Sample records for pelagic crinoid fossil

  1. Pelagic crinoids (Roveacrinida, Crinoidea) discovered in the Neogene of Poland.

    PubMed

    Gorzelak, Przemysław; Salamon, Mariusz A; Ferré, Bruno

    2011-10-01

    Until recently, it has been assumed that pelagic crinoids, the roveacrinids (Roveacrinida, Crinoidea), became extinct during the Cretaceous-Paleogene boundary event. Recent finds of well-preserved roveacrinidal remains (brachials and radials) in the Danian (Early Paleogene) of Poland showed that they survived into the earliest Cenozoic. This group was thus characterized as a "dead clade walking". Here, we present fossil evidence that these pelagic crinoids survived in Poland until at least the Middle Miocene (Badenian, ca. 14 Myr ago)-more than 50 Myr after their supposed extinction. These Miocene roveacrinids constitute the first documented evidence of Roveacrinida in strata of Neogene age, thus prolonging the stratigraphic range of pelagic crinoids. This find characterizes the order as a "Lazarus taxon" rather than a "dead clade walking" group.

  2. Fossil Crinoids

    NASA Astrophysics Data System (ADS)

    Hess, Hans; Ausich, William I.; Brett, Carlton E.; Simms, Michael J.

    1999-10-01

    Crinoids have graced the oceans for more than 500 million years. Among the most attractive fossils, crinoids had a key role in the ecology of marine communities through much of the fossil record, and their remains are prominent rock forming constituents of many limestones. This is the first comprehensive volume to bring together their form and function, classification, evolutionary history, occurrence, preservation and ecology. The main part of the book is devoted to assemblages of intact fossil crinoids, which are described in their geological setting in twenty-three chapters ranging from the Ordovician to the Tertiary. The final chapter deals with living sea lilies and feather stars. The volume is exquisitely illustrated with abundant photographs and line drawings of crinoids from sites around the world. This authoritative account recreates a fascinating picture of fossil crinoids for paleontologists, geologists, evolutionary and marine biologists, ecologists and amateur fossil collectors.

  3. Fossil Crinoids

    NASA Astrophysics Data System (ADS)

    Hess, Hans; Ausich, William I.; Brett, Carlton E.; Simms, Michael J.

    2003-01-01

    Crinoids have graced the oceans for more than 500 million years. Among the most attractive fossils, crinoids had a key role in the ecology of marine communities through much of the fossil record, and their remains are prominent rock forming constituents of many limestones. This is the first comprehensive volume to bring together their form and function, classification, evolutionary history, occurrence, preservation and ecology. The main part of the book is devoted to assemblages of intact fossil crinoids, which are described in their geological setting in twenty-three chapters ranging from the Ordovician to the Tertiary. The final chapter deals with living sea lilies and feather stars. The volume is exquisitely illustrated with abundant photographs and line drawings of crinoids from sites around the world. This authoritative account recreates a fascinating picture of fossil crinoids for paleontologists, geologists, evolutionary and marine biologists, ecologists and amateur fossil collectors.

  4. Computational Fluid Dynamics Analysis of the Fossil Crinoid Encrinus liliiformis (Echinodermata: Crinoidea)

    PubMed Central

    Dynowski, Janina F.; Nebelsick, James H.; Klein, Adrian; Roth-Nebelsick, Anita

    2016-01-01

    Crinoids, members of the phylum Echinodermata, are passive suspension feeders and catch plankton without producing an active feeding current. Today, the stalked forms are known only from deep water habitats, where flow conditions are rather constant and feeding velocities relatively low. For feeding, they form a characteristic parabolic filtration fan with their arms recurved backwards into the current. The fossil record, in contrast, provides a large number of stalked crinoids that lived in shallow water settings, with more rapidly changing flow velocities and directions compared to the deep sea habitat of extant crinoids. In addition, some of the fossil representatives were possibly not as flexible as today’s crinoids and for those forms alternative feeding positions were assumed. One of these fossil crinoids is Encrinus liliiformis, which lived during the middle Triassic Muschelkalk in Central Europe. The presented project investigates different feeding postures using Computational Fluid Dynamics to analyze flow patterns forming around the crown of E. liliiformis, including experimental validation by Particle Image Velocimetry. The study comprises the analysis of different flow directions, velocities, as well as crown orientations. Results show that inflow from lateral and oral leads to direct transport of plankton particles into the crown and onto the oral surface. With current coming from the “rear” (aboral) side of the crinoid, the conical opening of the crown produces a backward oriented flow in its wake that transports particles into the crown. The results suggest that a conical feeding position may have been less dependent on stable flow conditions compared to the parabolic filtration fan. It is thus assumed that the conical feeding posture of E. liliiformis was suitable for feeding under dynamically changing flow conditions typical for the shallow marine setting of the Upper Muschelkalk. PMID:27243221

  5. Predator-induced macroevolutionary trends in Mesozoic crinoids

    PubMed Central

    Gorzelak, Przemysław; Salamon, Mariusz A.; Baumiller, Tomasz K.

    2012-01-01

    Sea urchins are a major component of recent marine communities where they exert a key role as grazers and benthic predators. However, their impact on past marine organisms, such as crinoids, is hard to infer in the fossil record. Analysis of bite mark frequencies on crinoid columnals and comprehensive genus-level diversity data provide unique insights into the importance of sea urchin predation through geologic time. These data show that over the Mesozoic, predation intensity on crinoids, as measured by bite mark frequencies on columnals, changed in step with diversity of sea urchins. Moreover, Mesozoic diversity changes in the predatory sea urchins show a positive correlation with diversity of motile crinoids and a negative correlation with diversity of sessile crinoids, consistent with a crinoid motility representing an effective escape strategy. We contend that the Mesozoic diversity history of crinoids likely represents a macroevolutionary response to changes in sea urchin predation pressure and that it may have set the stage for the recent pattern of crinoid diversity in which motile forms greatly predominate and sessile forms are restricted to deep-water refugia. PMID:22509040

  6. Troglomorphism in the middle Triassic crinoids from Poland.

    PubMed

    Brom, Krzysztof R; Brachaniec, Tomasz; Salamon, Mariusz A

    2015-10-01

    In this paper, we document the Middle Triassic marine fauna recovered from the fissure/cave system of Stare Gliny (southern Poland) developed in the Devonian host dolomite. The fossils are mostly represented by in situ preserved and small-sized holdfasts of crinoids (Crinoidea) that are attached to the cave walls. Other fossils found in the cave infills include articulated brittle stars and brachiopods. Our findings constitute the oldest Mesozoic evidence for troglophile crinoids. We suggest that troglomorphism in these echinoderms was likely related to protection against predation, which underscores the magnitude of anti-predatory adaptations to increased predation pressure that occurred during the Early Mesozoic Marine Revolution.

  7. Persistent and widespread occurrence of bioactive quinone pigments during post-Paleozoic crinoid diversification

    PubMed Central

    Wolkenstein, Klaus

    2015-01-01

    Secondary metabolites often play an important role in the adaptation of organisms to their environment. However, little is known about the secondary metabolites of ancient organisms and their evolutionary history. Chemical analysis of exceptionally well-preserved colored fossil crinoids and modern crinoids from the deep sea suggests that bioactive polycyclic quinones related to hypericin were, and still are, globally widespread in post-Paleozoic crinoids. The discovery of hypericinoid pigments both in fossil and in present-day representatives of the order Isocrinida indicates that the pigments remained almost unchanged since the Mesozoic, also suggesting that the original color of hypericinoid-containing ancient crinoids may have been analogous to that of their modern relatives. The persistent and widespread occurrence, spatially as well as taxonomically, of hypericinoid pigments in various orders during the adaptive radiation of post-Paleozoic crinoids suggests a general functional importance of the pigments, contributing to the evolutionary success of the Crinoidea. PMID:25730856

  8. Discovery of dense aggregations of stalked crinoids in Izu-Ogasawara trench, Japan.

    PubMed

    Oji, Tatsuo; Ogawa, Yujiro; Hunter, Aaron W; Kitazawa, Kota

    2009-06-01

    Stalked crinoids are recognized as living fossils that typically inhabit modern deep-water environments exceeding 100 m. Previous records of stalked crinoids from hadal depths (exceeding 6000 m) are extremely rare, and no in-situ information has been available. We show here that stalked crinoids live densely on rocky substrates at depths over 9000 m in the Izu-Ogasawara Trench off the eastern coast of Japan, evidenced by underwater photos and videos taken by a remotely operated vehicle. This is the deepest in-situ observation of stalked crinoids and demonstrates that crinoid meadows can exist at hadal depths close to the deepest ocean floor, in a fashion quite similar to populations observed in shallower depths.

  9. Reassessing the improbability of a muscular crinoid stem

    NASA Astrophysics Data System (ADS)

    Gorzelak, Przemysław; Głuchowski, Edward; Salamon, Mariusz A.

    2014-08-01

    Muscular articulations in modern stalked crinoids are only present in the arms. Although it has been suggested that certain coiled-stemmed fossil taxa may have been functionally adapted to utilize muscles, evidence supporting this interpretation is lacking. Here, we use cathodoluminescence and SEM to reveal the skeletal microstructure of the enigmatic coiled-stemmed taxon Ammonicrinus (Flexibilia). Based on the well-established link between skeletal microstructure and the nature of infilling soft tissues in modern echinoderms, we reconstructed the palaeoanatomy of the Middle Devonian ammonicrinids. We show that their median columnals with elongated lateral columnal enclosure extensions (LCEE) have stereom microstructure unexpectedly resembling that in the crinoid muscular arm plates. In particular, large ligamentary facets, that are present on each side of a transverse ridge, are mainly comprised of fine galleried stereom that is indicative of the mutable collagenous tissues. In contrast, fine labyrinthic stereom, commonly associated with muscles, is situated in the periphery on each side of the surface of elongated LCEE. Our findings thus strongly suggest that the muscles may have also been present in the stem of ammonicrinids. These results reassess the previous hypotheses about evolution of muscles in crinoids and provide new insights into the mode of life of Ammonicrinus.

  10. Reassessing the improbability of a muscular crinoid stem

    PubMed Central

    Gorzelak, Przemysław; Głuchowski, Edward; Salamon, Mariusz A.

    2014-01-01

    Muscular articulations in modern stalked crinoids are only present in the arms. Although it has been suggested that certain coiled-stemmed fossil taxa may have been functionally adapted to utilize muscles, evidence supporting this interpretation is lacking. Here, we use cathodoluminescence and SEM to reveal the skeletal microstructure of the enigmatic coiled-stemmed taxon Ammonicrinus (Flexibilia). Based on the well-established link between skeletal microstructure and the nature of infilling soft tissues in modern echinoderms, we reconstructed the palaeoanatomy of the Middle Devonian ammonicrinids. We show that their median columnals with elongated lateral columnal enclosure extensions (LCEE) have stereom microstructure unexpectedly resembling that in the crinoid muscular arm plates. In particular, large ligamentary facets, that are present on each side of a transverse ridge, are mainly comprised of fine galleried stereom that is indicative of the mutable collagenous tissues. In contrast, fine labyrinthic stereom, commonly associated with muscles, is situated in the periphery on each side of the surface of elongated LCEE. Our findings thus strongly suggest that the muscles may have also been present in the stem of ammonicrinids. These results reassess the previous hypotheses about evolution of muscles in crinoids and provide new insights into the mode of life of Ammonicrinus. PMID:25116414

  11. A camerate-rich late carboniferous (Moscovian) crinoid fauna from volcanic conglomerate, Xinjiang, People's Republic of China

    USGS Publications Warehouse

    Lane, N.G.; Waters, J.A.; Maples, C.G.; Marcus, S.A.; Liao, Z.-T.

    1996-01-01

    A low-diversity camerate-rich crinoid fauna from the Qijiagou Formation, Taoshigo Valley near Turpan, Xinjiang-Uygar Autonomous Region, China was collected during field work in May, 1993. The crinoid fauna is dominated by species of Platycrinites. Other camerate crinoids include a species in the Paragaricocrinidae, Actinocrinites, a hexacrinitid, and an acrocrinoid. The only other non-North American occurrence of this latter family is Springeracrocrinus from the Moscovian of Russia. In addition to the camerates, there are several advanced cladid inadunates more typical of Upper Carboniferous crinoid faunas, including an erisocrinoid (possibly Sinocrinus), Graphiocrinus, ?Cromyocrinus, and an agassizocrinoid (Petschoracrinus) represented by partly fused infrabasal cones. A single radial plate with angustary facet may represent a cyathocrinoid, There also is a catillocrinoid, assigned here to Paracatillocrinus. The fauna, which resembles Moscovian crinoids described from Russia, is preserved in graded volcanic conglomeratic debris flows that overlie a carbonate mound and contain clasts up to 3 m in dimension. The crinoids are fragmentary, with many calyces seemingly torn into two or three pieces and dumped in with the pyroclastic debris. Camerates are represented by large thecal scraps consisting of numerous plates, or by large individual plates or circlets. Other fossils include rare solitary rugose corals, tabular bryozoans, Neospirifer, and other fragmentary brachiopods. We suspect that the crinoids may have been swept off of a nearby carbonate mound and deposited as debris-flow bedload.

  12. The oldest iocrinid crinoids from the Early/Middle Ordovician of China: Possible paleogeographic implications

    NASA Astrophysics Data System (ADS)

    Lin, Jih-Pai; Ausich, William I.; Balinski, Andrzej; Bergström, Stig M.; Sun, Yuanlin

    2018-01-01

    This study reports new crinoid material, which is identified as Muicrinus dawanensis new genus and new species from South China, representing the oldest known iocrinid in the world. These new fossils, which are characterized by bearing simple primibrachials and a helically coiled column, are from the lower-middle part of the Dawan Formation, which ranges from the upper Floian (Lower Ordovician) to the lower Dapingian (Middle Ordovician) (∼470 Ma). A total of 11 related taxa with 80 characters were selected to conduct phylogenetic analyses. Our results indicated that South China specimens are closely related to ones that are endemic to Laurentia. Synapomorphies shared between species in those two cratons include elongate supraradial plates and isotomous arm branching. Furthermore, the phylogenetic closeness of endemic taxa that only occurred in two cratons leads us to suggest a geographic connection between those regions during deep time. Combining biogeographic evidence reported from Cambrian studies, this study supports the "missing-link" configuration of Rodinia during the breakup phase. Based on the crinoid global biodiversity pattern updated here, the potential for discovery of new Paleozoic crinoid faunas in China is very high.

  13. Oral Region Homologies in Paleozoic Crinoids and Other Plesiomorphic Pentaradial Echinoderms

    PubMed Central

    Kammer, Thomas W.; Sumrall, Colin D.; Zamora, Samuel; Ausich, William I.; Deline, Bradley

    2013-01-01

    The phylogenetic relationships between major groups of plesiomorphic pentaradial echinoderms, the Paleozoic crinoids, blastozoans, and edrioasteroids, are poorly understood because of a lack of widely recognized homologies. Here, we present newly recognized oral region homologies, based on the Universal Elemental Homology model for skeletal plates, in a wide range of fossil taxa. The oral region of echinoderms is mainly composed of the axial, or ambulacral, skeleton, which apparently evolved more slowly than the extraxial skeleton that forms the majority of the body. Recent phylogenetic hypotheses have focused on characters of the extraxial skeleton, which may have evolved too rapidly to preserve obvious homologies across all these groups. The axial skeleton conserved homologous suites of characters shared between various edrioasteroids and specific blastozoans, and between other blastozoans and crinoids. Although individual plates can be inferred as homologous, no directly overlapping suites of characters are shared between edrioasteroids and crinoids. Six different systems of mouth (peristome) plate organization (Peristomial Border Systems) are defined. These include four different systems based on the arrangement of the interradially-positioned oral plates and their peristomial cover plates, where PBS A1 occurs only in plesiomorphic edrioasteroids, PBS A2 occurs in plesiomorphic edrioasteroids and blastozoans, and PBS A3 and PBS A4 occur in blastozoans and crinoids. The other two systems have radially-positioned uniserial oral frame plates in construction of the mouth frame. PBS B1 has both orals and uniserial oral frame plates and occurs in edrioasterid and possibly edrioblastoid edrioasteroids, whereas PBS B2 has exclusively uniserial oral frame plates and is found in isorophid edrioasteroids and imbricate and gogiid blastozoans. These different types of mouth frame construction offer potential synapomorphies to aid in parsimony-based phylogenetics for

  14. Intragranular Recrystallization and Lattice Reorientation of Calcite Grains in Experimentally Deformed Crinoids and Trilobites

    NASA Astrophysics Data System (ADS)

    Kim, N.; Takahashi, M.; Shigematsu, N.; Ree, J. H.; Jung, H.

    2017-12-01

    Intragranular recrystallization, including subgrain-rotation-recrystallization (SGR) and nucleation (and growth) of new grains along boundaries of deformation twins and bands, is an important process leading to grain-size reduction and causing rheological change depending on deformation condition. Despite of its importance, the detailed processes of intragranular recrystallization are still somewhat unclear. We deformed a limestone using triaxial testing machine at AIST of Japan at temperature of 500 700 °, strain rate of 10-4 10-5 s-1, confining pressure of 200 MPa and strain of up to 30%, to explore intragranular recrystallization processes of calcite. The limestone contains two abundant fossils, crinoid and trilobite. The crinoids are mono- or poly-crystalline. We focus on the monocrystalline crinoids with a coarser grain size ( 700 μm). The trilobites are polycrystalline and much finer-grained ( 7 μm) with initially a strong c-axis preferred orientation. At a lower temperature condition, subgrains develop both in twin and host domains of crinoids and evolve into new grains by SGR. At a higher temperature, recrystallized grains have irregular grain boundaries and bimodal grain-size distribution, implying grain-boundary migration (GBM) recrystallization. At a lower temperature, new grains nucleating and growing along twin boundaries inherit lattice orientation of twin domain, and with the nucleation site and usually a smaller grain size, they can be distinguished from new grains by SGR. At a higher temperature, however, the distinction is difficult at present due to extensive GBM. For the trilobites, there is only local GBM with no significant change in grain size, and flattening of grains reflects the bulk strain at a lower temperature. At a higher temperature, individual grains of the trilobites are equi-axed with weakened LPO, although the strain of trilobites is higher than bulk strain. These microfabrics suggest that the dominant deformation mechanism of

  15. New model of succession of Middle and Late Pennsylvanian fossil communities in north Texas, Mid-Continent, and Appalachians with implications on black shale controversy

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Boardman, D.R. II; Yancey, T.E.; Mapes, R.H.

    1983-03-01

    A new model for the succession of Pennsylvanian fossil communities, preserved in cyclothems, is proposed on the basis of more than 200 fossil localities in the Mid-Continent, Appalachians, and north Texas. Early models for Mid-Continent cyclothems placed the black shales in shallow water, with maximum transgression at the fusulinid-bearing zone in the overlying limestone. The most recent model proposed that the black phosphatic shales, which commonly occur between two subtidal carbonates, are widespread and laterally continuous over great distances and represent maximum transgression. The black phosphatic shales contain: ammonoids; inarticulate brachiopods; radiolarians; conularids; shark material and abundant and diverse conodonts.more » The black shales grade vertically and laterally into dark gray-black shales which contain many of the same pelagic and epipelagic forms found in the phosphatic black shales. This facies contains the deepest water benthic community. Most of these forms are immature, pyritized, and generally are preserved as molds. The dark gray-black facies grades into a medium gray shale facies which contains a mature molluscan fauna. The medium gray shale grades into a lighter gray facies, which is dominated by brachiopods, crinoids, and corals, with occasional bivalves and gastropods. (These facies are interpreted as being a moderate to shallow depth shelf community). The brachiopid-crinoid community is succeeded by shallow water communities which may have occupied shoreline, lagoonal, bay, interdeltaic, or shallow prodeltaic environments.« less

  16. Micro- to nanostructure and geochemistry of extant crinoidal echinoderm skeletons.

    PubMed

    Gorzelak, P; Stolarski, J; Mazur, M; Meibom, A

    2013-01-01

    This paper reports the results of micro- to nanostructural and geochemical analyses of calcitic skeletons from extant deep-sea stalked crinoids. Fine-scale (SEM, FESEM, AFM) observations show that the crinoid skeleton is composed of carbonate nanograins, about 20-100 nm in diameter, which are partly separated by what appears to be a few nm thick organic layers. Sub-micrometre-scale geochemical mapping of crinoid ossicles using a NanoSIMS ion microprobe, combined with synchrotron high-spatial-resolution X-ray micro-fluorescence (μ-XRF) maps and X-ray absorption near-edge structure spectroscopy (XANES) show that high Mg concentration in the central region of the stereom bars correlates with the distribution of S-sulphate, which is often associated with sulphated polysaccharides in biocarbonates. These data are consistent with biomineralization models suggesting a close association between organic components (including sulphated polysaccharides) and Mg ions. Additionally, geochemical analyses (NanoSIMS, energy dispersive spectroscopy) reveal that significant variations in Mg occur at many levels: within a single stereom trabecula, within a single ossicle and within a skeleton of a single animal. Together, these data suggest that physiological factors play an important role in controlling Mg content in crinoid skeletons and that great care should be taken when using their skeletons to reconstruct, for example, palaeotemperatures and Mg/Ca palaeo-variations of the ocean. © 2012 Blackwell Publishing Ltd.

  17. Fossil preservation and the stratigraphic ranges of taxa

    NASA Technical Reports Server (NTRS)

    Foote, M.; Raup, D. M.

    1996-01-01

    The incompleteness of the fossil record hinders the inference of evolutionary rates and patterns. Here, we derive relationships among true taxonomic durations, preservation probability, and observed taxonomic ranges. We use these relationships to estimate original distributions of taxonomic durations, preservation probability, and completeness (proportion of taxa preserved), given only the observed ranges. No data on occurrences within the ranges of taxa are required. When preservation is random and the original distribution of durations is exponential, the inference of durations, preservability, and completeness is exact. However, reasonable approximations are possible given non-exponential duration distributions and temporal and taxonomic variation in preservability. Thus, the approaches we describe have great potential in studies of taphonomy, evolutionary rates and patterns, and genealogy. Analyses of Upper Cambrian-Lower Ordovician trilobite species, Paleozoic crinoid genera, Jurassic bivalve species, and Cenozoic mammal species yield the following results: (1) The preservation probability inferred from stratigraphic ranges alone agrees with that inferred from the analysis of stratigraphic gaps when data on the latter are available. (2) Whereas median durations based on simple tabulations of observed ranges are biased by stratigraphic resolution, our estimates of median duration, extinction rate, and completeness are not biased.(3) The shorter geologic ranges of mammalian species relative to those of bivalves cannot be attributed to a difference in preservation potential. However, we cannot rule out the contribution of taxonomic practice to this difference. (4) In the groups studied, completeness (proportion of species [trilobites, bivalves, mammals] or genera [crinoids] preserved) ranges from 60% to 90%. The higher estimates of completeness at smaller geographic scales support previous suggestions that the incompleteness of the fossil record reflects loss of

  18. Explosive diversification following a benthic to pelagic shift in freshwater fishes

    PubMed Central

    2013-01-01

    Background Interspecific divergence along a benthic to pelagic habitat axis is ubiquitous in freshwater fishes inhabiting lentic environments. In this study, we examined the influence of this habitat axis on the macroevolution of a diverse, lotic radiation using mtDNA and nDNA phylogenies for eastern North America’s most species-rich freshwater fish clade, the open posterior myodome (OPM) cyprinids. We used ancestral state reconstruction to identify the earliest benthic to pelagic transition in this group and generated fossil-calibrated estimates of when this shift occurred. This transition could have represented evolution into a novel adaptive zone, and therefore, we tested for a period of accelerated lineage accumulation after this historical habitat shift. Results Ancestral state reconstructions inferred a similar and concordant region of our mtDNA and nDNA based gene trees as representing the shift from benthic to pelagic habitats in the OPM clade. Two independent tests conducted on each gene tree suggested an increased diversification rate after this inferred habitat transition. Furthermore, lineage through time analyses indicated rapid early cladogenesis in the clade arising after the benthic to pelagic shift. Conclusions A burst of diversification followed the earliest benthic to pelagic transition during the radiation of OPM cyprinids in eastern North America. As such, the benthic/pelagic habitat axis has likely influenced the generation of biodiversity across disparate freshwater ecosystems. PMID:24341464

  19. Explosive diversification following a benthic to pelagic shift in freshwater fishes.

    PubMed

    Hollingsworth, Phillip R; Simons, Andrew M; Fordyce, James A; Hulsey, C Darrin

    2013-12-17

    Interspecific divergence along a benthic to pelagic habitat axis is ubiquitous in freshwater fishes inhabiting lentic environments. In this study, we examined the influence of this habitat axis on the macroevolution of a diverse, lotic radiation using mtDNA and nDNA phylogenies for eastern North America's most species-rich freshwater fish clade, the open posterior myodome (OPM) cyprinids. We used ancestral state reconstruction to identify the earliest benthic to pelagic transition in this group and generated fossil-calibrated estimates of when this shift occurred. This transition could have represented evolution into a novel adaptive zone, and therefore, we tested for a period of accelerated lineage accumulation after this historical habitat shift. Ancestral state reconstructions inferred a similar and concordant region of our mtDNA and nDNA based gene trees as representing the shift from benthic to pelagic habitats in the OPM clade. Two independent tests conducted on each gene tree suggested an increased diversification rate after this inferred habitat transition. Furthermore, lineage through time analyses indicated rapid early cladogenesis in the clade arising after the benthic to pelagic shift. A burst of diversification followed the earliest benthic to pelagic transition during the radiation of OPM cyprinids in eastern North America. As such, the benthic/pelagic habitat axis has likely influenced the generation of biodiversity across disparate freshwater ecosystems.

  20. The base of the Campanian: a magnetostratigraphic definition, integrated biostratigraphy and isotope stratigraphy

    NASA Astrophysics Data System (ADS)

    Wagreich, Michael; Dinarès-Turell, Jaume; Wolfgring, Erik; Omer Yilmaz, Ismail

    2017-04-01

    The base of the Campanian, the longest stage of the Late Cretaceous, is still not defined by a GSSP (Global Boundary Stratotype Section and Point). Furthermore, no consensus exists about the boundary criterion: macrofossils like ammonites and inoceramids suffer from bioprovincialism. The LAD of the crinoid Marsupites testudinarius was proposed as the primary fossil marker. But this marker is mainly a chalk facies fossil, which is rare to absent in pelagic low-latitudes, absent in oceanic sections, and the LAD may be influenced by palaeoenvironmental and palaecological settings. More recently, a boundary defined by magnetostratigraphy, i.e. the base of Chron C33r after the Long Cretaceous Normal Chron C34n, has been discussed. Western Tethyan pelagic sections from Austria and Turkey provide means for integrating various stratigraphic methods. A robust magnetostratigraphic signal was found at the Postalm section (Gosau Group, Northern Calcareous Alps, Austria) which may serve as a reference section for the northwestern Tethys. Including nearby complementary sections, palaeomagnetic data can be integrated with strontium isotope stratigraphy and stable isotope data, planktonic foraminifera and calcareous nannoplankton biostratigraphy, , and ammonite, crinoid and inoceramid data. The Postalm section shows a deepening trend from upper Santonian conglomerates and grey shelf marls to pelagic bathyal red marly limestones of Campanian age. Palaeomagnetic data allow identifying the top of Chron C34n and the following reversal in the lower part of the red marly limestones. A 1 m-thick interval of high magnetic susceptibility is present at the end of C34n. Two of the main suggested biomarkers to pinpoint the Santonian-Campanian boundary, i.e. the last occurrence of the planktonic foraminifer Dicarinella asymetrica and the first occurrence of the nannofossil Broinsonia parca parca, occur in close proximity to the reversal, which is suggested herein as the primary marker event

  1. Depositional facies and sequence stratigraphy of a Lower Carboniferous bryozoan-crinoidal carbonate ramp in the Illinois Basin, mid-continent USA

    USGS Publications Warehouse

    Lasemi, Z.; Norby, R.D.; Treworgy, J.D.

    1998-01-01

    The Lower Carboniferous Fort Payne and Ullin Formations in the Illinois Basin form the transgressive and highstand systems tracts that were deposited in a carbonate ramp setting. During deposition of the Ullin Limestone, biotic communities dominated by fenestrate bryozoans and echinoderms (primarily crinoids) proliferated, possibly in response to global tectonic, biological, and oceanographic events that affected bathymetry and nutrient supply. The Fort Payne Formation consists of a dark grey-brown, siliceous and argillaceous lime mudstone in the lower part (transgressive systems tract) and a very fine-grained wackestone to packstone with rare mud mounds in the upper part (early high-stand), and was deposited in an outer ramp to basinal environment. During deposition of the lower Ullin Limestone (mostly early highstand), bryozoan-crinoidal build-ups accreted both laterally and vertically into several relatively large carbonate banks, which were partly surrounded by siliceous Fort Payne sea. Bryozoans (primarily fenestrates) were especially prevalent during the late stage of bank development and formed mud-free bioherms up to 120 m thick. In places, carbonate mud mounds also formed during the early stage of bank deposition. Bioherm development declined during deposition of the upper Ullin Limestone (late highstand), and a broad, storm-dominated carbonate ramp was established that became the site for widespread deposition of bryozoan-crinoidal sandwaves. Gradual shallowing led to ooid formation at the end of Ullin deposition. This sequence was terminated by a relative rise in sea level that resulted in deposition of the transgressive facies of the lower part of the overlying Salem Limestone. The depositional style and the nature of skeletal material of the Fort Payne and Ullin Formations are similar to those of cool-water carbonates. A deep-water setting along with upwelling of cool, nutrient-rich oceanic waters may have been responsible for the proliferation of

  2. Phylogeny of Comatulidae (Echinodermata: Crinoidea: Comatulida): a new classification and an assessment of morphological characters for crinoid taxonomy.

    PubMed

    Summers, Mindi M; Messing, Charles G; Rouse, Greg W

    2014-11-01

    Comatulidae Fleming, 1828 (previously, and incorrectly, Comasteridae A.H. Clark, 1908a), is a group of feather star crinoids currently divided into four accepted subfamilies, 21 genera and approximately 95 nominal species. Comatulidae is the most commonly-encountered and species-rich crinoid group on shallow tropical coral reefs, particularly in the Indo-western Pacific region (IWP). We conducted a molecular phylogenetic analysis of the group with concatenated data from up to seven genes for 43 nominal species spanning 17 genera and all subfamilies. Basal nodes returned low support, but maximum likelihood, maximum parsimony, and Bayesian analyses were largely congruent, permitting an evaluation of current taxonomy and analysis of morphological character transformations. Two of the four current subfamilies were paraphyletic, whereas 15 of the 17 included genera returned as monophyletic. We provide a new classification with two subfamilies, Comatulinae and Comatellinae n. subfamily Summers, Messing, & Rouse, the former containing five tribes. We revised membership of analyzed genera to make them all clades and erected Anneissia n. gen. Summers, Messing, & Rouse. Transformation analyses for morphological features generally used in feather star classification (e.g., ray branching patterns, articulations) and those specifically for Comatulidae (e.g., comb pinnule form, mouth placement) were labile with considerable homoplasy. These traditional characters, in combination, allow for generic diagnoses, but in most cases we did not recover apomorphies for subfamilies, tribes, and genera. New morphological characters that will be informative for crinoid taxonomy and identification are still needed. DNA sequence data currently provides the most reliable method of identification to the species-level for many taxa of Comatulidae. Copyright © 2014 Elsevier Inc. All rights reserved.

  3. A new Secondary metabolites of the crinoid (Comanthina schlegeli) associated fungus Alternaria brassicae 93.

    PubMed

    Zhang, Liu-Hong; Wang, Hong-Wei; Xu, Jia-Yi; Li, Jing; Liu, Lan

    2016-10-01

    Fungus Alternaria brassicae 93 isolated from crinoid (Comanthina schlegeli), which was collected from the South China Sea. Six compounds were isolated from A. brassicae 93, including one new compound (1), along with five known compounds, ochratoxin A methyl ester (2), cis-4-hydroxym-ellein (3), (R)-7-hydroxymellein (4), trans-2-anhydromevalonic (5) and protocatechuic acid (6). Their structures were determined by spectroscopic methods and comparison with reported data. Cytotoxicity against two human cancer cell lines and antibacterial activity against twelve aquatic bacteria of compound 1 were also tested.

  4. A fossil unicorn crestfish (Teleostei, Lampridiformes, Lophotidae) from the Eocene of Iran.

    PubMed

    Davesne, Donald

    2017-01-01

    Lophotidae, or crestfishes, is a family of rare deep-sea teleosts characterised by an enlarged horn-like crest on the forehead. They are poorly represented in the fossil record, by only three described taxa. One specimen attributed to Lophotidae has been described from the pelagic fauna of the middle-late Eocene Zagros Basin, Iran. Originally considered as a specimen of the fossil lophotid † Protolophotus , it is proposed hereby as a new genus and species † Babelichthys olneyi , gen. et sp. nov., differs from the other fossil lophotids by its relatively long and strongly projecting crest, suggesting a close relationship with the modern unicorn crestfish, Eumecichthys . This new taxon increases the diversity of the deep-sea teleost fauna to which it belongs, improving our understanding of the taxonomic composition of the early Cenozoic mesopelagic ecosystems.

  5. Four Aromatic Sulfates with an Inhibitory Effect against HCV NS3 Helicase from the Crinoid Alloeocomatella polycladia.

    PubMed

    Hermawan, Idam; Furuta, Atsushi; Higashi, Masahiro; Fujita, Yoshihisa; Akimitsu, Nobuyoshi; Yamashita, Atsuya; Moriishi, Kohji; Tsuneda, Satoshi; Tani, Hidenori; Nakakoshi, Masamichi; Tsubuki, Masayoshi; Sekiguchi, Yuji; Noda, Naohiro; Tanaka, Junichi

    2017-04-11

    Bioassay-guided separation of a lipophilic extract of the crinoid Alloeocomatella polycladia , inhibiting the activity of HCV NS3 helicase, yielded two groups of molecules: cholesterol sulfate and four new aromatic sulfates 1 - 4 . The structures of the aromatics were elucidated by spectroscopic analysis in addition to theoretical studies. The aromatic sulfates 1 - 4 showed moderate inhibition against NS3 helicase with IC 50 values of 71, 95, 7, and 5 μM, respectively.

  6. A fossil unicorn crestfish (Teleostei, Lampridiformes, Lophotidae) from the Eocene of Iran

    PubMed Central

    2017-01-01

    Lophotidae, or crestfishes, is a family of rare deep-sea teleosts characterised by an enlarged horn-like crest on the forehead. They are poorly represented in the fossil record, by only three described taxa. One specimen attributed to Lophotidae has been described from the pelagic fauna of the middle-late Eocene Zagros Basin, Iran. Originally considered as a specimen of the fossil lophotid †Protolophotus, it is proposed hereby as a new genus and species †Babelichthys olneyi, gen. et sp. nov., differs from the other fossil lophotids by its relatively long and strongly projecting crest, suggesting a close relationship with the modern unicorn crestfish, Eumecichthys. This new taxon increases the diversity of the deep-sea teleost fauna to which it belongs, improving our understanding of the taxonomic composition of the early Cenozoic mesopelagic ecosystems. PMID:28674642

  7. Four Aromatic Sulfates with an Inhibitory Effect against HCV NS3 Helicase from the Crinoid Alloeocomatella polycladia

    PubMed Central

    Hermawan, Idam; Furuta, Atsushi; Higashi, Masahiro; Fujita, Yoshihisa; Akimitsu, Nobuyoshi; Yamashita, Atsuya; Moriishi, Kohji; Tsuneda, Satoshi; Tani, Hidenori; Nakakoshi, Masamichi; Tsubuki, Masayoshi; Sekiguchi, Yuji; Noda, Naohiro; Tanaka, Junichi

    2017-01-01

    Bioassay-guided separation of a lipophilic extract of the crinoid Alloeocomatella polycladia, inhibiting the activity of HCV NS3 helicase, yielded two groups of molecules: cholesterol sulfate and four new aromatic sulfates 1–4. The structures of the aromatics were elucidated by spectroscopic analysis in addition to theoretical studies. The aromatic sulfates 1–4 showed moderate inhibition against NS3 helicase with IC50 values of 71, 95, 7, and 5 μM, respectively. PMID:28398249

  8. Early post-metamorphic, Carboniferous blastoid reveals the evolution and development of the digestive system in echinoderms.

    PubMed

    Rahman, Imran A; Waters, Johnny A; Sumrall, Colin D; Astolfo, Alberto

    2015-10-01

    Inferring the development of the earliest echinoderms is critical to uncovering the evolutionary assembly of the phylum-level body plan but has long proven problematic because early ontogenetic stages are rarely preserved as fossils. Here, we use synchrotron tomography to describe a new early post-metamorphic blastoid echinoderm from the Carboniferous (approx. 323 Ma) of China. The resulting three-dimensional reconstruction reveals a U-shaped tubular structure in the fossil interior, which is interpreted as the digestive tract. Comparisons with the developing gut of modern crinoids demonstrate that crinoids are an imperfect analogue for many extinct groups. Furthermore, consideration of our findings in a phylogenetic context allows us to reconstruct the evolution and development of the digestive system in echinoderms more broadly; there was a transition from a straight to a simple curved gut early in the phylum's evolution, but additional loops and coils of the digestive tract (as seen in crinoids) were not acquired until much later. © 2015 The Author(s).

  9. Pelagic larval duration predicts extinction risk in a freshwater fish clade.

    PubMed

    Douglas, Morgan; Keck, Benjamin P; Ruble, Crystal; Petty, Melissa; Shute, J R; Rakes, Patrick; Hulsey, C Darrin

    2013-01-01

    Pelagic larval duration (PLD) can influence evolutionary processes ranging from dispersal to extinction in aquatic organisms. Using estimates of PLD obtained from species of North American darters (Percidae: Etheostomatinae), we demonstrate that this freshwater fish clade exhibits surprising variation in PLD. Comparative analyses provide some evidence that higher stream gradients favour the evolution of shorter PLD. Additionally, similar to patterns in the marine fossil record in which lower PLD is associated with greater extinction probability, we found a reduced PLD in darter lineages was evolutionarily associated with extinction risk. Understanding the causes and consequences of PLD length could lead to better management and conservation of organisms in our increasingly imperiled aquatic environments.

  10. Benthic Foraminifera, Food in the Deep Sea, and Limits to Bentho-Pelagic Coupling

    NASA Astrophysics Data System (ADS)

    Thomas, E.; Boscolo-Galazzo, F.; Arreguin-Rodrigu, G. J.; Ortiz, S.; Alegret, L.

    2015-12-01

    The deep-sea is the largest habitat on Earth, contains highly diverse biota, but is very little known. Many of its abundant benthic biota (e.g., nematodes) are not preserved in the fossil record. Calcareous and agglutinated benthic foraminifera (unicellular eukaryotes, Rhizaria; efficient dispersers) and ostracodes (Animalia, Crustacea; non-efficient dispersers) are the most common organisms providing a fossil record of deep-sea environments. Very little food is supplied to the deep-sea, because organic matter produced by photosynthesis is largely degraded before it arrives at the seafloor. Only a few % of organic matter is carried to the ocean bottom by 'marine snow', with its particle size and behavior in the water column controlled by surface ecosystem structure, including type of dominant primary producers (diatoms, cyanobacteria). Food supply and its seasonality are generally seen as the dominant control on benthic assemblages (combined with oxygenation), providing bentho-pelagic coupling between primary and benthic productivity. Benthic foraminiferal assemblages (composition and density) thus are used widely to estimate past productivity, especially during episodes of global climate change, ocean acidification, and mass extinction of primary producers. We show that some environmental circumstances may result in interrupting bentho-pelagic coupling, e.g. through lateral supply of organic matter along continental margins (adding more refractory organic matter), through trophic focusing and/or fine particle winnowing on seamounts (giving an advantage to suspension feeders), and through carbonate undersaturation (giving advantage to infaunal over epifaunal calcifyers). In addition, increased remineralization of organic matter combined with increased metabolic rates may cause assemblages to reflect more oligotrophic conditions at stable primary productivity during periods of global warming. As a result, benthic foraminiferal accumulation rates must be carefully

  11. 76 FR 72643 - Western Pacific Pelagic Fisheries; Closure of the Hawaii Shallow-Set Pelagic Longline Fishery Due...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-11-25

    .... 080225267-91393-03] RIN 0648-XA370 Western Pacific Pelagic Fisheries; Closure of the Hawaii Shallow- Set...: Temporary rule; fishery closure. SUMMARY: NMFS closes the shallow-set pelagic longline fishery north of the Equator for all vessels registered under the Hawaii longline limited access program. The shallow-set...

  12. Piscivore diet response to a collapse in pelagic prey populations

    USGS Publications Warehouse

    Zeug, Steven; Feyrer, Frederick; Brodsky, Annie; Melgo, Jenny

    2017-01-01

    Pelagic fish populations in the upper San Francisco Estuary have experienced significant declines since the turn of the century; a pattern known as the pelagic organism decline (POD). This study investigated food habits of piscivorous fishes over two consecutive fall seasons following the decline of pelagic fish prey. Specifically, this study addressed the contribution of pelagic versus benthic prey to piscivorous fish diets, including the frequency of predation on special-status pelagic species, and the spatial variability in prey consumption. The piscivore community was dominated by Striped Bass and also included small numbers of Sacramento Pikeminnow and Largemouth Bass. Overall, pelagic prey items contributed less than 10% of the diet by weight in both years, whereas pre-POD studies gleaned from the literature found contributions of 39–100%, suggesting a major switch from pelagic to benthic prey resources. Between-year variation in piscivore diets reflected differences in environmental conditions associated with variation in freshwater outflow. No special status fish species were detected in any of the piscivore stomachs examined. The consequences of this pelagic to benthic diet shift warrants further investigation to understand its ecological relevance.

  13. Conservation of deep pelagic biodiversity.

    PubMed

    Robison, Bruce H

    2009-08-01

    The deep ocean is home to the largest ecosystems on our planet. This vast realm contains what may be the greatest number of animal species, the greatest biomass, and the greatest number of individual organisms in the living world. Humans have explored the deep ocean for about 150 years, and most of what is known is based on studies of the deep seafloor. In contrast, the water column above the deep seabed comprises more than 90% of the living space, yet less than 1% of this biome has been explored. The deep pelagic biota is the largest and least-known major faunal group on Earth despite its obvious importance at the global scale. Pelagic species represent an incomparable reservoir of biodiversity. Although we have yet to discover and describe the majority of these species, the threats to their continued existence are numerous and growing. Conserving deep pelagic biodiversity is a problem of global proportions that has never been addressed comprehensively. The potential effects of these threats include the extensive restructuring of entire ecosystems, changes in the geographical ranges of many species, large-scale elimination of taxa, and a decline in biodiversity at all scales. This review provides an initial framework of threat assessment for confronting the challenge of conserving deep pelagic biodiversity; and it outlines the need for baseline surveys and protected areas as preliminary policy goals.

  14. A new genus of endoparasitic copepods (Cyclopoida: Enterognathidae), forming a gall in the calyx of deep-sea crinoids.

    PubMed

    Ohtsuka, Susumu; Kitazawa, Kota; Boxshall, Geoffrey A

    2010-08-01

    A new genus and species of cyclopoid copepod belonging to the family Enterognathidae, Parenterognathus troglodytes, is described from a gall on the calyx of the deep-sea crinoid Glyptometra crassa (Clark, 1912) collected at depths of 775-787 m off Kumano-nada, middle Japan. The new genus can be distinguished from the three known genera of the family by body tagmosis and by the segmentation and armature of the appendages. This is the first record of this family from the Pacific Ocean. This family seems to be host-specific to relatively basal deuterostomes, such as echinoderms and hemichordates. The evolutionary transformation and history of the Enterognathidae are briefly discussed.

  15. Preservation of high primary porosity in Paleozoic crinoidal and bioclastic grainstones: Mississippian and Permian subsurface examples from western Canada and Texas

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Eby, D.E.; Kirkby, K.C.

    1991-03-01

    Limestone reservoirs with anomalously high porosity and permeability exist in several Paleozoic basinal settings in western North America. They are commonly either crinoidal or bioclastic grainstones that escaped the varied diagenetic environments that most carbonate shelves and slopes experience during sea-level fluctuations. Thus, these diagenetically 'conservative' settings can preserve most of the primary depositional porosity in calcarenites because of sedimentation and burial within normal marine basinal waters. These excellent reservoirs are often encased in deep marine, highly organic shales which may serve as both source rock and seal. Several subsurface examples of remarkable primary porosity occur within Mississippian Waulsortian buildupsmore » of western Canada and north-center Texas. Buildups within the Pekisko Formation of Alberta, for instance, contain abundant flanking crinoidal/bryozoan grainstones with up to 25% primary porosity and 10 d permeability. Muddy core facies with productive flank and capping calcarenites are up to 300 ft (90 m) thick and 600-1,200 ft (182-364 m) in diameter. The Bowar 'reef complex' in the Chappel Formation of Stephens County, Texas (2.6 MMBO to date) is similar in many respects to the Canadian buildups. Diagenetic 'conservatism' has also promoted excellent reservoir quality within the outer slope to basinal lower Permian bioclastic grain-flow deposit ('Wolfcamp detrital') plays of the eastern Midland basin where significant diagenesis was arrested after basinal deposition. Interpretations of these examples based upon core and outcrop control provide analogs for future exploration and exploitation in highly porous basinal carbonate reservoirs.« less

  16. Accommodating Dynamic Oceanographic Processes and Pelagic Biodiversity in Marine Conservation Planning

    PubMed Central

    Grantham, Hedley S.; Game, Edward T.; Lombard, Amanda T.; Hobday, Alistair J.; Richardson, Anthony J.; Beckley, Lynnath E.; Pressey, Robert L.; Huggett, Jenny A.; Coetzee, Janet C.; van der Lingen, Carl D.; Petersen, Samantha L.; Merkle, Dagmar; Possingham, Hugh P.

    2011-01-01

    Pelagic ecosystems support a significant and vital component of the ocean's productivity and biodiversity. They are also heavily exploited and, as a result, are the focus of numerous spatial planning initiatives. Over the past decade, there has been increasing enthusiasm for protected areas as a tool for pelagic conservation, however, few have been implemented. Here we demonstrate an approach to plan protected areas that address the physical and biological dynamics typical of the pelagic realm. Specifically, we provide an example of an approach to planning protected areas that integrates pelagic and benthic conservation in the southern Benguela and Agulhas Bank ecosystems off South Africa. Our aim was to represent species of importance to fisheries and species of conservation concern within protected areas. In addition to representation, we ensured that protected areas were designed to consider pelagic dynamics, characterized from time-series data on key oceanographic processes, together with data on the abundance of small pelagic fishes. We found that, to have the highest likelihood of reaching conservation targets, protected area selection should be based on time-specific data rather than data averaged across time. More generally, we argue that innovative methods are needed to conserve ephemeral and dynamic pelagic biodiversity. PMID:21311757

  17. Upper Campanian calciclastic turbidite sequences from the Hacımehmet area (eastern Pontides, NE Turkey): integrated biostratigraphy and microfacies analysis

    NASA Astrophysics Data System (ADS)

    Sari, Bilal; Kandemir, Raif; Özer, Sacit; Walaszczyk, Ireneusz; Görmüş, Muhittin; Demircan, Huriye; Yilmaz, Cemil

    2014-12-01

    The upper Campanian (Cretaceous) of the Hacımehmet area (south of the city of Trabzon; Eastern Pontides) is mainly composed of calciclastic turbidites. The basinal unit of the 119 m thick succession includes thin red pelagic limestone interlayers and conglomerates dominated by volcanic clasts. The overlying upper slope and lower slope units of the sequence consist of an alternation of allochthonous calcarenite/calcirudite beds and pelagic marls and mudstones. Calcarenite/calcirudite beds dominate the upper slope unit of the succession and are composed of transported material, including benthic foraminifers, red algae, bryozoan, crinoid and rudist fragments, inoceramid bivalve prisms and neritic and pelagic carbonate lithoclasts. The occurrence of Helicorbitoides boluensis (Sirel) extracted from the calcarenite/calcirudite beds indicates a Campanian age. Identifiable rudists such as Joufia reticulata Boehm, Bournonia cf. anatolica Ozer, Biradiolites cf. bulgaricus Pamouktchiev and ?Biradiolites sp. from the upper slope unit of the succession indicate a late Campanian- Maastrichtian age. The planktonic foraminifers within the red pelagic limestone beds, marls and mudstones throughout the succession consist mainly of Campanian-Maastrichtian forms and suggest mainly basinal depositional conditions. The presence of Radotruncana cf. calcarata (Cushman) accompanied by Globotruncanita elevata (Brotzen) in the basinal unit of the succession indicates an early late Campanian age for the lower part of the succession. Inoceramid bivalves have been collected from the upper part of the succession. The fauna is dominated by `Inoceramus' tenuilineatus Hall and Meek, 1854 and Cataceramus haldemensis (Giers, 1964); other taxa recognised are: `Inoceramus' algeriensis Heinz, 1932, Platyceramus vanuxemi (Meek and Hayden, 1860), `Inoceramus' cf. nebrascensis Owen, 1852, Cataceramus aff. barabini (Morton, 1834), Cataceramus gandjaensis (Aliev, 1956), and `Inoceramus' sp.; the assemblage

  18. Fractal Hypothesis of the Pelagic Microbial Ecosystem-Can Simple Ecological Principles Lead to Self-Similar Complexity in the Pelagic Microbial Food Web?

    PubMed

    Våge, Selina; Thingstad, T Frede

    2015-01-01

    Trophic interactions are highly complex and modern sequencing techniques reveal enormous biodiversity across multiple scales in marine microbial communities. Within the chemically and physically relatively homogeneous pelagic environment, this calls for an explanation beyond spatial and temporal heterogeneity. Based on observations of simple parasite-host and predator-prey interactions occurring at different trophic levels and levels of phylogenetic resolution, we present a theoretical perspective on this enormous biodiversity, discussing in particular self-similar aspects of pelagic microbial food web organization. Fractal methods have been used to describe a variety of natural phenomena, with studies of habitat structures being an application in ecology. In contrast to mathematical fractals where pattern generating rules are readily known, however, identifying mechanisms that lead to natural fractals is not straight-forward. Here we put forward the hypothesis that trophic interactions between pelagic microbes may be organized in a fractal-like manner, with the emergent network resembling the structure of the Sierpinski triangle. We discuss a mechanism that could be underlying the formation of repeated patterns at different trophic levels and discuss how this may help understand characteristic biomass size-spectra that hint at scale-invariant properties of the pelagic environment. If the idea of simple underlying principles leading to a fractal-like organization of the pelagic food web could be formalized, this would extend an ecologists mindset on how biological complexity could be accounted for. It may furthermore benefit ecosystem modeling by facilitating adequate model resolution across multiple scales.

  19. Fractal Hypothesis of the Pelagic Microbial Ecosystem—Can Simple Ecological Principles Lead to Self-Similar Complexity in the Pelagic Microbial Food Web?

    PubMed Central

    Våge, Selina; Thingstad, T. Frede

    2015-01-01

    Trophic interactions are highly complex and modern sequencing techniques reveal enormous biodiversity across multiple scales in marine microbial communities. Within the chemically and physically relatively homogeneous pelagic environment, this calls for an explanation beyond spatial and temporal heterogeneity. Based on observations of simple parasite-host and predator-prey interactions occurring at different trophic levels and levels of phylogenetic resolution, we present a theoretical perspective on this enormous biodiversity, discussing in particular self-similar aspects of pelagic microbial food web organization. Fractal methods have been used to describe a variety of natural phenomena, with studies of habitat structures being an application in ecology. In contrast to mathematical fractals where pattern generating rules are readily known, however, identifying mechanisms that lead to natural fractals is not straight-forward. Here we put forward the hypothesis that trophic interactions between pelagic microbes may be organized in a fractal-like manner, with the emergent network resembling the structure of the Sierpinski triangle. We discuss a mechanism that could be underlying the formation of repeated patterns at different trophic levels and discuss how this may help understand characteristic biomass size-spectra that hint at scale-invariant properties of the pelagic environment. If the idea of simple underlying principles leading to a fractal-like organization of the pelagic food web could be formalized, this would extend an ecologists mindset on how biological complexity could be accounted for. It may furthermore benefit ecosystem modeling by facilitating adequate model resolution across multiple scales. PMID:26648929

  20. Onondage pinnacle reefs in New York State

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Friedman, G.M.

    1995-09-01

    Onondaga pinnacle reefs, part of the Onondaga Formation, developed in an epeiric setting of the lowermost Middle Devonian (Eifelian). The reefs were initiated as coral-crinoidal mounds in the Edgecliff Member of the formation. Whereas most Devonian reefs are composed of rugose corals. Coral is the predominant kind of fossil, followed by crinoids, brachiopods, mollusks, undifferentiated skeletal debris, and possible sponges. The initial mineralogy of the corals is inferred to have been calcite. The porosity of these reefs is almost unique among reef reservoirs. most reefs produce from secondary or diagenetic porosity; by contrast Onondaga reefs display primary intracoralline or frameworkmore » porosity. Between framework builders and/or skeletal particles cryptocrystalline/microcrystalline cement fills pores. As observed in modern reefs this kind of cement resembles micrite, but probable formed as high-magnesian calcite in a high-energy setting. Syntaxial or rim cement common lines crinoid particles. Some of these pinnacle reefs, formerly gas producers, are presently under development as gas-storage reservoirs.« less

  1. Crinoids of the stalked family Hyocrinidae (Echinodermata) collected by the USNS Eltanin in the Southern Ocean.

    PubMed

    Roux, Michel

    2015-07-27

    Rare species only known from one or a few specimens, mainly collected from abyssal and hadal zones in the Southern Ocean are represented in a collection of stalked crinoids attributed to the family Hyocrinidae (Crinoidea, Echinodermata). The species studied here include Belyaevicrinus latipinnulus Mironov & Sorokina, 1998, Feracrinus heinzelleri Bohn, 2012, Ptilocrinus brucei Vaney, 1908 and Thalassocrinus clausus Mironov & Sorokina, 1998. This collection, mainly collected during the 1960s USNS Eltanin cruises, is housed at the Smithsonian Institution National Museum of Natural History in Washington DC. A detailed study of this material using SEM observations of ossicle articular facets and pinnule architecture allows the emendation of species diagnoses, and a better description of intraspecific variation and of changes in characters through ontogeny. Geographical and depth range extension of species are reported. A gonad with submature ovocytes of 100-150 µm was exceptionally preserved at the base of a genital pinnule of P. brucei. This species likely has a lecithotrophic larval development.

  2. Land Use Affects Carbon Sources to the Pelagic Food Web in a Small Boreal Lake

    PubMed Central

    Rinta, Päivi; van Hardenbroek, Maarten; Jones, Roger I.; Kankaala, Paula; Rey, Fabian; Szidat, Sönke; Wooller, Matthew J.; Heiri, Oliver

    2016-01-01

    Small humic forest lakes often have high contributions of methane-derived carbon in their food webs but little is known about the temporal stability of this carbon pathway and how it responds to environmental changes on longer time scales. We reconstructed past variations in the contribution of methanogenic carbon in the pelagic food web of a small boreal lake in Finland by analyzing the stable carbon isotopic composition (δ13C values) of chitinous fossils of planktivorous invertebrates in sediments from the lake. The δ13C values of zooplankton remains show several marked shifts (approx. 10 ‰), consistent with changes in the proportional contribution of carbon from methane-oxidizing bacteria in zooplankton diets. The results indicate that the lake only recently (1950s) obtained its present state with a high contribution of methanogenic carbon to the pelagic food web. A comparison with historical and palaeobotanical evidence indicates that this most recent shift coincided with agricultural land-use changes and forestation of the lake catchment and implies that earlier shifts may also have been related to changes in forest and land use. Our study demonstrates the sensitivity of the carbon cycle in small forest lakes to external forcing and that the effects of past changes in local land use on lacustrine carbon cycling have to be taken into account when defining environmental and ecological reference conditions in boreal headwater lakes. PMID:27487044

  3. 50 CFR 665.817 - American Samoa pelagic fishery area management.

    Code of Federal Regulations, 2010 CFR

    2010-10-01

    ... 50 Wildlife and Fisheries 9 2010-10-01 2010-10-01 false American Samoa pelagic fishery area management. 665.817 Section 665.817 Wildlife and Fisheries FISHERY CONSERVATION AND MANAGEMENT, NATIONAL OCEANIC AND ATMOSPHERIC ADMINISTRATION, DEPARTMENT OF COMMERCE (CONTINUED) FISHERIES IN THE WESTERN PACIFIC Western Pacific Pelagic Fisheries § 665.81...

  4. Widespread kelp-derived carbon in pelagic and benthic nearshore fishes

    USGS Publications Warehouse

    von Biela, Vanessa R.; Newsome, Seth D.; Bodkin, James L.; Kruse, Gordon H.; Zimmerman, Christian E.

    2016-01-01

    Kelp forests provide habitat for diverse and abundant fish assemblages, but the extent to which kelp provides a source of energy to fish and other predators is unclear. To examine the use of kelp-derived energy by fishes we estimated the contribution of kelp- and phytoplankton-derived carbon using carbon (δ13C) and nitrogen (δ15N) isotopes measured in muscle tissue. Benthic-foraging kelp greenling (Hexagrammos decagrammus) and pelagic-foraging black rockfish (Sebastes melanops) were collected at eight sites spanning ∼35 to 60°N from the California Current (upwelling) to Alaska Coastal Current (downwelling) in the northeast Pacific Ocean. Muscle δ13C values were expected to be higher for fish tissue primarily derived from kelp, a benthic macroalgae, and lower for tissue primarily derived from phytoplankton, pelagic microalgae. Muscle δ13C values were higher in benthic-feeding kelp greenling than in pelagic-feeding black rockfish at seven of eight sites, indicating more kelp-derived carbon in greenling as expected. Estimates of kelp carbon contributions ranged from 36 to 89% in kelp greenling and 32 to 65% in black rockfish using carbon isotope mixing models. Isotopic evidence suggests that these two nearshore fishes routinely derive energy from kelp and phytoplankton, across coastal upwelling and downwelling systems. Thus, the foraging mode of nearshore predators has a small influence on their ultimate energy source as energy produced by benthic macroalgae and pelagic microalgae were incorporated in fish tissue regardless of feeding mode and suggest strong and widespread benthic-pelagic coupling. Widespread kelp contributions to benthic- and pelagic-feeding fishes suggests that kelp energy provides a benefit to nearshore fishes and highlights the potential for kelp and fish production to be linked.

  5. Trophic structure of pelagic species in the northwestern Mediterranean Sea

    NASA Astrophysics Data System (ADS)

    Albo-Puigserver, Marta; Navarro, Joan; Coll, Marta; Layman, Craig A.; Palomera, Isabel

    2016-11-01

    Ecological knowledge of food web interactions within pelagic marine communities is often limited, impairing our capabilities to manage these ecologically and economically important marine fish species. Here we used stable isotope analyses to investigate trophic interactions in the pelagic ecosystem of the northwestern Mediterranean Sea during 2012 and 2013. Our results suggest that European sardine, Sardina pilchardus, and anchovy, Engraulis encrasicolus, are consumers located at relatively low levels of the pelagic food web. Unexpectedly, the round sardinella, Sardinella aurita, appeared to be located at a higher trophic level than the other small pelagic fish species, although previous studies found similarity in their diets. Isotope data suggested that trophic niches of species within the genera Trachurus spp. and Scomber spp., were distinct. Atlantic bonito Sarda sarda, European hake Merluccius merluccius and European squid Loligo vulgaris, appeared to feed at higher trophic levels than other species. Despite some intraspecific seasonal variability for some species, community trophic structure appeared relatively stable through the year. These data provide an important step for developing models of food web dynamics in the northwestern Mediterranean Sea.

  6. Seamounts are hotspots of pelagic biodiversity in the open ocean.

    PubMed

    Morato, Telmo; Hoyle, Simon D; Allain, Valerie; Nicol, Simon J

    2010-05-25

    The identification of biodiversity hotspots and their management for conservation have been hypothesized as effective ways to protect many species. There has been a significant effort to identify and map these areas at a global scale, but the coarse resolution of most datasets masks the small-scale patterns associated with coastal habitats or seamounts. Here we used tuna longline observer data to investigate the role of seamounts in aggregating large pelagic biodiversity and to identify which pelagic species are associated with seamounts. Our analysis indicates that seamounts are hotspots of pelagic biodiversity. Higher species richness was detected in association with seamounts than with coastal or oceanic areas. Seamounts were found to have higher species diversity within 30-40 km of the summit, whereas for sets close to coastal habitat the diversity was lower and fairly constant with distance. Higher probability of capture and higher number of fish caught were detected for some shark, billfish, tuna, and other by-catch species. The study supports hypotheses that seamounts may be areas of special interest for management for marine pelagic predators.

  7. Seamounts are hotspots of pelagic biodiversity in the open ocean

    PubMed Central

    Morato, Telmo; Hoyle, Simon D.; Allain, Valerie; Nicol, Simon J.

    2010-01-01

    The identification of biodiversity hotspots and their management for conservation have been hypothesized as effective ways to protect many species. There has been a significant effort to identify and map these areas at a global scale, but the coarse resolution of most datasets masks the small-scale patterns associated with coastal habitats or seamounts. Here we used tuna longline observer data to investigate the role of seamounts in aggregating large pelagic biodiversity and to identify which pelagic species are associated with seamounts. Our analysis indicates that seamounts are hotspots of pelagic biodiversity. Higher species richness was detected in association with seamounts than with coastal or oceanic areas. Seamounts were found to have higher species diversity within 30–40 km of the summit, whereas for sets close to coastal habitat the diversity was lower and fairly constant with distance. Higher probability of capture and higher number of fish caught were detected for some shark, billfish, tuna, and other by-catch species. The study supports hypotheses that seamounts may be areas of special interest for management for marine pelagic predators. PMID:20448197

  8. Acoustic assessment of pelagic lanktivores, 2016

    USGS Publications Warehouse

    Holden, Jeremy P.; Connerton, Michael J.; Weidel, Brian C.

    2017-01-01

    Alewife (Alosa pseudoharengus) and Rainbow Smelt (Osmerus mordax) are the most abundant pelagic planktivores in Lake Ontario (Weidel et al 2017), and the most important prey for salmon and trout, making up greater than 90% of the diet of the top predator, Chinook salmon (Lantry 2001, Brandt 1986), and supporting a multimillion dollar sportfishery. Alewife are also important prey for warm water predators, notably Walleye (Sander vitreus). Abundance of Alewife and smelt has declined since the 1980s, likely due to reduced nutrient loading, proliferation of invasive dreissenid mussels, and predation by stocked salmon and trout. Cisco (Coregonus artedi), a native planktivore, historically dominated the offshore pelagic prey fish of Lake Ontario, but their populations were severely reduced in the mid-20th century due to overfishing and competition with Alewife and smelt. Remnant populations of Cisco still exist, mostly in the eastern basin, and Cisco produce periodic strong year classes once or twice per decade (Owens et al 2003, most recently in 2012 and 2014 (OMNRF, 2017).Alewife (Alosa pseudoharengus) and Rainbow Smelt (Osmerus mordax) are the most abundant pelagic planktivores in Lake Ontario (Weidel et al 2017), and the most important prey for salmon and trout, making up greater than 90% of the diet of the top predator, Chinook salmon (Lantry 2001, Brandt 1986), and supporting a multimillion dollar sportfishery. Alewife are also important prey for warm water predators, notably Walleye (Sander vitreus). Abundance of Alewife and smelt has declined since the 1980s, likely due to reduced nutrient loading, proliferation of invasive dreissenid mussels, and predation by stocked salmon and trout. Cisco (Coregonus artedi), a native planktivore, historically dominated the offshore pelagic prey fish of Lake Ontario, but their populations were severely reduced in the mid-20th century due to overfishing and competition with Alewife and smelt. Remnant populations of Cisco still

  9. Pelage Variation and Distribution of the Black Lion Tamarin, Leontopithecus chrysopygus.

    PubMed

    Garbino, Guilherme Siniciato Terra; Rezende, Gabriela Cabral; Valladares-Padua, Claudio

    2016-01-01

    The black lion tamarin, Leontopithecus chrysopygus, is endemic to the Atlantic Forest of the interior of the state of São Paulo in Brazil. Since the discovery of the species, authors have described variations in the pelage of L. chrysopygus, but the nature of this variation has remained unknown. We examined museum specimens and living animals to see if the pelage of L. chrysopygus varies individually, geographically or ontogenetically. We reviewed all known locality records of the species, and concluded that the pelage of L. chrysopygus varies individually and ontogenetically, but not geographically. The extent of the golden chromogenetic fields is larger in younger individuals, and the pelage becomes darker after successive moults. We examined 2 juveniles that presented a pelage similar to adult L. caissara. Geographically, there is a clear break in the phenotypes attributed to L. caissara and L. chrysopygus, precluding clinal variation and corroborating the specific status of the former. The separation between these 2 species is coincident with a tectonic fault known as the 'Guapiara lineament', and may have been caused by a recent activation of this geological formation. We also suggest a historical contact zone between L. chrysopygus and the marmoset Callithrix aurita in the upper Rio Tietê region. © 2016 S. Karger AG, Basel.

  10. 76 FR 12340 - Proposed Information Collection; Comment Request; Large Pelagic Fishing Survey

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-03-07

    ... DEPARTMENT OF COMMERCE National Oceanic and Atmospheric Administration Proposed Information Collection; Comment Request; Large Pelagic Fishing Survey AGENCY: National Oceanic and Atmospheric..., (301) 713-2328 or [email protected] SUPPLEMENTARY INFORMATION: I. Abstract The Large Pelagic Fishing...

  11. Olfaction Contributes to Pelagic Navigation in a Coastal Shark.

    PubMed

    Nosal, Andrew P; Chao, Yi; Farrara, John D; Chai, Fei; Hastings, Philip A

    2016-01-01

    How animals navigate the constantly moving and visually uniform pelagic realm, often along straight paths between distant sites, is an enduring mystery. The mechanisms enabling pelagic navigation in cartilaginous fishes are particularly understudied. We used shoreward navigation by leopard sharks (Triakis semifasciata) as a model system to test whether olfaction contributes to pelagic navigation. Leopard sharks were captured alongshore, transported 9 km offshore, released, and acoustically tracked for approximately 4 h each until the transmitter released. Eleven sharks were rendered anosmic (nares occluded with cotton wool soaked in petroleum jelly); fifteen were sham controls. Mean swimming depth was 28.7 m. On average, tracks of control sharks ended 62.6% closer to shore, following relatively straight paths that were significantly directed over spatial scales exceeding 1600 m. In contrast, tracks of anosmic sharks ended 37.2% closer to shore, following significantly more tortuous paths that approximated correlated random walks. These results held after swimming paths were adjusted for current drift. This is the first study to demonstrate experimentally that olfaction contributes to pelagic navigation in sharks, likely mediated by chemical gradients as has been hypothesized for birds. Given the similarities between the fluid three-dimensional chemical atmosphere and ocean, further research comparing swimming and flying animals may lead to a unifying paradigm explaining their extraordinary navigational abilities.

  12. Spatial Variability of Benthic-Pelagic Coupling in an Estuary Ecosystem: Consequences for Microphytobenthos Resuspension Phenomenon

    PubMed Central

    Ubertini, Martin; Lefebvre, Sébastien; Gangnery, Aline; Grangeré, Karine; Le Gendre, Romain; Orvain, Francis

    2012-01-01

    The high degree of physical factors in intertidal estuarine ecosystem increases material processing between benthic and pelagic compartments. In these ecosystems, microphytobenthos resuspension is a major phenomenon since its contribution to higher trophic levels can be highly significant. Understanding the sediment and associated microphytobenthos resuspension and its fate in the water column is indispensable for measuring the food available to benthic and pelagic food webs. To identify and hierarchize the physical/biological factors potentially involved in MPB resuspension, the entire intertidal area and surrounding water column of an estuarine ecosystem, the Bay des Veys, was sampled during ebb tide. A wide range of physical parameters (hydrodynamic regime, grain size of the sediment, and suspended matter) and biological parameters (flora and fauna assemblages, chlorophyll) were analyzed to characterize benthic-pelagic coupling at the bay scale. Samples were collected in two contrasted periods, spring and late summer, to assess the impact of forcing variables on benthic-pelagic coupling. A mapping approach using kriging interpolation enabled us to overlay benthic and pelagic maps of physical and biological variables, for both hydrological conditions and trophic indicators. Pelagic Chl a concentration was the best predictor explaining the suspension-feeders spatial distribution. Our results also suggest a perennial spatio-temporal structure of both benthic and pelagic compartments in the ecosystem, at least when the system is not imposed to intense wind, with MPB distribution controlled by both grain size and bathymetry. The benthic component appeared to control the pelagic one via resuspension phenomena at the scale of the bay. Co-inertia analysis showed closer benthic-pelagic coupling between the variables in spring. The higher MPB biomass observed in summer suggests a higher contribution to filter-feeders diets, indicating a higher resuspension effect in

  13. Adaptation of pelage color and pigment variations in Israeli subterranean blind mole rats, Spalax ehrenbergi [corrected].

    PubMed

    Singaravelan, Natarajan; Raz, Shmuel; Tzur, Shay; Belifante, Shirli; Pavlicek, Tomas; Beiles, Avigdor; Ito, Shosuke; Wakamatsu, Kazumasa; Nevo, Eviatar

    2013-01-01

    Concealing coloration in rodents is well established. However, only a few studies examined how soil color, pelage color, hair-melanin content, and genetics (i.e., the causal chain) synergize to configure it. This study investigates the causal chain of dorsal coloration in Israeli subterranean blind mole rats, Spalax ehrenbergi. We examined pelage coloration of 128 adult animals from 11 populations belonging to four species of Spalax ehrenbergi superspecies (Spalax galili, Spalax golani, Spalax carmeli, and Spalax judaei) and the corresponding coloration of soil samples from the collection sites using a digital colorimeter. Additionally, we quantified hair-melanin contents of 67 animals using HPLC and sequenced the MC1R gene in 68 individuals from all four mole rat species. Due to high variability of soil colors, the correlation between soil and pelage color coordinates was weak and significant only between soil hue and pelage lightness. Multiple stepwise forward regression revealed that soil lightness was significantly associated with all pelage color variables. Pelage color lightness among the four species increased with the higher southward aridity in accordance to Gloger's rule (darker in humid habitats and lighter in arid habitats). Darker and lighter pelage colors are associated with darker basalt and terra rossa, and lighter rendzina soils, respectively. Despite soil lightness varying significantly, pelage lightness and eumelanin converged among populations living in similar soil types. Partial sequencing of the MC1R gene identified three allelic variants, two of which were predominant in northern species (S. galili and S. golani), and the third was exclusive to southern species (S. carmeli and S. judaei), which might have caused the differences found in pheomelanin/eumelanin ratio. Darker dorsal pelage in darker basalt and terra rossa soils in the north and lighter pelage in rendzina and loess soils in the south reflect the combined results of crypsis and

  14. Fossilized bioelectric wire - the trace fossil Trichichnus

    NASA Astrophysics Data System (ADS)

    Kędzierski, M.; Uchman, A.; Sawlowicz, Z.; Briguglio, A.

    2014-12-01

    The trace fossil Trichichnus is proposed as an indicator of fossil bioelectric bacterial activity at the interface oxic - anoxic zone of marine sediments. This fulfils the idea that such processes, commonly found in the modern realm, should be also present in the geological past. Trichichnus is an exceptional trace fossil due to its very thin diameter (mostly less than 1 mm) and common pyritic filling. It is ubiquitous in some fine-grained sediments, where it has been interpreted as a burrow formed deeper than any other trace fossils, below the redox boundary. Trichichnus formerly referred to as deeply burrowed invertebrates, has been found as remnant of a fossilized intrasediment bacterial mat that is pyritized. As visualized in 3-D by means of X-ray computed microtomography scanner, Trichichnus forms dense filamentous fabric, which reflects that produced by modern large, mat-forming, sulphide-oxidizing bacteria, belonging mostly to Trichichnus-related taxa, which are able to house a complex bacterial consortium. Several stages of Trichichnus formation, including filamentous, bacterial mat and its pyritization, are proposed to explain an electron exchange between oxic and suboxic/anoxic layers in the sediment. Therefore, Trichichnus can be considered a fossilized "electric wire".

  15. Fossilized bioelectric wire - the trace fossil Trichichnus

    NASA Astrophysics Data System (ADS)

    Kędzierski, M.; Uchman, A.; Sawlowicz, Z.; Briguglio, A.

    2015-04-01

    The trace fossil Trichichnus is proposed as an indicator of fossil bioelectric bacterial activity at the oxic-anoxic interface zone of marine sediments. This fulfils the idea that such processes, commonly found in the modern realm, should be also present in the geological past. Trichichnus is an exceptional trace fossil due to its very thin diameter (mostly less than 1 mm) and common pyritic filling. It is ubiquitous in some fine-grained sediments, where it has been interpreted as a burrow formed deeper than any other trace fossils, below the redox boundary. Trichichnus, formerly referred to as deeply burrowed invertebrates, has been found as remnant of a fossilized intrasediment bacterial mat that is pyritized. As visualized in 3-D by means of X-ray computed microtomography scanner, Trichichnus forms dense filamentous fabric, which reflects that it is produced by modern large, mat-forming, sulfide-oxidizing bacteria, belonging mostly to Thioploca-related taxa, which are able to house a complex bacterial consortium. Several stages of Trichichnus formation, including filamentous, bacterial mat and its pyritization, are proposed to explain an electron exchange between oxic and suboxic/anoxic layers in the sediment. Therefore, Trichichnus can be considered a fossilized "electric wire".

  16. Pelagic nekton abundance and distribution in the northern Sacramento–San Joaquin Delta, California

    USGS Publications Warehouse

    Feyrer, Frederick; Slater, Steven B.; Portz, Donald E.; Odom, Darren; Morgan-King, Tara L.; Brown, Larry R.

    2017-01-01

    Knowledge of the habitats occupied by species is fundamental for the development of effective conservation and management actions. The collapse of pelagic fish species in the Sacramento–San Joaquin Delta, California, has triggered a need to better understand factors that drive their distribution and abundance. A study was conducted in summer–fall 2014 in an attempt to identify physical and biological habitat conditions that drive the abundance and distribution of pelagic species in the northern region of the system. The study was conducted in the three largest channels in the northern Sacramento–San Joaquin Delta by dimension, volume, and flow capacity. The pelagic community was dominated by three nonnative species, Siberian prawn Exopalaemon modestus, which comprised 56% of the total number of organisms, and two fish species, Threadfin Shad Dorosoma petenense and Mississippi Silversides Menidia audens, which together comprised 43% of the total number of organisms. Total fish and total shrimp abundance were sensitive to the most extreme values of turbidity and temperature encountered and positively associated with total zooplankton biomass. The results suggested that habitat conditions in terminal channels, historically a common feature on the landscape, support higher abundances of pelagic species and zooplankton than open-ended channels. These results provide resource managers with useful information on the habitat associations of pelagic species and on how the future distribution and abundance of pelagic species will likely change in response to climate or other ecological factors.

  17. Ecological Insights from Pelagic Habitats Acquired Using Active Acoustic Techniques.

    PubMed

    Benoit-Bird, Kelly J; Lawson, Gareth L

    2016-01-01

    Marine pelagic ecosystems present fascinating opportunities for ecological investigation but pose important methodological challenges for sampling. Active acoustic techniques involve producing sound and receiving signals from organisms and other water column sources, offering the benefit of high spatial and temporal resolution and, via integration into different platforms, the ability to make measurements spanning a range of spatial and temporal scales. As a consequence, a variety of questions concerning the ecology of pelagic systems lend themselves to active acoustics, ranging from organism-level investigations and physiological responses to the environment to ecosystem-level studies and climate. As technologies and data analysis methods have matured, the use of acoustics in ecological studies has grown rapidly. We explore the continued role of active acoustics in addressing questions concerning life in the ocean, highlight creative applications to key ecological themes ranging from physiology and behavior to biogeography and climate, and discuss emerging avenues where acoustics can help determine how pelagic ecosystems function.

  18. Abundance and tidal behaviour of pelagic fish in the gateway to the Wadden Sea

    NASA Astrophysics Data System (ADS)

    Couperus, Bram; Gastauer, Sven; Fässler, Sascha M. M.; Tulp, Ingrid; van der Veer, Henk W.; Poos, Jan Jaap

    2016-03-01

    The shallow coast of The Netherlands is an important habitat for small pelagic fish. They form one of the major links between plankton and the higher trophic levels. Predatory fish, sea mammals and birds rely on small pelagic fish as a major food source. Currently, monitoring of fish in the Dutch coastal zone mainly focuses on demersal species, using bottom trawls and fykes. Four hydro-acoustic surveys were carried out in May and October 2010/2011 in the Marsdiep area, a relatively deep tidal inlet in the western Wadden Sea, to quantify abundances of pelagic fish. The aims of this study were to (1) describe temporal and vertical variations in fish distribution and school dimensions in relation to tide, and (2) estimate biomass of pelagic fish and their proportion to total fish biomass. The biomass of pelagic fish in the Marsdiep area ranged between 23 and 411 kg/ha. These were mainly sprat, but also young herring, anchovy and pilchard. The fish was scattered in small schools with volumes smaller than 5m3 and concentrated in the top 10 m below the surface. There was a clear effect of tidal cycle on school volume and fish abundance, with larger densities and larger schools at high tide compared to low tide. In May, sandeel contributed substantially to the pelagic assemblage, whereas in October sandeel was absent in the trawl catches, most likely because they stayed buried in the seabed from late summer to spring. The presence of pilchard and anchovy confirmed their re-establishment in the Southern North Sea and Wadden Sea. The abundance of pelagic fish exceeded the biomass of demersal fish in the western Wadden Sea by an order of magnitude. This finding is relevant for ecosystem studies. The fact that this study suggests that small pelagics outnumber demersal species to such a large extent calls for a rethinking of the allocation of monitoring effort in the Dutch coastal zone.

  19. Marquee Fossils

    ERIC Educational Resources Information Center

    Clary, Renee; Wandersee, James

    2008-01-01

    Professors of an online graduate-level paleontology class developed the concept of marquee fossils--fossils that have one or more unique characteristics that capture the attention and direct observation of students. In the classroom, Marquee fossils integrate the geology, biology, and environmental science involved in the study of fossilized…

  20. Paleoecology of the Devonian-Mississippian black-shale sequence in eastern Kentucky with an atlas of some common fossils

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Barron, L.S.; Ettensohn, F.R.

    The Devonian-Mississippian black-shale sequence of eastern North America is a distinctive stratigraphic interval generally characterized by low clastic influx, high organic production in the water column, anaerobic bottom conditions, and the relative absence of fossil evidence for biologic activity. The laminated black shales which constitute most of the black-shale sequence are broken by two major sequences of interbedded greenish-gray, clayey shales which contain bioturbation and pyritized micromorph invertebrates. The black shales contain abundant evidence of life from upper parts of the water column such as fish fossils, conodonts, algae and other phytoplankton; however, there is a lack of evidence ofmore » benthic life. The rare brachiopods, crinoids, and molluscs that occur in the black shales were probably epiplanktic. A significant physical distinction between the environment in which the black sediments were deposited and that in which the greenish-gray sediments were deposited was the level of dissolved oxygen. The laminated black shales point to anaerobic conditions and the bioturbated greenish-gray shales suggest dysaerobic to marginally aerobic-dysaerobic conditions. A paleoenvironmental model in which quasi-estuarine circulation compliments and enhances the effect of a stratified water column can account for both depletion of dissolved oxygen in the bottom environments and the absence of oxygen replenishment during black-shale deposition. Periods of abundant clastic influx from fluvial environments to the east probably account for the abundance of clays in the greenish-gray shale as well as the small amounts of oxygen necessary to support the depauparate, opportunistic, benthic faunas found there. These pulses of greenish-gray clastics were short-lived and eventually were replaced by anaerobic conditions and low rates of clastic sedimentation which characterized most of black-shale deposition.« less

  1. Fossil Explorers

    ERIC Educational Resources Information Center

    Moran, Sean; McLaughlin, Cheryl; MacFadden, Bruce; Jacobbe, Elizabeth; Poole, Michael

    2015-01-01

    Many young learners are fascinated with fossils, particularly charismatic forms such as dinosaurs and giant sharks. Fossils provide tangible, objective evidence of life that lived millions of years ago. They also provide a timescale of evolution not typically appreciated by young learners. Fossils and the science of paleontology can, therefore,…

  2. Longitudinal differentiation among pelagic populations in a planktic foraminifer

    PubMed Central

    Ujiié, Yurika; Asami, Takahiro; de Garidel-Thoron, Thibault; Liu, Hui; Ishitani, Yoshiyuki; de Vargas, Colomban

    2012-01-01

    Evolutionary processes in marine plankton have been assumed to be dependent on the oceanic circulation system, which transports plankton between populations in marine surface waters. Gene flow facilitated by oceanic currents along longitudinal gradients may efficiently impede genetic differentiation of pelagic populations in the absence of confounding marine environmental effects. However, how responsible oceanic currents are for the geographic distribution and dispersal of plankton is poorly understood. We examined the phylogeography of the planktic foraminifer Pulleniatina obliquiloculata in the Indo-Pacific Warm Pool (IPWP) by using partial small subunit ribosomal DNA (SSU rDNA) sequences. We found longitudinal clines in the frequencies of three distinct genetic types in the IPWP area. These frequencies were correlated with environmental factors that are characteristic of three water masses in the IPWP. Noteworthy, populations inhabiting longitudinally distant water masses at the Pacific and Indian sides of the IPWP were genetically different, despite transportation of individuals via oceanic currents. These results demonstrate that populations of pelagic plankton have diverged genetically among different water masses within a single climate zone. Changes of the oceanic circulation system could have impacted the geographic patterns of dispersal and divergence of pelagic plankton. PMID:22957176

  3. Radioactivity in fossils at the Hagerman Fossil Beds National Monument.

    PubMed

    Farmer, C Neal; Kathren, Ronald L; Christensen, Craig

    2008-08-01

    Since 1996, higher than background levels of naturally occurring radioactivity have been documented in both fossil and mineral deposits at Hagerman Fossil Beds National Monument in south-central Idaho. Radioactive fossil sites occur primarily within an elevation zone of 900-1000 m above sea level and are most commonly found associated with ancient river channels filled with sand. Fossils found in clay rich deposits do not exhibit discernable levels of radioactivity. Out of 300 randomly selected fossils, approximately three-fourths exhibit detectable levels of natural radioactivity ranging from 1 to 2 orders of magnitude above ambient background levels when surveyed with a portable hand held Geiger-Muller survey instrument. Mineral deposits in geologic strata also show above ambient background levels of radioactivity. Radiochemical lab analysis has documented the presence of numerous natural radioactive isotopes. It is postulated that ancient groundwater transported radioactive elements through sand bodies containing fossils which precipitated out of solution during the fossilization process. The elevated levels of natural radioactivity in fossils may require special precautions to ensure that exposures to personnel from stored or displayed items are kept as low as reasonably achievable (ALARA).

  4. Fossilized bioelectric wire – the trace fossil Trichichnus

    PubMed Central

    Kędzierski, M.; Uchman, A.; Sawlowicz, Z.; Briguglio, A.

    2015-01-01

    The trace fossil Trichichnus is proposed as an indicator of fossil bioelectric bacterial activity at the oxic–anoxic interface zone of marine sediments. This fulfils the idea that such processes, commonly found in the modern realm, should be also present in the geological past. Trichichnus is an exceptional trace fossil due to its very thin diameter (mostly less than 1 mm) and common pyritic filling. It is ubiquitous in some fine-grained sediments, where it has been interpreted as a burrow formed deeper than any other trace fossils, below the redox boundary. Trichichnus, formerly referred to as deeply burrowed invertebrates, has been found as remnant of a fossilized intrasediment bacterial mat that is pyritized. As visualized in 3-D by means of X-ray computed microtomography scanner, Trichichnus forms dense filamentous fabric, which reflects that it is produced by modern large, mat-forming, sulfide-oxidizing bacteria, belonging mostly to Thioploca-related taxa, which are able to house a complex bacterial consortium. Several stages of Trichichnus formation, including filamentous, bacterial mat and its pyritization, are proposed to explain an electron exchange between oxic and suboxic/anoxic layers in the sediment. Therefore, Trichichnus can be considered a fossilized “electric wire”. PMID:26290671

  5. Trace elements in oceanic pelagic communities in the western Indian Ocean.

    PubMed

    Bodin, Nathalie; Lesperance, Dora; Albert, Rona; Hollanda, Stephanie; Michaud, Philippe; Degroote, Maxime; Churlaud, Carine; Bustamante, Paco

    2017-05-01

    The mineral composition of target and non-target pelagic fish caught by purse-seiners and longliners in the western-central Indian Ocean was determined. From the 10 essential elements analysed, selenium and zinc showed the highest concentrations in swordfish and blue marlin while Indian mackerel appeared as a good source of copper, iron and chrome. All catch had levels of lead and cadmium, two toxic elements, below the maximum sanitary limits. Although some concerns were raised regarding mercury concentrations in the largest species (wahoo, swordfish and blue marlin), molar ratios of mercury and selenium indicate that all oceanic pelagic fish from the western-central Indian Ocean are safe for human consumption. This study also gives insights on the relationships between the levels of essential and toxic elements in fish muscle and the size, trophic position and diet sources of the studied pelagic species. Copyright © 2017 Elsevier Ltd. All rights reserved.

  6. The role of the male copulatory organs in the colonization of the pelagic by shrimp-like eucarids

    NASA Astrophysics Data System (ADS)

    Lunina, Anastasia; Vereshchaka, Alexander

    2017-03-01

    Five major taxa of shrimp-like eucarid are recognized: the order Euphausiacea, the suborder Dendrobranchiata, and three infraorders of the suborder Pleocyemata (Caridea, Procarididea, and Stenopodidea). These animals are very successful in the colonization of the pelagic but the role of the sexual structures (the appendix masculina and the petasma) in this process remains uncertain. We revise the vertical distribution of pelagic species and genera within the shrimp-like eucarids and the presence and types of the male copulatory organs. We further test the hypothesis that the pelagic habitat is associated with the elaborated male copulatory organs and discuss possible mechanisms of association. We also consider possible effects of the depth and coastal factors. In the turbulent and fluid pelagic, successful copulation depends on the perfect fixation and possible stimulation of mates during spermatophore transfer and thus on the development and elaboration of the copulatory structures. The presence of the appendix masculina is necessary for copulation in the pelagic and thus for colonization of the pelagic. The 'family success' in colonizing the pelagic depends on the presence of the elaborate petasma, whilst the 'species success' is not so obviously related to the presence of this organ. The colonization of the water column by the eucarids may basically occur at the family level and be associated with the elaboration of the petasma. Within monophyletic pelagic groups, a greater development of the petasma is observed in the epi- and mesopelagic; some reduction of this organ in the bathypelagic may be related to the reduction in current velocities and turbulence. Simplification of the petasma is also observed in coastal waters. The proposed hypothesis is invariant to phylogenetic models and suggests an explanation to why (but not in which consequence) the elaborated copulative organs have been developed/retained in the pelagic or reduced/lost in the benthic.

  7. Vertical ecology of the pelagic ocean: classical patterns and new perspectives.

    PubMed

    Sutton, T T

    2013-12-01

    Applications of acoustic and optical sensing and intensive, discrete-depth sampling, in concert with collaborative international research programmes, have substantially advanced knowledge of pelagic ecosystems in the 17 years since the 1996 Deepwater Fishes Symposium of the Fisheries Society of the British Isles. Although the epipelagic habitat is the best-known, and remote sensing and high-resolution modelling allow near-synoptic investigation of upper layer biophysical dynamics, ecological studies within the mesopelagic and deep-demersal habitats have begun to link lower and upper trophic level processes. Bathypelagic taxonomic inventories are far from complete, but recent projects (e.g. MAR-ECO and CMarZ, supported by the Census of Marine Life programme) have quantitatively strengthened distribution patterns previously described for fishes and have provided new perspectives. Synthesis of net and acoustic studies suggests that the biomass of deep-pelagic fishes may be two to three orders of magnitude greater than the total global commercial fisheries landings. Discrete-depth net sampling has revealed relatively high pelagic fish biomass below 1000 m in some regions, and that gelatinous zooplankton may be key energy vectors for deep-pelagic fish production. Lastly, perhaps, the most substantive paradigm shift is that vertical connectivity among fishes across classical depth zones is prevalent- suggesting that a whole-water column approach is warranted for deep ocean conservation and management. © 2013 The Fisheries Society of the British Isles.

  8. Feeding strategies and ecological roles of three predatory pelagic fish in the western Mediterranean Sea

    NASA Astrophysics Data System (ADS)

    Navarro, Joan; Sáez-Liante, Raquel; Albo-Puigserver, Marta; Coll, Marta; Palomera, Isabel

    2017-06-01

    Knowing the feeding ecology of marine predators is pivotal to developing an understanding of their ecological role in the ecosystem and determining the trophic relationships between them. Despite the ecological importance of predatory pelagic fish species, research on these species in the Mediterranean Sea is limited. Here, by combining analyses of stomach contents and stable isotope values, we examined the feeding strategies of swordfish, Xiphias gladius, little tunny, Euthynnus alletteratus and Atlantic bonito, Sarda sarda, in the western Mediterranean Sea. We also compared the trophic niche and trophic level of these species with published information of other sympatric pelagic predators present in the ecosystem. Results indicated that, although the diet of the three species was composed mainly by fin-fish species, a clear segregation in their main feeding strategies was found. Swordfish showed a generalist diet including demersal species such as blue whiting, Micromesistius poutassou, and European hake, Merluccius merluccius, and pelagic fin-fish such as barracudina species (Arctozenus risso and Lestidiops jayakari) or small pelagic fish species. Little tunny and Atlantic bonito were segregated isotopically between them and showed a diet basically composed of anchovy, Engraulis encrasicolus, and round sardinella, Sardinella aurita, and sardines, Sardina pilchardus, respectively. This trophic segregation, in addition to potential segregation by depth, is likely a mechanism that allows their potential coexistence within the same pelagic habitat. When the trophic position of these three predatory pelagic fish species is compared with other pelagic predators such as bluefin tuna, Thunnus thynnus, and dolphinfish, Coryphaena hippurus, present in the western Mediterranean Sea, we found that they show similar intermediate trophic position in the ecosystem. In conclusion, the combined stomach and isotopic results highlight, especially for little tunny and Atlantic

  9. 50 CFR 622.373 - Limited access system for charter vessel/headboat permits for Gulf coastal migratory pelagic fish.

    Code of Federal Regulations, 2013 CFR

    2013-10-01

    .../headboat permits for Gulf coastal migratory pelagic fish. 622.373 Section 622.373 Wildlife and Fisheries... for Gulf coastal migratory pelagic fish. (a) No applications for additional charter vessel/headboat permits for Gulf coastal migratory pelagic fish will be accepted. Existing permits may be renewed, are...

  10. 50 CFR 622.373 - Limited access system for charter vessel/headboat permits for Gulf coastal migratory pelagic fish.

    Code of Federal Regulations, 2014 CFR

    2014-10-01

    .../headboat permits for Gulf coastal migratory pelagic fish. 622.373 Section 622.373 Wildlife and Fisheries... for Gulf coastal migratory pelagic fish. (a) No applications for additional charter vessel/headboat permits for Gulf coastal migratory pelagic fish will be accepted. Existing permits may be renewed, are...

  11. Impacts of gold mine waste disposal on a tropical pelagic ecosystem.

    PubMed

    Brewer, D T; Morello, E B; Griffiths, S; Fry, G; Heales, D; Apte, S C; Venables, W N; Rothlisberg, P C; Moeseneder, C; Lansdell, M; Pendrey, R; Coman, F; Strzelecki, J; Jarolimek, C V; Jung, R F; Richardson, A J

    2012-12-01

    We used a comparative approach to investigate the impact of the disposal of gold mine tailings into the ocean near the Lihir mine (Niolam Island, Papua New Guinea). We found abundance and diversity of zooplankton, micronekton and pelagic fish to be similar or higher in the mine region compared to the reference site. We also found relatively high trace metal concentrations in lower trophic level groups, especially zooplankton, near the mine discharge, but few differences in tissue concentrations of micronekton, baitfish and pelagic fish between the two regions. Biomagnification of some trace metals by micronekton, and of mercury by fish was evident in both regions. We conclude that ocean mine waste disposal at Niolam Island has a local impact on the smaller and less mobile pelagic communities in terms of trace metal concentrations, but has little effect on the abundance and biodiversity of the local food web. Crown Copyright © 2012. Published by Elsevier Ltd. All rights reserved.

  12. Pelagic Biocarbonates: Assessing the "Forgotten" Fine Fraction

    NASA Astrophysics Data System (ADS)

    Brummer, G. J. A.

    2016-02-01

    Biocarbonates play an important role in the global carbon cycle and cover over half of the ocean floor. Biocarbonates in the open ocean are best known from planktonic foraminifera, which are relatively large (>150µm), heavy and few and coccoliths, which are very small (<32µm), light and abundant. Both of these components are relatively well studied. The size fraction in between adult foraminifera and coccoliths (32-150µm: the so-called fine fraction) consists of a large but poorly known mixture of particles, which is genarlly assumed to consist primarily of "juvenile" planktonic foraminifera, with minor amounts of calcareous dinoflagellates and various others less well-known microfossils. Abundance, diversity, mass and composition within the fine fraction are poorly constrained, as is the response to acidification/dissolution. This lack of knowledge primarily reflects the gap in size fraction studied by the different disciplinary approaches and techniques, which are not suited for identifying and quantifying these intermediate groups. Comparative ontogeny of planktonic foraminifera now shows that this fine fraction in sediments does not consist of "juveniles" as in the living plankton, but is dominated by mature specimens of small-sized species. First estimates indicate that these small species not only account for about one third of the number of species of planktonic foraminifera but also form about one third of their shell flux and global carbonate production in weight. Still, we hardly know anything on seasonality, depth habitat, shell composition (isotopes, trace metal incorporation), potential autotrophic symbionts, molecular genetics and geological range of these clearly very important species. Estimates from well-preserved sediments, show that the important role of these minute foraminiferal planktonic species may hold for much of the 180Ma long fossil record, opening a new research field pertaining to both modern and past pelagic ecosystems and the role

  13. Stable pelagic vertebrate community structure through extreme Paleogene greenhouse conditions

    NASA Astrophysics Data System (ADS)

    Sibert, E. C.; Friedman, M.; Hull, P. M.; Hunt, G.; Norris, R. D.

    2016-02-01

    The species composition (structure) and energy transfer (function) of an ecosystem is reflected by the presence and type of consumers that it supports. Here we use ichthyoliths, microfossil fish teeth and shark denticles, to assess the ecological variability of the pelagic fish community structure and composition from the Late Cretaceous to the middle Eocene from a drill core in the South Pacific gyre (DSDP Site 596). We find that the overall vertebrate community structure, as measured by the relative abundance of sharks to ray-finned fishes, has a punctuated change at the Cretaceous/Paleogene mass extinction. The vertebrate community structure remained stable throughout the Paleogene despite a five-fold increase in overall abundance of ichthyoliths during the extreme greenhouse of the Early Eocene. Further, we use a novel system to quantify the morphological variation in fish teeth. We find that the morphospace occupied by the tooth assemblage is conserved throughout the interval, with a slight expansion following the Cretaceous-Paleogene mass extinction, and the evolution of a distinct morphotype-group around the Paleocene-Eocene boundary. While there are elevated rates of morphotype origination and extinction following the Cretaceous-Paleogene mass extinction, the extreme greenhouse warming of the Early Eocene and associated increase in fish production produce near-zero origination and extinction rates. The relative stability in composition of the pelagic vertebrate community during intervals of extreme climate change and across large ranges of total fish accumulation, suggests that pelagic ecosystem structure is robust to climate events, and that the overall structure of the pelagic fish community may be decoupled from both climate and ecosystem function.

  14. "Fossil" Forecasting.

    ERIC Educational Resources Information Center

    Brody, Michael J.; deOnis, Ann

    2001-01-01

    Presents a density study in which students calculate the density of limestone substrate to determine if the specimen contains any fossils. Explains how to make fossils and addresses national standards. (YDS)

  15. Beyond the vent: New perspectives on hydrothermal plumes and pelagic biology

    NASA Astrophysics Data System (ADS)

    Phillips, Brennan T.

    2017-03-01

    Submarine hydrothermal vent fields introduce buoyant plumes of chemically altered seawater to the deep-sea water column. Chemoautotrophic microbes exploit this energy source, facilitating seafloor-based primary production that evidence suggests may transfer to pelagic consumers. While most hydrothermal plumes have relatively small volumes, there are recent examples of large-scale plume events associated with periods of eruptive activity, which have had a pronounced effect on water-column biology. This correlation suggests that hydrothermal plumes may have influenced basin-scale ocean chemistry during periods of increased submarine volcanism during the Phanerozoic eon. This paper synthesizes a growing body of scientific evidence supporting the hypothesis that hydrothermal plumes are the energetic basis of unique deep-sea pelagic food webs. While many important questions remain concerning the biology of hydrothermal plumes, this discussion is not present in ongoing management efforts related to seafloor massive sulfide (SMS) mining. Increased research efforts, focused on high-resolution surveys of midwater biology relative to plume structures, are recommended to establish baseline conditions and monitor the impact of future mining-based disturbances to the pelagic biosphere.

  16. Increased oceanic microplastic debris enhances oviposition in an endemic pelagic insect.

    PubMed

    Goldstein, Miriam C; Rosenberg, Marci; Cheng, Lanna

    2012-10-23

    Plastic pollution in the form of small particles (diameter less than 5 mm)--termed 'microplastic'--has been observed in many parts of the world ocean. They are known to interact with biota on the individual level, e.g. through ingestion, but their population-level impacts are largely unknown. One potential mechanism for microplastic-induced alteration of pelagic ecosystems is through the introduction of hard-substrate habitat to ecosystems where it is naturally rare. Here, we show that microplastic concentrations in the North Pacific Subtropical Gyre (NPSG) have increased by two orders of magnitude in the past four decades, and that this increase has released the pelagic insect Halobates sericeus from substrate limitation for oviposition. High concentrations of microplastic in the NPSG resulted in a positive correlation between H. sericeus and microplastic, and an overall increase in H. sericeus egg densities. Predation on H. sericeus eggs and recent hatchlings may facilitate the transfer of energy between pelagic- and substrate-associated assemblages. The dynamics of hard-substrate-associated organisms may be important to understanding the ecological impacts of oceanic microplastic pollution.

  17. Unexpected hydrogen isotope variation in oceanic pelagic seabirds

    USGS Publications Warehouse

    Ostrom, Peggy H.; Wiley, Anne E.; Rossman, Sam; Stricker, Craig A.; James, Helen F.

    2014-01-01

    Hydrogen isotopes have significantly enhanced our understanding of the biogeography of migratory animals. The basis for this methodology lies in predictable, continental patterns of precipitation δD values that are often reflected in an organism's tissues. δD variation is not expected for oceanic pelagic organisms whose dietary hydrogen (water and organic hydrogen in prey) is transferred up the food web from an isotopically homogeneous water source. We report a 142% range in the δD values of flight feathers from the Hawaiian petrel (Pterodroma sandwichensis), an oceanic pelagic North Pacific species, and inquire about the source of that variation. We show δD variation between and within four other oceanic pelagic species: Newell's shearwater (Puffinus auricularis newellii), Black-footed albatross (Phoebastria nigripes), Laysan albatross (Phoebastria immutabilis) and Buller's shearwater (Puffinus bulleri). The similarity between muscle δD values of hatch-year Hawaiian petrels and their prey suggests that trophic fractionation does not influence δD values of muscle. We hypothesize that isotopic discrimination is associated with water loss during salt excretion through salt glands. Salt load differs between seabirds that consume isosmotic squid and crustaceans and those that feed on hyposmotic teleost fish. In support of the salt gland hypothesis, we show an inverse relationship between δD and percent teleost fish in diet for three seabird species. Our results demonstrate the utility of δD in the study of oceanic consumers, while also contributing to a better understanding of δD systematics, the basis for one of the most commonly utilized isotope tools in avian ecology.

  18. Warming shelf seas drive the subtropicalization of European pelagic fish communities.

    PubMed

    Montero-Serra, Ignasi; Edwards, Martin; Genner, Martin J

    2015-01-01

    Pelagic fishes are among the most ecologically and economically important fish species in European seas. In principle, these pelagic fishes have potential to demonstrate rapid abundance and distribution shifts in response to climatic variability due to their high adult motility, planktonic larval stages, and low dependence on benthic habitat for food or shelter during their life histories. Here, we provide evidence of substantial climate-driven changes to the structure of pelagic fish communities in European shelf seas. We investigated the patterns of species-level change using catch records from 57,870 fisheries-independent survey trawls from across European continental shelf region between 1965 and 2012. We analysed changes in the distribution and rate of occurrence of the six most common species, and observed a strong subtropicalization of the North Sea and Baltic Sea assemblages. These areas have shifted away from cold-water assemblages typically characterized by Atlantic herring and European sprat from the 1960s to 1980s, to warmer-water assemblages including Atlantic mackerel, Atlantic horse mackerel, European pilchard and European anchovy from the 1990s onwards. We next investigated if warming sea temperatures have forced these changes using temporally comprehensive data from the North Sea region. Our models indicated the primary driver of change in these species has been sea surface temperatures in all cases. Together, these analyses highlight how individual species responses have combined to result in a dramatic subtropicalization of the pelagic fish assemblage of the European continental shelf. © 2014 John Wiley & Sons Ltd.

  19. Modes of fossil preservation

    USGS Publications Warehouse

    Schopf, J.M.

    1975-01-01

    The processes of geologic preservation are important for understanding the organisms represented by fossils. Some fossil differences are due to basic differences in organization of animals and plants, but the interpretation of fossils has also tended to be influenced by modes of preservation. Four modes of preservation generally can be distinguished: (1) Cellular permineralization ("petrifaction") preserves anatomical detail, and, occasionally, even cytologic structures. (2) Coalified compression, best illustrated by structures from coal but characteristic of many plant fossils in shale, preserves anatomical details in distorted form and produces surface replicas (impressions) on enclosing matrix. (3) Authigenic preservation replicates surface form or outline (molds and casts) prior to distortion by compression and, depending on cementation and timing, may intergrade with fossils that have been subject to compression. (4) Duripartic (hard part) preservation is characteristic of fossil skeletal remains, predominantly animal. Molds, pseudomorphs, or casts may form as bulk replacements following dissolution of the original fossil material, usually by leaching. Classification of the kinds of preservation in fossils will aid in identifying the processes responsible for modifying the fossil remains of both animals and plants. ?? 1975.

  20. Enamel ultrastructure in fossil cetaceans (Cetacea: Archaeoceti and Odontoceti).

    PubMed

    Loch, Carolina; Kieser, Jules A; Fordyce, R Ewan

    2015-01-01

    The transition from terrestrial ancestry to a fully pelagic life profoundly altered the body systems of cetaceans, with extreme morphological changes in the skull and feeding apparatus. The Oligocene Epoch was a crucial time in the evolution of cetaceans when the ancestors of modern whales and dolphins (Neoceti) underwent major diversification, but details of dental structure and evolution are poorly known for the archaeocete-neocete transition. We report the morphology of teeth and ultrastructure of enamel in archaeocetes, and fossil platanistoids and delphinoids, ranging from late Oligocene (Waitaki Valley, New Zealand) to Pliocene (Caldera, Chile). Teeth were embedded in epoxy resin, sectioned in cross and longitudinal planes, polished, etched, and coated with gold palladium for scanning electron microscopy (SEM) observation. SEM images showed that in archaeocetes, squalodontids and Prosqualodon (taxa with heterodont and nonpolydont/limited polydont teeth), the inner enamel was organized in Hunter-Schreger bands (HSB) with an outer layer of radial enamel. This is a common pattern in most large-bodied mammals and it is regarded as a biomechanical adaptation related to food processing and crack resistance. Fossil Otekaikea sp. and delphinoids, which were polydont and homodont, showed a simpler structure, with inner radial and outer prismless enamel. Radial enamel is regarded as more wear-resistant and has been retained in several mammalian taxa in which opposing tooth surfaces slide over each other. These observations suggest that the transition from a heterodont and nonpolydont/limited polydont dentition in archaeocetes and early odontocetes, to homodont and polydont teeth in crownward odontocetes, was also linked to a marked simplification in the enamel Schmelzmuster. These patterns probably reflect functional shifts in food processing from shear-and-mastication in archaeocetes and early odontocetes, to pierce-and-grasp occlusion in crownward odontocetes, with

  1. Enamel Ultrastructure in Fossil Cetaceans (Cetacea: Archaeoceti and Odontoceti)

    PubMed Central

    Loch, Carolina; Fordyce, R. Ewan

    2015-01-01

    The transition from terrestrial ancestry to a fully pelagic life profoundly altered the body systems of cetaceans, with extreme morphological changes in the skull and feeding apparatus. The Oligocene Epoch was a crucial time in the evolution of cetaceans when the ancestors of modern whales and dolphins (Neoceti) underwent major diversification, but details of dental structure and evolution are poorly known for the archaeocete-neocete transition. We report the morphology of teeth and ultrastructure of enamel in archaeocetes, and fossil platanistoids and delphinoids, ranging from late Oligocene (Waitaki Valley, New Zealand) to Pliocene (Caldera, Chile). Teeth were embedded in epoxy resin, sectioned in cross and longitudinal planes, polished, etched, and coated with gold palladium for scanning electron microscopy (SEM) observation. SEM images showed that in archaeocetes, squalodontids and Prosqualodon (taxa with heterodont and nonpolydont/limited polydont teeth), the inner enamel was organized in Hunter-Schreger bands (HSB) with an outer layer of radial enamel. This is a common pattern in most large-bodied mammals and it is regarded as a biomechanical adaptation related to food processing and crack resistance. Fossil Otekaikea sp. and delphinoids, which were polydont and homodont, showed a simpler structure, with inner radial and outer prismless enamel. Radial enamel is regarded as more wear-resistant and has been retained in several mammalian taxa in which opposing tooth surfaces slide over each other. These observations suggest that the transition from a heterodont and nonpolydont/limited polydont dentition in archaeocetes and early odontocetes, to homodont and polydont teeth in crownward odontocetes, was also linked to a marked simplification in the enamel Schmelzmuster. These patterns probably reflect functional shifts in food processing from shear-and-mastication in archaeocetes and early odontocetes, to pierce-and-grasp occlusion in crownward odontocetes, with

  2. Similarity principles for the biology of pelagic animals

    PubMed Central

    Barenblatt, G. I.; Monin, A. S.

    1983-01-01

    A similarity principle is formulated according to which the statistical pattern of the pelagic population is identical in all scales sufficiently large in comparison with the molecular one. From this principle, a power law is obtained analytically for the pelagic animal biomass distribution over the animal sizes. A hypothesis is presented according to which, under fixed external conditions, the oxygen exchange intensity of an animal is governed only by its mass and density and by the specific absorbing capacity of the animal's respiratory organ. From this hypothesis a power law is obtained by the method of dimensional analysis for the exchange intensity mass dependence. The known empirical values of the exponent of this power law are interpreted as an indication that the oxygen-absorbing organs of the animals can be represented as so-called fractal surfaces. In conclusion the biological principle of the decrease in specific exchange intensity with increase in animal mass is discussed. PMID:16593327

  3. Life cycle ecophysiology of small pelagic fish and climate-driven changes in populations

    NASA Astrophysics Data System (ADS)

    Peck, Myron A.; Reglero, Patricia; Takahashi, Motomitsu; Catalán, Ignacio A.

    2013-09-01

    Due to their population characteristics and trophodynamic role, small pelagic fishes are excellent bio-indicators of climate-driven changes in marine systems world-wide. We argue that making robust projections of future changes in the productivity and distribution of small pelagics will require a cause-and-effect understanding of historical changes based upon physiological principles. Here, we reviewed the ecophysiology of small pelagic (clupeiform) fishes including a matrix of abiotic and biotic extrinsic factors (e.g., temperature, salinity, light, and prey characteristics) and stage-specific vital rates: (1) adult spawning, (2) survival and development of eggs and yolk sac larvae, and (3) feeding and growth of larvae, post-larvae and juveniles. Emphasis was placed on species inhabiting Northwest Pacific and Northeast Atlantic (European) waters for which summary papers are particularly scarce compared to anchovy and sardine in upwelling systems. Our review revealed that thermal niches (optimal and sub-optimal ranges in temperatures) were species- and stage-specific but that temperature effects only partly explained observed changes in the distribution and/or productivity of populations in the Northwest Pacific and Northeast Atlantic; changes in temperature may be necessary but not sufficient to induce population-level shifts. Prey availability during the late larval and early juvenile period was a common, density-dependent mechanism linked to fluctuations in populations but recruitment mechanisms were system-specific suggesting that generalizations of climate drivers across systems should be avoided. We identified gaps in knowledge regarding basic elements of the growth physiology of each life stage that will require additional field and laboratory study. Avenues of research are recommended that will aid the development of models that provide more robust, physiological-based projections of the population dynamics of these and other small pelagic fish. In our

  4. Reported middle Paleozoic fossils and new geochronological data from the southern and central Appalachians: Disposable outrageous hypothesis or justification for major revision of tectonic history

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Hatcher, R.D. Jr.

    Recently published interpretations of fossil fragments from the Walden Creek Group (Ocoee Supergroup) suggesting that these rocks are middle Paleozoic (Devonian to Early Carboniferous), and new geochronological data that yield late Paleozoic age dates on rocks and major faults in the Blue Ridge and piedmont, if taken alone, would permit speculation that most of the deformation and metamorphism affecting this part of the orogen is Alleghanian. The two Ordovician clastic wedges (Sevier, Llanvirn, and Martinsburg, Caradoc-Ashgill) and the Carboniferous-Permian wedge(s), along with many radiometric ages on plutons, indicate uplift and sediment dispersal from the interior of the southern and centralmore » Appalachians (SCA) that may have resulted from Taconian and Alleghanian deformation. Combining the reproducible fossil evidence, including that from Alabama and a recently discovered crinoid fragment from the upper part of the Murphy belt sequence, with the most current geochronological data requires that peak metamorphism and penetrative deformation be at least Devonian or younger at the southwestern end of the orogen, and Late Ordovician or younger in the Carolinas and northern Georgia. Zircon ages reported from large thrust and dextral strike-slip faults bounding the Pine Mountain window indicate all of the faults there may be Alleghanian, except the younger sinistral Mesozoic faults, and requires that both metamorphism and penetrative deformation there also be Alleghanian. As in New England, the southern Appalachian Alleghanian metamorphic core is now known to be much more extensive. The older data require that the Taconian and perhaps the Acadian orogenies were significant events in the SCA, but these new data reconfirm the dominance of Alleghanian continent-continent collision processes here.« less

  5. The oldest fossil mushroom.

    PubMed

    Heads, Sam W; Miller, Andrew N; Crane, J Leland; Thomas, M Jared; Ruffatto, Danielle M; Methven, Andrew S; Raudabaugh, Daniel B; Wang, Yinan

    2017-01-01

    A new fossil mushroom is described and illustrated from the Lower Cretaceous Crato Formation of northeast Brazil. Gondwanagaricites magnificus gen. et sp. nov. is remarkable for its exceptional preservation as a mineralized replacement in laminated limestone, as all other fossil mushrooms are known from amber inclusions. Gondwanagaricites represents the oldest fossil mushroom to date and the first fossil mushroom from Gondwana.

  6. The oldest fossil mushroom

    PubMed Central

    Miller, Andrew N.; Crane, J. Leland; Thomas, M. Jared; Ruffatto, Danielle M.; Methven, Andrew S.; Raudabaugh, Daniel B.; Wang, Yinan

    2017-01-01

    A new fossil mushroom is described and illustrated from the Lower Cretaceous Crato Formation of northeast Brazil. Gondwanagaricites magnificus gen. et sp. nov. is remarkable for its exceptional preservation as a mineralized replacement in laminated limestone, as all other fossil mushrooms are known from amber inclusions. Gondwanagaricites represents the oldest fossil mushroom to date and the first fossil mushroom from Gondwana. PMID:28591180

  7. Widespread kelp-derived carbon in pelagic and benthic nearshore fishes suggested by stable isotope analysis

    NASA Astrophysics Data System (ADS)

    von Biela, Vanessa R.; Newsome, Seth D.; Bodkin, James L.; Kruse, Gordon H.; Zimmerman, Christian E.

    2016-11-01

    Kelp forests provide habitat for diverse and abundant fish assemblages, but the extent to which kelp provides a source of energy to fish and other predators is unclear. To examine the use of kelp-derived energy by fishes we estimated the contribution of kelp- and phytoplankton-derived carbon using carbon (δ13C) and nitrogen (δ15N) isotopes measured in muscle tissue. Benthic-foraging kelp greenling (Hexagrammos decagrammus) and pelagic-foraging black rockfish (Sebastes melanops) were collected at eight sites spanning ∼35 to 60°N from the California Current (upwelling) to Alaska Coastal Current (downwelling) in the northeast Pacific Ocean. Muscle δ13C values were expected to be higher for fish tissue primarily derived from kelp, a benthic macroalgae, and lower for tissue primarily derived from phytoplankton, pelagic microalgae. Muscle δ13C values were higher in benthic-feeding kelp greenling than in pelagic-feeding black rockfish at seven of eight sites, indicating more kelp-derived carbon in greenling as expected. Estimates of kelp carbon contributions ranged from 36 to 89% in kelp greenling and 32 to 65% in black rockfish using carbon isotope mixing models. Isotopic evidence suggests that these two nearshore fishes routinely derive energy from kelp and phytoplankton, across coastal upwelling and downwelling systems. Thus, the foraging mode of nearshore predators has a small influence on their ultimate energy source as energy produced by benthic macroalgae and pelagic microalgae were incorporated in fish tissue regardless of feeding mode and suggest strong and widespread benthic-pelagic coupling. Widespread kelp contributions to benthic- and pelagic-feeding fishes suggests that kelp energy provides a benefit to nearshore fishes and highlights the potential for kelp and fish production to be linked.

  8. Increased oceanic microplastic debris enhances oviposition in an endemic pelagic insect

    PubMed Central

    Goldstein, Miriam C.; Rosenberg, Marci; Cheng, Lanna

    2012-01-01

    Plastic pollution in the form of small particles (diameter less than 5 mm)—termed ‘microplastic’—has been observed in many parts of the world ocean. They are known to interact with biota on the individual level, e.g. through ingestion, but their population-level impacts are largely unknown. One potential mechanism for microplastic-induced alteration of pelagic ecosystems is through the introduction of hard-substrate habitat to ecosystems where it is naturally rare. Here, we show that microplastic concentrations in the North Pacific Subtropical Gyre (NPSG) have increased by two orders of magnitude in the past four decades, and that this increase has released the pelagic insect Halobates sericeus from substrate limitation for oviposition. High concentrations of microplastic in the NPSG resulted in a positive correlation between H. sericeus and microplastic, and an overall increase in H. sericeus egg densities. Predation on H. sericeus eggs and recent hatchlings may facilitate the transfer of energy between pelagic- and substrate-associated assemblages. The dynamics of hard-substrate-associated organisms may be important to understanding the ecological impacts of oceanic microplastic pollution. PMID:22573831

  9. Will My Fossil Float?

    ERIC Educational Resources Information Center

    Riesser, Sharon; Airey, Linda

    1993-01-01

    Explains how young students can be introduced to fossils. Suggests books to read and science activities including "Fossils to Eat" where students make fossils from peanut butter, honey, and powdered milk. (PR)

  10. Fossil Energy Program

    NASA Astrophysics Data System (ADS)

    McNeese, L. E.

    1981-01-01

    Increased utilization of coal and other fossil fuel alternatives as sources of clean energy is reported. The following topics are discussed: coal conversion development, chemical research and development, materials technology, component development and process evaluation studies, technical support to major liquefaction projects, process analysis and engineering evaluations, fossil energy environmental analysis, flue gas desulfurization, solid waste disposal, coal preparation waste utilization, plant control development, atmospheric fluidized bed coal combustor for cogeneration, TVA FBC demonstration plant program technical support, PFBC systems analysis, fossil fuel applications assessments, performance assurance system support for fossil energy projects, international energy technology assessment, and general equilibrium models of liquid and gaseous fuel supplies.

  11. 50 CFR 622.274 - Pelagic longline closed areas.

    Code of Federal Regulations, 2013 CFR

    2013-10-01

    ... ATMOSPHERIC ADMINISTRATION, DEPARTMENT OF COMMERCE FISHERIES OF THE CARIBBEAN, GULF OF MEXICO, AND SOUTH ATLANTIC Dolphin and Wahoo Fishery Off the Atlantic States § 622.274 Pelagic longline closed areas. (a) If... dolphin or wahoo— (1) In the Northeastern United States closed area from June 1 through June 30 each year...

  12. 50 CFR 622.274 - Pelagic longline closed areas.

    Code of Federal Regulations, 2014 CFR

    2014-10-01

    ... ATMOSPHERIC ADMINISTRATION, DEPARTMENT OF COMMERCE FISHERIES OF THE CARIBBEAN, GULF OF MEXICO, AND SOUTH ATLANTIC Dolphin and Wahoo Fishery Off the Atlantic States § 622.274 Pelagic longline closed areas. (a) If... dolphin or wahoo— (1) In the Northeastern United States closed area from June 1 through June 30 each year...

  13. Organic molecules as chemical fossils - The molecular fossil record

    NASA Technical Reports Server (NTRS)

    Eglinton, G.

    1983-01-01

    The study of biochemical clues to the early earth and the origin of life is discussed. The methods used in such investigation are described, including the extraction, fractionation, and analysis of geolipids and the analysis of kerogen. The occurrence of molecular fossils in the geological record is examined, discussing proposed precursor-product relationships and the molecular assessment of deep sea sediments, ancient sediments, and crude petroleums. Alterations in the molecular record due to diagenesis and catagenesis are considered, and the use of microbial lipids as molecular fossils is discussed. The results of searches for molecular fossils in Precambrian sediments are assessed.

  14. Spatial and temporal changes in the Barents Sea pelagic compartment during the recent warming

    NASA Astrophysics Data System (ADS)

    Eriksen, Elena; Skjoldal, Hein Rune; Gjøsæter, Harald; Primicerio, Raul

    2017-02-01

    The Barents Sea has experienced substantial warming over the last few decades with expansion of relatively warm Atlantic water and reduction in sea ice. Based on a review of relevant literature and additional analyses, we report changes in the pelagic compartment associated with this warming using data from autumn surveys (acoustic capelin, 0-group fish, and ecosystem surveys). We estimated biomass for 25 components of the pelagic community, including macroplankton, 0-group fish, and juvenile and adult pelagic fish, were examined for spatial and temporal variation over the period 1993-2013. The estimated total biomass of the investigated pelagic compartment, not including mesozooplankton, ranged between about 6 and 30 million tonnes wet weight with an average of 17 million tonnes over the 21-years period. Krill was the dominant biomass component (63%), whereas pelagic fish (capelin, polar cod and herring) made up 26% and 0-group fish 11% of the biomass on average. The spatial distribution of biomass showed a broad-scale pattern reflecting differences in distribution of the main pelagic fishes (capelin in the north, polar cod in the east, and herring in the south) and transport of krill and 0-group fish with the Atlantic water flowing into the southern Barents Sea. Dividing the Barents Sea into six regions, the highest average biomass values were found in the Southwestern and South-Central subareas (about 4 million tonnes in each), with krill as the main component. Biomass was also high in the North-Central subarea (about 3 million tonnes) where capelin was the major contributor. The total estimated biomass of the pelagic compartment remained relatively stable during each of two main periods (before and after 2004), but increased by a factor of two from around 11 million tonnes in the first to around 23 million tonnes in the last period. The pronounced increase reflected the warming between the relatively cold 1990s and the warmer 2000s and was driven mainly by an

  15. An early Oligocene fossil demonstrates treeshrews are slowly evolving "living fossils".

    PubMed

    Li, Qiang; Ni, Xijun

    2016-01-14

    Treeshrews are widely considered a "living model" of an ancestral primate, and have long been called "living fossils". Actual fossils of treeshrews, however, are extremely rare. We report a new fossil species of Ptilocercus treeshrew recovered from the early Oligocene (~34 Ma) of China that represents the oldest definitive fossil record of the crown group of treeshrews and nearly doubles the temporal length of their fossil record. The fossil species is strikingly similar to the living Ptilocercus lowii, a species generally recognized as the most plesiomorphic extant treeshrew. It demonstrates that Ptilocercus treeshrews have undergone little evolutionary change in their morphology since the early Oligocene. Morphological comparisons and phylogenetic analysis support the long-standing idea that Ptilocercus treeshrews are morphologically conservative and have probably retained many characters present in the common stock that gave rise to archontans, which include primates, flying lemurs, plesiadapiforms and treeshrews. This discovery provides an exceptional example of slow morphological evolution in a mammalian group over a period of 34 million years. The persistent and stable tropical environment in Southeast Asia through the Cenozoic likely played a critical role in the survival of such a morphologically conservative lineage.

  16. FOSSIL2 energy policy model documentation: FOSSIL2 documentation

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    None

    1980-10-01

    This report discusses the structure, derivations, assumptions, and mathematical formulation of the FOSSIL2 model. Each major facet of the model - supply/demand interactions, industry financing, and production - has been designed to parallel closely the actual cause/effect relationships determining the behavior of the United States energy system. The data base for the FOSSIL2 program is large, as is appropriate for a system dynamics simulation model. When possible, all data were obtained from sources well known to experts in the energy field. Cost and resource estimates are based on DOE data whenever possible. This report presents the FOSSIL2 model at severalmore » levels. Volumes II and III of this report list the equations that comprise the FOSSIL2 model, along with variable definitions and a cross-reference list of the model variables. Volume III lists the model equations and a one line definition for equations, in a short, readable format.« less

  17. Evaluating analytical approaches for estimating pelagic fish biomass using simulated fish communities

    USGS Publications Warehouse

    Yule, Daniel L.; Adams, Jean V.; Warner, David M.; Hrabik, Thomas R.; Kocovsky, Patrick M.; Weidel, Brian C.; Rudstam, Lars G.; Sullivan, Patrick J.

    2013-01-01

    Pelagic fish assessments often combine large amounts of acoustic-based fish density data and limited midwater trawl information to estimate species-specific biomass density. We compared the accuracy of five apportionment methods for estimating pelagic fish biomass density using simulated communities with known fish numbers that mimic Lakes Superior, Michigan, and Ontario, representing a range of fish community complexities. Across all apportionment methods, the error in the estimated biomass generally declined with increasing effort, but methods that accounted for community composition changes with water column depth performed best. Correlations between trawl catch and the true species composition were highest when more fish were caught, highlighting the benefits of targeted trawling in locations of high fish density. Pelagic fish surveys should incorporate geographic and water column depth stratification in the survey design, use apportionment methods that account for species-specific depth differences, target midwater trawling effort in areas of high fish density, and include at least 15 midwater trawls. With relatively basic biological information, simulations of fish communities and sampling programs can optimize effort allocation and reduce error in biomass estimates.

  18. 77 FR 15284 - Fisheries of the Caribbean, Gulf of Mexico, and South Atlantic; Coastal Migratory Pelagic...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2012-03-15

    ... Migratory Pelagic Resources of the Gulf of Mexico and South Atlantic; Closure AGENCY: National Marine...: Temporary rule; closure. SUMMARY: NMFS closes the commercial sector of the coastal migratory pelagic fishery... king mackerel resource. DATES: This rule is effective 12:01 a.m., local time, March 14, 2012, through...

  19. 75 FR 53874 - Fisheries of the Exclusive Economic Zone Off Alaska; Pelagic Shelf Rockfish for Vessels...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2010-09-02

    .... 0910131362-0087-02] RIN 0648-XY71 Fisheries of the Exclusive Economic Zone Off Alaska; Pelagic Shelf Rockfish for Vessels Participating in the Rockfish Entry Level Fishery in the Central Regulatory Area of the... for pelagic shelf rockfish for vessels participating in the rockfish entry level fishery in the...

  20. Fossil Fuels.

    ERIC Educational Resources Information Center

    Crank, Ron

    This instructional unit is one of 10 developed by students on various energy-related areas that deals specifically with fossil fuels. Some topics covered are historic facts, development of fuels, history of oil production, current and future trends of the oil industry, refining fossil fuels, and environmental problems. Material in each unit may…

  1. Ediacara Fossils

    ERIC Educational Resources Information Center

    Science Teacher, 2005

    2005-01-01

    Now, a research team from Virginia Tech and Nanjing Institute of Geology and Paleontology has discovered uniquely well-preserved fossil forms from 550-million-year-old rocks of the Ediacaran Period. The research appears in the Proceedings of the National Academy of Sciences. The discovery of these unusually preserved fossils reveals unprecedented…

  2. The Pelagics Habitat Analysis Module (PHAM): Decision Support Tools for Pelagic Fisheries

    NASA Astrophysics Data System (ADS)

    Armstrong, E. M.; Harrison, D. P.; Kiefer, D.; O'Brien, F.; Hinton, M.; Kohin, S.; Snyder, S.

    2009-12-01

    PHAM is a project funded by NASA to integrate satellite imagery and circulation models into the management of commercial and threatened pelagic species. Specifically, the project merges data from fishery surveys, and fisheries catch and effort data with satellite imagery and circulation models to define the habitat of each species. This new information on habitat will then be used to inform population distribution and models of population dynamics that are used for management. During the first year of the project, we created two prototype modules. One module, which was developed for the Inter-American Tropical Tuna Commission, is designed to help improve information available to manage the tuna fisheries of the eastern Pacific Ocean. The other module, which was developed for the Coastal Pelagics Division of the Southwest Fishery Science Center, assists management of by-catch of mako, blue, and thresher sharks along the Californian coast. Both modules were built with the EASy marine geographic information system, which provides a 4 dimensional (latitude, longitude, depth, and time) home for integration of the data. The projects currently provide tools for automated downloading and geo-referencing of satellite imagery of sea surface temperature, height, and chlorophyll concentrations; output from JPL’s ECCO2 global circulation model and its ROM California current model; and gridded data from fisheries and fishery surveys. It also provides statistical tools for defining species habitat from these and other types of environmental data. These tools include unbalanced ANOVA, EOF analysis of satellite imagery, and multivariate search routines for fitting fishery data to transforms of the environmental data. Output from the projects consists of dynamic maps of the distribution of the species that are driven by the time series of satellite imagery and output from the circulation models. It also includes relationships between environmental variables and recruitment. During

  3. The importance of benthic-pelagic coupling for marine ecosystem functioning in a changing world.

    PubMed

    Griffiths, Jennifer R; Kadin, Martina; Nascimento, Francisco J A; Tamelander, Tobias; Törnroos, Anna; Bonaglia, Stefano; Bonsdorff, Erik; Brüchert, Volker; Gårdmark, Anna; Järnström, Marie; Kotta, Jonne; Lindegren, Martin; Nordström, Marie C; Norkko, Alf; Olsson, Jens; Weigel, Benjamin; Žydelis, Ramunas; Blenckner, Thorsten; Niiranen, Susa; Winder, Monika

    2017-06-01

    Benthic-pelagic coupling is manifested as the exchange of energy, mass, or nutrients between benthic and pelagic habitats. It plays a prominent role in aquatic ecosystems, and it is crucial to functions from nutrient cycling to energy transfer in food webs. Coastal and estuarine ecosystem structure and function are strongly affected by anthropogenic pressures; however, there are large gaps in our understanding of the responses of inorganic nutrient and organic matter fluxes between benthic habitats and the water column. We illustrate the varied nature of physical and biological benthic-pelagic coupling processes and their potential sensitivity to three anthropogenic pressures - climate change, nutrient loading, and fishing - using the Baltic Sea as a case study and summarize current knowledge on the exchange of inorganic nutrients and organic material between habitats. Traditionally measured benthic-pelagic coupling processes (e.g., nutrient exchange and sedimentation of organic material) are to some extent quantifiable, but the magnitude and variability of biological processes are rarely assessed, preventing quantitative comparisons. Changing oxygen conditions will continue to have widespread effects on the processes that govern inorganic and organic matter exchange among habitats while climate change and nutrient load reductions may have large effects on organic matter sedimentation. Many biological processes (predation, bioturbation) are expected to be sensitive to anthropogenic drivers, but the outcomes for ecosystem function are largely unknown. We emphasize how improved empirical and experimental understanding of benthic-pelagic coupling processes and their variability are necessary to inform models that can quantify the feedbacks among processes and ecosystem responses to a changing world. © 2017 The Authors. Global Change Biology Published by John Wiley & Sons Ltd.

  4. Internal wave deposits in Jurassic Kermanshah pelagic carbonates and radiolarites (Kermanshah area, West Iran)

    NASA Astrophysics Data System (ADS)

    Abdi, Asad; Gharaie, Mohamad Hosein Mahmudy; Bádenas, Beatriz

    2014-12-01

    We report eventites generated by turbulence events triggered by breaking internal waves in Jurassic pelagic muds deposited in a graben area located between the Arabian and Bisotoun carbonate platforms, at the Kermanshah basin (West Iran). The 43 m-thick studied Pliensbachian-Aalenian succession at Kermanshah includes sponge spicule-radiolarian limestones and cherts with cm- to dm-thick intercalations of pyroclastic beds and coarse-grained deposits formed by neritic-derived grains and reworked pelagic material. Breaking of internal waves in localized areas reworked the available sediment on sea floor, including the erosion of cohesive pelagic muds and the resuspension of neritic-derived grains, which were resedimented from the Bisotoun platform most probably by storms or turbidity currents. The generated internal wave deposits include: flat- and round pebble limestone conglomerates, formed by deposition of pelagic clasts and neritic-derived grains near the breaker zone; laminated packstone-grainstones deposited by high-energy, upslope (swash) and downslope (backswash) flows; cm-thick packstone-grainstones with asymmetrical starved ripples and hummocy crossstratification, generated downdip by waning of backwash flows and internal wave oscillatory flows. These internal wave deposits predominate in the Pliensbachian-early Toarcian, and were related to internal waves developed along a thermocline linked to climate warming and excited by submarine volcanic eruptions, storms or tectonic shaking.

  5. A mathematical model of algae growth in a pelagic-benthic coupled shallow aquatic ecosystem.

    PubMed

    Zhang, Jimin; Shi, Junping; Chang, Xiaoyuan

    2018-04-01

    A coupled system of ordinary differential equations and partial differential equations is proposed to describe the interaction of pelagic algae, benthic algae and one essential nutrient in an oligotrophic shallow aquatic ecosystem with ample supply of light. The existence and uniqueness of non-negative steady states are completely determined for all possible parameter range, and these results characterize sharp threshold conditions for the regime shift from extinction to coexistence of pelagic and benthic algae. The influence of environmental parameters on algal biomass density is also considered, which is an important indicator of algal blooms. Our studies suggest that the nutrient recycling from loss of algal biomass may be an important factor in the algal blooms process; and the presence of benthic algae may limit the pelagic algal biomass density as they consume common resources even if the sediment nutrient level is high.

  6. [Effects of submarine topography and water depth on distribution of pelagic fish community in minnan-taiwan bank fishing ground].

    PubMed

    Fang, Shuimei; Yang, Shengyun; Zhang, Chengmao; Zhu, Jinfu

    2002-11-01

    According to the fishing record of the light-seine information vessel in Minnan-Taiwan bank ground during 1989 to 1999, the effects of submarine topography and water depth on distribution of pelagic fish community in Minnan-Taiwan bank fishing ground was studied. The results showed that the pelagic fish distributed concentratively, while the submarine topography and water depth varied widely, but in different fishing regions, the distribution of pelagic fishes was uneven. The distribution of fishing yield increased from north to south, and closed up from sides of the bank to south or north in the regions. Pelagic fish distributed mainly in mixed water in the southern Taiwan Strait, and in warm water in the Taiwan Strait. The central fishing grounds were at high salt regions. Close gathering regions of pelagic fish or central fishing ground would be varied with the seasonal variation of mixed water in the southern Taiwan Strait and warm water in the Taiwan Strait. Central fishing ground was not only related to submarine topography and water depth, but also related to wind direction, wind-power and various water systems. In the fishing ground, the gathering depth of pelagic fish was 30-60 m in spring and summer, and 40-80 m in autumn and winter.

  7. [Trophic niche partitioning of pelagic sharks in Central Eastern Pacific inferred from stable isotope analysis.

    PubMed

    Li, Yun Kai; Gao, Xiao di; Wang, Lin Yu; Fang, Lin

    2018-01-01

    As the apex predators of the open ocean ecosystems, pelagic sharks play important roles in stabilizing the marine food web through top-down control. Stable isotope analysis is a powerful tool to investigate the feeding ecology. The carbon and nitrogen isotope ratios can be used to trace food source and evaluate the trophic position of marine organisms. In this study, the isotope values of 130 pelagic sharks from 8 species in Central Eastern Pacific were analyzed and their trophic position and niche were calculated to compare the intra/inter-specific resource partitioning in the Central Eastern Pacific ecosystem. The results exhibited significant differences in both carbon and nitrogen isotope values among the shark species. The trophic levels ranged from 4.3 to 5.4 in the Central Eastern Pacific shark community. The trophic niche of blue sharks and shortfin mako sharks showed no overlap with the other shark species, exhibiting unique ecological roles in the open ocean food web. These data highlighted the diverse roles among pelagic sharks, supporting previous findings that this species is not trophically redundant and the trophic niche of pelagic sharks can not be simply replaced by those of other top predator species.

  8. FOSSIL2 energy policy model documentation: FOSSIL2 documentation

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    None

    1980-10-01

    This report discusses the structure, derivations, assumptions, and mathematical formulation of the FOSSIL2 model. Each major facet of the model - supply/demand interactions, industry financing, and production - has been designed to parallel closely the actual cause/effect relationships determining the behavior of the United States energy system. The data base for the FOSSIL2 program is large, as is appropriate for a system dynamics simulation model. When possible, all data were obtained from sources well known to experts in the energy field. Cost and resource estimates are based on DOE data whenever possible. This report presents the FOSSIL2 model at severalmore » levels. Volumes II and III of this report list the equations that comprise the FOSSIL2 model, along with variable definitions and a cross-reference list of the model variables. Volume II provides the model equations with each of their variables defined, while Volume III lists the equations, and a one line definition for equations, in a shorter, more readable format.« less

  9. 76 FR 37761 - Fisheries Off West Coast States; Coastal Pelagic Species Fisheries; Amendment 13 to the Coastal...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-06-28

    ...; Amendment 13 to the Coastal Pelagic Species Fishery Management Plan; Annual Catch Limits AGENCY: National... the Coastal Pelagic Species (CPS) Fishery Management Plan (FMP). This proposed rule will implement... in place to set and adjust fishery specifications and management measures and modifies this framework...

  10. Fossil Simulation in the Classroom

    ERIC Educational Resources Information Center

    Hoehn, Robert G.

    1977-01-01

    Describes classroom science demonstrations and experiments that simulate the process of fossil formation. Lists materials, procedures and suggestions for successful activities. Includes ten student activities (coral fossils, leaf fossils, leaf scars, carbonization, etc.). Describes a fossil game in which students work in pairs. (CS)

  11. Feeding ecology of pelagic fish larvae and juveniles in slope waters of the Gulf of Mexico.

    PubMed

    Wells, R J D; Rooker, J R

    2009-11-01

    Stable isotope ratios of carbon (delta13C) and nitrogen (delta15N) were used to investigate feeding patterns of larval and early juvenile pelagic fishes in slope waters of the Gulf of Mexico. Contribution of organic matter supplied to fishes and trophic position within this pelagic food web was estimated in 2007 and 2008 by comparing dietary signatures of the two main producers in this ecosystem: phytoplankton [based on particulate organic matter (POM)] and Sargassum spp. Stable isotope ratios of POM and pelagic Sargassum spp. were significantly different from one another with delta13C values of POM depleted by 3-6 per thousand and delta15N values enriched by 2 relative to Sargassum spp. Stable isotope ratios were significantly different among the five pelagic fishes examined: blue marlin Makaira nigricans, dolphinfish Coryphaena hippurus, pompano dolphinfish Coryphaena equiselis, sailfish Istiophorus platypterus and swordfish Xiphias gladius. Mean delta13C values ranged almost 2 among fishes and were most depleted in I. platypterus. In addition, mean delta15N values ranged 4-5 with highest mean values found for both C. hippurus and C. equiselis and the lowest mean value for M. nigricans during both years. Increasing delta13C or delta15N with standard length suggested that shifts in trophic position and diet occurred during early life for several species examined. Results of a two-source mixing model suggest approximately an equal contribution of organic matter by both sources (POM=55%; pelagic Sargassum spp.=45%) to the early life stages of pelagic fishes examined. Contribution of organic matter, however, varied among species, and sensitivity analyses indicated that organic source estimates changed from 2 to 13% for a delta(13)C fractionation change of +/-0.25 per thousand or a delta15N fractionation change of +/-1.0 per thousand relative to original fractionation values.

  12. Seasonal changes of the pelagic fish assemblage in a temperate estuary

    NASA Astrophysics Data System (ADS)

    Hagan, S. M.; Able, K. W.

    2003-01-01

    The pelagic (i.e. fishes that spend most of their time swimming and feeding in the water column) fish assemblage was examined within a temperate estuary to determine the seasonal patterns of abundance, biomass, richness, and species composition. These measures were related to abiotic measures including temperature, salinity, water transparency, and depth. Pelagic fish (n<350 000) were sampled weekly from August 1995 through December 1996 in Great Bay in New Jersey, USA, with a pop net (5 m diameter, 5 m length, 3 mm cod end). The most frequently occurring and abundant species included Clupea harengus (frequency of occurrence, 27.4%; abundance, 80.5% of total catch), Menidia menidia (frequency of occurrence, 83.5%; abundance, 10.6% of total catch), Anchoa mitchilli (frequency of occurrence, 66.5%; abundance, 6.0% of total catch), and Anchoa hepsetus (frequency of occurrence, 23.1%; abundance, 1.9% of total catch). There were strong seasonal trends in species richness, total abundance, and total biomass with peaks in spring, summer, and autumn, and very low values for these parameters in winter. In addition, there were five seasonal groupings within this pelagic fish assemblage that were primarily, but not exclusively, represented by young-of-the-year (age 0+). Winter (December-March) was represented by adult Gasterosteus aculeatus and age 1+ by M. menidia. Spring (April-early June) was overwhelmingly dominated by age 0+ C. harengus. Early summer (late June-early July) was relatively species rich and characterized by age 0+ of Brevoortia tyrannus, Pomatomus saltatrix, Alosa pseudoharengus, and age 1+ of A. mitchilli. Late summer (late July-August) remained relatively species rich and included mostly age 0+ of A. mitchilli, A. hepsetus, and M. menidia. Autumn (September-November) continued to have high catches of age 0+ of M. menidia and A. mitchilli. As expected for temperate estuaries, the best of the measured abiotic predictors of these groupings included

  13. 50 CFR 665.815 - Pelagic longline seabird mitigation measures.

    Code of Federal Regulations, 2010 CFR

    2010-10-01

    ... 50 Wildlife and Fisheries 9 2010-10-01 2010-10-01 false Pelagic longline seabird mitigation measures. 665.815 Section 665.815 Wildlife and Fisheries FISHERY CONSERVATION AND MANAGEMENT, NATIONAL... and livers in accordance with paragraph (a)(2)(i) of this section; (v) When using basket-style...

  14. Evolution of the Sauropterygian Labyrinth with Increasingly Pelagic Lifestyles.

    PubMed

    Neenan, James M; Reich, Tobias; Evers, Serjoscha W; Druckenmiller, Patrick S; Voeten, Dennis F A E; Choiniere, Jonah N; Barrett, Paul M; Pierce, Stephanie E; Benson, Roger B J

    2017-12-18

    Sauropterygia, a successful clade of marine reptiles abundant in aquatic ecosystems of the Mesozoic, inhabited nearshore to pelagic habitats over >180 million years of evolutionary history [1]. Aquatic vertebrates experience strong buoyancy forces that allow movement in a three-dimensional environment, resulting in structural convergences such as flippers and fish-like bauplans [2, 3], as well as convergences in the sensory systems. We used computed tomographic scans of 19 sauropterygian species to determine how the transition to pelagic lifestyles influenced the evolution of the endosseous labyrinth, which houses the vestibular sensory organ of balance and orientation [4]. Semicircular canal geometries underwent distinct changes during the transition from nearshore Triassic sauropterygians to the later, pelagic plesiosaurs. Triassic sauropterygians have dorsoventrally compact, anteroposteriorly elongate labyrinths, resembling those of crocodylians. In contrast, plesiosaurs have compact, bulbous labyrinths, sharing some features with those of sea turtles. Differences in relative labyrinth size among sauropterygians correspond to locomotory differences: bottom-walking [5, 6] placodonts have proportionally larger labyrinths than actively swimming taxa (i.e., all other sauropterygians). Furthermore, independent evolutionary origins of short-necked, large-headed "pliosauromorph" body proportions among plesiosaurs coincide with reductions of labyrinth size, paralleling the evolutionary history of cetaceans [7]. Sauropterygian labyrinth evolution is therefore correlated closely with both locomotory style and body proportions, and these changes are consistent with isolated observations made previously in other marine tetrapods. Our study presents the first virtual reconstructions of plesiosaur endosseous labyrinths and the first large-scale, quantitative study detailing the effects of increasingly aquatic lifestyles on labyrinth morphology among marine reptiles. Copyright

  15. Structure-activity relationships of polyphenols to prevent lipid oxidation in pelagic fish muscle.

    PubMed

    Pazos, Manuel; Iglesias, Jacobo; Maestre, Rodrigo; Medina, Isabel

    2010-10-27

    The influence of polymerization (number of monomers) and galloylation (content of esterified gallates) of oligomeric catechins (proanthocyanidins) on their effectiveness to prevent lipid oxidation in pelagic fish muscle was evaluated. Non-galloylated oligomers of catechin with diverse mean polymerization (1.9-3.4 monomeric units) were extracted from pine (Pinus pinaster) bark. Homologous fractions with galloylation ranging from 0.25 to <1 gallate group per molecule were obtained from grape (Vitis vinifera) and witch hazel (Hamamelis virginiana). The results showed the convenience of proanthocyanidins with medium size (2-3 monomeric units) and low galloylation degree (0.15-0.25 gallate group/molecule) to inhibit lipid oxidation in pelagic fish muscle. These optimal structural characteristics of proanthocyanidins were similar to those lately reported in fish oil-in-water emulsions using phosphatidylcholine as emulsifier. This finding suggests that the antioxidant behavior of polyphenols in muscle-based foods can be mimicked in emulsions prepared with phospholipids as emulsifier agents. The present data give relevant information to achieve an optimum use of polyphenols in pelagic fish muscle.

  16. Plastic ingestion by pelagic and demersal fish from the North Sea and Baltic Sea.

    PubMed

    Rummel, Christoph D; Löder, Martin G J; Fricke, Nicolai F; Lang, Thomas; Griebeler, Eva-Maria; Janke, Michael; Gerdts, Gunnar

    2016-01-15

    Plastic ingestion by marine biota has been reported for a variety of different taxa. In this study, we investigated 290 gastrointestinal tracts of demersal (cod, dab and flounder) and pelagic fish species (herring and mackerel) from the North and Baltic Sea for the occurrence of plastic ingestion. In 5.5% of all investigated fishes, plastic particles were detected, with 74% of all particles being in the microplastic (<5mm) size range. The polymer types of all found particles were analysed by means of Fourier transform infrared (FT-IR) spectroscopy. Almost 40% of the particles consisted of polyethylene (PE). In 3.4% of the demersal and 10.7% of the pelagic individuals, plastic ingestion was recorded, showing a significantly higher ingestion frequency in the pelagic feeders. The condition factor K was calculated to test differences in the fitness status between individuals with and without ingested plastic, but no direct effect was detected. Copyright © 2015 Elsevier Ltd. All rights reserved.

  17. Ecological diversification associated with the benthic-to-pelagic transition by North American minnows.

    PubMed

    Burress, E D; Holcomb, J M; Tan, M; Armbruster, J W

    2017-03-01

    Ecological opportunity is often regarded as a key factor that explains why diversity is unevenly distributed across life. Colonization of novel environments or adaptive zones may promote diversification. North American minnows exhibit an ancestral benthic-to-pelagic habitat shift that coincided with a burst in diversification. Here, we evaluate the phenotypic and ecological implications of this habitat shift by assessing craniofacial and dietary traits among 34 species and testing for morphology-diet covariation, convergence and adaptive optima. There were several instances of morphology-diet covariation such as correlations between mouth angle and the consumption of terrestrial insects and between relative gut length and the consumption of algae. After accounting for size and phylogenetic nonindependence, benthic species had longer heads, longer snouts, eyes positioned higher on their head, smaller mouth angles and longer digestive tracts than pelagic minnows. Benthic minnows also consumed more algae but less terrestrial insects, by volume, than pelagic minnows. Lastly, there were three distinct evolutionary regimes and more convergence in morphology and dietary characteristics than expected under a Brownian motion model of evolution. These findings indicate that colonization of the pelagic zone by minnows involved myriad phenotypic and dietary changes associated with exploitation of terrestrial subsidies. Thus, minnows exhibit phenotype-dietary covariation, an expansion of ecological roles and a burst in diversification rates in response to the ecological opportunity afforded by the colonization of a novel habitat. © 2016 European Society For Evolutionary Biology. Journal of Evolutionary Biology © 2016 European Society For Evolutionary Biology.

  18. Deepwater Horizon crude oil impacts the developing hearts of large predatory pelagic fish

    PubMed Central

    Incardona, John P.; Gardner, Luke D.; Linbo, Tiffany L.; Brown, Tanya L.; Esbaugh, Andrew J.; Mager, Edward M.; Stieglitz, John D.; French, Barbara L.; Labenia, Jana S.; Laetz, Cathy A.; Tagal, Mark; Sloan, Catherine A.; Elizur, Abigail; Benetti, Daniel D.; Grosell, Martin; Block, Barbara A.; Scholz, Nathaniel L.

    2014-01-01

    The Deepwater Horizon disaster released more than 636 million L of crude oil into the northern Gulf of Mexico. The spill oiled upper surface water spawning habitats for many commercially and ecologically important pelagic fish species. Consequently, the developing spawn (embryos and larvae) of tunas, swordfish, and other large predators were potentially exposed to crude oil-derived polycyclic aromatic hydrocarbons (PAHs). Fish embryos are generally very sensitive to PAH-induced cardiotoxicity, and adverse changes in heart physiology and morphology can cause both acute and delayed mortality. Cardiac function is particularly important for fast-swimming pelagic predators with high aerobic demand. Offspring for these species develop rapidly at relatively high temperatures, and their vulnerability to crude oil toxicity is unknown. We assessed the impacts of field-collected Deepwater Horizon (MC252) oil samples on embryos of three pelagic fish: bluefin tuna, yellowfin tuna, and an amberjack. We show that environmentally realistic exposures (1–15 µg/L total PAH) cause specific dose-dependent defects in cardiac function in all three species, with circulatory disruption culminating in pericardial edema and other secondary malformations. Each species displayed an irregular atrial arrhythmia following oil exposure, indicating a highly conserved response to oil toxicity. A considerable portion of Gulf water samples collected during the spill had PAH concentrations exceeding toxicity thresholds observed here, indicating the potential for losses of pelagic fish larvae. Vulnerability assessments in other ocean habitats, including the Arctic, should focus on the developing heart of resident fish species as an exceptionally sensitive and consistent indicator of crude oil impacts. PMID:24706825

  19. Biogeochemical significance of pelagic ecosystem function: an end-Cretaceous case study

    PubMed Central

    Penman, Donald E.; Rae, James W. B.

    2016-01-01

    Pelagic ecosystem function is integral to global biogeochemical cycling, and plays a major role in modulating atmospheric CO2 concentrations (pCO2). Uncertainty as to the effects of human activities on marine ecosystem function hinders projection of future atmospheric pCO2. To this end, events in the geological past can provide informative case studies in the response of ecosystem function to environmental and ecological changes. Around the Cretaceous–Palaeogene (K–Pg) boundary, two such events occurred: Deccan large igneous province (LIP) eruptions and massive bolide impact at the Yucatan Peninsula. Both perturbed the environment, but only the impact coincided with marine mass extinction. As such, we use these events to directly contrast the response of marine biogeochemical cycling to environmental perturbation with and without changes in global species richness. We measure this biogeochemical response using records of deep-sea carbonate preservation. We find that Late Cretaceous Deccan volcanism prompted transient deep-sea carbonate dissolution of a larger magnitude and timescale than predicted by geochemical models. Even so, the effect of volcanism on carbonate preservation was slight compared with bolide impact. Empirical records and geochemical models support a pronounced increase in carbonate saturation state for more than 500 000 years following the mass extinction of pelagic carbonate producers at the K–Pg boundary. These examples highlight the importance of pelagic ecosystems in moderating climate and ocean chemistry. PMID:27114586

  20. Robustness of fossil fish teeth for seawater neodymium isotope reconstructions under variable redox conditions in an ancient shallow marine setting

    NASA Astrophysics Data System (ADS)

    Huck, Claire E.; van de Flierdt, Tina; Jiménez-Espejo, Francisco J.; Bohaty, Steven M.; Röhl, Ursula; Hammond, Samantha J.

    2016-03-01

    Fossil fish teeth from pelagic open ocean settings are considered a robust archive for preserving the neodymium (Nd) isotopic composition of ancient seawater. However, using fossil fish teeth as an archive to reconstruct seawater Nd isotopic compositions in different sedimentary redox environments and in terrigenous-dominated, shallow marine settings is less proven. To address these uncertainties, fish tooth and sediment samples from a middle Eocene section deposited proximal to the East Antarctic margin at Integrated Ocean Drilling Program Site U1356 were analyzed for major and trace element geochemistry, and Nd isotopes. Major and trace element analyses of the sediments reveal changing redox conditions throughout deposition in a shallow marine environment. However, variations in the Nd isotopic composition and rare earth element (REE) patterns of the associated fish teeth do not correspond to redox changes in the sediments. REE patterns in fish teeth at Site U1356 carry a typical mid-REE-enriched signature. However, a consistently positive Ce anomaly marks a deviation from a pure authigenic origin of REEs to the fish tooth. Neodymium isotopic compositions of cleaned and uncleaned fish teeth fall between modern seawater and local sediments and hence could be authigenic in nature, but could also be influenced by sedimentary fluxes. We conclude that the fossil fish tooth Nd isotope proxy is not sensitive to moderate changes in pore water oxygenation. However, combined studies on sediments, pore waters, fish teeth, and seawater are needed to fully understand processes driving the reconstructed signature from shallow marine sections in proximity to continental sources.

  1. Status of pelagic prey fishes in Lake Michigan, 2013

    USGS Publications Warehouse

    Warner, David M.; Farha, Steven A.; O'Brien, Timothy P.; Ogilvie, Lynn; Claramunt, Randall M.; Hanson, Dale

    2014-01-01

    Acoustic surveys were conducted in late summer/early fall during the years 1992-1996 and 2001-2013 to estimate pelagic prey fish biomass in Lake Michigan. Midwater trawling during the surveys as well as target strength provided a measure of species and size composition of the fish community for use in scaling acoustic data and providing species-specific abundance estimates. The 2013 survey consisted of 27 acoustic transects (546 km total) and 31 midwater trawl tows. Mean prey fish biomass was 6.1 kg/ha (relative standard error, RSE = 11%) or 29.6 kilotonnes (kt = 1,000 metric tons), which was similar to the estimate in 2012 (31.1 kt) and 23.5% of the long-term (18 years) mean. The numeric density of the 2013 alewife year class was 6% of the time series average and this year-class contributed 4% of total alewife biomass (5.2 kg/ha, RSE = 12%). Alewife ≥age-1 comprised 96% of alewife biomass. In 2013, alewife comprised 86% of total prey fish biomass, while rainbow smelt and bloater were 4 and 10% of total biomass, respectively. Rainbow smelt biomass in 2013 (0.24 kg/ha, RSE = 17%) was essentially identical to the rainbow smelt biomass in 2012 and was 6% of the long term mean. Bloater biomass in 2013 was 0.6 kg/ha, only half the 2012 biomass, and 6% of the long term mean. Mean density of small bloater in 2013 (29 fish/ha, RSE = 29%) was lower than peak values observed in 2007-2009 and was 23% of the time series mean. In 2013, pelagic prey fish biomass in Lake Michigan was similar to Lake Huron, but pelagic community composition differs in the two lakes, with Lake Huron dominated by bloater.

  2. Metagenomic Analysis of Viral Communities in (Hado)Pelagic Sediments

    PubMed Central

    Yoshida, Mitsuhiro; Takaki, Yoshihiro; Eitoku, Masamitsu; Nunoura, Takuro; Takai, Ken

    2013-01-01

    In this study, we analyzed viral metagenomes (viromes) in the sedimentary habitats of three geographically and geologically distinct (hado)pelagic environments in the northwest Pacific; the Izu-Ogasawara Trench (water depth = 9,760 m) (OG), the Challenger Deep in the Mariana Trench (10,325 m) (MA), and the forearc basin off the Shimokita Peninsula (1,181 m) (SH). Virus abundance ranged from 106 to 1011 viruses/cm3 of sediments (down to 30 cm below the seafloor [cmbsf]). We recovered viral DNA assemblages (viromes) from the (hado)pelagic sediment samples and obtained a total of 37,458, 39,882, and 70,882 sequence reads by 454 GS FLX Titanium pyrosequencing from the virome libraries of the OG, MA, and SH (hado)pelagic sediments, respectively. Only 24−30% of the sequence reads from each virome library exhibited significant similarities to the sequences deposited in the public nr protein database (E-value <10−3 in BLAST). Among the sequences identified as potential viral genes based on the BLAST search, 95−99% of the sequence reads in each library were related to genes from single-stranded DNA (ssDNA) viral families, including Microviridae, Circoviridae, and Geminiviridae. A relatively high abundance of sequences related to the genetic markers (major capsid protein [VP1] and replication protein [Rep]) of two ssDNA viral groups were also detected in these libraries, thereby revealing a high genotypic diversity of their viruses (833 genotypes for VP1 and 2,551 genotypes for Rep). A majority of the viral genes predicted from each library were classified into three ssDNA viral protein categories: Rep, VP1, and minor capsid protein. The deep-sea sedimentary viromes were distinct from the viromes obtained from the oceanic and fresh waters and marine eukaryotes, and thus, deep-sea sediments harbor novel viromes, including previously unidentified ssDNA viruses. PMID:23468952

  3. Metagenomic analysis of viral communities in (hado)pelagic sediments.

    PubMed

    Yoshida, Mitsuhiro; Takaki, Yoshihiro; Eitoku, Masamitsu; Nunoura, Takuro; Takai, Ken

    2013-01-01

    In this study, we analyzed viral metagenomes (viromes) in the sedimentary habitats of three geographically and geologically distinct (hado)pelagic environments in the northwest Pacific; the Izu-Ogasawara Trench (water depth = 9,760 m) (OG), the Challenger Deep in the Mariana Trench (10,325 m) (MA), and the forearc basin off the Shimokita Peninsula (1,181 m) (SH). Virus abundance ranged from 10(6) to 10(11) viruses/cm(3) of sediments (down to 30 cm below the seafloor [cmbsf]). We recovered viral DNA assemblages (viromes) from the (hado)pelagic sediment samples and obtained a total of 37,458, 39,882, and 70,882 sequence reads by 454 GS FLX Titanium pyrosequencing from the virome libraries of the OG, MA, and SH (hado)pelagic sediments, respectively. Only 24-30% of the sequence reads from each virome library exhibited significant similarities to the sequences deposited in the public nr protein database (E-value <10(-3) in BLAST). Among the sequences identified as potential viral genes based on the BLAST search, 95-99% of the sequence reads in each library were related to genes from single-stranded DNA (ssDNA) viral families, including Microviridae, Circoviridae, and Geminiviridae. A relatively high abundance of sequences related to the genetic markers (major capsid protein [VP1] and replication protein [Rep]) of two ssDNA viral groups were also detected in these libraries, thereby revealing a high genotypic diversity of their viruses (833 genotypes for VP1 and 2,551 genotypes for Rep). A majority of the viral genes predicted from each library were classified into three ssDNA viral protein categories: Rep, VP1, and minor capsid protein. The deep-sea sedimentary viromes were distinct from the viromes obtained from the oceanic and fresh waters and marine eukaryotes, and thus, deep-sea sediments harbor novel viromes, including previously unidentified ssDNA viruses.

  4. Reorganization of a marine trophic network along an inshore-offshore gradient due to stronger pelagic-benthic coupling in coastal areas

    NASA Astrophysics Data System (ADS)

    Kopp, Dorothée; Lefebvre, Sébastien; Cachera, Marie; Villanueva, Maria Ching; Ernande, Bruno

    2015-01-01

    Recent theoretical considerations have highlighted the importance of the pelagic-benthic coupling in marine food webs. In continental shelf seas, it was hypothesized that the trophic network structure may change along an inshore-offshore gradient due to weakening of the pelagic-benthic coupling from coastal to offshore areas. We tested this assumption empirically using the eastern English Channel (EEC) as a case study. We sampled organisms from particulate organic matter to predatory fishes and used baseline-corrected carbon and nitrogen stable isotope ratios (δ13C and δ15N) to determine their trophic position. First, hierarchical clustering on δ13C and δ15N coupled to bootstrapping and estimates of the relative contribution of pelagic and benthic carbon sources to consumers' diet showed that, at mesoscale, the EEC food web forms a continuum of four trophic levels with trophic groups spread across a pelagic and a benthic trophic pathway. Second, based on the same methods, a discrete approach examined changes in the local food web structure across three depth strata in order to investigate the inshore-offshore gradient. It showed stronger pelagic-benthic coupling in shallow coastal areas mostly due to a reorganization of the upper consumers relative to the two trophic pathways, benthic carbon sources being available to pelagic consumers and, reciprocally, pelagic sources becoming accessible to benthic species. Third a continuous approach examined changes in the mean and variance of upper consumers' δ13C and δ15N with depth. It detected a significant decrease in δ13C variance and a significant increase in δ15N variance as depth increases. A theoretical two-source mixing model showed that an inshore-offshore decrease in the pelagic-benthic coupling was a sufficient condition to produce the δ13C variance pattern, thus supporting the conclusions of the discrete approach. These results suggest that environmental gradients such as the inshore-offshore one should

  5. Impact of El Niño events on pelagic fisheries in Peruvian waters

    NASA Astrophysics Data System (ADS)

    Ñiquen, Miguel; Bouchon, Marilú

    2004-03-01

    Using data from stock assesment surveys on pelagic resources during El Niño events of 1972/73, 1982/83, 1997/98, we analyze biological changes on pelagic ecosystems and pelagic fisheries during different stages of development of El Niño phenomenon: emergence, full, final and post-Niño. Results indicate changes in spatial distribution of resources, their concentration and size structure. In anchovy (Engraulis ringens) a decrease in biomass was observed, which was estimated at 1.2 million tons in September 1998, the lowest throughout the 1990s. This resource showed an asymmetric distribution towards the south of Peru. Other pelagic resources increased their biomass during or after Niño events, primarily sardine (Sardinops sagax), jack mackerel (Trachurus murphyi), pacific mackerel (Scomber japonicus), and longnose anchovy (Anchoa nasus). At the end of the El Niño phenomenon we found less productivity but more diversity in the pelagic ecosystem. During the 1997/98 El Niño, the diversity index (Manual de Ecologia, 1a Edition, Editorial Trillas, Mexico, 267pp) increased from 0.87 to 1.23-1.70. In both the emergence stage and fully developed stages of El Niño we found large numbers of sardine and longnose anchovy present simultaneously. Size structure of sardine, jack mackerel, and pacific mackerel showed an increase in juveniles. Anchovy during El Niño showed a single modal group composed of adults, but the post-Niño phase indicated an increase in juveniles with an average length of 6-7 cm. In El Niño conditions spawning among anchovy was low, but among sardine and pacific mackerel it was high. We observed, for the first time during full spawning, juvenile sardines with a total length of 18-20 cm. The anchovy spawning season during the post-Niño phase was considerably lengthened, from April to December 1998. Drastic change occurred in fisheries when monospecific fisheries, based on anchovy before El Niño, became multispecific fisheries based on sardine, jack

  6. Arthropod visual predators in the early pelagic ecosystem: evidence from the Burgess Shale and Chengjiang biotas.

    PubMed

    Vannier, J; García-Bellido, D C; Hu, S-X; Chen, A-L

    2009-07-22

    Exceptional fossil specimens with preserved soft parts from the Maotianshan Shale (ca 520 Myr ago) and the Burgess Shale (505 Myr ago) biotas indicate that the worldwide distributed bivalved arthropod Isoxys was probably a non-benthic visual predator. New lines of evidence come from the functional morphology of its powerful prehensile frontal appendages that, combined with large spherical eyes, are thought to have played a key role in the recognition and capture of swimming or epibenthic prey. The swimming and steering of this arthropod was achieved by the beating of multiple setose exopods and a flap-like telson. The appendage morphology of Isoxys indicates possible phylogenetical relationships with the megacheirans, a widespread group of assumed predator arthropods characterized by a pre-oral 'great appendage'. Evidence from functional morphology and taphonomy suggests that Isoxys was able to migrate through the water column and was possibly exploiting hyperbenthic niches for food. Although certainly not unique, the case of Isoxys supports the idea that off-bottom animal interactions such as predation, associated with complex feeding strategies and behaviours (e.g. vertical migration and hunting) were established by the Early Cambrian. It also suggests that a prototype of a pelagic food chain had already started to build-up at least in the lower levels of the water column.

  7. Understanding the structure and functioning of polar pelagic ecosystems to predict the impacts of change

    PubMed Central

    Drinkwater, K. F.; Grant, S. M.; Heymans, J. J.; Hofmann, E. E.; Hunt, G. L.; Johnston, N. M.

    2016-01-01

    The determinants of the structure, functioning and resilience of pelagic ecosystems across most of the polar regions are not well known. Improved understanding is essential for assessing the value of biodiversity and predicting the effects of change (including in biodiversity) on these ecosystems and the services they maintain. Here we focus on the trophic interactions that underpin ecosystem structure, developing comparative analyses of how polar pelagic food webs vary in relation to the environment. We highlight that there is not a singular, generic Arctic or Antarctic pelagic food web, and, although there are characteristic pathways of energy flow dominated by a small number of species, alternative routes are important for maintaining energy transfer and resilience. These more complex routes cannot, however, provide the same rate of energy flow to highest trophic-level species. Food-web structure may be similar in different regions, but the individual species that dominate mid-trophic levels vary across polar regions. The characteristics (traits) of these species are also different and these differences influence a range of food-web processes. Low functional redundancy at key trophic levels makes these ecosystems particularly sensitive to change. To develop models for projecting responses of polar ecosystems to future environmental change, we propose a conceptual framework that links the life histories of pelagic species and the structure of polar food webs. PMID:27928038

  8. Understanding the structure and functioning of polar pelagic ecosystems to predict the impacts of change.

    PubMed

    Murphy, E J; Cavanagh, R D; Drinkwater, K F; Grant, S M; Heymans, J J; Hofmann, E E; Hunt, G L; Johnston, N M

    2016-12-14

    The determinants of the structure, functioning and resilience of pelagic ecosystems across most of the polar regions are not well known. Improved understanding is essential for assessing the value of biodiversity and predicting the effects of change (including in biodiversity) on these ecosystems and the services they maintain. Here we focus on the trophic interactions that underpin ecosystem structure, developing comparative analyses of how polar pelagic food webs vary in relation to the environment. We highlight that there is not a singular, generic Arctic or Antarctic pelagic food web, and, although there are characteristic pathways of energy flow dominated by a small number of species, alternative routes are important for maintaining energy transfer and resilience. These more complex routes cannot, however, provide the same rate of energy flow to highest trophic-level species. Food-web structure may be similar in different regions, but the individual species that dominate mid-trophic levels vary across polar regions. The characteristics (traits) of these species are also different and these differences influence a range of food-web processes. Low functional redundancy at key trophic levels makes these ecosystems particularly sensitive to change. To develop models for projecting responses of polar ecosystems to future environmental change, we propose a conceptual framework that links the life histories of pelagic species and the structure of polar food webs. © 2016 The Authors.

  9. Fossilization of melanosomes via sulfurization.

    PubMed

    McNamara, Maria E; van Dongen, Bart E; Lockyer, Nick P; Bull, Ian D; Orr, Patrick J

    2016-05-01

    Fossil melanin granules (melanosomes) are an important resource for inferring the evolutionary history of colour and its functions in animals. The taphonomy of melanin and melanosomes, however, is incompletely understood. In particular, the chemical processes responsible for melanosome preservation have not been investigated. As a result, the origins of sulfur-bearing compounds in fossil melanosomes are difficult to resolve. This has implications for interpretations of original colour in fossils based on potential sulfur-rich phaeomelanosomes. Here we use pyrolysis gas chromatography mass spectrometry (Py-GCMS), fourier transform infrared spectroscopy (FTIR) and time of flight secondary ion mass spectrometry (ToF-SIMS) to assess the mode of preservation of fossil microstructures, confirmed as melanosomes based on the presence of melanin, preserved in frogs from the Late Miocene Libros biota (NE Spain). Our results reveal a high abundance of organosulfur compounds and non-sulfurized fatty acid methyl esters in both the fossil tissues and host sediment; chemical signatures in the fossil tissues are inconsistent with preservation of phaeomelanin. Our results reflect preservation via the diagenetic incorporation of sulfur, i.e. sulfurization (natural vulcanization), and other polymerization processes. Organosulfur compounds and/or elevated concentrations of sulfur have been reported from melanosomes preserved in various invertebrate and vertebrate fossils and depositional settings, suggesting that preservation through sulfurization is likely to be widespread. Future studies of sulfur-rich fossil melanosomes require that the geochemistry of the host sediment is tested for evidence of sulfurization in order to constrain interpretations of potential phaeomelanosomes and thus of original integumentary colour in fossils.

  10. Restoring Fossil Creek

    ERIC Educational Resources Information Center

    Flaccus, Kathleen; Vlieg, Julie; Marks, Jane C.; LeRoy, Carri J.

    2004-01-01

    Fossil Creek had been dammed for the past 90 years, and plans were underway to restore the stream. The creek runs through Central Arizona and flows from the high plateaus to the desert, cutting through the same formations that form the Grand Canyon. This article discusses the Fossil Creek monitoring project. In this project, students and teachers…

  11. Recruitment phenology and pelagic larval duration in Caribbean amphidromous fishes

    USGS Publications Warehouse

    Engman, Augustin C.; Kwak, Thomas J.; Fischer, Jesse R.

    2017-01-01

    Amphidromous fishes are major components of oceanic tropical island stream ecosystems, such as those of the Caribbean island, Puerto Rico. Fishes with this life history face threats related to the requirement for connectivity between freshwater and marine environments during early life stages. Pelagic larval duration and recruitment phenology are 2 early life-history processes that are crucial for the biology, ecology, conservation, and management of amphidromous fishes. However, these processes are understudied in the Caribbean in general and have never been quantified in Puerto Rico. We quantified recruit abundance, recruitment phenology, and pelagic larval duration of several Caribbean amphidromous fish species in multiple rivers in Puerto Rico and explored the effects of environmental variables on recruit abundances. Two fish taxa—sirajo goby (Sicydium spp.) and River Goby (Awaous banana)—were exceptionally abundant as postlarvae and recruited to Caribbean rivers in pulsed migration episodes that were periodic at annual and lunar scales. Sirajo goby and River Goby recruit abundances varied among rivers, were greater at sunrise than at sunset, and were positively related to river discharge. The pelagic larval duration of 4 fish taxa ranged from a minimum of 28 d to a maximum of 103 d with means between 43 ± 7 d (SD) and 65 ± 11 d. We identified the last-quarter moon phase during the months of June through January as periods of maximum amphidromous fish recruitment to freshwater streams. The results and conclusions of our study can be applied to identify critical times to maintain river–ocean connectivity and stream flow for the benefit of the amphidromous fish population dynamics, stream ecology, and natural resources of the Caribbean.

  12. Children's Ideas about Fossils and Foundational Concepts Related to Fossils

    ERIC Educational Resources Information Center

    Borgerding, Lisa A.; Raven, Sara

    2018-01-01

    Many standards documents and learning progressions recommend evolution learning in elementary grades. Given young children's interest in dinosaurs and other fossils, fossil investigations can provide a rich entry into evolutionary biology for young learners. Educational psychology literature has addressed children's reasoning about foundational…

  13. Cross-habitat effects shape the ecosystem consequences of co-invasion by a pelagic and a benthic consumer.

    PubMed

    Fryxell, David C; Diluzio, Amber R; Friedman, Maya A; Menge, Nicklaus A; Palkovacs, Eric P

    2016-10-01

    Invasive species can have major impacts on ecosystems, yet little work has addressed the combined effects of multiple invaders that exploit different habitats. Two common invaders in aquatic systems are pelagic fishes and crayfishes. Pelagic-oriented fish effects are typically strong on the pelagic food web, whereas crayfish effects are strong on the benthic food web. Thus, co-invasion may generate strong ecological responses in both habitats. We tested the effects of co-invasion on experimental pond ecosystems using two widespread invasive species, one pelagic (western mosquitofish) and one benthic (red swamp crayfish). As expected, mosquitofish had strong effects on the pelagic food web, reducing the abundance of Daphnia and causing a strong trophic cascade (increase in phytoplankton). Crayfish had strong effects on the benthic food web, reducing the abundance of benthic filamentous algae. Yet, we also found evidence for important cross-habitat effects. Mosquitofish treatments reduced the biomass of benthic filamentous algae, and crayfish treatments increased Daphnia and phytoplankton abundance. Combined effects of mosquitofish and crayfish were primarily positively or negatively additive, and completely offsetting for some responses, including gross primary production (GPP). Though co-invasion did not affect GPP, it strongly shifted primary production from the benthos into the water column. Effects on snail abundance revealed an interaction; snail abundance decreased only in the presence of both invaders. These results suggest that cross-habitat effects of co-invaders may lead to a variety of ecological outcomes; some of which may be unpredictable based on an understanding of each invader alone.

  14. Fossils of parasites: what can the fossil record tell us about the evolution of parasitism?

    PubMed

    Leung, Tommy L F

    2017-02-01

    Parasites are common in many ecosystems, yet because of their nature, they do not fossilise readily and are very rare in the geological record. This makes it challenging to study the evolutionary transition that led to the evolution of parasitism in different taxa. Most studies on the evolution of parasites are based on phylogenies of extant species that were constructed based on morphological and molecular data, but they give us an incomplete picture and offer little information on many important details of parasite-host interactions. The lack of fossil parasites also means we know very little about the roles that parasites played in ecosystems of the past even though it is known that parasites have significant influences on many ecosystems. The goal of this review is to bring attention to known fossils of parasites and parasitism, and provide a conceptual framework for how research on fossil parasites can develop in the future. Despite their rarity, there are some fossil parasites which have been described from different geological eras. These fossils include the free-living stage of parasites, parasites which became fossilised with their hosts, parasite eggs and propagules in coprolites, and traces of pathology inflicted by parasites on the host's body. Judging from the fossil record, while there were some parasite-host relationships which no longer exist in the present day, many parasite taxa which are known from the fossil record seem to have remained relatively unchanged in their general morphology and their patterns of host association over tens or even hundreds of millions of years. It also appears that major evolutionary and ecological transitions throughout the history of life on Earth coincided with the appearance of certain parasite taxa, as the appearance of new host groups also provided new niches for potential parasites. As such, fossil parasites can provide additional data regarding the ecology of their extinct hosts, since many parasites have

  15. Characterizing driver-response relationships in marine pelagic ecosystems for improved ocean management.

    PubMed

    Hunsicker, Mary E; Kappel, Carrie V; Selkoe, Kimberly A; Halpern, Benjamin S; Scarborough, Courtney; Mease, Lindley; Amrhein, Alisan

    2016-04-01

    Scientists and resource managers often use methods and tools that assume ecosystem components respond linearly to environmental drivers and human stressors. However, a growing body of literature demonstrates that many relationships are-non-linear, where small changes in a driver prompt a disproportionately large ecological response. We aim to provide a comprehensive assessment of the relationships between drivers and ecosystem components to identify where and when non-linearities are likely to occur. We focused our analyses on one of the best-studied marine systems, pelagic ecosystems, which allowed us to apply robust statistical techniques on a large pool of previously published studies. In this synthesis, we (1) conduct a wide literature review on single driver-response relationships in pelagic systems, (2) use statistical models to identify the degree of non-linearity in these relationships, and (3) assess whether general patterns exist in the strengths and shapes of non-linear relationships across drivers. Overall we found that non-linearities are common in pelagic ecosystems, comprising at least 52% of all driver-response relation- ships. This is likely an underestimate, as papers with higher quality data and analytical approaches reported non-linear relationships at a higher frequency (on average 11% more). Consequently, in the absence of evidence for a linear relationship, it is safer to assume a relationship is non-linear. Strong non-linearities can lead to greater ecological and socioeconomic consequences if they are unknown (and/or unanticipated), but if known they may provide clear thresholds to inform management targets. In pelagic systems, strongly non-linear relationships are often driven by climate and trophodynamic variables but are also associated with local stressors, such as overfishing and pollution, that can be more easily controlled by managers. Even when marine resource managers cannot influence ecosystem change, they can use information

  16. Right-handed fossil humans.

    PubMed

    Lozano, Marina; Estalrrich, Almudena; Bondioli, Luca; Fiore, Ivana; Bermúdez de Castro, José-Maria; Arsuaga, Juan Luis; Carbonell, Eudald; Rosas, Antonio; Frayer, David W

    2017-11-01

    Fossil hominids often processed material held between their upper and lower teeth. Pulling with one hand and cutting with the other, they occasionally left impact cut marks on the lip (labial) surface of their incisors and canines. From these actions, it possible to determine the dominant hand used. The frequency of these oblique striations in an array of fossil hominins documents the typically modern pattern of 9 right- to 1 left-hander. This ratio among living Homo sapiens differs from that among chimpanzees and bonobos and more distant primate relatives. Together, all studies of living people affirm that dominant right-handedness is a uniquely modern human trait. The same pattern extends deep into our past. Thus far, the majority of inferred right-handed fossils come from Europe, but a single maxilla from a Homo habilis, OH-65, shows a predominance of right oblique scratches, thus extending right-handedness into the early Pleistocene of Africa. Other studies show right-handedness in more recent African, Chinese, and Levantine fossils, but the sample compiled for non-European fossil specimens remains small. Fossil specimens from Sima del los Huesos and a variety of European Neandertal sites are predominately right-handed. We argue the 9:1 handedness ratio in Neandertals and the earlier inhabitants of Europe constitutes evidence for a modern pattern of handedness well before the appearance of modern Homo sapiens. © 2017 Wiley Periodicals, Inc.

  17. Composite biostratigraphy and microfacies analysis of the Upper Jurassic - Lower Cretaceous carbonate platform to slope successions in Sivrihisar (Eskişehir) region (NW Turkey, Pontides): Remarks on the palaeogeographic evolution of the Western Sakarya Zo

    NASA Astrophysics Data System (ADS)

    Atasoy, Serdar G.; Altıner, Demir; Okay, Aral I.

    2017-04-01

    , microencrusters, worm tubes, bivalve, crinoid and echinoid fragments). These deposits represent the background pelagic deposition on the slope. The slope facies are mainly composed of bioclastic-peloidal/ bioclastic-intraclastic packstone, rudstone-grainstone, bioclastic-lithoclastic floatstone-rudstone and reflect generally the increase in the amount of platform derived material (benthic foraminifera, microencrusters, worm tubes, corals, sponges, bryozoa). The matrix of these coarse grained deposits also contains pelagic taxa (calpionellids, radiolaria, Saccocoma sp., Globochaete sp., Pithonella sp., aptyhci). The slope facies are sometimes intercalataed with the toe-of-slope type facies indicating quiescence periods. The shallow marine carbonate platform deposits are characterized by peloidal-intraclastic poorly washed grainstone with bioclasts, bioclastic mudstone-wackestone, intraclastic packstone-rudstone and contain several shallow marine fossils (benthic foraminifera, encrustres and rare echnoid, bivalve and coral fragments) without any pelagic taxa. These carbonates are interpreted as back-reef platform deposits that should not be far away from the platform margin due to the co-occurence of Protopeneroplis striata and Mohlerina basiliensis, abundant in the shelf edge and reefal areas with the complex benthic foraminifera, Labyrinthina mirabilis common in lagoonal areas. If the position of the studied sections with respect to the EBBCP is considered, the studied basin and slope facies should represent the southern platform margin and slope environments of this carbonate platform that faced an ocean to the south during the Jurassic-Cretaceous. The slope and basinal facies overthrusted by the shallow marine deposits in a region situated to the south of the main İzmir-Ankara-Erzincan (İAE) suture suggests an important disruption and shortening of the EBBCP margin and slope deposits related to the closure of the İAE ocean.

  18. The non-uniformity of fossil preservation.

    PubMed

    Holland, Steven M

    2016-07-19

    The fossil record provides the primary source of data for calibrating the origin of clades. Although minimum ages of clades are given by the oldest preserved fossil, these underestimate the true age, which must be bracketed by probabilistic methods based on multiple fossil occurrences. Although most of these methods assume uniform preservation rates, this assumption is unsupported over geological timescales. On geologically long timescales (more than 10 Myr), the origin and cessation of sedimentary basins, and long-term variations in tectonic subsidence, eustatic sea level and sedimentation rate control the availability of depositional facies that preserve the environments in which species lived. The loss of doomed sediments, those with a low probability of preservation, imparts a secular trend to fossil preservation. As a result, the fossil record is spatially and temporally non-uniform. Models of fossil preservation should reflect this non-uniformity by using empirical estimates of fossil preservation that are spatially and temporally partitioned, or by using indirect proxies of fossil preservation. Geologically, realistic models of preservation will provide substantially more reliable estimates of the origination of clades.This article is part of the themed issue 'Dating species divergences using rocks and clocks'. © 2016 The Author(s).

  19. The non-uniformity of fossil preservation

    PubMed Central

    2016-01-01

    The fossil record provides the primary source of data for calibrating the origin of clades. Although minimum ages of clades are given by the oldest preserved fossil, these underestimate the true age, which must be bracketed by probabilistic methods based on multiple fossil occurrences. Although most of these methods assume uniform preservation rates, this assumption is unsupported over geological timescales. On geologically long timescales (more than 10 Myr), the origin and cessation of sedimentary basins, and long-term variations in tectonic subsidence, eustatic sea level and sedimentation rate control the availability of depositional facies that preserve the environments in which species lived. The loss of doomed sediments, those with a low probability of preservation, imparts a secular trend to fossil preservation. As a result, the fossil record is spatially and temporally non-uniform. Models of fossil preservation should reflect this non-uniformity by using empirical estimates of fossil preservation that are spatially and temporally partitioned, or by using indirect proxies of fossil preservation. Geologically, realistic models of preservation will provide substantially more reliable estimates of the origination of clades. This article is part of the themed issue ‘Dating species divergences using rocks and clocks’. PMID:27325828

  20. Fossilization of feathers

    NASA Astrophysics Data System (ADS)

    Davis, Paul G.; Briggs, Derek E. G.

    1995-09-01

    Scanning electron microscopy of feathers has revealed evidence that a bacterial glycocalyx (a network of exocellular polysaccharide fibers) played a role in promoting their fossilization in some cases. This mode of preservation has not been reported in other soft tissues. The majority of fossil feathers are preserved as carbonized traces. More rarely, bacteria on the surface are replicated by authigenic minerals (bacterial autolithification). The feathers of Archaeopteryx are preserved mainly by imprintation following early lithification of the substrate and decay of the feather. Lacustrine settings provide the most important taphonomic window for feather preservation. Preservation in terrestrial and normal-marine settings involves very different processes (in amber and in authigenically mineralized coprolites, respectively). Therefore, there may be a significant bias in the avian fossil record in favor of inland water habitats.

  1. Pelagic communities of the South West Indian Ocean seamounts: R/V Dr Fridtjof Nansen Cruise 2009-410

    NASA Astrophysics Data System (ADS)

    Rogers, A. D.; Alvheim, O.; Bemanaja, E.; Benivary, D.; Boersch-Supan, P.; Bornman, T. G.; Cedras, R.; Du Plessis, N.; Gotheil, S.; Høines, A.; Kemp, K.; Kristiansen, J.; Letessier, T.; Mangar, V.; Mazungula, N.; Mørk, T.; Pinet, P.; Pollard, R.; Read, J.; Sonnekus, T.

    2017-02-01

    The seamounts of the southern Indian Ocean remain some of the most poorly studied globally and yet have been subject to deep-sea fishing for decades and may face new exploitation through mining of seabed massive sulphides in the future. As an attempt to redress the knowledge deficit on deep-sea benthic and pelagic communities associated mainly with the seamounts of the South West Indian Ridge two cruises were undertaken to explore the pelagic and benthic ecology in 2009 and 2011 respectively. In this volume are presented studies on pelagic ecosystems around six seamounts, five on the South West Indian Ridge, including Atlantis Bank, Sapmer Seamount, Middle of What Seamount, Melville Bank and Coral Seamount and one un-named seamount on the Madagascar Ridge. In this paper, existing knowledge on the seamounts of the southwestern Indian Ocean is presented to provide context for the studies presented in this volume. An account of the overall aims, approaches and methods used primarily on the 2009 cruise are presented including metadata associated with sampling and some of the limitations of the study. Sampling during this cruise included physical oceanographic measurements, multibeam bathymetry, biological acoustics, and net sampling of phytoplankton, macrozooplankton and micronekton/nekton. The studies that follow reveal new data on the physical oceanography of this dynamic region of the oceans, and the important influence of water masses on the pelagic ecology associated with the seamounts of the South West Indian Ridge. New information on the pelagic fauna of the region fills an important biogeographic gap for the mid- to high-latitudes of the oceans of the southern hemisphere.

  2. Hypoxia Impacts on Food Web Linkages in a Pelagic Ecosystem

    NASA Astrophysics Data System (ADS)

    Sato, M.; Horne, J. K.; Parker-Stetter, S. L.; Essington, T.; Keister, J. E.; Moriarty, P.; Li, L.

    2016-02-01

    Low dissolved oxygen (DO), or hypoxia, causes significant disturbances on aquatic organisms, but the consequences for key food web linkages is not well understood. Here, we tested how the intensity of low DO events governs the degree of spatial overlap between pelagic zooplanktivorous fish and their zooplankton prey, fish feeding rates, and community compositions of zooplankton. We hypothesized that the greater sensitivity of fish to DO compared to zooplankton would lead to diminished spatial overlap at moderate DO and reduced feeding rates of fish, while severe hypoxia would amplify spatial overlap by preventing zooplankton from using deep refuge habitats leading to increased fish feeding rates. We also hypothesized shifts in zooplankton community composition towards less energetically profitable taxa such as small copepods and gelatinous species. We used a combination of multifrequency acoustic and net sampling for detecting distributions and abundance of zooplankton and pelagic fish in Hood Canal, WA, a seasonally hypoxic fjord. We employed a sampling design which paired hypoxic regions of Hood Canal with normoxic regions sampled prior to, during, and after the onset of hypoxia in two years. Contrary to our hypotheses, we found that fish and zooplankton did not change their horizontal and vertical distributions during periods and in locations with low DO levels. Consequently, the vertical overlap between fish and zooplankton did not change with DO. Fish feeding rates and the dominant zooplankton prey did not change with hypoxia events. The apparent resilience of fish to low DO in our system may be explained by decreased metabolic oxygen demand due to cool temperatures, increased availability and accessibility to their prey in low DO waters, or potential increase in predation risk at shallower depth. This study highlights the importance of both temperature and DO, instead of hypoxia threshold alone, in evaluating the impacts of hypoxia on pelagic communities.

  3. Influence of variable rates of neritic carbonate deposition on atmospheric carbon dioxide and pelagic sediments

    NASA Technical Reports Server (NTRS)

    Walker, J. C.; Opdyke, B. C.

    1995-01-01

    Short-term imbalances in the global cycle of shallow water calcium carbonate deposition and dissolution may be responsible for much of the observed Pleistocene change in atmospheric carbon dioxide content. However, any proposed changes in the alkalinity balance of the ocean must be reconciled with the sedimentary record of deep-sea carbonates. The possible magnitude of the effect of shallow water carbonate deposition on the dissolution of pelagic carbonate can be tested using numerical simulations of the global carbon cycle. Boundary conditions can be defined by using extant shallow water carbonate accumulation data and pelagic carbonate deposition/dissolution data. On timescales of thousands of years carbonate deposition versus dissolution is rarely out of equilibrium by more than 1.5 x 10(13) mole yr-1. Results indicate that the carbonate chemistry of the ocean is rarely at equilibrium on timescales less than 10 ka. This disequilibrium is probably due to sea level-induced changes in shallow water calcium carbonate deposition/dissolution, an interpretation that does not conflict with pelagic sedimentary data from the central Pacific.

  4. The original colours of fossil beetles

    PubMed Central

    McNamara, Maria E.; Briggs, Derek E. G.; Orr, Patrick J.; Noh, Heeso; Cao, Hui

    2012-01-01

    Structural colours, the most intense, reflective and pure colours in nature, are generated when light is scattered by complex nanostructures. Metallic structural colours are widespread among modern insects and can be preserved in their fossil counterparts, but it is unclear whether the colours have been altered during fossilization, and whether the absence of colours is always real. To resolve these issues, we investigated fossil beetles from five Cenozoic biotas. Metallic colours in these specimens are generated by an epicuticular multi-layer reflector; the fidelity of its preservation correlates with that of other key cuticular ultrastructures. Where these other ultrastructures are well preserved in non-metallic fossil specimens, we can infer that the original cuticle lacked a multi-layer reflector; its absence in the fossil is not a preservational artefact. Reconstructions of the original colours of the fossils based on the structure of the multi-layer reflector show that the preserved colours are offset systematically to longer wavelengths; this probably reflects alteration of the refractive index of the epicuticle during fossilization. These findings will allow the former presence, and original hue, of metallic structural colours to be identified in diverse fossil insects, thus providing critical evidence of the evolution of structural colour in this group. PMID:21957131

  5. The original colours of fossil beetles.

    PubMed

    McNamara, Maria E; Briggs, Derek E G; Orr, Patrick J; Noh, Heeso; Cao, Hui

    2012-03-22

    Structural colours, the most intense, reflective and pure colours in nature, are generated when light is scattered by complex nanostructures. Metallic structural colours are widespread among modern insects and can be preserved in their fossil counterparts, but it is unclear whether the colours have been altered during fossilization, and whether the absence of colours is always real. To resolve these issues, we investigated fossil beetles from five Cenozoic biotas. Metallic colours in these specimens are generated by an epicuticular multi-layer reflector; the fidelity of its preservation correlates with that of other key cuticular ultrastructures. Where these other ultrastructures are well preserved in non-metallic fossil specimens, we can infer that the original cuticle lacked a multi-layer reflector; its absence in the fossil is not a preservational artefact. Reconstructions of the original colours of the fossils based on the structure of the multi-layer reflector show that the preserved colours are offset systematically to longer wavelengths; this probably reflects alteration of the refractive index of the epicuticle during fossilization. These findings will allow the former presence, and original hue, of metallic structural colours to be identified in diverse fossil insects, thus providing critical evidence of the evolution of structural colour in this group.

  6. Climate change impacts on leatherback turtle pelagic habitat in the Southeast Pacific

    NASA Astrophysics Data System (ADS)

    Willis-Norton, Ellen; Hazen, Elliott L.; Fossette, Sabrina; Shillinger, George; Rykaczewski, Ryan R.; Foley, David G.; Dunne, John P.; Bograd, Steven J.

    2015-03-01

    Eastern Pacific populations of the leatherback turtle (Dermochelys coriacea) have declined by over 90% during the past three decades. The decline is primarily attributed to human pressures, including unsustainable egg harvest, development on nesting beaches, and by-catch mortality. In particular, the effects of climate change may impose additional stresses upon already threatened leatherback populations. This study analyzes how the pelagic habitat of Eastern Pacific leatherbacks may be affected by climate change over the next century. This population adheres to a persistent migration pattern; following nesting at Playa Grande, Costa Rica, individuals move rapidly through equatorial currents and into foraging habitat within the oligotrophic South Pacific Gyre. Forty-six nesting females were fitted with satellite tags. Based on the turtle positions, ten environmental variables were sampled along the tracks. Presence/absence habitat models were created to determine the oceanographic characteristics of the preferred turtle habitat. Core pelagic habitat was characterized by relatively low sea surface temperatures and chlorophyll-a. Based on these habitat models, we predicted habitat change using output from the Geophysical Fluid Dynamics Laboratory prototype Earth System Model under the Special Report on Emissions Scenario A2 (business-as-usual). Although the model predicted both habitat losses and gains throughout the region, we estimated that overall the core pelagic habitat of the Eastern Pacific leatherback population will decline by approximately 15% within the next century. This habitat modification might increase pressure on a critically endangered population, possibly forcing distributional shifts, behavioral changes, or even extinction.

  7. Persistence of an unusual pelagic zooplankton assemblage in a clear, mountain lake

    USGS Publications Warehouse

    Larson, G.L.; Hoffman, R.L.; C. David, McIntire

    2002-01-01

    The planktonic zooplankton assemblage in Mowich Lake, Mount Rainier National Park (MORA), was composed almost entirely of rotifers in 1966 and 1967. Adult pelagic crustacean taxa were rare. Their paucity was attributed to predation by kokanee salmon (Oncorhynchus nerka), which had been stocked in 1961. During a park-wide survey of 24 lakes in 1988, Mowich Lake was the only one that did not contain at least one planktonic crustacean species. Given the apparent persistence of the unusual pelagic zooplankton assemblage in Mowich Lake, the first objective of this study was to document the interannual variation in the taxonomic structure of the zooplankton assemblages in the lake from 1988 through 1999. A second objective was to determine if it was possible to predict the taxonomic composition of the pelagic crustacean zooplankton assemblage in Mowich Lake prior to the stocking of kokanee salmon. The Mowich Lake zooplankton assemblages in 1988-1999 were consistent with those in 1966 and 1967. Crustacean taxa were extremely rare, but they included most of the primary taxa collected from 23 MORA lakes surveyed in 1988. Nonetheless, the 1988 collections showed that the September rotifer assemblage in Mowich Lake was similar to 10 of the 24 lakes sampled. Seven of the 10 lakes were dominated by cladocerans, primarily Daphnia rosea and Holopedium gibberum. Therefore, it appeared that either one or both of these species may have numerically dominated the crustacean zooplankton assemblage in the lake prior to 1961.

  8. Persistence of an unusual pelagic zooplankton assemblage in a clear mountain lake

    USGS Publications Warehouse

    Larson, Gary L.; Hoffman, Robert L.; McIntire, C.D.

    2002-01-01

    The planktonic zooplankton assemblage in Mowich Lake, Mount Rainier National Park (MORA), was composed almost entirely of rotifers in 1966 and 1967. Adult pelagic crustacean taxa were rare. Their paucity was attributed to predation by kokanee salmon (Oncorhynchus nerka), which had been stocked in 1961. During a park-wide survey of 24 lakes in 1988, Mowich Lake was the only one that did not contain at least one planktonic crustacean species. Given the apparent persistence of the unusual pelagic zooplankton assemblage in Mowich Lake, the first objective of this study was to document the interannual variation in the taxonomic structure of the zooplankton assemblages in the lake from 1988 through 1999. A second objective was to determine if it was possible to predict the taxonomic composition of the pelagic crustacean zooplankton assemblage in Mowich Lake prior to the stocking of kokanee salmon. The Mowich Lake zooplankton assemblages in 1988a??1999 were consistent with those in 1966 and 1967. Crustacean taxa were extremely rare, but they included most of the primary taxa collected from 23 MORA lakes surveyed in 1988. Nonetheless, the 1988 collections showed that the September rotifer assemblage in Mowich Lake was similar to 10 of the 24 lakes sampled. Seven of the 10 lakes were dominated by cladocerans, primarily Daphnia rosea and Holopedium gibberum. Therefore, it appeared that either one or both of these species may have numerically dominated the crustacean zooplankton assemblage in the lake prior to 1961.

  9. The collapse of pelagic fishes in the upper San Francisco estuary

    USGS Publications Warehouse

    Sommer, T.; Armor, C.; Baxter, R.; Breuer, R.; Brown, L.; Chotkowski, M.; Culberson, S.; Feyrer, F.; Gingras, M.; Herbold, B.; Kimmerer, W.; Mueller-Solger, A.; Nobriga, M.; Souza, K.

    2007-01-01

    Although the pelagic fish community of the upper San Francisco Estuary historically has shown substantial variability, a recent collapse has captured the attention of resource managers, scientists, legislators, and the general public. The ecological and management consequences of the decline are most serious for delta smelt (Hypomesus transpacificus), a threatened species whose narrow range overlaps with large water diversions that supply water to over 25 million people. The decline occurred despite recent moderate hydrology, which typically results in at least modest recruitment, and investments of hundreds of millions of dollars in habitat restoration and environmental water allocations to support native fishes. In response to the pelagic fish collapse, an ambitious multi-agency research team has been working since 2005 to evaluate the causes of the decline, which likely include a combination of factors: stock-recruitment effects, a decline in habitat quality, increased mortality rates, and reduced food availability due to invasive species.

  10. Bioaccumulation of mercury in pelagic fishes from the northern Gulf of Mexico

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Cai, Yang J.; Rooker, Jay R.; Gill, Gary A.

    2007-03-30

    Total mercury (Hg) concentration was determined in the tissues of 10 pelagic fishes in the northern Golf of Mexico, and dietary tracers (stable isotopes and fatty acids) were used to evaluate the relationship between Hg and feeding history.

  11. Earth's early fossil record: Why not look for similar fossils on Mars?

    NASA Technical Reports Server (NTRS)

    Awramik, Stanley M.

    1989-01-01

    The oldest evidence of life on Earth is discussed with attention being given to the structure and formation of stromatolites and microfossils. Fossilization of microbes in calcium carbonate or chert media is discussed. In searching for fossil remains on Mars, some lessons learned from the study of Earth's earliest fossil record can be applied. Certain sedimentary rock types and sedimentary rock configurations should be targeted for investigation and returned by the Martian rover and ultimately by human explorers. Domical, columnar to wavy laminated stratiform sedimentary rocks that resemble stromatolites should be actively sought. Limestone, other carbonates, and chert are the favored lithology. Being macroscopic, stromatolites might be recognized by an intelligent unmanned rover. In addition, black, waxy chert with conchoidal fracture should be sought. Chert is by far the preferred lithology for the preservation of microbes and chemical fossils. Even under optimal geological conditions (little or no metamorphism or tectonic alteration, excellent outcrops, and good black chert) and using experienced field biogeologists, the chances of finding well preserved microbial remains in chert are very low.

  12. Potential impacts of the Deepwater Horizon oil spill on large pelagic fishes

    NASA Astrophysics Data System (ADS)

    Frias-Torres, Sarrah; Bostater, Charles R., Jr.

    2011-11-01

    Biogeographical analyses provide insights on how the Deepwater Horizon oil spill impacted large pelagic fishes. We georeferenced historical ichthyoplankton surveys and published literature to map the spawning and larval areas of bluefin tuna, swordfish, blue marlin and whale shark sightings in the Gulf of Mexico with daily satellite-derived images detecting surface oil. The oil spill covered critical areas used by large pelagic fishes. Surface oil was detected in 100% of the northernmost whale shark sightings, in 32.8 % of the bluefin tuna spawning area and 38 % of the blue marlin larval area. No surface oil was detected in the swordfish spawning and larval area. Our study likely underestimates the extend of the oil spill due to satellite sensors detecting only the upper euphotic zone and the use of dispersants altering crude oil density, but provides a previously unknown spatio-temporal analysis.

  13. Vertical structure, biomass and topographic association of deep-pelagic fishes in relation to a mid-ocean ridge system

    NASA Astrophysics Data System (ADS)

    Sutton, T. T.; Porteiro, F. M.; Heino, M.; Byrkjedal, I.; Langhelle, G.; Anderson, C. I. H.; Horne, J.; Søiland, H.; Falkenhaug, T.; Godø, O. R.; Bergstad, O. A.

    2008-01-01

    The assemblage structure and vertical distribution of deep-pelagic fishes relative to a mid-ocean ridge system are described from an acoustic and discrete-depth trawling survey conducted as part of the international Census of Marine Life field project MAR-ECO < http://www.mar-eco.no>. The 36-station, zig-zag survey along the northern Mid-Atlantic Ridge (MAR; Iceland to the Azores) covered the full depth range (0 to >3000 m), from the surface to near the bottom, using a combination of gear types to gain a more comprehensive understanding of the pelagic fauna. Abundance per volume of deep-pelagic fishes was highest in the epipelagic zone and within the benthic boundary layer (BBL; 0-200 m above the seafloor). Minimum fish abundance occurred at depths below 2300 m but above the BBL. Biomass per volume of deep-pelagic fishes over the MAR reached a maximum within the BBL, revealing a previously unknown topographic association of a bathypelagic fish assemblage with a mid-ocean ridge system. With the exception of the BBL, biomass per volume reached a water column maximum in the bathypelagic zone between 1500 and 2300 m. This stands in stark contrast to the general "open-ocean" paradigm that biomass decreases exponentially from the surface downwards. As much of the summit of the MAR extends into this depth layer, a likely explanation for this mid-water maximum is ridge association. Multivariate statistical analyses suggest that the dominant component of deep-pelagic fish biomass over the northern MAR was a wide-ranging bathypelagic assemblage that was remarkably consistent along the length of the ridge from Iceland to the Azores. Integrating these results with those of previous studies in oceanic ecosystems, there appears to be adequate evidence to conclude that special hydrodynamic and biotic features of mid-ocean ridge systems cause changes in the ecological structure of deep-pelagic fish assemblages relative to those at the same depths over abyssal plains. Lacking

  14. Natural Product Molecular Fossils.

    PubMed

    Falk, Heinz; Wolkenstein, Klaus

    The natural products synthesized by organisms that were living a long time ago gave rise to their molecular fossils. These can consist of either the original unchanged compounds or they may undergo peripheral transformations in which their skeletons remain intact. In cases when molecular fossils can be traced to their organismic source, they are termed "geological biomarkers".This contribution describes apolar and polar molecular fossils and, in particular biomarkers, along the lines usually followed in organic chemistry textbooks, and points to their bioprecursors when available. Thus, the apolar compounds are divided in linear and branched alkanes followed by alicyclic compounds and aromatic and heterocyclic molecules, and, in particular, the geoporphyrins. The polar molecular fossils contain as functional groups or constituent units ethers, alcohols, phenols, carbonyl groups, flavonoids, quinones, and acids, or are polymers like kerogen, amber, melanin, proteins, or nucleic acids. The final sections discuss the methodology used and the fundamental processes encountered by the biomolecules described, including diagenesis, catagenesis, and metagenesis.

  15. The legacy of fossil fuels.

    PubMed

    Armaroli, Nicola; Balzani, Vincenzo

    2011-03-01

    Currently, over 80% of the energy used by mankind comes from fossil fuels. Harnessing coal, oil and gas, the energy resources contained in the store of our spaceship, Earth, has prompted a dramatic expansion in energy use and a substantial improvement in the quality of life of billions of individuals in some regions of the world. Powering our civilization with fossil fuels has been very convenient, but now we know that it entails severe consequences. We treat fossil fuels as a resource that anyone anywhere can extract and use in any fashion, and Earth's atmosphere, soil and oceans as a dump for their waste products, including more than 30 Gt/y of carbon dioxide. At present, environmental legacy rather than consistence of exploitable reserves, is the most dramatic problem posed by the relentless increase of fossil fuel global demand. Harmful effects on the environment and human health, usually not incorporated into the pricing of fossil fuels, include immediate and short-term impacts related to their discovery, extraction, transportation, distribution, and burning as well as climate change that are spread over time to future generations or over space to the entire planet. In this essay, several aspects of the fossil fuel legacy are discussed, such as alteration of the carbon cycle, carbon dioxide rise and its measurement, greenhouse effect, anthropogenic climate change, air pollution and human health, geoengineering proposals, land and water degradation, economic problems, indirect effects on the society, and the urgent need of regulatory efforts and related actions to promote a gradual transition out of the fossil fuel era. While manufacturing sustainable solar fuels appears to be a longer-time perspective, alternatives energy sources already exist that have the potential to replace fossil fuels as feedstocks for electricity production. Copyright © 2011 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim.

  16. Regeneration of the digestive system in the crinoid Himerometra robustipinna occurs by transdifferentiation of neurosecretory-like cells

    PubMed Central

    Kalacheva, Nadezhda V.; Eliseikina, Marina G.; Frolova, Lidia T.

    2017-01-01

    The structure and regeneration of the digestive system in the crinoid Himerometra robustipinna (Carpenter, 1881) were studied. The gut comprises a spiral tube forming radial lateral processes, which gives it a five-lobed shape. The digestive tube consists of three segments: esophagus, intestine, and rectum. The epithelia of these segments have different cell compositions. Regeneration of the gut after autotomy of the visceral mass progresses very rapidly. Within 6 h after autotomy, an aggregation consisting of amoebocytes, coelomic epithelial cells and juxtaligamental cells (neurosecretory neurons) forms on the inner surface of the skeletal calyx. At 12 h post-autotomy, transdifferentiation of the juxtaligamental cells starts. At 24 h post-autotomy these cells undergo a mesenchymal-epithelial-like transition, resulting in the formation of the luminal epithelium of the gut. Specialization of the intestinal epithelial cells begins on day 2 post-autotomy. At this stage animals acquire the mouth and anal opening. On day 4 post-autotomy the height of both the enterocytes and the visceral mass gradually increases. Proliferation does not play any noticeable role in gut regeneration. The immersion of animals in a 10−7 M solution of colchicine neither stopped formation of the lost structures nor caused accumulation of mitoses in tissues. Weakly EdU-labeled nuclei were observed in the gut only on day 2 post-autotomy and were not detected at later regeneration stages. Single mitotically dividing cells were recorded during the same period. It is concluded that juxtaligamental cells play a major role in gut regeneration in H. robustipinna. The main mechanisms of morphogenesis are cell migration and transdifferentiation. PMID:28753616

  17. Regeneration of the digestive system in the crinoid Himerometra robustipinna occurs by transdifferentiation of neurosecretory-like cells.

    PubMed

    Kalacheva, Nadezhda V; Eliseikina, Marina G; Frolova, Lidia T; Dolmatov, Igor Yu

    2017-01-01

    The structure and regeneration of the digestive system in the crinoid Himerometra robustipinna (Carpenter, 1881) were studied. The gut comprises a spiral tube forming radial lateral processes, which gives it a five-lobed shape. The digestive tube consists of three segments: esophagus, intestine, and rectum. The epithelia of these segments have different cell compositions. Regeneration of the gut after autotomy of the visceral mass progresses very rapidly. Within 6 h after autotomy, an aggregation consisting of amoebocytes, coelomic epithelial cells and juxtaligamental cells (neurosecretory neurons) forms on the inner surface of the skeletal calyx. At 12 h post-autotomy, transdifferentiation of the juxtaligamental cells starts. At 24 h post-autotomy these cells undergo a mesenchymal-epithelial-like transition, resulting in the formation of the luminal epithelium of the gut. Specialization of the intestinal epithelial cells begins on day 2 post-autotomy. At this stage animals acquire the mouth and anal opening. On day 4 post-autotomy the height of both the enterocytes and the visceral mass gradually increases. Proliferation does not play any noticeable role in gut regeneration. The immersion of animals in a 10-7 M solution of colchicine neither stopped formation of the lost structures nor caused accumulation of mitoses in tissues. Weakly EdU-labeled nuclei were observed in the gut only on day 2 post-autotomy and were not detected at later regeneration stages. Single mitotically dividing cells were recorded during the same period. It is concluded that juxtaligamental cells play a major role in gut regeneration in H. robustipinna. The main mechanisms of morphogenesis are cell migration and transdifferentiation.

  18. 50 CFR 229.36 - Atlantic Pelagic Longline Take Reduction Plan (PLTRP).

    Code of Federal Regulations, 2014 CFR

    2014-10-01

    ... information on marine mammal interactions, fishing operations, marine mammal life history information, and..., or intend to do so, you must call NMFS Southeast Fisheries Science Center (SEFSC), 1-888-254-2558, at... from the Director, NMFS Southeast Fishery Science Center to use a pelagic longline exceeding 20 nm (37...

  19. 50 CFR 229.36 - Atlantic Pelagic Longline Take Reduction Plan (PLTRP).

    Code of Federal Regulations, 2012 CFR

    2012-10-01

    ... information on marine mammal interactions, fishing operations, marine mammal life history information, and..., or intend to do so, you must call NMFS Southeast Fisheries Science Center (SEFSC), 1-888-254-2558, at... from the Director, NMFS Southeast Fishery Science Center to use a pelagic longline exceeding 20 nm (37...

  20. 50 CFR 229.36 - Atlantic Pelagic Longline Take Reduction Plan (PLTRP).

    Code of Federal Regulations, 2013 CFR

    2013-10-01

    ... information on marine mammal interactions, fishing operations, marine mammal life history information, and..., or intend to do so, you must call NMFS Southeast Fisheries Science Center (SEFSC), 1-888-254-2558, at... from the Director, NMFS Southeast Fishery Science Center to use a pelagic longline exceeding 20 nm (37...

  1. 50 CFR 229.36 - Atlantic Pelagic Longline Take Reduction Plan (PLTRP).

    Code of Federal Regulations, 2011 CFR

    2011-10-01

    ... information on marine mammal interactions, fishing operations, marine mammal life history information, and..., or intend to do so, you must call NMFS Southeast Fisheries Science Center (SEFSC), 1-888-254-2558, at... from the Director, NMFS Southeast Fishery Science Center to use a pelagic longline exceeding 20 nm (37...

  2. 50 CFR 229.36 - Atlantic Pelagic Longline Take Reduction Plan (PLTRP).

    Code of Federal Regulations, 2010 CFR

    2010-10-01

    ... information on marine mammal interactions, fishing operations, marine mammal life history information, and..., or intend to do so, you must call NMFS Southeast Fisheries Science Center (SEFSC), 1-888-254-2558, at... from the Director, NMFS Southeast Fishery Science Center to use a pelagic longline exceeding 20 nm (37...

  3. Arthropod visual predators in the early pelagic ecosystem: evidence from the Burgess Shale and Chengjiang biotas

    PubMed Central

    Vannier, J.; García-Bellido, D.C.; Hu, S.-X.; Chen, A.-L.

    2009-01-01

    Exceptional fossil specimens with preserved soft parts from the Maotianshan Shale (ca 520 Myr ago) and the Burgess Shale (505 Myr ago) biotas indicate that the worldwide distributed bivalved arthropod Isoxys was probably a non-benthic visual predator. New lines of evidence come from the functional morphology of its powerful prehensile frontal appendages that, combined with large spherical eyes, are thought to have played a key role in the recognition and capture of swimming or epibenthic prey. The swimming and steering of this arthropod was achieved by the beating of multiple setose exopods and a flap-like telson. The appendage morphology of Isoxys indicates possible phylogenetical relationships with the megacheirans, a widespread group of assumed predator arthropods characterized by a pre-oral ‘great appendage’. Evidence from functional morphology and taphonomy suggests that Isoxys was able to migrate through the water column and was possibly exploiting hyperbenthic niches for food. Although certainly not unique, the case of Isoxys supports the idea that off-bottom animal interactions such as predation, associated with complex feeding strategies and behaviours (e.g. vertical migration and hunting) were established by the Early Cambrian. It also suggests that a prototype of a pelagic food chain had already started to build-up at least in the lower levels of the water column. PMID:19403536

  4. Looking at Fossils in New Ways

    ERIC Educational Resources Information Center

    Flannery, Maura C.

    2005-01-01

    Existing fossils could be studied from a different prospective with the use of new methods of analysis for gathering more information. The new techniques of studying fossils binds the new and the old techniques and information and provides another way to look at fossils.

  5. Storms in the deep: tempestite- and beach-like deposits in pelagic sequences (Middle-Upper Jurassic, Subbetic, South of Spain)

    NASA Astrophysics Data System (ADS)

    Pomar, Luis; Molina, Jose M.; Ruiz-Ortiz, Pedro A.; Vera, Juan A.

    2017-04-01

    Fine-peloidal- to coarse oolitic-bioclastic grainstones with hummocky cross stratification (HCS) occur interbedded in Middle-Upper Jurassic pelagic lime-mudstone successions (Betic ranges, Southern Spain). These strata were deposited in pelagic troughs and swells, away from continental areas, in the Southern Iberian Continental Margin of the Western Tethys. Previously interpreted as tempestites, mainly due to the attribution of the HCS to surface storm waves, they are now reinterpreted as the product of turbulence in deeper conditions. Among many, some selected examples are here presented. All of them share: 1) Grainstone beds are interbedded with pelagic mudstones and marls 2) Grainstone components were reworked by oscillatory flows superimposed to unidirectional tractive flows (unidirectional ripple lamination and HCS). 3) Components were either derived from shallow-water environments (e.g., ooids), or produced in pelagic conditions (e.g., radiolarians, Saccocoma, peloids, etc). 4) Although surface-storm tempestite flows can be required to bring downslope components from shallow-water settings, the grainstone beds reflect sediment reworking at a depth dominated by fine pelagic sedimentation. 5) Internal waves propagating along a pycnocline and breaking against a sloping surface are the best candidate to induce the sedimentary structures and sediment organization that characterize these grainstone beds. The examples here presented (Middle-Upper Jurassic of the Subbetic) include: a) Peloid grainstones interbedded with radiolarite marls deposited on the flanks of volcanic guyots. The interbedded lime muds and marls contain 'filaments', sponge spicules and radiolarians. b) Peloid-bioclastic (radiolarians, Saccocoma, etc.) grainstone beds with HCS, interbedded with pelagic lime muds. c) Coarse oolitic grainstone unit, encased in pelagic marls, with wedge-shaped crossbed-sets with gently seaward-dipping parallel lamination, and sets of low-angle up-slope dipping

  6. Remotely sensed wind speed predicts soaring behaviour in a wide-ranging pelagic seabird.

    PubMed

    Gibb, Rory; Shoji, Akiko; Fayet, Annette L; Perrins, Chris M; Guilford, Tim; Freeman, Robin

    2017-07-01

    Global wind patterns affect flight strategies in many birds, including pelagic seabirds, many of which use wind-powered soaring to reduce energy costs during at-sea foraging trips and migration. Such long-distance movement patterns are underpinned by local interactions between wind conditions and flight behaviour, but these fine-scale relationships are far less well understood. Here we show that remotely sensed ocean wind speed and direction are highly significant predictors of soaring behaviour in a migratory pelagic seabird, the Manx shearwater ( Puffinus puffinus ). We used high-frequency GPS tracking data (10 Hz) and statistical behaviour state classification to identify two energetic modes in at-sea flight, corresponding to flap-like and soar-like flight. We show that soaring is significantly more likely to occur in tailwinds and crosswinds above a wind speed threshold of around 8 m s -1 , suggesting that these conditions enable birds to reduce metabolic costs by preferentially soaring over flapping. Our results suggest a behavioural mechanism by which wind conditions may shape foraging and migration ecology in pelagic seabirds, and thus indicate that shifts in wind patterns driven by climate change could impact this and other species. They also emphasize the emerging potential of high-frequency GPS biologgers to provide detailed quantitative insights into fine-scale flight behaviour in free-living animals. © 2017 The Author(s).

  7. Faunas of Mississippian oolitic limestones: Evidence from Salem Limestone, southern Indiana

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Feldman, H.R.

    In the Salem Limestone of southern Indiana, a correlation exists between the faunal assemblage and abundance of grains with superficial oolitic coatings in grainstones. Coarse, poorly sorted fossiliferous grainstones are dominated by an echinoderm-bryozoan-brachiopod assemblage of fossils with few mollusks. The presence of large whole fossils, such as articulated crinoid calyces, suggest limited transport of skeletal components. Grainstones, dominated by grains with superficial coatings, and foraminifers tend to contain a diverse mollusk-dominated assemblage of gastropods, bivalves, rostroconchs, chitins, and scaphopods. These fossils are disarticulated, but generally are not fragmented even though many of them are thin and delicate. Echinoderms, brachiopods,more » and bryozoans are repsented in the mollusk-domdinated assemblage almost exclusively by well-rounded and coated fragments, suggesting that they are not in situ. The presence of similar molluscan assemblages in other Mississippian coated-grain grainstones from Alabama (the Monteagle Limestone) and Oklahoma (an unnamed limestone) indicates that the assemblage may have been wide-spread. Mississippian grainstones dominated by oolites (which are not prominent in the Salem) generally have very few fossils.« less

  8. Elasmobranch bycatch in the Italian Adriatic pelagic trawl fishery

    PubMed Central

    Fortuna, Caterina Maria; Moro, Fabrizio; Sala, Antonello

    2018-01-01

    Elasmobranchs are among the most threatened long-lived marine species worldwide, and incidental capture is a major source of mortality. The northern central Adriatic Sea, though one of the most overfished basins of the Mediterranean Sea, supports a very valuable marine biodiversity, including elasmobranchs. This study assesses the impact of the northern central Adriatic pelagic trawl fishery on common smooth-hound (Mustelus mustelus), spiny dogfish (Squalus acanthias), common eagle ray (Myliobatis aquila), and pelagic stingray (Pteroplatytrygon violacea) by examining incidental catches recorded between 2006 and 2015. The distribution of bycatch events was evaluated using geo-referenced data. Generalized Linear Models were computed to standardize the catch of the four species and to predict the relative abundance of bycatch events. Data analysis shows that most bycatch events involving all four species occurred in the northern Adriatic Sea. The models predicted significant, distinct temporal patterns of standardized catches in line with previous investigations. Water depth, season, and fishing region were the best predictors to explain bycatch events. The present data suggest that the northern Adriatic may be an important nursery area for several elasmobranchs. They also highlight the urgent need for a better understanding of the interactions between elasmobranchs and fisheries to develop and apply suitable, ad hoc management measures. PMID:29377920

  9. Elasmobranch bycatch in the Italian Adriatic pelagic trawl fishery.

    PubMed

    Bonanomi, Sara; Pulcinella, Jacopo; Fortuna, Caterina Maria; Moro, Fabrizio; Sala, Antonello

    2018-01-01

    Elasmobranchs are among the most threatened long-lived marine species worldwide, and incidental capture is a major source of mortality. The northern central Adriatic Sea, though one of the most overfished basins of the Mediterranean Sea, supports a very valuable marine biodiversity, including elasmobranchs. This study assesses the impact of the northern central Adriatic pelagic trawl fishery on common smooth-hound (Mustelus mustelus), spiny dogfish (Squalus acanthias), common eagle ray (Myliobatis aquila), and pelagic stingray (Pteroplatytrygon violacea) by examining incidental catches recorded between 2006 and 2015. The distribution of bycatch events was evaluated using geo-referenced data. Generalized Linear Models were computed to standardize the catch of the four species and to predict the relative abundance of bycatch events. Data analysis shows that most bycatch events involving all four species occurred in the northern Adriatic Sea. The models predicted significant, distinct temporal patterns of standardized catches in line with previous investigations. Water depth, season, and fishing region were the best predictors to explain bycatch events. The present data suggest that the northern Adriatic may be an important nursery area for several elasmobranchs. They also highlight the urgent need for a better understanding of the interactions between elasmobranchs and fisheries to develop and apply suitable, ad hoc management measures.

  10. Soft-Bodied Fossils Are Not Simply Rotten Carcasses - Toward a Holistic Understanding of Exceptional Fossil Preservation: Exceptional Fossil Preservation Is Complex and Involves the Interplay of Numerous Biological and Geological Processes.

    PubMed

    Parry, Luke A; Smithwick, Fiann; Nordén, Klara K; Saitta, Evan T; Lozano-Fernandez, Jesus; Tanner, Alastair R; Caron, Jean-Bernard; Edgecombe, Gregory D; Briggs, Derek E G; Vinther, Jakob

    2018-01-01

    Exceptionally preserved fossils are the product of complex interplays of biological and geological processes including burial, autolysis and microbial decay, authigenic mineralization, diagenesis, metamorphism, and finally weathering and exhumation. Determining which tissues are preserved and how biases affect their preservation pathways is important for interpreting fossils in phylogenetic, ecological, and evolutionary frameworks. Although laboratory decay experiments reveal important aspects of fossilization, applying the results directly to the interpretation of exceptionally preserved fossils may overlook the impact of other key processes that remove or preserve morphological information. Investigations of fossils preserving non-biomineralized tissues suggest that certain structures that are decay resistant (e.g., the notochord) are rarely preserved (even where carbonaceous components survive), and decay-prone structures (e.g., nervous systems) can fossilize, albeit rarely. As we review here, decay resistance is an imperfect indicator of fossilization potential, and a suite of biological and geological processes account for the features preserved in exceptional fossils. © 2017 The Authors. BioEssays Published by WILEY Periodicals, Inc.

  11. Effects of changing nutrient inputs on the ratio of small pelagic fish stock and phytoplankton biomass in the Black Sea

    NASA Astrophysics Data System (ADS)

    Yunev, Oleg A.; Velikova, Violeta; Carstensen, Jacob

    2017-10-01

    Significant increases in nitrogen and phosphorus inputs to the Black Sea in the second half of the 20th century caused eutrophication and drastically decreasing Si:N and Si:P ratios. Combined with climate change, overfishing of top predators and a huge outbreak of the non-indigenous ctenophore Mnemiopsis, the pelagic food web was strongly modified and its efficiency for channeling primary production to higher trophic levels substantially reduced. We used the ratio between small pelagic fish stock and phytoplankton biomass on the Danube shelf and in the open Black Sea to investigate long-term changes in food web functioning. The ratio had 1) highest values for the pre-eutrophication period when diatoms and copepods dominated the pelagic food web ('muscle food chain'), 2) decreased during the eutrophication period with stronger prevalence of autotrophic pico- and nanophytoplankton, bacteria, heterotrophic nanoflagellates, microzooplankton, Noctiluca and jellyfish ('jelly food chain' with increased importance of the microbial loop), 3) lowest values during the ecological crisis (1989-1992), when small pelagic fish stocks collapsed, and 4) increased after 1993, indicating that the ecosystem went out of the crisis and exhibited a trend of recovery. However, in the last period (1993-2008) the ratio remained close to values observed in the middle eutrophication phase, suggesting that the ecosystem was far from fully recovered. Since early 2000s, fluctuating pelagic fish stocks, with a tendency to decreasing fish landing again, have been observed in the Black Sea. Additionally, the quality of food for the small pelagic fish has deteriorated due to warming trends and the legacy of eutrophication, giving support for the 'jelly food chain', exhibiting low energy transfer and prevalence of organisms with high respiration rate and low nutritional value.

  12. Fossil energy program

    NASA Astrophysics Data System (ADS)

    McNeese, L. E.

    1981-12-01

    The progress made during the period from July 1 through September 30 for the Oak Ridge National Laboratory research and development projects in support of the increased utilization of coal and other fossil fuels as sources of clean energy is reported. The following topics are discussed: coal conversion development, chemical research and development, materials technology, fossil energy materials program, liquefaction projects, component development, process analysis, environmental control technology, atmospheric fluidized bed combustion, underground coal gasification, coal preparation and waste utilization.

  13. Deep pelagic food web structure as revealed by in situ feeding observations.

    PubMed

    Choy, C Anela; Haddock, Steven H D; Robison, Bruce H

    2017-12-06

    Food web linkages, or the feeding relationships between species inhabiting a shared ecosystem, are an ecological lens through which ecosystem structure and function can be assessed, and thus are fundamental to informing sustainable resource management. Empirical feeding datasets have traditionally been painstakingly generated from stomach content analysis, direct observations and from biochemical trophic markers (stable isotopes, fatty acids, molecular tools). Each approach carries inherent biases and limitations, as well as advantages. Here, using 27 years (1991-2016) of in situ feeding observations collected by remotely operated vehicles (ROVs), we quantitatively characterize the deep pelagic food web of central California within the California Current, complementing existing studies of diet and trophic interactions with a unique perspective. Seven hundred and forty-three independent feeding events were observed with ROVs from near-surface waters down to depths approaching 4000 m, involving an assemblage of 84 different predators and 82 different prey types, for a total of 242 unique feeding relationships. The greatest diversity of prey was consumed by narcomedusae, followed by physonect siphonophores, ctenophores and cephalopods. We highlight key interactions within the poorly understood 'jelly web', showing the importance of medusae, ctenophores and siphonophores as key predators, whose ecological significance is comparable to large fish and squid species within the central California deep pelagic food web. Gelatinous predators are often thought to comprise relatively inefficient trophic pathways within marine communities, but we build upon previous findings to document their substantial and integral roles in deep pelagic food webs. © 2017 The Authors.

  14. Deep pelagic food web structure as revealed by in situ feeding observations

    PubMed Central

    Haddock, Steven H. D.; Robison, Bruce H.

    2017-01-01

    Food web linkages, or the feeding relationships between species inhabiting a shared ecosystem, are an ecological lens through which ecosystem structure and function can be assessed, and thus are fundamental to informing sustainable resource management. Empirical feeding datasets have traditionally been painstakingly generated from stomach content analysis, direct observations and from biochemical trophic markers (stable isotopes, fatty acids, molecular tools). Each approach carries inherent biases and limitations, as well as advantages. Here, using 27 years (1991–2016) of in situ feeding observations collected by remotely operated vehicles (ROVs), we quantitatively characterize the deep pelagic food web of central California within the California Current, complementing existing studies of diet and trophic interactions with a unique perspective. Seven hundred and forty-three independent feeding events were observed with ROVs from near-surface waters down to depths approaching 4000 m, involving an assemblage of 84 different predators and 82 different prey types, for a total of 242 unique feeding relationships. The greatest diversity of prey was consumed by narcomedusae, followed by physonect siphonophores, ctenophores and cephalopods. We highlight key interactions within the poorly understood ‘jelly web’, showing the importance of medusae, ctenophores and siphonophores as key predators, whose ecological significance is comparable to large fish and squid species within the central California deep pelagic food web. Gelatinous predators are often thought to comprise relatively inefficient trophic pathways within marine communities, but we build upon previous findings to document their substantial and integral roles in deep pelagic food webs. PMID:29212727

  15. The first Loranthaceae fossils from Africa

    PubMed Central

    2018-01-01

    Abstract An ongoing re-investigation of the early Miocene Saldanha Bay (South Africa) palynoflora, using combined light and scanning electron microscopy (single grain method), is revealing several pollen types new to the African fossil record. One of the elements identified is Loranthaceae pollen. These grains represent the first and only fossil record of Loranthaceae in Africa. The fossil pollen grains resemble those produced by the core Lorantheae and are comparable to recent Asian as well as some African taxa/lineages. Molecular and fossil signals indicate that Loranthaceae dispersed into Africa via Asia sometime during the Eocene. The present host range of African Loranthaceae and the composition of the palynoflora suggest that the fossil had a range of potential host taxa to parasitise during the early Miocene in the Saldanha Bay region. PMID:29780299

  16. Fisheries conservation on the high seas: linking conservation physiology and fisheries ecology for the management of large pelagic fishes

    PubMed Central

    Horodysky, Andrij Z.; Cooke, Steven J.; Graves, John E.; Brill, Richard W.

    2016-01-01

    Populations of tunas, billfishes and pelagic sharks are fished at or over capacity in many regions of the world. They are captured by directed commercial and recreational fisheries (the latter of which often promote catch and release) or as incidental catch or bycatch in commercial fisheries. Population assessments of pelagic fishes typically incorporate catch-per-unit-effort time-series data from commercial and recreational fisheries; however, there have been notable changes in target species, areas fished and depth-specific gear deployments over the years that may have affected catchability. Some regional fisheries management organizations take into account the effects of time- and area-specific changes in the behaviours of fish and fishers, as well as fishing gear, to standardize catch-per-unit-effort indices and refine population estimates. However, estimates of changes in stock size over time may be very sensitive to underlying assumptions of the effects of oceanographic conditions and prey distribution on the horizontal and vertical movement patterns and distribution of pelagic fishes. Effective management and successful conservation of pelagic fishes requires a mechanistic understanding of their physiological and behavioural responses to environmental variability, potential for interaction with commercial and recreational fishing gear, and the capture process. The interdisciplinary field of conservation physiology can provide insights into pelagic fish demography and ecology (including environmental relationships and interspecific interactions) by uniting the complementary expertise and skills of fish physiologists and fisheries scientists. The iterative testing by one discipline of hypotheses generated by the other can span the fundamental–applied science continuum, leading to the development of robust insights supporting informed management. The resulting species-specific understanding of physiological abilities and tolerances can help to improve stock

  17. Bayesian phylogenetic estimation of fossil ages.

    PubMed

    Drummond, Alexei J; Stadler, Tanja

    2016-07-19

    Recent advances have allowed for both morphological fossil evidence and molecular sequences to be integrated into a single combined inference of divergence dates under the rule of Bayesian probability. In particular, the fossilized birth-death tree prior and the Lewis-Mk model of discrete morphological evolution allow for the estimation of both divergence times and phylogenetic relationships between fossil and extant taxa. We exploit this statistical framework to investigate the internal consistency of these models by producing phylogenetic estimates of the age of each fossil in turn, within two rich and well-characterized datasets of fossil and extant species (penguins and canids). We find that the estimation accuracy of fossil ages is generally high with credible intervals seldom excluding the true age and median relative error in the two datasets of 5.7% and 13.2%, respectively. The median relative standard error (RSD) was 9.2% and 7.2%, respectively, suggesting good precision, although with some outliers. In fact, in the two datasets we analyse, the phylogenetic estimate of fossil age is on average less than 2 Myr from the mid-point age of the geological strata from which it was excavated. The high level of internal consistency found in our analyses suggests that the Bayesian statistical model employed is an adequate fit for both the geological and morphological data, and provides evidence from real data that the framework used can accurately model the evolution of discrete morphological traits coded from fossil and extant taxa. We anticipate that this approach will have diverse applications beyond divergence time dating, including dating fossils that are temporally unconstrained, testing of the 'morphological clock', and for uncovering potential model misspecification and/or data errors when controversial phylogenetic hypotheses are obtained based on combined divergence dating analyses.This article is part of the themed issue 'Dating species divergences using

  18. Bayesian phylogenetic estimation of fossil ages

    PubMed Central

    Drummond, Alexei J.; Stadler, Tanja

    2016-01-01

    Recent advances have allowed for both morphological fossil evidence and molecular sequences to be integrated into a single combined inference of divergence dates under the rule of Bayesian probability. In particular, the fossilized birth–death tree prior and the Lewis-Mk model of discrete morphological evolution allow for the estimation of both divergence times and phylogenetic relationships between fossil and extant taxa. We exploit this statistical framework to investigate the internal consistency of these models by producing phylogenetic estimates of the age of each fossil in turn, within two rich and well-characterized datasets of fossil and extant species (penguins and canids). We find that the estimation accuracy of fossil ages is generally high with credible intervals seldom excluding the true age and median relative error in the two datasets of 5.7% and 13.2%, respectively. The median relative standard error (RSD) was 9.2% and 7.2%, respectively, suggesting good precision, although with some outliers. In fact, in the two datasets we analyse, the phylogenetic estimate of fossil age is on average less than 2 Myr from the mid-point age of the geological strata from which it was excavated. The high level of internal consistency found in our analyses suggests that the Bayesian statistical model employed is an adequate fit for both the geological and morphological data, and provides evidence from real data that the framework used can accurately model the evolution of discrete morphological traits coded from fossil and extant taxa. We anticipate that this approach will have diverse applications beyond divergence time dating, including dating fossils that are temporally unconstrained, testing of the ‘morphological clock', and for uncovering potential model misspecification and/or data errors when controversial phylogenetic hypotheses are obtained based on combined divergence dating analyses. This article is part of the themed issue ‘Dating species divergences

  19. A novel system for embryo-larval toxicity testing of pelagic fish: Applications for impact assessment of Deepwater Horizon crude oil.

    PubMed

    Stieglitz, John D; Mager, Edward M; Hoenig, Ronald H; Alloy, Matthew; Esbaugh, Andrew J; Bodinier, Charlotte; Benetti, Daniel D; Roberts, Aaron P; Grosell, Martin

    2016-11-01

    Key differences in the developmental process of pelagic fish embryos, in comparison to embryos of standard test fish species, present challenges to obtaining sufficient control survival needed to successfully perform traditional toxicity testing bioassays. Many of these challenges relate to the change in buoyancy, from positive to negative, of pelagic fish embryos that occurs just prior to hatch. A novel exposure system, the pelagic embryo-larval exposure chamber (PELEC), has been developed to conduct successful bioassays on the early life stages (ELSs; embryos/larvae) of pelagic fish. Using this unique recirculating upwelling system, it was possible to significantly improve control survival in pelagic fish ELS bioassays compared to commonly used static exposure methods. Results demonstrate that control performance of mahi-mahi (Coryphaena hippurus) embryos in the PELEC system, measured as percent survival after 96-hrs, significantly outperformed agitated static exposure and static exposure systems. Similar significant improvements in 72-hr control survival were obtained with yellowfin tuna (Thunnus albacares). The PELEC system was subsequently used to test the effects of photo-induced toxicity of crude oil to mahi-mahi ELSs over the course of 96-hrs. Results indicate a greater than 9-fold increase in toxicity of Deepwater Horizon (DWH) crude oil during co-exposure to ambient sunlight compared to filtered ambient sunlight, revealing the importance of including natural sunlight in 96-hr DWH crude oil bioassays as well as the PELEC system's potential application in ecotoxicological assessments. Copyright © 2016 Elsevier Ltd. All rights reserved.

  20. Persistent predator-prey dynamics revealed by mass extinction.

    PubMed

    Sallan, Lauren Cole; Kammer, Thomas W; Ausich, William I; Cook, Lewis A

    2011-05-17

    Predator-prey interactions are thought by many researchers to define both modern ecosystems and past macroevolutionary events. In modern ecosystems, experimental removal or addition of taxa is often used to determine trophic relationships and predator identity. Both characteristics are notoriously difficult to infer in the fossil record, where evidence of predation is usually limited to damage from failed attacks, individual stomach contents, one-sided escalation, or modern analogs. As a result, the role of predation in macroevolution is often dismissed in favor of competition and abiotic factors. Here we show that the end-Devonian Hangenberg event (359 Mya) was a natural experiment in which vertebrate predators were both removed and added to an otherwise stable prey fauna, revealing specific and persistent trophic interactions. Despite apparently favorable environmental conditions, crinoids diversified only after removal of their vertebrate consumers, exhibiting predatory release on a geological time scale. In contrast, later Mississippian (359-318 Mya) camerate crinoids declined precipitously in the face of increasing predation pressure from new durophagous fishes. Camerate failure is linked to the retention of obsolete defenses or "legacy adaptations" that prevented coevolutionary escalation. Our results suggest that major crinoid evolutionary phenomena, including rapid diversification, faunal turnover, and species selection, might be linked to vertebrate predation. Thus, interactions observed in small ecosystems, such as Lotka-Volterra cycles and trophic cascades, could operate at geologic time scales and higher taxonomic ranks. Both trophic knock-on effects and retention of obsolete traits might be common in the aftermath of predator extinction.

  1. Sustainability of Fossil Fuels

    NASA Astrophysics Data System (ADS)

    Lackner, K. S.

    2002-05-01

    For a sustainable world economy, energy is a bottleneck. Energy is at the basis of a modern, technological society, but unlike materials it cannot be recycled. Energy or more precisely "negentropy" (the opposite of entropy) is always consumed. Thus, one either accepts the use of large but finite resources or must stay within the limits imposed by dilute but self-renewing resources like sunlight. The challenge of sustainable energy is exacerbated by likely growth in world energy demand due to increased population and increased wealth. Most of the world still has to undergo the transition to a wealthy, stable society with the near zero population growth that characterizes a modern industrial society. This represents a huge unmet demand. If ten billion people were to consume energy like North Americans do today, world energy demand would be ten times higher. In addition, technological advances while often improving energy efficiency tend to raise energy demand by offering more opportunity for consumption. Energy consumption still increases at close to the 2.3% per year that would lead to a tenfold increase over the course of the next century. Meeting future energy demands while phasing out fossil fuels appears extremely difficult. Instead, the world needs sustainable or nearly sustainable fossil fuels. I propose the following definition of sustainable under which fossil fuels would well qualify: The use of a technology or resource is sustainable if the intended and unintended consequences will not force its abandonment within a reasonable planning horizon. Of course sustainable technologies must not be limited by resource depletion but this is only one of many concerns. Environmental impacts, excessive land use, and other constraints can equally limit the use of a technology and thus render it unsustainable. In the foreseeable future, fossil fuels are not limited by resource depletion. However, environmental concerns based on climate change and other environmental

  2. Heterogeneous distribution of pelagic sediments incoming the Japan Trench possibly controlling slip propagation on shallow plate boundary fault

    NASA Astrophysics Data System (ADS)

    Yamaguchi, A.; Nakamura, Y.; Fukuchi, R.; Kurano, H.; Ikehara, K.; Kanamatsu, T.; Arai, K.; Usami, K.; Ashi, J.

    2017-12-01

    Catastrophic tsunami of the 2011 Tohoku Earthquake was triggered by large coseismic slip reached to the Japan Trench axis (e.g. Fujiwara et al., 2011, Science; Kodaira et al., 2012, Nature Geoscience). Results of the IODP Expedition 343 (JFAST) suggest that small friction of smectite-rich pelagic clay caused slip propagation on shallow plate boundary fault (Ujiie et al., 2013, Science; Kameda et al., 2015, Geology; Moore et al., 2015, Geosphere). On the other hand, JAMSTEC high-resolution seismic profiles show that incoming sediments have large heterogeneities in thicknesses, and two areas of extremely thin sediments on the Pacific Plate (thickness less than 100 m) were found at around 39°N (Nakamura et al., AGU 2017, this session). To reconcile whether the smectite-rich pelagic clay even exists in these areas, we sampled surface sediments during the R/V Shinsei Maru KS-15-3 cruise. Seven piston cores were retrieved from seaward trench slope, horst, graben, and graben edge. Core lithologies are mainly diatomaceous ooze/clay including tephra layers, not resemble to pelagic clays discovered in JFAST. Ages of tephra layers were estimated by correlating mineral assemblages and refractive indices of volcanic glasses to Japanese widespread tephras. Averaged sedimentation rates of seaward trench slope, horst, graben, and graben edge are estimated to be 25-30, 6.5-20, 45, 0.9 cm/kyr, respectively. These sedimentation rates imply that sediments on seaward trench slope and horst have been deposited in the last 160-500 kyr, suggesting that entire pelagic sediments, including smectite-rich pelagic clay, have been removed by some reasons in the last 0.5 million years. Possible reason for such modification of sediment is near-trench igneous activity known as petit-spot volcanism (Hirano et al., 2006, Science). The lack of smectite-rich pelagic clay near 39°N of the Japan Trench is consistent with results of tsunami inversions proposing shallow large coseismic slip propagated

  3. Temporal variation in pelagic food chain length in response to environmental change

    PubMed Central

    Ruiz-Cooley, Rocio I.; Gerrodette, Tim; Fiedler, Paul C.; Chivers, Susan J.; Danil, Kerri; Ballance, Lisa T.

    2017-01-01

    Climate variability alters nitrogen cycling, primary productivity, and dissolved oxygen concentration in marine ecosystems. We examined the role of this variability (as measured by six variables) on food chain length (FCL) in the California Current (CC) by reconstructing a time series of amino acid–specific δ15N values derived from common dolphins, an apex pelagic predator, and using two FCL proxies. Strong declines in FCL were observed after the 1997–1999 El Niño Southern Oscillation (ENSO) event. Bayesian models revealed longer FCLs under intermediate conditions for surface temperature, chlorophyll concentration, multivariate ENSO index, and total plankton volume but not for hypoxic depth and nitrate concentration. Our results challenge the prevalent paradigm that suggested long-term stability in the food web structure in the CC and, instead, reveal that pelagic food webs respond strongly to disturbances associated with ENSO events, local oceanography, and ongoing changes in climate. PMID:29057322

  4. Assessing trace element diffusion models in fossil and sub-fossil bone

    NASA Astrophysics Data System (ADS)

    Suarez, C. A.; Kohn, M. J.

    2012-12-01

    Three different diffusion models have been proposed to explain trace element uptake during fossilization of bone: diffusion-adsorption (DA), diffusion-recrystallization (DR), and double-medium diffusion (DMD). Theoretically, differences in trace element profiles, particularly the rare earth elements (REE) and U, can discriminate among these possibilities. In this study, we tested which model best explains natural samples by analyzing trace element profiles in natural bone using laser-ablation inductively-coupled plasma mass spectrometry (LA-ICP-MS). Fossil bones ranging in age from a few ka to over 100 Ma were analyzed along traverses from the outer cortical edge to the inner marrow cavity margin. Forty major, minor and trace elements were analyzed, notably Ca, P, transition metals, Sr, Ba, REE, U, Th and Pb. Spatial and analytical resolutions were ~10 μm and ~100 ppb respectively. Many specimens show commonly observed exponential decreases in REE from the outer edge and marrow cavity, with relatively homogeneous U distributions. Yet, most significantly, specimens from American Falls (last interglacial) and Duck Point (last glacial maximum) show distinctive U plateaus adjacent to the outer and inner cortical bone margins. Whereas exponential profiles can be produced by different uptake processes, such plateaus are diagnostic of a DR mechanism. Our work is consistent with recent investigation of trace element diffusivities in modern fresh and deproteinated bone. These studies show similar diffusion rates for REE and U, so the profound disparity in U vs. REE profiles in most fossils cannot result solely from differences in volume diffusion within the context of DA and DMD. Rather, as a recrystallization front propagates into bone, the bone appears to encode changing soil water compositions with earlier vs. later compositions reflected in the bone margin vs. interior. Soil water U concentrations apparently remain nearly fixed during fossilization, whereas REE are

  5. Microbiological assessment along the fish production chain of the Norwegian pelagic fisheries sector--Results from a spot sampling programme.

    PubMed

    Svanevik, Cecilie Smith; Roiha, Irja Sunde; Levsen, Arne; Lunestad, Bjørn Tore

    2015-10-01

    Microbes play an important role in the degradation of fish products, thus better knowledge of the microbiological conditions throughout the fish production chain may help to optimise product quality and resource utilisation. This paper presents the results of a ten-year spot sampling programme (2005-2014) of the commercially most important pelagic fish species harvested in Norway. Fish-, surface-, and storage water samples were collected from fishing vessels and processing factories. Totally 1,181 samples were assessed with respect to microbiological quality, hygiene and food safety. We introduce a quality and safety assessment scheme for fresh pelagic fish recommending limits for heterotrophic plate counts (HPC), thermos tolerant coliforms, enterococci and Listeria monocytogenes. According to the scheme, in 25 of 41 samplings, sub-optimal conditions were found with respect to quality, whereas in 21 and 9 samplings, samples were not in compliance concerning hygiene and food safety, respectively. The present study has revealed that the quality of pelagic fish can be optimised by improving the hygiene conditions at some critical points at an early phase of the production chain. Thus, the proposed assessment scheme may provide a useful tool for the industry to optimise quality and maintain consumer safety of pelagic fishery products. Copyright © 2015 The Authors. Published by Elsevier Ltd.. All rights reserved.

  6. FOSSIL2 energy policy model documentation: generic structures of the FOSSIL2 model

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    None

    1980-10-01

    This report discusses the structure, derivations, assumptions, and mathematical formulation of the FOSSIL2 model. Each major facet of the model - supply/demand interactions, industry financing, and production - has been designed to parallel closely the actual cause/effect relationships determining the behavior of the United States energy system. The data base for the FOSSIL2 program is large. When possible, all data were obtained from sources well known to experts in the energy field. Cost and resource estimates are based on DOE data whenever possible. This report presents the FOSSIL2 model at several levels. In Volume I, an overview of the basicmore » structures, assumptions, and behavior of the FOSSIL2 model is presented so that the reader can understand the results of various policy tests. The discussion covers the three major building blocks, or generic structures, used to construct the model: supply/demand balance; finance and capital formation; and energy production. These structures reflect the components and interactions of the major processes within each energy industry that directly affect the dynamics of fuel supply, demand, and price within the energy system as a whole.« less

  7. Selective Preservation of Fossil Ghost Fish

    NASA Astrophysics Data System (ADS)

    Meacham, Amanda

    2016-04-01

    A unique type of fossil fish preservation has been discovered in the Angelo Member (Fossil Lake) of the Green River Formation. The Angelo Member is a predominately evaporative deposit dominated by dolomite, but contains facies of fossiliferous laminated calcimicrite. Fossil fish occurring in two beds conspicuously lack bones. Fish in the lower bed are only preserved as organic material, including skin, pigments, and eyes. Fish in the upper bed have three-dimensional etching where bones once existed but also contain skin, pigments, and eyes. The top third of the upper bed often contains calcite crystals that are pseudomorphs after trona and possibly halite. Preliminary mineralogical analysis and mapping of evaporate facies suggests that this unique preservation may be related to lake geochemical conditions, such as high pH and alkalinity. To our knowledge, this is the first time this type of preservation has been observed and studied. Fossils and sediments within these beds are being studied both vertically and laterally through the one-meter thick sequence containing the fossil fish using XRD, isotopic, SEM, thin section, and total organic carbon analysis. Nine quarries, 0.5-1 meter square, were excavated for both fossils and rock samples along with 17 additional rock sample locations across an approximately 25-kilometer square region. This investigation has the capability of reconstructing the paleoenvironment and lake chemistry of Fossil Lake during the deposition of the "ghost-fish" beds and solving the mystery of the "missing bones" and the unusual process of preservation.

  8. Long-Term Monitoring of Dolphin Biosonar Activity in Deep Pelagic Waters of the Mediterranean Sea.

    PubMed

    Caruso, Francesco; Alonge, Giuseppe; Bellia, Giorgio; De Domenico, Emilio; Grammauta, Rosario; Larosa, Giuseppina; Mazzola, Salvatore; Riccobene, Giorgio; Pavan, Gianni; Papale, Elena; Pellegrino, Carmelo; Pulvirenti, Sara; Sciacca, Virginia; Simeone, Francesco; Speziale, Fabrizio; Viola, Salvatore; Buscaino, Giuseppa

    2017-06-28

    Dolphins emit short ultrasonic pulses (clicks) to acquire information about the surrounding environment, prey and habitat features. We investigated Delphinidae activity over multiple temporal scales through the detection of their echolocation clicks, using long-term Passive Acoustic Monitoring (PAM). The Istituto Nazionale di Fisica Nucleare operates multidisciplinary seafloor observatories in a deep area of the Central Mediterranean Sea. The Ocean noise Detection Experiment collected data offshore the Gulf of Catania from January 2005 to November 2006, allowing the study of temporal patterns of dolphin activity in this deep pelagic zone for the first time. Nearly 5,500 five-minute recordings acquired over two years were examined using spectrogram analysis and through development and testing of an automatic detection algorithm. Echolocation activity of dolphins was mostly confined to nighttime and crepuscular hours, in contrast with communicative signals (whistles). Seasonal variation, with a peak number of clicks in August, was also evident, but no effect of lunar cycle was observed. Temporal trends in echolocation corresponded to environmental and trophic variability known in the deep pelagic waters of the Ionian Sea. Long-term PAM and the continued development of automatic analysis techniques are essential to advancing the study of pelagic marine mammal distribution and behaviour patterns.

  9. Title: Biogenic Magnetite Prevails in Oxic Pelagic Red Clay Core in the South Pacific Gyre

    NASA Astrophysics Data System (ADS)

    Shimono, T.; Yamazaki, T.

    2012-12-01

    Magnetotactic bacteria have been observed in wide variety of environments, including soils, freshwater lakes, and marine sediments, since Blakemore (1975) first described in 1975. Magnetotactic bacteria, which most commonly live within the oxic-anoxic transition zone (OATZ) of aquatic environments, produce intracellular crystals of magnetic minerals, specifically magnetite or greigite. It is considered that the magnetite/greigite crystals facilitate the bacteria's search for optimal conditions within the sharp chemical gradients of the OATZ. Petermann and Bleil (1993) reported living magnetotactic bacteria in pelagic and hemipelagic sediments near OATZ in the eastern South Atlantic at water depths to about 3,000 m, but they couldn't find actively swimming magnetotactic bacteria in sediments of deeper water depths. The South Pacific Gyre (SPG) is far from continents and the lowest productivity region on Earth. IODP site U1365 (water depth ~5,700 m) cored pelagic red clay of 75.5 m thick above ~100 Ma basement (except for the chart layer from ~42 to 63.5 m) in the western edge of the SPG. The core mainly consists of iron rich clay. The color is dark reddish and/or dark brown throughout the core. We conducted a paleomagnetic and environmental rock magnetic study of the pelagic clay core. The magnetostratigraphy revealed the top 5 m sediments cover the last 5 My, and sedimentation rate decreases downward from 1.7 to 0.6 m/m.y. Geochemical measurements of pore water indicate that dissolved oxygen was present throughout the core (>50 μM). Thus oxygen penetrates through the entire sediment column to the sediment/basalt interface, and there is no OATZ. Magnetic mineral assemblage of this core is dominated by biogenic magnetite despite no OATZ. First-order reversal curve (FORC) diagrams of all specimens have a narrow central ridge along the Hc axis with very small vertical spread. This indicates very weak magnetostatic interaction (Roberts et al., 2000), and is the

  10. Effects of in utero retinoic acid exposure on mouse pelage hair follicle development.

    PubMed

    García-Fernández, Rosa A; Pérez-Martínez, Claudia; Escudero-Diez, Alfredo; García-Iglesias, Maria J

    2002-06-01

    We investigated in vivo the histological and immunohistochemical responses of mouse hair pelage follicle morphogenesis to prenatal exposure to a potentially nonteratogenic dose of all-trans-retinoic acid (RA), as a basis studying the preventive effect of RA on adult mouse skin carcinogenesis. In pregnant mice, a single oral dose of RA at 30 mg kg-1 body weight given on day 11.5 of gestation caused no RA-induced changes in the morphology or temporal expression patterns of keratins during pelage hair follicle morphogenesis. The only differential effect of RA was a statistically significant increase in the number of BrdU-positive nuclei in hair bulbs from RA exposed fetuses compared with nonexposed mice. The absence of adverse RA effects suggests that this experimental design may represent a valuable protocol for use in studies on the in vivo effects of this retinoid on different skin diseases.

  11. Pelagic photoferrotrophy and iron cycling in a modern ferruginous basin.

    PubMed

    Llirós, Marc; García-Armisen, Tamara; Darchambeau, François; Morana, Cédric; Triadó-Margarit, Xavier; Inceoğlu, Özgül; Borrego, Carles M; Bouillon, Steven; Servais, Pierre; Borges, Alberto V; Descy, Jean-Pierre; Canfield, Don E; Crowe, Sean A

    2015-09-08

    Iron-rich (ferruginous) ocean chemistry prevailed throughout most of Earth's early history. Before the evolution and proliferation of oxygenic photosynthesis, biological production in the ferruginous oceans was likely driven by photoferrotrophic bacteria that oxidize ferrous iron {Fe(II)} to harness energy from sunlight, and fix inorganic carbon into biomass. Photoferrotrophs may thus have fuelled Earth's early biosphere providing energy to drive microbial growth and evolution over billions of years. Yet, photoferrotrophic activity has remained largely elusive on the modern Earth, leaving models for early biological production untested and imperative ecological context for the evolution of life missing. Here, we show that an active community of pelagic photoferrotrophs comprises up to 30% of the total microbial community in illuminated ferruginous waters of Kabuno Bay (KB), East Africa (DR Congo). These photoferrotrophs produce oxidized iron {Fe(III)} and biomass, and support a diverse pelagic microbial community including heterotrophic Fe(III)-reducers, sulfate reducers, fermenters and methanogens. At modest light levels, rates of photoferrotrophy in KB exceed those predicted for early Earth primary production, and are sufficient to generate Earth's largest sedimentary iron ore deposits. Fe cycling, however, is efficient, and complex microbial community interactions likely regulate Fe(III) and organic matter export from the photic zone.

  12. The Quaternary fossil-pollen record and global change

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Grimm, E.C.

    Fossil pollen provide one of the most valuable records of vegetation and climate change during the recent geological past. Advantages of the fossil-pollen record are that deposits containing fossil pollen are widespread, especially in areas having natural lakes, that fossil pollen occurs in continuous stratigraphic sequences spanning millennia, and that fossil pollen occurs in quantitative assemblages permitting a multivariate approach for reconstructing past vegetation and climates. Because of stratigraphic continuity, fossil pollen records climate cycles on a wide range of scales, from annual to the 100 ka Milankovitch cycles. Receiving particular emphasis recently are decadal to century scale changes, possiblemore » from the sediments of varved lakes, and late Pleistocene events on a 5--10 ka scale possibly correlating with the Heinrich events in the North Atlantic marine record or the Dansgaard-Oeschger events in the Greenland ice-core record. Researchers have long reconstructed vegetation and climate by qualitative interpretation of the fossil-pollen record. Recently quantitative interpretation has developed with the aid of large fossil-pollen databases and sophisticated numerical models. In addition, fossil pollen are important climate proxy data for validating General Circulation Models, which are used for predicting the possible magnitude future climate change. Fossil-pollen data also contribute to an understanding of ecological issues associated with global climate change, including questions of how and how rapidly ecosystems might respond to abrupt climate change.« less

  13. Fossil and non-fossil source contributions to atmospheric carbonaceous aerosols during extreme spring grassland fires in Eastern Europe

    NASA Astrophysics Data System (ADS)

    Ulevicius, V.; Byčenkienė, S.; Bozzetti, C.; Vlachou, A.; Plauškaitė, K.; Mordas, G.; Dudoitis, V.; Abbaszade, G.; Remeikis, V.; Garbaras, A.; Masalaite, A.; Blees, J.; Fröhlich, R.; Dällenbach, K. R.; Canonaco, F.; Slowik, J. G.; Dommen, J.; Zimmermann, R.; Schnelle-Kreis, J.; Salazar, G. A.; Agrios, K.; Szidat, S.; El Haddad, I.; Prévôt, A. S. H.

    2015-09-01

    In early spring the Baltic region is frequently affected by high pollution events due to biomass burning in that area. Here we present a comprehensive study to investigate the impact of biomass/grass burning (BB) on the evolution and composition of aerosol in Preila, Lithuania, during springtime open fires. Non-refractory submicron particulate matter (NR-PM1) was measured by an Aerodyne aerosol chemical speciation monitor (ACSM) and a source apportionment with the multilinear engine (ME-2) running the positive matrix factorization (PMF) model was applied to the organic aerosol fraction to investigate the impact of biomass/grass burning. Satellite observations over regions of biomass burning activity supported the results and identification of air mass transport to the area of investigation. Sharp increases in biomass burning tracers, such as levoglucosan up to 683 ng m-3 and black carbon (BC) up to 17 μg m-3 were observed during this period. A further separation between fossil and non-fossil primary and secondary contributions was obtained by coupling ACSM PMF results and radiocarbon (14C) measurements of the elemental (EC) and organic (OC) carbon fractions. Non-fossil organic carbon (OCnf) was the dominant fraction of PM1, with the primary (POCnf) and secondary (SOCnf) fractions contributing 26-44 % and 13-23 % to the TC, respectively. 5-8 % of the TC had a primary fossil origin (POCf), whereas the contribution of fossil secondary organic carbon (SOCf) was 4-13 %. Non-fossil EC (ECnf) and fossil EC (ECf) ranged from 13-24 % and 7-12 %, respectively. Isotope ratio of stable carbon and nitrogen isotopes were used to distinguish aerosol particles associated with solid and liquid fossil fuel burning.

  14. Fossil and non-fossil source contributions to atmospheric carbonaceous aerosols during extreme spring grassland fires in Eastern Europe

    NASA Astrophysics Data System (ADS)

    Ulevicius, Vidmantas; Byčenkienė, Steigvilė; Bozzetti, Carlo; Vlachou, Athanasia; Plauškaitė, Kristina; Mordas, Genrik; Dudoitis, Vadimas; Abbaszade, Gülcin; Remeikis, Vidmantas; Garbaras, Andrius; Masalaite, Agne; Blees, Jan; Fröhlich, Roman; Dällenbach, Kaspar R.; Canonaco, Francesco; Slowik, Jay G.; Dommen, Josef; Zimmermann, Ralf; Schnelle-Kreis, Jürgen; Salazar, Gary A.; Agrios, Konstantinos; Szidat, Sönke; El Haddad, Imad; Prévôt, André S. H.

    2016-05-01

    In early spring the Baltic region is frequently affected by high-pollution events due to biomass burning in that area. Here we present a comprehensive study to investigate the impact of biomass/grass burning (BB) on the evolution and composition of aerosol in Preila, Lithuania, during springtime open fires. Non-refractory submicron particulate matter (NR-PM1) was measured by an Aerodyne aerosol chemical speciation monitor (ACSM) and a source apportionment with the multilinear engine (ME-2) running the positive matrix factorization (PMF) model was applied to the organic aerosol fraction to investigate the impact of biomass/grass burning. Satellite observations over regions of biomass burning activity supported the results and identification of air mass transport to the area of investigation. Sharp increases in biomass burning tracers, such as levoglucosan up to 683 ng m-3 and black carbon (BC) up to 17 µg m-3 were observed during this period. A further separation between fossil and non-fossil primary and secondary contributions was obtained by coupling ACSM PMF results and radiocarbon (14C) measurements of the elemental (EC) and organic (OC) carbon fractions. Non-fossil organic carbon (OCnf) was the dominant fraction of PM1, with the primary (POCnf) and secondary (SOCnf) fractions contributing 26-44 % and 13-23 % to the total carbon (TC), respectively. 5-8 % of the TC had a primary fossil origin (POCf), whereas the contribution of fossil secondary organic carbon (SOCf) was 4-13 %. Non-fossil EC (ECnf) and fossil EC (ECf) ranged from 13-24 and 7-13 %, respectively. Isotope ratios of stable carbon and nitrogen isotopes were used to distinguish aerosol particles associated with solid and liquid fossil fuel burning.

  15. 76 FR 40836 - Fisheries Off West Coast States; Coastal Pelagic Species Fisheries; Closure

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-07-12

    .... 0912281446-0111-02] RIN 0648-XA554 Fisheries Off West Coast States; Coastal Pelagic Species Fisheries; Closure AGENCY: National Marine Fisheries Service (NMFS), National Oceanic and Atmospheric Administration..., Pacific sardine may be harvested only as part of the live bait fishery or incidental to other fisheries...

  16. 75 FR 33733 - Fisheries Off West Coast States; Coastal Pelagic Species Fisheries; Closure

    Federal Register 2010, 2011, 2012, 2013, 2014

    2010-06-15

    .... 0912281446-0111-02 RIN 0648-XW90 Fisheries Off West Coast States; Coastal Pelagic Species Fisheries; Closure AGENCY: National Marine Fisheries Service (NMFS), National Oceanic and Atmospheric Administration (NOAA... sardine can only be harvested as part of the live bait fishery or incidental to other fisheries; the...

  17. 77 FR 50952 - Fisheries Off West Coast States; Coastal Pelagic Species Fisheries; Closure

    Federal Register 2010, 2011, 2012, 2013, 2014

    2012-08-23

    .... 120312182-2239-02] RIN 0648-XC166 Fisheries Off West Coast States; Coastal Pelagic Species Fisheries; Closure AGENCY: National Marine Fisheries Service (NMFS), National Oceanic and Atmospheric Administration..., Pacific sardine may be harvested only as part of the live bait fishery or incidental to other fisheries...

  18. 75 FR 42610 - Fisheries Off West Coast States; Coastal Pelagic Species Fisheries; Closure

    Federal Register 2010, 2011, 2012, 2013, 2014

    2010-07-22

    .... 0912281446-0111-02] RIN 0648-XX54 Fisheries Off West Coast States; Coastal Pelagic Species Fisheries; Closure AGENCY: National Marine Fisheries Service (NMFS), National Oceanic and Atmospheric Administration (NOAA..., Pacific sardine can only be harvested as part of the live bait fishery or incidental to other fisheries...

  19. 75 FR 59156 - Fisheries Off West Coast States; Coastal Pelagic Species Fisheries; Closure

    Federal Register 2010, 2011, 2012, 2013, 2014

    2010-09-27

    .... 0912281446-0111-02] RIN 0648-XY79 Fisheries Off West Coast States; Coastal Pelagic Species Fisheries; Closure AGENCY: National Marine Fisheries Service (NMFS), National Oceanic and Atmospheric Administration (NOAA..., Pacific sardine can only be harvested as part of the live bait fishery or incidental to other fisheries...

  20. 78 FR 51097 - Fisheries Off West Coast States; Coastal Pelagic Species Fisheries; Closure

    Federal Register 2010, 2011, 2012, 2013, 2014

    2013-08-20

    .... 121210694-3514-02] RIN 0648-XC783 Fisheries Off West Coast States; Coastal Pelagic Species Fisheries; Closure AGENCY: National Marine Fisheries Service (NMFS), National Oceanic and Atmospheric Administration..., Pacific sardine may be harvested only as part of the live bait fishery or incidental to other fisheries...

  1. 76 FR 58720 - Fisheries Off West Coast States; Coastal Pelagic Species Fisheries; Closure

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-09-22

    .... 0912281446-0111-02] RIN 0648-XA709 Fisheries Off West Coast States; Coastal Pelagic Species Fisheries; Closure AGENCY: National Marine Fisheries Service (NMFS), National Oceanic and Atmospheric Administration..., 2012, Pacific sardine may be harvested only as part of the live bait fishery or incidental to other...

  2. Fossil Groups as Cosmological Labs

    NASA Astrophysics Data System (ADS)

    D'Onghia, Elena

    Optical and X-ray measurements of fossil groups (FGs) suggest that they are old and relaxed systems. If FGs are assembled at higher redshift, there is enough time for intermediate-luminosity galaxies to merge, resulting in the formation of the brightest group galaxy (BGG). We carry out the first, systematic study of a large sample of FGs, the "FOssil Group Origins'' (FOGO) based on an International Time Project at the Roque de los Muchachos Observatory. For ten FOGO FGs we have been awarded time at SUZAKU Telescope to measure the temperature of the hot intragroup gas (IGM). For these systems we plan to evaluate and correlate their X-ray luminosity and X-ray temperature, Lx-Tx, optical luminosity and X-ray temperature, Lopt-Tx, and group velocity dispersion with their X-ray temperature, sigma V-Tx, as compared to the non fossil systems. By combining these observations with state-of-art cosmological hydrodynamical simulations we will open a new window into the study of the IGM and the nature of fossil systems. Our proposed work will be of direct relevance for the understanding and interpretation of data from several NASA science missions. Specifically, the scaling relations obtained from these data combined with our predictions obtained using state-of-the-art hydrodynamical simulation numerical adopting a new hydrodynamical scheme will motivate new proposal on CHANDRA X-ray telescope for fossil groups and clusters. We will additionally create a public Online Planetarium Show. This will be an educational site, containing an interactive program called: "A Voyage to our Universe''. In the show we will provide observed images of fossil groups and similar images and movies obtained from the numerical simulations showing their evolution. The online planetarium show will be a useful reference and an interactive educational tool for both students and the public.

  3. Diagenesis of lower Cretaceous pelagic carbonates, North Atlantic: Paleoceanographic signals obscured

    USGS Publications Warehouse

    Frank, T.D.; Arthur, M.A.; Dean, W.E.

    1999-01-01

    The stable isotope and minor element geochemistry of Neocomian (Lower Cretaceous) pelagic carbonates of the North Atlantic Basin (Deep Sea Drilling Project Sites 105, 367, 387, 391, and 603) was examined to develop a diagenetic model for pelagic limestones. In particular, we hoped to test the fidelity of whole-rock geochemical records as paleoceanographic indicators for pelagic deposits of pre-Aptian age, in which individual microfossils are not available for analysis. Data indicate that in addition to depth of burial, rhythmic variations in primary carbonate content have strongly controlled diagenetic patterns and associated geochemical signatures in these Neocomian sequences. Samples become increasingly depleted in Sr and 18O with increasing CaCO3 content. Within individual sedimentary sections, substantial decreases in Sr/Ca ratios and ??18O values are evident over a range of 4 to 98% CaCO3. However, even over a relatively narrow range of 50 to 98% CaCO3 a 2.5%c variation in ??18O values and a change of a factor of 1.7 in Sr/Ca ratios are observed. Carbon isotope compositions do not vary as extensively with CaCO3 content, but carbonate-rich intervals tend to be relatively depleted in 13C. Petrographic analysis reveals that these geochemical patterns are related to the transfer of CaCO3 from carbonate-poor intervals (calcareous shales and marlstones) to adjacent carbonate-rich intervals (limestones) during burial compaction and pressure solution. This process results in the addition of diagenetic cement to carbonate-rich intervals to produce a bulk composition that is relatively depleted in Sr and 18O and, at the same time, enables the retention of more-or-less primary carbonate that is relatively enriched in Sr and 18O in adjacent carbonate-poor intervals. Thus, although cyclic variations in CaCO3 content are primary in the Neocomian sequences examined, measured variations in Sr/Ca ratios and ??18O values are not and, as such, do not provide reliable proxies for

  4. Relatively Recent Evolution of Pelage Coloration in Colobinae: Phylogeny and Phylogeography of Three Closely Related Langur Species

    PubMed Central

    Nadler, Tilo; Liu, Guangjian; Sun, Tao; Huang, Chengming; Zhou, Qihai; Zhou, Jiang; Que, Tengcheng; Wang, Ziming; Roos, Christian; Li, Ming

    2013-01-01

    To understand the evolutionary processes leading to the diversity of Asian colobines, we report here on a phylogenetic, phylogeographical and population genetic analysis of three closely related langurs, Trachypithecus francoisi, T. poliocephalus and T. leucocephalus, which are all characterized by different pelage coloration predominantly on the head and shoulders. Therefore, we sequenced a 395 bp long fragment of the mitochondrial control region from 178 T. francoisi, 54 T. leucocephalus and 19 T. poliocephalus individuals, representing all extant populations of these three species. We found 29 haplotypes in T. francoisi, 12 haplotypes in T. leucocephalus and three haplotypes in T. poliocephalus. T. leucocephalus and T. poliocephalus form monophyletic clades, which are both nested within T. francoisi, and diverged from T. francoisi recently, 0.46-0.27 (T. leucocephalus) and 0.50-0.25 million years ago (T. poliocephalus). Thus, T. francoisi appears as a polyphyletic group, while T. leucocephalus and T. poliocephalus are most likely independent descendents of T. francoisi that are both physically separated from T. francoisi populations by rivers, open sea or larger habitat gaps. Since T. francoisi populations show no variability in pelage coloration, pelage coloration in T. leucocephalus and T. poliocephalus is most likely the result of new genetic mutations after the split from T. francoisi and not of the fixation of different characters derived from an ancestral polymorphism. This case study highlights that morphological changes for example in pelage coloration can occur in isolated populations in relatively short time periods and it provides a solid basis for studies in related species. Nevertheless, to fully understand the evolutionary history of these three langur species, nuclear loci should be investigated as well. PMID:23613895

  5. Travels with the Fossil Hunters

    NASA Astrophysics Data System (ADS)

    Whybrow, Peter J.

    2000-04-01

    Whether dodging bullets in West Africa, or rabid dogs in Pakistan, surviving yak-butter tea in Tibet, or eating raw fish in China, the life of a globe-trotting fossil hunter is often hazardous and always filled with surprises. Travels with the Fossil Hunters lets readers share the wonder, joys of discovery, and excitement of these intrepid scientists. Packed with more than 100 beautiful, full-color photographs, the volume takes readers on twelve expeditions to remote parts of the world in search of diverse fossil remains, from those of dinosaurs to human ancestors. Each expedition by paleontologists from London's Natural History Museum reveals the problems and challenges of working in extreme conditions, from the deserts of the Sahara and Yemen to the frozen wastes of Antarctica, from the mountains of India to the forests of Latvia. Along the way they also describe the paleontology and geology of the countries they visit and the scientific reasons for their expeditions. With a foreword from Sir David Attenborough and an introduction from Richard Fortey, this fascinating book will appeal to amateur and professional fossil hunters alike and to readers interested in accounts of exotic locales. Peter Whybrow is a research scientist at the Natural History Museum, London. His research interests include Arabian Miocene vertebrates, paleoclimates, paleogeography, and biotic diversity. He is senior editor with A. Hill of Fossil Vertebrates of Arabia (Yale University Press, New Haven, 1999).

  6. Traversing the mountaintop: world fossil fuel production to 2050

    PubMed Central

    Nehring, Richard

    2009-01-01

    During the past century, fossil fuels—petroleum liquids, natural gas and coal—were the dominant source of world energy production. From 1950 to 2005, fossil fuels provided 85–93% of all energy production. All fossil fuels grew substantially during this period, their combined growth exceeding the increase in world population. This growth, however, was irregular, providing for rapidly growing per capita production from 1950 to 1980, stable per capita production from 1980 to 2000 and rising per capita production again after 2000. During the past half century, growth in fossil fuel production was essentially limited by energy demand. During the next half century, fossil fuel production will be limited primarily by the amount and characteristics of remaining fossil fuel resources. Three possible scenarios—low, medium and high—are developed for the production of each of the fossil fuels to 2050. These scenarios differ primarily by the amount of ultimate resources estimated for each fossil fuel. Total fossil fuel production will continue to grow, but only slowly for the next 15–30 years. The subsequent peak plateau will last for 10–15 years. These production peaks are robust; none of the fossil fuels, even with highly optimistic resource estimates, is projected to keep growing beyond 2050. World fossil fuel production per capita will thus begin an irreversible decline between 2020 and 2030. PMID:19770156

  7. Traversing the mountaintop: world fossil fuel production to 2050.

    PubMed

    Nehring, Richard

    2009-10-27

    During the past century, fossil fuels--petroleum liquids, natural gas and coal--were the dominant source of world energy production. From 1950 to 2005, fossil fuels provided 85-93% of all energy production. All fossil fuels grew substantially during this period, their combined growth exceeding the increase in world population. This growth, however, was irregular, providing for rapidly growing per capita production from 1950 to 1980, stable per capita production from 1980 to 2000 and rising per capita production again after 2000. During the past half century, growth in fossil fuel production was essentially limited by energy demand. During the next half century, fossil fuel production will be limited primarily by the amount and characteristics of remaining fossil fuel resources. Three possible scenarios--low, medium and high--are developed for the production of each of the fossil fuels to 2050. These scenarios differ primarily by the amount of ultimate resources estimated for each fossil fuel. Total fossil fuel production will continue to grow, but only slowly for the next 15-30 years. The subsequent peak plateau will last for 10-15 years. These production peaks are robust; none of the fossil fuels, even with highly optimistic resource estimates, is projected to keep growing beyond 2050. World fossil fuel production per capita will thus begin an irreversible decline between 2020 and 2030.

  8. 76 FR 40674 - Fisheries in the Western Pacific; Pelagic Fisheries; Purse Seine Prohibited Areas Around American...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-07-11

    ... Atmospheric Administration (NOAA), Commerce. ACTION: Disapproval of fishery ecosystem plan amendment and... Fishery Ecosystem Plan for Pelagic Fisheries of the Western Pacific (FEP) that would have prohibited purse...

  9. ZOOPLANKTON LIFE CYCLES: DIRECT DOCUMENTATION OF PELAGIC BIRTHS AND DEATHS RELATIVE TO DIAPAUSING EGG PRODUCTION

    EPA Science Inventory

    Full-season demographics (pelagic births and deaths, diapause egg production) provide valuable insights into species persistence and dispersal success. The spiny cladoceran (Bythotrephes) possesses morphological adaptations (spines and thick-walled diapause eggs) that lessen impa...

  10. 76 FR 11969 - Fisheries Off West Coast States; Coastal Pelagic Species Fisheries; Annual Specifications

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-03-04

    ... DEPARTMENT OF COMMERCE National Oceanic and Atmospheric Administration 50 CFR Part 660 [Docket No. 110111018-1095-02] RIN 0648-XA109 Fisheries Off West Coast States; Coastal Pelagic Species Fisheries; Annual Specifications AGENCY: National Marine Fisheries Service (NMFS), National Oceanic and Atmospheric Administration...

  11. Fossil Leaves and Fossil Leaf n-Alkanes: Reconstructing the First Closed Canopied Rainforests

    NASA Astrophysics Data System (ADS)

    Graham, H. V.; Freeman, K. H.

    2013-12-01

    Although the age and location is disputed, the rise of the first closed-canopy forest is likely linked with the expansion of angiosperms in the late Cretacous or early Cenozoic. The carbon isotope 'canopy effect' reflects the extent of canopy closure, and is well documented in δ13C values of the leaves and leaf lipids in modern forests. To test the extent of canopy closure among the oldest documented angiosperm tropical forests, we analyzed isotopic characteristics of leaf fossils and leaf waxes from the Guaduas and Cerrejón Formations. The Guaduas Fm. (Maastrichtian) contains some of the earliest angiosperm fossils in the Neotropics, and both leaf morphology and pollen records at this site suggest an open-canopy structure. The Cerrejón Fm. (Paleocene) contains what are believed to be the first recorded fossil leaves from a closed-canopy forest. We analyzed the bulk carbon isotope content (δ13Cleaf) of 199 fossil leaves, as well as the n-alkane concentration and chain-length distribution, and δ13C of alkanes (δ13Clipid) of 73 fossil leaves and adjacent sediment samples. Fossil leaves are dominated by eudicots and include ten modern plant families (Apocynaceae, Bombaceae, Euphorbaceae, Fabaceae, Lauraceae, Malvaceae, Meliaceae, Menispermaceae, Moraceae, Sapotaceae). We interpreted extent of canopy coverage based on the range of δ13Cleaf values. The narrow range of δ13C values in leaves from the Guaduas Fm (2.7‰) is consistent with an open canopy. A significantly wider range in values (6.3‰) suggests a closed-canopy signature for site 0315 of the Cerrejón Fm,. In contrast, at Site 0318, a lacustrine deposit, leaves had a narrow range (3.3‰) in δ13C values, and this is not consistent with a closed-canopy, but is consistent with leaf assemblages from a forest edge. Leaves that accumulate in lake sediments tend to be biased toward plants living at the lake edge, which do not experience closed-canopy conditions, and do not express the isotopic

  12. Progress of fossil fuel science

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Demirbas, M.F.

    2007-07-01

    Coal is the most abundant and widely distributed fossil fuel. More than 45% of the world's electricity is generated from coal, and it is the major fuel for generating electricity worldwide. The known coal reserves in the world are enough for more than 215 years of consumption, while the known oil reserves are only about 39 times of the world's consumption and the known natural gas reserves are about 63 times of the world's consumption level in 1998. In recent years, there have been effective scientific investigations on Turkish fossil fuels, which are considerable focused on coal resources. Coal ismore » a major fossil fuel source for Turkey. Turkish coal consumption has been stable over the past decade and currently accounts for about 24% of the country's total energy consumption. Lignite coal has had the biggest share in total fossil fuel production, at 43%, in Turkey. Turkish researchers may investigate ten broad pathways of coal species upgrading, such as desulfurization and oxydesulfurization, pyrolysis and hydropyrolysis, liquefaction and hydroliquefaction, extraction and supercritical fluid extraction, gasification, oxidation, briquetting, flotation, and structure identification.« less

  13. 76 FR 73517 - Fisheries in the Eastern Pacific Ocean; Pelagic Fisheries; Vessel Identification Requirements

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-11-29

    ... [Docket No. 110218143-1606-02] RIN 0648-BA49 Fisheries in the Eastern Pacific Ocean; Pelagic Fisheries... Conservation and Management of Highly Migratory Fish Stocks in the Western and Central Pacific Ocean... Western and Central Pacific Fisheries Commission (WCPFC). DATES: These regulations become effective on...

  14. Macroevolutionary developmental biology: Embryos, fossils, and phylogenies.

    PubMed

    Organ, Chris L; Cooper, Lisa Noelle; Hieronymus, Tobin L

    2015-10-01

    The field of evolutionary developmental biology is broadly focused on identifying the genetic and developmental mechanisms underlying morphological diversity. Connecting the genotype with the phenotype means that evo-devo research often considers a wide range of evidence, from genetics and morphology to fossils. In this commentary, we provide an overview and framework for integrating fossil ontogenetic data with developmental data using phylogenetic comparative methods to test macroevolutionary hypotheses. We survey the vertebrate fossil record of preserved embryos and discuss how phylogenetic comparative methods can integrate data from developmental genetics and paleontology. Fossil embryos provide limited, yet critical, developmental data from deep time. They help constrain when developmental innovations first appeared during the history of life and also reveal the order in which related morphologies evolved. Phylogenetic comparative methods provide a powerful statistical approach that allows evo-devo researchers to infer the presence of nonpreserved developmental traits in fossil species and to detect discordant evolutionary patterns and processes across levels of biological organization. © 2015 Wiley Periodicals, Inc.

  15. The argonaut shell: gas-mediated buoyancy control in a pelagic octopus

    PubMed Central

    Finn, Julian K.; Norman, Mark D.

    2010-01-01

    Argonauts (Cephalopoda: Argonautidae) are a group of rarely encountered open-ocean pelagic octopuses with benthic ancestry. Female argonauts inhabit a brittle ‘paper nautilus’ shell, the role of which has puzzled naturalists for millennia. The primary role attributed to the shell has been as a receptacle for egg deposition and brooding. Our observations of wild argonauts have revealed that the thin calcareous shell also functions as a hydrostatic structure, employed by the female argonaut to precisely control buoyancy at varying depths. Female argonauts use the shell to ‘gulp’ a measured volume of air at the sea surface, seal off the captured gas using flanged arms and forcefully dive to a depth where the compressed gas buoyancy counteracts body weight. This process allows the female argonaut to attain neutral buoyancy at depth and potentially adjust buoyancy to counter the increased (and significant) weight of eggs during reproductive periods. Evolution of this air-capture strategy enables this negatively buoyant octopus to survive free of the sea floor. This major shift in life mode from benthic to pelagic shows strong evolutionary parallels with the origins of all cephalopods, which attained gas-mediated buoyancy via the closed-chambered shells of the true nautiluses and their relatives. PMID:20484241

  16. The argonaut shell: gas-mediated buoyancy control in a pelagic octopus.

    PubMed

    Finn, Julian K; Norman, Mark D

    2010-10-07

    Argonauts (Cephalopoda: Argonautidae) are a group of rarely encountered open-ocean pelagic octopuses with benthic ancestry. Female argonauts inhabit a brittle 'paper nautilus' shell, the role of which has puzzled naturalists for millennia. The primary role attributed to the shell has been as a receptacle for egg deposition and brooding. Our observations of wild argonauts have revealed that the thin calcareous shell also functions as a hydrostatic structure, employed by the female argonaut to precisely control buoyancy at varying depths. Female argonauts use the shell to 'gulp' a measured volume of air at the sea surface, seal off the captured gas using flanged arms and forcefully dive to a depth where the compressed gas buoyancy counteracts body weight. This process allows the female argonaut to attain neutral buoyancy at depth and potentially adjust buoyancy to counter the increased (and significant) weight of eggs during reproductive periods. Evolution of this air-capture strategy enables this negatively buoyant octopus to survive free of the sea floor. This major shift in life mode from benthic to pelagic shows strong evolutionary parallels with the origins of all cephalopods, which attained gas-mediated buoyancy via the closed-chambered shells of the true nautiluses and their relatives.

  17. Diagenesis of echinoderm skeletons: Constraints on paleoseawater Mg/Ca reconstructions

    NASA Astrophysics Data System (ADS)

    Gorzelak, Przemysław; Krzykawski, Tomasz; Stolarski, Jarosław

    2016-09-01

    One of the most profound environmental changes thought to be reflected in chemical composition of numerous geological archives is Mg/Ca ratio of the seawater, which has varied dramatically throughout the Phanerozoic. Echinoderms that today typically form high magnesium calcite skeletons are increasingly being utilized as a proxy for interpreting secular changes in seawater chemistry. However, accurate characterization of the diagenetic changes of their metastable high magnesium calcite skeletons is a prerequisite for assessing their original, major-element geochemical composition. Here we expand the existing models of diagenesis of echinoderm skeleton by integration of various analytical methods that up to now rarely have been used to assess the diagenetic changes of fossil echinoderms. We validated the preservation of a suite of differently preserved echinoderm ossicles, mostly crinoids, ranging in age from the Cambrian through Recent. In 13 of 99 fossil echinoderm ossicles we found well-preserved porous microstructure (stereom), non-luminescent behaviour or blotchy dark color in cathodoluminescence, and distinct nanostructural features (layered and nanocomposite structure). Moreover, in representatives of such preserved samples, distribution of sulphates associated with organic matter is identical to those in Recent echinoderms. Only such ossicles, despite of local micrometer-scale diagenetic changes, were herein considered well-preserved, retaining their original major-element skeletal composition. By contrast, majority of samples show transformation to the stable low magnesium calcite that leads to obliteration of the primary geochemical and micro/nanostructural features and is accompanied with increase in cathodoluminescence emission intensity. Using only well-preserved fossil echinoderm samples, we found purely random variation in Mg/Ca in echinoderm skeletons through the observed time series; any periodicities in echinoderm skeletal Mg/Ca ratio which might

  18. Combining phylogenomics and fossils in higher-level squamate reptile phylogeny: molecular data change the placement of fossil taxa.

    PubMed

    Wiens, John J; Kuczynski, Caitlin A; Townsend, Ted; Reeder, Tod W; Mulcahy, Daniel G; Sites, Jack W

    2010-12-01

    Molecular data offer great potential to resolve the phylogeny of living taxa but can molecular data improve our understanding of relationships of fossil taxa? Simulations suggest that this is possible, but few empirical examples have demonstrated the ability of molecular data to change the placement of fossil taxa. We offer such an example here. We analyze the placement of snakes among squamate reptiles, combining published morphological data (363 characters) and new DNA sequence data (15,794 characters, 22 nuclear loci) for 45 living and 19 fossil taxa. We find several intriguing results. First, some fossil taxa undergo major changes in their phylogenetic position when molecular data are added. Second, most fossil taxa are placed with strong support in the expected clades by the combined data Bayesian analyses, despite each having >98% missing cells and despite recent suggestions that extensive missing data are problematic for Bayesian phylogenetics. Third, morphological data can change the placement of living taxa in combined analyses, even when there is an overwhelming majority of molecular characters. Finally, we find strong but apparently misleading signal in the morphological data, seemingly associated with a burrowing lifestyle in snakes, amphisbaenians, and dibamids. Overall, our results suggest promise for an integrated and comprehensive Tree of Life by combining molecular and morphological data for living and fossil taxa.

  19. Lake Roosevelt Fisheries Evaluation Program, Part C; Lake Roosevelt Pelagic Fish Study: Washington Department of Fish and Wildlife, 1998 Annual Report.

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Baldwin, Casey; Polacek, Matt; Bonar, Scott

    2002-11-01

    Pelagic fishes, such as kokanee and rainbow trout, provide an important fishery in Lake Roosevelt; however, spawner returns and creel results have been below management goals in recent years. Our objective was to identify factors that potentially limit pelagic fish production in Lake Roosevelt including entrainment, food limitation, piscivory, and other abiotic factors. We estimated the ratio of total fish entrained through Grand Coulee Dam to the pelagic fish abundance for September and October, 1998. If the majority of these fish were pelagic species, then entrainment averaged 10-13% of pelagic fish abundance each month. This rate of entrainment could imposemore » considerable losses to pelagic fish populations on an annual basis. Therefore, estimates of species composition of entrained fish will be important in upcoming years to estimate the proportion of stocked pelagic fish lost through the dam. Food was not limiting for kokanee or rainbow trout populations since growth rates were high and large zooplankton were present in the reservoir. Estimates of survival for kokanee were low (< 0.01 annual) and unknown for rainbow trout. We estimated that the 1997 standing stock biomass of large (>1.1 mm) Daphnia could have supported 0.08 annual survival by kokanee and rainbow trout before fish consumption would have exceeded available biomass during late winter and early spring. Therefore, if recruitment goals are met in the future there may be a bottleneck in food supply for pelagic planktivores. Walleye and northern pikeminnow were the primary piscivores of salmonids in 1996 and 1997. Predation on salmonid prey was rare for rainbow trout and not detected for burbot or smallmouth bass. Northern pikeminnow had the greatest individual potential as a salmonid predator due to their high consumptive demand; however, their overall impact was limited because of their low relative abundance. We modeled the predation impact of 273,524 walleye in 1996, and 39,075 northern

  20. Probe into the Internal Mechanism of Interlanguage Fossilization

    ERIC Educational Resources Information Center

    Huang, Qian

    2009-01-01

    Interlanguage fossilization is normal for second language acquisition. It is also a hotspot for studies on theory of foreign language acquisition. Many reasons cause the interlanguage fossilization. This paper probes into the internal mechanism of interlanguage fossilization from five aspects, namely the physiological aspect, the psychological…

  1. Assimilation of Diazotrophic Nitrogen into Pelagic Food Webs

    PubMed Central

    Woodland, Ryan J.; Holland, Daryl P.; Beardall, John; Smith, Jonathan; Scicluna, Todd; Cook, Perran L. M.

    2013-01-01

    The fate of diazotrophic nitrogen (ND) fixed by planktonic cyanobacteria in pelagic food webs remains unresolved, particularly for toxic cyanophytes that are selectively avoided by most herbivorous zooplankton. Current theory suggests that ND fixed during cyanobacterial blooms can enter planktonic food webs contemporaneously with peak bloom biomass via direct grazing of zooplankton on cyanobacteria or via the uptake of bioavailable ND (exuded from viable cyanobacterial cells) by palatable phytoplankton or microbial consortia. Alternatively, ND can enter planktonic food webs post-bloom following the remineralization of bloom detritus. Although the relative contribution of these processes to planktonic nutrient cycles is unknown, we hypothesized that assimilation of bioavailable ND (e.g., nitrate, ammonium) by palatable phytoplankton and subsequent grazing by zooplankton (either during or after the cyanobacterial bloom) would be the primary pathway by which ND was incorporated into the planktonic food web. Instead, in situ stable isotope measurements and grazing experiments clearly documented that the assimilation of ND by zooplankton outpaced assimilation by palatable phytoplankton during a bloom of toxic Nodularia spumigena Mertens. We identified two distinct temporal phases in the trophic transfer of ND from N. spumigena to the plankton community. The first phase was a highly dynamic transfer of ND to zooplankton with rates that covaried with bloom biomass while bypassing other phytoplankton taxa; a trophic transfer that we infer was routed through bloom-associated bacteria. The second phase was a slowly accelerating assimilation of the dissolved-ND pool by phytoplankton that was decoupled from contemporaneous variability in N. spumigena concentrations. These findings provide empirical evidence that ND can be assimilated and transferred rapidly throughout natural plankton communities and yield insights into the specific processes underlying the propagation of ND

  2. Assimilation of diazotrophic nitrogen into pelagic food webs.

    PubMed

    Woodland, Ryan J; Holland, Daryl P; Beardall, John; Smith, Jonathan; Scicluna, Todd; Cook, Perran L M

    2013-01-01

    The fate of diazotrophic nitrogen (N(D)) fixed by planktonic cyanobacteria in pelagic food webs remains unresolved, particularly for toxic cyanophytes that are selectively avoided by most herbivorous zooplankton. Current theory suggests that N(D) fixed during cyanobacterial blooms can enter planktonic food webs contemporaneously with peak bloom biomass via direct grazing of zooplankton on cyanobacteria or via the uptake of bioavailable N(D) (exuded from viable cyanobacterial cells) by palatable phytoplankton or microbial consortia. Alternatively, N(D) can enter planktonic food webs post-bloom following the remineralization of bloom detritus. Although the relative contribution of these processes to planktonic nutrient cycles is unknown, we hypothesized that assimilation of bioavailable N(D) (e.g., nitrate, ammonium) by palatable phytoplankton and subsequent grazing by zooplankton (either during or after the cyanobacterial bloom) would be the primary pathway by which N(D) was incorporated into the planktonic food web. Instead, in situ stable isotope measurements and grazing experiments clearly documented that the assimilation of N(D) by zooplankton outpaced assimilation by palatable phytoplankton during a bloom of toxic Nodularia spumigena Mertens. We identified two distinct temporal phases in the trophic transfer of N(D) from N. spumigena to the plankton community. The first phase was a highly dynamic transfer of N(D) to zooplankton with rates that covaried with bloom biomass while bypassing other phytoplankton taxa; a trophic transfer that we infer was routed through bloom-associated bacteria. The second phase was a slowly accelerating assimilation of the dissolved-N(D) pool by phytoplankton that was decoupled from contemporaneous variability in N. spumigena concentrations. These findings provide empirical evidence that N(D) can be assimilated and transferred rapidly throughout natural plankton communities and yield insights into the specific processes underlying

  3. Primate diversification inferred from phylogenies and fossils.

    PubMed

    Herrera, James P

    2017-12-01

    Biodiversity arises from the balance between speciation and extinction. Fossils record the origins and disappearance of organisms, and the branching patterns of molecular phylogenies allow estimation of speciation and extinction rates, but the patterns of diversification are frequently incongruent between these two data sources. I tested two hypotheses about the diversification of primates based on ∼600 fossil species and 90% complete phylogenies of living species: (1) diversification rates increased through time; (2) a significant extinction event occurred in the Oligocene. Consistent with the first hypothesis, analyses of phylogenies supported increasing speciation rates and negligible extinction rates. In contrast, fossils showed that while speciation rates increased, speciation and extinction rates tended to be nearly equal, resulting in zero net diversification. Partially supporting the second hypothesis, the fossil data recorded a clear pattern of diversity decline in the Oligocene, although diversification rates were near zero. The phylogeny supported increased extinction ∼34 Ma, but also elevated extinction ∼10 Ma, coinciding with diversity declines in some fossil clades. The results demonstrated that estimates of speciation and extinction ignoring fossils are insufficient to infer diversification and information on extinct lineages should be incorporated into phylogenetic analyses. © 2017 The Author(s). Evolution © 2017 The Society for the Study of Evolution.

  4. Proceedings: 1990 fossil plant cycling conference

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Not Available

    1991-12-01

    Fossil plant cycling continues to be a key issue for many electric utilities. EPRI's previous cycling workshops, held in 1983, 1985, and 1987, allowed utilities to benefit from collective industry experience in the conversion of baseload fossil units to cyclic operation. Continued improvements in equipment, retrofits, diagnostics, and controls were highlighted at the 1990 conference. The objective is to provide a forum for utility discussions of the cycling operation of fossil fuel power plants. Potomac Electric Power Company (PEPCO) hosted the 1990 EPRI Fossil Fuel Cycling Conference in Washington, DC, on December 4--6, 1990. More than 130 representatives from utilities,more » vendors, government agencies, universities, and industry associations attended the conference. Following the general session, technical sessions covered such topics as plant modifications, utility retrofit experience, cycling economics, life assessment, controls, environmental controls, and energy storage. Attendees also toured PEPCO's Potomac River generating station, the site of an earlier EPRI cycling conversion study.« less

  5. Effectiveness of tori line use to reduce seabird bycatch in pelagic longline fishing

    PubMed Central

    Domingo, Andrés; Abreu, Martin; Forselledo, Rodrigo; Yates, Oliver

    2017-01-01

    Industrial longline fisheries cause the death of large numbers of seabirds annually. Various mitigation measures have been proposed, including the use of tori lines. In this study the efficiency of a single tori line to reduce seabird bycatch was tested on pelagic longline vessels (25-37m length). Thirteen fishing trips were carried out in the area and season of the highest bycatch rates recorded in the southwest Atlantic (2009–2011). We deployed two treatments in random order: sets with a tori line and without a tori line (control treatment). The use of a tori line significantly reduced seabird bycatch rates. Forty three and seven birds were captured in the control (0.85 birds/1,000 hooks, n = 49 sets) and in the tori line treatment (0.13 birds/1,000 hooks, n = 51 sets), respectively. In 47% of the latter sets the tori line broke either because of entanglement with the longline gear or by tension. This diminished the tori line effectiveness; five of the seven captures during sets where a tori line was deployed were following ruptures. Nine additional trips were conducted with a tori line that was modified to reduce entanglements (2012–2016). Seven entanglements were recorded in 73 longline sets. The chance of a rupture on these trips was 4% (95% c.l. = 1–18%) of that during 2009–2011. This work shows that the use of a tori line reduces seabird bycatch in pelagic longline fisheries and is a practice suitable for medium size vessels (~25-40m length). Because the study area has historically very high bycatch rates at global level, this tori line design is potentially useful to reduce seabird bycatch in many medium size pelagic longline vessel fishing in the southern hemisphere. PMID:28886183

  6. 75 FR 1023 - International Fisheries Regulations; Fisheries in the Western Pacific; Pelagic Fisheries; Hawaii...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2010-01-08

    .... 080225267-91393-03] RIN 0648-AW49 International Fisheries Regulations; Fisheries in the Western Pacific; Pelagic Fisheries; Hawaii-based Shallow-set Longline Fishery; Correction AGENCY: National Marine Fisheries... process is preserved for closing the Hawaii-based shallow-set longline fishery as a result of the fishery...

  7. Comparing primate crania: The importance of fossils.

    PubMed

    Fleagle, John G; Gilbert, Christopher C; Baden, Andrea L

    2016-10-01

    Extant primate crania represent a small subset of primate crania that have existed. The main objective here is to examine how the inclusion of fossil crania changes our understanding of primate cranial diversity relative to analyses of extant primates. We hypothesize that fossil taxa will change the major axes of cranial shape, occupy new areas of morphospace, change the relative diversity of major primate clades, and fill in notable gaps separating major primate taxa/clades. Eighteen 3D landmarks were collected on 157 extant and fossil crania representing 90 genera. Data were subjected to a Generalized Procrustes Analysis then principal components analysis. Relative diversity between clades was assessed using an F-statistic. Fossil taxa do not significantly alter major axes of cranial shape, but they do occupy unique areas of morphospace, change the relative diversity between clades, and fill in notable gaps in primate cranial evolution. Strepsirrhines remain significantly less diverse than anthropoids. Fossil hominins fill the gap in cranial morphospace between extant great apes and modern humans. The morphospace outlined by living primates largely includes that occupied by fossil taxa, suggesting that the cranial diversity of living primates generally encompasses the total diversity that has evolved in this Order. The evolution of the anthropoid cranium was a significant event allowing anthropoids to achieve significantly greater cranial diversity compared to strepsirrhines. Fossil taxa fill in notable gaps within and between clades, highlighting their transitional nature and eliminating the appearance of large morphological distances between extant taxa, particularly in the case of extant hominids. © 2016 Wiley Periodicals, Inc.

  8. Expansion of oxygen minimum zones may reduce available habitat for tropical pelagic fishes

    NASA Astrophysics Data System (ADS)

    Stramma, Lothar; Prince, Eric D.; Schmidtko, Sunke; Luo, Jiangang; Hoolihan, John P.; Visbeck, Martin; Wallace, Douglas W. R.; Brandt, Peter; Körtzinger, Arne

    2012-01-01

    Climate model predictions and observations reveal regional declines in oceanic dissolved oxygen, which are probably influenced by global warming. Studies indicate ongoing dissolved oxygen depletion and vertical expansion of the oxygen minimum zone (OMZ) in the tropical northeast Atlantic Ocean. OMZ shoaling may restrict the usable habitat of billfishes and tunas to a narrow surface layer. We report a decrease in the upper ocean layer exceeding 3.5mll-1 dissolved oxygen at a rate of <=1myr-1 in the tropical northeast Atlantic (0-25°N, 12-30°W), amounting to an annual habitat loss of ~5.95×1013m3, or 15% for the period 1960-2010. Habitat compression and associated potential habitat loss was validated using electronic tagging data from 47 blue marlin. This phenomenon increases vulnerability to surface fishing gear for billfishes and tunas, and may be associated with a 10-50% worldwide decline of pelagic predator diversity. Further expansion of the Atlantic OMZ along with overfishing may threaten the sustainability of these valuable pelagic fisheries and marine ecosystems.

  9. Real-time distribution of pelagic fish: combining hydroacoustics, GIS and spatial modelling at a fine spatial scale.

    PubMed

    Muška, Milan; Tušer, Michal; Frouzová, Jaroslava; Mrkvička, Tomáš; Ricard, Daniel; Seďa, Jaromír; Morelli, Federico; Kubečka, Jan

    2018-03-29

    Understanding spatial distribution of organisms in heterogeneous environment remains one of the chief issues in ecology. Spatial organization of freshwater fish was investigated predominantly on large-scale, neglecting important local conditions and ecological processes. However, small-scale processes are of an essential importance for individual habitat preferences and hence structuring trophic cascades and species coexistence. In this work, we analysed the real-time spatial distribution of pelagic freshwater fish in the Římov Reservoir (Czechia) observed by hydroacoustics in relation to important environmental predictors during 48 hours at 3-h interval. Effect of diurnal cycle was revealed of highest significance in all spatial models with inverse trends between fish distribution and predictors in day and night in general. Our findings highlighted daytime pelagic fish distribution as highly aggregated, with general fish preferences for central, deep and highly illuminated areas, whereas nighttime distribution was more disperse and fish preferred nearshore steep sloped areas with higher depth. This turnover suggests prominent movements of significant part of fish assemblage between pelagic and nearshore areas on a diel basis. In conclusion, hydroacoustics, GIS and spatial modelling proved as valuable tool for predicting local fish distribution and elucidate its drivers, which has far reaching implications for understanding freshwater ecosystem functioning.

  10. Plastic particles in coastal pelagic ecosystems of the Northeast Pacific ocean.

    PubMed

    Doyle, Miriam J; Watson, William; Bowlin, Noelle M; Sheavly, Seba B

    2011-02-01

    The purpose of this study was to examine the distribution, abundance and characteristics of plastic particles in plankton samples collected routinely in Northeast Pacific ecosystems, and to contribute to the development of ideas for future research into the occurrence and impact of small plastic debris in marine pelagic ecosystems. Plastic debris particles were assessed from zooplankton samples collected as part of the National Oceanic and Atmospheric Administration's (NOAA) ongoing ecosystem surveys during two research cruises in the Southeast Bering Sea in the spring and fall of 2006 and four research cruises off the U.S. west coast (primarily off southern California) in spring, summer and fall of 2006, and in January of 2007. Nets with 0.505 mm mesh were used to collect surface samples during all cruises, and sub-surface samples during the four cruises off the west coast. The 595 plankton samples processed indicate that plastic particles are widely distributed in surface waters. The proportion of surface samples from each cruise that contained particles of plastic ranged from 8.75 to 84.0%, whereas particles were recorded in sub-surface samples from only one cruise (in 28.2% of the January 2007 samples). Spatial and temporal variability was apparent in the abundance and distribution of the plastic particles and mean standardized quantities varied among cruises with ranges of 0.004-0.19 particles/m³, and 0.014-0.209 mg dry mass/m³. Off southern California, quantities for the winter cruise were significantly higher, and for the spring cruise significantly lower than for the summer and fall surveys (surface data). Differences between surface particle concentrations and mass for the Bering Sea and California coast surveys were significant for pair-wise comparisons of the spring but not the fall cruises. The particles were assigned to three plastic product types: product fragments, fishing net and line fibers, and industrial pellets; and five size categories: <1 mm

  11. Organic preservation of fossil musculature with ultracellular detail

    PubMed Central

    McNamara, Maria; Orr, Patrick J.; Kearns, Stuart L.; Alcalá, Luis; Anadón, Pere; Peñalver-Mollá, Enrique

    2010-01-01

    The very labile (decay-prone), non-biomineralized, tissues of organisms are rarely fossilized. Occurrences thereof are invaluable supplements to a body fossil record dominated by biomineralized tissues, which alone are extremely unrepresentative of diversity in modern and ancient ecosystems. Fossil examples of extremely labile tissues (e.g. muscle) that exhibit a high degree of morphological fidelity are almost invariably replicated by inorganic compounds such as calcium phosphate. There is no consensus as to whether such tissues can be preserved with similar morphological fidelity as organic remains, except when enclosed inside amber. Here, we report fossilized musculature from an approximately 18 Myr old salamander from lacustrine sediments of Ribesalbes, Spain. The muscle is preserved organically, in three dimensions, and with the highest fidelity of morphological preservation yet documented from the fossil record. Preserved ultrastructural details include myofilaments, endomysium, layering within the sarcolemma, and endomysial circulatory vessels infilled with blood. Slight differences between the fossil tissues and their counterparts in extant amphibians reflect limited degradation during fossilization. Our results provide unequivocal evidence that high-fidelity organic preservation of extremely labile tissues is not only feasible, but likely to be common. This is supported by the discovery of similarly preserved tissues in the Eocene Grube Messel biota. PMID:19828545

  12. Dinosaur Fossil, Leonardo, at Ellington Field

    NASA Image and Video Library

    2008-03-14

    Documentation of NASA's partnership with the Houston Museum of Natural Science and Montana's Great Plains Dinosaur Museum to provide an insulated facility at Ellington Field in which the Leonardo Project Team was able to X-ray Leonardo, a 77 million year old dinosaur fossil. View of the Brachylophosaurus fossil called Leonardo.

  13. Interested in Pelagic Food Webs? BCO-DMO has your Data.

    NASA Astrophysics Data System (ADS)

    Chandler, C. L.; Groman, R. C.; Kinkade, D.; Rauch, S.; Allison, M. D.; Gegg, S. R.; Shepherd, A.; Wiebe, P. H.; Glover, D. M.

    2016-02-01

    Interdisciplinary research collaborations that address complex, global research themes such as the interactive effects of global warming and studies of pelagic food webs require access to a broad range of data types from all disciplines of oceanography, from all platforms (e.g. ships, gliders, floats, moorings), with the in situ observations complementing and being complemented by laboratory and model results. In an effort to build a comprehensive database of marine ecosystem research data, the National Science Foundation (NSF) funded the Biological and Chemical Oceanography Data Management Office (BCO-DMO; bco-dmo.org) to support the data management requirements of investigators funded by the NSF's Polar Programs (PLR) and Biological and Chemical Oceanography Sections (OCE). Since 2006, investigators funded by NSF PLR and OCE have been working with support from BCO-DMO data scientists, to build a data system that now includes the full range of ocean biogeochemistry data resulting from decades of research. In addition to data from recently funded PIs, the BCO-DMO data system also serves data from legacy programs (e.g. US Joint Global Ocean Flux Study and US Global Ocean Ecosystem Dynamics). The data are open-access, available for download in a variety of user-selectable formats, and accompanied by sufficient documentation to enable re-use. This presentation will highlight the diversity of data available from the BCO-DMO system and demonstrate some of the features that enable discovery, access and download of data relevant to studies of pelagic food webs.

  14. Reconstructing Carotenoid-Based and Structural Coloration in Fossil Skin.

    PubMed

    McNamara, Maria E; Orr, Patrick J; Kearns, Stuart L; Alcalá, Luis; Anadón, Pere; Peñalver, Enrique

    2016-04-25

    Evidence of original coloration in fossils provides insights into the visual communication strategies used by ancient animals and the functional evolution of coloration over time [1-7]. Hitherto, all reconstructions of the colors of reptile integument and the plumage of fossil birds and feathered dinosaurs have been of melanin-based coloration [1-6]. Extant animals also use other mechanisms for producing color [8], but these have not been identified in fossils. Here we report the first examples of carotenoid-based coloration in the fossil record, and of structural coloration in fossil integument. The fossil skin, from a 10 million-year-old colubrid snake from the Late Miocene Libros Lagerstätte (Teruel, Spain) [9, 10], preserves dermal pigment cells (chromatophores)-xanthophores, iridophores, and melanophores-in calcium phosphate. Comparison with chromatophore abundance and position in extant reptiles [11-15] indicates that the fossil snake was pale-colored in ventral regions; dorsal and lateral regions were green with brown-black and yellow-green transverse blotches. Such coloration most likely functioned in substrate matching and intraspecific signaling. Skin replicated in authigenic minerals is not uncommon in exceptionally preserved fossils [16, 17], and dermal pigment cells generate coloration in numerous reptile, amphibian, and fish taxa today [18]. Our discovery thus represents a new means by which to reconstruct the original coloration of exceptionally preserved fossil vertebrates. Copyright © 2016 Elsevier Ltd. All rights reserved.

  15. Studying the Relative Strengths of Environmental Factors that Influence Echinoderm Body Size Trends

    NASA Astrophysics Data System (ADS)

    Low, A.; Randhawa, S.; Heim, N. A.; Payne, J.

    2013-12-01

    Body size is often a useful metric in observing how a clade responds to environmental changes. Previous research has uncovered how environmental factors such as carbon dioxide and oxygen levels influence body size evolution. However, we wanted to look into how these natural factors interact and which factors seem to have a stronger relative influence on echinoderm body size. We analyzed carbon dioxide levels, a proxy for paleotemperature, oxygen levels, and sea level. Our research process involved measuring and calculating the volume of Phanerozoic echinoderm fossils recorded in the Treatise on Invertebrate Paleontology, plotting their mean volumes over various natural factors, and using statistical tools such as correlation tests and the PaleoTS statistical analysis software to compare the relative strengths of these factors. Furthermore, we divided our data into the following three subsets to uncover more specific relationships: 1) A set that included all data of the phylum Echinodermata 2) A set that focused on the two classes with the most recorded data, Echinoidea and Crinoidea 3) A set that focused on the crinoid specimens that originated in the Paleozoic and in the post-Paleozoic. In the first subset, echinoderms had the strongest correlation with carbon dioxide, a proxy for temperature, and possessed a weaker correlation with oxygen. In the second subset, we discovered that the echinoid data also possessed a strong correlation with carbon dioxide and a weaker correlation with oxygen. For crinoids, we found that the class as a whole showed no strong correlation with any measured environmental factors. However, when we divided the crinoids based on age, we found that both Paleozoic and post-Paleozoic crinoids individually correlated strongly with sea level. However, some uncertainty with this correlation arose as the comparison of the environmental correlate models suggested that an unbiased random walk was the best fit for the data. This stands as a sharp

  16. Instabilities in biofilms: The Kinneyia Fossil

    NASA Astrophysics Data System (ADS)

    Thomas, K. R.; Goehring, L.; Porada, H.; Wittig, R.; Herminghaus, S.

    2012-12-01

    Kinneyia structures are a wrinkle-type fossil pattern most often observed in ancient siliclastic sediment surfaces. Characterised by millimetre-scale ripples with flat-topped crests, these fossils are generally found in areas that were formally littoral habitats. Thin-section observations indicate that Kinneyia formed in surfaces covered by ancient microbial mats. However, to date there has been no conclusive explanation as to the process involved in the formation of these fossils. We propose that the key mechanism involved in the formation of the Kinneyia pattern is a Kelvin-Helmholtz-type instability induced in a viscoelastic film under flowing water. A ripple corrugation is spontaneously induced in the film, which grows in amplitude over time. Such a mechanism is expected to result in a ripple instability with a wavelength proportional to the thickness of the film. Experiments carried out using viscoelastic models microbial mats confirm this prediction, showing a wavelength roughly three times the thickness of the film. The behaviour is independent on the viscoelastic properties of the film. This model corresponds well with the fossil records, which show a reduced wavelength at the boundaries of the fossilised structures where the mat is expected to have been thinner. Fossils were collected from two ancient shallow sea bed sites in Namibia from the terminal Proterozoic period. The fossils are seen to overlay a storm deposit of 15-30cm in thickness. The ripples form on top of this deposit in the veneer, which sedimented after the storm event. Analysis of the shape of the fossilised patterns indicates a similar relationship between the wavelength and amplitude of the ripples to that observed experimentally. A strong directional dependence of the ripples is also observed.

  17. Cerium anomaly at microscale in fossils.

    PubMed

    Gueriau, Pierre; Mocuta, Cristian; Bertrand, Loïc

    2015-09-01

    Patterns in rare earth element (REE) concentrations are essential instruments to assess geochemical processes in Earth and environmental sciences. Excursions in the "cerium anomaly" are widely used to inform on past redox conditions in sediments. This proxy resources to the specificity of cerium to adopt both the +III and +IV oxidation states, while most rare earths are purely trivalent and share very similar reactivity and transport properties. In practical terms, the level of cerium anomaly is established through elemental point quantification and profiling. All these models rely on a supposed homogeneity of the cerium oxidation state within the samples. However, this has never been demonstrated, whereas the cerium concentration can significantly vary within a sample, as shown for fossils, which would vastly complicate interpretation of REE patterns. Here, we report direct micrometric mapping of Ce speciation through synchrotron X-ray absorption spectroscopy and production of local rare earth patterns in paleontological fossil tissues through X-ray fluorescence mapping. The sensitivity of the approach is demonstrated on well-preserved fishes and crustaceans from the Late Cretaceous (ca. 95 million years (Myr) old). The presence of Ce under the +IV form within the fossil tissues is attributed to slightly oxidative local conditions of burial and agrees well with the limited negative cerium anomaly observed in REE patterns. The [Ce(IV)]/[Ce(tot)] ratio appears remarkably stable at the microscale within each fossil and is similar between fossils from the locality. Speciation maps were obtained from an original combination of synchrotron microbeam X-ray fluorescence, absorption spectroscopy, and diffraction, together with light and electron microscopy. This work also highlights the need for more systematic studies of cerium geochemistry at the microscale in paleontological contexts, in particular across fossil histologies.

  18. 78 FR 18249 - Fisheries Off West Coast States; Coastal Pelagic Species Fisheries; Annual Specifications

    Federal Register 2010, 2011, 2012, 2013, 2014

    2013-03-26

    ... limit (ACL), harvest guideline (HG), annual catch target (ACT) and associated annual reference points... Coastal Pelagic Species (CPS) Fishery Management Plan (FMP). The 2012-2013 ACL or maximum HG for Pacific... difference between the ACL and ACT (10,128 mt) as a set aside for incidental landings in other CPS fisheries...

  19. 78 FR 79388 - Western Pacific Pelagic Fisheries; Catch and Effort Limits for the U.S. Participating Territories

    Federal Register 2010, 2011, 2012, 2013, 2014

    2013-12-30

    ...NMFS announces that the Western Pacific Fishery Management Council proposes to amend the Fishery Ecosystem Plan for Pelagic Fisheries of the Western Pacific Region. If approved, Amendment 7 would establish a management framework and process for specifying fishing catch and effort limits and accountability measures for pelagic fisheries in the U.S. Pacific territories (American Samoa, Guam, and the Commonwealth of the Northern Mariana Islands). The framework would authorize the government of each territory to allocate a portion of its specified catch or effort limit to a U.S. fishing vessel or vessels through a specified fishing agreement, and establish criteria, which a specified fishing agreement must satisfy. The framework also includes measures to ensure accountability for adhering to fishing catch and effort limits.

  20. Fossil Energy: Drivers and Challenges.

    NASA Astrophysics Data System (ADS)

    Friedmann, Julio

    2007-04-01

    Concerns about rapid economic growth, energy security, and global climate change have created a new landscape for fossil energy exploration, production, and utilization. Since 85% of primary energy supply comes from fossil fuels, and 85% of greenhouse gas emissions come from fossil fuel consumption, new and difficult technical and political challenges confront commercial, governmental, and public stakeholders. As such, concerns over climate change are explicitly weighed against security of international and domestic energy supplies, with economic premiums paid for either or both. Efficiency improvements, fuel conservation, and deployment of nuclear and renewable supplies will help both concerns, but are unlikely to offset growth in the coming decades. As such, new technologies and undertakings must both provide high quality fossil energy with minimal environmental impacts. The largest and most difficult of these undertakings is carbon management, wherein CO2 emissions are sequestered indefinitely at substantial incremental cost. Geological formations provide both high confidence and high capacity for CO2 storage, but present scientific and technical challenges. Oil and gas supply can be partially sustained and replaced through exploitation of unconventional fossil fuels such as tar-sands, methane hydrates, coal-to-liquids, and oil shales. These fuels provide enormous reserves that can be exploited at current costs, but generally require substantial energy to process. In most cases, the energy return on investment (EROI) is dropping, and unconventional fuels are generally more carbon intensive than conventional, presenting additional carbon management challenges. Ultimately, a large and sustained science and technology program akin to the Apollo project will be needed to address these concerns. Unfortunately, real funding in energy research has dropped dramatically (75%) in the past three decades, and novel designs in fission and fusion are not likely to provide any

  1. Divergence time estimates of mammals from molecular clocks and fossils: relevance of new fossil finds from India.

    PubMed

    Prasad, G V R

    2009-11-01

    This paper presents a brief review of recent advances in the classification of mammals at higher levels using fossils and molecular clocks. It also discusses latest fossil discoveries from the Cretaceous - Eocene (66-55 m.y.) rocks of India and their relevance to our current understanding of placental mammal origins and diversifications.

  2. Status and trends of pelagic prey fishes in Lake Huron, 2012

    USGS Publications Warehouse

    Warner, David M.; O'Brien, Timothy P.; Farha, Steve A.; Schaeffer, Jeff; Lenart, Stephen

    2012-01-01

    The USGS Great Lakes Science Center (GLSC) conducted acoustic/midwater trawl surveys of Lake Huron during 1997 and annually during 2004-2012. The 2012 survey was conducted during September and October, and included transects in Lake Huron’s Main Basin, Georgian Bay, and North Channel. Pelagic fish density (638 fish/ha) was lower in 2012 compared to 2011, with density in 2012 only 34% of 2011. Total biomass in 2012 was 74% of the 2011 value. Alewife Alosa pseudoharengus remained nearly absent, and only one cisco Coregonus artedi was captured. Rainbow smelt Osmerus mordax density was only 31% of the 2011 density. Bloater Coregonus hoyi density was less than half the 2011 density, mostly as a result of lower density of small bloater. Density and biomass of large bloater in 2012 were similar to 2011 levels. During 2012 we observed significantly higher fish biomass in North Channel than in the Main Basin or Georgian Bay. Prey availability during 2013 will likely be similar to 2012. Lake Huron now has pelagic fish biomass similar to that observed in recent lakewide acoustic surveys of Lake Michigan and Lake Superior, but species composition differs in the three lakes. There is an increasing diversity and prevalence of native species gradient from Lake Michigan to Lake Superior, with Lake Huron being intermediate in the prevalence of native fish species like coregonines and emerald shiner Notropis atherinoides.

  3. Status and trends of pelagic prey fishes in Lake Huron, 2012

    USGS Publications Warehouse

    Warner, David M.; O'Brien, Timothy P.; Farha, Steve A.; Schaeffer, Jeff; Lenart, Stephen

    2013-01-01

    The USGS Great Lakes Science Center (GLSC) conducted acoustic/midwater trawl surveys of Lake Huron during 1997 and annually during 2004-2012. The 2012 survey was conducted during September and October, and included transects in Lake Huron’s Main Basin, Georgian Bay, and North Channel. Pelagic fish density (638 fish/ha) was lower in 2012 compared to 2011, with density in 2012 only 34% of 2011. Total biomass in 2012 was 74% of the 2011 value. Alewife Alosa pseudoharengus remained nearly absent, and only one cisco Coregonus artedi was captured. Rainbow smelt Osmerus mordax density was only 31% of the 2011 density. Bloater Coregonus hoyi density was less than half the 2011 density, mostly as a result of lower density of small bloater. Density and biomass of large bloater in 2012 were similar to 2011 levels. During 2012 we observed significantly higher fish biomass in North Channel than in the Main Basin or Georgian Bay. Prey availability during 2013 will likely be similar to 2012. Lake Huron now has pelagic fish biomass similar to that observed in recent lakewide acoustic surveys of Lake Michigan and Lake Superior, but species composition differs in the three lakes. There is an increasing diversity and prevalence of native species gradient from Lake Michigan to Lake Superior, with Lake Huron being intermediate in the prevalence of native fish species like coregonines and emerald shiner Notropis atherinoides.

  4. Effect of Macondo Prospect 252 Oil on Microbiota Associated with Pelagic Sargassum in the Northern Gulf of Mexico.

    PubMed

    Torralba, Manolito G; Franks, James S; Gomez, Andres; Yooseph, Shibu; Nelson, Karen E; Grimes, D Jay

    2017-01-01

    The environmental impact of major oil spills on marine microorganisms has yet to be thoroughly investigated using molecular biology techniques. The Deepwater Horizon (DWH) drilling rig explosion of 2010 affected an approximately 176,000 km 2 surface area of the Gulf of Mexico (GOM) when an estimated 210 million gallons of oil from the Macondo Prospect spilled into the environment. Pelagic Sargassum, a complex of two surface drifting species (Sargassum natans and Sargassum fluitans) of marine brown macroalgae and a critically important habitat in the GOM ecosystem, was suffused by Macondo Prospect 252 oil released during the DWH event. Using 16S rRNA PCR and Roche 454 pyrosequencing, the effect of the oil on the bacterial population associated with pelagic Sargassum and contiguous waters was examined by comparing sequence data generated from samples collected from oiled and non-oiled locations in the northern GOM. Sequence data showed similar microbial composition in Sargassum regardless of exposure to oil primarily dominated by five phyla; Proteobacteria, Bacteroidetes, Actinobacteria, Verrucomicrobia, and unclassified bacteria. The microbial composition in water samples was significantly less diverse than for Sargassum and consisted primarily of Proteobacteria, Firmicutes, and Bacteroidetes. Due to the evenly distributed abundance of microbial species on oiled and non-oiled pelagic Sargassum, study findings indicate that DWH spilled oil had minimal effect on the composition and diversity of the microbial community associated with Sargassum and contiguous waters. However, higher abundances of Sulfitobacter and one species of Psychrobacter were found in oiled water samples when compared to non-oiled water samples indicating some effect of DHW oil in the microbial composition of seawater. Though there are a number of marine studies using molecular biology approaches, this is the first molecular examination of the impact of the DWH oil spill on bacterial communities

  5. Redefinition of the Meramecian/Chesterian boundary (Mississippian)

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Maples, C.G.; Waters, J.A.

    1987-07-01

    The Meramecian/Chesterian (Mississippi) boundary in the type area is currently defined as the highest occurrence of the crinoid Platycrinites penicillus and the lowest occurrence of the crinoid genera Agassizocrinus and Talarocrinus. Because these taxa have not been reported outside eastern North America, attempts have been made to use conodonts and Foraminifera to extend the Meramecian/Chesterian boundary outside the type area. Unfortunately, changes in conodont and foraminiferal assemblages do not coincide with the currently defined Meramecian/Chesterian boundary, and use of these fossil groups does not allow exact placement of the boundary outside of eastern North America. The authors suggest that themore » Meramecian/Chesterian boundary be redefined as the boundary between Foraminiferal Zones 15 and 16i of the Mamet scheme, irrespective of the occurrence of Platycrinites penicillus. This change in definition places the Ste. Genevieve Limestone (Genevievian Stage) in the lowermost part of the Chesterian Series, with which it is biotically and sedimentologically more allied than with the underlying St. Louis Limestone. This change provides a sound biostratigraphic base for correlating the Meramecian/Chesterian boundary outside of the type area. Foraminifera, conodonts, brachiopods, and corals all show significant changes at or slightly below the St. Louis/Ste. Genevieve contact rather than at the Genevievian/Chesterian contact. 52 references.« less

  6. Biostratigraphy and petrography of upper Paleozoic rocks of Sierra Las Pintas, northern Baja California

    NASA Astrophysics Data System (ADS)

    Navas-Parejo, Pilar; Lara-Peña, R. Aaron; Torres-Martínez, Miguel Angel; Martini, Michelangelo

    2018-07-01

    A transported crinoid fauna is herein described for the first time in the Paleozoic succession cropping out in the Sierra Las Pintas, northern Baja California, northwestern Mexico. The fossil association includes Heterostelechus texanus Moore and Jeffords, Preptopremnum laeve? Moore and Jeffords, and Mooreanteris perforatus Moore and Jeffords, which indicates a Middle Pennsylvanian-early Permian time-averaged age. The studied area corresponds with the northernmost outcrop of definitely late Paleozoic deep-water facies in northwestern Mexico and the southern United States. Petrographic analyses indicate that the studied metasandstones were primarily derived from high-grade metamorphic rocks and from a shallow-water platform environment dominated by crinoid meadows. These results allow the correlation of the studied metasedimentary rocks with the Carboniferous Rancho Nuevo Formation of the Sonora allochthon, which crops out in central Sonora. The Sonora allochthon includes an Early Ordovician-Late Pennsylvanian sedimentary succession that was deposited in the oceanic basin located south of the Laurentian craton. Therefore, upper Paleozoic metasedimentary rocks of the Sierra Las Pintas were deposited along the same continental margin of Laurentia as those rocks in the Sonora allochthon, and were mostly derived from metamorphic rocks of the continental craton and by the typical Carboniferous encrinites, which characterize the shallow-water rocks of central and northern Sonora.

  7. Dental development in living and fossil orangutans.

    PubMed

    Smith, Tanya M

    2016-05-01

    Numerous studies have investigated molar development in extant and fossil hominoids, yet relatively little is known about orangutans, the only great ape with an extensive fossil record. This study characterizes aspects of dental development, including cuspal enamel daily secretion rate, long-period line periodicities, cusp-specific molar crown formation times and extension rates, and initiation and completion ages in living and fossil orangutan postcanine teeth. Daily secretion rate and periodicities in living orangutans are similar to previous reports, while crown formation times often exceed published values, although direct comparisons are limited. One wild Bornean individual died at 4.5 years of age with fully erupted first molars (M1s), while a captive individual and a wild Sumatran individual likely erupted their M1s around five or six years of age. These data underscore the need for additional samples of orangutans of known sex, species, and developmental environment to explore potential sources of variation in molar emergence and their relationship to life history variables. Fossil orangutans possess larger crowns than living orangutans, show similarities in periodicities, and have faster daily secretion rate, longer crown formation times, and slower extension rates. Molar crown formation times exceed reported values for other fossil apes, including Gigantopithecus blacki. When compared to African apes, both living and fossil orangutans show greater cuspal enamel thickness values and periodicities, resulting in longer crown formation times and slower extension rates. Several of these variables are similar to modern humans, representing examples of convergent evolution. Molar crown formation does not appear to be equivalent among extant great apes or consistent within living and fossil members of Pongo or Homo. Copyright © 2016 The Author. Published by Elsevier Ltd.. All rights reserved.

  8. Multilocus phylogeny, divergence times, and a major role for the benthic-to-pelagic axis in the diversification of grunts (Haemulidae).

    PubMed

    Tavera, Jose; Acero P, Arturo; Wainwright, Peter C

    2018-04-01

    We present a phylogenetic analysis with divergence time estimates, and an ecomorphological assessment of the role of the benthic-to-pelagic axis of diversification in the history of haemulid fishes. Phylogenetic analyses were performed on 97 grunt species based on sequence data collected from seven loci. Divergence time estimation indicates that Haemulidae originated during the mid Eocene (54.7-42.3 Ma) but that the major lineages were formed during the mid-Oligocene 30-25 Ma. We propose a new classification that reflects the phylogenetic history of grunts. Overall the pattern of morphological and functional diversification in grunts appears to be strongly linked with feeding ecology. Feeding traits and the first principal component of body shape strongly separate species that feed in benthic and pelagic habitats. The benthic-to-pelagic axis has been the major axis of ecomorphological diversification in this important group of tropical shoreline fishes, with about 13 transitions between feeding habitats that have had major consequences for head and body morphology. Copyright © 2017 Elsevier Inc. All rights reserved.

  9. [Infrared spectroscopy and XRD studies of coral fossils].

    PubMed

    Chen, Quan-li; Zhou, Guan-min; Yin, Zuo-wei

    2012-08-01

    Coral fossil is an old remain of multicellular animal on the earth, and formed by various geological processes. The structural characteristics and compositions of the coral fossils with different color and radial texture on the surface were studied by infrared absorption spectroscopy and X-ray powder diffraction analyses. The results show that the studied coral fossils mainly are composed of SiO2, and the radial microstructure characterized by the calcareous coral cross-section is preserved. It is formed by metasomatism by SiO2. The infrared absorption spectra of the coral fossil with different color and texture are essentially the same, showing typical infrared absorption spectra of the quartz jade. XRD analysis shows that the main components of the coral fossils with different color and texture are consistent and mainly composed of SiO2 with a trace amount of other minerals and without CaCO3.

  10. Fossil energy biotechnology: A research needs assessment. Final report

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Not Available

    1993-11-01

    The Office of Program Analysis of the US Department of Energy commissioned this study to evaluate and prioritize research needs in fossil energy biotechnology. The objectives were to identify research initiatives in biotechnology that offer timely and strategic options for the more efficient and effective uses of the Nation`s fossil resource base, particularly the early identification of new and novel applications of biotechnology for the use or conversion of domestic fossil fuels. Fossil energy biotechnology consists of a number of diverse and distinct technologies, all related by the common denominator -- biocatalysis. The expert panel organized 14 technical subjects intomore » three interrelated biotechnology programs: (1) upgrading the fuel value of fossil fuels; (2) bioconversion of fossil feedstocks and refined products to added value chemicals; and, (3) the development of environmental management strategies to minimize and mitigate the release of toxic and hazardous petrochemical wastes.« less

  11. Using Strong Gravitational Lensing to Identify Fossil Group Progenitors

    NASA Astrophysics Data System (ADS)

    Johnson, Lucas E.; Irwin, Jimmy A.; White, Raymond E., III; Wong, Ka-Wah; Maksym, W. Peter; Dupke, Renato A.; Miller, Eric D.; Carrasco, Eleazar R.

    2018-04-01

    Fossil galaxy systems are classically thought to be the end result of galaxy group/cluster evolution, as galaxies experiencing dynamical friction sink to the center of the group potential and merge into a single, giant elliptical that dominates the rest of the members in both mass and luminosity. Most fossil systems discovered lie within z < 0.2, which leads to the question, what were these systems’ progenitors? Such progenitors are expected to have imminent or ongoing major merging near the brightest group galaxy that, when concluded, will meet the fossil criteria within the look forward time. Since strong gravitational lensing preferentially selects groups merging along the line of sight, or systems with a high mass concentration like fossil systems, we searched the CASSOWARY survey of strong-lensing events with the goal of determining whether lensing systems have any predisposition to being fossil systems or progenitors. We find that ∼13% of lensing groups are identified as traditional fossils while only ∼3% of nonlensing control groups are. We also find that ∼23% of lensing systems are traditional fossil progenitors compared to ∼17% for the control sample. Our findings show that strong-lensing systems are more likely to be fossil/pre-fossil systems than comparable nonlensing systems. Cumulative galaxy luminosity functions of the lensing and nonlensing groups also indicate a possible, fundamental difference between strong-lensing and nonlensing systems’ galaxy populations, with lensing systems housing a greater number of bright galaxies even in the outskirts of groups.

  12. Detection of pelagic habitat hotspots for skipjack tuna in the Gulf of Bone-Flores Sea, southwestern Coral Triangle tuna, Indonesia

    PubMed Central

    Farhum, Aisjah; Safruddin, Safruddin; Selamat, Muhammad Banda; Sudirman, Sudirman; Nurdin, Nurjannah; Syamsuddin, Mega; Ridwan, Muhammad; Saitoh, Sei-Ichi

    2017-01-01

    Using remote sensing of sea surface temperature (SST), sea surface height anomaly (SSHA) and chlorophyll-a (Chl-a) together with catch data, we investigated the detection and persistence of important pelagic habitat hotspots for skipjack tuna in the Gulf of Bone-Flores Sea, Indonesia. We analyzed the data for the period between the northwest and southeast monsoon 2007–2011. A pelagic hotspot index was constructed from a model of multi-spectrum satellite-based oceanographic data in relation to skipjack fishing performance. Results showed that skipjack catch per unit efforts (CPUEs) increased significantly in areas of highest pelagic hotspot indices. The distribution and dynamics of habitat hotspots were detected by the synoptic measurements of SST, SSHA and Chl-a ranging from 29.5° to 31.5°C, from 2.5 to 12.5 cm and from 0.15 to 0.35 mg m-3, respectively. Total area of hotspots consistently peaked in May. Validation of skipjack CPUE predicted by our model against observed data from 2012 was highly significant. The key pelagic habitat corresponded with the Chl-a front, which could be related to the areas of relatively high prey abundance (enhanced feeding opportunity) for skipjack. We found that the area and persistence of the potential skipjack habitat hotspots for the 5 years were clearly identified by the 0.2 mg m-3 Chl-a isopleth, suggesting that the Chl-a front provides a key oceanographic indicator for global understanding on skipjack tuna habitat hotspots in the western tropical Pacific Ocean, especially within Coral Triangle tuna. PMID:28968405

  13. Detection of pelagic habitat hotspots for skipjack tuna in the Gulf of Bone-Flores Sea, southwestern Coral Triangle tuna, Indonesia.

    PubMed

    Zainuddin, Mukti; Farhum, Aisjah; Safruddin, Safruddin; Selamat, Muhammad Banda; Sudirman, Sudirman; Nurdin, Nurjannah; Syamsuddin, Mega; Ridwan, Muhammad; Saitoh, Sei-Ichi

    2017-01-01

    Using remote sensing of sea surface temperature (SST), sea surface height anomaly (SSHA) and chlorophyll-a (Chl-a) together with catch data, we investigated the detection and persistence of important pelagic habitat hotspots for skipjack tuna in the Gulf of Bone-Flores Sea, Indonesia. We analyzed the data for the period between the northwest and southeast monsoon 2007-2011. A pelagic hotspot index was constructed from a model of multi-spectrum satellite-based oceanographic data in relation to skipjack fishing performance. Results showed that skipjack catch per unit efforts (CPUEs) increased significantly in areas of highest pelagic hotspot indices. The distribution and dynamics of habitat hotspots were detected by the synoptic measurements of SST, SSHA and Chl-a ranging from 29.5° to 31.5°C, from 2.5 to 12.5 cm and from 0.15 to 0.35 mg m-3, respectively. Total area of hotspots consistently peaked in May. Validation of skipjack CPUE predicted by our model against observed data from 2012 was highly significant. The key pelagic habitat corresponded with the Chl-a front, which could be related to the areas of relatively high prey abundance (enhanced feeding opportunity) for skipjack. We found that the area and persistence of the potential skipjack habitat hotspots for the 5 years were clearly identified by the 0.2 mg m-3 Chl-a isopleth, suggesting that the Chl-a front provides a key oceanographic indicator for global understanding on skipjack tuna habitat hotspots in the western tropical Pacific Ocean, especially within Coral Triangle tuna.

  14. 85 million years of pelagic ecosystem evolution: Pacific Ocean deep-sea ichthyolith records reveal fish community dynamics and a long-term decline in sharks

    NASA Astrophysics Data System (ADS)

    Sibert, E. C.; Norris, R. D.; Cuevas, J. M.; Graves, L. G.

    2015-12-01

    The structure and productivity of open ocean consumers has undergone major changes over the past 85 million years. Here, we present the first long-term detailed records of pelagic fish and sharks utilizing the record of ichthyoliths (teeth and dermal scales) from the deep Pacific Ocean. While the North and South Pacific Oceans show similar patterns throughout the 85 million year history, the North Pacific ichthyolith accumulation is significantly higher than the South Pacific, suggesting that the basin has been a more productive region for tens of millions of years. Fish and sharks were not abundant in the Pacific gyres until ~75 million years ago (Ma) suggesting that neither group was quantitatively important in oligotrophic pelagic food webs prior to the latest Cretaceous. Relative to ray-finned fish, sharks were common in the ancient ocean. Most ichthyolith assemblages have >50% shark dermal scales (denticles), but denticle abundance has been declining in both absolute and relative abundance since the Cretaceous-Paleogene (K/Pg) mass extinction. The accumulation rate of ichthyoliths of both sharks and ray-finned fish was highest in the Early Eocene, during the peak of the Cenozoic 'greenhouse' climate where production of shark dermal denticles and fish teeth increased almost five times over Paleocene production rates. Ichthyolith fluxes fell with cooler climates in the later Eocene and Oligocene, but fish production is almost always higher than in the Cretaceous and Paleocene reflecting the expanded ecological roles and importance of pelagic fish in marine ecosystems. Shark denticle production fell to less than half that of the Cretaceous by 20 Ma when it dropped abruptly to near-zero levels. Currently denticles make up <2% of the ichthyolith assemblages when present at all. Ecologically, pelagic sharks appear to be falling as major pelagic consumers over the Late Cretaceous and Cenozoic, and particularly over the past 20 Ma, perhaps reflecting demographic

  15. 78 FR 23540 - Fisheries in the Western Pacific; American Samoa Pelagic Longline Limited Entry Program

    Federal Register 2010, 2011, 2012, 2013, 2014

    2013-04-19

    ... individual wage record, ownership title, vessel registration, or other official documents. The documents must show either ownership of a vessel that was used to fish in the EEZ around American Samoa, or evidence... the Western Pacific; American Samoa Pelagic Longline Limited Entry Program AGENCY: National Marine...

  16. The uptake of macroplastic & microplastic by demersal & pelagic fish in the Northeast Atlantic around Scotland.

    PubMed

    Murphy, Fionn; Russell, Marie; Ewins, Ciaran; Quinn, Brian

    2017-09-15

    This study reports plastic ingestion in various fish found from coastal and offshore sites in Scottish marine waters. Coastal samples consisted of three demersal flatfish species (n=128) collected from the East and West coasts of Scotland. Offshore samples consisted of 5 pelagic species and 4 demersal species (n=84) collected from the Northeast Atlantic. From the coastal fish sampled, 47.7% of the gastrointestinal tracts contained macroplastic and microplastic. Of the 84 pelagic and demersal offshore fish, only 2 (2.4%) individuals from different species had ingested plastic identified as a clear polystyrene fibre and a black polyamide fibre. The average number of plastic items found per fish from all locations that had ingested plastic was 1.8 (±1.7) with polyamide (65.3%), polyethylene terephthalate (14.4%) and acrylic (14.4%) being the three most commonly found plastics. This study adds to the existing data on macroplastic and microplastic ingestion in fish species. Crown Copyright © 2017. Published by Elsevier Ltd. All rights reserved.

  17. Larval assemblages of large and medium-sized pelagic species in the Straits of Florida

    NASA Astrophysics Data System (ADS)

    Richardson, David E.; Llopiz, Joel K.; Guigand, Cedric M.; Cowen, Robert K.

    2010-07-01

    Critical gaps in our understanding of the distributions, interactions, life histories and preferred habitats of large and medium-size pelagic fishes severely constrain the implementation of ecosystem-based, spatially structured fisheries management approaches. In particular, spawning distributions and the environmental characteristics associated with the early life stages are poorly documented. In this study, we consider the diversity, assemblages, and associated habitat of the larvae of large and medium-sized pelagic species collected during 2 years of monthly surveys across the Straits of Florida. In total, 36 taxa and 14,295 individuals were collected, with the highest diversity occurring during the summer and in the western, frontal region of the Florida Current. Only a few species (e.g. Thunnus obesus, T. alalunga, Tetrapturus pfluegeri) considered for this study were absent. Small scombrids (e.g. T. atlanticus, Katsuwonus pelamis, Auxis spp.) and gempylids dominated the catch and were orders of magnitude more abundant than many of the rare species (e.g. Thunnus thynnus,Kajikia albida). Both constrained (CCA) and unconstrained (NMDS) multivariate analyses revealed a number of species groupings including: (1) a summer Florida edge assemblage (e.g. Auxis spp., Euthynnus alleterattus, Istiophorus platypterus); (2) a summer offshore assemblage (e.g. Makaira nigricans, T. atlanticus, Ruvettus pretiosus, Lampris guttatus); (3) an ubiquitous assemblage (e.g. K. pelamis, Coryphaena hippurus, Xiphias gladius); and (4) a spring/winter assemblage that was widely dispersed in space (e.g. trachipterids). The primary environmental factors associated with these assemblages were sea-surface temperature (highest in summer-early fall), day length (highest in early summer), thermocline depth (shallowest on the Florida side) and fluorescence (highest on the Florida side). Overall, the results of this study provide insights into how a remarkable diversity of pelagic species

  18. THE NATURE OF FOSSIL GALAXY GROUPS: ARE THEY REALLY FOSSILS?

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    La Barbera, F.; Sorrentino, G.; De Carvalho, R. R.

    We use SDSS-DR4 photometric and spectroscopic data out to redshift z {approx} 0.1 combined with ROSAT All Sky Survey X-ray data to produce a sample of 25 fossil groups (FGs), defined as bound systems dominated by a single, luminous elliptical galaxy with extended X-ray emission. We examine possible biases introduced by varying the parameters used to define the sample, and the main pitfalls are also discussed. The spatial density of FGs, estimated via the V/V {sub MAX} test, is 2.83 x 10{sup -6} h {sup 3} {sub 75} Mpc{sup -3} for L{sub X} > 0.89 x 10{sup 42} h {supmore » -2} {sub 75} erg s{sup -1} consistent with Vikhlinin et al., who examined an X-ray overluminous elliptical galaxy sample (OLEG). We compare the general properties of FGs identified here with a sample of bright field ellipticals generated from the same data set. These two samples show no differences in the distribution of neighboring faint galaxy density excess, distance from the red sequence in the color-magnitude diagram, and structural parameters such as a {sub 4} and internal color gradients. Furthermore, examination of stellar populations shows that our 25 FGs have similar ages, metallicities, and {alpha}-enhancement as the bright field ellipticals, undermining the idea that these systems represent fossils of a physical mechanism that occurred at high redshift. Our study reveals no difference between FGs and field ellipticals, suggesting that FGs might not be a distinct family of true fossils, but rather the final stage of mass assembly in the universe.« less

  19. Fossil Cores In The Kepler Data

    NASA Astrophysics Data System (ADS)

    Jackson, Brian

    Most gas giant exoplanets with orbital periods < few days are unstable against tidal decay and may be tidally disrupted before their host stars leave the main sequence. These gas giants probably contain rocky/icy cores, and so their cores will be stranded near their progenitor's Roche limit (few hours orbital period). These fossil cores will evade the Kepler mission's transit search because it is focused on periods > 0.5 days, but finding these fossil cores would provide unprecedented insights into planetary interiors and formation ? e.g., they would be a smoking gun favoring formation of gas giants via core accretion. We propose to search for and characterize fossil cores in the Kepler dataset. We will vet candidates using the Kepler photometry and auxiliary data, collect ground-based spectra of the host stars and radial-velocity (RV) and adaptive optics (AO) data to corroborate candidates. We will also constrain stellar tidal dissipation efficiencies (parameterized by Q) by determining our survey's completeness, elucidating dynamical origins and evolution of exoplanets even if we find no fossil cores. Our preliminary search has already found several dozen candidates, so the proposed survey has a high likelihood of success.

  20. Fossil Crustaceans as Parasites and Hosts.

    PubMed

    Klompmaker, Adiël A; Boxshall, Geoff A

    2015-01-01

    Numerous crustacean lineages have independently moved into parasitism as a mode of life. In modern marine ecosystems, parasitic crustaceans use representatives from many metazoan phyla as hosts. Crustaceans also serve as hosts to a rich diversity of parasites, including other crustaceans. Here, we show that the fossil record of such parasitic interactions is sparse, with only 11 examples, one dating back to the Cambrian. This may be due to the limited preservation potential and small size of parasites, as well as to problems with ascribing traces to parasitism with certainty, and to a lack of targeted research. Although the confirmed stratigraphic ranges are limited for nearly every example, evidence of parasitism related to crustaceans has become increasingly more complete for isopod-induced swellings in decapods so that quantitative analyses can be carried out. Little attention has yet been paid to the origin of parasitism in deep time, but insight can be generated by integrating data on fossils with molecular studies on modern parasites. In addition, there are other traces left by parasites that could fossilize, but have not yet been recognized in the fossil record. Copyright © 2015 Elsevier Ltd. All rights reserved.

  1. Fossil Energy organization restructured

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Not Available

    The Department of Energy has restructured its fossil energy organization to accommodate increases in activity and visibility of the President's $2.5 billion clean coal technology initiative. The realignment also includes changes in the coal research and development program and in supporting staff functions. In the coal program, changes in the organization include the establishment of two associate deputy assistant secretaries, both reporting to the deputy Assistant Secretary for Coal Technology. One associate deputy assistant secretary will oversee the Clean Coal Technology Program. A second associate deputy assistant secretary will manage the coal research and development program. An organizational chart illustratesmore » the new fossil energy headquarters organization.« less

  2. The largest fossil rodent

    PubMed Central

    Rinderknecht, Andrés; Blanco, R. Ernesto

    2008-01-01

    The discovery of an exceptionally well-preserved skull permits the description of the new South American fossil species of the rodent, Josephoartigasia monesi sp. nov. (family: Dinomyidae; Rodentia: Hystricognathi: Caviomorpha). This species with estimated body mass of nearly 1000 kg is the largest yet recorded. The skull sheds new light on the anatomy of the extinct giant rodents of the Dinomyidae, which are known mostly from isolated teeth and incomplete mandible remains. The fossil derives from San José Formation, Uruguay, usually assigned to the Pliocene–Pleistocene (4–2 Myr ago), and the proposed palaeoenvironment where this rodent lived was characterized as an estuarine or deltaic system with forest communities. PMID:18198140

  3. Synthesizing and databasing fossil calibrations: divergence dating and beyond

    PubMed Central

    Ksepka, Daniel T.; Benton, Michael J.; Carrano, Matthew T.; Gandolfo, Maria A.; Head, Jason J.; Hermsen, Elizabeth J.; Joyce, Walter G.; Lamm, Kristin S.; Patané, José S. L.; Phillips, Matthew J.; Polly, P. David; Van Tuinen, Marcel; Ware, Jessica L.; Warnock, Rachel C. M.; Parham, James F.

    2011-01-01

    Divergence dating studies, which combine temporal data from the fossil record with branch length data from molecular phylogenetic trees, represent a rapidly expanding approach to understanding the history of life. National Evolutionary Synthesis Center hosted the first Fossil Calibrations Working Group (3–6 March, 2011, Durham, NC, USA), bringing together palaeontologists, molecular evolutionists and bioinformatics experts to present perspectives from disciplines that generate, model and use fossil calibration data. Presentations and discussions focused on channels for interdisciplinary collaboration, best practices for justifying, reporting and using fossil calibrations and roadblocks to synthesis of palaeontological and molecular data. Bioinformatics solutions were proposed, with the primary objective being a new database for vetted fossil calibrations with linkages to existing resources, targeted for a 2012 launch. PMID:21525049

  4. Suzaku Finds "Fossil" Fireballs from Supernovae

    NASA Image and Video Library

    2017-12-08

    Suzaku Finds "Fossil" Fireballs from Supernovae In a supernova remnant known as the Jellyfish Nebula, Suzaku detected X-rays from fully ionized silicon and sulfur -- an imprint of higher-temperature conditions immediately following the star's explosion. The nebula is about 65 light-years across. (12/30/2009) Credit: JAXA/NASA/Suzaku To learn more go to: www.nasa.gov/mission_pages/astro-e2/news/fossil-fireballs...

  5. The Ecuadorian Artisanal Fishery for Large Pelagics: Species Composition and Spatio-Temporal Dynamics

    PubMed Central

    Martínez-Ortiz, Jimmy; Aires-da-Silva, Alexandre M.; Lennert-Cody, Cleridy E.; Maunder, Mark N.

    2015-01-01

    The artisanal fisheries of Ecuador operate within one of the most dynamic and productive marine ecosystems of the world. This study investigates the catch composition of the Ecuadorian artisanal fishery for large pelagic fishes, including aspects of its spatio-temporal dynamics. The analyses of this study are based on the most extensive dataset available to date for this fishery: a total of 106,963 trip-landing inspection records collected at its five principal ports during 2008 ‒ 2012. Ecuadorian artisanal fisheries remove a substantial amount of biomass from the upper trophic-level predatory fish community of the eastern tropical Pacific Ocean. It is estimated that at least 135 thousand metric tons (mt) (about 15.5 million fish) were landed in the five principal ports during the study period. The great novelty of Ecuadorian artisanal fisheries is the “oceanic-artisanal” fleet component, which consists of mother-ship (nodriza) boats with their towed fiber-glass skiffs (fibras) operating with pelagic longlines. This fleet has fully expanded into oceanic waters as far offshore as 100°W, west of the Galapagos Archipelago. It is estimated that nodriza operations produce as much as 80% of the total catches of the artisanal fishery. The remainder is produced by independent fibras operating in inshore waters with pelagic longlines and/or surface gillnets. A multivariate regression tree analysis was used to investigate spatio-environmental effects on the nodriza fleet (n = 6,821 trips). The catch species composition of the nodriza fleet is strongly influenced by the northwesterly circulation of the Humboldt Current along the coast of Peru and its associated cold waters masses. The target species and longline gear-type used by nodrizas change seasonally with the incursion of cool waters (< 25°C) from the south and offshore. During this season, dolphinfish (Coryphaena hippurus) dominates the catches. However, in warmer waters, the fishery changes to tuna

  6. The Ecuadorian Artisanal Fishery for Large Pelagics: Species Composition and Spatio-Temporal Dynamics.

    PubMed

    Martínez-Ortiz, Jimmy; Aires-da-Silva, Alexandre M; Lennert-Cody, Cleridy E; Maunder, Mark N

    2015-01-01

    The artisanal fisheries of Ecuador operate within one of the most dynamic and productive marine ecosystems of the world. This study investigates the catch composition of the Ecuadorian artisanal fishery for large pelagic fishes, including aspects of its spatio-temporal dynamics. The analyses of this study are based on the most extensive dataset available to date for this fishery: a total of 106,963 trip-landing inspection records collected at its five principal ports during 2008 ‒ 2012. Ecuadorian artisanal fisheries remove a substantial amount of biomass from the upper trophic-level predatory fish community of the eastern tropical Pacific Ocean. It is estimated that at least 135 thousand metric tons (mt) (about 15.5 million fish) were landed in the five principal ports during the study period. The great novelty of Ecuadorian artisanal fisheries is the "oceanic-artisanal" fleet component, which consists of mother-ship (nodriza) boats with their towed fiber-glass skiffs (fibras) operating with pelagic longlines. This fleet has fully expanded into oceanic waters as far offshore as 100°W, west of the Galapagos Archipelago. It is estimated that nodriza operations produce as much as 80% of the total catches of the artisanal fishery. The remainder is produced by independent fibras operating in inshore waters with pelagic longlines and/or surface gillnets. A multivariate regression tree analysis was used to investigate spatio-environmental effects on the nodriza fleet (n = 6,821 trips). The catch species composition of the nodriza fleet is strongly influenced by the northwesterly circulation of the Humboldt Current along the coast of Peru and its associated cold waters masses. The target species and longline gear-type used by nodrizas change seasonally with the incursion of cool waters (< 25°C) from the south and offshore. During this season, dolphinfish (Coryphaena hippurus) dominates the catches. However, in warmer waters, the fishery changes to tuna

  7. Exploring the chemical composition of pelagic tar collected in the North Atlantic Ocean

    NASA Astrophysics Data System (ADS)

    Green, H. S.; Reddy, C. M.; Valentine, D. L.; Aeppli, C.; Swarthout, B.; Sharpless, C.; Joyce, P.; Meyer, A. W.; Fuller, S. A.; Gosselin, K.

    2016-12-01

    Pelagic tarballs have been linked to multiple sources and their abundances follow notable historical and geographic trends. An overwhelming number of studies point to operational discharges (cargo washing) as the main source of pelagic tar. In a recent review article, Warnock et al. (2015) summarized that the abundance of tar balls has decreased over the last 30 years. The decreasing trend of tarballs has been attributed to the MARPOL 73/78 Annex I legislation, which was created from conventions held in 1973 and again in 1978 to respond to several tanker accidents and other pollution-related inputs. Two of the studies supporting the "MARPOL 73/78 effect" were based on the historical record of tarballs collected in the North Atlantic Ocean by the Sea Education Association (SEA; Woods Hole, MA). To supplement the SEA record, we performed a series of geochemical analyses on 100 of the SEA samples collected from 1988 to 2014. Bulk and gas chromatographic (GC) analyses revealed that the samples were highly variable. For example, the amount of material that could be dissolved in organic solvent but not measured by gas chromatography (referred to as the % GC amenable, a proxy on the distribution of compound classes that compose the tar) ranged from 10 to 80%, although skewed to values less than 40%. Another parameter, based on the GC data, was the perecentage of the resolved relative to the unresolved signal spanned from 0.1 to 1.8. Nine of the sampes would not dissolve in organic solvents and appear to be soot or coal. This study has an operation limitation as we choose to only examine samples > 1 cm (relative to samples smaller than 1mm). Our approach was based on the assumption that these samples were the most unlikely to be weathered and hence retain the genetic features of the initially released tar. While this study does not have the capacity to test confidently the MARPOL 73/78 effect, it does show that pelagic tars are highly variable, which in turn, have

  8. Magnetic resonance spectroscopy and imaging for the study of fossils.

    PubMed

    Giovannetti, Giulio; Guerrini, Andrea; Salvadori, Piero A

    2016-07-01

    Computed tomography (CT) has long been used for investigating palaeontological specimens, as it is a nondestructive technique which avoids the need to dissolve or ionize the fossil sample. However, magnetic resonance spectroscopy (MRS) and magnetic resonance imaging (MRI) have recently gained ground as analytical tools for examination of palaeontological samples, by nondestructively providing information about the structure and composition of fossils. While MRI techniques are able to reveal the three-dimensional geometry of the trace fossil, MRS can provide information on the chemical composition of the samples. The multidimensional nature of MR (magnetic resonance) signals has potential to provide rich three-dimensional data on the palaeontological specimens and also to help in elucidating paleopathological and paleoecological questions. In this work the verified applications and the emerging uses of MRI and MRS in paleontology are reviewed, with particular attention to fossil spores, fossil plants, ambers, fossil invertebrates, and fossil vertebrate studies. Copyright © 2016 Elsevier Inc. All rights reserved.

  9. The fossil record of the sixth extinction.

    PubMed

    Plotnick, Roy E; Smith, Felisa A; Lyons, S Kathleen

    2016-05-01

    Comparing the magnitude of the current biodiversity crisis with those in the fossil record is difficult without an understanding of differential preservation. Integrating data from palaeontological databases with information on IUCN status, ecology and life history characteristics of contemporary mammals, we demonstrate that only a small and biased fraction of threatened species (< 9%) have a fossil record, compared with 20% of non-threatened species. We find strong taphonomic biases related to body size and geographic range. Modern species with a fossil record tend to be large and widespread and were described in the 19(th) century. The expected magnitude of the current extinction based only on species with a fossil record is about half of that of one based on all modern species; values for genera are similar. The record of ancient extinctions may be similarly biased, with many species having originated and gone extinct without leaving a tangible record. © 2016 John Wiley & Sons Ltd/CNRS.

  10. Coastal Microstructure: From Active Overturn to Fossil Turbulence

    NASA Astrophysics Data System (ADS)

    Tau Leung, Pak

    2011-11-01

    The Remote Anthropogenic Sensing Program was a five year effort (2001- 2005) to examine subsurface phenomena related to a sewage outfall off the coast of Oahu, Hawaii. This research has implications for basic ocean hydrodynamics, particularly for a greatly improved understanding of the evolution of turbulent patches. It was the first time a microstructure measurement was used to study such a buoyancy-driven turbulence generated by a sea-floor diffuser. In 2004, two stations were selected to represent the near field and ambient conditions. They have nearly identical bathymetrical and hydrographical features and provide an ideal environment for a control experiment. Repeated vertical microstructure measurements were performed at both stations for 20 days. A time series of physical parameters was collected and used for statistical analysis. After comparing the data from both stations, it can be concluded that the turbulent mixing generated by the diffuser contributes to the elevated dissipation rate observed in the pycnocline and bottom boundary layer. To further understand the mixing processes in both regions, data were plotted on a Hydrodynamic Phase Diagram. The overturning stages of the turbulent patches are identified by Hydrodynamic Phase Diagram. This technique provides detailed information on the evolution of the turbulent patches from active overturns to fossilized scalar microstructures in the water column. Results from this study offer new evidence to support the fossil turbulence theory. This study concluded that: 1. Field Data collected near a sea-floor outfall diffuser show that turbulent patches evolve from active (overturning) to fossil (buoyancy-inhibited) stages, consistent with the process of turbulent patch evolution proposed by fossil turbulence theory. 2. The data show that active (overturning) and fossil (buoyancy-inhibited) patches have smaller length scales than the active+fossil (intermediate) stage of patch evolution, consistent with fossil

  11. Are the oldest 'fossils', fossils

    NASA Technical Reports Server (NTRS)

    Schopf, J. W.

    1976-01-01

    A comparative statistical study has been carried out on populations of modern algae, Precambrian algal microfossils, the 'organized elements' of the Orgueil carbonaceous meteorite, and the oldest microfossil-like objects now known (spheroidal bodies from the Fig Tree and Onverwacht Groups of the Swaziland Supergroup, South Africa). The distribution patterns exhibited by the more than 3000 m.y.-old Swaziland microstructures bear considerable resemblance to those of the abiotic 'organized elements' but differ rather markedly from those exhibited by younger, assuredly biogenic, populations. Based on these comparisons, it is concluded that the Swaziland spheroids could be, at least in part, of nonbiologic origin; these oldest known fossil-like microstructures should not be regarded as constituting firm evidence of Archean life.

  12. Contemporaneous trace and body fossils from a late Pleistocene Lakebed in Victoria, Australia, allow assessment of bias in the fossil record.

    PubMed

    Camens, Aaron Bruce; Carey, Stephen Paul

    2013-01-01

    The co-occurrence of vertebrate trace and body fossils within a single geological formation is rare and the probability of these parallel records being contemporaneous (i.e. on or near the same bedding plane) is extremely low. We report here a late Pleistocene locality from the Victorian Volcanic Plains in south-eastern Australia in which demonstrably contemporaneous, but independently accumulated vertebrate trace and body fossils occur. Bite marks from a variety of taxa are also present on the bones. This site provides a unique opportunity to examine the biases of these divergent fossil records (skeletal, footprints and bite marks) that sampled a single fauna. The skeletal record produced the most complete fauna, with the footprint record indicating a markedly different faunal composition with less diversity and the feeding traces suggesting the presence, amongst others, of a predator not represented by either the skeletal or footprint records. We found that the large extinct marsupial predator Thylacoleo was the only taxon apparently represented by all three records, suggesting that the behavioral characteristics of large carnivores may increase the likelihood of their presence being detected within a fossil fauna. In contrast, Diprotodon (the largest-ever marsupial) was represented only by trace fossils at this site and was absent from the site's skeletal record, despite its being a common and easily detected presence in late Pleistocene skeletal fossil faunas elsewhere in Australia. Small mammals absent from the footprint record for the site were represented by skeletal fossils and bite marks on bones.

  13. Contemporaneous Trace and Body Fossils from a Late Pleistocene Lakebed in Victoria, Australia, Allow Assessment of Bias in the Fossil Record

    PubMed Central

    Camens, Aaron Bruce; Carey, Stephen Paul

    2013-01-01

    The co-occurrence of vertebrate trace and body fossils within a single geological formation is rare and the probability of these parallel records being contemporaneous (i.e. on or near the same bedding plane) is extremely low. We report here a late Pleistocene locality from the Victorian Volcanic Plains in south-eastern Australia in which demonstrably contemporaneous, but independently accumulated vertebrate trace and body fossils occur. Bite marks from a variety of taxa are also present on the bones. This site provides a unique opportunity to examine the biases of these divergent fossil records (skeletal, footprints and bite marks) that sampled a single fauna. The skeletal record produced the most complete fauna, with the footprint record indicating a markedly different faunal composition with less diversity and the feeding traces suggesting the presence, amongst others, of a predator not represented by either the skeletal or footprint records. We found that the large extinct marsupial predator Thylacoleo was the only taxon apparently represented by all three records, suggesting that the behavioral characteristics of large carnivores may increase the likelihood of their presence being detected within a fossil fauna. In contrast, Diprotodon (the largest-ever marsupial) was represented only by trace fossils at this site and was absent from the site's skeletal record, despite its being a common and easily detected presence in late Pleistocene skeletal fossil faunas elsewhere in Australia. Small mammals absent from the footprint record for the site were represented by skeletal fossils and bite marks on bones. PMID:23301008

  14. Benthic-Pelagic Coupling in Biogeochemical and Climate Models: Existing Approaches, Recent developments and Roadblocks

    NASA Astrophysics Data System (ADS)

    Arndt, Sandra

    2016-04-01

    Marine sediments are key components in the Earth System. They host the largest carbon reservoir on Earth, provide the only long term sink for atmospheric CO2, recycle nutrients and represent the most important climate archive. Biogeochemical processes in marine sediments are thus essential for our understanding of the global biogeochemical cycles and climate. They are first and foremost, donor controlled and, thus, driven by the rain of particulate material from the euphotic zone and influenced by the overlying bottom water. Geochemical species may undergo several recycling loops (e.g. authigenic mineral precipitation/dissolution) before they are either buried or diffuse back to the water column. The tightly coupled and complex pelagic and benthic process interplay thus delays recycling flux, significantly modifies the depositional signal and controls the long-term removal of carbon from the ocean-atmosphere system. Despite the importance of this mutual interaction, coupled regional/global biogeochemical models and (paleo)climate models, which are designed to assess and quantify the transformations and fluxes of carbon and nutrients and evaluate their response to past and future perturbations of the climate system either completely neglect marine sediments or incorporate a highly simplified representation of benthic processes. On the other end of the spectrum, coupled, multi-component state-of-the-art early diagenetic models have been successfully developed and applied over the past decades to reproduce observations and quantify sediment-water exchange fluxes, but cannot easily be coupled to pelagic models. The primary constraint here is the high computation cost of simulating all of the essential redox and equilibrium reactions within marine sediments that control carbon burial and benthic recycling fluxes: a barrier that is easily exacerbated if a variety of benthic environments are to be spatially resolved. This presentation provides an integrative overview of

  15. The evolution of jaw protrusion mechanics is tightly coupled to bentho-pelagic divergence in damselfishes (Pomacentridae).

    PubMed

    Cooper, W James; Carter, Casey B; Conith, Andrew J; Rice, Aaron N; Westneat, Mark W

    2017-02-15

    Most species-rich lineages of aquatic organisms have undergone divergence between forms that feed from the substrate (benthic feeding) and forms that feed from the water column (pelagic feeding). Changes in trophic niche are frequently accompanied by changes in skull mechanics, and multiple fish lineages have evolved highly specialized biomechanical configurations that allow them to protrude their upper jaws toward the prey during feeding. Damselfishes (family Pomacentridae) are an example of a species-rich lineage with multiple trophic morphologies and feeding ecologies. We sought to determine whether bentho-pelagic divergence in the damselfishes is tightly coupled to changes in jaw protrusion ability. Using high-speed video recordings and kinematic analysis, we examined feeding performance in 10 species that include three examples of convergence on herbivory, three examples of convergence on omnivory and two examples of convergence on planktivory. We also utilized morphometrics to characterize the feeding morphology of an additional 40 species that represent all 29 damselfish genera. Comparative phylogenetic analyses were then used to examine the evolution of trophic morphology and biomechanical performance. We find that pelagic-feeding damselfishes (planktivores) are strongly differentiated from extensively benthic-feeding species (omnivores and herbivores) by their jaw protrusion ability, upper jaw morphology and the functional integration of upper jaw protrusion with lower jaw abduction. Most aspects of cranial form and function that separate these two ecological groups have evolved in correlation with each other and the evolution of the functional morphology of feeding in damselfishes has involved repeated convergence in form, function and ecology. © 2017. Published by The Company of Biologists Ltd.

  16. WHERE ARE THE FOSSILS OF THE FIRST GALAXIES? II. TRUE FOSSILS, GHOST HALOS, AND THE MISSING BRIGHT SATELLITES

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Bovill, Mia S.; Ricotti, Massimo, E-mail: msbovill@astro.umd.edu

    We use a new set of cold dark matter simulations of the local universe to investigate the distribution of fossils of primordial dwarf galaxies within and around the Milky Way. Throughout, we build upon previous results showing agreement between the observed stellar properties of a subset of the ultra-faint dwarfs and our simulated fossils. Here, we show that fossils of the first galaxies have galactocentric distributions and cumulative luminosity functions consistent with observations. In our model, we predict {approx}300 luminous satellites orbiting the Milky Way, 50%-70% of which are well-preserved fossils. Within the Milky Way virial radius, the majority ofmore » these fossils have luminosities L{sub V} < 10{sup 6} L{sub sun}. Despite our multidimensional agreement with observations at low masses and luminosities, the primordial model produces an overabundance of bright dwarf satellites (L{sub V} > 10{sup 4} L{sub sun}) with respect to observations where observations are nearly complete. The 'bright satellite problem' is most evident in the outer parts of the Milky Way. We estimate that, although relatively bright, the primordial stellar populations are very diffuse, producing a population with surface brightnesses below surveys' detection limits, and are easily stripped by tidal forces. Although we cannot yet present unmistakable evidence for the existence of the fossils of first galaxies in the Local Group, the results of our studies suggest observational strategies that may demonstrate their existence: (1) the detection of 'ghost halos' of primordial stars around isolated dwarfs would prove that stars formed in minihalos (M < 10{sup 8} M{sub sun}) before reionization and strongly suggest that at least a fraction of the ultra-faint dwarfs are fossils of the first galaxies; and (2) the existence of a yet unknown population of {approx}150 Milky Way ultra-faints with half-light radii r{sub hl} {approx} 100-1000 pc and luminosities L{sub V} < 10{sup 4} L{sub sun

  17. Forty Years Later: Updating the Fossilization Hypothesis

    ERIC Educational Resources Information Center

    Han, ZhaoHong

    2013-01-01

    A founding concept in second language acquisition (SLA) research, fossilization has been fundamental to understanding second language (L2) development. The Fossilization Hypothesis, introduced in Selinker's seminal text (1972), has thus been one of the most influential theories, guiding a significant bulk of SLA research for four decades; 2012…

  18. Where to Dig for Fossils: Combining Climate-Envelope, Taphonomy and Discovery Models

    PubMed Central

    Block, Sebastián; Saltré, Frédérik; Rodríguez-Rey, Marta; Fordham, Damien A.; Unkel, Ingmar; Bradshaw, Corey J. A.

    2016-01-01

    Fossils represent invaluable data to reconstruct the past history of life, yet fossil-rich sites are often rare and difficult to find. The traditional fossil-hunting approach focuses on small areas and has not yet taken advantage of modelling techniques commonly used in ecology to account for an organism’s past distributions. We propose a new method to assist finding fossils at continental scales based on modelling the past distribution of species, the geological suitability of fossil preservation and the likelihood of fossil discovery in the field, and apply it to several genera of Australian megafauna that went extinct in the Late Quaternary. Our models predicted higher fossil potentials for independent sites than for randomly selected locations (mean Kolmogorov-Smirnov statistic = 0.66). We demonstrate the utility of accounting for the distribution history of fossil taxa when trying to find the most suitable areas to look for fossils. For some genera, the probability of finding fossils based on simple climate-envelope models was higher than the probability based on models incorporating current conditions associated with fossil preservation and discovery as predictors. However, combining the outputs from climate-envelope, preservation, and discovery models resulted in the most accurate predictions of potential fossil sites at a continental scale. We proposed potential areas to discover new fossils of Diprotodon, Zygomaturus, Protemnodon, Thylacoleo, and Genyornis, and provide guidelines on how to apply our approach to assist fossil hunting in other continents and geological settings. PMID:27027874

  19. Where to Dig for Fossils: Combining Climate-Envelope, Taphonomy and Discovery Models.

    PubMed

    Block, Sebastián; Saltré, Frédérik; Rodríguez-Rey, Marta; Fordham, Damien A; Unkel, Ingmar; Bradshaw, Corey J A

    2016-01-01

    Fossils represent invaluable data to reconstruct the past history of life, yet fossil-rich sites are often rare and difficult to find. The traditional fossil-hunting approach focuses on small areas and has not yet taken advantage of modelling techniques commonly used in ecology to account for an organism's past distributions. We propose a new method to assist finding fossils at continental scales based on modelling the past distribution of species, the geological suitability of fossil preservation and the likelihood of fossil discovery in the field, and apply it to several genera of Australian megafauna that went extinct in the Late Quaternary. Our models predicted higher fossil potentials for independent sites than for randomly selected locations (mean Kolmogorov-Smirnov statistic = 0.66). We demonstrate the utility of accounting for the distribution history of fossil taxa when trying to find the most suitable areas to look for fossils. For some genera, the probability of finding fossils based on simple climate-envelope models was higher than the probability based on models incorporating current conditions associated with fossil preservation and discovery as predictors. However, combining the outputs from climate-envelope, preservation, and discovery models resulted in the most accurate predictions of potential fossil sites at a continental scale. We proposed potential areas to discover new fossils of Diprotodon, Zygomaturus, Protemnodon, Thylacoleo, and Genyornis, and provide guidelines on how to apply our approach to assist fossil hunting in other continents and geological settings.

  20. Finding fossils in new ways: an artificial neural network approach to predicting the location of productive fossil localities.

    PubMed

    Anemone, Robert; Emerson, Charles; Conroy, Glenn

    2011-01-01

    Chance and serendipity have long played a role in the location of productive fossil localities by vertebrate paleontologists and paleoanthropologists. We offer an alternative approach, informed by methods borrowed from the geographic information sciences and using recent advances in computer science, to more efficiently predict where fossil localities might be found. Our model uses an artificial neural network (ANN) that is trained to recognize the spectral characteristics of known productive localities and other land cover classes, such as forest, wetlands, and scrubland, within a study area based on the analysis of remotely sensed (RS) imagery. Using these spectral signatures, the model then classifies other pixels throughout the study area. The results of the neural network classification can be examined and further manipulated within a geographic information systems (GIS) software package. While we have developed and tested this model on fossil mammal localities in deposits of Paleocene and Eocene age in the Great Divide Basin of southwestern Wyoming, a similar analytical approach can be easily applied to fossil-bearing sedimentary deposits of any age in any part of the world. We suggest that new analytical tools and methods of the geographic sciences, including remote sensing and geographic information systems, are poised to greatly enrich paleoanthropological investigations, and that these new methods should be embraced by field workers in the search for, and geospatial analysis of, fossil primates and hominins. Copyright © 2011 Wiley-Liss, Inc.

  1. Comparative evaluation of solar, fission, fusion, and fossil energy resources. Part 4: Energy from fossil fuels

    NASA Technical Reports Server (NTRS)

    Williams, J. R.

    1974-01-01

    The conversion of fossil-fired power plants now burning oil or gas to burn coal is discussed along with the relaxation of air quality standards and the development of coal gasification processes to insure a continued supply of gas from coal. The location of oil fields, refining areas, natural gas fields, and pipelines in the U.S. is shown. The technologies of modern fossil-fired boilers and gas turbines are defined along with the new technologies of fluid-bed boilers and MHD generators.

  2. Offshore Windfarm Impact on Pelagic Primary Production in the Southern North Sea

    NASA Astrophysics Data System (ADS)

    Slavik, Kaela; Zhang, Wenyan; Lemmen, Carsten; Wirtz, Kai

    2016-04-01

    As society struggles to find solutions to mitigate global warming, the demand for renewable energy technology has increased. Especially investment in offshore wind energy has proliferated in the European Union, with projections over the next 15 years estimating an over 40 fold increase in total offshore wind electricity. Though built with the goal of reducing the environmental impacts associated with traditional energy production, the long-term ecological impacts of offshore windfarm structures is not yet well understood. The consequences are of particular importance in the southern North Sea, where the expansion of offshore windfarms is focused. Our study investigates how the gradual accumulation of epifaunal biomass on submerged substrate at offshore windfarms impacts ecosystem services in the southern North Sea. Biofouling is governed predominately by the filter feeder Mytilus edulis, which, as an ecological engineer, will further alter the surrounding benthic and pelagic environment. We reconstruct the distribution of benthic filter feeders in the SNS and generate scenarios of increased potential distribution based on available information of Mytilus edulis settlement at turbines and of turbine locations. These maps are coupled through the MOSSCO (Modular Coupling System for Shelves and Coasts) to state-of-the-art and high resolution hydrodynamic and ecosystem models. We find a substantial change in pelagic primary production as a result of additional Mytilus edulis growth at offshore windfarms.

  3. 76 FR 10778 - Fisheries of the Caribbean, Gulf of Mexico, and South Atlantic; Coastal Migratory Pelagic...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-02-28

    ... Migratory Pelagic Resources of the Gulf of Mexico and South Atlantic; Closure AGENCY: National Marine...: Temporary rule; closure. SUMMARY: NMFS closes the commercial sector for king mackerel in the Florida east coast subzone. This closure is necessary to protect the Gulf king mackerel resource. DATES: The closure...

  4. 76 FR 65662 - Fisheries of the Caribbean, Gulf of Mexico, and South Atlantic; Coastal Migratory Pelagic...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-10-24

    ... Migratory Pelagic Resources in the Gulf of Mexico and Atlantic Region; Amendment 18 AGENCY: National Marine... rule; request for comments. SUMMARY: NMFS proposes regulations to implement Amendment 18 to the Fishery..., Amendment 18 would set allocations for Atlantic cobia and establish control rules for king mackerel, Spanish...

  5. Fossil vs. non-fossil sources of fine carbonaceous aerosols in four Chinese cities during the extreme winter haze episode in 2013

    NASA Astrophysics Data System (ADS)

    Zhang, Y.-L.; Huang, R.-J.; El Haddad, I.; Ho, K.-F.; Cao, J.-J.; Han, Y.; Zotter, P.; Bozzetti, C.; Daellenbach, K. R.; Canonaco, F.; Slowik, J. G.; Salazar, G.; Schwikowski, M.; Schnelle-Kreis, J.; Abbaszade, G.; Zimmermann, R.; Baltensperger, U.; Prévôt, A. S. H.; Szidat, S.

    2014-10-01

    During winter 2013, extremely high concentrations (i.e. 4-20 times higher than the World Health Organization guideline) of PM2.5 (particulate matter with an aerodynamic diameter <2.5 μm) were reported in several large cities in China. In this work, source apportionment of fine carbonaceous aerosols during this haze episode was conducted at four major cities in China including Xian, Beijing, Shanghai and Guangzhou. An effective statistical analysis of a combined dataset from elemental carbon (EC) and organic carbon (OC), radiocarbon (14C) and biomass-burning marker measurements using Latin-hypercube sampling allowed a quantitative source apportionment of carbonaceous aerosols. We found that fossil emissions from coal combustion and vehicle exhaust dominated EC with a mean contribution of 75 ± 8% at all sites. The remaining 25 ± 8% was exclusively attributed to biomass combustion, consistent with the measurements of biomass-burning markers such as anhydrosugars (levoglucosan and mannosan) and water-soluble potassium (K+). With a combination of the levoglucosan-to-mannosan and levoglucosan-to-K+ ratios, the major source of biomass burning in winter in China is suggested to be combustion of crop residues. The contribution of fossil sources to OC was highest in Beijing (58 ± 5%) and decreased from Shanghai (49 ± 2%) to Xian (38 ± 3%) and Guangzhou (35 ± 7%). Generally, a larger fraction of fossil OC was rather from secondary origins than primary sources for all sites. Non-fossil sources accounted on average for 55 ± 10% and 48 ± 9% of OC and TC, respectively, which suggests that non-fossil emissions were very important contributors of urban carbonaceous aerosols in China. The primary biomass-burning emissions accounted for 40 ± 8%, 48 ± 18%, 53 ± 4% and 65 ± 26% of non-fossil OC for Xian, Beijing, Shanghai and Guangzhou, respectively. Other non-fossil sources excluding primary biomass-burning were mainly attributed to formation of secondary organic carbon

  6. Fossil vs. non-fossil sources of fine carbonaceous aerosols in four Chinese cities during the extreme winter haze episode of 2013

    NASA Astrophysics Data System (ADS)

    Zhang, Y.-L.; Huang, R.-J.; El Haddad, I.; Ho, K.-F.; Cao, J.-J.; Han, Y.; Zotter, P.; Bozzetti, C.; Daellenbach, K. R.; Canonaco, F.; Slowik, J. G.; Salazar, G.; Schwikowski, M.; Schnelle-Kreis, J.; Abbaszade, G.; Zimmermann, R.; Baltensperger, U.; Prévôt, A. S. H.; Szidat, S.

    2015-02-01

    During winter 2013, extremely high concentrations (i.e., 4-20 times higher than the World Health Organization guideline) of PM2.5 (particulate matter with an aerodynamic diameter < 2.5 μm) mass concentrations (24 h samples) were found in four major cities in China including Xi'an, Beijing, Shanghai and Guangzhou. Statistical analysis of a combined data set from elemental carbon (EC), organic carbon (OC), 14C and biomass-burning marker measurements using Latin hypercube sampling allowed a quantitative source apportionment of carbonaceous aerosols. Based on 14C measurements of EC fractions (six samples each city), we found that fossil emissions from coal combustion and vehicle exhaust dominated EC with a mean contribution of 75 ± 8% across all sites. The remaining 25 ± 8% was exclusively attributed to biomass combustion, consistent with the measurements of biomass-burning markers such as anhydrosugars (levoglucosan and mannosan) and water-soluble potassium (K+). With a combination of the levoglucosan-to-mannosan and levoglucosan-to-K+ ratios, the major source of biomass burning in winter in China is suggested to be combustion of crop residues. The contribution of fossil sources to OC was highest in Beijing (58 ± 5%) and decreased from Shanghai (49 ± 2%) to Xi'an (38 ± 3%) and Guangzhou (35 ± 7%). Generally, a larger fraction of fossil OC was from secondary origins than primary sources for all sites. Non-fossil sources accounted on average for 55 ± 10 and 48 ± 9% of OC and total carbon (TC), respectively, which suggests that non-fossil emissions were very important contributors of urban carbonaceous aerosols in China. The primary biomass-burning emissions accounted for 40 ± 8, 48 ± 18, 53 ± 4 and 65 ± 26% of non-fossil OC for Xi'an, Beijing, Shanghai and Guangzhou, respectively. Other non-fossil sources excluding primary biomass burning were mainly attributed to formation of secondary organic carbon (SOC) from non-fossil precursors such as biomass

  7. Occurrence of microplastics in the gastrointestinal tract of pelagic and demersal fish from the English Channel.

    PubMed

    Lusher, A L; McHugh, M; Thompson, R C

    2013-02-15

    Microplastics are present in marine habitats worldwide and laboratory studies show this material can be ingested, yet data on abundance in natural populations is limited. This study documents microplastics in 10 species of fish from the English Channel. 504 Fish were examined and plastics found in the gastrointestinal tracts of 36.5%. All five pelagic species and all five demersal species had ingested plastic. Of the 184 fish that had ingested plastic the average number of pieces per fish was 1.90±0.10. A total of 351 pieces of plastic were identified using FT-IR Spectroscopy; polyamide (35.6%) and the semi-synthetic cellulosic material, rayon (57.8%) were most common. There was no significant difference between the abundance of plastic ingested by pelagic and demersal fish. Hence, microplastic ingestion appears to be common, in relatively small quantities, across a range of fish species irrespective of feeding habitat. Further work is needed to establish the potential consequences. Copyright © 2012 Elsevier Ltd. All rights reserved.

  8. Studies on food organisms of pelagic fishes as revealed by the 1979 North Atlantic Eel Expedition

    NASA Astrophysics Data System (ADS)

    Appelbaum, S.

    1982-09-01

    The extent to which pelagic fishes occurring in the Sargasso Sea and adjacent parts of the Atlantic prey on leptocephali (Anguilliformes) was investigated. Most of the fishes examined (c. 95%) were collected using a commercial pelagical trawl. The stomach contents of about 1000 fishes (25 species of 10 families), mostly belonging to the suborders Myctophoidei, Stomiatoidei and the order Anguilliformes, were examined. The remains of invertebrates, mainly crustaceans, molluscs, tunicates, chaetognaths, and siphonophores were found in 28.8 % of the stomachs. Fishes, mostly myctophids or fish remains, were observed in 11.2 % of the stomachs; 18.7 % contained unidentified items and 40.6 % were empty. Leptocephali ( Ariosoma spp. and Gnathophis sp.) were found in the alimentary tract of 0.5 % of the fishes examined, exclusively represented by the myctophid, Ceratoscopelus warmingii. This report indicates that the Sargasso Sea population of Anguilla leptocephali, economically the most important eel, is not seriously affected by predation of oceanic fish species considered in this study.

  9. Giving the early fossil record of sponges a squeeze.

    PubMed

    Antcliffe, Jonathan B; Callow, Richard H T; Brasier, Martin D

    2014-11-01

    Twenty candidate fossils with claim to be the oldest representative of the Phylum Porifera have been re-analysed. Three criteria are used to assess each candidate: (i) the diagnostic criteria needed to categorize sponges in the fossil record; (ii) the presence, or absence, of such diagnostic features in the putative poriferan fossils; and (iii) the age constraints for the candidate fossils. All three criteria are critical to the correct interpretation of any fossil and its placement within an evolutionary context. Our analysis shows that no Precambrian fossil candidate yet satisfies all three of these criteria to be a reliable sponge fossil. The oldest widely accepted candidate, Mongolian silica hexacts from c. 545 million years ago (Ma), are here shown to be cruciform arsenopyrite crystals. The oldest reliable sponge remains are siliceous spicules from the basal Cambrian (Protohertzina anabarica Zone) Soltanieh Formation, Iran, which are described and analysed here in detail for the first time. Extensive archaeocyathan sponge reefs emerge and radiate as late as the middle of the Fortunian Stage of the Cambrian and demonstrate a gradual assembly of their skeletal structure through this time coincident with the evolution of other metazoan groups. Since the Porifera are basal in the Metazoa, their presence within the late Proterozoic has been widely anticipated. Molecular clock calibration for the earliest Porifera and Metazoa should now be based on the Iranian hexactinellid material dated to c. 535 Ma. The earliest convincing fossil sponge remains appeared at around the time of the Precambrian-Cambrian boundary, associated with the great radiation events of that interval. © 2014 The Authors. Biological Reviews © 2014 Cambridge Philosophical Society.

  10. Internal waves and modern and ancient hiatuses in pelagic caps of Pacific guyots and seamounts

    NASA Astrophysics Data System (ADS)

    Mitchell, Neil; Simmons, Harper; Lear, Carrie

    2013-04-01

    Locations of recent non-deposition and ancient hiatuses in the pelagic caps of guyots and seamounts are compared with paleotemperature and physiographic information to speculate on the character of internal tidal waves in the upper Pacific Ocean through the Cenozoic. Internal tidal waves are generated where the ocean barotropic tide passes over the Hawaiian and other major ridges in the Pacific basin. Drill core and geophysical evidence for sediment accumulation, non-deposition or erosion are used to classify broadly sites as either accumulating or eroding/non-depositing in the recent geological past. When these classified sites are compared against results of a numerical model of the internal tide field (Simmons, Ocean Mod. 2008), the sites accumulating particles over the past few million years are all found to lie away from beams of the modeled internal tide, while those that have not been accumulating are in areas of high internal wave energy. Given the correspondence to modern internal wave conditions, we examine whether internal tides can explain ancient hiatuses at the drill sites. For example, Late Cenozoic pelagic caps on guyots among the Marshall Islands contain two hiatuses of broadly similar age, but the dates of the first pelagic sediments deposited following each hiatus do not correlate between guyots, suggesting that they originate not from universal factors (e.g., water chemistry) but local, probably physical factors, such as internal tides. We investigate how changing boundary conditions such as ocean temperature and basin physiography may have affected the geometry and vigour of internal tides through the Cenozoic. Changes in the geometry of ridges underlying the Solomon, Bonin and Marianas Island chains caused by plate tectonics and subsidence may be responsible for sediment hiatuses at these far-field guyot sites.

  11. Internal waves and modern and ancient hiatuses in pelagic caps of Pacific guyots and seamounts

    NASA Astrophysics Data System (ADS)

    Mitchell, N. C.; Simmons, H. L.; Lear, C. H.

    2012-12-01

    Locations of recent non-deposition and ancient hiatuses in the pelagic caps of guyots and seamounts are compared with paleotemperature and physiographic information to speculate on the character of internal tidal waves in the upper Pacific Ocean through the Cenozoic. Internal tidal waves are generated where the ocean barotopic tide passes over the Hawaiian and other major ridges in the Pacific basin. Drill core and geophysical evidence for sediment accumulation, non-deposition or erosion are used to classify broadly sites as either accumulating or eroding/non-depositing in the recent geological past. When these classified sites are compared against results of a numerical model of the internal tide field (Simmons, Ocean Mod. 2008), the sites accumulating particles over the past few million years are all found to lie away from beams of the modeled internal tide, while those that have not been accumulating are in areas of high internal wave energy. Given the correspondence to modern internal wave conditions, we examine whether internal tides can explain ancient hiatuses at the drill sites. For example, Late Cenozoic pelagic caps on guyots among the Marshall Islands contain two hiatuses of broadly similar age, but the dates of the first pelagic sediments deposited following each hiatus do not correlate between guyots, suggesting that they originate not from universal factors (e.g., water chemistry) but local, probably physical factors, such as internal tides. We investigate how changing boundary conditions such as ocean temperature and basin physiography may have affected the geometry and vigour of internal tides through the Cenozoic. Changes in the geometry of ridges underlying the Solomon, Bonin and Marianas Island chains caused by plate tectonics and subsidence may be responsible for sediment hiatuses at these far-field guyot sites.

  12. Impact of biology knowledge on the conservation and management of large pelagic sharks.

    PubMed

    Yokoi, Hiroki; Ijima, Hirotaka; Ohshimo, Seiji; Yokawa, Kotaro

    2017-09-06

    Population growth rate, which depends on several biological parameters, is valuable information for the conservation and management of pelagic sharks, such as blue and shortfin mako sharks. However, reported biological parameters for estimating the population growth rates of these sharks differ by sex and display large variability. To estimate the appropriate population growth rate and clarify relationships between growth rate and relevant biological parameters, we developed a two-sex age-structured matrix population model and estimated the population growth rate using combinations of biological parameters. We addressed elasticity analysis and clarified the population growth rate sensitivity. For the blue shark, the estimated median population growth rate was 0.384 with a range of minimum and maximum values of 0.195-0.533, whereas those values of the shortfin mako shark were 0.102 and 0.007-0.318, respectively. The maturity age of male sharks had the largest impact for blue sharks, whereas that of female sharks had the largest impact for shortfin mako sharks. Hypotheses for the survival process of sharks also had a large impact on the population growth rate estimation. Both shark maturity age and survival rate were based on ageing validation data, indicating the importance of validating the quality of these data for the conservation and management of large pelagic sharks.

  13. Energy properties of solid fossil fuels and solid biofuels

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Holubcik, Michal, E-mail: michal.holubcik@fstroj.uniza.sk; Jandacka, Jozef, E-mail: jozef.jandacka@fstroj.uniza.sk; Kolkova, Zuzana, E-mail: zuzana.kolkova@rc.uniza.sk

    The paper deals about the problematic of energy properties of solid biofuels in comparison with solid fossil fuels. Biofuels are alternative to fossil fuels and their properties are very similar. During the experiments were done in detail experiments to obtain various properties of spruce wood pellets and wheat straw pellets like biofuels in comparison with brown coal and black coal like fossil fuels. There were tested moisture content, volatile content, fixed carbon content, ash content, elementary analysis (C, H, N, S content) and ash fusion temperatures. The results show that biofuels have some advantages and also disadvantages in comparison withmore » solid fossil fuels.« less

  14. Eumetazoan fossils in terminal Proterozoic phosphorites?

    PubMed Central

    Xiao, Shuhai; Yuan, Xunlai; Knoll, Andrew H.

    2000-01-01

    Phosphatic sedimentary rocks preserve a record of early animal life different from and complementary to that provided by Ediacaran fossils in terminal Proterozoic sandstones and shales. Phosphorites of the Doushantuo Formation, South China, contain eggs, egg cases, and stereoblastulae that document animals of unspecified phylogenetic position; small fossils containing putative spicules may specifically record the presence of sponges. Microfossils recently interpreted as the preserved gastrulae of cnidarian and bilaterian metazoans can alternatively be interpreted as conventional algal cysts and/or egg cases modified by diagenetic processes known to have had a pervasive influence on Doushantuo phosphorites. Regardless of this interpretation, evidence for Doushantuo eumetazoans is provided by millimeter-scale tubes that display tabulation and apical budding characteristic of some Cnidaria, especially the extinct tabulates. Like some Ediacaran remains, these small, benthic, colonial fossils may represent stem-group eumetazoans or stem-group cnidarians that lived in the late Proterozoic ocean. PMID:11095754

  15. Potential for travertine formation: Fossil Creek, Arizona

    Treesearch

    John Malusa; Steven T. Overby; Roderic A. Parnell

    2003-01-01

    Chemical analyses of water emanating from Fossil Springs in Central Arizona were conducted to predict changes in travertine deposition related to changes in stream discharge caused by diversion for hydroelectric power generation. During spring of 1996, water was sampled at 15 locations during normal seepage flow in a 6.7 km reach below Fossil Springs and at full...

  16. Source and sink of fluid in pelagic siliceous sediments along a cold subduction plate boundary

    NASA Astrophysics Data System (ADS)

    Yamaguchi, Asuka; Hina, Shoko; Hamada, Yohei; Kameda, Jun; Hamahashi, Mari; Kuwatani, Tatsu; Shimizu, Mayuko; Kimura, Gaku

    2016-08-01

    Subduction zones where old oceanic plate underthrusting occurs are characterized by thick pelagic sediments originating from planktonic ooze as well as cold thermal conditions. For a better understanding of dehydration from pelagic sediments and fluid behavior, which would play a key role in controlling the dynamics in the shallow portion of the subduction zone, as observed in the 2011 Tohoku earthquake and tsunami, we investigate cherts in a Jurassic accretionary complex in Japan. The microstructure and microchemistry of these cherts indicate dissolution of SiO2 from a pressure solution seam and precipitation of SiO2 to the ;white chert layer,; which would act as a fluid conduit. The amount of water necessary to precipitate SiO2 in the white chert is 102 times larger than that produced by compaction and silica/clay diagenesis. Other fluid sources, such as hydrated oceanic crust or oceanic mantle, are necessary to account for this discrepancy in the fluid budget. A large amount of external fluid likely contributed to rising pore pressure along cold plate boundaries.

  17. Genome sequence, population history, and pelage genetics of the endangered African wild dog (Lycaon pictus).

    PubMed

    Campana, Michael G; Parker, Lillian D; Hawkins, Melissa T R; Young, Hillary S; Helgen, Kristofer M; Szykman Gunther, Micaela; Woodroffe, Rosie; Maldonado, Jesús E; Fleischer, Robert C

    2016-12-09

    The African wild dog (Lycaon pictus) is an endangered African canid threatened by severe habitat fragmentation, human-wildlife conflict, and infectious disease. A highly specialized carnivore, it is distinguished by its social structure, dental morphology, absence of dewclaws, and colorful pelage. We sequenced the genomes of two individuals from populations representing two distinct ecological histories (Laikipia County, Kenya and KwaZulu-Natal Province, South Africa). We reconstructed population demographic histories for the two individuals and scanned the genomes for evidence of selection. We show that the African wild dog has undergone at least two effective population size reductions in the last 1,000,000 years. We found evidence of Lycaon individual-specific regions of low diversity, suggestive of inbreeding or population-specific selection. Further research is needed to clarify whether these population reductions and low diversity regions are characteristic of the species as a whole. We documented positive selection on the Lycaon mitochondrial genome. Finally, we identified several candidate genes (ASIP, MITF, MLPH, PMEL) that may play a role in the characteristic Lycaon pelage.

  18. Estimating times of extinction in the fossil record

    PubMed Central

    Marshall, Charles R.

    2016-01-01

    Because the fossil record is incomplete, the last fossil of a taxon is a biased estimate of its true time of extinction. Numerous methods have been developed in the palaeontology literature for estimating the true time of extinction using ages of fossil specimens. These methods, which typically give a confidence interval for estimating the true time of extinction, differ in the assumptions they make and the nature and amount of data they require. We review the literature on such methods and make some recommendations for future directions. PMID:27122005

  19. Estimating times of extinction in the fossil record.

    PubMed

    Wang, Steve C; Marshall, Charles R

    2016-04-01

    Because the fossil record is incomplete, the last fossil of a taxon is a biased estimate of its true time of extinction. Numerous methods have been developed in the palaeontology literature for estimating the true time of extinction using ages of fossil specimens. These methods, which typically give a confidence interval for estimating the true time of extinction, differ in the assumptions they make and the nature and amount of data they require. We review the literature on such methods and make some recommendations for future directions. © 2016 The Author(s).

  20. The first darter (Aves: Anhingidae) fossils from India (late Pliocene).

    PubMed

    Stidham, Thomas; Patnaik, Rajeev; Krishan, Kewal; Singh, Bahadur; Ghosh, Abhik; Singla, Ankita; Kotla, Simran S

    2017-01-01

    New fossils from the latest Pliocene portion of the Tatrot Formation exposed in the Siwalik Hills of northern India represent the first fossil record of a darter (Anhingidae) from India. The darter fossils possibly represent a new species, but the limited information on the fossil record of this group restricts their taxonomic allocation. The Pliocene darter has a deep pit on the distal face of metatarsal trochlea IV not reported in other anhingids, it has an open groove for the m. flexor perforatus et perforans digiti II tendon on the hypotarsus unlike New World anhingid taxa, and these darter specimens are the youngest of the handful of Neogene records of the group from Asia. These fossil specimens begin to fill in a significant geographic and temporal gap in the fossil record of this group that is largely known from other continents and other time periods. The presence of a darter and pelican (along with crabs, fish, turtles, and crocodilians) in the same fossil-bearing horizon strongly indicates the past presence of a substantial water body (large pond, lake, or river) in the interior of northern India in the foothills of the Himalayan Mountains.

  1. The Fossil Calibration Database-A New Resource for Divergence Dating.

    PubMed

    Ksepka, Daniel T; Parham, James F; Allman, James F; Benton, Michael J; Carrano, Matthew T; Cranston, Karen A; Donoghue, Philip C J; Head, Jason J; Hermsen, Elizabeth J; Irmis, Randall B; Joyce, Walter G; Kohli, Manpreet; Lamm, Kristin D; Leehr, Dan; Patané, Josés L; Polly, P David; Phillips, Matthew J; Smith, N Adam; Smith, Nathan D; Van Tuinen, Marcel; Ware, Jessica L; Warnock, Rachel C M

    2015-09-01

    Fossils provide the principal basis for temporal calibrations, which are critical to the accuracy of divergence dating analyses. Translating fossil data into minimum and maximum bounds for calibrations is the most important-often least appreciated-step of divergence dating. Properly justified calibrations require the synthesis of phylogenetic, paleontological, and geological evidence and can be difficult for nonspecialists to formulate. The dynamic nature of the fossil record (e.g., new discoveries, taxonomic revisions, updates of global or local stratigraphy) requires that calibration data be updated continually lest they become obsolete. Here, we announce the Fossil Calibration Database (http://fossilcalibrations.org), a new open-access resource providing vetted fossil calibrations to the scientific community. Calibrations accessioned into this database are based on individual fossil specimens and follow best practices for phylogenetic justification and geochronological constraint. The associated Fossil Calibration Series, a calibration-themed publication series at Palaeontologia Electronica, will serve as a key pipeline for peer-reviewed calibrations to enter the database. © The Author(s) 2015. Published by Oxford University Press, on behalf of the Society of Systematic Biologists. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

  2. 75 FR 12169 - Fisheries of the Caribbean, Gulf of Mexico, and South Atlantic; Coastal Migratory Pelagic...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2010-03-15

    ... Migratory Pelagic Resources of the Gulf of Mexico and South Atlantic; Commercial King and Spanish Mackerel... levels of participation in the commercial king and Spanish mackerel components of the coastal migratory... date for king mackerel and March 31, 2010, as a possible control date for Spanish mackerel. These dates...

  3. 77 FR 4272 - Fisheries of the Caribbean, Gulf of Mexico, and South Atlantic; Coastal Migratory Pelagic...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2012-01-27

    ... group Spanish mackerel in or from the exclusive economic zone (EEZ) in the southern zone to 1,500 lb...: The fishery for coastal migratory pelagic fish (king mackerel, Spanish mackerel, cero, cobia, little... Atlantic migratory group of Spanish mackerel (65 FR 41015, July 3, 2000). Atlantic migratory group Spanish...

  4. 76 FR 9692 - Fisheries of the Caribbean, Gulf of Mexico, and South Atlantic; Coastal Migratory Pelagic...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-02-22

    ... group Spanish mackerel in or from the exclusive economic zone (EEZ) in the southern zone to 1,500 lb...: The fishery for coastal migratory pelagic fish (king mackerel, Spanish mackerel, cero, cobia, little....76 million kg) for the Atlantic migratory group of Spanish mackerel. Atlantic migratory group Spanish...

  5. Radiation and speciation of pelagic organisms during periods of global warming: the case of the common minke whale, Balaenoptera acutorostrata.

    PubMed

    Pastene, Luis A; Goto, Mutsuo; Kanda, Naohisa; Zerbini, Alexandre N; Kerem, Dan; Watanabe, Kazuo; Bessho, Yoshitaka; Hasegawa, Masami; Nielsen, Rasmus; Larsen, Finn; Palsbøll, Per J

    2007-04-01

    How do populations of highly mobile species inhabiting open environments become reproductively isolated and evolve into new species? We test the hypothesis that elevated ocean-surface temperatures can facilitate allopatry among pelagic populations and thus promote speciation. Oceanographic modelling has shown that increasing surface temperatures cause localization and reduction of upwelling, leading to fragmentation of feeding areas critical to pelagic species. We test our hypothesis by genetic analyses of populations of two closely related baleen whales, the Antarctic minke whale (Balaenoptera bonaerensis) and common minke whale (Balaenoptera acutorostrata) whose current distributions and migration patterns extent are largely determined by areas of consistent upwelling with high primary production. Phylogeographic and population genetic analyses of mitochondrial DNA control-region nucleotide sequences collected from 467 whales sampled in four different ocean basins were employed to infer the evolutionary relationship among populations of B. acutorostrata by rooting an intraspecific phylogeny with a population of B. bonaerensis. Our findings suggest that the two species diverged in the Southern Hemisphere less than 5 million years ago (Ma). This estimate places the speciation event during a period of extended global warming in the Pliocene. We propose that elevated ocean temperatures in the period facilitated allopatric speciation by disrupting the continuous belt of upwelling maintained by the Antarctic Circumpolar Current. Our analyses revealed that the current populations of B. acutorostrata likely diverged after the Pliocene some 1.5 Ma when global temperatures had decreased and presumably coinciding with the re-establishment of the polar-equatorial temperature gradient that ultimately drives upwelling. In most population samples, we detected genetic signatures of exponential population expansions, consistent with the notion of increasing carrying capacity

  6. Higher mass-independent isotope fractionation of methylmercury in the pelagic food web of Lake Baikal (Russia).

    PubMed

    Perrot, Vincent; Pastukhov, Mikhail V; Epov, Vladimir N; Husted, Søren; Donard, Olivier F X; Amouroux, David

    2012-06-05

    Mercury undergoes several transformations that influence its stable isotope composition during a number of environmental and biological processes. Measurements of Hg isotopic mass-dependent (MDF) and mass-independent fractionation (MIF) in food webs may therefore help to identify major sources and processes leading to significant bioaccumulation of methylmercury (MeHg). In this work, δ(13)C, δ(15)N, concentration of Hg species (MeHg, inorganic Hg), and stable isotopic composition of Hg were determined at different trophic levels of the remote and pristine Lake Baikal ecosystem. Muscle of seals and different fish as well as amphipods, zooplankton, and phytoplankton were specifically investigated. MDF during trophic transfer of MeHg leading to enrichment of heavier isotopes in the predators was clearly established by δ(202)Hg measurements in the pelagic prey-predator system (carnivorous sculpins and top-predator seals). Despite the low concentrations of Hg in the ecosystem, the pelagic food web reveals very high MIF Δ(199)Hg (3.15-6.65‰) in comparison to coastal fish (0.26-1.65‰) and most previous studies in aquatic organisms. Trophic transfer does not influence MIF signature since similar Δ(199)Hg was observed in sculpins (4.59 ± 0.55‰) and seal muscles (4.62 ± 0.60‰). The MIF is suggested to be mainly controlled by specific physical and biogeochemical characteristics of the water column. The higher level of MIF in pelagic fish of Lake Baikal is mainly due to the bioaccumulation of residual MeHg that is efficiently turned over and photodemethylated in deep oligotrophic and stationary (i.e., long residence time) freshwater columns.

  7. Pelagic sea snakes dehydrate at sea

    PubMed Central

    Lillywhite, Harvey B.; Sheehy, Coleman M.; Brischoux, François; Grech, Alana

    2014-01-01

    Secondarily marine vertebrates are thought to live independently of fresh water. Here, we demonstrate a paradigm shift for the widely distributed pelagic sea snake, Hydrophis (Pelamis) platurus, which dehydrates at sea and spends a significant part of its life in a dehydrated state corresponding to seasonal drought. Snakes that are captured following prolonged periods without rainfall have lower body water content, lower body condition and increased tendencies to drink fresh water than do snakes that are captured following seasonal periods of high rainfall. These animals do not drink seawater and must rehydrate by drinking from a freshwater lens that forms on the ocean surface during heavy precipitation. The new data based on field studies indicate unequivocally that this marine vertebrate dehydrates at sea where individuals may live in a dehydrated state for possibly six to seven months at a time. This information provides new insights for understanding water requirements of sea snakes, reasons for recent declines and extinctions of sea snakes and more accurate prediction for how changing patterns of precipitation might affect these and other secondarily marine vertebrates living in tropical oceans. PMID:24648228

  8. Proceedings of the sixth annual conference on fossil energy materials. Fossil Energy AR and TD Mateials Program

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Cole, N.C.; Judkins, R.R.

    1992-07-01

    The Sixth Annual Conference on Fossil Energy Materials was held in Oak Ridge, Tennessee, on May 12--14, 1992. The meeting was sponsored by the US Department of Energy`s Office of Fossil Energy through the Advanced Research and Technology Development (AR&TD) Materials Program, and ASM International. The objective of the AR&TD Materials Program is to conduct research and development on materials for longer-term fossil energy applications as well as for generic needs of various fossil fuel technologies. The management of the Program has been decentralized to the DOE Field Office, Oak Ridge with Oak Ridge National Laboratory (ORNL) as the technicalmore » support contractor. The research is performed by staff members at ORNL and by a substantial number of researchers at other national laboratories, universities, and in private industry. The work is divided into the following categories: (1) ceramics, (2) development and corrosion resistance of iron aluminide, advanced austenitic and chromium-niobium alloys, and (3) technology assessment and technology transfer. This conference is held each year to review the work on all of the projects of the Program. The agenda for the meeting is given in Appendix A, and a list of attendees is presented in Appendix B. ASM International cosponsored the conference, for which we are especially grateful.« less

  9. Impacts of hypoxia on the structure and processes in the pelagic community (zooplankton, macro-invertebrates and fish)

    NASA Astrophysics Data System (ADS)

    Ekau, W.; Auel, H.; Pörtner, H.-O.; Gilbert, D.

    2009-05-01

    Dissolved oxygen (DO) concentration in the water column is an environmental parameter that is crucial for the successful development of many pelagic organisms. Hypoxia tolerance and threshold values are species- and stage-specific and can vary enormously. While some fish species may suffer from oxygen values of less than 3 ml L-1 and show impact on growth, development and behaviour, other organisms such as euphausiids may survive DO levels as low as 0.1 ml L-1. A change in the average or the minimum or maximum DO in an area may have significant impacts on the survival of certain species and hence on the species composition in the ecosystem with consequent changes in trophic pathways and productivity. Evidence of the deleterious effects of oxygen depletion on species of the pelagic realm is scarce, particularly in terms of the effect of low oxygen on development, recruitment and patterns of migration and distribution. While planktonic organisms have to cope with different DOs and find adaptive mechanisms, nektonic species may avoid areas of inconvenient DO and develop adapted migrational strategies. Planktonic organisms may only be able to escape vertically, above or beneath the Oxygen Minimum Zone (OMZ). In shallow areas only the surface layer can serve as a refuge, in deep waters many organisms have developed vertical migration strategies to use, pass and cope with the OMZ. This paper elucidates the role of DO for different taxa in the pelagic realm and the consequences of low oxygen for foodweb structure and system productivity.

  10. Spatial scale, means and gradients of hydrographic variables define pelagic seascapes of bluefin and bullet tuna spawning distribution.

    PubMed

    Alvarez-Berastegui, Diego; Ciannelli, Lorenzo; Aparicio-Gonzalez, Alberto; Reglero, Patricia; Hidalgo, Manuel; López-Jurado, Jose Luis; Tintoré, Joaquín; Alemany, Francisco

    2014-01-01

    Seascape ecology is an emerging discipline focused on understanding how features of the marine habitat influence the spatial distribution of marine species. However, there is still a gap in the development of concepts and techniques for its application in the marine pelagic realm, where there are no clear boundaries delimitating habitats. Here we demonstrate that pelagic seascape metrics defined as a combination of hydrographic variables and their spatial gradients calculated at an appropriate spatial scale, improve our ability to model pelagic fish distribution. We apply the analysis to study the spawning locations of two tuna species: Atlantic bluefin and bullet tuna. These two species represent a gradient in life history strategies. Bluefin tuna has a large body size and is a long-distant migrant, while bullet tuna has a small body size and lives year-round in coastal waters within the Mediterranean Sea. The results show that the models performance incorporating the proposed seascape metrics increases significantly when compared with models that do not consider these metrics. This improvement is more important for Atlantic bluefin, whose spawning ecology is dependent on the local oceanographic scenario, than it is for bullet tuna, which is less influenced by the hydrographic conditions. Our study advances our understanding of how species perceive their habitat and confirms that the spatial scale at which the seascape metrics provide information is related to the spawning ecology and life history strategy of each species.

  11. Hydroacoustical evidence of the expansion of pelagic swarms of Munida gregaria (Decapoda, Munididae) in the Beagle Channel and the Argentine Patagonian Shelf, and its relationship with habitat features

    NASA Astrophysics Data System (ADS)

    Diez, Mariano J.; Cabreira, Ariel G.; Madirolas, Adrián; Lovrich, Gustavo A.

    2016-08-01

    Squat lobsters are highly diversified and widespread decapods, of which only three species form pelagic swarms. Here we infer the expansion of Munida gregaria populations in the Beagle Channel and the Argentine Patagonian Shelf by means of acoustic surveys of pelagic swarms. We also describe the habitat characteristics in which these swarms occur. Acoustic data was collected during three multidisciplinary scientific cruises on board of the R/V Puerto Deseado during 2009, 2012 and 2014. Despite differences in the environmental conditions between the two surveyed areas, between 2009 and 2014 pelagic swarms increased their occurrence and abundance both in the Beagle Channel and on the Argentine Patagonian Shelf. Towards the end of the studied period, pelagic swarms of M. gregaria occurred in new locations, supporting the notion of a population expansion. Within the Beagle Channel swarm expansions were more marked than on the Patagonian Shelf. We here postulate that M. gregaria expansions occur in association with productive areas of the Argentine continental shelf, such as frontal zones, favoured by the squat lobster phenotypic plasticity that permit to exploit resources in both the neritic and benthic environments. At a regional scale on the Patagonian Shelf, three main groups of pelagic swarms of M. gregaria were clearly associated to respective frontal zones. The information presented here is necessary to understand fluctuations in both distribution and abundance patterns of a key species on the Argentine continental shelf. These fluctuations could be direct or indirect indicators of changes in the ecosystem.

  12. Structure and evolution of fossil H II regions

    NASA Technical Reports Server (NTRS)

    Mccray, R.; Schwarz, J.

    1971-01-01

    The structure and evolution of a fossil H II region created by a burst of ionizing radiation from a supernova is considered. The cooling time scale for the shell is about 10 to the 6th power years. Superposition of million-year-old fossil H II regions may account for the temperature and ionization of the interstellar medium. Fossil H II regions are unstable to growth of thermal condensations. Highly ionized filamentary structures form and dissipate in about 10,000 years. Partially ionized clouds form and dissipate in about 10 to the 6th power years.

  13. Microbial Fossils Detected in Desert Varnish

    NASA Technical Reports Server (NTRS)

    Flood, B. E.; Allen, C.; Longazo, T.

    2003-01-01

    Mars Global Surveyor Thermal Emission Spectrometer data indicate regions with significant levels of hematite (_Fe2O3). Fe-oxides, like hematite, can form as aqueous mineral precipitates and as such may preserve microscopic fossils or other biosignatures. Several potential terrestrial analogues to martian hematite like hydrothermal vents have preserved microfossils. Microbial fossilization in Fe-oxides is often a function of biomineralization. For example, goethite (FeO2H) encrustation of fungal mycelia from the mid-Tertiary preserved fungal morphologies such that their genera could be determined.

  14. Fossilization and Learning Strategies in Second Language Acquisition.

    ERIC Educational Resources Information Center

    Sims, William R.

    1989-01-01

    In interlanguage, the transitional state reaching from one's native language to a given target language, phonological, morphological, syntactic, lexical, sociocultural, or psycholinguistic errors may be generated and systematized by the process of fossilization. Depending on the amount of time needed for remediation, fossilized features may be…

  15. Fossil-Fuel C02 Emissions Database and Exploration System

    NASA Astrophysics Data System (ADS)

    Krassovski, M.; Boden, T.

    2012-04-01

    Fossil-Fuel C02 Emissions Database and Exploration System Misha Krassovski and Tom Boden Carbon Dioxide Information Analysis Center Oak Ridge National Laboratory The Carbon Dioxide Information Analysis Center (CDIAC) at Oak Ridge National Laboratory (ORNL) quantifies the release of carbon from fossil-fuel use and cement production each year at global, regional, and national spatial scales. These estimates are vital to climate change research given the strong evidence suggesting fossil-fuel emissions are responsible for unprecedented levels of carbon dioxide (CO2) in the atmosphere. The CDIAC fossil-fuel emissions time series are based largely on annual energy statistics published for all nations by the United Nations (UN). Publications containing historical energy statistics make it possible to estimate fossil-fuel CO2 emissions back to 1751 before the Industrial Revolution. From these core fossil-fuel CO2 emission time series, CDIAC has developed a number of additional data products to satisfy modeling needs and to address other questions aimed at improving our understanding of the global carbon cycle budget. For example, CDIAC also produces a time series of gridded fossil-fuel CO2 emission estimates and isotopic (e.g., C13) emissions estimates. The gridded data are generated using the methodology described in Andres et al. (2011) and provide monthly and annual estimates for 1751-2008 at 1° latitude by 1° longitude resolution. These gridded emission estimates are being used in the latest IPCC Scientific Assessment (AR4). Isotopic estimates are possible thanks to detailed information for individual nations regarding the carbon content of select fuels (e.g., the carbon signature of natural gas from Russia). CDIAC has recently developed a relational database to house these baseline emissions estimates and associated derived products and a web-based interface to help users worldwide query these data holdings. Users can identify, explore and download desired CDIAC

  16. Research needs for finely resolved fossil carbon emissions

    USGS Publications Warehouse

    Gurney, K.; Ansley, W.; Mendoza, D.; Petron, G.; Frost, G.; Gregg, J.; Fischer, M.; Pataki, Diane E.; Ackerman, K.; Houweling, S.; Corbin, K.; Andres, R.; Blasing, T.J.

    2007-01-01

    Scientific research on the global carbon cycle has emerged as a high priority in biogeochemistry, climate studies, and global change policy. The emission of carbon dioxide (CO2) from fossil fuel combustion is a dominant driver of the current net carbon fluxes between the land, the oceans, and the atmosphere, and it is a key contributor to the rise in modern radiative forcing. Contrary to a commonly held perception, our quantitative knowledge about these emissions is insufficient to satisfy current scientific and policy needs. A more highly spatially and temporally resolved quantification of the social and economic drivers of fossil fuel combustion, and the resulting CO2 emissions, is essential to supporting scientific and policy progress. In this article, a new community of emissions researchers called the CO2 Fossil Fuel Emission Effort (CO2FFEE) outlines a research agenda to meet the need for improved fossil fuel CO2 emissions information and solicits comment from the scientific community and research agencies.

  17. Identifying Fossil Shell Resources via Geophysical Surveys: Chesapeake Bay Region, Virginia

    DTIC Science & Technology

    2016-05-01

    ER D C/ CH L TR -1 6- 4 Chesapeake Fossil Shell Survey Identifying Fossil Shell Resources via Geophysical Surveys: Chesapeake Bay Region...other technical reports published by ERDC, visit the ERDC online library at http://acwc.sdp.sirsi.net/client/default. Chesapeake Fossil Shell...Survey ERDC/CHL TR-16-4 May 2016 Identifying Fossil Shell Resources via Geophysical Surveys: Chesapeake Bay Region, Virginia Heidi M. Wadman and Jesse

  18. Status of Pelagic Prey Fishes in Lake Michigan, 2014

    USGS Publications Warehouse

    Warner, David M.; Farha, Steven A.; Claramunt, Randall M.; Hanson, Dale; O'Brien, Timothy P.

    2015-01-01

    Acoustic surveys were conducted in late summer/early fall during the years 1992-1996 and 2001-2014 to estimate pelagic prey fish biomass in Lake Michigan. Midwater trawling during the surveys as well as target strength provided a measure of species and size composition of the fish community for use in scaling acoustic data and providing species-specific abundance estimates. The 2014 survey consisted of 27 acoustic transects (603 km total) and 31 midwater trawl tows. Four additional transects were sampled in Green Bay but were not included in lakewide estimates. Mean prey fish biomass was 6.5 kg/ha [31.7 kilotonnes (kt = 1,000 metric tons)], equivalent to 69.9 million pounds, which was similar to the estimate in 2013 (29.6 kt) and 25% of the long-term (19 years) mean. The numeric density of the 2014 alewife year-class was 3% of the time series average and was the lowest observed in the 19 years of sampling. This year-class contributed <1% of total alewife biomass (4.6 kg/ha). Alewife ≥age-1 comprised 99.5% of alewife biomass. Numeric density of alewife in Green Bay was more than three times that of the main lake. In 2014, alewife comprised 71% of total prey fish biomass, while rainbow smelt and bloater were 1% and 28% of total biomass, respectively. Rainbow smelt biomass in 2014 (0.08 kg/ha) was 66% lower than in 2013, 2% of the long-term mean, and lower than in any previous year. Bloater biomass in 2014 was 1.8 kg/ha, nearly three times more than the 2013 biomass, and 20% of the long-term mean. Mean density of small bloater in 2014 (122 fish/ha) was lower than peak values observed in 2007-2009 but was similar to the time series mean (124 fish/ha). In 2014, pelagic prey fish biomass in Lake Michigan was 71% of that in Lake Huron (all basins), where the community is dominated by bloater.

  19. Status of pelagic prey fishes in Lake Michigan, 2012

    USGS Publications Warehouse

    Warner, David M.; O'Brien, Timothy P.; Farha, Steve A.; Claramunt, Randall M.; Hanson, Dale

    2012-01-01

    Acoustic surveys were conducted in late summer/early fall during the years 1992-1996 and 2001-2012 to estimate pelagic prey fish biomass in Lake Michigan. Midwater trawling during the surveys as well as target strength provided a measure of species and size composition of the fish community for use in scaling acoustic data and providing species-specific abundance estimates. The 2012 survey consisted of 26 acoustic transects (576 km total) and 31 midwater tows. Mean total prey fish biomass was 6.4 kg/ha (relative standard error, RSE = 15%) or 31 kilotonnes (kt = 1,000 metric tons), which was 1.5 times the estimate for 2011 and 22% of the long-term mean. The increase from 2011 resulted from increased biomass of age-0 alewife, age-1 or older alewife, and large bloater. The abundance of the 2012 alewife year class was similar to the average, and this year-class contributed 35% of total alewife biomass (4.9 kg/ha, RSE = 17%), while the 2010 alewife year-class contributed 58%. The 2010 year class made up 89% of age-1 or older alewife biomass. In 2012, alewife comprised 77% of total prey fish biomass, while rainbow smelt and bloater were 4 and 19% of total biomass, respectively. Rainbow smelt biomass in 2012 (0.25 kg/ha, RSE = 17%) was 40% of the rainbow smelt biomass in 2011 and 5% of the long term mean. Bloater biomass was much lower (1.2 kg/ha, RSE = 12%) than in the 1990s, and mean density of small bloater in 2012 (191 fish/ha, RSE = 24%) was lower than peak values observed in 2007-2009. In 2012, pelagic prey fish biomass in Lake Michigan was similar to Lake Superior and Lake Huron. Prey fish biomass remained well below the Fish Community Objectives target of 500-800 kt, and key native species remain absent or rare.

  20. Stratigraphy and facies development of the marine Late Devonian near the Boulongour Reservoir, northwest Xinjiang, China

    NASA Astrophysics Data System (ADS)

    Suttner, Thomas J.; Kido, Erika; Chen, Xiuqin; Mawson, Ruth; Waters, Johnny A.; Frýda, Jiří; Mathieson, David; Molloy, Peter D.; Pickett, John; Webster, Gary D.; Frýdová, Barbora

    2014-02-01

    Late Devonian to Early Carboniferous stratigraphic units within the 'Zhulumute' Formation, Hongguleleng Formation (stratotype), 'Hebukehe' Formation and the Heishantou Formation near the Boulongour Reservoir in northwestern Xinjiang are fossil-rich. The Hongguleleng and 'Hebukehe' formations are biostratigraphically well constrained by microfossils from the latest Frasnian linguiformis to mid-Famennian trachytera conodont biozones. The Hongguleleng Formation (96.8 m) is characterized by bioclastic argillaceous limestones and marls (the dominant facies) intercalated with green spiculitic calcareous shales. It yields abundant and highly diverse faunas of bryozoans, brachiopods and crinoids with subordinate solitary rugose corals, ostracods, trilobites, conodonts and other fish teeth. The succeeding 'Hebukehe' Formation (95.7 m) consists of siltstones, mudstones, arenites and intervals of bioclastic limestone (e.g. 'Blastoid Hill') and cherts with radiolarians. A diverse ichnofauna, phacopid trilobites, echinoderms (crinoids and blastoids) together with brachiopods, ostracods, bryozoans and rare cephalopods have been collected from this interval. Analysis of geochemical data, microfacies and especially the distribution of marine organisms, which are not described in detail here, but used for facies analysis, indicate a deepening of the depositional environment at the Boulongour Reservoir section. Results presented here concern mainly the sedimentological and stratigraphical context of the investigated section. Additionally, one Late Devonian palaeo-oceanic and biotic event, the Upper Kellwasser Event is recognized near the section base.

  1. Fossil moonseeds from the Paleogene of West Gondwana (Patagonia, Argentina).

    PubMed

    Jud, Nathan A; Iglesias, Ari; Wilf, Peter; Gandolfo, Maria A

    2018-06-08

    The fossil record is critical for testing biogeographic hypotheses. Menispermaceae (moonseeds) are a widespread family with a rich fossil record and alternative hypotheses related to their origin and diversification. The family is well-represented in Cenozoic deposits of the northern hemisphere, but the record in the southern hemisphere is sparse. Filling in the southern record of moonseeds will improve our ability to evaluate alternative biogeographic hypotheses. Fossils were collected from the Salamanca (early Paleocene, Danian) and the Huitrera (early Eocene, Ypresian) formations in Chubut Province, Argentina. We photographed them using light microscopy, epifluorescence, and scanning electron microscopy and compared the fossils with similar extant and fossil Menispermaceae using herbarium specimens and published literature. We describe fossil leaves and endocarps attributed to Menispermaceae from Argentinean Patagonia. The leaves are identified to the family, and the endocarps are further identified to the tribe Cissampelideae. The Salamancan endocarp is assigned to the extant genus Stephania. These fossils significantly expand the known range of Menispermaceae in South America, and they include the oldest (ca. 64 Ma) unequivocal evidence of the family worldwide. Our findings highlight the importance of West Gondwana in the evolution of Menispermaceae during the Paleogene. Currently, the fossil record does not discern between a Laurasian or Gondwanan origin; however, it does demonstrate that Menispermaceae grew well outside the tropics by the early Paleocene. The endocarps' affinity with Cissampelideae suggests that diversification of the family was well underway by the earliest Paleocene. © 2018 The Authors. American Journal of Botany is published by Wiley Periodicals, Inc. on behalf of the Botanical Society of America.

  2. The inorganic constituents of echinoderms

    USGS Publications Warehouse

    Clarke, F.W.; Wheeler, W.C.

    1915-01-01

    In a recent paper on the composition of crinoid skeletons we showed that crinoids contain large quantities of magnesia, and that its proportion varies with the temperature of the water in which the creatures live. This result was so novel and surprising that it seemed desirable to examine other echinoderms and to ascertain whether they showed the same characteristics and regularity. A number of sea urchins and starfishes were therefore studied, their inorganic constituents being analyzed in the same manner as those of the crinoids

  3. Beyond fossil fuel–driven nitrogen transformations

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Chen, Jingguang G.; Crooks, Richard M.; Seefeldt, Lance C.

    Nitrogen is fundamental to all of life and many industrial processes. The interchange of nitrogen oxidation states in the industrial production of ammonia, nitric acid, and other commodity chemicals is largely powered by fossil fuels. Here, a key goal of contemporary research in the field of nitrogen chemistry is to minimize the use of fossil fuels by developing more efficient heterogeneous, homogeneous, photo-, and electrocatalytic processes or by adapting the enzymatic processes underlying the natural nitrogen cycle. These approaches, as well as the challenges involved, are discussed in this Review.

  4. Beyond fossil fuel–driven nitrogen transformations

    DOE PAGES

    Chen, Jingguang G.; Crooks, Richard M.; Seefeldt, Lance C.; ...

    2018-05-25

    Nitrogen is fundamental to all of life and many industrial processes. The interchange of nitrogen oxidation states in the industrial production of ammonia, nitric acid, and other commodity chemicals is largely powered by fossil fuels. Here, a key goal of contemporary research in the field of nitrogen chemistry is to minimize the use of fossil fuels by developing more efficient heterogeneous, homogeneous, photo-, and electrocatalytic processes or by adapting the enzymatic processes underlying the natural nitrogen cycle. These approaches, as well as the challenges involved, are discussed in this Review.

  5. 43 CFR 8224.1 - Use of the Fossil Forest Research Natural Area.

    Code of Federal Regulations, 2011 CFR

    2011-10-01

    ... 43 Public Lands: Interior 2 2011-10-01 2011-10-01 false Use of the Fossil Forest Research Natural...) BUREAU OF LAND MANAGEMENT, DEPARTMENT OF THE INTERIOR RECREATION PROGRAMS PROCEDURES Fossil Forest Research Natural Area § 8224.1 Use of the Fossil Forest Research Natural Area. (a) Fossils may be collected...

  6. 43 CFR 8224.1 - Use of the Fossil Forest Research Natural Area.

    Code of Federal Regulations, 2012 CFR

    2012-10-01

    ... 43 Public Lands: Interior 2 2012-10-01 2012-10-01 false Use of the Fossil Forest Research Natural...) BUREAU OF LAND MANAGEMENT, DEPARTMENT OF THE INTERIOR RECREATION PROGRAMS PROCEDURES Fossil Forest Research Natural Area § 8224.1 Use of the Fossil Forest Research Natural Area. (a) Fossils may be collected...

  7. 43 CFR 8224.1 - Use of the Fossil Forest Research Natural Area.

    Code of Federal Regulations, 2013 CFR

    2013-10-01

    ... 43 Public Lands: Interior 2 2013-10-01 2013-10-01 false Use of the Fossil Forest Research Natural...) BUREAU OF LAND MANAGEMENT, DEPARTMENT OF THE INTERIOR RECREATION PROGRAMS PROCEDURES Fossil Forest Research Natural Area § 8224.1 Use of the Fossil Forest Research Natural Area. (a) Fossils may be collected...

  8. 43 CFR 8224.1 - Use of the Fossil Forest Research Natural Area.

    Code of Federal Regulations, 2014 CFR

    2014-10-01

    ... 43 Public Lands: Interior 2 2014-10-01 2014-10-01 false Use of the Fossil Forest Research Natural...) BUREAU OF LAND MANAGEMENT, DEPARTMENT OF THE INTERIOR RECREATION PROGRAMS PROCEDURES Fossil Forest Research Natural Area § 8224.1 Use of the Fossil Forest Research Natural Area. (a) Fossils may be collected...

  9. Fossil group origins. VIII. RX J075243.6+455653 a transitionary fossil group

    NASA Astrophysics Data System (ADS)

    Aguerri, J. A. L.; Longobardi, A.; Zarattini, S.; Kundert, A.; D'Onghia, E.; Domínguez-Palmero, L.

    2018-01-01

    Context. It is thought that fossil systems are relics of structure formation in the primitive Universe. They are galaxy aggregations that have assembled their mass at high redshift with few or no subsequent accretion. Observationally these systems are selected by large magnitude gaps between their 1st and 2nd ranked galaxies (Δm12). Nevertheless, there is still debate over whether or not this observational criterium selects dynamically evolved ancient systems. Aims: We have studied the properties of the nearby fossil group RX J075243.6+455653 in order to understand the mass assembly of this system. Methods: Deep spectroscopic observations allow us to construct the galaxy luminosity function (LF) of RX J075243.6+455653 down to Mr*+6. The analysis of the faint-end of the LF in groups and clusters provides valuable information about the mass assembly of the system. In addition, we have analyzed the nearby large-scale structure around this group. Results: We identified 26 group members within r200 0.96 Mpc. These galaxies are located at Vc = 15551 ± 65 km s-1 and have a velocity dispersion of σc = 333 ± 46 km s-1. The X-ray luminosity of the group is LX = 2.2 × 1043 h70-2 erg s-1, resulting in a mass of M = 4.2 × 1013 h70-1 within 0.5r200. The group has Δm12 = 2.1 within 0.5r200, confirming the fossil nature of this system. RX J075243.6+455653 has a central brightest group galaxy (BGG) with Mr = -22.67, one of the faintest BGGs observed in fossil systems. The LF of the group shows a flat faint-end slope (α = -1.08 ± 0.33). This low density of dwarf galaxies is confirmed by the low value of the dwarf-to-giant ratio (DGR = 0.99 ± 0.49) for this system. Both the lack of dwarf galaxies and the low luminosity of the BGG suggests that RX J075243.6+455653 still has to accrete mass from its nearby environment. This mass accretion will be achieved because it is the dominant structure of a rich environment formed by several groups of galaxies (15) within 7 Mpc from the

  10. Total-Evidence Dating under the Fossilized Birth–Death Process

    PubMed Central

    Zhang, Chi; Stadler, Tanja; Klopfstein, Seraina; Heath, Tracy A.; Ronquist, Fredrik

    2016-01-01

    Bayesian total-evidence dating involves the simultaneous analysis of morphological data from the fossil record and morphological and sequence data from recent organisms, and it accommodates the uncertainty in the placement of fossils while dating the phylogenetic tree. Due to the flexibility of the Bayesian approach, total-evidence dating can also incorporate additional sources of information. Here, we take advantage of this and expand the analysis to include information about fossilization and sampling processes. Our work is based on the recently described fossilized birth–death (FBD) process, which has been used to model speciation, extinction, and fossilization rates that can vary over time in a piecewise manner. So far, sampling of extant and fossil taxa has been assumed to be either complete or uniformly at random, an assumption which is only valid for a minority of data sets. We therefore extend the FBD process to accommodate diversified sampling of extant taxa, which is standard practice in studies of higher-level taxa. We verify the implementation using simulations and apply it to the early radiation of Hymenoptera (wasps, ants, and bees). Previous total-evidence dating analyses of this data set were based on a simple uniform tree prior and dated the initial radiation of extant Hymenoptera to the late Carboniferous (309 Ma). The analyses using the FBD prior under diversified sampling, however, date the radiation to the Triassic and Permian (252 Ma), slightly older than the age of the oldest hymenopteran fossils. By exploring a variety of FBD model assumptions, we show that it is mainly the accommodation of diversified sampling that causes the push toward more recent divergence times. Accounting for diversified sampling thus has the potential to close the long-discussed gap between rocks and clocks. We conclude that the explicit modeling of fossilization and sampling processes can improve divergence time estimates, but only if all important model aspects

  11. Total-Evidence Dating under the Fossilized Birth-Death Process.

    PubMed

    Zhang, Chi; Stadler, Tanja; Klopfstein, Seraina; Heath, Tracy A; Ronquist, Fredrik

    2016-03-01

    Bayesian total-evidence dating involves the simultaneous analysis of morphological data from the fossil record and morphological and sequence data from recent organisms, and it accommodates the uncertainty in the placement of fossils while dating the phylogenetic tree. Due to the flexibility of the Bayesian approach, total-evidence dating can also incorporate additional sources of information. Here, we take advantage of this and expand the analysis to include information about fossilization and sampling processes. Our work is based on the recently described fossilized birth-death (FBD) process, which has been used to model speciation, extinction, and fossilization rates that can vary over time in a piecewise manner. So far, sampling of extant and fossil taxa has been assumed to be either complete or uniformly at random, an assumption which is only valid for a minority of data sets. We therefore extend the FBD process to accommodate diversified sampling of extant taxa, which is standard practice in studies of higher-level taxa. We verify the implementation using simulations and apply it to the early radiation of Hymenoptera (wasps, ants, and bees). Previous total-evidence dating analyses of this data set were based on a simple uniform tree prior and dated the initial radiation of extant Hymenoptera to the late Carboniferous (309 Ma). The analyses using the FBD prior under diversified sampling, however, date the radiation to the Triassic and Permian (252 Ma), slightly older than the age of the oldest hymenopteran fossils. By exploring a variety of FBD model assumptions, we show that it is mainly the accommodation of diversified sampling that causes the push toward more recent divergence times. Accounting for diversified sampling thus has the potential to close the long-discussed gap between rocks and clocks. We conclude that the explicit modeling of fossilization and sampling processes can improve divergence time estimates, but only if all important model aspects

  12. Fauna and paleoecological setting of the La Meseta Formation (Eocene), Antarctica

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Feldmann, R.M.; Wiedman, L.A.; Zinsmeister, W.J.

    The La Meseta Formation, an Eocene sandstone from Seymour Island, Palmer Peninsula, Antarctica, has yielded a diverse fossil assemblage of body and trace fossils representative of a cool temperate, littoral to shallow sublittoral habitat. Over 61 taxa of macroinvertebrates, excluding gastropod body fossils, and more than 18 ichnogenera collected from the La Meseta represent the largest, most comprehensive and most diverse assemblage of Paleogene fossils from Antarctica. Included in the body fossil assemblage are species representative of at least 26 taxa of bivales, four taxa of echinoids, two of crinoids, two of ophiuroids, two of asteroids, one inarticulate and fourmore » articulate brachiopods, two barnacles, six decapod crustaceans, two cyclostome and two cheilostome bryozoans, a scaphopod and one coral. The traces include several burrow forms characteristic of the Skolithos ichnofacies of Seilacher (1967), several halo and rind burrows, gastropod predation borings, and abundant examples of teredid bivalve borings in lithified wood.Autecological analyses of the preserved organisms and environmental interpretations of the ichnogenera indicate a littoral to very shallow sublittoral environment of deposition, generally above wave base, for the la Meseta Sandstone. Modern congeneric descendants of the body fossils are known to inhabit both deep water and shallow water habitats. Of the 20 extant genera of bivalves reported from the La Meseta, 19 generally occur only in cool temperate habitats. Only one genus is known to occur south of 60/sup 0/. Most of the shallow water forms are known from cool temperate, austral regimes.« less

  13. Impacts of hypoxia on the structure and processes in pelagic communities (zooplankton, macro-invertebrates and fish)

    NASA Astrophysics Data System (ADS)

    Ekau, W.; Auel, H.; Pörtner, H.-O.; Gilbert, D.

    2010-05-01

    Dissolved oxygen (DO) concentration in the water column is an environmental parameter that is crucial for the successful development of many pelagic organisms. Hypoxia tolerance and threshold values are species- and stage-specific and can vary enormously. While some fish species may suffer from oxygen values of less than 3 mL O2 L-1 through impacted growth, development and behaviour, other organisms such as euphausiids may survive DO levels as low as 0.1 mL O2 L-1. A change in the average or the range of DO may have significant impacts on the survival of certain species and hence on the species composition in the ecosystem with consequent changes in trophic pathways and productivity. Evidence for the deleterious effects of oxygen depletion on pelagic species is scarce, particularly in terms of the effect of low oxygen on development, recruitment and patterns of migration and distribution. While planktonic organisms have to cope with variable DOs and exploit adaptive mechanisms, nektonic species may avoid areas of unfavourable DO and develop adapted migration strategies. Planktonic organisms may only be able to escape vertically, above or beneath the Oxygen Minimum Zone (OMZ). In shallow areas only the surface layer can serve as a refuge, but in deep waters many organisms have developed vertical migration strategies to use, pass through and cope with the OMZ. This paper elucidates the role of DO for different taxa in the pelagic realm and the consequences of low oxygen for foodweb structure and system productivity. We describe processes in two contrasting systems, the semi-enclosed Baltic Sea and the coastal upwelling system of the Benguela Current to demonstrate the consequences of increasing hypoxia on ecosystem functioning and services.

  14. Long-term impact of bottom trawling on pelagic-benthic coupling in the southern North Sea (German Bight)

    NASA Astrophysics Data System (ADS)

    Friedrich, Jana; van Beusekom, Justus E. E.; Neumann, Andreas; Naderipour, Celine; Janssen, Felix; Ahmerkamp, Soeren; Holtappels, Moritz; Schueckel, Ulrike

    2016-04-01

    The southern North Sea, and the German Bight, has been systematically bottom-trawled at least since the late 19th century (Christiansen, 2009; Reiss et al., 2009; Kröncke 2011; Emeis et al., 2015, Neumann et al., 2016). As a result, benthic habitats and benthic biogenic structures created by bivalves, polychaetes and hydroids where destroyed or reduced. The parallel removal of hard substrate (gravel and boulders) avoids the resettlement of hard-substrate depended species. For example, the Oyster ground, a huge oyster bank a hundred years ago (Olsen, 1883), turned into a muddy depression today. In addition, shallow depth of max 40 m, strong tidal currents and frequent storms result in a high-energy environment with low sedimentation rates and recurrent sediment resuspension. The decrease in benthic filtering capacity by disturbance in epifauna and bottom roughness (Callaway et al., 2007) apparently influence pelagic-benthic coupling of biogeochemical fluxes. Heip et al. (1995) indicate that benthic respiration at depths prevailing in the German Bight accounts for 10-40% of total respiration, whereas pelagic respiration accounts for 60-90%. Previous estimates are in the middle of this range (Heip et al., 1995). To test these hypotheses and to assess the partitioning of benthic and pelagic processes, and the factors influencing organic matter mineralization, we measured pelagic production and respiration based on Winkler titration, in-situ benthic fluxes using chamber landers, we did ex-situ incubations of intact sediment cores and analysed still images from a towed benthic video sled. In addition, O2 fluxes in permeable sediments were estimated by integrating the volumetric rate measurements of the upper sediment layer over in-situ microsensor-measured O2 penetration depth. Our current results show significant seasonality in benthic respiration, with highest rates in summer and lowest rates in winter. No significant differences in total benthic respiration rates

  15. Mud fossils

    USGS Publications Warehouse

    ,

    1997-01-01

    At the close of the 18th century, the haze of fantasy and mysticism that tended to obscure the true nature of the Earth was being swept away. Careful studies by scientists showed that rocks had diverse origins. Some rock layers, containing clearly identifiable fossil remains of fish and other forms of aquatic animal and plant life, originally formed in the ocean. Other layers, consisting of sand grains winnowed clean by the pounding surf, obviously formed as beach deposits that marked the shorelines of ancient seas.

  16. Urea and lipid extraction treatment effects on δ(15)N and δ(13)C values in pelagic sharks.

    PubMed

    Li, Yunkai; Zhang, Yuying; Hussey, Nigel E; Dai, Xiaojie

    2016-01-15

    Stable isotope analysis (SIA) provides a powerful tool to investigate diverse ecological questions for marine species, but standardized values are required for comparative assessments. For elasmobranchs, their unique osmoregulatory strategy involves retention of (15)N-depleted urea in body tissues and this may bias δ(15)N values. This may be a particular problem for large predatory species, where δ(15)N discrimination between predator and consumed prey can be small. We evaluated three treatments (deionized water rinsing [DW], chloroform/methanol [LE] and combined chloroform/methanol and deionized water rinsing [LE+DW]) applied to white muscle tissue of 125 individuals from seven pelagic shark species to (i) assess urea and lipid effects on stable isotope values determined by IRMS and (ii) investigate mathematical normalization of these values. For all species examined, the δ(15)N values and C:N ratios increased significantly following all three treatments, identifying that urea removal is required prior to SIA of pelagic sharks. The more marked change in δ(15)N values following DW (1.3 ± 0.4‰) and LE+DW (1.2 ± 0.6‰) than following LE alone (0.7 ± 0.4‰) indicated that water rinsing was more effective at removing urea. The DW and LE+DW treatments lowered the %N values, resulting in an increase in C:N ratios from the unexpected low values of <2.6 in bulk samples to ~3.1 ± 0.1, the expected value of protein. The δ(13)C values of all species also increased significantly following LE and LE+DW treatments. Given the mean change in δ(15)N(1.2 ± 0.6‰) and δ(13)C values (0.7 ± 0.4‰) across pelagic shark species, it is recommended that muscle tissue samples be treated with LE+DW to efficiently extract both urea and lipids to standardize isotopic values. Mathematical normalization of urea and lipid-extracted δ(15)N(LE+DW) and δ(13)C(LE+DW) values using the lipid-extracted δ(15)N(LE) and δ(13)C(LE) data were established for all pelagic shark species

  17. Deep questions about the nature of early-life signals: a commentary on Lister (1673) ‘A description of certain stones figured like plants’

    PubMed Central

    Brasier, Martin

    2015-01-01

    In 1673, Martin Lister explored the preservation of ‘St Cuthbert's beads’ plus other fossil crinoid remains from approximately 350 Ma Carboniferous limestone in northern England. He used taphonomic evidence (transport, disarticulation, burial and cementation) to infer an origin as petrified plant remains, in contrast with his views expressed elsewhere that fossil mollusc shells could have formed abiogenically, by ‘plastic forces’ within rock. Lister also observed pentagonal symmetry, now seen as characteristic of living echinoderm skeletons. A postscript from John Ray supports Lister's ‘taphonomic’ observations and accepts the biogenicity of these fossil ‘vegetables’. Ray then concluded with a prophecy, predicting the discovery of comparable living fossils in remote ocean waters. These early discussions compare with current debates about the character of candidate microfossils from the early Earth and Mars. Interesting biomorphs are now tested against the abiogenic null hypotheses, making use of features such as those pioneered by Lister, including evidence for geological context, rules for growth and taphonomy. Advanced techniques now allow us to extend this list of criteria to include the nanoscale mapping of biology-like behaviour patterns plus metabolic pathways. Whereas the science of palaeobiology once began with tests for biogenicity, the same is now true for geobiology and astrobiology. This commentary was written to celebrate the 350th anniversary of the journal Philosophical Transactions of the Royal Society. PMID:25750150

  18. 76 FR 39789 - Fisheries of the Exclusive Economic Zone Off Alaska; Northern Rockfish and Pelagic Shelf Rockfish...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-07-07

    .... 101126522-0640-02] RIN 0648-XA546 Fisheries of the Exclusive Economic Zone Off Alaska; Northern Rockfish and Pelagic Shelf Rockfish for Trawl Catcher Vessels Participating in the Entry Level Rockfish Fishery in the...) for trawl catcher vessels participating in the entry level rockfish fishery in the Central Regulatory...

  19. Small pelagics in a changing ocean: biological responses of sardine early stages to warming

    PubMed Central

    Faleiro, Filipa; Pimentel, Marta; Pegado, Maria Rita; Bispo, Regina; Lopes, Ana Rita; Diniz, Mário S.; Rosa, Rui

    2016-01-01

    Small pelagic fishes are known to respond rapidly to changes in ocean climate. In this study, we evaluate the effects of future environmental warming (+2°C) during the early ontogeny of the European sardine, Sardina pilchardus. Warming reduced the survival of 30-day-old larvae by half. Length at hatching increased with temperature as expected, but no significant effect was observed on the length and growth at 30 days post-hatching. Warming did not significantly affect the thermal tolerance of sardine larvae, even though the mean lethal temperature increased by 1°C. In the warm conditions, sardine larvae showed signs of thermal stress, indicated by a pronounced increase in larval metabolism (Q10 = 7.9) and a 45% increase in the heat shock response. Lipid peroxidation was not significantly affected by the higher temperature, even though the mean value doubled. Warming did not affect the time larvae spent swimming, but decreased by 36% the frequency of prey attacks. Given the key role of these small pelagics in the trophic dynamics off the Western Iberian upwelling ecosystem, the negative effects of warming on the early stages may have important implications for fish recruitment and ecosystem structure. PMID:27293764

  20. New Ediacara fossils preserved in marine limestone and their ecological implications.

    PubMed

    Chen, Zhe; Zhou, Chuanming; Xiao, Shuhai; Wang, Wei; Guan, Chengguo; Hua, Hong; Yuan, Xunlai

    2014-02-25

    Ediacara fossils are central to our understanding of animal evolution on the eve of the Cambrian explosion, because some of them likely represent stem-group marine animals. However, some of the iconic Ediacara fossils have also been interpreted as terrestrial lichens or microbial colonies. Our ability to test these hypotheses is limited by a taphonomic bias that most Ediacara fossils are preserved in sandstones and siltstones. Here we report several iconic Ediacara fossils and an annulated tubular fossil (reconstructed as an erect epibenthic organism with uniserial arranged modular units), from marine limestone of the 551-541 Ma Dengying Formation in South China. These fossils significantly expand the ecological ranges of several key Ediacara taxa and support that they are marine organisms rather than terrestrial lichens or microbial colonies. Their close association with abundant bilaterian burrows also indicates that they could tolerate and may have survived moderate levels of bioturbation.

  1. Caught in the act: the first record of copulating fossil vertebrates.

    PubMed

    Joyce, Walter G; Micklich, Norbert; Schaal, Stephan F K; Scheyer, Torsten M

    2012-10-23

    The behaviour of fossil organisms can typically be inferred only indirectly, but rare fossil finds can provide surprising insights. Here, we report from the Eocene Messel Pit Fossil Site between Darmstadt and Frankfurt, Germany numerous pairs of the fossil carettochelyid turtle Allaeochelys crassesculpta that represent for the first time among fossil vertebrates couples that perished during copulation. Females of this taxon can be distinguished from males by their relatively shorter tails and development of plastral kinesis. The preservation of mating pairs has important taphonomic implications for the Messel Pit Fossil Site, as it is unlikely that the turtles would mate in poisonous surface waters. Instead, the turtles initiated copulation in habitable surface waters, but perished when their skin absorbed poisons while sinking into toxic layers. The mating pairs from Messel are therefore more consistent with a stratified, volcanic maar lake with inhabitable surface waters and a deadly abyss.

  2. Diagnosing Homo sapiens in the fossil record.

    PubMed

    Stringer, Christopher Brian; Buck, Laura Tabitha

    2014-01-01

    Diagnosing Homo sapiens is a critical question in the study of human evolution. Although what constitutes living members of our own species is straightforward, in the fossil record this is still a matter of much debate. The issue is complicated by questions of species diagnoses and ideas about the mode by which a new species is born, by the arguments surrounding the behavioural and cognitive separateness of the species, by the increasing appreciation of variation in the early African H. sapiens record and by new DNA evidence of hybridization with extinct species. This study synthesizes thinking on the fossils, archaeology and underlying evolutionary models of the last several decades with recent DNA results from both H. sapiens and fossil species. It is concluded that, although it may not be possible or even desirable to cleanly partition out a homogenous morphological description of recent H. sapiens in the fossil record, there are key, distinguishing morphological traits in the cranium, dentition and pelvis that can be usefully employed to diagnose the H. sapiens lineage. Increasing advances in retrieving and understanding relevant genetic data provide a complementary and perhaps potentially even more fruitful means of characterizing the differences between H. sapiens and its close relatives.

  3. Modeling the hook depth distribution of pelagic longlining in the equatorial area of Indian Ocean

    NASA Astrophysics Data System (ADS)

    Song, Liming; Li, Jie; Gao, Panfeng; Zhou, Ji; Xu, Liuxiong

    2012-12-01

    A survey was conducted in the equatorial area of Indian Ocean for a better understanding of the dynamics of hook depth distribution of pelagic longline fishery. We determined the relationship between hook depth and vertical shear of current coefficiency, wind speed, hook position code, sine of wind angle, sine of angle of attack and weight of messenger weight. We identified the hook depth models by the analysis of covariance with a general linear model. The results showed that the wind effect on the hook depth can be ignored from October to November in the survey area; the surface current effect on the hook depth can be ignored; the equatorial undercurrent is the key factor for the hook depth in Indian Ocean; and there is a negative correlation between the hook depth and vertical shear of current and angle of attack. It was also found that the deeper the hook was set, the higher hook depth shoaling was. The proposed model improves the accuracy of the prediction of hook depth, which can be used to estimate the vertical distribution of pelagic fish in water column.

  4. The Great Fossil Fiasco: Teaching about Peer Review.

    ERIC Educational Resources Information Center

    Gift, Nancy; Krasny, Marianne

    2003-01-01

    Describes a lesson that engages middle school students in learning about peer review. Uses the article "Archaeoraptor Fossil Trail," which was published in the November, 1999 issue of "National Geographic" as an example of a real life story of how peer review forces scientists to critically re-examine a fossil discovery. (SOE)

  5. Survival of fossils under extreme shocks induced by hypervelocity impacts.

    PubMed

    Burchell, M J; McDermott, K H; Price, M C; Yolland, L J

    2014-08-28

    Experimental data are shown for survival of fossilized diatoms undergoing shocks in the GPa range. The results were obtained from hypervelocity impact experiments which fired fossilized diatoms frozen in ice into water targets. After the shots, the material recovered from the target water was inspected for diatom fossils. Nine shots were carried out, at speeds from 0.388 to 5.34 km s(-1), corresponding to mean peak pressures of 0.2-19 GPa. In all cases, fragmented fossilized diatoms were recovered, but both the mean and the maximum fragment size decreased with increasing impact speed and hence peak pressure. Examples of intact diatoms were found after the impacts, even in some of the higher speed shots, but their frequency and size decreased significantly at the higher speeds. This is the first demonstration that fossils can survive and be transferred from projectile to target in hypervelocity impacts, implying that it is possible that, as suggested by other authors, terrestrial rocks ejected from the Earth by giant impacts from space, and which then strike the Moon, may successfully transfer terrestrial fossils to the Moon.

  6. Survival of fossils under extreme shocks induced by hypervelocity impacts

    PubMed Central

    Burchell, M. J.; McDermott, K. H.; Price, M. C.; Yolland, L. J.

    2014-01-01

    Experimental data are shown for survival of fossilized diatoms undergoing shocks in the GPa range. The results were obtained from hypervelocity impact experiments which fired fossilized diatoms frozen in ice into water targets. After the shots, the material recovered from the target water was inspected for diatom fossils. Nine shots were carried out, at speeds from 0.388 to 5.34 km s−1, corresponding to mean peak pressures of 0.2–19 GPa. In all cases, fragmented fossilized diatoms were recovered, but both the mean and the maximum fragment size decreased with increasing impact speed and hence peak pressure. Examples of intact diatoms were found after the impacts, even in some of the higher speed shots, but their frequency and size decreased significantly at the higher speeds. This is the first demonstration that fossils can survive and be transferred from projectile to target in hypervelocity impacts, implying that it is possible that, as suggested by other authors, terrestrial rocks ejected from the Earth by giant impacts from space, and which then strike the Moon, may successfully transfer terrestrial fossils to the Moon. PMID:25071234

  7. 76 FR 23964 - Fisheries in the Western Pacific; Pelagic Fisheries; Purse Seine Prohibited Areas Around American...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-04-29

    ... animals, such as pelagic fishes and sea turtles, tend to congregate to naturally-occurring floating... American Samoa enclosed by straight lines connecting the following coordinates: Point S. latitude W. longitude AS-3-A 11[deg]12[min] 172[deg]18[min] AS-3-B 12[deg]12[min] 169[deg]56[min] and from Point AS-3-A...

  8. Likelihood of Tree Topologies with Fossils and Diversification Rate Estimation.

    PubMed

    Didier, Gilles; Fau, Marine; Laurin, Michel

    2017-11-01

    Since the diversification process cannot be directly observed at the human scale, it has to be studied from the information available, namely the extant taxa and the fossil record. In this sense, phylogenetic trees including both extant taxa and fossils are the most complete representations of the diversification process that one can get. Such phylogenetic trees can be reconstructed from molecular and morphological data, to some extent. Among the temporal information of such phylogenetic trees, fossil ages are by far the most precisely known (divergence times are inferences calibrated mostly with fossils). We propose here a method to compute the likelihood of a phylogenetic tree with fossils in which the only considered time information is the fossil ages, and apply it to the estimation of the diversification rates from such data. Since it is required in our computation, we provide a method for determining the probability of a tree topology under the standard diversification model. Testing our approach on simulated data shows that the maximum likelihood rate estimates from the phylogenetic tree topology and the fossil dates are almost as accurate as those obtained by taking into account all the data, including the divergence times. Moreover, they are substantially more accurate than the estimates obtained only from the exact divergence times (without taking into account the fossil record). We also provide an empirical example composed of 50 Permo-Carboniferous eupelycosaur (early synapsid) taxa ranging in age from about 315 Ma (Late Carboniferous) to 270 Ma (shortly after the end of the Early Permian). Our analyses suggest a speciation (cladogenesis, or birth) rate of about 0.1 per lineage and per myr, a marginally lower extinction rate, and a considerable hidden paleobiodiversity of early synapsids. [Extinction rate; fossil ages; maximum likelihood estimation; speciation rate.]. © The Author(s) 2017. Published by Oxford University Press, on behalf of the Society

  9. Life in changing fluids: A critical appraisal of swimming animals before the Cambrian.

    PubMed

    Gold, David

    2018-05-03

    Molecular and fossil data place the initial diversification of animals in the Neoproterozoic, though there remains too much enough uncertainty to produce an exact chronology. This is unfortunate, as the Neoproterozoic represents a period of intense climate change, including multiple global glaciation events as well changes to ocean chemistry and oxygen content. Several authors have suggested that the coevolution of animals and their environment was tightly coupled, but such hypotheses rest on the presence of swimming (pelagic) species. In this paper, I review the evidence for pelagic animals during the Neoproterozoic. I conclude that there are very few groups of planktotrophic swimming animals that were likely to have existed at this time, with the possible exception of medusozoan cnidarians (jellyfish). Ultimately, hypotheses connecting Earth and animal evolution in the Neoproterozoic need to be tested with more geochemical work, fossil discoveries, and refinement of molecular clocks targeted on the relevant groups.

  10. Fossil human remains from Bolomor Cave (Valencia, Spain).

    PubMed

    Arsuaga, Juan Luis; Fernández Peris, Josep; Gracia-Téllez, Ana; Quam, Rolf; Carretero, José Miguel; Barciela González, Virginia; Blasco, Ruth; Cuartero, Felipe; Sañudo, Pablo

    2012-05-01

    Systematic excavations carried out since 1989 at Bolomor Cave have led to the recovery of four Pleistocene human fossil remains, consisting of a fibular fragment, two isolated teeth, and a nearly complete adult parietal bone. All of these specimens date to the late Middle and early Late Pleistocene (MIS 7-5e). The fibular fragment shows thick cortical bone, an archaic feature found in non-modern (i.e. non-Homo sapiens) members of the genus Homo. Among the dental remains, the lack of a midtrigonid crest in the M(1) represents a departure from the morphology reported for the majority of Neandertal specimens, while the large dimensions and pronounced shoveling of the marginal ridges in the C(1) are similar to other European Middle and late Pleistocene fossils. The parietal bone is very thick, with dimensions that generally fall above Neandertal fossils and resemble more closely the Middle Pleistocene Atapuerca (SH) adult specimens. Based on the presence of archaic features, all the fossils from Bolomor are attributed to the Neandertal evolutionary lineage. Copyright © 2012 Elsevier Ltd. All rights reserved.

  11. 76 FR 3587 - Standards of Performance for Fossil-Fuel-Fired, Electric Utility, Industrial-Commercial...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-01-20

    ... Standards of Performance for Fossil-Fuel-Fired, Electric Utility, Industrial-Commercial-Institutional, and... Fossil fuel-fired electric utility steam generating units. Federal Government 22112 Fossil fuel-fired... 22112 Fossil fuel-fired electric utility steam generating units owned by municipalities. 921150 Fossil...

  12. New Ediacara fossils preserved in marine limestone and their ecological implications

    PubMed Central

    Chen, Zhe; Zhou, Chuanming; Xiao, Shuhai; Wang, Wei; Guan, Chengguo; Hua, Hong; Yuan, Xunlai

    2014-01-01

    Ediacara fossils are central to our understanding of animal evolution on the eve of the Cambrian explosion, because some of them likely represent stem-group marine animals. However, some of the iconic Ediacara fossils have also been interpreted as terrestrial lichens or microbial colonies. Our ability to test these hypotheses is limited by a taphonomic bias that most Ediacara fossils are preserved in sandstones and siltstones. Here we report several iconic Ediacara fossils and an annulated tubular fossil (reconstructed as an erect epibenthic organism with uniserial arranged modular units), from marine limestone of the 551–541 Ma Dengying Formation in South China. These fossils significantly expand the ecological ranges of several key Ediacara taxa and support that they are marine organisms rather than terrestrial lichens or microbial colonies. Their close association with abundant bilaterian burrows also indicates that they could tolerate and may have survived moderate levels of bioturbation. PMID:24566959

  13. The Fossil Fuel Divestment Movement: An Ethical Dilemma for the Geosciences?

    NASA Astrophysics Data System (ADS)

    Greene, C. H.; Kammen, D. M.

    2014-12-01

    For over 200 years, fossil fuels have been the basis for an industrial revolution that has delivered a level of prosperity to modern society unimaginable during the previous 5000 years of human civilization. However, society's dependence on fossil fuels is coming to an end for two reasons. The first reason is because our fossil fuel reserves are running out, oil in this century, natural gas during the next century, and coal a few centuries later. The second reason is because fossil fuels are having a devastating impact on the habitability of our planet, disrupting our climate system and acidifying our oceans. So the question is not whether we will discontinue using fossil fuels, but rather whether we will stop using them before they do irreparable damage to the Earth's life-support systems. Within our geoscience community, climate scientists have determined that a majority of existing fossil fuel reserves must remain unburned if dangerous climate change and ocean acidification are to be avoided. In contrast, Exxon-Mobil, Shell, and other members of the fossil fuel industry are pursuing a business model that assumes all of their reserves will be burned and will not become stranded assets. Since the geosciences have had a long and mutually beneficial relationship with the fossil fuel industry, this inherent conflict between climate science and industrial interests presents an ethical dilemma for many geoscientists. This conflict is further heightened by the fossil fuel divestment movement, which is underway at over 400 college and university campuses around the world. This presentation will explore some of the ethical and financial issues being raised by the divestment movement from a geoscientist's perspective.

  14. THE XMM CLUSTER SURVEY: THE STELLAR MASS ASSEMBLY OF FOSSIL GALAXIES

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Harrison, Craig D.; Miller, Christopher J.; Richards, Joseph W.

    This paper presents both the result of a search for fossil systems (FSs) within the XMM Cluster Survey and the Sloan Digital Sky Survey and the results of a study of the stellar mass assembly and stellar populations of their fossil galaxies. In total, 17 groups and clusters are identified at z < 0.25 with large magnitude gaps between the first and fourth brightest galaxies. All the information necessary to classify these systems as fossils is provided. For both groups and clusters, the total and fractional luminosity of the brightest galaxy is positively correlated with the magnitude gap. The brightestmore » galaxies in FSs (called fossil galaxies) have stellar populations and star formation histories which are similar to normal brightest cluster galaxies (BCGs). However, at fixed group/cluster mass, the stellar masses of the fossil galaxies are larger compared to normal BCGs, a fact that holds true over a wide range of group/cluster masses. Moreover, the fossil galaxies are found to contain a significant fraction of the total optical luminosity of the group/cluster within 0.5 R{sub 200}, as much as 85%, compared to the non-fossils, which can have as little as 10%. Our results suggest that FSs formed early and in the highest density regions of the universe and that fossil galaxies represent the end products of galaxy mergers in groups and clusters.« less

  15. Pelagic effects of offshore wind farm foundations in the stratified North Sea

    NASA Astrophysics Data System (ADS)

    Floeter, Jens; van Beusekom, Justus E. E.; Auch, Dominik; Callies, Ulrich; Carpenter, Jeffrey; Dudeck, Tim; Eberle, Sabine; Eckhardt, André; Gloe, Dominik; Hänselmann, Kristin; Hufnagl, Marc; Janßen, Silke; Lenhart, Hermann; Möller, Klas Ove; North, Ryan P.; Pohlmann, Thomas; Riethmüller, Rolf; Schulz, Sabrina; Spreizenbarth, Stefan; Temming, Axel; Walter, Bettina; Zielinski, Oliver; Möllmann, Christian

    2017-08-01

    A recent increase in the construction of Offshore Wind Farms (OWFs) has initiated numerous environmental impact assessments and monitoring programs. These focus on sea mammals, seabirds, benthos or demersal fish, but generally ignore any potential effects OWFs may have on the pelagic ecosystem. The only work on the latter has been through modelling analyses, which predict localised impacts like enhanced vertical mixing leading to a decrease in seasonal stratification, as well as shelf-wide changes of tidal amplitudes. Here we provide for the first-time empirical bio-physical data from an OWF. The data were obtained by towing a remotely operated vehicle (TRIAXUS ROTV) through two non-operating OWFs in the summer stratified North Sea. The undulating TRIAXUS transects provided high-resolution CTD data accompanied by oxygen and chlorophyll-a measurements. We provide empirical indication that vertical mixing is increased within the OWFs, leading to a doming of the thermocline and a subsequent transport of nutrients into the surface mixed layer (SML). Nutrients were taken up rapidly because underwater photosynthetically active radiation (PAR) enabled net primary production in the entire water column, especially within submesoscale chlorophyll-a pillars that were observed at regular intervals within the OWF regions. Video Plankton Recorder (VPR) images revealed distinct meroplankton distribution patterns in a copepod-dominated plankton community. Hydroacoustic records did not show any OWF effects on the distribution of pelagic fish. The results of a pre-OWF survey show however, that it is difficult to fully separate the anthropogenic impacts from the natural variability.

  16. Dating Fossil Pollen: A Simulation.

    ERIC Educational Resources Information Center

    Sheridan, Philip

    1992-01-01

    Describes a hands-on simulation in which students determine the age of "fossil" pollen samples based on the pollen types present when examined microscopically. Provides instructions for the preparation of pollen slides. (MDH)

  17. Bill Lowrie In The Apennines U Reading - The Pelagic Record of Geomagnetic Reversals

    NASA Astrophysics Data System (ADS)

    Alvarez, Walter

    Twenty five years ago, Bill Lowrie and I, along with Mike Arthur, Al Fischer, Gio- vanni Napoleone, Isabella Premoli Silva and Bill Roggenthen, published a set of five papers in the Geological Society of America Bulletin (March 1977), reporting a re- markable new source of information on the geomagnetic polarity time scale. The re- versal sequence was already known back to the Late Cretaceous on the basis of marine magnetic anomalies, but only as a sequence of longer and shorter polarity intervals, a kind of fingerprint with almost no age calibration. At Gubbio, in the Umbrian Apen- nines of Italy, we discovered that the polarity intervals are also recorded in pelagic limestones, deposited quietly at moderate oceanic depths at rates of order 10 m/Myr. and these limestones are literally made of fossils, notably the planktic foraminifera which are the best age correlation tool for the last 100 Myr. The reversal sequence was now datable. You can make a discovery like this either by looking for it, as Al Fischer did U hoping ° that such a record would be present and waiting until magnetometers improved enough to make it possible to measure these very weakly magnetic rocks U or by stumbling ° on it as Bill and I did. We went to the Apennines hoping to measure paleomagnetically a tectonic rotation of the Italian crust. Digital spinner magnetometers had just become available and Bill found that he could measure the remanence of the Apennine pelagic limestones I had been studying in the field. Tectonic rotation of the Italian crust turned out to be very difficult to detect, because interbed slip was a major complication. But we accidentally sampled both normal and reversed beds in the Scaglia rossa limestone on our first trip, and back in the lab we recognized that we had a polarity record in front of us. The microfossils made it a datable record, which raised great excitement among our colleagues at Lamont, where sea-floor magnetic reversals were the key to tectonic

  18. Fossil nutlets of Boraginaceae from the continental Eocene of Hamada of Méridja (southwestern Algeria): The first fossil of the Borage family in Africa.

    PubMed

    Hammouda, Sid Ahmed; Weigend, Maximilian; Mebrouk, Fateh; Chacón, Juliana; Bensalah, Mustapha; Ensikat, Hans-Jürgen; Adaci, Mohammed

    2015-12-01

    The Paleogene deposits of the Hamada of Méridja, southwestern Algeria, are currently dated as lower-to-middle Eocene in age based on fossil gastropods and charophytes. Here we report the presence of fruits that can be assigned to the Boraginaceae s.str., apparently representing the first fossil record for this family in Africa, shedding new light on the historical biogeography of this group. Microscopic studies of the fossil nutlets were carried out and compared to extant Boraginaceae nutlets, and to types reported in the literature for this family. The fossils are strikingly similar in general size and morphology, particularly in the finer details of the attachment scar and ornamentation, to nutlets of extant representatives of the Boraginaceae tribe Echiochileae, and especially the genus Ogastemma. We believe that these nutlets represent an extinct member of this lineage. The Ogastemma-like fossils indicate that the Echiochileae, which are most diverse in northern Africa and southwestern Asia, have a long history in this region, dating back to the Eocene. This tribe corresponds to the basal-most clade in Boraginaceae s.str., and the fossils described here agree well with an assumed African origin of the family and the Boraginales I, providing an important additional calibration point for dating the phylogenies of this clade. © 2015 Botanical Society of America.

  19. Spatial, Temporal, and Habitat-Related Variation in Abundance of Pelagic Fishes in the Gulf of Mexico: Potential Implications of the Deepwater Horizon Oil Spill

    PubMed Central

    Rooker, Jay R.; Kitchens, Larissa L.; Dance, Michael A.; Wells, R. J. David; Falterman, Brett; Cornic, Maëlle

    2013-01-01

    Time-series data collected over a four-year period were used to characterize patterns of abundance for pelagic fishes in the northern Gulf of Mexico (GoM) before (2007–2009) and after (2010) the Deepwater Horizon oil spill. Four numerically dominant pelagic species (blackfin tuna, blue marlin, dolphinfish, and sailfish) were included in our assessment, and larval density of each species was lower in 2010 than any of the three years prior to the oil spill, although larval abundance in 2010 was often statistically similar to other years surveyed. To assess potential overlap between suitable habitat of pelagic fish larvae and surface oil, generalized additive models (GAMs) were developed to evaluate the influence of ocean conditions on the abundance of larvae from 2007–2009. Explanatory variables from GAMs were then linked to environmental data from 2010 to predict the probability of occurrence for each species. The spatial extent of surface oil overlapped with early life habitat of each species, possibly indicating that the availability of high quality habitat was affected by the DH oil spill. Shifts in the distribution of spawning adults is another factor known to influence the abundance of larvae, and the spatial occurrence of a model pelagic predator (blue marlin) was characterized over the same four-year period using electronic tags. The spatial extent of oil coincided with areas used by adult blue marlin from 2007–2009, and the occurrence of blue marlin in areas impacted by the DH oil spill was lower in 2010 relative to pre-spill years. PMID:24130759

  20. 13C-isotope analyses reveal that chemolithoautotrophic Gamma- and Epsilonproteobacteria feed a microbial food web in a pelagic redoxcline of the central Baltic Sea.

    PubMed

    Glaubitz, Sabine; Lueders, Tillmann; Abraham, Wolf-Rainer; Jost, Günter; Jürgens, Klaus; Labrenz, Matthias

    2009-02-01

    Marine pelagic redoxclines are zones of high dark CO(2) fixation rates, which can correspond up to 30% of the surface primary production. However, despite this significant contribution to the pelagic carbon cycle, the identity of most chemolithoautotrophic organisms is still unknown. Therefore, the aim of this study was to directly link the dark CO(2) fixation capacity of a pelagic redoxcline in the central Baltic Sea (Landsort Deep) with the identity of the main chemolithoautotrophs involved. Our approach was based on the analysis of natural carbon isotope signatures in fatty acid methyl esters (FAMEs) and on measurements of CO(2) incorporation in (13)C-bicarbonate pulse experiments. The incorporation of (13)C into chemolithoautotrophic cells was investigated by rRNA-based stable isotope probing (RNA-SIP) and FAME analysis after incubation for 24 and 72 h under in situ conditions. Our results demonstrated that fatty acids indicative of Proteobacteria were significantly enriched in (13)C slightly below the chemocline. RNA-SIP analyses revealed that two different Gammaproteobacteria and three closely related Epsilonproteobacteria of the Sulfurimonas cluster were active dark CO(2)-fixing microorganisms, with a time-dependent community shift between these groups. Labelling of Archaea was not detectable, but after 72 h of incubation the (13)C-label had been transferred to a potentially bacterivorous ciliate related to Euplotes sp. Thus, RNA-SIP provided direct evidence for the contribution of chemolithoautotrophic production to the microbial food web in this marine pelagic redoxcline, emphasizing the importance of dark CO(2)-fixing Proteobacteria within this habitat.

  1. 75 FR 38937 - Fisheries of the Exclusive Economic Zone Off Alaska; Northern Rockfish and Pelagic Shelf Rockfish...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2010-07-07

    .... 0910131362-0087-02] RIN 0648-XX34 Fisheries of the Exclusive Economic Zone Off Alaska; Northern Rockfish and Pelagic Shelf Rockfish for Trawl Catcher Vessels Participating in the Entry Level Rockfish Fishery in the... participating in the entry level rockfish fishery in the Central Regulatory Area of the Gulf of Alaska (GOA...

  2. RX J1548.9+0851, a fossil cluster?

    NASA Astrophysics Data System (ADS)

    Eigenthaler, P.; Zeilinger, W. W.

    2012-04-01

    Context. Fossil galaxy groups are spatially extended X-ray sources with X-ray luminosities above L{X, bol ≥ 1042 h50-2} erg s-1 and a central elliptical galaxy dominating the optical, the second-brightest galaxy being at least 2 mag fainter in the R band. Whether these systems are a distinct class of objects resulting from exceptional formation and evolution histories is still unclear, mainly due to the small number of objects studied so far, mostly lacking spectroscopy of group members for group membership confirmation and a detailed kinematical analysis. Aims: To complement the scarce sample of spectroscopically studied fossils down to their faint galaxy populations, the fossil candidate RX J1548.9+0851 (z = 0.072) is studied in this work. Our results are compared with existing data from fossils in the literature. Methods: We use ESO VLT VIMOS multi-object spectroscopy to determine redshifts of the faint galaxy population and study the luminosity-weighted dynamics and luminosity function of the system. The full-spectrum fitting package ULySS is used to determine ages and metallicities of group members. VIMOS imaging data are used to study the morphology of the central elliptical. Results: We identify 40 group members spectroscopically within the central 300 kpc of the system and find 31 additional redshifts from the literature, resulting in a total number of 54 spectroscopically confirmed group members within 1 Mpc. RX J1548.9+0851 is made up of two bright ellipticals in the central region with a magnitude gap of Δm1,2 = 1.34 in the SDSS r' band leaving the definition of RX J1548.9+0851 being a fossil to the assumption of the virial radius. We find a luminosity-weighted velocity dispersion of 568 km s-1 and a mass of 2.5 × 1014 M⊙ for the system confirming previous studies that revealed fossils to be massive. An average mass-to-light ratio of M/L 400 M⊙/L⊙ is derived from the SDSS g', r', and i' bands. The central elliptical is well-fitted by a pure de

  3. Latitudinal and photic effects on diel foraging and predation risk in freshwater pelagic ecosystems

    USGS Publications Warehouse

    Hansen, Adam G.; Beauchamp, David A.

    2014-01-01

    summer into autumn. 4. Model results show that diel-seasonal foraging and predation risk in freshwater pelagic ecosystems changes considerably with latitude, turbidity and cloud cover. These changes alter the structure of pelagic predator–prey interactions, and in turn, the broader role of pelagic consumers in habitat coupling in lakes. 

  4. Increased Oxygenation of the Oceans Since the Mid-Cenozoic as Constrained by Cr/Co and Os/Ir Ratios in Oxic Pelagic Sediments

    NASA Astrophysics Data System (ADS)

    Hu, M.; Lee, C.

    2005-12-01

    In terms of redox, the marine sediments can be roughly divided into anoxic to suboxic sediments on the margins and oxic sediments in pelagic (open ocean) environments. The relative amounts of anoxic/suboxic sediments being deposited at any given time could be related to biological productivity and/or the efficiency of the ocean circulation system. How the depositional area of anoxic/suboxic deposition has changed through time is thus of concern. One way to track redox conditions is to investigate variations in the concentrations of redox sensitive trace metals. Most studies along these lines have focused on anoxic sediments. However, one problem with using anoxic sediments to study the global oceans is that such sediments are typically deposited in somewhat isolated basins, whose redox conditions may vary from basin to basin. An alternative approach, taken here, is to examine redox-sensitive elemental ratios in oxic pelagic sediments. This is motivated by the fact that pelagic sediments are more likely to reflect average ocean chemistry. In addition, the redox-sensitive metal contents of oxic sediments represent the complement to anoxic sediments. Choosing an appropriate redox-sensitive elemental ratio which eliminates dilution/concentration effects, requires the identification of trace metals that are preferentially precipitated in oxic conditions and those precipitated in more reducing conditions. Overall elemental behaviors were estimated by comparing hydrogenous or authigenic burial fluxes of various trace metals at given pelagic ODP sites to global riverine input fluxes. If the pelagic burial fluxes of a given element are significantly smaller than the riverine input flux, other burial outputs are implied, and it is hypothesized here that this element may precipitate in reducing conditions, such as in oceanic margin. If, on the other hand, the pelagic burial flux is equal to or greater than the riverine input flux, the implication is that oxic pelagic

  5. Chemical, experimental, and morphological evidence for diagenetically altered melanin in exceptionally preserved fossils.

    PubMed

    Colleary, Caitlin; Dolocan, Andrei; Gardner, James; Singh, Suresh; Wuttke, Michael; Rabenstein, Renate; Habersetzer, Jörg; Schaal, Stephan; Feseha, Mulugeta; Clemens, Matthew; Jacobs, Bonnie F; Currano, Ellen D; Jacobs, Louis L; Sylvestersen, Rene Lyng; Gabbott, Sarah E; Vinther, Jakob

    2015-10-13

    In living organisms, color patterns, behavior, and ecology are closely linked. Thus, detection of fossil pigments may permit inferences about important aspects of ancient animal ecology and evolution. Melanin-bearing melanosomes were suggested to preserve as organic residues in exceptionally preserved fossils, retaining distinct morphology that is associated with aspects of original color patterns. Nevertheless, these oblong and spherical structures have also been identified as fossilized bacteria. To date, chemical studies have not directly considered the effects of diagenesis on melanin preservation, and how this may influence its identification. Here we use time-of-flight secondary ion mass spectrometry to identify and chemically characterize melanin in a diverse sample of previously unstudied extant and fossil taxa, including fossils with notably different diagenetic histories and geologic ages. We document signatures consistent with melanin preservation in fossils ranging from feathers, to mammals, to amphibians. Using principal component analyses, we characterize putative mixtures of eumelanin and phaeomelanin in both fossil and extant samples. Surprisingly, both extant and fossil amphibians generally exhibit melanosomes with a mixed eumelanin/phaeomelanin composition rather than pure eumelanin, as assumed previously. We argue that experimental maturation of modern melanin samples replicates diagenetic chemical alteration of melanin observed in fossils. This refutes the hypothesis that such fossil microbodies could be bacteria, and demonstrates that melanin is widely responsible for the organic soft tissue outlines in vertebrates found at exceptional fossil localities, thus allowing for the reconstruction of certain aspects of original pigment patterns.

  6. Chemical, experimental, and morphological evidence for diagenetically altered melanin in exceptionally preserved fossils

    PubMed Central

    Colleary, Caitlin; Dolocan, Andrei; Gardner, James; Singh, Suresh; Wuttke, Michael; Rabenstein, Renate; Habersetzer, Jörg; Schaal, Stephan; Feseha, Mulugeta; Clemens, Matthew; Jacobs, Bonnie F.; Currano, Ellen D.; Jacobs, Louis L.; Sylvestersen, Rene Lyng; Gabbott, Sarah E.; Vinther, Jakob

    2015-01-01

    In living organisms, color patterns, behavior, and ecology are closely linked. Thus, detection of fossil pigments may permit inferences about important aspects of ancient animal ecology and evolution. Melanin-bearing melanosomes were suggested to preserve as organic residues in exceptionally preserved fossils, retaining distinct morphology that is associated with aspects of original color patterns. Nevertheless, these oblong and spherical structures have also been identified as fossilized bacteria. To date, chemical studies have not directly considered the effects of diagenesis on melanin preservation, and how this may influence its identification. Here we use time-of-flight secondary ion mass spectrometry to identify and chemically characterize melanin in a diverse sample of previously unstudied extant and fossil taxa, including fossils with notably different diagenetic histories and geologic ages. We document signatures consistent with melanin preservation in fossils ranging from feathers, to mammals, to amphibians. Using principal component analyses, we characterize putative mixtures of eumelanin and phaeomelanin in both fossil and extant samples. Surprisingly, both extant and fossil amphibians generally exhibit melanosomes with a mixed eumelanin/phaeomelanin composition rather than pure eumelanin, as assumed previously. We argue that experimental maturation of modern melanin samples replicates diagenetic chemical alteration of melanin observed in fossils. This refutes the hypothesis that such fossil microbodies could be bacteria, and demonstrates that melanin is widely responsible for the organic soft tissue outlines in vertebrates found at exceptional fossil localities, thus allowing for the reconstruction of certain aspects of original pigment patterns. PMID:26417094

  7. Emissions Scenarios and Fossil-fuel Peaking

    NASA Astrophysics Data System (ADS)

    Brecha, R.

    2008-12-01

    Intergovernmental Panel on Climate Change (IPCC) emissions scenarios are based on detailed energy system models in which demographics, technology and economics are used to generate projections of future world energy consumption, and therefore, of greenhouse gas emissions. Built into the assumptions for these scenarios are estimates for ultimately recoverable resources of various fossil fuels. There is a growing chorus of critics who believe that the true extent of recoverable fossil resources is much smaller than the amounts taken as a baseline for the IPCC scenarios. In a climate optimist camp are those who contend that "peak oil" will lead to a switch to renewable energy sources, while others point out that high prices for oil caused by supply limitations could very well lead to a transition to liquid fuels that actually increase total carbon emissions. We examine a third scenario in which high energy prices, which are correlated with increasing infrastructure, exploration and development costs, conspire to limit the potential for making a switch to coal or natural gas for liquid fuels. In addition, the same increasing costs limit the potential for expansion of tar sand and shale oil recovery. In our qualitative model of the energy system, backed by data from short- and medium-term trends, we have a useful way to gain a sense of potential carbon emission bounds. A bound for 21st century emissions is investigated based on two assumptions: first, that extractable fossil-fuel resources follow the trends assumed by "peak oil" adherents, and second, that little is done in the way of climate mitigation policies. If resources, and perhaps more importantly, extraction rates, of fossil fuels are limited compared to assumptions in the emissions scenarios, a situation can arise in which emissions are supply-driven. However, we show that even in this "peak fossil-fuel" limit, carbon emissions are high enough to surpass 550 ppm or 2°C climate protection guardrails. Some

  8. Fossil fuels in the 21st century.

    PubMed

    Lincoln, Stephen F

    2005-12-01

    An overview of the importance of fossil fuels in supplying the energy requirements of the 21st century, their future supply, and the impact of their use on global climate is presented. Current and potential alternative energy sources are considered. It is concluded that even with substantial increases in energy derived from other sources, fossil fuels will remain a major energy source for much of the 21st century and the sequestration of CO2 will be an increasingly important requirement.

  9. Shallow water mud-mounds of the Early Devonian Buchan Group, East Gippsland, Australia

    NASA Astrophysics Data System (ADS)

    Tosolini, A.-M. P.; Wallace, M. W.; Gallagher, S. J.

    2012-12-01

    The Lower Devonian Rocky Camp Member of the Murrindal Limestone, Buchan Group of southeastern Australia consists of a series of carbonate mud-mounds and smaller lagoonal bioherms. The Rocky Camp mound is the best exposed of the mud-mounds and has many characteristics in common with Waulsortian (Carboniferous) mounds. Detailed paleoecological and sedimentological studies indicate that the mound initially accumulated in the photic zone, in contrast to most of the previously recorded mud-mounds. Five facies are present in the mud-mound: a Dasycladacean Wackestone Facies at the base of the mound represents a moderate energy, shallow water bank environment within the photic zone. A Crinioidal Wackestone Facies was deposited in a laterally equivalent foreslope setting. A Poriferan-Crinoidal Mudstone Facies developed in a quiet, deeper water, lee-side mound setting associated with a minor relative sea-level rise. A Stromatoporoid-Coralline Packstone Facies in the upper part of the mound deposited in a high-energy, fair-weather wave base, mound-front environment. The crest of the mound is represented by a Crinoidal-Receptaculitid Packstone Facies indicative of a moderate-energy mound-top environment in the photic zone, sheltered by the mound-front stromatoporoid-coral communities. A mound flank facies is present on the southern side of the mound and this consists of high-energy crinoidal grainstones. Mud-mound deposition was terminated by a transgression that deposited dark gray, fossil-poor marl of the overlying Taravale Formation. The Rocky Camp mound appears to have originated in shallow water photic zone conditions and grew into a high-energy environment, with the mound being eventually colonized by corals and stromatoporoids. The indications of a high-energy environment during later mound growth (growth form of colonial metazoans and grainstones of the flanking facies) suggest that the micrite in the mound was autochthonous and implies the presence of an energy

  10. Changing Concepts of the Nature and Significance of Fossils.

    ERIC Educational Resources Information Center

    Gregory, Joseph T.

    1984-01-01

    Provides information on early written accounts of fossils and contrasts historical phases regarding their organic or inorganic origin. Topic areas discussed include the deluge as a stimulant to geological investigation, stratigraphic sequences and earth history, extinction, faunal succession and organic evolution, and fossil evidence of…

  11. Social aggregation in the pelagic zone with special reference to fish and invertebrates.

    PubMed

    Ritz, David A; Hobday, Alistair J; Montgomery, John C; Ward, Ashley J W

    2011-01-01

    Aggregations of organisms, ranging from zooplankton to whales, are an extremely common phenomenon in the pelagic zone; perhaps the best known are fish schools. Social aggregation is a special category that refers to groups that self-organize and maintain cohesion to exploit benefits such as protection from predators, and location and capture of resources more effectively and with greater energy efficiency than could a solitary individual. In this review we explore general aggregation principles, with specific reference to pelagic organisms; describe a range of new technologies either designed for studying aggregations or that could potentially be exploited for this purpose; report on the insights gained from theoretical modelling; discuss the relationship between social aggregation and ocean management; and speculate on the impact of climate change. Examples of aggregation occur in all animal phyla. Among pelagic organisms, it is possible that repeated co-occurrence of stable pairs of individuals, which has been established for some schooling fish, is the likely precursor leading to networks of social interaction and more complex social behaviour. Social network analysis has added new insights into social behaviour and allows us to dissect aggregations and to examine how the constituent individuals interact with each other. This type of analysis is well advanced in pinnipeds and cetaceans, and work on fish is progressing. Detailed three-dimensional analysis of schools has proved to be difficult, especially at sea, but there has been some progress recently. The technological aids for studying social aggregation include video and acoustics, and have benefited from advances in digitization, miniaturization, motion analysis and computing power. New techniques permit three-dimensional tracking of thousands of individual animals within a single group which has allowed novel insights to within-group interactions. Approaches using theoretical modelling of aggregations have

  12. Biofilm and Diatom Succession on Polyethylene (PE) and Biodegradable Plastic Bags in Two Marine Habitats: Early Signs of Degradation in the Pelagic and Benthic Zone?

    PubMed Central

    Laforsch, Christian; Weber, Miriam

    2015-01-01

    The production of biodegradable plastic is increasing. Given the augmented littering of these products an increasing input into the sea is expected. Previous laboratory experiments have shown that degradation of plastic starts within days to weeks. Little is known about the early composition and activity of biofilms found on biodegradable and conventional plastic debris and its correlation to degradation in the marine environment. In this study we investigated the early formation of biofilms on plastic shopper bags and its consequences for the degradation of plastic. Samples of polyethylene and biodegradable plastic were tested in the Mediterranean Sea for 15 and 33 days. The samples were distributed equally to a shallow benthic (sedimentary seafloor at 6 m water depth) and a pelagic habitat (3 m water depth) to compare the impact of these different environments on fouling and degradation. The amount of biofilm increased on both plastic types and in both habitats. The diatom abundance and diversity differed significantly between the habitats and the plastic types. Diatoms were more abundant on samples from the pelagic zone. We anticipate that specific surface properties of the polymer types induced different biofilm communities on both plastic types. Additionally, different environmental conditions between the benthic and pelagic experimental site such as light intensity and shear forces may have influenced unequal colonisation between these habitats. The oxygen production rate was negative for all samples, indicating that the initial biofilm on marine plastic litter consumes oxygen, regardless of the plastic type or if exposed in the pelagic or the benthic zone. Mechanical tests did not reveal degradation within one month of exposure. However, scanning electron microscopy (SEM) analysis displayed potential signs of degradation on the plastic surface, which differed between both plastic types. This study indicates that the early biofilm formation and composition

  13. Biofilm and Diatom Succession on Polyethylene (PE) and Biodegradable Plastic Bags in Two Marine Habitats: Early Signs of Degradation in the Pelagic and Benthic Zone?

    PubMed

    Eich, Andreas; Mildenberger, Tobias; Laforsch, Christian; Weber, Miriam

    2015-01-01

    The production of biodegradable plastic is increasing. Given the augmented littering of these products an increasing input into the sea is expected. Previous laboratory experiments have shown that degradation of plastic starts within days to weeks. Little is known about the early composition and activity of biofilms found on biodegradable and conventional plastic debris and its correlation to degradation in the marine environment. In this study we investigated the early formation of biofilms on plastic shopper bags and its consequences for the degradation of plastic. Samples of polyethylene and biodegradable plastic were tested in the Mediterranean Sea for 15 and 33 days. The samples were distributed equally to a shallow benthic (sedimentary seafloor at 6 m water depth) and a pelagic habitat (3 m water depth) to compare the impact of these different environments on fouling and degradation. The amount of biofilm increased on both plastic types and in both habitats. The diatom abundance and diversity differed significantly between the habitats and the plastic types. Diatoms were more abundant on samples from the pelagic zone. We anticipate that specific surface properties of the polymer types induced different biofilm communities on both plastic types. Additionally, different environmental conditions between the benthic and pelagic experimental site such as light intensity and shear forces may have influenced unequal colonisation between these habitats. The oxygen production rate was negative for all samples, indicating that the initial biofilm on marine plastic litter consumes oxygen, regardless of the plastic type or if exposed in the pelagic or the benthic zone. Mechanical tests did not reveal degradation within one month of exposure. However, scanning electron microscopy (SEM) analysis displayed potential signs of degradation on the plastic surface, which differed between both plastic types. This study indicates that the early biofilm formation and composition

  14. Material flow analysis of fossil fuels in China during 2000-2010.

    PubMed

    Wang, Sheng; Dai, Jing; Su, Meirong

    2012-01-01

    Since the relationship between the supply and demand of fossil fuels is on edge in the long run, the contradiction between the economic growth and limited resources will hinder the sustainable development of the Chinese society. This paper aims to analyze the input of fossil fuels in China during 2000-2010 via the material flow analysis (MFA) that takes hidden flows into account. With coal, oil, and natural gas quantified by MFA, three indexes, consumption and supply ratio (C/S ratio), resource consumption intensity (RCI), and fossil fuels productivity (FFP), are proposed to reflect the interactions between population, GDP, and fossil fuels. The results indicated that in the past 11 years, China's requirement for fossil fuels has been increasing continuously because of the growing mine productivity in domestic areas, which also leads to a single energy consumption structure as well as excessive dependence on the domestic exploitation. It is advisable to control the fossil fuels consumption by energy recycling and new energy facilities' popularization in order to lead a sustainable access to nonrenewable resources and decrease the soaring carbon emissions.

  15. Assessing global fossil fuel availability in a scenario framework

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Bauer, Nico; Hilaire, Jérôme; Brecha, Robert J.

    This study assesses global, long-term economic availability of coal, oil and gas within the Shared Socio-economic Pathway (SSP) scenario framework considering alternative assumptions as to highly uncertain future developments of technology, policy and the economy. Diverse sets of trajectories are formulated varying the challenges to mitigation and adaptation of climate change. The potential CO2 emissions from fossil fuels make it a crucial element subject to deep uncertainties. The analysis is based on a well-established data set of cost-quantity combinations that assumes favorable techno-economic developments, but ignores additional constraints on the extraction sector. This study significantly extends that analysis to includemore » alternative assumptions for the fossil fuel sector consistent with the SSP scenario families and applies these filters to the original data set, thus resulting in alternative cumulative fossil fuel availability curves. In a Middle-of-the-Road scenario, low cost fossil fuels embody carbon consistent with a RCP6.0 emission profile, if all the CO2 were emitted freely during the 21st century. In scenarios with high challenges to mitigation, the assumed embodied carbon in low-cost fossil fuels can trigger a RCP8.5 scenario; low mitigation challenges scenarios are still consistent with a RCP4.5 scenario.« less

  16. Measurements of turbulence and fossil turbulence near ampere seamount

    NASA Astrophysics Data System (ADS)

    Gibson, Carl H.; Nabatov, Valeriy; Ozmidov, Rostislav

    1993-10-01

    Measurements of temperature and velocity microstructure near and downstream of a shallow seamount are used to compare fossil turbulence versus non-fossil turbulence models for the evolution of turbulence microstructure patches in the stratified ocean. According to non-fossil oceanic turbulence models, all overturn length scales LT of the microstructure grow and collapse in constant proportion to each other and to the turbulence energy (Oboukov) scale LO and the inertial buoyancy (Ozmidov) scale L R≡(ɛ/N 3) 1/2 of the patches; that is, with LTrms ≈1.2 LR and viscous dissipation rate ɛ ≈ ɛ 0∗. According to the Gibson fossil turbulence model, all microstructure originates from completely active turbulence with ɛ ⩾ ɛ 0 ≈ 3L T2N 3(≈ 28ɛ 0∗) and L T/√6 ≈ L Trms, but this rapidly decays into a more persistent active-fossil state with ɛ0⩾ ɛ⩾ ɛF ≈ 30 vN2, where N is the buoyancy frequency and v is the kinematic viscosity and, without further energy supply, finally reaches a completely fossil turbulence hydrodynamic state of internal wave motions, with ɛ ⩽ ɛF. The last turbulence eddies, with ɛ ≈ ɛF, vanish at a buoyant-inertial-viscous (fossil Kolmogorov) scale LKF that is much smaller than the remnant overturn scales LT for large ɛ0/ ɛF ratios. These density, temperature, and salinity overturns with LT ≈ 0.6 LR0 ≫ 0.6 LR persist as turbulence fossils (by retaining the memory of ɛo) and collapse very slowly. In the near wake below the summit depth of Ampere seamount, a much larger proportion of completely active turbulence patches was found than is usually found in the ocean interior away from sources. Dissipation rates ɛ and turbulence activity coefficients A T ≡ (ɛ/ɛ 0) 1/2 of microstructure patches were found to decrease downstream, suggesting that the active turbulence indicated by the patches with AT ⩾ 1 was caused by the presence of the seamount as a turbulence source. Therefore, the turbulence and mixing

  17. Partially incorrect fossil data augment analyses of discrete trait evolution in living species.

    PubMed

    Puttick, Mark N

    2016-08-01

    Ancestral state reconstruction of discrete character traits is often vital when attempting to understand the origins and homology of traits in living species. The addition of fossils has been shown to alter our understanding of trait evolution in extant taxa, but researchers may avoid using fossils alongside extant species if only few are known, or if the designation of the trait of interest is uncertain. Here, I investigate the impacts of fossils and incorrectly coded fossils in the ancestral state reconstruction of discrete morphological characters under a likelihood model. Under simulated phylogenies and data, likelihood-based models are generally accurate when estimating ancestral node values. Analyses with combined fossil and extant data always outperform analyses with extant species alone, even when around one quarter of the fossil information is incorrect. These results are especially pronounced when model assumptions are violated, such as when there is a trend away from the root value. Fossil data are of particular importance when attempting to estimate the root node character state. Attempts should be made to include fossils in analysis of discrete traits under likelihood, even if there is uncertainty in the fossil trait data. © 2016 The Authors.

  18. Modelling the pelagic nitrogen cycle and vertical particle flux in the Norwegian sea

    NASA Astrophysics Data System (ADS)

    Haupt, Olaf J.; Wolf, Uli; v. Bodungen, Bodo

    1999-02-01

    A 1D Eulerian ecosystem model (BIological Ocean Model) for the Norwegian Sea was developed to investigate the dynamics of pelagic ecosystems. The BIOM combines six biochemical compartments and simulates the annual nitrogen cycle with specific focus on production, modification and sedimentation of particles in the water column. The external forcing and physical framework is based on a simulated annual cycle of global radiation and an annual mixed-layer cycle derived from field data. The vertical resolution of the model is given by an exponential grid with 200 depth layers, allowing specific parameterization of various sinking velocities, breakdown of particles and the remineralization processes. The aim of the numerical experiments is the simulation of ecosystem dynamics considering the specific biogeochemical properties of the Norwegian Sea, for example the life cycle of the dominant copepod Calanus finmarchicus. The results of the simulations were validated with field data. Model results are in good agreement with field data for the lower trophic levels of the food web. With increasing complexity of the organisms the differences increase between simulated processes and field data. Results of the numerical simulations suggest that BIOM is well adapted to investigate a physically controlled ecosystem. The simulation of grazing controlled pelagic ecosystems, like the Norwegian Sea, requires adaptations of parameterization to the specific ecosystem features. By using seasonally adaptation of the most sensible processes like utilization of light by phytoplankton and grazing by zooplankton results were greatly improved.

  19. Insect diversity in the fossil record

    NASA Technical Reports Server (NTRS)

    Labandeira, C. C.; Sepkoski, J. J. Jr; Sepkoski JJ, J. r. (Principal Investigator)

    1993-01-01

    Insects possess a surprisingly extensive fossil record. Compilation of the geochronologic ranges of insect families demonstrates that their diversity exceeds that of preserved vertebrate tetrapods through 91 percent of their evolutionary history. The great diversity of insects was achieved not by high origination rates but rather by low extinction rates comparable to the low rates of slowly evolving marine invertebrate groups. The great radiation of modern insects began 245 million years ago and was not accelerated by the expansion of angiosperms during the Cretaceous period. The basic trophic machinery of insects was in place nearly 100 million years before angiosperms appeared in the fossil record.

  20. A new compilation of stomach content data for commercially-important pelagic fish species in the Northeast Atlantic

    NASA Astrophysics Data System (ADS)

    Pinnegar, J. K.; Goñi, N.; Trenkel, V. M.; Arrizabalaga, H.; Melle, W.; Keating, J.; Óskarsson, G.

    2014-04-01

    There is increasing demand for information on predator-prey interactions in the ocean as a result of legislative commitments aimed at achieving sustainable exploitation. However, comprehensive datasets are lacking for many fish species and this has hampered development of multispecies fisheries models and the formulation of effective food-web indicators. This work describes a new compilation of stomach content data for five pelagic fish species (herring, blue whiting, mackerel, albacore and bluefin tuna) sampled across the northeast Atlantic and submitted to the PANGAEA open-access data portal (www.pangaea.de). We provide detailed descriptions of sample origin and of the corresponding database structures. We describe the main results in terms of diet composition and predator-prey relationships. The feeding preferences of small pelagic fish (herring, blue whiting, mackerel) were sampled over a very broad geographic area within the North Atlantic basin, from Greenland in the west, to the Lofoten Islands in the east and from the Bay of Biscay northwards to the Arctic. This analysis revealed significant differences in the prey items selected in different parts of the region at different times of year. Tunas (albacore and bluefin) were sampled in the Bay of Biscay and Celtic Sea. Dominant prey items for these species varied by location, year and season. This data compilation exercise represents one of the largest and most wide-ranging ever attempted for pelagic fish in the north Atlantic. The earliest data included in the database were collected in 1864, whereas the most recent were collected in 2012.Datasets are available at doi:10.1594/PANGAEA.820041 and doi:10.1594/PANGAEA.826992.

  1. A new compilation of stomach content data for commercially important pelagic fish species in the northeast Atlantic

    NASA Astrophysics Data System (ADS)

    Pinnegar, J. K.; Goñi, N.; Trenkel, V. M.; Arrizabalaga, H.; Melle, W.; Keating, J.; Óskarsson, G.

    2015-02-01

    There is increasing demand for information on predator-prey interactions in the ocean as a result of legislative commitments aimed at achieving sustainable exploitation. However, comprehensive data sets are lacking for many fish species and this has hampered development of multispecies fisheries models and the formulation of effective food-web indicators. This work describes a new compilation of stomach content data for five pelagic fish species (herring, blue whiting, mackerel, albacore and bluefin tuna) sampled across the northeast Atlantic and submitted to the PANGAEA open-access data portal (www.pangaea.de). We provide detailed descriptions of sample origin and of the corresponding database structures. We describe the main results in terms of diet composition and predator-prey relationships. The feeding preferences of small pelagic fish (herring, blue whiting, mackerel) were sampled over a very broad geographic area within the North Atlantic basin, from Greenland in the west, to the Lofoten Islands in the east and from the Bay of Biscay northwards to the Arctic. This analysis revealed significant differences in the prey items selected in different parts of the region at different times of year. Tunas (albacore and bluefin) were sampled in the Bay of Biscay and Celtic Sea. Dominant prey items for these species varied by location, year and season. This data compilation exercise represents one of the largest and most wide-ranging ever attempted for pelagic fish in the North Atlantic. The earliest data included in the database were collected in 1864, whereas the most recent were collected in 2012. Data sets are available at doi:10.1594/PANGAEA.820041 and doi:10.1594/PANGAEA.826992.

  2. Time Trends of Persistent Organic Pollutants in Benthic and Pelagic Indicator Fishes from Puget Sound, Washington, USA.

    PubMed

    West, James E; O'Neill, Sandra M; Ylitalo, Gina M

    2017-08-01

    We modeled temporal trends in polychlorinated biphenyls (PCBs), polybrominated diphenyl ethers (PBDEs), and dichlorodiphenyltrichloroethane and its metabolites (DDTs) in two indicator fish species representing benthic and pelagic habitats in Puget Sound, Washington, USA. English sole (Parophrys vetulus, benthic) index sites and larger-scale Pacific herring (Clupea pallasii, pelagic) foraging areas represented a wide range of possible contamination conditions, with sampling locations situated adjacent to watersheds exhibiting high, medium and low development. Consistency in analytical data throughout the study was maintained by either calculating method-bias-correction factors on paired samples as methods evolved or by analyzing older archived samples by current methods. PCBs declined moderately in two herring stocks from a low-development basin (2.3 and 4.0% annual rate of decline) and showed no change in the highly developed and moderately developed basins during a 16- to 21-year period. PCBs increased in English sole from four of ten sites (2.9-7.1%), and the remaining six exhibited no significant change. PBDEs and DDTs declined significantly in all herring stocks (4.2-8.1%), although analytical challenges warrant caution in interpreting DDT results. PBDEs declined in English sole from two high-development and one low-development site (3.7-7.2%) and remained unchanged in the remaining seven. DDTs increased in English sole from one high-development site (Tacoma City Waterway) and declined in two high-development and one low development site. As with herring, analytical challenges warrant caution in interpreting the English sole DDT results. It is likely that source controls and mitigation efforts have contributed to the declines in PBDEs and DDTs overall, whereas PCBs appear to have persisted, especially in the pelagic food web, despite bans in PCB production and use.

  3. Education Program on Fossil Resources Including Coal

    NASA Astrophysics Data System (ADS)

    Usami, Masahiro

    Fossil fuels including coal play a key role as crucial energies in contributing to economic development in Asia. On the other hand, its limited quantity and the environmental problems causing from its usage have become a serious global issue and a countermeasure to solve such problems is very much demanded. Along with the pursuit of sustainable development, environmentally-friendly use of highly efficient fossil resources should be therefore, accompanied. Kyushu-university‧s sophisticated research through long years of accumulated experience on the fossil resources and environmental sectors together with the advanced large-scale commercial and empirical equipments will enable us to foster cooperative research and provide internship program for the future researchers. Then, this program is executed as a consignment business from the Ministry of Economy, Trade and Industry from 2007 fiscal year to 2009 fiscal year. The lecture that uses the textbooks developed by this program is scheduled to be started a course in fiscal year 2010.

  4. Do fossil vertebrate biominerals hold the key to the Palaeozoic climate?

    NASA Astrophysics Data System (ADS)

    Žigaitė, Ž.

    2012-04-01

    Fossil vertebrate hard tissues - teeth and dermoskeleton - are considered among the most geochemically stable biominerals, and therefore are widely used for palaeoenvironmental and palaeoclimatic reconstructions. Elemental and isotopic compositions of fossil dental tissues may provide unique palaeoenvironmental information, ranging from the diet and trophic positions on a food chain, to the palaeosalinity and water temperatures of ancient seas. However, the post-mortem alteration and re-crystallisation of fossil hard tissues may hamper these interpretations. Chemical composition and isotopic equilibrium of the biomineral change readily at any time from the earliest diagenesis to the final laboratory acid treatment during the fossil preparation. This is why particular attention shall be given to the preservation of fossil tissues, evaluating carefully the level of possible alteration in the primary geochemical composition. Pre-evaluation of fossil preservation can be made by semi-quantitative spot geochemistry analyses on fine polished teeth and scale thin sections using Energy Dispersive X-ray Spectroscopy (EDS), and help to preview the chemical composition of biomineral. The Electron Backscatter Diffractometry (EBSD) is useful to examine the cristallinity and possible structural alterations. In addition, rare earth element (REE) abundances can be measured in situ within the fine fossil tissues (such as enamel) using Laser Ablation Inductively Coupled Plasma Mass-spectrometry (LA-ICP-MS), giving evidence on the selective geochemical resilience between separate vertebrate hard tissues. Therefore, in order to decipher the geochemical signal correctly, the evaluation of preservation is a necessary starting point to any further studies of fossil biomineral geochemistry.

  5. Estimating past precipitation and temperature from fossil ostracodes

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Smith, A.J.; Forester, R.M.

    1994-12-31

    The fossil records of certain aquatic organisms provide a way of obtaining meaningful estimates of past temperature and precipitation. These estimates of past environmental conditions are derived from multivariate statistical methods that are in turn based on the modern biogeographic distributions and environmental tolerances of the biota of interest. These estimates are helpful in conducting slimate studies as part of the Yucca Mountain site characterization. Ostracodes are microscopic crustaceans that produce bivalved calcite shells which are easily fossilized in the sediments of the lakes and wetlands in which the animals lived. The modern biogeographic distribution and environmental conditions of livingmore » ostracodes are the basis for the interpretation of the past environmental conditions of the fossil ostracodes. The major assumption in this method of interpretation is that the environmental tolerances of ostracodes have not changed substantially over thousands of years. Two methods using these modern analogs to determine past environmental conditions are the modern analog method and the range method. The range method also considers the information provided by fossil ostracode assemblages that have no modern analog in today`s world.« less

  6. Dataset for analysing the relationships among economic growth, fossil fuel and non-fossil fuel consumption.

    PubMed

    Asafu-Adjaye, John; Byrne, Dominic; Alvarez, Maximiliano

    2017-02-01

    The data presented in this article are related to the research article entitled 'Economic Growth, Fossil Fuel and Non-Fossil Consumption: A Pooled Mean Group Analysis using Proxies for Capital' (J. Asafu-Adjaye, D. Byrne, M. Alvarez, 2016) [1]. This article describes data modified from three publicly available data sources: the World Bank׳s World Development Indicators (http://databank.worldbank.org/data/reports.aspx?source=world-development-indicators), the U.S. Energy Information Administration׳s International Energy Statistics (http://www.eia.gov/cfapps/ipdbproject/IEDIndex3.cfm?tid=44&pid=44&aid=2) and the Barro-Lee Educational Attainment Dataset (http://www.barrolee.com). These data can be used to examine the relationships between economic growth and different forms of energy consumption. The dataset is made publicly available to promote further analyses.

  7. Pelagic seabird surveys in the Tuamotu and Gambier Archipelagos, French Polynesia

    USGS Publications Warehouse

    Vanderwerf, Eric A.; Pierce, Ray J.; Gill, Verena A.; Wragg, Graham; Raust, Philippe; Tibbitts, T. Lee

    2006-01-01

    We conducted pelagic seabird surveys in the Gambier and Tuamotu Archipelagos in the southeastern Pacific Ocean totaling 40 hours during 7-27 March 2003 and 22.5 hours during 22-27 July 2001. We used a 300-m-wide strip transect to estimate seabird density, and we estimated relative abundance of birds at all distances. In 2001, we observed a total of 326 birds of 18 species. The mean relative abundance of all birds was 14.3 ?? 3.1/h. Red-footed Booby Sula sula was the most abundant species (5.6/h), followed by White Tern Gygis alba (3.4/h), and Great Crested or Swift Tern Sterna bergii (1.2/h). In 2003, we observed a total of 1463 birds of 25 species. The mean relative abundance of all birds was 36.6 ?? 11.4/h and the mean density of all birds was 4.14 ?? 0.72/km2. Brown Noddy Anous stolidus was the most abundant species (17.6/h, 1.5/km2), followed by White Tern (8.4/h, 1.3/km2), and Red-footed Booby (4.6/h, 0.8/km2). Several globally or locally rare species were observed infrequently, including Phoenix Petrel Pterodroma alba (0.1/h, 0.03/km2). Distribution of birds was uneven, with long periods of no birds punctuated by occasional feeding flocks. In 2003, species diversity was related to length of observation period, with more species observed on longer segments (r2 = 0.58, F1.5 = 6.03, P = 0.05). Although the duration and extent of our surveys were limited, these data are valuable because little published information is available on pelagic seabirds in southeastern Polynesia.

  8. Pelagic origin and fate of sedimenting particles in the Norwegian Sea

    NASA Astrophysics Data System (ADS)

    Bathmann, Ulrich V.; Peinert, Rolf; Noji, Thomas T.; Bodungen, Bodo V.

    A 17 month record of vertical particle flux of dry weight, carbonate and organic carbon were 25.8, 9.4 and 2.4g.m -2y -1, respectively. Parallel to trap deployments, pelagic system structure was recorded with high vertical and temporal resolution. Within a distinct seasonal cycle of vertical particle flux, zooplankton faecal pellets of various sizes, shapes and contents were collected by the traps in different proportions and quantities throughout the year (range: 0-4,500 10 3m -2d -1). The remains of different groups of organisms showed distinct seasonal variations in abundance. In early summer there was a small maximum in the diatom flux and this was followed by pulses of tinntinids, radiolarians, foraminiferans and pteropods between July and November. Food web interactions in the water column were important in controlling the quality and quantity of sinking materials. For example, changes in the population structure of dominant herbivores, the break-down of regenerating summer populations of microflagellates and protozooplankton and the collapse of a pteropod dominated community, each resulted in marked sedimentation pulses. These data from the Norwegian Sea indicate those mechanisms which either accelerate or counteract loss of material via sedimentation. These involve variations in the structure of the pelagic system and they operatè on long (e.g. annual plankton succession) and short (e.g. the end of new production, sporadic grazing of swarm feeders) time scales. Connecting investigation of the water column with a high resolution in time in parallel with drifting sediment trap deployments and shipboard experiments with the dominant zooplankters is a promising approach for giving a better understanding of both the origin and the fate of material sinking to the sea floor.

  9. Testing New Proxies for Photosymbiosis in the Fossil Record

    NASA Astrophysics Data System (ADS)

    Tornabene, C.; Martindale, R. C.; Schaller, M. F.

    2015-12-01

    Photosymbiosis is a mutualistic relationship that many corals have developed with dinoflagellates called zooxanthellae. The dinoflagellates, of the genus Symbiodinium, photosynthesize and provide corals with most of their energy, while in turn coral hosts live in waters where zooxanthellae have optimal exposure to sunlight. Thanks to this relationship, symbiotic corals calcify faster than non-symbiotic corals. Photosymbiosis is therefore considered the evolutionary innovation that allowed corals to become major reef-builders through geological time.This relationship is extremely difficult to study. Zooxanthellae, which are housed in the coral tissue, are not preserved in fossil coral skeletons, thus determining whether corals had symbionts requires a robust proxy. In order to address this critical question, the goal of this research is to test new proxies for ancient photosymbiosis. Currently the project is focused on assessing the nitrogen (δ15N) isotopes of corals' organic matrices, sensu Muscatine et al. (2005), as well as carbon and oxygen (δ13C, δ18O) isotopes of fossil coral skeletons. Samples from Modern, Pleistocene, Oligocene and Triassic coral skeletons were analyzed to test the validity of these proxies. Coral samples comprise both (interpreted) symbiotic and non-symbiotic fossil corals from the Oligocene and Triassic as well as symbiotic fossil corals from the Modern and Pleistocene to corroborate our findings with the results of Muscatine et al. (2005). Samples were tested for diagenesis through petrographic and scanning electron microscope (SEM) analyses to avoid contamination. Additionally, a novel technique that has not yet been applied to the fossil record was tested. The technique aims to recognize dinosterol, a dinoflagellate biomarker, in both modern and fossil coral samples. The premise of this proxy is that symbiotic corals should contain the dinoflagellate biomarker, whereas those lacking symbionts should lack dinosterol. Results from this

  10. Modeled distribution and abundance of a pelagic seabird reveal trends in relation to fisheries

    USGS Publications Warehouse

    Renner, Martin; Parrish, Julia K.; Piatt, John F.; Kuletz, Kathy J.; Edwards, Ann E.; Hunt, George L.

    2013-01-01

    The northern fulmar Fulmarus glacialis is one of the most visible and widespread seabirds in the eastern Bering Sea and Aleutian Islands. However, relatively little is known about its abundance, trends, or the factors that shape its distribution. We used a long-term pelagic dataset to model changes in fulmar at-sea distribution and abundance since the mid-1970s. We used an ensemble model, based on a weighted average of generalized additive model (GAM), multivariate adaptive regression splines (MARS), and random forest models to estimate the pelagic distribution and density of fulmars in the waters of the Aleutian Archipelago and Bering Sea. The most important predictor variables were colony effect, sea surface temperature, distribution of fisheries, location, and primary productivity. We calculated a time series from the ratio of observed to predicted values and found that fulmar at-sea abundance declined from the 1970s to the 2000s at a rate of 0.83% (± 0.39% SE) per annum. Interpolating fulmar densities on a spatial grid through time, we found that the center of fulmar distribution in the Bering Sea has shifted north, coinciding with a northward shift in fish catches and a warming ocean. Our study shows that fisheries are an important, but not the only factor, shaping fulmar distribution and abundance trends in the eastern Bering Sea and Aleutian Islands.

  11. Eocene Patagonia fossils of the daisy family.

    PubMed

    Barreda, V D; Palazzesi, L; Tellería, M C; Katinas, L; Crisci, J V; Bremer, K; Passalia, M G; Corsolini, R; Rodríguez Brizuela, R; Bechis, F

    2010-09-24

    Fossil capitula and pollen grains of Asteraceae from the Eocene of Patagonia, southern Argentina, exhibit morphological features recognized today in taxa, such as Mutisioideae and Carduoideae, that are phylogenetically close to the root of the asteracean tree. This fossil supports the hypothesis of a South American origin of Asteraceae and an Eocene age of divergence and suggests that an ancestral stock of Asteraceae may have formed part of a geoflora developed in southern Gondwana before the establishment of effective dispersal barriers within this landmass.

  12. A synthesis of carbon dioxide emissions from fossil-fuel combustion

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Andres, Robert Joseph; Boden, Thomas A; Breon, F.-M.

    2012-01-01

    This synthesis discusses the emissions of carbon dioxide from fossil-fuel combustion and cement production. While much is known about these emissions, there is still much that is unknown about the details surrounding these emissions. This synthesis explores 5 our knowledge of these emissions in terms of why there is concern about them; how they are calculated; the major global efforts on inventorying them; their global, regional, and national totals at different spatial and temporal scales; how they are distributed on global grids (i.e. maps); how they are transported in models; and the uncertainties associated with these different aspects of themore » emissions. The magnitude of emissions 10 from the combustion of fossil fuels has been almost continuously increasing with time since fossil fuels were first used by humans. Despite events in some nations specifically designed to reduce emissions, or which have had emissions reduction as a byproduct of other events, global total emissions continue their general increase with time. Global total fossil-fuel carbon dioxide emissions are known to within 10% uncertainty (95% 15 confidence interval). Uncertainty on individual national total fossil-fuel carbon dioxide emissions range from a few percent to more than 50 %. The information discussed in this manuscript synthesizes global, regional and national fossil-fuel carbon dioxide emissions, their distributions, their transport, and the associated uncertainties.« less

  13. Divesting from Fossil Fuels Makes Sense Morally… and Financially

    ERIC Educational Resources Information Center

    Cleveland, Cutler J.; Reibstein, Richard

    2015-01-01

    Should university endowments divest from fossil fuels? A public discussion of this question has seen some university presidents issuing statements that they would not divest--that investments should not be used for "political action." Many universities hold large endowments that have significant positions in fossil fuel companies or…

  14. Evidence for foraging -site fidelity and individual foraging behavior of pelagic cormorants rearing chicks in the gulf of Alaska

    USGS Publications Warehouse

    Kotzerka, J.; Hatch, Shyla A.; Garthe, S.

    2011-01-01

    The Pelagic Cormorant (Phalacrocorax pelagicus) is the most widespread cormorant in the North Pacific, but little is known about its foraging and diving behavior. However, knowledge of seabirds' foraging behavior is important to understanding their function in the marine environment. In 2006, using GPS dataloggers, we studied the foraging behavior of 14 male Pelagic Cormorants rearing chicks on Middleton Island, Alaska. For foraging, the birds had high fidelity to a small area 8 km north of the colony. Within that area, the cormorants' diving activity was of two distinct kinds-near-surface dives (1-6 m) and benthic dives (28-33 m). Individuals were consistent in the depths of their dives, either mostly shallow or mostly deep. Few showed no depth preference. Dive duration, time at maximum depth, and pauses at the water surface between consecutive dives were shorter for shallow dives than for deep dives. The cormorants made dives of both types throughout the day, but the frequency of deep dives increased toward evening. Maximum foraging range was 9 km; maximum total distance traveled per trip was 43.4 km. Trip durations ranged from 0.3 to 7.7 hr. Maximum depth of a dive was 42.2 m, and duration of dives ranged from 4 to 120 sec. We found that Pelagic Cormorants at Middleton Island were faithful to one particular foraging area and individuals dived in distinct patterns. Distinct, specialized foraging behavior may be advantageous in reducing intra- and interspecific competition but may also render the species vulnerable to changing environmental conditions. Copyright ?? The Cooper Ornithological Society 2011.

  15. Fossil butterflies, calibration points and the molecular clock (Lepidoptera: Papilionoidea).

    PubMed

    Jong, Rienk DE

    2017-05-25

    Fossil butterflies are extremely rare. Yet, they are the only direct evidence of the first appearance of particular characters and as such, they are crucial for calibrating a molecular clock, from which divergence ages are estimated. In turn, these estimates, in combination with paleogeographic information, are most important in paleobiogeographic considerations. The key issue here is the correct allocation of fossils on the phylogenetic tree from which the molecular clock is calibrated.The allocation of a fossil on a tree should be based on an apomorphic character found in a tree based on extant species, similar to the allocation of a new extant species. In practice, the latter is not done, at least not explicitly, on the basis of apomorphy, but rather on overall similarity or on a phylogenetic analysis, which is not possible for most butterfly fossils since they usually are very fragmentary. Characters most often preserved are in the venation of the wings. Therefore, special attention is given to possible apomorphies in venational characters in extant butterflies. For estimation of divergence times, not only the correct allocation of the fossil on the tree is important, but also the tree itself influences the outcome as well as the correct determination of the age of the fossil. These three aspects are discussed.        All known butterfly fossils, consisting of 49 taxa, are critically reviewed and their relationship to extant taxa is discussed as an aid for correctly calibrating a molecular clock for papilionoid Lepidoptera. In this context some aspects of age estimation and biogeographic conclusions are briefly mentioned in review. Specific information has been summarized in four appendices.

  16. Stable isotope analysis of vertebrae reveals ontogenetic changes in habitat in an endothermic pelagic shark

    PubMed Central

    Carlisle, Aaron B.; Goldman, Kenneth J.; Litvin, Steven Y.; Madigan, Daniel J.; Bigman, Jennifer S.; Swithenbank, Alan M.; Kline, Thomas C.; Block, Barbara A.

    2015-01-01

    Ontogenetic changes in habitat are driven by shifting life-history requirements and play an important role in population dynamics. However, large portions of the life history of many pelagic species are still poorly understood or unknown. We used a novel combination of stable isotope analysis of vertebral annuli, Bayesian mixing models, isoscapes and electronic tag data to reconstruct ontogenetic patterns of habitat and resource use in a pelagic apex predator, the salmon shark (Lamna ditropis). Results identified the North Pacific Transition Zone as the major nursery area for salmon sharks and revealed an ontogenetic shift around the age of maturity from oceanic to increased use of neritic habitats. The nursery habitat may reflect trade-offs between prey availability, predation pressure and thermal constraints on juvenile endothermic sharks. The ontogenetic shift in habitat coincided with a reduction of isotopic niche, possibly reflecting specialization upon particular prey or habitats. Using tagging data to inform Bayesian isotopic mixing models revealed that adult sharks primarily use neritic habitats of Alaska yet receive a trophic subsidy from oceanic habitats. Integrating the multiple methods used here provides a powerful approach to retrospectively study the ecology and life history of migratory species throughout their ontogeny. PMID:25621332

  17. Hydroacoustic estimates of abundance and spatial distribution of pelagic prey fishes in western Lake Superior

    USGS Publications Warehouse

    Mason, Doran M.; Johnson, Timothy B.; Harvey, Chris J.; Kitchell, James F.; Schram, Stephen T.; Bronte, Charles R.; Hoff, Michael H.; Lozano, Stephen J.; Trebitz, Anett S.; Schreiner, Donald R.; Lamon, E. Conrad; Hrabik, Thomas R.

    2005-01-01

    Lake herring (Coregonus artedi) and rainbow smelt (Osmerus mordax) are a valuable prey resource for the recovering lake trout (Salvelinus namaycush) in Lake Superior. However, prey biomass may be insufficient to support the current predator demand. In August 1997, we assessed the abundance and spatial distribution of pelagic coregonines and rainbow smelt in western Lake Superior by combining a 120 kHz split beam acoustics system with midwater trawls. Coregonines comprised the majority of the midwater trawl catches and the length distributions for trawl caught fish coincided with estimated sizes of acoustic targets. Overall mean pelagic prey fish biomass was 15.56 kg ha−1 with the greatest fish biomass occurring in the Apostle Islands region (27.98 kg ha−1), followed by the Duluth Minnesota region (20.22 kg ha−1), and with the lowest biomass occurring in the open waters of western Lake Superior (9.46 kg ha−1). Biomass estimates from hydroacoustics were typically 2–134 times greater than estimates derived from spring bottom trawl surveys. Prey fish biomass for Lake Superior is about order of magnitude less than acoustic estimates for Lakes Michigan and Ontario. Discrepancies observed between bioenergetics-based estimates of predator consumption of coregonines and earlier coregonine biomass estimates may be accounted for by our hydroacoustic estimates.

  18. A revised checklist of Nepticulidae fossils (Lepidoptera) indicates an Early Cretaceous origin.

    PubMed

    Doorenweerd, Camiel; Nieukerken, Erik J Van; Sohn, Jae-Cheon; Labandeira, Conrad C

    2015-05-27

    With phylogenetic knowledge of Lepidoptera rapidly increasing, catalysed by increasingly powerful molecular techniques, the demand for fossil calibration points to estimate an evolutionary timeframe for the order is becoming an increasingly pressing issue. The family Nepticulidae is a species rich, basal branch within the phylogeny of the Lepidoptera, characterized by larval leaf-mining habits, and thereby represents a potentially important lineage whose evolutionary history can be established more thoroughly with the potential use of fossil calibration points. Using our experience with extant global Nepticulidae, we discuss a list of characters that may be used to assign fossil leaf mines to Nepticulidae, and suggest useful methods for classifying relevant fossil material. We present a checklist of 79 records of Nepticulidae representing adult and leaf-mine fossils mentioned in literature, often with multiple exemplars constituting a single record. We provide our interpretation of these fossils. Two species now are included in the collective generic name Stigmellites: Stigmellites resupinata (Krassilov, 2008) comb. nov. (from Ophiheliconoma) and Stigmellites almeidae (Martins-Neto, 1989) comb. nov. (from Nepticula). Eleven records are for the first time attributed to Nepticulidae. After discarding several dubious records, including one possibly placing the family at a latest Jurassic position, we conclude that the oldest fossils likely attributable to Nepticulidae are several exemplars representing a variety of species from the Dakota Formation (USA). The relevant strata containing these earliest fossils are now dated at 102 Ma (million years ago) in age, corresponding to the latest Albian Stage of the Early Cretaceous. Integration of all records in the checklist shows that a continuous presence of nepticulid-like leaf mines preserved as compression-impression fossils and by amber entombment of adults have a fossil record extending to the latest Early Cretaceous.

  19. Testing the molecular clock using mechanistic models of fossil preservation and molecular evolution

    PubMed Central

    2017-01-01

    Molecular sequence data provide information about relative times only, and fossil-based age constraints are the ultimate source of information about absolute times in molecular clock dating analyses. Thus, fossil calibrations are critical to molecular clock dating, but competing methods are difficult to evaluate empirically because the true evolutionary time scale is never known. Here, we combine mechanistic models of fossil preservation and sequence evolution in simulations to evaluate different approaches to constructing fossil calibrations and their impact on Bayesian molecular clock dating, and the relative impact of fossil versus molecular sampling. We show that divergence time estimation is impacted by the model of fossil preservation, sampling intensity and tree shape. The addition of sequence data may improve molecular clock estimates, but accuracy and precision is dominated by the quality of the fossil calibrations. Posterior means and medians are poor representatives of true divergence times; posterior intervals provide a much more accurate estimate of divergence times, though they may be wide and often do not have high coverage probability. Our results highlight the importance of increased fossil sampling and improved statistical approaches to generating calibrations, which should incorporate the non-uniform nature of ecological and temporal fossil species distributions. PMID:28637852

  20. Testing the molecular clock using mechanistic models of fossil preservation and molecular evolution.

    PubMed

    Warnock, Rachel C M; Yang, Ziheng; Donoghue, Philip C J

    2017-06-28

    Molecular sequence data provide information about relative times only, and fossil-based age constraints are the ultimate source of information about absolute times in molecular clock dating analyses. Thus, fossil calibrations are critical to molecular clock dating, but competing methods are difficult to evaluate empirically because the true evolutionary time scale is never known. Here, we combine mechanistic models of fossil preservation and sequence evolution in simulations to evaluate different approaches to constructing fossil calibrations and their impact on Bayesian molecular clock dating, and the relative impact of fossil versus molecular sampling. We show that divergence time estimation is impacted by the model of fossil preservation, sampling intensity and tree shape. The addition of sequence data may improve molecular clock estimates, but accuracy and precision is dominated by the quality of the fossil calibrations. Posterior means and medians are poor representatives of true divergence times; posterior intervals provide a much more accurate estimate of divergence times, though they may be wide and often do not have high coverage probability. Our results highlight the importance of increased fossil sampling and improved statistical approaches to generating calibrations, which should incorporate the non-uniform nature of ecological and temporal fossil species distributions. © 2017 The Authors.

  1. Clustering fossils in solid inflation

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Akhshik, Mohammad, E-mail: m.akhshik@ipm.ir

    In solid inflation the single field non-Gaussianity consistency condition is violated. As a result, the long tenor perturbation induces observable clustering fossils in the form of quadrupole anisotropy in large scale structure power spectrum. In this work we revisit the bispectrum analysis for the scalar-scalar-scalar and tensor-scalar-scalar bispectrum for the general parameter space of solid. We consider the parameter space of the model in which the level of non-Gaussianity generated is consistent with the Planck constraints. Specializing to this allowed range of model parameter we calculate the quadrupole anisotropy induced from the long tensor perturbations on the power spectrum ofmore » the scalar perturbations. We argue that the imprints of clustering fossil from primordial gravitational waves on large scale structures can be detected from the future galaxy surveys.« less

  2. The phylogeny of fossil whip spiders.

    PubMed

    Garwood, Russell J; Dunlop, Jason A; Knecht, Brian J; Hegna, Thomas A

    2017-04-21

    Arachnids are a highly successful group of land-dwelling arthropods. They are major contributors to modern terrestrial ecosystems, and have a deep evolutionary history. Whip spiders (Arachnida, Amblypygi), are one of the smaller arachnid orders with ca. 190 living species. Here we restudy one of the oldest fossil representatives of the group, Graeophonus anglicus Pocock, 1911 from the Late Carboniferous (Duckmantian, ca. 315 Ma) British Middle Coal Measures of the West Midlands, UK. Using X-ray microtomography, our principal aim was to resolve details of the limbs and mouthparts which would allow us to test whether this fossil belongs in the extant, relict family Paracharontidae; represented today by a single, blind species Paracharon caecus Hansen, 1921. Tomography reveals several novel and significant character states for G. anglicus; most notably in the chelicerae, pedipalps and walking legs. These allowed it to be scored into a phylogenetic analysis together with the recently described Paracharonopsis cambayensis Engel & Grimaldi, 2014 from the Eocene (ca. 52 Ma) Cambay amber, and Kronocharon prendinii Engel & Grimaldi, 2014 from Cretaceous (ca. 99 Ma) Burmese amber. We recovered relationships of the form ((Graeophonus (Paracharonopsis + Paracharon)) + (Charinus (Stygophrynus (Kronocharon (Charon (Musicodamon + Paraphrynus)))))). This tree largely reflects Peter Weygoldt's 1996 classification with its basic split into Paleoamblypygi and Euamblypygi lineages; we were able to score several of his characters for the first time in fossils. Our analysis draws into question the monophyly of the family Charontidae. Our data suggest that Graeophonus is a crown group amblypygid, and falls within a monophyletic Paleoamblypgi clade, but outside the family Paracharontidae (= Paracharonopsis + Paracharon). Our results also suggest a new placement for the Burmese amber genus Kronocharon, a node further down from its original position. Overall, we offer a

  3. Spectroscopic studies of wood fossils from the Crato Formation, Cretaceous Period.

    PubMed

    da Silva, J H; Freire, P T C; Abagaro, B T O; Silva, J A F; Saraiva, G D; de Lima, F J; Barros, O A; Bantim, R A; Saraiva, A A F; Viana, B C

    2013-11-01

    In this work we study two types of wood fossils (Gymnosperms, Araucariaceae) from the Crato Formation of Araripe Basin in Brazil, from the Cretaceous Period. The samples were characterized by Raman and infrared spectroscopies, X-ray diffraction and scanning electron microscopy. The results obtained by different techniques showed that although the rocks surrounding the fossils have predominantly the same constitution - calcite - however, the formation processes of these types of wood fossils are quite different. One of the fossils, denominated as light wood, is predominantly composed of gypsum, while the other fossil, the dark wood, is rich in amorphous carbon, possibly the kerogen type. Implications relative to the environment where the plants lived millions years ago are also given. Finally, the results highlight the constitution of one of the most important paleontological sites of the Cretaceous Period in the South America. Copyright © 2013 Elsevier B.V. All rights reserved.

  4. The late Middle Pleistocene hominin fossil record of eastern Asia: synthesis and review.

    PubMed

    Bae, Christopher J

    2010-01-01

    Traditionally, Middle Pleistocene hominin fossils that cannot be allocated to Homo erectus sensu lato or modern H. sapiens have been assigned to different specific taxa. For example, in eastern Asia, these hominin fossils have been classified as archaic, early, or premodern H. sapiens. An increasing number of Middle Pleistocene hominin fossils are currently being assigned to H. heidelbergensis. This is particularly the case for the African and European Middle Pleistocene hominin fossil record. There have been suggestions that perhaps the eastern Asian late Middle Pleistocene hominins can also be allocated to the H. heidelbergensis hypodigm. In this article, I review the current state of the late Middle Pleistocene hominin fossil record from eastern Asia and examine the various arguments for assigning these hominins to the different specific taxa. The two primary conclusions drawn from this review are as follows: 1) little evidence currently exists in the eastern Asian Middle Pleistocene hominin fossil record to support their assignment to H. heidelbergensis; and 2) rather than add to the growing list of hominin fossil taxa by using taxonomic names like H. daliensis for northeast Asian fossils and H. mabaensis for Southeast Asian fossils, it is better to err on the side of caution and continue to use the term archaic H. sapiens to represent all of these hominin fossils. What should be evident from this review is the need for an increase in the quality and quantity of the eastern Asian hominin fossil data set. Fortunately, with the increasing number of large-scale multidisciplinary paleoanthropological field and laboratory research projects in eastern Asia, the record is quickly becoming better understood. Copyright © 2010 Wiley-Liss, Inc.

  5. Reconstructing the past: methods and techniques for the digital restoration of fossils

    PubMed Central

    2016-01-01

    During fossilization, the remains of extinct organisms are subjected to taphonomic and diagenetic processes. As a result, fossils show a variety of preservational artefacts, which can range from small breaks and cracks, disarticulation and fragmentation, to the loss and deformation of skeletal structures and other hard parts. Such artefacts can present a considerable problem, as the preserved morphology of fossils often forms the basis for palaeontological research. Phylogenetic and taxonomic studies, inferences on appearance, ecology and behaviour and functional analyses of fossil organisms strongly rely on morphological information. As a consequence, the restoration of fossil morphology is often a necessary prerequisite for further analyses. Facilitated by recent computational advances, virtual reconstruction and restoration techniques offer versatile tools to restore the original morphology of fossils. Different methodological steps and approaches, as well as software are outlined and reviewed here, and advantages and disadvantages are discussed. Although the complexity of the restorative processes can introduce a degree of interpretation, digitally restored fossils can provide useful morphological information and can be used to obtain functional estimates. Additionally, the digital nature of the restored models can open up possibilities for education and outreach and further research. PMID:27853548

  6. Status of pelagic prey fishes and pelagic macroinvertebrates in Lake Michigan, 2008

    USGS Publications Warehouse

    Warner, David M.; Claramunt, Randall M.; Holuszko, Jeffrey D.; Desorcie, Timothy J.

    2009-01-01

    Acoustic surveys were conducted in late summer/early fall during the years 1992-1996 and 2001-2008 to estimate pelagic prey fish biomass in Lake Michigan. Midwater trawling during the surveys provided a measure of species and size composition of the fish community for use in scaling acoustic data and providing species-specific abundance estimates. In 2005, we began sampling Mysis diluviana during the survey. The 2008 survey provided data from 24 acoustic transects (734 km), 33 midwater tows, and 39 mysid tows. Mean total prey fish biomass was 15.3 kg/ha (relative standard error, RSE = 7.6%) or ~82 kilotonnes (kt, 1,000 metric tons), which was 1.9 times higher than the estimate for 2007 but 78% lower than the long-term mean. The increase from 2007 was because of increased biomass of age-1 and age-3 alewife. The 2008 alewife year-class contributed ~12% of total alewife biomass (11.0 kg/ha, RSE = 9.0%), while the 2007 and 2005 alewife year-classes contributed ~33% and 35%, respectively. In 2008, alewife comprised 72% of total biomass, while rainbow smelt and bloater were 11 and 17% of total biomass, respectively. Rainbow smelt biomass in 2008 (1.6 kg/ha, RSE = 10.6%) was identical to the biomass in 2007 (1.6 kg/ha). Bloater biomass was again much lower (2.6 kg/ha, RSE = 15.2%) than in the 1990s, but mean density of small bloater in 2008 (534 fish/ha, RSE = 10.9) was the highest observed in any acoustic survey on record. Prey fish biomass remained well below the Fish Community Objectives target of 500-800 kt and only alewife and small bloater are above or near long-term mean biomass levels. Mysis diluviana remains relatively abundant. Mean density ranged from 185 ind./m2 (RSE = 6.8%) in 2005 to 112 ind./m2 (RSE = 5.1%) in 2007, but there was not a statistically significant difference among years.

  7. Improving large-scale groundwater models by considering fossil gradients

    NASA Astrophysics Data System (ADS)

    Schulz, Stephan; Walther, Marc; Michelsen, Nils; Rausch, Randolf; Dirks, Heiko; Al-Saud, Mohammed; Merz, Ralf; Kolditz, Olaf; Schüth, Christoph

    2017-05-01

    Due to limited availability of surface water, many arid to semi-arid countries rely on their groundwater resources. Despite the quasi-absence of present day replenishment, some of these groundwater bodies contain large amounts of water, which was recharged during pluvial periods of the Late Pleistocene to Early Holocene. These mostly fossil, non-renewable resources require different management schemes compared to those which are usually applied in renewable systems. Fossil groundwater is a finite resource and its withdrawal implies mining of aquifer storage reserves. Although they receive almost no recharge, some of them show notable hydraulic gradients and a flow towards their discharge areas, even without pumping. As a result, these systems have more discharge than recharge and hence are not in steady state, which makes their modelling, in particular the calibration, very challenging. In this study, we introduce a new calibration approach, composed of four steps: (i) estimating the fossil discharge component, (ii) determining the origin of fossil discharge, (iii) fitting the hydraulic conductivity with a pseudo steady-state model, and (iv) fitting the storage capacity with a transient model by reconstructing head drawdown induced by pumping activities. Finally, we test the relevance of our approach and evaluated the effect of considering or ignoring fossil gradients on aquifer parameterization for the Upper Mega Aquifer (UMA) on the Arabian Peninsula.

  8. Persistent organic pollutants in fat of three species of Pacific pelagic longline caught sea turtles: Accumulation in relation to ingested plastic marine debris

    USGS Publications Warehouse

    Clukey, Katharine; Lepczyk, Christopher A.; Balazs, George H.; Work, Thierry M.; Li, Qing X.; Bachman, Melanie J.; Lynch, Jennifer M.

    2017-01-01

    In addition to eating contaminated prey, sea turtles may be exposed to persistent organic pollutants (POPs) from ingesting plastic debris that has absorbed these chemicals. Given the limited knowledge about POPs in pelagic sea turtles and how plastic ingestion influences POP exposure, our objectives were to: 1) provide baseline contaminant levels of three species of pelagic Pacific sea turtles; and 2) assess trends of contaminant levels in relation to species, sex, length, body condition and capture location. In addition, we hypothesized that if ingesting plastic is a significant source of POP exposure, then the amount of ingested plastic may be correlated to POP concentrations accumulated in fat. To address our objectives we compared POP concentrations in fat samples to previously described amounts of ingested plastic from the same turtles. Fat samples from 25 Pacific pelagic sea turtles [2 loggerhead (Caretta caretta), 6 green (Chelonia mydas) and 17 olive ridley (Lepidochelys olivacea) turtles] were analyzed for 81 polychlorinated biphenyls (PCBs), 20 organochlorine pesticides, and 35 brominated flame-retardants. The olive ridley and loggerhead turtles had higher ΣDDTs (dichlorodiphenyltrichloroethane and metabolites) than ΣPCBs, at a ratio similar to biota measured in the South China Sea and southern California. Green turtles had a ratio close to 1:1. These pelagic turtles had lower POP levels than previously reported in nearshore turtles. POP concentrations were unrelated to the amounts of ingested plastic in olive ridleys, suggesting that their exposure to POPs is mainly through prey. In green turtles, concentrations of ΣPCBs were positively correlated with the number of plastic pieces ingested, but these findings were confounded by covariance with body condition index (BCI). Green turtles with a higher BCI had eaten more plastic and also had higher POPs. Taken together, our findings suggest that sea turtles accumulate most POPs through their prey rather

  9. Beyond fossil fuel-driven nitrogen transformations.

    PubMed

    Chen, Jingguang G; Crooks, Richard M; Seefeldt, Lance C; Bren, Kara L; Bullock, R Morris; Darensbourg, Marcetta Y; Holland, Patrick L; Hoffman, Brian; Janik, Michael J; Jones, Anne K; Kanatzidis, Mercouri G; King, Paul; Lancaster, Kyle M; Lymar, Sergei V; Pfromm, Peter; Schneider, William F; Schrock, Richard R

    2018-05-25

    Nitrogen is fundamental to all of life and many industrial processes. The interchange of nitrogen oxidation states in the industrial production of ammonia, nitric acid, and other commodity chemicals is largely powered by fossil fuels. A key goal of contemporary research in the field of nitrogen chemistry is to minimize the use of fossil fuels by developing more efficient heterogeneous, homogeneous, photo-, and electrocatalytic processes or by adapting the enzymatic processes underlying the natural nitrogen cycle. These approaches, as well as the challenges involved, are discussed in this Review. Copyright © 2018 The Authors, some rights reserved; exclusive licensee American Association for the Advancement of Science. No claim to original U.S. Government Works.

  10. Benthic-Pelagic Coupling: Effects on Nematode Communities along Southern European Continental Margins

    PubMed Central

    Pape, Ellen; Jones, Daniel O. B.; Manini, Elena; Bezerra, Tania Nara; Vanreusel, Ann

    2013-01-01

    Along a west-to-east axis spanning the Galicia Bank region (Iberian margin) and the Mediterranean basin, a reduction in surface primary productivity and in seafloor flux of particulate organic carbon was mirrored in the in situ organic matter quantity and quality within the underlying deep-sea sediments at different water depths (1200, 1900 and 3000 m). Nematode standing stock (abundance and biomass) and genus and trophic composition were investigated to evaluate downward benthic-pelagic coupling. The longitudinal decline in seafloor particulate organic carbon flux was reflected by a reduction in benthic phytopigment concentrations and nematode standing stock. An exception was the station sampled at the Galicia Bank seamount, where despite the maximal particulate organic carbon flux estimate, we observed reduced pigment levels and nematode standing stock. The strong hydrodynamic forcing at this station was believed to be the main cause of the local decoupling between pelagic and benthic processes. Besides a longitudinal cline in nematode standing stock, we noticed a west-to-east gradient in nematode genus and feeding type composition (owing to an increasing importance of predatory/scavenging nematodes with longitude) governed by potential proxies for food availability (percentage of nitrogen, organic carbon, and total organic matter). Within-station variability in generic composition was elevated in sediments with lower phytopigment concentrations. Standing stock appeared to be regulated by sedimentation rates and benthic environmental variables, whereas genus composition covaried only with benthic environmental variables. The coupling between deep-sea nematode assemblages and surface water processes evidenced in the present study suggests that it is likely that climate change will affect the composition and function of deep-sea nematodes. PMID:23565176

  11. Metabarcoding reveals environmental factors influencing spatio-temporal variation in pelagic micro-eukaryotes.

    PubMed

    Brannock, Pamela M; Ortmann, Alice C; Moss, Anthony G; Halanych, Kenneth M

    2016-08-01

    Marine environments harbour a vast diversity of micro-eukaryotic organisms (protists and other small eukaryotes) that play important roles in structuring marine ecosystems. However, micro-eukaryote diversity is not well understood. Likewise, knowledge is limited regarding micro-eukaryote spatial and seasonal distribution, especially over long temporal scales. Given the importance of this group for mobilizing energy from lower trophic levels near the base of the food chain to larger organisms, assessing community stability, diversity and resilience is important to understand ecosystem health. Herein, we use a metabarcoding approach to examine pelagic micro-eukaryote communities over a 2.5-year time series. Bimonthly surface sampling (July 2009 to December 2011) was conducted at four locations within Mobile Bay (Bay) and along the Alabama continental shelf (Shelf). Alpha-diversity only showed significant differences in Shelf sites, with the greatest differences observed between summer and winter. Beta-diversity showed significant differences in community composition in relation to season and the Bay was dominated by diatoms, while the Shelf was characterized by dinoflagellates and copepods. The northern Gulf of Mexico is heavily influenced by the Mobile River Basin, which brings low-salinity nutrient-rich water mostly during winter and spring. Community composition was correlated with salinity, temperature and dissolved silicate. However, species interactions (e.g. predation and parasitism) may also contribute to the observed variation, especially on the Shelf, which warrants further exploration. Metabarcoding revealed clear patterns in surface pelagic micro-eukaryote communities that were consistent over multiple years, demonstrating how these techniques could be greatly beneficial to ecological monitoring and management over temporal scales. © 2016 John Wiley & Sons Ltd.

  12. High-resolution δ13C record of fossil wood and bulk organic matter from a deep Oligocene lacustrine succession, Bach Long Vi Island, Vietnam

    NASA Astrophysics Data System (ADS)

    Rizzi, M.; Schovsbo, N. H.; Fyhn, M. B. W.; Korte, C.

    2017-12-01

    We present a high-resolution stable isotope record based on bulk organic matter (δ13Corg) and fossil wood (δ13Cwood) originating from Oligocene deep lacustrine sediments cored on the Bach Long Vi Island, northern Gulf of Tonkin, offshore Vietnam. The sediments are exceptionally well preserved. They are thus excellently suited for a detailed stratigraphical analysis of the stable isotope record and as proxy for environmental and climatic changes within this period. The sediments were deposited in rapid subsiding, narrow and elongated fault-bound graben (Fyhn and Phach, 2015) and are represented by deep pelagic lacustrine organic-rich mud interrupted by numerous density-flow deposits (Hovikoski et al., 2016). The density-flow deposits contain abundant fragments of fossil wood. Therefore it was possible to obtain 262 coalified wood fragments together with 1063 bulk organic samples throughout the span of the core. This allowed to establish a high resolution stable C isotope record (δ13Corg and δ13Cwood). In addition 2464 handheld XRF determinations were carried out to further characterize the depositional environment (Rizzi et al., 2017). The organic carbon isotope trend from the 500 m core succession provides insight into the palaeoenvironmental changes of the lake during the Oligocene. Both, global and local factors control the δ13C variations. The aim of the study is to obtain pure global δ13Corg and δ13Cwood signals that would allow comparison of the studied sediments with coeval syn-rift successions in the South China Sea region and other parts of the world. [1] Fyhn and Phach (2015) Tectonics, 34(2): 290-312. [2] Hovikoski et al. (2016) Journal of Sedimentary Research, 86(8): 982-1007. [3] Rizzi et al. (2017) EGU General Assembly Abstract EGU 2017-17584.

  13. Non-integumentary melanosomes can bias reconstructions of the colours of fossil vertebrate skin

    NASA Astrophysics Data System (ADS)

    McNamara, Maria; Kaye, Jonathan; Benton, Mike; Orr, Patrick

    2017-04-01

    The soft tissues of many fossil vertebrates preserve melanosomes - micron-scale organelles used to inform on original integumentary coloration and the evolution of visual signalling strategies through time. In extant vertebrates, however, melanosomes also occur in internal tissues, and hence melanosomes preserved in fossils may not derive solely from the integument. Here, by analyzing the internal tissues of extant and fossil frogs, we show that non-integumentary melanosomes are extremely abundant; they are usually localised to the torso in fossils but can also occur in the limbs, presumably due to dispersal during decay. Melanosomes from the body outlines of fossils cannot, therefore, reliably inform on integumentary coloration. Crucially, non-integumentary and integumentary melanosomes differ in geometry in both fossil and modern frogs and, in fossils, occur as discrete layers. Analysis of melanosome geometry, distribution and size-specific layering is required to differentiate integumentary from non-integumentary melanosomes and is essential to any attempt to reconstruct the original colours of vertebrate skin.

  14. The age for the fossil-bearing Tabbowa beds in Sri Lanka

    NASA Astrophysics Data System (ADS)

    Chang, S. C.; Dassanayake, S.; Wang, J.

    2014-12-01

    Well-preserved terrestrial fossils, mainly including conifers, cycads and ferns, were discovered from the Tabbowa beds in northwestern Sri Lanka. The high diversity and abundance of plants and insects from these Jurassic sediments provide a unique window to understand floral evolution and plant-insect co-evolution in the Mesozoic. For example, unearthed fossils from the Tabbowa beds indicate that leaf feeding and dwelling insects played a significant role in the Jurassic ecosystem. For another example, feeding and chewing marks on leaves allow studying insect behavior and paleo-ecology. Additionally, the recent discoveries of Otozamites latiphyllus and Otozamites tabbowensis from these sediments provide evidence that Bennettitales, an extinct order of seed plants, widely spread in the Gondwana during the Jurassic period. Although most fossils are yet to be well studied, and only few of the fossil occurrences have been published in western journals, plant fossils from the Tabbowa beds have great potential for substantially increasing our knowledge of Jurassic terrestrial ecosystems. The fossil-bearing Tabbowa beds are mainly composed of sandstone, siltstone, and mudstone with occasional thin bands of nodular limestone. Until now, radio-isotopic age determinations for the fossil-rich Tabbowa beds are lacking. In this study, we investigate the geological and geochronological setting of this area by dating detrital zircons from the Tabbowa beds. The age data will allow testing several hypotheses regarding the plant evolution, the basin development of this region.

  15. Fossilized Mammalian Erythrocytes Associated With a Tick Reveal Ancient Piroplasms.

    PubMed

    Poinar, George

    2017-07-01

    Ticks transmit a variety of pathogenic organisms to vertebrates, especially mammals. The fossil record of such associations is extremely rare. An engorged nymphal tick of the genus Ambylomma in Dominican amber was surrounded by erythrocytes from its mammalian host. Some of the exposed erythrocytes contained developmental stages of a hemoprotozoan resembling members of the Order Piroplasmida. The fossil piroplasm is described, its stages compared with those of extant piroplasms, and reasons provided why the mammalian host could have been a primate. The parasites were also found in the gut epithelial cells and body cavity of the fossil tick. Aside from providing the first fossil mammalian red blood cells and the first fossil intraerythrocytic hemoparasites, the present discovery shows that tick-piroplasm associations were already well established in the Tertiary. This discovery provides a timescale that can be used in future studies on the evolution of the Piroplasmida. © The Authors 2017. Published by Oxford University Press on behalf of Entomological Society of America. All rights reserved. For Permissions, please email: journals.permissions@oup.com Version of Record, first published online March 20, 2017 with fixed content and layout in compliance with Art. 8.1.3.2 ICZN.

  16. The Dynamics of Vertical Migration in the Oceanic Gulf of Mexico after Deepwater Horizon: Active Linkage of Large Vertebrates and Deep-Pelagic Nekton

    NASA Astrophysics Data System (ADS)

    Sutton, T.; Cook, A.; Frank, T. M.; Boswell, K. M.; Vecchione, M.; Judkins, H.; Romero, I.

    2016-02-01

    Toothed whales, smaller cetaceans, seabirds, and epipelagic gamefishes rely on deep-pelagic (meso- and bathypelagic) nekton as primary or secondary prey. This trophic interaction is mediated by downward and upward vertical movements (e.g., sperm whale diving and lanternfishes migration, respectively). This interaction also links particle-feeding lower trophic levels with top predators in a manner that spans the gamut of depth domains. This is particularly important with respect to a whole-water column disturbance such as the Deepwater Horizon oil spill (DWHOS). Here we present highly resolved vertical distribution and migration data collected during a large-scale, NOAA-supported, deep-pelagic (0-1500 m) survey in 2011, along with data collected during ongoing GoMRI-supported DEEPEND consortium surveys. The deep-pelagic nekton community of the Gulf of Mexico is a complex mixture of migrating, non-migrating, and partially migrating assemblages that connect surface waters with depths in excess of 1000 m. Major patterns of vertical distribution for 400+ species of fishes, cephalopods, and macrocrustaceans, the primary prey of many important species of oceanic vertebrates living near-surface, will be summarized and quantified with the goal of highlighting potential vectors of anthropogenic contamination transfer in the deep-pelagial, the Gulf's largest ecosystem.

  17. Using more than the oldest fossils: dating osmundaceae with three Bayesian clock approaches.

    PubMed

    Grimm, Guido W; Kapli, Paschalia; Bomfleur, Benjamin; McLoughlin, Stephen; Renner, Susanne S

    2015-05-01

    A major concern in molecular clock dating is how to use information from the fossil record to calibrate genetic distances from DNA sequences. Here we apply three Bayesian dating methods that differ in how calibration is achieved-"node dating" (ND) in BEAST, "total evidence" (TE) dating in MrBayes, and the "fossilized birth-death" (FBD) in FDPPDiv-to infer divergence times in the royal ferns. Osmundaceae have 16-17 species in four genera, two mainly in the Northern Hemisphere and two in South Africa and Australasia; they are the sister clade to the remaining leptosporangiate ferns. Their fossil record consists of at least 150 species in ∼17 genera. For ND, we used the five oldest fossils, whereas for TE and FBD dating, which do not require forcing fossils to nodes and thus can use more fossils, we included up to 36 rhizomes and frond compression/impression fossils, which for TE dating were scored for 33 morphological characters. We also subsampled 10%, 25%, and 50% of the 36 fossils to assess model sensitivity. FBD-derived divergence ages were generally greater than those inferred from ND; two of seven TE-derived ages agreed with FBD-obtained ages, the others were much younger or much older than ND or FBD ages. We prefer the FBD-derived ages because they best fit the Osmundales fossil record (including Triassic fossils not used in our study). Under the preferred model, the clade encompassing extant Osmundaceae (and many fossils) dates to the latest Paleozoic to Early Triassic; divergences of the extant species occurred during the Neogene. Under the assumption of constant speciation and extinction rates, the FBD approach yielded speciation and extinction rates that overlapped those obtained from just neontological data. However, FBD estimates of speciation and extinction are sensitive to violations in the assumption of continuous fossil sampling; therefore, these estimates should be treated with caution. © The Author(s) 2014. Published by Oxford University Press

  18. Absolute measures of the completeness of the fossil record

    NASA Technical Reports Server (NTRS)

    Foote, M.; Sepkoski, J. J. Jr; Sepkoski JJ, J. r. (Principal Investigator)

    1999-01-01

    Measuring the completeness of the fossil record is essential to understanding evolution over long timescales, particularly when comparing evolutionary patterns among biological groups with different preservational properties. Completeness measures have been presented for various groups based on gaps in the stratigraphic ranges of fossil taxa and on hypothetical lineages implied by estimated evolutionary trees. Here we present and compare quantitative, widely applicable absolute measures of completeness at two taxonomic levels for a broader sample of higher taxa of marine animals than has previously been available. We provide an estimate of the probability of genus preservation per stratigraphic interval, and determine the proportion of living families with some fossil record. The two completeness measures use very different data and calculations. The probability of genus preservation depends almost entirely on the Palaeozoic and Mesozoic records, whereas the proportion of living families with a fossil record is influenced largely by Cenozoic data. These measurements are nonetheless highly correlated, with outliers quite explicable, and we find that completeness is rather high for many animal groups.

  19. First record of Podocarpoid fossil wood in South China

    PubMed Central

    Li, Long; Jin, Jian-Hua; Quan, Cheng; Oskolski, Alexei A.

    2016-01-01

    A new species of fossil conifer wood, Podocarpoxylon donghuaiense sp. nov., is described from the late Eocene of Nadu Formation in Baise Basin of the Guangxi Province, South China. This fossil wood is characterized by distinct growth rings, circular to oval tracheids in cross section, 1–2-seriate opposite pits on radial tracheid walls, uniseriate (rarely biseriate) rays, smooth end walls of ray parenchyma cells, and the absence of resin ducts, suggesting its affinity to Podocarpaceae. The new species is distinctive from other Cenozoic woods ascribed to this family by the combination of distinctive growth rings, the absence of axial parenchyma, the occurrence of bordered pits on tangential tracheid walls, and the occurrence of 3–4 cuppressoid or taxodioid pits on cross-fields. This represents the first record of podocarpoid fossil wood in South China and provides fossil evidence for the early dispersal and diversification of Podocarpaceae in eastern Asia as well as for mild temperate seasonal climate in this region during the late Eocene. PMID:27571780

  20. Building Fossils in the Elementary School and Writing about Them Using Computers.

    ERIC Educational Resources Information Center

    Schlenker, Richard M.; Yoshida, Sarah

    This material describes a fossil-building activity using sea shells, chicken bones, and plaster for grade one through three students. Related process skills, vocabulary, computer principles, time requirements, and materials are listed. Two methods of building the fossils are discussed. After building the fossils, classes may be divided into pairs…

  1. Fossil group origins. III. The relation between optical and X-ray luminosities

    NASA Astrophysics Data System (ADS)

    Girardi, M.; Aguerri, J. A. L.; De Grandi, S.; D'Onghia, E.; Barrena, R.; Boschin, W.; Méndez-Abreu, J.; Sánchez-Janssen, R.; Zarattini, S.; Biviano, A.; Castro-Rodriguez, N.; Corsini, E. M.; del Burgo, C.; Iglesias-Páramo, J.; Vilchez, J. M.

    2014-05-01

    Aims: This study is part of the Fossil group origins (FOGO) project which aims to carry out a systematic and multiwavelength study of a large sample of fossil systems. Here we focus on the relation between the optical luminosity (Lopt) and X-ray luminosity (LX). Methods: Out of a total sample of 28 candidate fossil systems, we consider a sample of 12 systems whose fossil classification has been confirmed by a companion study. They are compared with the complementary sample of 16 systems whose fossil nature has not been confirmed and with a subsample of 102 galaxy systems from the RASS-SDSS galaxy cluster survey. Fossil and normal systems span the same redshift range 0 fossil system, the LX in the 0.1-2.4 keV band is computed using data from the ROSAT All Sky Survey to be comparable to the estimates of the comparison sample. For each fossil and normal system we homogeneously compute Lopt in the r-band within the characteristic cluster radius, using data from the Sloan Digital Sky Survey Data Release 7. Results: We sample the LX-Lopt relation over two orders of magnitude in LX. Our analysis shows that fossil systems are not statistically distinguishable from the normal systems through the 2D Kolmogorov-Smirnov test nor the fit of the LX-Lopt relation. Thus, the optical luminosity of the galaxy system does strongly correlate with the X-ray luminosity of the hot gas component, independently of whether the system is fossil or not. We discuss our results in comparison with previous literature. Conclusions: We conclude that our results are consistent with the classical merging scenario of the brightest galaxy formed via merger/cannibalism of other group galaxies with conservation of the optical light. We find no evidence for a peculiar state of the hot intracluster medium. Tables 1 and 2 are available in electronic form at http://www.aanda.org

  2. Fossils of reionization in the local group

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Gnedin, Nickolay Y.; /Fermilab /KICP, Chicago /Chicago U., EFI; Kravtsov, Andrey V.

    We use a combination of high-resolution gas dynamics simulations of high-redshift dwarf galaxies and dissipationless simulations of a Milky Way sized halo to estimate the expected abundance and spatial distribution of the dwarf satellite galaxies that formed most of their stars around z {approx} 8 and evolved only little since then. Such galaxies can be considered as fossils of the reionization era, and studying their properties could provide a direct window into the early, pre-reionization stages of galaxy formation. We show that 5-15% of the objects existing at z {approx} 8 do indeed survive until the present in the MWmore » like environment without significant evolution. This implies that it is plausible that the fossil dwarf galaxies do exist in the Local Group. Because such galaxies form their stellar systems early during the period of active merging and accretion, they should have spheroidal morphology regardless of their current distance from the host galaxy. We show that both the expected luminosity function and spatial distribution of dark matter halos which are likely to host fossil galaxies agree reasonably well with the observed distributions of the luminous (L{sub V} > 10{sup 6} Lsun) Local Group fossil candidates near the host galaxy (d<200 kpc). However, the predicted abundance is substantially larger (by a factor of 2-3) for fainter galaxies (L{sub V} < 10{sup 6} Lsun) at larger distances (d>300 kpc). We discuss several possible explanations for this discrepancy.« less

  3. Unprecedented Proliferation of Novel Pelagic Sargassum Form has Implications for Ecosystem Function and Regional Diversity in the Caribbean

    NASA Astrophysics Data System (ADS)

    Siuda, A. N.; Schell, J. M.; Goodwin, D. S.

    2016-02-01

    Pelagic Sargassum is a planktonic macroalgae comprised of two species, S. fluitans and S. natans, each exhibiting a variety of morphological forms; it is found throughout the North Atlantic, Caribbean Sea, and Gulf of Mexico. Drifting open ocean Sargassum provides essential habitat and food resources to organisms across multiple trophic levels, from resident shrimp to migratory sea turtles. Historic observations, including Sea Education Association's (SEA) 22-year field sampling dataset, indicate that S. natans-I and S. fluitans-III are most common and that S. natans-VIII is rare. Furthermore, SEA's long-term record shows very low pelagic Sargassum abundance in the Eastern Caribbean in contrast to the Sargasso Sea. During April 2014, Sargassum began washing ashore along Caribbean coastlines in unprecedented quantities. Shipboard observations of the recent inundation event occurred November 2014 to May 2015. In total, 30.5 kg of pelagic Sargassum was collected in 92.6% of surface neuston tows, sorted and weighed by morphological form. Notably, the predominant Sargassum form observed during the 2014/15 event is S. natans-VIII, a documented change in Sargassum diversity. Strong spatial patterns were also observed, with S. natans-VIII dominant in the Western Tropical Atlantic (87.3% wet weight) and Eastern Caribbean (95.3% wet weight) and S. natans-I most common in the South Sargasso Sea (87.5% wet weight). S. fluitans-III was observed in low abundance across all regions. These sudden assemblage and abundance changes, the biophysical mechanisms of which are not yet understood, have significant ecological consequences at multiple scales. Impacts to associated mobile fauna diversity and community structure, dependent fisheries and iconic species, and coastal ecosystem function will echo throughout the Caribbean, and should comprise focal areas of future research efforts across the region.

  4. 50 CFR Figure 5 to Part 679 - Kodiak Island Closure Status for Vessels Using Non-pelagic Trawl Gear

    Code of Federal Regulations, 2012 CFR

    2012-10-01

    ... 50 Wildlife and Fisheries 13 2012-10-01 2012-10-01 false Kodiak Island Closure Status for Vessels Using Non-pelagic Trawl Gear 5 Figure 5 to Part 679 Wildlife and Fisheries FISHERY CONSERVATION AND MANAGEMENT, NATIONAL OCEANIC AND ATMOSPHERIC ADMINISTRATION, DEPARTMENT OF COMMERCE (CONTINUED) FISHERIES OF THE EXCLUSIVE ECONOMIC ZONE OFF ALASKA P...

  5. 50 CFR Figure 5 to Part 679 - Kodiak Island Closure Status for Vessels Using Non-pelagic Trawl Gear

    Code of Federal Regulations, 2013 CFR

    2013-10-01

    ... 50 Wildlife and Fisheries 13 2013-10-01 2013-10-01 false Kodiak Island Closure Status for Vessels Using Non-pelagic Trawl Gear 5 Figure 5 to Part 679 Wildlife and Fisheries FISHERY CONSERVATION AND MANAGEMENT, NATIONAL OCEANIC AND ATMOSPHERIC ADMINISTRATION, DEPARTMENT OF COMMERCE (CONTINUED) FISHERIES OF THE EXCLUSIVE ECONOMIC ZONE OFF ALASKA P...

  6. 50 CFR Figure 5 to Part 679 - Kodiak Island Closure Status for Vessels Using Non-pelagic Trawl Gear

    Code of Federal Regulations, 2010 CFR

    2010-10-01

    ... 50 Wildlife and Fisheries 9 2010-10-01 2010-10-01 false Kodiak Island Closure Status for Vessels Using Non-pelagic Trawl Gear 5 Figure 5 to Part 679 Wildlife and Fisheries FISHERY CONSERVATION AND MANAGEMENT, NATIONAL OCEANIC AND ATMOSPHERIC ADMINISTRATION, DEPARTMENT OF COMMERCE (CONTINUED) FISHERIES OF THE EXCLUSIVE ECONOMIC ZONE OFF ALASKA Pt...

  7. 50 CFR Figure 5 to Part 679 - Kodiak Island Closure Status for Vessels Using Non-pelagic Trawl Gear

    Code of Federal Regulations, 2011 CFR

    2011-10-01

    ... 50 Wildlife and Fisheries 11 2011-10-01 2011-10-01 false Kodiak Island Closure Status for Vessels Using Non-pelagic Trawl Gear 5 Figure 5 to Part 679 Wildlife and Fisheries FISHERY CONSERVATION AND MANAGEMENT, NATIONAL OCEANIC AND ATMOSPHERIC ADMINISTRATION, DEPARTMENT OF COMMERCE (CONTINUED) FISHERIES OF THE EXCLUSIVE ECONOMIC ZONE OFF ALASKA P...

  8. Preservation of three-dimensional anatomy in phosphatized fossil arthropods enriches evolutionary inference.

    PubMed

    Schwermann, Achim H; Dos Santos Rolo, Tomy; Caterino, Michael S; Bechly, Günter; Schmied, Heiko; Baumbach, Tilo; van de Kamp, Thomas

    2016-02-05

    External and internal morphological characters of extant and fossil organisms are crucial to establishing their systematic position, ecological role and evolutionary trends. The lack of internal characters and soft-tissue preservation in many arthropod fossils, however, impedes comprehensive phylogenetic analyses and species descriptions according to taxonomic standards for Recent organisms. We found well-preserved three-dimensional anatomy in mineralized arthropods from Paleogene fissure fillings and demonstrate the value of these fossils by utilizing digitally reconstructed anatomical structure of a hister beetle. The new anatomical data facilitate a refinement of the species diagnosis and allowed us to reject a previous hypothesis of close phylogenetic relationship to an extant congeneric species. Our findings suggest that mineralized fossils, even those of macroscopically poor preservation, constitute a rich but yet largely unexploited source of anatomical data for fossil arthropods.

  9. Microbial Fossilization in Mineralizing Environments: Relevance for Mars "EXOPALEONTOLOGY"

    NASA Technical Reports Server (NTRS)

    Farmer, Jack D.; DesMarais, David J.; Morrison, David (Technical Monitor)

    1994-01-01

    The goals of post-Viking exobiology include the search for a Martian fossil record. How can we optimize future exploration efforts to search for fossils on Mars? The Precambrian fossil record indicates that key factors for the long-term preservation of microbial fossils include: 1) the rapid entombment and/or replacement of organisms and organic matter by fine-grained, stable mineral phases (e.g. silica, phosphate, and to a lesser extent, carbonate), 2) low-permeability host sediments (maintaining a closed chemical system during early diagenesis), and 3) shallow burial (maintaining post-depositional temperatures and pressures within the stability range for complex organic molecules). Modem terrestrial environments where early mineralization commonly occurs in association with microbial organisms include: subaerial thermal springs and shallow hydrothermal systems, sub-lacustrine springs and evaporites of alkaline lakes, and subsoil environments where hardpans (e.g. calcretes, silcretes) and duricrusts form. Studies of microbial fossilization in such environments provide important insights preservation patterns in Precambrian rocks, while also playing a role in the development of strategies for Mars exopaleontology. The refinement of site priorities for Mars exopaleontology is expected to benefit greatly from high resolution imaging and altimetry acquired during upcoming orbital missions, and especially infrared and gamma ray spectral data needed for determining surface composition. In anticipation of future orbital missions, constraints for identifying high priority mineral deposits on Mars are being developed through analog remote sensing studies of key mineralizing environments on Earth.

  10. Hexabromocyclododecane affects benthic-pelagic coupling in an experimental ecosystem.

    PubMed

    Bradshaw, Clare; Näslund, Johan; Hansen, Joakim; Kozlowsky-Suzuki, Betina; Sundström, Bo; Gustafsson, Kerstin

    2015-11-01

    Hexabromocyclododecane (HBCDD) is an additive brominated flame retardant and a recognized PBT chemical. However, little is known about its effects on coastal species, and even less on ecosystem effects. We investigated the dose-response effects of HBCDD over 8 months in 1000 L experimental mesocosms assembled from coastal Baltic Sea ecosystem components. HBCDD was added via spiked plankton material and a range of structural and functional endpoints were measured during the experiment. Increasing HBCDD concentration decreased the biomass of large Macoma balthica, resulting in a decreased recirculation of nutrients to the water. Changes in plankton communities were also observed, either due to direct toxic HBCDD effects or indirect via changes in benthic-pelagic coupling of nutrients. Such complex ecosystem responses can only be quantified and understood by using realistic experimental set-ups, and including knowledge of system-specific ecological interactions. This is the first study of HBCDD effects on ecosystem level. Copyright © 2015 Elsevier Ltd. All rights reserved.

  11. Meso- and bathy-pelagic fish parasites at the Mid-Atlantic Ridge (MAR): Low host specificity and restricted parasite diversity

    NASA Astrophysics Data System (ADS)

    Klimpel, Sven; Busch, Markus Wilhelm; Sutton, Tracey; Palm, Harry Wilhelm

    2010-04-01

    Seven meso- and bathy-pelagic fish species from the Mid-Atlantic Ridge (MAR) were firstly studied for fish parasites and feeding ecology. With a total of seven parasite species, the 247 meso- and bathy-pelagic deep-sea fish specimens belonging to the families Melamphaidae (3 spp.), Myctophidae (3 spp.) and Stomiidae (1 sp.) revealed low parasite diversity. The genetically identified nematodes Anisakis simplex (s.s.) and Anisakis pegreffii from the body cavity, liver and muscles of Myctophum punctatum were the most abundant parasites, reaching a prevalence of 91.4% and mean intensity of 3.1 (1-14). Anisakis sp. (unidentified) infected Chauliodus sloani and Poromitra crassiceps. Bothriocephalidean and tetraphyllidean cestode larvae infected Benthosema glaciale, the latter also occurring in C. sloani and Scopelogadus beanii, at low prevalences. Adult parasites at low infection rates included the digenean Lethadena sp. (2.9%), and the two copepod species Sarcotretes scopeli (5.7%) and Tautochondria dolichoura (5.3-11.4%). The myctophid Lampanyctus macdonaldi and the melamphaid Scopelogadus mizolepis mizolepis were free of parasites. Analyses of the stomach contents revealed crustaceans, especially copepods and euphausiids for the myctophids and also amphipods for the melamphaids as predominant prey items. While all stomachs showing distinct content comprising often unidentified 'tissue' (possibly gelatinous zooplankton), only C. sloani preyed upon fish. Though this feeding habit would enable transfer of a variety of crustacean-transmitted parasites into the fish, the parasite fauna in the meso- and bathy-pelagic fish was species poor. All observed parasites showed low host specificity, demonstrating no distinct pattern of host-parasite co-evolution. The MAR is no barrier for the parasite distribution in the North Atlantic meso- and bathy-pelagial.

  12. Material Flow Analysis of Fossil Fuels in China during 2000–2010

    PubMed Central

    Wang, Sheng; Dai, Jing; Su, Meirong

    2012-01-01

    Since the relationship between the supply and demand of fossil fuels is on edge in the long run, the contradiction between the economic growth and limited resources will hinder the sustainable development of the Chinese society. This paper aims to analyze the input of fossil fuels in China during 2000–2010 via the material flow analysis (MFA) that takes hidden flows into account. With coal, oil, and natural gas quantified by MFA, three indexes, consumption and supply ratio (C/S ratio), resource consumption intensity (RCI), and fossil fuels productivity (FFP), are proposed to reflect the interactions between population, GDP, and fossil fuels. The results indicated that in the past 11 years, China's requirement for fossil fuels has been increasing continuously because of the growing mine productivity in domestic areas, which also leads to a single energy consumption structure as well as excessive dependence on the domestic exploitation. It is advisable to control the fossil fuels consumption by energy recycling and new energy facilities' popularization in order to lead a sustainable access to nonrenewable resources and decrease the soaring carbon emissions. PMID:23365525

  13. A comprehensive database of quality-rated fossil ages for Sahul's Quaternary vertebrates.

    PubMed

    Rodríguez-Rey, Marta; Herrando-Pérez, Salvador; Brook, Barry W; Saltré, Frédérik; Alroy, John; Beeton, Nicholas; Bird, Michael I; Cooper, Alan; Gillespie, Richard; Jacobs, Zenobia; Johnson, Christopher N; Miller, Gifford H; Prideaux, Gavin J; Roberts, Richard G; Turney, Chris S M; Bradshaw, Corey J A

    2016-07-19

    The study of palaeo-chronologies using fossil data provides evidence for past ecological and evolutionary processes, and is therefore useful for predicting patterns and impacts of future environmental change. However, the robustness of inferences made from fossil ages relies heavily on both the quantity and quality of available data. We compiled Quaternary non-human vertebrate fossil ages from Sahul published up to 2013. This, the FosSahul database, includes 9,302 fossil records from 363 deposits, for a total of 478 species within 215 genera, of which 27 are from extinct and extant megafaunal species (2,559 records). We also provide a rating of reliability of individual absolute age based on the dating protocols and association between the dated materials and the fossil remains. Our proposed rating system identified 2,422 records with high-quality ages (i.e., a reduction of 74%). There are many applications of the database, including disentangling the confounding influences of hypothetical extinction drivers, better spatial distribution estimates of species relative to palaeo-climates, and potentially identifying new areas for fossil discovery.

  14. Impacts of curatorial and research practices on the preservation of fossil hominid remains.

    PubMed

    Le Cabec, Adeline; Toussaint, Michel

    2017-12-30

    Fossil remains are the only physical evidence of past forms of life which researchers can use to study the evolutionary biology of a species, especially regarding the human lineage. We review and consider the way in which the conditions surrounding a fossil's discovery and its use for scientific research impacts its long-term preservation. The deterioration of the body starts soon after death, continues in the sediments and only a subsample of the anatomical elements will persist and may finally be unearthed by archeologists. From their recovery onwards, fossil remains are exposed to many sources of further damage: from handling, restoration, measuring to invasive sampling. On the one hand, curators are faced with the inherent challenge of balancing their responsibility to protect fossil specimens with allowing researchers to perform specific analyses or invasive sampling detrimental to the preservation of the fossil. On the other hand, scientists may find their analyses complicated by multiple factors including taphonomy, or restoration techniques (e.g., consolidants, cleaning chemicals). We provide several historical examples illustrating the complex nature of the factors acting on fossil preservation. We discuss concerns about producing and sharing (digital) data from fossils. Finally, we also suggest and support some curatorial practices which maximize the traceability of treatments underwent by a fossil.

  15. Hydrogen production econometric studies. [hydrogen and fossil fuels

    NASA Technical Reports Server (NTRS)

    Howell, J. R.; Bannerot, R. B.

    1975-01-01

    The current assessments of fossil fuel resources in the United States were examined, and predictions of the maximum and minimum lifetimes of recoverable resources according to these assessments are presented. In addition, current rates of production in quads/year for the fossil fuels were determined from the literature. Where possible, costs of energy, location of reserves, and remaining time before these reserves are exhausted are given. Limitations that appear to hinder complete development of each energy source are outlined.

  16. Climate variability and change scenarios for a marine commodity: Modelling small pelagic fish, fisheries and fishmeal in a globalized market

    NASA Astrophysics Data System (ADS)

    Merino, Gorka; Barange, Manuel; Mullon, Christian

    2010-04-01

    The world's small pelagic fish populations, their fisheries, fishmeal and fish oil production industries and markets are part of a globalised production and consumption system. The potential for climate variability and change to alter the balance in this system is explored by means of bioeconomic models at two different temporal scales, with the objective of investigating the interactive nature of environmental and human-induced changes on this globalised system. Short-term (interannual) environmental impacts on fishmeal production are considered by including an annual variable production rate on individual small pelagic fish stocks over a 10-year simulation period. These impacts on the resources are perceived by the fishmeal markets, where they are confronted by two aquaculture expansion hypotheses. Long-term (2080) environmental impacts on the same stocks are estimated using long-term primary production predictions as proxies for the species' carrying capacities, rather than using variable production rates, and are confronted on the market side by two alternative fishmeal management scenarios consistent with IPCC-type storylines. The two scenarios, World Markets and Global Commons, are parameterized through classic equilibrium solutions for a global surplus production bioeconomic model, namely maximum sustainable yield and open access, respectively. The fisheries explicitly modelled in this paper represent 70% of total fishmeal production, thus encapsulating the expected dynamics of the global production and consumption system. Both short and long-term simulations suggest that the sustainability of the small pelagic resources, in the face of climate variability and change, depends more on how society responds to climate impacts than on the magnitude of climate alterations per se.

  17. Peaches Preceded Humans: Fossil Evidence from SW China

    PubMed Central

    Su, Tao; Wilf, Peter; Huang, Yongjiang; Zhang, Shitao; Zhou, Zhekun

    2015-01-01

    Peach (Prunus persica, Rosaceae) is an extremely popular tree fruit worldwide, with an annual production near 20 million tons. Peach is widely thought to have origins in China, but its evolutionary history is largely unknown. The oldest evidence for the peach has been Chinese archaeological records dating to 8000–7000 BP. Here, we report eight fossil peach endocarps from late Pliocene strata of Kunming City, Yunnan, southwestern China. The fossils are identical to modern peach endocarps, including size comparable to smaller modern varieties, a single seed, a deep dorsal groove, and presence of deep pits and furrows. These fossils show that China has been a critical region for peach evolution since long before human presence, much less agriculture. Peaches evolved their modern morphology under natural selection, presumably involving large, frugivorous mammals such as primates. Much later, peach size and variety increased through domestication and breeding. PMID:26610240

  18. Fossil Liposcelididae and the lice ages (Insecta: Psocodea)

    PubMed Central

    Grimaldi, David; Engel, Michael S

    2005-01-01

    Fossilized, winged adults belonging to the psocopteran family Liposcelididae are reported in amber from the mid-Cretaceous (ca 100 Myr) of Myanmar (described as Cretoscelis burmitica, gen. et sp. n.) and the Miocene (ca 20 Myr) of the Dominican Republic (Belaphopsocus dominicus sp. n.). Cretoscelis is an extinct sister group to all other Liposcelididae and the family is the free-living sister group to the true lice (order Phthiraptera, all of which are ectoparasites of birds and mammals). A phylogenetic hypothesis of relationships among genera of Liposcelididae, including fossils, reveals perfect correspondence between the chronology of fossils and cladistic rank of taxa. Lice and Liposcelididae minimally diverged 100 Myr, perhaps even in the earliest Cretaceous 145 Myr or earlier, in which case the hosts of lice would have been early mammals, early birds and possibly other feathered theropod dinosaurs, as well as haired pterosaurs. PMID:16537135

  19. Peaches Preceded Humans: Fossil Evidence from SW China

    NASA Astrophysics Data System (ADS)

    Su, Tao; Wilf, Peter; Huang, Yongjiang; Zhang, Shitao; Zhou, Zhekun

    2015-11-01

    Peach (Prunus persica, Rosaceae) is an extremely popular tree fruit worldwide, with an annual production near 20 million tons. Peach is widely thought to have origins in China, but its evolutionary history is largely unknown. The oldest evidence for the peach has been Chinese archaeological records dating to 8000-7000 BP. Here, we report eight fossil peach endocarps from late Pliocene strata of Kunming City, Yunnan, southwestern China. The fossils are identical to modern peach endocarps, including size comparable to smaller modern varieties, a single seed, a deep dorsal groove, and presence of deep pits and furrows. These fossils show that China has been a critical region for peach evolution since long before human presence, much less agriculture. Peaches evolved their modern morphology under natural selection, presumably involving large, frugivorous mammals such as primates. Much later, peach size and variety increased through domestication and breeding.

  20. Ocean-wide tracking of pelagic sharks reveals extent of overlap with longline fishing hotspots.

    PubMed

    Queiroz, Nuno; Humphries, Nicolas E; Mucientes, Gonzalo; Hammerschlag, Neil; Lima, Fernando P; Scales, Kylie L; Miller, Peter I; Sousa, Lara L; Seabra, Rui; Sims, David W

    2016-02-09

    Overfishing is arguably the greatest ecological threat facing the oceans, yet catches of many highly migratory fishes including oceanic sharks remain largely unregulated with poor monitoring and data reporting. Oceanic shark conservation is hampered by basic knowledge gaps about where sharks aggregate across population ranges and precisely where they overlap with fishers. Using satellite tracking data from six shark species across the North Atlantic, we show that pelagic sharks occupy predictable habitat hotspots of high space use. Movement modeling showed sharks preferred habitats characterized by strong sea surface-temperature gradients (fronts) over other available habitats. However, simultaneous Global Positioning System (GPS) tracking of the entire Spanish and Portuguese longline-vessel fishing fleets show an 80% overlap of fished areas with hotspots, potentially increasing shark susceptibility to fishing exploitation. Regions of high overlap between oceanic tagged sharks and longliners included the North Atlantic Current/Labrador Current convergence zone and the Mid-Atlantic Ridge southwest of the Azores. In these main regions, and subareas within them, shark/vessel co-occurrence was spatially and temporally persistent between years, highlighting how broadly the fishing exploitation efficiently "tracks" oceanic sharks within their space-use hotspots year-round. Given this intense focus of longliners on shark hotspots, our study argues the need for international catch limits for pelagic sharks and identifies a future role of combining fine-scale fish and vessel telemetry to inform the ocean-scale management of fisheries.

  1. Ocean-wide tracking of pelagic sharks reveals extent of overlap with longline fishing hotspots

    PubMed Central

    Queiroz, Nuno; Humphries, Nicolas E.; Hammerschlag, Neil; Miller, Peter I.; Sousa, Lara L.; Seabra, Rui; Sims, David W.

    2016-01-01

    Overfishing is arguably the greatest ecological threat facing the oceans, yet catches of many highly migratory fishes including oceanic sharks remain largely unregulated with poor monitoring and data reporting. Oceanic shark conservation is hampered by basic knowledge gaps about where sharks aggregate across population ranges and precisely where they overlap with fishers. Using satellite tracking data from six shark species across the North Atlantic, we show that pelagic sharks occupy predictable habitat hotspots of high space use. Movement modeling showed sharks preferred habitats characterized by strong sea surface-temperature gradients (fronts) over other available habitats. However, simultaneous Global Positioning System (GPS) tracking of the entire Spanish and Portuguese longline-vessel fishing fleets show an 80% overlap of fished areas with hotspots, potentially increasing shark susceptibility to fishing exploitation. Regions of high overlap between oceanic tagged sharks and longliners included the North Atlantic Current/Labrador Current convergence zone and the Mid-Atlantic Ridge southwest of the Azores. In these main regions, and subareas within them, shark/vessel co-occurrence was spatially and temporally persistent between years, highlighting how broadly the fishing exploitation efficiently “tracks” oceanic sharks within their space-use hotspots year-round. Given this intense focus of longliners on shark hotspots, our study argues the need for international catch limits for pelagic sharks and identifies a future role of combining fine-scale fish and vessel telemetry to inform the ocean-scale management of fisheries. PMID:26811467

  2. Changing Abundance of Magnetofossil Morphologies in Pelagic Red Clay Around Minamitorishima, Western North Pacific

    NASA Astrophysics Data System (ADS)

    Usui, Yoichi; Yamazaki, Toshitsugu; Saitoh, Masafumi

    2017-12-01

    Recent investigations have discovered an unexpected abundance of magnetofossils in oxic pelagic red clays. These have potential to serve as paleoenvironmental tracers in otherwise nonfossiliferous sediments. Here, we report on variations in the abundance and morphology of magnetofossils in red clay from the western North Pacific. Magnetic measurements revealed that magnetofossils dominate the magnetic mineral assemblage of the sediments. An endmember analysis of isothermal remanent magnetization acquisition curves, supplemented by an analysis of S ratios, indicates that the magnetic assemblage can be unmixed into three endmembers, two corresponding to magnetofossils and one to terrigenous magnetic minerals. Direct counting of magnetofossil morphologies under a transmission electron microscope shows that the two magnetofossil endmembers differentiate equant magnetofossils and bullet-shaped magnetofossils, respectively. The stratigraphic variation of the endmember contributions revealed that the equant magnetofossils are dominant for the most part, while an interval at around 7 m in core depth shows higher abundance of the bullet-shaped magnetofossils. This may reflect enhanced organic carbon flux to the sediments. The organic carbon content is low throughout the sediments, and it does not show any change corresponding to the increase of bullet-shaped magnetofossils, pointing at extensive remineralization of the organic carbon. On the basis of lithostratigraphic correlation to nearby drilling sites, we tentatively estimate the age of the bullet-shaped magnetofossil increase as sometime between ˜75 and 25 Ma. These results suggest that environmental information can be obtained from magnetofossils in pelagic red clay.

  3. Climate Change Impacts to North Pacific Pelagic Habitat Are Projected to Lower Carrying Capacity

    NASA Astrophysics Data System (ADS)

    Woodworth-Jefcoats, P. A.; Polovina, J. J.; Drazen, J.

    2016-02-01

    We use output from a suite of CMIP5 earth system models to explore the impacts of climate change on marine fisheries over the 21st century. Ocean temperatures from both the historical and RCP 8.5 projections are integrated over the upper 200 m of the water column to characterize thermal habitat in the epipelagic realm. We find that across all models the projected temperature increases lead to a redistribution of thermal habitat: temperatures that currently represent the majority of North Pacific pelagic habitat are replaced by temperatures several degrees warmer. Additionally, all models project the emergence of new thermal habitat that exceeds present-day maximum temperatures. Spatially, present-day thermal habitat retreats northward and contracts eastward as warmer habitat in the southern and western North Pacific expands. In addition to these changes in thermal habitat, zooplankton densities are projected to decline across much of the North Pacific. Taken together, warming temperatures and declining zooplankton densities create the potential for mismatches in metabolic demand and supply through the 21st century. We find that carrying capacity for tropical tunas and other commercially valuable pelagic fish may be especially vulnerable to the impacts of climate change. The waters projected to see the greatest redistribution of thermal habitat and greatest declines in zooplankton densities are primarily those targeted by the Hawaii-based and international longline fleets. Fishery managers around the North Pacific will need to incorporate these impacts of climate change into future management strategies.

  4. The taphonomy of unmineralised Palaeozoic fossils preserved as siliciclastic moulds and casts, and their utility in assessing the interaction between environmental change and the fossil record

    NASA Astrophysics Data System (ADS)

    MacGabhann, Breandán; Schiffbauer, James; Hagadorn, James; Van Roy, Peter; Lynch, Edward; Morrsion, Liam; Murray, John

    2015-04-01

    The enhanced preservation potential of biomineralised tissues in fossil organisms is a key factor in their utility in the investigation of palaeoenvironmental change on fossil ecosystems. By contrast, the considerably lower preservation potential of entirely unmineralised organisms severely reduces the utility of their temporal and spatial distribution in such analyses. However, understanding the taphonomic processes which lead to the preservation of such soft-bodied fossils may be an under-appreciated source of information, particularly in the case of specimens preserved as moulds and casts in coarser siliciclastic sediments. This information potential is well demonstrated by fossil eldonids, a Cambrian to Devonian clade of unmineralised asymmetrical discoidal basal or stem deuterostomes, with an apparently conservative biology and no clear palaeoenvironmental or biogeographical controls on their distribution. We investigated the taphonomic processes involved in the preservation of fossil eldonids as moulds and casts on bedding surfaces and within event beds from sandstones of the Ordovician Tafilalt lagerstätte in south-eastern Morocco, and from siltstones of the Devonian West Falls Group of New York, USA. Laser Raman microspectroscopy, SEM BSE imaging and EDS elemental mapping of fossil specimens reveals that moulded biological surfaces are coated by a fossil surface veneer primarily consisting of mixed iron oxides and oxyhydroxides (including pseudomorphs after pyrite), and aluminosilicate clay minerals. Moreover, comparison to fossil eldonids preserved as carbonaceous compressions in the Burgess Shale reveals that the biological structures preserved in the Tafilalt and New York specimens - the dorsal surface and a coiled sac containing the digestive tract - represent only specific portions of the anatomy of the complete animal. We suggest that the preserved remains were the only parts of these eldonid organisms composed primarily of complex organic

  5. Fossil Microorganisms in Archaean

    NASA Technical Reports Server (NTRS)

    Astafleva, Marina; Hoover, Richard; Rozanov, Alexei; Vrevskiy, A.

    2006-01-01

    Ancient Archean and Proterozoic rocks are the model objects for investigation of rocks comprising astromaterials. The first of Archean fossil microorganisms from Baltic shield have been reported at the last SPIE Conference in 2005. Since this confeence biomorphic structures have been revealed in Archean rocks of Karelia. It was determined that there are 3 types of such bion structures: 1. structures found in situ, in other words microorganisms even-aged with rock matrix, that is real Archean fossils biomorphic structures, that is to say forms inhabited early formed rocks, and 3. younger than Archean-Protherozoic minerali microorganisms, that is later contamination. We made attempt to differentiate these 3 types of findings and tried to understand of burial of microorganisms. The structures belongs (from our point of view) to the first type, or real Archean, forms were under examination. Practical investigation of ancient microorganisms from Green-Stone-Belt of Northern Karelia turns to be very perspective. It shows that even in such ancient time as Archean ancient diverse world existed. Moreover probably such relatively highly organized cyanobacteria and perhaps eukaryotic formes existed in Archean world.

  6. Zeolites replacing plant fossils in the Denver formation, Lakewood, Colorado.

    USGS Publications Warehouse

    Modreski, P.J.; Verbeek, E.R.; Grout, M.A.

    1984-01-01

    Well-developed crystals of heulandite and stilbite, within fossil wood, occur in sedimentary rocks in Lakewood, Jefferson County. The rocks belong to the Denver formation, a locally fossiliferous deposit of fluvial claystone, siltstone, sandstone and conglomerate, containing some volcanic mudflows (andesitic) of late Cretaceous to Palaeocene age. Altered volcanic glass released Na and Ca into the ground-water and subsequently zeolites were crystallized in the open spaces between grains and within fossil plant structures. Minor pyrite, quartz (jasper), calcite and apatite also occur as replacements of fossil wood. Similar zeolite occurrences in other areas are reviewed.-R.S.M.

  7. Turbulence and fossil turbulence lead to life in the universe

    NASA Astrophysics Data System (ADS)

    Gibson, Carl H.

    2013-07-01

    Turbulence is defined as an eddy-like state of fluid motion where the inertial-vortex forces of the eddies are larger than all the other forces that tend to damp the eddies out. Fossil turbulence is a perturbation produced by turbulence that persists after the fluid ceases to be turbulent at the scale of the perturbation. Because vorticity is produced at small scales, turbulence must cascade from small scales to large, providing a consistent physical basis for Kolmogorovian universal similarity laws. Oceanic and astrophysical mixing and diffusion are dominated by fossil turbulence and fossil turbulent waves. Observations from space telescopes show turbulence and vorticity existed in the beginning of the universe and that their fossils persist. Fossils of big bang turbulence include spin and the dark matter of galaxies: clumps of ∼1012 frozen hydrogen planets that make globular star clusters as seen by infrared and microwave space telescopes. When the planets were hot gas, they hosted the formation of life in a cosmic soup of hot-water oceans as they merged to form the first stars and chemicals. Because spontaneous life formation according to the standard cosmological model is virtually impossible, the existence of life falsifies the standard cosmological model.

  8. Turbulence and Fossil Turbulence lead to Life in the Universe

    NASA Astrophysics Data System (ADS)

    Gibson, Carl H.

    2012-03-01

    Turbulence is defined as an eddy-like state of fluid motion where the inertial-vortex forces of the eddies are larger than all the other forces that tend to damp the eddies out. Fossil turbulence is a perturbation produced by turbulence that persists after the fluid ceases to be turbulent at the scale of the perturbation. Because vorticity is produced at small scales, turbulence must cascade from small scales to large, providing a consistent physical basis for Kolmogorovian universal similarity laws. Oceanic and astrophysical mixing and diffusion are dominated by fossil turbulence and fossil turbulent waves. Observations from space telescopes show turbulence and vorticity existed in the beginning of the universe and that their fossils persist. Fossils of big bang turbulence include spin and the dark matter of galaxies: clumps of ~ 1012 frozen hydrogen planets that make globular star clusters as seen by infrared and microwave space telescopes. When the planets were hot gas, they hosted the formation of life in a cosmic soup of hot- water oceans as they merged to form the first stars and chemicals. Because spontaneous life formation according to the standard cosmological model is virtually impossible, the existence of life falsifies the standard cosmological model.

  9. The oldest Mahonia (Berberidaceae) fossil from East Asia and its biogeographic implications.

    PubMed

    Huang, Jian; Su, Tao; Lebereton-Anberrée, Julie; Zhang, Shi-Tao; Zhou, Zhe-Kun

    2016-03-01

    Interpretation of the biogeography of the genus Mahonia (Berberidaceae) is limited by the lack of fossil records in East Asia. Compressed fossil foliage, described here as Mahonia mioasiatica sp. nov., were collected from the Upper Miocene Xiaolongtan Formation in Wenshan, Yunnan, southwest China. These specimens represent the oldest reliable fossil record of Mahonia in East Asia. This new fossil species shows a general similarity to Group Orientales and is most similar to the extant eastern Asian Mahonia conferta. Considering other fossil evidence of Mahonia, we propose a migration route of this genus to Asia over the North Atlantic Land Bridge rather than the Bering Land Bridge. Our results also suggest that North America, Europe and East Asia have been successive centers of diversity for the genus, as a consequence of diversification in Group Orientales potentially related to historical climate change.

  10. Habitat use and diel vertical migration of bigeye thresher shark: Overlap with pelagic longline fishing gear.

    PubMed

    Coelho, Rui; Fernandez-Carvalho, Joana; Santos, Miguel N

    2015-12-01

    Pelagic longliners targeting swordfish and tunas in oceanic waters regularly capture sharks as bycatch, including currently protected species as the bigeye thresher, Alopias superciliosus. Fifteen bigeye threshers were tagged with pop-up satellite archival tags (PSATs) in 2012-2014 in the tropical northeast Atlantic, with successful transmissions received from 12 tags for a total of 907 tracking days. Marked diel vertical movements were recorded on all specimens, with most of the daytime spent in deeper colder water (mean depth = 353 m, SD = 73; mean temperature = 10.7 °C, SD = 1.8) and nighttime spent in warmer water closer to the surface (mean depth = 72 m, SD = 54; mean temperature = 21.9 °C, SD = 3.7). The operating depth of the pelagic longline gear was measured with Minilog Temperature and Depth Recorders (TDRs), and the overlap with habitat utilization was calculated. Overlap is taking place mainly during the night and is higher for juveniles. The results presented herein can be used as inputs for Ecological Risk Assessments for bigeye threshers captured in oceanic tuna fisheries, and serve as a basis for efficient management and conservation of this vulnerable shark species. Copyright © 2015 Elsevier Ltd. All rights reserved.

  11. Oldest record of Metrosideros (Myrtaceae): Fossil flowers, fruits, and leaves from Australia.

    PubMed

    Tarran, Myall; Wilson, Peter G; Hill, Robert S

    2016-04-01

    Myrtaceous fossil capsular fruits and flowers from the northwest of Tasmania, in the Early Oligocene-aged Little Rapid River (LRR) deposit, are described. The reproductive organs are found in association with Myrtaceous leaves previously thought to belong to a fleshy-fruited genus, Xanthomyrtus at both LRR, and an Eocene Tasmanian site at Hasties, which are reassessed with fresh morphological evidence. Standard Light Microscopy (LM) and Scanning Electron Microscopy (SEM) were used to investigate cuticular characters and an auto-montage camera system was used to take high-resolution images of fossil and extant fruits. Fossils are identified using a nearest living relative (NLR) approach. The fossil fruits and flowers share a number of characters with genera of capsular-fruited Myrtaceae, in particular sharing several synapomorphies with species of Metrosideros subg. Metrosideros (tribe: Metrosidereae). The fossil is here described, and named Metrosideros leunigii, sp. nov. This research establishes the presence of Metrosideros (aff. subg. Metrosideros) in the Eocene-Oligocene (∼40-30 mya) of Tasmania, Australia. This is the first fossil record of Metrosideros in Australia, as well as the oldest conclusive fossil record, and may provide evidence for an Australian origin of the genus. It is also yet another example of extinction in the Tertiary of a group of plants on the Australian mainland that is only found today on nearby Pacific landmasses. © 2016 Botanical Society of America.

  12. Fossil resource and energy security dynamics in conventional and carbon-constrained worlds

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    McCollum, David; Bauer, Nico; Calvin, Katherine V.

    Fossil resource endowments and the future development of fossil fuel prices are important factors that will critically influence the nature and direction of the global energy system. In this paper we analyze a multi-model ensemble of long-term energy and emissions scenarios that were developed within the framework of the EMF27 integrated assessment model inter-comparison exercise. The diverse nature of these models highlights large uncertainties in the likely development of fossil resource (coal, oil, and natural gas) consumption, trade, and prices over the course of the twenty-first century and under different climate policy frameworks. We explore and explain some of themore » differences across scenarios and models and compare the scenario results with fossil resource estimates from the literature. A robust finding across the suite of IAMs is that the cumulative fossil fuel consumption foreseen by the models is well within the bounds of estimated recoverable reserves and resources. Hence, fossil resource constraints are, in and of themselves, unlikely to limit future GHG emissions. Our analysis also shows that climate mitigation policies could lead to a major reallocation of financial flows between regions, in terms of expenditures on fossil fuels and carbon, and can help to alleviate near-term energy security concerns via the reductions in oil imports and increases in energy system diversity they will help to motivate.« less

  13. Teacher Training and Authentic Scientific Research Utilizing Cretaceous Fossil Resources

    NASA Astrophysics Data System (ADS)

    Danch, J. M.

    2016-12-01

    The readily accessible Cretaceous fossil beds of central New Jersey provide an excellent opportunity for both teacher training in the utilization of paleontological resources in the classroom and authentic scientific student research at the middle and high school levels. Woodbridge Township New Jersey School District teachers participated in field trips to various fossiliferous sites to obtain photographic and video data and invertebrate and vertebrate fossil specimens for use in the classroom. Teachers were also presented with techniques allowing them to mentor students in performing authentic paleontological research. Students participated in multi-year research projects utilizing Cretaceous fossils collected in the field and presented their findings at science fairs and symposia. A workshop for K - 12 teachers statewide was developed for the New Jersey Science Convention providing information about New Jersey fossil resources and allowing participants to obtain, study and classify specimens. Additionally, the workshop provided participants with the information necessary for them to plan and conduct their own field trips.

  14. Impacts of Wind and Solar on Fossil-Fueled Generators: Preprint

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Lew, D.; Brinkman, G.; Kumar, N.

    2012-08-01

    High penetrations of wind and solar power will impact the operations of the remaining generators on the power system. Regional integration studies have shown that wind and solar may cause fossil-fueled generators to cycle on and off and ramp down to part load more frequently and potentially more rapidly. Increased cycling, deeper load following, and rapid ramping may result in wear-and-tear impacts on fossil-fueled generators that lead to increased capital and maintenance costs, increased equivalent forced outage rates, and degraded performance over time. Heat rates and emissions from fossil-fueled generators may be higher during cycling and ramping than during steady-statemore » operation. Many wind and solar integration studies have not taken these increased cost and emissions impacts into account because data have not been available. This analysis considers the cost and emissions impacts of cycling and ramping of fossil-fueled generation to refine assessments of wind and solar impacts on the power system.« less

  15. Freshwater Fossil Pearls from the Nihewan Basin, Early Early Pleistocene.

    PubMed

    Li, Su-Ping; Yao, Pei-Yi; Li, Jin-Feng; Ferguson, David Kay; Min, Long-Rui; Chi, Zhen-Qing; Wang, Yong; Yao, Jian-Xin; Sha, Jin-Geng

    2016-01-01

    Fossil blister pearls attached to the shells of an Anodonta mollusk from China, early Early Pleistocene, are reported here for the first time. The pearls were investigated in detail using a variety of methods. Micro-CT scanning of the fossil pearls was carried out to discover the inner structure and the pearl nucleus. Using CTAn software, changes in the gray levels of the biggest pearl, which reflect the changing density of the material, were investigated. The results provide us with some clues on how these pearls were formed. Sand grains, shell debris or material with a similar density could have stimulated the development of these pearls. X-ray diffraction analysis of one fossil pearl and the shell to which it was attached reveals that only aragonite exists in both samples. The internal structures of our fossil shells and pearls were investigated using a Scanning Electron Microscope. These investigations throw some light on pearl development in the past.

  16. From Fossil Parasitoids to Vectors: Insects as Parasites and Hosts.

    PubMed

    Nagler, Christina; Haug, Joachim T

    2015-01-01

    Within Metazoa, it has been proposed that as many as two-thirds of all species are parasitic. This propensity towards parasitism is also reflected within insects, where several lineages independently evolved a parasitic lifestyle. Parasitic behaviour ranges from parasitic habits in the strict sense, but also includes parasitoid, phoretic or kleptoparasitic behaviour. Numerous insects are also the host for other parasitic insects or metazoans. Insects can also serve as vectors for numerous metazoan, protistan, bacterial and viral diseases. The fossil record can report this behaviour with direct (parasite associated with its host) or indirect evidence (insect with parasitic larva, isolated parasitic insect, pathological changes of host). The high abundance of parasitism in the fossil record of insects can reveal important aspects of parasitic lifestyles in various evolutionary lineages. For a comprehensive view on fossil parasitic insects, we discuss here different aspects, including phylogenetic systematics, functional morphology and a direct comparison of fossil and extant species. Copyright © 2015 Elsevier Ltd. All rights reserved.

  17. Fossil organic carbon in wastewater and its fate in treatment plants.

    PubMed

    Law, Yingyu; Jacobsen, Geraldine E; Smith, Andrew M; Yuan, Zhiguo; Lant, Paul

    2013-09-15

    This study reports the presence of fossil organic carbon in wastewater and its fate in wastewater treatment plants. The findings pinpoint the inaccuracy of current greenhouse gas accounting guidelines which defines all organic carbon in wastewater to be of biogenic origin. Stable and radiocarbon isotopes ((13)C and (14)C) were measured throughout the process train in four municipal wastewater treatment plants equipped with secondary activated sludge treatment. Isotopic mass balance analyses indicate that 4-14% of influent total organic carbon (TOC) is of fossil origin with concentrations between 6 and 35 mg/L; 88-98% of this is removed from the wastewater. The TOC mass balance analysis suggests that 39-65% of the fossil organic carbon from the influent is incorporated into the activated sludge through adsorption or from cell assimilation while 29-50% is likely transformed to carbon dioxide (CO2) during secondary treatment. The fossil organic carbon fraction in the sludge undergoes further biodegradation during anaerobic digestion with a 12% decrease in mass. 1.4-6.3% of the influent TOC consists of both biogenic and fossil carbon is estimated to be emitted as fossil CO2 from activated sludge treatment alone. The results suggest that current greenhouse gas accounting guidelines, which assume that all CO2 emission from wastewater is biogenic may lead to underestimation of emissions. Copyright © 2013 Elsevier Ltd. All rights reserved.

  18. Economic value of U.S. fossil fuel electricity health impacts.

    PubMed

    Machol, Ben; Rizk, Sarah

    2013-02-01

    Fossil fuel energy has several externalities not accounted for in the retail price, including associated adverse human health impacts, future costs from climate change, and other environmental damages. Here, we quantify the economic value of health impacts associated with PM(2.5) and PM(2.5) precursors (NO(x) and SO(2)) on a per kilowatt hour basis. We provide figures based on state electricity profiles, national averages and fossil fuel type. We find that the economic value of improved human health associated with avoiding emissions from fossil fuel electricity in the United States ranges from a low of $0.005-$0.013/kWh in California to a high of $0.41-$1.01/kWh in Maryland. When accounting for the adverse health impacts of imported electricity, the California figure increases to $0.03-$0.07/kWh. Nationally, the average economic value of health impacts associated with fossil fuel usage is $0.14-$0.35/kWh. For coal, oil, and natural gas, respectively, associated economic values of health impacts are $0.19-$0.45/kWh, $0.08-$0.19/kWh, and $0.01-$0.02/kWh. For coal and oil, these costs are larger than the typical retail price of electricity, demonstrating the magnitude of the externality. When the economic value of health impacts resulting from air emissions is considered, our analysis suggests that on average, U.S. consumers of electricity should be willing to pay $0.24-$0.45/kWh for alternatives such as energy efficiency investments or emission-free renewable sources that avoid fossil fuel combustion. The economic value of health impacts is approximately an order of magnitude larger than estimates of the social cost of carbon for fossil fuel electricity. In total, we estimate that the economic value of health impacts from fossil fuel electricity in the United States is $361.7-886.5 billion annually, representing 2.5-6.0% of the national GDP. Published by Elsevier Ltd.

  19. First report of fossil "keratose" demosponges in Phanerozoic carbonates: preservation and 3-D reconstruction.

    PubMed

    Luo, Cui; Reitner, Joachim

    2014-06-01

    Fossil record of Phanerozoic non-spicular sponges, beside of being important with respect to the lineage evolution per se, could provide valuable references for the investigation of Precambrian ancestral animal fossils. However, although modern phylogenomic studies resolve non-spicular demosponges as the sister group of the remaining spiculate demosponges, the fossil record of the former is extremely sparse or unexplored compared to that of the latter; the Middle Cambrian Vauxiidae Walcott 1920, is the only confirmed fossil taxon of non-spicular demosponges. Here, we describe carbonate materials from Devonian (Upper Givetian to Lower Frasnian) bioherms of northern France and Triassic (Anisian) microbialites of Poland that most likely represent fossil remnants of keratose demosponges. These putative fossils of keratose demosponges are preserved as automicritic clumps. They are morphologically distinguishable from microbial fabrics but similar to other spiculate sponge fossils, except that the skeletal elements consist of fibrous networks instead of assembled spicules. Consistent with the immunological behavior of sponges, these fibrous skeletons often form a rim at the edge of the automicritic aggregate, separating the inner part of the aggregate from foreign objects. To confirm the architecture of these fibrous networks, two fossil specimens and a modern thorectid sponge for comparison were processed for three-dimensional (3-D) reconstruction using serial grinding tomography. The resulting fossil reconstructions are three-dimensionally anastomosing, like modern keratose demosponges, but their irregular and nonhierarchical meshes indicate a likely verongid affinity, although a precise taxonomic conclusion cannot be made based on the skeletal architecture alone. This study is a preliminary effort, but an important start to identify fossil non-spicular demosponges in carbonates and to re-evaluate their fossilization potential.

  20. Range contraction in large pelagic predators

    PubMed Central

    Worm, Boris; Tittensor, Derek P.

    2011-01-01

    Large reductions in the abundance of exploited land predators have led to significant range contractions for those species. This pattern can be formalized as the range–abundance relationship, a general macroecological pattern that has important implications for the conservation of threatened species. Here we ask whether similar responses may have occurred in highly mobile pelagic predators, specifically 13 species of tuna and billfish. We analyzed two multidecadal global data sets on the spatial distribution of catches and fishing effort targeting these species and compared these with available abundance time series from stock assessments. We calculated the effort needed to reliably detect the presence of a species and then computed observed range sizes in each decade from 1960 to 2000. Results suggest significant range contractions in 9 of the 13 species considered here (between 2% and 46% loss of observed range) and significant range expansions in two species (11–29% increase). Species that have undergone the largest declines in abundance and are of particular conservation concern tended to show the largest range contractions. These include all three species of bluefin tuna and several marlin species. In contrast, skipjack tuna, which may have increased its abundance in the Pacific, has also expanded its range size. These results mirror patterns described for many land predators, despite considerable differences in habitat, mobility, and dispersal, and imply ecological extirpation of heavily exploited species across parts of their range. PMID:21693644

  1. Seeking carotenoid pigments in amber-preserved fossil feathers.

    PubMed

    Thomas, Daniel B; Nascimbene, Paul C; Dove, Carla J; Grimaldi, David A; James, Helen F

    2014-06-09

    Plumage colours bestowed by carotenoid pigments can be important for visual communication and likely have a long evolutionary history within Aves. Discovering plumage carotenoids in fossil feathers could provide insight into the ecology of ancient birds and non-avian dinosaurs. With reference to a modern feather, we sought chemical evidence of carotenoids in six feathers preserved in amber (Miocene to mid-Cretaceous) and in a feather preserved as a compression fossil (Eocene). Evidence of melanin pigmentation and microstructure preservation was evaluated with scanning electron and light microscopies. We observed fine microstructural details including evidence for melanin pigmentation in the amber and compression fossils, but Raman spectral bands did not confirm the presence of carotenoids in them. Carotenoids may have been originally absent from these feathers or the pigments may have degraded during burial; the preservation of microstructure may suggest the former. Significantly, we show that carotenoid plumage pigments can be detected without sample destruction through an amber matrix using confocal Raman spectroscopy.

  2. Seeking carotenoid pigments in amber-preserved fossil feathers

    NASA Astrophysics Data System (ADS)

    Thomas, Daniel B.; Nascimbene, Paul C.; Dove, Carla J.; Grimaldi, David A.; James, Helen F.

    2014-06-01

    Plumage colours bestowed by carotenoid pigments can be important for visual communication and likely have a long evolutionary history within Aves. Discovering plumage carotenoids in fossil feathers could provide insight into the ecology of ancient birds and non-avian dinosaurs. With reference to a modern feather, we sought chemical evidence of carotenoids in six feathers preserved in amber (Miocene to mid-Cretaceous) and in a feather preserved as a compression fossil (Eocene). Evidence of melanin pigmentation and microstructure preservation was evaluated with scanning electron and light microscopies. We observed fine microstructural details including evidence for melanin pigmentation in the amber and compression fossils, but Raman spectral bands did not confirm the presence of carotenoids in them. Carotenoids may have been originally absent from these feathers or the pigments may have degraded during burial; the preservation of microstructure may suggest the former. Significantly, we show that carotenoid plumage pigments can be detected without sample destruction through an amber matrix using confocal Raman spectroscopy.

  3. Tool making, hand morphology and fossil hominins

    PubMed Central

    Marzke, Mary W.

    2013-01-01

    Was stone tool making a factor in the evolution of human hand morphology? Is it possible to find evidence in fossil hominin hands for this capability? These questions are being addressed with increasingly sophisticated studies that are testing two hypotheses; (i) that humans have unique patterns of grip and hand movement capabilities compatible with effective stone tool making and use of the tools and, if this is the case, (ii) that there exist unique patterns of morphology in human hands that are consistent with these capabilities. Comparative analyses of human stone tool behaviours and chimpanzee feeding behaviours have revealed a distinctive set of forceful pinch grips by humans that are effective in the control of stones by one hand during manufacture and use of the tools. Comparative dissections, kinematic analyses and biomechanical studies indicate that humans do have a unique pattern of muscle architecture and joint surface form and functions consistent with the derived capabilities. A major remaining challenge is to identify skeletal features that reflect the full morphological pattern, and therefore may serve as clues to fossil hominin manipulative capabilities. Hominin fossils are evaluated for evidence of patterns of derived human grip and stress-accommodation features. PMID:24101624

  4. Tool making, hand morphology and fossil hominins.

    PubMed

    Marzke, Mary W

    2013-11-19

    Was stone tool making a factor in the evolution of human hand morphology? Is it possible to find evidence in fossil hominin hands for this capability? These questions are being addressed with increasingly sophisticated studies that are testing two hypotheses; (i) that humans have unique patterns of grip and hand movement capabilities compatible with effective stone tool making and use of the tools and, if this is the case, (ii) that there exist unique patterns of morphology in human hands that are consistent with these capabilities. Comparative analyses of human stone tool behaviours and chimpanzee feeding behaviours have revealed a distinctive set of forceful pinch grips by humans that are effective in the control of stones by one hand during manufacture and use of the tools. Comparative dissections, kinematic analyses and biomechanical studies indicate that humans do have a unique pattern of muscle architecture and joint surface form and functions consistent with the derived capabilities. A major remaining challenge is to identify skeletal features that reflect the full morphological pattern, and therefore may serve as clues to fossil hominin manipulative capabilities. Hominin fossils are evaluated for evidence of patterns of derived human grip and stress-accommodation features.

  5. Cell differentiation and germ-soma separation in Ediacaran animal embryo-like fossils

    NASA Astrophysics Data System (ADS)

    Chen, Lei; Xiao, Shuhai; Pang, Ke; Zhou, Chuanming; Yuan, Xunlai

    2014-12-01

    Phosphorites of the Ediacaran Doushantuo Formation (~600 million years old) yield spheroidal microfossils with a palintomic cell cleavage pattern. These fossils have been variously interpreted as sulphur-oxidizing bacteria, unicellular protists, mesomycetozoean-like holozoans, green algae akin to Volvox, and blastula embryos of early metazoans or bilaterian animals. However, their complete life cycle is unknown and it is uncertain whether they had a cellularly differentiated ontogenetic stage, making it difficult to test their various phylogenetic interpretations. Here we describe new spheroidal fossils from black phosphorites of the Doushantuo Formation that have been overlooked in previous studies. These fossils represent later developmental stages of previously published blastula-like fossils, and they show evidence for cell differentiation, germ-soma separation, and programmed cell death. Their complex multicellularity is inconsistent with a phylogenetic affinity with bacteria, unicellular protists, or mesomycetozoean-like holozoans. Available evidence also indicates that the Doushantuo fossils are unlikely crown-group animals or volvocine green algae. We conclude that an affinity with cellularly differentiated multicellular eukaryotes, including stem-group animals or algae, is likely but more data are needed to constrain further the exact phylogenetic affinity of the Doushantuo fossils.

  6. Cell differentiation and germ-soma separation in Ediacaran animal embryo-like fossils.

    PubMed

    Chen, Lei; Xiao, Shuhai; Pang, Ke; Zhou, Chuanming; Yuan, Xunlai

    2014-12-11

    Phosphorites of the Ediacaran Doushantuo Formation (∼600 million years old) yield spheroidal microfossils with a palintomic cell cleavage pattern. These fossils have been variously interpreted as sulphur-oxidizing bacteria, unicellular protists, mesomycetozoean-like holozoans, green algae akin to Volvox, and blastula embryos of early metazoans or bilaterian animals. However, their complete life cycle is unknown and it is uncertain whether they had a cellularly differentiated ontogenetic stage, making it difficult to test their various phylogenetic interpretations. Here we describe new spheroidal fossils from black phosphorites of the Doushantuo Formation that have been overlooked in previous studies. These fossils represent later developmental stages of previously published blastula-like fossils, and they show evidence for cell differentiation, germ-soma separation, and programmed cell death. Their complex multicellularity is inconsistent with a phylogenetic affinity with bacteria, unicellular protists, or mesomycetozoean-like holozoans. Available evidence also indicates that the Doushantuo fossils are unlikely crown-group animals or volvocine green algae. We conclude that an affinity with cellularly differentiated multicellular eukaryotes, including stem-group animals or algae, is likely but more data are needed to constrain further the exact phylogenetic affinity of the Doushantuo fossils.

  7. Proceedings of the Fifteenth Annual Conference on Fossil Energy Materials

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Judkins, R.R.

    2002-02-12

    The Fifteenth Annual Conference on Fossil Energy Materials was held in Knoxville, Tennessee, on April 30 through May 2, 2001. The meeting was sponsored by the U.S. Department of Energy's (DOE) Office of Fossil Energy through the Advanced Research Materials Program (ARM). The objective of the ARM Program is to conduct research and development on materials for longer-term fossil energy applications, as well as for generic needs of various fossil fuel technologies. The management of the program has been decentralized to the DOE Oak Ridge Operations Office and Oak Ridge National Laboratory (ORNL). The research is performed by staff membersmore » at ORNL and by researchers at other national laboratories, universities, and in private industry. The work is divided into the following categories: (1) structural, ceramics, (2) new alloys and coatings, (3) functional materials, and (4) technology development and transfer. These proceedings were produced primarily from electronic files provided by the authors. They have been neither refereed nor extensively edited. However, most of the papers have already undergone technical review within the individual organizations before submission to the Program Office. The proceedings are available on the Fossil Energy home page at http://www.ornl.gov/fossil (Workshops and Conferences). The successful completion of the conference and publication of the proceedings has required help from several people. The organizers wish to thank Angela Beach of the ORNL Conference Office for her help in the many arrangements, and the numerous staff and support personnel associated with the conference. Finally, we express our sincere appreciation to the authors whose efforts are the very basis of the conference.« less

  8. The Gulf of Cádiz pelagic ecosystem: A review

    NASA Astrophysics Data System (ADS)

    García Lafuente, Jesús; Ruiz, Javier

    2007-08-01

    The Gulf of Cádiz, strategically situated between the North Atlantic Ocean and the Mediterranean Sea, has been the focus of attention of a few oceanographic studies dealing with the deep circulation in order to understand the dynamics of the dense plume of Mediterranean water. Much less attention has been paid to the surface pelagic layer which holds important living resources of commercial and ecological interest. This overview summarizes the recent advances that have been made concerning the regional oceanography of the northern half of this important basin from an interdisciplinary point of view. Probably the most relevant oceanographic feature of the basin is its strong seasonality, which is linked to the meteorologically-induced seasonality of the eastern boundary current system of the North Atlantic. The prominent cape Santa Maria divides the continental shelf off the southern Iberian Peninsula in two shelves of different shape that hold different oceanographic processes, which in turn determine the characteristics of the pelagic ecosystem. Mass and energy inputs from land as well as tidally-driven processes makes the wider eastern shelf be more productive while the narrower western shelf, cut by a sharp submarine canyon, is under the influence of the almost-permanent upwelling spot off cape San Vicente. Under easterlies, the west-going, warm coastal countercurrent that is observed in the eastern shelf may invade the western shelf thus connecting biologically both shelves in an east-to-west direction. Westerlies induce generalised upwelling off the southern Iberia Peninsula, which adds to the almost-permanent one off cape San Vicente and generates an upwelling jet that moves eastwards. Cape Santa Maria may deflect this flow by generating a cold filament that extends southward and diverts water from the western shelf to the open ocean. This pattern of circulation hampers the biological connection between shelves in the west-to-east direction, which is

  9. Preservation of three-dimensional anatomy in phosphatized fossil arthropods enriches evolutionary inference

    PubMed Central

    Schwermann, Achim H; dos Santos Rolo, Tomy; Caterino, Michael S; Bechly, Günter; Schmied, Heiko; Baumbach, Tilo; van de Kamp, Thomas

    2016-01-01

    External and internal morphological characters of extant and fossil organisms are crucial to establishing their systematic position, ecological role and evolutionary trends. The lack of internal characters and soft-tissue preservation in many arthropod fossils, however, impedes comprehensive phylogenetic analyses and species descriptions according to taxonomic standards for Recent organisms. We found well-preserved three-dimensional anatomy in mineralized arthropods from Paleogene fissure fillings and demonstrate the value of these fossils by utilizing digitally reconstructed anatomical structure of a hister beetle. The new anatomical data facilitate a refinement of the species diagnosis and allowed us to reject a previous hypothesis of close phylogenetic relationship to an extant congeneric species. Our findings suggest that mineralized fossils, even those of macroscopically poor preservation, constitute a rich but yet largely unexploited source of anatomical data for fossil arthropods. DOI: http://dx.doi.org/10.7554/eLife.12129.001 PMID:26854367

  10. Arthropod colonization of land--linking molecules and fossils in oribatid mites (Acari, Oribatida).

    PubMed

    Schaefer, Ina; Norton, Roy A; Scheu, Stefan; Maraun, Mark

    2010-10-01

    Terrestrial fossils that document the early colonization of land are scarce for >100 my after the Cambrian explosion. This raises the question whether life on land did not exist or just did not fossilize. With a molecular dating technique, we analyzed the origin of terrestrial chelicerate microarthropods (Acari, Oribatida) which have a fossil record since the Middle Devonian that is exceptional among soil animals. Our results suggest that oribatid mites originated in the Precambrian (571+/-37 mya) and that the radiation of basal groups coincides with the gap in the terrestrial fossil record between the Cambrian explosion and the earliest fossilized records of continental ecosystems. Further, they suggest that the colonization of land started via the interstitial, approximately 150 my earlier than the oldest fossils of terrestrial ecosystems. Overall, the results imply that omnivorous and detritivorous arthropods formed a major component in early terrestrial food webs, thereby facilitating the invasion of terrestrial habitats by later colonizers of higher trophic levels. Copyright 2010 Elsevier Inc. All rights reserved.

  11. On the Acceptability of Funding from Fossil Energy Companies

    NASA Astrophysics Data System (ADS)

    Frumhoff, P. C.; Goldman, G. T.

    2016-12-01

    Vigorous debates within the American Geophysical Union over the acceptability of funding from ExxonMobil sit within a broader societal debate over the climate responsibilities of fossil energy companies in a carbon constrained world. This has been fueled by recent evidence that leading companies have invested heavily in efforts to sow doubt about climate science in order to avoid regulation of their products. This talk examines the nature of this evidence and the ethical stances that AGU and other scientific societies and organizations might take toward acceptability of funding from fossil energy companies in light of it. The results of a climate responsibility scorecard, assessing a sample of leading oil, gas and coal companies against specific criteria for a more responsible fossil energy company, also will be presented.

  12. Role of non-fossil energy in meeting China's energy and climate target for 2020

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Zhou, Sheng; Tong, Qing; Yu, Sha

    2012-12-01

    China is the largest energy consumer and CO2 emitter in the world. The Chinese government faces growing challenges of ensuring energy security and reducing greenhouse gas emissions. To address these two issues, the Chinese government has announced two ambitious domestic indicative autonomous mitigation targets for 2020: increasing the ratio of non-fossil energy to 15% and reducing carbon dioxide emissions per unit of GDP by 40-45% from 2005 levels. To explore the role of non-fossil energy in achieving these two targets, this paper first provides an overview of current status of non-fossil energy development in China; then gives a brief reviewmore » of GDP and primary energy consumption; next assesses in detail the role of the non fossil energy in 2020, including the installed capacity and electricity generation of non-fossil energy sources, the share and role of non-fossil energy in the electricity structure, emissions reduction resulting from the shift to non-fossil energy, and challenges for accomplishing the mitigation targets in 2020 ; finally, conclusions and policy measures for non-fossil energy development are proposed.« less

  13. Investments in Fossil Energy Technology: How the Government's Fossil Energy R&D Program Has Made a Difference

    DOE R&D Accomplishments Database

    1997-03-01

    America has the technological capacity to change its energy future. There is no reason, for example, why our nation must continue following a path of rising oil imports when billions of barrels of crude oil remain in domestic oil fields. There is no reason why we cannot continue to use our abundant supplies of high-value, low-cost coal when we have the scientific know-how to remove virtually all of its pollutants and reduce greenhouse gas emissions. There is no reason why we cannot turn increasingly to clean-burning natural gas and tap the huge supplies we know exist within our borders. We remain a nation rich in the fuels that have powered economic growth. Today 85 percent of the energy we use to heat our homes and businesses, generate our electricity, and fuel our vehicles comes from coal, petroleum and natural gas. As we move toward a new century, the contributions of these fuels will grow. By 2015, the United States is likely to require nearly 20 percent more energy than it uses today, and fossil fuels are projected to supply almost 88 percent of the energy Americans will consume. We have the scientific know-how to continue using our fossil fuel wealth without fear of environmental damage or skyrocketing costs. The key is technology - developing cutting edge concepts that are beyond the private sector's current capabilities. Some of the most important innovations in America's energy industry are the results of investments in the Federal government's fossil energy research and development programs. Today, our air and water are cleaner, our economy is stronger, and our industries are more competitive in the global market because these programs have produced results. This booklet summarizes many of these achievements. It is not a comprehensive list by any means. Still, it provides solid evidence that the taxpayers' investment in government fossil energy research has paid real and measurable dividends.

  14. Rule Fossilization: A Tentative Model

    ERIC Educational Resources Information Center

    Vigil, Neddy A.; Oller, John W.

    1976-01-01

    A cybernetic model of factors involved in the fossilization of grammatical and lexical forms in learner grammars is offered. A distinction is made between affective and cognitive dimensions of a multidimensional channel of human communication; and the effect of expected and unexpected feedback on these two dimensions is discussed. (Author/POP)

  15. Leaf fossils of the ancient Tasmanian relict Microcachrys (Podocarpaceae) from New Zealand.

    PubMed

    Carpenter, Raymond J; Jordan, Gregory J; Mildenhall, Dallas C; Lee, Daphne E

    2011-07-01

    Microcachrys tetragona (Podocarpaceae), endemic to the mountains of Tasmania, represents the only remaining taxon of one of the world's most ancient and widely distributed conifer lineages. Remarkably, however, despite its ∼150 Myr heritage, our understanding of the fossil history of this lineage is based almost entirely on the pollen record. Fossils of Microcachrys are especially important in light of recent molecular phylogenetic and dating evidence. This evidence dates the Microcachrys lineage to the Mesozoic and does not support the traditional placement of Microcachrys as sister to the southeastern Australian genus Pherosphaera. We undertook comparative studies of the foliage architecture, cuticle, and paleoecology of newly discovered fossils from the Oligo-Miocene of New Zealand and M. tetragona and discussed the importance of Microcachrys in the context of Podocarpaceae phylogeny. The fossils represent the oldest and first extra-Australian macrofossils of Microcachrys and are described as the new foliage species M. novae-zelandiae. These fossils confirm that the distinctive opposite decussate phyllotaxy of the genus is at least as old as the Oligo-Miocene and contribute to evidence that Microcachrys plants were sometimes important components of oligotrophic swampy habitats. Leaf fossils of Microcachrys closely comparable with the only extant species confirm that this lineage had a much wider past distribution. The fossil record and recent molecular phylogenetic studies, including that of Microcachrys, also serve to emphasize the important status of Tasmania as a refugium for seed plant lineages.

  16. The shape of pterosaur evolution: evidence from the fossil record.

    PubMed

    Dyke, G J; McGowan, A J; Nudds, R L; Smith, D

    2009-04-01

    Although pterosaurs are a well-known lineage of Mesozoic flying reptiles, their fossil record and evolutionary dynamics have never been adequately quantified. On the basis of a comprehensive data set of fossil occurrences correlated with taxon-specific limb measurements, we show that the geological ages of pterosaur specimens closely approximate hypothesized patterns of phylogenetic divergence. Although the fossil record has expanded greatly in recent years, collectorship still approximates a sigmoid curve over time as many more specimens (and thus taxa) still remain undiscovered, yet our data suggest that the pterosaur fossil record is unbiased by sites of exceptional preservation (lagerstätte). This is because as new species are discovered the number of known formations and sites yielding pterosaur fossils has also increased - this would not be expected if the bulk of the record came from just a few exceptional faunas. Pterosaur morphological diversification is, however, strongly age biased: rarefaction analysis shows that peaks of diversity occur in the Late Jurassic and Early Cretaceous correlated with periods of increased limb disparity. In this respect, pterosaurs appear unique amongst flying vertebrates in that their disparity seems to have peaked relatively late in clade history. Comparative analyses also show that there is little evidence that the evolutionary diversification of pterosaurs was in any way constrained by the appearance and radiation of birds.

  17. Combining fossil and molecular data to date the diversification of New World Primates.

    PubMed

    Schrago, C G; Mello, B; Soares, A E R

    2013-11-01

    Recent methodological advances in molecular dating associated with the growing availability of sequence data have prompted the study of the evolution of New World Anthropoidea in recent years. Motivated by questions regarding historical biogeography or the mode of evolution, these works aimed to obtain a clearer scenario of Platyrrhini origins and diversification. Although some consensus was found, disputed issues, especially those relating to the evolutionary affinities of fossil taxa, remain. The use of fossil taxa for divergence time analysis is traditionally restricted to the provision of calibration priors. However, new analytical approaches have been developed that incorporate fossils as terminals and, thus, directly assign ages to the fossil tips. In this study, we conducted a combined analysis of molecular and morphological data, including fossils, to derive the timescale of New World anthropoids. Differently from previous studies that conducted total-evidence analysis of molecules and morphology, our approach investigated the morphological clock alone. Our results corroborate the hypothesis that living platyrrhines diversified in the last 20 Ma and that Miocene Patagonian fossils compose an independent evolutionary radiation that diversified in the late Oligocene. When compared to the node ages inferred from the molecular timescale, the inclusion of fossils augmented the precision of the estimates for nodes constrained by the fossil tips. We show that morphological data can be analysed using the same methodological framework applied in relaxed molecular clock studies. © 2013 The Authors. Journal of Evolutionary Biology © 2013 European Society For Evolutionary Biology.

  18. Children are likely to suffer most from our fossil fuel addiction.

    PubMed

    Perera, Frederica P

    2008-08-01

    The periods of fetal and child development arguably represent the stages of greatest vulnerability to the dual impacts of fossil fuel combustion: the multiple toxic effects of emitted pollutants (polycyclic aromatic hydrocarbons, particles, sulfur oxides, nitrogen oxides, metals) and the broad health impacts of global climate change attributable in large part to carbon dioxide released by fossil fuel burning. In this commentary I highlight current scientific evidence indicating that the fetus and young child are at heightened risk of developmental impairment, asthma, and cancer from fossil fuel pollutants and from the predicted effects of climate disruption such as heat waves, flooding, infectious disease, malnutrition, and trauma. Increased risk during early development derives from the inherently greater biologic vulnerability of the developing fetus and child and from their long future lifetime, during which early insults can potentially manifest as adult as well as childhood disease. I cite recent reports concluding that reducing dependence on fossil fuel and promoting clean and sustainable energy is economically feasible. Although much has been written separately about the toxicity of fossil fuel burning emissions and the effects of climate change on health, these two faces of the problem have not been viewed together with a focus on the developing fetus and child. Adolescence and old age are also periods of vulnerability, but the potential for both immediate and long-term adverse effects is greatest when exposure occurs prenatally or in the early years. Consideration of the full spectrum of health risks to children from fossil fuel combustion underscores the urgent need for environmental and energy policies to reduce fossil fuel dependence and maximize the health benefits to this susceptible population. We do not have to leave our children a double legacy of ill health and ecologic disaster.

  19. Children Are Likely to Suffer Most from Our Fossil Fuel Addiction

    PubMed Central

    Perera, Frederica P.

    2008-01-01

    Background The periods of fetal and child development arguably represent the stages of greatest vulnerability to the dual impacts of fossil fuel combustion: the multiple toxic effects of emitted pollutants (polycyclic aromatic hydrocarbons, particles, sulfur oxides, nitrogen oxides, metals) and the broad health impacts of global climate change attributable in large part to carbon dioxide released by fossil fuel burning. Objectives In this commentary I highlight current scientific evidence indicating that the fetus and young child are at heightened risk of developmental impairment, asthma, and cancer from fossil fuel pollutants and from the predicted effects of climate disruption such as heat waves, flooding, infectious disease, malnutrition, and trauma. Increased risk during early development derives from the inherently greater biologic vulnerability of the developing fetus and child and from their long future lifetime, during which early insults can potentially manifest as adult as well as childhood disease. I cite recent reports concluding that reducing dependence on fossil fuel and promoting clean and sustainable energy is economically feasible. Discussion Although much has been written separately about the toxicity of fossil fuel burning emissions and the effects of climate change on health, these two faces of the problem have not been viewed together with a focus on the developing fetus and child. Adolescence and old age are also periods of vulnerability, but the potential for both immediate and long-term adverse effects is greatest when exposure occurs prenatally or in the early years. Conclusions Consideration of the full spectrum of health risks to children from fossil fuel combustion underscores the urgent need for environmental and energy policies to reduce fossil fuel dependence and maximize the health benefits to this susceptible population. We do not have to leave our children a double legacy of ill health and ecologic disaster. PMID:18709169

  20. Ocean warming has a greater effect than acidification on the early life history development and swimming performance of a large circumglobal pelagic fish.

    PubMed

    Watson, Sue-Ann; Allan, Bridie J M; McQueen, David E; Nicol, Simon; Parsons, Darren M; Pether, Stephen M J; Pope, Stephen; Setiawan, Alvin N; Smith, Neville; Wilson, Carly; Munday, Philip L

    2018-05-22

    Ocean warming and acidification are serious threats to marine life; however, their individual and combined effects on large pelagic and predatory fishes are poorly understood. We determined the effects of projected future temperature and carbon dioxide (CO 2 ) levels on survival, growth, morphological development and swimming performance on the early life stages of a large circumglobal pelagic fish, the yellowtail kingfish Seriola lalandi. Eggs, larvae and juveniles were reared in cross-factored treatments of temperature (21 and 25°C) and pCO 2 (500 and 985 μatm) from fertilisation to 25 days post hatching (dph). Temperature had the greatest effect on survival, growth and development. Survivorship was lower, but growth and morphological development were faster at 25°C, with surviving fish larger and more developed at 1, 11 and 21 dph. Elevated pCO 2 affected size at 1 dph, but not at 11 or 21 dph, and did not affect survival or morphological development. Elevated temperature and pCO 2 had opposing effects on swimming performance at 21 dph. Critical swimming speed (U crit ) was increased by elevated temperature but reduced by elevated pCO 2 . Additionally, elevated temperature increased the proportion of individuals that responded to a startle stimulus, reduced latency to respond and increased maximum escape speed, potentially due to the more advanced developmental stage of juveniles at 25°C. By contrast, elevated pCO 2 reduced the distance moved and average speed in response to a startle stimulus. Our results show that higher temperature is likely to be the primary driver of global change impacts on kingfish early life history; however, elevated pCO 2 could affect critical aspects of swimming performance in this pelagic species. Our findings will help parameterise and structure fisheries population dynamics models and improve projections of impacts to large pelagic fishes under climate change scenarios to better inform adaptation and mitigation responses

  1. Experimental analysis of decay biases in the fossil record of lobopodians

    NASA Astrophysics Data System (ADS)

    Murdock, Duncan; Gabbott, Sarah; Purnell, Mark

    2016-04-01

    If fossils are to realize their full potential in reconstructing the tree of life we must understand how our view of ancient organisms is obscured by taphonomic filters of decay and preservation. In most cases, processes of decay will leave behind either nothing or only the most decay resistant body parts, and even in those rare instances where soft tissues are fossilized we cannot assume that the resulting fossil, however exquisite, represents a faithful anatomical representation of the animal as it was in life.Recent experiments have shown that the biases introduced by decay can be far from random; in chordates, for example, the most phylogenetically informative characters are also the most decay-prone, resulting in 'stemward slippage'. But how widespread is this phenomenon, and are there other non-random biases linked to decay? Intuitively, we make assumptions about the likelihood of different kinds of characters to survive and be preserved, with knock-on effects for anatomical and phylogenetic interpretations. To what extent are these assumptions valid? We combine our understanding of the fossil record of lobopodians with insights from decay experiments of modern onychophorans (velvet worms) to test these assumptions. Our analysis demonstrates that taphonomically informed tests of character interpretations have the potential to improve phylogenetic resolution. This approach is widely applicable to the fossil record - allowing us to ground-truth some of the assumptions involved in describing exceptionally preserved fossil material.

  2. Freshly excavated fossil bones are best for amplification of ancient DNA.

    PubMed

    Pruvost, Mélanie; Schwarz, Reinhard; Correia, Virginia Bessa; Champlot, Sophie; Braguier, Séverine; Morel, Nicolas; Fernandez-Jalvo, Yolanda; Grange, Thierry; Geigl, Eva-Maria

    2007-01-16

    Despite the enormous potential of analyses of ancient DNA for phylogeographic studies of past populations, the impact these analyses, most of which are performed with fossil samples from natural history museum collections, has been limited to some extent by the inefficient recovery of ancient genetic material. Here we show that the standard storage conditions and/or treatments of fossil bones in these collections can be detrimental to DNA survival. Using a quantitative paleogenetic analysis of 247 herbivore fossil bones up to 50,000 years old and originating from 60 different archeological and paleontological contexts, we demonstrate that freshly excavated and nontreated unwashed bones contain six times more DNA and yield twice as many authentic DNA sequences as bones treated with standard procedures. This effect was even more pronounced with bones from one Neolithic site, where only freshly excavated bones yielded results. Finally, we compared the DNA content in the fossil bones of one animal, a approximately 3,200-year-old aurochs, excavated in two separate seasons 57 years apart. Whereas the washed museum-stored fossil bones did not permit any DNA amplification, all recently excavated bones yielded authentic aurochs sequences. We established that during the 57 years when the aurochs bones were stored in a collection, at least as much amplifiable DNA was lost as during the previous 3,200 years of burial. This result calls for a revision of the postexcavation treatment of fossil bones to better preserve the genetic heritage of past life forms.

  3. SUZAKU OBSERVATIONS OF THE X-RAY BRIGHTEST FOSSIL GROUP ESO 3060170

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Su, Yuanyuan; White, Raymond E. III; Miller, Eric D., E-mail: ysu@crimson.ua.edu

    2013-10-01

    'Fossil' galaxy groups, each dominated by a relatively isolated giant elliptical galaxy, have many properties intermediate between groups and clusters of galaxies. We used the Suzaku X-ray observatory to observe the X-ray brightest fossil group, ESO 3060170, out to R{sub 200}, in order to better elucidate the relation between fossil groups, normal groups, and clusters. We determined the intragroup gas temperature, density, and metal abundance distributions and derived the entropy, pressure, and mass profiles for this group. The entropy and pressure profiles in the outer regions are flatter than in simulated clusters, similar to what is seen in observations ofmore » massive clusters. This may indicate that the gas is clumpy and/or the gas has been redistributed. Assuming hydrostatic equilibrium, the total mass is estimated to be ∼1.7 × 10{sup 14} M{sub ☉} within a radius R{sub 200} of ∼1.15 Mpc, with an enclosed baryon mass fraction of 0.13. The integrated iron mass-to-light ratio of this fossil group is larger than in most groups and comparable to those of clusters, indicating that this fossil group has retained the bulk of its metals. A galaxy luminosity density map on a scale of 25 Mpc shows that this fossil group resides in a relatively isolated environment, unlike the filamentary structures in which typical groups and clusters are embedded.« less

  4. Morphometric variation of extant platyrrhine molars: taxonomic implications for fossil platyrrhines

    PubMed Central

    Nova Delgado, Mónica; Galbany, Jordi

    2016-01-01

    The phylogenetic position of many fossil platyrrhines with respect to extant ones is not yet clear. Two main hypotheses have been proposed: the layered or successive radiations hypothesis suggests that Patagonian fossils are Middle Miocene stem platyrrhines lacking modern descendants, whereas the long lineage hypothesis argues for an evolutionary continuity of all fossil platyrrhines with the extant ones. Our geometric morphometric analysis of a 15 landmark-based configuration of platyrrhines’ first and second lower molars suggest that morphological stasis may explain the reduced molar shape variation observed. Platyrrhine lower molar shape might be a primitive retention of the ancestral state affected by strong ecological constraints throughout the radiation of the main platyrrhine families. The Patagonian fossil specimens showed two distinct morphological patterns of lower molars, Callicebus—like and Saguinus—like, which might be the precursors of the extant forms, whereas the Middle Miocene specimens, though showing morphological resemblances with the Patagonian fossils, also displayed new, derived molar patterns, Alouatta—like and Pitheciinae—like, thereby suggesting that despite the overall morphological stasis of molars, phenotypic diversification of molar shape was already settled during the Middle Miocene. PMID:27190704

  5. Unlocking the early fossil record of the arthropod central nervous system

    PubMed Central

    Edgecombe, Gregory D.; Ma, Xiaoya; Strausfeld, Nicholas J.

    2015-01-01

    Extant panarthropods (euarthropods, onychophorans and tardigrades) are hallmarked by stunning morphological and taxonomic diversity, but their central nervous systems (CNS) are relatively conserved. The timing of divergences of the ground pattern CNS organization of the major panarthropod clades has been poorly constrained because of a scarcity of data from their early fossil record. Although the CNS has been documented in three-dimensional detail in insects from Cenozoic ambers, it is widely assumed that these tissues are too prone to decay to withstand other styles of fossilization or geologically older preservation. However, Cambrian Burgess Shale-type compressions have emerged as sources of fossilized brains and nerve cords. CNS in these Cambrian fossils are preserved as carbon films or as iron oxides/hydroxides after pyrite in association with carbon. Experiments with carcasses compacted in fine-grained sediment depict preservation of neural tissue for a more prolonged temporal window than anticipated by decay experiments in other media. CNS and compound eye characters in exceptionally preserved Cambrian fossils predict divergences of the mandibulate and chelicerate ground patterns by Cambrian Stage 3 (ca 518 Ma), a dating that is compatible with molecular estimates for these splits. PMID:26554038

  6. Upward revision of global fossil fuel methane emissions based on isotope database.

    PubMed

    Schwietzke, Stefan; Sherwood, Owen A; Bruhwiler, Lori M P; Miller, John B; Etiope, Giuseppe; Dlugokencky, Edward J; Michel, Sylvia Englund; Arling, Victoria A; Vaughn, Bruce H; White, James W C; Tans, Pieter P

    2016-10-06

    Methane has the second-largest global radiative forcing impact of anthropogenic greenhouse gases after carbon dioxide, but our understanding of the global atmospheric methane budget is incomplete. The global fossil fuel industry (production and usage of natural gas, oil and coal) is thought to contribute 15 to 22 per cent of methane emissions to the total atmospheric methane budget. However, questions remain regarding methane emission trends as a result of fossil fuel industrial activity and the contribution to total methane emissions of sources from the fossil fuel industry and from natural geological seepage, which are often co-located. Here we re-evaluate the global methane budget and the contribution of the fossil fuel industry to methane emissions based on long-term global methane and methane carbon isotope records. We compile the largest isotopic methane source signature database so far, including fossil fuel, microbial and biomass-burning methane emission sources. We find that total fossil fuel methane emissions (fossil fuel industry plus natural geological seepage) are not increasing over time, but are 60 to 110 per cent greater than current estimates owing to large revisions in isotope source signatures. We show that this is consistent with the observed global latitudinal methane gradient. After accounting for natural geological methane seepage, we find that methane emissions from natural gas, oil and coal production and their usage are 20 to 60 per cent greater than inventories. Our findings imply a greater potential for the fossil fuel industry to mitigate anthropogenic climate forcing, but we also find that methane emissions from natural gas as a fraction of production have declined from approximately 8 per cent to approximately 2 per cent over the past three decades.

  7. Hydroxylamine hydrochloride-acetic acid-soluble and -insoluble fractions of pelagic sediment: Readsorption revisited

    USGS Publications Warehouse

    Piper, D.Z.; Wandless, G.A.

    1992-01-01

    The extraction of the rare earth elements (REE) from deep-ocean pelagic sediment, using hydroxylamine hydrochloride-acetic acid, leads to the separation of approximately 70% of the bulk REE content into the soluble fraction and 30% into the insoluble fraction. The REE pattern of the soluble fraction, i.e., the content of REE normalized to average shale on an element-by-element basis and plotted against atomic number, resembles the pattern for seawater, whereas the pattern, as well as the absolute concentrations, in the insoluble fraction resembles the North American shale composite. These results preclude significant readsorption of the REE by the insoluble phases during the leaching procedure.

  8. Implications of a fossil stickleback assemblage for Darwinian gradualism.

    PubMed

    Bell, M A

    2009-11-01

    Darwin postulated that a complete fossil record would contain numerous gradual transitions between ancestral and descendant species, but 150 years after publication of The Origin of Species, few such transitions have materialized. The fossil stickleback Gasterosteus doryssus and the deposit in which it occurs provide excellent conditions to detect such transitions. Abundant, well-preserved fossils occur in a stratigraphic setting with fine temporal resolution. The paleoecology of G. doryssus resembles the ecology of modern lakes that harbour the phenotypically similar three-spined stickleback Gasterosteus aculeatus. Gasterosteus aculeatus are primitively highly armoured, but G. doryssus comprised two contemporaneous biological species with relatively weak armour, including a near-shore, benthic feeder (benthic) and an offshore planktivore (limnetic). The benthic species expanded its range into the limnetic zone of the lake, where it apparently switched to planktivory and evolved reduced armour within c. 5000 years in response to directional selection. Although gradual evolution of mean phenotypes occurred, a single major gene caused much of evolutionary change of the pelvic skeleton. Thus, Darwin's expectation that transitions between species in the fossil record would be gradual was met at a fine time scale, but for pelvic structure, a well-studied trait, his expectation that gradual change would depend entirely on numerous, small, heritable differences among individuals was incorrect.

  9. D Central Line Extraction of Fossil Oyster Shells

    NASA Astrophysics Data System (ADS)

    Djuricic, A.; Puttonen, E.; Harzhauser, M.; Mandic, O.; Székely, B.; Pfeifer, N.

    2016-06-01

    Photogrammetry provides a powerful tool to digitally document protected, inaccessible, and rare fossils. This saves manpower in relation to current documentation practice and makes the fragile specimens more available for paleontological analysis and public education. In this study, high resolution orthophoto (0.5 mm) and digital surface models (1 mm) are used to define fossil boundaries that are then used as an input to automatically extract fossil length information via central lines. In general, central lines are widely used in geosciences as they ease observation, monitoring and evaluation of object dimensions. Here, the 3D central lines are used in a novel paleontological context to study fossilized oyster shells with photogrammetric and LiDAR-obtained 3D point cloud data. 3D central lines of 1121 Crassostrea gryphoides oysters of various shapes and sizes were computed in the study. Central line calculation included: i) Delaunay triangulation between the fossil shell boundary points and formation of the Voronoi diagram; ii) extraction of Voronoi vertices and construction of a connected graph tree from them; iii) reduction of the graph to the longest possible central line via Dijkstra's algorithm; iv) extension of longest central line to the shell boundary and smoothing by an adjustment of cubic spline curve; and v) integration of the central line into the corresponding 3D point cloud. The resulting longest path estimate for the 3D central line is a size parameter that can be applied in oyster shell age determination both in paleontological and biological applications. Our investigation evaluates ability and performance of the central line method to measure shell sizes accurately by comparing automatically extracted central lines with manually collected reference data used in paleontological analysis. Our results show that the automatically obtained central line length overestimated the manually collected reference by 1.5% in the test set, which is deemed

  10. Fossil Energy Planning for Navajo Nation

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Acedo, Margarita

    This project includes fossil energy transition planning to find optimal solutions that benefit the Navajo Nation and stakeholders. The majority of the tribe’s budget currently comes from fossil energy-revenue. The purpose of this work is to assess potential alternative energy resources including solar photovoltaics and biomass (microalgae for either biofuel or food consumption). This includes evaluating carbon-based reserves related to the tribe’s resources including CO 2 emissions for the Four Corners generating station. The methodology for this analysis will consist of data collection from publicly available data, utilizing expertise from national laboratories and academics, and evaluating economic, health, and environmentalmore » impacts. Finally, this report will highlight areas of opportunities to implement renewable energy in the Navajo Nation by presenting the technology requirements, cost, and considerations to energy, water, and environment in an educational structure.« less

  11. REE compositions in fossil vertebrate dental tissues indicate biomineral preservation

    NASA Astrophysics Data System (ADS)

    Žigaite, Ž.; Kear, B.; Pérez-Huerta, A.; Jeffries, T.; Blom, H.

    2012-04-01

    Rare earth element (REE) abundances have been measured in a number of Palaeozoic and Mesozoic dental tissues using Laser Ablation Inductively Coupled Plasma Mass-spectrometry (LA-ICP-MS). Fossil vertebrates analysed comprise scales and tesserae of Silurian and Devonian acanthodians, chondrichthyans, galeaspids, mongolepids, thelodonts, as well as teeth of Cretaceous lungfish and marine reptiles. The evaluation of fossil preservation level has been made by semi-quantitative spot geochemistry analyses on fine polished teeth and scale thin sections, using Energy Dispersive X-ray Spectroscopy (EDS). Fossil teeth and scales with significant structure and colour alteration have shown elevated heavy element concentrations, and the silicification of bioapatite has been common in their tissues. Stable oxygen isotope measurements (δ18O) of bulk biomineral have been conducted in parallel, and showed comparatively lower heavy oxygen values in the same fossil tissues with stronger visible alteration. Significant difference in REE concentrations has been observed between the dentine and enamel of Cretaceous plesiosaurs, suggesting the enamel to be more geochemically resistant to diagenetic overprint.

  12. Fossil slabs attached to unsubducted fragments of the Farallon plate.

    PubMed

    Wang, Yun; Forsyth, Donald W; Rau, Christina J; Carriero, Nina; Schmandt, Brandon; Gaherty, James B; Savage, Brian

    2013-04-02

    As the Pacific-Farallon spreading center approached North America, the Farallon plate fragmented into a number of small plates. Some of the microplate fragments ceased subducting before the spreading center reached the trench. Most tectonic models have assumed that the subducting oceanic slab detached from these microplates close to the trench, but recent seismic tomography studies have revealed a high-velocity anomaly beneath Baja California that appears to be a fossil slab still attached to the Guadalupe and Magdalena microplates. Here, using surface wave tomography, we establish the lateral extent of this fossil slab and show that it is correlated with the distribution of high-Mg andesites thought to derive from partial melting of the subducted oceanic crust. We also reinterpret the high seismic velocity anomaly beneath the southern central valley of California as another fossil slab extending to a depth of 200 km or more that is attached to the former Monterey microplate. The existence of these fossil slabs may force a reexamination of models of the tectonic evolution of western North America over the last 30 My.

  13. Oxygen: A Fundamental Property Regulating Pelagic Ecosystem Structure in the Coastal Southeastern Tropical Pacific

    PubMed Central

    Bertrand, Arnaud; Chaigneau, Alexis; Peraltilla, Salvador; Ledesma, Jesus; Graco, Michelle; Monetti, Florian; Chavez, Francisco P.

    2011-01-01

    Background In the southeastern tropical Pacific anchovy (Engraulis ringens) and sardine (Sardinops sagax) abundance have recently fluctuated on multidecadal scales and food and temperature have been proposed as the key parameters explaining these changes. However, ecological and paleoecological studies, and the fact that anchovies and sardines are favored differently in other regions, raise questions about the role of temperature. Here we investigate the role of oxygen in structuring fish populations in the Peruvian upwelling ecosystem that has evolved over anoxic conditions and is one of the world's most productive ecosystems in terms of forage fish. This study is particularly relevant given that the distribution of oxygen in the ocean is changing with uncertain consequences. Methodology/Principal Findings A comprehensive data set is used to show how oxygen concentration and oxycline depth affect the abundance and distribution of pelagic fish. We show that the effects of oxygen on anchovy and sardine are opposite. Anchovy flourishes under relatively low oxygen conditions while sardine avoid periods/areas with low oxygen concentration and restricted habitat. Oxygen consumption, trophic structure and habitat compression play a fundamental role in fish dynamics in this important ecosystem. Conclusions/Significance For the ocean off Peru we suggest that a key process, the need to breathe, has been neglected previously. Inclusion of this missing piece allows the development of a comprehensive conceptual model of pelagic fish populations and change in an ocean ecosystem impacted by low oxygen. Should current trends in oxygen in the ocean continue similar effects may be evident in other coastal upwelling ecosystems. PMID:22216315

  14. Oxygen: a fundamental property regulating pelagic ecosystem structure in the coastal southeastern tropical Pacific.

    PubMed

    Bertrand, Arnaud; Chaigneau, Alexis; Peraltilla, Salvador; Ledesma, Jesus; Graco, Michelle; Monetti, Florian; Chavez, Francisco P

    2011-01-01

    In the southeastern tropical Pacific anchovy (Engraulis ringens) and sardine (Sardinops sagax) abundance have recently fluctuated on multidecadal scales and food and temperature have been proposed as the key parameters explaining these changes. However, ecological and paleoecological studies, and the fact that anchovies and sardines are favored differently in other regions, raise questions about the role of temperature. Here we investigate the role of oxygen in structuring fish populations in the Peruvian upwelling ecosystem that has evolved over anoxic conditions and is one of the world's most productive ecosystems in terms of forage fish. This study is particularly relevant given that the distribution of oxygen in the ocean is changing with uncertain consequences. A comprehensive data set is used to show how oxygen concentration and oxycline depth affect the abundance and distribution of pelagic fish. We show that the effects of oxygen on anchovy and sardine are opposite. Anchovy flourishes under relatively low oxygen conditions while sardine avoid periods/areas with low oxygen concentration and restricted habitat. Oxygen consumption, trophic structure and habitat compression play a fundamental role in fish dynamics in this important ecosystem. For the ocean off Peru we suggest that a key process, the need to breathe, has been neglected previously. Inclusion of this missing piece allows the development of a comprehensive conceptual model of pelagic fish populations and change in an ocean ecosystem impacted by low oxygen. Should current trends in oxygen in the ocean continue similar effects may be evident in other coastal upwelling ecosystems. © 2011 Bertrand et al.

  15. Trace fossils of microbial colonization on Mars: Criteria for search and for sample return

    NASA Technical Reports Server (NTRS)

    Friedmann, E. I.

    1988-01-01

    The recent discovery of microbial trace-fossil formation in the frigid Ross Desert of Antarctica suggests that early primitive life on Mars may have left behind similar signatures. These trace fossils are apparent as chemical or physical changes in rock (or sediment) structure (or chemistry) caused by the activity of organisms. Life on Mars, if it ever existed, almost certainly did not evolve above the level of microorganisms, and this should be considered in search for fossil life. For the reasons detailed here, microbial trace fossils seem to be a better and more realistic target for search than would be true microbial fossils (remnants of cellular structures).

  16. Transitional fossils and the origin of turtles

    PubMed Central

    Lyson, Tyler R.; Bever, Gabe S.; Bhullar, Bhart-Anjan S.; Joyce, Walter G.; Gauthier, Jacques A.

    2010-01-01

    The origin of turtles is one of the most contentious issues in systematics with three currently viable hypotheses: turtles as the extant sister to (i) the crocodile–bird clade, (ii) the lizard–tuatara clade, or (iii) Diapsida (a clade composed of (i) and (ii)). We reanalysed a recent dataset that allied turtles with the lizard–tuatara clade and found that the inclusion of the stem turtle Proganochelys quenstedti and the ‘parareptile’ Eunotosaurus africanus results in a single overriding morphological signal, with turtles outside Diapsida. This result reflects the importance of transitional fossils when long branches separate crown clades, and highlights unexplored issues such as the role of topological congruence when using fossils to calibrate molecular clocks. PMID:20534602

  17. A comprehensive database of quality-rated fossil ages for Sahul’s Quaternary vertebrates

    PubMed Central

    Rodríguez-Rey, Marta; Herrando-Pérez, Salvador; Brook, Barry W.; Saltré, Frédérik; Alroy, John; Beeton, Nicholas; Bird, Michael I.; Cooper, Alan; Gillespie, Richard; Jacobs, Zenobia; Johnson, Christopher N.; Miller, Gifford H.; Prideaux, Gavin J.; Roberts, Richard G.; Turney, Chris S.M.; Bradshaw, Corey J.A.

    2016-01-01

    The study of palaeo-chronologies using fossil data provides evidence for past ecological and evolutionary processes, and is therefore useful for predicting patterns and impacts of future environmental change. However, the robustness of inferences made from fossil ages relies heavily on both the quantity and quality of available data. We compiled Quaternary non-human vertebrate fossil ages from Sahul published up to 2013. This, the FosSahul database, includes 9,302 fossil records from 363 deposits, for a total of 478 species within 215 genera, of which 27 are from extinct and extant megafaunal species (2,559 records). We also provide a rating of reliability of individual absolute age based on the dating protocols and association between the dated materials and the fossil remains. Our proposed rating system identified 2,422 records with high-quality ages (i.e., a reduction of 74%). There are many applications of the database, including disentangling the confounding influences of hypothetical extinction drivers, better spatial distribution estimates of species relative to palaeo-climates, and potentially identifying new areas for fossil discovery. PMID:27434208

  18. Trophic transfer of persistent organic pollutants through a pelagic food web: The case of Lake Como (Northern Italy).

    PubMed

    Mazzoni, Michela; Boggio, Emanuela; Manca, Marina; Piscia, Roberta; Quadroni, Silvia; Bellasi, Arianna; Bettinetti, Roberta

    2018-05-30

    Despite DDT and PCB having been banned for about 40 years, they are still detectable in the environment. In the present research we specifically investigated the trophic transfer of these organochlorine contaminants (OC) through a pelagic food web of a deep lake in Northern Italy (Lake Como) over time. Zooplankton and fish were sampled each season of a year and OC concentrations and the carbon and nitrogen isotopic ratios were measured. By using stable isotopes, the direct trophic relationship between pelagic zooplankton and zooplanktivorous fish was confirmed for Alosa agone only in summer. Based on this result, the biomagnification factor normalized on the trophic level (BMF TL ) for organic contaminants was calculated. BMF TL values were within the range 0.9-1.9 for DDT isomers and 1.6-4.9 for some PCB congeners (PCB 95, PCB 101, PCB 149, PCB 153, PCB 138 - present both in zooplankton and in fish and representing >60% of the PCB contamination), confirming the biomagnification of these compounds in one of the two zooplanktivorous fish species of the lake. Copyright © 2018. Published by Elsevier B.V.

  19. New Mesozoic and Cenozoic fossils from Ecuador: Invertebrates, vertebrates, plants, and microfossils

    NASA Astrophysics Data System (ADS)

    Cadena, Edwin A.; Mejia-Molina, Alejandra; Brito, Carla M.; Peñafiel, Sofia; Sanmartin, Kleber J.; Sarmiento, Luis B.

    2018-04-01

    Ecuador is well known for its extensive extant biodiversity, however, its paleobiodiversity is still poorly explored. Here we report seven new Mesozoic and Cenozoic fossil localities from the Pacific coast, inter-Andean depression and Napo basin of Ecuador, including vertebrates, invertebrates, plants, and microfossils. The first of these localities is called El Refugio, located near the small town of Chota, Imbabura Province, from where we report several morphotypes of fossil leaves and a mycetopodid freshwater mussel of the Upper Miocene Chota Formation. A second site is also located near the town of Chota, corresponding to potentially Pleistocene to Holocene lake deposits from which we report the occurrence of leaves and fossil diatoms. A third locality is at the Pacific coast of the country, near Rocafuerte, a town in Esmeraldas Province, from which we report a late Miocene palm leaf. We also report the first partially articulated skull with teeth from a Miocene scombridid (Mackerels) fish from El Cruce locality, and completely preserved seeds from La Pila locality, both sites from Manabí Province. Two late Cretaceous fossil sites from the Napo Province, one near Puerto Napo showing a good record of fossil shrimps and a second near the town of Loreto shows the occurrence of granular amber and small gymnosperms seeds and cuticles. All these new sites and fossils show the high potential of the sedimentary sequences and basins of Ecuador for paleontological studies and for a better understanding of the fossil record of the country and northern South America.

  20. 76 FR 3517 - Standards of Performance for Fossil-Fuel-Fired, Electric Utility, Industrial-Commercial...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-01-20

    ... Standards of Performance for Fossil-Fuel-Fired, Electric Utility, Industrial-Commercial-Institutional, and... following: Category NAICS \\1\\ Examples of regulated entities Industry 221112 Fossil fuel-fired electric utility steam generating units. Federal Government 22112 Fossil fuel-fired electric utility steam...