore in a deep sea fan without much interruption. The product of this sediment is the Surveyor Fan and Channel system. Four tributary channels form the head of the Surveyor Channel complex and merge into the main channel trunk 200 km from the shelf edge. We integrate drill core and seismic reflection data to study the evolution of these tributaries in order to decipher glacial history of the southern Alaskan margin since the mid-Pleistocene (1.2 Ma). Updated age models from Integrated Ocean Drilling Program Expedition 341 Sites U1417 and U1418 provide a higher resolution chronology of sediment delivery to the Surveyor Fan than previous studies. We regionally extended the mapping of seismic subunits previously identified by Exp. 341 scientists at sites U1417 and U1418 and analyzed regional patterns of sediment deposition. Two-way travel time (isopach) maps of the three subunits show a trend of sediment depocenter shifting to the east since 1.2 Ma, where the Yakutat and Alsek tributaries have increasing sediment flux through time. Changes in sediment flux in each system represent the changes in locations and amplitudes of glacial ice over successive glacial intervals. Additionally, seismic analysis of channel geomorphology shows that each system contains distinct geomorphological evolutions. Since glacial erosion provides the sediment for the fan, the history of glacial ice onshore can be inferred from seismic geomorphology, where changes in glacial ice affect sediment supply and therefore shifts in depocenters and sedimentation pathways. This study shows an interaction between glacial activity onshore and deep sea fan sediment

  3. Continuous proxy measurements reveal large mercury fluxes from glacial and forested watersheds in Alaska.

    PubMed

    Vermilyea, Andrew W; Nagorski, Sonia A; Lamborg, Carl H; Hood, Eran W; Scott, Durelle; Swarr, Gretchen J

    2017-12-01

    In this study, a stream from a glacially dominated watershed and one from a wetland, temperate forest dominated watershed in southeast Alaska were continuously monitored for turbidity and fluorescence from dissolved organic matter (FDOM) while grab samples for unfiltered (UTHg), particulate (PTHg), and filtered mercury (FTHg) where taken over three 4-day periods (May snowmelt, July glacial melt, and September rainy season) during 2010. Strong correlations were found between FDOM and UTHg concentrations in the wetland, temperate forest watershed (r 2 =0.81), while turbidity and UTHg were highly correlated in the glacially dominated watershed (r 2 =0.82). Both of these parameters (FDOM and turbidity) showed stronger correlations than concentration-discharge relationships for UTHg (r 2 =0.55 for glacial stream, r 2 =0.38 for wetland/forest stream), thus allowing for a more precise determination of temporal variability in UTHg concentrations and fluxes. The association of mercury with particles and dissolved organic matter (DOM) appears to depend on the watershed characteristics, such as physical weathering and biogeochemical processes regulating mercury transport. Thus employing watershed-specific proxies for UTHg (such as FDOM and turbidity) can be effective for quantifying mercury export from watersheds with variable landcover. The UTHg concentration in the forest/wetland stream was consistently higher than in the glacial stream, in which most of the mercury was associated with particles; however, due to the high specific discharge from the glacial stream during the melt season, the watershed area normalized flux of mercury from the glacial stream was 3-6 times greater than the wetland/forest stream for the three sampling campaigns. The annual specific flux for the glacial watershed was 19.9gUTHgkm -2 y -1 , which is higher than any non-mining impacted stream measured to date. This finding indicates that glacial watersheds of southeast Alaska may be important

  4. Quaternary geology of the Boston area: Glacial events from Lake Charles to Lake Aberjona

    USGS Publications Warehouse

    Stone, Byron D.; Lane, John W.

    2014-01-01

    The multiple-glacial and glaciomarine Quaternary history of the Boston, Massachusetts area has been known generally since the earliest studies of the then newly recognized glacial deposits described by Prof. Louis Agassiz in the late1840’s and fossil marine shells in the drift in the 1850’s. Attention then turned to possible glacial erosional effects on the preglacial bedrock physiography, as related to rock units and structure, and to the challenges of defining useful physical and lithic characteristics of the drift by Prof. W.O. Crosby and others, 1880-1900. The problems of deducing the relative stratigraphic order among such small, fossil-barren surficial sedimentary deposits, and extending knowledge gained from studies of postulated ancient glacial lakes to a regional understanding of the history of many lakes during the retreat of the ice sheet required field work and use of geologic maps. With the advent of modern topographic maps in the 1880’s, the early period of discovery included field studies of glacial lake deposits in local river basins in the Boston region, basins that drain northward, thereby creating glacial lake basins dammed by the ice margin as it retreated to the north. Guided by M.I.T. and Harvard professors W.O. Crosby, N.S. Shaler, J.B. Woodworth, W.M. Davis, and others in the 1880-1920 period, the first Quaternary glacial stratigraphers were students (e.g. Crosby and Grabau, 1896, Clapp, 1905, Fuller 1905, Goldthwaite 1906, Grabau, 1906, Taylor, Tight).

  5. Monitoring the dynamics of glacial lakes in the High Mountain Asia region through time series Landsat images

    NASA Astrophysics Data System (ADS)

    Zhang, M.; Chen, F.

    2017-12-01

    Glacial lakes have been developing dramatically in the High Mountain Asia (HMA) region associated with human activities and persistent climatic warming. This leads to increased probability of glacial lake outburst floods (GLOF), pose potential threats to the downstream lives and properties of people. However, comprehensive information is lacking about the annual distribution, evolution and the driving mechanism of glacial lakes in the entire HMA due to the low accessibility and harsh natural conditions, with most studies focused either on certain portion of this region or at most several time intervals effort at monitoring glacial lakes at coarse resolution remote sensing. In this research, we produce yearly map of glacial lake extents in HMA from 2008 to 2016 using Landsat series satellites images, and further study the formation, distribution and dynamics of glacial lakes. In total 6197 and 8256 glacial lakes were detected in 2008 and 2016, respectively, mainly located at altitudes between 4400 m and 5600 m. The annual expansion rate is approximately 4.68 % from 2008 to 2016. To explore the cause of rapid expansion for some typical glacial lakes, we investigated their changing patterns through long-term expansion rates measured from change in shoreline positions. The results show that glacial lake expansion rates at some points change substantially (> 30 m/yr) and the formation of proglacial lakes may be dominated by different orientation-driving forces from parent glacier. The accelerating rate of ice and snow melting from glacier caused by global warming are primary contributor to glacial lake growth. The results may provide information for understanding the mechanism of lake dynamics, which also facilitate the scientific recognition of the potential glacial lakes hazards in this region.

  6. Effect of en-glacial water on ice sheet temperatures in a warming climate - a model approach

    NASA Astrophysics Data System (ADS)

    Phillips, T. P.; Rajaram, H.; Steffen, K.

    2009-12-01

    Each summer, significant amount of melt is generated in the ablation zones of large glaciers and ice sheets. This melt does not run off on the surface of the glacier or ice sheet. In fact a significant fraction enters the glacier and flows through en-glacial and sub-glacial hydrologic systems. Correspondingly, the en-glacial and sub-glacial hydrologic systems are brought to a temperature close to the pressure melting point of ice. The thermal influence of these hydrologic processes is seldom incorporated in heat transfer models for glaciers and ice sheets. In a warming climate, as melt water generation is amplified, en-glacial and sub-glacial hydrologic processes can influence the thermal dynamics of an ice sheet significantly, a feedback which is missed in current models. Although the role of refreezing melt water in the firn of the accumulation zone is often accounted for to explain warmer near-surface temperatures, the role of melt water flow within a glacier is not considered in large ice sheet models. We propose a simple parameterization of the influence of en-glacial and sub-glacial hydrology on the thermal dynamics of ice sheets, in the form of a dual-column model. Our model basically modifies the classical Budd column model for temperature variations in ice sheets by introducing an interaction with an en-glacial column, where the temperature is brought to the melting point during the melt season, and winter-time refreezing is influenced by latent heat effects associated with water retained within the en-glacial and sub-glacial systems. A cryo-hydraulic heat exchange coefficient ς is defined, as a parameter that quantifies this interaction. The parameter ς is related to k/R^2, where R is the characteristic spacing between en-glacial passages. The general behavior of the dual-column model is influenced by the competition between cooling by horizontal advection and warming by cryo-hydraulic exchange. We present a dimensionless parameter to quantify this

  7. Disablist Bullying in Northern Ireland and the Republic of Ireland: An Investigation of Student Teachers' Knowledge, Experience and Confidence

    ERIC Educational Resources Information Center

    Purdy, Noel; Mc Guckin, Conor

    2014-01-01

    This study explored the knowledge, experience and confidence of student teachers from Northern Ireland and the Republic of Ireland in relation to disablist bullying. Adopting a mixed methodological approach of four focus groups (N = 18) and a pencil-and-paper questionnaire (N = 257), the study explored the students knowledge, experience and…

  8. Phylogeographic Insights into a Peripheral Refugium: The Importance of Cumulative Effect of Glaciation on the Genetic Structure of Two Endemic Plants

    PubMed Central

    Zecca, Giovanni; Minuto, Luigi

    2016-01-01

    Quaternary glaciations and mostly last glacial maximum have shaped the contemporary distribution of many species in the Alps. However, in the Maritime and Ligurian Alps a more complex picture is suggested by the presence of many Tertiary paleoendemisms and by the divergence time between lineages in one endemic species predating the Late Pleistocene glaciation. The low number of endemic species studied limits the understanding of the processes that took place within this region. We used species distribution models and phylogeographical methods to infer glacial refugia and to reconstruct the phylogeographical pattern of Silene cordifolia All. and Viola argenteria Moraldo & Forneris. The predicted suitable area for last glacial maximum roughly fitted current known distribution. Our results suggest that separation of the major clades predates the last glacial maximum and the following repeated glacial and interglacial periods probably drove differentiations. The complex phylogeographical pattern observed in the study species suggests that both populations and genotypes extinction was minimal during the last glacial maximum, probably due to the low impact of glaciations and to topographic complexity in this area. This study underlines the importance of cumulative effect of previous glacial cycles in shaping the genetic structure of plant species in Maritime and Ligurian Alps, as expected for a Mediterranean mountain region more than for an Alpine region. PMID:27870888

  9. Millennial Climatic Fluctuations Are Key to the Structure of Last Glacial Ecosystems

    PubMed Central

    Huntley, Brian; Allen, Judy R. M.; Collingham, Yvonne C.; Hickler, Thomas; Lister, Adrian M.; Singarayer, Joy; Stuart, Anthony J.; Sykes, Martin T.; Valdes, Paul J.

    2013-01-01

    Whereas fossil evidence indicates extensive treeless vegetation and diverse grazing megafauna in Europe and northern Asia during the last glacial, experiments combining vegetation models and climate models have to-date simulated widespread persistence of trees. Resolving this conflict is key to understanding both last glacial ecosystems and extinction of most of the mega-herbivores. Using a dynamic vegetation model (DVM) we explored the implications of the differing climatic conditions generated by a general circulation model (GCM) in “normal” and “hosing” experiments. Whilst the former approximate interstadial conditions, the latter, designed to mimic Heinrich Events, approximate stadial conditions. The “hosing” experiments gave simulated European vegetation much closer in composition to that inferred from fossil evidence than did the “normal” experiments. Given the short duration of interstadials, and the rate at which forest cover expanded during the late-glacial and early Holocene, our results demonstrate the importance of millennial variability in determining the character of last glacial ecosystems. PMID:23613985

  10. Millennial climatic fluctuations are key to the structure of last glacial ecosystems.

    PubMed

    Huntley, Brian; Allen, Judy R M; Collingham, Yvonne C; Hickler, Thomas; Lister, Adrian M; Singarayer, Joy; Stuart, Anthony J; Sykes, Martin T; Valdes, Paul J

    2013-01-01

    Whereas fossil evidence indicates extensive treeless vegetation and diverse grazing megafauna in Europe and northern Asia during the last glacial, experiments combining vegetation models and climate models have to-date simulated widespread persistence of trees. Resolving this conflict is key to understanding both last glacial ecosystems and extinction of most of the mega-herbivores. Using a dynamic vegetation model (DVM) we explored the implications of the differing climatic conditions generated by a general circulation model (GCM) in "normal" and "hosing" experiments. Whilst the former approximate interstadial conditions, the latter, designed to mimic Heinrich Events, approximate stadial conditions. The "hosing" experiments gave simulated European vegetation much closer in composition to that inferred from fossil evidence than did the "normal" experiments. Given the short duration of interstadials, and the rate at which forest cover expanded during the late-glacial and early Holocene, our results demonstrate the importance of millennial variability in determining the character of last glacial ecosystems.

  11. The seasonal sea-ice zone in the glacial Southern Ocean as a carbon sink.

    PubMed

    Abelmann, Andrea; Gersonde, Rainer; Knorr, Gregor; Zhang, Xu; Chapligin, Bernhard; Maier, Edith; Esper, Oliver; Friedrichsen, Hans; Lohmann, Gerrit; Meyer, Hanno; Tiedemann, Ralf

    2015-09-18

    Reduced surface-deep ocean exchange and enhanced nutrient consumption by phytoplankton in the Southern Ocean have been linked to lower glacial atmospheric CO2. However, identification of the biological and physical conditions involved and the related processes remains incomplete. Here we specify Southern Ocean surface-subsurface contrasts using a new tool, the combined oxygen and silicon isotope measurement of diatom and radiolarian opal, in combination with numerical simulations. Our data do not indicate a permanent glacial halocline related to melt water from icebergs. Corroborated by numerical simulations, we find that glacial surface stratification was variable and linked to seasonal sea-ice changes. During glacial spring-summer, the mixed layer was relatively shallow, while deeper mixing occurred during fall-winter, allowing for surface-ocean refueling with nutrients from the deep reservoir, which was potentially richer in nutrients than today. This generated specific carbon and opal export regimes turning the glacial seasonal sea-ice zone into a carbon sink.

  12. The seasonal sea-ice zone in the glacial Southern Ocean as a carbon sink

    PubMed Central

    Abelmann, Andrea; Gersonde, Rainer; Knorr, Gregor; Zhang, Xu; Chapligin, Bernhard; Maier, Edith; Esper, Oliver; Friedrichsen, Hans; Lohmann, Gerrit; Meyer, Hanno; Tiedemann, Ralf

    2015-01-01

    Reduced surface–deep ocean exchange and enhanced nutrient consumption by phytoplankton in the Southern Ocean have been linked to lower glacial atmospheric CO2. However, identification of the biological and physical conditions involved and the related processes remains incomplete. Here we specify Southern Ocean surface–subsurface contrasts using a new tool, the combined oxygen and silicon isotope measurement of diatom and radiolarian opal, in combination with numerical simulations. Our data do not indicate a permanent glacial halocline related to melt water from icebergs. Corroborated by numerical simulations, we find that glacial surface stratification was variable and linked to seasonal sea-ice changes. During glacial spring–summer, the mixed layer was relatively shallow, while deeper mixing occurred during fall–winter, allowing for surface-ocean refueling with nutrients from the deep reservoir, which was potentially richer in nutrients than today. This generated specific carbon and opal export regimes turning the glacial seasonal sea-ice zone into a carbon sink. PMID:26382319

  13. Predicting Pleistocene climate from vegetation

    NASA Astrophysics Data System (ADS)

    Loehle, C.

    2006-10-01

    Climates at the Last Glacial Maximum have been inferred from fossil pollen assemblages, but these inferred climates are colder than those produced by climate simulations. Biogeographic evidence also argues against these inferred cold climates. The recolonization of glaciated zones in eastern North America following the last ice age produced distinct biogeographic patterns. It has been assumed that a wide zone south of the ice was tundra or boreal parkland (Boreal-Parkland Zone or BPZ), which would have been recolonized from southern refugia as the ice melted, but the patterns in this zone differ from those in the glaciated zone, which creates a major biogeographic anomaly. In the glacial zone, there are few endemics but in the BPZ there are many across multiple taxa. In the glacial zone, there are the expected gradients of genetic diversity with distance from the ice-free zone, but no evidence of this is found in the BPZ. Many races and related species exist in the BPZ which would have merged or hybridized if confined to the same refugia. Evidence for distinct southern refugia for most temperate species is lacking. Extinctions of temperate flora were rare. The interpretation of spruce as a boreal climate indicator may be mistaken over much of the region if the spruce was actually an extinct temperate species. All of these anomalies call into question the concept that climates in the zone south of the ice were very cold or that temperate species had to migrate far to the south. Similar anomalies exist in Europe and on tropical mountains. An alternate hypothesis is that low CO2 levels gave an advantage to pine and spruce, which are the dominant trees in the BPZ, and to herbaceous species over trees, which also fits the observed pattern. Most temperate species could have survived across their current ranges at lower abundance by retreating to moist microsites. These would be microrefugia not easily detected by pollen records, especially if most species became rare

  14. Surface exposure chronology of the Waimakariri glacial sequence in the Southern Alps of New Zealand: Implications for MIS-2 ice extent and LGM glacial mass balance

    NASA Astrophysics Data System (ADS)

    Rother, Henrik; Shulmeister, James; Fink, David; Alexander, David; Bell, David

    2015-11-01

    During the late Quaternary, the Southern Alps of New Zealand experienced multiple episodes of glaciation with large piedmont glaciers reaching the coastal plains in the west and expanding into the eastern alpine forelands. Here, we present a new 10Be exposure age chronology for a moraine sequence in the Waimakariri Valley (N-Canterbury), which has long been used as a reference record for correlating glacial events across New Zealand and the wider Southern Hemisphere. Our data indicate that the Waimakariri glacier reached its maximum last glaciation extent prior to ∼26 ka well before the global last glaciation maximum (LGM). This was followed by a gradual reduction in ice volume and the abandonment of the innermost LGM moraines at about 17.5 ka. Significantly, we find that during its maximum extent, the Waimakariri glacier overflowed the Avoca Plateau, previously believed to represent a mid-Pleistocene glacial surface (i.e. MIS 8). At the same time, the glacier extended to a position downstream of the Waimakariri Gorge, some 15 km beyond the previously mapped LGM ice limit. We use a simple steady-state mass balance model to test the sensitivity of past glacial accumulation to various climatic parameters, and to evaluate possible climate scenarios capable of generating the ice volume required to reach the full local-LGM extent. Model outcomes indicate that under New Zealand's oceanic setting, a cooling of 5 °C, assuming modern precipitation levels, or a cooling of 6.5 °C, assuming a one third reduction in precipitation, would suffice to drive the Waimakariri glacier to the eastern alpine forelands (Canterbury Plains). Our findings demonstrate that the scale of LGM glaciation in the Waimakariri Valley and adjacent major catchments, both in terms of ice volume and downvalley ice extent, has been significantly underestimated. Our observation that high-lying glacial surfaces, so far believed to represent much older glacial episodes, were glaciated during the LGM

  15. Are mesophotic coral ecosystems distinct communities and can they serve as refugia for shallow reefs?

    NASA Astrophysics Data System (ADS)

    Semmler, Robert F.; Hoot, Whitney C.; Reaka, Marjorie L.

    2017-06-01

    We analyzed an extensive dataset of over 9000 benthic and suprabenthic species found throughout the Gulf of Mexico (GoMx) to assess whether mesophotic coral ecosystems represent distinct assemblages and evaluate their potential to serve as refugia for shallow reef communities. We assessed community structure of the overall benthic community from 0 to 300 m via non-metric multidimensional scaling (NMDS) of species presence across depth bands. We used the Jaccard index of similarity to calculate the proportion of shared species between adjacent depth bands, measure species turnover with depth, and assess taxonomic overlap between shallow reefs versus progressively deeper depth bands. NMDS ordinations showed that the traditionally defined mesophotic range (30-150 m) as a whole is not a distinct community. In contrast, taxonomically distinct communities, determined by hierarchical clustering, were found at 0-70, 60-120, 110-200, and 190-300 m. Clustering highlighted an important separation in the benthic community at 60 m, which was especially important for actinopterygian fishes. Species turnover between adjacent depths decreased with depth for all taxa combined and individual taxa, with peaks at 60, 90-120, and 190-200 m. Fishes showed lower turnover from shallow to upper mesophotic depths (0-50 m) than all taxa combined, a substantial peak at 60 m, followed by a precipitous and continued decline in turnover thereafter. Taxonomic overlap between shallow (0-20 m) and progressively deeper zones declined steadily with depth in all taxa and individual taxa, suggesting that mid- and lower mesophotic habitats have less (but not inconsequential) potential to serve as refugia (60-150 m, 15-25% overlap with shallow habitats) than upper mesophotic zones (30-60 m, 30-45% overlap with shallow habitats) for all taxa combined. We conclude that the traditional mesophotic zone is home to three ecological communities in the GoMx, one that is confluent with shallow reefs, a distinct

  16. Trends in Irish-Medium Education in the Republic of Ireland and Northern Ireland since 1920: Shifting Agents and Explanations

    ERIC Educational Resources Information Center

    McAdory, Sara E.; Janmaat, Jan Germen

    2015-01-01

    Some recent studies have suggested a significant bottom-up or parental component to recent movements for autochthonous minority language-medium education (MLME). This study takes MLME as the outcome of interest and seeks to explain trends in Irish-medium education (IME) in the Republic of Ireland and Northern Ireland since 1920--a unique…

  17. Relationships between Attitudes to Irish, Social Class, Religion and National Identity in the Republic of Ireland and Northern Ireland

    ERIC Educational Resources Information Center

    Riagain, Padraig O.

    2007-01-01

    Research on language attitudes in the Republic of Ireland has been greatly influenced by stratification theories. That is to say, differences in attitudes are seen to reflect the positions individuals occupy in the social structure. Research on language attitudes in Northern Ireland is less developed, but has tended to view such attitudes as…

  18. Implementing Major Trauma Audit in Ireland.

    PubMed

    Deasy, Conor; Cronin, Marina; Cahill, Fiona; Geary, Una; Houlihan, Patricia; Woodford, Maralyn; Lecky, Fiona; Mealy, Ken; Crowley, Philip

    2016-01-01

    There are 27 receiving trauma hospitals in the Republic of Ireland. There has not been an audit system in place to monitor and measure processes and outcomes of care. The National Office of Clinical Audit (NOCA) is now working to implement Major Trauma Audit (MTA) in Ireland using the well-established National Health Service (NHS) UK Trauma Audit and Research Network (TARN). The aim of this report is to highlight the implementation process of MTA in Ireland to raise awareness of MTA nationally and share lessons that may be of value to other health systems undertaking the development of MTA. The National Trauma Audit Committee of the Royal College of Surgeons in Ireland, consisting of champions and stakeholders in trauma care, in 2010 advised on the adaptation of TARN for Ireland. In 2012, the Emergency Medicine Program endorsed TARN and in setting up the National Emergency Medicine Audit chose MTA as the first audit project. A major trauma governance group was established representing stakeholders in trauma care, a national project co-ordinator was recruited and a clinical lead nominated. Using Survey Monkey, the chief executives of all trauma receiving hospitals were asked to identify their hospital's trauma governance committee, trauma clinical lead and their local trauma data co-ordinator. Hospital Inpatient Enquiry systems were used to identify to hospitals an estimate of their anticipated trauma audit workload. There are 25 of 27 hospitals now collecting data using the TARN trauma audit platform. These hospitals have provided MTA Clinical Leads, allocated data co-ordinators and incorporated MTA reports formally into their clinical governance, quality and safety committee meetings. There has been broad acceptance of the NOCA escalation policy by hospitals in appreciation of the necessity for unexpected audit findings to stimulate action. Major trauma audit measures trauma patient care processes and outcomes of care to drive quality improvement at hospital and

  19. Critical insolation-CO2 relation for diagnosing past and future glacial inception.

    PubMed

    Ganopolski, A; Winkelmann, R; Schellnhuber, H J

    2016-01-14

    The past rapid growth of Northern Hemisphere continental ice sheets, which terminated warm and stable climate periods, is generally attributed to reduced summer insolation in boreal latitudes. Yet such summer insolation is near to its minimum at present, and there are no signs of a new ice age. This challenges our understanding of the mechanisms driving glacial cycles and our ability to predict the next glacial inception. Here we propose a critical functional relationship between boreal summer insolation and global carbon dioxide (CO2) concentration, which explains the beginning of the past eight glacial cycles and might anticipate future periods of glacial inception. Using an ensemble of simulations generated by an Earth system model of intermediate complexity constrained by palaeoclimatic data, we suggest that glacial inception was narrowly missed before the beginning of the Industrial Revolution. The missed inception can be accounted for by the combined effect of relatively high late-Holocene CO2 concentrations and the low orbital eccentricity of the Earth. Additionally, our analysis suggests that even in the absence of human perturbations no substantial build-up of ice sheets would occur within the next several thousand years and that the current interglacial would probably last for another 50,000 years. However, moderate anthropogenic cumulative CO2 emissions of 1,000 to 1,500 gigatonnes of carbon will postpone the next glacial inception by at least 100,000 years. Our simulations demonstrate that under natural conditions alone the Earth system would be expected to remain in the present delicately balanced interglacial climate state, steering clear of both large-scale glaciation of the Northern Hemisphere and its complete deglaciation, for an unusually long time.

  20. Deglaciation of the James Bay Lowlands and Northern Abitibi: Insights on Late-Glacial Ice Readvances and Drainage of Glacial Lake Ojibway

    NASA Astrophysics Data System (ADS)

    Roy, M.; Veillette, J. J.; Dell'Oste, F.

    2008-12-01

    Deglaciation in the James Bay region was marked by the scission of the Laurentide ice sheet margin into the Hudson dome to the west and the New-Quebec dome to the east, which subsequently retreated northward, in contact with the waters of glacial Lake Ojibway. Previous work based on air photo-interpretation and field observations indicate that ice retreat in the region was highly dynamic, with the occurrence of at least three ice readvances into the basin of Lake Objiway prior to the final deglaciation, and the incursion of the post- glacial Tyrrell Sea at ~8 ka (Hardy, 1976). Our investigations of stratigraphic sections exposed along the Harricana, Nottaway, Broadback, and Rupert rivers in the lowlands of Quebec indicate that only part of these events are preserved in these sedimentary sequences. The base of the late-glacial sequence generally consists of a carbonate-bearing clayey readvance till that lies on older tills of the last glacial cycle, or truncate Lake Ojibway glaciolacustrine sediments. None of the sections showed more than one till of the three (Cochrane I, Rupert, Cochrane II) readvances documented in the region. Nonetheless, an extensive Ojibway sequence located just south from the lowlands shows three intervals with significant increases in detrital carbonate and coarsening of the varve sequence that can be linked with these late-glacial surges. In the lowlands, the readvance till is commonly capped by a thick sequence of Ojibway varves. The contact between the glaciolacustrine sediments and the overlying Tyrrell Sea marine deposits is marked by a ~50 cm-thick horizon composed at the bottom of thinly laminated reddish and grey silt beds containing abundant rounded clay balls, overlain by coarser silts and fine sands with disseminated clasts. This horizon is here interpreted to reflect the abrupt drainage of Lake Ojibway. Recent radiocarbon dating of mollusks and foraminifers from the uppermost part of this horizon yielded ages of ~7.7 ka and ~8

  1. The status of Phytophthora ramorum in Ireland

    Treesearch

    Carmel O?Connor; Elizabeth Gosling

    2008-01-01

    This paper reports on the first 2 years of data collected to study the ecology of Phytophthora ramorum in Ireland. Since spring 2005, sampling has been carried out for the presence of the pathogen in soil and watercourses from 11 susceptible forest sites in Ireland, using a rapid DNA method in conjunction with morphological identification methods....

  2. Glacial morphology and depositional sequences of the Antarctic Continental Shelf

    USGS Publications Warehouse

    ten Brink, Uri S.; Schneider, Christopher

    1995-01-01

    Proposes a simple model for the unusual depositional sequences and morphology of the Antarctic continental shelf. It considers the regional stratal geometry and the reversed morphology to be principally the results of time-integrated effects of glacial erosion and sedimentation related to the location of the ice grounding line. The model offers several guidelines for stratigraphic interpretation of the Antarctic shelf and a Northern Hemisphere shelf, both of which were subject to many glacial advances and retreats. -Authors

  3. Differentiating Hydrothermal, Pedogenic, and Glacial Weathering in a Cold Volcanic Mars-Analog Environment

    NASA Technical Reports Server (NTRS)

    Scudder, N. A.; Horgan, B.; Havig, J.; Rutledge, A.; Rampe, E. B.; Hamilton, T.

    2016-01-01

    Although the current cold, dry environment of Mars extends back through much of its history, its earliest periods experienced significant water- related surface activity. Both geomorphic features (e.g., paleolakes, deltas, and river valleys) and hydrous mineral detections (e.g., clays and salts) have historically been interpreted to imply a "warm and wet" early Mars climate. More recently, atmospheric modeling studies have struggled to produce early climate conditions with temperatures above 0degC, leading some studies to propose a "cold and icy" early Mars dominated by widespread glaciation with transient melting. However, the alteration mineralogy produced in subglacial environments is not well understood, so the extent to which cold climate glacial weathering can produce the diverse alteration mineralogy observed on Mars is unknown. This summer, we will be conducting a field campaign in a glacial weathering environment in the Cascade Range, OR in order to determine the types of minerals that these environments produce. However, we must first disentangle the effects of glacial weathering from other significant alteration processes. Here we attempt a first understanding of glacial weathering by differentiating rocks and sediments weathered by hydrothermal, pedogenic, and glacial weathering processes in the Cascades volcanic range.

  4. Evidence of strong ocean heating during glacial periods

    NASA Astrophysics Data System (ADS)

    Zimov, S. A.; Zimov, N.

    2013-12-01

    Numerous hypotheses have addressed glacial-interglacial climatic dynamics, but none of them explain the sharp 25C temperature increase in Greenland in the last deglaciation (Cuffey et al. 1995; Dahl-Jensen et al. 1998). These robust data were obtained through analyzing the temperature profile in the Greenland ice sheet where cold from the last glaciation is preserved in the depth of the glacial sheet. We suggest that during glaciations the ocean accumulated energy: interior ocean water heated up to ~20-30C and during deglaciation this energy is released. In the analogy with reconstructing the ice sheet temperature profiles, the most reliable proof of ocean interior warming during the last glaciation is the heat flux profiles in the bottom sediments. In the final reports based on temperature measurements conducted during the DSDP (Deep Sea Drilling Project) it is stated that heat flux in the bottom sediments doesn't vary with depth and consequently there were no substantial temperature changes in the ocean interior during the last glacial cycle, and heat flux on the surface of the ocean bottom is the geothermal heat flux (Erickson et al., 1975, Hyndman et al., 1987). However, we have critically investigated data in all initial reports of all deep sea drilling projects and have noticed that all temperature data show that heat flow decreases strongly with depth (a minimum of 40 mW/m2), i.e. most of the heat flux detected on the surface of the ocean floor is not the geothermal heat flux but remaining heat that bottom sediments release. Sharp shifts in heat flow are seen within boreholes at depths crossing gas hydrate bottom. All this means that during the last glacial period interior water temperature was on 25-30C degrees warmer. Conversely, in isolated seas heat flow in the sediments shows little change with depth.

  5. Glacial History of Southernmost South America and Implications for Movement of the Westerlies and Antarctic Frontal Zone

    NASA Astrophysics Data System (ADS)

    Kaplan, M. R.; Fogwill, C. J.; Hulton, N. R.; Sugden, D. E.; Peter, K. W.

    2004-12-01

    The ~1 Myr glacial geologic record in southern South American is one of the few available terrestrial paleoclimate proxies at orbital and suborbital time scales in the middle latitudes of the Southern Hemisphere. Presently, southernmost Patagonia lies about 3\\deg north of the Antarctic frontal zone and within the middle latitude westerlies and the climate is controlled by the surrounding maritime conditions. Thus, the long-term glacial record provides insight into the history of climatic boundaries over the middle and high latitude southern ocean, including the upwind SE Pacific Ocean, tectonic-glacial evolution of the Andes, and global climate. To date, cosmogenic nuclide and 14C dating have focused on glacial fluctuations between 51 and 53\\deg S (Torres del Paine to northern Tierra del Fuego) during the last glacial cycle, including the late glacial period. At least 4 advances occurred between ca. 25 and 17 ka, with the maximum expansion of ice ca. 25-24 ka. Major deglaciation commenced after ca. 17.5 ka, which was interrupted by a major glacial-climate event ca. 14-12 ka. Modelling experiments suggest that the ice mass needed to form the glacial maximum moraines required about a 6\\deg cooling and a slight drying relative to the present. Such a fundamental temperature reduction, despite high summer isolation, strongly suggests northward movement of the westerlies and the polar front on millennial timescales. The Patagonian record also indicates that on orbital timescales equatorward movement of climate boundaries and glacial growth was in phase with major Northern Hemisphere ice volume change, despite high local summer insolation. At suborbital timescales, the picture is more complex. While major facets of the last glacial maximum appear to be in phase between the hemispheres, at least some late glacial events may be in step with Antarctic climate change. Present and future research will further constrain the timing of glacial events over the last 1 Myr and

  6. Towards an improved inventory of Glacial Lake Outburst Floods in the Himalayas

    NASA Astrophysics Data System (ADS)

    Veh, Georg; Walz, Ariane; Korup, Oliver; Roessner, Sigrid

    2016-04-01

    The retreat of glaciers in the Himalayas and the associated release of meltwater have prompted the formation and growth of thousands of glacial lakes in the last decades. More than 2,200 of these lakes have developed in unconsolidated moraine material. These lakes can drain in a single event, producing potentially destructive glacial lake outburst floods (GLOFs). Only 44 GLOFs in the Himalayas have been documented in more detail since the 1930s, and evidence for a change, let alone an increase, in the frequency of these flood events remains elusive. The rare occurrence of GLOFs is counterintuitive to our hypothesis that an increasing amount of glacial lakes has to be consistent with a rising amount of outburst floods. Censoring bias affects the GLOF record, such that mostly larger floods with commensurate impact have been registered. Existing glacial lake inventories are also of limited help for the identification of GLOFs, as they were created in irregular time steps using different methodological approach and covering different regional extents. We discuss the key requirements for generating a more continuous, close to yearly time series of glacial lake evolution for the Himalayan mountain range using remote sensing data. To this end, we use sudden changes in glacial lake areas as the key diagnostic of dam breaks and outburst floods, employing the full archive of cloud-free Landsat data (L5, L7 and L8) from 1988 to 2015. SRTM and ALOS World 3D topographic data further improve the automatic detection of glacial lakes in an alpine landscape that is often difficult to access otherwise. Our workflow comprises expert-based classification of water bodies using thresholds and masks from different spectral indices and band ratios. A first evaluation of our mapping approach suggests that GLOFs reported during the study period could be tracked independently by a significant reduction of lake size between two subsequent Landsat scenes. This finding supports the feasibility

  7. Early warning method of Glacial Lake Outburst Floods based on temperature and rainfall

    NASA Astrophysics Data System (ADS)

    Liu, Jingjing; Su, Pengcheng; Cheng, Zunlan

    2017-04-01

    Glacial lake outburst floods (GLOFs) are serious disasters in glacial areas. At present, glaciers are retreating while glacial lake area and the outburst risk increases due to the global warming. Therefore, the research of early warning method of GLOFs is important to prevent and reduce the disasters. This paper provides an early warning method using the temperature and rainfall as indices. The daily growth rate of positive antecedent accumulative temperature and the antecedent thirty days accumulative precipitation are calculated for 21 events of GLOF before 2010, based on data from the 21 meteorological stations nearby. The result shows that all the events are above the curve, TV = -0.0193RDC + 3.0018, which can be taken as the early warning threshold curve. This has been verified by the GLOF events in the Ranzeaco glacial lake on 2013-07-05.

  8. Glacial Age Correlations and Pedogenesis Rates at Long Valley, Costilla Masif, Northern New Mexico

    NASA Astrophysics Data System (ADS)

    Feldman, A. D.

    2017-12-01

    New Mexico represents the southernmost extent of glacial activity in the United States. As such an enhanced understanding of glacial climate cycles in the region as expressed through the relict landscapes they leave behind can enhance our understanding of the evolution of high altitude landscapes and soils throughout the Quaternary period. The Sangre De Cristo mountain range in northern New Mexico exhibits some of the southernmost expansion of glacial activity in the Southwest during the Quaternary; yet the range has had only limited correlation of its glacial chronology performed to date. In this study a detailed investigation into soil pedogenesis on relict moraine features is used to fit the Long Valley glacial sequence extending eastward from the Costilla Masif into the established Rocky Mountain glacial chronology. Analyzed soil development characteristics are particle size, organic carbon, and iron oxide distributions including total iron, ferric iron, ferrous iron, citrate dithionite, hydroxylamine for amorphous ferrihydrite, and pyrophosphate for organically bound iron. In addition, soils developement will be analyzed in situ for computation of a modified Harden soil profile development index. A secondary purpose of the study is to establish better constraints on the rates of soil pedogenesis in these high altitude glacial features. Soil profile developement and pedogenesis rates will be compared with previously published data from areas both further south in the Sangre De Cristo's as well as throughout the more northern sections of the Rocky Mountains to correlate moraine ages as well as to constrain how the particular climate of the Long Valley has affected soil development during the Quaternary.

  9. Glacial loess or shoreface sands: a re-interpretation of the Upper Ordovician (Ashgillian) glacial Ammar Formation, Southern Jordan

    NASA Astrophysics Data System (ADS)

    Turner, B. R.; Makhlouf, I. M.; Armstrong, H. A.

    2003-04-01

    Upper Ordovician (Ashgillian) glacial deposits of the Ammar Formation, Southern Jordan, comprise locally deformed, structureless fine sandstone, incised by glacial channels filled by braided outwash plain sandstones and transgressive marine mudstones. The structureless sandstones, previously interpreted as a glacial rock flour or loessite derived from the underlying undisturbed sandstones, differ significantly from typical loessite and contain hitherto unrecognised sedimentary structures, including hummocky cross-stratification. The sandstones, which grade laterally and vertically into stratigraphically equivalent undeformed marginal marine sandstones, are interpreted as a deformed facies of the underlying sandstones, deposited in a similar high energy shoreface environment. Although deformation of the shoreface sandstones was post-depositional, the origin of the deformation, and its confinement to the Jebel Ammar area is unknown. Deformation due to the weight of the overlying ice is unlikely as the glaciofluvial channels are now thought to have been cut by tunnel valley activity not ice. A more likely mechanism is post-glacial crustal tectonics. Melting of ice caps is commonly associated with intraplate seismicity and the development of an extensional crustal stress regime around the perimeter of ice caps; the interior is largely aseismic because the weight of the ice supresses seismic activity and faulting. Since southern Jordan lay close to the ice cap in Saudi Arabia it may have been subjected to postglacial seismicity and crustal stress, which induced ground shaking, reduced overburden pressure, increased hydrostatic pressure and possibly reactivation of existing tectonic faults. This resulted in liquefaction and extensive deformation of the sediments, which show many characteristics of seismites, generated by earthquake shocks. Since the glaciation was a very short-lived event (0.2-1 Ma), deglaciation and associated tectonism triggering deformation, lasted

  10. Glacial weathering, sulfide oxidation, and global carbon cycle feedbacks.

    PubMed

    Torres, Mark A; Moosdorf, Nils; Hartmann, Jens; Adkins, Jess F; West, A Joshua

    2017-08-15

    Connections between glaciation, chemical weathering, and the global carbon cycle could steer the evolution of global climate over geologic time, but even the directionality of feedbacks in this system remain to be resolved. Here, we assemble a compilation of hydrochemical data from glacierized catchments, use this data to evaluate the dominant chemical reactions associated with glacial weathering, and explore the implications for long-term geochemical cycles. Weathering yields from catchments in our compilation are higher than the global average, which results, in part, from higher runoff in glaciated catchments. Our analysis supports the theory that glacial weathering is characterized predominantly by weathering of trace sulfide and carbonate minerals. To evaluate the effects of glacial weathering on atmospheric pCO 2 , we use a solute mixing model to predict the ratio of alkalinity to dissolved inorganic carbon (DIC) generated by weathering reactions. Compared with nonglacial weathering, glacial weathering is more likely to yield alkalinity/DIC ratios less than 1, suggesting that enhanced sulfide oxidation as a result of glaciation may act as a source of CO 2 to the atmosphere. Back-of-the-envelope calculations indicate that oxidative fluxes could change ocean-atmosphere CO 2 equilibrium by 25 ppm or more over 10 ky. Over longer timescales, CO 2 release could act as a negative feedback, limiting progress of glaciation, dependent on lithology and the concentration of atmospheric O 2 Future work on glaciation-weathering-carbon cycle feedbacks should consider weathering of trace sulfide minerals in addition to silicate minerals.

  11. Glacial weathering, sulfide oxidation, and global carbon cycle feedbacks

    PubMed Central

    Torres, Mark A.; Moosdorf, Nils; Hartmann, Jens; Adkins, Jess F.

    2017-01-01

    Connections between glaciation, chemical weathering, and the global carbon cycle could steer the evolution of global climate over geologic time, but even the directionality of feedbacks in this system remain to be resolved. Here, we assemble a compilation of hydrochemical data from glacierized catchments, use this data to evaluate the dominant chemical reactions associated with glacial weathering, and explore the implications for long-term geochemical cycles. Weathering yields from catchments in our compilation are higher than the global average, which results, in part, from higher runoff in glaciated catchments. Our analysis supports the theory that glacial weathering is characterized predominantly by weathering of trace sulfide and carbonate minerals. To evaluate the effects of glacial weathering on atmospheric pCO2, we use a solute mixing model to predict the ratio of alkalinity to dissolved inorganic carbon (DIC) generated by weathering reactions. Compared with nonglacial weathering, glacial weathering is more likely to yield alkalinity/DIC ratios less than 1, suggesting that enhanced sulfide oxidation as a result of glaciation may act as a source of CO2 to the atmosphere. Back-of-the-envelope calculations indicate that oxidative fluxes could change ocean–atmosphere CO2 equilibrium by 25 ppm or more over 10 ky. Over longer timescales, CO2 release could act as a negative feedback, limiting progress of glaciation, dependent on lithology and the concentration of atmospheric O2. Future work on glaciation–weathering–carbon cycle feedbacks should consider weathering of trace sulfide minerals in addition to silicate minerals. PMID:28760954

  12. Terrestrial cosmogenic nuclide surface exposure dating of the oldest glacial successions in the Himalayan orogen: Ladakh Range, northern India

    USGS Publications Warehouse

    Owen, L.A.; Caffee, M.W.; Bovard, K.R.; Finkel, R.C.; Sharma, M.C.

    2006-01-01

    Terrestrial cosmogenic nuclide surface exposure dating of moraine boulders and alluvial fan sediments define the timing of five glacial advances over at least the last five glacial cycles in the Ladakh Range of the Transhimalaya. The glacial stages that have been identified are: the Indus Valley glacial stage, dated at older than 430 ka; the Leh glacial stage occurring in the penultimate glacial cycle or older; the Karglacial stage, occurring during the early part of the last glacial cycle; the Bazgo glacial stage, at its maximum during the middle of the last glacial cycle; and the early Holocene Khalling glacial stage. The exposure ages of the Indus Valley moraines are the oldest observed to date throughout the Himalayan orogen. We observe a pattern of progressively more restricted glaciation during the last five glacial cycles, likely indicating a progressive reduction in the moisture supply necessary to sustain glaciation. A possible explanation is that uplift of Himalayan ranges to the south and/or of the Karakoram Mountains to the west of the region may have effectively blocked moisture supply by the south Asian summer monsoon and mid-latitude westerlies, respectively. Alternatively, this pattern of glaciation may reflect a trend of progressively less extensive glaciation in mountain regions that has been observed globally throughout the Pleistocene. ?? 2006 Geological Society of America.

  13. Microbial genesis, life and death in glacial ice.

    PubMed

    Price, P Buford

    2009-01-01

    Arguments are given that terrestrial RNA and DNA may have originated in a frozen environment more than 4 billion years ago. Scenarios are developed for atmospheric transport of microbes onto glacial ice, their adaptation to subzero temperatures in the ice, and their incorporation into one of three habitats - liquid veins, mineral grain surfaces, or isolated inside 1 of the crystals that make up polycrystalline ice. The Arrhenius dependence of microbial metabolic rate on temperature is shown to match that required to repair damage owing to spontaneous DNA depurination and amino acid racemization. Even for the oldest glacial ice, microbial lifetime is shown not to be shortened by radiation damage from 238U, 232Th, or 40K in mineral dust in ice, by phage-induced lysis, or by penetrating cosmic radiation. Instead, death of those cells adapted to the hostile conditions in glacial ice is probably due to exhaustion of available nutrients. By contrast, in permafrost microbial death is more likely due to alpha-particle radiation damage from U and Th in the soil and rocks intermixed with ice. For residence times in ice longer than a million years, spore formers may be unable to compete in longevity with vegetative cells that are able to repair DNA damage via survival metabolism.

  14. Holocene re-colonisation, central-marginal distribution and habitat specialisation shape population genetic patterns within an Atlantic European grass species.

    PubMed

    Harter, D E V; Jentsch, A; Durka, W

    2015-05-01

    Corynephorus canescens (L.) P.Beauv. is an outbreeding, short-lived and wind-dispersed grass species, highly specialised on scattered and disturbance-dependent habitats of open sandy sites. Its distribution ranges from the Iberian Peninsula over Atlantic regions of Western and Central Europe, but excludes the two other classical European glacial refuge regions on the Apennine and Balkan Peninsulas. To investigate genetic patterns of this uncommon combination of ecological and biogeographic species characteristics, we analysed AFLP variation among 49 populations throughout the European distribution range, expecting (i) patterns of SW European glacial refugia and post-glacial expansion to the NE; (ii) decreasing genetic diversity from central to marginal populations; and (iii) interacting effects of high gene flow and disturbance-driven genetic drift. Decreasing genetic diversity from SW to NE and distinct gene pool clustering imply refugia on the Iberian Peninsula and in western France, from where range expansion originated towards the NE. High genetic diversity within and moderate genetic differentiation among populations, and a significant pattern of isolation-by-distance indicate a gene flow drift equilibrium within C. canescens, probably due to its restriction to scattered and dynamic habitats and limited dispersal distances. These features, as well as the re-colonisation history, were found to affect genetic diversity gradients from central to marginal populations. Our study emphasises the need for including the specific ecology into analyses of species (re-)colonisation histories and range centre-margin analyses. To account for discontinuous distributions, new indices of marginality were tested for their suitability in studies of centre-periphery gradients. © 2014 German Botanical Society and The Royal Botanical Society of the Netherlands.

  15. Model-based analysis supports interglacial refugia over long-dispersal events in the diversification of two South American cactus species

    PubMed Central

    Perez, M F; Bonatelli, I A S; Moraes, E M; Carstens, B C

    2016-01-01

    Pilosocereus machrisii and P. aurisetus are cactus species within the P. aurisetus complex, a group of eight cacti that are restricted to rocky habitats within the Neotropical savannas of eastern South America. Previous studies have suggested that diversification within this complex was driven by distributional fragmentation, isolation leading to allopatric differentiation, and secondary contact among divergent lineages. These events have been associated with Quaternary climatic cycles, leading to the hypothesis that the xerophytic vegetation patches which presently harbor these populations operate as refugia during the current interglacial. However, owing to limitations of the standard phylogeographic approaches used in these studies, this hypothesis was not explicitly tested. Here we use Approximate Bayesian Computation to refine the previous inferences and test the role of different events in the diversification of two species within P. aurisetus group. We used molecular data from chloroplast DNA and simple sequence repeats loci of P. machrisii and P. aurisetus, the two species with broadest distribution in the complex, in order to test if the diversification in each species was driven mostly by vicariance or by long-dispersal events. We found that both species were affected primarily by vicariance, with a refuge model as the most likely scenario for P. aurisetus and a soft vicariance scenario most probable for P. machrisii. These results emphasize the importance of distributional fragmentation in these species, and add support to the hypothesis of long-term isolation in interglacial refugia previously proposed for the P. aurisetus species complex diversification. PMID:27071846

  16. Glacial melt water in Greenland - A renewable resource for the future

    NASA Astrophysics Data System (ADS)

    Alther, G. R.; Ruedisili, L. C.; Stauber, H.; Kollbrunner, C. F.

    1981-06-01

    Glacial melt water in Greenland can be used as a renewable resource for generating electricity (a yearly estimate of 60-115 GW), and it can serve as a supplementary source for drinking and irrigation, metallurgical processing, and the manufacturing of liquid hydrogen as fuel. Southern Greenland is particularly suited for this melt water hydropower project, having high precipitation and summer temperatures, large quantities of melt water, natural 'nunatak' dams, and coastal ranges with steep gradients. Transportation of the generated energy is proposed in the form of sea cables and overland transmission lines, hydrogen gas pipelines, and tankers for liquid hydrogen transport. A hypothetical glacial power station is schematically illustrated, and production costs are calculated. The glacial melt project would serve as an economical source of energy with minimal damage to the environment.

  17. Interrogating medical tourism: Ireland, abortion, and mobility rights.

    PubMed

    Gilmartin, Mary; White, Allen

    2011-01-01

    Medical tourism in Ireland, like in many Western states, is built around assumptions about individual agency, choice, possibility, and mobility. One specific form of medical tourism—the flow of women from Ireland traveling in order to secure an abortion—disrupts and contradicts these assumptions. One legacy of the bitter, contentious political and legal battles surrounding abortion in Ireland in the 1980s and 1990s has been securing the right of mobility for all pregnant Irish citizens to cross international borders to secure an abortion. However, these mobility rights are contingent upon nationality, social class, and race, and they have enabled successive Irish governments to avoid any responsibility for providing safe, legal, and affordable abortion services in Ireland. Nearly twenty years after the X case discussed here, the pregnant female body moving over international borders—entering and leaving the state—is still interpreted as problematic and threatening to the Irish state.

  18. Geochemistry of glacial sediments in the area of the Bend massive sulfide deposit, north-central Wisconsin

    USGS Publications Warehouse

    Woodruff, L.G.; Attig, J.W.; Cannon, W.F.

    2004-01-01

    Geochemical exploration in northern Wisconsin has been problematic because of thick glacial overburden and complex stratigraphic record of glacial history. To assess till geochemical exploration in an area of thick glacial cover and complex stratigraphy samples of glacial materials were collected from cores from five rotasonic boreholes near a known massive sulfide deposit, the Bend deposit in north-central Wisconsin. Diamond drilling in the Bend area has defined a long, thin zone of mineralization at least partly intersected at the bedrock surface beneath 30-40 m of unconsolidated glacial sediments. The bedrock surface has remnant regolith and saprolite resulting from pre-Pleistocene weathering. Massive sulfide and mineralized rock collected from diamond drill core from the deposit contain high (10s to 10,000s ppm) concentrations of Ag, As, Au, Bi, Cu, Hg, Se, Te, and Tl. Geochemical properties of the glacial stratigraphic units helped clarify the sequence and source areas of several glacial ice advances preserved in the section. At least two till sheets are recognized. Over the zone of mineralization, saprolite and preglacial alluvial and lacustrine samples are preserved on the bedrock surface in a paleoriver valley. The overlying till sheet is a gray, silty carbonate till with a source hundreds of kilometers to the northwest of the study area. This gray till is overlain by red, sandy till with a source to the north in Proterozoic rocks of the Lake Superior area. The complex glacial stratigraphy confounds down-ice geochemical till exploration. The presence of remnant saprolite, preglacial sediment, and far-traveled carbonate till minimized glacial erosion of mineralized material. As a result, little evidence of down-ice glacial dispersion of lithologic or mineralogic indicators of Bend massive sulfide mineralization was found in the samples from the rotasonic cores. This study points out the importance of determining glacial stratigraphy and history, and

  19. Weak overturning circulation and increased iron fertilization maximized carbon storage in the glacial ocean

    NASA Astrophysics Data System (ADS)

    Muglia, J.; Skinner, L.; Schmittner, A.

    2017-12-01

    Circulation changes have been suggested to play an important role in the sequestration of atmospheric CO2 in the glacial ocean. However, previous studies have resulted in contradictory results regarding the strength of the Atlantic Meridional Overturning Circulation (AMOC) and three-dimensional, quantitative reconstructions of the glacial ocean constrained by multiple proxies remain lacking. Here we simulate the modern and glacial ocean using a coupled, global, three-dimensional, physical-biogeochemical model constrained simultaneously by d13C, radiocarbon, and d15N to explore the effects of AMOC differences and Southern Ocean iron fertilization on the distributions of these isotopes and ocean carbon storage. We show that d13C and radiocarbon data sparsely sampled at the locations of existing glacial sediment cores can be used to reconstruct the modern AMOC accurately. Applying this method to the glacial ocean we find that a surprisingly weak (6-9 Sv or about half of today's) and shallow AMOC maximizes carbon storage and best reproduces the sediment data. Increasing the atmospheric soluble iron flux in the model's Southern Ocean intensifies export production, carbon storage, and improves agreement with d13C and d15N reconstructions. Our best fitting model is a significant improvement compared with previous studies. It suggests that a weak and shallow AMOC and enhanced iron fertilization conspired to maximize carbon storage in the glacial ocean.

  20. Animal Health Ireland: providing national leadership and coordination of non-regulatory animal health issues in Ireland.

    PubMed

    More, S J; Doherty, M L; Downey, L; McKenzie, K; Devitt, C; O'Flaherty, J

    2011-12-01

    Livestock production plays an important role in the Irish economy. Regulatory animal health issues are the responsibility of government, but until recently there has been no national coordination of non-regulatory animal health issues. This gap has recently been filled with the establishment of Animal Health Ireland (AHI), a not-for-profit, partnership-based organisation providing national leadership and coordination of non-regulatory animal health issues in Ireland. Animal Health Ireland provides benefits to livestock producers and processors by providing the knowledge, education and coordination required to establish effective control strategies, both on-farm and nationally. This paper presents a brief overview of the context for AHI, and of its establishment and initial activities. Non-regulatory animal health issues have been prioritised. A series of work programmes (each focusing on a high-priority issue) have been established. Partnership is critical to success, both for AHI as an organisation and for effective farm-level transfer of knowledge. This model for national leadership and coordination of non-regulatory animal health issues may be of relevance elsewhere.

  1. 9 CFR 93.432 - Cattle from the Republic of Ireland.

    Code of Federal Regulations, 2011 CFR

    2011-01-01

    ... consignor and consignee, and descriptions of the cattle, including breed, ages, markings, and tattoo and... 9 Animals and Animal Products 1 2011-01-01 2011-01-01 false Cattle from the Republic of Ireland... Cattle from the Republic of Ireland. (a) All cattle to be imported from the Republic of Ireland shall be...

  2. 9 CFR 93.432 - Cattle from the Republic of Ireland.

    Code of Federal Regulations, 2014 CFR

    2014-01-01

    ... consignor and consignee, and descriptions of the cattle, including breed, ages, markings, and tattoo and... 9 Animals and Animal Products 1 2014-01-01 2014-01-01 false Cattle from the Republic of Ireland... Cattle from the Republic of Ireland. (a) All cattle to be imported from the Republic of Ireland shall be...

  3. 9 CFR 93.432 - Cattle from the Republic of Ireland.

    Code of Federal Regulations, 2012 CFR

    2012-01-01

    ... consignor and consignee, and descriptions of the cattle, including breed, ages, markings, and tattoo and... 9 Animals and Animal Products 1 2012-01-01 2012-01-01 false Cattle from the Republic of Ireland... Cattle from the Republic of Ireland. (a) All cattle to be imported from the Republic of Ireland shall be...

  4. 9 CFR 93.432 - Cattle from the Republic of Ireland.

    Code of Federal Regulations, 2010 CFR

    2010-01-01

    ... consignor and consignee, and descriptions of the cattle, including breed, ages, markings, and tattoo and... 9 Animals and Animal Products 1 2010-01-01 2010-01-01 false Cattle from the Republic of Ireland... Cattle from the Republic of Ireland. (a) All cattle to be imported from the Republic of Ireland shall be...

  5. 9 CFR 93.432 - Cattle from the Republic of Ireland.

    Code of Federal Regulations, 2013 CFR

    2013-01-01

    ... consignor and consignee, and descriptions of the cattle, including breed, ages, markings, and tattoo and... 9 Animals and Animal Products 1 2013-01-01 2013-01-01 false Cattle from the Republic of Ireland... Cattle from the Republic of Ireland. (a) All cattle to be imported from the Republic of Ireland shall be...

  6. A profile of physiotherapy supply in Ireland.

    PubMed

    Eighan, James; Walsh, Brendan; Smith, Samantha; Wren, Maev-Ann; Barron, Steve; Morgenroth, Edgar

    2018-04-13

    The lack of information on public and private physiotherapy supply in Ireland makes current and future resource allocation decisions difficult. This paper estimates the supply of physiotherapists in Ireland and profiles physiotherapists across acute and non-acute sectors, and across public and private practice. It examines geographic variation in physiotherapist supply, examining the implications of controlling for healthcare need. Physiotherapist headcounts are estimated using Health Service Personnel Census (HSPC) and Irish Society of Chartered Physiotherapists (ISCP) Register data. Headcounts are converted to whole-time equivalents (WTEs) using the HSPC and a survey of ISCP members to account for full- and part-time working practices. Non-acute supply per 10,000 population in each county is estimated to examine geographic inequalities and the raw population is adjusted in turn for a range of need indicators. An estimated 3172 physiotherapists were practising in Ireland in 2015; 6.8 physiotherapists per 10,000, providing an estimated 2620 WTEs. Females accounted for 74% of supply. Supply was greater in the non-acute sector; 1774 WTEs versus 846 WTEs in the acute sector. Physiotherapists in the acute sector were located mainly in publicly financed institutions (89%) with an even public/private split observed in the non-acute sector. Non-acute physiotherapist supply is unequally distributed across Ireland (Gini coefficient = 0.12; 95% CI 0.08-0.15), and inequalities remain after controlling for variations in healthcare needs across counties. The supply of physiotherapists in Ireland is 30% lower than the EU-28 average. Substantial inequality in the distribution of physiotherapists across counties is observed.

  7. Phylogeographic structure in long-tailed voles (Rodentia: Arvicolinae) belies the complex Pleistocene history of isolation, divergence, and recolonization of Northwest North America's fauna.

    PubMed

    Sawyer, Yadéeh E; Cook, Joseph A

    2016-09-01

    Quaternary climate fluctuations restructured biodiversity across North American high latitudes through repeated episodes of range contraction, population isolation and divergence, and subsequent expansion. Identifying how species responded to changing environmental conditions not only allows us to explore the mode and tempo of evolution in northern taxa, but also provides a basis for forecasting future biotic response across the highly variable topography of western North America. Using a multilocus approach under a Bayesian coalescent framework, we investigated the phylogeography of a wide-ranging mammal, the long-tailed vole, Microtus longicaudus . We focused on populations along the North Pacific Coast to refine our understanding of diversification by exploring the potentially compounding roles of multiple glacial refugia and more recent fragmentation of an extensive coastal archipelago. Through a combination of genetic data and species distribution models (SDMs), we found that historical climate variability influenced contemporary genetic structure, with multiple isolated locations of persistence (refugia) producing multiple divergent lineages (Beringian or northern, southeast Alaska or coastal, and southern or continental) during glacial advances. These vole lineages all occur along the North Pacific Coast where the confluence of numerous independent lineages in other species has produced overlapping zones of secondary contact, collectively a suture zone. Finally, we detected high levels of neoendemism due to complex island geography that developed in the last 10,000 years with the rising sea levels of the Holocene.

  8. Rethinking the history of common walnut (Juglans regia L.) in Europe: Its origins and human interactions.

    PubMed

    Pollegioni, Paola; Woeste, Keith; Chiocchini, Francesca; Del Lungo, Stefano; Ciolfi, Marco; Olimpieri, Irene; Tortolano, Virginia; Clark, Jo; Hemery, Gabriel E; Mapelli, Sergio; Malvolti, Maria Emilia

    2017-01-01

    Common walnut (Juglans regia L) is an economically important species cultivated worldwide for its high-quality wood and nuts. It is generally accepted that after the last glaciation J. regia survived and grew in almost completely isolated stands in Asia, and that ancient humans dispersed walnuts across Asia and into new habitats via trade and cultural expansion. The history of walnut in Europe is a matter of debate, however. In this study, we estimated the genetic diversity and structure of 91 Eurasian walnut populations using 14 neutral microsatellites. By integrating fossil pollen, cultural, and historical data with population genetics, and approximate Bayesian analysis, we reconstructed the demographic history of walnut and its routes of dispersal across Europe. The genetic data confirmed the presence of walnut in glacial refugia in the Balkans and western Europe. We conclude that human-mediated admixture between Anatolian and Balkan walnut germplasm started in the Early Bronze Age, and between western Europe and the Balkans in eastern Europe during the Roman Empire. A population size expansion and subsequent decline in northeastern and western Europe was detected in the last five centuries. The actual distribution of walnut in Europe resulted from the combined effects of expansion/contraction from multiple refugia after the Last Glacial Maximum and its human exploitation over the last 5,000 years.

  9. Rethinking the history of common walnut (Juglans regia L.) in Europe: Its origins and human interactions

    PubMed Central

    Pollegioni, Paola; Woeste, Keith; Chiocchini, Francesca; Del Lungo, Stefano; Ciolfi, Marco; Olimpieri, Irene; Tortolano, Virginia; Clark, Jo; Hemery, Gabriel E.; Mapelli, Sergio; Malvolti, Maria Emilia

    2017-01-01

    Common walnut (Juglans regia L) is an economically important species cultivated worldwide for its high-quality wood and nuts. It is generally accepted that after the last glaciation J. regia survived and grew in almost completely isolated stands in Asia, and that ancient humans dispersed walnuts across Asia and into new habitats via trade and cultural expansion. The history of walnut in Europe is a matter of debate, however. In this study, we estimated the genetic diversity and structure of 91 Eurasian walnut populations using 14 neutral microsatellites. By integrating fossil pollen, cultural, and historical data with population genetics, and approximate Bayesian analysis, we reconstructed the demographic history of walnut and its routes of dispersal across Europe. The genetic data confirmed the presence of walnut in glacial refugia in the Balkans and western Europe. We conclude that human-mediated admixture between Anatolian and Balkan walnut germplasm started in the Early Bronze Age, and between western Europe and the Balkans in eastern Europe during the Roman Empire. A population size expansion and subsequent decline in northeastern and western Europe was detected in the last five centuries. The actual distribution of walnut in Europe resulted from the combined effects of expansion/contraction from multiple refugia after the Last Glacial Maximum and its human exploitation over the last 5,000 years. PMID:28257470

  10. The Distribution and Magnitude of Glacial Erosion on 103-year Timescales at Engabreen, Norway

    NASA Astrophysics Data System (ADS)

    Rand, C.; Goehring, B. M.

    2017-12-01

    We derive the magnitudes of glacial erosion integrated over 103-year timescales across a transect transverse to the direction of ice flow at Engabreen, Norway. Understanding the distribution of glacial erosion is important for several reasons, including sediment budgeting to fjord environments, development of robust landscape evolution models, and if a better understanding between erosion and ice-bed interface properties (e.g., sliding rate, basal water pressure) can be developed, we can use records of glacial erosion to infer glaciological properties that can ultimately benefit models of past and future glaciers. With few exceptions, measurements of glacial erosion are limited to the historical past and even then are rare owing to the difficulty of accessing the glacier bed. One method proven useful in estimating glacial erosion on 103-year timescales is to measure the remaining concentrations of cosmogenic nuclides that accumulate in exposed bedrock during periods of retracted glacier extent and are removed by glacial erosion and radioactive decay during ice cover. Here we will present measurements of 14C and 10Be measured in proglacial bedrock from Engabreen. Our transects are ca. 600 and 400 meters in front of the modern ice front, and based on historical imagery, was ice covered until the recent past. Initial 10Be results show an increase in concentrations of nearly an order of magnitude from the samples near the center of the glacial trough to those on the lateral margin, consistent with conceptual models of glacial erosion parameterized in terms of sliding velocity. Naïve exposure ages that assume no subglacial erosion range from 0.22 - 9.04 ka. More importantly, we can estimate erosion depths by assuming zero erosion of the highest concentration sample along the two transects and calculate the amount of material removed to yield the lower concentrations elsewhere along the two transects. Results indicate minimum erosion depths of 1-183 cm for most ice

  11. Glacial heritage: knowledge, inventory and promotion in the Chablais area (France, Switzerland)

    NASA Astrophysics Data System (ADS)

    Perret, A.; Reynard, E.; Delannoy, J.-J.

    2012-04-01

    This study is part of an Interreg IVA project (www.123chablais.com) dealing with the promotion of different types of natural and cultural heritage in the Chablais area (French and Swiss Prealps) and is linked to the candidature of the French Chablais territory for the European Geoparks Network. The objective of the study is to develop a strategy for the promotion of the glacial heritage (landforms, deposits) in an area where the geomorphological features are highly influenced by glacial history and where key concepts in the Quaternary sciences were developed (e.g. the theory of multiple glaciations by Morlot in 1859), but that is now nearly completely deglaciated. The challenge is to find solutions to explain why the glacial heritage is so important for the regional economy and how it influences the life of inhabitants (e.g. Evian and Thonon mineral water, extraction industry, landscape and tourism), even if glaciers are not so impressive than in other parts of the Alps. The research is divided in three parts. (1) The first one aims to enhance knowledge on glacial landforms and deposits. The study area, that is quite large, has been intensively studied for more than two centuries; nevertheless, some parts have been only poorly studied. Intensive field survey was carried out to fill in the gaps of knowledge and some landforms, such as erratic boulders, have been dated in order to establish a chronology of deglaciation. All of these different elements have been included in a Geographic Information System with the aim of establishing maps of glacial stages in the Chablais area. (2) From this, an inventory of glacial geosites has been carried out, using the assessment method developed by Reynard et al. (2007). A specific focus has been on the assessment of the potential of the selected sites for educational purposes and geotourist promotion. (3) The last part has been the preparation of adapted educational and promotional supports. In particular, an exhibition will be

  12. Extraterrestrial accretion and glacial cycles

    NASA Technical Reports Server (NTRS)

    Muller, R. A.

    1994-01-01

    We propose that the approx. 100-k.y. cycle seen in terrestrial glaciation is due to changes in meteor flux that come from changes in the Earth's orbit. This model can explain a 70-k.y. 'anomalous' period in climate data and the apparent discrepancy between present extraterrestrial fluxes and those in oceanic sediments. It can be tested by measuring Ir densities in sediments and ice during glacials and interglacials.

  13. Population dynamic of the extinct European aurochs: genetic evidence of a north-south differentiation pattern and no evidence of post-glacial expansion.

    PubMed

    Mona, Stefano; Catalano, Giulio; Lari, Martina; Larson, Greger; Boscato, Paolo; Casoli, Antonella; Sineo, Luca; Di Patti, Carolina; Pecchioli, Elena; Caramelli, David; Bertorelle, Giorgio

    2010-03-26

    The aurochs (Bos primigenius) was a large bovine that ranged over almost the entirety of the Eurasian continent and North Africa. It is the wild ancestor of the modern cattle (Bos taurus), and went extinct in 1627 probably as a consequence of human hunting and the progressive reduction of its habitat. To investigate in detail the genetic history of this species and to compare the population dynamics in different European areas, we analysed Bos primigenius remains from various sites across Italy. Fourteen samples provided ancient DNA fragments from the mitochondrial hypervariable region. Our data, jointly analysed with previously published sequences, support the view that Italian aurochsen were genetically similar to modern bovine breeds, but very different from northern/central European aurochsen. Bayesian analyses and coalescent simulations indicate that the genetic variation pattern in both Italian and northern/central European aurochsen is compatible with demographic stability after the last glaciation. We provide evidence that signatures of population expansion can erroneously arise in stable aurochsen populations when the different ages of the samples are not taken into account. Distinct groups of aurochsen probably inhabited Italy and northern/central Europe after the last glaciation, respectively. On the contrary, Italian and Fertile Crescent aurochsen likely shared several mtDNA sequences, now common in modern breeds. We argue that a certain level of genetic homogeneity characterized aurochs populations in Southern Europe and the Middle East, and also that post-glacial recolonization of northern and central Europe advanced, without major demographic expansions, from eastern, and not southern, refugia.

  14. Population dynamic of the extinct European aurochs: genetic evidence of a north-south differentiation pattern and no evidence of post-glacial expansion

    PubMed Central

    2010-01-01

    Background The aurochs (Bos primigenius) was a large bovine that ranged over almost the entirety of the Eurasian continent and North Africa. It is the wild ancestor of the modern cattle (Bos taurus), and went extinct in 1627 probably as a consequence of human hunting and the progressive reduction of its habitat. To investigate in detail the genetic history of this species and to compare the population dynamics in different European areas, we analysed Bos primigenius remains from various sites across Italy. Results Fourteen samples provided ancient DNA fragments from the mitochondrial hypervariable region. Our data, jointly analysed with previously published sequences, support the view that Italian aurochsen were genetically similar to modern bovine breeds, but very different from northern/central European aurochsen. Bayesian analyses and coalescent simulations indicate that the genetic variation pattern in both Italian and northern/central European aurochsen is compatible with demographic stability after the last glaciation. We provide evidence that signatures of population expansion can erroneously arise in stable aurochsen populations when the different ages of the samples are not taken into account. Conclusions Distinct groups of aurochsen probably inhabited Italy and northern/central Europe after the last glaciation, respectively. On the contrary, Italian and Fertile Crescent aurochsen likely shared several mtDNA sequences, now common in modern breeds. We argue that a certain level of genetic homogeneity characterized aurochs populations in Southern Europe and the Middle East, and also that post-glacial recolonization of northern and central Europe advanced, without major demographic expansions, from eastern, and not southern, refugia. PMID:20346116

  15. New OBS network deployment offshore Ireland

    NASA Astrophysics Data System (ADS)

    Le Pape, Florian; Bean, Chris; Craig, David; Jousset, Philippe; Horan, Clare; Hogg, Colin; Donne, Sarah; McCann, Hannah; Möllhoff, Martin; Kirk, Henning; Ploetz, Aline

    2016-04-01

    With the presence of the stormy NE Atlantic, Ireland is ideally located to investigate further our understanding of ocean generated microseisms and use noise correlation methods to develop seismic imaging in marine environments as well as time-lapse monitoring. In order to study the microseismic activity offshore Ireland, 10 Broad Band Ocean Bottom Seismographs (OBSs) units including hydrophones have been deployed in January 2016 across the shelf offshore Donegal and out into the Rockall Trough. This survey represents the first Broadband passive study in this part of the NE Atlantic. The instruments will be recovered in August 2016 providing 8 months worth of data to study microseisms but also the offshore seismic activity in the area. One of the main goal of the survey is to investigate the spatial and temporal distributions of dominant microseism source regions, close to the microseism sources. Additionally we will study the coupling of seismic and acoustic signals at the sea bed and its evolution in both the deep water and continental shelf areas. Furthermore, the survey also aims to investigate further the relationship between sea state conditions (e.g. wave height, period), seafloor pressure variations and seismic data recorded on both land and seafloor. Finally, the deployed OBS network is also the first ever attempt to closely monitor local offshore earthquakes in Ireland. Ireland seismicity although relatively low can reduce slope stability and poses the possibility of triggering large offshore landslides and local tsunamis.

  16. Use of glacial fronts by narwhals (Monodon monoceros) in West Greenland

    PubMed Central

    Moon, Twila; Hauser, Donna D. W.; McGovern, Richard; Heide-Jørgensen, Mads Peter; Dietz, Rune; Hudson, Ben

    2016-01-01

    Glacial fronts are important summer habitat for narwhals (Monodon monoceros); however, no studies have quantified which glacial properties attract whales. We investigated the importance of glacial habitats using telemetry data from n = 15 whales tagged in September of 1993, 1994, 2006 and 2007 in Melville Bay, West Greenland. For 41 marine-terminating glaciers, we estimated (i) narwhal presence/absence, (ii) number of 24 h periods spent at glaciers and (iii) the fraction of narwhals that visited each glacier (at 5, 7 and 10 km) in autumn. We also compiled data on glacier width, ice thickness, ice velocity, front advance/retreat, area and extent of iceberg discharge, bathymetry, subglacial freshwater run-off and sediment flux. Narwhal use of glacial habitats expanded in the 2000s probably due to reduced summer fast ice and later autumn freeze-up. Using a generalized multivariate framework, glacier ice front thickness (vertical height in the water column) was a significant covariate in all models. A negative relationship with glacier velocity was included in several models and glacier front width was a significant predictor in the 2000s. Results suggest narwhals prefer glaciers with potential for higher ambient freshwater melt over glaciers with silt-laden discharge. This may represent a preference for summer freshwater habitat, similar to other Arctic monodontids. PMID:27784729

  17. Community Development in Ireland.

    ERIC Educational Resources Information Center

    Lee, Anna

    2003-01-01

    In the last decade, community development in Ireland has emphasized social and economic inclusion, regeneration, and civic participation. Continuing challenges include designation of diverse community representatives, demand for increased administrative efficiency, and management of mandates and accountability. There are more community development…

  18. Creating the Social Foundations for Apprenticeship in Ireland

    ERIC Educational Resources Information Center

    Nyhan, Barry; Ayres, Paul

    2009-01-01

    Purpose: The purpose of this paper is to discuss Irelands national apprenticeship programme, introduced in 1993, in the context of the country's evolving economic and social policies. Design/methodology/approach: A critical analysis is undertaken of the industrial climate in Ireland, which prevented the introduction of a national apprenticeship…

  19. Estuarine removal of glacial iron and implications for iron fluxes to the ocean

    USGS Publications Warehouse

    Schroth, Andrew W.; Crusius, John; Hoyer, Ian; Campbell, Robert

    2014-01-01

    While recent work demonstrates that glacial meltwater provides a substantial and relatively labile flux of the micronutrient iron to oceans, the role of high-latitude estuary environments as a potential sink of glacial iron is unknown. Here we present the first quantitative description of iron removal in a meltwater-dominated estuary. We find that 85% of “dissolved” Fe is removed in the low-salinity region of the estuary along with 41% of “total dissolvable” iron associated with glacial flour. We couple these findings with hydrologic and geochemical data from Gulf of Alaska (GoA) glacierized catchments to calculate meltwater-derived fluxes of size and species partitioned Fe to the GoA. Iron flux data indicate that labile iron in the glacial flour and associated Fe minerals dominate the meltwater contribution to the Fe budget of the GoA. As such, GoA nutrient cycles and related ecosystems could be strongly influenced by continued ice loss in its watershed.

  20. A Comparative Examination of Schools' Responses to Bereavement and the Associated Needs of the School Community in Galway, West of Ireland and Derry, Northern Ireland

    ERIC Educational Resources Information Center

    McGovern, Marguerita; Tracey, Anne

    2010-01-01

    The aim and objective of this study is to examine and compare how schools in Galway, Republic of Ireland and Derry in the North of Ireland (cities located within two independent jurisdictions in Ireland) manage and respond to bereavement. To carry out a survey of schools, the "Loss in Schools" questionnaire is considered the most…

  1. Palaeoceanography. Antarctic stratification and glacial CO2.

    PubMed

    Keeling, R F; Visbeck, M

    2001-08-09

    One way of accounting for lowered atmospheric carbon dioxide concentrations during Pleistocene glacial periods is by invoking the Antarctic stratification hypothesis, which links the reduction in CO2 to greater stratification of ocean surface waters around Antarctica. As discussed by Sigman and Boyle, this hypothesis assumes that increased stratification in the Antarctic zone (Fig. 1) was associated with reduced upwelling of deep waters around Antarctica, thereby allowing CO2 outgassing to be suppressed by biological production while also allowing biological production to decline, which is consistent with Antarctic sediment records. We point out here, however, that the response of ocean eddies to increased Antarctic stratification can be expected to increase, rather than reduce, the upwelling rate of deep waters around Antarctica. The stratification hypothesis may have difficulty in accommodating eddy feedbacks on upwelling within the constraints imposed by reconstructions of winds and Antarctic-zone productivity in glacial periods.

  2. Gigantic landslides versus glacial deposits: on origin of large hummock deposits in Alai Valley, Northern Pamir

    NASA Astrophysics Data System (ADS)

    Reznichenko, Natalya

    2015-04-01

    As glaciers are sensitive to local climate, their moraines position and ages are used to infer past climates and glacier dynamics. These chronologies are only valid if all dated moraines are formed as the result of climatically driven advance and subsequent retreat. Hence, any accurate palaeoenvironmental reconstruction requires thorough identification of the landform genesis by complex approach including geomorphological, sedimentological and structural landform investigation. Here are presented the implication of such approach for the reconstruction of the mega-hummocky deposits formation both of glacial and landslide origin in the glaciated Alai Valley of the Northern Pamir with further discussion on these and similar deposits validity for palaeoclimatic reconstructions. The Tibetan Plateau valleys are the largest glaciated regions beyond the ice sheets with high potential to provide the best geological record of glacial chronologies and, however, with higher probabilities of the numerous rock avalanche deposits including those that were initially considered of glacial origin (Hewitt, 1999). The Alai Valley is the largest intermountain depression in the upper reaches of the Amudarja River basin that has captured numerous unidentified extensive hummocky deposits descending from the Zaalai Range of Northern Pamir, covering area in more than 800 km2. Such vast hummocky deposits are usually could be formed either: 1) glacially by rapid glacial retreat due to the climate signal or triggered a-climatically glacial changes, such as glacial surge or landslide impact, or 2) during the landslide emplacement. Combination of sediment tests on agglomerates forming only in rock avalanche material (Reznichenko et al., 2012) and detailed geomorphological and sedimentological descriptions of these deposits allowed reconstructing the glacial deposition in the Koman and Lenin glacial catchments with identification of two gigantic rock avalanches and their relation to this glacial

  3. Towards a geology training and outreach centre in western Ireland

    NASA Astrophysics Data System (ADS)

    Lacchia, Anthea; Haughton, Peter; Shannon, Patrick

    2017-04-01

    An outreach and education centre is in the initial phases of development for the coastal area of County Clare, western Ireland. The high Carboniferous sea cliffs of the Loop Head area provide a rich training ground for geoscientists from industry and academia. The cliffs offer a unique, margin-scale perspective of a sedimentary basin fill succession that developed during the height of the Late Palaeozoic glaciation. The rocks, about which there is a long legacy of research, record several glacial cycles, associated with significant eustatic changes in sea level. For geoscientists working with or in industry, the value of the area lies in its analogy with hydrocarbon-bearing, deltaic to deep-water sedimentary successions on several continental margins, such as Miocene and Pliocene successions in the Gulf of Mexico. A programme of behind-outcrop drilling involving UCD and Statoil has acquired over 1350 m of core from 12 boreholes behind the sea cliffs. This core is already being used in training and research both in UCD and at Statoil. The coastal cliffs are also visited by tourists and special interest groups, such as birdwatchers. It is envisioned that the centre will involve the local community and wider public, facilitating links between geoscience, energy and environment. Transport of cores and training materials to the centre, where they will be made available to visiting field parties, is planned for this year. Progress to date, including public engagement activities with schools and at conferences as well as audience research and public consultation, and future plans will be outlined.

  4. Invertebrate Metacommunity Structure and Dynamics in an Andean Glacial Stream Network Facing Climate Change

    PubMed Central

    Cauvy-Fraunié, Sophie; Espinosa, Rodrigo; Andino, Patricio; Jacobsen, Dean; Dangles, Olivier

    2015-01-01

    Under the ongoing climate change, understanding the mechanisms structuring the spatial distribution of aquatic species in glacial stream networks is of critical importance to predict the response of aquatic biodiversity in the face of glacier melting. In this study, we propose to use metacommunity theory as a conceptual framework to better understand how river network structure influences the spatial organization of aquatic communities in glacierized catchments. At 51 stream sites in an Andean glacierized catchment (Ecuador), we sampled benthic macroinvertebrates, measured physico-chemical and food resource conditions, and calculated geographical, altitudinal and glaciality distances among all sites. Using partial redundancy analysis, we partitioned community variation to evaluate the relative strength of environmental conditions (e.g., glaciality, food resource) vs. spatial processes (e.g., overland, watercourse, and downstream directional dispersal) in organizing the aquatic metacommunity. Results revealed that both environmental and spatial variables significantly explained community variation among sites. Among all environmental variables, the glacial influence component best explained community variation. Overland spatial variables based on geographical and altitudinal distances significantly affected community variation. Watercourse spatial variables based on glaciality distances had a unique significant effect on community variation. Within alpine catchment, glacial meltwater affects macroinvertebrate metacommunity structure in many ways. Indeed, the harsh environmental conditions characterizing glacial influence not only constitute the primary environmental filter but also, limit water-borne macroinvertebrate dispersal. Therefore, glacier runoff acts as an aquatic dispersal barrier, isolating species in headwater streams, and preventing non-adapted species to colonize throughout the entire stream network. Under a scenario of glacier runoff decrease, we

  5. Enhanced East Pacific Rise hydrothermal activity during the last two glacial terminations

    NASA Astrophysics Data System (ADS)

    Lund, D. C.; Asimow, P. D.; Farley, K. A.; Rooney, T. O.; Seeley, E.; Jackson, E. W.; Durham, Z. M.

    2016-01-01

    Mid-ocean ridge magmatism is driven by seafloor spreading and decompression melting of the upper mantle. Melt production is apparently modulated by glacial-interglacial changes in sea level, raising the possibility that magmatic flux acts as a negative feedback on ice-sheet size. The timing of melt variability is poorly constrained, however, precluding a clear link between ridge magmatism and Pleistocene climate transitions. Here we present well-dated sedimentary records from the East Pacific Rise that show evidence of enhanced hydrothermal activity during the last two glacial terminations. We suggest that glacial maxima and lowering of sea level caused anomalous melting in the upper mantle and that the subsequent magmatic anomalies promoted deglaciation through the release of mantle heat and carbon at mid-ocean ridges.

  6. Early warming of tropical South America at the last glacial-interglacial transition.

    PubMed

    Seltzer, G O; Rodbell, D T; Baker, P A; Fritz, S C; Tapia, P M; Rowe, H D; Dunbar, R B

    2002-05-31

    Glaciation in the humid tropical Andes is a sensitive indicator of mean annual temperature. Here, we present sedimentological data from lakes beyond the glacial limit in the tropical Andes indicating that deglaciation from the Last Glacial Maximum led substantial warming at high northern latitudes. Deglaciation from glacial maximum positions at Lake Titicaca, Peru/Bolivia (16 degrees S), and Lake Junin, Peru (11 degrees S), occurred 22,000 to 19,500 calendar years before the present, several thousand years before the Bølling-Allerød warming of the Northern Hemisphere and deglaciation of the Sierra Nevada, United States (36.5 degrees to 38 degrees N). The tropical Andes deglaciated while climatic conditions remained regionally wet, which reflects the dominant control of mean annual temperature on tropical glaciation.

  7. Computer modelling of anthelmintic resistance and worm control outcomes for refugia-based nematode control strategies in Merino ewes in Western Australia.

    PubMed

    Cornelius, M P; Jacobson, C; Dobson, R; Besier, R B

    2016-04-15

    This study utilised computer simulation modelling (Risk Management Model for Nematodes) to investigate the impact of different parasite refugia scenarios on the development of anthelmintic resistance and worm control effectiveness. The simulations were conducted for adult ewe flocks in a Mediterranean climatic region over a 20 year time period. Factors explored in the simulation exercise were environment (different weather conditions), drug efficacy, the percentage of the flock left untreated, the timing of anthelmintic treatments, the initial worm egg count, and the number of drenches per annum. The model was run with variable proportions of the flock untreated (0, 10, 20, 30, 40 and 50%), with ewes selected at random so that reductions in the mean worm burden or egg count were proportional to the treated section of the flock. Treatments to ewes were given either in summer (December; low refugia potential, hence highly selective) or autumn (March; less selective due to a greater refugia potential), and the use of different anthelmintics was simulated to indicate the difference between active ingredients of different efficacy. Each model scenario was run for two environments, specifically a lower rainfall area (more selective) and a higher rainfall area (less selective) within a Mediterranean climatic zone, characterised by hot, dry summers and cool, wet winters. Univariate general linear models with least square difference post-hoc tests were used to examine differences between means of factors. The results confirmed that leaving a proportion of sheep in a flock untreated was effective in delaying the development of anthelmintic resistance, with as low as 10% of a flock untreated sufficient to significantly delay resistance, although this strategy was associated with a small reduction in worm control. Administering anthelmintics in autumn rather than summer was also effective in delaying the development of anthelmintic resistance in the lower rainfall

  8. Comparing Terrestrial Organic Carbon Cycle Dynamics in Interglacial and Glacial Climates in the South American Tropics

    NASA Astrophysics Data System (ADS)

    Fornace, K. L.; Galy, V.; Hughen, K. A.

    2014-12-01

    The application of compound-specific radiocarbon dating to molecular biomarkers has allowed for tracking of specific organic carbon pools as they move through the environment, providing insight into complex processes within the global carbon cycle. Here we use this technique to investigate links between glacial-interglacial climate change and terrestrial organic carbon cycling in the catchments of Cariaco Basin and Lake Titicaca, two tropical South American sites with well-characterized climate histories since the last glacial period. By comparing radiocarbon ages of terrestrial biomarkers (leaf wax compounds) with deposition ages in late glacial and Holocene sediments, we are able to gauge the storage time of these compounds in the catchments in soils, floodplains, etc. before transport to marine or lacustrine sediments. We are also able to probe the effects of temperature and hydrologic change individually by taking advantage of opposite hydrologic trends at the two sites: while both were colder during the last glacial period, precipitation at Titicaca decreased from the last glacial period to the Holocene, but the late glacial was marked by drier conditions at Cariaco. Preliminary data from both sites show a wide range of apparent ages of long-chain n-fatty acids (within error of 0 to >10,000 years older than sediment), with the majority showing ages on the order of several millennia at time of deposition and age generally increasing with chain length. While late glacial leaf waxes appear to be older relative to sediment than those deposited in the Holocene at both sites, at Cariaco we find a ~2-3 times larger glacial-interglacial age difference than at Titicaca. We hypothesize that at Titicaca the competing influences of wetter and colder conditions during the last glacial period, which respectively tend to increase and decrease the rate of organic carbon turnover on land, served to minimize the contrast between glacial and interglacial leaf wax storage time

  9. Termination of the Last Glacial Maximum sea-level lowstand: The Sunda-Shelf data revisited

    NASA Astrophysics Data System (ADS)

    Hanebuth, T. J. J.; Stattegger, K.; Bojanowski, A.

    2009-03-01

    The sea-level history around the last glaciation is in the focus of recent, controversial debates. A profound understanding of sea-level changes during this time interval is, however, essential since sea level is a central parameter in the climate system as well as a major force on continental margin sedimentation. Here, we present a seismic record together with augmented data from the Sunda Shelf [Hanebuth, T.J.J., Stattegger, K., Saito, Y., 2002. The architecture of the central Sunda Shelf (SE Asia) recorded by shallow-seismic surveying. Geo-Marine Letters 22, 86-94.] and compare our results in a careful evaluation with the sparse existing data sets of global validity, i.e. the Bonaparte Gulf record [Yokoyama, Y., Lambeck, K., DeDeckker, P., Johnston, P., Fifield, L.K., 2000. Timing of the Last Glacial Maximum from observed sea-level minima. Nature 406, 713-716.; Yokoyama, Y., De Deckker, P., Lambeck, K., Johnston, P., Fifield, L.K., 2001. Sea-level at the Last Glacial Maximum: evidence from nortwestern Australia to constrain ice volumes for oxygen isotope stage 2. Paleogeography Paleoclimatology Paleoecology 165, 281-297.], the Barbados coral record [Fairbanks, R.G., 1989. A 17,000-year glacio-eustatic sea level record: influence of glacial melting dates on the Younger Dryas event and deep ocean circulation. Nature 342, 637-642.; Peltier, W.R., Fairbanks, R.G., 2006. Global glacial ice volume and Last Glacial Maximum duration from an extended Barbados sea level record. Quaternary Science Reviews 25 (23-24), 3322-3337.] and the latest numerical model of continental deglaciation [Peltier, W.R., Fairbanks, R.G., 2006. Global glacial ice volume and Last Glacial Maximum duration from an extended Barbados sea level record. Quaternary Science Reviews 25 (23-24), 3322-3337.]. Sea level seems to have been lower shortly prior to the conventional Last Glacial Maximum (LGM; 21-19 cal kyr BP). The time interval around this glacial lowstand is not covered by ages from the

  10. Cytogeography of the Humifusa clade of Opuntia s.s. Mill. 1754 (Cactaceae, Opuntioideae, Opuntieae): correlations with pleistocene refugia and morphological traits in a polyploid complex

    PubMed Central

    Majure, Lucas C.; Judd, Walter S.; Soltis, Pamela S.; Soltis, Douglas E.

    2012-01-01

    southwestern U.S. (including all southwestern U.S. diploids and polyploids). In addition, tetraploid Opuntia humifusa s.l., which occurs primarily in the eastern U.S., is resolved in the southwestern diploid clade instead of with the southeastern diploid clade that includes diploid Opuntia humifusa s.l. Our results not only provide evidence for the polyphyletic nature of Opuntia humifusa and Opuntia macrorhiza, suggesting that each of these represents more than one species, but also demonstrate the high frequency of polyploidy in the Humifusa clade and the major role that genome duplication has played in the diversification of this lineage of Opuntia s.s. Our data also suggest that the southeastern and southwestern U.S. may represent glacial refugia for diploid members of this clade and that the clade as a whole should be considered a mature polyploid species complex. Widespread polyploids are likely derivatives of secondary contact among southeastern and southwestern diploid taxa as a result of the expansion and contraction of suitable habitat during the Pleistocene following glacial and interglacial events. PMID:24260652

  11. Education Policy Outlook: Ireland

    ERIC Educational Resources Information Center

    Pont, Beatriz; Figueroa, Diana Toledo; Zapata, Juliana; Fraccola, Sylvain

    2013-01-01

    This policy profile on education in Ireland is part of the "Education Policy Outlook" series, which presents comparative analysis of education policies and reforms across the Organisation for Economic Co-operation and Development (OECD) countries. Building on the OECD's substantial comparative and sectoral knowledge base, the series…

  12. Assessing Glacial Lake Outburst Flood Hazard in the Nepal Himalayas using Satellite Imagery and Hydraulic Models

    NASA Astrophysics Data System (ADS)

    Rounce, D.; McKinney, D. C.

    2015-12-01

    The last half century has witnessed considerable glacier melt that has led to the formation of large glacial lakes. These glacial lakes typically form behind terminal moraines comprising loose boulders, debris, and soil, which are susceptible to fail and cause a glacial lake outburst flood (GLOF). These lakes also act as a heat sink that accelerates glacier melt and in many cases is accompanied by rapid areal expansion. As these glacial lakes continue to grow, their hazard also increases due to the increase in potential flood volume and the lakes' proximity to triggering events such as avalanches and landslides. Despite the large threat these lakes may pose to downstream communities, there are few detailed studies that combine satellite imagery with hydraulic models to present a holistic understanding of the GLOF hazard. The aim of this work is to assess the GLOF hazard of glacial lakes in Nepal using a holistic approach based on a combination of satellite imagery and hydraulic models. Imja Lake will be the primary focus of the modeling efforts, but the methods will be developed in a manner that is transferable to other potentially dangerous glacial lakes in Nepal.

  13. Local Communities and Glacial Lake Outburst Flood Mitigation: Lessons from Peru

    NASA Astrophysics Data System (ADS)

    Carey, Mark

    2010-05-01

    Discourse in recent years among scientists and non-scientists increasingly promotes the involvement of local people in hazard mitigation, including inhabitants of floodplains in valleys below moraine-dammed glacial lakes. Despite advances in understanding human vulnerability to glacial lake outburst floods, there has been much less research on how these vulnerable populations are involved (or ignored) in the actual outburst flood mitigation process. Which groups should be involved? Are they in fact participating? Is that involvement successful? Peru's Cordillera Blanca mountain range provides an ideal site to help answer these questions because its moraine-dammed glacial lakes have produced more than a dozen outburst floods since ~1860. After floods in 1941, 1945, and 1950 killed approximately 6,000, the national government created a state agency, which still exists today, to monitor glacial lakes and prevent future outburst floods. Using this region as a case study to answer the above questions, this paper has three components. First, it provides historical examples of local people's participation in disaster mitigation, but shows that the outcome of such local involvement frequently turned out differently than scientists, engineers, and planners anticipated. Second, it shows the challenges and difficulties of involving local groups. Recent efforts in workshops, aid projects, and government programs show only limited success in community participation in disaster mitigation agendas. Third, the paper suggests that in many cases local indigenous people, as icons of the Andean region but often not the most vulnerable group, are disproportionately victimized and tacitly invited into disaster mitigation discussions. Poor urban residents inhabiting floodplains are often neglected, even though they are the most vulnerable to outburst floods. As other world regions such as the Himalayas increasingly contend with potential glacial lake outburst floods, these lessons from

  14. Laurentide glacial landscapes: the role of ice streams

    USGS Publications Warehouse

    Patterson, C.J.

    1998-01-01

    Glacial landforms of the North American prairie can be divided into two suites that result from different styles of ice flow: 1) a lowland suite of level-to-streamlined till consistent with formation beneath ice streams, and 2) an upland and lobe-margin suite of thick, hummocky till and glacial thrust blocks consistent with formation at ice-stream and ice-lobe margins. Southern Laurentide ice lobes hypothetically functioned as outlets of ice streams. Broad branching lowlands bounded by escarpments mark the stable positions of the ice streams that fed the lobes. If the lobes and ice streams were similar to modern ice streams, their fast flow was facilitated by high subglacial water pressure. Favorable geology and topography in the midcontinent encouraged nonuniform ice flow and controlled the location of ice streams and outlet lobes.

  15. Small Rural Schools in Ireland: Problems and Possibilities.

    ERIC Educational Resources Information Center

    Sugrue, Ciaran

    This paper provides an overview of practices in small rural elementary schools in Ireland and recent trends related to school size. There are 3,200 "ordinary" elementary schools in the Republic of Ireland serving children aged 4-12 in eight levels: two preschool levels and grades 1-6. System-wide policies with implications for small…

  16. INFOMAR, Ireland's National Seabed Mapping Programme; Sharing Valuable Insights.

    NASA Astrophysics Data System (ADS)

    Judge, M. T.; McGrath, F.; Cullen, S.; Verbruggen, K.

    2017-12-01

    Following the successful high-resolution deep-sea mapping carried out as part of the Irish National Seabed Survey (INSS), a strategic, long term programme was established: INtegrated mapping FOr the sustainable development of Ireland MArine Resources (INFOMAR). Funded by Ireland's Department of Communication, Climate Action and Environment, INFOMAR comprises a multi-platform approach to completing Ireland's marine mapping, and is a key action in the integrated marine plan, Harnessing Our Ocean Wealth. Co-managed by Geological Survey Ireland and the Marine Institute, the programme has three work strands: Data Acquisition; Data Exchange and Integration; Value Added Exploitation.The Data Acquisition strand includes collection of geological, hydrographic, oceanographic, habitat and heritage datasets that underpin sustainable development and management of Ireland's marine resources. INFOMAR operates a free data policy; data and outputs are delivered online through the Data Exchange and Integration strand. Uses of data and outputs are wide-ranging and multipurpose. In order to address the evolution and diversification of user requirements, further data product development is facilitated through the Value Added Exploitation strand.Ninety percent of Ireland's territory lies offshore. Therefore, strategic national seabed mapping continues to provide critical, high-resolution baseline datasets for numerous economic sectors and societal needs. From these we can glean important geodynamic knowledge of Ireland's vast maritime territory. INFOMAR remains aligned with national and European policies and directives. Exemplified by our commitment to EMODnet, a European Commission funded project that supports the collection, standardisation and sharing of available marine information, data and data products across all European Seas. As EMODnet Geology Minerals leaders we have developed a framework for mapping marine minerals. Furthermore, collaboration with the international research

  17. Hydraulic properties of three types of glacial deposits in Ohio

    USGS Publications Warehouse

    Strobel, M.L.

    1993-01-01

    The effects of thickness, grain size, fractures, weathering, and atmosphericconditions on vertical ground-water flow in glacial deposits were studied at three sites that represent ground moraine, end moraine, and lacustrine depositional environments. Vertical hydraulic conductivities computed from pumped-well tests were 3.24 x 10-1 to 6.47 x 10-1 ft/d (feet per day) at the site representing end moraine and 1.17 ft/d at the site representing lacustrine deposits. Analysis of test data for the ground moraine site did not yield estimates of hydraulic conductivities, but did indicate that ground water flows through the total thickness of deposits in response to discharge from a lower gravel unit. Vertical hydraulic conductivities computed from pumped-well tests of nested wells and data from drill-core analyses indicate that fractures affect the migration of ground water downward through the glacial deposits at these sites. Flow through glacial deposits is complex; it is controlled by fractures, gram-size distribution, clay content, thickness, and degree of weathering, and atmospheric conditions.

  18. Changing climatic response: a conceptual model for glacial cycles and the Mid-Pleistocene Transition

    NASA Astrophysics Data System (ADS)

    Daruka, I.; Ditlevsen, P. D.

    2014-03-01

    Milankovitch's astronomical theory of glacial cycles, attributing ice age climate oscillations to orbital changes in Northern Northern-Hemisphere insolation, is challenged by the paleoclimatic record. The climatic response to the variations in insolation is far from trivial. In general the glacial cycles are highly asymmetric in time, with slow cooling from the interglacials to the glacials (inceptions) and very rapid warming from the glacials to the interglacials (terminations). We shall refer to this fast-slow dynamics as the "saw-tooth" shape of the paleoclimatic record. This is non-linearly related to the time-symmetric variations in the orbital forcing. However, the most pronounced challenge to the Milankovitch theory is the Mid-Pleistocene Transition (MPT) occurring about one million years ago. During that event, the prevailing 41 kyr glacial cycles, corresponding to the almost harmonic obliquity cycle were replaced by longer saw-tooth shaped cycles with a time scale around 100 kyr. The MPT must have been driven by internal changes in climate response, since it does not correspond to any apparent changes in the orbital forcing. In order to identify possible mechanisms causing the observed changes in glacial dynamics, it is relevant to study simplified models with the capability of generating temporal behavior similar to the observed records. We present a simple oscillator type model approach, with two variables, a temperature anomaly and an ice volume analogous, climatic memory term. The generalization of the ice albedo feedback is included in terms of an effective multiplicative coupling between this latter climatic memory term (representing the internal degrees of freedom) and the external drive. The simple model reproduces the temporal asymmetry of the late Pleistocene glacial cycles and suggests that the MPT can be explained as a regime shift, aided by climatic noise, from a period 1 frequency locking to the obliquity cycle to a period 2-3 frequency

  19. Trinity mysteries: university, elite schooling and sport in Ireland.

    PubMed

    Finn, Gerry P T

    2010-01-01

    The development of sport in Ireland was, contrary to some arguments, highly influenced by English examples and Anglo-Irish institutions. Trinity College and prestigious Irish schools did have an impact, as did the number of Irish students sent to England for public school or university education. Athleticism was evident in Ireland as it was in England. Although the development of soccer did follow a slightly different trajectory from other sports, as was also the case in both England and Scotland, this does not mean that it departed from this broad evolutionary model of Irish sport. Yet this was Ireland: and Ireland was different. As opposition to British rule intensified, forms of sporting participation took on more and more of a national symbolism. The outcome was the emergence of a very potent form of athleticism: an Irish athleticism for an Irish people.

  20. Quantitative Morphometric Analysis of Terrestrial Glacial Valleys and the Application to Mars

    NASA Astrophysics Data System (ADS)

    Allred, Kory

    Although the current climate on Mars is very cold and dry, it is generally accepted that the past environments on the planet were very different. Paleo-environments may have been warm and wet with oceans and rivers. And there is abundant evidence of water ice and glaciers on the surface as well. However, much of that comes from visual interpretation of imagery and other remote sensing data. For example, some of the characteristics that have been utilized to distinguish glacial forms are the presence of landscape features that appear similar to terrestrial glacial landforms, constraining surrounding topography, evidence of flow, orientation, elevation and valley shape. The main purpose of this dissertation is to develop a model that uses quantitative variables extracted from elevation data that can accurately categorize a valley basin as either glacial or non-glacial. The application of this model will limit the inherent subjectivity of image analysis by human interpretation. The model developed uses hypsometric attributes (elevation-area relationship), a newly defined variable similar to the equilibrium line altitude for an alpine glacier, and two neighborhood search functions intended to describe the valley cross-sectional curvature, all based on a digital elevation model (DEM) of a region. The classification model uses data-mining techniques trained on several terrestrial mountain ranges in varied geologic and geographic settings. It was applied to a select set of previously catalogued locations on Mars that resemble terrestrial glaciers. The results suggest that the landforms do have a glacial origin, thus supporting much of the previous research that has identified the glacial landforms. This implies that the paleo-environment of Mars was at least episodically cold and wet, probably during a period of increased planetary obliquity. Furthermore, the results of this research and the implications thereof add to the body of knowledge for the current and past

  1. Forest refugia in Western and Central Africa as 'museums' of Mesozoic biodiversity.

    PubMed

    Murienne, Jérôme; Benavides, Ligia R; Prendini, Lorenzo; Hormiga, Gustavo; Giribet, Gonzalo

    2013-02-23

    The refugial speciation model, or 'species pump', is widely accepted in the context of tropical biogeography and has been advocated as an explanation for present species distributions in tropical Western and Central Africa. In order to test this hypothesis, a phylogeny of the cryptic arachnid order Ricinulei, based on four nuclear and mitochondrial DNA markers, was inferred. This ancient clade of litter-dwelling arthropods, endemic to the primary forests of Western and Central Africa and the Neotropics, might provide insights into the mode and tempo of evolution in Africa. Twenty-six African ricinuleid specimens were sampled from eight countries spanning the distribution of Ricinulei on the continent, and analysed together with Neotropical samples plus other arachnid outgroups. The phylogenetic and molecular dating results suggest that Ricinulei diversified in association with the fragmentation of Gondwana. The early diversification of Ricinoides in Western and Central Africa around 88 (±33) Ma fits old palaeogeographical events better than recent climatic fluctuations. Unlike most recent molecular studies, these results agree with fossil evidence, suggesting that refugia may have acted as 'museums' conserving ancient diversity rather than as engines generating diversity during successive episodes of climatic fluctuation in Africa.

  2. Climate refugia: The physical, hydrologic and disturbance basis

    NASA Astrophysics Data System (ADS)

    Holden, Z. A.; Maneta, M. P.; Forthofer, J.

    2015-12-01

    Projected changes in global climate and associated shifts in vegetation have increased interest in understanding species persistence at local scales. We examine the climatic and physical factors that could mediate changes in the distribution of vegetation in regions of complex topography. Using massive networks of low-cost temperature and humidity sensors, we developed topographically-resolved daily historical gridded temperature data for the US Northern Rockies. We used the WindNinja model to create daily historical wind speed maps across the same domain. Using a spatially distributed ecohydrology model (ECH2O) we examine separately the sensitivity of modeled evapotranspiration and soil moisture to wind, radiation, soil properties, minimum temperature and humidity. A suite of physical factors including lower wind speeds, cold air drainage, solar shading and increased soil depth reduce evapotranspiration and increase late season moisture availability in valley bottoms. Evapotranspiration shows strong sensitivity to spatial variability in surface wind speed, suggesting that sheltering effects from winds may be an important factor contributing to mountain refugia. Fundamental to our understanding of patterns of vegetation change is the role of stand-replacing wildfires, which modify the physical environment and subsequent patterns of species persistence and recruitment. Using satellite-derived maps of burn severity for recent fires in the US Northern Rockies we examined relationships between wind speed, cold air drainage potential and soil depth and the occurrence of unburned and low severity fire. Severe fire is less likely to occur in areas with high cold air drainage potential and low wind speeds, suggesting that sheltered valley bottoms have mediated the severity of recent wildfires. Our finding highlight the complex physical mechanisms by which mountain weather and climate mediate fire-induced vegetation changes in the US Northern Rocky Mountains.

  3. Use of glacial fronts by narwhals (Monodon monoceros) in West Greenland.

    PubMed

    Laidre, Kristin L; Moon, Twila; Hauser, Donna D W; McGovern, Richard; Heide-Jørgensen, Mads Peter; Dietz, Rune; Hudson, Ben

    2016-10-01

    Glacial fronts are important summer habitat for narwhals (Monodon monoceros); however, no studies have quantified which glacial properties attract whales. We investigated the importance of glacial habitats using telemetry data from n = 15 whales tagged in September of 1993, 1994, 2006 and 2007 in Melville Bay, West Greenland. For 41 marine-terminating glaciers, we estimated (i) narwhal presence/absence, (ii) number of 24 h periods spent at glaciers and (iii) the fraction of narwhals that visited each glacier (at 5, 7 and 10 km) in autumn. We also compiled data on glacier width, ice thickness, ice velocity, front advance/retreat, area and extent of iceberg discharge, bathymetry, subglacial freshwater run-off and sediment flux. Narwhal use of glacial habitats expanded in the 2000s probably due to reduced summer fast ice and later autumn freeze-up. Using a generalized multivariate framework, glacier ice front thickness (vertical height in the water column) was a significant covariate in all models. A negative relationship with glacier velocity was included in several models and glacier front width was a significant predictor in the 2000s. Results suggest narwhals prefer glaciers with potential for higher ambient freshwater melt over glaciers with silt-laden discharge. This may represent a preference for summer freshwater habitat, similar to other Arctic monodontids. © 2016 The Author(s).

  4. Family and Family Change in Ireland: An Overview

    ERIC Educational Resources Information Center

    Canavan, John

    2012-01-01

    In Ireland, historically and in the current era, family has been a central concern for society and the State. This article provides a descriptive overview of family life in Ireland and of major family-related changes over the past 40 years. It presents a general framework of analysis within which these changes can be understood, considers the…

  5. Glacial magnetite dissolution in abyssal NW Pacific sediments - evidence for carbon trapping?

    NASA Astrophysics Data System (ADS)

    Korff, Lucia; von Dobeneck, Tilo; Frederichs, Thomas; Kasten, Sabine; Kuhn, Gerhard; Gersonde, Rainer; Diekmann, Bernhard

    2016-04-01

    The abyssal North Pacific Ocean's large volume, depth, and terminal position on the deep oceanic conveyor make it a candidate site for deep carbon trapping as postulated by climate theory to explain the massive glacial drawdown of atmospheric CO2. As the major basins of the North Pacific have depths of 5500-6500m, far below the modern and glacial Calcite Compensation Depths (CCD), these abyssal sediments are carbonate-free and therefore not suitable for carbonate-based paleoceanographic proxy reconstructions. Instead, paleo-, rock and environmental magnetic methods are generally well applicable to hololytic abyssal muds and clays. In 2009, the international paleoceanographic research cruise SO 202 INOPEX ('Innovative North Pacific Experiment') of the German RV SONNE collected two ocean-spanning EW sediment core transects of the North Pacific and Bering Sea recovering a total of 50 piston and gravity cores from 45 sites. Out of seven here considered abyssal Northwest Pacific piston cores collected at water depths of 5100 to 5700m with mostly coherent shipboard susceptibility logs, the 20.23m long SO202-39-3, retrieved from 5102 m water depth east of northern Shatsky Rise (38°00.70'N, 164°26.78'E), was rated as the stratigraphically most promising record of the entire core transect and selected for detailed paleo- and environmental magnetic, geochemical and sedimentological investigations. This core was dated by correlating its RPI and Ba/Ti records to well-dated reference records and obviously provides a continuous sequence of the past 940 kyrs. The most striking orck magnetic features are coherent magnetite-depleted zones corresponding to glacial periods. In the interglacial sections, detrital, volcanic and even submicron bacterial magnetite fractions are excellently preserved. These alternating magnetite preservation states seem to reflect dramatic oxygenation changes in the deep North Pacific Ocean and hint at large-scale benthic glacial carbon trapping

  6. Underreporting of viral encephalitis and viral meningitis, Ireland, 2005-2008.

    PubMed

    Kelly, Tara A; O'Lorcain, Piaras; Moran, Joanne; Garvey, Patricia; McKeown, Paul; Connell, Jeff; Cotter, Suzanne

    2013-01-01

    Viral encephalitis (VE) and viral meningitis (VM) have been notifiable infectious diseases under surveillance in the Republic of Ireland since 1981. Laboratories have reported confirmed cases by detection of viral nucleic acid in cerebrospinal fluid since 2004. To determine the prevalence of these diseases in Ireland during 2005-2008, we analyzed 3 data sources: Hospital In-patient Enquiry data (from hospitalized following patients discharge) accessed through Health Intelligence Ireland, laboratory confirmations from the National Virus Reference Laboratory, and events from the Computerised Infectious Disease Reporting surveillance system. We found that the national surveillance system underestimates the incidence of these diseases in Ireland with a 10-fold higher VE hospitalization rate and 3-fold higher VM hospitalization rate than the reporting rate. Herpesviruses were responsible for most specified VE and enteroviruses for most specified VM from all 3 sources. Recommendations from this study have been implemented to improve the surveillance of these diseases in Ireland.

  7. Friis Hills glacial history: an international collaboration to examine Miocene climate in Antarctica

    NASA Astrophysics Data System (ADS)

    Halberstadt, A. R. W.; Kowalewski, D. E.

    2016-12-01

    The Friis Hills, Antarctica (western McMurdo Dry Valleys) contain unique, well-preserved records of Miocene climate. These terrestrial deposits hold geomorphic clues for deciphering the glacial history in a region directly adjacent to the East Antarctic Ice Sheet. Stacked till sheets, interbedded with lake sediments and non-glacial deposits, reveal a complex history of ice flow and erosion throughout multiple glacial-interglacial cycles (Lewis and Ashworth, 2015). Fossiliferous beds containing Nothofagus, diatoms, algal cells, pollen, insects, and mosses provide past climatological constraints. The Friis Hills sustained multiple alpine glaciations as well as full ice-sheet development, recording glacial drainage reorganization and evidence of previous ice configurations that possibly overrode the Transantarctic Mountains (Lewis and Ashworth, 2015) exposing only scattered nunataks (i.e. a portion of Friis Hills). Lack of chronological control has previously hindered efforts to link the Friis Hills glacial history with regional context; a tephra deposit at the base of the glacial drifts currently provides a single age constraint within the drift deposits. To build upon previous studies, an international collaboration between the USAP, Antarctic New Zealand, and the Italian Antarctic community proposes to core a paleo-lake in the center of the Friis Hills in November 2016, thereby acquiring one of the oldest continuous sedimentological records within the McMurdo Dry Valleys. Here we report discoveries from this year's fieldwork, and reconstruct paleoenvironment at the periphery of the East Antarctic Ice Sheet for the mid-early Miocene, a critical time when marine isotopic records indicate dramatic ice fluctuations. Ash within the sediment core stratigraphy will provide a more robust chronology for the region, and will also suggest possible outcrop locations of corresponding ash deposits to pursue while in the field. We anticipate that the Friis Hills stratigraphy will

  8. Using Distributed Temperature Sensing to Locate and Quantify Thermal Refugia: Insights Into Radiative & Hydrologic Processes

    NASA Astrophysics Data System (ADS)

    Bond, R. M.; Stubblefield, A. P.

    2012-12-01

    Stream temperature plays a critical role in determining the overall structure and function of stream ecosystems. Aquatic fauna are particularly vulnerable to projected increases in the magnitude and duration of elevated stream temperatures from global climate change. Northern California cold water salmon and trout fisheries have been declared thermally impacted by the California State Water Resources Control Board. This study employed Distributed Temperature Sensing (DTS) to detect stream heating and cooling at one meter resolution along a one kilometer section of the North Fork of the Salmon River, a tributary of the Klamath River, northern California, USA. The Salmon River has an extensive legacy of hydraulic gold mining tailing which have been reworked into large gravel bars; creating shallow wide runs, possibly filling in pools and disrupting riparian vegetation recruitment. Eight days of temperature data were collected at 15 minute intervals during July 2012. Three remote weather stations were deployed during the study period. The main objectives of this research were: one, quantify thermal inputs that create and maintain thermal refugia for cold water fishes; two, investigate the role of riparian and topographic shading in buffering peak summer temperatures; and three, create and validate a physically based stream heating model to predict effects of riparian management, drought, and climate change on stream temperature. DTS was used to spatially identify cold water seeps and quantify their contribution to the stream's thermal regime. Along the one kilometer reach, hyporheic flow was identified using DTS. The spring was between 16-18°C while the peak mainstem temperature above the spring reached a maximum of 23°C. The study found a diel heating cycle of 5°C with a Maximum Weekly Average Temperature (MWAT) of over 22°C; exceeding salmon and trout protective temperature standards set by USEPA Region 10. Twenty intensive fish counts over five days were

  9. Interlobate comparison of glacial-depositional style as evidenced by small-relief glacial landscape features in Illinois, Indiana, and Ohio, utilizing SIR-B

    NASA Technical Reports Server (NTRS)

    Johnson, W. H.; Bleuer, N. K.; Fraser, G. S.; Totten, S. M.

    1984-01-01

    The objectives and expected results of an investigation of the use of the Shuttle Imaging Radar-B (SIR-B) as a basic tool in the recognition and mapping of glacial landforms are discussed. The main goals are: (1) to evaluate the ability of SIR-B to delineate varying sizes, shapes, and relief of surface forms; (2) to compare and contrast SIR-B imagery with selected Seasat SAR imagery; (3) to utilize SIR-B imagery synergistically with available SEASAT SAR, LANDSAT RBV, and other imagery sources to identify and map suites of glacial landforms; and (4) eventually to interpret the suites in terms of ice dynamics and conditions of deglaciation, to relate them to the stratigraphic record, and to evaluate interactions of the major lobes and sublobes.

  10. Origin of last-glacial loess in the western Yukon-Tanana Upland, central Alaska, USA

    USGS Publications Warehouse

    Muhs, Daniel; Pigati, Jeffrey S.; Budahn, James R.; Skipp, Gary L.; Bettis, E. Arthur; Jensen, Britta

    2018-01-01

    Loess is widespread over Alaska, and its accumulation has traditionally been associated with glacial periods. Surprisingly, loess deposits securely dated to the last glacial period are rare in Alaska, and paleowind reconstructions for this time period are limited to inferences from dune orientations. We report a rare occurrence of loess deposits dating to the last glacial period, ~19 ka to ~12 ka, in the Yukon-Tanana Upland. Loess in this area is very coarse grained (abundant coarse silt), with decreases in particle size moving south of the Yukon River, implying that the drainage basin of this river was the main source. Geochemical data show, however, that the Tanana River valley to the south is also a likely distal source. The occurrence of last-glacial loess with sources to both the south and north is explained by both regional, synoptic-scale winds from the northeast and opposing katabatic winds that could have developed from expanded glaciers in both the Brooks Range to the north and the Alaska Range to the south. Based on a comparison with recent climate modeling for the last glacial period, seasonality of dust transport may also have played a role in bringing about contributions from both northern and southern sources.

  11. The influence of glacial meltwater on alpine aquatic ecosystems: a review.

    PubMed

    Slemmons, Krista E H; Saros, Jasmine E; Simon, Kevin

    2013-10-01

    The recent and rapid recession of alpine glaciers over the last 150 years has major implications for associated aquatic communities. Glacial meltwater shapes many of the physical features of high altitude lakes and streams, producing turbid environments with distinctive hydrology patterns relative to nival systems. Over the past decade, numerous studies have investigated the chemical and biological effects of glacial meltwater on freshwater ecosystems. Here, we review these studies across both lake and stream ecosystems. Focusing on alpine regions mainly in the Northern Hemisphere, we present examples of how glacial meltwater can affect habitat by altering physical and chemical features of aquatic ecosystems, and review the subsequent effects on the biological structure and function of lakes and streams. Collectively or separately, these factors can drive the overall distribution, diversity and behavior of primary producers, triggering cascading effects throughout the food web. We conclude by proposing areas for future research, particularly in regions where glaciers are soon projected to disappear.

  12. Ice stream reorganization and glacial retreat on the northwest Greenland shelf

    NASA Astrophysics Data System (ADS)

    Newton, A. M. W.; Knutz, P. C.; Huuse, M.; Gannon, P.; Brocklehurst, S. H.; Clausen, O. R.; Gong, Y.

    2017-08-01

    Understanding conditions at the grounding-line of marine-based ice sheets is essential for understanding ice sheet evolution. Offshore northwest Greenland, knowledge of the Last Glacial Maximum (LGM) ice sheet extent in Melville Bugt was previously based on sparse geological evidence. This study uses multibeam bathymetry, combined with 2-D and 3-D seismic reflection data, to present a detailed landform record from Melville Bugt. Seabed landforms include mega-scale glacial lineations, grounding-zone wedges, iceberg scours, and a lateral shear margin moraine, formed during the last glacial cycle. The geomorphology indicates that the LGM ice sheet reached the shelf edge before undergoing flow reorganization. After retreat of 80 km across the outer shelf, the margin stabilized in a mid-shelf position, possibly during the Younger Dryas (12.9-11.7 ka). The ice sheet then decoupled from the seafloor and retreated to a coast-proximal position. This landform record provides an important constraint on deglaciation history offshore northwest Greenland.

  13. Climate change: potential implications for Ireland's biodiversity

    NASA Astrophysics Data System (ADS)

    Donnelly, Alison

    2018-03-01

    A national biodiversity and climate change adaptation plan is being developed for Ireland by the Department of Communications, Climate Action, and Environment. In order to inform such a plan, it was necessary to review and synthesize some of the recent literature pertaining to the impact of climate change on biodiversity in Ireland. Published research on this topic fell within three broad categories: (i) changes in the timing of life-cycle events (phenology) of plants, birds, and insects; (ii) changes in the geographic range of some bird species; and (iii) changes in the suitable climatic zones of key habitats and species. The synthesis revealed evidence of (i) a trend towards earlier spring activity of plants, birds, and insects which may result in a change in ecosystem function; (ii) an increase in the number of bird species; and (iii) both increases and decreases in the suitable climatic area of key habitats and species, all of which are expected to impact Ireland's future biodiversity. This process identified data gaps and limitations in available information both of which could be used to inform a focused research strategy. In addition, it raises awareness of the potential implications of climate change for biodiversity in Ireland and elsewhere and demonstrates the need for biodiversity conservation plans to factor climate change into future designs.

  14. Climate change: potential implications for Ireland's biodiversity.

    PubMed

    Donnelly, Alison

    2018-03-12

    A national biodiversity and climate change adaptation plan is being developed for Ireland by the Department of Communications, Climate Action, and Environment. In order to inform such a plan, it was necessary to review and synthesize some of the recent literature pertaining to the impact of climate change on biodiversity in Ireland. Published research on this topic fell within three broad categories: (i) changes in the timing of life-cycle events (phenology) of plants, birds, and insects; (ii) changes in the geographic range of some bird species; and (iii) changes in the suitable climatic zones of key habitats and species. The synthesis revealed evidence of (i) a trend towards earlier spring activity of plants, birds, and insects which may result in a change in ecosystem function; (ii) an increase in the number of bird species; and (iii) both increases and decreases in the suitable climatic area of key habitats and species, all of which are expected to impact Ireland's future biodiversity. This process identified data gaps and limitations in available information both of which could be used to inform a focused research strategy. In addition, it raises awareness of the potential implications of climate change for biodiversity in Ireland and elsewhere and demonstrates the need for biodiversity conservation plans to factor climate change into future designs.

  15. Multi-millennial-scale climate variability in Antarctica during the past seven glacial periods

    NASA Astrophysics Data System (ADS)

    Kawamura, K.

    2009-12-01

    Climate variability on 1,000- to 10,000 -year timescales and associated interhemispheric seesaw during the last glacial period have been documented in a variety of paleoclimatic records. However, the frequency, magnitude, cause and prerequisites for the older glacial periods are still uncertain. We here present a new 720,000-year ice core record from Dome Fuji, East Antarctica. The agreement between the Dome Fuji and Dome C isotopic temperature records indicates homogeneous climate variability across the East Antarctic plateau throughout the past 720 kyr. By combining the two temperature proxy records, we identified persistent multi-millennial-scale Antarctic events over the past seven glacial periods. With a fully coupled atmosphere-ocean general circulation model, it is suggested that the prerequisite for the bipolar seesaw is the combination of a cold background climate and freshwater input into the northern North Atlantic. With our identification criteria, the mean repetition period of the large Antarctic events increased from 6 kyr in the older three glacial periods to 8 kyr in the younger four glacial periods. Low frequency variations (repetition period of >10 kyr) occur in the early parts of the last four glacial periods (i.e. after Mid-Brunhes climatic shift), suggesting a role of insolation forcing on the large bipolar events in the recent glacial periods. Dome Fuji Ice Core Project members (listed in alphabetical order): Ayako Abe-Ouchi, Yutaka Ageta, Shuji Aoki, Nobuhiko Azuma, Yoshiyuki Fujii, Koji Fujita, Shuji Fujita, Kotaro Fukui, Teruo Furukawa, Atsushi Furusaki, Kumiko Goto-Azuma, Ralf Greve, Motohiro Hirabayashi, Takeo Hondoh, Akira Hori, Shinichiro Horikawa, Kazuho Horiuchi, Makoto Igarashi, Yoshinori Iizuka, Takao Kameda, Kokichi Kamiyama, Hiroshi Kanda, Kenji Kawamura, Mika Kohno, Takayuki Kuramoto, Yuki Matsushi, Morihiro Miyahara, Takayuki Miyake, Atsushi Miyamoto, Hideaki Motoyama, Yasuo Nagashima, Yoshiki Nakayama, Takakiyo Nakazawa, Fumio

  16. All Christians? Experiences of science educators in Northern Ireland

    NASA Astrophysics Data System (ADS)

    Murphy, Colette; Hickey, Ivor; Beggs, Jim

    2010-03-01

    In this paper we respond to Staver's article (this issue) on an attempt to resolve the discord between science and religion. Most specifically, we comment on Staver's downplaying of difference between Catholics and Protestants in order to focus on the religion-science question. It is our experience that to be born into one or other of these traditions in some parts of the world (especially Northern Ireland) resulted in starkly contrasting opportunities, identities and practices in becoming and being science educators. The paper starts with a short contextual background to the impact of religion on schooling and higher education in Northern Ireland. We then explore the lives and careers of three science/religious educators in Northern Ireland: Catholic (Jim) and Protestant (Ivor) males who are contemporaries and whose experience spans pre-Troubles to post-conflict and a Catholic female (Colette) who moved to Northern Ireland during the Troubles as a teenager. Finally, we discuss the situation regarding the teaching of creationism and evolution in Northern Ireland—an issue has recently generated high public interest. The Chair of the Education Committee of the Northern Ireland Assembly recently stated that "creationism is not for the RE class because I believe that it can stand scientific scrutiny and that is a debate which I am quite happy to encourage and be part of…" (News Letter 2008). It could be the case that the evolution debate is being fuelled as a deliberate attempt to undermine some of the post-conflict collaboration projects between schools and communities in Northern Ireland.

  17. Road accident facts Ireland, 2002

    DOT National Transportation Integrated Search

    2003-11-01

    This report covers all road traffic accidents reported to : the Garda Sochna, where details were forwarded to the : National Roads Authority, involving fatalities, personal : injury or material damage which occurred on public : roads in Ireland (...

  18. Simulated Impact of Glacial Runoff on CO2 Uptake in the Gulf of Alaska

    NASA Astrophysics Data System (ADS)

    Pilcher, Darren J.; Siedlecki, Samantha A.; Hermann, Albert J.; Coyle, Kenneth O.; Mathis, Jeremy T.; Evans, Wiley

    2018-01-01

    The Gulf of Alaska (GOA) receives substantial summer freshwater runoff from glacial meltwater. The alkalinity of this runoff is highly dependent on the glacial source and can modify the coastal carbon cycle. We use a regional ocean biogeochemical model to simulate CO2 uptake in the GOA under different alkalinity-loading scenarios. The GOA is identified as a current net sink of carbon, though low-alkalinity tidewater glacial runoff suppresses summer coastal carbon uptake. Our model shows that increasing the alkalinity generates an increase in annual CO2 uptake of 1.9-2.7 TgC/yr. This transition is comparable to a projected change in glacial runoff composition (i.e., from tidewater to land-terminating) due to continued climate warming. Our results demonstrate an important local carbon-climate feedback that can significantly increase coastal carbon uptake via enhanced air-sea exchange, with potential implications to the coastal ecosystems in glaciated areas around the world.

  19. Glacial morphology in the Chinese Pamir: Connections among climate, erosion, topography, lithology and exhumation

    NASA Astrophysics Data System (ADS)

    Schoenbohm, Lindsay M.; Chen, Jie; Stutz, Jamey; Sobel, Edward R.; Thiede, Rasmus C.; Kirby, Benjamin; Strecker, Manfred R.

    2014-09-01

    Modification of the landscape by glacial erosion reflects the dynamic interplay of climate through temperature, precipitation, and prevailing wind direction, and tectonics through rock uplift and exhumation rate, lithology, and range and fault geometry. We investigate these relationships in the northeast Pamir Mountains using mapping and dating of moraines and terraces to determine the glacial history. We analyze modern glacial morphology to determine glacier area, spacing, headwall relief, debris cover, and equilibrium line altitude (ELA) using the area x altitude balance ratio (AABR), toe-to-headwall altitude ratio (THAR) and toe-to-summit altitude method (TSAM) for 156 glaciers and compare this to lithologic, tectonic, and climatic data. We observe a pronounced asymmetry in glacial ELA, area, debris cover, and headwall relief that we interpret to reflect both structural and climatic control: glaciers on the downwind (eastern) side of the range are larger, more debris covered, have steeper headwalls, and tend to erode headward, truncating the smaller glaciers of the upwind, fault-controlled side of the range. We explain this by the transfer of moisture deep into the range as wind-blown or avalanched snow and by limitations imposed on glacial area on the upwind side of the range by the geometry of the Kongur extensional system (KES). The correspondence between rapid exhumation along the KES and maxima in glacier debris cover and headwall relief and minimums in all measures of ELA suggest that taller glacier headwalls develop in a response to more rapid exhumation rates. However, we find that glaciers in the Muji valley did not extend beyond the range front until at least 43 ka, in contrast to extensive glaciation since 300 ka in the south around the high peaks, a pattern which does not clearly reflect uplift rate. Instead, the difference in glacial history and the presence of large peaks (Muztagh Ata and Kongur Shan) with flanking glaciers likely reflects

  20. Age evaluation and causation of rock-slope failures along the western margin of the Antrim Lava Group (ALG), Northern Ireland, based on cosmogenic isotope (36Cl) surface exposure dating

    NASA Astrophysics Data System (ADS)

    Southall, David W.; Wilson, Peter; Dunlop, Paul; Schnabel, Christoph; Rodés, Ángel; Gulliver, Pauline; Xu, Sheng

    2017-05-01

    The temporal pattern of postglacial rock-slope failure in a glaciated upland area of Ireland (the western margin of the Antrim Lava Group) was evaluated using both 36Cl exposure dating of surface boulders on run-out debris and 14C dating of basal organic soils from depressions on the debris. The majority of the 36Cl ages ( 21-15 ka) indicate that major failures occurred during or immediately following local deglaciation ( 18-17 ka). Other ages ( 14-9 ka) suggest some later, smaller-scale failures during the Lateglacial and/or early Holocene. The 14C ages (2.36-0.15 cal ka BP) indicate the very late onset of organic accumulation and do not provide close limiting age constraints. Rock-slope failure during or immediately following local deglaciation was probably in response to some combination of glacial debuttressing, slope steepening and paraglacial stress release. Later failures may have been triggered by seismic activity associated with glacio-isostatic crustal uplift and/or permafrost degradation consequent upon climate change. The 36Cl ages support the findings of previous studies that show the deglacial - Lateglacial period in northwest Ireland and Scotland to have been one of enhanced rock-slope failure. Table S2 Concentrations of main elements (as oxides) etc.

  1. Glacial Ice Deposits in Mid-Latitudes of Mars

    NASA Image and Video Library

    2010-03-02

    NASA Mars Reconnaissance Orbiter has detected widespread deposits of glacial ice in the mid-latitudes of Mars. This map of a region known as Deuteronilus Mensae, in the northern hemisphere, shows locations of the detected ice deposits in blue.

  2. Keep your feet warm? A cryptic refugium of trees linked to a geothermal spring in an ocean of glaciers.

    PubMed

    Carcaillet, Christopher; Latil, Jean-Louis; Abou, Sébastien; Ali, Adam; Ghaleb, Bassam; Magnin, Frédéric; Roiron, Paul; Aubert, Serge

    2018-06-01

    Up to now, the most widely accepted idea of the periglacial environment is that of treeless ecosystems such as the arctic or the alpine tundra, also called the tabula rasa paradigm. However, several palaeoecological studies have recently challenged this idea, that is, treeless environments in periglacial areas where all organisms would have been exterminated near the glacier formed during the Last Glacial Maximum, notably in the Scandinavian mountains. In the Alps, the issue of glacial refugia of trees remains unanswered. Advances in glacier reconstructions show that ice domes did not cover all upper massifs, but glaciers filled valleys. Here, we used fossils of plant and malacofauna from a travertine formation located in a high mountain region to demonstrate that trees (Pinus, Betula) grew with grasses during the Lateglacial-Holocene transition, while the glacier fronts were 200-300 m lower. The geothermal travertine started to accumulate more than 14,500 years ago, but became progressively more meteogene about 11,500 years ago due to a change in groundwater circulation. With trees, land snails (gastropods) associated to woody or open habitats and aquatic mollusc were also present at the onset of the current interglacial, namely the Holocene. The geothermal spring, due to warm water and soil, probably favoured woody glacial ecosystems. This new finding of early tree growth, combined with other scattered proofs of the tree presence before 11,000 years ago in the western Alps, changes our view of the tree distribution in periglacial environments, supporting the notion of tree refugia on nunataks in an ocean of glaciers. Therefore, the tabula rasa paradigm must be revisited because it has important consequences on the global changes, including postglacial plant migrations and biogeochemical cycles. © 2018 John Wiley & Sons Ltd.

  3. Vocational Training in Ireland.

    ERIC Educational Resources Information Center

    Dooney, Roy; Dunne, Paul

    This monograph, one of a series of studies of vocational education in the countries of the European Communities, describes the vocational training system in Ireland. The study was compiled from existing statistics and descriptions, and most figures cited refer to 1984. The report is organized in eight chapters. Chapter 1 covers population,…

  4. The development of counselling psychology in Ireland

    PubMed Central

    Connolly, Allison; O'Callaghan, Dermot; O'Brien, Owen; Broderick, John; Long, Catherine; O'Grady, Ian

    2014-01-01

    This paper discusses the distinctive nature of the specialism of counselling psychology and outlines the development of the discipline in Ireland in the context of international developments and its recognition as a professional branch of applied psychology. Today, counselling psychologists are employed in varied clinical and non-clinical settings including health and mental health services (statutory, private and voluntary sector) along with education, forensic, justice, industry and private practices. Counselling psychologist is the primary professional identity of many practising psychologists in Ireland and the Psychological Society of Ireland's Division of Counselling Psychology is the main affiliation of at least 179 members. With its focus on facilitating personal and interpersonal functioning across the life span and its emphasis on the therapeutic process, the specialism continues to bridge the disciplines of psychology, counselling and psychotherapy. In this article, some of the challenges still faced by counselling psychology are explored as it navigates its way through the changing landscape of further development and evolution. PMID:26494940

  5. Glacial seismology

    NASA Astrophysics Data System (ADS)

    Aster, R. C.; Winberry, J. P.

    2017-12-01

    Seismic source and wave propagation studies contribute to understanding structure, transport, fracture mechanics, mass balance, and other processes within glaciers and surrounding environments. Glaciogenic seismic waves readily couple with the bulk Earth, and can be recorded by seismographs deployed at local to global ranges. Although the fracturing, ablating, melting, and/or highly irregular environment of active glaciers can be highly unstable and hazardous, informative seismic measurements can commonly be made at stable proximal ice or rock sites. Seismology also contributes more broadly to emerging studies of elastic and gravity wave coupling between the atmosphere, oceans, solid Earth, and cryosphere, and recent scientific and technical advances have produced glaciological/seismological collaborations across a broad range of scales and processes. This importantly includes improved insight into the responses of cryospheric systems to changing climate and other environmental conditions. Here, we review relevant fundamental physics and glaciology, and provide a broad review of the current state of glacial seismology and its rapidly evolving future directions.

  6. Glacial seismology.

    PubMed

    Aster, R C; Winberry, J P

    2017-12-01

    Seismic source and wave propagation studies contribute to understanding structure, transport, fracture mechanics, mass balance, and other processes within glaciers and surrounding environments. Glaciogenic seismic waves readily couple with the bulk Earth, and can be recorded by seismographs deployed at local to global ranges. Although the fracturing, ablating, melting, and/or highly irregular environment of active glaciers can be highly unstable and hazardous, informative seismic measurements can commonly be made at stable proximal ice or rock sites. Seismology also contributes more broadly to emerging studies of elastic and gravity wave coupling between the atmosphere, oceans, solid Earth, and cryosphere, and recent scientific and technical advances have produced glaciological/seismological collaborations across a broad range of scales and processes. This importantly includes improved insight into the responses of cryospheric systems to changing climate and other environmental conditions. Here, we review relevant fundamental physics and glaciology, and provide a broad review of the current state of glacial seismology and its rapidly evolving future directions.

  7. Feedbacks between subglacial dynamics and long-term glacial landscape evolution (Invited)

    NASA Astrophysics Data System (ADS)

    Brædstrup, C. F.; Egholm, D. L.; Ugelvig, S. V.; Christensen, A. D.; Andersen, J. L.

    2011-12-01

    Several well-known glacial landforms (such as U-shaped troughs and cirques) are associated with characteristic length scales, indicating that the viscosity of the ice and the stress gradients associated with ice flow exert first-order controls on their formation. The evolution of these glacial landforms has so far mostly been explored using phenomenological models that simply link the subglacial erosion rate to sliding or ice discharge. In order to improve our understanding of the causal links between the glacial landforms and the physics of the subglacial environment, we have performed computational experiments with a higher-order ice sheet model (Egholm et al., 2009) capable of simulating the long-term evolution of subglacial dynamics at a high spatial resolution. The orientation and magnitude of subglacial stress components depend not only on ice thickness and ice surface gradients, but also on the details of the bed topography and the regional variations in ice flow velocity. As glaciers erode their beds and modify the morphology of glaciated valleys, the subglacial dynamics therefore change with important implications for the sliding patterns and the continued erosion rates. We focus this presentation on feedbacks between the evolving bed topography and the subglacial erosion patterns. We have performed our experiments with different sliding and erosion laws, including highly non-linear rules representing coulomb-type slip at the bed (Schoof, 2010) and a quarrying model associated to the level of cavitation (Iverson, 2012). The highly non-linear computational experiments are made possible by new and very efficient GPU-accelerated multigrid algorithms. The computational experiments show that higher-order stress effects associated with local changes to the bed gradient provide important stabilizing effects for example in overdeepenings and near topographic steps. The experiments also show how a narrow and meandering pre-glacial valley represents a much more

  8. Feedbacks between subglacial dynamics and long-term glacial landscape evolution (Invited)

    NASA Astrophysics Data System (ADS)

    Brædstrup, C. F.; Egholm, D. L.; Ugelvig, S. V.; Christensen, A. D.; Andersen, J. L.

    2013-12-01

    Several well-known glacial landforms (such as U-shaped troughs and cirques) are associated with characteristic length scales, indicating that the viscosity of the ice and the stress gradients associated with ice flow exert first-order controls on their formation. The evolution of these glacial landforms has so far mostly been explored using phenomenological models that simply link the subglacial erosion rate to sliding or ice discharge. In order to improve our understanding of the causal links between the glacial landforms and the physics of the subglacial environment, we have performed computational experiments with a higher-order ice sheet model (Egholm et al., 2009) capable of simulating the long-term evolution of subglacial dynamics at a high spatial resolution. The orientation and magnitude of subglacial stress components depend not only on ice thickness and ice surface gradients, but also on the details of the bed topography and the regional variations in ice flow velocity. As glaciers erode their beds and modify the morphology of glaciated valleys, the subglacial dynamics therefore change with important implications for the sliding patterns and the continued erosion rates. We focus this presentation on feedbacks between the evolving bed topography and the subglacial erosion patterns. We have performed our experiments with different sliding and erosion laws, including highly non-linear rules representing coulomb-type slip at the bed (Schoof, 2010) and a quarrying model associated to the level of cavitation (Iverson, 2012). The highly non-linear computational experiments are made possible by new and very efficient GPU-accelerated multigrid algorithms. The computational experiments show that higher-order stress effects associated with local changes to the bed gradient provide important stabilizing effects for example in overdeepenings and near topographic steps. The experiments also show how a narrow and meandering pre-glacial valley represents a much more

  9. Paleolimnology of Lake Ontario: AN Assessment of Glacial Meltwater Influx

    NASA Astrophysics Data System (ADS)

    Hladyniuk, R.; Longstaffe, F. J.

    2010-12-01

    The timing and extent of glacial meltwater outbursts from the Laurentide Ice Sheet (LIS) may provide insight into their potential role in initiating and/or sustaining the Younger Dryas (YD) cooling event. It has been previously proposed that meltwater from the LIS suppressed thermohaline circulation in the Atlantic Ocean, leading to an abrupt change in climate (Broecker et al. 1989). Several pathways for transport of glacial meltwater to the Atlantic Ocean have been suggested in the past, including eastern flow through the St. Lawrence River system and discharge into the Arctic Ocean via a northwestern outlet (Murton et al. 2010). Glacial meltwater contributions to Lake Ontario and its ancient equivalents during the last ~14,000 cal BP have been evaluated using the oxygen-isotope compositions of ostracode shells from three sediment cores in Lake Ontario. Glacial Lake Iroquois (~12,500 cal BP) δ18O values as low as -18‰ suggest significant contribution of glacial meltwater runoff from the LIS. This glacial sediment is characterized by occasional grains of sand and gravel. These ice-rafted particles indicate how far icebergs floated and suggest close proximity to the LIS. Early Lake Ontario sediment (~12,000 cal BP) exhibits thicker laminations, suggestive of increased winter ice cover and perhaps a colder climate, and is characterized by slightly lower δ18O values (-19.5‰). The end of glacial-dominated sedimentation at ~11,800 cal BP is demarcated by a significant increase in lakewater δ18O values (-12.0‰), reflecting mixing between regional precipitation in the watershed and upstream inflow into Lake Ontario. At ~10,800 cal BP, the δ18O value of Lake Ontario decreased to ~-15‰. This change reflects the main Algonquin highstand in Lake Huron, which flooded into Lake Ontario from both the Fenelon Falls and Port Huron outlets at this time. Shortly thereafter, the opening of the North Bay outlet and isostatic rebound at the Port Huron outlet limited

  10. Glacial evolution of the Ampato Volcanic Complex (Peru)

    NASA Astrophysics Data System (ADS)

    Alcalá, J.; Palacios, D.; Zamorano, J. J.; Vázquez, L.

    2009-04-01

    Ice masses on the Western range of the Central Andes are a main source of water resources and act as a geoindicator of variations in the climate of the tropics (Mark, 2008). The study of their evolution is of particular interest since they are situated in the transition zone between the tropical and mid-latitude circulation areas of the atmosphere (Zech et al., 2007). The function of this transition area is currently under debate, and understanding it is essential for the development of global climate models (Kull et al, 2008; Mark, 2008). However our understanding of the evolution of glaciers and their paleoclimatic factors for this sector of the Central Andes is still at a very basic level. This paper presents initial results of a study on the glacial evolution of the Ampato volcanic complex (15°24´- 15° 51´ S, 71° 51´ - 73° W; 6288 m a.s.l.) located in the Western Range of the Central Andes in Southern Peru, 70 km NW of the city of Arequipa. The main objectives are to identify the number of glacial phases the complex has undergone using geomorphological criteria to define a time frame for each phase, based on cosmogenic 36Cl dating of a sequence of moraine deposits; and to estimate the glacier Equilibrium Line Altitude (ELA) of each phase. The Ampato volcanic complex is formed by 3 great andesitic stratovolcanoes, the Nevados HualcaHualca-Sabancaya-Ampato, which started forming between the late Miocene and early Quaternary (Bulmer et al., 1999), aligned N-S and with summits covered with glaciers. The Sabancaya volcano is fully active, with its latest eruption occurring in 2001. Glacial landforms were identified and mapped using photointerpretation of vertical aerial photographs from 1955 (1:35,000 scale, National Geographic Institute of Peru), oblique photographs from 1943 (Aerophotographical Service of Peru), and a geo-referenced high-resolution Mrsid satellite image from 2000 (NASA). This cartography was corrected and improved through fieldwork. It was

  11. Evolution of supra-glacial lakes across the Greenland Ice Sheet

    NASA Astrophysics Data System (ADS)

    Sundal, A. V.; Shepherd, A.; Nienow, P.; Hanna, E.; Palmer, S.; Huybrechts, P.

    2009-04-01

    We have used 268 cloud-free Moderate-resolution Imaging Spectroradiometer (MODIS) images spanning the 2003 and 2005-2007 melt seasons to study the seasonal evolution of supra-glacial lakes in three different regions of the Greenland Ice Sheet. Lake area estimates were obtained by developing an automated classification method for their identification based on 250 m resolution MODIS surface reflectance observations. Widespread supra-glacial lake formation and drainage is observed across the ice sheet, with a 2-3 weeks delay in the evolution of total supra-glacial lake area in the northern areas compared to the south-west. The onset of lake growth varies by up to one month inter-annually, and lakes form and drain at progressively higher altitudes during the melt season. A correlation was found between the annual peak in total lake area and modelled annual runoff across all study areas. Our results indicate that, in a future warmer climate (Meehl et al., 2007), Greenland supra-glacial lakes can be expected to form at higher altitudes and over a longer time period than is presently the case, expanding the area and time period over which connections between the ice sheet surface and base may be established (Das et al., 2008) with potential consequences for ice sheet discharge (Zwally et al., 2002). Das, S., Joughin, M., Behn, M., Howat, I., King, M., Lizarralde, D., & Bhatia, M. (2008). Fracture propagation to the base of the Greenland Ice Sheet during supra-glacial lake drainage. Science, 5877, 778-781. Meehl, G.A., Stocker, T.F., Collins W.D., Friedlingstein, P., Gaye, A.T., Gregory, J.M., Kitoh, A., Knutti, R., Murphy, J.M., Noda, A., Raper, S.C.B., Watterson, I.G., Weaver, A.J. & Zhao, Z.C. (2007). Global Climate Projections. In: Climate Change 2007: The Physical Science Basis. Contribution of Working Group I to the Fourth Assessment Report of the Intergovernmental Panel on Climate Change [Solomon, S., D. Qin, M. Manning, Z. Chen, M. Marquis, K.B. Averyt, M. Tignor

  12. Insolation-driven 100 kyr glacial cycles and millennial climate change

    NASA Astrophysics Data System (ADS)

    Abe-Ouchi, A.; Saito, F.; Kawamura, K.; Raymo, M. E.; Okuno, J.; Takahashi, K.; Blatter, H.

    2013-12-01

    The waxing and waning of Northern Hemisphere ice sheets over the past one million years is dominated by an approximately 100-kyr periodicity and a sawtooth pattern (gradual growth and fast termination). Milankovitch theory proposes that summer insolation at high northern latitudes drives the glacial cycles, and statistical tests demonstrated that the glacial cycles are indeed linked to eccentricity, obliquity and precession cycles. However, insolation alone cannot explain the strong 100 kyr cycle which presumably arises through internal climatic feedbacks. Prior work with conceptual models, for example, showed that glacial terminations are associated with the build-up of Northern Hemisphere 'excess ice', but the physical mechanisms of 100-kyr cycle at work remain unclear. Here, using comprehensive climate and ice sheet models, we show that the ~100-kyr periodicity is explained by insolation and internal feedback amongst the climate, ice sheet and lithosphere/asthenosphere system (reference). We found that equilibrium states of ice sheets exhibit hysteresis responses to summer insolation, and that the shape and position of the hysteresis loop play a key role in determining the periodicities of glacial cycles. The hysteresis loop of the North American ice sheet is such that, after its inception, the ice sheet mass balance remains mostly positive or neutral through several precession cycles whose amplitude decreases towards an eccentricity minimum. The larger the ice sheet grows and extends towards lower latitudes, the smaller is the insolation required to turn the mass balance to negative. Therefore, once the large ice sheet is established, only a moderate increase in insolation can trigger a negative mass balance, leading to a complete retreat within several thousand years, due to the delayed isostatic rebound. The effect of ocean circulation and millennial scale climate change are not playing the dominant role for determing the 100kyr cycle, but are effective for

  13. Climatic Instability and Regional Glacial Advances in the Late Ediacaran

    NASA Astrophysics Data System (ADS)

    Hannah, J. L.; Stein, H. J.; Marolf, N.; Bingen, B.

    2014-12-01

    The Ediacaran Period closed out the environmentally raucous Neoproterozoic Era with the last of multiple glacial events and the first ephemeral glimmer of multicellular life. As such, evolution of Earth's biosphere and the marine environments that nurtured this nascent biota are of particular interest. Because the Ediacaran biota appear in the stratigraphic record just above tillites in many localities, inferences are naturally drawn to link glaciation to bioevolution. Here we review known controls on the timing and extent of the late Ediacaran Gaskier and Varanger glacial events, bolstered by new constraints on the Moelv tillite of South Norway. The elusive mid-Ediacaran glacial strata are poorly dated, patchy in distribution, and relatively limited in thickness. The type Gaskier glaciogenic units in Newfoundland are 582 to 584 Ma, based on U-Pb zircon ages from intercalated ash beds [1]. The Varanger glaciogenic deposits in northern Norway, in contrast, remain only roughly constrained to ca. 630 to 560 Ma. Post-Gaskier negative carbon isotope excursions (CIEs) have been reported from multiple localities in both China and SW United States, suggesting climatic instability in the late Ediacaran. Although most localities lack solid geochronology, paleontologic constraints place the Hongtiegou glacial diamictite and accompanying CIE in the Chaidam Basin, NW China, in the latest Ediacaran, ca. 555 Ma [2]. We previously suggested that the Moelv tillite in south Norway was roughly equivalent to the Gaskier, based on an imprecise Re-Os age of ~560 Ma [3] for the underlying Biri shale. Reanalysis of these data shows that the upper part of the shale section was disturbed by a redox front during the Caledonian orogeny. The undisturbed lower part of the section yields a more precise Model 1 isochron age of 559.5 ± 6.2 Ma, clearly post-dating the Gaskier event well outside analytical uncertainty. These new results bolster arguments that the Gaskier glaciation was not a global

  14. Inherent characteristics of sawtooth cycles can explain different glacial periodicities

    NASA Astrophysics Data System (ADS)

    Omta, Anne Willem; Kooi, Bob W.; van Voorn, George A. K.; Rickaby, Rosalind E. M.; Follows, Michael J.

    2016-01-01

    At the Mid-Pleistocene Transition about 1 Ma, the dominant periodicity of the glacial-interglacial cycles shifted from 40 to 100 kyr. Here, we use a previously developed mathematical model to investigate the possible dynamical origin of these different periodicities. The model has two variables, one of which exhibits sawtooth oscillations, resembling the glacial-interglacial cycles, whereas the other variable exhibits spikes at the rapid transitions. When applying a sinusoidal forcing with a fixed period, there emerges a rich variety of cycles with different periodicities, each being a multiple of the forcing period. Furthermore, the dominant periodicity of the system can change, while the forcing periodicity remains fixed, due to either random variations or different frequency components of the orbital forcing. Two key relationships stand out as predictions to be tested against observations: (1) the amplitude and the periodicity of the cycles are approximately linearly proportional to each other, a relationship that is also found in the δ ^{18}O temperature proxy. (2) The magnitude of the spikes increases with increasing periodicity and amplitude of the sawtooth. This prediction could be used to identify one or more currently hidden spiking variables driving the glacial-interglacial transitions. Essentially, the quest would be for any proxy record, concurrent with a dynamical model prediction, that exhibits deglacial spikes which increase at times when the amplitude/periodicity of the glacial cycles increases. In the specific context of our calcifier-alkalinity mechanism, the records of interest would be calcifier productivity and calcite accumulation. We believe that such a falsifiable hypothesis should provide a strong motivation for the collection of further records.

  15. ICT Policy and Implementation in Education: Cases in Canada, Northern Ireland and Ireland

    ERIC Educational Resources Information Center

    Austin, Roger; Hunter, Bill

    2013-01-01

    Countries with similar levels of economic development often implement different education ICT policies. Much of the existing research attributes such differences to economic and political factors. In this paper, we examine the development of ICT policy and implementation in the two parts of Ireland and in two Canadian provinces and find that…

  16. Telling Tales--Cruelty and Abuse in Schooling in Ireland

    ERIC Educational Resources Information Center

    Jeffers, Gerry

    2016-01-01

    The report of the Commission to Inquire into Child Abuse (Government of Ireland, 2009)--the Ryan Report--shocked Ireland and the wider world with its chilling descriptions of abuse that was systemic, pervasive, chronic, excessive, arbitrary and endemic. Subsequent debate has, rightly, centred on the "religious" arena, highlighting the…

  17. Role of Marine Biology in Glacial-Interglacial CO2 Cycles

    NASA Astrophysics Data System (ADS)

    Kohfeld, Karen E.; Le Quéré, Corinne; Harrison, Sandy P.; Anderson, Robert F.

    2005-04-01

    It has been hypothesized that changes in the marine biological pump caused a major portion of the glacial reduction of atmospheric carbon dioxide by 80 to 100 parts per million through increased iron fertilization of marine plankton, increased ocean nutrient content or utilization, or shifts in dominant plankton types. We analyze sedimentary records of marine productivity at the peak and the middle of the last glacial cycle and show that neither changes in nutrient utilization in the Southern Ocean nor shifts in plankton dominance explain the CO2 drawdown. Iron fertilization and associated mechanisms can be responsible for no more than half the observed drawdown.

  18. Vestiges of Glacial Action in Ostrava: Their Significance for and Application in Geotourism

    NASA Astrophysics Data System (ADS)

    Duraj, Miloš; Niemiec, Dominik; Cheng, Xianfeng; Koleňák, Petr

    2017-12-01

    The territory of Northern Moravia and Silesia is outstanding from the geological point of view. The abundance of different mineral resources has largely contributed to the intense development of the territory, particularly in the 19th century. Mineral resources were discovered already in the pre-historic period, when pre-historic man found coal at the coal seam exposures in Ostrava-Landek. They also used some raw materials that had been transported there by glacial action of the last Saale glaciation. Flint fragments and other travelled material may be frequently found in many localities to date. Large pieces that are called glacial boulders have been removed and exhibited for more than a century in many towns of the region. These vestiges of glacial action represent one of the many stages the Earth has passed through its history. At present, such findings mainly have an aesthetic function. Particularly interesting specimens have been protected as national monuments. The geomorphology of Ostrava has been responsible for the findings of the largest glacial boulders within the Czech Republic. Many of the formations are fascinating specimens that enrich the list of numerous geomontane sights in the City of Ostrava.

  19. Glacial geology of the Shingobee River headwaters area, north-central Minnesota

    USGS Publications Warehouse

    Melchior, Robert C.

    2014-01-01

    During middle and late Wisconsin time in the Shingobee River headwaters area, the Laurentide Wadena lobe, Hewitt and Itasca phases, produced terminal and ground moraine along with a variety of associated glacial features. The stratigraphic record is accessible and provides details of depositional mode as well as principal glacial events during the advance and retreat of middle and late Wisconsin ice tongues. Geomorphic features such as tunnel valleys, stream terraces, and postglacial stream cuts formed by erosional events persist to the present day. Middle Wisconsin Hewitt phase deposits are the oldest and include drumlins, ground moraine, boulder pavements, and outwash. Together, these deposits suggest a wet-based, periodically surging glacier in a subpolar thermal state. Regional permafrost and deposition from retreating ice are inferred between the end of the Hewitt phase and the advance of late Wisconsin Itasca phase ice. Itasca phase glaciation occurred as a contemporaneous pair of adjacent ice tongues whose contrasting moraine styles suggest independent flow modes. The western (Shingobee) portion of the Itasca moraine contains composite ridges, permafrost phenomena, hill-hole pairs, and debris flows. By contrast, eastern (Onigum) moraine deposits generally lack glaciotectonic features and consist almost exclusively of mud and debris flows. Near the end of the Itasca phase, large-scale hill-hole pairs developed in the Shingobee division, and debris flows from the Onigum division blocked the preexisting Shingobee tunnel valley to form glacial lake Willobee. Postglacial streams formed deep valleys as glacial lake Willobee catastrophically drained. Dates based on temperature trends in Greenland ice cores are proposed for prominent glacial events in the Shingobee area. This report proposes that Hewitt phase glaciation occurred between 27.2 and 23.6 kiloannum and Itasca phase glaciation between 22.8 and 14.7 kiloannum. Des Moines lobe (Younger Dryas) glaciation

  20. The Effects of Salt Water on Mechanical Properties of Glacial Ice

    NASA Astrophysics Data System (ADS)

    Holt, R. A.; McCarthy, C.

    2017-12-01

    An improved understanding of the mechanical properties of glacial ice, including factors that may change them, is essential for understanding vulnerability of ice sheets to climate change. It is understood that the temperature of intruding subglacial seawater affects the melting of glacial ice and therefore destabilizes ice shelves, but we hypothesize that seawater bathing the bottom of the glacier may also influence mechanical properties such as friction and elastic modulus. We undertook experiments to determine how the presence of saline solution at grain boundaries of ice might lead to weaker behavior. We created an ice sample by finely grinding and sieving seed ice, pressing it into a rectangular mold, and flooding with a 3.5wt% saline solution. We then quickly brought it to subsolidus (-22°) to completely freeze. The bulk composition of the sample was determined by refractive index to be 0.28wt%. Microstructural characterization of the sample indicates that, above the solidus, the melt phase was located at grain triple junctions and along grain boundaries. To test the frictional behavior of ice with saline sliding against rock, we used a cryo-biaxial apparatus designed to simulate the basal sliding of glacial ice. The experiments were run in the double direct configuration at 100 KPa normal stress and at T=-5°. The results demonstrate that ice containing a liquid saline solution has lower steady state friction than pure ice at the same conditions, and therefore can slip at a faster velocity. In addition to the bi-axial experiment we determined the elastic properties using an ultrasonic velocity testing system. P waves velocities through the saline ice sample were consistent with published values (Spencer et al., 1968, JGR). We also used both measured and estimated values to calculate the Young's modulus. We found that ice containing salt water has a lower Young's modulus than that of pure ice. Salt water significantly changes the mechanical properties of