Sample records for stand density determination

  1. New Method for Determining the Relative Stand Density of Forest Inventory Plots

    Treesearch

    Christopher W. Woodall; Patrick D. Miles

    2006-01-01

    Determining the relative density of Forest Inventory and Analysis plots is complicated by the various species and tree size combinations in the Nation?s forested ecosystems. Stand density index (SDI), although developed for use in even-aged monocultures, has been used for stand density assessment in largescale forest inventories. To improve application of SDI in uneven...

  2. Determining maximum stand density index in mixed species stands for strategic-scale stocking assessments

    Treesearch

    Chris W. Woodall; Patrick D. Miles; John S. Vissage

    2005-01-01

    Stand density index (SDI), although developed for use in even-aged monocultures, has been used for assessing stand density in large-scale forest inventories containing diverse tree species and size distributions. To improve application of SDI in unevenaged, mixed species stands present in large-scale forest inventories, trends in maximum SDI across diameter classes...

  3. How to formulate and solve "optimal stand density over time" problems for even-aged stands using dynamic programming.

    Treesearch

    Chung M. Chen; Dietmar W. Rose; Rolfe A. Leary

    1980-01-01

    Describes how dynamic programming can be used to solve optimal stand density problems when yields are given by prior simulation or by a new stand growth equation that is a function of the decision variable. Formulations of the latter type allow use of a calculus-based search procedure; they determine exact optimal residual density at each stage.

  4. Southern pine beetle in loblolly pine: simulating within stand interactions using the process model SPBLOBTHIN

    Treesearch

    Brian Strom; J. R. Meeker; J. Bishir; James Roberds; X. Wan

    2016-01-01

    Pine stand density is a key determinant of damage resulting from attacks by the southern pine beetle (SPB, Dendroctonus frontalis Zimm.). High-density stands of maturing loblolly pine (Pinus taeda L.) are at high risk for losses to SPB, and reducing stand density is the primary tool available to forest managers for preventing and mitigating damage. Field studies are...

  5. Effects of thinning on drought vulnerability and climate response in north temperate forest ecosystems

    USGS Publications Warehouse

    D’Amato, Anthony W.; Bradford, John B.; Fraver, Shawn; Palik, Brian J.

    2013-01-01

    Reducing tree densities through silvicultural thinning has been widely advocated as a strategy for enhancing resistance and resilience to drought, yet few empirical evaluations of this approach exist. We examined detailed dendrochronological data from a long-term (>50 yrs) replicated thinning experiment to determine if density reductions conferred greater resistance and/or resilience to droughts, assessed by the magnitude of stand-level growth reductions. Our results suggest that thinning generally enhanced drought resistance and resilience; however, this relationship showed a pronounced reversal over time in stands maintained at lower tree densities. Specifically, lower-density stands exhibited greater resistance and resilience at younger ages (49 years), yet exhibited lower resistance and resilience at older ages (76 years), relative to higher-density stands. We attribute this reversal to significantly greater tree sizes attained within the lower-density stands through stand development, which in turn increased tree-level water demand during the later droughts. Results from response-function analyses indicate that thinning altered growth-climate relationships, such that higher-density stands were more sensitive to growing-season precipitation relative to lower-density stands. These results confirm the potential of density management to moderate drought impacts on growth, and they highlight the importance of accounting for stand structure when predicting climate-change impacts to forest systems.

  6. Effects of thinning on drought vulnerability and climate response in north temperate forest ecosystems.

    PubMed

    D'Amato, Anthony W; Bradford, John B; Fraver, Shawn; Palik, Brian J

    2013-12-01

    Reducing tree densities through silvicultural thinning has been widely advocated as a strategy for enhancing resistance and resilience to drought, yet few empirical evaluations of this approach exist. We examined detailed dendrochronological data from a long-term (> 50 years) replicated thinning experiment to determine if density reductions conferred greater resistance and/or resilience to droughts, assessed by the magnitude of stand-level growth reductions. Our results suggest that thinning generally enhanced drought resistance and resilience; however, this relationship showed a pronounced reversal over time in stands maintained at lower tree densities. Specifically, lower-density stands exhibited greater resistance and resilience at younger ages (49 years), yet exhibited lower resistance and resilience at older ages (76 years), relative to higher-density stands. We attribute this reversal to significantly greater tree sizes attained within the lower-density stands through stand development, which in turn increased tree-level water demand during the later droughts. Results from response-function analyses indicate that thinning altered growth-climate relationships, such that higher-density stands were more sensitive to growing-season precipitation relative to lower-density stands. These results confirm the potential of density management to moderate drought impacts on growth, and they highlight the importance of accounting for stand structure when predicting climate-change impacts to forests.

  7. Effects of planting density on the Biomass partitioning of intensively managed loblolly pine stands on the Piedmont and upper Coastal plain of Georgia

    Treesearch

    Rodney E. Will; Nikhil Narahari; Robert O. Teskey; Barry D. Shiver; Matthew Wosotowsky

    2006-01-01

    Increased planting density enhances overall stand growth by increasing resource capture and use. However, planting density also may affect the proportion of biomass partitioned to stem growth, a main factor controlling stand growth and yield. During the fourth growing season, we determined the biomass partitioned to leaf, branch, stem, and fine root (> 0.5mm) of...

  8. Crown management and stand density

    Treesearch

    Thomas J. Dean; V. Clark Baldwin

    1996-01-01

    Determination of optimal stand-density continues to be a difficult problem. A trial cannot be established on every combination of soils, topography, and climate possible across the range of a widely distributed species such as loblolly pine, and continual advancements in nutrition and vegetation management, breeding, and utilization make established trials obsolete....

  9. Effects of planting density and genotype on loblolly pine stands growing in the mountains of southeastern Oklahoma

    Treesearch

    Rodney E. Will; Thomas C. Hennessey; Thomas B. Lynch; Robert Heinemann; Randal Holeman; Dennis Wilson; Keith Anderson; Gregory Campbell

    2013-01-01

    We determined the effects of planting density (4- by 4-, 6- by 6-, 8- by 8-, and 10- by 10-foot spacing) on stand-level height, diameter at breast height, stem volume, basal area, and periodic annual increment for two loblolly pine (Pinus taeda L.) seed sources. Seed sources for the 25-year-old stands were a North Carolina seed source (NCC 8-01) and...

  10. Stand quality management of a water oak plantation in Louisiana: preliminary results following thinning

    Treesearch

    James S. Meadows; Daniel A., Jr. Skojac

    2010-01-01

    Stand quality management is a new guiding principle in which thinning prescriptions are based on tree quality rather than on residual stand density. We recently initiated a series of hardwood thinning studies to determine the effects of four stand quality management thinning prescriptions on both stand-level and individual-tree-level growth, quality, and value: (1) no...

  11. [Rapid prediction of annual ring density of Paulownia elongate standing tress using near infrared spectroscopy].

    PubMed

    Jiang, Ze-Hui; Wang, Yu-Rong; Fei, Ben-Hua; Fu, Feng; Hse, Chung-Yun

    2007-06-01

    Rapid prediction of annual ring density of Paulownia elongate standing trees using near infrared spectroscopy was studied. It was non-destructive to collect the samples for trees, that is, the wood cores 5 mm in diameter were unthreaded at the breast height of standing trees instead of fallen trees. Then the spectra data were collected by autoscan method of NIR. The annual ring density was determined by mercury immersion. And the models were made and analyzed by the partial least square (PLS) and full cross validation in the 350-2 500 nm wavelength range. The results showed that high coefficients were obtained between the annual ring and the NIR fitted data. The correlation coefficient of prediction model was 0.88 and 0.91 in the middle diameter and bigger diameter, respectively. Moreover, high coefficients of correlation were also obtained between annual ring density laboratory-determined and the NIR fitted data in the middle diameter of Paulownia elongate standing trees, the correlation coefficient of calibration model and prediction model were 0.90 and 0.83, and the standard errors of calibration (SEC) and standard errors of prediction(SEP) were 0.012 and 0.016, respectively. The method can simply, rapidly and non-destructively estimate the annual ring density of the Paulownia elongate standing trees close to the cutting age.

  12. Snow accumulation under various forest stand densities at Tenderfoot Creek Experimental Forest, Montana, USA

    Treesearch

    Chadwick A. Moore; Ward W. McCaughey

    1997-01-01

    Snow accumulation in forested watersheds is controlled by climate, elevation, topographic factors and vegetation structure. Conifers affect snow accumulation principally by intercepting snow with the canopy which may later be sublimated. Various tree, stand, species and canopy densities of a subalpine fir habitat (ALBANASC) in central Montana were studied to determine...

  13. Thinning ponderosa pine (Pinus ponderosa) stands reduces mortality while maintaining stand productivity

    Treesearch

    Jianwei Zhang; Martin W. Ritchie; Douglas A. Maguire; William W. Oliver

    2013-01-01

    We analyzed 45-yr data collected from three ponderosa pine (Pinus ponderosa Lawson & C. Lawson) levels-of-growing-stock installations in Oregon (OR) and northern California (CA), USA, to determine the effect of stand density regimes on stand productivity and mortality. We found that periodic annual increment (PAI) of diameter, basal area (BA...

  14. Effects of planting density and genotype on canopy size, canopy structure, and growth of 25-year-old loblolly pine stands in southeastern Oklahoma

    Treesearch

    Thomas C. Hennessey; Rodney E. Will; Thomas B. Lynch; Robert Heinemann; Randal Holeman; Dennis Wilson; Keith Anderson; Gregory Campbell

    2013-01-01

    Leaf biomass and its display within the canopy are important driving variables of stand growth because they reflect a tree or stand’s capacity to intercept radiation, reduce carbon dioxide, and transpire water. We determined the effects of planting density (4- by 4-, 6- by 6-, 8- by 8-, and 10- by 10-foot spacing) on annual needle fall biomass, intercepted radiation,...

  15. Impact of forest type and management strategy on avian densities in the Mississippi Alluvial Valley, USA

    USGS Publications Warehouse

    Twedt, D.J.; Wilson, R.R.; Henne-Kerr, J.L.; Hamilton, R.B.

    1999-01-01

    Avian territory densities were determined from 20 Breeding Bird Censuses in mature (>30 years) bottomland hardwood stand: and 18 Breeding Bird Censuses in young (6-9 years old) cottonwood (Populas deltoides) plantations in the Mississippi Alluvial Valley. Avian species richness, diversity, and territory density were greater (p 0.05). Even so, detrended correspondence analysis based on avian territory densities readily segregated forest types and silvicultural treatments. Timber harvest within bottomland hardwood stands resulted in a shift in bird communities toward those found in cottonwood stands by increasing the densities of early-successional species such as Indigo Bunting (Passerina cyanea), Yellow-breasted Chat (Icteria virens), and Common Yellowthroat (Geothlypis trichas). Conversely, regenerating cottonwood stands from root sprouts, rather than planting stem cuttings, resulted in a shift in bird communities toward those found in bottomland hardwood stands by increasing densities of species such as White-eyed Vireo (Vireo griseus) and Wood Thrush (Hylocichla mustelina). Tree species diversity, angular canopy cover, and midstory density were positively associated with bird species assemblages in bottomland hardwood stands, whereas vegetation density at ground level was positively associated with bird communities in cottonwood plantations. Conversion of agricultural fields to short-rotation cottonwood plantations results in increased breeding bird populations by adding up to 140 additional territories 40 ha-1. Even so, relative conservation values, derive, from indicator species analysis and Partners in Flight concern scores, suggest that mature bottomland hardwood forests are twice as 'valuable' for bird conservation as are cottonwood plantations.

  16. A critical analysis of methods for rapid and nondestructive determination of wood density in standing trees

    Treesearch

    Shan Gao; Xiping Wang; Michael C. Wiemann; Brian K. Brashaw; Robert J. Ross; Lihai Wang

    2017-01-01

    Key message Field methods for rapid determination of wood density in trees have evolved from increment borer, torsiometer, Pilodyn, and nail withdrawal into sophisticated electronic tools of resistance drilling measurement. A partial resistance drilling approach coupled with knowledge of internal tree density distribution may...

  17. Pre-outbreak forest conditions mediate the effects of spruce beetle outbreaks on fuels in subalpine forests of Colorado.

    PubMed

    Mietkiewicz, Nathan; Kulakowski, Dominik; Veblen, Thomas T

    2018-03-01

    Over the past 30 years, forest disturbances have increased in size, intensity, and frequency globally, and are predicted to continue increasing due to climate change, potentially relaxing the constraints of vegetation properties on disturbance regimes. However, the consequences of the potentially declining importance of vegetation in determining future disturbance regimes are not well understood. Historically, bark beetles preferentially attack older trees and stands in later stages of development. However, as climate warming intensifies outbreaks by promoting growth of beetle populations and compromising tree defenses, smaller diameter trees and stands in early stages of development now are being affected by outbreaks. To date, no study has considered how stand age and other pre-outbreak forest conditions mediate the effects of outbreaks on surface and aerial fuel arrangements. We collected fuels data across a chronosequence of post-outbreak sites affected by spruce beetle (SB) between the 1940s and the 2010s, stratified by young (<130 yr) and old (>130 yr) post-fire stands. Canopy and surface fuel loads were calculated for each tree and stand, and available crown fuel load, crown bulk density, and canopy bulk densities were estimated. Canopy bulk density and density of live canopy individuals were reduced in all stands affected by SB, though foliage loss was proportionally greater in old stands as compared to young stands. Fine surface fuel loads in young stands were three times greater shortly (<30 yr) following outbreak as compared to young stands not affected by outbreak, after which the abundance of fine surface fuels decreased to below endemic (i.e., non-outbreak) levels. In both young and old stands, the net effect of SB outbreaks during the 20th and 21st centuries reduced total canopy fuels and increased stand-scale spatial heterogeneity of canopy fuels following outbreak. Importantly, the decrease in canopy fuels following outbreaks was greater in young post-fire stands than in older stands, suggesting that SB outbreaks may more substantially reduce risk of active crown fire when they affect stands in earlier stages of development. The current study shows that the effects of SB outbreaks on forest structure and on fuel profiles are strongly contingent on pre-outbreak conditions as determined by pre-outbreak disturbance history. © 2018 by the Ecological Society of America.

  18. Twenty-four years after theYellowstone Fires: Are postfire lodgepole pine stands converging in structure and function?

    PubMed

    Turner, Monica G; Whitby, Timothy G; Tinker, Daniel B; Romme, William H

    2016-05-01

    Disturbance and succession have long been of interest in ecology, but how landscape patterns of ecosystem structure and function evolve following large disturbances is poorly understood. After nearly 25 years, lodgepole pine (Pinus contorta var. latifolia) forests that regenerated after the 1988 Yellowstone Fires (Wyoming, USA) offer a prime opportunity to track the fate of disturbance-created heterogeneity in stand structure and function in a wilderness setting. In 2012, we resampled 72 permanent plots to ask (1) How have postfire stand structure and function changed between 11 and 24 yr postfire, and what variables explain these patterns and changes? (2) How has landscape-level (among-stand) variability in postfire stand structure and function changed between 11 and 24 yr postfire? We expected to see evidence of convergence beginning to emerge, but also that initial postfire stem density would still determine trajectories of biomass accumulation. After 24 yr, postfire lodgepole pine density remained very high (mean = 21,738 stems/ha, range = 0-344,067 stems/ha). Stem density increased in most plots between 11 and 24 yr postfire, but declined sharply where 11-yr-postfire stem density was > 72,000 stems/ha. Stems were small in high-density stands, but stand-level lodgepole pine leaf area, foliage biomass, and live aboveground biomass increased over time and with increasing stem density. After 24 yr, mean annual lodgepole pine aboveground net primary production (ANPP) was high (mean = 5 Mg · ha⁻¹ · yr⁻¹, range = 0-16.5 Mg · ha⁻¹ · yr⁻¹). Among stands, lodgepole pine ANPP increased with stem density, which explained 69% of the variation; another 8% of the variation was explained by environmental covariates. Early patterns of postfire lodgepole pine regeneration, which were contingent on prefire serotiny and fire severity, remained the dominant driver of stand structure and function. We observed mechanisms that would lead to convergence in stem density (structure) over time, but it was landscape variation in functional variables that declined substantially. Stand structure and function have not converged across the burned landscape, but our evidence suggests function will converge sooner than structure.

  19. Cereal Leaf Beetle (Coleoptera: Chrysomelidae) Regional Dispersion and Relationship With Wheat Stand Denseness.

    PubMed

    Reisig, Dominic D; Bacheler, Jack S; Herbert, D Ames; Heiniger, Ron; Kuhar, Thomas; Malone, Sean; Philips, Chris; Tilley, M Scott

    2017-06-01

    Cereal leaf beetle, Oulema melanopus L., is a pest of small grains and the literature conflicts on whether it is more abundant in sparse or dense stands of wheat. Our objectives were to determine the impact of stand denseness on cereal leaf beetle abundance and to investigate the regional dispersion of cereal leaf beetles across North Carolina and Virginia. One-hundred twenty fields were sampled across North Carolina and Virginia during 2011 for stand denseness, and cereal leaf beetle eggs, larvae, and adults. Two small-plot wheat experiments were planted in North Carolina using a low and a high seeding rate. Main plots were split, with one receiving a single nitrogen application and one receiving two. Egg density, but not larva or adult density, was positively correlated with stand denseness in the regional survey. Furthermore, regional spatial patterns of aggregation were noted for both stand denseness and egg number. In the small-plot experiments, seeding rate influenced stand denseness, but not nitrogen application. In one experiment, egg densities per unit area were higher in denser wheat, while in the other experiment, egg densities per tiller were lower in denser wheat. Larvae were not influenced by any factor. Overall, there were more cereal leaf beetle eggs in denser wheat stands. Previous observations that sparse stands of wheat are more prone to cereal leaf beetle infestation can be attributed to the fact that sparser stands have fewer tillers, which increases the cereal leaf beetle to tiller ratio compared with denser stands. © The Authors 2017. Published by Oxford University Press on behalf of Entomological Society of America. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

  20. Complex mountain terrain and disturbance history drive variation in forest aboveground live carbon density in the western Oregon Cascades, USA

    PubMed Central

    Zald, Harold S.J.; Spies, Thomas A.; Seidl, Rupert; Pabst, Robert J.; Olsen, Keith A.; Steel, E. Ashley

    2016-01-01

    Forest carbon (C) density varies tremendously across space due to the inherent heterogeneity of forest ecosystems. Variation of forest C density is especially pronounced in mountainous terrain, where environmental gradients are compressed and vary at multiple spatial scales. Additionally, the influence of environmental gradients may vary with forest age and developmental stage, an important consideration as forest landscapes often have a diversity of stand ages from past management and other disturbance agents. Quantifying forest C density and its underlying environmental determinants in mountain terrain has remained challenging because many available data sources lack the spatial grain and ecological resolution needed at both stand and landscape scales. The objective of this study was to determine if environmental factors influencing aboveground live carbon (ALC) density differed between young versus old forests. We integrated aerial light detection and ranging (lidar) data with 702 field plots to map forest ALC density at a grain of 25 m across the H.J. Andrews Experimental Forest, a 6369 ha watershed in the Cascade Mountains of Oregon, USA. We used linear regressions, random forest ensemble learning (RF) and sequential autoregressive modeling (SAR) to reveal how mapped forest ALC density was related to climate, topography, soils, and past disturbance history (timber harvesting and wildfires). ALC increased with stand age in young managed forests, with much greater variation of ALC in relation to years since wildfire in old unmanaged forests. Timber harvesting was the most important driver of ALC across the entire watershed, despite occurring on only 23% of the landscape. More variation in forest ALC density was explained in models of young managed forests than in models of old unmanaged forests. Besides stand age, ALC density in young managed forests was driven by factors influencing site productivity, whereas variation in ALC density in old unmanaged forests was also affected by finer scale topographic conditions associated with sheltered sites. Past wildfires only had a small influence on current ALC density, which may be a result of long times since fire and/or prevalence of non-stand replacing fire. Our results indicate that forest ALC density depends on a suite of multi-scale environmental drivers mediated by complex mountain topography, and that these relationships are dependent on stand age. The high and context-dependent spatial variability of forest ALC density has implications for quantifying forest carbon stores, establishing upper bounds of potential carbon sequestration, and scaling field data to landscape and regional scales. PMID:27041818

  1. Reineke’s stand density index: a quantitative and non-unitless measure of stand density

    Treesearch

    Curtis L. VanderSchaaf

    2013-01-01

    When used as a measure of relative density, Reineke’s stand density index (SDI) can be made unitless by relating the current SDI to a standard density but when used as a quantitative measure of stand density SDI is not unitless. Reineke’s SDI relates the current stand density to an equivalent number of trees per unit area in a stand with a quadratic mean diameter (Dq)...

  2. Relation of quantity of seed sown and density of seedlings to the development and survival of forest planting stock

    Treesearch

    W. G. Wahlenberg

    1929-01-01

    It is obvious that seedlings grown in dense stands can not develop so well as those grown without crowding. Nurserymen naturally wish to avoid injury to their stock from crowding, but they also desire to utilize their soil space as fully as possible. The optimum density of stand for each species and age class of nursery stock can be determined within reasonably close...

  3. Thinning results from a mixed upland hardwood stand after 35 years

    Treesearch

    Ronald J., Jr. Myers; Kenneth R. Roeder; W. Henry McNab

    2008-01-01

    A long-term study of precommercial thinning was installed in a 6-year-old oak-dominated stand regenerated by clearcutting in the southern Appalachian Mountains of North Carolina. Three levels of residual stand density were tested: control (no thinning), and 200, and 400 residual trees per acre (TPA). Objectives of the study were to determine the response of an upland...

  4. Sap Flow Estimate Of Watershed-Scale Transpiration

    NASA Astrophysics Data System (ADS)

    Kumagai, T.; Aoki, S.; Shimizu, T.; Otsuki, K.

    2006-12-01

    The present study examined how to obtain sufficient information to extrapolate watershed-scale transpiration in a Japanese cedar (Cryptomeria japonica D. Don) forest from sap flow measurements of available individual trees. In this study, we conducted measurements of tree biometrics and tree-to-tree and radial variations in xylem sap flux density (Fd) in two different stand plots, an upper slope plot (UP) and lower slope plot (LP), during the growing season with significant variations in environmental factors. The manner in which the mean stand sap flux density (JS) and tree stem allometric relationship (diameter at breast height (DBH) versus sapwood area (AS_tree)) vary between the two stands along the slope of the watershed was then investigated. After these analyses, appropriate sample sizes for estimations of representative JS values in the stand were also determined. The results demonstrated that a unique or general function allowed description of the allometric relationship along the slope, but the data for its formulation needed to be obtained in both UP and LP. They also revealed that JS in UP and LP were almost the same during the study period despite differences in tree density and size between the two plots. This implies that JS measured in a partial stand within a watershed is a reasonable estimator of the values of other stands, and that stand sapwood area calculated by AS_tree is a strong determinant of water-use in a forest watershed. To estimate JS in both an UP and LP, at least 10 trees should be sampled, but not necessarily more than this.

  5. Stand density index in uneven-aged ponderosa pine stands

    Treesearch

    C.W. Woodall; C.E. Fiedler; K.S. Milner

    2003-01-01

    Stand density index (SDI) was developed to quantify relative stand density in even-aged stands. Application of SDI in uneven-aged stands has been described mathematically but not justified biologically. Diameter-class trends in SDI and sapwood area across 14 uneven-aged ponderosa pine (Pinus ponderosa Dougl. ex P. & C. Laws.) stands in eastern...

  6. Stand structure and growth of Abies magnifica responded to five thinning levels in northeastern California, USA

    Treesearch

    Jianwei Zhang; William W. Oliver

    2006-01-01

    A 60-year old red fir stand with 23,950 stems per ha was thinned to five stand densities. Thinning occurred in 1972, 1976, and 1980. Height, DBH, and crown characteristics were measured seven times at four- to seven-year intervals from 1972 to 2002. Tree rings were measured retrospectively to determine growth of individual years. Periodic annual increment (PAI) was...

  7. DOE Office of Scientific and Technical Information (OSTI.GOV)

    Zarnoch, Stanley J.; Blake, John I.; Parresol, Bernard R.

    Snags are standing dead trees that are an important component in the nesting habitat of birds and other species. Although snag availability is believed to limit populations in managed and non-managed forests, little data are available to evaluate the relative effect of stand conditions and management on snag occurrence. We analyzed point sample data from an intensive forest inventory within an 80,000 ha landscape for four major forest types to support the hypotheses that routine low-intensity prescribed fire would increase, and thinning would decrease, snag occurrence. We employed path analysis to define a priori causal relationships to determine the directmore » and indirect effects of site quality, age, relative stand density index and fire for all forest types and thinning effects for loblolly pine and longleaf pine. Stand age was an important direct effect for loblolly pine, mixed pine-hardwoods and hardwoods, but not for longleaf pine. Snag occurrence in loblolly pine was increased by prescribed fire and decreased by thinning which confirmed our initial hypotheses. Although fire was not important in mixed pine-hardwoods, it was for hardwoods but the relationship depended on site quality. For longleaf pine the relative stand density index was the dominant variable affecting snag occurrence, which increased as the density index decreased. Site quality, age and thinning had significant indirect effects on snag occurrence in longleaf pine through their effects on the density index. Although age is an important condition affecting snag occurrence for most forest types, path analysis revealed that fire and density management practices within certain forest types can also have major beneficial effects, particularly in stands less than 60 years old.« less

  8. Natural seed fall in white pine (Pinus strobes L.) stands of varying density

    Treesearch

    Raymond E. Graber

    1970-01-01

    Seed fall was observed in three stands of mature white pines at stand basal-area densities of 80, 120, and 187 square feet per acre. It was found that the intermediate-density stand produced nearly 50 percent more seed than the stands of other densities. During a good seed year this stand produced 59 pounds of dry sound seed per acre. Most of the seeds were dispersed...

  9. Development of a stand density index equation for slash pine stands

    Treesearch

    Paul F. Doruska

    2002-01-01

    Stand density index (SDI) is commonly used as the basis for density management guides for even-aged forest stands. Many tree species follow the same self-thinning trajectory, allowing for the use of stand density index in such guides. Slash pine (Pinus elliottii Englem.) has been shown to depart from the self-thinning trajectory exhibited by other...

  10. Harvesting small trees for bio-energy

    Treesearch

    John Klepac; Robert Rummer; Jason Thompson

    2011-01-01

    A conventional whole-tree logging operation consisting of 4-wheeled and 3-wheeled saw-head feller-bunchers, two grapple skidders and a chipper that produces dirty chips was monitored across several stands and machine performance evaluated. Stands were inventoried to determine density, volume, and basal area per acre and will be used to relate machine performance to...

  11. Consistent definition and application of Reineke's Stand Density Index in silviculture and stand projection

    Treesearch

    John D. Shaw; James N. Long

    2010-01-01

    Reineke’s Stand Density Index (SDI) has been available to silviculturists for over 75 years, but application of this stand metric has been inconsistent. Originally described as a measurement of relative density in single-species, even-aged stands, it has since been generalized for use in uneven-aged stands and mixed-species stands. However, methods used to establish...

  12. The relative density of forests in the United States

    Treesearch

    Christopher W. Woodall; Charles H. Perry; Patrick D. Miles

    2006-01-01

    A relative stand density assessment technique, using the mean specific gravity of all trees in a stand to predict its maximum stand density index (SDI) and subsequently its relative stand density (current SDI divided by maximum SDI), was used to estimate the relative density of forests across the United States using a national-scale forest inventory. Live tree biomass...

  13. Are prescribed fire and thinning dominant processes affecting snag occurrence at a landscape scale?

    DOE PAGES

    Zarnoch, Stanley J.; Blake, John I.; Parresol, Bernard R.

    2014-11-01

    Snags are standing dead trees that are an important component in the nesting habitat of birds and other species. Although snag availability is believed to limit populations in managed and non-managed forests, little data are available to evaluate the relative effect of stand conditions and management on snag occurrence. We analyzed point sample data from an intensive forest inventory within an 80,000 ha landscape for four major forest types to support the hypotheses that routine low-intensity prescribed fire would increase, and thinning would decrease, snag occurrence. We employed path analysis to define a priori causal relationships to determine the directmore » and indirect effects of site quality, age, relative stand density index and fire for all forest types and thinning effects for loblolly pine and longleaf pine. Stand age was an important direct effect for loblolly pine, mixed pine-hardwoods and hardwoods, but not for longleaf pine. Snag occurrence in loblolly pine was increased by prescribed fire and decreased by thinning which confirmed our initial hypotheses. Although fire was not important in mixed pine-hardwoods, it was for hardwoods but the relationship depended on site quality. For longleaf pine the relative stand density index was the dominant variable affecting snag occurrence, which increased as the density index decreased. Site quality, age and thinning had significant indirect effects on snag occurrence in longleaf pine through their effects on the density index. Although age is an important condition affecting snag occurrence for most forest types, path analysis revealed that fire and density management practices within certain forest types can also have major beneficial effects, particularly in stands less than 60 years old.« less

  14. Winter wheat stand density determination and yield estimates from handheld and airborne scanners. [Montana

    NASA Technical Reports Server (NTRS)

    Aase, J. K.; Millard, J. P.; Siddoway, F. H. (Principal Investigator)

    1982-01-01

    Radiance measurements from handheld (Exotech 100-A) and air-borne (Daedalus DEI 1260) radiometers were related to wheat (Triticum aestivum L.) stand densities (simulated winter wheat winterkill) and to grain yield for a field located 11 km northwest of Sidney, Montana, on a Williams loam soil (fine-loamy, mixed Typic Argiborolls) where a semidwarf hard red spring wheat cultivar was needed to stand. Radiances were measured with the handheld radiometer on clear mornings throughout the growing season. Aircraft overflight measurements were made at the end of tillering and during the early stem extension period, and the mid-heading period. The IR/red ratio and normalized difference vegetation index were used in the analysis. The aircraft measurements corroborated the ground measurements inasmuch as wheat stand densities were detected and could be evaluated at an early enough growth stage to make management decision. The aircraft measurements also corroborated handheld measurements when related to yield prediction. The IR/red ratio, although there was some growth stage dependency, related well to yield when measured from just past tillering until about the watery-ripe stage.

  15. Harvest operations for density management: planning requirements, production, costs, stand damage, and recommendations

    Treesearch

    Loren D. Kellogg; Stephen J. Pilkerton

    2013-01-01

    Since the early 1990s, several studies have been undertaken to determine the planning requirements, productivity, costs, and residual stand damage of harvest operations in thinning treatments designed to promote development of complex forest structure in order to enhance ecological functioning and biological diversity. Th ese studies include the Oregon State...

  16. Natural regeneration of interior douglas-fir in the northern Rocky Mountains

    Treesearch

    Dennis E. Ferguson; Clinton E. Carlson

    1991-01-01

    The regeneration period is a crucial time in the life of a stand because it determines species composition and tree density for the new stand. Silviculturists prescribe treatments to favor adequate and timely conifer regeneration of economically important species that grow well. This paper presents research results that will help silviculturists identify factors...

  17. A simple index of stand density for Douglas-fir.

    Treesearch

    R.O. Curtis

    1982-01-01

    The expression RD = G/(Dg½), where G is basal area and Dg is quadratic mean stand diameter, provides a simple and convenient scale of relative stand density for Douglas-fir, equivalent to other generally accepted diameter-based stand density measures.

  18. Maximum stand density for ponderosa pine and red and white fir in northern California

    Treesearch

    William.W. Oliver; Fabian C.C. Uzoh

    1997-01-01

    Why are forest managers interested in quantifying maximum stand density? Nearly all conceivable management objectives dictate a stand density less than a biological maximum. Certainly, the notion that thinning dense stands increases growth on the remaining trees and reduces mortality is well-established in the literature. The interest in quantifying maximum stand...

  19. Stand Density and Canopy Gaps

    Treesearch

    Boris Zeide

    2004-01-01

    Estimation of stand density is based on a relationship between number of trees and their average diameter in fully stocked stands. Popular measures of density (Reineke’s stand density index and basal area) assume that number of trees decreases as a power function of diameter. Actually, number of trees drops faster than predicted by the power function because the number...

  20. Distribution-of-cut guides for thinning in Allegheny hardwoods: a review

    Treesearch

    Christopher A. Nowak; David A. Marquis

    1997-01-01

    Distribution-of-cut guidelines describe the amount of stand density to be removed from broad size classes of trees to attain a target residual stand density and stand structure. Current guides for thinning Allegheny hardwoods recommend that 75 percent of the cut relative stand density be taken from below the average stand diameter and 25 percent from above. These...

  1. Variable-density thinning in coast redwood: a comparison of marking strategies to attain stand variability

    Treesearch

    Kevin L. O' Hara; Lathrop P. Leonard; Christopher R. Keyes

    2012-01-01

    Variable-density thinning (VDT) is an emerging thinning method that attempts to enhance stand structural heterogeneity by deliberately thinning at different intensities throughout a stand. VDT may create stands with dense areas, open areas, and other areas that may be intermediate in density. Subsequent stand development forms a more varied structure than is...

  2. The glass transition temperature of thin films: A molecular dynamics study for a bead-spring model.

    PubMed

    Stevenson, Craig S; Curro, John G; McCoy, John D

    2017-05-28

    Molecular dynamics simulations were carried out on free-standing liquid films of different thicknesses h using a bead-spring model of 10 beads per chain. The glass transition temperatures, T g , of the various films were determined from plots of the internal energy versus temperature. We used these simulations to test the validity of our earlier conjecture that the glass transition of a confined liquid could be approximated by pre-averaging over the non-uniform density profile of the film. Using the density profiles from our simulations, we computed the average density of the free-standing films as a function of temperature. In all our film simulations we found, within the error of the simulation, that T g of the film occurred at the same density (or packing fraction) as the bulk system at the bulk glass transition temperature T g B . By equating these densities at their respective glass transition temperatures, as suggested by the simulations, we deduced that T g /T g B is proportional to h 0 /h. This is consistent with previous simulations and experimental data. Moreover, the parameter h 0 is determinable in our model from the density profile of the films.

  3. Restoration of Aspen-Dominated Ecosystem in the Lake States

    Treesearch

    Douglas M. Stone; John D. Elioff; Donald V. Potter; Donald B. Peterson; Robert Wagner

    2001-01-01

    A reserve tree method(RTM)of harvesting was installed in six 70-to75-year-old aspen-dominated stands to determine if retaining 10 to 15 dominant aspen per acre would decrease sucker density to facilitate restoration of a conifer component. A reserve shelterwood cut was applied to three additional stands to evaluate performance of white pine planted under 50% crown...

  4. Restoration of aspen-dominated ecosystems in the Lake States

    Treesearch

    Douglas M. Stone; John D. Elioff; Donald V. Potter; Donald B. Peterson; Robert Wagner

    2000-01-01

    A reserve tree method (RTM) of harvesting was installed in six 70-to75-year old aspen-dominated stands to determine if retaining 10 to 15 dominant aspen per acre would decrease sucker density to facilitate restoration of a conifer component. A reserve shelterwood cut was applied to three additional stands to evaluate performance of white pine planted under 50% crown...

  5. Restoration of aspen-dominated ecosystems in the Lake States

    Treesearch

    Douglas M. Stone; John D. Elioff; Donald V. Potter; Donald B. Peterson; Robert Wagner

    2001-01-01

    A reserve tree method (RTM) of harvesting was installed in six 70- to 75-yearold aspen-dominated stands to determine if retaining 10 to 15 dominant aspen per acre would decrease sucker density to facilitate restoration of a conifer component. A reserve shelterwood cut was applied to three additional stands to evaluate performance of white pine planted under 50% crown...

  6. Forest floor fuels in red and jack pine stands

    Treesearch

    James K. Brown

    1966-01-01

    An investigation to determine the quantity and density of forest floor fuels in red pine (Pinus resinosa Ait.) and jack pine (Pinus banksiana Lamb.) stands was conducted on National Forests in Michigan and Minnesota. The study was designed to answer three questions: How much fuel per acre exits in individual layers of the forest floor? How reliably can weight of...

  7. Design Difficulties in Stand Density Studies

    Treesearch

    Frank A. Bennett

    1969-01-01

    Designing unbiased stand density studies is difficult. An acceptable sample requires stratification of the plots of age, site, and density. When basal area, percent stocking, or Reineke's stand density index is used as the density measure, this stratification forces a high negative correlation between site and number of trees per acre. Mortality in trees per acre...

  8. Reineke's Stand Density Index: Where are we and where do we go from here?

    Treesearch

    John D. Shaw

    2006-01-01

    In recent years there has been renewed interest in Reineke's Stand Density Index (SDI). Although originally described as a measurement of relative density in single-species, even-aged stands, it has since been generalized for use in uneven-aged stands and its use in multi-species stands is an active area of investigation. Some investigators use a strict definition...

  9. Carbon balance and productivity of Lemna gibba, a candidate plant for CELSS

    NASA Technical Reports Server (NTRS)

    Gale, J.; Smernoff, D. T.; Macler, B. A.; Macelroy, R. D.

    1989-01-01

    The photosynthesis and productivity of Lemna gibba is analyzed for CELSS based plant growth. Net photosynthesis of Lemna gibba is determined as a function of incident photosynthetic photon flux (PPF), with the light coming from above, below, or from both directions. Light from below is about 75 percent as effective as from above when the stand is sparse, but much less so with dense stands. High rates of photosynthesis are measured at 750 micromol / sq m per sec PPF and 1500 micromol/ mol CO2 at densities up to 660 g fresh weight (FW)/ sq m with young cultures. The analysis includes diagrams illustrating the net photosynthesis response to bilateral lighting of a sparse stand of low assimilate Lemna gibba; the effect of stand density on the net photosynthesis response to bilateral lighting of high assimilate Lemna gibba; the net photosynthesis response to ambient CO2 of sparse stands of Lemna gibba; and the time course of net photosynthesis and respiration per unit chamber and per unit dry weight of Lemna gibba.

  10. Density, ages, and growth rates in old-growth and young-growth forests in coastal Oregon

    USGS Publications Warehouse

    Tappeiner, J. C.; Huffman, D.; Spies, T.; Bailey, John D.

    1997-01-01

    We studied the ages and diameter growth rates of trees in former Douglas-fir (Pseudotsuga menziesii (Mirb.)Franco) old-growth stands on 10 sites and compared them with young-growth stands (50-70 years old, regenerated after timber harvest) in the Coast Range of western Oregon. The diameters and diameter growth rates for the first 100 years of trees in the old-growth stands were significantly greater than those in the young-growth stands. Growth rates in the old stands were comparable with those from long-term studies of young stands in which density is about 100-120 trees/ha; often young-growth stand density is well over 500 trees/ha. Ages of large trees in the old stands ranged from 100 to 420 years; ages in young stands varied by only about 5 to 10 years. Apparently, regeneration of old-growth stands on these sites occurred over a prolonged period, and trees grew at low density with little self-thinning; in contrast, after timber harvest, young stands may develop with high density of trees with similar ages and considerable self-thinning. The results suggest that thinning may be needed in dense young stands where the management objective is to speed development of old-growth characteristics.

  11. Seasonal variation in surface fuel moisture between unthinned and thinned mixed conifer forest, northern California, USA

    Treesearch

    Becky L. Estes; Eric E. Knapp; Carl N. Skinner; Fabian C. C. Uzoh

    2012-01-01

    Reducing stand density is often used as a tool for mitigating the risk of high-intensity crown fires. However, concern has been expressed that opening stands might lead to greater drying of surface fuels, contributing to increased fire risk. The objective of this study was to determine whether woody fuel moisture differed between unthinned and thinned mixed-conifer...

  12. Is self-thinning in ponderosa pine ruled by Dendroctonus bark beetles?

    Treesearch

    William W. Oliver

    1995-01-01

    Stand density of even-aged stands of ponderosa pine in California seems to be ruled by Dendroctonus bark beetles, rather than the suppressioninduced mortality common for other tree species. Size-density trajectories were plotted for 155 permanent plots in both plantations and natural stands. Bark beetle kills created a limiting Stand Density Index of...

  13. Detectability of landscape effects on recolonization increases with regional population density

    PubMed Central

    Liman, Anna-Sara; Dalin, Peter; Björkman, Christer

    2015-01-01

    Variation in population size over time can influence our ability to identify landscape-moderated differences in community assembly. To date, however, most studies at the landscape scale only cover snapshots in time, thereby overlooking the temporal dynamics of populations and communities. In this paper, we present data that illustrate how temporal variation in population density at a regional scale can influence landscape-moderated variation in recolonization and population buildup in disturbed habitat patches. Four common insect species, two omnivores and two herbivores, were monitored over 8 years in 10 willow short-rotation coppice bio-energy stands with a four-year disturbance regime (coppice cycle). The population densities in these regularly disturbed stands were compared to densities in 17 undisturbed natural Salix cinerea (grey willow) stands in the same region. A time series approach was used, utilizing the natural variation between years to statistically model recolonization as a function of landscape composition under two different levels of regional density. Landscape composition, i.e. relative amount of forest vs. open agricultural habitats, largely determined the density of re-colonizing populations following willow coppicing in three of the four species. However, the impact of landscape composition was not detectable in years with low regional density. Our results illustrate that landscape-moderated recolonization can change over time and that considering the temporal dynamics of populations may be crucial when designing and evaluating studies at landscape level. PMID:26257881

  14. Detectability of landscape effects on recolonization increases with regional population density.

    PubMed

    Liman, Anna-Sara; Dalin, Peter; Björkman, Christer

    2015-07-01

    Variation in population size over time can influence our ability to identify landscape-moderated differences in community assembly. To date, however, most studies at the landscape scale only cover snapshots in time, thereby overlooking the temporal dynamics of populations and communities. In this paper, we present data that illustrate how temporal variation in population density at a regional scale can influence landscape-moderated variation in recolonization and population buildup in disturbed habitat patches. Four common insect species, two omnivores and two herbivores, were monitored over 8 years in 10 willow short-rotation coppice bio-energy stands with a four-year disturbance regime (coppice cycle). The population densities in these regularly disturbed stands were compared to densities in 17 undisturbed natural Salix cinerea (grey willow) stands in the same region. A time series approach was used, utilizing the natural variation between years to statistically model recolonization as a function of landscape composition under two different levels of regional density. Landscape composition, i.e. relative amount of forest vs. open agricultural habitats, largely determined the density of re-colonizing populations following willow coppicing in three of the four species. However, the impact of landscape composition was not detectable in years with low regional density. Our results illustrate that landscape-moderated recolonization can change over time and that considering the temporal dynamics of populations may be crucial when designing and evaluating studies at landscape level.

  15. Effects of wildlife forestry on abundance of breeding birds in bottomland hardwood forests of Louisiana

    USGS Publications Warehouse

    Norris, Jennifer L.; Chamberlain, Michael J.; Twedt, Daniel J.

    2009-01-01

    Effects of silvicultural activities on birds are of increasing interest because of documented national declines in breeding bird populations for some species and the potential that these declines are in part due to changes in forest habitat. Silviculturally induced disturbances have been advocated as a means to achieve suitable forest conditions for priority wildlife species in bottomland hardwood forests. We evaluated how silvicultural activities on conservation lands in bottomland hardwood forests of Louisiana, USA, influenced species-specific densities of breeding birds. Our data were from independent studies, which used standardized point-count surveys for breeding birds in 124 bottomland hardwood forest stands on 12 management areas. We used Program DISTANCE 5.0, Release 2.0 (Thomas et al. 2006) to estimate density for 43 species with > 50 detections. For 36 of those species we compared density estimates among harvest regimes (individual selection, group selection, extensive harvest, and no harvest). We observed 10 species with similar densities in those harvest regimes compared with densities in stands not harvested. However, we observed 10 species that were negatively impacted by harvest with greater densities in stands not harvested, 9 species with greater densities in individual selection stands, 4 species with greater densities in group selection stands, and 4 species with greater densities in stands receiving an extensive harvest (e.g., > 40% canopy removal). Differences in intensity of harvest influenced densities of breeding birds. Moreover, community-wide avian conservation values of stands subjected to individual and group selection, and stands not harvested, were similar to each other and greater than that of stands subjected to extensive harvest that removed > 40% canopy cover. These results have implications for managers estimating breeding bird populations, in addition to predicting changes in bird communities as a result of prescribed and future forest management practices.

  16. Modeling of SAR returns from a red pine stand

    NASA Technical Reports Server (NTRS)

    Lang, R. H.; Kilic, O.; Chauhan, N. S.; Ranson, J.

    1992-01-01

    Bright P-band radar returns from red pine forests have been observed on synthetic aperture radar (SAR) images in Bangor, Maine. A plot of red pine trees was selected for the characterization and modeling to understand the cause of the high P-band returns. The red pine stand under study consisted of mature trees. Diameter at breast height (DBH) measurements were made to determine stand density as a function of tree diameter. Soil moisture and bulk density measurements were taken along with ground rough surface profiles. Detailed biomass measurements of the needles, shoots, branches, and trunks were also taken. These site statistics have been used in a distorted Born approximation model of the forest. Computations indicate that the direct-reflected or the double-bounce contributions from the ground are responsible for the high observed P-band returns for HH polarization.

  17. Growth and yield considerations and implications for alternative density management objectives and approaches

    Treesearch

    David Marshall

    2013-01-01

    Density management through thinning is the most important tool foresters have to aff ect stand development and stand structure of existing stands. Reducing stand density by thinning increases the growing space and resource availability (e.g., light, water, and nutrients) for the remaining trees. Th is can result in increased average tree growth. More available site...

  18. The Influence of Residual Stand Densities on Regeneration in Sugar Maple Stands

    Treesearch

    Carl H. Tubbs

    1968-01-01

    Studies of regeneration 2, 5, and 10 years after cutting mature and overmature sugar maple stands to several residual densities show that (1) sugar maple is still the predominant species under all stand densities (2) nearly all regeneration reaching larger size classes became established before cutting (3) heavier cuttings (30, 50, and 70 square feet) are more rapidly...

  19. Preliminary thinning guidelines using stand density index for the maintenance of uneven-aged pinyon-juniper ecosystems

    Treesearch

    Douglas H. Page

    2008-01-01

    This paper demonstrates how Stand Density Index may be used to guide postthinning stand structure for the sustainable management of pinyon-juniper ecosystems. The post-thinning residual stand density can be varied to achieve various management objectives. Uneven-aged management is recommended, where possible, as a better approximation of the natural development process...

  20. Seasonal and cumulative loblolly pine development under two stand density and fertility levels through four growing seasons

    Treesearch

    James D. Haywood

    1994-01-01

    A loblolly pine stand was subjected to two cultural treatments to determine treatment effects in the 9th through 12th growing seasons. Thining resulted in less spring height growth in the 9th and 10th growing seasons than no thinning, but thinning resulted in more diameter growth each year. Fertilization increased height and diameter growth beginning in the 10th...

  1. Stand-density study of spruce-hemlock stands in southeastern Alaska.

    Treesearch

    Donald J. DeMars

    2000-01-01

    The lack of growth and yield information for young even-aged western hemlock (Tsuga heterophylla(Raf.) Sarg.)-Sitka spruce (Picea sitchensis (Bong.) Carr.) stands in southeastern Alaska served as the impetus for a long-term stand-density study begun in 1974. The study has followed permanent growth plots in managed stands under...

  2. Influence of stand density on soil CO2 efflux for a Pinus densiflora forest in Korea.

    PubMed

    Noh, Nam Jin; Son, Yowhan; Lee, Sue Kyoung; Yoon, Tae Kyung; Seo, Kyung Won; Kim, Choonsig; Lee, Woo-Kyun; Bae, Sang Won; Hwang, Jaehong

    2010-07-01

    We investigated the influence of stand density [938 tree ha(-1) for high stand density (HD), 600 tree ha(-1) for medium stand density (MD), and 375 tree ha(-1) for low stand density (LD)] on soil CO(2) efflux (R (S)) in a 70-year-old natural Pinus densiflora S. et Z. forest in central Korea. Concurrent with R (S) measurements, we measured litterfall, total belowground carbon allocation (TBCA), leaf area index (LAI), soil temperature (ST), soil water content (SWC), and soil nitrogen (N) concentration over a 2-year period. The R (S) (t C ha(-1) year(-1)) and leaf litterfall (t C ha(-1) year(-1)) values varied with stand density: 6.21 and 2.03 for HD, 7.45 and 2.37 for MD, and 6.96 and 2.23 for LD, respectively. In addition, R (S) was correlated with ST (R (2) = 0.77-0.80, P < 0.001) and SWC (R (2) = 0.31-0.35, P < 0.001). It appeared that stand density influenced R (S) via changes in leaf litterfall, LAI and SWC. Leaf litterfall (R (2) = 0.71), TBCA (R (2) = 0.64-0.87), and total soil N contents in 2007 (R (2) = 0.94) explained a significant amount of the variance in R (S) (P < 0.01). The current study showed that stand density is one of the key factors influencing R (S) due to the changing biophysical and environmental factors in P. densiflora.

  3. Changes in very fine root respiration and morphology with time since last fire in a boreal forest

    NASA Astrophysics Data System (ADS)

    Makita, Naoki; Pumpanen, Jukka; Köster, Kajar; Berninger, Frank

    2016-04-01

    We examined the physiological and morphological responses of individual fine root segments in boreal forests stands with different age since the last fire to determine changes in specific fine root respiration and morphological traits during forest succession. We investigated the respiration of fine roots divided into three diameter classes (<0.5, 0.5-1.0, and 1.0-2.0 mm) in a Finnish boreal Pinus sylvestris L. in forest stands with 5, 45, 63, and 155 years since the last fire. Specific respiration rates of <0.5 mm roots in 155-year-old stands were 74%, 38%, and 31% higher than in 5-, 45-, and 63-year-old stands, respectively. However, the respiration rates of thicker diameter roots did not significantly change among stands with respect to time after fire. Similarly, fire disturbance had a strong impact on morphological traits of <0.5 mm roots, but not on thicker roots. Root respiration rates correlated positively with specific root length (length per unit mass) and negatively with root tissue density (mass per unit volume) in all stand ages. The linear regression lines fitted to the relationships between root respiration and specific root length or root tissue density showed significantly higher intercepts in 63- and 155-year-old than in 5-year-old stands. Significant shifts in the intercept of the common slope of respiration vs. morphology indicate the different magnitude of the changes in physiological performance among the fire age class. Despite a specific small geographic area, we suggest that the recovery of boreal forests following wildfire induces a strategy that favors carbon investment in nutrient and water exploitation efficiency with consequences for higher respiration, length, and lower tissue density of very fine roots.

  4. A stand density management diagram for sawtimber-sized mixed upland central hardwoods

    Treesearch

    J.A., Jr. Kershaw; B.C. Fischer

    1991-01-01

    Data from 190 CFI plots located in southern and west-central Indiana are used to develop a stand density diagram for sawtimber-sized mixed upland hardwoods in the Central States. The stand density diagram utilizes the concepts of self-thinning to establish a maximum size-density curve, and the stocking standards of Gingrich (1967) to formulate imtermediate stocking...

  5. Growth of lodgepole pine thinned to various densities on two sites with differing productivities in central Oregon.

    Treesearch

    P.H. Cochran; Walter G. Dahms

    2000-01-01

    Concave curvilinear decreases in diameter growth occurred with increasing stand densities. A convex curvilinear increase in gross growth of basal area and total cubic volume took place with increasing stand density. Maximum cumulative net cubic (total and merchantable) and board-foot yields varied curvilinearly with stand density. These yields peaked at intermediate...

  6. Effect of diameter limits and stand structure on relative density indices: a case study

    Treesearch

    Robert O. Curtis

    2010-01-01

    An understory of shade-tolerant species often develops in stands in the Douglas-fir region of western Washington and Oregon and can have a disproportionate effect on relative density indices, such as Reineke stand density index and Curtis relative density. The effects of such understories and of other departures from The even-aged condition are illustrated with...

  7. Differences between standing and downed dead tree wood density reduction factors: A comparison across decay classes and tree species

    Treesearch

    Mark E. Harmon; Christopher W. Woodall; Becky Fasth; Jay Sexton; Misha Yatkov

    2011-01-01

    Woody detritus or dead wood is an important part of forest ecosystems and has become a routine facet of forest monitoring and inventory. Biomass and carbon estimates of dead wood depend on knowledge of species- and decay class-specifi c density or density reduction factors. While some progress has been made in determining these parameters for dead and downed trees (DD...

  8. Drivers of aboveground wood production in a lowland tropical forest of West Africa: teasing apart the roles of tree density, tree diversity, soil phosphorus, and historical logging.

    PubMed

    Jucker, Tommaso; Sanchez, Aida Cuni; Lindsell, Jeremy A; Allen, Harriet D; Amable, Gabriel S; Coomes, David A

    2016-06-01

    Tropical forests currently play a key role in regulating the terrestrial carbon cycle and abating climate change by storing carbon in wood. However, there remains considerable uncertainty as to whether tropical forests will continue to act as carbon sinks in the face of increased pressure from expanding human activities. Consequently, understanding what drives productivity in tropical forests is critical. We used permanent forest plot data from the Gola Rainforest National Park (Sierra Leone) - one of the largest tracts of intact tropical moist forest in West Africa - to explore how (1) stand basal area and tree diversity, (2) past disturbance associated with past logging, and (3) underlying soil nutrient gradients interact to determine rates of aboveground wood production (AWP). We started by statistically modeling the diameter growth of individual trees and used these models to estimate AWP for 142 permanent forest plots. We then used structural equation modeling to explore the direct and indirect pathways which shape rates of AWP. Across the plot network, stand basal area emerged as the strongest determinant of AWP, with densely packed stands exhibiting the fastest rates of AWP. In addition to stand packing density, both tree diversity and soil phosphorus content were also positively related to productivity. By contrast, historical logging activities negatively impacted AWP through the removal of large trees, which contributed disproportionately to productivity. Understanding what determines variation in wood production across tropical forest landscapes requires accounting for multiple interacting drivers - with stand structure, tree diversity, and soil nutrients all playing a key role. Importantly, our results also indicate that logging activities can have a long-lasting impact on a forest's ability to sequester and store carbon, emphasizing the importance of safeguarding old-growth tropical forests.

  9. Effect of plantation density on kraft pulp production from red pine (Pinus resinosa Ait.)

    Treesearch

    J.Y. Zhu; G.C. Myers

    2006-01-01

    Red pine (Pinus resinosa Ait.) butt logs from 38 year old research plots were used to study the effect of plantation stand density on kraft pulp production. Results indicate that plantation stand density can affect pulp yield, unrefined pulp mean fibre length, and the response of pulp fibre length to pulp refining. However, the effect of plantation stand density on...

  10. Leaf dynamics in growth and reproduction of Xanthium canadense as influenced by stand density

    PubMed Central

    Ogawa, Takahiro; Oikawa, Shimpei; Hirose, Tadaki

    2015-01-01

    Background and Aims Leaf longevity is controlled by the light gradient in the canopy and also by the nitrogen (N) sink strength in the plant. Stand density may influence leaf dynamics through its effects on light gradient and on plant growth and reproduction. This study tests the hypothesis that the control by the light gradient is manifested more in the vegetative period, whereas the opposite is true when the plant becomes reproductive and develops a strong N sink. Methods Stands of Xanthium canadense were established at two densities. Emergence, growth and death of every leaf on the main stem and branches, and plant growth and N uptake were determined from germination to full senescence. Mean residence time and dry mass productivity were calculated per leaf number, leaf area, leaf mass and leaf N (collectively termed ‘leaf variables’) in order to analyse leaf dynamics and its effect on plant growth. Key Results Branching and reproductive activities were higher at low than at high density. Overall there was no significant difference in mean residence time of leaf variables between the two stands. However, early leaf cohorts on the main stem had a longer retention time at low density, whereas later cohorts had a longer retention time at high density. Branch leaves emerged earlier and tended to live longer at low than at high density. Leaf efficiencies, defined as carbon export per unit investment of leaf variables, were higher at low density in all leaf variables except for leaf number. Conclusions In the vegetative phase of plant growth, the light gradient strongly controls leaf longevity, whereas later the effects of branching and reproductive activities become stronger and over-rule the effect of light environment. As leaf N supports photosynthesis and also works as an N source for plant development, N use is pivotal in linking leaf dynamics with plant growth and reproduction. PMID:26248476

  11. Longleaf pine cone production in relation to site index, stand age, and stand density

    Treesearch

    Thomas Croker

    1973-01-01

    Few cones were produced in stands less than 30 years old. In stands 30 to 70 years in age, production seemed best at timber densities of about 30 square feet of basal area per acre, and tended to increase with increasing site index.

  12. Maximum growth potential in loblolly pine: results from a 47-year-old spacing study in Hawaii

    Treesearch

    Lisa J. Samuelson; Thomas L. Eberhardt; John R. Butnor; Tom A. Stokes; Kurt H. Johnsen

    2010-01-01

    Growth, allocation to woody root biomass, wood properties, leaf physiology, and shoot morphology were examined in a 47-year-old loblolly pine (Pinus taeda L.) density trial located in Maui, Hawaii, to determine if stands continued to carry the high density, basal area, and volume reported at younger ages and to identify potential factors controlling...

  13. Individual tree growth response to variable-density thinning in coastal Pacific Northwest forests.

    Treesearch

    Scott D.s Roberts; Constance A. Harrington

    2008-01-01

    We examined 5-year basal area growth of nearly 2600 trees in stem-mapped plots at five locations differing in site characteristics, species composition, and management history on the Olympic Peninsula in Western Washington, USA. Our objectives were to determine if internal edges, the boundaries within the stand between components of the variable-density thinning,...

  14. Monitoring forest land from high altitude and from space

    NASA Technical Reports Server (NTRS)

    1971-01-01

    Forest inventory, forest stress, and standardization and calibration studies are presented. These include microscale photointerpretation of forest and nonforest land classes, multiseasonal film densities for automated forest and nonforest land classification, trend and spread of bark beetle infestations from 1968 through 1971, aerial photography for determining optimum levels of stand density to reduce such infestations, use of airborne spectrometers and multispectral scanners for previsual detection of Ponderosa pine trees under stress from insects and diseases, establishment of an earth resources technology satellite test site in the Black Hills and the identification of natural resolution targets, detection of root disease impact on forest stands by sequential orbital and suborbital multispectral photography, and calibration of color aerial photography.

  15. Long-Term Trends In Loblolly Pine Productivity And Stand Characteristics In Response To Stand Density And Fertilization In The Western Gulf Region

    Treesearch

    M.A. Sword; J. L. Chambers; Z. Tang; T. J. Dean; J. C. Goelz

    2002-01-01

    Two levels each of fertilization and stand density were established to create four environments in a 7-year-old loblolly pine plantation on a N and P deficient western Gulf Coastal Plain site in Louisiana. Levels of fertilization were no fertilization and application of 120 lb N and 134 lb P/ac. Levels of stand density were the original stocking (1,210 trees/ac), and...

  16. Vertical ground reaction force during standing and walking: Are they related to bone mineral density left-right asymmetries?

    PubMed

    Brozgol, Marina; Arbiv, Mira; Mirelman, Anat; Herman, Talia; Hausdorff, Jeffrey M; Vaisman, Nachum

    2017-05-01

    Osteoporosis is a systemic skeletal disease that is characterized by reduced bone mass, deterioration of bone tissue and skeletal fragility. The purpose of the current study was to determine whether asymmetrical femur bone mineral density (BMD) is associated with asymmetrical gait and standing. We compared measures of gait and standing asymmetry in subjects with (n=38) and without (n=11) significant left-right differences in BMD. Participants walked for 72m at their comfortable speed and stood quietly for 60s while outfitted with pressure-sensitive insoles. Based on the pressure measurements, indices of standing and gait asymmetry were determined. Gait Asymmetry (GA) indices of maximum ground reaction force (GRF) and stance time were significantly higher in the asymmetrical BMD group, compared to the symmetrical group (p<0.03). During quiet standing, maximal GRF was twice as high in those with BMD asymmetry, compared to those without, although this difference was not statistically significant (p=0.10). These preliminary findings indicate that femur BMD asymmetry and gait asymmetry are interrelated in otherwise healthy adults. Nutrition, metabolism and lifestyle are known contributors to BMD; typically, they affect bone health symmetrically. We suggest, therefore, that the BMD asymmetry may be due to previous changes in the loading pattern during walking that might have led to asymmetric bone deterioration. Future larger scale and prospective studies are needed to identify the mechanisms underlying the relationship between standing, gait and BMD and to explore whether gait training and exercises that target gait symmetry might help to reduce BMD asymmetry. Copyright © 2017 Elsevier B.V. All rights reserved.

  17. Effects of two-age management and clearcutting on songbird density and reproductive success

    Treesearch

    Jeffery V. Nichols; Petra Bohall Wood

    1995-01-01

    We examined density and reproductive success of passerine species on 7 uncut forest stands and on 12 stands harvested 10-14 years ago on the Monongahela National Forest of West Virginia (6 clearcut stands and 6 stands harvested using 2-age management). In 2-age management, stands resemble a shelterwood cut with 10-30 overstory trees/acre left uncut. Uncut periphery...

  18. Relationships between the attributes of standing live and dead down trees in forests of the Lake States

    Treesearch

    Christopher W. Woodall; Linda Nagel

    2007-01-01

    Refining the understanding of the relationship between a stand's standing live and down dead trees in terms of size, density, and biomass attributes may aid efforts to predict fuel loadings based on standing tree attributes. Therefore, the objective of this study was to compare down dead and standing live tree attributes (size, density, and biomass) for inventory...

  19. Stand quality management in a late-rotation, red oak-sweetgum stand in east Mississippi: preliminary results following thinning

    Treesearch

    James S. Meadows; Daniel A. Skojac

    2012-01-01

    Stand quality management is a new management strategy in which thinning prescriptions are based solely on tree quality rather than a quantitative level of residual stand density. As long as residual density falls within fairly broad limits, prescriptions are based on tree quality alone. We applied four thinning prescriptions based on stand quality management, along...

  20. Stand density, tree social status and water stress influence allocation in height and diameter growth of Quercus petraea (Liebl.).

    PubMed

    Trouvé, Raphaël; Bontemps, Jean-Daniel; Seynave, Ingrid; Collet, Catherine; Lebourgeois, François

    2015-10-01

    Even-aged forest stands are competitive communities where competition for light gives advantages to tall individuals, thereby inducing a race for height. These same individuals must however balance this competitive advantage with height-related mechanical and hydraulic risks. These phenomena may induce variations in height-diameter growth relationships, with primary dependences on stand density and tree social status as proxies for competition pressure and access to light, and on availability of local environmental resources, including water. We aimed to investigate the effects of stand density, tree social status and water stress on the individual height-circumference growth allocation (Δh-Δc), in even-aged stands of Quercus petraea Liebl. (sessile oak). Within-stand Δc was used as surrogate for tree social status. We used an original long-term experimental plot network, set up in the species production area in France, and designed to explore stand dynamics on a maximum density gradient. Growth allocation was modelled statistically by relating the shape of the Δh-Δc relationship to stand density, stand age and water deficit. The shape of the Δh-Δc relationship shifted from linear with a moderate slope in open-grown stands to concave saturating with an initial steep slope in closed stands. Maximum height growth was found to follow a typical mono-modal response to stand age. In open-grown stands, increasing summer soil water deficit was found to decrease height growth relative to radial growth, suggesting hydraulic constraints on height growth. A similar pattern was found in closed stands, the magnitude of the effect however lowering from suppressed to dominant trees. We highlight the high phenotypic plasticity of growth in sessile oak trees that further adapt their allocation scheme to their environment. Stand density and tree social status were major drivers of growth allocation variations, while water stress had a detrimental effect on height in the Δh-Δc allocation. © The Author 2015. Published by Oxford University Press. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

  1. Sap flux density and stomatal conductance of European beech and common oak trees in pure and mixed stands during the summer drought of 2003

    NASA Astrophysics Data System (ADS)

    Jonard, F.; André, F.; Ponette, Q.; Vincke, C.; Jonard, M.

    2011-10-01

    SummarySap flux density of European beech and common oak trees was determined from sap flow measurements in pure and mixed stands during the summer drought of 2003. Eight trees per species and per stand were equipped with sap flow sensors. Soil water content was monitored in each stand at different depths by using time-domain reflectometry (TDR). Leaf area index and vertical root distribution were also investigated during the growing season. From sap flux density ( SFD) data, mean stomatal conductance of individual trees ( G s) was calculated by inverting the Penman-Monteith equation. Linear mixed models were developed to analyse the effects of species and stand type (pure vs. mixed) on SFD and G s and on their sensitivity to environmental variables (vapour pressure deficit ( D), incoming solar radiation ( R G), and relative extractable water ( REW)). For reference environmental conditions, we did not find any tree species or stand type effects on SFD. The sensitivity of SFD to D was higher for oak than for beech in the pure stands ( P < 0.0001) but the mixing of species reduced it for oak and increased it for beech, so that the sensitivity of SFD to D became higher for beech than for oak in the mixed stand ( P < 0.0001). At reference conditions, G s was significantly higher for beech compared to oak (2.1 and 1.8 times in the pure and mixed stand, respectively). This was explained by a larger beech sapwood-to-leaf area ratio compared to oak. The sensitivity of G s to REW was higher for beech than for oak and was ascribed to a higher vulnerability of beech to air embolism and to a more sensitive stomatal regulation. The sensitivity of beech G s to REW was lower in the mixed than in the pure stand, which could be explained by a better sharing of the resources in the mixture, by facilitation processes (hydraulic lift), and by a rainfall partitioning in favour of beech.

  2. [Carbon density distribution characteristics and influencing factors in aerially seeded Pinus massoniana plantations].

    PubMed

    Pan, Ping; Han, Tian Yi; OuYang, Xun Zhi; Liu, Yuan Qiu; Zang, Hao; Ning, Jin Kui; Yang, Yang

    2017-12-01

    The distribution characteristics of carbon density under aerially seeded Pinus massoniana plantations in Ganzhou City of Jiangxi Province were studied. Total 15 factors, including site, stand, understory vegetation, litter and so on were selected to establish a relationship model between stand carbon density and influencing factors, and the main influencing factors were also screened. The results showed that the average carbon density was 98.29 t·hm -2 at stand level with soil layer (49.58 t·hm -2 ) > tree layer (45.25 t·hm -2 ) > understory vegetation layer (2.23 t·hm -2 ) > litter layer (1.23 t·hm -2 ). Significantly positive correlations were found among the tree, litter and soil layers, but not among the other layers. The main factors were tree density, avera-ge diameter at breast height (DBH), soil thickness, slope position, stand age and canopy density to affect carbon density in aerially seeded P. massoniana plantations. The partial correlation coefficients of the six main factors ranged from 0.331 to 0.434 with significance by t test. The multiple correlation coefficient of quantitative model I reached 0.796 with significance by F test (F=9.28). For stand density, the best tree density and canopy density were 1500-2100 plants·hm -2 and 0.4-0.7, respectively. The moderate density was helpful to improve ecosystem carbon sequestration. The carbon density increased with increasing stand age, DBH and soil thickness, and was higher in lower than middle and upper slope positions.

  3. Structural Characteristics of an Old-Growth Coast Redwood Stand in Mendocino County, California

    Treesearch

    Gregory A. Giusti

    2007-01-01

    This paper compares stand characteristics of Old Growth coastal redwood stand densities and forest structure found throughout the northern tier of the range of coast redwood (Sequoia sempervirens). Tree densities are relatively low compared to commercially managed stands of coast redwood. Tree size classes distributions vary from 254cm...

  4. Are self-thinning constraints needed in a tree-specific mortality model?

    Treesearch

    Robert A. Monserud; Thomas Ledermann; Hubert Sterba

    2005-01-01

    Can a tree-specific mortality model elicit expected forest stand density dynamics without imposing stand-level constraints such as Reineke's maximum stand density index (SDImax) or the -3/2 power law of self-thinning? We examine this emergent properties question using the Austrian stand simulator PROGNAUS. This simulator was chosen...

  5. Stand density relationships

    Treesearch

    John C. Tappeiner

    2013-01-01

    Th inning stands (managing their densities) aff ects the development of trees and understory plants as individuals, as well as stand-level characteristics like structure, microclimate, and stand growth, habitat for various species, and fuel and potential fi re severity. Th ese characteristics and the rate of changes are aff ected by thinning severity—the reduction in...

  6. Relationship between built environment attributes and physical function in Japanese community-dwelling older adults.

    PubMed

    Soma, Yuki; Tsunoda, Kenji; Kitano, Naruki; Jindo, Takashi; Tsuji, Taishi; Saghazadeh, Mahshid; Okura, Tomohiro

    2017-03-01

    To explore the relationships between the built environment and older adults' physical function. The present cross-sectional study carried out in 2010-2012 used data drawn from 509 community-dwelling older adults aged 65-86 years living in Kasama City, a Japanese rural region. We evaluated physical function with the following performance tests: grip strength, sit-to-stand, timed up & go and walking speed. Using geographic information systems, we measured population density and the number of destinations related to daily life, community centers, medical facilities and recreational facilities within participants' neighborhoods. After adjusting for potential confounders, we found lower population density was related to poor performance of sit-to-stand and walking speed in both sexes, and grip strength in women (trend P < 0.05). A lower number of daily life-related destinations was related to poor performance of sit-to-stand and walking speed in men, and grip strength and sit-to-stand in women. Similarly, the number of community centers was related to walking speed in both sexes. The number of medical and recreational facilities was also related to some physical performance in both sexes. A lower land use mix score, calculated by principal component analysis, was related to lower performance of sit-to-stand and walking speed in men, and grip strength and sit-to-stand in women. The present study suggests that, although there are some sex differences, low population density, land use mix, and fewer daily life-related destinations, community centers, medical facilities and recreational facilities are negative determinants of physical function. Geriatr Gerontol Int 2017; 17: 382-390. © 2016 Japan Geriatrics Society.

  7. Gross Basal Area Growth of Northern White-Cedar is Independent of Stand Density Over A Wide Range

    Treesearch

    Bruce W. Foltz; William F. Johnston

    1968-01-01

    A 20-year study in Wisconsin swamp stand of northern white-cedar indicates that after a second thinning gross basal area growth is independent of stand densities ranging from 90 to 225 square feet of basal area per acre. Mortality, net growth, and ingrowth were also independent of density in the thinned plots.

  8. Effects of stand density on top height estimation for ponderosa pine

    Treesearch

    Martin Ritchie; Jianwei Zhang; Todd Hamilton

    2012-01-01

    Site index, estimated as a function of dominant-tree height and age, is often used as an expression of site quality. This expression is assumed to be effectively independent of stand density. Observation of dominant height at two different ponderosa pine levels-of-growing-stock studies revealed that top height stability with respect to stand density depends on the...

  9. Growth of ponderosa pine thinned to different stocking levels in central Oregon: 30-year results.

    Treesearch

    P.H. Cochran; James W. Barrett

    1999-01-01

    Periodic annual increments (PAI) for survivor diameters decreased curvilinearly with increasing stand density. Gross volume and basal areas PAIs increased linearly with increasing stand density. Growth of basal area and volume for the 20 largest trees per acre were reduced curvilinearly with increasing stand density. Bark beetles were the primary cause of mortality. No...

  10. Effects of age and stand density of mother trees on early Pinus thunbergii seedling establishment in the coastal zone, China.

    PubMed

    Mao, Peili; Han, Guangxuan; Wang, Guangmei; Yu, Junbao; Shao, Hongbo

    2014-01-01

    Effects of age and stand density of mother tree on seed germination, seedling biomass allocation, and seedling growth of Pinus thunbergii were studied. The results showed that age of mother tree did not have significant influences on seed germination, but it was significant on seedling biomass allocation and growth. Seedlings from the minimum and maximum age of mother tree had higher leaf mass ratio and lower root mass ratio than from the middle age of mother tree. Moreover, they also had higher relative height growth rate and slenderness, which were related to their biomass allocation. Stand density of mother tree mainly demonstrated significant effects on seed germination and seedling growth. Seed from higher stand density of mother tree did not decrease germination rate, but had higher mean germination time, indicating that it delayed germination process. Seedlings of higher stand density of mother tree showed higher relative height growth rate and slenderness. These traits of offspring from higher stand density of mother tree were similar to its mother, indicating significant environmental maternal effects. So, mother tree identity of maternal age and environments had important effects on natural regeneration of the coastal P. thunbergii forest.

  11. Effects of Age and Stand Density of Mother Trees on Early Pinus thunbergii Seedling Establishment in the Coastal Zone, China

    PubMed Central

    Mao, Peili; Han, Guangxuan; Wang, Guangmei; Yu, Junbao; Shao, Hongbo

    2014-01-01

    Effects of age and stand density of mother tree on seed germination, seedling biomass allocation, and seedling growth of Pinus thunbergii were studied. The results showed that age of mother tree did not have significant influences on seed germination, but it was significant on seedling biomass allocation and growth. Seedlings from the minimum and maximum age of mother tree had higher leaf mass ratio and lower root mass ratio than from the middle age of mother tree. Moreover, they also had higher relative height growth rate and slenderness, which were related to their biomass allocation. Stand density of mother tree mainly demonstrated significant effects on seed germination and seedling growth. Seed from higher stand density of mother tree did not decrease germination rate, but had higher mean germination time, indicating that it delayed germination process. Seedlings of higher stand density of mother tree showed higher relative height growth rate and slenderness. These traits of offspring from higher stand density of mother tree were similar to its mother, indicating significant environmental maternal effects. So, mother tree identity of maternal age and environments had important effects on natural regeneration of the coastal P. thunbergii forest. PMID:24955404

  12. The Relation of Growth to Stand Density in Natural Loblolly Pine Stands

    Treesearch

    K.F. Wenger; T.C. Evans; T. Lotti; R.W. Cooper; E.V. Brender

    1958-01-01

    This is a progress report of a regional study on growing-space requirements for natural stands of loblolly pine (Pinus taeda L.).A primary objective is to measure the effects of residual stand density, obtained naturally or by cutting, during intermediate ages, upon volume yield and total production. By imposing real values and costs upon...

  13. Stocking equations for regeneration in mixed oak stands

    Treesearch

    Songlin Fei; Kim C. Steiner; James C. Finley

    2007-01-01

    Regeneration stocking equations for mixed-oak stands were developed based on data collected from nearly 14,000 plots in the central Appalachians. Maximum stand density was identified by plotting aggregate height against number of seedlings per plot, and was used as the reference level of the average maximum stand density (100 percent stocking or A-level stocking)....

  14. Are self-thinning contraints needed in a tree-specific mortality model.

    Treesearch

    Robert A. Monserud; Thomas Ledermann; Hubert Sterba

    2005-01-01

    Can a tree-specific mortality model elicit expected forest stand density dynamics without imposing stand-level constraints such as Reineke's maximum stand density index (SDI,) or the -312 power law of self-thinning? We examine this emergent properties question using the Austrian stand simulator PROGNAUS. This simulator was chosen specifically because it does not...

  15. Home Range, Habitat Selection, and Population Dynamics of Southern Flying Squirrels in Managed Forests in Arkansas

    Treesearch

    James F. Taulman; Kimberly G. Smith

    2004-01-01

    Abstract After experimental harvests on 18 mature pine-hardwood stands in 6 replicated groups, flying squirrels changed from uniform pre-harvest patterns of nest box use to concentrating on protected greenbelt (GB) areas on harvested stands. Squirrel densities declined on all harvested stands as densities increased on three control stands. Fifty...

  16. Coronal loop seismology using damping of standing kink oscillations by mode coupling

    NASA Astrophysics Data System (ADS)

    Pascoe, D. J.; Goddard, C. R.; Nisticò, G.; Anfinogentov, S.; Nakariakov, V. M.

    2016-05-01

    Context. Kink oscillations of solar coronal loops are frequently observed to be strongly damped. The damping can be explained by mode coupling on the condition that loops have a finite inhomogeneous layer between the higher density core and lower density background. The damping rate depends on the loop density contrast ratio and inhomogeneous layer width. Aims: The theoretical description for mode coupling of kink waves has been extended to include the initial Gaussian damping regime in addition to the exponential asymptotic state. Observation of these damping regimes would provide information about the structuring of the coronal loop and so provide a seismological tool. Methods: We consider three examples of standing kink oscillations observed by the Atmospheric Imaging Assembly (AIA) of the Solar Dynamics Observatory (SDO) for which the general damping profile (Gaussian and exponential regimes) can be fitted. Determining the Gaussian and exponential damping times allows us to perform seismological inversions for the loop density contrast ratio and the inhomogeneous layer width normalised to the loop radius. The layer width and loop minor radius are found separately by comparing the observed loop intensity profile with forward modelling based on our seismological results. Results: The seismological method which allows the density contrast ratio and inhomogeneous layer width to be simultaneously determined from the kink mode damping profile has been applied to observational data for the first time. This allows the internal and external Alfvén speeds to be calculated, and estimates for the magnetic field strength can be dramatically improved using the given plasma density. Conclusions: The kink mode damping rate can be used as a powerful diagnostic tool to determine the coronal loop density profile. This information can be used for further calculations such as the magnetic field strength or phase mixing rate.

  17. Legacy of Pre-Disturbance Spatial Pattern Determines Early Structural Diversity following Severe Disturbance in Montane Spruce Forests

    PubMed Central

    Bače, Radek; Svoboda, Miroslav; Janda, Pavel; Morrissey, Robert C.; Wild, Jan; Clear, Jennifer L.; Čada, Vojtěch; Donato, Daniel C.

    2015-01-01

    Background Severe canopy-removing disturbances are native to many temperate forests and radically alter stand structure, but biotic legacies (surviving elements or patterns) can lend continuity to ecosystem function after such events. Poorly understood is the degree to which the structural complexity of an old-growth forest carries over to the next stand. We asked how pre-disturbance spatial pattern acts as a legacy to influence post-disturbance stand structure, and how this legacy influences the structural diversity within the early-seral stand. Methods Two stem-mapped one-hectare forest plots in the Czech Republic experienced a severe bark beetle outbreak, thus providing before-and-after data on spatial patterns in live and dead trees, crown projections, down logs, and herb cover. Results Post-disturbance stands were dominated by an advanced regeneration layer present before the disturbance. Both major species, Norway spruce (Picea abies) and rowan (Sorbus aucuparia), were strongly self-aggregated and also clustered to former canopy trees, pre-disturbance snags, stumps and logs, suggesting positive overstory to understory neighbourhood effects. Thus, although the disturbance dramatically reduced the stand’s height profile with ~100% mortality of the canopy layer, the spatial structure of post-disturbance stands still closely reflected the pre-disturbance structure. The former upper tree layer influenced advanced regeneration through microsite and light limitation. Under formerly dense canopies, regeneration density was high but relatively homogeneous in height; while in former small gaps with greater herb cover, regeneration density was lower but with greater heterogeneity in heights. Conclusion These findings suggest that pre-disturbance spatial patterns of forests can persist through severe canopy-removing disturbance, and determine the spatial structure of the succeeding stand. Such patterns constitute a subtle but key legacy effect, promoting structural complexity in early-seral forests as well as variable successional pathways and rates. This influence suggests a continuity in spatial ecosystem structure that may well persist through multiple forest generations. PMID:26421726

  18. Stand and tree characteristics influencing density of fir engraver beetle attack scars in white fir

    Treesearch

    George T. Ferrell

    1973-01-01

    Density of embedded scars resulting from old fir engraver (Scolytus ventralis Lec.) attacks was assessed in 3,430 bole cross sections cut from 603 white firs growing in both cutover and in virgin old-growth stands in northern California. Mean scar density was directly related to percentage of mature stand removed by logging and was higher in...

  19. Impacts of four decades of stand density management treatments on wood properties of loblolly pine

    Treesearch

    M.A. Blazier; A. Clark; J.M. Mahon; M.R. Strub; R.F. Daniels; L.R. Schimleck

    2013-01-01

    Stand density management is a powerful silvicultural tool for manipulating stand volumes, but it has the potential to alter key wood properties. At a site in northcentral Louisiana, five density management regimes were conducted over a 45-year period. At age 49, a stratified sample of trees was destructively harvested for crown length, taper, and specific gravity...

  20. Stand age and fine root biomass, distribution and morphology in a Norway spruce chronosequence in southeast Norway.

    PubMed

    Børja, Isabella; De Wit, Heleen A; Steffenrem, Arne; Majdi, Hooshang

    2008-05-01

    We assessed the influence of stand age on fine root biomass and morphology of trees and understory vegetation in 10-, 30-, 60- and 120-year-old Norway spruce stands growing in sandy soil in southeast Norway. Fine root (< 1, 1-2 and 2-5 mm in diameter) biomass of trees and understory vegetation (< 2 mm in diameter) was sampled by soil coring to a depth of 60 cm. Fine root morphological characteristics, such as specific root length (SRL), root length density (RLD), root surface area (RSA), root tip number and branching frequency (per unit root length or mass), were determined based on digitized root data. Fine root biomass and morphological characteristics related to biomass (RLD and RSA) followed the same tendency with chronosequence and were significantly higher in the 30-year-old stand and lower in the 10-year-old stand than in the other stands. Among stands, mean fine root (< 2 mm) biomass ranged from 49 to 398 g m(-2), SLR from 13.4 to 19.8 m g(-1), RLD from 980 to 11,650 m m(-3) and RSA from 2.4 to 35.4 m(2) m(-3). Most fine root biomass of trees was concentrated in the upper 20 cm of the mineral soil and in the humus layer (0-5 cm) in all stands. Understory fine roots accounted for 67 and 25% of total fine root biomass in the 10- and 120-year-old stands, respectively. Stand age had no affect on root tip number or branching frequency, but both parameters changed with soil depth, with increasing number of root tips and decreasing branching frequency with increasing soil depth for root fractions < 2 mm in diameter. Specific (mass based) root tip number and branching density were highest for the finest roots (< 1 mm) in the humus layer. Season (spring or fall) had no effect on tree fine root biomass, but there was a small and significant increase in understory fine root biomass in fall relative to spring. All morphological characteristics showed strong seasonal variation, especially the finest root fraction, with consistently and significantly higher values in spring than in fall. We conclude that fine root biomass, especially in the finest fraction (< 1 mm in diameter), is strongly dependent on stand age. Among stands, carbon concentration in fine root biomass was highest in the 30-year-old stand, and appeared to be associated with the high tree and canopy density during the early stage of stand development. Values of RLD and RSA, morphological features indicative of stand nutrient-uptake efficiency, were higher in the 30-year-old stand than in the other stands.

  1. Estimating structural attributes of Douglas-fir/western hemlock forest stands from Landsat and SPOT imagery

    NASA Technical Reports Server (NTRS)

    Cohen, Warren B.; Spies, Thomas A.

    1992-01-01

    Relationships between spectral and texture variables derived from SPOT HRV 10 m panchromatic and Landsat TM 30 m multispectral data and 16 forest stand structural attributes is evaluated to determine the utility of satellite data for analysis of hemlock forests west of the Cascade Mountains crest in Oregon and Washington, USA. Texture of the HRV data was found to be strongly related to many of the stand attributes evaluated, whereas TM texture was weakly related to all attributes. Data analysis based on regression models indicates that both TM and HRV imagery should yield equally accurate estimates of forest age class and stand structure. It is concluded that the satellite data are a valuable source for estimation of the standard deviation of tree sizes, mean size and density of trees in the upper canopy layers, a structural complexity index, and stand age.

  2. Seed Production in Southern Appalachian Oak Stands

    Treesearch

    Donald E. Beck; David F. Olson

    1968-01-01

    Acorn production in mixed oak stands of different densities varied considerably among stands, years, and species. Density had no effect on total acorn production. Although numerous acorns were initiated each year, the majority either failed to develop fully or were damaged by insects and animals.

  3. The impact of wood ants (Formica rufa) on the distribution and abundance of ground beetles (Coleoptera: Carabidae) in a Scots pine plantation.

    PubMed

    Hawes, C; Stewart, A; Evans, H

    2002-05-01

    The importance of wood ants (Formica rufa) in determining the community structure (defined as the relative abundance of component species) and small-scale distribution of carabids was examined in a mature Scots pine stand in the New Forest, southern England. Carabids and wood ants were sampled by pitfall trapping throughout the forest stand from March to September 1998. The abundance of individual carabid species were modelled using vegetation type (grass or bracken), litter depth and wood ant density as independent explanatory variables. Models were fitted using a maximum-likelihood method (GLIM v.3.77; Baker 1985) with the assumption of a Poisson distribution, using a log-link function. Areas of high wood ant density were characterised by low abundance and species richness of carabids and high percentage dominance by the most commonly sampled species, Abax parallelepipedus. The extent and type of vegetation cover was found to influence the distribution and abundance of certain carabid species but only in areas where the density of wood ants was low. Large-bodied species occurred more frequently in bracken-dominated patches where the litter layer was deeper and the density of potential prey items was higher. Wood ant density was found to be the most important determinant of carabid species abundance in the study site.

  4. Influence of overstory on snow depth and density in hemlock-spruce stands: implications for management of deer habitat in Southeastern Alaska.

    Treesearch

    Thomas A. Hanley; Cathy L. Rose

    1987-01-01

    Snow depth and density were measured in 33 stands of western hemlock-Sitka spruce (Tsuga heterophylla [Rat] Sarg.-Picea sitchensis [Bong.] Carr.) over a 3-year period. The stands, near Juneau, Alaska, provided broad ranges of species composition, age, over-story canopy coverage, tree density, and wood volume. Stepwise multiple regression analyses indicated that both...

  5. Stand density guides for predicting growth of forest tress of southwest Idaho

    Treesearch

    Douglas D. Basford; John Sloan; Joy Roberts

    2010-01-01

    This paper presents a method for estimating stand growth from stand density and average diameter in stands of pure and mixed species in Southwest Idaho. The methods are adapted from a model developed for Douglas-fir, ponderosa pine, and lodgepole pine on the Salmon National Forest. Growth data were derived from ponderosa pine increment cores taken from sample plots on...

  6. Regulating Stand Density by Precommercial Thinning in Naturally Regenerated Loblolly Pine Stands: Evaluation of Management and Economic Opportunities

    Treesearch

    David J. Moorhead; Coleman W. Dangerfield; M. Boyd Edwards

    1997-01-01

    The economic performance of converting 13-year-old, overstocked (>3,000 trees per acre), naturally regenerated pine stands using precommercial thinning at a cost of $140 per acre was modeled for 25-, 35-, and 50-year rotations. The stand density was reduced to 283 trees per acre. Subsequent management scenarios recovered establishment and management costs through...

  7. Response of tree growth and species coexistence to density and species evenness in a young forest plantation with two competing species.

    PubMed

    Collet, Catherine; Ningre, François; Barbeito, Ignacio; Arnaud, Anthony; Piboule, Alexandre

    2014-03-01

    There is considerable evidence for the presence of positive species diversity-productivity relationships in plant populations, but the population parameters determining the type and strength of the relationship are poorly defined. Relationships between species evenness and tree survival or species coexistence are not well established. The objective of this study was to quantify the joint effects of density and species evenness on tree productivity and species coexistence. A 12-year-old experimental tree plantation mixing two species according to a double gradient of density and species proportion was used. A neighbourhood approach was employed and descriptors of local competition were used to model individual tree growth. Fagus sylvatica and Acer pseudoplatanus were used as model species, as they can be considered as ecologically equivalent in their young stages. Density and tree size were primary factors determining individual growth and stand productivity. Species identity had a significant, but less pronounced, role. Stand productivity was highest when species evenness was close to 1 and slightly lower in uneven mixtures. The reduction in stand productivity when species evenness decreased was of similar magnitude irrespective of which species became dominant, indicating symmetric effects for the two species. When examining individual tree growth in response to species proportion for each species separately, it was observed for both species that individual trees exhibited greater growth in uneven mixtures in which the other species was more frequent. The results suggest that mixtures of these two functionally similar species have the highest production at maximum evenness, indicating a complementary effect between them. The presence of a mixture combines both stabilizing mechanisms (individuals from both species show higher growth when surrounded by individuals from the other species) and equalizing mechanisms (the two species have very similar growth curves) that, in turn, determine the species' relative dominance. These processes should act to ensure the long-term coexistence of species.

  8. Response of tree growth and species coexistence to density and species evenness in a young forest plantation with two competing species

    PubMed Central

    Collet, Catherine; Ningre, François; Barbeito, Ignacio; Arnaud, Anthony; Piboule, Alexandre

    2014-01-01

    Background and Aims There is considerable evidence for the presence of positive species diversity–productivity relationships in plant populations, but the population parameters determining the type and strength of the relationship are poorly defined. Relationships between species evenness and tree survival or species coexistence are not well established. The objective of this study was to quantify the joint effects of density and species evenness on tree productivity and species coexistence. Methods A 12-year-old experimental tree plantation mixing two species according to a double gradient of density and species proportion was used. A neighbourhood approach was employed and descriptors of local competition were used to model individual tree growth. Fagus sylvatica and Acer pseudoplatanus were used as model species, as they can be considered as ecologically equivalent in their young stages. Key Results Density and tree size were primary factors determining individual growth and stand productivity. Species identity had a significant, but less pronounced, role. Stand productivity was highest when species evenness was close to 1 and slightly lower in uneven mixtures. The reduction in stand productivity when species evenness decreased was of similar magnitude irrespective of which species became dominant, indicating symmetric effects for the two species. When examining individual tree growth in response to species proportion for each species separately, it was observed for both species that individual trees exhibited greater growth in uneven mixtures in which the other species was more frequent. Conclusions The results suggest that mixtures of these two functionally similar species have the highest production at maximum evenness, indicating a complementary effect between them. The presence of a mixture combines both stabilizing mechanisms (individuals from both species show higher growth when surrounded by individuals from the other species) and equalizing mechanisms (the two species have very similar growth curves) that, in turn, determine the species' relative dominance. These processes should act to ensure the long-term coexistence of species. PMID:24323248

  9. Tree density and permafrost thaw depth influence water limitations on stomatal conductance in Siberian Arctic boreal forests

    NASA Astrophysics Data System (ADS)

    Kropp, H.; Loranty, M. M.; Natali, S.; Kholodov, A. L.; Alexander, H. D.; Zimov, N.

    2017-12-01

    Boreal forests may experience increased water stress under global climate change as rising air temperatures increase evaporative demand and decrease soil moisture. Increases in plant water stress can decrease stomatal conductance, and ultimately, decrease primary productivity. A large portion of boreal forests are located in Siberia, and are dominated by deciduous needleleaf trees, Larix spp. We investigated the variability and drivers of canopy stomatal conductance in upland Larix stands with different stand density that arose from differing fire severity. Our measurements focus on an open canopy stand with low tree density and deep permafrost thaw depth, and a closed canopy stand with high tree density and shallow permafrost thaw depth. We measured canopy stomatal conductance, soil moisture, and micrometeorological variables. Our results demonstrate that canopy stomatal conductance was significantly lower in the closed canopy stand with a significantly higher sensitivity to increases in atmospheric evaporative demand. Canopy stomatal conductance in both stands was tightly coupled to precipitation that occurred over the previous week; however, the closed canopy stand showed a significantly greater sensitivity to increases in precipitation compared to the open canopy stand. Differences in access to deep versus shallow soil moisture and the physical characteristics of the soil profile likely contribute to differences in sensitivity to precipitation between the two stands. Our results indicate that Larix primary productivity may be highly sensitive to changes in evaporative demand and soil moisture that can result of global climate change. However, the effect of increasing air temperatures and changes in precipitation will differ significantly depending on stand density, thaw depth, and the hydraulic characteristics of the soil profile.

  10. Relating Demographic Characteristics of a Small Mammal to Remotely Sensed Forest-Stand Condition

    PubMed Central

    Lada, Hania; Thomson, James R.; Cunningham, Shaun C.; Mac Nally, Ralph

    2014-01-01

    Many ecological systems around the world are changing rapidly in response to direct (land-use change) and indirect (climate change) human actions. We need tools to assess dynamically, and over appropriate management scales, condition of ecosystems and their responses to potential mitigation of pressures. Using a validated model, we determined whether stand condition of floodplain forests is related to densities of a small mammal (a carnivorous marsupial, Antechinus flavipes) in 60 000 ha of extant river red gum (Eucalyptus camaldulensis) forests in south-eastern Australia in 2004, 2005 and 2011. Stand condition was assessed remotely using models built from ground assessments of stand condition and satellite-derived reflectance. Other covariates, such as volumes of fallen timber, distances to floods, rainfall and life stages were included in the model. Trapping of animals was conducted at 272 plots (0.25 ha) across the region. Densities of second-year females (i.e. females that had survived to a second breeding year) and of second-year females with suckled teats (i.e. inferred to have been successful mothers) were higher in stands with the highest condition. There was no evidence of a relationship with stand condition for males or all females. These outcomes show that remotely-sensed estimates of stand condition (here floodplain forests) are relatable to some demographic characteristics of a small mammal species, and may provide useful information about the capacity of ecosystems to support animal populations. Over-regulation of large, lowland rivers has led to declines in many facets of floodplain function. If management of water resources continues as it has in recent decades, then our results suggest that there will be further deterioration in stand condition and a decreased capacity for female yellow-footed antechinuses to breed multiple times. PMID:24621967

  11. Macroscopic time and altitude distribution of plasma turbulence induced in ionospheric modification experiments

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Rose, H.; Dubois, D.; Russell, D.

    1996-03-01

    This is the final report of a three-year Laboratory-Directed Research and Development (LDRD) project at the Los Alamos National Laboratory (LANL). This research concentrated on the time dependence of the heater, induced-turbulence, and electron-density profiles excited in the ionosphere by a powerful radio-frequency heater wave. The macroscopic density is driven by the ponderomotive pressure and the density self-consistently determines the heater propagation. For typical parameters of the current Arecibo heater, a dramatic quasi-periodic behavior was found. For about 50 ms after turn-on of the heater wave, the turbulence is concentrated at the first standing-wave maximum of the heater near reflectionmore » altitude. From 50--100 ms the standing-wave pattern drops by about 1--2 km in altitude and the quasi-periodicity reappears at the higher altitudes with a period of roughly 50 ms. This behavior is due to the half-wavelength density depletion grating that is set up by the ponderomotive pressure at the maxima of the heater standing-wave pattern. Once the grating is established the heater can no longer propagate to higher altitudes. The grating is then unsupported by the heater at these altitudes and decays, allowing the heater to propagate again and initiate another cycle. For stronger heater powers, corresponding to the Arecibo upgrade and the HAARP heater now under construction, the effects are much more dramatic.« less

  12. Relative density: the key to stocking assessment in regional analysis—a forest survey viewpoint.

    Treesearch

    Colin D. MacLean

    1979-01-01

    Relative density is a measure of tree crowding compared to a reference level such as normal density. This stand attribute, when compared to management standards, indicates adequacy of stocking. The Pacific Coast Forest Survey Unit assesses the relative density of each stand sampled by summing the individual density contributions of each tree tallied, thus quantifying...

  13. Investigating the performance of LiDAR-derived biomass information in hydromechanic slope stability modelling

    NASA Astrophysics Data System (ADS)

    Schmaltz, Elmar; Steger, Stefan; Bogaard, Thom; Van Beek, Rens; Glade, Thomas

    2017-04-01

    Hydromechanic slope stability models are often used to assess the landslide susceptibility of hillslopes. Some of these models are able to account for vegetation related effects when assessing slope stability. However, spatial information of required vegetation parameters (especially of woodland) that are defined by land cover type, tree species and stand density are mostly underrepresented compared to hydropedological and geomechanical parameters. The aim of this study is to assess how LiDAR-derived biomass information can help to distinguish distinct tree stand-immanent properties (e.g. stand density and diversity) and further improve the performance of hydromechanic slope stability models. We used spatial vegetation data produced from sophisticated algorithms that are able to separate single trees within a stand based on LiDAR point clouds and thus allow an extraordinary detailed determination of the aboveground biomass. Further, this information is used to estimate the species- and stand-related distribution of the subsurface biomass using an innovative approach to approximate root system architecture and development. The hydrological tree-soil interactions and their impact on the geotechnical stability of the soil mantle are then reproduced in the dynamic and spatially distributed slope stability model STARWARS/PROBSTAB. This study highlights first advances in the approximation of biomechanical reinforcement potential of tree root systems in tree stands. Based on our findings, we address the advantages and limitations of highly detailed biomass information in hydromechanic modelling and physically based slope failure prediction.

  14. Evaluating potential impacts of species conversion on transpiration in the Piedmont of North Carolina

    NASA Astrophysics Data System (ADS)

    Boggs, J.; Treasure, E.; Simpson, G.; Domec, J.; Sun, G.; McNulty, S.

    2010-12-01

    Land management practices that include species conversion or vegetation manipulation can have consequences to surface water availability, groundwater recharge, streamflow generation, and water quality through altering the transpiration processes in forested watersheds. Our objective in this study is to compare stand water use or transpiration in a piedmont mixed hardwood stand (i.e., present stand) to five hypothetical single species stands (i.e., management scenarios), [Quercus spp. (oak), Acer Rubrum (red maple), Liquidambar styraciflua (sweetgum), Liriodendron tulipifera (tulip poplar), and Pinus Taeda (loblolly pine]. Since October 2007, six watersheds with a flume or v-notch weir installed at the watershed outlet have been monitored for baseline streamflow rates (mm d-1). In the summer of 2010, five trees from each of the above species were instrumented with sap flow sensors in the riparian upland of one watershed to develop linkages between stand stream runoff and transpiration. The sap flow or thermal heat dissipation method was used to calculate tree sap flux density for the mixed hardwood stand. Tree sapwood area and stand tree density were then used to compute stand transpiration rates, mm d-1, from June - August 2010. The parameters of the hypothetical single species stands were based on values determined from mixed hardwood stand conditions (e.g., the same stand sapwood area and stand tree density were applied to each option). The diameter at beast height of the monitored trees ranged from 10 cm to 38 cm with a water use range of 1.8 kg d-1 to 104 kg d-1. From our preliminary data, we found daily transpiration from the mixed hardwood stand (2.8 mm d-1 ± 0.06) was significantly (p < 0.05) lower than daily transpiration from the red maple (3.7 mm d-1 ± 0.14) and tulip poplar (3.5 mm d-1 ± 0.12) single species stand management option and significantly (p < 0.05) higher than the loblolly pine (2.3 mm d-1 ± 0.08), sweetgum (2.1 mm d-1 ± 0.08) and oak spp. (1.4 mm d-1 ± 0.04) option. Given that our data represent growing season conditions, these daily transpiration differences are likely a result of physical and physiological differences related to species canopy properties or root distribution and functions. Daily streamflow rates could be reduced by as much as 40% in the red maple scenario because of the increase in daily transpiration. This reduction in flow could have long-term implications and risk to water quality conditions and aquatic species habitat. We will continue to monitor transpiration rates in this mixed hardwood stand to quantify the seasonal variability in water use.

  15. Modelling the effect of autotoxicity on density-dependent phytotoxicity.

    PubMed

    Sinkkonen, A

    2007-01-21

    An established method to separate resource competition from chemical interference is cultivation of monospecific, even-aged stands. The stands grow at several densities and they are exposed to homogenously spread toxins. Hence, the dose received by individual plants is inversely related to stand density. This results in distinguishable alterations in dose-response slopes. The method is often recommended in ecological studies of allelopathy. However, many plant species are known to release autotoxic compounds. Often, the probability of autotoxicity increases as sowing density increases. Despite this, the possibility of autotoxicity is ignored when experiments including monospecific stands are designed and when their results are evaluated. In this paper, I model mathematically how autotoxicity changes the outcome of dose-response slopes as different densities of monospecific stands are grown on homogenously phytotoxic substrata. Several ecologically reasonable relations between plant density and autotoxin exposure are considered over a range of parameter values, and similarities between different relations are searched for. The models indicate that autotoxicity affects the outcome of density-dependent dose-response experiments. Autotoxicity seems to abolish the effects of other phytochemicals in certain cases, while it may augment them in other cases. Autotoxicity may alter the outcome of tests using the method of monospecific stands even if the dose of autotoxic compounds per plant is a fraction of the dose of non-autotoxic phytochemicals with similar allelopathic potential. Data from the literature support these conclusions. A faulty null hypothesis may be accepted if the autotoxic potential of a test species is overlooked in density-response experiments. On the contrary, if test species are known to be non-autotoxic, the method of monospecific stands does not need fine-tuning. The results also suggest that the possibility of autotoxicity should be investigated in many density-response bioassays that are made with even-aged plants, and that measure plant growth or germination.

  16. Semi-automated stand delineation in Mediterranean Pinus sylvestris plantations through segmentation of LiDAR data: The influence of pulse density

    NASA Astrophysics Data System (ADS)

    Varo-Martínez, Mª Ángeles; Navarro-Cerrillo, Rafael M.; Hernández-Clemente, Rocío; Duque-Lazo, Joaquín

    2017-04-01

    Traditionally, forest-stand delineation has been assessed based on orthophotography. The application of LiDAR has improved forest management by providing high-spatial-resolution data on the vertical structure of the forest. The aim of this study was to develop and test a semi-automated algorithm for stands delineation in a plantation of Pinus sylvestris L. using LiDAR data. Three specific objectives were evaluated, i) to assess two complementary LiDAR metrics, Assmann dominant height and basal area, for the characterization of the structure of P. sylvestris Mediterranean forests based on object-oriented segmentation, ii) to evaluate the influence of the LiDAR pulse density on forest-stand delineation accuracy, and iii) to investigate the algorithmś effectiveness in the delineation of P. sylvestris stands for map prediction of Assmann dominant height and basal area. Our results show that it is possible to generate accurate P. sylvestris forest-stand segmentations using multiresolution or mean shift segmentation methods, even with low-pulse-density LiDAR - which is an important economic advantage for forest management. However, eCognition multiresolution methods provided better results than the OTB (Orfeo Tool Box) for stand delineation based on dominant height and basal area estimations. Furthermore, the influence of pulse density on the results was not statistically significant in the basal area calculations. However, there was a significant effect of pulse density on Assmann dominant height [F2,9595 = 5.69, p = 0.003].for low pulse density. We propose that the approach shown here should be considered for stand delineation in other large Pinus plantations in Mediterranean regions with similar characteristics.

  17. Effects of thinning intensities on transpiration and productivity of 50-year-old Pinus koraeinsis stands

    NASA Astrophysics Data System (ADS)

    Park, Juhan; Kim, Taekyu; Moon, Minkyu; Cho, Sungsik; Ryu, Daun; Kim, Hyun Seok

    2015-04-01

    This study investigated the effects of thinning intensities on stand transpiration and productivity of 50-year-old Korean pine forests for two years. Forest thinning, which removes some fraction of trees from stand, is widely conducted for reducing competition between remaining trees, improving tree productivity, reducing the risk of natural fire, and thus maintaining healthy forest. Forest thinning alters the microclimatic conditions such as radiation distribution within canopy, vapor pressure deficit, and amount of available soil water. These changes influence on the tree water use, and related productivity. Thinning was conducted on March, 2012 with two intensities (Control, Light-thinning (20%), and Heavy-thinning (40% of tree density)). Transpiration was estimated from sap flux density, which was measured with Granier-type thermal dissipation sensors. Tree diameter growth was measured with dendrometer, and converted to tree productivity using allometric equations developed specifically in our study sites. The climatic conditions showed little differences between two years. During the first growing season after thinning, stand transpiration was ca. 20% and 42% lower on light-thinning and heavy-thinning stand, respectively, even though sap flux density were higher in thinned stand. The difference in stand transpiration among treatments showed seasonal trends, so it was larger on summer when soil moisture was abundant due to monsoon, but was diminished on spring and autumn when soil moisture was limited. Tree-level productivity increased ca. 8% and 21% on light-thinning and heavy thinning stand, respectively. However, stand net primary production was ca. 20% lower on light-thinning stand, and ca. 31% on heavy-thinning stand. As a result, water use efficiency increased only in heavy-thinning stand. During the second growing season after thinning, stand transpiration was ca. 19% lower on light-thinning stand, and ca. 37% lower on heavy-thinning stand. The reduction of stand transpiration difference in heavy-thinning stand was caused mainly by increase in sap flux density. Trees in thinned stand showed higher productivity, but the magnitude was ca. 4% on light-thinning stand, and ca. 27% on heavy-thinning stand. Stand net primary production was ca. 23% lower on light-thinning stand, and ca. 28% on heavy-thinning stand. As a result, heavy-thinning stand showed highest water use efficiency. These results indicate that there are differences in biological reactions with thinning intensities.

  18. Influence of hardwood midstory and pine species on pine bole arthropods

    Treesearch

    Christopher S. Collins; Richard N. Conner; Daniel Saenz

    2002-01-01

    Arthropod density on the boles of loblolly pines (Pinus taeda) was compared between a stand with and stand without hardwood midstory and between a stand of loblolly and shortleaf pines (P. echinata) in the Stephen E Austin Experimental Forest, Nacogdoches Co., Texas, USA from September 1993 through July 1994. Arthropod density was...

  19. Maintenance cost, toppling risk and size of trees in a self-thinning stand.

    PubMed

    Larjavaara, Markku

    2010-07-07

    Wind routinely topples trees during storms, and the likelihood that a tree is toppled depends critically on its allometry. Yet none of the existing theories to explain tree allometry consider wind drag on tree canopies. Since leaf area index in crowded, self-thinning stands is independent of stand density, the drag force per unit land can also be assumed to be independent of stand density, with only canopy height influencing the total toppling moment. Tree stem dimensions and the self-thinning biomass can then be computed by further assuming that the risk of toppling over and stem maintenance per unit land area are independent of stand density, and that stem maintenance cost is a linear function of stem surface area and sapwood volume. These assumptions provide a novel way to understand tree allometry and lead to a self-thinning line relating tree biomass and stand density with a power between -3/2 and -2/3 depending on the ratio of maintenance of sapwood and stem surface. (c) 2010 Elsevier Ltd. All rights reserved.

  20. Predicting Stand and Stock Tables from a Spacing Study in Naturally Regenerated Longleaf Pine

    Treesearch

    Robert M. Farrar

    1985-01-01

    A prediction system is presented whereby stand and stock tables are calculated for young natural longleaf pine stands of varying initial density. Tables can be output for stand conditions of 10 to 20 years of age, 300 to 1,500 initial trees per acre (at age lo), and 70 to 80 feet in site index (index age 50). The system also allows one to translate from density...

  1. Modeling the size-density relationship in direct-seeded slash pine stands

    Treesearch

    Quang V. Cao; Thomas J. Dean; V. Clark Baldwin

    2000-01-01

    The relationship between quadratic mean diameter and tree density appeared curvilinear on a log–log scale, based on data from direct-seeded slash pine (Pinus elliotti var. elliotti Engelm.) stands. The self-thinning trajectory followed a straight line for high tree density levels and then turned away from this line as tree density...

  2. Thinning of young Douglas-fir forests decreases density of northern flying squirrels in the Oregon Cascades

    USGS Publications Warehouse

    Manning, Tom; Hagar, Joan C.; McComb, Brenda C.

    2012-01-01

    Large-scale commercial thinning of young forests in the Pacific Northwest is currently promoted on public lands to accelerate the development of late-seral forest structure for the benefit of wildlife species such as northern spotted owls (Strix occidentalis caurina) and their prey, including the northern flying squirrel (Glaucomys sabrinus). Attempts to measure the impact of commercial thinning on northern flying squirrels have mostly addressed short-term effects (2–5 years post-thinning) and the few published studies of longer-term results have been contradictory. We measured densities of northern flying squirrels 11–13 years after thinning of young (55–65 years) Douglas-fir forest stands in the Cascade Range of Oregon, as part of the Young Stand Thinning & Diversity Study. The study includes four replicate blocks, each consisting of an unthinned control stand and one stand each of the following thinning treatments: Heavy Thin; Light Thin; and Light Thin with Gaps. Thinning decreased density of northern flying squirrels, and squirrel densities were significantly lower in heavily thinned stands than in more lightly thinned stands. Regression analysis revealed a strong positive relationship of flying squirrel density with density of large (>30 cm diameter) standing dead trees and a negative relationship with percent cover of low understory shrubs. Maintaining sufficient area and connectivity of dense, closed canopy forest is recommended as a strategy to assure that long-term goals of promoting late-seral structure do not conflict with short-term habitat requirements of this important species.

  3. Sapwood area ofPinus contorta stands as a function of mean size and density.

    PubMed

    Long, James N; Dean, Thomas J

    1986-09-01

    An indirect test of the relationship between leaf area and the combination of mean size and density is made in stands of lodgepole pine (Pinus contorta Dougl.). Total sapwood cross-sectional area of these stands is a function of the product of density and mean diameter raised to an exponent of about 1.6. Results from other studies, representing four species, suggest that this relationship between sapwood area and the combination of mean size and density may be general. The implications of the relationship are discussed in the context of evapotranspiration, competition and self-thinning.

  4. Opposing effects of fire severity on climate feedbacks in Siberian larch forests

    NASA Astrophysics Data System (ADS)

    Loranty, M. M.; Alexander, H. D.; Natali, S.; Kropp, H.; Mack, M. C.; Bunn, A. G.; Davydov, S. P.; Erb, A.; Kholodov, A. L.; Schaaf, C.; Wang, Z.; Zimov, N.; Zimov, S. A.

    2017-12-01

    Boreal larch forests in northeastern Siberia comprise nearly 25% of the continuous permafrost zone. Structural and functional changes in these ecosystems will have important climate feedbacks at regional and global scales. Like boreal ecosystems in North America, fire is an important determinant of landscape scale forest distribution, and fire regimes are intensifying as climate warms. In Siberian larch forests are dominated by a single tree species, and there is evidence that fire severity influences post-fire forest density via impacts on seedling establishment. The extent to which these effects occur, or persist, and the associated climate feedbacks are not well quantified. In this study we use forest stand inventories, in situ observations, and satellite remote sensing to examine: 1) variation in forest density within and between fire scars, and 2) changes in land surface albedo and active layer dynamics associated with forest density variation. At the landscape scale we observed declines in Landsat derived albedo as forests recovered in the first several decades after fire, though canopy cover varied widely within and between individual fire scars. Within an individual mid-successional fire scar ( 75 years) we observed canopy cover ranging from 15-90% with correspondingly large ranges of albedo during periods of snow cover, and relatively small differences in albedo during the growing season. We found an inverse relationship between canopy density and soil temperature within this fire scar; high-density low-albedo stands had cooler soils and shallower active layers, while low-density stands had warmer soils and deeper active layers. Intensive energy balance measurements at a high- and low- density site show that canopy cover alters the magnitude and timing of ground heat fluxes that affect active layer properties. Our results show that fire impacts on stand structure in Siberian larch forests affect land surface albedo and active layer dynamics in ways that may lead to opposing climate feedbacks. At effectively large scales these changes constitute positive and negative climate feedbacks, respectively. Accurate predictive understanding of terrestrial Arctic climate feedbacks requires improved knowledge regarding the ecological consequences of changing fire regimes in Siberian boreal forests.

  5. HDL: The "Good" Cholesterol

    MedlinePlus

    ... and LDL (bad) cholesterol: HDL stands for high-density lipoproteins. It is called the "good" cholesterol because ... cholesterol from your body. LDL stands for low-density lipoproteins. It is called the "bad" cholesterol because ...

  6. LDL: The "Bad" Cholesterol

    MedlinePlus

    ... and HDL (good) cholesterol: LDL stands for low-density lipoproteins. It is called the "bad" cholesterol because ... cholesterol in your arteries. HDL stands for high-density lipoproteins. It is called the "good" cholesterol because ...

  7. Tree and understory responses to variable-density thinning in western Washington.

    Treesearch

    Constance A. Harrington; Scott D. Roberts; Leslie C. Brodie

    2005-01-01

    The Olympic Habitat Development Study was initiated in 1994 to evaluate whether active management in 35- to 70-year-old stands could accelerate development of stand structures and plant and animal communities associated with late-successional forests. The study used a variable-density thinning prescription as the main tool to alter stand structure; the prescription...

  8. Growth and soil moisture in thinned lodgepole pine.

    Treesearch

    Walter G. Dahms

    1971-01-01

    A lodgepole pine levels-of-growing-stock study showed that trees growing at lower stand densities had longer crowns and grew more rapidly in diameter but did not grow significantly faster in height. Gross cubic-volume increment decreased with decreasing stand density. The decrease was small per unit of density at the higher densities but much greater at the lower...

  9. Above-ground biomass and structure of pristine Siberian Scots pine forests as controlled by competition and fire.

    PubMed

    Wirth, C; Schulze, E-D; Schulze, W; von Stünzner-Karbe, D; Ziegler, W; Miljukova, I M; Sogatchev, A; Varlagin, A B; Panvyorov, M; Grigoriev, S; Kusnetzova, W; Siry, M; Hardes, G; Zimmermann, R; Vygodskaya, N N

    1999-10-01

    The study presents a data set of above-ground biomass (AGB), structure, spacing and fire regime, for 24 stands of pristine Siberian Scots pine (Pinus sylvestris) forests with lichens (n = 20) or Vaccinium/mosses (n = 4) as ground cover, along four chronosequences. The stands of the "lichen" site type (LT) were stratified into three chronosequences according to stand density and fire history. Allometric equations were established from 90 sample trees for stem, coarse branch, fine branch, twig and needle biomass. The LT stands exhibited a low but sustained biomass accumulation until a stand age of 383 years. AGB reached only 6-10 kg dw  m -2 after 200 years depending on stand density and fire history compared to 20 kg dw  m -2 in the "Vaccinium" type (VT) stands. Leaf area index (LAI) in the LT stands remained at 0.5-1.5 and crown cover was 30-60%, whereas LAI reached 2.5 and crown cover was >100% in the VT stands. Although nearest-neighbour analyses suggested the existence of density-dependent mortality, fire impact turned out to have a much stronger effect on density dynamics. Fire scar dating and calculation of mean and initial fire return intervals revealed that within the LT stands differences in structure and biomass were related to the severity of fire regimes, which in turn was related to the degree of landscape fragmentation by wetlands. Self-thinning analysis was used to define the local carrying capacity for biomass. A series of undisturbed LT stands was used to characterise the upper self-thinning boundary. Stands that had experienced a moderate fire regime were positioned well below the self-thinning boundary in a distinct fire-thinning band of reduced major axis regression slope -0.26. We discuss how this downward shift resulted from alternating phases of density reduction by fire and subsequent regrowth. We conclude that biomass in Siberian Scots pine forests is strongly influenced by fire and that climate change will affect ecosystem functions predominantly via changes in fire regimes.

  10. Leaf δ18O of remaining trees is affected by thinning intensity in a semiarid pine forest.

    PubMed

    Moreno-Gutiérrez, Cristina; Barberá, Gonzalo G; Nicolás, Emilio; DE Luis, Martín; Castillo, Víctor M; Martínez-Fernández, Faustino; Querejeta, José I

    2011-06-01

    Silvicultural thinning usually improves the water status of remaining trees in water-limited forests. We evaluated the usefulness of a dual stable isotope approach (δ¹³C, δ¹⁸O) for comparing the physiological performance of remaining trees between forest stands subjected to two different thinning intensities (moderate versus heavy) in a 60-year-old Pinus halepensis Mill. plantation in semiarid southeastern Spain. We measured bulk leaf δ¹³C and δ¹⁸O, foliar elemental concentrations, stem water content, stem water δ¹⁸O (δ¹⁸O(stem water)), tree ring widths and leaf gas exchange rates to assess the influence of forest stand density on tree performance. Remaining trees in low-density stands (heavily thinned) showed lower leaf δ¹⁸O, and higher stomatal conductance (g(s)), photosynthetic rate and radial growth than those in moderate-density stands (moderately thinned). By contrast, leaf δ¹³C, intrinsic water-use efficiency, foliar elemental concentrations and δ¹⁸O(stem water) were unaffected by stand density. Lower foliar δ¹⁸O in heavily thinned stands reflected higher g(s) of remaining trees due to decreased inter-tree competition for water, whereas higher photosynthetic rate was largely attributable to reduced stomatal limitation to CO₂ uptake. The dual isotope approach provided insight into the early (12 months) effects of stand density manipulation on the physiological performance of remaining trees. © 2011 Blackwell Publishing Ltd.

  11. Stand dynamics in 60-year-old Allegheny hardwoods after thinning

    Treesearch

    Gary W. Miller

    1997-01-01

    Stand dynamics and tree growth in even-aged hardwood stands can be influenced by manipulating relative stand density, species composition, and stand structure. Land managers need quantitative information on the effect of vegetation manipulation to prescribe stand treatments that are appropriate for specific management objectives. Sixty-year-old stands composed of black...

  12. Relation of initial spacing and relative stand density indices to stand characteristics in a Douglas-fir plantation spacing trial

    Treesearch

    Robert O. Curtis; Sheel Bansal; Constance A. Harrington

    2016-01-01

    This report presents updated information on a 1981 Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco var. menziesii ) plantation spacing trial at 33 years from planting. Stand statistics at the most recent measurement were compared for initial spacing of 1 through 6 meters and associated relative densities. There was no clear...

  13. Rotated sigmoid structures in managed uneven-aged northern hardwood stands: a look at the Burr Type III distribution

    Treesearch

    Jeffrey H. Gove; Mark J. Ducey; William B. Leak; Lianjun Zhang

    2008-01-01

    Stand structures from a combined density manipulation and even- to uneven-aged conversion experiment on the Bartlett Experimental Forest (New Hampshire, USA) were examined 25 years after initial treatment for rotated sigmoidal diameter distributions. A comparison was made on these stands between two probability density functions for fitting these residual structures:...

  14. Growth of northern hardwoods in New England: a 25-year update

    Treesearch

    William B. Leak; Jeffrey H. Gove

    2008-01-01

    Twenty-five-year results from a study of four stand density levels and three percentages of sawtimber in a beech-red maple-birch-hemlock stand in New Hampshire showed that moderate stand densities of 60- to 80-ft2/ac with 25- to 30-ft2 of sawtimber produced the best growth responses. Ingrowth was dominated by beech, red...

  15. Variable density thinning promotes variable structural responses 14 years after treatment in the Pacific Northwest

    Treesearch

    John L. Willis; Scott D. Roberts; Constance A. Harrington

    2018-01-01

    Young stands are commonly assumed to require centuries to develop into late-successional forest habitat. This viewpoint reflects the fact that young stands often lack many of the structural features that define late-successional habitat, and that these features derive from complex stand dynamics that are difficult to mimic with forest management. Variable density...

  16. A density management diagram for Scots pine (Pinus sylvestris L.): A tool for assessing the forest's protective effect

    Treesearch

    Giorgio Vacchiano; Renzo Motta; James N. Long; John D. Shaw

    2008-01-01

    Density management diagrams (DMD) are graphical tools used in the design of silvicultural regimes in even-aged forests. They depict the relationship between stand density, average tree size, stand yield and dominant height, based upon relevant ecological and allometric relationships such as the self-thinning rule, the yield-density effect, and site index curves. DMD...

  17. Estimation of Reineke and Volume-Based Maximum Size-Density Lines For Shortleaf Pine

    Treesearch

    Thomas B. Lynch; Robert F. Wittwer; Douglas J. Stevenson

    2004-01-01

    Maximum size-density relationships for Reineke's stand density index as well as for a relationship based on average tree volume were fitted to data from more than a decade of annual remeasurements of plots in unthinned naturally occurring shor tleaf pine in southeaster n Oklahoma. Reineke's stand density index is based on a maximum line of the form log(N) = a...

  18. Size-density metrics, leaf area, and productivity in eastern white pine

    Treesearch

    J. C. Innes; M. J. Ducey; J. H. Gove; W. B. Leak; J. P. Barrett

    2005-01-01

    Size-density metrics are used extensively for silvicultural planning; however, they operate on biological assumptions that remain relatively untested. Using data from 12 even-aged stands of eastern white pine (Pinus strobus L.) growing in southern New Hampshire, we compared size-density metrics with stand productivity and its biological components,...

  19. Influence of Tree Species Composition and Community Structure on Carbon Density in a Subtropical Forest

    PubMed Central

    Hu, Yanqiu; Su, Zhiyao; Li, Wenbin; Li, Jingpeng; Ke, Xiandong

    2015-01-01

    We assessed the impact of species composition and stand structure on the spatial variation of forest carbon density using data collected from a 4-ha plot in a subtropical forest in southern China. We found that 1) forest biomass carbon density significantly differed among communities, reflecting a significant effect of community structure and species composition on carbon accumulation; 2) soil organic carbon density increased whereas stand biomass carbon density decreased across communities, indicating that different mechanisms might account for the accumulation of stand biomass carbon and soil organic carbon in the subtropical forest; and 3) a small number of tree individuals of the medium- and large-diameter class contributed predominantly to biomass carbon accumulation in the community, whereas a large number of seedlings and saplings were responsible for a small proportion of the total forest carbon stock. These findings demonstrate that both biomass carbon and soil carbon density in the subtropical forest are sensitive to species composition and community structure, and that heterogeneity in species composition and stand structure should be taken into account to ensure accurate forest carbon accounting. PMID:26317523

  20. Influence of Tree Species Composition and Community Structure on Carbon Density in a Subtropical Forest.

    PubMed

    Hu, Yanqiu; Su, Zhiyao; Li, Wenbin; Li, Jingpeng; Ke, Xiandong

    2015-01-01

    We assessed the impact of species composition and stand structure on the spatial variation of forest carbon density using data collected from a 4-ha plot in a subtropical forest in southern China. We found that 1) forest biomass carbon density significantly differed among communities, reflecting a significant effect of community structure and species composition on carbon accumulation; 2) soil organic carbon density increased whereas stand biomass carbon density decreased across communities, indicating that different mechanisms might account for the accumulation of stand biomass carbon and soil organic carbon in the subtropical forest; and 3) a small number of tree individuals of the medium- and large-diameter class contributed predominantly to biomass carbon accumulation in the community, whereas a large number of seedlings and saplings were responsible for a small proportion of the total forest carbon stock. These findings demonstrate that both biomass carbon and soil carbon density in the subtropical forest are sensitive to species composition and community structure, and that heterogeneity in species composition and stand structure should be taken into account to ensure accurate forest carbon accounting.

  1. Does Hawaiian native forest conserve water? Lower uptake rates at tree and stand scale by Metrosideros polymorpha relative to plantation species

    NASA Astrophysics Data System (ADS)

    Kagawa, A. K.; Sack, L.; Duarte, T. K.; James, S. A.

    2007-12-01

    Native plants are often claimed to be conservative water users that enhance groundwater recharge compared to faster-growing non-native species that tend to dominate watersheds. This argument would have implications for motivating conservation and restoration of native forest in Hawai'i. However, few studies have examined differences in native and non-native plant transpiration (water use) at species or at stand level. Our aim was determine whether species matter to stand-level water use. We measured plant transpiration in a continuous mosaic of native forest and non-native tree plantation in Honaunau, Hawaii, focusing on endemic dominant tree Metrosideros polymorpha, alien timber trees Eucalyptus saligna and Fraxinus uhdei, and dominant understory Cibotium tree ferns. We measured xylem sap flow for six individuals of each species continuously for over eight weeks, and we estimated stand water use by scaling up these measurements using stand sapwood area and tree fern leaf area values obtained through vegetation surveys. Native forest dominant Metrosideros had the lowest rates of whole-tree daily water use at 8 kg day-1 (200kg m-2sapwood day-1), less than half the daily rates for Eucalyptus or Fraxinus; Metrosideros also had the lowest maximum transpiration rates of the three tree species. At the stand level, Fraxinus-dominated stands had higher water use than Eucalyptus- and Metrosideros- dominated stands due to the species' high sap flow rates, five-fold greater sapwood allocation, and the stands' two-fold greater dominant tree density. In Metrosideros-dominated stands, high Cibotium tree fern leaf area contributed to nearly 60% of water use, indicating the fern's critical role in forest water balance. Stand water use was influenced by factors at various scales, including species composition, stem density, tree sizes, and tree species' sapwood allocation, and was affected significantly by understory contributions. These findings highlight the importance of constituent species in forest water use, and in the case of this Hawaiian forest, indicate conservative water use by native forest.

  2. Density of bunches of native bluebunch wheatgrass and alien crested wheatgrass

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Rickard, W.H.

    The density of bunches of bluebunch wheatgrass in a natural undisturbed stand averaged 3.28 per m/sup 2/ as compared to 2.96 per m/sup 2/ for a nearby stand of crested wheatgrass that was planted 30 years ago. Bunch density was similar in both stands indicating that spacing is a response to an environment deficient in soil water. Bunches of crested wheatgrass on the average weighed 3.5 times more than bunches of bluebunch wheatgrass and they also produced a greater weight of seedheads.

  3. Cultural intensity and planting density effects on individual tree stem growth, stand and crown attributes, and stand dynamics in thinned loblolly pine plantations during the age 12- to age 15- year period in the Upper Coastal Plain and Piedmont of the Southeastern United States

    Treesearch

    Evan Johnson; Michael Kane; Dehai Zhao; Robert Teskey

    2015-01-01

    Three existing loblolly pine (Pinus taeda L.) installations in the Plantation Management Research Cooperative's Upper Coastal Plain/Piedmont Culture Density Study were used to examine the effects of two cultural intensities, four initial planting densities, and their interactions on stem growth at the individual tree level from age 12 to 15 years and at the stand...

  4. The potential of acoustics to determine family differences for wood quality in a loblolly pine (Pinus taeda L.) trial

    Treesearch

    E M Raley; Jennifer H Myszewski; T D Byram

    2007-01-01

    Acoustics have been used to determine wood quality attributes in both standing timber and sawn lumber. Sonic transmission data are collected non-destructively and can act as a surrogate for stiffness, they are directly related to modulus of elasticity (MOE) and closely related to differences in microfibril angle (MFA). Together with wood density, MFA and MOE are the...

  5. Shortleaf pine reproduction abundance and growth in pine-oak stands in the Missouri Ozarks

    Treesearch

    Elizabeth M. Blizzard; Doyle Henken; John M. Kabrick; Daniel C. Dey; David R. Larsen; David Gwaze

    2007-01-01

    We conducted an operational study to evaluate effect of site preparation treatments on pine reproduction density and the impact of overstory basal area and understory density on pine reproduction height and basal diameter in pine-oak stands in the Missouri Ozarks. Stands were harvested to or below B-level stocking, but patchiness of the oak decline lead to some plots...

  6. Snag densities in old-growth stands on the Gasquet Ranger District, Six Rivers National Forest, California

    Treesearch

    Thomas M. Jimerson

    1989-01-01

    Baseline levels for densities of snags (standing dead trees) wered etermined in undisturbed old-growth stands on the Gasquet Ranger District. Six Riven National Forest, California. Snag species, number, diameter at breast height (d.b.h.), height, cavity type, cavity use, decay class, and snag origin were recorded on 317 plots over a 2-year period. The 2121 snags...

  7. Overstory density affects field performance of underplanted red oak (Quercus rubra L.) in Ontario

    Treesearch

    Daniel C. Dey; William C. Parker

    1997-01-01

    Red oak seedlings were underplanted in a closed-canopy mature northern hardwood stand and an adjacent shelterwood in central Ontario. Overstory density effects on seedling survival and growth were assessed 2 yr after planting. After 2 yr, seedling survival was 90% in the uncut stand and over 99% in the shelterwood. Seedlings in the uncut stand experienced negligible or...

  8. Stump sprout dynamics in response to reductions in stand density for nine upland hardwood species in the southern Appalachian Mountains

    Treesearch

    Tara L. Keyser; Stanley J. Zarnoch

    2014-01-01

    Much about stump sprout dynamics of upland hardwood trees species has been obtained in clearcuts. Information on the response of stump sprouts to alternative silvicultural treatments, including treatments that manipulate stand density and stand structure is lacking. In this study we examined the influence of harvest season and levels of basal area reduction on the...

  9. Cone Characteristics and Seed Quality 10 Years After An Uneven-Aged Regeneration Cut In Shortleaf Pine Stands

    Treesearch

    Kenneth J. Grayson; Robert F. Wittwer; Michael G. Shelton

    2002-01-01

    Cone characteristics and seed quality for 16 released (stand density 14 square meters per hectare) and 16 unreleased (stand density 28 square meters per hectare) shortleaf pine (Pinus echinata Mill.) trees were described by d.b.h. class (28, 33, 38, 43 centimeters) and crown position (upper south, upper north, lower south, and lower north). The 38-...

  10. Occurrence of termites (Isoptera) on living and standing dead trees in a tropical dry forest in Mexico.

    PubMed

    Calderón-Cortés, Nancy; Escalera-Vázquez, Luis H; Oyama, Ken

    2018-01-01

    Termites play a key role as ecosystem engineers in numerous ecological processes though their role in the dynamics of wood degradation in tropical dry forests, particularly at the level of the crown canopy, has been little studied. In this study, we analysed the occurrence of termites in the forest canopy by evaluating the density and proportion of living and standing dead trees associated with termites in deciduous and riparian habitats of the tropical dry forest in Chamela, Mexico. The results indicated that 60-98% of standing dead trees and 23-59% of living trees in Chamela were associated with termites. In particular, we found that the density of standing dead trees was higher in deciduous forests (0.057-0.066 trees/m 2 ) than in riparian forests (0.022 and 0.027 trees/m 2 ), even though the proportion of trees was not significantly different among habitats. Additionally, we found a higher density of trees associated with termites in trees of smaller size classes (0.01-0.09 trees/m 2 ) than in larger class sizes (0-0.02 trees/m 2 ). Interestingly, 72% of variation in the density of trees associated with termites is explained by the density of standing dead trees. Overall, these results indicate that standing dead tree availability might be the main factor regulating termite populations in Chamela forest and suggest that termites could play a key role in the decomposition of above-ground dead wood, mediating the incorporation of suspended and standing dead wood into the soil.

  11. Occurrence of termites (Isoptera) on living and standing dead trees in a tropical dry forest in Mexico

    PubMed Central

    Escalera-Vázquez, Luis H.; Oyama, Ken

    2018-01-01

    Termites play a key role as ecosystem engineers in numerous ecological processes though their role in the dynamics of wood degradation in tropical dry forests, particularly at the level of the crown canopy, has been little studied. In this study, we analysed the occurrence of termites in the forest canopy by evaluating the density and proportion of living and standing dead trees associated with termites in deciduous and riparian habitats of the tropical dry forest in Chamela, Mexico. The results indicated that 60–98% of standing dead trees and 23–59% of living trees in Chamela were associated with termites. In particular, we found that the density of standing dead trees was higher in deciduous forests (0.057–0.066 trees/m2) than in riparian forests (0.022 and 0.027 trees/m2), even though the proportion of trees was not significantly different among habitats. Additionally, we found a higher density of trees associated with termites in trees of smaller size classes (0.01–0.09 trees/m2) than in larger class sizes (0–0.02 trees/m2). Interestingly, 72% of variation in the density of trees associated with termites is explained by the density of standing dead trees. Overall, these results indicate that standing dead tree availability might be the main factor regulating termite populations in Chamela forest and suggest that termites could play a key role in the decomposition of above-ground dead wood, mediating the incorporation of suspended and standing dead wood into the soil. PMID:29785342

  12. SPRUCE S1 Bog Fine-root Production and Standing Crop Assessed With Minirhizotrons in the Southern and Northern Ends of the S1 Bog

    DOE Data Explorer

    Iversen, C. M. [Oak Ridge National Laboratory, Oak Ridge, Tennessee, USA; Childs, J. [Oak Ridge National Laboratory, Oak Ridge, Tennessee, USA; Norby, R. J. [Oak Ridge National Laboratory, Oak Ridge, Tennessee, USA; Garrett, A. [Oak Ridge National Laboratory, Oak Ridge, Tennessee, USA; Martin, A. [Oak Ridge National Laboratory, Oak Ridge, Tennessee, USA; Spence, J. [Oak Ridge National Laboratory, Oak Ridge, Tennessee, USA; Ontl, T. A. [Oak Ridge National Laboratory, Oak Ridge, Tennessee, USA

    2017-01-01

    This data set reports fine-root peak growth and standing crop measurements from a forested, ombrotrophic bog as determined using non-destructive minirhizotron technology. Minirhizotron images were collected throughout the growing seasons of 2011 and 2012 at the southern and northern ends of the S1 bog across gradients of tree density in paired hummock and hollow microtopography. The dominant woody species in the bog, and focus of the investigation, were trees Picea mariana and Larix laricina, and ericaceous shrubs Rhododendron groenlandicum and Chamaedaphne calyculata.

  13. The Effect of Density on the Height-Diameter Relationship

    Treesearch

    Boris Zeide; Curtis Vanderschaaf

    2002-01-01

    Using stand density along with mean diameter to predict average height increases the proportion of explained variance. This result, obtained from permanent plots established in a loblolly pine plantation thinned to different levels, makes sense. We know that due to competition, trees with the same diameter are taller in denser stands. Diameter and density are not only...

  14. Stand conditions associated with tree regeneration in sierran mixed-conifer forests.

    Treesearch

    Andrew N. Gray; Harold S.J. Zald; Ruth A. Kern; Malcolm North

    2005-01-01

    Fire suppression has significantly increased canopy cover, litter depth, and stem density in many western forests, altering microsite conditions that affect tree seedling establishment. We conducted studies in a mixed-conifer forest in the Sierra Nevada, California, to determine relationships between established understory trees and microsite quality, and to examine...

  15. Crown size relationships for black willow in the Lower Mississippi Alluvial Valley

    Treesearch

    Jamie L. Schuler; Bradley Woods; Joshua Adams; Ray Souter

    2015-01-01

    Growing space requirements derived from maximum and minimum crown sizes have been identified for many southern hardwood species. These requirements help managers assess stocking levels, schedule intermediate treatments, and even assist in determining planting densities. Throughout the Mississippi Alluvial Valley, black willow (Salix nigra Marsh.) stands are common...

  16. Determination of Fluid Density and Viscosity by Analyzing Flexural Wave Propagations on the Vibrating Micro-Cantilever

    PubMed Central

    Kim, Deokman; Hong, Seongkyeol; Park, Junhong

    2017-01-01

    The determination of fluid density and viscosity using most cantilever-based sensors is based on changes in resonant frequency and peak width. Here, we present a wave propagation analysis using piezoelectrically excited micro-cantilevers under distributed fluid loading. The standing wave shapes of microscale-thickness cantilevers partially immersed in liquids (water, 25% glycerol, and acetone), and nanoscale-thickness microfabricated cantilevers fully immersed in gases (air at three different pressures, carbon dioxide, and nitrogen) were investigated to identify the effects of fluid-structure interactions to thus determine the fluid properties. This measurement method was validated by comparing with the known fluid properties, which agreed well with the measurements. The relative differences for the liquids were less than 4.8% for the densities and 3.1% for the viscosities, and those for the gases were less than 6.7% for the densities and 7.3% for the viscosities, showing better agreements in liquids than in gases. PMID:29077005

  17. A tree-by-tree measure of site utilization for grand fir related to stand density index

    Treesearch

    Albert R. Stage

    1968-01-01

    The usefulness of stand density index (SDI) has been limited by lack of a way to partition its nonlinear expression into additive components to describe the relative stocking of a stand by species or quality classes. In this paper, a linear equation is derived to permit such a partition. A closely related expression for grand fir stocking is given that retains the...

  18. Computerized algorithms for partial cuts

    Treesearch

    R.L. Ernst; S.L. Stout

    1991-01-01

    Stand density, stand structure (diameter distribution), and species composition are all changed by intermediate treatments in forest stands. To use computer stand-growth simulators to assess the effects of different treatments on stand growth and development, users must be able to duplicate silviculturally realistic treatments in the simulator. In this paper, we review...

  19. Dynamics of dense direct-seeded stands of southern pines

    Treesearch

    J.C.G. Goelz

    2006-01-01

    Direct seeding of southern pines is an effective method of artificial regeneration, producing extremely dense stands when survival exceeds expectations. Long-term studies of dense direct-seeded stands provide ideal data for exploring development of stands as they approach the limit of maximum stand density. I present data from seven studies with ages of stands ranging...

  20. Estimating cavity tree abundance using nearest neighbor imputation methods for western Oregon and Washington forests

    Treesearch

    Hailemariam Temesgen; Tara M. Barrett; Greg Latta

    2008-01-01

    Cavity trees contribute to diverse forest structure and wildlife habitat. For a given stand, the size and density of cavity trees indicate its diversity, complexity, and suitability for wildlife habitat. Size and density of cavity trees vary with stand age, density, and structure. Using Forest Inventory and Analysis (FIA) data collected in western Oregon and western...

  1. Effects of low-density thinning in a declining white pine stand in Maine

    Treesearch

    William B. Leak; Mariko. Yamasaki

    2013-01-01

    Low-density (32 ft2/acre residual basal area) and medium-low density (60 ft2/acre residual basal area) thinnings were studied over a 4-year period in a declining white pine stand on the Massabesic Experimental Forest in southern Maine. Gross basal area growth at 60 ft2 was about three-fourths the rate...

  2. Controls on stand transpiration and soil water utilization along a tree density gradient in a Neotropical savanna

    Treesearch

    Sandra J. Bucci; Fabian G. Scholz; Guillermo Goldstein; William A. Hoffmann; Frederick C. Meinzer; Augusto C. Franco; Thomas Giambelluca; Fernando Miralles-Wilhelm

    2008-01-01

    Environmental controls of stand-level tree transpiration (E) and seasonal patterns of soil water utilization were studied in five central Brazilian savanna (Cerrado) sites differing in tree density. Tree density of Cerrado vegetation in the study area consistently changes along topographic gradients from ~1,000 trees ha-1 in open savannas (campo...

  3. Effect of Experimentally Manipulated Fire Regimes on the Response of Forests to Drought

    NASA Astrophysics Data System (ADS)

    Refsland, T. K.; Knapp, B.; Fraterrigo, J.

    2017-12-01

    Climate change is expected to increase drought stress in many forests and alter fire regimes. Fire can reduce tree density and thus competition for limited water, but the effects of changing fire regimes on forest productivity during drought remain poorly understood. We measured the annual ring-widths of adult oak (Quercus spp.) trees in Mark Twain National Forest, Missouri USA that experienced unburned, annual or periodic (every 4 years) surface fire treatments from 1951 - 2015. Severe drought events were identified using the BILJOU water balance model. We determined the effect of fire treatment on stand-level annual growth rates as well as stand-level resistance and resilience to drought, defined as the drought-induced reduction in growth and post-drought recovery in growth, respectively. During favorable wet years, annual and periodic fire treatments reduced annual growth rates by approximately 10-15% relative to unburned controls (P < 0.001). Stand-level growth rates declined 22-40% during drought events (P < 0.001), but fire-driven changes to stand basal area had no effect on the resistance or resilience of trees to drought. The decline in annual growth rates of burned stands during favorable wet years was likely caused by increased nitrogen (N) limitation in burned plots. After 60 years of treatment, burned plots experienced 30% declines in total soil N relative to unburned plots. Our finding that drought resistance and resilience were similar across all treatments suggest that fire-driven reductions in stand density may have negligible effects on soil moisture availability during drought. Our results highlight that climate-fire interactions can have important long-term effects on forest productivity.

  4. Proceedings of the National Silviculture Workshop: Density of Stocking Control; Eugene, Oregon; October 13-15, 1976

    Treesearch

    Jack H. Usher; Daniel B. Jones; A. R. Stage; Benjamin A. Roach; Gilbert B. Schubert; Darrell W. Crawford; Gilbert H. Schubert; Walter Fox; Edward A. Smith; Richard E. Lowrey Sofes; Richard F. Watt

    1976-01-01

    The 1976 National Silviculture Workshop was held in Eugene, Oregon, on October 13-15, 1976. The objectives were to discuss second growth management of individual stands, with particular emphasis on the control of stand density.

  5. Transition from Propagating Polariton Solitons to a Standing Wave Condensate Induced by Interactions

    NASA Astrophysics Data System (ADS)

    Sich, M.; Chana, J. K.; Egorov, O. A.; Sigurdsson, H.; Shelykh, I. A.; Skryabin, D. V.; Walker, P. M.; Clarke, E.; Royall, B.; Skolnick, M. S.; Krizhanovskii, D. N.

    2018-04-01

    We explore phase transitions of polariton wave packets, first, to a soliton and then to a standing wave polariton condensate in a multimode microwire system, mediated by nonlinear polariton interactions. At low excitation density, we observe ballistic propagation of the multimode polariton wave packets arising from the interference between different transverse modes. With increasing excitation density, the wave packets transform into single-mode bright solitons due to effects of both intermodal and intramodal polariton-polariton scattering. Further increase of the excitation density increases thermalization speed, leading to relaxation of the polariton density from a solitonic spectrum distribution in momentum space down to low momenta, with the resultant formation of a nonequilibrium condensate manifested by a standing wave pattern across the whole sample.

  6. Transition from Propagating Polariton Solitons to a Standing Wave Condensate Induced by Interactions.

    PubMed

    Sich, M; Chana, J K; Egorov, O A; Sigurdsson, H; Shelykh, I A; Skryabin, D V; Walker, P M; Clarke, E; Royall, B; Skolnick, M S; Krizhanovskii, D N

    2018-04-20

    We explore phase transitions of polariton wave packets, first, to a soliton and then to a standing wave polariton condensate in a multimode microwire system, mediated by nonlinear polariton interactions. At low excitation density, we observe ballistic propagation of the multimode polariton wave packets arising from the interference between different transverse modes. With increasing excitation density, the wave packets transform into single-mode bright solitons due to effects of both intermodal and intramodal polariton-polariton scattering. Further increase of the excitation density increases thermalization speed, leading to relaxation of the polariton density from a solitonic spectrum distribution in momentum space down to low momenta, with the resultant formation of a nonequilibrium condensate manifested by a standing wave pattern across the whole sample.

  7. Relation of initial spacing and relative stand density indices to stand characteristics in a Douglas-fir plantation spacing trial

    USGS Publications Warehouse

    Curtis, Robert O.; Bansal, Sheel; Harrington, Constance A.

    2016-01-01

    This report presents updated information on a 1981 Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco var. menziesii) plantation spacing trial at 33 years from planting. Stand statistics at the most recent measurement were compared for initial spacing of 1 through 6 meters and associated relative densities. There was no clear relationship of spacing to top height. Diameter, live crown ratio, and percent survival increased with spacing; basal area and relative density decreased with increase in spacing. Volume in trees ≥ 4 cm diameter was greatest at 2 m spacing, while utilizable volume (trees ≥20 cm dbh) was greatest at 4 m spacing. Live crown ratio decreased and total crown projectional area increased with increasing relative density indices. Total crown projectional area was more closely related to relative density than to basal area.

  8. Estimating stand structure using discrete-return lidar: an example from low density, fire prone ponderosa pine forests

    USGS Publications Warehouse

    Hall, S. A.; Burke, I.C.; Box, D. O.; Kaufmann, M. R.; Stoker, Jason M.

    2005-01-01

    The ponderosa pine forests of the Colorado Front Range, USA, have historically been subjected to wildfires. Recent large burns have increased public interest in fire behavior and effects, and scientific interest in the carbon consequences of wildfires. Remote sensing techniques can provide spatially explicit estimates of stand structural characteristics. Some of these characteristics can be used as inputs to fire behavior models, increasing our understanding of the effect of fuels on fire behavior. Others provide estimates of carbon stocks, allowing us to quantify the carbon consequences of fire. Our objective was to use discrete-return lidar to estimate such variables, including stand height, total aboveground biomass, foliage biomass, basal area, tree density, canopy base height and canopy bulk density. We developed 39 metrics from the lidar data, and used them in limited combinations in regression models, which we fit to field estimates of the stand structural variables. We used an information–theoretic approach to select the best model for each variable, and to select the subset of lidar metrics with most predictive potential. Observed versus predicted values of stand structure variables were highly correlated, with r2 ranging from 57% to 87%. The most parsimonious linear models for the biomass structure variables, based on a restricted dataset, explained between 35% and 58% of the observed variability. Our results provide us with useful estimates of stand height, total aboveground biomass, foliage biomass and basal area. There is promise for using this sensor to estimate tree density, canopy base height and canopy bulk density, though more research is needed to generate robust relationships. We selected 14 lidar metrics that showed the most potential as predictors of stand structure. We suggest that the focus of future lidar studies should broaden to include low density forests, particularly systems where the vertical structure of the canopy is important, such as fire prone forests.

  9. Seasonal Lateral Root Growth of Juvenile Loblolly Pine After Thinning and Fertilization on Gulf Coastal Plain Site

    Treesearch

    Mary Anne Sword; James D. Haywood; C. Dan Andries

    1998-01-01

    In 1989, two levels each of stand density and fertilization were factorially established in an 8-year-old loblolly pine plantation on a P-deficient site. Levels of stand density were nonthinned at 2,732 trees per hectare and thinned at 721 trees per hectare. Fertilizer levels were none or application of 150 kilograms P plus 135 kilograms N per hectare. In 1994, stand...

  10. Stand density, stand structure, and species composition in transition oak stands of northwestern Pennsylvania

    Treesearch

    S.L. Stout

    1991-01-01

    Transition stands, those containing species associated with both the northern hardwood and oak-hickory forest types, are important to forest diversity in northwestern Pennsylvania. These stands have high value for a variety of forest uses, including timber production, wildlife habitat, and aesthetics. Diameter distributions are characteristically stratified by species...

  11. Reducing stand densities in immature and mature stands, Applegate Watershed, Southwest Oregon.

    Treesearch

    Marty L. Main; Michael P. Amaranthus

    1996-01-01

    Abstract Throughout the Applegate watershed, dense, overstocked, immature stands of mixed conifers and hardwoods and declining stands of mature conifers present significant and complex silvicultural problems. Stand stagnation is common, as is loss of large-diameter conifers from insects and wildfire. Treatments designed to maintain or encourage development of large-...

  12. Response of old-growth conifers to reduction in stand density in western Oregon forests

    USGS Publications Warehouse

    Latham, P.; Tappeiner, J. C.

    2002-01-01

    The positive growth response of healthy young trees to density reduction is well known. In contrast, large old trees are usually thought to be intrinsically limited in their ability to respond to increased growing space; therefore, density reduction is seldom used in stands of old-growth trees. We tested the null hypothesis that old-growth trees are incapable of responding with increased growth following density reduction. The diameter growth response of 271 Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco), ponderosa pine (Pinus ponderosa Dougl. ex Laws) and sugar pine (Pinus lambertiana Dougl.) trees ranging in age from 158 to 650 years was examined 20 to 50 years after density reduction. Density reduction involved either light thinning with removal of less vigorous trees, or shelterwood treatments in which overstory trees were not removed. Ratios of basal area growth after treatment to basal area growth before treatment, and several other measures of growth, all indicated that the old trees sometimes benefited and were not harmed by density reduction. Growth increased by 10% or more for 68% of the trees in treated stands, and nearly 30% of trees increased growth by over 50%. This growth response persisted for at least 20 years. During this 20-year period, only three trees in treated stands (1.5%) exhibited a rapid decrease in growth, whereas growth decreased in 64% of trees in untreated stands. The length of time before a growth response to density reduction occurred varied from 5 to 25 years, with the greatest growth response often occurring 20 to 25 years after treatment. These results have important implications both for the basic biology of aging in woody plants as well as for silvicultural practices in forests with old-growth trees.

  13. Melting slope of MgO from molecular dynamics and density functional theory

    NASA Astrophysics Data System (ADS)

    Tangney, Paul; Scandolo, Sandro

    2009-09-01

    We combine density functional theory (DFT) with molecular dynamics simulations based on an accurate atomistic force field to calculate the pressure derivative of the melting temperature of magnesium oxide at ambient pressure—a quantity for which a serious disagreement between theory and experiment has existed for almost 15 years. We find reasonable agreement with previous DFT results and with a very recent experimental determination of the slope. We pay particular attention to areas of possible weakness in theoretical calculations and conclude that the long-standing discrepancy with experiment could only be explained by a dramatic failure of existing density functionals or by flaws in the original experiment.

  14. Stand density management of southern bottomland hardwoods

    Treesearch

    J.C.G. Goelz; J.S. Meadows

    1997-01-01

    We present tools to guide stand density management of southern bottomiand hardwoods and we provide guidance in their implementation. We present stocking guides for southern bottomland hardwoods and variants for associated forest types, water tupelo (Nyssa aquarica L.) and sweetgum (Liquidumbar styraciflua L.). The A-line...

  15. Implementing optimal thinning strategies

    Treesearch

    Kurt H. Riitters; J. Douglas Brodie

    1984-01-01

    Optimal thinning regimes for achieving several management objectives were derived from two stand-growth simulators by dynamic programming. Residual mean tree volumes were then plotted against stand density management diagrams. The results supported the use of density management diagrams for comparing, checking, and implementing the results of optimization analyses....

  16. Short- and long-term influence of stand density on soil microbial communities in ponderosa pine forests

    Treesearch

    Steven T. Overby

    2009-01-01

    Soil microbial communities process plant detritus and returns nutrients needed for plant growth. Increased knowledge of this intimate linkage between plant and soil microbial communities will provide a better understanding of ecosystem response to changing abiotic and biotic conditions. This dissertation consists of three studies to determine soil microbial community...

  17. Snag longevity in relation to wildfire and postfire salvage logging

    Treesearch

    Robin E. Russell; Victoria A. Saab; Jonathan G. Dudley; Jay J. Rotella

    2006-01-01

    Snags create nesting, foraging, and roosting habitat for a variety of wildlife species. Removal of snags through postfire salvage logging reduces the densities and size classes of snags remaining after wildfire. We determined important variables associated with annual persistence rates (the probability a snag remains standing from 1 year to the next) of large conifer...

  18. A density management diagram for longleaf pine stands with application to red-cockaded woodpecker habitat

    Treesearch

    John D. Shaw; James N. Long

    2007-01-01

    We developed a density management diagram (DMD) for longleaf pine (Pinus palustris P. Mill.) using data from Forest Inventory and Analysis plots. Selection criteria were for purity, defined as longleaf pine basal area (BA) that is 90% or more of plot BA, and even-agedness, as defined by a ratio between two calculations of stand density index. The...

  19. Can live tree size-density relationships provide a mechanism for predicting down and dead tree resources?

    Treesearch

    Christopher Woodall; James Westfall

    2009-01-01

    Live tree size-density relationships in forests have long provided a framework for understanding stand dynamics. There has been little examination of the relationship between the size-density attributes of live and standing/down dead trees (e.g., number and mean tree size per unit area, such information could help in large-scale efforts to estimate dead wood resources...

  20. Results after 20 years from a western larch levels-of-growing-stock study.

    Treesearch

    K.W. Seidel

    1987-01-01

    The 20-year growth response from a levels-of-growing-stock study in an even-aged western larch stand in eastern Oregon, first thinned at age 33, showed that trees growing at low stand densities grew more rapidly in diameter than trees in high-density plots. Height growth was relatively uniform among density levels. Both basal-area and total cubic-volume increment...

  1. Experimental observation of standing wave effect in low-pressure very-high-frequency capacitive discharges

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Liu, Yong-Xin; Gao, Fei; Liu, Jia

    2014-07-28

    Radial uniformity measurements of plasma density were carried out by using a floating double probe in a cylindrical (21 cm in electrode diameter) capacitive discharge reactor driven over a wide range of frequencies (27–220 MHz). At low rf power, a multiple-node structure of standing wave effect was observed at 130 MHz. The secondary density peak caused by the standing wave effect became pronounced and shifts toward the axis as the driving frequency further to increase, indicative of a much more shortened standing-wave wavelength. With increasing rf power, the secondary density peak shift toward the radial edge, namely, the standing-wave wavelength was increased,more » in good qualitative agreement with the previous theory and simulation results. At higher pressures and high frequencies, the rf power was primarily deposited at the periphery of the electrode, due to the fact that the waves were strongly damped as they propagated from the discharge edge into the center.« less

  2. A preliminary model of yellow-poplar seedling establishment two years after a growing season prescribed fire in southern Appalachian oak stands

    Treesearch

    Henry McNab

    2016-01-01

    Factors affecting the density and distribution of yellow-poplar regeneration after a single growing season prescribed fire were studied in mature upland oak stands in the southern Appalachian Mountains. In burned and unburned stands, density of one and two year old yellow-poplar seedlings was inventoried within 50 m from isolated yellow-poplar canopy seed trees in...

  3. Analysis of standing sound waves using holographic interferometry

    NASA Astrophysics Data System (ADS)

    Russell, Daniel A.; Parker, David E.; Hughes, Russell S.

    2009-08-01

    Optical holographic interferometry was used to study standing sound waves in air inside a resonance tube driven by a small loudspeaker at one end. The front face of the resonance tube was constructed with plexiglass, allowing optical interrogation of the tube interior. The object beam of the holographic setup was directed through the plexiglass and reflected off the back wall of the resonator. When driven at resonance, the fluctuations in the air density at the antinodes altered the refractive index of the air in the tube, causing interference patterns in the resulting holographic images. Real-time holography was used to determine resonance frequencies and to measure the wavelengths of the standing waves. Time-average holography was used to observe the effect of increasing the sound pressure level on the resulting fringe pattern. A simple theory was developed to successfully predict the fringe pattern.

  4. Spatial cross-correlation of undisturbed, natural shortleaf pine stands in northern Georgia

    Treesearch

    Robin M. Reich; Raymond L. Czaplewski; William A. Bechtold

    1994-01-01

    In this study a cross-correlation statistic is used to analyse the spatial relationship among stand characteristics of natural, undisturbed shortleaf pine stands sampled during 1961-72 and 1972-82 in northern Georgia. Stand characteristics included stand age, site index, tree density, hardwood competition, and mortality. In each time period, the spatial cross-...

  5. Changing stand structure and regional growth reductions in Georgia's natural pine stands

    Treesearch

    W.A. Bechtold; G.A. Ruark; F.T. Lloyd

    1991-01-01

    Forest Inventory and Analysis (FIA) data indicate reductions in the growth of naturally regenerated pines in Georgia between the two latest measurement periods (1961-1972 vs. 1972-1982). Analysis of Covariance was used to adjust stand-level basal area growth rates for differences between periods in stand age, stand density, site index, mortality, and hardwood...

  6. The application of a geometric optical canopy reflectance model to semiarid shrub vegetation

    NASA Technical Reports Server (NTRS)

    Franklin, Janet; Turner, Debra L.

    1992-01-01

    Estimates are obtained of the average plant size and density of shrub vegetation on the basis of SPOT High Resolution Visible Multispectral imagery from Chihuahuan desert areas, using the Li and Strahler (1985) model. The aggregated predictions for a number of stands within a class were accurate to within one or two standard errors of the observed average value. Accuracy was highest for those classes of vegetation where the nonrandom scrub pattern was characterized for the class on the basis of the average coefficient of the determination of density.

  7. Propagation Modelling and Decision Aids for Communications, Radar and Navigation Systems (La Modelisation de la Propagation et Aides a la Decision Pour les Sysemes de elecommunicaions, de Radar et de Navigation)

    DTIC Science & Technology

    1994-09-01

    the refractive index i. can be density, temperature , ion composition, ionospheric determined from a simplified form of the Appleton- electric field...see Cannon 119941. the electron density profile is based upon the underlying neutral composition. temperature and wind together with electric field...in many of the newer HF predictions decision software , NSSDC/WDC-A-R&S 90-19, National Space aids. They also provide a very useful stand alone

  8. Tree-, stand- and site-specific controls on landscape-scale patterns of transpiration

    NASA Astrophysics Data System (ADS)

    Hassler, Sibylle; Markus, Weiler; Theresa, Blume

    2017-04-01

    Transpiration is a key process in the hydrological cycle and a sound understanding and quantification of transpiration and its spatial variability is essential for management decisions as well as for improving the parameterisation of hydrological and soil-vegetation-atmosphere transfer models. For individual trees, transpiration is commonly estimated by measuring sap flow. Besides evaporative demand and water availability, tree-specific characteristics such as species, size or social status control sap flow amounts of individual trees. Within forest stands, properties such as species composition, basal area or stand density additionally affect sap flow, for example via competition mechanisms. Finally, sap flow patterns might also be influenced by landscape-scale characteristics such as geology, slope position or aspect because they affect water and energy availability; however, little is known about the dynamic interplay of these controls. We studied the relative importance of various tree-, stand- and site-specific characteristics with multiple linear regression models to explain the variability of sap velocity measurements in 61 beech and oak trees, located at 24 sites spread over a 290 km2-catchment in Luxembourg. For each of 132 consecutive days of the growing season of 2014 we modelled the daily sap velocities of these 61 trees and determined the importance of the different predictors. Results indicate that a combination of tree-, stand- and site-specific factors controls sap velocity patterns in the landscape, namely tree species, tree diameter, the stand density, geology and aspect. Compared to these predictors, spatial variability of atmospheric demand and soil moisture explains only a small fraction of the variability in the daily datasets. However, the temporal dynamics of the explanatory power of the tree-specific characteristics, especially species, are correlated to the temporal dynamics of potential evaporation. Thus, transpiration estimates at the landscape scale would benefit from not only considering hydro-meteorological drivers, but also including tree, stand and site characteristics in order to improve the spatial representation of transpiration for hydrological and soil-vegetation-atmosphere transfer models.

  9. Logging impacts on forest structure and seedling dynamics in a Prioria copaifera (Fabaceae) dominated tropical rain forest (Talamanca, Costa Rica).

    PubMed

    Valverde-Barrantes, Oscar J; Rocha, Oscar J

    2014-03-01

    The factors that determine the existence of tropical forests dominated by a single species (monodominated forests) have been the subject of debate for a long time. It has been hypothesized that the low frequency of disturbances in monodominated forests and the tolerance to shade of the monodominant species are two important factors explaining the prolonged dominance of a single species. We determined the role of these two factors by examining the effects of logging activities on the floristic composition and seedling dynamics in a Prioria copaifera dominated forest in Southeastern Costa Rica. We determined the floristic composition for trees > or = 2.5cm DBH and the associated recruitment, survival and mortality of tree canopy seedlings in two sites logged two (L-02) and 12 years (L-12) prior to sampling and an unlogged forest (ULF). Our results showed that L-02 stands had lower species richness (25 species) than the L-12 and ULF stands (49 and 46 species, respectively). As expected, we found significant logging effects on the canopy structure of the altered forests, particularly when comparing the L-02 and the ULF stands. Seedling density was higher in ULF (0.96 seedlings/ m2) than in the L-02 and L-12 stands (0.322 and 0.466 seedlings/m2, respectively). However, seedling mortality was higher in the ULF stands (54%) than in the L-02 (26%) and L-12 (15%) stands. P. macroloba in L-02 was the only species with abundant regeneration under P. copaifera in L-02 stand, where it accounted for 35% of the seedlings. Despite the reduction in seedling abundance observed after logging, P. copaifera seems to maintain large seedling populations in these forests, suggesting that this species maintains its dominance after logging disturbances. Our findings challenge the hypothesis that the regeneration of monodominant species is not likely to occur under heavily disturbed canopy conditions.

  10. Seasonal and cumulative loblolly pine development under two stand density and fertility levels

    Treesearch

    James D. Haywood

    1992-01-01

    An 8 year-old loblolly pine (Pinus taeda L.) stand was subjected to two cultural treatments for examination of seasonal and cumulative pine development. In the first treatment, pine density was either reduced by removal cutting to 2% trees per acre, at a 12- by 124 spacing, or left uncut with an original density of 1,210 trees per acre at a 6- by 6-...

  11. CARBON ISOTOPE DISCRIMINATION AND GROWTH RESPONSE OF OLD PINUS PONDEROSA TREES TO STAND DENSITY REDUCTIONS

    EPA Science Inventory

    Stand density reductions have been proposed as a method by which old-growth ponderosa pine (Pinus ponderosa) forests of North America can be converted back to pre-1900 conditions, thereby reducing the danger of catastrophic forest fires and insect attacks while increasing product...

  12. [Effects of litterfall and root input on soil physical and chemical properties in Pinus massoniana plantations in Three Gorges Reservoir Area, China].

    PubMed

    Ge, Xiao-Gai; Huang, Zhi-Lin; Cheng, Rui-Mei; Zeng, Li-Xiong; Xiao, Wen-Fa; Tan, Ben-Wang

    2012-12-01

    An investigation was made on the soil physical and chemical properties in different-aged Pinus massoniana plantations in Three Gorges Reservoir Area under effects of litterfall and roots. The annual litter production in mature stand was 19.4% and 65.7% higher than that in nearly mature and middle-aged stands, respectively. The litter standing amount was in the sequence of mature stand > middle-aged stand > nearly mature stand, while the litter turnover coefficient was in the order of nearly mature stand (0.51) > mature stand (0.40) > middle-aged stand (0.36). The total root biomass, live root biomass, and dead root biomass were the highest in middle-aged stand, and the lowest in nearly mature stand. In middle-aged stand, soil total porosity was the highest, and soil bulk density was the lowest. Soil organic matter and total nitrogen contents were in the order of mature stand > middle-aged stand > nearly mature stand, soil nitrate nitrogen occupied a larger proportion of soil mineral N in nearly mature stand, while ammonium nitrogen accounted more in middle-aged and mature stands. In nearly mature stand, litter production was moderate but turnover coefficient was the highest, and soil nutrient contents were the lowest. In middle-aged stand, root biomass and soil total porosity were the highest, and soil bulk density were the lowest. In mature stand, root biomass was lower while soil nutrient contents were the highest. The increase of root biomass could improve soil physical properties.

  13. Thinning increases growth of 60-year-old cherry-maple stands in West Virginia

    Treesearch

    Neil I. Lamson; Neil I. Lamson

    1985-01-01

    In north-central West Virginia, previously unmanaged 60-year-old cherrymaple stands were thinned to 60 percent relative stand density. Thinning reduced mortality, redistributed growth onto fewer, larger stems, and increased individual tree growth. Five-year periodic basal-area growth per acre was 1.2 times greater in thinned stands than in unthinned stands. Periodic...

  14. Differences in Fine-Root Biomass of Trees and Understory Vegetation among Stand Types in Subtropical Forests

    PubMed Central

    Fu, Xiaoli; Wang, Jianlei; Di, Yuebao; Wang, Huimin

    2015-01-01

    Variation of total fine-root biomass among types of tree stands has previously been attributed to the characteristics of the stand layers. The effects of the understory vegetation on total fine-root biomass are less well studied. We examined the variation of total fine-root biomass in subtropical tree stands at two sites of Datian and Huitong in China. The two sites have similar humid monsoon climate but different soil organic carbon. One examination compared two categories of basal areas (high vs. low basal area) in stands of single species. A second examination compared single-species and mixed stands with comparable basal areas. Low basal area did not correlate with low total fine-root biomass in the single-species stands. The increase in seedling density but decrease in stem density for the low basal area stands at Datian and the quite similar stand structures for the basal-area contrast at Huitong helped in the lack of association between basal area and total fine-root biomass at the two sites, respectively. The mixed stands also did not yield higher total fine-root biomasses. In addition to the lack of niche complementarity between tree species, the differences in stem and seedling densities and the belowground competition between the tree and non-tree species also contributed to the similarity of the total fine-root biomasses in the mixed and single-species stands. Across stand types, the more fertile site Datian yielded higher tree, non-tree and total fine-root biomasses than Huitong. However, the contribution of non-tree fine-root biomass to the total fine-root biomass was higher at Huitong (29.4%) than that at Datian (16.7%). This study suggests that the variation of total fine-root biomass across stand types not only was associated with the characteristics of trees, but also may be highly dependent on the understory layer. PMID:26047358

  15. Differences in Fine-Root Biomass of Trees and Understory Vegetation among Stand Types in Subtropical Forests.

    PubMed

    Fu, Xiaoli; Wang, Jianlei; Di, Yuebao; Wang, Huimin

    2015-01-01

    Variation of total fine-root biomass among types of tree stands has previously been attributed to the characteristics of the stand layers. The effects of the understory vegetation on total fine-root biomass are less well studied. We examined the variation of total fine-root biomass in subtropical tree stands at two sites of Datian and Huitong in China. The two sites have similar humid monsoon climate but different soil organic carbon. One examination compared two categories of basal areas (high vs. low basal area) in stands of single species. A second examination compared single-species and mixed stands with comparable basal areas. Low basal area did not correlate with low total fine-root biomass in the single-species stands. The increase in seedling density but decrease in stem density for the low basal area stands at Datian and the quite similar stand structures for the basal-area contrast at Huitong helped in the lack of association between basal area and total fine-root biomass at the two sites, respectively. The mixed stands also did not yield higher total fine-root biomasses. In addition to the lack of niche complementarity between tree species, the differences in stem and seedling densities and the belowground competition between the tree and non-tree species also contributed to the similarity of the total fine-root biomasses in the mixed and single-species stands. Across stand types, the more fertile site Datian yielded higher tree, non-tree and total fine-root biomasses than Huitong. However, the contribution of non-tree fine-root biomass to the total fine-root biomass was higher at Huitong (29.4%) than that at Datian (16.7%). This study suggests that the variation of total fine-root biomass across stand types not only was associated with the characteristics of trees, but also may be highly dependent on the understory layer.

  16. Long-term effects of clearcutting on tree species composition in an oak-hickory forest

    Treesearch

    Jessica A. Yeagle; John W. Groninger

    2006-01-01

    In 1973, a silvicultural clearcut, with and without a post-harvest herbicide treatment, was performed on an upland oak-hickory forest in southern Illinois. Prior to harvest, permanent plots were established, and a survey was conducted to determine stand structure and composition. In 2003, a post-harvest survey was performed using the permanent plots. Relative density...

  17. 93 Years of stand density and land-use legacy research at the Coulter Ranch Study Site

    Treesearch

    Andrew J. Sanchez Meador; Margaret M. Moore

    2008-01-01

    In 1913, the Fort Valley Experimental Forest initiated an unprecedented case-study experiment to determine the effects of harvesting methods on tree regeneration and growth on a ponderosa pine-Gambel oak forest at Coulter Ranch in northern Arizona. The harvesting methods examined were seed-tree, group selection, and light selection. In addition, the effects of...

  18. Oak savanna restoration: Oak response to fire and thinning through 28 years

    Treesearch

    Ronald E. Masters; Jack R. Waymire

    2012-01-01

    We used a small plot study on Pushmataha Wildlife Management Area in southeast Oklahoma to determine the efficacy of fire frequency and thinning as management tools for restoration of oak savanna, oak woodlands, pine-bluestem woodlands, and pine savanna for application on a landscape scale. On selected experimental units, we initially reduced stand density to favor...

  19. Growth in relation to canopy light interception in a red pine (Pinus resinosa) thinning study

    Treesearch

    Beverly E. Law; Kurt H. Riitters; Lewis F. Ohmann

    1992-01-01

    Growth data from the most recent 5 years of a 40-year thinning study in an even-aged red pine (Pinus resinosa) forest in cutfoot sioux experimental forest, Minnesota, were used with intercepted photosynthetically active radiation (IPAR) data to determine the relationship between light interception and growth for a range ofstand densities. Stand basal...

  20. Effect of silvicultural practice and wood type on loblolly pine particleboard and medium density fiberboard properties

    Treesearch

    Todd F. Shupe; Chung Y. Hse; Elvin T. Choong; Leslie H. Groom

    1999-01-01

    he objective of this study was to determine the effect of five different silvicultural strategies and wood type on mechanical and physical properties of loblolly pine (Pinus taeda L.) particleboard and fiberboard. The furnish was prepared in an unconventional manner from innerwood and outerwood veneer for each stand. Modulus of rupture (MOR)...

  1. EFFECT OF A SAUSAGE OSCILLATION ON RADIO ZEBRA-PATTERN STRUCTURES IN A SOLAR FLARE

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Yu, Sijie; Yan, Yihua; Nakariakov, V. M., E-mail: sjyu@nao.cas.cn

    2016-07-20

    Sausage modes that are axisymmetric fast magnetoacoustic oscillations of solar coronal loops are characterized by variation of the plasma density and magnetic field, and hence cause time variations of the electron plasma frequency and cyclotron frequency. The latter parameters determine the condition for the double plasma resonance (DPR), which is responsible for the appearance of zebra-pattern (ZP) structures in time spectra of solar type IV radio bursts. We perform numerical simulations of standing and propagating sausage oscillations in a coronal loop modeled as a straight, field-aligned plasma slab, and determine the time variation of the DPR layer locations. Instant valuesmore » of the plasma density and magnetic field at the DPR layers allowed us to construct skeletons of the time variation of ZP stripes in radio spectra. In the presence of a sausage oscillation, the ZP structures are shown to have characteristic wiggles with the time period prescribed by the sausage oscillation. Standing and propagating sausage oscillations are found to have different signatures in ZP patterns. We conclude that ZP wiggles can be used for the detection of short-period sausage oscillations and the exploitation of their seismological potential.« less

  2. Estimating Stand Height and Tree Density in Pinus taeda plantations using in-situ data, airborne LiDAR and k-Nearest Neighbor Imputation.

    PubMed

    Silva, Carlos Alberto; Klauberg, Carine; Hudak, Andrew T; Vierling, Lee A; Liesenberg, Veraldo; Bernett, Luiz G; Scheraiber, Clewerson F; Schoeninger, Emerson R

    2018-01-01

    Accurate forest inventory is of great economic importance to optimize the entire supply chain management in pulp and paper companies. The aim of this study was to estimate stand dominate and mean heights (HD and HM) and tree density (TD) of Pinus taeda plantations located in South Brazil using in-situ measurements, airborne Light Detection and Ranging (LiDAR) data and the non- k-nearest neighbor (k-NN) imputation. Forest inventory attributes and LiDAR derived metrics were calculated at 53 regular sample plots and we used imputation models to retrieve the forest attributes at plot and landscape-levels. The best LiDAR-derived metrics to predict HD, HM and TD were H99TH, HSD, SKE and HMIN. The Imputation model using the selected metrics was more effective for retrieving height than tree density. The model coefficients of determination (adj.R2) and a root mean squared difference (RMSD) for HD, HM and TD were 0.90, 0.94, 0.38m and 6.99, 5.70, 12.92%, respectively. Our results show that LiDAR and k-NN imputation can be used to predict stand heights with high accuracy in Pinus taeda. However, furthers studies need to be realized to improve the accuracy prediction of TD and to evaluate and compare the cost of acquisition and processing of LiDAR data against the conventional inventory procedures.

  3. [Characteristics of main layer and regeneration layer of Haloxylon ammodendron plantations at different ages on the southern edge of the Gurbantunggut Desert, Northwest China].

    PubMed

    Chen, Qi Min; Luo, Qing Hong; Ning, Hu Sen; Zhao, Cheng Yi; Duan, Wen Biao

    2017-03-18

    The population structure characteristics, natural regeneration, and the influential factors of Haloxylon ammodendron plantations at six different stand ages on the southern edge of the Gurbantunggut Desert were studied. The results showed that H. ammodendron plantation at the stand age of 7 could naturally regenerate. At the stand age of 17, the densities of the seedlings (<30 cm height), saplings (30≤H<50 height), and small trees (≥50 cm height) reached optimal class, and the mean height and base diameter of the small tress reached 1.10 m and 1.91 cm, respectively. The parent trees in H. ammodendron plantation at the stand age of 20 grew best. The height of 35% individuals grew up to 2.50-3.00 m, and the basal stem diameter of 23.1% individuals grew up to 8.00-10.00 cm. The height and diameter growth of the parent trees in H. ammodendron plantation at the stand age of 33 apparently declined, but the regeneration ability by natural seed dispersal was still strong. The regeneration density of natural seed dispersal showed the greatest correlation with the available nitrogen content in 0-100 cm soil layer (0.87), followed by the soil rapidly available phosphorus content (0.84) and the soil water content (0.79). The soils with pH 8.1-8.6 did not limit the nutrient growth of the regeneration layer. In the main stand layer, the individual density of whole regeneration layer showed the greatest correlation with the biomass of the parent trees (0.77), while the density of regeneration layer of the small trees showed the greatest correlation with the planting density (0.71) and the age of the parent trees (0.70).

  4. Direct sampling for stand density index

    Treesearch

    Mark J. Ducey; Harry T. Valentine

    2008-01-01

    A direct method of estimating stand density index in the field, without complex calculations, would be useful in a variety of silvicultural situations. We present just such a method. The approach uses an ordinary prism or other angle gauge, but it involves deliberately "pushing the point" or, in some cases, "pulling the point." This adjusts the...

  5. Standing Waves in a Nonuniform Medium

    ERIC Educational Resources Information Center

    Gluck, Paul

    2011-01-01

    A recent note in this journal presented a demonstration of standing waves along a cord consisting of two parts having different material densities, showing different sized wavelengths in each part. A generalization of that experiment to a continuously varying linear mass density is to vibrate a strip of material with gradually varying width (mass…

  6. Influence of stand density on stem quality in pole-size northern hardwoods.

    Treesearch

    Richard M. Godman; David J. Books

    1971-01-01

    Relates the type and frequency of limbs and limb-related defects in the first two logs of five hardwood species to residual basal area 15 years after initial cutting. Also discusses other tree characteristics influenced by stand density and the applicability of present silvicultural guides to improve stem quality.

  7. Regeneration of a coastal pine (Pinus thunbergii Parl.) forest 11 years after thinning, Niigata, Japan.

    PubMed

    Zhu, Jiaojun; Gonda, Yutaka; Yu, Lizhong; Li, Fengqin; Yan, Qiaoling; Sun, Yirong

    2012-01-01

    To examine the effects of thinning intensity on wind vulnerability and regeneration in a coastal pine (Pinus thunbergii) forest, thinning with intensities of 20%, 30% and 50% was conducted in December 1997; there was an unthinned treatment as the control (total 8 stands). We re-measured the permanent sites to assess the regeneration characteristics 11 years after thinning. In the 50% thinned stand, seedlings aged from 2 to 10 years exhibited the highest pine seedling density and growth. The age composition ranged from 1-3 years with densities of 9.9 and 5.1 seedlings m(-2) in 30% and 20% thinned stands; only 1-year-old seedlings with a density of 6.1 seedlings m(-2) in the unthinned stand. Similar trends were found for the regeneration of broadleaved species such as Robinia pseudoacacia and Prunus serrulata. We speculate that the canopy openness and moss coverage contributed to the regeneration success in the 50% thinned stand, while the higher litter depth and lack of soil moisture induced the regeneration failure in the unthinned stand. The stands thinned at 20% or 30% were less favourable for pine regeneration than the stands thinned at 50%. Therefore, thinning with less than 30% canopy openness (20% and 30% thinned stands) should be avoided, and thinning at higher than 30% canopy openness (50% thinned stand, approximately 1500 stems ha(-1) at ages 40-50 years) is suggested for increasing regeneration in the coastal pine forest. The implications of thinning-based silviculture in the coastal pine forest management are also discussed. The ongoing development of the broadleaved seedlings calls for further observations.

  8. The selection system of silviculture in spruce-fir stands - procedures, early results, and comparisons with unmanaged stands

    Treesearch

    Robert M. Frank; Barton M. Blum

    1978-01-01

    Early results after 20 years of record keeping indicate that spruce-fir stands will respond to the selection system of silviculture. Stand quality is improved, species composition can be altered, diameter-class distribution approaches a stated goal, stand density is controlled, and yields are increased. Selection silviculture in spruce-fir can now be compared to early...

  9. Modelling available crown fuel for Pinus pinaster Ait. stands in the "Cazorla, Segura and Las Villas Natural Park" (Spain).

    PubMed

    Molina, Juan Ramón; Rodríguez y Silva, Francisco; Mérida, Enrique; Herrera, Miguel Ángel

    2014-11-01

    One of the main limiting aspects in the application of crown fire models at landscape scale has been the uncertainty derived to describe canopy fuel stratum. Available crown fuel and canopy bulk density are essential in order to simulate crown fire behaviour and are of potential use in the evaluation of silvicultural treatments. Currently, the more accurate approach to estimate these parameters is to develop allometric models from common stand inventory data. In this sense, maritime pine (Pinus pinaster Aiton) trees were destructively sampled in the South of the Iberian Peninsula, covering natural and artificial stands. Crown fine fuel was separated into size classes and allometric equations that estimate crown fuel load by biomass fractions were developed. Available crown fuel was determined according to the fuel load differences between un-burned and burned trees with similar characteristics. Taking our destructive post-fire inventory into account, available crown fuel was estimated as the sum of needles biomass, 87.63% of the twigs biomass and 62.79% of the fine branches biomass. In spite of the differences between natural and artificial stands, generic models explained 82% (needles biomass), 89% (crown fuel), 92% (available crown fuel) and 94% (canopy bulk density) of the observed variation. Inclusion of the fitted models in fire management decision-making can provide a decision support system for assessing the potential crown fire of different silvicultural alternatives. Copyright © 2014 Elsevier Ltd. All rights reserved.

  10. Timber Management of NASA's John C. Stennis Space Station

    NASA Technical Reports Server (NTRS)

    Carr, Hugh; Johnson, Gary; Smoot, James; Cohan, Tyrus; O'Connor, Tina

    2001-01-01

    John C. Stennis Space Center occupies over 48560 hectares or just over 120000 acres in southwestern Mississippi and southeastern Louisiana near border of the two states. This area includes the Stennis Fee Area, which is the Center itself and the Acoustic Buffer Zone (ABZ) surrounding the Center. The ABZ is owned by several government, public and private entities and managed accordingly. The Fee Area and ABZ include wetlands, pine woodlands and areas of mixed floral species. The included maps detail the delineation of land in and around the Stennis Space Center. Areas owned by the federal government are divided into three stands to facilitate timber management. The stands are classified by species, density and average size to determine several timber in management issues including the schedule for controlled burns, economic viability of stands, and the creation of low impact recreational areas such as scenic trails or bike paths.

  11. Which Bow Shock Theory, Gasdynamic or Magnetohydrodynamic, Better Explains CME Stand-off Distance Ratios from LASCO-C2 Observations ?

    NASA Astrophysics Data System (ADS)

    Lee, Jae-Ok; Moon, Y.-J.; Lee, Jin-Yi; Kim, R.-S.; Cho, K.-S.

    2017-03-01

    It is generally believed that fast coronal mass ejections (CMEs) can generate their associated shocks, which are characterized by faint structures ahead of CMEs in white-light coronagraph images. In this study, we examine whether the observational stand-off distance ratio, defined as the CME stand-off distance divided by its radius, can be explained by bow shock theories. Of 535 SOHO/LASCO CMEs (from 1996 to 2015) with speeds greater than 1000 km s-1 and angular widths wider than 60°, we select 18 limb CMEs with the following conditions: (1) their Alfvénic Mach numbers are greater than one under Mann’s magnetic field and Saito’s density distributions; and (2) the shock structures ahead of the CMEs are well identified. We determine observational CME stand-off distance ratios by using brightness profiles from LASCO-C2 observations. We compare our estimates with theoretical stand-off distance ratios from gasdynamic (GD) and magnetohydrodynamic (MHD) theories. The main results are as follows. Under the GD theory, 39% (7/18) of the CMEs are explained in the acceptable ranges of adiabatic gamma (γ) and CME geometry. Under the MHD theory, all the events are well explained when we consider quasi-parallel MHD shocks with γ = 5/3. When we use polarized brightness (pB) measurements for coronal density distributions, we also find similar results: 8% (1/12) under GD theory and 100% (12/12) under MHD theory. Our results demonstrate that the bow shock relationships based on MHD theory are more suitable than those based on GD theory for analyzing CME-driven shock signatures.

  12. Which Bow Shock Theory, Gasdynamic or Magnetohydrodynamic, Better Explains CME Stand-off Distance Ratios from LASCO-C2 Observations ?

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Lee, Jae-Ok; Moon, Y.-J.; Lee, Jin-Yi

    It is generally believed that fast coronal mass ejections (CMEs) can generate their associated shocks, which are characterized by faint structures ahead of CMEs in white-light coronagraph images. In this study, we examine whether the observational stand-off distance ratio, defined as the CME stand-off distance divided by its radius, can be explained by bow shock theories. Of 535 SOHO /LASCO CMEs (from 1996 to 2015) with speeds greater than 1000 km s{sup −1} and angular widths wider than 60°, we select 18 limb CMEs with the following conditions: (1) their Alfvénic Mach numbers are greater than one under Mann’s magneticmore » field and Saito’s density distributions; and (2) the shock structures ahead of the CMEs are well identified. We determine observational CME stand-off distance ratios by using brightness profiles from LASCO-C2 observations. We compare our estimates with theoretical stand-off distance ratios from gasdynamic (GD) and magnetohydrodynamic (MHD) theories. The main results are as follows. Under the GD theory, 39% (7/18) of the CMEs are explained in the acceptable ranges of adiabatic gamma ( γ ) and CME geometry. Under the MHD theory, all the events are well explained when we consider quasi-parallel MHD shocks with γ = 5/3. When we use polarized brightness (pB) measurements for coronal density distributions, we also find similar results: 8% (1/12) under GD theory and 100% (12/12) under MHD theory. Our results demonstrate that the bow shock relationships based on MHD theory are more suitable than those based on GD theory for analyzing CME-driven shock signatures.« less

  13. Standing Kink modes in three-dimensional coronal loops

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Pascoe, D. J.; De Moortel, I., E-mail: dpascoe@mcs.st-andrews.ac.uk

    2014-04-01

    So far, the straight flux tube model proposed by Edwin and Roberts is the most commonly used tool in practical coronal seismology, in particular, to infer values of the (coronal) magnetic field from observed, standing kink mode oscillations. In this paper, we compare the period predicted by this basic model with three-dimensional (3D) numerical simulations of standing kink mode oscillations, as the period is a crucial parameter in the seismological inversion to determine the magnetic field. We perform numerical simulations of standing kink modes in both straight and curved 3D coronal loops and consider excitation by internal and external drivers.more » The period of oscillation for the displacement of dense coronal loops is determined by the loop length and the kink speed, in agreement with the estimate based on analytical theory for straight flux tubes. For curved coronal loops embedded in a magnetic arcade and excited by an external driver, a secondary mode with a period determined by the loop length and external Alfvén speed is also present. When a low number of oscillations is considered, these two periods can result in a single, non-resolved (broad) peak in the power spectrum, particularly for low values of the density contrast for which the two periods will be relatively similar. In that case (and for this particular geometry), the presence of this additional mode would lead to ambiguous seismological estimates of the magnetic field strength.« less

  14. Quantitative analysis of American woodcock nest and brood habitat

    USGS Publications Warehouse

    Bourgeois, A.; Keppie, Daniel M.; Owen, Ray B.

    1977-01-01

    Sixteen nest and 19 brood sites of American woodcock (Philohela minoI) were examined in northern lower Michigan between 15 April and 15 June 1974 to determine habitat structure associated with these sites. Woodcock hens utilized young, second-growth forest stands which were similar in species composition for both nesting and brood rearing. A multi-varIate discriminant function analysis revealed a significant (P< 0.05) difference, however, in habitat structure. Nest habitat was characterized by lower tree density (2176 trees/ha) and basal area (8.6 m2/ha), by being close to forest openings (7 m) and by being situated on dry, relatively well drained sites. In contrast, woodcock broods were located in sites that had nearly twice the tree density (3934 trees/hal and basal area (16.5 m2/ha), was located over twice as far from forest openings (18 m) and generally occurred on damp sites, near (8 m) standing water. Importance of the habitat features to the species and possible management implications are discussed.

  15. Canopy transpiration for two Japanese cypress forests with contrasting structures

    NASA Astrophysics Data System (ADS)

    Tsuruta, K.; Komatsu, H.; Kume, T.; Shinohara, Y.; Otsuki, K.

    2012-12-01

    Canopy transpiration (EC) could have large variations among stands with different structures. To evaluate a difference in EC between stands with different structures for Japanese cypress, we observed EC using the sap flow technique in two stands with contrasting structures (age was 19 year and 99 year, mean diameter at breast height was 13.5 cm and 44.6 cm, stem density was 2100 trees ha-1 and 350 trees ha-1, respectively) for 5 months under the same meteorological condition. The mean stand sap flux density (JS) for measurement period and stand sapwood area (AS_stand) for the old stand (0.43 m3 m-2 day-1 and 15.2 m2 ha-1) were lower than those for the young stand (0.62 m3 m-2 day-1 and 20.4 m2 ha-1) by 31.1 % and 25.4 %, respectively. EC is calculated as a product of JS and AS_stand. Therefore the EC in the old stand was lower than that in the young stand by 50 %. We calculated the contribution of the reference JS for a given meteorological conditions (JSref) and the response of JS to the meteorological conditions (JSresp) in the two stands, and examined which is a primary factor for the difference of EC between the two studied stands. The JSresp for the young stand were not considerably different from that for the old stand, whereas JSref for the young stand was greater than that for the old stand. This indicates that JSref (not JSresp) was the primary cause for the difference of EC between the two stands. Further studies observing EC from stands with various structures are needed to generalize our conclusions.

  16. Yellowstone wolf (Canis lupus) denisty predicted by elk (Cervus elaphus) biomass

    USGS Publications Warehouse

    Mech, L. David; Barber-Meyer, Shannon

    2015-01-01

    The Northern Range (NR) of Yellowstone National Park (YNP) hosts a higher prey biomass density in the form of elk (Cervus elaphus L., 1758) than any other system of gray wolves (Canis lupus L., 1758) and prey reported. Therefore, it is important to determine whether that wolf–prey system fits a long-standing model relating wolf density to prey biomass. Using data from 2005 to 2012 after elk population fluctuations dampened 10 years subsequent to wolf reintroduction, we found that NR prey biomass predicted wolf density. This finding and the trajectory of the regression extend the validity of the model to prey densities 19% higher than previous data and suggest that the model would apply to wolf–prey systems of even higher prey biomass.

  17. Early Fluid and Protein Shifts in Men During Water Immersion

    NASA Technical Reports Server (NTRS)

    Hinghofer-Szalkay, H.; Harrison, M. H.; Greenleaf, J. E.

    1987-01-01

    High precision blood and plasma densitometry was used to measure transvascular fluid shifts during water immersion to the neck. Six men (28-49 years) undertook 30 min of standing immersion in water at 35.0 +/- 0.2 C; immersion was preceded by 30 min control standing in air at 28 +/- 1 C. Blood was sampled from an antecubital catheter for determination of Blood Density (BD), Plasma Density (PD), Haematocrit (Ht), total Plasma Protein Concentration (PPC), and Plasma Albumin Concentration (PAC). Compared to control, significant decreases (p less than 0.01) in all these measures were observed after 20 min immersion. At 30 min, plasma volume had increased by 11.0 +/- 2.8%; the average density of the fluid shifted from extravascular fluid into the vascular compartment was 1006.3 g/l; albumin moved with the fluid and its albumin concentration was about one-third of the plasma protein concentration during early immersion. These calculations are based on the assumption that the F-cell ratio remained unchanged. No changes in erythrocyte water content during immersion were found. Thus, immersion-induced haemodilution is probably accompanied by protein (mainly albumin) augmentation which accompanies the intra-vascular fluid shift.

  18. Wave propagation and noncollisional heating in neutral loop and helicon discharges

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Celik, Y.; Crintea, D. L.; Luggenhoelscher, D.

    2011-02-15

    Heating mechanisms in two types of magnetized low pressure rf (13.56 MHz) discharges are investigated: a helicon discharge and a neutral loop discharge. Radial B-dot probe measurements demonstrate that the neutral loop discharge is sustained by helicon waves as well. Axial B-dot probe measurements reveal standing wave and beat patterns depending on the dc magnetic field strength and plasma density. In modes showing a strong wave damping, the plasma refractive index attains values around 100, leading to electron-wave interactions. In strongly damped modes, the radial plasma density profiles are mainly determined by power absorption of the propagating helicon wave, whereasmore » in weakly damped modes, inductive coupling dominates. Furthermore, an azimuthal diamagnetic drift is identified. Measurements of the helicon wave phase demonstrate that initial plane wave fronts are bent during their axial propagation due to the inhomogeneous density profile. A developed analytical standing wave model including Landau damping reproduces very well the damping of the axial helicon wave field. This comparison underlines the theory whereupon Landau damping of electrons traveling along the field lines at speeds close to the helicon phase velocity is the main damping mechanism in both discharges.« less

  19. Does variable-density thinning increase wind damage in conifer stands on the Olympic Peninsula?

    Treesearch

    S.D. Roberts; C.A. Harrington; K.R. Buermeyer

    2007-01-01

    Silvicultural treatments designed to enhance stand structural diversity may result in increased wind damage. The ability to avoid conditions that might lead to excessive wind damage would benefit forest managers. We analyzed wind damage following implementation of a variable-density thinning at four sites on the Olympic National Forest in northwest Washington. The...

  20. Crown development: an index of stand density

    Treesearch

    John E. Krajicek; Kenneth A. Brinkman

    1957-01-01

    A study of crown characteristics of several hundred open-grown oak, hickory, and Norway spruce trees in Iowa revealed a high correlation between stem diameter and crown area. Consideration of this relationship led to the idea that perhaps here was a realistic way to measure and evaluate stand density or stocking. If, given unlimited growing space, trees of a certain...

  1. Growth and mortality of ponderosa pine poles thinned to various densities in the Blue Mountains of Oregon.

    Treesearch

    P.H. Cochran; James W. Barrett

    1995-01-01

    Growth and mortality in a ponderosa pine (Pinus ponderosa Dougl. ex Laws.) stand were investigated for 24 years. High mortality rates from mountain pine beetle (Dendroctonus ponderosae Hopkins) occurred on some plots where values for stand density index exceeded 140. Periodic annual increments for quadratic mean diameters...

  2. Effects of silviculture on neotropical migratory birds in central and southeastern oak pine forests

    Treesearch

    James G. Dickson; Frank R. Thompson; Richard N. Conner; Kathleen E. Franzreb

    1993-01-01

    Avian communities that are associated with forest habitat attributes are affected by silvicultural and other stand influences. Some species have specific habitat requirements, whereas others occupy a broad range of vegetative conditions. In general, bird species richness and density are positively related to stand foliage volume and diversity. Bird density and...

  3. A 3D stand generator for central Appalachian hardwood forests

    Treesearch

    Jingxin Wang; Yaoxiang Li; Gary W. Miller

    2002-01-01

    A 3-dimensional (3D) stand generator was developed for central Appalachian hardwood forests. It was designed for a harvesting simulator to examine the interactions of stand, harvest, and machine. The Component Object Model (COM) was used to design and implement the program. Input to the generator includes species composition, stand density, and spatial pattern. Output...

  4. 93 years of stand density and land-use legacy research at the Coulter Ranch Study Site (P-53)

    Treesearch

    Andrew J. Sanchez Meador; Margaret M. Moore

    2008-01-01

    In 1913, the Fort Valley Experimental Forest initiated an unprecedented case-study experiment to determine the effects of harvesting methods on tree regeneration and growth on a ponderosa pine-Gambel oak forest at Coulter Ranch in northern Arizona. The harvesting methods examined were seed-tree, group selection, and light selection. In addition, the effects of...

  5. Patch to landscape patterns in post fire recruitment of a serotinous conifer

    USGS Publications Warehouse

    Ne'eman, Gidi; Fotheringham, C.J.; Keeley, J.E.

    1999-01-01

    Obligate seeding species are highly specialized to fire disturbance and many conifers such as cypress, which are adapted to high intensity stand-replacing fires, have canopy seed banks stored in serotinous cones. Resilience of these trees to fire disturbance is a function of disturbance frequency and one focus of this study was to determine the effect of patch age on postfire recruitment. A second focus was to determine the extent to which fire induced a landscape level change in the location of the forest boundary. Prior to a fire in 1994, a large Cupressus sargentii forest was a mosaic landscape of different aged patches of nearly pure cypress bordered by chaparral. Patches less than 60 years of age were relatively dense with roughly one tree every 1-2 m2 but older patches had thinned to one tree every 3-15 m2. Older trees had substantially greater canopy cone crops but the stand level seed bank size was not significantly correlated with stand age. Fire-dependent obligate seeding species are sensitive to fire return interval because of potential changes in the size of seed banks - facing both a potential 'immaturity risk' and a 'senescence risk'. At our site, C. sargentii regeneration was substantial in stands as young as 20 years, suggesting that fire return interval would need to be shorter than this to pose any significant risk. Reduced seedling recruitment in stands nearly 100 years of age may indicate risk from senescence is greater, however, even the lowest density seedling recruitment was many times greater than the density of mature forests - thus this cypress would appear to be resilient to a wide range of fire return intervals. Changes in landscape patterning of forest and chaparral are unlikely except after fire. Factors that inhibit tree establishment within the shrubland, as well as factors that affect shrub establishment within the forest border likely affect the 'permeability' of this ecotone. After the 1994 fire this boundary appeared to be stable in that cypress recruited best within the shadow of burned canopies and cypress were weak invaders of adjacent shrublands.

  6. Comparison of the evapotranspiration and its components before and after thinning in Japanese cedar and Japanese cypress forest

    NASA Astrophysics Data System (ADS)

    Tateishi, Makiko; Xiang, Yang; Matsuda, Hiroki; Saito, Takami; Sun, Haotian; Otsuki, Kyoichi; Kasahara, Tamao; Onda, Yuichi

    2014-05-01

    Water source area of Japan is often covered by forest, and 40 % of forest cover is coniferous plantation. Thinning has become a major tool in the management of plantation in recent years, but its effects on water cycle and its components are yet to be evaluated well. In this study, we investigated the changes in evapotranspiration and its components, including stand transpiration and canopy interception loss, after thinning in 50 years old Japanese cedar and Japanese cypress plantation at Yayama experimental catchment in Fukuoka, Japan. We established study plot in each Japanese cedar and Japanese cypress stand. Sap flow measurement was conducted for evaluating stand transpiration in each plot. Through fall and stem flow were also monitored to estimate canopy interception loss. The experiments were conducted over two years. During the measurements, 50 % of trees were thinned randomly in entire catchment, which has an area of 2.98 ha. Stem density was changed from 3945 to 1977 trees per ha after thinning. The reduction of daily stand transpiration in the studied Japanese cedar and cypress stands after thinning were 31.6 % and 48.2 % under the same condition of microclimate, respectively. These values were comparable to the changes in total sapwood area, 34.2 % and 44.5 %, and sap flow density did not change after thinning. It implies that sapwood area is a primary determinant of stand transpiration. Canopy interception ratios were 27 % and 26 % for Japanese cedar and cypress before thinning, and the ratios decreased to 24 % and 21 % after thinning, respectively. Thus, we obtained the changes in annual evapotranspiration and its components at catchment scale by using observation and models. The changes in partitioning of evapotranspiration is also discussed. The evapotranspiration before and after thinning were also compared to water balance data in this study site.

  7. [Vertical variation in stoichiometric relationships of soil carbon, nitrogen and phosphorus in five forest types in the Maoershan region, Northeast China.

    PubMed

    Zhang, Tai Dong; Wang, Chuan Kuan; Zhang, Quan Zhi

    2017-10-01

    Five forests under diverse site conditions but under identical climate in the Maoershan region of Northeast China were sampled for measuring contents of soil carbon (C), nitrogen (N), and phosphorus (P), soil bulk density, and soil thickness by soil profile horizons. The stands included two plantations (i.e., Pinus koraiensis and Larix gmelinii plantations) and three broadleaved forests (i.e., Quercus mongolica stand, Populus davidiana Betula platyphylla mixed stand, and hardwood stand). Our aim was to examine vertical distribution of the content, density, and stoichio metry of soil C, N and P for the five forest types. The results showed that the contents and densities of soil C, N and P differed significantly among the forest types, with the maxima of the soil C and N at both O and A horizons occurring in the hardwood stand. The contents of C and N decreased significantly with increasing soil depth in all the stands. P content decreased significantly only in the broadleaved stands, and P content had no significant difference among different soil layers in the coniferous stands. The soil C/N at the A horizon, N/P at the O horizon, and the C/P at A and B horizons were significantly different among the forest types. The soil C and N linearly correlated significantly across all the forest types without significant differences in the slopes and intercepts, and the soil N and P, or the soil C and P correlated significantly only in the broadleaved stands. This result suggested that the C-N coupling relationship tended to converge across the forest types, and the N-P and C-P relationships varied with forest types.

  8. A comparison of litter production in young and old baldcypress (Taxodium distichum L.) stands at Caddo Lake, Texas

    USGS Publications Warehouse

    McCoy, John W.; Draugelis-Dale, Rassa O.; Keeland, Bobby D.; Darville, Roy

    2010-01-01

    Aboveground primary productivity for cypress forests was assessed from measurements of litter production in two age groups and in two hydrological regimes (standing water and free-flowing). Caddo Lake, located in northeast Texas on the Texas-Louisiana border, offered a unique study site since it is dominated by extensive stands composed entirely of Taxodium distichum (L.) Rich, (baldcypress) in different age groups. Young stands (approximately 100 years old) are found along the shoreline and on shallow flooded islands. Old stands (-150 to 300 years old) are found in deeper water where they were continuously flooded. Litter production over three years from October 1998 to September 2001 was measured. Litter consisting of leaves, twigs, bark, reproductive parts, and Tillandsia usneoides (L.) L. (Spanish moss) was collected monthly using 0.5 m2 floating traps. Tree diameters were measured within 200 m2 circular plots in each stand. The young stands supported densities greater than 2,000 stems/ha and a mean stand basal area of 72.3 m2/ha, whereas old stands supported lower densities of about 500 stems/ha but with a similar mean stand basal area of 73.3 m2/ha. There was a significant difference between old and young stands for overall yearly litter production, averaging about 670 g/m2/yr in the young stands and 460 g/m2/yr in the old stands. Leaves and twigs were significantly greater in the young stands, while reproductive parts were higher in old stands. Litter collections between years or hydrological regimes were not significantly different.

  9. Breeding birds in managed forests on public conservation lands in the Mississippi Alluvial Valley

    USGS Publications Warehouse

    Twedt, Daniel J.; Wilson, R. Randy

    2017-01-01

    Managers of public conservation lands in the Mississippi Alluvial Valley have implemented forest management strategies to improve bottomland hardwood habitat for target wildlife species. Through implementation of various silvicultural practices, forest managers have sought to attain forest structural conditions (e.g., canopy cover, basal area, etc.) within values postulated to benefit wildlife. We evaluated data from point count surveys of breeding birds on 180 silviculturally treated stands (1049 counts) that ranged from 1 to 20 years post-treatment and 134 control stands (676 counts) that had not been harvested for >20 years. Birds detected during 10-min counts were recorded within four distance classes and three time intervals. Avian diversity was greater on treated stands than on unharvested stands. Of 42 commonly detected species, six species including Prothonotary Warbler (Prothonotaria citrea) and Acadian Flycatcher (Empidonax virescens) were indicative of control stands. Similarly, six species including Indigo Bunting (Passerina cyanea) and Yellow-breasted Chat (Icteria virens) were indicative of treated stands. Using a removal model to assess probability of detection, we evaluated occupancy of bottomland forests at two spatial scales (stands and points within occupied stands). Wildlife-forestry treatment improved predictive models of species occupancy for 18 species. We found years post treatment (range = 1–20), total basal area, and overstory canopy were important species-specific predictors of occupancy, whereas variability in basal area was not. In addition, we used a removal model to estimate species-specific probability of availability for detection, and a distance model to estimate effective detection radius. We used these two estimated parameters to derive species densities and 95% confidence intervals for treated and unharvested stands. Avian densities differed between treated and control stands for 16 species, but only Common Yellowthroat (Geothlypis trichas) and Yellow-breasted Chat had greater densities on treated stands.

  10. Estimating canopy bulk density and canopy base height for conifer stands in the interior Western United States using the Forest Vegetation Simulator Fire and Fuels Extension.

    Treesearch

    Seth Ex; Frederick Smith; Tara Keyser; Stephanie Rebain

    2017-01-01

    The Forest Vegetation Simulator Fire and Fuels Extension (FFE-FVS) is often used to estimate canopy bulk density (CBD) and canopy base height (CBH), which are key indicators of crown fire hazard for conifer stands in the Western United States. Estimated CBD from FFE-FVS is calculated as the maximum 4 m running mean bulk density of predefined 0.3 m thick canopy layers (...

  11. BLM Density Management and Riparian Buffer Study: Establishment Report and Study Plan

    USGS Publications Warehouse

    Cissel, John H.; Anderson, P.D.; Olson, Deanna H.; Puettmann, Klaus; Berryman, Shanti; Chan, Samuel; Thompson, Charley

    2006-01-01

    The Bureau of Land Management (BLM), Pacific Northwest Research Station (PNW), U.S. Geological Survey (USGS), and Oregon State University (OSU) established the BLM Density Management and Riparian Buffer Study (DMS) in 1994 to demonstrate and test options for young stand management to meet Northwest Forest Plan objectives in western Oregon. The primary objectives of the DMS are to evaluate the effects of alternative forest density management treatments in young stands on the development of important late-successional forest habitat attributes and to assess the combined effects of density management and alternative riparian buffer widths on aquatic and riparian ecosystems. The DMS consists of three integrated studies: initial thinning, rethinning, and riparian buffer widths. The initial thinning study was installed in 50- to 80-year-old stands that had never been commercially thinned. Four stand treatments of 30-60 acres each were established at each of seven study sites: (1) unthinned control, (2) high density retention [120 trees per acre (TPA)], (3) moderate density retention (80 TPA), and (4) variable density retention (40-120 TPA). Small (1/4 to 1 acre in size) leave islands were included in all treatments except the control, and small patch cuts (1/4 to 1 acre in size) were included in the moderate and variable density treatments. An eighth site, Callahan Creek, contains a partial implementation of the study design. The rethinning study was installed in four 70- to 90-year-old stands that previously had been commercially thinned. Each study stand was split into two parts: one part as an untreated control and the other part as a rethinning (30-60 TPA). The riparian buffer study was nested within the moderate density retention treatment at each of the eight initial thinning study sites and two rethinning sites. Alternative riparian buffer widths included: (1) streamside retention (one tree canopy width, or 20-25 feet), (2) variable width (follows topographic and vegetative breaks, 50 feet slope distance minimum), (3) one full site-potential tree height (approximately 220 feet), and (4) two full tree heights (approximately 440 feet). A second round of density management manipulations are now being planned for implementation beginning in 2009. Stem density will be reduced in the high, moderate, and variable density treatments and most existing riparian buffers, leave islands, and patch cuts will remain in place. Remeasurement, data management, and analysis are ongoing for three long-term, core components of the DMS: vegetation, microclimate, and aquatic vertebrates. In addition, several short-term collaborative studies have been completed on these sites, including leave island effectiveness as refugia, treatment response of terrestrial and aquatic arthropods, and smaller-scale studies of fungal, lichen, and bryophyte community response. Additional collaborative studies are encouraged on DMS sites.

  12. Nesting ecology of boreal forest birds following a massive outbreak of spruce beetles

    USGS Publications Warehouse

    Matsuoka, S.M.; Handel, C.M.

    2007-01-01

    We studied breeding dark-eyed juncos (Junco hyemalis), yellow-rumped warblers (Dendroica coronata), and spruce-nesting birds from 1997 to 1998 among forests with different levels of spruce (Picea spp.) mortality following an outbreak of spruce beetles (Dendroctonus rufipennis) in Alaska, USA. We identified species using live and beetle-killed spruce for nest sites and monitored nests to determine how the outbreak influenced avian habitat selection and reproduction. We tested predictions that 1) nesting success of ground-nesting juncos would increase with spruce mortality due to proliferation of understory vegetation available to conceal nests from predators, 2) nesting success of canopy-nesting warblers would decrease with spruce mortality due to fewer live spruce in which to conceal nests, and 3) both species would alter nest-site selection in response to disturbance. Juncos did not benefit from changes in understory vegetation; nesting success in highly disturbed stands (46%) was comparable to that in undisturbed habitats throughout their range. In stands with low spruce mortality, nesting success of juncos was low (5%) and corresponded with high densities of red squirrels (Tamiasciurus hudsonicus). Yellow-rumped warblers nested exclusively in spruce, but success did not vary with spruce mortality. As disturbance increased, nesting warblers switched from selecting forest patches with high densities of live white spruce (Picea glauca) to patches with beetle-killed spruce. Warblers also placed nests in large-diameter live or beetle-killed spruce, depending on which was more abundant in the stand, with no differences in nesting success. Five of the 12 other species of spruce-nesting birds also used beetle-killed spruce as nest sites. Because beetle-killed spruce can remain standing for >50 years, even highly disturbed stands provide an important breeding resource for boreal forest birds. We recommend that boreal forest managers preserve uncut blocks of infested forest within managed forest landscapes and practice partial harvest of beetle-killed spruce rather than commercial clear-cutting of infested stands in order to sustain breeding bird populations until natural reforestation occurs. Because breeding densities do not always reflect fitness, assessing impacts of a massive natural disturbance should include measuring impacts of changes in vegetation on both reproductive success and predator–prey dynamics.

  13. Financial analysis of early stand treatments in southwest Oregon.

    Treesearch

    Helge Eng; K. Norman Johnson; Roger D. Fight

    1990-01-01

    Management guidelines for economically efficient early stand treatments were developed by identifying treatments that would maximize financial returns over the rotation for coast Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco var. menziesii) in southwest Oregon. Short rotations and low stand densities (trees per acre) gave...

  14. The effects of stand structure after thinning on the growth of an Allegheny hardwood stand

    Treesearch

    David A. Marquis; Richard L. Ernst

    1991-01-01

    A 50-year-old Allegheny hardwood stand in which the crown canopy had stratified into distinct species groups was thinned to 60% relative density leaving dramatically different stand structures and species composition. Treatments included combined thinning, thin from middle, thin from above, thin from below, and unthinned control. Individual tree growth was stimulated...

  15. New methods for estimating parameters of weibull functions to characterize future diameter distributions in forest stands

    Treesearch

    Quang V. Cao; Shanna M. McCarty

    2006-01-01

    Diameter distributions in a forest stand have been successfully characterized by use of the Weibull function. Of special interest are cases where parameters of a Weibull distribution that models a future stand are predicted, either directly or indirectly, from current stand density and dominant height. This study evaluated four methods of predicting the Weibull...

  16. Drought induced tree mortality and ensuing bark beetle outbreaks in southwestern pinyon-juniper woodlands

    Treesearch

    Michael J. Clifford; Monique E. Rocca; Robert Delph; Paulette L. Ford; Neil S. Cobb

    2008-01-01

    The current drought and ensuing bark beetle outbreaks during 2002 to 2004 in the Southwest have greatly increased tree mortality in pinyon-juniper woodlands. We studied causes and consequences of the drought-induced mortality. First, we tested the paradigm that high stand densities in pinyon-juniper woodlands would increase tree mortality. Stand densities did not...

  17. A density management diagram for even-aged Sierra Nevada mixed-conifer stands

    Treesearch

    James N. Long; John D. Shaw

    2012-01-01

    We have developed a density management diagram (DMD) for even-aged mixed-conifer stands in the Sierra Nevada Mountains using forest inventory and analysis (FIA) data. Analysis plots were drawn from FIA plots in California, southern Oregon, and western Nevada which included those conifer species associated with the mixed-conifer forest type. A total of 204 plots met the...

  18. Seasonal photosynthesis and water relations of juvenile loblolly pine relative to stand density and canopy position

    Treesearch

    Zhenmin Tang; Jim L. Chambers; Mary A. Sword Sayer; James P. Barnett

    2003-01-01

    To assess the effects of stand density and canopy environment on tree physiology, we measured gas exchange responses of the same needle age class of 16-year-old loblolly pines (Pinus taeda L.) in thinned (512 trees ha-1) and non-thinned treatment plots (2,863 trees ha-1) in central Louisiana....

  19. Effects of variable-density thinning on understory diversity and heterogeneity in young Douglas-fir forests.

    Treesearch

    Juliann E. Aukema; Andrew B. Carey

    2008-01-01

    Nine years after variable-density thinning (VDT) on the Forest Ecosystem Study, we examined low understory vegetation in 60 plots of eight stands (four pairs of VDT and control). We compared native, exotic, ruderal, and nonforest species richness among the stands. We used clustering, ordination, and indicator species analysis to look for distinctive patches of plant...

  20. Snow breakage in a pole-sized ponderosa pine plantation ... more damage at high stand-densities

    Treesearch

    Robert F. Powers; William W. Oliver

    1970-01-01

    Damage by snow breakage to pole-sized ponderosa pine (Pinus pondvosa Laws.) increased as stand density increased. In a plantation on the west slope of California's Sierra Nevada, the tallest trees were most often broken. Thinning in the sapling stage is recommended as a preventative measure in dense plantations subject to heavy snowfall.

  1. Required sample size for monitoring stand dynamics in strict forest reserves: a case study

    Treesearch

    Diego Van Den Meersschaut; Bart De Cuyper; Kris Vandekerkhove; Noel Lust

    2000-01-01

    Stand dynamics in European strict forest reserves are commonly monitored using inventory densities of 5 to 15 percent of the total surface. The assumption that these densities guarantee a representative image of certain parameters is critically analyzed in a case study for the parameters basal area and stem number. The required sample sizes for different accuracy and...

  2. The Influence of Stand Density and Structure on Growth of Northern Hardwoods in New England

    Treesearch

    Dale S. Solomon

    1977-01-01

    Growth of northern hardwoods over a 10-year period was studied in plots that were treated to produce residual densities of 40, 60, 80, and 100 square feet of basal area per acre with stand structures of 30, 45 and 60 percent sawtimber. Both diameter and basal area growth are tabulated by treatment and species.

  3. Snag density varies with intensity of timber harvest and human access

    Treesearch

    Michael J. Wisdom; Lisa J. Bate

    2008-01-01

    Many species of vertebrates depend on snags (standing dead trees) for persistence, and limited research suggests that snag density is lower in areas of intensive timber harvest and increased human access. While intensive timber harvest is one source of potential snag loss, ease of human access to forest stands may also facilitate loss via firewood cutting of snags....

  4. Current Status of Mimosa pigra L. Infestation in Peninsular Malaysia

    PubMed Central

    Mansor, Asyraf; Crawley, Micheal J.

    2011-01-01

    The status and distribution of Mimosa pigra L., a semi-aquatic invasive species in Peninsular Malaysia, were continuously assessed between 2004 and 2007. This assessment investigated its population stand density and related weed management activities. In total, 106 sites of 6 main habitat types i.e., construction site (CS), dam/ reservoir (DM), forest reserve (FR), plantation (PL), river bank/waterway (RB) and roadside (RD) were assessed, and 55 sites were recorded with M. pigra populations. A CS is the most likely habitat to be infested with M. pigra (16 out of 18 assessed sites have this weed), whereas none of the FR visited were found to harbour M. pigra. In terms of population stand density, 41 populations were in the low range of stand density (individual plant of ≤5 m−2), compared to only 9 populations in the high range of stand density (individual plant of >10 m−2). In general, the current impact of M. pigra infestation on natural habitats is relatively low, as its distribution is only confined to disturbed areas. However, continuous monitoring of this weed species is highly recommended, especially in the riparian zone and wetland habitats. PMID:24575208

  5. Thinning regimes and initial spacing for Eucalyptus plantations in Brazil.

    PubMed

    Ferraz Filho, Antonio C; Mola-Yudego, Blas; González-Olabarria, José R; Scolforo, José Roberto S

    2018-01-01

    This study focuses on the effects of different thinning regimes on clonal Eucalyptus plantations growth. Four different trials, planted in 1999 and located in Bahia and Espírito Santo States, were used. Aside from thinning, initial planting density, and post thinning fertilization application were also evaluated. Before canopy closure, and therefore before excessive competition between trees took place, it was found that stands planted under low densities (667 trees per hectare) presented a lower mortality proportion when compared to stand planted under higher densities (1111 trees per hectare). However, diameter growth prior to thinning operations was not statistically different between these two densities, presenting an overall mean of 4.9 cm/year. After canopy closure and the application of the thinning treatments, it was found that thinning regimes beginning early in the life of the stand and leaving a low number of residual trees presented the highest diameter and height growth. Unthinned treatments and thinning regimes late in the life of the stand (after 5.5 years), leaving a large number of residual trees presented the highest values of basal area production. The choice of the best thinning regime for Eucalyptus clonal material will vary according to the plantation objective.

  6. The resolved layer of a collisionless, high beta, supercritical, quasi-perpendicular shock wave. I - Rankine-Hugoniot geometry, currents, and stationarity

    NASA Technical Reports Server (NTRS)

    Scudder, J. D.; Aggson, T. L.; Mangeney, A.; Lacombe, C.; Harvey, C. C.

    1986-01-01

    Data collected by the ISEE dual-spacecraft mission (on November 7, 1977) on a slowly moving, supercritical, high-beta, quasi-perpendicular bow shock are presented, and the local geometry, spatial scales, and stationarity of this shock wave are assessed in a self-consistent Rankine-Hugoniot-constrained frame of reference. Included are spatial profiles of the ac and dc magnetic and electric fields, electron and proton fluid velocities, current densities, electron and proton number densities, temperatures, pressures, and partial densities of the reflected protons. The observed layer profile is shown to be nearly phase standing and one-dimensional in a Rankine-Hugoniot frame, empirically determined by the magnetofluid parameters outside the layer proper.

  7. DOE Office of Scientific and Technical Information (OSTI.GOV)

    Brook, I.M.

    Five Thalassia communities with high blade density (greater than 3,000 blades/m/sup 2/) were sampled by suction dredge in April 1973. Four sites were in south Biscayne Bay, and one was at Murray Key in the Everglades National Park on the southwest coast of Florida. Macrofaunal abundance ranged from 292 to 10,728 individuals/m/sup 2/. It is postulated that a high standing crop of seagrass may not be the primary determining factor in faunal abundance.

  8. Tall shrub dynamics in northern Minnesota aspen and conifer forests.

    Treesearch

    James C. Galogh; David F. Grigal

    1988-01-01

    Tall shrub dynamics were examined in upland stands in northern Minnesota. Mortality rates of shrub stems did not differ among the stands. Shrub stem regeneration did differ among the stands and was related to overstory characteristics, soil moisture, and soil nutrients. Stem density was regulated by annual regeneration.

  9. Management of spruce-fir in even-aged stands in the central Rocky Mountains

    Treesearch

    Robert R. Alexander; Carleton B. Edminster

    1980-01-01

    Potential production of Engelmann spruce and subalpine fir in the central Rocky Mountains is simulated for vario.us combinations of stand density, site quality, ages, and thinning schedules. Such estimates are needed to project future development of stands managed in different ways for various uses.

  10. Stand development and silviculture in bottomland hardwoods

    Treesearch

    J. Steven Meadows

    1993-01-01

    Silviculture for the production of high-quality timber in southern bottomland hardwood forests involves the application of environmentally sound practices in order to enhance the growth and quality of both individual trees and stands. To accomplish this purpose, silvicultural practices are typically used to regulate stand density, species composition, and stem quality...

  11. Optimum stand prescriptions for ponderosa pine

    Treesearch

    David W. Hann; J. Douglas Brodie; Kurt H. Riitters

    1983-01-01

    Two examples for a northern Arizona ponderosa pine stand illustrate the usefulness of dynamic programming in making silvicultural decisions. The first example analyzes the optimal planting density for bare land, while the second examines the optimal precommercial thinning intensity for a 43-year-old stand. Bot hexamples assume that the manager's primary objective...

  12. Stand dynamics of Douglas-fir 20 years after precommercial thinning and nitrogen fertilization on a poor-quality site

    Treesearch

    Richard E. Miller; Timothy B. Harrington; Harry W. Anderson

    2016-01-01

    Amendment of soil nutrients through fertilization is used to increase wood production of forest stands. Subsequent allocation of growth to individual trees and the resulting increase in stand volume and value, however, depend on stand density at the time of treatment. Our primary research question was: To what extent can volume growth per acre in precommercially...

  13. Basal area or stocking percent: which works best in controlling density in natural shortleaf pine stands

    Treesearch

    Ivan L. Sander

    1986-01-01

    Results from a shortleaf pine thinning study in Missouri show that continually thinning a stand to the same basal area will eventually create an understocked stand and reduce yields. Using stocking percent to control thinning intensity allows basal area to increase as stands get older. The best yield should occur when shortleaf pine is repeatedly thinned to 60 percent...

  14. 3D numerical modelling of a willow vegetated river/floodplain system

    NASA Astrophysics Data System (ADS)

    Wilson, C. A. M. E.; Yagci, O.; Rauch, H.-P.; Olsen, N. R. B.

    2006-07-01

    SummaryUsing a three-dimensional finite volume code with standard k- ɛ turbulence closure the hydraulic impact of willow stands ( Salix alba and Salix fragilis) on the velocity distribution was modelled. The additional hydraulic resistance of the willow stands was modelled separately to the bed resistance using a drag force term that was introduced into the Navier-Stokes equations. Two flood events of varying magnitude and stages of plant development were simulated using this approach. The river comprises an asymmetric compound channel with vegetated floodplain of reach length 170 m. The willow development has been monitored annually and this information was used to define the density of the willow stands (average number per m 2) and its variation as a function of stand height. During both flood events the willow stands were submerged and in pronation. The willow stands were modelled in bending as well as in their undisturbed vertical state. Modelling the willow stands as vertical or in bending was found to have a major impact on the computed velocity profiles. The impact of using a drag-force approach based on a non-uniform projected area distribution was found to be greater when the plants are modelled vertically than when the plants are modelled in high degrees of bending. In field studies involving flexible plants without leaves, the determination of the drag coefficient is of less importance compared to the need to quantify the degree by which plants are in pronation.

  15. Seasonal development of loblolly pine lateral roots in response to stand density and fertilization

    Treesearch

    M.A. Sword

    1998-01-01

    In 1989, two levels each of stand density and fertilization treatments were factorially established in a 9-year-old loblolly pine plantation on a P-deficient Gulf Coastal Plain site in Rapides Parish, Louisiana, USA. In 1995, a second thinning was conducted on the previously thinned plots and fertilizer was re-applied to the previously fertilized plots. The morphology...

  16. Yellow pine regeneration as a function of fire severity and post-burn stand structure in the southern Appalachian Mountains

    Treesearch

    Michael A. Jenkins; Robert N. Klein; Virginia L. McDaniel

    2011-01-01

    We used pre- and post-burn fire effects data from six prescribed burns to examine post-burn threshold effects of stand structure (understory density, overstory density, shrub cover, duff depth, and total fuel load) on the regeneration of yellow pine (Pinus subgenus Diploxylon) seedlings and cover of herbaceous vegetation in six prescribed-fire management units located...

  17. Growth of western larch after thinning from above and below to several density levels: 10-year results.

    Treesearch

    K.W. Seidel

    1980-01-01

    The 10-year growth of a 55-year-old, even-aged western larch (Larix occidentalis Nutt.) stand after it was thinned from above and below to a wide range of stocking levels was measured in eastern Oregon. Basal area and volume growth increased as stand density increased for both thinning methods. Despite heavy infestations of the larch casebearer...

  18. The density and distribution of Sitka spruce and western hemlock seedling banks in partially harvested stands in southeast Alaska

    Treesearch

    Louise S.Y. Levy; Robert L. Deal; John C. Tappeiner

    2010-01-01

    This study’s objective was to document and describe the current seedling bank of Sitka spruce (Picea sitchensis (Bong.) Carr.) and western hemlock (Tsuga heterophylla (Raf.) Sarg.) stands in southeast Alaska that were partially cut between 1900 and 1984. We investigated the following: (1) What are seedling bank densities? (2)...

  19. Growth and yield of western larch in response to several density levels and two thinning methods: 15-year results.

    Treesearch

    K.W. Seidel

    1986-01-01

    The 15-year growth response from a levels-of-growing-stock study in an even-aged western larch (Larix occidentalis Nutt.) stand, first thinned from above and below at age 55, was measured in northeastern Oregon. Basal area and volume growth increased with stand density for both thinning methods, whereas diameter growth decreased. Attacks of the...

  20. Timber Resources Inventory and Monitoring Joint Research Project

    NASA Technical Reports Server (NTRS)

    Hill, C. L.

    1985-01-01

    Primary objectives were to develop remote sensing analysis techniques for extracting forest related information from LANDSAT Multispectral Scanner (MMS) and Thematic Mapper data and to determine the extent to which International Paper Company information needs can be addressed with remote sensing information. The company actively manages 8.4 million acres of forest land. Traditionally, their forest inventories, updated on a three year cycle, are conducted through field surveys and aerial photography. The results reside in a digital forest data base containing 240 descriptive parameteres for individual forest stands. The information in the data base is used to develop seasonal and long range management strategies. Forest stand condition assessements (species composition, age, and density stratification) and identification of silvicultural activities (site preparation, planting, thinning, and harvest) are addressed.

  1. Association of unipedal standing time and bone mineral density in community-dwelling Japanese women.

    PubMed

    Sakai, A; Toba, N; Takeda, M; Suzuki, M; Abe, Y; Aoyagi, K; Nakamura, T

    2009-05-01

    Bone mineral density (BMD) and physical performance of the lower extremities decrease with age. In community-dwelling Japanese women, unipedal standing time, timed up and go test, and age are associated with BMD while in women aged 70 years and over, unipedal standing time is associated with BMD. The aim of this study was to clarify whether unipedal standing time is significantly associated with BMD in community-dwelling women. The subjects were 90 community-dwelling Japanese women aged 54.7 years. BMD of the second metacarpal bone was measured by computed X-ray densitometry. We measured unipedal standing time as well as timed up and go test to assess physical performance of the lower extremities. Unipedal standing time decreased with increased age. Timed up and go test significantly correlated with age. Low BMD was significantly associated with old age, short unipedal standing time, and long timed up and go test. Stepwise regression analysis revealed that age, unipedal standing time, and timed up and go test were significant factors associated with BMD. In 21 participants aged 70 years and over, body weight and unipedal standing time, but not age, were significantly associated with BMD. BMD and physical performance of the lower extremities decrease with older age. Unipedal standing time, timed up and go test, and age are associated with BMD in community-dwelling Japanese women. In women aged 70 years and over, unipedal standing time is significantly associated with BMD.

  2. Remote sensing of Myriophyllum spicatum L. in a shallow, eutrophic lake

    NASA Technical Reports Server (NTRS)

    Gustafson, T. D.; Adams, M. S.

    1973-01-01

    An aerial 35 mm system was used for the acquisition of vertical color and color infrared imagery of the submergent aquatic macrophytes of Lake Wingra, Wisconsin. A method of photographic interpretation of stem density classes is tested for its ability to make standing crop biomass estimates of Myriophyllum spicatum. The results of film image density analysis are significantly correlated with stem densities and standing crop biomass of Myriophyllum and with the biomass of Oedogonium mats. Photographic methods are contrasted with conventional harvest procedures for efficiency and accuracy.

  3. Carbon storage in old-growth forests of the Mid-Atlantic: toward better understanding the eastern forest carbon sink.

    PubMed

    McGarvey, Jennifer C; Thompson, Jonathan R; Epstein, Howard E; Shugart, Herman H

    2015-02-01

    Few old-growth stands remain in the matrix of secondary forests that dominates the eastern North American landscape. These remnant stands offer insight on the potential carbon (C) storage capacity of now-recovering secondary forests. We surveyed the remaining old-growth forests on sites characteristic of the general Mid-Atlantic United States and estimated the size of multiple components of forest C storage. Within and between old-growth stands, variability in C density is high and related to overstory tree species composition. The sites contain 219 ± 46 Mg C/ha (mean ± SD), including live and dead aboveground biomass, leaf litter, and the soil O horizon, with over 20% stored in downed wood and snags. Stands dominated by tulip poplar (Liriodendron tulipifera) store the most live biomass, while the mixed oak (Quercus spp.) stands overall store more dead wood. Total C density is 30% higher (154 Mg C/ha), and dead wood C density is 1800% higher (46 Mg C/ha) in the old-growth forests than in the surrounding younger forests (120 and 5 Mg C/ha, respectively). The high density of dead wood in old growth relative to secondary forests reflects a stark difference in historical land use and, possibly, the legacy of the local disturbance (e.g., disease) history. Our results demonstrate the potential for dead wood to maintain the sink capacity of secondary forests for many decades to come.

  4. Standing beat-to-beat blood pressure variability is reduced among fallers in the Malaysian Elders Longitudinal Study.

    PubMed

    Goh, Choon-Hian; Ng, Siew-Cheok; Kamaruzzaman, Shahrul Bahyah; Chin, Ai-Vyrn; Tan, Maw Pin

    2017-10-01

    The aim of this study was to determine the relationship between falls and beat-to-beat blood pressure (BP) variability.Continuous noninvasive BP measurement is as accurate as invasive techniques. We evaluated beat-to-beat supine and standing BP variability (BPV) using time and frequency domain analysis from noninvasive continuous BP recordings.A total of 1218 older adults were selected. Continuous BP recordings obtained were analyzed to determine standard deviation (SD) and root mean square of real variability (RMSRV) for time domain BPV and fast-Fourier transform low frequency (LF), high frequency (HF), total power spectral density (PSD), and LF:HF ratio for frequency domain BPV.Comparisons were performed between 256 (21%) individuals with at least 1 fall in the past 12 months and nonfallers. Fallers were significantly older (P = .007), more likely to be female (P = .006), and required a longer time to complete the Timed-Up and Go test (TUG) and frailty walk test (P ≤ .001). Standing systolic BPV (SBPV) was significantly lower in fallers compared to nonfallers (SBPV-SD, P = .016; SBPV-RMSRV, P = .033; SBPV-LF, P = .003; SBPV-total PSD, P = .012). Nonfallers had significantly higher supine to standing ratio (SSR) for SBPV-SD, SBPV-RMSRV, and SBPV-total PSD (P = .017, P = .013, and P = .009). In multivariate analyses, standing BPV remained significantly lower in fallers compared to nonfallers after adjustment for age, sex, diabetes, frailty walk, and supine systolic BP. The reduction in frequency-domain SSR among fallers was attenuated by supine systolic BP, TUG, and frailty walk.In conclusion, reduced beat-to-beat BPV while standing is independently associated with increased risk of falls. Changes between supine and standing BPV are confounded by supine BP and walking speed.

  5. Analyzing the management and disturbance in European forest based on self-thinning theory

    NASA Astrophysics Data System (ADS)

    Yan, Y.; Gielen, B.; Schelhaas, M.; Mohren, F.; Luyssaert, S.; Janssens, I. A.

    2012-04-01

    There is increasing awareness that natural and anthropogenic disturbance in forests affects exchange of CO2, H2O and energy between the ecosystem and the atmosphere. Consequently quantification of land use and disturbance intensity is one of the next steps needed to improve our understanding of the carbon cycle, its interactions with the atmosphere and its main drivers at local as well as at global level. The conventional NPP-based approaches to quantify the intensity of land management are limited because they lack a sound ecological basis. Here we apply a new way of characterising the degree of management and disturbance in forests using the self- thinning theory and observations of diameter at breast height and stand density. We used plot level information on dominant tree species, diameter at breast height, stand density and soil type from the French national forest inventory from 2005 to 2010. Stand density and diameter at breast height were used to parameterize the intercept of the self-thinning relationship and combined with theoretical slope to obtain an upper boundary for stand productivity given its density. Subsequently, we tested the sensitivity of the self-thinning relationship for tree species, soil type, climate and other environmental characteristics. We could find statistical differences in the self-thinning relationship between species and soil types, mainly due to the large uncertainty of the parameter estimates. Deviation from the theoretical self-thinning line defined as DBH=αN-3/4, was used as a proxy for disturbances, allowing to make spatially explicit maps of forest disturbance over France. The same framework was used to quantify the density-DBH trajectory of even-aged stand management of beech and oak over France. These trajectories will be used as a driver of forest management in the land surface model ORCHIDEE.

  6. High Energy Density All Solid State Asymmetric Pseudocapacitors Based on Free Standing Reduced Graphene Oxide-Co3O4 Composite Aerogel Electrodes.

    PubMed

    Ghosh, Debasis; Lim, Joonwon; Narayan, Rekha; Kim, Sang Ouk

    2016-08-31

    Modern flexible consumer electronics require efficient energy storage devices with flexible free-standing electrodes. We report a simple and cost-effective route to a graphene-based composite aerogel encapsulating metal oxide nanoparticles for high energy density, free-standing, binder-free flexible pseudocapacitive electrodes. Hydrothermally synthesized Co3O4 nanoparticles are successfully housed inside the microporous graphene aerogel network during the room temperature interfacial gelation at the Zn surface. The resultant three-dimensional (3D) rGO-Co3O4 composite aerogel shows mesoporous quasiparallel layer stack morphology with a high loading of Co3O4, which offers numerous channels for ion transport and a 3D interconnected network for high electrical conductivity. All solid state asymmetric pseudocapacitors employing the composite aerogel electrodes have demonstrated high areal energy density of 35.92 μWh/cm(2) and power density of 17.79 mW/cm(2) accompanied by excellent cycle life.

  7. Foraging and nesting habitat of breeding male northern goshawks in the laurentian mixed forest province, Minnesota

    USGS Publications Warehouse

    Boal, C.W.; Andersen, D.E.; Kennedy, P.L.

    2005-01-01

    We used radiotelemetry to examine foraging habitat preferences of 17 breeding, male northern goshawks (Accipiter gentilis) in Minnesota from 1998-2000. We assessed habitat preference using radio relocation points and 50-m radius buffers of radio relocation points. Our data suggested that foraging male goshawks used early-successional upland conifer stands (???25 yrs old), early-successional upland deciduous stands (???50 yrs old), late-successional upland conifer stands (???50 yrs old), and late-successional upland deciduous stands (???50 yrs old) more frequently than expected based on the abundance of these vegetation types in the landscape. The 2 most available stand types, early-successional upland deciduous (<25 yrs old) and all ages of late-successional lowland conifer stands, were used less than expected by foraging goshawks. Late-successional lowland deciduous stands (???50 yrs old) were used in proportion to availability. Although analysis of relocation points suggested early-successional upland deciduous stands (25-49 yrs old) and late-successional upland conifer stands (???50 yrs old) were used in proportion to availability, analysis of buffers around relocation points indicated that these stand types were also used more than expected by foraging goshawks. Regardless of vegetation community type, stands used by goshawks were structurally similar with high canopy and understory stem densities, high canopy closure, substantial shrub cover, and large amounts of woody debris. Nest stands consisted of taller and larger diameter canopy trees and fewer understory trees than foraging stands, but stands were otherwise similar in structural features, suggesting goshawks used similar stands for nesting and foraging but that they tended to select the most mature stands for nesting. A commonality among nesting and foraging stands was the presence of open spaces between the canopy and understory foliage, and between understory and shrub layer foliage. In our study area, these spaces may have served as relatively unobstructed flight paths where foraging and nesting stands possessed stem densities at the upper end of that reported for goshawk habitat.

  8. Does initial spacing influence crown and hydraulic architecture of Eucalyptus marginata?

    PubMed

    Grigg, A H; Macfarlane, C; Evangelista, C; Eamus, D; Adams, M A

    2008-05-01

    Long-term declines in rainfall in south-western Australia have resulted in increased interest in the hydraulic characteristics of jarrah (Eucalyptus marginata Donn ex Smith) forest established in the region's drinking water catchments on rehabilitated bauxite mining sites. We hypothesized that in jarrah forest established on rehabilitated mine sites: (1) leaf area index (L) is independent of initial tree spacing; and (2) more densely planted trees have less leaf area for the same leaf mass, or the same sapwood area, and have denser sapwood. Initial stand densities ranged from about 600 to 9000 stems ha(-1), and trees were 18 years old at the time of sampling. Leaf area index was unaffected by initial stand density, except in the most sparsely stocked stands where L was 1.2 compared with 2.0-2.5 in stands at other spacings. The ratio of leaf area to sapwood area (A(l):A(s)) was unaffected by tree spacing or tree size and was 0.2 at 1.3 m height and 0.25 at the crown base. There were small increases in sapwood density and decreases in leaf specific area with increased spacing. Tree diameter or basal area was a better predictor of leaf area than sapwood area. At the stand scale, basal area was a good predictor of L (r(2) = 0.98, n = 15) except in the densest stands. We conclude that the hydraulic attributes of this forest type are largely independent of initial tree spacing, thus simplifying parameterization of stand and catchment water balance models.

  9. Field experience silvicultural cleaning project in young spruce and fir stands in central Nova Scotia

    Treesearch

    Theodore C. Tryon; Thomas W. Hartranft

    1977-01-01

    Silvicultural cleaning production varied from .15 to .34 acres per man day using light weight chain saws in young Spruce and Fir stands in Central Nova Scotia. Direct labor and saw costs, in cleaning young softwood stands in Nova Scotia, can be expected to range generally from $55.00 to $90.00 per acre, depending on crew experience, stand density, and equipment used....

  10. Response of six non-native invasive plant species to wildfires in the northern Rocky Mountains, USA

    Treesearch

    Dennis E. Ferguson; Christine L. Craig

    2010-01-01

    This paper presents early results on the response of six non-native invasive plant species to eight wildfires on six National Forests (NFs) in the northern Rocky Mountains, USA. Stratified random sampling was used to choose 224 stands based on burn severity, habitat type series, slope steepness, stand height, and stand density. Data for this report are from 219 stands...

  11. Long-term development of regeneration under longleaf pine seedtree and shelterwood stands

    Treesearch

    William D. Boyer

    1993-01-01

    Well-stocked mature longleaf pine (Pinus palustris Mill.) stands were cut tofive residual basal areas in 1957, namely 9,18,2 7.36, and 45 ft2 per ac, to observe the effect of stand density on seed production and seedling establishment. Seedlings, mainly from the 195.5 or 1961 seed crops, were established in treated stands. All pines on net 0.9 ac...

  12. Growth and bole quality responses to thinning in a red oak-sweetgum stand in southeastern Arkansas: nine-year results

    Treesearch

    James S. Meadows

    2012-01-01

    Science-based guidelines for thinning in southern bottomland hardwood stands are inadequate. To address this need, we established a series of thinning studies based on stand density management in hardwood stands on minor streambottom sites across the South. In the third study in this series, four thinning treatments were applied to a poletimber-sized, red oak-sweetgum...

  13. Structure of Pine Stands in the Southeast

    Treesearch

    William A. Bechtold; Gregory A. Ruark

    1988-01-01

    Distributional and statistical information associated with stand age, site index, basal area per acre, number of stems per acre, and stand density index is reported for major pine cover types of the Southeastern United States. Means, standard deviations, and ranges of these variables are listed by State and physiographic region for loblolly, slash, longleaf, pond,...

  14. Growth response of managed uneven-aged northern conifer stands

    Treesearch

    Dale S. Solomon; Robert M. Frank

    1983-01-01

    The growth response of trees in spruce-fir-hemlock stands was recorded from plots that were managed to control stand density, species composition, length of harvest interval, and salvage of mortality. Basal area, volume, and diameter increment are presented by species and size classification for harvesting intervals of 5, 10, and 20 years.

  15. Stereo photo guide for estimating canopy fuel characteristics in conifer stands

    Treesearch

    Joe H. Scott; Elizabeth D. Reinhardt

    2005-01-01

    Stereo photographs, hemispherical photographs, and stand data are presented with associated biomass and canopy fuel characteristics for five Interior West conifer stands. Canopy bulk density, canopy base height, canopy biomass by component, available canopy fuel load, and vertical distribution of canopy fuel are presented for each plot at several stages of sampling,...

  16. Black bear (Ursus americanus Pallas) feeding damage across timber harvest edges in northern California coast redwood (Sequoia sempervirens[D. Don] Endl.) forests, USA

    USGS Publications Warehouse

    Russell, W.H.; Carnell, K.; McBride, J.R.

    2001-01-01

    Feeding damage to trees by black bears (Ursus americanus Pallas) was recorded in proximity to timber harvest edges in harvested and old-growth stands of coast redwood (Sequoia sempervirens [D. Don] Endl.) in northern California, USA. Bears exhibited distinct preference in their feeding patterns related to stand structure and composition and to distance from the timber-harvest edge. Most damage was recorded within regenerating stands. Regression analysis indicated that density of damaged trees was negatively correlated with distance from timber harvest edges within old-growth stands. A significant negative correlation was also found between the density of trees damaged by bears and habitat diversity (H') as measured by the Shannon diversity index. In addition, bears exhibited preference for pole-size trees (dbh = 10-50 cm) over all other size classes, and coast redwood over other species. In general, damage by bears appeared to act as a natural thinning agent in even-aged stands. No damage was recorded in old-growth stands except in close proximity to the timber-harvest edge where subcanopy recruitment was high.

  17. Growth and development of ponderosa pine on sites of contrasting productivities: relative importance of stand density and shrub competition effects

    Treesearch

    Jianwei Zhang; William W. Oliver; Matt D. Busse

    2006-01-01

    Effects of stand density and shrub competition on growth and development were compared across a gradient of study sites. Challenge, the most productive site, is located in the foothills of the Sierra Nevada, northern California. Pringle Falls is of intermediate productivity in the rain shadow of the central Oregon Cascades. Trough Springs Ridge is the poorest site...

  18. Growth and yield of western larch: 15-year results of a levels-of-growing-stock study.

    Treesearch

    K.W. Seidel

    1982-01-01

    The 15-year growth response from a levels-of-growing-stock study in an even-aged western larch (Larix occidentalis Nutt.) stand in northeastern Oregon, first thinned at age 33, showed that trees growing at lower stand densities grew more rapidly in diameter but did not grow faster in height than trees in high density plots. Both basal area and...

  19. Changes in tree density do not influence epicormic branching of yellow-poplar

    Treesearch

    H. Clay Smith

    1977-01-01

    Epicormic branching was studied in a West Virginia yellow-poplar stand thinned to various tree density levels. Study trees in the 55- to 60-year-old second-growth stand were primarily codominant in crown class with 32 to 48 feet of log height. Eight-year study results indicated that yellow-poplar trees in this age class and locale could be thinned without serious loss...

  20. Approximation of the breast height diameter distribution of two-cohort stands by mixture models III Kernel density estimators vs mixture models

    Treesearch

    Rafal Podlaski; Francis A. Roesch

    2014-01-01

    Two-component mixtures of either the Weibull distribution or the gamma distribution and the kernel density estimator were used for describing the diameter at breast height (dbh) empirical distributions of two-cohort stands. The data consisted of study plots from the Å wietokrzyski National Park (central Poland) and areas close to and including the North Carolina section...

  1. Do understorey or overstorey traits drive tree encroachment on a drained raised bog?

    PubMed

    Jagodziński, A M; Horodecki, P; Rawlik, K; Dyderski, M K

    2017-07-01

    One of the most important threats to peatland ecosystems is drainage, resulting in encroachment of woody species. Our main aim was to check which features - overstorey or understorey vegetation - are more important for shaping the seedling bank of pioneer trees colonising peatlands (Pinus sylvestris and Betula pubescens). We hypothesised that tree stand parameters will be more important predictors of natural regeneration density than understorey vegetation parameters, and the former will be negatively correlated with species diversity and richness and also with functional richness and functional dispersion, which indicate a high level of habitat filtering. The study was conducted in the 'Zielone Bagna' nature reserve (NW Poland). We assessed the structure of tree stands and natural regeneration (of B. pubescens and P. sylvestris) and vegetation species composition. Random forest and DCA were applied to assess relationships between variables studied. Understorey vegetation traits affected tree seedling density (up to 0.5-m height) more than tree stand traits. Density of older seedlings depended more on tree stand traits. We did not find statistically significant relationships between natural regeneration densities and functional diversity components, except for functional richness, which was positively correlated with density of the youngest tree seedlings. Seedling densities were higher in plots with lower functional dispersion and functional divergence, which indicated that habitat filtering is more important than competition. Presence of an abundant seedling bank is crucial for the process of woody species encroachment on drained peatlands, thus its dynamics should be monitored in protected areas. © 2017 German Botanical Society and The Royal Botanical Society of the Netherlands.

  2. Influence of Crown Biomass Estimators and Distribution on Canopy Fuel Characteristics in Ponderosa Pine Stands of the Black Hills

    Treesearch

    Tara Keyser; Frederick Smith

    2009-01-01

    Two determinants of crown fire hazard are canopy bulk density (CBD) and canopy base height (CBH). The Fire and Fuels Extension to the Forest Vegetation Simulator (FFE-FVS) is a model that predicts CBD and CBH. Currently, FFE-FVS accounts for neither geographic variation in tree allometries nor the nonuniform distribution of crown mass when one is estimating CBH and CBD...

  3. Predicting breeding bird occurrence by stand- and microhabitat-scale features in even-aged stands in the Central Appalachians

    USGS Publications Warehouse

    McDermott, M.E.; Wood, P.B.; Miller, G.W.; Simpson, B.T.

    2011-01-01

    Spatial scale is an important consideration when managing forest wildlife habitat, and models can be used to improve our understanding of these habitats at relevant scales. Our objectives were to determine whether stand- or microhabitat-scale variables better predicted bird metrics (diversity, species presence, and abundance) and to examine breeding bird response to clearcut size and age in a highly forested landscape. In 2004-2007, vegetation data were collected from 62 even-aged stands that were 3.6-34.6. ha in size and harvested in 1963-1990 on the Monongahela National Forest, WV, USA. In 2005-2007, we also surveyed birds at vegetation plots. We used classification and regression trees to model breeding bird habitat use with a suite of stand and microhabitat variables. Among stand variables, elevation, stand age, and stand size were most commonly retained as important variables in guild and species models. Among microhabitat variables, medium-sized tree density and tree species diversity most commonly predicted bird presence or abundance. Early successional and generalist bird presence, abundance, and diversity were better predicted by microhabitat variables than stand variables. Thus, more intensive field sampling may be required to predict habitat use for these species, and management may be needed at a finer scale. Conversely, stand-level variables had greater utility in predicting late-successional species occurrence and abundance; thus management decisions and modeling at this scale may be suitable in areas with a uniform landscape, such as our study area. Our study suggests that late-successional breeding bird diversity can be maximized long-term by including harvests >10. ha in size into our study area and by increasing tree diversity. Some harvesting will need to be incorporated regularly, because after 15 years, the study stands did not provide habitat for most early successional breeding specialists. ?? 2010 Elsevier B.V.

  4. Estimating the Longwave Radiation Underneath the Forest Canopy in Snow-dominated Setting

    NASA Astrophysics Data System (ADS)

    Zhou, Y.; Kumar, M.; Link, T. E.

    2017-12-01

    Forest canopies alter incoming longwave radiation at the land surface, thus influencing snow cover energetics. The snow surface receives longwave radiation from the sky as well as from surrounding vegetation. The longwave radiation from trees is determined by its skin temperature, which shows significant heterogeneity depending on its position and morphometric attributes. Here our goal is to derive an effective tree temperature that can be used to estimate the longwave radiation received by the land surface pixel. To this end, we implement these three steps: 1) derive a relation between tree trunk surface temperature and the incident longwave radiation, shortwave radiation, and air temperature; 2) develop an inverse model to calculate the effective temperature by establishing a relationship between the effective temperature and the actual tree temperature; and 3) estimate the effective temperature using widely measured variables, such as solar radiation and forest density. Data used to derive aforementioned relations were obtained at the University of Idaho Experimental Forest, in northern Idaho. Tree skin temperature, incoming longwave radiation, solar radiation received by the tree surface, and air temperature were measured at an isolated tree and a tree within a homogeneous forest stand. Longwave radiation received by the land surface and the sky view factors were also measured at the same two locations. The calculated effective temperature was then compared with the measured tree trunk surface temperature. Additional longwave radiation measurements with pyrgeometer arrays were conducted under forests with different densities to evaluate the relationship between effective temperature and forest density. Our preliminary results show that when exposed to direct shortwave radiation, the tree surface temperature shows a significant difference from the air temperature. Under cloudy or shaded conditions, the tree surface temperature closely follows the air temperature. The effective tree temperature follows the air temperature in a dense forest stand, although it is significantly larger than the air temperature near the isolated tree. This discrepancy motivates us to explore ways to represent the effective tree temperature for stands with different densities.

  5. Rehabilitation of Understocked Loblolly-Shortleaf Pine Stands - IV. Natural and Planted Seedling/Sapling Stands

    Treesearch

    James B. Baker; Michael G. Shelton

    1998-01-01

    A 3- to 6 yr-old naturally regenerated, even-aged loblollypine (Pinus taeda L.) stand and a 5- yr-old loblolly pine plantation on good sites (SIbb = 85 to 90 ft at 50 yr ) were cut to density levels of 50, 90, 180,270, and 360 seedlings and/or saplings/ac. Two pine release treatments (none and individual tree release with a herbicide) were applied to the natural stand...

  6. The effect of low-magnitude whole body vibration on bone density and microstructure in men and women with chronic motor complete paraplegia

    PubMed Central

    Wuermser, Lisa-Ann; Beck, Lisa A.; Lamb, Jeffry L.; Atkinson, Elizabeth J.; Amin, Shreyasee

    2015-01-01

    Objective To examine the effect of low-magnitude whole body vibration on bone density and microstructure in women and men with chronic motor complete paraplegia. Methods We studied nine subjects (four women and five men) with motor complete paraplegia of 2 years duration or more, age 20–50 years. Subjects were instructed to stand on a low-magnitude vibration plate within a standing frame for 20 minutes per day, 5 days a week, and for 6 months. Bone density at the proximal femur by dual-energy X-ray absorptiometry and bone microstructure at the distal tibia by high-resolution peripheral quantitative computed tomography were assessed at four timepoints over 12 months (baseline, at 3 months and 6 months while on intervention, and after 6 months off intervention). Results Standing on the low-magnitude vibration plate with a standing frame was well tolerated by participants. However, most subjects did not show an improvement in bone density or microstructure after 6 months of intervention, or any relevant changes 6 months following the discontinuation of the low-magnitude vibration. Conclusion We were unable to identify an improvement in either bone density or microstructure following 6 months use of a low-magnitude vibration plate in women or men with chronic motor complete paraplegia. Longer duration of use may be necessary, or it is possible that this intervention is of limited benefit following chronic spinal cord injury. PMID:24621040

  7. The effect of low-magnitude whole body vibration on bone density and microstructure in men and women with chronic motor complete paraplegia.

    PubMed

    Wuermser, Lisa-Ann; Beck, Lisa A; Lamb, Jeffry L; Atkinson, Elizabeth J; Amin, Shreyasee

    2015-03-01

    To examine the effect of low-magnitude whole body vibration on bone density and microstructure in women and men with chronic motor complete paraplegia. We studied nine subjects (four women and five men) with motor complete paraplegia of 2 years duration or more, age 20-50 years. Subjects were instructed to stand on a low-magnitude vibration plate within a standing frame for 20 minutes per day, 5 days a week, and for 6 months. Bone density at the proximal femur by dual-energy X-ray absorptiometry and bone microstructure at the distal tibia by high-resolution peripheral quantitative computed tomography were assessed at four timepoints over 12 months (baseline, at 3 months and 6 months while on intervention, and after 6 months off intervention). Standing on the low-magnitude vibration plate with a standing frame was well tolerated by participants. However, most subjects did not show an improvement in bone density or microstructure after 6 months of intervention, or any relevant changes 6 months following the discontinuation of the low-magnitude vibration. We were unable to identify an improvement in either bone density or microstructure following 6 months use of a low-magnitude vibration plate in women or men with chronic motor complete paraplegia. Longer duration of use may be necessary, or it is possible that this intervention is of limited benefit following chronic spinal cord injury.

  8. Effect of thinning on height and diameter growth of oak & yellow-poplar saplings

    Treesearch

    Rufus H., Jr. Allen; David A. Marquis; David A. Marquis

    1970-01-01

    Studying the response to thinning of a 7- to 9-year-old upland hardwood sapling stand, we found that height growth of yellow-poplar and oak trees was markedly reduced by heavy thinning. This suggests that stand density should be carefully controlled to achieve maximum benefit from thinnings in very young stands.

  9. Eccentricity and fluting in young–growth western hemlock in Oregon.

    Treesearch

    Ryan Singleton; Dean S. DeBell; David D. Marshall; Barbara L. Gartner

    2004-01-01

    Stem irregularities can influence estimates of tree and stand attributes, efficiency of manufacturing processes, and quality of wood products. Eccentricity and fluting were characterized in young, managed western hemlock stands in the Oregon Coast Range. Sixty-one trees were selected from pure western hemlock stands across a range of age, site, and densities. The trees...

  10. Vegetative composition in forested areas following application of desired forest condition treatments

    Treesearch

    Trent A. Danley; Andrew W. Ezell; Emily B. Schultz; John D. Hodges

    2015-01-01

    Desired forest conditions, or DFCs, are recently created parameters which strive to create diverse stands of hardwoods of various species and age classes, along with varying densities and canopy gaps, through the use of uneven-aged silvicultural methods and repeated stand entries. Little research has been conducted to examine residual stand composition and hardwood...

  11. Effects of deer exclosures on oak regeneration in closed-canopy stands

    Treesearch

    Angela M. Yuska; Kim C. Steiner; James C. Finley

    2008-01-01

    Studies of the effects of high deer densities on forest regeneration have shown altered species composition and reduced diversity in stands regenerating after harvest. The effects of browsing in fully stocked, undisturbed stands are less well known but important, as establishment of seedlings of oaks and other species prior to disturbance is very important for self-...

  12. Alteration Of Nutrient Status By Manipulation Of Composition And Density In A Shortleaf Pine-Hardwood Stand

    Treesearch

    Hal O. Liechty; Valerie L. Sawyer; Michael G. Shelton

    2002-01-01

    Abstract - Uneven-aged management is used to promote adequate pine reproduction and control species composition of shortleaf pine (Pinus echinata Mill.)-hardwood stands in the Interior Highlands of the southern United States. The modification of pine-hardwood composition in these stands has the potential to alter nutrient pools and availability since...

  13. Stand dynamics of mixed red alder-conifer forests of southeast Alaska.

    Treesearch

    Robert L. Deal; Paul E. Hennon; Ewa H. Orlikowska; David V. D' Amore

    2004-01-01

    Stand structure and dynamics were evaluated in mixed red alder (Alnus rubra Bong.) - conifer forests of southeast Alaska. We assessed stand development, tree density, total basal area, diameter distribution of live and dead trees, height distribution of live trees, and mean diameter of all and largest conifers in 40-year-old red alder - conifer...

  14. Residue distribution and biomass recovery following biomass harvest of plantation pine

    Treesearch

    Johnny Grace III; John Klepac; S. Taylor; Dana Mitchell

    2016-01-01

    Forest biomass is anticipated to play a significant role in addressing an alternative energy supply. However, the efficiencies of current state-of-the-art recovery systems operating in forest biomass harvests are still relatively unknown. Forest biomass harvest stands typically have higher stand densities and smaller diameter trees than conventional stands which may...

  15. Performance of a Tracked Feller-Buncher with a Shear Head Operating in Small-Diameter Pine

    Treesearch

    J. Klepac

    2013-01-01

    A Tigercat 845D tracked feller-buncher equipped with a shear head was evaluated while performing a clearcut in a 15-year old Loblolly pine (Pinus taeda) plantation and a 18-year old natural stand. Mean density of the plantation was 573 TPA (Trees per Acre) while the natural stand averaged 328 TPA, with a slightly higher density of 390 TPA in the study area. Total cycle...

  16. Leaf area and structural changes after thinning in even-aged Picea rubens and Abies balsamea stands in Maine, USA

    Treesearch

    R. Justin DeRose; Robert S. Seymour

    2012-01-01

    We tested the hypothesis that changes in leaf area index (LAIm2 m-2) and mean stand diameter following thinning are due to thinning type and residual density. The ratios of pre- to postthinning diameter and LAI were used to assess structural changes between replicated crown, dominant, and low thinning treatments to 33% and 50% residual density in even-aged Picea rubens...

  17. Investigation on effect of Populus alba stands distance on density of pests and their natural enemies population under poplar/alfalfa agroforestry system.

    PubMed

    Khabir, Z H; Sadeghi, S E; Hanifeh, S; Eivazi, A

    2009-01-15

    This study was carried out in order to distinguish the effect of agroforestry system (combination of agriculture and forestry) on pests and natural enemy's population in poplar research station. Wood is one of the first substances that naturally was used for a long period of time. Forage is an important production of natural resources too. Some factors such as proper lands deficit, lack of economy, pest and disease attacks and faced production of these materials with serious challenges. Agroforestry is a method for decrease of the mentioned problems. The stands of poplar had have planted by complete randomized design with 4 treatments (stand distance) of poplar/alfalfa include 3x4, 3x6.7, 3x8, 3x10 m and 2 control treatments, alfalfa and poplar. The results showed that Chaitophorus populeti had the highest density in poplar and 3x10 m treatments. Monosteira unicostata is another insect pest that had most density in 3x10 m treatment. And alfalfa had high density of Chrysoperla carnea. The density of Coccinella septempunctata, were almost equal in all treatments.

  18. The Role of Remote Sensing in Assessing Forest Biomass in Appalachian South Carolina

    NASA Technical Reports Server (NTRS)

    Shain, W.; Nix, L.

    1982-01-01

    Information is presented on the use of color infrared aerial photographs and ground sampling methods to quantify standing forest biomass in Appalachian South Carolina. Local tree biomass equations are given and subsequent evaluation of stand density and size classes using remote sensing methods is presented. Methods of terrain analysis, environmental hazard rating, and subsequent determination of accessibility of forest biomass are discussed. Computer-based statistical analyses are used to expand individual cover-type specific ground sample data to area-wide cover type inventory figures based on aerial photographic interpretation and area measurement. Forest biomass data are presented for the study area in terms of discriminant size classes, merchantability limits, accessibility (as related to terrain and yield/harvest constraints), and potential environmental impact of harvest.

  19. Unipedal standing exercise and hip bone mineral density in postmenopausal women: a randomized controlled trial.

    PubMed

    Sakai, Akinori; Oshige, Toshihisa; Zenke, Yukichi; Yamanaka, Yoshiaki; Nagaishi, Hitoshi; Nakamura, Toshitaka

    2010-01-01

    The aim of this study was to test the effect of unipedal standing exercise on bone mineral density (BMD) of the hip in postmenopausal women. Japanese postmenopausal women (n = 94) were assigned at random to an exercise or control group (no exercise). The 6-month exercise program consisted of standing on a single foot for 1 min per leg 3 times per day. BMD of the hip was measured by dual-energy X-ray absorptiometry. There was no significant difference in age and baseline hip BMD between the exercise group (n = 49) and control group (n = 45). Exercise did not improve hip BMD compared with the control group. Stepwise regression analysis identified old age as a significant determinant (p = 0.034) of increased hip total BMD at 6 months after exercise. In 31 participants aged >/=70 years, the exercise group (n = 20) showed significant increase in the values of hip BMD at the areas of total (p = 0.008), intertrochanteric (p = 0.023), and Ward's triangle (p = 0.032). The same parameters were decreased in the control group (n = 11). The percent changes in hip BMD of the exercise group were not significantly different from those of the control group either in the participants with low baseline hip total BMD (<80% of the young adult mean) or high baseline hip total BMD (> or =80% of the young adult mean). In conclusion, unipedal standing exercise for 6 months did not improve hip BMD in Japanese postmenopausal women. Effect of exercise on hip total BMD was age dependent. In participants aged > or =70 years, the exercise significantly increased hip total BMD.

  20. Efficacy of "Verbenone Plus" for protecting ponderosa pine trees and stands from Dendroctonus brevicomis (Coleoptera: Curculionidae) attack in British Columbia and California.

    PubMed

    Fettig, Christopher J; McKelvey, Stephen R; Dabney, Christopher P; Huber, Dezene P W; Lait, Cameron G; Fowler, Donald L; Borden, John H

    2012-10-01

    The western pine beetle, Dendroctonus brevicomis LeConte (Coleoptera: Curculionidae, Scolytinae), is a major cause of ponderosa pine, Pinus ponderosa Douglas ex Lawson, mortality in much of western North America. We review several years of research that led to the identification of Verbenone Plus, a novel four-component semiochemcial blend [acetophenone, (E)-2-hexen-1-ol + (Z)-2-hexen-1-ol, and (-)-verbenone] that inhibits the response of D. brevicomis to attractant-baited traps, and examine the efficacy of Verbenone Plus for protecting individual trees and forest stands from D. brevicomis infestations in British Columbia and California. In all experiments, semiochemicals were stapled around the bole of treated trees at approximately equal to 2 m in height. (-)-Verbenone alone had no effect on the density of total attacks and successful attacks by D. brevicomis on attractant-baited P. ponderosa, but significantly increased the percentage of pitchouts (unsuccessful D. brevicomis attacks). Verbenone Plus significantly reduced the density of D. brevicomis total attacks and D. brevicomis successful attacks on individual trees. A significantly higher percentage of pitchouts occurred on Verbenone Plus-treated trees. The application of Verbenone Plus to attractant-baited P. ponderosa significantly reduced levels of tree mortality. In stand protection studies, Verbenone Plus significantly reduced the percentage of trees mass attacked by D. brevicomis in one study, but in a second study no significant treatment effect was observed. Future research should concentrate on determining optimal release rates and spacings of release devices in stand protection studies, and expansion of Verbenone Plus into other systems where verbenone alone has not provided adequate levels of tree protection.

  1. Bioimpedance for the spot measurement of tissue density

    NASA Astrophysics Data System (ADS)

    Dylke, E. S.; Ward, L. C.; Stannard, C.; Leigh, A.; Kilbreath, S. L.

    2013-04-01

    Long-standing lymphoedema is characterised by tissues changes which are currently not detectable using bioimpedance spectroscopy. It has been suggested that a combination of bipolar and tetrapolar measurements may be used to detect these tissues changes for a single site in the transverse direction. This was technique was trialled in a group of control participants with no history of lymphoedema or recent upper limb trauma. Repeated spot measurements were done without removal of electrodes to determine biological variability as well as with removal of electrodes to determine technical reproducibility. The inter-limb spot ratio of the controls was then compared to that of a number of women previously diagnosed with secondary lymphoedema in the forearm. Biological variability was not found to greatly influence repeated measures but only moderate technical reliability was found despite excellent co-efficient of variation for the majority of the measurements. A difference was seen between those with more severe swelling and the controls. This novel technique shows promise in detecting tissue changes associated with long-standing lymphoedema.

  2. Experimental investigation of standing wave effect in dual-frequency capacitively coupled argon discharges: role of a low-frequency source

    NASA Astrophysics Data System (ADS)

    Zhao, Kai; Liu, Yong-Xin; Kawamura, E.; Wen, De-Qi; Lieberman, M. A.; Wang, You-Nian

    2018-05-01

    It is well known that the plasma non-uniformity caused by the standing wave effect has brought about great challenges for plasma material processing. To improve the plasma uniformity, a low-frequency (LF) power source is introduced into a 100 MHz very-high-frequency (VHF) capacitively coupled argon plasma reactor. The effect of the LF parameters (LF voltage amplitude ϕ L and LF source f L) on the radial profile of plasma density has been investigated by utilizing a hairpin probe. The result at a low pressure (1 Pa) is compared to the one obtained by a 2D fluid-analytical capacitively coupled plasma model, showing good agreement in the plasma density radial profile. The experimental results show that the plasma density profile exhibits different dependences on ϕ L and f L at different gas pressures/electrode driven types (i.e., the two rf sources are applied on one electrode (case I) and separate electrodes (case II)). At low pressures (e.g., 8 Pa), the pronounced standing wave effect revealed in a VHF discharge can be suppressed at a relatively high ϕ L or a low f L in case I, because the HF sheath heating is largely weakened due to strong modulation by the LF source. By contrast, ϕ L and f L play insignificant roles in suppressing the standing wave effect in case II. At high pressures (e.g., 20 Pa), the opposite is true. The plasma density radial profile is more sensitive to ϕ L and f L in case II than in case I. In case II, the standing wave effect is surprisingly enhanced with increasing ϕ L at higher pressures; however, the center-high density profile caused by the standing wave effect can be compensated by increasing f L due to the enhanced electrostatic edge effect dominated by the LF source. In contrast, the density radial profile shows a much weaker dependence on ϕ L and f L in case I at higher pressures. To understand the different roles of ϕ L and f L, the electron excitation dynamics in each case are analyzed based on the measured spatio-temporal distributions of the electron-impact excitation rate by phase resolved optical emission spectroscopy.

  3. Climate and species functional traits influence maximum live tree stocking in the Lake States, USA

    Treesearch

    Mark J. Ducey; Christopher W. Woodall; Andrés Bravo-Oviedo

    2017-01-01

    Quantifying the density of live trees in forest stands and partitioning it between species or other stand components is critical for predicting forest dynamics and responses to management, as well as understanding the impacts of stand composition and structure on productivity. As plant traits such as shade tolerance have been proven to refine understanding of plant...

  4. Crop-tree release thinning in 65-year-old commercial cherry-maple stands (5-year results)

    Treesearch

    H. Clay Smith; Gary W. Miller; Neil I. Lamson

    1994-01-01

    Crop-tree release was applied to a 65-year-old cherry-maple stand in north central West Virginia. Criteria were developed for selecting crop trees for high quality sawtimber and veneer products. Five-year stand growth, mortality, and ingrowth using basal areas, volume, relative density, and number of trees were discussed for the treatments.

  5. The role of silvicultural thinning in eastern forests threatened by hemlock woolly adelgid (Adelges tsugae)

    Treesearch

    Mary Ann Fajvan

    2008-01-01

    In order to increase hemlock survivability in stands threatened by hemlock woolly adelgid (HWA), a new study is developing silvicultural thinning guidelines to reduce stand densities, reallocate resources, and increase hemlock vigor across a range of stand types and structures before HWA invasion. The 7 study areas are all geographically similar in that they regularly...

  6. Silvicultural potential for pre-comercial treatment in northern forest type

    Treesearch

    H. W.,Jr Hocker

    1977-01-01

    It is proposed that pre-commercial thinning of young northern hardwood, oak, white pine and spruce-fir stands be carried out using appropriate stocking guides to regulate stand density. Thinning should be carried out when stands are-between 1" and 2" dbh. Pruning of eastern white pine is recommended, while pruning of spruce and yellow birch seems feasible,...

  7. The densest loblolly pine stand and its silvicultural implications

    Treesearch

    Boris Zeide; John Stephens

    2010-01-01

    Estimation of stand density index has been based on the assumption that the only cause of mortality in fully stocked stands is diameter growth. For example, when average diameter increases by 1 percent, a fixed proportion (1.6 percent) of trees must die, regardless of age, average tree size, and other factors. This balance between growth and mortality entails the...

  8. Effect of stand densities on stand dynamics in white fir (Abies concolor) forests in northeast California, USA

    Treesearch

    Jianwei Zhang; William W. Oliver; Martin W. Ritchie

    2007-01-01

    Six white fir trials were established during the last half-century to develop guidelines for managing the existing natural stands across northeastern California. We analyzed data collected from these trials at about 5-year intervals since their installation. Because these trials were independent of each other, the design and treatments varied among the sites....

  9. Effect of a Spaced Thinning in Mature Lodgepole Pine on Within-Stand Microclimate and Fine Fuel Moisture Content

    Treesearch

    R. J. Whitehead; G. L. Russo; B. C. Hawkes; S. W. Taylor; B. N. Brown; H. J. Barclay; R. A. Benton

    2006-01-01

    Thinning mature forest stands to wide spacing is prescribed to reduce crown bulk density and likelihood of severe crown fire behaviour. However, it may adversely affect surface fuel load, moisture content and within-stand wind, which influence surface fire behaviour and crowning potential. Comparison of a mature lodgepole pine (Pinus contorta Dougl....

  10. Stocking levels and underlying assumptions for uneven-aged Ponderosa Pine stands.

    Treesearch

    P.H. Cochran

    1992-01-01

    Potential Problems With Q-Values Many ponderosa pine stands have a limited number of size classes, and it may be desirable to carry very large trees through several cutting cycles. Large numbers of trees below commercial size are not needed to provide adequate numbers of future replacement trees. Under these conditions, application of stand density index (SDI) can have...

  11. Growth and yield of all-aged Douglas-fir -- western hemlock forest stands: a matrix model with stand diversity effects.

    Treesearch

    Jingjing Liang; Joseph Buonglorno; Robert A. Monserud

    2005-01-01

    A density-dependent matrix model was developed for Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) -- western hemlock (Tsuga heterophylla (Raf.) Sarg.) forest stands in the Pacific Northwest of the United States. The model predicted the number and volume of trees for 4 species groups and 19 diameter classes. The parameters...

  12. Characteristics of Five Climax Stands in New Hampshire

    Treesearch

    W.B. Leak; W.B. Leak

    1987-01-01

    Analysis of species composition, stand density, and diameter distribution in five climax or old-growth stands in New Hampshire indicates that the northern hardwood climax is characterized by at least 65 to 70 percent tolerant hardwoods, 130 square feet basal area per acre, and "q" ratios (2-inch diameter classes) of 1.3 to 1.4. Climax sprucehemlock have at...

  13. Thinning cherry-maple stands in West Virginia: 5-year results

    Treesearch

    Neil I. Lamson; H. Clay. Smith; H. Clay. Smith

    1988-01-01

    In northern West Virginia, 60-year-old cherry-maple stands were thinned to 75,60, and 45 percent relative stand density. Analysis of 5-year growth data showed that basal-area growth was not reduced by thinning. Cubic-foot and board-foot volume growth decreased slightly. Individual-tree growth of all trees, dominant/codominant trees, and the 50 largest diameter trees...

  14. Light intensity related to stand density in mature stands of the western white pine type

    Treesearch

    C. A. Wellner

    1948-01-01

    Where tolerance of forest trees or subordinate vegetation is a factor in management, the forester needs a simple field method of Estimating or forecasting light intensities in forest stands. The following article describes a method developed for estimating light intensity beneath the canopy in western white pine forests which may have application in other types.

  15. Relating P-band AIRSAR backscatter to forest stand parameters

    NASA Technical Reports Server (NTRS)

    Wang, Yong; Melack, John M.; Davis, Frank W.; Kasischke, Eric S.; Christensen, Norman L., Jr.

    1993-01-01

    As part of research on forest ecosystems, the Jet Propulsion Laboratory (JPL) and collaborating research teams have conducted multi-season airborne synthetic aperture radar (AIRSAR) experiments in three forest ecosystems including temperate pine forest (Duke, Forest, North Carolina), boreal forest (Bonanza Creek Experimental Forest, Alaska), and northern mixed hardwood-conifer forest (Michigan Biological Station, Michigan). The major research goals were to improve understanding of the relationships between radar backscatter and phenological variables (e.g. stand density, tree size, etc.), to improve radar backscatter models of tree canopy properties, and to develop a radar-based scheme for monitoring forest phenological changes. In September 1989, AIRSAR backscatter data were acquired over the Duke Forest. As the aboveground biomass of the loblolly pine forest stands at Duke Forest increased, the SAR backscatter at C-, L-, and P-bands increased and saturated at different biomass levels for the C-band, L-band, and P-band data. We only use the P-band backscatter data and ground measurements here to study the relationships between the backscatter and stand density, the backscatter and mean trunk dbh (diameter at breast height) of trees in the stands, and the backscatter and stand basal area.

  16. Hydrological states and the resilience of deltaic forested wetlands

    NASA Astrophysics Data System (ADS)

    Keim, R.; Allen, S. T.

    2017-12-01

    The flooding regime constitutes a set of chronic disturbances that are largely responsible for ecosystem structure. However, disturbances do not always constitute stresses to plants that survive because of adaptations to flooded conditions. We examine baldcypress-water tupelo forested wetlands in the delta of the Mississippi River as a case study in mechanisms by which hydrologic change shapes wetland ecosystem change, supported by experimental evidence from remote sensing, tree-ring and other field studies, and meta-analysis across the literature. Decreased hydrologic variability caused by water control structures has reduced the frequency of flood events that increase growth of baldcypress and favor its establishment by reducing competition from other species. Hydrologic modifications that lead to semi-permanent, stagnant flooding constitute semi-permanent disturbance that prevents regeneration of any trees, reduces growth of established trees, and reduces stand density by causing mortality of some trees. However, baldcypress trees in low-density stands appear to be generally adapted for long-term survival in stagnant conditions. Thus, initial decreases in stand density after impoundment do not necessarily portend continued conversion away from forest because reduced inter-tree competition is a negative feedback on mortality. Overall, a natural hydrologic regime with high variability in riverine flooding favors denser stands with greater diversity of tree species, and the present, controlled hydrologic regime that has largely eliminated riverine flooding favors open stands. Sea-level rise will increase salinity that quickly leads to forest conversion to marsh, but will also increase stagnant, freshwater flooding further inland. These drivers of hydrologic change reduce carbon assimilation by forests, both by reduced stand-level productivity and decreased forested area.

  17. Estimating Evapotranspiration Of Orange Orchards Using Surface Renewal And Remote Sensing Techniques

    NASA Astrophysics Data System (ADS)

    Consoli, S.; Russo, A.; Snyder, R.

    2006-08-01

    Surface renewal (SR) analysis was utilized to calculate sensible heat flux density from high frequency temperature measurements above orange orchard canopies during 2005 in eastern Sicily (Italy). The H values were employed to estimate latent heat flux density (LE) using measured net radiation (Rn) and soil heat flux density (G) in the energy balance (EB) equation. Crop coefficients were determined by calculating the ratio Kc=ETa/ETo, with reference ETo derived from the daily Penman-Monteith equation. The estimated daily Kc values showed an average of about 0.75 for canopy covers having about 70% ground shading and 80% of PAR light interception. Remote sensing estimates of Kc and ET fluxes were compared with those measured by SR-EB. IKONOS satellite estimates of Kc and NDVI were linearly correlated for the orchard stands.

  18. Scaling up and error analysis of transpiration for Populus euphratica in a desert riparian forest

    NASA Astrophysics Data System (ADS)

    Si, J.; Li, W.; Feng, Q.

    2013-12-01

    Water consumption information of the forest stand is the most important factor for regional water resources management. However, water consumption of individual trees are usually measured based on the limited sample trees , so, it is an important issue how to realize eventual scaling up of data from a series of sample trees to entire stand. Estimation of sap flow flux density (Fd) and stand sapwood area (AS-stand) are among the most critical factors for determining forest stand transpiration using sap flow measurement. To estimate Fd, the various links in sap flow technology have great impact on the measurement of sap flow, to estimate AS-stand, an appropriate indirect technique for measuring each tree sapwood area (AS-tree) is required, because it is impossible to measure the AS-tree of all trees in a forest stand. In this study, Fd was measured in 2 mature P. euphratic trees at several radial depths, 0~10, 10~30mm, using sap flow sensors with the heat ratio method, the relationship model between AS-tree and stem diameter (DBH), growth model of AS-tree were established, using investigative original data of DBH, tree-age, and AS-tree. The results revealed that it can achieve scaling up of transpiration from sample trees to entire forest stand using AS-tree and Fd, however, the transpiration of forest stand (E) will be overvalued by 12.6% if using Fd of 0~10mm, and it will be underestimated by 25.3% if using Fd of 10~30mm, it implied that major uncertainties in mean stand Fd estimations are caused by radial variations in Fd. E will be obviously overvalued when the AS-stand is constant, this result imply that it is the key to improve the prediction accuracy that how to simulate the AS-stand changes in the day scale; They also showed that the potential errors in transpiration with a sample size of approximately ≥30 were almost stable for P.euphrtica, this suggests that to make an allometric equation it might be necessary to sample at least 30 trees.

  19. Fine root biomass in relation to site and stand characteristics in Norway spruce and Scots pine stands.

    PubMed

    Helmisaari, Heljä-Sisko; Derome, John; Nöjd, Pekka; Kukkola, Mikko

    2007-10-01

    Variations in fine root biomass of trees and understory in 16 stands throughout Finland were examined and relationships to site and stand characteristics determined. Norway spruce fine root biomass varied between 184 and 370 g m(-2), and that of Scots pine ranged between 149 and 386 g m(-2). In northern Finland, understory roots and rhizomes (< 2 mm diameter) accounted for up to 50% of the stand total fine root biomass. Therefore, the fine root biomass of trees plus understory was larger in northern Finland in stands of both tree species, resulting in a negative relationship between fine root biomass and the temperature sum and a positive relationship between fine root biomass and the carbon:nitrogen ratio of the soil organic layer. The foliage:fine root ratio varied between 2.1 and 6.4 for Norway spruce and between 0.8 and 2.2 for Scots pine. The ratio decreased for both Norway spruce and Scots pine from south to north, as well as from fertile to more infertile site types. The foliage:fine root ratio of Norway spruce was related to basal area and stem surface area. The strong positive correlations of these three parameters with fine root nitrogen concentration implies that more fine roots are needed to maintain a certain amount of foliage when nutrient availability is low. No significant relationships were found between stand parameters and fine root biomass at the stand level, but the relationships considerably improved when both fine root biomass and stand parameters were calculated for the mean tree in the stand. When the northern and southern sites were analyzed separately, fine root biomass per tree of both species was significantly correlated with basal area and stem surface area per tree. Basal area, stem surface area and stand density can be estimated accurately and easily. Thus, our results may have value in predicting fine root biomass at the tree and stand level in boreal Norway spruce and Scots pine forests.

  20. Predicting forested catchment evapotranspiration and streamflow from stand sapwood area and Aridity Index

    NASA Astrophysics Data System (ADS)

    Lane, Patrick

    2016-04-01

    Estimating the water balance of ungauged catchments has been the subject of decades of research. An extension of the fundamental problem of estimating the hydrology is then understanding how do changes in catchment attributes affect the water balance component? This is a particular issue in forest hydrology where vegetation exerts such a strong influence on evapotranspiration (ET), and consequent streamflow (Q). Given the primacy of trees in the water balance, and the potential for change to species and density through logging, fire, pests and diseases and drought, methods that directly relate ET/Q to vegetation structure, species, and stand density are very powerful. Plot studies on tree water use routinely use sapwood area (SA) to calculate transpiration and upscale to the stand/catchment scale. Recent work in south eastern Australian forests have found stand-wide SA to be linearly correlated (R2 = 0.89) with long term mean annual loss (P-Q), and hence, long term mean annual catchment streamflow. Robust relationships can be built between basal area (BA), tree density and stand SA. BA and density are common forest inventory measurements. Until now, no research has related the fundamental stand attribute of SA to streamflow. The data sets include catchments that have been thinned and with varying age classes. Thus far these analyses have been for energy limited systems in wetter forest types. SA has proven to be a more robust biometric than leaf area index which varies seasonally. That long term ET/Q is correlated with vegetation conforms to the Budyko framework. Use of a downscaled (20 m) Aridity Index (AI) has shown distinct correlations with stand SA, and therefore T. Structural patterns at a the hillslope scale not only correlate with SA and T, but also with interception (I) and forest floor evaporation (Es). These correlations between AI and I and Es have given R2 > 0.8. The result of these studies suggest an ability to estimate mean annual ET fluxes at sub hillslope scale using mappable attributes (AI, forest inventory data). Advances in forest inventory techniques, including LiDAR, mean stand attributes can increasingly be mapped over large areas. If combined with process measurements, these mapped attributes provide a powerful platform for simple but robust modelling at the sub-hillslope scale, including exploring hinge points of stand vulnerability to the drier, hotter climate predicted for SE Australia where energy limited systems may face water limitation.

  1. Computing Competition for Light in the GREENLAB Model of Plant Growth: A Contribution to the Study of the Effects of Density on Resource Acquisition and Architectural Development

    PubMed Central

    Cournède, Paul-Henry; Mathieu, Amélie; Houllier, François; Barthélémy, Daniel; de Reffye, Philippe

    2008-01-01

    Background and Aims The dynamical system of plant growth GREENLAB was originally developed for individual plants, without explicitly taking into account interplant competition for light. Inspired by the competition models developed in the context of forest science for mono-specific stands, we propose to adapt the method of crown projection onto the x–y plane to GREENLAB, in order to study the effects of density on resource acquisition and on architectural development. Methods The empirical production equation of GREENLAB is extrapolated to stands by computing the exposed photosynthetic foliage area of each plant. The computation is based on the combination of Poisson models of leaf distribution for all the neighbouring plants whose crown projection surfaces overlap. To study the effects of density on architectural development, we link the proposed competition model to the model of interaction between functional growth and structural development introduced by Mathieu (2006, PhD Thesis, Ecole Centrale de Paris, France). Key Results and Conclusions The model is applied to mono-specific field crops and forest stands. For high-density crops at full cover, the model is shown to be equivalent to the classical equation of field crop production ( Howell and Musick, 1985, in Les besoins en eau des cultures; Paris: INRA Editions). However, our method is more accurate at the early stages of growth (before cover) or in the case of intermediate densities. It may potentially account for local effects, such as uneven spacing, variation in the time of plant emergence or variation in seed biomass. The application of the model to trees illustrates the expression of plant plasticity in response to competition for light. Density strongly impacts on tree architectural development through interactions with the source–sink balances during growth. The effects of density on tree height and radial growth that are commonly observed in real stands appear as emerging properties of the model. PMID:18037666

  2. Computing competition for light in the GREENLAB model of plant growth: a contribution to the study of the effects of density on resource acquisition and architectural development.

    PubMed

    Cournède, Paul-Henry; Mathieu, Amélie; Houllier, François; Barthélémy, Daniel; de Reffye, Philippe

    2008-05-01

    The dynamical system of plant growth GREENLAB was originally developed for individual plants, without explicitly taking into account interplant competition for light. Inspired by the competition models developed in the context of forest science for mono-specific stands, we propose to adapt the method of crown projection onto the x-y plane to GREENLAB, in order to study the effects of density on resource acquisition and on architectural development. The empirical production equation of GREENLAB is extrapolated to stands by computing the exposed photosynthetic foliage area of each plant. The computation is based on the combination of Poisson models of leaf distribution for all the neighbouring plants whose crown projection surfaces overlap. To study the effects of density on architectural development, we link the proposed competition model to the model of interaction between functional growth and structural development introduced by Mathieu (2006, PhD Thesis, Ecole Centrale de Paris, France). The model is applied to mono-specific field crops and forest stands. For high-density crops at full cover, the model is shown to be equivalent to the classical equation of field crop production (Howell and Musick, 1985, in Les besoins en eau des cultures; Paris: INRA Editions). However, our method is more accurate at the early stages of growth (before cover) or in the case of intermediate densities. It may potentially account for local effects, such as uneven spacing, variation in the time of plant emergence or variation in seed biomass. The application of the model to trees illustrates the expression of plant plasticity in response to competition for light. Density strongly impacts on tree architectural development through interactions with the source-sink balances during growth. The effects of density on tree height and radial growth that are commonly observed in real stands appear as emerging properties of the model.

  3. Shrub succession on eight mixed-severity wildfires in western Montana, northeastern Oregon, and northern Idaho

    Treesearch

    Dennis E. Ferguson; John C. Byrne

    2016-01-01

    The response of 28 shrub species to wildfire burn severity was assessed for 8 wildfires on 6 national forests in the northern Rocky Mountains, USA. Stratified random sampling was used to choose 224 stands based on burn severity, habitat type series, slope steepness, stand height, and stand density, which resulted in 896 plots measured at approximately 2-year intervals...

  4. Structure of northern spotted owl nest stands and their historical conditions on the eastern slope of the Pacific Northwest Cascades, USA.

    Treesearch

    Richard Everett; D. Schellhaas; D. Spurbeck; [and others].

    1997-01-01

    The northern spotted (Strix occidentalis caurina) uses a wide array of nesting habitat throughout its current range and successfully reproduces in a variety of stand types on the eastern slope of the Pacific Northwest Cascades. The species has the ability to utilize dynamic forest stands that continue to undergo significant changes in tree density...

  5. Some effects of stand density and deer browsing on reproduction in an Adirondack hardwood stand

    Treesearch

    Robert O. Curtis; F.M. Rushmore

    1958-01-01

    The northern hardwood stands of the Adirondack region of New York constitute a major natural resource, the basis of one of the few year-round industries of the area. However, knowledge of their silviculture and potential productivity is limited. The results of an exploratory study begun in 1938 provide some information about the problems involved in the management of...

  6. Effects of repeated precommercial thinnings in central hardwood sapling stands

    Treesearch

    Donald E. Hilt; Martin E. Dale

    1982-01-01

    Precommercial thinnings were repeated four times in a central hardwood sapling stand beginning at age 8 and ending at age 22. Treated plots were thinned on an area-wide basis to specified density levels of 30-, 50-, and 70-percent stocking. The species composition of all stems in the stand was altered somewhat by thinning, but similar trends occurred on control plots....

  7. Loblolly pine foliar patterns and growth dynamics at age 12 in response to planting density and cultural intensity

    Treesearch

    Madison Katherine Akers; Michael Kane; Dehai Zhao; Richard F. Daniels; Robert O. Teskey

    2015-01-01

    Examining the role of foliage in stand development across a range of stand structures provides a more detailed understanding of the processes driving productivity and allows further development of process-based models for prediction. Productivity changes observed at the stand scale will be the integration of changes at the individual tree scale, but few studies have...

  8. Stand Density Effects on Biomass Allocation Patterns and Subsequent Soil Nitrogen Demand

    Treesearch

    Christopher A. Dicus; Thomas J. Dean

    1998-01-01

    Growth and yield data from a loblolly pine plantation in southeastern Louisiana were obtained yearly from 1993 to 1996 on numbered trees within two stands initially planted on a 1.22- by 1.22-meter spacing, and two stands planted on a 2.44- by 2.44-meter spacing. Using allometric equations derived from a 1994 on-site destructive harvest, cited nitrogen...

  9. Volume growth and response to thinning and fertilizing of Douglas-fir stands in southwestern Oregon.

    Treesearch

    R.E. Miller; G.W. Clendenen; D. Bruce

    1987-01-01

    From data for 114 thinning and fertilizing trials in forests of southwestern Oregon and northern California with 5 or more years of observation, we produced equations to estimate gross cubic volume growth of 10- to 70-year-old Douglas-fir stands. These equations use stand escriptors (breast-height age, site index, and relative density) and treatment descriptors to...

  10. Estimating loblolly pine size-density trajectories across a range of planting densities

    Treesearch

    Curtis L. VanderSchaaf; Harold E. Burkhart

    2013-01-01

    Size-density trajectories on the logarithmic (ln) scale are generally thought to consist of two major stages. The first is often referred to as the density-independent mortality stage where the probability of mortality is independent of stand density; in the second, often referred to as the density-dependent mortality or self-thinning stage, the probability of...

  11. Soil phosphorus functional fractions and tree tissue nutrient concentrations influenced by stand density in subtropical Chinese fir plantation forests

    PubMed Central

    Fang, Xiang-Min; Zhang, Xiu-Lan; Zong, Ying-Ying; Zhang, Yang; Wan, Song-Ze; Bu, Wen-Sheng

    2017-01-01

    Stand density regulation is an important measure of plantation forest management, and phosphorus (P) is often the limiting factor of tree productivity, especially in the subtropics and tropics. However, the stand density influence on ecosystem P cycling is unclear in Chinese fir (Cunninghamia lanceolata) plantations of subtropical China. We collected rhizosphere and bulk soils, leaves and twigs with different ages and roots with different orders to measure P and nitrogen (N) variables in Chinese fir plantations with low density (LDCF) and high density (HDCF) at Fujian and Hunan provinces of subtropical China. Rhizosphere soil labile P, slow P, occluded P and extractable P were higher in LDCF than HDCF at two sites. Meanwhile, P and N concentrations of 1-year-old leaves and twigs were higher in LDCF than HDCF and leaf N/P ratio generally increased with increasing leaf age at two sites. Rhizosphere vs. bulk soil labile P and occluded P were greater in LDCF than HDCF at Fujian. Nitrogen resorption efficiencies (NRE) of leaves and twigs were higher in LDCF than HDCF at Fujian, while their P resorption efficiencies (PRE) were not different between two densities at two sites. The average NRE of leaves (41.7%) and twigs (65.6%) were lower than the corresponding PRE (67.8% and 78.0%, respectively). Our results suggest that reducing stem density in Chinese fir plantations might be helpful to increase soil active P supplies and meet tree nutrient requirements. PMID:29073278

  12. Influence of light-dark schedules and stocking density on behaviour, risk of leg problems and occurrence of chronic fear in broilers.

    PubMed

    Sanotra, G S; Lund, J Damkjer; Vestergaard, K S

    2002-07-01

    1. The aims of this study were to determine (1) the effect of light-dark schedules on the walking ability, the risk of tibial dyschondroplasia (TD) as well as the duration of tonic immobility (TI) reactions in commercial broiler flocks and (2) the effect of a daily dark period and reduced density on the behaviour of broiler chickens. 2. Experiment 1. Group 1 had a 2 to 8 h daily dark period from 2 to 26 d of age (light-dark programme A) at a stocking density of 28.4 chicks/m2. Group 2 had 8 h of darkness daily from 2 to 38 d of age (light-dark programme B) at 24 chicks/m2. The control group had 24 h continuous light at 28.4 chicks/m2. 3. Experiment 2. Behaviour was studied with and without a daily 8 h dark period and at high (30 chicks/m2) and low (18 chicks/m2) stocking densities. 4. Programme B reduced the prevalence of impaired walking ability, corresponding to gait score > 2, when compared with controls. The effect on walking ability corresponding to gait score > 0 approached significance. 5. Both light-dark programmes reduced the occurrence of TD. Programme B (combined with reduced stocking density), however, had the greater effect. 6. Both light-dark programmes reduced the duration of TI, compared with controls (mean = 426 s) Programme B resulted in a larger reduction (alpha = -156.9 s) than programme A (alpha = -117.0). 7. The proportions of chicks drinking, eating, pecking, scratching, standing and performing vertical wing-shakes increased--both when the 8 h dark period and the reduced stocking density were applied separately and in combination (experiment 2). 8. For all behaviours, except standing, the effect of the dark period was largest in broilers kept at the high stocking density (d 40).

  13. Field-based evaluations of sampling techniques to support long-term monitoring of riparian ecosystems along wadeable streams on the Colorado Plateau

    USGS Publications Warehouse

    Scott, Michael L.; Reynolds, Elizabeth W.

    2007-01-01

    Compared to 5-m by 20-m tree quadrats, belt transects were shown to provide similar estimates of stand structure (stem density and stand basal area) in less than 30 percent of the time. Further, for the streams sampled, there were no statistically significant differences in stem density and basal area estimates between 10-m and 20-m belt transects and the smaller belts took approximately half the time to sample. There was, however, high variance associated with estimates of stand structure for infrequently occurring stems, such as large, relict or legacy riparian trees. Legacy riparian trees occurred in limited numbers at all sites sampled. A reachscale population census of these trees indicated that the 10-m belt transects tended to underestimate both stem density and basal area for these riparian forest elements and that a complete reach-scale census of legacy trees averaged less than one hour per site.

  14. Nanoporous distributed Bragg reflectors on free-standing nonpolar m-plane GaN

    NASA Astrophysics Data System (ADS)

    Mishkat-Ul-Masabih, Saadat; Luk, Ting Shan; Rishinaramangalam, Ashwin; Monavarian, Morteza; Nami, Mohsen; Feezell, Daniel

    2018-01-01

    We report the fabrication of m-plane nanoporous distributed Bragg reflectors (DBRs) on free-standing GaN substrates. The DBRs consist of 15 pairs of alternating undoped and highly doped n-type ([Si] = ˜3.7 × 1019 cm-3) GaN. Electrochemical (EC) etching was performed to convert the highly doped regions into a porous material, consequently reducing the effective refractive index of the layers. We demonstrate a DBR with peak reflectance greater than 98% at 450 nm with a stopband width of ˜72 nm. The polarization ratio of an incident polarized light source remains identical after reflection from the DBR, verifying that there is no drop in the polarization ratio due to the interfaces between the porous layers. We also quantify the porosity under various EC bias conditions for layers with different doping concentrations. The bias voltage controls the average pore diameter, while the pore density is primarily determined by the doping concentration. The results show that nanoporous DBRs on nonpolar free-standing GaN are promising candidates for high-reflectance, lattice-matched DBR mirrors for GaN-based resonant cavity devices.

  15. Effectiveness of management interventions on forest carbon stock in planted forests in Nepal.

    PubMed

    Dangal, Shambhu Prasad; Das, Abhoy Kumar; Paudel, Shyam Krishna

    2017-07-01

    Nepal has successfully established more than 370,000 ha of plantations, mostly with Pinus patula, in the last three and a half decades. However, intensive management of these planted forests is very limited. Despite the fact that the Kyoto Convention in 1997 recognized the role of plantations for forest-carbon sequestration, there is still limited knowledge on the effects of management practices and stand density on carbon-sequestration of popular plantation species (i.e. Pinus patula) in Nepal. We carried out case studies in four community forests planted between 1976 and 1990 to assess the impacts of management on forest carbon stocks. The study found that the average carbon stock in the pine plantations was 217 Mg C ha -1 , and was lower in forests with intensively managed plantations (214.3 Mg C ha -1 ) than in traditionally managed plantations (219 Mg C ha -1 ). However, it was the reverse in case of soil carbon, which was higher (78.65 Mg C ha -1 ) in the forests with intensive management. Though stand density was positively correlated with carbon stock, the proportionate increment in carbon stock was lower with increasing stand density, as carbon stock increased by less than 25% with a doubling of stand density (300-600). The total carbon stock was higher in plantations aged between 25 and 30 years compared to those aged between 30 and 35 years. Copyright © 2017 Elsevier Ltd. All rights reserved.

  16. Floor space covered by broiler chickens kept at stocking densities according to Council Directive 2007/43/EC.

    PubMed

    Giersberg, M F; Hartung, J; Kemper, N; Spindler, B

    2016-07-30

    It is controversially discussed whether the stocking densities set by the EU Directive 2007/43/EC allow a species-appropriate housing of broiler chickens. To calculate the exact area broilers occupy due to their physical size and shape, planimetric measurements using a colour-contrast method were carried out. In total, 1949 photographs of standing and 1482 of squatting chickens, taken from a top view, were analysed. A computer program counted the pixels representing the previously weighed animal in the photograph and calculated the animal area. The average area covered by chickens with 400 g live weight was 116.64±13.12 cm(2) in a standing and 138.61±12.92 cm(2) in a squatting position. These areas increased linearly as a function of live weight to 452.57±58.89 cm(2) (R(2)=0.90 standing) and 513.54±42.70 cm(2) (R(2)=0.82 squatting) at the end of the study (3200 g live weight). Squatting chickens occupied more space compared with a standing position in most of the tested weight classes (P<0.05). Depending on target weights, stocking densities and body positions, broilers occupied 48.5-77.7 per cent of 1 m(2) Thus, from a physical point of view, simultaneous resting is possible at any stocking density provided by the EU Directive and at common target weights. British Veterinary Association.

  17. Wetland biomass crops: Studies in natural and managed stands

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Andrews, N.J.

    1989-01-01

    Wetlands dominated by Typha spp. and other emergent macrophytes are some of the most productive natural systems in the temperate zone. The subject of this thesis is the selection and management of potential emergent biomass crops and is presented in the following three chapters: (1) The Productivity of Typha spp. in Managed Stands in Minnesota, (II) The Establishment of Typha spp. on a Northern Minnesota Peatland, and (III) Wetland Species Yield Comparisons. Typha rhizomes were planted in a series of 1.5 m{sup 2} paddies. Variables investigated included planting density, fertilizer application rate and seasonal changes in above and below-ground biomass,more » shoot height and density. Yield increased with increased planting density. Maximum shoot dry weight occurred in August while maximum below ground dry weight was obtained in October. Trial plots were established on a northern Minnesota peatland using both rhizomes and seed. The influence of planting density, substrate preparation and fertilizer application on the seasonal development and productivity of trial plots was investigated. Through extensive literature surveys and sampling natural stands, five species were identified for further screening including: Carex atherodes, Phragmites australis, Scirpus fluviatilis, Sparganium eurycarpum and Spartina pectinata. These species, along with planting stock from five productive Typha stands were planted in 1.5 m{sup 2} paddies to compare productivity and growth characteristics under identical conditions. Trial plots of Phragmites, Scirpus and Sparganium were also established on excavated peatland plots.« less

  18. Interactive effects of historical logging and fire exclusion on ponderosa pine forest structure in the northern Rockies.

    PubMed

    Naficy, Cameron; Sala, Anna; Keeling, Eric G; Graham, Jon; DeLuca, Thomas H

    2010-10-01

    Increased forest density resulting from decades of fire exclusion is often perceived as the leading cause of historically aberrant, severe, contemporary wildfires and insect outbreaks documented in some fire-prone forests of the western United States. Based on this notion, current U.S. forest policy directs managers to reduce stand density and restore historical conditions in fire-excluded forests to help minimize high-severity disturbances. Historical logging, however, has also caused widespread change in forest vegetation conditions, but its long-term effects on vegetation structure and composition have never been adequately quantified. We document that fire-excluded ponderosa pine forests of the northern Rocky Mountains logged prior to 1960 have much higher average stand density, greater homogeneity of stand structure, more standing dead trees and increased abundance of fire-intolerant trees than paired fire-excluded, unlogged counterparts. Notably, the magnitude of the interactive effect of fire exclusion and historical logging substantially exceeds the effects of fire exclusion alone. These differences suggest that historically logged sites are more prone to severe wildfires and insect outbreaks than unlogged, fire-excluded forests and should be considered a high priority for fuels reduction treatments. Furthermore, we propose that ponderosa pine forests with these distinct management histories likely require distinct restoration approaches. We also highlight potential long-term risks of mechanical stand manipulation in unlogged forests and emphasize the need for a long-term view of fuels management.

  19. Community-wide spatial and temporal discordances of seed-seedling shadows in a tropical rainforest.

    PubMed

    Rother, Débora Cristina; Pizo, Marco Aurélio; Siqueira, Tadeu; Rodrigues, Ricardo Ribeiro; Jordano, Pedro

    2015-01-01

    Several factors decrease plant survival throughout their lifecycles. Among them, seed dispersal limitation may play a major role by resulting in highly aggregated (contagious) seed and seedling distributions entailing increased mortality. The arrival of seeds, furthermore, may not match suitable environments for seed survival and, consequently, for seedling establishment. In this study, we investigated spatio-temporal patterns of seed and seedling distribution in contrasting microhabitats (bamboo and non-bamboo stands) from the Brazilian Atlantic Forest. Spatial distribution patterns, spatial concordance between seed rain and seedling recruitment between subsequent years in two fruiting seasons (2004-2005 and 2007-2009), and the relation between seeds and seedlings with environmental factors were examined within a spatially-explicit framework. Density and species richness of both seeds and seedlings were randomly distributed in non-bamboo stands, but showed significant clustering in bamboo stands. Seed and seedling distributions showed across-year inconsistency, suggesting a marked spatial decoupling of the seed and seedling stages. Generalized linear mixed effects models indicated that only seed density and seed species richness differed between stand types while accounting for variation in soil characteristics. Our analyses provide evidence of marked recruitment limitation as a result of the interplay between biotic and abiotic factors. Because bamboo stands promote heterogeneity in the forest, they are important components of the landscape. However, at high densities, bamboos may limit recruitment for the plant community by imposing marked discordances of seed arrival and early seedling recruitment.

  20. [Population structure of soil arthropod in different age Pinus massoniana plantations].

    PubMed

    Tan, Bo; Wu, Fu-zhong; Yang, Wan-qin; Zhang, Jian; Xu, Zhen-feng; Liu, Yang; Gou, Xiao-lin

    2013-04-01

    An investigation was conducted on the population structure of soil arthropod community in the 3-, 8-, 14-, 31-, and 40-years old Pinus massoniana plantations in the upper reaches of the Yangtze River in spring (May) and autumn (October), 2011, aimed to search for the scientific management of the plantation. A total of 4045 soil arthropods were collected, belonging to 57 families. Both the individual density and the taxonomic group number of the soil arthropod community decreased obviously with increasing soil depth, and this trend increased with increasing stand age. The dominant groups and ordinary groups of the soil arthropod community varied greatly with the stand age of P. massoniana plantation, and a significant difference (P<0.05) was observed in the individual density and taxonomic group number among different age P. massoniana plantations. In comparison with other stand age P. massoniana plantations, 3years old P. massoniana plantation had a significant difference in the structure and diversity of soil arthropod community, and the similarity index of the soil arthropod community was lower. The individual density, taxonomic group number, and diversity of soil arthropod community were the highest in 8-years old P. massoniana plantation, and then, decreased obviously with increasing stand age. It was suggested that the land fertility of the P. massoniana plantations could be degraded with increasing stand age, and it would be appropriate to make artificial regulation and restoration in 8-years old P. massoniana plantation.

  1. Soil variation and sampling intensity under red pine and aspen in Minnesota.

    Treesearch

    David H. Alban

    1974-01-01

    In red pine or aspen stands only two soil samples were needed to estimate (+/- 10%, 95% confidence) pH, bulk density, or sand, but 25 to 60 samples were required to estimate N, P, K, Ca, Mg, available water, or silt + clay. To estimate most forest floor properties required 30 to 50 samples in red pine stands, but only about half as many in aspen stands.

  2. Long-term precommercial thinning effects on Larix occidentalis (western larch) tree and stand characteristics

    Treesearch

    Michael S. Schaedel; Andrew J. Larson; David L. R. Affleck; R. Travis Belote; John M. Goodburn; David K. Wright; Elaine Kennedy Sutherland

    2017-01-01

    Precommercial thinning (PCT) is used to increase tree size and shorten harvest rotation time. Short-term results from PCT studies often show a trade-off between individual-tree growth and net stand yield, while longer-term effects of PCT on tree growth and stand yield are less well documented. We used a 54-year-old PCT study to test long-term effects of forest density...

  3. Recent changes in the estimation of standing dead tree biomass and carbon stocks in the U.S. forest inventory

    Treesearch

    Grant M. Domke; Christopher W. Woodall; James E. Smith

    2012-01-01

    Until recently, standing dead tree biomass and carbon (C) has been estimated as a function of live tree growing stock volume in the U.S. Forest Service, Forest Inventory and Analysis (FIA) Program. Traditional estimates of standing dead tree biomass/C attributes were based on merchantability standards that did not reflect density reductions or structural loss due to...

  4. Commercial thinning in small-diameter aspen stands in northern Minnesota: study establishment report

    Treesearch

    Daniel W. Gilmore; Jennifer D. Glenn; Michael E. Ostry; John C. Zasada; Michael A. Benedict

    2006-01-01

    In the spring of 1999, a long-term study was established to examine the physical and biological aspects of thinning young aspen stands in Minnesota. Three aspen stands ranging in age from 25 to 35 years were selected on lands owned by the State of Minnesota and UPM Kymmene. Two thinning treatments (low and high density) and an unthinned control were installed at each...

  5. Use of the Weibull function to predict future diameter distributions from current plot data

    Treesearch

    Quang V. Cao

    2012-01-01

    The Weibull function has been widely used to characterize diameter distributions in forest stands. The future diameter distribution of a forest stand can be predicted by use of a Weibull probability density function from current inventory data for that stand. The parameter recovery approach has been used to “recover” the Weibull parameters from diameter moments or...

  6. Simulation of dynamics of southern pine beetle hazard rating with respect to silvicultural treatment and stand development

    Treesearch

    D. J. Leduc; J. C. G. Goelz

    2010-01-01

    The hazard of southern pine beetle (SPB) infestations is affected by characteristics such as stand density, stand age, site quality, and tree size. COMPUTE P-LOB is a model that simulates the growth and development of loblolly pine plantations in the west gulf coastal plain. P-LOB was rewritten as COMPUTE SPB-Lob to update it for current operating systems and to...

  7. Forest volume-to-biomass models and estimates of mass for live and standing dead trees of U.S. forests.

    Treesearch

    James E. Smith; Linda S. Heath; Jennifer C. Jenkins

    2003-01-01

    Includes methods and equations for nationally consistent estimates of tree-mass density at the stand level (Mg/ha) as predicted by growing-stock volumes reported by the USDA Forest Service for forests of the conterminous United States. Developed for use in FORCARB, a carbon budget model for U.S. forests, the equations also are useful for converting plot-, stand- and...

  8. Individual tree- versus stand-level approaches to thinning: is it a choice of one or the other, or a combination of both?

    Treesearch

    Christopher A. Nowak

    1995-01-01

    Thinning guidelines have existed for most eastern hardwood forests for 20 to 30 years. While these guidelines are presented in varying degrees of detail, they generally all contain recommendations on levels of residual stand density and stand structure, along with information on crop tree requirements. Recent attempts have been made to simplify thinning guidelines by...

  9. Stand density-a factor affecting stem quality of young hardwoods

    Treesearch

    David L. Sonderman; David L. Sonderman

    1985-01-01

    A 14-year-old mixed oak stand was thinned in 1977 to stocking levels of 30, 50, and 60 percent and a control. From this stand, 117 trees were selected and their stem-related defects recorded. Six years later these same trees were reevaluated. The number of limb defects per square foot of surface area increased substantially more in the heavily thinned plots than in the...

  10. The role of red alder in riparian forest structure along headwater streams in southeastern Alaska

    USGS Publications Warehouse

    Orlikowska, E.H.; Deal, R.L.; Hennon, P.E.; Wipfli, M.S.

    2004-01-01

    We assessed the influence of red alder on tree species composition, stand density, tree size distribution, tree mortality, and potential for producing large conifers, in 38-42 yr old riparian forests along 13 headwater streams in the Maybeso and Harris watersheds on Prince of Wales Island, Alaska. Red alder ranged from 0 to 53% of the total live basal area of the stands. Tree density, basal area of live and dead trees, and mean diameter of live conifers were not significantly related to the percent of alder as a proportion of total stand live basal area within these riparian forests. The mean diameter of the 100 largest conifers per hectare (the largest trees) was similar among different sites and appeared unrelated to the amount of alder in the stands. The mean diameter of dead conifers increased slightly with increasing proportion of red alder. Most dead trees were small and died standing. Red alder was much more concentrated immediately along stream margins (within 0-1 m distance from the stream bank vs. > 1 m). The presence of red alder did not inhibit the production of large-diameter conifers, and both alder and conifers provided small woody debris for fishless headwater streams in southeastern Alaska. Red alder is an important structural component of young-growth riparian stands.

  11. Oscillations of a standing shock wave generated by the Richtmyer-Meshkov instability

    NASA Astrophysics Data System (ADS)

    Mikaelian, Karnig O.

    2016-07-01

    In a typical Richtmyer-Meshkov experiment a fast moving flat shock strikes a stationary perturbed interface between fluids A and B creating a transmitted and a reflected shock, both of which are perturbed. We propose shock tube experiments in which the reflected shock is stationary in the laboratory. Such a standing perturbed shock undergoes well-known damped oscillations. We present the conditions required for producing such a standing shock wave, which greatly facilitates the measurement of the oscillations and their rate of damping. We define a critical density ratio Rcritical, in terms of the adiabatic indices of the two fluids, and a critical Mach number Mscritical of the incident shock wave, which produces a standing reflected wave. If the initial density ratio R of the two fluids is less than Rcritical then a standing shock wave is possible at Ms=Mscritical . Otherwise a standing shock is not possible and the reflected wave always moves in the direction opposite the incident shock. Examples are given for present-day operating shock tubes with sinusoidal or inclined interfaces. We consider the effect of viscosity, which affects the damping rate of the oscillations. We point out that nonlinear bubble and spike amplitudes depend relatively weakly on the viscosity of the fluids and that the interface area is a better diagnostic.

  12. Densities of breeding birds and changes in vegetation in an alaskan boreal forest following a massive disturbance by spruce beetles

    USGS Publications Warehouse

    Matsuoka, S.M.; Handel, C.M.; Ruthrauff, D.R.

    2001-01-01

    We examined bird and plant communities among forest stands with different levels of spruce mortality following a large outbreak of spruce beetles (Dendroctonus rufipennis (Kirby)) in the Copper River Basin, Alaska. Spruce beetles avoided stands with black spruce (Picea mariana) and selectively killed larger diameter white spruce (Picea glauca), thereby altering forest structure and increasing the dominance of black spruce in the region. Alders (Alnus sp.) and crowberry (Empetrum nigrum) were more abundant in areas with heavy spruce mortality, possibly a response to the death of overstory spruce. Grasses and herbaceous plants did not proliferate as has been recorded following outbreaks in more coastal Alaskan forests. Two species closely tied to coniferous habitats, the tree-nesting Ruby-crowned Kinglet (Regulus calendula) and the red squirrel (Tamiasciurus hudsonicus), a major nest predator, were less abundant in forest stands with high spruce mortality than in low-mortality stands. Understory-nesting birds as a group were more abundant in forest stands with high levels of spruce mortality, although the response of individual bird species to tree mortality was variable. Birds breeding in stands with high spruce mortality likely benefited reproductively from lower squirrel densities and a greater abundance of shrubs to conceal nests from predators.

  13. Agricultural legacy, climate, and soil influence the restoration and carbon potential of woody regrowth in Australia.

    PubMed

    Dwyer, John M; Fensham, Rod J; Buckley, Yvonne M

    2010-10-01

    Opportunities for dual restoration and carbon benefits from naturally regenerating woody ecosystems in agricultural landscapes have been highlighted recently. The restoration capacity of woody ecosystems depends on the magnitude and duration of ecosystem modification, i.e., the "agricultural legacy." However, this legacy may not influence carbon sequestration in the same way as restoration because carbon potential depends primarily on biomass accumulation, with little consideration of other attributes and functions of the ecosystem. Our present study simultaneously assesses the restoration and carbon potential of Acacia harpophylla regrowth, an extensive regrowth ecosystem in northeastern Australia. We used a landscape-scale survey of A. harpophylla regrowth to test the following hypotheses: (1) management history, in combination with climatic and edaphic factors, has long-term effects on stem densities, and (2) higher-density stands have lower restoration and carbon potential, which is also influenced by climatic and edaphic factors. We focused on the restoration of forest structure, which was characterized using stem density, aboveground biomass, stem heights, and stem diameters. Data were analyzed using multilevel models within the hierarchical Bayesian model (HBM) framework. We found strong support for both hypotheses. Repeated attempts at clearing Brigalow (A. harpophylla ecosystem) regrowth increases stem densities, and these densities remain high over the long term, particularly in high-rainfall areas and on gilgaied, high-clay soils (hypothesis 1). In models testing hypothesis 2, interactions between stem density and stand age indicate that higher-density stands have slower biomass accumulation and structural development in the long term. After accounting for stem density and stand age, annual rainfall had a positive effect on biomass accumulation and structural development. Other climate and soil variables were retained in the various models but had weaker effects. Spatial extrapolations of the HBMs indicated that the central and eastern parts of the study region are most suitable for biomass accumulation; however, these may not correspond to the areas that historically supported the highest biomass Brigalow forests. We conclude that carbon and restoration goals are largely congruent within areas of similar climate. At the regional scale, however, spatial prioritization of restoration and carbon projects may only be aligned where carbon benefits will be high.

  14. Influence of Anthropogenic Disturbances on Stand Structural Complexity in Andean Temperate Forests: Implications for Managing Key Habitat for Biodiversity.

    PubMed

    Caviedes, Julián; Ibarra, José Tomás

    2017-01-01

    Forest attributes and their abundances define the stand structural complexity available as habitat for faunal biodiversity; however, intensive anthropogenic disturbances have the potential to degrade and simplify forest stands. In this paper we develop an index of stand structural complexity and show how anthropogenic disturbances, namely fire, logging, livestock, and their combined presence, affect stand structural complexity in a southern Global Biodiversity Hotspot. From 2011 to 2013, we measured forest structural attributes as well as the presence of anthropogenic disturbances in 505 plots in the Andean zone of the La Araucanía Region, Chile. In each plot, understory density, coarse woody debris, number of snags, tree diameter at breast height, and litter depth were measured, along with signs of the presence of anthropogenic disturbances. Ninety-five percent of the plots showed signs of anthropogenic disturbance (N = 475), with the combined presence of fire, logging, and livestock being the most common disturbance (N = 222; 44% of plots). The lowest values for the index were measured in plots combining fire, logging, and livestock. Undisturbed plots and plots with the presence of relatively old fires (> 70 years) showed the highest values for the index of stand structural complexity. Our results suggest that secondary forests < 70-year post-fire event, with the presence of habitat legacies (e.g. snags and CWD), can reach a structural complexity as high as undisturbed plots. Temperate forests should be managed to retain structural attributes, including understory density (7.2 ± 2.5 # contacts), volume of CWD (22.4 ± 25.8 m3/ha), snag density (94.4 ± 71.0 stems/ha), stand basal area (61.2 ± 31.4 m2/ha), and litter depth (7.5 ± 2.7 cm). Achieving these values will increase forest structural complexity, likely benefiting a range of faunal species in South American temperate forests.

  15. Influence of Anthropogenic Disturbances on Stand Structural Complexity in Andean Temperate Forests: Implications for Managing Key Habitat for Biodiversity

    PubMed Central

    2017-01-01

    Forest attributes and their abundances define the stand structural complexity available as habitat for faunal biodiversity; however, intensive anthropogenic disturbances have the potential to degrade and simplify forest stands. In this paper we develop an index of stand structural complexity and show how anthropogenic disturbances, namely fire, logging, livestock, and their combined presence, affect stand structural complexity in a southern Global Biodiversity Hotspot. From 2011 to 2013, we measured forest structural attributes as well as the presence of anthropogenic disturbances in 505 plots in the Andean zone of the La Araucanía Region, Chile. In each plot, understory density, coarse woody debris, number of snags, tree diameter at breast height, and litter depth were measured, along with signs of the presence of anthropogenic disturbances. Ninety-five percent of the plots showed signs of anthropogenic disturbance (N = 475), with the combined presence of fire, logging, and livestock being the most common disturbance (N = 222; 44% of plots). The lowest values for the index were measured in plots combining fire, logging, and livestock. Undisturbed plots and plots with the presence of relatively old fires (> 70 years) showed the highest values for the index of stand structural complexity. Our results suggest that secondary forests < 70-year post-fire event, with the presence of habitat legacies (e.g. snags and CWD), can reach a structural complexity as high as undisturbed plots. Temperate forests should be managed to retain structural attributes, including understory density (7.2 ± 2.5 # contacts), volume of CWD (22.4 ± 25.8 m3/ha), snag density (94.4 ± 71.0 stems/ha), stand basal area (61.2 ± 31.4 m2/ha), and litter depth (7.5 ± 2.7 cm). Achieving these values will increase forest structural complexity, likely benefiting a range of faunal species in South American temperate forests. PMID:28068349

  16. Thickness-dependent phase transition in graphite under high magnetic field

    NASA Astrophysics Data System (ADS)

    Taen, Toshihiro; Uchida, Kazuhito; Osada, Toshihito

    2018-03-01

    Various electronic phases emerge when applying high magnetic fields in graphite. However, the origin of a semimetal-insulator transition at B ≃30 T is still not clear, while an exotic density-wave state is theoretically proposed. In order to identify the electronic state of the insulator phase, we investigate the phase transition in thin-film graphite samples that were fabricated on silicon substrate by a mechanical exfoliation method. The critical magnetic fields of the semimetal-insulator transition in thin-film graphite shift to higher magnetic fields, accompanied by a reduction in temperature dependence. These results can be qualitatively reproduced by a density-wave model by introducing a quantum size effect. Our findings establish the electronic state of the insulator phase as a density-wave state standing along the out-of-plane direction, and help determine the electronic states in other high-magnetic-field phases.

  17. A ponderosa pine natural area reveals its secrets

    USGS Publications Warehouse

    Allen, Craig D.; Mac, Michael J.; Opler, Paul A.; Puckett Haecker, Catherine E.; Doran, Peter D.

    1998-01-01

    Monument Canyon Research Natural Area preserves an unlogged 259-hectare stand of old-growth ponderosa pine in the Jemez Mountains of New Mexico. This preserve, established in 1932, is the oldest research natural area in the state. This two-tiered forest displays an old-growth density of 100 stems per hectare (Muldavin et al. 1995), with an understory thicket of stagnant saplings and poles that raises the total stand density to an average of 5,954 stems per hectare, with concentrations as high as 21,617 stems per hectare (Fig. 1).

  18. Measuring Effective Leaf Area Index, Foliage Profile, and Stand Height in New England Forest Stands Using a Full-Waveform Ground-Based Lidar

    NASA Technical Reports Server (NTRS)

    Zhao, Feng; Yang, Xiaoyuan; Schull, Mithcell A.; Roman-Colon, Miguel O.; Yao, Tian; Wang, Zhuosen; Zhang, Qingling; Jupp, David L. B.; Lovell, Jenny L.; Culvenor, Darius; hide

    2011-01-01

    Effective leaf area index (LAI) retrievals from a scanning, ground-based, near-infrared (1064 nm) lidar that digitizes the full return waveform, the Echidna Validation Instrument (EVI), are in good agreement with those obtained from both hemispherical photography and the Li-Cor LAI-2000 Plant Canopy Analyzer. We conducted trials at 28 plots within six stands of hardwoods and conifers of varying height and stocking densities at Harvard Forest, Massachusetts, Bartlett Experimental Forest, New Hampshire, and Howland Experimental Forest, Maine, in July 2007. Effective LAI values retrieved by four methods, which ranged from 3.42 to 5.25 depending on the site and method, were not significantly different ( b0.1 among four methods). The LAI values also matched published values well. Foliage profiles (leaf area with height) retrieved from the lidar scans, although not independently validated, were consistent with stand structure as observed and as measured by conventional methods. Canopy mean top height, as determined from the foliage profiles, deviated from mean RH100 values obtained from the Lidar Vegetation Imaging Sensor (LVIS) airborne large-footprint lidar system at 27 plots by .0.91 m with RMSE=2.04 m, documenting the ability of the EVI to retrieve stand height. The Echidna Validation Instrument is the first realization of the Echidna lidar concept, devised by Australia's Commonwealth Scientific and Industrial Research Organization (CSIRO), for measuring forest structure using full-waveform, ground-based, scanning lidar.

  19. [Correlations between standing trees trunk decay degree and soil physical-chemical properties in Korean pine-broadleaved mixed forest in Xiao Xing'an Mountains of Northeast China].

    PubMed

    Sun, Tian-Yong; Wang, Li-Hai; Sun, Mo-Long

    2013-07-01

    Standing trees decay often causes vast loss of timber resources. To investigate the correlations between the standing trees decay and the site conditions is of importance to scientifically and reasonably manage forests and to decrease wood resources loss. By using Resistograph and meter ruler, a measurement was made on the decay degree of the trunk near root and the diameter at breast height (DBH) of 15 mature Korean pine standing trees in a Korean pine-broadleaved mixed forest in Xiao Xing' an Mountains in May, 2011. In the meantime, soil samples were collected from the root zones of standing trees and the upslope and downslope 5 meters away from the trunks, respectively. Five physical-chemical properties including moisture content, bulk density, total porosity, pH value, and organic matter content of the soil samples were tested. The regression equations concerning the trunk decay degree of the standing trees, their DBH, and the 5 soil properties were established. The results showed that the trunk decay degree of the mature Korean pine standing trees had higher correlations with the bulk density, total porosity, pH value, and organic matter content (R = 0.687), and significant positive correlation with the moisture content (R = 0.507) of the soils at the root zones of standing trees, but less correlation with the 5 properties of the soils at both upslope and downslope 5 meters away from the trunks. The trunk decay degree was decreased when the soil moisture content was below 18.4%. No significant correlation was observed between the trunk decay degree of mature Korean pine standing trees and the tree age.

  20. A hydroeconomic modeling framework for optimal integrated management of forest and water

    NASA Astrophysics Data System (ADS)

    Garcia-Prats, Alberto; del Campo, Antonio D.; Pulido-Velazquez, Manuel

    2016-10-01

    Forests play a determinant role in the hydrologic cycle, with water being the most important ecosystem service they provide in semiarid regions. However, this contribution is usually neither quantified nor explicitly valued. The aim of this study is to develop a novel hydroeconomic modeling framework for assessing and designing the optimal integrated forest and water management for forested catchments. The optimization model explicitly integrates changes in water yield in the stands (increase in groundwater recharge) induced by forest management and the value of the additional water provided to the system. The model determines the optimal schedule of silvicultural interventions in the stands of the catchment in order to maximize the total net benefit in the system. Canopy cover and biomass evolution over time were simulated using growth and yield allometric equations specific for the species in Mediterranean conditions. Silvicultural operation costs according to stand density and canopy cover were modeled using local cost databases. Groundwater recharge was simulated using HYDRUS, calibrated and validated with data from the experimental plots. In order to illustrate the presented modeling framework, a case study was carried out in a planted pine forest (Pinus halepensis Mill.) located in south-western Valencia province (Spain). The optimized scenario increased groundwater recharge. This novel modeling framework can be used in the design of a "payment for environmental services" scheme in which water beneficiaries could contribute to fund and promote efficient forest management operations.

  1. Variation in wood density by stand origin and log position for loblolly pine sawtimber in the coastal plain of Arkansas

    Treesearch

    Paul F. Doruska; David W. Patterson; Travis E. Posey

    2006-01-01

    A study was undertaken to investigate and report scaling factor variation for loblolly pine sawtimber in the Coastal Plain of Arkansas. Scaling factors for butt logs averaged 65.6 pounds per cubic foot for trees in stands of naturally seeded origin and 65.0 pounds per cubic foot for plantation trees. These scaling factors were not significantly different by stand...

  2. Forest regeneration composition and development in upland, mixed-oak forests.

    PubMed

    Fei, Songlin; Gould, Peter J; Steiner, Kim C; Finley, James C; McDill, Marc E

    2005-12-01

    Advance regeneration in 52 mature mixed-oak stands was analyzed and described. Red maple (Acer rubrum L.) was the most abundant species in the study area. Among oak (Quercus) species, northern red oak (Q. rubra L.) was the most abundant within the Allegheny Plateau physiographic province, whereas chestnut oak (Q. montana L.) was the most abundant within the Ridge and Valley physiographic province. Sixteen stands, for which data are available through the fourth growing season following harvest, were used to describe stand development. Cumulative height, a composite measure of size and density, was used to describe early stand development. Black gum (Nyssa sylvatica Marsh.) and black birch (Betula lenta L.) had dramatic increases in stand density and cumulative height after overstory removal. Cumulative height of northern red oak and chestnut oak showed a faster positive response to overstory removal than red maple. Oak retained its dominance in cumulative height for at least 4 years after harvest. Red maple nevertheless remained the most abundant tree species after overstory removal. Our results suggest that the principal advantage of red maple regeneration is its ability to accumulate in large numbers prior to harvest.

  3. [Vertical distribution of fuels in Pinus yunnanensis forest and related affecting factors].

    PubMed

    Wang, San; Niu, Shu-Kui; Li, De; Wang, Jing-Hua; Chen, Feng; Sun, Wu

    2013-02-01

    In order to understand the effects of fuel loadings spatial distribution on forest fire kinds and behaviors, the canopy fuels and floor fuels of Pinus yunnanensis forests with different canopy density, diameter at breast height (DBH), tree height, and stand age and at different altitude, slope grade, position, and aspect in Southwest China were taken as test objects, with the fuel loadings and their spatial distribution characteristics at different vertical layers compared and the fire behaviors in different stands analyzed. The relationships between the fuel loadings and the environmental factors were also analyzed by canonical correspondence analysis (CCA). In different stands, there existed significant differences in the vertical distribution of fuels. Pinus yunnanensis-Qak-Syzygium aromaticum, Pinus yunnanensis-oak, and Pinus yunnanensis forests were likely to occur floor fire but not crown fire, while Pinus yunnanensis-Platycladus orientalis, Pinus yunnanensis-Keteleeria fortune, and Keteleeria fortune-Pinus yunnanensis were not only inclined to occur floor fire, but also, the floor fire could be easily transformed into crown fire. The crown fuels were mainly affected by the stand age, altitude, DBH, and tree height, while the floor fuels were mainly by the canopy density, slope grade, altitude, and stand age.

  4. Fluctuations around equilibrium laws in ergodic continuous-time random walks.

    PubMed

    Schulz, Johannes H P; Barkai, Eli

    2015-06-01

    We study occupation time statistics in ergodic continuous-time random walks. Under thermal detailed balance conditions, the average occupation time is given by the Boltzmann-Gibbs canonical law. But close to the nonergodic phase, the finite-time fluctuations around this mean are large and nontrivial. They exhibit dual time scaling and distribution laws: the infinite density of large fluctuations complements the Lévy-stable density of bulk fluctuations. Neither of the two should be interpreted as a stand-alone limiting law, as each has its own deficiency: the infinite density has an infinite norm (despite particle conservation), while the stable distribution has an infinite variance (although occupation times are bounded). These unphysical divergences are remedied by consistent use and interpretation of both formulas. Interestingly, while the system's canonical equilibrium laws naturally determine the mean occupation time of the ergodic motion, they also control the infinite and Lévy-stable densities of fluctuations. The duality of stable and infinite densities is in fact ubiquitous for these dynamics, as it concerns the time averages of general physical observables.

  5. Potential effects of forest management on surface albedo

    NASA Astrophysics Data System (ADS)

    Otto, J.; Bréon, F.-M.; Schelhaas, M.-J.; Pinty, B.; Luyssaert, S.

    2012-04-01

    Currently 70% of the world's forests are managed and this figure is likely to rise due to population growth and increasing demand for wood based products. Forest management has been put forward by the Kyoto-Protocol as one of the key instruments in mitigating climate change. For temperate and boreal forests, the effects of forest management on the stand-level carbon balance are reasonably well understood, but the biophysical effects, for example through changes in the albedo, remain elusive. Following a modeling approach, we aim to quantify the variability in albedo that can be attributed to forest management through changes in canopy structure and density. The modelling approach chains three separate models: (1) a forest gap model to describe stand dynamics, (2) a Monte-Carlo model to estimate the probability density function of the optical path length of photons through the canopy and (3) a physically-based canopy transfer model to estimate the interaction between photons and leaves. The forest gap model provides, on a monthly time step the position, height, diameter, crown size and leaf area index of individual trees. The Monte-Carlo model computes from this the probability density function of the distance a photon travels through crown volumes to determine the direct light reaching the forest floor. This information is needed by the canopy transfer model to calculate the effective leaf area index - a quantity that allows it to correctly represent a 3D process with a 1D model. Outgoing radiation is calculated as the result of multiple processes involving the scattering due to the canopy layer and the forest floor. Finally, surface albedo is computed as the ratio between incident solar radiation and calculated outgoing radiation. The study used two time series representing thinning from below of a beech and a Scots pine forest. The results show a strong temporal evolution in albedo during stand establishment followed by a relatively stable albedo once the canopy is closed. During this period, albedo is affected for a short time by forest operations. The modelling approach allowed us to estimate the importance of ground vegetation in the stand albedo. Given that ground vegetation depends on the light reaching the forest floor, ground vegetation could act as a natural buffer to dampen changes in albedo, allowing the stand to maintain optimal leaf temperature. Consequently, accounting for only the carbon balance component of forest management ignores albedo impacts and is thus likely to yield biased estimates of the climate benefits of forest ecosystems.

  6. Recovery of aboveground plant biomass and productivity after fire in mesic and dry black spruce forests of interior Alaska

    USGS Publications Warehouse

    Mack, M.C.; Treseder, K.K.; Manies, K.L.; Harden, J.W.; Schuur, E.A.G.; Vogel, J.G.; Randerson, J.T.; Chapin, F. S.

    2008-01-01

    Plant biomass accumulation and productivity are important determinants of ecosystem carbon (C) balance during post-fire succession. In boreal black spruce (Picea mariana) forests near Delta Junction, Alaska, we quantified aboveground plant biomass and net primary productivity (ANPP) for 4 years after a 1999 wildfire in a well-drained (dry) site, and also across a dry and a moderately well-drained (mesic) chronosequence of sites that varied in time since fire (2 to ???116 years). Four years after fire, total biomass at the 1999 burn site had increased exponentially to 160 ?? 21 g m-2 (mean ?? 1SE) and vascular ANPP had recovered to 138 ?? 32 g m-2 y -1, which was not different than that of a nearby unburned stand (160 ?? 48 g m-2 y-1) that had similar pre-fire stand structure and understory composition. Production in the young site was dominated by re-sprouting graminoids, whereas production in the unburned site was dominated by black spruce. On the dry and mesic chronosequences, total biomass pools, including overstory and understory vascular and non-vascular plants, and lichens, increased logarithmically (dry) or linearly (mesic) with increasing site age, reaching a maximum of 2469 ?? 180 (dry) and 4008 ?? 233 g m-2 (mesic) in mature stands. Biomass differences were primarily due to higher tree density in the mesic sites because mass per tree was similar between sites. ANPP of vascular and non-vascular plants increased linearly over time in the mesic chronosequence to 335 ?? 68 g m-2 y -1 in the mature site, but in the dry chronosequence it peaked at 410 ?? 43 g m-2 y-1 in a 15-year-old stand dominated by deciduous trees and shrubs. Key factors regulating biomass accumulation and production in these ecosystems appear to be the abundance and composition of re-sprouting species early in succession, the abundance of deciduous trees and shrubs in intermediate aged stands, and the density of black spruce across all stand ages. A better understanding of the controls over these factors will help predict how changes in climate and fire regime will affect the carbon balance of Interior Alaska. ?? 2008 Springer Science+Business Media, LLC.

  7. Time spent in sedentary posture is associated with waist circumference and cardiovascular risk.

    PubMed

    Tigbe, W W; Granat, M H; Sattar, N; Lean, M E J

    2017-05-01

    The relationship between metabolic risk and time spent sitting, standing and stepping has not been well established. The present study aimed to determine associations of objectively measured time spent siting, standing and stepping, with coronary heart disease (CHD) risk. A cross-sectional study of healthy non-smoking Glasgow postal workers, n=111 (55 office workers, 5 women, and 56 walking/delivery workers, 10 women), who wore activPAL physical activity monitors for 7 days. Cardiovascular risks were assessed by metabolic syndrome categorisation and 10-year PROCAM (prospective cardiovascular Munster) risk. Mean (s.d.) age was 40 (8) years, body mass index 26.9 (3.9) kg m -2 and waist circumference 95.4 (11.9) cm. Mean (s.d.) high-density lipoprotein cholesterol (HDL cholesterol) 1.33 (0.31), low-density lipoprotein cholesterol 3.11 (0.87), triglycerides 1.23 (0.64) mmol l -1 and 10-year PROCAM risk 1.8 (1.7)%. The participants spent mean (s.d.) 9.1 (1.8) h per day sedentary, 7.6 (1.2) h per day sleeping, 3.9 (1.1) h per day standing and 3.3 (0.9) h per day stepping, accumulating 14 708 (4984) steps per day in 61 (25) sit-to-stand transitions per day. In univariate regressions-adjusting for age, sex, family history of CHD, shift worked, job type and socioeconomic status-waist circumference (P=0.005), fasting triglycerides (P=0.002), HDL cholesterol (P=0.001) and PROCAM risk (P=0.047) were detrimentally associated with sedentary time. These associations remained significant after further adjustment for sleep, standing and stepping in stepwise regression models. However, after further adjustment for waist circumference, the associations were not significant. Compared with those without the metabolic syndrome, participants with the metabolic syndrome were significantly less active-fewer steps, shorter stepping duration and longer time sitting. Those with no metabolic syndrome features walked >15 000 steps per day or spent >7 h per day upright. Longer time spent in sedentary posture is significantly associated with higher CHD risk and larger waist circumference.

  8. Canopy transpiration of two black locust (Robinia pseudoacacia) plantations with different ages in semi-arid Loess Plateau, China

    NASA Astrophysics Data System (ADS)

    Jiao, L.

    2015-12-01

    Black locust (Robinia pseudoacacia) was widely planted to control soil erosion and restore degraded ecosystem in Loess Plateau. The water use of the plantations was concerned due to its potential effects on hydrological cycle and regional water resource. Although some studies estimated canopy transpiration (Ec) of the mature black locust plantation, variation in Ec in plantations with different ages was not clear. In this study, we selected two plantations with different ages (12 years and 27 years, denoted as young stand and mature stand, respectively) in similar topographical conditions in Yangjuangou catchment in the central of Loess Plateau. Sap flux density (Fd) and tree biometrics were measured in each stand during the growing season in 2014. Soil water content (SWC) in each plot and meteorological variables in the catchment were simultaneously monitored. Tree transpiration (Et) was derived from Fd and tree sapwood area (As). Canopy transpiration (Ec) was estimated by a product of mean stand sap flux density (Js) and stand total sapwood area (AST). The mean Fd of mature trees was 2-fold larger than that of young trees.However, tree-to-tree variation in Fd among sampled trees within mature stand was evident compared to that within young stand. Mean Et in mature stand was higher than that in young stand. Ec in mature stand was significant higher than that in young stand,with cumulative value of 54 mm and 27 mm respectively. This is attributed to higher Js in mature stand although AST in young is slightly higher than that in mature stand. The patterns of daily Ec during the growing season were similar in both stands during the study period. A exponential saturation model can explain the responses of Ec to vapor deficit pressure (VPD) and solar radiation (Rs) in both stands.The relationship between Ec and SWC was not detected. Our finding suggested that stand age should be taken into consideration when estimated vegetation water use in this region. Further researches on variations in Ec of black locust plantations along a chronosequence and responses of Ec to drought should be conducted to provide useful information on water resource and forest managements in this region.

  9. Characteristic of Secondary Flow Caused by Local Density Change in Standing Acoustic Fields

    NASA Astrophysics Data System (ADS)

    Tonsho, Kazuyuki; Hirosawa, Takuya; Kusakawa, Hiroshi; Kuwahara, Takuo; Tanabe, Mitsuaki

    Secondary flow is a flow which is caused by the interference between standing acoustic fields and local density change. The behavior of the secondary flow depends on the location of the given local density change in the standing acoustic fields. When the density change is given at the middle of a velocity node and the neighboring velocity anti-node (middle point) or when it is given at the velocity anti-node in standing acoustic fields, the secondary flow shows particular behavior. Characteristic of the secondary flow at the two positions was predicted by numerical simulations. It was examined from these simulations whether the driving mechanism of the flow can be explained by the kind of acoustic radiation force that has been proposed so far. The predicted secondary flow was verified by experiments. For both the simulations and experiments, the standing acoustic fields generated in a cylinder are employed. In the experiments, the acoustic fields are generated by two loud speakers that are vibrated in same phase in a chamber. The employed resonance frequency is about 1000 Hz. The chamber is filled with air of room temperature and atmospheric pressure. In the numerical simulations and experiments, the local density change is given by heating or cooling. Because the secondary flow is influenced by buoyancy, the numerical simulations were done without taking gravity force into account and a part of the experiments were done by the microgravity condition using a drop tower. As a result of the simulations, at the middle point, the heated air was blown toward the node and the cooled air was blown toward the anti-node. It is clarified that the secondary flow is driven by the expected kind of acoustic radiation force. At the anti-node, both the heated and cooled air expands perpendicular to the traveling direction of the sound wave. The driving mechanism of the secondary flow can not be explained by the acoustic radiation force, and a detailed analysis is done. Through the comparison between experimental and numerical results, it was verified that the secondary flow is qualitatively predictable by the numerical simulations.

  10. [Effects of tree species diversity on fine-root biomass and morphological characteristics in subtropical Castanopsis carlesii forests].

    PubMed

    Wang, Wei-Wei; Huang, Jin-Xue; Chen, Feng; Xiong, De-Cheng; Lu, Zheng-Li; Huang, Chao-Chao; Yang, Zhi-Jie; Chen, Guang-Shui

    2014-02-01

    Fine roots in the Castanopsis carlesii plantation forest (MZ), the secondary forest of C. carlesii through natural regeneration with anthropogenic promotion (AR), and the secondary forest of C. carlesii through natural regeneration (NR) in Sanming City, Fujian Province, were estimated by soil core method to determine the influence of tree species diversity on biomass, vertical distribution and morphological characteristics of fine roots. The results showed that fine root biomass for the 0-80 cm soil layer in the MZ, AR and NR were (182.46 +/- 10.81), (242.73 +/- 17.85) and (353.11 +/- 16.46) g x m(-2), respectively, showing an increased tendency with increasing tree species diversity. In the three forests, fine root biomass was significantly influenced by soil depth, and fine roots at the 0-10 cm soil layer accounted for more than 35% of the total fine root biomass. However, the interaction of stand type and soil depth on fine-root distribution was not significant, indicating no influence of tree species diversity on spatial niche segregation in fine roots. Root surface area density and root length density were the highest in NR and lowest in the MZ. Specific root length was in the order of AR > MZ > NR, while specific root surface area was in the order of NR > MZ > AR. There was no significant interaction of stand type and soil depth on specific root length and specific root surface area. Fine root morphological plasticity at the stand level had no significant response to tree species diversity.

  11. Anatomical explanations for acute depressions in radial pattern of axial sap flow in two diffuse-porous mangrove species: implications for water use.

    PubMed

    Zhao, Hewei; Yang, Shengchang; Guo, Xudong; Peng, Congjiao; Gu, Xiaoxuan; Deng, Chuanyuan; Chen, Luzhen

    2018-02-01

    Mangrove species have developed uniquely efficient water-use strategies in order to survive in highly saline and anaerobic environments. Herein, we estimated the stand water use of two diffuse-porous mangrove species of the same age, Sonneratia apetala Buch. Ham and Sonneratia caseolaris (L.) Engl., growing in a similar intertidal environment. Specifically, to investigate the radial patterns of axial sap flow density (Js) and understand the anatomical traits associated with them, we measured axial sap flow density in situ together with micromorphological observations. A significant decrease of Js was observed for both species. This result was accompanied by the corresponding observations of wood structure and blockages in xylem sapwood, which appeared to influence and, hence, explained the acute radial reductions of axial sap flow in the stems of both species. However, higher radial resistance in sapwood of S. caseolaris caused a steeper decline of Js radially when compared with S. apetala, thus explaining the latter's more efficient use of water. Without first considering acute reductions in Js into the sapwood from the outer bark, a total of ~55% and 51% of water use would have been overestimated, corresponding to average discrepancies in stand water use of 5.6 mm day-1 for S. apetala trees and 2.5 mm day-1 for S. caseolaris trees. This suggests that measuring radial pattern of Js is a critical factor in determining whole-tree or stand water use. © The Author(s) 2018. Published by Oxford University Press. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

  12. Association mapping of rice cold germination with the USDA mini-core

    USDA-ARS?s Scientific Manuscript database

    Assuring stand establishment is a critical first step in optimizing rice crop yields. Plant stand density can impact yield potential, incidence of some diseases, weed competition, and grain quality. Most rice production in the Southern USA is drill seeded in the spring. Planting can occur as early a...

  13. Free-standing, well-aligned ordered mesoporous carbon nanofibers on current collectors for high-power micro-supercapacitors.

    PubMed

    Kang, Eunae; Jeon, Gumhye; Kim, Jin Kon

    2013-07-21

    The mesoporous carbon nanofiber arrays that stand on carbon-gold double-layer current collectors are synthesized by self-assembly of a PS-b-PEO copolymer and resol in AAO templates for a high-power micro-supercapacitor at high current densities.

  14. Fifteen-year results from six cutting methods in second-growth northern hardwoods.

    Treesearch

    Gayne G. Erdmann; Robert R. Oberg

    1973-01-01

    Presents and compares stand growth and yield information from three single-tree selection cuts, a crop-tree release treatment, an 8-inch diameter limit cut, and an uncut control. Discusses the influence of stand density on basal area growth, cubic volume growth, and board-foot volume growth.

  15. Uneven-aged silviculture and management in the western United States: Proceedings of an In-service Workshop; Redding, California; October 19-21, 1976

    Treesearch

    Carl M. Berntsen; A. P. Mustian; Carter B. Gibbs; David A. Marquis; Donald T. Gordon; Jerry F. Franklin; Marvin W. Foiles; Robert O. Curtis; Dale O. Hall; Robert R. Alexander; Carleton B. Edminster; Robert E. Phares

    1977-01-01

    The 1976 National Silviculture Workshop was held in Eugene, Oregon, on October 13-15, 1976. The objectives were to discuss second growth management of individual stands, with particular emphasis on the control of stand density.

  16. Prohexadione-calcium improves stand density and yield of alfalfa interseeded into silage corn

    USDA-ARS?s Scientific Manuscript database

    Interseeded alfalfa (Medicago sativa L.) could serve as a dual-purpose crop to provide groundcover for silage corn (Zea mays L.) and forage during subsequent years of production, but interspecific competition often leads to poor stands of alfalfa and unsatisfactory yields of corn. Four experiments e...

  17. Height growth and site index curves for managed even-aged stands of ponderosa pine in the Pacific Northwest

    Treesearch

    James W. Barrett

    1978-01-01

    This paper presents height growth and site index curves and equations for even-aged, managed stands of ponderosa pine east of the Cascade Range in Oregon and Washington where height growth has not been suppressed by high density or related factors.

  18. A Test of the Exponential Distribution for Stand Structure Definition in Uneven-aged Loblolly-Shortleaf Pine Stands

    Treesearch

    Paul A. Murphy; Robert M. Farrar

    1981-01-01

    In this study, 588 before-cut and 381 after-cut diameter distributions of uneven-aged loblolly-shortleaf pinestands were fitted to two different forms of the exponential probability density function. The left truncated and doubly truncated forms of the exponential were used.

  19. Aspen Ecology in Rocky Mountain National Park: Age Distribution, Genetics, and the Effects of Elk Herbivory

    USGS Publications Warehouse

    Zeigenfuss, Linda C.; Binkley, Dan; Tuskan, Gerald A.; Romme, William H.; Yin, Tongming; DiFazio, Stephen; Singer, Francis J.

    2008-01-01

    Lack of recruitment and canopy replacement of aspen (Populus tremuloides) stands that grow on the edges of grasslands on the low-elevation elk (Cervus elaphus) winter range of Rocky Mountain National Park (RMNP) in Colorado have been a cause of concern for more than 70 years. We used a combination of traditional dendrochronology and genetic techniques as well as measuring the characteristics of regenerating and nonregenerating stands on the elk winter range to determine when and under what conditions and estimated elk densities these stands established and through what mechanisms they may regenerate. The period from 1975 to 1995 at low elevation on the east side had 80-95 percent fewer aspen stems than would be expected based on the trend from 1855 through 1965. The age structure of aspen in the park indicates that the interacting effects of fires, elk population changes, and livestock grazing had more-or-less consistent effects on aspen from 1855 to 1965. The lack of a significant change in aspen numbers in recent decades in the higher elevation and west side parts of the park supports the idea that the extensive effects of elk browsing have been more important in reducing aspen numbers than other factors. The genetic variation of aspen populations in RMNP is high at the molecular level. We expected to find that most patches of aspen in the park were composed of a single clone of genetically identical trees, but in fact just 7 percent of measured aspen patches consisted of a single clone. A large frequency of polyploid (triploid and tetraploid) genotypes were found on the low elevation, east-side elk winter range. Nonregenerating aspen stands on the winter range had greater annual offtake, shorter saplings, and lower density of mid-height (1.5-2.5 m) saplings than regenerating stands. Overwinter elk browsing, however, did not appear to inhibit the leader length of aspen saplings. The winter range aspen stands of RMNP appear to be highly resilient in the face of very intense herbivory by elk and harsh environmental conditions. Conservation efforts through fencing protection and decreased elk browsing pressure are already being planned as part of the park's new elk management plan. If these efforts are undertaken, conditions that encourage stem recruitment to the tree canopy will likely result and the continued survival of these aspen stands will be enhanced.

  20. Pollen flow in fragmented landscapes maintains genetic diversity following stand-replacing disturbance in a neotropical pioneer tree, Vochysia ferruginea Mart.

    PubMed

    Davies, S J; Cavers, S; Finegan, B; White, A; Breed, M F; Lowe, A J

    2015-08-01

    In forests with gap disturbance regimes, pioneer tree regeneration is typically abundant following stand-replacing disturbances, whether natural or anthropogenic. Differences in pioneer tree density linked to disturbance regime can influence pollinator behaviour and impact on mating patterns and genetic diversity of pioneer populations. Such mating pattern shifts can manifest as higher selfing rates and lower pollen diversity in old growth forest populations. In secondary forest, where more closely related pollen donors occur, an increase in biparental inbreeding is a potential problem. Here, we investigate the consequences of secondary forest colonisation on the mating patterns and genetic diversity of open-pollinated progeny arrays for the long-lived, self-compatible pioneer tree, Vochysia ferruginea, at two Costa Rican sites. Five microsatellite loci were screened across adult and seed cohorts from old growth forest with lower density, secondary forest with higher density, and isolated individual trees in pasture. Progeny from both old growth and secondary forest contexts were predominantly outcrossed (tm=1.00) and experienced low levels of biparental inbreeding (tm-ts=0.00-0.04). In contrast to predictions, our results indicated that the mating patterns of V. ferruginea are relatively robust to density differences between old growth and secondary forest stands. In addition, we observed that pollen-mediated gene flow possibly maintained the genetic diversity of open-pollinated progeny arrays in stands of secondary forest adults. As part of a natural resource management strategy, we suggest that primary forest remnants should be prioritised for conservation to promote restoration of genetic diversity during forest regeneration.

  1. Modal Damping Ratio and Optimal Elastic Moduli of Human Body Segments for Anthropometric Vibratory Model of Standing Subjects.

    PubMed

    Gupta, Manoj; Gupta, T C

    2017-10-01

    The present study aims to accurately estimate inertial, physical, and dynamic parameters of human body vibratory model consistent with physical structure of the human body that also replicates its dynamic response. A 13 degree-of-freedom (DOF) lumped parameter model for standing person subjected to support excitation is established. Model parameters are determined from anthropometric measurements, uniform mass density, elastic modulus of individual body segments, and modal damping ratios. Elastic moduli of ellipsoidal body segments are initially estimated by comparing stiffness of spring elements, calculated from a detailed scheme, and values available in literature for same. These values are further optimized by minimizing difference between theoretically calculated platform-to-head transmissibility ratio (TR) and experimental measurements. Modal damping ratios are estimated from experimental transmissibility response using two dominant peaks in the frequency range of 0-25 Hz. From comparison between dynamic response determined form modal analysis and experimental results, a set of elastic moduli for different segments of human body and a novel scheme to determine modal damping ratios from TR plots, are established. Acceptable match between transmissibility values calculated from the vibratory model and experimental measurements for 50th percentile U.S. male, except at very low frequencies, establishes the human body model developed. Also, reasonable agreement obtained between theoretical response curve and experimental response envelop for average Indian male, affirms the technique used for constructing vibratory model of a standing person. Present work attempts to develop effective technique for constructing subject specific damped vibratory model based on its physical measurements.

  2. Organic horizon and mineral soil mercury along three clear-cut forest chronosequences across the northeastern USA.

    PubMed

    Richardson, Justin B; Petrenko, Chelsea L; Friedland, Andrew J

    2017-12-01

    Mercury (Hg) is a globally distributed pollutant trace metal that has been increasing in terrestrial environments due to rising anthropogenic emissions. Vegetation plays an important role in Hg sequestration in forested environments, but increasing tree removal for biofuels and wood products may affect this process. The long-term effect of clear-cutting on forest soil Hg remains uncertain, since most studies are limited to measuring changes for < 10 years following a single harvest event. The chronosequence approach, which substitutes space for time using forest stands of different ages since clear-cutting, allows for investigation of processes occurring over decades to centuries. Here, we utilized three clear-cut forest soil chronosequences across the northeastern USA to understand Hg accumulation and retention over several decades. Total Hg concentrations and pools were quantified for five soil depth increments along three chronosequences. Our results showed Hg concentrations and pools decreased in the initial 20 years following clear-cutting. Mineral soil Hg pools decreased 21-53% (7-14 mg m -2 ) between 1-5-year-old stands and 15-25-year-old stands but mineral soil Hg pools recovered in 55-140-year-old stands to similar values as measured in 1-5-year-old stands. Our study is one of the first to demonstrate a decrease and recovery in Hg pool size. These changes in Hg did not correspond with changes in bulk density, soil C, or pH. We utilized a simple two-box model to determine how different Hg fluxes affected organic and mineral soil horizon Hg pools. Our simple model suggests that changes in litterfall and volatilization rates could have caused the observed changes in organic horizon Hg pools. However, only increases in leaching could reproduce observed decreases to mineral soil Hg pools. Further studies are needed to determine the mechanism of Hg loss from forest soils following clear-cutting.

  3. Accounting for density reduction and structural loss in standing dead trees: Implications for forest biomass and carbon stock estimates in the United States

    Treesearch

    Grant M. Domke; Christopher W. Woodall; James E. Smith

    2011-01-01

    Standing dead trees are one component of forest ecosystem dead wood carbon (C) pools, whose national stock is estimated by the U.S. as required by the United Nations Framework Convention on Climate Change. Historically, standing dead tree C has been estimated as a function of live tree growing stock volume in the U.S.'s National Greenhouse Gas Inventory. Initiated...

  4. Species composition, diameter distribution, and crown class at initiation of a thinning study of pole-size hardwood stands in the Hoosier National Forest

    Treesearch

    Ryan L. Woods; Douglass F. Jacobs

    2008-01-01

    During the spring of 2007, a low thinning was implemented in stands on the Hoosier National Forest that had been clearcut harvested between 1975 and 1979; treatments consisted of 60- and 75-percent residual stocking, as well as control plots with no thinning. The 60-percent treatment increased the relative oak density per acre in all stands with the exception of one...

  5. Oscillations of a standing shock wave generated by the Richtmyer-Meshkov instability

    DOE PAGES

    Mikaelian, Karnig O.

    2016-07-13

    In a typical Richtmyer-Meshkov experiment a fast moving flat shock strikes a stationary perturbed interface between fluids A and B creating a transmitted and a reflected shock, both of which are perturbed. We propose shock tube experiments in which the reflected shock is stationary in the laboratory. Such a standing perturbed shock undergoes well-known damped oscillations. We present the conditions required for producing such a standing shock wave, which greatly facilitates the measurement of the oscillations and their rate of damping. We define a critical density ratio R critical, in terms of the adiabatic indices of the two fluids, andmore » a critical Mach number M critical s of the incident shock wave, which produces a standing reflected wave. If the initial density ratio R of the two fluids is less than R critical then a standing shock wave is possible at M s=M critical s. Otherwise a standing shock is not possible and the reflected wave always moves in the direction opposite the incident shock. Examples are given for present-day operating shock tubes with sinusoidal or inclined interfaces. We consider the effect of viscosity, which affects the damping rate of the oscillations. Furthermore, we point out that nonlinear bubble and spike amplitudes depend relatively weakly on the viscosity of the fluids and that the interface area is a better diagnostic.« less

  6. Snag densities in relation to human access and associated management factors in forests of Northeastern Oregon, USA

    Treesearch

    Lisa J. Bate; Michael J. Wisdom; Barbara C. Wales

    2007-01-01

    A key element of forest management is the maintenance of sufficient densities of snags (standing dead trees) to support associated wildlife. Management factors that influence snag densities, however, are numerous and complex. Consequently, accurate methods to estimate and model snag densities are needed. Using data collected in 2002 and Current Vegetation Survey (CVS)...

  7. Using a Density-Management Diagram to Develop Thinning Schedules for Loblolly Pine Plantations

    Treesearch

    Thomas J. Dean; V. Clark Baldwin

    1993-01-01

    A method for developing thinning schedules using a density-management diagram is presented. A density-management diagram is a form of stocking chart based on patterns of natural stand development. The diagram allows rotation diameter and the upper and lower limits of growing stock to be easily transformed into before and after thinning densities. Site height lines on...

  8. Plant interactions alter the predictions of metabolic scaling theory.

    PubMed

    Lin, Yue; Berger, Uta; Grimm, Volker; Huth, Franka; Weiner, Jacob

    2013-01-01

    Metabolic scaling theory (MST) is an attempt to link physiological processes of individual organisms with macroecology. It predicts a power law relationship with an exponent of -4/3 between mean individual biomass and density during density-dependent mortality (self-thinning). Empirical tests have produced variable results, and the validity of MST is intensely debated. MST focuses on organisms' internal physiological mechanisms but we hypothesize that ecological interactions can be more important in determining plant mass-density relationships induced by density. We employ an individual-based model of plant stand development that includes three elements: a model of individual plant growth based on MST, different modes of local competition (size-symmetric vs. -asymmetric), and different resource levels. Our model is consistent with the observed variation in the slopes of self-thinning trajectories. Slopes were significantly shallower than -4/3 if competition was size-symmetric. We conclude that when the size of survivors is influenced by strong ecological interactions, these can override predictions of MST, whereas when surviving plants are less affected by interactions, individual-level metabolic processes can scale up to the population level. MST, like thermodynamics or biomechanics, sets limits within which organisms can live and function, but there may be stronger limits determined by ecological interactions. In such cases MST will not be predictive.

  9. Effects of the seasonal flooding on riparian soil seed bank in the Three Gorges Reservoir Region: a case study in Shanmu River.

    PubMed

    Zhang, Miao; Chen, Fangqing; Chen, Shaohua; Wang, Yajin; Wang, Jianzhu

    2016-01-01

    The water-level fluctuation in the Three Gorges Reservoir Region has changed dramatically as a result of the hydroelectric project for flood control and power generation. The riparian seasonal hydrological environment also has changed from summer flooding with winter drought to summer drought with winter flooding. The changes of riparian seed bank and vegetation were investigated to determine the effects of the seasonal flooding on the composition and spatial distribution of riparian soil seed bank and the similarity of seed bank to standing vegetation. We conducted intensive riparian soil sampling (525 samples) along altitude gradient in the Shanmu River, a tributary of the Yangzi River in the reservoir region of China. Seed bank density, species richness and composition of soil seed bank were examined using the seedling-emergence method. The seasonal hydrological conditions resulted in a decrease in species diversity and an increase in the distribution heterogeneity of the soil seed bank. The soil seed bank was composed of 48 species from 22 families and 40 genera. Most species were annual and perennial herbaceous Polygonaceae, Asteraceae, and Poaceae. Rumex dentatus was the predominant species accounting for 27.0 % of the total seeds. Diversity and composition of the seed bank changed along an altitude gradient and soil depth. Maximum species richness was found in the top soil layer at 165 m and 175 m above sea level. The mean overall seed density of the soil seed bank was 13,475.3 ind m(-2). Density and the number of seeds increased initially and then decreased with increased altitude. Maximum seed density (22,500.2 ind m(-2)) was found at 165 m above sea level in the intermediately flooded riverbank, with the seed number accounting for 27.8 % of the total soil seed bank. Average seed density declined significantly with soil depth. The similarity of seed bank to standing vegetation was relatively high. The environmental heterogeneity created by the wide range and seasonal flooding led to the changes in biodiversity and seed density along altitude gradient. The seasonal flooding also led to the increase in the similarity of seed bank to standing vegetation as their composition both degraded. The seasonal flooding due to the dam reshape the composition and spatial distribution of riparian soil seed bank and limit the vegetation to a grassland dominated by a few annuals and perennials in the Three Gorges Reservoir Region.

  10. Regulation and control of intracellular algae (= zooxanthellae) in hard corals

    PubMed Central

    Jones, R. J.; Yellowlees, D.

    1997-01-01

    To examine algal (= zooxanthellae) regulation and control, and the factors determining algal densities in hard corals, the zooxanthellae mitotic index and release rates were regularly determined in branch tips from a colony of a staghorn coral, Acropora formosa, recovering from a coral 'bleaching' event (the stress-related dissociation of the coral–algal symbiosis). Mathematical models based upon density-dependent decreases in the algal division frequency and increases in algal release rates during the post-bleaching recovery period accurately predict the observed recovery period (ca. 20 weeks). The models suggest that (i) the colony recovered its algal population from the division of the remaining zooxanthellae, and (ii) the continual loss of zooxanthellae significantly slowed the recovery of the coral. Possible reasons for the 'paradoxical' loss of healthy zooxanthellae from the bleached coral are discussed in terms of endodermal processes occurring in the recovering coral and the redistribution of newly formed zooxanthellae to aposymbiotic host cells. At a steady-state algal density of 2.1 x 106 zooxanthellae cm-2 at the end of the recovery period, the zooxanthellae would have to form a double layer of cells in the coral tissues, consistent with microscopic observations. Neighbouring colonies of A. formosa with inherently higher algal densities possess proportionately smaller zooxanthellae. Results suggest that space availability and the size of the algal symbionts determines the algal densities in the coral colonies. The large increases in the algal densities reported in corals exposed to elevated nutrient concentrations (i.e between a two- and five-fold increase in the algal standing stock) are not consistent with this theory. We suggest that increases of this magnitude are a product of the experimental conditions: reasons for this statement are discussed. We propose that the stability of the coral–algal symbiosis under non-stress conditions, and the constancy of zooxanthellae densities in corals reported across growth form, depth and geographic range, are related to space availability limiting algal densities. However, at these densities, zooxanthellae have attributes consistent with nutrient limitation.

  11. 5 CFR 351.401 - Determining retention standing.

    Code of Federal Regulations, 2012 CFR

    2012-01-01

    ... 5 Administrative Personnel 1 2012-01-01 2012-01-01 false Determining retention standing. 351.401 Section 351.401 Administrative Personnel OFFICE OF PERSONNEL MANAGEMENT CIVIL SERVICE REGULATIONS REDUCTION IN FORCE Scope of Competition § 351.401 Determining retention standing. Each agency shall...

  12. 5 CFR 351.401 - Determining retention standing.

    Code of Federal Regulations, 2010 CFR

    2010-01-01

    ... 5 Administrative Personnel 1 2010-01-01 2010-01-01 false Determining retention standing. 351.401 Section 351.401 Administrative Personnel OFFICE OF PERSONNEL MANAGEMENT CIVIL SERVICE REGULATIONS REDUCTION IN FORCE Scope of Competition § 351.401 Determining retention standing. Each agency shall...

  13. 5 CFR 351.401 - Determining retention standing.

    Code of Federal Regulations, 2014 CFR

    2014-01-01

    ... 5 Administrative Personnel 1 2014-01-01 2014-01-01 false Determining retention standing. 351.401 Section 351.401 Administrative Personnel OFFICE OF PERSONNEL MANAGEMENT CIVIL SERVICE REGULATIONS REDUCTION IN FORCE Scope of Competition § 351.401 Determining retention standing. Each agency shall...

  14. 5 CFR 351.401 - Determining retention standing.

    Code of Federal Regulations, 2013 CFR

    2013-01-01

    ... 5 Administrative Personnel 1 2013-01-01 2013-01-01 false Determining retention standing. 351.401 Section 351.401 Administrative Personnel OFFICE OF PERSONNEL MANAGEMENT CIVIL SERVICE REGULATIONS REDUCTION IN FORCE Scope of Competition § 351.401 Determining retention standing. Each agency shall...

  15. 5 CFR 351.401 - Determining retention standing.

    Code of Federal Regulations, 2011 CFR

    2011-01-01

    ... 5 Administrative Personnel 1 2011-01-01 2011-01-01 false Determining retention standing. 351.401 Section 351.401 Administrative Personnel OFFICE OF PERSONNEL MANAGEMENT CIVIL SERVICE REGULATIONS REDUCTION IN FORCE Scope of Competition § 351.401 Determining retention standing. Each agency shall...

  16. Structural and compositional controls on transpiration in 40- and 450-year-old riparian forests in western Oregon, USA.

    PubMed

    Moore, Georgianne W; Bond, Barbara J; Jones, Julia A; Phillips, Nathan; Meinzer, Federick C

    2004-05-01

    Large areas of forests in the Pacific Northwest are being transformed to younger forests, yet little is known about the impact this may have on hydrological cycles. Previous work suggests that old trees use less water per unit leaf area or sapwood area than young mature trees of the same species in similar environments. Do old forests, therefore, use less water than young mature forests in similar environments, or are there other structural or compositional components in the forests that compensate for tree-level differences? We investigated the impacts of tree age, species composition and sapwood basal area on stand-level transpiration in adjacent watersheds at the H.J. Andrews Forest in the western Cascades of Oregon, one containing a young, mature (about 40 years since disturbance) conifer forest and the other an old growth (about 450 years since disturbance) forest. Sap flow measurements were used to evaluate the degree to which differences in age and species composition affect water use. Stand sapwood basal area was evaluated based on a vegetation survey for species, basal area and sapwood basal area in the riparian area of two watersheds. A simple scaling exercise derived from estimated differences in water use as a result of differences in age, species composition and stand sapwood area was used to estimate transpiration from late June through October within the entire riparian area of these watersheds. Transpiration was higher in the young stand because of greater sap flux density (sap flow per unit sapwood area) by age class and species, and greater total stand sapwood area. During the measurement period, mean daily sap flux density was 2.30 times higher in young compared with old Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) trees. Sap flux density was 1.41 times higher in young red alder (Alnus rubra Bong.) compared with young P. menziesii trees, and was 1.45 times higher in old P. menziesii compared with old western hemlock (Tsuga heterophylla (Raf.) Sarg.) trees. Overall, sapwood basal area was 21% higher in the young stand than in the old stand. In the old forest, T. heterophylla is an important co-dominant, accounting for 58% of total sapwood basal area, whereas P. menziesii is the only dominant conifer in the young stand. Angiosperms accounted for 36% of total sapwood basal area in the young stand, but only 7% in the old stand. For all factors combined, we estimated 3.27 times more water use by vegetation in the riparian area of the young stand over the measurement period. Tree age had the greatest effect on stand differences in water use, followed by differences in sapwood basal area, and finally species composition. The large differences in transpiration provide further evidence that forest management alters site water balance via elevated transpiration in vigorous young stands.

  17. Comparison of snag densities among regeneration treatments in mixed pine-hardwood forests

    Treesearch

    Roger W. Perry; Ronald E. Thill

    2013-01-01

    Standing dead trees (snags) are an important component of forest ecosystems, providing foraging, nesting, and roosting substrate for a variety of vertebrates. We examined the effects of four forest regeneration treatments on residual snag density and compared those with densities found in unharvested, naturally regenerated forests (controls) during the second, fourth,...

  18. Thinning Jeffrey pine stands to reduce susceptibility to bark beetle infestations in California, U.S.A.

    Treesearch

    Christopher J. Fettig; Christopher J. Hayes; Karen J. Jones; Stephen R. McKelvey; Sylvia L. Mori; Sheri L. Smith

    2012-01-01

    Bark beetles (Coleoptera: Curculionidae, Scolytinae) are commonly recognized as important tree mortality agents in coniferous forests of the western U.S.A.High stand density is consistently associated with bark beetle infestations in western coniferous forests, and therefore, thinning has long been advocated as a...

  19. Establishment and growth of native hardwood and conifer seedlings underplanted in thinned Douglas-fir stands.

    Treesearch

    Kathleen G. Maas-Hebner; William H. Emmingham; David L. Larson; Samuel S. Chan

    2005-01-01

    Five conifers and two hardwoods native to the Pacific Northwest were planted under four overstory densities of 30-year-old plantations of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) in the Oregon Coast Range, USA. Stand treatments were unthinned (547 trees ha-1), narrow thin (252 trees ha-1),...

  20. A silvicultural guide for developing a sugarbush

    Treesearch

    Kenneth F. Lancaster; Russell S. Walters; Frederick M. Laing; Raymond T. Foulds

    1974-01-01

    A practical guide for the management of a sugarbush. Guidelines are established for the manipulation of stand density and stocking to promote the development of healthy vigorous trees with deep, wide crowns, the necessary attributes for highest possible yield of sugar-rich sap. Specific treatments are prescribed for sapling, poletimber and small sawtimber stands and a...

  1. LANDIS PRO: a landscape model that predicts forest composition and structure changes at regional scales

    Treesearch

    Wen J. Wang; Hong S. He; Jacob S. Fraser; Frank R. Thompson; Stephen R. Shifley; Martin A. Spetich

    2014-01-01

    LANDIS PRO predicts forest composition and structure changes incorporating species-, stand-, and landscape-scales processes at regional scales. Species-scale processes include tree growth, establishment, and mortality. Stand-scale processes contain density- and size-related resource competition that regulates self-thinning and seedling establishment. Landscapescale...

  2. Growth and yield in Eucalyptus globulus

    Treesearch

    James A. Rinehart; Richard B. Standiford

    1983-01-01

    A study of the major Eucalyptus globulus stands throughout California conducted by Woodbridge Metcalf in 1924 provides a complete and accurate data set for generating variable site-density yield models. Two models were developed using linear regression techniques. Model I depicts a linear relationship between age and yield best used for stands between five and fifteen...

  3. Sustaining recruitment of oak reproduction in uneven-aged stands in the Ozark Highlands.

    Treesearch

    David R. Larsen; Edward F. Loewenstein; Paul S. Johnson

    1999-01-01

    Successful application of the single-tree selection system in Ozark oak forests depends on sustaining adequate recruitment of reproduction into the overstory. In turn, this requires maintaining stand density at ecologically appropriate levels. The ecological requirements for oak recruitment are discussed and guiding curves are presented that meet those requirements...

  4. Area change reporting using the desktop FIADB

    Treesearch

    Patrick D. Miles; Mark H. Hansen

    2012-01-01

    The estimation of area change between two FIA inventories is complicated by the "mapping" of subplots. Subplots can be subdivided or mapped into forest and nonforest conditions, and forest conditions can be further mapped based on distinct changes in reserved status, owner group, forest type, stand-size class, regeneration status, and stand density. The...

  5. Forest Stand Canopy Structure Attribute Estimation from High Resolution Digital Airborne Imagery

    Treesearch

    Demetrios Gatziolis

    2006-01-01

    A study of forest stand canopy variable assessment using digital, airborne, multispectral imagery is presented. Variable estimation involves stem density, canopy closure, and mean crown diameter, and it is based on quantification of spatial autocorrelation among pixel digital numbers (DN) using variogram analysis and an alternative, non-parametric approach known as...

  6. Functional response of ungulate browsers in disturbed eastern hemlock forests

    USGS Publications Warehouse

    DeStefano, Stephen

    2015-01-01

    Ungulate browsing in predator depleted North American landscapes is believed to be causing widespread tree recruitment failures. However, canopy disturbances and variations in ungulate densities are sources of heterogeneity that can buffer ecosystems against herbivory. Relatively little is known about the functional response (the rate of consumption in relation to food availability) of ungulates in eastern temperate forests, and therefore how “top down” control of vegetation may vary with disturbance type, intensity, and timing. This knowledge gap is relevant in the Northeastern United States today with the recent arrival of hemlock woolly adelgid (HWA; Adelges tsugae) that is killing eastern hemlocks (Tsuga canadensis) and initiating salvage logging as a management response. We used an existing experiment in central New England begun in 2005, which simulated severe adelgid infestation and intensive logging of intact hemlock forest, to examine the functional response of combined moose (Alces americanus) and white-tailed deer (Odocoileus virginianus) foraging in two different time periods after disturbance (3 and 7 years). We predicted that browsing impacts would be linear or accelerating (Type I or Type III response) in year 3 when regenerating stem densities were relatively low and decelerating (Type II response) in year 7 when stem densities increased. We sampled and compared woody regeneration and browsing among logged and simulated insect attack treatments and two intact controls (hemlock and hardwood forest) in 2008 and again in 2012. We then used AIC model selection to compare the three major functional response models (Types I, II, and III) of ungulate browsing in relation to forage density. We also examined relative use of the different stand types by comparing pellet group density and remote camera images. In 2008, total and proportional browse consumption increased with stem density, and peaked in logged plots, revealing a Type I response. In 2012, stem densities were greatest in girdled plots, but proportional browse consumption was highest at intermediate stem densities in logged plots, exhibiting a Type III (rather than a Type II) functional response. Our results revealed shifting top–down control by herbivores at different stages of stand recovery after disturbance and in different understory conditions resulting from logging vs. simulated adelgid attack. If forest managers wish to promote tree regeneration in hemlock stands that is more resistant to ungulate browsers, leaving HWA-infested stands unmanaged may be a better option than preemptively logging them.

  7. Models for Predicting the Biomass of Cunninghamialanceolata Trees and Stands in Southeastern China

    PubMed Central

    Saeed, Sajjad

    2017-01-01

    Using existing equations to estimate the biomass of a single tree or a forest stand still involves large uncertainties. In this study, we developed individual-tree biomass models for Chinese Fir (Cunninghamia lanceolata.) stands in Fujian Province, southeast China, by using 74 previously established models that have been most commonly used to estimate tree biomass. We selected the best fit models and modified them. The results showed that the published model ln(B(Biomass)) = a + b * ln(D) + c * (ln(H))2 + d * (ln(H))3 + e * ln(WD) had the best fit for estimating the tree biomass of Chinese Fir stands. Furthermore, we observed that variables D(diameter at breast height), H (height), and WD(wood density)were significantly correlated with the total tree biomass estimation model. As a result, a natural logarithm structure gave the best estimates for the tree biomass structure. Finally, when a multi-step improvement on tree biomass model was performed, the tree biomass model with Tree volume(TV), WD and biomass wood density conversion factor (BECF),achieved the highest simulation accuracy, expressed as ln(TB) = −0.0703 + 0.9780 * ln(TV) + 0.0213 * ln(WD) + 1.0166 * ln(BECF). Therefore, when TV, WD and BECF were combined with tree biomass volume coefficient bi for Chinese Fir, the stand biomass (SB)model included both volume(SV) and coefficient bi variables of the stand as follows: bi = Exp(−0.0703+0.9780*ln(TV)+0.0213 * ln(WD)+1.0166*ln(BECF)). The stand biomass model is SB = SV/TV * bi. PMID:28095512

  8. Models for Predicting the Biomass of Cunninghamialanceolata Trees and Stands in Southeastern China.

    PubMed

    Guangyi, Mei; Yujun, Sun; Saeed, Sajjad

    2017-01-01

    Using existing equations to estimate the biomass of a single tree or a forest stand still involves large uncertainties. In this study, we developed individual-tree biomass models for Chinese Fir (Cunninghamia lanceolata.) stands in Fujian Province, southeast China, by using 74 previously established models that have been most commonly used to estimate tree biomass. We selected the best fit models and modified them. The results showed that the published model ln(B(Biomass)) = a + b * ln(D) + c * (ln(H))2 + d * (ln(H))3 + e * ln(WD) had the best fit for estimating the tree biomass of Chinese Fir stands. Furthermore, we observed that variables D(diameter at breast height), H (height), and WD(wood density)were significantly correlated with the total tree biomass estimation model. As a result, a natural logarithm structure gave the best estimates for the tree biomass structure. Finally, when a multi-step improvement on tree biomass model was performed, the tree biomass model with Tree volume(TV), WD and biomass wood density conversion factor (BECF),achieved the highest simulation accuracy, expressed as ln(TB) = -0.0703 + 0.9780 * ln(TV) + 0.0213 * ln(WD) + 1.0166 * ln(BECF). Therefore, when TV, WD and BECF were combined with tree biomass volume coefficient bi for Chinese Fir, the stand biomass (SB)model included both volume(SV) and coefficient bi variables of the stand as follows: bi = Exp(-0.0703+0.9780*ln(TV)+0.0213 * ln(WD)+1.0166*ln(BECF)). The stand biomass model is SB = SV/TV * bi.

  9. Compensating effect of sap velocity for stand density leads to uniform hillslope-scale forest transpiration across a steep valley cross-section

    NASA Astrophysics Data System (ADS)

    Renner, Maik; Hassler, Sibylle; Blume, Theresa; Weiler, Markus; Hildebrandt, Anke; Guderle, Marcus; Schymanski, Stan; Kleidon, Axel

    2016-04-01

    Roberts (1983) found that forest transpiration is relatively uniform across different climatic conditions and suggested that forest transpiration is a conservative process compensating for environmental heterogeneity. Here we test this hypothesis at a steep valley cross-section composed of European Beech in the Attert basin in Luxemburg. We use sapflow, soil moisture, biometric and meteorological data from 6 sites along a transect to estimate site scale transpiration rates. Despite opposing hillslope orientation, different slope angles and forest stand structures, we estimated relatively similar transpiration responses to atmospheric demand and seasonal transpiration totals. This similarity is related to a negative correlation between sap velocity and site-average sapwood area. At the south facing sites with an old, even-aged stand structure and closed canopy layer, we observe significantly lower sap velocities but similar stand-average transpiration rates compared to the north-facing sites with open canopy structure, tall dominant trees and dense understorey. This suggests that plant hydraulic co-ordination allows for flexible responses to environmental conditions leading to similar transpiration rates close to the water and energy limits despite the apparent heterogeneity in exposition, stand density and soil moisture. References Roberts, J. (1983). Forest transpiration: A conservative hydrological process? Journal of Hydrology 66, 133-141.

  10. Wood density of young-growth western hemlock: relation to ring age, radial growth, stand density, and site quality.

    Treesearch

    Dean S. DeBell; Ryan Singleton; Barbara L. Gartner; David D. Marshall

    2004-01-01

    Breast-high stem sections were sampled from 56 western hemlock (Tsuga heterophylla (Raf.) Sarg.) trees growing in 15 plots representing a wide range of tree and site conditions in northwestern Oregon. Growth and wood density traits of individual rings were measured via X-ray densitometry, and relationships of ring density and its components to age...

  11. 78 FR 41942 - Standing Rock Sioux Tribe; Major Disaster and Related Determinations

    Federal Register 2010, 2011, 2012, 2013, 2014

    2013-07-12

    .... FEMA-4123-DR; Docket ID FEMA-2013-0001] Standing Rock Sioux Tribe; Major Disaster and Related... Presidential declaration of a major disaster for the Standing Rock Sioux Tribe (FEMA-4123-DR), dated June 25...''), as follows: I have determined that the damage to the lands associated with the Standing Rock Sioux...

  12. Succession after stand replacing disturbances by fire, wind throw, and insects in the dark Taiga of Central Siberia.

    PubMed

    Schulze, E-D; Wirth, C; Mollicone, D; Ziegler, W

    2005-11-01

    The dark taiga of Siberia is a boreal vegetation dominated by Picea obovata, Abies sibirica, and Pinus sibirica during the late succession. This paper investigates the population and age structure of 18 stands representing different stages after fire, wind throw, and insect damage. To our knowledge, this is the first time that the forest dynamics of the Siberian dark taiga is described quantitatively in terms of succession, and age after disturbance, stand density, and basal area. The basis for the curve-linear age/diameter relation of trees is being analyzed. (1) After a stand-replacing fire Betula dominates (4,000 trees) for about 70 years. Although tree density of Betula decreases rapidly, basal area (BA) reached >30 m2/ha after 40 years. (2) After fire, Abies, Picea, and Pinus establish at the same time as Betula, but grow slower, continue to gain height and eventually replace Betula. Abies has the highest seedling number (about 1,000 trees/ha) and the highest mortality. Picea establishes with 100-400 trees/ha, it has less mortality, but reached the highest age (>350 years, DBH 51 cm). Picea is the most important indicator for successional age after disturbance. Pinus sibirica is an accompanying species. The widely distributed "mixed boreal forest" is a stage about 120 years after fire reaching a BA of >40 m2/ha. (3) Wind throw and insect damage occur in old conifer stands. Betula does not establish. Abies initially dominates (2,000-6,000 trees/ha), but Picea becomes dominant after 150-200 years since Abies is shorter lived. (4) Without disturbance the forest develops into a pure coniferous canopy (BA 40-50 m2/ha) with a self-regenerating density of 1,000 coniferous canopy trees/ha. There is no collapse of old-growth stands. The dark taiga may serve as an example in which a limited set to tree species may gain dominance under certain disturbance conditions without ever getting monotypic.

  13. Acoustic radiation force on a rigid elliptical cylinder in plane (quasi)standing waves

    NASA Astrophysics Data System (ADS)

    Mitri, F. G.

    2015-12-01

    The acoustic radiation force on a 2D elliptical (non-circular) cylinder centered on the axis of wave propagation of plane quasi-standing and standing waves is derived, based on the partial-wave series expansion (PWSE) method in cylindrical coordinates. A non-dimensional acoustic radiation force function, which is the radiation force per unit length, per characteristic energy density and per unit cross-sectional surface of the ellipse, is defined in terms of the scattering coefficients that are determined by applying the Neumann boundary condition for an immovable surface. A system of linear equations involving a single numerical integration procedure is solved by matrix inversion. Numerical simulations showing the transition from the quasi-standing to the (equi-amplitude) standing wave behaviour are performed with particular emphasis on the aspect ratio a/b, where a and b are the ellipse semi-axes, as well as the dimensionless size parameter kb (where k is the wavenumber), without the restriction to a particular range of frequencies. It is found that at high kb values > 1, the radiation force per length with broadside incidence is larger, whereas the opposite situation occurs in the long-wavelength limit (i.e., kb < 1). The results are particularly relevant in acoustic levitation of elliptical cylinders, the acoustic stabilization of liquid columns in a host medium, acousto-fluidics devices, and other particle dynamics applications to name a few. Moreover, the formalism presented here may be effectively applied to compute the acoustic radiation force on other 2D surfaces of arbitrary shape such as super-ellipses, Chebyshev cylindrical particles, or other non-circular geometries.

  14. Free-standing 3D polyaniline-CNT/Ni-fiber hybrid electrodes for high-performance supercapacitors

    NASA Astrophysics Data System (ADS)

    Li, Yuan; Fang, Yuzhu; Liu, Hong; Wu, Xiaoming; Lu, Yong

    2012-04-01

    Free-standing 3D macroscopic polyaniline (PANi)-carbon nanotube (CNT)-nickel fiber hybrids have been developed, and they deliver high specific capacitance (725 F g-1 at 0.5 A g-1) and high energy density at high rates (~22 W h kg-1 at 2000 W kg-1, based on total electrode mass) with good cyclability.Free-standing 3D macroscopic polyaniline (PANi)-carbon nanotube (CNT)-nickel fiber hybrids have been developed, and they deliver high specific capacitance (725 F g-1 at 0.5 A g-1) and high energy density at high rates (~22 W h kg-1 at 2000 W kg-1, based on total electrode mass) with good cyclability. Electronic supplementary information (ESI) available: Experimental details on preparation, characterization, and electrochemical testing; Fig. S1-S8, Schemes S1 and S2. See DOI: 10.1039/c2nr30252g

  15. Assessment of a model of forest dynamics under contrasting climate and disturbance regimes in the Pacific Northwest [FORCLIM

    USGS Publications Warehouse

    Busing, Richard T.; Solomon, Allen M.

    2005-01-01

    An individual-based model of forest dynamics (FORCLIM) was tested for its ability to simulate forest composition and structure in the Pacific Northwest region of North America. Simulation results across gradients of climate and disturbance were compared to forest survey data from several vegetation zones in western Oregon. Modelled patterns of tree species composition, total basal area and stand height across climate gradients matched those in the forest survey data. However, the density of small stems (<50 cm DBH) was underestimated by the model. Thus actual size-class structure and other density-based parameters of stand structure were not simulated with high accuracy. The addition of partial-stand disturbances at moderate frequencies (<0.01 yr-1) often improved agreement between simulated and actual results. Strengths and weaknesses of the FORCLIM model in simulating forest dynamics and structure in the Pacific Northwest are discussed.

  16. Competition amplifies drought stress in forests across broad climatic and compositional gradients

    USGS Publications Warehouse

    Gleason, Kelly; Bradford, John B.; Bottero, Alessandra; D'Amato, Tony; Fraver, Shawn; Palik, Brian J.; Battaglia, Michael; Iverson, Louis R.; Kenefic, Laura; Kern, Christel C.

    2017-01-01

    Forests around the world are experiencing increasingly severe droughts and elevated competitive intensity due to increased tree density. However, the influence of interactions between drought and competition on forest growth remains poorly understood. Using a unique dataset of stand-scale dendrochronology sampled from 6405 trees, we quantified how annual growth of entire tree populations responds to drought and competition in eight, long-term (multi-decadal), experiments with replicated levels of density (e.g., competitive intensity) arrayed across a broad climatic and compositional gradient. Forest growth (cumulative individual tree growth within a stand) declined during drought, especially during more severe drought in drier climates. Forest growth declines were exacerbated by high density at all sites but one, particularly during periods of more severe drought. Surprisingly, the influence of forest density was persistent overall, but these density impacts were greater in the humid sites than in more arid sites. Significant density impacts occurred during periods of more extreme drought, and during warmer temperatures in the semi-arid sites but during periods of cooler temperatures in the humid sites. Because competition has a consistent influence over growth response to drought, maintaining forests at lower density may enhance resilience to drought in all climates.

  17. [Effects of land use changes on soil water conservation in Hainan Island, China].

    PubMed

    Wen, Zhi; Zhao, He; Liu, Lei; OuYang, Zhi Yun; Zheng, Hua; Mi, Hong Xu; Li, Yan Min

    2017-12-01

    In tropical areas, a large number of natural forests have been transformed into other plantations, which affected the water conservation function of terrestrial ecosystems. In order to clari-fy the effects of land use changes on soil water conservation function, we selected four typical land use types in the central mountainous region of Hainan Island, i.e., natural forests with stand age greater than 100 years (VF), secondary forests with stand age of 10 years (SF), areca plantations with stand age of 12 years (AF) and rubber plantations with stand age of 35 years (RF). The effects of land use change on soil water holding capacity and water conservation (presented by soil water index, SWI) were assessed. The results showed that, compared with VF, the soil water holding capacity index of other land types decreased in the top soil layer (0-10 cm). AF had the lowest soil water holding capacity in all soil layers. Soil water content and maximum water holding capacity were significantly related to canopy density, soil organic matter and soil bulk density, which indicated that canopy density, soil organic matter and compactness were important factors influencing soil water holding capacity. Compared to VF, soil water conservation of SF, AF and RF were reduced by 27.7%, 54.3% and 11.5%, respectively. The change of soil water conservation was inconsistent in different soil layers. Vegetation canopy density, soil organic matter and soil bulk density explained 83.3% of the variance of soil water conservation. It was suggested that land use conversion had significantly altered soil water holding capacity and water conservation function. RF could keep the soil water better than AF in the research area. Increasing soil organic matter and reducing soil compaction would be helpful to improve soil water holding capacity and water conservation function in land management.

  18. Impact Of Standing Water On Saltstone Placement II - Hydraulic Conductivity Data

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Cozzi, A. D.; Pickenheim, B. R.

    2012-12-06

    The amount of water present during placement and subsequent curing of saltstone has the potential to impact several properties important for grout quality. An active drain water system can remove residual standing water and expose the surface of the placed saltstone to air. Oxidation of the saltstone may result in an increase in the leachability of redox sensitive elements. A dry surface can lead to cracking, causing an increase in hydraulic conductivity. An inactive drain water system can allow standing water that generates unnecessary hydrostatic head on the vault walls. Standing water that cannot be removed via the drain systemmore » will be available for potential incorporation into subsequent grout placements. The objective of this work is to study the impact of standing water on grout quality pertaining to disposal units. A series of saltstone mixes was prepared and cured at ambient temperature to evaluate the impact of standing water on saltstone placement. The samples were managed to control drying effects on leachability by either exposing or capping the samples. The water to premix ratio was varied to represent a range of processing conditions. Samples were analyzed for density, leachability, and hydraulic conductivity. Report SRNL-STI-2012-00546 was issued detailing the experimental procedure, results, and conclusions related to density and leachability. In the previous report, it was concluded that: density tends to increase toward the bottom of the samples. This effect is pronounced with excess bleed water; drying of the saltstone during curing leads to decreased Leachability Index (more leaching) for potassium, sodium, rhenium, nitrite, and nitrate; there is no noticeable effect on saltstone oxidation/leachability by changing the water to premix ratio (over the range studied), or by pouring into standing water (when tested up to 10 volume percent). The hydraulic conductivity data presented in this report show that samples cured exposed to the atmosphere had about three orders of magnitude higher hydraulic conductivity than any of the other samples. Considering these data, along with the results presented in the previous report, leads to the conclusion that small changes in water to premix ratio and the inclusion of up to 10 volume percent standing water should not be expected to have a detrimental effect on saltstone grout quality. The hydraulic conductivity results further demonstrate that curing in a moist environment is critical to maintaining saltstone quality.« less

  19. Fine root dynamics in lodgepole pine and white spruce stands along productivity gradients in reclaimed oil sands sites.

    PubMed

    Jamro, Ghulam Murtaza; Chang, Scott X; Naeth, M Anne; Duan, Min; House, Jason

    2015-10-01

    Open-pit mining activities in the oil sands region of Alberta, Canada, create disturbed lands that, by law, must be reclaimed to a land capability equivalent to that existed before the disturbance. Re-establishment of forest cover will be affected by the production and turnover rate of fine roots. However, the relationship between fine root dynamics and tree growth has not been studied in reclaimed oil sands sites. Fine root properties (root length density, mean surface area, total root biomass, and rates of root production, turnover, and decomposition) were assessed from May to October 2011 and 2012 using sequential coring and ingrowth core methods in lodgepole pine (Pinus contorta Dougl.) and white spruce (Picea glauca (Moench.) Voss) stands. The pine and spruce stands were planted on peat mineral soil mix placed over tailings sand and overburden substrates, respectively, in reclaimed oil sands sites in Alberta. We selected stands that form a productivity gradient (low, medium, and high productivities) of each tree species based on differences in tree height and diameter at breast height (DBH) increments. In lodgepole pine stands, fine root length density and fine root production, and turnover rates were in the order of high > medium > low productivity sites and were positively correlated with tree height and DBH and negatively correlated with soil salinity (P < 0.05). In white spruce stands, fine root surface area was the only parameter that increased along the productivity gradient and was negatively correlated with soil compaction. In conclusion, fine root dynamics along the stand productivity gradients were closely linked to stand productivity and were affected by limiting soil properties related to the specific substrate used for reconstructing the reclaimed soil. Understanding the impact of soil properties on fine root dynamics and overall stand productivity will help improve land reclamation outcomes.

  20. Cross-scale interactions affect tree growth and intrinsic water ...

    EPA Pesticide Factsheets

    1. We investigated the potential of cross-scale interactions to affect the outcome of density reduction in a large-scale silvicultural experiment. 2. We measured tree growth and intrinsic water-use efficiency (iWUE) based on stable carbon isotopes (13C) to investigate the impacts of thinning across a range of progressively finer spatial scales: site, stand, hillslope position, and neighborhood position. In particular, we focused on the influence of thinning beyond the boundaries of thinned stands to include impacts on downslope and neighboring stands across sites varying in soil moisture. 3. Trees at the wet site responded to thinning with increased growth when compared with trees at the dry site. Additionally, trees in thinned stands at the dry site responded with increased iWUE while trees in thinned stands at the wet site showed no difference in iWUE compared to unthinned stands. 4. We hypothesized that water is not the primary limiting factor for growth at our sites, but that thinning released other resources, such as growing space or nutrients to drive the growth response. At progressively finer spatial scales we found that the responses of trees was not driven by hillslope location (i.e., downslope of thinning) but to changes in local neighborhood tree density. 5. The results of this study demonstrated that water can be viewed as an “agent” that allows us to investigate cross-scale interactions as it links coarse to finer spatial scales and vice ver

  1. Energy density of lake whitefish Coregonus clupeaformis in Lakes Huron and Michigan

    USGS Publications Warehouse

    Pothoven, S.A.; Nalepa, T.F.; Madenjian, C.P.; Rediske, R.R.; Schneeberger, P.J.; He, J.X.

    2006-01-01

    We collected lake whitefish Coregonus clupeaformis off Alpena and Tawas City, Michigan, USA in Lake Huron and off Muskegon, Michigan USA in Lake Michigan during 2002–2004. We determined energy density and percent dry weight for lake whitefish from both lakes and lipid content for Lake Michigan fish. Energy density increased with increasing fish weight up to 800 g, and then remained relatively constant with further increases in fish weight. Energy density, adjusted for weight, was lower in Lake Huron than in Lake Michigan for both small (≤800 g) and large fish (>800 g). Energy density did not differ seasonally for small or large lake whitefish or between adult male and female fish. Energy density was strongly correlated with percent dry weight and percent lipid content. Based on data from commercially caught lake whitefish, body condition was lower in Lake Huron than Lake Michigan during 1981–2003, indicating that the dissimilarity in body condition between the lakes could be long standing. Energy density and lipid content in 2002–2004 in Lake Michigan were lower than data for comparable sized fish collected in 1969–1971. Differences in energy density between lakes were attributed to variation in diet and prey energy content as well as factors that affect feeding rates such as lake whitefish density and prey abundance.

  2. Best density and structure for uneven-aged northern hardwood management in New England

    Treesearch

    William B. Leak

    2003-01-01

    Choice of the best residual density (basal area per acre) and structure (diameter distribution) for uneven-aged management of northern hardwoods is a complex decision that depends on the manager's decision rules, product objectives, site conditions, and - perhaps most important - current stand conditions. In contrast to other recommendations on residual density...

  3. Western spruce budworm as related to stand characteristics in the bitterroot national forest

    Treesearch

    Carroll B. Williams; Patrick J. Shea; Gerald S. Walton

    1971-01-01

    Relation of population density to certain stand conditions and damage indicators was analyzed in four drainages on the Bitterroot National Forest of Montana. Western spruce budworm (Choristoneura occidentalis Freeman) populations were strongly related to plot basal area, tree species, and tree crown levels, and also to current and past levels of tree defoliation....

  4. Wood volume increment in thinned, 50- to 55-year-old, mixed-species Allegheny hardwoods

    Treesearch

    Christopher A. Nowak

    1996-01-01

    A thinning study in 50- to 55-year-old, even-aged, mixed species Allegheny hardwoods produced highly variable merchantable stemwood volume increment responses. Regression equations relating parameters of stand growth (ingrowth, mortality, survivor growth, net growth, and gross growth) to relative stand density had R2 values ranging from 0.07 to 0...

  5. Log sampling methods and software for stand and landscape analyses.

    Treesearch

    Lisa J. Bate; Torolf R. Torgersen; Michael J. Wisdom; Edward O. Garton; Shawn C. Clabough

    2008-01-01

    We describe methods for efficient, accurate sampling of logs at landscape and stand scales to estimate density, total length, cover, volume, and weight. Our methods focus on optimizing the sampling effort by choosing an appropriate sampling method and transect length for specific forest conditions and objectives. Sampling methods include the line-intersect method and...

  6. Forest restoration at Redwood National Park: exploring prescribed fire alternatives to second-growth management: a case study

    Treesearch

    Eamon Engber; Jason Teraoka; Phil van Mantgem

    2017-01-01

    Almost half of Redwood National Park is comprised of second-growth forests characterized by high stand density, deficient redwood composition, and low understory biodiversity. Typical structure of young redwood stands impedes the recovery of old-growth conditions, such as dominance of redwood (Sequoia sempervirens (D. Don) Endl.), distinct...

  7. Lumber recovery from small-diameter ponderosa pine from Flagstaff, Arizona

    Treesearch

    Eini C. Lowell; David W. Green

    2001-01-01

    Thousands of acres of densely stocked ponderosa pine forests surround Flagstaff, AZ. These stands are at high risk of fire, insect, and disease outbreak. Stand density management activity can be expensive, but product recovery from the thinned material could help defray removal costs. This project evaluated the yield and economic return of lumber recovered from small-...

  8. 76 FR 65681 - Black Hills National Forest, Mystic Ranger District, South Dakota, Calumet Project Area

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-10-24

    ... acres of pine stands using a variety of methods to treat MPB infested stands, reduce the overall density... actions proposed are in direct response to management direction provided by the Black Hills National Forest Land and Resource Management Plan (Forest Plan). The site specific actions are designed, based on...

  9. Forcasting Shortleaf Pine Seed Crops in the Ouachita Mountains

    Treesearch

    Michael G. Shelton; Robert F. Wittwer

    2004-01-01

    We field tested a cone-rating system to forecast seed crops from 1993 to 1996 in 28 shortleaf pine (Pinus echinata Mill.) stands, which represented a wide range of stand conditions. Sample trees were visually assigned to one of three cone-density classes based on cone spacing, occurrence of cones in clusters, and distribution of cones within the...

  10. More than just timber: silvicultural options and ecosystem services from managed southern pine stands

    Treesearch

    Don C. Bragg; Jamie L. Schuler; Matthew H Pelkki; D. Andrew Scott; James M. Guldin

    2015-01-01

    The dramatic decline of timber harvests on public lands in the western United States (U.S.) has helped intensify silviculture in the southern U.S. Today, intensive southern pine management usually involves establishment of plantations using site preparation, genetically improved seedlings, chemical fertilization and competition control, early stand density regulation,...

  11. Season of burn and hardwood development in young longleaf pine stands

    Treesearch

    William D. Boyer

    1993-01-01

    Abstract.Four treatmems--bienniaI burns in winter, spring, and summer, and a no-bum control-were applied in plots in naturally established stands of longleaf pine (Pinus palustris) Mill.). Treatments commenced in 1974, when the pines were 15 years old, and the most recent observations were made in 1992. Midstory hardwood density...

  12. Responses of hardwood advace regeneration to seasonal prescribed fires in oak-dominated shelterwood stands

    Treesearch

    Patrick H. Brose; David H. Van Lear

    1998-01-01

    Effects of seasonal prescribed fires of varying intensities on density, mortality, stem form, height, and height growth of hardwood advance regeneration were investigated. Three mixed-hardwood stands on productive upland sites were cut using a shelterwood technique, each forming a block of spring burn, summer burn, winter burn, and control treatments. Advance...

  13. Diameter Growth 0f a Slash Pine Spacing Study Five Years After Being Thinned to a Constant Stand Density Index

    Treesearch

    Jamie C. Schexnayder; Thomas J. Dean; V. Clark Baldwin

    2002-01-01

    Abstract - In 1994, a 17-year old, slash pine (Pinus elliottii var. elliottii) spacing study was thinned to evaluate the influence of prethinning stand conditions on diameter growth after thinning. Diameter growth and crown dimensions measured just prior to thinning showed that diameter growth was positively...

  14. A frontier shortleaf pine stand in the old-growth Cross Timbers of Oklahoma

    Treesearch

    K. Chris Cerny; David W. Stahle; Don C. Bragg

    2016-01-01

    We investigated an old-growth oak-shortleaf pine (Pinus echinata) stand of high ecological integrity in east-central Oklahoma located west of the continuous native distribution of shortleaf pine. With the exception of an abundance of shortleaf pine, the basal area (17.2 m2/ha), density (559.6 trees/ha), and species...

  15. Diet of southern toads (Bufo terrestris) in loblolly pine (Pinus taeda) stands subject to coarse woody debris manipulations

    Treesearch

    Kurtis R. Moseley; Steven B. Castleberry; James L. Hanula; W. Mark Ford

    2004-01-01

    In the southeastern United States, coarse woody debris (CWD) typically harbors high densities of invertebrates. However, its importance as a foraging substrate for southeastern amphibians is relatively unknown. We examined effects of CWD manipulations on diet composition of southern toads (Bufo tmrestns) in upland loblolly pine (Pinus taeda) stands...

  16. Development of a central hardwood stand following whole-tree clearcutting in Connecticut

    Treesearch

    C. Wayne Martin

    1995-01-01

    Little information is available concerning the initial stages of forest regeneration following intensive harvesting of central hardwood stands in the northeastern part of the range. Establishment of commercial species, density, and rate of biomass accumulation of the regeneration are of major concern to both foresters and landowners contemplating a harvest. To help...

  17. Effect of stand density and structure on the abundance of northern red oak advance reproduction

    Treesearch

    Gary W. Miller

    1997-01-01

    Regenerating northern red oak (Quercus rubra L.) on high-quality growing sites is a continuing problem in the central Appalachian region. Competing species usually exhibit faster height growth after regeneration harvests compared to oak reproduction. The probability of advance oak reproduction becoming codominant in the new stand is positively...

  18. The Shadow Knows: Inferring the Density Distribution of a Nonuniform Medium from Its Standing Wave Pattern

    ERIC Educational Resources Information Center

    Binder, Philippe; Cunnyngham, Ian

    2012-01-01

    In a recent note in this journal, Gluck presents a beautiful demonstration of the standing wave generated by a strip of material with linearly varying width (a trapezoid). As expected, the resulting wave envelope (and its shadow) showed a varying wavelength--smaller as the strip width gets larger.

  19. Growing stock levels in even-aged ponderosa pine

    Treesearch

    Clifford A. Myers

    1967-01-01

    Growth of the most widely distributed pine in North America is under joint study by the western Forest and Range Experiment Stations of the U. S. Forest Service. Young, even-aged ponderosa pine (Pinus ponderosa Laws.) stands are being examined over a wide range of tree sizes, stand densities, and site index. The single plan that co-...

  20. Long-term trends in height growth of jack pine in North Central Ontario

    Treesearch

    J.C.G. Goelz; T.E. Burk

    1998-01-01

    Although most investigations of long-term growth trends of trees involve description of radial growth of trees, investigation of height growth of dominant and codominant trees also warrants attention for two significant reasons -- the dependent variable is largely independent of stand density and it represents an index of stand productivity. Residuals from a height...

  1. Silvicultural methods for regenerating oaks

    Treesearch

    F. Bryan Clark; Richard F. Watt

    1971-01-01

    Advance reproduction is the key to forming the new oak stand. However, the size or strength of the advance stems is just as important as number. Most oak stands approaching maturity have enough advance reproduction, but many do not. In such cases, harvest cuttings must be delayed and overstory densities regulated to favor the establishment and development of new oaks...

  2. Effects of alternative treatments on canopy fuel characteristics in five conifer stands

    Treesearch

    Joe H. Scott; Elizabeth D. Reinhardt

    2007-01-01

    A detailed study of canopy fuel characteristics in five different forest types provided a unique dataset for simulating the effects of various stand manipulation treatments on canopy fuels. Low thinning, low thinning with commercial dbh limit, and crown thinning had similar effects on canopy bulk density (CBD) and canopy fuel load (CFL...

  3. Distribution and dynamics of hayscented fern following stand harvest

    Treesearch

    Songlin Fei; Peter J. Gould; Melanie J. Kaeser; Kim C. Steiner

    2008-01-01

    The distribution and dynamics of hayscented fern were examined as part of a large-scale study of oak regeneration in Pennsylvania. The study included 69 stands covering 3,333 acres in two physiographic provinces. Hayscented fern was more widely distributed and occurred at higher densities in the Allegheny Plateau physiographic provinces versus the Ridge and Valley...

  4. The carbon consequences of thinning techniques: stand structure makes a difference

    Treesearch

    Coeli Hoover; Susan Stout

    2007-01-01

    Using results from a 25-year study of thinning in a northwestern Pennsylvania Allegheny hardwood stand, we assess whether and how thinning method affected carbon sequestration and merchantable volume production. Plots were thinned to similar residual relative density by removing trees from different portions of the diameter distribution. Plots that were thinned from...

  5. Forest inventory and management-based visual preference models of southern pine stands

    Treesearch

    Victor A. Rudis; James H. Gramann; Edward J. Ruddell; Joanne M. Westphal

    1988-01-01

    Statistical models explaining students' ratings of photographs of within stand forest scenes were constructed for 99 forest inventory plots in east Texas pine and oak-pine forest types. Models with parameters that are sensitive to visual preference yet compatible with forest management and timber inventories are presented. The models suggest that the density of...

  6. Extrafloral Nectaries in Aspen (Populus tremuloides): Heritable Genetic Variation and Herbivore-induced Expression

    PubMed Central

    Wooley, Stuart C.; Donaldson, Jack R.; Gusse, Adam C.; Lindroth, Richard L.; Stevens, Michael T.

    2007-01-01

    Background and Aims A wide variety of plants produce extrafloral nectaries (EFNs) that are visited by predatory arthropods. But very few studies have investigated the relationship between plant genetic variation and EFNs. The presence of foliar EFNs is highly variable among different aspen (Populus tremuloides) genotypes and the EFNs are visited by parasitic wasps and predatory flies. The aim here was to determine the heritability of EFNs among aspen genotypes and age classes, possible trade-offs between direct and indirect defences, EFN induction following herbivory, and the relationship between EFNs and predatory insects. Methods EFN density was quantified among aspen genotypes in Wisconsin on trees of different ages and broad-sense heritability from common garden trees was calculated. EFNs were also quantified in natural aspen stands in Utah. From the common garden trees foliar defensive chemical levels were quantified to evaluate their relationship with EFN density. A defoliation experiment was performed to determine if EFNs can be induced in response to herbivory. Finally, predatory arthropod abundance among aspen trees was quantified to determine the relationship between arthropod abundance and EFNs. Key Results Broad-sense heritability for expression (0·74–0·82) and induction (0·85) of EFNs was high. One-year-old trees had 20% greater EFN density than 4-year-old trees and more than 50% greater EFN density than ≥10-year-old trees. No trade-offs were found between foliar chemical concentrations and EFN density. Predatory fly abundance varied among aspen genotypes, but predatory arthropod abundance and average EFN density were not related. Conclusions Aspen extrafloral nectaries are strongly genetically determined and have the potential to respond rapidly to evolutionary forces. The pattern of EFN expression among different age classes of trees appears to follow predictions of optimal defence theory. The relationship between EFNs and predators likely varies in relation to multiple temporal and environmental factors. PMID:17951361

  7. Coronal loop seismology using damping of standing kink oscillations by mode coupling. II. additional physical effects and Bayesian analysis

    NASA Astrophysics Data System (ADS)

    Pascoe, D. J.; Anfinogentov, S.; Nisticò, G.; Goddard, C. R.; Nakariakov, V. M.

    2017-04-01

    Context. The strong damping of kink oscillations of coronal loops can be explained by mode coupling. The damping envelope depends on the transverse density profile of the loop. Observational measurements of the damping envelope have been used to determine the transverse loop structure which is important for understanding other physical processes such as heating. Aims: The general damping envelope describing the mode coupling of kink waves consists of a Gaussian damping regime followed by an exponential damping regime. Recent observational detection of these damping regimes has been employed as a seismological tool. We extend the description of the damping behaviour to account for additional physical effects, namely a time-dependent period of oscillation, the presence of additional longitudinal harmonics, and the decayless regime of standing kink oscillations. Methods: We examine four examples of standing kink oscillations observed by the Atmospheric Imaging Assembly (AIA) onboard the Solar Dynamics Observatory (SDO). We use forward modelling of the loop position and investigate the dependence on the model parameters using Bayesian inference and Markov chain Monte Carlo (MCMC) sampling. Results: Our improvements to the physical model combined with the use of Bayesian inference and MCMC produce improved estimates of model parameters and their uncertainties. Calculation of the Bayes factor also allows us to compare the suitability of different physical models. We also use a new method based on spline interpolation of the zeroes of the oscillation to accurately describe the background trend of the oscillating loop. Conclusions: This powerful and robust method allows for accurate seismology of coronal loops, in particular the transverse density profile, and potentially reveals additional physical effects.

  8. Effects of cultural intensity and density regime treatment on post-thinning loblolly pine individual tree DBH increment in the lower coastal plain of the southeastern United States

    Treesearch

    John T. Perren; Michael Kane; Dehai Zhao; Richard Daniels

    2016-01-01

    Thinning is a well understood concept used to manage density dependent factors at the stand level. This study evaluates the effect of planting density, cultural intensity, and thinning treatment on loblolly pine post-thinning individual tree development. The Lower Coastal Plain Culture/Density Study, has four initial densities, in combination with two cultural...

  9. [Effects of stand density on understory species diversity and soil physicochemical properties of Pinus massoniana plantation].

    PubMed

    Sun, Qian Hui; Wu, Xia; Wang, Mei Zhen; Zhang, Liu Hua; Yao, Xiao Lan; Qi, Jin Qiu; Hao, Jian Feng

    2018-03-01

    We analyzed understory species diversity, soil physicochemical traits and their relationships in the 25-year-old non-commercial Pinus massoniana plantations with five different stand densities, i.e., 1057, 1136, 1231, 1383 and 1515 trees·hm -2 , in Wenfeng Mountain, Xinjin District, Sichuan Province, China. The results showed that a total of 110 species were found, belonging to 57 families and 98 genera. With the increase of tree density, the understory species showed a succession pattern from positive to moderate to shady. Different densities had significant effects on the contents of total potassium and organic matter in the soils. With the increase ofdensity, the contents of organic matter and total potassium in understory vegetation first increased and then decreased. The trends of the relationship between both diversity and soil physiochemical characteristics and tree density were similar. Both of them increased with the increase of density, with the maximum value presented at the density of 1136 trees·hm -2 . The concentrations of total phosphorus, available potassium, total potassium and total nitrogen was closely related to plant diversity index. The results suggested that the density at 1136 trees·hm -2 would be more beneficial to improve the stability of species diversity and soil fertility of P. massoniana non-commercial plantations in Wenfeng Mountain.

  10. Status and habitat relationships of northern flying squirrels on Mount Desert Island, Maine

    USGS Publications Warehouse

    O'Connell, A.F.; Servello, F.; Higgins, J.; Halteman, W.

    2001-01-01

    Northern (Glaucomys sabrinus) and southern (G. volans) flying squirrels occur in Maine, but there is uncertainty about range overlap in southcentral Maine where the southern flying squirrel reaches its geographic range limit. We surveyed flying squirrels on Mount Desert Island (MDI), located along the central Maine coast, to update the current status and distribution of these species. We captured only northern flying squirrels, and populations (> 2 individuals) were located in two conifer stands and one mixed conifer-hardwood stand. All three stands were located in relatively older forests, outside a large area burned in a 1947 fire. Tree diameters were similar between trap stations with and without captures, under-story density was low overall, and there was a trend of higher seedling density at capture locations. Low understory density may allow squirrels more effective gliding movements between trees, which may enhance predator avoidance. Although the southern flying squirrel was reported from MDI numerous times during the 20th century, no voucher specimens exist, and species identification and localities have been poorly documented. Future surveys on MDI should consider collection of voucher specimens to validate subsequent survey efforts and effectively document changes in local biodiversity.

  11. Linking tree demography to climate change feedbacks: fire, larch forests, and carbon pools of the Siberian Arctic

    NASA Astrophysics Data System (ADS)

    Alexander, H. D.; Loranty, M. M.; Natali, S.; Pena, H., III; Ludwig, S.; Spektor, V.; Davydov, S. P.; Zimov, N.; Mack, M. C.

    2017-12-01

    Fire severity is increasing in larch forests of the Siberian Arctic as climate warms, and initial fire impacts on tree demographic processes could be an especially important determinant of long-term forest structure and carbon (C) dynamics. We hypothesized that (1) larch forest regrowth post-fire is largely determined by residual soil organic layer (SOL) depth because of the SOL's role as a seedbed and thermal regulator, and (2) changes in post-fire larch recruitment impact C accumulation through stand density impacts on understory microclimate and permafrost thaw. We tested these hypotheses by (1) experimentally creating a soil burn severity gradient in a Cajander larch (Larix cajanderi Mayr.) forest near Cherskiy, Russia and (2) quantifying C pools across a stand density gradient within a 75-year old fire scar. From 2012-2015, we added larch seeds to plots burned at different severities and monitored recruitment along with permafrost and active layer (i.e., subject to annual freeze-thaw) conditions (SOL depth, temperature, moisture, and thaw depth). Across the density gradient, we inventoried larch trees and harvested ground-layer vegetation to estimate aboveground contribution to C pools. We quantified woody debris C pools and sampled belowground C pools (soil, fine roots, and coarse roots) in the organic + upper (0-10 cm) mineral soil. Larch recruits were rare in unburned and low severity plots, but a total of 6 new germinants m-2 were tallied in moderate and high severity plots during the study. Seedling survival for > 1 year was only 40 and 25% on moderate and high severity treatments, respectively, but yielded net larch recruitment of 2 seedlings m-2, compared to 0.3 seedlings m-2 on low severity plots. Density of both total and established recruits increased with decreasing residual SOL depth, which correlated with increased soil temperature, moisture, and thaw depth. At 75-year post-fire, total C pools increased with increased larch density, largely due to increased tree aboveground C pools and decreased ground-layer vegetation C pools, which corresponded to higher canopy cover, cooler soils, and shallower active layer depths. Our findings highlight the potential for a climate-driven increase in fire severity to alter tree recruitment, successional dynamics, and C cycling in Siberian larch forests.

  12. Reproductive ecology and stand structure of Joshua tree forests across climate gradients of the Mojave Desert

    PubMed Central

    Hoines, Joshua

    2018-01-01

    Climate change is restructuring plant populations and can result in range shifts depending on responses at various life stages of plants. In 2013, a widespread and episodic flowering event provided an opportunity to characterize how Joshua tree’s reproductive success and population structure vary in response to the climate variability across its range. We examined the reproductive success and stand structure of 10 Joshua tree populations distributed across the Mojave Desert. Joshua tree density varied by more than an order of magnitude across sites. At 8 of the 10 sites, nearly 80% of the Joshua trees were in bloom, and at the other two 40% were in bloom. The range of seed production and fruit set across the study populations varied by more than an order of magnitude. Fruit production occurred at all of our study sites suggesting that yucca moth pollinators were present at our sites. Increasing temperature had strong positive correlations with the number of trees in bloom (R2 = 0.42), inflorescences per tree (R2 = 0.37), and fruit mass (R2 = 0.77) and seed size (R2 = 0.89. In contrast, temperature was negatively correlated with Joshua tree stand density (R2 = -0.80). Positive correlations between temperature and greater flower and seed production suggest that warming may positively affect Joshua Tree reproduction while negative relationships between temperature and stand density are suggestive of potential constraints of warmer temperatures on establishment success. PMID:29474414

  13. Reproductive ecology and stand structure of Joshua tree forests across climate gradients of the Mojave Desert.

    PubMed

    St Clair, Samuel B; Hoines, Joshua

    2018-01-01

    Climate change is restructuring plant populations and can result in range shifts depending on responses at various life stages of plants. In 2013, a widespread and episodic flowering event provided an opportunity to characterize how Joshua tree's reproductive success and population structure vary in response to the climate variability across its range. We examined the reproductive success and stand structure of 10 Joshua tree populations distributed across the Mojave Desert. Joshua tree density varied by more than an order of magnitude across sites. At 8 of the 10 sites, nearly 80% of the Joshua trees were in bloom, and at the other two 40% were in bloom. The range of seed production and fruit set across the study populations varied by more than an order of magnitude. Fruit production occurred at all of our study sites suggesting that yucca moth pollinators were present at our sites. Increasing temperature had strong positive correlations with the number of trees in bloom (R2 = 0.42), inflorescences per tree (R2 = 0.37), and fruit mass (R2 = 0.77) and seed size (R2 = 0.89. In contrast, temperature was negatively correlated with Joshua tree stand density (R2 = -0.80). Positive correlations between temperature and greater flower and seed production suggest that warming may positively affect Joshua Tree reproduction while negative relationships between temperature and stand density are suggestive of potential constraints of warmer temperatures on establishment success.

  14. Measuring Young’s modulus the easy way, and tracing the effects of measurement uncertainties

    NASA Astrophysics Data System (ADS)

    Nunn, John

    2015-09-01

    The speed of sound in a solid is determined by the density and elasticity of the material. Young’s modulus can therefore be calculated once the density and the speed of sound in the solid are measured. The density can be measured relatively easily, and the speed of sound through a rod can be measured very inexpensively by setting up a longitudinal standing wave and using a microphone to record its frequency. This is a simplified version of a technique called ‘impulse excitation’. It is a good educational technique for school pupils. This paper includes the description and the free provision of custom software to calculate the frequency spectrum of a recorded sound so that the resonant peaks can be readily identified. Discussion on the effect of measurement uncertainties is included to help the more thorough experimental student improve the accuracy of his method. The technique is sensitive enough to be able to detect changes in the elasticity modulus with a temperature change of just a few degrees.

  15. Multiscale habitat selection by Ruffed Grouse at low population densities

    USGS Publications Warehouse

    Zimmerman, G.S.; Gutierrez, R.J.; Thogmartin, W.E.; Banerjee, S.

    2009-01-01

    Theory suggests habitats should be chosen according to their relative evolutionary benefits and costs. It has been hypothesized that aspen (Populus spp.) forests provide optimal habitat for Ruffed Grouse (Bonasa umbellus). We used the low phase of a grouse population's cycle to assess the prediction that grouse should occupy aspen and avoid other forest types at low population density because of the presumptive fitness benefits of aspen. On the basis of our observations, we predict how the Ruffed Grouse population will increase in different forest types during the next cycle. In conifer (Pinus spp., Abies balsamea, Picea spp.)-dominated and mixed aspen-conifer landscapes, grouse densities were highest where forest types were evenly distributed. Within these landscapes, male Ruffed Grouse selected young aspen stands that were large and round or square. Although Ruffed Grouse selected young aspen stands strongly, contrary to prediction, they also used other forest types even when young aspen stands remained unoccupied. The relative densities of Ruffed Grouse in aspen and conifer forests indicated that the aspen forest's carrying capacities for grouse was higher than the conifer forest's at least during the low and declining phases of the grouse's cycle. On the basis of our observations, we predict that Ruffed Grouse populations in aspen-dominated landscapes will have higher population densities and fluctuate more than will populations in conifer-dominated landscapes. We suggest that studies of avian habitat selection would benefit from knowledge about the relative densities among habitats at differing population sizes because this information could provide insight into the role of habitat in regulating populations and clarify inferences from studies about habitat quality for birds. ?? 2009 by The Cooper Ornithological Society. All rights reserved.

  16. Variability in understory evapotranspiration with overstory density in Siberian larch forests

    NASA Astrophysics Data System (ADS)

    Tobio, A.; Loranty, M. M.; Kropp, H.; Pena, H., III; Alexander, H. D.; Natali, S.; Kholodov, A. L.

    2016-12-01

    Arctic ecosystems are changing rapidly in response to amplified rates of climate change. Increased vegetation productivity, altered ecosystem carbon and hydrologic cycling, and increased wildfire severity are among the key responses to changing permafrost and climate conditions. Boreal larch forests in northeastern Siberia are a critical but understudied ecosystem affected by these modifications. Understory vegetation in these ecosystems, which typically have low canopy cover, may account for half of all water fluxes. Despite the potential importance of the understory for ecosystem water exchange, there has been relatively little research examining variability in understory evapotranspiration in boreal larch forests. In particular, the water balance of understory shrubs and mosses is largely undefined and could provide insight on how understory vegetation and our changing climate interact. This is especially important because both observed increases in vegetation productivity and wildfire severity could lead to increases in forests density, altering the proportional contributions of over- and understory vegetation to whole ecosystem evapotranspiration. In order to better understand variability in understory evapotranspiration we measured in larch forests with differing overstory density and permafrost conditions that likely vary as a consequence of fire severity. We used the static chamber technique to measure fluxes across a range of understory vegetation types and environmental conditions. In general, we found that the understory vegetation in low density stands transpires more than that in high density stands. This tends to be correlated with a larger amount of aboveground biomass in the low density stands, and an increase in solar radiation, due to less shading by overstory trees. These results will help us to better understand water balances, evapotranspiration variability, and productivity changes associated with climate on understory vegetation. Additionally, our results will help understand how fire regime shifts may alter understory contributions to ecosystem evapotranspiration in Siberian larch forests.

  17. Elastic and fracture properties of free-standing amorphous ALD Al2O3 thin films measured with bulge test

    NASA Astrophysics Data System (ADS)

    Rontu, Ville; Nolvi, Anton; Hokkanen, Ari; Haeggström, Edward; Kassamakov, Ivan; Franssila, Sami

    2018-04-01

    We have investigated elastic and fracture properties of amorphous Al2O3 thin films deposited by atomic layer deposition (ALD) with bulge test technique using a free-standing thin film membrane and extended applicability of bulge test technique. Elastic modulus was determined to be 115 GPa for a 50 nm thick film and 170 GPa for a 15 nm thick film. Residual stress was 142 MPa in the 50 nm Al2O3 film while it was 116 MPa in the 15 nm Al2O3 film. Density was 3.11 g cm‑3 for the 50 nm film and 3.28 g cm‑3 for the 15 nm film. Fracture strength at 100 hPa s‑1 pressure ramp rate was 1.72 GPa for the 50 nm film while for the 15 nm film it was 4.21 GPa, almost 2.5-fold. Fracture strength was observed to be positively strain-rate dependent. Weibull moduli of these films were very high being around 50. The effective volume of a circular film in bulge test was determined from a FEM model enabling future comparison of fracture strength data between different techniques.

  18. Stand-structural effects on Heterobasidion abietinum-related mortality following drought events in Abies pinsapo.

    PubMed

    Linares, Juan Carlos; Camarero, Jesús Julio; Bowker, Matthew A; Ochoa, Victoria; Carreira, José Antonio

    2010-12-01

    Climate change may affect tree-pathogen interactions. This possibility has important implications for drought-prone forests, where stand dynamics and disease pathogenicity are especially sensitive to climatic stress. In addition, stand structural attributes including density-dependent tree-to-tree competition may modulate the stands' resistance to drought events and pathogen outbreaks. To assess the effects of stand structure on root-rot-related mortality after severe droughts, we focused on Heterobasidion abietinum mortality in relict Spanish stands of Abies pinsapo, a drought-sensitive fir. We compared stand attributes and tree spatial patterns in three plots with H. abietinum root-rot disease and three plots without root-rot. Point-pattern analyses were used to investigate the scale and extent of mortality patterns and to test hypotheses related to the spread of the disease. Dendrochronology was used to date the year of death and to assess the association between droughts and growth decline. We applied a structural equation modelling approach to test if tree mortality occurs more rapidly than predicted by a simple distance model when trees are subjected to high tree-to-tree competition and following drought events. Contrary to expectations of drought mortality, the effect of precipitation on the year of death was strong and negative, indicating that a period of high precipitation induced an earlier tree death. Competition intensity, related to the size and density of neighbour trees, also induced an earlier tree death. The effect of distance to the disease focus was negligible except in combination with intensive competition. Our results indicate that infected trees have decreased ability to withstand drought stress, and demonstrate that tree-to-tree competition and fungal infection act as predisposing factors of forest decline and mortality.

  19. Power spectral analysis of heart rate in hyperthyroidism.

    PubMed

    Cacciatori, V; Bellavere, F; Pezzarossa, A; Dellera, A; Gemma, M L; Thomaseth, K; Castello, R; Moghetti, P; Muggeo, M

    1996-08-01

    The aim of the present study was to evaluate the impact of hyperthyroidism on the cardiovascular system by separately analyzing the sympathetic and parasympathetic influences on heart rate. Heart rate variability was evaluated by autoregressive power spectral analysis. This method allows a reliable quantification of the low frequency (LF) and high frequency (HF) components of the heart rate power spectral density; these are considered to be under mainly sympathetic and pure parasympathetic control, respectively. In 10 newly diagnosed untreated hyperthyroid patients with Graves' disease, we analyzed power spectral density of heart rate cyclic variations at rest, while lying, and while standing. In addition, heart rate variations during deep breathing, lying and standing, and Valsalva's maneuver were analyzed. The results were compared to those obtained from 10 age-, sex-, and body mass index-matched control subjects. In 8 hyperthyroid patients, the same evaluation was repeated after the induction of stable euthyroidism by methimazole. Heart rate power spectral analysis showed a sharp reduction of HF components in hyperthyroid subjects compared to controls [lying, 13.3 +/- 4.1 vs. 32.0 +/- 5.6 normalized units (NU; P < 0.01); standing, 6.0 +/- 2.7 vs. 15.0 +/- 4.0 NU (P < 0.01); mean +/- SEM]. On the other hand components were comparable in the 2 groups (lying, 64.0 +/- 6.9 vs. 62.0 +/- 6.5 NU; standing, 77.0 +/- 6.5 vs. 78.0 +/- 5.4 NU). Hence, the LF/HF ratio, which is considered an index of sympathovagal balance, was increased in hyperthyroid subjects while both lying (11.3 +/- 4.5 vs. 3.5 +/- 1.1; P < 0.05) and standing (54.0 +/- 12.6 vs. 9.8 +/- 2.6; P < 0.02). This parameter was positively correlated with both T3 (r = 0.61; P < 0.05) and free T4 (r = 0.63; P < 0.05) serum levels. Among traditional cardiovascular autonomic tests, the reflex response of heart rate during lying to standing was significantly lower in hyperthyroid patients than in controls (1.12 +/- 0.03 vs. 1.31 +/- 0.04; P < 0.002). No statistically significant difference in reflex responses between the two groups was found in deep breathing or Valsalva's maneuver. In the 8 patients reexamined after methimazole treatment, we observed complete normalization of altered cardiovascular parameters, with slight predominance of the vagal component compared with controls. These results suggest that thyroid hormone excess may determine reduced parasympathetic activity and, thus, a relative hypersympathetic tone.

  20. Mapping of past stand-level forest disturbances and estimation of time since disturbance using simulated spaceborne LiDAR data

    NASA Astrophysics Data System (ADS)

    Sanchez Lopez, N.; Hudak, A. T.; Boschetti, L.

    2017-12-01

    Explicit information on the location, the size or the time since disturbance (TSD) at the forest stand level complements field inventories, improves the monitoring of forest attributes and the estimation of biomass and carbon stocks. Even-aged stands display homogenous structural parameters that have often been used as a proxy of stand age. Consequently, performing object-oriented analysis on Light Detection and Ranging (LiDAR) data has potential to detect historical stand-replacing disturbances. Recent research has shown good results in the delineation of forest stands as well as in the prediction of disturbance occurrence and TSD using airborne LiDAR data. Nevertheless, the use of airborne LiDAR for systematic monitoring of forest stands is limited by the sporadic availability of data and its high cost compared to satellite instruments. NASA's forthcoming Global Ecosystem Dynamics Investigations (GEDI) mission will provide systematically data on the vertical structure of the vegetation, but its use presents some challenges compared to the common discrete-return airborne LiDAR. GEDI will be a waveform instrument, hence the summary metrics will be different to those obtained with airborne LiDAR, and the sampling configuration could limit the utility of the data, especially on heterogeneous landscapes. The potential use of GEDI data for forest characterization at the stand level would therefore depend on the predictive power of the GEDI footprint metrics, and on the density of point samples relative to forest stand size (i.e. the number of observation/footprints per stand).In this study, we assess the performance of simulated GEDI-derived metrics for stand characterization and estimation of TSD, and the point density needed to adequately identify forest stands, which translates - due to the fixed sampling configuration - into the minimum temporal interval needed to collect a sufficient number of points. The study area was located in the Clear Creek, Selway River, and Elk Creek watersheds ( 54,000 ha) within the Nez Perce-Clearwater National Forest in Idaho, where airborne LiDAR and reference maps on TSD were available. Simulated GEDI footprints and waveforms were obtained from airborne LiDAR point clouds and the results were compared to similar analysis performed with airborne LiDAR.

  1. Biomass production and potential water stress increase with planting density in four highly productive clonal Eucalyptus genotypes

    Treesearch

    Rodrigo Hakamada; Robert M. Hubbard; Silvio Ferraz; Jose Luiz Stape; Cristiane Lemos

    2017-01-01

    The choice of planting density and tree genotype are basic decisions when establishing a forest stand. Understanding the interaction between planting density and genotype, and their relationship with biomass production and potential water stress, is crucial as forest managers are faced with a changing climate. However, few studies have investigated this relationship,...

  2. A new type of density-management diagram for slash pine plantations

    Treesearch

    Curtis L. VanderSchaaf

    2006-01-01

    Many Density-Management Diagrams (DMD) have been developed for conifer species throughout the world based on stand density index (SDI). The diagrams often plot the logarithm of average tree size (volume, weight, or quadratic mean diameter) over the logarithm of trees per unit area. A new type of DMD is presented for slash pine (Pinus elliottii var elliottii)...

  3. Influence of residual stand density on regeneration of northern hardwoods

    Treesearch

    W. B. Leak; Dale S. Solomon

    1975-01-01

    In a study of hardwood regeneration, experimental plots were treated to produce residual densities of 40, 60, 80, and 100 square feet of basal area per acre with 30, 45, and 60 percent sawtimber. After 9 years, numbers of stems and percent stocking were estimated for each species and they were rated for reproductive potential under different residual densities. Data...

  4. Effects of fish density and river fertilization on algal standing stocks, invertebrates communities, and fish production in an Arctic River

    USGS Publications Warehouse

    Deegan, Linda A.; Peterson, B.J.; Golden, H.; McIvor, C.C.; Miller, M.C.

    1997-01-01

    This study examined the relative importance of bottom-up and top-down controls of an arctic stream food web by simultaneous manipulation of the top predator and nutrient availability. We created a two-step trophic system (algae to insects) by removal of the top predator (Arctic grayling, Thymallus arcticus) in fertilized and control stream reaches. Fish abundance was also increased 10 times to examine the effect of high fish density on stream ecosystem dynamics and fish. We measured the response of epilithic algae, benthic and drifting insects, and fish to nutrient enrichment and to changes in fish density. Insect grazers had little effect on algae and fish had little effect on insects. In both the control and fertilized reaches, fish growth, energy storage, and reproductive response of females declined with increased fish density. Fish growth and energy storage were more closely correlated with per capita insect availability than with per capita algal standing stock

  5. Ecosystem carbon density and allocation across a chronosequence of longleaf pine forests.

    PubMed

    Samuelson, Lisa J; Stokes, Thomas A; Butnor, John R; Johnsen, Kurt H; Gonzalez-Benecke, Carlos A; Martin, Timothy A; Cropper, Wendell P; Anderson, Pete H; Ramirez, Michael R; Lewis, John C

    2017-01-01

    Forests can partially offset greenhouse gas emissions and contribute to climate change mitigation, mainly through increases in live biomass. We quantified carbon (C) density in 20 managed longleaf pine (Pinus palustris Mill.) forests ranging in age from 5 to 118 years located across the southeastern United States and estimated above- and belowground C trajectories. Ecosystem C stock (all pools including soil C) and aboveground live tree C increased nonlinearly with stand age and the modeled asymptotic maxima were 168 Mg C/ha and 80 Mg C/ha, respectively. Accumulation of ecosystem C with stand age was driven mainly by increases in aboveground live tree C, which ranged from <1 Mg C/ha to 74 Mg C/ha and comprised <1% to 39% of ecosystem C. Live root C (sum of below-stump C, ground penetrating radar measurement of lateral root C, and live fine root C) increased with stand age and represented 4-22% of ecosystem C. Soil C was related to site index, but not to stand age, and made up 39-92% of ecosystem C. Live understory C, forest floor C, downed dead wood C, and standing dead wood C were small fractions of ecosystem C in these frequently burned stands. Stand age and site index accounted for 76% of the variation in ecosystem C among stands. The mean root-to-shoot ratio calculated as the average across all stands (excluding the grass-stage stand) was 0.54 (standard deviation of 0.19) and higher than reports for other conifers. Long-term accumulation of live tree C, combined with the larger role of belowground accumulation of lateral root C than in other forest types, indicates a role of longleaf pine forests in providing disturbance-resistant C storage that can balance the more rapid C accumulation and C removal associated with more intensively managed forests. Although other managed southern pine systems sequester more C over the short-term, we suggest that longleaf pine forests can play a meaningful role in regional forest C management. © 2016 by the Ecological Society of America.

  6. Biogeographical patterns of biomass allocation in leaves, stems, and roots in China's forests.

    PubMed

    Zhang, Hao; Wang, Kelin; Xu, Xianli; Song, Tongqing; Xu, Yanfang; Zeng, Fuping

    2015-11-03

    To test whether there are general patterns in biomass partitioning in relation to environmental variation when stand biomass is considered, we investigated biomass allocation in leaves, stems, and roots in China's forests using both the national forest inventory data (2004-2008) and our field measurements (2011-2012). Distribution patterns of leaf, stem, and root biomass showed significantly different trends according to latitude, longitude, and altitude, and were positively and significantly correlated with stand age and mean annual precipitation. Trade-offs among leaves, stems, and roots varied with forest type and origin and were mainly explained by stand biomass. Based on the constraints of stand biomass, biomass allocation was also influenced by forest type, origin, stand age, stand density, mean annual temperature, precipitation, and maximum temperature in the growing season. Therefore, after stand biomass was accounted for, the residual variation in biomass allocation could be partially explained by stand characteristics and environmental factors, which may aid in quantifying carbon cycling in forest ecosystems and assessing the impacts of climate change on forest carbon dynamics in China.

  7. Biogeographical patterns of biomass allocation in leaves, stems, and roots in China’s forests

    PubMed Central

    Zhang, Hao; Wang, Kelin; Xu, Xianli; Song, Tongqing; Xu, Yanfang; Zeng, Fuping

    2015-01-01

    To test whether there are general patterns in biomass partitioning in relation to environmental variation when stand biomass is considered, we investigated biomass allocation in leaves, stems, and roots in China’s forests using both the national forest inventory data (2004–2008) and our field measurements (2011–2012). Distribution patterns of leaf, stem, and root biomass showed significantly different trends according to latitude, longitude, and altitude, and were positively and significantly correlated with stand age and mean annual precipitation. Trade-offs among leaves, stems, and roots varied with forest type and origin and were mainly explained by stand biomass. Based on the constraints of stand biomass, biomass allocation was also influenced by forest type, origin, stand age, stand density, mean annual temperature, precipitation, and maximum temperature in the growing season. Therefore, after stand biomass was accounted for, the residual variation in biomass allocation could be partially explained by stand characteristics and environmental factors, which may aid in quantifying carbon cycling in forest ecosystems and assessing the impacts of climate change on forest carbon dynamics in China. PMID:26525117

  8. Modeling biophysical properties of broad-leaved stands in the hyrcanian forests of Iran using fused airborne laser scanner data and ultraCam-D images

    NASA Astrophysics Data System (ADS)

    Mohammadi, Jahangir; Shataee, Shaban; Namiranian, Manochehr; Næsset, Erik

    2017-09-01

    Inventories of mixed broad-leaved forests of Iran mainly rely on terrestrial measurements. Due to rapid changes and disturbances and great complexity of the silvicultural systems of these multilayer forests, frequent repetition of conventional ground-based plot surveys is often cost prohibitive. Airborne laser scanning (ALS) and multispectral data offer an alternative or supplement to conventional inventories in the Hyrcanian forests of Iran. In this study, the capability of a combination of ALS and UltraCam-D data to model stand volume, tree density, and basal area using random forest (RF) algorithm was evaluated. Systematic sampling was applied to collect field plot data on a 150 m × 200 m sampling grid within a 1100 ha study area located at 36°38‧- 36°42‧N and 54°24‧-54°25‧E. A total of 308 circular plots (0.1 ha) were measured for calculation of stand volume, tree density, and basal area per hectare. For each plot, a set of variables was extracted from both ALS and multispectral data. The RF algorithm was used for modeling of the biophysical properties using ALS and UltraCam-D data separately and combined. The results showed that combining the ALS data and UltraCam-D images provided a slight increase in prediction accuracy compared to separate modeling. The RMSE as percentage of the mean, the mean difference between observed and predicted values, and standard deviation of the differences using a combination of ALS data and UltraCam-D images in an independent validation at 0.1-ha plot level were 31.7%, 1.1%, and 84 m3 ha-1 for stand volume; 27.2%, 0.86%, and 6.5 m2 ha-1 for basal area, and 35.8%, -4.6%, and 77.9 n ha-1 for tree density, respectively. Based on the results, we conclude that fusion of ALS and UltraCam-D data may be useful for modeling of stand volume, basal area, and tree density and thus gain insights into structural characteristics in the complex Hyrcanian forests.

  9. Analysis and comparison of nonlinear tree height prediction strategies for Douglas-fir forest.

    Treesearch

    H. Temesgen; V.J. Monleon; D.W. Hann

    2008-01-01

    Using an extensive Douglas-fir data set from southwest Oregon, we examined the (I) performance and suitability of selected prediction strategies, (2) contribution of relative position and stand-density measures in improving tree height (h) prediction values, and (3) effect of different subsampling designs to fill in missing h values in a new stand using a regional...

  10. The population and seasonal dynamics of weevils developing in the soil of birch stands

    Treesearch

    E. Kula

    2003-01-01

    Curculionidae developing in the soil of birch stands in an air-polluted region were classified using the method of soil photoeclectors on the basis of their population dynamics (1986-2000) and phenology of their emergence from where they developed. In the course of 15 years we saw two evident culminations in the population density of Polydrusus undatus...

  11. Using the Forest Vegetation Simulator to reconstruct historical stand conditions in the Colorado Front Range

    Treesearch

    Paula J. Fornwalt; Merrill R. Kaufmann; Laurie S. Huckaby; Jason M. Stoker

    2002-01-01

    Presettlement ponderosa pine/Douglas-fir forests of the Colorado Front Range were open and heterogeneous. Logging, grazing, and fire suppression over past 100 to150 years have altered stand structure by changing diameter distributions and increasing overstory density. In an effort to guide forest restoration toward presettlement conditions, we are currently using the...

  12. Potential fire behavior is reduced following forest restoration treatments

    Treesearch

    Peter Z. Fule; Charles McHugh; Thomas A. Heinlein; W. Wallace Covington

    2001-01-01

    Potential fire behavior was compared under dry, windy weather conditions in 12 ponderosa pine stands treated with alternative thinning prescriptions in the wildland/urban interface of Flagstaff, Arizona. Prior to thinning, stands averaged 474 trees/ acre, 158 ft2/acre basal area, crown bulk density 0.0045 lb/ft3, and crown base height 19.2 ft. Three thinning treatments...

  13. Stand-yield prediction for managed Ocala sand pine

    Treesearch

    D.L. Rockwood; B. Yang; K.W. Outcalt

    1997-01-01

    Sand pine is a very important species in Florida, producing significant quantities of fiber. The purpose of this study was to develop the site index and stand-level growth and yield equations managers need to make informed decisions. Data were collected from 35 seeded plots of Ocala sand pine covering a range of site indexes, ages, and densities in 1982-83. These plots...

  14. Stand density index as a tool to assess the maximization of forest carbon and biomass

    Treesearch

    Christopher W. Woodall; Anthony W. D’Amato; John B. Bradford; Andrew O. Finley

    2012-01-01

    Given the ability of forests to mitigate greenhouse gas emissions and provide feedstocks to energy utilities, there is an emerging need to assess forest biomass/carbon accretion opportunities over large areas. Techniques for objectively quantifying stand stocking of biomass/carbon are lacking for large areas given the complexity of tree species composition in the U.S....

  15. Regeneration Development Across a Range of Reproduction Cutting Methods in Shortleaf Pine and Pine-Hardwood Stands in the Interior Highlands

    Treesearch

    James M. Guldin; James B. Baker; Michael G. Shelton

    2004-01-01

    Abstract - Density, milacre stocking, and height of shortleaf pine ( Pinus echinata Mill.) regeneration under 13 reproduction cutting methods were measured after 5 growing seasons across a range of reproduction cutting treatments in shortleaf pine and pine-hardwood stands in the Interior Highlands of Arkansas and Oklahoma. A subset...

  16. Partial Cutting and Establishment of Artificial Nuttall Oak Regeneration in the Mississippi Alluvial Plain

    Treesearch

    Benjamin P. Ware; Emile S. Gardiner

    2004-01-01

    In the fall of 1998, we harvested a mixed bottomland hardwood stand in Sharkey County, MS, to establish two replicates of three residual stocking levels (0, 25, and 50 percent). We compared the various stand densities for suitability in establishment and growth of artificial oak regeneration. Following the overstory harvest, treatment plots received chain saw site...

  17. Changes in stand structure and composition after restoration treatments in low elevation dry forests of northeastern Oregon.

    Treesearch

    Andrew Youngblood; Kerry L. Metlen; Kent Coe

    2006-01-01

    In many fire-dependent forests in the United States, changes occurring in the last century have resulted in overstory structures, conifer densities, down woody structure and understory plant communities that deviate from those described historically. With these changes, many forests are presumed to be unsustainable. Broad-scale treatments are proposed to promote stand...

  18. Mortality among Seed Trees in Longleaf Pine Shelterwood Stands

    Treesearch

    William D. Boyer

    1970-01-01

    Mortality of longieaf pine (Pinus palustris Mill.) seed trees was recorded in 27 regeneration areas ranging from North Carolina to Louisiana. Annual mortality averaged 0.7 percent before, and 1.9 percent after a seed cut reduced stand density to about 30 square feet of basal area per acre. On a per-acre basis, however, annual losses averaged 0....

  19. Evaluation of sapling height and density after clearcutting and group selection in the Missouri Ozarks

    Treesearch

    Guerric T. Good; Benjamin O. Knapp; Lance A. Vickers; David R. Larsen; John M. Kabrick

    2017-01-01

    Silvicultural decisions often affect the development and characteristics of a stand. Silvicultural regeneration events can have immediate and gradual impacts on stand development. The objective of this study was to evaluate the effects of two silvicultural regeneration methods, clearcutting and group selection, on the composition of trees that are likely to recruit to...

  20. Forage production after hardwood control in a southern pine-hardwood stand

    Treesearch

    Robert M. Blair

    1971-01-01

    After hardwoods were removed from an all-aged pine-hardwood stand in central Louisiana, herbage available in late summer increased for 2 to 4 years, then declined rapidly as the density and growth of pine seedlings increased. Browse production increased for 6 to 8 years, but together with herbage produciton returned to pretreatment levels within 10 years. Competition...

  1. Stand-level bird response to experimental forest management in the Missouri Ozarks

    Treesearch

    Sarah W. Kendrick; Paul A. Porneluzi; Frank R. Thompson; Dana L. Morris; Janet M. Haslerig; John Faaborg

    2015-01-01

    Long-term landscape-scale experiments allow for the detection of effects of silviculture on bird abundance. Manipulative studies allow for strong inference on effects and confirmation of patterns from observational studies.We estimated bird-territory density within forest stands (2.89-62 ha) for 19 years of the Missouri Ozark Forest Ecosystem Project (MOFEP), a 100-...

  2. Forest stand structure and pattern of old-growth western hemlock/Douglas-fir and mixed-conifer forests

    Treesearch

    Malcolm North; Jiquan Chen; Brian Oakley; Bo Song; Mark Rudnicki; Andrew Gray; Jim Innes

    2004-01-01

    With fire suppression, many western forests are expected to have fewer gaps and higher stem density of shade-tolerant species as light competition becomes a more significant influence on stand pattern and composition. We compared species composition, structure, spatial pattern, and environmental factors such as light and soil moisture between two old-growth forests:...

  3. Clonal expansion and seedling recruitment of Oregon grape (Berberis nervosa) in Douglas-fir (Pseudotsuga menziesii) forests: Comparisons with salal (Gaultheria shallon)

    USGS Publications Warehouse

    Huffman, David W.; Tappeiner, John C.

    1997-01-01

    Seedling regeneration and morphology of Oregon grape (Berberis nervosa Pursh) and salal (Gaultheria shallon Pursh) were studied in thinned and unthinned Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) stands in the central Coast Range, Oregon. Above- and below-ground growth of both species were significantly and negatively correlated with stand density. Oregon grape appears to have less potential for vegetative spread than does salal. It produced two to three times fewer rhizome extensions, and rhizome extensions were only half as long as those of salal. Oregon grape seedlings were common in areas of moss ground cover among patches of the two species. Salal seedlings were restricted to decaying logs. Seedling densities of Oregon grape in thinned stands were more than six times those in unthinned stands. For Oregon grape, understory establishment is accomplished by seedling establishment and recruitment of new genets. In contrast, salal maintains itself in forest understories primarily through vegetative growth, since its seedling establishment is restricted mainly to decayed wood. Continual recruitment of new aerial stems or ramets enables Oregon grape to maintain a dense cover once it is established in the understory.

  4. Soil seed bank in different habitats of the Eastern Desert of Egypt.

    PubMed

    Gomaa, Nasr H

    2012-04-01

    The floristic composition and species diversity of the germinable soil seed bank were studied in three different habitats (desert salinized land, desert wadi, and reclaimed land) in the Eastern Desert of Egypt. Moreover, the degree of similarity between the seed bank and the above-ground vegetation was determined. The seed bank was studied in 40 stands representing the three habitats. Ten soil samples (each 25 × 20 cm and 5 cm depth) were randomly taken per stand. The seed bank was investigated by the seedling emergence method. Some 61 species belonging to 21 families and 54 genera were identified in the germinable seed bank. The recorded species include 43 annuals and 18 perennials. Ordination of stands by Detrended Correspondence Analysis (DCA) indicates that the stands of the three habitats are markedly distinguishable and show a clear pattern of segregation on the ordination planes. This indicates variations in the species composition among habitats. The results also demonstrate significant associations between the floristic composition of the seed bank and edaphic factors such as CaCO3, electrical conductivity, organic carbon and soil texture. The reclaimed land has the highest values of species richness, Shannon-index of diversity and the density of the germinable seed bank followed by the habitats of desert wadi and desert salinized land. Motyka's similarity index between the seed bank and the above-ground vegetation is significantly higher in reclaimed land (75.1%) compared to desert wadi (38.4%) and desert salinized land (36.5%).

  5. Take me to your leader: does early successional nonhost vegetation spatially inhibit Pissodes strobi (Coleoptera: Curculionidae)?

    PubMed

    Koopmans, Jordan M; De La Giroday, Honey-Marie C; Lindgren, B Staffan; Aukema, Brian H

    2009-08-01

    The spatial influences of host and nonhost trees and shrubs on the colonization patterns of white pine weevil Pissodes strobi (Peck) were studied within a stand of planted interior hybrid spruce [Picea glauca (Moench) Voss x Picea engelmannii (Parry) ex Engelm.]. Planted spruce accounted for one third of all trees within the stand, whereas the remaining two thirds were comprised of early-successional nonhost vegetation, such as alder (Alnus spp.), paper birch (Betula papyrifera Marsh.), black cottonwood [Populus balsamifera ssp. trichocarpa (T. Ng.) Brayshaw], lodgepole pine [Pinus contorta (Dougl.) ex Loud.], trembling aspen (Populus tremuloides Michx), willow (Salix spp.), and Canadian buffaloberry [Shepherdia canadensis (L.) Nutt.]. Unlike the spruce trees, nonhost vegetation in the stand was not uniformly distributed. Spatial point process models showed that Canadian buffaloberry, paper birch, black cottonwoood, and trembling aspen had negative associations with damage caused by the weevil, even though the density of the insects' hosts in these areas did not change. Moreover, knowing the locations of these nonhost trees provided as much, or more, inference about the locations of weevil-attacked trees as knowing the locations of suitable or preferred host trees (i.e., those larger in size). Nonhost volatiles, the alteration of soil composition, and overstory shade are discussed as potential explanatory factors for the patterns observed. New research avenues are suggested to determine whether nonhost vegetation in early successional stands might be an additional tool in the management of these insects in commercially important forests.

  6. Longer-term effects of selective thinning on microarthropod communities in a late-successional coniferous forest

    USGS Publications Warehouse

    Peck, R.W.; Niwa, C.G.

    2005-01-01

    Microarthropod densities within late-successional coniferous forests thinned 16-41 yr before sampling were compared with adjacent unthinned stands to identify longer term effects of thinning on this community. Soil and forest floor layers were sampled separately on eight paired sites. Within the forest floor oribatid, mesostigmatid, and to a marginal extent, prostigmatid mites, were reduced in thinned stands compared with unthinned stands. No differences were found for Collembola in the forest floor or for any mite suborder within the soil. Family level examination of mesostigmatid and prostigmatid mites revealed significant differences between stand types for both horizons. At the species level, thinning influenced numerous oribatid mites and Collembola. For oribatid mites, significant or marginally significant differences were found for seven of 15 common species in the forest floor and five of 16 common species in soil. Collembola were affected less, with differences found for one of 11 common species in the forest floor and three of 13 common species in soil. Multivariate analysis of variance and ordination indicated that forest thinning had little influence on the composition of oribatid mite and collembolan communities within either the forest floor or soil. Differences in microclimate or in the accumulation of organic matter on the forest floor were likely most responsible for the observed patterns of abundance. Considering the role that microarthropods play in nutrient cycling, determining the functional response of a wide range of taxa to thinning may be important to effective ecosystem management.

  7. Determining stocking, forest type and stand-size class from forest inventory data

    Treesearch

    Mark H. Hansen; Jerold T. Hahn

    1992-01-01

    This paper describes the procedures used by North Central Forest Experiment Station's Forest Inventory and Analysis Work Unit (NCFIA) in determining stocking, forest type, and stand-size class. The stocking procedure assigns a portion of the stocking to individual trees measured on NCFIA 10-point field plots. Stand size and forest type are determined as functions...

  8. Acoustic radiation force on a rigid elliptical cylinder in plane (quasi)standing waves

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Mitri, F. G., E-mail: F.G.Mitri@ieee.org

    2015-12-07

    The acoustic radiation force on a 2D elliptical (non-circular) cylinder centered on the axis of wave propagation of plane quasi-standing and standing waves is derived, based on the partial-wave series expansion (PWSE) method in cylindrical coordinates. A non-dimensional acoustic radiation force function, which is the radiation force per unit length, per characteristic energy density and per unit cross-sectional surface of the ellipse, is defined in terms of the scattering coefficients that are determined by applying the Neumann boundary condition for an immovable surface. A system of linear equations involving a single numerical integration procedure is solved by matrix inversion. Numericalmore » simulations showing the transition from the quasi-standing to the (equi-amplitude) standing wave behaviour are performed with particular emphasis on the aspect ratio a/b, where a and b are the ellipse semi-axes, as well as the dimensionless size parameter kb (where k is the wavenumber), without the restriction to a particular range of frequencies. It is found that at high kb values > 1, the radiation force per length with broadside incidence is larger, whereas the opposite situation occurs in the long-wavelength limit (i.e., kb < 1). The results are particularly relevant in acoustic levitation of elliptical cylinders, the acoustic stabilization of liquid columns in a host medium, acousto-fluidics devices, and other particle dynamics applications to name a few. Moreover, the formalism presented here may be effectively applied to compute the acoustic radiation force on other 2D surfaces of arbitrary shape such as super-ellipses, Chebyshev cylindrical particles, or other non-circular geometries.« less

  9. Solar cycle variation of geosynchronous plasma mass density derived from the frequency of standing Alfvén waves

    NASA Astrophysics Data System (ADS)

    Takahashi, Kazue; Denton, Richard E.; Singer, Howard J.

    2010-07-01

    We have studied the solar cycle variation of equatorial plasma mass density ρeq in the plasma trough at geosynchronous altitude. The density was indirectly determined from the frequency, fT3, of the third harmonic of toroidal standing Alfvén waves detected over a 12 year period from 1980 to 1991 with magnetometers on five Geostationary Operational Environmental Satellites (GOES). Realistic models of the ambient magnetic field and field line mass distribution were used in numerically solving the wave equation to relate fT3 to ρeq. Scanning the magnetometer data in a 30 min time window that moved forward in 10 min steps, we obtained 228,382 fT3 samples equivalent to 1586 days of data. The detection rate of fT3 is highest (˜50%) in the prenoon sector, and fT3 and ρeq samples from this sector were used to examine their dependence on F10.7, Kp, and Dst. Overall, F10.7 exhibits the highest correlation with fT3 and ρeq, implying that the solar UV/EUV control of ion production at the ionospheric height is strongly reflected in mass density variations at geosynchronous orbit. Using 27 day medians computed excluding periods of plasmasphere expansion to geosynchronous orbit and geomagnetic storm, we obtained the empirical formula fT3 (mHz) = 38 - 0.097F10.7 and logρeq (amu cm-3) = 0.42 + 0.0039F10.7, where F10.7 is given in the solar flux units 10-22 W · m-2 · Hz-1. This last formula means that with the 27 day F10.7 in the range of 68-255 in the selected solar cycle, the mass density varied by a factor of ˜5 from ˜5 to ˜26 amu cm-3. During extremely quiet times (Kp averaged using a 3 day time scale <1), for which the plasmasphere may extend out to geosynchronous orbit, and during storm periods (Dst < -50 nT), the mass density may be enhanced beyond these values.

  10. Interference and economic threshold level of little seed canary grass in wheat under different sowing times.

    PubMed

    Hussain, Saddam; Khaliq, Abdul; Matloob, Amar; Fahad, Shah; Tanveer, Asif

    2015-01-01

    Little seed canary grass (LCG) is a pernicious weed of wheat crop causing enormous yield losses. Information on the interference and economic threshold (ET) level of LCG is of prime significance to rationalize the use of herbicide for its effective management in wheat fields. The present study was conducted to quantify interference and ET density of LCG in mid-sown (20 November) and late-sown (10 December) wheat. Experiment was triplicated in randomized split-plot design with sowing dates as the main plots and LCG densities (10, 20, 30, and 40 plants m(-2)) as the subplots. Plots with two natural infestations of weeds including and excluding LCG were maintained for comparing its interference in pure stands with designated densities. A season-long weed-free treatment was also run. Results indicated that composite stand of weeds, including LCG, and density of 40 LCG plants m(-2) were more competitive with wheat, especially when crop was sown late in season. Maximum weed dry biomass was attained by composite stand of weeds including LCG followed by 40 LCG plants m(-2) under both sowing dates. Significant variations in wheat growth and yield were observed under the influence of different LCG densities as well as sowing dates. Presence of 40 LCG plants m(-2) reduced wheat yield by 28 and 34% in mid- and late-sown wheat crop, respectively. These losses were much greater than those for infestation of all weeds, excluding LCG. Linear regression model was effective in simulating wheat yield losses over a wide range of LCG densities, and the regression equations showed good fit to observed data. The ET levels of LCG were 6-7 and 2.2-3.3 plants m(-2) in mid- and late-sown wheat crop, respectively. Herbicide should be applied in cases when LCG density exceeds these levels under respective sowing dates.

  11. The effects of planting density and cultural intensity on loblolly pine crown characteristics at age twelve

    Treesearch

    Madison Akers; Michael Kane; Robert Teskey; Richard Daniels; Dehai Zhao; Santosh Subedi

    2012-01-01

    Twelve-year old loblolly pine (Pinus taeda L.) stands were analyzed for the effects of planting density and cultural intensity on tree and crown attributes. Four study installations were located in the Piedmont and Upper Coastal Plain regions of the U.S. South. The treatments included six planting densities (740, 1480, 2220, 2960, 3700, 4440 trees...

  12. Regeneration Patterns in Canopy Gaps of Mixed-oak Forests of the Southern Appalachians: Influences of Topographic Position and Evergreen Understory

    Treesearch

    Barton D. Clinton; Lindsay R. Boring; Wayne T. Swank

    1994-01-01

    Canopy gaps in southern Appalachian mixed-oak forests were assessed for the effects of topographic, gap and stand variables on density of wood seedlings. Seedling density was significantly correlated with percent slope and positively with gap age (l-5 yr). Density varied substantially among topographic positions and increased with gap size. Species richness...

  13. Age-related effects on leaf area/sapwood area relationships, canopy transpiration and carbon gain of Norway spruce stands (Picea abies) in the Fichtelgebirge, Germany.

    PubMed

    Köstner, B; Falge, E; Tenhunen, J D

    2002-06-01

    Stand age is an important structural determinant of canopy transpiration (E(c)) and carbon gain. Another more functional parameter of forest structure is the leaf area/sapwood area relationship, A(L)/A(S), which changes with site conditions and has been used to estimate leaf area index of forest canopies. The interpretation of age-related changes in A(L)/A(S) and the question of how A(L)/A(S) is related to forest functions are of current interest because they may help to explain forest canopy fluxes and growth. We conducted studies in mature stands of Picea abies (L.) Karst. varying in age from 40 to 140 years, in tree density from 1680 to 320 trees ha(-1), and in tree height from 15 to 30 m. Structural parameters were measured by biomass harvests of individual trees and stand biometry. We estimated E(c) from scaled-up xylem sap flux of trees, and canopy-level fluxes were predicted by a three-dimensional microclimate and gas exchange model (STANDFLUX). In contrast to pine species, A(L)/A(S) of P. abies increased with stand age from 0.26 to 0.48 m(2) cm(-2). Agreement between E(c) derived from scaled-up sap flux and modeled canopy transpiration was obtained with the same parameterization of needle physiology independent of stand age. Reduced light interception per leaf area and, as a consequence, reductions in net canopy photosynthesis (A(c)), canopy conductance (g(c)) and E(c) were predicted by the model in the older stands. Seasonal water-use efficiency (WUE = A(c)/E(c)), derived from scaled-up sap flux and stem growth as well as from model simulation, declined with increasing A(L)/A(S) and stand age. Based on the different behavior of age-related A(L)/A(S) in Norway spruce stands compared with other tree species, we conclude that WUE rather than A(L)/A(S) could represent a common age-related property of all species. We also conclude that, in addition to hydraulic limitations reducing carbon gain in old stands, a functional change in A(L)/A(S) that is related to reduced light interception per leaf area provides another potential explanation for reduced carbon gain in old stands of P. abies, even when hydraulic constraints increase in response to changes in canopy architecture and aging.

  14. Free standing Cu2Te, new anode material for sodium-ion battery

    NASA Astrophysics Data System (ADS)

    Sarkar, Ananta; Mallick, Md. Mofasser; Panda, Manas Ranjan; Vitta, Satish; Mitra, Sagar

    2018-05-01

    Sodium-ion battery is the most popular alternative to lithium-ion energy storage system due to its low cost and huge abundant resources throughout the world. Although recent literature showed cathode materials for sodium ion battery performs almost equivalent to lithium-ion counterpart but the anode of this sodium-ion battery is in premature state. Here, we introduced free-standing copper telluride (Cu2Te), a new anode materials for sodium-ion battery. For making the electrode we did not use any conductive carbon or current collector which increase the volumetric density as well as reduce the cost of the cell. This metallic Cu2Te alloy exhibited a high reversible capacity of ˜275 mAh g-1 at 50 mA g-1 current density and ˜200 mAh g-1 at higher current density of 100 mA g-1, operating between 0.1 to 2.0 V.

  15. Windows of opportunity: white-tailed deer and the dynamics of northern hardwood forests of the northeastern US

    USGS Publications Warehouse

    Sage, R.W.; Porter, W.F.; Underwood, H.B.

    2003-01-01

    Herbivory, lighting regimes, and site conditions are among the most important determinants of forest regeneration success, but these are affected by a host of other factors such as weather, predation, human exploitation, pathogens, wind and fire. We draw together > 50 years of research on the Huntington Wildlife Forest in the central Adirondack Mountains of New York to explore regeneration of northern hardwoods. A series of studies each of which focused on a single factor failed to identify the cause of regeneration failure. However, integration of these studies led to broader understanding of the process of forest stand development and identified at least three interacting factors: lighting regime, competing vegetation and selective browsing by white-tailed deer (Odocoileus virginianus). The diverse 100-200 year-old hardwood stands present today probably reflect regeneration during periods of low deer density (< 2.0 deer/km super(2)) and significant forest disturbance. If this hypothesis is correct, forest managers can mimic these 'natural windows of opportunity' through manipulation of a few sensitive variables in the system. Further, these manipulations can be conducted on a relatively small geographic scale. Control of deer densities on a scale of 500 ha and understory American beech (Fagus grandifolia) on a scale of < 100 ha in conjunction with an even-aged regeneration system consistently resulted in successful establishment of desirable hardwood regeneration.

  16. Mammographic breast density in recent and longer-standing ethiopian immigrants to israel.

    PubMed

    Sklair-Levy, Miri; Segev, Anat; Sella, Tamar; Calderon-Margalit, Ronit; Zippel, Douglas

    2018-04-23

    High breast density is associated with an increased risk of breast cancer development. Little is known concerning ethnic variations in breast density and its relevant contributing factors. We aimed to study breast density among Ethiopian immigrants to Israel in comparison with Israeli-born women and to determine any effect on breast density of the length of residency in the immigrant population. Mammographic breast density using the BI-RADS system was estimated and compared between 77 women of Ethiopian origin who live in Israel and 177 Israeli-born controls. Logistic regression analysis was performed to estimate the odds ratios (OR) for high density (BI-RADS score ≥ 3) vs low density (BI-RADS score < 3) cases, comparing the 2 origin groups. Ethiopian-born women had a crude OR of 0.15 (95% CI: 0.08-0.26) for high breast density compared with Israeli-born women. Adjustments for various cofounders did not affect the results. Time since immigration to Israel seemed to modify the relationship, with a stronger association for women who immigrated within 2 years prior to mammography (OR:0.07, 95% CI: 0.03-0.17) as opposed to women with a longer residency stay in Israel (OR:0.23, 95% CI:0.10-0.50). Adjustments of various confounders did not alter these findings. Breast density in Ethiopian immigrants to Israel is significantly lower than that of Israeli-born controls. Our study suggests a positive association between time since immigration and breast density. Future studies are required to define the possible effects of dietary change on mammographic density following immigration. © 2018 Wiley Periodicals, Inc.

  17. Forest production dynamics along a wood density spectrum in eastern US forests

    Treesearch

    C.W. Woodall; M.B. Russell; B.F. Walters; A.W. D' Amato; K. Zhu; S.S. Saatchi

    2015-01-01

    Emerging plant economics spectrum theories were confirmed across temperate forest systems of the eastern US where the use of a forest stand's mean wood density elucidated forest volume and biomass production dynamics integrating aspects of climate, tree mortality/growth, and rates of site occupancy.

  18. Within and Between Canopy Variabilit of Foliar Nitrogen Concentration for Loblolly and Slash Pine Stands Planted at Different Densities

    Treesearch

    Rodney E. Will; Greg Barron-Gafford; Robert O. Teskey; Barry D. Shiver

    2004-01-01

    Mid-summer foliar nitrogen concentrations (N) were measured at three canopy positions (upper, middle, lower), two foliage ages per canopy position (current-year and 1-year-old), and two flushes per age class (first flush and second flush) in 4-year-old loblolly (Pinus taeda L.) and slash pine (Pinus elliottii Engelm.) stands...

  19. Growth model for uneven-aged loblolly pine stands : simulations and management implications

    Treesearch

    C.-R. Lin; J. Buongiorno; Jeffrey P. Prestemon; K. E. Skog

    1998-01-01

    A density-dependent matrix growth model of uneven-aged loblolly pine stands was developed with data from 991 permanent plots in the southern United States. The model predicts the number of pine, soft hardwood, and hard hardwood trees in 13 diameter classes, based on equations for ingrowth, upgrowth, and mortality. Projections of 6 to 10 years agreed with the growth...

  20. Shortleaf Pine ecosystem restoration: impacts on soils and woody debris in the Ouachita mountains of the southern United States

    Treesearch

    Hal O. Liechty; Kenneth R. Luckow; Jessica Seifert Daniel; Daniel A. Marion; Martin Spetich; James M. Guldin

    2004-01-01

    A number of organizations and government agencies have been involved with restoration of overstocked shortleaf pine-hardwood stands to shortleaf pine-bluestem ecosystems in the Ouachita Mountains of the southern United States. These restoration efforts entail the reduction of stand density by harvesting and midstory competition control as well as the reintroduction of...

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