Phototropic solar tracking in sunflower plants: an integrative perspective.

PubMed

Kutschera, Ulrich; Briggs, Winslow R

2016-01-01

One of the best-known plant movements, phototropic solar tracking in sunflower (Helianthus annuus), has not yet been fully characterized. Two questions are still a matter of debate. (1) Is the adaptive significance solely an optimization of photosynthesis via the exposure of the leaves to the sun? (2) Is shade avoidance involved in this process? In this study, these concepts are discussed from a historical perspective and novel insights are provided. Results from the primary literature on heliotropic growth movements led to the conclusion that these responses cease before anthesis, so that the flowering heads point to the East. Based on observations on 10-week-old plants, the diurnal East-West oscillations of the upper fifth of the growing stem and leaves in relation to the position of the sun (inclusive of nocturnal re-orientation) were documented, and photon fluence rates on the leaf surfaces on clear, cloudy and rainy days were determined. In addition, the light-response curve of net CO2 assimilation was determined on the upper leaves of the same batch of plants, and evidence for the occurrence of shade-avoidance responses in growing sunflower plants is summarized. Only elongating, vegetative sunflower shoots and the upper leaves perform phototropic solar tracking. Photon fluence response and CO2 assimilation measurements cast doubt on the 'photosynthesis-optimization hypothesis' as the sole explanation for the evolution of these plant movements. We suggest that the shade-avoidance response, which maximizes light-driven CO2 assimilation, plays a major role in solar tracking populations of competing sunflower plants, and an integrative scheme of these growth movements is provided. © The Author 2015. Published by Oxford University Press on behalf of the Annals of Botany Company. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

Remote sensing of future competitors: Impacts on plant defenses

PubMed Central

Izaguirre, Miriam M.; Mazza, Carlos A.; Biondini, Mariela; Baldwin, Ian T.; Ballaré, Carlos L.

2006-01-01

Far-red radiation (FR) reflected by green tissues is a key signal that plants use to detect the proximity of future competitors. Perception of increased levels of FR elicits a suite of responses collectively known as the shade-avoidance syndrome, which includes increased stem elongation, production of erect leaves, and reduced lateral branching. These responses improve the access to light for plants that occur in crowded populations. Responses to the proximity of competitors are known to affect the susceptibility to disease and predation in several organisms, including social animals. However, the impacts of warning signals of competition on the expression of defenses have not been explicitly investigated in plants. In the experiments reported here, we show that reflected FR induced a dramatic down-regulation of chemical defenses in wild tobacco (Nicotiana longiflora). FR altered the expression of several defense-related genes, inhibited the accumulation of herbivore-induced phenolic compounds, and augmented the performance of the specialist herbivore Manduca sexta. Complementary studies with tomato suggested that the effects of FR on defenses are mediated by the photoreceptor phytochrome B. The central implication of these results is that shade-intolerant species such as wild tobacco and tomato activate functional changes that affect their ability to cope with herbivore attack in response to phytochrome signals of future competition, even in the absence of real competition for resources. These findings suggest that competition overshadowed herbivory during the evolution of this group of species and add a new axis to the definition of the shade-avoidance syndrome. PMID:16632610

Initial characterization of shade avoidance response suggests functional diversity between Populus phytochrome B genes.

DOE Office of Scientific and Technical Information (OSTI.GOV)

Karve, Abhijit A; Weston, David; Jawdy, Sara

2012-01-01

Shade avoidance signaling in higher plants involves perception of the incident red/far-red (R/FR) light by phytochromes and the modulation of downstream transcriptional networks to regulate developmental plasticity in relation to heterogeneous light environments. In this study, we characterized the expression and functional features of Populus phytochrome (PHY) gene family as well as the transcriptional responses of Populus to the changes in R/FR light. Expression data indicated that PHYA is the predominant PHY in the dark grown Populus seedling whereas PHYBs are most abundant in mature tissue types. Out of three Populus PHYs, PHYA is light labile and localized to cytosolmore » in dark whereas both PHYB1 and PHYB2 are light stable and are localized to nucleus in mesophyll protoplasts. When expressed in Arabidopsis, PHYB1 rescued Arabidopsis phyB mutant phenotype whereas PHYB2 did not, suggesting functional diversification between these two gene family members. However, phenotypes of transgenic Populus lines with altered expression of PHYB1, PHYB2 or both and the expression of candidate shade response genes in these transgenic lines suggest that PHYB1 and PHYB2 may have distinct yet overlapping functions. The RNAseq results and analysis of Populus exposed to enriched-FR light indicate that genes associated in cell wall modification and brassinosteroid signaling were induced under far red light. Overall our data indicate that Populus transcriptional responses are at least partially conserved with Arabidopsis.« less

Perception of low red:far-red ratio compromises both salicylic acid- and jasmonic acid-dependent pathogen defences in Arabidopsis.

PubMed

de Wit, Mieke; Spoel, Steven H; Sanchez-Perez, Gabino F; Gommers, Charlotte M M; Pieterse, Corné M J; Voesenek, Laurentius A C J; Pierik, Ronald

2013-07-01

In dense stands of plants, such as agricultural monocultures, plants are exposed simultaneously to competition for light and other stresses such as pathogen infection. Here, we show that both salicylic acid (SA)-dependent and jasmonic acid (JA)-dependent disease resistance is inhibited by a simultaneously reduced red:far-red light ratio (R:FR), the early warning signal for plant competition. Conversely, SA- and JA-dependent induced defences did not affect shade-avoidance responses to low R:FR. Reduced pathogen resistance by low R:FR was accompanied by a strong reduction in the regulation of JA- and SA-responsive genes. The severe inhibition of SA-responsive transcription in low R:FR appeared to be brought about by the repression of SA-inducible kinases. Phosphorylation of the SA-responsive transcription co-activator NPR1, which is required for full induction of SA-responsive transcription, was indeed reduced and may thus play a role in the suppression of SA-mediated defences by low R:FR-mediated phytochrome inactivation. Our results indicate that foraging for light through the shade-avoidance response is prioritised over plant immune responses when plants are simultaneously challenged with competition and pathogen attack. © 2013 The Authors The Plant Journal © 2013 John Wiley & Sons Ltd.

Differential response of Scots pine seedlings to variable intensity and ratio of red and far-red light.

PubMed

Razzak, Abdur; Ranade, Sonali Sachin; Strand, Åsa; García-Gil, M R

2017-08-01

We investigated the response to increasing intensity of red (R) and far-R (FR) light and to a decrease in R:FR ratio in Pinus sylvestris L. (Scots pine) seedling. The results showed that FR high-irradiance response for hypocotyl elongation may be present in Scots pine and that this response is enhanced by increasing light intensity. However, both hypocotyl inhibition and pigment accumulation were more strongly affected by the R light compared with FR light. This is in contrast to previous reports in Arabidopsis thaliana (L.) Heynh. In the angiosperm, A. thaliana R light shows an overall milder effect on inhibition of hypocotyl elongation and on pigment biosynthesis compared with FR suggesting conifers and angiosperms respond very differently to the different light regimes. Scots pine shade avoidance syndrome with longer hypocotyls, shorter cotyledons and lower chlorophyll content in response to shade conditions resembles the response observed in A. thaliana. However, anthocyanin accumulation increased with shade in Scots pine, which again differs from what is known in angiosperms. Overall, the response of seedling development and physiology to R and FR light in Scots pine indicates that the regulatory mechanism for light response may differ between gymnosperms and angiosperms. © 2017 John Wiley & Sons Ltd.

Adolescents’ use of purpose built shade in secondary schools: cluster randomised controlled trial

PubMed Central

White, Vanessa; Wakefield, Melanie A; Jamsen, Kris M; White, Victoria; Livingston, Patricia M; English, Dallas R; Simpson, Julie A

2009-01-01

Objective To examine whether students use or avoid newly shaded areas created by shade sails installed at schools. Design Cluster randomised controlled trial with secondary schools as the unit of randomisation. Setting 51 secondary schools with limited available shade, in Australia, assessed over two spring and summer terms. Participants Students outside at lunch times. Intervention Purpose built shade sails were installed in winter 2005 at full sun study sites to increase available shade for students in the school grounds. Main outcome measure Mean number of students using the primary study sites during weekly observations at lunch time. Results Over the study period the mean change in students using the primary study site from pre-test to post-test was 2.63 (95% confidence interval 0.87 to 4.39) students in intervention schools and −0.03 (−1.16 to 1.09) students in control schools. The difference in mean change between groups was 2.67 (0.65 to 4.68) students (P=0.011). Conclusions Students used rather than avoided newly shaded areas provided by purpose built shade sails at secondary schools in this trial, suggesting a practical means of reducing adolescents’ exposure to ultraviolet radiation. Trial registration Exempt. PMID:19223344

Sprinklers and shade cool cows and reduce insect-avoidance behavior in pasture-based dairy systems.

PubMed

Kendall, P E; Verkerk, G A; Webster, J R; Tucker, C B

2007-08-01

The body temperature of dairy cows in pastoral systems during summer reaches a peak during and following the p.m. milking. Shade and sprinklers can be used separately or in combination at the milking parlor to reduce heat load. Farmers anecdotally report that the use of sprinklers reduces irritation from insects that occurs while cows are waiting for milking. Once daily, we assessed the effectiveness of short-term exposure to shade and sprinklers for cooling cows [via respiration rate and body (vaginal) temperature] and reducing insect-avoidance behaviors before the p.m. milking in a pasture-based dairy system. Head position was measured as an indicator of whether cattle were avoiding water from the sprinklers. Forty-eight Holstein-Friesian dairy cows were divided into 12 groups (4 cows per group, n = 3 groups/treatment) and were exposed to 1 of 4 treatments for 90 min before the p.m. milking: 1) shade, 2) sprinklers, 3) shade and sprinklers, or 4) uncooled control. Respiration rate was reduced by 30% with shade alone compared with controls [54 vs. 78 +/- 2.3 ( +/- SED) breaths/min, respectively]. Sprinklers alone (30 +/- 2.3 breaths/min) and the combined effects of shade and sprinklers (24 +/- 2.3 breaths/min) reduced the respiration rate by 60 and 67%, respectively, compared with controls. Shaded cows had lower body temperatures during the 90-min treatment period compared with controls (shade: 38.6 degrees C; shade and sprinklers: 38.6 degrees C; control: 38.9 +/- 0.09 degrees C). The decrease in body temperature of cows under sprinklers was more marked than for shade alone and remained lower for at least 4 h after milking (sprinklers: 38.7 degrees C; shade and sprinklers: 38.6 degrees C; shade: 38.9 degrees C; control: 39.2 +/- 0.10 degrees C). The sprinkler treatment reduced the number of tail flicks (control: 12.6 vs. sprinklers: 6.6 +/- 2.4 flicks/min) and hoof stamps (control: 4.4 vs. sprinkler: 2.2 +/- 0.5 stamps/min). Cows exposed to sprinklers spent more time with their heads lowered compared with cows in the shaded and control treatments. The reductions in body temperature and respiration rate attributable to shade and sprinklers were greatest when the temperature-humidity index and heat-load index were > or = 69 and 77, respectively, and cows benefited from cooling when these levels were exceeded.

Green light in photomorphogenic development

NASA Astrophysics Data System (ADS)

Maruhnich, Stefanie Anne

Light quality, quantity, and duration provide essential environmental cues that shape plant growth and development. Over the last century, researchers have worked to discover how plants sense, integrate, and respond to red, blue, and far-red light. Green light is often considered a “benign” wavelength with little to no effect in plant development. However, sparse experiments in the literature demonstrate that green effects are often counterintuitive to normal light responses and oppose red- and blue-light-induced responses. Green light effects on plant growth and development are described here through the use of custom, tunable LED, light-emitting diode, chambers. These light sources allow for specific light qualities and quantities to be administered. The effects of green wavebands were assessed when red and blue photomorphogenic systems were active to answer the question: Are the effects of an inhibitor (green light) more evident in the presence of inducers (red and blue light)? In seedlings, supplemental green light increased hypocotyl elongation opposite to classical inhibition of hypocotyl elongation associated with growth in light and induced by red and blue wavebands. Results indicate that added green light induced a reversion of light-grown phenotypes. In mature plants, supplemental green light induced phenotypes typical of the shade-avoidance syndrome, including elongated petioles, smaller leaf areas, and leaf hyponasty. These responses are typical of lower-light conditions or far-red enriched environments. Contrary to far-red-light-induced shade-avoidance, data indicate green delays flowering. In Arabidopsis and strawberry plants, anthocyanin levels also decreased when green light was added to red and blue light treatments, which is again opposite to normal light-induced phenotypes. Photoreceptor mutants were tested and indicate green light effects in early development are cryptochromedependent. However, green-light-induced shade-avoidance responses were cryptochrome-independent. A candidate gene approach was used to identify other elements required for green light sensing and/or response. Defects in some green light responses were observed for mutants in CCD8/Max4, a putative carotenoid cleavage enzyme with high sequence similarity to a critical enzyme in animal vision. These data support a role for green light in plant development which opposes normal light-induced responses and indicate the existence of at least two green light sensing systems.

Characterization of tub4P287L, a β‐tubulin mutant, revealed new aspects of microtubule regulation in shade

PubMed Central

Yu, Jie; Qiu, Hong; Liu, Xin; Wang, Meiling; Gao, Yongli; Chory, Joanne

2015-01-01

Abstract When sun plants, such as Arabidopsis thaliana, are under canopy shade, elongation of stems/petioles will be induced as one of the most prominent responses. Plant hormones mediate the elongation growth. However, how environmental and hormonal signals are translated into cell expansion activity that leads to the elongation growth remains elusive. Through forward genetic study, we identified shade avoidance2 (sav2) mutant, which contains a P287L mutation in β‐TUBULIN 4. Cortical microtubules (cMTs) play a key role in anisotropic cell growth. Hypocotyls of sav2 are wild type‐like in white light, but are short and highly swollen in shade and dark. We showed that shade not only induces cMT rearrangement, but also affects cMT stability and dynamics of plus ends. Even though auxin and brassinosteroids are required for shade‐induced hypocotyl elongation, they had little effect on shade‐induced rearrangement of cMTs. Blocking auxin transport suppressed dark phenotypes of sav2, while overexpressing EB1b‐GFP, a microtubule plus‐end binding protein, rescued sav2 in both shade and dark, suggesting that tub4P287L represents a unique type of tubulin mutation that does not affect cMT function in supporting cell elongation, but may affect the ability of cMTs to respond properly to growth promoting stimuli. PMID:25899068

Plant proximity perception dynamically modulates hormone levels and sensitivity in Arabidopsis.

PubMed

Bou-Torrent, Jordi; Galstyan, Anahit; Gallemí, Marçal; Cifuentes-Esquivel, Nicolás; Molina-Contreras, Maria José; Salla-Martret, Mercè; Jikumaru, Yusuke; Yamaguchi, Shinjiro; Kamiya, Yuji; Martínez-García, Jaime F

2014-06-01

The shade avoidance syndrome (SAS) refers to a set of plant responses initiated after perception by the phytochromes of light enriched in far-red colour reflected from or filtered by neighbouring plants. These varied responses are aimed at anticipating eventual shading from potential competitor vegetation. In Arabidopsis thaliana, the most obvious SAS response at the seedling stage is the increase in hypocotyl elongation. Here, we describe how plant proximity perception rapidly and temporally alters the levels of not only auxins but also active brassinosteroids and gibberellins. At the same time, shade alters the seedling sensitivity to hormones. Plant proximity perception also involves dramatic changes in gene expression that rapidly result in a new balance between positive and negative factors in a network of interacting basic helix-loop-helix proteins, such as HFR1, PAR1, and BIM and BEE factors. Here, it was shown that several of these factors act as auxin- and BR-responsiveness modulators, which ultimately control the intensity or degree of hypocotyl elongation. It was deduced that, as a consequence of the plant proximity-dependent new, dynamic, and local balance between hormone synthesis and sensitivity (mechanistically resulting from a restructured network of SAS regulators), SAS responses are unleashed and hypocotyls elongate. © The Author 2014. Published by Oxford University Press on behalf of the Society for Experimental Biology.

Variation in light-intercepting area and photosynthetic rate of sun and shade shoots of two Picea species in relation to the angle of incoming light.

PubMed

Ishii, Hiroaki; Hamada, Yoko; Utsugi, Hajime

2012-10-01

We investigated the effects of sun- and shade-shoot architecture on the photosynthetic rates of two Picea species by applying light from various angles in the laboratory. Compared with sun shoots, shade shoots were characterized by lower mass allocation per light-intercepting area, less leaf mass per shoot mass, less mutual shading among leaves and more efficient allocation of chlorophyll to photosynthesis. The shoot silhouette to total leaf area ratio (STAR(ϕ)) decreased with increasing shoot inclination angle (ϕ, the shoot axis angle relative to the projection plane) and was consistently higher for the shade shoots. Morphological and physiological characteristics of the shade shoots resulted in maximum rates of net photosynthesis at ϕ = 0° (P(max,0)) similar to that of the sun shoots when expressed on a leaf mass, total leaf area and chlorophyll basis. When the angle of incoming light was varied, P(max,ϕ) per total leaf area (P(max,ϕ )/A(T)) of the shade shoots increased linearly with increasing STAR(ϕ), while P(max,ϕ) per shoot silhouette area did not change. In contrast, the response of the sun shoots was non-linear, and an optimum angle of incoming light was determined. Our results suggest that shade-shoot morphology is adaptive for utilizing diffuse light incoming from various angles, while sun-shoot morphology is adaptive for avoiding the negative effects of strong direct radiation and for enhancing light diffusion into the canopy. We propose that the angle of incoming light should be taken into account when estimating photosynthetic rates of sun shoots of conifer trees in the field.

Nicotiana plumbaginifolia hlg mutants have a mutation in a PHYB-type phytochrome gene: they have elongated hypocotyls in red light, but are not elongated as adult plants.

PubMed

Hudson, M; Robson, P R; Kraepiel, Y; Caboche, M; Smith, H

1997-11-01

Two new allelic mutants of Nicotiana plumbaginifolia have been isolated which display a hypocotyl which is long (hlg) when seedlings are grown in continuous white light (W). This can be accounted for by the decreased response to red light (R) of the hypocotyl elongation rate in these mutants. Responses to other wavelengths are unaffected in the mutants. When grown in white light, mature hlg mutants are not elongated with respect to the wild-type; they also bolt and flower later. The shade-avoidance responses to red/far red ratio (R:FR) are intact in these mutants. Both mutants are deficient in phyB-like polypeptide that is immunodetectable in the wild-type; both have wild-type levels of a phyA-like polypeptide. These alleles are inherited in a partially dominant manner, and correspond to single-base missense mutations in a gene highly homologous to N. tabacum PHYB, which codes for a phytochrome B-type photoreceptor. One allele, hlg-1, has an introduced amino acid substitution; this may define a residue essential for phytochrome protein stability. The other allele, hlg-2, has a stop codon introduced C-terminal to the chromophore binding domain. As these phyB mutants are unaffected in shade-avoidance responses, but deficient in perception of R, it is concluded that the phyB absent in these mutants is responsible for R perception in the N. plumbaginifolia seedling, but is not a R:FR sensor in light-grown plants.

A Light-Regulated Genetic Module Was Recruited to Carpel Development in Arabidopsis following a Structural Change to SPATULA[W

PubMed Central

Reymond, Mathieu C.; Brunoud, Géraldine; Chauvet, Aurélie; Martínez-Garcia, Jaime F.; Martin-Magniette, Marie-Laure; Monéger, Françoise; Scutt, Charles P.

2012-01-01

A key innovation of flowering plants is the female reproductive organ, the carpel. Here, we show that a mechanism that regulates carpel margin development in the model flowering plant Arabidopsis thaliana was recruited from light-regulated processes. This recruitment followed the loss from the basic helix-loop-helix transcription factor SPATULA (SPT) of a domain previously responsible for its negative regulation by phytochrome. We propose that the loss of this domain was a prerequisite for the light-independent expression in female reproductive tissues of a genetic module that also promotes shade avoidance responses in vegetative organs. Striking evidence for this proposition is provided by the restoration of wild-type carpel development to spt mutants by low red/far-red light ratios, simulating vegetation shade, which we show to occur via phytochrome B, PHYTOCHROME INTERACTING FACTOR4 (PIF4), and PIF5. Our data illustrate the potential of modular evolutionary events to generate rapid morphological change and thereby provide a molecular basis for neo-Darwinian theories that describe this nongradualist phenomenon. Furthermore, the effects shown here of light quality perception on carpel development lead us to speculate on the potential role of light-regulated mechanisms in plant organs that, like the carpel, form within the shade of surrounding tissues. PMID:22851763

Reverse bias protected solar array with integrated bypass battery

NASA Technical Reports Server (NTRS)

Landis, Geoffrey A (Inventor)

2012-01-01

A method for protecting the photovoltaic cells in a photovoltaic (PV) array from reverse bias damage by utilizing a rechargeable battery for bypassing current from a shaded photovoltaic cell or group of cells, avoiding the need for a bypass diode. Further, the method mitigates the voltage degradation of a PV array caused by shaded cells.

In response to partial plant shading, the lack of phytochrome A does not directly induce leaf senescence but alters the fine-tuning of chlorophyll biosynthesis

PubMed Central

Brouwer, Bastiaan; Gardeström, Per; Keech, Olivier

2014-01-01

Phytochrome is thought to control the induction of leaf senescence directly, however, the signalling and molecular mechanisms remain unclear. In the present study, an ecophysiological approach was used to establish a functional connection between phytochrome signalling and the physiological processes underlying the induction of leaf senescence in response to shade. With shade it is important to distinguish between complete and partial shading, during which either the whole or only a part of the plant is shaded, respectively. It is first shown here that, while PHYB is required to maintain chlorophyll content in a completely shaded plant, only PHYA is involved in maintaining the leaf chlorophyll content in response to partial plant shading. Second, it is shown that leaf yellowing associated with strong partial shading in phyA-mutant plants actually correlates to a decreased biosynthesis of chlorophyll rather than to an increase of its degradation. Third, it is shown that the physiological impact of this decreased biosynthesis of chlorophyll in strongly shaded phyA-mutant leaves is accompanied by a decreased capacity to adjust the Light Compensation Point. However, the increased leaf yellowing in phyA-mutant plants is not accompanied by an increase of senescence-specific molecular markers, which argues against a direct role of PHYA in inducing leaf senescence in response to partial shade. In conclusion, it is proposed that PHYA, but not PHYB, is essential for fine-tuning the chlorophyll biosynthetic pathway in response to partial shading. In turn, this mechanism allows the shaded leaf to adjust its photosynthetic machinery to very low irradiances, thus maintaining a positive carbon balance and repressing the induction of leaf senescence, which can occur under prolonged periods of shade. PMID:24604733

Plasticity in seedling morphology, biomass allocation and physiology among ten temperate tree species in response to shade is related to shade tolerance and not leaf habit.

PubMed

Chmura, D J; Modrzyński, J; Chmielarz, P; Tjoelker, M G

2017-03-01

Mechanisms of shade tolerance in tree seedlings, and thus growth in shade, may differ by leaf habit and vary with ontogeny following seed germination. To examine early responses of seedlings to shade in relation to morphological, physiological and biomass allocation traits, we compared seedlings of 10 temperate species, varying in their leaf habit (broadleaved versus needle-leaved) and observed tolerance to shade, when growing in two contrasting light treatments - open (about 20% of full sunlight) and shade (about 5% of full sunlight). We analyzed biomass allocation and its response to shade using allometric relationships. We also measured leaf gas exchange rates and leaf N in the two light treatments. Compared to the open treatment, shading significantly increased traits typically associated with high relative growth rate (RGR) - leaf area ratio (LAR), specific leaf area (SLA), and allocation of biomass into leaves, and reduced seedling mass and allocation to roots, and net assimilation rate (NAR). Interestingly, RGR was not affected by light treatment, likely because of morphological and physiological adjustments in shaded plants that offset reductions of in situ net assimilation of carbon in shade. Leaf area-based rates of light-saturated leaf gas exchange differed among species groups, but not between light treatments, as leaf N concentration increased in concert with increased SLA in shade. We found little evidence to support the hypothesis of a increased plasticity of broadleaved species compared to needle-leaved conifers in response to shade. However, an expectation of higher plasticity in shade-intolerant species than in shade-tolerant ones, and in leaf and plant morphology than in biomass allocation was supported across species of contrasting leaf habit. © 2016 German Botanical Society and The Royal Botanical Society of the Netherlands.

Twenty-Two Years of Warming, Fertilisation and Shading of Subarctic Heath Shrubs Promote Secondary Growth and Plasticity but Not Primary Growth

PubMed Central

Campioli, Matteo; Leblans, Niki; Michelsen, Anders

2012-01-01

Most manipulation experiments simulating global change in tundra were short-term or did not measure plant growth directly. Here, we assessed the growth of three shrubs (Cassiope tetragona, Empetrum hermaphroditum and Betula nana) at a subarctic heath in Abisko (Northern Sweden) after 22 years of warming (passive greenhouses), fertilisation (nutrients addition) and shading (hessian fabric), and compare this to observations from the first decade of treatment. We assessed the growth rate of current-year leaves and apical stem (primary growth) and cambial growth (secondary growth), and integrated growth rates with morphological measurements and species coverage. Primary- and total growth of Cassiope and Empetrum were unaffected by manipulations, whereas growth was substantially reduced under fertilisation and shading (but not warming) for Betula. Overall, shrub height and length tended to increase under fertilisation and warming, whereas branching increased mostly in shaded Cassiope. Morphological changes were coupled to increased secondary growth under fertilisation. The species coverage showed a remarkable increase in graminoids in fertilised plots. Shrub response to fertilisation was positive in the short-term but changed over time, likely because of an increased competition with graminoids. More erected postures and large, canopies (requiring enhanced secondary growth for stem reinforcement) likely compensated for the increased light competition in Empetrum and Cassiope but did not avoid growth reduction in the shade intolerant Betula. The impact of warming and shading on shrub growth was more conservative. The lack of growth enhancement under warming suggests the absence of long-term acclimation for processes limiting biomass production. The lack of negative effects of shading on Cassiope was linked to morphological changes increasing the photosynthetic surface. Overall, tundra shrubs showed developmental plasticity over the longer term. However, such plasticity was associated clearly with growth rate trends only in fertilised plots. PMID:22511968

Stomatal control and leaf thermal and hydraulic capacitances under rapid environmental fluctuations.

PubMed

Schymanski, Stanislaus J; Or, Dani; Zwieniecki, Maciej

2013-01-01

Leaves within a canopy may experience rapid and extreme fluctuations in ambient conditions. A shaded leaf, for example, may become exposed to an order of magnitude increase in solar radiation within a few seconds, due to sunflecks or canopy motions. Considering typical time scales for stomatal adjustments, (2 to 60 minutes), the gap between these two time scales raised the question whether leaves rely on their hydraulic and thermal capacitances for passive protection from hydraulic failure or over-heating until stomata have adjusted. We employed a physically based model to systematically study effects of short-term fluctuations in irradiance on leaf temperatures and transpiration rates. Considering typical amplitudes and time scales of such fluctuations, the importance of leaf heat and water capacities for avoiding damaging leaf temperatures and hydraulic failure were investigated. The results suggest that common leaf heat capacities are not sufficient to protect a non-transpiring leaf from over-heating during sunflecks of several minutes duration whereas transpirative cooling provides effective protection. A comparison of the simulated time scales for heat damage in the absence of evaporative cooling with observed stomatal response times suggested that stomata must be already open before arrival of a sunfleck to avoid over-heating to critical leaf temperatures. This is consistent with measured stomatal conductances in shaded leaves and has implications for water use efficiency of deep canopy leaves and vulnerability to heat damage during drought. Our results also suggest that typical leaf water contents could sustain several minutes of evaporative cooling during a sunfleck without increasing the xylem water supply and thus risking embolism. We thus submit that shaded leaves rely on hydraulic capacitance and evaporative cooling to avoid over-heating and hydraulic failure during exposure to typical sunflecks, whereas thermal capacitance provides limited protection for very short sunflecks (tens of seconds).

Cacao Cultivation under Diverse Shade Tree Cover Allows High Carbon Storage and Sequestration without Yield Losses

PubMed Central

Abou Rajab, Yasmin; Leuschner, Christoph; Barus, Henry; Tjoa, Aiyen; Hertel, Dietrich

2016-01-01

One of the main drivers of tropical forest loss is their conversion to oil palm, soy or cacao plantations with low biodiversity and greatly reduced carbon storage. Southeast Asian cacao plantations are often established under shade tree cover, but are later converted to non-shaded monocultures to avoid resource competition. We compared three co-occurring cacao cultivation systems (3 replicate stands each) with different shade intensity (non-shaded monoculture, cacao with the legume Gliricidia sepium shade trees, and cacao with several shade tree species) in Sulawesi (Indonesia) with respect to above- and belowground biomass and productivity, and cacao bean yield. Total biomass C stocks (above- and belowground) increased fivefold from the monoculture to the multi-shade tree system (from 11 to 57 Mg ha-1), total net primary production rose twofold (from 9 to 18 Mg C ha-1 yr-1). This increase was associated with a 6fold increase in aboveground biomass, but only a 3.5fold increase in root biomass, indicating a clear shift in C allocation to aboveground tree organs with increasing shade for both cacao and shade trees. Despite a canopy cover increase from 50 to 93%, cacao bean yield remained invariant across the systems (variation: 1.1–1.2 Mg C ha-1 yr-1). The monocultures had a twice as rapid leaf turnover suggesting that shading reduces the exposure of cacao to atmospheric drought, probably resulting in greater leaf longevity. Thus, contrary to general belief, cacao bean yield does not necessarily decrease under shading which seems to reduce physical stress. If planned properly, cacao plantations under a shade tree cover allow combining high yield with benefits for carbon sequestration and storage, production system stability under stress, and higher levels of animal and plant diversity. PMID:26927428

Cacao Cultivation under Diverse Shade Tree Cover Allows High Carbon Storage and Sequestration without Yield Losses.

PubMed

Abou Rajab, Yasmin; Leuschner, Christoph; Barus, Henry; Tjoa, Aiyen; Hertel, Dietrich

2016-01-01

One of the main drivers of tropical forest loss is their conversion to oil palm, soy or cacao plantations with low biodiversity and greatly reduced carbon storage. Southeast Asian cacao plantations are often established under shade tree cover, but are later converted to non-shaded monocultures to avoid resource competition. We compared three co-occurring cacao cultivation systems (3 replicate stands each) with different shade intensity (non-shaded monoculture, cacao with the legume Gliricidia sepium shade trees, and cacao with several shade tree species) in Sulawesi (Indonesia) with respect to above- and belowground biomass and productivity, and cacao bean yield. Total biomass C stocks (above- and belowground) increased fivefold from the monoculture to the multi-shade tree system (from 11 to 57 Mg ha-1), total net primary production rose twofold (from 9 to 18 Mg C ha-1 yr-1). This increase was associated with a 6fold increase in aboveground biomass, but only a 3.5fold increase in root biomass, indicating a clear shift in C allocation to aboveground tree organs with increasing shade for both cacao and shade trees. Despite a canopy cover increase from 50 to 93%, cacao bean yield remained invariant across the systems (variation: 1.1-1.2 Mg C ha-1 yr-1). The monocultures had a twice as rapid leaf turnover suggesting that shading reduces the exposure of cacao to atmospheric drought, probably resulting in greater leaf longevity. Thus, contrary to general belief, cacao bean yield does not necessarily decrease under shading which seems to reduce physical stress. If planned properly, cacao plantations under a shade tree cover allow combining high yield with benefits for carbon sequestration and storage, production system stability under stress, and higher levels of animal and plant diversity.

Reconciling species-level vs plastic responses of evergreen leaf structure to light gradients: shade leaves punch above their weight.

PubMed

Lusk, Christopher H; Onoda, Yusuke; Kooyman, Robert; Gutiérrez-Girón, Alba

2010-04-01

*When grown in a common light environment, the leaves of shade-tolerant evergreen trees have a larger leaf mass per unit area (LMA) than their light-demanding counterparts, associated with differences in lifespan. Yet plastic responses of LMA run counter to this pattern: shade leaves have smaller LMA than sun leaves, despite often living longer. *We measured LMA and cell wall content, and conducted punch and shear tests, on sun and shade leaves of 13 rainforest evergreens of differing shade tolerance, in order to understand adaptation vs plastic responses of leaf structure and biomechanics to shade. *Species shade tolerance and leaf mechanical properties correlated better with cell wall mass per unit area than with LMA. Growth light environment had less effect on leaf mechanics than on LMA: shade leaves had, on average, 40% lower LMA than sun leaves, but differences in work-to-shear, and especially force-to-punch, were smaller. This was associated with a slightly larger cell wall fraction in shade leaves. *The persistence of shade leaves might reflect unattractiveness to herbivores because they yield smaller benefits (cell contents per area) per unit fracture force than sun leaves. In forest trees, cell wall fraction and force-to-punch are more robust correlates of species light requirements than LMA.

Ecophysiological and Anatomical Mechanisms behind the Nurse Effect: Which Are More Important? A Multivariate Approach for Cactus Seedlings

PubMed Central

Delgado-Sánchez, Pablo; Yáñez-Espinosa, Laura; Jiménez-Bremont, Juan Francisco; Chapa-Vargas, Leonardo; Flores, Joel

2013-01-01

Background Cacti establish mostly occurs under the canopy of nurse plants which provide a less stressful micro-environment, although mechanisms underlying this process are unknown. The impact of the combination of light and watering treatments on Opuntia streptacantha (Cactaceae) seedlings was examined. Methods/Principal Findings Ecophysiological [titratable acidity, osmotic potential (‘solute potential’, Ψs), relative growth rate (RGR) and their components (NAR, SLA, and LWR)], anatomical (chloroplast density, chloroplast frequency, and cell area), and environmental [photosynthetic photon flux density (PPFD) and air temperature] sets of variables were analyzed, assessing relationships between them and measuring the intensity of the relationships. Three harvests were carried out at days 15, 30, and 45. Ψs and acidity content were the most important responses for seedling establishment. The main anatomical and environmental variables were chloroplast density and water availability, respectively. Opuntia streptacantha seedlings establish better in the shade-watering treatment, due to higher Ψs and acidity, unaffected chloroplasts, and lower PPFD. In addition, the chloroplasts of cells under high-light and non-watering treatment were clumped closer to the center of the cytosol than those under shade-drought, to avoid photoinhibition and/or to better distribute or utilize the penetrating light in the green plant tissue. Conclusions Opuntia seedlings grow better under the shade, although they can tolerate drought in open spaces by increasing and moving chloroplasts and avoiding drastic decreases in their Ψs. This tolerance could have important implications for predicting the impact of climate change on natural desert regeneration, as well as for planning reforestation-afforestation practices, and rural land uses. PMID:24312310

The acclimation of Tilia cordata stomatal opening in response to light, and stomatal anatomy to vegetational shade and its components.

PubMed

Aasamaa, Krõõt; Aphalo, Pedro José

2017-02-01

Stomatal anatomical traits and rapid responses to several components of visible light were measured in Tilia cordata Mill. seedlings grown in an open, fully sunlit field (C-set), or under different kinds of shade. The main questions were: (i) stomatal responses to which visible light spectrum regions are modified by growth-environment shade and (ii) which separate component of vegetational shade is most effective in eliciting the acclimation effects of the full vegetational shade. We found that stomatal opening in response to red or green light did not differ between the plants grown in the different environments. Stomatal response to blue light was increased (in comparison with that of C-set) in the leaves grown in full vegetational shade (IABW-set), in attenuated UVAB irradiance (AB-set) or in decreased light intensity (neutral shade) plus attenuated UVAB irradiance (IAB-set). In all sets, the addition of green light-two or four times stronger-into induction light barely changed the rate of the blue-light-stimulated stomatal opening. In the AB-set, stomatal response to blue light equalled the strong IABW-set response. In attenuated UVB-grown leaves, stomatal response fell midway between IABW- and C-set results. Blue light response by neutral shade-grown leaves did not differ from that of the C-set, and the response by the IAB-set did not differ from that of the AB-set. Stomatal size was not modified by growth environments. Stomatal density and index were remarkably decreased only in the IABW- and IAB-sets. It was concluded that differences in white light responses between T. cordata leaves grown in different light environments are caused only by their different blue light response. Differences in stomatal sensitivity are not dependent on altered stomatal anatomy. Attenuated UVAB irradiance is the most efficient component of vegetational shade in stimulating acclimation of stomata, whereas decreased light intensity plays a minor role. © The Author 2016. Published by Oxford University Press. All rights reserved. For Permissions, please e-mail: journals.permissions@oup.com.

Interplay of HD-Zip II and III transcription factors in auxin-regulated plant development.

PubMed

Turchi, L; Baima, S; Morelli, G; Ruberti, I

2015-08-01

The homeodomain-leucine zipper (HD-Zip) class of transcription factors is unique to plants. HD-Zip proteins bind to DNA exclusively as dimers recognizing dyad symmetric sequences and act as positive or negative regulators of gene expression. On the basis of sequence homology in the HD-Zip DNA-binding domain, HD-Zip proteins have been grouped into four families (HD-Zip I-IV). Each HD-Zip family can be further divided into subfamilies containing paralogous genes that have arisen through genome duplication. Remarkably, all the members of the HD-Zip IIγ and -δ clades are regulated by light quality changes that induce in the majority of the angiosperms the shade-avoidance response, a process regulated at multiple levels by auxin. Intriguingly, it has recently emerged that, apart from their function in shade avoidance, the HD-Zip IIγ and -δ transcription factors control several auxin-regulated developmental processes, including apical embryo patterning, lateral organ polarity, and gynoecium development, in a white-light environment. This review presents recent advances in our understanding of HD-Zip II protein function in plant development, with particular emphasis on the impact of loss-of-function HD-Zip II mutations on auxin distribution and response. The review also describes evidence demonstrating that HD-Zip IIγ and -δ genes are directly and positively regulated by HD-Zip III transcription factors, primary determinants of apical shoot development, known to control the expression of several auxin biosynthesis, transport, and response genes. Finally, the interplay between HD-Zip II and III transcription factors in embryo apical patterning and organ polarity is discussed. © The Author 2015. Published by Oxford University Press on behalf of the Society for Experimental Biology. All rights reserved. For permissions, please email: journals.permissions@oup.com.

Responses of legumes and grasses to non-, moderate, and dense shade in Missouri, USA. I. Forage yield and its species-level plasticity

Treesearch

Kejia Pang; J.W. Van Sambeek; Nadia E. Navarrete-Tindall; Chung-Ho Lin; Shibu Jose; H. E. Garrett

2017-01-01

Annual screenings of forage grasses and legumes for shade tolerance were conducted from 1996 to 2001 in the outdoor Shade Tolerance Screening Laboratory at the Horticulture and Agroforestry Research Center, University of Missouri. Forty-three forages were grown under non-shade (100% of full sunlight), moderate shade (45%), and dense shade (20%) without competition for...

Partial shading of lateral branches affects growth, and foliage nitrogen- and water-use efficiencies in the conifer Cunninghamia lanceolata growing in a warm monsoon climate.

PubMed

Dong, Tingfa; Li, Junyu; Zhang, Yuanbin; Korpelainen, Helena; Niinemets, Ülo; Li, Chunyang

2015-06-01

The degree to which branches are autonomous in their acclimation responses to alteration in light environment is still poorly understood. We investigated the effects of shading of the sapling crown of Cunninghamia lanceolata (Lamb.) Hook on the whole-tree and mid-crown branch growth and current-year foliage structure and physiology. Four treatments providing 0, 50, 75 and 90% shading compared with full daylight (denoted as Treatment(0), Treatment(50%), Treatment(75%) and Treatment(90%), and Shaded(0), Shaded(50%), Shaded(75%) and Shaded(90%) for the shaded branches and Sunlit(0), Sunlit(50%), Sunlit(75%) and Sunlit(90%) for the opposite sunlit branches under natural light conditions, respectively), were applied over two consecutive growing seasons. Shading treatments decreased the growth of basal stem diameter, leaf dry mass per unit leaf area, stomatal conductance, transpiration rate, the ratio of water-soluble to structural leaf nitrogen content, photosynthetic nitrogen-use efficiency and instantaneous and long-term (estimated from carbon isotope composition) water-use efficiency in shaded branches. Differences between shaded and sunlit branches increased with increasing severity and duration of shading. A non-autonomous, partly compensatory behavior of non-shaded branches was observed for most traits, thus reflecting the dependence between the traits of sunlit branches and the severity of shading of the opposite crown half. The results collectively indicated that tree growth and branch and leaf acclimation responses of C. lanceolata are not only affected by the local light environment, but also by relative within-crown light conditions. We argue that such a non-autonomous branch response to changes in light conditions can improve whole-tree resource optimization. These results contribute to better understanding of tree growth and utilization of water and nitrogen under heterogeneous light conditions within tree canopies. © The Author 2015. Published by Oxford University Press. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

Responses of legumes and grasses to non-, moderate, and dense shade in Missouri, USA. II. Forage quality and its species-level plasticity

Treesearch

Kejia Pang; J.W. Van Sambeek; Nadia E. Navarrete-Tindall; Chung-Ho Lin; Shibu Jose; H. E. Garrett

2017-01-01

From a series of shade tolerance screening trials conducted in an outdoor Shade Tolerance Screening Laboratory, 22 forages (16 grasses and 6 legumes) were selected for quality evaluation. The forages were grown under non-shade (100% of full sun, the control), moderate shade (45%), and dense shade (20%) with adequate water and nutrients and free of competition from...

Assessment of color parameters of composite resin shade guides using digital imaging versus colorimeter.

PubMed

Yamanel, Kivanc; Caglar, Alper; Özcan, Mutlu; Gulsah, Kamran; Bagis, Bora

2010-12-01

This study evaluated the color parameters of resin composite shade guides determined using a colorimeter and digital imaging method. Four composite shade guides, namely: two nanohybrid (Grandio [Voco GmbH, Cuxhaven, Germany]; Premise [KerrHawe SA, Bioggio, Switzerland]) and two hybrid (Charisma [Heraeus Kulzer, GmbH & Co. KG, Hanau, Germany]; Filtek Z250 [3M ESPE, Seefeld, Germany]) were evaluated. Ten shade tabs were selected (A1, A2, A3, A3,5, A4, B1, B2, B3, C2, C3) from each shade guide. CIE Lab values were obtained using digital imaging and a colorimeter (ShadeEye NCC Dental Chroma Meter, Shofu Inc., Kyoto, Japan). The data were analyzed using two-way analysis of variance and Bonferroni post hoc test. Overall, the mean ΔE values from different composite pairs demonstrated statistically significant differences when evaluated with the colorimeter (p < 0.001) but there was no significant difference with the digital imaging method (p = 0.099). With both measurement methods in total, 80% of the shade guide pairs from different composites (97/120) showed color differences greater than 3.7 (moderately perceptible mismatch), and 49% (59/120) had obvious mismatch (ΔE > 6.8). For all shade pairs evaluated, the most significant shade mismatches were obtained between Grandio-Filtek Z250 (p = 0.021) and Filtek Z250-Premise (p = 0.01) regarding ΔE mean values, whereas the best shade match was between Grandio-Charisma (p = 0.255) regardless of the measurement method. The best color match (mean ΔE values) was recorded for A1, A2, and A3 shade pairs in both methods. When proper object-camera distance, digital camera settings, and suitable illumination conditions are provided, digital imaging method could be used in the assessment of color parameters. Interchanging use of shade guides from different composite systems should be avoided during color selection. © 2010, COPYRIGHT THE AUTHORS. JOURNAL COMPILATION © 2010, WILEY PERIODICALS, INC.

Sapling leaf trait responses to light, tree height and soil nutrients for three conifer species of contrasting shade tolerance.

PubMed

Lilles, Erica B; Astrup, Rasmus; Lefrançois, Marie-Lou; David Coates, K

2014-12-01

We developed models to describe the responses of four commonly examined leaf traits (mass per area, weight, area and nitrogen (N) concentration) to gradients of light, soil nutrients and tree height in three conifer species of contrasting shade tolerance. Our observational dataset from the sub-boreal spruce forests of British Columbia included subalpine fir (Abies lasioscarpa [Hook.] Nutt; high shade tolerance), interior spruce (Picea glauca × Picea engelmannii [Moench] Voss; intermediate shade tolerance) and lodgepole pine (Pinus contorta Dougl. ex Loud. var. latifolia; low shade tolerance) saplings from 0.18 to 4.87 m tall, in 8-98% of total incident light, from field sites with <17.6 kg ha(-1) to >46.8 kg ha(-1) total dissolved N. Leaf weights and areas showed strong positive responses to light and height, but little or no response to soil nutrients. Parameter estimates indicated that the shape of leaf weight and area responses to light corresponded with shade tolerance ranking for the three species; pine had the most linear response whereas spruce and fir had asymptotic responses. Leaf N concentration responded positively to soil nutrients, negatively to light and idiosyncratically to height. The negative effect of light was only apparent on sites of high soil nutrient availability, and parameter estimates for the shape of the negative response also corresponded to shade tolerance ranking (apine = -0.79, aspruce = -0.15, afir = -0.07). Of the traits we measured, leaf mass per area showed the least response to light, soil nutrient and height gradients. Although it is a common practice in comparisons across many species, characterizing these conifers by mean values of their leaf traits would miss important intraspecific variation across environmental and size gradients. In these forests, parameter estimates representing the intraspecific variability of leaf trait responses can be used to understand relative shade tolerances. © The Author 2014. Published by Oxford University Press. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

Light-Mediated Hormonal Regulation of Plant Growth and Development.

PubMed

de Wit, Mieke; Galvão, Vinicius Costa; Fankhauser, Christian

2016-04-29

Light is crucial for plant life, and perception of the light environment dictates plant growth, morphology, and developmental changes. Such adjustments in growth and development in response to light conditions are often established through changes in hormone levels and signaling. This review discusses examples of light-regulated processes throughout a plant's life cycle for which it is known how light signals lead to hormonal regulation. Light acts as an important developmental switch in germination, photomorphogenesis, and transition to flowering, and light cues are essential to ensure light capture through architectural changes during phototropism and the shade avoidance response. In describing well-established links between light perception and hormonal changes, we aim to give insight into the mechanisms that enable plants to thrive in variable light environments.

Dynamic facade module prototype development for solar radiation prevention in high rise building

NASA Astrophysics Data System (ADS)

Sega Sufia Purnama, Muhammad; Sutanto, Dalhar

2018-03-01

Solar radiation is an aspect that high rise building must avoid. The problem is, if high rise building facade can’t overcome, the solar thermal will come in the building, and its affects on the increasing of room temperature above comfort range. A type of additional facade element that could solve solar thermal in high rise building is adding a sun shading. A dynamic facade is a shade plane in high rise building that can moved or changed on outside condition such as solar movement and intensity. This research will discuss the dynamic facade module prototype development in high rise building in Jakarta. This research will be finish through some step. (1) Static shading shadow simulation. (2) Dynamic facade concept design development. (3) Dynamic shading shadow simulation. (4) Making of dynamic facade module prototype. (5) Field test for the dynamic facade module prototype. The dynamic facade in Jakarta case will be effective to solve solar transmission in high rise building rather than static facade.

Non-linear effects of drought under shade: reconciling physiological and ecological models in plant communities.

PubMed

Holmgren, Milena; Gómez-Aparicio, Lorena; Quero, José Luis; Valladares, Fernando

2012-06-01

The combined effects of shade and drought on plant performance and the implications for species interactions are highly debated in plant ecology. Empirical evidence for positive and negative effects of shade on the performance of plants under dry conditions supports two contrasting theoretical models about the role of shade under dry conditions: the trade-off and the facilitation hypotheses. We performed a meta-analysis of field and greenhouse studies evaluating the effects of drought at two or more irradiance levels on nine response variables describing plant physiological condition, growth, and survival. We explored differences in plant response across plant functional types, ecosystem types and methodological approaches. The data were best fit using quadratic models indicating a humped-back shape response to drought along an irradiance gradient for survival, whole plant biomass, maximum photosynthetic capacity, stomatal conductance and maximal photochemical efficiency. Drought effects were ameliorated at intermediate irradiance, becoming more severe at higher or lower light levels. This general pattern was maintained when controlling for potential variations in the strength of the drought treatment among light levels. Our quantitative meta-analysis indicates that dense shade ameliorates drought especially among drought-intolerant and shade-tolerant species. Wet tropical species showed larger negative effects of drought with increasing irradiance than semiarid and cold temperate species. Non-linear responses to irradiance were stronger under field conditions than under controlled greenhouse conditions. Non-linear responses to drought along the irradiance gradient reconciliate opposing views in plant ecology, indicating that facilitation is more likely within certain range of environmental conditions, fading under deep shade, especially for drought-tolerant species.

Climbing plants in a temperate rainforest understorey: searching for high light or coping with deep shade?

PubMed

Valladares, Fernando; Gianoli, Ernesto; Saldaña, Alfredo

2011-08-01

While the climbing habit allows vines to reach well-lit canopy areas with a minimum investment in support biomass, many of them have to survive under the dim understorey light during certain stages of their life cycle. But, if the growth/survival trade-off widely reported for trees hold for climbing plants, they cannot maximize both light-interception efficiency and shade avoidance (i.e. escaping from the understorey). The seven most important woody climbers occurring in a Chilean temperate evergreen rainforest were studied with the hypothesis that light-capture efficiency of climbers would be positively associated with their abundance in the understorey. Species abundance in the understorey was quantified from their relative frequency and density in field plots, the light environment was quantified by hemispherical photography, the photosynthetic response to light was measured with portable gas-exchange analyser, and the whole shoot light-interception efficiency and carbon gain was estimated with the 3-D computer model Y-plant. Species differed in specific leaf area, leaf mass fraction, above ground leaf area ratio, light-interception efficiency and potential carbon gain. Abundance of species in the understorey was related to whole shoot features but not to leaf level features such as specific leaf area. Potential carbon gain was inversely related to light-interception efficiency. Mutual shading among leaves within a shoot was very low (<20 %). The abundance of climbing plants in this southern rainforest understorey was directly related to their capacity to intercept light efficiently but not to their potential carbon gain. The most abundant climbers in this ecosystem match well with a shade-tolerance syndrome in contrast to the pioneer-like nature of climbers observed in tropical studies. The climbers studied seem to sacrifice high-light searching for coping with the dim understorey light.

Communicating risk information: the influence of graphical display format on quantitative information perception-Accuracy, comprehension and preferences.

PubMed

Price, Melanie; Cameron, Rachel; Butow, Phyllis

2007-12-01

Statistical health risk information has proved notoriously confusing and difficult to understand. While past research indicates that presenting risk information in a frequency format is superior to relative risk and probability formats, the optimal characteristics of frequency formats are still unclear. The aim of this study is to determine the features of 1000 person frequency diagrams (pictographs) which result in the greatest speed and accuracy of graphical perception. Participants estimated the difference in chance of survival when taking or not taking Drug A, on a pictograph format, varying by mode (one-graph/two-graph), direction (vertical/horizontal), and shading (shaded/unshaded), and their preferences for the different formats. Their understanding of different components of the 1000 person diagram was assessed. Responses were timed and scored for accuracy. Horizontal pictographs were perceived faster and more accurately than vertical formats. Two-graph pictographs were perceived faster than one-graph formats. Shading reduced response time in two-graph formats, but increased response times in one-graph formats. Shaded and one-graph pictographs were preferred. As shading and one-graph formats were preferred, further clarification as to why shading negatively impacts on response times in the one-graph format is warranted. Horizontal pictographs are optimal.

Effects of street tree shade on asphalt concrete pavement performance

Treesearch

E.G. McPherson; J. Muchnick

2005-01-01

Forty-eight street segments were paired into 24 high-and low-shade pairs in Modesto, California, U.S. Field data were collected to calculate a Pavement Condition Index (PCI) and Tree Shade Index (TSI) for each segment. Statistical analyses found that greater PCI was associated with greater TSI, indicating that tree shade was partially responsible for reduced pavement...

Prevalence of Colour Vision Anomalies Amongst Dental Professionals and its Effect on Shade Matching of Teeth.

PubMed

Khosla, Amrit; Maini, Anuj Paul; Wangoo, Anuj; Singh, Sukhman; Mehar, Damanpreet Kaur

2017-01-01

The success of a restoration is dependent on accurate shade matching of teeth leading to studies evaluating the factors affecting the perception of shades. Colour vision anomalies including colour blindness have been found to exist in the population and it has been thought to be a potential factor affecting the colour perception ability. The present study was done to evaluate the prevalence of colour vision anomalies and its effect on matching of shades of teeth. A total of 147 dental professionals were randomly selected for the study and were first tested for visual acuity using the Snellen's Eye Chart so as to carry on the study with only those operators who had a vision of 6/6. Then, the Ishihara's colour charts were used to test the operators for colour vision handicap. In the last stage of the study, test for accuracy of shade selection was done using the Vitapan Classical shade guide. The shade guide tabs were covered to avoid bias. Percentage was used to calculate the prevalence of colour vision handicap and its effect on matching of shades of teeth as compared to normal vision, which was evaluated using Chi square test. Nineteen operators had colour vision anomalies out of hundred operators and only two operators presented with colour blindness. Colour vision anomaly was more prevalent than colour blindness and it was also found that it was more prevalent in males than females. The difference between the accuracy of shade matching between the operators with normal vision and colour vision defect and operators with normal vision and colour blindness was statistically not significant. Colour blindness and colour vision handicap are rare conditions, with the latter being more common in the population. According to our study, it was concluded that no statistically significant difference existed amongst the operators with normal vision and colour vision anomaly or operators with normal vision and colour blindness during the matching of shades of teeth.

Wintering birds avoid warm sunshine: predation and the costs of foraging in sunlight.

PubMed

Carr, Jennie M; Lima, Steven L

2014-03-01

Wintering birds can gain significant thermal benefits by foraging in direct sunlight. However, exposure to bright sunlight might make birds easier to detect by predators and may also cause visual glare that can reduce a bird's ability to monitor the environment. Thus, birds likely experience a trade-off between the thermal benefits and predation-related costs of foraging in direct sunlight. To examine this possible thermoregulation-predation trade-off, we monitored the behavior of mixed-species flocks of wintering emberizid sparrows foraging in alternating strips of sunlight and shade. On average, these sparrows routinely preferred to forage in the shade, despite midday air temperatures as much as 30 °C below their thermoneutral zone. This preference for shade was strongest at relatively high temperatures when the thermal benefits of foraging in sunlight were reduced, suggesting a thermoregulation-predation trade-off. Glare could be reduced if birds faced away from the sun while feeding in direct sunlight, but we found that foraging birds tended to face southward (the direction of the sun). We speculate that other factors, such as the likely direction of predator approach, may explain this southerly orientation, particularly if predators use solar glare to their advantage during an attack. This interpretation is supported by the fact that birds had the weakest southerly orientation on cloudy days. Wintering birds may generally avoid foraging in direct sunlight to minimize their risk of predation. However, given the thermal benefits of sunshine, such birds may benefit from foraging in habitats that provide a mosaic of sunlit and shaded microhabitats.

In High-Light-Acclimated Coffee Plants the Metabolic Machinery Is Adjusted to Avoid Oxidative Stress Rather than to Benefit from Extra Light Enhancement in Photosynthetic Yield

PubMed Central

Martins, Samuel C. V.; Araújo, Wagner L.; Tohge, Takayuki; Fernie, Alisdair R.; DaMatta, Fábio M.

2014-01-01

Coffee (Coffea arabica L.) has been traditionally considered as shade-demanding, although it performs well without shade and even out-yields shaded coffee. Here we investigated how coffee plants adjust their metabolic machinery to varying light supply and whether these adjustments are supported by a reprogramming of the primary and secondary metabolism. We demonstrate that coffee plants are able to adjust its metabolic machinery to high light conditions through marked increases in its antioxidant capacity associated with enhanced consumption of reducing equivalents. Photorespiration and alternative pathways are suggested to be key players in reductant-consumption under high light conditions. We also demonstrate that both primary and secondary metabolism undergo extensive reprogramming under high light supply, including depression of the levels of intermediates of the tricarboxylic acid cycle that were accompanied by an up-regulation of a range of amino acids, sugars and sugar alcohols, polyamines and flavonoids such as kaempferol and quercetin derivatives. When taken together, the entire dataset is consistent with these metabolic alterations being primarily associated with oxidative stress avoidance rather than representing adjustments in order to facilitate the plants from utilizing the additional light to improve their photosynthetic performance. PMID:24733284

Photosensory transduction in unicellular eukaryotes: a comparison between related ciliates Blepharisma japonicum and Stentor coeruleus and photoreceptor cells of higher organisms.

PubMed

Sobierajska, Katarzyna; Fabczak, Hanna; Fabczak, Stanisław

2006-06-01

Blepharisma japonicum and Stentor coeruleus are related ciliates, conspicuous by their photosensitivity. They are capable of avoiding illuminated areas in the surrounding medium, gathering exclusively in most shaded places (photodispersal). Such behaviour results mainly from motile photophobic response occurring in ciliates. This light-avoiding response is observed during a relatively rapid increase in illumination intensity (light stimulus) and consists of cessation of cell movement, a period of backward movement (ciliary reversal), followed by a forward swimming, usually in a new direction. The photosensitivity of ciliates is ascribed to their photoreceptor system, composed of pigment granules, containing the endogenous photoreceptor -- blepharismin in Blepharisma japonicum, and stentorin in Stentor coeruleus. A light stimulus, applied to both ciliates activates specific stimulus transduction processes leading to the electrical changes at the plasma membrane, correlated with a ciliary reversal during photophobic response. These data indicate that both ciliates Blepharisma japonicum and Stentor coeruleus, the lower eukaryotes, are capable of transducing the perceived light stimuli in a manner taking place in some photoreceptor cells of higher eukaryotes. Similarities and differences concerning particular stages of light transduction in eukaryotes at different evolutional levels are discussed in this article.

Fearing shades of grey: individual differences in fear responding towards generalisation stimuli.

PubMed

Arnaudova, Inna; Krypotos, Angelos-Miltiadis; Effting, Marieke; Kindt, Merel; Beckers, Tom

2017-09-01

Individual differences in fear generalisation have been proposed to play a role in the aetiology and/or maintenance of anxiety disorders, but few data are available to directly support that claim. The research that is available has focused mostly on generalisation of peripheral and central physiological fear responses. Far less is known about the generalisation of avoidance, the behavioural component of fear. In two experiments, we evaluated how neuroticism, a known vulnerability factor for anxiety, modulates an array of fear responses, including avoidance tendencies, towards generalisation stimuli (GS). Participants underwent differential fear conditioning, in which one conditioned stimulus (CS+) was repeatedly paired with an aversive outcome (shock; unconditioned stimulus, US), whereas another was not (CS-). Fear generalisation was observed across measures in Experiment 1 (US expectancy and evaluative ratings) and Experiment 2 (US expectancy, evaluative ratings, skin conductance, startle responses, safety behaviours), with overall highest responding to the CS+, lowest to the CS- and intermediate responding to the GSs. Neuroticism had very little impact on fear generalisation (but did affect GS recognition rates in Experiment 1), in line with the idea that fear generalisation is largely an adaptive process.

Shade response of a full size TESSERA module

NASA Astrophysics Data System (ADS)

Slooff, Lenneke H.; Carr, Anna J.; de Groot, Koen; Jansen, Mark J.; Okel, Lars; Jonkman, Rudi; Bakker, Jan; de Gier, Bart; Harthoorn, Adriaan

2017-08-01

A full size TESSERA shade tolerant module has been made and was tested under various shadow conditions. The results show that the dedicated electrical interconnection of cells result in an almost linear response under shading. Furthermore, the voltage at maximum power point is almost independent of the shadow. This decreases the demand on the voltage range of the inverter. The increased shadow linearity results in a calculated increase in annual yield of about 4% for a typical Dutch house.

Seasonal contrasts in the response of coffee ants to agroforestry shade-tree management.

PubMed

Teodoro, A V; Sousa-Souto, L; Klein, A-M; Tscharntke, T

2010-12-01

In many tropical landscapes, agroforestry systems are the last forested ecosystems, providing shade, having higher humidity, mitigating potential droughts, and possessing more species than any other crop system. Here, we tested the hypothesis that higher levels of shade and associated humidity in agroforestry enhance coffee ant richness more during the dry than rainy season, comparing ant richness in 22 plots of three coffee agroforestry types in coastal Ecuador: simple-shade agroforests (intensively managed with low tree species diversity), complex-shade agroforests (extensively managed with intermediate tree species diversity) and abandoned coffee agroforests (abandoned for 10-15 yr and resembling secondary forests). Seasonality affected responses of ant richness but not composition to agroforestry management, in that most species were observed in abandoned coffee agroforests in the dry season. In the rainy season, however, most species were found in simple-shade agroforests, and complex agroforestry being intermediate. Foraging coffee ants species composition did not change differently according to agroforestry type and season. Results show that shade appears to be most important in the dry seasons, while a mosaic of different land-use types may provide adequate environmental conditions to ant species, maximizing landscape-wide richness throughout the year. © 2010 Entomological Society of America

Physiological responses of feedlot heifers provided access to different levels of shade

USDA-ARS?s Scientific Manuscript database

Heat stress has a significant impact on all livestock and poultry species causing economic losses and animal well-being concerns. Providing shade is one heat-abatement strategy that has been studied for years. Material selected to provide shade for animals greatly influences the overall stress reduc...

Species-Independent Down-Regulation of Leaf Photosynthesis and Respiration in Response to Shading: Evidence from Six Temperate Tree Species

PubMed Central

Chen, Anping; Lichstein, Jeremy W.; Osnas, Jeanne L. D.; Pacala, Stephen W.

2014-01-01

The ability to down-regulate leaf maximum net photosynthetic capacity (Amax) and dark respiration rate (Rdark) in response to shading is thought to be an important adaptation of trees to the wide range of light environments that they are exposed to across space and time. A simple, general rule that accurately described this down-regulation would improve carbon cycle models and enhance our understanding of how forest successional diversity is maintained. In this paper, we investigated the light response of Amax and Rdark for saplings of six temperate forest tree species in New Jersey, USA, and formulated a simple model of down-regulation that could be incorporated into carbon cycle models. We found that full-sun values of Amax and Rdark differed significantly among species, but the rate of down-regulation (proportional decrease in Amax or Rdark relative to the full-sun value) in response to shade was not significantly species- or taxon-specific. Shade leaves of sun-grown plants appear to follow the same pattern of down-regulation in response to shade as leaves of shade-grown plants. Given the light level above a leaf and one species-specific number (either the full-sun Amax or full-sun Rdark), we provide a formula that can accurately predict the leaf's Amax and Rdark. We further show that most of the down regulation of per unit area Rdark and Amax is caused by reductions in leaf mass per unit area (LMA): as light decreases, leaves get thinner, while per unit mass Amax and Rdark remain approximately constant. PMID:24727745

The effects of ultraviolet radiation on growth and bleaching in three species of Hawaiian coral

DOE Office of Scientific and Technical Information (OSTI.GOV)

Goodman, G.D.

1990-01-09

Long term exposure to ultraviolet radiation is harmful to many organisms, including hermatypic corals, which obtain much of their nutrition from photosynthetic zooxanthellae. Therefore, increased UV radiation from atmospheric ozone depletion could inhibit growth of such corals. Moreover, coral bleaching, which has been attributed to loss of pigment and/or expulsion of zooxanthellae, may be a specific response to UV light. Does UV-A reduce skeletal growth or influence population density and pigment content of zooxanthellae In addition, do zooxanthellae migrate to shaded areas of the colony to avoid ultraviolet light Using alizarin red stain and suitable filters, I compared the stainmore » and suitable filters, I compared the effects of UV-A (320-400nm) and full-spectrum UV (280-400nm) on the skeletal growth of two Hawaiian corals, Montipora verrucosa, Pocillopora damicornis, in situ. In the perforate corals, M. Verrucosa and Porites compressa, I measured concentration of zooxanthellae and their chlorophyll content to quantify bleaching in response to UV light. Reduction in skeletal growth by the two corals in response to different ranges of UV light appears to be species specific. Bleaching by UV appears to be characterized by an initial loss of pigment followed by the expulsion and migration of the zooxanthellae to shaded areas of the colony. Differences in tolerance and adaptation to decreasing ozone levels and increasing UV light should confer a competitive advantage on various species and morphologies of reef-building corals.« less

Phenotypic Plasticity in Reproductive Traits of the Perennial Shrub Ulex europaeus in Response to Shading: A Multi-Year Monitoring of Cultivated Clones.

PubMed

Atlan, Anne; Hornoy, Benjamin; Delerue, Florian; Gonzalez, Maya; Pierre, Jean-Sébastien; Tarayre, Michèle

2015-01-01

Phenotypic plasticity may be advantageous for plants to be able to rapidly cope with new and changing environments associated with climate change or during biological invasions. This is especially true for perennial plants, as they may need a longer period to respond genetically to selective pressures than annuals, and also because they are more likely to experience environmental changes during their lifespan. However, few studies have explored the plasticity of the reproductive life history traits of woody perennial species. This study focuses on a woody shrub, Ulex europaeus (common gorse), and on the response of its reproductive traits to one important environmental factor, shading. The study was performed on clones originating from western France (within the native range of this invasive species) and grown for seven years. We compared traits of plants grown in a shade treatment (with two successive shade levels) vs. full natural light. The traits monitored included flowering onset, pod production and seed predation. All traits studied responded to shading, exhibiting various levels of plasticity. In particular, dense shade induced a radical but reversible decrease in flower and pod production, while moderate shade had little effect on reproductive traits. The magnitude of the response to dense shade depended on the genotype, showing a genetically based polymorphism of plasticity. The level of plasticity also showed substantial variations between years, and the effect of environmental variations was cumulative over time. This suggests that plasticity can influence the lifetime fitness of U. Europaeus and is involved in the capacity of the species to grow under contrasting environmental conditions.

Phenotypic Plasticity in Reproductive Traits of the Perennial Shrub Ulex europaeus in Response to Shading: A Multi-Year Monitoring of Cultivated Clones

PubMed Central

Atlan, Anne; Hornoy, Benjamin; Delerue, Florian; Gonzalez, Maya; Pierre, Jean-Sébastien; Tarayre, Michèle

2015-01-01

Phenotypic plasticity may be advantageous for plants to be able to rapidly cope with new and changing environments associated with climate change or during biological invasions. This is especially true for perennial plants, as they may need a longer period to respond genetically to selective pressures than annuals, and also because they are more likely to experience environmental changes during their lifespan. However, few studies have explored the plasticity of the reproductive life history traits of woody perennial species. This study focuses on a woody shrub, Ulex europaeus (common gorse), and on the response of its reproductive traits to one important environmental factor, shading. The study was performed on clones originating from western France (within the native range of this invasive species) and grown for seven years. We compared traits of plants grown in a shade treatment (with two successive shade levels) vs. full natural light. The traits monitored included flowering onset, pod production and seed predation. All traits studied responded to shading, exhibiting various levels of plasticity. In particular, dense shade induced a radical but reversible decrease in flower and pod production, while moderate shade had little effect on reproductive traits. The magnitude of the response to dense shade depended on the genotype, showing a genetically based polymorphism of plasticity. The level of plasticity also showed substantial variations between years, and the effect of environmental variations was cumulative over time. This suggests that plasticity can influence the lifetime fitness of U. Europaeus and is involved in the capacity of the species to grow under contrasting environmental conditions. PMID:26383627

Physiological responses of spring rapeseed (Brassica napus) to red/far-red ratios and irradiance during pre- and post-flowering stages.

PubMed

Rondanini, Deborah P; del Pilar Vilariño, Maria; Roberts, Marcos E; Polosa, Marina A; Botto, Javier F

2014-12-01

Early shade signals promote the shade avoidance syndrome (SAS) which causes, among others, petiole and shoot elongation and upward leaf position. In spite of its relevance, these photomorphogenic responses have not been deeply studied in rapeseed (Brassica napus). In contrast to other crops like maize and wheat, rapeseed has a complex developmental phenotypic pattern as it evolves from an initial rosette to the main stem elongation and an indeterminate growth of floral raceme. In this work, we analyzed (1) morphological and physiological responses at individual level due to low red/far-red (R/FR) ratio during plant development, and (2) changes in biomass allocation, grain yield and composition at crop level in response to high R/FR ratio and low irradiance in two modern spring rapeseed genotypes. We carried out pot and field experiments modifying R/FR ratios and irradiance at vegetative or reproductive stages. In pot experiments, low R/FR ratio increased the petiole and lamina length, upward leaf position and also accelerated leaf senescence. Furthermore, low R/FR ratio reduced main floral raceme and increased floral branching with higher remobilization of soluble carbohydrates from the stems. In field experiments, low irradiance during post-flowering reduced grain yield, harvest index and grain oil content, and high R/FR ratio reaching the crop partially alleviated such effects. We conclude that photomorphogenic signals are integrated early during the vegetative growth, and irradiance has stronger effects than R/FR signals at rapeseed crop level. © 2014 Scandinavian Plant Physiology Society.

Prevalence of Colour Vision Anomalies Amongst Dental Professionals and its Effect on Shade Matching of Teeth

PubMed Central

Maini, Anuj Paul; Wangoo, Anuj; Singh, Sukhman; Mehar, Damanpreet Kaur

2017-01-01

Abstract Introduction The success of a restoration is dependent on accurate shade matching of teeth leading to studies evaluating the factors affecting the perception of shades. Colour vision anomalies including colour blindness have been found to exist in the population and it has been thought to be a potential factor affecting the colour perception ability. Aim The present study was done to evaluate the prevalence of colour vision anomalies and its effect on matching of shades of teeth. Materials and Methods A total of 147 dental professionals were randomly selected for the study and were first tested for visual acuity using the Snellen’s Eye Chart so as to carry on the study with only those operators who had a vision of 6/6. Then, the Ishihara’s colour charts were used to test the operators for colour vision handicap. In the last stage of the study, test for accuracy of shade selection was done using the Vitapan Classical shade guide. The shade guide tabs were covered to avoid bias. Percentage was used to calculate the prevalence of colour vision handicap and its effect on matching of shades of teeth as compared to normal vision, which was evaluated using Chi square test. Results Nineteen operators had colour vision anomalies out of hundred operators and only two operators presented with colour blindness. Colour vision anomaly was more prevalent than colour blindness and it was also found that it was more prevalent in males than females. The difference between the accuracy of shade matching between the operators with normal vision and colour vision defect and operators with normal vision and colour blindness was statistically not significant. Conclusion Colour blindness and colour vision handicap are rare conditions, with the latter being more common in the population. According to our study, it was concluded that no statistically significant difference existed amongst the operators with normal vision and colour vision anomaly or operators with normal vision and colour blindness during the matching of shades of teeth. PMID:28274040

Karrikins: Regulators Involved in Phytohormone Signaling Networks during Seed Germination and Seedling Development

PubMed Central

Meng, Yongjie; Shuai, Haiwei; Luo, Xiaofeng; Chen, Feng; Zhou, Wenguan; Yang, Wenyu; Shu, Kai

2017-01-01

Seed germination and early seedling establishment are critical stages during a plant’s life cycle. These stages are precisely regulated by multiple internal factors, including phytohormones and environmental cues such as light. As a family of small molecules discovered in wildfire smoke, karrikins (KARs) play a key role in various biological processes, including seed dormancy release, germination regulation, and seedling establishment. KARs show a high similarity with strigolactone (SL) in both chemical structure and signaling transduction pathways. Current evidence shows that KARs may regulate seed germination by mediating the biosynthesis and/or signaling transduction of abscisic acid (ABA), gibberellin (GA) and auxin [indoleacetic acid (IAA)]. Interestingly, KARs regulate seed germination differently in different species. Furthermore, the promotion effect on seedling establishment implies that KARs have a great potential application in alleviating shade avoidance response, which attracts more and more attention in plant molecular biology. In these processes, KARs may have complicated interactions with phytohormones, especially with IAA. In this updated review, we summarize the current understanding of the relationship between KARs and SL in the chemical structure, signaling pathway and the regulation of plant growth and development. Further, the crosstalk between KARs and phytohormones in regulating seed germination and seedling development and that between KARs and IAA during shade responses are discussed. Finally, future challenges and research directions for the KAR research field are suggested. PMID:28174573

Shade material evaluation using a cattle response model and meteorological instrumentation.

PubMed

Eigenberg, Roger A; Brown-Brandl, Tami M; Nienaber, John A

2010-11-01

Shade structures are often considered as one method of reducing stress in feedlot cattle. Selection of a suitable shade material can be difficult without data that quantify material effectiveness for stress reduction. A summer study was conducted during 2007 using instrumented shade structures in conjunction with meteorological measurements to estimate relative effectiveness of various shade materials. Shade structures were 3.6 m by 6.0 m by 3.0 m high at the peak and 2.0 m high at the sides. Polyethylene shade cloth was used in three of the comparisons and consisted of effective coverings of 100%, 60% with a silver reflective coating, and 60% black material with no reflective coating. Additionally, one of the structures was fitted with a poly snow fence with an effective shade of about 30%. Each shade structure contained a solar radiation meter and a black globe thermometer to measure radiant energy received under the shade material. Additionally, meteorological data were collected as a non-shaded treatment and included temperature, humidity, wind speed, and solar radiation. Data analyses was conducted using a physiological model based on temperature, humidity, solar radiation and wind speed; a second model using black globe temperatures, relative humidity, and wind speed was used as well. Analyses of the data revealed that time spent in the highest stress category was reduced by all shade materials. Moreover, significant differences (P < 0.05) existed between all shade materials (compared to no-shade) for hourly summaries during peak daylight hours and for 'full sun' days.

Shade material evaluation using a cattle response model and meteorological instrumentation.

PubMed

Eigenberg, Roger A; Brown-Brandl, Tami M; Nienaber, John A

2009-11-01

Shade structures are often considered as one method of reducing stress in feedlot cattle. Selection of a suitable shade material can be difficult without data that quantify material effectiveness for stress reduction. A summer study was conducted during 2007 using instrumented shade structures in conjunction with meteorological measurements to estimate relative effectiveness of various shade materials. Shade structures were 3.6 m by 6.0 m by 3.0 m high at the peak and 2.0 m high at the sides. Polyethylene shade cloth was used in three of the comparisons and consisted of effective coverings of 100%, 60% with a silver reflective coating, and 60% black material with no reflective coating. Additionally, one of the structures was fitted with a poly snow fence with an effective shade of about 30%. Each shade structure contained a solar radiation meter and a black globe thermometer to measure radiant energy received under the shade material. Additionally, meteorological data were collected as a non-shaded treatment and included temperature, humidity, wind speed, and solar radiation. Data analyses was conducted using a physiological model based on temperature, humidity, solar radiation and wind speed; a second model using black globe temperatures, relative humidity, and wind speed was used as well. Analyses of the data revealed that time spent in the highest stress category was reduced by all shade materials. Moreover, significant differences (P < 0.05) existed between all shade materials (compared to no-shade) for hourly summaries during peak daylight hours and for 'full sun' days.

Shade material evaluation using a cattle response model and meteorological instrumentation.

PubMed

Eigenberg, Roger A; Brown-Brandl, Tami M; Nienaber, John A

2010-09-01

Shade structures are often considered as one method of reducing stress in feedlot cattle. Selection of a suitable shade material can be difficult without data that quantify material effectiveness for stress reduction. A summer study was conducted during 2007 using instrumented shade structures in conjunction with meteorological measurements to estimate relative effectiveness of various shade materials. Shade structures were 3.6 m x 6.0 m x 3.0 m high at the peak and 2.0 m high at the sides. Polyethylene shade cloth was used in three of the comparisons and consisted of effective coverings of 100%, 60% with a silver reflective coating, and 60% black material with no reflective coating. Additionally, one of the structures was fitted with a poly snow fence with an effective shade of about 30%. Each shade structure contained a solar radiation meter and a black globe thermometer to measure radiant energy received under the shade material. Additionally, meteorological data were collected as a non-shaded treatment and included temperature, humidity, wind speed, and solar radiation. Data analyses was conducted using a physiological model based on temperature, humidity, solar radiation and wind speed; a second model using black globe temperatures, relative humidity, and wind speed was used as well. Analyses of the data revealed that time spent in the highest stress category was reduced by all shade materials. Moreover, significant differences (P < 0.05) existed between all shade materials (compared to no-shade) for hourly summaries during peak daylight hours and for 'full sun' days.

The Ratio of Leaf to Total Photosynthetic Area Influences Shade Survival and Plastic Response to Light of Green‐stemmed Leguminous Shrub Seedlings

PubMed Central

VALLADARES, FERNANDO; HERNÁNDEZ, LIBERTAD G.; DOBARRO, IKER; GARCÍA‐PÉREZ, CRISTINA; SANZ, RUBÉN; PUGNAIRE, FRANCISCO I.

2003-01-01

Different plant species and organs within a plant differ in their plastic response to light. These responses influence their performance and survival in relation to the light environment, which may range from full sunlight to deep shade. Plasticity, especially with regard to physiological features, is linked to a greater capacity to exploit high light and is usually low in shade‐tolerant species. Among photosynthetic organs, green stems, which represent a large fraction of the total photosynthetic area of certain species, are hypothesized to be less capable of adjustment to light than leaves, because of biomechanical and hydraulic constraints. The response to light by leaves and stems of six species of leguminous, green‐stemmed shrubs from dry and high‐light environments was studied by growing seedlings in three light environments: deep shade, moderate shade and sun (3, 30 and 100 % of full sunlight, respectively). Survival in deep shade ranged from 2 % in Retama sphaerocarpa to 74 % in Ulex europaeus. Survival was maximal at moderate shade in all species, ranging from 80 to 98 %. The six species differed significantly in their ratio of leaf to total photosynthetic area, which influenced their light response. Survival in deep shade increased significantly with increasing ratio of leaf to total photosynthetic area, and decreased with increasing plasticity in net photosynthesis and dark respiration. Responses to light differed between stems and leaves within each species. Mean phenotypic plasticity for the variables leaf or stem specific mass, chlorophyll content, chlorophyll a/b ratio, and carotenoid to chlorophyll ratio of leaves, was inversely related to that of stems. Although mean plasticity of stems increased with the ratio of leaf to total photosynthetic area, the mean plasticity of leaves decreased. Shrubs with green stems and a low ratio of leaf to total photosynthetic area are expected to be restricted to well‐lit habitats, at least during the seedling stage, owing to their inefficient light capture and the low plasticity of their stems. PMID:12646502

Is shade for horses a comfort resource or a minimum requirement?

PubMed

Holcomb, K E

2017-09-01

Shade or shelter as protection from extremes of weather is required for horses at agricultural research and teaching facilities and is recommended or required by many states, professional organizations, and industry groups. The focus of this paper is the recent research on the responses of horses to hot, sunny weather, which has begun to provide scientific evidence that characterizes how and when shade is used and any benefits shade confers on horses. These behavioral and physiological findings support provision of shade as a resource for thermal comfort and the expression of normal behavior that should be included as a standard of best care practices for healthy adult horses living in the environmental conditions reviewed, rather than an absolute minimum care requirement. Additional research is required for horses living under other environmental conditions, for very young or old horses, horses in very poor body condition, or those with compromised health to determine if their responses to hot, sunny weather differ from those presented here.

Response of naturally regenerated and underplanted white oak (Quercus alba L.) seedlings 6 years following midstory removal

Treesearch

Jared M. Craig; John M. Lhotka; Jeffrey W. Stringer

2013-01-01

Historically, the abundance of red maple and shade-tolerant understory species was limited by fire and other disturbances. In the absence of periodic disturbance, regeneration of oaks with intermediate shade tolerance has been hindered due to inadequate light conditions created by shade-tolerant midstory trees. Research suggests that midstory removal in oak-dominated...

Climbing plants in a temperate rainforest understorey: searching for high light or coping with deep shade?

PubMed Central

Valladares, Fernando; Gianoli, Ernesto; Saldaña, Alfredo

2011-01-01

Background and Aims While the climbing habit allows vines to reach well-lit canopy areas with a minimum investment in support biomass, many of them have to survive under the dim understorey light during certain stages of their life cycle. But, if the growth/survival trade-off widely reported for trees hold for climbing plants, they cannot maximize both light-interception efficiency and shade avoidance (i.e. escaping from the understorey). The seven most important woody climbers occurring in a Chilean temperate evergreen rainforest were studied with the hypothesis that light-capture efficiency of climbers would be positively associated with their abundance in the understorey. Methods Species abundance in the understorey was quantified from their relative frequency and density in field plots, the light environment was quantified by hemispherical photography, the photosynthetic response to light was measured with portable gas-exchange analyser, and the whole shoot light-interception efficiency and carbon gain was estimated with the 3-D computer model Y-plant. Key Results Species differed in specific leaf area, leaf mass fraction, above ground leaf area ratio, light-interception efficiency and potential carbon gain. Abundance of species in the understorey was related to whole shoot features but not to leaf level features such as specific leaf area. Potential carbon gain was inversely related to light-interception efficiency. Mutual shading among leaves within a shoot was very low (<20 %). Conclusions The abundance of climbing plants in this southern rainforest understorey was directly related to their capacity to intercept light efficiently but not to their potential carbon gain. The most abundant climbers in this ecosystem match well with a shade-tolerance syndrome in contrast to the pioneer-like nature of climbers observed in tropical studies. The climbers studied seem to sacrifice high-light searching for coping with the dim understorey light. PMID:21685433

Periphyton response to long-term nutrient enrichment in a shaded headwater stream

Treesearch

Jennifer L. Greenwood; Amy D. Rosemond

2009-01-01

We maintained elevated but moderate concentrations of nitrogen and phosphorus continuously for 2 years in a heavily shaded headwater stream and compared effects on stream periphyton with a reference...

Salinity stress constrains photosynthesis in Fraxinus ornus more when growing in partial shading than in full sunlight: consequences for the antioxidant defence system

PubMed Central

Fini, Alessio; Guidi, Lucia; Giordano, Cristiana; Baratto, Maria Camilla; Ferrini, Francesco; Brunetti, Cecilia; Calamai, Luca; Tattini, Massimiliano

2014-01-01

Background and Aims A major challenge in plant ecophysiology is understanding the effects of multiple sub-optimal environmental conditions on plant performance. In most Mediterranean areas soil salinity builds up during the summer because of low availability of soil water coupled with hot temperatures. Although sunlight and soil salinity may strongly interact in determining a plant's performance, this has received relatively little attention. Methods Two-year-old seedlings of Fraxinus ornus were grown outdoors in pots during a Mediterranean summer in either 45 % (shaded plants) or 100 % (sun plants) sunlight irradiance and were supplied with either deionized water or deionized water plus 75 mm NaCl. Morpho-anatomical traits, water and ionic relations, gas exchange and photosystem II performance, concentrations of individual carotenoids, activity of antioxidant enzymes, concentrations of ascorbic acid and individual polyphenols were measured in leaves. Leaf oxidative stress and damage were estimated by in vivo analysis of stable free radicals and ultrastructural analyses. Key Results Leaf concentrations of potentially toxic ions did not markedly differ in shaded or sun plants in response to salinity. Leaves of sun plants displayed superior water use efficiency compared with leaves of shaded plants, irrespective of salinity treatment, and had both better stomatal control and higher CO2 carboxylation efficiency than leaves of shaded plants. In the salt-treated groups, the adverse effects of excess midday irradiance were greater in shade than in sun plants. The activity of enzymes responsible for detoxifying hydrogen peroxide decreased in shaded plants and increased in sun plants as a result of salinity stress. In contrast, the activity of guaiacol peroxidase and the concentration of phenylpropanoids increased steeply in response to salinity in shaded plants but were unaffected in sun plants. Conclusions It is concluded that salinity may constrain the performance of plants growing under partial shading more severely than that of plants growing under full sun during summer. The results suggest co-ordination within the antioxidant defence network aimed at detoxifying salt-induced generation of reactive oxygen species. PMID:25006177

Response of the Fine Root Production, Phenology, and Turnover Rate of Six Shrub Species from a Subtropical Forest to a Soil Moisture Gradient and Shading

NASA Astrophysics Data System (ADS)

Fu, X.; Dai, X.; Wang, H.

2015-12-01

Knowledge of the fine root dynamics of different life forms in forest ecosystems is critical to understanding how the overall belowground carbon cycling is affected by climate change. However, our current knowledge regarding how endogenous or exogenous factors regulate the root dynamics of understory vegetation is limited. We selected a suite of study sites representing different habitats with gradients of soil moisture and solar radiation (shading or no shading). We assessed the fine root production phenology, the total fine root production, and the turnover among six understory shrub species in a subtropical climate, and examined the responses of the fine root dynamics to gradients in the soil moisture and solar radiation. The shrubs included three evergreen species, Loropetalum chinense, Vaccinium bracteatum, and Adinandra millettii, and three deciduous species, Serissa serissoides, Rubus corchorifolius, and Lespedeza davidii. We observed that variations in the annual fine root production and turnover among species were significant in the deciduous group but not in the evergreen group. Notably, V. bracteatum and S. serissoides presented the greatest responses in terms of root phenology to gradients in the soil moisture and shading: high-moisture habitat led to a decrease and shade led to an increase in fine root production during spring. Species with smaller fine roots of the 1st+2nd-order diameter presented more sensitive responses in terms of fine root phenology to a soil moisture gradient. Species with a higher fine root nitrogen-to -carbon ratio exhibited more sensitive responses in terms of fine root annual production to shading. Soil moisture and shading did not change the annual fine root production as much as the turnover rate. The fine root dynamics of some understory shrubs varied significantly with soil moisture and solar radiation status and may be different from tree species. Our results emphasize the need to study the understory fine root dynamics in the achievement of a complete understanding of the overall belowground carbon cycling in a forest ecosystem, particularly ecosystems in which the understory fine root highly contributes to the belowground biomass.

Does greater specific leaf area plasticity help plants to maintain a high performance when shaded?

PubMed

Liu, Yanjie; Dawson, Wayne; Prati, Daniel; Haeuser, Emily; Feng, Yanhao; van Kleunen, Mark

2016-12-01

It is frequently assumed that phenotypic plasticity can be very advantageous for plants, because it may increase environmental tolerance (fitness homeostasis). This should, however, only hold for plastic responses that are adaptive, i.e. increase fitness. Numerous studies have shown shade-induced increases in specific leaf area (SLA), and there is wide consensus that this plastic response optimizes light capture and thus has to be adaptive. However, it has rarely been tested whether this is really the case. In order to identify whether SLA plasticity does contribute to the maintenance of high biomass of plant species under shaded conditions, a meta-analytical approach was employed. The data set included 280 species and 467 individual studies from 32 publications and two unpublished experiments. Plants increased their SLA by 55·4 % on average when shaded, while they decreased their biomass by 59·9 %. Species with a high SLA under high-light control conditions showed a significantly greater ability to maintain biomass production under shade overall. However, in contrast to the expectation of a positive relationship between SLA plasticity and maintenance of plant biomass, the results indicated that species with greater SLA plasticity were less able to maintain biomass under shade. Although a high SLA per se contributes to biomass homeostasis, there was no evidence that plasticity in SLA contributes to this. Therefore, it is argued that some of the plastic changes that are frequently thought to be adaptive might simply reflect passive responses to the environment, or result as by-products of adaptive plastic responses in other traits. © The Author 2016. Published by Oxford University Press on behalf of the Annals of Botany Company. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

Allocation strategies of savanna and forest tree seedlings in response to fire and shading: outcomes of a field experiment

NASA Astrophysics Data System (ADS)

Gignoux, Jacques; Konaté, Souleymane; Lahoreau, Gaëlle; Le Roux, Xavier; Simioni, Guillaume

2016-12-01

The forest-savanna ecotone may be very sharp in fire-prone areas. Fire and competition for light play key roles in its maintenance, as forest and savanna tree seedlings are quickly excluded from the other ecosystem. We hypothesized a tradeoff between seedling traits linked to fire resistance and to competition for light to explain these exclusions. We compared growth- and survival-related traits of two savanna and two forest species in response to shading and fire in a field experiment. To interpret the results, we decomposed our broad hypothesis into elementary tradeoffs linked to three constraints, biomass allocation, plant architecture, and shade tolerance, that characterize both savanna and adjacent forest ecosystems. All seedlings reached similar biomasses, but forest seedlings grew taller. Savanna seedlings better survived fire after topkill and required ten times less biomass than forest seedlings to survive. Finally, only savanna seedlings responded to shading. Although results were consistent with the classification of our species as mostly adapted to shade tolerance, competition for light in the open, and fire tolerance, they raised new questions: how could savanna seedlings survive better with a 10-times lower biomass than forest seedlings? Is their shade intolerance sufficient to exclude them from forest understory?

Season-dependent and independent responses of Mediterranean scrub to light conditions.

PubMed

Zunzunegui, María; Díaz-Barradas, Mari Cruz; Jáuregui, Juan; Rodríguez, Herminia; Álvarez-Cansino, Leonor

2016-05-01

Semi-arid plant species cope with excess of solar radiation with morphological and physiological adaptations that assure their survival when other abiotic stressors interact. At the leaf level, sun and shade plants may differ in the set of traits that regulate environmental stressors. Here, we evaluated if leaf-level physiological seasonal response of Mediterranean scrub species (Myrtus communis, Halimium halimifolium, Rosmarinus officinalis, and Cistus salvifolius) depended on light availability conditions. We aimed to determine which of these responses prevailed independently of the marked seasonality of Mediterranean climate, to define a leaf-level strategy in the scrub community. Thirty six leaf response variables - involving gas exchange, water status, photosystem II photochemical efficiency, photosynthetic pigments and leaf structure - were seasonally measured in sun exposed and shaded plants under field conditions. Physiological responses showed a common pattern throughout the year, in spite of the marked seasonality of the Mediterranean climate and of species-specific differences in the response to light intensity. Variables related to light use, CO2 assimilation, leaf pigment content, and LMA (leaf mass area) presented differences that were consistent throughout the year, although autumn was the season with greater contrast between sun and shade plants. Our data suggest that in Mediterranean scrub shade plants the lutein pool could have an important role in the photoprotection of the photosynthetic tissues. There was a negative linear correlation between the ratio lutein/total chlorophylls and the majority of leaf level variables. The combined effect of abiotic stress factors (light and drought or light and cold) was variable-specific, in some cases enhancing differences between sun and shade plants, while in others leading to unified strategies in all scrub species. Copyright © 2016 Elsevier Masson SAS. All rights reserved.

Structure and function of homodomain-leucine zipper (HD-Zip) proteins.

PubMed

Elhiti, Mohamed; Stasolla, Claudio

2009-02-01

Homeodomain-leucine zipper (HD-Zip) proteins are transcription factors unique to plants and are encoded by more than 25 genes in Arabidopsis thaliana. Based on sequence analyses these proteins have been classified into four distinct groups: HD-Zip I-IV. HD-Zip proteins are characterized by the presence of two functional domains; a homeodomain (HD) responsible for DNA binding and a leucine zipper domain (Zip) located immediately C-terminal to the homeodomain and involved in protein-protein interaction. Despite sequence similarities HD-ZIP proteins participate in a variety of processes during plant growth and development. HD-Zip I proteins are generally involved in responses related to abiotic stress, abscisic acid (ABA), blue light, de-etiolation and embryogenesis. HD-Zip II proteins participate in light response, shade avoidance and auxin signalling. Members of the third group (HD-Zip III) control embryogenesis, leaf polarity, lateral organ initiation and meristem function. HD-Zip IV proteins play significant roles during anthocyanin accumulation, differentiation of epidermal cells, trichome formation and root development.

[Response of photosynthetic characteristics of peanut seedlings leaves to low light].

PubMed

Zhang, Kun; Wan, Yong-shan; Liu, Feng-zhen; Zhang, Er-qun; Wang, Su

2009-12-01

To investigate the effects of shading and light recovery on the photosynthetic characteristics of peanut seedlings leaves, different shading treatments including no shading, 27% shading, 43% shading, and 77% shading were performed with black sunshade net at the seedling stage of two peanut cultivars Fenghua 1 and Fenghua 2, with related parameters determined. It was shown that with the increase of shading degree, the leaf chlorophyll content, actual PSII photochemical efficiency under irradiance (phi(PS II)), and maximum PS II photochemical efficiency (Fv/Fm) of test cultivars increased, while the Chl a/b ratio and photosynthetic rate (Pn) decreased. On the first day after light recovery, the Pn and stomatal conductance (Gs) decreased while the intercellular CO2 concentration (Ci) increased with increasing shading degree when measured under high light, but the Pn increased and the Gs and Ci decreased with increasing shading degree when measured under low light. The ratio of Pn measured under low light to that measured under high light increased significantly. With increasing shading degree, the light compensation point, light saturation point, CO2 compensation point, CO2 saturation point, and carboxylation efficiency decreased, while the apparent quantum yield increased. After the removal of shading, the Pn, phi(PS II), and Fv/Fm under natural light decreased immediately, but increased gradually 3-5 days after. 15 days after light recovery, the Pn, phi(PS II) and Fv/Fm in treatment 27% shading recovered to the level of no shading. As for the other treatments, the restored extent differed with shading degree and test variety. In the same treatments, the leaf chlorophyll content, Pn and phi(PS II) of Fenghua 1 were higher than those of Fenghua 2. The results demonstrated that shading at seedling stage improved the capabilities of test varieties in using low light, but reduced the capabilities in using high light.

Response of Artemisia annua L. to shade and manure fertilizer application in lowland altitude

NASA Astrophysics Data System (ADS)

Permana, H. H.; Widyastuti, Y.; Samanhudi; Yunus, A.

2018-03-01

Artemisia is a plant producing artemisinin substance which is the main compound in the treatment of malaria. Artemisia comes from China, usually grows wild in native habitats in the plains with an altitude of 1,000-1,500 meters above the sea level. Artemisia development efforts in Indonesia hampered by limited land with the required altitude due to their competition with vegetable crops. Based on this reason, this research is conducted to observe the growth of artemisia planted in lowland with the help of shade and manure. This study aims to determine the level of shade and best manure on the growth of Artemisia. Research conducted at the Laboratory of the Faculty of Agriculture UNS Jumantono using nested design with two factors, shade as main factor and manure fertilizer as sub factor. The data analysis used F test with confidence level of 5%, if significant, then continued with DMRT (Duncan Multiple Range Test). The results showed the treatment of shade gave no difference in growth within 50% shade, 75% shade as well as without shade treatment. Goat manure fertilizer gave the highest result and able to increase plant height, number of branches, flower weight and root volume.

Patterns of shading tolerance determined from experimental light reduction studies of seagrasses

EPA Science Inventory

An extensive review of the experimental literature on seagrass shading evaluated the relationship between experimental light reductions, duration of experiment and seagrass response metrics to determine whether there were consistent statistical patterns. There were highly signif...

Salinity stress constrains photosynthesis in Fraxinus ornus more when growing in partial shading than in full sunlight: consequences for the antioxidant defence system.

PubMed

Fini, Alessio; Guidi, Lucia; Giordano, Cristiana; Baratto, Maria Camilla; Ferrini, Francesco; Brunetti, Cecilia; Calamai, Luca; Tattini, Massimiliano

2014-09-01

A major challenge in plant ecophysiology is understanding the effects of multiple sub-optimal environmental conditions on plant performance. In most Mediterranean areas soil salinity builds up during the summer because of low availability of soil water coupled with hot temperatures. Although sunlight and soil salinity may strongly interact in determining a plant's performance, this has received relatively little attention. Two-year-old seedlings of Fraxinus ornus were grown outdoors in pots during a Mediterranean summer in either 45 % (shaded plants) or 100 % (sun plants) sunlight irradiance and were supplied with either deionized water or deionized water plus 75 mm NaCl. Morpho-anatomical traits, water and ionic relations, gas exchange and photosystem II performance, concentrations of individual carotenoids, activity of antioxidant enzymes, concentrations of ascorbic acid and individual polyphenols were measured in leaves. Leaf oxidative stress and damage were estimated by in vivo analysis of stable free radicals and ultrastructural analyses. Leaf concentrations of potentially toxic ions did not markedly differ in shaded or sun plants in response to salinity. Leaves of sun plants displayed superior water use efficiency compared with leaves of shaded plants, irrespective of salinity treatment, and had both better stomatal control and higher CO2 carboxylation efficiency than leaves of shaded plants. In the salt-treated groups, the adverse effects of excess midday irradiance were greater in shade than in sun plants. The activity of enzymes responsible for detoxifying hydrogen peroxide decreased in shaded plants and increased in sun plants as a result of salinity stress. In contrast, the activity of guaiacol peroxidase and the concentration of phenylpropanoids increased steeply in response to salinity in shaded plants but were unaffected in sun plants. It is concluded that salinity may constrain the performance of plants growing under partial shading more severely than that of plants growing under full sun during summer. The results suggest co-ordination within the antioxidant defence network aimed at detoxifying salt-induced generation of reactive oxygen species. © The Author 2014. Published by Oxford University Press on behalf of the Annals of Botany Company. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

Kynol/Nomex Fabrics for Fire Retardant Shipboard Utility Uniforms

DTIC Science & Technology

1988-06-01

changes shade permanentlj" from grey to black. The activation temperature of each set ranged from 220 F to 2800F in increments of 100F. The response of the...temperature value. When a tape reaches its activation temperature it changes shade permanent l from grey to black. The activation temperature of each...are normally only available in their natural gold color, to a suitable blue shade . g. Potential cost of Kynol/Nomex uniforms versus the FRT cotton

Photosynthetic response of Eriophorum vaginatum to in situ shrub shading in tussock tundra of northern Alaska

NASA Astrophysics Data System (ADS)

Anderson-Smith, A.; Pattison, R.; Sullivan, P.; Welker, J. M.

2009-12-01

Eriophorum vaginatum (Cotton Grass) is an important component of moist acidic tussock tundra, a plant community that appears to be undergoing changes in species composition associated with climate warming. This species is one of the most abundant in the arctic tundra, and provides important forage for caribou in their calving grounds on the Arctic Coastal Plain and along their migratory route through the foothills of Alaska. Recently, remote sensing data, repeat photography and plot-level measurements have indicated that shrub abundance is increasing while Eriophorum abundance is either constant or decreasing. One possible explanation for the reduction of Eriophorum while Betula nana is increasing, is that lower light levels in the taller Betula canopy may be constraining Eriophorum photosynthesis and subsequently reducing plant growth. This study measured the effect of shading on the light response of Eriphorum leaf photosynthesis in four different sites near Toolik Lake Alaska during the summer of 2009. Measurements were taken in: 1) a shrub patch within the drift zone of the ITEX long term snow fence experiment, 2) an LTER shade house (50% shading) built in 1989, 3) water track site 1 and water track site 2 (i.e. control areas with no experimental manipulations) Average photosynthetic rates for Eriophorum at a light level of 800 PAR varied from 3.8 to 10.9 umol m-2 s-1 and were not significantly different in shaded and unshaded areas. This study indicates that shading by shrubs does not appear to be altering the light response of Eriophorum nor does long-term shading by itself eliminate Eriophorum from the community. An alternative explanation for the decline of Eriophorum while Betula increases in abundance under changing climates may be related to plant and soil mineral nutrition, plant water relations or biotic processes involving herbivores.

Growth responses and accumulation of soluble sugars in Inga marginata Wild. (Fabaceae) subjected to flooding under contrasting light conditions.

PubMed

Bender, B; Capellesso, E S; Lottici, M E; Sentkovski, J; Mielniczki-Pereira, A A; Rosa, L M G; Sausen, T L

2017-01-01

Flood events in riparian forests of southern Brazil, can be characterized as unpredictable and of low magnitude with an average duration of less than 15 days. Inga marginata is an evergreen tree which grows in Southeast South America on a wide range of environments, including riparian forests. In this paper, the interactive effects of the light environment and soil flooding on morphological parameters of I. marginata were examined. Seedlings were acclimated in two contrasting light conditions: sun or shade for 30 days. Sun and shade plants were subjected to soil flooding for two periods; five or 15 days. After 5 days, the interaction between flooding and light did not affect growth, chlorophyll content and dry mass or the root-shoot ratio. After 15 days, flooded plants from the sunny treatment had a lower shoot dry mass compared to control sun plants and flooded plants from the shaded treatment. Moreover, the higher dry mass observed for shade plants compared to sun plants, following flooding, can also be directly associated with a higher content of soluble sugars. Shade plants of I. marginata showed a greater acclimation to soil waterlogging. This acclimation appears to be associated with a larger accumulation of soluble sugars compared to non-flooded plants. The responses observed on the shade plants appear to be decisive to indicate the use of I. marginata in degraded areas.

Detecting Subtle Shifts in Ecosystem Functioning in a Dynamic Estuarine Environment

PubMed Central

Pratt, Daniel R.; Lohrer, Andrew M.; Thrush, Simon F.; Hewitt, Judi E.; Townsend, Michael; Cartner, Katie; Pilditch, Conrad A.; Harris, Rachel J.; van Colen, Carl; Rodil, Iván F.

2015-01-01

Identifying the effects of stressors before they impact ecosystem functioning can be challenging in dynamic, heterogeneous ‘real-world’ ecosystems. In aquatic systems, for example, reductions in water clarity can limit the light available for photosynthesis, with knock-on consequences for secondary consumers, though in naturally turbid wave-swept estuaries, detecting the effects of elevated turbidity can be difficult. The objective of this study was to investigate the effects of shading on ecosystem functions mediated by sandflat primary producers (microphytobenthos) and deep-dwelling surface-feeding macrofauna (Macomona liliana; Bivalvia, Veneroida, Tellinidae). Shade cloths (which reduced incident light intensity by ~80%) were deployed on an exposed, intertidal sandflat to experimentally stress the microphytobenthic community associated with the sediment surface. After 13 weeks, sediment properties, macrofauna and fluxes of oxygen and inorganic nutrients across the sediment-water interface were measured. A multivariate metric of ecosystem function (MF) was generated by combining flux-based response variables, and distance-based linear models were used to determine shifts in the drivers of ecosystem function between non-shaded and shaded plots. No significant differences in MF or in the constituent ecosystem function variables were detected between the shaded and non-shaded plots. However, shading reduced the total explained variation in MF (from 64% in non-shaded plots to 15% in shaded plots) and affected the relative influence of M. liliana and other explanatory variables on MF. This suggests that although shade stress may shift the drivers of ecosystem functioning (consistent with earlier investigations of shading effects on sandflat interaction networks), ecosystem functions appear to have a degree of resilience to those changes. PMID:26214854

Effects of shading on starch pasting characteristics of indica hybrid rice (Oryza sativa L.).

PubMed

Wang, Li; Deng, Fei; Ren, Wan-Jun; Yang, Wen-Yu

2013-01-01

Rice is an important staple crop throughout the world, but environmental stress like low-light conditions can negatively impact crop yield and quality. Using pot experiments and field experiments, we studied the effects of shading on starch pasting viscosity and starch content with six rice varieties for three years, using the Rapid Visco Analyser to measure starch pasting viscosity. Shading at different growth stages and in different rice varieties all affected the starch pasting characteristics of rice. The effects of shading on starch pasting viscosity at middle and later growth stages were greater than those at earlier stages. Shading enhanced breakdown but reduced hold viscosity and setback at tillering-elongation stage. Most pasting parameters changed significantly with shading after elongation stage. Furthermore, the responses of different varieties to shading differed markedly. The change scope of starch pasting viscosity in Dexiang 4103 was rather small after heading, while that in IIyou 498 and Gangyou 906 was small before heading. We observed clear tendencies in peak viscosity, breakdown, and pasting temperature of the five rice varieties with shading in 2010 and 2011. Correlation analysis indicated that the rice amylose content was negatively correlated with breakdown, but was positively correlated with setback. Based on our results, IIyou 498, Gangyou 906, and Dexiang 4103 had higher shade endurance, making these varieties most suitable for high-quality rice cultivation in low-light regions.

Effects of Shading on Starch Pasting Characteristics of Indica Hybrid Rice (Oryza sativa L.)

PubMed Central

Ren, Wan-Jun; Yang, Wen-Yu

2013-01-01

Rice is an important staple crop throughout the world, but environmental stress like low-light conditions can negatively impact crop yield and quality. Using pot experiments and field experiments, we studied the effects of shading on starch pasting viscosity and starch content with six rice varieties for three years, using the Rapid Visco Analyser to measure starch pasting viscosity. Shading at different growth stages and in different rice varieties all affected the starch pasting characteristics of rice. The effects of shading on starch pasting viscosity at middle and later growth stages were greater than those at earlier stages. Shading enhanced breakdown but reduced hold viscosity and setback at tillering-elongation stage. Most pasting parameters changed significantly with shading after elongation stage. Furthermore, the responses of different varieties to shading differed markedly. The change scope of starch pasting viscosity in Dexiang 4103 was rather small after heading, while that in IIyou 498 and Gangyou 906 was small before heading. We observed clear tendencies in peak viscosity, breakdown, and pasting temperature of the five rice varieties with shading in 2010 and 2011. Correlation analysis indicated that the rice amylose content was negatively correlated with breakdown, but was positively correlated with setback. Based on our results, IIyou 498, Gangyou 906, and Dexiang 4103 had higher shade endurance, making these varieties most suitable for high-quality rice cultivation in low-light regions. PMID:23861872

The shade avoidance syndrome in Aribdopsis: the antoagonistic role of phtochrome a and B differentiates vegetation

USDA-ARS?s Scientific Manuscript database

Light limitation caused by dense vegetation is one of the greatest threats to plant survival in natural environments. Plants detect such neighboring vegetation as a reduction in the red to far-red ratio (R:FR) of the incoming light. The low R:FR signal, perceived by phytochromes, initiates a set of ...

Sunburn and sun-protective behaviors among adults with and without previous nonmelanoma skin cancer: a population-based study

PubMed Central

Fischer, Alexander H.; Wang, Timothy S.; Yenokyan, Gayane; Kang, Sewon; Chien, Anna L.

2016-01-01

Background Individuals with previous nonmelanoma skin cancer (NMSC) are at increased risk for subsequent skin cancer, and should therefore limit UV exposure. Objective To determine whether individuals with previous NMSC engage in better sun protection than those with no skin cancer history. Methods We pooled self-reported data (2005 and 2010 National Health Interview Surveys) from US non-Hispanic white adults (758 with and 34,161 without previous NMSC). We calculated adjusted prevalence odds ratios (aPOR) and 95% confidence intervals (95% CI), taking into account the complex survey design. Results Individuals with previous NMSC versus no history of NMSC had higher rates of frequent use of shade (44.3% versus 27.0%; aPOR=1.41; 1.16–1.71), long sleeves (20.5% versus 7.7%; aPOR=1.55; 1.21–1.98), a wide-brimmed hat (26.1% versus 10.5%; aPOR=1.52; 1.24–1.87), and sunscreen (53.7% versus 33.1%; aPOR=2.11; 95% CI=1.73–2.59), but did not have significantly lower odds of recent sunburn (29.7% versus 40.7%; aPOR=0.95; 0.77–1.17). Among subjects with previous NMSC, recent sunburn was inversely associated with age, sun avoidance, and shade but not sunscreen. Limitations Self-reported cross-sectional data and unavailable information quantifying regular sun exposure. Conclusion Physicians should emphasize sunburn prevention when counseling patients with previous NMSC, especially younger adults, focusing on shade and sun avoidance over sunscreen. PMID:27198078

Regulation of assimilate partitioning by daylength and spectral quality

DOE Office of Scientific and Technical Information (OSTI.GOV)

Britz, S.J.

1994-12-31

Photosynthesis is the process by which plants utilize light energy to assimilate and transform carbon dioxide into products that support growth and development. The preceding review provides an excellent summary of photosynthetic mechanisms and diurnal patterns of carbon metabolism with emphasis on the importance of gradual changes in photosynthetically-active radiation at dawn and dusk. In addition to these direct effects of irradiance, there are indirect effects of light period duration and spectral quality on carbohydrate metabolism and assimilate partitioning. Both daylength and spectral quality trigger developmental phenomena such as flowering (e.g., photoperiodism) and shade avoidance responses, but their effects onmore » partitioning of photoassimilates in leaves are less well known. Moreover, the adaptive significance to the plants of such effects is sometimes not clear.« less

Solar collector with altitude tracking

DOEpatents

Barak, Amitzur Z.

1977-01-01

A device is provided for turning a solar collector about an east-west horizontal axis so that the collector is tilted toward the sun as the EWV altitude of the sun varies each day. It includes one or more heat responsive elements and a shading means aligned so that within a range of EWV altitudes of the sun during daylight hours the shading means shades the element or elements while during the rest of the daylight hours the elements or elements are heated by the sun to assume heated, stable states. Mechanical linkage between the collector and the element is responsive to the states of the element or elements to tilt the collector in accordance with variations in the EWV altitude of the sun.

The epidermis coordinates auxin-induced stem growth in response to shade

PubMed Central

Procko, Carl; Burko, Yogev; Long, Jeff A.; Chory, Joanne

2016-01-01

Growth of a complex multicellular organism requires coordinated changes in diverse cell types. These cellular changes generate organs of the correct size, shape, and functionality. In plants, the growth hormone auxin induces stem elongation in response to shade; however, which cell types of the stem perceive the auxin signal and contribute to organ growth is poorly understood. Here, we blocked the transcriptional response to auxin within specific tissues to show that auxin signaling is required in many cell types for correct hypocotyl growth in shade, with a key role for the epidermis. Combining genetic manipulations in Arabidopsis thaliana with transcriptional profiling of the hypocotyl epidermis from Brassica rapa, we show that auxin acts in the epidermis in part by inducing activity of the locally acting, growth-promoting brassinosteroid pathway. Our findings clarify cell-specific auxin function in the hypocotyl and highlight the complexity of cell type interactions within a growing organ. PMID:27401556

Unravelling the nocturnal appearance of bogue Boops boops shoals in the anthropogenically modified shallow littoral.

PubMed

Mavraki, N; Georgiadis, M; Koutsikopoulos, C; Tzanatos, E

2016-05-01

In the present study the role of the nocturnal migration of bogue Boops boops shoals to anthropogenically modified shallow littoral locations was examined, evaluating four alternative hypotheses: (1) feeding, (2) reproduction, (3) attraction of B. boops to artificial light and (4) concealment in the darkness related to predation avoidance. All hypotheses apart from predation avoidance were rejected, as B. boops tended to concentrate in shaded locations of wider illuminated areas, a finding not only important concerning fish behaviour, but also with significant management implications. © 2016 The Fisheries Society of the British Isles.

Genetic variation in response to shade in coastal Douglas-fir.

Treesearch

J. Bradley St. Clair; Richard A. Sniezko

1999-01-01

Tree improvement programs have generally relied on testing families in open light environments. With increased interest in multiaged silvicultural systems, some people have questioned whether families selected in the open are appropriate for planting in the shade. We grew Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco var. menziesii...

Disease Vector Ecology Profile: Ecuador

DTIC Science & Technology

1998-12-01

fleas, has not been reported since 1985; however, extensive outbreaks in neighboring Peru occurred in 1992-94. Avoid rodent -infested areas and use PPM...throughout the year ( Peru ). Heavy shade in permanent bodies of water with abundant floatage. Commonly bites humans but apparently prefers rodents ...Vectors of Arboviruses other than Dengue or Yellow Fever in the Amazon Basin and Associated Northwestern Regions of South America

Lightness, chroma, and hue distributions of a shade guide as measured by a spectroradiometer.

PubMed

Lee, Yong-Keun; Yu, Bin; Lim, Ho-Nam

2010-09-01

The color attributes of commercially available shade guides have been measured by spectrophotometers (SP), which are designed to measure flat surfaces. However, there is limited information on the color distribution of shade guides as measured by spectroradiometers (SR), which are capable of measuring the color of curved surfaces. The purpose of this study was to determine the distributions of lightness (CIE L*) and chroma (C*(ab)) step intervals between adjacent shade tabs of a shade guide based on the lightness, chroma, and hue attributes measured by an SR. Lightness, chroma, hue angle, and CIE a* and b* values of the shade tabs (n=26) from a shade guide (Vitapan 3D-Master) were measured by an SR under daylight conditions. The distributions of the ratios in lightness and chroma of each tab compared with the lowest lightness tab or the lowest chroma tab were determined. The values for each color parameter were analyzed by a 3-way ANOVA with the factors of lightness, chroma, and hue designations of the shade tabs (alpha=.05). The chroma and CIE a* and b* values were influenced by the lightness, chroma, and hue designations of the shade tabs (P<.001); however, the lightness and hue angle were influenced by the lightness and hue designations, but not by the chroma designation. Distributions for the CIE a* and b* values, in each lightness group, corresponded with the chroma designation. However, the intervals in the lightness and chroma scales between adjacent tabs were not uniform. The intervals in the color parameters between adjacent shade tabs were not uniform based on SR measurements. Therefore, a shade guide in which shade tabs are more equally spaced by the color attributes, based on the values as measured by an SR along with observers' responses with respect to the equality of the intervals, should be devised. Copyright © 2010 The Editorial Council of the Journal of Prosthetic Dentistry. Published by Mosby, Inc. All rights reserved.

Modeling canopy-level productivity: is the "big-leaf" simplification acceptable?

NASA Astrophysics Data System (ADS)

Sprintsin, M.; Chen, J. M.

2009-05-01

The "big-leaf" approach to calculating the carbon balance of plant canopies assumes that canopy carbon fluxes have the same relative responses to the environment as any single unshaded leaf in the upper canopy. Widely used light use efficiency models are essentially simplified versions of the big-leaf model. Despite its wide acceptance, subsequent developments in the modeling of leaf photosynthesis and measurements of canopy physiology have brought into question the assumptions behind this approach showing that big leaf approximation is inadequate for simulating canopy photosynthesis because of the additional leaf internal control on carbon assimilation and because of the non-linear response of photosynthesis on leaf nitrogen and absorbed light, and changes in leaf microenvironment with canopy depth. To avoid this problem a sunlit/shaded leaf separation approach, within which the vegetation is treated as two big leaves under different illumination conditions, is gradually replacing the "big-leaf" strategy, for applications at local and regional scales. Such separation is now widely accepted as a more accurate and physiologically based approach for modeling canopy photosynthesis. Here we compare both strategies for Gross Primary Production (GPP) modeling using the Boreal Ecosystem Productivity Simulator (BEPS) at local (tower footprint) scale for different land cover types spread over North America: two broadleaf forests (Harvard, Massachusetts and Missouri Ozark, Missouri); two coniferous forests (Howland, Maine and Old Black Spruce, Saskatchewan); Lost Creek shrubland site (Wisconsin) and Mer Bleue petland (Ontario). BEPS calculates carbon fixation by scaling Farquhar's leaf biochemical model up to canopy level with stomatal conductance estimated by a modified version of the Ball-Woodrow-Berry model. The "big-leaf" approach was parameterized using derived leaf level parameters scaled up to canopy level by means of Leaf Area Index. The influence of sunlit/shaded leaf separation on GPP prediction was evaluated accounting for the degree of the deviation of 3-dimensional leaf spatial distribution from the random case. More specifically, we compared and evaluated the behavior of both models showing the advantages of sunlit/shaded leaf separation strategy over a simplified big-leaf approach. Keywords: canopy photosynthesis, leaf area index, clumping index, remote sensing.

A reverse genetics approach identifies novel mutants in light responses and anthocyanin metabolism in petunia.

PubMed

Berenschot, Amanda S; Quecini, Vera

2014-01-01

Flower color and plant architecture are important commercially valuable features for ornamental petunias (Petunia x hybrida Vilm.). Photoperception and light signaling are the major environmental factors controlling anthocyanin and chlorophyll biosynthesis and shade-avoidance responses in higher plants. The genetic regulators of these processes were investigated in petunia by in silico analyses and the sequence information was used to devise a reverse genetics approach to probe mutant populations. Petunia orthologs of photoreceptor, light-signaling components and anthocyanin metabolism genes were identified and investigated for functional conservation by phylogenetic and protein motif analyses. The expression profiles of photoreceptor gene families and of transcription factors regulating anthocyanin biosynthesis were obtained by bioinformatic tools. Two mutant populations, generated by an alkalyting agent and by gamma irradiation, were screened using a phenotype-independent, sequence-based method by high-throughput PCR-based assay. The strategy allowed the identification of novel mutant alleles for anthocyanin biosynthesis (CHALCONE SYNTHASE) and regulation (PH4), and for light signaling (CONSTANS) genes.

The shady side of solar protection.

PubMed

Parsons, P G; Neale, R; Wolski, P; Green, A

1998-04-06

To determine the value of shade in protecting humans from solar ultraviolet (UV) radiation. Measurement with photometers of protection factors for ultraviolet B radiation (UVB) and for total solar radiation for different types of trees and other structures during the summer months (1995-1997) in south-east Queensland. (The protection ratio is the ratio of the intensity of UVB or total solar radiation in direct sunlight to that in shade.) For summer sun at midday, the mean (SD) UV protection ratio for the shade of trees (n = 65) was 4.21 (1.36) on a horizontal surface and 1.33 (0.30) on a vertical surface. In contrast, the mean (SD) protection ratio for total solar energy (primarily infrared) was much higher (12.1 [1.4]). Trees common in recreational areas in Australia (eucalypts: UV protection ratio, 3.52 [0.79]; Norfolk Island pines: UV protection ratio, 3.72 [0.98]) offered reduced protection compared with trees with more dense foliage (UV protection ratio, 5.48 [1.44]). Over a whole day, measurement of shade by trees and other structures showed that the UV protection ratio was lower in the morning and afternoon. Shade from awnings, buildings and hats gave similar results to those for trees. Both at midday and over a whole day satisfactory protection (UV protection ratio > 15) was obtained only in shade which eliminated exposure to the sky as well as to direct sunlight; for example, in thickly wooded areas and under low, widely overhanging structures. Most forms of shade, while useful, offer people insufficient protection from solar UV. A fair-skinned person sheltering under a tree could suffer sunburn after less than one hour. There is a need for appropriate design of structural shade, use of other solar protection measures in conjunction with shade, and research on behavioural responses to shade.

Sunburn and sun-protective behaviors among adults with and without previous nonmelanoma skin cancer (NMSC): A population-based study.

PubMed

Fischer, Alexander H; Wang, Timothy S; Yenokyan, Gayane; Kang, Sewon; Chien, Anna L

2016-08-01

Individuals with previous nonmelanoma skin cancer (NMSC) are at increased risk for subsequent skin cancer, and should therefore limit ultraviolet exposure. We sought to determine whether individuals with previous NMSC engage in better sun protection than those with no skin cancer history. We pooled self-reported data (2005 and 2010 National Health Interview Surveys) from US non-Hispanic white adults (758 with and 34,161 without previous NMSC). We calculated adjusted prevalence odds ratios (aPOR) and 95% confidence intervals (CI), taking into account the complex survey design. Individuals with previous NMSC versus no history of NMSC had higher rates of frequent use of shade (44.3% vs 27.0%; aPOR 1.41; 95% CI 1.16-1.71), long sleeves (20.5% vs 7.7%; aPOR 1.55; 95% CI 1.21-1.98), a wide-brimmed hat (26.1% vs 10.5%; aPOR 1.52; 95% CI 1.24-1.87), and sunscreen (53.7% vs 33.1%; aPOR 2.11; 95% CI 1.73-2.59), but did not have significantly lower odds of recent sunburn (29.7% vs 40.7%; aPOR 0.95; 95% CI 0.77-1.17). Among those with previous NMSC, recent sunburn was inversely associated with age, sun avoidance, and shade but not sunscreen. Self-reported cross-sectional data and unavailable information quantifying regular sun exposure are limitations. Physicians should emphasize sunburn prevention when counseling patients with previous NMSC, especially younger adults, focusing on shade and sun avoidance over sunscreen. Copyright © 2016 American Academy of Dermatology, Inc. Published by Elsevier Inc. All rights reserved.

Differential responses of invasive Celastrus orbiculatus (Celastraceae) and native C. scandens to changes in light quality.

PubMed

Leicht, Stacey A; Silander, John A

2006-07-01

When plants are subjected to leaf canopy shade in forest understories or from neighboring plants, they not only experience reduced light quantity, but light quality in lowered red : far red light (R : FR). Growth and other developmental responses of plants in reduced R : FR can vary and are not consistent across species. We compared how an invasive liana, Celastrus orbiculatus, and its closely related native congener, C. scandens, responded to changes in the R : FR under controlled, simulated understory conditions. We measured a suite of morphological and growth attributes under control, neutral shading, and low R : FR light treatments. Celastrus orbiculatus showed an increase in height, aboveground biomass, and total leaf mass in reduced R : FR treatments as compared to the neutral shade, while C. scandens had increased stem diameter, single leaf area, and leaf mass to stem mass ratio. These differences provide a mechanistic understanding of the ability of C. orbiculatus to increase height and actively forage for light resources in forest understories, while C. scandens appears unable to forage for light and instead depends upon a light gap forming. The plastic growth response of C. orbiculatus in shaded conditions points to its success in forested habitats where C. scandens is largely absent.

Differential responses of invasive Celastrus orbiculatus (Celastraceae) and native C. scandens to changes in light quality

USGS Publications Warehouse

Leicht, S.A.; Silander, J.A.

2006-01-01

When plants are subjected to leaf canopy shade in forest understories or from neighboring plants, they not only experience reduced light quantity, but light quality in lowered red:far red light (R:FR). Growth and other developmental responses of plants in reduced R:FR can vary and are not consistent across species. We compared how an invasive liana, Celastrus orbiculatus, and its closely related native congener, C. scandens, responded to changes in the R:FR under controlled, simulated understory conditions. We measured a suite of morphological and growth attributes under control, neutral shading, and low R:FR light treatments. Celastrus orbiculatus showed an increase in height, aboveground biomass, and total leaf mass in reduced R:FR treatments as compared to the neutral shade, while C. scandens had increased stem diameter, single leaf area, and leaf mass to stem mass ratio. These differences provide a mechanistic understanding of the ability of C. orbiculatus to increase height and actively forage for light resources in forest understories, while C. scandens appears unable to forage for light and instead depends upon a light gap forming. The plastic growth response of C. orbiculatus in shaded conditions points to its success in forested habitats where C. scandens is largely absent.

Differential responses of double petal and multi petal jasmine to shading: I. Photosynthetic characteristics and chloroplast ultrastructure.

PubMed

Deng, Yanming; Li, Congcong; Shao, Qingsong; Ye, Xiaoqing; She, Jianming

2012-06-01

A double petal (DP) and a multi petal (MP) type jasmine (Jasminum sambac Ait.) growth and flowering was known largely affected by different levels of irradiance. Here, our objective was to determine the effects of shade on photosynthesis related characteristics and chloroplast ultrastructure of these two types. In both types, net photosynthetic rate (Pn), stomatal conductance (g(s)) and transpiration rate increased with decreasing irradiance from 100% to 20%, while both maximum and variable fluorescence showed a steady increase, and photochemical and nonphotochemical quenching indexes declined. At each conducted time, chlorophyll a, b and carotenoids contents in DP type shaded leaves increased whereas those in MP type decreased at 5% irradiance (considered as extreme shade). The maximum photochemical efficiency of photosystem II of DP plants showed subtle changes but that of MP plants declined by shading thereafter 21 days of treatment. Observation of chloroplast ultrastructure showed its best development in the leaves of DP and MP types mostly from 50% to 20% irradiance (considered as weak and moderate shade, respectively). At each shade treatment, Pn, g(s) and water use efficiency of DP-jasmine were always higher than those of MP-jasmine, thus the shade tolerance ability of the former was higher than that of the latter. The results showed that full sunlight and 5% natural irradiance caused photoinhibition and light deficiency of jasmine plants respectively, and modulating chloroplast development by the more numbers of thylakoids and grana to contain more photosynthetic pigments is an important shade tolerance mechanism of DP type. Copyright © 2012 Elsevier Masson SAS. All rights reserved.

Constrained Response Surface Optimisation and Taguchi Methods for Precisely Atomising Spraying Process

NASA Astrophysics Data System (ADS)

Luangpaiboon, P.; Suwankham, Y.; Homrossukon, S.

2010-10-01

This research presents a development of a design of experiment technique for quality improvement in automotive manufacturing industrial. The quality of interest is the colour shade, one of the key feature and exterior appearance for the vehicles. With low percentage of first time quality, the manufacturer has spent a lot of cost for repaired works as well as the longer production time. To permanently dissolve such problem, the precisely spraying condition should be optimized. Therefore, this work will apply the full factorial design, the multiple regression, the constrained response surface optimization methods or CRSOM, and Taguchi's method to investigate the significant factors and to determine the optimum factor level in order to improve the quality of paint shop. Firstly, 2κ full factorial was employed to study the effect of five factors including the paint flow rate at robot setting, the paint levelling agent, the paint pigment, the additive slow solvent, and non volatile solid at spraying of atomizing spraying machine. The response values of colour shade at 15 and 45 degrees were measured using spectrophotometer. Then the regression models of colour shade at both degrees were developed from the significant factors affecting each response. Consequently, both regression models were placed into the form of linear programming to maximize the colour shade subjected to 3 main factors including the pigment, the additive solvent and the flow rate. Finally, Taguchi's method was applied to determine the proper level of key variable factors to achieve the mean value target of colour shade. The factor of non volatile solid was found to be one more additional factor at this stage. Consequently, the proper level of all factors from both experiment design methods were used to set a confirmation experiment. It was found that the colour shades, both visual at 15 and 45 angel of measurement degrees of spectrophotometer, were nearly closed to the target and the defective at quality gate was also reduced from 0.35 WDPV to 0.10 WDPV. This reveals that the objective of this research is met and this procedure can be used as quality improvement guidance for paint shop of automotive vehicle.

Antioxidant defences and oxidative damage in salt-treated olive plants under contrasting sunlight irradiance.

PubMed

Melgar, Juan Carlos; Guidi, Lucia; Remorini, Damiano; Agati, Giovanni; Degl'innocenti, Elena; Castelli, Silvana; Camilla Baratto, Maria; Faraloni, Cecilia; Tattini, Massimiliano

2009-09-01

The interactive effects of root-zone salinity and sunlight on leaf biochemistry, with special emphasis on antioxidant defences, were analysed in Olea europaea L. cv. Allora, during the summer period. Plants were grown outside under 15% (shade plants) or 100% sunlight (sun plants) and supplied with 0 or 125 mM NaCl. The following measurements were performed: (1) the contribution of ions and soluble carbohydrates to osmotic potentials; (2) the photosystem II (PSII) photochemistry and the photosynthetic pigment concentration; (3) the concentration and the tissue-specific distribution of leaf flavonoids; (4) the activity of antioxidant enzymes; and (5) the leaf oxidative damage. The concentrations of Na(+) and Cl(-) were significantly greater in sun than in shade leaves, as also observed for the concentration of the 'antioxidant' sugar-alcohol mannitol. The de-epoxidation state of violaxanthin-cycle pigments increased in response to salinity stress in sun leaves. This finding agrees with a greater maximal PSII photochemistry (F(v)/F(m)) at midday, detected in salt-treated than in control plants, growing in full sunshine. By contrast, salt-treated plants in the shade suffered from midday depression in F(v)/F(m) to a greater degree than that observed in control plants. The high concentration of violaxanthin-cycle pigments in sun leaves suggests that zeaxanthin may protect the chloroplast from photo-oxidative damage, rather than dissipating excess excitation energy via non-photochemical quenching mechanisms. Dihydroxy B-ring-substituted flavonoid glycosides accumulate greatly in the mesophyll, not only in the epidermal cells, in response to high sunlight. The activity of antioxidant enzymes varied little because of sunlight irradiance, but declined sharply in response to high salinity in shade leaves. Interestingly, control and particularly salt-treated plants in the shade underwent greater oxidative damage than their sunny counterparts. These findings, which conform to the evolution of O. europaea in sunny environments, suggest that under partial shading, the antioxidant defence system may be ineffective to counter salt-induced oxidative damage.

Residential building energy conservation and avoided power plant emissions by urban and community trees in the United States

Treesearch

David J. Nowak; Nathaniel Appleton; Alexis Ellis; Eric Greenfield

2017-01-01

Urban trees and forests alter building energy use and associated emissions from power plants by shading buildings, cooling air temperatures and altering wind speeds around buildings. Field data on urban trees were combined with local urban/community tree and land cover maps, modeling of tree effects on building energy use and pollutant emissions, and state energy and...

"Magnetic" termite mound surfaces are oriented to suit wind and shade conditions.

PubMed

Jacklyn, Peter M

1992-09-01

The termites Amitermes meridionalis and A. laurensis construct remarkable meridional or "magnetic" mounds in northern Australia. These mounds vary geographically in mean orientation in a manner that suggests such variation is an adaptive response to local environmental conditions. Theoretical modelling of solar irradiance and mound rotation experiments show that maintenance of an eastern face temperature plateau during the dry season is the most likely physical basis for the mound orientation response. Subsequent heat transfer analysis shows that habitat wind speed and shading conditions also affect face temperature gradients such as the rate of eastern face temperature change. It is then demonstrated that the geographic variation in mean mound orientation follows the geographic variation in long-term wind speed and shading conditions across northern Australia such that an eastern face temperature plateau is maintained in all locations.

Color distribution of a shade guide in the value, chroma, and hue scale.

PubMed

Ahn, Jin-Soo; Lee, Yong-Keun

2008-07-01

Shade tabs in a shade guide are matched to teeth in the order of value, hue, and chroma; therefore, information on the distribution of shade tabs is essential for clinical application of a shade guide. However, there is limited information on the color distribution as sorted by these 3 parameters of a recently introduced shade guide. The purposes of this study were to determine the color distributions of tabs from a shade guide in the value (CIE L*), chroma (C*(ab)), and hue scale, and to determine the distribution of step intervals between adjacent tabs by value and chroma. The color of shade tabs (n=29) from a shade guide (Vitapan 3D-Master) was measured to determine the distribution of shade tabs by the value, chroma, hue angle, and CIE a* and b* values. The distribution of the ratios of the value and the chroma of each tab, when compared with the lowest value tab or the lowest chroma tab, was also determined. The data for each color parameter were analyzed by a 3-way ANOVA with the factors of value, chroma, and hue designations of the tabs (alpha=.05). The value, chroma, hue angle, and CIE a* and b* values were influenced by the value, chroma, and hue designations of shade tabs (P<.001). The distributions of the chroma of the tabs within the same value group were relatively ordered, but the values of different value groups overlapped in several instances. Distributions for the CIE a* and b* values reflected the chroma designations in each value group. In the same value group, L, M, and R hue designations corresponded with the manufacturer's stated hue, such as a yellow hue for the L designation and a red hue for the R designation. The distance in the value and chroma scales between adjacent tabs was not uniform. The color distribution of the Vitapan 3D-Master shade guide was more ordered than previously reported color distributions of other, traditional shade guides. However, the interval in the color parameters between adjacent tabs was not uniform; therefore, shade tabs spaced equally, according to the color parameters, should be studied based on the observer's response data.

DOES SUNLIGHT ENHANCE THE EFFECTIVENESS OF AVIAN PREENING FOR ECTOPARASITE CONTROL?

PubMed Central

Koop, Jennifer A. H.; Huber, Sarah K.; Clayton, Dale H.

2013-01-01

Preening is a bird’s first line of defense against harmful ectoparasites. Ectoparasites, in turn, have evolved adaptations for avoiding preening such as hardened exoskeletons and escape behavior. Earlier work suggests that some groups of ectoparasites, such as feather lice, leave hiding places in feathers that are exposed to direct sunlight, making them more vulnerable to preening. It is, therefore, conceivable that birds may choose to preen in direct sunlight, assuming it improves the effectiveness of preening. Using mourning doves and their feather lice, we tested 2 related hypotheses; (1) that birds with access to direct sunlight preen more often than birds in shade, and (2) that birds with access to direct sunlight are more effective at controlling their ectoparasites than birds in shade. To test these hypotheses, we conducted an experiment in which we manipulated both sunlight and preening ability. Our results provided no support for either hypothesis, i.e., birds given the opportunity to preen in direct sunlight did not preen significantly more often, or more effectively, than did birds in shade. Thus, the efficiency of preening for ectoparasite control appears to be independent of light intensity, at least in the case of mourning doves and their feather lice. PMID:21942474

Orbit and geometry constraints on the design and operation of a long-life SIRTF mission. [Shuttle Infrared Telescope Facility

NASA Technical Reports Server (NTRS)

Jackson, R. W.

1984-01-01

For a long-life SIRTF mission, the ability of the telescope to observe targets everywhere in the sky is an important requirement. For low-inclination orbits, a telescope aperture shade must be designed for Sun and Earth Limb avoidance angles of 50 deg to 60 deg to prevent unwanted radiation from entering the telescope. The minimum orbit inclination depends on the Earth Limb avoidance angle. About 30 percent of the sky will be prohibited for observations during any day in orbit, with about 100 days in orbit required to observe the entire sky.

The effects of shade, fertilizer, and pruning on eastern hemlock trees and hemlock woolly adelgid

Treesearch

Thomas McAvoy; Ryan Mays; Nels Johnson; Scott Salom

2017-01-01

Hemlock woolly adelgid (HWA), Adelges tsugae Annand, an invasive insect native to the Pacific Northwest and Asia, is responsible for widespread health decline and mortality of native hemlocks (Tsuga spp.) in the eastern United States. Shading and fertilizer has been found to affect the survival and health of both HWA and...

Understory Response to Disturbance: An Investigation of Prescribed Burning and Understory Removal Treatments

Treesearch

Benjamin J. Dolan; George R. Parker

2004-01-01

Lack of disturbance in the Central Hardwood Region has caused a decrease in abundance of shade-intolerant species, such as oaks (Quercus spp.) and hickories (Carya spp.), while shade-tolerant species have proliferated in the understory. The goal of this research is to determine how two disturbances, prescribed fire and mechanical...

Understory response to disturbance: an investigation of prescribed burning and understory thinning treatments

Treesearch

Benjamin J. Dolan; George R. Parker

2003-01-01

Lack of disturbance in the Central Hardwood Region has caused a decrease in abundance of shade-intolerant species, such as oaks (Quercus spp.) and hickories (Carya spp.), in the forest understory, while shade-tolerant species have proliferated. The goal of this research is to determine how two disturbances, prescribed fire and...

Response of container-grown flowering dogwood cultivars to sun/shade production regime, 2015

USDA-ARS?s Scientific Manuscript database

Flowering dogwood, Cornus florida, ‘Cherokee Brave™’ and ‘Cherokee Princess’ were grown in #5 nursery containers in an amended 100% bark substrate. Treatments were assigned based on exposure time to a full sun/shade condition during the growing season: 1) plants grown in full sun, 2) plants grown in...

Evaluation of spectral light management on growth of container-grown willow oak, nuttall oak and summer red maple

USDA-ARS?s Scientific Manuscript database

Plant response to blue, red, gray or black shade cloth was evaluated with willow oak (Quercus phellos L.), Nuttall oak (Quercus nuttallii Palmer, Nuttall) and Summer Red maple (Acer rubrum L. ‘Summer Red’) liners. Light transmitted through the colored shade cloth had no influence on germination of ...

Response of container-grown flowering dogwood cultivars to powdery mildew under sun/shade production and different fertilizer regimes

USDA-ARS?s Scientific Manuscript database

Container-grown dogwoods are an important product in the nursery and landscape industry and rank third in the US in nursery sales of ornamental trees. Dogwoods are a challenging crop to produce in container culture mainly due to fertilizer, water, and shade management. Our experiments indicate that ...

Shading responses of carbon allocation dynamics in mountain grassland

NASA Astrophysics Data System (ADS)

Bahn, M.; Lattanzi, F. A.; Brueggemann, N.; Siegwolf, R. T.; Richter, A.

2012-12-01

Carbon (C) allocation strongly influences plant and soil processes. Global environmental changes can alter source - sink relations of plants with potential implications for C allocation. Short-term C allocation dynamics in ecosystems and their responses to environmental changes are still poorly understood. To analyze effects of assimilate supply (i.e. C source strength) on ecosystem C allocation dynamics and the role of non-structural carbohydrates, canopy sections of a mountain meadow were pulse labeled with 13CO2 and subsequently shaded for a week or left unshaded (control). Tracer dynamics in above- and belowground sucrose and starch pools were analysed and coupled using compartmental modelling. The hypothesis was tested that shading affects tracer dynamics in non-structural carbohydrates and diminishes the transfer of recently assimilated C to roots and their storage pools. In unshaded plots up to 40% of assimilated C was routed through short-term storage in shoot starch and sucrose to buffer day / night cycles in photosynthesis. Shoot- and root sucrose and shoot starch were kinetically closely related pools. The tracer dynamics of the modelled root sucrose pool corresponded well with those in soil CO2 efflux. Root starch played no role in buffering day / night cycles and likely acted as a seasonal store. Shading strongly reduced sucrose and starch concentrations in shoots but not roots and resulted in a massive reduction of leaf respiration, while root respiration was much less diminished. Shading affected tracer dynamics in sucrose and starch of shoots: shoot starch rapidly lost tracer, while sucrose transiently increased its tracer content. Surprisingly, shading did not alter the dynamics of root carbohydrates. Even under severe C limitation after one week of shading, tracer C continued to be incorporated in root starch. Also the amount of 13C incorporated in phospholipid fatty acids of soil microbial communities was not reduced by shading, though its residence time followed a changed pattern, suggesting an influence of C source strength on the utilization and turnover of recent plant-derived C. These findings will be discussed in the broader context of plant and ecosystem carbon allocation, with particular reference to the concepts of 'source versus sink strength' and 'passive versus active C storage'.

Duration and intensity of shade differentially affects mycorrhizal growth- and phosphorus uptake responses of Medicago truncatula

PubMed Central

Konvalinková, Tereza; Püschel, David; Janoušková, Martina; Gryndler, Milan; Jansa, Jan

2015-01-01

Plant and fungal partners in arbuscular mycorrhizal symbiosis trade mineral nutrients for carbon, with the outcome of this relationship for plant growth and nutrition being highly context-dependent and changing with the availability of resources as well as with the specific requirements of the different partners. Here we studied how the model legume Medicago truncatula, inoculated or not with a mycorrhizal fungus Rhizophagus irregularis, responded to a gradient of light intensities applied over different periods of time, in terms of growth, phosphorus nutrition and the levels of root colonization by the mycorrhizal fungus. Short-term (6 d) shading, depending on its intensity, resulted in a rapid decline of phosphorus uptake to the shoots of mycorrhizal plants and simultaneous accumulation of phosphorus in the roots (most likely in the fungal tissues), as compared to the non-mycorrhizal controls. There was, however, no significant change in the levels of mycorrhizal colonization of roots due to short-term shading. Long-term (38 d) shading, depending on its intensity, provoked a multitude of plant compensatory mechanisms, which were further boosted by the mycorrhizal symbiosis. Mycorrhizal growth- and phosphorus uptake benefits, however, vanished at 10% of the full light intensity applied over a long-term. Levels of root colonization by the mycorrhizal fungus were significantly reduced by long-term shading. Our results indicate that even short periods of shade could have important consequences for the functioning of mycorrhizal symbiosis in terms of phosphorus transfer between the fungus and the plants, without any apparent changes in root colonization parameters or mycorrhizal growth response, and call for more focused research on temporal dynamics of mycorrhizal functioning under changing environmental conditions. PMID:25763002

Analyzing the Effect of Spinning Process Variables on Draw Frame Blended Cotton Mélange Yarn Quality

NASA Astrophysics Data System (ADS)

Ray, Suchibrata; Ghosh, Anindya; Banerjee, Debamalya

2018-06-01

An investigation has been made to study the effect of important spinning process variables namely shade depth, ring frame spindle speed and yarn twist multiplier (TM) on various yarn quality parameters like unevenness, strength, imperfection, elongation at break and hairiness index of draw frame blended cotton mélange yarn. Three factors Box and Behnken design of experiment has been used to conduct the study. The quadratic regression model is used to device the statistical inferences about sensitivity of the yarn quality parameters to the different process variables. The response surfaces are constructed for depicting the geometric representation of yarn quality parameters plotted as a function of process variables. Analysis of the results show that yarn strength of draw frame blended cotton mélange yarn is significantly affected by shade depth and TM. Yarn unevenness is affected by shade depth and ring frame spindle speed. Yarn imperfection level is mainly influenced by the shade depth and spindle speed. The shade depth and yarn TM have shown significant impact on yarn hairiness index.

Analyzing the Effect of Spinning Process Variables on Draw Frame Blended Cotton Mélange Yarn Quality

NASA Astrophysics Data System (ADS)

Ray, Suchibrata; Ghosh, Anindya; Banerjee, Debamalya

2017-12-01

An investigation has been made to study the effect of important spinning process variables namely shade depth, ring frame spindle speed and yarn twist multiplier (TM) on various yarn quality parameters like unevenness, strength, imperfection, elongation at break and hairiness index of draw frame blended cotton mélange yarn. Three factors Box and Behnken design of experiment has been used to conduct the study. The quadratic regression model is used to device the statistical inferences about sensitivity of the yarn quality parameters to the different process variables. The response surfaces are constructed for depicting the geometric representation of yarn quality parameters plotted as a function of process variables. Analysis of the results show that yarn strength of draw frame blended cotton mélange yarn is significantly affected by shade depth and TM. Yarn unevenness is affected by shade depth and ring frame spindle speed. Yarn imperfection level is mainly influenced by the shade depth and spindle speed. The shade depth and yarn TM have shown significant impact on yarn hairiness index.

Jasmonate and Phytochrome A Signaling in Arabidopsis Wound and Shade Responses Are Integrated through JAZ1 Stability[C][W

PubMed Central

Robson, Frances; Okamoto, Haruko; Patrick, Elaine; Harris, Sue-Ré; Wasternack, Claus; Brearley, Charles; Turner, John G.

2010-01-01

Jasmonate (JA) activates plant defense, promotes pollen maturation, and suppresses plant growth. An emerging theme in JA biology is its involvement in light responses; here, we examine the interdependence of the JA- and light-signaling pathways in Arabidopsis thaliana. We demonstrate that mutants deficient in JA biosynthesis and signaling are deficient in a subset of high irradiance responses in far-red (FR) light. These mutants display exaggerated shade responses to low, but not high, R/FR ratio light, suggesting a role for JA in phytochrome A (phyA) signaling. Additionally, we demonstrate that the FR light–induced expression of transcription factor genes is dependent on CORONATINE INSENSITIVE1 (COI1), a central component of JA signaling, and is suppressed by JA. phyA mutants had reduced JA-regulated growth inhibition and VSP expression and increased content of cis-(+)-12-oxophytodienoic acid, an intermediate in JA biosynthesis. Significantly, COI1-mediated degradation of JASMONATE ZIM DOMAIN1-β-glucuronidase (JAZ1-GUS) in response to mechanical wounding and JA treatment required phyA, and ectopic expression of JAZ1-GUS resulted in exaggerated shade responses. Together, these results indicate that JA and phyA signaling are integrated through degradation of the JAZ1 protein, and both are required for plant responses to light and stress. PMID:20435902

Response to Lisovski et al.

USGS Publications Warehouse

Streby, Henry M.; Kramer, Gunnar R.; Peterson, Sean M.; Lehman, Justin A.; Buehler, David A.; Andersen, David

2018-01-01

Lisovski et al. [1] describe the widely recognized limitations of light-level geolocator data for identifying short-distance latitudinal movements, recommend that caution be used when interpreting such data, intimated that we did not use such caution and argued that environmental shading likely explained the Golden-winged Warbler (Vermivora chrysoptera) movements described in our 2015 report [2] . Lisovski et al. [1] conclude that the bird movements we reported could not be disentangled from estimation error in stationary animals caused by environmental shading. We argue that, to the contrary, these hypotheses can easily be disentangled because the premise that environmental shading caused synchronous and parallel error among geolocators is false. With their assertion that our location estimates could be biased by >3,500 km on a day with no observable local sources of shading, Lisovski et al. [1] have taken a position of incredulity toward all geolocator-based animal movement data published to date.

Assessing heat load in drylot dairy cattle: Refining on-farm sampling methodology.

PubMed

Tresoldi, Grazyne; Schütz, Karin E; Tucker, Cassandra B

2016-11-01

Identifying dairy cattle experiencing heat stress and adopting appropriate mitigation strategies can improve welfare and profitability. However, little is known about how cattle use heat abatement resources (shade, sprayed water) on drylot dairies. It is also unclear how often we need to observe animals to measure high heat load, or the relevance of specific aspects of this response, particularly in terms of panting. Our objectives were to describe and determine sampling intervals to measure cattle use of heat abatement resources, respiration rate (RR) and panting characteristics (drooling, open mouth, protruding tongue), and to evaluate the relationship between the latter 2. High-producing cows were chosen from 4 drylots (8 cows/dairy, n=32) and observed for at least 5.9h (1000 to 1800h, excluding milking) when air temperature, humidity, and the combined index averaged 33°C, 30%, and 79, respectively. Use of heat abatement resources was recorded continuously; RR and the presence and absence of each panting characteristic were recorded every 5min. From the observed values, estimates using the specified sub-sampling intervals were calculated for heat abatement resource use (1, 5, 10, 15, 20, 30, 60, 90, and 120min), and for RR and panting (10, 15, 20, 30, 60, 90, and 120min). Estimates and observed values were compared using linear regression. Sampling intervals were considered accurate if they met 3 criteria: R 2 ≥0.9, intercept=0, and slope=1. The relationship between RR and each panting characteristic was analyzed using mixed models. Cows used shade (at corral or over feed bunk) and feed bunk area (where water was sprayed) for about 90 and 50% of the observed time, respectively, and used areas with no cooling for 2min at a time, on average. Cows exhibited drooling (34±4% of observations) more often than open mouth and protruding tongue (11±3 and 8±3% of observations, respectively). Respiration rate varied depending on the presence of panting (with vs. without drool present: 97±3 vs. 74±3 breaths/min; open vs. closed mouth: 104±4 vs. 85±4 breaths/min; protruding vs. non-protruding tongue: 105±5 vs. 91±5 breaths/min). Accurate estimates were obtained when using sampling intervals ≤90min for RR, ≤60min for corral shade and sprayed water use, and ≤30min for drooling. In a hot and dry climate, cows kept in drylots had higher RR when showing panting characteristics than when these were absent, and used shade extensively, avoiding areas with no cooling. In general, 30min intervals were most efficient for measuring heat load responses. Copyright © 2016 American Dairy Science Association. Published by Elsevier Inc. All rights reserved.

RNAseq reveals weed-induced PIF3-like as a candidate target to manipulate weed stress response in soybean.

PubMed

Horvath, David P; Hansen, Stephanie A; Moriles-Miller, Janet P; Pierik, Ronald; Yan, Changhui; Clay, David E; Scheffler, Brian; Clay, Sharon A

2015-07-01

Weeds reduce yield in soybeans (Glycine max) through incompletely defined mechanisms. The effects of weeds on the soybean transcriptome were evaluated in field conditions during four separate growing seasons. RNASeq data were collected from six biological samples of soybeans growing with or without weeds. Weed species and the methods to maintain weed-free controls varied between years to mitigate treatment effects, and to allow detection of general soybean weed responses. Soybean plants were not visibly nutrient- or water-stressed. We identified 55 consistently downregulated genes in weedy plots. Many of the downregulated genes were heat shock genes. Fourteen genes were consistently upregulated. Several transcription factors including a PHYTOCHROME INTERACTING FACTOR 3-like gene (PIF3) were included among the upregulated genes. Gene set enrichment analysis indicated roles for increased oxidative stress and jasmonic acid signaling responses during weed stress. The relationship of this weed-induced PIF3 gene to genes involved in shade avoidance responses in Arabidopsis provide evidence that this gene may be important in the response of soybean to weeds. These results suggest that the weed-induced PIF3 gene will be a target for manipulating weed tolerance in soybean. No claim to original US government works New Phytologist © 2015 New Phytologist Trust.

Changes in light intensity reveal a major role for carbon balance in Arabidopsis responses to high temperature.

PubMed

Vasseur, François; Pantin, Florent; Vile, Denis

2011-09-01

High temperature (HT) is a major limiting factor for plant productivity. Because some responses to HT, notably hyponasty, resemble those encountered in low light (LL), we hypothesized that plant responses to HT are under the control of carbon balance. We analysed the interactive effects of HT and irradiance level on hyponasty and a set of traits related to plant growth in natural accessions of Arabidopsis thaliana and mutants affected in heat dissipation through transpiration (NCED6-OE, ost2) and starch metabolism (pgm). HT induced hyponasty, reduced plant growth and modified leaf structure. LL worsened the effects of HT, while increasing light restored trait values close to levels observed at control temperature. Leaf temperature per se did not play a major role in the observed responses. By contrast, a major role of carbon balance was supported by hyponastic growth of pgm, as well as morphological, physiological (photosynthesis, sugar and starch contents) and transcriptional data. Carbon balance could be a common sensor of HT and LL, leading to responses specific of the shade avoidance syndrome. Hyponasty and associated changes in plant traits could be key traits conditioning plant performance under competition for light, particularly in warm environments. © 2011 Blackwell Publishing Ltd.

Response of sun-grown and shade-grown northern red oak seedlings to outplanting in clearcuts and shelterwoods in North Alabama

Treesearch

Callie Jo Schweitzer; Emile S. Gardiner; David L. Loftis

2006-01-01

The primary objective of this study was to determine if greenhouse light environment would affect outplanting success for northern red oak (Quercus rubra L.) in clearcuts and shelterwoods. In 2002, northern red oak seedlings were grown from acorns under full-ambient (sun) and half-ambient (shade) light conditions in a greenhouse. Seedlings grown...

The effects of providing portable shade at pasture on dairy cow behavior and physiology.

PubMed

Palacio, S; Bergeron, R; Lachance, S; Vasseur, E

2015-09-01

Access to pasture has advantages for cows such as reduced lameness and improved udder health, but also may expose cows to stressors such as extreme heat. The objective of this study was to understand how portable shade affected physiological and behavioral responses of pastured dairy cows in a Canadian summer. Over 8wk, a total of 24 lactating Holstein cows were separated into 2 treatments, one with access to shade and a control without access to shade. The cows were pastured in groups of 4, with 3 field sections per treatment. Instantaneous scan sampling of behaviors (drinking, lying, grazing, other) performed in the shade or not were recorded every 5min for 3h/d during the hottest part of the day (peak hours: 1130-1530h) 3d/wk. Ambient temperature, humidity, and vaginal temperature were recorded at 10-min intervals. Daily milk production was also recorded. Differences between treatments by week were analyzed using the generalized linear mixed model with group as random effect and treatment as fixed effect. Cows with shade access were observed at the water trough up to 6.42 times less and lying down up to 1.75 times more. Cows with shade access grazed up to 1.5 times more but only when the temperature-humidity index was above their comfort threshold (≥72) during the hottest part of the day (wk 2). Cows sought shade when it was made available, but spent less than half of their time observed (%) in the shade (40.8±4.67) with the exception of wk 2 when most of the time was spent under the shade (74.3±4.77). Daily lying time was highest during peak hours for cows with shade access. However, no overall difference in total lying time between the 2 treatments was observed. No differences were found in vaginal temperature or milk production between treatments with the exception of wk 1 for daily milk production, which was higher for cows in the control treatment. In conclusion, cows sought shade when it was provided at pasture, whereas cows without access to shade seemed to alter their behavior to cope with heat stress, as seen from the lack of physiological differences between treatments. The results indicate that providing cows with access to pasture under a temperate climate does not seem to have any detrimental physiological or production effects and providing them with shade can potentially decrease production costs and help with water conservation strategies as fewer cows were observed at the water when shade was provided. Copyright © 2015 American Dairy Science Association. Published by Elsevier Inc. All rights reserved.

De novo assembly and characterization of fruit transcriptome in Litchi chinensis Sonn and analysis of differentially regulated genes in fruit in response to shading

PubMed Central

2013-01-01

Background Litchi (Litchi chinensis Sonn.) is one of the most important fruit trees cultivated in tropical and subtropical areas. However, a lack of transcriptomic and genomic information hinders our understanding of the molecular mechanisms underlying fruit set and fruit development in litchi. Shading during early fruit development decreases fruit growth and induces fruit abscission. Here, high-throughput RNA sequencing (RNA-Seq) was employed for the de novo assembly and characterization of the fruit transcriptome in litchi, and differentially regulated genes, which are responsive to shading, were also investigated using digital transcript abundance(DTA)profiling. Results More than 53 million paired-end reads were generated and assembled into 57,050 unigenes with an average length of 601 bp. These unigenes were annotated by querying against various public databases, with 34,029 unigenes found to be homologous to genes in the NCBI GenBank database and 22,945 unigenes annotated based on known proteins in the Swiss-Prot database. In further orthologous analyses, 5,885 unigenes were assigned with one or more Gene Ontology terms, 10,234 hits were aligned to the 24 Clusters of Orthologous Groups classifications and 15,330 unigenes were classified into 266 Kyoto Encyclopedia of Genes and Genomes pathways. Based on the newly assembled transcriptome, the DTA profiling approach was applied to investigate the differentially expressed genes related to shading stress. A total of 3.6 million and 3.5 million high-quality tags were generated from shaded and non-shaded libraries, respectively. As many as 1,039 unigenes were shown to be significantly differentially regulated. Eleven of the 14 differentially regulated unigenes, which were randomly selected for more detailed expression comparison during the course of shading treatment, were identified as being likely to be involved in the process of fruitlet abscission in litchi. Conclusions The assembled transcriptome of litchi fruit provides a global description of expressed genes in litchi fruit development, and could serve as an ideal repository for future functional characterization of specific genes. The DTA analysis revealed that more than 1000 differentially regulated unigenes respond to the shading signal, some of which might be involved in the fruitlet abscission process in litchi, shedding new light on the molecular mechanisms underlying organ abscission. PMID:23941440

Effect of Late Planting and Shading on Cellulose Synthesis during Cotton Fiber Secondary Wall Development

PubMed Central

Chen, Ji; Lv, Fengjuan; Liu, Jingran; Ma, Yina; Wang, Youhua; Chen, Binglin; Meng, Yali; Zhou, Zhiguo; Oosterhuis, Derrick M.

2014-01-01

Cotton-rapeseed or cotton-wheat double cropping systems are popular in the Yangtze River Valley and Yellow River Valley of China. Due to the competition of temperature and light resources during the growing season of double cropping system, cotton is generally late-germinating and late-maturing and has to suffer from the coupling of declining temperature and low light especially in the late growth stage. In this study, late planting (LP) and shading were used to fit the coupling stress, and the coupling effect on fiber cellulose synthesis was investigated. Two cotton (Gossypium hirsutum L.) cultivars were grown in the field in 2010 and 2011 at three planting dates (25 April, 25 May and 10 June) each with three shading levels (normal light, declined 20% and 40% PAR). Mean daily minimum temperature was the primary environmental factor affected by LP. The coupling of LP and shading (decreased cellulose content by 7.8%–25.5%) produced more severe impacts on cellulose synthesis than either stress alone, and the effect of LP (decreased cellulose content by 6.7%–20.9%) was greater than shading (decreased cellulose content by 0.7%–5.6%). The coupling of LP and shading hindered the flux from sucrose to cellulose by affecting the activities of related cellulose synthesis enzymes. Fiber cellulose synthase genes expression were delayed under not only LP but shading, and the coupling of LP and shading markedly postponed and even restrained its expression. The decline of sucrose-phosphate synthase activity and its peak delay may cause cellulose synthesis being more sensitive to the coupling stress during the later stage of fiber secondary wall development (38–45 days post-anthesis). The sensitive difference of cellulose synthesis between two cultivars in response to the coupling of LP and shading may be mainly determined by the sensitiveness of invertase, sucrose-phosphate synthase and cellulose synthase. PMID:25133819

The Photochemical Reflectance Index from Directional Cornfield Reflectances: Observations and Simulations

NASA Technical Reports Server (NTRS)

Cheng, Yen-Ben; Middleton, Elizabeth M.; Zhang, Qingyuan; Corp, Lawrence A.; Dandois, Jonathan; Kustas, William P.

2012-01-01

The two-layer Markov chain Analytical Canopy Reflectance Model (ACRM) was linked with in situ hyperspectral leaf optical properties to simulate the Photochemical Reflectance Index (PRI) for a corn crop canopy at three different growth stages. This is an extended study after a successful demonstration of PRI simulations for a cornfield previously conducted at an early vegetative growth stage. Consistent with previous in situ studies, sunlit leaves exhibited lower PRI values than shaded leaves. Since sunlit (shaded) foliage dominates the canopy in the reflectance hotspot (coldspot), the canopy PRI derived from field hyperspectral observations displayed sensitivity to both view zenith angle and relative azimuth angle at all growth stages. Consequently, sunlit and shaded canopy sectors were most differentiated when viewed along the azimuth matching the solar principal plane. These directional PRI responses associated with sunlit/shaded foliage were successfully reproduced by the ACRM. As before, the simulated PRI values from the current study were closer to in situ values when both sunlit and shaded leaves were utilized as model input data in a two-layer mode, instead of a one-layer mode with sunlit leaves only. Model performance as judged by correlation between in situ and simulated values was strongest for the mature corn crop (r = 0.87, RMSE = 0.0048), followed by the early vegetative stage (r = 0.78; RMSE = 0.0051) and the early senescent stage (r = 0.65; RMSE = 0.0104). Since the benefit of including shaded leaves in the scheme varied across different growth stages, a further analysis was conducted to investigate how variable fractions of sunlit/shaded leaves affect the canopy PRI values expected for a cornfield, with implications for 20 remote sensing monitoring options. Simulations of the sunlit to shaded canopy ratio near 50/50 +/- 10 (e.g., 60/40) matching field observations at all growth stages were examined. Our results suggest in the importance of the sunlit/shaded fraction and canopy structure in understanding and interpreting PRI.

Fine Root Abundance and Dynamics of Stone Pine (Pinus cembra) at the Alpine Treeline Is Not Impaired by Self-shading.

PubMed

Kubisch, Petra; Leuschner, Christoph; Coners, Heinz; Gruber, Andreas; Hertel, Dietrich

2017-01-01

Low temperatures are crucial for the formation of the alpine treeline worldwide. Since soil temperature in the shade of tree canopies is lower than in open sites, it was assumed that self-shading may impair the trees' root growth performance. While experiments with tree saplings demonstrate root growth impairment at soil temperatures below 5-7°C, field studies exploring the soil temperature - root growth relationship at the treeline are missing. We recorded soil temperature and fine root abundance and dynamics in shaded and sun-exposed areas under canopies of isolated Pinus cembra trees at the alpine treeline. In contrast to the mentioned assumption, we found more fine root biomass and higher fine root growth in colder than in warmer soil areas. Moreover, colder areas showed higher fine root turnover and thus lower root lifespan than warmer places. We conclude that P. cembra balances enhanced fine root mortality in cold soils with higher fine root activity and by maintaining higher fine root biomass, most likely as a response to shortage in soil resource supply. The results from our study highlight the importance of in situ measurements on mature trees to understand the fine root response and carbon allocation pattern to the thermal growth conditions at the alpine treeline.

Ensemble composition and activity levels of insectivorous bats in response to management intensification in coffee agroforestry systems.

PubMed

Williams-Guillén, Kimberly; Perfecto, Ivette

2011-01-26

Shade coffee plantations have received attention for their role in biodiversity conservation. Bats are among the most diverse mammalian taxa in these systems; however, previous studies of bats in coffee plantations have focused on the largely herbivorous leaf-nosed bats (Phyllostomidae). In contrast, we have virtually no information on how ensembles of aerial insectivorous bats--nearly half the Neotropical bat species--change in response to habitat modification. To evaluate the effects of agroecosystem management on insectivorous bats, we studied their diversity and activity in southern Chiapas, Mexico, a landscape dominated by coffee agroforestry. We used acoustic monitoring and live captures to characterize the insectivorous bat ensemble in forest fragments and coffee plantations differing in the structural and taxonomic complexity of shade trees. We captured bats of 12 non-phyllostomid species; acoustic monitoring revealed the presence of at least 12 more species of aerial insectivores. Richness of forest bats was the same across all land-use types; in contrast, species richness of open-space bats increased in low shade, intensively managed coffee plantations. Conversely, only forest bats demonstrated significant differences in ensemble structure (as measured by similarity indices) across land-use types. Both overall activity and feeding activity of forest bats declined significantly with increasing management intensity, while the overall activity, but not feeding activity, of open-space bats increased. We conclude that diverse shade coffee plantations in our study area serve as valuable foraging and commuting habitat for aerial insectivorous bats, and several species also commute through or forage in low shade coffee monocultures.

Canopy gradients in leaf functional traits for species that differ in growth strategies and shade tolerance.

PubMed

Coble, Adam P; Fogel, Marilyn L; Parker, Geoffrey G

2017-10-01

In temperate deciduous forests, vertical gradients in leaf mass per area (LMA) and area-based leaf nitrogen (Narea) are strongly controlled by gradients in light availability. While there is evidence that hydrostatic constraints on leaf development may diminish LMA and Narea responses to light, inherent differences among tree species may also influence leaf developmental and morphological response to light. We investigated vertical gradients in LMA, Narea and leaf carbon isotope composition (δ13C) for three temperate deciduous species (Carpinus caroliniana Walter, Fagus grandifolia Ehrh., Liriodendron tulipifera L.) that differed in growth strategy (e.g., indeterminate and determinate growth), shade tolerance and leaf area to sapwood ratio (Al:As). Leaves were sampled across a broad range of light conditions within three vertical layers of tree crowns to maximize variation in light availability at each height and to minimize collinearity between light and height. All species displayed similar responses to light with respect to Narea and δ13C, but not for LMA. Light was more important for gradients in LMA for the shade-tolerant (C. caroliniana) and -intolerant (L. tulipifera) species with indeterminate growth, and height (e.g., hydrostatic gradients) and light were equally important for the shade-tolerant (F. grandifolia) species with determinate growth. Fagus grandifolia had a higher morphological plasticity in response to light, which may offer a competitive advantage in occupying a broader range of light conditions throughout the canopy. Differences in responses to light and height for the taller tree species, L. tulipifera and F. grandifolia, may be attributed to differences in growth strategy or Al:As, which may alter morphological and functional responses to light availability. While height was important in F. grandifolia, height was no more robust in predicting LMA than light in any of the species, confirming the strong role of light availability in determining LMA for temperate deciduous species. © The Author 2017. Published by Oxford University Press. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

Crassulacean acid metabolism in the shade. Studies on an epiphytic fern, Pyrrosia longifolia, and other rainforest species from Australia.

PubMed

Winter, K; Osmond, C B; Hubick, K T

1986-01-01

Crassulacean acid metabolism (CAM) was studied in a tropical epiphytic fern, Pyrrosia longifolia, from a fully sun-exposed and from a very shaded site in Northern Queensland, Australia. Measurements of instantaneous net CO 2 exchange showed carbon gain via CO 2 dark fixation with some net CO 2 uptake also occuring during late afternoon, in both sun and shade fronds. Maximum rates of net CO 2 uptake and the nocturnal increase in titratable acidity were lower in shade than in sun fronds. δ 13 C values of sun and shade fronds were not significantly different, and ranged between-14 and-15‰ suggesting that, in the long term, carbon gain was mainly via CO 2 dark fixation. Sun fronds had a higher light compensation point of photosynthesis than shade fronds but the same quantum yield. Yet there was no acclimation of photosynthetic O 2 evolution, (measured at 5% CO 2 ) in sun and shade fronds and photosynthesis saturated at between 200 and 400 μmol quanta m -2 s -1 . Use of higher light intensities for photosynthesis of sun fronds was probably precluded by low nutrient availability. Total nitrogen was less than 1% of dry weight in fully expanded sun and shade fronds. Exposure of shade fronds to full sunlight for 6 h led to a 60% decline in the quantum yield of photosynthesis and to a decline in variable fluorescence measured at room temperature. Photoinhibition by high light was also observed in Hoya nicholsoniae, a rainforest climber growing in deep shade. This species also exhibited CAM as demonstrated by nocturnal net CO 2 uptake, nocturnal acidification and a δ 13 C value of-14‰. Photosynthetic O 2 evolution in this species was saturated at 2.5% of full sunlight. Two species of Dendrobium (Orchidaceae) from sun-exposed sites, one species exhibiting CAM and the other one exhibiting net CO 2 uptake exclusively during daytime via conventional C 3 photosynthesis, showed similar light response curves and the same quantum yield for photosynthetic O 2 evolution.

Association of Indoor Tanning Frequency With Risky Sun Protection Practices and Skin Cancer Screening.

PubMed

Fischer, Alexander H; Wang, Timothy S; Yenokyan, Gayane; Kang, Sewon; Chien, Anna L

2016-10-12

Indoor tanning is prevalent among young adults and women and is associated with increased risk of melanoma. Evidence suggests that indoor tanners may be more inclined to adopt poor photoprotective practices that further increase their risk of skin cancer; however, gaps in the literature exist in young adults and by indoor tanning frequency. To examine the association between indoor tanning frequency and behaviors related to skin cancer prevention and to investigate whether these associations vary by age group or sex. Cross-sectional population-based study of US 2015 National Health Interview Survey data including 10 262 non-Hispanic white adults aged 18 to 60 years without a history of skin cancer. Rare/never use of sunscreen, protective clothing, shade; multiple sunburns within the past year; previous full-body skin examination. Of the 10 262 individuals in our study population (49% female; median age, 39 y), 787 (7.0%) reported having tanned indoors in the past year. Among individuals aged 18 to 34 years, frequent indoor tanners (≥10 times in the past year) were more likely to report never/rare use of protective clothing (adjusted prevalence ratio [aPR], 1.28; 95% CI, 1.10-1.49) and shade (aPR, 1.32; 95% CI, 1.03-1.70), compared with individuals who did not tan indoors. Among women aged 18 to 60 years, those who frequently tanned indoors were more likely to rarely/never use sunscreen (aPR, 1.34; 95% CI, 1.11-1.62), protective clothing (aPR, 1.27; 95% CI, 1.15-1.42), and shade (aPR, 1.54; 95% CI, 1.25-1.90) on a warm sunny day, as well as more likely to report multiple sunburns in the past year (aPR, 1.21; 95% CI, 1.00-1.45) compared with those who did not tan indoors. Individuals who tanned indoors in the past year were not significantly more likely to have undergone a previous full-body skin examination in any subpopulation examined. Individuals who tan indoors often exhibited a concurrent tendency to sunburn, avoid sun protection, and avoid skin cancer screening. Thus, the findings highlight that in addition to tanning bed avoidance, it is critical to emphasize sun protection and skin cancer screening in individuals who tan indoors.

Leaf non-structural carbohydrate allocation and C:N:P stoichiometry in response to light acclimation in seedlings of two subtropical shade-tolerant tree species.

PubMed

Xie, Hongtao; Yu, Mukui; Cheng, Xiangrong

2018-03-01

Light availability greatly affects plant growth and development. In shaded environments, plants must respond to reduced light intensity to ensure a regular rate of photosynthesis to maintain the dynamic balance of nutrients, such as leaf non-structural carbohydrates (NSCs), carbon (C), nitrogen (N) and phosphorus (P). To improve our understanding of the nutrient utilization strategies of understory shade-tolerant plants, we compared the variations in leaf NSCs, C, N and P in response to heterogeneous controlled light conditions between two subtropical evergreen broadleaf shade-tolerant species, Elaeocarpus sylvestris (E. sylvestris) and Illicium henryi (I. henryi). Light intensity treatments were applied at five levels (100%, 52%, 33%, 15% and 6% full sunlight) for 30 weeks to identify the effects of reduced light intensity on leaf NSC allocation patterns and leaf C:N:P stoichiometry characteristics. We found that leaf soluble sugar, starch and NSC concentrations in E. sylvestris showed decreasing trends with reduced light intensity, whereas I. henryi presented slightly increasing trends from 100% to 15% full sunlight and then significant decreases at extremely low light intensity (6% full sunlight). The soluble sugar/starch ratio of E. sylvestris decreased with decreasing light intensity, whereas that of I. henryi remained stable. Moreover, both species exhibited increasing trends in leaf N and P concentrations but limited leaf N:P and C:P ratio fluctuations with decreasing light intensity, revealing their adaptive strategies for poor light environments and their growth strategies under ideal light environments. There were highly significant correlations between leaf NSC variables and C:N:P stoichiometric variables in both species, revealing a trade-off in photosynthesis production between leaf NSC and carbon allocation. Thus, shade-tolerant plants readjusted their allocation of leaf NSCs, C, N and P in response to light acclimation. Redundancy analysis showed that leaf morphological features of both E. sylvestris and I. henryi affected their corresponding leaf nutrient traits. These results improve our understanding of the dynamic balance between leaf NSCs and leaf C, N and P components in the nutritional metabolism of shade-tolerant plants. Two species of understory shade-tolerant plants responded differently to varying light intensities in terms of leaf non-structural carbohydrate allocation and the utilization of carbon, nitrogen and phosphorus to balance nutritional metabolism and adapt to environmental stress. Copyright © 2018 Elsevier Masson SAS. All rights reserved.

Functional traits and plasticity in response to light in seedlings of four Iberian forest tree species.

PubMed

Sánchez-Gómez, David; Valladares, Fernando; Zavala, Miguel A

2006-11-01

We investigated the differential roles of physiological and morphological features on seedling survivorship along an experimental irradiance gradient in four dominant species of cool temperate-Mediterranean forests (Quercus robur L., Quercus pyrenaica Willd., Pinus sylvestris L. and Pinus pinaster Ait.). The lowest photochemical efficiency (F(v)/F(m) in dark-adapted leaves) was reached in deep shade (1% of full sunlight) in all species except Q. robur, which had the lowest photochemical efficiency in both deep shade and 100% of full sunlight. Species differed significantly in their survival in 1% of full sunlight but exhibited similar survivorship in 6, 20 and 100% of full sunlight. Shade-tolerant oaks had lower leaf area ratios, shoot to root ratios, foliage allocation ratios and higher rates of allocation to structural biomass (stem plus thick roots) than shade-intolerant pines. Overall phenotypic plasticity for each species, estimated as the difference between the minimum and the maximum mean values of the ecophysiological variables studied at the various irradiances divided by the maximum mean value of those variables, was inversely correlated with shade tolerance. Observed morphology, allocation and plasticity conformed to a conservative resource-use strategy, although observed differences in specific leaf area, which was higher in shade-tolerant species, supported a carbon gain maximization strategy. Lack of a congruent suite of traits underlying shade tolerance in the studied species provides evidence of adaptation to multiple selective forces. Although the study was based on only four species, the importance of ecophysiological variables as determinants of interspecific differences in survival in limiting light was demonstrated.

Ecological filtering by a dominant herb selects for shade tolerance in the tree seedling community of coastal dune forest.

PubMed

Tsvuura, Zivanai; Griffiths, Megan E; Gunton, Richard M; Franks, Peter J; Lawes, Michael J

2010-12-01

The regeneration niche is commonly partitioned along a gradient from shade-tolerant to shade-intolerant species to explain plant community assembly in forests. We examined the shade tolerance of tree seedlings in a subtropical coastal forest to determine whether the ecological filtering effect of a dominant, synchronously monocarpic herb (Isoglossa woodii) selects for species at either end of the light response continuum during the herb's vegetative and reproductive phases. Photosynthetic characteristics of seedlings of 20 common tree species and the herb were measured. Seedlings were grown in the greenhouse at 12-14% irradiance, and their light compensation points measured using an open-flow gas exchange system. The light compensation points for the tree species were low, falling within a narrow range from 2.1 ± 0.8 μmol m(-2) s(-1) in Celtis africana to 6.4 ± 0.7 μmol m(-2) s(-1) in Allophylus natalensis, indicating general shade tolerance, consistent with a high and narrow range of apparent quantum yield among species (0.078 ± 0.002 mol CO(2) mol(-1) photon). Rates of dark respiration were significantly lower in a generalist pioneer species (Acacia karroo) than in a forest pioneer (C. africana), or in late successional phase forest species. We argue that the general shade tolerance, and phenotypic clustering of shade tolerance, in many tree species from several families in this system, is a result of ecological filtering by the prevailing low light levels beneath the I. woodii understorey, which excludes most light-demanding species from the seedling community.

Impact of shade on outdoor thermal comfort—a seasonal field study in Tempe, Arizona

NASA Astrophysics Data System (ADS)

Middel, Ariane; Selover, Nancy; Hagen, Björn; Chhetri, Nalini

2016-12-01

Shade plays an important role in designing pedestrian-friendly outdoor spaces in hot desert cities. This study investigates the impact of photovoltaic canopy shade and tree shade on thermal comfort through meteorological observations and field surveys at a pedestrian mall on Arizona State University's Tempe campus. During the course of 1 year, on selected clear calm days representative of each season, we conducted hourly meteorological transects from 7:00 a.m. to 6:00 p.m. and surveyed 1284 people about their thermal perception, comfort, and preferences. Shade lowered thermal sensation votes by approximately 1 point on a semantic differential 9-point scale, increasing thermal comfort in all seasons except winter. Shade type (tree or solar canopy) did not significantly impact perceived comfort, suggesting that artificial and natural shades are equally efficient in hot dry climates. Globe temperature explained 51 % of the variance in thermal sensation votes and was the only statistically significant meteorological predictor. Important non-meteorological factors included adaptation, thermal comfort vote, thermal preference, gender, season, and time of day. A regression of subjective thermal sensation on physiological equivalent temperature yielded a neutral temperature of 28.6 °C. The acceptable comfort range was 19.1 °C-38.1 °C with a preferred temperature of 20.8 °C. Respondents exposed to above neutral temperature felt more comfortable if they had been in air-conditioning 5 min prior to the survey, indicating a lagged response to outdoor conditions. Our study highlights the importance of active solar access management in hot urban areas to reduce thermal stress.

Phenotypic plasticity of Neonotonia wightii and Pueraria phaseoloides grown under different light intensities.

PubMed

Santos, Leonardo D T; Da Cruz, Leandro R; Dos Santos, Samuel A; Sant'anna-Santos, Bruno F; Dos Santos, Izabela T; De Oliveira, Ariane M; Barros, Rodrigo E; Santos, Márcia V; Faria, Rodrigo M

2015-03-01

Plants have the ability to undergo morphophysiological changes based on availability of light. The present study evaluated biomass accumulation, leaf morphoanatomy and physiology of Neonotonia wightii and Pueraria phaseoloides grown in full sunlight, as well as in 30% and 50% shade. Two assays were performed, one for each species, using a randomized block design with 10 replicates. A higher accumulation of fresh mass in the shoot of the plants was observed for both species under cultivation in 50% shade, while no differences were detected between the full sunlight and 30% shade. N. wightii and P. phaseoloides showed increase in area and reduction in thickness leaf when cultivated in 50% shade. There were no changes in photosynthetic rate, stomatal conductance, water use efficiency and evapotranspiration of P. phaseoloides plants because growth environment. However, the shade treatments caused alterations in physiological parameters of N. wightii. In both species, structural changes in the mesophyll occurred depending on the availability of light; however, the amount of leaf blade tissue remained unaltered. Despite the influence of light intensity variation on the morphophysiological plasticity of N. wightii and P. phaseoloides, no effects on biomass accumulation were observed in response to light.

[Response of photosynthesis and growth to weak light regime in different Adzuki bean (Vigna angularis) varieties].

PubMed

Zou, Chang-ming; Wang, Yun-qing; Cao, Wei-dong; Liu, Ying; Zhang, Xiao-hong; Tang, Shan

2015-12-01

In order to determine the adaptability of Adzuki beans as the interplanting crops in fruit yards, field and pot experimental treatments with full natural light and weak light (48% of full natural light) regimes were conducted to test the shade tolerance and physiological responses of three Adzuki bean varieties including Funan green Vigna angularis (FGVA), early-mature black V. angularis (EBVA) and late-mature black V. angularis (LBVA). The leaf photosynthetic characteristic parameters, photosynthetic pigment contents and the activity of RuBPCase were measured during the first bloom stage. The response of growth to weak light was likewise studied. The results showed that the photosynthetic characteristic parameters, i.e., the maximum net photosynthetic rate, light saturation point and light compensation point of the three Adzuki bean varieties under the weak light stress changed differently. The weak light stress induced the reduction of net photosynthetic rate, water use efficiency and RuBPCase activity of the three Adzuki bean varieties significantly. The contents of chlorophyll a and chlorophyll b in leaves of FGVA increased significantly, while Chl a/b and carotenoid content in the leaves decreased significantly after shading. But the other two varieties did not change obviously in photosynthetic pigments content after shading. The weak light changed the growth of the three Adzuki bean varieties, such as decreasing dry matter yield and dry matter accumulation efficiency, reducing root nodule and root-shoot ratio, debasing leaves quantity and leaf area index. The first bloom stage and maturing stage of FGVA advanced, while that of EBVA delayed under weak light. However, flowers were not strong enough to seed for LBVA under the weak light. In conclusion, according to the photosynthetic characteristics changes after shading, as well as the growth status, we concluded that the shade tolerance of the three Adzuki beans was ranked as FGVA>EBVA>LBVA.

[Allergic responses to date palm and pecan pollen in Israel].

PubMed

Waisel, Y; Keynan, N; Gil, T; Tayar, D; Bezerano, A; Goldberg, A; Geller-Bernstein, C; Dolev, Z; Tamir, R; Levy, I

1994-03-15

Date palm (Phoenix dactylifera) and pecan (Carya illinoensis) trees are commonly planted in Israel for fruit, for shade, or as ornamental plants. Pollen grains of both species are allergenic; however, the extent of exposure to such pollen and the incidence of allergic response have not been studied here. We therefore investigated skin-test responses to pollen extracts of 12 varieties of palm and 9 of pecan in 705 allergic patients living in 3 cities and 19 rural settlements. Sensitivity to the pollen extracts of both species was much higher among residents of rural than of urban communities. Moreover, there was a definite relationship between the abundance of these trees in a region and the incidence of skin responders to their pollen. Sensitivity was frequent in settlements rich in these 2 species, such as those with nearby commercial date or pecan plantations. In general, sensitivity to date pollen extracts was lower than to pecan. However, differences in skin responses to pollen extracts of various clones were substantiated. Air sampling revealed that pollen pollution decreased considerably with distance from the trees. At approximately 100 m from a source concentrations of airborne pollen were low. Since planting of male palm and pecan trees in population centers would increase pollen pollution, it should be avoided.

An RNA-Seq Analysis of Grape Plantlets Grown in vitro Reveals Different Responses to Blue, Green, Red LED Light, and White Fluorescent Light

PubMed Central

Li, Chun-Xia; Xu, Zhi-Gang; Dong, Rui-Qi; Chang, Sheng-Xin; Wang, Lian-Zhen; Khalil-Ur-Rehman, Muhammad; Tao, Jian-Min

2017-01-01

Using an RNA sequencing (RNA-seq) approach, we analyzed the differentially expressed genes (DEGs) and physiological behaviors of “Manicure Finger” grape plantlets grown in vitro under white, blue, green, and red light. A total of 670, 1601, and 746 DEGs were identified in plants exposed to blue, green, and red light, respectively, compared to the control (white light). By comparing the gene expression patterns with the growth and physiological responses of the grape plantlets, we were able to link the responses of the plants to light of different spectral wavelengths and the expression of particular sets of genes. Exposure to red and green light primarily triggered responses associated with the shade-avoidance syndrome (SAS), such as enhanced elongation of stems, reduced investment in leaf growth, and decreased chlorophyll levels accompanied by the expression of genes encoding histone H3, auxin repressed protein, xyloglucan endotransglycosylase/hydrolase, the ELIP protein, and microtubule proteins. Furthermore, specific light treatments were associated with the expression of a large number of genes, including those involved in the glucan metabolic pathway and the starch and sucrose metabolic pathways; these genes were up/down-regulated in ways that may explain the increase in the starch, sucrose, and total sugar contents in the plants. Moreover, the enhanced root growth and up-regulation of the expression of defense genes accompanied with SAS after exposure to red and green light may be related to the addition of 30 g/L sucrose to the culture medium of plantlets grown in vitro. In contrast, blue light induced the up-regulation of genes related to microtubules, serine carboxypeptidase, chlorophyll synthesis, and sugar degradation and the down-regulation of auxin-repressed protein as well as a large number of resistance-related genes that may promote leaf growth, improve chlorophyll synthesis and chloroplast development, increase the ratio of chlorophyll a (chla)/chlorophyll b (chlb), and decrease the ratio of carbohydrates to proteins in plants. Although exposure to red and green light seems to impose “shade stress” on the plantlets, growth under blue light is comparable to growth observed under white or broad-spectrum light. PMID:28197159

Ambient occlusion effects for combined volumes and tubular geometry.

PubMed

Schott, Mathias; Martin, Tobias; Grosset, A V Pascal; Smith, Sean T; Hansen, Charles D

2013-06-01

This paper details a method for interactive direct volume rendering that computes ambient occlusion effects for visualizations that combine both volumetric and geometric primitives, specifically tube-shaped geometric objects representing streamlines, magnetic field lines or DTI fiber tracts. The algorithm extends the recently presented the directional occlusion shading model to allow the rendering of those geometric shapes in combination with a context providing 3D volume, considering mutual occlusion between structures represented by a volume or geometry. Stream tube geometries are computed using an effective spline-based interpolation and approximation scheme that avoids self-intersection and maintains coherent orientation of the stream tube segments to avoid surface deforming twists. Furthermore, strategies to reduce the geometric and specular aliasing of the stream tubes are discussed.

Ambient Occlusion Effects for Combined Volumes and Tubular Geometry

PubMed Central

Schott, Mathias; Martin, Tobias; Grosset, A.V. Pascal; Smith, Sean T.; Hansen, Charles D.

2013-01-01

This paper details a method for interactive direct volume rendering that computes ambient occlusion effects for visualizations that combine both volumetric and geometric primitives, specifically tube-shaped geometric objects representing streamlines, magnetic field lines or DTI fiber tracts. The algorithm extends the recently presented the directional occlusion shading model to allow the rendering of those geometric shapes in combination with a context providing 3D volume, considering mutual occlusion between structures represented by a volume or geometry. Stream tube geometries are computed using an effective spline-based interpolation and approximation scheme that avoids self-intersection and maintains coherent orientation of the stream tube segments to avoid surface deforming twists. Furthermore, strategies to reduce the geometric and specular aliasing of the stream tubes are discussed. PMID:23559506

Adaptive smart simulator for characterization and MPPT construction of PV array

NASA Astrophysics Data System (ADS)

Ouada, Mehdi; Meridjet, Mohamed Salah; Dib, Djalel

2016-07-01

Partial shading conditions are among the most important problems in large photovoltaic array. Many works of literature are interested in modeling, control and optimization of photovoltaic conversion of solar energy under partial shading conditions, The aim of this study is to build a software simulator similar to hard simulator and to produce a shading pattern of the proposed photovoltaic array in order to use the delivered information to obtain an optimal configuration of the PV array and construct MPPT algorithm. Graphical user interfaces (Matlab GUI) are built using a developed script, this tool is easy to use, simple, and has a rapid of responsiveness, the simulator supports large array simulations that can be interfaced with MPPT and power electronic converters.

Evidence that enforced sunlight exposure can cause hyperthermia in cattle ingesting low levels of ergot of rye (Claviceps purpurea), when air temperature and humidity conditions are only moderate.

PubMed

Bourke, C A

2003-09-01

To determine the rectal temperature response of cattle, following the oral administration of ergot of rye (Claviceps purpurea), under pen conditions of enforced sunlight compared with those of enforced shade. Hereford cross steers were divided into two groups of 18. One group was dosed once, on a Monday morning, with finely ground rye grass ergots at a rate of 180 mg/kg body weight and held in the sun for 7 h each day until Friday afternoon. The other group was not dosed but was similarly held in the sun during the same period. Their rectal temperatures were measured early morning and mid afternoon, from Monday to Friday inclusive. The process was repeated for each group, but this time they were held in the shade. The four treatment options were run concurrently by conducting the experiments over 6 weeks and using 3 animals in each treatment group, each week. The thermic response over all weeks, of the ergot treated, sunlight exposed cattle, was deemed greater than for the other groups, based on the following four parameters. The increase in rectal temperature between early morning and mid afternoon, the size of the mid afternoon rectal temperature rise, the difference between the maximum mid afternoon rectal temperature recorded by an animal in the sun compared with that recorded by the same animal in the shade, and finally the number of animals in a treatment group that recorded rectal temperatures > 40.00 degrees C. The difference in the daily increase in body temperature between the ergot treated, sun exposed cattle and the ergot treated, shaded cattle, was greater than that observed between the sun exposed and shade restricted control cattle. Nine of 18 ergot treated and sun exposed cattle developed hyperthermia; no cattle in the other three groups did. Some sunlight exposed cattle, dosed with a low amount of ergot of rye, can experience a body temperature elevation above the normal range, even under mild ambient temperature and humidity conditions. Sunlight exposure causes an increase in body temperature in normal cattle that is probably attributable to solar radiant heat. Ergot ingestion causes an increase in body temperature in shaded cattle that is probably attributable to ambient temperature and humidity effects. This response can be increased by sun exposure and this increase is attributable to solar radiant heat effects and possibly also solar radiation. A response to radiant heat is consistent with the known physiological effects of ingested ergot alkaloids, but a response to solar radiation is not.

Physiological Analysis of Phototropic Responses in Arabidopsis.

PubMed

Zeidler, Mathias

2016-01-01

Plants utilize light as sole energy source. To maximize light capture they are able to detect the light direction and orient themselves towards the light source. This phototropic response is mediated by the plant blue light photoreceptors phototropin1 and 2 (phot1 and phot2). Although fully differentiated plants also exhibit this response it can be best observed in etiolated seedlings. Differences in light between the illuminated and shaded site of a seedling stem lead to changes in the auxin-distribution, resulting in cell elongation on the shaded site. Since phototropism connects light perception, signaling, and auxin transport, it is of great interest to analyze this response with a fast and simple method.Here we describe a method to analyze the phototropic response of Arabidopsis seedlings. With numerous mutants available, its fast germination and its small size Arabidopsis is well suited for this analysis. Different genotypes can be simultaneously probed in less than a week.

Post-veraison sunlight exposure induces MYB-mediated transcriptional regulation of anthocyanin and flavonol synthesis in berry skins of Vitis vinifera

PubMed Central

Matus, José Tomás; Loyola, Rodrigo; Vega, Andrea; Peña-Neira, Alvaro; Bordeu, Edmundo; Arce-Johnson, Patricio; Alcalde, José Antonio

2009-01-01

Anthocyanins, flavan-3-ols, and flavonols are the three major classes of flavonoid compounds found in grape berry tissues. Several viticultural practices increase flavonoid content in the fruit, but the underlying genetic mechanisms responsible for these changes have not been completely deciphered. The impact of post-veraison sunlight exposure on anthocyanin and flavonol accumulation in grape berry skin and its relation to the expression of different transcriptional regulators known to be involved in flavonoid synthesis was studied. Treatments consisting of removing or moving aside the basal leaves which shade berry clusters were applied. Shading did not affect sugar accumulation or gene expression of HEXOSE TRANSPORTER 1, although in the leaf removal treatment, these events were retarded during the first weeks of ripening. Flavonols were the most drastically reduced flavonoids following shading and leaf removal treatments, related to the reduced expression of FLAVONOL SYNTHASE 4 and its putative transcriptional regulator MYB12. Anthocyanin accumulation and the expression of CHS2, LDOX, OMT, UFGT, MYBA1, and MYB5a genes were also affected. Other regulatory genes were less affected or not affected at all by these treatments. Non-transcriptional control mechanisms for flavonoid synthesis are also suggested, especially during the initial stages of ripening. Although berries from the leaf removal treatment received more light than shaded fruits, malvidin-3-glucoside and total flavonol content was reduced compared with the treatment without leaf removal. This work reveals that flavonol-related gene expression responds rapidly to field changes in light levels, as shown by the treatment in which shaded fruits were exposed to light in the late stages of ripening. Taken together, this study establishes MYB-specific responsiveness for the effect of sun exposure and sugar transport on flavonoid synthesis. PMID:19129169

Post-veraison sunlight exposure induces MYB-mediated transcriptional regulation of anthocyanin and flavonol synthesis in berry skins of Vitis vinifera.

PubMed

Matus, José Tomás; Loyola, Rodrigo; Vega, Andrea; Peña-Neira, Alvaro; Bordeu, Edmundo; Arce-Johnson, Patricio; Alcalde, José Antonio

2009-01-01

Anthocyanins, flavan-3-ols, and flavonols are the three major classes of flavonoid compounds found in grape berry tissues. Several viticultural practices increase flavonoid content in the fruit, but the underlying genetic mechanisms responsible for these changes have not been completely deciphered. The impact of post-veraison sunlight exposure on anthocyanin and flavonol accumulation in grape berry skin and its relation to the expression of different transcriptional regulators known to be involved in flavonoid synthesis was studied. Treatments consisting of removing or moving aside the basal leaves which shade berry clusters were applied. Shading did not affect sugar accumulation or gene expression of HEXOSE TRANSPORTER 1, although in the leaf removal treatment, these events were retarded during the first weeks of ripening. Flavonols were the most drastically reduced flavonoids following shading and leaf removal treatments, related to the reduced expression of FLAVONOL SYNTHASE 4 and its putative transcriptional regulator MYB12. Anthocyanin accumulation and the expression of CHS2, LDOX, OMT, UFGT, MYBA1, and MYB5a genes were also affected. Other regulatory genes were less affected or not affected at all by these treatments. Non-transcriptional control mechanisms for flavonoid synthesis are also suggested, especially during the initial stages of ripening. Although berries from the leaf removal treatment received more light than shaded fruits, malvidin-3-glucoside and total flavonol content was reduced compared with the treatment without leaf removal. This work reveals that flavonol-related gene expression responds rapidly to field changes in light levels, as shown by the treatment in which shaded fruits were exposed to light in the late stages of ripening. Taken together, this study establishes MYB-specific responsiveness for the effect of sun exposure and sugar transport on flavonoid synthesis.

Attenuation of the jasmonate burst, plant defensive traits, and resistance to specialist monarch caterpillars on shaded common milkweed (Asclepias syriaca).

PubMed

Agrawal, Anurag A; Kearney, Emily E; Hastings, Amy P; Ramsey, Trey E

2012-07-01

Plant responses to herbivory and light competition are often in opposing directions, posing a potential conflict for plants experiencing both stresses. For sun-adapted species, growing in shade typically makes plants more constitutively susceptible to herbivores via reduced structural and chemical resistance traits. Nonetheless, the impact of light environment on induced resistance has been less well-studied, especially in field experiments that link physiological mechanisms to ecological outcomes. Accordingly, we studied induced resistance of common milkweed (Asclepias syriaca, a sun-adapted plant), and linked hormonal responses, resistance traits, and performance of specialist monarch caterpillars (Danaus plexippus) in varying light environments. In natural populations, plants growing under forest-edge shade showed reduced levels of resistance traits (lower leaf toughness, cardenolides, and trichomes) and enhanced light-capture traits (higher specific leaf area, larger leaves, and lower carbon-to-nitrogen ratio) compared to paired plants in full sun. In a field experiment repeated over two years, only milkweeds growing in full sun exhibited induced resistance to monarchs, whereas plants growing in shade were constitutively more susceptible and did not induce resistance. In a more controlled field experiment, plant hormones were higher in the sun (jasmonic acid, salicylic acid, abscisic acid, indole acidic acid) and were induced by herbivory (jasmonic acid and abscisic acid). In particular, the jasmonate burst following herbivory was halved in plants raised in shaded habitats, and this correspondingly reduced latex induction (but not cardenolide induction). Thus, we provide a mechanistic basis for the attenuation of induced plant resistance in low resource environments. Additionally, there appears to be specificity in these interactions, with light-mediated impacts on jasmonate-induction being stronger for latex exudation than cardenolides.

The effect of interspecific variation in photosynthetic plasticity on 4-year growth rate and 8-year survival of understorey tree seedlings in response to gap formations in a cool-temperate deciduous forest.

PubMed

Oguchi, Riichi; Hiura, Tsutom; Hikosaka, Kouki

2017-08-01

Gap formation increases the light intensity in the forest understorey. The growth responses of seedlings to the increase in light availability show interspecific variation, which is considered to promote biodiversity in forests. At the leaf level, some species increase their photosynthetic capacity in response to gap formation, whereas others do not. Here we address the question of whether the interspecific difference in the photosynthetic response results in the interspecific variation in the growth response. If so, the interspecific difference in photosynthetic response would also contribute to species coexistence in forests. We also address the further relevant question of why some species do not increase their photosynthetic capacity. We assumed that some cost of photosynthetic plasticity may constrain acquisition of the plasticity in some species, and hypothesized that species with larger photosynthetic plasticity exhibit better growth after gap formation and lower survivorship in the shade understorey of a cool-temperate deciduous forest. We created gaps by felling canopy trees and studied the relationship between the photosynthetic response and the subsequent growth rate of seedlings. Naturally growing seedlings of six deciduous woody species were used and their mortality was examined for 8 years. The light-saturated rate of photosynthesis (Pmax) and the relative growth rate (RGR) of the seedlings of all study species increased at gap plots. The extent of these increases varied among the species. The stimulation of RGR over 4 years after gap formation was strongly correlated with change in photosynthetic capacity of newly expanded leaves. The increase in RGR and Pmax correlated with the 8-year mortality at control plots. These results suggest a trade-off between photosynthetic plasticity and the understorey shade tolerance. Gap-demanding species may acquire photosynthetic plasticity, sacrificing shade tolerances, whereas gap-independent species may acquire shade tolerances, sacrificing photosynthetic plasticity. This strategic difference among species would contribute to species coexistence in cool-temperate deciduous forests. © The Author 2017. Published by Oxford University Press. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

Color blindness and contrast perception in cuttlefish (Sepia officinalis) determined by a visual sensorimotor assay.

PubMed

Mäthger, Lydia M; Barbosa, Alexandra; Miner, Simon; Hanlon, Roger T

2006-05-01

We tested color perception based upon a robust behavioral response in which cuttlefish (Sepia officinalis) respond to visual stimuli (a black and white checkerboard) with a quantifiable, neurally controlled motor response (a body pattern). In the first experiment, we created 16 checkerboard substrates in which 16 grey shades (from white to black) were paired with one green shade (matched to the maximum absorption wavelength of S. officinalis' sole visual pigment, 492 nm), assuming that one of the grey shades would give a similar achromatic signal to the tested green. In the second experiment, we created a checkerboard using one blue and one yellow shade whose intensities were matched to the cuttlefish's visual system. In both assays it was tested whether cuttlefish would show disruptive coloration on these checkerboards, indicating their ability to distinguish checkers based solely on wavelength (i.e., color). Here, we show clearly that cuttlefish must be color blind, as they showed non-disruptive coloration on the checkerboards whose color intensities were matched to the Sepia visual system, suggesting that the substrates appeared to their eyes as uniform backgrounds. Furthermore, we show that cuttlefish are able to perceive objects in their background that differ in contrast by approximately 15%. This study adds support to previous reports that S. officinalis is color blind, yet the question of how cuttlefish achieve "color-blind camouflage" in chromatically rich environments still remains.

Fine Root Abundance and Dynamics of Stone Pine (Pinus cembra) at the Alpine Treeline Is Not Impaired by Self-shading

PubMed Central

Kubisch, Petra; Leuschner, Christoph; Coners, Heinz; Gruber, Andreas; Hertel, Dietrich

2017-01-01

Low temperatures are crucial for the formation of the alpine treeline worldwide. Since soil temperature in the shade of tree canopies is lower than in open sites, it was assumed that self-shading may impair the trees’ root growth performance. While experiments with tree saplings demonstrate root growth impairment at soil temperatures below 5–7°C, field studies exploring the soil temperature – root growth relationship at the treeline are missing. We recorded soil temperature and fine root abundance and dynamics in shaded and sun-exposed areas under canopies of isolated Pinus cembra trees at the alpine treeline. In contrast to the mentioned assumption, we found more fine root biomass and higher fine root growth in colder than in warmer soil areas. Moreover, colder areas showed higher fine root turnover and thus lower root lifespan than warmer places. We conclude that P. cembra balances enhanced fine root mortality in cold soils with higher fine root activity and by maintaining higher fine root biomass, most likely as a response to shortage in soil resource supply. The results from our study highlight the importance of in situ measurements on mature trees to understand the fine root response and carbon allocation pattern to the thermal growth conditions at the alpine treeline. PMID:28469633

Impact of Photovoltaic Canopy Shade on Outdoor Thermal Comfort in a Hot Desert City

NASA Astrophysics Data System (ADS)

Middel, Ariane; Selover, Nancy; Hagen, Björn; Chhetri, Nalini

2016-04-01

Shade plays an important role in designing pedestrian-friendly outdoor spaces in hot desert cities. This study investigates the impact of photovoltaic canopy shade on thermal comfort through microclimate observations and field surveys at a pedestrian mall on Arizona State University's Tempe campus. Six stationary sensors under solar canopies and in nearby sun-exposed and tree-shaded locations monitored 5-min temperature and humidity for a year. On selected clear calm days representative of each season, we conducted hourly microclimate transects from 7:00AM to 6:00PM and surveyed 1284 people about their thermal perception, comfort, and preferences. Shade lowered thermal sensation votes by approximately 1 point on the Likert scale, increasing thermal comfort in all seasons except winter. The shade type (tree or solar canopy) did not significantly impact perceived comfort, suggesting that artificial and natural shade are equally efficient in semi-arid desert environments. Globe temperature explained 50% of the variance in thermal sensation votes and was the only statistically significant meteorological predictor. Important non-meteorological factors include adaptation level, gender, thermal comfort vote, thermal preference, season, and time of day. A regression of perceived comfort on Physiological Equivalent Temperature yielded a neutral temperature of 28.6°C. The acceptable comfort range was 19.1°C-38.1°C with a preferred temperature of 20.8°C. Respondents exposed to above neutral temperatures felt more comfortable if they had been in air-conditioning 5 minutes prior to the survey, indicating a lagged response to outdoor conditions. Our study highlights the importance of active solar access management in hot urban areas.

Effect of clonal integration on nitrogen cycling in rhizosphere of rhizomatous clonal plant, Phyllostachys bissetii, under heterogeneous light.

PubMed

Li, Yang; Chen, Jing-Song; Xue, Ge; Peng, Yuanying; Song, Hui-Xing

2018-07-01

Clonal integration plays an important role in clonal plant adapting to heterogeneous habitats. It was postulated that clonal integration could exhibit positive effects on nitrogen cycling in the rhizosphere of clonal plant subjected to heterogeneous light conditions. An in-situ experiment was conducted using clonal fragments of Phyllostachys bissetii with two successive ramets. Shading treatments were applied to offspring or mother ramets, respectively, whereas counterparts were treated to full sunlight. Rhizomes between two successive ramets were either severed or connected. Extracellular enzyme activities and nitrogen turnover were measured, as well as soil properties. Abundance of functional genes (archaeal or bacterial amoA, nifH) in the rhizosphere of shaded, offspring or mother ramets were determined using quantitative polymerase chain reaction. Carbon or nitrogen availabilities were significantly influenced by clonal integration in the rhizosphere of shaded ramets. Clonal integration significantly increased extracellular enzyme activities and abundance of functional genes in the rhizosphere of shaded ramets. When rhizomes were connected, higher nitrogen turnover (nitrogen mineralization or nitrification rates) was exhibited in the rhizosphere of shaded offspring ramets. However, nitrogen turnover was significantly decreased by clonal integration in the rhizosphere of shaded mother ramets. Path analysis indicated that nitrogen turnover in the rhizosphere of shaded, offspring or mother ramets were primarily driven by the response of soil microorganisms to dissolved organic carbon or nitrogen. This unique in-situ experiment provided insights into the mechanism of nutrient recycling mediated by clonal integration. It was suggested that effects of clonal integration on the rhizosphere microbial processes were dependent on direction of photosynthates transport in clonal plant subjected to heterogeneous light conditions. Copyright © 2018 Elsevier B.V. All rights reserved.

Impact of shade on outdoor thermal comfort-a seasonal field study in Tempe, Arizona.

PubMed

Middel, Ariane; Selover, Nancy; Hagen, Björn; Chhetri, Nalini

2016-12-01

Shade plays an important role in designing pedestrian-friendly outdoor spaces in hot desert cities. This study investigates the impact of photovoltaic canopy shade and tree shade on thermal comfort through meteorological observations and field surveys at a pedestrian mall on Arizona State University's Tempe campus. During the course of 1 year, on selected clear calm days representative of each season, we conducted hourly meteorological transects from 7:00 a.m. to 6:00 p.m. and surveyed 1284 people about their thermal perception, comfort, and preferences. Shade lowered thermal sensation votes by approximately 1 point on a semantic differential 9-point scale, increasing thermal comfort in all seasons except winter. Shade type (tree or solar canopy) did not significantly impact perceived comfort, suggesting that artificial and natural shades are equally efficient in hot dry climates. Globe temperature explained 51 % of the variance in thermal sensation votes and was the only statistically significant meteorological predictor. Important non-meteorological factors included adaptation, thermal comfort vote, thermal preference, gender, season, and time of day. A regression of subjective thermal sensation on physiological equivalent temperature yielded a neutral temperature of 28.6 °C. The acceptable comfort range was 19.1 °C-38.1 °C with a preferred temperature of 20.8 °C. Respondents exposed to above neutral temperature felt more comfortable if they had been in air-conditioning 5 min prior to the survey, indicating a lagged response to outdoor conditions. Our study highlights the importance of active solar access management in hot urban areas to reduce thermal stress.

Hierarchical rendering of trees from precomputed multi-layer z-buffers

DOE Office of Scientific and Technical Information (OSTI.GOV)

Max, N.

1996-02-01

Chen and Williams show how precomputed z-buffer images from different fixed viewing positions can be reprojected to produce an image for a new viewpoint. Here images are precomputed for twigs and branches at various levels in the hierarchical structure of a tree, and adaptively combined, depending on the position of the new viewpoint. The precomputed images contain multiple z levels to avoid missing pixels in the reconstruction, subpixel masks for anti-aliasing, and colors and normals for shading after reprojection.

Tolerance to solar ultraviolet-B radiation in the citrus red mite, an upper surface user of host plant leaves.

PubMed

Fukaya, Midori; Uesugi, Ryuji; Ohashi, Hirokazu; Sakai, Yuta; Sudo, Masaaki; Kasai, Atsushi; Kishimoto, Hidenari; Osakabe, Masahiro

2013-01-01

Plant-dwelling mites are potentially exposed to solar ultraviolet-B (UVB) radiation that causes deleterious and often lethal effects, leading most mites to inhabit the lower (underside) leaf surfaces. However, in species of spider mite belonging to the Genus Panonychus, a substantial portion of individuals occur on upper leaf surfaces. We investigated whether the upper leaf surfaces of citrus trees are favorable for P. citri, and to what extent they are tolerant to UVB radiation. If eggs are not adequately protected from UVB damage, females may avoid ovipositing on the upper surfaces of sunny leaves. To test this, we conducted laboratory experiments using a UVB lamp, and semioutdoor manipulative experiments. As a result, P. citri eggs are tolerant to UVB. Field studies revealed that the ratio of eggs and adult females on upper leaf surfaces were larger for shaded than for sunny leaves. However, 64-89% of eggs hatched successfully even on sunny upper leaf surfaces. Nutritional evaluation revealed that whether on sunny or shaded leaves, in fecundity and juvenile development P. citri reaped the fitness benefits of upper leaf surfaces. Consequently, P. citri is tolerant to UVB damage, and inhabiting the upper surfaces of shaded leaves is advantageous to this mite. © 2012 Wiley Periodicals, Inc. Photochemistry and Photobiology © 2012 The American Society of Photobiology.

Reliability and accuracy of four dental shade-matching devices.

PubMed

Kim-Pusateri, Seungyee; Brewer, Jane D; Davis, Elaine L; Wee, Alvin G

2009-03-01

There are several electronic shade-matching instruments available for clinical use, but the reliability and accuracy of these instruments have not been thoroughly investigated. The purpose of this in vitro study was to evaluate the reliability and accuracy of 4 dental shade-matching instruments in a standardized environment. Four shade-matching devices were tested: SpectroShade, ShadeVision, VITA Easyshade, and ShadeScan. Color measurements were made of 3 commercial shade guides (Vitapan Classical, Vitapan 3D-Master, and Chromascop). Shade tabs were placed in the middle of a gingival matrix (Shofu GUMY) with shade tabs of the same nominal shade from additional shade guides placed on both sides. Measurements were made of the central region of the shade tab positioned inside a black box. For the reliability assessment, each shade tab from each of the 3 shade guide types was measured 10 times. For the accuracy assessment, each shade tab from 10 guides of each of the 3 types evaluated was measured once. Differences in reliability and accuracy were evaluated using the Standard Normal z test (2 sided) (alpha=.05) with Bonferroni correction. Reliability of devices was as follows: ShadeVision, 99.0%; SpectroShade, 96.9%; VITA Easyshade, 96.4%; and ShadeScan, 87.4%. A significant difference in reliability was found between ShadeVision and ShadeScan (P=.008). All other comparisons showed similar reliability. Accuracy of devices was as follows: VITA Easyshade, 92.6%; ShadeVision, 84.8%; SpectroShade, 80.2%; and ShadeScan, 66.8%. Significant differences in accuracy were found between all device pairs (P<.001) for all comparisons except for SpectroShade versus ShadeVision (P=.033). Most devices had similar high reliability (over 96%), indicating predictable shade values from repeated measurements. However, there was more variability in accuracy among devices (67-93%), and differences in accuracy were seen with most device comparisons.

Reduction of shading-derived artifacts in skin chromophore imaging without measurements or assumptions about the shape of the subject

NASA Astrophysics Data System (ADS)

Yoshida, Kenichiro; Nishidate, Izumi; Ojima, Nobutoshi; Iwata, Kayoko

2014-01-01

To quantitatively evaluate skin chromophores over a wide region of curved skin surface, we propose an approach that suppresses the effect of the shading-derived error in the reflectance on the estimation of chromophore concentrations, without sacrificing the accuracy of that estimation. In our method, we use multiple regression analysis, assuming the absorbance spectrum as the response variable and the extinction coefficients of melanin, oxygenated hemoglobin, and deoxygenated hemoglobin as the predictor variables. The concentrations of melanin and total hemoglobin are determined from the multiple regression coefficients using compensation formulae (CF) based on the diffuse reflectance spectra derived from a Monte Carlo simulation. To suppress the shading-derived error, we investigated three different combinations of multiple regression coefficients for the CF. In vivo measurements with the forearm skin demonstrated that the proposed approach can reduce the estimation errors that are due to shading-derived errors in the reflectance. With the best combination of multiple regression coefficients, we estimated that the ratio of the error to the chromophore concentrations is about 10%. The proposed method does not require any measurements or assumptions about the shape of the subjects; this is an advantage over other studies related to the reduction of shading-derived errors.

Effects of shade tab arrangement on the repeatability and accuracy of shade selection.

PubMed

Yılmaz, Burak; Yuzugullu, Bulem; Cınar, Duygu; Berksun, Semih

2011-06-01

Appropriate and repeatable shade matching using visual shade selection remains a challenge for the restorative dentist. The purpose of this study was to evaluate the effect of different arrangements of a shade guide on the repeatability and accuracy of visual shade selection by restorative dentists. Three Vitapan Classical shade guides were used for shade selection. Seven shade tabs from one shade guide were used as target shades for the testing (A1, A4, B2, B3, C2, C4, and D3); the other 2 guides were used for shade selection by the subjects. One shade guide was arranged according to hue and chroma and the second was arranged according to value. Thirteen male and 22 female restorative dentists were asked to match the target shades using shade guide tabs arranged in the 2 different orders. The sessions were performed twice with each guide in a viewing booth. Collected data were analyzed with Fisher's exact test to compare the accuracy and repeatability of the shade selection (α=.05). There were no significant differences observed in the accuracy or repeatability of the shade selection results obtained with the 2 different arrangements. When the hue/chroma-ordered shade guide was used, 58% of the shade selections were accurate. This ratio was 57.6% when the value-ordered shade guide was used. The observers repeated 55.5% of the selections accurately with the hue/chroma-ordered shade guide and 54.3% with the value-ordered shade guide. The accuracy and repeatability of shade selections by restorative dentists were similar when different arrangements (hue/chroma-ordered and value-ordered) of the Vitapan Classical shade guide were used. Copyright © 2011 The Editorial Council of the Journal of Prosthetic Dentistry. Published by Mosby, Inc. All rights reserved.

Adaptive smart simulator for characterization and MPPT construction of PV array

DOE Office of Scientific and Technical Information (OSTI.GOV)

Ouada, Mehdi, E-mail: mehdi.ouada@univ-annaba.org; Meridjet, Mohamed Salah; Dib, Djalel

2016-07-25

Partial shading conditions are among the most important problems in large photovoltaic array. Many works of literature are interested in modeling, control and optimization of photovoltaic conversion of solar energy under partial shading conditions, The aim of this study is to build a software simulator similar to hard simulator and to produce a shading pattern of the proposed photovoltaic array in order to use the delivered information to obtain an optimal configuration of the PV array and construct MPPT algorithm. Graphical user interfaces (Matlab GUI) are built using a developed script, this tool is easy to use, simple, and hasmore » a rapid of responsiveness, the simulator supports large array simulations that can be interfaced with MPPT and power electronic converters.« less

Responses of four Indo-West Pacific seagrass species to shading.

PubMed

Collier, Catherine J; Waycott, Michelle; Ospina, Ana Giraldo

2012-01-01

Seagrasses of the Great Barrier Reef predominantly occur in coastal regions where terrestrial inputs modify water quality and photosynthetic light is highly variable. Responses to shading were tested for Cymodocea serrulata, Halodule uninervis, Thalassia hemprichii and Zostera muelleri. In aquaria, four light treatments - high (66% surface light), moderate (31%), low (14%) and very low light (1%) treatments - were applied for 102d. Stress responses in the low and very low light treatments occurred in the following sequence: metabolic and physiological changes (reduced growth, increased pigment concentrations and photosynthetic efficiency); shedding (leaf loss, shoot loss) and production of new, altered tissue (leaves with reduced length, width and thickness). Complete shoot loss was projected after 76 (Z. muelleri) to 130d (T. hemprichii). Responses were slower in the low than in the very low treatment, therefore, efforts to minimize water quality degradation will be rewarded with delayed impacts to seagrasses. Copyright © 2011 Elsevier Ltd. All rights reserved.

Lizards fail to plastically adjust nesting behavior or thermal tolerance as needed to buffer populations from climate warming.

PubMed

Telemeco, Rory S; Fletcher, Brooke; Levy, Ofir; Riley, Angela; Rodriguez-Sanchez, Yesenia; Smith, Colton; Teague, Collin; Waters, Amanda; Angilletta, Michael J; Buckley, Lauren B

2017-03-01

Although observations suggest the potential for phenotypic plasticity to allow adaptive responses to climate change, few experiments have assessed that potential. Modeling suggests that Sceloporus tristichus lizards will need increased nest depth, shade cover, or embryonic thermal tolerance to avoid reproductive failure resulting from climate change. To test for such plasticity, we experimentally examined how maternal temperatures affect nesting behavior and embryonic thermal sensitivity. The temperature regime that females experienced while gravid did not affect nesting behavior, but warmer temperatures at the time of nesting reduced nest depth. Additionally, embryos from heat-stressed mothers displayed increased sensitivity to high-temperature exposure. Simulations suggest that critically low temperatures, rather than high temperatures, historically limit development of our study population. Thus, the plasticity needed to buffer this population has not been under selection. Plasticity will likely fail to compensate for ongoing climate change when such change results in novel stressors. © 2016 John Wiley & Sons Ltd.

A Temporarily Red Light-Insensitive Mutant of Tomato Lacks a Light-Stable, B-Like Phytochrome.

PubMed Central

Van Tuinen, A.; Kerckhoffs, LHJ.; Nagatani, A.; Kendrick, R. E.; Koornneef, M.

1995-01-01

We have selected four recessive mutants in tomato (Lycopersicon esculentum Mill.) that, under continuous red light (R), have long hypocotyls and small cotyledons compared to wild type (WT), a phenotype typical of phytochrome B (phyB) mutants of other species. These mutants, which are allelic, are only insensitive to R during the first 2 days upon transition from darkness to R, and therefore we propose the gene symbol tri (temporarily red light insensitive). White light-grown mutant plants have a more elongated growth habit than that of the WT. An immunochemically and spectrophotometrically detectable phyB-like polypeptide detectable in the WT is absent or below detection limits in the tri1 mutant. In contrast to the absence of an elongation growth response to far-red light (FR) given at the end of the daily photoperiod (EODFR) in all phyB-deficient mutants so far characterized, the tri1 mutant responds to EODFR treatment. The tri1 mutant also shows a strong response to supplementary daytime far-red light. We propose that the phyB-like phytochrome deficient in the tri mutants plays a major role during de-etiolation and that other light-stable phytochromes can regulate the EODFR and shade-avoidance responses in tomato. PMID:12228517

Effects of shade on physiological changes, oxidative stress, and total antioxidant power in Thai Brahman cattle.

PubMed

Aengwanich, Worapol; Kongbuntad, Watee; Boonsorn, Thongchai

2011-09-01

The purpose of this study was to assess the effects of artificial shade, tree shade, and no shade on physiological changes, oxidative stress, and total antioxidant power in Thai Brahman cattle. Twenty-one cattle were divided into three groups: cattle maintained under artificial shade, under tree shade, and without shade. On days 1, 7, 14, 21, and 28 of the experimental period, after the cattle were set in individual stalls for 2 h, physiological changes, thiobarbituric acid reactive substances (TBARS), and total antioxidant power were investigated. The results revealed that the respiratory rate, heart rate, sweat rate and the neutrophil/lymphocyte ratio of the no-shade cattle were significantly higher than those of cattle maintained under artificial shade and tree shade (P < 0.05). During the early period of heat exposure, the total antioxidant power of the no-shade cattle was lower than those of cattle maintained under artificial shade and tree shade, but the total antioxidant power of cattle maintained under artificial shade and tree shade were not different (P > 0.05). However, rectal temperature and packed cell volume of the cattle in all groups did not differ (P > 0.05). These results showed that artificial shade and tree shade can protect cattle from sunlight compared to no shade, and that the effectiveness of tree shade for sunlight protection is at an intermediate level.

Microclimatological and Physiological Controls of Stomatal Conductance and Transpiration of Co-Occurring Seedlings with Varying Shade Tolerance

NASA Astrophysics Data System (ADS)

Siegert, C. M.; Levia, D. F.

2010-12-01

Forest ecosystems provide a significant portion of fresh water to the hydrologic cycle through transpiration, the majority of which is supplied by saplings and mature trees. However, a smaller, yet measurable, proportion is also supplied by seedlings. The contribution of seedlings is dependent upon physiological characteristics of the species, whose range of habitat is ultimately controlled by microclimate. The objectives of this study were to (1) observe meteorological conditions of two forest microlimates and (2) assess the intra- and interspecific stomatal conductance and transpiration responses of naturally occurring seedlings of varying shade tolerance. Naturally established seedlings in a deciduous forest understory and an adjacent clearing were monitored throughout the 2008 growing season in southeastern Pennsylvania (39°49'N, 75°43'W). Clear spatial and temporal trends of stomatal conductance and transpiration were observed throughout this study. The understory microclimate conditions overall had a lower degree of variability and had consistently lower mean quantum flux density, air temperature, vapor pressure deficit, volumetric water content, and soil temperature than the clearing plot. Shade tolerant understory seedlings (Fagus grandifolia Ehrh. (American beech) and Prunus serotina L. (black cherry)) had significantly lower mean monthly rates of water loss (p = 0.05) than shade intolerant clearing seedlings (P. serotina and Liriodendron tulipifera L. (yellow poplar)). Additionally, water loss by shade grown P. serotina was significantly lower (p = 0.05) than by sun grown P. serotina. Significant intraspecific responses (p = 0.05) were also observed on a monthly basis, with the exception of L. tulipifera. These findings indicate that physiological differences, specifically shade tolerance, play an important role in determining rates of stomatal conductance and transpiration in tree seedlings. To a lesser degree, microclimate variability was also shown to influence rates stomatal conductance and transpiration (3.6% and 7.8% in the understory and 8.2% and 23.2% in the clearing, respectively). Field validations are critical to developing better models and forest management strategies and therefore the results of this study may serve to validate those obtained in previous studies conducted largely under controlled conditions.

Responses of leaf structure and photosynthetic properties to intra-canopy light gradients: a common garden test with four broadleaf deciduous angiosperm and seven evergreen conifer tree species.

PubMed

Wyka, Tomasz P; Oleksyn, J; Zytkowiak, R; Karolewski, P; Jagodziński, A M; Reich, P B

2012-09-01

Spectra of leaf traits in northern temperate forest canopies reflect major differences in leaf longevity between evergreen conifers and deciduous broadleaf angiosperms, as well as plastic modifications caused by within-crown shading. We investigated (1) whether long-lived conifer leaves exhibit similar intra-canopy plasticity as short-lived broadleaves, and (2) whether global interspecific relationships between photosynthesis, nitrogen, and leaf structure identified for sun leaves adequately describe leaves differentiated in response to light gradients. We studied structural and photosynthetic properties of intra-tree sun and shade foliage in adult trees of seven conifer and four broadleaf angiosperm species in a common garden in Poland. Shade leaves exhibited lower leaf mass-per-area (LMA) than sun leaves; however, the relative difference was smaller in conifers than in broadleaves. In broadleaves, LMA was correlated with lamina thickness and tissue density, while in conifers, it was correlated with thickness but not density. In broadleaves, but not in conifers, reduction of lamina thickness was correlated with a thinner palisade layer. The more conservative adjustment of conifer leaves could result from a combination of phylogenetic constraints, contrasting leaf anatomies and shoot geometries, but also from functional requirements of long-lived foliage. Mass-based nitrogen concentration (N(mass)) was similar between sun and shade leaves, and was lower in conifers than in deciduous broadleaved species. Given this, the smaller LMA in shade corresponded with a lower area-based N concentration (N(area)). In evergreen conifers, LMA and N(area) were less powerful predictors of area-based photosynthetic rate (A (max(area))) in comparison with deciduous broadleaved angiosperms. Multiple regression for sun and shade leaves showed that, in each group, A (max(mass)) was related to N(mass) but not to LMA, whereas LMA became a significant codeterminant of A (max(mass)) in analysis combining both groups. Thus, a fundamental mass-based relationship between photosynthesis, nitrogen, and leaf structure reported previously also exists in a dataset combining within-crown and across-functional type variation.

Development of transgenic crops based on photo-biotechnology.

PubMed

Ganesan, Markkandan; Lee, Hyo-Yeon; Kim, Jeong-Il; Song, Pill-Soon

2017-11-01

The phenotypes associated with plant photomorphogenesis such as the suppressed shade avoidance response and de-etiolation offer the potential for significant enhancement of crop yields. Of many light signal transducers and transcription factors involved in the photomorphogenic responses of plants, this review focuses on the transgenic overexpression of the photoreceptor genes at the uppermost stream of the signalling events, particularly phytochromes, crytochromes and phototropins as the transgenes for the genetic engineering of crops with improved harvest yields. In promoting the harvest yields of crops, the photoreceptors mediate the light regulation of photosynthetically important genes, and the improved yields often come with the tolerance to abiotic stresses such as drought, salinity and heavy metal ions. As a genetic engineering approach, the term photo-biotechnology has been coined to convey the idea that the greater the photosynthetic efficiency that crop plants can be engineered to possess, the stronger the resistance to biotic and abiotic stresses. Development of GM crops based on photoreceptor transgenes (mainly phytochromes, crytochromes and phototropins) is reviewed with the proposal of photo-biotechnology that the photoreceptors mediate the light regulation of photosynthetically important genes, and the improved yields often come with the added benefits of crops' tolerance to environmental stresses. © 2016 John Wiley & Sons Ltd.

Does triacylglycerol (TAG) serve a photoprotective function in plant leaves? An examination of leaf lipids under shading and drought.

PubMed

Marchin, Renée M; Turnbull, Tarryn L; Deheinzelin, Audrey I; Adams, Mark A

2017-11-01

Plant survival in many ecosystems requires tolerance of large radiation loads, unreliable water supply and suboptimal soil fertility. We hypothesized that increased production of neutral lipids (triacylglycerols, TAGs) in plant leaves is a mechanism for dissipating excess radiation energy. In a greenhouse experiment, we combined drought and shade treatments and examined responses among four species differing in life form, habitat, and drought- and shade-tolerance. We also present a lipid extraction protocol suitable for sclerophyllous leaves of native Australian trees (e.g. Acacia, Eucalyptus). Fluorescence measurements indicated that plants exposed to full sunlight experienced mild photoinhibition during our experiment. Accumulation of TAGs did not follow photosynthetic capacity, but instead, TAG concentration increased with non-photochemical quenching. This suggests that plants under oxidative stress may increase biosynthesis of TAGs. Moderate drought stress resulted in a 60% reduction in TAG concentration in wheat (Triticum aestivum). Shading had no effect on TAGs, but increased concentrations of polar lipids in leaves; for example, acclimation to shade in Austrodanthonia spp., a native Australian grass, resulted in a 60% increase in associated polar lipids and higher foliar chlorophyll concentrations. Shading also reduced the digalactosyldiacylglycerol:monogalactosyldiacylglycerol (DGDG:MGDG) ratio in leaves, with a corresponding increase in the degree of unsaturation and thus fluidity of thylakoid membranes of chloroplasts. Our results suggest that prevention of photodamage may be coordinated with accumulation of TAGs, although further research is required to determine if TAGs serve a photoprotective function in plant leaves. © 2017 Scandinavian Plant Physiology Society.

The impact of translucent fabric shades and control strategies on energy savings and visual quality

NASA Astrophysics Data System (ADS)

Wankanapon, Pimonmart

Translucent fabric shades provide opportunities for building occupants to control sunlight penetration for heat reduction, thermal comfort, and visual quality. Regulating shades affects building energy and can potentially reduce the size of mechanical cooling systems. Shades are not normally included in energy model studies during the design process, even though shades potential impact energy use. This is because the occupants normally leave shades closed a large fraction of the time, but models are generally performed with no shades. Automatic shade control is now available, so it is necessary to understand the impact of shades on visual quality and their energy saving potential in order to optimize their overall performance. There are very limited studies that have address shades and their integrated performance on energy consumption and visual quality. Most of these do not reflected modern shade types and their application. The goals of this study are: First, to determine the impact of shades on total, heating, cooling and lighting energy savings with different design and operation parameters. Second, to study and develop different automatic shade control strategies to promote and optimize energy savings and visual quality. A simulation-based approach using EnergyPlus in a parametric study provide better understanding energy savings under different shade conditions. The parametric runs addressed various building parameters such as geometry, orientation, site climate, glazing/shade properties, and shade control strategies with integrated lighting control. The impact of shades was determined for total building and space heating, cooling and lighting energy savings. The effect of shades on visual quality was studied using EnergyPlus, AGI32 and DAYSIM for several indices such as daylight glare index (DGI), work plane illuminance, luminance ratios and view. Different shade control strategies and integrated lighting control were considered with two translucent fabric shade colors. The results clearly show the benefit of automatic shade control strategies with integrated lighting control over a condition when shades are closed all day. The main contributor to the total energy savings is from lighting energy savings, followed by cooling energy savings. Shades provide greater benefit in a hot climate and in a moderate climate than in a cold climate. Different control strategies provide savings in the range of 7-35% for annual total space energy with higher savings with light colored shades. Control strategies of shades should be selected and optimized based on climate, orientation, window area, and window/shade properties. High performance glazings, when equipped with shades, show lower energy savings when compared to standard glazings. High transmittance/reflectance shades, such as white shades, perform better than dark shades in most of the cases due to higher lighting energy savings obtained with the automatic electric lighting control and the resulting cooling energy savings from rejection of some solar energy and a reduction in the heat from lights. A South orientation showed the least benefit of automatic control of shades when compare to other orientations due to the large fraction of time shades are required to provide visual comfort. Under automatic shade control, energy savings are higher the more often the shades can be raised. The different automatic control strategies present tradeoffs between energy savings and comfort. With regard to visual quality, daylight quality assessments on view, glare, luminance ratios, and UDI can be used to assess shade control strategies. Automatic shade control can increase the number of view hours while controlling sunlight penetration. With automatic shade control, more daylight hours can be provided within the beneficial range of 100-2000 lux compared to shades that are closed all day. For a person facing the window, discomfort glare is likely to increase the more often the shades are raised. Keeping the shades down ensures an acceptable glare condition, but limits energy savings. Luminance ratios are another metric that can be used to assess shade performance. With white shades, the luminance ratios between the task and proximate surfaces are improved. Dark shades help improve the luminance ratios between the task and distant surfaces. When the shades are left open, even with no direct sunlight in the space, task to window luminance ratios will often exceed 1:10.

Loss of photosynthetic efficiency in the shade. An Achilles heel for the dense modern stands of our most productive C4 crops?

PubMed Central

Pignon, Charles P.; Jaiswal, Deepak; McGrath, Justin M.

2017-01-01

Abstract The wild progenitors of major C4 crops grew as individuals subjected to little shading. Today they are grown in dense stands where most leaves are shaded. Do they maintain photosynthetic efficiency in these low light conditions produced by modern cultivation? The apparent maximum quantum yield of CO2 assimilation (ΦCO2max,app), a key determinant of light-limited photosynthesis, has not been systematically studied in field stands of C4 crops. ΦCO2max,app was derived from the initial slope of the response of leaf CO2 uptake (A) to photon flux (Q). Leaf fractional light absorptance (α) was measured to determine the absolute maximum quantum yield of CO2 assimilation on an absorbed light basis (ΦCO2max,abs). Light response curves were determined on sun and shade leaves of 49 field plants of Miscanthus × giganteus and Zea mays following canopy closure. ΦCO2max,app and ΦCO2max,abs declined significantly by 15–27% (P<0.05) with canopy depth. Experimentally, leaf age was shown unlikely to cause this loss. Modeling canopy CO2 assimilation over diurnal courses suggested that the observed decline in ΦCO2max,app with canopy depth costs 10% of potential carbon gain. Overcoming this limitation could substantially increase the productivity of major C4 crops. PMID:28110277

A comparison of 2 evaporative cooling systems on a commercial dairy farm in Saudi Arabia.

PubMed

Ortiz, X A; Smith, J F; Villar, F; Hall, L; Allen, J; Oddy, A; Al-Haddad, A; Lyle, P; Collier, R J

2015-12-01

Efficacy of 2 cooling systems (Korral Kool, KK, Korral Kool Inc., Mesa, AZ; FlipFan dairy system, FF, Schaefer Ventilation Equipment LLC, Sauk Rapids, MN) was estimated utilizing 400 multiparous Holstein dairy cows randomly assigned to 1 of 4 cooled California-style shade pens (2 shade pens per cooling system). Each shaded pen contained 100 cows (days in milk=58±39, milk production=56±18 kg/d, and lactation=3±1). Production data (milk yield and reproductive performance) were collected during 3mo (June-August, 2013) and physiological responses (core body temperature, respiration rates, surface temperatures, and resting time) were measured in June and July to estimate responses of cows to the 2 different cooling systems. Water and electricity consumption were recorded for each system. Cows in the KK system displayed slightly lower respiration rates in the month of June and lower surface temperatures in June and July. However, no differences were observed in the core body temperature of cows, resting time, feed intake, milk yield, services/cow, and conception rate between systems. The FF system used less water and electricity during this study. In conclusion, both cooling systems (KK and FF) were effective in mitigating the negative effects of heat stress on cows housed in arid environments, whereas the FF system consumed less water and electricity and did not require use of curtains on the shade structure. Copyright © 2015 American Dairy Science Association. Published by Elsevier Inc. All rights reserved.

Effects of low light on photosynthetic properties, antioxidant enzyme activity, and anthocyanin accumulation in purple pak-choi (Brassica campestris ssp. Chinensis Makino).

PubMed

Zhu, Hongfang; Li, Xiaofeng; Zhai, Wen; Liu, Yang; Gao, Qianqian; Liu, Jinping; Ren, Li; Chen, Huoying; Zhu, Yuying

2017-01-01

Anthocyanins are secondary metabolites that contribute to red, blue, and purple colors in plants and are affected by light, but the effects of low light on the physiological responses of purple pak-choi plant leaves are still unclear. In this study, purple pak-choi seedlings were exposed to low light by shading with white gauze and black shading in a phytotron. The responses in terms of photosynthetic properties, carbohydrate metabolism, antioxidant enzyme activity, anthocyanin biosynthetic enzyme activity, and the relative chlorophyll and anthocyanin content of leaves were measured. The results showed that chlorophyll b, intracellular CO2 content, stomatal conductance and antioxidant activities of guaiacol peroxidase, catalase and superoxide dismutase transiently increased in the shade treatments at 5 d. The malondialdehyde content also increased under low light stress, which damages plant cells. With the extension of shading time (at 15 d), the relative chlorophyll a, anthocyanin and soluble protein contents, net photosynthetic rate, transpiration rate, stomata conductance, antioxidant enzyme activities, and activities of four anthocyanin biosynthetic enzymes decreased significantly. Thus, at the early stage of low light treatment, the chlorophyll b content increased to improve photosynthesis. When the low light treatment was extended, antioxidant enzyme activity and the activity of anthocyanin biosynthesis enzymes were inhibited, causing the purple pak-choi seedlings to fade from purple to green. This study provides valuable information for further deciphering genetic mechanisms and improving agronomic traits in purple pak-choi under optimal light requirements.

How do Mediterranean shrub species cope with shade? Ecophysiological response to different light intensities.

PubMed

Díaz-Barradas, M C; Zunzunegui, M; Alvarez-Cansino, L; Esquivias, M P; Valera, J; Rodríguez, H

2018-03-01

Under natural conditions, light exposure for Mediterranean shrubs can be highly variable, especially during cloudy days or under a canopy, and can interfere with other environmental factors such as temperature and water availability. With the aim of decoupling the effect of radiation and temperature from water availability, we conducted an experiment where two perennial and three summer semi-deciduous shrub species were subjected to different levels of irradiation. In order to follow plant responses to light exposure, we measured gas exchange, photosystem II photochemical efficiency, photosynthetic pigments and leaf mass area in spring and summer. Results showed that all study species presented a plastic response to different light conditions, and that light-related traits varied in a coordinated manner. Summer semi-deciduous species exhibited a more opportunistic response, with higher photosynthesis rates in full sun, but under shade conditions, the two strategies presented similar assimilation rates. Stomatal conductance did not show such a drastic response as photosynthetsis, being related to changes in WUE. Daily cycles of F v /F m revealed a slight photoinhibitory response during summer, mainly in perennial species. In all cases photosynthetic pigments adjusted to the radiation level; leaves had lower chlorophyll content, higher pool of xanthophylls and higher proportion of the de-epoxydaded state of xanthophylls under sun conditions. Lutein content increased in relation to the xanthophyll pool under shade conditions. Our results evidenced that radiation is an important driving factor controlling morphological and physiological status of Mediterranean shrub species, independently of water availability. Summer semi-deciduous species exhibit a set of traits with higher response variability, maximising their photosynthetic assimilation under different sun conditions. © 2017 German Society for Plant Sciences and The Royal Botanical Society of the Netherlands.

In vitro model to evaluate reliability and accuracy of a dental shade-matching instrument.

PubMed

Kim-Pusateri, Seungyee; Brewer, Jane D; Dunford, Robert G; Wee, Alvin G

2007-11-01

There are several electronic shade-matching instruments available for clinical use; unfortunately, there are limited acceptable in vitro models to evaluate their reliability and accuracy. The purpose of this in vitro study was to evaluate the reliability and accuracy of a dental clinical shade-matching instrument. Using the shade-matching instrument (ShadeScan), color measurements were made of 3 commercial shade guides (VITA Classical, VITA 3D-Master, and Chromascop). Shade tabs were selected and placed in the middle of a gingival matrix (Shofu Gummy), with tabs of the same nominal shade from additional shade guides placed on both sides. Measurements were made of the central region of the shade tab inside a black box. For the reliability assessment, each shade tab from each of the 3 shade guide types was measured 10 times. For the accuracy assessment, each shade tab from 10 guides of each of the 3 types evaluated was measured once. Reliability, accuracy, and 95% confidence intervals were calculated for each shade tab. Differences were determined by 1-way ANOVA followed by the Bonferroni multiple comparison procedure. Reliability of ShadeScan was as follows: VITA Classical = 95.0%, VITA 3D-Master = 91.2%, and Chromascop = 76.5%. Accuracy of ShadeScan was as follows: VITA Classical = 65.0%, VITA 3D-Master = 54.2%, Chromascop = 84.5%. This in vitro study showed a varying degree of reliability and accuracy for ShadeScan, depending on the type of shade guide system used.

Tooth shade measurements under standard and nonstandard illumination and their agreement with skin color.

PubMed

Al-Dwairi, Ziad; Shaweesh, Ashraf; Kamkarfar, Sohrab; Kamkarfar, Shahrzad; Borzabadi-Farahani, Ali; Lynch, Edward

2014-01-01

The purpose of this study was to examine the relationship between skin color (shade) and tooth shade under standard and nonstandard illumination sources. Four hundred Jordanian participants (200 males, 200 females, 20 to 50 years of age) were studied. Skin colors were assessed and categorized using the L'Oreal and Revlon foundation shade guides (light, medium, dark). The Vita Pan Classical Shade Guide (VPCSG; Vident) and digital Vita EasyShade Intraoral Dental Spectrophotometer (VESIDS; Vident) were used to select shades in the middle thirds of maxillary central incisors; tooth shades were classified into four categories (highest, high, medium, low). Significant gender differences were observed for skin colors (P = .000) and tooth shade guide systems (P = .001 and .050 for VPCSG and VESIDS, respectively). The observed agreement was 100% and 93% for skin and tooth shade guides, respectively. The corresponding kappa statistic values were 1.00 and 0.79, respectively (substantial agreement, P < .001). The observed agreement between skin color and tooth shades (VPCSG and VESIDS) was approximately 50%. The digital tooth shade guide system can be a satisfactory substitute for classical tooth shade guides and clinical shade matching. There was only moderate agreement between skin color and tooth shade.

Prevalence of Tooth Shade and its Correlation with Skin Colour - A Cross-sectional Study.

PubMed

Vadavadagi, Suneel V; Kumari, K V Halini; Choudhury, Gopal Krishna; Vilekar, Abhishek Madhukar; Das, Sitansu Sekhar; Jena, Debkant; Kataraki, Bharat; B L, Bhavana

2016-02-01

Aesthetics has become an important issue in modern society. Tooth shade is one of the factors in determining aesthetics. Studies have revealed that tooth shade is influenced by age, gender, eye colour, skin colour and other factors. The present study was aimed to assess the prevalence of tooth shade and its correlation with skin colour. A total of 300 subjects aged 18-20 years were evaluated for tooth shade using Vitapan - 3D shade guide. Anterior teeth were checked under natural light and facial skin colour by Lakme liquid foundation make up as a shade guide. Data was analysed using chi square test and spearman's correlation. Out of 300 students, 114 (38.00%) had A2 tooth shade; the least prevalent tooth shade among Chitradurga population was C1 (4.00%). There was a positive correlation between tooth shade and skin colour which was found to be statistically significant (p <0.05). The most prevalent tooth shade among Chitradurga population was A2 and least was C1. There was a significant correlation between tooth shade and skin colour with lighter skin tone subjects having lighter tooth shade hence skin colour can be used as a guide for shade selection.

Effect of individual shades on reliability and validity of observers in colour matching.

PubMed

Lagouvardos, P E; Diamanti, H; Polyzois, G

2004-06-01

The effect of individual shades in shade guides, on the reliability and validity of measurements in a colour matching process is very important. Observer's agreement on shades and sensitivity/specificity of shades, can give us an estimate of shade's effect on observer's reliability and validity. In the present study, a group of 16 students, matched 15 shades of a Kulzer's guide and 10 human incisors to Kulzer's and/or Vita's shade tabs, in 4 different tests. The results showed shades I, B10, C40, A35 and A10 were those with the highest reliability and validity values. In conclusion, a) the matching process with shades of different materials was not accurate enough, b) some shades produce a more reliable and valid match than others and c) teeth are matched with relative difficulty.

Shade guide optimization--a novel shade arrangement principle for both ceramic and composite shade guides when identifying composite test objects.

PubMed

Østervemb, Niels; Jørgensen, Jette Nedergaard; Hørsted-Bindslev, Preben

2011-02-01

The most widely used shade guide for composite materials is made of ceramic and arranged according to a non-proven method. There is a need for a composite shade guide using a scientifically based arrangement principle. To compare the shade tab arrangement of the Vitapan Classical shade guide and an individually made composite shade guide using both the originally proposed arrangement principle and arranged according to ΔE2000 values with hue group division. An individual composite shade guide made from Filtek Supreme XT body colors was compared to the Vitapan Classical shade guide. Twenty-five students matched color samples made from Filtek Supreme XT body colors using the two shade guides arranged after the two proposed principles--four shade guides in total. Age, sequence, gender, time, and number of correct matches were recorded. The proposed visually optimal composite shade guide was both fastest and had the highest number of correct matches. Gender was significantly associated with time used for color sampling but not regarding the number of correct shade matches. A composite shade guide is superior compared to the ceramic Vitapan Classical guide when using composite test objects. A rearrangement of the shade guide according to hue, subdivided according to ΔE2000, significantly reduces the time needed to take a color sample and increases the number of correct shade matches. Total color difference in relation to the lightest tab with hue group division is recommended as a possible and universally applicable mode of tab arrangement in dental color standards. Moreover, a shade guide made of the composite materials itself is to be preferred as both a faster and more accurate method of determining color. © 2011, COPYRIGHT THE AUTHORS. JOURNAL COMPILATION © 2011, WILEY PERIODICALS, INC.

EXAMINING THE PREFERENCE FOR SHADE STRUCTURES IN FARMED GREEN SEA TURTLES (CHELONIA MYDAS) AND SHADE'S EFFECT ON GROWTH AND TEMPERATURES.

PubMed

Keller, Martha; Mustin, Walter

2017-03-01

The Cayman Turtle Farm raises thousands of green sea turtles ( Chelonia mydas ) annually under aquaculture conditions. Historically, the turtles have been raised in tanks without routine access to a shade structure. The purpose of this study was to determine the effects of adding a shade structure on curved carapace length (CCL) and weight gain of green sea turtles. In addition, water and cloacal temperatures were compared across treatment groups and shade cover preferences observed. Ninety turtles were split equally into three treatment groups for this 8-wk study. In the first group turtles were kept in tanks in full sun, the second group in half-shaded tanks, and the third group in tanks completely covered with shade cloth. Time-lapse cameras mounted above half-shaded tanks were used to determine turtle shade structure preferences throughout the day. There were no differences in CCL among treatment groups. Significant increases in weights were noted in turtles kept in full sun and half-shaded tanks versus the fully shaded tanks. Significantly higher water and cloacal turtle temperatures were noted in the full-sun tank compared with the half-shaded or completely shaded tanks. A significantly lower number of turtles was observed in the sun in the half-shaded tanks, indicating a possible preference by turtles for a shade structure. Results suggest that providing shade structures for sea turtles results in a significant decrease in both overall water temperature as well as a reduction in maximum high daily temperatures. Results also suggest that turtles exhibit a preference for shade structures when it is provided as an option. From these results, we recommend that a shade structure be provided when housing green sea turtles in outdoor enclosures.

Relationships between ambient conditions, thermal status, and feed intake of cattle during summer heat stress with access to shade.

PubMed

Curtis, A K; Scharf, B; Eichen, P A; Spiers, D E

2017-01-01

Heat stress in feedlot cattle is known to reduce their performance. The challenge comes in determining reliable predictors of current and near-future changes in thermal status and performance. A 42-d study, using crossbred (Bos taurus) steers was conducted during summer months (July through August) to identify best environmental determinants of rumen temperature (T rumen ) and feed intake (FI) in feedlot cattle with access to shade. A further goal was to define the relationship between T rumen and FI. Shade coverage was approximately 50%, and all animals were provided standard feedlot diets and water ad libitum. Intraruminal telemetric boluses recorded T rumen several times each hour. Ear tags, telemetrically connected to a feed monitoring system, provided FI data using RFID technology. Data loggers recorded ambient conditions in sun and shade, along with black globe temperature. Regression analyses identified daylight black globe and air temperatures in shade, with one hour delays, as the best predictors of T rumen . Prediction of FI was much less reliable. Unexpectedly, T rumen was not superior to ambient variables in predicting FI. Maximum daily temperature humidity index, calculated using BG in sun with a 5-d lag, was the best significant predictor of FI. These results indicate for feedlot cattle that although air temperature alone in the shade may be the best predictor of T rumen in the heat, black globe temperature in the sun may be a better determinant of feed intake over time. Additional studies are needed to verify the delayed FI response which seems unusually long. Copyright © 2016 Elsevier Ltd. All rights reserved.

Light history modulates antioxidant and photosynthetic responses of biofilms to both natural (light) and chemical (herbicides) stressors.

PubMed

Bonnineau, Chloé; Sague, Irene Gallardo; Urrea, Gemma; Guasch, Helena

2012-05-01

In multiple stress situations, the co-occurrence of environmental and chemical factors can influence organisms' ability to cope with toxicity. In this context, the influence of light adaptation on the response of freshwater biofilms to sudden light changes or to herbicides exposure was investigated by determining various parameters: diatom community composition, photosynthetic parameters, chlorophyll a content, antioxidant enzyme activities. Biofilms were grown in microcosms under sub-optimal, saturating, and high light intensities and showed already described characteristics of shade/light adaptation (community structure, photosynthetic adaptation, etc.). Light history modulated antioxidant and photosynthetic responses of biofilms to the stress caused by short-term exposure to sudden light changes or to herbicides. First biofilms adapted to sub-optimal light intensity (shade-adapted) were found to be more sensitive to an increase in light intensity than high-light adapted ones to a reduction in light intensity. Second, while light history influenced biofilms' response to glyphosate, it had little influence on biofilms' response to copper and none on its response to oxyfluorfen. Indeed glyphosate exposure led to a stronger decrease in photosynthetic efficiency of shade-adapted biofilms (EC(50) = 11.7 mg L(-1)) than of high-light adapted communities (EC(50) = 35.6 mg L(-1)). Copper exposure led to an activation of ascorbate peroxidase (APX) in biofilms adapted to sub-optimal and saturating light intensity while the protein content decreased in all biofilms exposed to copper. Oxyfluorfen toxicity was independent of light history provoking an increase in APX activity. In conclusion this study showed that both previous exposure to contaminants and physical habitat characteristics might influence community tolerance to disturbances strongly.

Content of antioxidative caffeoylquinic acid derivatives in field-grown Ligularia fischeri (Ledeb.) Turcz and responses to sunlight.

PubMed

Kim, Sang Min; Jeon, Je-Seung; Kang, Suk Woo; Jung, Yu-Jin; Ly, Lin Na; Um, Byung-Hun

2012-06-06

Ligularia fischeri (Ledeb.) Turcz, a commercial leafy vegetable, contains caffeoylquinic acid derivatives (CQAs) as major phenolic constituents. The HPLC chromatograms of leaf extracts collected from different areas in Korea showed a significant variation in CQA amount, and two tri-O-caffeoylquinic acids (triCQAs) were purified and structurally identified by NMR and MS from this plant. Radical scavenging activities among CQAs were found to be increased in proportion to the number of caffeoyl groups. Since this plant prefers damp and shady growth conditions, the effects of sunlight were investigated by growing plantlets in sunlight and shade for four weeks. Greater leaf thickness and higher phenolic contents were found for leaves grown in sunlight than in shade. Four major CQAs-5-mono-O-caffeoylquinic acid (5-monoCQA), and 3,4-, 3,5-, and 4,5-di-O-caffeoylquinic acid (diCQA)-were induced by solar irradiation, whereas the content of these compounds decreased steadily in shade leaves. The leaves of L. fischeri clearly showed adaptation responses to sunlight, and these characteristics can be exploited for cultivation of this plant for potential use as a nutraceutical and functional food.

Evaluation of Dental Shade Guide Variability Using Cross-Polarized Photography.

PubMed

Gurrea, Jon; Gurrea, Marta; Bruguera, August; Sampaio, Camila S; Janal, Malvin; Bonfante, Estevam; Coelho, Paulo G; Hirata, Ronaldo

2016-01-01

This study evaluated color variability in the A hue between the VITA Classical (VITA Zahnfabrik) shade guide and four other VITA-coded ceramic shade guides using a Canon EOS 60D camera and software (Photoshop CC, Adobe). A total of 125 photographs were taken, 5 per shade tab for each of 5 shades (A1 to A4) from the following shade guides: VITA Classical (control), IPS e.max Ceram (Ivoclar Vivadent), IPS d.SIGN (Ivoclar Vivadent), Initial ZI (GC), and Creation CC (Creation Willi Geller). Photos were processed with Adobe Photoshop CC to allow standardized evaluation of hue, chroma, and value between shade tabs. None of the VITA-coded shade tabs fully matched the VITA Classical shade tab for hue, chroma, or value. The VITA-coded shade guides evaluated herein showed an overall unmatched shade in all tabs when compared with the control, suggesting that shade selection should be made using the guide produced by the manufacturer of the ceramic intended for the final restoration.

Microarray and growth analyses identify differences and similarities of early corn response to weeds, shade, and nitrogen stress

USDA-ARS?s Scientific Manuscript database

Weed interference with crop growth is often attributed to water, nutrient, or light competition; however, specific physiological responses to these stresses are not well described. This study’s objective was to compare growth, yield, and gene expression responses of corn to nitrogen (N), low light (...

Removal of nutrient limitations in forest gaps enhances growth rate and resistance to cavitation in subtropical canopy tree species differing in shade tolerance.

PubMed

Villagra, Mariana; Campanello, Paula I; Montti, Lia; Goldstein, Guillermo

2013-03-01

A 4-year fertilization experiment with nitrogen (N) and phosphorus (P) was carried out in natural gaps of a subtropical forest in northeastern Argentina. Saplings of six dominant canopy species differing in shade tolerance were grown in five control and five N + P fertilized gaps. Hydraulic architectural traits such as wood density, the leaf area to sapwood area ratio (LA : SA), vulnerability to cavitation (P50) and specific and leaf-specific hydraulic conductivity were measured, as well as the relative growth rate, specific leaf area (SLA) and percentage of leaf damage by insect herbivores. Plant growth rates and resistance to drought-induced embolisms increased when nutrient limitations were removed. On average, the P50 of control plants was -1.1 MPa, while the P50 of fertilized plants was -1.6 MPa. Wood density and LA : SA decreased with N + P additions. A trade-off between vulnerability to cavitation and efficiency of water transport was not observed. The relative growth rate was positively related to the total leaf surface area per plant and negatively related to LA : SA, while P50 was positively related to SLA across species and treatments. Plants with higher growth rates and higher total leaf area in fertilized plots were able to avoid hydraulic dysfunction by becoming less vulnerable to cavitation (more negative P50). Two high-light-requiring species exhibited relatively low growth rates due to heavy herbivore damage. Contrary to expectations, shade-tolerant plants with relatively high resistance to hydraulic dysfunction and reduced herbivory damage were able to grow faster. These results suggest that during the initial phase of sapling establishment in gaps, species that were less vulnerable to cavitation and exhibited reduced herbivory damage had faster realized growth rates than less shade-tolerant species with higher potential growth rates. Finally, functional relationships between hydraulic traits and growth rate across species and treatments were maintained regardless of soil nutrient status.

Intrarater repeatability of shade selections with two shade guides.

PubMed

Hammad, Ihab A

2003-01-01

Color research has shown that shade guides do not always represent the color of natural teeth. Moreover, visual evaluation has been found to be unreliable and inconsistent. This investigation evaluated the effects of 2 shade guides on the intrarater repeatability (reliability) of prosthodontists and general practitioners with regard to shade selection. Ten prosthodontists and ten general practitioners (all men, 35-45 years old) with an average practice experience of 14 years participated in this study. Examiners were tested to eliminate color blindness. Each clinician used Vita Lumin Vacuum and Vitapan 3D-Master shade guides to determine the shades of the maxillary right canines of 20 patients following a standard protocol. The identification codes of the shade tabs were masked to prevent shade memory. All teeth were polished before each shade selection, and the selection process was standardized for controlled lighting and procedures. Shade selections were randomly repeated 1 month later by the same practitioners on the same group of patients in accordance with the same shade-selection protocol. Analysis of variance and t tests for individual comparisons among means were performed (P<.05). Significant interactions were found between the effects of shade guide system and specialty training on intrarater repeatability (P<.0001, analysis of variance). The intrarater repeatability of prosthodontists was significantly higher than that of general practitioners when the Vita Lumin Vacuum shade guide was used (P<.0001, t test). Use of the Vitapan 3D-Master shade guide significantly improved the intrarater repeatability of general practitioners compared with the Vita Lumin Vacuum shade guide (P<.0005). This improvement was not significant, however, among prosthodontists (P=.2861). Within the limitations of this study, the prosthodontists demonstrated superior intrarater repeatability in shade selection, especially when the Vita Lumin Vacuum shade guide was used. Use of the Vitapan 3D-Master shade guide notably improved intrarater repeatability among the general practitioners.

Effects of water level, shade and time on germination and growth of freshwater marsh plants along a simulated successional gradient

USGS Publications Warehouse

Kellogg, C.H.; Bridgham, S.D.; Leicht, S.A.

2003-01-01

1. We examined the effect of soil organic content (1.4, 3.6, 7.2% soil C), water level (+2, -1, -4 cm from soil surface) and duration (13 or 33 days) on 10 species that varied in abundance during succession in freshwater marshes. We also determined the effect of shade (0, 40, 80% shade) and soil organic content (1.4 and 7.2% soil C) on germination of six species over 62 days with water 0.5 cm below the soil surface. 2. Water level consistently affected species germination on both dates. Above-ground biomass was generally higher with increasing organic content of soil, but shade had little effect on germination or height. 3 The hydrologic zone in which species were found in the field was a good indicator of the response of germination to hydrology. Both early successional species and species wide-spread across the successional gradient show similar germination on all organic contents, while later successional species appear to germinate best at higher organic contents. 4. Successional changes in soils are capable of affecting plant community development, independent of disturbance.

Response of an algal assemblage to nutrient enrichment and shading in a Hawaiian stream

USGS Publications Warehouse

Stephens, S.H.; Brasher, A.M.D.; Smith, C.M.

2012-01-01

To investigate the effects of nitrate enrichment, phosphate enrichment, and light availability on benthic algae, nutrient-diffusing clay flowerpots were colonized with algae at two sites in a Hawaiian stream during spring and autumn 2002 using a randomized factorial design. The algal assemblage that developed under the experimental conditions was investigated by determining biomass (ash-free dry mass and chlorophyll a concentrations) and composition of the diatom assemblage. In situ pulse amplitude-modulated fluorometry was also used to model photosynthetic rate of the algal assemblage. Algal biomass and maximum photosynthetic rate were significantly higher at the unshaded site than at the shaded site. These parameters were higher at the unshaded site with either nitrate, or to a lesser degree, nitrate plus phosphate enrichment. Analysis of similarity of diatom assemblages showed significant differences between shaded and unshaded sites, as well as between spring and autumn experiments, but not between nutrient treatments. However, several individual species of diatoms responded significantly to nitrate enrichment. These results demonstrate that light availability (shaded vs. unshaded) is the primary limiting factor to algal growth in this stream, with nitrogen as a secondary limiting factor. ?? 2011 Springer Science+Business Media B.V.

Sun protection policies and practices in New Zealand primary schools.

PubMed

Reeder, Anthony I; Jopson, Janet A; Gray, Andrew

2012-02-10

For schools with primary age students, to report the percentages meeting specific requirements of the New Zealand SunSmart Schools Accreditation Programme (SSAP). Schools were randomly selected, within geographic regions, from the Ministry of Education schools database. A questionnaire, mailed to school principals, assessed schools regarding 12 criteria for accreditation: policy, information, hats, 'play in the shade', sunscreen, clothing, role modelling, curriculum, planning, rescheduling, shade provision and review. Post-stratification weights (for achieving each criterion) were used to compensate for oversampling within some regions and differential response rates between regions, using the number of schools per region. 388 schools (representative in socioeconomic decile, size and type) participated. Less than 4% fully met accreditation criteria. Clothing (42%), curriculum delivery and shade (each 54%) requirements were met by the fewest schools. Staff role modelling (92%) was the most commonly met. Schools with uniforms tended to have more protective clothing expectations. Ongoing promotion is needed to consolidate gains and encourage comprehensive sun protection through policies, practices, environment and curriculum. Staff role modelling requirements may be strengthened by implementing existing occupational guidelines for mitigating UVR hazards. There is a need to further assist schools, particularly regarding sun protective clothing, curriculum delivery and environmental shade.

Reproductive success in varying light environments: direct and indirect effects of light on plants and pollinators.

PubMed

Kilkenny, Francis F; Galloway, Laura F

2008-03-01

Plant populations often exist in spatially heterogeneous environments. Light level can directly affect plant reproductive success through resource availability or by altering pollinator behavior. It can also indirectly influence reproductive success by determining floral display size which may in turn influence pollinator attraction. We evaluated direct and indirect effects of light availability and measured phenotypic selection on phenological traits that may enhance pollen receipt in the insect-pollinated herb Campanulastrum americanum. In a natural population, plants in the sun had larger displays and received 7 times more visits than plants in the shade. Using experimental arrays to separate the direct effects of irradiance on insects from their response to display size, we found more visits to plants in the sun than in the shade, but no association between number of visits each flower received and display size. Plants in the sun were not pollen limited but pollen-augmented shade flowers produced 50% more seeds than open-pollinated flowers. Phenological traits, which may influence pollen receipt, were not under direct selection in the sun. However, earlier initiation and a longer duration of flowering were favored in the shade, which may enhance visitation in this pollen-limited habitat.

Evaluation of a standard shade guide for color change after disinfection.

PubMed

Pohjola, Randall M; Hackman, Steven T; Browning, William D

2007-09-01

To determine if surface disinfectants cause a change in the shade perception of a standard Classic Vitapan shade guide. Consistency in shade selection for dental restorations involves many factors, and one of the most important is the shade tabs used in the selection process. Ten shade tabs each of shades B2, D2, C1, and A3.5 were selected from the Classic Vitapan shade guide (Vident). All tabs were measured with the EasyShade shade device (Vident) at baseline. Three tabs of each shade were set aside as controls. The other 7 tabs of each shade were treated with the surface disinfectant Cavicide (Metrex Research) for 480 cycles to simulate a year's usage. After each 480 cycles, all the tabs were again measured with the EasyShade. This process was repeated to simulate 2 and 3 years of use. The data were analyzed to calculate the delta E 2000 for any change. A statistically significant increase was observed in the value (L*) and chroma (C*) after 2 and 3 years of simulated treatments. These changes were not perceptible to the clinician. The authors suggest that 1 standard shade guide be set aside to compare against those in clinical use to determine when they should be replaced.

A conflict-based model of color categorical perception: evidence from a priming study.

PubMed

Hu, Zhonghua; Hanley, J Richard; Zhang, Ruiling; Liu, Qiang; Roberson, Debi

2014-10-01

Categorical perception (CP) of color manifests as faster or more accurate discrimination of two shades of color that straddle a category boundary (e.g., one blue and one green) than of two shades from within the same category (e.g., two different shades of green), even when the differences between the pairs of colors are equated according to some objective metric. The results of two experiments provide new evidence for a conflict-based account of this effect, in which CP is caused by competition between visual and verbal/categorical codes on within-category trials. According to this view, conflict arises because the verbal code indicates that the two colors are the same, whereas the visual code indicates that they are different. In Experiment 1, two shades from the same color category were discriminated significantly faster when the previous trial also comprised a pair of within-category colors than when the previous trial comprised a pair from two different color categories. Under the former circumstances, the CP effect disappeared. According to the conflict-based model, response conflict between visual and categorical codes during discrimination of within-category pairs produced an adjustment of cognitive control that reduced the weight given to the categorical code relative to the visual code on the subsequent trial. Consequently, responses on within-category trials were facilitated, and CP effects were reduced. The effectiveness of this conflict-based account was evaluated in comparison with an alternative view that CP reflects temporary warping of perceptual space at the boundaries between color categories.

Evaluation of an optimized shade guide made from porcelain powder mixtures.

PubMed

Wang, Peng; Wei, Jiaqiang; Li, Qing; Wang, Yining

2014-12-01

Color errors associated with current shade guides and problems with color selection and duplication are still challenging for restorative dentists. The purpose of this study was to evaluate an optimized shade guide for visual shade duplication. Color distributions (L*, a*, and b*) of the maxillary left central incisors of 236 participants, whose ages ranged from 20 to 60, were measured with a spectrophotometer. Based on this color map, an optimized shade guide was designed with 14 shade tabs evenly distributed within the given color range of the natural incisors. The shade tabs were fabricated with porcelain powder mixtures and conventional laboratory procedures. A comparison of shade duplication by using the optimized and Vitapan Classical shade guides was conducted. Thirty Chinese participants were involved, and the colors of the left maxillary incisors were selected by using 2 shade guides. Metal ceramic crowns were fabricated according to the results of the shade selection. The colors of the shade tabs, natural teeth, and the ceramic crowns were measured with a spectrophotometer. The color differences among the natural teeth, the shade tabs, and the corresponding metal ceramic crowns were calculated and analyzed (α=.017). Significant differences were found in both phases of shade determination and shade duplication (P<.017). The total number of color errors with the optimized shade guide was 3.5, which was significantly less than that of Vitapan, 5.1 (P<.001). The optimized shade guide system improved performance not only in the color selection phase but also in the color of the fabricated crowns. Copyright © 2014 Editorial Council for the Journal of Prosthetic Dentistry. Published by Elsevier Inc. All rights reserved.

Sun shield

NASA Technical Reports Server (NTRS)

Frank, Arthur M. (Inventor); Derespinis, Silvio F. (Inventor); Mockovciak, John, Jr. (Inventor)

1987-01-01

A shading device which is capable of compactly storing a flexible shade on a biased, window shade type spring roller is disclosed. It is controlled to deliver the shade selectively to either its operative shading or compact storage orientation.

Shade determination using camouflaged visual shade guides and an electronic spectrophotometer.

PubMed

Kvalheim, S F; Øilo, M

2014-03-01

The aim of the present study was to compare a camouflaged visual shade guide to a spectrophotometer designed for restorative dentistry. Two operators performed analyses of 66 subjects. One central upper incisor was measured four times by each operator; twice with a camouflaged visual shade guide and twice with a spectrophotometer Both methods had acceptable repeatability rates, but the electronic shade determination showed higher repeatability. In general, the electronically determined shades were darker than the visually determined shades. The use of a camouflaged visual shade guide seems to be an adequate method to reduce operator bias.

Shaded-Color Picture Generation of Computer-Defined Arbitrary Shapes

NASA Technical Reports Server (NTRS)

Cozzolongo, J. V.; Hermstad, D. L.; Mccoy, D. S.; Clark, J.

1986-01-01

SHADE computer program generates realistic color-shaded pictures from computer-defined arbitrary shapes. Objects defined for computer representation displayed as smooth, color-shaded surfaces, including varying degrees of transparency. Results also used for presentation of computational results. By performing color mapping, SHADE colors model surface to display analysis results as pressures, stresses, and temperatures. NASA has used SHADE extensively in sign and analysis of high-performance aircraft. Industry should find applications for SHADE in computer-aided design and computer-aided manufacturing. SHADE written in VAX FORTRAN and MACRO Assembler for either interactive or batch execution.

Influence of illuminants on the color distribution of shade guides.

PubMed

Park, Ji-Hoon; Lee, Yong-Keun; Lim, Bum-Soon

2006-12-01

Although a shade tab in a shade guide is matched to a natural tooth in the order of value, hue, and chroma, there are limited data on the color distribution of currently available shade guides sorted by these 3 parameters. Furthermore, spectrophotometric color measurements of shade tabs differ depending on the standard illuminant employed. The purpose of this study was to determine the color distributions of 2 shade guides in value (CIE L( *)), chroma (C( *)(ab)) and hue angle (h(o)) scale relative to the standard illuminants D(65), A, and F2. Color of shade tabs (n=36) from 2 shade guides (Vita Lumin and Chromascop) were measured, and the distributions for CIE L( *), C( *)(ab) and h(o) values were compared. Color differences of shade tabs depending on the illuminant were calculated. The distributions of the ratios of CIE L( *) and C( *)(ab) values of each shade tab compared with the lowest value tab or the lowest chroma tab were determined. The data for the value, chroma, and hue angle within each shade guide were analyzed with a 2-way ANOVA with the factors of shade designation and type of illuminant (alpha=.05). Color difference caused by change of illuminant was analyzed with a 2-way ANOVA with the factors of shade designation and pair of illuminants compared (alpha=.05). The Scheffe multiple comparison test was performed as a post hoc test. CIE L( *), C( *)(ab) and h(o) values were influenced by shade designation and type of illuminant in both shade guides. Color difference caused by change of the illuminant was influenced by the shade designation and pair of illuminants compared. The order of mean color differences of 16 Vita Lumin shade tabs by pairs of illuminants compared was as follows: DeltaE( *)(ab) (D(65)/F2) = 1.63 3.7). Color distribution of 2 shade guides by the value and chroma was not logical.

Evaluation of reliability and validity of three dental color-matching devices.

PubMed

Tsiliagkou, Aikaterini; Diamantopoulou, Sofia; Papazoglou, Efstratios; Kakaboura, Afrodite

2016-01-01

To assess the repeatability and accuracy of three dental color-matching devices under standardized and freehand measurement conditions. Two shade guides (Vita Classical A1-D4, Vita; and Vita Toothguide 3D-Master, Vita), and three color-matching devices (Easyshade, Vita; SpectroShade, MHT Optic Research; and ShadeVision, X-Rite) were used. Five shade tabs were selected from the Vita Classical A1-D4 (A2, A3.5, B1, C4, D3), and five from the Vita Toothguide 3D-Master (1M1, 2R1.5, 3M2, 4L2.5, 5M3) shade guides. Each shade tab was recorded 15 continuous, repeated times with each device under two different measurement conditions (standardized, and freehand). Both qualitative (color shade) and quantitative (L, a, and b) color characteristics were recorded. The color difference (ΔE) of each recorded value with the known values of the shade tab was calculated. The repeatability of each device was evaluated by the coefficient of variance. The accuracy of each device was determined by comparing the recorded values with the known values of the reference shade tab (one sample t test; α = 0.05). The agreement between the recorded shade and the reference shade tab was calculated. The influence of the parameters (devices and conditions) on the parameter ΔE was investigated (two-way ANOVA). Comparison of the devices was performed with Bonferroni pairwise post-hoc analysis. Under standardized conditions, repeatability of all three devices was very good, except for ShadeVision with Vita Classical A1-D4. Accuracy ranged from good to fair, depending on the device and the shade guide. Under freehand conditions, repeatability and accuracy for Easyshade and ShadeVision were negatively influenced, but not for SpectroShade, regardless of the shade guide. Based on the total of the color parameters assessed per device, SpectroShade was the most reliable of the three color-matching devices studied.

Heat stress in cows at pasture and benefit of shade in a temperate climate region

NASA Astrophysics Data System (ADS)

Veissier, Isabelle; Van laer, Eva; Palme, Rupert; Moons, Christel P. H.; Ampe, Bart; Sonck, Bart; Andanson, Stéphane; Tuyttens, Frank A. M.

2017-11-01

Under temperate climates, cattle are often at pasture in summer and are not necessarily provided with shade. We aimed at evaluating in a temperate region (Belgium) to what extent cattle may suffer from heat stress (measured through body temperature, respiration rate and panting score, cortisol or its metabolites in milk, and feces on hot days) and at assessing the potential benefits of shade. During the summer of 2012, 20 cows were kept on pasture without access to shade. During the summer of 2011, ten cows had access to shade (young trees with shade cloth hung between them), whereas ten cows had no access. Climatic conditions were quantified by the Heat Load Index (HLI). In animals without access to shade respiration rates, panting scores, rectal temperatures, and milk cortisol concentrations increased as HLI increased in both 2011 and 2012. Fecal cortisol metabolites varied with HLI in 2011 only. When cattle had access to shade, their use of shade increased as the HLI increased. This effect was more pronounced during the last part of the summer, possibly due to better acquaintance with the shade construction. In this case, shade use increased to 65% at the highest HLI (79). Shade tempered the effects on respiration, rectal temperature, and fecal cortisol metabolites. Milk cortisol was not influenced by HLI for cows using shade for > 10% of the day. Therefore, even in temperate areas, cattle may suffer from heat when they are at pasture in summer and providing shade can reduce such stress.

Role of riparian shade on the fish assemblage of a reservoir littoral

USGS Publications Warehouse

Raines, C. D.; Miranda, Leandro E.

2016-01-01

Research into the effects of shade on reservoir fish assemblages is lacking, with most investigations focused on streams. Unlike many streams, the canopy in a reservoir shades only a narrow fringe of water adjacent to the shoreline, and may not have the influential effect on the aquatic environment reported in streams. We compared fish assemblages between shaded and unshaded sites in a shallow reservoir. Overall species richness (gamma diversity) was higher in shaded sites, and fish assemblage composition differed between shaded and unshaded sites. Average light intensity was 66 % lower in shaded sites, and differences in average temperature and dissolved oxygen were small. Unlike streams where shade can have large effects on water physicochemistry, in reservoirs shade-related differences in fish assemblages seemed to be linked principally to differences in light intensity. Diversity in light intensity in shaded and unshaded sites in reservoirs can create various mosaics of light-based habitats that enable diversity of species assemblages. Managing to promote the habitat diversity provided by shade may require coping with the artificial nature of reservoir riparian zones and water level fluctuations.

Shade improves survival rate of outplanted 2-0 red fir seedlings

Treesearch

Donald T. Gordon

1970-01-01

Effect of shade on 2-0 red fir seedlings outplanted on the Tahoe National Forest, central California, was studied by comparing five treatments: no shade, low shade for 1 year and for 2 years, and high shade for 1 year and for 2 years. Seedlings benefited from shading-even under apparently excellent survival conditions in the first season. The extra cost of artificial...

Effects of a shade-matching light and background color on reliability in tooth shade selection.

PubMed

Najafi-Abrandabadi, Siamak; Vahidi, Farhad; Janal, Malvin N

2018-01-01

The purpose of this study was to evaluate the effects of a shade-matching light (Rite-Lite-2, AdDent) and different viewing backgrounds on reliability in a test of shade tab matching. Four members of the Prosthodontic faculty matched 10 shade tabs selected for a range of shades against the shade guide. All raters were tested for color blindness and were calibrated prior to the study. Matching took place under four combinations of conditions: with operatory light or the shade-matching light, and using either a pink or a blue background. Reliability was quantified with the kappa statistic, separately for agreement of value, hue, and chroma for each shade tab. In general, raters showed fair to moderate levels of agreement when judging the value of the shade tabs, but could not agree on the hue and chroma of the stimuli. The pink background led to higher levels of agreement than the blue background, and the shade-matching light improved agreement when used in conjunction with the pink but not the blue background. Moderate levels of agreement were found in matching shade tab value. Agreement was generally better when using the pink rather than the blue background, regardless of light source. The use of the shade-matching light tended to amplify the advantage of the pink background.

Shading effect on generative characters of upland red rice of Southeast Sulawesi, Indonesia

NASA Astrophysics Data System (ADS)

Muhidin; Syam’un, E.; Kaimuddin; Musa, Y.; Sadimantara, G. R.; Usman; Leomo, S.; Rakian, T. C.

2018-05-01

Upland red rice (Oryza sativa) contains anthocyanin, a phenolic compounds that can act as antioxidants and functional food for human dietary. The content of functional food on upland red rice is influenced by shading condition, but the production is also influenced by environmental condition, especially the availability of light. The study aims is to assess and analyze the effect of shade on the growth and production of upland red rice. The research was conducted using the quantitative method to obtain the optimal shading condition that can increase the rice anthocyanin content and relatively high production. The research was arranged in split plot design, with shade as main plot and the different of cultivar as sub plot with three replications. The shading treatment consist of 4 levels as follows: n1=shade level < 25 %, n2=shade level 25-50%, n3=shade level 50-75% and n4=shade level > 75%. The cultivar tested were (v1) = Labandiri, (v2) = Jangkobembe, (v3) = Ranggohitam, and (v4) = Paedara. The rice planted in between teak wood trees with different age and level of canopy. The research reveals that shades had an effect in decreasing plant production (the higher level of shade, the higher the decrease level of production), but the shades can improve the quality of red rice through the increase of anthocyanin content.

Uniform versus Asymmetric Shading Mediates Crown Recession in Conifers

PubMed Central

Schoonmaker, Amanda L.; Lieffers, Victor J.; Landhäusser, Simon M.

2014-01-01

In this study we explore the impact of asymmetrical vs. uniform crown shading on the mortality and growth of upper and lower branches within tree crowns, for two conifer species: shade intolerant lodgepole pine (Pinus contorta) and shade tolerant white spruce (Picea glauca). We also explore xylem hydraulics, foliar nutrition, and carbohydrate status as drivers for growth and expansion of the lower and upper branches in various types of shading. This study was conducted over a two-year period across 10 regenerating forest sites dominated by lodgepole pine and white spruce, in the lower foothills of Alberta, Canada. Trees were assigned to one of four shading treatments: (1), complete uniform shading of the entire tree, (2) light asymmetric shading where the lower 1/4–1/3 of the tree crown was shaded, (3) heavy asymmetric shading as in (2) except with greater light reduction and (4) control in which no artificial shading occurred and most of the entire crown was exposed to full light. Asymmetrical shading of only the lower crown had a larger negative impact on the bud expansion and growth than did uniform shading, and the effect was stronger in pine relative to spruce. In addition, lower branches in pine also had lower carbon reserves, and reduced xylem-area specific conductivity compared to spruce. For both species, but particularly the pine, the needles of lower branches tended to store less C than upper branches in the asymmetric shade, which could suggest a movement of reserves away from the lower branches. The implications of these findings correspond with the inherent shade tolerance and self-pruning behavior of these conifers and supports a carbon based mechanism for branch mortality – mediated by an asymmetry in light exposure of the crown. PMID:25136823

Uniform versus asymmetric shading mediates crown recession in conifers.

PubMed

Schoonmaker, Amanda L; Lieffers, Victor J; Landhäusser, Simon M

2014-01-01

In this study we explore the impact of asymmetrical vs. uniform crown shading on the mortality and growth of upper and lower branches within tree crowns, for two conifer species: shade intolerant lodgepole pine (Pinus contorta) and shade tolerant white spruce (Picea glauca). We also explore xylem hydraulics, foliar nutrition, and carbohydrate status as drivers for growth and expansion of the lower and upper branches in various types of shading. This study was conducted over a two-year period across 10 regenerating forest sites dominated by lodgepole pine and white spruce, in the lower foothills of Alberta, Canada. Trees were assigned to one of four shading treatments: (1), complete uniform shading of the entire tree, (2) light asymmetric shading where the lower 1/4-1/3 of the tree crown was shaded, (3) heavy asymmetric shading as in (2) except with greater light reduction and (4) control in which no artificial shading occurred and most of the entire crown was exposed to full light. Asymmetrical shading of only the lower crown had a larger negative impact on the bud expansion and growth than did uniform shading, and the effect was stronger in pine relative to spruce. In addition, lower branches in pine also had lower carbon reserves, and reduced xylem-area specific conductivity compared to spruce. For both species, but particularly the pine, the needles of lower branches tended to store less C than upper branches in the asymmetric shade, which could suggest a movement of reserves away from the lower branches. The implications of these findings correspond with the inherent shade tolerance and self-pruning behavior of these conifers and supports a carbon based mechanism for branch mortality--mediated by an asymmetry in light exposure of the crown.

"Double crap!" abuse and harmed identity in Fifty Shades of Grey.

PubMed

Bonomi, Amy E; Altenburger, Lauren E; Walton, Nicole L

2013-09-01

While intimate partner violence (IPV) affects 25% of women and impairs health, current societal conditions-including the normalization of abuse in popular culture such as novels, film, and music-create the context to support such violence. Fifty Shades of Grey, a best-selling novel, depicts a "romantic" and "erotic" relationship involving 28-year-old megamillionaire, Christian Grey, and a 22-year-old college student, Anastasia Steele. We argue that the relationship is characterized by IPV, which is harmful to Anastasia. All authors engaged in iterative readings of the text, and wrote narrative summaries to elucidate themes. Validity checks included double review of the first eight chapters of the novel to establish consistency in our analysis approach, iterative discussions in-person and electronically to arbitrate discrepancies, and review of our analysis with other abuse and sexual practice experts. To characterize IPV, we used the U.S. Centers for Disease Control and Prevention's definitions of emotional abuse (intimidation/threats; isolation; stalking; and humiliation) and sexual violence (forced sex acts/contact against a person's will, including using alcohol/drugs or intimidation/pressure). To characterize harm, we used Smith's conceptualizations of perceived threat, managing, altered identity, yearning, entrapment, and disempowerment experienced by abused women. Emotional abuse is present in nearly every interaction, including: stalking (Christian deliberately follows Anastasia and appears in unusual places, uses a phone and computer to track Anastasia's whereabouts, and delivers expensive gifts); intimidation (Christian uses intimidating verbal and nonverbal behaviors, such as routinely commanding Anastasia to eat and threatening to punish her); and isolation (Christian limits Anastasia's social contact). Sexual violence is pervasive-including using alcohol to compromise Anastasia's consent, as well as intimidation (Christian initiates sexual encounters when genuinely angry, dismisses Anastasia's requests for boundaries, and threatens her). Anastasia experiences reactions typical of abused women, including: constant perceived threat ("my stomach churns from his threats"); altered identity (describes herself as a "pale, haunted ghost"); and stressful managing (engages in behaviors to "keep the peace," such as withholding information about her social whereabouts to avoid Christian's anger). Anastasia becomes disempowered and entrapped in the relationship as her behaviors become mechanized in response to Christian's abuse. Our analysis identified patterns in Fifty Shades that reflect pervasive intimate partner violence-one of the biggest problems of our time. Further, our analysis adds to a growing body of literature noting dangerous violence standards being perpetuated in popular culture.

Acclimation of Plant Populations to Shade: Photosynthesis, Respiration, and Carbon Use Efficiency

NASA Technical Reports Server (NTRS)

Frantz, Jonathan M.; Bugbee, Bruce

2005-01-01

Cloudy days cause an abrupt reduction in daily photosynthetic photon flux (PPF), but we have a poor understanding of how plants acclimate to this change. We used a unique lo-chamber, steady-state, gas-exchange system to continuously measure daily photosynthesis and night respiration of populations of a starch accumulator [tomato (Lycopersicone scukntum Mill. cv. Micro-Tina)] and a sucrose accumulator [lettuce (Latuca sativa L ev. Grand Rapids)] over 42 days. AI1 measurements were done at elevated CO2, (1200micr-/mol) avoid any CO2 limitations and included both shoots and roots. We integrated photosynthesis and respiration measurements separately to determine daily net carbon gain and carbon use efficiency (CUE) as the ratio of daily net C gain to total day-time C fixed over the 42-day period. After 16 to 20 days of growth in constant PPF, plants in some chambers were subjected to an abrupt PPF reduction to simulate shade or a series of cloudy days. The immediate effect and the long term acclimation rate w'ere assessed from canopy quantum yield and carbon use efficiency. The effect of shade on carbon use efficiency and acclimation was much slower than predicted by widely used growth models. It took 12 days for tomato populations to recover their original CUE and lettuce CUE never completely acclimated. Tomatoes, the starch accumulator, acclimated to low light more rapidly than lettuce, the sucrose accumulator. Plant growth models should be modified to include the photosynthesis/respiration imbalance and resulting inefficiency of carbon gain associated with changing PIT conditions on cloudy days.

Transcriptomics of shading-induced and NAA-induced abscission in apple (Malus domestica) reveals a shared pathway involving reduced photosynthesis, alterations in carbohydrate transport and signaling and hormone crosstalk

PubMed Central

2011-01-01

Background Naphthaleneacetic acid (NAA), a synthetic auxin analogue, is widely used as an effective thinner in apple orchards. When applied shortly after fruit set, some fruit abscise leading to improved fruit size and quality. However, the thinning results of NAA are inconsistent and difficult to predict, sometimes leading to excess fruit drop or insufficient thinning which are costly to growers. This unpredictability reflects our incomplete understanding of the mode of action of NAA in promoting fruit abscission. Results Here we compared NAA-induced fruit drop with that caused by shading via gene expression profiling performed on the fruit abscission zone (FAZ), sampled 1, 3, and 5 d after treatment. More than 700 genes with significant changes in transcript abundance were identified from NAA-treated FAZ. Combining results from both treatments, we found that genes associated with photosynthesis, cell cycle and membrane/cellular trafficking were downregulated. On the other hand, there was up-regulation of genes related to ABA, ethylene biosynthesis and signaling, cell wall degradation and programmed cell death. While the differentially expressed gene sets for NAA and shading treatments shared only 25% identity, NAA and shading showed substantial similarity with respect to the classes of genes identified. Specifically, photosynthesis, carbon utilization, ABA and ethylene pathways were affected in both NAA- and shading-induced young fruit abscission. Moreover, we found that NAA, similar to shading, directly interfered with leaf photosynthesis by repressing photosystem II (PSII) efficiency within 10 minutes of treatment, suggesting that NAA and shading induced some of the same early responses due to reduced photosynthesis, which concurred with changes in hormone signaling pathways and triggered fruit abscission. Conclusions This study provides an extensive transcriptome study and a good platform for further investigation of possible regulatory genes involved in the induction of young fruit abscission in apple, which will enable us to better understand the mechanism of fruit thinning and facilitate the selection of potential chemicals for the thinning programs in apple. PMID:22003957

Transcriptomics of shading-induced and NAA-induced abscission in apple (Malus domestica) reveals a shared pathway involving reduced photosynthesis, alterations in carbohydrate transport and signaling and hormone crosstalk.

PubMed

Zhu, Hong; Dardick, Chris D; Beers, Eric P; Callanhan, Ann M; Xia, Rui; Yuan, Rongcai

2011-10-17

Naphthaleneacetic acid (NAA), a synthetic auxin analogue, is widely used as an effective thinner in apple orchards. When applied shortly after fruit set, some fruit abscise leading to improved fruit size and quality. However, the thinning results of NAA are inconsistent and difficult to predict, sometimes leading to excess fruit drop or insufficient thinning which are costly to growers. This unpredictability reflects our incomplete understanding of the mode of action of NAA in promoting fruit abscission. Here we compared NAA-induced fruit drop with that caused by shading via gene expression profiling performed on the fruit abscission zone (FAZ), sampled 1, 3, and 5 d after treatment. More than 700 genes with significant changes in transcript abundance were identified from NAA-treated FAZ. Combining results from both treatments, we found that genes associated with photosynthesis, cell cycle and membrane/cellular trafficking were downregulated. On the other hand, there was up-regulation of genes related to ABA, ethylene biosynthesis and signaling, cell wall degradation and programmed cell death. While the differentially expressed gene sets for NAA and shading treatments shared only 25% identity, NAA and shading showed substantial similarity with respect to the classes of genes identified. Specifically, photosynthesis, carbon utilization, ABA and ethylene pathways were affected in both NAA- and shading-induced young fruit abscission. Moreover, we found that NAA, similar to shading, directly interfered with leaf photosynthesis by repressing photosystem II (PSII) efficiency within 10 minutes of treatment, suggesting that NAA and shading induced some of the same early responses due to reduced photosynthesis, which concurred with changes in hormone signaling pathways and triggered fruit abscission. This study provides an extensive transcriptome study and a good platform for further investigation of possible regulatory genes involved in the induction of young fruit abscission in apple, which will enable us to better understand the mechanism of fruit thinning and facilitate the selection of potential chemicals for the thinning programs in apple.

Consistency in color parameters of a commonly used shade guide.

PubMed

Tashkandi, Esam

2010-01-01

The use of shade guides to assess the color of natural teeth subjectively remains one of the most common means for dental shade assessment. Any variation in the color parameters of the different shade guides may lead to significant clinical implications. Particularly, since the communication between the clinic and the dental laboratory is based on using the shade guide designation. The purpose of this study was to investigate the consistency of the L∗a∗b∗ color parameters of a sample of a commonly used shade guide. The color parameters of a total of 100 VITAPAN Classical Vacuum shade guide (VITA Zahnfabrik, Bad Säckingen, Germany(were measured using a X-Rite ColorEye 7000A Spectrophotometer (Grand Rapids, Michigan, USA). Each shade guide consists of 16 tabs with different designations. Each shade tab was measured five times and the average values were calculated. The ΔE between the average L∗a∗b∗ value for each shade tab and the average of the 100 shade tabs of the same designation was calculated. Using the Student t-test analysis, no significant differences were found among the measured sample. There is a high consistency level in terms of color parameters of the measured VITAPAN Classical Vacuum shade guide sample tested.

A scaleable methodology for assessing the impacts of urban shade on the summer electricity use of residential homes

NASA Astrophysics Data System (ADS)

Taylor, Robert Vanderlei

Our cities are experiencing unprecedented growth while net global temperatures continue to trend warmer making sustainable urban development and energy conservation pressing public issues. This research explores how urban landscaping -- in particular trees and buildings -- affect summer electricity use in residential homes. I studied the interactions of urban shade and temperature to explore how vegetation distribution and intensity could play a meaningful role in heat mitigation in urban environments. Only a few studies have reconciled modeled electricity savings from tree shade with actual electricity consumption data. This research proposes a methodology for modeling the isolated effects of urban shade (tree shade vs building shade) on buildings' summertime electricity consumption from micro to mesoscales, empirically validating the modeled shade with actual electricity billing data, and comparing the electric energetic impact of tree shade effects with building shade effects. This proposed methodology seeks to resolve three primary research questions: 1) What are the modeled quantities of urban shade associated with the area of interest (AOI)? 2) To what extent do the effects of shading from trees and buildings mitigate summertime heat in the AOI? 2) To what extent do the shade effects from trees and buildings reduce summertime electricity consumption in the AOI?

Heat stress in cows at pasture and benefit of shade in a temperate climate region.

PubMed

Veissier, Isabelle; Van Laer, Eva; Palme, Rupert; Moons, Christel P H; Ampe, Bart; Sonck, Bart; Andanson, Stéphane; Tuyttens, Frank A M

2018-04-01

Under temperate climates, cattle are often at pasture in summer and are not necessarily provided with shade. We aimed at evaluating in a temperate region (Belgium) to what extent cattle may suffer from heat stress (measured through body temperature, respiration rate and panting score, cortisol or its metabolites in milk, and feces on hot days) and at assessing the potential benefits of shade. During the summer of 2012, 20 cows were kept on pasture without access to shade. During the summer of 2011, ten cows had access to shade (young trees with shade cloth hung between them), whereas ten cows had no access. Climatic conditions were quantified by the Heat Load Index (HLI). In animals without access to shade respiration rates, panting scores, rectal temperatures, and milk cortisol concentrations increased as HLI increased in both 2011 and 2012. Fecal cortisol metabolites varied with HLI in 2011 only. When cattle had access to shade, their use of shade increased as the HLI increased. This effect was more pronounced during the last part of the summer, possibly due to better acquaintance with the shade construction. In this case, shade use increased to 65% at the highest HLI (79). Shade tempered the effects on respiration, rectal temperature, and fecal cortisol metabolites. Milk cortisol was not influenced by HLI for cows using shade for > 10% of the day. Therefore, even in temperate areas, cattle may suffer from heat when they are at pasture in summer and providing shade can reduce such stress.

Assessment of microclimate conditions under artificial shades in a ginseng field.

PubMed

Lee, Kyu Jong; Lee, Byun-Woo; Kang, Je Yong; Lee, Dong Yun; Jang, Soo Won; Kim, Kwang Soo

2016-01-01

Knowledge on microclimate conditions under artificial shades in a ginseng field would facilitate climate-aware management of ginseng production. Weather data were measured under the shade and outside the shade at two fields located in Gochang-gun and Jeongeup-si, Korea, in 2011 and 2012 seasons to assess temperature and humidity conditions under the shade. An empirical approach was developed and validated for the estimation of leaf wetness duration (LWD) using weather measurements outside the shade as inputs to the model. Air temperature and relative humidity were similar between under the shade and outside the shade. For example, temperature conditions favorable for ginseng growth, e.g., between 8°C and 27°C, occurred slightly less frequently in hours during night times under the shade (91%) than outside (92%). Humidity conditions favorable for development of a foliar disease, e.g., relative humidity > 70%, occurred slightly more frequently under the shade (84%) than outside (82%). Effectiveness of correction schemes to an empirical LWD model differed by rainfall conditions for the estimation of LWD under the shade using weather measurements outside the shade as inputs to the model. During dew eligible days, a correction scheme to an empirical LWD model was slightly effective (10%) in reducing estimation errors under the shade. However, another correction approach during rainfall eligible days reduced errors of LWD estimation by 17%. Weather measurements outside the shade and LWD estimates derived from these measurements would be useful as inputs for decision support systems to predict ginseng growth and disease development.

Assessment of microclimate conditions under artificial shades in a ginseng field

PubMed Central

Lee, Kyu Jong; Lee, Byun-Woo; Kang, Je Yong; Lee, Dong Yun; Jang, Soo Won; Kim, Kwang Soo

2015-01-01

Background Knowledge on microclimate conditions under artificial shades in a ginseng field would facilitate climate-aware management of ginseng production. Methods Weather data were measured under the shade and outside the shade at two fields located in Gochang-gun and Jeongeup-si, Korea, in 2011 and 2012 seasons to assess temperature and humidity conditions under the shade. An empirical approach was developed and validated for the estimation of leaf wetness duration (LWD) using weather measurements outside the shade as inputs to the model. Results Air temperature and relative humidity were similar between under the shade and outside the shade. For example, temperature conditions favorable for ginseng growth, e.g., between 8°C and 27°C, occurred slightly less frequently in hours during night times under the shade (91%) than outside (92%). Humidity conditions favorable for development of a foliar disease, e.g., relative humidity > 70%, occurred slightly more frequently under the shade (84%) than outside (82%). Effectiveness of correction schemes to an empirical LWD model differed by rainfall conditions for the estimation of LWD under the shade using weather measurements outside the shade as inputs to the model. During dew eligible days, a correction scheme to an empirical LWD model was slightly effective (10%) in reducing estimation errors under the shade. However, another correction approach during rainfall eligible days reduced errors of LWD estimation by 17%. Conclusion Weather measurements outside the shade and LWD estimates derived from these measurements would be useful as inputs for decision support systems to predict ginseng growth and disease development. PMID:26843827

Recreating the shading effects of ship wake induced turbidity to test acclimation responses in the seagrass Thalassia hemprichii

NASA Astrophysics Data System (ADS)

Browne, Nicola K.; Yaakub, Siti Maryam; Tay, Jason K. L.; Todd, Peter A.

2017-12-01

Elevated sediment delivery and resuspension in coastal waters from human activities such as shipping can have detrimental effects on seagrass health by limiting light penetration. Managing seagrasses requires knowledge of their light acclamatory abilities so guidelines for coastal activities (e.g. ship movements) that influence sediment dynamics can be created. Guidelines typically focus on ensuring that seagrasses are able to meet their minimal light requirements (MLR). MLRs can be achieved by different light regimes, but it remains unknown whether a chronically low yet stable light regime is less or more detrimental than a highly variable regime with periods of extreme low to no light. To test this, we compared the physiological and morphological responses of Thalassia hemprichii among three light regimes: an open control (30-40% ambient light), a shaded control with (11-15% ambient light), and a fluctuating shade (4-30% ambient light). The MLR for the T. hemprichii we studied was lower (4-10% ambient light) than previous reports (mean = 18%) illustrating enhanced light acclimation in Singapore's chronically turbid waters. Seagrass shoots in the shaded control, however, exhibited significantly more morphological stress symptoms, with reduced shoot growth and lower below ground biomass. These data suggest that for seagrass exposed to periods of acute light stress, energetic costs associated with photo-acclimation to more variable light regimes can be offset if the plant can meet its daily light requirements during periods of high light. Management of seagrass beds should incorporate regular light monitoring and move towards an adaptive feedback-based approach to ensure the long-term viability of these vulnerable ecosystems.

Genotypic Diversity and Short-term Response to Shading Stress in a Threatened Seagrass: Does Low Diversity Mean Low Resilience?

PubMed

Evans, Suzanna M; Vergés, Adriana; Poore, Alistair G B

2017-01-01

Seagrasses that are predominantly clonal often have low levels of genetic variation within populations and predicting their response to changing conditions requires an understanding of whether genetic variation confers increased resistance to environmental stressors. A higher level of genetic diversity is assumed to benefit threatened species due to the increased likelihood of those populations having genotypes that can persist under environmental change. To test this idea, we conducted an in situ shading experiment with six geographically distinct meadows of the threatened seagrass Posidonia australis that vary in genetic diversity. Different genotypes within meadows varied widely in their physiological and growth responses to reduced light during a simulated short-term turbidity event. The majority of meadows were resistant to the sudden reduction in light availability, but a small subset of meadows with low genotypic diversity were particularly vulnerable to the early effects of shading, showing substantially reduced growth rates after only 3 weeks. Using the photosynthetic performance (maximum quantum yield) of known genotypes, we simulated meadows of varying genetic diversity to show that higher diversity can increase meadow resilience to stress by ensuring a high probability of including a high-performing genotype. These results support the hypothesis that complementarity among genotypes enhances the adaptive capacity of a population, and have significant implications for the conservation of declining P. australis meadows close to the species range edge on the east coast of Australia, where the genotypic diversity is low.

Absence of sex differential plasticity to light availability during seed maturation in Geranium sylvaticum.

PubMed

Varga, Sandra; Laaksonen, Ester; Siikamäki, Pirkko; Kytöviita, Minna-Maarit

2015-01-01

Sex-differential plasticity (SDP) hypothesis suggests that since hermaphrodites gain fitness through both pollen and seed production they may have evolved a higher degree of plasticity in their reproductive strategy compared to females which achieve fitness only through seed production. SDP may explain the difference in seed production observed between sexes in gynodioecious species in response to resource (nutrients or water) availability. In harsh environments, hermaphrodites decrease seed production whereas females keep it relatively similar regardless of the environmental conditions. Light availability can be also a limiting resource and thus could theoretically affect differently female and hermaphrodite seed output even though this ecological factor has been largely overlooked. We tested whether the two sexes in the gynodioecious species Geranium sylvaticum differ in their tolerance to light limitation during seed maturation in the field. We used a fully factorial block experiment exposing female and hermaphrodite plants to two different light environments (control and shade) after their peak flowering period. Specifically, we measured fruit and seed production in response to decreased light availability and compared it between the sexes. Shading reduced the number of fruits and seeds produced, but the decrease was similar between the sexes. Furthermore, shading delayed seed production by three days in both sexes, but did not affect seed mass, seed P content, or the probability of re-flowering the following year. Our results give no evidence for reproductive SDP in response to light during seed maturation.

Genotypic Diversity and Short-term Response to Shading Stress in a Threatened Seagrass: Does Low Diversity Mean Low Resilience?

PubMed Central

Evans, Suzanna M.; Vergés, Adriana; Poore, Alistair G. B.

2017-01-01

Seagrasses that are predominantly clonal often have low levels of genetic variation within populations and predicting their response to changing conditions requires an understanding of whether genetic variation confers increased resistance to environmental stressors. A higher level of genetic diversity is assumed to benefit threatened species due to the increased likelihood of those populations having genotypes that can persist under environmental change. To test this idea, we conducted an in situ shading experiment with six geographically distinct meadows of the threatened seagrass Posidonia australis that vary in genetic diversity. Different genotypes within meadows varied widely in their physiological and growth responses to reduced light during a simulated short-term turbidity event. The majority of meadows were resistant to the sudden reduction in light availability, but a small subset of meadows with low genotypic diversity were particularly vulnerable to the early effects of shading, showing substantially reduced growth rates after only 3 weeks. Using the photosynthetic performance (maximum quantum yield) of known genotypes, we simulated meadows of varying genetic diversity to show that higher diversity can increase meadow resilience to stress by ensuring a high probability of including a high-performing genotype. These results support the hypothesis that complementarity among genotypes enhances the adaptive capacity of a population, and have significant implications for the conservation of declining P. australis meadows close to the species range edge on the east coast of Australia, where the genotypic diversity is low. PMID:28855915

Effects of sunlight exposure on grapevine powdery mildew development.

PubMed

Austin, Craig N; Wilcox, Wayne F

2012-09-01

Natural and artificially induced shade increased grapevine powdery mildew (Erysiphe necator) severity in the vineyard, with foliar disease severity 49 to 75% higher relative to leaves in full sun, depending on the level of natural shading experienced and the individual experiment. Cluster disease severities increased by 20 to 40% relative to those on check vines when ultraviolet (UV) radiation was filtered from sunlight reaching vines in artificial shading experiments. Surface temperatures of leaves in full sunlight averaged 5 to 8°C higher than those in natural shade, and in one experiment, filtering 80% of all wavelengths of solar radiation, including longer wavelengths responsible for heating irradiated tissues, increased disease more than filtering UV alone. In controlled environment experiments, UV-B radiation reduced germination of E. necator conidia and inhibited both colony establishment (hyphal formation and elongation) and maturity (latent period). Inhibitory effects of UV-B radiation were significantly greater at 30°C than at 20 or 25°C. Thus, sunlight appears to inhibit powdery mildew development through at least two mechanisms, i.e., (i) UV radiation's damaging effects on exposed conidia and thalli of the pathogen; and (ii) elevating temperatures of irradiated tissues to a level supraoptimal or inhibitory for pathogen development. Furthermore, these effects are synergistic at temperatures near the upper threshold for disease development.

Establishing a Wild, Ex Situ Population of a Critically Endangered Shade-Tolerant Rainforest Conifer: A Translocation Experiment

PubMed Central

Zimmer, Heidi C.; Offord, Catherine A.; Auld, Tony D.; Baker, Patrick J.

2016-01-01

Translocation can reduce extinction risk by increasing population size and geographic range, and is increasingly being used in the management of rare and threatened plant species. A critical determinant of successful plant establishment is light environment. Wollemia nobilis (Wollemi pine) is a critically endangered conifer, with a wild population of 83 mature trees and a highly restricted distribution of less than 10 km2. We used under-planting to establish a population of W. nobilis in a new rainforest site. Because its optimal establishment conditions were unknown, we conducted an experimental translocation, planting in a range of different light conditions from deeply shaded to high light gaps. Two years after the experimental translocation, 85% of plants had survived. There were two distinct responses: very high survival (94%) but very low growth, and lower survival (69%) and higher growth, associated with initial plant condition. Overall survival of translocated W. nobilis was strongly increased in planting sites with higher light, in contrast to previous studies demonstrating long-term survival of wild W. nobilis juveniles in deep shade. Translocation by under-planting may be useful in establishing new populations of shade-tolerant plant species, not least by utilizing the range of light conditions that occur in forest understories. PMID:27403527

Establishing a Wild, Ex Situ Population of a Critically Endangered Shade-Tolerant Rainforest Conifer: A Translocation Experiment.

PubMed

Zimmer, Heidi C; Offord, Catherine A; Auld, Tony D; Baker, Patrick J

2016-01-01

Translocation can reduce extinction risk by increasing population size and geographic range, and is increasingly being used in the management of rare and threatened plant species. A critical determinant of successful plant establishment is light environment. Wollemia nobilis (Wollemi pine) is a critically endangered conifer, with a wild population of 83 mature trees and a highly restricted distribution of less than 10 km2. We used under-planting to establish a population of W. nobilis in a new rainforest site. Because its optimal establishment conditions were unknown, we conducted an experimental translocation, planting in a range of different light conditions from deeply shaded to high light gaps. Two years after the experimental translocation, 85% of plants had survived. There were two distinct responses: very high survival (94%) but very low growth, and lower survival (69%) and higher growth, associated with initial plant condition. Overall survival of translocated W. nobilis was strongly increased in planting sites with higher light, in contrast to previous studies demonstrating long-term survival of wild W. nobilis juveniles in deep shade. Translocation by under-planting may be useful in establishing new populations of shade-tolerant plant species, not least by utilizing the range of light conditions that occur in forest understories.

Metabolic Flux Redirection and Transcriptomic Reprogramming in the Albino Tea Cultivar ‘Yu-Jin-Xiang’ with an Emphasis on Catechin Production

PubMed Central

Liu, Guo-Feng; Han, Zhuo-Xiao; Feng, Lin; Gao, Li-Ping; Gao, Ming-Jun; Gruber, Margaret Y.; Zhang, Zhao-Liang; Xia, Tao; Wan, Xiao-Chun; Wei, Shu

2017-01-01

In this study, shade-induced conversion from a young pale/yellow leaf phenotype to a green leaf phenotype was studied using metabolic and transcriptomic profiling and the albino cultivar ‘Yu-Jin-Xiang’ (‘YJX’) of Camellia sinensis for a better understanding of mechanisms underlying the phenotype shift and the altered catechin and theanine production. Shaded leaf greening resulted from an increase in leaf chlorophyll and carotenoid abundance and chloroplast development. A total of 1,196 differentially expressed genes (DEGs) were identified between the ‘YJX’ pale and shaded green leaves, and these DEGs affected ‘chloroplast organization’ and ‘response to high light’ besides many other biological processes and pathways. Metabolic flux redirection and transcriptomic reprogramming were found in flavonoid and carotenoid pathways of the ‘YJX’ pale leaves and shaded green leaves to different extents compared to the green cultivar ‘Shu-Cha-Zao’. Enhanced production of the antioxidant quercetin rather than catechin biosynthesis was correlated positively with the enhanced transcription of FLAVONOL SYNTHASE and FLAVANONE/FLAVONOL HYDROXYLASES leading to quercetin accumulation and negatively correlated to suppressed LEUCOANTHOCYANIDIN REDUCTASE, ANTHOCYANIDIN REDUCTASE and SYNTHASE leading to catechin biosynthesis. The altered levels of quercetin and catechins in ‘YJX’ will impact on its tea flavor and health benefits. PMID:28332598

Differential responses of Bolivian timber species to prescribed fire and other gap treatments

Treesearch

Debora K. Kennard; Francis E. Putz

2005-01-01

We followed the establishment and growth response of 13 commercial tree species to canopy opening, above-ground biomass removal, and experimental burns of low and high intensities in a lowland dry forest in Bolivia. Three patterns of response to treatments were observed among the most abundant commercial tree species. (1) Shade-intolerant species regenerated mostly...

Shade-Induced Action Potentials in Helianthus annuus L. Originate Primarily from the Epicotyl

PubMed Central

Stephens, Nicholas R; Cleland, Robert E; Van Volkenburgh, Elizabeth

2006-01-01

Repeated observations that shading (a drastic reduction in illumination rate) increased the generation of spikes (rapidly reversed depolarizations) in leaves and stems of many cucumber and sunflower plants suggests a phenomenon widespread among plant organs and species. Although shaded leaves occasionally generate spikes and have been suggested to trigger systemic action potentials (APs) in sunflower stems, we never found leaf-generated spikes to propagate out of the leaf and into the stem. On the contrary, our data consistently implicate the epicotyl as the location where most spikes and APs (propagating spikes) originate. Microelectrode studies of light and shading responses in mesophyll cells of leaf strips and in epidermis/cortex cells of epicotyl segments confirm this conclusion and show that spike induction is not confined to intact plants. 90% of the epicotyl-generated APs undergo basipetal propagation to the lower epicotyl, hypocotyl and root. They propagate with an average rate of 2 ± 0.3 mm s−1 and always undergo a large decrement from the hypocotyl to the root. The few epicotyl-derived APs that can be tracked to leaf blades (< 10%) undergo either a large decrement or fail to be transmitted at all. Occasionally (5% of the observations) spikes were be generated in hypocotyl and lower epicotyl that moved towards the upper epicotyl unaltered, decremented, or amplified. This study confirms that plant APs arise to natural, nontraumatic changes. In simultaneous recordings with epicotyl growth, AP generation was found to parallel the acceleration of stem growth under shade. The possible relatedness of both processes must be further investigated. PMID:19521471

Influence of light source, polarization, education, and training on shade matching quality.

PubMed

Clary, Jacqueline A; Ontiveros, Joe C; Cron, Stanley G; Paravina, Rade D

2016-07-01

Many factors influence the quality of shade selection, and isolating how significantly each of these factors influences results is difficult. The purpose of this in vitro study was to compare results of shade matching using handheld lights with or without a polarizing filter with results obtained using a professional viewing booth and to analyze the influence of education and training on shade selection outcome. A total of 96 third-year dental students (evaluators) were randomly separated into 4 groups. Each group was assigned 1 of 2 handheld shade-matching devices (lights) with or without a polarizing filter. Each group performed a shade matching exercise using the handheld light or a professional viewing booth. The exercise consisted of matching shade tabs placed in a typodont to a commercial shade guide. Each group repeated this procedure 4 times over a 9-week period. A lecture on shade matching was presented at the fifth week of the study, between "before" and "after" shade matching procedures. Shade matching scores with handheld lights (7.8) were higher than scores of shade matching with the viewing booth (7.2). The mean scores for before (7.2) and after (7.8) shade matching (with education and training in between) were significantly different. The combined effect of light and education and training improved the shade matching score by 1.2, from 6.8 in the before sessions using the viewing booth to 8.0 in the after sessions using handheld lights. A 21% increase in the number of evaluators who selected 1 of 4 best matches was recorded, 10% for handheld lights versus viewing booth after education and training versus before sessions and 11% between after sessions using handheld lights versus before sessions using viewing booth. Within the limits of the study, the shade matching scores with handheld lights were significantly better than the results obtained using a viewing booth (P<.01). Using a handheld light with or without a polarizing filter did not influence shade matching results. Mean shade matching scores were significantly better after education and training (P<.01). Light combined with education and training resulted in the greatest increase in shade matching quality. Copyright © 2016 Editorial Council for the Journal of Prosthetic Dentistry. Published by Elsevier Inc. All rights reserved.

Achieving the desired colour in discoloured teeth, using leucite-based CAD-CAM laminate systems.

PubMed

Turgut, Sedanur; Bagis, Bora; Ayaz, Elif Aydogan

2014-01-01

To evaluate the cumulative effect of the abutment tooth and resin cement colour on the resultant optical properties of porcelain laminate veneers (PLVs) fabricated with leucite-based CAD/CAM blocks with different shades and thicknesses. A total of 224 ceramic specimens were prepared from the IPS Empress CAD with four different shades of HT-A1, LT-A1, MT-A1 and Bl-1. Resin composite discs were prepared with shade A3.5. For the cementation, 4 different shades of light-cure resin cements were chosen. L*, a*, and b* values, as well as the chroma (C) and hue (h) values of each cemented ceramic and the A1 shade table, were recorded. L*, Cab* and hab* values of the cemented ceramics were influenced by ceramic shade, ceramic thickness, cement shade, and interaction terms of the three variables. There were significant differences between the 1-mm-thick ceramic veneers that exhibited higher L* and lower Cab* values compared with veneers that were 0.5mm in thickness. Using the Tr shade cement resulted in lower L* and higher Cab* values for all thicknesses and ceramic shades, whereas the WO shade cement resulted in higher values. The selected colour of a laminate restoration is significantly affected by the ceramic shade, ceramic thickness and resin cement shade. Using the WO shade resin cement seems to be more effective in masking the discoloured abutment tooth. Clinicians should also select the translucency level of the ceramic block with the shade when a leucite-based CAD/CAM system is chosen to treat a discoloured tooth. Copyright © 2013 Elsevier Ltd. All rights reserved.

Loss of thermal refugia near equatorial range limits.

PubMed

Lima, Fernando P; Gomes, Filipa; Seabra, Rui; Wethey, David S; Seabra, Maria I; Cruz, Teresa; Santos, António M; Hilbish, Thomas J

2016-01-01

This study examines the importance of thermal refugia along the majority of the geographical range of a key intertidal species (Patella vulgata Linnaeus, 1758) on the Atlantic coast of Europe. We asked whether differences between sun-exposed and shaded microhabitats were responsible for differences in physiological stress and ecological performance and examined the availability of refugia near equatorial range limits. Thermal differences between sun-exposed and shaded microhabitats are consistently associated with differences in physiological performance, and the frequency of occurrence of high temperatures is most probably limiting the maximum population densities supported at any given place. Topographical complexity provides thermal refugia throughout most of the distribution range, although towards the equatorial edges the magnitude of the amelioration provided by shaded microhabitats is largely reduced. Importantly, the limiting effects of temperature, rather than being related to latitude, seem to be tightly associated with microsite variability, which therefore is likely to have profound effects on the way local populations (and consequently species) respond to climatic changes. © 2015 John Wiley & Sons Ltd.

Genotype x shade effects for western hemlock.

Treesearch

G.R. Johnson; C. Cartwright

2005-01-01

Western hemlock (Tsuga heterophylla (Raf.) Sarg.) families were grown under different levels of shade for 2 or 3 years at two nursery sites to determine whether families performed differently relative to one another in the different shade environments. Differences were found both for levels of shade and families, but no family x shade interaction...

Dental Shade Guide Variability for Hues B, C, and D Using Cross-Polarized Photography.

PubMed

Sampaio, Camila S; Gurrea, Jon; Gurrea, Marta; Bruguera, August; Atria, Pablo J; Janal, Malvin; Bonfante, Estevam A; Coelho, Paulo G; Hirata, Ronaldo

2018-04-20

This study evaluated the color variability of hues B, C, and D between the VITA Classical shade guide (Vita Zahnfabrik) and four other VITA-coded ceramic shade guides using a digital camera (Canon EOS 60D) and computer software (Adobe Photoshop CC). A cross-polarizing filter was used to standardize external light sources influencing color match. A total of 275 pictures were taken, 5 per shade tab, for 11 shades (B1, B2, B3, B4, C1, C2, C3, C4, D2, D3, and D4), from the following shade guides: VITA Classical (control); IPS e.max Ceram (Ivoclar Vivadent); IPS d.SIGN (Ivoclar Vivadent); Initial ZI (GC); and Creation CC (Creation Willi Geller). Pictures were evaluated using Adobe Photoshop CC for standardization of hue, chroma, and value between shade tabs. The VITA-coded shade guides evaluated here showed an overall unmatched shade in all their tabs when compared to the control, suggesting that shade selection should be made with the corresponding manufacturer guide of the ceramic intended for the final restoration.

Slow induction of photosynthesis on shade to sun transitions in wheat may cost at least 21% of productivity.

PubMed

Taylor, Samuel H; Long, Stephen P

2017-09-26

Wheat is the second most important direct source of food calories in the world. After considerable improvement during the Green Revolution, increase in genetic yield potential appears to have stalled. Improvement of photosynthetic efficiency now appears a major opportunity in addressing the sustainable yield increases needed to meet future food demand. Effort, however, has focused on increasing efficiency under steady-state conditions. In the field, the light environment at the level of individual leaves is constantly changing. The speed of adjustment of photosynthetic efficiency can have a profound effect on crop carbon gain and yield. Flag leaves of wheat are the major photosynthetic organs supplying the grain of wheat, and will be intermittently shaded throughout a typical day. Here, the speed of adjustment to a shade to sun transition in these leaves was analysed. On transfer to sun conditions, the leaf required about 15 min to regain maximum photosynthetic efficiency. In vivo analysis based on the responses of leaf CO 2 assimilation ( A ) to intercellular CO 2 concentration ( c i ) implied that the major limitation throughout this induction was activation of the primary carboxylase of C3 photosynthesis, ribulose-1,5-bisphosphate carboxylase/oxygenase (Rubisco). This was followed in importance by stomata, which accounted for about 20% of the limitation. Except during the first few seconds, photosynthetic electron transport and regeneration of the CO 2 acceptor molecule, ribulose-1,5-bisphosphate (RubP), did not affect the speed of induction. The measured kinetics of Rubisco activation in the sun and de-activation in the shade were predicted from the measurements. These were combined with a canopy ray tracing model that predicted intermittent shading of flag leaves over the course of a June day. This indicated that the slow adjustment in shade to sun transitions could cost 21% of potential assimilation.This article is part of the themed issue 'Enhancing photosynthesis in crop plants: targets for improvement'. © 2017 The Authors.

Adaptive Criterion Setting in Perceptual Decision Making

ERIC Educational Resources Information Center

Stuttgen, Maik C.; Yildiz, Ali; Gunturkun, Onur

2011-01-01

Pigeons responded in a perceptual categorization task with six different stimuli (shades of gray), three of which were to be classified as "light" or "dark", respectively. Reinforcement probability for correct responses was varied from 0.2 to 0.6 across blocks of sessions and was unequal for correct light and dark responses. Introduction of a new…

Evaluation of different shades to improve dairy cattle well-being in Argentina

NASA Astrophysics Data System (ADS)

Valtorta, S. E.; Leva, Perla E.; Gallardo, Miriam R.

Two tree shades (TS1 and TS2) and an artificial shade structure (AS) were evaluated using black globe temperatures (BGTs) to assess their effectiveness in reducing heat load. The artificial structure consisted of a black woven polypropylene cloth providing 80% shade, mounted on 2.5-m-high eucalyptus posts. The work was carried out at Rafaela Experimental Station, Argentina, during the summer (January and February) 1994. BGTs and floor temperatures were measured in concrete floor holding pens with and without artifical shade. The results showed no difference between TS1, TS2 and AS, their average BGTs being 30.2 (SD 0.58), 29.0 (SD 0.70) and 30.2 (SD 0.74)°C, respectively. BGTs under all three shades were significantly lower (P<0.01) than the average BGT recorded outside: 35.5 (SD 1.12)°C. Average BGTs in holding pens were 32.4 (SD 1.38) and 39.9 (SD 1.91)°C for shaded and non-shaded areas (P<0.01). The corresponding average floor temperatures were 27.8 (SD 0.68)°C and 47.7 (SD 2.13)°C (P<0.01). To assess the effects of shade on animal well-being, afternoon rectal temperatures (RT) and respiratory rate (respirations per minute, RR) of lactating cows were recorded twice a week. Rectal temperatures were significantly higher for non-shaded cows (P<0.01), mean RT being 40.1 (SD 0.65)°C vs 39.3 (SD 0.42)°C for the shaded animals. Corresponding RRs were 78.9 (SD 18.0) and 60.7 (SD 10.6) (P<0.05). It was concluded that: (1) tree and artificial shades produced similar effects, (2) shading the holding pen with an 80% shading cloth was effective in reducing heat load and floor temperatures, and (3) access to shade in our pasture-based system improved animal well-being.

A Novel Two-Step Method for Screening Shade Tolerant Mutant Plants via Dwarfism

PubMed Central

Li, Wei; Katin-Grazzini, Lorenzo; Krishnan, Sanalkumar; Thammina, Chandra; El-Tanbouly, Rania; Yer, Huseyin; Merewitz, Emily; Guillard, Karl; Inguagiato, John; McAvoy, Richard J.; Liu, Zongrang; Li, Yi

2016-01-01

When subjected to shade, plants undergo rapid shoot elongation, which often makes them more prone to disease and mechanical damage. Shade-tolerant plants can be difficult to breed; however, they offer a substantial benefit over other varieties in low-light areas. Although perennial ryegrass (Lolium perenne L.) is a popular species of turf grasses because of their good appearance and fast establishment, the plant normally does not perform well under shade conditions. It has been reported that, in turfgrass, induced dwarfism can enhance shade tolerance. Here we describe a two-step procedure for isolating shade tolerant mutants of perennial ryegrass by first screening for dominant dwarf mutants, and then screening dwarf plants for shade tolerance. The two-step screening process to isolate shade tolerant mutants can be done efficiently with limited space at early seedling stages, which enables quick and efficient isolation of shade tolerant mutants, and thus facilitates development of shade tolerant new cultivars of turfgrasses. Using the method, we isolated 136 dwarf mutants from 300,000 mutagenized seeds, with 65 being shade tolerant (0.022%). When screened directly for shade tolerance, we recovered only four mutants from a population of 150,000 (0.003%) mutagenized seeds. One shade tolerant mutant, shadow-1, was characterized in detail. In addition to dwarfism, shadow-1 and its sexual progeny displayed high degrees of tolerance to both natural and artificial shade. We showed that endogenous gibberellin (GA) content in shadow-1 was higher than wild-type controls, and shadow-1 was also partially GA insensitive. Our novel, simple and effective two-step screening method should be applicable to breeding shade tolerant cultivars of turfgrasses, ground covers, and other economically important crop plants that can be used under canopies of existing vegetation to increase productivity per unit area of land. PMID:27752260

Evaluation of different shades to improve dairy cattle well-being in Argentina.

PubMed

Valtorta, S E; Leva, P E; Gallardo, M R

1997-11-01

Two tree shades (TS1 and TS2) and an artificial shade structure (AS) were evaluated using black globe temperatures (BGTs) to assess their effectiveness in reducing heat load. The artificial structure consisted of a black woven polypropylene cloth providing 80% shade, mounted on 2.5-m-high eucalyptus posts. The work was carried out at Rafaela Experimental Station, Argentina, during the summer (January and February) 1994. BGTs and floor temperatures were measured in concrete floor holding pens with and without artificial shade. The results showed no difference between TS1, TS2 and AS, their average BGTs being 30.2 (SD 0.58), 29.0 (SD 0.70) and 30.2 (SD 0.74) degrees C, respectively. BGTs under all three shades were significantly lower (P < 0.01) than the average BGT recorded outside: 35.5 (SD 1.12) degrees C. Average BGTs in holding pens were 32.4 (SD 1.38) and 39.9 (SD 1.91) degrees C for shaded and non-shaded areas (P < 0.01). The corresponding average floor temperatures were 27.8 (SD 0.68) degrees C and 47.7 (SD 2.13) degrees C (P < 0.01). To assess the effects of shade on animal well-being, afternoon rectal temperatures (RT) and respiratory rate (respirations per minute, RR) of lactating cows were recorded twice a week. Rectal temperatures were significantly higher for non-shaded cows (P < 0.01), mean RT being 40.1 (SD 0.65) degrees C vs 39.3 (SD 0.42) degrees C for the shaded animals. Corresponding RRs were 78.9 (SD 18.0) and 60.7 (SD 10.6) (P < 0.05). It was concluded that: (1) tree and artificial shades produced similar effects, (2) shading the holding pen with an 80% shading cloth was effective in reducing heat load and floor temperatures, and (3) access to shade in our pasture-based system improved animal well-being.

Comparison of accuracies of an intraoral spectrophotometer and conventional visual method for shade matching using two shade guide systems.

PubMed

Parameswaran, Vidhya; Anilkumar, S; Lylajam, S; Rajesh, C; Narayan, Vivek

2016-01-01

This in vitro study compared the shade matching abilities of an intraoral spectrophotometer and the conventional visual method using two shade guides. The results of previous investigations between color perceived by human observers and color assessed by instruments have been inconclusive. The objectives were to determine accuracies and interrater agreement of both methods and effectiveness of two shade guides with either method. In the visual method, 10 examiners with normal color vision matched target control shade tabs taken from the two shade guides (VITAPAN Classical™ and VITAPAN 3D Master™) with other full sets of the respective shade guides. Each tab was matched 3 times to determine repeatability of visual examiners. The spectrophotometric shade matching was performed by two independent examiners using an intraoral spectrophotometer (VITA Easyshade™) with five repetitions for each tab. Results revealed that visual method had greater accuracy than the spectrophotometer. The spectrophotometer; however, exhibited significantly better interrater agreement as compared to the visual method. While VITAPAN Classical shade guide was more accurate with the spectrophotometer, VITAPAN 3D Master shade guide proved better with visual method. This in vitro study clearly delineates the advantages and limitations of both methods. There were significant differences between the methods with the visual method producing more accurate results than the spectrophotometric method. The spectrophotometer showed far better interrater agreement scores irrespective of the shade guide used. Even though visual shade matching is subjective, it is not inferior and should not be underrated. Judicious combination of both techniques is imperative to attain a successful and esthetic outcome.

New entrance shade design for SIRTF

NASA Technical Reports Server (NTRS)

Davis, Paul K.

1989-01-01

A new design for the entrance shade for the Space Infrared Telescope Facility (SIRTF) is presented. The evolution of the entrance shade began with a simple frustum, symmetrical about the telescope axis, when SIRTF was expected to be Shuttle-attached. With the change to a free-flying SIRTF this frustum was cut off at an angle. The telescope will be operated so that whenever not in the earth's shadow the high side is kept toward the sun. However, the entrance shade interior itself will be so warm that the optics, including the secondary mirror and its mechanisms and support structure, will be restricted to the rear part of the barrel, termed the aftbaffle, which is shaded from the interior of the entrance shade by the forebaffle. This is best accomplished by the most recent design in which the axis of the entrance shade is offset from the telescope axis. This results in a shorter entrance shade, shorter forebaffle, and a shaded region within the barrel which is symmetrical about the telescope axis. All of these are advantageous.

Responses of Primula vulgaris to light quality in the maternal and germination environments.

PubMed

Marin, M; Blandino, C; Laverack, G; Toorop, P; Powell, A A

2018-05-22

In the model species Arabidopsis thaliana phytochromes mediate dormancy and germination responses to seasonal cues experienced during seed maturation on the maternal plants. However, the effect of the maternal light environment on seed germination in native wild species has not been studied well. This is particularly important given its practical application in the context of environmental restoration, when there can be marked changes in the canopy. Plants of Primula vulgaris were grown in the field over two vegetative seasons under four shading treatments from low to high ratio of red to far-red light (R:FR). Leaf and seed traits were assessed in response to the light treatments. The germination of seeds from these four maternal environments (pre-dispersal) was investigated at seven light and five temperature treatments (post-dispersal). Thinner leaves, larger leaf area and greater chlorophyll content were found in plants growing in reduced R:FR. Shading in the maternal environment led to increased seed size and yield, although the conditions experienced by the maternal plants had no effect on seed germination. Seeds responded strongly to the cues experienced in their immediate germination environment. Germination was always enhanced under higher R:FR conditions. The observed phenotypic trait variation plays a major role in the ability of P. vulgaris to grow in a wide range of light conditions. However, the increased germination capacity in response to a higher R:FR for all maternal environments suggests potential for seedling establishment under vegetative shade only in the presence of canopy gaps. This article is protected by copyright. All rights reserved. This article is protected by copyright. All rights reserved.

Impacts of Topographic Shading on Surface Energy Balance of High Mountain Asia Glaciers

NASA Astrophysics Data System (ADS)

Olson, M.; Rupper, S.

2016-12-01

Topographic shading plays an important role in the energy balance of valley glaciers. While previous studies incorporate shading of varying complexity in surface energy balance models, to date, no large-scale studies have explored in depth the effects of topographic shading on glacier surface energy balance, and how these vary geographically within High Mountain Asia (HMA). Here we develop a model to examine the variability in potential insolation during the summer melt season using the ASTER GDEM and multi-hour solar geometry to simulate topographic shading on an idealized glacier. Shading is calculated in simulations utilizing a range of slopes, aspects, and latitudes. We test glacier mass balance sensitivity to these parameters for a suite of glaciers throughout HMA. Our results show that shading impacts on glaciers in HMA are highly variable across different geographic regions, but that they are largely predictable based on topographic characteristics such as slope and aspect. For example, we find in regions with steep topography and high relief that shading frequently dominates in the ablation zone rather than the accumulation zone, contrary to the findings of some previous studies. In these regions, topographic shading may play a more significant role in glacier energy balance. These results will better define the effects of topographic shading on surface energy balance, and improve model accuracy within HMA. Additionally, this topographic shading model provides a framework to quantify how shading effects vary for advancing or retreating glaciers as they respond to fluctuations in climate across HMA.

Evolutionary Agroecology: the potential for cooperative, high density, weed-suppressing cereals.

PubMed

Weiner, Jacob; Andersen, Sven B; Wille, Wibke K-M; Griepentrog, Hans W; Olsen, Jannie M

2010-09-01

Evolutionary theory can be applied to improve agricultural yields and/or sustainability, an approach we call Evolutionary Agroecology. The basic idea is that plant breeding is unlikely to improve attributes already favored by millions of years of natural selection, whereas there may be unutilized potential in selecting for attributes that increase total crop yield but reduce plants' individual fitness. In other words, plant breeding should be based on group selection. We explore this approach in relation to crop-weed competition, and argue that it should be possible to develop high density cereals that can utilize their initial size advantage over weeds to suppress them much better than under current practices, thus reducing or eliminating the need for chemical or mechanical weed control. We emphasize the role of density in applying group selection to crops: it is competition among individuals that generates the 'Tragedy of the Commons', providing opportunities to improve plant production by selecting for attributes that natural selection would not favor. When there is competition for light, natural selection of individuals favors a defensive strategy of 'shade avoidance', but a collective, offensive 'shading' strategy could increase weed suppression and yield in the high density, high uniformity cropping systems we envision.

Shade selection performed by novice dental professionals and colorimeter.

PubMed

Klemetti, E; Matela, A-M; Haag, P; Kononen, M

2006-01-01

The objective of this study was to test inter-observer variability in shade selection for porcelain restorations, using three different shade guides: Vita Lumin Vacuum, Vita 3D-Master and Procera. Nineteen young dental professionals acted as observers. The results were also compared with those of a digital colorimeter (Shade Eye Ex; Shofu, Japan). Regarding repeatability, no significant differences were found between the three shade guides, although repeatability was relatively low (33-43%). Agreement with the colorimetric results was also low (8-34%). In conclusion, shade selection shows moderate to great inter-observer variation. In teaching and standardizing the shade selection procedure, a digital colorimeter may be a useful educational tool.

Satisfaction of Dental Students, Faculty, and Patients with Tooth Shade-Matching Using a Spectrophotometer.

PubMed

Ballard, Erin; Metz, Michael J; Harris, Bryan T; Metz, Cynthia J; Chou, Jang-Ching; Morton, Dean; Lin, Wei-Shao

2017-05-01

The aims of this study were to evaluate dental students' clinical shade-matching outcomes (from subjective use of shade guide) with an objective electronic shade-matching tool (spectrophotometer); to assess patients', students', and supervising faculty members' satisfaction with the clinical shade-matching outcomes; and to assess clinicians' support for use of the spectrophotometer to improve esthetic outcomes. A total of 103 volunteer groups, each consisting of patient, dental student, and supervising faculty member at the University of Louisville, were recruited to participate in the study in 2015. Using the spectrophotometer, clinical shade-matching outcome (ΔE clinical ) and laboratory shade-matching outcome (ΔE laboratory ) were calculated. Two five-point survey items were used to assess the groups' satisfaction with the clinical shade-matching outcome and support for an objective electronic shade-matching tool in the student clinic. The results showed that both ΔE clinical (6.5±2.4) and ΔE laboratory (4.3±2.0) were outside the clinical acceptability threshold ΔE values of 2.7, when visual shade-matching method (subjective usage of shade guide) was used to fabricate definitive restorations. Characteristics of the patients, dental students, supervising faculty members, and restorations had minimal to no effect on the ΔE clinical The patients, dental students, and supervising faculty members generally had positive opinions about the clinical shade-matching outcome, despite the increased ΔE clinical observed. Overall, clinical shade-matching outcomes in this school need further improvement, but the patients' positive opinions may indicate the need to revisit the acceptability threshold ΔE value of 2.7 in the academic setting.

Comparison of accuracies of an intraoral spectrophotometer and conventional visual method for shade matching using two shade guide systems

PubMed Central

Parameswaran, Vidhya; Anilkumar, S.; Lylajam, S.; Rajesh, C.; Narayan, Vivek

2016-01-01

Background and Objectives: This in vitro study compared the shade matching abilities of an intraoral spectrophotometer and the conventional visual method using two shade guides. The results of previous investigations between color perceived by human observers and color assessed by instruments have been inconclusive. The objectives were to determine accuracies and interrater agreement of both methods and effectiveness of two shade guides with either method. Methods: In the visual method, 10 examiners with normal color vision matched target control shade tabs taken from the two shade guides (VITAPAN Classical™ and VITAPAN 3D Master™) with other full sets of the respective shade guides. Each tab was matched 3 times to determine repeatability of visual examiners. The spectrophotometric shade matching was performed by two independent examiners using an intraoral spectrophotometer (VITA Easyshade™) with five repetitions for each tab. Results: Results revealed that visual method had greater accuracy than the spectrophotometer. The spectrophotometer; however, exhibited significantly better interrater agreement as compared to the visual method. While VITAPAN Classical shade guide was more accurate with the spectrophotometer, VITAPAN 3D Master shade guide proved better with visual method. Conclusion: This in vitro study clearly delineates the advantages and limitations of both methods. There were significant differences between the methods with the visual method producing more accurate results than the spectrophotometric method. The spectrophotometer showed far better interrater agreement scores irrespective of the shade guide used. Even though visual shade matching is subjective, it is not inferior and should not be underrated. Judicious combination of both techniques is imperative to attain a successful and esthetic outcome. PMID:27746599

Patterns of shading tolerance determined from experimental ...

EPA Pesticide Factsheets

An extensive review of the experimental literature on seagrass shading evaluated the relationship between experimental light reductions, duration of experiment and seagrass response metrics to determine whether there were consistent statistical patterns. There were highly significant linear relationships of both percent biomass and percent shoot density reduction versus percent light reduction (versus controls), although unexplained variation in the data were high. Duration of exposure affected extent of response for both metrics, but was more clearly a factor in biomass response. Both biomass and shoot density showed linear responses to duration of light reduction for treatments 60%. Unexplained variation was again high, and greater for shoot density than biomass. With few exceptions, regressions of both biomass and shoot density on light reduction for individual species and for genera were statistically significant, but also tended to show high degrees of variability in data. Multivariate regressions that included both percent light reduction and duration of reduction as dependent variables increased the percentage of variation explained in almost every case. Analysis of response data by seagrass life history category (Colonizing, Opportunistic, Persistent) did not yield clearly separate response relationships in most cases. Biomass tended to show somewhat less variation in response to light reduction than shoot density, and of the two, may be the prefe

Increased growth of young citrus trees under reduced radiation load in a semi-arid climate.

PubMed

Raveh, E; Cohen, S; Raz, T; Yakir, D; Grava, A; Goldschmidt, E E

2003-01-01

This study investigated the effects of radiation heat-load reduction by shading on the growth and development of citrus trees in a warm subtropical region. The experiment was conducted from mid-June until late October when daily maximal air temperature averaged 29.3 degrees C. Two-year-old de-fruited Murcott tangor (Citrus reticulata BlancoxCitrus sinensis (L.) Osb.) trees were grown under 30% or 60% shade tunnels, or 60% flat shade (providing midday shade only), using highly reflective aluminized nets. Non-shaded trees were used as the control. Shading reduced direct more than diffuse radiation. Daily radiation was reduced by 35% for the 30% Tunnel and 60% Flat treatments, and by 55% for the 60% Tunnel. Two days of intensive measurement showed that shading increased average sunlit leaf conductance by 44% and photosynthesis by 29%. Shading did not significantly influence root and stem dry weight growth, but it increased the increment in leaf dry weight during the three month period by an average of 28% relative to the control, while final tree height in the 30% Tunnel treatment exceeded the control by 35%. Shoot to root and shoot mass ratios increased and root mass ratio decreased due to shading because of the increase in leaf dry weight. Shading increased starch concentration in leaves while the shadiest treatment, 60% Tunnel, decreased starch concentration in the roots. Carbon isotope ratio (delta(13)C) of exposed leaves that developed under shading was significantly reduced by 1.9 per thousand in the 60% Tunnel, indicating that shading increased CO(2) concentrations at the chloroplasts (C(c)), as would be expected from increased conductance. Substomatal CO(2) concentrations, C(i), computed from leaf net CO(2) assimilation rate and conductance values, also indicate that shading increases internal CO(2) concentrations. Based on tree dry mass, tree height, and total carbohydrates fractions, the 30% Tunnel and the 60% Flat were the optimal shade treatments.

Properly placed shade trees reduce summertime electricity bills in Sacramento, California

Treesearch

Geoffery H. Donovan; David R. Butry

2009-01-01

The discovery that shade trees can reduce home cooling costs is hardly surprising. Anybody who has sat under a tree on a warm summer day understands the shade benefit of trees. However, quantifying the effect a shade tree has on home energy use and carbon footprint, and identifying the optimal location for a shade tree, is less straightforward. Past studies that have...

[Effects of shading in summer and autumn on physiological and biochemical characteristics of 'Huangjinya' in Shandong Province, China].

PubMed

Tian, Yue Yue; Zhang, Li Xia; Zhang, Zheng Qun; Qiao, Ming Ming; Fan, Yan Gen

2017-03-18

In order to ensure the suitable shade model for 'Huangjinya' tea plant in Shandong Province, black or blue shading net at 55%, 70% or 85% shading rates was selected to recover tea garden in summer and autumn, then micro-climate of tea garden, leaf color, chlorophyll fluorescence parameters, growth status and biochemical composition of tea shoots were investigated.The results showed that compared with the control, light intensity and air temperature in tea garden, leaf temperature of tea plants in different shading treatments significantly decreased, while air humidity in tea garden increased. The contents of chlorophyll in the tea leaves were obviously increased with increasing the shading rate, which resulted in the leaf color becoming green. The yellowing characteristics and biochemical quality of 'Huangjinya' tea plants could be well kept in 55% shading treatments. In 70% shading treatments, 'Huangjinya' tea plants had better growth situation and higher yield with no photo-inhibition. Compared with the blue shading treatments, black shading treatments could obviously promote the growth of 'Huangjinya' tea plants, keep yellowing characteristics, and improve the quality. Therefore, the 70% black shading treatment (daily PAR values of 1.2-3.5 ten thousand lx) was appropriate for promoting the growth of 'Huangjinya' tea plants at the seedling stage. For mature tea plants, the 55% black shading treatment (daily PAR values of 1.8-5.5 ten thousand lx) could be used to keep the yellowing characteristics and to improve biochemical quality effectively, so as to give full play to its variety characteristics, to achieve goal of high quality and high yield.

Effects of shading on plant growth, flower quality and photosynthetic capacity of Rosa hybrida

NASA Astrophysics Data System (ADS)

Hou, Wei; Luo, Ya; Wang, Xiaorong; Chen, Qing; Sun, Bo; Wang, Yan; Liu, Zejing; Tang, Haoru; Zhang, Yong

2018-04-01

With the acceleration of the urbanization process, numbers of garden plants are grown in shaded environment. Rose (Rosa hybrida) is one of the most important ornamental plants used in urban green spaces. In the present study, effects of shading on plant growth, flower quality and photosynthetic capacity of R. hybrida were investigated. The results showed that shade not only delayed the initial flowering date, but also prolonged the flowering time. The physiological indices, including flower fresh weight, total anthocyanins and flavonoids contents, were lower in plants grown with shade compared with those grown in sun exposure. Overall, R. hybrida seemed to be a shade adapted plant which could be successfully grown in urban green spaces or understory with slight shade.

76 FR 41248 - Notice of Proposed Administrative Settlement Pursuant to the Comprehensive Environmental Response...

Federal Register 2010, 2011, 2012, 2013, 2014

2011-07-13

... responses to any comments received will be available for public inspection at 1445 Ross Avenue, Dallas... available for public inspection at 1445 Ross Avenue, Dallas, Texas 75202-2733. A copy of the proposed settlement may be obtained from Kevin Shade, 1445 Ross Avenue, Dallas, Texas 75202-2733 or by calling (214...

Color-Matching and Blending-Effect of Universal Shade Bulk-Fill-Resin-Composite in Resin-Composite-Models and Natural Teeth.

PubMed

Abdelraouf, Rasha M; Habib, Nour A

2016-01-01

Objectives . To assess visually color-matching and blending-effect (BE) of a universal shade bulk-fill-resin-composite placed in resin-composite-models with different shades and cavity sizes and in natural teeth (extracted and patients' teeth). Materials and Methods . Resin-composite-discs (10 mm × 1 mm) were prepared of universal shade composite and resin-composite of shades: A1, A2, A3, A3.5, and A4. Spectrophotometric-color-measurement was performed to calculate color-difference (Δ E ) between the universal shade and shaded-resin-composites discs and determine their translucency-parameter (TP). Visual assessment was performed by seven normal-color-vision-observers to determine the color-matching between the universal shade and each shade, under Illuminant D65. Color-matching visual scoring (VS) values were expressed numerically (1-5): 1: mismatch/totally unacceptable, 2: Poor-Match/hardly acceptable, 3: Good-Match/acceptable, 4: Close-Match/small-difference, and 5: Exact-Match/no-color-difference. Occlusal cavities of different sizes were prepared in teeth-like resin-composite-models with shades A1, A2, A3, A3.5, and A4. The cavities were filled by the universal shade composite. The same scale was used to score color-matching between the fillings and composite-models. BE was calculated as difference in mean-visual-scores in models and that of discs. Extracted teeth with two different class I-cavity sizes as well as ten patients' lower posterior molars with occlusal caries were prepared, filled by universal shade composite, and assessed similarly. Results . In models, the universal shade composite showed close matching in the different cavity sizes and surrounding shades (4 ≤ VS < 5) (BE = 0.6-2.9 in small cavities and 0.5-2.8 in large cavities). In extracted teeth, there was good-to-close color-matching (VS = 3.7-4.4 in small cavities, BE = 2.5-3.2) (VS = 3-3.5, BE = 1.8-2.3 in large cavities). In patients' molars, the universal shade composite showed good-matching (VS = 3-3.3, BE = -0.9-2.1). Conclusions . Color-matching of universal shade resin-composite was satisfactory rather than perfect in patients' teeth.

Is leaf dry matter content a better predictor of soil fertility than specific leaf area?

PubMed Central

Hodgson, J. G.; Montserrat-Martí, G.; Charles, M.; Jones, G.; Wilson, P.; Shipley, B.; Sharafi, M.; Cerabolini, B. E. L.; Cornelissen, J. H. C.; Band, S. R.; Bogard, A.; Castro-Díez, P.; Guerrero-Campo, J.; Palmer, C.; Pérez-Rontomé, M. C.; Carter, G.; Hynd, A.; Romo-Díez, A.; de Torres Espuny, L.; Royo Pla, F.

2011-01-01

Background and Aims Specific leaf area (SLA), a key element of the ‘worldwide leaf economics spectrum’, is the preferred ‘soft’ plant trait for assessing soil fertility. SLA is a function of leaf dry matter content (LDMC) and leaf thickness (LT). The first, LDMC, defines leaf construction costs and can be used instead of SLA. However, LT identifies shade at its lowest extreme and succulence at its highest, and is not related to soil fertility. Why then is SLA more frequently used as a predictor of soil fertility than LDMC? Methods SLA, LDMC and LT were measured and leaf density (LD) estimated for almost 2000 species, and the capacity of LD to predict LDMC was examined, as was the relative contribution of LDMC and LT to the expression of SLA. Subsequently, the relationships between SLA, LDMC and LT with respect to soil fertility and shade were described. Key Results Although LD is strongly related to LDMC, and LDMC and LT each contribute equally to the expression of SLA, the exact relationships differ between ecological groupings. LDMC predicts leaf nitrogen content and soil fertility but, because LT primarily varies with light intensity, SLA increases in response to both increased shade and increased fertility. Conclusions Gradients of soil fertility are frequently also gradients of biomass accumulation with reduced irradiance lower in the canopy. Therefore, SLA, which includes both fertility and shade components, may often discriminate better between communities or treatments than LDMC. However, LDMC should always be the preferred trait for assessing gradients of soil fertility uncoupled from shade. Nevertheless, because leaves multitask, individual leaf traits do not necessarily exhibit exact functional equivalence between species. In consequence, rather than using a single stand-alone predictor, multivariate analyses using several leaf traits is recommended. PMID:21948627

Light quality characterization under climate screens and shade nets for controlled-environment agriculture.

PubMed

Kotilainen, Titta; Robson, T Matthew; Hernández, Ricardo

2018-01-01

Climate screens are typically used inside glass greenhouses to improve control of humidity and temperature, and thus reduce energy expenditure. Shade nets are more appropriate to use, either with or without polyethylene cladding, at locations less-reliant on climate control, but where protection against hail, wind and excessive solar radiation might be needed. In addition, insect screens and nets can be employed to hinder insect pests and other invertebrates entering either type of production environment, and to keep invertebrates used in pest management contained inside. Screens and nets both transmit sunlight in a wavelength-specific manner, giving them the potential to affect plant morphology and physiology. Screens and nets of various colours and nominal shading factors have been described and studied; however, detailed measurements of their spectral characteristics are scarce. We measured solar spectral photon-irradiance and its attenuation by climate screens, shade nets, insect nets, greenhouse glass, and polyethylene covers. Our aim was to elucidate the effects of different patterns, colours, and shading factors, on light quality in production environments. Our measurements reveal that there are large differences both in the fraction of global irradiance attenuated and spectral ratios received under materials that are otherwise superficially similar in terms of their appearance and texture. We suggest that the type of spectral characterization that we performed is required to fully interpret the results of research examining plant responses to different types of screen and net. These data on spectral irradiance would benefit material manufacturers, researchers, growers, and horticultural consultants, enabling material selection to better match the solutions sought by growers and their desired outcomes regarding plant performance.

Is leaf dry matter content a better predictor of soil fertility than specific leaf area?

PubMed

Hodgson, J G; Montserrat-Martí, G; Charles, M; Jones, G; Wilson, P; Shipley, B; Sharafi, M; Cerabolini, B E L; Cornelissen, J H C; Band, S R; Bogard, A; Castro-Díez, P; Guerrero-Campo, J; Palmer, C; Pérez-Rontomé, M C; Carter, G; Hynd, A; Romo-Díez, A; de Torres Espuny, L; Royo Pla, F

2011-11-01

Specific leaf area (SLA), a key element of the 'worldwide leaf economics spectrum', is the preferred 'soft' plant trait for assessing soil fertility. SLA is a function of leaf dry matter content (LDMC) and leaf thickness (LT). The first, LDMC, defines leaf construction costs and can be used instead of SLA. However, LT identifies shade at its lowest extreme and succulence at its highest, and is not related to soil fertility. Why then is SLA more frequently used as a predictor of soil fertility than LDMC? SLA, LDMC and LT were measured and leaf density (LD) estimated for almost 2000 species, and the capacity of LD to predict LDMC was examined, as was the relative contribution of LDMC and LT to the expression of SLA. Subsequently, the relationships between SLA, LDMC and LT with respect to soil fertility and shade were described. Although LD is strongly related to LDMC, and LDMC and LT each contribute equally to the expression of SLA, the exact relationships differ between ecological groupings. LDMC predicts leaf nitrogen content and soil fertility but, because LT primarily varies with light intensity, SLA increases in response to both increased shade and increased fertility. Gradients of soil fertility are frequently also gradients of biomass accumulation with reduced irradiance lower in the canopy. Therefore, SLA, which includes both fertility and shade components, may often discriminate better between communities or treatments than LDMC. However, LDMC should always be the preferred trait for assessing gradients of soil fertility uncoupled from shade. Nevertheless, because leaves multitask, individual leaf traits do not necessarily exhibit exact functional equivalence between species. In consequence, rather than using a single stand-alone predictor, multivariate analyses using several leaf traits is recommended.

Shared and divergent pathways for flower abscission are triggered by gibberellic acid and carbon starvation in seedless Vitis vinifera L.

PubMed

Domingos, Sara; Fino, Joana; Cardoso, Vânia; Sánchez, Claudia; Ramalho, José C; Larcher, Roberto; Paulo, Octávio S; Oliveira, Cristina M; Goulao, Luis F

2016-02-01

Abscission is a highly coordinated developmental process by which plants control vegetative and reproductive organs load. Aiming at get new insights on flower abscission regulation, changes in the global transcriptome, metabolome and physiology were analyzed in 'Thompson Seedless' grapevine (Vitis vinifera L.) inflorescences, using gibberellic acid (GAc) spraying and shading as abscission stimuli, applied at bloom. Natural flower drop rates increased from 63.1% in non-treated vines to 83% and 99% in response to GAc and shade treatments, respectively. Both treatments had a broad effect on inflorescences metabolism. Specific impacts from shade included photosynthesis inhibition, associated nutritional stress, carbon/nitrogen imbalance and cell division repression, whereas GAc spraying induced energetic metabolism simultaneously with induction of nucleotide biosynthesis and carbon metabolism, therefore, disclosing alternative mechanisms to regulate abscission. Regarding secondary metabolism, changes in flavonoid metabolism were the most represented metabolic pathways in the samples collected following GAc treatment while phenylpropanoid and stilbenoid related pathways were predominantly affected in the inflorescences by the shade treatment. However, both GAc and shade treated inflorescences revealed also shared pathways, that involved the regulation of putrescine catabolism, the repression of gibberellin biosynthesis, the induction of auxin biosynthesis and the activation of ethylene signaling pathways and antioxidant mechanisms, although often the quantitative changes occurred on specific transcripts and metabolites of the pathways. Globally, the results suggest that chemical and environmental cues induced contrasting effects on inflorescence metabolism, triggering flower abscission by different mechanisms and pinpointing the participation of novel abscission regulators. Grapevine showed to be considered a valid model to study molecular pathways of flower abscission competence acquisition, noticeably responding to independent stimuli.

Avian response to shade‐layer restoration in coffee plantations in Puerto Rico

USGS Publications Warehouse

Irizarry, Amarilys D.; Collazo, Jaime A.; Pacifici, Krishna; Reich, Brian J.; Battle, Kathryn E.

2018-01-01

Documenting the evolving processes associated with habitat restoration and how long it takes to detect avian demographic responses is crucial to evaluate the success of restoration initiatives and to identify ways to improve their effectiveness. The importance of this endeavor prompted the U.S. Fish and Wildlife Service and the USDA Natural Resources Conservation Service to evaluate their sun‐to‐shade coffee restoration program in Puerto Rico initiated in 2003. We quantified the responses of 12 resident avian species using estimates of local occupancy and extinction probabilities based on surveys conducted in 2015–2017 at 65 restored farms grouped according to time‐since‐initial‐restoration (TSIR): new (2011–2014), intermediate (2007–2010), and old (2003–2006). We also surveyed 40 forest sites, which served as reference sites. Vegetation complexity increased with TSIR, ranging between 35 and 40% forest cover in farms 6–9 years TSIR. Forest specialists (e.g. Loxigilla portoricencis) exhibited highest average occupancy in farms initially classified as intermediate (6–9 years) and old (>10 years), paralleling occupancy in secondary forests. Occupancy of open‐habitat specialists (e.g. Tiaris olivaceus) was more variable, but higher in recently restored farms. Restoring the shade layer has the potential to heighten ecological services derived from forest specialists (e.g. frugivores) without losing the services of many open‐habitat specialists (e.g. insectivores). Annual local extinction probability for forest specialists decreased with increasing habitat complexity, strengthening the potential value of shade restoration as a tool to enhance habitat for avifauna that evolved in forested landscapes.

Predictors of sun protection behaviours and sunburn among Australian adolescents.

PubMed

Pettigrew, Simone; Jongenelis, Michelle; Strickland, Mark; Minto, Carolyn; Slevin, Terry; Jalleh, Geoffrey; Lin, Chad

2016-07-13

Excessive sun exposure and sunburn increase individuals' risk of skin cancer. It is especially important to prevent sunburn in childhood due to the higher relative risk of skin cancer across the life span compared to risk associated with sunburn episodes experienced later in life. This study examined demographic and attitudinal factors associated with engagement in a range of sun protection behaviours (wearing a hat, wearing protective clothing, staying in the shade, and staying indoors during the middle of the day) and the frequency of sunburn among Western Australian adolescents to provide insights of relevance for future sun protection campaigns. Cross-sectional telephone surveys were conducted annually with Western Australians between 2005/06 and 2014/15. The results from 4150 adolescents aged 14-17 years were used to conduct a path analysis of factors predicting various sun protection behaviours and sunburn. Significant primary predictors of the sun protection behaviours included in the study were skin type (sun sensitivity), gender, tanning-related attitudes and behaviours, and perceived relevance of public service advertisements that advocate sun protection. Of the four sun protection behaviours investigated, staying in the shade and staying indoors during the middle of the day were associated with a lower frequency of sunburn. There is a particular need to target sun protection messages at adolescent males who are less likely to engage in the most effective sun protection behaviours and demonstrate an increased propensity to experience sunburn. The results suggest that such future sun protection messages should include a focus on the importance of staying in the shade or indoors during periods of high UV radiation to increase awareness of the efficacy of these methods of avoiding skin cancer.

Herbivores modify selection on plant functional traits in a temperate rainforest understory.

PubMed

Salgado-Luarte, Cristian; Gianoli, Ernesto

2012-08-01

There is limited evidence regarding the adaptive value of plant functional traits in contrasting light environments. It has been suggested that changes in these traits in response to light availability can increase herbivore susceptibility. We tested the adaptive value of plant functional traits linked with carbon gain in contrasting light environments and also evaluated whether herbivores can modify selection on these traits in each light environment. In a temperate rainforest, we examined phenotypic selection on functional traits in seedlings of the pioneer tree Aristotelia chilensis growing in sun (canopy gap) and shade (forest understory) and subjected to either natural herbivory or herbivore exclusion. We found differential selection on functional traits depending on light environment. In sun, there was positive directional selection on photosynthetic rate and relative growth rate (RGR), indicating that selection favors competitive ability in a high-resource environment. Seedlings with high specific leaf area (SLA) and intermediate RGR were selected in shade, suggesting that light capture and conservative resource use are favored in the understory. Herbivores reduced the strength of positive directional selection acting on SLA in shade. We provide the first demonstration that natural herbivory rates can change the strength of selection on plant ecophysiological traits, that is, attributes whose main function is resource uptake. Research addressing the evolution of shade tolerance should incorporate the selective role of herbivores.

[Effects of light intensities after anthesis on the photosynthetic characteristics and chloroplast ultrastructure in mesophyll cell of summer maize (Zea mays L. )].

PubMed

Gao, Jia; Cui, Hai Yan; Shi, Jian Guo; Dong, Shu Ting; Liu, Peng; Zhao, Bin; Zhang, Ji Wang

2018-03-01

We examined the changes of photosynthetic characteristics and chloroplast ultrastructure in mesophyll cell of summer maize in response to different light intensities in the field, with the summer maize hybrid Denghai 605 as experimental material. Two treatments of both shading (S) and increasing light (L) from flowering to physiological maturity stage were designed, with the ambient sunlight treatment as control (CK). Under shading treatment, poorly developed thylakoid structure, blurry lamellar structure, loose granum, large gap between slices and warping granum were the major characteristics in chloroplast. Meanwhile, photosynthetic rate (P n ), transpiration rate, stomatal conductance, chlorophyll content, and actual photo-chemical efficiency (Φ PSII ) decreased, whereas the maximal photochemical efficiency and non-photochemical quenching increased, which resulted in decreases in grain yield under shading treatment. However, a better development was observed in chloroplasts for L treatment, with the number of grana and lamellae increased and lamellae arranged compactly. In addition, P n and Φ PSII increased under L treatment, which increased grain yield. The chloroplast arrangement dispersed in mesophyll cells and chloroplast ultrastructure was destroyed after shading, and then chlorophyll synthesis per unit leaf area and photosynthetic capacity decreased. In contrast, the number of grana and lamellae increased and lamellae arranged compactly after increasing light, which are beneficial for corn yield.

No time for candy: passionfruit (Passiflora edulis) plants down-regulate damage-induced extra floral nectar production in response to light signals of competition.

PubMed

Izaguirre, Miriam M; Mazza, Carlos A; Astigueta, María S; Ciarla, Ana M; Ballaré, Carlos L

2013-09-01

Plant fitness is often defined by the combined effects of herbivory and competition, and plants must strike a delicate balance between their ability to capture limiting resources and defend against herbivore attack. Many plants use indirect defenses, such as volatile compounds and extra floral nectaries (EFN), to attract canopy arthropods that are natural enemies of herbivorous organisms. While recent evidence suggests that upon perception of low red to far-red (R:FR) ratios, which signal the proximity of competitors, plants down-regulate resource allocation to direct chemical defenses, it is unknown if a similar phytochrome-mediated response occurs for indirect defenses. We evaluated the interactive effects of R:FR ratio and simulated herbivory on nectar production by EFNs of passion fruit (Passiflora edulis f. flavicarpa). The activity of petiolar EFNs dramatically increased in response to simulated herbivory and hormonal treatment with methyl jasmonate (MeJA). Low R:FR ratios, which induced a classic "shade-avoidance" repertoire of increased stem elongation in P. edulis, strongly suppressed the EFN response triggered by simulated herbivory or MeJA application. Strikingly, the EFN response to wounding and light quality was localized to the branches that received the treatments. In vines like P. edulis, a local response would allow the plants to precisely adjust their light harvesting and defense phenotypes to the local conditions encountered by individual branches when foraging for resources in patchy canopies. Consistent with the emerging paradigm that phytochrome regulation of jasmonate signaling is a central modulator of adaptive phenotypic plasticity, our results demonstrate that light quality is a strong regulator of indirect defenses.

MYBs affect the variation in the ratio of anthocyanin and flavanol in fruit peel and flesh in response to shade.

PubMed

Lu, Yanfen; Bu, Yufen; Hao, Suxiao; Wang, Yaru; Zhang, Jie; Tian, Ji; Yao, Yuncong

2017-03-01

Fruit pigment accumulation, which represents an important indicator of nutrient quality and appearance value, is often affected by low light under rain, cloud, fog and haze conditions during the veraison period. It is not known whether continuous low light interferes with the production and accumulation of secondary metabolites in veraison fruit. In this paper, we measured pigments and the transcriptional level of genes related to secondary metabolites, i.e., flavonoid biosynthesis in the peel and flesh of Malus crabapple 'Radiant' fruit in response to normal light and shade from 10th July to 30th August. The results showed crosstalk between the flavonoid biosynthetic genes and the involvement of key transcription factors such as McMYB4, McMYB7, McMYB10, and McMYB16 in the regulation of the ratio of anthocyanins and flavanols, which accounted for the different colouration of the fruit peel and flesh under shade conditions. A model is proposed for the regulation of the flavonoid pathway in the peel and flesh of 'Radiant' fruit based on our study results. Moreover, the molecular mechanism for 'Radiant' fruit colouration provides reference information for understanding the light regulatory mechanism involved in the biosynthesis of flavonoids and for designing the next generation of apple breeding. Copyright © 2017 Elsevier B.V. All rights reserved.

A statistical light use efficiency model explains 85% variations in global GPP

NASA Astrophysics Data System (ADS)

Jiang, C.; Ryu, Y.

2016-12-01

Photosynthesis is a complicated process whose modeling requires different levels of assumptions, simplification, and parameterization. Among models, light use efficiency (LUE) model is highly compact but powerful in monitoring gross primary production (GPP) from satellite data. Most of LUE models adopt a multiplicative from of maximum LUE, absorbed photosynthetically active radiation (APAR), and temperature and water stress functions. However, maximum LUE is a fitting parameter with large spatial variations, but most studies only use several biome dependent constants. In addition, stress functions are empirical and arbitrary in literatures. Moreover, meteorological data used are usually coarse-resolution, e.g., 1°, which could cause large errors. Finally, sunlit and shade canopy have completely different light responses but little considered. Targeting these issues, we derived a new statistical LUE model from a process-based and satellite-driven model, the Breathing Earth System Simulator (BESS). We have already derived a set of global radiation (5-km resolution), carbon and water fluxes (1-km resolution) products from 2000 to 2015 from BESS. By exploring these datasets, we found strong correlation between APAR and GPP for sunlit (R2=0.84) and shade (R2=0.96) canopy, respectively. A simple model, only driven by sunlit and shade APAR, was thus built based on linear relationships. The slopes of the linear function act as effective LUE of global ecosystem, with values of 0.0232 and 0.0128 umol C/umol quanta for sunlit and shade canopy, respectively. When compared with MPI-BGC GPP products, a global proxy of FLUXNET data, BESS-LUE achieved an overall accuracy of R2 = 0.85, whereas original BESS was R2 = 0.83 and MODIS GPP product was R2 = 0.76. We investigated spatiotemporal variations of the effective LUE. Spatially, the ratio of sunlit to shade values ranged from 0.1 (wet tropic) to 4.5 (dry inland). By using maps of sunlit and shade effective LUE the accuracy of BESS-LUE further reached R2 = 0.88. Temporally, both sunlit and shade effective LUE had seasonal peak values in NH summer, and both showed significant increasing trends. Overall, BESS-LUE exhibited promising potential in global GPP mapping. We are going to evaluate it using FLUXNET2015 database and satellite solar Induced Fluorescence (SIF) data.

Does gender and experience influence shade matching quality?

PubMed

Haddad, Helene J; Jakstat, Holger A; Arnetzl, Gerwin; Borbely, Judit; Vichi, Alessandro; Dumfahrt, Herbert; Renault, Patrick; Corcodel, Nicoleta; Pohlen, Bostjan; Marada, Gyula; de Parga, Juan A Martinez Vazquez; Reshad, Mamaly; Klinke, Thomas U; Hannak, Wolfgang B; Paravina, Rade D

2009-01-01

To evaluate the influence of gender and level of experience on shade matching quality. A study was simultaneously performed at 15 universities located in 9 countries. A total of 614 color normal participants completed all phases of the experiment. Among them, there were 305 females and 309 males, 319 dental students and 295 dental professionals. A lecture on color matching in dentistry was given to all participants. Initial training was performed using Toothguide Trainer software (TT), while Toothguide Training Box (TTB) was used for both training and testing of participants' shade matching results. The test task was to successively match 15 shade guide tabs with the corresponding shade guide. The shade matching score for each participant was computed as a sum of color differences (SigmaDeltaE(ab)(*) score) between target tabs and selected tabs. Lower scores corresponded to better shade matching results and vice versa. Means and standard deviations were calculated. Mann-Whitney U test was used for statistical analysis of the data (alpha=0.05). The mean shade matching score (S.D.) for all participants was 41 (21). The score for female and male participants was 38 (20) and 44 (21), respectfully (p<0.001). The difference in scores between dental students, 42 (20), and dental professionals, 39 (21), was not statistically significant. Within the limitations of this study, females achieved significantly better shade matching results than males, indicating that gender plays an important role in shade matching. The level of experience was not found to be significant factor in shade matching.

Assessment of the influence of gender and skin color on the preference of tooth shade in Saudi population.

PubMed

Labban, Nawaf; Al-Otaibi, Hanan; Alayed, Abdulaziz; Alshankiti, Khaled; Al-Enizy, Mohammad A

2017-07-01

The aim of the study was to evaluate the influence of gender and skin color on the preference of different teeth shades in general population. Two standardized frontal smile photographs of male and female subjects were manipulated using photoshop to represent 4 skin colors [(type II, III, IV, and V) (Fitzpatrick scale)]. The teeth shades under each skin color were digitally manipulated to represent one of 6 teeth shades (BL1, BL2, BL3, BL4, B1 and A1). A questionnaire assessed demographic characteristics (age, nationality, gender, education level, occupation, and income) along with the satisfaction of their smiles. Male and female set of pictures with combination of skin colors and teeth shades were presented and participants were asked to select the most esthetically pleasing teeth shade with regard to gender and skin color. Cross-tabulations and chi-squared tests were used to perform the statistical analyses ( α  = 0.05). Three hundred and thirty-six (60.4% male; 39.6% female) individuals participated in the study. The difference in the preferred teeth shades was significant among the male and female photographs across all skin colors ( p  < 0.05). Lighter teeth shades were preferred among female subjects compared to male subjects with the same skin color. In addition, lighter teeth shades were preferred among subjects with a lighter skin color and vice versa ( p  < 0.05). Gender and skin color influences the perception of teeth shades among general population. Therefore, lighter tooth shades (BL1, BL2) for lighter skin color and comparatively darker tooth shades (BL4, B1, A1) for darker skin individuals should be prescribed as these are perceived as natural among Saudi population.

[Effect of ceramic thickness and resin cement shades on final color of heat-pressed ceramic veneers].

PubMed

Ren, D F; Zhan, K R; Chen, X D; Xing, W Z

2017-02-09

Objective: To analyze the effect of ceramic materials thickness and resin cement shades on the final color of ceramic veneers in the discolored teeth, and to investigate the color agreement of try-in pastes to the corresponding resin cements. Methods: Sixty artificial maxillary central incisor teeth (C2 shade) were used to simulate the natural discolored teeth and prepared according to veneer tooth preparation protocol. Veneers of different thickness in the body region (0.50 and 0.75 mm) were fabricated using ceramic materials (LT A2 shade, IPS e.max Press). The ceramic veneer specimens were bonded to the artificial teeth using the 6 shades of resin cements (Variolink Veneer: shades of LV-3, LV-2, HV+3; RelyX™ Veneer: shades of TR, A3, WO) ( n= 5). A clinical spectrophotometer was used to measure the color parameters of ceramic veneers at the cervical, body and incisal regions. Color changes of veneers before and after cementation were calculated and registered as ΔE1, and the changes between try-in paste and the corresponding resin cements were registered as ΔE2. Results: Three-way ANOVA indicated that ΔE1 and ΔE2 values were significantly affected by the ceramic thickness, resin cement shades and measuring regions ( P< 0.05). The ΔE1 values of six shades ranged from 0.59-8.27. The ΔE1 values were more than 2.72 when the ceramic veneers were cemented with resin cements in shades of HV+3 and WO. The ΔE2 values of six shades ranged from 0.60-2.56. The shades of HV+3, WO and A3 resin cements were more than 1.60. Conclusions: Different thickness of ceramic materials, resin cement shades and measuring regions could affect the final color of ceramic veneers. The color differences of some resin cements and corresponding try-in pastes might be observed in clinical practice.

Lightness, chroma and hue differences on visual shade matching.

PubMed

Pecho, Oscar E; Pérez, María M; Ghinea, Razvan; Della Bona, Alvaro

2016-11-01

To analyze the influence of lightness, chroma and hue differences on visual shade matching performed by dental students. 100 dental students (DS) volunteers with normal vision participated in the study. A spectroradiometer (SP) was used to measure the spectral reflectance of 4 extracted human upper central incisors (UCI 1-4) and shade tabs from Vita Classical (VC) and Vita Toothguide 3D-Master (3D) shade guides. Measurements were performed over a gray background, inside a viewing booth and under D65 illuminant (diffuse/0° geometry). Color parameters (L*, a*, b*, C* and h°) were calculated. DS used VC and 3D to visually select the best shade match for each UCI. CIE metric differences (Δa * ,Δb * ,ΔL ' , ΔC ' and ΔH ' ) and CIEDE2000(2:1:1) lightness (ΔE L ), chroma (ΔE C ) and hue (ΔE H ) differences were obtained from each UCI and the first three shades selected by DS and the first option using CIELAB, CIEDE2000(1:1:1) and CIEDE2000(2:1:1) color difference metrics. The closest CIELAB color-discrimination ellipsoid (from RIT-DuPont visual color-difference data) to each UCI was selected for the analysis of visual shade matching. DS showed a preference for shades with lower chroma (ΔC ' and ΔE C ) and/or hue (ΔH ' and ΔE H ) values instead of shades with lower lightness values (ΔL ' and ΔE L ). Most best visual matches were near the tolerance ellipsoid centered on tooth shade. This study is an attempt to partially explain the inconsistencies between visual and instrumental shade matching and the limitations of shade guides. Visual shade matching was driven by color differences with lower chroma and hue values. Copyright © 2016 The Academy of Dental Materials. Published by Elsevier Ltd. All rights reserved.

Influence of Different Enamel Shades and Thickness on Chroma and Value of Dentin Vita Shade: An in vitro Comparative Assessment Study.

PubMed

Hajira, Noor Saira Wajid Najma; Mehta, Deepak; Ashwini, P; Meena, N; Usha, H L

2015-04-01

The aim of the present study is to determine the influence of different enamel shades of various thickness on chroma and value of vita shade of dentin. Three enamel composite resin shades (Enamel white, grey and neutral) and one dentin shade (A 2) from A melogen Plus (Ultradent) was used. Ninety Enamel disk specimens of 0.5, 0.75 and 1 mm thickness and 10 mm in diameter for each shade and 90 dentin disk specimens of 2 mm in thickness and 10 mm in diameter was used for the study. The spectrophotometric values of the dentin shade with and without enamel specimens were recorded and the values were converted to CIEL*a*b values. Statistical analysis was done using Pearson correlation coefficients to verify the effect of thickness on Chroma and value, and the significance was evaluated by one-way ANOVA and Tukey post hoc test. Two way ANOVA and Tukey post hoc was done to verify the variation within the groups. Results revealed a significant positive correlation between thickness and chroma and a negative correlation between thickness and value. There was a statistically significant variation in between the groups. All groups produced a significant increase in chroma with increase in thickness of enamel shade upto a thickness of 0.75 mm after which the behavior of each shade was erratic. Hence, the optimum thickness would be 0.75 mm. All groups produced a significant decrease in value with increase in thickness of enamel shade. Enamel white produced the greatest reduction in value, enamel neutral the least and enamel grey demonstrated an intermediate result. There is a need to have a knowledge of the effect on chroma and value when dentin is layered with different enamel shades, it is also important to understand the effect of these enamel shades at different thicknesses to better control the color and reproduce esthetic simulating natural teeth.

[Studies on the shade-endurance capacity of Glycyrrhiza uralensis].

PubMed

Wei, Sheng-li; Wang, Wen-quan; Chen, Xiu-hua; Qin, Shu-ying; Chen, Xiu-tian

2005-01-01

To study the shade-endurance property of Glycyrrhiza uralensis and provide rationale for the practice of inter-cropping G. uralensis with trees. Black shading nets were used to provide five different environments of light intensities (light penetration rates of 100%, 75%, 65%, 50% and 25%, respectively). To assess the shade-endurance capacity of G. uralensis, several aspects were evaluated, including growth characters, physiological and ecological characters, biomass, and chemical contents. G. uralensis is a light-favored plant. The growth indices such as plant height, stem diameter, leaves number, root diameter, biomass, and daily average photosynthetic rate (Pn) are highest when light permeation rate is 100%. All these indices decrease when light intensity decreases. However, G. uralensis possesses shade-endurance capacity to some degree; it adapts to the shading environment by increasing the leaf area and chlorophyll contents. Shading has no obvious effect on the absolute light energy utilization rate (Eu) or Fv/Fm ratio. The influence of shading on the chemical contents of G. uralensis is obvious.

[Study on the matching of the shade between beverage and modifying porcelain shade guide].

PubMed

Yao, Jiang-wu; Li, Shui-gen; Lin, Jin-ying

2007-10-01

To evaluate the matching of the shade between beverage and modifying porcelain shade guide according to Munsell color order system, thus to provide the reference basis for selecting modifying porcelain to mimic the stain of natural tooth by technician. The shade of Vita Akzent, Vita Interno, Shofu Vintage & Unibond and Noritake Super Porcelain EX-3 shade tabs as well as 15 kinds of beverage were measured according to Munsell color order system on Color-Eye 7000A spectrophotometer. The difference of the frequency of approximate hue, value, and chroma between shade tabs and beverage were compared by calculating the Fisher exact probabilities. The frequency of approximate hue between 4 kinds of shade tabs and 15 kinds of bev-erage was significant different (P<0.05), while the frequency of approximate value, and chroma was not significant different (P>0.05). Except the hue, the color distribution of 4 kinds of shade tabs was similar to that of 15 kinds of beverage. But the color of beverage also can be approximately matched by any kind of modifying porcelain by mixing porcelain powder of appropriate hue, value, and chroma.

External shading devices for energy efficient building

NASA Astrophysics Data System (ADS)

Shahdan, M. S.; Ahmad, S. S.; Hussin, M. A.

2018-02-01

External shading devices on a building façade is an important passive design strategy as they reduce solar radiation. Although studies have proven the benefits of external shading devices, many are designed solely for aesthetic purposes without fully considering its high potential to reduce solar radiation and glare. Furthermore, explorations into shading devices by the design team are mostly left too late in the design development phases. Hence, the paper looks into the effectiveness of external shading devices on a building towards more energy efficient building. The study aims to analyse the effects of various configurations of external shading devices towards the energy consumption of a case study building based on computer simulations. This study uses Building Information Modelling (BIM) through Autodesk Revit software as simulation tool. The constant variables for the simulation are the orientation of the building, types of glazing used by the building and the internal loads of the building. Whereas, the manipulated variable is the types of shading device used. The data were sorted according to the categories and translated into a chart. Analysis of the findings indicate that shading devices with different configurations show significant results in the energy consumption and the best configuration is the egg-crate shading devices. The study recommends that the consideration for shading device as a passive design strategy needs to be developed at the early stage of the building design.

Heterotrophic respiration in drained tropical peat temperatures influenced by shading gradient

NASA Astrophysics Data System (ADS)

Jauhiainen, Jyrki; Kerojoki, Otto; Silvennoinen, Hanna; Limin, Suwido; Vasander, Harri

2015-04-01

Lowland peatlands in Southeast Asia constitute a highly concentrated carbon (C) pool of global significance. These peatlands have formed over periods of several millennia by forest vegetation tolerant to flooding and poor substrates. Uncontrollable drainage and reoccurring wild fires in lack of management after removal of forest cover has impaired the C-storing functions in large reclaimed areas. Intergovernmental Panel on Climate Change (IPCC) reporting sees drained tropical organic soils as one of the largest greenhouse gas emissions releasing terrestrial systems. Vast areas of deforested tropical peatlands do not receive noteworthy shading by vegetation, which increases the amount of solar radiation reaching the peat surface. We studied heterotrophic carbon dioxide (CO2), nitrous oxide (N2O) and methane (CH4) fluxes in tropical peat in conditions, where; (i) peat temperatures were modified by artificial shading (no shade, 28%, 51% and 90% from the full sun exposure), (ii) root respiration was minimized, (iii) nutrient availability for peat decomposer community was changed (NPK fertilization of 0 and 313 kg ha-1). The experiment was repeated at two over 20 years ago drained fallow agricultural- and degraded sites in Central Kalimantan, Indonesia. Enhanced shading created a lasting decrease in peat temperatures, and decreased diurnal temperature fluctuations, in comparison to less shaded plots. The largest peat temperature difference was between the unshaded and 90% shaded peat surface, where the average temperatures within the topmost 50-cm peat profile differed 3 °C, and diurnal temperatures at 5 cm depth varied up to 4.2 °C in the unshaded and 0.4 °C in the 90% shaded conditions. Highest impacts on the heterotrophic CO2 fluxes caused by the treatments were on agricultural land, where 90% shading from the full exposure resulted in a 33% lower CO2 emission average on the unfertilised plots and a 66% lower emission average on the fertilised plots. Correlation between peat temperature and CO2 flux suggested an approximately 8% (unfertilised) and 25% (fertilised) emissions change for each 1 °C temperature change at 5 cm depth on the agricultural land. CO2 flux responses to the treatments remained low or were inconsistent over the peat temperature range.. Fertilised conditions negatively correlated with N2O efflux with increases in temperature, suggesting a 12-36% lower efflux for each 1 °C increase in peat temperature (at 5 cm depth) at the sites. Despite the apparently similar landscapes of fallow agricultural land and degraded peatland sites, the differences in greenhouse gas dynamics are expected to be an outcome of the long-term management differences. Based on the results it is possible to seek management practices that prolong timespan for using drained tropical peat for cultivation, simultaneously reduce negative climate impacts created from peat substrate carbon loss, and also improve greenhouse gas monitoring techniques at field.

Relationship between natural tooth shade and skin colour.

PubMed

Nourbakhsh, M; Mousavinejad, N; Adli, A R; Harati, M

2013-06-01

The purpose of this study was to assess the correlation of skin colour and tooth shade. One hundred and twenty six individuals aging between 18 to 25 years participated in this study. Colour of the maxillary central incisors was examined by VITA easy shade. Tooth shades were assigned to four ordinal values. Nivea Beauty Protect Foundation shade sample was used as a guide to assess facial skin colour Shin colours were also assigned to four ordinal values. Spearman test revealed that there was a significant relationship between tooth shade and skin colour Total co-relation factor was 51.6% (p <0 .01). Co-relation factors were 57% for women and 27% for men (p <0 .01). The highest tooth shade prevalence belonged to the second group and the highest skin colour prevalence was also in the second skin colour group.

Using Google Earth to Assess Shade for Sun Protection in Urban Recreation Spaces: Methods and Results.

PubMed

Gage, R; Wilson, N; Signal, L; Barr, M; Mackay, C; Reeder, A; Thomson, G

2018-05-16

Shade in public spaces can lower the risk of and sun burning and skin cancer. However, existing methods of auditing shade require travel between sites, and sunny weather conditions. This study aimed to evaluate the feasibility of free computer software-Google Earth-for assessing shade in urban open spaces. A shade projection method was developed that uses Google Earth street view and aerial images to estimate shade at solar noon on the summer solstice, irrespective of the date of image capture. Three researchers used the method to separately estimate shade cover over pre-defined activity areas in a sample of 45 New Zealand urban open spaces, including 24 playgrounds, 12 beaches and 9 outdoor pools. Outcome measures included method accuracy (assessed by comparison with a subsample of field observations of 10 of the settings) and inter-rater reliability. Of the 164 activity areas identified in the 45 settings, most (83%) had no shade cover. The method identified most activity areas in playgrounds (85%) and beaches (93%) and was accurate for assessing shade over these areas (predictive values of 100%). Only 8% of activity areas at outdoor pools were identified, due to a lack of street view images. Reliability for shade cover estimates was excellent (intraclass correlation coefficient of 0.97, 95% CI 0.97-0.98). Google Earth appears to be a reasonably accurate and reliable and shade audit tool for playgrounds and beaches. The findings are relevant for programmes focused on supporting the development of healthy urban open spaces.

Reliability of conventional shade guides in teeth color determination.

PubMed

Todorović, Ana; Todorović, Aleksandar; Gostović, Aleksandra Spadijer; Lazić, Vojkan; Milicić, Biljana; Djurisić, Slobodan

2013-10-01

Color matching in prosthodontic therapy is a very important task because it influences the esthetic value of dental restorations. Visual shade matching represents the most frequently applied method in clinical practice. Instrumental measurements provide objective and quantified data in color assessment of natural teeth and restorations. In instrumental shade analysis, the goal is to achieve the smallest deltaE value possible, indicating the most accurate shade match. The aim of this study was to evaluate the reliability of commercially available ceramic shade guides. VITA Easyshade spectrophotometer (VITA, Germany) was used for instrumental color determination. Utilizing this device, color samples of ten VITA Classical and ten VITA 3D - Master shade guides were analyzed. Each color sample from all shade guides was measured three times and the basic parameters of color quality were examined: deltaL, deltaC, deltaH, deltaE, deltaElc. Based on these parameters spectrophotometer marks the shade matching as good, fair or adjust. After performing 1,248 measurements of ceramic color samples, frequency of evaluations adjust, fair and good were statistically significantly different between VITA Classical and VITA 3D Master shade guides (p = 0.002). There were 27.1% cases scored as adjust, 66.3% as fair and 6.7% as good. In VITA 3D - Master shade guides 30.9% cases were evaluated as adjust, 66.4% as fair and 2.7% cases as good. Color samples from different shade guides, produced by the same manufacturer, show variability in basic color parameters, which once again proves the lack of precision and nonuniformity of the conventional method.

Measured and estimated performance of a fleet of shaded photovoltaic systems with string and module-level inverters

DOE PAGES

MacAlpine, Sara; Deline, Chris; Dobos, Aron

2017-03-16

Shade obstructions can significantly impact the performance of photovoltaic (PV) systems. Although there are many models for partially shaded PV arrays, there is a lack of information available regarding their accuracy and uncertainty when compared with actual field performance. This work assesses the recorded performance of 46 residential PV systems, equipped with either string-level or module-level inverters, under a variety of shading conditions. We compare their energy production data to annual PV performance predictions, with a focus on the practical models developed here for National Renewable Energy Laboratory's system advisor model software. This includes assessment of shade extent on eachmore » PV system by using traditional onsite surveys and newer 3D obstruction modelling. The electrical impact of shade is modelled by either a nonlinear performance model or assumption of linear impact with shade extent, depending on the inverter type. When applied to the fleet of residential PV systems, performance is predicted with median annual bias errors of 2.5% or less, for systems with up to 20% estimated shading loss. The partial shade models are not found to add appreciable uncertainty to annual predictions of energy production for this fleet of systems but do introduce a monthly root-mean-square error of approximately 4%-9% due to seasonal effects. Here the use of a detailed 3D model results in similar or improved accuracy over site survey methods, indicating that, with proper description of shade obstructions, modelling of partially shaded PV arrays can be done completely remotely, potentially saving time and cost.« less

Conservation in Brazil's Chocolate Forest: The Unlikely Persistence of the Traditional Cocoa Agroecosystem.

PubMed

JOHNS

1999-01-01

/ In southern Bahia, Brazil, the traditional cocoa agroecosystem with a dense shade canopy of native trees is now recognized as a secondary conservation route for highly endangered Atlantic Rainforest species. This "chocolate forest" of the densely shaded farms persists despite a massive 20-year Brazilian government modernization program in which shade was seen as a chief impediment to raising cocoa production. The objective of this study was to determine how this traditional agroecosystem endured. Although dense shade limits cocoa yield, it provides several agroecological benefits: control of insect pests and weeds, microclimate stability, and soil fertility maintenance. A keycomponent of modernization efforts was a shade-tree removal program designed to maximize cocoa production by using low shade and fertilizer while substituting agrochemicals for many beneficial roles of the overhead trees. This research found that many farmers rejected, or only partially accepted, the shade reduction process although it promised much higher cocoa yield and profit. Farmers employing a wide range of shading were interviewed, and it was found that decisions to remove or maintain the shade trees were linked to both agroecological and risk-minimization factors. Farmers' perceptions of the agroecological functions of the shade trees and individual willingness to entertain the economic risk associated with substituting agrochemicals for these were important. A less-profitable, but lower-risk approach of occasional fertilizer and agrochemical use with the traditional shade intact was a rational and widespread choice. Policies designed to maintain the traditional agroecosystem through the current economic crisis should heed the multiple functions of the overhead trees. KEY WORDS: Conservation; Brazil; Atlantic Rainforest; Cocoa; Agroecology; Risk; Agroforestry

A novel two-step method for screening shade tolerant mutant plants via dwarfism

USDA-ARS?s Scientific Manuscript database

When subjected to shade, plants undergo rapid shoot elongation, which often makes them more prone to disease and mechanical damage. It has been reported that, in turfgrass, induced dwarfism can enhance shade tolerance. Here, we describe a two-step procedure for isolating shade tolerant mutants of ...

Imaging system design for improved information capacity

NASA Technical Reports Server (NTRS)

Fales, C. L.; Huck, F. O.; Samms, R. W.

1984-01-01

Shannon's theory of information for communication channels is used to assess the performance of line-scan and sensor-array imaging systems and to optimize the design trade-offs involving sensitivity, spatial response, and sampling intervals. Formulations and computational evaluations account for spatial responses typical of line-scan and sensor-array mechanisms, lens diffraction and transmittance shading, defocus blur, and square and hexagonal sampling lattices.

Predicting Fire Susceptibility in the Forests of Amazonia

NASA Technical Reports Server (NTRS)

Nepstad, Daniel C.; Brown, I. Foster; Setzer, Alberto

2000-01-01

Although fire is the single greatest threat to the ecological integrity of Amazon forests, our ability to predict the occurrence of Amazon forest fires is rudimentary. Part of the difficulty encountered in making such predictions is the remarkable capacity of Amazon forests to tolerate drought by tapping moisture stored in deep soil. These forests can avoid drought-induced leaf shedding by withdrawing moisture to depths of 8 meters and more. Hence, the absorption of deep soil moisture allows these forests to maintain their leaf canopies following droughts of several months duration, thereby maintaining the deep shade and high relative humidity of the forest interior that prevents these ecosystems from burning. But the drought- and fire-avoidance that is conferred by this deep-rooting phenomenon is not unlimited. During successive years of drought, such as those provoked by El Nino episodes, deep soil moisture can be depleted, and drought-induced leaf shedding begins. The goal of this project was to incorporate this knowledge of Amazon forest fire ecology into a predictive model of forest flammability.

Retractable Sun Shade

NASA Technical Reports Server (NTRS)

Frank, A.; Derespinis, S. F.; Mockovciak, John, Jr.

1986-01-01

Window-shade type spring roller contains blanket, taken up by rotating cylindrical frame and held by frame over area to be shaded. Blanket made of tough, opaque polyimide material. Readily unfurled by mechanism to protect space it encloses from Sun. Blanket forms arched canopy over space and allows full access to it from below. When shading not needed, retracted mechanism stores blanket compactly. Developed for protecting sensitive Space Shuttle payloads from direct sunlight while cargo-bay doors open. Adapted to shading of greenhouses, swimming pools, and boats.

Thiamethoxam as a seed treatment alters the physiological response of maize (Zea mays) seedlings to neighbouring weeds.

PubMed

Afifi, Maha; Lee, Elizabeth; Lukens, Lewis; Swanton, Clarence

2015-04-01

Thiamethoxam is a broad-spectrum neonicotinoid insecticide that, when applied to seed, has been observed to enhance seedling vigour under environmental stress conditions. Stress created by the presence of neighbouring weeds is known to trigger the accumulation of hydrogen peroxide (H2 O2 ) in maize seedling tissue. No previous work has explored the effect of thiamethoxam as a seed treatment on the physiological response of maize seedlings emerging in the presence of neighbouring weeds. Thiamethoxam was found to enhance seedling vigour and to overcome the expression of typical shade avoidance characteristics in the presence of neighbouring weeds. These results were attributed to maintenance of the total phenolics content, 1,1-diphenyl-2-picryl-hydrazyl (DPPH) radical scavenging activity and anthocyanin and lignin contents. These findings were also associated with the activation of scavenging genes, which reduced the accumulation of H2 O2 and the subsequent damage caused by lipid peroxidation in maize seedlings originating from treated seeds even when exposed to neighbouring weeds. These results suggest the possibility of exploring new chemistries and modes of action as novel seed treatments to upregulate free radical scavenging genes and to maintain the antioxidant system within plants. Such an approach may provide an opportunity to enhance crop competitiveness with weeds. © 2014 Society of Chemical Industry.

Tree Size Inequality Reduces Forest Productivity: An Analysis Combining Inventory Data for Ten European Species and a Light Competition Model.

PubMed

Bourdier, Thomas; Cordonnier, Thomas; Kunstler, Georges; Piedallu, Christian; Lagarrigues, Guillaume; Courbaud, Benoit

2016-01-01

Plant structural diversity is usually considered as beneficial for ecosystem functioning. For instance, numerous studies have reported positive species diversity-productivity relationships in plant communities. However, other aspects of structural diversity such as individual size inequality have been far less investigated. In forests, tree size inequality impacts directly tree growth and asymmetric competition, but consequences on forest productivity are still indeterminate. In addition, the effect of tree size inequality on productivity is likely to vary with species shade-tolerance, a key ecological characteristic controlling asymmetric competition and light resource acquisition. Using plot data from the French National Geographic Agency, we studied the response of stand productivity to size inequality for ten forest species differing in shade tolerance. We fitted a basal area stand production model that included abiotic factors, stand density, stand development stage and a tree size inequality index. Then, using a forest dynamics model we explored whether mechanisms of light interception and light use efficiency could explain the tree size inequality effect observed for three of the ten species studied. Size inequality negatively affected basal area increment for seven out of the ten species investigated. However, this effect was not related to the shade tolerance of these species. According to the model simulations, the negative tree size inequality effect could result both from reduced total stand light interception and reduced light use efficiency. Our results demonstrate that negative relationships between size inequality and productivity may be the rule in tree populations. The lack of effect of shade tolerance indicates compensatory mechanisms between effect on light availability and response to light availability. Such a pattern deserves further investigations for mixed forests where complementarity effects between species are involved. When studying the effect of structural diversity on ecosystem productivity, tree size inequality is a major facet that should be taken into account.

Physiological and morphological responses of the temperate seagrass Zostera muelleri to multiple stressors: investigating the interactive effects of light and temperature.

PubMed

York, Paul H; Gruber, Renee K; Hill, Ross; Ralph, Peter J; Booth, David J; Macreadie, Peter I

2013-01-01

Understanding how multiple environmental stressors interact to affect seagrass health (measured as morphological and physiological responses) is important for responding to global declines in seagrass populations. We investigated the interactive effects of temperature stress (24, 27, 30 and 32°C) and shading stress (75, 50, 25 and 0% shade treatments) on the seagrass Zostera muelleri over a 3-month period in laboratory mesocosms. Z. muelleri is widely distributed throughout the temperate and tropical waters of south and east coasts of Australia, and is regarded as a regionally significant species. Optimal growth was observed at 27°C, whereas rapid loss of living shoots and leaf mass occurred at 32°C. We found no difference in the concentration of photosynthetic pigments among temperature treatments by the end of the experiment; however, up-regulation of photoprotective pigments was observed at 30°C. Greater levels of shade resulting in high photochemical efficiencies, while elevated irradiance suppressed effective quantum yield (ΔF/FM'). Chlorophyll fluorescence fast induction curves (FIC) revealed that the J step amplitude was significantly higher in the 0% shade treatment after 8 weeks, indicating a closure of PSII reaction centres, which likely contributed to the decline in ΔF/FM' and photoinhibition under higher irradiance. Effective quantum yield of PSII (ΔF/FM') declined steadily in 32°C treatments, indicating thermal damage. Higher temperatures (30°C) resulted in reduced above-ground biomass ratio and smaller leaves, while reduced light led to a reduction in leaf and shoot density, above-ground biomass ratio, shoot biomass and an increase in leaf senescence. Surprisingly, light and temperature had few interactive effects on seagrass health, even though these two stressors had strong effects on seagrass health when tested in isolation. In summary, these results demonstrate that populations of Z. muelleri in south-eastern Australia are sensitive to small chronic temperature increases and light decreases that are predicted under future climate change scenarios.

Influence of shading on ornamental and physiological characteristics during flower development of groundcover rose (Rosa hybrida L.)

NASA Astrophysics Data System (ADS)

Hou, Wei; Luo, Ya; Wang, Xiaorong; Chen, Qing; Sun, Bo; Wang, Yan; Liu, Zejing; Tang, Haoru; Zhang, Yong

2018-04-01

The objective of the present investigation was to study the effect of shading on flower quality during flower development and photosynthetic capacity of groundcover rose (Rosa hybrida L.). The results showed that shade significantly increased flower diameter, levels of soluble protein and soluble sugar, total carotenoids content and superoxide dismutase (SOD) activity, while contents of malondialdehyde (MDA) and total anthocyanins in shaded flowers were significantly decreased as compared to sun-exposed flowers. However, no significant changes were observed in petal color parameters L*, a*, b* and C* between sun exposure and shade treatment plants at each flower developmental stage. Therefore, groundcover rose seemed to have the capacity to shade condition through auto-regulation. These results could provide us with a theoretical basis for further application of groundcover rose in the greening of urban spaces and an understanding of the mechanisms behind the changes induced by shade.

An in vitro study to evaluate the difference in shade between commercially available shade guides and glazed porcelain.

PubMed

Manimaran, P; Sadan, D Sai

2016-10-01

Smile is one of the most important interactive communication skills of a person. A smile is the key factor for an aesthetic appearance. Hence aesthetics is one of the motivating factor for the patients to seek dental care. Correction of unaesthetic appearance gives a positive effect to the self esteem of the patient. The aim of this study was to compare the difference in the shade between the commercially available shade guides namely Vita Classical And Ivoclar Chromascop and the fired porcelain samples fabricated using Vita Zahnfabrik VMK 95 and Ivoclar Classic Materials respectively. The objective of this study was to obtain a matching brand of material that has a particular shade tab among the brands used. To conclude, Ivoclar material matched the chromascop shade guide better than the vita material matched the vita classic shade guide.

Shading reduces growth of longleaf and loblolly pine seedlings in containers

Treesearch

James P. Barnett

1989-01-01

Development of longleaf (Pinus palustris Mill.) and loblolly (P. taeda L.) pine seediings growing under three light conditions--full sunlight, 30% shade, and 50% shade--was evaluated. Although there was little difference between development of seedlings in 30% and 50% shade, those grown is full sunlight were signiticantly larger...

Mechanosensing of stem bending and its interspecific variability in five neotropical rainforest species.

PubMed

Coutand, Catherine; Chevolot, Malia; Lacointe, André; Rowe, Nick; Scotti, Ivan

2010-02-01

In rain forests, sapling survival is highly dependent on the regulation of trunk slenderness (height/diameter ratio): shade-intolerant species have to grow in height as fast as possible to reach the canopy but also have to withstand mechanical loadings (wind and their own weight) to avoid buckling. Recent studies suggest that mechanosensing is essential to control tree dimensions and stability-related morphogenesis. Differences in species slenderness have been observed among rainforest trees; the present study thus investigates whether species with different slenderness and growth habits exhibit differences in mechanosensitivity. Recent studies have led to a model of mechanosensing (sum-of-strains model) that predicts a quantitative relationship between the applied sum of longitudinal strains and the plant's responses in the case of a single bending. Saplings of five different neotropical species (Eperua falcata, E. grandiflora, Tachigali melinonii, Symphonia globulifera and Bauhinia guianensis) were subjected to a regimen of controlled mechanical loading phases (bending) alternating with still phases over a period of 2 months. Mechanical loading was controlled in terms of strains and the five species were subjected to the same range of sum of strains. The application of the sum-of-strain model led to a dose-response curve for each species. Dose-response curves were then compared between tested species. The model of mechanosensing (sum-of-strain model) applied in the case of multiple bending as long as the bending frequency was low. A comparison of dose-response curves for each species demonstrated differences in the stimulus threshold, suggesting two groups of responses among the species. Interestingly, the liana species B. guianensis exhibited a higher threshold than other Leguminosae species tested. This study provides a conceptual framework to study variability in plant mechanosensing and demonstrated interspecific variability in mechanosensing.

The effect of shade on chlorophyll and anthocyanin content of upland red rice

NASA Astrophysics Data System (ADS)

Muhidin; Syam'un, E.; Kaimuddin; Musa, Y.; Sadimantara, G. R.; Usman; Leomo, S.; Rakian, T. C.

2018-02-01

Upland red rice (Oryza sativa) is a staple food and contains anthocyanin, which can act as antioxidants, plays an important role both for the plant itself and for human health. Levels of antioxidants in rice can be affected by the availability of light. The results showed that the difference of shade, cultivar, and interaction both significantly affect the content of chlorophyll a, chlorophyll b and total chlorophyll. The results also showed that shade could increase chlorophyll in all cultivars tested. The highest levels of chlorophyll a were present in the moderate shade level (n2), then decreased at the shelter level (n3) and increased again at high levels (n4). While on chlorophyll content b, it appears that shade increased chlorophyll b in all cultivars tested and this increase was linear to the increase of shade. The shade treatment may increase the anthocyanin content and the increase depending on the type of cultivar. Increased levels of anthocyanin highest due to shade occurred on Jangkobembe cultivar. The original level of anthocyanin on Jangkobembe cultivar averaged 0.096 mg g-1 increased to 2.487 mg g-1 or increased 26 fold. It is concluded that the shade had a significant effect on the chlorophyll and anthocyanin content.

[Research on the designing method of a special shade guide for tooth whitening].

PubMed

Xu, Yingxin

2015-10-01

To investigate a method of designing an accurate and scientific shade guide, especially used for judging the effect of tooth whitening, by analyzing the colorimetric values of discolored teeth statistically. One hundred thirty-six pictures of patients who had been receiving the Beyond cold light whitening treatment from February 2009 to July 2014 were analyzed, including 25 tetracycline teeth, 61 mottled-enamel teeth, and 50 yellow teeth. The colorimetric values of discolored teeth were measured. The L* values of shade tabs were calculated by hierarchical clustering of those of discolored teeth. The a* and b* values of shade tabs were the mean of those observed for discolored teeth. Accordingly, different shade guides were designed for each type of discolored teeth, and the effects were evaluated. A statistically significant difference in colorimetric values was found among the three types of discolored teeth. Compared with the Vitapan Classical shade guide, the shade guides designed through the present method were more scientific and accurate in judging the effect of tooth whitening. Moreover, the arrangement of shade tabs was more logical, and the color difference between shade tabs and discolored teeth was smaller. The proposed designing method is theoretically feasible, although its clinical effect has yet to be proven.

The effect of shade and vermicompost application on yield and flavonoid levels of Tempuyung (Sonchus arvensis)

NASA Astrophysics Data System (ADS)

Putri, D. P.; Widyastuti, Y.; Dewi, W. S.; Yunus, A.

2018-03-01

This study aims to determine the effect of shade and vermicompost on yield and level of flavonoids in Tempuyung (Sonchus arvensis). The study was conducted in May- August 2016, in Tegal Gede Village, Kab. Karanganyar. Secondary metabolite content analysis was performed in B2P2TOOT laboratory, Tawangmangu. This study used a Completely Randomized Design (CRD) arranged in a split plot. Treatment consists of shade as main plot and the dose of vermicompost as sub plot. Shade treatments are 0% (without shade), 50% and 75%. The dose of vermicompost are 0 g / polybag (control), 250 g / polybag, 500 g / polybag and 750 g / polybag. Each treatment was repeated 3 times and each in experimental unit consist of 2 sample plants. The results showed that the higher levels of shade tend to decrease vegetative growth of plants. Moreover, vermicompost give significant effect on leaf weight, l number, wet and dry weight. The combination of 0% shade and 750 gram vermicompost gave the highest value (P <0.05) to leaf weight ± 186.68 g, wet weight ± 252.08 g and dry weight ± 30,76 gTempuyung. The combination of 50% shade with 0 g vermicompost and 75% shade with 250 g vermicompost show to increasing the content of flavonoid compounds.

Segmentation-based retrospective shading correction in fluorescence microscopy E. coli images for quantitative analysis

NASA Astrophysics Data System (ADS)

Mai, Fei; Chang, Chunqi; Liu, Wenqing; Xu, Weichao; Hung, Yeung S.

2009-10-01

Due to the inherent imperfections in the imaging process, fluorescence microscopy images often suffer from spurious intensity variations, which is usually referred to as intensity inhomogeneity, intensity non uniformity, shading or bias field. In this paper, a retrospective shading correction method for fluorescence microscopy Escherichia coli (E. Coli) images is proposed based on segmentation result. Segmentation and shading correction are coupled together, so we iteratively correct the shading effects based on segmentation result and refine the segmentation by segmenting the image after shading correction. A fluorescence microscopy E. Coli image can be segmented (based on its intensity value) into two classes: the background and the cells, where the intensity variation within each class is close to zero if there is no shading. Therefore, we make use of this characteristics to correct the shading in each iteration. Shading is mathematically modeled as a multiplicative component and an additive noise component. The additive component is removed by a denoising process, and the multiplicative component is estimated using a fast algorithm to minimize the intra-class intensity variation. We tested our method on synthetic images and real fluorescence E.coli images. It works well not only for visual inspection, but also for numerical evaluation. Our proposed method should be useful for further quantitative analysis especially for protein expression value comparison.

Growth and production of new superior rice varieties in the shade intensity

NASA Astrophysics Data System (ADS)

Alridiwirsah; Harahap, E. M.; Akoeb, E. N.; Hanum, H.

2018-02-01

Shade intensity is one of the most important requirements for plant growth, affecting growth, development, survival, and crop productivity. This study aims to evaluate the growth and productiom of New Superior Rice Varieties In The shade Intensity. This study was conducted in Balai Pengkajian Teknologi Pertanian, Pagar Merbau, Deli Serdang, North Sumatra. The research used completely randomized design with twofactors. The shade intensity (N) were 25%, 50% and no shade intensity as a control. Whereas new superior rice varieties were V1: Inpara 2, V2: Suluttan Unsrat 2, V3: Inpari Mugibat, V4: Inpari Sidenuk, V5: Mekongga, V6: Ciherang, V7:Inpari 10, V8: Inpari 3, V9: Inpari 4, V10: Inpari 30, dan V11: Cibogo. The result indicated that new superior rice varietiesshowedsignificant effectonthe growth and productionvariablesuch as leaf area, where Inpari Sidenuk variety was the highest among the varieties. Total chorophyll, the highest was found on Inpari variety. Number of tillers and plant height where the highest was found on Ciherang variety. The shade intensity showed significant effect on leaf area, where 25% shade intensity was the highest. Total chlorophyll, the highest was found on 50% shade intensity, number of tillers, the highest was found on no shade intensity.

Shading decreases the abundance of the herbivorous California horn snail, Cerithidea californica

USGS Publications Warehouse

Lorda, Julio; Lafferty, Kevin D.

2012-01-01

Most of the intertidal zone in estuaries of California, USA and Baja California, Mexico is covered with vascular vegetation. Shading by these vascular plants influences abiotic and biotic processes that shape benthic community assemblages. We present data on the effects of shading on the California horn snail, Cerithidea californica. This species is important because it is the most common benthic macrofaunal species in these systems and acts as an obligate intermediate host of several species of rematode parasites that infect several other species. Using observational and experimental studies, we found a negative effect of shade on the distribution and abundance of the California horn snail. We hypothesized that shading reduces the abundance of the epipelic diatoms that the snails feeds on, causing snails to leave haded areas. We observed a negative relationship between vascular plant cover, sub-canopy light levels, and snail density in Mugu Lagoon. Then we experimentally manipulated light regimes, by clipping vegetation and adding shade structures, and found higher snail densities at higher light levels. In Goleta Slough, we isolated the effect of shade from vegetation by documenting a negative relationship between the shade created by two bridges and diatom and snail densities. We also found that snails moved the greatest distances over shaded channel banks compared to unshaded channel banks. Further, we documented the effect of water depth and channel bank orientation on shading in this system. An additional effect of shading is the reduction of temperature, providing an alternative explanation for some of our results. These results broaden our knowledge of how variation in the light environment influences the ecology of estuarine ecosystems.

Could digital imaging be an alternative for digital colorimeters?

PubMed

Caglar, Alper; Yamanel, Kivanc; Gulsahi, Kamran; Bagis, Bora; Ozcan, Mutlu

2010-12-01

This study evaluated the colour parameters of composite and ceramic shade guides determined using a colorimeter and digital imaging method with illuminants at different colour temperatures. Two different resin composite shade guides, namely Charisma (Heraeus Kulzer) and Premise (Kerr Corporation), and two different ceramic shade guides, Vita Lumin Vacuum (VITA Zahnfabrik) and Noritake (Noritake Co.), were evaluated at three different colour temperatures (2,700 K, 2,700-6,500 K, and 6500 K) of illuminants. Ten shade tabs were selected (A1, A2, A3, A3,5, A4, B1, B2, B3, C2 and C3) from each shade guide. CIE Lab values were obtained using digital imaging and a colorimeter (ShadeEye NCC Dental Chroma Meter, Shofu Inc.). The data were analysed using two-way ANOVA, and Pearson's correlation. While mean L* values of both composite and ceramic shade guides were not affected from the colour temperature, L* values obtained with the colorimeter showed significantly lower values than those of the digital imaging (p < 0.01). At combined 2,700-6500 K colour temperature, the means of a* values obtained from colorimeter and digital imaging did not show significant differences (p > 0.05). For both composite and ceramic shade guides, L* and b* values obtained from colorimeter and digital imaging method presented a high level of correlation. High-level correlations were also acquired for a* values in all shade guides except for the Charisma composite shade guide. Digital imaging method could be an alternative for the colorimeters unless the proper object-camera distance, digital camera settings and suitable illumination conditions could be supplied. However, variations in shade guides, especially for composites, may affect the correlation.

Shade matching assisted by digital photography and computer software.

PubMed

Schropp, Lars

2009-04-01

To evaluate the efficacy of digital photographs and graphic computer software for color matching compared to conventional visual matching. The shade of a tab from a shade guide (Vita 3D-Master Guide) placed in a phantom head was matched to a second guide of the same type by nine observers. This was done for twelve selected shade tabs (tests). The shade-matching procedure was performed visually in a simulated clinic environment and with digital photographs, and the time spent for both procedures was recorded. An alternative arrangement of the shade tabs was used in the digital photographs. In addition, a graphic software program was used for color analysis. Hue, chroma, and lightness values of the test tab and all tabs of the second guide were derived from the digital photographs. According to the CIE L*C*h* color system, the color differences between the test tab and tabs of the second guide were calculated. The shade guide tab that deviated least from the test tab was determined to be the match. Shade matching performance by means of graphic software was compared with the two visual methods and tested by Chi-square tests (alpha= 0.05). Eight of twelve test tabs (67%) were matched correctly by the computer software method. This was significantly better (p < 0.02) than the performance of the visual shade matching methods conducted in the simulated clinic (32% correct match) and with photographs (28% correct match). No correlation between time consumption for the visual shade matching methods and frequency of correct match was observed. Shade matching assisted by digital photographs and computer software was significantly more reliable than by conventional visual methods.

Effect of abutment shade, ceramic thickness, and coping type on the final shade of zirconia all-ceramic restorations: in vitro study of color masking ability.

PubMed

Oh, Seon-Hee; Kim, Seok-Gyu

2015-10-01

The aim of the study was to evaluate the effect of abutment shade, ceramic thickness, and coping type on the final shade of zirconia all-ceramic restorations. Three different types of disk-shaped zirconia coping specimens (Lava, Cercon, Zirkonzahn: ø10 mm × 0.4 mm) were fabricated and veneered with IPS e.max Press Ceram (shade A2), for total thicknesses of 1 and 1.5 mm. A total of sixty zirconia restoration specimens were divided into six groups based on their coping types and thicknesses. The abutment specimens (ø10 mm × 7 mm) were prepared with gold alloy, base metal (nickel-chromium) alloy, and four different shades (A1, A2, A3, A4) of composite resins. The average L*, a*, b* values of the zirconia specimens on the six abutment specimens were measured with a dental colorimeter, and the statistical significance in the effects of three variables was analyzed by using repeated measures analysis of variance (α=.05).The average shade difference (ΔE) values of the zirconia specimens between the A2 composite resin abutment and other abutments were also evaluated. The effects of zirconia specimen thickness (P<.001), abutment shade (P<.001), and type of zirconia copings (P<.003) on the final shade of the zirconia restorations were significant. The average ΔE value of Lava specimens (1 mm) between the A2 composite resin and gold alloy abutments was higher (close to the acceptability threshold of 5.5 ΔE) than th ose between the A2 composite resin and other abutments. This in-vitro study demonstrated that abutment shade, ceramic thickness, and coping type affected the resulting shade of zirconia restorations.

Potential energy savings with exterior shades in large office buildings and the impact of discomfort glare

DOE Office of Scientific and Technical Information (OSTI.GOV)

Hoffmann, Sabine; Lee, Eleanor

Exterior shades are highly efficient for reducing solar load in commercial buildings. Their impact on net energy use depends on the annual energy balance of heating, cooling, fan and lighting energy. This paper discusses the overall energy use intensity of various external shading systems for a prototypical large office building split into the different types of energy use and for different orientations and window sizes. Lighting energy was calculated for a constant lighting power as well as for dimmed lighting fixtures (daylighting control). In Section 3, slat angles and solar cut-off angles were varied for fixed exterior slat shading systems.more » While the most light-blocking shades performed best for the case without daylighting controls, the optimum cut-off angle with daylighting controls was found to be 30 deg for the office building prototype used in Chicago and Houston. For large window-to-wall (WWR) ratios, window related annual energy use could be reduced by at least 70 % without daylighting control and by a minimum of 86 % with daylighting control in average over all orientations. The occurrence of discomfort glare was is considered in Section 4 of the paper, which looks at the performance of commercially available exterior shading systems when an interior shade is used in addition to the exterior shade during hours when occupants would experience discomfort glare. Glare control impacts overall energy use intensity significantly for exterior shades with high transmittance, especially when daylighting controls are used. In these cases, exterior shades are only beneficial for window-to-wall areas ≥ 45% in the hot Houston climate. For smaller windows and in a heating/cooling climate like Chicago, exterior shades can increase energy consumption« less

Evaluating factors that affect the shade-matching ability of dentists, dental staff members and laypeople.

PubMed

Capa, Nuray; Malkondu, Ozlem; Kazazoglu, Ender; Calikkocaoglu, Senih

2010-01-01

The authors conducted a study to evaluate the influence of dentists' and nondentists' experience, age, sex, eye color and use of eyeglasses or contact lenses on tooth shade-matching ability. The authors included 120 participants in this study conducted in Istanbul (periodontists, oral and maxillofacial surgeons, orthodontists, endodontists, pediatric dentists, prosthodontists, restorative dentists, general dentists in private practice, dental technicians, dental assistants, dental assistant students and laypeople). The authors assigned participants to one of three groups: group 1 was composed of prosthodontists, restorative dentists and dental technicians; group 2 consisted of other dental specialists and general dentists; and group 3 included dental assistants, dental assistant students and laypeople. The authors asked participants to match the shades of three artificial maxillary right central incisors (Vitapan acrylic teeth [shades 2L1.5, 1M2, 2R1.5], Vita Zahnfabrik, Bad Säckingen, Germany) by using a shade guide system (Vita Toothguide 3D-Master, Vita Zahnfabrik). They calculated shade matching for the three color components (value, hue, chroma) and analyzed the results by using a chi(2) test. The rate of success in matching the shade for IM2 was 53.3 percent for participants in group 1, 30 percent for participants in group 2 and 20 percent for participants in group 3 (P = .017). However, there were no significant differences between the three groups for shades 2L1.5 and 2R1.5. Professional experience (P = .003) and age (P = .027) were associated with shade-matching success for tooth shade 2L1.5 only. The results showed no statistically significant differences with respect to sex, eye color or use of eyeglasses or contact lenses. Dental care professionals who routinely performed restorative procedures matched the shades better than did participants in other groups. Professional experience was associated positively with the outcome, while sex, eye color and use of eyeglasses or contact lenses did not have any effect on shade-matching results. To improve shade-matching skills, clinicians should participate in hands-on courses, continuing education classes and other training programs.

Colour parameters and shade correspondence of CAD-CAM ceramic systems.

PubMed

Della Bona, Alvaro; Pecho, Oscar E; Ghinea, Razvan; Cardona, Juan C; Pérez, María M

2015-06-01

To evaluate colour differences between (1) CAD-CAM ceramic systems considering shades A1, A2 and A3 and the corresponding nominal shade of VC (Vita Classical shade guide) and (2) shades A1-A2, A2-A3 and A1, A2 and A3 within the same ceramic system. Samples of shades A1, A2 and A3 were fabricated (n=5) from CAD-CAM ceramic blocks (IPS e.max(®) CAD LT and HT, IPS Empress(®) CAD LT and HT, Paradigm™ C, and VITABLOCS(®) Mark II) and polished to 1.0±0.01mm in thickness. Spectral reflectance and colour coordinates were measured using a spectroradiometer inside a viewing booth using the CIE D65 illuminant and the d/0° geometry. Spectral reflectance curves were compared using VAF coefficient and were statistically analyzed using Kruskal-Wallis and the Mann-Whitney U test (α=0.05). Colour coordinates were statistically analyzed using one-way ANOVA, Tukey's test with Bonferroni correction (α=0.001). All colour differences (ΔEab(*) and ΔE00) were analyzed through comparisons with the PT - perceptibility and AT - acceptability thresholds for dental ceramics. ΔE between ceramic systems and its corresponding shade ranged from 6.32 to 13.42 (ΔEab(*)) and 4.48 to 9.30 (ΔE00). ΔE between shades A1-A2, A2-A3 and A1, A2 and A3 ranged, respectively, 1.93-4.82, 1.22-5.59 and 3.63-8.84 (ΔEab(*)); 1.54-3.87, 1.03-3.90 and 2.95-6.51 (ΔE00). Considering the corresponding nominal shade from VC, none of the ceramic systems showed colour differences below the AT. In addition, some ceramic systems showed colour differences below AT (shades A1-A2 and A2-A3) and below PT (shades A2-A3). Careful adjustments should be made to the final shade of CAD-CAM ceramic restorations to reach a clinically acceptable shade match. Copyright © 2015 Elsevier Ltd. All rights reserved.

From Charles Darwin's botanical country-house studies to modern plant biology.

PubMed

Kutschera, U; Briggs, W R

2009-11-01

As a student of theology at Cambridge University, Charles Darwin (1809-1882) attended the lectures of the botanist John S. Henslow (1796-1861). This instruction provided the basis for his life-long interest in plants as well as the species question. This was a major reason why in his book On the Origin of Species, which was published 150 years ago, Darwin explained his metaphorical phrase 'struggle for life' with respect to animals and plants. In this article, we review Darwin's botanical work with reference to the following topics: the struggle for existence in the vegetable kingdom with respect to the phytochrome-mediated shade avoidance response; the biology of flowers and Darwin's plant-insect co-evolution hypothesis; climbing plants and the discovery of action potentials; the power of movement in plants and Darwin's conflict with the German plant physiologist Julius Sachs; and light perception by growing grass coleoptiles with reference to the phototropins. Finally, we describe the establishment of the scientific discipline of Plant Biology that took place in the USA 80 years ago, and define this area of research with respect to Darwin's work on botany and the physiology of higher plants.

Chlorophyll fluorescence kinetics, photosynthetic activity, and pigment composition of blue-shade and half-shade leaves as compared to sun and shade leaves of different trees.

PubMed

Lichtenthaler, Hartmut K; Babani, Fatbardha; Navrátil, Martin; Buschmann, Claus

2013-11-01

The chlorophyll (Chl) fluorescence induction kinetics, net photosynthetic CO2 fixation rates P N, and composition of photosynthetic pigments of differently light exposed leaves of several trees were comparatively measured to determine the differences in photosynthetic activity and pigment adaptation of leaves. The functional measurements were carried out with sun, half-shade and shade leaves of seven different trees species. These were: Acer platanoides L., Ginkgo biloba L., Fagus sylvatica L., Platanus x acerifolia Willd., Populus nigra L., Quercus robur L., Tilia cordata Mill. In three cases (beech, ginkgo, and oak), we compared the Chl fluorescence kinetics and photosynthetic rates of blue-shade leaves of the north tree crown receiving only blue sky light but no direct sunlight with that of sun leaves. In these cases, we also determined in detail the pigment composition of all four leaf types. In addition, we determined the quantum irradiance and spectral irradiance of direct sunlight, blue skylight as well as the irradiance in half shade and full shade. The results indicate that sun leaves possess significantly higher mean values for the net CO2 fixation rates P N (7.8-10.7 μmol CO2 m(-2) s(-1) leaf area) and the Chl fluorescence ratio R Fd (3.85-4.46) as compared to shade leaves (mean P N of 2.6-3.8 μmol CO2 m(-2) s(-1) leaf area.; mean R Fd of 1.94-2.56). Sun leaves also exhibit higher mean values for the pigment ratio Chl a/b (3.14-3.31) and considerably lower values for the weight ratio total chlorophylls to total carotenoids, (a + b)/(x + c), (4.07-4.25) as compared to shade leaves (Chl a/b 2.62-2.72) and (a + b)/(x + c) of 5.18-5.54. Blue-shade and half-shade leaves have an intermediate position between sun and shade leaves in all investigated parameters including the ratio F v/F o (maximum quantum yield of PS2 photochemistry) and are significantly different from sun and shade leaves but could not be differentiated from each other. The mean values of the Chl fluorescence decrease ratio R Fd of blue-shade and half-shade leaves fit well into the strong linear correlation with the net photosynthetic rates P N of sun and shade leaves, thus unequivocally indicating that the determination of the Chl fluorescence decrease ratio R Fd is a fast and indirect measurement of the photosynthetic activity of leaves. The investigations clearly demonstrate that the photosynthetic capacity and pigment composition of leaves and chloroplasts strongly depend on the amounts and quality of light received by the leaves.

Teaching of color in predoctoral and postdoctoral dental education in 2009.

PubMed

Paravina, Rade D; O'Neill, Paula N; Swift, Edward J; Nathanson, Dan; Goodacre, Charles J

2010-01-01

The goal of the study was to determine the current status of the teaching of color in dental education at both the predoctoral (Pre-D) and postdoctoral (Post-D) levels. A cross-sectional web-based survey, containing 27 multiple choice, multiple best and single best answers was created. Upon receiving the administrative approval, dental faculty involved in the teaching of color to Pre-D or Post-D dental students from around the world (N=205), were administered a survey. Statistical analysis of differences between Pre-D and Post-D was performed using Chi-square test (α=0.05). A total of 130 responses were received (response rate 63.4%); there were 70 responses from North America, 40 from Europe, 10 from South America, nine from Asia and one from Africa. A course on "color" or "color in dentistry" was included in the dental curriculum of 80% of Pre-D programs and 82% of Post-D programs. The number of hours dedicated to color-related topics was 4.0±2.4 for Pre-D and 5.5±2.9 for Post-D, respectively (p<0.01). Topics associated with tooth color, shade matching method, tooth whitening, and teaching of appearance parameters other than color, were frequently taught. Significant differences were recorded between the number of hours dedicated to teaching of color at predoctoral and postdoctoral level. The same is true for the prosthodontics and restorative courses, teaching on negative after images; color rendering index, Bleachedguide 3D-Master shade guide, digital camera and lens selection, composite resins, and maxillofacial prosthetic materials. Except for the restorative courses and composite resins, significantly higher results were recorded for Post-D programs. Vitapan Classical and 3D-Master were the most frequently taught shade guides. Copyright © 2010 Elsevier Ltd. All rights reserved.

Tree seedlings respond to both light and soil nutrients in a Patagonian evergreen-deciduous forest.

PubMed

Promis, Alvaro; Allen, Robert B

2017-01-01

Seedlings of co-occurring species vary in their response to resource availability and this has implications for the conservation and management of forests. Differential shade-tolerance is thought to influence seedling performance in mixed Nothofagus betuloides-Nothofagus pumilio forests of Patagonia. However, these species also vary in their soil nutrient requirements. To determine the effects of light and soil nutrient resources on small seedlings we examined responses to an experimental reduction in canopy tree root competition through root trenching and restricting soil nutrient depletion through the addition of fertilizer. To understand the effect of light these treatments were undertaken in small canopy gaps and nearby beneath undisturbed canopy with lower light levels. Seedling diameter growth was greater for N. pumilio and height growth was greater for N. betuloides. Overall, diameter and height growth were greater in canopy gaps than beneath undisturbed canopy. Such growths were also greater with fertilizer and root trenching treatments, even beneath undisturbed canopy. Seedling survival was lower under such treatments, potentially reflecting thinning facilitated by resource induced growth. Finally, above-ground biomass did not vary among species although the less shade tolerant N. pumilio had higher below-ground biomass and root to shoot biomass ratio than the more shade tolerant N. betuloides. Above- and below-ground biomass were higher in canopy gaps so that the root to shoot biomass ratio was similar to that beneath undisturbed canopy. Above-ground biomass was also higher with fertilizer and root trenching treatments and that lowered the root to shoot biomass ratio. Restricting soil nutrient depletion allowed seedlings of both species to focus their responses above-ground. Our results support a view that soil nutrient resources, as well as the more commonly studied light resources, are important to seedlings of Nothofagus species occurring on infertile soils.

Auxin-to-Gibberellin Ratio as a Signal for Light Intensity and Quality in Regulating Soybean Growth and Matter Partitioning

PubMed Central

Yang, Feng; Fan, Yuanfang; Wu, Xiaoling; Cheng, Yajiao; Liu, Qinlin; Feng, Lingyang; Chen, Junxu; Wang, Zhonglin; Wang, Xiaochun; Yong, Taiwen; Liu, Weiguo; Liu, Jiang; Du, Junbo; Shu, Kai; Yang, Wenyu

2018-01-01

The intensity and quality (red to far-red (R/Fr) ratio) of light directly affect growth of plant under shading. Gibberellins (GAs) and auxin [indole-3-acetic acid (IAA)] play important roles in mediating the shading adaptive responses of plants. Thus, the intensity and quality of the uncoupling light from shading were assessed to identify the influence of each component on the morphology and matter distribution of the leaf, stem, and petiole. This assessment was based on the changes in endogenous Gibberellin 1 (GA1) and IAA levels. Soybean plants were grown in a growth chamber with four treatments [normal (N), N+Fr, low (L), and L+Fr light]. Results revealed that the reductions in photosynthetically active radiation (PAR) and R/Fr ratio equally increased height and stem mass fractions (SMFs) of the soybean seedling. The light intensity significantly influenced the dry mass per unit area and mass fraction of soybean leaves, whereas the light quality regulated the petiole elongation and mass fraction. Low R/Fr ratio (high Fr light) increased the soybean biomass by improving the photosynthetic assimilation rate and quantum yield of photosystem II. In addition, the IAA and GA1 levels in the leaf, stem, and petiole did not reflect the growth response trends of each tissue toward light intensity and quality; however, trends of the IAA-to-GA1 content ratios were similar to those of the growth and matter allocation of each soybean tissue under different light environments. Therefore, the response of growth and matter allocation of soybean to light intensity and quality may be regulated by the IAA-to-GA1 content ratio in the tissues of the soybean plant. PMID:29441084

Carbon allocation to growth and storage in two evergreen species of contrasting successional status.

PubMed

Piper, Frida I; Sepúlveda, Paulina; Bustos-Salazar, Angela; Zúñiga-Feest, Alejandra

2017-05-01

A prevailing hypothesis in forest succession is that shade-tolerant species grow more slowly than shade-intolerant species, across light conditions, because they prioritize carbon (C) allocation to storage. We examined this hypothesis in a confamilial pair of species, including one of the fastest-growing tree species in the world ( Eucalyptus globulus ) and a shade-tolerant, slow-growing species ( Luma apiculata ). Seedlings were subjected to one out of four combinations of light (high vs. low) and initial defoliation (90% defoliated vs. nondefoliated) for four months. Growth, C storage concentration in different organs, leaf shedding, and lateral shoot formation were measured at the end of the experiment. Eucalyptus globulus grew faster than L. apiculata in high light, but not in low light. Both species had lower C storage concentration in low than in high light, but similar C storage concentrations in each light condition. Defoliation had no effect on C storage, except in the case of the old leaves of both species, which showed lower C storage levels in response to defoliation. Across treatments, leaf shedding was 96% higher in E. globulus than in L. apiculata while, in contrast, lateral shoot formation was 87% higher in L. apiculata . In low light, E. globulus prioritized C storage instead of growth, whereas L. apiculata prioritized growth and lateral branching. Our results suggest that shade tolerance depends on efficient light capture rather than C conservation traits. © 2017 Botanical Society of America.

The least-squares mixing models to generate fraction images derived from remote sensing multispectral data

NASA Technical Reports Server (NTRS)

Shimabukuro, Yosio Edemir; Smith, James A.

1991-01-01

Constrained-least-squares and weighted-least-squares mixing models for generating fraction images derived from remote sensing multispectral data are presented. An experiment considering three components within the pixels-eucalyptus, soil (understory), and shade-was performed. The generated fraction images for shade (shade image) derived from these two methods were compared by considering the performance and computer time. The derived shade images are related to the observed variation in forest structure, i.e., the fraction of inferred shade in the pixel is related to different eucalyptus ages.

Energy Savings Calculations for Heat Island Reduction Strategies in Baton Rouge, Sacramento and Salt Lake City

DOE Office of Scientific and Technical Information (OSTI.GOV)

Konopacki, S.; Akbari, H.

2000-03-01

In 1997, the US Environmental Protection Agency (EPA) established the ''Heat Island Reduction Initiative'', to quantify the potential benefits of Heat Island Reduction (HIR) strategies (i.e., shade trees, reflective roofs, reflective pavements and urban vegetation) to reduce cooling energy use in buildings, lower the ambient air temperature and improve urban air quality in cities, and reduce CO2 emissions from power plants. Under this initiative, the Urban Heat Island Pilot Project (UHIPP) was created with the objective to investigate the potential of HIR strategies in residential and commercial buildings in three initial UHIPP cities: Baton Rouge, Sacramento and Salt Lake City.more » This paper summarizes our efforts to calculate the annual energy savings, peak power avoidance and annual C02 reduction of HIR strategies in the three initial cities. In this analysis, we focused on three building types that offer most savings potential: single-family residence, office and retail store. Each building type was characterized in detail by old or new construction and with a gas furnace or an electric heat pump. We defined prototypical building characteristics for each building type and simulated the impact of HIR strategies on building cooling and heating energy use and peak power demand using the DOE-2.IE model. Our simulations included the impact of (1) strategically-placed shade trees near buildings [direct effect], (2) use of high-albedo roofing material on building [direct effect], (3) combined strategies I and 2 [direct effect], (4) urban reforestation with high-albedo pavements and building surfaces [indirect effect] and (5) combined strategies 1, 2 and 4 [direct and indirect effects]. We then estimated the total roof area of air-conditioned buildings in each city using readily obtainable data to calculate the metropolitan-wide impact of HIR strategies. The results show, that in Baton Rouge, potential annual energy savings of $15M could be realized by rate-payers from the combined direct and indirect effects of HIR strategies. Additionally, peak power avoidance is estimated at 133 MW and the reduction in annual carbon emissions at 41 kt. In Sacramento, the potential annual energy savings is estimated at $26M, with an avoidance of 486 MW in peak power and a reduction in annual carbon of 92 kt. In Salt Lake City, the potential annual energy savings is estimated at $4M, with an avoidance of 85 MW in peak power and a reduction in annual carbon of 20 kt.« less

Beampattern control of a microphone array to minimize secondary source contamination.

PubMed

Jordan, Peter; Fitzpatrick, John A; Meskell, Craig

2003-10-01

A null-steering technique is adapted and applied to a linear delay-and-sum beamformer in order to measure the noise generated by one of the propellers of a 1/8 scale twin propeller aircraft model. The technique involves shading the linear array using a set of weights, which are calculated according to the locations onto which the nulls need to be steered (in this case onto the second propeller). The technique is based on an established microwave antenna theory, and uses a plane-wave, or far field formulation in order to represent the response of the array by an nth-order polynomial, where n is the number of array elements. The roots of this polynomial correspond to the minima of the array response, and so by an appropriate choice of roots, a polynomial can be generated, the coefficients of which are the weights needed to achieve the prespecified set of null positions. It is shown that, for the technique to work with actual data, the cross-spectral matrix must be conditioned before array shading is implemented. This ensures that the shading function is not distorted by the intrinsic element weighting which can occur as a result of the directional nature of aeroacoustic systems. A difference of 6 dB between measurements before and after null steering shows the technique to have been effective in eliminating the contribution from one of the propellers, thus providing a quantitative measure of the acoustic energy from the other.

Seeing the landscape for the trees: Metrics to guide riparian shade management in river catchments

NASA Astrophysics Data System (ADS)

Johnson, Matthew F.; Wilby, Robert L.

2015-05-01

Rising water temperature (Tw) due to anthropogenic climate change may have serious consequences for river ecosystems. Conservation and/or expansion of riparian shade could counter warming and buy time for ecosystems to adapt. However, sensitivity of river reaches to direct solar radiation is highly heterogeneous in space and time, so benefits of shading are also expected to be site specific. We use a network of high-resolution temperature measurements from two upland rivers in the UK, in conjunction with topographic shade modeling, to assess the relative significance of landscape and riparian shade to the thermal behavior of river reaches. Trees occupy 7% of the study catchments (comparable with the UK national average) yet shade covers 52% of the area and is concentrated along river corridors. Riparian shade is most beneficial for managing Tw at distances 5-20 km downstream from the source of the rivers where discharge is modest, flow is dominated by near-surface hydrological pathways, there is a wide floodplain with little landscape shade, and where cumulative solar exposure times are sufficient to affect Tw. For the rivers studied, we find that approximately 0.5 km of complete shade is necessary to off-set Tw by 1°C during July (the month with peak Tw) at a headwater site; whereas 1.1 km of shade is required 25 km downstream. Further research is needed to assess the integrated effect of future changes in air temperature, sunshine duration, direct solar radiation, and downward diffuse radiation on Tw to help tree planting schemes achieve intended outcomes.

Porcelain thickness and cement shade effects on the colour and translucency of porcelain veneering materials.

PubMed

Kürklü, Duygu; Azer, Shereen S; Yilmaz, Burak; Johnston, William M

2013-11-01

Purposes of this in vitro study include evaluating colour changes in combinations of feldspathic porcelain and cement resulting from different thicknesses of porcelain and different shades of composite luting agent, and evaluating relative translucency parameter (RTP) values. Porcelain discs of shade A1 at nominal thicknesses of 0.5 and 1.0mm were bonded to cements of three shades in a factorial design. Colours were calculated for CIE D65 Illuminant and Standard Human Observer on black, grey and white backings. A colour difference (CD) was calculated of each possible pair of different porcelain thickness values for the same cement shade and each possible pair of different cement shades for the same porcelain thickness. RTP was analyzed by ANOVA and selected pairwise comparisons. All mean CDs studied were perceptible and most were at or greater than the clinical acceptability threshold, with the notable exception that the mean CDs and their confidence limits were below the clinical acceptability threshold for a change in porcelain thickness when utilizing the Clear cement shade. Variation in the shade of the resin luting cement will result in CDs which are near or beyond clinical acceptability. A decrease in porcelain thickness did significantly increase RTP when bonded to the resin cement shades studied. Changes in porcelain thickness or cement shade may adversely affect basic aesthetic properties of these materials. Development of methods for analyzing aesthetic effects over greater ranges of thickness for these materials would improve the prognosis for using these materials. Copyright © 2013 Elsevier Ltd. All rights reserved.

Tanning Shade Gradations of Models in Mainstream Fitness and Muscle Enthusiast Magazines: Implications for Skin Cancer Prevention in Men.

PubMed

Basch, Corey H; Hillyer, Grace Clarke; Ethan, Danna; Berdnik, Alyssa; Basch, Charles E

2015-07-01

Tanned skin has been associated with perceptions of fitness and social desirability. Portrayal of models in magazines may reflect and perpetuate these perceptions. Limited research has investigated tanning shade gradations of models in men's versus women's fitness and muscle enthusiast magazines. Such findings are relevant in light of increased incidence and prevalence of melanoma in the United States. This study evaluated and compared tanning shade gradations of adult Caucasian male and female model images in mainstream fitness and muscle enthusiast magazines. Sixty-nine U.S. magazine issues (spring and summer, 2013) were utilized. Two independent reviewers rated tanning shade gradations of adult Caucasian male and female model images on magazines' covers, advertisements, and feature articles. Shade gradations were assessed using stock photographs of Caucasian models with varying levels of tanned skin on an 8-shade scale. A total of 4,683 images were evaluated. Darkest tanning shades were found among males in muscle enthusiast magazines and lightest among females in women's mainstream fitness magazines. By gender, male model images were 54% more likely to portray a darker tanning shade. In this study, images in men's (vs. women's) fitness and muscle enthusiast magazines portrayed Caucasian models with darker skin shades. Despite these magazines' fitness-related messages, pro-tanning images may promote attitudes and behaviors associated with higher skin cancer risk. To date, this is the first study to explore tanning shades in men's magazines of these genres. Further research is necessary to identify effects of exposure to these images among male readers. © The Author(s) 2014.

[Effects of shading at different phases of grain-filling on wheat grain protein components contents and processing quality].

PubMed

Shi, Yu; Chen, Mao-xue; Yu, Zhen-wen; Xu, Zhen-zhu

2011-10-01

Taking three wheat cultivars Jimai 20 (strong gluten), Taishan 23 (medium gluten), and Ningmai 9 (weak gluten) as test materials, a field experiment was conducted to examine the effects of shading at different phases of grain-filling on the grain protein components contents and processing quality. Four treatments were installed, i. e., no shading (S0), shading at early grain-filling phase (from 0 day after anthesis (DAA) to 11 DAA; S1), shading at medium grain-filling phase (from 12 DAA to 23 DAA; S2), and shading at late grain-filling phase (from 24 DAA to 35 DAA; S3). No significant differences were observed in the grain albumin+globulin contents of the three cultivars among the four treatments. Shading increased the grain HMW-GS, LMW-GS, gluten, glutenin, and total protein contents of Jimai 20 and Taishan 23 significantly, and the increments were higher in treatment S2 than in other shading treatments. Treatments S2 and S3 increased the grain protein components contents of Ningmai 9 significantly. Comparing with the control, shading decreased the grain yield significantly, but increased the dough development time, dough stability time, and sedimentation volume, especially for treatment S2, which suggested that the wheat grain quality had a close relationship with the light intensity at medium phase of grain-filling. Overall, the regulation effect of shading at grain-filling stage on the wheat grain yield, grain protein components contents, and indices values of grain processing quality for the test cultivars was in the order of Jimai 20 > Taishan 23 > Ningmai 9.

Characterizing visual asymmetries in contrast perception using shaded stimuli.

PubMed

Chacón, José; Castellanos, Miguel Ángel; Serrano-Pedraza, Ignacio

2015-01-01

Previous research has shown a visual asymmetry in shaded stimuli where the perceived contrast depended on the polarity of their dark and light areas (Chacón, 2004). In particular, circles filled out with a top-dark luminance ramp were perceived with higher contrast than top-light ones although both types of stimuli had the same physical contrast. Here, using shaded stimuli, we conducted four experiments in order to find out if the perceived contrast depends on: (a) the contrast level, (b) the type of shading (continuous vs. discrete) and its degree of perceived three-dimensionality, (c) the orientation of the shading, and (d) the sign of the perceived contrast alterations. In all experiments the observers' tasks were to equate the perceived contrast of two sets of elements (usually shaded with opposite luminance polarity), in order to determine the subjective equality point. Results showed that (a) there is a strong difference in perceived contrast between circles filled out with luminance ramp top-dark and top-light that is similar for different contrast levels; (b) we also found asymmetries in contrast perception with different shaded stimuli, and this asymmetry was not related with the perceived three-dimensionality but with the type of shading, being greater for continuous-shading stimuli; (c) differences in perceived contrast varied with stimulus orientation, showing the maximum difference on vertical axis with a left bias consistent with the bias found in previous studies that used visual-search tasks; and (d) asymmetries are consistent with an attenuation in perceived contrast that is selective for top-light vertically-shaded stimuli.

[Effects of soil moisture content and light intensity on the plant growth and leaf physiological characteristics of squash].

PubMed

Du, She-ni; Bai, Gang-shuan; Liang, Yin-li

2011-04-01

A pot experiment with artificial shading was conducted to study the effects of soil moisture content and light intensity on the plant growth and leaf physiological characteristics of squash variety "Jingyingyihao". Under all test soil moisture conditions, 30% shading promoted the growth of "Jingyingyihao", with the highest yield at 70% - 80% soil relative moisture contents. 70% shading inhibited plant growth severely, only flowering and not bearing fruits, no economic yield produced. In all treatments, there was a similar water consumption trend, i. e., both the daily and the total water consumption decreased with increasing shading and decreasing soil moisture content. Among all treatments, 30% shading and 70% - 80% soil relative moisture contents had the highest water use efficiency (2.36 kg mm(-1) hm(-2)) and water output rate (1.57 kg mm(-1) hm(-2)). The net photosynthetic rate, transpiration rate, stomatal conductance, and chlorophyll content of squash leaves decreased with increasing shading, whereas the intercellular CO2 concentration was in adverse. The leaf protective enzyme activity and proline content decreased with increasing shading, and the leaf MAD content decreased in the order of 70% shading, natural radiation, and 30% shading. Under the three light intensities, the change characteristics of squash leaf photosynthesis, protective enzyme activity, and proline and MAD contents differed with the increase of soil relative moisture content.

Stream shading, summer streamflow and maximum water temperature following intense wildfire in headwater streams

Treesearch

Michael Amaranthus; Howard Jubas; David Arthur

1989-01-01

Adjacent headwater streams were monitored for postfire shade, summer streamflow and maximum water temperature following the 40,000 ha Silver Complex fire in southern Oregon. Average postfire shade (30 percent) for the three streams was considerably less than prefire shade (est.>90 percent). Dramatic increases in direct solar radiation resulted in large but variable...

An appraisal of the classic forest succession paradigm with the shade tolerance index.

PubMed

Lienard, Jean; Florescu, Ionut; Strigul, Nikolay

2015-01-01

In this paper we revisit the classic theory of forest succession that relates shade tolerance and species replacement and assess its validity to understand patch-mosaic patterns of forested ecosystems of the USA. We introduce a macroscopic parameter called the "shade tolerance index" and compare it to the classic continuum index in southern Wisconsin forests. We exemplify shade tolerance driven succession in White Pine-Eastern Hemlock forests using computer simulations and analyzing approximated chronosequence data from the USDA FIA forest inventory. We describe this parameter across the last 50 years in the ecoregions of mainland USA, and demonstrate that it does not correlate with the usual macroscopic characteristics of stand age, biomass, basal area, and biodiversity measures. We characterize the dynamics of shade tolerance index using transition matrices and delimit geographical areas based on the relevance of shade tolerance to explain forest succession. We conclude that shade tolerance driven succession is linked to climatic variables and can be considered as a primary driving factor of forest dynamics mostly in central-north and northeastern areas in the USA. Overall, the shade tolerance index constitutes a new quantitative approach that can be used to understand and predict succession of forested ecosystems and biogeographic patterns.

Comparison of shade matching by visual observation and an intraoral dental colorimeter.

PubMed

Li, Q; Wang, Y N

2007-11-01

The purpose of this study was to compare the applicability of two shade-matching approaches: Vintage Halo shade guide (visual method) and Shofu ShadeEye NCC colorimeter (instrumental method). Twenty participants' maxillary left central incisors were evaluated. Corresponding metal ceramic crowns were fabricated with each shade-matching approach. The colour distributions (L*, a* and b*) of the middle third region of each tooth and corresponding metal ceramic crowns were spectrophotometrically assessed. The colour difference (DeltaE) and colour distributions (DeltaL*, Deltaa* and Deltab*) between the tooth and the corresponding crowns were calculated. We found that the colour differences of both groups fell within the clinical unacceptable range (DeltaE > 2.75). Regarding DeltaE and the three colour distributions, no significant difference was found, expect for a* (P < 0.01). The shade matching difficulty degree was analysed through the agreements of visual shade selection. Within easy matching cases, the instrumental method achieved better results (P = 0.041). In conclusion, it is suggested that the reliability of shade matching can be ensured by neither the colorimeter nor the visual approach. However, the colorimeter can achieve better results within easy matching cases.

The effect of irradiance on the carbon balance and tissue characteristics of five herbaceous species differing in shade-tolerance

PubMed Central

Pons, Thijs L.; Poorter, Hendrik

2014-01-01

The carbon balance is defined here as the partitioning of daily whole-plant gross CO2 assimilation (A) in C available for growth and C required for respiration (R). A scales positively with growth irradiance and there is evidence for an irradiance dependence of R as well. Here we ask if R as a fraction of A is also irradiance dependent, whether there are systematic differences in C-balance between shade-tolerant and shade-intolerant species, and what the causes could be. Growth, gas exchange, chemical composition and leaf structure were analyzed for two shade-tolerant and three shade-intolerant herbaceous species that were hydroponically grown in a growth room at five irradiances from 20 μmol m−2 s−1 (1.2 mol m−2 day−1) to 500 μmol m−2 s−1 (30 mol m−2 day−1). Growth analysis showed little difference between species in unit leaf rate (dry mass increase per unit leaf area) at low irradiance, but lower rates for the shade-tolerant species at high irradiance, mainly as a result of their lower light-saturated rate of photosynthesis. This resulted in lower relative growth rates in these conditions. Daily whole-plant R scaled with A in a very tight manner, giving a remarkably constant R/A ratio of around 0.3 for all but the lowest irradiance. Although some shade-intolerant species showed tendencies toward a higher R/A and inefficiencies in terms of carbon and nitrogen investment in their leaves, no conclusive evidence was found for systematic differences in C-balance between the shade-tolerant and intolerant species at the lowest irradiance. Leaf tissue of the shade-tolerant species was characterized by high dry matter percentages, C-concentration and construction costs, which could be associated with a better defense in shade environments where leaf longevity matters. We conclude that shade-intolerant species have a competitive advantage at high irradiance due to superior potential growth rates, but that shade-tolerance is not necessarily associated with a better C-balance at low irradiance. Under those conditions tolerance to other stresses is probably more important for the performance of shade-tolerant species. PMID:24550922

75 FR 52724 - Procurement List Additions

Federal Register 2010, 2011, 2012, 2013, 2014

2010-08-27

...: 8405-00-FAB-0201--Man's, S/S, Air Force Shade 1550, Size 13\\1/ 2\\ NSN: 8405-00-FAB-0202--Man's, S/S, Air Force Shade 1550, Size 14 NSN: 8405-00-FAB-0203--Man's, S/S, Air Force Shade 1550, Size 14 \\1/ 2\\ NSN: 8405-00-FAB-0204--Man's, S/S, Air Force Shade 1550, Size 15 NSN: 8405-00-FAB-0212--Man's, S/S...

Ranking the shade tolerance of forty-five candidate groundcovers for agroforestry plantings

Treesearch

J.W. Van Sambeek; N.E. Navarrete-Tindall; H.E. Garrett; C.-H. Lin; R.L. McGraw; D.C. Wallace

2007-01-01

Several large-scale screening trials evaluating native and introduced herbaceous ground covers have been conducted in the last half century. Most trials have used shade cloth to evaluate growth of potted plants under moderate shade (45 to 55 percent of full sunlight) similar to what might be found in many agroforestry practices and heavy shade (20 to 30 percent of full...

An appraisal of the classic forest succession paradigm with the shade tolerance index

Treesearch

Jean Lienard; Ionut Florescu; Nikolay Strigul

2015-01-01

We revisit the classic theory of forest succession that relates shade tolerance and species replacement and assess its validity to understand patch-mosaic patterns of forested ecosystems of the USA. We introduce a macroscopic parameter called the “shade tolerance index” and compare it to the classic continuum index in southern Wisconsin forests. We exemplify shade...

WETLAND INVERTEBRATE COMMUNITY RESPONSES TO VARYING EMERGENT LITTER IN A PRAIRIE POTHOLE EMERGENT MARSH

EPA Science Inventory

Plant litter produced in the interior of dense emergent stands may directly or indirectly influence invertebrate communities. Low litter may provide structure and refuge to invertebrates while high litter may shade out vegetation and algae and decrease oxygen concentrations. With...

A CLIMATE-RESPONSIVE ADAPTIVE CONTROL FOR A COMBINATION PASSIVE SOLAR SHADING AND NATURAL VENTILATION

EPA Science Inventory

Currently, nations around the globe are facing striking concerns regarding energy consumption. In the United States, we face increasing demands that will cause increasing fuel prices thus ultimately higher-energy cost. The future could be eased by reduce energy consumption ...

Sun protective behaviors and vitamin D levels in the US population: NHANES 2003–2006

PubMed Central

Linos, Eleni; Keiser, Elizabeth; Kanzler, Matthew; Sainani, Kristin L.; Lee, Wayne; Vittinghoff, Eric; Chren, Mary-Margaret; Tang, Jean Y.

2013-01-01

Background Sun protection is recommended for skin cancer prevention, yet little is known about the role of sun protection on vitamin D levels. Our aim was to investigate the relationship between different types of sun protective behaviors and serum 25(OH)D levels in the general US population. Methods Cross-sectional, nationally representative survey of 5,920 adults aged 18–60 years in the US National Health and Nutrition Examination Survey 2003–2006. We analyzed questionnaire responses on sun protective behaviors: staying in the shade, wearing long sleeves, wearing a hat, using sunscreen and SPF level. Analyses were adjusted for multiple confounders of 25(OH)D levels and stratified by race. Our primary outcome measures were serum 25(OH)D levels (ng/ml) measured by radioimmunoassay and vitamin D deficiency, defined as 25(OH)D levels <20 ng/ml. Results Staying in the shade and wearing long sleeves were significantly associated with lower 25(OH)D levels. Subjects who reported frequent use of shade on a sunny day had −3.5 ng/ml (ptrend < 0.001) lower 25(OH)D levels compared to subjects who reported rare use. Subjects who reported frequent use of long sleeves had −2.2 ng/ml (ptrend = 0.001) lower 25(OH)D levels. These associations were strongest for whites, and did not reach statistical significance among Hispanics or blacks. White participants who reported frequently staying in the shade or wearing long sleeves had double the odds of vitamin D deficiency compared with those who rarely did so. Neither wearing a hat nor using sunscreen was associated with low 25(OH)D levels or vitamin D deficiency. Conclusions White individuals who protect themselves from the sun by seeking shade or wearing long sleeves may have lower 25(OH)D levels and be at risk for vitamin D deficiency. Frequent sunscreen use does not appear to be linked to vitamin D deficiency in this population. PMID:22045154

Too hot to die? The effects of vegetation shading on past, present, and future activity budgets of two diurnal skinks from arid Australia.

PubMed

Grimm-Seyfarth, Annegret; Mihoub, Jean-Baptiste; Henle, Klaus

2017-09-01

Behavioral thermoregulation is an important mechanism allowing ectotherms to respond to thermal variations. Its efficiency might become imperative for securing activity budgets under future climate change. For diurnal lizards, thermal microhabitat variability appears to be of high importance, especially in hot deserts where vegetation is highly scattered and sensitive to climatic fluctuations. We investigated the effects of a shading gradient from vegetation on body temperatures and activity timing for two diurnal, terrestrial desert lizards, Ctenotus regius, and Morethia boulengeri , and analyzed their changes under past, present, and future climatic conditions. Both species' body temperatures and activity timing strongly depended on the shading gradient provided by vegetation heterogeneity. At high temperatures, shaded locations provided cooling temperatures and increased diurnal activity. Conversely, bushes also buffered cold temperature by saving heat. According to future climate change scenarios, cooler microhabitats might become beneficial to warm-adapted species, such as C. regius, by increasing the duration of daily activity. Contrarily, warmer microhabitats might become unsuitable for less warm-adapted species such as M. boulengeri for which midsummers might result in a complete restriction of activity irrespective of vegetation. However, total annual activity would still increase provided that individuals would be able to shift their seasonal timing towards spring and autumn. Overall, we highlight the critical importance of thermoregulatory behavior to buffer temperatures and its dependence on vegetation heterogeneity. Whereas studies often neglect ecological processes when anticipating species' responses to future climate change the strongest impact of a changing climate on terrestrial ectotherms in hot deserts is likely to be the loss of shaded microhabitats rather than the rise in temperature itself. We argue that conservation strategies aiming at addressing future climate changes should focus more on the cascading effects of vegetation rather than on shifts of species distributions predicted solely by climatic envelopes.

Sun and Shade leaves, SIF, and Photosynthetic Capacity

NASA Astrophysics Data System (ADS)

Berry, J. A.; Badgley, G.

2016-12-01

Recent advances in retrieval of solar induced chlorophyll fluorescence (SIF) have opened up new possibilities for remote sensing of canopy physiology and structure. To date most of the emphasis has been placed on SIF as an indicator of stress and photosynthetic capacity. However, it is clear that canopy structure can also have an influence. To this point, simulations of SIF in land surface models tend to under predict observed variation in SIF. Also, large, systematic differences in SIF from different canopy types seem to correlate well with the photosynthetic capacity of these canopies. SIF emissions from pampered crops can be several-fold that from evergreen, needle-leaf forests. Yet, these may have similar vegetation indices and absorb a similar fraction of incident PAR. SIF photons produced in a conifer canopy do have a lower probability of escaping its dense, clumped foliage. However, this does not explain the correlated differences in photosynthetic rate and SIF. It is useful, in this regard, to consider the separate contributions of sun and shade leaves to the SIF emitted by a canopy. Sun leaves tend to be displayed to intercept the direct solar beam, and these highly illuminated leaves are often visible from above the canopy. Sun leaves produce more SIF and a large fraction of it escapes. Therefore, the intensity of SIF may be a sensitive indicator of the partitioning of absorbed PAR to sun and shade leaves. Many models account tor the different photosynthetic capacity of sun and shade leaves in calculating canopy responses. However, the fraction of leaves in each category is usually parameterized by an assumed leaf angle distribution (e.g. spherical). In reality, the sun/shade fraction can vary over a wide range, and it has been difficult to measure. SIF and possibly near-IR reflectance of canopies can be used to specify this key parameter with obvious importance to understanding photosynthetic rate.

Reliability of visual and instrumental color matching.

PubMed

Igiel, Christopher; Lehmann, Karl Martin; Ghinea, Razvan; Weyhrauch, Michael; Hangx, Ysbrand; Scheller, Herbert; Paravina, Rade D

2017-09-01

The aim of this investigation was to evaluate intra-rater and inter-rater reliability of visual and instrumental shade matching. Forty individuals with normal color perception participated in this study. The right maxillary central incisor of a teaching model was prepared and restored with 10 feldspathic all-ceramic crowns of different shades. A shade matching session consisted of the observer (rater) visually selecting the best match by using VITA classical A1-D4 (VC) and VITA Toothguide 3D Master (3D) shade guides and the VITA Easyshade Advance intraoral spectrophotometer (ES) to obtain both VC and 3D matches. Three shade matching sessions were held with 4 to 6 weeks between sessions. Intra-rater reliability was assessed based on the percentage of agreement for the three sessions for the same observer, whereas the inter-rater reliability was calculated as mean percentage of agreement between different observers. The Fleiss' Kappa statistical analysis was used to evaluate visual inter-rater reliability. The mean intra-rater reliability for the visual shade selection was 64(11) for VC and 48(10) for 3D. The corresponding ES values were 96(4) for both VC and 3D. The percentages of observers who matched the same shade with VC and 3D were 55(10) and 43(12), respectively, while corresponding ES values were 88(8) for VC and 92(4) for 3D. The results for visual shade matching exhibited a high to moderate level of inconsistency for both intra-rater and inter-rater comparisons. The VITA Easyshade Advance intraoral spectrophotometer exhibited significantly better reliability compared with visual shade selection. This study evaluates the ability of observers to consistently match the same shade visually and with a dental spectrophotometer in different sessions. The intra-rater and inter-rater reliability (agreement of repeated shade matching) of visual and instrumental tooth color matching strongly suggest the use of color matching instruments as a supplementary tool in everyday dental practice to enhance the esthetic outcome. © 2017 Wiley Periodicals, Inc.

Effect of abutment shade, ceramic thickness, and coping type on the final shade of zirconia all-ceramic restorations: in vitro study of color masking ability

PubMed Central

Oh, Seon-Hee

2015-01-01

PURPOSE The aim of the study was to evaluate the effect of abutment shade, ceramic thickness, and coping type on the final shade of zirconia all-ceramic restorations. MATERIALS AND METHODS Three different types of disk-shaped zirconia coping specimens (Lava, Cercon, Zirkonzahn: ø10 mm × 0.4 mm) were fabricated and veneered with IPS e.max Press Ceram (shade A2), for total thicknesses of 1 and 1.5 mm. A total of sixty zirconia restoration specimens were divided into six groups based on their coping types and thicknesses. The abutment specimens (ø10 mm × 7 mm) were prepared with gold alloy, base metal (nickel-chromium) alloy, and four different shades (A1, A2, A3, A4) of composite resins. The average L*, a*, b* values of the zirconia specimens on the six abutment specimens were measured with a dental colorimeter, and the statistical significance in the effects of three variables was analyzed by using repeated measures analysis of variance (α=.05).The average shade difference (ΔE) values of the zirconia specimens between the A2 composite resin abutment and other abutments were also evaluated. RESULTS The effects of zirconia specimen thickness (P<.001), abutment shade (P<.001), and type of zirconia copings (P<.003) on the final shade of the zirconia restorations were significant. The average ΔE value of Lava specimens (1 mm) between the A2 composite resin and gold alloy abutments was higher (close to the acceptability threshold of 5.5 ΔE) than th ose between the A2 composite resin and other abutments. CONCLUSION This in-vitro study demonstrated that abutment shade, ceramic thickness, and coping type affected the resulting shade of zirconia restorations. PMID:26576252

The effect of enamel porcelain thickness on color and the ability of a shade guide to prescribe chroma.

PubMed

Jarad, F D; Moss, B W; Youngson, C C; Russell, M D

2007-04-01

To test the null hypothesis that there is no color change when enamel porcelain thickness is changed and to evaluate the ability of a shade guide to prescribe chroma. Three shades (3M1, 3M2 and 3M3) were selected from a Vitapan 3D master shade guide. Five disk specimens were prepared for each shade, consisting of three layers (opacious dentin, dentin and enamel) at thicknesses of 0.6, 0.8 and 0.6mm, respectively. The color of each disk was measured using a spectrophotometer. Enamel porcelain was reduced in thickness to 0.3mm and porcelain disks were remeasured. Reducing the enamel thickness of porcelain disk specimens significantly increased L) (p<0.05), b*, metric chroma and hue angle (p<0.001). For the three shades studied (3M1, 3M2 and 3M3) L* values were not significantly different (p>0.05) and chroma increased for 3M1 with the lowest chroma to 3M3 with the highest chroma, which is in line with the shade guide specifications. Although statistically significant (p<0.001) changes in hue angle between the three shades were small (less than 3 degrees overall). The difference in chroma between the three shades 3M1, 3M2 and 3M3 was greatest for the thin enamel layer than the thick enamel layer. A change in enamel thickness from 0.6 to 0.3mm resulted in a three-unit change in L* and metric chroma and a 4 degrees change in hue angle. A change in enamel porcelain thickness will have a greater effect on higher chromatic shades than those with lower chroma. The ability of the shade guide to prescribe chroma was demonstrated but this could be offset by an anomalous enamel thickness.

Modeling shade tree use by beef cattle as a function of black globe temperature and time of day

NASA Astrophysics Data System (ADS)

Foust, Amanda M.; Headlee, William L.

2017-12-01

Increasing temperatures associated with global climate change threaten to disrupt agricultural systems such as beef production, yet relatively little is known about the use of natural tree shade to mitigate the negative effects of heat stress on beef cattle. In this study, we evaluated how temperature and time of day influenced the utilization of tree shade in relation to coloration, orientation, and behavior of beef cattle in a pasture system. Temperatures in shade and direct sunlight were measured using black globe temperature (BGT) data loggers. Time-lapse images from game cameras were used to obtain counts of shade usage, coloration, orientation, and behavior of cattle throughout the daytime hours. In general, we found that shade utilization and most of the predominating orientations and behaviors differed significantly ( P < 0.05) by both time of day (Hour) and BGT in direct sunlight (BGTsun), while interactions between these two effects (Hour × BGTsun) were often nonsignificant. The mean percentage of the herd using shade was highest in mid-morning (87-96%) and early afternoon (97%), but also increased with BGTsun regardless of the time of day; these trends were similar for both dark- and light-colored cattle. Lying down was the dominant behavior exhibited in the shade, while foraging was the most prevalent behavior in the sun. When herd shade usage was lowest in mid- to late-afternoon (<1%) we also observed an increase in the use of heat-mitigating orientations in the sun (37-47%). We discuss some practical implications of these results, including the potential use of temperature thresholds to interpret cattle behaviors and shade usage.

Modeling shade tree use by beef cattle as a function of black globe temperature and time of day.

PubMed

Foust, Amanda M; Headlee, William L

2017-12-01

Increasing temperatures associated with global climate change threaten to disrupt agricultural systems such as beef production, yet relatively little is known about the use of natural tree shade to mitigate the negative effects of heat stress on beef cattle. In this study, we evaluated how temperature and time of day influenced the utilization of tree shade in relation to coloration, orientation, and behavior of beef cattle in a pasture system. Temperatures in shade and direct sunlight were measured using black globe temperature (BGT) data loggers. Time-lapse images from game cameras were used to obtain counts of shade usage, coloration, orientation, and behavior of cattle throughout the daytime hours. In general, we found that shade utilization and most of the predominating orientations and behaviors differed significantly (P < 0.05) by both time of day (Hour) and BGT in direct sunlight (BGT sun ), while interactions between these two effects (Hour × BGT sun ) were often nonsignificant. The mean percentage of the herd using shade was highest in mid-morning (87-96%) and early afternoon (97%), but also increased with BGT sun regardless of the time of day; these trends were similar for both dark- and light-colored cattle. Lying down was the dominant behavior exhibited in the shade, while foraging was the most prevalent behavior in the sun. When herd shade usage was lowest in mid- to late-afternoon (<1%) we also observed an increase in the use of heat-mitigating orientations in the sun (37-47%). We discuss some practical implications of these results, including the potential use of temperature thresholds to interpret cattle behaviors and shade usage.

Real-time Supervised Detection of Pink Areas in Dermoscopic Images of Melanoma: Importance of Color Shades, Texture and Location

PubMed Central

Kaur, Ravneet; Albano, Peter P.; Cole, Justin G.; Hagerty, Jason; LeAnder, Robert W.; Moss, Randy H.; Stoecker, William V.

2015-01-01

Background/Purpose Early detection of malignant melanoma is an important public health challenge. In the USA, dermatologists are seeing more melanomas at an early stage, before classic melanoma features have become apparent. Pink color is a feature of these early melanomas. If rapid and accurate automatic detection of pink color in these melanomas could be accomplished, there could be significant public health benefits. Methods Detection of three shades of pink (light pink, dark pink, and orange pink) was accomplished using color analysis techniques in five color planes (red, green, blue, hue and saturation). Color shade analysis was performed using a logistic regression model trained with an image set of 60 dermoscopic images of melanoma that contained pink areas. Detected pink shade areas were further analyzed with regard to the location within the lesion, average color parameters over the detected areas, and histogram texture features. Results Logistic regression analysis of a separate set of 128 melanomas and 128 benign images resulted in up to 87.9% accuracy in discriminating melanoma from benign lesions measured using area under the receiver operating characteristic curve. The accuracy in this model decreased when parameters for individual shades, texture, or shade location within the lesion were omitted. Conclusion Texture, color, and lesion location analysis applied to multiple shades of pink can assist in melanoma detection. When any of these three details: color location, shade analysis, or texture analysis were omitted from the model, accuracy in separating melanoma from benign lesions was lowered. Separation of colors into shades and further details that enhance the characterization of these color shades are needed for optimal discrimination of melanoma from benign lesions. PMID:25809473

Accurate shade image matching by using a smartphone camera.

PubMed

Tam, Weng-Kong; Lee, Hsi-Jian

2017-04-01

Dental shade matching by using digital images may be feasible when suitable color features are properly manipulated. Separating the color features into feature spaces facilitates favorable matching. We propose using support vector machines (SVM), which are outstanding classifiers, in shade classification. A total of 1300 shade tab images were captured using a smartphone camera with auto-mode settings and no flash. The images were shot at angled distances of 14-20cm from a shade guide at a clinic equipped with light tubes that produced a 4000K color temperature. The Group 1 samples comprised 1040 tab images, for which the shade guide was randomly positioned in the clinic, and the Group 2 samples comprised 260 tab images, for which the shade guide had a fixed position in the clinic. Rectangular content was cropped manually on each shade tab image and further divided into 10×2 blocks. The color features extracted from the blocks were described using a feature vector. The feature vectors in each group underwent SVM training and classification by using the "leave-one-out" strategy. The top one and three accuracies of Group 1 were 0.86 and 0.98, respectively, and those of Group 2 were 0.97 and 1.00, respectively. This study provides a feasible technique for dental shade classification that uses the camera of a mobile device. The findings reveal that the proposed SVM classification might outperform the shade-matching results of previous studies that have performed similarity measurements of ΔE levels or used an S, a*, b* feature set. Copyright © 2016 Japan Prosthodontic Society. Published by Elsevier Ltd. All rights reserved.

Shade tree diversity and aboveground carbon stocks in Theobroma cacao agroforestry systems: implications for REDD+ implementation in a West African cacao landscape.

PubMed

Dawoe, Evans; Asante, Winston; Acheampong, Emmanuel; Bosu, Paul

2016-12-01

The promotion of cacao agroforestry is one of the ways of diversifying farmer income and creating incentives through their inclusion in REDD+ interventions. We estimated the aboveground carbon stocks in cacao and shade trees, determined the floristic diversity of shade trees and explored the possibility of implementing REDD+ interventions in cacao landscapes. Using replicated multi-site transect approach, data were collected from nine 1-ha plots established on 5 km long transects in ten cacao growing districts in Ghana West Africa. Biomass of cacao and shade trees was determined using allometric equations. One thousand four hundred and one (1401) shade trees comprising 109 species from 33 families were recorded. Total number of species ranged from 34 to 49. Newbouldia laevis (Bignoniacea) was the most frequently occurring specie and constituted 43.2 % of all shade trees. The most predominant families were Sterculiaceae and Moraceae (10 species each), followed by Meliaceae and Mimosaceae (8 species each) and Caesalpiniacaea (6 species). Shannon diversity indices (H', H max and J') and species richness were low compared to other similar studies. Shade tree densities ranged from 16.2 ± 3.0 to 22.8 ± 1.7 stems ha -1 and differed significantly between sites. Carbon stocks of shade trees differed between sites but were similar in cacao trees. The average C stock in cacao trees was 7.45 ± 0.41 Mg C ha -1 compared with 8.32 ± 1.15 Mg C ha -1 in the shade trees. Cacao landscapes in Ghana have the potential of contributing to forest carbon stocks enhancement by increasing the stocking density of shade trees to recommended levels.

Real-time supervised detection of pink areas in dermoscopic images of melanoma: importance of color shades, texture and location.

PubMed

Kaur, R; Albano, P P; Cole, J G; Hagerty, J; LeAnder, R W; Moss, R H; Stoecker, W V

2015-11-01

Early detection of malignant melanoma is an important public health challenge. In the USA, dermatologists are seeing more melanomas at an early stage, before classic melanoma features have become apparent. Pink color is a feature of these early melanomas. If rapid and accurate automatic detection of pink color in these melanomas could be accomplished, there could be significant public health benefits. Detection of three shades of pink (light pink, dark pink, and orange pink) was accomplished using color analysis techniques in five color planes (red, green, blue, hue, and saturation). Color shade analysis was performed using a logistic regression model trained with an image set of 60 dermoscopic images of melanoma that contained pink areas. Detected pink shade areas were further analyzed with regard to the location within the lesion, average color parameters over the detected areas, and histogram texture features. Logistic regression analysis of a separate set of 128 melanomas and 128 benign images resulted in up to 87.9% accuracy in discriminating melanoma from benign lesions measured using area under the receiver operating characteristic curve. The accuracy in this model decreased when parameters for individual shades, texture, or shade location within the lesion were omitted. Texture, color, and lesion location analysis applied to multiple shades of pink can assist in melanoma detection. When any of these three details: color location, shade analysis, or texture analysis were omitted from the model, accuracy in separating melanoma from benign lesions was lowered. Separation of colors into shades and further details that enhance the characterization of these color shades are needed for optimal discrimination of melanoma from benign lesions. © 2015 John Wiley & Sons A/S. Published by John Wiley & Sons Ltd.

A Custom Made Intrinsic Silicone Shade Guide for Indian Population

PubMed Central

Behanam, Mohammed; Ahila, S.C.; Jei, J. Brintha

2016-01-01

Introduction Replication of natural skin colour in maxillofacial prosthesis has been traditionally done using trial and error method, as concrete shade guides are unavailable till date. Hence a novel custom made intrinsic silicone shade guide has been attempted for Indian population. Aim Reconstruction of maxillofacial defects is challenging, as achieving an aesthetic result is not always easy. A concoction of a novel intrinsic silicone shade guide was contemplated for the study and its reproducibility in clinical practice was analysed. Materials and Methods Medical grade room temperature vulcanising silicone was used for the fabrication of shade tabs. The shade guide consisted of three main groups I, II and III which were divided based upon the hues yellow, red and blue respectively. Five distinct intrinsic pigments were added in definite proportions to subdivide each group of different values from lighter to darker shades. A total number of 15 circular shade tabs comprised the guide. To validate the usage of the guide, visual assessment of colour matching was done by four investigators to investigate the consent of perfect colour correspondence. Data was statistically analysed using kappa coefficients. Results The kappa values were found to be 0.47 to 0.78 for yellow based group I, 0.13 to 0.65 for red based group II, and 0.07 to 0.36 for blue based group III. This revealed that the shade tabs of yellow and red based hues matched well and showed a statistically good colour matching. Conclusion This intrinsic silicone shade guide can be effectively utilised for fabrication of maxillofacial prosthesis with silicone in Indian population. A transparent colour formula with definite proportioning of intrinsic pigments is provided for obtaining an aesthetic match to skin tone. PMID:27190946

VALIDATING the Accuracy of Sighten's Automated Shading Tool

DOE Office of Scientific and Technical Information (OSTI.GOV)

Solar companies - including installers, financiers, and distributors - leverage Sighten software to deliver accurate shading calculations and solar proposals. Sighten recently partnered with Google Project Sunroof to provide automated remote shading analysis directly within the Sighten platform. The National Renewable Energy Laboratory (NREL), in partnership with Sighten, independently verified the accuracy of Sighten's remote-shading solar access values (SAVs) on an annual basis for locations in Los Angeles, California, and Denver, Colorado.

An Appraisal of the Classic Forest Succession Paradigm with the Shade Tolerance Index

PubMed Central

Lienard, Jean; Florescu, Ionut; Strigul, Nikolay

2015-01-01

In this paper we revisit the classic theory of forest succession that relates shade tolerance and species replacement and assess its validity to understand patch-mosaic patterns of forested ecosystems of the USA. We introduce a macroscopic parameter called the “shade tolerance index” and compare it to the classic continuum index in southern Wisconsin forests. We exemplify shade tolerance driven succession in White Pine-Eastern Hemlock forests using computer simulations and analyzing approximated chronosequence data from the USDA FIA forest inventory. We describe this parameter across the last 50 years in the ecoregions of mainland USA, and demonstrate that it does not correlate with the usual macroscopic characteristics of stand age, biomass, basal area, and biodiversity measures. We characterize the dynamics of shade tolerance index using transition matrices and delimit geographical areas based on the relevance of shade tolerance to explain forest succession. We conclude that shade tolerance driven succession is linked to climatic variables and can be considered as a primary driving factor of forest dynamics mostly in central-north and northeastern areas in the USA. Overall, the shade tolerance index constitutes a new quantitative approach that can be used to understand and predict succession of forested ecosystems and biogeographic patterns. PMID:25658092

Interior shadings for office indoor visual comfort in humid climate region

NASA Astrophysics Data System (ADS)

Dinapradipta, Asri; Sudarma, Erwin; Defiana, Ima; Erwindi, Collinthia

2018-03-01

As part of the fenestration system, the interior shadings have also a role to control the indoor environment to maintain indoor visual comfort. As the occupants have personal access to control these, their control behavior then, might enhance or even worsen indoor comfort performance. The controlling behavior might not only influence indoor comfort performance but can also indicate the success or failure of interior shading as a control device element. This paper is intended to report control behavior patterns, as represented by the variety of the slats’ openings of two types of interior shading i.e. Venetian and Vertical blinds and to analyze these on the concurrent impacts to indoor office building’s indoor illuminance and luminance distribution. The purpose of this research is to figure out the shading control patterns as well as to examine the effectiveness of these two types of interior shadings to control indoor visual environment. This study is a quantitative research using experimentation on the slats’ opening of two types of shadings at two identical office rooms. The research results suggested that both types of blinds seem unsuitable for gaining proper illumination values at work planes in humid tropics area. However, these shadings demonstrate good performance for luminance distribution except for that of the closed Venetian blinds.

Optimization and Comparison of Different Digital Mammographic Tomosynthesis Reconstruction Methods

DTIC Science & Technology

2007-04-01

physical measurements of impulse response analysis, modulation transfer function (MTF) and noise power spectrum (NPS). (Months 5- 12). 1.2.1. Simulate...added: projection images with simulated impulse and the 1/r2 shading difference. Other system blur and noise issues were not addressed in this paper...spectrum (NPS), Noise -equivalent quanta (NEQ), impulse response, Back Projection (BP) 1. INTRODUCTION Digital breast tomosynthesis is a new

Light accelerates plant responses to warming.

PubMed

De Frenne, Pieter; Rodríguez-Sánchez, Francisco; De Schrijver, An; Coomes, David A; Hermy, Martin; Vangansbeke, Pieter; Verheyen, Kris

2015-08-17

Competition for light has profound effects on plant performance in virtually all terrestrial ecosystems. Nowhere is this more evident than in forests, where trees create environmental heterogeneity that shapes the dynamics of forest-floor communities(1-3). Observational evidence suggests that biotic responses to both anthropogenic global warming and nitrogen pollution may be attenuated by the shading effects of trees and shrubs(4-9). Here we show experimentally that tree shade is slowing down changes in below-canopy communities due to warming. We manipulated levels of photosynthetically active radiation, temperature and nitrogen, alone and in combination, in a temperate forest understorey over a 3-year period, and monitored the composition of the understorey community. Light addition, but not nitrogen enrichment, accelerated directional plant community responses to warming, increasing the dominance of warmth-preferring taxa over cold-tolerant plants (a process described as thermophilization(6,10-12)). Tall, competitive plants took greatest advantage of the combination of elevated temperature and light. Warming of the forest floor did not result in strong community thermophilization unless light was also increased. Our findings suggest that the maintenance of locally closed canopy conditions could reduce, at least temporarily, warming-induced changes in forest floor plant communities.

Shading Vita In-ceram YZ substructures: influence on value and chroma, part II.

PubMed

Devigus, A; Lombardi, G

2004-10-01

In this study, the influence of differently shaded substructures made of Y-TZP from different manufacturers (without, with 0.5 mm or 1.0 mm ceramic veneer with Vita VM9 Base Dentin) on the lightness, the saturation, and the color shade (= value, chroma, and hue) was measured and assessed with a spectral photometer (Easy Shade, Vita) on a clinical case in the mouth and on models of different color. Shading substructures made of Y-TZP facilitated adaptation to the basic shade and reduced the layer thickness of the veneer ceramic required to achieve the desired color. This should be done in the future as a matter of routine. In this way, tooth substance can be better conserved when preparing the teeth, and the esthetic result is not diminished. The material used for the production of the working models (plaster or plastic) should be tooth colored and in terms of lightness--analogous to the shading of the substructure--in order to facilitate clinical color measurement and quality control in the laboratory.

The effects of shade on performance, carcass classes and behaviour of heat-stressed feedlot cattle at the finisher phase.

PubMed

Blaine, K L; Nsahlai, Ignatius Verla

2011-03-01

The study aimed to determine the impact of shade on the performance, carcass class and behaviour using 146 crossbred steers and bulls during the finishing phase on a commercial feedlot in February. Treatments were (1) shade and (2) no shade. Average daily gain (ADG), DMI, and feed efficiency were measured, and hot carcass weight (HCW) and grades were collected at slaughter. The proportion of animals within a pen engaged in various behavioural activities (standing, lying, feeding and panting) was recorded. Five randomly selected individual animals per treatment were monitored every 2 min between 0700 and 1600 hours to determine the time spent on each of the above activities. Shade improved the final body weight (P < 0.0001), ADG (P = 0.079), and HCW by 8.33 kg (P < 0.05). Shade increased (P < 0.05) the feeding activity but decreased (P < 0.05) panting behaviour. Shade conferred an economic benefit of R2.03 head(-1)week(-1), and thus would reduce heat stress and increase the feedlot profitability.

Improved color matching of metal ceramic restorations. Part II: Procedures for visual communication.

PubMed

Sorensen, J A; Torres, T J

1987-12-01

Most ceramic restorations are fabricated in a location remote from the dental office. Successful fabrication of matching life-like ceramic restorations necessitates a collaborative effort between the dentist and the ceramist. To meet the demands for visual communication of shade and surface texture, the following steps are recommended. 1. A means of communicating and recording surface texture that facilitates blending the restorations with the natural dentition should be used. 2. The system should use an esthetics prescription form that functions with the Shade Indicator Chart system to relate the shade of opaque, body, and incisal porcelains and their arrangement to the ceramist. 3. An easily made identification mold to form shade tabs is needed. 4. Identification shade tabs should be made to verify and document shade formulations selected with the Shade Indicator Chart system. 5. Methods for precisely mapping and reproducing individual characterization patterns are needed. This information permits the visualization of the end result, allowing the artistic expression of the ceramist to create vital-appearing restorations intrinsically and in harmony with the natural dentition.

Camp Unit Design Guidelines.

ERIC Educational Resources Information Center

Hultsman, John T.; Cottrell, Richard L.

This document provides a set of generalized guidelines for the design of units in large family campgrounds. Managers of recreational lands have two responsibilities and goals: to protect the natural resources, and to provide an enjoyable experience for users. With these goals in mind, unique variables to each unit such as shade, site aesthetics,…

Seasonal flight activity and distribution of metallic woodboring beetles (Coleoptera: Buprestidae) collected in North Carolina and Tennessee

USDA-ARS?s Scientific Manuscript database

Metallic wood boring insects (Coleoptera: Buprestidae) species are responsible for high levels of host plant injury to deciduous shade and flowering trees in commercial nurseries, urban forests, and managed landscapes. Ornamental plant producers in the southeastern U.S. have ranked borers, includin...

Compositional changes in 'Bartlett' pear ( Pyrus communis L.) cell wall polysaccharides as affected by sunlight conditions.

PubMed

Raffo, María D; Ponce, Nora M A; Sozzi, Gabriel O; Vicente, Ariel R; Stortz, Carlos A

2011-11-23

Preharvest conditions can have a great impact on fruit quality attributes and postharvest responses. Firmness is an important quality attribute in pear, and excessive softening increases susceptibility to bruising and decay, thus limiting fruit postharvest life. Textural characteristics of fruits are determined at least in part by cell wall structure and disassembly. Few studies have analyzed the influence of fruit preharvest environment in softening, cell wall composition, and degradation. In the current work 'Bartlett' pears grown either facing the sun (S) or in the shade (H) were harvested and stored for 13 days at 20 °C. An evaluation of fruit soluble solids, acidity, color, starch degradation, firmness, cell wall yield, pectin and matrix glycan solubilization, depolymerization, and monosaccharide composition was carried out. Sun-exposed pears showed more advanced color development and similar levels of starch degradation, sugars, and acids than shaded fruit. Sunlight-grown pears were at harvest firmer than shade-grown pears. Both fruit groups softened during storage at 20 °C, but even after ripening, sun-exposed pears remained firmer. Sunlight exposure did not have a great impact on pectin molecular weight. Instead, at harvest a higher proportion of water-solubilized uronic acids and alkali-solubilized neutral sugars and a larger mean molecular size of tightly bound glycans was found in sun-exposed pears. During ripening cell wall catabolism took place in both sun- and shade-grown pears, but pectin solubilization was clearly delayed in sun-exposed fruit. This was associated with decreased removal of RG I-arabinan side chains rather than with reduced depolymerization.

Photosynthesis, photoprotection, and growth of shade-tolerant tropical tree seedlings under full sunlight.

PubMed

Krause, G Heinrich; Winter, Klaus; Matsubara, Shizue; Krause, Barbara; Jahns, Peter; Virgo, Aurelio; Aranda, Jorge; García, Milton

2012-09-01

High solar radiation in the tropics is known to cause transient reduction in photosystem II (PSII) efficiency and CO(2) assimilation in sun-exposed leaves, but little is known how these responses affect the actual growth performance of tropical plants. The present study addresses this question. Seedlings of five woody neotropical forest species were cultivated under full sunlight and shaded conditions. In full sunlight, strong photoinhibition of PSII at midday was documented for the late-successional tree species Ormosia macrocalyx and Tetragastris panamensis and the understory/forest gap species, Piper reticulatum. In leaves of O. macrocalyx, PSII inhibition was accompanied by substantial midday depression of net CO(2) assimilation. Leaves of all species had increased pools of violaxanthin-cycle pigments. Other features of photoacclimation, such as increased Chl a/b ratio and contents of lutein, β-carotene and tocopherol varied. High light caused strong increase of tocopherol in leaves of T. panamensis and another late-successional species, Virola surinamensis. O. macrocalyx had low contents of tocopherol and UV-absorbing substances. Under full sunlight, biomass accumulation was not reduced in seedlings of T. panamensis, P. reticulatum, and V. surinamensis, but O. macrocalyx exhibited substantial growth inhibition. In the highly shade-tolerant understory species Psychotria marginata, full sunlight caused strongly reduced growth of most individuals. However, some plants showed relatively high growth rates under full sun approaching those of seedlings at 40 % ambient irradiance. It is concluded that shade-tolerant tropical tree seedlings can achieve efficient photoacclimation and high growth rates in full sunlight.

Tradeoffs between income, biodiversity, and ecosystem functioning during tropical rainforest conversion and agroforestry intensification

PubMed Central

Steffan-Dewenter, Ingolf; Kessler, Michael; Barkmann, Jan; Bos, Merijn M.; Buchori, Damayanti; Erasmi, Stefan; Faust, Heiko; Gerold, Gerhard; Glenk, Klaus; Gradstein, S. Robbert; Guhardja, Edi; Harteveld, Marieke; Hertel, Dietrich; Höhn, Patrick; Kappas, Martin; Köhler, Stefan; Leuschner, Christoph; Maertens, Miet; Marggraf, Rainer; Migge-Kleian, Sonja; Mogea, Johanis; Pitopang, Ramadhaniel; Schaefer, Matthias; Schwarze, Stefan; Sporn, Simone G.; Steingrebe, Andrea; Tjitrosoedirdjo, Sri S.; Tjitrosoemito, Soekisman; Twele, André; Weber, Robert; Woltmann, Lars; Zeller, Manfred; Tscharntke, Teja

2007-01-01

Losses of biodiversity and ecosystem functioning due to rainforest destruction and agricultural intensification are prime concerns for science and society alike. Potentially, ecosystems show nonlinear responses to land-use intensification that would open management options with limited ecological losses but satisfying economic gains. However, multidisciplinary studies to quantify ecological losses and socioeconomic tradeoffs under different management options are rare. Here, we evaluate opposing land use strategies in cacao agroforestry in Sulawesi, Indonesia, by using data on species richness of nine plant and animal taxa, six related ecosystem functions, and on socioeconomic drivers of agroforestry expansion. Expansion of cacao cultivation by 230% in the last two decades was triggered not only by economic market mechanisms, but also by rarely considered cultural factors. Transformation from near-primary forest to agroforestry had little effect on overall species richness, but reduced plant biomass and carbon storage by ≈75% and species richness of forest-using species by ≈60%. In contrast, increased land use intensity in cacao agroforestry, coupled with a reduction in shade tree cover from 80% to 40%, caused only minor quantitative changes in biodiversity and maintained high levels of ecosystem functioning while doubling farmers' net income. However, unshaded systems further increased income by ≈40%, implying that current economic incentives and cultural preferences for new intensification practices put shaded systems at risk. We conclude that low-shade agroforestry provides the best available compromise between economic forces and ecological needs. Certification schemes for shade-grown crops may provide a market-based mechanism to slow down current intensification trends. PMID:17360392

Tradeoffs between income, biodiversity, and ecosystem functioning during tropical rainforest conversion and agroforestry intensification.

PubMed

Steffan-Dewenter, Ingolf; Kessler, Michael; Barkmann, Jan; Bos, Merijn M; Buchori, Damayanti; Erasmi, Stefan; Faust, Heiko; Gerold, Gerhard; Glenk, Klaus; Gradstein, S Robbert; Guhardja, Edi; Harteveld, Marieke; Hertel, Dietrich; Höhn, Patrick; Kappas, Martin; Köhler, Stefan; Leuschner, Christoph; Maertens, Miet; Marggraf, Rainer; Migge-Kleian, Sonja; Mogea, Johanis; Pitopang, Ramadhaniel; Schaefer, Matthias; Schwarze, Stefan; Sporn, Simone G; Steingrebe, Andrea; Tjitrosoedirdjo, Sri S; Tjitrosoemito, Soekisman; Twele, André; Weber, Robert; Woltmann, Lars; Zeller, Manfred; Tscharntke, Teja

2007-03-20

Losses of biodiversity and ecosystem functioning due to rainforest destruction and agricultural intensification are prime concerns for science and society alike. Potentially, ecosystems show nonlinear responses to land-use intensification that would open management options with limited ecological losses but satisfying economic gains. However, multidisciplinary studies to quantify ecological losses and socioeconomic tradeoffs under different management options are rare. Here, we evaluate opposing land use strategies in cacao agroforestry in Sulawesi, Indonesia, by using data on species richness of nine plant and animal taxa, six related ecosystem functions, and on socioeconomic drivers of agroforestry expansion. Expansion of cacao cultivation by 230% in the last two decades was triggered not only by economic market mechanisms, but also by rarely considered cultural factors. Transformation from near-primary forest to agroforestry had little effect on overall species richness, but reduced plant biomass and carbon storage by approximately 75% and species richness of forest-using species by approximately 60%. In contrast, increased land use intensity in cacao agroforestry, coupled with a reduction in shade tree cover from 80% to 40%, caused only minor quantitative changes in biodiversity and maintained high levels of ecosystem functioning while doubling farmers' net income. However, unshaded systems further increased income by approximately 40%, implying that current economic incentives and cultural preferences for new intensification practices put shaded systems at risk. We conclude that low-shade agroforestry provides the best available compromise between economic forces and ecological needs. Certification schemes for shade-grown crops may provide a market-based mechanism to slow down current intensification trends.

Design of a radiator shade for testing in a simulated lunar environment

NASA Technical Reports Server (NTRS)

Huff, Jaimi; Remington, Randy; Tang, Toan

1992-01-01

The National Aeronautics and Space Administration (NASA) and The Universities Space Research Association (USRA) have chosen the parabolic/catenary concept from their sponsored Fall 1991 lunar radiation shade project for further testing and development. NASA asked the design team to build a shading device and support structure for testing in a vacuum chamber. Besides the support structure for the catenary shading device, the design team was asked to develop a system for varying the shade shape so that the device can be tested at different focal lengths. The design team developed concept variants and combined the concept variants to form overall designs. Using a decision matrix, an overall design was selected by the team from several overall design alternatives. Concept variants were developed for three primary functions. The three functions were structural support, shape adjustments, and end shielding. The shade adjustment function was divided into two sub-functions, arc length adjustment, and width adjustment.

3D Shape Perception in Posterior Cortical Atrophy: A Visual Neuroscience Perspective

PubMed Central

Gillebert, Céline R.; Schaeverbeke, Jolien; Bastin, Christine; Neyens, Veerle; Bruffaerts, Rose; De Weer, An-Sofie; Seghers, Alexandra; Sunaert, Stefan; Van Laere, Koen; Versijpt, Jan; Vandenbulcke, Mathieu; Salmon, Eric; Todd, James T.; Orban, Guy A.

2015-01-01

Posterior cortical atrophy (PCA) is a rare focal neurodegenerative syndrome characterized by progressive visuoperceptual and visuospatial deficits, most often due to atypical Alzheimer's disease (AD). We applied insights from basic visual neuroscience to analyze 3D shape perception in humans affected by PCA. Thirteen PCA patients and 30 matched healthy controls participated, together with two patient control groups with diffuse Lewy body dementia (DLBD) and an amnestic-dominant phenotype of AD, respectively. The hierarchical study design consisted of 3D shape processing for 4 cues (shading, motion, texture, and binocular disparity) with corresponding 2D and elementary feature extraction control conditions. PCA and DLBD exhibited severe 3D shape-processing deficits and AD to a lesser degree. In PCA, deficient 3D shape-from-shading was associated with volume loss in the right posterior inferior temporal cortex. This region coincided with a region of functional activation during 3D shape-from-shading in healthy controls. In PCA patients who performed the same fMRI paradigm, response amplitude during 3D shape-from-shading was reduced in this region. Gray matter volume in this region also correlated with 3D shape-from-shading in AD. 3D shape-from-disparity in PCA was associated with volume loss slightly more anteriorly in posterior inferior temporal cortex as well as in ventral premotor cortex. The findings in right posterior inferior temporal cortex and right premotor cortex are consistent with neurophysiologically based models of the functional anatomy of 3D shape processing. However, in DLBD, 3D shape deficits rely on mechanisms distinct from inferior temporal structural integrity. SIGNIFICANCE STATEMENT Posterior cortical atrophy (PCA) is a neurodegenerative syndrome characterized by progressive visuoperceptual dysfunction and most often an atypical presentation of Alzheimer's disease (AD) affecting the ventral and dorsal visual streams rather than the medial temporal system. We applied insights from fundamental visual neuroscience to analyze 3D shape perception in PCA. 3D shape-processing deficits were affected beyond what could be accounted for by lower-order processing deficits. For shading and disparity, this was related to volume loss in regions previously implicated in 3D shape processing in the intact human and nonhuman primate brain. Typical amnestic-dominant AD patients also exhibited 3D shape deficits. Advanced visual neuroscience provides insight into the pathogenesis of PCA that also bears relevance for vision in typical AD. PMID:26377458

In vitro and in vivo evaluations of three computer-aided shade matching instruments.

PubMed

Yuan, Kun; Sun, Xiang; Wang, Fu; Wang, Hui; Chen, Ji-hua

2012-01-01

This study evaluated the accuracy and reliability of three computer-aided shade matching instruments (Shadepilot, VITA Easyshade, and ShadeEye NCC) using both in vitro and in vivo models. The in vitro model included the measurement of five VITA Classical shade guides. The in vivo model utilized three instruments to measure the central region of the labial surface of maxillary right central incisors of 85 people. The accuracy and reliability of the three instruments in these two evaluating models were calculated. Significant differences were observed in the accuracy of instruments both in vitro and in vivo. No significant differences were found in the reliability of instruments between and within the in vitro and the in vivo groups. VITA Easyshade was significantly different in accuracy between in vitro and in vivo models, while no significant difference was found for the other two instruments. Shadepilot was the only instrument tested in the present study that showed high accuracy and reliability both in vitro and in vivo. Significant differences were observed in the L*a*b* values of the 85 natural teeth measured using three instruments in the in vivo assessment. The pair-agreement rates of shade matching among the three instruments ranged from 37.7% to 48.2%, and the incidence of identical shade results shared by all three instruments was 25.9%. As different L*a*b* values and shade matching results were reported for the same tooth, a combination of the evaluated shade matching instruments and visual shade confirmation is recommended for clinical use.

A field measure of the shade fraction

NASA Technical Reports Server (NTRS)

Gillespie, Alan R.; Smith, Milton O.; Sabol, Donald E.

1992-01-01

'Shade' has a technical definition peculiar to linear spectral mixture analysis of imaging spectrometer data: it is the reduction in radiance from a surface due to lighting conditions and geometry, and includes topographic shading described by photometric functions as well as shadowing at all scales. 'Shade' is an important constituent of nearly all remotely sensed images, and is one endmember resolved in spectral mixture analysis, where it is represented as a fraction of the measured radiance and a characteristic spectrum. This spectrum is typically the null vector, provided the data have been corrected for atmospheric and instrument effects: i.e., 'shade' is the radiance from an ideal black surface. In topographic shading, irradiance is reduced - typically in proportion to cos(i), where i (incidence angle) is the angle between the sun and the local surface normal vectors. Therefore, the radiance is lowered by a multiplicative factor. Shadowing occurs when i is greater than 90 deg, or when sunlight is blocked by adjacent high terrain; the only irradiance is down-welling skylight and bounce light from adjacent terrain. In spectral mixture analysis, 'shade' is regarded as an additive term. In this regard, it is an accurate description of the proportion of a scene that consists of ideal shadows ('checkerboard mixing'); however, 'shade' represents the multiplicative cos(i) factor as well, as here it should be interpreted as the proportion of shadow that would darken the scene an equivalent amount. In either case, the 'shade' fraction is lessened by adjacency effects, because the scene has a non-zero reflectivity instead of the ideal black surface generally assumed.

Seedling transplants reveal species-specific responses of high-elevation tropical treeline trees to climate change.

PubMed

Rehm, Evan M; Feeley, Kenneth J

2016-08-01

The elevations at which tropical treelines occur are believed to represent the point where low mean temperatures limit the growth of upright woody trees. Consequently, tropical treelines are predicted to shift to higher elevations with global warming. However, treelines throughout the tropics have remained stationary despite increasing global mean temperatures. The goal of the study reported here was to build a more comprehensive understanding of the effects of mean temperature, low-temperature extremes, shading, and their interactions on seedling survival at tropical treelines. We conducted a seedling transplant study using three dominant canopy-forming treeline species in the southern tropical Andes. We found species-specific differences and contrasting responses in seedling survival to changes in mean temperature. The most abundant naturally occurring species at the seedling stage outside the treeline, Weinmannia fagaroides, showed a negative relationship between the survival of transplanted seedlings and mean temperature, the opposite of a priori expectations. Conversely, Clethra cuneata showed increased survival at higher mean temperatures, but survival also increased with higher absolute low temperatures and the presence of shade. Finally, the survival of Gynoxys nitida seedlings was insensitive to temperature but increased under shade. These findings show that multiple factors can determine the upper distributional limit of species forming the current tropical treeline. As such, predictions of future local and regional tropical treeline shifts may need to consider several factors beyond changes in mean temperature. If the treeline remains stationary and cloud forests are unable to expand into higher elevations, there may be severe species loss in this biodiversity hotspot.

Impacts of sediments on coral energetics: partitioning the effects of turbidity and settling particles.

PubMed

Junjie, Reef K; Browne, Nicola K; Erftemeijer, Paul L A; Todd, Peter A

2014-01-01

Sediment loads have long been known to be deleterious to corals, but the effects of turbidity and settling particles have not previously been partitioned. This study provides a novel approach using inert silicon carbide powder to partition and quantify the mechanical effects of sediment settling versus reduced light under a chronically high sedimentary regime on two turbid water corals commonly found in Singapore (Galaxea fascicularis and Goniopora somaliensis). Coral fragments were evenly distributed among three treatments: an open control (30% ambient PAR), a shaded control (15% ambient PAR) and sediment treatment (15% ambient PAR; 26.4 mg cm(-2) day(-1)). The rate of photosynthesis and respiration, and the dark-adapted quantum yield were measured once a week for four weeks. By week four, the photosynthesis to respiration ratio (P/R ratio) and the photosynthetic yield (Fv/Fm) had fallen by 14% and 3-17% respectively in the shaded control, contrasting with corals exposed to sediments whose P/R ratio and yield had declined by 21% and 18-34% respectively. The differences in rates between the shaded control and the sediment treatment were attributed to the mechanical effects of sediment deposition. The physiological response to sediment stress differed between species with G. fascicularis experiencing a greater decline in the net photosynthetic yield (13%) than G. somaliensis (9.5%), but a smaller increase in the respiration rates (G. fascicularis = 9.9%, G. somaliensis  = 14.2%). These different physiological responses were attributed, in part, to coral morphology and highlighted key physiological processes that drive species distribution along high to low turbidity and depositional gradients.

Behavioral thermoregulation in a group of zoo-housed colobus monkeys (Colobus guereza).

PubMed

Wark, Jason D; Kuhar, Christopher W; Lukas, Kristen E

2014-01-01

Although wild primates are known to modify behavior in response to thermal stress, less is known about behavioral thermoregulation in zoo-housed primates. Zoo exhibits expose individuals to unique thermal environments and may constrain the thermoregulatory strategies available to individual animals. In this study, we observed a group of seven colobus monkeys (Colobus guereza) living on a concrete "Monkey Island" style exhibit that featured limited shade and limited arboreal space. Behaviors were recorded using continuous focal animal sampling (n = 63 days, 97.7 hr). Logistic regression revealed 23°C was the temperature at which monkeys began resting more in shade than in sun. When temperatures exceeded 23°C, animals spent more time in open sitting postures with limbs extended from the body; sat less frequently in closed, hunched postures; spent more time in social contact; and performed more self-directed behaviors. Exhibit use also shifted under higher temperatures, with more time spent in areas with shade and lower surface temperatures. Lastly, when provided with access to an indoor holding area, the colobus monkeys spent more than half the time indoors when temperatures exceeded 23°C, yet only 10% of their time indoors when the temperature was below this value. Although postural changes have been reported in wild colobus, the postural and other behavioral changes observed in the current study occurred at temperatures lower than expected based on the published thermoneutral zone of colobus monkeys and highlight the importance of considering the specific thermoregulatory responses of zoo animals. © 2014 Wiley Periodicals, Inc.

Leaf physiology and biomass allocation of backcross hybrid American chestnut (Castanea dentata) seedlings in response to light and water availability.

PubMed

Brown, Caleb E; Mickelbart, Michael V; Jacobs, Douglass F

2014-12-01

Partial canopy cover promotes regeneration of many temperate forest trees, but the consequences of shading on seedling drought resistance are unclear. Reintroduction of blight-resistant American chestnut (Castanea dentata (Marsh.) Borkh.) into eastern North American forests will often occur on water-limited sites and under partial canopy cover. We measured leaf pre-dawn water potential (Ψpd), leaf gas exchange, and growth and biomass allocation of backcross hybrid American chestnut seedlings from three orchard sources grown under different light intensities (76, 26 and 8% full photosynthetically active radiation (PAR)) and subjected to well-watered or mid-season water-stressed conditions. Seedlings in the water-stress treatment were returned to well-watered conditions after wilting to examine recovery. Seedlings growing under medium- and high-light conditions wilted at lower leaf Ψpd than low-light seedlings. Recovery of net photosynthesis (Anet) and stomatal conductance (gs) was greater in low and medium light than in high light. Seed source did not affect the response to water stress or light level in most cases. Between 26 and 8% full PAR, light became limiting to the extent that the effects of water stress had no impact on some growth and morphological traits. We conclude that positive and negative aspects of shading on seedling drought tolerance and recovery are not mutually exclusive. Partial shade may help American chestnut tolerate drought during early establishment through effects on physiological conditioning. © The Author 2014. Published by Oxford University Press. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

Clinical evaluation of a new 10% carbamide peroxide tooth-whitening agent.

PubMed

Barnes, D M; Kihn, P W; Romberg, E; George, D; DePaola, L; Medina, E

1998-10-01

NUPRO Gold Tooth Whitening System was evaluated for efficacy according to the proposed ADA guidelines for acceptance. Sixty participants with discolored anterior teeth participated in a 14-day, double-blind, clinical trial. The participants were matched for age, gender, and oral health status and were given either a placebo gel without the active agent or the NUPRO Gold active gel, which they wore in a custom-fabricated mouth guard for home use. The shade of each participant's maxillary anterior teeth was evaluated using a value-oriented Vita Lumin Vacuum Shade Guide before the study. The same shade guide was used to determine shade changes. Time of use of the agent and potential side effects, such as tooth and gingival hypersensitivity and tissue irritation, were assessed at all recall examinations and were recorded by participants in daily diaries. The average shade change for the placebo users was less than one shade. The average shade change for the NUPRO Gold users was 6.96 shades. Tooth hypersensitivity varied from none to severe. Tissue irritation was minimal. The results of these evaluations indicate that NUPRO Gold is effective as a tooth-whitening system, when administered properly under the supervision of a dentist, with commonly reported side effects of transient tooth sensitivity and minimal gingival sensitivity. Little or no change in tissue health was noted. This study was supported by Dentsply Preventive Care (York, Pennsylvania).

Leaf traits show different relationships with shade tolerance in moist versus dry tropical forests.

PubMed

Poorter, Lourens

2009-03-01

Shade tolerance is the central paradigm for understanding forest succession and dynamics, but there is considerable debate as to what the salient features of shade tolerance are, whether adult leaves show similar shade adaptations to seedling leaves, and whether the same leaf adaptations are found in forests under different climatic control. Here, adult leaf and metamer traits were measured for 39 tree species from a tropical moist semi-evergreen forest (1580 mm rain yr(-1)) and 41 species from a dry deciduous forest (1160 mm yr(-1)) in Bolivia. Twenty-six functional traits were measured and related to species regeneration light requirements.Adult leaf traits were clearly associated with shade tolerance. Different, rather than stronger, shade adaptations were found for moist compared with dry forest species. Shade adaptations exclusively found in the evergreen moist forest were related to tough and persistent leaves, and shade adaptations in the dry deciduous forest were related to high light interception and water use.These results suggest that, for forests differing in rainfall seasonality, there is a shift in the relative importance of functional leaf traits and performance trade-offs that control light partitioning. In the moist evergreen forest leaf traits underlying the growth-survival trade-off are important, whereas in the seasonally deciduous forest leaf traits underlying the growth trade-off between low and high light might become important.

Effects of shade on welfare and meat quality of grazing sheep under high ambient temperature.

PubMed

Liu, H W; Cao, Y; Zhou, D W

2012-12-01

This study was conducted to evaluate the effects of providing shade on growth performance, welfare, and meat quality of grazing sheep under high ambient temperature. A total of 120 healthy male Ujumqin wool sheep (a local breed; BW = 18.7 ± 1.27 kg; 14 wk old) were randomly and equally divided into shaded and unshaded treatments with 3 pens per treatment. Sheep were grazed on an unshaded pastureland from 0600 to 1000 h and 1400 to 1800 h. During other times, sheep were confined in shaded or unshaded pens. Body weight was recorded on d 1 and 42 of the experiment. Rectal temperature and respiration rate were recorded on d 7, 14, 21, 28, 35, and 42. At end of the trial, sheep were blood sampled and slaughtered to collect meat samples. Respiration rate was greater (P < 0.05) in the unshaded sheep than shaded sheep on d 14, 21, 28, 35, and 42 of the trial whereas no significant differences were found on d 7. Moreover, no differences were observed in final BW, ADG, or rectal temperature throughout the trial. The pH at 24 h postmortem (pH(24)) and cooking loss were greater (P < 0.01) in unshaded than shaded sheep. On the contrary, lightness (L*), redness (a*) and yellowness (b*) values at 24 h postmortem were lower (P < 0.05) in unshaded versus shaded sheep. The sheep in the unshaded group had a greater (P < 0.05) cortisol concentration compared with the shaded group. Sheep in the shaded group had lower creatine kinase activity (P < 0.01) as well as observed for glucose (P < 0.05), triiodothyronine (P < 0.01), and thyroxine (P < 0.05) concentrations and white blood cell count (P < 0.05). Compared with the unshaded group, sheep in the shaded group had a greater lymphocytes (LYM) count (P < 0.05). In contrast, the opposite was true for neutrophils (NEU) count (P < 0.01) and NEU:LYM ratio (P < 0.01). In conclusion, the shade cloth, although not enhancing ADG, improved meat quality traits and certain stress parameters in grazing sheep reared under high ambient temperature.

Seedling growth and biomass allocation in relation to leaf habit and shade tolerance among 10 temperate tree species.

PubMed

Modrzyński, Jerzy; Chmura, Daniel J; Tjoelker, Mark G

2015-08-01

Initial growth of germinated seeds is an important life history stage, critical for establishment and succession in forests. Important questions remain regarding the differences among species in early growth potential arising from shade tolerance. In addition, the role of leaf habit in shaping relationships underlying shade tolerance-related differences in seedling growth remains unresolved. In this study we examined variation in morphological and physiological traits among seedlings of 10 forest tree species of the European temperate zone varying in shade tolerance and leaf habit (broadleaved winter-deciduous species vs needle-leaved conifers) during a 10-week period. Seeds were germinated and grown in a controlled environment simulating an intermediate forest understory light environment to resolve species differences in initial growth and biomass allocation. In the high-resource experimental conditions during the study, seedlings increased biomass allocation to roots at the cost of leaf biomass independent of shade tolerance and leaf habit. Strong correlations between relative growth rate (RGR), net assimilation rate (NAR), leaf area ratio (LAR), specific leaf area (SLA) and leaf mass fraction (LMF) indicate that physiology and biomass allocation were equally important determinants of RGR as plant structure and leaf morphology among these species. Our findings highlight the importance of seed mass- and seed size-related root morphology (specific root length-SRL) for shade tolerance during early ontogeny. Leaf and plant morphology (SLA, LAR) were more successful in explaining variation among species due to leaf habit than shade tolerance. In both broadleaves and conifers, shade-tolerant species had lower SRL and greater allocation of biomass to stems (stem mass fraction). Light-seeded shade-intolerant species with greater SRL had greater RGR in both leaf habit groups. However, the greatest plant mass was accumulated in the group of heavy-seeded shade-tolerant broadleaves. The results of our study suggest that the combinations of plant attributes enhancing growth under high light vary with shade tolerance, but differ between leaf habit groups. © The Author 2015. Published by Oxford University Press. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

Biomass Allocation of Stoloniferous and Rhizomatous Plant in Response to Resource Availability: A Phylogenetic Meta-Analysis

PubMed Central

Xie, Xiu-Fang; Hu, Yu-Kun; Pan, Xu; Liu, Feng-Hong; Song, Yao-Bin; Dong, Ming

2016-01-01

Resource allocation to different functions is central in life-history theory. Plasticity of functional traits allows clonal plants to regulate their resource allocation to meet changing environments. In this study, biomass allocation traits of clonal plants were categorized into absolute biomass for vegetative growth vs. for reproduction, and their relative ratios based on a data set including 115 species and derived from 139 published literatures. We examined general pattern of biomass allocation of clonal plants in response to availabilities of resource (e.g., light, nutrients, and water) using phylogenetic meta-analysis. We also tested whether the pattern differed among clonal organ types (stolon vs. rhizome). Overall, we found that stoloniferous plants were more sensitive to light intensity than rhizomatous plants, preferentially allocating biomass to vegetative growth, aboveground part and clonal reproduction under shaded conditions. Under nutrient- and water-poor condition, rhizomatous plants were constrained more by ontogeny than by resource availability, preferentially allocating biomass to belowground part. Biomass allocation between belowground and aboveground part of clonal plants generally supported the optimal allocation theory. No general pattern of trade-off was found between growth and reproduction, and neither between sexual and clonal reproduction. Using phylogenetic meta-analysis can avoid possible confounding effects of phylogeny on the results. Our results shown the optimal allocation theory explained a general trend, which the clonal plants are able to plastically regulate their biomass allocation, to cope with changing resource availability, at least in stoloniferous and rhizomatous plants. PMID:27200071

Influence of resin cement shade on the color and translucency of ceramic veneers

PubMed Central

HERNANDES, Daiana Kelly Lopes; ARRAIS, Cesar Augusto Galvão; de LIMA, Erick; CESAR, Paulo Francisco; RODRIGUES, José Augusto

2016-01-01

ABSTRACT Objective This in vitro study evaluated the effect of two different shades of resin cement (RC- A1 and A3) layer on color change, translucency parameter (TP), and chroma of low (LT) and high (HT) translucent reinforced lithium disilicate ceramic laminates. Material and Methods One dual-cured RC (Variolink II, A1- and A3-shade, Ivoclar Vivadent) was applied to 1-mm thick ceramic discs to create thin RC films (100 µm thick) under the ceramics. The RC was exposed to light from a LED curing unit. Color change (ΔE) of ceramic discs was measured according to CIEL*a*b* system with a standard illuminant D65 in reflectance mode in a spectrophotometer, operating in the light range of 360-740 nm, equipped with an integrating sphere. The color difference between black (B) and white (W) background readings was used for TP analysis, while chroma was calculated by the formula C* ab=(a*2+b*2)½. ΔE of 3.3 was set as the threshold of clinically unacceptable. The results were evaluated by two-way ANOVA followed by Tukey's post hoc test. Results HT ceramics showed higher ΔE and higher TP than LT ceramics. A3-shade RC promoted higher ΔE than A1-shade cement, regardless of the ceramic translucency. No significant difference in TP was noted between ceramic discs with A1- and those with A3-shade cement. Ceramic with underlying RC showed lower TP than discs without RC. HT ceramics showed lower chroma than LT ceramics, regardless of the resin cement shade. The presence of A3-shade RC resulted in higher chroma than the presence of A1-shade RC. Conclusions Darker underlying RC layer promoted more pronounced changes in ceramic translucency, chroma, and shade of high translucent ceramic veneers. These differences may not be clinically differentiable. PMID:27556211

Influence of resin cement shade on the color and translucency of ceramic veneers.

PubMed

Hernandes, Daiana Kelly Lopes; Arrais, Cesar Augusto Galvão; Lima, Erick de; Cesar, Paulo Francisco; Rodrigues, José Augusto

2016-01-01

This in vitro study evaluated the effect of two different shades of resin cement (RC- A1 and A3) layer on color change, translucency parameter (TP), and chroma of low (LT) and high (HT) translucent reinforced lithium disilicate ceramic laminates. One dual-cured RC (Variolink II, A1- and A3-shade, Ivoclar Vivadent) was applied to 1-mm thick ceramic discs to create thin RC films (100 µm thick) under the ceramics. The RC was exposed to light from a LED curing unit. Color change (ΔE) of ceramic discs was measured according to CIEL*a*b* system with a standard illuminant D65 in reflectance mode in a spectrophotometer, operating in the light range of 360-740 nm, equipped with an integrating sphere. The color difference between black (B) and white (W) background readings was used for TP analysis, while chroma was calculated by the formula C*ab=(a*2+b*2)½. ΔE of 3.3 was set as the threshold of clinically unacceptable. The results were evaluated by two-way ANOVA followed by Tukey's post hoc test. HT ceramics showed higher ΔE and higher TP than LT ceramics. A3-shade RC promoted higher ΔE than A1-shade cement, regardless of the ceramic translucency. No significant difference in TP was noted between ceramic discs with A1- and those with A3-shade cement. Ceramic with underlying RC showed lower TP than discs without RC. HT ceramics showed lower chroma than LT ceramics, regardless of the resin cement shade. The presence of A3-shade RC resulted in higher chroma than the presence of A1-shade RC. Darker underlying RC layer promoted more pronounced changes in ceramic translucency, chroma, and shade of high translucent ceramic veneers. These differences may not be clinically differentiable.

Species abundance and potential biological control services in shade vs. sun coffee in Puerto Rico

USGS Publications Warehouse

Borkhataria, Rena R.; Collazo, Jaime A.; Groom, Martha J.

2012-01-01

Birds, lizards and insects were surveyed in three sun and three shade coffee plantations in Puerto Rico to provide a comprehensive comparison of biodiversity between plantations types and to identify potential interrelationships (e.g., biological or natural control services) between members of each taxon and coffee pests. Abundance of avian species, including insectivorous species, was significantly higher in shade coffee. Anolis cristatellus and A. stratulus were significantly more abundant in sun plantations whereas A. gundlachi and A. evermanni were detected more frequently in shade plantations. Insects in the orders Coleoptera, Diptera, Hemiptera, Hymenoptera, Lepidoptera, Neuroptera, and Psocoptera were significantly more abundant in shade coffee, while orthopterans were more abundant in sun. The coffee leaf miner (Leucoptera coffeela) and the flatid planthopper (Petrusa epilepsis) did not differ significantly between plantation types, nor did the abundance of the wasp complex that parasitizes the coffee leaf miner. These findings confirmed that shade plantations harbor a wide array of elements of biodiversity; but sun plantations may also harbor many elements of biodiversity, and in some cases, in higher abundance than in shade plantations.

Effect of shade on Arabica coffee berry disease development: Toward an agroforestry system to reduce disease impact.

PubMed

Mouen Bedimo, J A; Njiayouom, I; Bieysse, D; Ndoumbè Nkeng, M; Cilas, C; Nottéghem, J L

2008-12-01

Coffee berry disease (CBD), caused by Colletotrichum kahawae, is a major constraint for Arabica coffee cultivation in Africa. The disease is specific to green berries and can lead to 60% harvest losses. In Cameroon, mixed cropping systems of coffee with other crops, such as fruit trees, are very widespread agricultural practices. Fruit trees are commonly planted at random on coffee farms, providing a heterogeneous shading pattern for coffee trees growing underneath. Based on a recent study of CBD, it is known that those plants can reduce disease incidence. To assess the specific effect of shade, in situ and in vitro disease development was compared between coffee trees shaded artificially by a net and trees located in full sunlight. In the field, assessments confirmed a reduction in CBD on trees grown under shade compared with those grown in full sunlight. Artificial inoculations in the laboratory showed that shade did not have any effect on the intrinsic susceptibility of coffee berries to CBD. Coffee shading mainly acts on environmental parameters in limiting disease incidence. In addition to reducing yield losses, agroforestry system may also be helpful in reducing chemical control of the disease and in diversifying coffee growers' incomes.

Physiological and Morphological Responses of the Temperate Seagrass Zostera muelleri to Multiple Stressors: Investigating the Interactive Effects of Light and Temperature

PubMed Central

York, Paul H.; Gruber, Renee K.; Hill, Ross; Ralph, Peter J.; Booth, David J.; Macreadie, Peter I.

2013-01-01

Understanding how multiple environmental stressors interact to affect seagrass health (measured as morphological and physiological responses) is important for responding to global declines in seagrass populations. We investigated the interactive effects of temperature stress (24, 27, 30 and 32°C) and shading stress (75, 50, 25 and 0% shade treatments) on the seagrass Zostera muelleri over a 3-month period in laboratory mesocosms. Z. muelleri is widely distributed throughout the temperate and tropical waters of south and east coasts of Australia, and is regarded as a regionally significant species. Optimal growth was observed at 27°C, whereas rapid loss of living shoots and leaf mass occurred at 32°C. We found no difference in the concentration of photosynthetic pigments among temperature treatments by the end of the experiment; however, up-regulation of photoprotective pigments was observed at 30°C. Greater levels of shade resulting in high photochemical efficiencies, while elevated irradiance suppressed effective quantum yield (ΔF/FM’). Chlorophyll fluorescence fast induction curves (FIC) revealed that the J step amplitude was significantly higher in the 0% shade treatment after 8 weeks, indicating a closure of PSII reaction centres, which likely contributed to the decline in ΔF/FM’ and photoinhibition under higher irradiance. Effective quantum yield of PSII (ΔF/FM’) declined steadily in 32°C treatments, indicating thermal damage. Higher temperatures (30°C) resulted in reduced above-ground biomass ratio and smaller leaves, while reduced light led to a reduction in leaf and shoot density, above-ground biomass ratio, shoot biomass and an increase in leaf senescence. Surprisingly, light and temperature had few interactive effects on seagrass health, even though these two stressors had strong effects on seagrass health when tested in isolation. In summary, these results demonstrate that populations of Z. muelleri in south-eastern Australia are sensitive to small chronic temperature increases and light decreases that are predicted under future climate change scenarios. PMID:24124551

Response of the physiological parameters of mango fruit (transpiration, water relations and antioxidant system) to its light and temperature environment.

PubMed

Léchaudel, Mathieu; Lopez-Lauri, Félicie; Vidal, Véronique; Sallanon, Huguette; Joas, Jacques

2013-04-15

Depending on the position of the fruit in the tree, mango fruit may be exposed to high temperature and intense light conditions that may lead to metabolic and physiological disorders and affect yield and quality. The present study aimed to determine how mango fruit adapted its functioning in terms of fruit water relations, epicarp characteristics and the antioxidant defence system in peel, to environmental conditions. The effect of contrasted temperature and light conditions was evaluated under natural solar radiation and temperature by comparing well-exposed and shaded fruit at three stages of fruit development. The sun-exposed and shaded peels of the two sides of the well-exposed fruit were also compared. Depending on fruit position within the canopy and on the side of a well-exposed fruit, the temperature gradient over a day affected fruit characteristics such as transpiration, as revealed by the water potential gradient as a function of the treatments, and led to a significant decrease in water conductance for well-exposed fruits compared to fruits within the canopy. Changes in cuticle thickness according to fruit position were consistent with those of fruit water conductance. Osmotic potential was also affected by climatic environment and harvest stage. Environmental conditions that induced water stress and greater light exposure, like on the sunny side of well-exposed fruit, increased the hydrogen peroxide, malondialdehyde and total and reduced ascorbate contents, as well as SOD, APX and MDHAR activities, regardless of the maturity stage. The lowest values were measured in the peel of the shaded fruit, that of the shaded side of well-exposed fruit being intermediate. Mango fruits exposed to water-stress-induced conditions during growth adapt their functioning by reducing their transpiration. Moreover, oxidative stress was limited as a consequence of the increase in antioxidant content and enzyme activities. This adaptive response of mango fruit to its climatic environment during growth could affect postharvest behaviour and quality. Copyright © 2012 Elsevier GmbH. All rights reserved.

Shifting roles of heterotrophy and autotrophy in coral energetics under varying turbidity.

PubMed

Anthony; Fabricius

2000-09-20

Suspended particulate matter (SPM) strongly alters the trophic environment of photosymbiotic aquatic organisms. At high particles loads, phototrophic energy gains can be diminished due to light absorption by suspended particles, and stress from particle abrasion or deposition on tissues. However, energy gains are enhanced if organisms are able to use SPM as a food source. For photosymbiotic benthic suspension feeders, increases in SPM concentrations may require both phototrophic and heterotrophic acclimation to sustain a positive energy balance. This study provides an experimental analysis of the effects of contrasting light and SPM regimes on the energy budget (scope for growth) of two zooxanthellate corals (Goniastrea retiformis and Porites cylindrica). Using a factorial design in a flow-through tank system, corals were exposed for 2 months to shaded and unshaded conditions (equivalent to 3-4 m depth at 4 and 16 mg dry weight SPM l(-1), respectively) and a range of controlled SPM loads with a natural organic content ( approximately 3% w/w). In G. retiformis, rates of particle ingestion were a linear function of SPM concentration within a broad range (1-30 mg dry weight l(-1)). After 2 months of shading, photosynthetic acclimation was significant in G. retiformis, but did not compensate for the reduced light level, as daily respiration exceeded daily photosynthesis. However, in response to the prolonged shading, G. retiformis more than doubled its rate of particle feeding. At high SPM treatments (16 mg dw l(-1)), sediment feeding by this species compensated fully for the 35-47% lower phototrophy in the shaded treatment. Due to both photo- and heterotrophic plasticity, G. retiformis gained tissue and skeletal mass at all experimental levels of light and SPM. In contrast, rates of particle intake by P. cylindrica contributed <10% to the energy budget in shaded and <3% in unshaded conditions. Feeding rates of P. cylindrica were half-saturated at approximately 3 mg dry weight l(-1), and four- to eight-fold lower than those of G. retiformis. Skeletal growth was sustained, but tissue mass and lipid contents declined in shaded and high-SPM treatments, and carbon loss due to shading by SPM was not compensated for by particle feeding. Thus, due to a lack of photo- and heterotrophic plasticity, periods of high turbidity resulted in energy deficiency in P. cylindrica, and high turbidity conditions appeared physiologically unsustainable for this species. This study is the first to show heterotrophic plasticity in a symbiotic coral, and to show that such plasticity can offset stress from high particle loads. It demonstrates that changes in the trophic mode of some coral species are a mechanism for sustaining a positive energy balance in turbid environments, thereby broadening their physiological niche.

Relevant optical properties for direct restorative materials.

PubMed

Pecho, Oscar E; Ghinea, Razvan; do Amaral, Erika A Navarro; Cardona, Juan C; Della Bona, Alvaro; Pérez, María M

2016-05-01

To evaluate relevant optical properties of esthetic direct restorative materials focusing on whitened and translucent shades. Enamel (E), body (B), dentin (D), translucent (T) and whitened (Wh) shades for E (WhE) and B (WhB) from a restorative system (Filtek Supreme XTE, 3M ESPE) were evaluated. Samples (1 mm thick) were prepared. Spectral reflectance (R%) and color coordinates (L*, a*, b*, C* and h°) were measured against black and white backgrounds, using a spectroradiometer, in a viewing booth, with CIE D65 illuminant and d/0° geometry. Scattering (S) and absorption (K) coefficients and transmittance (T%) were calculated using Kubelka-Munk's equations. Translucency (TP) and opalescence (OP) parameters and whiteness index (W*) were obtained from differences of CIELAB color coordinates. R%, S, K and T% curves from all shades were compared using VAF (Variance Accounting For) coefficient with Cauchy-Schwarz inequality. Color coordinates and optical parameters were statistically analyzed using one-way ANOVA, Tukey's test with Bonferroni correction (α=0.0007). Spectral behavior of R% and S were different for T shades. In addition, T shades showed the lowest R%, S and K values, as well as the highest T%, TP an OP values. In most cases, WhB shades showed different color and optical properties (including TP and W*) than their corresponding B shades. WhE shades showed similar mean W* values and higher mean T% and TP values than E shades. When using whitened or translucent composites, the final color is influenced not only by the intraoral background but also by the color and optical properties of multilayers used in the esthetic restoration. Copyright © 2016 Academy of Dental Materials. Published by Elsevier Ltd. All rights reserved.

Effects of combining ozone and hydrogren peroxide on tooth bleaching: A clinical study.

PubMed

Al-Omiri, Mahmoud K; Hassan, Ra'ed S Abul; AlZarea, Bader K; Lynch, Edward

2016-10-01

The purpose of this study was to evaluate the efficiency of bleaching after combining ozone and 38% H2O2 in comparison to the sole use of 38% H2O2. Consecutive 26 participants (13 males and 13 females) were recruited into this study. They were randomly allocated into 2 groups (n=13 for each group). In group 1 (test group); the participants' upper anterior teeth were treated with 38% H2O2 for 20min then the teeth were exposed to ozone for 60s (healOzone(®) X4, KaVo Dental, Biberach, Germany). In group 2 (controls); the upper anterior teeth were treated with 20min of 38% H2O2 only. The shade of teeth was evaluated by recording the L* a* b* values and Vita Classic shades at study baseline and after bleaching in both groups. The statistically significant changes were set at P≤0.05. Tooth sensitivity and teeth shades were comparable between groups at study baseline (p>0.05). Controls reported more tooth sensitivity following bleaching (p<0.001). Teeth achieved better Vita shades, higher L* values (lighter shades), and lower a* and b* values (lighter shades) after bleaching in both groups (P≤0.05). However, teeth bleached with H2O2 and ozone achieved better Vita shades, higher L* values and lower a* values (lighter shades) than those bleached with H2O2 alone (p<0.001). Changes in b* values were not significantly different between groups. Bleaching with 38% H2O2 for 20min followed by 60s of ozone application would result in teeth with lighter shades than bleaching with 38% H2O2 alone. Copyright © 2016 Elsevier Ltd. All rights reserved.

Is shade beneficial for mediterranean shrubs experiencing periods of extreme drought and late-winter frosts?

PubMed

Valladares, Fernando; Zaragoza-Castells, Joana; Sánchez-Gómez, David; Matesanz, Silvia; Alonso, Beatriz; Portsmuth, Angelika; Delgado, Antonio; Atkin, Owen K

2008-12-01

Plants are naturally exposed to multiple, frequently interactive stress factors, most of which are becoming more severe due to global change. Established plants have been reported to facilitate the establishment of juvenile plants, but net effects of plant-plant interactions are difficult to assess due to complex interactions among environmental factors. An investigation was carried out in order to determine how two dominant evergreen shrubs (Quercus ilex and Arctostaphylos uva-ursi) co-occurring in continental, Mediterranean habitats respond to multiple abiotic stresses and whether the shaded understorey conditions ameliorate the negative effects of drought and winter frosts on the physiology of leaves. Microclimate and ecophysiology of sun and shade plants were studied at a continental plateau in central Spain during 2004-2005, with 2005 being one of the driest and hottest years on record; several late-winter frosts also occurred in 2005. Daytime air temperature and vapour pressure deficit were lower in the shade than in the sun, but soil moisture was also lower in the shade during the spring and summer of 2005, and night-time temperatures were higher in the shade. Water potential, photochemical efficiency, light-saturated photosynthesis, stomatal conductance and leaf 13C composition differed between sun and shade individuals throughout the seasons, but differences were species specific. Shade was beneficial for leaf-level physiology in Q. ilex during winter, detrimental during spring for both species, and of little consequence in summer. The results suggest that beneficial effects of shade can be eclipsed by reduced soil moisture during dry years, which are expected to be more frequent in the most likely climate change scenarios for the Mediterranean region.

Is Shade Beneficial for Mediterranean Shrubs Experiencing Periods of Extreme Drought and Late-winter Frosts?

PubMed Central

Valladares, Fernando; Zaragoza-Castells, Joana; Sánchez-Gómez, David; Matesanz, Silvia; Alonso, Beatriz; Portsmuth, Angelika; Delgado, Antonio; Atkin, Owen K.

2008-01-01

Background and Aims Plants are naturally exposed to multiple, frequently interactive stress factors, most of which are becoming more severe due to global change. Established plants have been reported to facilitate the establishment of juvenile plants, but net effects of plant–plant interactions are difficult to assess due to complex interactions among environmental factors. An investigation was carried out in order to determine how two dominant evergreen shrubs (Quercus ilex and Arctostaphylos uva-ursi) co-occurring in continental, Mediterranean habitats respond to multiple abiotic stresses and whether the shaded understorey conditions ameliorate the negative effects of drought and winter frosts on the physiology of leaves. Methods Microclimate and ecophysiology of sun and shade plants were studied at a continental plateau in central Spain during 2004–2005, with 2005 being one of the driest and hottest years on record; several late-winter frosts also occurred in 2005. Key Results Daytime air temperature and vapour pressure deficit were lower in the shade than in the sun, but soil moisture was also lower in the shade during the spring and summer of 2005, and night-time temperatures were higher in the shade. Water potential, photochemical efficiency, light-saturated photosynthesis, stomatal conductance and leaf 13C composition differed between sun and shade individuals throughout the seasons, but differences were species specific. Shade was beneficial for leaf-level physiology in Q. ilex during winter, detrimental during spring for both species, and of little consequence in summer. Conclusions The results suggest that beneficial effects of shade can be eclipsed by reduced soil moisture during dry years, which are expected to be more frequent in the most likely climate change scenarios for the Mediterranean region. PMID:18819947

Rationale, design, and baseline data of a cross-national randomized trial on the effect of built shade in public parks for sun protection.

PubMed

Buller, David B; Dobbinson, Suzanne; English, Dallas R; Wakefield, Melanie; Buller, Mary Klein

2017-04-01

Environments can be structured to reduce solar ultraviolet radiation (UV) exposure to prevent skin cancer. A prospective randomized trial is being conducted to test whether introducing shade sails in passive recreation areas (PRAs) in public parks will increase use and decrease UV exposure in the shaded compared to unshaded PRAs. Shade effects will be compared between Melbourne, Australia and Denver, USA. The trial enrolled 145 public parks with PRAs suitable for shade construction and randomized parks to intervention or control in a 1:3 ratio. Use of PRAs and UV levels were recorded at each park by trained observers during 30-minute periods on four weekend days in each of two summers (pretest and posttest). Shade sails were constructed between the summers. Given low numbers of users at pretest, the outcome measure was modified to use of the PRA (use vs. no use) and unit of analysis to the individual observation. Observations (n=580) occurred on average 29days from the summer solstice and 55min from solar noon in warm weather (mean=26.2°C) with some cloud cover but only slight or no wind. Typically, PRAs had benches and picnic tables and were located near playgrounds. PRAs were in use during 13.3% of observations (mean=0.41 users). UV over 30-minutes at the PRA boundary (mean=3.2 standard erythemal dose [SED]) and center (mean=3.3 SED) was high. Shade for skin cancer prevention has been understudied. This study will address this gap by determining whether purpose-built shade structures promote greater use of shaded areas within public parks. Copyright © 2017 Elsevier Inc. All rights reserved.

Light compensation points in shade-grown seedlings of deciduous broadleaf tree species with different successional traits raised under elevated CO2.

PubMed

Kitao, M; Hida, T; Eguchi, N; Tobita, H; Utsugi, H; Uemura, A; Kitaoka, S; Koike, T

2016-01-01

We measured leaf photosynthetic traits in shade-grown seedlings of four tree species native to northern Japan, raised under an elevated CO2 condition, to investigate the effects of elevated CO2 on shade tolerance of deciduous broadleaf tree species with different successional traits. We considered Betula platyphylla var. japonica and Betula maximowicziana as pioneer species, Quercus mongolica var. crispula as a mid-successional species, and Acer mono as a climax species. The plants were grown under shade conditions (10% of full sunlight) in a CO2 -regulated phytotron. Light compensation points (LCPs) decreased in all tree species when grown under elevated CO2 (720 μmol·mol(-1) ), which were accompanied by higher apparent quantum yields but no photosynthetic down-regulation. LCPs in Q. mongolica and A. mono grown under elevated CO2 were lower than those in the two pioneer birch species. The LCP in Q. mongolica seedlings was not different from that of A. mono in each CO2 treatment. However, lower dark respiration rates were observed in A. mono than in Q. mongolica, suggesting higher shade tolerance in A. mono as a climax species in relation to carbon loss at night. Thus, elevated CO2 may have enhanced shade tolerance by lowering LCPs in all species, but the ranking of shade tolerance related to successional traits did not change among species under elevated CO2 , i.e. the highest shade tolerance was observed in the climax species (A. mono), followed by a gap-dependent species (Q. mongolica), while lower shade tolerance was observed in the pioneer species (B. platyphylla and B. maximowicziana). © 2015 German Botanical Society and The Royal Botanical Society of the Netherlands.

Analysis of Shade Matching in Natural Dentitions Using Intraoral Digital Spectrophotometer in LED and Filtered LED Light Sources.

PubMed

Chitrarsu, Vijai Krishnan; Chidambaranathan, Ahila Singaravel; Balasubramaniam, Muthukumar

2017-10-31

To evaluate the shade matching capabilities in natural dentitions using Vita Toothguide 3D-Master and an intraoral digital spectrophotometer (Vita Easyshade Advance 4.0) in various light sources. Participants between 20 and 40 years old with natural, unrestored right maxillary central incisors, no history of bleaching, orthodontic treatment, or malocclusion and no rotations were included. According to their shades, subjects were randomly selected and grouped into A1, A2, and A3. A total of 100 participants (50 male and 50 female) in each group were chosen for this study. Shade selection was made between 10 am and 2 pm for all light sources. The same examiner selected the shade of natural teeth with Vita Toothguide 3D-Master under natural light within 2 minutes. Once the Vita Toothguide 3D-Masterwas matched with the maxillary right central incisor, the L*, a*, and b* values, chroma, and hue were recorded with Vita Easyshade Advance 4.0 by placing it on the shade tab under the same light source. The values were statistically analyzed using one-way ANOVA and Tukey's HSD post hoc test with SPSS v22.0 software. The mean ∆E* ab values for shades A1, A2, and A3 for groups 1, 2, and 3 were statistically significantly different from each other (p < 0.001). The intraoral digital spectrophotometer showed statistically significant differences in shade matching compared to Vita Toothguide 3D-Master. Incandescent light showed more accurate shade matching than the filtered LED, LED, and daylight. © 2017 by the American College of Prosthodontists.

Shade-grown coffee in Puerto Rico: Opportunities to preserve biodiversity while reinvigorating a struggling agricultural commodity

USGS Publications Warehouse

Borkhataria, R.; Collazo, J.A.; Groom, Martha J.; Jordan-Garcia, A.

2012-01-01

Shade-grown coffee contributes to biodiversity conservation and has many ecological benefits. We reviewed historical trends in coffee production and interviewed 100 coffee growers in 1999 to determine current management practices and attitudes toward the cultivation of sun and shade coffee in Puerto Rico. We discuss the outlook for the coffee industry in the 21st century and implications for biodiversity conservation, hoping lessons from Puerto Rico will apply to the international coffee industry. Throughout the 20th century, government intervention, including subsidies and technical assistance, supported coffee farming in Puerto Rico. In an effort to modernize coffee production and increase yields, the conversion from shade to sun coffee plantations was encouraged. Despite government support, the amount of land devoted to this once dominant agricultural commodity declined markedly between 1982 and 2007 (84%), due to labor shortages, low income, and catastrophic hurricanes. We found that a return to shaded plantations would be embraced by most farmers. Growers of shaded coffee were generally happier with their cultivation practices (89.3% satisfied) than growers of sun coffee (60.9% satisfied), valued biodiversity, and were willing to cultivate coffee under shade if given similar incentives to those received for farming sun coffee. The future of the coffee industry in Puerto Rico may depend on government programs that capitalize upon emerging markets for sustainably produced, shade-grown coffee. We conclude that where governments have close ties to the coffee industry, they should strive to wed economic development with the conservation of biodiversity and associated ecological services by providing support and incentives for the production of shade coffee. ?? 2011 Elsevier B.V.

Rationale, Design, and Baseline Data of a Cross-national Randomized Trial on the Effect of Built Shade in Public Parks for Sun Protection

PubMed Central

Buller, David B.; Dobbinson, Suzanne; English, Dallas R.; Wakefield, Melanie; Buller, Mary Klein

2017-01-01

Environments can be structured to reduce solar ultraviolet radiation (UV) exposure to prevent skin cancer. A prospective randomized trial is being conducted to test whether introducing shade sales in passive recreation areas (PRAs) in public parks will increase use and decrease UV exposure in the shaded compared to unshaded PRAs. Shade effects will be compared between Melbourne, Australia and Denver, USA. The trial enrolled 144 public parks with PRAs suitable for shade construction and randomized parks to intervention or control in a 1:3 ratio. Use of PRAs and UV levels were recorded at each park by trained observers during 30-minute periods on four weekend days in each of two summers (pretest and posttest). Shade sails were constructed between the summers. Given low numbers of users at pretest, the outcome measure was modified to use of the PRA (use v. no use) and unit of analysis to the individual observation. Observations (n=580) occurred on average 29 days from the summer solstice and 55 minutes from solar noon in warm weather (mean=26.2°C) with some cloud cover but only slight or no wind. Typically, PRAs had benches and picnic tables and were located near playgrounds. PRAs were in use during 13.3% of observations (mean=0.41 users). UV over 30-minutes at the PRA boundary (mean=3.2 standard erythemal dose [SED]) and center (mean=3.3 SED) was high. Shade for skin cancer prevention has been understudied. This study will address this gap by determining whether purpose-built shade structures promote greater use of shaded areas within public parks. PMID:28185996

Shade images of forested areas obtained from Landsat MSS data

NASA Technical Reports Server (NTRS)

Shimabukuro, Yosio Edemir; Smith, James A.

1989-01-01

The objective of this report is to generate a shade (shadow) image of forested areas from Landsat MSS data by implementing a linear mixing model, where shadow is considered as one of the primary components in a pixel. The shade images are related to the observed variation in forest structure; i.e., the proportion of inferred shadow in a pixel is related to different forest ages, forest types, and tree crown cover. The constrained least-squares method is used to generate shade images for forest of eucalyptus and vegetation of 'cerrado' over the Itapeva study area in Brazil. The resulted shade images may explain the difference on ages for forest of eucalyptus and the difference on tree crown cover for vegetation of cerrado.

[Effects of shading on two Sphagnum species growth and their interactions].

PubMed

Ma, Jin-Ze; Bu, Zhao-Jun; Zheng, Xing-Xing; Li, Shan-Lin; Zeng, Jing; Zhao, Gao-Lin

2012-02-01

Taking Sphagnum palustre and S. fallax as test materials, this paper studied their growth and interactions under shading. In monoculture, shading promoted the height growth of S. palustre markedly, but had no effect on the growth of S. fallax and the biomass and branching of S. palustre. In mixed culture, S. fallax suppressed the increase of biomass and branching of S. palustre, while S. palustre had no effects on S. fallax. With the increase of shading stress, the competition of neighbour on S. fallax intensified. When the stress increased further, neighbor effect on S. fallax tended to be positive. However, the effect of neighbour on S. palustre was always competitive and did not change with the increase of shading stress.

Redwood seedling responses to light patterns and intensities

Treesearch

Ronald W. Boldenow; Joe R. McBride

2017-01-01

Coast redwood (Sequoia sempervirens (D. Don) Endl.) seedlings were grown from seed in controlled environments with 16 hour photoperiods using three light patterns that mimicked full shade (constant light level), intermittent high light such as long duration sun flecks (low light with 15 minutes of intense light every 2 hours), and large...

Understory structure in a 23-year-old Acacia koa forest and 2-year growth responses to silvicultural treatments

Treesearch

Paul G. Scowcroft; Janis E. Haraguchi; David M. Fujii

2008-01-01

Restoration of degraded Acacia koa forests in Hawaii often involves mechanical scarification to stimulate germination of seed buried in the soil and to suppress vegetation that competes with shade intolerant A. koa. Resulting even-age stands are gradually colonized by other plant species, but understory...

Plant Pigment Identification: A Classroom and Outreach Activity

ERIC Educational Resources Information Center

Garber, Kathleen C. A.; Odendaal, Antoinette Y.; Carlson, Erin E.

2013-01-01

Anthocyanins are a class of pigments responsible for the bright colors of many flowers, fruits, and vegetables typically resulting in shades of red, blue, and purple. Students were asked to perform an activity to enable them to identify which anthocyanin was present in one of several possible plant materials through a hands-on activity. Students…

Effects of simulated eutrophication and overfishing on algae and invertebrate settlement in a coral reef of Koh Phangan, Gulf of Thailand.

PubMed

Stuhldreier, Ines; Bastian, Pepe; Schönig, Eike; Wild, Christian

2015-03-15

Coral reefs in the Gulf of Thailand are highly under-investigated regarding responses to anthropogenic stressors. Thus, this study simulated overfishing and eutrophication using herbivore exclosure cages and slow-release fertilizer to study the in-situ effects on benthic algae and invertebrate settlement in a coral reef of Koh Phangan, Thailand. Settlement of organisms and the development of organic matter on light-exposed and shaded tiles were quantified weekly/biweekly over a study period of 12 weeks. Simulated eutrophication did not significantly influence response parameters, while simulated overfishing positively affected dry mass, turf algae height and fleshy macroalgae occurrence on light-exposed tiles. On shaded tiles, settlement of crustose coralline algae decreased, while abundances of ascidians increased compared to controls. An interactive effect of both stressors was not observed. These results hint to herbivory as actual key controlling factor on the benthic community, and fleshy macroalgae together with ascidians as potential bioindicators for local overfishing. Copyright © 2015 Elsevier Ltd. All rights reserved.

Effect of shade on animal welfare, growth performance, and carcass characteristics in large pens of beef cattle fed a beta agonist in a commercial feedlot.

PubMed

Hagenmaier, J A; Reinhardt, C D; Bartle, S J; Thomson, D U

2016-12-01

Feedlot cattle ( = 1,395; BW = 568 ± 43 kg) were used to evaluate the effects of shade on animal welfare, growth performance, and carcass quality during the summer of 2013 in a Kansas commercial feedlot. Seven lots of predominately black steers and heifers (4 and 3, respectively) visually determined to be approaching the final mo on feed were identified, randomly gate-sorted, and allocated to pens located across the feed alley from each other to receive 1 of 2 treatments: 1) Shade (mean shade area = 1.5 m/ animal) or 2) No shade. Shade was provided using a 13-ounce polyethylene fabric and pens were oriented northwest to southeast. The mean starting date was June 13 and the mean days on feed for lots while on the study was 38 d. Cattle were fed a 77.67% DM steam-flaked corn-based diet and had ad libitum access to water throughout the duration of the trial. Zilpaterol hydrochloride (ZIL) was included in the finishing ration at an inclusion rate of 8.3 mg/kg of DM for the last 20 d on feed with a 3 d withdrawal period. Pen floor temperatures (PFT) were measured using an infrared thermometer and prevalence of cattle open-mouth breathing (OMB) was recorded on a pen basis. In addition to shade treatment, the effect of temperature humidity index (THI) on PFT and OMB was analyzed by classifying days as either "Alert" (THI < 79) or "Danger" (THI > 79). On the day of slaughter, pens within a replicate were kept separate through all stages of the marketing channel from loading at the feedlot until stunning at the plant. Pen served as the experimental unit for all measurements. There was a THI × shade treatment interaction for PFT and OMB ( < 0.001) where days classified as "Danger" increased PFT and prevalence of OMB compared to "Alert" days in unshaded but not shaded cattle. Shaded cattle had greater DMI ( = 0.01); however, unshaded cattle had greater G:F ( = 0.05) and therefore no differences were observed in ADG ( = 0.39). Shaded cattle had greater dressing percentage ( = 0.01), although HCW, LM area, fat thickness, marbling score, and quality grade did not differ between treatments ( > 0.05). Heat stress, a significant animal welfare concern and cause of reduced performance in feedlot cattle during the final phase of the feeding period, was alleviated in shaded cattle and illustrates the importance of shade provision as 1 tool to protect the welfare and increase feed consumption in large pens of feedlot cattle during hot summer months.

Separate localization of light signal perception for sun or shade type chloroplast and palisade tissue differentiation in Chenopodium album.

PubMed

Yano, S; Terashima, I

2001-12-01

Physiological and ecological characteristics of sun and shade leaves have been compared in detail, but their developmental processes, in particular their light sensory mechanisms, are still unknown. This study compares the development of sun and shade leaves of Chenopodium album L., paying special attention to the light sensory site. We hypothesized that mature leaves sense the light environment, and that this information determines anatomy of new leaves. To examine this hypothesis, we shaded plants partially. In the low-light apex treatment (LA), the shoot apex with developing leaves was covered by a cap made of a shading screen and received photosynthetically active photon flux density (PPFD) of 60 micromol m(-2 )s(-1), while the remaining mature leaves were exposed to 360 micromol m(-2 )s(-1). In the high-light apex treatment (HA), the apex was exposed while the mature leaves were covered by a shade screen. After these treatments for 6 d, we analyzed leaf anatomy and chloroplast ultrastructure. The anatomy of LA leaves with a two-layered palisade tissue was similar to that of sun leaves, while their chloroplasts were shade-type with thick grana. The anatomy of HA leaves and shade leaves was similar and both had one-layered palisade tissue, while chloroplasts of HA leaves were sun-type having thin grana. These results clearly demonstrate that new leaves differentiate depending on the light environment of mature leaves, while chloroplasts differentiate depending on the local light environment.

Application of a digital technique in evaluating the reliability of shade guides.

PubMed

Cal, E; Sonugelen, M; Guneri, P; Kesercioglu, A; Kose, T

2004-05-01

There appears to be a need for a reliable method for quantification of tooth colour and analysis of shade. Therefore, the primary objective of this study was to show the applicability of graphic software in colour analysis and secondly to investigate the reliability of commercial shade guides produced by the same manufacturer, using this digital technique. After confirming the reliability and reproducibility of the digital method by using self-assessed coloured images, three shade guides of the same manufacturer were photographed in daylight and in studio environments with a digital camera and saved in tagged image file format (TIFF) format. Colour analysis of each photograph was performed using the Adobe Photoshop 4.0 graphic program. Luminosity, and red, green, blue (L and RGB) values of each shade tab of each shade guide were measured and the data were subjected to statistical analysis using the repeated measure Anova test. The L and RGB values of the images taken in daylight differed significantly from those of the images taken in studio environment (P < 0.05). In both environments, the luminosity and red values of the shade tabs were significantly different from each other (P < 0.05). It was concluded that, when the environmental conditions were kept constant, the Adobe Photoshop 4.0 colour analysis program could be used to analyse the colour of images. On the other hand, the results revealed that the accuracy of shade tabs widely being used in colour matching should be readdressed.

Effect of altitude on biochemical composition and quality of green arabica coffee beans can be affected by shade and postharvest processing method.

PubMed

Worku, Mohammed; de Meulenaer, Bruno; Duchateau, Luc; Boeckx, Pascal

2018-03-01

Although various studies have assessed altitude, shade and postharvest processing effects on biochemical content and quality of coffee beans, data on their interactions are scarce. The individual and interactive effects of these factors on the caffeine, chlorogenic acids (CGA) and sucrose contents as well as physical and sensory qualities of green coffee beans from large plantations in southwestern Ethiopia were evaluated. Caffeine and CGA contents decreased with increasing altitude; they respectively declined 0.12 and 1.23gkg -1 100m -1 . Sucrose content increased with altitude; however, the altitude effect was significant for wet-processed beans (3.02gkg -1 100m -1 ), but not for dry-processed beans (0.36g kg -1 100m -1 ). Similarly, sucrose content increased with altitude with much stronger effect for coffee grown without shade (2.11gkg -1 100m -1 ) compared to coffee grown under shade (0.93gkg -1 100m -1 ). Acidity increased with altitude when coffee was grown under shade (0.22 points 100m -1 ), but no significant altitude effect was observed on coffee grown without shade. Beans grown without shade showed a higher physical quality score for dry (37.2) than for wet processing (29.1). These results generally underline the complex interaction effects between altitude and shade or postharvest processing on biochemical composition and quality of green arabica coffee beans. Copyright © 2017. Published by Elsevier Ltd.

Testing the air pollution tolerances of shade tree cultivars

DOE Office of Scientific and Technical Information (OSTI.GOV)

Karnosky, D.J.

1978-01-01

A program in progress to test the relative air pollution tolerances of some commonly planted shade tree cultivars is described. The relative sulfur dioxide and ozone tolerances of several cultivars of acer, fagus, fraxinus, ginkgo, gleditsia, platanus, and quercus species are being determined by examination of foliar response to short-term, high-concentration fumigations. In a follow-up study, the extent of foliar injury, growth reduction, and mortality caused by air pollution will be examined for the same cultivars grown in test plantings in and around New York City. The correlation of the relative sulfur dioxide and ozone tolerances, as determined by themore » controlled-fumigation tests, with urban survival and growth will be useful in determining the adequacy of acute exposure fumigations in testing the air pollution tolerances of the tree species under study.« less

Lack of Physiological Depth Patterns in Conspecifics of Endemic Antarctic Brown Algae: A Trade-Off between UV Stress Tolerance and Shade Adaptation?

PubMed Central

Gómez, Iván; Huovinen, Pirjo

2015-01-01

A striking characteristic of endemic Antarctic brown algae is their broad vertical distribution. This feature is largely determined by the shade adaptation in order to cope with the seasonal variation in light availability. However, during spring-summer months, when light penetrates deep in the water column these organisms have to withstand high levels of solar radiation, including UV. In the present study we examine the light use characteristics in parallel to a potential for UV tolerance (measured as content of phenolic compounds, antioxidant activity and maximum quantum yield of fluorescence) in conspecific populations of four Antarctic brown algae (Ascoseira mirabilis, Desmarestia menziesii, D. anceps and Himantothallus grandifolius) distributed over a depth gradient between 5 and 30 m. The main results indicated that a) photosynthetic efficiency was uniform along the depth gradient in all the studied species, and b) short-term (6 h) exposure to UV radiation revealed a high tolerance measured as chlorophyll fluorescence, phlorotannin content and antioxidant capacity. Multivariate analysis of similarity indicated that light requirements for photosynthesis, soluble phlorotannins and antioxidant capacity are the variables determining the responses along the depth gradient in all the studied species. The suite of physiological responses of algae with a shallower distribution (A. mirabilis and D. menziesii) differed from those with deeper vertical range (D. anceps and H. grandifolius). These patterns are consistent with the underwater light penetration that defines two zones: 0–15 m, with influence of UV radiation (1% of UV-B and UV-A at 9 m and 15 m respectively) and a zone below 15 m marked by PAR incidence (1% up to 30 m). These results support the prediction that algae show a UV stress tolerance capacity along a broad depth range according to their marked shade adaptation. The high contents of phlorotannins and antioxidant potential appear to be strongly responsible for the lack of clear depth patterns in light demand characteristics and UV tolerance. PMID:26252953

Lights Off for Arbuscular Mycorrhiza: On Its Symbiotic Functioning under Light Deprivation

PubMed Central

Konvalinková, Tereza; Jansa, Jan

2016-01-01

Plants are often exposed to shade over different time scales and this may substantially affect not only their own growth, but also development and functioning of the energetically dependent organisms. Among those, the root symbionts such as arbuscular mycorrhizal (AM) fungi and rhizobia represent particularly important cases—on the one hand, they consume a significant share of plant carbon (C) budget and, on the other, they generate a number of important nutritional feedbacks on their plant hosts, often resulting in a net positive effect on their host growth and/or fitness. Here we discuss our previous results comparing mycorrhizal performance under different intensities and durations of shade (Konvalinková et al., 2015) in a broader context of previously published literature. Additionally, we review publicly available knowledge on the root colonization and mycorrhizal growth responses in AM plants under light deprivation. Experimental evidence shows that sudden and intensive decrease of light availability to a mycorrhizal plant triggers rapid deactivation of phosphorus transfer from the AM fungus to the plant already within a few days, implying active and rapid response of the AM fungus to the energetic status of its plant host. When AM plants are exposed to intensive shading on longer time scales (weeks to months), positive mycorrhizal growth responses (MGR) are often decreasing and may eventually become negative. This is most likely due to the high C cost of the symbiosis relative to the C availability, and failure of plants to fully compensate for the fungal C demand under low light. Root colonization by AM fungi often declines under low light intensities, although the active role of plants in regulating the extent of root colonization has not yet been unequivocally demonstrated. Quantitative information on the rates and dynamics of C transfer from the plant to the fungus is mostly missing, as is the knowledge on the involved molecular mechanisms. Therefore, these subjects deserve particular attention in the future. PMID:27375642

Lights Off for Arbuscular Mycorrhiza: On Its Symbiotic Functioning under Light Deprivation.

PubMed

Konvalinková, Tereza; Jansa, Jan

2016-01-01

Plants are often exposed to shade over different time scales and this may substantially affect not only their own growth, but also development and functioning of the energetically dependent organisms. Among those, the root symbionts such as arbuscular mycorrhizal (AM) fungi and rhizobia represent particularly important cases-on the one hand, they consume a significant share of plant carbon (C) budget and, on the other, they generate a number of important nutritional feedbacks on their plant hosts, often resulting in a net positive effect on their host growth and/or fitness. Here we discuss our previous results comparing mycorrhizal performance under different intensities and durations of shade (Konvalinková et al., 2015) in a broader context of previously published literature. Additionally, we review publicly available knowledge on the root colonization and mycorrhizal growth responses in AM plants under light deprivation. Experimental evidence shows that sudden and intensive decrease of light availability to a mycorrhizal plant triggers rapid deactivation of phosphorus transfer from the AM fungus to the plant already within a few days, implying active and rapid response of the AM fungus to the energetic status of its plant host. When AM plants are exposed to intensive shading on longer time scales (weeks to months), positive mycorrhizal growth responses (MGR) are often decreasing and may eventually become negative. This is most likely due to the high C cost of the symbiosis relative to the C availability, and failure of plants to fully compensate for the fungal C demand under low light. Root colonization by AM fungi often declines under low light intensities, although the active role of plants in regulating the extent of root colonization has not yet been unequivocally demonstrated. Quantitative information on the rates and dynamics of C transfer from the plant to the fungus is mostly missing, as is the knowledge on the involved molecular mechanisms. Therefore, these subjects deserve particular attention in the future.

Congruence between Spouses’ Perceptions and Observers’ Ratings of Responsiveness: The Role of Attachment Avoidance

PubMed Central

Beck, Lindsey A.; Pietromonaco, Paula R.; DeVito, Cassandra C.; Powers, Sally I.; Boyle, Alysia M.

2016-01-01

Although close relationships require partners to depend on one another for mutual responsiveness, avoidantly-attached individuals are especially averse to risking such dependency. The authors propose that both avoidant and non-avoidant individuals perceive signs of their own and their partners’ responsiveness in ways that reflect motivated perceptions of dependency. The present research examined how the interplay between spouses’ attachment avoidance and observed responsive behaviors during marital conflict shaped perceptions of their own and their partners’ responsiveness. Newlywed couples attempted to resolve a relationship conflict, then reported perceptions of their own and their partners’ responsiveness during the conflict. Observers also coded both partners’ responsive behaviors during the conflict. Avoidant husbands perceived themselves as less responsive than did observers; avoidant wives’ perceptions that their husbands were less responsive matched observers’ ratings. The discussion highlights the role of gender role norms in understanding links between attachment and responsiveness. PMID:24132245

Contrasting impacts of light reduction on sediment biogeochemistry in deep- and shallow-water tropical seagrass assemblages (Green Island, Great Barrier Reef).

PubMed

Schrameyer, Verena; York, Paul H; Chartrand, Kathryn; Ralph, Peter J; Kühl, Michael; Brodersen, Kasper Elgetti; Rasheed, Michael A

2018-05-01

Seagrass meadows increasingly face reduced light availability as a consequence of coastal development, eutrophication, and climate-driven increases in rainfall leading to turbidity plumes. We examined the impact of reduced light on above-ground seagrass biomass and sediment biogeochemistry in tropical shallow- (∼2 m) and deep-water (∼17 m) seagrass meadows (Green Island, Australia). Artificial shading (transmitting ∼10-25% of incident solar irradiance) was applied to the shallow- and deep-water sites for up to two weeks. While above-ground biomass was unchanged, higher diffusive O 2 uptake (DOU) rates, lower O 2 penetration depths, and higher volume-specific O 2 consumption (R) rates were found in seagrass-vegetated sediments as compared to adjacent bare sand (control) areas at the shallow-water sites. In contrast, deep-water sediment characteristics did not differ between bare sand and vegetated sites. At the vegetated shallow-water site, shading resulted in significantly lower hydrogen sulphide (H 2 S) levels in the sediment. No shading effects were found on sediment biogeochemistry at the deep-water site. Overall, our results show that the sediment biogeochemistry of shallow-water (Halodule uninervis, Syringodium isoetifolium, Cymodocea rotundata and C. serrulata) and deep-water (Halophila decipiens) seagrass meadows with different species differ in response to reduced light. The light-driven dynamics of the sediment biogeochemistry at the shallow-water site could suggest the presence of a microbial consortium, which might be stimulated by photosynthetically produced exudates from the seagrass, which becomes limited due to lower seagrass photosynthesis under shaded conditions. Copyright © 2018 Elsevier Ltd. All rights reserved.

Environmental control of reproductive phenology and the effect of pollen supplementation on resource allocation in the cleistogamous weed, Ruellia nudiflora (Acanthaceae).

PubMed

Munguía-Rosas, Miguel A; Parra-Tabla, Victor; Ollerton, Jeff; Cervera, J Carlos

2012-02-01

Mixed reproductive strategies may have evolved as a response of plants to cope with environmental variation. One example of a mixed reproductive strategy is dimorphic cleistogamy, where a single plant produces closed, obligately self-pollinated (CL) flowers and open, potentially outcrossed (CH) flowers. Frequently, optimal environmental conditions favour production of more costly CH structures whilst economical and reliable CL structures are produced under less favourable conditions. In this study we explore (1) the effect of light and water on the reproductive phenology and (2) the effect of pollen supplementation on resource allocation to seeds in the cleistogamous weed Ruellia nudiflora. Split-plot field experiments were carried out to assess the effect of shade (two levels: ambient light vs. a reduction of 50 %) and watering (two levels: non-watered vs. watered) on the onset, end and duration of the production of three reproductive structures: CH flowers, CH fruit and CL fruit. We also looked at the effect of these environmental factors on biomass allocation to seeds (seed weight) from obligately self-pollinated flowers (CL), open-pollinated CH flowers and pollen-supplemented CH flowers. CH structures were produced for a briefer period and ended earlier under shaded conditions. These conditions also resulted in an earlier production of CL fruit. Shaded conditions also produced greater biomass allocation to CH seeds receiving extra pollen. Sub-optimal (shaded) conditions resulted in a briefer production period of CH structures whilst these same conditions resulted in an earlier production of CL structures. However, under sub-optimal conditions, plants also allocated more resources to seeds sired from CH flowers receiving large pollen loads. Earlier production of reproductive structures and relatively larger seed might improve subsequent success of CL and pollen-supplemented CH seeds, respectively.

Radiometer footprint model to estimate sunlit and shaded components for row crops

USDA-ARS?s Scientific Manuscript database

This paper describes a geometric model for computing the relative proportion of sunlit vegetation, shaded vegetation, sunlit soil, and shaded soil appearing in a circular or elliptical radiometer footprint for row crops, where the crop rows were modeled as continuous ellipses. The model was validate...

Shade effects on growth, flowering, and fruit of apple

USDA-ARS?s Scientific Manuscript database

Light is a critical resource needed by plants for growth and reproduction. A major portion of the apple (Malus xdomestica Borkh.) tree’s canopy is subjected to shade during most daylight hours each day and such shade may affect productivity. The current research determined effects of morning, afte...

Shape from Shading in Pigeons

ERIC Educational Resources Information Center

Cook, Robert G.; Qadri, Muhammad A. J.; Kieres, Art; Commons-Miller, Nicholas

2012-01-01

Light is the origin of vision. The pattern of shading reflected from object surfaces is one of several optical features that provide fundamental information about shape and surface orientation. To understand how surface and object shading is processed by birds, six pigeons were tested with differentially illuminated convex and concave curved…

Influence of shading on container-grown flowering dogwoods

USDA-ARS?s Scientific Manuscript database

Bare root dogwoods can be successfully grown when transplanted into a container production system. Shade treatments regardless of color or density did have an effect on the plant growth of Cherokee Brave™ and Cherokee Princess dogwood. Plants grown under 50% black and 50% white shade had more heigh...

PowerFilm PowerShade Fixed Site Solar System Cost Reduction Plan

DTIC Science & Technology

2014-07-31

2337 230’^ St Ames, lA 50014 JL’Olt^lO:pi𔄁oq7 j pageii Executive summary Power film and its partners have developed a Generation II PowerShade...which meets or exceeds the goals of this development contract, increasing power and improving the ROI of the power shade significantly while improving a...number of the human factors. Comparing the Gen II 1.8 KW PowerShade to the equivalent power Gen I unit (Gen I Medium), The cost has been reduced

Geometric and shading correction for images of printed materials using boundary.

PubMed

Brown, Michael S; Tsoi, Yau-Chat

2006-06-01

A novel technique that uses boundary interpolation to correct geometric distortion and shading artifacts present in images of printed materials is presented. Unlike existing techniques, our algorithm can simultaneously correct a variety of geometric distortions, including skew, fold distortion, binder curl, and combinations of these. In addition, the same interpolation framework can be used to estimate the intrinsic illumination component of the distorted image to correct shading artifacts. We detail our algorithm for geometric and shading correction and demonstrate its usefulness on real-world and synthetic data.

Scale model testing of drogues for free drifting buoys

NASA Technical Reports Server (NTRS)

Vachon, W. A.

1973-01-01

Instrumented model drogue tests were conducted in a ship model towing tank. The purpose of the tests was to observe and measure deployment and drag characteristics of such shapes as parachutes, crossed vanes, and window shades which may be employed in conjunction with free drifting buoys. Both Froude and Reynolds scaling laws were applied while scaling to full scale relative velocities of from 0 to 0.2 knots. A weighted window shade drogue is recommended because of its performance, high drag coefficient, simplicity, and low cost. Detailed theoretical performance curves are presented for parachutes, crossed vanes, and window shade drogues. Theoretical estimates of depth locking accuracy and buoy-induced dynamic loads pertinent to window shade drogues are presented as a design aid. An example of a window shade drogue design is presented.

Aircraft geometry verification with enhanced computer generated displays

NASA Technical Reports Server (NTRS)

Cozzolongo, J. V.

1982-01-01

A method for visual verification of aerodynamic geometries using computer generated, color shaded images is described. The mathematical models representing aircraft geometries are created for use in theoretical aerodynamic analyses and in computer aided manufacturing. The aerodynamic shapes are defined using parametric bi-cubic splined patches. This mathematical representation is then used as input to an algorithm that generates a color shaded image of the geometry. A discussion of the techniques used in the mathematical representation of the geometry and in the rendering of the color shaded display is presented. The results include examples of color shaded displays, which are contrasted with wire frame type displays. The examples also show the use of mapped surface pressures in terms of color shaded images of V/STOL fighter/attack aircraft and advanced turboprop aircraft.

Color Compatibility of Gingival Shade Guides and Gingiva-Colored Dental Materials with Healthy Human Gingiva.

PubMed

Sarmast, Nima D; Angelov, Nikola; Ghinea, Razvan; Powers, John M; Paravina, Rade D

The CIELab and CIEDE2000 coverage error (ΔE* COV and ΔE' COV , respectively) of basic shades of different gingival shade guides and gingiva-colored restorative dental materials (n = 5) was calculated as compared to a previously compiled database on healthy human gingiva. Data were analyzed using analysis of variance with Tukey-Kramer multiple-comparison test (P < .05). A 50:50% acceptability threshold of 4.6 for ΔE* and 4.1 for ΔE' was used to interpret the results. ΔE* COV / ΔE' COV ranged from 4.4/3.5 to 8.6/6.9. The majority of gingival shade guides and gingiva-colored restorative materials exhibited statistically significant coverage errors above the 50:50% acceptability threshold and uneven shade distribution.

Accuracy of shade matching performed by colour blind and normal dental students using 3D Master and Vita Lumin shade guides.

PubMed

Vafaee, F; Rakhshan, V; Vafaei, M; Khoshhal, M

2012-03-01

The purpose of this study was to investigate whether 3D Master or VitaLumin shade guides could improve colour selection in individuals with normal and defective colour vision. First, colour perception of 260 dental students was evaluated. Afterwards, 9 colour blind and 9 matched normal subjects tried to detect colours of 10 randomly selected tabs from each kit and the correct/false answers were counted. Of the colour-defective subjects, 47.8% and 33.3% correctly detected the shade using 3D Master and VitaLumin, respectively. These statistics were 62.2% and 42.2% in normal subjects. In normal participants, but not in colour blind ones, 3D Master significantly improved shade matching accuracy compared to VitaLumin.

A biological timer in the fat body comprising Blimp-1, βFtz-f1 and Shade regulates pupation timing in Drosophila melanogaster.

PubMed

Akagi, Kazutaka; Sarhan, Moustafa; Sultan, Abdel-Rahman S; Nishida, Haruka; Koie, Azusa; Nakayama, Takumi; Ueda, Hitoshi

2016-07-01

During the development of multicellular organisms, many events occur with precise timing. In Drosophila melanogaster, pupation occurs about 12 h after puparium formation and its timing is believed to be determined by the release of a steroid hormone, ecdysone (E), from the prothoracic gland. Here, we demonstrate that the ecdysone-20-monooxygenase Shade determines pupation timing by converting E to 20-hydroxyecdysone (20E) in the fat body, which is the organ that senses nutritional status. The timing of shade expression is determined by its transcriptional activator βFtz-f1. The βftz-f1 gene is activated after a decline in the expression of its transcriptional repressor Blimp-1, which is temporally expressed around puparium formation in response to a high titer of 20E. The expression level and stability of Blimp-1 is critical for the precise timing of pupation. Thus, we propose that Blimp-1 molecules function like sand in an hourglass in this precise developmental timer system. Furthermore, our data suggest that a biological advantage results from both the use of a transcriptional repressor for time determination and the association of developmental timing with nutritional status of the organism. © 2016. Published by The Company of Biologists Ltd.

Reliability and accuracy of Crystaleye spectrophotometric system.

PubMed

Chen, Li; Tan, Jian Guo; Zhou, Jian Feng; Yang, Xu; Du, Yang; Wang, Fang Ping

2010-01-01

to develop an in vitro shade-measuring model to evaluate the reliability and accuracy of the Crystaleye spectrophotometric system, a newly developed spectrophotometer. four shade guides, VITA Classical, VITA 3D-Master, Chromascop and Vintage Halo NCC, were measured with the Crystaleye spectrophotometer in a standardised model, ten times for 107 shade tabs. The shade-matching results and the CIE L*a*b* values of the cervical, body and incisal regions for each measurement were automatically analysed using the supporting software. Reliability and accuracy were calculated for each shade tab both in percentage and in colour difference (ΔE). Difference was analysed by one-way ANOVA in the cervical, body and incisal regions. range of reliability was 88.81% to 98.97% and 0.13 to 0.24 ΔE units, and that of accuracy was 44.05% to 91.25% and 1.03 to 1.89 ΔE units. Significant differences in reliability and accuracy were found between the body region and the cervical and incisal regions. Comparisons made among regions and shade guides revealed that evaluation in ΔE was prone to disclose the differences. measurements with the Crystaleye spectrophotometer had similar, high reliability in different shade guides and regions, indicating predictable repeated measurements. Accuracy in the body region was high and less variable compared with the cervical and incisal regions.

Increased light-use efficiency sustains net primary productivity of shaded coffee plants in agroforestry system.

PubMed

Charbonnier, Fabien; Roupsard, Olivier; le Maire, Guerric; Guillemot, Joannès; Casanoves, Fernando; Lacointe, André; Vaast, Philippe; Allinne, Clémentine; Audebert, Louise; Cambou, Aurélie; Clément-Vidal, Anne; Defrenet, Elsa; Duursma, Remko A; Jarri, Laura; Jourdan, Christophe; Khac, Emmanuelle; Leandro, Patricia; Medlyn, Belinda E; Saint-André, Laurent; Thaler, Philippe; Van Den Meersche, Karel; Barquero Aguilar, Alejandra; Lehner, Peter; Dreyer, Erwin

2017-08-01

In agroforestry systems, shade trees strongly affect the physiology of the undergrown crop. However, a major paradigm is that the reduction in absorbed photosynthetically active radiation is, to a certain extent, compensated by an increase in light-use efficiency, thereby reducing the difference in net primary productivity between shaded and non-shaded plants. Due to the large spatial heterogeneity in agroforestry systems and the lack of appropriate tools, the combined effects of such variables have seldom been analysed, even though they may help understand physiological processes underlying yield dynamics. In this study, we monitored net primary productivity, during two years, on scales ranging from individual coffee plants to the entire plot. Absorbed radiation was mapped with a 3D model (MAESPA). Light-use efficiency and net assimilation rate were derived for each coffee plant individually. We found that although irradiance was reduced by 60% below crowns of shade trees, coffee light-use efficiency increased by 50%, leaving net primary productivity fairly stable across all shade levels. Variability of aboveground net primary productivity of coffee plants was caused primarily by the age of the plants and by intraspecific competition among them (drivers usually overlooked in the agroforestry literature) rather than by the presence of shade trees. © 2017 John Wiley & Sons Ltd.

Shading Vita YZ substructures: influence on value and chroma, part I.

PubMed

Devigus, A; Lombardi, G

2004-07-01

All-ceramic restorations should reproduce as well as possible the color of the natural teeth to create a restoration in accordance with the esthetic wishes and ideas of the patient. The basic color of zirconium oxide is white to ivory. The color can be partially adapted by veneering it with ceramic materials. However, it would be better if the substructure could already be adapted to the basic color shade of the neighboring teeth. In this study, the influence of differently shaded frameworks made of Y-TZP by Vita and 3M ESPE and by Enrico Steger (without, with 0.5 mm, or with 1.0 mm ceramic veneer with Base Dentin) on the brightness, saturation, and color shade (= value, chroma, and hue) was measured and assessed with the aid of a spectral photometer (EasyShade, Vita) in a clinical case in the mouth and on the model. By adaptation to the basic shade, the shading of substructures made of Y-TZP can help to reduce the necessary layer thickness of the veneer ceramic to achieve the desired color and should be performed in the future as a matter of routine. In this way, more substance can be conserved when restoring the teeth without having to accept an impairment of the esthetic result.

Assessing the influence of skin color and tooth shade value on perceived smile attractiveness.

PubMed

Sabherwal, Ruchika S; Gonzalez, Juan; Naini, Farhad B

2009-06-01

The authors conducted a study to determine whether variations in skin color would influence perceptions of smile attractiveness for a given tooth shade value. Using a photograph of a woman smiling, the authors altered skin color to consist of four tones (fair, fair/medium, medium/dark and dark) and altered tooth shade value (brightness) to range from 00 (brightest) to 05 (darkest). Two groups of judges (70 dentists, 70 laypeople) completed a questionnaire and rated the images for smile attractiveness. For most tooth shade values (00, 01, 03, 04, 05), multiple mixed linear regression showed that variation in skin color influenced respondents' perceptions of smile attractiveness (P < .001). For images corresponding to tooth shade values 01, 02, 03, 04 and 05, men provided lower ratings than did women (P < .05). Attractiveness ratings increased with the age of participants (P < .05). Dentists rated images higher than did laypeople (P < .05). Variation in skin color for most tooth shade values influenced dentists' and laypeople's perceived smile attractiveness. Dentists and laypeople did not perceive the brightest tooth shade to be the most attractive, and they did not perceive all skin colors to be equally attractive with bright white teeth. Respondents perceived dark skin with bright white teeth and fair skin with dark teeth as relatively unattractive.

[Comparison of the color difference between teeth underwent cold light whitening and two kinds of shade guides].

PubMed

Xu, Y X

2018-06-18

To investigate which shade guide, Vitapan Classical or Vita Bleachedguide 3DMaster, is better matched with the color of teeth in judging whitening effect, by comparing the color difference between shade tabs and corresponding teeth underwent cold light tooth whitening. A total of 60 patients underwent Beyond cold light tooth whitening from May 2014 to April 2016. The patients were divided into two experimental groups according to the shade guide used. Vitapan Classical shade guide was used to judge whitening effect in one group, and Vita Bleachedguide 3DMaster shade guide was used in another. Shade matching was carried out before and after whitening in both the two groups, and the results were recorded by digital photographs. Shade matching procedures were carried out by two doctors independently. If they chose the same tab, it would be seen as the shade matching result; While if they chose different tabs, another doctor would be invited to make a decision. Photographs were taken in preset conditions: intraoral photos of the full dentition in the front, and the proportion of shooting was 1:3; aperture was F22; shutter speed was 1/200; intensity of flash was M/8; ISO value was 200. The photographs were analyzed by Photoshop software. Chromatic values were measured, and color difference values were calculated. Measuring of chromatic values was carried out by three doctors independently, and all the photos were measured twice by each doctor. Six measure results of each photo were recorded, and the maximum and the minimum were excluded, then the mean was seen as the final result. The color difference values were compared by independent-sample t test. Besides, changes of shade tabs after whitening in the two groups were recorded. Color difference value was 5.06±1.71 in Vitapan Classical group, and 3.39±1.36 in Vita Bleachedguide 3D-Master group. There was statistically significant difference between the two groups (t=4.68,P<0.001). Change of shade tabs was 3.63±1.75 in Vitapan Classical group, and 2.23±1.01 in Vita Bleachedguide 3DMaster group. Vita Bleachedguide 3D-Master is better matched with the color of teeth, so it is preferred in judging the effect of cold light tooth whitening.

Shade matching performance of normal and color vision-deficient dental professionals with standard daylight and tungsten illuminants.

PubMed

Gokce, Hasan Suat; Piskin, Bulent; Ceyhan, Dogan; Gokce, Sila Mermut; Arisan, Volkan

2010-03-01

The lighting conditions of the environment and visual deficiencies such as red-green color vision deficiency affect the clinical shade matching performance of dental professionals. The purpose of this study was to evaluate the shade matching performance of normal and color vision-deficient dental professionals with standard daylight and tungsten illuminants. Two sets of porcelain disc replicas of 16 shade guide tabs (VITA Lumin) were manufactured to exact L*a*b* values by using a colorimeter. Then these twin porcelain discs (13 mm x 2.4 mm) were mixed up and placed into a color-matching cabinet that standardized the lighting conditions for the observation tests. Normal and red-green color vision-deficient dental professionals were asked to match the 32 porcelain discs using standard artificial daylight D65 (high color temperature) and tungsten filament lamp light (T) (low color temperature) illuminants. The results were analyzed by repeated-measures ANOVA and paired and independent samples t tests for the differences between dental professionals and differences between the illuminants (alpha=.05). Regarding the sum of the correct shade match scores of all observations with both illuminants, the difference between normal vision and red-green color vision-deficient dental professional groups was not statistically significant (F=4.132; P=.054). However, the correct shade match scores of each group were significantly different for each illuminant (P<.005). The correct shade matching scores of normal color vision dental professionals were significantly higher with D65 illuminant (t=7.004; P<.001). Color matching scores of red-green color vision-deficient dental professionals (approximately 5.7 more pairs than with D65) were significantly higher with T illuminant (t=5.977; P<.001). CONCLUSIONS.: Within the limitations of this study, the shade matching performance of dental professionals was affected by color vision deficiency and the color temperature of the illuminant. The color vision-deficient group was notably unsuccessful with the D65 illuminant in shade matching. In contrast, there was a significant increase in the shade matching performance of the color vision-deficient group with T illuminant. The lower color temperature illuminant dramatically decreased the normal color vision groups' correct shade matching score. (c) 2010 The Editorial Council of the Journal of Prosthetic Dentistry. Published by Mosby, Inc. All rights reserved.

Mixed-conifer forests of central Oregon: effects of logging and fire exclusion vary with environment.

PubMed

Merschel, Andrew G; Spies, Thomas A; Heyerdahl, Emily K

Twentieth-century land management has altered the structure and composition of mixed-conifer forests and decreased their resilience to fire, drought, and insects in many parts of the Interior West. These forests occur across a wide range of environmental settings and historical disturbance regimes, so their response to land management is likely to vary across landscapes and among ecoregions. However, this variation has not been well characterized and hampers the development of appropriate management and restoration plans. We identified mixed-conifer types in central Oregon based on historical structure and composition, and successional trajectories following recent changes in land use, and evaluated how these types were distributed across environmental gradients. We used field data from 171 sites sampled across a range of environmental settings in two subregions: the eastern Cascades and the Ochoco Mountains. We identified four forest types in the eastern Cascades and four analogous types with lower densities in the Ochoco Mountains. All types historically contained ponderosa pine, but differed in the historical and modern proportions of shade-tolerant vs. shade-intolerant tree species. The Persistent Ponderosa Pine and Recent Douglas-fir types occupied relatively hot–dry environments compared to Recent Grand Fir and Persistent Shade Tolerant sites, which occupied warm–moist and cold–wet environments, respectively. Twentieth-century selective harvesting halved the density of large trees, with some variation among forest types. In contrast, the density of small trees doubled or tripled early in the 20th century, probably due to land-use change and a relatively cool, wet climate. Contrary to the common perception that dry ponderosa pine forests are the most highly departed from historical conditions, we found a greater departure in the modern composition of small trees in warm–moist environments than in either hot–dry or cold–wet environments. Furthermore, shade-tolerant trees began infilling earlier in cold–wet than in hot–dry environments and also in topographically shaded sites in the Ochoco Mountains. Our new classification could be used to prioritize management that seeks to restore structure and composition or create resilience in mixed-conifer forests of the region.

Drought tolerance associated with vertical stratification of two co-occurring epiphytic bromeliads in a tropical dry forest.

PubMed

Graham, Eric A; Andrade, Jose Luis

2004-05-01

Vertical stratification of epiphytes generally has not been reported for dry forests. For two epiphytic Crassulacean acid metabolism bromeliads that segregate vertically, it was hypothesized that different potentials for photoprotection or shade tolerance rather than drought tolerance is responsible for the observed stratification. The light environment, capacity for photoprotection, germination response to light quality, and responses to light and drought were thus examined for Tillandsia brachycaulos and T. elongata. Vertical and light-environment distributions differed for the two species but photoprotection and photodamage did not where they occurred at similar field locations; T. brachycaulos had a higher pigment acclimation to light. Tillandsia brachycaulos had higher acid accumulation under low light as opposed to T. elongata, which responded similarly to all but the highest light treatment. Tillandsia brachycaulos maintained positive total daily net CO(2) uptake through 30 d of drought; T. elongata had a total daily net CO(2) loss after 7 d of drought. The vertical stratification was most likely the result of the sensitivity to drought of T. elongata rather than differences in photoprotection or shade tolerance between the two species. Tillandsia elongata occurs in more exposed locations, which may be advantageous for rainfall interception and dew formation.

Fluorescence responses and photosynthetic rates of sunlit and shaded leaves of Italian alpine forest species: Summer 1997 ATOM-LIFT campaign

NASA Astrophysics Data System (ADS)

Kim, Moon S.; Cecchi, Giovanna; Chappelle, Emmett W.; Bazzani, Marco; McMurtrey, James E., III; Corp, Lawrence A.; Sandu, R.; Tirelli, Daniele

1998-07-01

Terrestrial vegetation studies were carried out in the Italian Northeastern Alps in Val Visdende. The measurement site was 15 Kilometers Northeast of the town of St. Stefano di Calore (Belluno), Italy. Measurements were acquired on a wooded site at the Italian Department of Forestry Station on species native to the Italian Alps. The species included spruce (Picea abies) and alder (Alnus incana) trees. Characterization was also made of the fluorescence responses of several under-story species such as Dactylorhiza fuchsii of the Orchidaceae family, Caltha palustris and Ranunculus ficaria of the Ranuncolcee family, and Trifolium pratense and Trifolium repens of the Leguminosae family. Terrestrial vegetation monitoring was conducted with the Italian FLIDAR remote sensing instrument mounted in a mobile van, the NASA/USDA Fluorescence Imaging System (FIS), and the Spectron SE-590 for optical properties. Photosynthetic CO2 gas exchange rates we made with LI-COR 6400 infrared gas analyzer. Pigments from the samples were extracted and analyzed with a Perkin Elmer Lamda 7 Spectrometer to determine pigment concentrations. Fluorescence responses were collected from vegetation samples grown under different ambient light regimes of sun-lit versus shaded. The vegetation showed different fluorescence characteristics. A fluorescence algorithm, (F740/F680)/F550, and rate of photosynthesis showed a strong linear relationship.

Light-mediated polarization of the PIN3 auxin transporter for the phototropic response in Arabidopsis.

PubMed

Ding, Zhaojun; Galván-Ampudia, Carlos S; Demarsy, Emilie; Łangowski, Łukasz; Kleine-Vehn, Jürgen; Fan, Yuanwei; Morita, Miyo T; Tasaka, Masao; Fankhauser, Christian; Offringa, Remko; Friml, Jiří

2011-04-01

Phototropism is an adaptation response, through which plants grow towards the light. It involves light perception and asymmetric distribution of the plant hormone auxin. Here we identify a crucial part of the mechanism for phototropism, revealing how light perception initiates auxin redistribution that leads to directional growth. We show that light polarizes the cellular localization of the auxin efflux carrier PIN3 in hypocotyl endodermis cells, resulting in changes in auxin distribution and differential growth. In the dark, high expression and activity of the PINOID (PID) kinase correlates with apolar targeting of PIN3 to all cell sides. Following illumination, light represses PINOID transcription and PIN3 is polarized specifically to the inner cell sides by GNOM ARF GTPase GEF (guanine nucleotide exchange factor)-dependent trafficking. Thus, differential trafficking at the shaded and illuminated hypocotyl side aligns PIN3 polarity with the light direction, and presumably redirects auxin flow towards the shaded side, where auxin promotes growth, causing hypocotyls to bend towards the light. Our results imply that PID phosphorylation-dependent recruitment of PIN proteins into distinct trafficking pathways is a mechanism to polarize auxin fluxes in response to different environmental and endogenous cues. © 2011 Macmillan Publishers Limited. All rights reserved

Characterization of cocoa production, income diversification and shade tree management along a climate gradient in Ghana.

PubMed

Abdulai, Issaka; Jassogne, Laurence; Graefe, Sophie; Asare, Richard; Van Asten, Piet; Läderach, Peter; Vaast, Philippe

2018-01-01

Reduced climatic suitability due to climate change in cocoa growing regions of Ghana is expected in the coming decades. This threatens farmers' livelihood and the cocoa sector. Climate change adaptation requires an improved understanding of existing cocoa production systems and farmers' coping strategies. This study characterized current cocoa production, income diversification and shade tree management along a climate gradient within the cocoa belt of Ghana. The objectives were to 1) compare existing production and income diversification between dry, mid and wet climatic regions, and 2) identify shade trees in cocoa agroforestry systems and their distribution along the climatic gradient. Our results showed that current mean cocoa yield level of 288kg ha-1yr-1 in the dry region was significantly lower than in the mid and wet regions with mean yields of 712 and 849 kg ha-1 yr-1, respectively. In the dry region, farmers diversified their income sources with non-cocoa crops and off-farm activities while farmers at the mid and wet regions mainly depended on cocoa (over 80% of annual income). Two shade systems classified as medium and low shade cocoa agroforestry systems were identified across the studied regions. The medium shade system was more abundant in the dry region and associated to adaptation to marginal climatic conditions. The low shade system showed significantly higher yield in the wet region but no difference was observed between the mid and dry regions. This study highlights the need for optimum shade level recommendation to be climatic region specific.

Effect of ceramic thickness and cement shade on the final shade after bonding using the 3D master system: a laboratory study.

PubMed

Montero, Javier; Gómez-Polo, Cristina

2016-06-01

The final color of a ceramic restoration is influenced by both the ceramic thickness and the cement shade. This study aims to evaluate the color stability according to the 3D Master System of e.max ceramic discs after bonding with different shades of luting agents. A total of 120 e.max.Press 2M1 HT ceramic discs (60 discs of 1-mm thick and 60 discs of 0.5 mm thick) and three different values of Variolink Veneer cement were used (-3, 0, +3) for the cementation process. An Easyshade compact device was used to measure color shade tabs, according to the 3D Master System, on the discs both before and after the cementation protocols. Bivariate and multivariate analyses were carried out with the spss v.21. After bonding with the different luting agents, only 30% remained as 2M1: specifically, 22% of the thinner discs and 37.3% of the thicker discs. In general, the effect of bonding increased the value and the chroma of the shade to a significant extent. Regression analyses revealed that the most significant predictor for all color parameters was cement shade, the thinner disc group bonded with -3 cement being the most unstable subgroup. According to the 3D Master System, the shade of the luting agent was the main predictor of the final color. However, the final color seems to be somewhat unpredictable, at least according to the modulating factors evaluated in the present study.

Characterization of cocoa production, income diversification and shade tree management along a climate gradient in Ghana

PubMed Central

Jassogne, Laurence; Graefe, Sophie; Asare, Richard; Van Asten, Piet; Läderach, Peter; Vaast, Philippe

2018-01-01

Reduced climatic suitability due to climate change in cocoa growing regions of Ghana is expected in the coming decades. This threatens farmers’ livelihood and the cocoa sector. Climate change adaptation requires an improved understanding of existing cocoa production systems and farmers’ coping strategies. This study characterized current cocoa production, income diversification and shade tree management along a climate gradient within the cocoa belt of Ghana. The objectives were to 1) compare existing production and income diversification between dry, mid and wet climatic regions, and 2) identify shade trees in cocoa agroforestry systems and their distribution along the climatic gradient. Our results showed that current mean cocoa yield level of 288kg ha-1yr-1 in the dry region was significantly lower than in the mid and wet regions with mean yields of 712 and 849 kg ha-1 yr-1, respectively. In the dry region, farmers diversified their income sources with non-cocoa crops and off-farm activities while farmers at the mid and wet regions mainly depended on cocoa (over 80% of annual income). Two shade systems classified as medium and low shade cocoa agroforestry systems were identified across the studied regions. The medium shade system was more abundant in the dry region and associated to adaptation to marginal climatic conditions. The low shade system showed significantly higher yield in the wet region but no difference was observed between the mid and dry regions. This study highlights the need for optimum shade level recommendation to be climatic region specific. PMID:29659629

DOE Office of Scientific and Technical Information (OSTI.GOV)

Deline, Chris; Meydbray, Jenya; Donovan, Matt

The 2012 NREL report 'Photovoltaic Shading Testbed for Module-Level Power Electronics' provides a standard methodology for estimating the performance benefit of distributed power electronics under partial shading conditions. Since the release of the report, experiments have been conducted for a number of products and for different system configurations. Drawing from these experiences, updates to the test and analysis methods are recommended. Proposed changes in data processing have the benefit of reducing the sensitivity to measurement errors and weather variability, as well as bringing the updated performance score in line with measured and simulated values of the shade recovery benefit ofmore » distributed PV power electronics. Also, due to the emergence of new technologies including sub-module embedded power electronics, the shading method has been extended to include power electronics that operate at a finer granularity than the module level. An update to the method is proposed to account for these emerging technologies that respond to shading differently than module-level devices. The partial shading test remains a repeatable test procedure that attempts to simulate shading situations as would be experienced by typical residential or commercial rooftop photovoltaic (PV) systems. Performance data for multiple products tested using this method are discussed, based on equipment from Enphase, Solar Edge, Maxim Integrated and SMA. In general, the annual recovery of shading losses from the module-level electronics evaluated is 25-35%, with the major difference between different trials being related to the number of parallel strings in the test installation rather than differences between the equipment tested. Appendix D data has been added in this update.« less

Shade Tree Diversity, Cocoa Pest Damage, Yield Compensating Inputs and Farmers' Net Returns in West Africa

PubMed Central

Daghela Bisseleua, Hervé Bertin; Fotio, Daniel; Yede; Missoup, Alain Didier; Vidal, Stefan

2013-01-01

Cocoa agroforests can significantly support biodiversity, yet intensification of farming practices is degrading agroforestry habitats and compromising ecosystem services such as biological pest control. Effective conservation strategies depend on the type of relationship between agricultural matrix, biodiversity and ecosystem services, but to date the shape of this relationship is unknown. We linked shade index calculated from eight vegetation variables, with insect pests and beneficial insects (ants, wasps and spiders) in 20 cocoa agroforests differing in woody and herbaceous vegetation diversity. We measured herbivory and predatory rates, and quantified resulting increases in cocoa yield and net returns. We found that number of spider webs and wasp nests significantly decreased with increasing density of exotic shade tree species. Greater species richness of native shade tree species was associated with a higher number of wasp nests and spider webs while species richness of understory plants did not have a strong impact on these beneficial species. Species richness of ants, wasp nests and spider webs peaked at higher levels of plant species richness. The number of herbivore species (mirid bugs and cocoa pod borers) and the rate of herbivory on cocoa pods decreased with increasing shade index. Shade index was negatively related to yield, with yield significantly higher at shade and herb covers<50%. However, higher inputs in the cocoa farms do not necessarily result in a higher net return. In conclusion, our study shows the importance of a diverse shade canopy in reducing damage caused by cocoa pests. It also highlights the importance of conservation initiatives in tropical agroforestry landscapes. PMID:23520451

Comparisons of Photosynthetic Responses of Xanthium strumarium and Helianthus annuus to Chronic and Acute Water Stress in Sun and Shade 1

PubMed Central

Ben, Gui-Ying; Osmond, C. Barry; Sharkey, Thomas D.

1987-01-01

We have examined the effects of mild, chronic water stress and acute water stress on two water stress sensitive plants, Xanthium strumarium and Helianthus annuus. Using a combination of the leaf disc O2 electrode to measure the light responses of photosynthesis and 77 K fluorescence to monitor damage to the primary photochemistry, we have found the following: (a) The CO2 saturated rate of photosynthesis at high light is the most water stress sensitive parameter measured. (b) The apparent quantum yield (moles O2 per mole photons) was slightly, if at all, affected by mild water stress (>−1.5 megapascals). (c) Severe water stress (<−1.5 megapascals) reduced the quantum yield of photosynthesis regardless of whether the stress was applied in sun or shade. The light independent reduction of quantum yield was not associated with a reduction in 77 K fluorescence (Fv/Fm) indicating that the quantum yield reduction was not the result of damage to primary photochemistry. (d) The diel fluctuation in 77 K fluorescence seen in sun-exposed control leaves was greatly exaggerated in water stressed leaves because of enhanced decline in 77 K fluorescence in the morning. The rate of recovery was similar in both control and water stressed leaves. Shaded leaves showed no change in 77 K fluorescence regardless of whether water stress was imposed or not. (e) The water stress sensitive plants used in these experiments did not recover from acute water stress severe enough to reduce the quantum yield or chronic water stress which lasted long enough that light dependent damage to primary photochemistry occurred. PMID:16665465

Comparisons of Photosynthetic Responses of Xanthium strumarium and Helianthus annuus to Chronic and Acute Water Stress in Sun and Shade.

PubMed

Ben, G Y; Osmond, C B; Sharkey, T D

1987-06-01

We have examined the effects of mild, chronic water stress and acute water stress on two water stress sensitive plants, Xanthium strumarium and Helianthus annuus. Using a combination of the leaf disc O(2) electrode to measure the light responses of photosynthesis and 77 K fluorescence to monitor damage to the primary photochemistry, we have found the following: (a) The CO(2) saturated rate of photosynthesis at high light is the most water stress sensitive parameter measured. (b) The apparent quantum yield (moles O(2) per mole photons) was slightly, if at all, affected by mild water stress (>-1.5 megapascals). (c) Severe water stress (<-1.5 megapascals) reduced the quantum yield of photosynthesis regardless of whether the stress was applied in sun or shade. The light independent reduction of quantum yield was not associated with a reduction in 77 K fluorescence (F(v)/F(m)) indicating that the quantum yield reduction was not the result of damage to primary photochemistry. (d) The diel fluctuation in 77 K fluorescence seen in sun-exposed control leaves was greatly exaggerated in water stressed leaves because of enhanced decline in 77 K fluorescence in the morning. The rate of recovery was similar in both control and water stressed leaves. Shaded leaves showed no change in 77 K fluorescence regardless of whether water stress was imposed or not. (e) The water stress sensitive plants used in these experiments did not recover from acute water stress severe enough to reduce the quantum yield or chronic water stress which lasted long enough that light dependent damage to primary photochemistry occurred.

Mesophyll conductance plays a central role in leaf functioning of Oleaceae species exposed to contrasting sunlight irradiance.

PubMed

Fini, Alessio; Loreto, Francesco; Tattini, Massimiliano; Giordano, Cristiana; Ferrini, Francesco; Brunetti, Cecilia; Centritto, Mauro

2016-05-01

The ability to modify mesophyll conductance (gm ) in response to changes in irradiance may be a component of the acclimation of plants to shade-sun transitions, thus influencing species-specific distributions along light-gradients, and the ecological niches for the different species. To test this hypothesis we grew three woody species of the Oleaceae family, the evergreen Phillyrea latifolia (sun-requiring), the deciduous Fraxinus ornus (facultative sun-requiring) and the hemi-deciduous Ligustrum vulgare (shade tolerant) at 30 or 100% sunlight irradiance. We show that neither mesophyll conductance calculated with combined gas exchange and chlorophyll fluorescence techniques (gm) nor CO2 assimilation significantly varied in F. ornus because of sunlight irradiance. This corroborates previous suggestions that species with high plasticity for light requirements, do not need to undertake extensive reorganization of leaf conductances to CO2 diffusion to adapt to different light environments. On the other hand, gm steeply declined in L. vulgare and increased in P. latifolia exposed to full-sun conditions. In these two species, leaf anatomical traits are in part responsible for light-driven changes in gm , as revealed by the correlation between gm and mesophyll conductance estimated by anatomical parameters (gmA). Nonetheless, gm was greatly overestimated by gmA when leaf metabolism was impaired because of severe light stress. We show that gm is maximum at the light intensity at which plant species have evolved and we conclude that gm actually plays a key role in the sun and shade adaptation of Mediterranean species. The limits of gmA in predicting mesophyll conductance are also highlighted. © 2015 Scandinavian Plant Physiology Society.

Impacts of Sediments on Coral Energetics: Partitioning the Effects of Turbidity and Settling Particles

PubMed Central

Junjie, Reef K.; Browne, Nicola K.; Erftemeijer, Paul L. A.; Todd, Peter A.

2014-01-01

Sediment loads have long been known to be deleterious to corals, but the effects of turbidity and settling particles have not previously been partitioned. This study provides a novel approach using inert silicon carbide powder to partition and quantify the mechanical effects of sediment settling versus reduced light under a chronically high sedimentary regime on two turbid water corals commonly found in Singapore (Galaxea fascicularis and Goniopora somaliensis). Coral fragments were evenly distributed among three treatments: an open control (30% ambient PAR), a shaded control (15% ambient PAR) and sediment treatment (15% ambient PAR; 26.4 mg cm−2 day−1). The rate of photosynthesis and respiration, and the dark-adapted quantum yield were measured once a week for four weeks. By week four, the photosynthesis to respiration ratio (P/R ratio) and the photosynthetic yield (Fv/Fm) had fallen by 14% and 3–17% respectively in the shaded control, contrasting with corals exposed to sediments whose P/R ratio and yield had declined by 21% and 18–34% respectively. The differences in rates between the shaded control and the sediment treatment were attributed to the mechanical effects of sediment deposition. The physiological response to sediment stress differed between species with G. fascicularis experiencing a greater decline in the net photosynthetic yield (13%) than G. somaliensis (9.5%), but a smaller increase in the respiration rates (G. fascicularis = 9.9%, G. somaliensis = 14.2%). These different physiological responses were attributed, in part, to coral morphology and highlighted key physiological processes that drive species distribution along high to low turbidity and depositional gradients. PMID:25197883

Depth-Related Effects on a Meiofaunal Community Dwelling in the Periphyton of a Mesotrophic Lake

PubMed Central

Kreuzinger-Janik, Bianca; Schroeder, Fabian; Majdi, Nabil; Traunspurger, Walter

2015-01-01

Periphyton is a complex assemblage of micro- and meiofauna embedded in the organic matrix that coats most submerged substrate in the littoral of lakes. The aim of this study was to better understand the consequences of depth-level fluctuation on a periphytic community. The effects of light and wave disturbance on the development of littoral periphyton were evaluated in Lake Erken (Sweden) using an experimental design that combined in situ shading with periphyton depth transfers. Free-living nematodes were a major contributor to the meiofaunal community. Their species composition was therefore used as a proxy to distinguish the contributions of light- and wave-related effects. The periphyton layer was much thicker at a depth of 30 cm than at 200 cm, as indicated by differences in the amounts of organic and phototrophic biomass and meiofaunal and nematode densities. A reduction of the depth-level of periphyton via a transfer from a deep to a shallow location induced rapid positive responses by its algal, meiofaunal, and nematode communities. The slower and weaker negative responses to the reverse transfer were attributed to the potentially higher resilience of periphytic communities to increases in the water level. In the shallow littoral of the lake, shading magnified the effects of phototrophic biomass erosion by waves, as the increased exposure to wave shear stress was not compensated for by an increase in photosynthesis. This finding suggests that benthic primary production will be strongly impeded in the shallow littoral zones of lakes artificially shaded by construction or embankments. However, regardless of the light constraints, an increased exposure to wave action had a generally positive short-term effect on meiofaunal density, by favoring the predominance of species able to anchor themselves to the substrate, especially the Chromadorid nematode Punctodora ratzeburgensis. PMID:26353016

29 CFR 780.506 - Dependence of exemption on shade-grown tobacco operations.

Code of Federal Regulations, 2010 CFR

2010-07-01

... 29 Labor 3 2010-07-01 2010-07-01 false Dependence of exemption on shade-grown tobacco operations. 780.506 Section 780.506 Labor Regulations Relating to Labor (Continued) WAGE AND HOUR DIVISION....506 Dependence of exemption on shade-grown tobacco operations. The exemption provided by section 13(a...

29 CFR 780.505 - Definition of “shade-grown tobacco.”

Code of Federal Regulations, 2010 CFR

2010-07-01

... Employment or Agricultural Employees in Processing Shade-Grown Tobacco; Exemption From Minimum Wage and... 29 Labor 3 2010-07-01 2010-07-01 false Definition of âshade-grown tobacco.â 780.505 Section 780.505 Labor Regulations Relating to Labor (Continued) WAGE AND HOUR DIVISION, DEPARTMENT OF LABOR...

Douglas-fir in northern California: effects of shade on germination, survival, and growth

Treesearch

Rudolph O. Strothmann

1972-01-01

Effects of four light intensities on germination, survival, and early growth of Douglas-fir (Pseudotsuga menziesii) were studied on south-facing cutblocks in northwestern California. Tested were four shade intensities: 0, 25, 50, and 75 percent. On seeded spots, 50 percent shade resulted in greatest germination and survival, being significantly...

Involvement of abscisic acid in correlative control of flower abscission in soybean

DOE Office of Scientific and Technical Information (OSTI.GOV)

Yarrow, G.L.

1985-01-01

Studies were carried out in three parts: (1) analysis of endogenous abscisic acid (ABA) in abscising and non-abscising flowers, (2) partitioning of radio-labelled ABA and photoassimilates within the soybean raceme, and (3) shading experiments, wherein endogenous levels, metabolism and partitioning of ABA were determined. Endogenous concentrations of ABA failed to show any consistent relationship to abscission of soybean flowers. Partitioning of radiolabelled ABA and photoassimilates displayed consistently higher sink strengths (% DPM) for both /sup 3/H-ABA and /sup 14/C-photoassimilates for non-abscising flowers than for abscising flowers within control racemes. Shading flowers with aluminum foil, 48 hrs prior to sampling, resultedmore » in lowered endogenous ABA concentrations at 12, 17 and 22 days after anthesis (DAA), but not at 0 or 4 DAA. No differences were found in the catabolism of /sup 3/H-ABA between shaded (abscising) and non-shaded (non-abscising) flowers. Reduced partitioning of ABA and photoassimilates to shaded flowers resulted when shades were applied at 0, 4, 12, and 17 DAA, but not at 22 DAA.« less

Partial shading of Cabernet Sauvignon and Shiraz vines altered wine color and mouthfeel attributes, but increased exposure had little impact.

PubMed

Joscelyne, Venetia L; Downey, Mark O; Mazza, Marica; Bastian, Susan E P

2007-12-26

Few studies have investigated the impact of vine shading on the sensory attributes of the resultant wine. This study examines the effects of canopy exposure levels on phenolic composition plus aroma, flavor, and mouthfeel aspects in wine. Wines were made from Cabernet Sauvignon and Shiraz grapes (Vitis vinifera L.) subjected to different levels of canopy exposure in a commercial vineyard in the Sunraysia region, Victoria, Australia. Canopy exposure treatments included control (standard vineyard practice), exposed (achieved with a foliage wire 600 mm above the top cordon), highly exposed (using a foliage wire with leaf plucking in the fruit zone), and shaded treatment (using 70% shade-cloth). Spectral and descriptive analyses showed that levels of anthocyanins, other phenolics, and perceived astringency were lower in wines made from shaded fruit; however, the reverse was generally not observed in wines of exposed and highly exposed fruit. Descriptive analysis also showed wines from the shaded fruit were different from other treatments for a number of flavor and aroma characters. These findings have implications for vineyard management practices.

Woody plant diversity and structure of shade-grown-coffee plantations in northern Chiapas, Mexico.

PubMed

Soto-Pinto, L; Romero-Alvarado, Y; Caballero-Nieto, J; Segura Warnholtz, G

2001-01-01

Shade-grown coffee is an agricultural system that contains some forest-like characteristics. However, structure and diversity are poorly known in shade coffee systems. In 61 coffee-growers' plots of Chiapas, Mexico, structural variables of shade vegetation and coffee yields were measured, recording species and their use. Coffee stands had five vegetation strata. Seventy seven woody species mostly used as wood were found (mean density 371.4 trees per hectare). Ninety percent were native species (40% of the local flora), the remaining were introduced species, mainly fruit trees/shrubs. Diametric distribution resembles that of a secondary forest. Principal Coordinates Analysis grouped plots in four classes by the presence of Inga, however the majority of plots are diverse. There was no difference in equitability among groups or coffee yields. Coffee yield was 835 g clean coffee per shrub, or ca. 1,668 kg ha-1. There is a significant role of shade-grown coffee as diversity refuge for woody plants and presumably associated fauna as well as an opportunity for shade-coffee growers to participate in the new biodiversity-friendly-coffee market.

Tracking reflector assembly for a skylight

DOE Office of Scientific and Technical Information (OSTI.GOV)

Dominquez, R.L.

1984-02-07

A tracking reflector assembly for a skylight includes a ring-shaped base member rotatably supported above the skylight by a plurality of rollers which engage a channel formed within an annular wall of the ring. A reflector is pivotally coupled to the ring for reflecting light into the skylight to supplement light which strikes the skylight directly. A vertical drive motor operates in response to a pair of photosensors for raising and lowering the reflector to follow changes in the angular elevation of the sun. The ring-shaped base member includes a toothed lower surface engaged by a gear coupled to amore » horizontal drive motor for rotating the ring-shaped base member in response to a third photosensor for following east-to-west movement of the sun. Each of the aforementioned photosensors is normally shaded and actuates the associated drive motor only when being struck by direct sunlight. A vertical limit switch limits the amount by which the reflector may be pivotally raised to avoid reflecting midday summer sunlight into the skylight. Another switch is responsive to closure of the reflector over the base member for preventing the vertical drive motor from attempting to further pivot the reflector downwardly. A fourth photosensor senses darkness resulting from sunset or heavy overcast conditions for pivoting the reflector downwardly and returning the base member and reflector to an easterly direction. A limit switch senses the return of the base member to the full east position for terminating actuation of the horizontal drive motor. A user operated switch selectively enables the tracking reflector assembly to operate automatically or causes the reflector to be lowered and the base member to be returned to the full east position until the user again enables the automatic control circuitry.« less

Dissecting molecular and physiological response mechanisms to high solar radiation in cyanic and acyanic leaves: a case study on red and green basil.

PubMed

Tattini, Massimiliano; Sebastiani, Federico; Brunetti, Cecilia; Fini, Alessio; Torre, Sara; Gori, Antonella; Centritto, Mauro; Ferrini, Francesco; Landi, Marco; Guidi, Lucia

2017-04-01

Photosynthetic performance and the expression of genes involved in light signaling and the biosynthesis of isoprenoids and phenylpropanoids were analysed in green ('Tigullio', TIG) and red ('Red Rubin', RR) basil. The aim was to detect the physiological and molecular response mechanisms to high sunlight. The attenuation of blue-green light by epidermal anthocyanins was shown to evoke shade-avoidance responses with consequential effects on leaf morpho-anatomical traits and gas exchange performance. Red basil had a lower mesophyll conductance, partially compensated by the less effective control of stomatal movements, in comparison with TIG. Photosynthesis decreased more in TIG than in RR in high sunlight, because of larger stomatal limitations and the transient impairment of PSII photochemistry. The methylerythritol 4-phosphate pathway promoted above all the synthesis and de-epoxidation of violaxanthin-cycle pigments in TIG and of neoxanthin and lutein in RR. This enabled the green leaves to process the excess radiant energy effectively, and the red leaves to optimize light harvesting and photoprotection. The greater stomatal closure observed in TIG than in RR was due to enhanced abscisic acid (ABA) glucose ester deglucosylation and reduced ABA oxidation, rather than to superior de novo ABA synthesis. This study shows a strong competition between anthocyanin and flavonol biosynthesis, which occurs at the level of genes regulating the oxidation of the C2-C3 bond in the dihydro-flavonoid skeleton. © The Author 2017. Published by Oxford University Press on behalf of the Society for Experimental Biology. All rights reserved. For permissions, please email: journals.permissions@oup.com.

EFFECTS OF LIGHT REDUCTION ON GROWTH OF THE SUBMERGED MACROPHYTE VALLISNERIA AMERICANA AND THE COMMUNITY OF ROOT-ASSOCIATED HETEROTROPHIC BACTERIA

EPA Science Inventory

A shading experiment was conducted over a growing season to measure the effects of light reduction on Vallisneria americana in Perdido Bay on the Florida-Alabama border, and to determine the response of heterotrophic bacteria in the rhizosphere. Plants subjected to 92% light redu...

75 FR 57463 - Notice of Proposed Administrative Settlement Pursuant to the Comprehensive Environmental Response...

Federal Register 2010, 2011, 2012, 2013, 2014

2010-09-21

... public inspection at 1445 Ross Avenue, Dallas, Texas 75202-2733. DATES: Comments must be submitted on or... relating to the settlement are available for public inspection at 1445 Ross Avenue, Dallas, Texas 75202-2733. A copy of the proposed settlement may be obtained from Kevin Shade, 1445 Ross Avenue, Dallas...

Initial response of pondberry released from heavy shade

Treesearch

Brian Roy Lockhart; Emile S. Gardiner; Theodor D. Leininger

2015-01-01

Pondberry [Lindera melissifolia (Walt.) Blume] is a federally endangered woody shrub in the Lauraceae family. The U.S. Fish and Wildlife Service developed a recovery plan in 1993 that emphasized the need to study pondberry biology and ecology and to develop management practices that would promote recovery and conservation of this species. We initiated a large-scale...

Self-shading associated with a skylight-blocked approach system for the measurement of water-leaving radiance and its correction.

PubMed

Shang, Zhehai; Lee, Zhongping; Dong, Qiang; Wei, Jianwei

2017-09-01

Self-shading associated with a skylight-blocked approach (SBA) system for the measurement of water-leaving radiance (L w ) and its correction [Appl. Opt.52, 1693 (2013)APOPAI0003-693510.1364/AO.52.001693] is characterized by Monte Carlo simulations, and it is found that this error is in a range of ∼1%-20% under most water properties and solar positions. A model for estimating this shading error is further developed, and eventually a scheme to correct this error based on the shaded measurements is proposed and evaluated. It is found that the shade-corrected value in the visible domain is within 3% of the true value, which thus indicates that we can obtain not only high precision but also high accuracy L w in the field with the SBA scheme.

Multiple Solutions for Reconfiguration to Address Partial Shading Losses in Solar Photovoltaic Arrays

NASA Astrophysics Data System (ADS)

Sharma, Nikesh; Pareek, Smita; Chaturvedi, Nitin; Dahiya, Ratna

2018-03-01

Solar photovoltaic (SPV) systems are steadily rising and considered as the best alternatives to meet the rising demand of energy. In developing countries like India, SPV’s contribution being a clean energy is the most favourable. However, experiences have shown that produced power of these systems is usually affected due to day, night, seasonal variations, insolation, partial shading conditions etc. Among these parameters, partial shading causes a huge reduction in output power of PV systems. This results in lack of confidence for this technology among users. Thus, it is important and a major challenge in PV systems to minimize the effect of partial shading on their energy production. The work in this paper aims to propose solutions for reconfiguration of solar photovoltaic arrays in order to reduce partial shading losses and thus to enhance power generation.

Partial Shade Stress Test for Thin-Film Photovoltaic Modules: Preprint

DOE Office of Scientific and Technical Information (OSTI.GOV)

Silverman, Timothy J.; Deceglie, Michael G.; Deline, Chris

2015-09-02

Partial shade of monolithic thin-film PV modules can cause reverse-bias conditions leading to permanent damage. In this work, we propose a partial shade stress test for thin-film PV modules that quantifies permanent performance loss. We designed the test with the aid of a computer model that predicts the local voltage, current and temperature stress that result from partial shade. The model predicts the module-scale interactions among the illumination pattern, the electrical properties of the photovoltaic material and the thermal properties of the module package. The test reproduces shading and loading conditions that may occur in the field. It accounts formore » reversible light-induced performance changes and for additional stress that may be introduced by light-enhanced reverse breakdown. We present simulated and experimental results from the application of the proposed test.« less

Developmental and reproductive performance of a specialist herbivore depend on seasonality of, and light conditions experienced by, the host plant

PubMed Central

Zachariades, Costas; Heshula, Lelethu U.; Hill, Martin P.

2018-01-01

Host plant phenology (as influenced by seasonality) and light-mediated changes in the phenotypic and phytochemical properties of leaves have been hypothesised to equivocally influence insect herbivore performance. Here, we examined the effects of seasonality, through host plant phenology (late growth-season = autumn vs flowering-season = winter) and light environment (shade vs full-sun habitat) on the leaf characteristics of the invasive alien plant, Chromolaena odorata. In addition, the performance of a specialist folivore, Pareuchaetes insulata, feeding on leaves obtained from both shaded and full-sun habitats during autumn and winter, was evaluated over two generations. Foliar nitrogen and magnesium contents were generally higher in shaded plants with much higher levels during winter. Leaf water content was higher in shaded and in autumn plants. Total non-structural carbohydrate (TNC) and phosphorus contents did not differ as a function of season, but were higher in shaded foliage compared to full-sun leaves. Leaf toughness was noticeably higher on plants growing in full-sun during winter. With the exception of shaded leaves in autumn that supported the best performance [fastest development, heaviest pupal mass, and highest growth rate and Host Suitability Index (HSI) score], full-sun foliage in autumn surprisingly also supported an improved performance of the moth compared to shaded or full-sun leaves in winter. Our findings suggest that shaded and autumn foliage are nutritionally more suitable for the growth and reproduction of P. insulata. However, the heavier pupal mass, increased number of eggs and higher HSI score in individuals that fed on full-sun foliage in autumn compared to their counterparts that fed on shaded or full-sun foliage in winter suggest that full-sun foliage during autumn is also a suitable food source for larvae of the moth. In sum, our study demonstrates that seasonal and light-modulated changes in leaf characteristics can affect insect folivore performance in ways that are not linear. PMID:29304104

The effect of various disinfectants on dental shade guides.

PubMed

Huang, Peterson Y; Masri, Radi; Romberg, Elaine; Driscoll, Carl F

2014-09-01

Dental shade guides are used to evaluate tooth color before prosthodontic procedures and are subjected to disinfection after use. The effect of disinfection on shade guides has not been thoroughly investigated. The purpose of this study was to evaluate the effect of disinfectants on the color of shade tabs. Changes in the color (ΔE) of VITA Classical Shade Guide tabs were measured with a VITA Easyshade spectrophotometer in the CIELAB system and calculated after being subjected to Cavicide, Asepticare TB, Sporicidin, and distilled water (control) over a simulated period of 2 years. Statistical analysis was accomplished by a 2-way analysis of variance followed by the Tukey honestly significant difference (HSD) test (α=.05). A significant difference was noted in the degree of shade tab color change, depending on the type of disinfectant used (F=153.2, P<.001). No significant difference was noted in the amount of shade tab color change that occurred after disinfection among the different shade tabs used (F=0.611, P=.865), nor was a significant interaction noted between the type of disinfectant and the different shade tabs used (F=0.7, P=.919). Asepticare TB showed the least significant amount of change (ΔE=0.401), and Sporicidin (ΔE=0.889) and the control (ΔE=0.969) showed significantly more color change than Asepticare TB but less than Cavicide (ΔE=1.198). The average total CIELAB color difference for 50% human perceptibility is approximately 1 unit (under standardized laboratory conditions). In the oral cavity, however, an average change of 3.7 ΔE units could still allow teeth to be perceived as having the same color. Therefore, although the results are statistically significant, they may not be clinically important. Copyright © 2014 Editorial Council for the Journal of Prosthetic Dentistry. Published by Elsevier Inc. All rights reserved.

Seeds Use Temperature Cues to Ensure Germination under Nurse-plant Shade in Xeric Kalahari Savannah

PubMed Central

Kos, Martijn; Poschlod, Peter

2007-01-01

Background and Aims In arid environments many plant species are found associated with the canopies of woody perennials. Favourable conditions for establishment under canopies are likely to be associated with shade, but under canopies shade is distributed patchily and differs in quality. Diurnal temperature fluctuations and maximum temperatures could be reliable indicators of safe sites. Here, an examination is made as to whether canopy-associated species use temperature cues to germinate in shade patches, rather than matrix areas between trees. Methods The study was carried out in arid southern Kalahari savannah (Republic of South Africa). Perennial and annual species associated with Acacia erioloba trees and matrix species were germinated at temperature regimes resembling shaded and unshaded conditions. Soil temperature was measured in the field. Key Results Germination of all fleshy-fruited perennial acacia-associated species and two annual acacia-associated species was inhibited by the temperature regime resembling unshaded conditions compared with at least one of the regimes resembling shaded conditions. Inhibition in perennials decreased with seed mass, probably reflecting that smaller seedlings are more vulnerable to drought. Germination of matrix species was not inhibited by the unshaded temperature regime and in several cases it increased germination compared with shaded temperature regimes or constant temperature. Using phylogenetically independent contrasts a significant positive relationship was found between canopy association and the germination at shade temperatures relative to unshaded temperatures. Conclusions The data support the hypothesis that canopy species have developed mechanisms to prevent germination in open sun conditions. The results and data from the literature show that inhibition of germination at temperature regimes characteristic of open sun conditions can be found in fleshy-fruited species of widely divergent taxonomic groups. It is predicted that germination mechanisms to detect canopy shade based on temperature cues are widespread in species depending on nurse plants, especially bird-dispersed species. PMID:17259226

Developmental and reproductive performance of a specialist herbivore depend on seasonality of, and light conditions experienced by, the host plant.

PubMed

Uyi, Osariyekemwen O; Zachariades, Costas; Heshula, Lelethu U; Hill, Martin P

2018-01-01

Host plant phenology (as influenced by seasonality) and light-mediated changes in the phenotypic and phytochemical properties of leaves have been hypothesised to equivocally influence insect herbivore performance. Here, we examined the effects of seasonality, through host plant phenology (late growth-season = autumn vs flowering-season = winter) and light environment (shade vs full-sun habitat) on the leaf characteristics of the invasive alien plant, Chromolaena odorata. In addition, the performance of a specialist folivore, Pareuchaetes insulata, feeding on leaves obtained from both shaded and full-sun habitats during autumn and winter, was evaluated over two generations. Foliar nitrogen and magnesium contents were generally higher in shaded plants with much higher levels during winter. Leaf water content was higher in shaded and in autumn plants. Total non-structural carbohydrate (TNC) and phosphorus contents did not differ as a function of season, but were higher in shaded foliage compared to full-sun leaves. Leaf toughness was noticeably higher on plants growing in full-sun during winter. With the exception of shaded leaves in autumn that supported the best performance [fastest development, heaviest pupal mass, and highest growth rate and Host Suitability Index (HSI) score], full-sun foliage in autumn surprisingly also supported an improved performance of the moth compared to shaded or full-sun leaves in winter. Our findings suggest that shaded and autumn foliage are nutritionally more suitable for the growth and reproduction of P. insulata. However, the heavier pupal mass, increased number of eggs and higher HSI score in individuals that fed on full-sun foliage in autumn compared to their counterparts that fed on shaded or full-sun foliage in winter suggest that full-sun foliage during autumn is also a suitable food source for larvae of the moth. In sum, our study demonstrates that seasonal and light-modulated changes in leaf characteristics can affect insect folivore performance in ways that are not linear.

European Code against Cancer 4th Edition: Ultraviolet radiation and cancer.

PubMed

Greinert, Rüdiger; de Vries, Esther; Erdmann, Friederike; Espina, Carolina; Auvinen, Anssi; Kesminiene, Ausrele; Schüz, Joachim

2015-12-01

Ultraviolet radiation (UVR) is part of the electromagnetic spectrum emitted naturally from the sun or from artificial sources such as tanning devices. Acute skin reactions induced by UVR exposure are erythema (skin reddening), or sunburn, and the acquisition of a suntan triggered by UVR-induced DNA damage. UVR exposure is the main cause of skin cancer, including cutaneous malignant melanoma, basal-cell carcinoma, and squamous-cell carcinoma. Skin cancer is the most common cancer in fair-skinned populations, and its incidence has increased steeply over recent decades. According to estimates for 2012, about 100,000 new cases of cutaneous melanoma and about 22,000 deaths from it occurred in Europe. The main mechanisms by which UVR causes cancer are well understood. Exposure during childhood appears to be particularly harmful. Exposure to UVR is a risk factor modifiable by individuals' behaviour. Excessive exposure from natural sources can be avoided by seeking shade when the sun is strongest, by wearing appropriate clothing, and by appropriately applying sunscreens if direct sunlight is unavoidable. Exposure from artificial sources can be completely avoided by not using sunbeds. Beneficial effects of sun or UVR exposure, such as for vitamin D production, can be fully achieved while still avoiding too much sun exposure and the use of sunbeds. Taking all the scientific evidence together, the recommendation of the 4th edition of the European Code Against Cancer for ultraviolet radiation is: "Avoid too much sun, especially for children. Use sun protection. Do not use sunbeds." Copyright © 2015 International Agency for Research on Cancer. Published by Elsevier Ltd. All rights reserved.

3D Shape Perception in Posterior Cortical Atrophy: A Visual Neuroscience Perspective.

PubMed

Gillebert, Céline R; Schaeverbeke, Jolien; Bastin, Christine; Neyens, Veerle; Bruffaerts, Rose; De Weer, An-Sofie; Seghers, Alexandra; Sunaert, Stefan; Van Laere, Koen; Versijpt, Jan; Vandenbulcke, Mathieu; Salmon, Eric; Todd, James T; Orban, Guy A; Vandenberghe, Rik

2015-09-16

Posterior cortical atrophy (PCA) is a rare focal neurodegenerative syndrome characterized by progressive visuoperceptual and visuospatial deficits, most often due to atypical Alzheimer's disease (AD). We applied insights from basic visual neuroscience to analyze 3D shape perception in humans affected by PCA. Thirteen PCA patients and 30 matched healthy controls participated, together with two patient control groups with diffuse Lewy body dementia (DLBD) and an amnestic-dominant phenotype of AD, respectively. The hierarchical study design consisted of 3D shape processing for 4 cues (shading, motion, texture, and binocular disparity) with corresponding 2D and elementary feature extraction control conditions. PCA and DLBD exhibited severe 3D shape-processing deficits and AD to a lesser degree. In PCA, deficient 3D shape-from-shading was associated with volume loss in the right posterior inferior temporal cortex. This region coincided with a region of functional activation during 3D shape-from-shading in healthy controls. In PCA patients who performed the same fMRI paradigm, response amplitude during 3D shape-from-shading was reduced in this region. Gray matter volume in this region also correlated with 3D shape-from-shading in AD. 3D shape-from-disparity in PCA was associated with volume loss slightly more anteriorly in posterior inferior temporal cortex as well as in ventral premotor cortex. The findings in right posterior inferior temporal cortex and right premotor cortex are consistent with neurophysiologically based models of the functional anatomy of 3D shape processing. However, in DLBD, 3D shape deficits rely on mechanisms distinct from inferior temporal structural integrity. Posterior cortical atrophy (PCA) is a neurodegenerative syndrome characterized by progressive visuoperceptual dysfunction and most often an atypical presentation of Alzheimer's disease (AD) affecting the ventral and dorsal visual streams rather than the medial temporal system. We applied insights from fundamental visual neuroscience to analyze 3D shape perception in PCA. 3D shape-processing deficits were affected beyond what could be accounted for by lower-order processing deficits. For shading and disparity, this was related to volume loss in regions previously implicated in 3D shape processing in the intact human and nonhuman primate brain. Typical amnestic-dominant AD patients also exhibited 3D shape deficits. Advanced visual neuroscience provides insight into the pathogenesis of PCA that also bears relevance for vision in typical AD. Copyright © 2015 Gillebert, Schaeverbeke et al.

The effect of different shades of specific luting agents and IPS empress ceramic thickness on overall color.

PubMed

Terzioğlu, Hakan; Yilmaz, Burak; Yurdukoru, Bengul

2009-10-01

The color stability of both porcelain and luting materials is very important for the esthetics of laminate veneers and all-ceramic crowns. The purpose of this study was to determine the effect of different shades of resin-based luting cement and the thickness of IPS Empress ceramics on the final color of the restorations. Resin-based dual-polymerized composite cement in two different shades (RelyX ARC) and ceramic disks of different thicknesses were selected for the study. Forty specimens (ten each of four different thicknesses: 0.5 mm, 1 mm, 2 mm, and 3 mm) were used for the evaluation. Initial specimen color parameters were determined in a Commission Internationale de l'Eclairage L*a*b* color order system with a colorimeter. Two different shades of the cement were prepared as polymerized layers and applied to one face of the specimens in order. Color changes were calculated between baseline color measurements and measurements after cementation. Color difference data were analyzed statistically. All specimens showed a significant color shift (DE > 3.7) after cementation regardless of the cement shade. However, the differences in the cement shade did not significantly affect the final color of the ceramic specimens for any thickness, and color shifts were not perceivable between the different shades of cement. (Int J Periodontics Restorative Dent 2009;29:499-505.).

Hydraulics and life history of tropical dry forest tree species: coordination of species' drought and shade tolerance.

PubMed

Markesteijn, Lars; Poorter, Lourens; Bongers, Frans; Paz, Horacio; Sack, Lawren

2011-07-01

Plant hydraulic architecture has been studied extensively, yet we know little about how hydraulic properties relate to species' life history strategies, such as drought and shade tolerance. The prevailing theories seem contradictory. We measured the sapwood (K(s) ) and leaf (K(l) ) hydraulic conductivities of 40 coexisting tree species in a Bolivian dry forest, and examined associations with functional stem and leaf traits and indices of species' drought (dry-season leaf water potential) and shade (juvenile crown exposure) tolerance. Hydraulic properties varied across species and between life-history groups (pioneers vs shade-tolerant, and deciduous vs evergreen species). In addition to the expected negative correlation of K(l) with drought tolerance, we found a strong, negative correlation between K(l) and species' shade tolerance. Across species, K(s) and K(l) were negatively correlated with wood density and positively with maximum vessel length. Consequently, drought and shade tolerance scaled similarly with hydraulic properties, wood density and leaf dry matter content. We found that deciduous species also had traits conferring efficient water transport relative to evergreen species. Hydraulic properties varied across species, corresponding to the classical trade-off between hydraulic efficiency and safety, which for these dry forest trees resulted in coordinated drought and shade tolerance across species rather than the frequently hypothesized trade-off. © 2011 The Authors. New Phytologist © 2011 New Phytologist Trust.

Realtime Hourly Data Table

Science.gov Websites

Tables View the latest hourly text summary CLICK ON UNDERLINED HOUR / SHADED BOX FOR THE LATEST CYCLE 00z Dump Tables View the latest rap text summary CLICK ON UNDERLINED HOUR / SHADED BOX FOR THE LATEST CYCLE Data Dump Tables View the latest model data text summary NAM GFS GDS CLICK ON UNDERLINED HOUR / SHADED

Shade Tolerance of Festuca paradoxa Desv., a Cool-Season Grass Native to North America

Treesearch

Nadia Navarrete-Tindall; Larry Mechlin; J. W. Van Sambeek

2003-01-01

Paradox grass (Festuca paradoxa Desv.) is a native cool-season grass found in prairies and forest openings. Paradox grass has not been included in tree plantings. To determine paradox grass adaptation to shaded environmmts, we established a pot experiment in the shade laboratory at the University of Missouri Horticulture and Agroforestry Research...

Evaluating relationships among tree growth rate, shade tolerance, and browse tolerance following disturbance in an eastern deciduous forest

Treesearch

Lisa M. Krueger; Chris J. Peterson; Alejandro Royo; Walter P. Carson

2009-01-01

Interspecific differences in shade tolerance among woody species are considered a primary driving force underlying forest succession. However, variation in shade tolerance may be only one of many interspecific differences that cause species turnover. For example, tree species may differ in their sensitivity to herbivory. Nonetheless,...

Shades of Pink: Preschoolers Make Meaning in a Reggio-Inspired Classroom

ERIC Educational Resources Information Center

Kim, Bo Sun

2012-01-01

Shades of Pink study describes how six preschoolers and their teacher engaged in a collaborative learning project through which they learned about the shades of a color--in this case, pink. As the children learned through experimenting and discussing their theories, they represented ideas using art as a tool for discovery and learning. The study…

Chlorophyll Fluorescence Is a Better Proxy for Sunlit Leaf Than Total Canopy Photosynthesis

NASA Astrophysics Data System (ADS)

Chen, J. M.; Wang, Z.; Zhang, F.; Mo, G.

2015-12-01

Chlorophyll fluorescence (CF) results from non-photochemical quenching during plant photosynthesis under excessive radiation. We explore the relationship between gross primary productivity (GPP) and CF using a process ecosystem model, which separates a vegetation canopy into sunlit and shaded leaf groups and simulates the total canopy GPP as the sum of sunlit and shaded leaf GPP. Using GOME-2 and GOSAT data acquired in 2010 over the global land surface, we found that measured CF signals gridded in 1 degree resolution are well correlated with simulated total GPP and its sunlit and shaded components, but the correlation coefficients (R) are largest for the sunlit GPP and smallest for shaded GPP. The seasonal R2 values vary from 0.57 to 0.74, 0.58 to 0.71, and 0.48 to 0.56 for sunlit, total and shaded GPP, respectively. The significance levels for these correlations are all greater than p<0.01. Averaged over the globe, the total simulated shaded GPP is 39% of the total GPP. Theoretically, CF from vegetation comes mostly from sunlit leaves. The significant correlation between measured canopy-level CF and the shaded GPP is likely due to the correlation between shaded and sunlit GPP as both increase with leaf area index. Our simulation confirms the validity of using canopy-level CF measurements to assess the total GPP as the first approximation, although these measurements are a consistently better indicator of sunlit GPP than total GPP. In previous studies, the R2 values for the correlation between CF and total GPP were found to range from 0.76 to 0.88, 0.56 to 0.78, and 0.57 to 0.77 for MPI-BGC, MODIS and CASA model results, respectively. These values are similar or larger than those for sunlit GPP simulated in our study, but are considerably larger than those for total GPP in our study because the correlation for total GPP is contaminated by the inclusion of shaded GPP. All these three models use canopy total light use efficiency without considering the differences between sunlit and shaded leaves, and therefore they mostly capture spatio-temporal variations in sunlit GPP. We therefore argue that solar-induced CF measured from vegetation is a better proxy of sunlit GPP than the total GPP, and the use of CF data for assessing the terrestrial carbon cycle can be improved when sunlit and shaded GPP are modelled separately.

Loudspeaker line array educational demonstration.

PubMed

Anderson, Brian E; Moser, Brad; Gee, Kent L

2012-03-01

This paper presents a physical demonstration of an audio-range line array used to teach interference of multiple sources in a classroom or laboratory exercise setting. Software has been developed that permits real-time control and steering of the array. The graphical interface permits a user to vary the frequency, the angular response by phase shading, and reduce sidelobes through amplitude shading. An inexpensive, eight-element loudspeaker array has been constructed to test the control program. Directivity measurements of this array in an anechoic chamber and in a large classroom are presented. These measurements have good agreement with theoretical directivity predictions, thereby allowing its use as a quantitative learning tool for advanced students as well as a qualitative demonstration of arrays in other settings. Portions of this paper are directed toward educators who may wish to implement a similar demonstration for their advanced undergraduate or graduate level course in acoustics. © 2012 Acoustical Society of America

Photosynthesis in chlorolichens: the influence of the habitat light regime.

PubMed

Piccotto, Massimo; Tretiach, Mauro

2010-11-01

The hypothesis that CO(2) gas exchange and chlorophyll a fluorescence (ChlaF) of lichens vary according to the light regimes of their original habitat, as observed in vascular plants, was tested by analysing the photosynthetic performance of 12 populations of seven dorsoventral, foliose lichens collected from open, south-exposed rocks to densely shaded forests. Light response curves were induced at optimum thallus water content and ChlaF emission curves at the species-specific photon flux at which the quantum yield of CO(2) assimilation is the highest and is saturating the photosynthetic process. Photosynthetic pigments were quantified in crude extracts. The results confirm that the maximum rate of gross photosynthesis is correlated with the chlorophyll content of lichens, which is influenced by light as well as by nitrogen availability. Like leaves, shade tolerant lichens emit more ChlaF than sun-loving ones, whereas the photosynthetic quantum conversion is higher in the latter.

The linear shade guide design of Vita 3D-master performs as well as the original design of the Vita 3D-master.

PubMed

Corcodel, N; Rammelsberg, P; Jakstat, H; Moldovan, O; Schwarz, S; Hassel, A J

2010-11-01

Visual tooth colour assessment by use of the Vita 3D-Master(®) (3D; Vita Zahnfabrik, Bad Säckingen, Germany) is well documented. To improve handling, a new linear arrangement of the shade tabs has been introduced (LG; Linearguide 3D-Master(®) ). The purpose of this study was to investigate whether the linear design has an effect on shade matching. Fifty-six students underwent identical, theoretical and practical training, by use of an Internet learning module [Toothguide Training Software(®) (TT)] and a standardised training programme [Toothguide Training Box(®) (TTB)]. Each student then matched 30 randomly chosen shade tabs presented in an intra-oral setting by a standardised device [Toothguide Check Box(®) (TCB)]; 15 matches were made using the 3D and 15 using the LG shade guide system, under a daylight lamp (840 matches for each guide). It was recorded to what extent the presented and selected shade tabs, or the lightness group of the tabs, matched, also the needed time for colour matching. The results showed that 35% of perfect matches were observed for the 3D and 32% for the LG. The lightness group was correct in 59% of cases for 3D and 56% for LG. Mean time needed for matching of tabs and lightness group was no different between groups (no significant difference for any assessment). Within the limitations of the study design, the colour assessment with regard to performance and time needed in shade matching was not different with the LG or the 3D. Therefore, the user should choose which shade tab arrangement is more applicable. © 2010 Blackwell Publishing Ltd.

Influence of Color Education and Training on Shade Matching Skills.

PubMed

Ristic, Ivan; Stankovic, Sasa; Paravina, Rade D

2016-09-01

To evaluate the influence of education and training on quality of tooth color matching. Dental students (N = 174), matched the color of eight shade tabs in a viewing booth, using VITA Linearguide 3D-Master shade guide. The experimental group had color education and training between the before and after session. The control group did not have any additional information in between two sessions. Color differences between the task tabs and selected tabs were calculated using CIE formulas. The score for the best match (smallest color difference) was 10 points, the 2nd best match 9 points, down to 1 point for the 10th best match. Means and standard deviations were calculated. Differences were analyzed using the Student t-test. Shade matching scores in the experimental group were significantly better after education and training (p < 0.001), with a mean score before and after shade matching sessions of 7.06 (1.19) and 8.43 (0.92), respectively. The percentage of students in the experimental group that selected one of three best matches increased 24.3%. The control group exhibited no significant improvement in the after session. Within the limitations of the study, education and training improved students' shade matching skills. While the vast majority of dental restorations and practically all restorations in the esthetic zone are tooth colored, the profession as a whole is far from perfect when it comes to accurate shade matching. Education and training can improve shade matching ability: enhanced esthetics of dental restorations, increased patient satisfaction, and a reduced number of color corrections are some of the notable benefits and rewards. (J Esthet Restor Dent 00:000-000, 2016) J Esthet Restor Dent 28:287-294, 2016). © 2016 Wiley Periodicals, Inc.

[Comparative analysis of some factors in tooth color matching].

PubMed

Török, Judit; Mauks, Levente; Márton, Sándor; Hegedűs, Csaba

2014-09-01

Recently (nowadays) to achieve a natural looking restoration is an ever increasing demand from the patients and also from the doctor side. To select the right color of the restoration matching the remaining natural teeth is always a challenging task. A clinical study was performed at the University of Debrecen Faculty of Dentristry with the help of dental students using two different shade guides. The study tested the influence of gender and knowledge of color science on shade matching. 78 students were asked to find the right matching color of the same upper canine to two different shade guides (Vitapan Classic and Vita 3D-Master) under standard condition. After informing the student about the basic principles of color the matching procedure was repeated. Results were analyzed statistically. In our study we found that gender does not influence the color choice. Matching accuracy is not increased by better knowledge of colors. We can conclude that significantly less students matched the proper color with Vitapan Classic shade guide after information of the property of colors without training the shade selection. Within the limitation of the study design it was concluded that not more students selected the proper color even after giving them information about colors, instructions about shade selections. For the same one canine several color were selected by the participants (6 types with Vitapan Classic and 19 types with Vita 3D-Master) which conformed that visual determination is not a reliably consistent way of the tooth shade selection. The Vita Company 1990s developed 3D-Master shade guide is not widely used, although we found the repeatability is more than 70%.

A continent-wide analysis of the shade requirements of red and western grey kangaroos

PubMed Central

Roberts, J. A.; Coulson, G.; Munn, A. J.; Kearney, M. R.

2016-01-01

ABSTRACT Foraging time may be constrained by a suite of phenomena including weather, which can restrict a species' activity and energy intake. This is recognized as pivotal for many species whose distributions are known to correlate with climate, including kangaroos, although such impacts are rarely quantified. We explore how differences in shade seeking, a thermoregulatory behavior, of 2 closely-related kangaroo species, Macropus rufus (red kangaroos) and M. fuliginosus (western grey kangaroos), might reflect differences in their distributions across Australia. We observed foraging and shade-seeking behavior in the field and, together with local weather observations, calculated threshold radiant temperatures (based on solar and infrared radiant heat loads) over which the kangaroos retreated to shade. We apply these calculated tolerance thresholds to hourly microclimatic estimates derived from daily-gridded weather data to predict activity constraints across the Australian continent over a 10-year period. M. fuliginosus spent more time than M. rufus in the shade (7.6 ± 0.7 h versus 6.4 ± 0.9 h) and more time foraging (11.8 ± 0.5 h vs. 10.0 ± 0.6 h), although total time resting was equivalent (∼8.2 h). M. rufus tolerated 19°C higher radiant temperatures than M. fuliginosus (89°C versus 70°C radiant temperature). Across Australia, we predicted M. fuliginosus to be more restricted to shade than M. rufus, with higher absolute shade requirements farther north. These results corroborate previous findings that M. rufus is more adept at dealing with heat than M. fuliginosus and indicate that M. rufus is less dependent on shade on a continental scale. PMID:27857963

[The fabrication and chroma study of a kind of custom shade guide with metal substrate].

PubMed

Chen, Ling; Liu, Xian; Xu, Li-qiang; Chao, Yong-lie; Liu, Wen-fang; Chen, Xiao-fang

2007-06-01

This study was to explore the chroma of a custom shade guide with metal substrate. A custom shade guide sample was fabricated using the central maxillary incisor as a model,which was corresponding to the colour of VITAPAN shade guide.The colour of each shade tab was measured with PR-650 spectrophotometer when every porcelain was added. From the opaque to glaze, the lightness was decreased and the chroma was increased distinctively; When the body porcelain were baked over their corresponding opaque, the lightness was decreased and the chroma was increased; When the enamel porcelain and the translucent porcelain were added on, the lightness and chroma were decreased slightly; the glaze could increase the lightness. The "add on" porcelain can affect the final color of the restoration, and the effects of each porcelain layer were different from each other.

Differences between the human eye and the spectrophotometer in the shade matching of tooth colour.

PubMed

Gómez-Polo, Cristina; Gómez-Polo, Miguel; Celemin-Viñuela, Alicia; Martínez Vázquez De Parga, Juan Antonio

2014-06-01

The aim of this work was to assess the agreement between instrumental and visual colour matching. Shade selection with the 3DMaster Toothguide (Vita-Zahnfabrik) was performed for 1361 maxillary central incisors and compared with the shade obtained with the EasyShade Compact (Vita-Zahnfabrik) spectrophotometer. We observed a greater correlation between the objective method and the subjective one in the colour dimension of lightness (Kappa 0.6587), followed by hue (Kappa 0.4337) and finally chroma (Kappa 0.3578). The colour dimension in which the greatest agreement is seen between the operator and the spectrophotometer is value or lightness. This study reveals differences between the measurement of colour via spectrophotometry and the visual shade selection method. According to our results, there is better agreement in the value or lightness colour dimension, which is the most important one in the choice of tooth colour. Copyright © 2014 Elsevier Ltd. All rights reserved.

[In vivo model to evaluate the accuracy of complete-tooth spectrophotometer for dental clinics].

PubMed

Liu, Feng; Yang, Jian; Xu, Tong-Kai; Xu, Ming-Ming; Ma, Yu

2011-02-01

To test ΔE between measured value and right value from the Crystaleye complete-tooth spectrophotometer, and to evaluate the accuracy rate of the spectrophotometer. Twenty prosthodontists participated in the study. Each of them used Vita 3D-Master shadeguide to do the shade matching, and used Crystaleye complete-tooth spectrophotometer (before and after the test training) tested the middle of eight fixed tabs from shadeguide in the dark box. The results of shade matching and spectrophotometer were recorded. The accuracy rate of shade matching and the spectrophotometer before and after training were calculated. The average accuracy rate of shade matching was 49%. The average accuracy rate of the spectrophotometer before and after training was 83% and 99%. The accuracy of the spectrophotometer was significant higher than that in shade matching, and training can improve the accuracy rate.

The effects of zilpaterol hydrochloride and shade on blood metabolites of finishing beef steers

USDA-ARS?s Scientific Manuscript database

The effects of feeding zilpaterol hydrochloride (ZH) and shade were evaluated on blood metabolites and lung score in finishing beef steers. Cattle were fed 0 or 8.33 mg/kg ZH for 21 d with a 3- or 4-d withdrawal before harvest and were housed in open or shaded pens. Blood samples and lung scores w...

Competition among surface roots in a selectively-logged, semi-deciduous forest in southeastern Mexico - effects on seedlings of two species of contrasting shade tolerance

Treesearch

Matthew Dickinson; D.F. Wigham

2013-01-01

Experimental manipulations of root competition on naturally established seedlings were conducted across canopy openness and soil depth gradients in a selectively-logged, semideciduous forest on limestone-derived soils in southeastern Mexico. We studied the relatively shade intolerant mahogany (Swietenia macrophylla, Meliaceae) and shade tolerant...

AVIAN RESOURCE USE IN DOMINICAN SHADE COFFEE PLANTATIONS

Treesearch

JOSEPH M. WUNDERLE. JR; STEVEN C. LATTAA

1998-01-01

We quantified foraging behavior of 19 bird species in shade coffee plantations in the Dominican Republic to document and evaluate their use of food resources in the shade overstory relative to the coffee understory. All species were observed foraging in the Znga “era overstory, and 18 of the 19 species had median foraging heights significantly above the median maximum...

Importance of Shading to Visitors Selecting a Campsite at Indian Boundary Campground in Tennessee

Treesearch

George A. James; Harold K. Cordell

1970-01-01

Campers at Indian Boundary Campground in Tenr.essee were interviewed during 1966 and 1967 to determine the amount of shading they preferred. The overstory in this campground was selectively thinned several years earlier. This gave campers a choice of campsites, ranging from those so heavily shaded that little sunlight reached the forest floor to...

Performance assessment of Vita Easy Shade spectrophotometer on colour measurement of aesthetic dental materials.

PubMed

AlGhazali, N; Burnside, G; Smith, R W; Preston, A J; Jarad, F D

2011-12-01

Four different shades were used to produce 20 samples of resin-based composite and 20 samples of porcelain to evaluate the performance ability of an intra oral test spectrophotometer compared to a reference spectrophotometer. The absolute colour coordinates CIELAB values measured with both spectrophotometers were significantly different (p < 0.001). However, a high correlation was found (p < 0.001) despite the low concordance noticed. The colour difference deltaE* values calculated between different shades also were significantly different between both spectrophotometers (p < 0.05). Therefore, the Easy Shade can be used in dental practice and dental research with some limitations.

Habitat use by the endangered Karner blue butterfly in oak woodlands: The influence of canopy cover

USGS Publications Warehouse

Grundel, Ralph; Pavlovic, Noel B.; Sulzman, Christina L.

1998-01-01

The Karner blue butterfly Lycaeides melissa samuelis is an endangered species residing in the Great Lakes and northeastern regions of the United States. Increased canopy cover is a major factor implicated in the decline of the Karner blue at many locales. Therefore, we examined how the butterfly's behavior varied with canopy cover. Adult males at Indiana Dunes National Lakeshore used habitat under canopy openings for nearly 90% of their activities; females used openings and shaded areas more equally. The frequency of oviposition on the sole host plant, wild lupine Lupinus perennis, was highest under 30–60% canopy cover even though lupine was more abundant in more open areas. Larvae fed preferentially on larger lupine plants and on lupines in denser patches. However, lupines were generally larger in the shade. Therefore, shade-related trade-offs existed between lupine abundance and distribution of larval feeding and oviposition. Also, heterogeneity of shading by sub-canopy woody vegetation was greater at oviposition sites than at sites where lupine did not grow. Given the importance of shade heterogeneity, a mixture of canopy openings and shade, on a scale similar to daily adult movement range, should be beneficial for this butterfly.

Hydraulic acclimation to shading in boreal conifers of varying shade tolerance.

PubMed

Schoonmaker, Amanda L; Hacke, Uwe G; Landhäusser, Simon M; Lieffers, Victor J; Tyree, Melvin T

2010-03-01

The purpose of this study was to determine how shading affects the hydraulic and wood-anatomical characteristics of four boreal conifers (Pinus banksiana, Pinus contorta, Picea glauca and Picea mariana) that differ in shade tolerance. Plants were grown in an open field and under a deciduous-dominated overstory for 6 years. Sapwood- and leaf-area specific conductivity, vulnerability curves, and anatomical measurements (light and scanning electron microscopy) were made on leading shoots from six to nine trees of each treatment combination. There was no difference in sapwood-area specific conductivity between open-grown and understory conifers, although two of four species had larger tracheid diameters in the open. Shaded conifers appeared to compensate for small diameter tracheids by changes in pit membrane structure. Scanning electron microscopy revealed that understory conifers had thinner margo strands, greater maximum pore size in the margo, and more torus extensions. All of these trends may contribute to inadequate sealing of the torus. This is supported by the fact that all species showed increased vulnerability to cavitation when grown in the understory. Although evaporative demand in an understory environment is low, a rapid change into fully exposed conditions could be detrimental for shaded conifers.

Metabolomic Analysis Using Ultra-Performance Liquid Chromatography-Quadrupole-Time of Flight Mass Spectrometry (UPLC-Q-TOF MS) Uncovers the Effects of Light Intensity and Temperature under Shading Treatments on the Metabolites in Tea

PubMed Central

Ma, Lifeng; Yi, Xiaoyun; Ruan, Jianyun

2014-01-01

To investigate the effect of light intensity and temperature on the biosynthesis and accumulation of quality-related metabolites, field grown tea plants were shaded by Black Net and Nano-insulating Film (with additional 2–4°C cooling effect) with un-shaded plants as a control. Young shoots were subjected to UPLC-Q-TOF MS followed by multivariate statistical analysis. Most flavonoid metabolites (mainly flavan-3-ols, flavonols and their glycosides) decreased significantly in the shading treatments, while the contents of chlorophyll, β-carotene, neoxanthin and free amino acids, caffeine, benzoic acid derivatives and phenylpropanoids increased. Comparison between two shading treatments indicated that the lower temperature under Nano shading decreased flavonols and their glycosides but increased accumulation of flavan-3-ols and proanthocyanidins. The comparison also showed a greater effect of temperature on galloylation of catechins than light intensity. Taken together, there might be competition for substrates between the up- and down-stream branches of the phenylpropanoid/flavonoid pathway, which was influenced by light intensity and temperature. PMID:25390340

Metabolomic analysis using ultra-performance liquid chromatography-quadrupole-time of flight mass spectrometry (UPLC-Q-TOF MS) uncovers the effects of light intensity and temperature under shading treatments on the metabolites in tea.

PubMed

Zhang, Qunfeng; Shi, Yuanzhi; Ma, Lifeng; Yi, Xiaoyun; Ruan, Jianyun

2014-01-01

To investigate the effect of light intensity and temperature on the biosynthesis and accumulation of quality-related metabolites, field grown tea plants were shaded by Black Net and Nano-insulating Film (with additional 2-4°C cooling effect) with un-shaded plants as a control. Young shoots were subjected to UPLC-Q-TOF MS followed by multivariate statistical analysis. Most flavonoid metabolites (mainly flavan-3-ols, flavonols and their glycosides) decreased significantly in the shading treatments, while the contents of chlorophyll, β-carotene, neoxanthin and free amino acids, caffeine, benzoic acid derivatives and phenylpropanoids increased. Comparison between two shading treatments indicated that the lower temperature under Nano shading decreased flavonols and their glycosides but increased accumulation of flavan-3-ols and proanthocyanidins. The comparison also showed a greater effect of temperature on galloylation of catechins than light intensity. Taken together, there might be competition for substrates between the up- and down-stream branches of the phenylpropanoid/flavonoid pathway, which was influenced by light intensity and temperature.

Climate and species functional traits influence maximum live tree stocking in the Lake States, USA

Treesearch

Mark J. Ducey; Christopher W. Woodall; Andrés Bravo-Oviedo

2017-01-01

Quantifying the density of live trees in forest stands and partitioning it between species or other stand components is critical for predicting forest dynamics and responses to management, as well as understanding the impacts of stand composition and structure on productivity. As plant traits such as shade tolerance have been proven to refine understanding of plant...

Different shades of gray: crafting a regulatory response to private equity buyouts in electricity

DOE Office of Scientific and Technical Information (OSTI.GOV)

Vaheesan, Sandeep

2009-01-15

Private equity has made a dramatic entry into the once-staid world of electric utilities. These investors, with their superior management expertise and greater access to capital, have the potential to improve utility operations and increase investment in badly needed generation and transmission facilities. Their acquisitions do not come without their share of risks, however. (author)

Capturing species-level drought responses in a temperate deciduous forest using ratios of photochemical reflectance indices between sunlit and shaded canopies

Treesearch

Taehee Hwang; Hamed Gholizadeh; Daniel A. Sims; Kimberly A. Novick; Edward R. Brzostek; Richard P. Phillips; Daniel T. Roman; Scott M. Robeson; Abdullah F. Rahman

2017-01-01

To classify trees along a spectrum of isohydric to anisohydric behavior is a promising new framework for identifying tree species' sensitivities to drought stress, directly related to the vulnerability of carbon uptake of terrestrial ecosystems with increased hydroclimate variability. Trees with isohydric strategies regulate stomatal conductance to maintain...

Fortifying the forest: Thinning and burning increase resistance to a bark beetle outbreak and promote forest resilience

Treesearch

Sharon M. Hood; Stephen Baker; Anna Sala

2016-01-01

Fire frequency in low-elevation coniferous forests in western North America has greatly declined since the late 1800s. In many areas, this has increased tree density and the proportion of shade-tolerant species, reduced resource availability, and increased forest susceptibility to forest insect pests and high-severity wildfire. In response, treatments are...

Repeated prescribed fires alter gap-phase regeneration in mixed-oak forests

Treesearch

Todd F. Hutchinson; Robert P. Long; Joanne Rebbeck; Elaine Kennedy Sutherland; Daniel A. Yaussy

2012-01-01

Oak dominance is declining in the central hardwoods region, as canopy oaks are being replaced by shade-tolerant trees that are abundant in the understory of mature stands. Although prescribed fire can reduce understory density, oak seedlings often fail to show increased vigor after fire, as the canopy remains intact. In this study, we examine the response of tree...

Douglas-fir survival and growth in response to spring planting date and depth

Treesearch

R. O. Strothmann

1971-01-01

Douglas-fir seedlings were planted by four methods in late February and late March on hot, south-facing slopes in northwestern California. Besides standard planting, the techniques used included deep planting at two different depths, and shading the lower stem. The differences in survival after 3 growing seasons were not statistically significant, but deep planting had...

Antelope bitterbrush and Scouler's willow response to a shelterwood harvest and prescribed burn in western Montana

Treesearch

Dayna M. Ayers; Donald J. Bedunah; Michael G. Harrington

1999-01-01

In many western Montana ponderosa pine (Pinus ponderosa) stands, fire suppression and past selective logging of large trees have resulted in conditions favoring succession to dense stands of shade-tolerant, but insect- and disease-prone Douglas-fir (Pseudotsuga menziesii). Stand thinning and understory prescribed burning have been proposed as surrogates for pre-Euro-...

Energy savings for heat-island reduction strategies in Chicago and Houston (including updates for Baton Rouge, Sacramento, and Salt Lake City)

DOE Office of Scientific and Technical Information (OSTI.GOV)

Konopacki, S.; Akbari, H.

2002-02-28

In 1997, the U.S. Environmental Protection Agency (EPA) established the ''Heat Island Reduction Initiative'' to quantify the potential benefits of Heat-Island Reduction (HIR) strategies (i.e., shade trees, reflective roofs, reflective pavements and urban vegetation) to reduce cooling-energy use in buildings, lower the ambient air temperature and improve urban air quality in cities, and reduce CO2 emissions from power plants. Under this initiative, the Urban Heat Island Pilot Project (UHIPP) was created with the objective of investigating the potential of HIR strategies in residential and commercial buildings in three initial UHIPP cities: Baton Rouge, LA; Sacramento, CA; and Salt Lake City,more » UT. Later two other cities, Chicago, IL and Houston, TX were added to the UHIPP. In an earlier report we summarized our efforts to calculate the annual energy savings, peak power avoidance, and annual CO2 reduction obtainable from the introduction of HIR strategies in the initial three cities. This report summarizes the results of our study for Chicago and Houston. In this analysis, we focused on three building types that offer the highest potential savings: single-family residence, office and retail store. Each building type was characterized in detail by vintage and system type (i.e., old and new building constructions, and gas and electric heat). We used the prototypical building characteristics developed earlier for each building type and simulated the impact of HIR strategies on building cooling- and heating-energy use and peak power demand using the DOE-2.1E model. Our simulations included the impact of (1) strategically-placed shade trees near buildings [direct effect], (2) use of high-albedo roofing material on the building [direct effect], (3) urban reforestation with high-albedo pavements and building surfaces [indirect effect] and (4) combined strategies 1, 2, and 3 [direct and indirect effects]. We then estimated the total roof area of air-conditioned buildings in each city using readily obtainable data to calculate the metropolitan-wide impact of HIR strategies. The results show that in Chicago, potential annual energy savings of $30M could be realized by ratepayers from the combined direct and indirect effects of HIR strategies. Additionally, peak power avoidance is estimated at 400 MW and the reduction in annual carbon emissions at 58 ktC. In Houston, the potential annual energy savings are estimated at $82M, with an avoidance of 730 MW in peak power and a reduction in annual carbon emissions of 170 ktC.« less

Seagrass canopy photosynthetic response is a function of canopy density and light environment: a model for Amphibolis griffithii.

PubMed

Hedley, John D; McMahon, Kathryn; Fearns, Peter

2014-01-01

A three-dimensional computer model of canopies of the seagrass Amphibolis griffithii was used to investigate the consequences of variations in canopy structure and benthic light environment on leaf-level photosynthetic saturation state. The model was constructed using empirical data of plant morphometrics from a previously conducted shading experiment and validated well to in-situ data on light attenuation in canopies of different densities. Using published values of the leaf-level saturating irradiance for photosynthesis, results show that the interaction of canopy density and canopy-scale photosynthetic response is complex and non-linear, due to the combination of self-shading and the non-linearity of photosynthesis versus irradiance (P-I) curves near saturating irradiance. Therefore studies of light limitation in seagrasses should consider variation in canopy structure and density. Based on empirical work, we propose a number of possible measures for canopy scale photosynthetic response that can be plotted to yield isoclines in the space of canopy density and light environment. These plots can be used to interpret the significance of canopy changes induced as a response to decreases in the benthic light environment: in some cases canopy thinning can lead to an equivalent leaf level light environment, in others physiological changes may also be required but these alone may be inadequate for canopy survival. By providing insight to these processes the methods developed here could be a valuable management tool for seagrass conservation during dredging or other coastal developments.

Seagrass Canopy Photosynthetic Response Is a Function of Canopy Density and Light Environment: A Model for Amphibolis griffithii

PubMed Central

Hedley, John D.; McMahon, Kathryn; Fearns, Peter

2014-01-01

A three-dimensional computer model of canopies of the seagrass Amphibolis griffithii was used to investigate the consequences of variations in canopy structure and benthic light environment on leaf-level photosynthetic saturation state. The model was constructed using empirical data of plant morphometrics from a previously conducted shading experiment and validated well to in-situ data on light attenuation in canopies of different densities. Using published values of the leaf-level saturating irradiance for photosynthesis, results show that the interaction of canopy density and canopy-scale photosynthetic response is complex and non-linear, due to the combination of self-shading and the non-linearity of photosynthesis versus irradiance (P-I) curves near saturating irradiance. Therefore studies of light limitation in seagrasses should consider variation in canopy structure and density. Based on empirical work, we propose a number of possible measures for canopy scale photosynthetic response that can be plotted to yield isoclines in the space of canopy density and light environment. These plots can be used to interpret the significance of canopy changes induced as a response to decreases in the benthic light environment: in some cases canopy thinning can lead to an equivalent leaf level light environment, in others physiological changes may also be required but these alone may be inadequate for canopy survival. By providing insight to these processes the methods developed here could be a valuable management tool for seagrass conservation during dredging or other coastal developments. PMID:25347849

Short- and long-term modulation of the lutein epoxide and violaxanthin cycles in two species of the Lauraceae: sweet bay laurel (Laurus nobilis L.) and avocado (Persea americana Mill.).

PubMed

Esteban, R; Jiménez, M S; Morales, D; Jiménez, E T; Hormaetxe, K; Becerril, J M; Osmond, B; García-Plazaola, J I

2008-05-01

Short- and long-term responses of the violaxanthin (V) and lutein epoxide (Lx) cycles were studied in two species of Lauraceae: sweet bay laurel (Laurus nobilis L.) and avocado (Persea americana L.). The Lx content exceeded the V content in shade leaves of both species. Both Lx and V were de-epoxidised on illumination, but only V was fully restored by epoxidation in low light. Violaxanthin was preferentially de-epoxidised in low light in L. nobilis. This suggests that Lx accumulates with leaf ageing, partly because its conversion to lutein is limited in shade. After exposure to strong light, shade leaves of avocado readjusted the total pools of alpha- and beta-xanthophyll cycles by de novo synthesis of antheraxanthin, zeaxanthin and lutein. This occurred in parallel with a sustained depression of F(v)/F(m). In Persea indica, a closely related but low Lx species, F(v)/F(m) recovered faster after a similar light treatment, suggesting the involvement of the Lx cycle in sustained energy dissipation. Furthermore, the seasonal correlation between non-reversible Lx and V photoconversions and pre-dawn F(v)/F(m) in sun leaves of sweet bay supported the conclusion that the Lx cycle is involved in a slowly reversible downregulation of photosynthesis analogous to the V cycle.

Shading aboveground L-joint and lap-joint tests : comparison of white pine and sugar maple test assemblies

Treesearch

Carol A. Clausen; Daniel L. Lindner

2011-01-01

Five-year performance ratings are presented for two types of untreated, uncoated wood joints (L and lap) in aboveground tests under shaded conditions. The effect of shading on moisture entrapment in pine and maple L and lap joints was evaluated in a moderate decay zone (Madison, Wisconsin). Variations were observed between wood species, visual ratings, joint type,...

Sacramento's parking lot shading ordinance: environmental and economic costs of compliance

Treesearch

E.G. McPherson

2001-01-01

A survey of 15 Sacramento parking lots and computer modeling were used to evaluate parking capacity and compliance with the 1983 ordinance requiring 50% shade of paved areas (PA) 15 years after development. There were 6% more parking spaces than required by ordinance, and 36% were vacant during peak use periods. Current shade was 14% with 44% of this amount provided by...

Shade factors for 149 taxa of in-leaf urban trees in the USA

Treesearch

E. Gregory McPherson; Qingfu Xiao; Natalie S. van Doorn; Nels Johnson; Shannon Albers; Paula J. Peper

2018-01-01

Shade factors, defined as the percentage of sky covered by foliage and branches within the perimeter of individual tree crowns, have been used to model the effects of trees on air pollutant uptake, building energy use and rainfall interception. For the past 30 years the primary source of shade factors was a database containing values from 47 species. In most...

The repeatability of an intraoral dental colorimeter.

PubMed

Tung, Francis F; Goldstein, Gary R; Jang, Sungkoo; Hittelman, Eugene

2002-12-01

Characterizing and reproducing color remain one of the most challenging aspects of dentistry. A relatively new intraoral colorimeter measures the color of natural teeth and metal-ceramic restorations and prints out a color recipe for the Vintage Halo Porcelain System. The reliability of the colorimeter is unknown. The purpose of this study was to evaluate the reliability of a contact dental colorimeter and to correlate the shade registered by the colorimeter with the shade selected by experienced clinicians. In part I of the study, 2 examiners (A and B) took 2 colorimeter measurements from the maxillary right central incisors of 11 subjects. The examiners were blinded to their own data and those of other investigators. The readings were repeated 3 weeks later with the same protocol. The Cronbach alpha reliability coefficient was used to analyze the collected data. In part II of the study, 2 experienced clinicians (examiners D and E) selected a shade from the classic Vita Lumin Vacuum shade guide for the maxillary right central incisors of the same 11 subjects. The clinicians were blinded to each other's selections and the colorimeter readings. It should be noted that the manufacturer of the colorimeter uses the terms shade, value, and hue to represent chroma, value, and hue, respectively, as defined in the Glossary of Prosthodontic Terms (J Prosthet Dent 1999;81:39-110). The reliability analysis results for each of the combined trials for shade, value, and hue were all >.94. The interexaminer reliability alpha values were >.9 for shade and value and.64 to.74 for hue. The interexaminer alpha represented the value range of each of 4 measurements. The intraexaminer reliability alpha values for shade, value, and hue were.99,.95, and.96 for examiner A and.99,.93, and.97 for examiner B, respectively. In part II of the study, the colorimeter agreed with itself 82% of the time, whereas clinicians agreed with each other on the selected shade 73% of the time. Selections made by the colorimeter and the clinicians matched 55% to 64% of the time. Within the limitations of this study, the colorimeter reliably measured the color of natural teeth.

Iterative CT shading correction with no prior information

NASA Astrophysics Data System (ADS)

Wu, Pengwei; Sun, Xiaonan; Hu, Hongjie; Mao, Tingyu; Zhao, Wei; Sheng, Ke; Cheung, Alice A.; Niu, Tianye

2015-11-01

Shading artifacts in CT images are caused by scatter contamination, beam-hardening effect and other non-ideal imaging conditions. The purpose of this study is to propose a novel and general correction framework to eliminate low-frequency shading artifacts in CT images (e.g. cone-beam CT, low-kVp CT) without relying on prior information. The method is based on the general knowledge of the relatively uniform CT number distribution in one tissue component. The CT image is first segmented to construct a template image where each structure is filled with the same CT number of a specific tissue type. Then, by subtracting the ideal template from the CT image, the residual image from various error sources are generated. Since forward projection is an integration process, non-continuous shading artifacts in the image become continuous signals in a line integral. Thus, the residual image is forward projected and its line integral is low-pass filtered in order to estimate the error that causes shading artifacts. A compensation map is reconstructed from the filtered line integral error using a standard FDK algorithm and added back to the original image for shading correction. As the segmented image does not accurately depict a shaded CT image, the proposed scheme is iterated until the variation of the residual image is minimized. The proposed method is evaluated using cone-beam CT images of a Catphan©600 phantom and a pelvis patient, and low-kVp CT angiography images for carotid artery assessment. Compared with the CT image without correction, the proposed method reduces the overall CT number error from over 200 HU to be less than 30 HU and increases the spatial uniformity by a factor of 1.5. Low-contrast object is faithfully retained after the proposed correction. An effective iterative algorithm for shading correction in CT imaging is proposed that is only assisted by general anatomical information without relying on prior knowledge. The proposed method is thus practical and attractive as a general solution to CT shading correction.

Stimulus conflict triggers behavioral avoidance.

PubMed

Dignath, David; Eder, Andreas B

2015-12-01

According to a recent extension of the conflict-monitoring theory, conflict between two competing response tendencies is registered as an aversive event and triggers a motivation to avoid the source of conflict. In the present study, we tested this assumption. Over five experiments, we examined whether conflict is associated with an avoidance motivation and whether stimulus conflict or response conflict triggers an avoidance tendency. Participants first performed a color Stroop task. In a subsequent motivation test, participants responded to Stroop stimuli with approach- and avoidance-related lever movements. These results showed that Stroop-conflict stimuli increased the frequency of avoidance responses in a free-choice motivation test, and also increased the speed of avoidance relative to approach responses in a forced-choice test. High and low proportions of response conflict in the Stroop task had no effect on avoidance in the motivation test. Avoidance of conflict was, however, obtained even with new conflict stimuli that had not been presented before in a Stroop task, and when the Stroop task was replaced with an unrelated filler task. Taken together, these results suggest that stimulus conflict is sufficient to trigger avoidance.

Radiant heat loss, an unexploited path for heat stress reduction in shaded cattle.

PubMed

Berman, A; Horovitz, T

2012-06-01

Reducing thermal radiation on shaded animals reduces heat stress independently of other means of stress relief. Radiant heat exchange was estimated as a function of climate, shade structure, and animal density. Body surface portion exposed to radiant sources in shaded environments was determined by geometrical relations to determine angles of view of radiation sources (roof underside, sky, sun-exposed ground, shaded ground) on the animal's surface. The relative representation of environment radiation sources on the body surface was determined. Animal thermal radiation balance was derived from radiant heat gained from radiation sources (including surrounding animals) and that lost from the animal surface. The animal environment was assumed to have different shade dimensions and temperatures. These were summed to the radiant heat balance of the cow. The data formed served to estimate the effect of changes in intensity of radiation sources, roof and shaded surface dimensions, and animal density on radiant heat balance (Rbal) of cattle. Roof height effect was expressed by effect of roof temperature on Rbal. Roof underside temperature (35 to 75°C) effect on Rbal was reduced by roof height. If roof height were 4m, an increase in its underside temperature from 35 to 75°C would increase mean Rbal from -63 to -2 W·m⁻², whereas if roof height were 10 m, Rbal would only increase from -99 to -88 W·m⁻². A hot ground temperature increase from 35 to 65°C reduced mean Rbal heat loss from -45 to 3 W·m⁻². Increasing the surface of the shaded area had only a minor effect on Rbal and on the effect of hot ground on Rbal. Increasing shade roof height reduced the effect of roof temperature on Rbal to minor levels when height was > 8m. Increasing the roof height from 4 to 10 m decreased Rbal from -32 to -94 W·m⁻². Increasing indirect radiation from 100 to 500 W·m⁻² was associated with an increase in Rbal from -135 to +23 W·m⁻². Their combined effects were lower Rbal with increasing roof height and a reduction in rate of decrease with increasing level of indirect radiation. Roof height as an Rbal attenuator declined with increasing indirect radiation level. The latter factor might be reduced by lowering roof surface radiation absorption and through roof heat transfer, as well as by use of shade structure elements to reduce indirect radiation in the shaded area. Radiant heat from the cow body surface may be reduced by lower cow density. Radiant heat attenuation may thus further elevate animal productivity in warm climates, with no associated operation costs. Copyright © 2012 American Dairy Science Association. Published by Elsevier Inc. All rights reserved.

Evaluation of the effects of plant-derived essential oils on central nervous system function using discrete shuttle-type conditioned avoidance response in mice.

PubMed

Umezu, Toyoshi

2012-06-01

Although plant-derived essential oils (EOs) have been used to treat various mental disorders, their central nervous system (CNS) acting effects have not been clarified. The present study compared the effects of 20 kinds of EOs with the effects of already-known CNS acting drugs to examine whether the EOs exhibited CNS stimulant-like effects, CNS depressant-like effects, or neither. All agents were tested using a discrete shuttle-type conditioned avoidance task in mice. Essential oils of peppermint and chamomile exhibited CNS stimulant-like effects; that is, they increased the response rate (number of shuttlings/min) of the avoidance response. Linden also increased the response rate, however, the effect was not dose-dependent. In contrast, EOs of orange, grapefruit, and cypress exhibited CNS depressant-like effects; that is, they decreased the response rate of the avoidance response. Essential oils of eucalyptus and rose decreased the avoidance rate (number of avoidance responses/number of avoidance trials) without affecting the response rate, indicating that they may exhibit some CNS acting effects. Essential oils of 12 other plants, including juniper, patchouli, geranium, jasmine, clary sage, neroli, lavender, lemon, ylang-ylang, niaouli, vetivert and frankincense had no effect on the avoidance response in mice. Copyright © 2011 John Wiley & Sons, Ltd.

Individual phenolic response and peroxidase activity in peel of differently sun-exposed apples in the period favorable for sunburn occurrence.

PubMed

Zupan, Anka; Mikulic-Petkovsek, Maja; Slatnar, Ana; Stampar, Franci; Veberic, Robert

2014-11-15

Extreme weather events like high solar radiation can cause stress in apple fruits (Malus domestica Borkh.). The aim of the study was to make a screening of individual phenols and peroxidase activity in apple peel as a response to sunburn and different sun-exposures in the period when weather conditions are suitable for sunburn occurrence. Apple fruits of 'Golden Delicious' and 'Braeburn' were sampled. Fruit temperature and color were measured prior HPLC-MS(2) and peroxidase activity analyses. Sunburned peel was darker and more yellow-red in comparison to healthy peel, which appeared yellow-green. Fruit temperature, total as well as individual flavonols and dihydrochalcones, total hydroxycinnamics and perixodase activity were highest in sunburned peel in comparison with healthy sun-exposed peel, furthermore both were different than shaded sides of both fruits and peel of apples inside the tree crown; moreover in sunburned peel dihydrochalcones were determined for the first time. Chlorogenic acid was up to 2.5 times higher, 3-hydroxy-phloretin-2'-O-xyloglucoside was up to 10 times higher and quercetin-3-galactoside was up to 33 times higher in sunburned peel, comparing to shaded sided peels. Flavanols did not show a distinct pattern. A deeper insight in phenolic response against environmental stress caused by high solar radiation and high air temperatures has been made. Copyright © 2014 Elsevier GmbH. All rights reserved.