Sample records for population density average

  1. The Effect of Inhibitory Neuron on the Evolution Model of Higher-Order Coupling Neural Oscillator Population

    PubMed Central

    Qi, Yi; Wang, Rubin; Jiao, Xianfa; Du, Ying

    2014-01-01

    We proposed a higher-order coupling neural network model including the inhibitory neurons and examined the dynamical evolution of average number density and phase-neural coding under the spontaneous activity and external stimulating condition. The results indicated that increase of inhibitory coupling strength will cause decrease of average number density, whereas increase of excitatory coupling strength will cause increase of stable amplitude of average number density. Whether the neural oscillator population is able to enter the new synchronous oscillation or not is determined by excitatory and inhibitory coupling strength. In the presence of external stimulation, the evolution of the average number density is dependent upon the external stimulation and the coupling term in which the dominator will determine the final evolution. PMID:24516505

  2. 77 FR 4014 - Takes of Marine Mammals Incidental to Specified Activities; Physical Oceanographic Studies in the...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2012-01-26

    ... chambers for a total discharge volume of 210 in\\3\\) with a 1,200 m long hydrophone streamer. GI guns will... require the Navy to use species-specific mean maximum densities, rather than the mean average densities... use mean maximum densities, rather than mean average densities. Marine mammal population density...

  3. [Population change of farmland rodent and the influences of climate and cultivation factors in Fengxian District of Shanghai, China].

    PubMed

    Fei, Lei; Wang, Zu-guo; Yao, Yao; Xu, Xiang-ming; Gu, Pin-qiang

    2015-02-01

    We analyzed the changes of rodent population from 1988 to 2013 in Fengxian District, Shanghai, and discussed the relevance of rodent population dynamics and climate, and ecological factors. Results showed that the average rat density was 1.3% from 1988 to 2013, and the dominant species was Apodemus agrarius. There was little change of density between years. The average composition ratio was 97.5%. The peak of rodent density appeared from February to May, highest in March. Average density and peak decreased obviously year after year. Numbers of adult rodents of 1st group were positively correlated, while that of adult rodents of 2nd group and elderly group negatively correlated with years. Rat age presented a shortened trend annually. Rodent density and temperature factors were negatively correlated except in January. Extreme warmth in February had a lagged, positive effect on the rat density. Precipitation factors had no significant correlation with the density of rodents. Relative humidity was positively correlated with the density, mostly. Planting structure adjustment and reduction of planting acreage were significantly correlated with the reduction of rodent density. Increased deratization rate was also correlated with the reduction.

  4. Metapopulation extinction risk: dispersal's duplicity.

    PubMed

    Higgins, Kevin

    2009-09-01

    Metapopulation extinction risk is the probability that all local populations are simultaneously extinct during a fixed time frame. Dispersal may reduce a metapopulation's extinction risk by raising its average per-capita growth rate. By contrast, dispersal may raise a metapopulation's extinction risk by reducing its average population density. Which effect prevails is controlled by habitat fragmentation. Dispersal in mildly fragmented habitat reduces a metapopulation's extinction risk by raising its average per-capita growth rate without causing any appreciable drop in its average population density. By contrast, dispersal in severely fragmented habitat raises a metapopulation's extinction risk because the rise in its average per-capita growth rate is more than offset by the decline in its average population density. The metapopulation model used here shows several other interesting phenomena. Dispersal in sufficiently fragmented habitat reduces a metapopulation's extinction risk to that of a constant environment. Dispersal between habitat fragments reduces a metapopulation's extinction risk insofar as local environments are asynchronous. Grouped dispersal raises the effective habitat fragmentation level. Dispersal search barriers raise metapopulation extinction risk. Nonuniform dispersal may reduce the effective fraction of suitable habitat fragments below the extinction threshold. Nonuniform dispersal may make demographic stochasticity a more potent metapopulation extinction force than environmental stochasticity.

  5. Estimating the population size and colony boundary of subterranean termites by using the density functions of directionally averaged capture probability.

    PubMed

    Su, Nan-Yao; Lee, Sang-Hee

    2008-04-01

    Marked termites were released in a linear-connected foraging arena, and the spatial heterogeneity of their capture probabilities was averaged for both directions at distance r from release point to obtain a symmetrical distribution, from which the density function of directionally averaged capture probability P(x) was derived. We hypothesized that as marked termites move into the population and given sufficient time, the directionally averaged capture probability may reach an equilibrium P(e) over the distance r and thus satisfy the equal mixing assumption of the mark-recapture protocol. The equilibrium capture probability P(e) was used to estimate the population size N. The hypothesis was tested in a 50-m extended foraging arena to simulate the distance factor of field colonies of subterranean termites. Over the 42-d test period, the density functions of directionally averaged capture probability P(x) exhibited four phases: exponential decline phase, linear decline phase, equilibrium phase, and postequilibrium phase. The equilibrium capture probability P(e), derived as the intercept of the linear regression during the equilibrium phase, correctly projected N estimates that were not significantly different from the known number of workers in the arena. Because the area beneath the probability density function is a constant (50% in this study), preequilibrium regression parameters and P(e) were used to estimate the population boundary distance 1, which is the distance between the release point and the boundary beyond which the population is absent.

  6. Photographic Identification of Humpback and Blue Whales off the US West Coast: Results and Updated Abundance Estimates from 2008 Field Season

    DTIC Science & Technology

    2009-12-01

    1990s and only being present part of the time off the US West Coast lowering the average density present (but not reflecting an actual population ...the average density present (but not reflecting an actual population decline). 15. SUBJECT TERMS 16. SECURITY CLASSIFICATION OF: 17. LIMITATION OF...due to our sample representing a decreasing proportion of this growing population leading to greater variation and possibly greater susceptibility to

  7. [Population density, age distribution and urbanisation as factors influencing the frequency of home visits--an analysis for Mecklenburg-West Pomerania].

    PubMed

    Heymann, R; Weitmann, K; Weiss, S; Thierfelder, D; Flessa, S; Hoffmann, W

    2009-07-01

    This study examines and compares the frequency of home visits by general practitioners in regions with a lower population density and regions with a higher population density. The discussion centres on the hypothesis whether the number of home visits in rural and remote areas with a low population density is, in fact, higher than in urbanised areas with a higher population density. The average age of the population has been considered in both cases. The communities of Mecklenburg West-Pomerania were aggregated into postal code regions. The analysis is based on these postal code regions. The average frequency of home visits per 100 inhabitants/km2 has been calculated via a bivariate, linear regression model with the population density and the average age for the postal code region as independent variables. The results are based on billing data of the year 2006 as provided by the Association of Statutory Health Insurance Physicians of Mecklenburg-Western Pomerania. In a second step a variable which clustered the postal codes of urbanised areas was added to a multivariate model. The hypothesis of a negative correlation between the frequency of home visits and the population density of the areas examined cannot be confirmed for Mecklenburg-Western Pomerania. Following the dichotomisation of the postal code regions into sparsely and densely populated areas, only the very sparsely populated postal code regions (less than 100 inhabitants/km2) show a tendency towards a higher frequency of home visits. Overall, the frequency of home visits in sparsely populated postal code regions is 28.9% higher than in the densely populated postal code regions (more than 100 inhabitants/km2), although the number of general practitioners is approximately the same in both groups. In part this association seems to be confirmed by a positive correlation between the average age in the individual postal code regions and the number of home visits carried out in the area. As calculated on the basis of the data at hand, only the very sparsely populated areas with a still gradually decreasing population show a tendency towards a higher frequency of home visits. According to the data of 2006, the number of home visits remains high in sparsely populated areas. It may increase in the near future as the number of general practitioners in these areas will gradually decrease while the number of immobile and older inhabitants will increase.

  8. [Natural forming causes of China population distribution].

    PubMed

    Fang, Yu; Ouyang, Zhi-Yun; Zheng, Hua; Xiao, Yi; Niu, Jun-Feng; Chen, Sheng-Bin; Lu, Fei

    2012-12-01

    The diverse natural environment in China causes the spatial heterogeneity of China population distribution. It is essential to understand the interrelations between the population distribution pattern and natural environment to enhance the understanding of the man-land relationship and the realization of the sustainable management for the population, resources, and environment. This paper analyzed the China population distribution by adopting the index of population density (PD) in combining with spatial statistic method and Lorenz curve, and discussed the effects of the natural factors on the population distribution and the interrelations between the population distribution and 16 indices including average annual precipitation (AAP), average annual temperature (AAT), average annual sunshine duration (AASD), precipitation variation (PV), temperature variation (TV), sunshine duration variation (SDV), relative humidity (RH), aridity index (AI), warmth index ( WI), > or = 5 degrees C annual accumulated temperature (AACT), average elevation (AE), relative height difference (RHD), surface roughness (SR), water system density (WSD), net primary productivity (NPP), and shortest distance to seashore (SDTS). There existed an obvious aggregation phenomenon in the population distribution in China. The PD was high in east China, medium in central China, and low in west China, presenting an obvious positive spatial association. The PD was significantly positively correlated with WSD, AAT, AAP, NPP, AACT, PV, RH, and WI, and significantly negatively correlated with RHD, AE, SDV, SR, and SDTS. The climate factors (AAT, WI, PV, and NPP), topography factors (SR and RHD), and water system factor (WSD) together determined the basic pattern of the population distribution in China. It was suggested that the monitoring of the eco-environment in the east China of high population density should be strengthened to avoid the eco-environmental degradation due to the expanding population, and the conservation of the eco-environment in the central and west China with vulnerable eco-environment should also be strengthened to enhance the population carrying ability of these regions and to mitigate the eco-environmental pressure in the east China of high population density.

  9. Area characteristics and admission rates of people with schizophrenia and affective disorders in a German rural catchment area.

    PubMed

    Losert, C; Schmauß, M; Becker, T; Kilian, R

    2012-12-01

    Studies in urban areas identified environmental risk factors for mental illness, but little research on this topic has been performed in rural areas. Hospital admission rates were computed for 174 rural municipalities in the catchment area of the state psychiatric hospital in Günzburg in years 2006 to 2009 and combined with structural and socio-economic data. Relationships of overall and diagnosis-specific admission rates with municipality characteristics were analysed by means of negative binomial regression models. Admission rates of patients with a diagnosis of schizophrenia and affective disorder combined decrease with increasing population growth, population density, average income and green areas, while admission rates are positively correlated with commuter balance, income inequality, unemployment rates and traffic areas. Admission rates for schizophrenia are negatively related to population growth, average income and agricultural areas, but positively related to mobility index, income inequality and unemployment rate. Admission rates for affective disorders are negatively related to population growth, population density, average income and green areas, while higher admission rates are correlated with commuter balance, high income inequality, unemployment rate and traffic-related areas. Effects of wealth, economic inequality, population density and structural area characteristics influence psychiatric admission rates also in rural areas.

  10. Correlations between U.S. county annual cancer incidence and population density.

    PubMed

    Vares, David Ae; St-Pierre, Linda S; Persinger, Michael A

    2015-01-01

    Population density implicitly involves specific distances between living individuals who exhibit biophysical forces and energies. Objective was to investigate major data bases of cancer incidence and population data to help understand the emergent properties of diseases that become apparent only when large populations and areas are considered. Correlation analyses of the annual incidence (years 2007 to 2011) of cancer in counties (2,885) of the U.S. and population densities were convergent with these quantitative predictions and suggested an inflection threshold around 50 people per square mile. The potential role of subtle or even "non-local" factors coupled to averaged population density in the viability and mortality of the human species may serve as alternative explanations to the attribution of malignancy to "chance" factors. Calculations indicated average distances between the electric force dipole of the brains or bodies of human beings generate forces known to affect DNA extension and when distributed over the Compton wavelength of the electron could produce energies sufficient to affect the binding of base nucleotides. An inclusive science of human ecology might benefit from considering subtle forces and energies associated with the individual members within the habitat that could determine the probability of cellular anomalies.

  11. Peromyscus ranges at high and low population densities

    USGS Publications Warehouse

    Stickel, L.F.

    1960-01-01

    Live-trapping studies at the Patuxent Wildlife Research Center, Maryland, showed that the ranges of wood mice were larger when the population density was lower and smaller when the population density was higher. When the population density was about 1.3 male mice per acre in June 1954, the average distance recorded between traps after four or more captures was 258 feet. When the population density was about 4.1 male mice per acre in June 1957, the average distance was 119 feet. Differences were statistically significant. Females were so scarce at the low that comparisons could not be made for them. Examples from the literature also show that home range of a species may vary with population density. Other examples show that the range may vary with habitat, breeding condition and food supply. These variations in range size reduce the reliability of censuses in which relative methods are used: Lines of traps sample the population of a larger area when ranges are large than they do when ranges are small. Direct comparisons therefore will err in some degree. Error may be introduced also when line-trap data are transformed to per acre figures on the basis of home-range estimates made by area-trapping at another place or time. Variation in range size also can make it necessary to change area-trapping plans, for larger quadrants are needed when ranges are larger. It my be necessary to set traps closer together when ranges are small than when ranges are large.

  12. Exposure time of oral rabies vaccine baits relative to baiting density and raccoon population density.

    PubMed

    Blackwell, Bradley F; Seamans, Thomas W; White, Randolph J; Patton, Zachary J; Bush, Rachel M; Cepek, Jonathan D

    2004-04-01

    Oral rabies vaccination (ORV) baiting programs for control of raccoon (Procyon lotor) rabies in the USA have been conducted or are in progress in eight states east of the Mississippi River. However, data specific to the relationship between raccoon population density and the minimum density of baits necessary to significantly elevate rabies immunity are few. We used the 22-km2 US National Aeronautics and Space Administration Plum Brook Station (PBS) in Erie County, Ohio, USA, to evaluate the period of exposure for placebo vaccine baits placed at a density of 75 baits/km2 relative to raccoon population density. Our objectives were to 1) estimate raccoon population density within the fragmented forest, old-field, and industrial landscape at PBS: and 2) quantify the time that placebo, Merial RABORAL V-RG vaccine baits were available to raccoons. From August through November 2002 we surveyed raccoon use of PBS along 19.3 km of paved-road transects by using a forward-looking infrared camera mounted inside a vehicle. We used Distance 3.5 software to calculate a probability of detection function by which we estimated raccoon population density from transect data. Estimated population density on PBS decreased from August (33.4 raccoons/km2) through November (13.6 raccoons/km2), yielding a monthly mean of 24.5 raccoons/km2. We also quantified exposure time for ORV baits placed by hand on five 1-km2 grids on PBS from September through October. An average 82.7% (SD = 4.6) of baits were removed within 1 wk of placement. Given raccoon population density, estimates of bait removal and sachet condition, and assuming 22.9% nontarget take, the baiting density of 75/ km2 yielded an average of 3.3 baits consumed per raccoon and the sachet perforated.

  13. Chaotic attractors and physical measures for some density dependent Leslie population models

    NASA Astrophysics Data System (ADS)

    Ugarcovici, Ilie; Weiss, Howard

    2007-12-01

    Following ecologists' discoveries, mathematicians have begun studying extensions of the ubiquitous age structured Leslie population model that allow some survival probabilities and/or fertility rates to depend on population densities. These nonlinear extensions commonly exhibit very complicated dynamics: through computer studies, some authors have discovered robust Hénon-like strange attractors in several families. Population biologists and demographers frequently wish to average a function over many generations and conclude that the average is independent of the initial population distribution. This type of 'ergodicity' seems to be a fundamental tenet in population biology. In this paper we develop the first rigorous ergodic theoretic framework for density dependent Leslie population models. We study two generation models with Ricker and Hassell (recruitment type) fertility terms. We prove that for some parameter regions these models admit a chaotic (ergodic) attractor which supports a unique physical probability measure. This physical measure, having full Lebesgue measure basin, satisfies in the strongest possible sense the population biologist's requirement for ergodicity in their population models. We use the celebrated work of Wang and Young 2001 Commun. Math. Phys. 218 1-97, and our results are the first applications of their method to biology, ecology or demography.

  14. Resolving the interactions between population density and air pollution emissions controls in the San Joaquin Valley, USA.

    PubMed

    Hixson, Mark; Mahmud, Abdullah; Hu, Jianlin; Kleeman, Michael J

    2012-05-01

    The effectiveness of emissions control programs designed to reduce concentrations of airborne particulate matter with an aerodynamic diameter < 2.5 microm (PM2.5) in California's San Joaquin Valley was studied in the year 2030 under three growth scenarios: low, medium, and high population density. Base-case inventories for each choice of population density were created using a coupled emissions modeling system that simultaneously considered interactions between land use and transportation, area source, and point source emissions. The ambient PM2.5 response to each combination of population density and emissions control was evaluated using a regional chemical transport model over a 3-week winter stagnation episode. Comparisons between scenarios were based on regional average and population-weighted PM2.5 concentrations. In the absence of any emissions control program, population-weighted concentrations of PM2.5 in the future San Joaquin Valley are lowest undergrowth scenarios that emphasize low population density. A complete ban on wood burning and a 90% reduction in emissions from food cooking operations and diesel engines must occur before medium- to high-density growth scenarios result in lower population-weighted concentrations of PM2.5. These trends partly reflect the fact that existing downtown urban cores that naturally act as anchor points for new high-density growth in the San Joaquin Valley are located close to major transportation corridors for goods movement. Adding growth buffers around transportation corridors had little impact in the current analysis, since the 8-km resolution of the chemical transport model already provided an artificial buffer around major emissions sources. Assuming that future emissions controls will greatly reduce or eliminate emissions from residential wood burning, food cooking, and diesel engines, the 2030 growth scenario using "as-planned" (medium) population density achieves the lowest population-weighted average PM2.5 concentration in the future San Joaquin Valley during a severe winter stagnation event. The San Joaquin Valley is one of the most heavily polluted air basins in the United States that are projected to experience strong population growth in the coming decades. The best plan to improve air quality in the region combines medium- or high-density population growth with rigorous emissions controls. In the absences of controls, high-density growth leads to increased population exposure to PM2.5 compared with low-density growth scenarios (urban sprawl).

  15. The many faces of population density.

    PubMed

    Mayor, Stephen J; Schaefer, James A

    2005-09-01

    Population density, one of the most fundamental demographic attributes, may vary systematically with spatial scale, but this scale-sensitivity is incompletely understood. We used a novel approach-based on fully censused and mapped distributions of eastern grey squirrel (Sciurus carolinensis) dreys, beaver (Castor canadensis) lodges, and moose (Alces alces)--to explore the scale-dependence of population density and its relationship to landscape features. We identified population units at several scales, both objectively, using cluster analysis, and arbitrarily, using artificial bounds centred on high-abundance sites. Densities declined with census area. For dreys, this relationship was stronger in objective versus arbitrary population units. Drey density was inconsistently related to patch area, a relationship that was positive for all patches but negative when non-occupied patches were excluded. Drey density was negatively related to the proportion of green-space and positively related to the density of buildings or roads, relationships that were accentuated at coarser scales. Mean drey densities were more sensitive to scale when calculated as organism-weighted versus area-weighted averages. Greater understanding of these scaling effects is required to facilitate comparisons of population density across studies.

  16. Population decline of the Elfin-woods Warbler Setophaga angelae in eastern Puerto Rico

    Treesearch

    W.J. Arendt; S.S. Qian; K. Mineard

    2013-01-01

    We estimated the population density of the globally threatened Elfin-woods Warbler Setophaga angelae within two forest types at different elevations in the Luquillo Experimental Forest in north-eastern Puerto Rico. Population densities ranged from 0.01 to 0.02 individuals/ha in elfin woodland and 0.06–0.26 individuals/ha in palo colorado forest in 2006, with average...

  17. Determining mutation density using Restriction Enzyme Sequence Comparative Analysis (RESCAN)

    USDA-ARS?s Scientific Manuscript database

    The average mutation density of a mutant population is a major consideration when developing resources for the efficient, cost-effective implementation of reverse genetics methods such as Targeting of Induced Local Lesions in Genomes (TILLING). Reliable estimates of mutation density can be achieved ...

  18. Multiple density dependence in two sub-populations of the amphipod Monoporeia affinis: a potential for alternative equilibria.

    PubMed

    Leonardsson, Kjell

    1994-02-01

    Possible mechanisms for differences in population densities and dynamics were investigated in the amphipod Monoporeia affinis at two deep sites in the northern Bothnian Sea. The two sites were sampled yearly for 10 years. Average sizes, growth and mortality of the different age-classes were estimated from the cohort structure of the two populations. Laboratory experiments also investigated the ability of the common predatory isopod Saduria entomon to cause densitydependent (DD) mortality of the prey M. affinis. At site A, 43 m depth, the average density of M. affinis was twice as high as at site B, 81 m depth. The fluctuations in density were asynchronous between the two sites. Recruitment and subadult sizes of Monoporeia affinis were density dependent at both sites. The main functional difference between the two populations seemed to be the DD mortality of the 1 + cohort that occurred only at the low-density site B. A corresponding DD mortality was found in the predation experiments at densities of 1 + m. affinis corresponding to those found at site B. The potential importance of the predator was also indicated by a significant negative correlation between the biomass of S. entomon and the rate of change in M. affinis density in the field. The similarities in the abiotic factors between the two sites suggested that differences in carrying capacity should be small. The results could be explained by the predation regulation hypothesis for the low-density population at site B, while at site A M. affinis seemed to be regulated by intra-specific competition and limited by predation. It is suggested that in this simple predator-prey system there is potential for the existence of alternative equilibria.

  19. 12 CFR 563e.12 - Definitions.

    Code of Federal Regulations, 2010 CFR

    2010-01-01

    ... poverty, unemployment, and population loss; or (B) Population size, density, and dispersion. Activities... OTS, based on the year-to-year change in the average of the Consumer Price Index for Urban Wage...

  20. Bayesian Inference on the Effect of Density Dependence and Weather on a Guanaco Population from Chile

    PubMed Central

    Zubillaga, María; Skewes, Oscar; Soto, Nicolás; Rabinovich, Jorge E.; Colchero, Fernando

    2014-01-01

    Understanding the mechanisms that drive population dynamics is fundamental for management of wild populations. The guanaco (Lama guanicoe) is one of two wild camelid species in South America. We evaluated the effects of density dependence and weather variables on population regulation based on a time series of 36 years of population sampling of guanacos in Tierra del Fuego, Chile. The population density varied between 2.7 and 30.7 guanaco/km2, with an apparent monotonic growth during the first 25 years; however, in the last 10 years the population has shown large fluctuations, suggesting that it might have reached its carrying capacity. We used a Bayesian state-space framework and model selection to determine the effect of density and environmental variables on guanaco population dynamics. Our results show that the population is under density dependent regulation and that it is currently fluctuating around an average carrying capacity of 45,000 guanacos. We also found a significant positive effect of previous winter temperature while sheep density has a strong negative effect on the guanaco population growth. We conclude that there are significant density dependent processes and that climate as well as competition with domestic species have important effects determining the population size of guanacos, with important implications for management and conservation. PMID:25514510

  1. Bayesian inference on the effect of density dependence and weather on a guanaco population from Chile.

    PubMed

    Zubillaga, María; Skewes, Oscar; Soto, Nicolás; Rabinovich, Jorge E; Colchero, Fernando

    2014-01-01

    Understanding the mechanisms that drive population dynamics is fundamental for management of wild populations. The guanaco (Lama guanicoe) is one of two wild camelid species in South America. We evaluated the effects of density dependence and weather variables on population regulation based on a time series of 36 years of population sampling of guanacos in Tierra del Fuego, Chile. The population density varied between 2.7 and 30.7 guanaco/km2, with an apparent monotonic growth during the first 25 years; however, in the last 10 years the population has shown large fluctuations, suggesting that it might have reached its carrying capacity. We used a Bayesian state-space framework and model selection to determine the effect of density and environmental variables on guanaco population dynamics. Our results show that the population is under density dependent regulation and that it is currently fluctuating around an average carrying capacity of 45,000 guanacos. We also found a significant positive effect of previous winter temperature while sheep density has a strong negative effect on the guanaco population growth. We conclude that there are significant density dependent processes and that climate as well as competition with domestic species have important effects determining the population size of guanacos, with important implications for management and conservation.

  2. Exploring the relationship between population density and maternal health coverage.

    PubMed

    Hanlon, Michael; Burstein, Roy; Masters, Samuel H; Zhang, Raymond

    2012-11-21

    Delivering health services to dense populations is more practical than to dispersed populations, other factors constant. This engenders the hypothesis that population density positively affects coverage rates of health services. This hypothesis has been tested indirectly for some services at a local level, but not at a national level. We use cross-sectional data to conduct cross-country, OLS regressions at the national level to estimate the relationship between population density and maternal health coverage. We separately estimate the effect of two measures of density on three population-level coverage rates (6 tests in total). Our coverage indicators are the fraction of the maternal population completing four antenatal care visits and the utilization rates of both skilled birth attendants and in-facility delivery. The first density metric we use is the percentage of a population living in an urban area. The second metric, which we denote as a density score, is a relative ranking of countries by population density. The score's calculation discounts a nation's uninhabited territory under the assumption those areas are irrelevant to service delivery. We find significantly positive relationships between our maternal health indicators and density measures. On average, a one-unit increase in our density score is equivalent to a 0.2% increase in coverage rates. Countries with dispersed populations face higher burdens to achieve multinational coverage targets such as the United Nations' Millennial Development Goals.

  3. Body density differences between negro and caucasian professional football players

    PubMed Central

    Adams, J.; Bagnall, K. M.; McFadden, K. D.; Mottola, M.

    1981-01-01

    Other workers have shown that the bone density for the average negro is greater than for the average caucasian. This would lead to greater values of body density for the average negro but it is confused because the average negro has a different body form (and consequently different proportions of body components) compared with the average caucasian. This study of body density of a group of professional Canadian football players investigates whether or not to separate negroes from caucasians when considering the formation of regression equations for prediction of body density. Accordingly, a group of 7 negroes and 7 caucasians were matched somatotypically and a comparison was made of their body density values obtained using a hydrostatic weighing technique and a closed-circuit helium dilution technique for measuring lung volumes. The results show that if somatotype is taken into account then no significant difference in body density values is found between negro and caucasian professional football players. The players do not have to be placed in separate groups but it remains to be seen whether or not these results apply to general members of the population. ImagesFigure 1 PMID:7317724

  4. Evolution of stochastic demography with life history tradeoffs in density-dependent age-structured populations.

    PubMed

    Lande, Russell; Engen, Steinar; Sæther, Bernt-Erik

    2017-10-31

    We analyze the stochastic demography and evolution of a density-dependent age- (or stage-) structured population in a fluctuating environment. A positive linear combination of age classes (e.g., weighted by body mass) is assumed to act as the single variable of population size, [Formula: see text], exerting density dependence on age-specific vital rates through an increasing function of population size. The environment fluctuates in a stationary distribution with no autocorrelation. We show by analysis and simulation of age structure, under assumptions often met by vertebrate populations, that the stochastic dynamics of population size can be accurately approximated by a univariate model governed by three key demographic parameters: the intrinsic rate of increase and carrying capacity in the average environment, [Formula: see text] and [Formula: see text], and the environmental variance in population growth rate, [Formula: see text] Allowing these parameters to be genetically variable and to evolve, but assuming that a fourth parameter, [Formula: see text], measuring the nonlinearity of density dependence, remains constant, the expected evolution maximizes [Formula: see text] This shows that the magnitude of environmental stochasticity governs the classical trade-off between selection for higher [Formula: see text] versus higher [Formula: see text] However, selection also acts to decrease [Formula: see text], so the simple life-history trade-off between [Formula: see text]- and [Formula: see text]-selection may be obscured by additional trade-offs between them and [Formula: see text] Under the classical logistic model of population growth with linear density dependence ([Formula: see text]), life-history evolution in a fluctuating environment tends to maximize the average population size. Published under the PNAS license.

  5. Bulk density of small meteoroids

    NASA Astrophysics Data System (ADS)

    Kikwaya, J.-B.; Campbell-Brown, M.; Brown, P. G.

    2011-06-01

    Aims: Here we report on precise metric and photometric observations of 107 optical meteors, which were simultaneously recorded at multiple stations using three different intensified video camera systems. The purpose is to estimate bulk meteoroid density, link small meteoroids to their parent bodies based on dynamical and physical density values expected for different small body populations, to better understand and explain the dynamical evolution of meteoroids after release from their parent bodies. Methods: The video systems used had image sizes ranging from 640 × 480 to 1360 × 1036 pixels, with pixel scales from 0.01° per pixel to 0.05° per pixel, and limiting meteor magnitudes ranging from Mv = +2.5 to +6.0. We find that 78% of our sample show noticeable deceleration, allowing more robust constraints to be placed on density estimates. The density of each meteoroid is estimated by simultaneously fitting the observed deceleration and lightcurve using a model based on thermal fragmentation, conservation of energy and momentum. The entire phase space of the model free parameters is explored for each event to find ranges of parameters which fit the observations within the measurement uncertainty. Results: (a) We have analysed our data by first associating each of our events with one of the five meteoroid classes. The average density of meteoroids whose orbits are asteroidal and chondritic (AC) is 4200 kg m-3 suggesting an asteroidal parentage, possibly related to the high-iron content population. Meteoroids with orbits belonging to Jupiter family comets (JFCs) have an average density of 3100 ± 300 kg m-3. This high density is found for all meteoroids with JFC-like orbits and supports the notion that the refractory material reported from the Stardust measurements of 81P/Wild 2 dust is common among the broader JFC population. This high density is also the average bulk density for the 4 meteoroids with orbits belonging to the Ecliptic shower-type class (ES) also related to JFCs. Both categories we suggest are chondritic based on their high bulk density. Meteoroids of HT (Halley type) orbits have a minimum bulk density value of 360+400-100 kg m-3 and a maximum value of 1510+400-900 kg m-3. This is consistent with many previous works which suggest bulk cometary meteoroid density is low. SA (Sun-approaching)-type meteoroids show a density spread from 1000 kg m-3 to 4000 kg m-3, reflecting multiple origins. (b) We found two different meteor showers in our sample: Perseids (10 meteoroids, ~11% of our sample) with an average bulk density of 620 kg m-3 and Northern Iota Aquariids (4 meteoroids) with an average bulk density of 3200 kg m-3, consistent with the notion that the NIA derive from 2P/Encke.

  6. Reproduction and population dynamics of Mastomys natalensis Smith, 1834 in an agricultural landscape in the Western Usambara Mountains, Tanzania.

    PubMed

    Makundi, Rhodes H; Massawe, Apia W; Mulungu, Loth S

    2007-12-01

    The multimammate rat, Mastomys natalensis Smith 1834, is a dominant species in agro-ecosystems in Sub-Saharan Africa, but adapts quickly to changes in non-agricultural landscape, particularly woodlands and forests. In this study we report on reproduction and population dynamics of M. natalensis in deforested high elevation localities in the Usambara Mountains, north-east Tanzania. We conducted Capture-Mark-Recapture studies in 2002-2004, and established that reproduction of M. natalensis takes place in the extended wet season between February and June, and the population density peaks in June-August. Reproduction cease in July to January and population density drops from July onwards. Reproduction and population density fluctuations are linked to the duration and amount of rainfall. In years when rainfall was below average and the wet season was short, the population density was significantly lower (below 10 animals/ha and 60 animals/ha in 2003 and 2004 respectively, compared to >100 animals/ha in 2002 when rainfall was above the seasonal average) (F(df 2,13)= 9.092, p < 0.01 for in between years variations and F(df 12,15)= 5.389, p < 0.01 for effect of cumulative annual rainfall on population density). These densities were much lower than in the lowland savannah habitats in central and southwest Tanzania. A comparison between the farmland/fallow mosaic fields and agro-forestry areas showed higher population densities in the former, which have similarities to the preferred habitats in the lowland savannahs. The increasing abundance of M. natalensis in the Usambara could have some consequences: M. natalensis is major pest and is involved in the plague cycle in the western Usambara Mountains. Mastomys natalensis is also a strong competitor and the impact on endemic rodent species, e.g. Lophuromys flavopunctatus and Praomys delectorum is unknown.

  7. Modeling the Response of Anopheles gambiae (Diptera: Culicidae) Populations in the Kenya Highlands to a Rise in Mean Annual Temperature.

    PubMed

    Wallace, Dorothy; Prosper, Olivia; Savos, Jacob; Dunham, Ann M; Chipman, Jonathan W; Shi, Xun; Ndenga, Bryson; Githeko, Andrew

    2017-03-01

    A dynamical model of Anopheles gambiae larval and adult populations is constructed that matches temperature-dependent maturation times and mortality measured experimentally as well as larval instar and adult mosquito emergence data from field studies in the Kenya Highlands. Spectral classification of high-resolution satellite imagery is used to estimate household density. Indoor resting densities collected over a period of one year combined with predictions of the dynamical model give estimates of both aquatic habitat and total adult mosquito densities. Temperature and precipitation patterns are derived from monthly records. Precipitation patterns are compared with average and extreme habitat estimates to estimate available aquatic habitat in an annual cycle. These estimates are coupled with the original model to produce estimates of adult and larval populations dependent on changing aquatic carrying capacity for larvae and changing maturation and mortality dependent on temperature. This paper offers a general method for estimating the total area of aquatic habitat in a given region, based on larval counts, emergence rates, indoor resting density data, and number of households.Altering the average daily temperature and the average daily rainfall simulates the effect of climate change on annual cycles of prevalence of An. gambiae adults. We show that small increases in average annual temperature have a large impact on adult mosquito density, whether measured at model equilibrium values for a single square meter of habitat or tracked over the course of a year of varying habitat availability and temperature. © The Authors 2016. Published by Oxford University Press on behalf of Entomological Society of America. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

  8. Identification of the population density of a species model with nonlocal diffusion and nonlinear reaction

    NASA Astrophysics Data System (ADS)

    Tuan, Nguyen Huy; Van Au, Vo; Khoa, Vo Anh; Lesnic, Daniel

    2017-05-01

    The identification of the population density of a logistic equation backwards in time associated with nonlocal diffusion and nonlinear reaction, motivated by biology and ecology fields, is investigated. The diffusion depends on an integral average of the population density whilst the reaction term is a global or local Lipschitz function of the population density. After discussing the ill-posedness of the problem, we apply the quasi-reversibility method to construct stable approximation problems. It is shown that the regularized solutions stemming from such method not only depend continuously on the final data, but also strongly converge to the exact solution in L 2-norm. New error estimates together with stability results are obtained. Furthermore, numerical examples are provided to illustrate the theoretical results.

  9. DOE Office of Scientific and Technical Information (OSTI.GOV)

    Kostova, T; Carlsen, T

    We present a study, based on simulations with SERDYCA, a spatially-explicit individual based model of rodent dynamics, on the connection between population persistence and the presence of inhomogeneities in the habitat. We are specifically interested on the effect that inhomogeneities that do not fragment the environment, have on population persistence. Our results suggest that a certain percentage of inhomogeneities can increase the average time to extinction of the population. Inhomogeneities decrease the population density and can increase the ratio of juveniles in the population thus providing a better chance for the population to restore itself after a severe period withmore » critically low population density. We call this the ''inhomogeneity localization effect''.« less

  10. The frequency and distribution of recent landslides in three montane tropical regions of Puerto Rico

    Treesearch

    Matthew C. Larsen; Angel J. Torres-Sanchez

    1998-01-01

    Landslides are common in steep mountainous areas of Puerto Rico where mean annual rainfall and the frequency of intense storms are high. Each year, landslides cause extensive damage to property and occasionally result in loss of life. Average population density is high, 422 peoplerkm2, and is increasing. This increase in population density is accompanied by growing...

  11. Exploring the relationship between population density and maternal health coverage

    PubMed Central

    2012-01-01

    Background Delivering health services to dense populations is more practical than to dispersed populations, other factors constant. This engenders the hypothesis that population density positively affects coverage rates of health services. This hypothesis has been tested indirectly for some services at a local level, but not at a national level. Methods We use cross-sectional data to conduct cross-country, OLS regressions at the national level to estimate the relationship between population density and maternal health coverage. We separately estimate the effect of two measures of density on three population-level coverage rates (6 tests in total). Our coverage indicators are the fraction of the maternal population completing four antenatal care visits and the utilization rates of both skilled birth attendants and in-facility delivery. The first density metric we use is the percentage of a population living in an urban area. The second metric, which we denote as a density score, is a relative ranking of countries by population density. The score’s calculation discounts a nation’s uninhabited territory under the assumption those areas are irrelevant to service delivery. Results We find significantly positive relationships between our maternal health indicators and density measures. On average, a one-unit increase in our density score is equivalent to a 0.2% increase in coverage rates. Conclusions Countries with dispersed populations face higher burdens to achieve multinational coverage targets such as the United Nations’ Millennial Development Goals. PMID:23170895

  12. High urban population density of birds reflects their timing of urbanization.

    PubMed

    Møller, Anders Pape; Diaz, Mario; Flensted-Jensen, Einar; Grim, Tomas; Ibáñez-Álamo, Juan Diego; Jokimäki, Jukka; Mänd, Raivo; Markó, Gábor; Tryjanowski, Piotr

    2012-11-01

    Living organisms generally occur at the highest population density in the most suitable habitat. Therefore, invasion of and adaptation to novel habitats imply a gradual increase in population density, from that at or below what was found in the ancestral habitat to a density that may reach higher levels in the novel habitat following adaptation to that habitat. We tested this prediction of invasion biology by analyzing data on population density of breeding birds in their ancestral rural habitats and in matched nearby urban habitats that have been colonized recently across a continental latitudinal gradient. We estimated population density in the two types of habitats using extensive point census bird counts, and we obtained information on the year of urbanization when population density in urban habitats reached levels higher than that of the ancestral rural habitat from published records and estimates by experienced ornithologists. Both the difference in population density between urban and rural habitats and the year of urbanization were significantly repeatable when analyzing multiple populations of the same species across Europe. Population density was on average 30 % higher in urban than in rural habitats, although density reached as much as 100-fold higher in urban habitats in some species. Invasive urban bird species that colonized urban environments over a long period achieved the largest increases in population density compared to their ancestral rural habitats. This was independent of whether species were anciently or recently urbanized, providing a unique cross-validation of timing of urban invasions. These results suggest that successful invasion of urban habitats was associated with gradual adaptation to these habitats as shown by a significant increase in population density in urban habitats over time.

  13. [Variation pattern and its affecting factors of three-dimensional landscape in urban residential community of Shenyang].

    PubMed

    Zhang, Pei-Feng; Hu, Yuan-Man; Xiong, Zai-Ping; Liu, Miao

    2011-02-01

    Based on the 1:10000 aerial photo in 1997 and the three QuickBird images in 2002, 2005, and 2008, and by using Barista software and GIS and RS techniques, the three-dimensional information of the residential community in Tiexi District of Shenyang was extracted, and the variation pattern of the three-dimensional landscape in the district during its reconstruction in 1997-2008 and related affecting factors were analyzed with the indices, ie. road density, greening rate, average building height, building height standard deviation, building coverage rate, floor area rate, building shape coefficient, population density, and per capita GDP. The results showed that in 1997-2008, the building area for industry decreased, that for commerce and other public affairs increased, and the area for residents, education, and medical cares basically remained stable. The building number, building coverage rate, and building shape coefficient decreased, while the floor area rate, average building height, height standard deviation, road density, and greening rate increased. Within the limited space of residential community, the containing capacity of population and economic activity increased, and the environment quality also improved to some extent. The variation degree of average building height increased, but the building energy consumption decreased. Population growth and economic development had positive correlations with floor area rate, road density, and greening rate, but negative correlation with building coverage rate.

  14. Effects of urban sprawl on obesity.

    PubMed

    Zhao, Zhenxiang; Kaestner, Robert

    2010-12-01

    In this paper, we examine the effect of changes in population density-urban sprawl-between 1970 and 2000 on BMI and obesity of residents in metropolitan areas in the U.S. We address the possible endogeneity of population density by using a two-step instrumental variables approach. We exploit the plausibly exogenous variation in population density caused by the expansion of the U.S. Interstate Highway System, which largely followed the original 1947 plan for the Interstate Highway System. We find a negative association between population density and obesity, and estimates are robust across a wide range of specifications. Estimates indicate that if the average metropolitan area had not experienced the decline in the proportion of population living in dense areas over the last 30 years, the rate of obesity would have been reduced by approximately 13%. Copyright © 2010 Elsevier B.V. All rights reserved.

  15. Fraser River sockeye salmon productivity and climate: A re-analysis that avoids an undesirable property of Ricker’s curve

    NASA Astrophysics Data System (ADS)

    McKinnell, Skip

    2008-05-01

    In descending order of importance, artificial spawning channels, density-dependent mortality, carryover mortality, and climate have significant influences on the average productivity of Fraser River sockeye salmon ( Oncorhynchus nerka). When factors that are known or have been hypothesized to affect Fraser River sockeye salmon productivity are included in a single analytical framework, no significant change in average productivity occurred in 1976/1977, however, beginning in 1989 average productivity was significantly lower. In the one lake (Chilko) in the Fraser River basin where pre-smolt survival can be distinguished from post-smolt survival, this decline arose from freshwater causes. After accounting for other factors that have a greater influence, Fraser River sockeye salmon productivity tends to be slightly lower in years when the intensity of the Aleutian low pressure region is stormier in winter, although the effect is not strongly expressed in any particular population. A footnote to the study was the realization that estimates of Ricker’s density-dependent mortality parameter, β, are influenced by both the numerical properties of the equation and by population biology; density-dependent and density-independent influences on the estimates of the parameter are confounded.

  16. A new approach on seismic mortality estimations based on average population density

    NASA Astrophysics Data System (ADS)

    Zhu, Xiaoxin; Sun, Baiqing; Jin, Zhanyong

    2016-12-01

    This study examines a new methodology to predict the final seismic mortality from earthquakes in China. Most studies established the association between mortality estimation and seismic intensity without considering the population density. In China, however, the data are not always available, especially when it comes to the very urgent relief situation in the disaster. And the population density varies greatly from region to region. This motivates the development of empirical models that use historical death data to provide the path to analyze the death tolls for earthquakes. The present paper employs the average population density to predict the final death tolls in earthquakes using a case-based reasoning model from realistic perspective. To validate the forecasting results, historical data from 18 large-scale earthquakes occurred in China are used to estimate the seismic morality of each case. And a typical earthquake case occurred in the northwest of Sichuan Province is employed to demonstrate the estimation of final death toll. The strength of this paper is that it provides scientific methods with overall forecast errors lower than 20 %, and opens the door for conducting final death forecasts with a qualitative and quantitative approach. Limitations and future research are also analyzed and discussed in the conclusion.

  17. Determining the functional form of density dependence: deductive approaches for consumer-resource systems having a single resource.

    PubMed

    Abrams, Peter A

    2009-09-01

    Consumer-resource models are used to deduce the functional form of density dependence in the consumer population. A general approach to determining the form of consumer density dependence is proposed; this involves determining the equilibrium (or average) population size for a series of different harvest rates. The relationship between a consumer's mortality and its equilibrium population size is explored for several one-consumer/one-resource models. The shape of density dependence in the resource and the shape of the numerical and functional responses all tend to be "inherited" by the consumer's density dependence. Consumer-resource models suggest that density dependence will very often have both concave and convex segments, something that is impossible under the commonly used theta-logistic model. A range of consumer-resource models predicts that consumer population size often declines at a decelerating rate with mortality at low mortality rates, is insensitive to or increases with mortality over a wide range of intermediate mortalities, and declines at a rapidly accelerating rate with increased mortality when mortality is high. This has important implications for management and conservation of natural populations.

  18. Water quality of Lake Arlington on Village Creek, north-central Texas; 1973 to 1981

    USGS Publications Warehouse

    Andrews, Freeman L.; Gibbons, Willard J.

    1983-01-01

    The densities and composition of algal populations varied seasonally. At site AC, total algae counts ranged from 200 to 240,000 cells per mi Hi liter and averaged about 50,000 cells per milliliter. At site FC, total algae counts ranged from 1,000 to 290,000 cells per milliliter and averaged about 56,000 cells per milliliter. Algal densities were greatest during the summer with blue-green algae being the predominant phyla.

  19. Effects of roadside habitat and fox density on a snow track survey for foxes in Ohio

    USGS Publications Warehouse

    Stanley, Thomas R.; Bart, Jonathan

    1991-01-01

    Many methods have been used to survey red fox (Vulpes vulpes) and gray fox (Urocyon dnereoargenteus) populations. However, none has proven entirely satisfactory, and wild foxes remain one of the most difficult economically important wildlife species to monitor. In this study we evaluated the reliability of a snow track survey method for foxes by investigating whether the average number of road crossings per fox is influenced by changes in roadside habitat or changes in fox density. Several snow track surveys were conducted in two Ohio counties during January and February, 1984. Data on roadside habitat, relative fox densities, and fox crossings were collected. Results suggested that changes in roadside habitat could influence the average number of crossings per fox and, therefore, changes in the index could occur independent of actual population changes. We found no evidence that crossings per fox varied with fox density, but further research is needed to substantiate this finding.

  20. Temporal changes in deep-sea sponge populations are correlated to changes in surface climate and food supply

    NASA Astrophysics Data System (ADS)

    Kahn, Amanda S.; Ruhl, Henry A.; Smith, Kenneth L.

    2012-12-01

    Density and average size of two species of abyssal sponges were analyzed at Station M (∼4100 m depth) over an 18-year time-series (1989-2006) using camera sled transects. Both sponge taxa share a similar plate-like morphology despite being within different families, and both showed similar variations in density and average body size over time, suggesting that the same factors may control the demographics of both species. Peaks in significant cross correlations between increases in particulate organic carbon flux and corresponding increases in sponge density occurred with a time lag of 13 months. Sponge density also fluctuated with changes in two climate indices: the NOI with a time lag of 18 months and NPGO with a time lag of 15 months. The results support previous suggestions that increased particulate organic carbon flux may induce recruitment or regeneration in deep-sea sponges. It is unknown whether the appearance of young individuals results from recruitment, regeneration, or both, but the population responses to seasonal and inter-annual changes in food supply demonstrate that sponge populations are dynamic and are capable of responding to inter-annual changes despite being sessile and presumably slow-growing.

  1. Population Dynamics of Dactylella oviparasitica and Heterodera schachtii: Toward a Decision Model for Sugar Beet Planting

    PubMed Central

    Yang, Jiue-in; Benecke, Scott; Jeske, Daniel R.; Rocha, Fernando S.; Smith Becker, Jennifer; Timper, Patricia; Ole Becker, J.

    2012-01-01

    A series of experiments were performed to examine the population dynamics of the sugarbeet cyst nematode, Heterodera schachtii, and the nematophagus fungus Dactylella oviparasitica. After two nematode generations, the population densities of H. schachtii were measured in relation to various initial infestation densities of both D. oviparasitica and H. schachtii. In general, higher initial population densities of D. oviparasitica were associated with lower final population densities of H. schachtii. Regression models showed that the initial densities of D. oviparasitica were only significant when predicting the final densities of H. schachtii J2 and eggs as well as fungal egg parasitism, while the initial densities of J2 were significant for all final H. schachtii population density measurements. We also showed that the densities of H. schachtii-associated D. oviparasitica fluctuate greatly, with rRNA gene numbers going from zero in most field-soil-collected cysts to an average of 4.24 x 108 in mature females isolated directly from root surfaces. Finally, phylogenetic analysis of rRNA genes suggested that D. oviparasitica belongs to a clade of nematophagous fungi that includes Arkansas Fungus strain L (ARF-L) and that these fungi are widely distributed. We anticipate that these findings will provide foundational data facilitating the development of more effective decision models for sugar beet planting. PMID:23481664

  2. Relationships between population density, fine-scale genetic structure, mating system and pollen dispersal in a timber tree from African rainforests.

    PubMed

    Duminil, J; Daïnou, K; Kaviriri, D K; Gillet, P; Loo, J; Doucet, J-L; Hardy, O J

    2016-03-01

    Owing to the reduction of population density and/or the environmental changes it induces, selective logging could affect the demography, reproductive biology and evolutionary potential of forest trees. This is particularly relevant in tropical forests where natural population densities can be low and isolated trees may be subject to outcross pollen limitation and/or produce low-quality selfed seeds that exhibit inbreeding depression. Comparing reproductive biology processes and genetic diversity of populations at different densities can provide indirect evidence of the potential impacts of logging. Here, we analysed patterns of genetic diversity, mating system and gene flow in three Central African populations of the self-compatible legume timber species Erythrophleum suaveolens with contrasting densities (0.11, 0.68 and 1.72 adults per ha). The comparison of inbreeding levels among cohorts suggests that selfing is detrimental as inbred individuals are eliminated between seedling and adult stages. Levels of genetic diversity, selfing rates (∼16%) and patterns of spatial genetic structure (Sp ∼0.006) were similar in all three populations. However, the extent of gene dispersal differed markedly among populations: the average distance of pollen dispersal increased with decreasing density (from 200 m in the high-density population to 1000 m in the low-density one). Overall, our results suggest that the reproductive biology and genetic diversity of the species are not affected by current logging practices. However, further investigations need to be conducted in low-density populations to evaluate (1) whether pollen limitation may reduce seed production and (2) the regeneration potential of the species.

  3. A bottom up approach to on-road CO2 emissions estimates: improved spatial accuracy and applications for regional planning.

    PubMed

    Gately, Conor K; Hutyra, Lucy R; Wing, Ian Sue; Brondfield, Max N

    2013-03-05

    On-road transportation is responsible for 28% of all U.S. fossil-fuel CO2 emissions. Mapping vehicle emissions at regional scales is challenging due to data limitations. Existing emission inventories use spatial proxies such as population and road density to downscale national or state-level data. Such procedures introduce errors where the proxy variables and actual emissions are weakly correlated, and limit analysis of the relationship between emissions and demographic trends at local scales. We develop an on-road emission inventory product for Massachusetts-based on roadway-level traffic data obtained from the Highway Performance Monitoring System (HPMS). We provide annual estimates of on-road CO2 emissions at a 1 × 1 km grid scale for the years 1980 through 2008. We compared our results with on-road emissions estimates from the Emissions Database for Global Atmospheric Research (EDGAR), with the Vulcan Product, and with estimates derived from state fuel consumption statistics reported by the Federal Highway Administration (FHWA). Our model differs from FHWA estimates by less than 8.5% on average, and is within 5% of Vulcan estimates. We found that EDGAR estimates systematically exceed FHWA by an average of 22.8%. Panel regression analysis of per-mile CO2 emissions on population density at the town scale shows a statistically significant correlation that varies systematically in sign and magnitude as population density increases. Population density has a positive correlation with per-mile CO2 emissions for densities below 2000 persons km(-2), above which increasing density correlates negatively with per-mile emissions.

  4. Visiting Patterns and Effects of Density at a Visitors' Center.

    ERIC Educational Resources Information Center

    Burrus-Bammel, Lei Lane; Bammel, Gene

    1986-01-01

    Reports on a study designed to investigate the relationship between the length of stay at a national forest visitor center and the density of the visitor population. Results indicate a significant negative correlation exists between the daily number of users and the average duration of their visits. (TW)

  5. Consequences of cannibalism and competition for food in a smallmouth bass population: An individual-based modeling study

    USGS Publications Warehouse

    Dong, Q.; DeAngelis, D.L.

    1998-01-01

    We used an individual-based modeling approach to study the consequences of cannibalism and competition for food in a freshwater fish population. We simulated the daily foraging, growth, and survival of the age-0 fish and older juvenile individuals of a sample population to reconstruct patterns of density dependence in the age-0 fish during the growth season. Cannibalism occurs as a part of the foraging process. For age-0 fish, older juvenile fish are both potential cannibals and competitors of food. We found that competition and cannibalism produced intraclass and interclass density dependence. Our modeling results suggested the following. (1) With low density of juvenile fish and weak interclass interactions, the age-0 fish recruitment shows a Beverton-Holt type of density dependence. (2) With high density of juvenile fish and strong interclass interactions, the age-0 fish recruitment shows a Ricker type of density dependence, and overcompensation occurs. (3) Interclass competition of food is responsible for much of the overcompensation. (4) Cannibalism intensifies the changes in the recruitment that are brought about by competition. Cannibalism can (a) generally reduce the recruitment, (b) particularly reduce the maximum level of recruitment, (c) cause overcompensation to occur at lower densities, and (d) produce a stronger overcompensation. (5) Growth is also a function of density. Cannibalism generally improves average growth of cannibals. (6) Variation in the lengths of age-0 fish increases with density and with a decreased average growth. These results imply that cannibalism and competition for food could strongly affect recruitment dynamics. Our model also showed that the rate of cannibalism either could be fairly even through the whole season or could vary dramatically. The individual-based modeling approach can help ecologists understand the mechanistic connection between daily behavioral and physiological processes operating at the level of individual organisms and seasonal patterns of population structure and dynamics. ?? Copyright by the American Fisheries Society 1998.

  6. Density-dependent regulation of population size in colonial breeders: Allee and buffer effects in the migratory Montagu's harrier.

    PubMed

    Soutullo, Alvaro; Limiñana, Rubén; Urios, Vicente; Surroca, Martín; A Gill, Jennifer

    2006-09-01

    Expanding populations offer an opportunity to uncover the processes driving spatial variation in distribution and abundance. Individual settlement decisions will be influenced by the availability and relative quality of patches, and by how these respond to changes in conspecific density. For example, conspecific presence can alter patch suitability through reductions in resource availability or territorial exclusion, leading to buffer effect patterns of disproportionate population expansion into poorer quality areas. However, conspecific presence can also enhance patch suitability through Allee effect processes, such as transmission of information about resources or improved predator detection and deterrence. Here, we explore the factors underlying the settlement pattern of a growing population of Montagu's harriers (Circus pygargus) in Spain. The population increased exponentially between 1981 and 2001, but stabilised between 2001 and 2004. This population increase occurred alongside a remarkable spatial expansion, with novel site use occurring prior to maximum densities in occupied sites being reached. However, no temporal trends in fecundity were observed and, within sites, average fecundity did not decline with increasing density. Across the population, variance in productivity did increase with population size, suggesting a complex pattern of density-dependent costs and benefits. We suggest that both Allee and buffer effects are operating in this system, with the benefits of conspecific presence counteracting density-dependent declines in resource availability or quality.

  7. Premature greying of the hair is not associated with low bone mineral density.

    PubMed

    Beardsworth, S A; Kearney, C E; Steel, S A; Newman, J; Purdie, D W

    1999-01-01

    In two recent case-control studies premature greying of the hair was associated with a lowering of bone mineral density (BMD) and osteopenia, suggesting that this might be a clinically useful risk marker for osteoporosis. We report a further re-examination of this proposal in 52 prematurely grey-haired women from East Yorkshire who responded to an advertisement inviting them for bone densitometry. Thirty-five had no clinical or drug history that could influence bone density. All were Caucasian with a mean age of 52.8 years. In the group as a whole the mean BMD values at the lumbar spine and femoral neck were no different from those of a young adult, but there was a trend toward a greater than average BMD than that of the local age-matched population (p = 0.097 and 0.218, respectively). Twenty women were premenopausal, with an average age of 45.3 years. Mean BMD values at the lumbar spine and femoral neck in this group were no different from those of young adults. There was, however, a trend toward a BMD greater than that of the local age-matched population at the femoral neck (p = 0.117). Fifteen women were postmenopausal with an average age of 62.9 years and an average age at menopause of 51.1 years. Mean BMD values at both the lumbar spine and femoral neck in this group were lower than those of young adults, but no different from those of the local age-matched population. In conclusion, our group of prematurely grey-haired women had average BMD for their age, and we are therefore unable to support the proposed clinical usefulness of premature greying as a risk marker for osteoporosis.

  8. Relationships between population density, fine-scale genetic structure, mating system and pollen dispersal in a timber tree from African rainforests

    PubMed Central

    Duminil, J; Daïnou, K; Kaviriri, D K; Gillet, P; Loo, J; Doucet, J-L; Hardy, O J

    2016-01-01

    Owing to the reduction of population density and/or the environmental changes it induces, selective logging could affect the demography, reproductive biology and evolutionary potential of forest trees. This is particularly relevant in tropical forests where natural population densities can be low and isolated trees may be subject to outcross pollen limitation and/or produce low-quality selfed seeds that exhibit inbreeding depression. Comparing reproductive biology processes and genetic diversity of populations at different densities can provide indirect evidence of the potential impacts of logging. Here, we analysed patterns of genetic diversity, mating system and gene flow in three Central African populations of the self-compatible legume timber species Erythrophleum suaveolens with contrasting densities (0.11, 0.68 and 1.72 adults per ha). The comparison of inbreeding levels among cohorts suggests that selfing is detrimental as inbred individuals are eliminated between seedling and adult stages. Levels of genetic diversity, selfing rates (∼16%) and patterns of spatial genetic structure (Sp ∼0.006) were similar in all three populations. However, the extent of gene dispersal differed markedly among populations: the average distance of pollen dispersal increased with decreasing density (from 200 m in the high-density population to 1000 m in the low-density one). Overall, our results suggest that the reproductive biology and genetic diversity of the species are not affected by current logging practices. However, further investigations need to be conducted in low-density populations to evaluate (1) whether pollen limitation may reduce seed production and (2) the regeneration potential of the species. PMID:26696137

  9. DOE Office of Scientific and Technical Information (OSTI.GOV)

    Kostova, T; Carlsen, T

    We present a study, based on simulations with SERDYCA, a spatially-explicit individual-based model of rodent dynamics, on the relation between population persistence and the presence of numerous isolated disturbances in the habitat. We are specifically interested in the effect of disturbances that do not fragment the environment on population persistence. Our results suggest that the presence of disturbances in the absence of fragmentation can actually increase the average time to extinction of the modeled population. The presence of disturbances decreases population density but can increase the chance for mating in monogamous species and consequently, the ratio of juveniles in themore » population. It thus provides a better chance for the population to restore itself after a severe period with critically low population density. We call this the ''disturbance-forced localization effect''.« less

  10. Sensitivity of MSE measurements on the beam atomic level population

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Ruiz, C., E-mail: carlos.ruiz@wisc.edu; Kumar, S. T. A.; Anderson, F. S. B.

    The effect of variation in atomic level population of a neutral beam on the Motional Stark Effect (MSE) measurements is investigated in the low density plasmas of HSX stellarator. A 30 KeV, 4 A, 3 ms hydrogen diagnostic neutral beam is injected into HSX plasmas of line averaged electron density ranging from 2 to 4 ⋅ 10{sup 18} m{sup −3} at a magnetic field of 1 T. For this density range, the excited level population of the hydrogen neutral beam is expected to undergo variations. Doppler shifted and Stark split H{sub α} and H{sub β} emissions from the beam aremore » simultaneously measured using two cross-calibrated spectrometers. The emission spectrum is simulated and fit to the experimental measurements and the deviation from a statistically populated beam is investigated.« less

  11. Density-Dependent Regulation of Brook Trout Population Dynamics along a Core-Periphery Distribution Gradient in a Central Appalachian Watershed

    PubMed Central

    Huntsman, Brock M.; Petty, J. Todd

    2014-01-01

    Spatial population models predict strong density-dependence and relatively stable population dynamics near the core of a species' distribution with increasing variance and importance of density-independent processes operating towards the population periphery. Using a 10-year data set and an information-theoretic approach, we tested a series of candidate models considering density-dependent and density-independent controls on brook trout population dynamics across a core-periphery distribution gradient within a central Appalachian watershed. We sampled seven sub-populations with study sites ranging in drainage area from 1.3–60 km2 and long-term average densities ranging from 0.335–0.006 trout/m. Modeled response variables included per capita population growth rate of young-of-the-year, adult, and total brook trout. We also quantified a stock-recruitment relationship for the headwater population and coefficients of variability in mean trout density for all sub-populations over time. Density-dependent regulation was prevalent throughout the study area regardless of stream size. However, density-independent temperature models carried substantial weight and likely reflect the effect of year-to-year variability in water temperature on trout dispersal between cold tributaries and warm main stems. Estimated adult carrying capacities decreased exponentially with increasing stream size from 0.24 trout/m in headwaters to 0.005 trout/m in the main stem. Finally, temporal variance in brook trout population size was lowest in the high-density headwater population, tended to peak in mid-sized streams and declined slightly in the largest streams with the lowest densities. Our results provide support for the hypothesis that local density-dependent processes have a strong control on brook trout dynamics across the entire distribution gradient. However, the mechanisms of regulation likely shift from competition for limited food and space in headwater streams to competition for thermal refugia in larger main stems. It also is likely that source-sink dynamics and dispersal from small headwater habitats may partially influence brook trout population dynamics in the main stem. PMID:24618602

  12. Density dependence in a recovering osprey population: demographic and behavioural processes.

    PubMed

    Bretagnolle, V; Mougeot, F; Thibault, J-C

    2008-09-01

    1. Understanding how density-dependent and independent processes influence demographic parameters, and hence regulate population size, is fundamental within population ecology. We investigated density dependence in growth rate and fecundity in a recovering population of a semicolonial raptor, the osprey Pandion haliaetus [Linnaeus, 1758], using 31 years of count and demographic data in Corsica. 2. The study population increased from three pairs in 1974 to an average of 22 pairs in the late 1990s, with two distinct phases during the recovery (increase followed by stability) and contrasted trends in breeding parameters in each phase. 3. We show density dependence in population growth rate in the second phase, indicating that the stabilized population was regulated. We also show density dependence in productivity (fledging success between years and hatching success within years). 4. Using long-term data on behavioural interactions at nest sites, and on diet and fish provisioning rate, we evaluated two possible mechanisms of density dependence in productivity, food depletion and behavioural interference. 5. As density increased, both provisioning rate and the size of prey increased, contrary to predictions of a food-depletion mechanism. In the time series, a reduction in fledging success coincided with an increase in the number of non-breeders. Hatching success decreased with increasing local density and frequency of interactions with conspecifics, suggesting that behavioural interference was influencing hatching success. 6. Our study shows that, taking into account the role of non-breeders, in particular in species or populations where there are many floaters and where competition for nest sites is intense, can improve our understanding of density-dependent processes and help conservation actions.

  13. The relationship between population density and cancer mortality in Taiwan.

    PubMed

    Yang, C Y; Hsieh, Y L

    1998-04-01

    Many investigators have examined urbanization gradients in cancer rates. The purpose of this report was to identify urban-rural trends in cancer mortality rates (1982-1991) for municipalities in Taiwan. For this purpose, Taiwan's municipalities were classified as rural, suburban, urban, or metropolitan, using population density as an ordinal indicator of the degree of urbanization. Average annual age-adjusted, site-specific cancer mortality rates were calculated for both sexes within each population density group. Significant increasing trends with more urbanization were observed in mortality rates for cancers of the lung, pancreas, and kidney among both males and females, as well as male prostate cancer, and female breast and ovary cancer. In addition, this study revealed a significant rural excess for nonmelanoma skin cancer among both males and females, as well as male non-Hodgkin's lymphoma, and cancers of the female bone, and female connective tissue. Analytic studies for sites with consistent urban-rural trends may be fruitful in identifying the aspect of population density, or other unmeasured factors, that contribute to these trends.

  14. Effects of the distant population density on spatial patterns of demographic dynamics

    NASA Astrophysics Data System (ADS)

    Tamura, Kohei; Masuda, Naoki

    2017-08-01

    Spatio-temporal patterns of population changes within and across countries have various implications. Different geographical, demographic and econo-societal factors seem to contribute to migratory decisions made by individual inhabitants. Focusing on internal (i.e. domestic) migration, we ask whether individuals may take into account the information on the population density in distant locations to make migratory decisions. We analyse population census data in Japan recorded with a high spatial resolution (i.e. cells of size 500×500 m) for the entirety of the country, and simulate demographic dynamics induced by the gravity model and its variants. We show that, in the census data, the population growth rate in a cell is positively correlated with the population density in nearby cells up to a distance of 20 km as well as that of the focal cell. The ordinary gravity model does not capture this empirical observation. We then show that the empirical observation is better accounted for by extensions of the gravity model such that individuals are assumed to perceive the attractiveness, approximated by the population density, of the source or destination cell of migration as the spatial average over a circle of radius ≈1 km.

  15. Effects of the distant population density on spatial patterns of demographic dynamics.

    PubMed

    Tamura, Kohei; Masuda, Naoki

    2017-08-01

    Spatio-temporal patterns of population changes within and across countries have various implications. Different geographical, demographic and econo-societal factors seem to contribute to migratory decisions made by individual inhabitants. Focusing on internal (i.e. domestic) migration, we ask whether individuals may take into account the information on the population density in distant locations to make migratory decisions. We analyse population census data in Japan recorded with a high spatial resolution (i.e. cells of size 500×500  m ) for the entirety of the country, and simulate demographic dynamics induced by the gravity model and its variants. We show that, in the census data, the population growth rate in a cell is positively correlated with the population density in nearby cells up to a distance of 20 km as well as that of the focal cell. The ordinary gravity model does not capture this empirical observation. We then show that the empirical observation is better accounted for by extensions of the gravity model such that individuals are assumed to perceive the attractiveness, approximated by the population density, of the source or destination cell of migration as the spatial average over a circle of radius ≈1 km.

  16. Effects of the distant population density on spatial patterns of demographic dynamics

    PubMed Central

    2017-01-01

    Spatio-temporal patterns of population changes within and across countries have various implications. Different geographical, demographic and econo-societal factors seem to contribute to migratory decisions made by individual inhabitants. Focusing on internal (i.e. domestic) migration, we ask whether individuals may take into account the information on the population density in distant locations to make migratory decisions. We analyse population census data in Japan recorded with a high spatial resolution (i.e. cells of size 500×500 m) for the entirety of the country, and simulate demographic dynamics induced by the gravity model and its variants. We show that, in the census data, the population growth rate in a cell is positively correlated with the population density in nearby cells up to a distance of 20 km as well as that of the focal cell. The ordinary gravity model does not capture this empirical observation. We then show that the empirical observation is better accounted for by extensions of the gravity model such that individuals are assumed to perceive the attractiveness, approximated by the population density, of the source or destination cell of migration as the spatial average over a circle of radius ≈1 km. PMID:28878987

  17. Use of non-invasive genetics to generate core-area population estimates of a threatened predator in the Superior National Forest, USA

    USGS Publications Warehouse

    Barber-Meyer, Shannon; Ryan, Daniel; Grosshuesch, David; Catton, Timothy; Malick-Wahls, Sarah

    2018-01-01

    core areas and averaged 52.3 (SD=8.3, range=43-59) during 2015-2017 in the larger core areas. We found no evidence for a decrease or increase in abundance during either period. Lynx density estimates were approximately 7-10 times lower than densities of lynx in northern populations at the low of the snowshoe hare (Lepus americanus) population cycle. To our knowledge, our results are the first attempt to estimate abundance, trend and density of lynx in Minnesota using non-invasive genetic capture-mark-recapture. Estimates such as ours provide useful benchmarks for future comparisons by providing a context with which to assess 1) potential changes in forest management that may affect lynx recovery and conservation, and 2) possible effects of climate change on the depth, density, and duration of annual snow cover and correspondingly, potential effects on snowshoe hares as well.

  18. Single-treatment porcine zona pellucida immunocontraception associated with reduction of a population of white-tailed deer (Odocoileus virginianus).

    PubMed

    Rutberg, Allen T; Naugle, Ricky E; Verret, Frank

    2013-12-01

    Previous reports have demonstrated gradual reductions of white-tailed deer (Odocoileus virginianus) populations through immunocontraception, with stabilization occurring after 2-4 yr of treatment, and subsequent reductions of 6-10% annually. These studies employed porcine zona pellucida (PZP) vaccines that required two initial treatments and annual retreatments. From 2005 to 2010, 258 adult and yearling female deer on Fripp Island, South Carolina, were treated with one of several PZP preparations designed to produce 2+ yr of effective contraception with a single treatment. These included several preparations of SpayVac and of native PZP-adjuvant emulsion plus PZP and QA-21 in timed-release pellets. Deer were chemically immobilized, ear-tagged, and administered initial treatments by hand in February-March. Some treated deer were boosted remotely with PZP-adjuvant emulsion 1.5 - 4.5 yr after initial treatments. Ground-based distance sampling was used to estimate deer population density at Fripp Island, a resort community, and at a relatively undeveloped neighboring control site, Hunting Island. Most vaccine preparations tested reduced fawning rates by 75% to 95% for at least 1 yr. From 2005 to 2011, deer density on Fripp Island declined by 50%, from 72 deer/km(2) to 36 deer/km(2), an average annual reduction of 11%. In contrast, population density on the Hunting Island control site fluctuated between 2005 and 2011, averaging 23 deer/km(2) (range, 19-28 deer/km(2)). Population declines on Fripp Island were associated with an increase in the proportion of treated females and with a progressive decrease in winter fawn:doe ratios, from 1.21 fawns/doe in 2005 to 0.19 fawns/doe in 2010. Winter fawn:doe ratios averaged 1.36 fawns/doe (range, 0.84 - 1.62 fawns/doe) at the Hunting Island control site. Annual survivorship averaged approximately 79% among ear-tagged females. The rate at which deer populations diminished in association with PZP treatments on Fripp Island was higher than that seen at other study sites, although the reasons for the more rapid decline on Fripp Island are not well understood.

  19. Grizzly bear density in Glacier National Park, Montana

    USGS Publications Warehouse

    Kendall, K.C.; Stetz, J.B.; Roon, David A.; Waits, L.P.; Boulanger, J.B.; Paetkau, David

    2008-01-01

    We present the first rigorous estimate of grizzly bear (Ursus arctos) population density and distribution in and around Glacier National Park (GNP), Montana, USA. We used genetic analysis to identify individual bears from hair samples collected via 2 concurrent sampling methods: 1) systematically distributed, baited, barbed-wire hair traps and 2) unbaited bear rub trees found along trails. We used Huggins closed mixture models in Program MARK to estimate total population size and developed a method to account for heterogeneity caused by unequal access to rub trees. We corrected our estimate for lack of geographic closure using a new method that utilizes information from radiocollared bears and the distribution of bears captured with DNA sampling. Adjusted for closure, the average number of grizzly bears in our study area was 240.7 (95% CI = 202–303) in 1998 and 240.6 (95% CI = 205–304) in 2000. Average grizzly bear density was 30 bears/1,000 km2, with 2.4 times more bears detected per hair trap inside than outside GNP. We provide baseline information important for managing one of the few remaining populations of grizzlies in the contiguous United States.

  20. Population density and racial differences in the performance of emergency medical services.

    PubMed

    David, Guy; Harrington, Scott E

    2010-07-01

    This paper analyzes the existence and scope of possible racial differences/disparities in the provision of emergency medical services (EMS) response capability (time from dispatch to arrival at the scene and level of training of the responding team) using data on approximately 120,000 cardiac incidents in the state of Mississippi during 1995-2004. The conceptual framework and empirical analysis focus on the likely effects of population density on the efficient production of EMS as a local public good subject to congestion, and on the need to control adequately for population density to avoid bias in testing for racial differences. Models that control for aggregate population density at the county-level indicate "reverse" disparities: faster estimated response times for African-Americans than for whites. When a refined county-level measure of population density is used that incorporates differences in African-American and white population density by Census tract, the reverse disparity in response times disappears. There also is little or no evidence of race-related differences in the certification level of EMS responders. However, there is evidence that, controlling for response time, African-Americans on average were significantly more likely to be deceased than whites upon EMS arrival at the scene. The overall results are germane to the debate over the scope of conditioning variables that should be included when testing for racial disparities in health care.

  1. Quantitative Relationship of Soil Texture with the Observed Population Density Reduction of Heterodera glycines after Annual Corn Rotation in Nebraska

    PubMed Central

    Pérez-Hernández, Oscar; Giesler, Loren J.

    2014-01-01

    Soil texture has been commonly associated with the population density of Heterodera glycines (soybean cyst nematode: SCN), but such an association has been mainly described in terms of textural classes. In this study, multivariate analysis and a generalized linear modeling approach were used to elucidate the quantitative relationship of soil texture with the observed SCN population density reduction after annual corn rotation in Nebraska. Forty-five commercial production fields were sampled in 2009, 2010, and 2011 and SCN population density (eggs/100 cm3 of soil) for each field was determined before (Pi) and after (Pf) annual corn rotation from ten 3 × 3-m sampling grids. Principal components analysis revealed that, compared with silt and clay, sand had a stronger association with SCN Pi and Pf. Cluster analysis using the average linkage method and confirmed through 1,000 bootstrap simulations identified two groups: one corresponding to predominant silt-and-clay fields and other to sand-predominant fields. This grouping suggested that SCN relative percent population decline was higher in the sandy than in the silt-and-clay predominant group. However, when groups were compared for their SCN population density reduction using Pf as the response, Pi as a covariate, and incorporating the year and field variability, a negative binomial generalized linear model indicated that the SCN population density reduction was not statistically different between the sand-predominant field group and the silt-and-clay predominant group. PMID:24987160

  2. 10 CFR 960.5-2-3 - Meteorology.

    Code of Federal Regulations, 2013 CFR

    2013-01-01

    ... operation and closure will not be likely to lead to radionuclide releases to an unrestricted area greater... with higher population densities than are the average for the region. (2) History of extreme weather...

  3. 10 CFR 960.5-2-3 - Meteorology.

    Code of Federal Regulations, 2014 CFR

    2014-01-01

    ... operation and closure will not be likely to lead to radionuclide releases to an unrestricted area greater... with higher population densities than are the average for the region. (2) History of extreme weather...

  4. 10 CFR 960.5-2-3 - Meteorology.

    Code of Federal Regulations, 2012 CFR

    2012-01-01

    ... operation and closure will not be likely to lead to radionuclide releases to an unrestricted area greater... with higher population densities than are the average for the region. (2) History of extreme weather...

  5. Electron energy distribution function in the divertor region of the COMPASS tokamak during neutral beam injection heating

    NASA Astrophysics Data System (ADS)

    Hasan, E.; Dimitrova, M.; Havlicek, J.; Mitošinková, K.; Stöckel, J.; Varju, J.; Popov, Tsv K.; Komm, M.; Dejarnac, R.; Hacek, P.; Panek, R.; the COMPASS Team

    2018-02-01

    This paper presents the results from swept probe measurements in the divertor region of the COMPASS tokamak in D-shaped, L-mode discharges, with toroidal magnetic field BT = 1.15 T, plasma current Ip = 180 kA and line-average electron densities varying from 2 to 8×1019 m-3. Using neutral beam injection heating, the electron energy distribution function is studied before and during the application of the beam. The current-voltage characteristics data are processed using the first-derivative probe technique. This technique allows one to evaluate the plasma potential and the real electron energy distribution function (respectively, the electron temperatures and densities). At the low average electron density of 2×1019 m-3, the electron energy distribution function is bi-Maxwellian with a low-energy electron population with temperatures 4-6 eV and a high-energy electron group 12-25 eV. As the line-average electron density is increased, the electron temperatures decrease. At line-average electron densities above 7×1019 m-3, the electron energy distribution function is found to be Maxwellian with a temperature of 6-8.5 eV. The effect of the neutral beam injection heating power in the divertor region is also studied.

  6. Effect of deer exclusion by fencing on abundance of Amblyomma americanum (Acari: Ixodidae) on Fire Island, New York, USA

    USGS Publications Warehouse

    Ginsberg, H.S.; Butler, M.; Zhioua, E.

    2002-01-01

    The effects of deer exclusion on northern populations of lone star ticks, Amblyomma americanum, were tested at the Lighthouse Tract, Fire Island, NY, USA, where densities of this species have increased recently. Game fencing was erected to exclude deer from two sites of roughly one ha each, and populations of nymphal and adult A. americanum within were compared with those at control sites outside the exclosures. Percent control of nymphs within vs. outside the exclosures averaged 48.4% in the four years post-treatment, compared to pretreatment values. Percent control varied markedly in different years, suggesting that factors in addition to deer densities had strong effects on population densities of A. americanum. Exclosures of this size did not control adult A. americanum. Effects of deer exclusion in this recently expanded northern population of A. americanum were similar to those that have been reported for southern populations of this species.

  7. Commonly rare and rarely common: comparing population abundance of invasive and native aquatic species.

    PubMed

    Hansen, Gretchen J A; Vander Zanden, M Jake; Blum, Michael J; Clayton, Murray K; Hain, Ernie F; Hauxwell, Jennifer; Izzo, Marit; Kornis, Matthew S; McIntyre, Peter B; Mikulyuk, Alison; Nilsson, Erika; Olden, Julian D; Papeş, Monica; Sharma, Sapna

    2013-01-01

    Invasive species are leading drivers of environmental change. Their impacts are often linked to their population size, but surprisingly little is known about how frequently they achieve high abundances. A nearly universal pattern in ecology is that species are rare in most locations and abundant in a few, generating right-skewed abundance distributions. Here, we use abundance data from over 24,000 populations of 17 invasive and 104 native aquatic species to test whether invasive species differ from native counterparts in statistical patterns of abundance across multiple sites. Invasive species on average reached significantly higher densities than native species and exhibited significantly higher variance. However, invasive and native species did not differ in terms of coefficient of variation, skewness, or kurtosis. Abundance distributions of all species were highly right skewed (skewness>0), meaning both invasive and native species occurred at low densities in most locations where they were present. The average abundance of invasive and native species was 6% and 2%, respectively, of the maximum abundance observed within a taxonomic group. The biological significance of the differences between invasive and native species depends on species-specific relationships between abundance and impact. Recognition of cross-site heterogeneity in population densities brings a new dimension to invasive species management, and may help to refine optimal prevention, containment, control, and eradication strategies.

  8. Commonly Rare and Rarely Common: Comparing Population Abundance of Invasive and Native Aquatic Species

    PubMed Central

    Hansen, Gretchen J. A.; Vander Zanden, M. Jake; Blum, Michael J.; Clayton, Murray K.; Hain, Ernie F.; Hauxwell, Jennifer; Izzo, Marit; Kornis, Matthew S.; McIntyre, Peter B.; Mikulyuk, Alison; Nilsson, Erika; Olden, Julian D.; Papeş, Monica; Sharma, Sapna

    2013-01-01

    Invasive species are leading drivers of environmental change. Their impacts are often linked to their population size, but surprisingly little is known about how frequently they achieve high abundances. A nearly universal pattern in ecology is that species are rare in most locations and abundant in a few, generating right-skewed abundance distributions. Here, we use abundance data from over 24,000 populations of 17 invasive and 104 native aquatic species to test whether invasive species differ from native counterparts in statistical patterns of abundance across multiple sites. Invasive species on average reached significantly higher densities than native species and exhibited significantly higher variance. However, invasive and native species did not differ in terms of coefficient of variation, skewness, or kurtosis. Abundance distributions of all species were highly right skewed (skewness>0), meaning both invasive and native species occurred at low densities in most locations where they were present. The average abundance of invasive and native species was 6% and 2%, respectively, of the maximum abundance observed within a taxonomic group. The biological significance of the differences between invasive and native species depends on species-specific relationships between abundance and impact. Recognition of cross-site heterogeneity in population densities brings a new dimension to invasive species management, and may help to refine optimal prevention, containment, control, and eradication strategies. PMID:24194883

  9. The population density of Lymnaea columella (Say, 1817) (Mollusca, Lymnaeidae) an intermediate host of Fasciola hepatica (Linnaeus, 1758), in the Caparaó microregion, ES, Brazil.

    PubMed

    D'Almeida, S C G; Freitas, D F; Carneiro, M B; Camargo, P F; Azevedo, J C; Martins, I V F

    2016-06-01

    The aim of this study was to monitor the population density of Lymnaea columella, an intermediate host of Fasciola hepatica, in various aquatic habitats and in drinking water in the area of the Instituto Federal de Educação, Ciência e Tecnologia do Espírito Santo, on Caparaó Microregion, municipality of Alegre, state of Espírito Santo, Brazil. Monthly samplings were performed at certain points between drainage areas and drinking water in cattle and goat production systems during the years 2010 to 2013. The mean temperature, precipitation and the frequency of samples of L. columella were analysed graphically according the monthly average during the study period. A total of 2,038 molluscs were collected, 1558 of which were L. columella, that predominated in all sampled points. The highest average of specimens observed for L. columella was in the years 2010 and 2013 (51.0), and occurred decreased in 2011 (19.8). The temperature and precipitation averaged is 23.7 °C and 141 mm/year, respectively. Rainfall peak occurred in March (2011, 2013) and November (2012), during these periods the population of L. columella growth. There was no significant difference in the relationship between the specimens observed with seasons (dry-wet), thus the population of L. columella remained stable and can be found throughout the year.

  10. Impact of kalbaishakhi storms on adult mosquito populations in Kolkata (Calcutta), India.

    PubMed

    Pramanik, M K; Raut, S K

    2003-01-01

    The effect of kalbaishakhi storms on the adult mosquito populations of Kolkata (Calcutta), India was studied in 2000-2001. It is revealed that the density of mosquitoes was reduced, on average by 59.6 and 10%, during the first and second 24 h after each kalbaishakhi storms, respectively. Copyright 2002 Elsevier Science B.V.

  11. Spatial distribution of U.S. household carbon footprints reveals suburbanization undermines greenhouse gas benefits of urban population density.

    PubMed

    Jones, Christopher; Kammen, Daniel M

    2014-01-21

    Which municipalities and locations within the United States contribute the most to household greenhouse gas emissions, and what is the effect of population density and suburbanization on emissions? Using national household surveys, we developed econometric models of demand for energy, transportation, food, goods, and services that were used to derive average household carbon footprints (HCF) for U.S. zip codes, cities, counties, and metropolitan areas. We find consistently lower HCF in urban core cities (∼ 40 tCO2e) and higher carbon footprints in outlying suburbs (∼ 50 tCO2e), with a range from ∼ 25 to >80 tCO2e in the 50 largest metropolitan areas. Population density exhibits a weak but positive correlation with HCF until a density threshold is met, after which range, mean, and standard deviation of HCF decline. While population density contributes to relatively low HCF in the central cities of large metropolitan areas, the more extensive suburbanization in these regions contributes to an overall net increase in HCF compared to smaller metropolitan areas. Suburbs alone account for ∼ 50% of total U.S. HCF. Differences in the size, composition, and location of household carbon footprints suggest the need for tailoring of greenhouse gas mitigation efforts to different populations.

  12. Population density of red langurs in Sabangau tropical peat-swamp forest, Central Kalimantan, Indonesia.

    PubMed

    Ehlers Smith, David A; Ehlers Smith, Yvette C

    2013-08-01

    Because of the large-scale destruction of Borneo's rainforests on mineral soils, tropical peat-swamp forests (TPSFs) are increasingly essential for conserving remnant biodiversity, particularly in the lowlands where the majority of habitat conversion has occurred. Consequently, effective strategies for biodiversity conservation are required, which rely on accurate population density and distribution estimates as a baseline. We sought to establish the first population density estimates of the endemic red langur (Presbytis rubicunda) in Sabangau TPSF, the largest remaining contiguous lowland forest-block on Borneo. Using Distance sampling principles, we conducted line transect surveys in two of Sabangau's three principle habitat sub-classes and calculated group density at 2.52 groups km⁻² (95% CI 1.56-4.08) in the mixed-swamp forest sub-class. Based on an average recorded group size of 6.95 individuals, population density was 17.51 ind km⁻², the second highest density recorded in this species. The accessible area of the tall-interior forest, however, was too disturbed to yield density estimates representative of the entire sub-class, and P. rubicunda was absent from the low-pole forest, likely as a result of the low availability of the species' preferred foods. This absence in 30% of Sabangau's total area indicates the importance of in situ population surveys at the habitat-specific level for accurately informing conservation strategies. We highlight the conservation value of TPSFs for P. rubicunda given the high population density and large areas remaining, and recommend 1) quantifying the response of P. rubicunda to the logging and burning of its habitats; 2) surveying degraded TPSFs for viable populations, and 3) effectively delineating TPSF sub-class boundaries from remote imagery to facilitate population estimates across the wider peat landscape, given the stark contrast in densities found across the habitat sub-classes of Sabangau. © 2013 Wiley Periodicals, Inc.

  13. The Relationship between Population Density and Cancer Mortality in Taiwan

    PubMed Central

    Hsieh, Ya‐Lun

    1998-01-01

    Many investigators have examined urbanization gradients in cancer rates. The purpose of this report was to identify urban‐rural trends in cancer mortality rates (1982–1991) for municipalities in Taiwan. For this purpose, Taiwan's municipalities were classified as rural, suburban, urban, or metropolitan, using population density as an ordinal indicator of the degree of urbanization. Average annual age‐adjusted, site‐specific cancer mortality rates were calculated for both sexes within each population density group. Significant increasing trends with more urbanization were observed in mortality rates for cancers of the lung, pancreas, and kidney among both males and females, as well as male prostate cancer, and female breast and ovary cancer. In addition, this study revealed a significant rural excess for nonmelanoma skin cancer among both males and females, as well as male non‐Hodgkin's lymphoma, and cancers of the female bone, and female connective tissue. Analytic studies for sites with consistent urban‐rural trends may be fruitful in identifying the aspect of population density, or other unmeasured factors, that contribute to these trends. PMID:9617339

  14. Determinants of the Transmission Variation of Hand, Foot and Mouth Disease in China.

    PubMed

    Zhao, Jijun; Li, Xinmin

    2016-01-01

    Severe outbreaks of hand, foot and mouth disease (HFMD) have occurred in China for decades. Our understanding of the HFMD transmission process and its determinants is still limited. In this paper, factors that affect the local variation of HFMD transmission process were studied. Three classes of factors, including meteorological, demographic and public health intervention factors, were carefully selected and their effects on HFMD transmission were investigated with Pearson's correlation coefficient and multiple linear regression models. The determining factors for the variation of HFMD transmission were different for the southeastern and the northwestern regions of China. In the northwest, fadeouts occurred yearly, and the average age at infection and the fadeout were negatively correlated with the population density. In the southeast, HFMD transmission was governed by the combined effects of the birth rate, the relative humidity and the interaction of the Health System Performance and the log of the population density. When the Health System Performance was low, HFMD transmission increased with the population density, but when the Health System Performance was high, the better health performance counteracted the transmission increase due to the higher population density.

  15. Cost-effectiveness of investing in sidewalks as a means of increasing physical activity: a RESIDE modelling study

    PubMed Central

    Veerman, J Lennert; Zapata-Diomedi, Belen; Gunn, Lucy; McCormack, Gavin R; Cobiac, Linda J; Mantilla Herrera, Ana Maria; Giles-Corti, Billie; Shiell, Alan

    2016-01-01

    Background Studies consistently find that supportive neighbourhood built environments increase physical activity by encouraging walking and cycling. However, evidence on the cost-effectiveness of investing in built environment interventions as a means of promoting physical activity is lacking. In this study, we assess the cost-effectiveness of increasing sidewalk availability as one means of encouraging walking. Methods Using data from the RESIDE study in Perth, Australia, we modelled the cost impact and change in health-adjusted life years (HALYs) of installing additional sidewalks in established neighbourhoods. Estimates of the relationship between sidewalk availability and walking were taken from a previous study. Multistate life table models were used to estimate HALYs associated with changes in walking frequency and duration. Sensitivity analyses were used to explore the impact of variations in population density, discount rates, sidewalk costs and the inclusion of unrelated healthcare costs in added life years. Results Installing and maintaining an additional 10 km of sidewalk in an average neighbourhood with 19 000 adult residents was estimated to cost A$4.2 million over 30 years and gain 24 HALYs over the lifetime of an average neighbourhood adult resident population. The incremental cost-effectiveness ratio was A$176 000/HALY. However, sensitivity results indicated that increasing population densities improves cost-effectiveness. Conclusions In low-density cities such as in Australia, installing sidewalks in established neighbourhoods as a single intervention is unlikely to cost-effectively improve health. Sidewalks must be considered alongside other complementary elements of walkability, such as density, land use mix and street connectivity. Population density is particularly important because at higher densities, more residents are exposed and this improves the cost-effectiveness. Health gain is one of many benefits of enhancing neighbourhood walkability and future studies might consider a more comprehensive assessment of its social value (eg, social cohesion, safety and air quality). PMID:27650762

  16. Reduced energy density of close-up diets decrease ruminal pH and increase concentration of volatile fatty acids postpartum in Holstein cows.

    PubMed

    Huang, Wenming; Tian, Yujia; Li, Shengli; Wu, Zhaohai; Cao, Zhijun

    2017-11-01

    The objective of this study was to determine the effect of reduced energy density of close-up diets on ruminal fermentation parameters in transition cows. Fourteen Holstein dry cows were blocked and assigned randomly to three groups fed a high energy density diet (HD, 1.62 Mcal of net energy for lactation (NE L )/kg dry matter (DM)), or a middle energy density diet (MD, 1.47 Mcal NE L /kg DM), or a low energy density diet (LD, 1.30 Mcal NE L /kg DM) prepartum, and were fed the same diet postpartum. The reduced energy density diets decreased the average dry matter intake (DMI) prepartum and tended to increase the DMI postpartum. The ruminal pH of the LD group was significantly higher prepartum and lower during the first week of lactation compared with the other two groups. The reduced energy density diet depressed the average ruminal concentration of propionate and butyrate prepartum, and increased the average concentration of total volatile fatty acids (VFA) postpartum. The LD group had higher populations of Butyrivibrio fibrisolvens and Ruminococcus flavefaciens relative to HD and MD groups on 7 days in milk. In conclusion, the cows fed reduced energy density diet prepartum had higher VFA concentration, but were more susceptible to subacute ruminal acidosis postpartum. © 2017 Japanese Society of Animal Science.

  17. Density and population viability of coastal marten: a rare and geographically isolated small carnivore

    PubMed Central

    Green, David S.; Levi, Taal

    2018-01-01

    Pacific martens (Martes caurina humboldtensis) in coastal forests of Oregon and northern California in the United States are rare and geographically isolated, prompting a petition for listing under the Endangered Species Act. If listed, regulations have the potential to influence land-use decisions on public and private lands, but no estimates of population size, density, or viability of remnant marten populations are available for evaluating their conservation status. We used GPS and VHF telemetry and spatial mark-resight to estimate home ranges, density, and population size of Pacific martens in the Oregon Dunes National Recreation Area, central coast Oregon, USA. We then estimated population viability at differing levels of human-caused mortality (e.g., vehicle mortality). Marten home ranges were small on average (females = 0.8 km2, males 1.5 km2) and density (1.13 martens/1 km2) was the highest reported for North American populations (M. caurina, M. americana). We estimated 71 adult martens (95% CRI [41–87]) across two subpopulations separated by a large barrier (Umpqua River). Using population viability analysis, extinction risk for a subpopulation of 30 martens, approximately the size of the subpopulation south of the Umpqua River, ranged from 32% to 99% with two or three annual human-caused mortalities within 30 years. Absent population expansion, limiting human-caused mortalities will likely have the greatest conservation impact. PMID:29637018

  18. Population density and suicide in Scotland.

    PubMed

    Stark, Cameron; Hopkins, Paddy; Gibbs, Diane; Belbin, Alan; Hay, Alistair

    2007-01-01

    Suicide rates among men have increased in Scotland while falling in neighbouring countries. A national suicide prevention strategy has been produced. Previous work found that some rural areas of Scotland had higher than average rates of male suicide and undetermined deaths. This article describes the association between population density and suicide and undetermined death rates in Scotland. Anonymised information on deaths from suicide and undetermined cause in Scotland were obtained from the General Registrar Office for 1981-1999, including information on postcode sector. Each postcode sector was assigned a deprivation and population density score. Loglinear models were used to examine the effects of time period (grouped into four periods), deprivation quintiles, population density (grouped into four categories) and their interactions in each sex in three age groups. A significance level of 5% was used throughout. Adjusted rate ratios and 95% confidence intervals were based on models that included only significant factors and interactions. In men, there were higher rate ratios in the most densely populated and least densely populated quartiles, with intermediate rate ratios in other areas. There was no association with population density in women aged less than 25 years, a similar pattern to men in 25-44 year old women, and lower rates in rural areas in older women. Higher levels of deprivation were associated with higher rate ratios of suicide in both sexes and all age groups. Rate ratios over time increased in younger men and women, remained stable in older men, and declined in older women. Deprivation is associated with higher rates of suicide and undetermined deaths at all levels of population density and in all age groups. The highest rates of suicide among men are in the most and least densely populated areas, after adjusting for deprivation. The effect is different among women, with no effect among younger women, and lower rates among older women in areas with lower population density.

  19. Does White Clover (Trifolium repens) Abundance in Temperate Pastures Determine Sitona obsoletus (Coleoptera: Curculionidae) Larval Populations?

    PubMed

    McNeill, Mark R; van Koten, Chikako; Cave, Vanessa M; Chapman, David; Hodgson, Hamish

    2016-01-01

    To determine if host plant abundance determined the size of clover root weevil (CRW) Sitona obsoletus larval populations, a study was conducted over 4 years in plots sown in ryegrass ( Lolium perenne ) (cv. Nui) sown at either 6 or 30 kg/ha and white clover ( Trifolium repens ) sown at a uniform rate of 8 kg/ha. This provided a range of % white clover content to investigate CRW population establishment and impacts on white clover survival. Larval sampling was carried out in spring (October) when larval densities are near their spring peak at Lincoln (Canterbury, New Zealand) with % clover measured in autumn (April) and spring (September) of each year. Overall, mean larval densities measured in spring 2012-2015 were 310, 38, 59, and 31 larvae m -2 , respectively. There was a significant decline in larval populations between 2012 and 2013, but spring populations were relatively uniform thereafter. The mean % white clover measured in autumns of 2012 to 2015 was 17, 10, 3, and 11%, respectively. In comparison, mean spring % white clover from 2012 to 2015, averaged c. 5% each year. Analysis relating spring (October) larval populations to % white clover measured in each plot in autumn (April) found the 2012 larval population to be statistically significantly larger in the ryegrass 6 kg/ha plots than 30 kg/ha plots. Thereafter, sowing rate had no significant effect on larval populations. From 2013 to 2015, spring larval populations had a negative relationship with the previous autumn % white clover with the relationship highly significant for the 2014 data. When CRW larval populations in spring 2013 to 2015 were predicted from the 2013 to 2015 autumn % white clover, respectively, based on their positive relationship in 2012, the predicted densities were substantially larger than those observed. Conversely, when 2015 spring larval data and % clover was regressed against 2012-2014 larval populations, observed densities tended to be higher than predicted, but the numbers came closer to predicted for the 2013 and 2014 populations. These differences are attributed to a CRW population decline that was not accounted by % white clover changes, the CRW decline most likely due to biological control by the Braconid endoparasitoid Microctonus aethiopoides , which showed incremental increases in parasitism between 2012 and 2015, which in 2015 averaged 93%.

  20. Consequences for conservation: population density and genetic effects on reproduction of an endangered lagomorph.

    PubMed

    Demay, Stephanie M; Becker, Penny A; Waits, Lisette P; Johnson, Timothy R; Rachlow, Janet L

    2016-04-01

    Understanding reproduction and mating systems is important for managers tasked with conserving vulnerable species. Genetic tools allow biologists to investigate reproduction and mating systems with high resolution and are particularly useful for species that are otherwise difficult to study in their natural environments. We conducted parentage analyses using 19 nuclear DNA microsatellite loci to assess the influence of population density, genetic diversity, and ancestry on reproduction, and to examine the mating system of pygmy rabbits (Brachylagus idahoensis) bred in large naturalized enclosures for the reintroduction and recovery of the endangered distinct population in central Washington, USA. Reproductive output for females and males decreased as population density and individual homozygosity increased. We identified an interaction indicating that male reproductive output decreased as genetic diversity declined at high population densities, but there was no effect at low densities. Males with high amounts (> 50%) of Washington ancestry had higher reproductive output than the other ancestry groups, while reproductive output was decreased for males with high northern Utah/Wyoming ancestry and females with high Oregon/Nevada ancestry. Females and males bred with an average of 3.8 and 3.6 mates per year, respectively, and we found no evidence of positive or negative assortative mating with regards to ancestry. Multiple paternity was confirmed in 81% of litters, and we report the first documented cases of juvenile breeding by pygmy rabbits. This study demonstrates how variation in population density, genetic diversity, and ancestry impact fitness for an endangered species being bred for conservation. Our results advance understanding of basic life history characteristics for a cryptic species that is difficult to study in the wild and provide lessons for managing populations of vulnerable species in captive and free-ranging populations.

  1. Male Kirtland's Warblers' patch-level response to landscape structure during periods of varying population size and habitat amounts

    USGS Publications Warehouse

    Donner, D.M.; Ribic, C.A.; Probst, J.R.

    2009-01-01

    Forest planners must evaluate how spatiotemporal changes in habitat amount and configuration across the landscape as a result of timber management will affect species' persistence. However, there are few long-term programs available for evaluation. We investigated the response of male Kirtland's Warbler (Dendroica kirtlandii) to 26 years of changing patch and landscape structure during a large, 26-year forestry-habitat restoration program within the warbler's primary breeding range. We found that the average density of male Kirtland's Warblers was related to a different combination of patch and landscape attributes depending on the species' regional population level and habitat amounts on the landscape (early succession jack pine (Pinus banksiana) forests; 15-42% habitat cover). Specifically, patch age and habitat regeneration type were important at low male population and total habitat amounts, while patch age and distance to an occupied patch were important at relatively high population and habitat amounts. Patch age and size were more important at increasing population levels and an intermediate amount of habitat. The importance of patch age to average male density during all periods reflects the temporal buildup and decline of male numbers as habitat suitability within the patch changed with succession. Habitat selection (i.e., preference for wildfire-regenerated habitat) and availability may explain the importance of habitat type and patch size during lower population and habitat levels. The relationship between male density and distance when there was the most habitat on the landscape and the male population was large and still increasing may be explained by the widening spatial dispersion of the increasing male population at the regional scale. Because creating or preserving habitat is not a random process, management efforts would benefit from more investigations of managed population responses to changes in spatial structure that occur through habitat gain rather than habitat loss to further our empirical understanding of general principles of the fragmentation process and habitat cover threshold effects within dynamic landscapes.

  2. Average glandular dose in paired digital mammography and digital breast tomosynthesis acquisitions in a population based screening program: effects of measuring breast density, air kerma and beam quality

    NASA Astrophysics Data System (ADS)

    Helge Østerås, Bjørn; Skaane, Per; Gullien, Randi; Catrine Trægde Martinsen, Anne

    2018-02-01

    The main purpose was to compare average glandular dose (AGD) for same-compression digital mammography (DM) and digital breast tomosynthesis (DBT) acquisitions in a population based screening program, with and without breast density stratification, as determined by automatically calculated breast density (Quantra™). Secondary, to compare AGD estimates based on measured breast density, air kerma and half value layer (HVL) to DICOM metadata based estimates. AGD was estimated for 3819 women participating in the screening trial. All received craniocaudal and mediolateral oblique views of each breasts with paired DM and DBT acquisitions. Exposure parameters were extracted from DICOM metadata. Air kerma and HVL were measured for all beam qualities used to acquire the mammograms. Volumetric breast density was estimated using Quantra™. AGD was estimated using the Dance model. AGD reported directly from the DICOM metadata was also assessed. Mean AGD was 1.74 and 2.10 mGy for DM and DBT, respectively. Mean DBT/DM AGD ratio was 1.24. For fatty breasts: mean AGD was 1.74 and 2.27 mGy for DM and DBT, respectively. For dense breasts: mean AGD was 1.73 and 1.79 mGy, for DM and DBT, respectively. For breasts of similar thickness, dense breasts had higher AGD for DM and similar AGD for DBT. The DBT/DM dose ratio was substantially lower for dense compared to fatty breasts (1.08 versus 1.33). The average c-factor was 1.16. Using previously published polynomials to estimate glandularity from thickness underestimated the c-factor by 5.9% on average. Mean AGD error between estimates based on measurements (air kerma and HVL) versus DICOM header data was 3.8%, but for one mammography unit as high as 7.9%. Mean error of using the AGD value reported in the DICOM header was 10.7 and 13.3%, respectively. Thus, measurement of breast density, radiation dose and beam quality can substantially affect AGD estimates.

  3. Average glandular dose in paired digital mammography and digital breast tomosynthesis acquisitions in a population based screening program: effects of measuring breast density, air kerma and beam quality.

    PubMed

    Østerås, Bjørn Helge; Skaane, Per; Gullien, Randi; Martinsen, Anne Catrine Trægde

    2018-01-25

    The main purpose was to compare average glandular dose (AGD) for same-compression digital mammography (DM) and digital breast tomosynthesis (DBT) acquisitions in a population based screening program, with and without breast density stratification, as determined by automatically calculated breast density (Quantra ™ ). Secondary, to compare AGD estimates based on measured breast density, air kerma and half value layer (HVL) to DICOM metadata based estimates. AGD was estimated for 3819 women participating in the screening trial. All received craniocaudal and mediolateral oblique views of each breasts with paired DM and DBT acquisitions. Exposure parameters were extracted from DICOM metadata. Air kerma and HVL were measured for all beam qualities used to acquire the mammograms. Volumetric breast density was estimated using Quantra ™ . AGD was estimated using the Dance model. AGD reported directly from the DICOM metadata was also assessed. Mean AGD was 1.74 and 2.10 mGy for DM and DBT, respectively. Mean DBT/DM AGD ratio was 1.24. For fatty breasts: mean AGD was 1.74 and 2.27 mGy for DM and DBT, respectively. For dense breasts: mean AGD was 1.73 and 1.79 mGy, for DM and DBT, respectively. For breasts of similar thickness, dense breasts had higher AGD for DM and similar AGD for DBT. The DBT/DM dose ratio was substantially lower for dense compared to fatty breasts (1.08 versus 1.33). The average c-factor was 1.16. Using previously published polynomials to estimate glandularity from thickness underestimated the c-factor by 5.9% on average. Mean AGD error between estimates based on measurements (air kerma and HVL) versus DICOM header data was 3.8%, but for one mammography unit as high as 7.9%. Mean error of using the AGD value reported in the DICOM header was 10.7 and 13.3%, respectively. Thus, measurement of breast density, radiation dose and beam quality can substantially affect AGD estimates.

  4. Towards the Development of a More Accurate Monitoring Procedure for Invertebrate Populations, in the Presence of an Unknown Spatial Pattern of Population Distribution in the Field

    PubMed Central

    Petrovskaya, Natalia B.; Forbes, Emily; Petrovskii, Sergei V.; Walters, Keith F. A.

    2018-01-01

    Studies addressing many ecological problems require accurate evaluation of the total population size. In this paper, we revisit a sampling procedure used for the evaluation of the abundance of an invertebrate population from assessment data collected on a spatial grid of sampling locations. We first discuss how insufficient information about the spatial population density obtained on a coarse sampling grid may affect the accuracy of an evaluation of total population size. Such information deficit in field data can arise because of inadequate spatial resolution of the population distribution (spatially variable population density) when coarse grids are used, which is especially true when a strongly heterogeneous spatial population density is sampled. We then argue that the average trap count (the quantity routinely used to quantify abundance), if obtained from a sampling grid that is too coarse, is a random variable because of the uncertainty in sampling spatial data. Finally, we show that a probabilistic approach similar to bootstrapping techniques can be an efficient tool to quantify the uncertainty in the evaluation procedure in the presence of a spatial pattern reflecting a patchy distribution of invertebrates within the sampling grid. PMID:29495513

  5. Predicting debris

    NASA Technical Reports Server (NTRS)

    Kessler, Donald J.

    1988-01-01

    The probable amount, sizes, and relative velocities of debris are discussed, giving examples of the damage caused by debris, and focusing on the use of mathematical models to forecast the debris environment and solar activity now and in the future. Most debris are within 2,000 km of the earth's surface. The average velocity of spacecraft-debris collisions varies from 9 km/sec at 30 degrees of inclination to 13 km/sec near polar orbits. Mathematical models predict a 5 percent per year increase in the large-fragment population, producing a small-fragment population increase of 10 percent per year until the year 2060, the time of critical density. A 10 percent increase in the large population would cause the critical density to be reached around 2025.

  6. Assessing the expected response to genomic selection of individuals and families in Eucalyptus breeding with an additive-dominant model.

    PubMed

    Resende, R T; Resende, M D V; Silva, F F; Azevedo, C F; Takahashi, E K; Silva-Junior, O B; Grattapaglia, D

    2017-10-01

    We report a genomic selection (GS) study of growth and wood quality traits in an outbred F 2 hybrid Eucalyptus population (n=768) using high-density single-nucleotide polymorphism (SNP) genotyping. Going beyond previous reports in forest trees, models were developed for different selection targets, namely, families, individuals within families and individuals across the entire population using a genomic model including dominance. To provide a more breeder-intelligible assessment of the performance of GS we calculated the expected response as the percentage gain over the population average expected genetic value (EGV) for different proportions of genomically selected individuals, using a rigorous cross-validation (CV) scheme that removed relatedness between training and validation sets. Predictive abilities (PAs) were 0.40-0.57 for individual selection and 0.56-0.75 for family selection. PAs under an additive+dominance model improved predictions by 5 to 14% for growth depending on the selection target, but no improvement was seen for wood traits. The good performance of GS with no relatedness in CV suggested that our average SNP density (~25 kb) captured some short-range linkage disequilibrium. Truncation GS successfully selected individuals with an average EGV significantly higher than the population average. Response to GS on a per year basis was ~100% more efficient than by phenotypic selection and more so with higher selection intensities. These results contribute further experimental data supporting the positive prospects of GS in forest trees. Because generation times are long, traits are complex and costs of DNA genotyping are plummeting, genomic prediction has good perspectives of adoption in tree breeding practice.

  7. Manitoba

    Atmospheric Science Data Center

    2013-04-17

    ... black spruce, jack pine, tamarack, willow, and birch. Human population density is sparse, averaging about 1 person per 10 square ... Big Sand Lake, above image center, was reported to be under control on June 20, whereas the plume at lower left, to the west of Gold Sand ...

  8. Association, effects and validation of polymorphisms within the NCAPG - LCORL locus located on BTA6 with feed intake, gain, meat and carcass traits in beef cattle

    USDA-ARS?s Scientific Manuscript database

    Background: In a previously reported genome-wide association study based on a high-density bovine SNP genotyping array, 8 SNP were nominally associated (P

  9. Density, distribution, and genetic structure of grizzly bears in the Cabinet-Yaak Ecosystem

    USGS Publications Warehouse

    Macleod, Amy C.; Boyd, Kristina L.; Boulanger, John; Royle, J. Andrew; Kasworm, Wayne F.; Paetkau, David; Proctor, Michael F.; Annis, Kim; Graves, Tabitha A.

    2016-01-01

    The conservation status of the 2 threatened grizzly bear (Ursus arctos) populations in the Cabinet-Yaak Ecosystem (CYE) of northern Montana and Idaho had remained unchanged since designation in 1975; however, the current demographic status of these populations was uncertain. No rigorous data on population density and distribution or analysis of recent population genetic structure were available to measure the effectiveness of conservation efforts. We used genetic detection data from hair corral, bear rub, and opportunistic sampling in traditional and spatial capture–recapture models to generate estimates of abundance and density of grizzly bears in the CYE. We calculated mean bear residency on our sampling grid from telemetry data using Huggins and Pledger models to estimate the average number of bears present and to correct our superpopulation estimates for lack of geographic closure. Estimated grizzly bear abundance (all sex and age classes) in the CYE in 2012 was 48–50 bears, approximately half the population recovery goal. Grizzly bear density in the CYE (4.3–4.5 grizzly bears/1,000 km2) was among the lowest of interior North American populations. The sizes of the Cabinet (n = 22–24) and Yaak (n = 18–22) populations were similar. Spatial models produced similar estimates of abundance and density with comparable precision without requiring radio-telemetry data to address assumptions of geographic closure. The 2 populations in the CYE were demographically and reproductively isolated from each other and the Cabinet population was highly inbred. With parentage analysis, we documented natural migrants to the Cabinet and Yaak populations by bears born to parents in the Selkirk and Northern Continental Divide populations. These events supported data from other sources suggesting that the expansion of neighboring populations may eventually help sustain the CYE populations. However, the small size, isolation, and inbreeding documented by this study demonstrate the need for comprehensive management designed to support CYE population growth and increased connectivity and gene flow with other populations.

  10. Population and prehistory III: food-dependent demography in variable environments.

    PubMed

    Lee, Charlotte T; Puleston, Cedric O; Tuljapurkar, Shripad

    2009-11-01

    The population dynamics of preindustrial societies depend intimately on their surroundings, and food is a primary means through which environment influences population size and individual well-being. Food production requires labor; thus, dependence of survival and fertility on food involves dependence of a population's future on its current state. We use a perturbation approach to analyze the effects of random environmental variation on this nonlinear, age-structured system. We show that in expanding populations, direct environmental effects dominate induced population fluctuations, so environmental variability has little effect on mean hunger levels, although it does decrease population growth. The growth rate determines the time until population is limited by space. This limitation introduces a tradeoff between population density and well-being, so population effects become more important than the direct effects of the environment: environmental fluctuation increases mortality, releasing density dependence and raising average well-being for survivors. We discuss the social implications of these findings for the long-term fate of populations as they transition from expansion into limitation, given that conditions leading to high well-being during growth depress well-being during limitation.

  11. Indoor tanning facility density in eighty U.S. cities.

    PubMed

    Palmer, Richard C; Mayer, Joni A; Woodruff, Susan I; Eckhardt, Laura; Sallis, James F

    2002-06-01

    The purpose of this study was to examine the number of tanning facilities in select U.S. cities. The twenty most populated cities from each of 4 U.S. regions were selected for the sample. For each city, data on the number of tanning facilities, climate, and general demographic profile were collected. Data for state tanning facility legislation also were collected. A tanning facility density variable was created by dividing the city's number of facilities by its population size. The 80 cities had an average of 50 facilities each. Results of linear regression analysis indicated that higher density was significantly associated with colder climate, lower median income, and higher proportion of Whites. These data indicate that indoor tanning facilities are prevalent in the environments of U.S. urban-dwellers. Cities having the higher density profile may be logical targets for interventions promoting less or safer use of these facilities.

  12. Increasing Youths' Exposure to a Tobacco Prevention Media Campaign in Rural and Low-Population-Density Communities

    PubMed Central

    Vallone, Donna M.; Allen, Jane A.; Cullen, Jennifer; Mowery, Paul D.; Xiao, Haijun; Dorrler, Nicole; Asche, Eric T.; Healton, Cheryl

    2009-01-01

    Objectives. We examined the effectiveness of a program to increase exposure to national “truth” tobacco countermarketing messages among youths in rural and low-population-density communities. Methods. A longitudinal survey of 2618 youths aged 12 to 17 years was conducted over 5 months in 8 media markets receiving supplemental advertising and 8 comparison markets receiving less than the national average of “truth” messages. Results. Confirmed awareness of “truth” increased from 40% to 71% among youths in treatment markets while remaining stable in comparison markets. Over 35% of all youths who were unaware of the campaign at baseline became aware of it as a direct result of the increased advertising. Youths living in rural and low-population-density communities were receptive to the campaign's messages. Conclusions. Through purchase of airtime in local broadcast media, the reach of a national tobacco countermarketing campaign was expanded among youths living in rural and low-population-density areas. This strategy of augmenting delivery of nationally broadcast antitobacco ads can serve as a model for leveraging limited tobacco control resources to increase the impact of evidence-based tobacco prevention campaigns. PMID:19833994

  13. Determinants of the Transmission Variation of Hand, Foot and Mouth Disease in China

    PubMed Central

    Li, Xinmin

    2016-01-01

    Severe outbreaks of hand, foot and mouth disease (HFMD) have occurred in China for decades. Our understanding of the HFMD transmission process and its determinants is still limited. In this paper, factors that affect the local variation of HFMD transmission process were studied. Three classes of factors, including meteorological, demographic and public health intervention factors, were carefully selected and their effects on HFMD transmission were investigated with Pearson’s correlation coefficient and multiple linear regression models. The determining factors for the variation of HFMD transmission were different for the southeastern and the northwestern regions of China. In the northwest, fadeouts occurred yearly, and the average age at infection and the fadeout were negatively correlated with the population density. In the southeast, HFMD transmission was governed by the combined effects of the birth rate, the relative humidity and the interaction of the Health System Performance and the log of the population density. When the Health System Performance was low, HFMD transmission increased with the population density, but when the Health System Performance was high, the better health performance counteracted the transmission increase due to the higher population density. PMID:27701445

  14. Density estimates of monarch butterflies overwintering in central Mexico

    PubMed Central

    Diffendorfer, Jay E.; López-Hoffman, Laura; Oberhauser, Karen; Pleasants, John; Semmens, Brice X.; Semmens, Darius; Taylor, Orley R.; Wiederholt, Ruscena

    2017-01-01

    Given the rapid population decline and recent petition for listing of the monarch butterfly (Danaus plexippus L.) under the Endangered Species Act, an accurate estimate of the Eastern, migratory population size is needed. Because of difficulty in counting individual monarchs, the number of hectares occupied by monarchs in the overwintering area is commonly used as a proxy for population size, which is then multiplied by the density of individuals per hectare to estimate population size. There is, however, considerable variation in published estimates of overwintering density, ranging from 6.9–60.9 million ha−1. We develop a probability distribution for overwinter density of monarch butterflies from six published density estimates. The mean density among the mixture of the six published estimates was ∼27.9 million butterflies ha−1 (95% CI [2.4–80.7] million ha−1); the mixture distribution is approximately log-normal, and as such is better represented by the median (21.1 million butterflies ha−1). Based upon assumptions regarding the number of milkweed needed to support monarchs, the amount of milkweed (Asclepias spp.) lost (0.86 billion stems) in the northern US plus the amount of milkweed remaining (1.34 billion stems), we estimate >1.8 billion stems is needed to return monarchs to an average population size of 6 ha. Considerable uncertainty exists in this required amount of milkweed because of the considerable uncertainty occurring in overwinter density estimates. Nevertheless, the estimate is on the same order as other published estimates. The studies included in our synthesis differ substantially by year, location, method, and measures of precision. A better understanding of the factors influencing overwintering density across space and time would be valuable for increasing the precision of conservation recommendations. PMID:28462031

  15. Density estimates of monarch butterflies overwintering in central Mexico

    USGS Publications Warehouse

    Thogmartin, Wayne E.; Diffendorfer, James E.; Lopez-Hoffman, Laura; Oberhauser, Karen; Pleasants, John M.; Semmens, Brice X.; Semmens, Darius J.; Taylor, Orley R.; Wiederholt, Ruscena

    2017-01-01

    Given the rapid population decline and recent petition for listing of the monarch butterfly (Danaus plexippus L.) under the Endangered Species Act, an accurate estimate of the Eastern, migratory population size is needed. Because of difficulty in counting individual monarchs, the number of hectares occupied by monarchs in the overwintering area is commonly used as a proxy for population size, which is then multiplied by the density of individuals per hectare to estimate population size. There is, however, considerable variation in published estimates of overwintering density, ranging from 6.9–60.9 million ha−1. We develop a probability distribution for overwinter density of monarch butterflies from six published density estimates. The mean density among the mixture of the six published estimates was ∼27.9 million butterflies ha−1 (95% CI [2.4–80.7] million ha−1); the mixture distribution is approximately log-normal, and as such is better represented by the median (21.1 million butterflies ha−1). Based upon assumptions regarding the number of milkweed needed to support monarchs, the amount of milkweed (Asclepias spp.) lost (0.86 billion stems) in the northern US plus the amount of milkweed remaining (1.34 billion stems), we estimate >1.8 billion stems is needed to return monarchs to an average population size of 6 ha. Considerable uncertainty exists in this required amount of milkweed because of the considerable uncertainty occurring in overwinter density estimates. Nevertheless, the estimate is on the same order as other published estimates. The studies included in our synthesis differ substantially by year, location, method, and measures of precision. A better understanding of the factors influencing overwintering density across space and time would be valuable for increasing the precision of conservation recommendations.

  16. Effect of mechanical fragmentation of sphagnum on population density and structure of micromycete communities

    NASA Astrophysics Data System (ADS)

    Semenova, T. A.; Golovchenko, A. V.

    2017-07-01

    The population density and taxonomic structure of micromycetes were monitored for six months in a model experiment with natural and mechanically fragmented (fine and coarse) samples of sphagnum. Sphagnum fragmentation favored an increase in the number of micromycetes only during the first week of the experiment. On the average, the number of micromycetes in fine-fragmented samples was two times greater than that in the coarse-fragmented samples. The diversity of micromycetes increased in the fragmented samples of sphagnum owing to the activation of some species, which remained in the inactive state as spores in the peat before fragmentation.

  17. Disaggregating measurement uncertainty from population variability and Bayesian treatment of uncensored results

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Strom, Daniel J.; Joyce, Kevin E.; Maclellan, Jay A.

    2012-04-17

    In making low-level radioactivity measurements of populations, it is commonly observed that a substantial portion of net results are negative. Furthermore, the observed variance of the measurement results arises from a combination of measurement uncertainty and population variability. This paper presents a method for disaggregating measurement uncertainty from population variability to produce a probability density function (PDF) of possibly true results. To do this, simple, justifiable, and reasonable assumptions are made about the relationship of the measurements to the measurands (the 'true values'). The measurements are assumed to be unbiased, that is, that their average value is the average ofmore » the measurands. Using traditional estimates of each measurement's uncertainty to disaggregate population variability from measurement uncertainty, a PDF of measurands for the population is produced. Then, using Bayes's theorem, the same assumptions, and all the data from the population of individuals, a prior PDF is computed for each individual's measurand. These PDFs are non-negative, and their average is equal to the average of the measurement results for the population. The uncertainty in these Bayesian posterior PDFs is all Berkson with no remaining classical component. The methods are applied to baseline bioassay data from the Hanford site. The data include 90Sr urinalysis measurements on 128 people, 137Cs in vivo measurements on 5,337 people, and 239Pu urinalysis measurements on 3,270 people. The method produces excellent results for the 90Sr and 137Cs measurements, since there are nonzero concentrations of these global fallout radionuclides in people who have not been occupationally exposed. The method does not work for the 239Pu measurements in non-occupationally exposed people because the population average is essentially zero.« less

  18. Housing price gradient and immigrant population: Data from the Italian real estate market.

    PubMed

    Antoniucci, Valentina; Marella, Giuliano

    2018-02-01

    The database presented here was collected by Antoniucci and Marella to analyze the correlation between the housing price gradient and the immigrant population in Italy during 2016. It may also be useful in other statistical analyses, be they on the real estate market or in another branches of social science. The data sample relates to 112 Italian provincial capitals. It provides accurate information on urban structure, and specifically on urban density. The two most significant variables are original indicators constructed from official data sources: the housing price gradient, or the ratio between average prices in the center and suburbs by city; and building density, which is the average number of housing units per residential building. The housing price gradient is calculated for the two residential sub-markets, new-build and existing units, providing an original and detailed sample of the Italian residential market. Rather than average prices, the housing price gradient helps to identify potential divergences in residential market trends. As well as house prices, two other data clusters are considered: socio-economic variables, which provide a framework of each city, in terms of demographic and economic information; and various data on urban structure, which are rarely included in the same database.

  19. The role of density dependence in growth patterns of ceded territory walleye populations of northern Wisconsin: Effects of changing management regimes

    USGS Publications Warehouse

    Sass, G.G.; Hewett, S.W.; Beard, T.D.; Fayram, A.H.; Kitchell, J.F.

    2004-01-01

    We assessed density-related changes in growth of walleye Sander vitreus in the ceded territory of northern Wisconsin from 1977 to 1999. We used asymptotic length (Lz), growth rate near t0 (??), and body condition as measures of walleye growth to determine the relationship between growth and density. Among lakes, there was weak evidence of density-dependent growth: adult density explained only 0-6% of the variability in the growth metrics. Within lakes, growth was density dependent. Lz, ??, and body condition of walleyes changing with density for 69, 28, and 62% of the populations examined, respectively. Our results suggest that walleye growth was density dependent within individual lakes. However, growth was not coherently density dependent among lakes, which was possibly due to inherent differences in the productivity, surface area, forage base, landscape position, species composition, and management regime of lakes in the ceded territory. Densities of adult walleyes averaged 8.3 fish/ha and did not change significantly during 1990-1999. Mean Lz and body condition of walleyes were signilicantly higher before 1990 than after 1990, which may indicate an increase in density due to changes in management regimes. The observed growth changes do not appear to be a consequence of the statewide 15-in minimum size limit adopted in 1990 but rather a response to the treaty rights management regime. We conclude that walleye growth has the potential to predict regional-scale adult walleye densities if lake-specific variables are included in a model to account for regional-scale differences among walleye populations and lakes.

  20. Geographical Detector-Based Risk Factors Assessment of the Hand-Foot-Mouth Disease in China

    NASA Astrophysics Data System (ADS)

    Huang, J.

    2017-12-01

    Background: Hand, foot and mouth disease(HFMD) is a common infectious disease, causing thousands of deaths among children in China. This study focused on analyzing the impacts of different populations and different industry structures on HFMD incidence in China. Methods: We collected HFMD cases from 2307 counties during May 2008 in China. The potential risk factors included: monthly mean temperature, monthly mean relative humidity, monthly precipitation, different population density, different industry structures. Geographical detector technique was used to analyze the main and interactive effect of potential risk factors on HFMD incidence. Result: Using risk detector, we found the most serious HFMD incidence mainly located in the Yangtze River delta and the Pearl River delta. When the temperature was high, the incidence of HFMD was also high. This finding indicates that there is a correlation between monthly mean temperature and the incidence of HFMD. Similar analysis was undertaken to analyze the correlation between other variables and the incidence of HFMD using the risk detector. Using factor detector, we found the effect of risk factors on the incidence of HFMD, and this was ranked by PD value as follows: density of children aged 0-9 years (0.25) > tertiary industry (0.23) > GDP (0.20) >middle school student density (0.13) > relative humidity (0.12) >average temperature (0.11) >first industry (0.05). Using ecological detector, we found that child density, tertiary industry, and GDP had a strong effect on the incidence of HFMD. Using interactive detector, we found that the interactive PD value of tertiary industry and child population density was 0.42, which of GDP and tertiary industry was 0.34, that of child population density and GDP was 0.35, and that of average temperature and relative humidity was 0.28. All of these interactive PD values appeared to be higher than any PD value of sole risk factors. The combinations of the above-mentioned risk factors could effectively explain spatial variability of the incidence of HFMD in China.

  1. Cost-effectiveness of investing in sidewalks as a means of increasing physical activity: a RESIDE modelling study.

    PubMed

    Veerman, J Lennert; Zapata-Diomedi, Belen; Gunn, Lucy; McCormack, Gavin R; Cobiac, Linda J; Mantilla Herrera, Ana Maria; Giles-Corti, Billie; Shiell, Alan

    2016-09-20

    Studies consistently find that supportive neighbourhood built environments increase physical activity by encouraging walking and cycling. However, evidence on the cost-effectiveness of investing in built environment interventions as a means of promoting physical activity is lacking. In this study, we assess the cost-effectiveness of increasing sidewalk availability as one means of encouraging walking. Using data from the RESIDE study in Perth, Australia, we modelled the cost impact and change in health-adjusted life years (HALYs) of installing additional sidewalks in established neighbourhoods. Estimates of the relationship between sidewalk availability and walking were taken from a previous study. Multistate life table models were used to estimate HALYs associated with changes in walking frequency and duration. Sensitivity analyses were used to explore the impact of variations in population density, discount rates, sidewalk costs and the inclusion of unrelated healthcare costs in added life years. Installing and maintaining an additional 10 km of sidewalk in an average neighbourhood with 19 000 adult residents was estimated to cost A$4.2 million over 30 years and gain 24 HALYs over the lifetime of an average neighbourhood adult resident population. The incremental cost-effectiveness ratio was A$176 000/HALY. However, sensitivity results indicated that increasing population densities improves cost-effectiveness. In low-density cities such as in Australia, installing sidewalks in established neighbourhoods as a single intervention is unlikely to cost-effectively improve health. Sidewalks must be considered alongside other complementary elements of walkability, such as density, land use mix and street connectivity. Population density is particularly important because at higher densities, more residents are exposed and this improves the cost-effectiveness. Health gain is one of many benefits of enhancing neighbourhood walkability and future studies might consider a more comprehensive assessment of its social value (eg, social cohesion, safety and air quality). Published by the BMJ Publishing Group Limited. For permission to use (where not already granted under a licence) please go to http://www.bmj.com/company/products-services/rights-and-licensing/

  2. The ATLAS3D Project - XXX. Star formation histories and stellar population scaling relations of early-type galaxies

    NASA Astrophysics Data System (ADS)

    McDermid, Richard M.; Alatalo, Katherine; Blitz, Leo; Bournaud, Frédéric; Bureau, Martin; Cappellari, Michele; Crocker, Alison F.; Davies, Roger L.; Davis, Timothy A.; de Zeeuw, P. T.; Duc, Pierre-Alain; Emsellem, Eric; Khochfar, Sadegh; Krajnović, Davor; Kuntschner, Harald; Morganti, Raffaella; Naab, Thorsten; Oosterloo, Tom; Sarzi, Marc; Scott, Nicholas; Serra, Paolo; Weijmans, Anne-Marie; Young, Lisa M.

    2015-04-01

    We present the stellar population content of early-type galaxies from the ATLAS3D survey. Using spectra integrated within apertures covering up to one effective radius, we apply two methods: one based on measuring line-strength indices and applying single stellar population (SSP) models to derive SSP-equivalent values of stellar age, metallicity, and alpha enhancement; and one based on spectral fitting to derive non-parametric star formation histories, mass-weighted average values of age, metallicity, and half-mass formation time-scales. Using homogeneously derived effective radii and dynamically determined galaxy masses, we present the distribution of stellar population parameters on the Mass Plane (MJAM, σe, R^maj_e), showing that at fixed mass, compact early-type galaxies are on average older, more metal-rich, and more alpha-enhanced than their larger counterparts. From non-parametric star formation histories, we find that the duration of star formation is systematically more extended in lower mass objects. Assuming that our sample represents most of the stellar content of today's local Universe, approximately 50 per cent of all stars formed within the first 2 Gyr following the big bang. Most of these stars reside today in the most massive galaxies (>1010.5 M⊙), which themselves formed 90 per cent of their stars by z ˜ 2. The lower mass objects, in contrast, have formed barely half their stars in this time interval. Stellar population properties are independent of environment over two orders of magnitude in local density, varying only with galaxy mass. In the highest density regions of our volume (dominated by the Virgo cluster), galaxies are older, alpha-enhanced, and have shorter star formation histories with respect to lower density regions.

  3. Clinal patterns in genetic variation for northern leopard frog (Rana pipiens): Conservation status and population histories

    USGS Publications Warehouse

    Stockwell, Craig A.; Fisher, Justin D.L.; McLean, Kyle I.

    2016-01-01

    The security of the northern leopard frog (Rana pipiens) varies spatially with populations east and west of North Dakota considered as secure and at risk, respectively. We used genetic markers to characterize the conservation status of northern leopard frog populations across North Dakota. We used multiple regression analyses and model selection to evaluate correlations of expected heterozygosity (HE) with the direct and additive effects of: i) geographic location,ii) wetland density and iii) average annual precipitation. There was lower genetic diversity in the western portion of the state due to lower levels of diversity for populations southwest of the Missouri River. This may reflect a refugial/colonization signature for the only non-glaciated area of North Dakota. Genetic diversity was also positively associated with wetland densities which is consistent with the reliance of this species on a mosaic of wetlands. Our findings suggest that populations in the southwestern part of North Dakota are of higher conservation concern, a finding consistent with the higher risk noted for northern leopard frog populations in most states west of North Dakota. Our findings also pose the hypothesis that climate change induced changes in wetland densities will reduce genetic diversity of northern leopard frog populations.

  4. Density Shock Waves in Confined Microswimmers

    NASA Astrophysics Data System (ADS)

    Tsang, Alan Cheng Hou; Kanso, Eva

    2016-01-01

    Motile and driven particles confined in microfluidic channels exhibit interesting emergent behavior, from propagating density bands to density shock waves. A deeper understanding of the physical mechanisms responsible for these emergent structures is relevant to a number of physical and biomedical applications. Here, we study the formation of density shock waves in the context of an idealized model of microswimmers confined in a narrow channel and subject to a uniform external flow. Interestingly, these density shock waves exhibit a transition from "subsonic" with compression at the back to "supersonic" with compression at the front of the population as the intensity of the external flow increases. This behavior is the result of a nontrivial interplay between hydrodynamic interactions and geometric confinement, and it is confirmed by a novel quasilinear wave model that properly captures the dependence of the shock formation on the external flow. These findings can be used to guide the development of novel mechanisms for controlling the emergent density distribution and the average population speed, with potentially profound implications on various processes in industry and biotechnology, such as the transport and sorting of cells in flow channels.

  5. The interaction between the spatial distribution of resource patches and population density: consequences for intraspecific growth and morphology.

    PubMed

    Jacobson, Bailey; Grant, James W A; Peres-Neto, Pedro R

    2015-07-01

    How individuals within a population distribute themselves across resource patches of varying quality has been an important focus of ecological theory. The ideal free distribution predicts equal fitness amongst individuals in a 1 : 1 ratio with resources, whereas resource defence theory predicts different degrees of monopolization (fitness variance) as a function of temporal and spatial resource clumping and population density. One overlooked landscape characteristic is the spatial distribution of resource patches, altering the equitability of resource accessibility and thereby the effective number of competitors. While much work has investigated the influence of morphology on competitive ability for different resource types, less is known regarding the phenotypic characteristics conferring relative ability for a single resource type, particularly when exploitative competition predominates. Here we used young-of-the-year rainbow trout (Oncorhynchus mykiss) to test whether and how the spatial distribution of resource patches and population density interact to influence the level and variance of individual growth, as well as if functional morphology relates to competitive ability. Feeding trials were conducted within stream channels under three spatial distributions of nine resource patches (distributed, semi-clumped and clumped) at two density levels (9 and 27 individuals). Average trial growth was greater in high-density treatments with no effect of resource distribution. Within-trial growth variance had opposite patterns across resource distributions. Here, variance decreased at low-population, but increased at high-population densities as patches became increasingly clumped as the result of changes in the levels of interference vs. exploitative competition. Within-trial growth was related to both pre- and post-trial morphology where competitive individuals were those with traits associated with swimming capacity and efficiency: larger heads/bodies/caudal fins and less angled pectoral fins. The different degrees of within-population growth variance at the same density level found here, as a function of spatial resource distribution, provide an explanation for the inconsistencies in within-site growth variance and population regulation often noted with regard to density dependence in natural landscapes. © 2015 The Authors. Journal of Animal Ecology © 2015 British Ecological Society.

  6. Relationship between ancient bridges and population dynamics in the lower Yangtze River Basin, China.

    PubMed

    Zhao, Yang; Jia, Xin; Lee, Harry F; Zhao, Hongqiang; Cai, Shuliang; Huang, Xianjin

    2017-01-01

    It has been suggested that population growth dynamics may be revealed by the geographic distribution and the physical structure of ancient bridges. Yet, this relationship has not been empirically verified. In this study, we applied the archaeological records for ancient bridges to reveal the population growth dynamics in the lower Yangtze River Basin in late imperial China. We investigated 89 ancient bridges in Yixing that were built during the Ming and Qing dynasties (AD1368-1911). Global Position System information and structure (length, width, and span) of those bridges was measured during our field investigations. Their distribution density was calculated by ArcGIS. The historical socio-economic dynamics of Yixing was inferred from the distribution and structure of ancient bridges. Based on the above information, the population growth dynamics in Yixing was projected. Our results show that 77 bridges were built in Yixing during the Qing dynasty, which is 6.41 times more than the number built during the Ming dynasty. In the Ming dynasty, bridges were built on pivotal routes; in the Qing dynasty, bridges were scattered across various places. Over the period, the density distribution of bridges shifted northwestward, while the average length and width of bridges decreased. The increasing number of bridges corresponded to population growth, largely attributable to massive clan migration from northern China during the Little Ice Age. The shift in the density distribution of bridges corresponded to the formation of settlements of large clans and the blossoming of Yixing Teapot handicrafts. The scattering and the reduction in average length and width of bridges was due to the dispersal of population and the associated formation of small settlements in the latter period. Our approach is innovative and robust, and could be employed to recover long-term historical population growth dynamics in other parts of China.

  7. Relationship between ancient bridges and population dynamics in the lower Yangtze River Basin, China

    PubMed Central

    Zhao, Yang; Lee, Harry F.; Zhao, Hongqiang; Cai, Shuliang; Huang, Xianjin

    2017-01-01

    It has been suggested that population growth dynamics may be revealed by the geographic distribution and the physical structure of ancient bridges. Yet, this relationship has not been empirically verified. In this study, we applied the archaeological records for ancient bridges to reveal the population growth dynamics in the lower Yangtze River Basin in late imperial China. We investigated 89 ancient bridges in Yixing that were built during the Ming and Qing dynasties (AD1368–1911). Global Position System information and structure (length, width, and span) of those bridges was measured during our field investigations. Their distribution density was calculated by ArcGIS. The historical socio-economic dynamics of Yixing was inferred from the distribution and structure of ancient bridges. Based on the above information, the population growth dynamics in Yixing was projected. Our results show that 77 bridges were built in Yixing during the Qing dynasty, which is 6.41 times more than the number built during the Ming dynasty. In the Ming dynasty, bridges were built on pivotal routes; in the Qing dynasty, bridges were scattered across various places. Over the period, the density distribution of bridges shifted northwestward, while the average length and width of bridges decreased. The increasing number of bridges corresponded to population growth, largely attributable to massive clan migration from northern China during the Little Ice Age. The shift in the density distribution of bridges corresponded to the formation of settlements of large clans and the blossoming of Yixing Teapot handicrafts. The scattering and the reduction in average length and width of bridges was due to the dispersal of population and the associated formation of small settlements in the latter period. Our approach is innovative and robust, and could be employed to recover long-term historical population growth dynamics in other parts of China. PMID:28792976

  8. Influences on Bythotrephes longimanus life-history characteristics in the Great Lakes

    USGS Publications Warehouse

    Pothoven, Steven A.; Vanderploeg, Henry A.; Warner, David M.; Schaeffer, Jeffrey S.; Ludsin, Stuart A.; Claramunt, Randall M.; Nalepa, Thomas F.

    2012-01-01

    We compared Bythotrephes population demographics and dynamics to predator (planktivorous fish) and prey (small-bodied crustacean zooplankton) densities at a site sampled through the growing season in Lakes Michigan, Huron, and Erie. Although seasonal average densities of Bythotrephes were similar across lakes (222/m2 Erie, 247/m2 Huron, 162/m2 Michigan), temporal trends in abundance differed among lakes. In central Lake Erie where Bythotrephes' prey assemblage was dominated by small individuals (60%), where planktivorous fish densities were high (14,317/ha), and where a shallow water column limited availability of a deepwater refuge, the Bythotrephes population was characterized by a small mean body size, large broods with small neonates, allocation of length increases mainly to the spine rather than to the body, and a late summer population decline. By contrast, in Lake Michigan where Bythotrephes' prey assemblage was dominated by large individuals (72%) and planktivorous fish densities were lower (5052/ha), the Bythotrephes population was characterized by a large mean body size (i.e., 37–55% higher than in Erie), small broods with large neonates, nearly all growth in body length occurring between instars 1 and 2, and population persistence into fall. Life-history characteristics in Lake Huron tended to be intermediate to those found in Lakes Michigan and Erie, reflecting lower overall prey and predator densities (1224/ha) relative to the other lakes. Because plasticity in life history can affect interactions with other species, our findings point to the need to understand life-history variation among Great Lakes populations to improve our ability to model the dynamics of these ecosystems.

  9. Atomic structure data based on average-atom model for opacity calculations in astrophysical plasmas

    NASA Astrophysics Data System (ADS)

    Trzhaskovskaya, M. B.; Nikulin, V. K.

    2018-03-01

    Influence of the plasmas parameters on the electron structure of ions in astrophysical plasmas is studied on the basis of the average-atom model in the local thermodynamic equilibrium approximation. The relativistic Dirac-Slater method is used for the electron density estimation. The emphasis is on the investigation of an impact of the plasmas temperature and density on the ionization stages required for calculations of the plasmas opacities. The level population distributions and level energy spectra are calculated and analyzed for all ions with 6 ≤ Z ≤ 32 occurring in astrophysical plasmas. The plasma temperature range 2 - 200 eV and the density range 2 - 100 mg/cm3 are considered. The validity of the method used is supported by good agreement between our values of ionization stages for a number of ions, from oxygen up to uranium, and results obtained earlier by various methods among which are more complicated procedures.

  10. NEWLY QUENCHED GALAXIES AS THE CAUSE FOR THE APPARENT EVOLUTION IN AVERAGE SIZE OF THE POPULATION

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Carollo, C. M.; Bschorr, T. J.; Lilly, S. J.

    2013-08-20

    We use the large COSMOS sample of galaxies to study in an internally self-consistent way the change in the number densities of quenched early-type galaxies (Q-ETGs) of a given size over the redshift interval 0.2 < z < 1 in order to study the claimed size evolution of these galaxies. In a stellar mass bin at 10{sup 10.5} < M{sub galaxy} < 10{sup 11} M{sub Sun }, we see no change in the number density of compact Q-ETGs over this redshift range, while in a higher mass bin at >10{sup 11} M{sub Sun }, where we would expect merging tomore » be more significant, we find a small decrease, by {approx}30%. In both mass bins, the increase of the median sizes of Q-ETGs with time is primarily caused by the addition to the size function of larger and more diffuse Q-ETGs. At all masses, compact Q-ETGs become systematically redder toward later epochs, with a (U - V) color difference which is consistent with a passive evolution of their stellar populations, indicating that they are a stable population that does not appreciably evolve in size. We find furthermore, at all epochs, that the larger Q-ETGs (at least in the lower mass bin) have average rest-frame colors that are systematically bluer than those of the more compact Q-ETGs, suggesting that the former are indeed younger than the latter. The idea that new, large, Q-ETGs are responsible for the observed growth in the median size of the population at a given mass is also supported by analysis of the sizes and number of the star-forming galaxies that are expected to be the progenitors of the new Q-ETGs over the same period. In the low mass bin, the new Q-ETGs appear to have {approx}30% smaller half-light radii than their star-forming progenitors. This is likely due to the fading of their disks after they cease star formation. Comparison with higher redshifts shows that the median size of newly quenched galaxies roughly scales, at constant mass, as (1 + z){sup -1}. We conclude that the dominant cause of the size evolution seen in the Q-ETG population is that the average sizes and thus stellar densities of individual Q-ETGs roughly scale with the average density of the universe at the time when they were quenched, and that subsequent size changes in individual objects, through merging or other processes, are of secondary importance, especially at masses below 10{sup 11} M{sub Sun}.« less

  11. Sampling Error in Relation to Cyst Nematode Population Density Estimation in Small Field Plots.

    PubMed

    Župunski, Vesna; Jevtić, Radivoje; Jokić, Vesna Spasić; Župunski, Ljubica; Lalošević, Mirjana; Ćirić, Mihajlo; Ćurčić, Živko

    2017-06-01

    Cyst nematodes are serious plant-parasitic pests which could cause severe yield losses and extensive damage. Since there is still very little information about error of population density estimation in small field plots, this study contributes to the broad issue of population density assessment. It was shown that there was no significant difference between cyst counts of five or seven bulk samples taken per each 1-m 2 plot, if average cyst count per examined plot exceeds 75 cysts per 100 g of soil. Goodness of fit of data to probability distribution tested with χ 2 test confirmed a negative binomial distribution of cyst counts for 21 out of 23 plots. The recommended measure of sampling precision of 17% expressed through coefficient of variation ( cv ) was achieved if the plots of 1 m 2 contaminated with more than 90 cysts per 100 g of soil were sampled with 10-core bulk samples taken in five repetitions. If plots were contaminated with less than 75 cysts per 100 g of soil, 10-core bulk samples taken in seven repetitions gave cv higher than 23%. This study indicates that more attention should be paid on estimation of sampling error in experimental field plots to ensure more reliable estimation of population density of cyst nematodes.

  12. Parasite transmission in social interacting hosts: Monogenean epidemics in guppies

    USGS Publications Warehouse

    Johnson, M.B.; Lafferty, K.D.; van, Oosterhout C.; Cable, J.

    2011-01-01

    Background: Infection incidence increases with the average number of contacts between susceptible and infected individuals. Contact rates are normally assumed to increase linearly with host density. However, social species seek out each other at low density and saturate their contact rates at high densities. Although predicting epidemic behaviour requires knowing how contact rates scale with host density, few empirical studies have investigated the effect of host density. Also, most theory assumes each host has an equal probability of transmitting parasites, even though individual parasite load and infection duration can vary. To our knowledge, the relative importance of characteristics of the primary infected host vs. the susceptible population has never been tested experimentally. Methodology/Principal Findings: Here, we examine epidemics using a common ectoparasite, Gyrodactylus turnbulli infecting its guppy host (Poecilia reticulata). Hosts were maintained at different densities (3, 6, 12 and 24 fish in 40 L aquaria), and we monitored gyrodactylids both at a population and individual host level. Although parasite population size increased with host density, the probability of an epidemic did not. Epidemics were more likely when the primary infected fish had a high mean intensity and duration of infection. Epidemics only occurred if the primary infected host experienced more than 23 worm days. Female guppies contracted infections sooner than males, probably because females have a higher propensity for shoaling. Conclusions/Significance: These findings suggest that in social hosts like guppies, the frequency of social contact largely governs disease epidemics independent of host density. ?? 2011 Johnson et al.

  13. Parasite transmission in social interacting hosts: Monogenean epidemics in guppies

    USGS Publications Warehouse

    Johnson, Mirelle B.; Lafferty, Kevin D.; van Oosterhout, Cock; Cable, Joanne

    2011-01-01

    Background Infection incidence increases with the average number of contacts between susceptible and infected individuals. Contact rates are normally assumed to increase linearly with host density. However, social species seek out each other at low density and saturate their contact rates at high densities. Although predicting epidemic behaviour requires knowing how contact rates scale with host density, few empirical studies have investigated the effect of host density. Also, most theory assumes each host has an equal probability of transmitting parasites, even though individual parasite load and infection duration can vary. To our knowledge, the relative importance of characteristics of the primary infected host vs. the susceptible population has never been tested experimentally. Methodology/Principal Findings Here, we examine epidemics using a common ectoparasite, Gyrodactylus turnbulli infecting its guppy host (Poecilia reticulata). Hosts were maintained at different densities (3, 6, 12 and 24 fish in 40 L aquaria), and we monitored gyrodactylids both at a population and individual host level. Although parasite population size increased with host density, the probability of an epidemic did not. Epidemics were more likely when the primary infected fish had a high mean intensity and duration of infection. Epidemics only occurred if the primary infected host experienced more than 23 worm days. Female guppies contracted infections sooner than males, probably because females have a higher propensity for shoaling. Conclusions/Significance These findings suggest that in social hosts like guppies, the frequency of social contact largely governs disease epidemics independent of host density.

  14. Ninespine Stickleback Abundance in Lake Michigan Increases After Dreissenid Mussel Invasion

    USGS Publications Warehouse

    Madenjian, Charles P.; Bunnell, David B.; Gorman, Owen T.

    2010-01-01

    Based on data from our annual lakewide bottom trawl survey of Lake Michigan, we determined that density of ninespine sticklebacks Pungitius pungitius increased from an average of 0.234 kg/ha during 1973–1995 to an average of 1.318 kg/ha during 1996–2007. This greater-than-fivefold increase in density coincided with the dreissenid mussel invasion of Lake Michigan. Intervention analysis revealed that ninespine stickleback density in Lake Michigan significantly increased between the two time periods. In contrast, based on data from our annual bottom trawl survey of U.S. waters of Lake Superior, ninespine stickleback density decreased from an average of 0.133 kg/ha during 1978–1999 to an average of only 0.026 kg/ha during 2000–2007. This greater-than-fivefold density decrease, which was found to be significant via intervention analysis, coincided with population recovery for both lean and fat morphotypes of lake trout Salvelinus namaycush in Lake Superior. In contrast to Lake Michigan, dreissenid mussels have not invaded Lake Superior on a lakewide basis. Thus, a comparison of these two lakes indicated that the increase in ninespine stickleback abundance in Lake Michigan was most likely attributable to the dreissenid mussel invasion. In addition, based on our correlation analysis, alewives Alosa pseudoharengus did not have an adverse effect on ninespine stickleback abundance in Lake Michigan. Perhaps the recent increase in biomass of green algae Cladophora spp. associated with the dreissenid mussel invasion improved spawning habitat quality for ninespine sticklebacks and led to their stepwise abundance increase in Lake Michigan beginning in 1996

  15. Decline of a Rare Moth at Its Last Known English Site: Causes and Lessons for Conservation.

    PubMed

    Baker, David; Barrett, Sinead; Beale, Colin M; Crawford, Terry J; Ellis, Sam; Gullett, Tallulah; Mayhew, Peter J; Parsons, Mark S; Relf, Penny; Robertson, Paul; Small, Julian; Wainwright, Dave

    2016-01-01

    The conditions required by rare species are often only approximately known. Monitoring such species over time can help refine management of their protected areas. We report population trends of a rare moth, the Dark Bordered Beauty Epione vespertaria (Linnaeus, 1767) (Lepidoptera: Geometridae) at its last known English site on a protected lowland heath, and those of its host-plant, Salix repens (L.) (Malpighiales: Salicaceae). Between 2007 and 2014, adult moth density reduced by an average of 30-35% annually over the monitored area, and its range over the monitored area contracted in concert. By comparing data from before this decline (2005) with data taken in 2013, we show that the density of host-plants over the monitored area reduced three-fold overall, and ten-fold in the areas of highest host-plant density. In addition, plants were significantly smaller in 2013. In 2005, moth larvae tended to be found on plants that were significantly larger than average at the time. By 2013, far fewer plants were of an equivalent size. This suggests that the rapid decline of the moth population coincides with, and is likely driven by, changes in the host-plant population. Why the host-plant population has changed remains less certain, but fire, frost damage and grazing damage have probably contributed. It is likely that a reduction in grazing pressure in parts of the site would aid host-plant recovery, although grazing remains an important site management activity. Our work confirms the value of constant monitoring of rare or priority insect species, of the risks posed to species with few populations even when their populations are large, of the potential conflict between bespoke management for species and generic management of habitats, and hence the value of refining our knowledge of rare species' requirements so that their needs can be incorporated into the management of protected areas.

  16. Increased population density of neurosurgeons associated with decreased risk of death from motor vehicle accidents in the United States.

    PubMed

    Desai, Atman; Bekelis, Kimon; Zhao, Wenyan; Ball, Perry A

    2012-09-01

    Motor vehicle accidents (MVAs) are a leading cause of death and disability in young people. Given that a major cause of death from MVAs is traumatic brain injury, and neurosurgeons hold special expertise in this area relative to other members of a trauma team, the authors hypothesized that neurosurgeon population density would be related to reduced mortality from MVAs across US counties. The Area Resource File (2009-2010), a national health resource information database, was retrospectively analyzed. The primary outcome variable was the 3-year (2004-2006) average in MVA deaths per million population for each county. The primary independent variable was the density of neurosurgeons per million population in the year 2006. Multiple regression analysis was performed, adjusting for population density of general practitioners, urbanicity of the county, and socioeconomic status of the county. The median number of annual MVA deaths per million population, in the 3141 counties analyzed, was 226 (interquartile range [IQR] 151-323). The median number of neurosurgeons per million population was 0 (IQR 0-0), while the median number of general practitioners per million population was 274 (IQR 175-410). Using an unadjusted analysis, each increase of 1 neurosurgeon per million population was associated with 1.90 fewer MVA deaths per million population (p < 0.001). On multivariate adjusted analysis, each increase of 1 neurosurgeon per million population was associated with 1.01 fewer MVA deaths per million population (p < 0.001), with a respective decrease in MVA deaths of 0.03 per million population for an increase in 1 general practitioner (p = 0.007). Rural location, persistent poverty, and low educational level were all associated with significant increases in the rate of MVA deaths. A higher population density of neurosurgeons is associated with a significant reduction in deaths from MVAs, a major cause of death nationally. This suggests that the availability of local neurosurgeons is an important factor in the overall likelihood of survival from an MVA, and therefore indicates the importance of promoting neurosurgical education and practice throughout the country.

  17. Why is the age-standardized incidence of low-trauma fractures rising in many elderly populations?

    PubMed

    Kannus, Pekka; Niemi, Seppo; Parkkari, Jari; Palvanen, Mika; Heinonen, Ari; Sievänen, Harri; Järvinen, Teppo; Khan, Karim; Järvinen, Markku

    2002-08-01

    Low-trauma fractures of elderly people are a major public health burden worldwide, and as the number and mean age of older adults in the population continue to increase, the number of fractures is also likely to increase. Epidemiologically, however, an additional concern is that, for unknown reasons, the age-standardized incidence (average individual risk) of fracture has also risen in many populations during the recent decades. Possible reasons for this rise include a birth cohort effect, deterioration in the average bone strength by time, and increased average risk of (serious) falls. Literature provides evidence that the rise is not due to a birth cohort effect, whereas no study shows whether bone fragility has increased during this relatively short period of time. This osteoporosis hypothesis could, however, be tested if researchers would now repeat the population measurements of bone mass and density that were made in the late 1980s and the 1990s. If such studies proved that women's and men's age-standardized mean values of bone mass and density have declined over time, the osteoporosis hypothesis would receive scientific support. The third explanation is based on the hypothesis that the number and/or severity of falls has risen in elderly populations during the recent decades. Although no study has directly tested this hypothesis, a great deal of indirect epidemiologic evidence supports this contention. For example, the age-standardized incidence of fall-induced severe head injuries, bruises and contusions, and joint distortions and dislocations has increased among elderly people similarly to the low-trauma fractures. The fall hypothesis could also be tested in the coming years because the 1990s saw many research teams reporting age- and sex-specific incidences of falling for elderly populations, and the same could be done now to provide data comparing the current incidence rates of falls with the earlier ones.

  18. Density-Dependent Recruitment and Diapause in the Spring-Feeding Generation of Hemlock Woolly Adelgid (Hemiptera: Adelgidae) in Western North America.

    PubMed

    Weed, Aaron S; Elkinton, Joseph S; Lany, Nina K

    2016-12-01

    Insect populations are affected by density-dependent and density-independent factors, and knowing how these factors affect long-term population growth is critical to pest management. In this study, we experimentally manipulated densities of the hemlock woolly adelgid on eastern and western hemlock trees in the western USA to evaluate the effects of density and host species on hemlock woolly adelgid crawler colonization. We then followed development of hemlock woolly adelgid on each hemlock species. Settlement of crawlers was strongly density-dependent and consistent between host species. In addition, a period of hot days that coincided with the settlement of hemlock woolly adelgid crawlers put our experimental and naturally occurring populations into diapause during April. Diapause resulted in one generation that yr in our experimental population. Analyses of long-term air temperature records indicated that diapause-inducing temperatures in April similar to those observed in our experiment have occurred rarely since 1909 and the frequency of these events has not changed over time. Prior work suggests that hemlock woolly adelgid completes two generations per yr in the western USA with a diapause occurring in the summer. This typical life history reflects the long-term influence of regional average seasonal temperature patterns on development and the timing of diapause-inducing temperatures. However, the timing of unseasonal weather, such as the hot days observed in our experiment, occasionally changes life history trajectories from this normal pattern. Our results show that density-dependent and density-independent factors have strong effects on generational mortality and life history of hemlock woolly adelgid that are important to its population dynamics and management. Published by Oxford University Press on behalf of Entomological Society of America 2016. This work is written by US Government employees and is in the public domain in the US.

  19. Effects of genotype and population density on growth performance, carcass characteristics, and cost-benefits of broiler chickens in north central Nigeria.

    PubMed

    Yakubu, Abdulmojeed; Ayoade, John A; Dahiru, Yakubu M

    2010-04-01

    The influence of genotype and stocking densities on growth performance, carcass qualities, and cost-benefits of broilers were examined in a 28-day trial. Two hundred and seven 4-week-old birds each of Anak Titan and Arbor Acre hybrid broiler types were randomly assigned to three stocking density treatments of 8.3, 11.1, and 14.3 birds/m(2) in a 2 x 3 factorial arrangement. Final body weight, average weekly body weight and average weekly feed intake were affected (P < 0.05) by strain, with higher means recorded for Arbor Acres. However, average weekly body weight gain and feed conversion ratio were similar (P > 0.05) in both genetic groups. The effect of placement density on some growth parameters did not follow a linear trend. Arbor Acres had significantly (P < 0.05) higher relative (%) fasted body, carcass, back, neck, and wing weights compared to Anak Titans. Housing density effect (P < 0.05) was observed for relative (%) fasted body, shank, and wing weights of birds. However, the relative weights of visceral organs of birds were not significantly (P > 0.05) influenced by genotype and housing density. The economic analysis revealed that higher gross margin was recorded for Arbor Acres compared to Anak Titans (euro 2.76 versus euro 2.19; P < 0.05, respectively). Conversely, stocking rate did not exert any influence (P > 0.05) on profit margin. Genotype x stocking density interaction effect was significant for some of the carcass indices investigated. It is concluded that under sub-humid conditions of a tropical environment, the use of Arbor Acre genetic type as well as a placement density of 14.3 birds/m(2) appeared to be more profitable.

  20. Monitoring planktivorous seabird populations: Validating surface counts of crevice-nesting auklets using mark-resight techniques

    USGS Publications Warehouse

    Sheffield, L.M.; Gall, Adrian E.; Roby, D.D.; Irons, D.B.; Dugger, K.M.

    2006-01-01

    Least Auklets (Aethia pusilla (Pallas, 1811)) are the most abundant species of seabird in the Bering Sea and offer a relatively efficient means of monitoring secondary productivity in the marine environment. Counting auklets on surface plots is the primary method used to track changes in numbers of these crevice-nesters, but counts can be highly variable and may not be representative of the number of nesting individuals. We compared average maximum counts of Least Auklets on surface plots with density estimates based on mark–resight data at a colony on St. Lawrence Island, Alaska, during 2001–2004. Estimates of breeding auklet abundance from mark–resight averaged 8 times greater than those from maximum surface counts. Our results also indicate that average maximum surface counts are poor indicators of breeding auklet abundance and do not vary consistently with auklet nesting density across the breeding colony. Estimates of Least Auklet abundance from mark–resight were sufficiently precise to meet management goals for tracking changes in seabird populations. We recommend establishing multiple permanent banding plots for mark–resight studies on colonies selected for intensive long-term monitoring. Mark–resight is more likely to detect biologically significant changes in size of auklet breeding colonies than traditional surface count techniques.

  1. 2D motility tracking of Pseudomonas putida KT2440 in growth phases using video microscopy

    PubMed Central

    Davis, Michael L.; Mounteer, Leslie C.; Stevens, Lindsey K.; Miller, Charles D.; Zhou, Anhong

    2011-01-01

    Pseudomonas putida KT2440 is a gram negative motile soil bacterium important in bioremediation and biotechnology. Thus, it is important to understand its motility characteristics as individuals and in populations. Population characteristics were determined using a modified Gompertz model. Video microscopy and imaging software were utilized to analyze two dimensional (2D) bacteria movement tracks to quantify individual bacteria behavior. It was determined that inoculum density increased the lag time as seeding densities decreased, and that the maximum specific growth rate decreased as seeding densities increased. Average bacterial velocity remained relatively similar throughout exponential growth phase (~20.9 µm/sec), while maximum velocities peak early in exponential growth phase at a velocity of 51.2 µm/sec. Pseudomonas putida KT2440 also favor smaller turn angles indicating they often continue in the same direction after a change in flagella rotation throughout the exponential growth phase. PMID:21334971

  2. Modeling the impact of the indigenous microbial population on the maximum population density of Salmonella on alfalfa.

    PubMed

    Rijgersberg, Hajo; Franz, Eelco; Nierop Groot, Masja; Tromp, Seth-Oscar

    2013-07-01

    Within a microbial risk assessment framework, modeling the maximum population density (MPD) of a pathogenic microorganism is important but often not considered. This paper describes a model predicting the MPD of Salmonella on alfalfa as a function of the initial contamination level, the total count of the indigenous microbial population, the maximum pathogen growth rate and the maximum population density of the indigenous microbial population. The model is parameterized by experimental data describing growth of Salmonella on sprouting alfalfa seeds at inoculum size, native microbial load and Pseudomonas fluorescens 2-79. The obtained model fits well to the experimental data, with standard errors less than ten percent of the fitted average values. The results show that the MPD of Salmonella is not only dictated by performance characteristics of Salmonella but depends on the characteristics of the indigenous microbial population like total number of cells and its growth rate. The model can improve the predictions of microbiological growth in quantitative microbial risk assessments. Using this model, the effects of preventive measures to reduce pathogenic load and a concurrent effect on the background population can be better evaluated. If competing microorganisms are more sensitive to a particular decontamination method, a pathogenic microorganism may grow faster and reach a higher level. More knowledge regarding the effect of the indigenous microbial population (size, diversity, composition) of food products on pathogen dynamics is needed in order to make adequate predictions of pathogen dynamics on various food products.

  3. VizieR Online Data Catalog: ATLAS3D Project. XXX (McDermid+, 2015)

    NASA Astrophysics Data System (ADS)

    McDermid, R. M.; Alatalo, K.; Blitz, L.; Bournaud, F.; Bureau, M.; Cappellari, M.; Crocker, A. F.; Davies, R. L.; Davis, T. A.; De Zeeuw, P. T.; Duc, P.-A.; Emsellem, E.; Khochfar, S.; Krajnovic, D.; Kuntschner, H.; Morganti, R.; Naab, T.; Oosterloo, T.; Sarzi, M.; Scott, N.; Serra, P.; Weijmans, A.-M.; Young, L. M.

    2015-09-01

    We present the stellar population content of early-type galaxies from the ATLAS3D survey. Using spectra integrated within apertures covering up to one effective radius, we apply two methods: one based on measuring line-strength indices and applying single stellar population (SSP) models to derive SSP-equivalent values of stellar age, metallicity, and alpha enhancement; and one based on spectral fitting to derive non-parametric star formation histories, mass-weighted average values of age, metallicity, and half-mass formation time-scales. Using homogeneously derived effective radii and dynamically determined galaxy masses, we present the distribution of stellar population parameters on the Mass Plane (MJAM, σe, Rmaje), showing that at fixed mass, compact early-type galaxies are on average older, more metal-rich, and more alpha-enhanced than their larger counterparts. From non-parametric star formation histories, we find that the duration of star formation is systematically more extended in lower mass objects. Assuming that our sample represents most of the stellar content of today's local Universe, approximately 50 percent of all stars formed within the first 2Gyr following the big bang. Most of these stars reside today in the most massive galaxies (>1010.5M⊙), which themselves formed 90 percent of their stars by z~2. The lower mass objects, in contrast, have formed barely half their stars in this time interval. Stellar population properties are independent of environment over two orders of magnitude in local density, varying only with galaxy mass. In the highest density regions of our volume (dominated by the Virgo cluster), galaxies are older, alpha-enhanced, and have shorter star formation histories with respect to lower density regions. (4 data files).

  4. Population dynamics of Amur tigers (Panthera tigris altaica) in Sikhote-Alin Biosphere Zapovednik: 1966-2012.

    PubMed

    Miquelle, Dale G; Smirnov, Evgeny N; Zaumyslova, Olga Yu; Soutyrina, Svetlana V; Johnson, Douglas H

    2015-07-01

    In 2010, the world's tiger (Panthera tigris) range countries agreed to the goal of doubling tiger numbers over 12 years, but whether such an increase is biologically feasible has not been assessed. Long-term monitoring of tigers in Sikhote-Alin Biosphere Zapovednik (SABZ), Russia provided an opportunity to determine growth rates of a recovering population. A 41-year growth phase was followed by a rapid decline in tiger numbers. Annual growth rates during the growth phase averaged 4.6%, beginning near 10% in the earliest years but quickly dropping below 5%. Sex ratio (females per male) mirrored growth rates, declining as population size increased. The rapid decline from 2009 to 2012 appeared to be tied to multiple factors, including poaching, severe winters and disease. Reproductive indicators of this population are similar to those of Bengal tiger populations, suggesting that growth rates may be similar. These results suggest that, first, tiger populations likely in general grow slowly: 3-5% yearly increases are realistic and larger growth rates are likely only when populations are highly depressed, mortality rates are low and prey populations are high relative to numbers of adult females. Second, while more research is needed, it should not be assumed that tiger populations with high prey densities will necessarily grow more quickly than populations with low prey densities. Third, while growth is slow, decline can be rapid. Fourth, because declines can happen so quickly, there is a constant need to monitor populations and be ready to respond with appropriate and timely conservation interventions if tiger populations are to remain secure. Finally, an average annual growth rate across all tiger populations of 6%, required to reach the Global Tiger Initiative's goal of doubling tiger numbers in 12 years, is a noble but unlikely scenario. © 2015 International Society of Zoological Sciences, Institute of Zoology/Chinese Academy of Sciences and Wiley Publishing Asia Pty Ltd.

  5. Evaluation of Observation-Fused Regional Air Quality Model Results for Population Air Pollution Exposure Estimation

    PubMed Central

    Chen, Gang; Li, Jingyi; Ying, Qi; Sherman, Seth; Perkins, Neil; Rajeshwari, Sundaram; Mendola, Pauline

    2014-01-01

    In this study, Community Multiscale Air Quality (CMAQ) model was applied to predict ambient gaseous and particulate concentrations during 2001 to 2010 in 15 hospital referral regions (HRRs) using a 36-km horizontal resolution domain. An inverse distance weighting based method was applied to produce exposure estimates based on observation-fused regional pollutant concentration fields using the differences between observations and predictions at grid cells where air quality monitors were located. Although the raw CMAQ model is capable of producing satisfying results for O3 and PM2.5 based on EPA guidelines, using the observation data fusing technique to correct CMAQ predictions leads to significant improvement of model performance for all gaseous and particulate pollutants. Regional average concentrations were calculated using five different methods: 1) inverse distance weighting of observation data alone, 2) raw CMAQ results, 3) observation-fused CMAQ results, 4) population-averaged raw CMAQ results and 5) population-averaged fused CMAQ results. It shows that while O3 (as well as NOx) monitoring networks in the HRR regions are dense enough to provide consistent regional average exposure estimation based on monitoring data alone, PM2.5 observation sites (as well as monitors for CO, SO2, PM10 and PM2.5 components) are usually sparse and the difference between the average concentrations estimated by the inverse distance interpolated observations, raw CMAQ and fused CMAQ results can be significantly different. Population-weighted average should be used to account spatial variation in pollutant concentration and population density. Using raw CMAQ results or observations alone might lead to significant biases in health outcome analyses. PMID:24747248

  6. Population dynamics of long-tailed ducks breeding on the Yukon-Kuskokwim Delta, Alaska

    USGS Publications Warehouse

    Schamber, Jason L.; Flint, Paul L.; Grand, J. Barry; Wilson, Heather M.; Morse, Julie A.

    2009-01-01

    Population estimates for long-tailed ducks in North America have declined by nearly 50% over the past 30 years. Life history and population dynamics of this species are difficult to ascertain, because the birds nest at low densities across a broad range of habitat types. Between 1991 and 2004, we collected information on productivity and survival of long-tailed ducks at three locations on the Yukon-Kuskokwim Delta. Clutch size averaged 7.1 eggs, and nesting success averaged 30%. Duckling survival to 30 days old averaged 10% but was highly variable among years, ranging from 0% to 25%. Apparent annual survival of adult females based on mark-recapture of nesting females was estimated at 74%. We combined these estimates of survival and productivity into a matrix-based population model, which predicted an annual population decline of 19%. Elasticities indicated that population growth rate (λ) was most sensitive to changes in adult female survival. Further, the relatively high sensitivity of λ to duckling survival suggests that low duckling survival may be a bottleneck to productivity in some years. These data represent the first attempt to synthesize a population model for this species. Although our analyses were hampered by the small sample sizes inherent in studying a dispersed nesting species, our model provides a basis for management actions and can be enhanced as additional data become available.

  7. The dynamics of fish populations in the Palancar stream,a small tributary of the river Guadalquivir, Spain

    NASA Astrophysics Data System (ADS)

    Bravo, Ramón; Soriguer, Mila C.; Villar, Noelia; Hernando, José A.

    2001-02-01

    The relationship between flooding and changes in the size distribution of fish populations in the Palancar stream confirms observations in other rivers. On average, density decreased by 36.2 % and biomass increased by 14.5 %, passing from a period of severe drought to one of heavier than normal rains. Precipitation is the most important of the many factors affecting the populations of the Palancar stream; the most evident changes all occurred after the drought. During the drought period, the marked seasonal fluctuation in flow was the most important factor regulating the population dynamics. Fish density and biomass varied in proportion to the water volume. During the rainy period, the studied section of the river was found to be an important reproduction and nursery area, with juveniles and individuals of reproduction age dominating. The presence of Micropterus salmoides, an introduced piscivorous species, is another factor affecting the population dynamics in the Palancar stream. The observed absence of age 0+ individuals of the dominant populations is considered a direct effect of predation.

  8. Estimating neuronal connectivity from axonal and dendritic density fields

    PubMed Central

    van Pelt, Jaap; van Ooyen, Arjen

    2013-01-01

    Neurons innervate space by extending axonal and dendritic arborizations. When axons and dendrites come in close proximity of each other, synapses between neurons can be formed. Neurons vary greatly in their morphologies and synaptic connections with other neurons. The size and shape of the arborizations determine the way neurons innervate space. A neuron may therefore be characterized by the spatial distribution of its axonal and dendritic “mass.” A population mean “mass” density field of a particular neuron type can be obtained by averaging over the individual variations in neuron geometries. Connectivity in terms of candidate synaptic contacts between neurons can be determined directly on the basis of their arborizations but also indirectly on the basis of their density fields. To decide when a candidate synapse can be formed, we previously developed a criterion defining that axonal and dendritic line pieces should cross in 3D and have an orthogonal distance less than a threshold value. In this paper, we developed new methodology for applying this criterion to density fields. We show that estimates of the number of contacts between neuron pairs calculated from their density fields are fully consistent with the number of contacts calculated from the actual arborizations. However, the estimation of the connection probability and the expected number of contacts per connection cannot be calculated directly from density fields, because density fields do not carry anymore the correlative structure in the spatial distribution of synaptic contacts. Alternatively, these two connectivity measures can be estimated from the expected number of contacts by using empirical mapping functions. The neurons used for the validation studies were generated by our neuron simulator NETMORPH. An example is given of the estimation of average connectivity and Euclidean pre- and postsynaptic distance distributions in a network of neurons represented by their population mean density fields. PMID:24324430

  9. Methods for Adjusting U.S. Geological Survey Rural Regression Peak Discharges in an Urban Setting

    USGS Publications Warehouse

    Moglen, Glenn E.; Shivers, Dorianne E.

    2006-01-01

    A study was conducted of 78 U.S. Geological Survey gaged streams that have been subjected to varying degrees of urbanization over the last three decades. Flood-frequency analysis coupled with nonlinear regression techniques were used to generate a set of equations for converting peak discharge estimates determined from rural regression equations to a set of peak discharge estimates that represent known urbanization. Specifically, urban regression equations for the 2-, 5-, 10-, 25-, 50-, 100-, and 500-year return periods were calibrated as a function of the corresponding rural peak discharge and the percentage of impervious area in a watershed. The results of this study indicate that two sets of equations, one set based on imperviousness and one set based on population density, performed well. Both sets of equations are dependent on rural peak discharges, a measure of development (average percentage of imperviousness or average population density), and a measure of homogeneity of development within a watershed. Average imperviousness was readily determined by using geographic information system methods and commonly available land-cover data. Similarly, average population density was easily determined from census data. Thus, a key advantage to the equations developed in this study is that they do not require field measurements of watershed characteristics as did the U.S. Geological Survey urban equations developed in an earlier investigation. During this study, the U.S. Geological Survey PeakFQ program was used as an integral tool in the calibration of all equations. The scarcity of historical land-use data, however, made exclusive use of flow records necessary for the 30-year period from 1970 to 2000. Such relatively short-duration streamflow time series required a nonstandard treatment of the historical data function of the PeakFQ program in comparison to published guidelines. Thus, the approach used during this investigation does not fully comply with the guidelines set forth in U.S. Geological Survey Bulletin 17B, and modifications may be needed before it can be applied in practice.

  10. 10 CFR 960.5-2-1 - Population density and distribution.

    Code of Federal Regulations, 2010 CFR

    2010-01-01

    ... SITES FOR A NUCLEAR WASTE REPOSITORY Preclosure Guidelines Preclosure Radiological Safety § 960.5-2-1... repository operation and closure, (1) the expected average radiation dose to members of the public within any...) Disqualifying conditions. A site shall be disqualified if— (1) Any surface facility of a repository would be...

  11. The inequitable distribution of tobacco outlet density: the role of income in two Black Mid-Atlantic geopolitical areas.

    PubMed

    Fakunle, D O; Milam, A J; Furr-Holden, C D; Butler, J; Thorpe, R J; LaVeist, T A

    2016-07-01

    Studies have shown that communities with higher concentrations of low-income racial and ethnic minorities correlate with a greater presence of tobacco outlets. Community-level income has consistently been among the strongest predictors of tobacco outlet density. This study analyzes two Maryland geopolitical areas with similar racial concentrations yet differing income levels in an attempt to disentangle the race-income relationship with tobacco outlet density. In this cross-sectional examination of tobacco outlet and census tract-level sociodemographic data, Baltimore City, Maryland, and Prince George's County, Maryland, were geocoded to determine tobacco outlet density. Tobacco outlet density was defined as the mean number of tobacco outlets per 1000 persons per census tract. Comparisons of tobacco outlet density and sociodemographic variables were analysed via two-sample t-tests, and the direct effect of sociodemographic variables on tobacco outlet density for each area was analysed via spatial lag regressions. Prince George's County, the area with the higher income level ($77,190 vs $43,571), has a significantly lower tobacco outlet density than Baltimore City (P < 0.001). Prince George's County has a 67.5% Black population and an average of 3.94 tobacco outlets per 1000 persons per tract. By contrast, Baltimore City has a 65.3% Black population and an average of 7.95 tobacco outlets per 1000 persons per tract. Spatial lag regression model results indicate an inverse relationship between income and tobacco outlet density in Baltimore City and Prince George's County (β = -0.03, P < 0.01 &β = -0.01, P = 0.02, respectively), and a significant interaction term indicating a greater magnitude in the relationship between income and tobacco outlet density in Baltimore City (β = -0.05, P < 0.01). Results suggest that higher socio-economic status, even in primarily underrepresented racial and ethnic geopolitical areas, is linked to lower tobacco outlet density. Copyright © 2016 The Royal Society for Public Health. Published by Elsevier Ltd. All rights reserved.

  12. Construction of a High-Density Genetic Map from RNA-Seq Data for an Arabidopsis Bay-0 × Shahdara RIL Population

    PubMed Central

    Serin, Elise A. R.; Snoek, L. B.; Nijveen, Harm; Willems, Leo A. J.; Jiménez-Gómez, Jose M.; Hilhorst, Henk W. M.; Ligterink, Wilco

    2017-01-01

    High-density genetic maps are essential for high resolution mapping of quantitative traits. Here, we present a new genetic map for an Arabidopsis Bayreuth × Shahdara recombinant inbred line (RIL) population, built on RNA-seq data. RNA-seq analysis on 160 RILs of this population identified 30,049 single-nucleotide polymorphisms (SNPs) covering the whole genome. Based on a 100-kbp window SNP binning method, 1059 bin-markers were identified, physically anchored on the genome. The total length of the RNA-seq genetic map spans 471.70 centimorgans (cM) with an average marker distance of 0.45 cM and a maximum marker distance of 4.81 cM. This high resolution genotyping revealed new recombination breakpoints in the population. To highlight the advantages of such high-density map, we compared it to two publicly available genetic maps for the same population, comprising 69 PCR-based markers and 497 gene expression markers derived from microarray data, respectively. In this study, we show that SNP markers can effectively be derived from RNA-seq data. The new RNA-seq map closes many existing gaps in marker coverage, saturating the previously available genetic maps. Quantitative trait locus (QTL) analysis for published phenotypes using the available genetic maps showed increased QTL mapping resolution and reduced QTL confidence interval using the RNA-seq map. The new high-density map is a valuable resource that facilitates the identification of candidate genes and map-based cloning approaches. PMID:29259624

  13. Community risk indicators for dental caries in school children: an ecologic study.

    PubMed

    Amstutz, R D; Rozier, R G

    1995-06-01

    A statewide survey of NC schoolchildren found wide variation in dental caries prevalence among sampled classrooms. This study examined factors associated with this variation using classrooms as a surrogate for the larger community, in order to identify community risk indicators (CRI). In all, 172 classrooms (3400 students) in Grades K-6 were available for analysis. Initially, 56 sociodemographic, environmental, health system, and clinical factors were evaluated for their association with caries prevalence (K-3: average dfs + DMFS; 4-6: average DMFS) using univariate and bivariate analyses. Of these, 21 factors met our criteria for evaluation using WLS multivariate regression. For Grades K-3 (n = 108), population density, parental education, and coastal residence were negatively associated with caries scores, while age, and medical and dental, Medicaid expenditures were positive. For Grades 4--6 (n = 64), age and fs:dfs ratio were positively associated with caries scores while population density, population:dentist ratio, and years of natural fluoride exposure were negative. CRIs for both models, when compared to individual models, explained a substantial portion of the variation in caries prevalence, 31% for Grades K-3 and 51% for Grades 4-6. Results suggest that a risk assessment model based on community rather than individual variables is feasible and further refinement may reveal factors useful in identifying high risk communities.

  14. The effect of area size and predation on the time to extinction of prairie vole populations. simulation studies via SERDYCA: a Spatially-Explicit Individual-Based Model of Rodent Dynamics

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Kostova, T; Carlsen, T

    2003-11-21

    We present a spatially-explicit individual-based computational model of rodent dynamics, customized for the prairie vole species, M. Ochrogaster. The model is based on trophic relationships and represents important features such as territorial competition, mating behavior, density-dependent predation and dispersal out of the modeled spatial region. Vegetation growth and vole fecundity are dependent on climatic components. The results of simulations show that the model correctly predicts the overall temporal dynamics of the population density. Time-series analysis shows a very good match between the periods corresponding to the peak population density frequencies predicted by the model and the ones reported in themore » literature. The model is used to study the relation between persistence, landscape area and predation. We introduce the notions of average time to extinction (ATE) and persistence frequency to quantify persistence. While the ATE decreases with decrease of area, it is a bell-shaped function of the predation level: increasing for 'small' and decreasing for 'large' predation levels.« less

  15. The StreamCat Dataset: Accumulated Attributes for NHDPlusV2 (Version 2.1) Catchments Riparian Buffer for the Conterminous United States: 2010 US Census Housing Unit and Population Density

    EPA Pesticide Factsheets

    This dataset represents the population and housing unit density within individual, local NHDPlusV2 catchments and upstream, contributing watersheds riparian buffers based on 2010 US Census data. Densities are calculated for every block group and watershed averages are calculated for every local NHDPlusV2 catchment(see Data Sources for links to NHDPlusV2 data and Census Data). This data set is derived from The TIGER/Line Files and related database (.dbf) files for the conterminous USA. It was downloaded as Block Group-Level Census 2010 SF1 Data in File Geodatabase Format (ArcGIS version 10.0). The landscape raster (LR) was produced based on the data compiled from the questions asked of all people and about every housing unit. The (block-group population / block group area) and (block-group housing units / block group area) were summarized by local catchment and by watershed to produce local catchment-level and watershed-level metrics as a continuous data type (see Data Structure and Attribute Information for a description).

  16. Population ecology of the mallard: II. Breeding habitat conditions, size of the breeding populations, and production indices

    USGS Publications Warehouse

    Pospahala, Richard S.; Anderson, David R.; Henny, Charles J.

    1974-01-01

    This report, the second in a series on a comprehensive analysis of mallard population data, provides information on mallard breeding habitat, the size and distribution of breeding populations, and indices to production. The information in this report is primarily the result of large-scale aerial surveys conducted during May and July, 1955-73. The history of the conflict in resource utilization between agriculturalists and wildlife conservation interests in the primary waterfowl breeding grounds is reviewed. The numbers of ponds present during the breeding season and the midsummer period and the effects of precipitation and temperature on the number of ponds present are analyzed in detail. No significant cycles in precipitation were detected and it appears that precipitation is primarily influenced by substantial seasonal and random components. Annual estimates (1955-73) of the number of mallards in surveyed and unsurveyed breeding areas provided estimates of the size and geographic distribution of breeding mallards in North America. The estimated size of the mallard breeding population in North America has ranged from a high of 14.4 million in 1958 to a low of 7.1 million in 1965. Generally, the mallard breeding population began to decline after the 1958 peak until 1962, and remained below 10 million birds until 1970. The decline and subsequent low level of the mallard population between 1959 and 1969 .generally coincided with a period of poor habitat conditions on the major breeding grounds. The density of mallards was highest in the Prairie-Parkland Area with an average of nearly 19.2 birds per square mile. The proportion of the continental mallard breeding population in the Prairie-Parkland Area ranged from 30% in 1962 to a high of 600/0 in 1956. The geographic distribution of breeding mallards throughout North America was significantly related to the number of May ponds in the Prairie-Parkland Area. Estimates of midsummer habitat conditions and indices to production from the July Production Survey were studied in detail. Several indices relating to production showed marked declines from west to east in the Prairie-Parkland Area, these are: (1) density of breeding mallards (per square mile and per May pond), (2) brood density (per square mile and per July pond), (3) average brood size (all species combined), and (4) brood survival from class II to class III. An index to late nesting and renesting efforts was highest during years when midsummer water conditions were good. Production rates of many ducks breeding in North America appear to be regulated by both density-dependent and density-independent factors. Spacing of birds in the Prairie-Parkland Area appeared to be a key factor in the density-dependent regulation of the population. The spacing mechanism, in conjunction with habitat conditions, influenced some birds to overfly the primary breeding grounds into less favorable habitats to the north and northwest where the production rate may be suppressed. The production rate of waterfowl in the Prairie Parkland Area seems to be independent of density (after emigration has taken place) because the production index appears to be a linear function of the number of breeding birds in the area. Similarly, the production rate of waterfowl in northern Saskatchewan and northern Manitoba appeared to be independent of density. Production indices in these northern areas appear to be a linear function of the size of the breeding population. Thus, the density and distribution of breeding ducks is probably regulated through a spacing mechanism that is at least partially dependent on measurable environmental factors. The result is a density-dependent process operating to ultimately effect the production and production rate of breeding ducks on a continent-wide basis. Continental production, and therefore the size of the fall population, is probably partially regulated by the number of birds that are distributed north and northwest into environments less favorable for successful reproduction. Thus, spacing of the birds in the Prairie-Parkland Area and the movement of a fraction of the birds out of the prime breeding areas may be key factors in the density-dependent regulation of the total mallard population.

  17. Intraspecific competition and high food availability are associated with insular gigantism in a lizard.

    PubMed

    Pafilis, Panayiotis; Meiri, Shai; Foufopoulos, Johannes; Valakos, Efstratios

    2009-09-01

    Resource availability, competition, and predation commonly drive body size evolution. We assess the impact of high food availability and the consequent increased intraspecific competition, as expressed by tail injuries and cannibalism, on body size in Skyros wall lizards (Podarcis gaigeae). Lizard populations on islets surrounding Skyros (Aegean Sea) all have fewer predators and competitors than on Skyros but differ in the numbers of nesting seabirds. We predicted the following: (1) the presence of breeding seabirds (providing nutrients) will increase lizard population densities; (2) dense lizard populations will experience stronger intraspecific competition; and (3) such aggression, will be associated with larger average body size. We found a positive correlation between seabird and lizard densities. Cannibalism and tail injuries were considerably higher in dense populations. Increases in cannibalism and tail loss were associated with large body sizes. Adult cannibalism on juveniles may select for rapid growth, fuelled by high food abundance, setting thus the stage for the evolution of gigantism.

  18. Constraining the H2 column density distribution at z ˜ 3 from composite DLA spectra

    NASA Astrophysics Data System (ADS)

    Balashev, S. A.; Noterdaeme, P.

    2018-07-01

    We present the detection of the average H2 absorption signal in the overall population of neutral gas absorption systems at z˜ 3 using composite absorption spectra built from the Sloan Digital Sky Survey-III damped Lyman α catalogue. We present a new technique to directly measure the H2 column density distribution function f_H_2(N) from the average H2 absorption signal. Assuming a power-law column density distribution, we obtain a slope β = -1.29 ± 0.06(stat) ± 0.10 (sys) and an incidence rate of strong H2 absorptions [with N(H2) ≳ 1018 cm-2] to be 4.0 ± 0.5(stat) ± 1.0 (sys) per cent in H I absorption systems with N(H I) ≥1020 cm-2. Assuming the same inflexion point where f_H_2(N) steepens as at z = 0, we estimate that the cosmological density of H2 in the column density range log N(H_2) (cm^{-2})= 18{-}22 is {˜ } 15 per cent of the total. We find one order of magnitude higher H2 incident rate in a sub-sample of extremely strong damped Lyman α absorption systems (DLAs) [log N(H I) (cm^{-2}) ≥ 21.7], which, together with the derived shape of f_H_2(N), suggests that the typical H I-H2 transition column density in DLAs is log N(H)(cm-2) ≳ 22.3 in agreement with theoretical expectations for the average (low) metallicity of DLAs at high-z.

  19. Constraining the H2 column density distribution at z˜3 from composite DLA spectra

    NASA Astrophysics Data System (ADS)

    Balashev, S. A.; Noterdaeme, P.

    2018-04-01

    We present the detection of the average H2 absorption signal in the overall population of neutral gas absorption systems at z ˜ 3 using composite absorption spectra built from the Sloan Digital Sky Survey-III damped Lyman-α catalogue. We present a new technique to directly measure the H2 column density distribution function f_H_2(N) from the average H2 absorption signal. Assuming a power-law column density distribution, we obtain a slope β = -1.29 ± 0.06(stat) ± 0.10 (sys) and an incidence rate of strong H2 absorptions (with N(H2) ≳ 1018 cm-2) to be 4.0 ± 0.5(stat) ± 1.0 (sys) % in H I absorption systems with N(H I)≥1020 cm-2. Assuming the same inflexion point where f_H_2(N) steepens as at z = 0, we estimate that the cosmological density of H2 in the column density range log N(H_2) (cm^{-2})= 18-22 is ˜15% of the total. We find one order of magnitude higher H2 incident rate in a sub-sample of extremely strong DLAs (log N(H I) (cm^{-2}) ≥ 21.7), which, together with the the derived shape of f_H_2(N), suggests that the typical H I-H2 transition column density in DLAs is log N(H)(cm-2) ≳ 22.3 in agreement with theoretical expectations for the average (low) metallicity of DLAs at high-z.

  20. Intraspecific variation in stomatal traits, leaf traits and physiology reflects adaptation along aridity gradients in a South African shrub

    PubMed Central

    Carlson, Jane E.; Adams, Christopher A.; Holsinger, Kent E.

    2016-01-01

    Background and Aims Trait–environment relationships are commonly interpreted as evidence for local adaptation in plants. However, even when selection analyses support this interpretation, the mechanisms underlying differential benefits are often unknown. This study addresses this gap in knowledge using the broadly distributed South African shrub Protea repens. Specifically, the study examines whether broad-scale patterns of trait variation are consistent with spatial differences in selection and ecophysiology in the wild. Methods In a common garden study of plants sourced from 19 populations, associations were measured between five morphological traits and three axes describing source climates. Trait–trait and trait–environment associations were analysed in a multi-response model. Within two focal populations in the wild, selection and path analyses were used to test associations between traits, fecundity and physiological performance. Key Results Across 19 populations in a common garden, stomatal density increased with the source population’s mean annual temperature and decreased with its average amount of rainfall in midsummer. Concordantly, selection analysis in two natural populations revealed positive selection on stomatal density at the hotter, drier site, while failing to detect selection at the cooler, moister site. Dry-site plants with high stomatal density also had higher stomatal conductances, cooler leaf temperatures and higher light-saturated photosynthetic rates than those with low stomatal density, but no such relationships were present among wet-site plants. Leaf area, stomatal pore index and specific leaf area in the garden also co-varied with climate, but within-population differences were not associated with fitness in either wild population. Conclusions The parallel patterns of broad-scale variation, differences in selection and differences in trait–ecophysiology relationships suggest a mechanism for adaptive differentiation in stomatal density. Densely packed stomata may improve performance by increasing transpiration and cooling, but predominately in drier, hotter climates. This study uniquely shows context-dependent benefits of stomatal density – a trait rarely linked to local adaptation in plants. PMID:26424782

  1. Global hydrobelts: improved reporting scale for water-related issues?

    NASA Astrophysics Data System (ADS)

    Meybeck, M.; Kummu, M.; Dürr, H. H.

    2012-08-01

    Questions related to water such as its availability, water needs or stress, or management, are mapped at various resolutions at the global scale. They are reported at many scales, mostly along political or continental boundaries. As such, they ignore the fundamental heterogeneity of the hydroclimate and the natural boundaries of the river basins. Here, we describe the continental landmasses according to eight global-scale hydrobelts strictly limited by river basins, defined at a 30' (0.5°) resolution. The belts were defined and delineated, based primarily on the annual average temperature (T) and runoff (q), to maximise interbelt differences and minimise intrabelt variability. The belts were further divided into 29 hydroregions based on continental limits. This new global puzzle defines homogeneous and near-contiguous entities with similar hydrological and thermal regimes, glacial and postglacial basin histories, endorheism distribution and sensitivity to climate variations. The Mid-Latitude, Dry and Subtropical belts have northern and southern analogues and a general symmetry can be observed for T and q between them. The Boreal and Equatorial belts are unique. The hydroregions (median size 4.7 Mkm2) contrast strongly, with the average q ranging between 6 and 1393 mm yr-1 and the average T between -9.7 and +26.3 °C. Unlike the hydroclimate, the population density between the North and South belts and between the continents varies greatly, resulting in pronounced differences between the belts with analogues in both hemispheres. The population density ranges from 0.7 to 0.8 p km-2 for the North American Boreal and some Australian hydroregions to 280 p km-2 for the Asian part of the Northern Mid-Latitude belt. The combination of population densities and hydroclimate features results in very specific expressions of water-related characteristics in each of the 29 hydroregions. Our initial tests suggest that hydrobelt and hydroregion divisions are often more appropriate for water-relative global analysis and reporting than conventional continental or political divisions.

  2. Spatial climate patterns explain negligible variation in strength of compensatory density feedbacks in birds and mammals.

    PubMed

    Herrando-Pérez, Salvador; Delean, Steven; Brook, Barry W; Cassey, Phillip; Bradshaw, Corey J A

    2014-01-01

    The use of long-term population data to separate the demographic role of climate from density-modified demographic processes has become a major topic of ecological investigation over the last two decades. Although the ecological and evolutionary mechanisms that determine the strength of density feedbacks are now well understood, the degree to which climate gradients shape those processes across taxa and broad spatial scales remains unclear. Intuitively, harsh or highly variable environmental conditions should weaken compensatory density feedbacks because populations are hypothetically unable to achieve or maintain densities at which social and trophic interactions (e.g., competition, parasitism, predation, disease) might systematically reduce population growth. Here we investigate variation in the strength of compensatory density feedback, from long-term time series of abundance over 146 species of birds and mammals, in response to spatial gradients of broad-scale temperature precipitation variables covering 97 localities in 28 countries. We use information-theoretic metrics to rank phylogenetic generalized least-squares regression models that control for sample size (time-series length) and phylogenetic non-independence. Climatic factors explained < 1% of the remaining variation in density-feedback strength across species, with the highest non-control, model-averaged effect sizes related to extreme precipitation variables. We could not link our results directly to other published studies, because ecologists use contrasting responses, predictors and statistical approaches to correlate density feedback and climate--at the expense of comparability in a macroecological context. Censuses of multiple populations within a given species, and a priori knowledge of the spatial scales at which density feedbacks interact with climate, seem to be necessary to determine cross-taxa variation in this phenomenon. Despite the availability of robust modelling tools, the appropriate data have not yet been gathered for most species, meaning that we cannot yet make any robust generalisations about how demographic feedbacks interact with climate.

  3. Spatial Climate Patterns Explain Negligible Variation in Strength of Compensatory Density Feedbacks in Birds and Mammals

    PubMed Central

    Herrando-Pérez, Salvador; Delean, Steven; Brook, Barry W.; Cassey, Phillip; Bradshaw, Corey J. A.

    2014-01-01

    The use of long-term population data to separate the demographic role of climate from density-modified demographic processes has become a major topic of ecological investigation over the last two decades. Although the ecological and evolutionary mechanisms that determine the strength of density feedbacks are now well understood, the degree to which climate gradients shape those processes across taxa and broad spatial scales remains unclear. Intuitively, harsh or highly variable environmental conditions should weaken compensatory density feedbacks because populations are hypothetically unable to achieve or maintain densities at which social and trophic interactions (e.g., competition, parasitism, predation, disease) might systematically reduce population growth. Here we investigate variation in the strength of compensatory density feedback, from long-term time series of abundance over 146 species of birds and mammals, in response to spatial gradients of broad-scale temperature precipitation variables covering 97 localities in 28 countries. We use information-theoretic metrics to rank phylogenetic generalized least-squares regression models that control for sample size (time-series length) and phylogenetic non-independence. Climatic factors explained < 1% of the remaining variation in density-feedback strength across species, with the highest non-control, model-averaged effect sizes related to extreme precipitation variables. We could not link our results directly to other published studies, because ecologists use contrasting responses, predictors and statistical approaches to correlate density feedback and climate – at the expense of comparability in a macroecological context. Censuses of multiple populations within a given species, and a priori knowledge of the spatial scales at which density feedbacks interact with climate, seem to be necessary to determine cross-taxa variation in this phenomenon. Despite the availability of robust modelling tools, the appropriate data have not yet been gathered for most species, meaning that we cannot yet make any robust generalisations about how demographic feedbacks interact with climate. PMID:24618822

  4. Photographic mark-recapture analysis of local dynamics within an open population of dolphins.

    PubMed

    Fearnbach, H; Durban, J; Parsons, K; Claridge, D

    2012-07-01

    Identifying demographic changes is important for understanding population dynamics. However, this requires long-term studies of definable populations of distinct individuals, which can be particularly challenging when studying mobile cetaceans in the marine environment. We collected photo-identification data from 19 years (1992-2010) to assess the dynamics of a population of bottlenose dolphins (Tursiops truncatus) restricted to the shallow (<7 m) waters of Little Bahama Bank, northern Bahamas. This population was known to range beyond our study area, so we adopted a Bayesian mixture modeling approach to mark-recapture to identify clusters of individuals that used the area to different extents, and we specifically estimated trends in survival, recruitment, and abundance of a "resident" population with high probabilities of identification. There was a high probability (p= 0.97) of a long-term decrease in the size of this resident population from a maximum of 47 dolphins (95% highest posterior density intervals, HPDI = 29-61) in 1996 to a minimum of just 24 dolphins (95% HPDI = 14-37) in 2009, a decline of 49% (95% HPDI = approximately 5% to approximately 75%). This was driven by low per capita recruitment (average approximately 0.02) that could not compensate for relatively low apparent survival rates (average approximately 0.94). Notably, there was a significant increase in apparent mortality (approximately 5 apparent mortalities vs. approximately 2 on average) in 1999 when two intense hurricanes passed over the study area, with a high probability (p = 0.83) of a drop below the average survival probability (approximately 0.91 in 1999; approximately 0.94, on average). As such, our mark-recapture approach enabled us to make useful inference about local dynamics within an open population of bottlenose dolphins; this should be applicable to other studies challenged by sampling highly mobile individuals with heterogeneous space use.

  5. Consequences of splitting whole-genome sequencing effort over multiple breeds on imputation accuracy.

    PubMed

    Bouwman, Aniek C; Veerkamp, Roel F

    2014-10-03

    The aim of this study was to determine the consequences of splitting sequencing effort over multiple breeds for imputation accuracy from a high-density SNP chip towards whole-genome sequence. Such information would assist for instance numerical smaller cattle breeds, but also pig and chicken breeders, who have to choose wisely how to spend their sequencing efforts over all the breeds or lines they evaluate. Sequence data from cattle breeds was used, because there are currently relatively many individuals from several breeds sequenced within the 1,000 Bull Genomes project. The advantage of whole-genome sequence data is that it carries the causal mutations, but the question is whether it is possible to impute the causal variants accurately. This study therefore focussed on imputation accuracy of variants with low minor allele frequency and breed specific variants. Imputation accuracy was assessed for chromosome 1 and 29 as the correlation between observed and imputed genotypes. For chromosome 1, the average imputation accuracy was 0.70 with a reference population of 20 Holstein, and increased to 0.83 when the reference population was increased by including 3 other dairy breeds with 20 animals each. When the same amount of animals from the Holstein breed were added the accuracy improved to 0.88, while adding the 3 other breeds to the reference population of 80 Holstein improved the average imputation accuracy marginally to 0.89. For chromosome 29, the average imputation accuracy was lower. Some variants benefitted from the inclusion of other breeds in the reference population, initially determined by the MAF of the variant in each breed, but even Holstein specific variants did gain imputation accuracy from the multi-breed reference population. This study shows that splitting sequencing effort over multiple breeds and combining the reference populations is a good strategy for imputation from high-density SNP panels towards whole-genome sequence when reference populations are small and sequencing effort is limiting. When sequencing effort is limiting and interest lays in multiple breeds or lines this provides imputation of each breed.

  6. Results of the IMO Video Meteor Network - April 2016

    NASA Astrophysics Data System (ADS)

    Molau, S.; Crivello, S.; Goncalves, R.; Saraiva, C.; Stomeo, E.; Kac, J.

    2016-10-01

    In 2016 April, a total of 78 video cameras of the IMO Video Meteor Network recorded more than 16 000 meteors in almost 7 700 hours of observing time. The flux density profile of the Lyrids 2016 is presented and compared to the average for the years 2011-2015. The flux density increased significantly as twilight set in on the morning of 2016 April 22. A similar increase was also seen in 2012. The population index of the Lyrids is also derived from observations around the shower maximum.

  7. Population ecology of polar bears in Davis Strait, Canada and Greenland

    USGS Publications Warehouse

    Peacock, Elizabeth; Taylor, Mitchell K.; Laake, Jeffrey L.; Stirling, Ian

    2013-01-01

    Until recently, the sea ice habitat of polar bears was understood to be variable, but environmental variability was considered to be cyclic or random, rather than progressive. Harvested populations were believed to be at levels where density effects were considered not significant. However, because we now understand that polar bear demography can also be influenced by progressive change in the environment, and some populations have increased to greater densities than historically lower numbers, a broader suite of factors should be considered in demographic studies and management. We analyzed 35 years of capture and harvest data from the polar bear (Ursus maritimus) subpopulation in Davis Strait, including data from a new study (2005–2007), to quantify its current demography. We estimated the population size in 2007 to be 2,158 ± 180 (SE), a likely increase from the 1970s. We detected variation in survival, reproductive rates, and age-structure of polar bears from geographic sub-regions. Survival and reproduction of bears in southern Davis Strait was greater than in the north and tied to a concurrent dramatic increase in breeding harp seals (Pagophilus groenlandicus) in Labrador. The most supported survival models contained geographic and temporal variables. Harp seal abundance was significantly related to polar bear survival. Our estimates of declining harvest recovery rate, and increasing total survival, suggest that the rate of harvest declined over time. Low recruitment rates, average adult survival rates, and high population density, in an environment of high prey density, but deteriorating and variable ice conditions, currently characterize the Davis Strait polar bears. Low reproductive rates may reflect negative effects of greater densities or worsening ice conditions.

  8. Spectral Index Properties of millijansky Radio Sources in ATLAS

    NASA Astrophysics Data System (ADS)

    Randall, Kate; Hopkins, A. M.; Norris, R. P.; Zinn, P.; Middelberg, E.; Mao, M. Y.; Sharp, R. G.

    2012-01-01

    At the faintest radio flux densities (S1.4GHz < 10 milliJansky (mJy)), the spectral index properties of radio sources are not well constrained. The bright radio source population (S1.4GHz > 10 mJy) is well studied and is predominantly comprised of AGN. At fainter flux densities, particularly into the microJansky regime, star-forming galaxies begin to dominate the radio source population. Understanding these faint radio source populations is essential for understanding galaxy evolution, and the link between AGN and star formation. Conflicting results have recently arisen regarding whether there is a flattening of the average spectral index between a low radio frequency (325 or 610 MHz) and 1.4 GHz at these faint flux densities. To explore this issue, we have investigated the spectral index properties of a new catalogue of 843 MHz radio sources in the ELAIS-S1 (the European Large Area ISO Survey - South 1 Region) field. Our results support previous work showing a tendency towards flatter radio spectra at fainter flux densities. This catalogue is cross-matched to the Australia Telescope Large Area Survey (ATLAS), the widest deep radio survey to date at 1.4 GHz, with complementary 2.3 GHz, optical and infrared Spitzer Wide-area Infra-Red Extragalactic data. The variation of spectral index properties have been explored as a function of redshift, luminosity and flux density. [These new measurements have been used to identify a population of faint Compact Steep Spectrum sources, thought to be one of the earliest stages of the AGN life-cycle. Exploring this population will aid us in understanding the evolution of AGN as a whole.

  9. Compensation and additivity of anthropogenic mortality: life-history effects and review of methods.

    PubMed

    Péron, Guillaume

    2013-03-01

    Demographic compensation, the increase in average individual performance following a perturbation that reduces population size, and, its opposite, demographic overadditivity (or superadditivity) are central processes in both population ecology and wildlife management. A continuum of population responses to changes in cause-specific mortality exists, of which additivity and complete compensation constitute particular points. The position of a population on that continuum influences its ability to sustain exploitation and predation. Here I describe a method for quantifying where a population is on the continuum. Based on variance-covariance formulae, I describe a simple metric for the rate of compensation-additivity. I synthesize the results from 10 wildlife capture-recapture monitoring programmes from the literature and online databases, reviewing current statistical methods and the treatment of common sources of bias. These results are used to test hypotheses regarding the effects of life-history strategy, population density, average cause-specific mortality and age class on the rate of compensation-additivity. This comparative analysis highlights that long-lived species compensate less than short-lived species and that populations below their carrying capacity compensate less than those above. © 2012 The Authors. Journal of Animal Ecology © 2012 British Ecological Society.

  10. State Education Finance and Governance Profile: Mississippi

    ERIC Educational Resources Information Center

    Poulin, Nicole S.

    2010-01-01

    This article presents the state education finance and governance profile of Mississippi. Mississippians compose 0.95% of the total U.S. population, and the average density of the state is 60.7 people per square mile. In terms of education finance, the property tax is the sole form of local revenue for public education in Mississippi. In 1997, the…

  11. 78 FR 77153 - Environmental Action Statement Screening Form and Proposed Amendment to the Candidate...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2013-12-20

    ... by domestic livestock, oil and gas development, wind energy development, loss of native rangelands to... birds protected on enrolled lands. If, on average, LEPC densities are about two birds per square mile in.../implementation area of this CCAA and an overall population of about 3,125 birds within the entire planning area...

  12. Specific Gravity Variation in a Lower Mississippi Valley Cottonwood Population

    Treesearch

    R. E. Farmer; J. R. Wilcox

    1966-01-01

    Specific gravity varied from 0,32 to 0.46, averaging 0.38. Most of the variation was associated with individual trees; samples within locations accounted for a smaller, but statistically significant, portion of the variation. Variation between locatians was not significant. It was concluded that individual high-density trees' should be sought throughout the...

  13. Mapping of Vegetation with the Geoinformation System and Determining of Carrying Capacity of the Pre-Urals Steppe area for a Newly Establishing Population of the Przewalski Horse Equus ferus przewalskii at the Orenburg State Nature Reserve

    NASA Astrophysics Data System (ADS)

    Fedorov, N. I.; Mikhailenko, O. I.; Zharkikh, T. L.; Bakirova, R. T.

    2018-01-01

    Mapping of the vegetation (1:25000) of the Pre-Urals Steppe area at the Orenburg State Nature Reserve was completed in 2016. A map created with the geoinformation system contains 1931 simple and complex polygons for 25 types of vegetation. In a drought year, the average stock of palatable vegetation of the whole area is estimated at 8380 tons dry weight. The estimation is based on the size of areas covered by different types of vegetation, their grass production, the correction coefficients for decreasing of pasture forage stocks in winter and decreasing of production of grass communities in dry years. Based on pasture forage stocks the area could tolerate the maximum population size of 1769 individuals of the Przewalski horse, their average density could be 0.11 horse per ha. Yet, as watering places for animals are limited in Pre-Urals Steppe, grazing pressures on the vegetation next to the water sources may increase in dry years. That is why the above-mentioned calculated maximum population size and density must be reduced at least by half until some additional watering places are established and monitoring of the grazing effect on the vegetation next to the places is carried out regularly. Thus, the maximum size of the population is estimated at 800 to 900 individuals, which is almost 1.5 times more than necessary to establish a self-sustained population of the Przewalski horse.

  14. A case-control study to assess the impact of mammographic density on breast cancer risk in women aged 40-49 at intermediate familial risk.

    PubMed

    Assi, Valentina; Massat, Nathalie J; Thomas, Susan; MacKay, James; Warwick, Jane; Kataoka, Masako; Warsi, Iqbal; Brentnall, Adam; Warren, Ruth; Duffy, Stephen W

    2015-05-15

    Mammographic density is a strong risk factor for breast cancer, but its potential application in risk management is not clear, partly due to uncertainties about its interaction with other breast cancer risk factors. We aimed to quantify the impact of mammographic density on breast cancer risk in women aged 40-49 at intermediate familial risk of breast cancer (average lifetime risk of 23%), in particular in premenopausal women, and to investigate its relationship with other breast cancer risk factors in this population. We present the results from a case-control study nested with the FH01 cohort study of 6,710 women mostly aged 40-49 at intermediate familial risk of breast cancer. One hundred and three cases of breast cancer were age-matched to one or two controls. Density was measured by semiautomated interactive thresholding. Absolute density, but not percent density, was a significant risk factor for breast cancer in this population after adjusting for area of nondense tissue (OR per 10 cm(2) = 1.07, 95% CI 1.00-1.15, p = 0.04). The effect was stronger in premenopausal women, who made up the majority of the study population. Absolute density remained a significant predictor of breast cancer risk after adjusting for age at menarche, age at first live birth, parity, past or present hormone replacement therapy, and the Tyrer-Cuzick 10-year relative risk estimate of breast cancer. Absolute density can improve breast cancer risk stratification and delineation of high-risk groups alongside the Tyrer-Cuzick 10-year relative risk estimate. © 2014 UICC.

  15. Intraspecific variation in stomatal traits, leaf traits and physiology reflects adaptation along aridity gradients in a South African shrub.

    PubMed

    Carlson, Jane E; Adams, Christopher A; Holsinger, Kent E

    2016-01-01

    Trait-environment relationships are commonly interpreted as evidence for local adaptation in plants. However, even when selection analyses support this interpretation, the mechanisms underlying differential benefits are often unknown. This study addresses this gap in knowledge using the broadly distributed South African shrub Protea repens. Specifically, the study examines whether broad-scale patterns of trait variation are consistent with spatial differences in selection and ecophysiology in the wild. In a common garden study of plants sourced from 19 populations, associations were measured between five morphological traits and three axes describing source climates. Trait-trait and trait-environment associations were analysed in a multi-response model. Within two focal populations in the wild, selection and path analyses were used to test associations between traits, fecundity and physiological performance. Across 19 populations in a common garden, stomatal density increased with the source population's mean annual temperature and decreased with its average amount of rainfall in midsummer. Concordantly, selection analysis in two natural populations revealed positive selection on stomatal density at the hotter, drier site, while failing to detect selection at the cooler, moister site. Dry-site plants with high stomatal density also had higher stomatal conductances, cooler leaf temperatures and higher light-saturated photosynthetic rates than those with low stomatal density, but no such relationships were present among wet-site plants. Leaf area, stomatal pore index and specific leaf area in the garden also co-varied with climate, but within-population differences were not associated with fitness in either wild population. The parallel patterns of broad-scale variation, differences in selection and differences in trait-ecophysiology relationships suggest a mechanism for adaptive differentiation in stomatal density. Densely packed stomata may improve performance by increasing transpiration and cooling, but predominately in drier, hotter climates. This study uniquely shows context-dependent benefits of stomatal density--a trait rarely linked to local adaptation in plants. © The Author 2015. Published by Oxford University Press on behalf of the Annals of Botany Company. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

  16. Butterfly Density and Behaviour in Uncut Hay Meadow Strips: Behavioural Ecological Consequences of an Agri-Environmental Scheme.

    PubMed

    Lebeau, Julie; Wesselingh, Renate A; Van Dyck, Hans

    2015-01-01

    Sparing zones from mowing has been proposed, and applied, to improve local conditions for survival and reproduction of insects in hay meadows. However, little is known about the efficiency of refuge zones and the consequences for local populations. We studied population densities of butterflies before and after mowing in the refuge zone of 15 meadows in 2009 and 2011. We also studied the behaviour of the meadow brown (Maniola jurtina) comparing nectar use, interactions and flights in the refuge zone before and after mowing. Densities of grassland butterflies in this zone doubled on average after mowing. The density of females of M. jurtina increased on average fourfold, while males showed a more modest increase. In line with the idea of increased scramble competition in the refuge zone after mowing, M. jurtina increased the time spent on nectar feeding, the preferred nectar source was visited more frequently, and females made more use of non-preferred nectar sources. Maniola jurtina did not interact more with conspecifics after mowing, but interactions lasted longer. Flight tracks did not change in linearity, but were faster and shorter after mowing. After mowing, only a part of the local grassland butterflies moved to the uncut refuge zone. The resulting concentration effect alters the time allocated to different activities, nectar use and movements. These aspects have been largely ignored for agri-environmental schemes and grassland management in nature reserves and raise questions about optimal quantities and quality of uncut refuge sites for efficient conservation of grassland arthropods in agricultural landscapes.

  17. A cost constraint alone has adverse effects on food selection and nutrient density: an analysis of human diets by linear programming.

    PubMed

    Darmon, Nicole; Ferguson, Elaine L; Briend, André

    2002-12-01

    Economic constraints may contribute to the unhealthy food choices observed among low socioeconomic groups in industrialized countries. The objective of the present study was to predict the food choices a rational individual would make to reduce his or her food budget, while retaining a diet as close as possible to the average population diet. Isoenergetic diets were modeled by linear programming. To ensure these diets were consistent with habitual food consumption patterns, departure from the average French diet was minimized and constraints that limited portion size and the amount of energy from food groups were introduced into the models. A cost constraint was introduced and progressively strengthened to assess the effect of cost on the selection of foods by the program. Strengthening the cost constraint reduced the proportion of energy contributed by fruits and vegetables, meat and dairy products and increased the proportion from cereals, sweets and added fats, a pattern similar to that observed among low socioeconomic groups. This decreased the nutritional quality of modeled diets, notably the lowest cost linear programming diets had lower vitamin C and beta-carotene densities than the mean French adult diet (i.e., <25% and 10% of the mean density, respectively). These results indicate that a simple cost constraint can decrease the nutrient densities of diets and influence food selection in ways that reproduce the food intake patterns observed among low socioeconomic groups. They suggest that economic measures will be needed to effectively improve the nutritional quality of diets consumed by these populations.

  18. Butterfly Density and Behaviour in Uncut Hay Meadow Strips: Behavioural Ecological Consequences of an Agri-Environmental Scheme

    PubMed Central

    Lebeau, Julie; Wesselingh, Renate A.; Van Dyck, Hans

    2015-01-01

    Sparing zones from mowing has been proposed, and applied, to improve local conditions for survival and reproduction of insects in hay meadows. However, little is known about the efficiency of refuge zones and the consequences for local populations. We studied population densities of butterflies before and after mowing in the refuge zone of 15 meadows in 2009 and 2011. We also studied the behaviour of the meadow brown (Maniola jurtina) comparing nectar use, interactions and flights in the refuge zone before and after mowing. Densities of grassland butterflies in this zone doubled on average after mowing. The density of females of M. jurtina increased on average fourfold, while males showed a more modest increase. In line with the idea of increased scramble competition in the refuge zone after mowing, M. jurtina increased the time spent on nectar feeding, the preferred nectar source was visited more frequently, and females made more use of non-preferred nectar sources. Maniola jurtina did not interact more with conspecifics after mowing, but interactions lasted longer. Flight tracks did not change in linearity, but were faster and shorter after mowing. After mowing, only a part of the local grassland butterflies moved to the uncut refuge zone. The resulting concentration effect alters the time allocated to different activities, nectar use and movements. These aspects have been largely ignored for agri-environmental schemes and grassland management in nature reserves and raise questions about optimal quantities and quality of uncut refuge sites for efficient conservation of grassland arthropods in agricultural landscapes. PMID:26284618

  19. Double Exposure and the Climate Gap: Changing demographics and extreme heat in Ciudad Juárez, Mexico

    PubMed Central

    Collins, Timothy W.; McDonald, Yolanda J.; Aldouri, Raed; Aboargob, Faraj; Eldeb, Abdelatif; Aguilar, María de Lourdes Romo; Velázquez-Angulo, Juárez Gilberto

    2013-01-01

    Scholars have recognized a climate gap, wherein poor communities face disproportionate impacts of climate change. Others have noted that climate change and economic globalization may mutually affect a region or social group, leading to double exposure. This paper investigates how current and changing patterns of neighborhood demographics are associated with extreme heat in the border city of Juárez, Mexico. Many Juárez neighborhoods are at-risk to triple exposures, in which residents suffer due to the conjoined effects of the global recession, drug war violence, and extreme heat. Due to impacts of the recession on maquiladora employment and the explosion of drug violence (since 2008), over 75% of neighborhoods experienced decreasing population density between 2000 and 2010 and the average neighborhood saw a 40% increase in the proportion of older adults. Neighborhoods with greater drops in population density and increases in the proportion of older residents over the decade are at significantly higher risk to extreme heat, as are neighborhoods with lower population density and lower levels of education. In this context, triple exposures are associated with a climate gap that most endangers lower socioeconomic status and increasingly older aged populations remaining in neighborhoods from which high proportions of residents have departed. PMID:25642135

  20. Double Exposure and the Climate Gap: Changing demographics and extreme heat in Ciudad Juárez, Mexico.

    PubMed

    Grineski, Sara E; Collins, Timothy W; McDonald, Yolanda J; Aldouri, Raed; Aboargob, Faraj; Eldeb, Abdelatif; Aguilar, María de Lourdes Romo; Velázquez-Angulo, Juárez Gilberto

    2015-02-01

    Scholars have recognized a climate gap, wherein poor communities face disproportionate impacts of climate change. Others have noted that climate change and economic globalization may mutually affect a region or social group, leading to double exposure. This paper investigates how current and changing patterns of neighborhood demographics are associated with extreme heat in the border city of Juárez, Mexico. Many Juárez neighborhoods are at-risk to triple exposures, in which residents suffer due to the conjoined effects of the global recession, drug war violence, and extreme heat. Due to impacts of the recession on maquiladora employment and the explosion of drug violence (since 2008), over 75% of neighborhoods experienced decreasing population density between 2000 and 2010 and the average neighborhood saw a 40% increase in the proportion of older adults. Neighborhoods with greater drops in population density and increases in the proportion of older residents over the decade are at significantly higher risk to extreme heat, as are neighborhoods with lower population density and lower levels of education. In this context, triple exposures are associated with a climate gap that most endangers lower socioeconomic status and increasingly older aged populations remaining in neighborhoods from which high proportions of residents have departed.

  1. Will altered climate affect a discrete population of the sea urchin Strongylocentrotus droebachiensis?

    NASA Astrophysics Data System (ADS)

    Nyhagen, Fredrik Oulie; Christie, Hartvig; Norderhaug, Kjell Magnus

    2018-02-01

    The Green sea urchin Strongylocentrotus droebachiensis is a cold-water species that in the 70s and 80s was the main culprit in the major grazing of macrophytes in the Norwegian fjords and coastal areas. Recent reports have questioned a decline in the S. droebachiensis in the Oslofjord, similar to what was observed to occur in the Norwegian coastal population of S. droebachiensis after the 1990s. In surveys from 2013 we collected data on urchin density, test diameter, early urchin settlement and Gonad Index and created a framework in order to investigate if the current Inner Oslofjord populations of S. droebachiensis are different from surveys conducted in 1979 and 1992. This study found significant differences in density, diameter and settlement over time. Highest density in 1992 was at 10 and 15 m, while being at 20 m in 2013. Urchin diameter at the station with most urchins was on average 5 cm in 1992 compared to 3 cm in 2013. Settlement differed both between the number of settlers between 1992 and 2013, and season for highest settlement (summer in 1992, and autumn in 2013). The inner Oslofjord sea urchin data indicate a still healthy population, although environmental data (as temperature and salinity) have changed to less favorable conditions for this species.

  2. Density of surgeons is significantly associated with reduced risk of deaths from motor vehicle crashes in US counties.

    PubMed

    Chang, David C; Eastman, Brent; Talamini, Mark A; Osen, Hayley B; Tran Cao, Hop S; Coimbra, Raul

    2011-05-01

    The concept of surgery and public health has been introduced in recent years, highlighting the impact of surgeons on improving public health outcomes, a relationship that has traditionally been ascribed to general practitioners. The purpose of this study is to quantify the effect of surgeon availability on deaths from motor vehicle crashes (MVC). Retrospective analysis of the Area Resource File from 2006 was performed. The primary outcome variable was the three-year (2001-2003) average in MVC deaths per 1 million population for each county. The primary independent variable was the density of surgeons per 1 million population in year 2003. Multiple linear regression analysis was performed, adjusting for density of general practitioners, urbanicity of the county, and socioeconomic status of the county. A total of 3,225 counties were analyzed. The median number of MVC deaths per million population was 226 (IQR 158-320). The median number of surgeon per million population was 55 (IQR 0-105), while the median number of general practitioners per million population was 424 (IQR 274-620). On unadjusted analysis, each increase of one surgeon per million population was associated with 0.38 fewer MVC deaths per million population (p < 0.001). On multivariate analysis, each increase of one surgeon per million population was significantly associated with 0.16 fewer MVC deaths per million population (p < 0.001). Rural location, persistent poverty, and low educational level were all associated with significant increases in MVC deaths. Higher density of surgeons is associated with significant reduction in deaths from MVCs. This highlights the need for 1) consideration of surgery as primary care and 2) development of inclusive surgical systems designed to provide care commensurate with patients' degree of injury. Copyright © 2011 American College of Surgeons. Published by Elsevier Inc. All rights reserved.

  3. A Comparison of Grizzly Bear Demographic Parameters Estimated from Non-Spatial and Spatial Open Population Capture-Recapture Models.

    PubMed

    Whittington, Jesse; Sawaya, Michael A

    2015-01-01

    Capture-recapture studies are frequently used to monitor the status and trends of wildlife populations. Detection histories from individual animals are used to estimate probability of detection and abundance or density. The accuracy of abundance and density estimates depends on the ability to model factors affecting detection probability. Non-spatial capture-recapture models have recently evolved into spatial capture-recapture models that directly include the effect of distances between an animal's home range centre and trap locations on detection probability. Most studies comparing non-spatial and spatial capture-recapture biases focussed on single year models and no studies have compared the accuracy of demographic parameter estimates from open population models. We applied open population non-spatial and spatial capture-recapture models to three years of grizzly bear DNA-based data from Banff National Park and simulated data sets. The two models produced similar estimates of grizzly bear apparent survival, per capita recruitment, and population growth rates but the spatial capture-recapture models had better fit. Simulations showed that spatial capture-recapture models produced more accurate parameter estimates with better credible interval coverage than non-spatial capture-recapture models. Non-spatial capture-recapture models produced negatively biased estimates of apparent survival and positively biased estimates of per capita recruitment. The spatial capture-recapture grizzly bear population growth rates and 95% highest posterior density averaged across the three years were 0.925 (0.786-1.071) for females, 0.844 (0.703-0.975) for males, and 0.882 (0.779-0.981) for females and males combined. The non-spatial capture-recapture population growth rates were 0.894 (0.758-1.024) for females, 0.825 (0.700-0.948) for males, and 0.863 (0.771-0.957) for both sexes. The combination of low densities, low reproductive rates, and predominantly negative population growth rates suggest that Banff National Park's population of grizzly bears requires continued conservation-oriented management actions.

  4. Evidence for density-dependent changes in growth, downstream movement, and size of Chinook salmon subyearlings in a large-river landscape

    USGS Publications Warehouse

    Connor, William P.; Tiffan, Kenneth F.; Plumb, John M.; Moffit, Christine M.

    2013-01-01

    We studied the growth rate, downstream movement, and size of naturally produced fall Chinook Salmon Oncorhynchus tshawytscha subyearlings (age 0) for 20 years in an 8th-order river landscape with regulated riverine upstream rearing areas and an impounded downstream migration corridor. The population transitioned from low to high abundance in association with U.S. Endangered Species Act and other federally mandated recovery efforts. The mean growth rate of parr in the river did not decline with increasing abundance, but during the period of higher abundance the timing of dispersal from riverine habitat into the reservoir averaged 17 d earlier and the average size at the time of downstream dispersal was smaller by 10 mm and 1.8 g. Changes in apparent abundance, measured by catch per unit effort, largely explained the time of dispersal, measured by median day of capture, in riverine habitat. The growth rate of smolts in the reservoir declined from an average of 0.6 to 0.2 g/d between the abundance periods because the reduction in size at reservoir entry was accompanied by a tendency to migrate rather than linger and by increasing concentrations of smolts in the reservoir. The median date of passage through the reservoir was 14 d earlier on average, and average smolt size was smaller by 38 mm and 22.0 g, in accordance with density-dependent behavioral changes reflected by decreased smolt growth. Unexpectedly, smolts during the high-abundance period had begun to reexpress the migration timing and size phenotypes observed before the river was impounded, when abundance was relatively high. Our findings provide evidence for density-dependent phenotypic change in a large river that was influenced by the expansion of a recovery program. Thus, this study shows that efforts to recover native fishes can have detectable effects in large-river landscapes. The outcome of such phenotypic change, which will be an important area of future research, can only be fully judged by examining the effect of the change on population viability and productivity.

  5. Bird population density estimated from acoustic signals

    USGS Publications Warehouse

    Dawson, D.K.; Efford, M.G.

    2009-01-01

    Many animal species are detected primarily by sound. Although songs, calls and other sounds are often used for population assessment, as in bird point counts and hydrophone surveys of cetaceans, there are few rigorous methods for estimating population density from acoustic data. 2. The problem has several parts - distinguishing individuals, adjusting for individuals that are missed, and adjusting for the area sampled. Spatially explicit capture-recapture (SECR) is a statistical methodology that addresses jointly the second and third parts of the problem. We have extended SECR to use uncalibrated information from acoustic signals on the distance to each source. 3. We applied this extension of SECR to data from an acoustic survey of ovenbird Seiurus aurocapilla density in an eastern US deciduous forest with multiple four-microphone arrays. We modelled average power from spectrograms of ovenbird songs measured within a window of 0??7 s duration and frequencies between 4200 and 5200 Hz. 4. The resulting estimates of the density of singing males (0??19 ha -1 SE 0??03 ha-1) were consistent with estimates of the adult male population density from mist-netting (0??36 ha-1 SE 0??12 ha-1). The fitted model predicts sound attenuation of 0??11 dB m-1 (SE 0??01 dB m-1) in excess of losses from spherical spreading. 5.Synthesis and applications. Our method for estimating animal population density from acoustic signals fills a gap in the census methods available for visually cryptic but vocal taxa, including many species of bird and cetacean. The necessary equipment is simple and readily available; as few as two microphones may provide adequate estimates, given spatial replication. The method requires that individuals detected at the same place are acoustically distinguishable and all individuals vocalize during the recording interval, or that the per capita rate of vocalization is known. We believe these requirements can be met, with suitable field methods, for a significant number of songbird species. ?? 2009 British Ecological Society.

  6. How Often Do Thermally Excited 630.0 nm Emissions Occur in the Polar Ionosphere?

    NASA Astrophysics Data System (ADS)

    Kwagala, Norah Kaggwa; Oksavik, Kjellmar; Lorentzen, Dag A.; Johnsen, Magnar G.

    2018-01-01

    This paper studies thermally excited emissions in the polar ionosphere derived from European Incoherent Scatter Svalbard radar measurements from the years 2000-2015. The peak occurrence is found around magnetic noon, where the radar observations show cusp-like characteristics. The ionospheric, interplanetary magnetic field and solar wind conditions favor dayside magnetic reconnection as the dominant driving process. The thermal emissions occur 10 times more frequently on the dayside than on the nightside, with an average intensity of 1-5 kR. For typical electron densities in the polar ionosphere (2 × 1011 m-3), we find the peak occurrence rate to occur for extreme electron temperatures (>3000 K), which is consistent with assumptions in literature. However, for extreme electron densities (>5 × 1011 m-3), we can now report on a completely new population of thermal emissions that may occur at much lower electron temperatures (˜2300 K). The empirical atmospheric model (NRLMSISE-00) suggests that the latter population is associated with enhanced neutral atomic oxygen densities.

  7. High-levels of microplastic pollution in a large, remote, mountain lake.

    PubMed

    Free, Christopher M; Jensen, Olaf P; Mason, Sherri A; Eriksen, Marcus; Williamson, Nicholas J; Boldgiv, Bazartseren

    2014-08-15

    Despite the large and growing literature on microplastics in the ocean, little information exists on microplastics in freshwater systems. This study is the first to evaluate the abundance, distribution, and composition of pelagic microplastic pollution in a large, remote, mountain lake. We quantified pelagic microplastics and shoreline anthropogenic debris in Lake Hovsgol, Mongolia. With an average microplastic density of 20,264 particles km(-2), Lake Hovsgol is more heavily polluted with microplastics than the more developed Lakes Huron and Superior in the Laurentian Great Lakes. Fragments and films were the most abundant microplastic types; no plastic microbeads and few pellets were observed. Household plastics dominated the shoreline debris and were comprised largely of plastic bottles, fishing gear, and bags. Microplastic density decreased with distance from the southwestern shore, the most populated and accessible section of the park, and was distributed by the prevailing winds. These results demonstrate that without proper waste management, low-density populations can heavily pollute freshwater systems with consumer plastics. Copyright © 2014 Elsevier Ltd. All rights reserved.

  8. Contingency and statistical laws in replicate microbial closed ecosystems.

    PubMed

    Hekstra, Doeke R; Leibler, Stanislas

    2012-05-25

    Contingency, the persistent influence of past random events, pervades biology. To what extent, then, is each course of ecological or evolutionary dynamics unique, and to what extent are these dynamics subject to a common statistical structure? Addressing this question requires replicate measurements to search for emergent statistical laws. We establish a readily replicated microbial closed ecosystem (CES), sustaining its three species for years. We precisely measure the local population density of each species in many CES replicates, started from the same initial conditions and kept under constant light and temperature. The covariation among replicates of the three species densities acquires a stable structure, which could be decomposed into discrete eigenvectors, or "ecomodes." The largest ecomode dominates population density fluctuations around the replicate-average dynamics. These fluctuations follow simple power laws consistent with a geometric random walk. Thus, variability in ecological dynamics can be studied with CES replicates and described by simple statistical laws. Copyright © 2012 Elsevier Inc. All rights reserved.

  9. 2D motility tracking of Pseudomonas putida KT2440 in growth phases using video microscopy.

    PubMed

    Davis, Michael L; Mounteer, Leslie C; Stevens, Lindsey K; Miller, Charles D; Zhou, Anhong

    2011-05-01

    Pseudomonas putida KT2440 is a gram negative motile soil bacterium important in bioremediation and biotechnology. Thus, it is important to understand its motility characteristics as individuals and in populations. Population characteristics were determined using a modified Gompertz model. Video microscopy and imaging software were utilized to analyze two dimensional (2D) bacteria movement tracks to quantify individual bacteria behavior. It was determined that inoculum density increased the lag time as seeding densities decreased, and that the maximum specific growth rate decreased as seeding densities increased. Average bacterial velocity remained relatively similar throughout the exponential growth phase (~20.9 μm/s), while maximum velocities peak early in the exponential growth phase at a velocity of 51.2 μm/s. P. putida KT2440 also favors smaller turn angles indicating that they often continue in the same direction after a change in flagella rotation throughout the exponential growth phase. Copyright © 2011 The Society for Biotechnology, Japan. Published by Elsevier B.V. All rights reserved.

  10. Modelling moose–forest interactions under different predation scenarios at Isle Royale National Park, USA

    USGS Publications Warehouse

    DeJager, Nathan R.; Rohweder, Jason; Miranda, Brian R.; Sturtevant, Brian R.; Fox, Timothy J.; Romanski, Mark C.

    2017-01-01

    Loss of top predators may contribute to high ungulate population densities and chronic over-browsing of forest ecosystems. However, spatial and temporal variability in the strength of interactions between predators and ungulates occurs over scales that are much shorter than the scales over which forest communities change, making it difficult to characterize trophic cascades in forest ecosystems. We applied the LANDIS-II forest succession model and a recently developed ungulate browsing extension to model how the moose population could interact with the forest ecosystem of Isle Royale National Park, USA, under three different wolf predation scenarios. We contrasted a 100-yr future without wolves (no predation) with two predation scenarios (weak, long-term average predation rates and strong, higher than average rates). Increasing predation rates led to lower peak moose population densities, lower biomass removal rates, and higher estimates of forage availability and landscape carrying capacity, especially during the first 40 yr of simulations. Thereafter, moose population density was similar for all predation scenarios, but available forage biomass and the carrying capacity of the landscape continued to diverge among predation scenarios. Changes in total aboveground live biomass and species composition were most pronounced in the no predation and weak predation scenarios. Consistent with smaller-scale studies, high browsing rates led to reductions in the biomass of heavily browsed Populus tremuloides, Betula papyrifera, and Abies balsamea, and increases in the biomass of unbrowsed Picea glauca and Picea mariana, especially after the simulation year 2050, when existing boreal hardwood stands at Isle Royale are projected to senesce. As a consequence, lower predation rates corresponded with a landscape that progressively shifted toward dominance by Picea glauca and Picea mariana, and lacking available forage biomass. Consistencies with previously documented small-scale successional shifts, and population estimates and trends that approximate those from this and other boreal forests that support moose provide some confidence that these dynamics represent a trophic cascade and therefore provide an important baseline against which to evaluate long-term and large-scale effects of alternative predator management strategies on ungulate populations and forest succession.

  11. High charge state carbon and oxygen ions in Earth's equatorial quasi-trapping region

    NASA Technical Reports Server (NTRS)

    Christon, S. P.; Hamilton, D. C.; Gloeckler, G.; Eastmann, T. E.

    1994-01-01

    Observations of energetic (1.5 - 300 keV/e) medium-to-high charge state (+3 less than or equal to Q less than or equal to +7) solar wind origin C and O ions made in the quasi-trapping region (QTR) of Earth's magnetosphere are compared to ion trajectories calculated in model equatorial magnetospheric magnetic and electric fields. These comparisons indicate that solar wind ions entering the QTR on the nightside as an energetic component of the plasma sheet exit the region on the dayside, experiencing little or no charge exchange on the way. Measurements made by the CHarge Energy Mass (CHEM) ion spectrometer on board the Active Magnetospheric Particle Tracer Explorer/Charge Composition Explorer (AMPTE/CCE) spacecraft at 7 less than L less than 9 from September 1984 to January 1989 are the source of the new results contained herein: quantitative long-term determination of number densities, average energies, energy spectra, local time distributions, and their variation with geomagnetic disturbance level as indexed by Kp. Solar wind primaries (ions with charge states unchanged) and their secondaries (ions with generally lower charge states produced from primaries in the magnetosphere via charge exchange)are observed throughout the QTR and have distinctly different local time variations that persist over the entire 4-year analysis interval. During Kp larger than or equal to 3 deg intervals, primary ion (e.g., O(+6)) densities exhibit a pronounced predawn maximum with average energy minimum and a broad near-local-noon density minimum with average energy maximum. Secondary ion (e.g., O(+5)) densities do not have an identifiable predawn peak, rather they have a broad dayside maximum peaked in local morning and a nightside minimum. During Kp less than or equal to 2(-) intervals, primary ion density peaks are less intense, broader in local time extent, and centered near midnight, while secondary ion density local time variations diminish. The long-time-interval baseline helps to refine and extend previous observations; for example, we show that ionospheric contribution to O(+3)) is negligible. Through comparison with model ion trajectories, we interpret the lack of pronounced secondary ion density peaks colocated with the primary density peaks to indicate that: (1) negligible charge exchange occurs at L greater than 7, that is, solar wind secondaries are produced at L less than 7, and (2) solar wind secondaries do not form a significant portion of the plasma sheet population injected into the QTR. We conclude that little of the energetic solar wind secondary ion population is recirculated through the magnetosphere.

  12. THE EVOLUTION OF THE STELLAR MASS FUNCTIONS OF STAR-FORMING AND QUIESCENT GALAXIES TO z = 4 FROM THE COSMOS/UltraVISTA SURVEY

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Muzzin, Adam; Franx, Marijn; Labbé, Ivo

    2013-11-01

    We present measurements of the stellar mass functions (SMFs) of star-forming and quiescent galaxies to z = 4 using a sample of 95,675 K{sub s} -selected galaxies in the COSMOS/UltraVISTA field. The SMFs of the combined population are in good agreement with previous measurements and show that the stellar mass density of the universe was only 50%, 10%, and 1% of its current value at z ∼ 0.75, 2.0, and 3.5, respectively. The quiescent population drives most of the overall growth, with the stellar mass density of these galaxies increasing as ρ{sub star}∝(1 + z){sup –4.7±0.4} since z = 3.5,more » whereas the mass density of star-forming galaxies increases as ρ{sub star}∝(1 + z){sup –2.3±0.2}. At z > 2.5, star-forming galaxies dominate the total SMF at all stellar masses, although a non-zero population of quiescent galaxies persists to z = 4. Comparisons of the K{sub s} -selected star-forming galaxy SMFs with UV-selected SMFs at 2.5 < z < 4 show reasonable agreement and suggest that UV-selected samples are representative of the majority of the stellar mass density at z > 3.5. We estimate the average mass growth of individual galaxies by selecting galaxies at fixed cumulative number density. The average galaxy with log(M{sub star}/M{sub ☉}) = 11.5 at z = 0.3 has grown in mass by only 0.2 dex (0.3 dex) since z = 2.0 (3.5), whereas those with log(M{sub star}/M{sub ☉}) = 10.5 have grown by >1.0 dex since z = 2. At z < 2, the time derivatives of the mass growth are always larger for lower-mass galaxies, which demonstrates that the mass growth in galaxies since that redshift is mass-dependent and primarily bottom-up. Lastly, we examine potential sources of systematic uncertainties in the SMFs and find that those from photo-z templates, stellar population synthesis modeling, and the definition of quiescent galaxies dominate the total error budget in the SMFs.« less

  13. Use of multiwavelength emission from hollow cathode lamp for measurement of state resolved atom density of metal vapor produced by electron beam evaporation

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Majumder, A.; Dikshit, B.; Bhatia, M. S.

    2008-09-15

    State resolved atom population of metal vapor having low-lying metastable states departs from equilibrium value. It needs to be experimentally investigated. This paper reports the use of hollow cathode lamp based atomic absorption spectroscopy technique to measure online the state resolved atom density (ground and metastable) of metal vapor in an atomic beam produced by a high power electron gun. In particular, the advantage of availability of multiwavelength emission in hollow cathode lamp is used to determine the atom density in different states. Here, several transitions pertaining to a given state have also been invoked to obtain the mean valuemore » of atom density thereby providing an opportunity for in situ averaging. It is observed that at higher source temperatures the atoms from metastable state relax to the ground state. This is ascribed to competing processes of atom-atom and electron-atom collisions. The formation of collision induced virtual source is inferred from measurement of atom density distribution profile along the width of the atomic beam. The total line-of-sight average atom density measured by absorption technique using hollow cathode lamp is compared to that measured by atomic vapor deposition method. The presence of collisions is further supported by determination of beaming exponent by numerically fitting the data.« less

  14. Blood Density Is Nearly Equal to Water Density: A Validation Study of the Gravimetric Method of Measuring Intraoperative Blood Loss.

    PubMed

    Vitello, Dominic J; Ripper, Richard M; Fettiplace, Michael R; Weinberg, Guy L; Vitello, Joseph M

    2015-01-01

    Purpose. The gravimetric method of weighing surgical sponges is used to quantify intraoperative blood loss. The dry mass minus the wet mass of the gauze equals the volume of blood lost. This method assumes that the density of blood is equivalent to water (1 gm/mL). This study's purpose was to validate the assumption that the density of blood is equivalent to water and to correlate density with hematocrit. Methods. 50 µL of whole blood was weighed from eighteen rats. A distilled water control was weighed for each blood sample. The averages of the blood and water were compared utilizing a Student's unpaired, one-tailed t-test. The masses of the blood samples and the hematocrits were compared using a linear regression. Results. The average mass of the eighteen blood samples was 0.0489 g and that of the distilled water controls was 0.0492 g. The t-test showed P = 0.2269 and R (2) = 0.03154. The hematocrit values ranged from 24% to 48%. The linear regression R (2) value was 0.1767. Conclusions. The R (2) value comparing the blood and distilled water masses suggests high correlation between the two populations. Linear regression showed the hematocrit was not proportional to the mass of the blood. The study confirmed that the measured density of blood is similar to water.

  15. High-Resolution Detection of Identity by Descent in Unrelated Individuals

    PubMed Central

    Browning, Sharon R.; Browning, Brian L.

    2010-01-01

    Detection of recent identity by descent (IBD) in population samples is important for population-based linkage mapping and for highly accurate genotype imputation and haplotype-phase inference. We present a method for detection of recent IBD in population samples. Our method accounts for linkage disequilibrium between SNPs to enable full use of high-density SNP data. We find that our method can detect segments of a length of 2 cM with moderate power and negligible false discovery rate in Illumina 550K data in Northwestern Europeans. We compare our method with GERMLINE and PLINK, and we show that our method has a level of resolution that is significantly better than these existing methods, thus extending the usefulness of recent IBD in analysis of high-density SNP data. We survey four genomic regions in a sample of UK individuals of European descent and find that on average, at a given location, our method detects IBD in 2.7 per 10,000 pairs of individuals in Illumina 550K data. We also present methodology and results for detection of homozygosity by descent (HBD) and survey the whole genome in a sample of 1373 UK individuals of European descent. We detect HBD in 4.7 individuals per 10,000 on average at a given location. Our methodology is implemented in the freely available BEAGLE software package. PMID:20303063

  16. An Effective Cuckoo Search Algorithm for Node Localization in Wireless Sensor Network.

    PubMed

    Cheng, Jing; Xia, Linyuan

    2016-08-31

    Localization is an essential requirement in the increasing prevalence of wireless sensor network (WSN) applications. Reducing the computational complexity, communication overhead in WSN localization is of paramount importance in order to prolong the lifetime of the energy-limited sensor nodes and improve localization performance. This paper proposes an effective Cuckoo Search (CS) algorithm for node localization. Based on the modification of step size, this approach enables the population to approach global optimal solution rapidly, and the fitness of each solution is employed to build mutation probability for avoiding local convergence. Further, the approach restricts the population in the certain range so that it can prevent the energy consumption caused by insignificant search. Extensive experiments were conducted to study the effects of parameters like anchor density, node density and communication range on the proposed algorithm with respect to average localization error and localization success ratio. In addition, a comparative study was conducted to realize the same localization task using the same network deployment. Experimental results prove that the proposed CS algorithm can not only increase convergence rate but also reduce average localization error compared with standard CS algorithm and Particle Swarm Optimization (PSO) algorithm.

  17. An Effective Cuckoo Search Algorithm for Node Localization in Wireless Sensor Network

    PubMed Central

    Cheng, Jing; Xia, Linyuan

    2016-01-01

    Localization is an essential requirement in the increasing prevalence of wireless sensor network (WSN) applications. Reducing the computational complexity, communication overhead in WSN localization is of paramount importance in order to prolong the lifetime of the energy-limited sensor nodes and improve localization performance. This paper proposes an effective Cuckoo Search (CS) algorithm for node localization. Based on the modification of step size, this approach enables the population to approach global optimal solution rapidly, and the fitness of each solution is employed to build mutation probability for avoiding local convergence. Further, the approach restricts the population in the certain range so that it can prevent the energy consumption caused by insignificant search. Extensive experiments were conducted to study the effects of parameters like anchor density, node density and communication range on the proposed algorithm with respect to average localization error and localization success ratio. In addition, a comparative study was conducted to realize the same localization task using the same network deployment. Experimental results prove that the proposed CS algorithm can not only increase convergence rate but also reduce average localization error compared with standard CS algorithm and Particle Swarm Optimization (PSO) algorithm. PMID:27589756

  18. Nesting success of Northern Pintails on the coastal Yukon-Kuskokwim Delta, Alaska

    USGS Publications Warehouse

    Flint, Paul L.; Grand, James B.

    1996-01-01

    We studied nesting chronology and success of Northern Pintails (Anas acuta) on the coastal Yukon-Kuskokwim Delta, Alaska during the summers of 1991-1993. We found a total of 795 nests during three annual searches of a 27.4 km2 area. Minimum nest density averaged 9.67 nests per km2. Nesting success varied among years and ranged from 43.12% in 1991 to 10.74% in 1993 (average 23.95%). Most nest loss was the result of predation and tidal flooding. Daily nest survival probability declined with nest initiation date in all three years and also varied with nest age in 1992. Clutch size averaged 7.63 ± 0.067 (SE) eggs per nest and was larger than reported for other populations of Northern Pintails. Clutch size declined during the 44-47 day nesting interval at a greater rate than reported for other populations of Northern Pintails. We conclude that sub-arctic and prairie nesting Northern Pintails have similar reproductive potentials.

  19. r- and K-selection in fluctuating populations is determined by the evolutionary trade-off between two fitness measures: Growth rate and lifetime reproductive success.

    PubMed

    Engen, Steinar; Saether, Bernt-Erik

    2017-01-01

    In a stable environment, evolution maximizes growth rates in populations that are not density regulated and the carrying capacity in the case of density regulation. In a fluctuating environment, evolution maximizes a function of growth rate, carrying capacity and environmental variance, tending to r-selection and K-selection under large and small environmental noise, respectively. Here we analyze a model in which birth and death rates depend on density through the same function but with independent strength of density dependence. As a special case, both functions may be linear, corresponding to logistic dynamics. It is shown that evolution maximizes a function of the deterministic growth rate r 0 and the lifetime reproductive success (LRS) R 0 , both defined at small densities, as well as the environmental variance. Under large noise this function is dominated by r 0 and average lifetimes are small, whereas R 0 dominates and lifetimes are larger under small noise. Thus, K-selection is closely linked to selection for large R 0 so that evolution tends to maximize LRS in a stable environment. Consequently, different quantities (r 0 and R 0 ) tend to be maximized at low and high densities, respectively, favoring density-dependent changes in the optimal life history. © 2016 The Author(s). Evolution © 2016 The Society for the Study of Evolution.

  20. Attributes for MRB_E2RF1 Catchments in Selected Major River Basins: Population Density, 2000

    USGS Publications Warehouse

    Wieczorek, Michael; LaMotte, Andrew E.

    2010-01-01

    This data set represents the average population density, in number of people per square kilometer multiplied by 10 for the year 2000, compiled for every MRB_E2RF1 catchment of selected Major River Basins (MRBs, Crawford and others, 2006). The source data set is the 2000 Population Density by Block Group for the Conterminous United States (Hitt, 2003). The MRB_E2RF1 catchments are based on a modified version of the U.S. Environmental Protection Agency's (USEPA) RF1_2 and include enhancements to support national and regional-scale surface-water quality modeling (Nolan and others, 2002; Brakebill and others, 2011). Data were compiled for every MRB_E2RF1 catchment for the conterminous United States covering covering New England and Mid-Atlantic (MRB1), South Atlantic-Gulf and Tennessee (MRB2), the Great Lakes, Ohio, Upper Mississippi, and Souris-Red-Rainy (MRB3), the Missouri (MRB4), the Lower Mississippi, Arkansas-White-Red, and Texas-Gulf (MRB5), the Rio Grande, Colorado, and the Great basin (MRB6), the Pacific Northwest (MRB7) river basins, and California (MRB8).

  1. Population dynamics of zebra mussels Dreissena polymorpha (Pallas, 1771) during the initial invasion of the Upper Mississippi River, USA

    USGS Publications Warehouse

    Cope, W.G.; Bartsch, M.R.; Hightower, J.E.

    2006-01-01

    The aim of this study was to document and model the population dynamics of zebra mussels Dreissena polymorpha (Pallas, 1771) in Pool 8 of the Upper Mississippi River (UMR), USA, for five consecutive years (1992-1996) following their initial discovery in September 1991. Artificial substrates (concrete blocks, 0.49 m2 surface area) were deployed on or around the first of May at two sites within each of two habitat types (main channel border and contiguous backwater). Blocks were removed monthly (30 ?? 10 d) from the end of May to the end of October to obtain density and growth information. Some blocks deployed in May 1995 were retrieved in April 1996 to obtain information about overwinter growth and survival. The annual density of zebra mussels in Pool 8 of the UMR increased from 3.5/m2 in 1992 to 14,956/m 2 in 1996. The average May-October growth rate of newly recruited individuals, based on a von Bertalanffy growth model fitted to monthly shell-length composition data, was 0.11 mm/d. Model estimates of the average survival rate varied from 21 to 100% per month. Estimated recruitment varied substantially among months, with highest levels occurring in September-October of 1994 and 1996, and in July of 1995. Recruitment and density in both habitat types increased by two orders of magnitude in 1996. Follow-up studies will be necessary to assess the long-term stability of zebra mussel populations in the UMR; this study provides the critical baseline information needed for those future comparisons. ?? Published by Oxford University Press on behalf of The Malacological Society of London 2006.

  2. Do food availability, parasitism, and stress have synergistic effects on red colobus populations living in forest fragments?

    PubMed

    Chapman, Colin A; Wasserman, Michael D; Gillespie, Thomas R; Speirs, Michaela L; Lawes, Michael J; Saj, Tania L; Ziegler, Toni E

    2006-12-01

    Identifying factors that influence animal density is a fundamental goal in ecology that has taken on new importance with the need to develop informed management plans. This is particularly the case for primates as the tropical forest that supports many species is being rapidly converted. We use a system of forest fragments adjacent to Kibale National Park, Uganda, to examine if food availability and parasite infections have synergistic affects on red colobus (Piliocolobus tephrosceles) abundance. Given that the size of primate populations can often respond slowly to environmental changes, we also examined how these factors influenced cortisol levels. To meet these objectives, we monitored gastrointestinal parasites, evaluated fecal cortisol levels, and determined changes in food availability by conducting complete tree inventories in eight fragments in 2000 and 2003. Red colobus populations declined by an average of 21% among the fragments; however, population change ranged from a 25% increase to a 57% decline. The cumulative basal area of food trees declined by an average of 29.5%; however, forest change was highly variable (a 2% gain to a 71% decline). We found that nematode prevalence averaged 58% among fragments (range 29-83%). The change in colobus population size was correlated both with food availability and a number of indices of parasite infections. A path analysis suggests that change in food availability has a strong direct effect on population size, but it also has an indirect effect via parasite infections. 2006 Wiley-Liss, Inc.

  3. Modeling mechanical inhomogeneities in small populations of proliferating monolayers and spheroids.

    PubMed

    Lejeune, Emma; Linder, Christian

    2018-06-01

    Understanding the mechanical behavior of multicellular monolayers and spheroids is fundamental to tissue culture, organism development, and the early stages of tumor growth. Proliferating cells in monolayers and spheroids experience mechanical forces as they grow and divide and local inhomogeneities in the mechanical microenvironment can cause individual cells within the multicellular system to grow and divide at different rates. This differential growth, combined with cell division and reorganization, leads to residual stress. Multiple different modeling approaches have been taken to understand and predict the residual stresses that arise in growing multicellular systems, particularly tumor spheroids. Here, we show that by using a mechanically robust agent-based model constructed with the peridynamic framework, we gain a better understanding of residual stresses in multicellular systems as they grow from a single cell. In particular, we focus on small populations of cells (1-100 s) where population behavior is highly stochastic and prior investigation has been limited. We compare the average strain energy density of cells in monolayers and spheroids using different growth and division rules and find that, on average, cells in spheroids have a higher strain energy density than cells in monolayers. We also find that cells in the interior of a growing spheroid are, on average, in compression. Finally, we demonstrate the importance of accounting for stochastic fluctuations in the mechanical environment, particularly when the cellular response to mechanical cues is nonlinear. The results presented here serve as a starting point for both further investigation with agent-based models, and for the incorporation of major findings from agent-based models into continuum scale models when explicit representation of individual cells is not computationally feasible.

  4. Ranging, Activity and Habitat Use by Tigers in the Mangrove Forests of the Sundarban.

    PubMed

    Naha, Dipanjan; Jhala, Yadvendradev V; Qureshi, Qamar; Roy, Manjari; Sankar, Kalyansundaram; Gopal, Rajesh

    2016-01-01

    The Sundarban of India and Bangladesh (about 6000 km²) are the only mangrove forests inhabited by a sizeable population of tigers. The adjoining area also supports one of the highest human densities and experiences severe human-tiger conflicts. We used GPS-Satellite and VHF radio-collars on 6 (3 males and 3 female) tigers to study their ranging patterns and habitat preference. The average home range (95% Fixed Kernel) for resident females was 56.4 (SE 5.69) and for males it was 110 (SE 49) km². Tigers crossed an average of 5 water channels > 30 meters per day with a mean width of 54 meters, whereas channels larger than 400 meters were rarely crossed. Tigers spent over 58% of their time within Phoenix habitat but compositional analysis showed a habitat preference of the order Avicennia-Sonneratia > Phoenix > Ceriops > Barren > Water. Average daily distance moved was 4.6 km (range 0.1-23). Activity of tigers peaked between 05:00 hours and 10:00 hours showing some overlap with human activity. Territory boundaries were demarcated by large channels which tigers intensively patrolled. Extra caution should be taken while fishing or honey collection during early morning in Avicennia-Sonneratia and Phoenix habitat types along wide channels to reduce human-tiger conflict. Considering home-range core areas as exclusive, tiger density was estimated at 4.6 (SE range 3.6 to 6.7) tigers/100 km2 giving a total population of 76 (SE range 59-110) tigers in the Indian Sundarban. Reluctance of tigers to cross wide water channels combined with increasing commercial boat traffic and sea level rise due to climate change pose a real threat of fragmenting the Sundarban tiger population.

  5. Ranging, Activity and Habitat Use by Tigers in the Mangrove Forests of the Sundarban

    PubMed Central

    Naha, Dipanjan; Jhala, Yadvendradev V.; Qureshi, Qamar; Roy, Manjari; Sankar, Kalyansundaram; Gopal, Rajesh

    2016-01-01

    The Sundarban of India and Bangladesh (about 6000 km²) are the only mangrove forests inhabited by a sizeable population of tigers. The adjoining area also supports one of the highest human densities and experiences severe human-tiger conflicts. We used GPS-Satellite and VHF radio-collars on 6 (3 males and 3 female) tigers to study their ranging patterns and habitat preference. The average home range (95% Fixed Kernel) for resident females was 56.4 (SE 5.69) and for males it was 110 (SE 49) km². Tigers crossed an average of 5 water channels > 30 meters per day with a mean width of 54 meters, whereas channels larger than 400 meters were rarely crossed. Tigers spent over 58% of their time within Phoenix habitat but compositional analysis showed a habitat preference of the order Avicennia-Sonneratia > Phoenix > Ceriops > Barren > Water. Average daily distance moved was 4.6 km (range 0.1–23). Activity of tigers peaked between 05:00 hours and 10:00 hours showing some overlap with human activity. Territory boundaries were demarcated by large channels which tigers intensively patrolled. Extra caution should be taken while fishing or honey collection during early morning in Avicennia-Sonneratia and Phoenix habitat types along wide channels to reduce human-tiger conflict. Considering home-range core areas as exclusive, tiger density was estimated at 4.6 (SE range 3.6 to 6.7) tigers/100 km2 giving a total population of 76 (SE range 59–110) tigers in the Indian Sundarban. Reluctance of tigers to cross wide water channels combined with increasing commercial boat traffic and sea level rise due to climate change pose a real threat of fragmenting the Sundarban tiger population. PMID:27049644

  6. Positive density-dependent reproduction regulated by local kinship and size in an understorey tropical tree

    PubMed Central

    Castilla, Antonio R.; Pope, Nathaniel; Jha, Shalene

    2016-01-01

    Background and Aims Global pollinator declines and continued habitat fragmentation highlight the critical need to understand reproduction and gene flow across plant populations. Plant size, conspecific density and local kinship (i.e. neighbourhood genetic relatedness) have been proposed as important mechanisms influencing the reproductive success of flowering plants, but have rarely been simultaneously investigated. Methods We conducted this study on a continuous population of the understorey tree Miconia affinis in the Forest Dynamics Plot on Barro Colorado Island in central Panama. We used spatial, reproductive and population genetic data to investigate the effects of tree size, conspecific neighbourhood density and local kinship on maternal and paternal reproductive success. We used a Bayesian framework to simultaneously model the effects of our explanatory variables on the mean and variance of maternal viable seed set and siring success. Key Results Our results reveal that large trees had lower proportions of viable seeds in their fruits but sired more seeds. We documented differential effects of neighbourhood density and local kinship on both maternal and paternal reproductive components. Trees in more dense neighbourhoods produced on average more viable seeds, although this positive density effect was influenced by variance-inflation with increasing local kinship. Neighbourhood density did not have significant effects on siring success. Conclusions This study is one of the first to reveal an interaction among tree size, conspecific density and local kinship as critical factors differentially influencing maternal and paternal reproductive success. We show that both maternal and paternal reproductive success should be evaluated to determine the population-level and individual traits most essential for plant reproduction. In addition to conserving large trees, we suggest the inclusion of small trees and the conservation of dense patches with low kinship as potential strategies for strengthening the reproductive status of tropical trees. PMID:26602288

  7. Positive density-dependent reproduction regulated by local kinship and size in an understorey tropical tree.

    PubMed

    Castilla, Antonio R; Pope, Nathaniel; Jha, Shalene

    2016-02-01

    Global pollinator declines and continued habitat fragmentation highlight the critical need to understand reproduction and gene flow across plant populations. Plant size, conspecific density and local kinship (i.e. neighbourhood genetic relatedness) have been proposed as important mechanisms influencing the reproductive success of flowering plants, but have rarely been simultaneously investigated. We conducted this study on a continuous population of the understorey tree Miconia affinis in the Forest Dynamics Plot on Barro Colorado Island in central Panama. We used spatial, reproductive and population genetic data to investigate the effects of tree size, conspecific neighbourhood density and local kinship on maternal and paternal reproductive success. We used a Bayesian framework to simultaneously model the effects of our explanatory variables on the mean and variance of maternal viable seed set and siring success. Our results reveal that large trees had lower proportions of viable seeds in their fruits but sired more seeds. We documented differential effects of neighbourhood density and local kinship on both maternal and paternal reproductive components. Trees in more dense neighbourhoods produced on average more viable seeds, although this positive density effect was influenced by variance-inflation with increasing local kinship. Neighbourhood density did not have significant effects on siring success. This study is one of the first to reveal an interaction among tree size, conspecific density and local kinship as critical factors differentially influencing maternal and paternal reproductive success. We show that both maternal and paternal reproductive success should be evaluated to determine the population-level and individual traits most essential for plant reproduction. In addition to conserving large trees, we suggest the inclusion of small trees and the conservation of dense patches with low kinship as potential strategies for strengthening the reproductive status of tropical trees. © The Author 2015. Published by Oxford University Press on behalf of the Annals of Botany Company. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

  8. [Genetic ecological monitoring in human populations: heterozygosity, mtDNA haplotype variation, and genetic load].

    PubMed

    Balanovskiĭ, O P; Koshel', S M; Zaporozhchenko, V V; Pshenichnov, A S; Frolova, S A; Kuznetsova, M A; Baranova, E E; Teuchezh, I E; Kuznetsova, A A; Romashkina, M V; Utevskaia, O M; Churnosov, M I; Villems, R; Balanovskaia, E V

    2011-11-01

    Yu. P. Altukhov suggested that heterozygosity is an indicator of the state of the gene pool. The idea and a linked concept of genetic ecological monitoring were applied to a new dataset on mtDNA variation in East European ethnic groups. Haplotype diversity (an analog of the average heterozygosity) was shown to gradually decrease northwards. Since a similar trend is known for population density, interlinked changes were assumed for a set of parameters, which were ordered to form a causative chain: latitude increases, land productivity decreases, population density decreases, effective population size decreases, isolation of subpopulations increases, genetic drift increases, and mtDNA haplotype diversity decreases. An increase in genetic drift increases the random inbreeding rate and, consequently, the genetic load. This was confirmed by a significant correlation observed between the incidence of autosomal recessive hereditary diseases and mtDNA haplotype diversity. Based on the findings, mtDNA was assumed to provide an informative genetic system for genetic ecological monitoring; e.g., analyzing the ecology-driven changes in the gene pool.

  9. Strong contribution of immigration to local population regulation: evidence from a migratory passerine.

    PubMed

    Schaub, Michael; Jakober, Hans; Stauber, Wolfgang

    2013-08-01

    A mechanistic understanding of the dynamics of populations requires knowledge about the variation of the underlying demographic rates and about the reasons for their variability. In geographically open populations, immigration is often necessary to prevent declines, but little is known about whether immigration can contribute to its regulation. We studied the dynamics of a Red-backed Shrike population (Lanius collurio) over 36 years in Germany with a Bayesian integrated population model. We estimated mean and temporal variability of population sizes, productivity, apparent survival, and immigration. We assessed how strongly the demographic rates were correlated with population growth to understand the demographic reasons of population change and how strongly the demographic rates were correlated with population size to identify possible density-dependent mechanisms. The shrike population varied between 35 and 74 breeding pairs but did not show a significant trend in population size over time (growth rate 1.002 +/- 0.001 [mean +/- SD]). Apparent survival of females (juveniles 0.06 +/- 0.01; adults 0.37 +/- 0.03) was lower than that of males (juveniles 0.10 +/- 0.01; adults 0.44 +/- 0.02). Immigration rates were substantial and higher in females (0.56 +/- 0.02) than in males (0.43 +/- 0.02), and average productivity was 2.76 +/- 0.14. Without immigration, the Red-backed Shrike population would have declined strongly. Immigration was the strongest driver for the number of females while local recruitment was the most important driver for the number of males. Immigration of both sexes and productivity, but not local recruitment and survival, were subject to density dependence. Density-dependent productivity was not effectively regulating the local population but may have contributed to regulate shrike populations at larger spatial scales. These findings suggest that immigration is not only an important component to prevent a geographically open population from decline, but that it can also contribute to its regulation.

  10. The Velocity and Density Distribution of Earth-Intersecting Meteoroids: Implications for Environment Models

    NASA Technical Reports Server (NTRS)

    Moorhead, A. V.; Brown, P. G.; Campbell-Brown, M. D.; Moser, D. E.; Blaauw, R. C.; Cooke, W. J.

    2017-01-01

    Meteoroids are known to damage spacecraft: they can crater or puncture components, disturb a spacecraft's attitude, and potentially create secondary electrical effects. Because the damage done depends on the speed, size, density, and direction of the impactor, accurate environment models are critical for mitigating meteoroid-related risks. Yet because meteoroid properties are derived from indirect observations such as meteors and impact craters, many characteristics of the meteoroid environment are uncertain. In this work, we present recent improvements to the meteoroid speed and density distributions. Our speed distribution is derived from observations made by the Canadian Meteor Orbit Radar. These observations are de-biased using modern descriptions of the ionization efficiency. Our approach yields a slower meteoroid population than previous analyses (see Fig. 1 for an example) and we compute the uncertainties associated with our derived distribution. We adopt a higher fidelity density distribution than that used by many older models. In our distribution, meteoroids with TJ less than 2 are assigned to a low-density population, while those with TJ greater than 2 have higher densities (see Fig. 2). This division and the distributions themselves are derived from the densities reported by Kikwaya et al. These changes have implications for the environment: for instance, the helion/antihelion sporadic sources have lower speeds than the apex and toroidal sources and originate from high-T(sub J) parent bodies. Our on-average slower and denser distributions thus imply that the helion and antihelion sources dominate the meteoroid environment even more completely than previously thought. Finally, for a given near-Earth meteoroid cratering rate, a slower meteoroid population produces a comparatively higher rate of satellite attitude disturbances.

  11. Response of a cotton rat population to increased density

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Bigler, W.J.; Lebetkin, E.H.; Cumbie, P.M.

    Experiments were conducted in old-field enclosures to define some of the responses of a confined, nonreproducing cotton rat population to increases in density. Successive groups of alien animals were randomly added to a monitored resident population on two occasions. Aliens exhibited high rates of mortality and weight loss. The alien males were particularly affected, only 18 percent surviving the experiment. Residents could not be accounted for by trapping as easily as aliens. Residents concentrated in discrete areas of preferred habitat yet ranged over the entire enclosure, while aliens stayed in heavy ground cover close to the walls. Analysis of themore » average distance between captures of each group on a daily basis revealed increases in resident movements and corresponding decreases in alien movements. Changes in spatial distribution, movement patterns, sex ratio and rats of mortality due to trap deaths and predation in the alien groups reflected behavioral interactions that operate as a form of social exclusion.« less

  12. Scaling and entropy in p-median facility location along a line

    NASA Astrophysics Data System (ADS)

    Gastner, Michael T.

    2011-09-01

    The p-median problem is a common model for optimal facility location. The task is to place p facilities (e.g., warehouses or schools) in a heterogeneously populated space such that the average distance from a person's home to the nearest facility is minimized. Here we study the special case where the population lives along a line (e.g., a road or a river). If facilities are optimally placed, the length of the line segment served by a facility is inversely proportional to the square root of the population density. This scaling law is derived analytically and confirmed for concrete numerical examples of three US interstate highways and the Mississippi River. If facility locations are permitted to deviate from the optimum, the number of possible solutions increases dramatically. Using Monte Carlo simulations, we compute how scaling is affected by an increase in the average distance to the nearest facility. We find that the scaling exponents change and are most sensitive near the optimum facility distribution.

  13. [Imbalance of distribution of physicians in the Arab world].

    PubMed

    el Matri, A

    1990-01-01

    The Arab world, as a whole, now has 8.5 medical doctors per 10,000 population. This average covers a great diversity of national situations; for example 0.6 doctors per 10,000 population in Somalia, and 17.5 doctors per 10,000 population in Qatar. From the viewpoint of medical staffing, the Arab countries can be grouped into four categories: (i) Countries with low medical density: their national staff is scarce, they do not have a long academic tradition, and they have to rely on foreign physicians (Yemen, Djibouti, Somalia, Mauritania). (ii) Countries with acceptable medical density: they have strengthened their medical training system during the last decades, their production of medical graduates is now fairly high, and there is seemingly a threat of oversupply (Algeria, Tunisia, Syria, Iraq). (iii) Countries with relatively high medical density: their medical staffing is below that of developed countries, but their economic possibilities are reduced, which leads to an oversupply; these countries are exporters of medical manpower (Egypt, Jordan). (iv) Countries that import medical manpower although their own medical density is quite high: their medical training is recent or non-existent and their health system relies to a great extent on foreign doctors (Saudi Arabia, Bahrain, Qatar, United Arab Emirates, Libya, Kuwait). They are all oil producers. In brief, a shortage and an oversupply of medical doctors coexist in the Arab world, and concerted action is required.

  14. Golden eagle population trends in the western United States: 1968-2010

    USGS Publications Warehouse

    Millsap, Brian A.; Zimmerman, Guthrie S.; Sauer, John R.; Nielson, Ryan M.; Otto, Mark; Bjerre, Emily; Murphy, Robert K.

    2013-01-01

    In 2009, the United States Fish and Wildlife Service promulgated permit regulations for the unintentional lethal take (anthropogenic mortality) and disturbance of golden eagles (Aquila chrysaetos). Accurate population trend and size information for golden eagles are needed so agency biologists can make informed decisions when eagle take permits are requested. To address this need with available data, we used a log-linear hierarchical model to average data from a late-summer aerial-line-transect distance-sampling survey (WGES) of golden eagles in the United States portions of Bird Conservation Region (BCR) 9 (Great Basin), BCR 10 (Northern Rockies), BCR 16 (Southern Rockies/Colorado Plateau), and BCR 17 (Badlands and Prairies) from 2006 to 2010 with late-spring, early summer Breeding Bird Survey (BBS) data for the same BCRs and years to estimate summer golden eagle population size and trends in these BCRs. We used the ratio of the density estimates from the WGES to the BBS index to calculate a BCR-specific adjustment factor that scaled the BBS index (i.e., birds per route) to a density estimate. Our results indicated golden eagle populations were generally stable from 2006 to 2010 in the 4 BCRs, with an estimated average rate of population change of −0.41% (95% credible interval [CI]: −4.17% to 3.40%) per year. For the 4 BCRs and years, we estimated annual golden eagle population size to range from 28,220 (95% CI: 23,250–35,110) in 2007 to 26,490 (95% CI: 21,760–32,680) in 2008. We found a general correspondence in trends between WGES and BBS data for these 4 BCRs, which suggested BBS data were providing useful trend information. We used the overall adjustment factor calculated from the 4 BCRs and years to scale BBS golden eagle counts from 1968 to 2005 for the 4 BCRs and for 1968 to 2010 for the 8 other BCRs (without WGES data) to estimate golden eagle population size and trends across the western United States for the period 1968 to 2010. In general, we noted slightly declining trends in southern BCRs and slightly increasing trends in northern BCRs. However, we estimated the average rate of golden eagle population change across all 12 BCRs for the period 1968–2010 as +0.40% per year (95% CI = −0.27% to 1.00%), suggesting a stable population. We also estimated the average rate of population change for the period 1990–2010 was +0.5% per year (95% CI = −0.33% to 1.3%). Our annual estimates of population size for the most recent decade range from 31,370 (95% CI: 25,450–39,310) in 2004 to 33,460 (95% CI: 27,380–41,710) in 2007. Our results clarify that golden eagles are not declining widely in the western United States. © 2013 The Wildlife Society.

  15. Diminishing returns and Malthus's first essay on population: theory and application.

    PubMed

    Hollander, S

    1989-06-01

    "This paper argues that Malthus's famous geometrical and arithmetical ratios do not necessarily imply diminishing returns. That average product would not necessarily fall with increased population density is immaterial to the polemical purpose of the Essay on Population which is satisfied by demonstrating the necessity for a reduction of the population growth below its maximum potential. But I do not deny the presence of the diminishing returns principle in the first Essay. Indeed, here is found the origin of Malthus's later land-scarcity based growth model involving a declining path of real wages in consequence of a decelerating rate of growth of labour demand." The author also considers Malthus's analysis of the situation in Britain in 1798. (SUMMARY IN FRE) excerpt

  16. The balance between adaptation to catalysts and competition radius shapes the total wealth, time variability and inequality

    NASA Astrophysics Data System (ADS)

    Davidovich, Hadar; Louzoun, Yoram

    2013-05-01

    The globalization of modern markets has led to the emergence of competition between producers in ever growing distances. This opens the interesting question in population dynamics of the effect of long-range competition. We here study a model of non-local competition to test the effect of the competition radius on the wealth distribution, using the framework of a stochastic birth-death process, with non-local interactions. We show that this model leads to non-trivial dynamics that can have implications in other domains of physics. Competition is studied in the context of the catalyst induced growth of autocatalytic agents, representing the growth of capital in the presence of investment opportunities. These agents are competing with all other agents in a given radius on growth possibilities. We show that a large scale competition leads to an extreme localization of the agents, where typically a single aggregate of agents can survive within a given competition radius. The survival of these aggregates is determined by the diffusion rates of the agents and the catalysts. For high and low agent diffusion rates, the agent population is always annihilated, while for intermediate diffusion rates, a finite agent population persists. Increasing the catalyst diffusion rate always leads to a decrease in the average agent population density. The extreme localization of the agents leads to the emergence of intermittent fluctuations, when a large aggregate of agents disappear. As the competition radius increases, so does the average agent density and its spatial variance as well as the volatility.

  17. Distribution of Brazilian dermatologists according to geographic location, population and HDI of municipalities: an ecological study*

    PubMed Central

    Schmitt, Juliano Vilaverde; Miot, Hélio Amante

    2014-01-01

    This study investigated the geographic distribution of dermatologists in Brazilian municipalities in relation to the population, regions of the country and human development index. We conducted an ecological study based on data from the 2010 census, the 2010 human development index, and the records of the Brazilian Society of Dermatology. 5565 municipalities and 6718 dermatologists were surveyed. Only 504 (9.1%) municipalities had dermatologists, and accounted for 56.2% of the Brazilian population. The smallest population size and lowest HDI rate that best discriminated municipalities that did not have dermatologists were found to be 28,000 and 0.71, respectively. The average population density of dermatologists in cities was 1/23.000 inhabitants, and variations were independently associated with the HDI, the population of the municipalities and the region of the country. PMID:25387516

  18. Demographics of an ornate box turtle population experiencing minimal human-induced disturbances

    USGS Publications Warehouse

    Converse, S.J.; Iverson, J.B.; Savidge, J.A.

    2005-01-01

    Human-induced disturbances may threaten the viability of many turtle populations, including populations of North American box turtles. Evaluation of the potential impacts of these disturbances can be aided by long-term studies of populations subject to minimal human activity. In such a population of ornate box turtles (Terrapene ornata ornata) in western Nebraska, we examined survival rates and population growth rates from 1981-2000 based on mark-recapture data. The average annual apparent survival rate of adult males was 0.883 (SE = 0.021) and of adult females was 0.932 (SE = 0.014). Minimum winter temperature was the best of five climate variables as a predictor of adult survival. Survival rates were highest in years with low minimum winter temperatures, suggesting that global warming may result in declining survival. We estimated an average adult population growth rate (????) of 1.006 (SE = 0.065), with an estimated temporal process variance (????2) of 0.029 (95% CI = 0.005-0.176). Stochastic simulations suggest that this mean and temporal process variance would result in a 58% probability of a population decrease over a 20-year period. This research provides evidence that, unless unknown density-dependent mechanisms are operating in the adult age class, significant human disturbances, such as commercial harvest or turtle mortality on roads, represent a potential risk to box turtle populations. ?? 2005 by the Ecological Society of America.

  19. Genome-Wide Single-Nucleotide Polymorphisms Discovery and High-Density Genetic Map Construction in Cauliflower Using Specific-Locus Amplified Fragment Sequencing

    PubMed Central

    Zhao, Zhenqing; Gu, Honghui; Sheng, Xiaoguang; Yu, Huifang; Wang, Jiansheng; Huang, Long; Wang, Dan

    2016-01-01

    Molecular markers and genetic maps play an important role in plant genomics and breeding studies. Cauliflower is an important and distinctive vegetable; however, very few molecular resources have been reported for this species. In this study, a novel, specific-locus amplified fragment (SLAF) sequencing strategy was employed for large-scale single nucleotide polymorphism (SNP) discovery and high-density genetic map construction in a double-haploid, segregating population of cauliflower. A total of 12.47 Gb raw data containing 77.92 M pair-end reads were obtained after processing and 6815 polymorphic SLAFs between the two parents were detected. The average sequencing depths reached 52.66-fold for the female parent and 49.35-fold for the male parent. Subsequently, these polymorphic SLAFs were used to genotype the population and further filtered based on several criteria to construct a genetic linkage map of cauliflower. Finally, 1776 high-quality SLAF markers, including 2741 SNPs, constituted the linkage map with average data integrity of 95.68%. The final map spanned a total genetic length of 890.01 cM with an average marker interval of 0.50 cM, and covered 364.9 Mb of the reference genome. The markers and genetic map developed in this study could provide an important foundation not only for comparative genomics studies within Brassica oleracea species but also for quantitative trait loci identification and molecular breeding of cauliflower. PMID:27047515

  20. Cometary ion dynamics observed in the close vicinity of comet 67P/Churyumov-Gerasimenko during the intermediate activity period

    NASA Astrophysics Data System (ADS)

    Berčič, L.; Behar, E.; Nilsson, H.; Nicolaou, G.; Wieser, G. Stenberg; Wieser, M.; Goetz, C.

    2018-06-01

    Aims: Cometary ions are constantly produced in the coma, and once produced they are accelerated and eventually escape the coma. We describe and interpret the dynamics of the cometary ion flow, of an intermediate active comet, very close to the nucleus and in the terminator plane. Methods: We analysed in situ ion and magnetic field measurements, and characterise the velocity distribution functions (mostly using plasma moments). We propose a statistical approach over a period of one month. Results: On average, two populations were observed, separated in phase space. The motion of the first is governed by its interaction with the solar wind farther upstream, while the second one is accelerated in the inner coma and displays characteristics compatible with an ambipolar electric field. Both populations display a consistent anti-sunward velocity component. Conclusions: Cometary ions born in different regions of the coma are seen close to the nucleus of comet 67P/Churyumov-Gerasimenko with distinct motions governed in one case by the solar wind electric field and in the other case by the position relative to the nucleus. A consistent anti-sunward component is observed for all cometary ions. An asymmetry is found in the average cometary ion density in a solar wind electric field reference frame, with higher density in the negative (south) electric field hemisphere. There is no corresponding signature in the average magnetic field strength.

  1. Potential impact of Chironomus plumosus larvae on hypolimnetic oxygen in the central basin of Lake Erie

    USGS Publications Warehouse

    Soster, Frederick M.; Matisoff, Gerald; Schloesser, Donald W.; Edwards, William J.

    2015-01-01

    Previous studies have indicated that burrow-irrigating infauna can increase sediment oxygen demand (SOD) and impact hypolimnetic oxygen in stratified lakes. We conducted laboratory microcosm experiments and computer simulations with larvae of the burrowing benthic midge Chironomus plumosus to quantify burrow oxygen uptake rates and subsequent contribution to sediment oxygen demand in central Lake Erie. Burrow oxygen uptake and water flow velocities through burrows were measured using oxygen microelectrodes and hot film anemometry, respectively. Burrow oxygen consumption averaged 2.66 × 10− 10 (SE = ± 7.82 × 10− 11) mol O2/burrow/s at 24 °C and 9.64 × 10− 10 (SE = ± 4.86 × 10− 10) mol O2/burrow/s at 15 °C. In sealed microcosm experiments, larvae increased SOD 500% at 24 °C (density = 1508/m2) and 375% at 15 °C (density = 864/m2). To further evaluate effects of densities of C. plumosus burrows on SOD we developed a 3-D transport reaction model of the process. Using experimental data and chironomid abundance data in faunal surveys in 2009 and 2010, we estimated that bioirrigation by a population of 140 larvae/m2 could account for between 2.54 × 10− 11 mol/L/s (model results) and 5.58 × 10− 11 mol/L/s (experimental results) of the average 4.22 × 10− 11 mol/L/s oxygen depletion rate between 1970 and 2003, which could have accounted for 60–132% of the oxygen decline. At present, it appears that the population density of this species may be an important factor in development of hypoxic or anoxic conditions in central Lake Erie.

  2. Serengeti real estate: density vs. fitness-based indicators of lion habitat quality.

    PubMed

    Mosser, Anna; Fryxell, John M; Eberly, Lynn; Packer, Craig

    2009-10-01

    Habitat quality is typically inferred by assuming a direct relationship between consumer density and resource abundance, although it has been suggested that consumer fitness may be a more accurate measure of habitat quality. We examined density vs. fitness-based measures of habitat quality for lions in the Serengeti National Park, Tanzania. A 40-year average of female reproductive success (yearling cubs per female) was best explained by proximity to river confluences, whereas patterns of productivity (yearling cubs per km(2)) and adult female density (individuals per km(2)) were associated with more general measures of habitat quality and areas of shelter in poor habitat. This suggests that density may not accurately distinguish between high-quality 'source' areas and low-quality sites that merely provide refuges for effectively non-reproductive individuals. Our results indicate that density may be a misleading indicator of real estate value, particularly for populations that do not conform to an ideal free distribution.

  3. Long-term dynamics of Typha populations

    USGS Publications Warehouse

    Grace, J.B.; Wetzel, R.G.

    1998-01-01

    The zonation of Typha populations in an experimental pond in Michigan was re-examined 15 years after the original sampling to gain insight into the long-term dynamics. Current distributions of Typha populations were also examined in additional experimental ponds at the site that have been maintained for 23 years. The zonation between T. latifolia and T. angustifolia in the previously studied pond 15 years after the initial sampling revealed that the density and distribution of shoots had not changed significantly. Thus, it appears that previously reported results (based on 7- year old populations) have remained consistent over time. Additional insight into the interaction between these two taxa was sought by comparing mixed and monoculture stands in five experimental ponds that have remained undisturbed for their 23-year history. The maximum depth of T. latifolia, the shallow- water species, was not significantly reduced when growing in the presence of the more flood tolerant T. angustifolia. In contrast, the minimum depth of T. angustifolia was reduced from 0 to 37 cm when in the presence of T. latifolia. When total populations were compared between monoculture and mixed stands, the average density of T. angustifolia shoots was 59.4 percent lower in mixed stands while the density of T. latifolia was 32 percent lower, with T. angustifolia most affected at shallow depths (reduced by 92 percent) and T. latifolia most affected at the deepest depths (reduced by 60 percent). These long-term observations indicate that competitive displacement between Typha taxa has remained stable over time.

  4. Handling times and saturating transmission functions in a snail-worm symbiosis.

    PubMed

    Hopkins, Skylar R; McGregor, Cari M; Belden, Lisa K; Wojdak, Jeremy M

    2018-06-16

    All dynamic species interaction models contain an assumption that describes how contact rates scale with population density. Choosing an appropriate contact-density function is important, because different functions have different implications for population dynamics and stability. However, this choice can be challenging, because there are many possible functions, and most are phenomenological and thus difficult to relate to underlying ecological processes. Using one such phenomenological function, we described a nonlinear relationship between field transmission rates and host density in a common snail-oligochaete symbiosis. We then used a well-known contact function from predator-prey models, the Holling Type II functional response, to describe and predict host snail contact rates in the laboratory. The Holling Type II functional response accurately described both the nonlinear contact-density relationship and the average contact duration that we observed. Therefore, we suggest that contact rates saturate with host density in this system because each snail contact requires a non-instantaneous handling time, and additional possible contacts do not occur during that handling time. Handling times and nonlinear contact rates might also explain the nonlinear relationship between symbiont transmission and snail density that we observed in the field, which could be confirmed by future work that controls for other potential sources of seasonal variation in transmission rates. Because most animal contacts are not instantaneous, the Holling Type II functional response might be broadly relevant to diverse host-symbiont systems.

  5. Mobile population dynamics and malaria vulnerability: a modelling study in the China-Myanmar border region of Yunnan Province, China.

    PubMed

    Chen, Tian-Mu; Zhang, Shao-Sen; Feng, Jun; Xia, Zhi-Gui; Luo, Chun-Hai; Zeng, Xu-Can; Guo, Xiang-Rui; Lin, Zu-Rui; Zhou, Hong-Ning; Zhou, Shui-Sen

    2018-04-29

    The China-Myanmar border region presents a great challenge in malaria elimination in China, and it is essential to understand the relationship between malaria vulnerability and population mobility in this region. A community-based, cross-sectional survey was performed in five villages of Yingjiang county during September 2016. Finger-prick blood samples were obtained to identify asymptomatic infections, and imported cases were identified in each village (between January 2013 and September 2016). A stochastic simulation model (SSM) was used to test the relationship between population mobility and malaria vulnerability, according to the mechanisms of malaria importation. Thirty-two imported cases were identified in the five villages, with a 4-year average of 1 case/year (range: 0-5 cases/year). No parasites were detected in the 353 blood samples from 2016. The median density of malaria vulnerability was 0.012 (range: 0.000-0.033). The average proportion of mobile members of the study population was 32.56% (range: 28.38-71.95%). Most mobile individuals lived indoors at night with mosquito protection. The SSM model fit the investigated data (χ 2  = 0.487, P = 0.485). The average probability of infection in the members of the population that moved to Myanmar was 0.011 (range: 0.0048-0.1585). The values for simulated vulnerability increased with greater population mobility in each village. A high proportion of population mobility was associated with greater malaria vulnerability in the China-Myanmar border region. Mobile population-specific measures should be used to decrease the risk of malaria re-establishment in China.

  6. Genetic diversity affects the strength of population regulation in a marine fish.

    PubMed

    Johnson, D W; Freiwald, J; Bernardi, G

    2016-03-01

    Variation is an essential feature of biological populations, yet much of ecological theory treats individuals as though they are identical. This simplifying assumption is often justified by the perception that variation among individuals does not have significant effects on the dynamics of whole populations. However, this perception may be skewed by a historic focus on studying single populations. A true evaluation of the extent to which among-individual variation affects the dynamics of populations requires the study of multiple populations. In this study, we examined variation in the dynamics of populations of a live-bearing, marine fish (black surfperch; Embiotoca jacksoni). In collaboration with an organization of citizen scientists (Reef Check California), we were able to examine the dynamics of eight populations that were distributed throughout approximately 700 km of coastline, a distance that encompasses much of this species' range. We hypothesized that genetic variation within a local population would be related to the intensity of competition and to the strength of population regulation. To test this hypothesis, we examined whether genetic diversity (measured by the diversity of mitochondrial DNA haplotypes) was related to the strength of population regulation. Low-diversity populations experienced strong density dependence in population growth rates and population sizes were regulated much more tightly than they were in high-diversity populations. Mechanisms that contributed to this pattern include links between genetic diversity, habitat use, and spatial crowding. On average, low-diversity populations used less of the available habitat and exhibited greater spatial clustering (and more intense competition) for a given level of density (measured at the scale of the reef). Although the populations we studied also varied with respect to exogenous characteristics (habitat complexity, densities of predators, and interspecific competitors), none of these characteristics was significantly related to the strength of population regulation. In contrast, an endogenous characteristic of the population (genetic diversity) explained 77% of the variation in the strength of population regulation (95% CI: 27-94%). Our results suggest that the genetic and phenotypic composition of populations can play a major role in their dynamics.

  7. ELUCID. IV. Galaxy Quenching and its Relation to Halo Mass, Environment, and Assembly Bias

    NASA Astrophysics Data System (ADS)

    Wang, Huiyuan; Mo, H. J.; Chen, Sihan; Yang, Yang; Yang, Xiaohu; Wang, Enci; van den Bosch, Frank C.; Jing, Yipeng; Kang, Xi; Lin, Weipeng; Lim, S. H.; Huang, Shuiyao; Lu, Yi; Li, Shijie; Cui, Weiguang; Zhang, Youcai; Tweed, Dylan; Wei, Chengliang; Li, Guoliang; Shi, Feng

    2018-01-01

    We examine the quenched fraction of central and satellite galaxies as a function of galaxy stellar mass, halo mass, and the matter density of their large-scale environment. Matter densities are inferred from our ELUCID simulation, a constrained simulation of the local universe sampled by SDSS, while halo masses and central/satellite classification are taken from the galaxy group catalog of Yang et al. The quenched fraction for the total population increases systematically with the three quantities. We find that the “environmental quenching efficiency,” which quantifies the quenched fraction as a function of halo mass, is independent of stellar mass. And this independence is the origin of the stellar mass independence of density-based quenching efficiency found in previous studies. Considering centrals and satellites separately, we find that the two populations follow similar correlations of quenching efficiency with halo mass and stellar mass, suggesting that they have experienced similar quenching processes in their host halo. We demonstrate that satellite quenching alone cannot account for the environmental quenching efficiency of the total galaxy population, and that the difference between the two populations found previously arises mainly from the fact that centrals and satellites of the same stellar mass reside, on average, in halos of different mass. After removing these effects of halo mass and stellar mass, there remains a weak, but significant, residual dependence on environmental density, which is eliminated when halo assembly bias is taken into account. Our results therefore indicate that halo mass is the prime environmental parameter that regulates the quenching of both centrals and satellites.

  8. Calibrating abundance indices with population size estimators of red back salamanders (Plethodon cinereus) in a New England forest

    PubMed Central

    Ellison, Aaron M.; Jackson, Scott

    2015-01-01

    Herpetologists and conservation biologists frequently use convenient and cost-effective, but less accurate, abundance indices (e.g., number of individuals collected under artificial cover boards or during natural objects surveys) in lieu of more accurate, but costly and destructive, population size estimators to detect and monitor size, state, and trends of amphibian populations. Although there are advantages and disadvantages to each approach, reliable use of abundance indices requires that they be calibrated with accurate population estimators. Such calibrations, however, are rare. The red back salamander, Plethodon cinereus, is an ecologically useful indicator species of forest dynamics, and accurate calibration of indices of salamander abundance could increase the reliability of abundance indices used in monitoring programs. We calibrated abundance indices derived from surveys of P. cinereus under artificial cover boards or natural objects with a more accurate estimator of their population size in a New England forest. Average densities/m2 and capture probabilities of P. cinereus under natural objects or cover boards in independent, replicate sites at the Harvard Forest (Petersham, Massachusetts, USA) were similar in stands dominated by Tsuga canadensis (eastern hemlock) and deciduous hardwood species (predominantly Quercus rubra [red oak] and Acer rubrum [red maple]). The abundance index based on salamanders surveyed under natural objects was significantly associated with density estimates of P. cinereus derived from depletion (removal) surveys, but underestimated true density by 50%. In contrast, the abundance index based on cover-board surveys overestimated true density by a factor of 8 and the association between the cover-board index and the density estimates was not statistically significant. We conclude that when calibrated and used appropriately, some abundance indices may provide cost-effective and reliable measures of P. cinereus abundance that could be used in conservation assessments and long-term monitoring at Harvard Forest and other northeastern USA forests. PMID:26020008

  9. Accurate heterogeneous dose calculation for lung cancer patients without high‐resolution CT densities

    PubMed Central

    Li, Jonathan G.; Liu, Chihray; Olivier, Kenneth R.; Dempsey, James F.

    2009-01-01

    The aim of this study was to investigate the relative accuracy of megavoltage photon‐beam dose calculations employing either five bulk densities or independent voxel densities determined by calibration of the CT Houndsfield number. Full‐resolution CT and bulk density treatment plans were generated for 70 lung or esophageal cancer tumors (66 cases) using a commercial treatment planning system with an adaptive convolution dose calculation algorithm (Pinnacle3, Philips Medicals Systems). Bulk densities were applied to segmented regions. Individual and population average densities were compared to the full‐resolution plan for each case. Monitor units were kept constant and no normalizations were employed. Dose volume histograms (DVH) and dose difference distributions were examined for all cases. The average densities of the segmented air, lung, fat, soft tissue, and bone for the entire set were found to be 0.14, 0.26, 0.89, 1.02, and 1.12 g/cm3, respectively. In all cases, the normal tissue DVH agreed to better than 2% in dose. In 62 of 70 DVHs of the planning target volume (PTV), agreement to better than 3% in dose was observed. Six cases demonstrated emphysema, one with bullous formations and one with a hiatus hernia having a large volume of gas. These required the additional assignment of density to the emphysemic lung and inflammatory changes to the lung, the regions of collapsed lung, the bullous formations, and the hernia gas. Bulk tissue density dose calculation provides an accurate method of heterogeneous dose calculation. However, patients with advanced emphysema may require high‐resolution CT studies for accurate treatment planning. PACS number: 87.53.Tf

  10. Blood Density Is Nearly Equal to Water Density: A Validation Study of the Gravimetric Method of Measuring Intraoperative Blood Loss

    PubMed Central

    Vitello, Dominic J.; Ripper, Richard M.; Fettiplace, Michael R.; Weinberg, Guy L.; Vitello, Joseph M.

    2015-01-01

    Purpose. The gravimetric method of weighing surgical sponges is used to quantify intraoperative blood loss. The dry mass minus the wet mass of the gauze equals the volume of blood lost. This method assumes that the density of blood is equivalent to water (1 gm/mL). This study's purpose was to validate the assumption that the density of blood is equivalent to water and to correlate density with hematocrit. Methods. 50 µL of whole blood was weighed from eighteen rats. A distilled water control was weighed for each blood sample. The averages of the blood and water were compared utilizing a Student's unpaired, one-tailed t-test. The masses of the blood samples and the hematocrits were compared using a linear regression. Results. The average mass of the eighteen blood samples was 0.0489 g and that of the distilled water controls was 0.0492 g. The t-test showed P = 0.2269 and R 2 = 0.03154. The hematocrit values ranged from 24% to 48%. The linear regression R 2 value was 0.1767. Conclusions. The R 2 value comparing the blood and distilled water masses suggests high correlation between the two populations. Linear regression showed the hematocrit was not proportional to the mass of the blood. The study confirmed that the measured density of blood is similar to water. PMID:26464949

  11. Effect of experimental manipulation on survival and recruitment of feral pigs

    USGS Publications Warehouse

    Hanson, L.B.; Mitchell, M.S.; Grand, J.B.; Jolley, D.B.; Sparklin, B.D.; Ditchkoff, S.S.

    2009-01-01

    Lethal removal is commonly used to reduce the density of invasive-species populations, presuming it reduces population growth rate; the actual effect of lethal removal on the vital rates contributing to population growth, however, is rarely tested. We implemented a manipulative experiment of feral pig (Sus scrofa) populations at Fort Benning, Georgia, USA, to assess the demographic effects of harvest intensity. Using markrecapture data, we estimated annual survival, recruitment, and population growth rates of populations in a moderately harvested area and a heavily harvested area for 200406. Population growth rates did not differ between the populations. The top-ranked model for survival included a harvest intensity effect; model-averaged survival was lower for the heavily harvested population than for the moderately harvested population. Increased immigration and reproduction likely compensated for the increased mortality in the heavily harvested population. We conclude that compensatory responses in feral pig recruitment can limit the success of lethal control efforts. ?? 2009 CSIRO.

  12. Impact of Typhoon Haiyan on a Philippine Tarsier Population.

    PubMed

    Gursky, Sharon; Salibay, Cristina; Grow, Nanda; Fields, Lori

    2017-01-01

    Over the last 2 decades the Philippine tarsier (Carlito syrichta aka Tarsius syrichta) has had its conservation status revised from Endangered to Data Deficient to Near Threatened. The last status change was based on a study of the species' population density, which suggested that a single natural catastrophe could potentially wipe out the Philippine tarsier. In 2013 typhoon Haiyan hit Bohol, one of the island strongholds for this species. In this study we compare the density of the Bohol tarsier population within the Philippine Tarsier and Wildlife Sanctuary before and after the typhoon. We demonstrate that the typhoon significantly affected the density of the Philippine tarsier in the sanctuary. Before the typhoon, tarsier density was approximately 157 individuals/km2 whereas after the typhoon the density was a mere 36 individuals/km2. Prior to the typhoon, more Philippine tarsiers were found in older secondary forest than in younger secondary forest, whereas after the typhoon all observed individuals were found in relatively younger secondary forest. Vegetation plots where we observed Philippine tarsiers prior to the typhoon contained a mean of 33 trees/m2, with a mean diameter at breast height (DBH) of 24 cm, and a mean height of 4 m. After the typhoon vegetation plots contained an average of 156 trees, had a mean DBH of 6 cm, and a mean height of 2 m. Based on the IUCN Red List criteria, the reduction and fluctuation in the density of this species suggests that the conservation status of the Philippine tarsier should be changed to Vulnerable. This study indicates natural disasters can have a significant effect on the extinction risk of primates, with implications for future effects of anthropogenic climate change. © 2017 S. Karger AG, Basel.

  13. Reducing energy intake and energy density for a sustainable diet: a study based on self-selected diets in French adults.

    PubMed

    Masset, Gabriel; Vieux, Florent; Verger, Eric Olivier; Soler, Louis-Georges; Touazi, Djilali; Darmon, Nicole

    2014-06-01

    Studies on theoretical diets are not sufficient to implement sustainable diets in practice because of unknown cultural acceptability. In contrast, self-selected diets can be considered culturally acceptable. The objective was to identify the most sustainable diets consumed by people in everyday life. The diet-related greenhouse gas emissions (GHGE) for self-selected diets of 1918 adults participating in the cross-sectional French national dietary survey Individual and National Survey on Food Consumption (INCA2) were estimated. "Lower-Carbon," "Higher-Quality," and "More Sustainable" diets were defined as having GHGE lower than the overall median value, a probability of adequate nutrition intake (PANDiet) score (a measure of the overall nutritional adequacy of a diet) higher than the overall median value, and a combination of both criteria, respectively. Diet cost, as a proxy for affordability, and energy density were also assessed. More Sustainable diets were consumed by 23% of men and 20% of women, and their GHGE values were 19% and 17% lower than the population average (mean) value, respectively. In comparison with the average value, Lower-Carbon diets achieved a 20% GHGE reduction and lower cost, but they were not sustainable because they had a lower PANDiet score. Higher-Quality diets were not sustainable because of their above-average GHGE and cost. More Sustainable diets had an above-average PANDiet score and a below-average energy density, cost, GHGE, and energy content; the energy share of plant-based products was increased by 20% and 15% compared with the average for men and women, respectively. A strength of this study was that most of the dimensions for "sustainable diets" were considered, ie, not only nutritional quality and GHGE but also affordability and cultural acceptability. A reduction in diet-related GHGE by 20% while maintaining high nutritional quality seems realistic. This goal could be achieved at no extra cost by reducing energy intake and energy density and increasing the share of plant-based products. © 2014 American Society for Nutrition.

  14. Extra-metabolic energy use and the rise in human hyper-density

    NASA Astrophysics Data System (ADS)

    Burger, Joseph R.; Weinberger, Vanessa P.; Marquet, Pablo A.

    2017-03-01

    Humans, like all organisms, are subject to fundamental biophysical laws. Van Valen predicted that, because of zero-sum dynamics, all populations of all species in a given environment flux the same amount of energy on average. Damuth’s ’energetic equivalence rule’ supported Van Valen´s conjecture by showing a tradeoff between few big animals per area with high individual metabolic rates compared to abundant small species with low energy requirements. We use metabolic scaling theory to compare variation in densities and individual energy use in human societies to other land mammals. We show that hunter-gatherers occurred at densities lower than the average for a mammal of our size. Most modern humans, in contrast, concentrate in large cities at densities up to four orders of magnitude greater than hunter-gatherers, yet consume up to two orders of magnitude more energy per capita. Today, cities across the globe flux greater energy than net primary productivity on a per area basis. This is possible by importing enormous amounts of energy and materials required to sustain hyper-dense, modern humans. The metabolic rift with nature created by modern cities fueled largely by fossil energy poses formidable challenges for establishing a sustainable relationship on a rapidly urbanizing, yet finite planet.

  15. Extra-metabolic energy use and the rise in human hyper-density.

    PubMed

    Burger, Joseph R; Weinberger, Vanessa P; Marquet, Pablo A

    2017-03-02

    Humans, like all organisms, are subject to fundamental biophysical laws. Van Valen predicted that, because of zero-sum dynamics, all populations of all species in a given environment flux the same amount of energy on average. Damuth's 'energetic equivalence rule' supported Van Valen´s conjecture by showing a tradeoff between few big animals per area with high individual metabolic rates compared to abundant small species with low energy requirements. We use metabolic scaling theory to compare variation in densities and individual energy use in human societies to other land mammals. We show that hunter-gatherers occurred at densities lower than the average for a mammal of our size. Most modern humans, in contrast, concentrate in large cities at densities up to four orders of magnitude greater than hunter-gatherers, yet consume up to two orders of magnitude more energy per capita. Today, cities across the globe flux greater energy than net primary productivity on a per area basis. This is possible by importing enormous amounts of energy and materials required to sustain hyper-dense, modern humans. The metabolic rift with nature created by modern cities fueled largely by fossil energy poses formidable challenges for establishing a sustainable relationship on a rapidly urbanizing, yet finite planet.

  16. Seasonal population dynamics of Homalodisca vitripennis (Hemiptera: Cicadellidae) in sweet orange trees maintained under continuous deficit irrigation.

    PubMed

    Krugner, Rodrigo; Groves, Russell L; Johnson, Marshall W; Flores, Arnel P; Hagler, James R; Morse, Joseph G

    2009-06-01

    A 2-yr study was conducted in a citrus orchard (Citrus sinensis L. Osbeck cultivar Valencia) to determine the influence of plant water stress on the population dynamics of glassy-winged sharpshooter, Homalodisca vitripennis (Germar). Experimental treatments included irrigation at 100% of the crop evapotranspiration rate (ET(c)) and continuous deficit-irrigation regimens at 80 and 60% ET(c). Microclimate and plant conditions monitored included temperature and humidity in the tree canopy, leaf surface temperature, water potential, and fruit quality and yield. Glassy-winged sharpshooter population densities and activity were monitored weekly by a combination of visual inspections, beat net sampling, and trapping. Glassy-winged sharpshooter populations were negatively affected by severe plant water stress; however, population densities were not linearly related to decreasing water availability in plants. Citrus trees irrigated at 60% ET(c) had significantly warmer leaves, lower xylem water potential, and consequently hosted fewer glassy-winged sharpshooter eggs, nymphs, and adults than trees irrigated at 80% ET(c). Citrus trees irrigated at 100% ET(c) hosted similar numbers of glassy-winged sharpshooter stages as trees irrigated at 60% ET(c) and a lower number of glassy-winged sharpshooter nymphs than the 80% ET(c) treatment, specifically during the nymphal density peak in mid-April to early July. Irrigation treatments did not affect populations of monitored natural enemies. Although the adult glassy-winged sharpshooter population was reduced, on average, by 50% in trees under severe water stress, the total number of fruit and number of fruit across several fruit grade categories were significantly lower in the 60% ET(c) than in the 80 and 100% ET(c) irrigation treatments.

  17. Effects of plantation density on wood density and anatomical properties of red pine (Pinus resinosa Ait.)

    Treesearch

    J. Y. Zhu; C. Tim Scott; Karen L. Scallon; Gary C. Myers

    2007-01-01

    This study demonstrated that average ring width (or average annual radial growth rate) is a reliable parameter to quantify the effects of tree plantation density (growth suppression) on wood density and tracheid anatomical properties. The average ring width successfully correlated wood density and tracheid anatomical properties of red pines (Pinus resinosa Ait.) from a...

  18. [Effective size of subpopulation of the early run sockeye salmon Oncorhynchus nerka from Azabach's Lake (Kamchatka): effect of density on variance of reproductive success].

    PubMed

    Efremov, V V; Parenskiĭ, V A

    2004-04-01

    Using Parensky's approach for estimating the number of breeding pairs, we determined effective subpopulation size Ne in early-run sockeye salmon Oncorhynchus nerka from Azabach'e Lake (Kamchatka) in 1977 through 1981. On average (over years and populations), biased sex ratio decreased Ne by 7% as compared to the number of fish on the spawning sites (Ni). High density reduced the Ne/Ni ratio by 62-66% because some fish were excluded from spawning. Dominance polygyny as compared to monogamy and random union of gametes could reduce Ne by about 17%.

  19. Spontaneous and controlled-diameter synthesis of single-walled and few-walled carbon nanotubes

    NASA Astrophysics Data System (ADS)

    Inoue, Shuhei; Lojindarat, Supanat; Kawamoto, Takahiro; Matsumura, Yukihiko; Charinpanitkul, Tawatchai

    2018-05-01

    In this study, we explored the spontaneous and controlled-diameter growth of carbon nanotubes. We evaluated the effects of catalyst density, reduction time, and a number of catalyst coating on the substrate (for multi-walled carbon nanotubes) on the diameter of single-walled carbon nanotubes and the number of layers in few-walled carbon nanotubes. Increasing the catalyst density and reduction time increased the diameters of the carbon nanotubes, with the average diameter increasing from 1.05 nm to 1.86 nm for single-walled carbon nanotubes. Finally, we succeeded in synthesizing a significant double-walled carbon nanotube population of 24%.

  20. A Comparison of Grizzly Bear Demographic Parameters Estimated from Non-Spatial and Spatial Open Population Capture-Recapture Models

    PubMed Central

    Whittington, Jesse; Sawaya, Michael A.

    2015-01-01

    Capture-recapture studies are frequently used to monitor the status and trends of wildlife populations. Detection histories from individual animals are used to estimate probability of detection and abundance or density. The accuracy of abundance and density estimates depends on the ability to model factors affecting detection probability. Non-spatial capture-recapture models have recently evolved into spatial capture-recapture models that directly include the effect of distances between an animal’s home range centre and trap locations on detection probability. Most studies comparing non-spatial and spatial capture-recapture biases focussed on single year models and no studies have compared the accuracy of demographic parameter estimates from open population models. We applied open population non-spatial and spatial capture-recapture models to three years of grizzly bear DNA-based data from Banff National Park and simulated data sets. The two models produced similar estimates of grizzly bear apparent survival, per capita recruitment, and population growth rates but the spatial capture-recapture models had better fit. Simulations showed that spatial capture-recapture models produced more accurate parameter estimates with better credible interval coverage than non-spatial capture-recapture models. Non-spatial capture-recapture models produced negatively biased estimates of apparent survival and positively biased estimates of per capita recruitment. The spatial capture-recapture grizzly bear population growth rates and 95% highest posterior density averaged across the three years were 0.925 (0.786–1.071) for females, 0.844 (0.703–0.975) for males, and 0.882 (0.779–0.981) for females and males combined. The non-spatial capture-recapture population growth rates were 0.894 (0.758–1.024) for females, 0.825 (0.700–0.948) for males, and 0.863 (0.771–0.957) for both sexes. The combination of low densities, low reproductive rates, and predominantly negative population growth rates suggest that Banff National Park’s population of grizzly bears requires continued conservation-oriented management actions. PMID:26230262

  1. The StreamCat Dataset: Accumulated Attributes for NHDPlusV2 Catchments (Version 2.1) for the Conterminous United States: 2010 US Census Housing Unit and Population Density

    EPA Pesticide Factsheets

    This dataset represents the population and housing unit density within individual, local NHDPlusV2 catchments and upstream, contributing watersheds based on 2010 US Census data. Densities are calculated for every block group and watershed averages are calculated for every local NHDPlusV2 catchment(see Data Sources for links to NHDPlusV2 data and Census Data). This data set is derived from The TIGER/Line Files and related database (.dbf) files for the conterminous USA. It was downloaded as Block Group-Level Census 2010 SF1 Data in File Geodatabase Format (ArcGIS version 10.0). The landscape raster (LR) was produced based on the data compiled from the questions asked of all people and about every housing unit. The (block-group population / block group area) and (block-group housing units / block group area) were summarized by local catchment and by watershed to produce local catchment-level and watershed-level metrics as a continuous data type (see Data Structure and Attribute Information for a description). Using a riparian buffer(see Process Steps), statistics were generated for areas within each catchment that are within 100 meters of the stream reach in an attempt to evaluate for the riparian zone.

  2. Use of geographic information systems in rabies vaccination campaigns.

    PubMed

    Grisi-Filho, José Henrique de Hildebrand e; Amaku, Marcos; Dias, Ricardo Augusto; Montenegro Netto, Hildebrando; Paranhos, Noemia Tucunduva; Mendes, Maria Cristina Novo Campos; Ferreira Neto, José Soares; Ferreira, Fernando

    2008-12-01

    To develop a method to assist in the design and assessment of animal rabies control campaigns. A methodology was developed based on geographic information systems to estimate the animal (canine and feline) population and density per census tract and per subregion (known as "Subprefeituras") in the city of São Paulo (Southeastern Brazil) in 2002. The number of vaccination units in a given region was estimated to achieve a certain proportion of vaccination coverage. Census database was used for the human population, as well as estimates ratios of dog:inhabitant and cat:inhabitant. Estimated figures were 1,490,500 dogs and 226,954 cats in the city, i.e. an animal population density of 1138.14 owned animals per km(2). In the 2002 campaign, 926,462 were vaccinated, resulting in a vaccination coverage of 54%. The estimated number of vaccination units to be able to reach a 70%-vaccination coverage, by vaccinating 700 animals per unit on average, was 1,729. These estimates are presented as maps of animal density according to census tracts and "Subprefeituras". The methodology used in the study may be applied in a systematic way to the design and evaluation of rabies vaccination campaigns, enabling the identification of areas of critical vaccination coverage.

  3. Rhizobacterial population density and nitrogen fixation in seagrass community of Gulf of Mannar, India.

    PubMed

    Raja, S; Thangaradjou, T; Sivakumar, K; Kannan, L

    2012-11-01

    Seagrass rhizosphere generally supports high bacterial population density which plays a major role in determining the nutrient cycles of the sea. Higher densities of total heterotrphic bacteria (26.3 x 10(6) CFU g(-1)), nitrogen fixing (27.3 x 10(3) CFUg(-1), ammonifying (44.66 x 10(6) MPN g(-1)) and nitrifying bacteria (42.33 X 10(6) MPN g(-1)) have been registered in the seagrass areas than the non seagrass area. In particular, all these rhizosphere microbial population was higher in Thalassia hemprichii. The rates of nitrogen fixation was recorded in the different species of seagrasses such as Enhalus acoroides (1.166 n mol g(-1) d(-1)), Halophila ovalis (0.166 n mol g(-1) d(-1)), Thalassia hemprichii(18.5 n mol g(-1) d(-1)), Cymodocea serrulata (10.5 n mol g(-1) d(-1)), Halodule uninervis (5.375 n mol g(-1) d(-1)) and Syringodium isoetifolium (0.666 n mol g(-1) d(-1)) using gas chromatography. The average nitrogen fixation by the seagrasses of Gulf of Mannar alone was estimated to be 7640.58 n mol m(-2) d(-1) and the contributions from the rhizosphere microbes will increase the quantity to many fold.

  4. Peptide crystal simulations reveal hidden dynamics

    PubMed Central

    Janowski, Pawel A.; Cerutti, David S.; Holton, James; Case, David A.

    2013-01-01

    Molecular dynamics simulations of biomolecular crystals at atomic resolution have the potential to recover information on dynamics and heterogeneity hidden in the X-ray diffraction data. We present here 9.6 microseconds of dynamics in a small helical peptide crystal with 36 independent copies of the unit cell. The average simulation structure agrees with experiment to within 0.28 Å backbone and 0.42 Å all-atom rmsd; a model refined against the average simulation density agrees with the experimental structure to within 0.20 Å backbone and 0.33 Å all-atom rmsd. The R-factor between the experimental structure factors and those derived from this unrestrained simulation is 23% to 1.0 Å resolution. The B-factors for most heavy atoms agree well with experiment (Pearson correlation of 0.90), but B-factors obtained by refinement against the average simulation density underestimate the coordinate fluctuations in the underlying simulation where the simulation samples alternate conformations. A dynamic flow of water molecules through channels within the crystal lattice is observed, yet the average water density is in remarkable agreement with experiment. A minor population of unit cells is characterized by reduced water content, 310 helical propensity and a gauche(−) side-chain rotamer for one of the valine residues. Careful examination of the experimental data suggests that transitions of the helices are a simulation artifact, although there is indeed evidence for alternate valine conformers and variable water content. This study highlights the potential for crystal simulations to detect dynamics and heterogeneity in experimental diffraction data, as well as to validate computational chemistry methods. PMID:23631449

  5. Productivity, mortality, and population trends of wolves in northeastern Minnesota

    USGS Publications Warehouse

    Mech, L.D.

    1977-01-01

    Population parameters, mortality causes, and mechanisms of a population decline were studied in wolves (Canis lupus lycaon) from 1968 to 1976 in the Superior National Forest. The main method was aerial radio-tracking of 129 wolves and their packmates. Due to a decline in white-tailed deer (Odocoileus virginianus), the wolf population decreased during most of the study. Average annual productivity varied from 1.5 to 3.3 pups per litter, and annual mortality rates from 7 to 65 percent. Malnutrition and intraspecific strife accounted equally for 58 percent of the mortality; human causes accounted for the remainder. As wolf numbers began to decline, pup starvation became apparent, followed by lower pup production, and then by increased intraspecific strife. At higher densities, adult pack wolves were the most secure members of the population, but as the population declined, they became the least secure because of intraspecific strife.

  6. Sink populations in carnivore management: cougar demography and immigration in a hunted population.

    PubMed

    Robinson, Hugh S; Wielgus, Robert B; Cooley, Hilary S; Cooley, Skye W

    2008-06-01

    Carnivores are widely hunted for both sport and population control, especially where they conflict with human interests. It is widely believed that sport hunting is effective in reducing carnivore populations and related human-carnivore conflicts, while maintaining viable populations. However, the way in which carnivore populations respond to harvest can vary greatly depending on their social structure, reproductive strategies, and dispersal patterns. For example, hunted cougar (Puma concolor) populations have shown a great degree of resiliency. Although hunting cougars on a broad geographic scale (> 2000 km2) has reduced densities, hunting of smaller areas (i.e., game management units, < 1000 km2), could conceivably fail because of increased immigration from adjacent source areas. We monitored a heavily hunted population from 2001 to 2006 to test for the effects of hunting at a small scale (< 1000 km2) and to gauge whether population control was achieved (lambda < or = 1.0) or if hunting losses were negated by increased immigration allowing the population to remain stable or increase (lambda > or = 1.0). The observed growth rate of 1.00 was significantly higher than our predicted survival/fecundity growth rates (using a Leslie matrix) of 0.89 (deterministic) and 0.84 (stochastic), with the difference representing an 11-16% annual immigration rate. We observed no decline in density of the total population or the adult population, but a significant decrease in the average age of independent males. We found that the male component of the population was increasing (observed male population growth rate, lambda(OM) = 1.09), masking a decrease in the female component (lambda(OF) = 0.91). Our data support the compensatory immigration sink hypothesis; cougar removal in small game management areas (< 1000 km2) increased immigration and recruitment of younger animals from adjacent areas, resulting in little or no reduction in local cougar densities and a shift in population structure toward younger animals. Hunting in high-quality habitats may create an attractive sink, leading to misinterpretation of population trends and masking population declines in the sink and surrounding source areas.

  7. Non-sky-averaged sensitivity curves for space-based gravitational-wave observatories

    NASA Astrophysics Data System (ADS)

    Vallisneri, Michele; Galley, Chad R.

    2012-06-01

    The signal-to-noise ratio (SNR) is used in gravitational-wave observations as the basic figure of merit for detection confidence and, together with the Fisher matrix, for the amount of physical information that can be extracted from a detected signal. SNRs are usually computed from a sensitivity curve, which describes the gravitational-wave amplitude needed by a monochromatic source of given frequency to achieve a threshold SNR. Although the term ‘sensitivity’ is used loosely to refer to the detector’s noise spectral density, the two quantities are not the same: the sensitivity includes also the frequency- and orientation-dependent response of the detector to gravitational waves and takes into account the duration of observation. For interferometric space-based detectors similar to LISA, which are sensitive to long-lived signals and have constantly changing position and orientation, exact SNRs need to be computed on a source-by-source basis. For convenience, most authors prefer to work with sky-averaged sensitivities, accepting inaccurate SNRs for individual sources and giving up control over the statistical distribution of SNRs for source populations. In this paper, we describe a straightforward end-to-end recipe to compute the non-sky-averaged sensitivity of interferometric space-based detectors of any geometry. This recipe includes the effects of spacecraft motion and of seasonal variations in the partially subtracted confusion foreground from Galactic binaries, and it can be used to generate a sampling distribution of sensitivities for a given source population. In effect, we derive error bars for the sky-averaged sensitivity curve, which provide a stringent statistical interpretation for previously unqualified statements about sky-averaged SNRs. As a worked-out example, we consider isotropic and Galactic-disk populations of monochromatic sources, as observed with the ‘classic LISA’ configuration. We confirm that the (standard) inverse-rms average sensitivity for the isotropic population remains the same whether or not the LISA orbits are included in the computation. However, detector motion tightens the distribution of sensitivities, so for 50% of sources the sensitivity is within 30% of its average. For the Galactic-disk population, the average and the distribution of the sensitivity for a moving detector turn out to be similar to the isotropic case.

  8. Population structure of an invasive parthenogenetic gastropod in coastal lakes and estuaries of northern KwaZulu-Natal, South Africa.

    PubMed

    Miranda, Nelson A F; Perissinotto, Renzo; Appleton, Christopher C

    2011-01-01

    Estuaries and coastal lakes receive little attention despite being heavily invaded by non-indigenous invasive species (NIS). In these situations, studies of population dynamics in invaded habitats can provide valuable insights into how NIS interact with new environments. Tarebia granifera is a prosobranch gastropod from south-east Asia which has invaded other sub-tropical parts of the world. This study addresses whether a small number of key environmental factors influences gastropod communities, and specifically how the population density and size structure of T. granifera were influenced by environmental change in estuaries and coastal lakes in southern Africa. T. granifera's density, number of brooded juveniles and size structure were measured at the St. Lucia Estuary, Mgobozeleni Estuary, Lake Sibaya and Lake Nhlange. Size structure was classified according to shell height (SH). All dissected individuals were found to be female and free from trematode infection. Salinity, water depth, temperature, and pH were the main factors correlated with population density of gastropod communities. T. granifera often reached densities well over 1000 ind. m(-2), displacing indigenous gastropods and becoming a dominant component of the benthic community. T. granifera successfully invaded estuaries despite frequent exposure to high salinity and desiccation, which could together eliminate >97% of the population. The persistence of T. granifera was ensured due to its high fecundity and the environmental tolerance of large adults (20-30 mm SH) which carried an average of 158±12.8 SD brooded juveniles. Repeat introductions were not essential for the success of this parthenogenetic NIS. There is a need for a broader study on the reproductive biology of T. granifera (including the previously overlooked "brood pouch ecology"), which affects population dynamics and may be relevant to other parthenogenetic NIS, such as Melanoides tuberculata and Potamopyrgus antipodarum.

  9. An ecological study of the locations of schools and child pedestrian injury collisions.

    PubMed

    LaScala, Elizabeth A; Gruenewald, Paul J; Johnson, Fred W

    2004-07-01

    Geographic studies of the incidence and prevalence of child pedestrian injury collisions in different community environments have been primarily descriptive and idiosyncratic, reflecting one or another likely determinant of the places where these injuries occur. The current study maintains that multiple determinants of child pedestrian injury collisions must be considered in evaluating the unique contributions of any one community feature to injury rates. These features include local characteristics of populations, such as rates of unemployment, and places, such as locations of schools. Schools are one stable geographic feature associated with regular, often concentrated periods of complex and congested traffic patterns. The objective of the present study was to examine annual rates of child pedestrian injury in four California communities with a focus on the unique contribution of schools to injury risk. We predicted that annual numbers of child pedestrian injury collisions (both in-school and summer combined) would be greater in communities with higher youth population densities, more unemployment, fewer high-income households, and higher traffic flow. It was hypothesized that youth population density and its interaction with the number of schools in a given area would be related to greater rates of child pedestrian collisions during in-school months. An ecological approach was taken that divided the four communities into 102 geographic units with an average of 6321 people residing in each unit. Archival data on traffic flow, number of child pedestrian injury collisions and locations of schools were obtained from state agencies. Individual-level data were obtained from a general population survey conducted in the communities. The results showed that annual numbers of injuries were greater in areas with higher youth population densities, more unemployment, fewer high-income households, and greater traffic flow. Annual numbers of injuries during in-school months were greater in areas containing middle schools and greater population densities of youth.

  10. Local variations in the timing of RSV epidemics.

    PubMed

    Noveroske, Douglas B; Warren, Joshua L; Pitzer, Virginia E; Weinberger, Daniel M

    2016-11-11

    Respiratory syncytial virus (RSV) is a primary cause of hospitalizations in children worldwide. The timing of seasonal RSV epidemics needs to be known in order to administer prophylaxis to high-risk infants at the appropriate time. We used data from the Connecticut State Inpatient Database to identify RSV hospitalizations based on ICD-9 diagnostic codes. Harmonic regression analyses were used to evaluate RSV epidemic timing at the county level and ZIP code levels. Linear regression was used to investigate associations between the socioeconomic status of a locality and RSV epidemic timing. 9,740 hospitalizations coded as RSV occurred among children less than 2 years old between July 1, 1997 and June 30, 2013. The earliest ZIP code had a seasonal RSV epidemic that peaked, on average, 4.64 weeks earlier than the latest ZIP code. Earlier epidemic timing was significantly associated with demographic characteristics (higher population density and larger fraction of the population that was black). Seasonal RSV epidemics in Connecticut occurred earlier in areas that were more urban (higher population density and larger fraction of the population that was). These findings could be used to better time the administration of prophylaxis to high-risk infants.

  11. Monoxenic liquid culture with Escherichia coli of the free-living nematode Panagrolaimus sp. (strain NFS 24-5), a potential live food candidate for marine fish and shrimp larvae.

    PubMed

    Ayub, Farhana; Seychelles, Laurent; Strauch, Olaf; Wittke, Martina; Ehlers, Ralf-Udo

    2013-09-01

    The free-living, bacterial-feeding nematode Panagrolaimus sp. (strain NFS 24-5) has potential for use as live food for marine shrimp and fish larvae. Mass production in liquid culture is a prerequisite for its commercial exploitation. Panagrolaimus sp. was propagated in monoxenic liquid culture on Escherichia coli and parameters, like nematode density, population dynamics and biomass were recorded and compared with life history table data. A mean maximum nematode density of 174,278 mL(-1) and a maximum of 251,000 mL(-1) were recorded on day 17 after inoculation. Highest average biomass was 40 g L(-1) at day 13. The comparison with life history table data indicated that the hypothetical potential of liquid culture is much higher than documented during this investigation. Nematode development is delayed in liquid culture and egg production per female is more than five times lower than reported from life history trait analysis. The latter assessed a nematode generation time of 7.1 days, whereas the process time at maximum nematode density in liquid culture was 16 days indicating that a reduction of the process time can be achieved by further investigating the influence of nematode inoculum density on population development. The results challenge future research to reduce process time and variability and improve population dynamics also during scale-up of the liquid culture process.

  12. Mathematical models for predicting human mobility in the context of infectious disease spread: introducing the impedance model.

    PubMed

    Sallah, Kankoé; Giorgi, Roch; Bengtsson, Linus; Lu, Xin; Wetter, Erik; Adrien, Paul; Rebaudet, Stanislas; Piarroux, Renaud; Gaudart, Jean

    2017-11-22

    Mathematical models of human mobility have demonstrated a great potential for infectious disease epidemiology in contexts of data scarcity. While the commonly used gravity model involves parameter tuning and is thus difficult to implement without reference data, the more recent radiation model based on population densities is parameter-free, but biased. In this study we introduce the new impedance model, by analogy with electricity. Previous research has compared models on the basis of a few specific available spatial patterns. In this study, we use a systematic simulation-based approach to assess the performances. Five hundred spatial patterns were generated using various area sizes and location coordinates. Model performances were evaluated based on these patterns. For simulated data, comparison measures were average root mean square error (aRMSE) and bias criteria. Modeling of the 2010 Haiti cholera epidemic with a basic susceptible-infected-recovered (SIR) framework allowed an empirical evaluation through assessing the goodness-of-fit of the observed epidemic curve. The new, parameter-free impedance model outperformed previous models on simulated data according to average aRMSE and bias criteria. The impedance model achieved better performances with heterogeneous population densities and small destination populations. As a proof of concept, the basic compartmental SIR framework was used to confirm the results obtained with the impedance model in predicting the spread of cholera in Haiti in 2010. The proposed new impedance model provides accurate estimations of human mobility, especially when the population distribution is highly heterogeneous. This model can therefore help to achieve more accurate predictions of disease spread in the context of an epidemic.

  13. Galaxy Downsizing Evidenced by Hybrid Evolutionary Tracks

    NASA Astrophysics Data System (ADS)

    Firmani, C.; Avila-Reese, V.

    2010-11-01

    The stellar-dark halo mass relation of galaxies at different redshifts, M s(M h, z), encloses relevant features concerning their physical processes and evolution. This sequence of relations, defined in the range 0 < z < 4, together with average Λ cold dark matter (ΛCDM) halo mass aggregation histories (MAHs), is used here for inferring average galaxian hybrid evolutionary tracks (GHETs), where "hybrid" remarks on the combination of observational (M s) and theoretical (M h) ingredients. As a result of our approach, a unified picture of stellar and halo mass buildup, population migration, and downsizing of galaxies as a function of mass is presented. The inferred average M s growth histories (GHETs) of the highest and lowest mass galaxies are definitively quite different from the average MAHs, M h(z), of the corresponding dark halos. Depending on how a given M h(z) compares with the mass at which the M s-to-M h ratio curve peaks at the epoch z, M hp(z), two evolutionary phases are evidenced: (1) galaxies in an active regime of M s growth when M h < M hp and (2) galaxies in a quiescent or passive regime when M h>M hp. The typical M s at which galaxies transit from the active (star-forming) to the quiescent regime, M tran, increases with z, log(M tran/M sun) ≈10.30 + 0.55z, making evident a population downsizing phenomenon. This result agrees with independent observational determinations based on the evolution of the galaxy stellar mass function decomposition into blue and red galaxy populations. The specific star formation rate (SSFR) predicted from the derivative of the GHET is consistent with direct measures of the SSFR for galaxies at different redshifts, though both sets of observational inferences are independent. The average GHETs of galaxies smaller than M tran at z = 0 (M s ≈1010.3 M sun, M h ≈1011.8 M sun) did not reach the quiescent regime, and for them, the lower the mass, the faster the later M s growth rate (downsizing in SSFR). The GHETs allow us to predict the transition rate in the number density of active to passive population; the predicted values agree with direct estimates of the growth rate in the number density for the (massive) red population up to z ~ 1. We show that ΛCDM-based models of disk galaxy evolution, including feedback-driven outflows, are able to reproduce the low-mass side of the M s-M h relation at z ~ 0, but at higher z strongly disagree with the GHETs: models do not reproduce the strong downsizing in SSFR and the high SSFR of low-mass galaxies.

  14. Quantifying temporal isolation: a modelling approach assessing the effect of flowering time differences on crop-to-weed pollen flow in sunflower

    PubMed Central

    Roumet, Marie; Cayre, Adeline; Latreille, Muriel; Muller, Marie-Hélène

    2015-01-01

    Flowering time divergence can be a crucial component of reproductive isolation between sympatric populations, but few studies have quantified its actual contribution to the reduction of gene flow. In this study, we aimed at estimating pollen-mediated gene flow between cultivated sunflower and a weedy conspecific sunflower population growing in the same field and at quantifying, how it is affected by the weeds' flowering time. For that purpose, we extended an existing mating model by including a temporal distance (i.e. flowering time difference between potential parents) effect on mating probabilities. Using phenological and genotypic data gathered on the crop and on a sample of the weedy population and its offspring, we estimated an average hybridization rate of approximately 10%. This rate varied strongly from 30% on average for weeds flowering at the crop flowering peak to 0% when the crop finished flowering and was affected by the local density of weeds. Our result also suggested the occurrence of other factors limiting crop-to-weed gene flow. This level of gene flow and its dependence on flowering time might influence the evolutionary fate of weedy sunflower populations sympatric to their crop relative. PMID:25667603

  15. The Gulf of Ambracia's Common Bottlenose Dolphins, Tursiops truncatus: A Highly Dense and yet Threatened Population.

    PubMed

    Gonzalvo, J; Lauriano, G; Hammond, P S; Viaud-Martinez, K A; Fossi, M C; Natoli, A; Marsili, L

    The common bottlenose dolphin (Tursiops truncatus) is the only cetacean present in the semiclosed waters of the Gulf of Ambracia, Western Greece. This increasingly degraded coastal ecosystem hosts one of the highest observed densities in the Mediterranean Sea for this species. Photo-identification data and tissue samples collected through skin-swabbing and remote biopsy sampling techniques during boat-based surveys conducted between 2006 and 2015 in the Gulf, were used to examine bottlenose dolphin abundance, population trends, site fidelity, genetic differentiation and toxicological status. Bottlenose dolphins showed high levels of year-round site fidelity throughout the 10-year study period. Dolphin population estimates mostly fell between 130 and 170 with CVs averaging about 10%; a trend in population size over the 10 years was a decline of 1.6% per year (but this was not significant). Genetic differentiation between the bottlenose dolphins of the Gulf and their conspecifics from neighbouring populations was detected, and low genetic diversity was found among individuals sampled. In addition, pesticides where identified as factors posing a real toxicological problem for local bottlenose dolphins. Therefore, in the Gulf of Ambracia, high dolphin density does not seem to be indicative of favourable conservation status or pristine habitat. © 2016 Elsevier Ltd All rights reserved.

  16. The 2006 Project Progress Report for 1987-099-00 Dworshak Kokanee Population and Entrainment Assessment (contract # 26850) is attached to project 2007-003-00, contract #31598. [POINTER

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    None

    2008-12-18

    During this contract, we continued testing underwater strobe lights to determine their effectiveness at repelling kokanee Oncorhynchus nerka away from Dworshak Dam. Strobe light tests were conducted on four nights from April 24-27, 2006, in front of the middle reservoir outlet (RO) 2. The density and distribution of fish, (thought to be mostly kokanee), were monitored with a split-beam echo sounder. We then compared fish counts and densities during nights when the lights were flashing to counts and densities during adjacent nights without the lights on. On two nights, April 25 and 27, 2006, when no lights were present, fishmore » counts near RO 2 averaged 12.4 fish and densities averaged 31.0 fish/ha. When strobe lights were turned on during the nights of April 24 and 26, mean counts dropped to 4.7 fish and densities dropped to 0.5 fish/ha. The decline in counts (62%) and densities (99%) was statistically significant (p = 0.009 and 0.002, respectively). Test results indicated that strobe lights were able to reduce fish densities by at least 50% in front of a discharging reservoir outlet, which would be sufficient to improve sport fish harvest. We also used split-beam hydroacoustics to monitor the kokanee population in Dworshak Reservoir during 2006. Estimated abundance of kokanee increased from the 2005 population estimate. Based on hydroacoustic surveys, we estimated approximately 5,815,000 kokanee (90% CI {+-} 27.6%) in Dworshak Reservoir in August 2006. This included 2,183,000 age-0 (90% CI {+-} 24.2%), 1,509,000 age-1 (90% CI {+-} 29.0%), and 2,124,000 age-2 (90% CI {+-} 27.6%) kokanee. This resulted in a density of age-2 kokanee above the management goal of 30-50 adults/ha. Entrainment sampling was conducted with fixed-site, split-beam hydroacoustics from May through September for a continuous 24 h period when dam operations permitted. The highest fish detection rates from entrainment assessments were found during dawn periods, unlike previous year's results, which were highest during nighttime. The lowest detection rate was found during the day period, which was consistent with previous findings. Fish detection rates were generally low during high discharges throughout the summer and highest during low discharges in May and June. Low detection rates were found during high discharge periods during drawdowns for anadromous fish flows in July and August, which resulted in low susceptibility to entrainment. Counts of spawning kokanee in four tributary streams totaled 29,743 fish. These data fit the previously developed relationship between spawner counts and adult kokanee abundance in the reservoir.« less

  17. Dworshak Kokanee Population and Engrainment Assessment : 2006 Annual Report, March 1, 2006 - February 28, 2007.

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Stark, Eric J.

    2008-12-18

    During this contract, we continued testing underwater strobe lights to determine their effectiveness at repelling kokanee Oncorhynchus nerka away from Dworshak Dam. Strobe light tests were conducted on four nights from April 24-27, 2006, in front of the middle reservoir outlet (RO) 2. The density and distribution of fish, (thought to be mostly kokanee), were monitored with a split-beam echo sounder. We then compared fish counts and densities during nights when the lights were flashing to counts and densities during adjacent nights without the lights on. On two nights, April 25 and 27, 2006, when no lights were present, fishmore » counts near RO 2 averaged 12.4 fish and densities averaged 31.0 fish/ha. When strobe lights were turned on during the nights of April 24 and 26, mean counts dropped to 4.7 fish and densities dropped to 0.5 fish/ha. The decline in counts (62%) and densities (99%) was statistically significant (p = 0.009 and 0.002, respectively). Test results indicated that strobe lights were able to reduce fish densities by at least 50% in front of a discharging reservoir outlet, which would be sufficient to improve sport fish harvest. We also used split-beam hydroacoustics to monitor the kokanee population in Dworshak Reservoir during 2006. Estimated abundance of kokanee increased from the 2005 population estimate. Based on hydroacoustic surveys, we estimated approximately 5,815,000 kokanee (90% CI {+-} 27.6%) in Dworshak Reservoir in August 2006. This included 2,183,000 age-0 (90% CI {+-} 24.2%), 1,509,000 age-1 (90% CI {+-} 29.0%), and 2,124,000 age-2 (90% CI {+-} 27.6%) kokanee. This resulted in a density of age-2 kokanee above the management goal of 30-50 adults/ha. Entrainment sampling was conducted with fixed-site, split-beam hydroacoustics from May through September for a continuous 24 h period when dam operations permitted. The highest fish detection rates from entrainment assessments were found during dawn periods, unlike previous year's results, which were highest during nighttime. The lowest detection rate was found during the day period, which was consistent with previous findings. Fish detection rates were generally low during high discharges throughout the summer and highest during low discharges in May and June. Low detection rates were found during high discharge periods during drawdowns for anadromous fish flows in July and August, which resulted in low susceptibility to entrainment. Counts of spawning kokanee in four tributary streams totaled 29,743 fish. These data fit the previously developed relationship between spawner counts and adult kokanee abundance in the reservoir.« less

  18. The 2005 Project Progress Report for 1987-099-00 Dworshak Kokanee Population and Entrainment Assessment (contract # 16791) is attached to project 1987-099-00, contract # 26850. [POINTER

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    None

    2008-12-18

    During this contract, we continued testing underwater strobe lights to determine their effectiveness at repelling kokanee Oncorhynchus nerka away from Dworshak Dam. We tested one set of nine strobe lights flashing at a rate of 360 flashes/min in front of turbine 3 while operating at higher discharges than previously tested. The density and distribution of fish, (thought to be mostly kokanee), were monitored with a split-beam echo sounder. We then compared fish counts and densities during nights when the lights were flashing to counts and densities during adjacent nights without the lights on. On five nights between January 31 andmore » February 28, 2006, when no lights were present, fish counts near turbine 3 averaged eight fish and densities averaged 91 fish/ha. When strobe lights were turned on during five adjacent nights during the same period, mean counts dropped to four fish and densities dropped to 35 fish/ha. The decline in counts (49%) was not statistically significant (p = 0.182), but decline in densities (62%) was significant (p = 0.049). There appeared to be no tendency for fish to habituate to the lights during the night. Test results indicated that strobe lights were able to reduce fish densities by at least 50% in front of turbines operating at higher discharges, which would be sufficient to improve sportfish harvest. We also used split-beam hydroacoustics to monitor the kokanee population in Dworshak Reservoir during 2005. Estimated abundance of kokanee decreased from the 2004 population estimate. Based on hydroacoustic surveys, we estimated 3,011,626 kokanee (90% CI {+-} 15.2%) in Dworshak Reservoir, July 2005. This included 2,135,986 age-0 (90% CI {+-} 15.9%), 769,175 age-1 (90% CI {+-} 16.0%), and 107,465 age-2 (90% CI {+-} 15.2%). Poor survival of kokanee from age-1 to age-2 continued to keep age-2 densities below the management goal of 30-50 adults/ha. Entrainment sampling was conducted with fixed-site split-beam hydroacoustics a minimum of two days per month for a continuous 24 h period when dam operations permitted. The highest fish detection rates from entrainment assessments were again found during nighttime periods and lowest during the day. Fish detection rates were low during high discharges throughout the spring and summer and highest during low discharges in September and November. High discharge during drawdowns for anadromous fish flows in July and August again resulted in low detection rates and susceptibility to entrainment. Index counts of spawning kokanee in four tributary streams totaled 12,742 fish. This data fits the previously developed relationship between spawner counts and adult kokanee abundance in the reservoir.« less

  19. Pollen-limited reproduction in blue oak: Implications for wind pollination in fragmented populations

    USGS Publications Warehouse

    Knapp, E.E.; Goedde, M.A.; Rice, K.J.

    2001-01-01

    Human activities are fragmenting forests and woodlands worldwide, but the impact of reduced tree population densities on pollen transfer in wind-pollinated trees is poorly understood. In a 4-year study, we evaluated relationships among stand density, pollen availability, and seed production in a thinned and fragmented population of blue oak (Quercus douglasii). Geographic coordinates were established and flowering interval determined for 100 contiguous trees. The number of neighboring trees within 60 m that released pollen during each tree's flowering period was calculated and relationships with acorn production explored using multiple regression. We evaluated the effects of female flower production, average temperature, and relative humidity during the pollination period, and number of pollen-producing neighbors on individual trees' acorn production. All factors except temperature were significant in at least one of the years of our study, but the combination of factors influencing acorn production varied among years. In 1996, a year of large acorn crop size, acorn production was significantly positively associated with number of neighboring pollen producers and density of female flowers. In 1997, 1998, and 1999, many trees produced few or no acorns, and significant associations between number of pollen-producing neighbors and acorn production were only apparent among moderately to highly reproductive trees. Acorn production by these reproductive trees in 1997 was significantly positively associated with number of neighboring pollen producers and significantly negatively associated with average relative humidity during the pollination period. In 1998, no analysis was possible, because too few trees produced a moderate to large acorn crop. Only density of female flowers was significantly associated with acorn production of moderately to highly reproductive trees in 1999. The effect of spatial scale was also investigated by conducting analyses with pollen producers counted in radii ranging from 30 m to 80 m. The association between number of pollen-producing neighbors and acorn production was strongest when neighborhood sizes of 60 m or larger were considered. Our results suggest that fragmentation and thinning of blue oak woodlands may reduce pollen availability and limit reproduction in this wind-pollinated species.

  20. Detection of the Small Magellanic Cloud in gamma-rays with  Fermi /LAT

    DOE PAGES

    Abdo, A. A.

    2010-11-01

    Context. The flux of gamma rays with energies greater than 100 MeV is dominated by diffuse emission coming from cosmic-rays (CRs) illuminating the interstellar medium (ISM) of our Galaxy through the processes of Bremsstrahlung, pion production and decay, and inverse-Compton scattering. The study of this diffuse emission provides insight into the origin and transport of cosmic rays. Aims. We searched for gamma-ray emission from the Small Magellanic Cloud (SMC) in order to derive constraints on the cosmic-ray population and transport in an external system with properties different from the Milky Way. Methods. We analysed the first 17 months of continuousmore » all-sky observations by the Large Area Telescope (LAT) of the Fermi mission to determine the spatial distribution, flux and spectrum of the gamma-ray emission from the SMC. We also used past radio synchrotron observations of the SMC to study the population of CR electrons specifically. Results. We obtained the first detection of the SMC in high-energy gamma rays, with an integrated >100 MeV flux of (3.7 ± 0.7) × 10 -8 ph cm -2 s -1, with additional systematic uncertainty of ≤16%. The emission is steady and from an extended source ~3° in size. It is not clearly correlated with the distribution of massive stars or neutral gas, nor with known pulsars or supernova remnants, but a certain correlation with supergiant shells is observed. Conclusions. The observed flux implies an upper limit on the average CR nuclei density in the SMC of ~15% of the value measured locally in the Milky Way. The population of high-energy pulsars of the SMC may account for a substantial fraction of the gamma-ray flux, which would make the inferred CR nuclei density even lower. The average density of CR electrons derived from radio synchrotron observations is consistent with the same reduction factor but the uncertainties are large. From our current knowledge of the SMC, such a low CR density does not seem to be due to a lower rate of CR injection and rather indicates a smaller CR confinement volume characteristic size.« less

  1. Detection of the Small Magellanic Cloud in gamma-rays with Fermi/LAT

    NASA Astrophysics Data System (ADS)

    Abdo, A. A.; Ackermann, M.; Ajello, M.; Baldini, L.; Ballet, J.; Barbiellini, G.; Bastieri, D.; Bechtol, K.; Bellazzini, R.; Berenji, B.; Blandford, R. D.; Bloom, E. D.; Bonamente, E.; Borgland, A. W.; Bouvier, A.; Brandt, T. J.; Bregeon, J.; Brez, A.; Brigida, M.; Bruel, P.; Buehler, R.; Buson, S.; Caliandro, G. A.; Cameron, R. A.; Caraveo, P. A.; Carrigan, S.; Casandjian, J. M.; Cecchi, C.; Çelik, Ö.; Charles, E.; Chekhtman, A.; Cheung, C. C.; Chiang, J.; Ciprini, S.; Claus, R.; Cohen-Tanugi, J.; Conrad, J.; Dermer, C. D.; de Palma, F.; Digel, S. W.; Silva, E. Do Couto E.; Drell, P. S.; Dubois, R.; Dumora, D.; Favuzzi, C.; Fegan, S. J.; Fukazawa, Y.; Funk, S.; Fusco, P.; Gargano, F.; Gasparrini, D.; Gehrels, N.; Germani, S.; Giglietto, N.; Giordano, F.; Giroletti, M.; Glanzman, T.; Godfrey, G.; Grenier, I. A.; Grondin, M.-H.; Grove, J. E.; Guiriec, S.; Hadasch, D.; Harding, A. K.; Hayashida, M.; Hays, E.; Horan, D.; Hughes, R. E.; Jean, P.; Jóhannesson, G.; Johnson, A. S.; Johnson, W. N.; Kamae, T.; Katagiri, H.; Kataoka, J.; Kerr, M.; Knödlseder, J.; Kuss, M.; Lande, J.; Latronico, L.; Lee, S.-H.; Lemoine-Goumard, M.; Llena Garde, M.; Longo, F.; Loparco, F.; Lovellette, M. N.; Lubrano, P.; Makeev, A.; Martin, P.; Mazziotta, M. N.; McEnery, J. E.; Michelson, P. F.; Mitthumsiri, W.; Mizuno, T.; Monte, C.; Monzani, M. E.; Morselli, A.; Moskalenko, I. V.; Murgia, S.; Nakamori, T.; Naumann-Godo, M.; Nolan, P. L.; Norris, J. P.; Nuss, E.; Ohsugi, T.; Okumura, A.; Omodei, N.; Orlando, E.; Ormes, J. F.; Panetta, J. H.; Parent, D.; Pelassa, V.; Pepe, M.; Pesce-Rollins, M.; Piron, F.; Porter, T. A.; Rainò, S.; Rando, R.; Razzano, M.; Reimer, A.; Reimer, O.; Reposeur, T.; Ripken, J.; Ritz, S.; Romani, R. W.; Sadrozinski, H. F.-W.; Sander, A.; Saz Parkinson, P. M.; Scargle, J. D.; Sgrò, C.; Siskind, E. J.; Smith, D. A.; Smith, P. D.; Spandre, G.; Spinelli, P.; Strickman, M. S.; Strong, A. W.; Suson, D. J.; Takahashi, H.; Takahashi, T.; Tanaka, T.; Thayer, J. B.; Thayer, J. G.; Thompson, D. J.; Tibaldo, L.; Torres, D. F.; Tosti, G.; Tramacere, A.; Uchiyama, Y.; Usher, T. L.; Vandenbroucke, J.; Vasileiou, V.; Vilchez, N.; Vitale, V.; Waite, A. P.; Wang, P.; Winer, B. L.; Wood, K. S.; Yang, Z.; Ylinen, T.; Ziegler, M.

    2010-11-01

    Context. The flux of gamma rays with energies greater than 100 MeV is dominated by diffuse emission coming from cosmic-rays (CRs) illuminating the interstellar medium (ISM) of our Galaxy through the processes of Bremsstrahlung, pion production and decay, and inverse-Compton scattering. The study of this diffuse emission provides insight into the origin and transport of cosmic rays. Aims: We searched for gamma-ray emission from the Small Magellanic Cloud (SMC) in order to derive constraints on the cosmic-ray population and transport in an external system with properties different from the Milky Way. Methods: We analysed the first 17 months of continuous all-sky observations by the Large Area Telescope (LAT) of the Fermi mission to determine the spatial distribution, flux and spectrum of the gamma-ray emission from the SMC. We also used past radio synchrotron observations of the SMC to study the population of CR electrons specifically. Results: We obtained the first detection of the SMC in high-energy gamma rays, with an integrated >100 MeV flux of (3.7±0.7) × 10-8 ph cm-2 s-1, with additional systematic uncertainty of ≤16%. The emission is steady and from an extended source ~3° in size. It is not clearly correlated with the distribution of massive stars or neutral gas, nor with known pulsars or supernova remnants, but a certain correlation with supergiant shells is observed. Conclusions: The observed flux implies an upper limit on the average CR nuclei density in the SMC of ~15% of the value measured locally in the Milky Way. The population of high-energy pulsars of the SMC may account for a substantial fraction of the gamma-ray flux, which would make the inferred CR nuclei density even lower. The average density of CR electrons derived from radio synchrotron observations is consistent with the same reduction factor but the uncertainties are large. From our current knowledge of the SMC, such a low CR density does not seem to be due to a lower rate of CR injection and rather indicates a smaller CR confinement volume characteristic size.

  2. The Socioecology of Territory Size and a "Work-Around" Hypothesis for the Adoption of Farming

    PubMed Central

    Freeman, Jacob

    2016-01-01

    This paper combines theory from ecology and anthropology to investigate variation in the territory sizes of subsistence oriented agricultural societies. The results indicate that population and the dependence of individuals within a society on “wild” foods partly determine the territory sizes of agricultural societies. In contrast, the productivity of an agroecosystem is not an important determinant of territory size. A comparison of the population-territory size scaling dynamics of agricultural societies and human foragers indicates that foragers and farmers face the same constraints on their ability to expand their territory and intensify their use of resources within a territory. However, the higher density of food in an agroecosystem allows farmers, on average, to live at much higher population densities than human foragers. These macroecological patterns are consistent with a “work-around hypothesis” for the adoption of farming. This hypothesis is that as residential groups of foragers increase in size, farming can sometimes better reduce the tension between an individual’s autonomy over resources and the need for social groups to function to provide public goods like defense and information. PMID:27391955

  3. Species richness and trait composition of butterfly assemblages change along an altitudinal gradient.

    PubMed

    Leingärtner, Annette; Krauss, Jochen; Steffan-Dewenter, Ingolf

    2014-06-01

    Species richness patterns along altitudinal gradients are well-documented ecological phenomena, yet very little data are available on how environmental filtering processes influence the composition and traits of butterfly assemblages at high altitudes. We have studied the diversity patterns of butterfly species at 34 sites along an altitudinal gradient ranging from 600 to 2,000 m a.s.l. in the National Park Berchtesgaden (Germany) and analysed traits of butterfly assemblages associated with dispersal capacity, reproductive strategies and developmental time from lowlands to highlands, including phylogenetic analyses. We found a linear decline in butterfly species richness along the altitudinal gradient, but the phylogenetic relatedness of the butterfly assemblages did not increase with altitude. Compared to butterfly assemblages at lower altitudes, those at higher altitudes were composed of species with larger wings (on average 9%) which laid an average of 68% more eggs. In contrast, egg maturation time in butterfly assemblages decreased by about 22% along the altitudinal gradient. Further, butterfly assemblages at higher altitudes were increasingly dominated by less widespread species. Based on our abundance data, but not on data in the literature, population density increased with altitude, suggesting a reversed density-distribution relationship, with higher population densities of habitat specialists in harsh environments. In conclusion, our data provide evidence for significant shifts in the composition of butterfly assemblages and for the dominance of different traits along the altitudinal gradient. In our study, these changes were mainly driven by environmental factors, whereas phylogenetic filtering played a minor role along the studied altitudinal range.

  4. Why is solar cycle 24 an inefficient producer of high-energy particle events?

    NASA Astrophysics Data System (ADS)

    Vainio, Rami; Raukunen, Osku; Tylka, Allan J.; Dietrich, William F.; Afanasiev, Alexandr

    2017-08-01

    Aims: The aim of the study is to investigate the reason for the low productivity of high-energy SEPs in the present solar cycle. Methods: We employ scaling laws derived from diffusive shock acceleration theory and simulation studies including proton-generated upstream Alfvén waves to find out how the changes observed in the long-term average properties of the erupting and ambient coronal and/or solar wind plasma would affect the ability of shocks to accelerate particles to the highest energies. Results: Provided that self-generated turbulence dominates particle transport around coronal shocks, it is found that the most crucial factors controlling the diffusive shock acceleration process are the number density of seed particles and the plasma density of the ambient medium. Assuming that suprathermal populations provide a fraction of the particles injected to shock acceleration in the corona, we show that the lack of most energetic particle events as well as the lack of low charge-to-mass ratio ion species in the present cycle can be understood as a result of the reduction of average coronal plasma and suprathermal densities in the present cycle over the previous one.

  5. Effect of horseshoe crab spawning density on nest disturbance and exhumation of eggs: A simulation study

    USGS Publications Warehouse

    Smith, D.R.

    2007-01-01

    Because the Delaware Bay horseshoe crab (Limulus polyphemus) population is managed to provide for dependent species, such as migratory shorebirds, there is a need to understand the process of egg exhumation and to predict eggs available to foraging shorebirds. A simple spatial model was used to simulate horseshoe crab spawning that would occur on a typical Delaware Bay beach during spring tide cycles to quantify density-dependent nest disturbance. At least 20% of nests and eggs were disturbed for levels of spawning greater than one third of the average density in Delaware Bay during 2004. Nest disturbance increased approximately linearly as spawning density increased from one half to twice the 2004 level. As spawning density increased further, the percentage of eggs that were disturbed reached an asymptote of 70% for densities up to 10 times the density in 2004. Nest disturbance was heaviest in the mid beach zone. Nest disturbance precedes entrainment and begins the process of exhumation of eggs to surface sediments. Model predictions were combined with observations from egg surveys to estimate a snap-shot exhumation rate of 5-9% of disturbed eggs. Because an unknown quantity of eggs were exhumed and removed from the beach prior to the survey, cumulative exhumation rate was likely to have been higher than the snap-shot estimate. Because egg exhumation is density-dependent, in addition to managing for a high population size, identification and conservation of beaches where spawning horseshoe crabs concentrate in high densities (i.e., hot spots) are important steps toward providing a reliable food supply for migratory shorebirds. ?? 2007 Estuarine Research Federation.

  6. Solar energy in the context of energy use, energy transportation and energy storage.

    PubMed

    MacKay, David J C

    2013-08-13

    Taking the UK as a case study, this paper describes current energy use and a range of sustainable energy options for the future, including solar power and other renewables. I focus on the area involved in collecting, converting and delivering sustainable energy, looking in particular detail at the potential role of solar power. Britain consumes energy at a rate of about 5000 watts per person, and its population density is about 250 people per square kilometre. If we multiply the per capita energy consumption by the population density, then we obtain the average primary energy consumption per unit area, which for the UK is 1.25 watts per square metre. This areal power density is uncomfortably similar to the average power density that could be supplied by many renewables: the gravitational potential energy of rainfall in the Scottish highlands has a raw power per unit area of roughly 0.24 watts per square metre; energy crops in Europe deliver about 0.5 watts per square metre; wind farms deliver roughly 2.5 watts per square metre; solar photovoltaic farms in Bavaria, Germany, and Vermont, USA, deliver 4 watts per square metre; in sunnier locations, solar photovoltaic farms can deliver 10 watts per square metre; concentrating solar power stations in deserts might deliver 20 watts per square metre. In a decarbonized world that is renewable-powered, the land area required to maintain today's British energy consumption would have to be similar to the area of Britain. Several other high-density, high-consuming countries are in the same boat as Britain, and many other countries are rushing to join us. Decarbonizing such countries will only be possible through some combination of the following options: the embracing of country-sized renewable power-generation facilities; large-scale energy imports from country-sized renewable facilities in other countries; population reduction; radical efficiency improvements and lifestyle changes; and the growth of non-renewable low-carbon sources, namely 'clean' coal, 'clean' gas and nuclear power. If solar is to play a large role in the future energy system, then we need new methods for energy storage; very-large-scale solar either would need to be combined with electricity stores or it would need to serve a large flexible demand for energy that effectively stores useful energy in the form of chemicals, heat, or cold.

  7. Pricing of common cosmetic surgery procedures: local economic factors trump supply and demand.

    PubMed

    Richardson, Clare; Mattison, Gennaya; Workman, Adrienne; Gupta, Subhas

    2015-02-01

    The pricing of cosmetic surgery procedures has long been thought to coincide with laws of basic economics, including the model of supply and demand. However, the highly variable prices of these procedures indicate that additional economic contributors are probable. The authors sought to reassess the fit of cosmetic surgery costs to the model of supply and demand and to determine the driving forces behind the pricing of cosmetic surgery procedures. Ten plastic surgery practices were randomly selected from each of 15 US cities of various population sizes. Average prices of breast augmentation, mastopexy, abdominoplasty, blepharoplasty, and rhytidectomy in each city were compared with economic and demographic statistics. The average price of cosmetic surgery procedures correlated substantially with population size (r = 0.767), cost-of-living index (r = 0.784), cost to own real estate (r = 0.714), and cost to rent real estate (r = 0.695) across the 15 US cities. Cosmetic surgery pricing also was found to correlate (albeit weakly) with household income (r = 0.436) and per capita income (r = 0.576). Virtually no correlations existed between pricing and the density of plastic surgeons (r = 0.185) or the average age of residents (r = 0.076). Results of this study demonstrate a correlation between costs of cosmetic surgery procedures and local economic factors. Cosmetic surgery pricing cannot be completely explained by the supply-and-demand model because no association was found between procedure cost and the density of plastic surgeons. © 2015 The American Society for Aesthetic Plastic Surgery, Inc. Reprints and permission: journals.permissions@oup.com.

  8. Modeling Parasite Dynamics on Farmed Salmon for Precautionary Conservation Management of Wild Salmon

    PubMed Central

    Rogers, Luke A.; Peacock, Stephanie J.; McKenzie, Peter; DeDominicis, Sharon; Jones, Simon R. M.; Chandler, Peter; Foreman, Michael G. G.; Revie, Crawford W.; Krkošek, Martin

    2013-01-01

    Conservation management of wild fish may include fish health management in sympatric populations of domesticated fish in aquaculture. We developed a mathematical model for the population dynamics of parasitic sea lice (Lepeophtheirus salmonis) on domesticated populations of Atlantic salmon (Salmo salar) in the Broughton Archipelago region of British Columbia. The model was fit to a seven-year dataset of monthly sea louse counts on farms in the area to estimate population growth rates in relation to abiotic factors (temperature and salinity), local host density (measured as cohort surface area), and the use of a parasiticide, emamectin benzoate, on farms. We then used the model to evaluate management scenarios in relation to policy guidelines that seek to keep motile louse abundance below an average three per farmed salmon during the March–June juvenile wild Pacific salmon (Oncorhynchus spp.) migration. Abiotic factors mediated the duration of effectiveness of parasiticide treatments, and results suggest treatment of farmed salmon conducted in January or early February minimized average louse abundance per farmed salmon during the juvenile wild salmon migration. Adapting the management of parasites on farmed salmon according to migrations of wild salmon may therefore provide a precautionary approach to conserving wild salmon populations in salmon farming regions. PMID:23577082

  9. Impact of minimum catch size on the population viability of Strombus gigas (Mesogastropoda: Strombidae) in Quintana Roo, Mexico.

    PubMed

    Peel, Joanne R; Mandujano, María del Carmen

    2014-12-01

    The queen conch Strombus gigas represents one of the most important fishery resources of the Caribbean but heavy fishing pressure has led to the depletion of stocks throughout the region, causing the inclusion of this species into CITES Appendix II and IUCN's Red-List. In Mexico, the queen conch is managed through a minimum fishing size of 200 mm shell length and a fishing quota which usually represents 50% of the adult biomass. The objectives of this study were to determine the intrinsic population growth rate of the queen conch population of Xel-Ha, Quintana Roo, Mexico, and to assess the effects of a regulated fishing impact, simulating the extraction of 50% adult biomass on the population density. We used three different minimum size criteria to demonstrate the effects of minimum catch size on the population density and discuss biological implications. Demographic data was obtained through capture-mark-recapture sampling, collecting all animals encountered during three hours, by three divers, at four different sampling sites of the Xel-Ha inlet. The conch population was sampled each month between 2005 and 2006, and bimonthly between 2006 and 2011, tagging a total of 8,292 animals. Shell length and lip thickness were determined for each individual. The average shell length for conch with formed lip in Xel-Ha was 209.39 ± 14.18 mm and the median 210 mm. Half of the sampled conch with lip ranged between 200 mm and 219 mm shell length. Assuming that the presence of the lip is an indicator for sexual maturity, it can be concluded that many animals may form their lip at greater shell lengths than 200 mm and ought to be considered immature. Estimation of relative adult abundance and densities varied greatly depending on the criteria employed for adult classification. When using a minimum fishing size of 200 mm shell length, between 26.2% and up to 54.8% of the population qualified as adults, which represented a simulated fishing impact of almost one third of the population. When conch extraction was simulated using a classification criteria based on lip thickness, it had a much smaller impact on the population density. We concluded that the best management strategy for S. gigas is a minimum fishing size based on a lip thickness, since it has lower impact on the population density, and given that selective fishing pressure based on size may lead to the appearance of small adult individuals with reduced fecundity. Furthermore, based on the reproductive biology and the results of the simulated fishing, we suggest a minimum lip thickness of ≥ 15 mm, which ensures the protection of reproductive stages, reduces the risk of overfishing, leading to non-viable density reduction.

  10. Correlation Between Pediatrician Supply and Public Health in Japan as Evidenced by Vaccination Coverage in 2010: Secondary Data Analysis

    PubMed Central

    Sakai, Rie; Fink, Günther; Wang, Wei; Kawachi, Ichiro

    2015-01-01

    Background In industrialized countries, assessment of the causal effect of physician supply on population health has yielded mixed results. Since the scope of child vaccination is an indicator of preventive health service utilization, this study investigates the correlation between vaccination coverage and pediatrician supply as a reflection of overall pediatric health during a time of increasing pediatrician numbers in Japan. Methods Cross-sectional data were collected from publicly available sources for 2010. Dependent variables were vaccination coverage for measles and diphtheria, pertussis, and tetanus (DPT) by region. The primary predictor of interest was number of pediatricians per 10 000-child population (pediatrician density) at the municipality level. Multivariate logistic regression models were used to estimate associations of interest, conditional on a large range of demographic and infrastructure-related factors as covariates, including non-pediatric physician density, total population, per capita income, occupation, unemployment rate, prevalence of single motherhood, number of hospital beds per capita, length of roads, crime rate, accident rate, and metropolitan area code as urban/rural status. The percentage of the population who completed college-level education or higher in 2010 was included in the model as a proxy for education level. Results Pediatrician density was positively and significantly associated with vaccination coverage for both vaccine series. On average, each unit of pediatrician density increased odds by 1.012 for measles (95% confidence interval, 1.010–1.015) and 1.019 for DPT (95% confidence interval, 1.016–1.022). Conclusions Policies increasing pediatrician supply contribute to improved preventive healthcare services utilization, such as immunizations, and presumably improved child health status in Japan. PMID:25817986

  11. The hELENa project - I. Stellar populations of early-type galaxies linked with local environment and galaxy mass

    NASA Astrophysics Data System (ADS)

    Sybilska, A.; Lisker, T.; Kuntschner, H.; Vazdekis, A.; van de Ven, G.; Peletier, R.; Falcón-Barroso, J.; Vijayaraghavan, R.; Janz, J.

    2017-09-01

    We present the first in a series of papers in The role of Environment in shaping Low-mass Early-type Nearby galaxies (hELENa) project. In this paper, we combine our sample of 20 low-mass early types (dEs) with 258 massive early types (ETGs) from the ATLAS3D survey - all observed with the SAURON integral field unit - to investigate early-type galaxies' stellar population scaling relations and the dependence of the population properties on local environment, extended to the low-σ regime of dEs. The ages in our sample show more scatter at lower σ values, indicative of less massive galaxies being affected by the environment to a higher degree. The shape of the age-σ relations for cluster versus non-cluster galaxies suggests that cluster environment speeds up the placing of galaxies on the red sequence. While the scaling relations are tighter for cluster than for the field/group objects, we find no evidence for a difference in average population characteristics of the two samples. We investigate the properties of our sample in the Virgo cluster as a function of number density (rather than simple clustrocentric distance) and find that dE ages correlate with the local density such that galaxies in regions of lower density are younger, likely because they are later arrivals to the cluster or have experienced less pre-processing in groups, and consequently used up their gas reservoir more recently. Overall, dE properties correlate more strongly with density than those of massive ETGs, which was expected as less massive galaxies are more susceptible to external influences.

  12. Hydrology and density feedbacks control the ecology of intermediate hosts of schistosomiasis across habitats in seasonal climates

    PubMed Central

    Perez-Saez, Javier; Mande, Theophile; Ceperley, Natalie; Bertuzzo, Enrico; Mari, Lorenzo; Rinaldo, Andrea

    2016-01-01

    We report about field and theoretical studies on the ecology of the aquatic snails (Bulinus spp. and Biomphalaria pfeifferi) that serve as obligate intermediate hosts in the complex life cycle of the parasites causing human schistosomiasis. Snail abundance fosters disease transmission, and thus the dynamics of snail populations are critically important for schistosomiasis modeling and control. Here, we single out hydrological drivers and density dependence (or lack of it) of ecological growth rates of local snail populations by contrasting novel ecological and environmental data with various models of host demography. Specifically, we study various natural and man-made habitats across Burkina Faso’s highly seasonal climatic zones. Demographic models are ranked through formal model comparison and structural risk minimization. The latter allows us to evaluate the suitability of population models while clarifying the relevant covariates that explain empirical observations of snail abundance under the actual climatic forcings experienced by the various field sites. Our results link quantitatively hydrological drivers to distinct population dynamics through specific density feedbacks, and show that statistical methods based on model averaging provide reliable snail abundance projections. The consistency of our ranking results suggests the use of ad hoc models of snail demography depending on habitat type (e.g., natural vs. man-made) and hydrological characteristics (e.g., ephemeral vs. permanent). Implications for risk mapping and space-time allocation of control measures in schistosomiasis-endemic contexts are discussed. PMID:27162339

  13. Hydrology and density feedbacks control the ecology of intermediate hosts of schistosomiasis across habitats in seasonal climates.

    PubMed

    Perez-Saez, Javier; Mande, Theophile; Ceperley, Natalie; Bertuzzo, Enrico; Mari, Lorenzo; Gatto, Marino; Rinaldo, Andrea

    2016-06-07

    We report about field and theoretical studies on the ecology of the aquatic snails (Bulinus spp. and Biomphalaria pfeifferi) that serve as obligate intermediate hosts in the complex life cycle of the parasites causing human schistosomiasis. Snail abundance fosters disease transmission, and thus the dynamics of snail populations are critically important for schistosomiasis modeling and control. Here, we single out hydrological drivers and density dependence (or lack of it) of ecological growth rates of local snail populations by contrasting novel ecological and environmental data with various models of host demography. Specifically, we study various natural and man-made habitats across Burkina Faso's highly seasonal climatic zones. Demographic models are ranked through formal model comparison and structural risk minimization. The latter allows us to evaluate the suitability of population models while clarifying the relevant covariates that explain empirical observations of snail abundance under the actual climatic forcings experienced by the various field sites. Our results link quantitatively hydrological drivers to distinct population dynamics through specific density feedbacks, and show that statistical methods based on model averaging provide reliable snail abundance projections. The consistency of our ranking results suggests the use of ad hoc models of snail demography depending on habitat type (e.g., natural vs. man-made) and hydrological characteristics (e.g., ephemeral vs. permanent). Implications for risk mapping and space-time allocation of control measures in schistosomiasis-endemic contexts are discussed.

  14. Stochastically generating tree diameter lists to populate forest stands based on the linkage variables forest type and stand age

    Treesearch

    Bernard R. Parresol; F. Thomas Lloyd

    2003-01-01

    Forest inventory data were used to develop a standage-driven, stochastic predictor of unit-area, frequency weighted lists of breast high tree diameters (DBH). The average of mean statistics from 40 simulation prediction sets of an independent 78-plot validation dataset differed from the observed validation means by 0.5 cm for DBH, and by 12 trees/h for density. The 40...

  15. A long-term evaluation of biopsy darts and DNA to estimate cougar density

    USGS Publications Warehouse

    Beausoleil, Richard A.; Clark, Joseph D.; Maletzke, Benjamin T.

    2016-01-01

    Accurately estimating cougar (Puma concolor) density is usually based on long-term research consisting of intensive capture and Global Positioning System collaring efforts and may cost hundreds of thousands of dollars annually. Because wildlife agency budgets rarely accommodate this approach, most infer cougar density from published literature, rely on short-term studies, or use hunter harvest data as a surrogate in their jurisdictions; all of which may limit accuracy and increase risk of management actions. In an effort to develop a more cost-effective long-term strategy, we evaluated a research approach using citizen scientists with trained hounds to tree cougars and collect tissue samples with biopsy darts. We then used the DNA to individually identify cougars and employed spatially explicit capture–recapture models to estimate cougar densities. Overall, 240 tissue samples were collected in northeastern Washington, USA, producing 166 genotypes (including recaptures and excluding dependent kittens) of 133 different cougars (8-25/yr) from 2003 to 2011. Mark–recapture analyses revealed a mean density of 2.2 cougars/100 km2 (95% CI=1.1-4.3) and stable to decreasing population trends (β=-0.048, 95% CI=-0.106–0.011) over the 9 years of study, with an average annual harvest rate of 14% (range=7-21%). The average annual cost per year for field sampling and genotyping was US$11,265 ($422.24/sample or $610.73/successfully genotyped sample). Our results demonstrated that long-term biopsy sampling using citizen scientists can increase capture success and provide reliable cougar-density information at a reasonable cost.

  16. Dynamics of an introduced and unexploited Lake Whitefish population in Lake Pend Oreille, Idaho

    USGS Publications Warehouse

    Hosack, Michael A.; Hansen, Michael J.; Horner, Ned J.

    2014-01-01

    To evaluate biological potential of a commercial fishery for an unexploited Lake Whitefish Coregonus clupeaformis population in Lake Pend Oreille, Idaho, we estimated population parameters related to production and yield. The length frequency based on trap-netting in autumn 2005 was normal with a mean of 448 mm TL, whereas the length frequency based on gillnetting in spring 2006 was bimodal with a mean of 390 mm TL. Sex composition was skewed toward females (0.66) during autumn trap-netting. Shape parameters β of weight–length models for females (β = 3.38) and males (β = 3.45) were similar to those of other unexploited populations. Instantaneous growth rates K for females (K = 0.144 per year) and males (K = 0.153 per year) were among the lowest for unexploited populations across the species’ range. Age at 50% maturity (females: 6.5 years; males: 6.0 years) and length at 50% maturity (females: 390 mm TL; males: 378 mm TL) were high for unexploited populations. The natural mortality rate M (0.149 per year, ages 11–36) was among the lowest observed for unexploited populations. Adult population density was lower than that of other populations based on total surface area (mean = 1.35 fish/ha; 95% confidence interval [CI] = 1.11–1.78 fish/ha) but was average based on lake area shallower than 70 m (4.07 fish/ha; 95% CI = 3.35–5.35 fish/ha). Population density of juveniles and adults averaged 84 fish/ha (95% CI = 52–143 fish/ha) over the entire surface area and 278 fish/ha (95% CI = 173–474 fish/ha) over depths shallower than 70 m. The difference between the low M of the unexploited population in Lake Pend Oreille (M = 0.149 per year; annual mortality rate A = 14%) and the high sustainable total mortality Z of exploited stocks in the Laurentian Great Lakes (Z = 1.204; A = 70%) suggests a large scope for sustainable fishing mortality F (1.055 per year; exploitation rate u = 61%) that is equivalent to a sustainable Lake Whitefish harvest of 55,000 individuals (50,000–60,000 individuals) and 49,000 kg (45,000–54,000 kg) from Lake Pend Oreille.

  17. Spectral density of mixtures of random density matrices for qubits

    NASA Astrophysics Data System (ADS)

    Zhang, Lin; Wang, Jiamei; Chen, Zhihua

    2018-06-01

    We derive the spectral density of the equiprobable mixture of two random density matrices of a two-level quantum system. We also work out the spectral density of mixture under the so-called quantum addition rule. We use the spectral densities to calculate the average entropy of mixtures of random density matrices, and show that the average entropy of the arithmetic-mean-state of n qubit density matrices randomly chosen from the Hilbert-Schmidt ensemble is never decreasing with the number n. We also get the exact value of the average squared fidelity. Some conjectures and open problems related to von Neumann entropy are also proposed.

  18. Focused study of interweaving hazards across the Caribbean

    NASA Astrophysics Data System (ADS)

    Braun, John J.; Mattioli, Glen S.; Calais, Eric; Carlson, David; Dixon, Timothy H.; Jackson, Michael E.; Kursinski, E. Robert; Mora-Paez, Hector; Miller, M. Meghan; Pandya, Rajul; Robertson, Richard; Wang, Guoquan

    2012-02-01

    The Caribbean is a region of lush vegetation, beaches, active volcanoes, and significant mountain ranges, all of which create a natural aesthetic that is recognized globally. Yet these very same features, molded through geological, oceanic, and atmospheric processes, also pose natural hazards for the developing countries in the Caribbean. The rise in population density, migration to coastal areas, and substandard building practices, combined with the threat of natural hazards, put the region's human population at risk for particularly devastating disasters. These demographic and social characteristics exist against a backdrop of the threat of an evolving climate, which produces a more vigorous hurricane environment and a rising average sea level.

  19. Estimation of linkage disequilibrium and analysis of genetic diversity in Korean chicken lines.

    PubMed

    Seo, Dongwon; Lee, Doo Ho; Choi, Nuri; Sudrajad, Pita; Lee, Seung-Hwan; Lee, Jun-Heon

    2018-01-01

    The development of genetic markers for animal breeding is an effective strategy to reduce the time and cost required to improve economically important traits. To implement genomic selection in the multibreed chicken population of Korea, an understanding of the linkage disequilibrium (LD) status of the target population is essential. In this study, we performed population genetic analyses to investigate LD decay, the effective population size, and breed diversity using 600K high-density single nucleotide polymorphism genotypes of 189 native chickens in 14 lines (including Korean native chicken, imported and adapted purebred and commercial chickens). The results indicated that commercial native chickens have less calculated LD (average, r2 = 0.13-0.26) and purebred native chickens have more calculated LD (average, r2 = 0.24-0.37) across the entire genome. The effective population sizes of the examined lines showed patterns opposite to those of population LD. The phylogeny and admixture analyses showed that commercial and purebred chickens were well distinguished, except for Rhode Island Red (RIR) purebred lines of NC (NIAS_RIR_C) and ND (NIAS_RIR_D). These lines are difficult to distinguish clearly because they originated from the same respective breeds. The results of this study may provide important information for the development of genetic markers that can be used in breeding to improve the economic traits of native chickens.

  20. Estimation of linkage disequilibrium and analysis of genetic diversity in Korean chicken lines

    PubMed Central

    Seo, Dongwon; Lee, Doo Ho; Choi, Nuri; Sudrajad, Pita; Lee, Seung-Hwan

    2018-01-01

    The development of genetic markers for animal breeding is an effective strategy to reduce the time and cost required to improve economically important traits. To implement genomic selection in the multibreed chicken population of Korea, an understanding of the linkage disequilibrium (LD) status of the target population is essential. In this study, we performed population genetic analyses to investigate LD decay, the effective population size, and breed diversity using 600K high-density single nucleotide polymorphism genotypes of 189 native chickens in 14 lines (including Korean native chicken, imported and adapted purebred and commercial chickens). The results indicated that commercial native chickens have less calculated LD (average, r2 = 0.13–0.26) and purebred native chickens have more calculated LD (average, r2 = 0.24–0.37) across the entire genome. The effective population sizes of the examined lines showed patterns opposite to those of population LD. The phylogeny and admixture analyses showed that commercial and purebred chickens were well distinguished, except for Rhode Island Red (RIR) purebred lines of NC (NIAS_RIR_C) and ND (NIAS_RIR_D). These lines are difficult to distinguish clearly because they originated from the same respective breeds. The results of this study may provide important information for the development of genetic markers that can be used in breeding to improve the economic traits of native chickens. PMID:29425208

  1. Obscuration-dependent Evolution of Active Galactic Nuclei

    NASA Astrophysics Data System (ADS)

    Buchner, Johannes; Georgakakis, Antonis; Nandra, Kirpal; Brightman, Murray; Menzel, Marie-Luise; Liu, Zhu; Hsu, Li-Ting; Salvato, Mara; Rangel, Cyprian; Aird, James; Merloni, Andrea; Ross, Nicholas

    2015-04-01

    We aim to constrain the evolution of active galactic nuclei (AGNs) as a function of obscuration using an X-ray-selected sample of ~2000 AGNs from a multi-tiered survey including the CDFS, AEGIS-XD, COSMOS, and XMM-XXL fields. The spectra of individual X-ray sources are analyzed using a Bayesian methodology with a physically realistic model to infer the posterior distribution of the hydrogen column density and intrinsic X-ray luminosity. We develop a novel non-parametric method that allows us to robustly infer the distribution of the AGN population in X-ray luminosity, redshift, and obscuring column density, relying only on minimal smoothness assumptions. Our analysis properly incorporates uncertainties from low count spectra, photometric redshift measurements, association incompleteness, and the limited sample size. We find that obscured AGNs with N H > 1022 cm-2 account for {77}+4-5% of the number density and luminosity density of the accretion supermassive black hole population with L X > 1043 erg s-1, averaged over cosmic time. Compton-thick AGNs account for approximately half the number and luminosity density of the obscured population, and {38}+8-7% of the total. We also find evidence that the evolution is obscuration dependent, with the strongest evolution around N H ≈ 1023 cm-2. We highlight this by measuring the obscured fraction in Compton-thin AGNs, which increases toward z ~ 3, where it is 25% higher than the local value. In contrast, the fraction of Compton-thick AGNs is consistent with being constant at ≈35%, independent of redshift and accretion luminosity. We discuss our findings in the context of existing models and conclude that the observed evolution is, to first order, a side effect of anti-hierarchical growth.

  2. Production of a high-efficiency TILLING population through polyploidization.

    PubMed

    Tsai, Helen; Missirian, Victor; Ngo, Kathie J; Tran, Robert K; Chan, Simon R; Sundaresan, Venkatesan; Comai, Luca

    2013-04-01

    Targeting Induced Local Lesions in Genomes (TILLING) provides a nontransgenic method for reverse genetics that is widely applicable, even in species where other functional resources are missing or expensive to build. The efficiency of TILLING, however, is greatly facilitated by high mutation density. Species vary in the number of mutations induced by comparable mutagenic treatments, suggesting that genetic background may affect the response. Allopolyploid species have often yielded higher mutation density than diploids. To examine the effect of ploidy, we autotetraploidized the Arabidopsis (Arabidopsis thaliana) ecotype Columbia, whose diploid has been used for TILLING extensively, and mutagenized it with 50 mm ethylmethane sulfonate. While the same treatment sterilized diploid Columbia, the tetraploid M1 plants produced good seed. To determine the mutation density, we searched 528 individuals for induced mutations in 15 genes for which few or no knockout alleles were previously available. We constructed tridimensional pools from the genomic DNA of M2 plants, amplified target DNA, and subjected them to Illumina sequencing. The results were analyzed with an improved version of the mutation detection software CAMBa that accepts any pooling scheme. This small population provided a rich resource with approximately 25 mutations per queried 1.5-kb fragment, including on average four severe missense and 1.3 truncation mutations. The overall mutation density of 19.4 mutations Mb(-1) is 4 times that achieved in the corresponding diploid accession, indicating that genomic redundancy engenders tolerance to high mutation density. Polyploidization of diploids will allow the production of small populations, such as less than 2,000, that provide allelic series from knockout to mild loss of function for virtually all genes.

  3. Health Insurance Marketplaces: Early Findings on Changes in Plan Availability and Premiums in Rural Places, 2014-2015.

    PubMed

    Barker, Abigail; McBride, Timothy D; Kemper, Leah M; Mueller, Keith

    2015-05-01

    The Patient Protection and Affordable Care Act established Health Insurance Marketplaces (HIMs) in all 50 states and the District of Columbia. This policy brief assesses the changes in HIMs from 2014 to 2015 in terms of choices offered and premiums charged, with emphasis on how these measures vary across rural and urban places. Key Findings. (1) In 74 percent of HIM rating areas, the number of firms operating increased by at least one, while the number of firms decreased in only about 6 percent of rating areas. Further, 64 percent of rating areas with fewer than 50 persons per square mile gained at least one firm. (2) There was no consistent pattern of premium increases with respect to rating area population density (used as a proxy here for the degree of "ruralness" of the rating areas). Nationally, rural areas are not experiencing higher premium increases than their urban counterparts. In fact, the lowest increases in second-lowest cost silver plan premiums occurred in the medium-density population rating areas of 51 to 300 persons per square mile. (3) Average adjusted premiums increased from 2014 to 2015 by 6.7 percent in Federally-Facilitated Marketplaces (FFMs) compared to just 1.4 percent in State-Based Marketplaces (SBMs). Regardless of SBM or FFM status, premium increases across the United States were negatively correlated with the number of firms entering the market. (4) Analysis of the most rural states, in terms of percentage of the population classified as nonmetropolitan, shows that, in general, premiums fell significantly in rural places where they had been rather high, and they increased in rural places where they had been rather low. The five rural states with the lowest premium increases had an average of 0.17 firms entering the market, while the five with the highest premium increases had an average of 0.50 firms exiting the market.

  4. Are Emissions of Restricted Halocarbons in the USA and Canada Still Globally Significant?

    NASA Astrophysics Data System (ADS)

    Hurst, D. F.; Romashkin, P. A.; Hall, B. D.; Elkins, J. W.; Lin, J. C.; Gerbig, C.; Daube, B. C.; Wofsy, S. C.

    2004-12-01

    The global manufacture of halocarbons regulated by the Montreal Protocol has dropped substantially in response to the January 1, 1996, production phase-out deadline (1994 for halons) for developed (Article 5) countries like the United States and Canada. Contemporary emissions of these ozone-depleting substances (ODS) emanate from ongoing production in developing countries and releases of banked halocarbons world-wide. ODS emissions in developing nations can be appraised from reported production figures, but not so for developed nations where recent manufacture is negligible. Emissions in the United States and Canada are increasingly difficult to estimate because of limited information about bank sizes and release rates in the post-production era. In addition, regional- or national-scale emission estimates should no longer be derived wholly from localized measurements because of the potentially patchy spatial distributions of modern emissions. We estimate ODS emissions in the USA and Canada from >1000 simultaneous, in situ measurements each of CO and six restricted halocarbons (CFC-11, CFC-12, CFC-113, methyl chloroform, carbon tetrachloride, and halon-1211) in and above the planetary boundary layer during the 2003 CO2 Budget and Regional Airborne - North America (COBRA-NA 2003) study. The data obtained during 87 flight hours are geographically extensive (>30,000 km) including two 11,000 km flight circuits across both countries. More than 50 pollution "events" with statistically significant ODS:CO emission ratios were sampled, and for each event we have determined a flux footprint using the Stochastic Time-Inverted Lagrangian Transport (STILT) model. The model also calculates footprint-weighted average population densities and CO fluxes which we convert to footprint-weighted average ODS fluxes using the measured ODS:CO emission ratios. Statistically robust relationships between footprint-averaged ODS fluxes and population densities for several ODS indicate that population-based extrapolations of these relationships to national levels are warranted. Emission estimates for the USA and Canada in 2003 will be presented and compared to the magnitudes of global emissions.

  5. Global hydrobelts and hydroregions: improved reporting scale for water-related issues?

    NASA Astrophysics Data System (ADS)

    Meybeck, M.; Kummu, M.; Dürr, H. H.

    2013-03-01

    Global-scale water issues such as its availability, water needs or stress, or management, are mapped at various resolutions and reported at many scales, mostly along political or continental boundaries. As such, they ignore the fundamental heterogeneity of hydroclimates and natural boundaries of river basins. Here we describe the continental landmasses at two levels: eight hydrobelts strictly limited by river basins, defined at a 30' (0.5°) resolution, which are decomposed on continents as 26 hydroregions. The belts were defined and delineated, based primarily on the annual average temperature (T) and run-off (q), to maximise inter-belt differences and minimise intra-belt variability. This new global puzzle defines homogeneous and near-contiguous entities with similar hydrological and thermal regimes, glacial and postglacial basin histories, endorheism distribution and sensitivity to climate variations. The mid-latitude, dry and subtropical belts have northern and southern analogues and a general symmetry can be observed for T and q between them. The boreal and equatorial belts are unique. Population density between belts and between the continents varies greatly, resulting in pronounced differences between the belts with analogues in both hemispheres. Hydroregions (median size 4.7 M km2) are highly contrasted, with the average q ranging between 6 and 1393 mm yr-1 and the average T between -9.7 and +26.3 °C, and a population density ranging from 0.7 to 0.8 p km-2 for the North American boreal region and some Australian hydroregions to 280 p km-2 for some Asian hydroregions. The population/run-off ratio, normalised to a reference pristine region, is used to map and quantify the global population at risk of severe water quality degradation. Our initial tests suggest that hydrobelt and hydroregion divisions are often more appropriate than conventional continental or political divisions for the global analysis of river basins within the Earth system and of water resources. The GIS files of the hydrobelts and hydroregions are available at the supplement of this article and at doi:10.1594/PANGAEA.806957 as well as geotypes.net.

  6. Walleye population and fishery responses after elimination of legal harvest on Escanaba Lake, Wisconsin

    USGS Publications Warehouse

    Haglund, Justin M.; Isermann, Daniel A.; Sass, Greg G.

    2016-01-01

    Implementing harvest regulations to eliminate or substantially reduce (≥90%) the exploitation of Walleyes Sander vitreus in recreational fisheries may increase population size structure, but these measures also could reduce angler effort because many Walleye anglers are harvest oriented. We analyzed data collected during 1995–2015 to determine whether Walleye population and fishery metrics in Escanaba Lake, Wisconsin, changed after a minimum TL limit of 71 cm with a one-fish daily bag limit was implemented in 2003. This change eliminated the legal harvest of Walleyes after several decades during which annual exploitation averaged 34%. We detected a significant increase in the loge density of adult females after the regulation change, but the loge density of all adults and adult males did not differ between periods. Mean TL of adult males was significantly greater after the regulation change, but the mean TL of females and the proportional size distribution of preferred-length fish (≥51 cm TL) were similar between periods. Sex-specific mean TLs at age 5 did not differ between periods. Loge density of age-0 Walleyes did not change after 2003, but variation in age-0 density was lower. Total angler effort and the effort for anglers targeting Walleyes were significantly lower (35% and 60% declines, respectively) after the regulation change, whereas catch rates for both angler categories did not differ between periods. Our results suggest that implementing highly restrictive regulations that greatly reduce or eliminate legal harvest will not always increase angler catch rates and population size structure. Highly restrictive regulations may also deter anglers from using a fishery when many other fisheries are available. Our findings are useful for fishery managers who may work with anglers holding the belief that lower exploitation is a potential remedy for low Walleye size structure, even when density and growth suggest that there is limited potential for improvement.

  7. High-density genetic map using whole-genome resequencing for fine mapping and candidate gene discovery for disease resistance in peanut.

    PubMed

    Agarwal, Gaurav; Clevenger, Josh; Pandey, Manish K; Wang, Hui; Shasidhar, Yaduru; Chu, Ye; Fountain, Jake C; Choudhary, Divya; Culbreath, Albert K; Liu, Xin; Huang, Guodong; Wang, Xingjun; Deshmukh, Rupesh; Holbrook, C Corley; Bertioli, David J; Ozias-Akins, Peggy; Jackson, Scott A; Varshney, Rajeev K; Guo, Baozhu

    2018-04-10

    Whole-genome resequencing (WGRS) of mapping populations has facilitated development of high-density genetic maps essential for fine mapping and candidate gene discovery for traits of interest in crop species. Leaf spots, including early leaf spot (ELS) and late leaf spot (LLS), and Tomato spotted wilt virus (TSWV) are devastating diseases in peanut causing significant yield loss. We generated WGRS data on a recombinant inbred line population, developed a SNP-based high-density genetic map, and conducted fine mapping, candidate gene discovery and marker validation for ELS, LLS and TSWV. The first sequence-based high-density map was constructed with 8869 SNPs assigned to 20 linkage groups, representing 20 chromosomes, for the 'T' population (Tifrunner × GT-C20) with a map length of 3120 cM and an average distance of 1.45 cM. The quantitative trait locus (QTL) analysis using high-density genetic map and multiple season phenotyping data identified 35 main-effect QTLs with phenotypic variation explained (PVE) from 6.32% to 47.63%. Among major-effect QTLs mapped, there were two QTLs for ELS on B05 with 47.42% PVE and B03 with 47.38% PVE, two QTLs for LLS on A05 with 47.63% and B03 with 34.03% PVE and one QTL for TSWV on B09 with 40.71% PVE. The epistasis and environment interaction analyses identified significant environmental effects on these traits. The identified QTL regions had disease resistance genes including R-genes and transcription factors. KASP markers were developed for major QTLs and validated in the population and are ready for further deployment in genomics-assisted breeding in peanut. © 2018 The Authors. Plant Biotechnology Journal published by Society for Experimental Biology and The Association of Applied Biologists and John Wiley & Sons Ltd.

  8. Estimating risks to aquatic life using quantile regression

    USGS Publications Warehouse

    Schmidt, Travis S.; Clements, William H.; Cade, Brian S.

    2012-01-01

    One of the primary goals of biological assessment is to assess whether contaminants or other stressors limit the ecological potential of running waters. It is important to interpret responses to contaminants relative to other environmental factors, but necessity or convenience limit quantification of all factors that influence ecological potential. In these situations, the concept of limiting factors is useful for data interpretation. We used quantile regression to measure risks to aquatic life exposed to metals by including all regression quantiles (τ  =  0.05–0.95, by increments of 0.05), not just the upper limit of density (e.g., 90th quantile). We measured population densities (individuals/0.1 m2) of 2 mayflies (Rhithrogena spp., Drunella spp.) and a caddisfly (Arctopsyche grandis), aqueous metal mixtures (Cd, Cu, Zn), and other limiting factors (basin area, site elevation, discharge, temperature) at 125 streams in Colorado. We used a model selection procedure to test which factor was most limiting to density. Arctopsyche grandis was limited by other factors, whereas metals limited most quantiles of density for the 2 mayflies. Metals reduced mayfly densities most at sites where other factors were not limiting. Where other factors were limiting, low mayfly densities were observed despite metal concentrations. Metals affected mayfly densities most at quantiles above the mean and not just at the upper limit of density. Risk models developed from quantile regression showed that mayfly densities observed at background metal concentrations are improbable when metal mixtures are at US Environmental Protection Agency criterion continuous concentrations. We conclude that metals limit potential density, not realized average density. The most obvious effects on mayfly populations were at upper quantiles and not mean density. Therefore, we suggest that policy developed from mean-based measures of effects may not be as useful as policy based on the concept of limiting factors.

  9. Estimating numbers of greater prairie-chickens using mark-resight techniques

    USGS Publications Warehouse

    Clifton, A.M.; Krementz, D.G.

    2006-01-01

    Current monitoring efforts for greater prairie-chicken (Tympanuchus cupido pinnatus) populations indicate that populations are declining across their range. Monitoring the population status of greater prairie-chickens is based on traditional lek surveys (TLS) that provide an index without considering detectability. Estimators, such as immigration-emigration joint maximum-likelihood estimator from a hypergeometric distribution (IEJHE), can account for detectability and provide reliable population estimates based on resightings. We evaluated the use of mark-resight methods using radiotelemetry to estimate population size and density of greater prairie-chickens on 2 sites at a tallgrass prairie in the Flint Hills of Kansas, USA. We used average distances traveled from lek of capture to estimate density. Population estimates and confidence intervals at the 2 sites were 54 (CI 50-59) on 52.9 km 2 and 87 (CI 82-94) on 73.6 km2. The TLS performed at the same sites resulted in population ranges of 7-34 and 36-63 and always produced a lower population index than the mark-resight population estimate with a larger range. Mark-resight simulations with varying male:female ratios of marks indicated that this ratio was important in designing a population study on prairie-chickens. Confidence intervals for estimates when no marks were placed on females at the 2 sites (CI 46-50, 76-84) did not overlap confidence intervals when 40% of marks were placed on females (CI 54-64, 91-109). Population estimates derived using this mark-resight technique were apparently more accurate than traditional methods and would be more effective in detecting changes in prairie-chicken populations. Our technique could improve prairie-chicken management by providing wildlife biologists and land managers with a tool to estimate the population size and trends of lekking bird species, such as greater prairie-chickens.

  10. Control of Carbon Nanotube Density and Tower Height in an Array

    NASA Technical Reports Server (NTRS)

    Delzeit, Lance D. (Inventor); Schipper, John F. (Inventor)

    2010-01-01

    A method for controlling density or tower height of carbon nanotube (CNT) arrays grown in spaced apart first and second regions on a substrate. CNTs having a first density range (or first tower height range) are grown in the first region using a first source temperature range for growth. Subsequently or simultaneously, CNTs having a second density range (or second tower height range), having an average density (or average tower height) in the second region different from the average density (or average tower height) for the first region, are grown in the second region, using supplemental localized hearing for the second region. Application for thermal dissipation and/or dissipation of electrical charge or voltage in an electronic device are discussed.

  11. Learning and adaptation in the management of waterfowl harvests

    USGS Publications Warehouse

    Johnson, Fred A.

    2011-01-01

    A formal framework for the adaptive management of waterfowl harvests was adopted by the U.S. Fish and Wildlife Service in 1995. The process admits competing models of waterfowl population dynamics and harvest impacts, and relies on model averaging to compute optimal strategies for regulating harvest. Model weights, reflecting the relative ability of the alternative models to predict changes in population size, are used in the model averaging and are updated each year based on a comparison of model predictions and observations of population size. Since its inception the adaptive harvest program has focused principally on mallards (Anas platyrhynchos), which constitute a large portion of the U.S. waterfowl harvest. Four competing models, derived from a combination of two survival and two reproductive hypotheses, were originally assigned equal weights. In the last year of available information (2007), model weights favored the weakly density-dependent reproductive hypothesis over the strongly density-dependent one, and the additive mortality hypothesis over the compensatory one. The change in model weights led to a more conservative harvesting policy than what was in effect in the early years of the program. Adaptive harvest management has been successful in many ways, but nonetheless has exposed the difficulties in defining management objectives, in predicting and regulating harvests, and in coping with the tradeoffs inherent in managing multiple waterfowl stocks exposed to a common harvest. The key challenge now facing managers is whether adaptive harvest management as an institution can be sufficiently adaptive, and whether the knowledge and experience gained from the process can be reflected in higher-level policy decisions.

  12. Population characteristics of a recovering US Virgin Islands red hind spawning aggregation following protection

    PubMed Central

    Nemeth, Richard S.

    2006-01-01

    Many species of groupers form spawning aggregations, dramatic events where 100s to 1000s of individuals gather annually at specific locations for reproduction. Spawning aggregations are often targeted by local fishermen, making them extremely vulnerable to over fishing. The Red Hind Bank Marine Conservation District located in St. Thomas, United States Virgin Islands, was closed seasonally in 1990 and closed permanently in 1999 to protect an important red hind Epinephelus guttatus spawning site. This study provides some of the first information on the population response of a spawning aggregation located within a marine protected area. Tag-and-release fishing and fish transects were used to evaluate population characteristics and habitat utilization patterns of a red hind spawning aggregation between 1999 and 2004. Compared with studies conducted before the permanent closure, the average size of red hind increased mostly during the seasonal closure period (10 cm over 12 yr), but the maximum total length of male red hind increased by nearly 7 cm following permanent closure. Average density and biomass of spawning red hind increased by over 60% following permanent closure whereas maximum spawning density more than doubled. Information from tag returns indicated that red hind departed the protected area following spawning and migrated 6 to 33 km to a ca. 500 km2 area. Protection of the spawning aggregation site may have also contributed to an overall increase in the size of red hind caught in the commercial fishery, thus increasing the value of the grouper fishery for local fishermen. PMID:16612415

  13. Survival and metamorphosis of low-density populations of larval sea lampreys (Petromyzon marinus) in streams following lampricide treatment

    USGS Publications Warehouse

    Johnson, Nicholas S.; Swink, William D.; Brenden, Travis O.; Slade, Jeffrey W.; Steeves, Todd B.; Fodale, Michael F.; Jones, Michael L.

    2014-01-01

    Sea lamprey Petromyzon marinus control in the Great Lakes primarily involves application of lampricides to streams where larval production occurs to kill larvae prior to their metamorphosing and entering the lakes as parasites (juveniles). Because lampricides are not 100% effective, larvae that survive treatment maymetamorphose before streams are again treated. Larvae that survive treatment have not beenwidely studied, so their dynamics are notwell understood.Wetagged and released larvae in six Great Lake tributaries following lampricide treatment and estimated vital demographic rates using multistate tag-recovery models. Model-averaged larval survivals ranged from 56.8 to 57.6%. Model-averaged adult recovery rates, which were the product of juvenile survivals and adult capture probabilities, ranged from 6.8 to 9.3%. Using stochastic simulations, we estimated production of juvenile sea lampreys from a hypothetical population of treatment survivors under different growth conditions based on parameter estimates from this research. For fast-growing populations, juvenile production peaked 2 years after treatment. For slow-growing populations, juvenile production was approximately one-third that of fast-growing populations,with production not peaking until 4 years after treatment. Our results suggest that dynamics (i.e., survival, metamorphosis) of residual larval populations are very similar to those of untreated larval populations. Consequently, residual populations do not necessarily warrant special consideration for the purpose of sea lamprey control and can be ranked for treatment along with other populations. Consecutive lampricide treatments, which are under evaluation by the sea lamprey control program, would bemost effective for reducing juvenile production in large, fast-growing populations.

  14. Pooled Results From 5 Validation Studies of Dietary Self-Report Instruments Using Recovery Biomarkers for Energy and Protein Intake

    PubMed Central

    Freedman, Laurence S.; Commins, John M.; Moler, James E.; Arab, Lenore; Baer, David J.; Kipnis, Victor; Midthune, Douglas; Moshfegh, Alanna J.; Neuhouser, Marian L.; Prentice, Ross L.; Schatzkin, Arthur; Spiegelman, Donna; Subar, Amy F.; Tinker, Lesley F.; Willett, Walter

    2014-01-01

    We pooled data from 5 large validation studies of dietary self-report instruments that used recovery biomarkers as references to clarify the measurement properties of food frequency questionnaires (FFQs) and 24-hour recalls. The studies were conducted in widely differing US adult populations from 1999 to 2009. We report on total energy, protein, and protein density intakes. Results were similar across sexes, but there was heterogeneity across studies. Using a FFQ, the average correlation coefficients for reported versus true intakes for energy, protein, and protein density were 0.21, 0.29, and 0.41, respectively. Using a single 24-hour recall, the coefficients were 0.26, 0.40, and 0.36, respectively, for the same nutrients and rose to 0.31, 0.49, and 0.46 when three 24-hour recalls were averaged. The average rate of under-reporting of energy intake was 28% with a FFQ and 15% with a single 24-hour recall, but the percentages were lower for protein. Personal characteristics related to under-reporting were body mass index, educational level, and age. Calibration equations for true intake that included personal characteristics provided improved prediction. This project establishes that FFQs have stronger correlations with truth for protein density than for absolute protein intake, that the use of multiple 24-hour recalls substantially increases the correlations when compared with a single 24-hour recall, and that body mass index strongly predicts under-reporting of energy and protein intakes. PMID:24918187

  15. Neighborhood Disparities in the Restaurant Food Environment.

    PubMed

    Martinez-Donate, Ana P; Espino, Jennifer Valdivia; Meinen, Amy; Escaron, Anne L; Roubal, Anne; Nieto, Javier; Malecki, Kristen

    2016-11-01

    Restaurant meals account for a significant portion of the American diet. Investigating disparities in the restaurant food environment can inform targeted interventions to increase opportunities for healthy eating among those who need them most. To examine neighborhood disparities in restaurant density and the nutrition environment within restaurants among a statewide sample of Wisconsin households. Households (N = 259) were selected from the 2009-2010 Survey of the Health of Wisconsin (SHOW), a population-based survey of Wisconsin adults. Restaurants in the household neighborhood were enumerated and audited using the Nutrition Environment Measures Survey for Restaurants (NEMS-R). Neighborhoods were defined as a 2- and 5-mile street-distance buffer around households in urban and non-urban areas, respectively. Adjusted linear regression models identified independent associations between sociodemographic household characteristics and neighborhood restaurant density and nutrition environment scores. On average, each neighborhood contained approximately 26 restaurants. On average, restaurants obtained 36.1% of the total nutrition environment points. After adjusting for household characteristics, higher restaurant density was associated with both younger and older household average age (P < .05), all white households (P = .01), and urban location (P < .001). Compared to rural neighborhoods, urban and suburban neighborhoods had slightly higher (ie, healthier) nutrition environment scores (P < .001). The restaurant food environment in Wisconsin neighborhoods varies by age, race, and urbanicity, but offers ample room for improvement across socioeconomic groups and urbanicity levels. Future research must identify policy and environmental interventions to promote healthy eating in all restaurants, especially in young and/or rural neighborhoods in Wisconsin.

  16. Ion Thermal Decoupling and Species Separation in Shock-Driven Implosions

    DOE PAGES

    Rinderknecht, Hans G.; Rosenberg, M. J.; Li, C. K.; ...

    2015-01-14

    Here, anomalous reduction of the fusion yields by 50% and anomalous scaling of the burn-averaged ion temperatures with the ion-species fraction has been observed for the first time in D 3He-filled shock-driven inertial confinement fusion implosions. Two ion kinetic mechanisms are used to explain the anomalous observations: thermal decoupling of the D and 3He populations and diffusive species separation. The observed insensitivity of ion temperature to a varying deuterium fraction is shown to be a signature of ion thermal decoupling in shock-heated plasmas. The burn-averaged deuterium fraction calculated from the experimental data demonstrates a reduction in the average core deuteriummore » density, as predicted by simulations that use a diffusion model. Accounting for each of these effects in simulations reproduces the observed yield trends.« less

  17. Modelling the effect of pyrethroid use intensity on mite population density for walnuts.

    PubMed

    Zhan, Yu; Fan, Siqi; Zhang, Minghua; Zalom, Frank

    2015-01-01

    Published studies relating pyrethroid use and subsequent mite outbreaks have largely been based on laboratory and field experiments, with some inferring a result of increased miticide use. The present study derived a mathematical model proposed to quantify the effect of pyrethroid use intensity on mite population density. The model was validated against and parameterized with actual field-level pyrethroid and miticide use data from 1995 to 2009 for California walnuts, where the miticide use intensity was a proxy of the mite population density. The parameterized model was MI = 1.61 - 0.89 · exp(-93.31PI) (RMSE = 0.13; R(2) = 0.69; P < 0.01), where PI and MI are the average pyrethroid and miticide use intensity in small intervals respectively. A three-range scheme was presented to quantify pesticide applications based on the change rate of MI to PI. Specific for California walnuts, the PI range of 0-0.025 kg ha(-1) was identified as the rapidly increasing range where MI increased vastly when PI increased. Results confirmed that more miticide was used, presumably to prevent or control mite resurgence when pyrethroids were applied, a practice that is not only costly but might be expected to aggravate mite resistance to miticides and increase risk associated with these chemicals to the environment and human health. © 2014 Society of Chemical Industry.

  18. Using ring width correlations to study the effects of plantation density on wood density and anatomical properties of red pine (Pinus resinosa Ait.)

    Treesearch

    J. Y. Zhu; C. T. Scott; K. L. Scallon; G. C. Myers

    2006-01-01

    This study demonstrated that average ring width (or average annual radial growth rate) is a reliable parameter to quantify the effects of tree plantation ndensity (growth suppression) on wood density and tracheid anatomical properties. The average ring width successfully correlated wood density and tracheid anatomical properties of red pines (Pinus resinosa Ait.) from...

  19. Population density of Sotalia guianensis (Cetacea: Delphinidae) in the Cananéia region, Southeastern Brazil.

    PubMed

    Havukainen, Liisa; Monteiro, Emygdio Leite de Araujo Fiho; Filla, Gislaine de Fatima

    2011-09-01

    Population density in cetaceans can be estimated through photo-identification, mark-recapture, land-based observations and visual estimative. We the aim to contribute with conservation strategies, we used line transects (distance method) to estimate the population density of the river dolphin, S. guianensis, in the estuarine region of Cananéia, Southeastern Brazil. The study, developed from May 2003 until April 2004, during dry and rainy seasons and different times of the day, included a sampling area divided into three sectors according to their proximity to the open sea: Sector I (the closest to the open sea); Sector II (with a large flow of fresh water and a salient declivity); and Sector III (with a large flow of fresh water and non salient declivity). Onboard random sampling was carried out in all three sectors, and dolphins seen from the bow to 90 degrees on both port and starboard sides, were registered along with their position and distance from the boat. The total density found was 12.41 ind/km2 (CV = 25.53%) with an average of 2.2 individuals per group for both periods of the day, morning and afternoon. Densities also varied between dry and rainy seasons, being lower in the first with 5.77 ind/km2 (CV = 27.87%) than in the second 20.28 ind/km2 (CV = 31.95%), respectively. Regarding the three sectors, a non-causal heterogeneous distribution was found: Sector I was the most populated (D = 33.10 ind/km2, CV = 13.34%), followed by Sector II (D = 7.8 ind/km2, CV = 21.07%) and Sector III (D = 3.04 ind/km2, CV = 34.04%). The aforementioned area, due to its proximity to the open sea, has the highest salinity level and therefore has the greatest chance of holding most of the marine fish schools which can be cornered by dolphins on high declivity areas during fishing activities. This suggests that food availability may be the most important factor on the river dolphin's distribution in the estuary. Similar studies will contribute to a better understanding of these populations and are essential for future conservation strategies.

  20. Probabilistic Model for Laser Damage to the Human Retina

    DTIC Science & Technology

    2012-03-01

    the beam. Power density may be measured in radiant exposure, J cm2 , or by irradiance , W cm2 . In the experimental database used in this study and...to quan- tify a binary response, either lethal or non-lethal, within a population such as insects or rats. In directed energy research, probit...value of the normalized Arrhenius damage integral. In a one-dimensional simulation, the source term is determined as a spatially averaged irradiance (W

  1. Population Density and Distribution of Wheat Bugs Infesting Durum Wheat in Sardinia, Italy

    PubMed Central

    Salis, Luigi; Goula, Marta; Izquierdo, Jordi; Gordún, Elena

    2013-01-01

    Wheat is a very important crop in Italy, and is infested by wheat bugs belonging to the genera Eurygaster (Hemiptera: Scutellaridae) and Aelia (Hemiptera: Pentatomidae). Many wheat bug infestations have been reported in the north, south, and center of Italy, both in the past as well as recently. The present study was carried out in Sardinia, Italy, during two years (2007 and 2008). The objective of this study was to determine the species and distribution of wheat bugs in durum wheat fields in Sardinia, and to estimate their population density in order to know the incidence of the pest on the island. Sampling took place twice a year (May and June) in three zones, representative of durum wheat cropping in the island. Four species of wheat bugs were found; the predominant species was Eurygaster austriaca (Schrank), followed by Aelia germari (Kuster), Eurygaster maura L., and Aelia acuminata L. The average density of wheat bugs was low (1.1 individuals/m2), but in certain areas it was above the damage threshold (4 individuals/m2). For this reason, the conclusion of the study is that this pest should be monitored in order to control outbreaks and prevent their further spread. PMID:23906035

  2. Population density and distribution of wheat bugs infesting durum wheat in Sardinia, Italy.

    PubMed

    Salis, Luigi; Goula, Marta; Izquierdo, Jordi; Gordún, Elena

    2013-01-01

    Wheat is a very important crop in Italy, and is infested by wheat bugs belonging to the genera Eurygaster (Hemiptera: Scutellaridae) and Aelia (Hemiptera: Pentatomidae). Many wheat bug infestations have been reported in the north, south, and center of Italy, both in the past as well as recently. The present study was carried out in Sardinia, Italy, during two years (2007 and 2008). The objective of this study was to determine the species and distribution of wheat bugs in durum wheat fields in Sardinia, and to estimate their population density in order to know the incidence of the pest on the island. Sampling took place twice a year (May and June) in three zones, representative of durum wheat cropping in the island. Four species of wheat bugs were found; the predominant species was Eurygaster austriaca (Schrank), followed by Aelia germari (Kuster), Eurygaster maura L., and Aelia acuminata L. The average density of wheat bugs was low (1.1 individuals/m²), but in certain areas it was above the damage threshold (4 individuals/m²). For this reason, the conclusion of the study is that this pest should be monitored in order to control outbreaks and prevent their further spread.

  3. The public health benefits of reducing fine particulate matter through conversion to cleaner heating fuels in New York City.

    PubMed

    Kheirbek, Iyad; Haney, Jay; Douglas, Sharon; Ito, Kazuhiko; Caputo, Steven; Matte, Thomas

    2014-12-02

    In recent years, both New York State and City issued regulations to reduce emissions from burning heating oil. To assess the benefits of these programs in New York City, where the density of emissions and vulnerable populations vary greatly, we simulated the air quality benefits of scenarios reflecting no action, partial, and complete phase-out of high-sulfur heating fuels using the Community MultiScale Air Quality (CMAQ) model conducted at a high spatial resolution (1 km). We evaluated the premature mortality and morbidity benefits of the scenarios within 42 city neighborhoods and computed benefits by neighborhood poverty status. The complete phase-out scenario reduces annual average fine particulate matter (PM2.5) by an estimated 0.71 μg/m(3) city-wide (average of 1 km estimates, 10-90th percentile: 0.1-1.6 μg/m(3)), avoiding an estimated 290 premature deaths, 180 hospital admissions for respiratory and cardiovascular disease, and 550 emergency department visits for asthma each year. The largest improvements were seen in areas of highest building and population density and the majority of benefits have occurred through the partial phase out of high-sulfur heating fuel already achieved. While emissions reductions were greatest in low-poverty neighborhoods, health benefits are estimated to be greatest in high-poverty neighborhoods due to higher baseline morbidity and mortality rates.

  4. Uncovering hidden variation in polyploid wheat.

    PubMed

    Krasileva, Ksenia V; Vasquez-Gross, Hans A; Howell, Tyson; Bailey, Paul; Paraiso, Francine; Clissold, Leah; Simmonds, James; Ramirez-Gonzalez, Ricardo H; Wang, Xiaodong; Borrill, Philippa; Fosker, Christine; Ayling, Sarah; Phillips, Andrew L; Uauy, Cristobal; Dubcovsky, Jorge

    2017-02-07

    Comprehensive reverse genetic resources, which have been key to understanding gene function in diploid model organisms, are missing in many polyploid crops. Young polyploid species such as wheat, which was domesticated less than 10,000 y ago, have high levels of sequence identity among subgenomes that mask the effects of recessive alleles. Such redundancy reduces the probability of selection of favorable mutations during natural or human selection, but also allows wheat to tolerate high densities of induced mutations. Here we exploited this property to sequence and catalog more than 10 million mutations in the protein-coding regions of 2,735 mutant lines of tetraploid and hexaploid wheat. We detected, on average, 2,705 and 5,351 mutations per tetraploid and hexaploid line, respectively, which resulted in 35-40 mutations per kb in each population. With these mutation densities, we identified an average of 23-24 missense and truncation alleles per gene, with at least one truncation or deleterious missense mutation in more than 90% of the captured wheat genes per population. This public collection of mutant seed stocks and sequence data enables rapid identification of mutations in the different copies of the wheat genes, which can be combined to uncover previously hidden variation. Polyploidy is a central phenomenon in plant evolution, and many crop species have undergone recent genome duplication events. Therefore, the general strategy and methods developed herein can benefit other polyploid crops.

  5. Biomass of freshwater turtles: a geographic comparison

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Congdon, J.D.; Greene, J.L.; Gibbons, J.W.

    1986-01-01

    Standing crop biomass of freshwater turtles and minimum annual biomass of egg production were calculated for marsh and farm pond habitats in South Caroling and in Michigan. The species in South Carolina included Chelydra serpentina, Deirochelys reticularia, Kinosternon subrubrum, Pseudemys floridana, P. scripta and Sternotherus odoratus. The species in Michigan were Chelydra serpentina, Chrysemys picta and Emydoidea blandingi. Biomass was also determined for a single species population of P. scripta on a barrier island near Charleston, South Carolina. Population density and biomass of Pseudemys scripta in Green Pond on Capers Island were higher than densities and biomass of the entiremore » six-species community studied on the mainland. In both the farm pond and marsh habitat in South Carolina P. scripta was the numerically dominant species and had the highest biomass. In Michigan, Chrysemys picta was the numerically dominant species; however, the biomass of Chelydra serpentina was higher. The three-species community in Michigan in two marshes (58 kg ha/sup -1/ and 46 kg ha/sup -1/) and farm ponds (23 kg ha/sup -1/) had lower biomasses than did the six-species community in a South Carolina marsh (73 kg/sup -1/). Minimum annual egg production by all species in South Carolina averaged 1.93 kg ha/sup -1/ and in Michigan averaged 2.89 kg ha/sup -1/ of marsh.« less

  6. Understanding relationships among abundance, extirpation, and climate at ecoregional scales.

    PubMed

    Beever, Erik A; Dobrowski, S Z; Long, J; Mynsberge, A R; Piekielek, N B

    2013-07-01

    Recent research on mountain-dwelling species has illustrated changes in species distributional patterns in response to climate change. Abundance of a species will likely provide an earlier warning indicator of change than will occupancy, yet relationships between abundance and climatic factors have received less attention. We tested whether predictors of counts of American pikas (Ochotona princeps) during surveys from the Great Basin region in 1994-1999 and 2003-2008 differed between the two periods. Additionally, we tested whether various modeled aspects of ecohydrology better predicted relative density than did average annual precipitation, and whether risk of site-wide extirpation predicted subsequent population counts of pikas. We observed several patterns of change in pika abundance at range edges that likely constitute early warnings of distributional shifts. Predictors of pika abundance differed strongly between the survey periods, as did pika extirpation patterns previously reported from this region. Additionally, maximum snowpack and growing-season precipitation resulted in better-supported models than those using average annual precipitation, and constituted two of the top three predictors of pika density in the 2000s surveys (affecting pikas perhaps via vegetation). Unexpectedly, we found that extirpation risk positively predicted subsequent population size. Our results emphasize the need to clarify mechanisms underlying biotic responses to recent climate change at organism-relevant scales, to inform management and conservation strategies for species of concern.

  7. Understanding relationships among abundance, extirpation,and climate at ecoregional scales

    USGS Publications Warehouse

    Beever, Erik A.; Solomon Dubrowski,; ,; ,; J. Long,; ,; A. Mysnberge,; Piekielek, N. B.

    2014-01-01

    Recent research on mountain-dwelling species has illustrated changes in species’ distributional patterns in response to climate change. Abundance of a species will likely provide an earlier warning indicator of change than will occupancy, yet relationships between abundance and climatic factors have received less attention. We tested whether predictors of counts of American pikas (Ochotona princeps) during surveys from the Great Basin region in 1994–1999 and 2003–2008 differed between the two periods. Additionally, we tested whether various modeled aspects of ecohydrology better predicted relative density than did average annual precipitation, and whether risk of site-wide extirpation predicted subsequent population counts of pikas. We observed several patterns of change in pika abundance at range edges that likely constitute early warnings of distributional shifts. Predictors of pika abundance differed strongly between the survey periods, as did pika extirpation patterns previously reported from this region. Additionally, maximum snowpack and growing-season precipitation resulted in better-supported models than those using average annual precipitation, and constituted two of the top three predictors of pika density in the 2000s surveys (affecting pikas perhaps via vegetation). Unexpectedly, we found that extirpation risk positively predicted subsequent population size. Our results emphasize the need to clarify mechanisms underlying biotic responses to recent climate change at organism-relevant scales, to inform management and conservation strategies for species of concern.

  8. Application of MODIS GPP to Forecast Risk of Hantavirus Pulmonary Syndrome Based on Fluctuations in Reservoir Population Density

    NASA Astrophysics Data System (ADS)

    Loehman, R.; Heinsch, F. A.; Mills, J. N.; Wagoner, K.; Running, S.

    2003-12-01

    Recent predictive models for hantavirus pulmonary syndrome (HPS) have used remotely sensed spectral reflectance data to characterize risk areas with limited success. We present an alternative method using gross primary production (GPP) from the MODIS sensor to estimate the effects of biomass accumulation on population density of Peromyscus maniculatus (deer mouse), the principal reservoir species for Sin Nombre virus (SNV). The majority of diagnosed HPS cases in North America are attributed to SNV, which is transmitted to humans through inhalation of excretions and secretions from infected rodents. A logistic model framework is used to evaluate MODIS GPP, temperature, and precipitation as predictors of P. maniculatus density at established trapping sites across the western United States. Rodent populations are estimated using monthly minimum number alive (MNA) data for 2000 through 2002. Both local meteorological data from nearby weather stations and 1.25 degree x 1 degree gridded data from the NASA DAO were used in the regression model to determine the spatial sensitivity of the response. MODIS eight-day GPP data (1-km resolution) were acquired and binned to monthly average and monthly sum GPP for 3km x 3km grids surrounding each rodent trapping site. The use of MODIS GPP to forecast HPS risk may result in a marked improvement over past reflectance-based risk area characterizations. The MODIS GPP product provides a vegetation dynamics estimate that is unique to disease models, and targets the fundamental ecological processes responsible for increased rodent density and amplified disease risk.

  9. Population dynamics and angler exploitation of the unique muskellunge population in Shoepack Lake, Voyageurs National Park, Minnesota

    USGS Publications Warehouse

    Frohnauer, N.K.; Pierce, C.L.; Kallemeyn, L.W.

    2007-01-01

    A unique population of muskellunge Esox masquinongy inhabits Shoepack Lake in Voyageurs National Park, Minnesota. Little is known about its status, dynamics, and angler exploitation, and there is concern for the long-term viability of this population. We used intensive sampling and mark-recapture methods to quantify abundance, survival, growth, condition, age at maturity and fecundity and angler surveys to quantify angler pressure, catch rates, and exploitation. During our study, heavy rain washed out a dam constructed by beavers Castor canadensis which regulates the water level at the lake outlet, resulting in a nearly 50% reduction in surface area. We estimated a population size of 1,120 adult fish at the beginning of the study. No immediate reduction in population size was detected in response to the loss of lake area, although there was a gradual, but significant, decline in population size over the 2-year study. Adults grew less than 50 mm per year, and relative weight (W r) averaged roughly 80. Anglers were successful in catching, on average, two fish during a full day of angling, but harvest was negligible. Shoepack Lake muskellunge exhibit much slower growth rates and lower condition, but much higher densities and angler catch per unit effort (CPUE), than other muskellunge populations. The unique nature, limited distribution, and location of this population in a national park require special consideration for management. The results of this study provide the basis for assessing the long-term viability of the Shoepack Lake muskellunge population through simulations of long-term population dynamics and genetically effective population size. ?? Copyright by the American Fisheries Society 2007.

  10. Favre-Averaged Turbulence Statistics in Variable Density Mixing of Buoyant Jets

    NASA Astrophysics Data System (ADS)

    Charonko, John; Prestridge, Kathy

    2014-11-01

    Variable density mixing of a heavy fluid jet with lower density ambient fluid in a subsonic wind tunnel was experimentally studied using Particle Image Velocimetry and Planar Laser Induced Fluorescence to simultaneously measure velocity and density. Flows involving the mixing of fluids with large density ratios are important in a range of physical problems including atmospheric and oceanic flows, industrial processes, and inertial confinement fusion. Here we focus on buoyant jets with coflow. Results from two different Atwood numbers, 0.1 (Boussinesq limit) and 0.6 (non-Boussinesq case), reveal that buoyancy is important for most of the turbulent quantities measured. Statistical characteristics of the mixing important for modeling these flows such as the PDFs of density and density gradients, turbulent kinetic energy, Favre averaged Reynolds stress, turbulent mass flux velocity, density-specific volume correlation, and density power spectra were also examined and compared with previous direct numerical simulations. Additionally, a method for directly estimating Reynolds-averaged velocity statistics on a per-pixel basis is extended to Favre-averages, yielding improved accuracy and spatial resolution as compared to traditional post-processing of velocity and density fields.

  11. Declining populations of the fingernail clam Musculium transversum in the upper Mississippi River

    USGS Publications Warehouse

    Wilson, D.M.; Naimo, T.J.; Weiner, J.G.; Anderson, R.V.; Sandheinrich, M.B.; Sparks, Richard E.

    1995-01-01

    We examined recent temporal trends in the abundance of fingernail clams Musculium transversum (formerly Sphaerium transversum) in the upper Mississippi River. Historical data on densities of fingernail clams were obtained from regional scientists and published literature. We also sampled benthos in six navigation pools in summer 1991, finding very few fingernail clams. The combined data set, including historical data and sampling results, extended from 1973 to 1992 and was sufficient to statistically evaluate trends in densities of fingernail clams in eight pools. Populations of fingernail clams declined significantly in five of the eight pools examined (Pools 2, 5, 7, 9, and 19), which spanned a 700-km reach of river from St. Paul, Minnesota, to Keokuk, Iowa. Densities in Pool 19, which had the longest historical record on fingernail clam abundance, averaged 30 000 m super(-2) in 1985 and progressively declined to zero in 1990. Combined data from all eight pools showed a significant decline in abundance of fingernail clams. An evaluation of potential causal factors led us to hypothesize that the population declines in Pools 2 to 9 were linked to point-source pollution rather than to dredging activity or commercial navigation traffic. In Pool 19, the declines of fingernail clams may have resulted from low-flow conditions during drought periods, but the causal mechanisms by which low flow influences fingernail clam abundance are unclear. The decrease in fingernail clam populations may adversely affect certain fish and wildlife, such as migrating lesser scaup Aythya affinis, which feed heavily on the small mollusk. Moreover, the decreases in populations of this pollution-sensitive mollusk may signal a large-scale deterioration in the health of this riverine ecosystem.

  12. Demography and decline of the Mentasta Caribou Herd, Alaska

    USGS Publications Warehouse

    Jenkins, Kurt J.; Barten, Neil L.

    2005-01-01

    We evaluated population trends in the Mentasta caribou (Rangifer tarandus (L., 1758)) herd in Wrangell a?? St. Elias National Park and Preserve, Alaska, from 1990 to 1997 and determined factors contributing to its decline. We postulated that predation-related mortality of adult females and juveniles was the proximate cause of the decline, and that survival of juvenile caribou reflected interactions with winter severity, calving distribution, timing of births, density of caribou, and physical condition of neonates at birth. The population declined at its greatest rate from 1990 to 1993 (r = a??0.32) and at a lower rate from 1994 to 1997 (r = a??0.09). Recruitment (number of calves/100 females during September) averaged 4/100 during the rapid population decline from 1990 to 1993 and 13/100 from 1994 to 1997. Parturition rate of adult females ranged from 65% to 97%. Survival of adult females and juveniles ranged from 0.77 to 0.86 and from 0.00 to 0.22, respectively. Approximately 43%, 59%, and 79% of all juvenile mortality occurred by 1, 2, and 4 weeks of age, respectively. We confirmed predation-related mortality as the primary proximate cause of population decline, with gray wolves (Canis lupus L., 1758), bears (species of the genus Ursus L., 1758), and other predators accounting for 57%, 38%, and 5%, respectively, of all juvenile mortality, and bears causing disproportionate mortality among 0- to 1-week-old neonates. We supported the hypotheses that timing of birth and habitat conditions at the birth site, particularly mottled snow patterns, affected vulnerability and survival of neonates, and birth mass affected survival of juveniles through summer. We speculate that the population will continue to decline before reaching a low-density equilibrium that is sustained by density-dependent changes in the functional responses of predators.

  13. A function accounting for training set size and marker density to model the average accuracy of genomic prediction.

    PubMed

    Erbe, Malena; Gredler, Birgit; Seefried, Franz Reinhold; Bapst, Beat; Simianer, Henner

    2013-01-01

    Prediction of genomic breeding values is of major practical relevance in dairy cattle breeding. Deterministic equations have been suggested to predict the accuracy of genomic breeding values in a given design which are based on training set size, reliability of phenotypes, and the number of independent chromosome segments ([Formula: see text]). The aim of our study was to find a general deterministic equation for the average accuracy of genomic breeding values that also accounts for marker density and can be fitted empirically. Two data sets of 5'698 Holstein Friesian bulls genotyped with 50 K SNPs and 1'332 Brown Swiss bulls genotyped with 50 K SNPs and imputed to ∼600 K SNPs were available. Different k-fold (k = 2-10, 15, 20) cross-validation scenarios (50 replicates, random assignment) were performed using a genomic BLUP approach. A maximum likelihood approach was used to estimate the parameters of different prediction equations. The highest likelihood was obtained when using a modified form of the deterministic equation of Daetwyler et al. (2010), augmented by a weighting factor (w) based on the assumption that the maximum achievable accuracy is [Formula: see text]. The proportion of genetic variance captured by the complete SNP sets ([Formula: see text]) was 0.76 to 0.82 for Holstein Friesian and 0.72 to 0.75 for Brown Swiss. When modifying the number of SNPs, w was found to be proportional to the log of the marker density up to a limit which is population and trait specific and was found to be reached with ∼20'000 SNPs in the Brown Swiss population studied.

  14. Study on Karst Rock Desertification by Human-Nature Interaction: A Case Study of Fengshan County of Guangxi, China

    NASA Astrophysics Data System (ADS)

    Zhou, G.; Su, C.; Zhang, R.; Shi, Y.; Liu, Y.; Ma, Y.

    2014-11-01

    Rocky desertification is a process of soil erosion leading to the bareness of bedrock, and is a kind of typical natural disaster in karst areas, which seriously constrains the developments of local society and economy. This paper studies the impact of human activities on the changes of rocky desertification. With the interpretation of remote sensing images of Fengshan County, GuangXi, China covering 1990, 1995 and 2005, this paper analyzes the relationship of the changes of social and economic index (including population density, agricultural population, rural per capital net income, and farmland) and the changes of rocky desertification in recent 15 years. The results indicate that: in recent 15 years, the average annual growth rate of rocky desertification, rate of change of population density, agricultural population, rural per capita net income, and farmland area are 8.7375 km2 /year, 0.7 person/km2.year, 1153 person/year, 85 Yuan/year, and -269.1 ha/year, respectively. The first three of social and economic indicators at different degrees of growth, show a positive correlation with rocky desertification change. However, farmland area, which has been diminishing, shows a negative correlation with rocky desertification. The influence of human activities for karst rocky desertification can be regarded as population growth - steep slope reclamation - the destruction of vegetation - erosion - rocky desertification - poverty.

  15. Local habitat conditions explain the variation in the strength of self-thinning in a stream salmonid

    PubMed Central

    Myrvold, Knut Marius; Kennedy, Brian P

    2015-01-01

    Self-thinning patterns are frequently used to describe density dependence in populations on timescales shorter than the organism's life span and have been used to infer carrying capacity of the environment. Among mobile animals, this concept has been used to document density dependence in stream salmonids, which compete over access to food and space. The carrying capacity, growth conditions, and initial cohort sizes often vary between streams and stream sections, which would influence the onset and strength of the density dependence. Despite much effort in describing habitat relationships in stream fishes, few studies have explicitly tested how the physical environment affects the slope of the thinning curves. Here, we investigate the prevalence and strength of self-thinning in juvenile stages of a steelhead (Oncorhynchus mykiss) population in Idaho, USA. Further, we investigate the roles of local physical habitat and metabolic constraints in explaining the variation in thinning curves among study sites in the watershed. Only yearling steelhead exhibited an overall significant thinning trend, but the slope of the mass–density relationship (−0.53) was shallower than predicted by theory and reported from empirical studies. There was no detectable relationship in subyearling steelhead. Certain abiotic factors explained a relatively large portion of the variation in the strength of the self-thinning among the study reaches. For subyearling steelhead, the slopes were negatively associated with the average water depth and flow velocity in the study sites, whereas slopes in yearlings were steeper in sites that incurred a higher metabolic cost. Our results show that the prevalence and strength of density dependence in natural fish populations can vary across heterogeneous watersheds and can be more pronounced during certain stages of a species' life history, and that environmental factors can mediate the extent to which density dependence is manifested in predictable ways. PMID:26380659

  16. Accurate genomic predictions for BCWD resistance in rainbow trout are achieved using low-density SNP panels: Evidence that long-range LD is a major contributing factor.

    PubMed

    Vallejo, Roger L; Silva, Rafael M O; Evenhuis, Jason P; Gao, Guangtu; Liu, Sixin; Parsons, James E; Martin, Kyle E; Wiens, Gregory D; Lourenco, Daniela A L; Leeds, Timothy D; Palti, Yniv

    2018-06-05

    Previously accurate genomic predictions for Bacterial cold water disease (BCWD) resistance in rainbow trout were obtained using a medium-density single nucleotide polymorphism (SNP) array. Here, the impact of lower-density SNP panels on the accuracy of genomic predictions was investigated in a commercial rainbow trout breeding population. Using progeny performance data, the accuracy of genomic breeding values (GEBV) using 35K, 10K, 3K, 1K, 500, 300 and 200 SNP panels as well as a panel with 70 quantitative trait loci (QTL)-flanking SNP was compared. The GEBVs were estimated using the Bayesian method BayesB, single-step GBLUP (ssGBLUP) and weighted ssGBLUP (wssGBLUP). The accuracy of GEBVs remained high despite the sharp reductions in SNP density, and even with 500 SNP accuracy was higher than the pedigree-based prediction (0.50-0.56 versus 0.36). Furthermore, the prediction accuracy with the 70 QTL-flanking SNP (0.65-0.72) was similar to the panel with 35K SNP (0.65-0.71). Genomewide linkage disequilibrium (LD) analysis revealed strong LD (r 2  ≥ 0.25) spanning on average over 1 Mb across the rainbow trout genome. This long-range LD likely contributed to the accurate genomic predictions with the low-density SNP panels. Population structure analysis supported the hypothesis that long-range LD in this population may be caused by admixture. Results suggest that lower-cost, low-density SNP panels can be used for implementing genomic selection for BCWD resistance in rainbow trout breeding programs. © 2018 The Authors. This article is a U.S. Government work and is in the public domain in the USA. Journal of Animal Breeding and Genetics published by Blackwell Verlag GmbH.

  17. [Population pressure: a factor of political destabilization].

    PubMed

    Tallon, F

    1993-04-01

    Political stability throughout the world appears to be greater in countries with slowly growing populations than in those with rapid growth. Population is not the only influence on political stability, however. The relationship between political stability and development is strong. The rich countries with the slowest growth are the most stable, while poor developing countries with rapid growth suffer from chronic instability. Demographic pressure and density are not the same thing and must be distinguished. A fragile environment like that of the Sahel will experience demographic pressure despite low density. Japan has a greater population density than Rwanda and little cultivable land, but the population has a high standard of living. demographic pressure is not comparable in Japan and Rwanda because Japan has slow population growth and stable democratic political institutions. The rate of growth seems to be a more important element in destabilization than density. Rapid growth creates enormous political tensions especially when profound ethnic divisions exist, and it complicates problems of government by encouraging rapid urbanization. The unbalanced age structures resulting from rapid growth hinder the satisfaction of employment, educational, and health care needs for the ever-increasing masses of young people. 49% of Rwanda's population is under 15 and 66% is under 25. Rwanda is already densely populated, with around 300 inhabitants/sq km, and its population is projected to double in 20 years. 95% of the population is dependent on agriculture, but by 1988 the average landholding per family was only 1.25 hectares and 58% of families did not grown sufficient food for household needs. Further reduction in the size of holdings or a growing landless population will have multiple consequences. Urban migration will inevitably increase, bringing with it all the problems so evident in other poor countries where the process is more advanced than in Rwanda. Chaotic urbanization is in itself a destabilizing force. The phenomenon of potential political destabilization due to demographic pressure had been analyzed according to different criteria. Unconstitutional sudden changes of government, incomplete civil and political liberties, violent conflicts between ethnic and other subgroups, and the frustrated aspirations of large numbers of young persons are among possible indicators of political instability. A report by the Population Crisis committee which evaluated these indicators in different countries placed Rwanda 39th among 120 countries for potential destabilization. But the period of observation avoided the 1973 disturbances and the 1990 war in Rwanda. The results would undoubtedly be considerably less favorable if the period of observation extended to the present.

  18. Bioavailability and possible benefits of wheat intake naturally enriched with selenium and its products.

    PubMed

    Djujić, I S; Jozanov-Stankov, O N; Milovac, M; Janković, V; Djermanović, V

    2000-12-01

    Bioavailability and possible benefits of wheat intake naturally enriched with selenium and its products was tested. Wheat obtained by application of an original combination and procedure for foliar supplementation of plants with Se was characterized on the average by five times higher content of Se, the main form being L-(+)-selenomethionine (SeMet). Substitution of Se-deficient wheat by wheat naturally enriched with Se and its products contributed to the increase of daily intake on the average by 18 microg (12-35 microg) in volunteers, which is more than 50% of the average daily intake. Six weeks after the beginning of its application, increased daily intake of Se brought about the increase of its concentration in the plasma of the examined persons by 53%, in their erythrocytes by 37%, in their hair by 44%, and in their urine by 54%. This result was comparable to the effect obtained in the course of an 8-wk daily intake of supplements with 100 microg Se in the form of enriched bakery yeast. Analysis of glutathione peroxidase (GSH-Px) activity in blood, thiobarbituric acid reactive substances (TBARS) in plasma, lipid parameters (total cholesterol, high-density lipoprotein cholesterol, low-density lipoprotein cholesterol, and triglycerides), and glucose in serum of volunteers showed that the increased Se intake induced increased GSH-Px activity in blood and decreased concentrations of TBARS, lipid parameters, and glucose in blood. Using only one crop (wheat enriched with Se), the existing deficiency of Se in our population can be alleviated. In this way, one-fourth of our population with lower Se intake than 21 microg/d will satisfy basal requirements, whereas one-half will become moderately deficient in Se instead of distinctly deficient in Se.

  19. The Strength of Selection against Neanderthal Introgression

    PubMed Central

    Juric, Ivan

    2016-01-01

    Hybridization between humans and Neanderthals has resulted in a low level of Neanderthal ancestry scattered across the genomes of many modern-day humans. After hybridization, on average, selection appears to have removed Neanderthal alleles from the human population. Quantifying the strength and causes of this selection against Neanderthal ancestry is key to understanding our relationship to Neanderthals and, more broadly, how populations remain distinct after secondary contact. Here, we develop a novel method for estimating the genome-wide average strength of selection and the density of selected sites using estimates of Neanderthal allele frequency along the genomes of modern-day humans. We confirm that East Asians had somewhat higher initial levels of Neanderthal ancestry than Europeans even after accounting for selection. We find that the bulk of purifying selection against Neanderthal ancestry is best understood as acting on many weakly deleterious alleles. We propose that the majority of these alleles were effectively neutral—and segregating at high frequency—in Neanderthals, but became selected against after entering human populations of much larger effective size. While individually of small effect, these alleles potentially imposed a heavy genetic load on the early-generation human–Neanderthal hybrids. This work suggests that differences in effective population size may play a far more important role in shaping levels of introgression than previously thought. PMID:27824859

  20. Implications of changes in tropical shifting cultivation intensification on land productivity and GHG-related biogeochemistry

    NASA Astrophysics Data System (ADS)

    Bustier, Bernard; Ngoy, Alfred; Pietsch, Stephan; Mosnier, Aline

    2017-04-01

    Traditional shifting cultivation used to be a sustainable type of land use for the subsistence of populations in tropical rainforests. The vast resource of moist tropical forests together with low population densities allowed for long fallow periods on sparsely distributed slash and burn parcels with large areas of untouched forest in between. Population growth and concomitant increase in land demand for subsistence as well as increasing infrastructure development for commercial forestry, cash crops and mining, however, altered the picture over recent decades. As a result, fallow periods were reduced due to lack of pristine land. In this study we use field data and modeling results from the Congo Basin to assess the impacts of reduced fallow periods on Carbon sequestration dynamics using a BGC model calibrated and validated with > 150 research plots distributed over the western Congo Basin and representing different management and land use histories. We find that the average carbon sequestration rate reduces over the number of cultivation cycles and that a reduction of the fallow from 10 years to 7 years reduce the average carbon sequestration between 13 and 21% and from 7 years to 4 years between 23 and 29% depending on soil fertility. Results will be discussed in the context of population growth and changes in environmetal conditions.

  1. Multispecies reaction-diffusion systems

    NASA Astrophysics Data System (ADS)

    Aghamohammadi, A.; Fatollahi, A. H.; Khorrami, M.; Shariati, A.

    2000-10-01

    Multispecies reaction-diffusion systems, for which the time evolution equations of correlation functions become a closed set, are considered. A formal solution for the average densities is found. Some special interactions and the exact time dependence of the average densities in these cases are also studied. For the general case, the large-time behavior of the average densities has also been obtained.

  2. Life history trait analysis of the entomopathogenic nematode Steinernema feltiae provides the basis for prediction of dauer juvenile yields in monoxenic liquid culture.

    PubMed

    Addis, Temesgen; Teshome, Asmamaw; Strauch, Olaf; Ehlers, Ralf-Udo

    2016-05-01

    Entomopathogenic nematodes (Steinernema spp.) are used in integrated pest management to control insect pests in cryptic environments. The nematodes are mass produced in monoxenic liquid culture with their symbiotic bacteria Xenorhabdus spp. For a better understanding of nematode population dynamics, the life history traits (LHTs) of the entomopathogenic nematode Steinernema feltiae were assessed at 25 °C by observing single pairs of male and female nematodes using a hanging drop technique. To investigate the influence of different food supplies on nematode reproduction, the LHTs were assessed with a daily supply of 5 ×, 10 × and 20 × 10(9) cells ml(-1) of the nematode's bacterial symbiont Xenorhabdus bovienii in semi-solid nematode growth gelrite (NGG) medium. Increasing bacterial density had a significant positive influence on the average number of offspring produced, which ranged from 359 to 813 per female. The intrinsic rate of natural increase r m, which ranges from 1.10 to 1.19 day(-1), was neither influenced by the bacterial density, nor was the mean generation time T (5.12-5.25 days) and population doubling time (PDT) (0.64-0.59 days). The average lifespan of reproductive females, which ranged from 6.7 to 7.3 days, was positively correlated with bacterial density. A positive correlation between female body volume and bacterial density was recorded (R = 0.67) as well as a significant positive correlation between female body size and offspring production (R = 0.89) in hanging drops. Whether these data can be used to predict nematode yields in liquid culture was tested. The total female body volume calculated as the average female body volume × total number of parental females per millilitre 3 days after nematode inoculation was positively correlated (R = 0.72) with nematode yields. The total female body volume on process day 3 is thus a good indicator for the estimation of nematode yield at the end of the process (12-15 days post dauer juvenile (DJ) inoculation) in both Erlenmeyer flasks and bioreactors. With a mean deviation of 9467 DJs ml(-1), the error resembles approximately 5 % of the final DJ yields.

  3. Dengue diversity across spatial and temporal scales: Local structure and the effect of host population size.

    PubMed

    Salje, Henrik; Lessler, Justin; Maljkovic Berry, Irina; Melendrez, Melanie C; Endy, Timothy; Kalayanarooj, Siripen; A-Nuegoonpipat, Atchareeya; Chanama, Sumalee; Sangkijporn, Somchai; Klungthong, Chonticha; Thaisomboonsuk, Butsaya; Nisalak, Ananda; Gibbons, Robert V; Iamsirithaworn, Sopon; Macareo, Louis R; Yoon, In-Kyu; Sangarsang, Areerat; Jarman, Richard G; Cummings, Derek A T

    2017-03-24

    A fundamental mystery for dengue and other infectious pathogens is how observed patterns of cases relate to actual chains of individual transmission events. These pathways are intimately tied to the mechanisms by which strains interact and compete across spatial scales. Phylogeographic methods have been used to characterize pathogen dispersal at global and regional scales but have yielded few insights into the local spatiotemporal structure of endemic transmission. Using geolocated genotype (800 cases) and serotype (17,291 cases) data, we show that in Bangkok, Thailand, 60% of dengue cases living <200 meters apart come from the same transmission chain, as opposed to 3% of cases separated by 1 to 5 kilometers. At distances <200 meters from a case (encompassing an average of 1300 people in Bangkok), the effective number of chains is 1.7. This number rises by a factor of 7 for each 10-fold increase in the population of the "enclosed" region. This trend is observed regardless of whether population density or area increases, though increases in density over 7000 people per square kilometer do not lead to additional chains. Within Thailand these chains quickly mix, and by the next dengue season viral lineages are no longer highly spatially structured within the country. In contrast, viral flow to neighboring countries is limited. These findings are consistent with local, density-dependent transmission and implicate densely populated communities as key sources of viral diversity, with home location the focal point of transmission. These findings have important implications for targeted vector control and active surveillance. Copyright © 2017, American Association for the Advancement of Science.

  4. Microbiologic progression of post-traumatic peritonitis.

    PubMed

    Nyström, P O; Skau, T; Johansson, L; Lennquist, S

    1985-08-01

    The natural course of post-traumatic peritonitis was studied in 17 pigs. Following a standardized missile wound to the abdomen the pigs were operated on and observed in continuous anesthesia for periods up to 3 days. Bacterial density was measured in peritoneal exudate, parietal peritoneum, laparotomy wound, and projectile entrance and exit wounds at this operation and also at relaparotomy, which terminated the experiments. The bacterial density showed significant perioperative fall, averaging X 1,905 in peritoneal exudate and X 40 in parietal peritoneum. From a low postoperative level (mean c. 10(2)/ml or /gm), bacterial density increased significantly in the observation period. In peritoneal exudate the increase was X 4,073 and in peritoneum X 2,630. In laparotomy and projectile entrance and exit wounds the corresponding figures were X 234, X 126, and X 15. The bacterial population's perioperative reduction and subsequent proliferation were interpreted as reflecting the host defense, which accordingly was limited in capacity and duration.

  5. Waterbird use of coastal impoundments and management implications in east-central Florida

    NASA Technical Reports Server (NTRS)

    Breininger, David R.; Smith, Rebecca B.

    1990-01-01

    Monthly surveys were conducted on Kennedy Space Center for one year to determine densities of waterbirds within impounded salt marshes that were predominantly open water with little emergent vegetation. The objective was to assess the importance of these impoundments to waterbirds, particularly wading birds, which are species of special conservation concern. Water-level management for mosquito control and waterfowl provided habitat for an abundance of ducks, shorebirds, coots, and wading birds. Average densities throughout the year for these groups were 5.26, 412, 2.80, and 2.20 birds/ha, respectively. The majority of waterfowl were present during the winter. Shorebirds were most common during spring migration. Wading bird densities increased with declining water level. Due to the extensive alteration and development of coastal wetlands in central Florida, properly managed impoundments may provide important feeding areas for maintaining certain waterbird populations.

  6. Allee effect: the story behind the stabilization or extinction of microbial ecosystem.

    PubMed

    Goswami, Madhurankhi; Bhattacharyya, Purnita; Tribedi, Prosun

    2017-03-01

    A population exhibiting Allee effect shows a positive correlation between population fitness and population size or density. Allee effect decides the extinction or conservation of a microbial population and thus appears to be an important criterion in population ecology. The underlying factor of Allee effect that decides the stabilization and extinction of a particular population density is the threshold or the critical density of their abundance. According to Allee, microbial populations exhibit a definite, critical or threshold density, beyond which the population fitness of a particular population increases with the rise in population density and below it, the population fitness goes down with the decrease in population density. In particular, microbial population displays advantageous traits such as biofilm formation, expression of virulence genes, spore formation and many more only at a high population density. It has also been observed that microorganisms exhibiting a lower population density undergo complete extinction from the residual microbial ecosystem. In reference to Allee effect, decrease in population density or size introduces deleterious mutations among the population density through genetic drift. Mutations are carried forward to successive generations resulting in its accumulation among the population density thus reducing its microbial fitness and thereby increasing the risk of extinction of a particular microbial population. However, when the microbial load is high, the chance of genetic drift is less, and through the process of biofilm formation, the cooperation existing among the microbial population increases that increases the microbial fitness. Thus, the high microbial population through the formation of microbial biofilm stabilizes the ecosystem by increasing fitness. Taken together, microbial fitness shows positive correlation with the ecosystem conservation and negative correlation with ecosystem extinction.

  7. Ensemble Averaged Probability Density Function (APDF) for Compressible Turbulent Reacting Flows

    NASA Technical Reports Server (NTRS)

    Shih, Tsan-Hsing; Liu, Nan-Suey

    2012-01-01

    In this paper, we present a concept of the averaged probability density function (APDF) for studying compressible turbulent reacting flows. The APDF is defined as an ensemble average of the fine grained probability density function (FG-PDF) with a mass density weighting. It can be used to exactly deduce the mass density weighted, ensemble averaged turbulent mean variables. The transport equation for APDF can be derived in two ways. One is the traditional way that starts from the transport equation of FG-PDF, in which the compressible Navier- Stokes equations are embedded. The resulting transport equation of APDF is then in a traditional form that contains conditional means of all terms from the right hand side of the Navier-Stokes equations except for the chemical reaction term. These conditional means are new unknown quantities that need to be modeled. Another way of deriving the transport equation of APDF is to start directly from the ensemble averaged Navier-Stokes equations. The resulting transport equation of APDF derived from this approach appears in a closed form without any need for additional modeling. The methodology of ensemble averaging presented in this paper can be extended to other averaging procedures: for example, the Reynolds time averaging for statistically steady flow and the Reynolds spatial averaging for statistically homogeneous flow. It can also be extended to a time or spatial filtering procedure to construct the filtered density function (FDF) for the large eddy simulation (LES) of compressible turbulent reacting flows.

  8. Density-dependent natural selection and trade-offs in life history traits.

    PubMed

    Mueller, L D; Guo, P Z; Ayala, F J

    1991-07-26

    Theories of density-dependent natural selection state that at extreme population densities evolution produces alternative life histories due to trade-offs. The trade-offs are presumed to arise because those genotypes with highest fitness at high population densities will not also have high fitness at low density and vice-versa. These predictions were tested by taking samples from six populations of Drosophila melanogaster kept at low population densities (r-populations) for nearly 200 generations and placing them in crowded cultures (K-populations). After 25 generations in the crowded cultures, the derived K-populations showed growth rate and productivity that at high densities were elevated relative to the controls, but at low density were depressed.

  9. Linking social and ecological systems to sustain coral reef fisheries.

    PubMed

    Cinner, Joshua E; McClanahan, Timothy R; Daw, Tim M; Graham, Nicholas A J; Maina, Joseph; Wilson, Shaun K; Hughes, Terence P

    2009-02-10

    The ecosystem goods and services provided by coral reefs are critical to the social and economic welfare of hundreds of millions of people, overwhelmingly in developing countries [1]. Widespread reef degradation is severely eroding these goods and services, but the socioeconomic factors shaping the ways that societies use coral reefs are poorly understood [2]. We examine relationships between human population density, a multidimensional index of socioeconomic development, reef complexity, and the condition of coral reef fish populations in five countries across the Indian Ocean. In fished sites, fish biomass was negatively related to human population density, but it was best explained by reef complexity and a U-shaped relationship with socioeconomic development. The biomass of reef fishes was four times lower at locations with intermediate levels of economic development than at locations with both low and high development. In contrast, average biomass inside fishery closures was three times higher than in fished sites and was not associated with socioeconomic development. Sustaining coral reef fisheries requires an integrated approach that uses tools such as protected areas to quickly build reef resources while also building capacities and capital in societies over longer time frames to address the complex underlying causes of reef degradation.

  10. Measurement and simulation of passive fast-ion D-alpha emission from the DIII-D tokamak

    DOE PAGES

    Bolte, Nathan G.; Heidbrink, William W.; Pace, David; ...

    2016-09-14

    Spectra of passive fast-ion D-alpha (FIDA) light from beam ions that charge exchange with background neutrals are measured and simulated. The fast ions come from three sources: ions that pass through the diagnostic sightlines on their first full orbit, an axisymmetric confined population, and ions that are expelled into the edge region by instabilities. A passive FIDA simulation (P-FIDASIM) is developed as a forward model for the spectra of the first-orbit fast ions and consists of an experimentally-validated beam deposition model, an ion orbit-following code, a collisional-radiative model, and a synthetic spectrometer. Model validation consists of the simulation of 86more » experimental spectra that are obtained using 6 different neutral beam fast-ion sources and 13 different lines of sight. Calibrated spectra are used to estimate the neutral density throughout the cross-section of the tokamak. The resulting 2D neutral density shows the expected increase toward each X-point with average neutral densities of 8 X 10 9 cm -3 at the plasma boundary and 1 X 10 11 cm -3 near the wall. Here, fast ions that are on passing orbits are expelled by the sawtooth instability more readily than trapped ions. In a sample discharge, approximately 1% of the fast-ion population is ejected into the high neutral density region per sawtooth crash.« less

  11. Rare variant APOC3 R19X is associated with cardio-protective profiles in a diverse population-based survey as part of the Epidemiologic Architecture for Genes Linked to Environment Study.

    PubMed

    Crawford, Dana C; Dumitrescu, Logan; Goodloe, Robert; Brown-Gentry, Kristin; Boston, Jonathan; McClellan, Bob; Sutcliffe, Cara; Wiseman, Rachel; Baker, Paxton; Pericak-Vance, Margaret A; Scott, William K; Allen, Melissa; Mayo, Ping; Schnetz-Boutaud, Nathalie; Dilks, Holli H; Haines, Jonathan L; Pollin, Toni I

    2014-12-01

    A founder mutation was recently discovered and described as conferring favorable lipid profiles and reduced subclinical atherosclerotic disease in a Pennsylvania Amish population. Preliminary data have suggested that this null mutation APOC3 R19X (rs76353203) is rare in the general population. To better describe the frequency and lipid profile in the general population, we as part of the Population Architecture using Genomics and Epidemiology I Study and the Epidemiological Architecture for Genes Linked to Environment Study genotyped rs76353203 in 1113 Amish participants from Ohio and Indiana and 19 613 participants from the National Health and Nutrition Examination Surveys (NHANES III, 1999 to 2002, and 2007 to 2008). We found no carriers among the Ohio and Indiana Amish. Of the 19 613 NHANES participants, we identified 31 participants carrying the 19X allele, for an overall allele frequency of 0.08%. Among fasting adults, the 19X allele was associated with lower triglycerides (n=7603; β=-71.20; P=0.007) and higher high-density lipoprotein cholesterol (n=8891; β=15.65; P=0.0002) and, although not significant, lower low-density lipoprotein cholesterol (n=6502; β= -4.85; P=0.68) after adjustment for age, sex, and race/ethnicity. On average, 19X allele participants had approximately half the triglyceride levels (geometric means, 51.3 to 69.7 versus 134.6 to 141.3 mg/dL), >20% higher high-density lipoprotein cholesterol levels (geometric means, 56.8 to 74.4 versus 50.38 to 53.36 mg/dL), and lower low-density lipoprotein cholesterol levels (geometric means, 104.5 to 128.6 versus 116.1 to 125.7 mg/dL) compared with noncarrier participants. These data demonstrate that APOC3 19X exists in the general US population in multiple racial/ethnic groups and is associated with cardio-protective lipid profiles. © 2014 American Heart Association, Inc.

  12. Macroscopic neural mass model constructed from a current-based network model of spiking neurons.

    PubMed

    Umehara, Hiroaki; Okada, Masato; Teramae, Jun-Nosuke; Naruse, Yasushi

    2017-02-01

    Neural mass models (NMMs) are efficient frameworks for describing macroscopic cortical dynamics including electroencephalogram and magnetoencephalogram signals. Originally, these models were formulated on an empirical basis of synaptic dynamics with relatively long time constants. By clarifying the relations between NMMs and the dynamics of microscopic structures such as neurons and synapses, we can better understand cortical and neural mechanisms from a multi-scale perspective. In a previous study, the NMMs were analytically derived by averaging the equations of synaptic dynamics over the neurons in the population and further averaging the equations of the membrane-potential dynamics. However, the averaging of synaptic current assumes that the neuron membrane potentials are nearly time invariant and that they remain at sub-threshold levels to retain the conductance-based model. This approximation limits the NMM to the non-firing state. In the present study, we newly propose a derivation of a NMM by alternatively approximating the synaptic current which is assumed to be independent of the membrane potential, thus adopting a current-based model. Our proposed model releases the constraint of the nearly constant membrane potential. We confirm that the obtained model is reducible to the previous model in the non-firing situation and that it reproduces the temporal mean values and relative power spectrum densities of the average membrane potentials for the spiking neurons. It is further ensured that the existing NMM properly models the averaged dynamics over individual neurons even if they are spiking in the populations.

  13. Validity criteria for Fermi's golden rule scattering rates applied to metallic nanowires.

    PubMed

    Moors, Kristof; Sorée, Bart; Magnus, Wim

    2016-09-14

    Fermi's golden rule underpins the investigation of mobile carriers propagating through various solids, being a standard tool to calculate their scattering rates. As such, it provides a perturbative estimate under the implicit assumption that the effect of the interaction Hamiltonian which causes the scattering events is sufficiently small. To check the validity of this assumption, we present a general framework to derive simple validity criteria in order to assess whether the scattering rates can be trusted for the system under consideration, given its statistical properties such as average size, electron density, impurity density et cetera. We derive concrete validity criteria for metallic nanowires with conduction electrons populating a single parabolic band subjected to different elastic scattering mechanisms: impurities, grain boundaries and surface roughness.

  14. A general diagram for estimating pore size of ultrafiltration and reverse osmosis membranes

    NASA Technical Reports Server (NTRS)

    Sarbolouki, M. N.

    1982-01-01

    A slit sieve model has been used to develop a general correlation between the average pore size of the upstream surface of a membrane and the molecular weight of the solute which it retains by better than 80%. The pore size is determined by means of the correlation using the high retention data from an ultrafiltration (UF) or a reverse osmosis (RO) experiment. The pore population density can also be calculated from the flux data via appropriate equations.

  15. Dynamics of a recovering Arctic bird population: the importance of climate, density dependence, and site quality

    USGS Publications Warehouse

    Bruggeman, Jason E.; Swem, Ted; Andersen, David E.; Kennedy, Patricia L.; Nigro, Debora A.

    2015-01-01

    Intrinsic and extrinsic factors affect vital rates and population-level processes, and understanding these factors is paramount to devising successful management plans for wildlife species. For example, birds time migration in response, in part, to local and broadscale climate fluctuations to initiate breeding upon arrival to nesting territories, and prolonged inclement weather early in the breeding season can inhibit egg-laying and reduce productivity. Also, density-dependent regulation occurs in raptor populations, as territory size is related to resource availability. Arctic Peregrine Falcons (Falco peregrinus tundrius; hereafter Arctic peregrine) have a limited and northern breeding distribution, including the Colville River Special Area (CRSA) in the National Petroleum Reserve–Alaska, USA. We quantified influences of climate, topography, nest productivity, prey habitat, density dependence, and interspecific competition affecting Arctic peregrines in the CRSA by applying the Dail-Madsen model to estimate abundance and vital rates of adults on nesting cliffs from 1981 through 2002. Arctic peregrine abundance increased throughout the 1980s, which spanned the population's recovery from DDT-induced reproductive failure, until exhibiting a stationary trend in the 1990s. Apparent survival rate (i.e., emigration; death) was negatively correlated with the number of adult Arctic peregrines on the cliff the previous year, suggesting effects of density-dependent population regulation. Apparent survival and arrival rates (i.e., immigration; recruitment) were higher during years with earlier snowmelt and milder winters, and apparent survival was positively correlated with nesting season maximum daily temperature. Arrival rate was positively correlated with average Arctic peregrine productivity along a cliff segment from the previous year and initial abundance was positively correlated with cliff height. Higher cliffs with documented higher productivity (presumably indicative of higher-quality habitat), are a priority for continued protection from potential nearby development and disturbance to minimize population-level impacts. Climate change may affect Arctic peregrines in multiple ways, including through access to more snow-free nest sites and a lengthened breeding season that may increase likelihood of nest success. Our work provides insight into factors affecting a population during and after recovery, and demonstrates how the Dail-Madsen model can be used for any unmarked population with multiple years of abundance data collected through repeated surveys.

  16. Dynamics of newly established elk populations

    USGS Publications Warehouse

    Sargeant, G.A.; Oehler, M.W.

    2007-01-01

    The dynamics of newly established elk (Cervus elaphus) populations can provide insights about maximum sustainable rates of reproduction, survival, and increase. However, data used to estimate rates of increase typically have been limited to counts and rarely have included complementary estimates of vital rates. Complexities of population dynamics cannot be understood without considering population processes as well as population states. We estimated pregnancy rates, survival rates, age ratios, and sex ratios for reintroduced elk at Theodore Roosevelt National Park, North Dakota, USA; combined vital rates in a population projection model; and compared model projections with observed elk numbers and population ratios. Pregnancy rates in January (early in the second trimester of pregnancy) averaged 54.1% (SE = 5.4%) for subadults and 91.0% (SE = 1.7%) for adults, and 91.6% of pregnancies resulted in recruitment at 8 months. Annual survival rates of adult females averaged 0.96 (95% CI = 0.94-0.98) with hunting included and 0.99 (95% CI = 0.97-0.99) with hunting excluded from calculations. Our fitted model explained 99.8% of past variation in population estimates and represents a useful new tool for short-term management planning. Although we found no evidence of temporal variation in vital rates, variation in population composition caused substantial variation in projected rates of increase (??=1.20-1.36). Restoring documented hunter harvests and removals of elk by the National Park Service led to a potential rate of ?? = 1.26. Greater rates of increase substantiated elsewhere were within the expected range of chance variation, given our model and estimates of vital rates. Rates of increase realized by small elk populations are too variable to support inferences about habitat quality or density dependence.

  17. DOE Office of Scientific and Technical Information (OSTI.GOV)

    Larsen, Peter H.; LaCommare, Kristina H.; Eto, Joseph H.

    Here, this study examines the relationship between annual changes in electricity reliability reported by a large cross-section of U.S. electricity distribution utilities over a period of 13 years and a broad set of potential explanatory variables, including weather and utility characteristics. We find statistically significant correlations between the average number of power interruptions experienced annually and above average wind speeds, precipitation, lightning strikes, and a measure of population density: customers per line mile. We also find significant relationships between the average number of minutes of power interruptions experienced and above average wind speeds, precipitation, cooling degree-days, and one strategy usedmore » to mitigate the impacts of severe weather: the amount of underground transmission and distribution line miles. Perhaps most importantly, we find a significant time trend of increasing annual average number of minutes of power interruptions over time—especially when interruptions associated with extreme weather are included. Lastly, the research method described in this analysis can provide a basis for future efforts to project long-term trends in reliability and the associated benefits of strategies to improve grid resiliency to severe weather—both in the U.S. and abroad.« less

  18. The association of Antarctic krill Euphausia superba with the under-ice habitat.

    PubMed

    Flores, Hauke; van Franeker, Jan Andries; Siegel, Volker; Haraldsson, Matilda; Strass, Volker; Meesters, Erik Hubert; Bathmann, Ulrich; Wolff, Willem Jan

    2012-01-01

    The association of Antarctic krill Euphausia superba with the under-ice habitat was investigated in the Lazarev Sea (Southern Ocean) during austral summer, autumn and winter. Data were obtained using novel Surface and Under Ice Trawls (SUIT), which sampled the 0-2 m surface layer both under sea ice and in open water. Average surface layer densities ranged between 0.8 individuals m(-2) in summer and autumn, and 2.7 individuals m(-2) in winter. In summer, under-ice densities of Antarctic krill were significantly higher than in open waters. In autumn, the opposite pattern was observed. Under winter sea ice, densities were often low, but repeatedly far exceeded summer and autumn maxima. Statistical models showed that during summer high densities of Antarctic krill in the 0-2 m layer were associated with high ice coverage and shallow mixed layer depths, among other factors. In autumn and winter, density was related to hydrographical parameters. Average under-ice densities from the 0-2 m layer were higher than corresponding values from the 0-200 m layer collected with Rectangular Midwater Trawls (RMT) in summer. In winter, under-ice densities far surpassed maximum 0-200 m densities on several occasions. This indicates that the importance of the ice-water interface layer may be under-estimated by the pelagic nets and sonars commonly used to estimate the population size of Antarctic krill for management purposes, due to their limited ability to sample this habitat. Our results provide evidence for an almost year-round association of Antarctic krill with the under-ice habitat, hundreds of kilometres into the ice-covered area of the Lazarev Sea. Local concentrations of postlarval Antarctic krill under winter sea ice suggest that sea ice biota are important for their winter survival. These findings emphasise the susceptibility of an ecological key species to changing sea ice habitats, suggesting potential ramifications on Antarctic ecosystems induced by climate change.

  19. The Association of Antarctic Krill Euphausia superba with the Under-Ice Habitat

    PubMed Central

    Flores, Hauke; van Franeker, Jan Andries; Siegel, Volker; Haraldsson, Matilda; Strass, Volker; Meesters, Erik Hubert; Bathmann, Ulrich; Wolff, Willem Jan

    2012-01-01

    The association of Antarctic krill Euphausia superba with the under-ice habitat was investigated in the Lazarev Sea (Southern Ocean) during austral summer, autumn and winter. Data were obtained using novel Surface and Under Ice Trawls (SUIT), which sampled the 0–2 m surface layer both under sea ice and in open water. Average surface layer densities ranged between 0.8 individuals m−2 in summer and autumn, and 2.7 individuals m−2 in winter. In summer, under-ice densities of Antarctic krill were significantly higher than in open waters. In autumn, the opposite pattern was observed. Under winter sea ice, densities were often low, but repeatedly far exceeded summer and autumn maxima. Statistical models showed that during summer high densities of Antarctic krill in the 0–2 m layer were associated with high ice coverage and shallow mixed layer depths, among other factors. In autumn and winter, density was related to hydrographical parameters. Average under-ice densities from the 0–2 m layer were higher than corresponding values from the 0–200 m layer collected with Rectangular Midwater Trawls (RMT) in summer. In winter, under-ice densities far surpassed maximum 0–200 m densities on several occasions. This indicates that the importance of the ice-water interface layer may be under-estimated by the pelagic nets and sonars commonly used to estimate the population size of Antarctic krill for management purposes, due to their limited ability to sample this habitat. Our results provide evidence for an almost year-round association of Antarctic krill with the under-ice habitat, hundreds of kilometres into the ice-covered area of the Lazarev Sea. Local concentrations of postlarval Antarctic krill under winter sea ice suggest that sea ice biota are important for their winter survival. These findings emphasise the susceptibility of an ecological key species to changing sea ice habitats, suggesting potential ramifications on Antarctic ecosystems induced by climate change. PMID:22384073

  20. Demographic Variables for Wild Asian Elephants Using Longitudinal Observations

    PubMed Central

    de Silva, Shermin; Webber, C. Elizabeth; Weerathunga, U. S.; Pushpakumara, T. V.; Weerakoon, Devaka K.; Wittemyer, George

    2013-01-01

    Detailed demographic data on wild Asian elephants have been difficult to collect due to habitat characteristics of much of the species’ remaining range. Such data, however, are critical for understanding and modeling population processes in this endangered species. We present data from six years of an ongoing study of Asian elephants (Elephas maximus) in Uda Walawe National Park, Sri Lanka. This relatively undisturbed population numbering over one thousand elephants is individually monitored, providing cohort-based information on mortality and reproduction. Reproduction was seasonal, such that most births occurred during the long inter-monsoon dry season and peaked in May. During the study, the average age at first reproduction was 13.4 years and the 50th percentile inter-birth interval was approximately 6 years. Birth sex ratios did not deviate significantly from parity. Fecundity was relatively stable throughout the observed reproductive life of an individual (ages 11–60), averaging between 0.13–0.17 female offspring per individual per year. Mortalities and injuries based on carcasses and disappearances showed that males were significantly more likely than females to be killed or injured through anthropogenic activity. Overall, however, most observed injuries did not appear to be fatal. This population exhibits higher fecundity and density relative to published estimates on other Asian elephant populations, possibly enhanced by present range constriction. Understanding the factors responsible for these demographic dynamics can shed insight on the future needs of this elephant population, with probable parallels to other populations in similar settings. PMID:24376581

  1. Demographic variables for wild Asian elephants using longitudinal observations.

    PubMed

    de Silva, Shermin; Webber, C Elizabeth; Weerathunga, U S; Pushpakumara, T V; Weerakoon, Devaka K; Wittemyer, George

    2013-01-01

    Detailed demographic data on wild Asian elephants have been difficult to collect due to habitat characteristics of much of the species' remaining range. Such data, however, are critical for understanding and modeling population processes in this endangered species. We present data from six years of an ongoing study of Asian elephants (Elephas maximus) in Uda Walawe National Park, Sri Lanka. This relatively undisturbed population numbering over one thousand elephants is individually monitored, providing cohort-based information on mortality and reproduction. Reproduction was seasonal, such that most births occurred during the long inter-monsoon dry season and peaked in May. During the study, the average age at first reproduction was 13.4 years and the 50(th) percentile inter-birth interval was approximately 6 years. Birth sex ratios did not deviate significantly from parity. Fecundity was relatively stable throughout the observed reproductive life of an individual (ages 11-60), averaging between 0.13-0.17 female offspring per individual per year. Mortalities and injuries based on carcasses and disappearances showed that males were significantly more likely than females to be killed or injured through anthropogenic activity. Overall, however, most observed injuries did not appear to be fatal. This population exhibits higher fecundity and density relative to published estimates on other Asian elephant populations, possibly enhanced by present range constriction. Understanding the factors responsible for these demographic dynamics can shed insight on the future needs of this elephant population, with probable parallels to other populations in similar settings.

  2. Ecology of a nesting red-shouldered hawk population

    USGS Publications Warehouse

    Stewart, R.E.

    1949-01-01

    An ecological study of a nesting Red-shouldered Hawk population was made over a 185 square mile area on the Coastal Plain of Maryland in 1947. The courting and nesting season extended from late February until late June.....During the nesting season a combination of fairly extensive flood-plain forest with adjacent clearings appears to meet the major ecological requirements of the Red-shouldered Hawk in this region. A total of 51 pairs was found in the study area, occupying about 42 square miles or 23% of the total area studied. The population density on the land that was suitable for this species was about 1 pair per .8 of a square mile, while the density for the entire study area would be only about 1 pair per 3.6 square miles.....Nests were spaced fairly evenly over most of the flood-plain forests, especially in areas where the width.of the flood plain was relatively constant. There was an inverse correlation between the width of the flood plain and the distances between nests in adjacent territories. The nests were all situated in fairly large trees and were from 28 feet to 77 feet above the ground, averaging 50. They were found in 14 different species of trees, all deciduous.....The Barred Owl and Red-shouldered Hawk were commonly associated together in the same lowland habitats. Other raptores were all largely restricted to upland habitats....The average number of young in 47 occupied nests following the hatching period was 2.7 with extremes of 1 and 4. Only 3 out of 52 nests (6%) were found deserted at this time....The food habits of nestling Red-shouldered Hawks are very diversified. They feed on many types of warm-blooded and cold-blooded vertebrates as well as invertebrates.

  3. High biodiversity on a deep-water reef in the eastern Fram Strait.

    PubMed

    Meyer, Kirstin S; Soltwedel, Thomas; Bergmann, Melanie

    2014-01-01

    We report on the distribution and abundance of megafauna on a deep-water rocky reef (1796-2373 m) in the Fram Strait, west of Svalbard. Biodiversity and population density are high, with a maximum average of 26.7±0.9 species m(-2) and 418.1±49.6 individuals m(-2) on the east side of the reef summit. These figures contrast with the surrounding abyssal plain fauna, with an average of only 18.1±1.4 species and 29.4±4.3 individuals m(-2) (mean ± standard error). The east side of the reef summit, where the highest richness and density of fauna are found, faces into the predominant bottom current, which likely increases in speed to the summit and serves as a source of particulate food for the numerous suspension feeders present there. We conclude that the observed faunal distribution patterns could be the result of hydrodynamic patterns and food availability above and around the reef. To our knowledge, this study is the first to describe the distribution and diversity of benthic fauna on a rocky reef in deep water.

  4. Potential of Crops Uncommon to Alabama for Management of Root-Knot and Soybean Cyst Nematodes

    PubMed Central

    Rodríguez-Kábana, R.; King, P. S.; Robertson, D. G.; Weaver, C. F.

    1988-01-01

    Vigna unguiculata, Cassia fasiculata, and Sesamum indicum did not support Meloidogyne arenaria, M. incognita, or Heterodera glycines race 4 in greenhouse studies with soils from peanut and soybean fields. Fagopyron eseulentum, Cyamopsis tetragonoloba, and Cucurbita pepo were hosts to the two Meloidogyne spp. but were nonhosts to H. glycines. Meloidogyne arenaria and M. incognita galled but reproduced poorly in the roots of three types of Amaranthus cruentus, and low densities of these two Meloidogyne spp. (< 10 second-stage juveniles/100 cm³ soil) occurred in soil cultivated with this crop. In a field study no juveniles of M. arenaria determined at peanut harvest were recovered from plots with Ricinus communis, Gossypium hirsutum, Aeschynomene americana, C. fasiculata, or S. indicum. Peanut plots averaged 120 juveniles/100 cm³ soil. Application of aldicarb (12 kg a.i./ha broadcast) in peanut resulted in an average of 27 juveniles/100 cm³ soil. Several crops were as effective as aldicarb treatment for reducing soil juvenile population densities of M. arenaria. PMID:19290317

  5. Advanced PIC-MCC simulation for the investigation of step-ionization effect in intermediate-pressure capacitively coupled plasmas

    NASA Astrophysics Data System (ADS)

    Kim, Jin Seok; Hur, Min Young; Kim, Chang Ho; Kim, Ho Jun; Lee, Hae June

    2018-03-01

    A two-dimensional parallelized particle-in-cell simulation has been developed to simulate a capacitively coupled plasma reactor. The parallelization using graphics processing units is applied to resolve the heavy computational load. It is found that the step-ionization plays an important role in the intermediate gas pressure of a few Torr. Without the step-ionization, the average electron density decreases while the effective electron temperature increases with the increase of gas pressure at a fixed power. With the step-ionization, however, the average electron density increases while the effective electron temperature decreases with the increase of gas pressure. The cases with the step-ionization agree well with the tendency of experimental measurement. The electron energy distribution functions show that the population of electrons having intermediate energy from 4.2 to 12 eV is relaxed by the step-ionization. Also, it was observed that the power consumption by the electrons is increasing with the increase of gas pressure by the step-ionization process, while the power consumption by the ions decreases with the increase of gas pressure.

  6. High population density enhances recruitment and survival of a harvested coral reef fish.

    PubMed

    Wormald, Clare L; Steele, Mark A; Forrester, Graham E

    2013-03-01

    A negative relationship between population growth and population density (direct density dependence) is necessary for population regulation and is assumed in most models of harvested populations. Experimental tests for density dependence are lacking for large-bodied, harvested fish because of the difficulty of manipulating population density over large areas. We studied a harvested coral reef fish, Lutjanus apodus (schoolmaster snapper), using eight large, isolated natural reefs (0.4-1.6 ha) in the Bahamas as replicates. An initial observational test for density dependence was followed by a manipulation of population density. The manipulation weakened an association between density and shelter-providing habitat features and revealed a positive effect of population density on recruitment and survival (inverse density dependence), but no effect of density on somatic growth. The snappers on an individual reef were organized into a few shoals, and we hypothesize that large shoals on high-density reefs were less vulnerable to large piscivores (groupers and barracudas) than the small shoals on low-density reefs. Reductions in predation risk for individuals in large social groups are well documented, but because snapper shoals occupied reefs the size of small marine reserves, these ecological interactions may influence the outcome of management actions.

  7. Effect of the target power density on high-power impulse magnetron sputtering of copper

    NASA Astrophysics Data System (ADS)

    Kozák, Tomáš

    2012-04-01

    We present a model analysis of high-power impulse magnetron sputtering of copper. We use a non-stationary global model based on the particle and energy conservation equations in two zones (the high density plasma ring above the target racetrack and the bulk plasma region), which makes it possible to calculate time evolutions of the averaged process gas and target material neutral and ion densities, as well as the fluxes of these particles to the target and substrate during a pulse period. We study the effect of the increasing target power density under conditions corresponding to a real experimental system. The calculated target current waveforms show a long steady state and are in good agreement with the experimental results. For an increasing target power density, an analysis of the particle densities shows a gradual transition to a metal dominated discharge plasma with an increasing degree of ionization of the depositing flux. The average fraction of target material ions in the total ion flux onto the substrate is more than 90% for average target power densities higher than 500 W cm-2 in a pulse. The average ionized fraction of target material atoms in the flux onto the substrate reaches 80% for a maximum average target power density of 3 kW cm-2 in a pulse.

  8. Adaptive harvest management for the Svalbard population of pink-footed geese: 2015 progress summary

    USGS Publications Warehouse

    Johnson, Fred A.; Madsen, Jesper

    2015-01-01

    This document describes progress to date on the development of an adaptive harvest management strategy for maintaining the Svalbard population of pink‐footed geese (Anser brachyrhynchus) near their agreed target level (60,000) by providing for sustainable harvests in Norway and Denmark. This report provides an assessment of the most recent monitoring information (1991-2014) and its implications for the harvest management strategy, and it is an update of an initial assessment for 2013-2015 (see http://pinkfootedgoose.aewa.info/). By combining varying hypotheses about survival and reproduction, a suite of nine models have been developed that represent a wide range of possibilities concerning the extent to which demographic rates are density dependent or independent. Current updated model weights suggest little evidence for density-dependent survival and reproduction, suggesting that the population may have recently experienced a release from density-dependent mechanisms, corresponding to the period of most rapid growth in population size. The optimal harvest strategy for the 2013–2015 hunting seasons prescribed a harvest quota of 15,000 per year. The harvest in the 2014 hunting season was 14,991, compared to 11,081 in 2013, mostly due to an increase in harvest in Denmark during January 2015. The percentage of young in the fall of 2014 was 10.3%, which is lower than average. The observed population size of 59,000 in May 2015 was much lower than expected. For the 2015 hunting season, observed population size and temperature days suggest that an emergency closure should be considered. In the event a harvest of 15,000 is maintained, predicted population size in May 2016 is 51,700 (95% CL: 41,600-64,300), based on observed TempDays = 9 in May 2015 and the most recent model weights. On the other hand, if the season were closed this year, we would expect a population size of 66,700 (95% CL: 53,600-82,900) in May 2016. A total harvest of 6,700 would be expected to result in a 2016 population size at goal (i.e., 60,000).

  9. Density regulation in Northeast Atlantic fish populations: Density dependence is stronger in recruitment than in somatic growth.

    PubMed

    Zimmermann, Fabian; Ricard, Daniel; Heino, Mikko

    2018-05-01

    Population regulation is a central concept in ecology, yet in many cases its presence and the underlying mechanisms are difficult to demonstrate. The current paradigm maintains that marine fish populations are predominantly regulated by density-dependent recruitment. While it is known that density-dependent somatic growth can be present too, its general importance remains unknown and most practical applications neglect it. This study aimed to close this gap by for the first time quantifying and comparing density dependence in growth and recruitment over a large set of fish populations. We fitted density-dependent models to time-series data on population size, recruitment and age-specific weight from commercially exploited fish populations in the Northeast Atlantic Ocean and the Baltic Sea. Data were standardized to enable a direct comparison within and among populations, and estimated parameters were used to quantify the impact of density regulation on population biomass. Statistically significant density dependence in recruitment was detected in a large proportion of populations (70%), whereas for density dependence in somatic growth the prevalence of density dependence depended heavily on the method (26% and 69%). Despite age-dependent variability, the density dependence in recruitment was consistently stronger among age groups and between alternative approaches that use weight-at-age or weight increments to assess growth. Estimates of density-dependent reduction in biomass underlined these results: 97% of populations with statistically significant parameters for growth and recruitment showed a larger impact of density-dependent recruitment on population biomass. The results reaffirm the importance of density-dependent recruitment in marine fishes, yet they also show that density dependence in somatic growth is not uncommon. Furthermore, the results are important from an applied perspective because density dependence in somatic growth affects productivity and catch composition, and therefore the benefits of maintaining fish populations at specific densities. © 2018 The Authors. Journal of Animal Ecology published by John Wiley & Sons Ltd on behalf of British Ecological Society.

  10. Variation of genetic diversity in a rapidly expanding population of the greater long-tailed hamster (Tscherskia triton) as revealed by microsatellites.

    PubMed

    Xu, Laixiang; Xue, Huiliang; Song, Mingjing; Zhao, Qinghua; Dong, Jingping; Liu, Juan; Guo, Yu; Xu, Tongqin; Cao, Xiaoping; Wang, Fusheng; Wang, Shuqing; Hao, Shushen; Yang, Hefang; Zhang, Zhibin

    2013-01-01

    Genetic diversity is essential for persistence of animal populations over both the short- and long-term. Previous studies suggest that genetic diversity may decrease with population decline due to genetic drift or inbreeding of small populations. For oscillating populations, there are some studies on the relationship between population density and genetic diversity, but these studies were based on short-term observation or in low-density phases. Evidence from rapidly expanding populations is lacking. In this study, genetic diversity of a rapidly expanding population of the Greater long-tailed hamsters during 1984-1990, in the Raoyang County of the North China Plain was studied using DNA microsatellite markers. Results show that genetic diversity was positively correlated with population density (as measured by % trap success), and the increase in population density was correlated with a decrease of genetic differentiation between the sub-population A and B. The genetic diversity tended to be higher in spring than in autumn. Variation in population density and genetic diversity are consistent between sub-population A and B. Such results suggest that dispersal is density- and season-dependent in a rapidly expanding population of the Greater long-tailed hamster. For typically solitary species, increasing population density can increase intra-specific attack, which is a driving force for dispersal. This situation is counterbalanced by decreasing population density caused by genetic drift or inbreeding as the result of small population size. Season is a major factor influencing population density and genetic diversity. Meanwhile, roads, used to be considered as geographical isolation, have less effect on genetic differentiation in a rapidly expanding population. Evidences suggest that gene flow (Nm) is positively correlated with population density, and it is significant higher in spring than that in autumn.

  11. Evidence for a Low Bulk Crustal Density for Mars from Gravity and Topography.

    PubMed

    Goossens, Sander; Sabaka, Terence J; Genova, Antonio; Mazarico, Erwan; Nicholas, Joseph B; Neumann, Gregory A

    2017-08-16

    Knowledge of the average density of the crust of a planet is important in determining its interior structure. The combination of high-resolution gravity and topography data has yielded a low density for the Moon's crust, yet for other terrestrial planets the resolution of the gravity field models has hampered reasonable estimates. By using well-chosen constraints derived from topography during gravity field model determination using satellite tracking data, we show that we can robustly and independently determine the average bulk crustal density directly from the tracking data, using the admittance between topography and imperfect gravity. We find a low average bulk crustal density for Mars, 2582 ± 209 kg m -3 . This bulk crustal density is lower than that assumed until now. Densities for volcanic complexes are higher, consistent with earlier estimates, implying large lateral variations in crustal density. In addition, we find indications that the crustal density increases with depth.

  12. A generic individual-based model to simulate morphogenesis, C-N acquisition and population dynamics in contrasting forage legumes.

    PubMed

    Louarn, Gaëtan; Faverjon, Lucas

    2018-04-18

    Individual-based models (IBMs) are promising tools to disentangle plant interactions in multi-species grasslands and foster innovative species mixtures. This study describes an IBM dealing with the morphogenesis, growth and C-N acquisition of forage legumes that integrates plastic responses from functional-structural plant models. A generic model was developed to account for herbaceous legume species with contrasting above- and below-ground morphogenetic syndromes and to integrate the responses of plants to light, water and N. Through coupling with a radiative transfer model and a three-dimensional virtual soil, the model allows dynamic resolution of competition for multiple resources at individual plant level within a plant community. The behaviour of the model was assessed on a range of monospecific stands grown along gradients of light, water and N availability. The model proved able to capture the diversity of morphologies encountered among the forage legumes. The main density-dependent features known about even-age plant populations were correctly anticipated. The model predicted (1) the 'reciprocal yield' law relating average plant mass to density, (2) a self-thinning pattern close to that measured for herbaceous species and (3) consistent changes in the size structure of plant populations with time and pedo-climatic conditions. In addition, plastic changes in the partitioning of dry matter, the N acquisition mode and in the architecture of shoots and roots emerged from the integration of plant responses to their local environment. This resulted in taller plants and thinner roots when competition was dominated by light, and shorter plants with relatively more developed root systems when competition was dominated by soil resources. A population dynamic model considering growth and morphogenesis responses to multiple resources heterogeneously distributed in the environment was presented. It should allow scaling plant-plant interactions from individual to community levels without the inconvenience of average plant models.

  13. Contributions of Caucasian-associated bone mass loci to the variation in bone mineral density in Vietnamese population.

    PubMed

    Ho-Pham, Lan T; Nguyen, Sing C; Tran, Bich; Nguyen, Tuan V

    2015-07-01

    Bone mineral density (BMD) is under strong genetic regulation, but it is not clear which genes are involved in the regulation, particularly in Asian populations. This study sought to determine the association between 29 genes discovered by Caucasian-based genome-wide association studies and BMD in a Vietnamese population. The study involved 564 Vietnamese men and women aged 18 years and over (average age: 47 years) who were randomly sampled from the Ho Chi Minh City. BMD at the femoral neck, lumbar spine, total hip and whole body was measured by DXA (Hologic QDR4500, Bedford, MA, USA). Thirty-two single nucleotide polymorphisms (SNPs) in 29 genes were genotyped using Sequenom MassARRAY technology. The magnitude of association between SNPs and BMD was analyzed by the linear regression model. The Bayesian model average method was used to identify SNPs that are independently associated with BMD. The distribution of genotypes of all, but two, SNPs was consistent with the Hardy-Weinberg equilibrium law. After adjusting for age, gender and weight, 3 SNPs were associated with BMD: rs2016266 (SP7 gene), rs7543680 (ZBTB40 gene), and rs1373004 (MBL2/DKK1 gene). Among the three genetic variants, the SNP rs2016266 had the strongest association, with each minor allele being associated with ~0.02 g/cm(2) increase in BMD at the femoral neck and whole body. Each of these genetic variant explained about 0.2 to 1.1% variance of BMD. All other SNPs were not significantly associated with BMD. These results suggest that genetic variants in the SP7, ZBTB40 and MBL2/DKK1 genes are associated with BMD in the Vietnamese population, and that the effect of these genes on BMD is likely to be modest. Copyright © 2015 Elsevier Inc. All rights reserved.

  14. Microbial diversity in a bagasse-based compost prepared for the production of Agaricus brasiliensis

    PubMed Central

    Silva, Cristina Ferreira; Azevedo, Raquel Santos; Braga, Claudia; da Silva, Romildo; Dias, Eustáquio Souza; Schwan, Rosane Freitas

    2009-01-01

    Edible mushrooms are renowned for their nutritional and medicinal properties and are thus of considerable commercial importance. Mushroom production depends on the chemical composition of the basic substrates and additional supplements employed in the compost as well as on the method of composting. In order to minimise the cost of mushroom production, considerable interest has been shown in the use of agro-industrial residues in the preparation of alternative compost mixtures. However, the interaction of the natural microbiota present in agricultural residues during the composting process greatly influences the subsequent colonisation by the mushroom. The aim of the present study was to isolate and identify the microbiota present in a sugar cane bagasse and coast-cross straw compost prepared for the production of Agaricus brasilienses. Composting lasted for 14 days, during which time the substrates and additives were mixed every 2 days, and this was followed by a two-step steam pasteurisation (55 - 65°C; 15 h each step). Bacteria, (mainly Bacillus and Paenibacillus spp. and members of the Enterobacteriaceae) were the predominant micro-organisms present throughout the composting process with an average population density of 3 x 108 CFU/g. Actinomycetes, and especially members of the genus Streptomyces, were well represented with a population density of 2 - 3 x 108 CFU/g. The filamentous fungi, however, exhibited much lower population densities and were less diverse than the other micro-organisms, although Aspergillus fumigatus was present during the whole composting process and after pasteurisation. PMID:24031404

  15. Assessment of Density Variations of Marine Sediments with Ocean and Sediment Depths

    PubMed Central

    Tenzer, R.; Gladkikh, V.

    2014-01-01

    We analyze the density distribution of marine sediments using density samples taken from 716 drill sites of the Deep Sea Drilling Project (DSDP). The samples taken within the upper stratigraphic layer exhibit a prevailing trend of the decreasing density with the increasing ocean depth (at a rate of −0.05 g/cm3 per 1 km). Our results confirm findings of published studies that the density nonlinearly increases with the increasing sediment depth due to compaction. We further establish a 3D density model of marine sediments and propose theoretical models of the ocean-sediment and sediment-bedrock density contrasts. The sediment density-depth equation approximates density samples with an average uncertainty of about 10% and better represents the density distribution especially at deeper sections of basin sediments than a uniform density model. The analysis of DSDP density data also reveals that the average density of marine sediments is 1.70 g/cm3 and the average density of the ocean bedrock is 2.9 g/cm3. PMID:24744686

  16. Forest cover influences dispersal distance of white-tailed deer

    USGS Publications Warehouse

    Long, E.S.; Diefenbach, D.R.; Rosenberry, C.S.; Wallingford, B.D.; Grund, M.D.

    2005-01-01

    Animal dispersal patterns influence gene flow, disease spread, population dynamics, spread of invasive species, and establishment of rare or endangered species. Although differences in dispersal distances among taxa have been reported, few studies have described plasticity of dispersal distance among populations of a single species. In 2002-2003, we radiomarked 308 juvenile (7- to 10-month-old), male white-tailed deer (Odocoileus virginianus) in 2 study areas in Pennsylvania. By using a meta-analysis approach, we compared dispersal rates and distances from these populations together with published reports of 10 other nonmigratory populations of white-tailed deer. Population density did not influence dispersal rate or dispersal distance, nor did forest cover influence dispersal rate. However, average (r2 = 0.94, P < 0.001, d.f. = 9) and maximum (r2 = 0.86, P = 0.001, d.f. = 7) dispersal distances of juvenile male deer were greater in habitats with less forest cover. Hence, dispersal behavior of this habitat generalist varies, and use of landscape data to predict population-specific dispersal distances may aid efforts to model population spread, gene flow, or disease transmission. ?? 2005 American Society of Mammalogists.

  17. Spatial capture-recapture models for search-encounter data

    USGS Publications Warehouse

    Royle, J. Andrew; Kery, Marc; Guelat, Jerome

    2011-01-01

    1. Spatial capture–recapture models make use of auxiliary data on capture location to provide density estimates for animal populations. Previously, models have been developed primarily for fixed trap arrays which define the observable locations of individuals by a set of discrete points. 2. Here, we develop a class of models for 'search-encounter' data, i.e. for detections of recognizable individuals in continuous space, not restricted to trap locations. In our hierarchical model, detection probability is related to the average distance between individual location and the survey path. The locations are allowed to change over time owing to movements of individuals, and individual locations are related formally by a model describing individual activity or home range centre which is itself regarded as a latent variable in the model. We provide a Bayesian analysis of the model in WinBUGS, and develop a custom MCMC algorithm in the R language. 3. The model is applied to simulated data and to territory mapping data for the Willow Tit from the Swiss Breeding Bird Survey MHB. While the observed density was 15 territories per nominal 1 km2 plot of unknown effective sample area, the model produced a density estimate of 21∙12 territories per square km (95% posterior interval: 17–26). 4. Spatial capture–recapture models are relevant to virtually all animal population studies that seek to estimate population size or density, yet existing models have been proposed mainly for conventional sampling using arrays of traps. Our model for search-encounter data, where the spatial pattern of searching can be arbitrary and may change over occasions, greatly expands the scope and utility of spatial capture–recapture models.

  18. Profound Effects of Population Density on Fitness-Related Traits in an Invasive Freshwater Snail

    PubMed Central

    Zachar, Nicholas; Neiman, Maurine

    2013-01-01

    Population density can profoundly influence fitness-related traits and population dynamics, and density dependence plays a key role in many prominent ecological and evolutionary hypotheses. Here, we evaluated how individual-level changes in population density affect growth rate and embryo production early in reproductive maturity in two different asexual lineages of Potamopyrgus antipodarum, a New Zealand freshwater snail that is an important model system for ecotoxicology and the evolution of sexual reproduction as well as a potentially destructive worldwide invader. We showed that population density had a major influence on individual growth rate and early-maturity embryo production, effects that were often apparent even when comparing treatments that differed in population density by only one individual. While individual growth rate generally decreased as population density increased, we detected a hump-shaped relationship between embryo production and density, with females from intermediate-density treatments producing the most embryos and females from low- and high-density treatments producing the fewest embryos. The two lineages responded similarly to the treatments, indicating that these effects of population density might apply more broadly across P. antipodarum. These results indicate that there are profound and complex relationships between population density, growth rate, and early-maturity embryo production in at least two lineages of this important model system, with potential implications for the study of invasive populations, research on the maintenance of sex, and approaches used in ecotoxicology. PMID:24278240

  19. Profound effects of population density on fitness-related traits in an invasive freshwater snail.

    PubMed

    Zachar, Nicholas; Neiman, Maurine

    2013-01-01

    Population density can profoundly influence fitness-related traits and population dynamics, and density dependence plays a key role in many prominent ecological and evolutionary hypotheses. Here, we evaluated how individual-level changes in population density affect growth rate and embryo production early in reproductive maturity in two different asexual lineages of Potamopyrgus antipodarum, a New Zealand freshwater snail that is an important model system for ecotoxicology and the evolution of sexual reproduction as well as a potentially destructive worldwide invader. We showed that population density had a major influence on individual growth rate and early-maturity embryo production, effects that were often apparent even when comparing treatments that differed in population density by only one individual. While individual growth rate generally decreased as population density increased, we detected a hump-shaped relationship between embryo production and density, with females from intermediate-density treatments producing the most embryos and females from low- and high-density treatments producing the fewest embryos. The two lineages responded similarly to the treatments, indicating that these effects of population density might apply more broadly across P. antipodarum. These results indicate that there are profound and complex relationships between population density, growth rate, and early-maturity embryo production in at least two lineages of this important model system, with potential implications for the study of invasive populations, research on the maintenance of sex, and approaches used in ecotoxicology.

  20. The Importance of Shorebirds to Energy Fluxes in a Food Web of a South European Estuary

    NASA Astrophysics Data System (ADS)

    Moreira, Francisco

    1997-01-01

    This paper presents the results of a study on the diets and energetic needs of shorebird populations in the Tagus Estuary (Portugal). The average biomass removed by the bird populations during 1 year was estimated as 1755 kg ash-free dry weight day -1, equivalent to a predation pressure of 103 kJ m -2year -1. Gulls ingested more than half of this total, with waders and ducks ingesting 15-20% each. Cormorants, egrets and flamingos each took less than 7% of the total ingested biomass, thus constituting secondary groups for energy transfers in the food web. Assuming an invertebrate production estimate of 853 kJ m -2year -1, the consumption efficiency of birds on the invertebrate populations was 12%. Twelve bird species took 90% of the total biomass ingested during winter by the bird populations, each specializing on one or two preferred prey species that constituted more than 75% of their ingested biomass. During winter, the Tagus bird community is characterized by its great dependence on one prey species, the bivalve Scrobicularia plana. This prey, taken either in the form of siphons or intact individuals, represented a significant percentage of the biomass taken by the bird populations (45%), and it is important for three reasons. Firstly, other bivalves taken by birds in North European estuaries are absent or occur in low densities in the Tagus. Secondly, Scrobiculariadensities in the Tagus are high compared with values observed further north. Thirdly, although high densities also occur in Northern Europe, these density peaks are occasional as there is a trend for the occurrence of irregular recruitment, whereas further south, there is regular recruitment. Gulls and waders took significant percentages of bivalve siphons. This is an interesting finding, as the predation of siphons is a form of grazing with important ecological implications for energy fluxes across the estuarine ecosystem and for the prey population.

  1. Importance of regional variation in conservation planning: A rangewide example of the Greater Sage-Grouse

    USGS Publications Warehouse

    Doherty, Kevin E.; Evans, Jeffrey S.; Coates, Peter S.; Juliusson, Lara; Fedy, Bradley C.

    2016-01-01

    We developed rangewide population and habitat models for Greater Sage-Grouse (Centrocercus urophasianus) that account for regional variation in habitat selection and relative densities of birds for use in conservation planning and risk assessments. We developed a probabilistic model of occupied breeding habitat by statistically linking habitat characteristics within 4 miles of an occupied lek using a nonlinear machine learning technique (Random Forests). Habitat characteristics used were quantified in GIS and represent standard abiotic and biotic variables related to sage-grouse biology. Statistical model fit was high (mean correctly classified = 82.0%, range = 75.4–88.0%) as were cross-validation statistics (mean = 80.9%, range = 75.1–85.8%). We also developed a spatially explicit model to quantify the relative density of breeding birds across each Greater Sage-Grouse management zone. The models demonstrate distinct clustering of relative abundance of sage-grouse populations across all management zones. On average, approximately half of the breeding population is predicted to be within 10% of the occupied range. We also found that 80% of sage-grouse populations were contained in 25–34% of the occupied range within each management zone. Our rangewide population and habitat models account for regional variation in habitat selection and the relative densities of birds, and thus, they can serve as a consistent and common currency to assess how sage-grouse habitat and populations overlap with conservation actions or threats over the entire sage-grouse range. We also quantified differences in functional habitat responses and disturbance thresholds across the Western Association of Fish and Wildlife Agencies (WAFWA) management zones using statistical relationships identified during habitat modeling. Even for a species as specialized as Greater Sage-Grouse, our results show that ecological context matters in both the strength of habitat selection (i.e., functional response curves) and response to disturbance.

  2. Genetic variation facilitates seedling establishment but not population growth rate of a perennial invader

    PubMed Central

    Li, Shou-Li; Vasemägi, Anti; Ramula, Satu

    2016-01-01

    Background and Aims Assessing the demographic consequences of genetic variation is fundamental to invasion biology. However, genetic and demographic approaches are rarely combined to explore the effects of genetic variation on invasive populations in natural environments. This study combined population genetics, demographic data and a greenhouse experiment to investigate the consequences of genetic variation for the population fitness of the perennial, invasive herb Lupinus polyphyllus. Methods Genetic and demographic data were collected from 37 L. polyphyllus populations representing different latitudes in Finland, and genetic variation was characterized based on 13 microsatellite loci. Associations between genetic variation and population size, population density, latitude and habitat were investigated. Genetic variation was then explored in relation to four fitness components (establishment, survival, growth, fecundity) measured at the population level, and the long-term population growth rate (λ). For a subset of populations genetic variation was also examined in relation to the temporal variability of λ. A further assessment was made of the role of natural selection in the observed variation of certain fitness components among populations under greenhouse conditions. Key Results It was found that genetic variation correlated positively with population size, particularly at higher latitudes, and differed among habitat types. Average seedling establishment per population increased with genetic variation in the field, but not under greenhouse conditions. Quantitative genetic divergence (QST) based on seedling establishment in the greenhouse was smaller than allelic genetic divergence (F′ST), indicating that unifying selection has a prominent role in this fitness component. Genetic variation was not associated with average survival, growth or fecundity measured at the population level, λ or its variability. Conclusions The study suggests that although genetic variation may facilitate plant invasions by increasing seedling establishment, it may not necessarily affect the long-term population growth rate. Therefore, established invasions may be able to grow equally well regardless of their genetic diversity. PMID:26420202

  3. Neosho madtom and other ictalurid populations in relation to hydrologic characteristics of an impounded Midwestern warmwater stream: Update

    USGS Publications Warehouse

    Bryan, Janice L.; Wildhaber, Mark L.; William B. Leeds, William B.; Dey, Rima

    2010-01-01

    The Neosho madtom, Noturus placidus, is a small (less than 75 millimeters in total length) ictalurid that is native to the main stems of the Neosho and Cottonwood Rivers in Kansas and Oklahoma and the Spring River in Kansas and Missouri. The Neosho madtom was federally listed as threatened by the U.S. Fish and Wildlife Service in May 1990. The U.S. Fish and Wildlife Service has been monitoring Neosho madtoms since 1991, and questioned whether or not Neosho madtom densities were affected by other catfish species, reservoirs, and hydrologic characteristics. Using the first 8 years of U.S. Fish and Wildlife Service monitoring data, Wildhaber and others (2000) analyzed whether or not Neosho madtom densities were related to these environmental characteristics. The goal of this report is to update these results with data from 1999 to 2008. The trends of Neosho madtom densities in respect to John Redmond Reservoir and other catfish species remains consistent with the previous report. In both the Neosho and Spring Rivers, Neosho madtoms had a significant positive association with all catfish species. Of those species tested, only in the population of Neosho madtoms were significantly different in density above verses below the John Redmond Reservoir after accounting for the yearly variation. The average density of Neosho madtoms at the streamgage immediately below the reservoir had the second lowest density compared to the other streamgages. The positive associations with Neosho madtoms that remained consistent from the previous report included the 1-, 3-, and 7-day minima discharges and the annual minimum discharge from the previous water year (water year prior to when the fish were sampled) and the 1-, 3-, 7-, and 30-day minima discharges from the current water year (same water year fish were sampled).

  4. Comparing methods to estimate Reineke’s maximum size-density relationship species boundary line slope

    Treesearch

    Curtis L. VanderSchaaf; Harold E. Burkhart

    2010-01-01

    Maximum size-density relationships (MSDR) provide natural resource managers useful information about the relationship between tree density and average tree size. Obtaining a valid estimate of how maximum tree density changes as average tree size changes is necessary to accurately describe these relationships. This paper examines three methods to estimate the slope of...

  5. The density dilemma: limitations on juvenile production in threatened salmon populations

    USGS Publications Warehouse

    Walters, Annika W.; Copeland, Timothy; Venditti, David A.

    2013-01-01

    Density-dependent processes have repeatedly been shown to have a central role in salmonid population dynamics, but are often assumed to be negligible for populations at low abundances relative to historical records. Density dependence has been observed in overall spring/summer Snake River Chinook salmon Oncorhynchus tshawytscha production, but it is not clear how patterns observed at the aggregate level relate to individual populations within the basin. We used a Bayesian hierarchical modelling approach to explore the degree of density dependence in juvenile production for nine Idaho populations. Our results indicate that density dependence is ubiquitous, although its strength varies between populations. We also investigated the processes driving the population-level pattern and found density-dependent growth and mortality present for both common life-history strategies, but no evidence of density-dependent movement. Overwinter mortality, spatial clustering of redds and limited resource availability were identified as potentially important limiting factors contributing to density dependence. The ubiquity of density dependence for these threatened populations is alarming as stability at present low abundance levels suggests recovery may be difficult without major changes. We conclude that density dependence at the population level is common and must be considered in demographic analysis and management.

  6. How many tigers Panthera tigris are there in Huai Kha Khaeng Wildlife Sanctuary, Thailand? An estimate using photographic capture-recapture sampling

    USGS Publications Warehouse

    Simcharoen, S.; Pattanavibool, A.; Karanth, K.U.; Nichols, J.D.; Kumar, N.S.

    2007-01-01

    We used capture-recapture analyses to estimate the density of a tiger Panthera tigris population in the tropical forests of Huai Kha Khaeng Wildlife Sanctuary, Thailand, from photographic capture histories of 15 distinct individuals. The closure test results (z = 0.39, P = 0.65) provided some evidence in support of the demographic closure assumption. Fit of eight plausible closed models to the data indicated more support for model Mh, which incorporates individual heterogeneity in capture probabilities. This model generated an average capture probability $\\hat p$ = 0.42 and an abundance estimate of $\\widehat{N}(\\widehat{SE}[\\widehat{N}])$ = 19 (9.65) tigers. The sampled area of $\\widehat{A}(W)(\\widehat{SE}[\\widehat{A}(W)])$ = 477.2 (58.24) km2 yielded a density estimate of $\\widehat{D}(\\widehat{SE}[\\widehat{D}])$ = 3.98 (0.51) tigers per 100 km2. Huai Kha Khaeng Wildlife Sanctuary could therefore hold 113 tigers and the entire Western Forest Complex c. 720 tigers. Although based on field protocols that constrained us to use sub-optimal analyses, this estimated tiger density is comparable to tiger densities in Indian reserves that support moderate prey abundances. However, tiger densities in well-protected Indian reserves with high prey abundances are three times higher. If given adequate protection we believe that the Western Forest Complex of Thailand could potentially harbour >2,000 wild tigers, highlighting its importance for global tiger conservation. The monitoring approaches we recommend here would be useful for managing this tiger population.

  7. Relationships between human population density and burned area at continental and global scales.

    PubMed

    Bistinas, Ioannis; Oom, Duarte; Sá, Ana C L; Harrison, Sandy P; Prentice, I Colin; Pereira, José M C

    2013-01-01

    We explore the large spatial variation in the relationship between population density and burned area, using continental-scale Geographically Weighted Regression (GWR) based on 13 years of satellite-derived burned area maps from the global fire emissions database (GFED) and the human population density from the gridded population of the world (GPW 2005). Significant relationships are observed over 51.5% of the global land area, and the area affected varies from continent to continent: population density has a significant impact on fire over most of Asia and Africa but is important in explaining fire over < 22% of Europe and Australia. Increasing population density is associated with both increased and decreased in fire. The nature of the relationship depends on land-use: increasing population density is associated with increased burned are in rangelands but with decreased burned area in croplands. Overall, the relationship between population density and burned area is non-monotonic: burned area initially increases with population density and then decreases when population density exceeds a threshold. These thresholds vary regionally. Our study contributes to improved understanding of how human activities relate to burned area, and should contribute to a better estimate of atmospheric emissions from biomass burning.

  8. Relationships between Human Population Density and Burned Area at Continental and Global Scales

    PubMed Central

    Bistinas, Ioannis; Oom, Duarte; Sá, Ana C. L.; Harrison, Sandy P.; Prentice, I. Colin; Pereira, José M. C.

    2013-01-01

    We explore the large spatial variation in the relationship between population density and burned area, using continental-scale Geographically Weighted Regression (GWR) based on 13 years of satellite-derived burned area maps from the global fire emissions database (GFED) and the human population density from the gridded population of the world (GPW 2005). Significant relationships are observed over 51.5% of the global land area, and the area affected varies from continent to continent: population density has a significant impact on fire over most of Asia and Africa but is important in explaining fire over < 22% of Europe and Australia. Increasing population density is associated with both increased and decreased in fire. The nature of the relationship depends on land-use: increasing population density is associated with increased burned are in rangelands but with decreased burned area in croplands. Overall, the relationship between population density and burned area is non-monotonic: burned area initially increases with population density and then decreases when population density exceeds a threshold. These thresholds vary regionally. Our study contributes to improved understanding of how human activities relate to burned area, and should contribute to a better estimate of atmospheric emissions from biomass burning. PMID:24358108

  9. Waterfowl production on the Woodworth Station in south-central North Dakota, 1965-1981

    USGS Publications Warehouse

    Higgins, K.F.; Kirsch, L.M.; Klett, A.T.; Miller, H.W.

    1992-01-01

    During 17 years of study at the Woodworth, North Dakota study area, the percent of 548 wetland basin with water during 1-15 May ranged from 8 to 87 and averaged 56; waterfowl pair densities varied from 19 to 56/km2 and averaged 40/km2. Pond occupancy by duck pairs averaged 37% during mid-May counts and 48% for late May and early June counts. A positive linear relation occurred between the estimated number of duck pairs and the percent of basins with water during 1-15 May.There were 3,339 duck nests found in grassland habitats from 1966 through 1981. Approximately 66% (85% Mayfield) of these were depredated or abandoned. Mammals caused 88% of nest failures. Half or more of the eventually successful clutches were unhatched by 10 July in 9 of 16 years. Haying would have disturbed or destroyed an average of 43%, 33%, 22%, 15%, and 9% of the duck nests if initiated on 10 July, 15 July, 20 July, 25 July, and 1 August, respectively.The total average size of completed clutch for all species was 29% smaller at the end of the nesting season than at the beginning, underscoring the importance of protecting early clutches.Production averaged 30 broods per 100 pairs of ducks and ranged from 15 to 61 broods per 100 pairs. Brood densities ranged from 10 to 63/km2 and averaged 12/km2. Mean brood size averaged 6.4 for all species. July broods averaged 7.2 ducklings and August broods 5.7 ducklings. Duckling loss averaged 2.6 per brood and 85% (2.2 ducklings) of this loss was estimated to occur during the first 14 days after hatch.Wetlands of all sizes and classes were important at some time to one species of duck or another. With the exception of some diving ducks, all species used a complex of sizes and classes of wetlands for space, food, and shelter necessary for nesting and brooding. Pair counts during 20 May-7 June were most indicative of the breeding population. A combination of two brood counts resulted in the best estimate of annual production. An average of only 50% of the total duck broods per year was counted during the 1-15 July surveys, which approximated the average time of the Service's July aerial surveys. During this study the area produced an average of 1 duck per 4 ha of upland and had a nest density of approximately 1 nest per 14 ha. Nest success rates averaged 35.1% (16.3% Mayfield). Predation was significantly reduced by good vegetative cover at nest sites. Seeded grasslands (dense nesting cover) yielded better production than native prairie or croplands. Seeded grasslands also produced 3 times more ducklings per unit area than adjacent native prairie and more than 14 times as many as adjacent, annually tilled croplands.Ducks generally showed higher nest densities and better nesting success when using growing grain crops than when nesting in standing or mulched stubble fields. Among native mixed-grass prairie and seeded grassland, production was enhanced by leaving fields idle or by treating them with periodic burning. Duck production was generally lowered by grazing field of native prairie but duck production on grazing lands was higher than in annually tilled croplands.

  10. Probability of lek collapse is lower inside sage-grouse Core Areas: Effectiveness of conservation policy for a landscape species.

    PubMed

    Spence, Emma Suzuki; Beck, Jeffrey L; Gregory, Andrew J

    2017-01-01

    Greater sage-grouse (Centrocercus urophasianus) occupy sagebrush (Artemisia spp.) habitats in 11 western states and 2 Canadian provinces. In September 2015, the U.S. Fish and Wildlife Service announced the listing status for sage-grouse had changed from warranted but precluded to not warranted. The primary reason cited for this change of status was that the enactment of new regulatory mechanisms was sufficient to protect sage-grouse populations. One such plan is the 2008, Wyoming Sage Grouse Executive Order (SGEO), enacted by Governor Freudenthal. The SGEO identifies "Core Areas" that are to be protected by keeping them relatively free from further energy development and limiting other forms of anthropogenic disturbances near active sage-grouse leks. Using the Wyoming Game and Fish Department's sage-grouse lek count database and the Wyoming Oil and Gas Conservation Commission database of oil and gas well locations, we investigated the effectiveness of Wyoming's Core Areas, specifically: 1) how well Core Areas encompass the distribution of sage-grouse in Wyoming, 2) whether Core Area leks have a reduced probability of lek collapse, and 3) what, if any, edge effects intensification of oil and gas development adjacent to Core Areas may be having on Core Area populations. Core Areas contained 77% of male sage-grouse attending leks and 64% of active leks. Using Bayesian binomial probability analysis, we found an average 10.9% probability of lek collapse in Core Areas and an average 20.4% probability of lek collapse outside Core Areas. Using linear regression, we found development density outside Core Areas was related to the probability of lek collapse inside Core Areas. Specifically, probability of collapse among leks >4.83 km from inside Core Area boundaries was significantly related to well density within 1.61 km (1-mi) and 4.83 km (3-mi) outside of Core Area boundaries. Collectively, these data suggest that the Wyoming Core Area Strategy has benefited sage-grouse and sage-grouse habitat conservation; however, additional guidelines limiting development densities adjacent to Core Areas may be necessary to effectively protect Core Area populations.

  11. A high-density consensus map of barley linking DArT markers to SSR, RFLP and STS loci and agricultural traits

    PubMed Central

    Wenzl, Peter; Li, Haobing; Carling, Jason; Zhou, Meixue; Raman, Harsh; Paul, Edie; Hearnden, Phillippa; Maier, Christina; Xia, Ling; Caig, Vanessa; Ovesná, Jaroslava; Cakir, Mehmet; Poulsen, David; Wang, Junping; Raman, Rosy; Smith, Kevin P; Muehlbauer, Gary J; Chalmers, Ken J; Kleinhofs, Andris; Huttner, Eric; Kilian, Andrzej

    2006-01-01

    Background Molecular marker technologies are undergoing a transition from largely serial assays measuring DNA fragment sizes to hybridization-based technologies with high multiplexing levels. Diversity Arrays Technology (DArT) is a hybridization-based technology that is increasingly being adopted by barley researchers. There is a need to integrate the information generated by DArT with previous data produced with gel-based marker technologies. The goal of this study was to build a high-density consensus linkage map from the combined datasets of ten populations, most of which were simultaneously typed with DArT and Simple Sequence Repeat (SSR), Restriction Enzyme Fragment Polymorphism (RFLP) and/or Sequence Tagged Site (STS) markers. Results The consensus map, built using a combination of JoinMap 3.0 software and several purpose-built perl scripts, comprised 2,935 loci (2,085 DArT, 850 other loci) and spanned 1,161 cM. It contained a total of 1,629 'bins' (unique loci), with an average inter-bin distance of 0.7 ± 1.0 cM (median = 0.3 cM). More than 98% of the map could be covered with a single DArT assay. The arrangement of loci was very similar to, and almost as optimal as, the arrangement of loci in component maps built for individual populations. The locus order of a synthetic map derived from merging the component maps without considering the segregation data was only slightly inferior. The distribution of loci along chromosomes indicated centromeric suppression of recombination in all chromosomes except 5H. DArT markers appeared to have a moderate tendency toward hypomethylated, gene-rich regions in distal chromosome areas. On the average, 14 ± 9 DArT loci were identified within 5 cM on either side of SSR, RFLP or STS loci previously identified as linked to agricultural traits. Conclusion Our barley consensus map provides a framework for transferring genetic information between different marker systems and for deploying DArT markers in molecular breeding schemes. The study also highlights the need for improved software for building consensus maps from high-density segregation data of multiple populations. PMID:16904008

  12. Air quality of Beijing (China) and Delhi (India) and impact on Human Health and Climate in Asia

    NASA Astrophysics Data System (ADS)

    Zheng, S.; Singh, R. P.; Wu, Y.; Wu, C.

    2015-12-01

    Air pollution has been estimated to represent a significant fraction of the total mortality attributable to 26 risk factors assessed by the World Health Organization global burden of disease project. Delhi is distributed over 1484 km2 with population density of 11297/km2 (as in 2011) and surrounded by highly industrialized National Capital region (NCR) with population density of 1050/km2. Beijing covers an area of 16,800 km2, with population density of 1300/km2 (upto 2014). It is located at the foothills of Yan Mountains and Taihang Mountains, in the North China Plain. Both these cities suffer with poor air quality and are severely affected by dense haze, fog and smog during summer and winter seasons. Earlier studies in developing countries have concentrated on limited air quality parameters. Detailed results from trace gases (O3, NO, NO2, and CO) and particulate matter (PM10 and PM2.5) in two Asian megacities, Delhi (India) and Beijing (China), will be presented. Trace gases and particulate matter in Beijing were collected at 31 sites during 2013-2014. The measurements in Delhi were carried out at 8 sites during October 2010 - March 2013. The annual average of PM10, PM2.5, O3, NO, NO2, and CO over Delhi in 2013 is 199 ug/m3, 123 ug/m3, 25.6 ppb, 21.5 ppb, 15.8 ppb, 1.7 ppb, respectively. The annual average of PM10, PM2.5, O3, NO2, CO, and SO2 over Beijing is 113 ug/m3, 85 ug/m3, 51 ug/m3, 46 ug/m3, 1.3 mg/m3, 23 ug/m3, respectively. The annual and seasonal variations of trace gases and particulate matter in Beijing and Delhi are also analyzed, as well as spatial changes of air pollution in these two cities. A comparative analysis in these two cities and the sources of pollution and their impact on human health and Asian climate will be discussed.

  13. Nutrient loads within the Sava River Catchment and comparison with load relations in the Baltic region

    NASA Astrophysics Data System (ADS)

    Levi, Lea; Cvetkovic, Vladimir; Destouni, Georgia

    2015-04-01

    This study compiles estimates of total nitrogen and phosphorus loads in the Sava River Catchment (SRC), investigates the load relations to human drivers of excess nutrient loading, and compares them with corresponding relations implied by data reported for the Baltic region. Nutrient load data, associated average discharge concentrations (ratio of load to water discharge) and their relations to human drivers are investigated across subcatchments of the SRC with different agricultural and population conditions. The Zagreb subcatchment, which has the smallest area but the highest population density and runoff among the investigated SRC subcatchments, exhibits the highest loads of both nitrogen and phosphorus. Overall for the SRC, results show high correlation (R2=0.93-0.95) of nutrient loads with population density and of concentrations with farmland share. A further question investigated here is then to what degree these relations are comparable with such relations found also for the Baltic region. The two regions are otherwise quite different in their climatic, agricultural and wastewater treatment conditions, so relation consistency, even if surprising, would be important in indicating some degree of relation transferability worthy of further investigation also in other regions. For the Baltic region corresponding correlations to those found in the SRC are in the range R2=0.79-0.88. In particular nitrogen and phosphorus concentration correlations with farmland share are qualitatively consistent between the regions. At the same time, phosphorus concentration correlation with population density shows quite different results between regions. Obtained results indicate a certain level of transferability of dependencies between the two regions and call for further detailed investigations on finer spatial-temporal scales.

  14. [Ecological Trendofthe Incidence of Tuberculosis in Mianyang City During 2004-2013].

    PubMed

    Zhang, Wen-Hao; Xiao, Chuan; Ren, Tao; Wang, Li-Ping; Wang, Lan; Yuan, Ping

    2016-09-01

    To determine the trend of the incidence of tuberculosis (TB) in Mianyang City during 2004-2013 and its ecological determinants. Linear correlations between TB incidence and ecological factors were analyzed using the data collected in Mianyang City from 2004 to 2013. A multivariate linear regression model was established to determine the ecological predictors of TB incidence. The incidence of TB in Mianyang City decreased over the period of 2004-2013. Economic development and increased health resources were negatively correlated with TB incidence. Population density was positively correlated with TB incidence. A multivariate linear regression equationon TB incidence ( y ) was established with the independent variables ( x₁ to x ₁₀) forming a component (using principal component analysis) to eliminate multicollinearity: y =117.692-1.467 x ₁-1.145 x ₂-1.961 x ₃-4.777 x ₄-2.690 x ₅-6.181 x ₆+82.234 x ₇-2.721 x ₈-0.351 x ₉-0.382 x ₁₀. The incidence of TB decreased with the increases of real GDP per capita ( x ₁), average wage of workers( x ₂), per capita disposable income of urban residents ( x ₃), rural per capita net income ( x ₄), per capita consumption expenditure of urban residents ( x ₅), per capita living consumption expenditure of rural residents ( x ₆), number of licensed (assistant) physicians per thousand population ( x ₈), urbanization rate ( x ₉),and per capita housing construction area of urban ( x ₁₀),while it increased with the increase of density of population ( x ₇). Socio-economic development, health resources and population density are predictors of TB incidence.

  15. Genetic relatedness of previously Plant-Variety-Protected commercial maize inbreds.

    PubMed

    Beckett, Travis J; Morales, A Jason; Koehler, Klaus L; Rocheford, Torbert R

    2017-01-01

    The emergence of high-throughput, high-density genotyping methods combined with increasingly powerful computing systems has created opportunities to further discover and exploit the genes controlling agronomic performance in elite maize breeding populations. Understanding the genetic basis of population structure in an elite set of materials is an essential step in this genetic discovery process. This paper presents a genotype-based population analysis of all maize inbreds whose Plant Variety Protection certificates had expired as of the end of 2013 (283 inbreds) as well as 66 public founder inbreds. The results provide accurate population structure information and allow for important inferences in context of the historical development of North American elite commercial maize germplasm. Genotypic data was obtained via genotyping-by-sequencing on 349 inbreds. After filtering for missing data, 77,314 high-quality markers remained. The remaining missing data (average per individual was 6.22 percent) was fully imputed at an accuracy of 83 percent. Calculation of linkage disequilibrium revealed that the average r2 of 0.20 occurs at approximately 1.1 Kb. Results of population genetics analyses agree with previously published studies that divide North American maize germplasm into three heterotic groups: Stiff Stalk, Non-Stiff Stalk, and Iodent. Principal component analysis shows that population differentiation is indeed very complex and present at many levels, yet confirms that division into three main sub-groups is optimal for population description. Clustering based on Nei's genetic distance provides an additional empirical representation of the three main heterotic groups. Overall fixation index (FST), indicating the degree of genetic divergence between the three main heterotic groups, was 0.1361. Understanding the genetic relationships and population differentiation of elite germplasm may help breeders to maintain and potentially increase the rate of genetic gain, resulting in higher overall agronomic performance.

  16. Is alloparenting helpful for Mednyi Island arctic foxes, Alopex lagopus semenovi?

    PubMed

    Kruchenkova, Elena P; Goltsman, Michael; Sergeev, Sergei; Macdonald, David W

    2009-04-01

    The Arctic Fox Alopex lagopus semenovi population on Mednyi Island is completely isolated and subsists largely by scavenging on seabird colonies, which have remained abundant and spatio-temporally predictable for many years. We compared population data at the beginning of 1976/1978 and some time after 1994-2005, finding an 85% decline in fox numbers due to disease, to assess the effect of population size on social structure. A total of 81 groups of known size and composition was observed during this 29-year period. Overall, helpers (usually non-lactating yearling females) occurred in 25.7% of groups, and in 32.4% of groups there were two or three lactating females. Female engagement in alloparental behaviour decreased, but not statistically significantly, after the decline in population density. Total food availability was apparently constant throughout the study period, and therefore, the amount available per individual was much higher later in the study. Both communally nursing females and helpers brought food and helped to guard the litter. However, the benefits of communal rearing were unclear. While cubs were left without guards significantly more rarely in the groups with an additional adult, the number of cubs weaned per lactating female was greater in groups with one (3.93 +/- 1.60), as opposed to two or three (3.06 +/- 0.92), lactating females. Survival of cubs to 1 year of age in the groups with two lactating females and/or with helpers was lower than that in the families with one lactating female without helpers (22.2% vs 32.2%). Fewer second-generation litters were born to foxes produced by composite families than to those produced by pairs. Reproductive adults producing by pairs had, on average, 1.23 (+/-1.72) second-generation litters. In groups that initially included additional adults, the average number of second-generation litters per reproductive female was 0.21 (+/-0.49) and 0.46 (+/-0.81) litters per male. Thus, according to three measures, increased group size had no apparent positive impact on reproductive success. The increased parental investment and enhanced guarding of the cubs in the larger families could be beneficial under conditions of high population density and a saturated biotope to which the island fox population was presumably adapted before the population crash in the late 1970s.

  17. Is alloparenting helpful for Mednyi Island arctic foxes, Alopex lagopus semenovi?

    NASA Astrophysics Data System (ADS)

    Kruchenkova, Elena P.; Goltsman, Michael; Sergeev, Sergei; MacDonald, David W.

    2009-04-01

    The Arctic Fox Alopex lagopus semenovi population on Mednyi Island is completely isolated and subsists largely by scavenging on seabird colonies, which have remained abundant and spatio-temporally predictable for many years. We compared population data at the beginning of 1976/1978 and some time after 1994-2005, finding an 85% decline in fox numbers due to disease, to assess the effect of population size on social structure. A total of 81 groups of known size and composition was observed during this 29-year period. Overall, helpers (usually non-lactating yearling females) occurred in 25.7% of groups, and in 32.4% of groups there were two or three lactating females. Female engagement in alloparental behaviour decreased, but not statistically significantly, after the decline in population density. Total food availability was apparently constant throughout the study period, and therefore, the amount available per individual was much higher later in the study. Both communally nursing females and helpers brought food and helped to guard the litter. However, the benefits of communal rearing were unclear. While cubs were left without guards significantly more rarely in the groups with an additional adult, the number of cubs weaned per lactating female was greater in groups with one (3.93 ± 1.60), as opposed to two or three (3.06 ± 0.92), lactating females. Survival of cubs to 1 year of age in the groups with two lactating females and/or with helpers was lower than that in the families with one lactating female without helpers (22.2% vs 32.2%). Fewer second-generation litters were born to foxes produced by composite families than to those produced by pairs. Reproductive adults producing by pairs had, on average, 1.23 (±1.72) second-generation litters. In groups that initially included additional adults, the average number of second-generation litters per reproductive female was 0.21 (±0.49) and 0.46 (±0.81) litters per male. Thus, according to three measures, increased group size had no apparent positive impact on reproductive success. The increased parental investment and enhanced guarding of the cubs in the larger families could be beneficial under conditions of high population density and a saturated biotope to which the island fox population was presumably adapted before the population crash in the late 1970s.

  18. Impact crater densities on volcanoes and coronae on venus: implications for volcanic resurfacing.

    PubMed

    Namiki, N; Solomon, S C

    1994-08-12

    The density of impact craters on large volcanoes on Venus is half the average crater density for the planet. The crater density on some classes of coronae is not significantly different from the global average density, but coronae with extensive associated volcanic deposits have lower crater densities. These results are inconsistent with both single-age and steady-state models for global resurfacing and suggest that volcanoes and coronae with associated volcanism have been active on Venus over the last 500 million years.

  19. Noise-driven bias in the non-local voter model

    NASA Astrophysics Data System (ADS)

    Minors, Kevin; Rogers, Tim; Yates, Christian A.

    2018-04-01

    Is it more effective to have a strong influence over a small domain, or a weaker influence over a larger one? Here, we introduce and analyse an off-lattice generalisation of the voter model, in which the range and strength of agents' influence are control parameters. We consider both low- and high-density regimes and, using distinct mathematical approaches, derive analytical predictions for the evolution of agent densities. We find that, even when the agents are equally persuasive on average, those whose influence is wider but weaker have an overall noise-driven advantage allowing them to reliably dominate the entire population. We discuss the implications of our results and the potential of our model (or adaptations thereof) to improve the understanding of political campaign strategies and the evolution of disease.

  20. Validity criteria for Fermi’s golden rule scattering rates applied to metallic nanowires

    NASA Astrophysics Data System (ADS)

    Moors, Kristof; Sorée, Bart; Magnus, Wim

    2016-09-01

    Fermi’s golden rule underpins the investigation of mobile carriers propagating through various solids, being a standard tool to calculate their scattering rates. As such, it provides a perturbative estimate under the implicit assumption that the effect of the interaction Hamiltonian which causes the scattering events is sufficiently small. To check the validity of this assumption, we present a general framework to derive simple validity criteria in order to assess whether the scattering rates can be trusted for the system under consideration, given its statistical properties such as average size, electron density, impurity density et cetera. We derive concrete validity criteria for metallic nanowires with conduction electrons populating a single parabolic band subjected to different elastic scattering mechanisms: impurities, grain boundaries and surface roughness.

  1. Determinant of Household Business Scale of Moa Buffaloes at Moa Island Southwest Maluku Regency

    NASA Astrophysics Data System (ADS)

    Lainsamputty, J.; Roessali, W.; Santosa, S. I.; Eddy, B. T.

    2018-02-01

    The objective of this research was to analyze factors that affect the business scale of Moa buffaloes at Moa Island, Regency of Southwest Maluku. The research used a survey method with multistage random sampling. The location chosen was the District of Moa Lakor at Moa Island based on its largest buffalo population. Respondents were randomly drawn in a total of 120 respondents. The variables measured were years of experience in rearing animals, costs of production, farmer’s participation in group activities, animal housing systems, farmer’s income and farmer’s age. The statistical test used was the multiple linear regressions. The results showed that the mean of business scale in the area of high density of buffaloes population was 12.6 AU, in the moderate was 12.4 AU and in the low was 11.0 AU. The average of production cost was IDR 1.893.536.00/year, the average of revenue was IDR14.083.333.00/year, while the average of income was IDR 12.189.797.00/year. The independent variables simultaneously influence the business scale (P<0.01). Partially, experience, costs of production, participation and housing systems had highly significant influences on the business scale (P<0.01).

  2. Oblate-Earth Effects on the Calculation of Ec During Spacecraft Reentry

    NASA Technical Reports Server (NTRS)

    Bacon, John B.; Matney, Mark

    2017-01-01

    The bulge in the Earth at its equator has been shown to lead to a clustering of natural decays biased to occur towards the equator and away from the orbit's extreme latitudes. Such clustering must be considered when predicting the Expectation of Casualty (Ec) during a natural decay, because of the corresponding clustering of the human population in the lower latitudes. This study expands upon prior work, and formalizes in a single empirical equation the correction that must be made to the calculation of the average exposed population density as a result of this effect. The equation is represented as a function of ballistic number and inclination of the entering spacecraft over the credible range of ballistic numbers.

  3. Population Density Modulates Drug Inhibition and Gives Rise to Potential Bistability of Treatment Outcomes for Bacterial Infections.

    PubMed

    Karslake, Jason; Maltas, Jeff; Brumm, Peter; Wood, Kevin B

    2016-10-01

    The inoculum effect (IE) is an increase in the minimum inhibitory concentration (MIC) of an antibiotic as a function of the initial size of a microbial population. The IE has been observed in a wide range of bacteria, implying that antibiotic efficacy may depend on population density. Such density dependence could have dramatic effects on bacterial population dynamics and potential treatment strategies, but explicit measures of per capita growth as a function of density are generally not available. Instead, the IE measures MIC as a function of initial population size, and population density changes by many orders of magnitude on the timescale of the experiment. Therefore, the functional relationship between population density and antibiotic inhibition is generally not known, leaving many questions about the impact of the IE on different treatment strategies unanswered. To address these questions, here we directly measured real-time per capita growth of Enterococcus faecalis populations exposed to antibiotic at fixed population densities using multiplexed computer-automated culture devices. We show that density-dependent growth inhibition is pervasive for commonly used antibiotics, with some drugs showing increased inhibition and others decreased inhibition at high densities. For several drugs, the density dependence is mediated by changes in extracellular pH, a community-level phenomenon not previously linked with the IE. Using a simple mathematical model, we demonstrate how this density dependence can modulate population dynamics in constant drug environments. Then, we illustrate how time-dependent dosing strategies can mitigate the negative effects of density-dependence. Finally, we show that these density effects lead to bistable treatment outcomes for a wide range of antibiotic concentrations in a pharmacological model of antibiotic treatment. As a result, infections exceeding a critical density often survive otherwise effective treatments.

  4. Population Density Modulates Drug Inhibition and Gives Rise to Potential Bistability of Treatment Outcomes for Bacterial Infections

    PubMed Central

    Maltas, Jeff; Brumm, Peter; Wood, Kevin B.

    2016-01-01

    The inoculum effect (IE) is an increase in the minimum inhibitory concentration (MIC) of an antibiotic as a function of the initial size of a microbial population. The IE has been observed in a wide range of bacteria, implying that antibiotic efficacy may depend on population density. Such density dependence could have dramatic effects on bacterial population dynamics and potential treatment strategies, but explicit measures of per capita growth as a function of density are generally not available. Instead, the IE measures MIC as a function of initial population size, and population density changes by many orders of magnitude on the timescale of the experiment. Therefore, the functional relationship between population density and antibiotic inhibition is generally not known, leaving many questions about the impact of the IE on different treatment strategies unanswered. To address these questions, here we directly measured real-time per capita growth of Enterococcus faecalis populations exposed to antibiotic at fixed population densities using multiplexed computer-automated culture devices. We show that density-dependent growth inhibition is pervasive for commonly used antibiotics, with some drugs showing increased inhibition and others decreased inhibition at high densities. For several drugs, the density dependence is mediated by changes in extracellular pH, a community-level phenomenon not previously linked with the IE. Using a simple mathematical model, we demonstrate how this density dependence can modulate population dynamics in constant drug environments. Then, we illustrate how time-dependent dosing strategies can mitigate the negative effects of density-dependence. Finally, we show that these density effects lead to bistable treatment outcomes for a wide range of antibiotic concentrations in a pharmacological model of antibiotic treatment. As a result, infections exceeding a critical density often survive otherwise effective treatments. PMID:27764095

  5. Spatial and Temporal Variation in Aedes aegypti and Aedes albopictus (Diptera: Culicidae) Numbers in the Yogyakarta Area of Java, Indonesia, With Implications for Wolbachia Releases.

    PubMed

    Tantowijoyo, W; Arguni, E; Johnson, P; Budiwati, N; Nurhayati, P I; Fitriana, I; Wardana, S; Ardiansyah, H; Turley, A P; Ryan, P; O'Neill, S L; Hoffmann, A A

    2016-01-01

    of mosquito vector populations, particularly through Wolbachia endosymbionts. The success of these strategies depends on understanding the dynamics of vector populations. In preparation for Wolbachia releases around Yogyakarta, we have studied Aedes populations in five hamlets. Adult monitoring with BioGent- Sentinel (BG-S) traps indicated that hamlet populations had different dynamics across the year; while there was an increase in Aedes aegypti (L.) and Aedes albopictus (Skuse) numbers in the wet season, species abundance remained relatively stable in some hamlets but changed markedly (>2 fold) in others. Local rainfall a month prior to monitoring partly predicted numbers of Ae. aegypti but not Ae. albopictus. Site differences in population size indicated by BG-S traps were also evident in ovitrap data. Egg or larval collections with ovitraps repeated at the same location suggested spatial autocorrelation (<250 m) in the areas of the hamlets where Ae. aegypti numbers were high. Overall, there was a weak negative association (r<0.43) between Ae. aegypti and Ae. albopictus numbers in ovitraps when averaged across collections. Ae. albopictus numbers in ovitraps and BG-S traps were positively correlated with vegetation around areas where traps were placed, while Ae. aegypti were negatively correlated with this feature. These data inform intervention strategies by defining periods when mosquito densities are high, highlighting the importance of local site characteristics on populations, and suggesting relatively weak interactions between Ae. aegypti and Ae. albopictus. They also indicate local areas within hamlets where consistently high mosquito densities may influence Wolbachia invasions and other interventions.

  6. The 6 "biggies".

    PubMed

    Myers, N

    1993-01-01

    The 6 developing nations with the largest populations in mid-1993 were China with 1.18 billion people, India 897 million, Indonesia 188 million, Brazil 152 million, Pakistan 122 million, and Bangladesh 114 million. Successful expansion of population planning programs in these nations could help defuse the population bomb. China's fertility rate has hardly declined for several years, and the year 2000 projection had to be revised from 1.2 billion to 1.29 billion people. India's family planning efforts have also stalled, and even if it attains the two-child family by the year 2015, population growth will not level out before the total reaches 1.9 billion. In India, Kerala State's population density matches that of Bangladesh, and per-capita income is among the lowest in the country. Yet, life expectancy is one quarter above the national average, infant mortality less than half, and literacy almost twice. And the fertility rate is down to 2.3, contrasting strongly with India's average of 3.9. Indonesia's population growth rate has plunged from 2.3% per year in the mid-1960s to 1.97% in the mid-1980s and to 1.7% today. Women now have an average of only three children. Brazil has achieved an annual population growth rate of only 1.5% and a fertility rate of 2.6, but fewer than 3 married women in 5 use modern contraception. There are an estimated 2 million illegal abortions a year. Part of the problem is the gulf between poverty and affluence. As a result of this gulf, child mortality remains the fourth highest in Latin America. Pakistan's population is projected to grow to 275 million people by the year 2025. The current growth rate is 3.1% per year. Family size is 6.7 children, the desired size is 4 children, and only 9% of married women use modern contraception. In Bangladesh, in 1975, the family size was 7 children. In 1993, it was fewer than 5. The spread of family-planning facilities boosted contraceptive use from 3% in 1971 to 40% in 1991.

  7. Microplastic contamination in Lake Winnipeg, Canada.

    PubMed

    Anderson, Philip J; Warrack, Sarah; Langen, Victoria; Challis, Jonathan K; Hanson, Mark L; Rennie, Michael D

    2017-06-01

    Microplastics are an emerging contaminant of concern in aquatic ecosystems. To better understand microplastic contamination in North American surface waters, we report for the first time densities of microplastics in Lake Winnipeg, the 11th largest freshwater body in the world. Samples taken 2014 to 2016 revealed similar or significantly greater microplastic densities in Lake Winnipeg compared with those reported in the Laurentian Great Lakes. Plastics in the lake were largely of secondary origin, overwhelmingly identified as fibres. We detected significantly greater densities of microplastics in the north basin compared to the south basin of the lake in 2014, but not in 2015 or 2016. Mean lake-wide densities across all years were comparable and not statistically different. Scanning electron microscopy with energy dispersive X-ray spectroscopy indicated that 23% of isolated particles on average were not plastic. While the ecological impact of microplastics on aquatic ecosystems is still largely unknown, our study contributes to the growing evidence that microplastic contamination is widespread even around sparsely-populated freshwater ecosystems, and provides a baseline for future study and risk assessments. Copyright © 2017 Elsevier Ltd. All rights reserved.

  8. Is aceticlastic methanogen composition in full-scale anaerobic processes related to acetate utilization capacity?

    PubMed

    Yilmaz, Vedat; Ince-Yilmaz, Ebru; Yilmazel, Yasemin Dilsad; Duran, Metin

    2014-06-01

    In this study, biomass samples were obtained from six municipal and nine industrial full-scale anaerobic processes to investigate whether the aceticlastic methanogen population composition is related to acetate utilization capacity and the nature of the wastewater treated, i.e. municipal sludge or industrial wastewater. Batch serum bottle tests were used to determine the specific acetate utilization rate (AUR), and a quantitative real-time polymerase chain reaction protocol was used to enumerate the acetate-utilizing Methanosaeta and Methanosarcina populations in the biomass samples. Methanosaeta was the dominant aceticlastic methanogen in all samples, except for one industrial wastewater-treating anaerobic process. However, Methanosarcina density in industrial biomass samples was higher than the Methanosarcina density in the municipal samples. The average AUR values of municipal and industrial wastewater treatment plant biomass samples were 10.49 and 10.65 mg CH3COO(-)/log(aceticlastic methanogen gene copy).d, respectively. One-way ANOVA test and principle component analysis showed that the acetate utilization capacities and aceticlastic methanogen community composition did not show statistically significant correlation among the municipal digesters and industrial wastewater-treating processes investigated.

  9. Intermittent burst of a super rogue wave in the breathing multi-soliton regime of an anomalous fiber ring cavity.

    PubMed

    Lee, Seungjong; Park, Kyoungyoon; Kim, Hyuntai; Vazquez-Zuniga, Luis Alonso; Kim, Jinseob; Jeong, Yoonchan

    2018-04-30

    We report the intermittent burst of a super rogue wave in the multi-soliton (MS) regime of an anomalous-dispersion fiber ring cavity. We exploit the spatio-temporal measurement technique to log and capture the shot-to-shot wave dynamics of various pulse events in the cavity, and obtain the corresponding intensity probability density function, which eventually unveils the inherent nature of the extreme events encompassed therein. In the breathing MS regime, a specific MS regime with heavy soliton population, the natural probability of pulse interaction among solitons and dispersive waves exponentially increases owing to the extraordinarily high soliton population density. Combination of the probabilistically started soliton interactions and subsequently accompanying dispersive waves in their vicinity triggers an avalanche of extreme events with even higher intensities, culminating to a burst of a super rogue wave nearly ten times stronger than the average solitons observed in the cavity. Without any cavity modification or control, the process naturally and intermittently recurs within a time scale in the order of ten seconds.

  10. Herd-level bovine tuberculosis risk factors: assessing the role of low-level badger population disturbance.

    PubMed

    Wright, David M; Reid, Neil; Ian Montgomery, W; Allen, Adrian R; Skuce, Robin A; Kao, Rowland R

    2015-08-17

    Bovine TB (bTB) is endemic in Irish cattle and has eluded eradication despite considerable expenditure, amid debate over the relative roles of badgers and cattle in disease transmission. Using a comprehensive dataset from Northern Ireland (>10,000 km(2); 29,513 cattle herds), we investigated interactions between host populations in one of the first large-scale risk factor analyses for new herd breakdowns to combine data on both species. Cattle risk factors (movements, international imports, bTB history, neighbours with bTB) were more strongly associated with herd risk than area-level measures of badger social group density, habitat suitability or persecution (sett disturbance). Highest risks were in areas of high badger social group density and high rates of persecution, potentially representing both responsive persecution of badgers in high cattle risk areas and effects of persecution on cattle bTB risk through badger social group disruption. Average badger persecution was associated with reduced cattle bTB risk (compared with high persecution areas), so persecution may contribute towards sustaining bTB hotspots; findings with important implications for existing and planned disease control programmes.

  11. Geospatial Analysis of Grey Wolf Movement Patterns

    NASA Astrophysics Data System (ADS)

    Sur, D.

    2017-12-01

    The grey wolf is a top predator that lives across a diverse habitat, ranging from Europe to North America. They often hunt in packs, preferring caribou, deer and elk as prey. Currently, many gray wolves live in Denali National Park and Preserve. In this study, several wolf packs were studied in three distinct regions of Denali. The purpose of my research was to investigate the links between wolf habitat, movement patterns, and prey thresholds. These are needed for projecting future population, growth and distribution of wolves in the studied region. I also investigated the effect wolves have on the ecological structure of the communities they inhabit. In the study I carried out a quantitative analysis of wolf population trends and daily distance movement by utilizing an analysis of variance (ANOVA) in the program JmpPro12 (SAS Institute, Crary, NC) to assess regional differences in pack size, wolf density, average daily distance moved. I found a clear link between the wolf habitat and prey thresholds; the habitat directly influences the types of prey available. However there was no link between the daily distance movement, the wolf habitat and prey density.

  12. Monitoring benthic foraminiferal dynamics at Bottsand coastal lagoon (western Baltic Sea)

    NASA Astrophysics Data System (ADS)

    Schönfeld, Joachim

    2018-04-01

    Benthic foraminifera from Bottsand coastal lagoon, western Baltic Sea, have been studied since the mid-1960s. They were monitored annually in late autumn since 2003 at the terminal ditch of the lagoon. There were 12 different species recognised, of which three have not been recorded during earlier investigations. Dominant species showed strong interannual fluctuations and a steady increase in population densities over the last decade. Elphidium incertum, a stenohaline species of the Baltic deep water fauna, colonised the Bottsand lagoon in 2016, most likely during a period of salinities >19 units and water temperatures of 18 °C on average in early autumn. The high salinities probably triggered their germination from a propagule bank in the ditch bottom sediment. The new E. incertum population showed densities higher by an order of magnitude than those of the indigenous species. The latter did not decline, revealing that E. incertum used another food source or occupied a different microhabitat. Elphidium incertum survived transient periods of lower salinities in late autumn 2017, though with reduced abundances, and became a regular faunal constituent at the Bottsand lagoon.

  13. Variation in the biomass density and demography of Antarctic krill in the vicinity of the South Shetland Islands during the 1999/2000 austral summer

    NASA Astrophysics Data System (ADS)

    Hewitt, R. P.; Kim, S.; Naganobu, M.; Gutierrez, M.; Kang, D.; Takao, Y.; Quinones, J.; Lee, Y.-H.; Shin, H.-C.; Kawaguchi, S.; Emery, J. H.; Demer, D. A.; Loeb, V. J.

    2004-06-01

    Vessels from Japan, Peru, and the USA conducted four sequential surveys designed to estimate the biomass density and demography of Antarctic krill in the vicinity of the South Shetland Islands between late December 1999 and early March 2000. The surveys were conducted during the same austral summer as the CCAMLR 2000 Survey in the Scotia Sea (Watkins et al., Deep-Sea Research, II, this issue [doi: 10.1016/j.dsr2.2004.06.010]), and the data were analyzed in a similar manner. Biomass densities were not significantly different between the surveys and averaged 49 g m -2. Maps of krill biomass indicate three areas of consistently high density: one near the eastern end of Elephant Island, one mid-way between Elephant Island and King George Island, and one near Cape Shirreff on the north side of Livingston Island. The areas of highest krill density appeared to move closer to the shelf break as the season progressed. This apparent movement was accompanied by a change in the demographic structure of the population, with smaller krill absent and a larger proportion of sexually mature animals present in late summer.

  14. Density dependence in demography and dispersal generates fluctuating invasion speeds

    PubMed Central

    Li, Bingtuan; Miller, Tom E. X.

    2017-01-01

    Density dependence plays an important role in population regulation and is known to generate temporal fluctuations in population density. However, the ways in which density dependence affects spatial population processes, such as species invasions, are less understood. Although classical ecological theory suggests that invasions should advance at a constant speed, empirical work is illuminating the highly variable nature of biological invasions, which often exhibit nonconstant spreading speeds, even in simple, controlled settings. Here, we explore endogenous density dependence as a mechanism for inducing variability in biological invasions with a set of population models that incorporate density dependence in demographic and dispersal parameters. We show that density dependence in demography at low population densities—i.e., an Allee effect—combined with spatiotemporal variability in population density behind the invasion front can produce fluctuations in spreading speed. The density fluctuations behind the front can arise from either overcompensatory population growth or density-dependent dispersal, both of which are common in nature. Our results show that simple rules can generate complex spread dynamics and highlight a source of variability in biological invasions that may aid in ecological forecasting. PMID:28442569

  15. Forecasting fish biomasses, densities, productions, and bioaccumulation potentials of mid-atlantic wadeable streams.

    PubMed

    Barber, M Craig; Rashleigh, Brenda; Cyterski, Michael J

    2016-01-01

    Regional fishery conditions of Mid-Atlantic wadeable streams in the eastern United States are estimated using the Bioaccumulation and Aquatic System Simulator (BASS) bioaccumulation and fish community model and data collected by the US Environmental Protection Agency's Environmental Monitoring and Assessment Program (EMAP). Average annual biomasses and population densities and annual productions are estimated for 352 randomly selected streams. Realized bioaccumulation factors (BAF) and biomagnification factors (BMF), which are dependent on these forecasted biomasses, population densities, and productions, are also estimated by assuming constant water exposures to methylmercury and tetra-, penta-, hexa-, and hepta-chlorinated biphenyls. Using observed biomasses, observed densities, and estimated annual productions of total fish from 3 regions assumed to support healthy fisheries as benchmarks (eastern Tennessee and Catskill Mountain trout streams and Ozark Mountains smallmouth bass streams), 58% of the region's wadeable streams are estimated to be in marginal or poor condition (i.e., not healthy). Using simulated BAFs and EMAP Hg fish concentrations, we also estimate that approximately 24% of the game fish and subsistence fishing species that are found in streams having detectable Hg concentrations would exceed an acceptable human consumption criterion of 0.185 μg/g wet wt. Importantly, such streams have been estimated to represent 78.2% to 84.4% of the Mid-Atlantic's wadeable stream lengths. Our results demonstrate how a dynamic simulation model can support regional assessment and trends analysis for fisheries. © 2015 SETAC.

  16. Spatial distributions of the red palm mite, Raoiella indica (Acari: Tenuipalpidae) on coconut and their implications for development of efficient sampling plans.

    PubMed

    Roda, A; Nachman, G; Hosein, F; Rodrigues, J C V; Peña, J E

    2012-08-01

    The red palm mite (Raoiella indica), an invasive pest of coconut, entered the Western hemisphere in 2004, then rapidly spread through the Caribbean and into Florida, USA. Developing effective sampling methods may aid in the timely detection of the pest in a new area. Studies were conducted to provide and compare intra tree spatial distribution of red palm mite populations on coconut in two different geographical areas, Trinidad and Puerto Rico, recently invaded by the mite. The middle stratum of a palm hosted significantly more mites than fronds from the upper or lower canopy and fronds from the lower stratum, on average, had significantly fewer mites than the two other strata. The mite populations did not vary within a frond. Mite densities on the top section of the pinna had significantly lower mite densities than the two other sections, which were not significantly different from each other. In order to improve future sampling plans for the red palm mite, the data was used to estimate the variance components associated with the various levels of the hierarchical sampling design. Additionally, presence-absence data were used to investigate the probability of no mites being present in a pinna section randomly chosen from a frond inhabited by mites at a certain density. Our results show that the most precise density estimate at the plantation level is to sample one pinna section per tree from as many trees as possible.

  17. Experimental test of an eco-evolutionary dynamic feedback loop between evolution and population density in the green peach aphid.

    PubMed

    Turcotte, Martin M; Reznick, David N; Daniel Hare, J

    2013-05-01

    An eco-evolutionary feedback loop is defined as the reciprocal impacts of ecology on evolutionary dynamics and evolution on ecological dynamics on contemporary timescales. We experimentally tested for an eco-evolutionary feedback loop in the green peach aphid, Myzus persicae, by manipulating initial densities and evolution. We found strong evidence that initial aphid density alters the rate and direction of evolution, as measured by changes in genotype frequencies through time. We also found that evolution of aphids within only 16 days, or approximately three generations, alters the rate of population growth and predicts density compared to nonevolving controls. The impact of evolution on population dynamics also depended on density. In one evolution treatment, evolution accelerated population growth by up to 10.3% at high initial density or reduced it by up to 6.4% at low initial density. The impact of evolution on population growth was as strong as or stronger than that caused by a threefold change in intraspecific density. We found that, taken together, ecological condition, here intraspecific density, alters evolutionary dynamics, which in turn alter concurrent population growth rate (ecological dynamics) in an eco-evolutionary feedback loop. Our results suggest that ignoring evolution in studies predicting population dynamics might lead us to over- or underestimate population density and that we cannot predict the evolutionary outcome within aphid populations without considering population size.

  18. Single Marker and Haplotype-Based Association Analysis of Semolina and Pasta Colour in Elite Durum Wheat Breeding Lines Using a High-Density Consensus Map.

    PubMed

    N'Diaye, Amidou; Haile, Jemanesh K; Cory, Aron T; Clarke, Fran R; Clarke, John M; Knox, Ron E; Pozniak, Curtis J

    2017-01-01

    Association mapping is usually performed by testing the correlation between a single marker and phenotypes. However, because patterns of variation within genomes are inherited as blocks, clustering markers into haplotypes for genome-wide scans could be a worthwhile approach to improve statistical power to detect associations. The availability of high-density molecular data allows the possibility to assess the potential of both approaches to identify marker-trait associations in durum wheat. In the present study, we used single marker- and haplotype-based approaches to identify loci associated with semolina and pasta colour in durum wheat, the main objective being to evaluate the potential benefits of haplotype-based analysis for identifying quantitative trait loci. One hundred sixty-nine durum lines were genotyped using the Illumina 90K Infinium iSelect assay, and 12,234 polymorphic single nucleotide polymorphism (SNP) markers were generated and used to assess the population structure and the linkage disequilibrium (LD) patterns. A total of 8,581 SNPs previously localized to a high-density consensus map were clustered into 406 haplotype blocks based on the average LD distance of 5.3 cM. Combining multiple SNPs into haplotype blocks increased the average polymorphism information content (PIC) from 0.27 per SNP to 0.50 per haplotype. The haplotype-based analysis identified 12 loci associated with grain pigment colour traits, including the five loci identified by the single marker-based analysis. Furthermore, the haplotype-based analysis resulted in an increase of the phenotypic variance explained (50.4% on average) and the allelic effect (33.7% on average) when compared to single marker analysis. The presence of multiple allelic combinations within each haplotype locus offers potential for screening the most favorable haplotype series and may facilitate marker-assisted selection of grain pigment colour in durum wheat. These results suggest a benefit of haplotype-based analysis over single marker analysis to detect loci associated with colour traits in durum wheat.

  19. Fixed-Precision Sequential Sampling Plans for Estimating Alfalfa Caterpillar, Colias lesbia, Egg Density in Alfalfa, Medicago sativa, Fields in Córdoba, Argentina

    PubMed Central

    Serra, Gerardo V.; Porta, Norma C. La; Avalos, Susana; Mazzuferi, Vilma

    2013-01-01

    The alfalfa caterpillar, Colias lesbia (Fabricius) (Lepidoptera: Pieridae), is a major pest of alfalfa, Medicago sativa L. (Fabales: Fabaceae), crops in Argentina. Its management is based mainly on chemical control of larvae whenever the larvae exceed the action threshold. To develop and validate fixed-precision sequential sampling plans, an intensive sampling programme for C. lesbia eggs was carried out in two alfalfa plots located in the Province of Córdoba, Argentina, from 1999 to 2002. Using Resampling for Validation of Sampling Plans software, 12 additional independent data sets were used to validate the sequential sampling plan with precision levels of 0.10 and 0.25 (SE/mean), respectively. For a range of mean densities of 0.10 to 8.35 eggs/sample, an average sample size of only 27 and 26 sample units was required to achieve a desired precision level of 0.25 for the sampling plans of Green and Kuno, respectively. As the precision level was increased to 0.10, average sample size increased to 161 and 157 sample units for the sampling plans of Green and Kuno, respectively. We recommend using Green's sequential sampling plan because it is less sensitive to changes in egg density. These sampling plans are a valuable tool for researchers to study population dynamics and to evaluate integrated pest management strategies. PMID:23909840

  20. Construction of an SNP-based high-density linkage map for flax (Linum usitatissimum L.) using specific length amplified fragment sequencing (SLAF-seq) technology.

    PubMed

    Yi, Liuxi; Gao, Fengyun; Siqin, Bateer; Zhou, Yu; Li, Qiang; Zhao, Xiaoqing; Jia, Xiaoyun; Zhang, Hui

    2017-01-01

    Flax is an important crop for oil and fiber, however, no high-density genetic maps have been reported for this species. Specific length amplified fragment sequencing (SLAF-seq) is a high-resolution strategy for large scale de novo discovery and genotyping of single nucleotide polymorphisms. In this study, SLAF-seq was employed to develop SNP markers in an F2 population to construct a high-density genetic map for flax. In total, 196.29 million paired-end reads were obtained. The average sequencing depth was 25.08 in male parent, 32.17 in the female parent, and 9.64 in each F2 progeny. In total, 389,288 polymorphic SLAFs were detected, from which 260,380 polymorphic SNPs were developed. After filtering, 4,638 SNPs were found suitable for genetic map construction. The final genetic map included 4,145 SNP markers on 15 linkage groups and was 2,632.94 cM in length, with an average distance of 0.64 cM between adjacent markers. To our knowledge, this map is the densest SNP-based genetic map for flax. The SNP markers and genetic map reported in here will serve as a foundation for the fine mapping of quantitative trait loci (QTLs), map-based gene cloning and marker assisted selection (MAS) for flax.

  1. Effect of selective logging on genetic diversity and gene flow in Cariniana legalis sampled from a cacao agroforestry system.

    PubMed

    Leal, J B; Santos, R P; Gaiotto, F A

    2014-01-28

    The fragments of the Atlantic Forest of southern Bahia have a long history of intense logging and selective cutting. Some tree species, such as jequitibá rosa (Cariniana legalis), have experienced a reduction in their populations with respect to both area and density. To evaluate the possible effects of selective logging on genetic diversity, gene flow, and spatial genetic structure, 51 C. legalis individuals were sampled, representing the total remaining population from the cacao agroforestry system. A total of 120 alleles were observed from the 11 microsatellite loci analyzed. The average observed heterozygosity (0.486) was less than the expected heterozygosity (0.721), indicating a loss of genetic diversity in this population. A high fixation index (FIS = 0.325) was found, which is possibly due to a reduction in population size, resulting in increased mating among relatives. The maximum (1055 m) and minimum (0.095 m) distances traveled by pollen or seeds were inferred based on paternity tests. We found 36.84% of unique parents among all sampled seedlings. The progenitors of the remaining seedlings (63.16%) were most likely out of the sampled area. Positive and significant spatial genetic structure was identified in this population among classes 10 to 30 m away with an average coancestry coefficient between pairs of individuals of 0.12. These results suggest that the agroforestry system of cacao cultivation is contributing to maintaining levels of diversity and gene flow in the studied population, thus minimizing the effects of selective logging.

  2. Assessing the roles of population density and predation risk in the evolution of offspring size in populations of a placental fish

    PubMed Central

    Schrader, Matthew; Travis, Joseph

    2012-01-01

    Population density is an ecological variable that is hypothesized to be a major agent of selection on offspring size. In high-density populations, high levels of intraspecific competition are expected to favor the production of larger offspring. In contrast, lower levels of intraspecific competition and selection for large offspring should be weaker and more easily overridden by direct selection for increased fecundity in low-density populations. Some studies have found associations between population density and offspring size consistent with this hypothesis. However, their interpretations are often clouded by a number of issues. Here, we use data from a 10-year study of nine populations of the least killifish, Heterandria formosa, to describe the associations of offspring size with habitat type, population density, and predation risk. We found that females from spring populations generally produced larger offspring than females from ponds; however, the magnitude of this difference varied among years. Across all populations, larger offspring were associated with higher densities and lower risks of predation. Interestingly, the associations between the two ecological variables (density and predation risk) and offspring size were largely independent of one another. Our results suggest that previously described genetic differences in offspring size are due to density-dependent natural selection. PMID:22957156

  3. Intraspecific competition and density dependence of food consumption and growth in Arctic charr.

    PubMed

    Amundsen, Per-Arne; Knudsen, Rune; Klemetsen, Anders

    2007-01-01

    1. Intraspecific competition for restricted food resources is considered to play a fundamental part in density dependence of somatic growth and other population characteristics, but studies simultaneously addressing the interrelationships between population density, food acquisition and somatic growth have been missing. 2. We explored the food consumption and individual growth rates of Arctic charr Salvelinus alpinus in a long-term survey following a large-scale density manipulation experiment in a subarctic lake. 3. Prior to the initiation of the experiment, the population density was high and the somatic growth rates low, revealing a severely overcrowded and stunted fish population. 4. During the 6-year period of stock depletion the population density of Arctic charr was reduced with about 75%, resulting in an almost twofold increase in food consumption rates and enhanced individual growth rates of the fish. 5. Over the decade following the density manipulation experiment, the population density gradually rose to intermediate levels, accompanied by corresponding reductions in food consumption and somatic growth rates. 6. The study revealed negative relationships with population density for both food consumption and individual growth rates, reflecting a strong positive correlation between quantitative food intake and somatic growth rates. 7. Both the growth and consumption rate relationships with population density were well described by negative power curves, suggesting that large density perturbations are necessary to induce improved feeding conditions and growth rates in stunted fish populations. 8. The findings demonstrate that quantitative food consumption represents the connective link between population density and individual growth rates, apparently being highly influenced by intraspecific competition for limited resources.

  4. Effects of high density on spacing behaviour and reproduction in Akodon azarae: A fencing experiment

    NASA Astrophysics Data System (ADS)

    Ávila, Belén; Bonatto, Florencia; Priotto, José; Steinmann, Andrea R.

    2016-01-01

    We studied the short term spacing behavioural responses of Pampean grassland mouse (Akodon azarae) with regard to population density in four 0.25 ha enclosures (two control and two experimental) in the 2011 breeding season. Based on the hypothesis that A. azarae breeding females exhibit spacing behaviour, and breeding males show a fusion spatial response, we tested the following predictions: (1) home range size and intrasexual overlap degree of females are independent of population density values; (2) at high population density, home range size of males decreases and the intrasexual home range overlap degree increases. To determine if female reproductive success decreases at high population density, we analyzed pregnancy rate, size and weight of litters, and period until fecundation in both low and high enclosure population density. We found that both males and females varied their home range size in relation to population density. Although male home ranges were always bigger than those of females in populations with high density, home range sizes of both sexes decreased. Females kept exclusive home ranges independent of density values meanwhile males decreased home range overlap in high breeding density populations. Although females produced litters of similar size in both treatments, weight of litter, pregnant rate and period until fecundation varied in relation to population density. Our results did not support the hypothesis that at high density females of A. azarae exhibit spacing behaviour neither that males exhibit a fusion spatial response.

  5. Causes and consequences of complex population dynamics in an annual plant, Cardamine pensylvanica

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Crone, E.E.

    1995-11-08

    The relative importance of density-dependent and density-independent factors in determining the population dynamics of plants has been widely debated with little resolution. In this thesis, the author explores the effects of density-dependent population regulation on population dynamics in Cardamine pensylvanica, an annual plant. In the first chapter, she shows that experimental populations of C. pensylvanica cycled from high to low density in controlled constant-environment conditions. These cycles could not be explained by external environmental changes or simple models of direct density dependence (N{sub t+1} = f[N{sub t}]), but they could be explained by delayed density dependence (N{sub t+1} = f[N{submore » t}, N{sub t+1}]). In the second chapter, she shows that the difference in the stability properties of population growth models with and without delayed density dependence is due to the presence of Hopf as well as slip bifurcations from stable to chaotic population dynamics. She also measures delayed density dependence due to effects of parental density on offspring quality in C. pensylvanica and shows that this is large enough to be the cause of the population dynamics observed in C. pensylvanica. In the third chapter, the author extends her analyses of density-dependent population growth models to include interactions between competing species. In the final chapter, she compares the effects of fixed spatial environmental variation and variation in population size on the evolutionary response of C. pensylvanica populations.« less

  6. Lipid levels among African and Middle-Eastern Bedouin populations.

    PubMed

    Dreiher, Jacob; Cohen, Arnon D; Weitzman, Shimon; Sharf, Amir; Shvartzman, Pesach

    2008-06-01

    Previous studies observed higher high-density lipoprotein (HDL) levels and lower triglycerides levels among people of African ancestry. The goal of this study was to characterize lipid levels in Bedouins of African vs. Middle-Eastern ethnicity. A cross-sectional study was conducted in a Bedouin primary care clinic in southern Israel, with 4470 listed individuals over the age of 21, of whom 402 (9%) were of African origin. A stratified random sample was included in the analysis. Associations between ethnicity, age, gender and lipid levels were assessed. Multiple linear regression and logistic regression models were used for multivariate analysis. The study included 261 African Bedouins and 406 Middle-Eastern Bedouins. (median age: 37 years, 58.6% females). The average total cholesterol and low-density lipoprotein (LDL) levels were 10 mg/dl lower among African Bedouins as compared to Middle-Eastern Bedouins (total cholesterol: 168.6 vs. 179.6 mg/dl, p<0.001; LDL: 99.5 vs. 109.0 mg/dl, respectively, p<0.001). Average triglycerides levels were 36 mg/dl lower among African Bedouins as compared to Middle-Eastern Bedouins (102.8 vs. 138.9 mg/dl, respectively, p<0.001). Average HDL levels were 3 mg/dl higher among African Bedouins as compared to Middle-Eastern Bedouins (48.3 vs. 44.6 mg/dl, respectively, p<0.001). A lower prevalence of dyslipidemia was found in African Bedouins, as compared with Middle-Eastern Bedouins.

  7. 42 CFR 505.5 - Loan criteria.

    Code of Federal Regulations, 2011 CFR

    2011-10-01

    ... meet the following criteria: (1) The hospital is located in a State that, based on population density, is defined as a rural State. A rural State is one of ten States with the lowest population density... prioritized beginning with the State with the lowest population density. Population density is determined...

  8. 42 CFR 505.5 - Loan criteria.

    Code of Federal Regulations, 2010 CFR

    2010-10-01

    ... meet the following criteria: (1) The hospital is located in a State that, based on population density, is defined as a rural State. A rural State is one of ten States with the lowest population density... prioritized beginning with the State with the lowest population density. Population density is determined...

  9. Do minority and poor neighborhoods have higher access to fast-food restaurants in the United States?

    PubMed Central

    James, Peter; Arcaya, Mariana C.; Parker, Devin M.; Tucker-Seeley, Reginald

    2016-01-01

    Background Disproportionate access to unhealthy foods in poor or minority neighborhoods may be a primary determinant of obesity disparities. We investigated whether fast-food access varies by Census block group (CBG) percent black and poverty. Methods We measured the average driving distance from each CBG population-weighted centroid to the five closest top ten fast-food chains and CBG percent black and percent below poverty Results Among 209,091 CBGs analyzed (95.1% of all US CBGs), CBG percent black was positively associated with fast-food access controlling for population density and percent poverty (average distance to fast food was 3.56 miles closer (95% CI: -3.64, -3.48) in CBGs with the highest versus lowest quartile of percentage of black residents). Poverty was not independently associated with fast-food access. The relationship between fast-food access and race was stronger in CBGs with higher levels of poverty (p for interaction <0.0001). Conclusions Predominantly black neighborhoods had higher access to fast-food while poverty was not an independent predictor of fast-food access. PMID:24945103

  10. Do minority and poor neighborhoods have higher access to fast-food restaurants in the United States?

    PubMed

    James, Peter; Arcaya, Mariana C; Parker, Devin M; Tucker-Seeley, Reginald D; Subramanian, S V

    2014-09-01

    Disproportionate access to unhealthy foods in poor or minority neighborhoods may be a primary determinant of obesity disparities. We investigated whether fast-food access varies by Census block group (CBG) percent black and poverty. We measured the average driving distance from each CBG population-weighted centroid to the five closest top ten fast-food chains and CBG percent black and percent below poverty. Among 209,091 CBGs analyzed (95.1% of all US CBGs), CBG percent black was positively associated with fast-food access controlling for population density and percent poverty (average distance to fast-food was 3.56 miles closer (95% CI: -3.64, -3.48) in CBGs with the highest versus lowest quartile of percentage of black residents). Poverty was not independently associated with fast-food access. The relationship between fast-food access and race was stronger in CBGs with higher levels of poverty (p for interaction <0.0001). Predominantly black neighborhoods had higher access to fast-food while poverty was not an independent predictor of fast-food access. Copyright © 2014 Elsevier Ltd. All rights reserved.

  11. Oblate-Earth Effects on the Calculation of Ec During Spacecraft Reentry

    NASA Technical Reports Server (NTRS)

    Bacon, John B.; Matney, Mark J.

    2017-01-01

    The bulge in the Earth at its equator has been shown to lead to a clustering of natural decays biased to occur towards the equator and away from the orbit's extreme latitudes. Such clustering must be considered when predicting the Expectation of Casualty (Ec) during a natural decay because of the clustering of the human population in the same lower latitudes. This study expands upon prior work, and formalizes the correction that must be made to the calculation of the average exposed population density as a result of this effect. Although a generic equation can be derived from this work to approximate the effects of gravitational and atmospheric perturbations on a final decay, such an equation averages certain important subtleties in achieving a best fit over all conditions. The authors recommend that direct simulation be used to calculate the true Ec for any specific entry as a more accurate method. A generic equation is provided, represented as a function of ballistic number and inclination of the entering spacecraft over the credible range of ballistic numbers.

  12. Sexual segregation in North American elk: the role of density dependence

    PubMed Central

    Stewart, Kelley M; Walsh, Danielle R; Kie, John G; Dick, Brian L; Bowyer, R Terry

    2015-01-01

    We investigated how density-dependent processes and subsequent variation in nutritional condition of individuals influenced both timing and duration of sexual segregation and selection of resources. During 1999–2001, we experimentally created two population densities of North American elk (Cervus elaphus), a high-density population at 20 elk/km2, and a low-density population at 4 elk/km2 to test hypotheses relative to timing and duration of sexual segregation and variation in selection of resources. We used multi-response permutation procedures to investigate patterns of sexual segregation, and resource selection functions to document differences in selection of resources by individuals in high- and low-density populations during sexual segregation and aggregation. The duration of sexual segregation was 2 months longer in the high-density population and likely was influenced by individuals in poorer nutritional condition, which corresponded with later conception and parturition, than at low density. Males and females in the high-density population overlapped in selection of resources to a greater extent than in the low-density population, probably resulting from density-dependent effects of increased intraspecific competition and lower availability of resources. PMID:25691992

  13. A geographic analysis of population density thresholds in the influenza pandemic of 1918-19.

    PubMed

    Chandra, Siddharth; Kassens-Noor, Eva; Kuljanin, Goran; Vertalka, Joshua

    2013-02-20

    Geographic variables play an important role in the study of epidemics. The role of one such variable, population density, in the spread of influenza is controversial. Prior studies have tested for such a role using arbitrary thresholds for population density above or below which places are hypothesized to have higher or lower mortality. The results of such studies are mixed. The objective of this study is to estimate, rather than assume, a threshold level of population density that separates low-density regions from high-density regions on the basis of population loss during an influenza pandemic. We study the case of the influenza pandemic of 1918-19 in India, where over 15 million people died in the short span of less than one year. Using data from six censuses for 199 districts of India (n=1194), the country with the largest number of deaths from the influenza of 1918-19, we use a sample-splitting method embedded within a population growth model that explicitly quantifies population loss from the pandemic to estimate a threshold level of population density that separates low-density districts from high-density districts. The results demonstrate a threshold level of population density of 175 people per square mile. A concurrent finding is that districts on the low side of the threshold experienced rates of population loss (3.72%) that were lower than districts on the high side of the threshold (4.69%). This paper introduces a useful analytic tool to the health geographic literature. It illustrates an application of the tool to demonstrate that it can be useful for pandemic awareness and preparedness efforts. Specifically, it estimates a level of population density above which policies to socially distance, redistribute or quarantine populations are likely to be more effective than they are for areas with population densities that lie below the threshold.

  14. A geographic analysis of population density thresholds in the influenza pandemic of 1918–19

    PubMed Central

    2013-01-01

    Background Geographic variables play an important role in the study of epidemics. The role of one such variable, population density, in the spread of influenza is controversial. Prior studies have tested for such a role using arbitrary thresholds for population density above or below which places are hypothesized to have higher or lower mortality. The results of such studies are mixed. The objective of this study is to estimate, rather than assume, a threshold level of population density that separates low-density regions from high-density regions on the basis of population loss during an influenza pandemic. We study the case of the influenza pandemic of 1918–19 in India, where over 15 million people died in the short span of less than one year. Methods Using data from six censuses for 199 districts of India (n=1194), the country with the largest number of deaths from the influenza of 1918–19, we use a sample-splitting method embedded within a population growth model that explicitly quantifies population loss from the pandemic to estimate a threshold level of population density that separates low-density districts from high-density districts. Results The results demonstrate a threshold level of population density of 175 people per square mile. A concurrent finding is that districts on the low side of the threshold experienced rates of population loss (3.72%) that were lower than districts on the high side of the threshold (4.69%). Conclusions This paper introduces a useful analytic tool to the health geographic literature. It illustrates an application of the tool to demonstrate that it can be useful for pandemic awareness and preparedness efforts. Specifically, it estimates a level of population density above which policies to socially distance, redistribute or quarantine populations are likely to be more effective than they are for areas with population densities that lie below the threshold. PMID:23425498

  15. Life history of the vulnerable endemic crayfish Cambarus (Erebicambarus) maculatus Hobbs and Pflieger, 1988 (Decapoda: Astacoidea: Cambaridae) in Missouri, USA

    USGS Publications Warehouse

    DiStefano, Robert J.; Westhoff, Jacob T.; Ames, Catlin W.; Rosenberger, Amanda E.

    2016-01-01

    The vulnerable freckled crayfish, Cambarus maculatus Hobbs and Pflieger, 1988, is endemic to only one drainage in eastern Missouri, USA, which is impacted by heavy metals mining and adjacent to a rapidly-expanding urban area. We studied populations of C. maculatus in two small streams for 25 months to describe annual reproductive cycles, and gather information about fecundity, sex ratio, size at maturity, size-class structure, and growth, capturing a monthly average of more than 50 individuals from each of the two study populations. Information about the density of the species at supplemental sampling streams was also obtained. The species exhibited traits consistent with a K-strategist life history; long-lived, slow-growing, with fewer but larger eggs than sympatric crayfish species. Breeding season occurred in mid- to late autumn, potentially extending into early winter. Egg brooding occurred primarily in May. Young of year were first observed in June. We estimated that these populations contained four to six size-classes, observed smaller individuals grew faster than larger individuals, and most became sexually mature in their second year of life. Densities of C. maculatus were low relative to several sympatric species of Orconectes Cope, 1872. Life history information presented herein will be important for anticipated future conservation efforts.

  16. Robust permanence for ecological equations with internal and external feedbacks.

    PubMed

    Patel, Swati; Schreiber, Sebastian J

    2018-07-01

    Species experience both internal feedbacks with endogenous factors such as trait evolution and external feedbacks with exogenous factors such as weather. These feedbacks can play an important role in determining whether populations persist or communities of species coexist. To provide a general mathematical framework for studying these effects, we develop a theorem for coexistence for ecological models accounting for internal and external feedbacks. Specifically, we use average Lyapunov functions and Morse decompositions to develop sufficient and necessary conditions for robust permanence, a form of coexistence robust to large perturbations of the population densities and small structural perturbations of the models. We illustrate how our results can be applied to verify permanence in non-autonomous models, structured population models, including those with frequency-dependent feedbacks, and models of eco-evolutionary dynamics. In these applications, we discuss how our results relate to previous results for models with particular types of feedbacks.

  17. Global Intraurban Intake Fractions for Primary Air Pollutants from Vehicles and Other Distributed Sources

    PubMed Central

    2012-01-01

    We model intraurban intake fraction (iF) values for distributed ground-level emissions in all 3646 global cities with more than 100 000 inhabitants, encompassing a total population of 2.0 billion. For conserved primary pollutants, population-weighted median, mean, and interquartile range iF values are 26, 39, and 14–52 ppm, respectively, where 1 ppm signifies 1 g inhaled/t emitted. The global mean urban iF reported here is roughly twice as large as previous estimates for cities in the United States and Europe. Intake fractions vary among cities owing to differences in population size, population density, and meteorology. Sorting by size, population-weighted mean iF values are 65, 35, and 15 ppm, respectively, for cities with populations larger than 3, 0.6–3, and 0.1–0.6 million. The 20 worldwide megacities (each >10 million people) have a population-weighted mean iF of 83 ppm. Mean intraurban iF values are greatest in Asia and lowest in land-rich high-income regions. Country-average iF values vary by a factor of 3 among the 10 nations with the largest urban populations. PMID:22332712

  18. Identification of QTLs associated with oil content in a high-oil Brassica napus cultivar and construction of a high-density consensus map for QTLs comparison in B. napus.

    PubMed

    Wang, Xiaodong; Wang, Hao; Long, Yan; Li, Dianrong; Yin, Yongtai; Tian, Jianhua; Chen, Li; Liu, Liezhao; Zhao, Weiguo; Zhao, Yajun; Yu, Longjiang; Li, Maoteng

    2013-01-01

    Increasing seed oil content is one of the most important goals in breeding of rapeseed (B. napus L.). To dissect the genetic basis of oil content in B. napus, a large and new double haploid (DH) population containing 348 lines was obtained from a cross between 'KenC-8' and 'N53-2', two varieties with >10% difference in seed oil content, and this population was named the KN DH population. A genetic linkage map consisting of 403 markers was constructed, which covered a total length of 1783.9 cM with an average marker interval of 4.4 cM. The KN DH population was phenotyped in eight natural environments and subjected to quantitative trait loci (QTL) analysis for oil content. A total of 63 identified QTLs explaining 2.64-17.88% of the phenotypic variation were identified, and these QTLs were further integrated into 24 consensus QTLs located on 11 chromosomes using meta-analysis. A high-density consensus map with 1335 marker loci was constructed by combining the KN DH map with seven other published maps based on the common markers. Of the 24 consensus QTLs in the KN DH population, 14 were new QTLs including five new QTLs in A genome and nine in C genome. The analysis revealed that a larger population with significant differences in oil content gave a higher power detecting new QTLs for oil content, and the construction of the consensus map provided a new clue for comparing the QTLs detected in different populations. These findings enriched our knowledge of QTLs for oil content and should be a potential in marker-assisted breeding of B. napus.

  19. Identification of QTLs Associated with Oil Content in a High-Oil Brassica napus Cultivar and Construction of a High-Density Consensus Map for QTLs Comparison in B. napus

    PubMed Central

    Long, Yan; Li, Dianrong; Yin, Yongtai; Tian, Jianhua; Chen, Li; Liu, Liezhao; Zhao, Weiguo; Zhao, Yajun; Yu, Longjiang; Li, Maoteng

    2013-01-01

    Increasing seed oil content is one of the most important goals in breeding of rapeseed (B. napus L.). To dissect the genetic basis of oil content in B. napus, a large and new double haploid (DH) population containing 348 lines was obtained from a cross between ‘KenC-8’ and ‘N53-2’, two varieties with >10% difference in seed oil content, and this population was named the KN DH population. A genetic linkage map consisting of 403 markers was constructed, which covered a total length of 1783.9 cM with an average marker interval of 4.4 cM. The KN DH population was phenotyped in eight natural environments and subjected to quantitative trait loci (QTL) analysis for oil content. A total of 63 identified QTLs explaining 2.64–17.88% of the phenotypic variation were identified, and these QTLs were further integrated into 24 consensus QTLs located on 11 chromosomes using meta-analysis. A high-density consensus map with 1335 marker loci was constructed by combining the KN DH map with seven other published maps based on the common markers. Of the 24 consensus QTLs in the KN DH population, 14 were new QTLs including five new QTLs in A genome and nine in C genome. The analysis revealed that a larger population with significant differences in oil content gave a higher power detecting new QTLs for oil content, and the construction of the consensus map provided a new clue for comparing the QTLs detected in different populations. These findings enriched our knowledge of QTLs for oil content and should be a potential in marker-assisted breeding of B. napus. PMID:24312482

  20. [The burden of disease attributed to low bone mineral density among population aged ≥40 years old in China, 1990 and 2013].

    PubMed

    Zhao, Z P; Ai, H H; Li, Y C; Wang, L M; Yin, P; Zhang, M; Deng, Q; Huang, Z J; Liu, J M; Liu, Y N; Gao, Y J; Zhou, M G

    2016-09-06

    Objective: To identify cause-specific death and attributed burden of low bone mineral density in China among population aged ≥40 years old , 1990 and 2013. Methods: By using data from Global Burden of Disease(GBD)2013, this study analyzed death caused by low mineral density, and disability-adjusted life years(DALY)among population aged 40 and above in China(not including Taiwan, China). This study also analyzed DALY by composition of injury which due to low bone mineral density. It also analyzed changes in DALY by provinces in China, 1990 and 2013. An average world population age-structure for the period 2000- 2025 was adopted to calculate the age standardized rates. Results: In 2013, there were 38.1 thousands male and 30.7 thousands female who aged 40 and above dead due to low bone mineral density in China. The burden of injury caused by low bone mineral density was more sever in male than female, which accounted for 1.525 million DALY in male and 0.873 million DALY in female. In 1990, low bone mineral density attributed transportation and accidental injury caused 0.794 million and 0.567 million DALY losses, respectively. In 2013, low bone mineral density attributed transportation and accidental injury caused 1.421 million and 0.951 million DALY losses, respectively. Compared to 1990, DALY losses caused by transportation and accidental injury, increased by 79.1% and 67.6%, respectively. In 1990, DALY rate losses due to low bone mineral density attributed transportation and accidental injury were 68.1 per 100 000 and 48.7 per 100 000, respectively. In 2013, DALY rate losses due to low bone mineral density attributed transportation and accidental injury were 102.0 per 100 000 and 68.2 per 100 000, respectively. Compared to 1990, DALY rates which caused by transportation and accidental injury, increased by 49.8% and 40.2%, respectively. According to the ranking of standardized DALY rate in 2013 by provinces, the top 3 provinces, which standardized DALYs attributed to low bone mineral density lost the most, were Zhejiang Province(2.6 per 100 000), Jiangsu Province(2.4 per 100 000), and Fujian Province(2.2 per 100 000). Compared to 1990, the standardized rate of DALY decreased in 27 provinces, while the DALY rate increased in only 6 provinces which included Ningxia Hui Autonomous Region, Qinghai Province, Hebei Province, Guangxi Zhuang Autonomous Region, and Henan Province and Xinjiang Uygur Autonomous Region. Conclusion: This study found that the burden of health losses attributed to it was higher in men than in women. Compared to 1990, DALY rates decreased in most of the provinces, however, the rates of losses of DALY which caused by transportation and accidental injury were still increasing.

  1. Recent trends in power system reliability and implications for evaluating future investments in resiliency

    DOE PAGES

    Larsen, Peter H.; LaCommare, Kristina H.; Eto, Joseph H.; ...

    2016-10-27

    Here, this study examines the relationship between annual changes in electricity reliability reported by a large cross-section of U.S. electricity distribution utilities over a period of 13 years and a broad set of potential explanatory variables, including weather and utility characteristics. We find statistically significant correlations between the average number of power interruptions experienced annually and above average wind speeds, precipitation, lightning strikes, and a measure of population density: customers per line mile. We also find significant relationships between the average number of minutes of power interruptions experienced and above average wind speeds, precipitation, cooling degree-days, and one strategy usedmore » to mitigate the impacts of severe weather: the amount of underground transmission and distribution line miles. Perhaps most importantly, we find a significant time trend of increasing annual average number of minutes of power interruptions over time—especially when interruptions associated with extreme weather are included. Lastly, the research method described in this analysis can provide a basis for future efforts to project long-term trends in reliability and the associated benefits of strategies to improve grid resiliency to severe weather—both in the U.S. and abroad.« less

  2. Does Historical Urban Density Explain the Variation in Per Capita Carbon Dioxide Emissions Across U.S. Cities?

    NASA Astrophysics Data System (ADS)

    Campbell, K. B.

    2013-12-01

    The shape a city takes can have long-term impacts. The built environment is durable, and urban infrastructure is costly to alter post-construction, so decisions made early in a city's history have a lasting effect. Cities are some of the biggest aggregate sources of CO2 emissions but are also the areas with the lowest per capita emissions. Even though per capita emissions in urban areas are less than their rural counterparts, the variation in emissions across cities is drastic and understanding this variation can improve the way we build and plan cities. Research has been conducted on how density correlates with per capita emissions, but little has been done on how historical growth has influenced emissions. Using historical census data and the Vulcan Project's fossil fuel CO2 emissions data product, I investigate in greater detail whether historical population density in U.S. cities has had a significant impact on future CO2 emissions in the urban area and in the surrounding region. The census data includes all places that have reported a population of over 100,000 people in any decennial census between 1790 and 2000 and the land area the year that the city first crosses that 100,000-population threshold. This data is used to create the historical density measure. The Vulcan CO2 emissions data is broken down by sector. For this project I use the residential, commercial, and transportation (on road and non-road) emissions sectors on a 10x10km grid in 2002. I also control for regional variation in heating and cooling days, current urban density, average house age, median income, and variation in residential heating (gas, electric, fuel oil, and coal) as these are all known correlates of carbon dioxide emissions. Understanding if historical density better explains the variation in per capita carbon dioxide emissions across cities will help urban planners and city governments decide if it is appropriate to regulate growth during the initial boom of a city, a regulation that can be costly.

  3. Modelling space of spread Dengue Hemorrhagic Fever (DHF) in Central Java use spatial durbin model

    NASA Astrophysics Data System (ADS)

    Ispriyanti, Dwi; Prahutama, Alan; Taryono, Arkadina PN

    2018-05-01

    Dengue Hemorrhagic Fever is one of the major public health problems in Indonesia. From year to year, DHF causes Extraordinary Event in most parts of Indonesia, especially Central Java. Central Java consists of 35 districts or cities where each region is close to each other. Spatial regression is an analysis that suspects the influence of independent variables on the dependent variables with the influences of the region inside. In spatial regression modeling, there are spatial autoregressive model (SAR), spatial error model (SEM) and spatial autoregressive moving average (SARMA). Spatial Durbin model is the development of SAR where the dependent and independent variable have spatial influence. In this research dependent variable used is number of DHF sufferers. The independent variables observed are population density, number of hospitals, residents and health centers, and mean years of schooling. From the multiple regression model test, the variables that significantly affect the spread of DHF disease are the population and mean years of schooling. By using queen contiguity and rook contiguity, the best model produced is the SDM model with queen contiguity because it has the smallest AIC value of 494,12. Factors that generally affect the spread of DHF in Central Java Province are the number of population and the average length of school.

  4. Field-based evaluations of sampling techniques to support long-term monitoring of riparian ecosystems along wadeable streams on the Colorado Plateau

    USGS Publications Warehouse

    Scott, Michael L.; Reynolds, Elizabeth W.

    2007-01-01

    Compared to 5-m by 20-m tree quadrats, belt transects were shown to provide similar estimates of stand structure (stem density and stand basal area) in less than 30 percent of the time. Further, for the streams sampled, there were no statistically significant differences in stem density and basal area estimates between 10-m and 20-m belt transects and the smaller belts took approximately half the time to sample. There was, however, high variance associated with estimates of stand structure for infrequently occurring stems, such as large, relict or legacy riparian trees. Legacy riparian trees occurred in limited numbers at all sites sampled. A reachscale population census of these trees indicated that the 10-m belt transects tended to underestimate both stem density and basal area for these riparian forest elements and that a complete reach-scale census of legacy trees averaged less than one hour per site.

  5. Individual Colorimetric Observer Model

    PubMed Central

    Asano, Yuta; Fairchild, Mark D.; Blondé, Laurent

    2016-01-01

    This study proposes a vision model for individual colorimetric observers. The proposed model can be beneficial in many color-critical applications such as color grading and soft proofing to assess ranges of color matches instead of a single average match. We extended the CIE 2006 physiological observer by adding eight additional physiological parameters to model individual color-normal observers. These eight parameters control lens pigment density, macular pigment density, optical densities of L-, M-, and S-cone photopigments, and λmax shifts of L-, M-, and S-cone photopigments. By identifying the variability of each physiological parameter, the model can simulate color matching functions among color-normal populations using Monte Carlo simulation. The variabilities of the eight parameters were identified through two steps. In the first step, extensive reviews of past studies were performed for each of the eight physiological parameters. In the second step, the obtained variabilities were scaled to fit a color matching dataset. The model was validated using three different datasets: traditional color matching, applied color matching, and Rayleigh matches. PMID:26862905

  6. Critical patch size generated by Allee effect in gypsy moth, Lymantria dispar (L.)

    Treesearch

    E. Vercken; A.M. Kramer; P.C. Tobin; J.M. Drake

    2011-01-01

    Allee effects are important dynamical mechanisms in small-density populations in which per capita population growth rate increases with density. When positive density dependence is sufficiently severe (a 'strong' Allee effect), a critical density arises below which populations do not persist. For spatially distributed populations subject to dispersal, theory...

  7. Macrocyclops albidus (Copepoda: cyclopidae) for the Biocontrol of Aedes albopictus and Culex pipiens in Italy.

    PubMed

    Veronesi, Rodolfo; Carrieri, Marco; Maccagnani, Bettina; Maini, Stefano; Bellini, Romeo

    2015-03-01

    The aim of our study was to assess the potential of Macrocyclops albidus as a biological control agent against the 1st and 2nd instars of Culex pipiens and Aedes albopictus. Under laboratory conditions of prey saturation (50 1st instars/copepod), an average of 58.98% of Cx. pipiens and 54.99% of Ae. albopictus larvae were killed by 1 copepod in 24 h. Trials run in big drums containing 200 liters of water showed that the M. albidus population, inoculated in April, efficiently controlled the mosquito population for the entire season. The predator was particularly effective against Ae. albopictus, as only 2 larvae of this species were found in the treated drums, compared to 814 larvae in untreated control drums throughout the study period. No difference was observed in the control efficacy between the 2 initial densities of copepods used. The reduction in Ae. albopictus density in the drums with 100 and 500 M. albidus with respect to the control drums was 99.90 ± 0.35% and 100.0 ± 0.0%, respectively. For Cx. pipiens, the reduction in density was 88.69 ± 13.51% and 84.65% in drums inoculated with 100 and 500 copepods, respectively. Macrocyclops albidus populations survived through the winter and continued to keep the mosquito population under control during the 2008 season. The M. albidus population developed very well both in drums placed in sunny and shaded areas and proved to be tolerant to both high (summer) and low (winter) temperatures. Trials performed on M. albidus survival in catch basins showed that after a few weeks, the copepod population dramatically decreased and subsequently disappeared. The main problem for copepod survival in catch basins seemed to be the low oxygen tension and accumulation of toxic substances, rather than copepods being flushed out in heavy rainfall episodes. During the period when copepods were present, they maintained the mosquito population under control; their partial disappearance from the catch basins, however, would require more recolonization intervention to maintain mosquito control during the season.

  8. Alcohol outlet density and violence: A geographically weighted regression approach.

    PubMed

    Cameron, Michael P; Cochrane, William; Gordon, Craig; Livingston, Michael

    2016-05-01

    We investigate the relationship between outlet density (of different types) and violence (as measured by police activity) across the North Island of New Zealand, specifically looking at whether the relationships vary spatially. We use New Zealand data at the census area unit (approximately suburb) level, on police-attended violent incidents and outlet density (by type of outlet), controlling for population density and local social deprivation. We employed geographically weighted regression to obtain both global average and locally specific estimates of the relationships between alcohol outlet density and violence. We find that bar and night club density, and licensed club density (e.g. sports clubs) have statistically significant and positive relationships with violence, with an additional bar or night club is associated with nearly 5.3 additional violent events per year, and an additional licensed club associated with 0.8 additional violent events per year. These relationships do not show significant spatial variation. In contrast, the effects of off-licence density and restaurant/café density do exhibit significant spatial variation. However, the non-varying effects of bar and night club density are larger than the locally specific effects of other outlet types. The relationships between outlet density and violence vary significantly across space for off-licences and restaurants/cafés. These results suggest that in order to minimise alcohol-related harms, such as violence, locally specific policy interventions are likely to be necessary. [Cameron MP, Cochrane W, Gordon C, Livingston M. Alcohol outlet density and violence: A geographically weighted regression approach. Drug Alcohol Rev 2016;35:280-288]. © 2015 Australasian Professional Society on Alcohol and other Drugs.

  9. Ecological drivers of guanaco recruitment: variable carrying capacity and density dependence.

    PubMed

    Marino, Andrea; Pascual, Miguel; Baldi, Ricardo

    2014-08-01

    Ungulates living in predator-free reserves offer the opportunity to study the influence of food limitation on population dynamics without the potentially confounding effects of top-down regulation or livestock competition. We assessed the influence of relative forage availability and population density on guanaco recruitment in two predator-free reserves in eastern Patagonia, with contrasting scenarios of population density. We also explored the relative contribution of the observed recruitment to population growth using a deterministic linear model to test the assumption that the studied populations were closed units. The observed densities increased twice as fast as our theoretical populations, indicating that marked immigration has taken place during the recovery phase experienced by both populations, thus we rejected the closed-population assumption. Regarding the factors driving variation in recruitment, in the low- to medium-density setting, we found a positive linear relationship between recruitment and surrogates of annual primary production, whereas no density dependence was detected. In contrast, in the high-density scenario, both annual primary production and population density showed marked effects, indicating a positive relationship between recruitment and per capita food availability above a food-limitation threshold. Our results support the idea that environmental carrying capacity fluctuates in response to climatic variation, and that these fluctuations have relevant consequences for herbivore dynamics, such as amplifying density dependence in drier years. We conclude that including the coupling between environmental variability in resources and density dependence is crucial to model ungulate population dynamics; to overlook temporal changes in carrying capacity may even mask density dependence as well as other important processes.

  10. Effective size of density-dependent two-sex populations: the effect of mating systems.

    PubMed

    Myhre, A M; Engen, S; SAEther, B-E

    2017-08-01

    Density dependence in vital rates is a key feature affecting temporal fluctuations of natural populations. This has important implications for the rate of random genetic drift. Mating systems also greatly affect effective population sizes, but knowledge of how mating system and density regulation interact to affect random genetic drift is poor. Using theoretical models and simulations, we compare N e in short-lived, density-dependent animal populations with different mating systems. We study the impact of a fluctuating, density-dependent sex ratio and consider both a stable and a fluctuating environment. We find a negative relationship between annual N e /N and adult population size N due to density dependence, suggesting that loss of genetic variation is reduced at small densities. The magnitude of this decrease was affected by mating system and life history. A male-biased, density-dependent sex ratio reduces the rate of genetic drift compared to an equal, density-independent sex ratio, but a stochastic change towards male bias reduces the N e /N ratio. Environmental stochasticity amplifies temporal fluctuations in population size and is thus vital to consider in estimation of effective population sizes over longer time periods. Our results on the reduced loss of genetic variation at small densities, particularly in polygamous populations, indicate that density regulation may facilitate adaptive evolution at small population sizes. © 2017 European Society For Evolutionary Biology. Journal of Evolutionary Biology © 2017 European Society For Evolutionary Biology.

  11. Population regulation in a changing environment: Long-term changes in growth, condition and survival of sprat, Sprattus sprattus L. in the Bristol Channel, UK

    NASA Astrophysics Data System (ADS)

    Henderson, Peter A.; Henderson, Rowena C.

    2017-02-01

    Sprat, Sprattus sprattus, is the dominant pelagic species in British inshore and estuarine waters. Within the Bristol Channel the population is almost totally composed of fish < 3 years old with the adults overwintering in Bridgwater Bay. Sprat follow regular seasonal migrations and occasionally form huge aggregations which together generate considerable between sample variability. Using a 36-year monthly time series collected in the Bristol Channel since 1980, together with two periods of intensive daily and weekly sampling, sprat growth is shown to have declined almost linearly over the last 36 years coincident with increasing late summer-autumn seawater temperatures. Longevity has also declined, with age 3 + sprat > 140 mm standard length lost to the population by 1999. Further, adult condition, measured as the average weight of a 103 mm standard length adult, declined rapidly from 13.7 g in 2007 to 9 g in 2011. Despite these changes, which would have reduced age-specific fecundity, a sign-rank test showed abundance of adult sprat has shown no long-term trend and Bulmer's test indicates density-dependent regulation is operating. While sprat recruitment is shown to be responding to the sunspot cycle, the North Atlantic Oscillation and sea water temperature, the impact of these variables on adult population density is damped because of density-dependent regulation. The result is that sprat respond to environmental change with large changes in their growth and condition, but the adult abundance is constrained and shows no long-term trend. Recruitment was modelled by combining a Ricker curve with terms for the response of sprat to solar activity, the North Atlantic Oscillation and spring temperature. It is shown that the stock-recruitment relationship does not form a simple curve, but is bounded within a region in which the upper and lower constraints are defined by environmental conditions. Within this bounded region the population trajectory under differing environmental regimes can be predicted.

  12. The statistics of peaks of Gaussian random fields. [cosmological density fluctuations

    NASA Technical Reports Server (NTRS)

    Bardeen, J. M.; Bond, J. R.; Kaiser, N.; Szalay, A. S.

    1986-01-01

    A set of new mathematical results on the theory of Gaussian random fields is presented, and the application of such calculations in cosmology to treat questions of structure formation from small-amplitude initial density fluctuations is addressed. The point process equation is discussed, giving the general formula for the average number density of peaks. The problem of the proper conditional probability constraints appropriate to maxima are examined using a one-dimensional illustration. The average density of maxima of a general three-dimensional Gaussian field is calculated as a function of heights of the maxima, and the average density of 'upcrossing' points on density contour surfaces is computed. The number density of peaks subject to the constraint that the large-scale density field be fixed is determined and used to discuss the segregation of high peaks from the underlying mass distribution. The machinery to calculate n-point peak-peak correlation functions is determined, as are the shapes of the profiles about maxima.

  13. Impact of Rotylenchulus reniformis on Cotton Yield as Affected by Soil Texture and Irrigation

    PubMed Central

    Herring, Stephanie L.; Heitman, Joshua L.

    2010-01-01

    The effects of soil type, irrigation, and population density of Rotylenchulus reniformis on cotton were evaluated in a two-year microplot experiment. Six soil types, Fuquay sand, Norfolk sandy loam, Portsmouth loamy sand, Muck, Cecil sandy loam, and Cecil sandy clay, were arranged in randomized complete blocks with five replications. Each block had numerous plots previously inoculated with R. reniformis and two or more noninoculated microplots per soil type, one half of which were irrigated in each replicate for a total of 240 plots. Greatest cotton lint yields were achieved in the Muck, Norfolk sandy loam, and Portsmouth loamy sand soils. Cotton yield in the Portsmouth loamy sand did not differ from the Muck soil which averaged the greatest lint yield per plot of all soil types. Cotton yield was negatively related to R. reniformis PI (initial population density) in all soil types except for the Cecil sandy clay which had the highest clay content. Supplemental irrigation increased yields in the higher yielding Muck, Norfolk sandy loam, and Portsmouth loamy sand soils compared to the lower yielding Cecil sandy clay, Cecil sandy loam, and Fuquay sand soils. The Portsmouth sandy loam was among the highest yielding soils, and also supported the greatest R. reniformis population density. Cotton lint yield was affected more by R. reniformis Pi with irrigation in the Portsmouth loamy sand soil with a greater influence of Pi on lint yield in irrigated plots than other soils. A significant first degree PI × irrigation interaction for this soil type confirms this observation. PMID:22736865

  14. Impact of Rotylenchulus reniformis on Cotton Yield as Affected by Soil Texture and Irrigation.

    PubMed

    Herring, Stephanie L; Koenning, Stephen R; Heitman, Joshua L

    2010-12-01

    The effects of soil type, irrigation, and population density of Rotylenchulus reniformis on cotton were evaluated in a two-year microplot experiment. Six soil types, Fuquay sand, Norfolk sandy loam, Portsmouth loamy sand, Muck, Cecil sandy loam, and Cecil sandy clay, were arranged in randomized complete blocks with five replications. Each block had numerous plots previously inoculated with R. reniformis and two or more noninoculated microplots per soil type, one half of which were irrigated in each replicate for a total of 240 plots. Greatest cotton lint yields were achieved in the Muck, Norfolk sandy loam, and Portsmouth loamy sand soils. Cotton yield in the Portsmouth loamy sand did not differ from the Muck soil which averaged the greatest lint yield per plot of all soil types. Cotton yield was negatively related to R. reniformis PI (initial population density) in all soil types except for the Cecil sandy clay which had the highest clay content. Supplemental irrigation increased yields in the higher yielding Muck, Norfolk sandy loam, and Portsmouth loamy sand soils compared to the lower yielding Cecil sandy clay, Cecil sandy loam, and Fuquay sand soils. The Portsmouth sandy loam was among the highest yielding soils, and also supported the greatest R. reniformis population density. Cotton lint yield was affected more by R. reniformis Pi with irrigation in the Portsmouth loamy sand soil with a greater influence of Pi on lint yield in irrigated plots than other soils. A significant first degree PI × irrigation interaction for this soil type confirms this observation.

  15. Solar Wind 0.1-1 keV Electrons in the Corotating Interaction Regions

    NASA Astrophysics Data System (ADS)

    Wang, L.; Tao, J.; Li, G.; Wimmer-Schweingruber, R. F.; Jian, L. K.; He, J.; Tu, C.; Tian, H.; Bale, S. D.

    2017-12-01

    Here we present a statistical study of the 0.1-1 keV suprathermal electrons in the undisturbed and compressed slow/fast solar wind, for the 71 corotating interaction regions (CIRs) with good measurements from the WIND 3DP and MFI instruments from 1995 to 1997. For each of these CIRs, we separate the strahl and halo electrons based on their different behaviors in pitch angle distributions in the undisturbed and compressed solar wind. We fit both the strahl and halo energy spectra to a kappa function with an index κ index and effective temperature Teff, and calculate the pitch-angle width at half-maximum (PAHM) of the strahl population. We also integrate the electron measurements between 0.1 and 1.0 keV to obtain the number density n and average energy Eavg for the strahl and halo populations. We find that for both the strahl and halo populations within and around these CIRs, the fitted κ index strongly correlates with Teff, similar to the quiet-time solar wind (Tao et al., ApJ, 2016). The number density of both the strahl and halo shows a strong positive correlation with the electron core temperature. The strahl number density ns is correlated with the magnitude of interplanetary magnetic field, and the strahl PAHM width is anti-correlated with the solar wind speed. These results suggest that the origin of strahl electrons from the solar corona is likely related to the electron core temperature and magnetic field strength, while the production of halo electrons in the interplanetary medium could depend on the solar wind velocity.

  16. Potential misuse of avian density as a conservation metric

    USGS Publications Warehouse

    Skagen, Susan K.; Yackel Adams, Amy A.

    2011-01-01

    Effective conservation metrics are needed to evaluate the success of management in a rapidly changing world. Reproductive rates and densities of breeding birds (as a surrogate for reproductive rate) have been used to indicate the quality of avian breeding habitat, but the underlying assumptions of these metrics rarely have been examined. When birds are attracted to breeding areas in part by the presence of conspecifics and when breeding in groups influences predation rates, the effectiveness of density and reproductive rate as indicators of habitat quality is reduced. It is beneficial to clearly distinguish between individual- and population-level processes when evaluating habitat quality. We use the term reproductive rate to refer to both levels and further distinguish among levels by using the terms per capita fecundity (number of female offspring per female per year, individual level) and population growth rate (the product of density and per capita fecundity, population level). We predicted how density and reproductive rate interact over time under density-independent and density-dependent scenarios, assuming the ideal free distribution model of how birds settle in breeding habitats. We predicted population density of small populations would be correlated positively with both per capita fecundity and population growth rate due to the Allee effect. For populations in the density-dependent growth phase, we predicted no relation between density and per capita fecundity (because individuals in all patches will equilibrate to the same success rate) and a positive relation between density and population growth rate. Several ecological theories collectively suggest that positive correlations between density and per capita fecundity would be difficult to detect. We constructed a decision tree to guide interpretation of positive, neutral, nonlinear, and negative relations between density and reproductive rates at individual and population levels. ?? 2010 Society for Conservation Biology.

  17. Electron temperatures within magnetic clouds between 2 and 4 AU: Voyager 2 observations

    NASA Astrophysics Data System (ADS)

    Sittler, E. C.; Burlaga, L. F.

    1998-08-01

    We have performed an analysis of Voyager 2 plasma electron observations within magnetic clouds between 2 and 4 AU identified by Burlaga and Behannon [1982]. The analysis has been confined to three of the magnetic clouds identified by Burlaga and Behannon that had high-quality data. The general properties of the plasma electrons within a magnetic cloud are that (1) the moment electron temperature anticorrelates with the electron density within the cloud, (2) the ratio Te/Tp tends to be >1, and (3) on average, Te/Tp~7.0. All three results are consistent with previous electron observations within magnetic clouds. Detailed analyses of the core and halo populations within the magnetic clouds show no evidence of either an anticorrelation between the core temperature TC and the electron density Ne or an anticorrelation between the halo temperature TH and the electron density. Within the magnetic clouds the halo component can contribute more than 50% of the electron pressure. The anticorrelation of Te relative to Ne can be traced to the density of the halo component relative to the density of the core component. The core electrons dominate the electron density. When the density goes up, the halo electrons contribute less to the electron pressure, so we get a lower Te. When the electron density goes down, the halo electrons contribute more to the electron pressure, and Te goes up. We find a relation between the electron pressure and density of the form Pe=αNeγ with γ~0.5.

  18. Double sampling to estimate density and population trends in birds

    USGS Publications Warehouse

    Bart, Jonathan; Earnst, Susan L.

    2002-01-01

    We present a method for estimating density of nesting birds based on double sampling. The approach involves surveying a large sample of plots using a rapid method such as uncorrected point counts, variable circular plot counts, or the recently suggested double-observer method. A subsample of those plots is also surveyed using intensive methods to determine actual density. The ratio of the mean count on those plots (using the rapid method) to the mean actual density (as determined by the intensive searches) is used to adjust results from the rapid method. The approach works well when results from the rapid method are highly correlated with actual density. We illustrate the method with three years of shorebird surveys from the tundra in northern Alaska. In the rapid method, surveyors covered ~10 ha h-1 and surveyed each plot a single time. The intensive surveys involved three thorough searches, required ~3 h ha-1, and took 20% of the study effort. Surveyors using the rapid method detected an average of 79% of birds present. That detection ratio was used to convert the index obtained in the rapid method into an essentially unbiased estimate of density. Trends estimated from several years of data would also be essentially unbiased. Other advantages of double sampling are that (1) the rapid method can be changed as new methods become available, (2) domains can be compared even if detection rates differ, (3) total population size can be estimated, and (4) valuable ancillary information (e.g. nest success) can be obtained on intensive plots with little additional effort. We suggest that double sampling be used to test the assumption that rapid methods, such as variable circular plot and double-observer methods, yield density estimates that are essentially unbiased. The feasibility of implementing double sampling in a range of habitats needs to be evaluated.

  19. Patterns of climate-induced density shifts of species: poleward shifts faster in northern boreal birds than in southern birds.

    PubMed

    Virkkala, Raimo; Lehikoinen, Aleksi

    2014-10-01

    Climate change has been shown to cause poleward range shifts of species. These shifts are typically demonstrated using presence-absence data, which can mask the potential changes in the abundance of species. Moreover, changes in the mean centre of weighted density of species are seldom examined, and comparisons between these two methods are even rarer. Here, we studied the change in the mean weighted latitude of density (MWLD) of 94 bird species in Finland, northern Europe, using data covering a north-south gradient of over 1000 km from the 1970s to the 2010s. The MWLD shifted northward on average 1.26 km yr(-1) , and this shift was significantly stronger in northern species compared to southern species. These shifts can be related to climate warming during the study period, because the annual temperature had increased more in northern Finland (by 1.7 °C) than in southern Finland (by 1.4 °C), although direct causal links cannot be shown. Density shifts of species distributed over the whole country did not differ from shifts in species situated on the edge of the species range in southern and northern species. This means that density shifts occur both in the core and on the edge of species distribution. The species-specific comparison of MWLD values with corresponding changes in the mean weighted latitude using presence-absence atlas data (MWL) revealed that the MWLD moved more slowly than the MWL in the atlas data in the southern species examined, but more rapidly in the northern species. Our findings highlight that population densities are also moving rapidly towards the poles and the use of presence-absence data can mask the shift of population densities. We encourage use of abundance data in studies considering the effects of climate change on biodiversity. © 2014 John Wiley & Sons Ltd.

  20. Predation and nutrients drive population declines in breeding waders.

    PubMed

    Møller, Anders Pape; Thorup, Ole; Laursen, Karsten

    2018-04-20

    Allee effects are defined as a decline in per capita fitness at low population density. We hypothesized that predation reduces population size of breeding waders and thereby the efficiency of predator deterrence, while total nitrogen through its effects on primary and secondary productivity increases population size. Therefore, nest predation could have negative consequences for population size because nest failure generally results in breeding dispersal and hence reduced local population density. To test these predictions, we recorded nest predation in five species of waders for 4,745 nests during 1987-2015 at the nature reserve Tipperne, Denmark. Predation rates were generally negatively related to conspecific and heterospecific population density, but positively related to overall population density of the entire wader community. Nest predation and population density were related to ground water level, management (grazing and mowing), and nutrients. High nest predation with a time lag of one year resulted in low overall breeding population density, while high nutrient levels resulted in higher population density. These two factors accounted for 86% of the variance in population size, presumably due to effects of nest predation on emigration, while nutrient levels increased the level of vegetation cover and the abundance of food in the surrounding brackish water. These findings are consistent with the hypothesis that predation may reduce population density through negative density dependence, while total nitrogen at adjacent shallow water may increase population size. Nest predation rates were reduced by high ground water level in March, grazing by cattle and mowing that affected access to and susceptibility of nests to predators. These effects can be managed to benefit breeding waders. © 2018 by the Ecological Society of America.

  1. 10 CFR 960.5-2-1 - Population density and distribution.

    Code of Federal Regulations, 2013 CFR

    2013-01-01

    ... 10 Energy 4 2013-01-01 2013-01-01 false Population density and distribution. 960.5-2-1 Section 960... Population density and distribution. (a) Qualifying condition. The site shall be located such that, during... specified in § 960.5-1(a)(1). (b) Favorable conditions. (1) A low population density in the general region...

  2. 10 CFR 960.5-2-1 - Population density and distribution.

    Code of Federal Regulations, 2012 CFR

    2012-01-01

    ... 10 Energy 4 2012-01-01 2012-01-01 false Population density and distribution. 960.5-2-1 Section 960... Population density and distribution. (a) Qualifying condition. The site shall be located such that, during... specified in § 960.5-1(a)(1). (b) Favorable conditions. (1) A low population density in the general region...

  3. 10 CFR 960.5-2-1 - Population density and distribution.

    Code of Federal Regulations, 2011 CFR

    2011-01-01

    ... 10 Energy 4 2011-01-01 2011-01-01 false Population density and distribution. 960.5-2-1 Section 960... Population density and distribution. (a) Qualifying condition. The site shall be located such that, during... specified in § 960.5-1(a)(1). (b) Favorable conditions. (1) A low population density in the general region...

  4. 10 CFR 960.5-2-1 - Population density and distribution.

    Code of Federal Regulations, 2014 CFR

    2014-01-01

    ... 10 Energy 4 2014-01-01 2014-01-01 false Population density and distribution. 960.5-2-1 Section 960... Population density and distribution. (a) Qualifying condition. The site shall be located such that, during... specified in § 960.5-1(a)(1). (b) Favorable conditions. (1) A low population density in the general region...

  5. Measuring Mars' Atmospheric Neutral Density from 160 to 220km with the MGS Electron Reflectometer

    NASA Astrophysics Data System (ADS)

    Lillis, R.; Engel, J.; Mitchell, D.; Brain, D.; Lin, R.; Bougher, S.; Acuna, M.

    2005-08-01

    The Magnetometer/Electron Reflectometer (MAG/ER) experiment aboard Mars Global Surveyor (MGS) samples the local electron population's distribution in energy and pitch angle (angle between electron velocity and local magnetic field direction) at the mapping orbit altitude of ˜400km. We develop a single-particle model of the electrons' interaction with the neutral atmosphere and motion along open field-lines connecting the solar wind to remnant crustal magnetization. Electron reflection from magnetic gradients and absorption due to inelastic collisons with atmospheric neutrals results in characteristic pitch angle (PA) distributions for open field lines. By assuming the validity of spherical harmonic expansions (Cain et al, 2003) in the strongest field regions of Mars (such as Terra Sirenum), we trace the electron paths and fit these PA distributions to our model to constrain the scale height and density of the neutral atmosphere in the region of greatest absorption, 160-220km. We analyse almost 3 martian years of MGS mapping Orbit Data and present the first measurements of Mars' neutral density above 180km. Although the uncertainties in single measurements are quite large, averaging over many measurements over a period of weeks allows us to see long-term trends. Major results are: 1) a mean density of 0.03 kg/km3 at 160km with a month-averaged variation of ˜40%, 2) a very strong annual seasonal variation, confirmed by periodogram and least-squares fit and 3) increasing seasonal density variability with distance from the equator. We see broad general agreement with predictions from Mars Thermosphere Global Circulation Model (MTGCM) simulations [Bougher et al, 2004] and with inferred densities from MGS Doppler tracking data [Tracadas et al, 2001]. Our results will help to constrain the upper boundaries of GCMs and assist orbital decay calculations for low-orbiting spacecraft, such as the 2005 Mars Reconnaissance Orbiter. We thank the NASA Jet Propulsion Laboratory for funding assistance for this research.

  6. Relationship between exposure to vector bites and antibody responses to mosquito salivary gland extracts.

    PubMed

    Fontaine, Albin; Pascual, Aurélie; Orlandi-Pradines, Eve; Diouf, Ibrahima; Remoué, Franck; Pagès, Frédéric; Fusaï, Thierry; Rogier, Christophe; Almeras, Lionel

    2011-01-01

    Mosquito-borne diseases are major health problems worldwide. Serological responses to mosquito saliva proteins may be useful in estimating individual exposure to bites from mosquitoes transmitting these diseases. However, the relationships between the levels of these IgG responses and mosquito density as well as IgG response specificity at the genus and/or species level need to be clarified prior to develop new immunological markers to assess human/vector contact. To this end, a kinetic study of antibody levels against several mosquito salivary gland extracts from southeastern French individuals living in three areas with distinct ecological environments and, by implication, distinct Aedes caspius mosquito densities were compared using ELISA. A positive association was observed between the average levels of IgG responses against Ae. caspius salivary gland extracts and spatial Ae. caspius densities. Additionally, the average level of IgG responses increased significantly during the peak exposure to Ae. caspius at each site and returned to baseline four months later, suggesting short-lived IgG responses. The species-specificity of IgG antibody responses was determined by testing antibody responses to salivary gland extracts from Cx. pipiens, a mosquito that is present at these three sites at different density levels, and from two other Aedes species not present in the study area (Ae. aegypti and Ae. albopictus). The IgG responses observed against these mosquito salivary gland extracts contrasted with those observed against Ae. caspius salivary gland extracts, supporting the existence of species-specific serological responses. By considering different populations and densities of mosquitoes linked to environmental factors, this study shows, for the first time, that specific IgG antibody responses against Ae. caspius salivary gland extracts may be related to the seasonal and geographical variations in Ae. caspius density. Characterisation of such immunological-markers may allow the evaluation of the effectiveness of vector-control strategies or estimation of the risk of vector-borne disease transmission.

  7. Relationship between Exposure to Vector Bites and Antibody Responses to Mosquito Salivary Gland Extracts

    PubMed Central

    Orlandi-Pradines, Eve; Diouf, Ibrahima; Remoué, Franck; Pagès, Frédéric; Fusaï, Thierry; Rogier, Christophe; Almeras, Lionel

    2011-01-01

    Mosquito-borne diseases are major health problems worldwide. Serological responses to mosquito saliva proteins may be useful in estimating individual exposure to bites from mosquitoes transmitting these diseases. However, the relationships between the levels of these IgG responses and mosquito density as well as IgG response specificity at the genus and/or species level need to be clarified prior to develop new immunological markers to assess human/vector contact. To this end, a kinetic study of antibody levels against several mosquito salivary gland extracts from southeastern French individuals living in three areas with distinct ecological environments and, by implication, distinct Aedes caspius mosquito densities were compared using ELISA. A positive association was observed between the average levels of IgG responses against Ae. caspius salivary gland extracts and spatial Ae. caspius densities. Additionally, the average level of IgG responses increased significantly during the peak exposure to Ae. caspius at each site and returned to baseline four months later, suggesting short-lived IgG responses. The species-specificity of IgG antibody responses was determined by testing antibody responses to salivary gland extracts from Cx. pipiens, a mosquito that is present at these three sites at different density levels, and from two other Aedes species not present in the study area (Ae. aegypti and Ae. albopictus). The IgG responses observed against these mosquito salivary gland extracts contrasted with those observed against Ae. caspius salivary gland extracts, supporting the existence of species-specific serological responses. By considering different populations and densities of mosquitoes linked to environmental factors, this study shows, for the first time, that specific IgG antibody responses against Ae. caspius salivary gland extracts may be related to the seasonal and geographical variations in Ae. caspius density. Characterisation of such immunological-markers may allow the evaluation of the effectiveness of vector-control strategies or estimation of the risk of vector-borne disease transmission. PMID:22195000

  8. The Frequency Detuning Correction and the Asymmetry of Line Shapes: The Far Wings of H2O-H2O

    NASA Technical Reports Server (NTRS)

    Ma, Q.; Tipping, R. H.; Hansen, James E. (Technical Monitor)

    2002-01-01

    A far-wing line shape theory which satisfies the detailed balance principle is applied to the H2O-H2O system. Within this formalism, two line shapes are introduced, corresponding to band-averages over the positive and negative resonance lines, respectively. Using the coordinate representation, the two line shapes can be obtained by evaluating 11-dimensional integrations whose integrands are a product of two factors. One depends on the interaction between the two molecules and is easy to evaluate. The other contains the density matrix of the system and is expressed as a product of two 3-dimensional distributions associated with the density matrices of the absorber and the perturber molecule, respectively. If most of the populated states are included in the averaging process, to obtain these distributions requires extensive computer CPU time, but only have to be computed once for a given temperature. The 11-dimensional integrations are evaluated using the Monte Carlo method, and in order to reduce the variance, the integration variables are chosen such that the sensitivity of the integrands on them is clearly distinguished.

  9. A Bayesian model averaging approach with non-informative priors for cost-effectiveness analyses.

    PubMed

    Conigliani, Caterina

    2010-07-20

    We consider the problem of assessing new and existing technologies for their cost-effectiveness in the case where data on both costs and effects are available from a clinical trial, and we address it by means of the cost-effectiveness acceptability curve. The main difficulty in these analyses is that cost data usually exhibit highly skew and heavy-tailed distributions, so that it can be extremely difficult to produce realistic probabilistic models for the underlying population distribution. Here, in order to integrate the uncertainty about the model into the analysis of cost data and into cost-effectiveness analyses, we consider an approach based on Bayesian model averaging (BMA) in the particular case of weak prior informations about the unknown parameters of the different models involved in the procedure. The main consequence of this assumption is that the marginal densities required by BMA are undetermined. However, in accordance with the theory of partial Bayes factors and in particular of fractional Bayes factors, we suggest replacing each marginal density with a ratio of integrals that can be efficiently computed via path sampling. Copyright (c) 2010 John Wiley & Sons, Ltd.

  10. Desertification and a shift of forest species in the West African Sahel

    USGS Publications Warehouse

    Gonzalez, Patrick

    2001-01-01

    Original field data show that forest species richness and tree density in the West African Sahel declined in the last half of the 20th century. Average forest species richness of areas of 4 km2 in Northwest Senegal fell from 64 ?? 2 species ca 1945 to 43 ?? 2 species in 1993, a decrease significant at p < 0.001. Densities of trees of height ???3 m declined from 10 ?? 0.3 trees ha-1 in 1954 to 7.8 ?? 0.3 trees ha-1 in 1989, also significant at p < 0.001. Standing wood biomass fell 2.1 t ha-1 in the period 1956-1993, releasing CO2 at a rate of 60 kgC person-1 yr-1. These changes have shifted vegetation zones toward areas of higher rainfall at an average rate of 500 to 600 m yr-1. Arid Sahel species have expanded in the north, tracking a concomitant retraction of mesic Sudan and Guinean species to the south. Multivariate analyses identify latitude and longitude, proxies for rainfall and temperature, as the most significant factors explaining tree and shrub distribution. The changes also decreased human carrying capacity to below actual population densities. The rural population of 45 people km-2 exceeded the 1993 carrying capacity, for firewood from shrubs, of 13 people km-2 (range 1 to 21 people km-2). As an adaptation strategy, ecological and socioeconomic factors favor the natural regeneration of local species over the massive plantation of exotic species. Natural regeneration is a traditional practice in which farmers select small field trees that they wish to raise to maturity, protect them, and prune them to promote rapid growth of the apical meristem. The results of this research provide evidence for desertification in the West African Sahel. These documented impacts of desertification foreshadow possible future effects of climate change.

  11. Integrated Assessment of Behavioral and Environmental Risk Factors for Lyme Disease Infection on Block Island, Rhode Island

    PubMed Central

    Krause, Peter J.; Niccolai, Linda; Steeves, Tanner; O’Keefe, Corrine Folsom; Diuk-Wasser, Maria A.

    2014-01-01

    Peridomestic exposure to Borrelia burgdorferi-infected Ixodes scapularis nymphs is considered the dominant means of infection with black-legged tick-borne pathogens in the eastern United States. Population level studies have detected a positive association between the density of infected nymphs and Lyme disease incidence. At a finer spatial scale within endemic communities, studies have focused on individual level risk behaviors, without accounting for differences in peridomestic nymphal density. This study simultaneously assessed the influence of peridomestic tick exposure risk and human behavior risk factors for Lyme disease infection on Block Island, Rhode Island. Tick exposure risk on Block Island properties was estimated using remotely sensed landscape metrics that strongly correlated with tick density at the individual property level. Behavioral risk factors and Lyme disease serology were assessed using a longitudinal serosurvey study. Significant factors associated with Lyme disease positive serology included one or more self-reported previous Lyme disease episodes, wearing protective clothing during outdoor activities, the average number of hours spent daily in tick habitat, the subject’s age and the density of shrub edges on the subject’s property. The best fit multivariate model included previous Lyme diagnoses and age. The strength of this association with previous Lyme disease suggests that the same sector of the population tends to be repeatedly infected. The second best multivariate model included a combination of environmental and behavioral factors, namely hours spent in vegetation, subject’s age, shrub edge density (increase risk) and wearing protective clothing (decrease risk). Our findings highlight the importance of concurrent evaluation of both environmental and behavioral factors to design interventions to reduce the risk of tick-borne infections. PMID:24416278

  12. Integrated assessment of behavioral and environmental risk factors for Lyme disease infection on Block Island, Rhode Island.

    PubMed

    Finch, Casey; Al-Damluji, Mohammed Salim; Krause, Peter J; Niccolai, Linda; Steeves, Tanner; O'Keefe, Corrine Folsom; Diuk-Wasser, Maria A

    2014-01-01

    Peridomestic exposure to Borrelia burgdorferi-infected Ixodes scapularis nymphs is considered the dominant means of infection with black-legged tick-borne pathogens in the eastern United States. Population level studies have detected a positive association between the density of infected nymphs and Lyme disease incidence. At a finer spatial scale within endemic communities, studies have focused on individual level risk behaviors, without accounting for differences in peridomestic nymphal density. This study simultaneously assessed the influence of peridomestic tick exposure risk and human behavior risk factors for Lyme disease infection on Block Island, Rhode Island. Tick exposure risk on Block Island properties was estimated using remotely sensed landscape metrics that strongly correlated with tick density at the individual property level. Behavioral risk factors and Lyme disease serology were assessed using a longitudinal serosurvey study. Significant factors associated with Lyme disease positive serology included one or more self-reported previous Lyme disease episodes, wearing protective clothing during outdoor activities, the average number of hours spent daily in tick habitat, the subject's age and the density of shrub edges on the subject's property. The best fit multivariate model included previous Lyme diagnoses and age. The strength of this association with previous Lyme disease suggests that the same sector of the population tends to be repeatedly infected. The second best multivariate model included a combination of environmental and behavioral factors, namely hours spent in vegetation, subject's age, shrub edge density (increase risk) and wearing protective clothing (decrease risk). Our findings highlight the importance of concurrent evaluation of both environmental and behavioral factors to design interventions to reduce the risk of tick-borne infections.

  13. Experimental Examination of Intraspecific Density-Dependent Competition during the Breeding Period in Monarch Butterflies (Danaus plexippus)

    PubMed Central

    Flockhart, D. T. Tyler; Martin, Tara G.; Norris, D. Ryan

    2012-01-01

    A central goal of population ecology is to identify the factors that regulate population growth. Monarch butterflies (Danaus plexippus) in eastern North America re-colonize the breeding range over several generations that result in population densities that vary across space and time during the breeding season. We used laboratory experiments to measure the strength of density-dependent intraspecific competition on egg laying rate and larval survival and then applied our results to density estimates of wild monarch populations to model the strength of density dependence during the breeding season. Egg laying rates did not change with density but larvae at high densities were smaller, had lower survival, and weighed less as adults compared to lower densities. Using mean larval densities from field surveys resulted in conservative estimates of density-dependent population reduction that varied between breeding regions and different phases of the breeding season. Our results suggest the highest levels of population reduction due to density-dependent intraspecific competition occur early in the breeding season in the southern portion of the breeding range. However, we also found that the strength of density dependence could be almost five times higher depending on how many life-stages were used as part of field estimates. Our study is the first to link experimental results of a density-dependent reduction in vital rates to observed monarch densities in the wild and show that the effects of density dependent competition in monarchs varies across space and time, providing valuable information for developing robust, year-round population models in this migratory organism. PMID:22984614

  14. Comparison of beverage consumption in adult populations from three different countries: do the international reference values allow establishing the adequacy of water and beverage intakes?

    PubMed

    Nissensohn, Mariela; Fuentes Lugo, Daniel; Serra-Majem, Lluis

    2016-07-13

    Recommendations of adequate total water intake (aTWI) have been proposed by the European Food Safety Agency (EFSA) and the Institute of Medicine (IOM)of the United States of America. However, there are differences in the approach used to support them: IOM recommendation is based on average intakes observed in NHANES III (Third National Health and Nutrition Examination Survey) and EFSA recommendation on a combination of observed intakes from 13 different European countries. Despite these recommendations of aTWI, the currently available scientifi c evidence is not sufficient to establish a cut-off value that would prevent disease, reduce the risk for chronic diseases or improve health status. To compare the average daily consumption of fluids (water and other beverages) in selective samples of population from Mexico, US and Spain, evaluating the quantity of fluid intake and understanding the contribution of each fluid type to the total fl uid intake. We also aim to determine if they reached adequate intake (AI) values, as defi ned by three different criteria: IOM, EFSA and water density. Three studies were compared: from Mexico, the National Health and Nutrition Survey conducted in 2012 (NHNS 2012); from US, the NHANES III 2005-2010 and from Spain the ANIBES study leaded in 2013. Different categories of beverages were used to establish the pattern of energy intake for each country. Only adult population was selected. TWI of each study was compared with EFSA and IOM AI recommendations, as well as applying the criterion of water density (mL/kcal). The American study obtained the higher value of total kcal/day from food and beverages (2,437 ± 13). Furthermore, the percentage of daily energy intake coming from beverages was, for American adults, 21%. Mexico was slightly behind with 19% and Spain ANIBES study registered only 12%. ANIBES showed signifi cantly low AI values for the overall population, but even more alarming in the case of males. Only 12% of men, in contrast with 21% of women, do satisfy the EFSA criterion. The IOM criterion reaches even less with higher recommended values for daily intake. In contrast, 60% of the American population reached the recommended intake of the IOM criterion. However, available data did not allow calculating the percentage reached by the EFSA criterion. Data from the Mexican study did not permit conducting comparisons with IOM or with EFSA. However, the water density criteria (mL/kcal) was higher than 1. There is a notable difference between all three populations in terms of TWI. Furthermore, within the same population, values of adequacy of TWI changed signifi cantly when they were assessed using different criteria. More scientifi c evidence is required for the production of better defined water intake recommendations in the future as well as more studies focusing on beverage consumption patterns in different settings.

  15. DENSITY-DEPENDENT EVOLUTION OF LIFE-HISTORY TRAITS IN DROSOPHILA MELANOGASTER.

    PubMed

    Bierbaum, Todd J; Mueller, Laurence D; Ayala, Francisco J

    1989-03-01

    Populations of Drosophila melanogaster were maintained for 36 generations in r- and K-selected environments in order to test the life-history predictions of theories on density-dependent selection. In the r-selection environment, populations were reduced to low densities by density-independent adult mortality, whereas populations in the K-selection environment were maintained at their carrying capacity. Some of the experimental results support the predictions or r- and K-selection theory; relative to the r-selected populations, the K-selected populations evolved an increased larval-to-adult viability, larger body size, and longer development time at high larval densities. Mueller and Ayala (1981) found that K-selected populations also have a higher rate of population growth at high densities. Other predictions of the thoery are contradicted by the lack of differences between the r and K populations in adult longevity and fecundity and a slower rate of development for r-selected individuals at low densities. The differences between selected populations in larval survivorship, larval-to-adult development time, and adult body size are strongly dependent on larval density, and there is a significant interaction between populations and larval density for each trait. This manifests an inadequacy of the theory on r- and K-selection, which does not take into account such interactions between genotypes and environments. We describe mechanisms that may explain the evolution of preadult life-history traits in our experiment and discuss the need for changes in theories of density-dependent selection. © 1989 The Society for the Study of Evolution.

  16. Low-energy ion distribution functions on a magnetically quiet day at geostationary altitude /L = 7/

    NASA Technical Reports Server (NTRS)

    Singh, N.; Raitt, W. J.; Yasuhara, F.

    1982-01-01

    Ion energy and pitch angle distribution functions are examined for a magnetically quiet day using averaged data from ATS 6. For both field-aligned and perpendicular fluxes, the populations have a mixture of characteristic energies, and the distribution functions can be fairly well approximated by Maxwellian distributions over three different energy bands in the range 3-600 eV. Pitch angle distributions varying with local time, and energy distributions are used to compute total ion density. Pitch angle scattering mechanisms responsible for the observed transformation of pitch angle distribution are examined, and it is found that a magnetic noise of a certain power spectral density belonging to the electromagnetic ion cyclotron mode near the ion cyclotron frequency can be effective in trapping the field aligned fluxes by pitch angle scattering.

  17. Drivers of symbiont diversity in freshwater snails: a comparative analysis of resource availability, community heterogeneity, and colonization opportunities

    PubMed Central

    McCaffrey, Keegan; Johnson, Pieter T. J.

    2017-01-01

    Decades of community ecology research have highlighted the importance of resource availability, habitat heterogeneity, and colonization opportunities in driving biodiversity. Less clear, however, is whether a similar suite of factors explains the diversity of symbionts. Here, we used a hierarchical dataset involving 12,712 freshwater snail hosts representing five species to test the relative importance of potential factors in driving symbiont richness. Specifically, we used model selection to assess the explanatory power of variables related to host species identity, resource availability (average body size, host density), ecological heterogeneity (richness of hosts and other taxa), and colonization opportunities (wetland size and amount of neighboring wetland area) on symbiont richness in 146 snail host populations in California, USA. We encountered a total of 24 taxa of symbionts, including both obligatory parasites such as digenetic trematodes as well as more commensal, mutualistic, or opportunistic groups such as aquatic insect larvae, annelids, and leeches. After validating richness estimates per host population using species accumulative curves, we detected positive effects on symbiont richness from host body size, total richness of the aquatic community, and colonization opportunities. Neither snail density nor the richness of snail species accounted for significant variation in symbiont diversity. Host species identity also affected symbiont richness, with higher gamma and average alpha diversity among more common host species and with higher local abundances. These findings highlight the importance of multiple, concurrent factors in driving symbiont richness that extend beyond epidemiological measures of host abundance or host diversity alone. PMID:28039528

  18. Esophageal eosinophilia is increased in rural areas with low population density: results from a national pathology database.

    PubMed

    Jensen, Elizabeth T; Hoffman, Kate; Shaheen, Nicholas J; Genta, Robert M; Dellon, Evan S

    2014-05-01

    Eosinophilic esophagitis (EoE) is an increasingly prevalent chronic disease arising from an allergy/immune-mediated process. Generally, the risk of atopic disease differs in rural and urban environments. The relationship between population density and EoE is unknown. Our aim was to assess the relationship between EoE and population density. We conducted a cross-sectional, case-control study of patients with esophageal biopsies in a US national pathology database between January 2009 and June 2012 to assess the relationship between population density and EoE. Using geographic information systems, the population density (individuals per square mile) was determined for each patient zip code. The odds of esophageal eosinophilia and EoE were estimated for each quintile of population density and adjusted for potential confounders. Sensitivity analyses were conducted with varying case definitions and to evaluate the potential for bias from endoscopy volume and patient factors. Of 292,621 unique patients in the source population, 89,754 had normal esophageal biopsies and 14,381 had esophageal eosinophilia with ≥15 eosinophils per high-power field. The odds of having esophageal eosinophilia increased with decreasing population density (P for trend <0.001). Compared with those in the highest quintile of population density, odds of having esophageal eosinophilia were significantly higher among those in the lowest quintile of population density (adjusted odds ratio (aOR) 1.27, 95% confidence interval (CI): 1.18, 1.36). A similar dose-response trend was observed across case definitions with increased odds of EoE in the lowest population density quintile (aOR 1.59, 95% CI: 1.45-1.76). Estimates were robust to sensitivity analyses. Population density is strongly and inversely associated with esophageal eosinophilia and EoE. This association is robust to varying case definitions and adjustment factors. Environmental exposures that are more prominent in rural areas may be relevant to the pathogenesis of EoE.

  19. Esophageal eosinophilia is increased in rural areas with low population density: Results from a national pathology database

    PubMed Central

    Jensen, Elizabeth T.; Hoffman, Kate; Shaheen, Nicholas J.; Genta, Robert M.; Dellon, Evan S.

    2015-01-01

    Objectives Eosinophilic esophagitis (EoE) is an increasingly prevalent chronic disease arising from an allergy/immune-mediated process. Generally, the risk of atopic disease differs in rural and urban environments. The relationship between population density and EoE is unknown. Our aim was to assess the relationship between EoE and population density. Methods : We conducted a cross-sectional, case-control study of patients with esophageal biopsies in a U.S. national pathology database between January 2009 and June 2012 to assess the relationship between population density and EoE. Using Geographic Information Systems (GIS), the population density (individuals/mile2) was determined for each patient zip code. The odds of esophageal eosinophilia and EoE were estimated for each quintile of population density and adjusted for potential confounders. Sensitivity analyses were conducted with varying case definitions and to evaluate the potential for bias from endoscopy volume and patient factors. Results Of 292,621 unique patients in the source population, 89,754 had normal esophageal biopsies and 14,381 had esophageal eosinophilia with ≥15 eosinophils per high-power field (eos/hpf). The odds of esophageal eosinophilia increased with decreasing population density (p for trend < 0.001). Compared to those in the highest quintile of population density, odds of esophageal eosinophilia were significantly higher amongst those in the lowest quintile of population density (aOR 1.27, 95% CI: 1.18, 1.36). A similar dose-response trend was observed across case definitions with odds of EoE increased in the lowest population density quintile (aOR 1.59, 95% CI: 1.45-1.76). Estimates were robust to sensitivity analyses. Conclusions Population density is strongly and inversely associated with esophageal eosinophilia and EoE. This association is robust to varying case definitions and adjustment factors. Environmental exposures more prominent in rural areas may be relevant to the pathogenesis of EoE. PMID:24667575

  20. Larval Population Density Alters Adult Sleep in Wild-Type Drosophila melanogaster but Not in Amnesiac Mutant Flies

    PubMed Central

    Chi, Michael W.; Griffith, Leslie C.; Vecsey, Christopher G.

    2014-01-01

    Sleep has many important biological functions, but how sleep is regulated remains poorly understood. In humans, social isolation and other stressors early in life can disrupt adult sleep. In fruit flies housed at different population densities during early adulthood, social enrichment was shown to increase subsequent sleep, but it is unknown if population density during early development can also influence adult sleep. To answer this question, we maintained Drosophila larvae at a range of population densities throughout larval development, kept them isolated during early adulthood, and then tested their sleep patterns. Our findings reveal that flies that had been isolated as larvae had more fragmented sleep than those that had been raised at higher population densities. This effect was more prominent in females than in males. Larval population density did not affect sleep in female flies that were mutant for amnesiac, which has been shown to be required for normal memory consolidation, adult sleep regulation, and brain development. In contrast, larval population density effects on sleep persisted in female flies lacking the olfactory receptor or83b, suggesting that olfactory signals are not required for the effects of larval population density on adult sleep. These findings show that population density during early development can alter sleep behavior in adulthood, suggesting that genetic and/or structural changes are induced by this developmental manipulation that persist through metamorphosis. PMID:25116571

  1. Larval Population Density Alters Adult Sleep in Wild-Type Drosophila melanogaster but Not in Amnesiac Mutant Flies.

    PubMed

    Chi, Michael W; Griffith, Leslie C; Vecsey, Christopher G

    2014-08-11

    Sleep has many important biological functions, but how sleep is regulated remains poorly understood. In humans, social isolation and other stressors early in life can disrupt adult sleep. In fruit flies housed at different population densities during early adulthood, social enrichment was shown to increase subsequent sleep, but it is unknown if population density during early development can also influence adult sleep. To answer this question, we maintained Drosophila larvae at a range of population densities throughout larval development, kept them isolated during early adulthood, and then tested their sleep patterns. Our findings reveal that flies that had been isolated as larvae had more fragmented sleep than those that had been raised at higher population densities. This effect was more prominent in females than in males. Larval population density did not affect sleep in female flies that were mutant for amnesiac, which has been shown to be required for normal memory consolidation, adult sleep regulation, and brain development. In contrast, larval population density effects on sleep persisted in female flies lacking the olfactory receptor or83b, suggesting that olfactory signals are not required for the effects of larval population density on adult sleep. These findings show that population density during early development can alter sleep behavior in adulthood, suggesting that genetic and/or structural changes are induced by this developmental manipulation that persist through metamorphosis.

  2. [Effect of the population density on growth and regeneration in the snail Achatina fulica].

    PubMed

    Sidel'nikov, A P; Stepanov, I I

    2000-01-01

    In the laboratory, the growth rate of the giant African snail Achatina fulica, as estimated by the weight and shell length was shown to decrease when the population density increased from 10 to 60 snails/m2 of the total terrarium area for five months. In the second experiment, when the population density increased from 48 to 193 snails/m2, the growth rate had already decreased by six weeks. In the groups with a high population density the feeding behavior was weakened, expressed by a greater amount of nonconsumed food, according to visual observations, than in the groups with lower population densities. At the population density of 10 to 60 snails/m2, the proliferative activity in the course of the optic tentacle regeneration, as expressed by the mitotic index, did not differ reliably within five months. In the second experiment, the mitotic indices at the population densities of 96 and 193 snails/m2 within 1.5 months exceeded that of 48 snails/m2. Recommendations are given concerning the population density from the viewpoint of commercial growth of the snails. It was proposed that, based on the analysis of the mechanism underlying the inhibition of feeding behavior in populations with extra high densities, one may develop a new approach to the production of chemical agents to control land snails as agricultural pests.

  3. A new bathyal sipunculan from Southern California, with ecological notes

    NASA Astrophysics Data System (ADS)

    Thompson, Bruce E.

    1980-11-01

    Golfingia (Nephasoma) nicolasi n. sp. is described. It is a long, slender species with a filiform introvert that is 6 to 7 times the length of the trunk. The species was often the numerically dominant taxon in samples collected from the San Nicolas Basin, California, and was also callected from several other basins off southern California. Analyses of several collections from the San Nicolas Basin show that the population was spatially patchy; temporal variation was also indicated but only one year was sampled adequately. Average population densities were highest at the base of the slopes descending into the basin from the highly productive Santa Rosa-Cortes Ridge and Tanner Bank. G. nicolasi appears to feed on the large amounts of organic detritus that accumulate from this source.

  4. Evolution of complex density-dependent dispersal strategies.

    PubMed

    Parvinen, Kalle; Seppänen, Anne; Nagy, John D

    2012-11-01

    The question of how dispersal behavior is adaptive and how it responds to changes in selection pressure is more relevant than ever, as anthropogenic habitat alteration and climate change accelerate around the world. In metapopulation models where local populations are large, and thus local population size is measured in densities, density-dependent dispersal is expected to evolve to a single-threshold strategy, in which individuals stay in patches with local population density smaller than a threshold value and move immediately away from patches with local population density larger than the threshold. Fragmentation tends to convert continuous populations into metapopulations and also to decrease local population sizes. Therefore we analyze a metapopulation model, where each patch can support only a relatively small local population and thus experience demographic stochasticity. We investigated the evolution of density-dependent dispersal, emigration and immigration, in two scenarios: adult and natal dispersal. We show that density-dependent emigration can also evolve to a nonmonotone, "triple-threshold" strategy. This interesting phenomenon results from an interplay between the direct and indirect benefits of dispersal and the costs of dispersal. We also found that, compared to juveniles, dispersing adults may benefit more from density-dependent vs. density-independent dispersal strategies.

  5. Population density and mortality among individuals in motor vehicle crashes.

    PubMed

    Gedeborg, Rolf; Thiblin, Ingemar; Byberg, Liisa; Melhus, Håkan; Lindbäck, Johan; Michaelsson, Karl

    2010-10-01

    To assess whether higher mortality rates among individuals in motor vehicle crashes in areas with low population density depend on injury type and severity or are related to the performance of emergency medical services (EMS). Prehospital and hospital deaths were studied in a population-based cohort of 41,243 motor vehicle crashes that occurred in Sweden between 1998 and 2004. The final multivariable analysis was restricted to 6884 individuals in motor vehicle crashes, to minimise the effects of confounding factors. Crude mortality rates following motor vehicle crashes were inversely related to regional population density. In regions with low population density, the unadjusted rate ratio for prehospital death was 2.2 (95% CI 1.9 to 2.5) and for hospital death 1.5 (95% CI 1.1 to 1.9), compared with a high-density population. However, after controlling for regional differences in age, gender and the type/severity of injuries among 6884 individuals in motor vehicle crashes, low population density was no longer associated with increased mortality. At 25 years of age, predicted prehospital mortality was 9% lower (95% CI 5% to 12%) in regions with low population density compared with high population density. This difference decreased with increasing age, but was still 3% lower (95% CI 0.5% to 5%) at 65 years of age. The inverse relationship between population density and mortality among individuals in motor vehicle crashes is related to pre-crash factors that influence the type and severity of injuries and not to differences in EMS.

  6. Density-dependent selection on mate search and evolution of Allee effects.

    PubMed

    Berec, Luděk; Kramer, Andrew M; Bernhauerová, Veronika; Drake, John M

    2018-01-01

    Sexually reproducing organisms require males and females to find each other. Increased difficulty of females finding mates as male density declines is the most frequently reported mechanism of Allee effects in animals. Evolving more effective mate search may alleviate Allee effects, but may depend on density regimes a population experiences. In particular, high-density populations may evolve mechanisms that induce Allee effects which become detrimental when populations are reduced and maintained at a low density. We develop an individual-based, eco-genetic model to study how mating systems and fitness trade-offs interact with changes in population density to drive evolution of the rate at which males or females search for mates. Finite mate search rate triggers Allee effects in our model and we explore how these Allee effects respond to such evolution. We allow a population to adapt to several population density regimes and examine whether high-density populations are likely to reverse adaptations attained at low densities. We find density-dependent selection in most of scenarios, leading to search rates that result in lower Allee thresholds in populations kept at lower densities. This mainly occurs when fecundity costs are imposed on mate search, and provides an explanation for why Allee effects are often observed in anthropogenically rare species. Optimizing selection, where the attained trait value minimizes the Allee threshold independent of population density, depended on the trade-off between search and survival, combined with monogamy when females were searching. Other scenarios led to runaway selection on the mate search rate, including evolutionary suicide. Trade-offs involved in mate search may thus be crucial to determining how density influences the evolution of Allee effects. Previous studies did not examine evolution of a trait related to the strength of Allee effects under density variation. We emphasize the crucial role that mating systems, fitness trade-offs and the evolving sex have in determining the density threshold for population persistence, in particular since evolution need not always take the Allee threshold to its minimum value. © 2017 The Authors. Journal of Animal Ecology © 2017 British Ecological Society.

  7. Effects of population density on corticosterone levels of prairie voles in the field

    PubMed Central

    Blondel, Dimitri V.; Wallace, Gerard N.; Calderone, Stefanie; Gorinshteyn, Marija; St. Mary, Colette M.; Phelps, Steven M.

    2015-01-01

    High population density is often associated with increased levels of stress-related hormones, such as corticosterone (CORT). Prairie voles (Microtus ochrogaster) are a socially monogamous species known for their large population density fluctuations in the wild. Although CORT influences the social behavior of prairie voles in the lab, the effect of population density on CORT has not previously been quantified in this species in the field. We validated a non-invasive hormone assay for measuring CORT metabolites in prairie vole feces. We then used semi-natural enclosures to experimentally manipulate population density, and measured density effects on male space use and fecal CORT levels. Our enclosures generated patterns of space use and social interaction that were consistent with previous prairie vole field studies. Contrary to the positive relationship between CORT and density typical of other taxa, we found that lower population densities (80 animals/ha) produced higher fecal CORT than high densities (240/ha). Combined with prior work in the lab and field, the data suggest that high prairie vole population densities indicate favorable environments, perhaps through reduced predation risk. Lastly, we found that field animals had lower fecal CORT levels than laboratory-living animals. The data emphasize the usefulness of prairie voles as models for integrating ecological, evolutionary and mechanistic questions in social behavior. PMID:26342968

  8. Paradoxical effects of density on measurement of copper tolerance in Silene paradoxa L.

    PubMed

    Capuana, Maurizio; Colzi, Ilaria; Buccianti, Antonella; Coppi, Andrea; Palm, Emily; Del Bubba, Massimo; Gonnelli, Cristina

    2018-01-01

    This work investigated if the assessment of tolerance to trace metals can depend on plant density in the experimental design. A non-metallicolous and a metallicolous populations of Silene paradoxa were hydroponically cultivated at increasing density and in both the absence (-Cu conditions) and excess of copper (+Cu conditions). In -Cu conditions, the metallicolous population showed a lower susceptibility to plant density in comparison to the non-metallicolous one, explained by a higher capacity of the metallicolous population to exploit resources. In +Cu conditions, an alleviating effect of increasing density was found in roots. Such effect was present to a greater extent in the non-metallicolous population, thus making the populations equally copper-tolerant at the highest density used. In shoots, an additive effect of increasing plant density to copper toxicity was reported. Its higher intensity in the metallicolous population reverted the copper tolerance relationship at the highest plant densities used. In both populations, a density-induced decrease in root copper accumulation was observed, thus concurring to the reported mitigation in +Cu conditions. Our work revealed the importance of density studies on the optimization of eco-toxicological bioassays and of metal tolerance assessment and it can be considered the first example of an alleviating effect of increasing plant number on copper stress in a metallophyte.

  9. Change-in-ratio density estimator for feral pigs is less biased than closed mark-recapture estimates

    USGS Publications Warehouse

    Hanson, L.B.; Grand, J.B.; Mitchell, M.S.; Jolley, D.B.; Sparklin, B.D.; Ditchkoff, S.S.

    2008-01-01

    Closed-population capture-mark-recapture (CMR) methods can produce biased density estimates for species with low or heterogeneous detection probabilities. In an attempt to address such biases, we developed a density-estimation method based on the change in ratio (CIR) of survival between two populations where survival, calculated using an open-population CMR model, is known to differ. We used our method to estimate density for a feral pig (Sus scrofa) population on Fort Benning, Georgia, USA. To assess its validity, we compared it to an estimate of the minimum density of pigs known to be alive and two estimates based on closed-population CMR models. Comparison of the density estimates revealed that the CIR estimator produced a density estimate with low precision that was reasonable with respect to minimum known density. By contrast, density point estimates using the closed-population CMR models were less than the minimum known density, consistent with biases created by low and heterogeneous capture probabilities for species like feral pigs that may occur in low density or are difficult to capture. Our CIR density estimator may be useful for tracking broad-scale, long-term changes in species, such as large cats, for which closed CMR models are unlikely to work. ?? CSIRO 2008.

  10. Spatial capture-recapture: a promising method for analyzing data collected using artificial cover objects

    USGS Publications Warehouse

    Sutherland, Chris; Munoz, David; Miller, David A.W.; Grant, Evan H. Campbell

    2016-01-01

    Spatial capture–recapture (SCR) is a relatively recent development in ecological statistics that provides a spatial context for estimating abundance and space use patterns, and improves inference about absolute population density. SCR has been applied to individual encounter data collected noninvasively using methods such as camera traps, hair snares, and scat surveys. Despite the widespread use of capture-based surveys to monitor amphibians and reptiles, there are few applications of SCR in the herpetological literature. We demonstrate the utility of the application of SCR for studies of reptiles and amphibians by analyzing capture–recapture data from Red-Backed Salamanders, Plethodon cinereus, collected using artificial cover boards. Using SCR to analyze spatial encounter histories of marked individuals, we found evidence that density differed little among four sites within the same forest (on average, 1.59 salamanders/m2) and that salamander detection probability peaked in early October (Julian day 278) reflecting expected surface activity patterns of the species. The spatial scale of detectability, a measure of space use, indicates that the home range size for this population of Red-Backed Salamanders in autumn was 16.89 m2. Surveying reptiles and amphibians using artificial cover boards regularly generates spatial encounter history data of known individuals, which can readily be analyzed using SCR methods, providing estimates of absolute density and inference about the spatial scale of habitat use.

  11. Optimal city size and population density for the 21st century.

    PubMed

    Speare A; White, M J

    1990-10-01

    The thesis that large scale urban areas result in greater efficiency, reduced costs, and a better quality of life is reexamined. The environmental and social costs are measured for different scales of settlement. The desirability and perceived problems of a particular place are examined in relation to size of place. The consequences of population decline are considered. New York city is described as providing both opportunities in employment, shopping, and cultural activities as well as a high cost of living, crime, and pollution. The historical development of large cities in the US is described. Immigration has contributed to a greater concentration of population than would have otherwise have occurred. The spatial proximity of goods and services argument (agglomeration economies) has changed with advancements in technology such as roads, trucking, and electronic communication. There is no optimal city size. The overall effect of agglomeration can be assessed by determining whether the markets for goods and labor are adequate to maximize well-being and balance the negative and positive aspects of urbanization. The environmental costs of cities increase with size when air quality, water quality, sewage treatment, and hazardous waste disposal is considered. Smaller scale and lower density cities have the advantages of a lower concentration of pollutants. Also, mobilization for program support is easier with homogenous population. Lower population growth in large cities would contribute to a higher quality of life, since large metropolitan areas have a concentration of immigrants, younger age distributions, and minority groups with higher than average birth rates. The negative consequences of decline can be avoided if reduction of population in large cities takes place gradually. For example, poorer quality housing can be removed for open space. Cities should, however, still attract all classes of people with opportunities equally available.

  12. Effects of flow regimes altered by dams on survival, population declines, and range-wide losses of California river-breeding frogs.

    PubMed

    Kupferberg, Sarah J; Palen, Wendy J; Lind, Amy J; Bobzien, Steve; Catenazzi, Alessandro; Drennan, Joe; Power, Mary E

    2012-06-01

    Widespread alteration of natural hydrologic patterns by large dams combined with peak demands for power and water delivery during summer months have resulted in frequent aseasonal flow pulses in rivers of western North America. Native species in these ecosystems have evolved with predictable annual flood-drought cycles; thus, their likelihood of persistence may decrease in response to disruption of the seasonal synchrony between stable low-flow conditions and reproduction. We evaluated whether altered flow regimes affected 2 native frogs in California and Oregon (U.S.A.) at 4 spatial and temporal extents. We examined changes in species distribution over approximately 50 years, current population density in 11 regulated and 16 unregulated rivers, temporal trends in abundance among populations occupying rivers with different hydrologic histories, and within-year patterns of survival relative to seasonal hydrology. The foothill yellow-legged frog (Rana boylii), which breeds only in flowing water, is more likely to be absent downstream of large dams than in free-flowing rivers, and breeding populations are on average 5 times smaller in regulated rivers than in unregulated rivers. Time series data (range = 8 - 19 years) from 5 populations of yellow-legged frogs and 2 populations of California red-legged frogs (R. draytonii) across a gradient of natural to highly artificial timing and magnitude of flooding indicate that variability of flows in spring and summer is strongly correlated with high mortality of early life stages and subsequent decreases in densities of adult females. Flow management that better mimics natural flow timing is likely to promote persistence of these species and others with similar phenology. ©2012 Society for Conservation Biology.

  13. The Next Generation Virgo Cluster Survey. XII. Stellar Populations and Kinematics of Compact, Low-mass Early-type Galaxies from Gemini GMOS-IFU Spectroscopy

    NASA Astrophysics Data System (ADS)

    Guérou, Adrien; Emsellem, Eric; McDermid, Richard M.; Côté, Patrick; Ferrarese, Laura; Blakeslee, John P.; Durrell, Patrick R.; MacArthur, Lauren A.; Peng, Eric W.; Cuillandre, Jean-Charles; Gwyn, Stephen

    2015-05-01

    We present Gemini Multi Object Spectrograph integral-field unit (GMOS-IFU) data of eight compact, low-mass early-type galaxies (ETGs) in the Virgo cluster. We analyze their stellar kinematics and stellar population and present two-dimensional maps of these properties covering the central 5″ × 7″ region. We find a large variety of kinematics, from nonrotating to highly rotating objects, often associated with underlying disky isophotes revealed by deep images from the Next Generation Virgo Cluster Survey. In half of our objects, we find a centrally concentrated younger and more metal-rich stellar population. We analyze the specific stellar angular momentum through the λR parameter and find six fast rotators and two slow rotators, one having a thin counterrotating disk. We compare the local galaxy density and stellar populations of our objects with those of 39 more extended low-mass Virgo ETGs from the SMAKCED survey and 260 massive (M > 1010 {{M}⊙ }) ETGs from the ATLAS3D sample. The compact low-mass ETGs in our sample are located in high-density regions, often close to a massive galaxy, and have, on average, older and more metal-rich stellar populations than less compact low-mass galaxies. We find that the stellar population parameters follow lines of constant velocity dispersion in the mass-size plane, smoothly extending the comparable trends found for massive ETGs. Our study supports a scenario where low-mass compact ETGs have experienced long-lived interactions with their environment, including ram-pressure stripping and gravitational tidal forces, that may be responsible for their compact nature.

  14. Endemic Dengue Associated with the Co-Circulation of Multiple Viral Lineages and Localized Density-Dependent Transmission

    PubMed Central

    Raghwani, Jayna; Rambaut, Andrew; Holmes, Edward C.; Hang, Vu Ty; Hien, Tran Tinh; Farrar, Jeremy; Wills, Bridget; Lennon, Niall J.; Birren, Bruce W.; Henn, Matthew R.; Simmons, Cameron P.

    2011-01-01

    Dengue is one of the most important infectious diseases of humans and has spread throughout much of the tropical and subtropical world. Despite this widespread dispersal, the determinants of dengue transmission in endemic populations are not well understood, although essential for virus control. To address this issue we performed a phylogeographic analysis of 751 complete genome sequences of dengue 1 virus (DENV-1) sampled from both rural (Dong Thap) and urban (Ho Chi Minh City) populations in southern Viet Nam during the period 2003–2008. We show that DENV-1 in Viet Nam exhibits strong spatial clustering, with likely importation from Cambodia on multiple occasions. Notably, multiple lineages of DENV-1 co-circulated in Ho Chi Minh City. That these lineages emerged at approximately the same time and dispersed over similar spatial regions suggests that they are of broadly equivalent fitness. We also observed an important relationship between the density of the human host population and the dispersion rate of dengue, such that DENV-1 tends to move from urban to rural populations, and that densely populated regions within Ho Chi Minh City act as major transmission foci. Despite these fluid dynamics, the dispersion rates of DENV-1 are relatively low, particularly in Ho Chi Minh City where the virus moves less than an average of 20 km/year. These low rates suggest a major role for mosquito-mediated dispersal, such that DENV-1 does not need to move great distances to infect a new host when there are abundant susceptibles, and imply that control measures should be directed toward the most densely populated urban environments. PMID:21655108

  15. Spatially Explicit Modeling Reveals Cephalopod Distributions Match Contrasting Trophic Pathways in the Western Mediterranean Sea

    PubMed Central

    Puerta, Patricia; Hunsicker, Mary E.; Quetglas, Antoni; Álvarez-Berastegui, Diego; Esteban, Antonio; González, María; Hidalgo, Manuel

    2015-01-01

    Populations of the same species can experience different responses to the environment throughout their distributional range as a result of spatial and temporal heterogeneity in habitat conditions. This highlights the importance of understanding the processes governing species distribution at local scales. However, research on species distribution often averages environmental covariates across large geographic areas, missing variability in population-environment interactions within geographically distinct regions. We used spatially explicit models to identify interactions between species and environmental, including chlorophyll a (Chla) and sea surface temperature (SST), and trophic (prey density) conditions, along with processes governing the distribution of two cephalopods with contrasting life-histories (octopus and squid) across the western Mediterranean Sea. This approach is relevant for cephalopods, since their population dynamics are especially sensitive to variations in habitat conditions and rarely stable in abundance and location. The regional distributions of the two cephalopod species matched two different trophic pathways present in the western Mediterranean Sea, associated with the Gulf of Lion upwelling and the Ebro river discharges respectively. The effects of the studied environmental and trophic conditions were spatially variant in both species, with usually stronger effects along their distributional boundaries. We identify areas where prey availability limited the abundance of cephalopod populations as well as contrasting effects of temperature in the warmest regions. Despite distributional patterns matching productive areas, a general negative effect of Chla on cephalopod densities suggests that competition pressure is common in the study area. Additionally, results highlight the importance of trophic interactions, beyond other common environmental factors, in shaping the distribution of cephalopod populations. Our study presents a valuable approach for understanding the spatially variant ecology of cephalopod populations, which is important for fisheries and ecosystem management. PMID:26201075

  16. Genetic structure, spatial organization, and dispersal in two populations of bat-eared foxes

    PubMed Central

    Kamler, Jan F; Gray, Melissa M; Oh, Annie; Macdonald, David W

    2013-01-01

    We incorporated radio-telemetry data with genetic analysis of bat-eared foxes (Otocyon megalotis) from individuals in 32 different groups to examine relatedness and spatial organization in two populations in South Africa that differed in density, home-range sizes, and group sizes. Kin clustering occurred only for female dyads in the high-density population. Relatedness was negatively correlated with distance only for female dyads in the high-density population, and for male and mixed-sex dyads in the low-density population. Home-range overlap of neighboring female dyads was significantly greater in the high compared to low-density population, whereas overlap within other dyads was similar between populations. Amount of home-range overlap between neighbors was positively correlated with genetic relatedness for all dyad-site combinations, except for female and male dyads in the low-density population. Foxes from all age and sex classes dispersed, although females (mostly adults) dispersed farther than males. Yearlings dispersed later in the high-density population, and overall exhibited a male-biased dispersal pattern. Our results indicated that genetic structure within populations of bat-eared foxes was sex-biased, and was interrelated to density and group sizes, as well as sex-biases in philopatry and dispersal distances. We conclude that a combination of male-biased dispersal rates, adult dispersals, and sex-biased dispersal distances likely helped to facilitate inbreeding avoidance in this evolutionarily unique species of Canidae. PMID:24101981

  17. Population density and youth antisocial behavior.

    PubMed

    Harden, K Paige; D'Onofrio, Brian M; Van Hulle, Carol; Turkheimer, Eric; Rodgers, Joseph L; Waldman, Irwin D; Lahey, Benjamin B

    2009-08-01

    Theoretical models concerning how neighborhood contexts adversely influence juvenile antisocial behavior frequently focus on urban neighborhoods; however, previous studies comparing urban and rural areas on the prevalence of youth antisocial behavior have yielded mixed results. The current study uses longitudinal data on the offspring of a nationally representative sample of mothers (N = 4,886) in the US. There was no relation between density and mother-reported child conduct problems across ages 4-13 years, but youth living in areas of greater population density exhibited more youth self-reported delinquency across 10-17 years. Families often moved to counties with greater or lesser population density, but longitudinal analyses treating population density as a time-varying covariate did not support the hypothesis that living in densely populated counties influenced youth delinquency. Rather, the association between population density and delinquency appears to be due to unmeasured selection variables that differ between families who live in more or less densely populated counties.

  18. Socio-Economic Instability and the Scaling of Energy Use with Population Size

    PubMed Central

    DeLong, John P.; Burger, Oskar

    2015-01-01

    The size of the human population is relevant to the development of a sustainable world, yet the forces setting growth or declines in the human population are poorly understood. Generally, population growth rates depend on whether new individuals compete for the same energy (leading to Malthusian or density-dependent growth) or help to generate new energy (leading to exponential and super-exponential growth). It has been hypothesized that exponential and super-exponential growth in humans has resulted from carrying capacity, which is in part determined by energy availability, keeping pace with or exceeding the rate of population growth. We evaluated the relationship between energy use and population size for countries with long records of both and the world as a whole to assess whether energy yields are consistent with the idea of an increasing carrying capacity. We find that on average energy use has indeed kept pace with population size over long time periods. We also show, however, that the energy-population scaling exponent plummets during, and its temporal variability increases preceding, periods of social, political, technological, and environmental change. We suggest that efforts to increase the reliability of future energy yields may be essential for stabilizing both population growth and the global socio-economic system. PMID:26091499

  19. Socio-Economic Instability and the Scaling of Energy Use with Population Size.

    PubMed

    DeLong, John P; Burger, Oskar

    2015-01-01

    The size of the human population is relevant to the development of a sustainable world, yet the forces setting growth or declines in the human population are poorly understood. Generally, population growth rates depend on whether new individuals compete for the same energy (leading to Malthusian or density-dependent growth) or help to generate new energy (leading to exponential and super-exponential growth). It has been hypothesized that exponential and super-exponential growth in humans has resulted from carrying capacity, which is in part determined by energy availability, keeping pace with or exceeding the rate of population growth. We evaluated the relationship between energy use and population size for countries with long records of both and the world as a whole to assess whether energy yields are consistent with the idea of an increasing carrying capacity. We find that on average energy use has indeed kept pace with population size over long time periods. We also show, however, that the energy-population scaling exponent plummets during, and its temporal variability increases preceding, periods of social, political, technological, and environmental change. We suggest that efforts to increase the reliability of future energy yields may be essential for stabilizing both population growth and the global socio-economic system.

  20. Genetic variation facilitates seedling establishment but not population growth rate of a perennial invader.

    PubMed

    Li, Shou-Li; Vasemägi, Anti; Ramula, Satu

    2016-01-01

    Assessing the demographic consequences of genetic variation is fundamental to invasion biology. However, genetic and demographic approaches are rarely combined to explore the effects of genetic variation on invasive populations in natural environments. This study combined population genetics, demographic data and a greenhouse experiment to investigate the consequences of genetic variation for the population fitness of the perennial, invasive herb Lupinus polyphyllus. Genetic and demographic data were collected from 37 L. polyphyllus populations representing different latitudes in Finland, and genetic variation was characterized based on 13 microsatellite loci. Associations between genetic variation and population size, population density, latitude and habitat were investigated. Genetic variation was then explored in relation to four fitness components (establishment, survival, growth, fecundity) measured at the population level, and the long-term population growth rate (λ). For a subset of populations genetic variation was also examined in relation to the temporal variability of λ. A further assessment was made of the role of natural selection in the observed variation of certain fitness components among populations under greenhouse conditions. It was found that genetic variation correlated positively with population size, particularly at higher latitudes, and differed among habitat types. Average seedling establishment per population increased with genetic variation in the field, but not under greenhouse conditions. Quantitative genetic divergence (Q(ST)) based on seedling establishment in the greenhouse was smaller than allelic genetic divergence (F'(ST)), indicating that unifying selection has a prominent role in this fitness component. Genetic variation was not associated with average survival, growth or fecundity measured at the population level, λ or its variability. The study suggests that although genetic variation may facilitate plant invasions by increasing seedling establishment, it may not necessarily affect the long-term population growth rate. Therefore, established invasions may be able to grow equally well regardless of their genetic diversity. © The Author 2015. Published by Oxford University Press on behalf of the Annals of Botany Company. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

  1. Initial glenoid fixation using two different reverse shoulder designs with an equivalent center of rotation in a low-density and high-density bone substitute.

    PubMed

    Stroud, Nicholas J; DiPaola, Matthew J; Martin, Brian L; Steiler, Cindy A; Flurin, Pierre-Henri; Wright, Thomas W; Zuckerman, Joseph D; Roche, Christopher P

    2013-11-01

    Numerous glenoid implant designs have been introduced into the global marketplace in recent years; however, little comparative biomechanical data exist to substantiate one design consideration over another. This study dynamically evaluated reverse shoulder glenoid baseplate fixation and compared the initial fixation associated with 2 reverse shoulder designs having an equivalent center of rotation in low-density and high-density bone substitute substrates. Significant differences in fixation were observed between implant designs, where the circular-porous reverse shoulder was associated with approximately twice the micromotion per equivalent test than the oblong-grit-blasted design. Additionally, 6 of the 7 circular-porous reverse shoulders failed catastrophically in the low-density bone model at an average of 2603 ± 981 cycles. None of the oblong-grit-blasted designs failed in the low-or high-density bone models and none of the circular-porous designs failed in the high-density bone models after 10,000 cycles of loading. These results demonstrate that significant differences in initial fixation exist between reverse shoulder implants having an equivalent center of rotation and suggest that design parameters, other than the position of the center of rotation, significantly affect fixation in low-density and high-density polyurethane bone substitutes. Subtle changes in glenoid baseplate design can dramatically affect fixation, particularly in low-density bone substitutes that are intended to simulate the bone quality of the recipient population for reverse shoulders. Copyright © 2013 Journal of Shoulder and Elbow Surgery Board of Trustees. Published by Mosby, Inc. All rights reserved.

  2. Response of bird species densities to habitat structure and fire history along a Midwestern open-forest gradient

    USGS Publications Warehouse

    Grundel, R.; Pavlovic, N.B.

    2007-01-01

    Oak savannas were historically common but are currently rare in the Midwestern United States. We assessed possible associations of bird species with savannas and other threatened habitats in the region by relating fire frequency and vegetation characteristics to seasonal densities of 72 bird species distributed across an open-forest gradient in northwestern Indiana. About one-third of the species did not exhibit statistically significant relationships with any combination of seven vegetation characteristics that included vegetation cover in five vertical strata, dead tree density, and tree height. For 40% of the remaining species, models best predicting species density incorporated tree density. Therefore, management based solely on manipulating tree density may not be an adequate strategy for managing bird populations along this open-forest gradient. Few species exhibited sharp peaks in predicted density under habitat conditions expected in restored savannas, suggesting that few savanna specialists occur among Midwestern bird species. When fire frequency, measured over fifteen years, was added to vegetation characteristics as a predictor of species density, it was incorporated into models for about one-quarter of species, suggesting that fire may modify habitat characteristics in ways that are important for birds but not captured by the structural habitat variables measured. Among those species, similar numbers had peaks in predicted density at low, intermediate, or high fire frequency. For species suggested by previous studies to have a preference for oak savannas along the open-forest gradient, estimated density was maximized at an average fire return interval of about one fire every three years. ?? The Cooper Ornithological Society 2007.

  3. Population density predicts outcome from out-of-hospital cardiac arrest in Victoria, Australia.

    PubMed

    Nehme, Ziad; Andrew, Emily; Cameron, Peter A; Bray, Janet E; Bernard, Stephen A; Meredith, Ian T; Smith, Karen

    2014-05-05

    To examine the impact of population density on incidence and outcome of out-of-hospital cardiac arrest (OHCA). Data were extracted from the Victorian Ambulance Cardiac Arrest Registry for all adult OHCA cases of presumed cardiac aetiology attended by the emergency medical service (EMS) between 1 January 2003 and 31 December 2011. Cases were allocated into one of five population density groups according to their statistical local area: very low density (≤ 10 people/km(2)), low density (11-200 people/km(2)), medium density (201-1000 people/km(2)), high density (1001-3000 people/km(2)), and very high density (> 3000 people/km(2)). Survival to hospital and survival to hospital discharge. The EMS attended 27 705 adult presumed cardiac OHCA cases across 204 Victorian regions. In 12 007 of these (43.3%), resuscitation was attempted by the EMS. Incidence was lower and arrest characteristics were consistently less favourable for lower population density groups. Survival outcomes, including return of spontaneous circulation, survival to hospital and survival to hospital discharge, were significantly poorer in less densely populated groups (P < 0.001 for all comparisons). When compared with very low density populations, the risk-adjusted odds ratios of surviving to hospital discharge were: low density, 1.88 (95% CI, 1.15-3.07); medium density, 2.49 (95% CI, 1.55-4.02); high density, 3.47 (95% CI, 2.20-5.48) and very high density, 4.32 (95% CI, 2.67-6.99). Population density is independently associated with survival after OHCA, and significant variation in the incidence and characteristics of these events are observed across the state.

  4. The Bright and Dark Sides of High-redshift Starburst Galaxies from Herschel and Subaru Observations

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Puglisi, A.; Rodighiero, G.; Rodríguez-Muñoz, L.

    2017-04-01

    We present rest-frame optical spectra from the FMOS-COSMOS survey of 12 z ∼ 1.6 Herschel starburst galaxies, with star formation rate (SFR) elevated by ×8, on average, above the star-forming main sequence (MS). Comparing the H α to IR luminosity ratio and the Balmer decrement, we find that the optically thin regions of the sources contain on average only ∼10% of the total SFR, whereas ∼90% come from an extremely obscured component that is revealed only by far-IR observations and is optically thick even in H α . We measure the [N ii]{sub 6583}/H α ratio, suggesting that the lessmore » obscured regions have a metal content similar to that of the MS population at the same stellar masses and redshifts. However, our objects appear to be metal-rich outliers from the metallicity–SFR anticorrelation observed at fixed stellar mass for the MS population. The [S ii]{sub 6732}/[S ii]{sub 6717} ratio from the average spectrum indicates an electron density n {sub e} ∼ 1100 cm{sup −3} , larger than what was estimated for MS galaxies but only at the 1.5 σ level. Our results provide supporting evidence that high- z MS outliers are analogous of local ULIRGs and are consistent with a major-merger origin for the starburst event.« less

  5. Environmental variability and population dynamics: Do European and North American ducks play by the same rules?

    USGS Publications Warehouse

    Pöysä, Hannu; Rintala, Jukka; Johnson, Douglas H.; Kauppinen, Jukka; Lammi, Esa; Nudds, Thomas D.; Väänänen, Veli-Matti

    2016-01-01

    Density dependence, population regulation, and variability in population size are fundamental population processes, the manifestation and interrelationships of which are affected by environmental variability. However, there are surprisingly few empirical studies that distinguish the effect of environmental variability from the effects of population processes. We took advantage of a unique system, in which populations of the same duck species or close ecological counterparts live in highly variable (north American prairies) and in stable (north European lakes) environments, to distinguish the relative contributions of environmental variability (measured as between-year fluctuations in wetland numbers) and intraspecific interactions (density dependence) in driving population dynamics. We tested whether populations living in stable environments (in northern Europe) were more strongly governed by density dependence than populations living in variable environments (in North America). We also addressed whether relative population dynamical responses to environmental variability versus density corresponded to differences in life history strategies between dabbling (relatively “fast species” and governed by environmental variability) and diving (relatively “slow species” and governed by density) ducks. As expected, the variance component of population fluctuations caused by changes in breeding environments was greater in North America than in Europe. Contrary to expectations, however, populations in more stable environments were not less variable nor clearly more strongly density dependent than populations in highly variable environments. Also, contrary to expectations, populations of diving ducks were neither more stable nor stronger density dependent than populations of dabbling ducks, and the effect of environmental variability on population dynamics was greater in diving than in dabbling ducks. In general, irrespective of continent and species life history, environmental variability contributed more to variation in species abundances than did density. Our findings underscore the need for more studies on populations of the same species in different environments to verify the generality of current explanations about population dynamics and its association with species life history.

  6. Environmental variability and population dynamics: do European and North American ducks play by the same rules?

    PubMed

    Pöysä, Hannu; Rintala, Jukka; Johnson, Douglas H; Kauppinen, Jukka; Lammi, Esa; Nudds, Thomas D; Väänänen, Veli-Matti

    2016-10-01

    Density dependence, population regulation, and variability in population size are fundamental population processes, the manifestation and interrelationships of which are affected by environmental variability. However, there are surprisingly few empirical studies that distinguish the effect of environmental variability from the effects of population processes. We took advantage of a unique system, in which populations of the same duck species or close ecological counterparts live in highly variable (north American prairies) and in stable (north European lakes) environments, to distinguish the relative contributions of environmental variability (measured as between-year fluctuations in wetland numbers) and intraspecific interactions (density dependence) in driving population dynamics. We tested whether populations living in stable environments (in northern Europe) were more strongly governed by density dependence than populations living in variable environments (in North America). We also addressed whether relative population dynamical responses to environmental variability versus density corresponded to differences in life history strategies between dabbling (relatively "fast species" and governed by environmental variability) and diving (relatively "slow species" and governed by density) ducks. As expected, the variance component of population fluctuations caused by changes in breeding environments was greater in North America than in Europe. Contrary to expectations, however, populations in more stable environments were not less variable nor clearly more strongly density dependent than populations in highly variable environments. Also, contrary to expectations, populations of diving ducks were neither more stable nor stronger density dependent than populations of dabbling ducks, and the effect of environmental variability on population dynamics was greater in diving than in dabbling ducks. In general, irrespective of continent and species life history, environmental variability contributed more to variation in species abundances than did density. Our findings underscore the need for more studies on populations of the same species in different environments to verify the generality of current explanations about population dynamics and its association with species life history.

  7. Dispersion patterns and sampling plans for Diaphorina citri (Hemiptera: Psyllidae) in citrus.

    PubMed

    Sétamou, Mamoudou; Flores, Daniel; French, J Victor; Hall, David G

    2008-08-01

    The abundance and spatial dispersion of Diaphorina citri Kuwayama (Hemiptera: Psyllidae) were studied in 34 grapefruit (Citrus paradisi Macfad.) and six sweet orange [Citrus sinensis (L.) Osbeck] orchards from March to August 2006 when the pest is more abundant in southern Texas. Although flush shoot infestation levels did not vary with host plant species, densities of D. citri eggs, nymphs, and adults were significantly higher on sweet orange than on grapefruit. D. citri immatures also were found in significantly higher numbers in the southeastern quadrant of trees than other parts of the canopy. The spatial distribution of D. citri nymphs and adults was analyzed using Iowa's patchiness regression and Taylor's power law. Taylor's power law fitted the data better than Iowa's model. Based on both regression models, the field dispersion patterns of D. citri nymphs and adults were aggregated among flush shoots in individual trees as indicated by the regression slopes that were significantly >1. For the average density of each life stage obtained during our surveys, the minimum number of flush shoots per tree needed to estimate D. citri densities varied from eight for eggs to four flush shoots for adults. Projections indicated that a sampling plan consisting of 10 trees and eight flush shoots per tree would provide density estimates of the three developmental stages of D. citri acceptable enough for population studies and management decisions. A presence-absence sampling plan with a fixed precision level was developed and can be used to provide a quick estimation of D. citri populations in citrus orchards.

  8. Linkage disequilibrium compared between five populations of domestic sheep.

    PubMed

    Meadows, Jennifer R S; Chan, Eva K F; Kijas, James W

    2008-09-30

    The success of genome-wide scans depends on the strength and magnitude of linkage disequilibrium (LD) present within the populations under investigation. High density SNP arrays are currently in development for the sheep genome, however little is known about the behaviour of LD in this livestock species. This study examined the behaviour of LD within five sheep populations using two LD metrics, D' and x2'. Four economically important Australian sheep flocks, three pure breeds (White Faced Suffolk, Poll Dorset, Merino) and a crossbred population (Merino x Border Leicester), along with an inbred Australian Merino museum flock were analysed. Short range LD (0 - 5 cM) was observed in all five populations, however the persistence with increasing distance and magnitude of LD varied considerably between populations. Average LD (x2') for markers spaced up to 20 cM exceeded the non-syntenic average within the White Faced Suffolk, Poll Dorset and Macarthur Merino. LD decayed faster within the Merino and Merino x Border Leicester, with LD below or consistent with observed background levels. Using marker-marker LD as a guide to the behaviour of marker-QTL LD, estimates of minimum marker spacing were made. For a 95% probability of detecting QTL, a microsatellite marker would be required every 0.1 - 2.5 centimorgans, depending on the population used. Sheep populations were selected which were inbred (Macarthur Merino), highly heterogeneous (Merino) or intermediate between these two extremes. This facilitated analysis and comparison of LD (x2') between populations. The strength and magnitude of LD was found to differ markedly between breeds and aligned closely with both observed levels of genetic diversity and expectations based on breed history. This confirmed that breed specific information is likely to be important for genome wide selection and during the design of successful genome scans where tens of thousands of markers will be required.

  9. Disentangling density-dependent dynamics using full annual cycle models and Bayesian model weight updating

    USGS Publications Warehouse

    Robinson, Orin J.; McGowan, Conor P.; Devers, Patrick K.

    2017-01-01

    Density dependence regulates populations of many species across all taxonomic groups. Understanding density dependence is vital for predicting the effects of climate, habitat loss and/or management actions on wild populations. Migratory species likely experience seasonal changes in the relative influence of density dependence on population processes such as survival and recruitment throughout the annual cycle. These effects must be accounted for when characterizing migratory populations via population models.To evaluate effects of density on seasonal survival and recruitment of a migratory species, we used an existing full annual cycle model framework for American black ducks Anas rubripes, and tested different density effects (including no effects) on survival and recruitment. We then used a Bayesian model weight updating routine to determine which population model best fit observed breeding population survey data between 1990 and 2014.The models that best fit the survey data suggested that survival and recruitment were affected by density dependence and that density effects were stronger on adult survival during the breeding season than during the non-breeding season.Analysis also suggests that regulation of survival and recruitment by density varied over time. Our results showed that different characterizations of density regulations changed every 8–12 years (three times in the 25-year period) for our population.Synthesis and applications. Using a full annual cycle, modelling framework and model weighting routine will be helpful in evaluating density dependence for migratory species in both the short and long term. We used this method to disentangle the seasonal effects of density on the continental American black duck population which will allow managers to better evaluate the effects of habitat loss and potential habitat management actions throughout the annual cycle. The method here may allow researchers to hone in on the proper form and/or strength of density dependence for use in models for conservation recommendations.

  10. Potential misuse of avian density as a conservation metric.

    PubMed

    Skagen, Susan K; Yackel Adams, Amy A

    2011-02-01

    Effective conservation metrics are needed to evaluate the success of management in a rapidly changing world. Reproductive rates and densities of breeding birds (as a surrogate for reproductive rate) have been used to indicate the quality of avian breeding habitat, but the underlying assumptions of these metrics rarely have been examined. When birds are attracted to breeding areas in part by the presence of conspecifics and when breeding in groups influences predation rates, the effectiveness of density and reproductive rate as indicators of habitat quality is reduced. It is beneficial to clearly distinguish between individual- and population-level processes when evaluating habitat quality. We use the term reproductive rate to refer to both levels and further distinguish among levels by using the terms per capita fecundity (number of female offspring per female per year, individual level) and population growth rate (the product of density and per capita fecundity, population level). We predicted how density and reproductive rate interact over time under density-independent and density-dependent scenarios, assuming the ideal free distribution model of how birds settle in breeding habitats. We predicted population density of small populations would be correlated positively with both per capita fecundity and population growth rate due to the Allee effect. For populations in the density-dependent growth phase, we predicted no relation between density and per capita fecundity (because individuals in all patches will equilibrate to the same success rate) and a positive relation between density and population growth rate. Several ecological theories collectively suggest that positive correlations between density and per capita fecundity would be difficult to detect. We constructed a decision tree to guide interpretation of positive, neutral, nonlinear, and negative relations between density and reproductive rates at individual and population levels. Journal compilation ©2010 Society for Conservation Biology. No claim to original US government works.

  11. Survival rates of female white-tailed deer on an industrial forest following a decline in population density

    Treesearch

    Shawn M. Crimmins; John W. Edwards; Patrick D. Keyser; James M. Crum; W. Mark Ford; Brad F. Miller; Tyler A. Campbell; Karl V. Miller

    2013-01-01

    With white-tailed deer (Odocoileus virginianus) populations at historically high levels throughout many North American forests, many current management activities are aimed at reducing deer populations. However, very little information exists on the ecology of low-density white-tailed deer populations or populations that have declined in density. We...

  12. Partitioning the sources of demographic variation reveals density-dependent nest predation in an island bird population

    PubMed Central

    Sofaer, Helen R; Sillett, T Scott; Langin, Kathryn M; Morrison, Scott A; Ghalambor, Cameron K

    2014-01-01

    Ecological factors often shape demography through multiple mechanisms, making it difficult to identify the sources of demographic variation. In particular, conspecific density can influence both the strength of competition and the predation rate, but density-dependent competition has received more attention, particularly among terrestrial vertebrates and in island populations. A better understanding of how both competition and predation contribute to density-dependent variation in fecundity can be gained by partitioning the effects of density on offspring number from its effects on reproductive failure, while also evaluating how biotic and abiotic factors jointly shape demography. We examined the effects of population density and precipitation on fecundity, nest survival, and adult survival in an insular population of orange-crowned warblers (Oreothlypis celata) that breeds at high densities and exhibits a suite of traits suggesting strong intraspecific competition. Breeding density had a negative influence on fecundity, but it acted by increasing the probability of reproductive failure through nest predation, rather than through competition, which was predicted to reduce the number of offspring produced by successful individuals. Our results demonstrate that density-dependent nest predation can underlie the relationship between population density and fecundity even in a high-density, insular population where intraspecific competition should be strong. PMID:25077023

  13. Partitioning the sources of demographic variation reveals density-dependent nest predation in an island bird population.

    PubMed

    Sofaer, Helen R; Sillett, T Scott; Langin, Kathryn M; Morrison, Scott A; Ghalambor, Cameron K

    2014-07-01

    Ecological factors often shape demography through multiple mechanisms, making it difficult to identify the sources of demographic variation. In particular, conspecific density can influence both the strength of competition and the predation rate, but density-dependent competition has received more attention, particularly among terrestrial vertebrates and in island populations. A better understanding of how both competition and predation contribute to density-dependent variation in fecundity can be gained by partitioning the effects of density on offspring number from its effects on reproductive failure, while also evaluating how biotic and abiotic factors jointly shape demography. We examined the effects of population density and precipitation on fecundity, nest survival, and adult survival in an insular population of orange-crowned warblers (Oreothlypis celata) that breeds at high densities and exhibits a suite of traits suggesting strong intraspecific competition. Breeding density had a negative influence on fecundity, but it acted by increasing the probability of reproductive failure through nest predation, rather than through competition, which was predicted to reduce the number of offspring produced by successful individuals. Our results demonstrate that density-dependent nest predation can underlie the relationship between population density and fecundity even in a high-density, insular population where intraspecific competition should be strong.

  14. Bone mineral density referral for dual-energy X-ray absorptiometry using quantitative ultrasound as a prescreening tool in postmenopausal women from the general population: a cost-effectiveness analysis.

    PubMed

    Marín, F; López-Bastida, J; Díez-Pérez, A; Sacristán, J A

    2004-03-01

    The aim of our study was to assess, from the perspective of the National Health Services in Spain, the cost-effectiveness of quantitative ultrasound (QUS) as a prescreen referral method for bone mineral density (BMD) assessment by dual-energy X-ray absorptiometry (DXA) in postmenopausal women of the general population. Using femoral neck DXA and heel QUS. We evaluated 267 consecutive postmenopausal women 65 years and older and attending primary care physician offices for any medical reason. Subjects were classified as osteoporotic or nonosteoporotic (normal or osteopenic) using the WHO definition for DXA. Effectiveness was assessed in terms of the sensitivity and specificity of the referral decisions based on the QUS measurement. Local costs were estimated from health services and actual resource used. Cost-effectiveness was evaluated in terms of the expected cost per true positive osteoporotic case detected. Baseline prevalence of osteoporosis evaluated by DXA was 55.8%. The sensitivity and specificity for the diagnosis of osteoporosis by QUS using the optimal cutoff thresholds for the estimated heel BMD T-score were 97% and 94%, respectively. The average cost per osteoporotic case detected based on DXA measurement alone was 23.85 euros. The average cost per osteoporotic case detected using QUS as a prescreen was 22.00 euros. The incremental cost-effectiveness of DXA versus QUS was 114.00 euros per true positive case detected. Our results suggest that screening for osteoporosis with QUS while applying strict cufoff values in postmenopausal women of the general population is not substantially more cost-effective than DXA alone for the diagnosis of osteoporosis. However, the screening strategy with QUS may be an option in those circumstances where the diagnosis of osteoporosis is deficient because of the difficulty in accessing DXA equipment.

  15. Estimation of residential radon exposure and definition of Radon Priority Areas based on expected lung cancer incidence.

    PubMed

    Elío, J; Crowley, Q; Scanlon, R; Hodgson, J; Zgaga, L

    2018-05-01

    Radon is a naturally occurring gas, classified as a Class 1 human carcinogen, being the second most significant cause of lung cancer after tobacco smoking. A robust spatial definition of radon distribution in the built environment is therefore essential for understanding the relationship between radon exposure and its adverse health effects on the general population. Using Ireland as a case study, we present a methodology to estimate an average indoor radon concentration and calculate the expected radon-related lung cancer incidence. We use this approach to define Radon Priority Areas at the administrative level of Electoral Divisions (EDs). Geostatistical methods were applied to a data set of almost 32,000 indoor radon measurements, sampled in Ireland between 1992 and 2013. Average indoor radon concentrations by ED range from 21 to 338 Bq m -3 , corresponding to an effective dose ranging from 0.8 to 13.3 mSv y -1 respectively. Radon-related lung cancer incidence by ED was calculated using a dose-effect model giving between 15 and 239 cases per million people per year, depending on the ED. Based on these calculations, together with the population density, we estimate that of the approximately 2,300 lung cancer cases currently diagnosed in Ireland annually, about 280 may be directly linked to radon exposure. This figure does not account for the synergistic effect of radon exposure with other factors (e.g. tobacco smoking), so likely represents a minimum estimate. Our approach spatially defines areas with the expected highest incidence of radon-related lung cancer, even though indoor radon concentrations for these areas may be moderate or low. We therefore recommend that both indoor radon concentration and population density by small area are considered when establishing national radon action plans. Copyright © 2018 Elsevier Ltd. All rights reserved.

  16. [Regional Differences in the Care of Patients with Neovascular Age-Related Macular Degeneration, Based on the Non-Interventional OCEAN Study].

    PubMed

    Ziemssen, F; Hufenbach, U; Wiedon, A; Scheffler, M; Bertelmann, T

    2016-12-01

    Background: The main cause of blindness in the elderly in Germany is neovascular age-related macular degeneration (nAMD). In the non-interventional OCEAN study, data were prospectively collected on the routine clinical care of patients treated with the drug ranibizumab. Patients: As part of an interim analysis within the ongoing study (NCT02194803), stratification was performed by the 17 regions of the German associations of panel physicians and by areas of different population density. Only data were analysed for patients for whom the first treatment with ranibizumab was documented. Results: A total of 5,606 patients were documented. The present manuscript reviews 2,658 treatment-naive patients with nAMD, documented by 324 ophthalmologists. Most patients receiving an intravitreal injection were female (60 %). The average patient was aged 77.7 ± 8.2 years at study start. The great majority of patients had statutory health insurance (91 %). At baseline, fluorescein angiography (FLA) was performed for 72 % of patients, while optical coherence tomography (OCT) was carried out for 76 %. A combination of both was performed for 54 % of patients, varying regionally from 26 % (Saxony-Anhalt) to 100 % (Berlin). The average waiting time between the first examination and the first injection was 20.0 ± 18.5 days. With different statistical models (ANOVA adjusted, with/without interactions), significant effects on treatment delay were found for district type (population density), federal state and type of specialist. Conclusion: No major regional differences were observed in the demographic characteristics of the patient population. The main regional disparities in the care of nAMD patients were in the application of diagnostic methods and the waiting times between the first examination and the first drug administration. The regional variations in treatment delays could clearly influence the risk of worse functional outcome. Georg Thieme Verlag KG Stuttgart · New York.

  17. Optimum 3D Matrix Stiffness for Maintenance of Cancer Stem Cells Is Dependent on Tissue Origin of Cancer Cells

    PubMed Central

    Jabbari, Esmaiel; Sarvestani, Samaneh K.; Daneshian, Leily; Moeinzadeh, Seyedsina

    2015-01-01

    Introduction The growth and expression of cancer stem cells (CSCs) depend on many factors in the tumor microenvironment. The objective of this work was to investigate the effect of cancer cells’ tissue origin on the optimum matrix stiffness for CSC growth and marker expression in a model polyethylene glycol diacrylate (PEGDA) hydrogel without the interference of other factors in the microenvironment. Methods Human MCF7 and MDA-MB-231 breast carcinoma, HCT116 colorectal and AGS gastric carcinoma, and U2OS osteosarcoma cells were used. The cells were encapsulated in PEGDA gels with compressive moduli in the 2-70 kPa range and optimized cell seeding density of 0.6x106 cells/mL. Micropatterning was used to optimize the growth of encapsulated cells with respect to average tumorsphere size. The CSC sub-population of the encapsulated cells was characterized by cell number, tumorsphere size and number density, and mRNA expression of CSC markers. Results The optimum matrix stiffness for growth and marker expression of CSC sub-population of cancer cells was 5 kPa for breast MCF7 and MDA231, 25 kPa for colorectal HCT116 and gastric AGS, and 50 kPa for bone U2OS cells. Conjugation of a CD44 binding peptide to the gel stopped tumorsphere formation by cancer cells from different tissue origin. The expression of YAP/TAZ transcription factors by the encapsulated cancer cells was highest at the optimum stiffness indicating a link between the Hippo transducers and CSC growth. The optimum average tumorsphere size for CSC growth and marker expression was 50 μm. Conclusion The marker expression results suggest that the CSC sub-population of cancer cells resides within a niche with optimum stiffness which depends on the cancer cells’ tissue origin. PMID:26168187

  18. Energy density of the Scottish diet estimated from food purchase data: relationship with socio-economic position and dietary targets.

    PubMed

    Barton, Karen L; Wrieden, Wendy L; Sherriff, Andrea; Armstrong, Julie; Anderson, Annie S

    2014-07-14

    Frequent consumption of energy-dense foods has been strongly implicated in the global increase of obesity. The World Cancer Research Fund suggests a population-level energy density (ED) goal for diets of 523 kJ/100 g (125 kcal/100 g) as desirable for reducing weight gain and related co-morbidities. However, there is limited information about the ED of diets of contemporary populations. The aims of the present study were to (1) estimate the mean ED of the Scottish diet, (2) assess differences in ED over time by socio-economic position, by household (HH) composition and for HH meeting dietary targets for fat and fruit and vegetables, and (3) assess the relationship between ED and the consumption of foods and nutrients, which are indicative of diet quality. ED of the diet was estimated from food (including milk) from UK food purchase survey data. The average ED of the Scottish diet was estimated as 718 kJ/100 g with no change between the survey periods 2001 and 2009. Individuals living in the most deprived areas had a higher mean ED than those living in the least deprived areas (737 v. 696 kJ/100 g). Single-parent HH had the highest mean ED (765 kJ/100 g) of all the HH surveyed. The mean ED of HH achieving dietary targets for fat and fruit and vegetables was 576 kJ/100 g compared with 731 kJ/100 g for non-achievers. HH within the lowest quintile of ED were, on average, closest to meeting most dietary guidelines. Food purchase data can be used to monitor the quality of the diet in terms of dietary ED of the population and subgroups defined by an area-based measure of socio-economic status.

  19. Projection of invertebrate populations in the headwater streams of a temperate catchment under a changing climate.

    PubMed

    Nukazawa, Kei; Arai, Ryosuke; Kazama, So; Takemon, Yasuhiro

    2018-06-14

    Climate change places considerable stress on riverine ecosystems by altering flow regimes and increasing water temperature. This study evaluated how water temperature increases under climate change scenarios will affect stream invertebrates in pristine headwater streams. The studied headwater-stream sites were distributed within a temperate catchment of Japan and had similar hydraulic-geographical conditions, but were subject to varying temperature conditions due to altitudinal differences (100 to 850 m). We adopted eight general circulation models (GCMs) to project air temperature under conservative (RCP2.6), intermediate (RCP4.5), and extreme climate scenarios (RCP8.5) during the near (2031-2050) and far (2081-2100) future. Using the water temperature of headwater streams computed by a distributed hydrological-thermal model as a predictor variable, we projected the population density of stream invertebrates in the future scenarios based on generalized linear models. The mean decrease in the temporally averaged population density of Plecoptera was 61.3% among the GCMs, even under RCP2.6 in the near future, whereas density deteriorated even further (90.7%) under RCP8.5 in the far future. Trichoptera density was also projected to greatly deteriorate under RCP8.5 in the far future. We defined taxa that exhibited temperature-sensitive declines under climate change as cold stenotherms and found that most Plecoptera taxa were cold stenotherms in comparison to other orders. Specifically, the taxonomic families that only distribute in Palearctic realm (e.g., Megarcys ochracea and Scopura longa) were selectively assigned, suggesting that Plecoptera family with its restricted distribution in the Palearctic might be a sensitive indicator of climate change. Plecoptera and Trichoptera populations in the headwaters are expected/anticipated to decrease over the considerable geographical range of the catchment, even under the RCP2.6 in the near future. Given headwater invertebrates play important roles in streams, such as contributing to watershed productivity, our results provide useful information for managing streams at the catchment-level. Copyright © 2018 Elsevier B.V. All rights reserved.

  20. The Luminosity Function of Fermi-detected Flat-Spectrum Radio Quasars

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Ajello, M.; Shaw, M.S.; Romani, R.W.

    2012-04-16

    Fermi has provided the largest sample of {gamma}-ray selected blazars to date. In this work we use a complete sample of FSRQs detected during the first year of operation to determine the luminosity function (LF) and its evolution with cosmic time. The number density of FSRQs grows dramatically up to redshift {approx}0.5-2.0 and declines thereafter. The redshift of the peak in the density is luminosity dependent, with more luminous sources peaking at earlier times; thus the LF of {gamma}-ray FSRQs follows a luminosity-dependent density evolution similarly to that of radio-quiet AGN. Also using data from the Swift Burst Alert Telescopemore » we derive the average spectral energy distribution of FSRQs in the 10 keV-100GeV band and show that there is no correlation of the peak {gamma}-ray luminosity with {gamma}-ray peak frequency. The coupling of the SED and LF allows us to predict that the contribution of FSRQs to the Fermi isotropic {gamma}-ray background is 9.3{sub -1.0}{sup +1.6}% ({+-}3% systematic uncertainty) in the 0.1-100GeV band. Finally we determine the LF of unbeamed FSRQs, finding that FSRQs have an average Lorentz factor of {gamma} = 11.7{sub -2.2}{sup +3.3}, that most are seen within 5{sup o} of the jet axis, and that they represent only {approx}0.1% of the parent population.« less

  1. Flavin Charge Transfer Transitions Assist DNA Photolyase Electron Transfer

    NASA Astrophysics Data System (ADS)

    Skourtis, Spiros S.; Prytkova, Tatiana; Beratan, David N.

    2007-12-01

    This contribution describes molecular dynamics, semi-empirical and ab-initio studies of the primary photo-induced electron transfer reaction in DNA photolyase. DNA photolyases are FADH--containing proteins that repair UV-damaged DNA by photo-induced electron transfer. A DNA photolyase recognizes and binds to cyclobutatne pyrimidine dimer lesions of DNA. The protein repairs a bound lesion by transferring an electron to the lesion from FADH-, upon photo-excitation of FADH- with 350-450 nm light. We compute the lowest singlet excited states of FADH- in DNA photolyase using INDO/S configuration interaction, time-dependent density-functional, and time-dependent Hartree-Fock methods. The calculations identify the lowest singlet excited state of FADH- that is populated after photo-excitation and that acts as the electron donor. For this donor state we compute conformationally-averaged tunneling matrix elements to empty electron-acceptor states of a thymine dimer bound to photolyase. The conformational averaging involves different FADH--thymine dimer confromations obtained from molecular dynamics simulations of the solvated protein with a thymine dimer docked in its active site. The tunneling matrix element computations use INDO/S-level Green's function, energy splitting, and Generalized Mulliken-Hush methods. These calculations indicate that photo-excitation of FADH- causes a π→π* charge-transfer transition that shifts electron density to the side of the flavin isoalloxazine ring that is adjacent to the docked thymine dimer. This shift in electron density enhances the FADH--to-dimer electronic coupling, thus inducing rapid electron transfer.

  2. On the averaging area for incident power density for human exposure limits at frequencies over 6 GHz

    NASA Astrophysics Data System (ADS)

    Hashimoto, Yota; Hirata, Akimasa; Morimoto, Ryota; Aonuma, Shinta; Laakso, Ilkka; Jokela, Kari; Foster, Kenneth R.

    2017-04-01

    Incident power density is used as the dosimetric quantity to specify the restrictions on human exposure to electromagnetic fields at frequencies above 3 or 10 GHz in order to prevent excessive temperature elevation at the body surface. However, international standards and guidelines have different definitions for the size of the area over which the power density should be averaged. This study reports computational evaluation of the relationship between the size of the area over which incident power density is averaged and the local peak temperature elevation in a multi-layer model simulating a human body. Three wave sources are considered in the frequency range from 3 to 300 GHz: an ideal beam, a half-wave dipole antenna, and an antenna array. 1D analysis shows that averaging area of 20 mm  ×  20 mm is a good measure to correlate with the local peak temperature elevation when the field distribution is nearly uniform in that area. The averaging area is different from recommendations in the current international standards/guidelines, and not dependent on the frequency. For a non-uniform field distribution, such as a beam with small diameter, the incident power density should be compensated by multiplying a factor that can be derived from the ratio of the effective beam area to the averaging area. The findings in the present study suggest that the relationship obtained using the 1D approximation is applicable for deriving the relationship between the incident power density and the local temperature elevation.

  3. Urbanization and Mental Health in China: Linking the 2010 Population Census with a Cross-Sectional Survey.

    PubMed

    Chen, Juan; Chen, Shuo; Landry, Pierre F

    2015-07-31

    Along with the rapid urbanization in China, the state of mental health also receives growing attention. Empirical measures, however, have not been developed to assess the impact of urbanization on mental health and the dramatic spatial variations. Innovatively linking the 2010 Chinese Population Census with a 2011 national survey of urban residents, we first assess the impact of urbanization on depressive symptoms measured by the Center of Epidemiological Studies Depression Scale (CES-D) of 1288 survey respondents. We then retrieve county-level characteristics from the 2010 Chinese Population Census that match the individual characteristics in the survey, so as to create a profile of the "average person" for each of the 2869 counties or city districts, and predict a county-specific CES-D score. We use this county-specific CES-D score to compute the CES-D score for the urban population at the prefectural level, and to demonstrate the dramatic spatial variations in urbanization and mental health across China: highly populated cities along the eastern coast such as Shenyang and Shanghai show high CES-D scores, as do cities in western China with high population density and a high proportion of educated ethnic minorities.

  4. Effects of population density on corticosterone levels of prairie voles in the field.

    PubMed

    Blondel, Dimitri V; Wallace, Gerard N; Calderone, Stefanie; Gorinshteyn, Marija; St Mary, Colette M; Phelps, Steven M

    2016-01-01

    High population density is often associated with increased levels of stress-related hormones, such as corticosterone (CORT). Prairie voles (Microtus ochrogaster) are a socially monogamous species known for their large population density fluctuations in the wild. Although CORT influences the social behavior of prairie voles in the lab, the effect of population density on CORT has not previously been quantified in this species in the field. We validated a non-invasive hormone assay for measuring CORT metabolites in prairie vole feces. We then used semi-natural enclosures to experimentally manipulate population density, and measured density effects on male space use and fecal CORT levels. Our enclosures generated patterns of space use and social interaction that were consistent with previous prairie vole field studies. Contrary to the positive relationship between CORT and density typical of other taxa, we found that lower population densities (80 animals/ha) produced higher fecal CORT than higher densities (240/ha). Combined with prior work in the lab and field, the data suggest that high prairie vole population densities indicate favorable environments, perhaps through reduced predation risk. Lastly, we found that field animals had lower fecal CORT levels than laboratory-living animals. The data emphasize the usefulness of prairie voles as models for integrating ecological, evolutionary, and mechanistic questions in social behavior. Copyright © 2015 Elsevier Inc. All rights reserved.

  5. Derivation of Poisson and Nernst-Planck equations in a bath and channel from a molecular model.

    PubMed

    Schuss, Z; Nadler, B; Eisenberg, R S

    2001-09-01

    Permeation of ions from one electrolytic solution to another, through a protein channel, is a biological process of considerable importance. Permeation occurs on a time scale of micro- to milliseconds, far longer than the femtosecond time scales of atomic motion. Direct simulations of atomic dynamics are not yet possible for such long-time scales; thus, averaging is unavoidable. The question is what and how to average. In this paper, we average a Langevin model of ionic motion in a bulk solution and protein channel. The main result is a coupled system of averaged Poisson and Nernst-Planck equations (CPNP) involving conditional and unconditional charge densities and conditional potentials. The resulting NP equations contain the averaged force on a single ion, which is the sum of two components. The first component is the gradient of a conditional electric potential that is the solution of Poisson's equation with conditional and permanent charge densities and boundary conditions of the applied voltage. The second component is the self-induced force on an ion due to surface charges induced only by that ion at dielectric interfaces. The ion induces surface polarization charge that exerts a significant force on the ion itself, not present in earlier PNP equations. The proposed CPNP system is not complete, however, because the electric potential satisfies Poisson's equation with conditional charge densities, conditioned on the location of an ion, while the NP equations contain unconditional densities. The conditional densities are closely related to the well-studied pair-correlation functions of equilibrium statistical mechanics. We examine a specific closure relation, which on the one hand replaces the conditional charge densities by the unconditional ones in the Poisson equation, and on the other hand replaces the self-induced force in the NP equation by an effective self-induced force. This effective self-induced force is nearly zero in the baths but is approximately equal to the self-induced force in and near the channel. The charge densities in the NP equations are interpreted as time averages over long times of the motion of a quasiparticle that diffuses with the same diffusion coefficient as that of a real ion, but is driven by the averaged force. In this way, continuum equations with averaged charge densities and mean-fields can be used to describe permeation through a protein channel.

  6. Are Barroom and Neighborhood Characteristics Independently Related to Local-Area Assaults?

    PubMed

    Morrison, Christopher; Mair, Christina F; Lee, Juliet P; Gruenewald, Paul J

    2015-12-01

    Two separate but complementary literatures examine bar-related violence: one has focused on barroom features, and the other has focused on features of neighborhoods near bars. This study unifies these 2 perspectives using a microenvironmental approach. In a purposive sample of 65 bars in 4 California cities, we used premise assessments to characterize the physical, social, and economic environments of barrooms (e.g., patron count, average pace of drinking, and restaurant service); and a combination of systematic social observation, census, and alcohol license data to characterize the neighborhoods in which they were located (e.g., physical disorder, alcohol outlet density, and median household income). Hierarchical Poisson models then assessed relationships between these features and counts of police-reported assaults within buffer areas around bars. Aspects of both barroom environments (more patrons, more dancing, and louder music) and neighborhood environments (greater bar density, greater physical disorder, lower population density, and lower income) were independently related to increased incidence of assaults. Preventive intervention to reduce bar-area violence may be directed at both bar environments (e.g., limiting the number of patrons) and neighborhood environments (e.g., limiting outlet density). Copyright © 2015 by the Research Society on Alcoholism.

  7. 42 CFR 505.5 - Loan criteria.

    Code of Federal Regulations, 2012 CFR

    2012-10-01

    ... that meet the following criteria: (1) The hospital is located in a State that, based on population density, is defined as a rural State. A rural State is one of ten States with the lowest population... prioritized beginning with the State with the lowest population density. Population density is determined...

  8. 42 CFR 505.5 - Loan criteria.

    Code of Federal Regulations, 2013 CFR

    2013-10-01

    ... that meet the following criteria: (1) The hospital is located in a State that, based on population density, is defined as a rural State. A rural State is one of ten States with the lowest population... prioritized beginning with the State with the lowest population density. Population density is determined...

  9. 42 CFR 505.5 - Loan criteria.

    Code of Federal Regulations, 2014 CFR

    2014-10-01

    ... that meet the following criteria: (1) The hospital is located in a State that, based on population density, is defined as a rural State. A rural State is one of ten States with the lowest population... prioritized beginning with the State with the lowest population density. Population density is determined...

  10. Size-resolved measurements of mixing state and cloud-nucleating ability of aerosols in Nanjing, China

    NASA Astrophysics Data System (ADS)

    Ma, Yan; Li, Shizheng; Zheng, Jun; Khalizov, Alexei; Wang, Xing; Wang, Zhen; Zhou, Yaoyao

    2017-09-01

    An integrated aerosol analytical system was deployed in Nanjing, a megacity in the Yangtze River Delta, to measure size-resolved aerosol mixing states, effective densities, cloud condensation nucleus (CCN) activities, and chemical composition in August 2013. It was found that aerosols were predominantly internally mixed. The average effective densities were 1.38 ± 0.09, 1.48 ± 0.08, and 1.53 ± 0.07 g cm-3 for 50, 80, and 120 nm particles, respectively. Although black carbon (BC) represented only 0.3%, 1.6%, and 3.3% of the particle mass, on average, it was present in 7%, 38%, and 47% of the total particle number concentration at 50, 80, and 120 nm, respectively, indicating that BC particles may contribute significantly to the total atmospheric aerosol population. Externally mixed BC was only occasionally observed with an effective density of 0.67-0.97 g cm-3. Aerosols sampled generally exhibited a relatively high CCN activity and hygroscopicity (κ = 0.35 ± 0.13). Both newly formed particles and freshly emitted BC particles were observed to age rapidly from photochemical processes, with a significant enhancement in the particle CCN activity and an increase in the effective density. Aerosols influenced by four different air masses presented similar CCN activation, indicating that CCN activation would be primarily dependent on the particle size rather than the particle origin (and hence original composition). Our results suggest that under highly active photochemical conditions as encountered in this study, particles from both local sources and regional transport can be rapidly converted into efficient CCN by photochemical aging, thereby making important contributions to the atmospheric CCN budget and exerting profound implications on aerosol indirect climate forcing.

  11. Similar Spectral Power Densities Within the Schumann Resonance and a Large Population of Quantitative Electroencephalographic Profiles: Supportive Evidence for Koenig and Pobachenko

    PubMed Central

    Saroka, Kevin S.; Vares, David E.; Persinger, Michael A.

    2016-01-01

    In 1954 and 1960 Koenig and his colleagues described the remarkable similarities of spectral power density profiles and patterns between the earth-ionosphere resonance and human brain activity which also share magnitudes for both electric field (mV/m) and magnetic field (pT) components. In 2006 Pobachenko and colleagues reported real time coherence between variations in the Schumann and brain activity spectra within the 6–16 Hz band for a small sample. We examined the ratios of the average potential differences (~3 μV) obtained by whole brain quantitative electroencephalography (QEEG) between rostral-caudal and left-right (hemispheric) comparisons of 238 measurements from 184 individuals over a 3.5 year period. Spectral densities for the rostral-caudal axis revealed a powerful peak at 10.25 Hz while the left-right peak was 1.95 Hz with beat-differences of ~7.5 to 8 Hz. When global cerebral measures were employed, the first (7–8 Hz), second (13–14 Hz) and third (19–20 Hz) harmonics of the Schumann resonances were discernable in averaged QEEG profiles in some but not all participants. The intensity of the endogenous Schumann resonance was related to the ‘best-of-fitness’ of the traditional 4-class microstate model. Additional measurements demonstrated real-time coherence for durations approximating microstates in spectral power density variations between Schumann frequencies measured in Sudbury, Canada and Cumiana, Italy with the QEEGs of local subjects. Our results confirm the measurements reported by earlier researchers that demonstrated unexpected similarities in the spectral patterns and strengths of electromagnetic fields generated by the human brain and the earth-ionospheric cavity. PMID:26785376

  12. Similar Spectral Power Densities Within the Schumann Resonance and a Large Population of Quantitative Electroencephalographic Profiles: Supportive Evidence for Koenig and Pobachenko.

    PubMed

    Saroka, Kevin S; Vares, David E; Persinger, Michael A

    2016-01-01

    In 1954 and 1960 Koenig and his colleagues described the remarkable similarities of spectral power density profiles and patterns between the earth-ionosphere resonance and human brain activity which also share magnitudes for both electric field (mV/m) and magnetic field (pT) components. In 2006 Pobachenko and colleagues reported real time coherence between variations in the Schumann and brain activity spectra within the 6-16 Hz band for a small sample. We examined the ratios of the average potential differences (~3 μV) obtained by whole brain quantitative electroencephalography (QEEG) between rostral-caudal and left-right (hemispheric) comparisons of 238 measurements from 184 individuals over a 3.5 year period. Spectral densities for the rostral-caudal axis revealed a powerful peak at 10.25 Hz while the left-right peak was 1.95 Hz with beat-differences of ~7.5 to 8 Hz. When global cerebral measures were employed, the first (7-8 Hz), second (13-14 Hz) and third (19-20 Hz) harmonics of the Schumann resonances were discernable in averaged QEEG profiles in some but not all participants. The intensity of the endogenous Schumann resonance was related to the 'best-of-fitness' of the traditional 4-class microstate model. Additional measurements demonstrated real-time coherence for durations approximating microstates in spectral power density variations between Schumann frequencies measured in Sudbury, Canada and Cumiana, Italy with the QEEGs of local subjects. Our results confirm the measurements reported by earlier researchers that demonstrated unexpected similarities in the spectral patterns and strengths of electromagnetic fields generated by the human brain and the earth-ionospheric cavity.

  13. Nearshore distribution and abundance of Dungeness crabs in Glacier Bay National Park, Alaska

    USGS Publications Warehouse

    O'Clair, Charles E.; Freese, J. Lincoln; Stone, Robert P.; Shirley, Thomas C.; Leder, Erica H.; Taggart, S. James; Kruse, Gordon H.; Engstrom, Daniel R.

    1995-01-01

    As part of an ongoing, multi-agency study to determine the effects of closure of the commercial fishery for Dungeness crabs, Cancer magister, on crab population structure we examined patterns of distribution and abundance of crabs in nearshore habitats at five locations in and near Glacier Bay National Park. Sampling was conducted in April and September 1992 and April 1993 prior to the anticipated closure of the fishery in the park. Divers censused crabs by sex and reproductive state (ovigerous/nonovigerous females) along belt transects (2m x 100m) laid perpendicular to shore in the depth range 0 m (mean lower low water) to 18 m.Preliminary results from the first three sampling periods revealed that the average densities of Dungeness crabs at the five locations ranged from 78 to 2012 crabs/ha. Crab densities differed between populations depending on sex, reproductive state of females and sampling period. Male crabs showed reduced densities at Gustavus Flats in April 1992 (P<0.01) and 1993 (P<0.001). Ovigerous females had greater density at Bartlett Cove in April 1993 (P<0.001). Sex ratios were frequently skewed toward females. At Bartlett Cove and Gustavus Flats females outnumbered males in April 1992 and 1993 (P<0.001). Most of the females at Bartlett Cove and Gustavus Flats in April 1992 and 1993 were ovigerous (P-0.001). Males tended to occupy greater depths than females in April 1992 (P<0.05) but not April 1993 (P-005). The mean depth of males shifted from deeper to shallower water between April and September 1992 (P<0.001). The depth distribution of ovigerous crabs did not differ from that of nonovigerous female crabs. Future research prior to the anticipated closure of the commercial Dungeness crab fishery in Glacier Bay will include a tagging study to determine the extent of crab movement and further study of the temporal as well as the spatial variability observed in the structure of these populations.

  14. Determinants of pika population density vs. occupancy in the Southern Rocky Mountains.

    PubMed

    Erb, Liesl P; Ray, Chris; Guralnick, Robert

    2014-04-01

    Species distributions are responding rapidly to global change. While correlative studies of local extinction have been vital to understanding the ecological impacts of global change, more mechanistic lines of inquiry are needed for enhanced forecasting. The current study assesses whether the predictors of local extinction also explain population density for a species apparently impacted by climate change. We tested a suite of climatic and habitat metrics as predictors of American pika (Ochotona princeps) relative population density in the Southern Rocky Mountains, USA. Population density was indexed as the density of pika latrine sites. Negative binomial regression and AICc showed that the best predictors of pika latrine density were patch area followed by two measures of vegetation quality: the diversity and relative cover of forbs. In contrast with previous studies of habitat occupancy in the Southern Rockies, climatic factors were not among the top predictors of latrine density. Populations may be buffered from decline and ultimately from extirpation at sites with high-quality vegetation. Conversely, populations at highest risk for declining density and extirpation are likely to be those in sites with poor-quality vegetation.

  15. Thermal and energetic constraints on ectotherm abundance: A global test using lizards

    USGS Publications Warehouse

    Buckley, L.B.; Rodda, G.H.; Jetz, W.

    2008-01-01

    Population densities of birds and mammals have been shown to decrease with body mass at approximately the same rate as metabolic rates increase, indicating that energetic needs constrain endotherm population densities. In ectotherms, the exponential increase of metabolic rate with body temperature suggests that environmental temperature may additionally constrain population densities. Here we test simple bioenergetic models for an ecologically important group of ectothermic vertebrates by examining 483 lizard populations. We find that lizard population densities decrease as a power law of body mass with a slope approximately inverse to the slope of the relationship between metabolic rates and body mass. Energy availability should limit population densities. As predicted, environmental productivity has a positive effect on lizard density, strengthening the relationship between lizard density and body mass. In contrast, the effect of environmental temperature is at most weak due to behavioral thermoregulation, thermal evolution, or the temperature dependence of ectotherm performance. Our results provide initial insights into how energy needs and availability differentially constrain ectotherm and endotherm density across broad spatial scales. ?? 2008 by the Ecological Society of America.

  16. Thermal and energetic constraints on ectotherm abundance: a global test using lizards.

    PubMed

    Buckley, Lauren B; Rodda, Gordon H; Jetz, Walter

    2008-01-01

    Population densities of birds and mammals have been shown to decrease with body mass at approximately the same rate as metabolic rates increase, indicating that energetic needs constrain endotherm population densities. In ectotherms, the exponential increase of metabolic rate with body temperature suggests that environmental temperature may additionally constrain population densities. Here we test simple bioenergetic models for an ecologically important group of ectothermic vertebrates by examining 483 lizard populations. We find that lizard population densities decrease as a power law of body mass with a slope approximately inverse to the slope of the relationship between metabolic rates and body mass. Energy availability should limit population densities. As predicted, environmental productivity has a positive effect on lizard density, strengthening the relationship between lizard density and body mass. In contrast, the effect of environmental temperature is at most weak due to behavioral thermoregulation, thermal evolution, or the temperature dependence of ectotherm performance. Our results provide initial insights into how energy needs and availability differentially constrain ectotherm and endotherm density across broad spatial scales.

  17. Probability of detecting band-tailed pigeons during call-broadcast versus auditory surveys

    USGS Publications Warehouse

    Kirkpatrick, C.; Conway, C.J.; Hughes, K.M.; Devos, J.C.

    2007-01-01

    Estimates of population trend for the interior subspecies of band-tailed pigeon (Patagioenas fasciata fasciata) are not available because no standardized survey method exists for monitoring the interior subspecies. We evaluated 2 potential band-tailed pigeon survey methods (auditory and call-broadcast surveys) from 2002 to 2004 in 5 mountain ranges in southern Arizona, USA, and in mixed-conifer forest throughout the state. Both auditory and call-broadcast surveys produced low numbers of cooing pigeons detected per survey route (x?? ??? 0.67) and had relatively high temporal variance in average number of cooing pigeons detected during replicate surveys (CV ??? 161%). However, compared to auditory surveys, use of call-broadcast increased 1) the percentage of replicate surveys on which ???1 cooing pigeon was detected by an average of 16%, and 2) the number of cooing pigeons detected per survey route by an average of 29%, with this difference being greatest during the first 45 minutes of the morning survey period. Moreover, probability of detecting a cooing pigeon was 27% greater during call-broadcast (0.80) versus auditory (0.63) surveys. We found that cooing pigeons were most common in mixed-conifer forest in southern Arizona and density of male pigeons in mixed-conifer forest throughout the state averaged 0.004 (SE = 0.001) pigeons/ha. Our results are the first to show that call-broadcast increases the probability of detecting band-tailed pigeons (or any species of Columbidae) during surveys. Call-broadcast surveys may provide a useful method for monitoring populations of the interior subspecies of band-tailed pigeon in areas where other survey methods are inappropriate.

  18. Face Value: Towards Robust Estimates of Snow Leopard Densities.

    PubMed

    Alexander, Justine S; Gopalaswamy, Arjun M; Shi, Kun; Riordan, Philip

    2015-01-01

    When densities of large carnivores fall below certain thresholds, dramatic ecological effects can follow, leading to oversimplified ecosystems. Understanding the population status of such species remains a major challenge as they occur in low densities and their ranges are wide. This paper describes the use of non-invasive data collection techniques combined with recent spatial capture-recapture methods to estimate the density of snow leopards Panthera uncia. It also investigates the influence of environmental and human activity indicators on their spatial distribution. A total of 60 camera traps were systematically set up during a three-month period over a 480 km2 study area in Qilianshan National Nature Reserve, Gansu Province, China. We recorded 76 separate snow leopard captures over 2,906 trap-days, representing an average capture success of 2.62 captures/100 trap-days. We identified a total number of 20 unique individuals from photographs and estimated snow leopard density at 3.31 (SE = 1.01) individuals per 100 km2. Results of our simulation exercise indicate that our estimates from the Spatial Capture Recapture models were not optimal to respect to bias and precision (RMSEs for density parameters less or equal to 0.87). Our results underline the critical challenge in achieving sufficient sample sizes of snow leopard captures and recaptures. Possible performance improvements are discussed, principally by optimising effective camera capture and photographic data quality.

  19. Face Value: Towards Robust Estimates of Snow Leopard Densities

    PubMed Central

    2015-01-01

    When densities of large carnivores fall below certain thresholds, dramatic ecological effects can follow, leading to oversimplified ecosystems. Understanding the population status of such species remains a major challenge as they occur in low densities and their ranges are wide. This paper describes the use of non-invasive data collection techniques combined with recent spatial capture-recapture methods to estimate the density of snow leopards Panthera uncia. It also investigates the influence of environmental and human activity indicators on their spatial distribution. A total of 60 camera traps were systematically set up during a three-month period over a 480 km2 study area in Qilianshan National Nature Reserve, Gansu Province, China. We recorded 76 separate snow leopard captures over 2,906 trap-days, representing an average capture success of 2.62 captures/100 trap-days. We identified a total number of 20 unique individuals from photographs and estimated snow leopard density at 3.31 (SE = 1.01) individuals per 100 km2. Results of our simulation exercise indicate that our estimates from the Spatial Capture Recapture models were not optimal to respect to bias and precision (RMSEs for density parameters less or equal to 0.87). Our results underline the critical challenge in achieving sufficient sample sizes of snow leopard captures and recaptures. Possible performance improvements are discussed, principally by optimising effective camera capture and photographic data quality. PMID:26322682

  20. The contributions of breast density and common genetic variation to breast cancer risk.

    PubMed

    Vachon, Celine M; Pankratz, V Shane; Scott, Christopher G; Haeberle, Lothar; Ziv, Elad; Jensen, Matthew R; Brandt, Kathleen R; Whaley, Dana H; Olson, Janet E; Heusinger, Katharina; Hack, Carolin C; Jud, Sebastian M; Beckmann, Matthias W; Schulz-Wendtland, Ruediger; Tice, Jeffrey A; Norman, Aaron D; Cunningham, Julie M; Purrington, Kristen S; Easton, Douglas F; Sellers, Thomas A; Kerlikowske, Karla; Fasching, Peter A; Couch, Fergus J

    2015-05-01

    We evaluated whether a 76-locus polygenic risk score (PRS) and Breast Imaging Reporting and Data System (BI-RADS) breast density were independent risk factors within three studies (1643 case patients, 2397 control patients) using logistic regression models. We incorporated the PRS odds ratio (OR) into the Breast Cancer Surveillance Consortium (BCSC) risk-prediction model while accounting for its attributable risk and compared five-year absolute risk predictions between models using area under the curve (AUC) statistics. All statistical tests were two-sided. BI-RADS density and PRS were independent risk factors across all three studies (P interaction = .23). Relative to those with scattered fibroglandular densities and average PRS (2(nd) quartile), women with extreme density and highest quartile PRS had 2.7-fold (95% confidence interval [CI] = 1.74 to 4.12) increased risk, while those with low density and PRS had reduced risk (OR = 0.30, 95% CI = 0.18 to 0.51). PRS added independent information (P < .001) to the BCSC model and improved discriminatory accuracy from AUC = 0.66 to AUC = 0.69. Although the BCSC-PRS model was well calibrated in case-control data, independent cohort data are needed to test calibration in the general population. © The Author 2015. Published by Oxford University Press. All rights reserved. For Permissions, please e-mail: journals.permissions@oup.com.

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